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<mods:title id="21C5325A113A249DC3F02324274DF5EB">Revision of the bathydemersal fish genus Pycnocraspedum (Ophidiidae; Teleostei), with description of two new species</mods:title>
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<taxonomicName id="4C754D449002FFDCFF77FE968BD2A9B2" ID-CoL="4QM82" authority="(Parr, 1933)" baseAuthorityName="Parr" baseAuthorityYear="1933" box="[151,660,366,393]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="19" pageNumber="462" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD59002FFDCFF77FE968BD2A9B2" bold="true" box="[151,660,366,393]" pageId="19" pageNumber="462">
<emphasis id="B901EAD59002FFDCFF77FE9688BEA9B3" bold="true" box="[151,504,366,393]" italics="true" pageId="19" pageNumber="462">Pycnocraspedum phyllosoma</emphasis>
(
<bibRefCitation id="EFE44B369002FFDCFDE7FE968BCAA9B2" author="Parr, A. E." box="[519,652,366,392]" pageId="19" pageNumber="462" pagination="1 - 51" refId="ref19495" refString="Parr, A. E. (1933) Deepsea Berycomorphi and Percomorphi from the waters around the Bahama and Bermuda Islands. Bulletin Bingham Oceanographic Collection, 3 (6), 1-51." type="journal article" year="1933">Parr, 1933</bibRefCitation>
)
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<subSubSection id="C36F654C9002FFDCFF77FE6B8B64A997" box="[151,546,402,429]" pageId="19" pageNumber="462" type="description">
<paragraph id="8BCA36C79002FFDCFF77FE6B8B64A997" blockId="19.[151,660,366,429]" box="[151,546,402,429]" pageId="19" pageNumber="462">
<figureCitation id="134E2A429002FFDCFF77FE6B8846A997" box="[151,256,403,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1527,1551]" captionTargetBox="[168,1411,202,1477]" captionTargetId="figure-17@3.[151,1436,181,1504]" captionTargetPageId="3" captionText="FIGURE 1. Radiograph figures of selected specimens of the genus Pycnocraspedum to visualize position of dorsal-fin rays in relation to first vertebra; “1” denotes the first dorsal-fin ray above the first vertebra; numbers with negative denotation represent dorsal-fin rays in front of the first vertebra. 1A) Pycnocraspedum armatum, USNM 227390; 1B) Pycnocraspedum phyllosoma USNM 227413; 1C) Pycnocraspedum squamipinne, ZMUC P771871; 1D) Pycnocraspedum fulvum (holotype), BSKU 27497." figureDoi="http://doi.org/10.5281/zenodo.15561735" httpUri="https://zenodo.org/record/15561735/files/figure.png" pageId="19" pageNumber="462">Figures 1</figureCitation>
,
<figureCitation id="134E2A429002FFDCFEEEFE6B885BA997" box="[270,285,403,429]" captionStart="FIGURE 3" captionStartId="11.[151,250,1786,1810]" captionTargetBox="[256,1328,181,1748]" captionTargetId="figure-17@11.[256,1331,181,1762]" captionTargetPageId="11" captionText="FIGURE 3. Detail figures and drawings of preopercular spines and lower preopercular pores in Pycnocraspedum; preopercular spines are numbered from 1 through 4 from the lower angle of the preoperculum upwards; abbreviation lpp = lower preopercular pore. 3A3B) Pycnocraspedum armatum, USNM 227412, 3A1-3A2 specimen 1 left side, 3A3 specimen 1 right side (mirror imaged), 3B specimen 2; 3C3D) Pycnocraspedum phyllosoma, 3C USNM 421587, 3D USNM 421586; 3E) Pycnocraspedum fulvum (holotype), BSKU 27497; 3F3G) Pycnocraspedum africanum sp. nov., 3F holotype, MNHN 2000-5691, 3G paratype, MNHN 2000-5692; 3H) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021; 3I3K) Pycnocraspedum squamipinne, 3I lectotype, BMNH 1890.7.31.1, 3J ZMUC P771871, 3K ZMUC P771737." figureDoi="http://doi.org/10.5281/zenodo.15561739" httpUri="https://zenodo.org/record/15561739/files/figure.png" pageId="19" pageNumber="462">3</figureCitation>
,
<figureCitation id="134E2A429002FFDCFECAFE6B887CA997" box="[298,314,403,429]" captionStart="FIGURE 4" captionStartId="13.[151,250,1811,1835]" captionTargetBox="[151,1436,352,1787]" captionTargetId="figure-76@13.[151,1436,345,1788]" captionTargetPageId="13" captionText="FIGURE 4. Head and body squamation details in Pycnocraspedum phyllosoma, USNM 421587; abbreviations used: aip-1aip- 3 = anterior infraorbital pores 13, asp = anterior supraorbital pore, pip-1 = posterior infraorbital pore 1; preopercular spines numbered. 4A) head squamation in lateral view; 4B) trunk squamation and every 10 scale row count marked by white dot; 4C) snout squamation and pores detail; 4D) head squamation in ventral view." figureDoi="http://doi.org/10.5281/zenodo.15561741" httpUri="https://zenodo.org/record/15561741/files/figure.png" pageId="19" pageNumber="462">4</figureCitation>
,
<figureCitation id="134E2A429002FFDCFEA6FE6A8810A997" box="[326,342,402,429]" captionStart="FIGURE 5" captionStartId="14.[151,250,1750,1774]" captionTargetBox="[245,1358,193,1715]" captionTargetId="figure-23@14.[222,1366,181,1726]" captionTargetPageId="14" captionText="FIGURE 5. Otoliths, inner faces (if not noted otherwise) shown from right side. 5A5C) Pycnocraspedum africanum sp. nov., 5A holotype, MNHN 2000-5691 (5A2 dorsal view), 5B paratype MNHN 2000-5692, 5C tentatively assigned specimen, IRSNB (5C2 dorsal view, 5C3 anterior view); 5D5E) Pycnocraspedum armatum, 5D USNM 227412 (5D2 ventral view), 5E USNM 227390 (5E2 ventral view); 5F) Pycnocraspedum fulvum (holotype), BSKU 27497 (5F2 ventral view, 5F3 anterior view); 5G) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021 (5G2 ventral view); 5H) Pycnocraspedum phyllosoma, USNM 421587 (5H2 ventral view, 5H3 anterior view); 5I5K) Pycnocraspedum squamipinne, 5I lectotype, BMNH 1890.7.31.1 (5I2 ventral view), 5J ZMUC P771871, 5L ZMUC P771739 (5L2 ventral view), 5K ZMUC P771738 (5K2 ventral view); 5M) †Pycnocraspedum cetonaense (holotype) SMF P.5694 (5M2 dorsal view, 5M3 anterior view)." figureDoi="http://doi.org/10.5281/zenodo.15561743" httpUri="https://zenodo.org/record/15561743/files/figure.png" pageId="19" pageNumber="462">5</figureCitation>
,
<figureCitation id="134E2A429002FFDCFE82FE6B8834A997" box="[354,370,403,429]" captionStart="FIGURE 9" captionStartId="20.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-23@20.[151,1435,181,1785]" captionTargetPageId="20" captionText="FIGURE 9. Pycnocraspedum phyllosoma (Parr, 1933); 9A, 9C) USNM 412587, 295 mm SL, 12.0832°N 68.8991°W; 9B) radiograph of USNM 227413, 230 mm SL, 20.85°N 71.52°, 512576 m." figureDoi="http://doi.org/10.5281/zenodo.15561751" httpUri="https://zenodo.org/record/15561751/files/figure.png" pageId="19" pageNumber="462">9</figureCitation>
,
<figureCitation id="134E2A429002FFDCFE9EFE6B88DBA997" box="[382,413,403,429]" captionStart="FIGURE 12" captionStartId="24.[151,250,899,923]" captionTargetBox="[153,1433,186,871]" captionTargetId="figure-23@24.[151,1436,181,875]" captionTargetPageId="24" captionText="FIGURE 12. Geographic distribution of the species of Pycnocraspedum. One symbol can represent more than one location." figureDoi="http://doi.org/10.5281/zenodo.15561755" httpUri="https://zenodo.org/record/15561755/files/figure.png" pageId="19" pageNumber="462">12</figureCitation>
;
<tableCitation id="C6F7037C9002FFDCFE48FE6B8B40A997" box="[424,518,403,429]" captionStart="TABLE 1" captionStartId="2.[152,244,151,175]" captionTargetBox="[163,1411,227,1782]" captionTargetId="graphics-48@2.[151,1437,256,1787]" captionText="TABLE 1. Selected keyparameters ofthecurrently 37recognizedophidiidgenera.Characterssharedwith Pycnocraspedum are highlighted. D/V = first dorsal-fin ray above vertebra." httpUri="http://table.plazi.org/id/DF0A664F9013FFCDFF78FF6F8BBBA8E9" pageId="19" pageNumber="462" tableUuid="DF0A664F9013FFCDFF78FF6F8BBBA8E9">Tables 1</tableCitation>
,
<tableCitation id="C6F7037C9002FFDCFDF4FE6B8B64A997" box="[532,546,403,429]" captionStart="TABLE 2" captionStartId="8.[152,244,720,744]" captionTargetPageId="8" captionText="TABLE 2. Meristic, morphometric and otolith characters of the species of the genus Pycnocraspedum. For abbreviations" httpUri="http://table.plazi.org/id/DF0A664F9019FFC7FF78FD288B4AAB36" pageId="19" pageNumber="462" tableUuid="DF0A664F9019FFC7FF78FD288B4AAB36">2</tableCitation>
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<taxonomicName id="4C754D449002FFDCFF77FE2F8B0EA9D5" ID-CoL="748GJ" authority="Parr, 1933: 44 - 46" authorityName="Parr" authorityPageNumber="44 - 46" authorityYear="1933" box="[151,584,471,495]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="19" pageNumber="462" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD59002FFDCFF77FE2F88CCA9D5" box="[151,394,471,495]" italics="true" pageId="19" pageNumber="462">Neobythites phyllosoma</emphasis>
<bibRefCitation id="EFE44B369002FFDCFE70FE208B0EA9D5" author="Parr, A. E." box="[400,584,471,495]" pageId="19" pageNumber="462" pagination="1 - 51" refId="ref19495" refString="Parr, A. E. (1933) Deepsea Berycomorphi and Percomorphi from the waters around the Bahama and Bermuda Islands. Bulletin Bingham Oceanographic Collection, 3 (6), 1-51." type="journal article" year="1933">Parr, 1933: 4446</bibRefCitation>
</taxonomicName>
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, fig. 20, off
<collectingCountry id="F36276579002FFDCFD24FE2F8A02A9D5" box="[708,836,471,495]" name="Turks Island" pageId="19" pageNumber="462">Turks Island</collectingCountry>
,
<collectingCountry id="F36276579002FFDCFCAFFE2F8AE8A9D5" box="[847,942,471,495]" name="Bahamas" pageId="19" pageNumber="462">Bahamas</collectingCountry>
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,
<geoCoordinate id="EE4150009002FFDCFBF9FE2F8D37A9D5" box="[1049,1137,471,495]" degrees="71" direction="west" minutes="11" orientation="longitude" pageId="19" pageNumber="462" precision="925" value="-71.183334">71°11'W</geoCoordinate>
, Pawnee St. 52.
<taxonomicName id="4C754D449002FFDCFF77FE0E8B02AA34" ID-CoL="4QM82" authority="(Parr, 1933)" baseAuthorityName="Parr" baseAuthorityYear="1933" box="[151,580,502,526]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="19" pageNumber="462" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD59002FFDCFF77FE0E8887AA34" box="[151,449,502,526]" italics="true" pageId="19" pageNumber="462">Pycnocraspedum phyllosoma</emphasis>
(
<bibRefCitation id="EFE44B369002FFDCFE2EFE0E8B7AAA34" author="Parr, A. E." box="[462,572,502,526]" pageId="19" pageNumber="462" pagination="1 - 51" refId="ref19495" refString="Parr, A. E. (1933) Deepsea Berycomorphi and Percomorphi from the waters around the Bahama and Bermuda Islands. Bulletin Bingham Oceanographic Collection, 3 (6), 1-51." type="journal article" year="1933">Parr, 1933</bibRefCitation>
)
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:
<bibRefCitation id="EFE44B369002FFDCFDAEFE0E8A67AA34" author="Nielsen, J. G. &amp; Cohen, D. M. &amp; Markle, D. F. &amp; Robins, C. R." box="[590,801,502,526]" pageId="19" pageNumber="462" refId="ref19264" refString="Nielsen, J. G., Cohen, D. M., Markle, D. F. &amp; Robins, C. R. (1999) An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiiform fishes known to date. FAO Fisheries Synopsis No. 125. Vol. 18. FAO, Rome, 178 pp." type="book" year="1999">
Nielsen
<emphasis id="B901EAD59002FFDCFD42FE0F8B93AA34" box="[674,725,502,526]" italics="true" pageId="19" pageNumber="462">et al.</emphasis>
(1999)
</bibRefCitation>
, Evseenko &amp; Okiyama (2006),
<bibRefCitation id="EFE44B369002FFDCFB81FE0E8C54AA34" author="Prokofiev, A. M." box="[1121,1298,502,526]" pageId="19" pageNumber="462" pagination="1019 - 1024" refId="ref19635" refString="Prokofiev, A. M. (2022) New data on the morphology and distribution of two rare species of Ophidiiformes: Diplacanthopoma japonicum (Bythitidae) and Pycnocraspedum microlepis (Neobythitidae). Voprosy Ikhtiologii, 62 (6), 680 - 685. [in Russian, English translation appeared in Journal of Ichthyology, 62 (6), 1019-1024] https://doi.org/10.1134/S0032945222060224" type="journal article" year="2022">Prokofiev (2022)</bibRefCitation>
.
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<emphasis id="B901EAD59002FFDCFF77FDC4883CAA6C" bold="true" box="[151,378,572,598]" pageId="19" pageNumber="462">Material examined</emphasis>
(
<specimenCount id="9D73FD4E9002FFDCFE6AFDC48B5FAA6C" box="[394,537,572,598]" count="4" pageId="19" pageNumber="462" type="generic">4 specimens</specimenCount>
, 130355 mm SL):
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<specimenCount id="9D73FD4E9002FFDCFD1BFDC48A3DAA6C" box="[763,891,572,598]" count="1" pageId="19" pageNumber="462" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE029002FFDCFC62FDC48A95AA6C" box="[898,979,572,598]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="462" type="Museum">USNM</collectionCode>
227413, 230 mm SL,
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(off
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),
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<location id="8EAA601C9002FFDCFE42FD9888B0AA40" LSID="urn:lsid:plazi:treatment:03DC87D19002FFDCFF77FE968A1FAFCD:8EAA601C9002FFDCFE42FD9888B0AA40" box="[418,502,608,634]" country="Caicos Island" latitude="20.85" longLatPrecision="760" longitude="-71.52" name="Oregon" pageId="19" pageNumber="462">Oregon</location>
cr. 101, st. 5427, 40 ft shrimp trawl, 512576 m,
<date id="FFCB10079002FFDCFBC1FD988DFCAA41" box="[1057,1210,608,635]" pageId="19" pageNumber="462" value="1965-05-27">
<collectingDate id="EF8FE9EF9002FFDCFBC1FD988DFCAA41" box="[1057,1210,608,635]" pageId="19" pageNumber="462" value="1965-05-27">27 May 1965</collectingDate>
</date>
</materialsCitation>
;
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<collectionCode id="ED64AE029002FFDCFAABFD988CDAAA40" box="[1355,1436,608,634]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="462" type="Museum">USNM</collectionCode>
421586, 355 mm SL,
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(off
<collectingCountry id="F36276579002FFDCFD27FD7C8A6FAAA4" box="[711,809,644,670]" name="Curaçao" pageId="19" pageNumber="462">Curacao</collectingCountry>
),
<collectionCode id="ED64AE029002FFDCFCDBFD7D8A19AAA5" box="[827,863,645,671]" country="Italy" name="Collection of Leptospira Strains" pageId="19" pageNumber="462">RV</collectionCode>
<location id="8EAA601C9002FFDCFC87FD7C8A81AAA4" LSID="urn:lsid:plazi:treatment:03DC87D19002FFDCFF77FE968A1FAFCD:8EAA601C9002FFDCFC87FD7C8A81AAA4" box="[871,967,644,670]" country="Curaçao" latitude="12.0832" longLatPrecision="7" longitude="-68.8991" municipality="Deep Reef Observation Project" name="Curasub" pageId="19" pageNumber="462">Curasub</location>
,
<collectingMunicipality id="6BAEACBD9002FFDCFC33FD7D8C76AAA4" box="[979,1328,644,671]" pageId="19" pageNumber="462">Deep Reef Observation Project</collectingMunicipality>
(
<collectionCode id="ED64AE029002FFDCFAA0FD7C8CD6AAA4" box="[1344,1424,644,670]" pageId="19" pageNumber="462">DROP</collectionCode>
),
<date id="FFCB10079002FFDCFF77FD51884AAAF8" box="[151,268,680,707]" pageId="19" pageNumber="462" value="2013-05">
<collectingDate id="EF8FE9EF9002FFDCFF77FD51884AAAF8" box="[151,268,680,707]" pageId="19" pageNumber="462" value="2013-05">May 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A9002FFDCFEF7FD508B1AAADC" collectingDate="2013-05" collectionCode="USNM, RV, DROP" country="Curaçao" latitude="12.0832" location="Curasub" longLatPrecision="7" longitude="-68.8991" municipality="Deep Reef Observation Project" pageId="19" pageNumber="462" specimenCount="1">
<specimenCount id="9D73FD4E9002FFDCFEF7FD5088D0AAF8" box="[279,406,680,706]" count="1" pageId="19" pageNumber="462" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE029002FFDCFE7CFD5088ABAAF8" box="[412,493,680,706]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="462" type="Museum">USNM</collectionCode>
412587, 295 mm SL,
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(off
<collectingCountry id="F36276579002FFDCFBFCFD508D38AAF8" box="[1052,1150,680,706]" name="Curaçao" pageId="19" pageNumber="462">Curacao</collectingCountry>
),
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<location id="8EAA601C9002FFDCFB5BFD508C5DAAF8" LSID="urn:lsid:plazi:treatment:03DC87D19002FFDCFF77FE968A1FAFCD:8EAA601C9002FFDCFB5BFD508C5DAAF8" box="[1211,1307,680,706]" country="Curaçao" latitude="12.0832" longLatPrecision="7" longitude="-68.8991" municipality="Deep Reef Observation Project" name="Curasub" pageId="19" pageNumber="462">Curasub</location>
,
<collectingMunicipality id="6BAEACBD9002FFDCFAC6FD518830AADC" pageId="19" pageNumber="462">Deep Reef Observation Project</collectingMunicipality>
(
<collectionCode id="ED64AE029002FFDCFE64FD348892AADC" box="[388,468,716,742]" pageId="19" pageNumber="462">DROP</collectionCode>
),
<date id="FFCB10079002FFDCFE07FD358B1AAADC" box="[487,604,716,743]" pageId="19" pageNumber="462" value="2013-05">
<collectingDate id="EF8FE9EF9002FFDCFE07FD358B1AAADC" box="[487,604,716,743]" pageId="19" pageNumber="462" value="2013-05">May 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A9002FFDCFD88FD348A0CAB30" collectingDate="1967-09-29" collectionCode="USNM, RV" country="Curaçao" elevation="448" latitude="25.97" location="Gerda" longLatPrecision="746" longitude="-78.48" municipality="Otter" pageId="19" pageNumber="462" specimenCount="1">
<specimenCount id="9D73FD4E9002FFDCFD88FD348BA0AADC" box="[616,742,716,742]" count="1" pageId="19" pageNumber="462" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE029002FFDCFD0DFD348A78AADC" box="[749,830,716,742]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="462" type="Museum">USNM</collectionCode>
470952, 130 mm SL,
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<geoCoordinate id="EE4150009002FFDCFB7CFD348C47AADC" box="[1180,1281,716,742]" degrees="78.48" direction="west" orientation="longitude" pageId="19" pageNumber="462" precision="555" value="-78.48">78.48°W</geoCoordinate>
,
<collectionCode id="ED64AE029002FFDCFAEBFD358C69AADD" box="[1291,1327,717,743]" country="Italy" name="Collection of Leptospira Strains" pageId="19" pageNumber="462">RV</collectionCode>
<location id="8EAA601C9002FFDCFAD6FD348C3CAADC" LSID="urn:lsid:plazi:treatment:03DC87D19002FFDCFF77FE968A1FAFCD:8EAA601C9002FFDCFAD6FD348C3CAADC" box="[1334,1402,716,742]" country="Curaçao" latitude="25.97" longLatPrecision="746" longitude="-78.48" municipality="Otter" name="Gerda" pageId="19" pageNumber="462">Gerda</location>
cr. 6720, st. 925, 10
<collectingMunicipality id="6BAEACBD9002FFDCFEB9FD0888D5AB30" box="[345,403,752,778]" pageId="19" pageNumber="462">Otter</collectingMunicipality>
trawl, 439457 m,
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<collectingDate id="EF8FE9EF9002FFDCFD8EFD088A0CAB30" box="[622,842,752,778]" pageId="19" pageNumber="462" value="1967-09-29">29 September 1967</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9002FFDCFF27FCEC8A5AABD8" pageId="19" pageNumber="462" type="diagnosis">
<paragraph id="8BCA36C79002FFDCFF27FCEC8A5AABD8" blockId="19.[151,1437,572,2039]" pageId="19" pageNumber="462">
<emphasis id="B901EAD59002FFDCFF27FCEC8804AB14" bold="true" box="[199,322,788,814]" pageId="19" pageNumber="462">Diagnosis.</emphasis>
Precaudal vertebrae 1213, total vertebrae 5254; dorsal-fin rays 96101; anal-fin rays 7073; pectoral-fin rays 2527; long gill rakers on first gill arch 45; pseudobranchial filaments 56; 34 dorsal-fin rays in front of first vertebra; HL 22.726.5% of SL; body depth at vent 2123% of SL; predorsal length 18.823% of SL; rear margin of preopercle with 3 spines, 2 lower and 1 upper, with flashy flap between spines 2 and 3; transversal scale rows above origin of anal fin 7278, and above lateral line 2021; otolith (
<quantity id="4C8D9B229002FFDCFBF4FC5C8D3FAB85" box="[1044,1145,932,959]" metricMagnitude="-2" metricUnit="m" metricValue="2.33" pageId="19" pageNumber="462" unit="mm" value="23.3">23.3 mm</quantity>
in length) OL:OH = 2.15, dorsal rim smooth, ventral rim symmetrical to dorsal rim.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9002FFDCFF27FC148DFBAEEC" pageId="19" pageNumber="462" type="description">
<paragraph id="8BCA36C79002FFDCFF27FC148878AC74" blockId="19.[151,1437,572,2039]" pageId="19" pageNumber="462">
<emphasis id="B901EAD59002FFDCFF27FC14881FAC3C" bold="true" box="[199,345,1004,1030]" pageId="19" pageNumber="462">Description.</emphasis>
Relatively robust fish with stout tapering tail and rounded snout.Maximal size of fishes investigated
<quantity id="4C8D9B229002FFDCFF77FBE889B5AC11" box="[151,243,1040,1067]" metricMagnitude="-1" metricUnit="m" metricValue="3.55" pageId="19" pageNumber="462" unit="mm" value="355.0">355 mm</quantity>
SL. Preopercular rim with 3 spines, 2 spines positioned at bend and 1 further up, with fleshy flap in between spines 2 and 3.
