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<mods:titleInfo id="4A1A4D9C49FFED3676F32ABFD51C19A3">
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<mods:title id="A8BD1ED57D88BE8F818098E5EE374D2A">Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India</mods:title>
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<mods:namePart id="1A264B844B82B7375DFA4FC9AD6DF345">Boruah, Bitupan</mods:namePart>
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<mods:affiliation id="E55FB98A4DC5DD7235260DD49EB392FE">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
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<mods:namePart id="77A0AAA55CF4EEF106EBEF40B67D43DF">Narayanan, Surya</mods:namePart>
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<mods:affiliation id="8A6251F72D6EDE949DB952FE6C05933E">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India</mods:affiliation>
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<mods:namePart id="3CFDCB9C96D0885621D5EFE3778D630A">Aravind, Neelavar Ananthram</mods:namePart>
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<mods:nameIdentifier id="EA122EFE14BAFE341B7AB35BB1BF4577" type="ORCID">0000-0002-4515-8421</mods:nameIdentifier>
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<mods:affiliation id="775A7F2AC48CFD635035BA712AFF1B0E">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India & Yenepoya Research Centre, Yenepoya University, Derlakatte, Mangalore, India</mods:affiliation>
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<mods:namePart id="8A81E6AE4AA6C6B3B406EE70265C7C1D">Lalronunga, Samuel</mods:namePart>
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<mods:nameIdentifier id="D174B567E7C2D29498EADBFFAB764338" type="ORCID">0000-0002-7542-4815</mods:nameIdentifier>
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<mods:affiliation id="47F11D477CB170D9F5FED5D85FCB582D">Holy Child Society, Nalkata, Tripura 799263, India</mods:affiliation>
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<mods:namePart id="1E969CFB87BD6C6DFE8F1AF746CC698F">Deepak, V.</mods:namePart>
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<mods:affiliation id="ED40530589F1784FAC94B2B33032D01C">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK & School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK</mods:affiliation>
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<mods:namePart id="D52E863A73E2D6C70D8E48C51286B294">Das, Abhijit</mods:namePart>
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<mods:nameIdentifier id="A05F43E527C8C3A24F2239BC1D4CE2E8" type="ORCID">0000-0002-5851-8457</mods:nameIdentifier>
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<mods:affiliation id="98BCB1FDD81AB137AD1C8AA4BA6DBB6F">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
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<mods:typeOfResource id="C976FFF8AC93C8171B8007B3CB07389D">text</mods:typeOfResource>
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<mods:titleInfo id="D4A492ADFB6A74B6B4ACBB5FDC8C497A">
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<mods:title id="1AD0D33CFA7AEC99514D0C6FD71E16E9">Vertebrate Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="65986592FFD48A62C8B20377ECBA78BB">
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<mods:date id="CBD448CC673DE4D73ED62ED40B74E462">2024</mods:date>
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<mods:number id="7C6443E89BC8C3DA40B0556C82588783">2024-07-29</mods:number>
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<mods:detail id="FD3951D5C3F2D7E3078F958A70E9FDDE" type="volume">
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<mods:number id="A64D5C46DCDC8EFFC455AEE705C347DA">74</mods:number>
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<mods:start id="9F2ABE0BE269BCEC3776B5652FDE678E">453</mods:start>
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<mods:end id="0AD350E609A63A12727AAB7E3DFC84B2">486</mods:end>
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<mods:classification id="8D7FBCEB2AA3DB5AAFC9105C45EDC9F3">journal article</mods:classification>
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<mods:identifier id="87A959E8AC1D2BAD327159359582F855" type="DOI">10.3897/vz.74.e124752</mods:identifier>
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<mods:identifier id="0C0EB2416EEE7AF56159FCAFEF69AD78" type="ZooBank">154CE236-EFA8-4411-834B-234A9B45A63F</mods:identifier>
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<treatment id="507D9749BFBD54018DADAD5538DE287C" ID-DOI="http://doi.org/10.5281/zenodo.13124039" ID-Zenodo-Dep="13124039" ID-arpha="507D9749-BFBD-5401-8DAD-AD5538DE287C" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C" httpUri="http://treatment.plazi.org/id/507D9749BFBD54018DADAD5538DE287C">
