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<mods:title id="E17A27EEBD9F6BA80A55033DE1C24E64">Acotylea (Platyhelminthes, Polycladida) from the southern and western Iberian Peninsula, with the description of five new species</mods:title>
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<mods:namePart id="53E8BB15AEF35B5C5E7BBBBCF57985A8">Pérez-García, Patricia</mods:namePart>
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<mods:namePart id="4605D140F6B6991A6C8A9444001399AF">Gouveia, Filipa</mods:namePart>
<mods:affiliation id="410CADC9475C6F85859BF0FE02989BCF">EPCV-Departamento de Ciências da Vida, Universidade Lusófona, Campo Grande, 376, 1749 - 024 Lisboa, Portugal</mods:affiliation>
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<mods:namePart id="E5ACCD54C470992D563A76165A4AAE97">Calado, Gonçalo</mods:namePart>
<mods:affiliation id="9A04A50F0CDFB3DCAC6DB55B6DB0669C">EPCV-Departamento de Ciências da Vida, Universidade Lusófona, Campo Grande, 376, 1749 - 024 Lisboa, Portugal &amp; MARE - Centro de Ciências do Mar e do Ambiente (FCT / NOVA), Campus de Caparica, Caparica, Portugal</mods:affiliation>
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<mods:namePart id="1E03A6BDF7231EFA547D3AAC17227F5B">Noreña, Carolina</mods:namePart>
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<mods:namePart id="915061295DFB227C3CC0FBF0B2759374">Cervera, Juan Lucas</mods:namePart>
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<mods:affiliation id="C7CA35E75F78FD3F0A815EE63455FF65">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales, Campus de Excelencia Internacional del Mar (CEI MAR), Av. República Saharahui, s / n, Ap. 40. 11510, Puerto Real (Cádiz), Spain &amp; Instituto Universitario de Investigación Marina (INMAR), Campus de Excelencia Internacional del Mar (CEI · MAR), Universidad de Cádiz, Av. República Saharahui, s / n, Ap. 40. 11510, Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:date id="8BD39412CC0A187DA1066D540B72F55F">2024</mods:date>
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<mods:number id="A45FE70DF678BCAEFFA54CF747FECE49">100</mods:number>
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<treatment id="0201B053C7EE5DFCA7D5F4952B7732A8" ID-DOI="http://doi.org/10.5281/zenodo.14052247" ID-Zenodo-Dep="14052247" ID-arpha="0201B053-C7EE-5DFC-A7D5-F4952B7732A8" LSID="urn:lsid:plazi:treatment:0201B053C7EE5DFCA7D5F4952B7732A8" httpUri="http://treatment.plazi.org/id/0201B053C7EE5DFCA7D5F4952B7732A8">
<subSubSection id="BD13A3495E91853C92B64F6188ECA7B9" type="nomenclature">
<paragraph id="5268B687391FA6509A2D26384D7485F7">
<taxonomicName id="F4B39075BF235041AC9EC3B39E605CA5" ID-CoL="38JTY" authority="Haswell, 1907" authorityName="Haswell" authorityYear="1907" family="Gnesiocerotidae" genus="Echinoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="celerrima">
<emphasis id="9D5EE5F49DA30C0BE79E109F7A27AD79" italics="true">Echinoplana celerrima</emphasis>
<bibRefCitation id="1FD530E1E88D5D57E3EA92B8FED75024" DOI="10.1111/j.1096-3642.1907.tb00455.x" author="Haswell WA" journalOrPublisher="Transactions of the Linnean Society of London" pagination="465-485" refId="B30" refString="Haswell WA (1907) Observations on Australian polyclads. Transactions of the Linnean Society of London 9: 465 - 485. https: // doi. org / 10.1111 / j. 1096 - 3642.1907. tb 00455. x" title="Observations on Australian polyclads." volume="9" year="1907">Haswell, 1907</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0B069B8204C6F765E342B58615FF783D" type="description">
<paragraph id="3AEE62CCF0817406EBD808FC6A233A66">
<figureCitation id="3EE221FD64E3227F5D766DC6076A10A6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Echinoplana celerrima (MNHNC MB 16-000120): A. Dorsal view of the living specimen; B. Detail of the eyes; C. Ventral view of the living specimen, showing the male copulatory apparatus." figureDoi="10.3897/zse.100.128211.figure10" httpUri="https://binary.pensoft.net/fig/1169823">Fig. 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EG1BI" type="material">
<paragraph id="DEB3A3A5A7DE36E85943FB4329C8FDD9">
<heading id="64554945FFD292FA434692E3AB3D4100" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="1009C0DD7AC06C1D852C0745927F677F">
<materialsCitation id="623AC7BCB43CCC89B7A32304924BAE27" collectingDate="2018-05-13" collectionCode="MNHNC" specimenCode="MB 16-000104" specimenCount="1">
<abbrev id="ABBRID0EM1BI" xlink_title="Museu Nacional de História Natural e da Ciência of Lisbon">
<collectionCode id="348AE2E76242B778946496C323766909">MNHNC</collectionCode>
</abbrev>
<specimenCode id="022A09D938736380BD048D67C983E4CB" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB 16-000104</specimenCode>
, Station 7,
<date id="97414BD0917621B8F7C4F26B685E1A1A" value="2018-05-13">
<collectingDate id="1FBE2E2125C1A60BC81AF786DE25DD0F" value="2018-05-13">13 May 2018</collectingDate>
</date>
,
<quantity id="F7C37B8D5ACB7EC01ED047F50E7B988B" metricMagnitude="-2" metricUnit="m" metricValue="1.6" unit="mm" value="16.0">16 mm</quantity>
long. All the measurements from the description refer to this specimen
</materialsCitation>
; •
<materialsCitation id="C237A518EDF5BF23D0CFFAC9355F47E4" collectingDate="2015-02-19" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="0865257450081FC9526F83C7CDF479D0" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="8055379003B4D455D32B1F34AF5782E5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3983, Station 11,
<date id="D6FDFD7421CCDC4346F5B8F32882B2A7" value="2015-02-19">
<collectingDate id="E6C73122F69554EB7E71437974CEF1A1" value="2015-02-19">19 February 2015</collectingDate>
</date>
,
<quantity id="19513D79F3D856A91250068B4927D40F" metricMagnitude="-3" metricUnit="m" metricValue="9.0" unit="mm" value="9.0">9 mm</quantity>
long
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EU1BI" type="Morphology">
<paragraph id="265E5CA13E19A96CB61B270F69B90A99">
<heading id="1313A8993E0CA1E614714649A40472EF" reason="title">Morphology.</heading>
</paragraph>
<paragraph id="95E6DCF374F3EF84D3CBDEE4F16C47D2">
Elongated worm, wider in the anterior margin with a blunt end (Fig.
<figureCitation id="05CA35C4FED952A5930A3F427219D59E" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Echinoplana celerrima (MNHNC MB 16-000120): A. Dorsal view of the living specimen; B. Detail of the eyes; C. Ventral view of the living specimen, showing the male copulatory apparatus." figureDoi="10.3897/zse.100.128211.figure10" httpUri="https://binary.pensoft.net/fig/1169823">10 A</figureCitation>
). Length between 9 and
<quantity id="5817D9EECD9D04712EAE5C793080308D" metricMagnitude="-2" metricUnit="m" metricValue="1.6" unit="mm" value="16.0">16 mm</quantity>
alive (12.5 ±
<quantity id="DFE962DD66A5440BFF55EE7292017FF0" metricMagnitude="-3" metricUnit="m" metricValue="4.95" unit="mm" value="4.95">4.95 mm</quantity>
). Dorsal colour caramel and translucent intestinal branches are easily observed and vary from brown to green. Central part of the body darkish brown. Tentacles absent. Tentacular and cerebral eye clusters present, distributed parallelly and in two long rows (Fig.
<figureCitation id="CE59DC32E2DF64A237A86419101A478D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Echinoplana celerrima (MNHNC MB 16-000120): A. Dorsal view of the living specimen; B. Detail of the eyes; C. Ventral view of the living specimen, showing the male copulatory apparatus." figureDoi="10.3897/zse.100.128211.figure10" httpUri="https://binary.pensoft.net/fig/1169823">10 B</figureCitation>
). Tentacular and cerebral eyes clustered in approximately 32 and 43 eyes, respectively. Pharynx ruffled, located in the second third of the body. Oral pore in the last third of the pharynx.
</paragraph>
<caption id="AA368DFD4EEA35846B78B8C7ED96113B" ID-DOI="10.3897/zse.100.128211.figure10" ID-arpha="391AD206-85FB-5C8F-B684-F05E9302473C" httpUri="https://binary.pensoft.net/fig/1169823" startId="F10">
<paragraph id="58770FEE4F5243D5EBE902515C08A8A5">
<label id="CF97E86AB8473A9B775FEA7D77D3F48F">Figure 10.</label>
</paragraph>
<paragraph id="50F37242340EA650A3F661E90E4DCD2E">
<taxonomicName id="5AD0BB039D80EA5ED7DC0428D4F12213" authorityName="Haswell" authorityYear="1907" family="Gnesiocerotidae" genus="Echinoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="celerrima">
<emphasis id="F8C4B7DA4D7C9BA2E7458B61D4E2B554" italics="true">Echinoplana celerrima</emphasis>
</taxonomicName>
(
<abbrev id="ABBRID0EV2BI" xlink_title="Museu Nacional de História Natural e da Ciência of Lisbon">MNHNC</abbrev>
MB 16-000120):
<emphasis id="10CECDDC29C39FBED34694D6442587ED" bold="true">A.</emphasis>
Dorsal view of the living specimen;
<emphasis id="6307FFD8463619ABFD7EB862CEDC3291" bold="true">B.</emphasis>
Detail of the eyes;
<emphasis id="75C83FC0EA545373C9D6A456D19E61D6" bold="true">C.</emphasis>
Ventral view of the living specimen, showing the male copulatory apparatus.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EE3BI" type="Reproductive system">
<paragraph id="7836AB88D058DB9DDE7490DC94A6205F">
<heading id="5C4969964CCB199F9203C6EF6BEB16AD" reason="title">Reproductive system.</heading>
</paragraph>
<paragraph id="430378424AECC6546378FF3BF6C941E0">
Genital pores separated (separated approximately
<quantity id="C6270E562DC16B627527B26322D7DBE5" metricMagnitude="-3" metricUnit="m" metricValue="2.26" unit="mm" value="2.26">2.26 mm</quantity>
). Male apparatus (
<quantity id="1E4E6C007EB5B755DCDE2B2949335862" metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
long) characterised by a seminal vesicle, an interpolated prostatic vesicle, and a conspicuous cirrus shown by transparency (Fig.
<figureCitation id="2D0CC68DF13F804BE39D3938B4C78884" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Echinoplana celerrima (MNHNC MB 16-000120): A. Dorsal view of the living specimen; B. Detail of the eyes; C. Ventral view of the living specimen, showing the male copulatory apparatus." figureDoi="10.3897/zse.100.128211.figure10" httpUri="https://binary.pensoft.net/fig/1169823">10 C</figureCitation>
). Female system with a large vagina (
<quantity id="A0A7DDE35B3464DC5888FC84AC416115" metricMagnitude="-3" metricUnit="m" metricValue="2.29" unit="mm" value="2.29">2.29 mm</quantity>
), a ductus vaginalis, and Langs vesicle in its distal region.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EO3BI" type="remarks">
<paragraph id="839241F35D14670A06B241C5AE2DCD09">
<heading id="1CC736F34D7136E4B9BD05C952F798C8" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="18C1CDC1CB7C41DBAAFCC0A72F1E1233">
<taxonomicName id="963F23E5105BD7301651CC675750FD11" authorityName="Haswell" authorityYear="1907" family="Gnesiocerotidae" genus="Echinoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="celerrima">
<emphasis id="410E2A7FF9516EE2AF01F212B7559F0F" italics="true">Echinoplana celerrima</emphasis>
</taxonomicName>
is the only species of the genus
<taxonomicName id="9D3626DE992EA4887A5E7BC3ABDE9011" authorityName="Haswell" authorityYear="1907" family="Gnesiocerotidae" genus="Echinoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="genus">
<emphasis id="96FE19238DD020F3EFCD62DB34CC0A3C" italics="true">Echinoplana</emphasis>
</taxonomicName>
described by
<bibRefCitation id="05D6580640F5624733D2129AE9D5DD56" DOI="10.1111/j.1096-3642.1907.tb00455.x" author="Haswell WA" firstAuthor="Haswell" journalOrPublisher="Transactions of the Linnean Society of London" pagination="465-485" refId="B30" refString="Haswell WA (1907) Observations on Australian polyclads. Transactions of the Linnean Society of London 9: 465485. https://doi.org/10.1111/j.1096-3642.1907.tb00455.x" title="Observations on Australian polyclads." volume="9" year="1907">Haswell (1907)</bibRefCitation>
from Port Jackson, Sidney,
<collectingCountry id="292F7EEE6DC7ECAE543F9C2B47F0D257" name="Australia">Australia</collectingCountry>
. After that,
<bibRefCitation id="311CDAED239CF0EB67324FD672B78928" author="Galleni L" firstAuthor="Galleni" journalOrPublisher="Atti della Società Toscana de Scienze Naturali di Pisa" pagination="139-148" refId="B18" refString="Galleni L (1978) Policladi delle coste toscane. III. Echinoplana celerrima Haswell, Planoceridae nuovo per il Mediterraneo e note sul genere. Echinoplana. Atti. Atti della Società Toscana de Scienze Naturali di Pisa 85: 139148." title="Policladi delle coste toscane. III. Echinoplana celerrima Haswell, Planoceridae nuovo per il Mediterraneo e note sul genere. Echinoplana. Atti." volume="85" year="1978">Galleni (1978)</bibRefCitation>
reported this species in the Port of Livorno,
<collectingCountry id="243546B0AD78B88DA8603F1D33EBEEB0" name="Italy">Italy</collectingCountry>
, which was the first record of the species in the Mediterranean. Later,
<bibRefCitation id="9E8632516CA4634B27BDA69448FDB7C7" DOI="10.11646/zootaxa.1560.1.1" author="Holleman J" firstAuthor="Holleman" issue="1" journalOrPublisher="Zootaxa" pagination="1-17" refId="B32" refString="Holleman J (2007) Some New Zealand Polyclads (Platyhelminthes, Polycladida). Zootaxa 1560 (1): 117. https://doi.org/10.11646/zootaxa.1560.1.1" title="Some New Zealand Polyclads (Platyhelminthes, Polycladida)." volume="1560" year="2007">Holleman (2007)</bibRefCitation>
reported this species on the Pacific coast on the North Island of
<collectingCountry id="779C9801124C0AFE6DDF2C786F719EF9" name="New Zealand">New Zealand</collectingCountry>
.
<taxonomicName id="3C9630BC2A9E976EB5824DDC9B5950DE" authorityName="Haswell" authorityYear="1907" family="Gnesiocerotidae" genus="Echinoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="celerrima">
<emphasis id="ECEA6F48106B7EA682BFC5C012F08186" italics="true">E. celerrima</emphasis>
</taxonomicName>
was also recorded in the lagoon of
<collectingCountry id="105435DD22FE58A8DD4A7DA5882873E5" name="Tunisia">Tunis</collectingCountry>
(
<bibRefCitation id="E30025A02A2707F6DFFD4FBF3752F0C1" DOI="10.11646/zootaxa.2195.1.3" author="Gammoudi" etAl="et al." firstAuthor="Gammoudi" journalOrPublisher="Zootaxa" pagination="43-60" refId="B22" refString="Gammoudi M, Tekaya S, Noreña C (2009) Contribution to the knowledge of Acotylean Polyclads (Platyhelminthes, Polycladida) from Tunisian Coasts. Zootaxa 2195: 4360. https://doi.org/10.11646/zootaxa.2195.1.3" title="Contribution to the knowledge of Acotylean Polyclads (Platyhelminthes, Polycladida) from Tunisian Coasts." volume="2195" year="2009">Gammoudi et al. 2009</bibRefCitation>
;
<bibRefCitation id="1E7A8E2FAE0EAB89B82D8BD4ECEC6DC1" DOI="10.11646/zootaxa.4263.1.5" author="Gammoudi" firstAuthor="Gammoudi" issue="1" journalOrPublisher="Zootaxa" pagination="120-138" refId="B24" refString="Gammoudi M, Garbouj M, Egger B, Tekaya S (2017) Updated inventory and distribution of free-living flatworms from Tunisian waters. Zootaxa 4263 (1): 120138. https://doi.org/10.11646/zootaxa.4263.1.5" title="Updated inventory and distribution of free-living flatworms from Tunisian waters." volume="4263" year="2017">2017</bibRefCitation>
) and now in the southern Iberian Peninsula (present study). Considering the geographical distance between the specimens from
<collectingCountry id="2D2DEE43A5573098C3B45DC94299B21B" name="Australia">Australia</collectingCountry>
,
<collectingCountry id="745BBC05E28DBBC68D64856F8DDB5EE4" name="New Zealand">New Zealand</collectingCountry>
, and the Mediterranean Sea, the contrast of information provided by the morphological characteristics and the DNA analyses (using nuclear and mitochondrial markers) is crucial to elucidating whether the populations belong to the same species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EF5BI" type="biology">
<paragraph id="499509198135D0D67D98A09067A434C5">
<heading id="94BD2514724C01B645FC2AED6595CE6C" reason="title">Biology.</heading>
</paragraph>
<paragraph id="00877DA27550093EB7C7291B3D9F9C61">
In this study, the specimens were collected under stones in the intertidal zone. Furthermore,
<taxonomicName id="12B4010E6755EE04053F05A2D54F6456" authorityName="Haswell" authorityYear="1907" family="Gnesiocerotidae" genus="Echinoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="celerrima">
<emphasis id="8291D3B07D86AFB82E16BF386A3DF671" italics="true">Echinoplana celerrima</emphasis>
</taxonomicName>
frequently inhabits areas with human activity, such as harbours (
<bibRefCitation id="E01BC51EC1C35E0EA1F0A8B2F94E431F" DOI="10.1111/j.1096-3642.1907.tb00455.x" author="Haswell" firstAuthor="Haswell" journalOrPublisher="Transactions of the Linnean Society of London" pagination="465-485" refId="B30" refString="Haswell WA (1907) Observations on Australian polyclads. Transactions of the Linnean Society of London 9: 465485. https://doi.org/10.1111/j.1096-3642.1907.tb00455.x" title="Observations on Australian polyclads." volume="9" year="1907">Haswell 1907</bibRefCitation>
,
<bibRefCitation id="E379CB4493F08A8B6B98B706EAD6AB88" author="Galleni" firstAuthor="Galleni" journalOrPublisher="Atti della Società Toscana de Scienze Naturali di Pisa" pagination="139-148" refId="B18" refString="Galleni L (1978) Policladi delle coste toscane. III. Echinoplana celerrima Haswell, Planoceridae nuovo per il Mediterraneo e note sul genere. Echinoplana. Atti. Atti della Società Toscana de Scienze Naturali di Pisa 85: 139148." title="Policladi delle coste toscane. III. Echinoplana celerrima Haswell, Planoceridae nuovo per il Mediterraneo e note sul genere. Echinoplana. Atti." volume="85" year="1978">Galleni 1978</bibRefCitation>
). Hence, the specimen of Sagres was precisely collected in a rocky area attached to a harbour zone, supporting one of the possible scenarios about the dispersion of this species (
<bibRefCitation id="3DEDEB6A41FA92C4D48449DBD93E272B" author="Prudhoe" firstAuthor="Prudhoe" journalOrPublisher="Records of South Australian Museum" pagination="361-384" refId="B66" refString="Prudhoe S (1982) Polyc 1 ad turbellarians from the southern eoasts of Australia. Records of South Australian Museum 18: 361384." title="Polyc 1 ad turbellarians from the southern eoasts of Australia." volume="18" year="1982">Prudhoe 1982</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EC6BI" type="distribution">
<paragraph id="E4701913FE5FC43B756CA9990B484E11">
<heading id="424A33CC7B4B91E347D2A4BA1B00CFE9" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="4E2A945D43E852FAF2027EB9E8D00DDC">
As previously explained,
<taxonomicName id="C5C7534CB3E198B811A5318B9C1F1012" authorityName="Haswell" authorityYear="1907" family="Gnesiocerotidae" genus="Echinoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="celerrima">
<emphasis id="61FB87DAF5ED7BB8D53D71E2A817D02B" italics="true">Echinoplana celerrima</emphasis>
</taxonomicName>
is widely distributed on the southeastern coast of
<collectingCountry id="896C3EBE16C2A74DA7F996BAD7F1F808" name="Australia">Australia</collectingCountry>
(
<bibRefCitation id="FAF7BBFAAA06969D8BF0411D34DDC1D7" DOI="10.1111/j.1096-3642.1907.tb00455.x" author="Haswell" firstAuthor="Haswell" journalOrPublisher="Transactions of the Linnean Society of London" pagination="465-485" refId="B30" refString="Haswell WA (1907) Observations on Australian polyclads. Transactions of the Linnean Society of London 9: 465485. https://doi.org/10.1111/j.1096-3642.1907.tb00455.x" title="Observations on Australian polyclads." volume="9" year="1907">Haswell 1907</bibRefCitation>
;
<bibRefCitation id="446D07C94072ADE880F03C9DEEFB0405" author="Prudhoe" firstAuthor="Prudhoe" journalOrPublisher="Records of South Australian Museum" pagination="361-384" refId="B66" refString="Prudhoe S (1982) Polyc 1 ad turbellarians from the southern eoasts of Australia. Records of South Australian Museum 18: 361384." title="Polyc 1 ad turbellarians from the southern eoasts of Australia." volume="18" year="1982">Prudhoe 1982</bibRefCitation>
;
<bibRefCitation id="B37BFC673BD5B17CE7EDD4DE356616E1" DOI="10.1007/s00227-008-1056-2" author="Johnston and Lee" firstAuthor="Johnston" journalOrPublisher="Marine Biology" pagination="555-561" refId="B38" refString="Johnston EL, Lee KM (2008) Brooding behaviour and reproductive success in two species of free-living simultaneous hermaphrodites. Marine Biology 155: 555561. https://doi.org/10.1007/s00227-008-1056-2" title="Brooding behaviour and reproductive success in two species of free-living simultaneous hermaphrodites." volume="155" year="2008">Johnston and Lee 2008</bibRefCitation>
;
<bibRefCitation id="E9B74A3A81BC2A6E9596B154AD48F2D0" DOI="10.11646/zootaxa.5024.1.1" author="Rodriguez" etAl="et al." firstAuthor="Rodriguez" issue="1" journalOrPublisher="Zootaxa" pagination="1-63" refId="B73" refString="Rodríguez J, Hutchings PA, Williamson JE (2021) Biodiversity o intertidal marine flatworms (Polycladida, Platyhelminthes) in southeastern Australia. Zootaxa 5024 (1): 163. https://doi.org/10.11646/zootaxa.5024.1.1" title="Biodiversity o intertidal marine flatworms (Polycladida, Platyhelminthes) in southeastern Australia." volume="5024" year="2021">Rodríguez et al. 2021</bibRefCitation>
), Livorno,
<collectingCountry id="C192CA3DC7A3404BE0B78A24F27ABC0C" name="Italy">Italy</collectingCountry>
(
<bibRefCitation id="DD95358869101DED77D2F97F060F9E5B" author="Galleni" firstAuthor="Galleni" journalOrPublisher="Atti della Società Toscana de Scienze Naturali di Pisa" pagination="139-148" refId="B18" refString="Galleni L (1978) Policladi delle coste toscane. III. Echinoplana celerrima Haswell, Planoceridae nuovo per il Mediterraneo e note sul genere. Echinoplana. Atti. Atti della Società Toscana de Scienze Naturali di Pisa 85: 139148." title="Policladi delle coste toscane. III. Echinoplana celerrima Haswell, Planoceridae nuovo per il Mediterraneo e note sul genere. Echinoplana. Atti." volume="85" year="1978">Galleni 1978</bibRefCitation>
),
<collectingCountry id="E45FF818E4F31FE598790C72089369FC" name="New Zealand">New Zealand</collectingCountry>
(
<bibRefCitation id="AF879AE23386C2A5D89521E4A12CA371" DOI="10.11646/zootaxa.1560.1.1" author="Holleman" firstAuthor="Holleman" issue="1" journalOrPublisher="Zootaxa" pagination="1-17" refId="B32" refString="Holleman J (2007) Some New Zealand Polyclads (Platyhelminthes, Polycladida). Zootaxa 1560 (1): 117. https://doi.org/10.11646/zootaxa.1560.1.1" title="Some New Zealand Polyclads (Platyhelminthes, Polycladida)." volume="1560" year="2007">Holleman 2007</bibRefCitation>
),
<collectingCountry id="D3210B6DB01D0832DB2E97A766AA2E5B" name="Tunisia">Tunisia</collectingCountry>
(
<bibRefCitation id="36B41ADD686BEA38CECB5C8E46F8DF91" DOI="10.11646/zootaxa.2195.1.3" author="Gammoudi" etAl="et al." firstAuthor="Gammoudi" journalOrPublisher="Zootaxa" pagination="43-60" refId="B22" refString="Gammoudi M, Tekaya S, Noreña C (2009) Contribution to the knowledge of Acotylean Polyclads (Platyhelminthes, Polycladida) from Tunisian Coasts. Zootaxa 2195: 4360. https://doi.org/10.11646/zootaxa.2195.1.3" title="Contribution to the knowledge of Acotylean Polyclads (Platyhelminthes, Polycladida) from Tunisian Coasts." volume="2195" year="2009">Gammoudi et al. 2009</bibRefCitation>
,
<bibRefCitation id="DAA02B72423754D07850A327084F9836" DOI="10.11646/zootaxa.4263.1.5" author="Gammoudi" firstAuthor="Gammoudi" issue="1" journalOrPublisher="Zootaxa" pagination="120-138" refId="B24" refString="Gammoudi M, Garbouj M, Egger B, Tekaya S (2017) Updated inventory and distribution of free-living flatworms from Tunisian waters. Zootaxa 4263 (1): 120138. https://doi.org/10.11646/zootaxa.4263.1.5" title="Updated inventory and distribution of free-living flatworms from Tunisian waters." volume="4263" year="2017">2017</bibRefCitation>
), and
<collectingRegion id="8A221154E7E145A564AB03D444E63232" country="Spain" name="Catalunya">Catalonia</collectingRegion>
,
<collectingCountry id="F69686AA2550E9FAEF7FCD21B3C234B5" name="Spain">Spain</collectingCountry>
(
<bibRefCitation id="CE63E1F5462C6CCA426EC7F1B7859118" author="Gammoudi and Tekaya" firstAuthor="Gammoudi" issue="1-4" journalOrPublisher="Bulletin de la Societé Zoologique de France" pagination="197-213" refId="B21" refString="Gammoudi M, Tekaya S (2012) Distribution in the western Mediterranean of some Polyclads (Platyhelminthes). Bulletin de la Societé Zoologique de France 137 (14): 197213." title="Distribution in the western Mediterranean of some Polyclads (Platyhelminthes)." volume="137" year="2012">Gammoudi and Tekaya 2012</bibRefCitation>
). Our specimens provide the first reports of this species on the North-Atlantic shores of
<collectingCountry id="2FA2BCAB17661C4A8A8C537EFBE3263F" name="Portugal">Portugal</collectingCountry>
and
<collectingRegion id="1591D76F38EDC6E82F7BF7E89FB6C1FB" country="Spain" name="Andalucia">Andalusia</collectingRegion>
(
<collectingCountry id="2CD1C2004829AB5426EF4C1ADB9E972C" name="Spain">Spain</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,126 @@
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<subSubSection id="5A016D278ACB560AADF5257E10D2BFF2" type="nomenclature">
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<taxonomicName id="2A8E4F9DCFC8550A3F32C6B3C8842FB6" authority="Pierre, Fl. Forest. Cochinch." authorityName="Pierre, Fl. Forest. Cochinch." class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="section" section="Macrostigma">
<emphasis id="24D456C97DE5FE288DAE549532F89CF0" italics="true">Garcinia section Macrostigma</emphasis>
Pierre, Fl. Forest. Cochinch.
</taxonomicName>
1 (6): 63. 1883;
<treatmentCitation id="3400F11DC9D7F8044C64A02828F6A949" author="M. Gaudeul et al." journal="PhytoKeys" page="94" score="9.729594737742866" volume="239" year="2024">
<bibCitation id="1F0A0467468F3080D21743949F902C31" author="M. Gaudeul et al." journal="PhytoKeys" page="94" score="9.729594737742866" volume="239" year="2024">
M. Gaudeul et al.,
<bibCitation id="772830D4235E572C00F0C19EDA84138C" journal="PhytoKeys" page="94" score="9.729594737742866" volume="239" year="2024">PhytoKeys 239: 94. 2024</bibCitation>
</bibCitation>
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.
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EGPAG" type="material">
<paragraph id="92014B29F04D9EDFEF96D3E26B784EBA">
<heading id="65EB76BDD05793EE87BDAE6B190ED8DD" reason="title">
<typeStatus id="EDFB63179601D42B44D6F22FA55C8B98">Type</typeStatus>
.
</heading>
</paragraph>
<paragraph id="55F7729CEDF937AAE5A66948FBA8ED48">
<taxonomicName id="301FD45BE5F0F4785E024C3BCC9375B8" authority="Miq., Ann. Mus. Bot. Lugduno-Batavi" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="latissima">
<emphasis id="26A5664B00FF47FF93E7A891DC038CAD" italics="true">Garcinia latissima</emphasis>
Miq., Ann. Mus. Bot. Lugduno-Batavi
</taxonomicName>
1: 209. 1864.
</paragraph>
<paragraph id="6CF121781273558F272D5CFB797DDEB2">
<taxonomicName id="C6787BC9F78D3D99152709F9804C4D58" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="section" section="Macrostigma">
<emphasis id="410693B0AB307F927535D06B2391E287" italics="true">Garcinia section Macrostigma</emphasis>
</taxonomicName>
is characterized by its axillary or terminal cymose inflorescences with two to many flowers (in a simple cyme or in fascicles of several simple cymes), or a solitary flower (in female flowers); 4 ( 5) - merous flowers [4 - merous, e. g.,
<taxonomicName id="65984099BC1A22978B600D81F7B30C82" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nuntasaenii">
<emphasis id="24DE55A3FB12B088648B0A5E69A10C06" italics="true">G. nuntasaenii</emphasis>
</taxonomicName>
; 5 - merous, e. g.,
<taxonomicName id="2A09CEBEF60194A6AA7F27A28F6C0029" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="phuongmaiensis">
<emphasis id="3AE447D6EF17959208F731B68ED88D78" italics="true">G. phuongmaiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="9A1AA3E0A6290AC38B7E1FDC350F7259" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="71A5474E75C2C1EC82B7F5362BB0B63F" italics="true">G. prainiana</emphasis>
</taxonomicName>
]; male flowers with numerous stamens united into a central column, sometimes lobed with lobes equaling number of petals [a single weakly 4 - lobed bundle, e. g.,
<taxonomicName id="24FFEEAC71F65AE38C749911C92B604E" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nuntasaenii">
<emphasis id="162D763E55E66E02E9FA7A5EE471D00A" italics="true">G. nuntasaenii</emphasis>
</taxonomicName>
], or into completely separate antepetalous bundles [united into 5 bundles, e. g.,
<taxonomicName id="761790AB3962D4D6688A91204B34CD36" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="phuongmaiensis">
<emphasis id="BD6019F7CFDC9219A2F4DB6021AD4E74" italics="true">G. phuongmaiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="AB3F7269F57AB7A36673CE8C8B85CC32" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="50C38D1EB449771FF0032DDD389C9C4C" italics="true">G. prainiana</emphasis>
</taxonomicName>
]; androecium often adnate to the petals to varying degrees; 2 - thecous anthers; pistillode absent in male flowers, but rudimentary or well-developed pistillode present in some species; ( 3) 48 - locular ovaries; unlobed or lobed and smooth or papillate stigmas; and fruits with a smooth surface, unlobed or faintly, shallowly or deeply lobed. Distinguishing morphological characters of this section based on
<bibRefCitation id="9CD3CA3D8D77F8FE5A9CAFDADEB47E92" DOI="10.3897/phytokeys.239.112563" author="Gaudeul" etAl="et al." firstAuthor="Gaudeul" journalOrPublisher="PhytoKeys" pagination="73-105" refId="B9" refString="Gaudeul M, Sweeney P, Munzinger J (2024) An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence. PhytoKeys 239: 73105. https://doi.org/10.3897/phytokeys.239.112563" title="An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence." volume="239" year="2024">Gaudeul et al. 2024</bibRefCitation>
, which includes additional information.
</paragraph>
<paragraph id="569A7F886F4FB29E33C6FA128F9FE28B">
A section of 29 species worldwide (
<bibRefCitation id="128169F016BFEE70DBD99BDB4E4B28F8" DOI="10.3897/phytokeys.239.112563" author="Gaudeul" etAl="et al." firstAuthor="Gaudeul" journalOrPublisher="PhytoKeys" pagination="73-105" refId="B9" refString="Gaudeul M, Sweeney P, Munzinger J (2024) An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence. PhytoKeys 239: 73105. https://doi.org/10.3897/phytokeys.239.112563" title="An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence." volume="239" year="2024">Gaudeul et al. 2024</bibRefCitation>
); two species in
<collectingCountry id="723532BE9BA2A9B68B70BB4E37368124" name="Thailand">Thailand</collectingCountry>
(i. e.,
<taxonomicName id="6CAFEBA7003C08834A0C844AF01A6696" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nuntasaenii">
<emphasis id="B9FFC450D3BE8E0FE7D4E3740F09DF28" italics="true">G. nuntasaenii</emphasis>
</taxonomicName>
and
<taxonomicName id="E62C7F1DBF31F179819F40DE64AF3D84" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="7956F29C0C86A2CEF19BA46783BAE447" italics="true">G. prainiana</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,217 @@
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<mods:namePart id="4D7734E2163FCAE88E886CEC3D8F1EC7">Gouveia, Filipa</mods:namePart>
<mods:affiliation id="D0BCA59A09E351DFC4E50B3C22B9FF0B">EPCV-Departamento de Ciências da Vida, Universidade Lusófona, Campo Grande, 376, 1749 - 024 Lisboa, Portugal</mods:affiliation>
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<paragraph id="255F1D80721D86BC23ACCCA8E85A18FE">
<taxonomicName id="4A1AF5C3C69250564CE8B64CA85377F6" ID-CoL="4FM4F" authority="Perez-Garcia, Norena &amp; Cervera, 2018" authorityName="Perez-Garcia, Norena &amp; Cervera" authorityYear="2018" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="caetaria">
<emphasis id="BF65F178325AA047DD87E6C54A453FA3" italics="true">Phaenoplana caetaria</emphasis>
Pérez-García, Noreña &amp; Cervera, 2018
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="DD02196EAA56FE478FAF974F142FECC0" type="description">
<paragraph id="8683F2830DA4B5C009B70966334CA314">
<figureCitation id="EDB752C663823AE48982DC9D97DC2E70" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Phaenoplana caetaria (Two living specimens of Station 16): A. Dorsal view of specimen A; B. Detail of the eyes; C. Dorsal view of specimen B; D. Detail of the eyes." figureDoi="10.3897/zse.100.128211.figure15" httpUri="https://binary.pensoft.net/fig/1169828">Fig. 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPTAK" type="material">
<paragraph id="A7435B6B58C1DE7A1C9EDA0375EE9676">
<heading id="7BDD9A39AEFF8CEA4ADE6F5DCAC96D9F" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="0A02FD1DA00E1310D3A482B20D93805E">
Specimen found in the field, Station 16,
<date id="96359F45E7C09D107E3E1EBDAF8D4590" value="2015-05-19">19 May 2015</date>
,
<quantity id="75431EC48B9D1BBE10ABC53E7027F546" metricMagnitude="-2" metricUnit="m" metricValue="1.3" unit="mm" value="13.0">13 mm</quantity>
; specimen found in the field, Station 16,
<date id="3E5C43E85E2CEA58F88FAD8212425F33" value="2015-05-19">19 May 2015</date>
,
<quantity id="DD5F8DDA9A46A3FDB05713CB61E576F5" metricMagnitude="-2" metricUnit="m" metricValue="2.1" unit="mm" value="21.0">21 mm</quantity>
; specimen found in the field, Station 17,
<date id="50B581C27DC00E9B6AD1EEAAB8D02D27" value="2015-04-19">19 April 2015</date>
,
<quantity id="C47EB4E661F2CE4F9658E33822107FCB" metricMagnitude="-2" metricUnit="m" metricValue="2.4" unit="mm" value="24.0">24 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUTAK" type="type locality">
<paragraph id="96A40F5E730E249D5F2BA6C123725412">
<heading id="5F6DF27ADBB06F9C6E2D689129F11204" reason="title">
<typeStatus id="6D1691B1FFE446A9A72C8F15056E6C86">Type</typeStatus>
locality.
</heading>
</paragraph>
<paragraph id="8DE2B15236EE11EBD423AA7FDE9B5198">
Punta Carnero, Cádiz,
<collectingCountry id="5BA1EFD2E4F69400C08D5C03BDF7D365" name="Spain">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZTAK" type="description">
<paragraph id="CDFB213111C1FF4AB2AB3CAFC49301BB">
<heading id="C8BA6C51C4C82041F7C0CF9AC7BB709B" reason="title">Description.</heading>
</paragraph>
<paragraph id="92D0F3EA4864218B013B9A5519E049AB">
<taxonomicName id="16BE8B245400088CE62D4F80559A420E" authorityName="Faubel" authorityYear="1983" family="Stylochoplanidae" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="family">Stylochoplanidae</taxonomicName>
with elongated body, wider anteriorly, and folded within the margins. Length between
<quantity id="9888129E1A466D591E404FCEB846ED0D" metricMagnitude="-2" metricUnit="m" metricValue="2.4" unit="mm" value="24.0">24 mm</quantity>
and
<quantity id="B0C89439918B1F076772A94C6A1FF40A" metricMagnitude="-2" metricUnit="m" metricValue="1.3" unit="mm" value="13.0">13 mm</quantity>
(19.3 ± 5.69). Colour of the dorsal surface light brown with yellowish spots (Fig.
<figureCitation id="CCC7FA18FCA69CB2687B181531B4AE6B" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Phaenoplana caetaria (Two living specimens of Station 16): A. Dorsal view of specimen A; B. Detail of the eyes; C. Dorsal view of specimen B; D. Detail of the eyes." figureDoi="10.3897/zse.100.128211.figure15" httpUri="https://binary.pensoft.net/fig/1169828">15 A, C</figureCitation>
). Ventral surface pale brown. Tentacles absent. Tentacular and cerebral eyes arranged in two clusters, with approximately 15 tentacular eyes and 27 cerebral eyes in each cluster (Fig.
<figureCitation id="8B70FC4B215F9F83C5DAA46BA8CB040A" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Phaenoplana caetaria (Two living specimens of Station 16): A. Dorsal view of specimen A; B. Detail of the eyes; C. Dorsal view of specimen B; D. Detail of the eyes." figureDoi="10.3897/zse.100.128211.figure15" httpUri="https://binary.pensoft.net/fig/1169828">15 B, D</figureCitation>
). Pharynx ruffled, positioned slightly anteriorly, between the first and second thirds of the body. Opening of the mouth in the last third of the pharynx. Genital pores separated. Male copulatory apparatus composed of a true seminal vesicle, an interpolated prostatic vesicle, and a penis papilla with a penis rod. Female complex consists of a well-developed external vagina (vagina bulbosa) and reduced Langs vesicle (for more details, see
<bibRefCitation id="9ACFD1AE7B3DE5C9698BB9D267E2E719" DOI="10.1007/s12526-018-0900-y" author="Perez-Garcia" etAl="et al." firstAuthor="Perez-Garcia" journalOrPublisher="Marine Biodiversity" pagination="1187-1195" refId="B63" refString="Pérez-García P, Noreña C, Cervera JL (2019) Two new acotylean flatworms (Polycladida) of two genera unrecorded in the Eastern Atlantic. Marine Biodiversity 49: 11871195. https://doi.org/10.1007/s12526-018-0900-y" title="Two new acotylean flatworms (Polycladida) of two genera unrecorded in the Eastern Atlantic." volume="49" year="2019">Pérez-García et al. 2019</bibRefCitation>
).
</paragraph>
<caption id="CA1AE6099228ADCEB1099D7F1A2FDAFF" ID-DOI="10.3897/zse.100.128211.figure15" ID-arpha="5298155F-C507-518B-A467-4AB6F03E49E5" httpUri="https://binary.pensoft.net/fig/1169828" startId="F15">
<paragraph id="CE14418472FABA94C5D9EE3337CA322F">
<label id="489D1B0CC2BCD733AE7C4B7D69CA8CE5">Figure 15.</label>
</paragraph>
<paragraph id="F99A7AAA8ACD47963F908E43463B3DD7">
<taxonomicName id="9624AC84B85822B4A46E9BBF2A92D3D0" authorityName="Perez-Garcia, Norena &amp; Cervera" authorityYear="2018" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="caetaria">
<emphasis id="37952A34ADDE15814B4499B29D085F39" italics="true">Phaenoplana caetaria</emphasis>
</taxonomicName>
(Two living specimens of Station 16):
<emphasis id="8DE3B77DC609A6A5C2630547BAD84EF5" bold="true">A.</emphasis>
Dorsal view of specimen A;
<emphasis id="38342F07A2AA31D5678FF148FE7F6F75" bold="true">B.</emphasis>
Detail of the eyes;
<emphasis id="5EDDCADF8A58DC126B892BBADC7B6FB4" bold="true">C.</emphasis>
Dorsal view of specimen B;
<emphasis id="AE5909BB1F2D93B6E5319B7899B6A915" bold="true">D.</emphasis>
Detail of the eyes.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQVAK" type="remarks">
<paragraph id="787DFCF6C23B03D6D3C2995457ACC546">
<heading id="ED5A06BDDC659661EC3F9A65ACC07129" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="2DB4E57468D8A3595673D3DC8BD445A8">
After the original description of
<taxonomicName id="0EE5D0EE6B45D0DD64DFFC6CC04274D2" authority="(Perez-Garcia et al., 2019)" baseAuthorityName="Perez-Garcia" baseAuthorityYear="2019" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="caetaria">
<emphasis id="3F1FAE65F7F53F33B4889D22AB37E478" italics="true">Phaenoplana caetaria</emphasis>
(
<bibRefCitation id="7B7DBEC088B5A81526B79E1894147922" DOI="10.1007/s12526-018-0900-y" author="Perez-Garcia" etAl="et al." firstAuthor="Perez-Garcia" journalOrPublisher="Marine Biodiversity" pagination="1187-1195" refId="B63" refString="Pérez-García P, Noreña C, Cervera JL (2019) Two new acotylean flatworms (Polycladida) of two genera unrecorded in the Eastern Atlantic. Marine Biodiversity 49: 11871195. https://doi.org/10.1007/s12526-018-0900-y" title="Two new acotylean flatworms (Polycladida) of two genera unrecorded in the Eastern Atlantic." volume="49" year="2019">Pérez-García et al., 2019</bibRefCitation>
)
</taxonomicName>
,
<bibRefCitation id="93F25FA5358621D0B097DD86DF5AD428" DOI="10.12782/specdiv.24.1" author="Oya Y &amp; Kajihara H" firstAuthor="Oya" journalOrPublisher="Species Diversity" pagination="1-6" refId="B57" refString="Oya Y, Kajihara H (2019) A New Species of Phaenoplana (Platyhelminthes: Polycladida) from the Ogasawara Islands. Species Diversity 24: 16. https://doi.org/10.12782/specdiv.24.1" title="A New Species of Phaenoplana (Platyhelminthes: Polycladida) from the Ogasawara Islands." volume="24" year="2019">Oya and Kajihara (2019)</bibRefCitation>
described a new species of
<taxonomicName id="B50C011D207239CB0F6BFFDFFFE86979" authorityName="Faubel" authorityYear="1983" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="genus">
<emphasis id="AA6D249016A2A8E62080D693543F1E4B" italics="true">Phaenoplana</emphasis>
</taxonomicName>
from
<collectingCountry id="CAD8C8714ABBE4091F4EA5F5FBFFC1B8" name="Japan">Japan</collectingCountry>
(
<taxonomicName id="8DC765D408607AC6AF23127004CB808B" authorityName="Oya &amp; Kajihara" authorityYear="2019" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="kopepe">
<emphasis id="DA76337642B0F93B819E1457EAFF523F" italics="true">P. kopepe</emphasis>
</taxonomicName>
), increasing the number of species in this genus to six. In this context,
<taxonomicName id="6433BDCD65B2B50D9D5A1D75A161476A" authorityName="Oya &amp; Kajihara" authorityYear="2019" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="kopepe">
<emphasis id="5E5D7C123C81205B131CFD6999DE8978" italics="true">P. kopepe</emphasis>
</taxonomicName>
is differentiated from
<taxonomicName id="9791B8C0B2BC095B29BAF88AD2C47AEB" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="caetaria">
<emphasis id="82C27DB266B1137BF9FD40898AF3DF0E" italics="true">P. caetaria</emphasis>
</taxonomicName>
by the presence of tentacles, a muscular bulb that surrounds the prostatic vesicle, the shaft of the penis, and the male atrium, a character that is missing in
<taxonomicName id="A1B735F0B7DA1BCD2A17E40C6DE1E31B" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="caetaria">
<emphasis id="42932E7E3A21F62B7B2C4C38E8E088F9" italics="true">P. caetaria</emphasis>
</taxonomicName>
, which shows instead a very characteristic twisted penis rod. Representatives of
<taxonomicName id="8517C91E68CE96D58CA3023E815CE2E7" authorityName="Faubel" authorityYear="1983" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="genus">
<emphasis id="0AC99FAA8F6494DBC0F4A2290FCA9DA3" italics="true">Phaenoplana</emphasis>
</taxonomicName>
have a predominantly Pacific distribution (5 known species) compared to the Mediterranean or Atlantic (two species), although the record of
<taxonomicName id="26DFC4A4519C89EBA7033B1582482ABE" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="caetaria">
<emphasis id="1C20655DA7AFE47A86AB2D1AC099AAD6" italics="true">P. caetaria</emphasis>
</taxonomicName>
shows a clear expansion of the specific distribution of
<taxonomicName id="EDA609DF807C98AD9A7F3FE32927417A" authorityName="Faubel" authorityYear="1983" family="Stylochoplanidae" genus="Phaenoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="genus">
<emphasis id="CF61145AE574DC448B298DAED4C5CC94" italics="true">Phaenoplana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVYAK" type="biology">
<paragraph id="4FF8C040ED1AA232F0878F30395BA7EF">
<heading id="877EBE9FE55863D2D4176D4E28B4EEEC" reason="title">Biology.</heading>
</paragraph>
<paragraph id="2B8E65BB8AF2F29673931B65311C8684">
This species is found under stones in the intertidal area of La Ballenera and Punta Carnero Beach, Cadiz,
<collectingCountry id="BF639B0B87672C92D6365DF3E5B2410D" name="Spain">Spain</collectingCountry>
.
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</treatment>
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@ -69,7 +71,7 @@ Chen &amp; Jiang
<heading id="A41A14BF41E7799213F760D362047137" reason="title">Type material.</heading>
</paragraph>
<paragraph id="EFCCCD20FB883FCB8E97F1AF0B220F7B">
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<typeStatus id="FF50484A4C40A3E5B35DE14A6888603C" type="holotype">Holotype</typeStatus>
</emphasis>
@ -84,7 +86,7 @@ Chen &amp; Jiang
;
<collectingCounty id="B3F97EEF4EE689F87A296D88690AD761">Mengla County</collectingCounty>
;
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<location id="17A837BD473D94EF71CDEB834BDA02D8" LSID="urn:lsid:plazi:treatment:48B14866403B522EBA3CC03522C48AB8:17A837BD473D94EF71CDEB834BDA02D8" country="China" county="Mengla County" latitude="21.898167" longLatPrecision="12" longitude="101.27866" municipality="Xishuangbanna" name="Menglun Town" stateProvince="Yunnan Province">Menglun Town</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EK4AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.278667,21.898167]}">
<geoCoordinate id="3D17CC2C038CE3B5DE14B3D56D544A47" degrees="21" direction="north" minutes="53.89" orientation="latitude" precision="9" value="21.898167">21 ° 53.89 ' N</geoCoordinate>
@ -100,10 +102,10 @@ a. s. l.
<date id="3C5E1FF608F5E22761F644DB8EF8F18A" value="2019-05-12">
<collectingDate id="B72246FE20BF8C7EFE248EE6E93D0332" value="2019-05-12">12 May. 2019</collectingDate>
</date>
; Z-l
<collectorName id="07FA4B6F8CCFEE6CC38432A6BF866D43">Bai</collectorName>
, Z-g
<collectorName id="A84AA6B5FC2401F9C400EA74BAB9B692">Chen</collectorName>
;
<collectorName id="515813903BD797612AE363B0F77F1268">Z-l Bai</collectorName>
,
<collectorName id="7A0AEF28F27E1D5EC039CF3338A092CF">Z-g Chen</collectorName>
,
<collectorName id="0718585291AACCE9B879C6BC86792CCA">C Wang</collectorName>
,
@ -112,11 +114,11 @@ leg.;
<abbrev id="ABBRID0EP4AE" xlink_title="Langfang Normal University">
<collectionCode id="D9C95D9F9D1E18FD4D8E93624FB87CE4">LFNU</collectionCode>
</abbrev>
<collectorName id="C9C7DFAEC6C99FA28E850F6D5CDB966C">Proap</collectorName>
202405-1 [on slide]
<specimenCode id="E82D2BAF9A8296F1D7D7EACDD338ACD6">Proap 202405-1</specimenCode>
[on slide]
</materialsCitation>
.
<materialsCitation id="7996DA676AF1DD6E4F8C8AF1B9C6031B" collectingDate="2019-05-12" collectionCode="LFNU" collectorName="Bai &amp; Chen &amp; C Wang &amp; H Yu &amp; Proap" country="China" county="Mengla County" elevation="568" latitude="21.898167" location="Menglun Town" longLatPrecision="12" longitude="101.27866" municipality="Xishuangbanna" specimenCount="15" specimenCount-female="15" stateProvince="Yunnan" typeStatus="paratype">
<materialsCitation id="7996DA676AF1DD6E4F8C8AF1B9C6031B" collectingDate="2019-05-12" collectionCode="LFNU" collectorName="Bai &amp; Chen &amp; C Wang &amp; H Yu &amp; Proap" country="China" county="Mengla County" elevation="568" latitude="21.898167" location="Menglun Town" longLatPrecision="12" longitude="101.27866" municipality="Xishuangbanna" specimenCode="Proap 202405-2, Proap 202405-7" specimenCount="15" specimenCount-female="15" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="7C973E0DACC36DEE244DB89525C3C26F" bold="true" italics="true">
<typeStatus id="167F49CEDC04D95A4A1F97B2D81AF76C" type="paratype">Paratypes</typeStatus>
</emphasis>
@ -124,7 +126,11 @@ leg.;
<specimenCount id="90CC9763E34DC1B8069CA1FB6DC0611C" count="8" type="female">8 ♀♀</specimenCount>
[
<specimenCount id="15A01BEB28505E1229183B8E225965C4" count="5" type="female">5 ♀♀</specimenCount>
on slides, Proap 202405-2 Proap 202405 - 7; •
on slides,
<specimenCode id="DB5EE18682D15B38791485DECDC564E0">Proap 202405-2</specimenCode>
<specimenCode id="4C9B8E3AAF90316B3900C67EF4C66D66">Proap 202405-7</specimenCode>
; •
<specimenCount id="B259488B7E28A147A26AF9C95A293D77" count="2" type="female">2 ♀♀</specimenCount>
destroyed for DNA extraction]; same data as holotype;
<abbrev id="ABBRID0EW4AE" xlink_title="Langfang Normal University">

View file

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</mods:name>
<mods:name id="24B4B6724FC9DC4B9B4776C03C78741F" type="personal">
<mods:role id="AA1068DC7CB2A98C1310D46969C561AC">
<mods:roleTerm id="44060EF5F61DB92D0B25759DCC64F2AE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C038EFBAFFF4F3748E7B86E510F10332">Gouveia, Filipa</mods:namePart>
<mods:affiliation id="A4734FFC1FF046C7CAC91ABE6E047270">EPCV-Departamento de Ciências da Vida, Universidade Lusófona, Campo Grande, 376, 1749 - 024 Lisboa, Portugal</mods:affiliation>
</mods:name>
<mods:name id="B131DB1589EA35A08E2BF64FB135C064" type="personal">
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</mods:role>
<mods:namePart id="7A710199515992E0CF4AC1079A83641F">Calado, Gonçalo</mods:namePart>
<mods:affiliation id="E2F6634BA845B61C702D44420A45995F">EPCV-Departamento de Ciências da Vida, Universidade Lusófona, Campo Grande, 376, 1749 - 024 Lisboa, Portugal &amp; MARE - Centro de Ciências do Mar e do Ambiente (FCT / NOVA), Campus de Caparica, Caparica, Portugal</mods:affiliation>
</mods:name>
<mods:name id="7539E0C90F9E443C6489573C4B5433FB" type="personal">
<mods:role id="DF8197FF5645CAD042F41471022292D7">
<mods:roleTerm id="9ADCBC52AF95776903CCF5440270567E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AF96BFA955E823E001F6C847101D651B">Noreña, Carolina</mods:namePart>
<mods:nameIdentifier id="379D237E51F4AD48AC39C87DE0C1FAD9" type="ORCID">0000-0002-4580-7460</mods:nameIdentifier>
<mods:affiliation id="A600C791D71D82DA216B0307963C8528">Departamento de Biodiversidad and Biología Evolutiva, Museo Nacional de Ciencias Naturales (CSIC), C / Jose Gutiérrez Abascal 2, Ap. 28006, Madrid, Spain</mods:affiliation>
</mods:name>
<mods:name id="5E850A16F02D10C36A4975392F5348CA" type="personal">
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<mods:roleTerm id="B5DC11BFCAABF00BB97B7467A414F202">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="236C5CC0AD461014CC93A64D953088A3">Cervera, Juan Lucas</mods:namePart>
<mods:nameIdentifier id="6DD5AC3BBED9DCA7A37456B4C50274FB" type="ORCID">0000-0002-8337-2867</mods:nameIdentifier>
<mods:affiliation id="CA29E6604B63B463494BDE2BF426ED20">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales, Campus de Excelencia Internacional del Mar (CEI MAR), Av. República Saharahui, s / n, Ap. 40. 11510, Puerto Real (Cádiz), Spain &amp; Instituto Universitario de Investigación Marina (INMAR), Campus de Excelencia Internacional del Mar (CEI · MAR), Universidad de Cádiz, Av. República Saharahui, s / n, Ap. 40. 11510, Puerto Real (Cádiz), Spain</mods:affiliation>
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<mods:typeOfResource id="970B09EAF4B52A51334A01AFE4BD7826">text</mods:typeOfResource>
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<mods:titleInfo id="D2AA5D783CD9DF73B2E883114D42D2C3">
<mods:title id="5BEEC3A7D1D3448240BE53E5AA2CDF82">Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part id="5EF24AF325E3A24BE3CBAC895F3C4657">
<mods:date id="8041BDDEBF3654212D7D18BA9FA2ED58">2024</mods:date>
<mods:detail id="9D9951EED915F50B982E26748B2762B9" type="pubDate">
<mods:number id="9836D28550070CD15F9E97B4ED0D4BBA">2024-11-07</mods:number>
</mods:detail>
<mods:detail id="11037A1A56D3AB4E9A11F6952FD82672" type="volume">
<mods:number id="A5C186A38A0EA723C2F0B31DA0537078">100</mods:number>
</mods:detail>
<mods:detail id="C60B46AC7463CD0A1B25760E8571FB27" type="issue">
<mods:number id="C2452CA73A7562CEB0A2B771D4820971">4</mods:number>
</mods:detail>
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<mods:start id="69AB6061A86E6148FA9B2EFCEC157240">1487</mods:start>
<mods:end id="5AC035ECDD0D7335AFF62391FA76A1F9">1513</mods:end>
</mods:extent>
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<mods:classification id="06B5DF53B9D1357A064EFBB42E52B40E">journal article</mods:classification>
<mods:identifier id="1EB01BD615923FB538F900C3B7BCCF22" type="DOI">10.3897/zse.100.128211</mods:identifier>
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<treatment id="4939B452948250D989766873C18F1310" ID-DOI="http://doi.org/10.5281/zenodo.14052243" ID-Zenodo-Dep="14052243" ID-arpha="4939B452-9482-50D9-8976-6873C18F1310" ID-zoobank="https://zoobank.org/9D19D6CC-7155-41D6-B6DB-39EE6E52F479" LSID="urn:lsid:plazi:treatment:4939B452948250D989766873C18F1310" LSID-ZBK="urn:lsid:zoobank.org:act:9D19D6CC-7155-41D6-B6DB-39EE6E52F479" httpUri="http://treatment.plazi.org/id/4939B452948250D989766873C18F1310">
<subSubSection id="D342AAE4915A6B31BC5EB73AA9588C76" type="nomenclature">
<paragraph id="C737C1E327DD8A0BA3F52DC47EF09284">
<taxonomicName id="D2FA37D1CE6AB39D042113C498340722" authorityName="Pérez-García &amp; Gouveia &amp; Calado &amp; Noreña &amp; Cervera" authorityYear="2024" family="Stylochidae" genus="Stylochus" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="erytheius" status="sp. nov.">
<emphasis id="3D9FBF7C524626CE12763D2804A72C5C" italics="true">Stylochus erytheius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="16A2766BDD87D7A867279BE6E066B561" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0DD76380E0E409CBE13B32DE89B71860" type="description">
<paragraph id="3E6010AAE49AC1AEE64FCAA59EAF0B8C">
<figureCitation id="7C264233C9619B92B475B745BCD8CBE4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stylochus erytheius sp. nov. (MNCN 4.01 / 3059 to 3113 and 4336 to 4535). Histological sections are antero-posteriorly oriented: A. Dorsal view of a living specimen; B. Ventral view; C. Detail of the tentacles and cerebral eyes; D. Histological section of the seminal vesicle; E. Histological section of the male and female copulatory organs; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; eg, extravesicular glands; ev, external vagina; fp, female pore; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; pp, penis papilla; pv, prostatic vesicle; sb, spermiducal bulbs; sv, seminal vesicle." figureDoi="10.3897/zse.100.128211.figure4" httpUri="https://binary.pensoft.net/fig/1169817">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGTAG" type="Holotype">
<paragraph id="6A2371C4A6A599DF97E93E1ED0912252">
<heading id="36664A04D694634560DB98E044E1924D" reason="title">
<typeStatus id="180342653A26370A89679BC838F02E98" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="841D59929E890DC5C5F753FF214DFB7D">
<materialsCitation id="BEDB9FA7A37720CF33708C6D81A46E27" collectingDate="2014-10-09" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1" typeStatus="holotype">
<specimenCode id="AB859D29D1C17E094240D5162981845D" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="9261C66E4A9C4E973FEE5DB76F8F327F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3059 to 3113 and 4336 to 4535, Station 11,
<date id="67E4EACE14B3563772A01934E7716300" value="2014-10-09">
<collectingDate id="78132FC2214F3237B1F0F81C043F818F" value="2014-10-09">9 October 2014</collectingDate>
</date>
,
<quantity id="67853C20C21A73CED2CDA80AB45B4244" metricMagnitude="-2" metricUnit="m" metricValue="7.1" unit="mm" value="71.0">71 mm</quantity>
long, sagittal sectioned into 254 slides.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQTAG" type="material">
<paragraph id="3DD212F6E907864E4054D66BA10A7DBA">
<heading id="4DB028E6D7494F842F2D5465B48D4048" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="652AE7DF24B914A21BFBD91DD7947346">
<materialsCitation id="16379AB287A446AC42F1146B612F0A84" collectingDate="2015-04-20" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="F840699B46566F445FD3A96927B53D5B" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="6C2EA335083EAA59217F802CFD044406" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3962, Station 11,
<date id="DAD97AC088E3A7AF45111493A3FB6642" value="2015-04-20">
<collectingDate id="20D43F5C90EDBBC657EE5744A3A875B2" value="2015-04-20">20 April 2015</collectingDate>
</date>
,
<quantity id="4D03F81992A86E7AF30D758FECA89827" metricMagnitude="-2" metricUnit="m" metricValue="3.5" unit="mm" value="35.0">35 mm</quantity>
long
</materialsCitation>
; •
<materialsCitation id="C1292138983CA3DCC2F89E4373A9A7C8" collectingDate="2015-04-20" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="A1F400B9FDEB498269944C772AACA3E2" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="2ABCBE4AB4D7A3955D6271C101034BD6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3964, Station 11,
<date id="5FE40462824C382E4681B4FD507A8B4A" value="2015-04-20">
<collectingDate id="927E70A0BDDBB070269263D5FA2F9E7B" value="2015-04-20">20 April 2015</collectingDate>
</date>
,
<quantity id="3F86D352D5985DF4F881D2D803C5A131" metricMagnitude="-2" metricUnit="m" metricValue="3.7" unit="mm" value="37.0">37 mm</quantity>
long
</materialsCitation>
; •
<materialsCitation id="2F802303AF3DF09EEF7171713B80CE7A" collectingDate="2018-02-02" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="FC0571FDD1BD624353FEC30D94FF77BD" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="DDF255891DF6452DB15F4C4C360EA704" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3966, Station 11,
<date id="7419A5C381E389583EBC1743DE294F47" value="2018-02-02">
<collectingDate id="2F2BC9ECAF77AA334DF812D658D756E7" value="2018-02-02">
2 February
<quantity id="9C1102E2BF08FD1F1F014F1A0A0050A3" metricMagnitude="3" metricUnit="m" metricValue="2.01812" unit="mm" value="2018120.0">2018, 120 mm</quantity>
</collectingDate>
</date>
long
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECUAG" type="diagnosis">
<paragraph id="6CDF08A5B1E23C30B691D7A3C5076757">
<heading id="23CC6D25F960B8AF156C398B3E86A308" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B852394DA5F49A09625DBD02CDCABC4B">Body shape is oval with rounded anterior and posterior ends. Orange-brown dorsal background; numerous dark brown and whitish dots scattered on the dorsal surface; visible line of white dots along the main body axis; ventral surface pale orange; gonopores well-separated; with spermiducal bulbs; opening of the sperm duct distally located; penis papilla short.</paragraph>
</subSubSection>
<subSubSection id="SECID0EHUAG" type="description">
<paragraph id="44E213CD702D2A3BD56C9D72B7B58C93">
<heading id="8B3E4A2CD70E7CCDC990E8A63E5BA288" reason="title">Description.</heading>
</paragraph>
<paragraph id="F60208E5BA205DC576492693A77699A7">
Oval-shaped
<taxonomicName id="1A50EB087D4196F55C95F49400DA3684" authorityName="Stimpson" authorityYear="1857" family="Stylochidae" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="family">Stylochidae</taxonomicName>
, with rounded anterior and posterior ends, fleshy,
<quantity id="33654A26BEB8AA9B3FA5755D6CABEF49" metricMagnitude="-2" metricUnit="m" metricValue="7.1" unit="mm" value="71.0">71 mm</quantity>
in length (65.75 ±
<quantity id="FBEB29BD06CF317A32A36CA45FD99B6D" metricMagnitude="-2" metricUnit="m" metricValue="3.976" unit="mm" value="39.76">39.76 mm</quantity>
) and
<quantity id="12A67CF3C034A74232423BF88DE9F7AC" metricMagnitude="-2" metricUnit="m" metricValue="6.3" unit="mm" value="63.0">63 mm</quantity>
in width. Margins mostly smooth. Background colouration orange-brown, with multiple dark brown and white dots on the dorsal surface (as shown in Fig.