</paragraph>
<paragraph id="8BCA36C79002FFDCFF27FBA08DEAACE4" blockId="19.[151,1437,572,2039]" pageId="19" pageNumber="462">
Meristics (
<specimenCount id="9D73FD4E9002FFDCFEA0FBA08889AC48" box="[320,463,1112,1138]" count="4" pageId="19" pageNumber="462" type="generic">4 specimens</specimenCount>
, not including juvenile
<typeStatus id="54CE88659002FFDCFD3BFBA08A7BAC48" box="[731,829,1112,1138]" pageId="19" pageNumber="462" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C754D449002FFDCFC85FBA18D45AC48" baseAuthorityName="Parr" baseAuthorityYear="1933" box="[869,1027,1112,1138]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="19" pageNumber="462" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD59002FFDCFC85FBA18D45AC48" box="[869,1027,1112,1138]" italics="true" pageId="19" pageNumber="462">P. phyllosoma</emphasis>
</taxonomicName>
): precaudal vertebrae (1213); total vertebrae 5254; dorsal-fin rays 96101; anal-fin rays 7073; pectoral-fin rays 2527; caudal-fin rays 10; D/V = -3 -4; D/A = 2930; V/A = 1617; long gill rakers on first gill arch 45; pseudobranchial filaments 56; transversal scale rows above origin of anal fin 7278, and above lateral line 2021; pyloric coeca 1821.
</paragraph>
<paragraph id="8BCA36C79002FFDCFF27FB1089FCAD54" blockId="19.[151,1437,572,2039]" pageId="19" pageNumber="462">
Morphometrics (
<specimenCount id="9D73FD4E9002FFDCFE64FB108B57AD38" box="[388,529,1256,1282]" count="4" pageId="19" pageNumber="462" type="generic">4 specimens</specimenCount>
) in % of SL: HL 22.726.5; body depth at vent 2123; diameter of eye 4.45.0; upper jaw length 11.414.5; interorbital width 6.58.1; pelvic-fin base to anal-fin base 22.226; predorsal length 18.823; preanal length 41.751; prepelvic length 19.127; pectoral-fin length 14.517.2; pelvic-fin length 11.5 13.
</paragraph>
<paragraph id="8BCA36C79002FFDCFF27FA80882BAEB4" blockId="19.[151,1437,572,2039]" pageId="19" pageNumber="462">
Otolith morphology (
<specimenCount id="9D73FD4E9002FFDCFE59FA808B70ADA8" box="[441,566,1400,1426]" count="1" pageId="19" pageNumber="462" type="generic">1 specimen</specimenCount>
from specimen of
<quantity id="4C8D9B229002FFDCFCE9FA808A20ADA9" box="[777,870,1400,1427]" metricMagnitude="-1" metricUnit="m" metricValue="2.95" pageId="19" pageNumber="462" unit="mm" value="295.0">295 mm</quantity>
SL). Size
<quantity id="4C8D9B229002FFDCFC3CFA808D06ADA9" box="[988,1088,1400,1427]" metricMagnitude="-2" metricUnit="m" metricValue="2.33" pageId="19" pageNumber="462" unit="mm" value="23.3">23.3 mm</quantity>
in length; OL:OH = 2.15; OH: OT = 2.4. Moderately thin, oval, elongated, anteriorly moderately rounded, posteriorly distinctly pointed, anterior and posterior tips positioned along median sulcus axis. Dorsal rim regularly curved, relatively high, smooth; ventral rim about as deep as dorsal rim high, smooth. Inner face distinctly bent/convex in horizontal and vertical directions, with long, undivided sulcus reaching relatively close to anterior and posterior rims of otolith. Sulcus with shallow, undivided, uniform colliculum with slightly concave dorsal rim and slightly convex ventral rim and rounded anterior and posterior tips. OL:CL = 1.25. Dorsal field smooth, without depression; ventral field smooth. Outer face nearly flat, rather smooth.
</paragraph>
<paragraph id="8BCA36C79002FFDCFF27F9608DFBAEEC" blockId="19.[151,1437,572,2039]" pageId="19" pageNumber="462">
Coloration. Live coloration pale with small dark spots in small individuals (
<bibRefCitation id="EFE44B369002FFDCFBA8F9608C65AE88" author="Girard, M. G. &amp; Mundy, B. C. &amp; Nonaka, A. &amp; Johnson, G. D." box="[1096,1315,1688,1714]" pageId="19" pageNumber="462" pagination="474 - 489" refId="ref18250" refString="Girard, M. G., Mundy, B. C., Nonaka, A. &amp; Johnson, G. D. (2023) Cusk-eel confusion: revisions of larval Luciobrotula and Pycnocraspedum and re-descriptions of two bythitid larvae (Ophidiiformes). Ichthyological Research, 70 (4), 474-489. https://doi.org/10.1007/s10228-023-00906-4" type="journal article" year="2023">
Girard
<emphasis id="B901EAD59002FFDCFB7EF9618D9DAE88" box="[1182,1243,1688,1714]" italics="true" pageId="19" pageNumber="462">et al.</emphasis>
2023
</bibRefCitation>
). Color of preserved large specimens medium to dark brown, with dusty dark brown coloration on scales.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9002FFDCFF27F9188A37AFB0" pageId="19" pageNumber="462" type="discussion">
<paragraph id="8BCA36C79002FFDCFF27F9188A37AFB0" blockId="19.[151,1437,572,2039]" pageId="19" pageNumber="462">
<emphasis id="B901EAD59002FFDCFF27F918880AAEC0" bold="true" box="[199,332,1760,1786]" pageId="19" pageNumber="462">Discussion.</emphasis>
<taxonomicName id="4C754D449002FFDCFEB5F9188BE6AEC0" baseAuthorityName="Parr" baseAuthorityYear="1933" box="[341,672,1760,1786]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="19" pageNumber="462" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD59002FFDCFEB5F9188BE6AEC0" box="[341,672,1760,1786]" italics="true" pageId="19" pageNumber="462">Pycnocraspedum phyllosoma</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C754D449002FFDCFCC2F9198AE3AEC0" authorityName=", Gosline" authorityYear="1954" box="[802,933,1761,1786]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="19" pageNumber="462" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD59002FFDCFCC2F9198AE3AEC0" box="[802,933,1761,1786]" italics="true" pageId="19" pageNumber="462">P. armatum</emphasis>
</taxonomicName>
in many traits and differs only in relatively subtle features such as the number of spines on the preopercle (3 vs. 4), 7278 transversal scale rows (vs. 6068), 1821 pyloric coeca (vs. 1216), and a relatively dark body coloration. Its otoliths show a symmetrical shape of dorsal and ventral rim while in other species the ventral rim is shallower than the dorsal rim or vice versa in
<taxonomicName id="4C754D449002FFDCFA68F8B58860AFB0" authorityName="Alcock" authorityYear="1889" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="19" pageNumber="462" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59002FFDCFA68F8B58860AFB0" italics="true" pageId="19" pageNumber="462">P. squamipinne</emphasis>
</taxonomicName>
, where the ventral rim is deeper than the dorsal rim.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9002FFDCFF27F86C8A1FAFCD" pageId="19" pageNumber="462" type="distribution">
<paragraph id="8BCA36C79002FFDCFF27F86C8A1FAFCD" blockId="19.[151,1437,572,2039]" pageId="19" pageNumber="462">
<emphasis id="B901EAD59002FFDCFF27F86C8827AF94" bold="true" box="[199,353,1940,1966]" pageId="19" pageNumber="462">Distribution.</emphasis>
<taxonomicName id="4C754D449002FFDCFE8BF86C8BFEAF94" baseAuthorityName="Parr" baseAuthorityYear="1933" box="[363,696,1940,1966]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="19" pageNumber="462" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD59002FFDCFE8BF86C8BFEAF94" box="[363,696,1940,1966]" italics="true" pageId="19" pageNumber="462">Pycnocraspedum phyllosoma</emphasis>
</taxonomicName>
is the only species of the genus known from the tropical West Atlantic while all other extant species of
<taxonomicName id="4C754D449002FFDCFDBDF8408A58AFE8" authorityName="Alcock" authorityYear="1889" box="[605,798,1976,2002]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="19" pageNumber="462" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59002FFDCFDBDF8408A58AFE8" box="[605,798,1976,2002]" italics="true" pageId="19" pageNumber="462">Pycnocraspedum</emphasis>
</taxonomicName>
occur in the Indo-West Pacific. The
<quantity id="4C8D9B229002FFDCFB53F8408C49AFE9" box="[1203,1295,1976,2003]" metricMagnitude="-1" metricUnit="m" metricValue="1.3599999999999999" pageId="19" pageNumber="462" unit="mm" value="136.0">136 mm</quantity>
SL
<typeStatus id="54CE88659002FFDCFADAF8408CDAAFE8" box="[1338,1436,1976,2002]" pageId="19" pageNumber="462" type="holotype">holotype</typeStatus>
was caught pelagic in the
<collectingCountry id="F36276579002FFDCFE59F8248B02AFCC" box="[441,580,2012,2038]" name="Turks Island" pageId="19" pageNumber="462">Turks Island</collectingCountry>
Passage, Pawnee St. 52.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,62 +1,62 @@
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@ -76,34 +76,36 @@
<figureCitation id="134E2A429004FFDAFF77FF438847A8EF" box="[151,257,187,213]" captionStart="FIGURE 3" captionStartId="11.[151,250,1786,1810]" captionTargetBox="[256,1328,181,1748]" captionTargetId="figure-17@11.[256,1331,181,1762]" captionTargetPageId="11" captionText="FIGURE 3. Detail figures and drawings of preopercular spines and lower preopercular pores in Pycnocraspedum; preopercular spines are numbered from 1 through 4 from the lower angle of the preoperculum upwards; abbreviation lpp = lower preopercular pore. 3A3B) Pycnocraspedum armatum, USNM 227412, 3A1-3A2 specimen 1 left side, 3A3 specimen 1 right side (mirror imaged), 3B specimen 2; 3C3D) Pycnocraspedum phyllosoma, 3C USNM 421587, 3D USNM 421586; 3E) Pycnocraspedum fulvum (holotype), BSKU 27497; 3F3G) Pycnocraspedum africanum sp. nov., 3F holotype, MNHN 2000-5691, 3G paratype, MNHN 2000-5692; 3H) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021; 3I3K) Pycnocraspedum squamipinne, 3I lectotype, BMNH 1890.7.31.1, 3J ZMUC P771871, 3K ZMUC P771737." figureDoi="http://doi.org/10.5281/zenodo.15561739" httpUri="https://zenodo.org/record/15561739/files/figure.png" pageId="21" pageNumber="464">Figures 3</figureCitation>
,
<figureCitation id="134E2A429004FFDAFEEEFF438858A8EC" box="[270,286,187,214]" captionStart="FIGURE 5" captionStartId="14.[151,250,1750,1774]" captionTargetBox="[245,1358,193,1715]" captionTargetId="figure-23@14.[222,1366,181,1726]" captionTargetPageId="14" captionText="FIGURE 5. Otoliths, inner faces (if not noted otherwise) shown from right side. 5A5C) Pycnocraspedum africanum sp. nov., 5A holotype, MNHN 2000-5691 (5A2 dorsal view), 5B paratype MNHN 2000-5692, 5C tentatively assigned specimen, IRSNB (5C2 dorsal view, 5C3 anterior view); 5D5E) Pycnocraspedum armatum, 5D USNM 227412 (5D2 ventral view), 5E USNM 227390 (5E2 ventral view); 5F) Pycnocraspedum fulvum (holotype), BSKU 27497 (5F2 ventral view, 5F3 anterior view); 5G) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021 (5G2 ventral view); 5H) Pycnocraspedum phyllosoma, USNM 421587 (5H2 ventral view, 5H3 anterior view); 5I5K) Pycnocraspedum squamipinne, 5I lectotype, BMNH 1890.7.31.1 (5I2 ventral view), 5J ZMUC P771871, 5L ZMUC P771739 (5L2 ventral view), 5K ZMUC P771738 (5K2 ventral view); 5M) †Pycnocraspedum cetonaense (holotype) SMF P.5694 (5M2 dorsal view, 5M3 anterior view)." figureDoi="http://doi.org/10.5281/zenodo.15561743" httpUri="https://zenodo.org/record/15561743/files/figure.png" pageId="21" pageNumber="464">5</figureCitation>
, 10, 12;
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,
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<figureCitation id="134E2A429004FFDAFECAFF43880EA8EF" box="[298,328,187,213]" captionStart="FIGURE 10" captionStartId="21.[151,250,1383,1407]" captionTargetBox="[171,1416,561,1349]" captionTargetId="figure-155@21.[151,1435,561,1359]" captionText="FIGURE 10. Pycnocraspedum rowleyensis sp. nov.; 10A) holotype, WAM P28058.021, 18°05'S, 118°10'E, 400 m; 10B) radiograph of holotype." figureDoi="http://doi.org/10.5281/zenodo.15595598" httpUri="https://zenodo.org/record/15595598/files/figure.png" pageId="21" pageNumber="464">10</figureCitation>
, 12;
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,
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<quantity id="4C8D9B229004FFDAFF77FEDF8998A97B" box="[151,222,295,321]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="21" pageNumber="464" unit="km" value="80.0">80 km</quantity>
southwest of
<collectingMunicipality id="6BAEACBD9004FFDAFE9AFEDF8B63A97B" box="[378,549,295,321]" pageId="21" pageNumber="464">Rowley Shoals</collectingMunicipality>
southwest of Rowley Shoals
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,
<quantity id="4C8D9B229004FFDAFDD1FEDF8B3CA978" box="[561,634,295,322]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="464" unit="m" value="400.0">
<elevation id="0058D1F49004FFDAFDD1FEDF8B3CA978" box="[561,634,295,322]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="464" unit="m" value="400.0">400 m</elevation>
@ -120,13 +122,15 @@ and
.
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<subSubSection id="C36F654C9004FFDAFF27FEB38807AA23" pageId="21" pageNumber="464" type="diagnosis">
<paragraph id="8BCA36C79004FFDAFF27FEB38807AA23" blockId="21.[151,1437,259,537]" pageId="21" pageNumber="464">
<emphasis id="B901EAD59004FFDAFF27FEB38804A95F" bold="true" box="[199,322,331,357]" pageId="21" pageNumber="464">Diagnosis.</emphasis>
Precaudal vertebrae 12, total vertebrae 52; dorsal-fin rays 87; anal-fin rays 69; pectoral-fin rays 25; long gill rakers on first gill arch 4; pseudobranchial filaments 45; 2 dorsal-fin rays in front of first vertebra; HL 25.3% of SL; body depth at vent 18.9% of SL; predorsal length 23% of SL; transverse scale rows above origin of anal fin 70, and above lateral line 17; broad expansion on preopercular angle with two spines, concave section above spine 2 without fleshy flap on hind margin of preopercle; otolith (
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in length) OL:OH = 2.1, dorsal rim crenulated.
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<emphasis id="B901EAD59004FFDAFF77FA9F8862AD45" bold="true" box="[151,292,1383,1407]" pageId="21" pageNumber="464">FIGURE 10.</emphasis>
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@ -151,7 +155,7 @@ in length) OL:OH = 2.1, dorsal rim crenulated.
<typeStatus id="54CE88659004FFDAFEC7FA7388C5AD99" box="[295,387,1419,1443]" pageId="21" pageNumber="464" type="holotype">holotype</typeStatus>
.