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<subSubSection id="C8A7230CE852ED7D1E95ED8BBE46E9C7" type="nomenclature">
|
||||
<paragraph id="72F3CB5D9BCE20CAB94346B42BE7087F">
|
||||
<heading id="8A42322A9212B8C13A4A21A2C9870F1D" reason="label">Expanded description of</heading>
|
||||
<taxonomicName id="C9CE8533B709300C820FD4517F01AB40" ID-CoL="8H73R" authority="Li, 2007" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
|
||||
<emphasis id="926891F2589AF1637DEA5CD2730D25FB" italics="true">Cyrtodactylus cayuensis</emphasis>
|
||||
<bibRefCitation id="2D910A302259869858BEE2C3DD5603BA" author="Li PP" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="733-737" refId="B22" refString="Li PP (2007) Description of a new subspecies of Cyrtodactylus khasiensis from China. Acta Zootaxonomica Sinica 32: 733 - 737." title="Description of a new subspecies of Cyrtodactylus khasiensis from China." volume="32" year="2007">Li, 2007</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="BA6A2F25E6CB04563CFA677285AD63BF" type="description">
|
||||
<paragraph id="0FC529ADE614032C5367AD68577BCF25">
|
||||
<figureCitation id="A5F206DB3B5C978AD34809A3BB205C9C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyrtodactylus cayuensis from India showing the variation in dorsal marking pattern. A WII-ADR 1219 (Balek), B WII-ADR 1681 (Ezengo), C WII-ADR 1199 (Jengging), D WII-ADR 1218 (Balek), E WII-ADR 454 (Jengging), F WII-ADR 1213 (Ramsing), G WII-ADR 698 (Potin), H WII-ADR 3017 (Glaw lake), I precloacal pores in WII-ADR 698, and J precloacal pores in WII-ADR 478. Scale bar: 10 mm. I, J not to scale." figureDoi="10.3897/vz.74.e124752.figure10" httpUri="https://binary.pensoft.net/fig/1102276">Figures 10</figureCitation>
|
||||
,
|
||||
<tableCitation id="6D24A2E86AE0C2BB86FADA220227FB43" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">S 3; Tables 2, S 4</tableCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EMZDK" type="Chresonyms">
|
||||
<paragraph id="CF2D1ACD8A411EB2D190E43B9602410A">
|
||||
<heading id="2A92FE860E2C1172119951CA0C4B6445" reason="title">Chresonyms.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="CA47BE8FC6EF9515DD067EF8372714AD">
|
||||
<taxonomicName id="B310044B4BA164D8C5AE41C35E2E3FD5" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="subSpecies" species="khasiensis" subSpecies="cayuensis">
|
||||
<emphasis id="72092101DE52188070AE7E9ADAB9905B" italics="true">Cyrtodactylus khasiensis cayuensis</emphasis>
|
||||
</taxonomicName>
|
||||
–
|
||||
<bibRefCitation id="249F5CC457CAD413AE0E30ED18C144AA" author="Li PP" firstAuthor="Li" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="733-737" refId="B22" refString="Li PP (2007) Description of a new subspecies of Cyrtodactylus khasiensis from China. Acta Zootaxonomica Sinica 32: 733–737." title="Description of a new subspecies of Cyrtodactylus khasiensis from China." volume="32" year="2007">Li (2007)</bibRefCitation>
|
||||
</paragraph>
|
||||
<paragraph id="20CA001A5EAAB3CABCFDAF5B722A59E6">
|
||||
<taxonomicName id="A3876E8EFFDDAB4EECD9EEC55EEC7309" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
|
||||
<emphasis id="1D4BC9D4D69FF84E72D38A5FEF5BFB16" italics="true">Cyrtodactylus cayuensis</emphasis>
|
||||
</taxonomicName>
|
||||
– Agarwal et al. (2018: 337)
|
||||
</paragraph>
|
||||
<paragraph id="201EED750B227F91DCED5A733F074AC2">
|
||||
<taxonomicName id="C0C0A5762A17572546C6FC46359DCCCA" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="arunachalensis">
|
||||
<emphasis id="26715360AD3BE9CD4CF4792D5E208CD4" italics="true">Cyrtodactylus arunachalensis</emphasis>
|
||||
</taxonomicName>
|
||||
–
|
||||
<bibRefCitation id="5036E5447E8BA7FE0583FF745F171D60" DOI="10.3897/evolsyst.5.61667" author="Mirza ZA & Bhosale H & Ansari F & Phansalkar P & Sawant M & Gowande G & Patel H" etAl="et al." firstAuthor="Mirza" journalOrPublisher="Evolutionary Systematics" pagination="13-23" refId="B28" refString="Mirza ZA, Bhosale H, Ansari F, Phansalkar P, Sawant M, Gowande G, Patel H (2021) A new species of geckos of the genus Cyrtodactylus Gray, 1827 from Arunachal Pradesh, India. Evolutionary Systematics 5: 13–23. https://doi.org/10.3897/evolsyst.5.61667" title="A new species of geckos of the genus Cyrtodactylus Gray, 1827 from Arunachal Pradesh, India." volume="5" year="2021">Mirza et al. (2021)</bibRefCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EC2DK" type="material">
|
||||
<paragraph id="D8BCC3A27E3FFBCF668E47B2EFD0CFE1">
|
||||
<heading id="4C3D6B323DC39CB2A574707C23F46EC7" reason="title">Materials examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="778113877C026504E58CADD12B84CDDE">
|
||||
(Fig.