<figureCitation id="A3A201600011C16377CBE78FE0EC2324" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stylochus erytheius sp. nov. (MNCN 4.01 / 3059 to 3113 and 4336 to 4535). Histological sections are antero-posteriorly oriented: A. Dorsal view of a living specimen; B. Ventral view; C. Detail of the tentacles and cerebral eyes; D. Histological section of the seminal vesicle; E. Histological section of the male and female copulatory organs; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; eg, extravesicular glands; ev, external vagina; fp, female pore; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; pp, penis papilla; pv, prostatic vesicle; sb, spermiducal bulbs; sv, seminal vesicle." figureDoi="10.3897/zse.100.128211.figure4" httpUri="https://binary.pensoft.net/fig/1169817">4 A</figureCitation>
). Additionally, a conspicuous line of white dots extends along the central axis of the body. Ventrally pale orange (Fig.
<figureCitation id="68B623F5F585AC194C872666C8F42095" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stylochus erytheius sp. nov. (MNCN 4.01 / 3059 to 3113 and 4336 to 4535). Histological sections are antero-posteriorly oriented: A. Dorsal view of a living specimen; B. Ventral view; C. Detail of the tentacles and cerebral eyes; D. Histological section of the seminal vesicle; E. Histological section of the male and female copulatory organs; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; eg, extravesicular glands; ev, external vagina; fp, female pore; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; pp, penis papilla; pv, prostatic vesicle; sb, spermiducal bulbs; sv, seminal vesicle." figureDoi="10.3897/zse.100.128211.figure4" httpUri="https://binary.pensoft.net/fig/1169817">4 B</figureCitation>
). Nuchal conical tentacles transparent, with tentacular eyes at the base and tips (Fig.
<figureCitation id="75A78D4B9179A1A5D59D044DF7471ADF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stylochus erytheius sp. nov. (MNCN 4.01 / 3059 to 3113 and 4336 to 4535). Histological sections are antero-posteriorly oriented: A. Dorsal view of a living specimen; B. Ventral view; C. Detail of the tentacles and cerebral eyes; D. Histological section of the seminal vesicle; E. Histological section of the male and female copulatory organs; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; eg, extravesicular glands; ev, external vagina; fp, female pore; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; pp, penis papilla; pv, prostatic vesicle; sb, spermiducal bulbs; sv, seminal vesicle." figureDoi="10.3897/zse.100.128211.figure4" httpUri="https://binary.pensoft.net/fig/1169817">4 C</figureCitation>
). Poorly visible cerebral eyes in two scattered clusters between the tentacles. Marginal eyes along the entire body margin. Pharynx ruffled centrally. Oral pore at the centre of the pharynx. Reddish, conspicuous gonopores well-separated (
<quantity id="B5CA1CA53F102D7B796113BE9364AC55" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.6 mm</quantity>
) posteriorly located.
</paragraph>
<caption id="BBBE652519C3847E37B3557E097E2E45" ID-DOI="10.3897/zse.100.128211.figure4" ID-arpha="9D26D7EF-AB67-59CD-9F70-66E5EB9902A2" httpUri="https://binary.pensoft.net/fig/1169817" startId="F4">
<paragraph id="3D64B94C5F3F2CB49B04AC62A2E3A6B1">
<label id="09E463CC2292109819E65A704CAE4D84">Figure 4.</label>
</paragraph>
<paragraph id="D0D5D9A19BBE0EA05D61F38E3745DC76">
<taxonomicName id="6C9232864DCDF777B9626E36B6AB26EC" authorityName="Pérez-García &amp; Gouveia &amp; Calado &amp; Noreña &amp; Cervera" authorityYear="2024" family="Stylochidae" genus="Stylochus" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="erytheius" status="sp. nov.">
<emphasis id="B8CD73E76FC0C55BCAA78F03BD3301E8" italics="true">Stylochus erytheius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B42F627557F3D95DA14F857C15386C4C" rank="species">sp. nov.</taxonomicNameLabel>
(
<named-content id="CBAF36DCF16AA5F6C2B139816360F0E0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01 / 3059 to 3113 and 4336 to 4535). Histological sections are antero-posteriorly oriented:
<emphasis id="88FA1518D8597A989D0FD4C81702B135" bold="true">A.</emphasis>
Dorsal view of a living specimen;
<emphasis id="F10BE0795AE628D4FD884999A09C1DC3" bold="true">B.</emphasis>
Ventral view;
<emphasis id="7DAC866776B4BDC3AFA9F50B99F14236" bold="true">C.</emphasis>
Detail of the tentacles and cerebral eyes;
<emphasis id="257C76B6EC5C540E80F6F44EBCF26B81" bold="true">D.</emphasis>
Histological section of the seminal vesicle;
<emphasis id="FE8462A1EB304274D25D8D1B949CD2C0" bold="true">E.</emphasis>
Histological section of the male and female copulatory organs;
<emphasis id="30BFA9AF84B9176A9A3A75CE81F8AABD" bold="true">F.</emphasis>
Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; eg, extravesicular glands; ev, external vagina; fp, female pore; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; pp, penis papilla; pv, prostatic vesicle; sb, spermiducal bulbs; sv, seminal vesicle.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EGWAG" type="Reproductive system">
<paragraph id="78F8FB72B97CFA4AA5FE08018D0F9506">
<heading id="28858C83E198FD685B65FBC5CAA7FDC3" reason="title">Reproductive system.</heading>
</paragraph>
<paragraph id="155EDC07C84EC4867DB977D12FB44F72">
Male copulatory apparatus comprises spermiducal bulbs, a tripartite seminal vesicle, and a free prostatic vesicle (Fig.
<figureCitation id="29B99CB35E7DA2DADF78F133FB1FB60E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stylochus erytheius sp. nov. (MNCN 4.01 / 3059 to 3113 and 4336 to 4535). Histological sections are antero-posteriorly oriented: A. Dorsal view of a living specimen; B. Ventral view; C. Detail of the tentacles and cerebral eyes; D. Histological section of the seminal vesicle; E. Histological section of the male and female copulatory organs; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; eg, extravesicular glands; ev, external vagina; fp, female pore; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; pp, penis papilla; pv, prostatic vesicle; sb, spermiducal bulbs; sv, seminal vesicle." figureDoi="10.3897/zse.100.128211.figure4" httpUri="https://binary.pensoft.net/fig/1169817">4 F</figureCitation>
). Spermiducal bulbs broad, run ventrally, and then posteriorly, turning dorsally before entering the seminal vesicle. Tripartite seminal vesicle anchor-like shape (Fig.
<figureCitation id="8E095068F531C32B92F760C4244DEA19" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stylochus erytheius sp. nov. (MNCN 4.01 / 3059 to 3113 and 4336 to 4535). Histological sections are antero-posteriorly oriented: A. Dorsal view of a living specimen; B. Ventral view; C. Detail of the tentacles and cerebral eyes; D. Histological section of the seminal vesicle; E. Histological section of the male and female copulatory organs; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; eg, extravesicular glands; ev, external vagina; fp, female pore; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; pp, penis papilla; pv, prostatic vesicle; sb, spermiducal bulbs; sv, seminal vesicle." figureDoi="10.3897/zse.100.128211.figure4" httpUri="https://binary.pensoft.net/fig/1169817">4 D, F</figureCitation>
), well developed. Prostatic vesicle, large and oval-shaped (
<quantity id="670999B86B8B07FF0C3C2574A10C0C18" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
long), belongs to the polyglandular (“
<emphasis id="13DDFE420AC4DCB8CAF00C862AB9636B" italics="true">djiboutiensis</emphasis>
”)
<typeStatus id="8C116F82B9656EAC85CF12610243B05A">type</typeStatus>
(
<bibRefCitation id="FB858D7AFB65FF8F82AD4D1FEF66B243" author="Faubel" firstAuthor="Faubel" journalOrPublisher="Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut" pagination="17-121" refId="B15" refString="Faubel A (1983) The Polycladida, Turbellaria. Proposal and stablishment of a new system. Part I. The Acotylea. Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut 80: 17121." title="The Polycladida, Turbellaria. Proposal and stablishment of a new system. Part I. The Acotylea." volume="80" year="1983">Faubel 1983</bibRefCitation>
). Muscular wall crossed by numerous extravesicular glands. The prostatic duct and the seminal duct join to form the short ejaculatory duct just before entering the penis papilla. Penis papilla elongated (
<quantity id="49DCB16131B05F90C5438C7C5879F49E" metricMagnitude="-4" metricUnit="m" metricValue="2.1" unit="mm" value="0.21">0.21 mm</quantity>
), opening into a sinuous male atrium (Fig.
<figureCitation id="DC1BB58E6B191CE288BF66F9A0D1668E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stylochus erytheius sp. nov. (MNCN 4.01 / 3059 to 3113 and 4336 to 4535). Histological sections are antero-posteriorly oriented: A. Dorsal view of a living specimen; B. Ventral view; C. Detail of the tentacles and cerebral eyes; D. Histological section of the seminal vesicle; E. Histological section of the male and female copulatory organs; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; eg, extravesicular glands; ev, external vagina; fp, female pore; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; pp, penis papilla; pv, prostatic vesicle; sb, spermiducal bulbs; sv, seminal vesicle." figureDoi="10.3897/zse.100.128211.figure4" httpUri="https://binary.pensoft.net/fig/1169817">4 E</figureCitation>
).
</paragraph>
<paragraph id="8BCE8EC505FFA205A2B6C3427483A7B5">The female copulatory apparatus simple and inconspicuous, formed by the vagina externa and interna. Without Langs vesicle. The oviducts open together into the posterior region of the vagina interna. Abundant shell and cement glands open into the vagina externa and distal region of the vagina interna.</paragraph>
</subSubSection>
<subSubSection id="SECID0E6WAG" type="type locality">
<paragraph id="3E6D38DECE6A7A8E9B87CA653174F5CE">
<heading id="E60332D463E00CE8731A56087CE49D24" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="EFE0D6CDEC18EE34B8A616A2075ED318">
La Caleta Beach, Cadiz,
<collectingCountry id="B0F4118BCEBEE558C720B20D92BAF2DA" name="Spain">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEXAG" type="etymology">
<paragraph id="4AFBCEBF814E51826CCD60E823E3F090">
<heading id="AB697A42E9DFD57629C25FA38E055BA9" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="55F2C2E22491C81F104D03CAB75CDBFD">
Named after Erytheia, a small island where the Phoenicians settled approximately 1100 BC. This area is where the beach of La Caleta is located, the
<typeStatus id="E3BFF70B1DD2449241B9FAF9AEAFD40D">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJXAG" type="remarks">
<paragraph id="4C83389E0DA6D7370138CC13C520E9E1">
<heading id="2102002EF14CC71FEE5E36DFBC631705" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="15E4B7F83F49CB3AAA5BFEC6C65FE394">
A discussion of this species is included together with
<taxonomicName id="20223F4B1C5875848839405A216E2FA3" authorityName="Pérez-García &amp; Gouveia &amp; Calado &amp; Noreña &amp; Cervera" authorityYear="2024" family="Stylochidae" genus="Stylochus" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="marimarensis" status="sp. nov.">
<emphasis id="671D6079D65B7F33D55C5AD40BC2FCFB" italics="true">S. marimarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3BFF4A01BAD63B7592F5E6E135A0361A" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1XAG" type="biology">
<paragraph id="F5FB67787A181A291300AD86D69C9729">
<heading id="D8ACF7DA635616A135D37369890EF397" reason="title">Biology.</heading>
</paragraph>
<paragraph id="29660DCC31372582F32E73E4516DCA9A">Specimens were collected from rocky substrates under stones in the intertidal zone.</paragraph>
</subSubSection>
<subSubSection id="SECID0E6XAG" type="distribution">
<paragraph id="EF7C903049B3D2BF7F5F7F7F1F3763D0">
<heading id="2590B830404C8C6C26D87C7C66B37E83" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="F4A327906A701B69EA975F9F3CEAD8A8">
This species has only been found in La Caleta, Cadiz,
<collectingCountry id="21342F4FBBBC9D9045EE931F11C94C49" name="Spain">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,120 @@
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<mods:title id="0FA0C53957E07206F76158DF6520862B">A review of the millipede genus Pseudodesmus Pocock, 1887 (Diplopoda, Platydesmida, Andrognathidae) from Vietnam, with descriptions of five new species and notes on its phylogeny</mods:title>
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<mods:affiliation id="78484C9EDC86BF7A0A4D3348946C71ED">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="F43F341B7678BDD70252D8D1AB0E5CA4">Nguyen, Lien T. P.</mods:namePart>
<mods:affiliation id="4E0B7349235B62C7215825B3FB88A0E1">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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<mods:affiliation id="3735244D97D97CA69E4E27ED96D5640A">University of Veterinary Medicine Budapest, István u. 2, 1078 Budapest, Hungary</mods:affiliation>
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<subSubSection id="3E33195F20AE7E4D3B4212625D094939" type="nomenclature">
<paragraph id="C57091E8FD00CE79DFF4491C784F8A66">
<heading id="C4DD5F332B8339369CC885AABC7CB5DC" reason="label">Genus</heading>
<taxonomicName id="AC3BC28B6D30F39601ABEA28BC41CDBE" authority="Pocock, 1887" authorityName="Pocock" authorityYear="1887" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Playtdesmida" phylum="Arthropoda" rank="genus">
<emphasis id="77FE556E96DBCC2EB72AE88D6C922B2F" italics="true">Pseudodesmus</emphasis>
<bibRefCitation id="EF90F8900BF374581E7A50FCEDD096CE" DOI="10.1080/00222938709460038" author="Pocock RI" pagination="1-222" refId="B28" refString="Pocock RI (1887) Description of a new genus and species of Polyzonidae. Annals and Magazine of Natural History ser. 5: 1 - 222. https: // doi. org / 10.1080 / 00222938709460038" title="Description of a new genus and species of Polyzonidae. Annals and Magazine of Natural History ser." volume="5" year="1887">Pocock, 1887</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELJAE" type="Type-species">
<paragraph id="DFE8ACE43EBBF9691AE710B711384A14">
<heading id="528D22A201394594F1B7157955B64C2F" reason="title">Type-species.</heading>
</paragraph>
<paragraph id="E9510FD0E9AE7A88B982E44267A89AB8">
<taxonomicName id="85A1D7149107A2DCC3A33030DDF0120C" authority="Pocock, 1887" authorityName="Pocock" authorityYear="1887" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="E5137D65FAD5D3A57001508E94BB5D25" italics="true">Pseudodesmus verrucosus</emphasis>
<bibRefCitation id="CD70285C805703C7022235E1793273A1" DOI="10.1080/00222938709460038" author="Pocock RI" pagination="1-222" refId="B28" refString="Pocock RI (1887) Description of a new genus and species of Polyzonidae. Annals and Magazine of Natural History ser. 5: 1 - 222. https: // doi. org / 10.1080 / 00222938709460038" title="Description of a new genus and species of Polyzonidae. Annals and Magazine of Natural History ser." volume="5" year="1887">Pocock, 1887</bibRefCitation>
</taxonomicName>
, by original designation. Known only from
<specimenCount id="765DB6CF900609B253C10E27A206F5C0" count="2" type="female">two female</specimenCount>
specimens from
<collectingRegion id="D53DB6B9A6A168E1968AD16D01FBC996" country="Malaysia" name="Perak">Perak State</collectingRegion>
, Peninsular
<collectingCountry id="6FF14AB7DEA6BFFD1F93E7D1845F5B5F" name="Malaysia">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3JAE" type="Historical diagnosis">
<paragraph id="C6E32F3BE07FB406A92F2DDE415E08BE">
<heading id="3A4D721F162AAC25BFA5A4F6D1198578" reason="title">Historical diagnosis.</heading>
</paragraph>
<paragraph id="35BAE22B7ACD0A4978701D786578F5D7">
Translated from Latin in
<bibRefCitation id="4C2A1B5815C6E153362FAD6C10E72C8F" DOI="10.1080/00222938709460038" author="Pocock RI" firstAuthor="Pocock" pagination="1-222" refId="B28" refString="Pocock RI (1887) Description of a new genus and species of Polyzonidae. Annals and Magazine of Natural History ser. 5: 1222. https://doi.org/10.1080/00222938709460038" title="Description of a new genus and species of Polyzonidae. Annals and Magazine of Natural History ser." volume="5" year="1887">Pocock (1887)</bibRefCitation>
:
<taxonomicName id="32ACCBCCBB47F5F2158B38F546581C0E" authorityName="Lucas" authorityYear="1843" class="Diplopoda" family="Platydesmidae" genus="Platydesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="genus">
<emphasis id="8A0A0288BBC36C8C6DC07A2AA79D61CA" italics="true">Platydesmus</emphasis>
</taxonomicName>
form. With a long body, bearing a series of tubercles above and keels below on each side. With a number of segments numbering greater than seventy; the last segment posteriorly not sharp; the segments, except for the first and last, bearing keels near the lower part of the side, rising close to level, and with a groove along the middle of the back. Each keel, except the first three, has a repugnatorial pore on the lateral margin. The first four segments have two legs each, the others are equipped with two pairs of legs; the last (and possibly the penultimate) lacking legs. The legs consist of six joints; the last joint of the leg is armed with a claw at the tip. The leg-bearing plates are free. The head is bent under the anterior segments, with a convex front; the anterior margin scarcely produced into a snout. Without eyes. Antennae consist of seven articles; located on the sides of the head; the last article being the smallest. The mandibles are hidden. The gnathochilarium is prominent; with large stipes; the jaws and cardines are not conspicuous; there are no lobes on the small, thin lamellae linguales; the mentum is hammer-shaped and large.
</paragraph>
<paragraph id="AD4137A2D928E0DC21F23605F6CB7DBB">
According to
<bibRefCitation id="00895618CD8E32C26A1C4440FE85D367" author="Wesener T" editor="Minelli A" firstAuthor="Wesener" journalOrPublisher="Vol. 2. Treatise on Zoology - Anatomy, Taxonomy, Biology. Brill, Leiden" pagination="382-482" refId="B41" refString="Wesener T (2015) Subterclass Colobognatha Brandt, 1834. In: Minelli A (Ed.) The Myriapoda. Vol. 2. Treatise on Zoology Anatomy, Taxonomy, Biology. Brill, Leiden, 382482." title="Subterclass Colobognatha Brandt, 1834." volumeTitle="The Myriapoda." year="2015">Wesener (2015)</bibRefCitation>
, the genus can be recognized by the following characters: “ Colour variable. Trunk wide, paraterga strongly elongated. Body length up to
<quantity id="992E3B4ED27037DF3597F2863A2C2B98" metricMagnitude="-2" metricUnit="m" metricValue="6.0" unit="mm" value="60.0">60 mm</quantity>
, with 6676 body rings. Collum with tubercles. Tergites densely pilose, strongly sculptured, with several smaller and one row of larger tubercles. Larger tubercles sometimes forming a ridge or resembling conic spines ”.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETKAE" type="remarks">
<paragraph id="83D1A90B4A52C45A8E4F25A9ACD1140B">
<heading id="E73FE23434B7863860701E6CB6453FAA" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="A9254A5CA6A0A7E3E4AE8716B29B713E">
The genus has been found only in Southeast Asia, but has been poorly studied. Since the description of its last species,
<taxonomicName id="80EA913DF6B5A613D179E72385673A72" authority="(Attems, 1953)" baseAuthorityName="Attems" baseAuthorityYear="1953" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="3FBD75582FD5F3FDA2877A4C65018962" italics="true">Pseudodesmus persimilis</emphasis>
(
<bibRefCitation id="0C6007A18554C44C737B032B6847BC7B" author="Attems C" journalOrPublisher=", série A" pagination="133-230" refId="B3" refString="Attems C (1953) Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939). Memoires du Museum national d'histoire naturelle, N. S., serie A 5: 133 - 230." title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938-1939). Mémoires du Muséum national dhistoire naturelle, N. S." volume="5" year="1953">Attems, 1953</bibRefCitation>
)
</taxonomicName>
, there has been very little taxonomic information about the genus.
</paragraph>
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<mods:title id="F17157B1A81D62D91D751386A7ED1FA7">A review of the millipede genus Pseudodesmus Pocock, 1887 (Diplopoda, Platydesmida, Andrognathidae) from Vietnam, with descriptions of five new species and notes on its phylogeny</mods:title>
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<mods:namePart id="53D0E905324E43E3E9D9F827A4565542">Nguyen, Anh D.</mods:namePart>
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<mods:affiliation id="799C3044BBA5183A21095E7B26CECE31">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="556B6927C98294F02A33AF3CC3E5DDA7">Nguyen, Lien T. P.</mods:namePart>
<mods:affiliation id="FEFA792EDF783525BCA1AA741AD21CD2">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="A8F75FECD952D4266ACFBAE998D800D4">Korsós, Zoltán</mods:namePart>
<mods:nameIdentifier id="85EE59DE46472352D35FC21D4D353961" type="ORCID">0000-0003-1545-5086</mods:nameIdentifier>
<mods:affiliation id="2A85A185A70D13DE37DD43A61D704F9D">University of Veterinary Medicine Budapest, István u. 2, 1078 Budapest, Hungary</mods:affiliation>
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<mods:title id="6CF20FBA80DB22D797385BEBD35AE380">Zoosystematics and Evolution</mods:title>
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<mods:part id="2C3B3603EBD701FA1A790EE9E58AB959">
<mods:date id="B0CEA79754F5A12B0DBB79F3651761FF">2024</mods:date>
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<mods:number id="7F6010373FDA08C84CB96F86272DB9A5">2024-11-07</mods:number>
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<mods:number id="95204F47C48EA17C3A7943222EF5D743">100</mods:number>
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<treatment id="5C8A7EFE68115280A5B44B8016067696" ID-DOI="http://doi.org/10.5281/zenodo.14052222" ID-Zenodo-Dep="14052222" ID-arpha="5C8A7EFE-6811-5280-A5B4-4B8016067696" ID-zoobank="https://zoobank.org/0E9D217B-0ED9-41E8-9712-960188EF700A" LSID="urn:lsid:zoobank.org:act:0E9D217B-0ED9-41E8-9712-960188EF700A" LSID-ZBK="urn:lsid:zoobank.org:act:0E9D217B-0ED9-41E8-9712-960188EF700A" httpUri="http://treatment.plazi.org/id/5C8A7EFE68115280A5B44B8016067696">
<subSubSection id="0852BC6145E5D9F9BDA6392832A7EF7A" type="nomenclature">
<paragraph id="8F89D3B26E1A374791CD95F734720D25">
<taxonomicName id="4DCA19684F5B0BD95B660E938FE22A94" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Playtdesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
<emphasis id="BF3E3B2C978BE53A1F360B565816A309" italics="true">Pseudodesmus bidoup</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4ABEA99E37B28E7BE1227EA726E20FEE" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0B288E7FFB60CA4D63B21EEF0507DC07" type="description">
<paragraph id="503A5A45FE5DEA6BEC79F6028E03F06A">
<figureCitation id="32C2123FCF5F27D8A5CBC4076E58D517" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Habitus, dorsal view; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure2" httpUri="https://binary.pensoft.net/fig/1169892">Figs 2</figureCitation>
,
<figureCitation id="A86CA0D39D47DF0BC43F06E1F2945844" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Collum and body rings 26, dorsal view; B. Midbody body rings, dorsal view; C. Anterior part of body, ventral view; D. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure3" httpUri="https://binary.pensoft.net/fig/1169893">3</figureCitation>
,
<figureCitation id="B506DD3FCA099BB88A5FF9EA01F66D8D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957). Gonopods. A. Right anterior gonopods, anterior view; B. Right posterior gonopod, distal part, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure4" httpUri="https://binary.pensoft.net/fig/1169894">4</figureCitation>
,
<figureCitation id="8929E499587A1E0A496C6D33304D88F2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957). Gonopods. SEM. A, B. Right anterior gonopods, ventral view (A), anterior view (B); C, D. Right posterior gonopod, ventral view (C), sublateral view (D). Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure5" httpUri="https://binary.pensoft.net/fig/1169895">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EASAE" type="type material">
<paragraph id="6D9B3AEBCFE5E722B7BECAEEC2B4F9B4">
<heading id="6FD8B2956ACD4D0094891DBA03F95AE8" reason="title">Type material.</heading>
</paragraph>
<paragraph id="0FCDD1B602648CD0EEF163B16BE9DED9">
<materialsCitation id="0779C929638E30FFB917AC7BC7DADA0D" collectingDate="2009-04-29" collectingDateMax="2009-05-09" collectingDateMin="2009-04-29" collectorName="Anh D. Nguyen" country="Vietnam" elevation="1650" latitude="12.11231" location="Bi Doup - Nui Ba National Park" longLatPrecision="5" longitude="108.6627" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="holotype">
<emphasis id="4DC4A6C0221FCB4E4BB1628B1FD99873" bold="true">
<emphasis id="E2EA48CB02D628794D8B07EE02405C43" bold="true" italics="true">
<typeStatus id="7747B35894A2B442E7A8491B5EE0C6E2" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="407C43123093202023C235218A10660A" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="B1E80F925BD6F54EEB8904A508AAFB0E" count="1" type="male">1 male</specimenCount>
;
<collectingRegion id="FC693A4F691CF5192F9739F38E18B91C" country="Vietnam" name="Lam Dong">Lam Dong Province</collectingRegion>
,
<location id="D75E8E3BD1702BC4EC6242A65F1C576D" LSID="urn:lsid:plazi:treatment:5C8A7EFE68115280A5B44B8016067696:D75E8E3BD1702BC4EC6242A65F1C576D" country="Vietnam" latitude="12.11231" longLatPrecision="5" longitude="108.6627" name="Bi Doup - Nui Ba National Park" stateProvince="Lam Dong">Bi Doup Nui Ba National Park</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOSAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[108.662700,12.112310]}">
<geoCoordinate id="AF13978EB5B46814D6942DB2217DE0E6" degrees="12.11231" direction="north" orientation="latitude" precision="1" value="12.11231">12.11231 ° N</geoCoordinate>
,
<geoCoordinate id="68736079E0A6FC8D7A783AA54204C581" degrees="108.6627" direction="east" orientation="longitude" precision="5" value="108.6627">108.6627 ° E</geoCoordinate>
</named-content>
;
<elevation id="32B3F80BFB20AC5D596DFE802D9ED655" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">
<quantity id="004B8A33A9F04017F2F04FD19D420154" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">1,5001,800 m</quantity>
a. s. l.
</elevation>
;
<date id="3AE9FA7F28B8F369808B25FC3E3A2BB1" value="2009-04-29" valueMax="2009-05-09" valueMin="2009-04-29">
<collectingDate id="27A6AD5F2FCE22C03E11BC1D8B3099EA" value="2009-04-29" valueMax="2009-05-09" valueMin="2009-04-29">29 April 9 May 2009</collectingDate>
</date>
;
<collectorName id="8C025AF3216149AB91BBBBC223ADD4D6">Anh D. Nguyen</collectorName>
leg; mixed forest;
<abbrev content-type="institution" id="ABBRID0ETSAE" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
957 H.
</materialsCitation>
</paragraph>
<paragraph id="091ED146B4DAA19CB9CD153EC8DC3E5E">
<materialsCitation id="6DD4881DE0F68B861E188E94845E8AFA" collectingDate="2009-04-29" collectingDateMax="2009-05-09" collectingDateMin="2009-04-29" collectorName="Anh D. Nguyen" country="Vietnam" elevation="1650" latitude="12.11231" location="Bi Doup - Nui Ba National Park" longLatPrecision="5" longitude="108.6627" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" stateProvince="Lam Dong" typeStatus="paratype">
<emphasis id="8B341C0BB34D288E6FE42AB339A0C252" bold="true">
<emphasis id="374B08E4B855FF619836D35F49EDB77E" bold="true" italics="true">
<typeStatus id="91114DDAEBB1A2CF36A9EFBE82173FED" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="CBD702C50A0CF3AF34E997E7DD38114E" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="B14E14E87057AA79FD3DCA07C86C3F55" count="1" type="female">1 female</specimenCount>
,
<specimenCount id="3850A13C64D18AD338BA0BCADBF078E9" count="1" type="juvenile">1 juvenile</specimenCount>
; same data as for the holotype;
<abbrev content-type="institution" id="ABBRID0E6SAE" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
957 P
</materialsCitation>
<materialsCitation id="E80C6DDE60F557699CC5A480A385BEF0" collectingDate="2009-04-29" collectingDateMax="2009-05-09" collectingDateMin="2009-04-29" collectionCode="HNHM" collectorName="Anh D. Nguyen" country="Vietnam" elevation="1650" latitude="12.11231" location="Bi Doup - Nui Ba National Park" longLatPrecision="5" longitude="108.6627" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCount id="ECDAA8A9993E5D7E379D4A399CE9CDE8" count="1" type="female">1 female</specimenCount>
; same data as for the holotype;
<named-content id="82D807876577991FAE4FF5697B00B641" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum-termeszettudomanyi-muzeum" xlink_title="Department of Zoology, Hungarian Natural History Museum, Budapest, Hungary">
<collectionCode id="09C0FBADA74AEDF9D8A8A4F3991572D6" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
</named-content>
</materialsCitation>
<materialsCitation id="0A82184C56C7FF5886E6859607E3F358" collectingDate="2004-04-15" collectingDateMax="2004-04-19" collectingDateMin="2004-04-15" collectorName="Anh D. Nguyen" country="Vietnam" elevation="830" latitude="15.23439" location="Lo Xo" longLatPrecision="1" longitude="107.73386" specimenCount="1" specimenCount-male="1" stateProvince="Kon Tum" typeStatus="paratype">
<specimenCount id="09E9A4933FA6A57E90DA1E5332DDE449" count="1" type="male">1 male</specimenCount>
(41 body rings);
<collectingRegion id="05BB6EF1B9AFEF11F08D1E2D88347961" country="Vietnam" name="Kon Tum">Kon Tum Province</collectingRegion>
,
<location id="6219B20990FB8189B7E631E5471DC9AC" LSID="urn:lsid:plazi:treatment:5C8A7EFE68115280A5B44B8016067696:6219B20990FB8189B7E631E5471DC9AC" country="Vietnam" latitude="15.23439" longLatPrecision="1" longitude="107.73386" name="Lo Xo" stateProvince="Kon Tum">Lo Xo</location>
pass;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMTAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.733860,15.234390]}">
<geoCoordinate id="F1AF6F34B9AE5CD95F18C01F1F6373BA" degrees="15.23439" direction="north" orientation="latitude" precision="1" value="15.23439">15.23439 ° N</geoCoordinate>
,
<geoCoordinate id="002CCE5CCDC8CF626B4F06E78E403882" degrees="107.73386" direction="east" orientation="longitude" precision="1" value="107.73386">107.73386 ° E</geoCoordinate>
</named-content>
;
<elevation id="CFCE161F5B4532AC9B182EFDEBE4844E" metricMagnitude="2" metricUnit="m" metricValue="8.3" unit="m" value="830.0">
<quantity id="1800A0947DAB389503B35F1AC1F85E07" metricMagnitude="2" metricUnit="m" metricValue="8.3" unit="m" value="830.0">830 m</quantity>
a. s. l.
</elevation>
;
<date id="AA6A9F46E6004429D0E475004F804462" value="2004-04-15" valueMax="2004-04-19" valueMin="2004-04-15">
<collectingDate id="4AB2682783C3817CCA0D215EAEC2A893" value="2004-04-15" valueMax="2004-04-19" valueMin="2004-04-15">1519 April 2004</collectingDate>
</date>
;
<collectorName id="2058E5FF1BBB9F8CCA550B2C1DB74057">Anh D. Nguyen</collectorName>
leg.; secondary forest;
<collectingMethod id="E6B76454884ED6BED93932D5A9AFF967">pitfall trap</collectingMethod>
;
<abbrev content-type="institution" id="ABBRID0ERTAE" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
961
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWTAE" type="diagnosis">
<paragraph id="F84A10954A6757B395E1557A1DA70C53">
<heading id="F5E3EC82BBCE4B13DB897684DA2D3839" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="47A9F4C21D695C3D76985116B47D61A0">Head slightly smaller than collum. Collum with two rows of tubercles 1 + 1 (2) smaller and 2 + 2 larger tubercles. Body smooth, neither setose nor pubescent, midbody rings with two rows of tubercles on metazonites, anterior row longer, extending to about midlength of paraterga, containing 5 + 5 tubercles; posterior row shorter and extending to base of paraterga, consisting of 3 + 3 tubercles. Posterior gonopods 6 - segmented, distally carrying four apical stylets.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2TAE" type="diagnosis">
<paragraph id="6304023984E494106B434E1A51312822">
<heading id="FDF5725A0FEAE9DF1BA014BC04CAE3C3" reason="title">Diagnosis remark.</heading>
</paragraph>
<paragraph id="D90E147B9DC6F01F4C84122E37127135">
The new species is easily distinguished from
<taxonomicName id="CAF1FD3DCD43D82024EE59A39FCCE988" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="camptotrichus">
<emphasis id="EA3D3D74456D17286093578FC4D4EA97" italics="true">P. camptotrichus</emphasis>
</taxonomicName>
by coloration in alcohol (light yellowish vs earth brown). It also differs from two previously known species,
<taxonomicName id="BAC36F4248AD75CA19F510D7BC5A3DE6" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="338726729307A2B025BD8534EEF6284F" italics="true">P. variegatus</emphasis>
</taxonomicName>
and
<taxonomicName id="7F06B16E37D737085264C88922811B91" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="camptotrichus">
<emphasis id="EFF4739B1DCC5354C1D763E7D39ACB54" italics="true">P. camptotrichus</emphasis>
</taxonomicName>
in having smaller size (number of male body rings: 35 vs 5356 and 3742; length:
<quantity id="04D56A606903EF0A928200CA2B87BC64" metricMagnitude="-3" metricUnit="m" metricValue="9.25" unit="mm" value="9.25">9.25 mm</quantity>
vs
<quantity id="A96F9D0B54474C062B1C8E1A68828FE4" metricMagnitude="-2" metricUnit="m" metricValue="3.2" unit="mm" value="32.0">32 mm</quantity>
and
<quantity id="68928396650EFCFAA3D4FC04D7D4A7A5" metricMagnitude="-2" metricUnit="m" metricValue="1.4" unit="mm" value="14.0">14 mm</quantity>
; width:
<quantity id="2688037DAABC097636EEB5E66A34BB83" metricMagnitude="-3" metricUnit="m" metricValue="1.63" unit="mm" value="1.63">1.63 mm</quantity>
vs
<quantity id="B8D88F53C9DC9AA9C9F04B9E80D33380" metricMagnitude="-3" metricUnit="m" metricValue="5.8" unit="mm" value="5.8">5.8 mm</quantity>
and 2.0 mm, respectively), number of metazonal tubercles (two rows of 5 + 5 and 3 + 3 vs two rows of 815 + 8 15 and 57 + 5 7, and two rows of 6 + 6 to 9 + 9 each row, respectively).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECVAE" type="description">
<paragraph id="7E1C8F3952F9FB79111D4DBCC8CA2B11">
<heading id="05C11505BFD514EEC3E87A7C8E16A5DE" reason="title">Description.</heading>
</paragraph>
<paragraph id="2119EBA69F80272393012D9822AA5D9C">
<emphasis id="B7FBEDD9A91C18AD91FA131845EC6AFC" bold="true">
Male
<typeStatus id="62EAB1CD7EE72192976623974E06DFEE" type="holotype">holotype</typeStatus>
.
</emphasis>
</paragraph>
<paragraph id="3C2772EF5E974D5984C312B33217E2D7">
<emphasis id="C6ADEEDD4E0B2DAEFEEE00FC270EE8A1" bold="true" italics="true">Measurements</emphasis>
: 35 body rings plus telson; length about
<quantity id="A276B45A7E304FAE2A067C184E135E12" metricMagnitude="-3" metricUnit="m" metricValue="9.25" unit="mm" value="9.25">9.25 mm</quantity>
, length of metazona about
<quantity id="9B07BF5D93FB631EAFB7D584357C544D" metricMagnitude="-4" metricUnit="m" metricValue="1.9" unit="mm" value="0.19">0.19 mm</quantity>
, width of metazona about
<quantity id="E3EA332EF3FA6586ED16BED5060E4E8E" metricMagnitude="-3" metricUnit="m" metricValue="1.63" unit="mm" value="1.63">1.63 mm</quantity>
, width of prozona about
<quantity id="6EB6668B42AE7890A71FF50DE01EC422" metricMagnitude="-3" metricUnit="m" metricValue="1.25" unit="mm" value="1.25">1.25 mm</quantity>
; ratio of width of head and width of collum = 0.93.
</paragraph>
<paragraph id="21692D587CAFDB7D0169C261E8A846FF">Coloration of ethanol-preserved specimen: whole body including legs and antenna light yellowish.</paragraph>
<paragraph id="43C706DEEA6D0036CFE0F274973B737F">
Head almost smooth, not setose, round-shaped. Antenna stout, clavate,
<emphasis id="CFE68F002130DEC24002E40A0F470535" italics="true">in situ</emphasis>
extending to body ring 4; antennomere 6&gt; 5&gt; 3&gt; 4 = 2&gt; 7 =
<quantity id="0C71F7872196B937C7D9758FEC8453E4" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
length. Collum with distinct, but small paraterga directed anterolaterad (Figs
<figureCitation id="44DADC2BB1FC9D6E441A574E718EBD97" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Habitus, dorsal view; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure2" httpUri="https://binary.pensoft.net/fig/1169892">2 B</figureCitation>
,
<figureCitation id="8DAA85910531409D17C5457F8592BA9E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Collum and body rings 26, dorsal view; B. Midbody body rings, dorsal view; C. Anterior part of body, ventral view; D. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure3" httpUri="https://binary.pensoft.net/fig/1169893">3 A</figureCitation>
); two rows of tubercles on collum: anterior row with 1 + 1 small tubercles, posterior row with 2 + 2 large tubercles including larger paramedian ones (Fig.
<figureCitation id="854F97B47620021F499AAC25E19D2CDE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Collum and body rings 26, dorsal view; B. Midbody body rings, dorsal view; C. Anterior part of body, ventral view; D. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure3" httpUri="https://binary.pensoft.net/fig/1169893">3 A</figureCitation>
). Body smooth, neither setose nor pubescent. Metatergites elevated medially and slightly declined laterad, so body in cross-section subtriangular. Terga 24 with a row of 24 + 2 4 large tubercles including 2 + 2 tubercles on paraterga (Figs
<figureCitation id="00F3753022DEDBAE3CB5EDBE03E4A8FE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Habitus, dorsal view; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure2" httpUri="https://binary.pensoft.net/fig/1169892">2 B</figureCitation>
,
<figureCitation id="2CCDF3086E271E0AB0CBFA6AC4B014B5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Collum and body rings 26, dorsal view; B. Midbody body rings, dorsal view; C. Anterior part of body, ventral view; D. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure3" httpUri="https://binary.pensoft.net/fig/1169893">3 A</figureCitation>
); other terga with two rows of tubercles, anterior row longer, extending to about midlength of paraterga, containing 5 + 5 tubercles; posterior row shorter and extending to base of paraterga, consisting of 34 + 3 4 tubercles. Paramedian tubercles considerably larger (Figs
<figureCitation id="78887FAAB5FA247E7B57349DCDB0F291" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Habitus, dorsal view; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure2" httpUri="https://binary.pensoft.net/fig/1169892">2 A C</figureCitation>
,
<figureCitation id="6D9C446872721F720AC8D30148CC90EB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Collum and body rings 26, dorsal view; B. Midbody body rings, dorsal view; C. Anterior part of body, ventral view; D. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure3" httpUri="https://binary.pensoft.net/fig/1169893">3 A, B</figureCitation>
).
</paragraph>
<caption id="95809EBC6BCC68F02B15F7AE77A40AD6" ID-DOI="10.3897/zse.100.132111.figure2" ID-arpha="30BD1615-1DCD-5BB9-A2F8-62E62260D196" httpUri="https://binary.pensoft.net/fig/1169892" startId="F2">
<paragraph id="7E17F3E8C4DE2A891E9496C080B58A41">
<label id="C023DEBCFD0C252B8253C9B4828AFAB2">Figure 2.</label>
</paragraph>
<paragraph id="8CA75620A94E6EE8419FD330852C8E73">
<taxonomicName id="B052AF678DEA59931C1C7FFA5055C918" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
<emphasis id="A6312999B0B710B5EE0B6AA63CC71B9E" italics="true">Pseudodesmus bidoup</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94BA3F38C7FDF7D6D041E24C614C9480" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EFXAE" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
957), under UV light.
<emphasis id="377C8ACE3DD7C5B67581388A356EBE8B" bold="true">A.</emphasis>
Habitus, dorsal view;
<emphasis id="A5515D5611FDCCE191FE1B02D8F4EE14" bold="true">B.</emphasis>
Anterior part of body, dorsal view;
<emphasis id="4C68CD021972022731A90D6526CB99DA" bold="true">C.</emphasis>
Posterior part of body, dorsal view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="1FAC3405A3FBEB0FF8476C74C23E60F2" ID-DOI="10.3897/zse.100.132111.figure3" ID-arpha="F1CC91CD-441A-5CE7-AB2F-E49B64ABF823" httpUri="https://binary.pensoft.net/fig/1169893" startId="F3">
<paragraph id="090FBA1B1476751F842FB03DC348D745">
<label id="5503985909B81C61F15FB4455223B5C7">Figure 3.</label>
</paragraph>
<paragraph id="FDCBD8DB245DDE211391AB802D474D06">
<taxonomicName id="DAAC8076FB48434F56ECEE32E40BECAD" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
<emphasis id="C50BCCC0A85E2CFB6060474CA89EC7A9" italics="true">Pseudodesmus bidoup</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89E6D2A520A7CCFFACF6E28D55523C73" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EIYAE" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
957), under UV light.
<emphasis id="3E0117424CE0C55B45E03EC37B89BB26" bold="true">A.</emphasis>
Collum and body rings 26, dorsal view;
<emphasis id="3537F3BEE922CAEB0921C74E2652CCF8" bold="true">B.</emphasis>
Midbody body rings, dorsal view;
<emphasis id="D121E5AD1FC3118DADB8F2DCD67D984C" bold="true">C.</emphasis>
Anterior part of body, ventral view;
<emphasis id="A6188DF772561DA6C2B8D89A0CC13906" bold="true">D.</emphasis>
Posterior part of body, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="040A051BEC450937E75FEF2644BE0BCF">
Paraterga slightly curved anteriad on body rings 25, lateral margins rounded (Fig.
<figureCitation id="421307E88790284CF8B2BEEBAAEFB806" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Habitus, dorsal view; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure2" httpUri="https://binary.pensoft.net/fig/1169892">2 A, B</figureCitation>
); increasingly less so curved the following body rings, slightly curved caudad on body rings 2527, and strongly curved caudad on body rings 2831; caudal corners of paraterga increasingly acute on 67 posteriormost body rings in front of telson; paraterga of penultimate body ring produced strongly caudad and flanking telson (Figs
<figureCitation id="D3A43D0FC5630C17876F61487CBC0EFA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Habitus, dorsal view; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure2" httpUri="https://binary.pensoft.net/fig/1169892">2 A C</figureCitation>
,
<figureCitation id="53D20F87B9E4F411BD307EAF40F55A2E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Collum and body rings 26, dorsal view; B. Midbody body rings, dorsal view; C. Anterior part of body, ventral view; D. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure3" httpUri="https://binary.pensoft.net/fig/1169893">3 A, B</figureCitation>
). All margins of paraterga serrated, with tiny setae.
</paragraph>
<paragraph id="F1A0D072C99CB83E176BD58BEFEBE35F">
Telson (Figs
<figureCitation id="806AD9060F00260F7ED9397118251F6D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Habitus, dorsal view; B. Anterior part of body, dorsal view; C. Posterior part of body, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure2" httpUri="https://binary.pensoft.net/fig/1169892">2 C</figureCitation>
,
<figureCitation id="0E7CD890C13EFC20B89FFACD009877C0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Collum and body rings 26, dorsal view; B. Midbody body rings, dorsal view; C. Anterior part of body, ventral view; D. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure3" httpUri="https://binary.pensoft.net/fig/1169893">3 D</figureCitation>
) short, caudal margin rounded; epiproct with 2 + 2 small / tiny tubercles at caudal margin. Paraprocts and hypoproct semi-circular.
</paragraph>
<paragraph id="104EC47F492CE266D003F24269E2DBFD">
Legs slender, shorter than mid-metazonal width, terminating before lateral margins of paraterga. Prefemur = tarsus&gt; femur&gt; postfemur = tibia in length. Claws normal. Coxal sacs present from body rings 328 (Fig.
<figureCitation id="F5609688C8A03F2BC8DC2FB564F8F475" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Collum and body rings 26, dorsal view; B. Midbody body rings, dorsal view; C. Anterior part of body, ventral view; D. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure3" httpUri="https://binary.pensoft.net/fig/1169893">3 D</figureCitation>
).
</paragraph>
<paragraph id="0B16D70EDA9571844513BB1B095FF47D">
<emphasis id="85566AA56A10C829E4E17FDC64104E01" bold="true" italics="true">Gonopods</emphasis>
: Two pairs of gonopods directed anteromesad. Anterior gonopods (Figs
<figureCitation id="9EB4532799DF65744151253F3B1BF41D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957). Gonopods. A. Right anterior gonopods, anterior view; B. Right posterior gonopod, distal part, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure4" httpUri="https://binary.pensoft.net/fig/1169894">4 A</figureCitation>
,
<figureCitation id="052CB2334A4BEC5879E2AC4E48BA1762" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957). Gonopods. SEM. A, B. Right anterior gonopods, ventral view (A), anterior view (B); C, D. Right posterior gonopod, ventral view (C), sublateral view (D). Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure5" httpUri="https://binary.pensoft.net/fig/1169895">5 A, B</figureCitation>
) 6 - segmented, covered with long setae; coxite broad, basal part sparsely covered with microgranulations; podomere 3 and ultimate podomere (
<emphasis id="9728F3521DE94AA5DCF66767442A1238" italics="true">po 6</emphasis>
) longest; ultimate podomere with distal long, thick setae (
<emphasis id="E6874948C3E5F22FEDD4E38C132C9FA4" italics="true">mse</emphasis>
). Posterior gonopods (Figs
<figureCitation id="B2EECD9F45EADCEEBDFB42C93C2A9C48" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957), under UV light. A. Collum and body rings 26, dorsal view; B. Midbody body rings, dorsal view; C. Anterior part of body, ventral view; D. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure3" httpUri="https://binary.pensoft.net/fig/1169893">3 B</figureCitation>
,
<figureCitation id="15EBCE92A00C60D2E1BD3877C8A30490" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pseudodesmus bidoup sp. nov. Holotype (IEBR-Myr 957). Gonopods. A. Right anterior gonopods, anterior view; B. Right posterior gonopod, distal part, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure4" httpUri="https://binary.pensoft.net/fig/1169894">4 C, D</figureCitation>
) 6 - segmented, covered with sparse long setae; coxite broadly large, basal part sparsely covered with microgranulations; podomeres 25 short and stout, somewhat equal in length; ultimate podomere (
<emphasis id="453B7EB2063FFC4751B77AAE9C43119C" italics="true">po 6</emphasis>
) longest, with four apical stylets (
<emphasis id="059E33E7B455BFD9726BA4EE32845AF8" italics="true">sty</emphasis>
).
</paragraph>
<caption id="11B462CDF070AC2B6040485E64FBEC27" ID-DOI="10.3897/zse.100.132111.figure4" ID-arpha="6167BDF1-12BD-5E43-A3F5-5BA929FE8485" httpUri="https://binary.pensoft.net/fig/1169894" startId="F4">
<paragraph id="63059CC4252BB599CDF1356831878ABB">
<label id="EB8ED6018049F9E174CD8DB9DB878C94">Figure 4.</label>
</paragraph>
<paragraph id="54D3604EF9BAC81976A6E10C375B845E">
<taxonomicName id="B66C4F6978E4B5F308A0DD8E5C832CF8" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
<emphasis id="2C96ECAD2A438E5F6E86259E8584568D" italics="true">Pseudodesmus bidoup</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8624F09DEE8452F6708DD840E644975F" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EK2AE" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
957). Gonopods.
<emphasis id="63CE5E04ACE192727612C3FF80B8947A" bold="true">A.</emphasis>
Right anterior gonopods, anterior view;
<emphasis id="F45DF39CBC55D955D656EF36939C33D5" bold="true">B.</emphasis>
Right posterior gonopod, distal part, ventral view.
<emphasis id="5557B4CF30EA9C2D73636CB6AF4E421C" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="5FA040CC8E7AD6F7A6B5804170DB3740" ID-DOI="10.3897/zse.100.132111.figure5" ID-arpha="13BC5385-0088-5B63-9D3F-AC28515E5480" httpUri="https://binary.pensoft.net/fig/1169895" startId="F5">
<paragraph id="9BA580550C9180FCFEE15C2328E95FE4">
<label id="2373721971EF5F9E118D70D978FA37E5">Figure 5.</label>
</paragraph>
<paragraph id="F265641349A8578FF34C1993DAC6DE7F">
<taxonomicName id="2C4B28AF1BA1C28E53975C9A015C6670" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
<emphasis id="B5553B8C92A2B76F07FBE2DB121C3EBA" italics="true">Pseudodesmus bidoup</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7A2E8EAE0D1EF0E1F94E9CFCCCCBF111" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EN3AE" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
957). Gonopods.
<abbrev id="ABBRID0ES3AE" xlink_title="Scanning electron microscope">SEM</abbrev>
.
<emphasis id="5586C66F60CA89139AC56055E7B7EF66" bold="true">A, B.</emphasis>
Right anterior gonopods, ventral view (
<emphasis id="93C62E9DB340A91EEDC093185509BACF" bold="true">A</emphasis>
), anterior view (
<emphasis id="9D8A530F02CBD20F38E591FBE049CB79" bold="true">B</emphasis>
);
<emphasis id="2D52C04D42AD8B888863C47786EAD90B" bold="true">C, D.</emphasis>
Right posterior gonopod, ventral view (
<emphasis id="30036C842BA994DBF72528E52D9DE4DD" bold="true">C</emphasis>
), sublateral view (
<emphasis id="CAAF6DAAE6976A47A41100D0D3D522C9" bold="true">D</emphasis>
).