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<emphasis id="B901EAD59004FFDAFF27FA0D8817AE35" bold="true" box="[199,337,1525,1551]" pageId="21" pageNumber="464">Description</emphasis>

View file

@ -0,0 +1,479 @@
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<taxonomicName id="4C754D449007FFD9FF77FDED8BF6AA15" ID-CoL="4QM83" authority="Alcock, 1889" authorityName="Alcock" authorityYear="1889" box="[151,688,533,560]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="465" phylum="Chordata" rank="species" species="squamipinne">
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<emphasis id="B901EAD59007FFD9FF77FDED8B4FAA15" bold="true" box="[151,521,533,560]" italics="true" pageId="22" pageNumber="465">Pycnocraspedum squamipinne</emphasis>
<bibRefCitation id="EFE44B369007FFD9FDF0FDED8BF6AA15" author="Alcock, A." box="[528,688,533,559]" pageId="22" pageNumber="465" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
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<figureCitation id="134E2A429007FFD9FF77FDC18846AA69" box="[151,256,569,595]" captionStart="FIGURE 1" captionStartId="3.[151,250,1527,1551]" captionTargetBox="[168,1411,202,1477]" captionTargetId="figure-17@3.[151,1436,181,1504]" captionTargetPageId="3" captionText="FIGURE 1. Radiograph figures of selected specimens of the genus Pycnocraspedum to visualize position of dorsal-fin rays in relation to first vertebra; “1” denotes the first dorsal-fin ray above the first vertebra; numbers with negative denotation represent dorsal-fin rays in front of the first vertebra. 1A) Pycnocraspedum armatum, USNM 227390; 1B) Pycnocraspedum phyllosoma USNM 227413; 1C) Pycnocraspedum squamipinne, ZMUC P771871; 1D) Pycnocraspedum fulvum (holotype), BSKU 27497." figureDoi="http://doi.org/10.5281/zenodo.15561735" httpUri="https://zenodo.org/record/15561735/files/figure.png" pageId="22" pageNumber="465">Figures 1</figureCitation>
,
<figureCitation id="134E2A429007FFD9FEEEFDC1885BAA69" box="[270,285,569,595]" captionStart="FIGURE 3" captionStartId="11.[151,250,1786,1810]" captionTargetBox="[256,1328,181,1748]" captionTargetId="figure-17@11.[256,1331,181,1762]" captionTargetPageId="11" captionText="FIGURE 3. Detail figures and drawings of preopercular spines and lower preopercular pores in Pycnocraspedum; preopercular spines are numbered from 1 through 4 from the lower angle of the preoperculum upwards; abbreviation lpp = lower preopercular pore. 3A3B) Pycnocraspedum armatum, USNM 227412, 3A1-3A2 specimen 1 left side, 3A3 specimen 1 right side (mirror imaged), 3B specimen 2; 3C3D) Pycnocraspedum phyllosoma, 3C USNM 421587, 3D USNM 421586; 3E) Pycnocraspedum fulvum (holotype), BSKU 27497; 3F3G) Pycnocraspedum africanum sp. nov., 3F holotype, MNHN 2000-5691, 3G paratype, MNHN 2000-5692; 3H) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021; 3I3K) Pycnocraspedum squamipinne, 3I lectotype, BMNH 1890.7.31.1, 3J ZMUC P771871, 3K ZMUC P771737." figureDoi="http://doi.org/10.5281/zenodo.15561739" httpUri="https://zenodo.org/record/15561739/files/figure.png" pageId="22" pageNumber="465">3</figureCitation>
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,
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,
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;
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,
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<emphasis id="B901EAD59007FFD9FF77FD8F8888AAB5" box="[151,462,631,655]" italics="true" pageId="22" pageNumber="465">Pycnocraspedum squamipinne</emphasis>
<bibRefCitation id="EFE44B369007FFD9FE35FD8F8BD0AAB5" author="Alcock, A." box="[469,662,631,655]" pageId="22" pageNumber="465" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889: 386</bibRefCitation>
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; figured in
<bibRefCitation id="EFE44B369007FFD9FCECFD8F8AD5AAB5" author="Alcock, A." box="[780,915,631,655]" pageId="22" pageNumber="465" pagination="136 - 156" refId="ref17617" refString="Alcock, A. (1898) Natural history notes from H. M. Indian Survey Steamer &quot; Investigator &quot;. Series II. No. 25. A note on the deep-sea fishes, with descriptions of some new genera and species, including another probably viviparous ophidioid. Annals Magazine Natural History, Series 7, 2 (8), 136-156. https://doi.org/10.1080/00222939808678029" type="journal article" year="1898">Alcock 1898</bibRefCitation>
: pl. 21, fig. 1 as
<taxonomicName id="4C754D449007FFD9FBA2FD8F8CC6AAAA" authority=", Bay" authorityName="Bay" box="[1090,1408,631,656]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="465" phylum="Chordata" rank="species" species="squamipinnis">
<emphasis id="B901EAD59007FFD9FBA2FD8F8C0FAAB5" box="[1090,1353,631,655]" italics="true" pageId="22" pageNumber="465">Neobythites squamipinnis</emphasis>
, Bay
</taxonomicName>
of Bengal,
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,
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, RV Investigator, depth 193 fathoms (
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).
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</paragraph>
<paragraph id="8BCA36C79007FFD9FF77FD4C8854AB30" blockId="22.[151,1436,631,778]" pageId="22" pageNumber="465">
<taxonomicName id="4C754D449007FFD9FF77FD4C8B20AAF7" ID-CoL="4QM83" authority="Alcock, 1889" authorityName="Alcock" authorityYear="1889" box="[151,614,692,717]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="22" pageNumber="465" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59007FFD9FF77FD4C8896AAF6" box="[151,464,692,716]" italics="true" pageId="22" pageNumber="465">Pycnocraspedum squamipinne</emphasis>
<bibRefCitation id="EFE44B369007FFD9FE38FD4C8B20AAF7" author="Alcock, A." box="[472,614,692,717]" pageId="22" pageNumber="465" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="EFE44B369007FFD9FD93FD4D8AC6AAF6" author="Menon, A. G. K. &amp; Yazdani, G. M." box="[627,896,692,717]" pageId="22" pageNumber="465" pagination="91 - 190" refId="ref18832" refString="Menon, A. G. K. &amp; Yazdani, G. M. (1968) Catalogue of type-specimens in the Zoological Survey of India, part 2 - Fishes. Records of the Zoological Survey of India, 61 (1 - 2), 91-190. https://doi.org/10.26515/rzsi/v61/i1-2/1963/161616" type="journal article" year="1968">Menon &amp; Yazdani (1968)</bibRefCitation>
,
<bibRefCitation id="EFE44B369007FFD9FC6DFD4D8DF0AAF7" author="Menon, A. G. K. &amp; Rama-Rao, K. V." box="[909,1206,693,717]" pageId="22" pageNumber="465" pagination="377 - 378" refId="ref18714" refString="Menon, A. G. K. &amp; Rama-Rao, K. V. (1970) Type-specimens of fishes described in the R. I. M. S. &quot; Investigator &quot; collections (1884 - 1926). Copeia, 1970 (2), 377-378. https://doi.org/10.2307/1441668" type="journal article" year="1970">Menon &amp; Rama-Rao (1970)</bibRefCitation>
,
<bibRefCitation id="EFE44B369007FFD9FB22FD4D884BAAD1" author="Menon, A. G. K. &amp; Rama-Rao, K. V." pageId="22" pageNumber="465" pagination="31 - 48" refId="ref18777" refString="Menon, A. G. K. &amp; Rama-Rao, K. V. (1975) A catalogue of type specimens of fishes described in the biological collections of R. I. M. S. &quot; Investigator &quot; during 1884 - 1926. Matsya, 1, 31-48." type="journal article" year="1975">Menon &amp; Rama-Rao (1975)</bibRefCitation>
,
<bibRefCitation id="EFE44B369007FFD9FEFBFD2B88BEAAD1" author="Nielsen, J. G. &amp; Cohen, D. M. &amp; Markle, D. F. &amp; Robins, C. R." box="[283,504,723,747]" pageId="22" pageNumber="465" refId="ref19264" refString="Nielsen, J. G., Cohen, D. M., Markle, D. F. &amp; Robins, C. R. (1999) An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiiform fishes known to date. FAO Fisheries Synopsis No. 125. Vol. 18. FAO, Rome, 178 pp." type="book" year="1999">
Nielsen
<emphasis id="B901EAD59007FFD9FE92FD2C88EFAAD1" box="[370,425,723,747]" italics="true" pageId="22" pageNumber="465">et al.</emphasis>
(1999)
</bibRefCitation>
, Evseenko &amp; Okiyama (2006),
<bibRefCitation id="EFE44B369007FFD9FCA9FD2B8D14AAD1" author="Psomadakis, P. N. &amp; Thein, H. &amp; Russell, B. C. &amp; Tun, M. T." box="[841,1106,723,747]" pageId="22" pageNumber="465" refId="ref19714" refString="Psomadakis, P. N., Thein, H., Russell, B. C. &amp; Tun, M. T. (2020) Field identification guide to the living marine resources of Myanmar. FAO species identification guide for fishery purposes. FAO and MOALI, Rome, xvii + 694 pp., 63 colour pls." type="book" year="2020">
Psomadakis
<emphasis id="B901EAD59007FFD9FC2CFD2C8D45AAD1" box="[972,1027,723,747]" italics="true" pageId="22" pageNumber="465">et al.</emphasis>
(2020)
</bibRefCitation>
,
<bibRefCitation id="EFE44B369007FFD9FBBFFD2C8C6DAAD1" author="Teena, T. K. &amp; Murugan, A. &amp; Kumar, A. T. T. &amp; Lal, K. K." box="[1119,1323,723,747]" pageId="22" pageNumber="465" pagination="77 - 83" refId="ref20468" refString="Teena, T. K., Murugan, A., Kumar, A. T. T. &amp; Lal, K. K. (2021) Redescription of a rare cusk eel, Pycnocraspedum squamipinne (Actinopterygii, Ophidiiformes, Ophidiidae) from Bay of Bengal. Acta Ichthyologica et Piscatoria, 51 (1), 77-83. https://doi.org/10.3897/aiep.51.63469" type="journal article" year="2021">
Teena
<emphasis id="B901EAD59007FFD9FB45FD2C8D9AAAD1" box="[1189,1244,723,747]" italics="true" pageId="22" pageNumber="465">et al.</emphasis>
(2021)
</bibRefCitation>
,
<bibRefCitation id="EFE44B369007FFD9FAD8FD2B884BAB30" author="Prokofiev, A. M." pageId="22" pageNumber="465" pagination="1019 - 1024" refId="ref19635" refString="Prokofiev, A. M. (2022) New data on the morphology and distribution of two rare species of Ophidiiformes: Diplacanthopoma japonicum (Bythitidae) and Pycnocraspedum microlepis (Neobythitidae). Voprosy Ikhtiologii, 62 (6), 680 - 685. [in Russian, English translation appeared in Journal of Ichthyology, 62 (6), 1019-1024] https://doi.org/10.1134/S0032945222060224" type="journal article" year="2022">Prokofiev (2022)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9007FFD9FF77FCC08C1BAC74" pageId="22" pageNumber="465" type="materials_examined">
<paragraph id="8BCA36C79007FFD9FF77FCC08C1BAC74" blockId="22.[151,1437,824,2039]" pageId="22" pageNumber="465">
<emphasis id="B901EAD59007FFD9FF77FCC08838AB68" bold="true" box="[151,382,824,850]" pageId="22" pageNumber="465">Material examined</emphasis>
(
<specimenCount id="9D73FD4E9007FFD9FE73FCC08B63AB68" box="[403,549,824,850]" count="6" pageId="22" pageNumber="465" type="generic">6 specimens</specimenCount>
,
<quantity id="4C8D9B229007FFD9FDD5FCC08B89AB69" box="[565,719,824,851]" metricMagnitude="-1" metricUnit="m" metricValue="1.825" metricValueMax="2.43" metricValueMin="1.22" pageId="22" pageNumber="465" unit="mm" value="182.5" valueMax="243.0" valueMin="122.0">122243 mm</quantity>
SL):
<materialsCitation id="3B1D3C9A9007FFD9FCF8FCC08B86AB4D" collectionCode="BMNH" collectorName="RV Investigator" elevation="353" latitude="20.291666" location="Bay of Bengal" longLatPrecision="868" longitude="88.833336" pageId="22" pageNumber="465" specimenCode="1890.7.31.1" specimenCount="1" typeStatus="lectotype">
<specimenCount id="9D73FD4E9007FFD9FCF8FCC08ADAAB68" box="[792,924,824,850]" count="1" pageId="22" pageNumber="465" type="generic">1 specimen</specimenCount>
<typeStatus id="54CE88659007FFD9FC47FCC08D57AB68" box="[935,1041,824,850]" pageId="22" pageNumber="465" type="lectotype">lectotype</typeStatus>
,
<collectionCode id="ED64AE029007FFD9FBC1FCC18D33AB69" box="[1057,1141,825,851]" pageId="22" pageNumber="465">BMNH</collectionCode>
<specimenCode id="DBD39EBC9007FFD9FB60FCC08C4EAB68" box="[1152,1288,824,850]" pageId="22" pageNumber="465">1890.7.31.1</specimenCode>
,
<geoCoordinate id="EE4150009007FFD9FAF8FCC08CDBAB68" box="[1304,1437,824,850]" degrees="20" direction="north" minutes="17" orientation="latitude" pageId="22" pageNumber="465" precision="15" seconds="30" value="20.291666">20°17'30&quot;N</geoCoordinate>
<geoCoordinate id="EE4150009007FFD9FF77FCA489B7AB4D" box="[151,241,860,887]" degrees="88" direction="east" minutes="50" orientation="longitude" pageId="22" pageNumber="465" precision="925" value="88.833336">88°50'E</geoCoordinate>
(
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),
<collectorName id="268053119007FFD9FE59FCA58B2AAB4C" box="[441,620,860,887]" pageId="22" pageNumber="465">RV Investigator</collectorName>
,
<quantity id="4C8D9B229007FFD9FD96FCA48B86AB4D" box="[630,704,860,887]" metricMagnitude="2" metricUnit="m" metricValue="3.53" pageId="22" pageNumber="465" unit="m" value="353.0">
<elevation id="0058D1F49007FFD9FD96FCA48B86AB4D" box="[630,704,860,887]" metricMagnitude="2" metricUnit="m" metricValue="3.53" pageId="22" pageNumber="465" unit="m" value="353.0">353 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A9007FFD9FD2BFCA48CD1ABA1" collectingDate="2015-05-03" collectionCode="ZMUC, R, V" collectorName="Dr. Fridtjof Nansen" country="Myanmar" elevation="129" latitude="18.266666" location="Bay of Bengal" longLatPrecision="1275" longitude="93.71667" pageId="22" pageNumber="465" specimenCount="1" typeStatus="lectotype">
<specimenCount id="9D73FD4E9007FFD9FD2BFCA48A0CAB4C" box="[715,842,860,886]" count="1" pageId="22" pageNumber="465" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE029007FFD9FCB0FCA48AE7AB4C" box="[848,929,860,886]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="22" pageNumber="465">ZMUC</collectionCode>
P
<quantity id="4C8D9B229007FFD9FC5DFCA48D3AAB4D" box="[957,1148,860,887]" metricMagnitude="5" metricUnit="m" metricValue="7.71737226" pageId="22" pageNumber="465" unit="mm" value="7.71737226E8">771737, 226 mm</quantity>
SL, off
<collectingCountry id="F36276579007FFD9FB38FCA58C01AB4D" box="[1240,1351,861,887]" name="Myanmar" pageId="22" pageNumber="465">Myanmar</collectingCountry>
,
<location id="8EAA601C9007FFD9FAB1FCA589AFABA0" LSID="urn:lsid:plazi:treatment:03DC87D19007FFD7FF77FDED899EAE7D:8EAA601C9007FFD9FAB1FCA589AFABA0" country="Myanmar" latitude="18.266666" longLatPrecision="1275" longitude="93.71667" name="Bay of Bengal" pageId="22" pageNumber="465">Bay of Bengal</location>
,
<geoCoordinate id="EE4150009007FFD9FF14FC788815ABA0" box="[244,339,896,922]" degrees="18" direction="north" minutes="16" orientation="latitude" pageId="22" pageNumber="465" precision="925" value="18.266666">18°16'N</geoCoordinate>
,
<geoCoordinate id="EE4150009007FFD9FEBEFC7888FFABA0" box="[350,441,896,922]" degrees="93" direction="east" minutes="43" orientation="longitude" pageId="22" pageNumber="465" precision="925" value="93.71667">93°43'E</geoCoordinate>
,
<collectionCode id="ED64AE029007FFD9FE25FC788891ABA0" box="[453,471,896,922]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="465">R</collectionCode>
/
<collectionCode id="ED64AE029007FFD9FE3FFC7888B4ABA0" box="[479,498,896,922]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="465" type="Museum">V</collectionCode>
<emphasis id="B901EAD59007FFD9FE19FC798B92ABA0" box="[505,724,896,922]" italics="true" pageId="22" pageNumber="465">
<collectorName id="268053119007FFD9FE19FC798B92ABA0" box="[505,724,896,922]" pageId="22" pageNumber="465">Dr. Fridtjof Nansen</collectorName>
</emphasis>
, survey 2015404, st.19, bottom trawl,
<quantity id="4C8D9B229007FFD9FB62FC788C45ABA1" box="[1154,1283,896,923]" metricMagnitude="2" metricUnit="m" metricValue="1.285" metricValueMax="1.3" metricValueMin="1.27" pageId="22" pageNumber="465" unit="m" value="128.5" valueMax="130.0" valueMin="127.0">
<elevation id="0058D1F49007FFD9FB62FC788C45ABA1" box="[1154,1283,896,923]" metricMagnitude="2" metricUnit="m" metricValue="1.285" metricValueMax="1.3" metricValueMin="1.27" pageId="22" pageNumber="465" unit="m" value="128.5" valueMax="130.0" valueMin="127.0">127130 m</elevation>
</quantity>
,
<date id="FFCB10079007FFD9FAEEFC788CD1ABA1" box="[1294,1431,896,923]" pageId="22" pageNumber="465" value="2015-05-03">
<collectingDate id="EF8FE9EF9007FFD9FAEEFC788CD1ABA1" box="[1294,1431,896,923]" pageId="22" pageNumber="465" value="2015-05-03">3 May 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A9007FFD9FF77FC5C8D02ABD9" collectingDate="2015-05-03" collectionCode="ZMUC, R, V" collectorName="Dr. Fridtjof Nansen &amp; St." country="Myanmar" elevation="468" latitude="18.233334" location="Bay of Bengal" longLatPrecision="1276" longitude="93.63333" pageId="22" pageNumber="465" specimenCode="P 771736" specimenCount="2" typeStatus="lectotype">