|
||||
<figureCitation id="E6C15AE00D7800CAE646D20BFBA5DCE9" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyrtodactylus cayuensis from India showing the variation in dorsal marking pattern. A WII-ADR 1219 (Balek), B WII-ADR 1681 (Ezengo), C WII-ADR 1199 (Jengging), D WII-ADR 1218 (Balek), E WII-ADR 454 (Jengging), F WII-ADR 1213 (Ramsing), G WII-ADR 698 (Potin), H WII-ADR 3017 (Glaw lake), I precloacal pores in WII-ADR 698, and J precloacal pores in WII-ADR 478. Scale bar: 10 mm. I, J not to scale." figureDoi="10.3897/vz.74.e124752.figure10" httpUri="https://binary.pensoft.net/fig/1102276">10</figureCitation>
|
||||
).
|
||||
<collectingRegion id="99A66E4C22AA6BF8E1F495EBA10EFD16" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
|
||||
:
|
||||
<specimenCount id="207B96D2CFF269066E33A683CAAD9077" count="1" type="male">one male</specimenCount>
|
||||
(WII-ADR 1219) and
|
||||
<specimenCount id="0A778DBEE8C1EFB27BFF071EC46F7DBA" count="1" type="female">one female</specimenCount>
|
||||
(WII-ADR 1218) collected from Balek village (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP2DK" specific-use="{"type":"Point","coordinates":[95.272100,28.062400]}">
|
||||
<geoCoordinate id="75F2584ACA713396B799F29F9F69F5DB" degrees="28.0624" direction="north" orientation="latitude" precision="5" value="28.0624">28.0624 ° N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="BCA63C1A6B619CF6C8232BCA11A09999" degrees="95.2721" direction="east" orientation="longitude" precision="5" value="95.2721">95.2721 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
; elevation
|
||||
<quantity id="C59C3841943AC9D1DFF398EB93A9851E" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450 m</quantity>
|
||||
a. s. l.), East Siang District on
|
||||
<date id="31BBBA95F5569F63A1A04F48C439847A" value="2021-10-29">29 October 2021</date>
|
||||
by Bitupan Boruah;
|
||||
<specimenCount id="29BB93CF6B2F8B184B29200C53119BAC" count="1" type="male">one male</specimenCount>
|
||||
(WII-ADR 1213) collected from Ramsing (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX2DK" specific-use="{"type":"Point","coordinates":[94.979500,28.656300]}">
|
||||
<geoCoordinate id="F8897D5244653885B97E9D1D28ECD03E" degrees="28.6563" direction="north" orientation="latitude" precision="5" value="28.6563">28.6563 ° N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="04D1B3D647C89A53608C01DB4B714064" degrees="94.9795" direction="east" orientation="longitude" precision="5" value="94.9795">94.9795 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
; elevation
|
||||
<quantity id="E2579E4E2FCD3554D27DA29809C53FF5" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
|
||||
a. s. l.), Mouling National Park, Upper Siang District on
|
||||
<date id="E92BB40106586A5FFB897F30283C97E7" value="2021-10-27">27 October 2021</date>
|
||||
by Bitupan Boruah;
|
||||
<specimenCount id="56B7E3F556DB4553B1339FE72622FB02" count="1" type="female">one female</specimenCount>
|
||||
(WII-ADR 1199) collected from Syrnyup stream (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E62DK" specific-use="{"type":"Point","coordinates":[95.030500,28.534000]}">
|
||||
<geoCoordinate id="BE2A74361CE825ABA05338B581420200" degrees="28.5340" direction="north" orientation="latitude" precision="5" value="28.534">28.5340 ° N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="73979654104F37809B98C863027B4DCC" degrees="95.0305" direction="east" orientation="longitude" precision="5" value="95.0305">95.0305 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
; elevation
|
||||
<quantity id="329965E1309EFA79A6BE2D2C4F30306D" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</quantity>
|
||||
a. s. l.), Jengging, Mouling National Park, Upper Siang District on
|
||||