<emphasis id="73689A874E54CB31A919C4014E069A02" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EJ4AE" type="variation">
<paragraph id="5F08DBDBAF48A2276BB9A69CC8F7F9DA">
<heading id="B4BCA49E666E957D6F4BA220E67D2692" reason="title">Variation.</heading>
</paragraph>
<paragraph id="3CF7D554D9D722BDC0183AF025CEC2D6">
<emphasis id="EEF725DD7A87C32FFC38B6668E095C21" bold="true" italics="true">
Female
<typeStatus id="FC2438AEE6AC02D648D8930A8C2145FF" type="paratype">paratype</typeStatus>
</emphasis>
: number of body rings 4344 plus telson, length ca.
<quantity id="876C503FC7C10AF3964DB6E296F67A17" metricMagnitude="-2" metricUnit="m" metricValue="1.4125" metricValueMax="1.45" metricValueMin="1.375" unit="mm" value="14.125" valueMax="14.5" valueMin="13.75">13.7514.50 mm</quantity>
, length of midbody metazonites about
<quantity id="5E8893A7B423091404E4933A238A3EEA" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" unit="mm" value="0.29" valueMax="0.31" valueMin="0.27">0.270.31 mm</quantity>
, width of midbody metazonite and prozonite ca.
<quantity id="FF25EDB4DDAAB8C4902F432110271549" metricMagnitude="-3" metricUnit="m" metricValue="2.375" metricValueMax="2.5" metricValueMin="2.25" unit="mm" value="2.375" valueMax="2.5" valueMin="2.25">2.252.50 mm</quantity>
and
<quantity id="292106D82CAB30C724E16067274BEDD7" metricMagnitude="-3" metricUnit="m" metricValue="1.595" metricValueMax="1.63" metricValueMin="1.56" unit="mm" value="1.595" valueMax="1.63" valueMin="1.56">1.561.63 mm</quantity>
, respectively; ratio of width of head and width of collum = 0.96. Nonsexual characters as in male.
</paragraph>
<paragraph id="212CBD6C9B5914A85A200FFFD5204B52">
The male sample
<abbrev content-type="institution" id="ABBRID0EW4AE" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
961 has strong tubercles, even on the collum, then on every body ring the two (1 + 1) median posterior tubercles form a strong paramedian crest along the entire body. Anterior tubercles are also the same but smaller. Color is uniformly yellowish, without a dark pattern.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E24AE" type="etymology">
<paragraph id="9D63A93FB7300B9FF673554FAF1E29B0">
<heading id="51CDFADD92145320EDF2522A70094AF4" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="32866B95BF254F5B21746317683C3C79">
Named after the “ Bi Doup ” mountain,
<typeStatus id="9D3DE308D0DC4BDC84FB8289D48B6750">type</typeStatus>
locality. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EA5AE" type="distribution">
<paragraph id="346D1F8CC791ECDA5F07AC48DA23AB64">
<heading id="0CC800DFC93F58B81D3378D05463F087" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="622A5C3EB3C69BBCDE7FFC6B9B9D4480">
The species has been found in mixed forests and regenerated forests in
<collectingRegion id="9B61C8E7E03E6B05BBD220C95D676981" country="Vietnam" name="Kon Tum">Kon Tum</collectingRegion>
and
<collectingRegion id="550CDFD0EBAD8C58789A311CBD2AC906" country="Vietnam" name="Lam Dong">Lam Dong</collectingRegion>
provinces, Highlands of
<collectingCountry id="40459EE8B91C918CC66D31364288F374" name="Vietnam">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EF5AE" type="DNA barcoding">
<paragraph id="EAFC2304458201E0CC5323DF42731435">
<heading id="EA52DD1E929CEF6893AAC473509F7C1A" reason="title">DNA barcoding.</heading>
</paragraph>
<paragraph id="40BC61C3AE61056AFC607C9810E0126E">
Sequencing a fragment of the
<abbrev id="ABBRID0EL5AE" xlink_title="cytochrome c oxidase subunit I">COI</abbrev>
gene failed.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,344 @@
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<paragraph id="61874ACDB0CC081656094DFD630A2285">
<taxonomicName id="F9CFCAD751EFBC4C54609DD32A7E4A0D" ID-CoL="XGKK" authority="(Lang, 1884) Faubel, 1983" authorityName="Faubel" authorityYear="1983" baseAuthorityName="Lang" baseAuthorityYear="1884" family="Stylochoplanidae" genus="Comoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="agilis">
<emphasis id="7E98F21DF070192A12D6B6B08EDFC911" italics="true">Comoplana agilis</emphasis>
(
<bibRefCitation id="4521792E72321CA0EBEC2D375EEB4227" DOI="10.5962/bhl.title.10545" author="Lang A" refId="B43" refString="Lang A (1884) Die Polycladen (Seeplanarien) des Golfes von Neapel und der angrenzenden Meeresabschnitte. Eine Monographie. Fauna und Flora des Golfes von Neapel 11. W Engelmann, Leipzig. https: // doi. org / 10.5962 / bhl. title. 10545" year="1884">Lang, 1884</bibRefCitation>
)
<bibRefCitation id="6FBA1D4483C052DF3CA50F4584B1ADFA" author="Faubel A" journalOrPublisher="Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut" pagination="17-121" refId="B15" refString="Faubel A (1983) The Polycladida, Turbellaria. Proposal and stablishment of a new system. Part I. The Acotylea. Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut 80: 17 - 121." title="The Polycladida, Turbellaria. Proposal and stablishment of a new system. Part I. The Acotylea." volume="80" year="1983">Faubel, 1983</bibRefCitation>
</taxonomicName>
</paragraph>
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<paragraph id="5818C3C0542AFFE2CC087E813B8680DF">
<figureCitation id="6673771F9AD72EEE3C6DA3759BA9436D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comoplana agilis (MNCN 4.01 / 34634 to 3635). Histological sections are postero-anteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the tentacles and eyes; C, D. Histological sections of the reproductive system; E. Sagittal reconstruction of the reproductive system. Abbreviations: c, cilia; ca, common atrium; cp, common pore; cg, cemental glands; ed, ejaculatory duct; el, epithelial lining; ev, external vagina; i, intestine; iv, internal vagina; lv, Langs vesicle; mw, muscular wall; ov, oviduct; ph, pharynx; pp, penis papilla; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure14" httpUri="https://binary.pensoft.net/fig/1169827">Fig. 14</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="2BEA7595165EA3E2B63879D9665F90DA">
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<taxonomicName id="E502514957958B48CDCD8E2776C24D0F" ID-CoL="539Q4" authority="Lang, 1884" authorityName="Lang" authorityYear="1884" family="Stylochoplanidae" genus="Stylochoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="agilis">
<emphasis id="DE4405BDF5B25A831593223E65DC292A" italics="true">Stylochoplana agilis</emphasis>
<bibRefCitation id="188EA8835C34029403D2252FB4AA2A79" DOI="10.5962/bhl.title.10545" author="Lang A" refId="B43" refString="Lang A (1884) Die Polycladen (Seeplanarien) des Golfes von Neapel und der angrenzenden Meeresabschnitte. Eine Monographie. Fauna und Flora des Golfes von Neapel 11. W Engelmann, Leipzig. https: // doi. org / 10.5962 / bhl. title. 10545" year="1884">Lang, 1884</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EXLAK" type="material">
<paragraph id="CC098B686E62B9E638973A58DD293559">
<heading id="B8D5E4DB41280CDE2DEDF2CC54A5309E" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="2AA8E170AA129C6392FB44FBA31ACD73">
<materialsCitation id="192DE47A9F1DE2978DEB1E2FCDF4D98E" collectingDate="2018-12-08" collectionCode="MNHNC" specimenCode="MB 16-000125" specimenCount="1">
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<collectionCode id="27E91CE307F976B76A7B8E9A2D01473F">MNHNC</collectionCode>
</abbrev>
<specimenCode id="EC017EE1054760263B55450461584B3D" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB 16-000125</specimenCode>
, Station 4,
<date id="50A2A9843D39DD57A250751ED3C5BE2E" value="2018-12-08">
<collectingDate id="EEBF5246718CF16E5DAD773AF043BBFB" value="2018-12-08">8 December 2018</collectingDate>
</date>
,
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long
</materialsCitation>
; •
<materialsCitation id="5EC68F87AB35A60A1CA61F9633246EB3" collectingDate="2018-05-16" collectionCode="MNHNC" specimenCode="MB 16-000126" specimenCount="1">
<abbrev id="ABBRID0EBMAK" xlink_title="Museu Nacional de História Natural e da Ciência of Lisbon">
<collectionCode id="9F4802FC77BB7A5A06017CA0868C7CBD">MNHNC</collectionCode>
</abbrev>
<specimenCode id="D4C98F46641D9EDC50B068F6C3A0C99E" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB 16-000126</specimenCode>
, Station 8,
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<collectingDate id="3025BEB9E3F6E53FC94A95DEB25A7226" value="2018-05-16">16 May 2018</collectingDate>
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,
<quantity id="612B5FFF2192209E4F4F4F8DBDE6514D" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
long
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; •
<materialsCitation id="3E03E834CCFE753564664550715A9E6A" collectingDate="2018-05-17" collectionCode="MNHNC" specimenCode="MB 16-000127" specimenCount="1">
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<collectionCode id="ACFD98B5A1590810EDEEE8CE6B52601F">MNHNC</collectionCode>
</abbrev>
<specimenCode id="C4BDEE086197149B7D8A915F0B3EEB76" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB 16-000127</specimenCode>
, Station 8,
<date id="1734A529A63DFCA5ECDA4D9D96567F57" value="2018-05-17">
<collectingDate id="D506F247DEF876E8D14B636629BBB40D" value="2018-05-17">17 May 2018</collectingDate>
</date>
,
<quantity id="0D9E1F0E66492404D7CB1EB3E5DADEAD" metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
long
</materialsCitation>
; •
<materialsCitation id="5CDB2CB1500F7EF7A3B3F6640497D8E3" collectingDate="2015-10-28" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="BF29C8D482E83F1816A5F6660955FF17" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="777502E21D4B9178D166A0A3D2F10B11" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3631 to 3633, Station 11,
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</date>
,
<quantity id="C7E85C78FDAFF1BE15BB5F39164A49E5" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
long, sagittally sectioned into 3 slides
</materialsCitation>
; •
<materialsCitation id="933831C8FEEB070A21689525F95C9E9D" collectingDate="2015-10-28" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="CA6F0D93052A29EBB5F5F98133FDE32D" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="749F5AE7695496094DACE7F83BBB82CD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 4085, Station 11,
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<collectingDate id="104550284B822001E49E199ADC419D4D" value="2015-10-28">28 October 2015</collectingDate>
</date>
,
<quantity id="08A2A518851F544B3FB5F69FF83AF6E6" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
long; and
</materialsCitation>
<materialsCitation id="D01BD570620049B47D4D0C9FB22C5657" collectingDate="2015-10-29" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="FEA45772511B5A2C77904057B4F17135" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="012EE0529688482A886783405F1CDF79" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 34634 to 3635, Station 11,
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</date>
,
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long, sagittally sectioned into 2 slides. All of the measurements from the description refer to this specimen
</materialsCitation>
; •
<materialsCitation id="64560DB09975A4B1CAC56D3822C43F4E" collectingDate="2018-06-19" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="509488E6AB3619393D5E66BBD89124E8" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
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4.01
</specimenCode>
/ 4086, Station 11,
<date id="DA739A17FE0DBE0A14DAB3A5FFD00B28" value="2018-06-19">
<collectingDate id="866E9C4BBD11DDDA49387DDDDCB0D63D" value="2018-06-19">19 June 2018</collectingDate>
</date>
,
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long
</materialsCitation>
; •
<materialsCitation id="330B21CB4F53F2A3D792505990C121F6" collectingDate="2018-06-19" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="C8D8DEA968EA396D0BF6341A5AE9BAE2" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="9FDBB5FFE48AC9B21952FA7B52A2B84B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 4087, Station 11,
<date id="53B3FBCA559C53F426403F62A8F08938" value="2018-06-19">
<collectingDate id="51FAE1E493516D13A153D65D76B8DDBF" value="2018-06-19">19 June 2018</collectingDate>
</date>
,
<quantity id="A389202871BB9D811C2BEA600F7735C8" metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
long
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4MAK" type="description">
<paragraph id="B3F1E8A17607EFBD6FADE41BEA6704F9">
<heading id="9483FAB8D0309CC44D40520057F5C6F3" reason="title">Description.</heading>
</paragraph>
<paragraph id="A06C34E33F06877A9C3501A94AA19E80">
small
<taxonomicName id="6CFB06D016E67B049C065BA0687B4BC2" authorityName="Faubel" authorityYear="1983" family="Stylochoplanidae" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="family">Stylochoplanidae</taxonomicName>
, with lengths varying between
<quantity id="D00CB665743374B08A41BCDE33E61E1F" metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
and
<quantity id="A26C8E650405DAF23E73C5D4B415AE3F" metricMagnitude="-3" metricUnit="m" metricValue="8.0" unit="mm" value="8.0">8 mm</quantity>
(4.33 ±
<quantity id="305EAA8FC595C35E04B857109EB82FE6" metricMagnitude="-3" metricUnit="m" metricValue="1.66" unit="mm" value="1.66">1.66 mm</quantity>
). Body shape elongated much wider in the anterior region and at the blunt posterior end (Fig.
<figureCitation id="E6D391D5C59DADC0D38F0F109451AB35" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comoplana agilis (MNCN 4.01 / 34634 to 3635). Histological sections are postero-anteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the tentacles and eyes; C, D. Histological sections of the reproductive system; E. Sagittal reconstruction of the reproductive system. Abbreviations: c, cilia; ca, common atrium; cp, common pore; cg, cemental glands; ed, ejaculatory duct; el, epithelial lining; ev, external vagina; i, intestine; iv, internal vagina; lv, Langs vesicle; mw, muscular wall; ov, oviduct; ph, pharynx; pp, penis papilla; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure14" httpUri="https://binary.pensoft.net/fig/1169827">14 A</figureCitation>
). Dorsal colour differs from dark brown to dark green, with white or cream spots along the central longitudinal area. Conspicuous, cylindrical tentacles located at the anterior end to the beginning of body narrowing. Tentacular eyes (approximately 6 eyes per cluster) inside the tentacles and cerebral eyes (16 eyes per cluster) around the base of the tentacles. (Fig.
<figureCitation id="391E3293A7FE3B046676EB987EA642F4" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comoplana agilis (MNCN 4.01 / 34634 to 3635). Histological sections are postero-anteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the tentacles and eyes; C, D. Histological sections of the reproductive system; E. Sagittal reconstruction of the reproductive system. Abbreviations: c, cilia; ca, common atrium; cp, common pore; cg, cemental glands; ed, ejaculatory duct; el, epithelial lining; ev, external vagina; i, intestine; iv, internal vagina; lv, Langs vesicle; mw, muscular wall; ov, oviduct; ph, pharynx; pp, penis papilla; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure14" httpUri="https://binary.pensoft.net/fig/1169827">14 B</figureCitation>
). Pharynx ruffled, located more or less central. Oral pore situated in the posterior part of the pharynx. Genital pores joining in a common atrium (Fig.
<figureCitation id="31C4E2AE3B08CC613C75FD7D8850D89A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comoplana agilis (MNCN 4.01 / 34634 to 3635). Histological sections are postero-anteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the tentacles and eyes; C, D. Histological sections of the reproductive system; E. Sagittal reconstruction of the reproductive system. Abbreviations: c, cilia; ca, common atrium; cp, common pore; cg, cemental glands; ed, ejaculatory duct; el, epithelial lining; ev, external vagina; i, intestine; iv, internal vagina; lv, Langs vesicle; mw, muscular wall; ov, oviduct; ph, pharynx; pp, penis papilla; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure14" httpUri="https://binary.pensoft.net/fig/1169827">14 C E</figureCitation>
).
</paragraph>
<caption id="D5BE9407EDE30684E2DC8B57EC94AFF7" ID-DOI="10.3897/zse.100.128211.figure14" ID-arpha="6360A5B9-D601-54A3-899D-8BF9B5361B92" httpUri="https://binary.pensoft.net/fig/1169827" startId="F14">
<paragraph id="DDBE67E4D94932AB97B0CA6D87FA5844">
<label id="BE45AC701DEEB2BEB6E444754A395682">Figure 14.</label>
</paragraph>
<paragraph id="00FB5BB01E64C09CEBC5BB722ADD708B">
<taxonomicName id="807C170D70FDA74B65B53111EC661459" authorityName="Faubel" authorityYear="1983" baseAuthorityName="Lang" baseAuthorityYear="1884" family="Stylochoplanidae" genus="Comoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="agilis">
<emphasis id="16A6F3B554F2ABDE8676B6EFF6E5148A" italics="true">Comoplana agilis</emphasis>
</taxonomicName>
(
<named-content id="74641A1F8A20F3A5D9BC32D0F1F09FAB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01 / 34634 to 3635). Histological sections are postero-anteriorly oriented:
<emphasis id="812C8D34CA61E58DF4FA7DCB8744EADB" bold="true">A.</emphasis>
Dorsal view of the living specimen;
<emphasis id="DBBF763269904D772051BB79AA0DFFAB" bold="true">B.</emphasis>
Detail of the tentacles and eyes;
<emphasis id="082064B4BB055986C2D4A69F67FECF45" bold="true">C, D.</emphasis>
Histological sections of the reproductive system;
<emphasis id="B613C3230D077EF5E5F5B37D12F014D7" bold="true">E.</emphasis>
Sagittal reconstruction of the reproductive system. Abbreviations: c, cilia; ca, common atrium; cp, common pore; cg, cemental glands; ed, ejaculatory duct; el, epithelial lining; ev, external vagina; i, intestine; iv, internal vagina; lv, Langs vesicle; mw, muscular wall; ov, oviduct; ph, pharynx; pp, penis papilla; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EYOAK" type="Reproductive system">
<paragraph id="291C32CB301325A77E2C6AD71D5C94A3">
<heading id="0CE21D111F5BD9B2D54F6609B38200AB" reason="title">Reproductive system.</heading>
</paragraph>
<paragraph id="33E01CAD65BDE1A42014AFB4D9172871">
Male copulatory system with true seminal vesicle, interpolated prostatic vesicle, and conical penis papilla (Fig.
<figureCitation id="303255B7EE8427A6E46715F306429D89" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comoplana agilis (MNCN 4.01 / 34634 to 3635). Histological sections are postero-anteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the tentacles and eyes; C, D. Histological sections of the reproductive system; E. Sagittal reconstruction of the reproductive system. Abbreviations: c, cilia; ca, common atrium; cp, common pore; cg, cemental glands; ed, ejaculatory duct; el, epithelial lining; ev, external vagina; i, intestine; iv, internal vagina; lv, Langs vesicle; mw, muscular wall; ov, oviduct; ph, pharynx; pp, penis papilla; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure14" httpUri="https://binary.pensoft.net/fig/1169827">14 C E</figureCitation>
). Vas deferens run ventrally and enter proximally into the seminal vesicle through a common duct. Seminal vesicle voluminous (
<quantity id="D2918A4A57871DCD5BCEECF455329FDC" metricMagnitude="-4" metricUnit="m" metricValue="1.2" unit="mm" value="0.12">0.12 mm</quantity>
long and
<quantity id="F7FF0BF91BC9386CDD0C8B1AE3FF5541" metricMagnitude="-4" metricUnit="m" metricValue="2.3" unit="mm" value="0.23">0.23 mm</quantity>
wide), kidney-shaped, and connected proximally to the prostatic vesicle. A narrow ejaculatory duct runs from the seminal vesicle to the lumen of the prostatic vesicle (Fig.
<figureCitation id="5BBFDD31EAE27B21B60873DE68A55CD9" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comoplana agilis (MNCN 4.01 / 34634 to 3635). Histological sections are postero-anteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the tentacles and eyes; C, D. Histological sections of the reproductive system; E. Sagittal reconstruction of the reproductive system. Abbreviations: c, cilia; ca, common atrium; cp, common pore; cg, cemental glands; ed, ejaculatory duct; el, epithelial lining; ev, external vagina; i, intestine; iv, internal vagina; lv, Langs vesicle; mw, muscular wall; ov, oviduct; ph, pharynx; pp, penis papilla; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure14" httpUri="https://binary.pensoft.net/fig/1169827">14 D</figureCitation>
). Prostatic vesicle more or less rounded and well-developed (
<quantity id="95D93B573435749A8EA2C76DB89DBD77" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
long and
<quantity id="EB2924A16D8948DB97E62DFDEBC60EC6" metricMagnitude="-4" metricUnit="m" metricValue="2.8" unit="mm" value="0.28">0.28 mm</quantity>
wide), with a thick muscular wall (
<quantity id="F664B3E4029D86DFCEDD9F1EFC35D42F" metricMagnitude="-5" metricUnit="m" metricValue="4.0" unit="mm" value="0.04">0.04 mm</quantity>
) pierced by extravesicular glands. Epithelial lining of the prostatic vesicle smooth. Penis papilla
<quantity id="293A2C76041692CD5D1525B0CA7B3436" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
long, with a sinuous trajectory to the deep and ciliated common atrium.
</paragraph>
<paragraph id="545FBBE2EC90C3A3BDDDEF77CFCB5D6D">
Female system with a narrow external and internal vagina, Langs vesicle present (Fig.
<figureCitation id="4F7553FA985EC6FCC694D9787E26CFF8" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comoplana agilis (MNCN 4.01 / 34634 to 3635). Histological sections are postero-anteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the tentacles and eyes; C, D. Histological sections of the reproductive system; E. Sagittal reconstruction of the reproductive system. Abbreviations: c, cilia; ca, common atrium; cp, common pore; cg, cemental glands; ed, ejaculatory duct; el, epithelial lining; ev, external vagina; i, intestine; iv, internal vagina; lv, Langs vesicle; mw, muscular wall; ov, oviduct; ph, pharynx; pp, penis papilla; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure14" httpUri="https://binary.pensoft.net/fig/1169827">14 D</figureCitation>
). External vagina runs dorsally and turns backwards, continuing to the internal vagina, which receives the common oviduct ventrally. Langs vesicle large, rounded (
<quantity id="B0989C54C1A919388580CB8FE2C5B5CC" metricMagnitude="-4" metricUnit="m" metricValue="1.5" unit="mm" value="0.15">0.15 mm</quantity>
long and
<quantity id="6DD62C9F8709667488FF3B89FE48D943" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
wide), and lies on the ventral side of the body (Fig.
<figureCitation id="2DA4D43FE53463D08CB315B934FBDB1A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comoplana agilis (MNCN 4.01 / 34634 to 3635). Histological sections are postero-anteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the tentacles and eyes; C, D. Histological sections of the reproductive system; E. Sagittal reconstruction of the reproductive system. Abbreviations: c, cilia; ca, common atrium; cp, common pore; cg, cemental glands; ed, ejaculatory duct; el, epithelial lining; ev, external vagina; i, intestine; iv, internal vagina; lv, Langs vesicle; mw, muscular wall; ov, oviduct; ph, pharynx; pp, penis papilla; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure14" httpUri="https://binary.pensoft.net/fig/1169827">14 C E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQPAK" type="biology">
<paragraph id="4D9C39EF92A414D2FDE74621DC79C7BC">
<heading id="168D461FBB0CD9BC2754D73E90C6489F" reason="title">Biology.</heading>
</paragraph>
<paragraph id="5EBDF324BD7EEC8B932EF91DEE509C88">
Due to its small size and colouration, this species is very cryptic on substrates such as stones or sand. The specimens studied were mainly found on algae such as
<taxonomicName id="CDD76DD21C20EB518C155FCE197FBFD9" class="Hydrozoa" family="Halopterididae" genus="Halopteris" kingdom="Animalia" order="Leptothecata" phylum="Cnidaria" rank="species" species="scoparia">
<emphasis id="768ECC6FD0A8D6F557227E3107AEEF19" italics="true">Halopteris scoparia</emphasis>
</taxonomicName>
,
<taxonomicName id="5BF6CFD154FF2FFCDDBD6C6918C65CA4" class="Insecta" family="Tettigoniidae" genus="Dictyota" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="dichotoma">
<emphasis id="C625673C9283C0D18167A03928DB79DF" italics="true">Dictyota dichotoma</emphasis>
</taxonomicName>
(Ochrophyta), and
<taxonomicName id="55E3C9A4F93C16E26C8E4D2D08FD966E" authority="(Rodophyta)" baseAuthorityName="Rodophyta" family="Corallinaceae" genus="Ellisolandia" kingdom="Plantae" order="Cryptonemiales" phylum="Rhodophyta" rank="species" species="elongata">
<emphasis id="616629A2D80588C2C44EFB19C9C6A139" italics="true">Ellisolandia elongata</emphasis>
(Rodophyta)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3QAK" type="distribution">
<paragraph id="91876100052C9BA498CB68C7A6878B3B">
<heading id="C44667A406BE7785BF94C7E32A2F6576" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="7C54D320498639EA50ADCBE8A9A179E2">
Widely known from the Mediterranean, this species was reported in Naples,
<collectingCountry id="27593BA7287AEC77DEF9D0C6BF148E30" name="Italy">Italy</collectingCountry>
(
<bibRefCitation id="A6DD67D88EEC97275B87A90ECAFFC5E1" DOI="10.5962/bhl.title.10545" author="Lang" firstAuthor="Lang" refId="B43" refString="Lang A (1884) Die Polycladen (Seeplanarien) des Golfes von Neapel und der angrenzenden Meeresabschnitte. Eine Monographie. Fauna und Flora des Golfes von Neapel 11. W Engelmann, Leipzig. https://doi.org/10.5962/bhl.title.10545" year="1884">Lang 1884</bibRefCitation>
), the coasts of Catalonia (
<bibRefCitation id="91710170576794087D4696D8D857FCD4" author="Novell" firstAuthor="Novell" refId="B55" refString="Novell C (2001) Contribution to the knowledge of polyclad Turbellaria of the Catalan coast. PhD Thesis, University of Barcelona, Spain. http://www.tesisenxarxa.net/TESIS_UB/AVAILABLE/TDX-0107104-121621//Tesis.PDF" url="http://www.tesisenxarxa.net/TESIS_UB/AVAILABLE/TDX-0107104-121621//Tesis.PDF" year="2001">Novell 2001</bibRefCitation>
), and the Atlantic coasts of Europe, such as Galicia (
<bibRefCitation id="06D3B59FA5357A482E83F6A7BD250397" DOI="10.1007/s13127-017-0344-4" author="Aguado" etAl="et al." firstAuthor="Aguado" issue="4" journalOrPublisher="Organisms Diversity &amp; Evolution" pagination="767-778" refId="B2" refString="Aguado MT, Noreña C, Alcaraz L, Marquina D, Brusa F, Damborenea C, Almon B, Bleidorn C, Grande C (2017) Phylogeny of Polycladida (Platyhelminthes) based on mtDNA data. Organisms Diversity &amp; Evolution 17 (4): 767778. https://doi.org/10.1007/s13127-017-0344-4" title="Phylogeny of Polycladida (Platyhelminthes) based on mtDNA data." volume="17" year="2017">Aguado et al. 2017</bibRefCitation>
), the North Sea (
<bibRefCitation id="EDDF48F405C9CD775BCF0763D7E3A8F3" author="Karez" firstAuthor="Karez" refId="B39" refString="Karez R (1991) Die Besiedlung sublitoraler Rotalgenam Felssockel von Helgoland. Dipl. Arb. Univ. Marburg, 143 pp." year="1991">Karez 1991</bibRefCitation>
;
<bibRefCitation id="894DDEE706747A56E62C7E2C15DDBE0B" DOI="10.1007/BF02366182" author="Harms" firstAuthor="Harms" journalOrPublisher="Helgoländer Wissenschaftliche Meeresuntersuchungen" pagination="1-34" refId="B29" refString="Harms J (1993) Check-list of species (algae, invertebrales and vertebrales) found in the vicinity of the island of Helgoland (North Sea, German Bight) - a review of recent records. Helgoländer Wissenschaftliche Meeresuntersuchungen 41: 134. https://doi.org/10.1007/BF02366182" title="Check-list of species (algae, invertebrales and vertebrales) found in the vicinity of the island of Helgoland (North Sea, German Bight) - a review of recent records." volume="41" year="1993">Harms 1993</bibRefCitation>
), and the British coasts (
<bibRefCitation id="31A5548845D00CC2B94558962A613F35" author="Howson and Picton" firstAuthor="Howson" journalOrPublisher="Ulster Museum and the marine conservation society, Belfast and Ross-on Wye Ulster museum publication" pagination="1-508" refId="B33" refString="Howson CM, Picton BE (1997) The species directory of the marine fauna and flora of the british isles and surrounding seas. Ulster Museum and the marine conservation society, Belfast and Ross-on Wye Ulster museum publication 276: 1508." title="The species directory of the marine fauna and flora of the british isles and surrounding seas." volume="276" year="1997">Howson and Picton 1997</bibRefCitation>
). This is the first report of this species in
<collectingCountry id="4A8E60E95AD103A2DE6A0959858F5464" name="Portugal">Portugal</collectingCountry>
and Andalusia.
</paragraph>
</subSubSection>
</treatment>
</document>

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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2D141BCC724BCBD8B7688713917BCFEA">journal article</mods:classification>
<mods:identifier id="E6E26FE18E0EB5CE33A709D60597330C" type="DOI">10.3897/phytokeys.248.130311</mods:identifier>
</mods:mods>
<treatment id="6D5026F191765A6DB8F6C721079CB3DC" ID-arpha="6D5026F1-9176-5A6D-B8F6-C721079CB3DC" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC" httpUri="http://treatment.plazi.org/id/6D5026F191765A6DB8F6C721079CB3DC">
<subSubSection id="55C94E8DE74F96817E259E2FA8930EEE" type="nomenclature">
<paragraph id="A0D9DA1A3F98B2BFB0C1A675C356943C">
<heading id="B3007F3243683449783E374B5BAC670E" reason="label">2.</heading>
<taxonomicName id="F3CAA8170811B8CE06BE0B80EFF426DB" ID-CoL="6JX2X" authority="King, J. Asiat. Soc. Bengal, Pt." class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="90C145E54BBBFCA75B950BAE9C8259DB" italics="true">Garcinia prainiana</emphasis>
King, J. Asiat. Soc. Bengal, Pt.
</taxonomicName>
2,
<bibCitation id="54471680C7EAF69CFA1A02F40D9FE8DF" issue="2" journal="Nat. Hist." page="171" score="4.075586588997398" volume="59" year="1890">Nat. Hist. 59 (2): 171. 1890</bibCitation>
; Vesque in A. DC. &amp; C. DC., Monogr. Phan. 8: 329. 1893; Ridl., Fl.
<bibCitation id="7FE2F9C4F9D2D7D1EC19DE38C27B44A4" journal="Malay Penins." page="180" score="0.004206541313292987" volume="1" year="1922">Malay Penins. 1: 180. 1922</bibCitation>
; Corner, Wayside Trees Mal. 1: 320. fig. 112. ed. 2. 1952; Corner &amp; Watan., Ill. Guide Trop. Pl.: t. 193. 1969; Whitmore in Whitmore, Tree Fl.
<bibCitation id="38E14D52166C7CF9B5043993D1B4B78C" journal="Malaya" page="220" score="0.125" volume="2" year="1973">Malaya 2: 220. 1973</bibCitation>
;
<bibCitation id="8CF2B413C06C247F75018D285C2AA1D5" author="I. M. Turner" issue="1" journal="Gard. Bull. Singapore" page="263" score="86.90123279129422" volume="47" year="1995">
I. M. Turner,
<bibCitation id="7804A84832C0A78927F66613B2952F57" issue="1" journal="Gard. Bull. Singapore" page="263" score="86.90123279129422" volume="47" year="1995">
Gard. Bull.
<collectingCountry id="B1950C3C8E5C7F0537DD34460EF1334E" name="Singapore">Singapore</collectingCountry>
47 (1): 263. 1995
</bibCitation>
</bibCitation>
; Ngernsaengsaruay et al., PeerJ, DOI 10.7717/peerj.16572: 28. figs 13, 14, 15. 2023.
</paragraph>
</subSubSection>
<subSubSection id="972F3B021E74B6DEF0841D748A8E242D" type="description">
<paragraph id="1900A9F723912A9A6A5BAF5BBD192A18">
<figureCitation id="9717E9D091084071070BCCB62282513B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Garcinia prainiana A, B branchlets, young leaves, and mature leaves C inflorescences with male flower buds and open male flower D branches and branchlets with mature and ripe fruits E branchlets, leaves, and mature and ripe fruits F ripe fruits (pericarp removed and in transverse section showing seeds with fleshy pulp). Photos: Chatchai Ngernsaengsaruay (A C, E, F), G Rawit Sichaikhan (D)." figureDoi="10.3897/phytokeys.248.130311.figure6" httpUri="https://binary.pensoft.net/fig/1169588">Fig. 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESCBG" type="material">
<paragraph id="29C5D013AD2075E023356ACE5922853C">
<heading id="EFE20B558347531D7DE12EE629BB4741" reason="title">
<typeStatus id="7205429A1D9120814C0B990F18193396">Type</typeStatus>
.
</heading>
</paragraph>
<paragraph id="D7D4426FF6CC58C96928925CB7D95EFE">
Peninsular
<collectingCountry id="8FA10E4B66B3C54CA2D56AA7EBDC78CD" name="Malaysia">Malaysia</collectingCountry>
<materialsCitation id="62ADDEB81FC343901838A4AEC649BD5B" collectingDate="1885-02" collectorName="B. Scortechini" location="Kuala Dipang" specimenCount="2" stateProvince="Perak" typeStatus="holotype">
<collectingRegion id="C3162C387001DA446B0BE43924279B2D" country="Malaysia" name="Perak">Perak</collectingRegion>
,
<location id="8122C685749604A9F29C8CDAF3254572" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:8122C685749604A9F29C8CDAF3254572" name="Kuala Dipang" stateProvince="Perak">Kuala Dipang</location>
(originally Kwala Dipang on the label; originally published Kwala Dynong ), fl.,
<date id="01BAF82A7683A2D2B3089987C8E32C18" value="1885-02">
<collectingDate id="DED3B51B9E407FAD154204E8C4312E3D" value="1885-02">February 1885</collectingDate>
</date>
,
<emphasis id="518A885C131AF965675E439DAEF95315" italics="true">
<collectorName id="0B41F542CB2402BCB3588B00E59C5FD4">B. Scortechini</collectorName>
1796
</emphasis>
(
<typeStatus id="0A07A7A3A490B4AA85DFDC10D5859E64" type="lectotype">lectotype</typeStatus>
designated by
<bibRefCitation id="3B46F5BD123FDF8C816DDA546517FE5C" DOI="10.7717/peerj.16572" author="Ngernsaengsaruay C &amp; Chanton P &amp; Chaiprasongsuk M &amp; Leksungnoen N" refId="B25" refString="Ngernsaengsaruay C, Chanton P, Chaiprasongsuk M, Leksungnoen N (2023 b) A taxonomic revision of Garcinia section Xanthochymus (Clusiaceae) in Thailand. PeerJ 11 (e 16572): 154. https://doi.org/10.7717/peerj.16572">Ngernsaengsaruay et al. 2023 b</bibRefCitation>
: 28, CAL [CAL 0000005844, photo seen];
<typeStatus id="AB32596660E884D327BBB55E323E309C" type="isolectotype">isolectotypes</typeStatus>
K [K 000677678!], P [P 04701324, photo seen])
</materialsCitation>
.
</paragraph>
<caption id="5D1124DD2F06E9B52D670E677EF64A7B" ID-DOI="10.3897/phytokeys.248.130311.figure6" ID-arpha="ADC0B841-A8EA-5DB3-833E-75751680C1C1" httpUri="https://binary.pensoft.net/fig/1169588" startId="F6">
<paragraph id="7B4CDA4F6E46D50050C0CAA2D88B10E9">
<label id="3CC834745F097F4FDC41FE0805FB2B7B">Figure 6.</label>
</paragraph>
<paragraph id="00A493033219B214621EA9746A645824">
<taxonomicName id="1F67C500FFFA043E2243523D68336F65" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="AE801F3AC4FF1F65837BAE4D8286FE22" italics="true">Garcinia prainiana</emphasis>
</taxonomicName>
<emphasis id="2B8BF9220AC263C0DAB373FF7B4AB114" bold="true">A, B</emphasis>
branchlets, young leaves, and mature leaves
<emphasis id="A2EB1FD663E86499338104325AB8CA7A" bold="true">C</emphasis>
inflorescences with male flower buds and open male flower
<emphasis id="935C26B28120712F287D66BE3218AB14" bold="true">D</emphasis>
branches and branchlets with mature and ripe fruits
<emphasis id="EE35A40BF3FF70F8A1C070047E70265F" bold="true">E</emphasis>
branchlets, leaves, and mature and ripe fruits
<emphasis id="4E6AA1EAADBF16ABBB81832D79061D5C" bold="true">F</emphasis>
ripe fruits (pericarp removed and in transverse section showing seeds with fleshy pulp). Photos: Chatchai Ngernsaengsaruay (
<emphasis id="B67DB661B680F65C8D448AAB18453D2D" bold="true">A C, E, F</emphasis>
), G Rawit Sichaikhan (
<emphasis id="49BFF90DD3A71EF86342EAF50BEFDD5A" bold="true">D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EEEBG" type="description">
<paragraph id="5714B0D8544B962BEB6F3A307887AD44">
<heading id="5519026257F5E5B210876153A42FF666" reason="title">Description.</heading>
</paragraph>
<paragraph id="1FD69D8FD57B41334C2F4973656E5679">
<emphasis id="45683939E7E3702E8BCD23733670AA29" bold="true" italics="true">Habit</emphasis>
evergreen trees,
<quantity id="B508DC1680F699C69530B1BE0D366876" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" unit="m" value="7.5" valueMax="12.0" valueMin="3.0">312 m</quantity>
tall,
<quantity id="30314FCBA18284BB281FC4D221C4C599" metricMagnitude="-1" metricUnit="m" metricValue="4.5" metricValueMax="7.5" metricValueMin="1.5" unit="cm" value="45.0" valueMax="75.0" valueMin="15.0">1575 cm</quantity>
GBH; latex white, sticky; branches decussate, horizontal or nearly horizontal; branchlets 4 - ridged, glabrous.
<emphasis id="0BDC8CF414F1679FCBC32DE9E50B9461" bold="true" italics="true">Bark</emphasis>
pale brown, grayish brown or blackish brown, smooth or slightly rough; inner bark pale yellow.
<emphasis id="7E17BD3F6A220B19C4067A661EB9B00B" bold="true" italics="true">Terminal bud</emphasis>
concealed between the bases of the uppermost pair of petioles.
<emphasis id="257380F2CE97F70F4CA4DF460DAD6DD9" bold="true" italics="true">Leaves</emphasis>
decussate; lamina elliptic, oblong or elliptic-oblong, sometimes narrowly oblong, 12.527.5 ×
<quantity id="ED8B25F1A6EF9587F84B7C2A49A5FA76" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="11.5" metricValueMin="5.5" unit="cm" value="8.5" valueMax="11.5" valueMin="5.5">5.511.5 cm</quantity>
, apex acute or obtuse, base subcordate, often subamplexicaul, margin repand and slightly revolute, coriaceous, bullate or slightly bullate, shiny dark green above, paler below, glabrous on both surfaces, midrib flattened above, raised as a prominent ridge below, secondary veins 920 each side, curving towards the margin and connected in distinct loops and united into an intramarginal vein, flattened above, raised below, conspicuous on both surfaces, intersecondary veins conspicuous, veinlets reticulate, visible on both surfaces, interrupted long wavy lines present, of differing lengths, nearly parallel to the midrib, running across the secondary veins to the apex, visible below; petiole green,
<quantity id="5C50EB20C7B17296182C113294C824F7" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="6.0" metricValueMin="1.5" unit="mm" value="3.75" valueMax="6.0" valueMin="1.5">1.56 mm</quantity>
long,
<quantity id="84E273D74352A00F0B614208DEB4E01D" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">25 mm</quantity>
in diam., not grooved, transversely rugose, glabrous, with a basal appendage clasping the branchlet; young leaves shiny pale green.
<emphasis id="2D5FA5C79FC760D446D5B7D6D9769390" bold="true" italics="true">Inflorescences</emphasis>
terminal, sometimes on short, leafless lateral branchlets, cymose, usually in dense fascicles of several to many flowers.
<emphasis id="2558815246E494B06CEE8CAFF26D9692" bold="true" italics="true">Flowers</emphasis>
unisexual, plants dioecious, 5 - merous, fully open flowers with spreading petals; bracteolate; sepals 5 and petals 5, quincuncial, coriaceous, glabrous.
<emphasis id="81637FD0C83D54E5E913AE762BF384AE" bold="true" italics="true">Male flowers</emphasis>
<quantity id="C2CAEAA5AF2C4A7E59D8AF92D3717CE3" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" unit="cm" value="3.0" valueMax="3.5" valueMin="2.5">2.53.5 cm</quantity>
in diam.; bracteoles pale green, triangular 2.34.5 ×
<quantity id="38E1131A2CE00934FA7F818E846AA8DB" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.7" metricValueMin="1.8" unit="mm" value="2.75" valueMax="3.7" valueMin="1.8">1.83.7 mm</quantity>
, apex acute, conduplicate with a central keel; pedicel pinkish green, reddish green or greenish red,
<quantity id="AEDB7BEB6A330BFB30E5953E31F6FA18" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">36 mm</quantity>
long,
<quantity id="1892F054178168425856084F42532672" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.54 mm</quantity>
in diam., widened at the apical part, terete, glabrous; sepals pinkish green, reddish green or greenish red, concave, broadly ovate or suborbicular 4.88 ×
<quantity id="DA2A7B4C428F8066C6C2911AFC23F00D" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="7.8" metricValueMin="5.0" unit="mm" value="6.4" valueMax="7.8" valueMin="5.0">57.8 mm</quantity>
, unequal, apex rounded; petals variable in color: pale yellow, yellowish pink, yellowish red, pinkish red, pink or red, broadly obovate or obovate, 0.81.4 ×
<quantity id="8B82E057F54D0D93058E37AE0F57FDEF" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" unit="cm" value="0.85" valueMax="1.1" valueMin="0.6">0.61.1 cm</quantity>
, subequal, sometimes unequal, apex rounded; stamens numerous, united into 5 bundles surrounding a pistillode, antepetalous,
<quantity id="5F168E88167D87A07CFC5F65A8C32468" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="4.2" metricValueMin="1.7" unit="mm" value="2.95" valueMax="4.2" valueMin="1.7">1.74.2 mm</quantity>
long, each bundle
<quantity id="A5234530E4E3CDB355BBB3AF93C75CA1" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="4.0" metricValueMin="1.2" unit="mm" value="2.6" valueMax="4.0" valueMin="1.2">1.24 mm</quantity>
wide, pale yellow, pink or red; filaments fused throughout their entire length; anthers yellow,
<quantity id="6DD46612A86E16568F01D2EF35F4BC1D" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" unit="mm" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">0.30.6 mm</quantity>
long; pistillode fungiform,
<quantity id="139A4A4946B15FF90611B50CF3D8ECFF" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" unit="mm" value="6.5" valueMax="7.5" valueMin="5.5">5.57.5 mm</quantity>
long; sterile stigma pale yellow, pink or red, sessile, convex, indistinctly lobed,
<quantity id="03D32A858C7345788217A6564E5258B6" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
in diam., papillate.
<emphasis id="68A8D67816F29E78AB4E2ABE50FDBE99" bold="true" italics="true">Female flowers</emphasis>
<quantity id="1CC8C2E5D0EF5296A501CAA0383D25AF" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" unit="cm" value="3.25" valueMax="4.0" valueMin="2.5">2.54 cm</quantity>
in diam.; bracteoles and pedicel same as in male flowers; sepals and petals same as or slightly larger than in male flowers; staminodes absent; pistil fungiform,
<quantity id="9FB288E47EECD0AEC67C33E6AE9059B4" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="8.5" metricValueMin="6.0" unit="mm" value="7.25" valueMax="8.5" valueMin="6.0">68.5 mm</quantity>
long; ovary pale green, depressed globose 46 ×
<quantity id="BFB234B8B3B6AD9F44F78BAFBFCF71ED" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" unit="mm" value="5.5" valueMax="6.5" valueMin="4.5">4.56.5 mm</quantity>
, unlobed, glabrous, 58 - locular; stigma pale yellow, pink or red, sessile, convex, weakly 58 - lobed or indistinctly lobed,
<quantity id="030FC7ADE2FE8240E9CA5B73A2B11808" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
in diam., papillate.
<emphasis id="70D60A1DE0FCF2D73EF2D0FB864D18D4" bold="true" italics="true">Fruits</emphasis>
berries, depressed globose or depressed subglobose, sometimes globose, 23.5 ×
<quantity id="0E4E4523A56393EA4A721B0E27057353" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="5.3" metricValueMin="2.0" unit="cm" value="3.65" valueMax="5.3" valueMin="2.0">25.3 cm</quantity>
, sometimes oblique, asymmetrical, unlobed, slightly concave or flattened at the apex, green, turning greenish yellow, bright yellow, orangish yellow and bright orange when ripe, smooth, glabrous, glossy, then exocarp becoming dark brownish black and slightly sinuously wrinkled when dry, pericarp
<quantity id="B47FB61EA5022FE31816E3E865F217B3" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="8.0" metricValueMin="3.5" unit="mm" value="5.75" valueMax="8.0" valueMin="3.5">3.58 mm</quantity>
thick, exocarp thin; persistent stigma dark brown or blackish brown, circular, button-like,
<quantity id="ECE11FB01B1945A0A11B1A3706255E66" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" unit="cm" value="0.85" valueMax="1.1" valueMin="0.6">0.61.1 cm</quantity>
in diam., slightly concave or flattened, weakly 58 - lobed or indistinctly lobed, papillate; persistent sepals pale green, turning yellowish green and orangish green, larger than in flowering material; fruiting stalk green, thick,
<quantity id="52723390DF4D49A50D9A7D462A92AB58" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" unit="cm" value="0.95" valueMax="1.5" valueMin="0.4">0.41.5 cm</quantity>
long,
<quantity id="95DC65200C41E062DD910E02E2B84A78" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">36 mm</quantity>
in diam.,
<emphasis id="A759D60CCEB2C0DF672B1F0F50E5250D" bold="true" italics="true">Seeds</emphasis>
16, often aborted, brown, broadly ellipsoid, ellipsoid or subglobose, 0.91.6 ×
<quantity id="8623DE03D63E3EF78FA36508FDE89143" metricMagnitude="-2" metricUnit="m" metricValue="1.0499999999999998" metricValueMax="1.4" metricValueMin="0.7" unit="cm" value="1.0499999999999998" valueMax="1.4" valueMin="0.7">0.71.4 cm</quantity>
, with pale orange fleshy pulp. The description of this species was taken from
<bibRefCitation id="73F9FCD941B330A2DAB0FAC813E24DAD" DOI="10.7717/peerj.16572" author="Ngernsaengsaruay C &amp; Chanton P &amp; Chaiprasongsuk M &amp; Leksungnoen N" refId="B25" refString="Ngernsaengsaruay C, Chanton P, Chaiprasongsuk M, Leksungnoen N (2023 b) A taxonomic revision of Garcinia section Xanthochymus (Clusiaceae) in Thailand. PeerJ 11 (e 16572): 154. https://doi.org/10.7717/peerj.16572">Ngernsaengsaruay et al. 2023 b</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOFBG" type="distribution">
<paragraph id="EB61EFAA13C5B5246B18FEB42D462964">
<heading id="7A3BC53082BF820C74B19554CED5280B" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="143FF9F96090F9F965A066BD6F900E07">
Known only from Peninsular
<collectingCountry id="C2C209436C08351B4D2650E545CADA76" name="Thailand">Thailand</collectingCountry>
and Peninsular
<collectingCountry id="84D925E3B4062E903650C0573297F21C" name="Malaysia">Malaysia</collectingCountry>
. It is widely distributed in Peninsular
<collectingCountry id="7DF9A43D4724E944641C71A5B3469EAC" name="Malaysia">Malaysia</collectingCountry>
(
<collectingRegion id="7AD369D313A504E688887D77A495EB8D" country="Malaysia" name="Perlis">Perlis</collectingRegion>
,
<collectingRegion id="EDD8F5AEFE3274BD91B8A75B2B98632F" country="Malaysia" name="Kedah">Kedah</collectingRegion>
,
<collectingRegion id="6E2E9639B5DD46822136AAB93CF3DFCA" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
,
<collectingRegion id="B473BEA58496B8072503C6738E830E38" country="Malaysia" name="Perak">Perak</collectingRegion>
,
<collectingRegion id="A3BC8C68CC1C88463558D954538E3424" country="Malaysia" name="Kelantan">Kelantan</collectingRegion>
,
<collectingRegion id="77662899DD0334C7AE63C63A5AFC2B14" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
,
<collectingRegion id="DABF3ADEDF2196F5CB79318325DE742B" country="Malaysia" name="Pahang">Pahang</collectingRegion>
,
<collectingRegion id="6F946A068DD4DF11AF456958930E4790" country="Malaysia" name="Selangor">Selangor</collectingRegion>
,
<collectingRegion id="92C4CC52F77C8F26A298782DC8E66B4B" country="Malaysia" name="Negeri Sembilan">Negeri Sembilan</collectingRegion>
,
<collectingCountry id="5CA810923D2011D79567936CF3292A01" name="Malaysia">Malacca</collectingCountry>
,
<collectingRegion id="4F2C6CE49287AE7D0B685AD6C99AC46A" country="Malaysia" name="Johor">Johor</collectingRegion>
) (
<bibRefCitation id="D1E56208F37D0D1342892EBE25737281" author="Corner" firstAuthor="Corner" refId="B6" refString="Corner EJH (1952) Wayside Trees of Malaya. 2 nd edn. The Government Printing Office, Singapore, 312320." year="1952">Corner 1952</bibRefCitation>
;
<bibRefCitation id="1E50FF29039616AAB4FD8D1CB0CA6BAE" author="Whitmore" editor="Whitmore TC" firstAuthor="Whitmore" journalOrPublisher="2. Wing Tai Cheung Printing Co. Ltd, Hong Kong" pagination="62-236" refId="B49" refString="Whitmore TC (1973) Guttiferae. In: Whitmore TC (Ed.) Tree Flora of Malaya Vol. 2. Wing Tai Cheung Printing Co. Ltd, Hong Kong, 62236." title="Guttiferae." volumeTitle="Tree Flora of Malaya Vol." year="1973">Whitmore 1973</bibRefCitation>
;
<bibRefCitation id="D413C1C2137F0C86090264236F2DE324" DOI="10.26492/gbs71(1).2019-01" author="Turner" firstAuthor="Turner" issue="1" journalOrPublisher="Gardens Bulletin (Singapore)" pagination="1-346" refId="B46" refString="Turner IM (1995) A catalogue of the vascular plants of Malaya. Gardens Bulletin (Singapore) 47 (1): 1346. https://doi.org/10.26492/gbs71 (1). 2019-01" title="A catalogue of the vascular plants of Malaya." volume="47" year="1995">Turner 1995</bibRefCitation>
;
<bibRefCitation id="211BD43C28124E97C983197CDE6A02F6" author="Azuan and Salma" firstAuthor="Azuan" refId="B2" refString="Azuan A, Salma I (2018) Fruit for the future: cerapu (Garcinia prainiana) [Abstract] in Conference: MARDI Science and Technology Exhibition (MSTE 2008) at Serdang, Selangor." year="2018">Azuan and Salma 2018</bibRefCitation>
). It can be found mainly in
<collectingRegion id="671880D0EA033F3F064B685F3B2F0710" country="Malaysia" name="Pahang">Pahang</collectingRegion>
,
<collectingRegion id="DBC8153BE4FE4E8A78091C5AC2933F5D" country="Malaysia" name="Perak">Perak</collectingRegion>
, and
<collectingRegion id="0EC034BE75C179F83D40067121403F46" country="Malaysia" name="Negeri Sembilan">Negeri Sembilan</collectingRegion>
(
<bibRefCitation id="624C4E84734E5A628B6E6B45BDADF7CE" author="Syazwani" firstAuthor="Syazwani" refId="B40" refString="Syazwani A (2020) Garcinia prainiana King (Clusiaceae). Malaysia Biodiversity Information System (MyBIS). https://www.mybis.gov.my/art/275" url="https://www.mybis.gov.my/art/275" year="2020">Syazwani 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIGBG" type="distribution">
<paragraph id="89DFA62C6038B8D23EF5C0ACB187EF8E">
<heading id="9E8DA72F261EEE04DE384C9D3B78851F" reason="title">
Distribution in
<collectingCountry id="6C45D2BAC9A10976BB039CE052558584" name="Thailand">Thailand</collectingCountry>
.
</heading>
</paragraph>
<paragraph id="02B78966FD3640D9C6F82EF5B8A44A06">
<emphasis id="26661048452A2B7E831101A1A8B9FF62" bold="true">Peninsular</emphasis>
:
<collectingRegion id="6F914844E150606A6E0D7FD0A86599DB" country="Thailand" name="Yala">Yala</collectingRegion>
(Than To),
<collectingRegion id="23796B5A54F66AE143F5579DCAD7EEB4" country="Thailand" name="Narathiwat">Narathiwat</collectingRegion>
(Waeng, Su-ngai Kolok).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQGBG" type="habitat">
<paragraph id="29C6A28E4F17FEA4E3A634D6FC4CC6A1">
<heading id="421E2C3339DFCE923BFA8683AFD6E3CE" reason="title">Habitat and ecology.</heading>
</paragraph>
<paragraph id="1BEAF9EAB776B01CB8D5917313AA95EE">
It is found in tropical lowland evergreen rain forests, occasionally along streams,
<quantity id="0E619C9017BE1D70CAEB6B15E08010D0" metricMagnitude="2" metricUnit="m" metricValue="1.15" metricValueMax="2.0" metricValueMin="0.3" unit="m" value="115.0" valueMax="200.0" valueMin="30.0">30200 m</quantity>
amsl. It is also cultivated in villages and botanical gardens.