<specimenCount id="9D73FD4E9007FFD9FF77FC5C8866AB84" box="[151,288,932,958]" count="2" pageId="22" pageNumber="465" type="generic">2 specimens</specimenCount>
<collectionCode id="ED64AE029007FFD9FEC5FC5C8830AB84" box="[293,374,932,958]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="22" pageNumber="465">ZMUC</collectionCode>
<specimenCode id="DBD39EBC9007FFD9FE9CFC5D88A2AB84" box="[380,484,932,959]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="22" pageNumber="465">P 771736</specimenCode>
and P 771738, 120
<quantity id="4C8D9B229007FFD9FD27FC5C8A67AB85" box="[711,801,932,959]" metricMagnitude="-1" metricUnit="m" metricValue="1.95" pageId="22" pageNumber="465" unit="mm" value="195.0">195 mm</quantity>
SL, off
<collectingCountry id="F36276579007FFD9FC9AFC5D8AAFAB85" box="[890,1001,933,959]" name="Myanmar" pageId="22" pageNumber="465">Myanmar</collectingCountry>
,
<location id="8EAA601C9007FFD9FC13FC5D8DD3AB84" LSID="urn:lsid:plazi:treatment:03DC87D19007FFD7FF77FDED899EAE7D:8EAA601C9007FFD9FC13FC5D8DD3AB84" box="[1011,1173,932,959]" country="Myanmar" latitude="18.233334" longLatPrecision="1276" longitude="93.63333" name="Bay of Bengal" pageId="22" pageNumber="465">Bay of Bengal</location>
,
<geoCoordinate id="EE4150009007FFD9FB7FFC5C8DB8AB84" box="[1183,1278,932,958]" degrees="18" direction="north" minutes="14" orientation="latitude" pageId="22" pageNumber="465" precision="925" value="18.233334">18°14'N</geoCoordinate>
,
<geoCoordinate id="EE4150009007FFD9FAE8FC5C8C25AB84" box="[1288,1379,932,958]" degrees="93" direction="east" minutes="38" orientation="longitude" pageId="22" pageNumber="465" precision="925" value="93.63333">93°38'E</geoCoordinate>
,
<collectionCode id="ED64AE029007FFD9FA8EFC5C8CC6AB84" box="[1390,1408,932,958]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="465">R</collectionCode>
/
<collectionCode id="ED64AE029007FFD9FA69FC5C8CDAAB84" box="[1417,1436,932,958]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="465" type="Museum">V</collectionCode>
<emphasis id="B901EAD59007FFD9FF77FC318835ABD8" box="[151,371,968,994]" italics="true" pageId="22" pageNumber="465">
<collectorName id="268053119007FFD9FF77FC318835ABD8" box="[151,371,968,994]" pageId="22" pageNumber="465">Dr. Fridtjof Nansen</collectorName>
</emphasis>
, survey 2015404,
<collectorName id="268053119007FFD9FDA2FC308B26ABD8" box="[578,608,968,994]" pageId="22" pageNumber="465">St.</collectorName>
18, bottom trawl,
<quantity id="4C8D9B229007FFD9FCCEFC308AE8ABD9" box="[814,942,968,995]" metricMagnitude="2" metricUnit="m" metricValue="4.675" metricValueMax="4.6899999999999995" metricValueMin="4.66" pageId="22" pageNumber="465" unit="m" value="467.5" valueMax="469.0" valueMin="466.0">
<elevation id="0058D1F49007FFD9FCCEFC308AE8ABD9" box="[814,942,968,995]" metricMagnitude="2" metricUnit="m" metricValue="4.675" metricValueMax="4.6899999999999995" metricValueMin="4.66" pageId="22" pageNumber="465" unit="m" value="467.5" valueMax="469.0" valueMin="466.0">466469 m</elevation>
</quantity>
,
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<collectingDate id="EF8FE9EF9007FFD9FC5AFC308D02ABD9" box="[954,1092,968,995]" pageId="22" pageNumber="465" value="2015-05-03">3 May 2015</collectingDate>
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;
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<specimenCount id="9D73FD4E9007FFD9FBAFFC308D88ABD8" box="[1103,1230,968,994]" count="1" pageId="22" pageNumber="465" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE029007FFD9FB34FC308C63ABD8" box="[1236,1317,968,994]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="22" pageNumber="465">ZMUC</collectionCode>
P
<quantity id="4C8D9B229007FFD9FAA1FC3089B1AC3D" metricMagnitude="5" metricUnit="m" metricValue="7.71739225" pageId="22" pageNumber="465" unit="mm" value="7.71739225E8">771739, 225 mm</quantity>
SL, off
<collectingCountry id="F36276579007FFD9FEBBFC15888CAC3D" box="[347,458,1005,1031]" name="Myanmar" pageId="22" pageNumber="465">Myanmar</collectingCountry>
,
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,
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,
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,
<collectionCode id="ED64AE029007FFD9FC86FC148A3EAC3C" box="[870,888,1004,1030]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="465">R</collectionCode>
/
<collectionCode id="ED64AE029007FFD9FC60FC148AD5AC3C" box="[896,915,1004,1030]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="465" type="Museum">V</collectionCode>
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</emphasis>
, survey 2015404, st. 42, bottom trawl,
<quantity id="4C8D9B229007FFD9FED7FBE888FFAC11" box="[311,441,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="2.755" metricValueMax="2.77" metricValueMin="2.7399999999999998" pageId="22" pageNumber="465" unit="m" value="275.5" valueMax="277.0" valueMin="274.0">
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</quantity>
,
<date id="FFCB10079007FFD9FE26FBE88B14AC11" box="[454,594,1040,1067]" pageId="22" pageNumber="465" value="2015-05-06">
<collectingDate id="EF8FE9EF9007FFD9FE26FBE88B14AC11" box="[454,594,1040,1067]" pageId="22" pageNumber="465" value="2015-05-06">6 May 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A9007FFD9FDBFFBE88C1CAC74" collectingDate="2018-08-29" collectionCode="ZMUC, R, V" collectorName="Dr. Fridtjof Nansen" country="Myanmar" elevation="229" latitude="16.433332" location="Bay of Bengal" longLatPrecision="1282" longitude="93.95" pageId="22" pageNumber="465" specimenCount="1" typeStatus="lectotype">
<specimenCount id="9D73FD4E9007FFD9FDBFFBE88B99AC10" box="[607,735,1040,1066]" count="1" pageId="22" pageNumber="465" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE029007FFD9FD07FBE88A7EAC10" box="[743,824,1040,1066]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="22" pageNumber="465">ZMUC</collectionCode>
P
<quantity id="4C8D9B229007FFD9FCB6FBE88D51AC11" box="[854,1047,1040,1067]" metricMagnitude="5" metricUnit="m" metricValue="7.7187119599999985" pageId="22" pageNumber="465" unit="mm" value="7.71871196E8">771871, 196 mm</quantity>
SL, off
<collectingCountry id="F36276579007FFD9FB97FBE98DA0AC11" box="[1143,1254,1041,1067]" name="Myanmar" pageId="22" pageNumber="465">Myanmar</collectingCountry>
,
<location id="8EAA601C9007FFD9FB11FBE98CDEAC10" LSID="urn:lsid:plazi:treatment:03DC87D19007FFD7FF77FDED899EAE7D:8EAA601C9007FFD9FB11FBE98CDEAC10" box="[1265,1432,1040,1067]" country="Myanmar" latitude="16.433332" longLatPrecision="1282" longitude="93.95" name="Bay of Bengal" pageId="22" pageNumber="465">Bay of Bengal</location>
,
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,
<geoCoordinate id="EE4150009007FFD9FEE2FBCC881BAC75" box="[258,349,1076,1103]" degrees="93" direction="east" minutes="57" orientation="longitude" pageId="22" pageNumber="465" precision="925" value="93.95">93°57'E</geoCoordinate>
,
<collectionCode id="ED64AE029007FFD9FE89FBCC883DAC74" box="[361,379,1076,1102]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="465">R</collectionCode>
/
<collectionCode id="ED64AE029007FFD9FE64FBCC88D1AC74" box="[388,407,1076,1102]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="465" type="Museum">V</collectionCode>
<emphasis id="B901EAD59007FFD9FE7EFBCD8B3CAC74" box="[414,634,1076,1102]" italics="true" pageId="22" pageNumber="465">
<collectorName id="268053119007FFD9FE7EFBCD8B3CAC74" box="[414,634,1076,1102]" pageId="22" pageNumber="465">Dr. Fridtjof Nansen</collectorName>
</emphasis>
, survey 2018411, st. 26, bottom trawl, 241
<quantity id="4C8D9B229007FFD9FB89FBCC8DF4AC75" box="[1129,1202,1076,1103]" metricMagnitude="2" metricUnit="m" metricValue="2.29" pageId="22" pageNumber="465" unit="m" value="229.0">
<elevation id="0058D1F49007FFD9FB89FBCC8DF4AC75" box="[1129,1202,1076,1103]" metricMagnitude="2" metricUnit="m" metricValue="2.29" pageId="22" pageNumber="465" unit="m" value="229.0">229 m</elevation>
</quantity>
,
<date id="FFCB10079007FFD9FB5EFBCC8C1CAC74" box="[1214,1370,1076,1102]" pageId="22" pageNumber="465" value="2018-08-29">
<collectingDate id="EF8FE9EF9007FFD9FB5EFBCC8C1CAC74" box="[1214,1370,1076,1102]" pageId="22" pageNumber="465" value="2018-08-29">29 Aug. 2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9007FFD9FF27FBA08BBEAD70" pageId="22" pageNumber="465" type="diagnosis">
<paragraph id="8BCA36C79007FFD9FF27FBA08BBEAD70" blockId="22.[151,1437,824,2039]" pageId="22" pageNumber="465">
<emphasis id="B901EAD59007FFD9FF27FBA08804AC48" bold="true" box="[199,322,1112,1138]" pageId="22" pageNumber="465">Diagnosis.</emphasis>
Precaudal vertebrae 1213, total vertebrae 4751; dorsal-fin rays 9092; anal-fin rays 5765; pectoral-fin rays 2530; long gill rakers on first gill arch 4; pseudobranchials 12; 13 dorsal-fin rays in front of first vertebra; HL 29.631.5% of SL; body depth at vent 17.318% of SL; predorsal length 2529% of SL; transversal scale rows above origin of anal fin 5258 (
<quantity id="4C8D9B229007FFD9FD8DFB3C8B96ACE4" box="[621,720,1220,1247]" metricMagnitude="0" metricUnit="m" metricValue="1.4986" metricValueMax="1.524" metricValueMin="1.4731999999999998" pageId="22" pageNumber="465" unit="in" value="59.0" valueMax="60.0" valueMin="58.0">5860 in</quantity>
<typeStatus id="54CE88659007FFD9FD36FB3C8A3EACE4" box="[726,888,1220,1246]" pageId="22" pageNumber="465" type="paralectotype">paralectotypes</typeStatus>
according to
<bibRefCitation id="EFE44B369007FFD9FBECFB3D8D8BACE4" author="Teena, T. K. &amp; Murugan, A. &amp; Kumar, A. T. T. &amp; Lal, K. K." box="[1036,1229,1220,1247]" pageId="22" pageNumber="465" pagination="77 - 83" refId="ref20468" refString="Teena, T. K., Murugan, A., Kumar, A. T. T. &amp; Lal, K. K. (2021) Redescription of a rare cusk eel, Pycnocraspedum squamipinne (Actinopterygii, Ophidiiformes, Ophidiidae) from Bay of Bengal. Acta Ichthyologica et Piscatoria, 51 (1), 77-83. https://doi.org/10.3897/aiep.51.63469" type="journal article" year="2021">
Teena
<emphasis id="B901EAD59007FFD9FBB5FB3D8DCAACE4" box="[1109,1164,1220,1246]" italics="true" pageId="22" pageNumber="465">et al.</emphasis>
2021
</bibRefCitation>
), and above lateral line 1317 (
<quantity id="4C8D9B229007FFD9FEFFFB1088C4AD38" box="[287,386,1256,1283]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.81" metricValueMin="2.794" pageId="22" pageNumber="465" unit="in" value="13.0" valueMax="15.0" valueMin="11.0">1115 in</quantity>
<typeStatus id="54CE88659007FFD9FE6AFB108B6AAD38" box="[394,556,1256,1282]" pageId="22" pageNumber="465" type="paralectotype">paralectotypes</typeStatus>
according to
<bibRefCitation id="EFE44B369007FFD9FD26FB118ACAAD38" author="Teena, T. K. &amp; Murugan, A. &amp; Kumar, A. T. T. &amp; Lal, K. K." box="[710,908,1256,1283]" pageId="22" pageNumber="465" pagination="77 - 83" refId="ref20468" refString="Teena, T. K., Murugan, A., Kumar, A. T. T. &amp; Lal, K. K. (2021) Redescription of a rare cusk eel, Pycnocraspedum squamipinne (Actinopterygii, Ophidiiformes, Ophidiidae) from Bay of Bengal. Acta Ichthyologica et Piscatoria, 51 (1), 77-83. https://doi.org/10.3897/aiep.51.63469" type="journal article" year="2021">
Teena
<emphasis id="B901EAD59007FFD9FCF0FB118A0FAD38" box="[784,841,1256,1282]" italics="true" pageId="22" pageNumber="465">et al.</emphasis>
2021
</bibRefCitation>
); pyloric coeca 1213; preopercle with 1 weak spine above first lower preopercular pore and broad concave section above; otoliths relatively small, at&gt;
<quantity id="4C8D9B229007FFD9FAD1FAF48CC6AD1D" box="[1329,1408,1292,1319]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="465" unit="mm" value="10.0">10 mm</quantity>
in length: OL:OH = 2.42.5, dorsal rim shallow, smooth.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9007FFD8FF27FAAC8D12A88B" lastPageId="23" lastPageNumber="466" pageId="22" pageNumber="465" type="description">
<paragraph id="8BCA36C79007FFD9FF27FAAC8C59AD8D" blockId="22.[151,1437,824,2039]" pageId="22" pageNumber="465">
<emphasis id="B901EAD59007FFD9FF27FAAC881FAD54" bold="true" box="[199,345,1364,1390]" pageId="22" pageNumber="465">Description.</emphasis>
Relatively robust fish with stout tapering tail and rounded snout.Maximal size of fishes investigated
<quantity id="4C8D9B229007FFD9FF77FA8089BEADA9" box="[151,248,1400,1427]" metricMagnitude="-1" metricUnit="m" metricValue="2.43" pageId="22" pageNumber="465" unit="mm" value="243.0">243 mm</quantity>
SL. Head stout, with straight, inclined dorsal profile. Preopercle with 1 weak spine above first lower preopercular pore, with broad concave section above and obtuse angle below second preopercular pore.
</paragraph>
<paragraph id="8BCA36C79007FFD9FF27FA388A3BAE7C" blockId="22.[151,1437,824,2039]" pageId="22" pageNumber="465">
Meristics (
<specimenCount id="9D73FD4E9007FFD9FEDEFA38888BADE0" box="[318,461,1472,1498]" count="6" pageId="22" pageNumber="465" type="generic">6 specimens</specimenCount>
): precaudal vertebrae 12 (1213); total vertebrae 49 (4751); dorsal-fin rays 92 (9092); anal-fin rays 65 (5765); pectoral-fin rays 28 (2530); caudal-fin rays 10; D/V = -2 (-1 -3); D/A = 2431; V/A = 1617; long gill rakers on first gill arch 4; pseudobranchial filaments 1 (12); transversal scale rows above origin of anal fin 5258, and above lateral line 1317; pyloric coeca 1213.
</paragraph>
<paragraph id="8BCA36C79007FFD9FF27F9A88B28AEEC" blockId="22.[151,1437,824,2039]" pageId="22" pageNumber="465">
Morphometrics (
<specimenCount id="9D73FD4E9007FFD9FE64F9A88B56AE50" box="[388,528,1616,1643]" count="5" pageId="22" pageNumber="465" type="generic">5 specimens</specimenCount>
) in % of SL: HL 29.6 (29.631.5); body depth at vent 17.3 (1718); diameter of eye 4.9 (4.55.1); upper jaw length 15.4 (15.015.4); interorbital width 5.2 (5.26.2); pelvic-fin base to anal-fin base 26 (2632.5); predorsal length 27 (2529); preanal length 4652; prepelvic length 2021.5; pectoral-fin length 15.2 (15.218); pelvic-fin length 11.414.
</paragraph>
<paragraph id="8BCA36C79007FFD9FF27F9188B32AFCC" blockId="22.[151,1437,824,2039]" pageId="22" pageNumber="465">
Otolith morphology (
<specimenCount id="9D73FD4E9007FFD9FE57F9188B01AEC0" box="[439,583,1760,1786]" count="3" pageId="22" pageNumber="465" type="generic">3 specimens</specimenCount>
). Size up to
<quantity id="4C8D9B229007FFD9FD30F9188A66AEC1" box="[720,800,1760,1787]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="465" unit="mm" value="12.0">12 mm</quantity>
in length; OL:OH =
<quantity id="4C8D9B229007FFD9FBECF9188D38AEC0" box="[1036,1150,1760,1787]" metricMagnitude="-2" metricUnit="m" metricValue="6.223" metricValueMax="6.35" metricValueMin="6.096" pageId="22" pageNumber="465" unit="in" value="2.45" valueMax="2.5" valueMin="2.4">2.42.5 in</quantity>
specimens larger than
<quantity id="4C8D9B229007FFD9FA60F9188985AF25" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="465" unit="mm" value="10.0">10 mm</quantity>
in length,
<quantity id="4C8D9B229007FFD9FEDDF8FC883BAF24" box="[317,381,1796,1822]" metricMagnitude="-2" metricUnit="m" metricValue="4.572" pageId="22" pageNumber="465" unit="in" value="1.8">1.8 in</quantity>
specimens smaller than
<quantity id="4C8D9B229007FFD9FD73F8FC8B93AF25" box="[659,725,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="22" pageNumber="465" unit="mm" value="6.0">6 mm</quantity>
in length; OH:OT = 2.62.8. Otolith relatively thin, elongated, anteriorly rounded, posteriorly rounded or slightly pointed; anterior and posterior tips along horizontal axis. Dorsal rim shallow, nearly flat or depressed at times, moderately lobed in small specimens; ventral rim shallow, regularly curved. All rims smooth. Inner face nearly flat, less bent than outer face. Sulcus moderately long, horizontal, with shallow, undivided, uniform colliculum with flat to straight dorsal rim and slightly convex ventral rim and rounded anterior and posterior tips. OL:CL = 1.51.6. Dorsal field without distinct depression; ventral field smooth. Outer face more convex than inner face, smooth.