<date id="74002D5F7A7C46E2989AE8955CB16213" value="2021-10-26">26 October 2021</date>
|
||||
by Bitupan Boruah;
|
||||
<specimenCount id="EAB5A5201F674947B54FDCDEF0964A98" count="2" type="male">two males</specimenCount>
|
||||
(WII-ADR 695 and WII-ADR 696) and
|
||||
<specimenCount id="FB615ABC298C53CF136A47668ADFB78A" count="2" type="female">two females</specimenCount>
|
||||
(WII-ADR 697 and WII-ADR 698) collected from Potin (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH3DK" specific-use="{"type":"Point","coordinates":[93.849700,27.347800]}">
|
||||
<geoCoordinate id="1BCAEB6108EB205809C1C1F14E4A1871" degrees="27.3478" direction="north" orientation="latitude" precision="5" value="27.3478">27.3478 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="6104781A052483D83FE2646A0385D20F" degrees="93.8497" direction="east" orientation="longitude" precision="5" value="93.8497">93.8497 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
, elevation
|
||||
<quantity id="EDCC4D11652435D48665772D276076BE" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</quantity>
|
||||
a. s. l.), Lower Subansiri District on
|
||||
<date id="3DB9D5066C035EB365FCE3DDDAD0EF60" value="2019-10-05">5 October 2019</date>
|
||||
by Bitupan Boruah;
|
||||
<specimenCount id="CB5AF0605439ECC8A81E9883A036C211" count="1" type="female">one female</specimenCount>
|
||||
(WII-ADR 3017) collected near Glaw lake (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP3DK" specific-use="{"type":"Point","coordinates":[96.445600,27.696000]}">
|
||||
<geoCoordinate id="F5115D947AFC324DBEF425E7401A6282" degrees="27.6960" direction="north" orientation="latitude" precision="5" value="27.696">27.6960 ° N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="9A314D5B0239E88C73437EAE8BC461A0" degrees="96.4456" direction="east" orientation="longitude" precision="5" value="96.4456">96.4456 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
; elevation
|
||||
<quantity id="B883FAB4D2C42BEFB836CC65304E01F5" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
|
||||
a. s. l.), Kamlang Tiger Reserve, Lohit District on
|
||||
<date id="0DCBC3E5C73A607C47E838073B1A0244" value="2022-09-02">2 September 2022</date>
|
||||
by Abhijit Das, Bitupan Boruah and Naitik G. Patel;
|
||||
<specimenCount id="5983550108D8B5E41FDE6B9164BB818A" count="1" type="male">one male</specimenCount>
|
||||
(WII-ADR 1682) and
|
||||
<specimenCount id="1AB4A844CCC6CBF52C5565AD354F2AEE" count="1" type="female">one female</specimenCount>
|
||||
(WII-ADR 1681) collected from Ezengo (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX3DK" specific-use="{"type":"Point","coordinates":[95.863800,28.156500]}">
|
||||
<geoCoordinate id="F70328F07D5627C626CF9F702A505183" degrees="28.1565" direction="north" orientation="latitude" precision="5" value="28.1565">28.1565 ° N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="D48505D8A2B7956AAB23489B3B2D144F" degrees="95.8638" direction="east" orientation="longitude" precision="5" value="95.8638">95.8638 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
; elevation
|
||||
<quantity id="E6094C6516B8B1883D3E9FD0551BCC67" metricMagnitude="2" metricUnit="m" metricValue="5.6" unit="m" value="560.0">560 m</quantity>
|
||||
a. s. l.), Lower Dibang Valley District on
|
||||
<date id="7D91172C428B060DC773E804AB60CC11" value="2022-08-03">3 August 2022</date>
|
||||
By Bitupan Boruah;
|
||||
<specimenCount id="36E563588AA5A6D34FEA761A5BE7CC3A" count="3" type="female">three females</specimenCount>
|
||||
(WII-ADR 453, WII-ADR 454 and WII-ADR 459) collected from Jengging (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E63DK" specific-use="{"type":"Point","coordinates":[95.053700,28.549900]}">
|
||||