</paragraph>
<paragraph id="951805AC3623F267B7F5B1A4C728A7D1">
In Peninsular
<collectingCountry id="3DFE06B71CC1F0272593C4E8C01044C9" name="Malaysia">Malaysia</collectingCountry>
, it occurs in lowland and hill forests, on hillsides and ridges up to elevations of
<quantity id="B1B3D896E8447A47E76E2C3CB740FE20" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1,000 m</quantity>
amsl. It is also cultivated in villages (
<bibRefCitation id="F7A9DAC4207E3DE90BFDBA74B9B40313" author="Whitmore" editor="Whitmore TC" firstAuthor="Whitmore" journalOrPublisher="2. Wing Tai Cheung Printing Co. Ltd, Hong Kong" pagination="62-236" refId="B49" refString="Whitmore TC (1973) Guttiferae. In: Whitmore TC (Ed.) Tree Flora of Malaya Vol. 2. Wing Tai Cheung Printing Co. Ltd, Hong Kong, 62236." title="Guttiferae." volumeTitle="Tree Flora of Malaya Vol." year="1973">Whitmore 1973</bibRefCitation>
;
<bibRefCitation id="08C92BB71BF7F615524AAB7D112290DC" author="Syazwani" firstAuthor="Syazwani" refId="B40" refString="Syazwani A (2020) Garcinia prainiana King (Clusiaceae). Malaysia Biodiversity Information System (MyBIS). https://www.mybis.gov.my/art/275" url="https://www.mybis.gov.my/art/275" year="2020">Syazwani 2020</bibRefCitation>
; from the specimen
<emphasis id="13D974DDF34BC82FE757313F9F894459" italics="true">T. C. Whitmore Kep. FRI 4018</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBHBG" type="phenology">
<paragraph id="1EA194EBBCB71B9472B6ACE00E702423">
<heading id="5D5467B52EE5086D18163544911F8F01" reason="title">Phenology.</heading>
</paragraph>
<paragraph id="59C712B459E8C2746793DF9D10C5674B">Flowering and fruiting more than once; flowering nearly throughout the year, usually in February to May; fruiting April to June and September to December.</paragraph>
</subSubSection>
<subSubSection id="SECID0EGHBG" type="conservation status">
<paragraph id="9A8D4E4CA5C71931D80EB2DB3F1FBC02">
<heading id="6FE2B14B784EB76EF94EAA5E742000D4" reason="title">Conservation status.</heading>
</paragraph>
<paragraph id="1FD7FFC121496F5554ABDAC011A1DA3C">
<abbrev id="ABBRID0EMHBG" xlink_title="Least concern">LC</abbrev>
(
<bibRefCitation id="B3BFB14113F426AD6701D6B2568D6D86" DOI="10.7717/peerj.16572" author="Ngernsaengsaruay C &amp; Chanton P &amp; Chaiprasongsuk M &amp; Leksungnoen N" refId="B25" refString="Ngernsaengsaruay C, Chanton P, Chaiprasongsuk M, Leksungnoen N (2023 b) A taxonomic revision of Garcinia section Xanthochymus (Clusiaceae) in Thailand. PeerJ 11 (e 16572): 154. https://doi.org/10.7717/peerj.16572">Ngernsaengsaruay et al. 2023 b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUHBG" type="etymology">
<paragraph id="C20761E9DB0111C86E38DD04C10154A9">
<heading id="F7B6C469D847E5317090592A76EA8352" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="43B0DA24A7182A7B7DD968C2532EA463">
The specific epithet of
<taxonomicName id="8379C30B1C73BF08EB8E462C4554BB3F" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="8FC9115F6FC9EA22E82222B64D8B1D96" italics="true">Garcinia prainiana</emphasis>
</taxonomicName>
refers to Sir David Prain (18571944), a British botanist, a herbarium curator of the Royal Botanic Garden, Calcutta (18871898), and a director of the Royal Botanic Gardens, Kew (19051922) (
<bibRefCitation id="7D1B67C62CDB70547501E4C2DFFC248B" DOI="10.5962/t.206493" author="Stafleu and Cowan" firstAuthor="Stafleu" refId="B34" refString="Stafleu FA, Cowan RS (1983) Taxonomic Literature: A Selective Guide to Botanical Publications and Collections with Dates, Commentaries and Types Vol. 4 [P Sak]. 2 nd edn., Bohn, Scheltema &amp; Holkema, Utrecht, The Netherlands. https://doi.org/10.5962/t.206493" year="1983">Stafleu and Cowan 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJIBG" type="Vernacular names">
<paragraph id="B562E0ADFA83A6732BB0E79EF7EF79E9">
<heading id="141FF80D6A2544F0871828782B953F7C" reason="title">Vernacular names.</heading>
</paragraph>
<paragraph id="D0F3691363770C25F42B8699E5CD1592">
<emphasis id="13455D6535E6C757F67FDF68D0A1F4D0" bold="true">Chupu</emphasis>
(จ ู ป ู) (Malay-Narathiwat); Cerapu, Chekau, Chepu, Cherapu, Cherpu, Cherupu, Chupak, Chupu, Kechupu, Kecupu, Menchepu, Menchupu (Malay); Button mangosteen (English).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERIBG" type="Uses">
<paragraph id="CD00F37AEAE147ADA8378217A3C37217">
<heading id="1D3E56D66E1DEE1390E90F7740F94A3C" reason="title">Uses.</heading>
</paragraph>
<paragraph id="0B2AD7E3C71A4C6D9314C53338DBFEB5">
<taxonomicName id="DB4815589FE7D29B972EC8F35F8CE1B0" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="E1062BF328C0DABA2483CE7E684757B6" italics="true">Garcinia prainiana</emphasis>
</taxonomicName>
is locally cultivated for its fruits in peninsular
<collectingCountry id="CB6B7497D69B315AEBC060F5832B35A6" name="Thailand">Thailand</collectingCountry>
. The fruits (pericarp and fleshy pulp surrounding the seeds) are edible and have a sour or sweet-sour taste. It is also grown in some botanical gardens as an ornamental plant to provide botanical education.
</paragraph>
<paragraph id="F5E13A52590C040EE3A2FB296CE0482F">
In Peninsular
<collectingCountry id="30461BC0E39F16801495DB4030460E15" name="Malaysia">Malaysia</collectingCountry>
, it is commonly cultivated in village gardens. The ripe fruits are edible and are sometimes used fresh in beverages (
<bibRefCitation id="5D6D1573D556F8FF02AF9023F0D98496" author="Allen" firstAuthor="Allen" refId="B1" refString="Allen BM (1965) Malayan Fruits: An Introduction to the Cultivated Species. Eastern Universities Press, Ltd., Singapore." year="1965">Allen 1965</bibRefCitation>
;
<bibRefCitation id="FCBDBB3748C42B117DBE65179B384C1D" author="Burkill" etAl="et al." firstAuthor="Burkill" refId="B4" refString="Burkill IH, Birtwistle W, Foxworthy FW, Scrivenor JB, Watson JG (1966) A Dictionary of the Economic Products of the Malay Peninsula Vol 1 (A H). Government of Malaysia and Singapore, Ministry of Agriculture and Co-Operatives, Kuala Lumpur, 10631074." year="1966">Burkill et al. 1966</bibRefCitation>
). The pulp of fruits has high antioxidant content of about 91.9 % and vitamin C content of about
<quantity id="436FCA6EB8E1B627C98BA1DFE2AFE0AA" metricMagnitude="-5" metricUnit="kg" metricValue="2.73" unit="mg" value="27.3">27.3 mg</quantity>
per
<quantity id="B7AFD4AFD5B263D47867DBA20290F273" metricMagnitude="-1" metricUnit="kg" metricValue="1.0" unit="g" value="100.0">100 g</quantity>
fresh weight (
<bibRefCitation id="615E43A8EB73E3FFBDFB6C29BAD3569B" author="Azuan and Salma" firstAuthor="Azuan" refId="B2" refString="Azuan A, Salma I (2018) Fruit for the future: cerapu (Garcinia prainiana) [Abstract] in Conference: MARDI Science and Technology Exhibition (MSTE 2008) at Serdang, Selangor." year="2018">Azuan and Salma 2018</bibRefCitation>
). In a traditional Malay recipe, the raw fruits are described as being cooked with dried fish (
<bibRefCitation id="6A4BA25084D50B974B20DBE390B8C2C6" author="Zawiah and Othaman" firstAuthor="Zawiah" refId="B50" refString="Zawiah N, Othaman H (2012) 99 Spesies Buah di FRIM. Institut Penyelidikan Perhutanan Malaysia." year="2012">Zawiah and Othaman 2012</bibRefCitation>
). The wood is used for house building (
<bibRefCitation id="62AF0D5C58DB08F7AEEFEFB196612474" author="Allen" firstAuthor="Allen" refId="B1" refString="Allen BM (1965) Malayan Fruits: An Introduction to the Cultivated Species. Eastern Universities Press, Ltd., Singapore." year="1965">Allen 1965</bibRefCitation>
;
<bibRefCitation id="F9FE22CACF5340C8B741BB2D599196BB" author="Burkill" etAl="et al." firstAuthor="Burkill" refId="B4" refString="Burkill IH, Birtwistle W, Foxworthy FW, Scrivenor JB, Watson JG (1966) A Dictionary of the Economic Products of the Malay Peninsula Vol 1 (A H). Government of Malaysia and Singapore, Ministry of Agriculture and Co-Operatives, Kuala Lumpur, 10631074." year="1966">Burkill et al. 1966</bibRefCitation>
). It is an excellent ornamental plant for use in landscape gardens in parks (
<bibRefCitation id="9FED6E6513616DBA885ED560685FC437" refId="B17" refString="National Parks Flora and Fauna Web (2023) Garcinia prainiana. National Parks Board. https://www.nparks.gov.sg/florafaunaweb" url="https://www.nparks.gov.sg/florafaunaweb" year="2023">National Parks Flora and Fauna Web 2023</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAKBG" type="notes">
<paragraph id="3E3533E52D4452903A92D5DB329FB438">
<heading id="E2AC2EF805DC2B70A5FA2DC40FFEDBE8" reason="title">Notes.</heading>
</paragraph>
<paragraph id="690DAA0F479CE794DE15696C2A048E07">
According to
<bibRefCitation id="D4BA8EC1252381E1A7CBE7B94B807811" DOI="10.7717/peerj.16572" author="Ngernsaengsaruay C &amp; Chanton P &amp; Chaiprasongsuk M &amp; Leksungnoen N" refId="B25" refString="Ngernsaengsaruay C, Chanton P, Chaiprasongsuk M, Leksungnoen N (2023 b) A taxonomic revision of Garcinia section Xanthochymus (Clusiaceae) in Thailand. PeerJ 11 (e 16572): 154. https://doi.org/10.7717/peerj.16572">Ngernsaengsaruay et al. (2023 b</bibRefCitation>
), the male flowers of
<taxonomicName id="FD7BBD26ABA2B0A94E2E9FDE8771C07A" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="9652BF7A5743B3EBF23EE83DE5FBF8FA" italics="true">Garcinia prainiana</emphasis>
</taxonomicName>
were reported to have a small ring-shaped disk surrounding the base of the pistillode. However, in this study, we re-examined the flowers, and a small ring-shaped disk is absent.
</paragraph>
<paragraph id="EE829E900C40C73029CFC1CAE95FA261">
<taxonomicName id="5C7B7F13C0717C4BA262EECE02D369F6" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="A596B861BDF2989344FD6CAEC19CD964" italics="true">Garcinia prainiana</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="EEAB14746603A78E1657EB46A4FAC37F" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="phuongmaiensis">
<emphasis id="DCBA989A38F6D3EDD2771118EC5C40D0" italics="true">G. phuongmaiensis</emphasis>
</taxonomicName>
in having coriaceous, bullate, shiny dark green, subcordate, subamplexicaul leaves with a short petiole; 5 - merous flowers; and numerous stamens, united into 5 antepetalous bundles surrounding a pistillode, but differs in relatively larger habit as a
<quantity id="C40ADC93253E36362C312A28EA0407C5" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" unit="m" value="7.5" valueMax="12.0" valueMin="3.0">312 m</quantity>
tall tree (vs smaller habit, shrubs,
<quantity id="77CAB05A2A29A32CCF833F93BA22A842" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">13 m</quantity>
tall); larger leaves, 12.527.5 ×
<quantity id="5549878B0115E2B5BE3944A3C6A3E0B0" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="11.5" metricValueMin="5.5" unit="cm" value="8.5" valueMax="11.5" valueMin="5.5">5.511.5 cm</quantity>
(vs smaller leaves, 411 ×
<quantity id="7B8847039CC8B3F582006AD60136740C" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" unit="cm" value="3.75" valueMax="5.0" valueMin="2.5">2.55 cm</quantity>
); larger flowers,
<quantity id="6DC7F4B63762971E9F4A45AB9FC40792" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" unit="cm" value="3.25" valueMax="4.0" valueMin="2.5">2.54 cm</quantity>
in diam. (vs smaller flowers, c.
<quantity id="6BA3A5FC962BA70FB0225CC6DD873AEA" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="cm" value="1.0">1 cm</quantity>
in diam.); variable in color of petals: pale yellow, yellowish pink, yellowish red, pinkish red, pink or red (vs white petals); staminodes absent (vs present); unlobed fruits, turning bright yellow, orangish yellow and bright orange when ripe (vs shallowly 34 - lobed fruits, turning bright red when ripe); seeds with pale orange fleshy pulp (vs seeds with white fleshy pulp); and is distributed in Peninsular
<collectingCountry id="6ACD502CFCCA80B9A0ED70AC82B0A1F9" name="Malaysia">Malaysia</collectingCountry>
and Peninsular
<collectingCountry id="FF09EA60EF1C38D9798205425DBB6903" name="Thailand">Thailand</collectingCountry>
(vs Central
<collectingCountry id="A3F2044C0BB68ACDE735C1E75A079ED2" name="Vietnam">Vietnam</collectingCountry>
). The morphological characteristics and distribution of
<taxonomicName id="814B8721B4318043D39ADD3063BE1BFE" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="phuongmaiensis">
<emphasis id="94D827FFFBE122381E378644A9968DBE" italics="true">G. phuongmaiensis</emphasis>
</taxonomicName>
were taken from
<bibRefCitation id="296382A1D274288BFAB7526EEE6E3056" DOI="10.1007/s12225-023-10077-0" author="Tuan DLA &amp; Toyama H &amp; Nguyen XMA &amp; Phan TTN &amp; Truong VB &amp; Dang VS" etAl="et al." firstAuthor="Tuan" issue="2" journalOrPublisher="Kew Bulletin" pagination="189-195" refId="B44" refString="Tuan DLA, Toyama H, Nguyen XMA, Phan TTN, Truong VB, Dang VS (2023) Garcinia phuongmaiensis, a new species of Garcinia sect. Xanthochymus (Clusiaceae) from central Vietnam. Kew Bulletin 78 (2): 189195. https://doi.org/10.1007/s12225-023-10077-0" title="Garcinia phuongmaiensis, a new species of Garcinia sect. Xanthochymus (Clusiaceae) from central Vietnam." volume="78" year="2023">Tuan et al. (2023)</bibRefCitation>
.
</paragraph>
<paragraph id="5681952C3117EE837A91FD503C601440">
<bibRefCitation id="F38418CA0CF1B3E1106E348C5F7D89F4" author="Vesque J" firstAuthor="Vesque" journalOrPublisher="Masson, Paris" refId="B48" refString="Vesque J (1893) Guttiferae. In: de Candolle ALPP, de Candolle ACP (Eds) Monographiae Phanerogamarum, Prodromi Nunc Continuatio, Nunc Revisio Vol. 8. Sumptibus G. Masson, Paris, 669 pp." title="Guttiferae. In: de Candolle ALPP, de Candolle ACP (Eds) Monographiae Phanerogamarum, Prodromi Nunc Continuatio, Nunc Revisio Vol. 8. Sumptibus G." year="1893">Vesque (1893)</bibRefCitation>
placed
<taxonomicName id="7F42FDFD71069E1E6DA93E141912EAB0" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="2B4507668380AE846B71AFC23A93988A" italics="true">Garcinia prainiana</emphasis>
</taxonomicName>
with species of
<emphasis id="2D4DEC1C4E6562029D3C587122B24F8A" italics="true">
G. section
<taxonomicName id="FD17A3E0511D10942DB66AB636C8EE8A" rank="section" section="Xanthochymus">Xanthochymus</taxonomicName>
</emphasis>
(
<emphasis id="71AA359AD654359879B1AD2066E17578" italics="true">subgenus Xanthochymus</emphasis>
), and this placement was followed by
<bibRefCitation id="F9F0394F56E3F9C06F47D23E18F90DD9" author="Jones SW" firstAuthor="Jones" journalOrPublisher="PhD Thesis, University of Leicester and British Museum (Natural History), London" refId="B12" refString="Jones SW (1980) Morphology and Major Taxonomy of Garcinia (Guttiferae). PhD Thesis, University of Leicester and British Museum (Natural History), London, 474 pp." title="Morphology and Major Taxonomy of Garcinia (Guttiferae)." year="1980">Jones (1980)</bibRefCitation>
. However, the flowers of
<taxonomicName id="15426B71418623255929E9200D0AFB36" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="7046DDE7146B2FC7CC3B72EC9CE6895B" italics="true">G. prainiana</emphasis>
</taxonomicName>
have a pistillode and lack receptacular disks and antepetalous appendages, unlike those found in other
<emphasis id="0B8CE8D61D6ED9BA0A232F439722799F" italics="true">
G. section
<taxonomicName id="7C48C8924B93EC96F01689A09FF2EE99" rank="section" section="Xanthochymus">Xanthochymus</taxonomicName>
</emphasis>
species (e. g.,
<taxonomicName id="AD3FB03776D2A75865E630F3FABB0BDF" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="dulcis">
<emphasis id="84F603FFB7F174002514A494A6C5C3F6" italics="true">G. dulcis</emphasis>
</taxonomicName>
,
<taxonomicName id="03F231E0304F8FDA5AD9AC9BB01A8C57" authority="Merr." authorityName="Merr." class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="subelliptica">
<emphasis id="98BCA4FC7DEA1AF8800B379B2FDA64A8" italics="true">G. subelliptica</emphasis>
Merr.
</taxonomicName>
) or other species in “ lineage A ”. These flowers, with staminal phalanges adnate to the petals, and
<taxonomicName id="42975FF9AFEAD85AEA3468F032994FB6" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="F7DCEF5166017A9AAEAF1B833A0350A7" italics="true">G. prainiana</emphasis>
</taxonomicName>
s branching, adaxial, exudate-containing canal pattern agree with the molecular data and support its placement within a subclade (clade 9) of “ lineage B ” with which it shares many features (
<bibRefCitation id="02A938FE55E9BB3BAD19EE4C8E0FE1B3" DOI="10.1086/591990" author="Sweeney" firstAuthor="Sweeney" issue="9" journalOrPublisher="International Journal of Plant Sciences" pagination="1288-1303" refId="B37" refString="Sweeney PW (2008) Phylogeny and floral diversity in the genus Garcinia (Clusiaceae) and relatives. International Journal of Plant Sciences 169 (9): 12881303. https://doi.org/10.1086/591990" title="Phylogeny and floral diversity in the genus Garcinia (Clusiaceae) and relatives." volume="169" year="2008">Sweeney 2008</bibRefCitation>
). More recently,
<bibRefCitation id="B803C02C80A1C24BE52367438A75151D" DOI="10.3897/phytokeys.239.112563" author="Gaudeul M &amp; Sweeney P &amp; Munzinger J" etAl="et al." firstAuthor="Gaudeul" journalOrPublisher="PhytoKeys" pagination="73-105" refId="B9" refString="Gaudeul M, Sweeney P, Munzinger J (2024) An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence. PhytoKeys 239: 73105. https://doi.org/10.3897/phytokeys.239.112563" title="An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence." volume="239" year="2024">Gaudeul et al. (2024)</bibRefCitation>
reported that in addition to molecular data,
<taxonomicName id="D8FB32C700C6B82F253A3554BA50CF93" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="prainiana">
<emphasis id="14456DD275972942C79A5180888E7E65" italics="true">G. prainiana</emphasis>
</taxonomicName>
has morphology that supports its placement into
<emphasis id="541905C0A08C1E4637FBFFA02B154744" italics="true">
G. section
<taxonomicName id="A2A02D587AB701A4145DE7529E4A30DB" rank="section" section="Macrostigma">Macrostigma</taxonomicName>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6PBG" type="material">
<paragraph id="6D31540183A16AB970D81EB31F145315">
<heading id="9B607A49A419F8D6032668BB780FE286" reason="title">Additional specimens examined.</heading>
</paragraph>
<paragraph id="A40876C11C8A05918D8E3A1D181DA86D">
<materialsCitation id="1727240609933420F20F01A318F5D2D7" collectingDate="2019-05-31" collectionCode="BKF" collectorName="C. Ngernsaengsaruay &amp; W. Boonthasak" country="Thailand" location="Chulachomklao Royal Military Academy" municipality="Phrueksaphan Thepparat Botanicical Garden" specimenCount="1" stateProvince="Nakhon Nayok">
<emphasis id="9CCD51A4E9B7D352D62EAD26C899DAB3" bold="true">
<collectingCountry id="A3B67D5D0F0CFF08A7960B32FB0DC968" name="Thailand">Thailand</collectingCountry>
. Central
</emphasis>
<collectingRegion id="4CE22DFAD1CE0EDA81085C97C57D97A3" country="Thailand" name="Nakhon Nayok">Nakhon Nayok</collectingRegion>
[
<collectingMunicipality id="AE05F21B5C77A12D482A49946A14A7F0">Phrueksaphan Thepparat Botanicical Garden</collectingMunicipality>
,
<location id="6D141B79D50FD4ECEEE214EEE7057F98" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:6D141B79D50FD4ECEEE214EEE7057F98" country="Thailand" municipality="Phrueksaphan Thepparat Botanicical Garden" name="Chulachomklao Royal Military Academy" stateProvince="Nakhon Nayok">Chulachomklao Royal Military Academy</location>
, cultivated,
<date id="F7DEE27CC25E121119DA35B798EB023F" value="2019-05-31">
<collectingDate id="E78E2374B3387000986CD597590DFBDD" value="2019-05-31">31 May 2019</collectingDate>
</date>
,
<emphasis id="9316ACBB927795D9A7B961EFBE4D30F3" italics="true">
<collectorName id="EAD7DE8DBE05CF8E357333436E1A6B6E">C. Ngernsaengsaruay</collectorName>
&amp;
<collectorName id="2600F58E340F4224C27AF4AA02200423">W. Boonthasak</collectorName>
G 30-31052019
</emphasis>
(
<collectionCode id="A78D2BD678306EF50C8BFDB576A9C6FE" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D7660596966CE90E131444BA006790DA" collectingDate="2022-02-16" collectionCode="BKF" collectorName="C. Ngernsaengsaruay" country="Thailand" county="Khao Chong Botanical Garden" location="Na Yong District" municipality="Chong Subdistrict" specimenCount="1" stateProvince="Trang">
<emphasis id="62C50E7B58E37B969B58B204A11BBFEA" bold="true">Peninsular</emphasis>
<collectingRegion id="BABF04A3361DED3372C3F9FABC2162A0" country="Thailand" name="Trang">Trang</collectingRegion>
[
<collectingCounty id="A145447880C913B782537EBF95DC69E9">Khao Chong Botanical Garden</collectingCounty>
,
<collectingMunicipality id="3C84F7AC44C4CB9B8D58F2963B5453AE">Chong Subdistrict</collectingMunicipality>
,
<location id="F3F1C67D49E4230417316FCC0727BDC1" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:F3F1C67D49E4230417316FCC0727BDC1" country="Thailand" county="Khao Chong Botanical Garden" municipality="Chong Subdistrict" name="Na Yong District" stateProvince="Trang">Na Yong District</location>
, cultivated,
<date id="1FF36ED7CAC56FE6C4B454FCF394DED5" value="2022-02-16">
<collectingDate id="775750BD75BE3A7AE675B21C46F6CB7A" value="2022-02-16">16 Feb 2022</collectingDate>
</date>
,
<emphasis id="C19D2924E0D0BEFC9A2859721C536BDE" italics="true">
<collectorName id="6A3B6600861903327A13301D66042D6A">C. Ngernsaengsaruay</collectorName>
et al. G 32-16022022
</emphasis>
(
<collectionCode id="EFE020CA1C657B5DC99A821EA902CE6A" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
)]
</materialsCitation>
<materialsCitation id="C50B77AE2AAA61586B8FC5C35385CF3E" collectingDate="2019-11-27" collectionCode="BKF" collectorName="C. Ngernsaengsaruay &amp; G. Sichaikhan" country="Thailand" county="Chulabhorn Phatthana" location="Than To District" municipality="Project" specimenCount="1" stateProvince="Yala">
<collectingRegion id="98AE5F0C48D6A9BC7D2D8E93E1B225CE" country="Thailand" name="Yala">Yala</collectingRegion>
[
<collectingCounty id="35F67D18F6230503065465354FD3D77E">Chulabhorn Phatthana</collectingCounty>
7
<collectingMunicipality id="2C9E34317BD03C6D44919451CE7D1D6A">Project</collectingMunicipality>
,
<location id="FB05B1B62266B3673A836A7F7CE88D13" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:FB05B1B62266B3673A836A7F7CE88D13" country="Thailand" county="Chulabhorn Phatthana" municipality="Project" name="Than To District" stateProvince="Yala">Than To District</location>
, near waterfall,
<date id="DCF07929A213A8FEAF350EBBB0D17230" value="2019-11-27">
<collectingDate id="FB034CD643C68149B6025E6AC4307C95" value="2019-11-27">27 Nov 2019</collectingDate>
</date>
,
<emphasis id="A0D77073CD2E99309749906F21A07546" italics="true">
<collectorName id="71317EBE1B44FFD28CDE8DA516742B2F">C. Ngernsaengsaruay</collectorName>
&amp;
<collectorName id="01104C379F628E2B1CE573616A1D00A0">G. Sichaikhan</collectorName>
G 31-27112019
</emphasis>
(
<collectionCode id="8C467EF1B8068033FA4B87C025F084D3" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
)]
</materialsCitation>
<materialsCitation id="EEC0761FC42048A168EC91D1C8778EDF" collectingDate="2002-04-20" collectingDateMax="2005-09-22" collectingDateMin="2002-04-20" collectionCode="BKF, QBG" collectorName="M. Poopath &amp; Hala-Bala Wildlife Sanctuary &amp; C. Niyomdham &amp; P. Puudjaa &amp; U. Upho" country="Thailand" county="Sanctuary" location="Lo Chut Subdistrict" municipality="Ban Bala" specimenCount="1" specimenCount-male="1" stateProvince="Narathiwat">
<collectingRegion id="AD78D72F943EC7DB48C631CB271F5B94" country="Thailand" name="Narathiwat">Narathiwat</collectingRegion>
[Hala-BalaWildlife
<collectingCounty id="64E2D6C34C1F654F07B40B58AB2D0E50">Sanctuary</collectingCounty>
,
<collectingMunicipality id="BC66AF5F55F2651B67BA5B6470954C3A">Ban Bala</collectingMunicipality>
,
<location id="C729EB135BE48BD9730254839C7CCF0F" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:C729EB135BE48BD9730254839C7CCF0F" country="Thailand" county="Sanctuary" municipality="Ban Bala" name="Lo Chut Subdistrict" stateProvince="Narathiwat">Lo Chut Subdistrict</location>
,
<location id="365706DEEF40E35B3FF6AB80F1CB47F0" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:365706DEEF40E35B3FF6AB80F1CB47F0" country="Thailand" county="Sanctuary" municipality="Ban Bala" name="Waeng District" stateProvince="Narathiwat">Waeng District</location>
,
<specimenCount id="D7BFCFC4ADA41AAF53F3AE09B74A8CFA" count="1" type="male"></specimenCount>
fl.,
<date id="5E17374D1E37E57365EB1AEFAFF79589" value="2005-05-13">
<collectingDate id="E7D629F00EC55DC72C13A2622873DBCB" value="2005-05-13">13 May 2005</collectingDate>
</date>
(as
<taxonomicName id="5EED286DA311ED90730D9C840570D526" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="C8E3E6F2457F69185885AFA19B43CDF9" italics="true">Garcinia</emphasis>
sp.
</taxonomicName>
),
<emphasis id="8A9ED86BF8336D6B61A1B29FE30A7E63" italics="true">
<collectorName id="A76504F244E579EB639B262227418EA4">M. Poopath</collectorName>
274
</emphasis>
(
<collectionCode id="2A25CAABE0878A32F3164F414FC2407A" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
) •
<collectorName id="4FFDB8D291AB63E9CC3B0DFB0F557B9E">Hala-Bala Wildlife Sanctuary</collectorName>
,
<location id="97EB502011B2E3365470AA3520092D6E" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:97EB502011B2E3365470AA3520092D6E" country="Thailand" county="Sanctuary" municipality="Ban Bala" name="Waeng District" stateProvince="Narathiwat">Waeng District</location>
, fr.,
<date id="9CB613E66932085B044B20C1E5972C62" value="2005-09-22">
<collectingDate id="359F22FCA797C547B06C75C11D4BAF75" value="2005-09-22">22 Sep 2005</collectingDate>
</date>
,
<emphasis id="2970D40992AAAA0EC44077BF04E31773" italics="true">
<collectorName id="D7DF12E24F2C3B4CCA8D74D8BB76E176">C. Niyomdham</collectorName>
&amp;
<collectorName id="5167BFB4FECC99D771F5BC6CB846E475">P. Puudjaa</collectorName>
7593
</emphasis>
(
<collectionCode id="D5AAB3C13602077FA592758D85EEA151" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
) •
<location id="6E426A1151FB0A17D3A1E11AFB0912D8" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:6E426A1151FB0A17D3A1E11AFB0912D8" country="Thailand" county="Sanctuary" municipality="Ban Bala" name="Su-ngai Kolok District" stateProvince="Narathiwat">Su-ngai Kolok District</location>
, fr.,
<date id="65785BF682D1CAD9E7C0F1C8B11D0D56" value="2002-04-20">
<collectingDate id="60F49FB556E9C15964F50B809F5B184B" value="2002-04-20">20 Apr 2002</collectingDate>
</date>
,
<emphasis id="5C61B967BB969098EC4FB114C2FEB353" italics="true">
<collectorName id="E4772C18F34175895D4D4DCF077A91EE">U. Upho</collectorName>
556
</emphasis>
(
<collectionCode id="8A48088330A03AE159BE82B14D74F0D1" country="Thailand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13640" lsid="urn:lsid:biocol.org:col:13640" name="Queen Sirikit Botanic Garden" type="">QBG</collectionCode>
) •
<location id="B1DB148C658D08C3B357F60C423FAAC8" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:B1DB148C658D08C3B357F60C423FAAC8" country="Thailand" county="Sanctuary" municipality="Ban Bala" name="Su-ngai Kolok District" stateProvince="Narathiwat">Su-ngai Kolok District</location>
, fl., cultivated,
<date id="36FF380B9D00FA39A6DE63E1578DE9BB" value="2003-05-20">
<collectingDate id="4E1C855CB6667C741373A613B139E792" value="2003-05-20">20 May 2003</collectingDate>
</date>
,
<emphasis id="8BEC3AFA39F414B3D7E6CE607B9DE7BA" italics="true">
<collectorName id="6B2F57C5E4D778DEB6A97AB946CB98DD">U. Upho</collectorName>
550
</emphasis>
(
<collectionCode id="1CE189EBD56E53B7D05A29641361F75A" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
)]
</materialsCitation>
.
</paragraph>
<paragraph id="C78E5F835D9344DD56F99FC0D4ABB080">
<emphasis id="D80B2103E77A3001674640413FB636ED" bold="true">
Peninsular
<materialsCitation id="C5F2BF7B9DA55EB10510652525638844" collectingDate="1896-12" collectingDateMax="1971-10-30" collectingDateMin="1896-12" collectorName="C. Curtis" country="Malaysia" location="Kwala Dipang" specimenCount="1" specimenCount-male="1" stateProvince="Perak">
<collectingCountry id="6BE728672A07843E4F3949D17F85DB4D" name="Malaysia">Malaysia</collectingCountry>
<collectingRegion id="3A91A6524AC2C00179D9478524573457" country="Malaysia" name="Perak">Perak</collectingRegion>
[
<location id="FC3415AE256EBE8F5F0014A6AEA3EF28" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:FC3415AE256EBE8F5F0014A6AEA3EF28" country="Malaysia" name="Kwala Dipang" stateProvince="Perak">Kwala Dipang</location>
,
<specimenCount id="66E57278B7F75CEB04C2D7FD07FA6AF5" count="1" type="male"></specimenCount>
fl.,
<date id="5FACCB9F0F94EF8771A0A114E338D380" value="1896-12">
<collectingDate id="5AE23B081B7986E85361C3527B67932E" value="1896-12">Dec 1896</collectingDate>
</date>
,
<emphasis id="DD09CF1762CCB08E63148E582F00DCBD" italics="true">
<collectorName id="0B314E1B06493740B4720B592B8B28BE">C. Curtis</collectorName>
3273
</emphasis>
(K [K 000677679]) • Kg Kepayang near Ipoh, fr.,
<date id="D41BD64A6428229D4A58FD7E474D2236" value="1971-10-30">
<collectingDate id="A45671CD56DCBD9AA84B3169AFB721B5" value="1971-10-30">30 Oct 1971</collectingDate>
</date>
,
<emphasis id="2C5719282BB0DD0B7C8920EB71D51200" italics="true">Syed Abu Bakar Kep. FRI 20440</emphasis>
(L [L 2417220])]
</materialsCitation>
</emphasis>
<materialsCitation id="34F4EDFC794DCC25865DDE310BE4E46D" collectingDate="1967-07-14" collectingDateMax="1994-06-02" collectingDateMin="1967-07-14" collectionCode="BKF, PSU" collectorName="T. C. Whitmore Kep. &amp; H. S. Loh Kep. &amp; K. M. Kochummen Kep. &amp; B. C. Stone &amp; M. Sidek &amp; A. Latiff &amp; A. Noorsiha &amp; H. Husmady" country="Malaysia" county="Su-ngai Bertam" location="Path" municipality="Kuala Mensum" specimenCount="1" specimenCount-female="1" stateProvince="Pahang">
<collectingRegion id="241AC0C37E1A09D48FFDFBB2DB6B0177" country="Malaysia" name="Pahang">Pahang</collectingRegion>
[
<collectingCounty id="315AC1CE11CC488F214E73C3F32F2ED1">Su-ngai Bertam</collectingCounty>
at
<collectingMunicipality id="F257803CF0AE3030872F4F1A0BF4FC64">Kuala Mensum</collectingMunicipality>
, fl.,
<date id="1AD362448E43A23C77AC5AFE626787AF" value="1971-06-02">
<collectingDate id="FC5DDFAAB2C6D94D51D78B50FED74A09" value="1971-06-02">2 Jun 1971</collectingDate>
</date>
,
<emphasis id="472BEC9BF6A74AE14F4F7D61FBFB1C5B" italics="true">
<collectorName id="4026F90B63A97DE214770CD0BEDFB292">T. C. Whitmore Kep.</collectorName>
FRI 20091
</emphasis>
(L [L 2417222]) •
<location id="61CAB44E7C845AAC376D6DD3B4C28FA4" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:61CAB44E7C845AAC376D6DD3B4C28FA4" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Path" stateProvince="Pahang">Path</location>
leading to
<location id="7968B9F9692D406B6FFB0B692B506D44" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:7968B9F9692D406B6FFB0B692B506D44" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Kuala Mensum" stateProvince="Pahang">Kuala Mensum</location>
from
<location id="3B4B1FA3D8DCF5140D4DCD101AE7CBAC" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:3B4B1FA3D8DCF5140D4DCD101AE7CBAC" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Boh Tea" stateProvince="Pahang">Boh Tea</location>
,
<location id="3CD3F9439FAE3E6FB65BDB0C6BAB6BCC" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:3CD3F9439FAE3E6FB65BDB0C6BAB6BCC" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Cameron Highlands" stateProvince="Pahang">Cameron Highlands</location>
, fr.
<date id="E0F944652CDF0BBA0F076033BBC8F642" value="1971-09-24">
<collectingDate id="06483F585175CE98DB4DB45B09F61A1B" value="1971-09-24">24 Sep 1971</collectingDate>
</date>
,
<emphasis id="5922F8F80915BF0B4819507C35CE9A34" italics="true">
<collectorName id="1610F429F6800BCCDD70A63DE3051657">H. S. Loh Kep.</collectorName>
FRI 19187
</emphasis>
(L [L 2417221]) •
<location id="B70885D13AD1431C86F52707AF529FE7" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:B70885D13AD1431C86F52707AF529FE7" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Cameron Highlands Road" stateProvince="Pahang">Cameron Highlands Road</location>
, fr.,
<date id="7FC8A2E0E7402C40066F666E18442F33" value="1982-01-18">
<collectingDate id="A9B2F52C7B608151948312E5398903F9" value="1982-01-18">18 Jan 1982</collectingDate>
</date>
,
<emphasis id="79985CEE0C3F4AE50ED36CBAB313E14E" italics="true">
<collectorName id="A544BA0430B1C39DFBCD5D7D2F5A8B36">K. M. Kochummen Kep.</collectorName>
FRI 29377
</emphasis>
(L [L 2417225]) •
<collectingRegion id="94FF4B5F7A2D80A581BA433AC198B8F5" country="Malaysia" name="Kelantan">Kelantan</collectingRegion>
[
<quantity id="1B910590E3D667587109331882783343" metricMagnitude="2" metricUnit="m" metricValue="8.04672" unit="mi" value="0.5">0.5 mile</quantity>
east of
<location id="25A0E8DA177E989A550F693EAF73CA21" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:25A0E8DA177E989A550F693EAF73CA21" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Gua Musang" stateProvince="Pahang">Gua Musang</location>
, fr.,
<date id="645ADDB19A360450790940121395515F" value="1967-07-14">
<collectingDate id="4DB2A3D8C83E165C89AD874279DD4309" value="1967-07-14">14 Jul 1967</collectingDate>
</date>
,
<emphasis id="30CC6430E1E147850E82062E04459911" italics="true">
<collectorName id="C9981D1AF9B219008B18B6900EF17078">T. C. Whitmore Kep.</collectorName>
FRI 4018
</emphasis>
(L [L 2417226]) •
<location id="750721BC4C5B81AC472DA6AEA7D93CB0" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:750721BC4C5B81AC472DA6AEA7D93CB0" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Su-ngai Lebir" stateProvince="Pahang">Su-ngai Lebir</location>
, below
<location id="B3A75E8C97EEFA6061D9105ACFDE3FE0" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:B3A75E8C97EEFA6061D9105ACFDE3FE0" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Kuala Relai" stateProvince="Pahang">Kuala Relai</location>
at
<location id="88148C9E9FFC8E6D2AE4ED984998B1CF" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:88148C9E9FFC8E6D2AE4ED984998B1CF" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Jentah" stateProvince="Pahang">Jentah</location>
, fl.,
<date id="8EB8AE4CC7439CEB4FE403426D6A64CF" value="1976-04-24">
<collectingDate id="109BB8F6498B5DB89B25AA6CC0656009" value="1976-04-24">24 Apr 1976</collectingDate>
</date>
,
<emphasis id="8B3879FCBE85BD40FD5AC80840EC416A" italics="true">
<collectorName id="57E35F13F74BA82A0AE9F0AE4BD8E423">B. C. Stone</collectorName>
&amp;
<collectorName id="07D6F8F3BA54723BFFD530F83AEF1B84">M. Sidek</collectorName>
12426
</emphasis>
(
<collectionCode id="465CE3818B7FF5B60512F68209468DCA" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
, L [L 2417224], US [US 02961246]) •
<location id="57F5DD061D49D7F7E23E9605B5CB8742" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:57F5DD061D49D7F7E23E9605B5CB8742" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Su-ngai Long" stateProvince="Pahang">Su-ngai Long</location>
off
<location id="834927D25927C944DB8470524EA47A61" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:834927D25927C944DB8470524EA47A61" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Su-ngai Pergau" stateProvince="Pahang">Su-ngai Pergau</location>
,
<location id="F61A031087746502B02F7893349C36F7" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:F61A031087746502B02F7893349C36F7" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Jeli" stateProvince="Pahang">Jeli</location>
, fr.,
<date id="19B6F407F99993050526E4C6EFCBE872" value="1986-09-26">
<collectingDate id="4155E28335DB6029ADC9B4C30B5643AC" value="1986-09-26">26 Sep 1986</collectingDate>
</date>
,
<emphasis id="D07CF7654987369ADF5D59CB914C95D7" italics="true">
<collectorName id="5B74F9382318CB772E502C397D2ACEDF">A. Latiff</collectorName>
et al. ALM 1856
</emphasis>
(L [L 3806490],
<collectionCode id="E94D707A4923D499BC4535977CF5A5F7" country="USA" name="Portland State University, Vertebrate Biology Museum" type="Museum">PSU</collectionCode>
) •
<location id="7F95E1CA39C1E3243A3E0A39B4CA5A67" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:7F95E1CA39C1E3243A3E0A39B4CA5A67" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Ketam" stateProvince="Pahang">Ketam</location>
,
<location id="69B580395D0A6EB445A60817909A754B" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:69B580395D0A6EB445A60817909A754B" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Cicar Tinggi" stateProvince="Pahang">Cicar Tinggi</location>
,
<location id="297141900D255186A72CD59E7D27B17F" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:297141900D255186A72CD59E7D27B17F" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Kampung Bata" stateProvince="Pahang">Kampung Bata</location>
,
<location id="51F509036B6707D971EFAB5B99BF365A" LSID="urn:lsid:plazi:treatment:6D5026F191765A6DB8F6C721079CB3DC:51F509036B6707D971EFAB5B99BF365A" country="Malaysia" county="Su-ngai Bertam" municipality="Kuala Mensum" name="Pasir Mas" stateProvince="Pahang">Pasir Mas</location>
,
<specimenCount id="87EB63A3CBC706509ACC7EF605F5BE0D" count="1" type="female"></specimenCount>
fl. &amp; fr.,
<date id="EB3C806424EEA67A8D2BD897A7706050" value="1992-08-01">
<collectingDate id="404FB1AE8DD351C894DDAB47DD51567C" value="1992-08-01">1 Aug 1992</collectingDate>
</date>
,
<emphasis id="0749B38F0EA7F22DF05DBDCECF2524FA" italics="true">
<collectorName id="54A669254CEF2C450BAAD2785E30E0CD">A. Noorsiha</collectorName>
et al. Kep. FRI 39214
</emphasis>
(L [L 3878683]) • Pasir Putih, fl.,
<date id="5D81A2FC3121C0E4D975C9F23257DB4E" value="1992-10-23">
<collectingDate id="774CB934B6F295366E9761C6155CD69B" value="1992-10-23">23 Oct 1992</collectingDate>
</date>
,
<emphasis id="EA107CC35C74D3040DF265ED825EDEFF" italics="true">
<collectorName id="F9602BBB975E971F7A3B2756835D76EA">H. Husmady</collectorName>
et al. Kep. FRI 39551
</emphasis>
(L [L 3806959]) • near Brooke Camp, Gua Musang, fl.,
<date id="E935D68E3B6A0852C0C5AF1D9178264C" value="1994-06-02">
<collectingDate id="519CBE34F8A31AEB53DB2EBF19787C93" value="1994-06-02">2 Jun 1994</collectingDate>
</date>
,
<emphasis id="60953E72809FC1F9B4C6171589B5BA28" italics="true">
<collectorName id="C65F399A19C30467A7924D17EF711FFF">H. Husmady</collectorName>
et al. Kep. FRI 41841
</emphasis>
(L [L 2417223])].
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,549 @@
<document id="30EC331F81E49D568ACD73E383E747EF" ID-DOI="10.3897/zse.100.132111" ID-ZooBank="B39508EB-6EB2-4CC0-88D8-B3E87BDCED02" ID-publisher-id="132111" URI-arpha="6D90141D-221E-5A1C-82D3-04B8878C6781" URI-zoobank="http://zoobank.org/B39508EB-6EB2-4CC0-88D8-B3E87BDCED02" article-type="research-article" checkinTime="1731000560999" checkinUser="pensoft" docAuthor="Nguyen, Anh D., Nguyen, Lien T. P. &amp; Korsós, Zoltán" docDate="2024" docId="6D56422D09F954BE972DB1DD2A1AE4AA" docLanguage="en" docName="Zoosystematics and Evolution 100(4): 1515-1541" docOrigin="Zoosystematics and Evolution 100 (4)" docSource="https://zse.pensoft.net/article/132111/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Pseudodesmus karstomus Nguyen, Nguyen &amp; Korsós, 2024, sp. nov." docType="treatment" docUuid="0EE484F0-C29F-439C-B717-5478C9838FEF" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="1541" masterDocId="6D90141D221E5A1C82D304B8878C6781" masterDocTitle="A review of the millipede genus Pseudodesmus Pocock, 1887 (Diplopoda, Platydesmida, Andrognathidae) from Vietnam, with descriptions of five new species and notes on its phylogeny" masterLastPageNumber="1541" masterPageNumber="1515" pageNumber="1515" updateTime="1731001090142" updateUser="ExternalLinkService">
<mods:mods id="2EE975CECAB32CDF7C71223AFCCB90CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="385127B7FEEA1AA73CE3F1734A0D5D72">
<mods:title id="4D12D7F3DF136FAD30DB16B28654160D">A review of the millipede genus Pseudodesmus Pocock, 1887 (Diplopoda, Platydesmida, Andrognathidae) from Vietnam, with descriptions of five new species and notes on its phylogeny</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="4AB928A0A09CC74FCE8113F8D28D3D82">Nguyen, Anh D.</mods:namePart>
<mods:nameIdentifier id="B651A2DBE2F9B4F1F77089A93C85C9E9" type="ORCID">0000-0001-9273-0040</mods:nameIdentifier>
<mods:affiliation id="858E37A54BFFDC0F8E2399D9A6A7BFFC">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:name id="A2D8B6F374B9D6B0C3C6DC534EB98462" type="personal">
<mods:role id="909F55BB243F507E8031352A7F9341BD">
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</mods:role>
<mods:namePart id="475A5F77543F8CB8F6E7F3D1945E5D0C">Nguyen, Lien T. P.</mods:namePart>
<mods:affiliation id="EDFAFE49F2E626EE2FE656B3E1C9F0BE">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="8F7AA8BF29C28E15561663C03C657546">Korsós, Zoltán</mods:namePart>
<mods:nameIdentifier id="09C84CB85CC135360CA20BA3C8BBC545" type="ORCID">0000-0003-1545-5086</mods:nameIdentifier>
<mods:affiliation id="CE4B5A50C59A088534249F5B048C03FA">University of Veterinary Medicine Budapest, István u. 2, 1078 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6909935528D9723B343D6A332AC340EC">text</mods:typeOfResource>
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<mods:part id="EAD1C83BC6A1E64893C00E778C548C3F">
<mods:date id="1B4CB2548D3B8D0AFFB24BB4C2779ACC">2024</mods:date>
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<treatment id="6D56422D09F954BE972DB1DD2A1AE4AA" ID-DOI="http://doi.org/10.5281/zenodo.14052229" ID-Zenodo-Dep="14052229" ID-arpha="6D56422D-09F9-54BE-972D-B1DD2A1AE4AA" ID-zoobank="https://zoobank.org/0EE484F0-C29F-439C-B717-5478C9838FEF" LSID="urn:lsid:zoobank.org:act:0EE484F0-C29F-439C-B717-5478C9838FEF" LSID-ZBK="urn:lsid:zoobank.org:act:0EE484F0-C29F-439C-B717-5478C9838FEF" httpUri="http://treatment.plazi.org/id/6D56422D09F954BE972DB1DD2A1AE4AA">
<subSubSection id="9329746EA4831D4C2159C9AA23123282" type="nomenclature">
<paragraph id="2DFC8CCB4C77ACC4093B335ABC432FE0">
<taxonomicName id="7A528E4A821BC1CC8D9659DE1E317B43" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Playtdesmida" phylum="Arthropoda" rank="species" species="karstomus" status="sp. nov.">
<emphasis id="20460A0B4D8948362EE72DB457ACB47E" italics="true">Pseudodesmus karstomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F573D6F9D558A5735F59B5967E6A2BC5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FE7448F9488F27F37E5E301B76ED3005" type="description">
<paragraph id="8956875E01413379863D77ADF3EE2841">
<figureCitation id="95EC96A4ABE051AC03147CCE37C8A446" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">Figs 11</figureCitation>
,
<figureCitation id="B474B6D2199AABA8CD57B2B6D581F605" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12</figureCitation>
,
<figureCitation id="A2254B38B932ECD4E49F5ECDAA8E5DB7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), gonopods. A. Left anterior gonopods, anterior view. B. Right posterior gonopods, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure13" httpUri="https://binary.pensoft.net/fig/1169903">13</figureCitation>
,
<figureCitation id="0405581F6982F672178F1B803C174F4D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), gonopods. SEM. A, B. Posterior gonopods, ventral view (A), anterior view (B); C, D. Anterior gonopods, ventral view (C), anterior view (D); E. Apical stylets of posterior gonopod, ventral view; F. Distal part of anterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure14" httpUri="https://binary.pensoft.net/fig/1169904">14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EORAG" type="material">
<paragraph id="A63002DC91BE83F86A0A97D82D23532E">
<heading id="38E3E3F40C2853A2FB907DC1C96730C0" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="7CE01DDB2E37BC345DE95FADBB53F38F">
<materialsCitation id="3E4854B71C86EC8B44147F6F04731CE3" collectingDate="2021-11-01" collectorName="A. D. Nguyen" country="Vietnam" county="Tra Linh District" latitude="22.7588" location="Thang Hen Lake" longLatPrecision="7" longitude="106.2958" municipality="Quoc Toan" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="holotype">
<emphasis id="8529C6F224FA19327AEF68A305054740" bold="true" italics="true">
<typeStatus id="772AD2B87D20CE2F492629EDB3893E30" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="44C40036709AC8F2C22341BB70C4B2B4" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="AB26BEDBFA7D7775A1CEF96C8312CBA9" count="1" type="male">1 male</specimenCount>
;
<collectingRegion id="45982E7E38FBB33761428E79AFD01D8C" country="Vietnam" name="Cao Bang">Cao Bang Province</collectingRegion>
,
<collectingCounty id="D5BE0629038D51DAE3F6534757C8F1EB">Tra Linh District</collectingCounty>
,
<collectingMunicipality id="DFE874DB9EE7CFC8FE412ECCAF0AD730">Quoc Toan</collectingMunicipality>
commune,
<location id="E32FEDD7CD3F712E7A1E0D0F705C7A27" LSID="urn:lsid:plazi:treatment:6D56422D09F954BE972DB1DD2A1AE4AA:E32FEDD7CD3F712E7A1E0D0F705C7A27" country="Vietnam" county="Tra Linh District" latitude="22.7588" longLatPrecision="7" longitude="106.2958" municipality="Quoc Toan" name="Thang Hen Lake" stateProvince="Cao Bang">Thang Hen Lake</location>
, near
<location id="7C6CFCB9FACDC4E2BB7048805D67B485" LSID="urn:lsid:plazi:treatment:6D56422D09F954BE972DB1DD2A1AE4AA:7C6CFCB9FACDC4E2BB7048805D67B485" country="Vietnam" county="Tra Linh District" latitude="22.7588" longLatPrecision="7" longitude="106.2958" municipality="Quoc Toan" name="Ham Huong cave" stateProvince="Cao Bang">Ham Huong cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3RAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.295800,22.758800]}">
<geoCoordinate id="E8329B83ED6C84844869355C4FF7C306" degrees="22.7588" direction="north" orientation="latitude" precision="5" value="22.7588">22.7588 ° N</geoCoordinate>
,
<geoCoordinate id="8A4B0D0682F1BE80D9D7D4100FF5A3D9" degrees="106.2958" direction="east" orientation="longitude" precision="5" value="106.2958">106.2958 ° E</geoCoordinate>
</named-content>
,
<date id="6EA8CCA8BB9BA1B8EC0DC9195FDF761F" value="2021-11-01">
<collectingDate id="118F9523A54507E30D01AFD14DC146FC" value="2021-11-01">1 Nov. 2021</collectingDate>
</date>
;
<collectorName id="FB80481174756F9821EBEEFE4AAAB92A">A. D. Nguyen</collectorName>
leg.;
<abbrev content-type="institution" id="ABBRID0EBSAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
959 H.
</materialsCitation>
</paragraph>
<paragraph id="5DC6677705EE30A853BACD822C954B52">
<materialsCitation id="71D084FFC405284DEDD128B602001A95" collectingDate="2021-11-01" collectorName="A. D. Nguyen" country="Vietnam" county="Tra Linh District" latitude="22.7588" location="Thang Hen Lake" longLatPrecision="7" longitude="106.2958" municipality="Quoc Toan" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="paratype">
<emphasis id="831EF6CA3767CB4A6FEB05D94F8A49AB" bold="true" italics="true">
<typeStatus id="2E975F5AAD49012FFFA2F7B0F7FB7E28" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="832E1F3FE15BB26D0747C37C2B7BF32B" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="EDE05510A607441A424B7377E8DD2455" count="1" type="male">1 male</specimenCount>
,
<specimenCount id="29ADE50015246167EE7CBAF23AAC4483" count="1" type="female">1 female</specimenCount>
same data as for the holotype;
<abbrev content-type="institution" id="ABBRID0ENSAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
959 P
</materialsCitation>
<materialsCitation id="B02E4621A76848B980D7CBD24195F648" collectingDate="2021-11-01" collectionCode="HNHM" collectorName="A. D. Nguyen" country="Vietnam" county="Tra Linh District" latitude="22.7588" location="Thang Hen Lake" longLatPrecision="7" longitude="106.2958" municipality="Quoc Toan" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang" typeStatus="paratype">
<specimenCount id="6613FD7821C312DD258B62388042BE68" count="1" type="male">1 male</specimenCount>
; same data as for the holotype;
<named-content id="F89C34703A87780FE1600A6C4D7618D5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum-termeszettudomanyi-muzeum" xlink_title="Department of Zoology, Hungarian Natural History Museum, Budapest, Hungary">
<collectionCode id="9EFF3855CED31A9391769B595091B113" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXSAG" type="diagnosis">
<paragraph id="9D1C30B81E9B8FB1DAFB2805948957E3">
<heading id="7547C4B704288BBBC1FF7ABDE469ED2A" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="1E2F432CF1E13F717C37D75BB7365550">Terga and base of paraterga marbled yellowish brown with some blackish tubercles on several body rings. Head triangular-shaped, slightly narrower than collum. Collum stout strongly bilobed, anteriorly excavated, with small 2 bumps in the middle, densely setose. Midbody metazonae with two rows of tubercles, anterior row with 78 + 7 8 tubercles, posterior one with 2 + 2 tubercles. Posterior gonopods 6 - segmented, with two apical stylets and two lateroapical stylets.</paragraph>
</subSubSection>
<subSubSection id="SECID0E3SAG" type="diagnosis">
<paragraph id="B11C67A921AB71BEB2AFAA81108CC0A4">
<heading id="B672B96E17A3055ADA0D14DFE6FD2CB8" reason="title">Diagnosis remarks.</heading>
</paragraph>
<paragraph id="C2ED2D82E8B8D4A475B995B2EB00B2AA">
The new species is easily distinguished from
<taxonomicName id="2D38FDFC791FB7373A7C9684D6B180A7" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="camptotrichus">
<emphasis id="68F9B49D80374B7E0D4AFDC5C913F735" italics="true">P. camptotrichus</emphasis>
</taxonomicName>
by coloration (marbled yellowish brown with some blackish tubercles vs earth brown), larger size (length:
<quantity id="660B79FE8A4B8D2FBD46DA7C5252ED6F" metricMagnitude="-2" metricUnit="m" metricValue="1.864" unit="mm" value="18.64">18.64 mm</quantity>
vs
<quantity id="64C44B93E6D45066B268C5CE7C9A3A38" metricMagnitude="-2" metricUnit="m" metricValue="1.4" unit="mm" value="14.0">14 mm</quantity>
; width:
<quantity id="78AD7A463891FCFCDE115FC4E03DD998" metricMagnitude="-3" metricUnit="m" metricValue="2.74" unit="mm" value="2.74">2.74 mm</quantity>
vs 2.0 mm), number of metazonal tubercles (two rows of 78 + 7 8 and 2 + 2 tubercles vs two rows of 6 + 6 to 9 + 9 each row).
</paragraph>
<paragraph id="A1A94E9B1B722E93BC4A4092783825B0">
It also differs from
<taxonomicName id="592A8A54AE02D32FF868834ED5B9C8AC" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="D3E62BE36FB6D3B13A228B2E6F207125" italics="true">P. variegatus</emphasis>
</taxonomicName>
in having smaller size (length:
<quantity id="3BE1FF6D457BF75FEBCD27161CEC7456" metricMagnitude="-2" metricUnit="m" metricValue="1.864" unit="mm" value="18.64">18.64 mm</quantity>
vs 32.0 mm; width:
<quantity id="222B655EF0AE91CAAA26BE1963E3FFD6" metricMagnitude="-3" metricUnit="m" metricValue="2.74" unit="mm" value="2.74">2.74 mm</quantity>
vs
<quantity id="796DE7C1D14B5065E966CC475E84FC18" metricMagnitude="-3" metricUnit="m" metricValue="5.8" unit="mm" value="5.8">5.8 mm</quantity>
), number of metazonal tubercles (two rows of 78 + 7 8 and 2 + 2 tubercles vs two rows of 815 + 8 15 and 57 + 5 7).
</paragraph>
<paragraph id="4FD2458CCD4F8999EB83753F47CFBBCD">
<taxonomicName id="7B188FFC974D5758D9A1E6DA1BB9EB54" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="karstomus" status="sp. nov.">
<emphasis id="DF987538AEC8C8B29A578C5479F99E9C" italics="true">Pseudodesmus karstomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3E26080C925F428FF1684D66D5E850C4" rank="species">sp. nov.</taxonomicNameLabel>
is differentiated from two above new species,
<taxonomicName id="3A31ADBCDB6265B2FB4A3A62E32458DC" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
<emphasis id="8A0EE0AE984605EDB07CE9198D5278CC" italics="true">P. bidoup</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29852890B51DEC4DA83F4E8AE25E320" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="5B894C1AE37539ADBC42BA97EAEC5529" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov">
<emphasis id="294E573DEC7D8F6F90ADC33029CF7501" italics="true">P. ngoclinh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C08AB435E6B20D5D823DCED683039C46" rank="species">sp. nov</taxonomicNameLabel>
in body size and number of gonopod stylets. It shares with
<taxonomicName id="76A617D95E6D270D1C6A122E38376A52" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
<emphasis id="3AAC3A96B16A1F33449F6BA6D8C73FEE" italics="true">P. bidoup</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE8D58DE7CEE1D549F0C3CD927DC3993" rank="species">sp. nov.</taxonomicNameLabel>
a posterior gonopod with four distal stylets, but two species are distinguished by body size (length:
<quantity id="D0DF9054CA2157A0C41F30FE5F860DD0" metricMagnitude="-2" metricUnit="m" metricValue="1.862" unit="mm" value="18.62">18.62 mm</quantity>
vs
<quantity id="250EA9297BDCEBE3F69F2A49FC142A07" metricMagnitude="-3" metricUnit="m" metricValue="9.25" unit="mm" value="9.25">9.25 mm</quantity>
; width:
<quantity id="24CD85022A5AC63317D6F1026EA54A3C" metricMagnitude="-3" metricUnit="m" metricValue="4.05" unit="mm" value="4.05">4.05 mm</quantity>
vs
<quantity id="54C8F5BC87CFA13A230DD2E30608F535" metricMagnitude="-3" metricUnit="m" metricValue="1.63" unit="mm" value="1.63">1.63 mm</quantity>
).