</paragraph>
<paragraph id="8BCA36C79006FFD8FF27FF6F8D12A88B" blockId="23.[151,1437,151,537]" box="[199,1108,151,177]" pageId="23" pageNumber="466">Coloration. Live coloration unknown. Color of preserved specimens light brown.</paragraph>
</subSubSection>
<subSubSection id="C36F654C9006FFD7FF27FF428B13AD4A" lastPageId="24" lastPageNumber="467" pageId="23" pageNumber="466" type="discussion">
<paragraph id="8BCA36C79006FFD8FF27FF428B4AAA23" blockId="23.[151,1437,151,537]" pageId="23" pageNumber="466">
<emphasis id="B901EAD59006FFD8FF27FF42880AA8EE" bold="true" box="[199,332,186,212]" pageId="23" pageNumber="466">Discussion.</emphasis>
<bibRefCitation id="EFE44B369006FFD8FEB2FF438B65A8EF" author="Teena, T. K. &amp; Murugan, A. &amp; Kumar, A. T. T. &amp; Lal, K. K." box="[338,547,187,213]" pageId="23" pageNumber="466" pagination="77 - 83" refId="ref20468" refString="Teena, T. K., Murugan, A., Kumar, A. T. T. &amp; Lal, K. K. (2021) Redescription of a rare cusk eel, Pycnocraspedum squamipinne (Actinopterygii, Ophidiiformes, Ophidiidae) from Bay of Bengal. Acta Ichthyologica et Piscatoria, 51 (1), 77-83. https://doi.org/10.3897/aiep.51.63469" type="journal article" year="2021">
Teena
<emphasis id="B901EAD59006FFD8FE7BFF438894A8EF" box="[411,466,187,213]" italics="true" pageId="23" pageNumber="466">et al.</emphasis>
(2021)
</bibRefCitation>
presented a re-description of
<taxonomicName id="4C754D449006FFD8FC8DFF438D51A8EE" authorityName="Alcock" authorityYear="1889" box="[877,1047,187,212]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="466" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59006FFD8FC8DFF438D51A8EE" box="[877,1047,187,212]" italics="true" pageId="23" pageNumber="466">P. squamipinne</emphasis>
</taxonomicName>
based on two of the
<specimenCount id="9D73FD4E9006FFD8FB1CFF438CDAA8EF" box="[1276,1436,187,213]" count="3" pageId="23" pageNumber="466" type="generic" typeStatus="syntype">three syntypes</specimenCount>
from the Bay of Bengal at ZSI (Kolkata) and 15 additional newly collected specimens off southern
<collectingCountry id="F36276579006FFD8FB3DFF278C5CA8C3" box="[1245,1306,223,249]" name="India" pageId="23" pageNumber="466">India</collectingCountry>
. The values given for the
<typeStatus id="54CE88659006FFD8FEC9FEFB88CDA927" box="[297,395,259,285]" pageId="23" pageNumber="466" type="syntype">syntypes</typeStatus>
(now
<typeStatus id="54CE88659006FFD8FE30FEFB8B32A927" box="[464,628,259,285]" pageId="23" pageNumber="466" type="paralectotype">paralectotypes</typeStatus>
) fall well into the range of the specimens studied by us, although they are larger at 270 and
<quantity id="4C8D9B229006FFD8FEBAFEDF88F1A97B" box="[346,439,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="2.77" pageId="23" pageNumber="466" unit="mm" value="277.0">277 mm</quantity>
SL than our specimens. The detailed figures of Teena
<emphasis id="B901EAD59006FFD8FBF8FEDF8D16A97B" box="[1048,1104,295,321]" italics="true" pageId="23" pageNumber="466">et al.</emphasis>
including the otolith of a
<quantity id="4C8D9B229006FFD8FA92FEDF8985A95F" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="466" unit="mm" value="200.0">200 mm</quantity>
SL specimen match our descriptions reasonably well. However, the maximal range given by them for meristics and morphometrics in their table 1 exceeds all values that we recognized for
<taxonomicName id="4C754D449006FFD8FC03FE978DCAA9B2" authorityName="Alcock" authorityYear="1889" box="[995,1164,367,392]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="466" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59006FFD8FC03FE978DCAA9B2" box="[995,1164,367,392]" italics="true" pageId="23" pageNumber="466">P. squamipinne</emphasis>
</taxonomicName>
. Possibly, the maximum value refers to their largest specimen of
<quantity id="4C8D9B229006FFD8FDB0FE6B8BEDA997" box="[592,683,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="3.8" pageId="23" pageNumber="466" unit="mm" value="380.0">380 mm</quantity>
SL, which is also of an unusual size for this relatively small species. The cause for this discrepancy is unknown, but one possible explanation could be that the specimen(s) in question represent another species of the genus, for instance
<taxonomicName id="4C754D449006FFD8FD36FE238A22A9CF" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[726,868,475,501]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="466" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD59006FFD8FD36FE238A22A9CF" box="[726,868,475,501]" italics="true" pageId="23" pageNumber="466">P. africanum</emphasis>
</taxonomicName>
. However, to resolve such speculation requires the review of the original specimens.
</paragraph>
<caption id="DF0A664F9006FFD8FF77F8E88C1EAF12" ID-DOI="http://doi.org/10.5281/zenodo.15561753" ID-Zenodo-Dep="15561753" box="[151,1368,1808,1832]" httpUri="https://zenodo.org/record/15561753/files/figure.png" pageId="23" pageNumber="466" startId="23.[151,250,1808,1832]" targetBox="[151,1435,561,1785]" targetPageId="23" targetType="figure">
<paragraph id="8BCA36C79006FFD8FF77F8E88C1EAF12" blockId="23.[151,1368,1808,1832]" box="[151,1368,1808,1832]" pageId="23" pageNumber="466">
<emphasis id="B901EAD59006FFD8FF77F8E88858AF12" bold="true" box="[151,286,1808,1832]" pageId="23" pageNumber="466">FIGURE 11.</emphasis>
<taxonomicName id="4C754D449006FFD8FEC5F8E88BADAF12" authority="Alcock, 1889" authorityName="Alcock" authorityYear="1889" box="[293,747,1808,1832]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="23" pageNumber="466" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59006FFD8FEC5F8E88B1CAF12" box="[293,602,1808,1832]" italics="true" pageId="23" pageNumber="466">Pycnocraspedum squamipinne</emphasis>
<bibRefCitation id="EFE44B369006FFD8FD80F8E88BADAF12" author="Alcock, A." box="[608,747,1808,1832]" pageId="23" pageNumber="466" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
</taxonomicName>
, lectotype, BMNH 1890.7.31.1, 20°17'30&quot;N 88°50'E, 353 m.
</paragraph>
</caption>
<caption id="DF0A664F9009FFD7FF77FC7B8CC5ABA1" ID-DOI="http://doi.org/10.5281/zenodo.15561755" ID-Zenodo-Dep="15561755" box="[151,1411,899,923]" httpUri="https://zenodo.org/record/15561755/files/figure.png" pageId="24" pageNumber="467" startId="24.[151,250,899,923]" targetBox="[153,1433,186,871]" targetPageId="24" targetType="figure">
<paragraph id="8BCA36C79009FFD7FF77FC7B8CC5ABA1" blockId="24.[151,1411,899,923]" box="[151,1411,899,923]" pageId="24" pageNumber="467">
<emphasis id="B901EAD59009FFD7FF77FC7B8866ABA1" bold="true" box="[151,288,899,923]" pageId="24" pageNumber="467">FIGURE 12.</emphasis>
Geographic distribution of the species of
<taxonomicName id="4C754D449009FFD7FD29FC7B8A31ABA1" authorityName="Alcock" authorityYear="1889" box="[713,887,899,923]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59009FFD7FD29FC7B8A31ABA1" box="[713,887,899,923]" italics="true" pageId="24" pageNumber="467">Pycnocraspedum</emphasis>
</taxonomicName>
. One symbol can represent more than one location.
</paragraph>
</caption>
<paragraph id="8BCA36C79009FFD7FF27FC168B13AD4A" blockId="24.[151,1437,1005,1607]" pageId="24" pageNumber="467">
As analyzed here,
<taxonomicName id="4C754D449009FFD7FE7EFC168B0DAC3D" authorityName="Alcock" authorityYear="1889" box="[414,587,1006,1031]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59009FFD7FE7EFC168B0DAC3D" box="[414,587,1006,1031]" italics="true" pageId="24" pageNumber="467">P. squamipinne</emphasis>
</taxonomicName>
shows a number of characters that distinguish it well from its congeners. These are the low number of vertebrae (4751 vs. 5159), the low number of anal-fin rays (5765 vs. 6879, except
<quantity id="4C8D9B229009FFD7FF77FBCD8997AC75" box="[151,209,1077,1103]" metricMagnitude="0" metricUnit="m" metricValue="1.6002" pageId="24" pageNumber="467" unit="in" value="63.0">63 in</quantity>
<taxonomicName id="4C754D449009FFD7FF39FBCE8804AC75" authorityName="Machida" authorityYear="1984" box="[217,322,1077,1103]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="fulvum">
<emphasis id="B901EAD59009FFD7FF39FBCE8804AC75" box="[217,322,1077,1103]" italics="true" pageId="24" pageNumber="467">P. fulvum</emphasis>
</taxonomicName>
), the low number of pseudobranchial filaments (12 vs. 47), the low number of transversal scale rows (5258) and scale rows above the lateral line (1317), the large head (HL in % of SL = 29.631.5 vs. 22.7 27.6), slender shape (body depth in % of SL = 1718 vs. 18.923.3), and the predorsal length in % of SL (2529 vs. 1725). The rear margin of the preopercle is relatively thin and weak with only one distinct spine (the second from below as described in the chapter “Key characters”;
<figureCitation id="134E2A429009FFD7FCF7FB3D8A1FACDA" box="[791,857,1221,1248]" captionStart="FIGURE 3" captionStartId="11.[151,250,1786,1810]" captionTargetBox="[256,1328,181,1748]" captionTargetId="figure-17@11.[256,1331,181,1762]" captionTargetPageId="11" captionText="FIGURE 3. Detail figures and drawings of preopercular spines and lower preopercular pores in Pycnocraspedum; preopercular spines are numbered from 1 through 4 from the lower angle of the preoperculum upwards; abbreviation lpp = lower preopercular pore. 3A3B) Pycnocraspedum armatum, USNM 227412, 3A1-3A2 specimen 1 left side, 3A3 specimen 1 right side (mirror imaged), 3B specimen 2; 3C3D) Pycnocraspedum phyllosoma, 3C USNM 421587, 3D USNM 421586; 3E) Pycnocraspedum fulvum (holotype), BSKU 27497; 3F3G) Pycnocraspedum africanum sp. nov., 3F holotype, MNHN 2000-5691, 3G paratype, MNHN 2000-5692; 3H) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021; 3I3K) Pycnocraspedum squamipinne, 3I lectotype, BMNH 1890.7.31.1, 3J ZMUC P771871, 3K ZMUC P771737." figureDoi="http://doi.org/10.5281/zenodo.15561739" httpUri="https://zenodo.org/record/15561739/files/figure.png" pageId="24" pageNumber="467">Fig. 3</figureCitation>
). Also, the otolith morphology is distinctly different from that of its congeners being characterized by a shallow, nearly flat dorsal rim, the inner face less bent than the outer face (vs. inner face strongly or more bent than outer face) and a relatively short sulcus (OL:CL = 1.51.6 vs. 1.31.45). Furthermore,
<taxonomicName id="4C754D449009FFD7FE4BFACA8B10AD71" authorityName="Alcock" authorityYear="1889" box="[427,598,1330,1355]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59009FFD7FE4BFACA8B10AD71" box="[427,598,1330,1355]" italics="true" pageId="24" pageNumber="467">P. squamipinne</emphasis>
</taxonomicName>
does not seem to grow to the sizes of its congeners (maximal verified size is SL
<quantity id="4C8D9B229009FFD7FF37FAAE8872AD4A" box="[215,308,1366,1392]" metricMagnitude="-1" metricUnit="m" metricValue="2.77" pageId="24" pageNumber="467" unit="mm" value="277.0">277 mm</quantity>
SL vs.
<quantity id="4C8D9B229009FFD7FE69FAAD8B58AD4A" box="[393,542,1365,1392]" metricMagnitude="-1" metricUnit="m" metricValue="3.825" metricValueMax="4.2" metricValueMin="3.45" pageId="24" pageNumber="467" unit="mm" value="382.5" valueMax="420.0" valueMin="345.0">345420 mm</quantity>
SL).
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9009FFD7FF27FA81899EAE7D" pageId="24" pageNumber="467" type="distribution">
<paragraph id="8BCA36C79009FFD7FF27FA81899EAE7D" blockId="24.[151,1437,1005,1607]" pageId="24" pageNumber="467">
<emphasis id="B901EAD59009FFD7FF27FA818827ADA9" bold="true" box="[199,353,1401,1427]" pageId="24" pageNumber="467">Distribution.</emphasis>
<taxonomicName id="4C754D449009FFD7FE88FA818BF9ADA9" authorityName="Alcock" authorityYear="1889" box="[360,703,1401,1427]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59009FFD7FE88FA818BF9ADA9" box="[360,703,1401,1427]" italics="true" pageId="24" pageNumber="467">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
has been mentioned across a wide distribution range in the Indo-West Pacific from East Africa to
<collectingCountry id="F36276579009FFD7FDE0FA668BF6AD8D" box="[512,688,1437,1464]" name="New Caledonia" pageId="24" pageNumber="467">New Caledonia</collectingCountry>
. However, in our study we found that all records from outside of the Bay of Bengal represent different species:
<taxonomicName id="4C754D449009FFD7FD95FA3A8A42ADE6" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[629,772,1474,1500]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD59009FFD7FD95FA3A8A42ADE6" box="[629,772,1474,1500]" italics="true" pageId="24" pageNumber="467">P. africanum</emphasis>
</taxonomicName>
in East Africa and mostly
<taxonomicName id="4C754D449009FFD7FBCFFA3A8DE9ADE1" authorityName=", Gosline" authorityYear="1954" box="[1071,1199,1474,1499]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD59009FFD7FBCFFA3A8DE9ADE1" box="[1071,1199,1474,1499]" italics="true" pageId="24" pageNumber="467">P. armatum</emphasis>
</taxonomicName>
elsewhere outside of the Bay of Bengal. It thus appears that
<taxonomicName id="4C754D449009FFD7FDA7FA1E8BB7ADC5" authorityName="Alcock" authorityYear="1889" box="[583,753,1510,1535]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59009FFD7FDA7FA1E8BB7ADC5" box="[583,753,1510,1535]" italics="true" pageId="24" pageNumber="467">P. squamipinne</emphasis>
</taxonomicName>
is endemic to the Bay of Bengal. Subject to review of certain specimens mentioned by
<bibRefCitation id="EFE44B369009FFD7FE54F9F28BCFAE1E" author="Teena, T. K. &amp; Murugan, A. &amp; Kumar, A. T. T. &amp; Lal, K. K." box="[436,649,1545,1572]" pageId="24" pageNumber="467" pagination="77 - 83" refId="ref20468" refString="Teena, T. K., Murugan, A., Kumar, A. T. T. &amp; Lal, K. K. (2021) Redescription of a rare cusk eel, Pycnocraspedum squamipinne (Actinopterygii, Ophidiiformes, Ophidiidae) from Bay of Bengal. Acta Ichthyologica et Piscatoria, 51 (1), 77-83. https://doi.org/10.3897/aiep.51.63469" type="journal article" year="2021">
Teena
<emphasis id="B901EAD59009FFD7FE1EF9F28B71AE19" box="[510,567,1545,1571]" italics="true" pageId="24" pageNumber="467">et al.</emphasis>
(2021)
</bibRefCitation>
it is possible that a second species of
<taxonomicName id="4C754D449009FFD7FBD6F9F18DB1AE19" authorityName="Alcock" authorityYear="1889" box="[1078,1271,1545,1571]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59009FFD7FBD6F9F18DB1AE19" box="[1078,1271,1545,1571]" italics="true" pageId="24" pageNumber="467">Pycnocraspedum</emphasis>
</taxonomicName>
may occur off
<collectingCountry id="F36276579009FFD7FF77F9D58992AE7D" box="[151,212,1581,1607]" name="India" pageId="24" pageNumber="467">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,181 @@
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<mods:title id="2E19D434847EB6CDAB082FBC13674174">Revision of the bathydemersal fish genus Pycnocraspedum (Ophidiidae; Teleostei), with description of two new species</mods:title>
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<subSubSection id="C36F654C9009FFD7FF77F9618A4AAE89" box="[151,780,1689,1716]" pageId="24" pageNumber="467" type="nomenclature">
<paragraph id="8BCA36C79009FFD7FF77F9618A4AAE89" blockId="24.[151,780,1689,1752]" box="[151,780,1689,1716]" pageId="24" pageNumber="467">
<heading id="D08281AB9009FFD7FF77F9618A4AAE89" bold="true" box="[151,780,1689,1716]" fontSize="11" level="1" pageId="24" pageNumber="467" reason="1">
<emphasis id="B901EAD59009FFD7FF77F9618A4AAE89" bold="true" box="[151,780,1689,1716]" pageId="24" pageNumber="467">
<taxonomicName id="4C754D449009FFD7FF46F9618A4AAE89" authority="(Schwarzhans, 1978)" baseAuthorityName="Schwarzhans" baseAuthorityYear="1978" box="[166,780,1689,1716]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="cetonaense">
<emphasis id="B901EAD59009FFD7FF46F9618B44AE8E" bold="true" box="[166,514,1689,1716]" italics="true" pageId="24" pageNumber="467">Pycnocraspedum cetonaense</emphasis>
(
<bibRefCitation id="EFE44B369009FFD7FDF1F9618A42AE89" author="Schwarzhans, W." box="[529,772,1689,1715]" pageId="24" pageNumber="467" pagination="1 - 52" refId="ref19902" refString="Schwarzhans, W. (1978) Otolithen aus dem Unter-Pliozan von Sud-Sizilien und aus der Toscana. Berliner Geowissenschaftliche Abhandlungen, 8, 1-52." type="journal article" year="1978">Schwarzhans, 1978</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9009FFD7FF77F94589B7AEE2" box="[151,241,1725,1752]" pageId="24" pageNumber="467" type="description">
<paragraph id="8BCA36C79009FFD7FF77F94589B7AEE2" blockId="24.[151,780,1689,1752]" box="[151,241,1725,1752]" pageId="24" pageNumber="467">
<figureCitation id="134E2A429009FFD7FF77F94589B7AEE2" box="[151,241,1725,1752]" captionStart="FIGURE 5" captionStartId="14.[151,250,1750,1774]" captionTargetBox="[245,1358,193,1715]" captionTargetId="figure-23@14.[222,1366,181,1726]" captionTargetPageId="14" captionText="FIGURE 5. Otoliths, inner faces (if not noted otherwise) shown from right side. 5A5C) Pycnocraspedum africanum sp. nov., 5A holotype, MNHN 2000-5691 (5A2 dorsal view), 5B paratype MNHN 2000-5692, 5C tentatively assigned specimen, IRSNB (5C2 dorsal view, 5C3 anterior view); 5D5E) Pycnocraspedum armatum, 5D USNM 227412 (5D2 ventral view), 5E USNM 227390 (5E2 ventral view); 5F) Pycnocraspedum fulvum (holotype), BSKU 27497 (5F2 ventral view, 5F3 anterior view); 5G) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021 (5G2 ventral view); 5H) Pycnocraspedum phyllosoma, USNM 421587 (5H2 ventral view, 5H3 anterior view); 5I5K) Pycnocraspedum squamipinne, 5I lectotype, BMNH 1890.7.31.1 (5I2 ventral view), 5J ZMUC P771871, 5L ZMUC P771739 (5L2 ventral view), 5K ZMUC P771738 (5K2 ventral view); 5M) †Pycnocraspedum cetonaense (holotype) SMF P.5694 (5M2 dorsal view, 5M3 anterior view)." figureDoi="http://doi.org/10.5281/zenodo.15561743" httpUri="https://zenodo.org/record/15561743/files/figure.png" pageId="24" pageNumber="467">Fig. 5M</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9009FFD7FF77F8FA8D4EAF4C" pageId="24" pageNumber="467" type="reference_group">
<paragraph id="8BCA36C79009FFD7FF77F8FA8A1BAF20" blockId="24.[151,1032,1794,1910]" box="[151,861,1794,1818]" pageId="24" pageNumber="467">
<treatmentCitationGroup id="AB6511E99009FFD7FF77F8FA8A1BAF20" box="[151,861,1794,1818]" pageId="24" pageNumber="467">
<emphasis id="B901EAD59009FFD7FF77F8FA8866AF20" box="[151,288,1794,1818]" italics="true" pageId="24" pageNumber="467">Brotulidarum</emphasis>
<treatmentCitation id="0AD410D69009FFD7FEC7F8FB8BF1AF20" author="Schwarzhans, W." box="[295,695,1794,1818]" page="38" pageId="24" pageNumber="467" year="1978">
<taxonomicName id="4C754D449009FFD7FEC7F8FB8BF1AF20" authority="Schwarzhans, 1978: 37 - 38" authorityName="Schwarzhans" authorityPageNumber="37 - 38" authorityYear="1978" box="[295,695,1794,1818]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="cetonaensis">
<emphasis id="B901EAD59009FFD7FEC7F8FB88DBAF20" box="[295,413,1795,1818]" italics="true" pageId="24" pageNumber="467">cetonaensis</emphasis>
<bibRefCitation id="EFE44B369009FFD7FE43F8FA8BF1AF20" author="Schwarzhans, W." box="[419,695,1794,1818]" pageId="24" pageNumber="467" pagination="1 - 52" refId="ref19902" refString="Schwarzhans, W. (1978) Otolithen aus dem Unter-Pliozan von Sud-Sizilien und aus der Toscana. Berliner Geowissenschaftliche Abhandlungen, 8, 1-52." type="journal article" year="1978">Schwarzhans, 1978: 3738</bibRefCitation>
</taxonomicName>
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, pl. 10, fig. 115.