<geoCoordinate id="98E6CAB88A3E1B58DEE3628294964A8B" degrees="28.5499" direction="north" orientation="latitude" precision="5" value="28.5499">28.5499 ° N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="7D9F321ECC03AE9DDE319284FD073E46" degrees="95.0537" direction="east" orientation="longitude" precision="5" value="95.0537">95.0537 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
; elevation
|
||||
<quantity id="C4F79E803110B8E81A1353CFCA55D0E8" metricMagnitude="2" metricUnit="m" metricValue="7.6" unit="m" value="760.0">760 m</quantity>
|
||||
a. s. l.) by Abhijit Das on
|
||||
<date id="3B5BC1D537033D4E28C6066DB9C3DBB1" value="2018-10-05">5 October 2018</date>
|
||||
;
|
||||
<specimenCount id="A35CB19E2045DBDF80E2832C93610F27" count="1" type="female">one female</specimenCount>
|
||||
(WII-ADR 478) and
|
||||
<specimenCount id="BDF452773A67C8CCEE1F3E25336696B3" count="1" type="male">one male</specimenCount>
|
||||
(WII-ADR 473) collected from
|
||||
<quantity id="515323FACB6D73565D972F6E03CEE122" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
|
||||
northwest to Pasighat (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH4DK" specific-use="{"type":"Point","coordinates":[95.268200,28.094500]}">
|
||||
<geoCoordinate id="7E5A365D295C37F2863527C9CA23DA69" degrees="28.0945" direction="north" orientation="latitude" precision="5" value="28.0945">28.0945 ° N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="28D82508028FD3D230C39435D8479B15" degrees="95.2682" direction="east" orientation="longitude" precision="5" value="95.2682">95.2682 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
; elevation
|
||||
<quantity id="5B9B9B3D86A396CEDD0E49D29616382A" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">410 m</quantity>
|
||||
a. s. l.) by Abhijit Das on
|
||||
<date id="EE44B8C8746D60186591CDB47D6ED123" value="2018-10-02">2 October 2018</date>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="9A6CE0EB7C8B2849EA5740F64A78FD75" ID-DOI="10.3897/vz.74.e124752.figure10" ID-arpha="85FD849D-6749-530E-B888-2043F84A1248" httpUri="https://binary.pensoft.net/fig/1102276" startId="F10">
|
||||
<paragraph id="1305AA44A36028E2F2F090602D725D3F">
|
||||
<label id="549BEF6610A56DC097A34246C80A069A">Figure 10.</label>
|
||||
</paragraph>
|
||||
<paragraph id="BB98C6501DFA8FAE3B2EFCBD1E774B18">
|
||||
<taxonomicName id="B9E732165BD3395A8530084AEEB6EC99" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
|
||||
<emphasis id="2D91098550B316A036CAAE3C1662303C" italics="true">Cyrtodactylus cayuensis</emphasis>
|
||||
</taxonomicName>
|
||||
from India showing the variation in dorsal marking pattern.
|
||||
<emphasis id="9DB13E9CF5B9CE758EB75D173582D238" bold="true">A</emphasis>
|
||||
WII-ADR 1219 (Balek),
|
||||
<emphasis id="AC25C0262069E488206AEC361CCC7F6A" bold="true">B</emphasis>
|
||||
WII-ADR 1681 (Ezengo),
|
||||
<emphasis id="1735B8C9054955D98F61025001FF9814" bold="true">C</emphasis>
|
||||
WII-ADR 1199 (Jengging),
|
||||
<emphasis id="189317A6F9AA7CE8373D7D7E385DCF63" bold="true">D</emphasis>
|
||||
WII-ADR 1218 (Balek),
|
||||
<emphasis id="FA0BC65F5AC74B70D4DFC503D9016530" bold="true">E</emphasis>
|
||||
WII-ADR 454 (Jengging),
|
||||
<emphasis id="27E3EC9C73D1F0D9A00B6F220C179E30" bold="true">F</emphasis>
|
||||
WII-ADR 1213 (Ramsing),
|
||||
<emphasis id="12B99593C367D72F5554E0912229323F" bold="true">G</emphasis>
|
||||
WII-ADR 698 (Potin),
|
||||
<emphasis id="E253066F32F753AED84417B3A86F6CA7" bold="true">H</emphasis>
|
||||
WII-ADR 3017 (Glaw lake),
|
||||
<emphasis id="F067B8AE1E672EEFCE3D7F33FFECF438" bold="true">I</emphasis>
|
||||
precloacal pores in WII-ADR 698, and
|
||||
<emphasis id="ED247180329B79ECB8B428552FEDF8C5" bold="true">J</emphasis>
|
||||
precloacal pores in WII-ADR 478. Scale bar: 10 mm.