<taxonomicName id="9663C58C151CC44ABE94B7867EB3E627" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="karstomus">
<emphasis id="500FD55644B47C4A1AD123EDCA8C31FF" italics="true">Pseudodesmus karstomus</emphasis>
</taxonomicName>
is also larger than
<taxonomicName id="5AF072B40AF5DC7F0F3830E8CDE2E397" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh">
<emphasis id="B687080DE8811BD77B460F97F7595A32" italics="true">P. ngoclinh</emphasis>
</taxonomicName>
(length:
<quantity id="D9E00DF5DB81FB11180AB6B9D534D8B3" metricMagnitude="-2" metricUnit="m" metricValue="1.862" unit="mm" value="18.62">18.62 mm</quantity>
vs
<quantity id="BACAF061B544E57D067C2B93FD100CEE" metricMagnitude="-2" metricUnit="m" metricValue="1.14" unit="mm" value="11.4">11.4 mm</quantity>
; width:
<quantity id="9B340F667C27A47C66C89EBB1FDA79C1" metricMagnitude="-3" metricUnit="m" metricValue="4.05" unit="mm" value="4.05">4.05 mm</quantity>
vs
<quantity id="F13F07FD08DFE9D36EE0CD61EB5F82F7" metricMagnitude="-3" metricUnit="m" metricValue="2.74" unit="mm" value="2.74">2.74 mm</quantity>
), and having more distal stylets in posterior gonopod (four vs two).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5VAG" type="description">
<paragraph id="8CE546A4F7C2DA6FC8539666B78F0DC7">
<heading id="83FE69A39E218CBE6481B225213D2F31" reason="title">Description.</heading>
</paragraph>
<paragraph id="9D7170F13CD57FA28F62EA95AB577D75">
<emphasis id="29A23EF9C431B95480DC97927AF9722C" bold="true">
Male
<typeStatus id="29279EE73EDD483F7DC48B203211E92F" type="holotype">holotype</typeStatus>
.
</emphasis>
</paragraph>
<paragraph id="70E8A25B8827871041839A7C3E650A58">
<emphasis id="5242FB52778B26AA0ADB1EB1574196E5" bold="true" italics="true">Measurements</emphasis>
: 61 body rings plus telson, length about
<quantity id="FDC54C4152B7203D24C21168B9C4A68A" metricMagnitude="-2" metricUnit="m" metricValue="1.864" unit="mm" value="18.64">18.64 mm</quantity>
, length of midbody metazona ca.
<quantity id="4E95D69BDC9B96F63A0A937FA53425D9" metricMagnitude="-4" metricUnit="m" metricValue="2.9" unit="mm" value="0.29">0.29 mm</quantity>
, width of midbody metazona ca.
<quantity id="6F906320735D209D16842BFBA10C9872" metricMagnitude="-3" metricUnit="m" metricValue="4.05" unit="mm" value="4.05">4.05 mm</quantity>
, width of midbody prozona ca.
<quantity id="2202E6B104122849293582E913673D3A" metricMagnitude="-3" metricUnit="m" metricValue="1.35" unit="mm" value="1.35">1.35 mm</quantity>
; ratio of width of head and width of collum = 0.98.
</paragraph>
<paragraph id="FCF5871B9E4065CC28E1E099E3CE8A64">
<emphasis id="D84E3DE50E47BF28B4E584734F0AA4EC" bold="true" italics="true">Coloration</emphasis>
of ethanol-preserved specimens: Terga and base of paraterga marbled yellowish brown with some blackish tubercles on several body rings. Collum and first 4 body rings yellow, 5
<superScript id="D84D128DFDE72DADB8C2EEF4434A7053">th</superScript>
weakly brownish, then increasing color pattern: 610
<superScript id="8A2EA873DBE9C4B4DB7996E8C6B45DFB">th</superScript>
light in the middle and brownish at the base of paraterga, 1112
<superScript id="0B6BBF99272CE75FEF29DF357DE64159">th</superScript>
dark brown in the middle and light borwn at the base of paraterga; then the same pattern repeated as 4 light + 3 brown, 5 light + 3 brown, 5 light + 3 brown, 5 light + 3 brown, 3 light + 2 brown, 2 light + 2 light brown, 3 light in the middle and brownish at the base of paraterga, then the ultimate 3 body rings plus telson and light yellow (Fig.
<figureCitation id="9B8EECF377E0235A261F77C6E9744649" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11 A C</figureCitation>
). Paraterga paler in preserved specimens, but pinkish in alive specimens.
</paragraph>
<caption id="DA987E0C9785030088E6A66DEE7203EC" ID-DOI="10.3897/zse.100.132111.figure11" ID-arpha="0F2B441D-7ECC-5CBD-909E-BA02C677E0DB" httpUri="https://binary.pensoft.net/fig/1169901" startId="F11">
<paragraph id="43E4FF7CD617AC3FA409B2A235858978">
<label id="58F5AFB1D191EBA98069474AC2D3A87F">Figure 11.</label>
</paragraph>
<paragraph id="05891EBDA5BBDD60E7B04B9B793E3001">
<taxonomicName id="0D8E1D58A5573745807191F940089BB1" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="karstomus" status="sp. nov.">
<emphasis id="E840592EB34EDDFE2CAD2646ED1D4DF1" italics="true">Pseudodesmus karstomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F782EC60EA62B347E22C7328A6B1A344" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0ERXAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
959 H), under white light.
<emphasis id="C49353B9B921B7A37F5A9D676948E4C8" bold="true">A.</emphasis>
Anterior part of body, dorsal view;
<emphasis id="B3B7906A8F736A36E1D19A9E4A22C615" bold="true">B.</emphasis>
Midbody body rings, dorsal view;
<emphasis id="A09368AD509D45F9904482678E90202C" bold="true">C.</emphasis>
Posterior part of body, dorsal view;
<emphasis id="68D59CBA0BEE88C195B0E1201C073445" bold="true">D.</emphasis>
Anterior part of body, ventral view;
<emphasis id="61E83445E21A719CFD036AA60262C2EC" bold="true">E.</emphasis>
Head and collum, ventral view;
<emphasis id="0E46096FF32932FACB68C77BB1F2EDC7" bold="true">F.</emphasis>
Posteriormost part of body, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="4C9E0E9F8C0DB3E893F246B2153FBC18">
Head (Figs
<figureCitation id="2AB7CE428B8A6783FAD6432DC8924C41" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11 D, E</figureCitation>
,
<figureCitation id="3D44D4AD50C4AAE22A77E72445F1ABD8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12 D, E</figureCitation>
) somewhat smooth, triangular-shaped. Antenna short, somewhat stout,
<emphasis id="444C50FDB0133C6EFC0A91CF9A2053DA" italics="true">in situ</emphasis>
extending to body ring 5; antennomere 6&gt; 5&gt; 3&gt; 4 = 2&gt; 7 =
<quantity id="0A0B8CA34D0872D3EF9E4DDB5C5ACEE1" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
length. Collum (Figs
<figureCitation id="0EED5D37B396B760314796A962E03388" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11 D, E</figureCitation>
,
<figureCitation id="FCB2457ED6DD32D6712B4315FD2DADCD" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12 D, E</figureCitation>
) stout strongly bilobed, anteriorly excavated, with two small bumps in the middle, densely setose. Paraterga very tiny, somewhat like lateral keels, caudal margin similar lateral keels.
</paragraph>
<caption id="2DA6912913AFAE488FF229582FDE9943" ID-DOI="10.3897/zse.100.132111.figure12" ID-arpha="469B7016-BD4C-5D35-B5AE-BCCD4A2A4879" httpUri="https://binary.pensoft.net/fig/1169902" startId="F12">
<paragraph id="DA376AC57BE8CB5D370ACAD8DE2F471A">
<label id="32CCB085C4053FABF91FEB30F553070B">Figure 12.</label>
</paragraph>
<paragraph id="3C9E9323E6EE322C781371E5D88A6CFC">
<taxonomicName id="0F59BC90896D7065CAE6F09C6FFE0014" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="karstomus" status="sp. nov.">
<emphasis id="DF2741ED032801F8193F2D09E932AEE0" italics="true">Pseudodesmus karstomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C5BB1EF5B15B8FC4B9746F5C94DFC345" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EOZAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
959 H), under UV light.
<emphasis id="8E69B4CB87421FC97F394E0DF1C1AAE8" bold="true">A.</emphasis>
Anterior part of body, dorsal view;
<emphasis id="3351A5BD6047F912DA6664198926195A" bold="true">B.</emphasis>
Midbody body rings, dorsal view;
<emphasis id="214A2ED38E2C7823FF4D03A2CA6CDF7E" bold="true">C.</emphasis>
Posterior part of body, dorsal view;
<emphasis id="34D0E388F1925022931648E66C7D3D03" bold="true">D.</emphasis>
Anterior part of body, ventral view;
<emphasis id="9CEB7D8A441017E3C01D1B88274C99B7" bold="true">E.</emphasis>
Head and collum, ventral view;
<emphasis id="B7B980FC60E8EA66C18197C214D06986" bold="true">F.</emphasis>
Posterior part of body, ventral vie. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="B5BF6D4D99E9567AFE77327A368A7665">
Tergum 2 with a row of 1 + 1 large, subtriangular tubercles, those tubercles reducing to rather small ones on subsequent terga. Terga 25 with one row, from 6
<superScript id="91D5A0ABE2762E77B8C0D9DF7ED14FA0">th</superScript>
onward with two rows of tubercles, anterior row with 78 + 7 8 tubercles, reaching the middle of paraterga, posterior row with only 2 + 2, never extending to paraterga; the 1 + 1 median tubercles (both anterior and posterior) transversally enlarged, elongated, oval (Figs
<figureCitation id="4FD004C8EBBA876EF8C3D7D0A616AB54" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11 A C</figureCitation>
,
<figureCitation id="A631EB5B9A74A8A1FD5A76EFA118C393" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12 A C</figureCitation>
).
</paragraph>
<paragraph id="875272753909720C69C45293921E63EE">
Paraterga in general very wide (Figs
<figureCitation id="E65992784D76FA553818D172019AC192" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11</figureCitation>
,
<figureCitation id="BABEFCDA39ECF40C981FE37B946E5979" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12</figureCitation>
). Paraterga 23 (Figs
<figureCitation id="B0DA43EDBD8DD5AB5665BEF139CF5025" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11 D, E</figureCitation>
,
<figureCitation id="7A90A2BC244DD0E6C7478C4855EEAF8A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12 D, E</figureCitation>
) anterad directed; paraterga 35 extending slightly lateroanterad, paraterga 658 laterad, 5961 lateroposteriad, 6263 posteriad. Paraterga 113 rectangular, from 14
<superScript id="E28924F1E52AD44729095DD76C76360D">th</superScript>
onwards tapering laterad, anterior edge increasingly bending backwards.
</paragraph>
<paragraph id="20583742A60BD5C02B40DE840E41CA6C">
Telson (Figs
<figureCitation id="00D5E74F0FC568FBE54172908CB06A0E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11 C, F</figureCitation>
,
<figureCitation id="5013C0CD4164D0F36356BF485CB08FDE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12 C, F</figureCitation>
) short, caudal margin rounded; epiproct with 3 + 3 setiferous tubercles at caudal margin (Figs
<figureCitation id="7488C8D93C33905478CB2D6EB555E2C8" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11 C, F</figureCitation>
,
<figureCitation id="BF3ACEF6BF521C39D3FEEA9B2E2C2798" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12 C, F</figureCitation>
). Paraproct and hypoproct semi-circular (Figs
<figureCitation id="6CC3D7F80A20296E3120009E572DDEC1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11 F</figureCitation>
,
<figureCitation id="D47F15A6FF2987A428B38CCBE00E40FA" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12 F</figureCitation>
).
</paragraph>
<paragraph id="77924007D34005B634146D0DBA18A49D">
Legs slender, shorter than paraterga, terminating before lateral paratergal margins. Claws normal. Coxal sacs present ventrally on body rings 359 (Figs
<figureCitation id="70451C6844E83ABD54D7ECC53273F487" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posteriormost part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure11" httpUri="https://binary.pensoft.net/fig/1169901">11 F</figureCitation>
,
<figureCitation id="F05FA61FF38164D414FDEA6736A6707A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), under UV light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Anterior part of body, ventral view; E. Head and collum, ventral view; F. Posterior part of body, ventral vie. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure12" httpUri="https://binary.pensoft.net/fig/1169902">12 F</figureCitation>
).
</paragraph>
<paragraph id="35F94FEF9C67ABF43F5D39ED71CBBE06">
<emphasis id="D313EA8063228A97CB8B640EB93EEC9E" bold="true" italics="true">Gonopods</emphasis>
(Figs
<figureCitation id="67E292406657ABB8C68A6292BA5D70A7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), gonopods. A. Left anterior gonopods, anterior view. B. Right posterior gonopods, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure13" httpUri="https://binary.pensoft.net/fig/1169903">13</figureCitation>
,
<figureCitation id="A518FF9A316C4EC1E5571693B1189E80" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), gonopods. SEM. A, B. Posterior gonopods, ventral view (A), anterior view (B); C, D. Anterior gonopods, ventral view (C), anterior view (D); E. Apical stylets of posterior gonopod, ventral view; F. Distal part of anterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure14" httpUri="https://binary.pensoft.net/fig/1169904">14</figureCitation>
): Two pairs of gonopods directed mesoanteriad. Anterior gonopods (Figs
<figureCitation id="86259B94EF437B1A5BF84D7678F799C2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), gonopods. A. Left anterior gonopods, anterior view. B. Right posterior gonopods, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure13" httpUri="https://binary.pensoft.net/fig/1169903">13 A</figureCitation>
,
<figureCitation id="FDEC02AAB4051CB6C2F6C5A1316C90D5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), gonopods. SEM. A, B. Posterior gonopods, ventral view (A), anterior view (B); C, D. Anterior gonopods, ventral view (C), anterior view (D); E. Apical stylets of posterior gonopod, ventral view; F. Distal part of anterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure14" httpUri="https://binary.pensoft.net/fig/1169904">14 A, B, F</figureCitation>
) 6 - segmented, covered with long setae; coxite (
<emphasis id="15C1F3FA9FEA5456CB80C72B3F0C59B2" italics="true">co</emphasis>
) broad, basal part sparsely covered with microgranulations; podomeres 25, short and stout; podomere 6 slightly longer than other podomeres, distally carrying macrosetae (
<emphasis id="2C61800AD9C7CEAAB37595E2DC435E7D" italics="true">mse</emphasis>
). Posterior gonopods (Figs
<figureCitation id="A4344E90CE3E72DD747854571C416A7A" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), gonopods. A. Left anterior gonopods, anterior view. B. Right posterior gonopods, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure13" httpUri="https://binary.pensoft.net/fig/1169903">13 B</figureCitation>
,
<figureCitation id="81149DCF5A7D274FCEBE8CA970E5C486" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Pseudodesmus karstomus sp. nov. Holotype (IEBR-Myr 959 H), gonopods. SEM. A, B. Posterior gonopods, ventral view (A), anterior view (B); C, D. Anterior gonopods, ventral view (C), anterior view (D); E. Apical stylets of posterior gonopod, ventral view; F. Distal part of anterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure14" httpUri="https://binary.pensoft.net/fig/1169904">14 C, D, E</figureCitation>
) 6 - segmented, covered with sparse long setae; coxite (
<emphasis id="04C58D19CB085F06C829C3C45C37506C" italics="true">co</emphasis>
) broad, basal part sparsely covered with microgranulations; podomeres 25 short and stout; ultimate podomere (
<emphasis id="65A5E04B0F631F66B2B6528F8FE3F189" italics="true">po 6</emphasis>
) longest, with two apical stylets and two lateroapical stylets (
<emphasis id="F218684D37CC09F84445DEA451538D2F" italics="true">sty</emphasis>
).
</paragraph>
<caption id="D581F848CBC73476169B65760C4C43C1" ID-DOI="10.3897/zse.100.132111.figure13" ID-arpha="2C43D11A-2FDC-52E4-8F02-FD4F54E8D8D1" httpUri="https://binary.pensoft.net/fig/1169903" startId="F13">
<paragraph id="8FF29F067CDF40135D1B25445742B929">
<label id="4E2D760BB29305E80B3FB7A9ED549F52">Figure 13.</label>
</paragraph>
<paragraph id="4E2DE7FD6B6FC99811A1DC25C30873B3">
<taxonomicName id="291CF8A65192B8781810924FD7197B6D" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="karstomus" status="sp. nov.">
<emphasis id="B9ECACF1EEC8899A4463989F1DE8C23F" italics="true">Pseudodesmus karstomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FED733AB57C8A21D0FD43D3728639462" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EE5AG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
959 H), gonopods.
<emphasis id="D87C3AB74AAFEF4F8A6D35399ABCAEE5" bold="true">A.</emphasis>
Left anterior gonopods, anterior view.
<emphasis id="3AFCE436681E6177A94982DD40D43D0C" bold="true">B.</emphasis>
Right posterior gonopods, anterior view.
<emphasis id="DC7CD054DA2E660E9004C19656F23457" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="7E4D9149577C2B8AC1CAE396BD704792" ID-DOI="10.3897/zse.100.132111.figure14" ID-arpha="810348C2-A39D-56D1-9444-3E64A8B7C519" httpUri="https://binary.pensoft.net/fig/1169904" startId="F14">
<paragraph id="E8BDFE7BAC3403417A3E3B868E97DB39">
<label id="5FB5846D84E0EE06823AA0A7985D1B98">Figure 14.</label>
</paragraph>
<paragraph id="EAA9533D07C99FB539860B75AE94A626">
<taxonomicName id="361C227E3B95CA009D19901FA51D6F18" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="karstomus" status="sp. nov.">
<emphasis id="094F9F3426CC9A1EF021DBA12A4AE860" italics="true">Pseudodesmus karstomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35590535292688304E5E32D1F442F193" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EH6AG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
959 H), gonopods.
<abbrev id="ABBRID0EM6AG" xlink_title="Scanning electron microscope">SEM</abbrev>
.
<emphasis id="CD4599A6D2DB8816AFEDFDA708D8D80E" bold="true">A, B.</emphasis>
Posterior gonopods, ventral view (
<emphasis id="ABD302C73427DEC45E448BEC49841E01" bold="true">A</emphasis>
), anterior view (
<emphasis id="AD83BF9A4410C86BD0338E3255CF9E56" bold="true">B</emphasis>
);
<emphasis id="1E744F97C713FC1416E7C69FB57C5CFA" bold="true">C, D.</emphasis>
Anterior gonopods, ventral view (
<emphasis id="D5B42951FF77EF933600189F6835389F" bold="true">C</emphasis>
), anterior view (
<emphasis id="E334AF2DF714D7103158C4025EE37977" bold="true">D</emphasis>
);
<emphasis id="5682DA2758D15C4533795864AF6D06E2" bold="true">E.</emphasis>
Apical stylets of posterior gonopod, ventral view;
<emphasis id="EF45077CE25F8C8CDEFD1C72A5251367" bold="true">F.</emphasis>
Distal part of anterior gonopod, anterior view.
<emphasis id="8A8713BA1A47155A37A2E2E891BFD539" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EHABG" type="variation">
<paragraph id="8DB8DF85DC065E8F7C18DD07BABC8411">
<heading id="CE62F3E026D45D9FE8EA4D3A97F8AC78" reason="title">Variation.</heading>
</paragraph>
<paragraph id="059B0D0F9E3AFFA7614695FD124D3194">
<emphasis id="8011D30039372C0EE2FB3C63A158B83A" bold="true" italics="true">
Male
<typeStatus id="F725C5C8DBBE6E8880343274CE6D31F4" type="paratype">paratypes</typeStatus>
</emphasis>
have 6263 plus telson, length about
<quantity id="4CF12C6EE036D43D813028DC035FA47F" metricMagnitude="-2" metricUnit="m" metricValue="1.7840000000000003" metricValueMax="1.7920000000000003" metricValueMin="1.7760000000000002" unit="mm" value="17.84" valueMax="17.92" valueMin="17.76">17.7617.92 mm</quantity>
, length of midbody metazoan ca.
<quantity id="BA89D520E35D062B220762EDF0F0EF91" metricMagnitude="-4" metricUnit="m" metricValue="3.1" metricValueMax="3.3" metricValueMin="2.9" unit="mm" value="0.31" valueMax="0.33" valueMin="0.29">0.290.33 mm</quantity>
, width of midbody metazona ca.
<quantity id="430E0E5A754161B47CDF6AF9F2FDFFDE" metricMagnitude="-3" metricUnit="m" metricValue="2.92" metricValueMax="2.94" metricValueMin="2.9" unit="mm" value="2.92" valueMax="2.94" valueMin="2.9">2.902.94 mm</quantity>
, width of midbody prozona ca.
<quantity id="7222FA64120A10A17CAA1ECCA5702844" metricMagnitude="-3" metricUnit="m" metricValue="1.41" metricValueMax="1.45" metricValueMin="1.37" unit="mm" value="1.41" valueMax="1.45" valueMin="1.37">1.371.45 mm</quantity>
; average ratio of width of head and width of collum = 0.89 (width of collum:
<quantity id="5EB8C05B435DBE60A1247EE32BADBA29" metricMagnitude="-4" metricUnit="m" metricValue="6.1" unit="mm" value="0.61">0.61 mm</quantity>
, width of head:
<quantity id="EB2EECD421C2755BD6A6195DFDA2A748" metricMagnitude="-4" metricUnit="m" metricValue="6.8" unit="mm" value="0.68">0.68 mm</quantity>
).
</paragraph>
<paragraph id="0B41E0652F9CE24F3BE31775ABDA3036">
<emphasis id="C9C000B91617FF079B7137F2F3D31430" bold="true" italics="true">
Female
<typeStatus id="5937B992569476C14CC60B7B3D01FE6C" type="paratype">paratype</typeStatus>
.
</emphasis>
66 body rings plus telson; length about
<quantity id="9D15A3EC8811F8830067AD7D42F37711" metricMagnitude="-2" metricUnit="m" metricValue="2.2359999999999998" unit="mm" value="22.36">22.36 mm</quantity>
, length of midbody metazoan ca.
<quantity id="21F93F180E548EC1582E29D1A04BBA8A" metricMagnitude="-4" metricUnit="m" metricValue="4.2" unit="mm" value="0.42">0.42 mm</quantity>
, width of midbody metazona ca.
<quantity id="144460D9DC2BA541215B84735E5FC417" metricMagnitude="-3" metricUnit="m" metricValue="4.06" unit="mm" value="4.06">4.06 mm</quantity>
, width of midbody prozona ca.
<quantity id="D450652AABE7C64AA9E1E37E15F89995" metricMagnitude="-3" metricUnit="m" metricValue="1.75" unit="mm" value="1.75">1.75 mm</quantity>
; ratio of width of head and width of collum = 0.9 (width of collum:
<quantity id="7854874CF38E2B094A17B4EF3530656C" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" unit="mm" value="0.72">0.72 mm</quantity>
, width of head:
<quantity id="628923918406A9A3D8C745C760C12FA2" metricMagnitude="-4" metricUnit="m" metricValue="6.5" unit="mm" value="0.65">0.65 mm</quantity>
). Nonsexual characters as in male.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZABG" type="etymology">
<paragraph id="265F46D11BC21E6EE217B6FE93D4F045">
<heading id="8A596EB320AFA5ABFE8A22B35EB22786" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="97C8ED192E74C8545822AB7022E5B422">
Named to emphasize the
<typeStatus id="5BB57DD32AFC9AE91F265C66E078AB5A">type</typeStatus>
locality, karst areas. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5ABG" type="distribution">
<paragraph id="151268511606A72B7E2B2F0A1A99ABF9">
<heading id="CD242435219928B9860304E462C02EBC" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="9B5114E18DA8D85D9F7B7C1613FA24AE">
The species was found in limestone mountain in
<collectingRegion id="37D1DB07DAFCA16E982025A3FB1D57E8" country="Vietnam" name="Cao Bang">Cao Bang Province</collectingRegion>
, northern
<collectingCountry id="8D7D3F07D381E52B96ED125F2A0E2FA3" name="Vietnam">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDBBG" type="DNA barcoding">
<paragraph id="89636E0B880A9112BCB2BC9EBBECB157">
<heading id="CA517ED54582D00970DA56F0A253C596" reason="title">DNA barcoding.</heading>
</paragraph>
<paragraph id="AC919F24F7DB18E07B11BED036FA6550">
A fragment of the
<abbrev id="ABBRID0EJBBG" xlink_title="cytochrome c oxidase subunit I">COI</abbrev>
gene is accessioned at
<collectionCode id="8855A454264AAD4F8BA1005D5896B0A0">NCBI</collectionCode>
GenBank with the following accession numbers:
<ext-link id="B5845B479AFC45651D7B5E89F74F1B01" ext-link-type="gen" xlink_href="PQ423224" xlink_type="simple">PQ 423224</ext-link>
<ext-link id="FAABC54B57F8471E1B170AE08B5A7FC0" ext-link-type="gen" xlink_href="PQ423225" xlink_type="simple">PQ 423225</ext-link>
. The new species has a close
<abbrev id="ABBRID0EXBBG" xlink_title="cytochrome c oxidase subunit I">COI</abbrev>
gene similarity with
<taxonomicName id="C6099891AD60C7E648F18CFFBBE68145" authorityName="Chamberlin" authorityYear="1922" class="Diplopoda" family="Andrognathidae" genus="Gosodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="claremontus">
<emphasis id="2416534C69229748C281DEE558797BE4" italics="true">Gosodesmus claremontus</emphasis>
</taxonomicName>
(
<ext-link id="8FBA7365B4987ABFDC73222F01E595FD" ext-link-type="gen" xlink_href="JX962723" xlink_type="simple">
<accessionNumber id="5ACE529E67DB2D18D51F66B8FE01FCF4" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JX962723">
<collectionCode id="7A10E01E33A8A307FAC79A7685787207">JX</collectionCode>
962723
</accessionNumber>
</ext-link>
) of 85.14 % in the
<collectionCode id="3F642662A01C3EC5DB22952F3BAACA6D">BLAST</collectionCode>
search.
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="182D09019DFCB702A55A7A6B4C729027" authority="Pierre, Fl. Forest. Cochinch." authorityName="Pierre, Fl. Forest. Cochinch." class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="section" section="Dicrananthera">
<emphasis id="0B8EA4C77AB4608C3B9F49E014D5750F" italics="true">Garcinia section Dicrananthera</emphasis>
Pierre, Fl. Forest. Cochinch.
</taxonomicName>
1 (5): 8. 1883;
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M. Gaudeul et al.,
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.
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<paragraph id="27AA0C03E6D3AA7177EC5F47062691DF">
<taxonomicName id="FC54E65EDC3361BE8196C30988029274" authority="Pierre, Fl. Forest. Cochinch." authorityName="Pierre, Fl. Forest. Cochinch." class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="thorelii">
<emphasis id="EBAD90A45921CA73E7BF80F24F795AA9" italics="true">Garcinia thorelii</emphasis>
Pierre, Fl. Forest. Cochinch.
</taxonomicName>
1 (4): t. 62. 1882.
</paragraph>
<paragraph id="101F9C6E8ADCB07E857D6049E420EE0D">
<taxonomicName id="89D70AB36217736266D742B9A9C7C1E0" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="section" section="Dicrananthera">
<emphasis id="B2B67516D536D21353B9EB1D365A652B" italics="true">Garcinia section Dicrananthera</emphasis>
</taxonomicName>
is distinguished by the presence of a pair of interpetiolar stipuliform structures (rare in
<taxonomicName id="16C73B9BD3D2DE56CA1F9FBF90CB87BD" authorityName=", Stevens" authorityYear="2007" class="Magnoliopsida" family="Clusiaceae" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="family">Clusiaceae</taxonomicName>
); leaves with prominent, widely spaced secondary veins and scalariform tertiary veins; axillary or terminal cymose inflorescences with three to many flowers [i. e.,
<taxonomicName id="5A7AA0BAC0A4E3EAE447141FE408E99F" authorityName="K. Armstr. &amp; P. W. Sweeney. Jones" authorityYear="1980" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="yaatapsap">
<emphasis id="9519ADFD30CC7CE8BC17D4007935C342" italics="true">G. yaatapsap</emphasis>
</taxonomicName>
(35 - flowered),
<taxonomicName id="4AF1209D4D157D4E570313AEE235F0F6" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="paucinervis">
<emphasis id="693FE357F09C525F9ADF7F3EB5C8C582" italics="true">G. paucinervis</emphasis>
</taxonomicName>
(410 - flowered),
<taxonomicName id="0C3C18031774DA886F1A19087FC0EF8B" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="stipulata">
<emphasis id="555E449F267385FB116B798DD17F84C1" italics="true">G. stipulata</emphasis>
</taxonomicName>
(46 - flowered),
<taxonomicName id="5F4AB25AD152A7C9FAFC980EA38F9DD7" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nujiangensis">
<emphasis id="82CACE38FAFE54203DF6B2AD5A0D8FCC" italics="true">G. nujiangensis</emphasis>
</taxonomicName>
(610 - flowered), and
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<emphasis id="3D4F46CCAD56AB13B9BE43994BB52CF6" italics="true">G. thorelii</emphasis>
</taxonomicName>
(2040 - flowered thyrse)]; flowers with 4 sepals and 4 petals; male flowers with numerous stamens united into an annular mass encircling and attached to the pistillode (i. e.,
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<emphasis id="5082A0F14D7A74C74F6485786A3B37C7" italics="true">G. stipulata</emphasis>
</taxonomicName>
and
<taxonomicName id="362FFF0AA3EEF01BB443BE5D2C01F57B" authorityName="K. Armstr. &amp; P. W. Sweeney. Jones" authorityYear="1980" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="yaatapsap">
<emphasis id="1296AC28522237E90638503403650718" italics="true">G. yaatapsap</emphasis>
</taxonomicName>
) or united into 4 bundles surrounding the pistillode (i. e.,
<taxonomicName id="6BBFC248D171D51496A3B5D1C67AEFBE" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nujiangensis">
<emphasis id="E44B91E8F625A78660E87F9F87214F8C" italics="true">G. nujiangensis</emphasis>
</taxonomicName>
,
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<emphasis id="3E168A85A4132A2614B52C51B377D086" italics="true">G. paucinervis</emphasis>
</taxonomicName>
, and
<taxonomicName id="CCCB6CD127984F377408B6EDB0FD9C3F" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="thorelii">
<emphasis id="AE34E20364C9FCA42B174A2A5231599C" italics="true">G. thorelii</emphasis>
</taxonomicName>
), 2 - thecous anthers; 12 - locular ovaries; unlobed and smooth stigmas; and fruits with a smooth surface and unlobed. Distinguishing morphological characters reported here for this section were taken from
<bibRefCitation id="67312F16E7C040671C485BB2553E3556" author="Pierre" firstAuthor="Pierre" refId="B28" refString="Pierre JBL (1882) Flore Forestiére de la Cochinchine Vol 1 (4). Octave Doin, Éditeur, Paris, t. 4964." year="1882">Pierre 1882</bibRefCitation>
,
<bibRefCitation id="9C17CC70B28DFBCA42DC07A4826C6F08" author="Pitard" editor="Lecomte PH" firstAuthor="Pitard" journalOrPublisher="1 (4). Masson et Cie, Éditeurs, Paris" pagination="289-448" refId="B30" refString="Pitard CJ (1910) Hypéricacées, Guttiféres, Ternstræmiacées et Stachyuracées. In: Lecomte PH (Ed.) Flore Générale de lIndo-Chine Vol. 1 (4). Masson et Cie, Éditeurs, Paris, 289448." title="Hypéricacées, Guttiféres, Ternstræmiacées et Stachyuracées." volumeTitle="Flore Générale de lIndo-Chine Vol." year="1910">Pitard 1910</bibRefCitation>
,
<bibRefCitation id="177D79F304021440E3CEC15BF4834C7C" author="Chun and How" firstAuthor="Chun" issue="1" journalOrPublisher="Acta Phytotaxonomica Sinica" pagination="1-25" refId="B5" refString="Chun WY, How FC (1956) Species novae arborum utilium Chinae meridionalis. Acta Phytotaxonomica Sinica 5 (1): 125." title="Species novae arborum utilium Chinae meridionalis." volume="5" year="1956">Chun and How 1956</bibRefCitation>
,
<bibRefCitation id="9853459005A22695BDF7EFF81C95158F" author="Jones" firstAuthor="Jones" journalOrPublisher="PhD Thesis, University of Leicester and British Museum (Natural History), London" refId="B12" refString="Jones SW (1980) Morphology and Major Taxonomy of Garcinia (Guttiferae). PhD Thesis, University of Leicester and British Museum (Natural History), London, 474 pp." title="Morphology and Major Taxonomy of Garcinia (Guttiferae)." year="1980">Jones 1980</bibRefCitation>
,
<bibRefCitation id="9DB03C18471A3A36D4D4901589A1AA01" author="Li" firstAuthor="Li" journalOrPublisher="Acta Phytotaxonomica Sinica" pagination="490-499" refId="B15" refString="Li YH (1981) Some new species of the genus Garcinia from South China. Acta Phytotaxonomica Sinica 19: 490499." title="Some new species of the genus Garcinia from South China." volume="19" year="1981">Li 1981</bibRefCitation>
,
<bibRefCitation id="F8D2473B8EA26217B92450E139E1C353" author="Li" editor="Wu CY &amp; Raven PH &amp; Hong DY" etAl="et al." firstAuthor="Li" journalOrPublisher="Science Press, Beijing, and Missouri Botanical Garden Press, St. Louis" pagination="40-47" refId="B16" refString="Li XW, Li J, Stevens P (2007) Garcinia (Clusiaceae). In: Wu CY, Raven PH, Hong DY (Eds) Flora of China Vol 13 (Clusiaceae through Araliaceae). Science Press, Beijing, and Missouri Botanical Garden Press, St. Louis, 4047." title="Garcinia (Clusiaceae)." volumeTitle="Flora of China Vol 13 (Clusiaceae through Araliaceae)." year="2007">Li et al. 2007</bibRefCitation>
,
<bibRefCitation id="60498D54D4383AB20DEB55C904C9BEBA" DOI="10.1007/978-3-540-32219-1_10" author="Stevens" editor="Kubitzki K" firstAuthor="Stevens" journalOrPublisher="Springer-Verlag, Berlin" pagination="48-66" refId="B36" refString="Stevens PF (2007) Clusiaceae - Guttiferae. In: Kubitzki K (Ed.) The Families and Genera of Vascular Plants Vol 9, Flowering Plants, Eudicots: Berberidopsidales, Buxales, Crossosomatales, Fabales pp, Geraniales, Gunnerales, Myrtales pp, Proteales, Saxifragales, Vitales, Zygophyllales, Clusiaceae alliance, Passifloraceae alliance, Dilleniaceae, Huaceae, Picramniaceae, Sabiaceae. Springer-Verlag, Berlin, 4866. https://doi.org/10.1007/978-3-540-32219-1_10" title="Clusiaceae - Guttiferae." volumeTitle="The Families and Genera of Vascular Plants Vol 9, Flowering Plants, Eudicots: Berberidopsidales, Buxales, Crossosomatales, Fabales pp, Geraniales, Gunnerales, Myrtales pp, Proteales, Saxifragales, Vitales, Zygophyllales, Clusiaceae alliance, Passifloraceae alliance, Dilleniaceae, Huaceae, Picramniaceae, Sabiaceae." year="2007">Stevens 2007</bibRefCitation>
,
<bibRefCitation id="67FC2879E548255D47DC67DD829BCDA4" DOI="10.3897/phytokeys.239.112563" author="Gaudeul" etAl="et al." firstAuthor="Gaudeul" journalOrPublisher="PhytoKeys" pagination="73-105" refId="B9" refString="Gaudeul M, Sweeney P, Munzinger J (2024) An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence. PhytoKeys 239: 73105. https://doi.org/10.3897/phytokeys.239.112563" title="An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence." volume="239" year="2024">Gaudeul et al. 2024</bibRefCitation>
, and from our observations.
</paragraph>
<paragraph id="7056131D9AD96C06D0C236023424A683">
A section of five species worldwide (
<bibRefCitation id="72EFDB8D02A2749842B96998EF2B6C11" DOI="10.3897/phytokeys.239.112563" author="Gaudeul" etAl="et al." firstAuthor="Gaudeul" journalOrPublisher="PhytoKeys" pagination="73-105" refId="B9" refString="Gaudeul M, Sweeney P, Munzinger J (2024) An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence. PhytoKeys 239: 73105. https://doi.org/10.3897/phytokeys.239.112563" title="An updated infrageneric classification of the pantropical species-rich genus Garcinia L. (Clusiaceae) and some insights into the systematics of New Caledonian species, based on molecular and morphological evidence." volume="239" year="2024">Gaudeul et al. 2024</bibRefCitation>
); one species in
<collectingCountry id="386BFAFF7A624E98416EB221EA631C4F" name="Thailand">Thailand</collectingCountry>
.
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<mods:affiliation id="44CFEE4025DFF9ED3E60026C2A4C34F8">University of Veterinary Medicine Budapest, István u. 2, 1078 Budapest, Hungary</mods:affiliation>
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<mods:typeOfResource id="E4283EA352E11ADA75A77EC7625C0DBA">text</mods:typeOfResource>
<mods:relatedItem id="D84E8CEA9281429A5DD3BC314DBF355D" type="host">
<mods:titleInfo id="AE1DC0654AEA16AA011FF6A62D979E3A">
<mods:title id="66A1E95AFCF451AB0B228999A01E09B5">Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part id="872433F1B238A17B4469C2C945C18C16">
<mods:date id="B86E6740F6D6076D978ACF371A7CB77D">2024</mods:date>
<mods:detail id="9676E5B0170C4666D330AEEE6A9DCA35" type="pubDate">
<mods:number id="7802ED0C88E8F5B1A6A939963E69BBF4">2024-11-07</mods:number>
</mods:detail>
<mods:detail id="A40285C162A702BDA8E1C30D544E2CD9" type="volume">
<mods:number id="A0972A4F7E9AC73293F6DC7F459B081E">100</mods:number>
</mods:detail>
<mods:detail id="0622B0DC75965C0141B203BA3DE423E2" type="issue">
<mods:number id="86BC6A64E9567D5F5F9ECFB8C196467D">4</mods:number>
</mods:detail>
<mods:extent id="A8F8BC0BB04326D0438C4E955C5CC13C" unit="page">
<mods:start id="0ADDB075549609ABC3B7D66DB4533F7E">1515</mods:start>
<mods:end id="E066203A00C923CA77DB7FDD6B358089">1541</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C6FB19D263B9074452EBDE206AF57198">journal article</mods:classification>
<mods:identifier id="F5B922944F0CD76317434E04B77D84F6" type="DOI">10.3897/zse.100.132111</mods:identifier>
<mods:identifier id="4879C39CD29D9A41C1796230A3793220" type="ZooBank">B39508EB-6EB2-4CC0-88D8-B3E87BDCED02</mods:identifier>
</mods:mods>
<treatment id="92A4751707F158C499FD3B94C2D1B998" ID-DOI="http://doi.org/10.5281/zenodo.14052233" ID-Zenodo-Dep="14052233" ID-arpha="92A47517-07F1-58C4-99FD-3B94C2D1B998" ID-zoobank="https://zoobank.org/2FD3A321-E3F2-419E-A200-4446A132B10A" LSID="urn:lsid:plazi:treatment:92A4751707F158C499FD3B94C2D1B998" LSID-ZBK="urn:lsid:zoobank.org:act:2FD3A321-E3F2-419E-A200-4446A132B10A" httpUri="http://treatment.plazi.org/id/92A4751707F158C499FD3B94C2D1B998">
<subSubSection id="76FFCDCF45C694B1C77CAF291391CD99" type="nomenclature">
<paragraph id="F69C680C1A19D4EA43488545D256D60E">
<taxonomicName id="835A0541C3EE713BE6866B3C0A338FB7" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Playtdesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="2F3CCCF33AE2DEDA2944C06802B9771A" italics="true">Pseudodesmus irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="624BB255AAB10403A82A1E3DECAB7E8E" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DB7883FBE81A28543D9CDA4D31C65830" type="description">
<paragraph id="95AC61DB027DCA6DBB9D8FFBD79631B9">
<figureCitation id="D4C05D747F6C78DBC36975B8FC042950" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">Figs 21</figureCitation>
,
<figureCitation id="C1E3904937A970D761D1BE3B011D60C4" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22</figureCitation>
,
<figureCitation id="FC3E325D6CA5D60EBF2DB13101800BEE" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. A. Left anterior gonopod, anterior view; B. Left posterior gonopod, sublateral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure23" httpUri="https://binary.pensoft.net/fig/1169913">23</figureCitation>
,
<figureCitation id="250B03AF395B4166D6693C28E8F05619" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. SEM. A, B. Left anterior gonopod, anterior view (A), ventral view (B); C. Distal part of left anterior gonopod, anterior view; D, E. Posterior gonopod and leg 11, ventral view (D), sublateral view (E); F. Distal part of posterior gonopod, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure24" httpUri="https://binary.pensoft.net/fig/1169914">24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ED1BG" type="material">
<paragraph id="E11A1E99A6914A3E50FB95B9151D4077">
<heading id="04E2C7C7DBB1B22490B74AA6B10EFB8B" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="070712A5CAC703A52ACE3488530D78ED">
<materialsCitation id="49374A688EF41B262508D28DDAF83464" collectingDate="2006-04-15" collectingDateMax="2006-04-24" collectingDateMin="2006-04-15" collectorName="A. D. Nguyen" country="Vietnam" elevation="1500" latitude="12.11124" location="Hon Ba Mt." longLatPrecision="1" longitude="108.98426" specimenCount="1" specimenCount-male="1" stateProvince="Khanh Hoa" typeStatus="holotype">
<emphasis id="A961241B165D41FC56ADF56525C9BFDC" bold="true" italics="true">
<typeStatus id="35D62E9349500536C4BF59B88FD87AEE" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="A5BE041FB1BEBA45DDDFBEA7B722B2E0" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="807AE7A2380DD8BF4FB54413BA991D2C" count="1" type="male">1 male</specimenCount>
;
<collectingRegion id="A2FA515AC9FC2A786BF659810B33B23C" country="Vietnam" name="Khanh Hoa">Khanh Hoa Province</collectingRegion>
,
<location id="2911AB3C3CBA63534BAEE12E6BFC04C7" LSID="urn:lsid:plazi:treatment:92A4751707F158C499FD3B94C2D1B998:2911AB3C3CBA63534BAEE12E6BFC04C7" country="Vietnam" latitude="12.11124" longLatPrecision="1" longitude="108.98426" name="Hon Ba Mt." stateProvince="Khanh Hoa">Hon Ba Mt.</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ER1BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[108.984260,12.111240]}">
<geoCoordinate id="68A2A8F4ADD3E45DFFF2C077A971C5AB" degrees="12.11124" direction="north" orientation="latitude" precision="1" value="12.11124">12.11124 ° N</geoCoordinate>
,
<geoCoordinate id="146C429CDD142418F932E22A0BA2E39C" degrees="108.98426" direction="east" orientation="longitude" precision="1" value="108.98426">108.98426 ° E</geoCoordinate>
</named-content>
;
<elevation id="D5DA4A84E2526B21F85E753982BF56C4" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
<quantity id="FB840B0B50D716B5937F761DB1A103E7" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1,500 m</quantity>
a. s. l.
</elevation>
;
<date id="FFBF79BCDB3C6D0CC1D4FF7F6C9CCA7D" value="2006-04-15" valueMax="2006-04-24" valueMin="2006-04-15">
<collectingDate id="11FCAAEC64C2A118AB6F239BA6AC6B13" value="2006-04-15" valueMax="2006-04-24" valueMin="2006-04-15">1524 Apr. 2006</collectingDate>
</date>
;
<collectorName id="F13C63EC4814B88798A72C32792A026B">A. D. Nguyen</collectorName>
leg.; primary forest;
<abbrev content-type="institution" id="ABBRID0EW1BG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
958 H.
</materialsCitation>
</paragraph>
<paragraph id="06C116DDDAF7C37FC70A739CAF5E2F75">
<materialsCitation id="6E30D8536BA40EA05C38D1130B3E062F" collectingDate="2006-04-15" collectingDateMax="2006-04-24" collectingDateMin="2006-04-15" collectorName="A. D. Nguyen" country="Vietnam" elevation="1500" latitude="12.11124" location="Hon Ba Mt." longLatPrecision="1" longitude="108.98426" specimenCount="1" specimenCount-male="1" stateProvince="Khanh Hoa" typeStatus="paratype">
<emphasis id="7060CC674070ECD3B6744AF8CE37EDA4" bold="true" italics="true">
<typeStatus id="C738A765EA0B3A282D7F483030322DA8" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="BFCDC57F6CD2A522E8A8D9FC7AEED731" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="CE61F84AE3502B9C432E4BDA437D3E39" count="1" type="male">1 male</specimenCount>
; same data as for holotype;
<abbrev content-type="institution" id="ABBRID0EC2BG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
958 P
</materialsCitation>
<materialsCitation id="E405F93C2297CC10E20F1B975F2517D6" collectingDate="2004-03-25" collectingDateMax="2004-04-11" collectingDateMin="2004-03-25" collectorName="A. D. Nguyen" country="Vietnam" elevation="1700" latitude="15.07071" location="Ngoc Linh Mt" longLatPrecision="1" longitude="107.97142" specimenCount="1" specimenCount-female="1" stateProvince="Kon Tum" typeStatus="paratype">
<specimenCount id="8946CDE967036F0CAE997D145AF0FC3B" count="1" type="female">1 female</specimenCount>
;
<collectingRegion id="1CDFB8321203ACB28BE752F502365F24" country="Vietnam" name="Kon Tum">Kon Tum Province</collectingRegion>
,
<location id="EE6923B08EF0375718027810DD839802" LSID="urn:lsid:plazi:treatment:92A4751707F158C499FD3B94C2D1B998:EE6923B08EF0375718027810DD839802" country="Vietnam" latitude="15.07071" longLatPrecision="1" longitude="107.97142" name="Ngoc Linh Mt" stateProvince="Kon Tum">Ngoc Linh Mt</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EK2BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.971420,15.070710]}">
<geoCoordinate id="37304AC4EF3B7FDF574ABE44C3627B4A" degrees="15.07071" direction="north" orientation="latitude" precision="1" value="15.07071">15.07071 ° N</geoCoordinate>
,
<geoCoordinate id="5DF52E8F526DA0F90972FB31B258FB8D" degrees="107.97142" direction="east" orientation="longitude" precision="1" value="107.97142">107.97142 ° E</geoCoordinate>
</named-content>
;
<elevation id="36F52CABC211C1A33B938D7A6FB4C77C" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<quantity id="382D83E3F908B2165542E7DD8A133B40" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1,700 m</quantity>
a. s. l.
</elevation>
;
<date id="37F78344CF7B80F99D2B6B8BCD6C3CDE" value="2004-03-25" valueMax="2004-04-11" valueMin="2004-03-25">
<collectingDate id="AC5AEC19A877DAF7E871DE9B98C72677" value="2004-03-25" valueMax="2004-04-11" valueMin="2004-03-25">25 Mar. 11 Apr. 2004</collectingDate>
</date>
,
<collectorName id="5C5C0D75F98A4BAEE3943D7FCA8C4209">A. D. Nguyen</collectorName>
leg.; regenerated forest;
<abbrev content-type="institution" id="ABBRID0EP2BG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
960
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EU2BG" type="diagnosis">
<paragraph id="06517F8696C6FB2254B27AD57219AD7D">
<heading id="C126A4C0EDDF84F9BDAD812F6F89C025" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="94D844B2511758CEF3916D654EB3F4E6">Head narrower than collum. Collum bilobed, caudal margin highly elevated with 3 + 3 tubercles with 3 + 3 tubercles, and with a row of 1 + 1 tubercles. Midbody ring with 2 distinct rows of tubercles, anterior row of 69 + 6 9 and posterior of 3 + 3. Paratergal margins with small / tiny dentations or notches. Posterior gonopods 6 - segmented, with three apical stylets.</paragraph>
</subSubSection>
<subSubSection id="SECID0EZ2BG" type="diagnosis">
<paragraph id="CE4B11782FCBD379D35E5E666E660167">
<heading id="18820DD803CAD07441F65C76CD3A88C8" reason="title">Diagnosis remarks.</heading>
</paragraph>
<paragraph id="722EB33E483565CCAEDDE3E954AB1012">
The new species is distinguished from
<taxonomicName id="652CBEC50E862E58F72A343B60A597A7" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="camptotrichus">
<emphasis id="C05AAAECEEE1B4BA209DA72CC12C3F95" italics="true">P. camptotrichus</emphasis>
</taxonomicName>
by coloration (irregularly yellow with brown spots vs earth brown), shorter length (
<quantity id="89A3544D888F2E903FD039C0C4A8439C" metricMagnitude="-2" metricUnit="m" metricValue="1.292" unit="mm" value="12.92">12.92 mm</quantity>
vs
<quantity id="4A5C382AEEAB2BB5A361494DBC2CDC7E" metricMagnitude="-2" metricUnit="m" metricValue="1.4" unit="mm" value="14.0">14 mm</quantity>
), slightly wider (
<quantity id="D97A0F4656D6E6741AD8BC174E121F72" metricMagnitude="-3" metricUnit="m" metricValue="2.39" unit="mm" value="2.39">2.39 mm</quantity>
vs 2.0 mm), having less number of tubercles in posterior metazonal row (3 + 3 vs 69 + 6 9). The new species is similar to
<taxonomicName id="5C5A78FBB7818C0839E643EDBCFD903F" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="0BD96D7B51E5AEF0536DC1F60885CB4A" italics="true">P. variegatus</emphasis>
</taxonomicName>
regarding body coloration (irregularly yellow with brown spots), but two species can be separated by body size (length:
<quantity id="7D952364108AC7B99192574B47C36B04" metricMagnitude="-2" metricUnit="m" metricValue="1.296" unit="mm" value="12.96">12.96 mm</quantity>
vs
<quantity id="3E15586925CECEBFA805A75AF8F37CA0" metricMagnitude="-2" metricUnit="m" metricValue="3.2" unit="mm" value="32.0">32 mm</quantity>
; width:
<quantity id="4D65BE99F37388C508CFA54EDF571392" metricMagnitude="-3" metricUnit="m" metricValue="2.39" unit="mm" value="2.39">2.39 mm</quantity>
vs
<quantity id="074CFAA1668693BC1224CC76EF249B70" metricMagnitude="-3" metricUnit="m" metricValue="5.8" unit="mm" value="5.8">5.8 mm</quantity>
) and and number of metazonal tubercles (two rows of 69 + 6 9 and 3 + 3 vs two rows of 815 + 8 15 and 57 + 5 7).
</paragraph>
<paragraph id="1C892FB59ADEB79C529BCA58835B36C5">
<taxonomicName id="EAB86053E8D4CBCAD58C2BD0C1DD9A38" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="40B771AB9AB5B7E40470A8CBFFE4E4D7" italics="true">Pseudodesmus irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FD9E428DD9EA076D0B15E1DBFD8A3EF0" rank="species">sp. nov.</taxonomicNameLabel>
is differentiated from above four new species,
<taxonomicName id="16EEB83CE5DD57F1D0A8FE4B43E6414F" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
<emphasis id="BC6DF077DE9D394887EE3DD0700D5242" italics="true">P. bidoup</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D371B82A1D43F3B9EC0CCBED22A193DC" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="731F6AE1D2B08DCE66676921EB4334D4" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov">
<emphasis id="7D70FCD230DF420E8F99048E841F03C0" italics="true">P. ngoclinh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F315AC10B4878F3738FD53BF8464A279" rank="species">sp. nov</taxonomicNameLabel>
,
<taxonomicName id="F846FF89687F032752455F90310984BC" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="karstomus" status="sp. nov.">
<emphasis id="BE464DDA84334345A325614109E60492" italics="true">P. karstomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="615082F07DE9CF68EA08BDF7F5777DBF" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="004E05020971C4C347BC4832B4BDC73A" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="condao" status="sp. nov.">
<emphasis id="3E9680D2AD3F9C98D5FD49F6B8702C75" italics="true">P. condao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBCB4E7D938FAA794C7C65DCA4240426" rank="species">sp. nov.</taxonomicNameLabel>
in body coloration of irregularly yellow with brown spots, body size and number of stylets (three vs four, two, four, and two, respectively).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EO5BG" type="description">
<paragraph id="76D01D1670F676C1B800E82F0971F42C">
<heading id="83CF53C61049AC080554EAB234C88C39" reason="title">Description.</heading>
</paragraph>
<paragraph id="1F233A94A28D4C1A640E79250D21E4E8">
<emphasis id="802FBEF239FA916488C992619A42BDAF" bold="true">
Male
<typeStatus id="34BE8C29AB1BA1CC548889EE8D3588CB" type="holotype">holotype</typeStatus>
.
</emphasis>
</paragraph>
<paragraph id="718B01DF5C3446917972955859EAC0A2">
<emphasis id="6ADCAB57E55781C4DDB41DA84C4C7EA2" bold="true" italics="true">Measurements</emphasis>
: 42 body rings plus telson, length about
<quantity id="388D46933AC9FF62AA59147669AFF5B9" metricMagnitude="-2" metricUnit="m" metricValue="1.296" unit="mm" value="12.96">12.96 mm</quantity>
, length of midbody metazona
<quantity id="50F4B5503AD3F6A48069A7CF6A8B6AA3" metricMagnitude="-4" metricUnit="m" metricValue="2.4" unit="mm" value="0.24">0.24 mm</quantity>
, width of midbody metazona
<quantity id="D527EAEC5F1A162EFD8779AF9375066D" metricMagnitude="-3" metricUnit="m" metricValue="2.39" unit="mm" value="2.39">2.39 mm</quantity>
, width of midbody prozona
<quantity id="BBDBE19BCA836217F1713B77B8007AB3" metricMagnitude="-3" metricUnit="m" metricValue="1.08" unit="mm" value="1.08">1.08 mm</quantity>
; ratio of width of head and width of collum = 0.74.
</paragraph>
<paragraph id="6C77CCFDBD860387FBF2D8A05F336273">
<emphasis id="8A5AF910AFED73F5128B8C7DA9628A06" bold="true" italics="true">Coloration</emphasis>
of ethanol-preserved specimens (Fig.
<figureCitation id="066538CBDE51A864F4F1D9D1FEC522A0" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. SEM. A, B. Left posterior gonopod, subventral view (A), anterior view (B); C. Anterior gonopods, anterior view; D. Left anterior gonopod, anterior view; E. Distal part of left posterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure20" httpUri="https://binary.pensoft.net/fig/1169910">20</figureCitation>
): complex and irregular: body rings 14 with none, body rings 57 with 2 brown spots, body ring 8 brown on the right, body ring 9 with two brown spots, body ring 10 brown on the right, body ring 11 brown on the left, body rings 1213 with two brown spots, body ring 14 none, body ring 15 brown on the left, body rings 1617 brown on the right, body ring 18 brown on left half, body ring 19 brown on right half, body ring 20 brown on left half, body ring 21 none, body ring 22 with left spot, body ring 23 with two brown spots, body rings 2426 brown on left half, body rings 2728 full dark, body ring 29 none, body rings 3032 brown left, body rings 3334 brown right, body rings 3536 full, body rings 3738 none, body ring 39 brown right, body rings 4042 light brown.
</paragraph>
<paragraph id="ED3E6A3206B9DA399DF7BE47AEE77000">
Head (Figs
<figureCitation id="25532EFAABB1847EAA99A669B5F71D9A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 B</figureCitation>
,
<figureCitation id="9C6E26D0D9B008F4BC60FF0D35991E41" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 B</figureCitation>
) narrower than collum in width, smooth, round-shaped. Antenna stout, clavate,
<emphasis id="EE8D3BB145792A81C740C0BB39CB50B2" italics="true">in situ</emphasis>
extending to body ring 6; antennomere 6&gt; 5&gt; 3&gt; 4 = 2&gt; 7 =
<quantity id="B5384444DC325EAF128AF6FC4BC27622" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
length (Figs
<figureCitation id="3D18ADC72CAB9BE5426AF7F2BB7972A5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 B</figureCitation>
,
<figureCitation id="3649B38DC65B84E3C75ADECE6E4848BD" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 B</figureCitation>
). Collum (Figs
<figureCitation id="67F4AAF36335D3C4E713C84E2AC87AF1" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 B</figureCitation>
,
<figureCitation id="D12254901AE1D727C6AC66A41E4F6E9C" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 B</figureCitation>
) bilobuled, caudal margin highly elevated with 3 + 3 tubercles, and with a row of 1 + 1 tubercles, paramedian one much larger than lateral one.