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</paragraph>
<paragraph id="8BCA36C79009FFD7FF77F8D98A3FAF03" blockId="24.[151,1032,1794,1910]" box="[151,889,1825,1849]" pageId="24" pageNumber="467">
<emphasis id="B901EAD59009FFD7FF40F8D988CBAF03" box="[160,397,1825,1849]" italics="true" pageId="24" pageNumber="467">
genus
<taxonomicName id="4C754D449009FFD7FF01F8D98BB6AF03" authority="(Schwarzhans, 1978)" authorityName="Nolf" authorityYear="1980" baseAuthorityName="Schwarzhans" baseAuthorityYear="1978" box="[225,752,1825,1849]" family="Ophidiidae" genus="Neobythitinorum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="genus">
Neobythitinorum
<emphasis id="B901EAD59009FFD7FE7AF8D98B56AF02" box="[410,528,1825,1848]" italics="true" pageId="24" pageNumber="467">cetonaensis</emphasis>
(
<bibRefCitation id="EFE44B369009FFD7FDFDF8D98BAEAF03" author="Schwarzhans, W." box="[541,744,1825,1849]" pageId="24" pageNumber="467" pagination="1 - 52" refId="ref19902" refString="Schwarzhans, W. (1978) Otolithen aus dem Unter-Pliozan von Sud-Sizilien und aus der Toscana. Berliner Geowissenschaftliche Abhandlungen, 8, 1-52." type="journal article" year="1978">Schwarzhans, 1978</bibRefCitation>
)
</taxonomicName>
</emphasis>
:
<bibRefCitation id="EFE44B369009FFD7FD19F8D98A33AF03" author="Nolf, D." box="[761,885,1825,1849]" pageId="24" pageNumber="467" pagination="71 - 195" refId="ref19421" refString="Nolf, D. (1980) Etude monographique des Otolithes des Ophidiiformes actuels et Revision des Especes fossils (Pisces, Teleostei). Mededelingen van de Werkgroep voor Tertiaire an Kwartaire Geologie, 17 (2), 71-195." type="journal article" year="1980">Nolf (1980)</bibRefCitation>
.
</paragraph>
<paragraph id="8BCA36C79009FFD7FF77F8B88A30AF6D" blockId="24.[151,1032,1794,1910]" box="[151,886,1855,1879]" pageId="24" pageNumber="467">
<taxonomicName id="4C754D449009FFD7FF77F8B88BD0AF6D" authority="(Schwarzhans, 1978)" baseAuthorityName="Schwarzhans" baseAuthorityYear="1978" box="[151,662,1855,1879]" family="Ophidiidae" genus="Paragenypterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="cetonaensis">
<emphasis id="B901EAD59009FFD7FF77F8B888F3AF6D" box="[151,437,1856,1879]" italics="true" pageId="24" pageNumber="467">Paragenypterus cetonaensis</emphasis>
(
<bibRefCitation id="EFE44B369009FFD7FE23F8C78BC8AF6D" author="Schwarzhans, W." box="[451,654,1855,1879]" pageId="24" pageNumber="467" pagination="1 - 52" refId="ref19902" refString="Schwarzhans, W. (1978) Otolithen aus dem Unter-Pliozan von Sud-Sizilien und aus der Toscana. Berliner Geowissenschaftliche Abhandlungen, 8, 1-52." type="journal article" year="1978">Schwarzhans, 1978</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="EFE44B369009FFD7FD7FF8C78A34AF6D" author="Schwarzhans, W." box="[671,882,1855,1879]" pageId="24" pageNumber="467" pagination="63 - 122" refId="ref19930" refString="Schwarzhans, W. (1981) Vergleichende morphologische Untersuchungen an rezenten und fossilen Otolithen der Ordnung Ophidiiformes. Berliner geowissenschaftliche Abhandlungen, A, 32, 63-122." type="journal article" year="1981">Schwarzhans (1981)</bibRefCitation>
.
</paragraph>
<paragraph id="8BCA36C79009FFD7FF77F8A68D4EAF4C" blockId="24.[151,1032,1794,1910]" box="[151,1032,1886,1910]" pageId="24" pageNumber="467">
<taxonomicName id="4C754D449009FFD7FF77F8A68BE4AF4C" authority="(Schwarzhans, 1978)" baseAuthorityName="Schwarzhans" baseAuthorityYear="1978" box="[151,674,1886,1910]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="24" pageNumber="467" phylum="Chordata" rank="species" species="cetonaensis">
<emphasis id="B901EAD59009FFD7FF77F8A68884AF4C" box="[151,450,1886,1910]" italics="true" pageId="24" pageNumber="467">Pycnocraspedum cetonaensis</emphasis>
(
<bibRefCitation id="EFE44B369009FFD7FE2FF8A68BDCAF4C" author="Schwarzhans, W." box="[463,666,1886,1910]" pageId="24" pageNumber="467" pagination="1 - 52" refId="ref19902" refString="Schwarzhans, W. (1978) Otolithen aus dem Unter-Pliozan von Sud-Sizilien und aus der Toscana. Berliner Geowissenschaftliche Abhandlungen, 8, 1-52." type="journal article" year="1978">Schwarzhans, 1978</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="EFE44B369009FFD7FD4CF8A68A39AF4C" author="Schwarzhans, W." box="[684,895,1886,1910]" pageId="24" pageNumber="467" pagination="71 - 98" refId="ref19961" refString="Schwarzhans, W. (1994) Sexual and ontogenetic dimorphism in otoliths of the family Ophidiidae. Cybium, 18, 71-98." type="journal article" year="1994">Schwarzhans (1994)</bibRefCitation>
;
<bibRefCitation id="EFE44B369009FFD7FC69F8A68D42AF4C" author="Nolf, D." box="[905,1028,1886,1910]" pageId="24" pageNumber="467" refId="ref19465" refString="Nolf, D. (2013) The Diversity of Fish Otoliths, Past and Present. The Royal Belgian Institute of Natural Sciences, Brussels, 359 pp." type="book" year="2013">Nolf (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9009FFD7FF77F85C88E8AFD8" pageId="24" pageNumber="467" type="materials_examined">
<paragraph id="8BCA36C79009FFD7FF77F85C88E8AFD8" blockId="24.[151,1437,1956,2019]" pageId="24" pageNumber="467">
<emphasis id="B901EAD59009FFD7FF77F85C883AAF84" bold="true" box="[151,380,1956,1982]" pageId="24" pageNumber="467">Material examined</emphasis>
(
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):
<materialsCitation id="3B1D3C9A9009FFD7FDCAF85C88ECAFD8" collectionCode="SMF" country="Italy" county="Early Pliocene" location="Cetona near Siena" municipality="Zanclean" pageId="24" pageNumber="467" specimenCode="P.5694" specimenCount="1" typeStatus="holotype">
The unique
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,
<collectionCode id="ED64AE029009FFD7FCC8F85C8A26AF84" box="[808,864,1956,1982]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="467">SMF</collectionCode>
<specimenCode id="DBD39EBC9009FFD7FC89F85C8AF1AF85" box="[873,951,1956,1983]" pageId="24" pageNumber="467">P.5694</specimenCode>
,
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,
<collectingCountry id="F36276579009FFD7FB46F85C8D9CAF84" box="[1190,1242,1956,1982]" name="Italy" pageId="24" pageNumber="467">Italy</collectingCountry>
,
<collectingMunicipality id="6BAEACBD9009FFD7FB08F85C8C14AF84" box="[1256,1362,1956,1982]" pageId="24" pageNumber="467">Zanclean</collectingMunicipality>
,
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, a fossil otolith
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9008FFD6FF27FF6E8A93A97B" pageId="25" pageNumber="468" type="discussion">
<paragraph id="8BCA36C79008FFD6FF27FF6E8A93A97B" blockId="25.[151,1436,150,321]" pageId="25" pageNumber="468">
<emphasis id="B901EAD59008FFD6FF27FF6E880AA88A" bold="true" box="[199,332,150,176]" pageId="25" pageNumber="468">Discussion.</emphasis>
<taxonomicName id="4C754D449008FFD6FEB5FF6F8BDDA88A" baseAuthorityName="Schwarzhans" baseAuthorityYear="1978" box="[341,667,151,177]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="25" pageNumber="468" phylum="Chordata" rank="species" species="cetonaense">
<emphasis id="B901EAD59008FFD6FEB5FF6F8BDDA88A" box="[341,667,151,177]" italics="true" pageId="25" pageNumber="468">Pycnocraspedum cetonaense</emphasis>
</taxonomicName>
was recovered from the Early Pliocene (about 5 mya) of
<collectingCountry id="F36276579008FFD6FAD7FF6F8C2CA88B" box="[1335,1386,151,177]" name="Italy" pageId="25" pageNumber="468">Italy</collectingCountry>
and represents the only known fossil record of the genus, based on a single large otolith. The otolith resembles in shape and proportions those of the extant
<taxonomicName id="4C754D449008FFD6FDC2FF278BF7A8C3" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[546,689,223,249]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="25" pageNumber="468" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD59008FFD6FDC2FF278BF7A8C3" box="[546,689,223,249]" italics="true" pageId="25" pageNumber="468">P. africanum</emphasis>
</taxonomicName>
but differs in the relatively wide sulcus and the presence of clearly separated ostial and caudal colliculi (vs. single, fused colliculum). This record also demonstrates that the genus
<taxonomicName id="4C754D449008FFD6FF77FEDF881EA97B" authorityName="Alcock" authorityYear="1889" box="[151,344,295,321]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="25" pageNumber="468" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59008FFD6FF77FEDF881EA97B" box="[151,344,295,321]" italics="true" pageId="25" pageNumber="468">Pycnocraspedum</emphasis>
</taxonomicName>
was even more widely distributed in the past than today.
</paragraph>
</subSubSection>
</treatment>
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View file

@ -1,70 +1,70 @@
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<emphasis id="B901EAD59016FFC8FF77F8838B2AAFAF" bold="true" box="[151,620,1914,1941]" pageId="7" pageNumber="450">
Key to the species of
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<taxonomicName id="4C754D449016FFC8FE7DF8828B2AAFAF" ID-CoL="75NT" authority="Alcock, 1889" authorityName="Alcock" authorityYear="1889" box="[413,620,1914,1941]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FE7DF8828B2AAFAF" bold="true" box="[413,620,1914,1941]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
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</emphasis>
@ -74,9 +74,9 @@ Key to the species of
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<paragraph id="8BCA36C79016FFC8FF77F8448B89AFC0" blockId="7.[151,1436,1980,2042]" pageId="7" pageNumber="450">
(see also
<tableCitation id="C6F7037C9016FFC8FF1EF8448816AFEC" box="[254,336,1980,2006]" captionStart="TABLE 2" captionStartId="8.[152,244,720,744]" captionText="TABLE 2. Meristic, morphometric and otolith characters of the species of the genus Pycnocraspedum. For abbreviations" pageId="7" pageNumber="450">Table 2</tableCitation>
<tableCitation id="C6F7037C9016FFC8FF1EF8448816AFEC" box="[254,336,1980,2006]" captionStart="TABLE 2" captionStartId="8.[152,244,720,744]" captionTargetPageId="8" captionText="TABLE 2. Meristic, morphometric and otolith characters of the species of the genus Pycnocraspedum. For abbreviations" httpUri="http://table.plazi.org/id/DF0A664F9019FFC7FF78FD288B4AAB36" pageId="7" pageNumber="450" tableUuid="DF0A664F9019FFC7FF78FD288B4AAB36">Table 2</tableCitation>
for the complete set of measurements and
<tableCitation id="C6F7037C9016FFC8FCCBF8448A3BAFEC" box="[811,893,1980,2006]" captionStart="TABLE 3" captionStartId="9.[152,244,964,988]" captionText="TABLE 3. Ranges of selected key counts and measurements of the species of the genus Pycnocraspedum. D/V = first dorsal-fin ray above vertebra. Colors indicate species for ease of comparison." pageId="7" pageNumber="450">Table 3</tableCitation>
<tableCitation id="C6F7037C9016FFC8FCCBF8448A3BAFEC" box="[811,893,1980,2006]" captionStart="TABLE 3" captionStartId="9.[152,244,964,988]" captionTargetBox="[151,1436,1044,2008]" captionTargetId="figure-207@9.[151,1436,1046,1970]" captionTargetPageId="9" captionText="TABLE 3. Ranges of selected key counts and measurements of the species of the genus Pycnocraspedum. D/V = first dorsal-fin ray above vertebra. Colors indicate species for ease of comparison." pageId="7" pageNumber="450">Table 3</tableCitation>
for measurement and meristic ranges of selected traits; data for
<taxonomicName id="4C754D449016FFC8FEDBF819888AAFC0" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[315,460,2016,2042]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD59016FFC8FEDBF819888AAFC0" box="[315,460,2016,2042]" italics="true" pageId="7" pageNumber="450">P. microlepis</emphasis>

View file

@ -0,0 +1,446 @@
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<heading id="D08281AB9017FFC9FF77F9A28B25AE4E" bold="true" box="[151,611,1626,1653]" fontSize="11" level="1" pageId="6" pageNumber="449" reason="1">
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Genus
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<emphasis id="B901EAD59017FFC9FF0EF9A288FBAE4F" bold="true" box="[238,445,1626,1653]" italics="true" pageId="6" pageNumber="449">Pycnocraspedum</emphasis>
<bibRefCitation id="EFE44B369017FFC9FE23F9A28B25AE4E" author="Alcock, A." box="[451,611,1626,1652]" pageId="6" pageNumber="449" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
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<taxonomicName id="4C754D449017FFC9FF77F9678890AE8D" ID-CoL="75NT" authority="Alcock, 1889" authorityName="Alcock" authorityYear="1889" box="[151,470,1695,1719]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="449" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59017FFC9FF77F9678803AE8D" box="[151,325,1695,1719]" italics="true" pageId="6" pageNumber="449">Pycnocraspedum</emphasis>
<bibRefCitation id="EFE44B369017FFC9FEABF9678890AE8D" author="Alcock, A." box="[331,470,1695,1719]" pageId="6" pageNumber="449" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
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:
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species by monotypy
<taxonomicName id="4C754D449017FFC9FD0FF9678DF0AE8D" ID-CoL="4QM83" authority="Alcock, 1889" authorityName="Alcock" authorityYear="1889" box="[751,1206,1695,1719]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="449" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59017FFC9FD0FF9678D62AE8D" box="[751,1060,1695,1719]" italics="true" pageId="6" pageNumber="449">Pycnocraspedum squamipinne</emphasis>
<bibRefCitation id="EFE44B369017FFC9FBCAF9678DF0AE8D" author="Alcock, A." box="[1066,1206,1695,1719]" pageId="6" pageNumber="449" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
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.
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<paragraph id="8BCA36C79017FFC9FF77F9468D4BAEEC" blockId="6.[151,1437,1695,1811]" box="[151,1037,1726,1750]" pageId="6" pageNumber="449">
<taxonomicName id="4C754D449017FFC9FF77F94688CAAEEC" ID-CoL="D54S4" authority="Matsubara, 1943" authorityName="Matsubara" authorityYear="1943" box="[151,396,1726,1750]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="449" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59017FFC9FF77F946899EAEEF" box="[151,216,1726,1749]" italics="true" pageId="6" pageNumber="449">Itatius</emphasis>
<bibRefCitation id="EFE44B369017FFC9FF3EF94688CAAEEC" author="Matsubara, K." box="[222,396,1726,1750]" pageId="6" pageNumber="449" pagination="37 - 81" refId="ref18682" refString="Matsubara, K. (1943) Ichthyological annotations from the depths of the sea Japan, I - VII. Journal of Sigenkagaku Kenkyusyo, 1, 37-81." type="journal article" year="1943">Matsubara, 1943</bibRefCitation>
</taxonomicName>
:
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species by monotypy
<taxonomicName id="4C754D449017FFC9FD44F9468D4FAEEC" ID-CoL="3QDWT" authority="Matsubara, 1943" authorityName="Matsubara" authorityYear="1943" box="[676,1033,1726,1750]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="449" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD59017FFC9FD44F9468A13AEEC" box="[676,853,1726,1750]" italics="true" pageId="6" pageNumber="449">Itatius microlepis</emphasis>
<bibRefCitation id="EFE44B369017FFC9FCBBF9468D4FAEEC" author="Matsubara, K." box="[859,1033,1726,1750]" pageId="6" pageNumber="449" pagination="37 - 81" refId="ref18682" refString="Matsubara, K. (1943) Ichthyological annotations from the depths of the sea Japan, I - VII. Journal of Sigenkagaku Kenkyusyo, 1, 37-81." type="journal article" year="1943">Matsubara, 1943</bibRefCitation>
</taxonomicName>
.
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<paragraph id="8BCA36C79017FFC9FF77F9258801AF29" blockId="6.[151,1437,1695,1811]" pageId="6" pageNumber="449">
<taxonomicName id="4C754D449017FFC9FF77F9258B4EAECF" authority="Schwarzhans, 1981" authorityName="Schwarzhans" authorityYear="1981" box="[151,520,1756,1781]" family="Ophidiidae" genus="Paragenypterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="449" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59017FFC9FF77F925887FAECE" box="[151,313,1757,1780]" italics="true" pageId="6" pageNumber="449">Paragenypterus</emphasis>
<bibRefCitation id="EFE44B369017FFC9FEA0F9248B4EAECF" author="Schwarzhans, W." box="[320,520,1756,1781]" pageId="6" pageNumber="449" pagination="63 - 122" refId="ref19930" refString="Schwarzhans, W. (1981) Vergleichende morphologische Untersuchungen an rezenten und fossilen Otolithen der Ordnung Ophidiiformes. Berliner geowissenschaftliche Abhandlungen, A, 32, 63-122." type="journal article" year="1981">Schwarzhans, 1981</bibRefCitation>
</taxonomicName>
:
<typeStatus id="54CE88659017FFC9FDF3F9258B79AECF" box="[531,575,1757,1781]" pageId="6" pageNumber="449">type</typeStatus>
species by monotypy
<emphasis id="B901EAD59017FFC9FCC3F9248D70AECE" box="[803,1078,1756,1780]" italics="true" pageId="6" pageNumber="449">Brotulidarum centonaensis</emphasis>
Schwarzans, 1978, a fossil otolith-based taxon.