|
||||
<emphasis id="810B37CBE3573B27FA9688CB82640A3E" bold="true">I</emphasis>
|
||||
,
|
||||
<emphasis id="1BD501DA45BE36B39D9EAC222F25D318" bold="true">J</emphasis>
|
||||
not to scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E35DK" type="Revised diagnosis">
|
||||
<paragraph id="279F2D38D7AB03B5EBFD59230388F4A0">
|
||||
<heading id="F38363A7982DA57E4FB53DF577CA96E2" reason="title">Revised diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="14EAAF97811352C282538C37C27E9989">
|
||||
Medium-sized gecko,
|
||||
<abbrev id="ABBRID0EC6DK" xlink_title="snout to vent length">SVL</abbrev>
|
||||
<quantity id="5429E424F9D710D30115D3E0C71974C0" metricMagnitude="-2" metricUnit="m" metricValue="7.234999999999999" metricValueMax="8.35" metricValueMin="6.12" unit="mm" value="72.35" valueMax="83.5" valueMin="61.2">61.2–83.5 mm</quantity>
|
||||
in adult males and
|
||||
<abbrev id="ABBRID0EG6DK" xlink_title="snout to vent length">SVL</abbrev>
|
||||
<quantity id="4A83F868424A4C4599230829F2B38DE2" metricMagnitude="-2" metricUnit="m" metricValue="7.15" metricValueMax="8.36" metricValueMin="5.9399999999999995" unit="mm" value="71.5" valueMax="83.6" valueMin="59.4">59.4–83.6 mm</quantity>
|
||||
in adult females; 8–13 supralabials and 8–12 infralabials; 18–26 rows of bluntly conical and feebly keeled enlarged tubercles across mid-dorsum; 27–38 paravertebral tubercles between the level of axilla and groin; 28–44 mid-ventral scales; 13–21 subdigital lamellae beneath the fourth digit of manus; 11–23 subdigital lamellae beneath the fourth digit of pes; colouration and marking pattern variable; dorsally greyish-brown, pale-brown or pale-yellowish-brown; five to eight dark-brown bands consisting of irregular shaped and sized enlarged spots on back between axilla and groin, or dorsum with dark-brown reticulation; tail with 9–13 dark and 9–12 light bands alternatively arranged on top.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EK6DK" type="material">
|
||||
<paragraph id="1EB40EEF3C92BD6E2077AA2E5DF2D727">
|
||||
<heading id="C6284181699E7E95ABE33566BB9C0E01" reason="title">Description based on newly collected materials.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8E7C302500064FB15E38599C6CEC4A2E">
|
||||
Medium-sized gecko,
|
||||
<abbrev id="ABBRID0EQ6DK" xlink_title="snout to vent length">SVL</abbrev>
|
||||
<quantity id="D2440B1CEACA9C72A8034E5323493F82" metricMagnitude="-2" metricUnit="m" metricValue="6.815" metricValueMax="7.51" metricValueMin="6.12" unit="mm" value="68.15" valueMax="75.1" valueMin="61.2">61.2–75.1 mm</quantity>
|
||||
in males (n = 6) and
|
||||
<abbrev id="ABBRID0EU6DK" xlink_title="snout to vent length">SVL</abbrev>
|
||||
<quantity id="02518CD647FD6FC9B19551A0E35D8F94" metricMagnitude="-2" metricUnit="m" metricValue="7.15" metricValueMax="8.36" metricValueMin="5.9399999999999995" unit="mm" value="71.5" valueMax="83.6" valueMin="59.4">59.4–83.6 mm</quantity>
|
||||
in females (n = 11); head moderately large (
|
||||
<abbrev id="ABBRID0EY6DK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0E36DK" xlink_title="snout to vent length">SVL</abbrev>
|
||||
= 0.25–0.28), dorsoventrally depressed, longer than width (
|
||||
<abbrev id="ABBRID0EAAEK" xlink_title="maximum head width">HW</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EEAEK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
|
||||
= 0.67–0.75), distinct from neck, broader at occipital region; snout rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat, a longitudinal furrow on dorsal surface of the snout, snout short (
|
||||
<abbrev id="ABBRID0EIAEK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EMAEK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
|
||||
= 0.38–0.43), longer than orbit (
|
||||
<abbrev id="ABBRID0EQAEK" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EUAEK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
|
||||
= 0.49–0.78); nostril opening directed posterolaterally; ear opening oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous; supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases; rostral wide, a short groove at the middle on top; granular scales at parietal region and occipital region intermixed with slightly large rounded granular tubercles, dense in occipital and temporal region and size increases towards nape; 9–13 supralabials present, size decreases towards angle of jaw; a series of scales, slightly larger than the loreal scales present above the supralabials; 8–12 infralabials, size decreases towards angle of jaw; two or three rows of slightly enlarged and narrow scales along the infralabials, posteriorly size of those decreases; rest of the gular scales are small, granular juxtaposed, nearly homogeneous, scales on throat are imbricate.