</paragraph>
<caption id="6FC915BBC12E5A9CC34CC0A5F0534755" ID-DOI="10.3897/zse.100.132111.figure21" ID-arpha="3BFBCC07-E890-575A-A50D-59E368F11E98" httpUri="https://binary.pensoft.net/fig/1169911" startId="F21">
<paragraph id="44D73A148AEDAFD9C86C84D3AF41E68E">
<label id="96BACA2E26B33F210F0AF1F3AC707BF7">Figure 21.</label>
</paragraph>
<paragraph id="618CD4BE16461E273AECDAEBC3535B3C">
<taxonomicName id="215E79C3E519AFF5629E953B51360736" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="DBA25AFBAADD4DA4F93100307B869172" italics="true">Pseudodesmus irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E523AD0F377784AAD24FC3895419141A" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EYAAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
958 H), under normal light.
<emphasis id="00A5D2704D0D9E9FBAB8583EF0CAEA20" bold="true">A, B.</emphasis>
Anterior part of body, dorsal view (
<emphasis id="F0942817E5530F410077E12BE2E86A67" bold="true">A</emphasis>
), ventral view (
<emphasis id="9A7C467E75C5C005353130A49F88776F" bold="true">B</emphasis>
);
<emphasis id="D5CD85EE7A85F6D9A07B092ACE5ECF29" bold="true">C, D.</emphasis>
Midbody body rings, dorsal view (
<emphasis id="C3B86C2D89B869BF56315E29A1CBAED8" bold="true">C</emphasis>
), ventral view (
<emphasis id="4DA344CA524C5DB67F647C1071A154DC" bold="true">D</emphasis>
);
<emphasis id="20A5A46E1E4C988AE8ADDF9E3AB137ED" bold="true">E, F.</emphasis>
Posterior part of body, dorsal view (
<emphasis id="C1255ED4102F79F8C9BC2BAB69725CF9" bold="true">E</emphasis>
), ventral view (
<emphasis id="9BEF0D801ECA9FDBED17F4718980202B" bold="true">F</emphasis>
). Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="343AE333CA58A3DCE1A2A46C9ABD55A7" ID-DOI="10.3897/zse.100.132111.figure22" ID-arpha="4F60D10F-21F1-5FCB-BB11-8B1451742D6B" httpUri="https://binary.pensoft.net/fig/1169912" startId="F22">
<paragraph id="F0AB55AD1C680939A7F05674229CC5B6">
<label id="CE1DC4B2568988F15749CC123AE06880">Figure 22.</label>
</paragraph>
<paragraph id="8A3BB8394EC26187117BC8ACCF936DE7">
<taxonomicName id="6DE6B1850173235F10F1E04F6F0481F9" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="9607A79F7F2F4234ADEA155ACAFD37A6" italics="true">Pseudodesmus irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BA5A5752F747CFD28BD229D254A16B26" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EHCAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
958 H), under UV light.
<emphasis id="DF183C5F1C02B11C17AF5CB628E7DA10" bold="true">A, B.</emphasis>
Anterior part of body, dorsal view (
<emphasis id="B7B8E3DA76BF84A066943B7F6312F08B" bold="true">A</emphasis>
), ventral view (
<emphasis id="A14E54B8A6039FF4C353C496782D30C5" bold="true">B</emphasis>
);
<emphasis id="7A2F7716B5DB0F88F9AE2403DA59CC90" bold="true">C.</emphasis>
Cross section of midbody body ring, posterior view;
<emphasis id="5544A4493A9FC70FCFD25A8B2B905743" bold="true">D.</emphasis>
Midbody body rings, dorsal view;
<emphasis id="8394E9391FAE743A072D6ED1DAD7C1D3" bold="true">E, F.</emphasis>
Posterior part of body, dorsal view (
<emphasis id="0BF3E1DC0FE54A09321F74ADE6646BB0" bold="true">E</emphasis>
), ventral view (
<emphasis id="B58579B50D9247C2FCF40D4A66A0EFA9" bold="true">F</emphasis>
). Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="98C8C486D36A52F6501B51D3D83CC9C6">
Body (Figs
<figureCitation id="0384762FF27C72577566968EC1D4002D" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21</figureCitation>
,
<figureCitation id="2061633F32DEB4D557BBF762A9134164" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22</figureCitation>
): ring 2 with two rows of 3 + 3 and 1 + 1 tubercles; ring 3 with two rows of 3 + 3 and 3 + 3 tubercles; other rings with 2 distinct rows of tubercles, anterior row of 6 + 6 (increasing to (9 + 9) on midbody rings), and posterior of 2 + 4 (body ring 4), 3 + 3 (ring 5), 3 + 2 (ring 6), 3 + 3 (on other rings). The last 5 rings with two rows of 3 + 3 and 3 + 3 tubercles (Fig.
<figureCitation id="A6255C721A6218C6106C5B222DDE0B02" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 E</figureCitation>
). Anterior rows extending to 2 / 3 paraterga. Paramedian tubercles on posterior rows much larger than ones on anterior rows.
</paragraph>
<paragraph id="693091F54DBA47DD9B1BD5A84D1A9CDF">
Paraterga well developed, slightly curved anteriad on body rings 23, lateral margins smooth (Figs
<figureCitation id="1762B618EC8C477FB9E8F20967D4B863" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 A, B</figureCitation>
,
<figureCitation id="6A004220E042E333674958B00228B343" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 A, B</figureCitation>
); increasingly less so on the following body rings, slightly curved caudad on body rings 2527, and strongly curved caudad on body rings 3740; paraterga of penultimate body ring produced strictly caudad and flanking telson (Figs
<figureCitation id="99BAED6CD845F7C238F587366F3289CF" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 E, F</figureCitation>
,
<figureCitation id="4853AF22948D0A0B5AEB5DE2B21FBB6F" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 E, F</figureCitation>
). Lateral margin of paraterga with small / tiny dentations or notches (Figs
<figureCitation id="9411B93EE652B8260EB37AC3DBAE8355" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21</figureCitation>
,
<figureCitation id="743952980C718A396521DF7E53447497" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22</figureCitation>
).
</paragraph>
<paragraph id="157D70479DDFEA81B40F70AC56A1EF2E">
Telson (Figs
<figureCitation id="F6304C1F2868C2D8CF96A0C5C5D033D3" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 E, F</figureCitation>
,
<figureCitation id="645B1610D76EEEA2906A3B7812CC3A87" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 E, F</figureCitation>
) short, caudal margin rounded; epiproct with 2 + 2 small / tiny tubercles at caudal margin. Paraprocts and hypoproct semi-circular.
</paragraph>
<paragraph id="B7D5ED95B2906C12A0CE122BBE54228A">
Legs slender, shorter than body ring width together with paraterga, terminating before lateral paratergal margins. Claws normal. Coxal sacs strong, present on body rings 332 (Fig.
<figureCitation id="A45C65EFC0F7A11606D3999279D56E68" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 F</figureCitation>
).
</paragraph>
<paragraph id="FD4D1A73AD25C48BEEDD16E1CD1A40A8">
<emphasis id="71358AAE26BE6B9CA49C13B0E1F88881" bold="true" italics="true">Gonopods</emphasis>
(Figs
<figureCitation id="0CF3380E799F78809638D68B2F5DD685" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. A. Left anterior gonopod, anterior view; B. Left posterior gonopod, sublateral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure23" httpUri="https://binary.pensoft.net/fig/1169913">23</figureCitation>
,
<figureCitation id="5A316F33988032BB255A4CF8FAA3F3D3" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. SEM. A, B. Left anterior gonopod, anterior view (A), ventral view (B); C. Distal part of left anterior gonopod, anterior view; D, E. Posterior gonopod and leg 11, ventral view (D), sublateral view (E); F. Distal part of posterior gonopod, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure24" httpUri="https://binary.pensoft.net/fig/1169914">24</figureCitation>
): Two pairs of gonopods directed anteriad. Anterior gonopods (Figs
<figureCitation id="E42736B52FCDAA425767618DBE69C676" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. A. Left anterior gonopod, anterior view; B. Left posterior gonopod, sublateral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure23" httpUri="https://binary.pensoft.net/fig/1169913">23 A</figureCitation>
,
<figureCitation id="27CC88799850710E6DEF20EB79F013DB" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. SEM. A, B. Left anterior gonopod, anterior view (A), ventral view (B); C. Distal part of left anterior gonopod, anterior view; D, E. Posterior gonopod and leg 11, ventral view (D), sublateral view (E); F. Distal part of posterior gonopod, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure24" httpUri="https://binary.pensoft.net/fig/1169914">24 A C</figureCitation>
) 6 - segmented, covered with long setae; coxite (
<emphasis id="18C9E09C5ED8B1E9A307DFC58529B498" italics="true">co</emphasis>
) broad, basal part sparsely covered with microgranulations; podomeres 25, short and stout; podomere 6 slightly longer than other podomeres, distally carrying macrosetae (
<emphasis id="C194DD37F0BCBC703C5F6C5A954C166F" italics="true">mse</emphasis>
). Posterior gonopods (Figs
<figureCitation id="1A3589A9A5982CEAAC35BF2061E6E56A" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. A. Left anterior gonopod, anterior view; B. Left posterior gonopod, sublateral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure23" httpUri="https://binary.pensoft.net/fig/1169913">23 B</figureCitation>
,
<figureCitation id="ABED24E3CA8C4256C57501F56A056498" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. SEM. A, B. Left anterior gonopod, anterior view (A), ventral view (B); C. Distal part of left anterior gonopod, anterior view; D, E. Posterior gonopod and leg 11, ventral view (D), sublateral view (E); F. Distal part of posterior gonopod, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure24" httpUri="https://binary.pensoft.net/fig/1169914">24 D F</figureCitation>
) 6 - segmented, covered with sparse long setae; coxite (
<emphasis id="2E71C5A71B095DA17B2D23A0E58BD7BD" italics="true">co</emphasis>
) broad, basal part sparsely covered with microgranulations; podomeres 25 short and stout; ultimate podomere (
<emphasis id="E927E08799E0082881A62B3B0C6516EF" italics="true">po 6</emphasis>
) longest, with three apical stylets (
<emphasis id="B69FC763E72295105C58AA4F7F416337" italics="true">sty</emphasis>
).
</paragraph>
<caption id="0A9E9451888D3018FEB7EFC98A7B3701" ID-DOI="10.3897/zse.100.132111.figure23" ID-arpha="D64B6B27-019F-54C0-BD85-2EE09DADBA95" httpUri="https://binary.pensoft.net/fig/1169913" startId="F23">
<paragraph id="169493012E1D3E5A2E220BC8CB922CAA">
<label id="FAED3053614D83ACD178D56E7EF70E3C">Figure 23.</label>
</paragraph>
<paragraph id="24B7FEE99E669F454AA47E2D70DEAF16">
<taxonomicName id="41DAC60934412D93DAB6BF0DAC9EF8D9" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="90588EE9C819FC6FDD456EEF990341BF" italics="true">Pseudodesmus irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C15E21951480CFB98C7752064190DBC5" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EVGAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
958 H), gonopods.
<emphasis id="8AD1AC9DA8BD065DD1FE38BD48CDFBA3" bold="true">A.</emphasis>
Left anterior gonopod, anterior view;
<emphasis id="2C91F3C24C7C0FDF3BB83664DFD11C41" bold="true">B.</emphasis>
Left posterior gonopod, sublateral view.
<emphasis id="74A386EE11228B0D631B0AA9F424F409" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="65CF122E46A0129575630D8B84033AE9" ID-DOI="10.3897/zse.100.132111.figure24" ID-arpha="A4B4F4D5-C781-5AC6-8AD4-108C2D0F22D0" httpUri="https://binary.pensoft.net/fig/1169914" startId="F24">
<paragraph id="9DB5F4399FF8C0C96415BE7B0538E34C">
<label id="0BF3673C9F181185B3A8B92AA687CA99">Figure 24.</label>
</paragraph>
<paragraph id="6FAD5032C28F18B8573F5FDAB4B9B01B">
<taxonomicName id="1D6E3BB144488C22CFE65F21098A9520" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
<emphasis id="47CB23D8B9085AC4FE2EA3405A508519" italics="true">Pseudodesmus irregularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6207951DE2B1C91A920EBBD6A2EE87C5" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EYHAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
958 H), gonopods.
<abbrev id="ABBRID0E4HAI" xlink_title="Scanning electron microscope">SEM</abbrev>
.
<emphasis id="AEB12D2C8E6F9A676EB99B706EA822C4" bold="true">A, B.</emphasis>
Left anterior gonopod, anterior view (
<emphasis id="3C6CD6C7B689E34D35CB36D3EBD8B60C" bold="true">A</emphasis>
), ventral view (
<emphasis id="3CBFC0CD2816C906B4E9BB00BE977B31" bold="true">B</emphasis>
);
<emphasis id="D069726F37FF74BA24DDA483840EDD30" bold="true">C.</emphasis>
Distal part of left anterior gonopod, anterior view;
<emphasis id="A395C70ED5417056DC26EA1CF47254EB" bold="true">D, E.</emphasis>
Posterior gonopod and leg 11, ventral view (
<emphasis id="CAFBA0D61C770A92EC8489D92B5774CF" bold="true">D</emphasis>
), sublateral view (
<emphasis id="44C70B8573E45103262994307FB5B288" bold="true">E</emphasis>
);
<emphasis id="F38311C5CD238CEE91FD1BF534BE73F8" bold="true">F.</emphasis>
Distal part of posterior gonopod, ventral view.
<emphasis id="4DF3BEF8A65C2E41F906B17DB1E28EBB" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EYIAI" type="variation">
<paragraph id="7CA18905F3F2EFE44AEF85327AC6E5AA">
<heading id="674365FE2F049A5BF622AD541AECC742" reason="title">Variation.</heading>
</paragraph>
<paragraph id="7AB555462AEEAC565A2B9C3670E18295">
<emphasis id="279C0EABE33C2CECED678AE70A3BC6DA" bold="true" italics="true">
Male
<typeStatus id="82179F7139BA2F95B45D06AB265B6E12" type="paratype">paratype</typeStatus>
</emphasis>
(
<emphasis id="C31713F87114B948F6A4BF2741AD9D25" bold="true">
<abbrev content-type="institution" id="ABBRID0EEJAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
958 P
</emphasis>
): 36 body rings plus telson, length about
<quantity id="3FF6D7CB536994CE66883865AC7B2B14" metricMagnitude="-3" metricUnit="m" metricValue="9.75" unit="mm" value="9.75">9.75 mm</quantity>
, length of midbody metazoan ca.
<quantity id="DBE0294014BC6B6F769ED43BA5A42C96" metricMagnitude="-4" metricUnit="m" metricValue="1.4" unit="mm" value="0.14">0.14 mm</quantity>
, width of midbody metazona ca.
<quantity id="D1346F46D89806C6E83A593722453E90" metricMagnitude="-3" metricUnit="m" metricValue="1.72" unit="mm" value="1.72">1.72 mm</quantity>
, width of midbody prozona ca.
<quantity id="B1BE29736A3B6FDE7D07EB569E47AEEF" metricMagnitude="-4" metricUnit="m" metricValue="8.2" unit="mm" value="0.82">0.82 mm</quantity>
; ratio of width of head and width of collum = 0.85.
</paragraph>
<paragraph id="EB0D102C15C2FBCBF5B44CBF3B09B284">
The female from Ngoc Linh Mts (
<emphasis id="F8E5FE013525FBE1BCB91C98649FCCB8" bold="true">
<abbrev content-type="institution" id="ABBRID0ENJAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
960
</emphasis>
) has 41 body rings plus telson; midbody width
<quantity id="32BF92EC18ABF8E97352406E0E521A8E" metricMagnitude="-3" metricUnit="m" metricValue="2.1" unit="mm" value="2.1">2.1 mm</quantity>
, length about
<quantity id="111A9C5A1043F49C515C259DBA9DB702" metricMagnitude="-2" metricUnit="m" metricValue="1.011" unit="mm" value="10.11">10.11 mm</quantity>
. Coloration with widening pattern repeated between 712, 1317, 1824, 2531, 3236 body rings (see also
<bibRefCitation id="001F83566BAA4F219CEAC754D11CE1CF" DOI="10.3897/ijm.6.1949" author="Enghoff" firstAuthor="Enghoff" journalOrPublisher="International Journal of Myriapodology" pagination="1-27" refId="B10" refString="Enghoff H (2011) Trans-segmental serial colour patterns in millipedes and their developmental interpretation (Diplopoda). International Journal of Myriapodology 6: 127. https://doi.org/10.3897/ijm.6.1949" title="Trans-segmental serial colour patterns in millipedes and their developmental interpretation (Diplopoda)." volume="6" year="2011">Enghoff 2011</bibRefCitation>
). Tubercles are small: on collum only one row, on 24 two weak rows (reaching only one row on paraterga), from body ring 5 onwards two rows, each 1 + 1 large tubercles in the middle, then anterior row 1013 small tubercles reaching edge of paraterga, in posterior row only 34 small tubercles, on the last five body rings anterior row is weaker but still with 810 small tubercles, in posterior row the median tubercles larger, with 4 small tubercles laterally.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXJAI" type="etymology">
<paragraph id="59292F5DDA7E4E6C03B4BD1FFBC6D850">
<heading id="B43539290021D2B67EC8ABD0BC834EAA" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="55BF7134A4914D8D860ACB97AC6E1E1F">To emphasize the irregular pattern of terga. Noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="SECID0E3JAI" type="distribution">
<paragraph id="68BD8110F705332DDEF24B5691E9FB8F">
<heading id="A81E860EB629FF31AA584E90816B503C" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="82349D04E083289A86E7E4E20DB3DF52">
The species has been only known from
<collectingRegion id="44A9A4EF17F489B324C39A30933F2625" country="Vietnam" name="Khanh Hoa">Khanh Hoa</collectingRegion>
and
<collectingRegion id="62020231B084070407464CB77E6BDF8E" country="Vietnam" name="Kon Tum">Kon Tum Province</collectingRegion>
, southcentral
<collectingCountry id="60FB2A2F77CDA04F014C0088F027E93E" name="Vietnam">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBKAI" type="DNA barcoding">
<paragraph id="929C51424D8580806C148122061687E1">
<heading id="7352C28E3BF4D9DF47D0F6F4D7E121FC" reason="title">DNA barcoding.</heading>
</paragraph>
<paragraph id="A9A7A0D2975CBEC53CA47E51BDD4BF0E">
Sequencing a fragment of the
<abbrev id="ABBRID0EHKAI" xlink_title="cytochrome c oxidase subunit I">COI</abbrev>
gene failed.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,498 @@
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<mods:title id="628F166086A624C283A1C9B44133BB3B">A review of the millipede genus Pseudodesmus Pocock, 1887 (Diplopoda, Platydesmida, Andrognathidae) from Vietnam, with descriptions of five new species and notes on its phylogeny</mods:title>
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<mods:namePart id="795D6CCF69693DF3783A49AC357D2D00">Nguyen, Anh D.</mods:namePart>
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<mods:affiliation id="4CF5EDDC48AD589E762E04CE4656BA3E">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="32DE3EE068788AFCD87553C19CC15B59">Nguyen, Lien T. P.</mods:namePart>
<mods:affiliation id="19B523CB0B3BD600A884B03C755D0429">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="E84A18376476D92B0E691EC44F97FD4C">Korsós, Zoltán</mods:namePart>
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<mods:affiliation id="D56F8F15D416442F37E17791F121477A">University of Veterinary Medicine Budapest, István u. 2, 1078 Budapest, Hungary</mods:affiliation>
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<treatment id="AEB045681A0F51989153C08ABF220F73" ID-DOI="http://doi.org/10.5281/zenodo.14052224" ID-Zenodo-Dep="14052224" ID-arpha="AEB04568-1A0F-5198-9153-C08ABF220F73" ID-zoobank="https://zoobank.org/3132C8AB-185A-4282-873E-274B50C0ECC7" LSID="urn:lsid:plazi:treatment:AEB045681A0F51989153C08ABF220F73" LSID-ZBK="urn:lsid:zoobank.org:act:3132C8AB-185A-4282-873E-274B50C0ECC7" httpUri="http://treatment.plazi.org/id/AEB045681A0F51989153C08ABF220F73">
<subSubSection id="287AE4E9240235D4F913442E124DCC2C" type="nomenclature">
<paragraph id="B363C50EE9EC712349D772FDEC5F842F">
<taxonomicName id="275618BB2BAF525E917942ADDE3972BB" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Playtdesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov.">
<emphasis id="943D21F583B0B087325439747D97BDE4" italics="true">Pseudodesmus ngoclinh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CA6F65EF2D192C94D532D6104833F75" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="EBEF41B6F782C6BFB168C874B3117678" type="description">
<paragraph id="FF3672C20D1F26175FBF10A6522DA29D">
<figureCitation id="6005FC18A1431C2538111E43E748618A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">Figs 6</figureCitation>
,
<figureCitation id="003E339EF2C44379398FE7206FB60357" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A. Anterior part of body, dorsal view; B. Head and collum, anterior view; C. Anterior half of body, ventral view; D. Anterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure7" httpUri="https://binary.pensoft.net/fig/1169897">7</figureCitation>
,
<figureCitation id="C0DD73C30D1160F4533CD38D028441D2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A, B. Midbody body rings, dorsal view (A), ventral view (B); C, D. Posterior half of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure8" httpUri="https://binary.pensoft.net/fig/1169898">8</figureCitation>
,
<figureCitation id="84EF704B70F67BB8D758F6A32A1C57E5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), gonopods. A. Left anterior gonopod, ventral view; B. Posterior gonopods, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure9" httpUri="https://binary.pensoft.net/fig/1169899">9</figureCitation>
,
<figureCitation id="9BFEAE3FA1928E99ECC017875DB154B1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), gonopods. SEM. A, B. Left anterior gonopod, ventral view (A), posterior view (B); C, D. Posterior gonopods, ventral view (C), anterior view (B). Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure10" httpUri="https://binary.pensoft.net/fig/1169900">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENAAG" type="material">
<paragraph id="440E4A807A20F0922A2EEC592F7378D9">
<heading id="9977C2667D028100CD1B7B327FBF5E8F" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="3D888D5B2B2511B27545490B3D4B9023">
<materialsCitation id="6C0ECA48653030664DAAA3AEC3BDF43C" collectingDate="2006-03-21" collectingDateMax="2006-04-09" collectingDateMin="2006-03-21" collectorName="Anh D. Nguyen" country="Vietnam" elevation="1850" latitude="15.07134" location="Ngoc Linh Mts" longLatPrecision="1" longitude="107.97158" specimenCount="1" specimenCount-male="1" stateProvince="Kon Tum" typeStatus="holotype">
<emphasis id="BF862DAFF27D17A45E458F2FB88D5A85" bold="true" italics="true">
<typeStatus id="B21AF6C9DF98BF10795495011C67FA93" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="71929D48A3A0CA23C8F14D088521E5D0" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="945E0C17E1664E0B9DA03189328CD748" count="1" type="male">1 male</specimenCount>
;
<collectingRegion id="6A4191CEF1D20DC024EF612AF53E901B" country="Vietnam" name="Kon Tum">Kon Tum Province</collectingRegion>
,
<location id="C5084DDF9806459CE63FB3CC75256604" LSID="urn:lsid:plazi:treatment:AEB045681A0F51989153C08ABF220F73:C5084DDF9806459CE63FB3CC75256604" country="Vietnam" latitude="15.07134" longLatPrecision="1" longitude="107.97158" name="Ngoc Linh Mts" stateProvince="Kon Tum">Ngoc Linh Mts</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2AAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.971580,15.071340]}">
<geoCoordinate id="A9AFD4F5084D56159B4B1A0EE17C12CF" degrees="15.07134" direction="north" orientation="latitude" precision="1" value="15.07134">15.07134 ° N</geoCoordinate>
,
<geoCoordinate id="925A5ADEA4A4E7A86CC8B851574880CB" degrees="107.97158" direction="east" orientation="longitude" precision="1" value="107.97158">107.97158 ° E</geoCoordinate>
</named-content>
;
<elevation id="1865D758E3AB4B02F2589E83DCE1EFC7" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">
<quantity id="9DB8B26A005394F28C3F55C249CC07F7" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1,8001,900 m</quantity>
a. s. l.
</elevation>
;
<date id="1CB1C934B49BB0033776112F999CC252" value="2006-03-21" valueMax="2006-04-09" valueMin="2006-03-21">
<collectingDate id="C174C2D2D960DEC03D647AC648E926F7" value="2006-03-21" valueMax="2006-04-09" valueMin="2006-03-21">21 Mar. 9 Apr. 2006</collectingDate>
</date>
;
<collectorName id="ED1D41D5E4906D37D81B310507E0A4A9">Anh D. Nguyen</collectorName>
leg.; primary forest, decaying wood, under the bark;
<abbrev content-type="institution" id="ABBRID0EABAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
986 H.
</materialsCitation>
</paragraph>
<paragraph id="A825E0BFA0CD5268CE22547FE33CA3F5">
<materialsCitation id="B9ED2375A1715D1E597A75F8B2F388C5" collectingDate="2006-03-21" collectingDateMax="2006-04-09" collectingDateMin="2006-03-21" collectorName="Anh D. Nguyen" country="Vietnam" elevation="1850" latitude="15.07134" location="Ngoc Linh Mts" longLatPrecision="1" longitude="107.97158" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Kon Tum" typeStatus="paratype">
<emphasis id="0BE03251DEAE4288AEA008E38A8C04FF" bold="true" italics="true">
<typeStatus id="48FBC3FF2DB04C579D5EFEA2B9072B4C" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="ECADBCD4FBFBC5B297A7BB43B94BE69A" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="5867760437E7003AD02839AE63354B4F" count="1" type="male">1 male</specimenCount>
,
<specimenCount id="EDE0A315CC5FB68D315B86EC5651C4C6" count="2" type="female">2 females</specimenCount>
; same data as for the holotype;
<abbrev content-type="institution" id="ABBRID0EMBAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
986 P
</materialsCitation>
<materialsCitation id="A9297D4ACE87F22BAF641FD4CF1B1B6B" collectingDate="2006-03-21" collectingDateMax="2006-04-09" collectingDateMin="2006-03-21" collectionCode="HNHM" collectorName="Anh D. Nguyen" country="Vietnam" elevation="1850" latitude="15.07134" location="Ngoc Linh Mts" longLatPrecision="1" longitude="107.97158" specimenCount="2" specimenCount-female="2" stateProvince="Kon Tum" typeStatus="paratype">
<specimenCount id="B2173A0A2D26C96E8C416C2073DC0642" count="2" type="female">2 females</specimenCount>
; same data as for the holotype;
<named-content id="5E4A09308EB206ED5A19A822D405962B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum-termeszettudomanyi-muzeum" xlink_title="Department of Zoology, Hungarian Natural History Museum, Budapest, Hungary">
<collectionCode id="E6C686E8D8FF6D071F72589930467A33" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWBAG" type="diagnosis">
<paragraph id="5D6FC9D81F868C453F35BE3F3FF934E5">
<heading id="FCCC046523E1657D1BDC749A2528F882" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="2DA22BC225A6A6AADD92B41E0B8F5A87">Head slightly broader than collum. Collum with distinct paraterga directed anterad, with two rows of 1 + 1 smaller and 2 + 2 larger tubercles. Body covered with dense minute pubescence. Midbody body rings with two rows of tubercles on metazona, posterior row of 45 + 4 5 tubercles, shorter and extending to base of paraterga; anterior row of 510 + 510 small tubercles, longer, extending to about midlength of paraterga. Posterior gonopods 6 - segmented, with two apical stylets.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2BAG" type="diagnosis">
<paragraph id="378247185024E4BB7A5673D7EC3D7A67">
<heading id="488A7908BF469479C24000F81916167E" reason="title">Diagnosis remarks.</heading>
</paragraph>
<paragraph id="5F0B35008497E083C5BCD1718F8D1F63">
The new species is easily distinguished from
<taxonomicName id="7D06F3EA5643A6B0BF1E34B8C7FE3CAB" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="camptotrichus">
<emphasis id="8EA77FFB60A099E12E4DACF55CA9778A" italics="true">P. camptotrichus</emphasis>
</taxonomicName>
by coloration (yellowish / brownish plus dark spots vs whole earth brown). It also differs from
<taxonomicName id="AA0082F55B7B6A0CEEE547982B57D856" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="C744F66C838E90B4CD4C76E0F8912FB2" italics="true">P. variegatus</emphasis>
</taxonomicName>
and
<taxonomicName id="72234C39C315E0CE2753EFD822F208DB" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="camptotrichus">
<emphasis id="219DC931F56C8AC3203F2D2FE1B5A41D" italics="true">P. camptotrichus</emphasis>
</taxonomicName>
in having smaller size (length:
<quantity id="57D94967898E4C778323DAA8D4FF9FD4" metricMagnitude="-2" metricUnit="m" metricValue="1.14" unit="mm" value="11.4">11.4 mm</quantity>
vs
<quantity id="E43F4FFAA3D14918288928B7E0FAAD3C" metricMagnitude="-2" metricUnit="m" metricValue="3.2" unit="mm" value="32.0">32 mm</quantity>
and
<quantity id="F09232F8B0FBAC8BDFE064D41A67BBA9" metricMagnitude="-2" metricUnit="m" metricValue="1.4" unit="mm" value="14.0">14 mm</quantity>
; width:
<quantity id="1B81AE0540AF8DA0A8AFA6AC45A46678" metricMagnitude="-3" metricUnit="m" metricValue="2.74" unit="mm" value="2.74">2.74 mm</quantity>
vs
<quantity id="8C5218718B31DBF60387E94CD2F427A8" metricMagnitude="-3" metricUnit="m" metricValue="5.8" unit="mm" value="5.8">5.8 mm</quantity>
and 2.0 mm, respestively), number of metazonal tubercles (two rows of 510 + 510 and 45 + 4 5 vs two rows of 815 + 8 15 and 57 + 5 7, and two rows of 6 + 6 to 9 + 9 each row, respectively).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECDAG" type="description">
<paragraph id="C936A0D7B3A3BFFC3559182D031BE631">
<heading id="BB89957BBD8733E2D185B32D67848806" reason="title">Description.</heading>
</paragraph>
<paragraph id="749DF58DC0B15CA9362E22EFB0D2568A">
<emphasis id="F8754BFDC4C1C60E1878D149D0C6EF95" bold="true">
Male
<typeStatus id="D93832348172EA2C7804017BE27F15A9" type="holotype">holotype</typeStatus>
.
</emphasis>
</paragraph>
<paragraph id="521C4AD1B3BE5FED510892CD9013B92E">
<emphasis id="1CFE9710BA229712E0C0EAC2EC50A61D" bold="true" italics="true">Measurements</emphasis>
: 51 body rings plus telson; length about
<quantity id="089A4850DB5538DA5BE825E6B701C58E" metricMagnitude="-2" metricUnit="m" metricValue="1.14" unit="mm" value="11.4">11.40 mm</quantity>
, length of midbody metazona about
<quantity id="199CDC8A047E7C68CA14ECB6C1EC1177" metricMagnitude="-4" metricUnit="m" metricValue="1.9" unit="mm" value="0.19">0.19 mm</quantity>
, width of midbody metazona about
<quantity id="AEEAA0DE52748F4CD91605B2DCEAA17B" metricMagnitude="-3" metricUnit="m" metricValue="2.74" unit="mm" value="2.74">2.74 mm</quantity>
, width of midbody prozona about
<quantity id="2BFD6ABB03B53CDE94B7FE268553011C" metricMagnitude="-3" metricUnit="m" metricValue="1.26" unit="mm" value="1.26">1.26 mm</quantity>
; ratio of width of head and width of collum = 1.05.
</paragraph>
<paragraph id="78B47408F98BF08210E88833739B5D64">Coloration of ethanol-preserved specimens: first 6 body rings including collum yellowish in the middle, then most body rings with a light median section, dark spots (on tubercles) irregularly on one side or both sides, dark spots on paraterga, asymmetrically, and almost every body ring different, sometimes repeating 34 body rings, last 56 body rings uniformly brownish.</paragraph>
<paragraph id="08ED568ABA4D125F956A5A20E12C70FF">
Head (Figs
<figureCitation id="B26CC3F43FB93E333967A276891C6FB6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">6 A, B, F</figureCitation>
,
<figureCitation id="68CD10121F3C3BB191109C54D76CE514" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A. Anterior part of body, dorsal view; B. Head and collum, anterior view; C. Anterior half of body, ventral view; D. Anterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure7" httpUri="https://binary.pensoft.net/fig/1169897">7 A D</figureCitation>
) round-shaped, moderately densely setose. Antenna stout, clavate,
<emphasis id="0F1C0A1B2E79280C6C2273DA5868C026" italics="true">in situ</emphasis>
reaching to body ring 5; antennomere 6&gt; 5&gt; 3&gt; 4 = 2&gt; 7 =
<quantity id="01979653B2E0B2673A7FF8B826971E54" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
length. Collum with distinct paraterga directed anterad, with two rows: anterior row with 1 + 1 tubercles, posterior row with 2 + 2 tubercles, paramedian ones being larger (Figs
<figureCitation id="CDFBDABC3955C45696337D16B981544A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">6 A</figureCitation>
,
<figureCitation id="510BECC6B16E29ECDAE747F5154AFA78" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A. Anterior part of body, dorsal view; B. Head and collum, anterior view; C. Anterior half of body, ventral view; D. Anterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure7" httpUri="https://binary.pensoft.net/fig/1169897">7 A, B</figureCitation>
). Body covered with dense minute pubescence. Slopes of dorsum steep, so body in cross-section subtriangular. Body rings 24 with a single row, each of 1 + 2, 1 + 1, 3 + 4 tubercles, respectively (Figs
<figureCitation id="1F8C2BCDF89E4612D33C5B6F4795063E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">6 A, C</figureCitation>
,
<figureCitation id="9E442F77B4EEEB9C01D668DE8BC80658" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A. Anterior part of body, dorsal view; B. Head and collum, anterior view; C. Anterior half of body, ventral view; D. Anterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure7" httpUri="https://binary.pensoft.net/fig/1169897">7 A</figureCitation>
). Midbody body rings with two rows of tubercles on metazona, posterior row of 45 + 4 5 tubercles, shorter and extending to base of paraterga; anterior row of 510 + 510 small tubercles, longer, extending to about midlength of paraterga. Paramedian tubercles considerably larger, and posterior paramedian tubercles much larger than anterior paramedian ones (Figs
<figureCitation id="1AF7CAAD0A39878FD638B2AF88DEB2EE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">6 D</figureCitation>
,
<figureCitation id="67958B15144B9611F978B89EB5BA3BA3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A, B. Midbody body rings, dorsal view (A), ventral view (B); C, D. Posterior half of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure8" httpUri="https://binary.pensoft.net/fig/1169898">8 A</figureCitation>
,
<figureCitation id="A3CE06BCB004A0FDFA4F9691954F89FF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A, B. Midbody body rings, dorsal view (A), ventral view (B); C, D. Posterior half of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure8" httpUri="https://binary.pensoft.net/fig/1169898">8 C</figureCitation>
). The last 6 body rings with only a row of 4 + 4 tubercles (Figs
<figureCitation id="1FBFB051F1AB78D384DB568AEF03BF98" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">6 E</figureCitation>
,
<figureCitation id="300B36DF562186D97EF03C0A0A5F7F60" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A, B. Midbody body rings, dorsal view (A), ventral view (B); C, D. Posterior half of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure8" httpUri="https://binary.pensoft.net/fig/1169898">8 C</figureCitation>
).
</paragraph>
<caption id="0AAB103706EC0104CA232E39E99CFEDB" ID-DOI="10.3897/zse.100.132111.figure6" ID-arpha="C1390437-C36A-5351-ABAE-E44E3D7DCCE5" httpUri="https://binary.pensoft.net/fig/1169896" startId="F6">
<paragraph id="1DE507676918F53B60C88CD5F031EBCA">
<label id="560EE35B6238D91BE51903A2763AEB92">Figure 6.</label>
</paragraph>
<paragraph id="9063C9E8D6C4FD0799BE88F1BB9B5B7C">
<taxonomicName id="44BA50F13500F0C3DDB087A88F64130E" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov.">
<emphasis id="7A05A1E346C1F7F04AE626E1BA1EDEA1" italics="true">Pseudodesmus ngoclinh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46FB425C6E1F2F806F3CF798E945471A" rank="species">sp. nov</taxonomicNameLabel>
, holotype (
<abbrev content-type="institution" id="ABBRID0EVFAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
986 H), under white light.
<emphasis id="CA76BDD59123156FF828E1B3D4682EE6" bold="true">A C.</emphasis>
Anterior part of body, dorsal view (
<emphasis id="6ED3F119D7CE9177081DDDC9F41A2752" bold="true">A, C</emphasis>
), ventral view (
<emphasis id="C3511F074B4AFDFD34B32805C343C541" bold="true">B</emphasis>
);
<emphasis id="E4A8293FA24370BFDF4C867603E0AE3E" bold="true">D.</emphasis>
Midbody body rings, ventral view;
<emphasis id="8CF798B58C76836B5C24F1F3D545074E" bold="true">E.</emphasis>
Posterior part of body dorsal view;
<emphasis id="F292336EB86C1D053127E30CB47EA1E6" bold="true">F.</emphasis>
Anterior half of body, ventral view;
<emphasis id="4E35F00C9D985912147B8C0F3CBCD75A" bold="true">G.</emphasis>
Posterior half of body, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="30369B9012564486DC3793C4D393A04A" ID-DOI="10.3897/zse.100.132111.figure7" ID-arpha="7561D817-3254-53A1-A97D-2CD30DF8CFE8" httpUri="https://binary.pensoft.net/fig/1169897" startId="F7">
<paragraph id="F3494E02861F72890EB9759F4223E6DE">
<label id="7CB19FF206523C9B9E5C79D33C718008">Figure 7.</label>
</paragraph>
<paragraph id="B1588093B69B0DD0A0CE37BDC0F751A1">
<taxonomicName id="416F83B7DEC836708CA491FCF8C072A2" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov.">
<emphasis id="EA9E3FB314D5826A82D8EB51002D1AB8" italics="true">Pseudodesmus ngoclinh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79F095D412DE68CF149D4BF80A44B4FC" rank="species">sp. nov</taxonomicNameLabel>
, holotype (
<abbrev content-type="institution" id="ABBRID0EAHAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
986 H), under UV light.
<emphasis id="2E404A452DA139E548958CFB2B620DC4" bold="true">A.</emphasis>
Anterior part of body, dorsal view;
<emphasis id="035F4F02492B6225C39F4A9E39A0102D" bold="true">B.</emphasis>
Head and collum, anterior view;
<emphasis id="BCA1B5F977FF428D6AEFAE371630321F" bold="true">C.</emphasis>
Anterior half of body, ventral view;
<emphasis id="55681C60F4AF3D864FEEFFA8F1811257" bold="true">D.</emphasis>
Anterior part of body, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="B357E0F3CAD6337C4685F2918DD79A69" ID-DOI="10.3897/zse.100.132111.figure8" ID-arpha="67C85753-0AF8-507D-8DC4-9BE44FEFBBC2" httpUri="https://binary.pensoft.net/fig/1169898" startId="F8">
<paragraph id="309F9847DC333D49A76AF3EF89622096">
<label id="456D90ADB91A01D0CE380106CCC742A2">Figure 8.</label>
</paragraph>
<paragraph id="C9501766C809A08B1854C74A889BC222">
<taxonomicName id="9858199837C91A3CBFB0034A85DFA127" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov.">
<emphasis id="B9567869C6924A7EDD11FE0FE02F3A03" italics="true">Pseudodesmus ngoclinh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CDFF31B7DF8D55567F30A27FBCB75AD" rank="species">sp. nov</taxonomicNameLabel>
, holotype (
<abbrev content-type="institution" id="ABBRID0EFIAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
986 H), under UV light.
<emphasis id="B46E6DC51781604B73EC1DC95C8BD557" bold="true">A, B.</emphasis>
Midbody body rings, dorsal view (
<emphasis id="4DDA55CFD9B8F05F2985DB818564AA54" bold="true">A</emphasis>
), ventral view (
<emphasis id="4DBB18A307F0BC3B68162A5794893755" bold="true">B</emphasis>
);
<emphasis id="D275E5F2D5FF368A5B1C7ACFDD0743B6" bold="true">C, D.</emphasis>
Posterior half of body, dorsal view (
<emphasis id="14A6F843F35555362DCA863087BD21C4" bold="true">C</emphasis>
), ventral view (
<emphasis id="4C21C0CF2CF72D3EAC6A95C5FD32272E" bold="true">D</emphasis>
). Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="5BB4D45CE3057DC9199AB269F41EBE8A">
Paraterga curved strongly anteriad on body rings 25, increasingly less so on body rings 610 (to body ring 11 on
<typeStatus id="11F70651EA58DFC2AD7F5437E8F15A29" type="paratype">paratype</typeStatus>
specimens), slightly curved caudad on body ring 11 (to body ring 12 on
<typeStatus id="4A657568495AB673AFCC8D8CA94EA422" type="paratype">paratype</typeStatus>
specimens) and following body rings; caudal curvature of paraterga increasingly clear on 67 posteriormost body rings in front of telson; paraterga of penultimate body ring produced strictly caudad and flanking telson (Figs
<figureCitation id="842CFD9AAF14A442CE652BD84644CE1B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">6 E, G</figureCitation>
,
<figureCitation id="B6CADF91CF66C244CF6786C3549B6D03" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A, B. Midbody body rings, dorsal view (A), ventral view (B); C, D. Posterior half of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure8" httpUri="https://binary.pensoft.net/fig/1169898">8 C, D</figureCitation>
). Caudal paratergal margins entire, without notches (Figs
<figureCitation id="0FAF5F876A9291F3CDAA27280306CACC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">6</figureCitation>
<figureCitation id="D17F1A4A3C82A7176CF2841FC866AF91" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A, B. Midbody body rings, dorsal view (A), ventral view (B); C, D. Posterior half of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure8" httpUri="https://binary.pensoft.net/fig/1169898">8</figureCitation>
).
</paragraph>
<paragraph id="C7558D3E07FBD70AEC5DB4F9411E731C">
Telson (Figs
<figureCitation id="97DE9AB9146026A2D807B6D1AACF6317" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">6 C</figureCitation>
,
<figureCitation id="05C1412186F1B32DB9E51DCA02A61A69" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A, B. Midbody body rings, dorsal view (A), ventral view (B); C, D. Posterior half of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure8" httpUri="https://binary.pensoft.net/fig/1169898">8 C, D</figureCitation>
) short, caudal margin rounded; epiproct with 2 + 2 small / tiny tubercles at caudal margin. Paraprocts and hypoproct semi-circular.
</paragraph>
<paragraph id="075F8FE456A5A3367D75C0426A443615">
Legs slender, shorter than width of body ring together with paraterga, terminating before lateral paratergal margins. Prefemur = tarsus&gt; femur&gt; postfemur = tibia in length. Claws normal. Coxal sacs present on body rings 341 (Figs
<figureCitation id="8C20BC61B96F81E3F325625624B78DDF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under white light. A C. Anterior part of body, dorsal view (A, C), ventral view (B); D. Midbody body rings, ventral view; E. Posterior part of body dorsal view; F. Anterior half of body, ventral view; G. Posterior half of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure6" httpUri="https://binary.pensoft.net/fig/1169896">6 G</figureCitation>
,
<figureCitation id="8775C167B9A4B78D8ED038A3207FF737" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), under UV light. A, B. Midbody body rings, dorsal view (A), ventral view (B); C, D. Posterior half of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure8" httpUri="https://binary.pensoft.net/fig/1169898">8 B</figureCitation>
).
</paragraph>
<paragraph id="DED5DEDA4439409FA03B136D8392CAC9">
<emphasis id="DBC05FD0B99D8D38C4C63ACF4E6BFD84" bold="true" italics="true">Gonopods</emphasis>
: Two pairs of gonopods directed mesoanteriad. Anterior gonopods (Figs
<figureCitation id="26D27B35B74E1DCACC344DCC1632D050" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), gonopods. A. Left anterior gonopod, ventral view; B. Posterior gonopods, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure9" httpUri="https://binary.pensoft.net/fig/1169899">9 A</figureCitation>
,
<figureCitation id="03A14E24E79F39D449B4E35CAE73C768" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), gonopods. SEM. A, B. Left anterior gonopod, ventral view (A), posterior view (B); C, D. Posterior gonopods, ventral view (C), anterior view (B). Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure10" httpUri="https://binary.pensoft.net/fig/1169900">10 A, B</figureCitation>
) 6 - segmented, covered with long setae; coxite broadly stout, basal part sparsely covered with microgranulations; podomeres 25, short and stout; podomere 6 slightly longer than other podomeres, distally carrying macrosetae (
<emphasis id="BB7619C75173EE8005EE42272684BF9C" italics="true">mse</emphasis>
). Posterior gonopods (Figs
<figureCitation id="F2F707DCB88F8FAEB91CE925FE005E75" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), gonopods. A. Left anterior gonopod, ventral view; B. Posterior gonopods, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure9" httpUri="https://binary.pensoft.net/fig/1169899">9 B</figureCitation>
,
<figureCitation id="83F6BB2E0E2EF390E40A39181699DBF1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Pseudodesmus ngoclinh sp. nov, holotype (IEBR-Myr 986 H), gonopods. SEM. A, B. Left anterior gonopod, ventral view (A), posterior view (B); C, D. Posterior gonopods, ventral view (C), anterior view (B). Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure10" httpUri="https://binary.pensoft.net/fig/1169900">10 C, D</figureCitation>
) 6 - segmented, covered with sparse long setae; coxite broad, basal part sparsely covered with microgranulations; podomeres 25 short and stout; ultimate podomere (
<emphasis id="B570C98F039CB6A3FAE2F81B934B3081" italics="true">po 6</emphasis>
) longest, with two apical stylets (
<emphasis id="179718817782926B6226653123872CAD" italics="true">sty</emphasis>
).
</paragraph>
<caption id="3B53255678701739501A4B0CE29D0D0C" ID-DOI="10.3897/zse.100.132111.figure9" ID-arpha="24BA9893-4169-56A9-A8DF-2FB7D1B537E5" httpUri="https://binary.pensoft.net/fig/1169899" startId="F9">
<paragraph id="B9DA6A62CC94244BDE2CB415E4E2C82D">
<label id="9FB535FD959CCC3F1E6E3AC4AE2E34EC">Figure 9.</label>
</paragraph>
<paragraph id="0378A1711472B77E85D6EBBD2D36F6D7">
<taxonomicName id="998DA6BDA65836B0F368C6B0CE504E18" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov.">
<emphasis id="7AEFF5F1124772A90BF5205F9AB916D4" italics="true">Pseudodesmus ngoclinh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="693390C7A0DA5FB5FE5DE8067E993036" rank="species">sp. nov</taxonomicNameLabel>
, holotype (
<abbrev content-type="institution" id="ABBRID0ERLAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
986 H), gonopods.
<emphasis id="8989876F9F0F7A5B37FF9913460500D0" bold="true">A.</emphasis>
Left anterior gonopod, ventral view;
<emphasis id="A0AA076ABE47E624338776C2251C595F" bold="true">B.</emphasis>
Posterior gonopods, anterior view.
<emphasis id="C8D8FA0F4116ED00E033B3742BB05C21" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="5A3A70E23DB19DFD06F34B089F0D60E4" ID-DOI="10.3897/zse.100.132111.figure10" ID-arpha="98F6F558-2D90-5D1C-A6C6-10489113CF36" httpUri="https://binary.pensoft.net/fig/1169900" startId="F10">
<paragraph id="08D1788FBFE8EE02F4AEB2EA4757308F">
<label id="F61EF8517ADCF06A74122F4C8C270AEA">Figure 10.</label>
</paragraph>
<paragraph id="D117BDC549C48FF2FC6946A3B768A7E5">
<taxonomicName id="0C499631B965106411208AF514E9FC80" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov.">
<emphasis id="E064220BE1D518CD8FA4E2E08409CC80" italics="true">Pseudodesmus ngoclinh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38FC4FDFAB2246EF6BFB759B76401DC" rank="species">sp. nov</taxonomicNameLabel>
, holotype (
<abbrev content-type="institution" id="ABBRID0EUMAG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
986 H), gonopods.
<abbrev id="ABBRID0EZMAG" xlink_title="Scanning electron microscope">SEM</abbrev>
.
<emphasis id="F099825E8A057CA441D8CD9CD2390D96" bold="true">A, B.</emphasis>
Left anterior gonopod, ventral view (
<emphasis id="E7A5D9D3E387E4D12A7887A480DA0244" bold="true">A</emphasis>
), posterior view (
<emphasis id="38FEAED6708E380D1C00FAA4D6E82AEB" bold="true">B</emphasis>
);
<emphasis id="BE7101ACA79BF229D952FC9B2803DB21" bold="true">C, D.</emphasis>
Posterior gonopods, ventral view (
<emphasis id="60D78BF5A5AE6B83AAABAE11C65DA0CC" bold="true">C</emphasis>
), anterior view (
<emphasis id="915EEB8FCBE61245AD5E8F933728DF77" bold="true">B</emphasis>
).
<emphasis id="695F8AE2FAC954E37F221BC73D625FF5" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQNAG" type="variation">
<paragraph id="1C42ECC066AA6DFFDC52C5B2CDE98ED8">
<heading id="A3473BB2CF0D7631F690BFD6F6C992A2" reason="title">Variation.</heading>
</paragraph>
<paragraph id="B02945FAA949A4452DCED0E4170938EC">
<emphasis id="F0E539D577E485A7C1F03C2FFFF832A2" bold="true" italics="true">
Male
<typeStatus id="F09715DEA68B1161958EB9A729AFF2D3" type="paratype">paratype</typeStatus>
</emphasis>
: 53 body rings + telson; length about
<quantity id="D962880A51B754D3616F960704D425D3" metricMagnitude="-2" metricUnit="m" metricValue="1.173" unit="mm" value="11.73">11.73 mm</quantity>
, length of midbody metazona about
<quantity id="FEF97ACAAF87C6A5C78AA4BDEE1C16A9" metricMagnitude="-4" metricUnit="m" metricValue="2.1" unit="mm" value="0.21">0.21 mm</quantity>
, width of midbody metazona about
<quantity id="5E8D54E9B425B2BE5E278347CB618D90" metricMagnitude="-3" metricUnit="m" metricValue="2.79" unit="mm" value="2.79">2.79 mm</quantity>
, width of midbody prozona about
<quantity id="2BB73284B4C45616B36AC86A445EEC47" metricMagnitude="-3" metricUnit="m" metricValue="1.28" unit="mm" value="1.28">1.28 mm</quantity>
.
</paragraph>
<paragraph id="A7C0F0F6C3C8B8A546330345BFF86E26">
<emphasis id="51551EEEFF2082DC179BC293F4236124" bold="true" italics="true">
Female
<typeStatus id="20729AFFFAFD12C707FF079FF430F4A0" type="paratype">paratypes</typeStatus>
.