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<emphasis id="B901EAD59017FFC9FF77F8B9884CAF61" bold="true" box="[151,266,1857,1883]" pageId="6" pageNumber="449">Diagnosis</emphasis>
(adopted and altered from
<bibRefCitation id="EFE44B369017FFC9FDA4F8B98A67AF61" author="Nielsen, J. G. &amp; Cohen, D. M. &amp; Markle, D. F. &amp; Robins, C. R." box="[580,801,1857,1883]" pageId="6" pageNumber="449" refId="ref19264" refString="Nielsen, J. G., Cohen, D. M., Markle, D. F. &amp; Robins, C. R. (1999) An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiiform fishes known to date. FAO Fisheries Synopsis No. 125. Vol. 18. FAO, Rome, 178 pp." type="book" year="1999">
Nielsen
<emphasis id="B901EAD59017FFC9FD43F8BA8B9BAF61" box="[675,733,1857,1883]" italics="true" pageId="6" pageNumber="449">et al.</emphasis>
1999
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). A genus of the subfamily
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characterized by the following combination of characters: Body short with large head; upper jaw ending behind eye; hind margin of preopercle with 1 to 4, spines; 2 median basibranchial tooth patches, sometimes accompanied by additional small single or pair of lateral patches; very small, closely packed teeth resembling sandpaper surface on dentary, premaxilla, vomer and palatines; 4 to 6 developed gill rakers on first gill arch; 23 lower preopercular pores; no upper preopercular pore; 1 to 7 small pseudobranchial filaments; pectoral-fin rays 24 to 28; precaudal vertebrae 11 to 14 (mostly 12 to 13); 0 to 5 dorsal-fin rays in front of first vertebra; otoliths slender with long, uniform sulcus. Head and body completely covered with small cycloid scales, including bases of dorsal, anal and pectoral fins.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C9016FFC8FF27FEFA8A6DAA93" pageId="7" pageNumber="450" type="description">
<paragraph id="8BCA36C79016FFC8FF27FEFA8861A95F" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="450">
<emphasis id="B901EAD59016FFC8FF27FEFA88F3A926" bold="true" box="[199,437,258,284]" pageId="7" pageNumber="450">General description</emphasis>
: A number of characters occur in all species of
<taxonomicName id="4C754D449016FFC8FC32FEFB8DD5A927" authorityName="Alcock" authorityYear="1889" box="[978,1171,259,285]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FC32FEFB8DD5A927" box="[978,1171,259,285]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
in the same manner. In order to reduce redundancy, the following description summarizes those characters that are valid in all the species here studied.
</paragraph>
<paragraph id="8BCA36C79016FFC8FF27FE978899AA07" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="450">Head stout, with straight, inclined dorsal profile. Opercle with sharp, slightly extruding spine. Eye moderately small. Maxilla extending far behind eye, strongly widened posteriorly upward and downward from axis and with a vertical posterior end. Narrow, poorly defined supramaxilla. Pores: 1 anterior supraorbital, no posterior supraorbital, 3 anterior infraorbital, 3 posterior infraorbital, 3 anterior mandibular, 3 posterior mandibular, 23 lower preopercular. Head completely covered with scales, including opercle, preopercle, cheeks, and maxilla. Scales on head larger than scales on body and peduncle.</paragraph>
<paragraph id="8BCA36C79016FFC8FF27FDBF8A6DAA93" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="450">Dentition. Minute sandpaper-like granular teeth on broad dentary and premaxillary bands, narrow palatine bands, and a narrow, boomerang-shaped vomerine patch. Basibranchial tooth patches 2 long median and 1 small lateral patch on left or right side, rarely a symmetrical pair.</paragraph>
</subSubSection>
<subSubSection id="C36F654C9016FFC8FF27FD4A8C6EAF13" pageId="7" pageNumber="450" type="discussion">
<paragraph id="8BCA36C79016FFC8FF27FD4A8CCCAC47" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="450">
<emphasis id="B901EAD59016FFC8FF27FD4A8809AAF6" bold="true" box="[199,335,690,716]" pageId="7" pageNumber="450">Discussion:</emphasis>
Following the synonymization of the genera
<taxonomicName id="4C754D449016FFC8FC60FD4B8DA4AAF7" authority="Alcock, 1890" authorityName="Alcock" authorityYear="1890" box="[896,1250,691,717]" family="Ophidiidae" genus="Tauredophidium" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FC60FD4B8D71AAF7" box="[896,1079,691,717]" italics="true" pageId="7" pageNumber="450">Tauredophidium</emphasis>
<bibRefCitation id="EFE44B369016FFC8FBA4FD4B8DA4AAF7" author="Alcock, A." box="[1092,1250,691,717]" pageId="7" pageNumber="450" pagination="197 - 222" refId="ref17546" refString="Alcock, A. (1890) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 16. On the bathybial fishes collected in the Bay of Bengal during the season 1889 - 1890. Annals Magazine Natural History, Series 6, 6 (33), 197-222. https://doi.org/10.1080/00222939008694027" type="journal article" year="1890">Alcock, 1890</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C754D449016FFC8FAC8FD4B8861AACB" authority="Cohen, 1961" authorityName="Cohen" authorityYear="1961" family="Ophidiidae" genus="Xyelacyba" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FAC8FD4B8CDBAAF7" box="[1320,1437,691,717]" italics="true" pageId="7" pageNumber="450">Xyelacyba</emphasis>
<bibRefCitation id="EFE44B369016FFC8FF77FD2F8861AACB" author="Cohen, D. M." box="[151,295,727,753]" pageId="7" pageNumber="450" pagination="288 - 292" refId="ref17823" refString="Cohen, D. M. (1961) A new genus and species of deepwater ophidiid fish from the Gulf of Mexico. Copeia, 288-292. https://doi.org/10.2307/1439802" type="book chapter" year="1961">Cohen, 1961</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="4C754D449016FFC8FE89FD2F8BD1AACB" authority="Gunther, 1878" authorityName="Gunther" authorityYear="1878" box="[361,663,727,753]" family="Ophidiidae" genus="Acanthonus" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FE89FD2F88ABAACB" box="[361,493,727,753]" italics="true" pageId="7" pageNumber="450">Acanthonus</emphasis>
<bibRefCitation id="EFE44B369016FFC8FE16FD2F8BD1AACB" author="Gunther, A." box="[502,663,727,753]" pageId="7" pageNumber="450" pagination="17 - 28" refId="ref18522" refString="Gunther, A. (1878) Preliminary notices of deep-sea fishes collected during the voyage of H. M. S. Challenger. Annals Magazine Natural History, Series 5, 2 (7), 17-28. https://doi.org/10.1080/00222937808682376" type="journal article" year="1878">Günther, 1878</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EFE44B369016FFC8FD24FD2F8AE7AACB" author="Girard, M. G. &amp; Nonaka, A. &amp; Baldwin, C. C. &amp; Johnson, G. D." box="[708,929,727,753]" pageId="7" pageNumber="450" pagination="20 - 42" refId="ref18326" refString="Girard, M. G., Nonaka, A., Baldwin, C. C. &amp; Johnson, G. D. (2024) Discovery and description of elaborate larval cusk-eels and the relationships among Acanthonus, Tauredophidium and Xyelacyba (Teleostei: Ophidiidae). In: Leis, J. M., Watson, W., Mundy, B. C. &amp; Konstantinidis, P. (Eds.), Early Life History and Biology of Marine Fishes: Research inspired by the work of H. Geoffrey Moser. NOAA Professional Paper NMFS, 24, pp. 20-42. https://doi.org/10.7755/PP.24.3" type="journal article" year="2024">
Girard
<emphasis id="B901EAD59016FFC8FCF5FD2F8A08AACB" box="[789,846,727,753]" italics="true" pageId="7" pageNumber="450">et al.</emphasis>
(2024)
</bibRefCitation>
and the establishment of
<emphasis id="B901EAD59016FFC8FB25FD2F8CDAAACB" box="[1221,1436,727,753]" italics="true" pageId="7" pageNumber="450">Tenuicephalichthys</emphasis>
<bibRefCitation id="EFE44B369016FFC8FF77FD0388AEAB2F" author="Schwarzhans, W. &amp; Moller, P. R." box="[151,488,763,789]" pageId="7" pageNumber="450" pagination="600" refId="ref20052" refString="Schwarzhans, W. &amp; Moller, P. R. (2024) Erratum to: Schwarzhans, W. &amp; Moller, P. R. (2021) Revision of the ' dragon-head' cusk eels of the genus Porogadus (Teleostei: Ophidiidae), with description of eight new species and one new genus. Zootaxa, 5492 (4), 600. https://doi.org/10.11646/zootaxa.5492.4.9" type="journal article" year="2024">Schwarzhans &amp; Møller 2024</bibRefCitation>
, the subfamily
<taxonomicName id="4C754D449016FFC8FD7EFD038A7BAB2F" box="[670,829,763,789]" class="Teleostei" family="Ophidiidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="subFamily" subFamily="Neobythitinae">Neobythitinae</taxonomicName>
now comprises 37 genera. The interrelationships of many of the neobythitine genera have remained elusive.
<bibRefCitation id="EFE44B369016FFC8FCC3FCE78AABAB03" author="Matsubara, K." box="[803,1005,799,825]" pageId="7" pageNumber="450" pagination="37 - 81" refId="ref18682" refString="Matsubara, K. (1943) Ichthyological annotations from the depths of the sea Japan, I - VII. Journal of Sigenkagaku Kenkyusyo, 1, 37-81." type="journal article" year="1943">Matsubara (1943)</bibRefCitation>
found
<taxonomicName id="4C754D449016FFC8FBA1FCE78DCFAB02" authorityName="Matsubara" authorityYear="1943" box="[1089,1161,799,824]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FBA1FCE78DCFAB02" box="[1089,1161,799,824]" italics="true" pageId="7" pageNumber="450">Itatius</emphasis>
</taxonomicName>
(syn.
<taxonomicName id="4C754D449016FFC8FB32FCE78CD5AB03" authorityName="Alcock" authorityYear="1889" box="[1234,1427,799,825]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FB32FCE78CD5AB03" box="[1234,1427,799,825]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
) resembling
<taxonomicName id="4C754D449016FFC8FEFFFCBB88E5AB67" authorityName="Goode &amp; Bean" authorityYear="1885" box="[287,419,835,861]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FEFFFCBB88E5AB67" box="[287,419,835,861]" italics="true" pageId="7" pageNumber="450">Neobythites</emphasis>
</taxonomicName>
, differing from
<taxonomicName id="4C754D449016FFC8FD86FCBB8BACAB67" authorityName="Goode &amp; Bean" authorityYear="1885" box="[614,746,835,861]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FD86FCBB8BACAB67" box="[614,746,835,861]" italics="true" pageId="7" pageNumber="450">Neobythites</emphasis>
</taxonomicName>
in: “1) bones of head soft; 2) pseudobranchiae absent; 3) developed gill-rakers on first gill-arch fewer than 5; 4) dorsal inserted above the upper angle of the gill-opening.”
<bibRefCitation id="EFE44B369016FFC8FF77FC738806AB9F" author="Gosline, W. A." box="[151,320,906,933]" pageId="7" pageNumber="450" pagination="68 - 83" refId="ref18451" refString="Gosline, W. A. (1954) Fishes killed by the 1950 eruption of Mauna Loa. Brotulidae. Pacific Science, 8, 68-83." type="journal article" year="1954">Gosline (1954)</bibRefCitation>
postulated the existence of a group of oviparous brotulid genera which he thought are intermediate between oviparous and viviparous genera and counted
<taxonomicName id="4C754D449016FFC8FCEAFC578A8DABF3" authorityName="Alcock" authorityYear="1889" box="[778,971,943,969]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FCEAFC578A8DABF3" box="[778,971,943,969]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D449016FFC8FBE4FC578D2FABF3" box="[1028,1129,943,969]" family="Ophidiidae" genus="Volcanus" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FBE4FC578D2FABF3" box="[1028,1129,943,969]" italics="true" pageId="7" pageNumber="450">Volcanus</emphasis>
</taxonomicName>
(syn.
<taxonomicName id="4C754D449016FFC8FB52FC578C05ABF3" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[1202,1347,943,969]" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FB52FC578C05ABF3" box="[1202,1347,943,969]" italics="true" pageId="7" pageNumber="450">Luciobrotula</emphasis>
</taxonomicName>
) to this group.
<bibRefCitation id="EFE44B369016FFC8FF0BFC2B88CDABD7" author="Cohen, D. M." box="[235,395,979,1005]" pageId="7" pageNumber="450" pagination="38 - 398" refId="ref17863" refString="Cohen, D. M. (1964) A review of the ophidioid fish genus Luciobrotula with the description of a new species from the western North Atlantic. Bulletin Marine Science, 38-398." type="book chapter" year="1964">Cohen (1964)</bibRefCitation>
added
<taxonomicName id="4C754D449016FFC8FE06FC2B8B02ABD6" authorityName="Heller &amp; Snodgrass" authorityYear="1903" box="[486,580,979,1004]" family="Ophidiidae" genus="Petrotyx" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FE06FC2B8B02ABD6" box="[486,580,979,1004]" italics="true" pageId="7" pageNumber="450">Petrotyx</emphasis>
</taxonomicName>
to this group and
<bibRefCitation id="EFE44B369016FFC8FCC7FC2B8D0CABD7" author="Cohen, D. M. &amp; Nielsen, J. G." box="[807,1098,979,1005]" pageId="7" pageNumber="450" pagination="1 - 72" refId="ref17901" refString="Cohen, D. M. &amp; Nielsen, J. G. (1978) Guide to the identification of genera of the fish order Ophidiiformes with a tentative classification of the order. NOAA Technical Report NMFS Circular, 417, 1-72. https://doi.org/10.5962/bhl.title.63242" type="journal article" year="1978">Cohen &amp; Nielsen (1978)</bibRefCitation>
considered
<taxonomicName id="4C754D449016FFC8FB3BFC2B8CDAABD7" authorityName="Alcock" authorityYear="1889" box="[1243,1436,979,1005]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FB3BFC2B8CDAABD7" box="[1243,1436,979,1005]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
belonging to a cluster with
<emphasis id="B901EAD59016FFC8FE3FFC0F8BA7AC2A" box="[479,737,1015,1041]" italics="true" pageId="7" pageNumber="450">
<taxonomicName id="4C754D449016FFC8FE3FFC0F8B35AC2B" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[479,627,1015,1041]" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">Luciobrotula</taxonomicName>
,
<taxonomicName id="4C754D449016FFC8FD63FC0F8BA7AC2A" authorityName="Heller &amp; Snodgrass" authorityYear="1903" box="[643,737,1015,1040]" family="Ophidiidae" genus="Petrotyx" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">Petrotyx</taxonomicName>
</emphasis>
and
<taxonomicName id="4C754D449016FFC8FCC0FC0F8AFFAC2B" authorityName="Cohen &amp; Nielsen" authorityYear="1978" box="[800,953,1015,1041]" family="Ophidiidae" genus="Spottobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FCC0FC0F8AFFAC2B" box="[800,953,1015,1041]" italics="true" pageId="7" pageNumber="450">Spottobrotula</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE44B369016FFC8FC2BFC0F8DC5AC2B" author="Machida, Y." box="[971,1155,1015,1041]" pageId="7" pageNumber="450" pagination="371 - 375" refId="ref18614" refString="Machida, Y. (1984) Ophidiidae. In: Okamura, O. &amp; Kitajima, T. (Eds.), Fishes of the Okinawa Trough and the adjacent waters. Vol. I. The intensive research of unexploited fishery resources on continental slopes. Japan Fisheries Resource Conservation Association, Tokyo, pp. 244 - 261 + 371-375." type="book chapter" year="1984">Machida (1984)</bibRefCitation>
found
<taxonomicName id="4C754D449016FFC8FB3BFC0F8CDAAC2B" authorityName="Alcock" authorityYear="1889" box="[1243,1436,1015,1041]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FB3BFC0F8CDAAC2B" box="[1243,1436,1015,1041]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
resembling
<taxonomicName id="4C754D449016FFC8FEFBFBE388D9AC0F" authorityName="Goode &amp; Bean" authorityYear="1885" box="[283,415,1051,1077]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FEFBFBE388D9AC0F" box="[283,415,1051,1077]" italics="true" pageId="7" pageNumber="450">Neobythites</emphasis>
</taxonomicName>
. Studies using molecular phylogenetic analysis of neobythitine taxa so far are few.