|
||||
</paragraph>
|
||||
<paragraph id="5EC4FE9B55FCB690B1E39E6948A35B2F">
|
||||
Habitus slender (
|
||||
<abbrev id="ABBRID0E1AEK" xlink_title="maximum body width">BW</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0E5AEK" xlink_title="snout to vent length">SVL</abbrev>
|
||||
= 0.16–0.22,
|
||||
<abbrev id="ABBRID0ECBEK" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EGBEK" xlink_title="snout to vent length">SVL</abbrev>
|
||||
= 0.41–0.47), dorsoventrally depressed; dorsal scales granular, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, continuing up to third or fifth segment of the tail, size of the tubercles increases towards posterior body, pronounced at sacrum and base of the tail, posteriormost rows indistinctly keeled; 15–21 rows of dorsal tubercles across mid dorsum; 27–38 paravertebral tubercles; ventrolateral fold present; ventral scales on chest and belly larger than those of dorsal, flat, smooth, cycloid, subimbricate to imbricate; 34–44 mid ventral scales; seven or nine distinct precloacal pores in male and 7–10 small precloacal pores in female;
|
||||
<abbrev id="ABBRID0EKBEK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
|
||||
followed by three to six unpored enlarged scales below.
|
||||
</paragraph>
|
||||
<paragraph id="FBA8A732B126673E687C88DCC784E8EF">
|
||||
Forelimbs and hindlimbs slender (
|
||||
<abbrev id="ABBRID0EQBEK" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EUBEK" xlink_title="snout to vent length">SVL</abbrev>
|
||||
= 0.13–0.15,
|
||||
<abbrev id="ABBRID0EYBEK" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0E3BEK" xlink_title="snout to vent length">SVL</abbrev>
|
||||
= 0.16–0.18); digits strongly inflected at the joints, all bearing large recurved claw and enlarged subdigital lamellae, dorsal scales on forelimbs heterogeneous, granular juxtaposed, smooth and subimbricate at distal end of forearm; forearm intermixed with a few rounded, large tubercles; dorsal scales of hindlimbs granular intermixed with large, rounded, bluntly conical tubercles; thigh scales towards lateral and dorsolateral side are smooth and subimbricate; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous, granular juxtaposed; scales on ventral side of hindlimbs slightly smaller than or nearly equal to those of belly, smooth, cycloid and subimbricate; scales on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
|
||||
</paragraph>
|
||||
<paragraph id="DAC23334F71077903DA787AFA3CF2B96">
|
||||
Tail slender (
|
||||
<abbrev id="ABBRID0ECCEK" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
|
||||
=
|
||||
<quantity id="8DDDC19420FDD395A98C56CA4A87ED6C" metricMagnitude="-2" metricUnit="m" metricValue="7.75" metricValueMax="9.7" metricValueMin="5.8" unit="mm" value="77.5" valueMax="97.0" valueMin="58.0">58–97 mm</quantity>
|
||||
), gradually tapering, segments indistinct; dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size; large feebly keeled scales up to third to fifth segment of the tail, those on basal segment are pronounced and posterior rows with indistinct keel; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; two to four bluntly conical spurs present at the base of the tail. Detailed morphological characteristics of the collected samples are provided in Table S 4.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EGCEK" type="Sequence divergence">
|
||||
<paragraph id="DA9A72FEE8B6FD2E213E3578AA3A395D">
|
||||
<heading id="2DE3283C476D34AD3E0B9365D77303C8" reason="title">Sequence divergence.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="9016B803E4DD2F3E4737426025F8AEDA">
|
||||
The intraspecific divergence within
|
||||
<taxonomicName id="1A740EADDDE34B2A45EA2685424B5BD1" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
|
||||
<emphasis id="CCD6C08266E14897DE53936551A9386D" italics="true">C. cayuensis</emphasis>
|
||||
</taxonomicName>
|
||||
ranges between 0.1–6.4 %. Among the two clades identified by
|
||||
<abbrev id="ABBRID0EXCEK" xlink_title="Bayesian Poisson Tree Processes">bPTP</abbrev>
|
||||
, both clades have an intraspecific divergence between 0.1–4.3 % and 0.2–2.4 %, respectively. The divergence between these two clades range between 3–6.4 %.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E2CEK" type="Colouration in life">
|
||||
<paragraph id="B503F7703BE9108064D81DB916B75B74">
|
||||