</emphasis>
4851 body rings + telson. Length
<emphasis id="D2C54C8D37C6962E85C2BB298869B563" italics="true">ca</emphasis>
<quantity id="D375527BE5253418C98652799154BA8D" metricMagnitude="-2" metricUnit="m" metricValue="1.2865" metricValueMax="1.3519999999999999" metricValueMin="1.221" unit="mm" value="12.865" valueMax="13.52" valueMin="12.21">12.2113.52 mm</quantity>
, width of midbody metazona about
<quantity id="45DA66AC55B4CE61B322599F1F150B6B" metricMagnitude="-3" metricUnit="m" metricValue="2.9299999999999997" metricValueMax="3.02" metricValueMin="2.84" unit="mm" value="2.9299999999999997" valueMax="3.02" valueMin="2.84">2.843.02 mm</quantity>
; width of prozona about
<quantity id="4F963E794C8C9F2BF975931A931510A2" metricMagnitude="-3" metricUnit="m" metricValue="1.32" unit="mm" value="1.32">1.32 mm</quantity>
; length of metazona about
<quantity id="7AB7CB4875B0B7AF378254BF34951016" metricMagnitude="-4" metricUnit="m" metricValue="2.6" unit="mm" value="0.26">0.26 mm</quantity>
, average ratio of width of head and width of collum = 1.14. Nonsexual characters as in male.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEOAG" type="etymology">
<paragraph id="0BACB939FDA33EFC24DEEFDEE0AE573C">
<heading id="D6C357F4C7EF4BCEF54A6A883DEE11BF" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="284C3B22CA4B7111929EAB44FD5E6B59">
Named after the “ Ngoc Linh ” Mountain, the
<typeStatus id="0B0EC2D6ED4E431B2161DD5DE2933E65">type</typeStatus>
locality. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJOAG" type="distribution">
<paragraph id="EE1C26792FCDD5069423B83BBADCCED3">
<heading id="A4AA815E391591755FD9685E50BDE81B" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="BC430CCA1699E67E25FCF84298F84E8F">
The species has been found in Ngoc Linh Mts at the high elevation of
<quantity id="697E25E74C850A55C1B64B1814004490" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">18001900 m</quantity>
a. s. l. in
<collectingRegion id="DA15034693CD4CAB93FDD291C18D1ACA" country="Vietnam" name="Kon Tum">Kon Tum Province</collectingRegion>
,
<collectingCountry id="FE904E3CC8E8A013636EFF34D7393B0D" name="Vietnam">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOOAG" type="DNA barcoding">
<paragraph id="F6EE9495E5D50A8FF8BFD552C5468038">
<heading id="F6F21EE39CA9E36ADE58F2FF430ED910" reason="title">DNA barcoding.</heading>
</paragraph>
<paragraph id="E74668D952FFD0F17A65B9BA814107CA">
A fragment of the
<abbrev id="ABBRID0EUOAG" xlink_title="cytochrome c oxidase subunit I">COI</abbrev>
gene is accessioned at
<collectionCode id="0B3E174C905E4F84F59604779D3AC63F">NCBI</collectionCode>
GenBank with the following accession numbers:
<ext-link id="90488A0E718EE09E901A40512694FE6C" ext-link-type="gen" xlink_href="PQ423222" xlink_type="simple">PQ 423222</ext-link>
<ext-link id="F75D443733220939D01D2BF47CCAE9E8" ext-link-type="gen" xlink_href="PQ423223" xlink_type="simple">PQ 423223</ext-link>
. The new species has a
<abbrev id="ABBRID0ECPAG" xlink_title="cytochrome c oxidase subunit I">COI</abbrev>
gene similarity with
<taxonomicName id="1636951708DC17EC1E7C9A2EE2F05015" authorityName="Cope" authorityYear="1869" class="Diplopoda" family="Andrognathidae" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="family">Andrognathidae</taxonomicName>
spp. (
<ext-link id="6A01422886B76612A2FCF9FA03713541" ext-link-type="gen" xlink_href="MF983566" xlink_type="simple">
<accessionNumber id="3F9753F0261B582180D6335DDA616877" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF983566">
<collectionCode id="151E00EFDBBBAC0A615980E09C3601B6">MF</collectionCode>
983566
</accessionNumber>
</ext-link>
and
<ext-link id="32CE362F97A7942D7CA1A4F399D090FB" ext-link-type="gen" xlink_href="MF983567" xlink_type="simple">
<accessionNumber id="CBEA8DD88DB4F3D781E2D63659544EC2" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF983567">MF 983567</accessionNumber>
</ext-link>
) of 89.13 % and 89.37 % identity, respectively.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,609 @@
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<mods:title id="76962352DF0F3EE2454C5593A766848B">Acotylea (Platyhelminthes, Polycladida) from the southern and western Iberian Peninsula, with the description of five new species</mods:title>
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<mods:namePart id="F90B8B290DAFED9C434FEE1F46AD79E9">Gouveia, Filipa</mods:namePart>
<mods:affiliation id="92038D266969DB2A4AA374B34E5B5F55">EPCV-Departamento de Ciências da Vida, Universidade Lusófona, Campo Grande, 376, 1749 - 024 Lisboa, Portugal</mods:affiliation>
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<taxonomicName id="749128A76022A1E2D90C0AA412FDBB89" authority="(Bock, 1913)" baseAuthorityName="Bock" baseAuthorityYear="1913" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="0E559FF3767580D231A60CD426187283" italics="true">Leptoplana mediterranea</emphasis>
(
<bibRefCitation id="15DA39351A78FBFA5CD9F6995BF3D24C" author="Bock S" journalOrPublisher="Zoologiska Bidrag fran Uppsala" pagination="31-344" refId="B6" refString="Bock S (1913) Studien uber Polycladen. Zoologiska Bidrag fran Uppsala 2: 31 - 344." title="Studien über Polycladen." volume="2" year="1913">Bock, 1913</bibRefCitation>
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<emphasis id="0CA6D9605DAB5345903E0673D9B064CD" italics="true">Leptoplana tremellaris mediterranea</emphasis>
<bibRefCitation id="942BBFD535DD4F67BEB11702070A90A2" author="Bock S" journalOrPublisher="Zoologiska Bidrag fran Uppsala" pagination="31-344" refId="B6" refString="Bock S (1913) Studien uber Polycladen. Zoologiska Bidrag fran Uppsala 2: 31 - 344." title="Studien über Polycladen." volume="2" year="1913">Bock, 1913</bibRefCitation>
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<heading id="7228F5C4FCE969980836F91B7B8D1FA3" reason="title">Material examined.</heading>
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long, sagittally sectioned into 3 slides
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long, sagittally sectioned into 4 slides
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long, sagittally sectioned into 18 slides
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<materialsCitation id="BB42E6ED64681F677CA3A98F318B614F" collectingDate="2016-09-08" collectionCode="MNHNC" specimenCode="MB 16-000112" specimenCount="1">
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long
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long
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<materialsCitation id="5D673C8D788BCA47BC5C601B461E4495" collectingDate="2018-10-28" collectionCode="MNHNC" specimenCode="MB 16-000122" specimenCount="1">
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long, sagittally sectioned into 13 slides
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<materialsCitation id="22C8D5D182DDC053072D03C1BA0F728E" collectingDate="2018-10-28" collectionCode="MNHNC" specimenCode="MB 16-000121" specimenCount="1">
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long, sagittally sectioned into 28 slides
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<materialsCitation id="7A6691A66C2FC8D36FB37188354B57A9" collectingDate="2018-11-10" collectionCode="MNHNC" specimenCode="MB 16-000123" specimenCount="1">
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long, sagittally sectioned into 50 slides
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<materialsCitation id="6EC8305DF87CBCB004B6490451C8978B" collectingDate="2019-02-23" collectionCode="MNHNC" specimenCode="MB 16-000116" specimenCount="1">
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<specimenCode id="716BC037AD3FF29D9744E64D6AC8C702" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB 16-000116</specimenCode>
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<collectingDate id="C29C272ABCA00064B004CFF66CA560EE" value="2019-02-23">23 February 2019</collectingDate>
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long, sagittally sectioned into 20 slides
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<materialsCitation id="F38B0716E60551671286077E727FEA1A" collectingDate="2019-02-23" collectionCode="MNHNC" specimenCode="MB 16-000117" specimenCount="1">
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<specimenCode id="ED68E0702BE76C842DFE239CF988F48D" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB 16-000117</specimenCode>
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<date id="65838934DBB5C0FAAF6A245ABCF10B53" value="2019-02-23">
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long, sagittally sectioned into 21 slides
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<materialsCitation id="A1FA018F63E5DC0264BD801E3A97E35F" collectingDate="2019-02-23" collectionCode="MNHNC" specimenCode="MB 16-000118" specimenCount="1">
<abbrev id="ABBRID0EWIBI" xlink_title="Museu Nacional de História Natural e da Ciência of Lisbon">
<collectionCode id="762A2C04A9DDEA396CF33C1B47F79268">MNHNC</collectionCode>
</abbrev>
<specimenCode id="3DB292909B8D677FF196F712714E49C7" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB 16-000118</specimenCode>
, Station 4,
<date id="DC33CDC262C7B204F342AD08042A4CB2" value="2019-02-23">
<collectingDate id="DA64A54FD83781C6691FE6D0191924A1" value="2019-02-23">23 February 2019</collectingDate>
</date>
,
<quantity id="CB01DFB4EE3703A94A1089E2251B3918" metricMagnitude="-2" metricUnit="m" metricValue="1.9" unit="mm" value="19.0">19 mm</quantity>
long, sagittally sectioned into 27 slides
</materialsCitation>
; •
<materialsCitation id="84CBD23DBC32B6FD47984D35EA3C2836" collectingDate="2019-02-23" collectionCode="MNHNC" specimenCode="MB 16-000119" specimenCount="1">
<abbrev id="ABBRID0E1IBI" xlink_title="Museu Nacional de História Natural e da Ciência of Lisbon">
<collectionCode id="CF1DBCB96F992D3D7921F693C118B4B9">MNHNC</collectionCode>
</abbrev>
<specimenCode id="180D3F80312E6B6C2DA8D58F64440BFE" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB 16-000119</specimenCode>
, Station 4,
<date id="62D4AB25FD8A07E600CDD49F110CDBF5" value="2019-02-23">
<collectingDate id="656A7D3CF3A5F3AB6E837F0DB2AD7BA2" value="2019-02-23">23 February 2019</collectingDate>
</date>
,
<quantity id="29BDCF07C11335350797A81B464E192D" metricMagnitude="-2" metricUnit="m" metricValue="1.3" unit="mm" value="13.0">13 mm</quantity>
long, sagittally sectioned into 17 slides
</materialsCitation>
; •
<materialsCitation id="9593E6A5541A9E90BD74583D994808CE" collectingDate="2019-02-23" collectionCode="MNHNC" specimenCode="MB 16-000120" specimenCount="1">
<abbrev id="ABBRID0E5IBI" xlink_title="Museu Nacional de História Natural e da Ciência of Lisbon">
<collectionCode id="9DDA8A8E8C94BB428FA9BD480F90A228">MNHNC</collectionCode>
</abbrev>
<specimenCode id="54EC75B2B0045BC58AEC4F2502316397" collectionCode="MB" country="Portugal" name="Universidade de Lisboa, Museu Bocage">MB 16-000120</specimenCode>
, Station 4,
<date id="43441175F014EF01D97962299C1A5C77" value="2019-02-23">
<collectingDate id="B59A8198BA3B40EE2C505F9A77560C16" value="2019-02-23">23 February 2019</collectingDate>
</date>
,
<quantity id="5871CD461043A91FFCC30EC3CD564F93" metricMagnitude="-2" metricUnit="m" metricValue="1.8" unit="mm" value="18.0">18 mm</quantity>
long, sagittally sectioned into 16 slides
</materialsCitation>
; •
<materialsCitation id="8A72CDFAE6AF2D4D4204826774A2BC92" collectingDate="2015-05-10" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="6915A61C689A2D56D61EBFC43A1FAB7F" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="C2CE34206E8C1C1FB03D20AA4A5963E6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3968 to 3981, Station 11,
<date id="CE3981AE38052650CA7CA9F7B3342FF4" value="2015-05-10">
<collectingDate id="CD189F57B66BD938DA9BF02F8FE3A6E1" value="2015-05-10">10 May 2015</collectingDate>
</date>
,
<quantity id="CBA857B233AB96D2FF5C49E027A76CEC" metricMagnitude="-2" metricUnit="m" metricValue="1.4" unit="mm" value="14.0">14 mm</quantity>
long, sagittally sectioned into 13 slides. All of the measurements from the description refer to this specimen
</materialsCitation>
; •
<materialsCitation id="4C003E7D45CD62F42EAD7E4602027D0D" collectingDate="2015-05-10" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="7935C06A01223CDCE21E16481FBD913F" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="05DEAD0551A35E73DCC3D0AD155A3E1D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3382 to 3393, Station 11,
<date id="2FE10E0371AEF20F801C9075F68C4FCD" value="2015-05-10">
<collectingDate id="CA1B4C99B73C0D5BD445C5736F977AA3" value="2015-05-10">10 May 2015</collectingDate>
</date>
,
<quantity id="16B892B6C164DEF6DAD6EC29E1D05AC6" metricMagnitude="-2" metricUnit="m" metricValue="1.1" unit="mm" value="11.0">11 mm</quantity>
long, sagittally sectioned into 12 slides
</materialsCitation>
; •
<materialsCitation id="E7AB0DE45B4887D7D896DE06B877226B" collectingDate="2016-04-11" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="2BF952DC793ADE839C370AB4F5FECB2B" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="5366BBE82BF7411595AAABE9C21EB5BF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3982, Station 11,
<date id="83454D86B99E034D49144F988156DBF0" value="2016-04-11">
<collectingDate id="77454811AFBC748D0C62882D261A4D1A" value="2016-04-11">11 April 2016</collectingDate>
</date>
,
<quantity id="E316F23E0E8B9040CAE6A4DB4D8DA6A2" metricMagnitude="-2" metricUnit="m" metricValue="1.1" unit="mm" value="11.0">11 mm</quantity>
long
</materialsCitation>
; •
<materialsCitation id="ECE35EAC192C9A15C66CB5D5E7793BFA" collectingDate="2013-12-06" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="32C265007B16943933D25B2CA1FF3057" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="7CDA9DE5A79B5D83932F6934889CACB8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3409 to 3416, Station 12,
<date id="DF1B38C03D969071D7071E55E05968EA" value="2013-12-06">
<collectingDate id="98274BD36DDC0B3EE831BFBFCA2B1D32" value="2013-12-06">6 December 2013</collectingDate>
</date>
,
<quantity id="53395105BDB3D4B55BA676345640C6E4" metricMagnitude="-2" metricUnit="m" metricValue="1.3" unit="mm" value="13.0">13 mm</quantity>
long, sagittally sectioned into 8 slides
</materialsCitation>
; •
<materialsCitation id="4E2D13B4A3F254D18122BA5ADE3035F4" collectingDate="2013-12-06" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="EEEB4CF4504EC9053FE0AB1C39E92CB4" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="C3F1C7CB7B5C7DDDEB50D8E7DECC0ABA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3417 to 3428, Station 12,
<date id="FF9F97F4B82FD97E33403C1946AAE3A8" value="2013-12-06">
<collectingDate id="C8930BC3AF9EE68CC51D98F55C085DE5" value="2013-12-06">6 December 2013</collectingDate>
</date>
,
<quantity id="31A61D9DC048994F8ECD979B06C915DF" metricMagnitude="-2" metricUnit="m" metricValue="1.5" unit="mm" value="15.0">15 mm</quantity>
long, sagittally sectioned into 12 slides
</materialsCitation>
; •
<materialsCitation id="C8ACDB497F720400FAC2A92ED1A32B18" collectingDate="2013-12-06" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="17CA541B116E0AE4D3288778BB3127C9" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="CB8C3C273169D5F02198EBFE9341D23E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3429 to 3447, Station 12,
<date id="DD7F8D5B14E34746D734B15F967A021D" value="2013-12-06">
<collectingDate id="7534ABD7863F960ED48F7EBD1D98E3FB" value="2013-12-06">6 December 2013</collectingDate>
</date>
,
<quantity id="1EB2221C88AFCECA504067CD0AC167D5" metricMagnitude="-2" metricUnit="m" metricValue="1.7" unit="mm" value="17.0">17 mm</quantity>
long, sagittally sectioned into 19 slides
</materialsCitation>
; •
<materialsCitation id="CE2F76A5F9B8180749D346437FAED3DD" collectingDate="2013-12-06" collectionCode="MNCN" specimenCode="MNCN 4.01" specimenCount="1">
<specimenCode id="996144691A0F144FE690E30EB15F21E1" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">
<named-content id="08394A80047ED66407740778FB305F39" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01
</specimenCode>
/ 3394 to 3408, Station 12,
<date id="2908345D6D717B22E1CFFCB0042C8DB0" value="2013-12-06">
<collectingDate id="E64C76ED9D8C8D179DA73E835FD93E0E" value="2013-12-06">6 December 2013</collectingDate>
</date>
,
<quantity id="9A5F9EA76184DFA65C10E204724C4E92" metricMagnitude="-2" metricUnit="m" metricValue="1.6" unit="mm" value="16.0">16 mm</quantity>
long, sagittally sectioned into 15 slides
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5JBI" type="description">
<paragraph id="1F0890978D69F896EAC9088D8661318B">
<heading id="677F9D7F4482175C95073FFB3D3CF58D" reason="title">Description.</heading>
</paragraph>
<paragraph id="0C6A93885D7A5D0482D30747DE0EA961">
Elongated worm, wider in the anterior end and narrower in the posterior end. Length between
<quantity id="4577F89E44B93D25A7B127C935BA0397" metricMagnitude="-2" metricUnit="m" metricValue="1.1" unit="mm" value="11.0">11 mm</quantity>
and
<quantity id="3F7BABAED7B0D102A037E42680328271" metricMagnitude="-2" metricUnit="m" metricValue="3.1" unit="mm" value="31.0">31 mm</quantity>
(18.42 ±
<quantity id="536AAF40DBE822039E1EA01A4F87109A" metricMagnitude="-3" metricUnit="m" metricValue="4.58" unit="mm" value="4.58">4.58 mm</quantity>
). Ground colour of the dorsal surface, beige brownish in the pharynx area (Fig.
<figureCitation id="B982688D14EB263A07CF19D7C9802A84" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Leptoplana mediterranea (MNCN 4.01 / 3968 to 3981) Histological sections are antero-posteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the eyes; C. Example of the environment where the specimen was found; D. Histological section of the male copulatory apparatus; E. Histological section of the female copulatory apparatus; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; d, diverticula; ev, external vagina; fp, female pore; gs, genital sucker; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; p, penis; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure8" httpUri="https://binary.pensoft.net/fig/1169821">8 A</figureCitation>
). Tentacles absent. A whitish colour is observed in the brain area, allowing clear observation of approx. 35 cerebral and 26 tentacular eye clusters (Fig.
<figureCitation id="51F300BC7E89131F96C05F871BDF5250" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Leptoplana mediterranea (MNCN 4.01 / 3968 to 3981) Histological sections are antero-posteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the eyes; C. Example of the environment where the specimen was found; D. Histological section of the male copulatory apparatus; E. Histological section of the female copulatory apparatus; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; d, diverticula; ev, external vagina; fp, female pore; gs, genital sucker; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; p, penis; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure8" httpUri="https://binary.pensoft.net/fig/1169821">8 B</figureCitation>
). Ruffled pharynx between the first and second thirds of the body, mouth opening in the last third. Genital pores separated (approximately
<quantity id="447C090BAC68A684276D5979CBDF1A21" metricMagnitude="-4" metricUnit="m" metricValue="3.3" unit="mm" value="0.33">0.33 mm</quantity>
) and located just behind the pharynx.
</paragraph>
<caption id="A34AD2D1256EE4BFD4B25D2CE7920F49" ID-DOI="10.3897/zse.100.128211.figure8" ID-arpha="DE4DD35D-B412-5DA8-923B-25FE917831F3" httpUri="https://binary.pensoft.net/fig/1169821" startId="F8">
<paragraph id="FE98ABD72B921EFC2D35BE334C76C866">
<label id="728FF1140BDCAB297DF4933C266DCF55">Figure 8.</label>
</paragraph>
<paragraph id="4E1DA98A3A1DDF6DFB3AC1F806D36F07">
<taxonomicName id="F82A4DF57AA605F26C2C3F49E86CD855" baseAuthorityName="Bock" baseAuthorityYear="1913" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="BDC0F391E40ACB9736BFE304A47EDD7D" italics="true">Leptoplana mediterranea</emphasis>
</taxonomicName>
(
<named-content id="61358764BE8F18E9EEAC0FDA5F1483E7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales de Madrid">MNCN</named-content>
4.01 / 3968 to 3981) Histological sections are antero-posteriorly oriented:
<emphasis id="7F2875EC6EE25C9FD3D89D5A14F30723" bold="true">A.</emphasis>
Dorsal view of the living specimen;
<emphasis id="4F0D2C22B7F91B096BB3446E7DC0EB7E" bold="true">B.</emphasis>
Detail of the eyes;
<emphasis id="ECF266FF495193D15788D94D329C8C94" bold="true">C.</emphasis>
Example of the environment where the specimen was found;
<emphasis id="77B72ACF08F483417F9F89EFE20892E4" bold="true">D.</emphasis>
Histological section of the male copulatory apparatus;
<emphasis id="350CA39935B87034B3B1D91D26CA2BEB" bold="true">E.</emphasis>
Histological section of the female copulatory apparatus;
<emphasis id="BEC4C9B026438F8C2213896B32F38933" bold="true">F.</emphasis>
Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; d, diverticula; ev, external vagina; fp, female pore; gs, genital sucker; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; p, penis; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EULBI" type="Reproductive system">
<paragraph id="93482AC0EAB647D14B2BDCB90638EB04">
<heading id="11026AD3BDB66009141FB9A64F862278" reason="title">Reproductive system.</heading>
</paragraph>
<paragraph id="2065F020292A9096B6DABD3603A8C441">
Male reproductive system with a true and large seminal vesicle (
<quantity id="43A1080BC8C34E5862158EA7C46AC58F" metricMagnitude="-4" metricUnit="m" metricValue="2.8" unit="mm" value="0.28">0.28 mm</quantity>
long ×
<quantity id="DF9CED40C14B9F395C5AD060A617DB5C" metricMagnitude="-4" metricUnit="m" metricValue="1.4" unit="mm" value="0.14">0.14 mm</quantity>
wide), interpolated prostatic vesicle, and an unarmed penis cirrus, all enclosed in a muscular bulb (Fig.
<figureCitation id="F5C08CDB59B34ACAC97930E0A6F19090" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Leptoplana mediterranea (MNCN 4.01 / 3968 to 3981) Histological sections are antero-posteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the eyes; C. Example of the environment where the specimen was found; D. Histological section of the male copulatory apparatus; E. Histological section of the female copulatory apparatus; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; d, diverticula; ev, external vagina; fp, female pore; gs, genital sucker; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; p, penis; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure8" httpUri="https://binary.pensoft.net/fig/1169821">8 F</figureCitation>
). Vas deferens ventrally positioned, joining together before entering into the seminal vesicle. Seminal vesicle oval-shaped (Fig.
<figureCitation id="418723A41A5568BA54A0EC01090B246F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Leptoplana mediterranea (MNCN 4.01 / 3968 to 3981) Histological sections are antero-posteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the eyes; C. Example of the environment where the specimen was found; D. Histological section of the male copulatory apparatus; E. Histological section of the female copulatory apparatus; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; d, diverticula; ev, external vagina; fp, female pore; gs, genital sucker; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; p, penis; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure8" httpUri="https://binary.pensoft.net/fig/1169821">8 D</figureCitation>
). Prostatic vesicle ventrally fastened to the seminal vesicle and separated by an epithelial common wall. Genital sucker located between the male and female pores.
</paragraph>
<paragraph id="0DFBCEFBCBBF08B165FED2BA2A7D4596">
Female complex simple and composed of a wide external vagina and a short and curved internal vagina. Langs vesicle rudimentary (Fig.
<figureCitation id="4E01BCCA2035AC2A1AFAF4D15267257A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Leptoplana mediterranea (MNCN 4.01 / 3968 to 3981) Histological sections are antero-posteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the eyes; C. Example of the environment where the specimen was found; D. Histological section of the male copulatory apparatus; E. Histological section of the female copulatory apparatus; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; d, diverticula; ev, external vagina; fp, female pore; gs, genital sucker; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; p, penis; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure8" httpUri="https://binary.pensoft.net/fig/1169821">8 E, F</figureCitation>
). Numerous shell glands surround the whole female system.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIMBI" type="remarks">
<paragraph id="C5A071061E964024987D459450F4D1FC">
<heading id="BAC1F6CFD367BDE692A074D134808237" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="73DFD49B5CE4010AD176C4FDE73D6941">
These diagnostic characters are present in the specimens collected during the present study, so this is the first report of the presence of
<taxonomicName id="D8B5CC2BED793B689457B9C4AE58CDC2" baseAuthorityName="Bock" baseAuthorityYear="1913" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="29BE9BE9A0ACD2126102DAB0E5BDFB95" italics="true">L. mediterranea</emphasis>
</taxonomicName>
outside the Mediterranean.
</paragraph>
<paragraph id="96A8030F591C32D22286E29FD0A0D1D1">
Likewise, it is possible that the specimens found by
<bibRefCitation id="AE5A91BF3D05E9DAB934C3CCD885396F" author="Saldanha L" firstAuthor="Saldanha" refId="B75" refString="Saldanha L (1974) Estudo do povoamento dos horizontes superiors da rocha litoral da costa da Arrabida (Portugal). Museu e Laboratório Zoológico e Antropológico. Faculdade de Ciências de Lisboa. Arquivos do Museu Bocage V (I), 382 pp." year="1974">Saldanha (1974)</bibRefCitation>
reported as
<taxonomicName id="325004D31F212FA2A4718AA1B9816611" baseAuthorityName="Muller OF" baseAuthorityYear="1773" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="tremellaris">
<emphasis id="931979AB7A7B00F6B33041EC3CC90403" italics="true">L. tremellaris</emphasis>
</taxonomicName>
belong to
<taxonomicName id="AC696E9CF878B94FB97A167CBFBB354F" baseAuthorityName="Bock" baseAuthorityYear="1913" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="6D509A5055D6119CEB4A6DCE5848A52F" italics="true">L. mediterranea</emphasis>
</taxonomicName>
as well since some specimens of
<taxonomicName id="34C3363C2F010A33D89C032BB8AC1962" baseAuthorityName="Bock" baseAuthorityYear="1913" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="249C033BE723F9864815DC87E6B7694C" italics="true">L. mediterranea</emphasis>
</taxonomicName>
were found close to Arrabida. However, this hypothesis needs to be confirmed with histological sections.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAOBI" type="biology">
<paragraph id="0E745E70CBC24F2E9608131E3ACE72EF">
<heading id="2F0CB2259513456381158ACDFFC03107" reason="title">Biology.</heading>
</paragraph>
<paragraph id="94D7615BA51F9A88F9D336400084A1C7">
Under rocks (Fig.
<figureCitation id="7B1ECD16613F76E0F6CD50CD6ECD0AE6" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Leptoplana mediterranea (MNCN 4.01 / 3968 to 3981) Histological sections are antero-posteriorly oriented: A. Dorsal view of the living specimen; B. Detail of the eyes; C. Example of the environment where the specimen was found; D. Histological section of the male copulatory apparatus; E. Histological section of the female copulatory apparatus; F. Sagittal reconstruction of the reproductive system. Abbreviations: cg, cemental glands; d, diverticula; ev, external vagina; fp, female pore; gs, genital sucker; i, intestine; iv, internal vagina; ma, male atrium; mp, male pore; ov, oviduct; p, penis; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; vd, vas deferens." figureDoi="10.3897/zse.100.128211.figure8" httpUri="https://binary.pensoft.net/fig/1169821">8 C</figureCitation>
, present study), but it was also found with
<taxonomicName id="89BC1FD826ABF719EE994A70FB639894" authorityName="J. V. Lamouroux" authorityYear="1809" baseAuthorityName="Forsskal" class="Ulvophyceae" family="Caulerpaceae" genus="Caulerpa" kingdom="Plantae" order="Bryopsidales" phylum="Chlorophyta" rank="species" species="prolifera">
<emphasis id="1938241944FBE8E387D252A2B76E1A3D" italics="true">Caulerpa prolifera</emphasis>
</taxonomicName>
(Chlorophyta) (
<bibRefCitation id="785503A9241B6A1C67ABA6C6612640E8" DOI="10.11646/zootaxa.3780.1.4" author="Marquina D &amp; Osca D &amp; Rodríguez J &amp; Fernández-Despiau E &amp; Noreña C" issue="1" journalOrPublisher="Zootaxa" pagination="108-134" refId="B45" refString="Marquina D, Osca D, Rodríguez J, Fernández-Despiau E, Noreña C (2014 a) State of knowledge of the Acotylea (Polycladida, Platyhelminthes) from the Mediterranean coasts of Spain: New records and new species. Zootaxa 3780 (1): 108134. https://doi.org/10.11646/zootaxa.3780.1.4" title="State of knowledge of the Acotylea (Polycladida, Platyhelminthes) from the Mediterranean coasts of Spain: New records and new species." volume="3780">Marquina et al. 2014 a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5OBI" type="distribution">
<paragraph id="0B792066C838BEFED9F2230778E66A0F">
<heading id="BFFE68171281B5A3F806189C9E15311B" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="934210DC0F594F5AACFDEF0B86D388BA">
Widely distributed along the Mediterranean coasts,
<taxonomicName id="77DB1044BF26E0CEE32E5FF52C92B647" baseAuthorityName="Bock" baseAuthorityYear="1913" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="1D7841D7C54905822D31D4B2211702D0" italics="true">L. mediterranea</emphasis>
</taxonomicName>
was reported as
<taxonomicName id="A084553CF16C149027A1924D0704BF5B" baseAuthorityName="Muller OF" baseAuthorityYear="1773" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="tremellaris">
<emphasis id="633A5B5542DB584C643B7354F9274DA2" italics="true">L. tremellaris</emphasis>
</taxonomicName>
“ forma
<taxonomicName id="D44CB7205A13A432826C97C7FC77B145" baseAuthorityName="Bock" baseAuthorityYear="1913" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B5D077382D997AD54B553CBD4A915875" italics="true">mediterranea</emphasis>
</taxonomicName>
” in Palermo (
<bibRefCitation id="191AD445EA3AC3B96A17371A9983637F" DOI="10.5962/bhl.title.10133" author="Grube" firstAuthor="Grube" refId="B27" refString="Grube AE (1840) Actinien, Echinodermen und Würmer des adriatischen und Mittelmeers, nach eigenen Sammlungenbeschrieben. Königsberg, 92 pp. https://doi.org/10.5962/bhl.title.10133" year="1840">Grube 1840</bibRefCitation>
), Naples (
<bibRefCitation id="F91060A7FEEF896C4C1E09CAC1B976D8" DOI="10.5962/bhl.title.10545" author="Lang" firstAuthor="Lang" refId="B43" refString="Lang A (1884) Die Polycladen (Seeplanarien) des Golfes von Neapel und der angrenzenden Meeresabschnitte. Eine Monographie. Fauna und Flora des Golfes von Neapel 11. W Engelmann, Leipzig. https://doi.org/10.5962/bhl.title.10545" year="1884">Lang 1884</bibRefCitation>
;
<bibRefCitation id="E08B3A743FECF2724F888DEC88FFFC21" author="Palombi" firstAuthor="Palombi" journalOrPublisher="Archivio Zoologico Italiano" pagination="1-45" refId="B60" refString="Palombi A (1936) Policladi liberi e commensali raccolti sulle coste del Sud Africa, della Florida e del Golfe di Napoli. Archivio Zoologico Italiano 23: 145." title="Policladi liberi e commensali raccolti sulle coste del Sud Africa, della Florida e del Golfe di Napoli." volume="23" year="1936">Palombi 1936</bibRefCitation>
), the Gulf of Lyon (
<bibRefCitation id="38D78F5C3062807AA8D3956364555AD4" author="Pruvot" firstAuthor="Pruvot" refId="B68" refString="Pruvot G (1897) Essai sur les Fonds et la Faune de la Manche occidentale (Côtes de Bretagne) comparés à ceux du Golfe du Lion. Archive de Zoologie Expérimentale et Générale 3 (5): 1920, 617660." year="1897">Pruvot 1897</bibRefCitation>
), Trieste (
<bibRefCitation id="6597638AF9D1F6F2A0D1109EACD58E01" author="Micoletzky" firstAuthor="Micoletzky" journalOrPublisher="Arbeiten aus dem Zoologischen Institut der Universität Wien" pagination="167-182" refId="B48" refString="Micoletzky H (1910) Die Turbellarienfauna des Golfes von Triest. Arbeiten aus dem Zoologischen Institut der Universität Wien 18: 167182." title="Die Turbellarienfauna des Golfes von Triest." volume="18" year="1910">Micoletzky 1910</bibRefCitation>
), Port Said (
<bibRefCitation id="37B4B5BFDCD2A432DE5017F24F79EECB" DOI="10.1111/j.1096-3642.1928.tb00208.x" author="Palombi" firstAuthor="Palombi" issue="5" journalOrPublisher="Transactions of the Zoological Society of London" pagination="579-631" refId="B59" refString="Palombi A (1928) Report on the Turbellaria (Cambridge Expedition Suez Canal 1924). Transactions of the Zoological Society of London 22 (5): 579631. https://doi.org/10.1111/j.1096-3642.1928.tb00208.x" title="Report on the Turbellaria (Cambridge Expedition Suez Canal 1924)." volume="22" year="1928">Palombi 1928</bibRefCitation>
), the Adriatic Sea (
<bibRefCitation id="AE6C7D9996366CBFB2A17C3FB95E6C0A" author="Steinbock" firstAuthor="Steinbock" journalOrPublisher="Thalassia" pagination="1-33" refId="B78" refString="Steinböck O (1933) Die Turbellarienfauna der Umgebung von Rovigno. Thalassia 1: 133." title="Die Turbellarienfauna der Umgebung von Rovigno." volume="1" year="1933">Steinböck 1933</bibRefCitation>
), the Italian coasts (
<bibRefCitation id="94DDED8FBA102AB68D705C7E1F819F7B" author="Galleni and Gremigni" editor="Adiyodi KG &amp; Adiyodi RG" firstAuthor="Galleni" journalOrPublisher="IV, Part A. Fertilization, development and parental care. John Wiley and Sons, New York" pagination="63-89" refId="B19" refString="Galleni L, Gremigni V (1989) Platyhelminthes - Turbellaria. In: Adiyodi KG, Adiyodi RG (Eds) Reproductive biology of invertebrates, Vol. IV, Part A. Fertilization, development and parental care. John Wiley and Sons, New York, 6389." title="Platyhelminthes - Turbellaria." volumeTitle="Reproductive biology of invertebrates, Vol." year="1989">Galleni and Gremigni 1989</bibRefCitation>
), and Catalonia (
<bibRefCitation id="64A128E5F2003CEDDDD30943001EBF3E" author="Novell" firstAuthor="Novell" refId="B55" refString="Novell C (2001) Contribution to the knowledge of polyclad Turbellaria of the Catalan coast. PhD Thesis, University of Barcelona, Spain. http://www.tesisenxarxa.net/TESIS_UB/AVAILABLE/TDX-0107104-121621//Tesis.PDF" url="http://www.tesisenxarxa.net/TESIS_UB/AVAILABLE/TDX-0107104-121621//Tesis.PDF" year="2001">Novell 2001</bibRefCitation>
.
<collectingCountry id="08643AFAE36C1914815468D2D7964CAF" name="Tunisia">Tunisia</collectingCountry>
(
<bibRefCitation id="6FA89343ADD569012578F4145315952F" DOI="10.11646/zootaxa.3178.1.4" author="Gammoudi" etAl="et al." firstAuthor="Gammoudi" journalOrPublisher="Zootaxa" pagination="45-56" refId="B23" refString="Gammoudi M, Egger B, Tekaya S, Noreña C (2012) The genus Leptoplana (Leptoplanidae, Polycladida) in the Mediterranean basin. Redescription of the species Leptoplana mediterranea (Bock 1913) comb. nov. Zootaxa 3178: 4556. https://doi.org/10.11646/zootaxa.3178.1.4" title="The genus Leptoplana (Leptoplanidae, Polycladida) in the Mediterranean basin. Redescription of the species Leptoplana mediterranea (Bock 1913) comb. nov." volume="3178" year="2012">Gammoudi et al. 2012</bibRefCitation>
,
<bibRefCitation id="18F5619D975CD4142C23AE88E6DB24F1" DOI="10.11646/zootaxa.4263.1.5" author="Gammoudi" firstAuthor="Gammoudi" issue="1" journalOrPublisher="Zootaxa" pagination="120-138" refId="B24" refString="Gammoudi M, Garbouj M, Egger B, Tekaya S (2017) Updated inventory and distribution of free-living flatworms from Tunisian waters. Zootaxa 4263 (1): 120138. https://doi.org/10.11646/zootaxa.4263.1.5" title="Updated inventory and distribution of free-living flatworms from Tunisian waters." volume="4263" year="2017">2017</bibRefCitation>
); Mar Menor, SE Spain (
<bibRefCitation id="99C99C41A57E209BE1A742D5F1F47E14" DOI="10.11646/zootaxa.3780.1.4" author="Marquina D &amp; Osca D &amp; Rodríguez J &amp; Fernández-Despiau E &amp; Noreña C" issue="1" journalOrPublisher="Zootaxa" pagination="108-134" refId="B45" refString="Marquina D, Osca D, Rodríguez J, Fernández-Despiau E, Noreña C (2014 a) State of knowledge of the Acotylea (Polycladida, Platyhelminthes) from the Mediterranean coasts of Spain: New records and new species. Zootaxa 3780 (1): 108134. https://doi.org/10.11646/zootaxa.3780.1.4" title="State of knowledge of the Acotylea (Polycladida, Platyhelminthes) from the Mediterranean coasts of Spain: New records and new species." volume="3780">Marquina et al. 2014 a</bibRefCitation>
); and the southern plus western Iberian Peninsula (present study). This is the first record of
<taxonomicName id="C6B0DAC8C543B4631D032695AA769C8A" baseAuthorityName="Bock" baseAuthorityYear="1913" family="Leptoplanidae" genus="Leptoplana" kingdom="Animalia" order="Polycladida" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="C4D2FD21B58E9805DEBBDB507C6220AB" italics="true">L. mediterranea</emphasis>
</taxonomicName>
in
<collectingRegion id="BCF52926231FF8587E54A57A75BBCF94" country="Portugal" name="Porto">Oporto</collectingRegion>
(
<collectingCountry id="6FDB64ECD7F50DFCB11014567852782D" name="Portugal">Portugal</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,628 @@
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<mods:title id="A46E724619CC8E298EB30E58F0E01A64">A review of the millipede genus Pseudodesmus Pocock, 1887 (Diplopoda, Platydesmida, Andrognathidae) from Vietnam, with descriptions of five new species and notes on its phylogeny</mods:title>
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<mods:affiliation id="421461B8FEFD93E0EED3CE746287719C">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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<emphasis id="152A5B364CC6BFF9FF3901DC861C0F36" italics="true">Pseudodesmus condao</emphasis>
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<taxonomicNameLabel id="948ED73DCF0A88043C91003B2A14825A" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="E9810B043A62BCCD9921911821D4BC0F">
<figureCitation id="59737BAB0D59090ADFCED5033100A719" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure15" httpUri="https://binary.pensoft.net/fig/1169905">Figs 15</figureCitation>
,
<figureCitation id="705F509D8113CB6792C39B3B896DF1D0" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Head and collum, ventral view; E. Anterior part of body, ventral view; F. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure16" httpUri="https://binary.pensoft.net/fig/1169906">16</figureCitation>
,
<figureCitation id="A65CC5B120A2ABD051BADDA9B1CCA6DD" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure17" httpUri="https://binary.pensoft.net/fig/1169907">17</figureCitation>
,
<figureCitation id="13F25BE0CC62EFE4403B4BD7B9A15A66" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Posterior part of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure18" httpUri="https://binary.pensoft.net/fig/1169908">18</figureCitation>
,
<figureCitation id="ECB51F61DC290EF8B6CBDA325A8A860C" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. A. Left anterior gonopods, anterior view; B. Left posterior gonopod, subventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure19" httpUri="https://binary.pensoft.net/fig/1169909">19</figureCitation>
,
<figureCitation id="8849C91A8C0652861BC8F4A045E9F51A" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. SEM. A, B. Left posterior gonopod, subventral view (A), anterior view (B); C. Anterior gonopods, anterior view; D. Left anterior gonopod, anterior view; E. Distal part of left posterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure20" httpUri="https://binary.pensoft.net/fig/1169910">20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMEBG" type="material">
<paragraph id="9F7A9D7510CCEE795E96DA4161267E86">
<heading id="A0B062E1F7E212ECA50E7F358E0F0C46" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="6C593E1497643CCD2BE16C45D4AAA888">
<materialsCitation id="06D3B1FD70FD88C2EB6A0A8443876F84" collectingDate="2019-11-07" collectorName="Con Dao NP &amp; A. D. Nguyen" country="Vietnam" elevation="37" latitude="8.70528" location="Ong Dung Beach" longLatPrecision="1" longitude="106.59158" specimenCount="1" specimenCount-male="1" stateProvince="Ba Ria-Vung Tau" typeStatus="holotype">
<emphasis id="C4B5D167E833167428EB4F978C7EF05D" bold="true" italics="true">
<typeStatus id="3CF7B68C17CFB593DAB222C27E1C4329" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="6964219FCFF03059B6845C8CA8253459" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="640140C254318734893EBDF13EDA9FD1" count="1" type="male">1 male</specimenCount>
;
<collectingRegion id="E1E1C412A69C0F0FABD95626F44153FC" country="Vietnam" name="Ba Ria-Vung Tau">Ba Ria-Vung Tau Province</collectingRegion>
,
<collectorName id="2D9E2B7EFAC6460FA18A71562958397B">
Con Dao
<abbrev content-type="institution" id="ABBRID0EXEBG" xlink_title="National Park">NP</abbrev>
</collectorName>
, on the way to
<location id="67B11901E29E41983B65F83D3DDE35E1" LSID="urn:lsid:plazi:treatment:D3009B8EC9725C80A3A17F06F682CA91:67B11901E29E41983B65F83D3DDE35E1" country="Vietnam" latitude="8.70528" longLatPrecision="1" longitude="106.59158" name="Ong Dung Beach" stateProvince="Ba Ria-Vung Tau">Ong Dung Beach</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6EBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.591580,8.705280]}">
<geoCoordinate id="71C8578EB4672470A3A12A4285445DCF" degrees="08.70528" direction="north" orientation="latitude" precision="1" value="8.70528">08.70528 ° N</geoCoordinate>
,
<geoCoordinate id="12F06CC6A02B6E3AB2B22E7391A84028" degrees="106.59158" direction="east" orientation="longitude" precision="1" value="106.59158">106.59158 ° E</geoCoordinate>
</named-content>
,
<elevation id="A81C8B37FF742D703B39BA5B547FE15C" metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">
<quantity id="A2FFBF375E56991424E7530393D681FF" metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">37 m</quantity>
a. s. l.
</elevation>
,
<date id="958BCBD0A11351FA159E44958F7950DE" value="2019-11-07">
<collectingDate id="52BD457F75DF5D831A7C691F7DD987EE" value="2019-11-07">7 Nov. 2019</collectingDate>
</date>
;
<collectorName id="9E52D1B441507BD38619BD99FB07ACF1">A. D. Nguyen</collectorName>
leg.; natural forest;
<abbrev content-type="institution" id="ABBRID0EEFBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
982 H.
</materialsCitation>
</paragraph>
<paragraph id="E69299EF32BA6CB77920D2528F4F3C6D">
<materialsCitation id="91B7479587E994A4166C5577B42C991F" collectingDate="2019-11-07" collectorName="Con Dao NP &amp; A. D. Nguyen" country="Vietnam" elevation="37" latitude="8.70528" location="Ong Dung Beach" longLatPrecision="1" longitude="106.59158" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Ba Ria-Vung Tau" typeStatus="paratype">
<emphasis id="6818B68C98293F1076C01E900695AA11" bold="true" italics="true">
<typeStatus id="0C88F93A75483BEC06B9549A02313FDE" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="BFE40B2387E20F53A6C6FB2D35E633E6" name="Vietnam">Vietnam</collectingCountry>
<specimenCount id="FC84A59D4B4FC5800E314AC43B0D440F" count="2" type="male">2 males</specimenCount>
,
<specimenCount id="140C4BC6666C4B627965D36DF004DA2F" count="2" type="female">2 females</specimenCount>
; same data as the holotype;
<abbrev content-type="institution" id="ABBRID0EQFBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
982 P
</materialsCitation>
<materialsCitation id="9CE721E41A82E03780A337E3990F4C37" collectingDate="2019-11-07" collectionCode="IEBR" collectorName="Nguyen A. D. &amp; Myr" country="Vietnam" elevation="37" latitude="8.70528" location="Ong Dung" longLatPrecision="1" longitude="106.59158" municipality="Con Dao" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Ba Ria-Vung Tau" typeStatus="paratype">
<specimenCount id="A44732B9AB8867115EA3E49B902304D6" count="3" type="male">3 males</specimenCount>
,
<specimenCount id="73567D1B274EC1E33567FF12BF785D62" count="1" type="female">1 female</specimenCount>
<collectingRegion id="D776B60AE4789CE9C234A99E6010B6B6" country="Vietnam" name="Ba Ria-Vung Tau">Ba Ria-Vung Tau Province</collectingRegion>
,
<collectingMunicipality id="DC7C560AB3E8A64BC4A03F3AC72E4E2B">Con Dao</collectingMunicipality>
<abbrev content-type="institution" id="ABBRID0EVFBG" xlink_title="National Park">NP</abbrev>
, on the way to
<location id="B4E3889644C55D0058ABFBA202678782" LSID="urn:lsid:plazi:treatment:D3009B8EC9725C80A3A17F06F682CA91:B4E3889644C55D0058ABFBA202678782" country="Vietnam" latitude="8.70528" longLatPrecision="1" longitude="106.59158" municipality="Con Dao" name="Ong Dung" stateProvince="Ba Ria-Vung Tau">Ong Dung</location>
beach,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4FBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.591580,8.705280]}">
<geoCoordinate id="C5D5029491A821DADB53B80649F1D732" degrees="08.70528" direction="north" orientation="latitude" precision="1" value="8.70528">08.70528 ° N</geoCoordinate>
,
<geoCoordinate id="C28221F20A05D5DCA188788BC124A168" degrees="106.59158" direction="east" orientation="longitude" precision="1" value="106.59158">106.59158 ° E</geoCoordinate>
</named-content>
,
<elevation id="3BC63568049DEDDF29495F6CFD957C86" metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">
<quantity id="640569460128FB3C0C93600C502749BA" metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">37 m</quantity>
a. s. l.
</elevation>
,
<date id="78354C8AC25460894F408DF41ED1CF0E" value="2019-11-07">
<collectingDate id="A8EFF3F1203B15EC85D7B8B63C0BDC54" value="2019-11-07">7 November 2019</collectingDate>
</date>
, leg.
<collectorName id="60856370A8072037FDF0CCBFC25EFBFD">Nguyen A. D.</collectorName>
; natural forest;
<collectionCode id="60D6793074DC4FDBE906431D14FB26A5">IEBR</collectionCode>
<collectorName id="FEE51AFE42CE712A43DAF0480FB846E1">Myr</collectorName>
821
</materialsCitation>
<materialsCitation id="8FB70F241418F814BA331407D1ED7A1A" collectingDate="2023-11-08" collectorName="A. D. Nguyen" country="Vietnam" elevation="37" latitude="8.70528" location="Ong Dung" longLatPrecision="1" longitude="106.59158" municipality="Con Dao" specimenCount="13" specimenCount-female="5" specimenCount-male="8" stateProvince="Ba Ria-Vung Tau" typeStatus="paratype">
<specimenCount id="DB88D4E09A0F9410DF0F616E5AB87CDD" count="8" type="male">8 males</specimenCount>
,
<specimenCount id="D753B7082DA39A9D4FF88318EE297490" count="5" type="female">5 females</specimenCount>
;
<collectingRegion id="34F7E8EE3EF359F5E954553455F1744D" country="Vietnam" name="Ba Ria-Vung Tau">Ba Ria-Vung Tau Province</collectingRegion>
,
<collectingMunicipality id="A7492C1977A8403B7DA0F497804B9795">Con Dao</collectingMunicipality>
<abbrev content-type="institution" id="ABBRID0ECGBG" xlink_title="National Park">NP</abbrev>
, on the way to
<location id="9097862A9D220B12A78B3BCEC738D600" LSID="urn:lsid:plazi:treatment:D3009B8EC9725C80A3A17F06F682CA91:9097862A9D220B12A78B3BCEC738D600" country="Vietnam" latitude="8.70528" longLatPrecision="1" longitude="106.59158" municipality="Con Dao" name="Ong Dung" stateProvince="Ba Ria-Vung Tau">Ong Dung</location>
beach,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKGBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.591580,8.705280]}">
<geoCoordinate id="CB5D8B655D1720B0E7DBD17806D1D661" degrees="08.70528" direction="north" orientation="latitude" precision="1" value="8.70528">08.70528 ° N</geoCoordinate>
,
<geoCoordinate id="2772E8A28AB824D7A510BA619F84A6DC" degrees="106.59158" direction="east" orientation="longitude" precision="1" value="106.59158">106.59158 ° E</geoCoordinate>
</named-content>
,
<elevation id="9B8D72E3428910174FF3AE9CE545B887" metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">
<quantity id="F20EA7DC0BEA459CEB8E98D18CE77057" metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">37 m</quantity>
a. s. l.
</elevation>
;
<date id="09DCA576C54FBD76A7D4CA92C73BF35D" value="2023-11-08">
<collectingDate id="AC73E27B61C515E03A1B7880DD6A3464" value="2023-11-08">8 Nov. 2023</collectingDate>
</date>
;
<collectorName id="299FDB145193CF89BDA10076C3E5002D">A. D. Nguyen</collectorName>
leg.; natural forest;
<abbrev content-type="institution" id="ABBRID0EPGBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
977
</materialsCitation>
<materialsCitation id="D1818D64AD7B061E9C6F2E5A09758C55" collectingDate="2023-11-08" collectionCode="HNHM" collectorName="A. D. Nguyen" country="Vietnam" elevation="37" latitude="8.70528" location="Ong Dung" longLatPrecision="1" longitude="106.59158" municipality="Con Dao" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Ba Ria-Vung Tau" typeStatus="paratype">
<specimenCount id="14C457F2ACD49A3575335B4C92E66708" count="2" type="male">2 males</specimenCount>
,
<specimenCount id="8F740BF8BAA313D318E5347B06DD308B" count="2" type="female">2 females</specimenCount>
; same data as for sample
<abbrev content-type="institution" id="ABBRID0EUGBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
977;
<named-content id="34F04F15227D5B38B1E7B63A6E526E68" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/hungarian-natural-history-museum-termeszettudomanyi-muzeum" xlink_title="Department of Zoology, Hungarian Natural History Museum, Budapest, Hungary">
<collectionCode id="E16FF3469C2BC975326945E03374D513" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" type="">HNHM</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5GBG" type="diagnosis">
<paragraph id="8C826A8C3D0D8281692B75B6F1F4E022">
<heading id="26E274C7A6BFED88ECF02FB6E62823F8" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="53D8588DD719811767677501CA8A8715">Color dark yellow, little darker in the middle. Head distinctly smaller than collum. Collum bilobed, each with three anterior lobes, no tubercles. Midbody terga with two rows, 34 + 34 large and 4 5 + 4 5 large tubercles in anterior and posterior rows, respectively. Posterior gonopods 6 - segmented, with two apical stylets.</paragraph>
</subSubSection>
<subSubSection id="SECID0EDHBG" type="diagnosis">
<paragraph id="DA02020A9CF0A4C7A3145958C2DD50AE">
<heading id="87E2C4889D32FA123F40FCE261A2E988" reason="title">Diagnosis remarks.</heading>
</paragraph>
<paragraph id="A4C68A7CA43FE17D7E2D7854BF1BBFA6">
The new species is very similar with
<taxonomicName id="F17A5386B6A6757A3DB672C8282428C2" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="camptotrichus">
<emphasis id="EA721E0A2B3E9FC5003016D411483762" italics="true">P. camptotrichus</emphasis>
</taxonomicName>
by coloration (dark yellow vs earth brown), but it differs from
<taxonomicName id="25D3AD8B3E490E8788F58EB79C413B97" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="FEB656CB58CDDDA8C88DDD47C2648796" italics="true">P. variegatus</emphasis>
</taxonomicName>
in body coloration (dark yellow vs yellow-brown with black spots). In addition, it is smaller than
<taxonomicName id="EFBF1156BD24A67546ACBD94897E9227" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="C16F95CD1CC624C93CB1BF1D4948D0B8" italics="true">P. variegatus</emphasis>
</taxonomicName>
and
<taxonomicName id="FB6BFC3996855C7098385C060B178A8E" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="camptotrichus">
<emphasis id="B8F1064A80A784DC8024ABCB52F22987" italics="true">P. camptotrichus</emphasis>
</taxonomicName>
(length:
<quantity id="15762CAC52FAC41CE641F85EE091F889" metricMagnitude="-3" metricUnit="m" metricValue="9.33" unit="mm" value="9.33">9.33 mm</quantity>
vs
<quantity id="DCE172978134C998F190844E3D64BAA7" metricMagnitude="-2" metricUnit="m" metricValue="3.2" unit="mm" value="32.0">32 mm</quantity>
and
<quantity id="FEA309FE9E6CDC503F5DC4AE7114DB40" metricMagnitude="-2" metricUnit="m" metricValue="1.4" unit="mm" value="14.0">14 mm</quantity>
; width:
<quantity id="FD828731E6ED1E3556214C35DB02DF78" metricMagnitude="-3" metricUnit="m" metricValue="1.57" unit="mm" value="1.57">1.57 mm</quantity>
vs
<quantity id="53836B6CBED3D508F48DA87EDC30E2CF" metricMagnitude="-3" metricUnit="m" metricValue="5.8" unit="mm" value="5.8">5.8 mm</quantity>
and 2.0 mm, respestively), having less number of male body rings (42 vs 5356 and 3742), and number of metazonal tubercles (two rows of 34 + 34 and 45 + 45 tubercles vs two rows of 815 + 815 and 57 + 57, and two rows of 69 + 69 each row, respectively).
</paragraph>
<paragraph id="4C1405DC6434440B12F3442F8D98277C">
<taxonomicName id="48CD8A09ADE6C75C091881F649D699BF" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="condao" status="sp. nov.">
<emphasis id="BFBA025C4284ECB36B4ABBE33C43FC51" italics="true">Pseudodesmus condao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2F0006C5465D99B3F00EAC06B1DE2F4" rank="species">sp. nov.</taxonomicNameLabel>
is differentiated from above three new species,
<taxonomicName id="9C43764F308A8D011136FF21D8024174" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
<emphasis id="CC57425851FCDC822E1588F0757E109A" italics="true">P. bidoup</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6840F1FCF88BF11F9D374FD54774283" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="F561615B92271387A12D054631090D75" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov">
<emphasis id="F759FF7A6BF9E91CED8A12F51751C805" italics="true">P. ngoclinh</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CBD41F338029F935A711327D6859E40A" rank="species">sp. nov</taxonomicNameLabel>
and
<taxonomicName id="8C8CD77D3F0C536FCE5747685A6E7A07" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="karstomus" status="sp. nov.">
<emphasis id="729AC9F7EC6AC4DF3BE93830159BAB0D" italics="true">P. karstomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECF9218CD0B4D9D8CA3C9C8F25A7C4BB" rank="species">sp. nov.</taxonomicNameLabel>
in body coloration of generally uniformly dark yellow, smaller body size and number of stylets (two vs four, two and four, respectively).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDKBG" type="description">
<paragraph id="B86731817815B46DF4905F375B230DBA">
<heading id="7D8F0870EF227AB781D71E5D881B07BF" reason="title">Description.</heading>
</paragraph>
<paragraph id="25AAE83A2F09BF2BCA1A78708B023C48">
<emphasis id="7C719A8ADFC3E87F20E120E9E65DF139" bold="true">
Male
<typeStatus id="B6CCC4D8E72A36CF8F7E61056DDA94AA" type="holotype">holotype</typeStatus>
.
</emphasis>
</paragraph>
<paragraph id="D4367E228C041235E76A922CED395BFF">
<emphasis id="A714164CB8EDE7CEA8E7012FF2619FEE" bold="true" italics="true">Measurements</emphasis>
: 42 body rings plus telson, length about
<quantity id="54B35D8D1AF3E44CBC49864A6D036B0E" metricMagnitude="-3" metricUnit="m" metricValue="9.33" unit="mm" value="9.33">9.33 mm</quantity>
, length of midbody metazona ca.
<quantity id="CF56A58597D3A53659C61AAB08319167" metricMagnitude="-4" metricUnit="m" metricValue="2.4" unit="mm" value="0.24">0.24 mm</quantity>
, width of midbody metazona ca.
<quantity id="CF69AF4CA73312B8248D5810B786433C" metricMagnitude="-3" metricUnit="m" metricValue="1.57" unit="mm" value="1.57">1.57 mm</quantity>
, width of midbody prozona ca.
<quantity id="B7B492919B261BF485251CA3E42DA45B" metricMagnitude="-4" metricUnit="m" metricValue="6.3" unit="mm" value="0.63">0.63 mm</quantity>
; ratio of width of head and width of collum = 0.89.
</paragraph>
<paragraph id="DAE86DEA4B46CB22FFD45E995029382A">
<emphasis id="F4D6D1C53B82749F098F03B3B24E9C97" bold="true" italics="true">Coloration</emphasis>
of ethanol-preserved specimens: dark yellow, little darker in the middle.
</paragraph>
<paragraph id="2E695D282EFB682494889216A15D43A2">
Head distinctly smaller than collum. Antenna stout, clavate,
<emphasis id="E1DA4A1946E438BEB2BD799FDAFA1DE6" italics="true">in situ</emphasis>
extending to body ring 4; antennomere 6&gt; 5&gt; 3&gt; 4 = 2&gt; 7 =
<quantity id="4C3823B4F8260594AA0259BCEBDB2E15" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
length. Collum (Figs
<figureCitation id="E2F4AA314662073B1781202C31BD3BAC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure15" httpUri="https://binary.pensoft.net/fig/1169905">15 A</figureCitation>
,
<figureCitation id="46D1AA4CF16920D3C134D706841A95A5" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Head and collum, ventral view; E. Anterior part of body, ventral view; F. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure16" httpUri="https://binary.pensoft.net/fig/1169906">16 A</figureCitation>
,
<figureCitation id="5C00087D2C4F2CE0E6AC9A32F85323FE" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure17" httpUri="https://binary.pensoft.net/fig/1169907">17 A</figureCitation>
,
<figureCitation id="1154BDBB08183E56FDEF0DE298E33FB9" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Posterior part of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure18" httpUri="https://binary.pensoft.net/fig/1169908">18 A</figureCitation>
) bilobed, each with three anterior lobes, no tubercles. Body rings 24 one row of tubercles, then each body ring with 2 rows, body rings 58 with 1 + 1 larger in the middle, then 2 + 2 larger tubercles, both in anterior and posterior rows (Figs
<figureCitation id="38F581C36CD25354813A45F18C1C1A48" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure15" httpUri="https://binary.pensoft.net/fig/1169905">15 A</figureCitation>
,
<figureCitation id="44921A236B667CE382C3A4EFB5461CBF" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Head and collum, ventral view; E. Anterior part of body, ventral view; F. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure16" httpUri="https://binary.pensoft.net/fig/1169906">16 A C</figureCitation>
,
<figureCitation id="AFF05E1F2A5D753ABDA004AF189790D0" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure17" httpUri="https://binary.pensoft.net/fig/1169907">17 A</figureCitation>
,
<figureCitation id="A38C49CB5AB8EEA95F236C469031D3BB" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Posterior part of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure18" httpUri="https://binary.pensoft.net/fig/1169908">18 A, C</figureCitation>
). Additionally, 23 small tubercles in both rows laterally, but only on tergites, not extending to paraterga (Figs
<figureCitation id="7371FEDC3627D267A029761676E8406C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure15" httpUri="https://binary.pensoft.net/fig/1169905">15 A</figureCitation>
,
<figureCitation id="8AF3F807BA554946BA08AF733A268D9D" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Head and collum, ventral view; E. Anterior part of body, ventral view; F. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure16" httpUri="https://binary.pensoft.net/fig/1169906">16 A C</figureCitation>
,
<figureCitation id="3A85FDE9BF9CDFB65D6060E0466938E0" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure17" httpUri="https://binary.pensoft.net/fig/1169907">17 A</figureCitation>
,
<figureCitation id="30422F43CDB5AD0AE1DBB1ED9218C2D2" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Posterior part of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure18" httpUri="https://binary.pensoft.net/fig/1169908">18 A, C</figureCitation>
).
</paragraph>
<caption id="F3CA22CF339EFF558B9ECF9E6BA00BD8" ID-DOI="10.3897/zse.100.132111.figure15" ID-arpha="04525CBA-8400-5557-B1A6-05149846E842" httpUri="https://binary.pensoft.net/fig/1169905" startId="F15">
<paragraph id="F0EE6E8FEE2B4A7DF88869BB49BB7112">
<label id="C84BA5BEC7966E292AB1E98161EB843F">Figure 15.</label>
</paragraph>
<paragraph id="3AB597B93AFC641C0D4985725313B5BF">
<taxonomicName id="AD68DAA0CF7E501FFCD0F2B345DCC2D0" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="condao" status="sp. nov.">
<emphasis id="ACB936A1491D77A38C2F0059D6A2402B" italics="true">Pseudodesmus condao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F569E989964C0327BBC859540B3C536" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EANBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
982 H), under white light.
<emphasis id="D822BAF988A67D652F7817F8EF5E903B" bold="true">A, B.</emphasis>
Habitus, dorsal view (
<emphasis id="E0721FB73156D405B5B9D91BBB479A66" bold="true">A</emphasis>
), ventral view (
<emphasis id="4AA65F067577F64E4E628F9B0BD802F4" bold="true">B</emphasis>
). Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="F7B3A63972069918BF2306688ABC7D5C" ID-DOI="10.3897/zse.100.132111.figure16" ID-arpha="72417E37-061C-594D-A74E-9409D78A251E" httpUri="https://binary.pensoft.net/fig/1169906" startId="F16">
<paragraph id="9B37E21046CFEB13A70374DE99D27D54">
<label id="161706C1B3410FBA84F6B5EE79B92770">Figure 16.</label>
</paragraph>
<paragraph id="7C52C1B1EE5B6AB6A96F12EC51E8D93D">
<taxonomicName id="AFC0A118781F8C02E23A16B5EB237AE3" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="condao" status="sp. nov.">
<emphasis id="52AFB03DC1D08D1E3D622B69BE081041" italics="true">Pseudodesmus condao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BC30163523104A3D054EAF780254FBBB" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EDOBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
982 H), under white light.
<emphasis id="FAA57B08C399AA96167595C23531AD80" bold="true">A.</emphasis>
Anterior part of body, dorsal view;
<emphasis id="2F4C30239E4A194C11F9FA93C89D3EAF" bold="true">B.</emphasis>
Midbody body rings, dorsal view;
<emphasis id="A61C84A9A5299CD39107450BD37110AC" bold="true">C.</emphasis>
Posterior part of body, dorsal view;
<emphasis id="5F9341DD59616F59F803FB48C68224F4" bold="true">D.</emphasis>
Head and collum, ventral view;
<emphasis id="654BCAE4F16F7B8BC33D7F349C0AEAD6" bold="true">E.</emphasis>
Anterior part of body, ventral view;
<emphasis id="3EA10B9200532039ACDA313C66E5DDF7" bold="true">F.</emphasis>
Posterior part of body, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="D35BED2C1DD8123D1B807CDF52299022" ID-DOI="10.3897/zse.100.132111.figure17" ID-arpha="6E334B3D-2FCD-57BC-9FD9-CB09565BC195" httpUri="https://binary.pensoft.net/fig/1169907" startId="F17">
<paragraph id="C0DD66778670258DE23133FFE087741A">
<label id="65227959BA647BAE4492102F448CF80C">Figure 17.</label>
</paragraph>
<paragraph id="A3C5D6F219F35E1F867753226E2243BF">
<taxonomicName id="1C15941656A2BC38BA3C53DB3DC2435C" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="condao" status="sp. nov.">
<emphasis id="57DC50AB48C646A3D660389CDC8012F8" italics="true">Pseudodesmus condao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EA4E83A9632DC2E9B232CE2634D8118" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EMPBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
982 H), under UV light.
<emphasis id="24D00846CCD3A3E09523BCD6732745D0" bold="true">A, B.</emphasis>
Habitus, dorsal view (
<emphasis id="7D6E5C11651142074D80930FAD186214" bold="true">A</emphasis>
), ventral view (
<emphasis id="72FD2314442449149F75D1D1FA8560A2" bold="true">B</emphasis>
). Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="C9F76EC6E063083A97C66B550E74317D" ID-DOI="10.3897/zse.100.132111.figure18" ID-arpha="24B422C5-436C-5688-8835-B044B9CB3CA2" httpUri="https://binary.pensoft.net/fig/1169908" startId="F18">
<paragraph id="AB7881D502151152AB23FA4A1620998E">
<label id="BAB45B98A71CC8E029FD8D7D4967EF26">Figure 18.</label>
</paragraph>
<paragraph id="02C71ED3B939642784E40D7204FD7C6B">
<taxonomicName id="1C67ADE4E0A7381D58BF5CF31519B2E3" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="condao" status="sp. nov.">
<emphasis id="5F9BAFB2BD1C712D7A3BF040D4334CE5" italics="true">Pseudodesmus condao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4450F9E29F95C50B978097637293F57" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EPQBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
982 H), under UV light.
<emphasis id="D5EC5F1EC53233C08E6DD6CA62860665" bold="true">A, B.</emphasis>
Anterior part of body, dorsal view (
<emphasis id="CFDE3D95879EBE11FFF44AAFA89CEC51" bold="true">A</emphasis>
), ventral view (
<emphasis id="F781C10C6D0FEBF78C444AE0EC3EC45A" bold="true">B</emphasis>
);
<emphasis id="02B40C550966CDAA50AB44E9F078080C" bold="true">C, D.</emphasis>
Posterior part of body, dorsal view (
<emphasis id="00086F92E3DDB0FC3CF21ECA31A0FFAE" bold="true">C</emphasis>
), ventral view (
<emphasis id="907F04A54951C4B4C64CC373E6364C48" bold="true">D</emphasis>
). Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="7C70072A3FD3738FE2E06C65A9C795C5">
Paraterga (Figs
<figureCitation id="86B5EFFE60AEAF5937F498755DF6967D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure15" httpUri="https://binary.pensoft.net/fig/1169905">15</figureCitation>
<figureCitation id="E14E059B132D58A22434E9724108CE23" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Posterior part of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure18" httpUri="https://binary.pensoft.net/fig/1169908">18</figureCitation>
) smooth, almost transparent, wider than in the other species, a little bit spoon form (the distal part is wider than the proximal part), especially in the anterior body ring (up to 10
<superScript id="09DC50EF630220B30D983B61547F2B95">th</superScript>
), later tapering towards the edge, anterior edge is bending backwards, posterior edge remains transverse. Anterior and lateral margin of paraterga serrated (Figs
<figureCitation id="EDA590D0D6D96F74F6700567EF2143F0" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure17" httpUri="https://binary.pensoft.net/fig/1169907">17 A</figureCitation>
,
<figureCitation id="FE6DCD99EAD3C9B28F44D6940B0D82F5" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Posterior part of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure18" httpUri="https://binary.pensoft.net/fig/1169908">18</figureCitation>
).
</paragraph>
<paragraph id="E37135F67BE341E8D03FAEB77A756EDC">
Telson (Figs
<figureCitation id="BBE0747E43F81D9A7EA9BCC07C1B91AA" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under white light. A. Anterior part of body, dorsal view; B. Midbody body rings, dorsal view; C. Posterior part of body, dorsal view; D. Head and collum, ventral view; E. Anterior part of body, ventral view; F. Posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure16" httpUri="https://binary.pensoft.net/fig/1169906">16 C, F</figureCitation>
,
<figureCitation id="E4909B72D3AFF88DAD13E4E89A685F5E" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure17" httpUri="https://binary.pensoft.net/fig/1169907">17 C, D</figureCitation>
) short, caudal margin rounded; epiproct with 2 + 2 small / tiny tubercles at caudal margin. Paraprocts and hypoproct semi-circular.
</paragraph>
<paragraph id="D7C0AA650518883A100F3B6DE471BB6F">
Legs slender, shorter than width of body ring together with paraterga, terminating before lateral paratergal margins. Claws normal. Coxal projections present until leg 71 (body ring 38) (Figs
<figureCitation id="8462A6A57081302D9436E8F64CC442F7" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Habitus, dorsal view (A), ventral view (B). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure17" httpUri="https://binary.pensoft.net/fig/1169907">17 B</figureCitation>
,
<figureCitation id="7F90C4136E3179C27AE67279EE976A94" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Posterior part of body, dorsal view (C), ventral view (D). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure18" httpUri="https://binary.pensoft.net/fig/1169908">18 D</figureCitation>
).
</paragraph>
<paragraph id="6CBF6D277D5EF289E4D64183EC253BE6">
<emphasis id="75A0034973C1F17692C9F548528F5C93" bold="true" italics="true">Gonopods</emphasis>
(Figs
<figureCitation id="A88D2335E032EDF17899FF54721F139A" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. A. Left anterior gonopods, anterior view; B. Left posterior gonopod, subventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure19" httpUri="https://binary.pensoft.net/fig/1169909">19</figureCitation>
,
<figureCitation id="45E5594CACE2FA4BDC0A45E707EE5ED1" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. SEM. A, B. Left posterior gonopod, subventral view (A), anterior view (B); C. Anterior gonopods, anterior view; D. Left anterior gonopod, anterior view; E. Distal part of left posterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure20" httpUri="https://binary.pensoft.net/fig/1169910">20</figureCitation>
): Two pairs of gonopods directed anteriad. Anterior gonopods (Figs
<figureCitation id="CC9344610E0A05F0671C9AA773FA0FE4" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. A. Left anterior gonopods, anterior view; B. Left posterior gonopod, subventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure19" httpUri="https://binary.pensoft.net/fig/1169909">19 A</figureCitation>
,
<figureCitation id="C0EA9E50CFDF29B0405E501C5A4266F2" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. SEM. A, B. Left posterior gonopod, subventral view (A), anterior view (B); C. Anterior gonopods, anterior view; D. Left anterior gonopod, anterior view; E. Distal part of left posterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure20" httpUri="https://binary.pensoft.net/fig/1169910">20 C, D</figureCitation>
) 6 - segmented, covered with long setae; coxite (
<emphasis id="9483FB455F683965B47571262604D933" italics="true">co</emphasis>
) broad, basal part sparsely covered with microgranulations; podomeres 25, short and stout; podomere 6 slightly longer than other podomeres, distally carrying macrosetae (
<emphasis id="668F05BB7B24DA86F83E48D0E3CBB8D6" italics="true">mse</emphasis>
). Posterior gonopods (Figs
<figureCitation id="1684BEE074235191FE8F653B88F138AD" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. A. Left anterior gonopods, anterior view; B. Left posterior gonopod, subventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure19" httpUri="https://binary.pensoft.net/fig/1169909">19 B</figureCitation>
,
<figureCitation id="4ABB309ADAF6FFEB194C901263CF4EA5" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. SEM. A, B. Left posterior gonopod, subventral view (A), anterior view (B); C. Anterior gonopods, anterior view; D. Left anterior gonopod, anterior view; E. Distal part of left posterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure20" httpUri="https://binary.pensoft.net/fig/1169910">20 A, B, E</figureCitation>
) 6 - segmented, covered with sparse long setae; coxite (
<emphasis id="AD080C448F4F11478E485E0BAE163F4E" italics="true">co</emphasis>
) broad, basal part sparsely covered with microgranulations; podomeres 25 short and stout; ultimate podomere (
<emphasis id="FD0D4C286B213FFF9C30F212F4A71921" italics="true">po 6</emphasis>
) longest, with two apical stylets.
</paragraph>
<caption id="CD234E9500E71CD4A12B9F46CCE6359A" ID-DOI="10.3897/zse.100.132111.figure19" ID-arpha="93C35AF0-5736-5ECF-864D-EB0044163596" httpUri="https://binary.pensoft.net/fig/1169909" startId="F19">
<paragraph id="E6C1FE731FD55F85AF92D0724CB386DB">
<label id="010854C63B85627CBB371EA826DE4AB5">Figure 19.</label>
</paragraph>
<paragraph id="DAAD14760D698F534FB35E9198F257E9">
<taxonomicName id="D505623D2996787565CD77C7D9C29476" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="condao" status="sp. nov.">
<emphasis id="9ADE27B8454B8E6A6B1F67E2A75DF43B" italics="true">Pseudodesmus condao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFCF69700F2F64EEB7C38C4B38318369" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EHUBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
982 H), gonopods.
<emphasis id="89D1074F8DA59938CDE5AA3B15BE7D94" bold="true">A.</emphasis>
Left anterior gonopods, anterior view;
<emphasis id="0AEC5848F379D7066139768B90E64682" bold="true">B.</emphasis>
Left posterior gonopod, subventral view.
<emphasis id="AC4778CA2EE96BA85E5E4B1899E977DD" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="13810BD042B2D07567ADDEFA4879A6A4" ID-DOI="10.3897/zse.100.132111.figure20" ID-arpha="26790970-466E-5CF0-89EF-E0E42BEB47E8" httpUri="https://binary.pensoft.net/fig/1169910" startId="F20">
<paragraph id="396CDD1346DA3C984C1FBEB73F3AD8D2">
<label id="C45A465F743B1738E6A4B6600E88141C">Figure 20.</label>
</paragraph>
<paragraph id="F391E925D3E68BBE238471DCFB0964A9">
<taxonomicName id="298A3169BE949B32C82E62BC0F2C7672" authorityName="Nguyen &amp; Nguyen &amp; Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="condao" status="sp. nov.">
<emphasis id="19ECD90EF6D43A18C78D07C004386B10" italics="true">Pseudodesmus condao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95B0F4F756969053B308476298AAE399" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (
<abbrev content-type="institution" id="ABBRID0EKVBG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
982 H), gonopods.
<abbrev id="ABBRID0EPVBG" xlink_title="Scanning electron microscope">SEM</abbrev>
.
<emphasis id="B4C04CC918DF0E3A73256D432E7EF915" bold="true">A, B.</emphasis>
Left posterior gonopod, subventral view (
<emphasis id="EFA985DE849A2A931B4D9EC583333623" bold="true">A</emphasis>
), anterior view (
<emphasis id="7FE4AC83AFA1B43506EF84E9AF8E6247" bold="true">B</emphasis>
);
<emphasis id="FB09B37112C473074F5203E075B1CABB" bold="true">C.</emphasis>
Anterior gonopods, anterior view;
<emphasis id="27A02D383B8FBB53347E20FF32C70A04" bold="true">D.</emphasis>
Left anterior gonopod, anterior view;
<emphasis id="BFC55C14CF26AAEBDBEB9AEF33B590DB" bold="true">E.</emphasis>
Distal part of left posterior gonopod, anterior view.
<emphasis id="161709D665115EAACFF6E61AC7B589B5" italics="true">Abbreviations</emphasis>
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EGWBG" type="variation">
<paragraph id="CA68716C14D6324308839B2FF4CC76FB">
<heading id="C324E48EDBD0AD16281DBBD82F6B61C0" reason="title">Variation.</heading>
</paragraph>
<paragraph id="B1DFF22E618A9849FDC72FA55D5DB315">
<emphasis id="890FD454E681F7F45DBC95FC3DFA196D" bold="true" italics="true">
Male
<typeStatus id="292EE6CE6384CB89717DD5D6B796226A" type="paratype">paratypes</typeStatus>
</emphasis>
: 4647 body rings plus telson, length about
<quantity id="02260930B8B30C541355639CBC43486D" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.119" metricValueMin="1.081" unit="mm" value="11.0" valueMax="11.19" valueMin="10.81">10.8111.19 mm</quantity>
, length of midbody metazona ca.
<quantity id="4B202070CF45878DCBB19011B9A81375" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" unit="mm" value="0.265" valueMax="0.28" valueMin="0.25">0.250.28 mm</quantity>
, width of midbody metazona ca.
<quantity id="D3310586795A64B6E03AA8E869EEDE52" metricMagnitude="-3" metricUnit="m" metricValue="1.7349999999999999" metricValueMax="1.77" metricValueMin="1.7" unit="mm" value="1.7349999999999999" valueMax="1.77" valueMin="1.7">1.701.77 mm</quantity>
, width of midbody prozona ca.
<quantity id="D76BEC61EC4CC9DB0061E94D7A037689" metricMagnitude="-4" metricUnit="m" metricValue="8.55" metricValueMax="9.6" metricValueMin="7.5" unit="mm" value="0.855" valueMax="0.96" valueMin="0.75">0.750.96 mm</quantity>
; ratio of width of head and width of collum = 0.830.88. (width of collum:
<quantity id="12DF80A45C8FE3EF163A2C176F67B0C6" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="7.4" metricValueMin="6.4" unit="mm" value="0.69" valueMax="0.74" valueMin="0.64">0.640.74 mm</quantity>
, width of head:
<quantity id="E07F7FEE8C95396571734E5678E202B2" metricMagnitude="-4" metricUnit="m" metricValue="5.9" metricValueMax="6.5" metricValueMin="5.3" unit="mm" value="0.59" valueMax="0.65" valueMin="0.53">0.530.65 mm</quantity>
).
</paragraph>
<paragraph id="DF4252FDCEDD5FD1322696BFAB7CD2E1">
<emphasis id="C5D4AC932B40257571CF97A6A1DFDAA5" bold="true" italics="true">
Female
<typeStatus id="A107615D5D7262EC405FDB28829B358B" type="paratype">paratypes</typeStatus>
</emphasis>
: 4849 body rings plus telson, length about
<quantity id="EE0304099005EC72E4CE50EC8AB6393B" metricMagnitude="-2" metricUnit="m" metricValue="1.236" metricValueMax="1.25" metricValueMin="1.222" unit="mm" value="12.36" valueMax="12.5" valueMin="12.22">12.2212.50 mm</quantity>
, length of midbody metazoan ca.
<quantity id="81FCE04101ADA177B0024D080D09F56C" metricMagnitude="-4" metricUnit="m" metricValue="2.6" unit="mm" value="0.26">0.26 mm</quantity>
, width of midbody metazona ca.
<quantity id="706FB2A34D37584186FB231337885CA8" metricMagnitude="-3" metricUnit="m" metricValue="1.76" metricValueMax="1.77" metricValueMin="1.75" unit="mm" value="1.76" valueMax="1.77" valueMin="1.75">1.751.77 mm</quantity>
, width of midbody prozona ca.
<quantity id="A90309A9CFB00B3A1C09A095A5663596" metricMagnitude="-4" metricUnit="m" metricValue="9.7" metricValueMax="9.8" metricValueMin="9.6" unit="mm" value="0.97" valueMax="0.98" valueMin="0.96">0.960.98 mm</quantity>
; ratio of width of head and width of collum = 0.84. (width of collum:
<quantity id="46301EDF57F6065475AABBADB5270454" metricMagnitude="-4" metricUnit="m" metricValue="6.8" metricValueMax="8.0" metricValueMin="5.6" unit="mm" value="0.68" valueMax="0.8" valueMin="0.56">0.560.80 mm</quantity>
, width of head:
<quantity id="3FEDD522A70834DB6E3602F413000787" metricMagnitude="-4" metricUnit="m" metricValue="5.7" metricValueMax="6.7" metricValueMin="4.699999999999999" unit="mm" value="0.57" valueMax="0.67" valueMin="0.47">0.470.67 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYWBG" type="etymology">
<paragraph id="315817884A657D29818ADDC57ABCD356">
<heading id="DAC2629D2662B5A0EB010AB49A264A24" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="FC61D9F2A8C8748DE76BBB008B43D756">
Named after its
<typeStatus id="BCE86DC959F48E15702F11BAB66A4A72">type</typeStatus>
locality, Con Dao National Park. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4WBG" type="distribution">
<paragraph id="631A4A78C2C6C258DA4B15F6DF9D2008">
<heading id="B7223C6553D345B7F5F9DCDAFEA68B82" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="9A0B31BE07F776957360CD21767BB5D8">
The species was found in evergreen forests in Con Dao National Park, Ba Ria Vung Tau Province, southern
<collectingCountry id="D67F3D5EF5F3759F3BA6ECC527196C1C" name="Vietnam">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECXBG" type="DNA barcoding">
<paragraph id="2EF2FE7C40BA2F564A6D2134D208FDF4">
<heading id="D82574D36BE556BBBF55D316516CE9BF" reason="title">DNA barcoding.</heading>
</paragraph>
<paragraph id="2C5BA43616287205BA00DDEA5746FCAB">
A fragment of the
<abbrev id="ABBRID0EIXBG" xlink_title="cytochrome c oxidase subunit I">COI</abbrev>
gene is accessioned at
<collectionCode id="A0A1C48456E96064A1FA50717B7D6B78">NCBI</collectionCode>
GenBank with the following accession numbers:
<ext-link id="2FC7B45DC7D1D0357C55ED53564DA18A" ext-link-type="gen" xlink_href="PQ423226" xlink_type="simple">PQ 423226</ext-link>
<ext-link id="F982905FB7727E5D0E6A9CED6EC11533" ext-link-type="gen" xlink_href="PQ423227" xlink_type="simple">PQ 423227</ext-link>
. The new species has a close
<abbrev id="ABBRID0EWXBG" xlink_title="cytochrome c oxidase subunit I">COI</abbrev>
gene similarity with
<taxonomicName id="D3E585A3096CC2BDE5A8092AC1963C63" authorityName="Chamberlin" authorityYear="1922" class="Diplopoda" family="Andrognathidae" genus="Gosodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="claremontus">
<emphasis id="182149D189EC6D5E8FF7A09FA0F8AF1F" italics="true">Gosodesmus claremontus</emphasis>
</taxonomicName>
(
<ext-link id="344B15CAF00541AFBB6F7F90B2341B7C" ext-link-type="gen" xlink_href="JX962723" xlink_type="simple">
<accessionNumber id="48C0ABC0D95F22301E7117C31C88BB99" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JX962723">
<collectionCode id="4AA402FB0699FDF36949956C75F8AE62">JX</collectionCode>
962723
</accessionNumber>
</ext-link>
) of 87.46 % in the
<collectionCode id="BC5D3E9D0E1E3C502208A464735DF1B6">BLAST</collectionCode>
search.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,497 @@
<document id="4D073002547300ED812CC67C56FE916B" ID-DOI="10.3897/phytokeys.248.130311" ID-publisher-id="130311" URI-arpha="5AD7E4B7-F0FC-513D-8CBF-FB0C2FDF56DB" article-type="research-article" checkinTime="1731001061380" checkinUser="pensoft" docAuthor="Ngernsaengsaruay, Chatchai &amp; Chanton, Pichet" docDate="2024" docId="D5FC3CF09433530A86FAD95ABDDD8B9A" docLanguage="en" docName="PhytoKeys 248: 339-360" docOrigin="PhytoKeys 248" docSource="https://phytokeys.pensoft.net/article/130311/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Garcinia nuntasaenii Ngerns. &amp; Suddee, Thai Forest Bull., Bot." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="360" masterDocId="5AD7E4B7F0FC513D8CBFFB0C2FDF56DB" masterDocTitle="A taxonomic revision of Garcinia sections Dicrananthera and Macrostigma (Clusiaceae) in Thailand" masterLastPageNumber="360" masterPageNumber="339" pageNumber="339" updateTime="1731001138325" updateUser="pensoft">
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<mods:title id="D251E4786FCB3B7685587A9F2089EE75">A taxonomic revision of Garcinia sections Dicrananthera and Macrostigma (Clusiaceae) in Thailand</mods:title>
</mods:titleInfo>
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<mods:namePart id="76FF1D1B579CCD9434ECE8E56FCE41EA">Ngernsaengsaruay, Chatchai</mods:namePart>
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<mods:namePart id="22BE479B32789B315B9EE7F1E149D0B7">Chanton, Pichet</mods:namePart>
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<mods:affiliation id="A715CD46699471908C99E210E9A5F4F4">Biodiversity Center, Kasetsart University (BDCKU), Chatuchak, Bangkok 10900, Thailand</mods:affiliation>
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</mods:mods>
<treatment id="D5FC3CF09433530A86FAD95ABDDD8B9A" ID-arpha="D5FC3CF0-9433-530A-86FA-D95ABDDD8B9A" LSID="urn:lsid:plazi:treatment:D5FC3CF09433530A86FAD95ABDDD8B9A" httpUri="http://treatment.plazi.org/id/D5FC3CF09433530A86FAD95ABDDD8B9A">
<subSubSection id="493205FC8BDF11CF2D42A44EE51A57A1" type="nomenclature">
<paragraph id="169A3BFEE969DE4E87884F767EAEA952">
<heading id="EC505637A72FABF980EE821ADE815CFE" reason="label">1.</heading>
<taxonomicName id="E599077B049B8D3E1CA24067DA3C1659" ID-CoL="6K924" authority="Ngerns. &amp; Suddee, Thai Forest Bull., Bot." authorityName="Ngerns. &amp; Suddee, Thai Forest Bull., Bot." class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nuntasaenii">
<emphasis id="0DC9E03C3571D20FFDA523A5DC681025" italics="true">Garcinia nuntasaenii</emphasis>
Ngerns. &amp; Suddee, Thai Forest Bull., Bot.
</taxonomicName>
44 (2): 134. figs 1, 2, 3. 2016; Tagane et al., Edinburgh J. Bot. 75 (1): 110. fig. 2 G. 2018.
</paragraph>
</subSubSection>
<subSubSection id="878D7C91C27649BA8378C8C498531581" type="description">
<paragraph id="8350D8C22CB8067F6A468FEEEC4F989E">
<figureCitation id="80EBE9980DC58B55FC3749C06291841D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Garcinia nuntasaenii A habit and habitat B branchlets and young leaves C terminal bud concealed between the bases of the uppermost pair of petioles D terminal inflorescences with flower buds E branchlets and male inflorescences in axils of fallen leaves with male flower buds and open male flowers F female flower (top view) G branchlets, leaves, and fruit. Photos: Chatchai Ngernsaengsaruay (A D, F), Manop Poopath (E, G)." figureDoi="10.3897/phytokeys.248.130311.figure5" httpUri="https://binary.pensoft.net/fig/1169587">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFYAG" type="material">
<paragraph id="7765E9E00567C5D9EB116288C251C0BA">
<heading id="77888039B0905EBFD08EFF5A734D9286" reason="title">
<typeStatus id="E64C983EA3AB5C8366F29EBD8BD3C017">Type</typeStatus>
.
</heading>
</paragraph>
<paragraph id="2F7800F5ECE1B814B2961FAE23C49359">
<materialsCitation id="AB16E6DDFF6D26686C9D929C63828BA7" collectingDate="2008-12-13" collectionCode="BKF" collectorName="N. Nuntasaen" country="Thailand" location="Phu Wua Wildlife Sanctuary" municipality="Bung Khla District" specimenCount="2" stateProvince="Bueng Kan" typeStatus="holotype">
<collectingCountry id="8D417F0EA31A8D116E70FFD88E4C6F9D" name="Thailand">Thailand</collectingCountry>
<collectingRegion id="6D8927EEFC260B5EA7E3F3F16E9CC790" country="Thailand" name="Bueng Kan">Bueng Kan Province</collectingRegion>
,
<collectingMunicipality id="C38C4823B5F5E0C1EB5EA6E956C1D139">Bung Khla District</collectingMunicipality>
,
<location id="666F0558E06D79B91125D3C292F47E96" LSID="urn:lsid:plazi:treatment:D5FC3CF09433530A86FAD95ABDDD8B9A:666F0558E06D79B91125D3C292F47E96" country="Thailand" municipality="Bung Khla District" name="Phu Wua Wildlife Sanctuary" stateProvince="Bueng Kan">Phu Wua Wildlife Sanctuary</location>
, fr.,
<date id="BF8E70D8E54C1F0F041E6A581E4BA23B" value="2008-12-13">
<collectingDate id="4CA6BCFD8FF3A9749B7B41FCE72B02DE" value="2008-12-13">13 Dec 2008</collectingDate>
</date>
,
<emphasis id="FB80069D55E6D54B427858028966CB00" italics="true">
<collectorName id="5D140C6135F6C2F28CD697682FAFD7FF">N. Nuntasaen</collectorName>
10
</emphasis>
(
<typeStatus id="C88456152F4F0CFC6FC249B94478FC33" type="holotype">holotype</typeStatus>
<collectionCode id="30BA36BB4EB42E3C8BAF89DE7A11F922" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
!;
<typeStatus id="B115FAF943AD900635B544F597F554A9" type="isotype">isotype</typeStatus>
<collectionCode id="9EA01E232A01F22BCC788DAAD2262D7B" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<caption id="FB53AB6F07028155B15806A489706A77" ID-DOI="10.3897/phytokeys.248.130311.figure5" ID-arpha="46F35E4A-58E1-587E-AD53-E61F2BEE9C0C" httpUri="https://binary.pensoft.net/fig/1169587" startId="F5">
<paragraph id="8B67E5A48E8E2A57370665ECE1304F45">
<label id="3845992D89BFF19503BCC1DA2811CDB7">Figure 5.</label>
</paragraph>
<paragraph id="10AA04BA0705455CAE466E5E1D50D74B">
<taxonomicName id="3CDE1D3A12B31A2818820CCB7EE89C36" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nuntasaenii">
<emphasis id="B2D8E75EB85D6B326FD983D3D09E19C3" italics="true">Garcinia nuntasaenii</emphasis>
</taxonomicName>
<emphasis id="10620F259725DC3D4512FFAB50011F21" bold="true">A</emphasis>
habit and habitat
<emphasis id="6B468D995CF97BB00D175D2251691FDD" bold="true">B</emphasis>
branchlets and young leaves
<emphasis id="106C8DD3CF094C60E464F43418740F84" bold="true">C</emphasis>
terminal bud concealed between the bases of the uppermost pair of petioles
<emphasis id="C108B5449AE87CEFD6ACB1C9AC41D88D" bold="true">D</emphasis>
terminal inflorescences with flower buds
<emphasis id="95A6EC66CB60954AC2FEA01F36DDF348" bold="true">E</emphasis>
branchlets and male inflorescences in axils of fallen leaves with male flower buds and open male flowers
<emphasis id="6DA1A48922803E9DD0C6914AD885896E" bold="true">F</emphasis>
female flower (top view)
<emphasis id="93B11E2194CF072A50EB20CFAA89823F" bold="true">G</emphasis>
branchlets, leaves, and fruit. Photos: Chatchai Ngernsaengsaruay (
<emphasis id="51FC46244CDC38F0F21BCA99D5D5240D" bold="true">A D, F</emphasis>
), Manop Poopath (
<emphasis id="B490406093673E45D842DF50194E2CC1" bold="true">E, G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EXZAG" type="description">
<paragraph id="C21BFA6D93C34CA8F7444105C6D940E4">
<heading id="EF72B2B22BD1C24FF39CD06A95D35885" reason="title">Description.</heading>
</paragraph>
<paragraph id="A2E01744900E47CDBB77D5CFA7B0FFFF">
<emphasis id="6D173C3805114ADC6841CC51B2F60ACD" bold="true" italics="true">Habit</emphasis>
shrub,
<quantity id="F84F0A251DD0017C8F60B2FC13A02578" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">12 m</quantity>
tall; latex white, turning pale yellow, sticky; branches decussate, horizontal or nearly horizontal; branchlets green, 4 - angular, glabrous.
<emphasis id="B90A3BCAA5235734665B467AF90DCDD1" bold="true" italics="true">Bark</emphasis>
green when young, turning dark brown when mature, smooth, usually lenticellate; inner bark pale yellow.
<emphasis id="A887C7215328C9FFF390AD23E1B83022" bold="true" italics="true">Terminal bud</emphasis>
concealed between the bases of the uppermost pair of petioles.
<emphasis id="B9CF8971F63F40F4821C8C4838D20C35" bold="true" italics="true">Leaves</emphasis>
decussate; lamina lanceolate-ovate, ovate or elliptic, 617 ×
<quantity id="21AD6B223472CC488F5654962E107433" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="7.5" metricValueMin="3.0" unit="cm" value="5.25" valueMax="7.5" valueMin="3.0">37.5 cm</quantity>
, apex acuminate or acute and rigid, base subcordate, margin thick, entire and slightly undulate (repand), coriaceous, slightly bullate or bullate, apical part of leaves conduplicate, twisted and recurved, glossy dark green above, paler below, glabrous on both surfaces, midrib shallowly grooved above, raised as a prominent ridge below, secondary veins 1220 each side, curving towards the margin and connected in distinct loops and united into an intramarginal vein, flattened above, slightly raised below, conspicuous on both surfaces, intersecondary veins conspicuous, veinlets reticulate, visible on both surfaces, interrupted long wavy lines present, of differing lengths, nearly parallel to the midrib, running across the secondary veins to the apex, visible below; petiole green,
<quantity id="7D9F787CD268B241E5E92B430D2CA3CF" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" unit="cm" value="0.6" valueMax="1.0" valueMin="0.2">0.21 cm</quantity>
long, not grooved, transversely rugose, glabrous, with a basal appendage clasping the branchlet; young leaves red.
<emphasis id="C3EB689A82E0BB8F74B2DA3193BBA237" bold="true" italics="true">Inflorescences</emphasis>
terminal or at leafless nodes (in axils of fallen leaves), cymose.
<emphasis id="57B2D435D568E7B2AA8E9058FB9D7BDA" bold="true" italics="true">Flowers</emphasis>
unisexual, plants dioecious, 4 - merous, fully open flowers
<quantity id="D2C8B0F2B44AFB8515AD84304259D83F" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" unit="cm" value="0.9" valueMax="1.0" valueMin="0.8">0.81 cm</quantity>
in diam.; bracteoles narrowly triangular or triangular, 210 ×
<quantity id="301B301510A8A3167102A4E58F843596" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 mm</quantity>
, apex acute or acuminate, somewhat thick; pedicels short; sepals 4 and petals 4, decussate, pale yellow or creamish white, somewhat thick; sepals obovate or elliptic, 36 ×
<quantity id="96253AE8CB97E0B108781367610EDD73" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.53.5 mm</quantity>
, concave, apex rounded; petals broadly obovate, 5.58.5 ×
<quantity id="FB49F81AB901AC2956494C7A3D749000" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="7.0" metricValueMin="3.5" unit="mm" value="5.25" valueMax="7.0" valueMin="3.5">3.57 mm</quantity>
, apex rounded, apical part recurved.
<emphasis id="8A1C3F9022D025F07CEBEF7A62AA8DFB" bold="true" italics="true">Flower buds</emphasis>
: sepals pink or pink-pale yellow.
<emphasis id="CCCD5FAF1A9BBA3CF86245C7E31F1F5C" bold="true" italics="true">Male flowers</emphasis>
in fascicles of several simple cymes; stamens numerous, united into a single weakly 4 - lobed bundle; filaments very short; anthers
<quantity id="B502B298A632E9E3CF1FDCC8EF88B0D3" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.30.5 mm</quantity>
long; pistillode usually absent.
<emphasis id="EC48609117F3A2F825431F8941A87F8F" bold="true" italics="true">Female flowers</emphasis>
usually in a cluster of 3 flowers (a simple cyme); staminodes numerous, united into a single weakly 4 - lobed bundle, surrounding the ovary; pistil fungiform; ovary depressed globose, 1.52 ×
<quantity id="2A43D64E02BE4538DF5BB95D80BD553D" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
, shallowly 46 - lobed, 46 - locular; stigma convex, hemispherical, 1.52 ×
<quantity id="EAA8C0C7C7D5BBC82572178D86D936B0" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.53.5 mm</quantity>
, papillate.
<emphasis id="D5612AE63DDA9391C9986CEA38CA387C" bold="true" italics="true">Fruits</emphasis>
berries, depressed globose, 0.50.7 ×
<quantity id="C7224DD95800F760CEC84970CFC520FF" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">12 cm</quantity>
, 46 - lobed, green with white dots, turning red when ripe, smooth, glabrous, glossy, with persistent sepals; persistent stigma circular, flat, radiately lobed or unlobed; fruiting stalk
<quantity id="EA4D39C7FEC7654C715DE17D19C41FB3" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
. long.
<emphasis id="CD4C7CD661937E3D649D90ED4B8ED4E7" bold="true" italics="true">Seeds</emphasis>
46, with fleshy pulp. The description of this species is based on
<bibRefCitation id="F7898072A32D0F28B7CC6862DEA104E7" DOI="10.20531/TFB.2016.44.2.09" author="Ngernsaengsaruay and Suddee" firstAuthor="Ngernsaengsaruay" issue="2" journalOrPublisher="Thai Forest Bulletin" pagination="134-139" refId="B20" refString="Ngernsaengsaruay C, Suddee S (2016) Garcinia nuntasaenii (Clusiaceae), a new species from Thailand. Thai Forest Bulletin 44 (2): 134139. https://doi.org/10.20531/TFB.2016.44.2.09" title="Garcinia nuntasaenii (Clusiaceae), a new species from Thailand." volume="44" year="2016">Ngernsaengsaruay and Suddee 2016</bibRefCitation>
, which includes additional information.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE2AG" type="distribution">
<paragraph id="B09F08BD339F9DAC6A43EBC26F1A1125">
<heading id="782A33C7B002EF3E3B546C62BC41A01A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="3CA8066ACDCD74FA0F1D769ED1712B89">
Central
<collectingCountry id="BA633C31281566C41B55A2F6B69318BB" name="Laos">Laos</collectingCountry>
and North-Eastern
<collectingCountry id="E6AA0E08171ACD4E37CA0E37C4663ADA" name="Thailand">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ2AG" type="distribution">
<paragraph id="EB9B2324DABC10F6EE9919469833B8D3">
<heading id="C2FF5337094A9D132C184BBC91E84576" reason="title">
Distribution in
<collectingCountry id="47E9C0372161ABDAB724B8407515B558" name="Thailand">Thailand</collectingCountry>
.
</heading>
</paragraph>
<paragraph id="C38978A9477B15E7D9CCE321E8936130">
<emphasis id="ACD39CC8D5BF3448ED34650B691D71D1" bold="true">North-Eastern</emphasis>
:
<collectingRegion id="5C8B7F54FE0F4A3E2B51C533581BE201" country="Thailand" name="Bueng Kan">Bueng Kan</collectingRegion>
,
<collectingRegion id="F9634006A205039E24B8B88E8EAA86CA" country="Thailand" name="Nakhon Phanom">Nakhon Phanom</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ER2AG" type="habitat">
<paragraph id="D46EFF48864BB8651B5010985CB408C8">
<heading id="23DB18B441B10730BE53DB7974EC13FA" reason="title">Habitat and ecology.</heading>
</paragraph>
<paragraph id="A11F83212AEAD72FBFB3C6A9A0900E84">
It is found in dry evergreen forests,
<quantity id="D98D775C32F18CD2294C52A174578A60" metricMagnitude="2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" unit="m" value="185.0" valueMax="220.0" valueMin="150.0">150220 m</quantity>
amsl.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EW2AG" type="phenology">
<paragraph id="D1E0E66B9C9C682E244077971E81229E">
<heading id="D1EF497530CEFA784795281AA50A3661" reason="title">Phenology.</heading>
</paragraph>
<paragraph id="67B741488EC670F8064B0743193248C4">Flowering and fruiting more than once; flowering in December to July; fruiting in December to April.</paragraph>
</subSubSection>
<subSubSection id="SECID0E22AG" type="conservation status">
<paragraph id="1DF3C9917083FFED9BD1E3F9CE14024A">
<heading id="57D490B89218DEB1FAB8BAE00D23F4C3" reason="title">Conservation status.</heading>
</paragraph>
<paragraph id="1F220E28A6499425C8F323553C27322F">
<taxonomicName id="2DB7D886A3B6C16A372F30F95EF20AFA" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nuntasaenii">
<emphasis id="462ED8BDBFC2C0487403FE723FBC6B97" italics="true">Garcinia nuntasaenii</emphasis>
</taxonomicName>
is known only from four localities in
<collectingRegion id="89CD091304AF892F09CC2BD57DADE43D" country="Thailand" name="Bueng Kan">Bueng Kan</collectingRegion>
and
<collectingRegion id="E6CE9267FD42029999986DC9912015DB" country="Thailand" name="Nakhon Phanom">Nakhon Phanom</collectingRegion>
Provinces, North-Eastern
<collectingCountry id="E952E84D4F92DE0978AC75B9673FE68A" name="Thailand">Thailand</collectingCountry>
and Nam Kading National Protected Area,
<collectingCountry id="AFCD5CBACC98B8C11A2FC828B382FDFE" name="Laos">Laos</collectingCountry>
. All occurrences are in small populations. It has a small
<abbrev id="ABBRID0EM3AG" xlink_title="Extent of Occurrence">EOO</abbrev>
of
<quantity id="A7CDDEC6DC4B86D29990E405CB3B225A" metricMagnitude="5" metricUnit="m" metricValue="7.9687" unit="km" value="796.87">796.87 km</quantity>
<superScript id="E8038B8C107B47DECB44382F9D01940A">2</superScript>
and a relatively small
<abbrev id="ABBRID0ES3AG" xlink_title="Area of Occupancy">AOO</abbrev>
of
<quantity id="63D0CE9B599A1201A967133AFAF20A3B" metricMagnitude="4" metricUnit="m" metricValue="1.6" unit="km" value="16.0">16 km</quantity>
<superScript id="5D68D7DB88ED7156C334556C5A6CC9EB">2</superScript>
. Because of its number of localities, and because the roots of this species are used for medicinal purpose by nearby villagers in
<collectingCountry id="1166DF02D12C7FDA91F7C8E11C5191DF" name="Thailand">Thailand</collectingCountry>
, the population is suspected to be declining. We therefore suggest the conservation status Endangered [
<abbrev id="ABBRID0EY3AG" xlink_title="Endangered">EN</abbrev>
B 2 ab (v)].
</paragraph>
</subSubSection>
<subSubSection id="SECID0E33AG" type="etymology">
<paragraph id="1D064B068082B2A903E0A766245FFD22">
<heading id="E34DF480FC8476EC05555C8866087F66" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="CCD58A3D707D393DC857A94A6CEB2CC1">
The specific epithet of
<taxonomicName id="E0080B5E8893A6A1D2DAC7800812BDE2" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nuntasaenii">
<emphasis id="49547D3A60CB742BB459A50D71C73632" italics="true">Garcinia nuntasaenii</emphasis>
</taxonomicName>
honours Mr Narong Nuntasaen, a staff member and a plant collector of
<collectionCode id="AF12939626D1BBD93F22996FCA31EF8D" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
, who collected the
<typeStatus id="8774758E84860C09F9DBF0E530E80CF4">type</typeStatus>
specimen.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EN4AG" type="Vernacular names">
<paragraph id="4413DB3C6DB481D7099CA0F77B32B9B6">
<heading id="C2CD54E3A1556404B77D2F5F1244A834" reason="title">Vernacular names.</heading>
</paragraph>
<paragraph id="7DE09DDB488A4D6DF20004918E28622C">
<emphasis id="47CEA9B05AECC3F1887ED7200611A43B" bold="true">Chang nga ek</emphasis>
(ช ้ างงาเอก) (
<collectingRegion id="F851FBCA80DDCE4B16BED0F8484AAECB" country="Thailand" name="Bueng Kan">Bueng Kan</collectingRegion>
,
<collectingRegion id="7650A017C87960EF97DD90C1E6ADAE1A" country="Thailand" name="Nakhon Phanom">Nakhon Phanom</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EV4AG" type="Uses">
<paragraph id="0BCC7FD716E4DA8170955CEFA99588C7">
<heading id="AB08197D3DB0410A724A1B39DA0079C3" reason="title">Uses.</heading>
</paragraph>
<paragraph id="343C3BDC61CE2CB58D07E7D6BC15FB65">
The fruits are edible (from the specimen
<emphasis id="63316FAA4BCFEF2E2522632CA6D23911" italics="true">M. Norsaengsri &amp; N. Tathana 8630</emphasis>
). The roots are locally used for medicinal purposes (Ngernsaengsaruay personal observation).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E44AG" type="notes">
<paragraph id="CDE39EE431504C9C2CCB6BDDA9C3198C">
<heading id="C3107386E08CA1FD70B2CB11CDCF69EE" reason="title">Note.</heading>
</paragraph>
<paragraph id="FFB4C5D3AE5DAA0AD99BCA1F0FB6DCCA">
<taxonomicName id="FB7A2DB42579F2D82BEDDCF5CA9274C6" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="nuntasaenii">
<emphasis id="6B2F7358E37E18255C4AA312217F8068" italics="true">Garcinia nuntasaenii</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F65B3D1A3BEDF3C9750949E221AA604C" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="phuongmaiensis">
<emphasis id="91DE18DBE9D36848BAD5D029CD0B5047" italics="true">G. phuongmaiensis</emphasis>
</taxonomicName>
in its habit (shrubs); sticky white latex, turning pale yellow when exposed to the air; 4 - angular branchlets, especially when young; coriaceous, bullate, shiny dark green, subcordate leaves with a short petiole; flower size, c.
<quantity id="8766DED0D3FAE77649276973208C62DD" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="cm" value="1.0">1 cm</quantity>
in diam.; and the color of fruits, turning red when ripe, but differs in having 4 - merous flowers [vs (4 ) 5 - merous flowers]; pale yellow or creamish white petals (vs white petals); 46 - lobed fruits (vs 34 - lobed fruits); and is also distributed in Central
<collectingCountry id="71A5C280757F9358BE04FD090D54B13C" name="Laos">Laos</collectingCountry>
and North-Eastern
<collectingCountry id="860FB4E7BF3CC948BD80438FEC1DB92A" name="Thailand">Thailand</collectingCountry>
(vs distributed in Central
<collectingCountry id="77ECD0B529DF79FB51B8C947380749C5" name="Vietnam">Vietnam</collectingCountry>
). The morphological characteristics and distribution of
<taxonomicName id="FF32FBA9E81669302F0F6F7678C70585" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="phuongmaiensis">
<emphasis id="4C69BA646C092C46EE84F1F22AE55BB5" italics="true">G. phuongmaiensis</emphasis>
</taxonomicName>
were taken from
<bibRefCitation id="7BB2885FD71D01A1CD9E41CAC9C05B6A" DOI="10.1007/s12225-023-10077-0" author="Tuan DLA &amp; Toyama H &amp; Nguyen XMA &amp; Phan TTN &amp; Truong VB &amp; Dang VS" etAl="et al." firstAuthor="Tuan" issue="2" journalOrPublisher="Kew Bulletin" pagination="189-195" refId="B44" refString="Tuan DLA, Toyama H, Nguyen XMA, Phan TTN, Truong VB, Dang VS (2023) Garcinia phuongmaiensis, a new species of Garcinia sect. Xanthochymus (Clusiaceae) from central Vietnam. Kew Bulletin 78 (2): 189195. https://doi.org/10.1007/s12225-023-10077-0" title="Garcinia phuongmaiensis, a new species of Garcinia sect. Xanthochymus (Clusiaceae) from central Vietnam." volume="78" year="2023">Tuan et al. (2023)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EI6AG" type="material">
<paragraph id="324AB500BFA5FF191E8C5C971F4B97F3">
<heading id="047A2264EC9EDFD4574A707CF3E09F15" reason="title">Additional specimens examined.</heading>
</paragraph>
<paragraph id="FEF9A63BCEC735F11B2A7647CFFE6C53">
<materialsCitation id="1B38CC4956134B0F7B78CB46340706F1" collectingDate="1996-11-08" collectingDateMax="2015-07-24" collectingDateMin="1996-11-08" collectionCode="BKF, QBG" collectorName="C. Niyomdham &amp; R. Pooma &amp; N. Nuntasaen &amp; M. Norsaengsri &amp; N. Tathana &amp; M. Poopath &amp; C. Ngernsaengsaruay &amp; N. Meeprom" country="Thailand" county="Phu Wua Wildlife Sanctuary" location="Upper Northeast Wild Plants Conservation Center" municipality="Bung Khla District" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Bueng Kan">
<emphasis id="9F8F49D5E4A1B2E012957A9133433148" bold="true">
<collectingCountry id="0353F9E8D70FDA2C536AB7990494B606" name="Thailand">Thailand</collectingCountry>
. North-Eastern
</emphasis>
<collectingRegion id="48A763A9EE87EDE23156B5788CE611F0" country="Thailand" name="Bueng Kan">Bueng Kan</collectingRegion>
[
<collectingCounty id="AA2F62BD3E4305AE26784F194E62AC15">Phu Wua Wildlife Sanctuary</collectingCounty>
,
<collectingMunicipality id="EA13EFB111F759D2FC44F1E3DEBEF10D">Bung Khla District</collectingMunicipality>
, fl.,
<date id="61E8AB3813E9FABC0E70CF00370B7FE8" value="1996-11-08">
<collectingDate id="06B63489F7DFA155BB9534C3AE4EC331" value="1996-11-08">8 Nov 1996</collectingDate>
</date>
(
<taxonomicName id="4C35F4A997CCDBBBEB6BDE8F1E20AA57" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="D59506B260B2AD19455926FB58718B25" italics="true">Garcinia</emphasis>
sp.
</taxonomicName>
),
<emphasis id="ECA6B09AB6E87D222E1974CF385D3BE9" italics="true">
<collectorName id="2FA370FB7FF423E0C47C5446E4CE897C">C. Niyomdham</collectorName>
4910
</emphasis>
(
<collectionCode id="83B24BF78EEF90D892B6FAC1B5D0D49F" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
) • ibid., fl.,
<date id="DFA05294EDB7E71012D759712FDB57D3" value="2001-08-27">
<collectingDate id="8A9D683CFF846A096DD8EC340965BBC3" value="2001-08-27">27 Aug. 2001</collectingDate>
</date>
(
<taxonomicName id="6ABA08743804CF839D4576773D4AD0CC" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="D63D4BB909539D6311699A6F464B8BBC" italics="true">Garcinia</emphasis>
sp.
</taxonomicName>
),
<emphasis id="AD578CC88DA11E900FC5FB02280A04A3" italics="true">
<collectorName id="9FD0543B2B9AEAEDE2021F7905338F59">R. Pooma</collectorName>
et al. 2791
</emphasis>
(
<collectionCode id="D5CCF5286D2CA0AEEA91364647FD5A33" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
) • ibid.,
<specimenCount id="4D2A1F204282890D19DF9F6B15A5D06D" count="1" type="male"></specimenCount>
fl.,
<date id="65916F09D0AF4A1A48E4DE07BDDF87C3" value="2008-01-01">
<collectingDate id="AA5244215E4B5002FE2536B934D1B7C0" value="2008-01-01">1 Jan 2008</collectingDate>
</date>
(spirit specimen),
<emphasis id="05CAA10411162B7C8BE2C35EDFF2C020" italics="true">
<collectorName id="E92FFB502FC98EDDF4D2BF8B39B6AE8E">N. Nuntasaen</collectorName>
11
</emphasis>
(
<collectionCode id="C9C864844081C2CA3A66D4D015DD5947" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
) • ibid., fr.,
<date id="BC51BDDAE202C4E7F7E5D6BB0D6DC04E" value="2011-12-27">
<collectingDate id="FB1D225E91B4DEF61D043DEA2FC2D49D" value="2011-12-27">27 Dec 2011</collectingDate>
</date>
(
<taxonomicName id="D34571974AB82A9407CF2DFDA6AEF802" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="EEB2C6B86DD49486090875115B39F820" italics="true">Garcinia</emphasis>
sp.
</taxonomicName>
),
<emphasis id="C0678DBE05D1A743961E8FDCFD0DB2F4" italics="true">
<collectorName id="0E83D736FEE30B416E57A009DF843595">M. Norsaengsri</collectorName>
&amp;
<collectorName id="42BDA5D016760A758A3049205DC86755">N. Tathana</collectorName>
8630
</emphasis>
(
<collectionCode id="8E767FA7E78B4A5C95D32F0E0238EF6C" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
,
<collectionCode id="314AE0DFFB1FB15E1B28C33681E68384" country="Thailand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13640" lsid="urn:lsid:biocol.org:col:13640" name="Queen Sirikit Botanic Garden" type="">QBG</collectionCode>
) • ibid.,
<date id="E361DF3EA4B23659A49D1635D07A5843" value="2015-02-12">
<collectingDate id="BD721E89E0F272DA41BC4E110A5FA134" value="2015-02-12">12 Feb 2015</collectingDate>
</date>
,
<emphasis id="3BB42810B7FEAB916F67CC4521C724A3" italics="true">
<collectorName id="08E70462712E90B7E2B3BB1819EB4B66">M. Poopath</collectorName>
981-1, 981-2
</emphasis>
(
<collectionCode id="05681D1510B73E3646DD69DC759083B5" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
) • ibid.,
<specimenCount id="0902EC89A94162506DD006E6E62E7016" count="1" type="female"></specimenCount>
fl.,
<date id="CDEC92F853D308635CE53F48F9CCF2D2" value="2015-07-24">
<collectingDate id="DEA92F6524D01DB1BEB31ED4A8BB6027" value="2015-07-24">24 Jul 2015</collectingDate>
</date>
,
<emphasis id="3D3293E13C24C4C53B2D0C8422EB560D" italics="true">
<collectorName id="75D69A712E9321D35C53A73153755EB1">C. Ngernsaengsaruay</collectorName>
&amp;
<collectorName id="715B1DF2F1A132F38C9FC30315471C24">N. Meeprom</collectorName>
754
</emphasis>
(
<collectionCode id="FD4626B3F0C0565B301528B9D8F64200" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
) • The
<location id="020DF0C6DF6694AFD9DF46B01C964F0C" LSID="urn:lsid:plazi:treatment:D5FC3CF09433530A86FAD95ABDDD8B9A:020DF0C6DF6694AFD9DF46B01C964F0C" country="Thailand" county="Phu Wua Wildlife Sanctuary" municipality="Bung Khla District" name="Upper Northeast Wild Plants Conservation Center" stateProvince="Bueng Kan">Upper Northeast Wild Plants Conservation Center</location>
,
<location id="AC8D49D2AFA15C7542E9CD3F32BDA39D" LSID="urn:lsid:plazi:treatment:D5FC3CF09433530A86FAD95ABDDD8B9A:AC8D49D2AFA15C7542E9CD3F32BDA39D" country="Thailand" county="Phu Wua Wildlife Sanctuary" municipality="Bung Khla District" name="Bung Khla District" stateProvince="Bueng Kan">Bung Khla District</location>
,
<specimenCount id="CE0F94B93453A8D314576ABB136EE518" count="1" type="male"></specimenCount>
fl.,
<date id="6193D0C38583FF60CE35D18A061DB1A2" value="2015-07-24">
<collectingDate id="3B760F5CF61441762631D00777925DDD" value="2015-07-24">24 Jul 2015</collectingDate>
</date>
,
<emphasis id="9B6C453D18E47A38D2DAB0BF8C1B2208" italics="true">
<collectorName id="FDC6F73D12394E2A2B977E57F50B4CB9">C. Ngernsaengsaruay</collectorName>
&amp;
<collectorName id="55D55CDB99E6F93942B4CF0AF6FD2B11">N. Meeprom</collectorName>
750, 751
</emphasis>
(
<collectionCode id="8B7F248E9CD3F02484FACB537F6E6CD0" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
) • ibid.,
<specimenCount id="B1B19C12CC6CBD155FF38C054D982A60" count="1" type="female"></specimenCount>
fl.,
<date id="A6021BF1CB244CF3C2C1283BBD25E10C" value="2015-07-24">
<collectingDate id="AC58B9023B9C1D5C1CBCB0264631D8E3" value="2015-07-24">24 Jul 2015</collectingDate>
</date>
,
<emphasis id="259EE36A30D897477E6DD307368D4973" italics="true">
<collectorName id="7AA40582AD25C2ACF8D66FBCEB848840">C. Ngernsaengsaruay</collectorName>
&amp;
<collectorName id="5349EB0B6C1252AC03D5EDD76A149732">N. Meeprom</collectorName>
752, 753
</emphasis>
(
<collectionCode id="2A49608F9755386867CA05BA2A1B16B4" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" type="Herbarium">BKF</collectionCode>
)]
</materialsCitation>
<materialsCitation id="9B31FF5D96AADFA281187FC489C9BEC8" country="Thailand" location="Phu Langka National Park" municipality="Ban Phaeng District" specimenCount="1" stateProvince="Nakhon Phanom">
<collectingRegion id="933FC3FF381677CA345A5ADC80E13D5B" country="Thailand" name="Nakhon Phanom">Nakhon Phanom</collectingRegion>
[
<collectingMunicipality id="029EC02213DEA5C24E163083155B4AA1">Ban Phaeng District</collectingMunicipality>
,
<location id="F570CF083866ABEB00A60E726C571104" LSID="urn:lsid:plazi:treatment:D5FC3CF09433530A86FAD95ABDDD8B9A:F570CF083866ABEB00A60E726C571104" country="Thailand" municipality="Ban Phaeng District" name="Phu Langka National Park" stateProvince="Nakhon Phanom">Phu Langka National Park</location>
(Narong Nuntasaen own observation)]
</materialsCitation>
.
</paragraph>
<paragraph id="54D73B93D2BD627AB543FE6E69E4F8A7">
<emphasis id="F2A50114A3DA65364C14363A50CC518D" bold="true">
<collectingCountry id="DD460C84727F47E348643159894A8B62" name="Laos">Laos</collectingCountry>
</emphasis>
• Bolikhamxay [Nam Kading National Protected Area,
<specimenCount id="BB4F427C1E655A58327A4F9ED1161F10" count="1" type="male"></specimenCount>
fl.,
<emphasis id="85A81E13B2A6BE5732F5EAE0C8A0A3F8" italics="true">L 8</emphasis>
• ibid., fl. &amp; fr.,
<emphasis id="0490B826EE81E251EFB43028ED0F1E92" italics="true">L 431</emphasis>
• ibid.,
<emphasis id="CE8B24518DE35E75A2DFEAA58B487C6B" italics="true">L 1034</emphasis>
(
<bibRefCitation id="1CFDB650725EF797E1A1A0BAFB975AE0" DOI="10.1017/S0960428617000361" author="Tagane" etAl="et al." firstAuthor="Tagane" issue="1" journalOrPublisher="Edinburgh Journal of Botany" pagination="107-116" refId="B41" refString="Tagane S, Souladeth P, Rueangruea S, Okabe N, Zhang M, Chayer S, Yang CJ, Yahara T (2018) Flora of Nam Kading National Protected Area II: 30 new records of angiosperms for Laos. Edinburgh Journal of Botany 75 (1): 107116. https://doi.org/10.1017/S0960428617000361" title="Flora of Nam Kading National Protected Area II: 30 new records of angiosperms for Laos." volume="75" year="2018">Tagane et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>