<heading id="ECD89F7288261A73881B22060188292D" reason="title">Colouration in life.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="943CF06A3C03576575EA9878FB86BC20">Dorsal colour and marking pattern variable (Fig. S 3). Head and body greyish-brown, pale-brown to pale-yellowish-brown on top; head on top irregularly mottled with dark-brown, slightly enlarged dark-brown irregular patches may be present on occipital area; dark-brown postorbital stripe and loreal stripe present; a light stripe above loreal stripe and postorbital stripe may be visible; a few pale-yellow spots may be present on temporal and occipital region; supraciliary yellowish-brown; neck with short dark-brown streak with irregular edge or with irregular shaped patches; trunk dorsally with five to eight dark-brown bands consisting of irregular shaped and sized enlarged spots and posteriorly pale-yellowish or light-edged, these bands mid dorsally interrupted, or dorsum with dark-brown reticulation; limbs dorsally greyish-brown or pale-yellowish-brown or pale-brown with distinct or indistinct dark-brown reticulation; a dark-brown band on sacrum, band mid dorsally intersected; tail dorsally with 9–13 dark and 9–12 light bands alternatively arranged, bands are with irregular edged, anterior one or two dark bands may be mid dorsally intersected, dark bands are broad and light bands are comparatively narrow, dark bands are posteriorly becomes black or more dark-brown, while light bands become whitish; light bands may have a few dark-brown spots.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EADEK" type="distribution">
|
||||
<paragraph id="C7C5E01291724B4421EF6507310D9001">
|
||||
<heading id="99BCA824112410B94FBB1510B8F0CB21" reason="title">Distribution and natural history.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="DE222B6093D082C936AFA56C8F3EF1C1">
|
||||
The
|
||||
<typeStatus id="56046C7452172E48D70B41084D1869C4">type</typeStatus>
|
||||
locality,
|
||||
<collectingRegion id="CCF01467387DFDCD2590598B4DA75269" country="China" name="Xizang">Xizang</collectingRegion>
|
||||
(
|
||||
<collectingRegion id="1640CA3865AE026CCA9C597449F4E094" country="China" name="Xizang">Tibet</collectingRegion>
|
||||
),
|
||||
<collectingCountry id="8050427475CEFE288A1D97D1C6BA890B" name="China">China</collectingCountry>
|
||||
by
|
||||
<taxonomicName id="23CBA55514CAAB750C404CA1A998EF80" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
|
||||
<emphasis id="24A584D64025CCE57CD9D1141E16793A" italics="true">C. cayuensis</emphasis>
|
||||
</taxonomicName>
|
||||
also appears to be widely distributed in
|
||||
<collectingRegion id="08238081176E002A0DBBC4B2A555A30B" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="3FB691275992F34A6FD5C90062EE3DA0" name="India">India</collectingCountry>
|
||||
(Fig.
|
||||
<figureCitation id="F1F9916961C1031B15599488433F545C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 B</figureCitation>
|
||||
), within an elevational range of
|
||||
<quantity id="311F248A55EFEF908B0F988579416614" metricMagnitude="2" metricUnit="m" metricValue="8.05" metricValueMax="12.0" metricValueMin="4.1" unit="m" value="805.0" valueMax="1200.0" valueMin="410.0">410–1200 m</quantity>
|
||||
a. s. l.
|
||||
<taxonomicName id="21F95172509E7678550E270A19BF4A61" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
|
||||
<emphasis id="2635E1D087874EAD98888F865ECF7601" italics="true">Cyrtodactylus cayuensis</emphasis>
|
||||
</taxonomicName>
|
||||
is widespread in the foothills of
|
||||
<collectingRegion id="1042A0AE8FD505A6BE5D1DECA5C17D17" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
|
||||
from the western part of the state, at Seijusa (Pakke Tiger Reserve) to the eastern part (Kamlang Tiger Reserve) and it has successfully spread across major tributaries of the Brahmaputra River, i. e., the Subansiri, Siang, Dibang, Lohit up to at least north of Noa-Dihing. Populations of
|
||||
<taxonomicName id="FEA8D256B6D9349F7821AA5E7C6B6957" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
|
||||
<emphasis id="54A2B80E2B18BC5089AAC0148D4A0312" italics="true">C. cayuensis</emphasis>
|
||||
</taxonomicName>
|
||||
have been recorded in degraded evergreen forest as well as in undisturbed semi-evergreen and evergreen forests.
|
||||
<taxonomicName id="8F899589036A29CFF3BB1F7E4839F34A" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
|
||||
<emphasis id="9F2F435888C14922D82894E7EAC63FD8" italics="true">Cyrtodactylus cayuensis</emphasis>
|
||||
</taxonomicName>
|
||||
is particularly abundant in the vegetation close to the picturesque Glaw Lake in the Kamlang Tiger Reserve. We observed individuals at night on shrubs and bedrocks (sedimentary rock) along small forest streams. We also encountered them on small trees up to a 2.0 m inside the forest at Jengging, Kamlang Tiger Reserve and Mehao Wildlife Sanctuary.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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