<bibRefCitation id="EFE44B369016FFC8FAB0FBE38860AC63" author="Moller, P. R. &amp; Knudsen, S. W. &amp; Schwarzhans, W. &amp; Nielsen, J. G." pageId="7" pageNumber="450" pagination="391 - 408" refId="ref19037" refString="Moller, P. R., Knudsen, S. W., Schwarzhans, W. &amp; Nielsen, J. G. (2016) A new classification of viviparous brotulas (Bythitidae) - with family status for Dinematichthyidae - based on molecular, morphological and fossil data. Molecular Phylogenetics and Evolution, 100, 391-408. https://doi.org/10.1016/j.ympev.2016.04.008" type="journal article" year="2016">
Møller
<emphasis id="B901EAD59016FFC8FF77FBC78994AC63" box="[151,210,1087,1113]" italics="true" pageId="7" pageNumber="450">et al.</emphasis>
(2016)
</bibRefCitation>
did not include
<taxonomicName id="4C754D449016FFC8FE07FBC78BEEAC63" authorityName="Alcock" authorityYear="1889" box="[487,680,1087,1113]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FE07FBC78BEEAC63" box="[487,680,1087,1113]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE44B369016FFC8FD59FBC78ADCAC63" author="Girard, M. G. &amp; Mundy, B. C. &amp; Nonaka, A. &amp; Johnson, G. D." box="[697,922,1087,1113]" pageId="7" pageNumber="450" pagination="474 - 489" refId="ref18250" refString="Girard, M. G., Mundy, B. C., Nonaka, A. &amp; Johnson, G. D. (2023) Cusk-eel confusion: revisions of larval Luciobrotula and Pycnocraspedum and re-descriptions of two bythitid larvae (Ophidiiformes). Ichthyological Research, 70 (4), 474-489. https://doi.org/10.1007/s10228-023-00906-4" type="journal article" year="2023">
Girard
<emphasis id="B901EAD59016FFC8FCEBFBC78A00AC63" box="[779,838,1087,1113]" italics="true" pageId="7" pageNumber="450">et al.</emphasis>
(2023)
</bibRefCitation>
resolved
<taxonomicName id="4C754D449016FFC8FBEDFBC78D88AC63" authorityName="Alcock" authorityYear="1889" box="[1037,1230,1087,1113]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FBEDFBC78D88AC63" box="[1037,1230,1087,1113]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
as sister taxon of
<taxonomicName id="4C754D449016FFC8FF77FB9B885DAC47" authorityName="Goode &amp; Bean" authorityYear="1885" box="[151,283,1123,1149]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FF77FB9B885DAC47" box="[151,283,1123,1149]" italics="true" pageId="7" pageNumber="450">Neobythites</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE44B369016FFC8FEC9FB9B8B5AAC47" author="Wong, M. - K. &amp; Chen, W. - J." box="[297,540,1123,1149]" pageId="7" pageNumber="450" pagination="1 - 13" refId="ref20545" refString="Wong, M. - K. &amp; Chen, W. - J. (2024) Exploring the phylogeny and depth evolution of cusk eels and their relatives (Ophidiiformes: Ophidioidei). Molecular Phylogenetics and Evolution, 199, 1-13. https://doi.org/10.1016/j.ympev.2024.108164" type="journal article" year="2024">Wong &amp; Chen (2024)</bibRefCitation>
showed
<taxonomicName id="4C754D449016FFC8FD9FFB9B8A06AC47" authorityName="Alcock" authorityYear="1889" box="[639,832,1123,1149]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FD9FFB9B8A06AC47" box="[639,832,1123,1149]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
in a clade with
<taxonomicName id="4C754D449016FFC8FC12FB9B8DCBAC47" box="[1010,1165,1123,1149]" family="Ophidiidae" genus="Brotulataenia" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FC12FB9B8DCBAC47" box="[1010,1165,1123,1149]" italics="true" pageId="7" pageNumber="450">Brotulataenia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D449016FFC8FB23FB9B8CC5AC46" authorityName="Alcock" authorityYear="1891" box="[1219,1411,1123,1148]" family="Ophidiidae" genus="Lamprogrammus" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FB23FB9B8CC5AC46" box="[1219,1411,1123,1148]" italics="true" pageId="7" pageNumber="450">Lamprogrammus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCA36C79016FFC8FF27FB7F8D46AE17" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="450">
A comparison of the diagnostic key characters of
<taxonomicName id="4C754D449016FFC8FCD3FB7F8AB2AC9B" authorityName="Alcock" authorityYear="1889" box="[819,1012,1159,1185]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FCD3FB7F8AB2AC9B" box="[819,1012,1159,1185]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
—developed gill rakers, number of precaudal vertebrae, origin of dorsal fin, presence/absence of preopercular spines, number of pectoral-fin rays, and otolith morphology—showed the highest degree of congruence with
<taxonomicName id="4C754D449016FFC8FC17FB378D3DACD3" authorityName="Goode &amp; Bean" authorityYear="1885" box="[1015,1147,1231,1257]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FC17FB378D3DACD3" box="[1015,1147,1231,1257]" italics="true" pageId="7" pageNumber="450">Neobythites</emphasis>
</taxonomicName>
(except origin of dorsal fin) and
<taxonomicName id="4C754D449016FFC8FF1AFB0B88D2AD36" authorityName="Alcock" authorityYear="1890" box="[250,404,1267,1292]" family="Ophidiidae" genus="Monomitopus" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FF1AFB0B88D2AD36" box="[250,404,1267,1292]" italics="true" pageId="7" pageNumber="450">Monomitopus</emphasis>
</taxonomicName>
(except origin of dorsal fin and developed gill rakers) (
<tableCitation id="C6F7037C9016FFC8FBF3FB0B8D2EAD37" box="[1043,1128,1267,1293]" captionStart="TABLE 1" captionStartId="2.[152,244,151,175]" captionTargetBox="[163,1411,227,1782]" captionTargetId="graphics-48@2.[151,1437,256,1787]" captionText="TABLE 1. Selected keyparameters ofthecurrently 37recognizedophidiidgenera.Characterssharedwith Pycnocraspedum are highlighted. D/V = first dorsal-fin ray above vertebra." httpUri="http://table.plazi.org/id/DF0A664F9013FFCDFF78FF6F8BBBA8E9" pageId="7" pageNumber="450" tableUuid="DF0A664F9013FFCDFF78FF6F8BBBA8E9">Table 1</tableCitation>
). Particularly the origin of the dorsal fin in front of the first vertebra is an almost unique character within
<taxonomicName id="4C754D449016FFC8FBC9FAEF8D89AD0B" authorityName="Berg" authorityYear="1940" box="[1065,1231,1303,1329]" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="order">Ophidiiformes</taxonomicName>
, shared only with
<taxonomicName id="4C754D449016FFC8FF77FAC3886EAD6F" authorityName="Nielsen" authorityYear="1986" box="[151,296,1339,1365]" family="Ophidiidae" genus="Leptobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FF77FAC3886EAD6F" box="[151,296,1339,1365]" italics="true" pageId="7" pageNumber="450">Leptobrotula</emphasis>
</taxonomicName>
. The dorsal-fin commences above the first vertebra in some species of the genera
<taxonomicName id="4C754D449016FFC8FB0DFAC38CD3AD6F" authorityName="Alcock" authorityYear="1889" box="[1261,1429,1339,1365]" family="Ophidiidae" genus="Glyptophidium" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FB0DFAC38CD3AD6F" box="[1261,1429,1339,1365]" italics="true" pageId="7" pageNumber="450">Glyptophidium</emphasis>
</taxonomicName>
,
<taxonomicName id="4C754D449016FFC8FF77FAA789B5AD43" authorityName="Bleeker" authorityYear="1857" box="[151,243,1375,1401]" family="Ophidiidae" genus="Sirembo" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FF77FAA789B5AD43" box="[151,243,1375,1401]" italics="true" pageId="7" pageNumber="450">Sirembo</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D449016FFC8FECDFAA78880AD43" authorityName="Cohen &amp; Nielsen" authorityYear="1978" box="[301,454,1375,1401]" family="Ophidiidae" genus="Spottobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FECDFAA78880AD43" box="[301,454,1375,1401]" italics="true" pageId="7" pageNumber="450">Spottobrotula</emphasis>
</taxonomicName>
otherwise it commences above the third vertebra or posteriorly in other ophidiiforms. A low number of developed gill rakers on the first gill arch is shared with several other genera such as
<taxonomicName id="4C754D449016FFC8FAF6FA7B8CD0ADA7" authorityName="Zugmayer" authorityYear="1911" box="[1302,1430,1411,1437]" family="Ophidiidae" genus="Barathrites" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FAF6FA7B8CD0ADA7" box="[1302,1430,1411,1437]" italics="true" pageId="7" pageNumber="450">Barathrites</emphasis>
</taxonomicName>
,
<taxonomicName id="4C754D449016FFC8FF77FA5F885CADFA" authorityName="Whitley" authorityYear="1941" box="[151,282,1447,1472]" family="Ophidiidae" genus="Dannevigia" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FF77FA5F885CADFA" box="[151,282,1447,1472]" italics="true" pageId="7" pageNumber="450">Dannevigia</emphasis>
</taxonomicName>
,
<emphasis id="B901EAD59016FFC8FEC6FA5F8B1DADFB" box="[294,603,1447,1473]" italics="true" pageId="7" pageNumber="450">
<taxonomicName id="4C754D449016FFC8FEC6FA5F8886ADFB" authorityName="Gill" authorityYear="1863" box="[294,448,1447,1473]" family="Ophidiidae" genus="Hoplobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">Hoplobrotula</taxonomicName>
,
<taxonomicName id="4C754D449016FFC8FE2AFA5F8B1DADFB" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[458,603,1447,1473]" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">Luciobrotula</taxonomicName>
</emphasis>
,
<taxonomicName id="4C754D449016FFC8FD88FA5F8B80ADFA" authorityName="Heller &amp; Snodgrass" authorityYear="1903" box="[616,710,1447,1472]" family="Ophidiidae" genus="Petrotyx" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FD88FA5F8B80ADFA" box="[616,710,1447,1472]" italics="true" pageId="7" pageNumber="450">Petrotyx</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D449016FFC8FD19FA5F8A13ADFB" authorityName="Bleeker" authorityYear="1857" box="[761,853,1447,1473]" family="Ophidiidae" genus="Sirembo" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FD19FA5F8A13ADFB" box="[761,853,1447,1473]" italics="true" pageId="7" pageNumber="450">Sirembo</emphasis>
</taxonomicName>
. Long, slender otoliths are also found in
<taxonomicName id="4C754D449016FFC8FAF6FA5F8CD0ADFA" authorityName="Goode &amp; Bean" authorityYear="1896" box="[1302,1430,1447,1472]" family="Ophidiidae" genus="Bassogigas" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FAF6FA5F8CD0ADFA" box="[1302,1430,1447,1472]" italics="true" pageId="7" pageNumber="450">Bassogigas</emphasis>
</taxonomicName>
,
<taxonomicName id="4C754D449016FFC8FF77FA338854ADDF" authorityName="Cohen &amp; Nielsen" authorityYear="1978" box="[151,274,1483,1509]" family="Ophidiidae" genus="Epetriodus" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FF77FA338854ADDF" box="[151,274,1483,1509]" italics="true" pageId="7" pageNumber="450">Epetriodus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C754D449016FFC8FEC0FA3388F7ADDF" authorityName="Smith &amp; Radcliffe" authorityYear="1913" box="[288,433,1483,1509]" family="Ophidiidae" genus="Luciobrotula" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FEC0FA3388F7ADDF" box="[288,433,1483,1509]" italics="true" pageId="7" pageNumber="450">Luciobrotula</emphasis>
</taxonomicName>
,
<taxonomicName id="4C754D449016FFC8FE20FA338B1CADDE" authorityName="Alcock" authorityYear="1890" box="[448,602,1483,1508]" family="Ophidiidae" genus="Monomitopus" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FE20FA338B1CADDE" box="[448,602,1483,1508]" italics="true" pageId="7" pageNumber="450">Monomitopus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D449016FFC8FD70FA338A52ADDF" authorityName="Goode &amp; Bean" authorityYear="1885" box="[656,788,1483,1509]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FD70FA338A52ADDF" box="[656,788,1483,1509]" italics="true" pageId="7" pageNumber="450">Neobythites</emphasis>
</taxonomicName>
, but only in
<taxonomicName id="4C754D449016FFC8FC43FA338D65ADDE" authorityName="Goode &amp; Bean" authorityYear="1896" box="[931,1059,1483,1508]" family="Ophidiidae" genus="Bassogigas" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FC43FA338D65ADDE" box="[931,1059,1483,1508]" italics="true" pageId="7" pageNumber="450">Bassogigas</emphasis>
</taxonomicName>
,
<taxonomicName id="4C754D449016FFC8FBD1FA338DF3ADDF" authorityName="Goode &amp; Bean" authorityYear="1885" box="[1073,1205,1483,1509]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FBD1FA338DF3ADDF" box="[1073,1205,1483,1509]" italics="true" pageId="7" pageNumber="450">Neobythites</emphasis>
</taxonomicName>
and in some species of
<taxonomicName id="4C754D449016FFC8FF58FA178814AE32" authorityName="Alcock" authorityYear="1890" box="[184,338,1519,1544]" family="Ophidiidae" genus="Monomitopus" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FF58FA178814AE32" box="[184,338,1519,1544]" italics="true" pageId="7" pageNumber="450">Monomitopus</emphasis>
</taxonomicName>
the slender otolith shape is paired with a long sulcus. This comparison would indeed support a relationship of
<taxonomicName id="4C754D449016FFC8FEA1F9EB8B44AE17" authorityName="Alcock" authorityYear="1889" box="[321,514,1555,1581]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FEA1F9EB8B44AE17" box="[321,514,1555,1581]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
with
<taxonomicName id="4C754D449016FFC8FDA1F9EB8B83AE17" authorityName="Goode &amp; Bean" authorityYear="1885" box="[577,709,1555,1581]" family="Ophidiidae" genus="Neobythites" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FDA1F9EB8B83AE17" box="[577,709,1555,1581]" italics="true" pageId="7" pageNumber="450">Neobythites</emphasis>
</taxonomicName>
as the most likely outcome.
</paragraph>
<paragraph id="8BCA36C79016FFC8FF27F9CE8C6EAF13" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="450">
<emphasis id="B901EAD59016FFC8FF27F9CE886EAE6A" bold="true" box="[199,296,1590,1616]" pageId="7" pageNumber="450">Species:</emphasis>
We consider all previously described 5 extant species as valid for
<taxonomicName id="4C754D449016FFC8FBE2F9CF8D85AE6B" authorityName="Alcock" authorityYear="1889" box="[1026,1219,1591,1617]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="B901EAD59016FFC8FBE2F9CF8D85AE6B" box="[1026,1219,1591,1617]" italics="true" pageId="7" pageNumber="450">Pycnocraspedum</emphasis>
</taxonomicName>
and describe 2 new species:
<taxonomicName id="4C754D449016FFC8FF17F9A388C3AE4F" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[247,389,1627,1653]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="africanum" status="sp. nov.">
<emphasis id="B901EAD59016FFC8FF17F9A388C3AE4F" box="[247,389,1627,1653]" italics="true" pageId="7" pageNumber="450">P. africanum</emphasis>
</taxonomicName>
<emphasis id="B901EAD59016FFC8FE6CF9A388A0AE4F" bold="true" box="[396,486,1627,1653]" pageId="7" pageNumber="450">
<taxonomicNameLabel id="A23257AE9016FFC8FE6CF9A388A0AE4F" box="[396,486,1627,1653]" pageId="7" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the Western Indian Ocean,
<taxonomicName id="4C754D449016FFC8FCB9F9A38DC6AE4F" authority="Gosline, 1954" authorityName=", Gosline" authorityYear="1954" box="[857,1152,1626,1653]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD59016FFC8FCB9F9A38A9FAE4E" box="[857,985,1627,1652]" italics="true" pageId="7" pageNumber="450">P. armatum</emphasis>
<bibRefCitation id="EFE44B369016FFC8FC00F9A38DC6AE4F" author="Gosline, W. A." box="[992,1152,1626,1653]" pageId="7" pageNumber="450" pagination="68 - 83" refId="ref18451" refString="Gosline, W. A. (1954) Fishes killed by the 1950 eruption of Mauna Loa. Brotulidae. Pacific Science, 8, 68-83." type="journal article" year="1954">Gosline, 1954</bibRefCitation>
</taxonomicName>
, widely distributed in the Western and Central Pacific as far east as to Hawaii and south to
<collectingCountry id="F36276579016FFC8FC64F9878D73AEA3" box="[900,1077,1663,1689]" name="New Caledonia" pageId="7" pageNumber="450">New Caledonia</collectingCountry>
,
<taxonomicName id="4C754D449016FFC8FBA2F9878C1BAEA3" authority="Machida, 1984" authorityName="Machida" authorityYear="1984" box="[1090,1373,1663,1689]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="fulvum">
<emphasis id="B901EAD59016FFC8FBA2F9878DEDAEA3" box="[1090,1195,1663,1689]" italics="true" pageId="7" pageNumber="450">P. fulvum</emphasis>
<bibRefCitation id="EFE44B369016FFC8FB54F9878C1BAEA3" author="Machida, Y." box="[1204,1373,1663,1689]" pageId="7" pageNumber="450" pagination="371 - 375" refId="ref18614" refString="Machida, Y. (1984) Ophidiidae. In: Okamura, O. &amp; Kitajima, T. (Eds.), Fishes of the Okinawa Trough and the adjacent waters. Vol. I. The intensive research of unexploited fishery resources on continental slopes. Japan Fisheries Resource Conservation Association, Tokyo, pp. 244 - 261 + 371-375." type="book chapter" year="1984">Machida, 1984</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F36276579016FFC8FF77F95B899CAE87" box="[151,218,1699,1725]" name="Japan" pageId="7" pageNumber="450">Japan</collectingCountry>
,
<taxonomicName id="4C754D449016FFC8FF04F95B8B0FAE87" authority="(Matsubara, 1943)" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[228,585,1699,1725]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD59016FFC8FF04F95B8832AE87" box="[228,372,1699,1725]" italics="true" pageId="7" pageNumber="450">P. microlepis</emphasis>
(
<bibRefCitation id="EFE44B369016FFC8FE63F95B8B07AE87" author="Matsubara, K." box="[387,577,1699,1725]" pageId="7" pageNumber="450" pagination="37 - 81" refId="ref18682" refString="Matsubara, K. (1943) Ichthyological annotations from the depths of the sea Japan, I - VII. Journal of Sigenkagaku Kenkyusyo, 1, 37-81." type="journal article" year="1943">Matsubara, 1943</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F36276579016FFC8FD6DF95B8B8BAE87" box="[653,717,1699,1725]" name="Japan" pageId="7" pageNumber="450">Japan</collectingCountry>
is considered nominally valid but could not be reassessed due to the apparent loss of the unique
<typeStatus id="54CE88659016FFC8FE0FF93F8B64AEDB" box="[495,546,1735,1761]" pageId="7" pageNumber="450">type</typeStatus>
,
<taxonomicName id="4C754D449016FFC8FDCCF93F8A15AEDB" authority="(Parr, 1933)" baseAuthorityName="Parr" baseAuthorityYear="1933" box="[556,851,1735,1761]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD59016FFC8FDCCF93F8B8EAEDB" box="[556,712,1735,1761]" italics="true" pageId="7" pageNumber="450">P. phyllosoma</emphasis>
(
<bibRefCitation id="EFE44B369016FFC8FD35F93F8A0DAEDB" author="Parr, A. E." box="[725,843,1735,1761]" pageId="7" pageNumber="450" pagination="1 - 51" refId="ref19495" refString="Parr, A. E. (1933) Deepsea Berycomorphi and Percomorphi from the waters around the Bahama and Bermuda Islands. Bulletin Bingham Oceanographic Collection, 3 (6), 1-51." type="journal article" year="1933">Parr, 1933</bibRefCitation>
)
</taxonomicName>
from the Caribbean,
<taxonomicName id="4C754D449016FFC8FBDEF93F8D9DAEDB" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[1086,1243,1735,1761]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="rowleyensis" status="sp. nov.">
<emphasis id="B901EAD59016FFC8FBDEF93F8D9DAEDB" box="[1086,1243,1735,1761]" italics="true" pageId="7" pageNumber="450">P. rowleyensis</emphasis>
</taxonomicName>
<emphasis id="B901EAD59016FFC8FB01F93F8C7CAEDB" bold="true" box="[1249,1338,1735,1761]" pageId="7" pageNumber="450">
<taxonomicNameLabel id="A23257AE9016FFC8FB01F93F8C7CAEDB" box="[1249,1338,1735,1761]" pageId="7" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from off northwestern
<collectingCountry id="F36276579016FFC8FECEF91388DEAF3F" box="[302,408,1771,1797]" name="Australia" pageId="7" pageNumber="450">Australia</collectingCountry>
, and the
<typeStatus id="54CE88659016FFC8FE18F9138B6EAF3F" box="[504,552,1771,1797]" pageId="7" pageNumber="450">type</typeStatus>
species
<taxonomicName id="4C754D449016FFC8FD64F9138A8EAF3F" authority="Alcock, 1889" authorityName="Alcock" authorityYear="1889" box="[644,968,1771,1797]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD59016FFC8FD64F9138A68AF3E" box="[644,814,1771,1796]" italics="true" pageId="7" pageNumber="450">P. squamipinne</emphasis>
<bibRefCitation id="EFE44B369016FFC8FCD2F9138A8EAF3F" author="Alcock, A." box="[818,968,1771,1797]" pageId="7" pageNumber="450" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
</taxonomicName>
from the Bay of Bengal. In addition, there is one fossil otolith-based species from the Early Pliocene of
<collectingCountry id="F36276579016FFC8FCA5F8F78A3BAF13" box="[837,893,1807,1833]" name="Italy" pageId="7" pageNumber="450">Italy</collectingCountry>
:
<taxonomicName id="4C754D449016FFC8FC67F8F78C65AF13" authority="(Schwarzhans, 1978)" baseAuthorityName="Schwarzhans" baseAuthorityYear="1978" box="[903,1315,1807,1833]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="centonaense">
<emphasis id="B901EAD59016FFC8FC67F8F78D6BAF12" box="[903,1069,1807,1832]" italics="true" pageId="7" pageNumber="450">P. centonaense</emphasis>
(
<bibRefCitation id="EFE44B369016FFC8FBDCF8F78C5DAF13" author="Schwarzhans, W." box="[1084,1307,1807,1833]" pageId="7" pageNumber="450" pagination="1 - 52" refId="ref19902" refString="Schwarzhans, W. (1978) Otolithen aus dem Unter-Pliozan von Sud-Sizilien und aus der Toscana. Berliner Geowissenschaftliche Abhandlungen, 8, 1-52." type="journal article" year="1978">Schwarzhans, 1978</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="47613FFB5B06E19053F468AF4249F9DF" ID-DOI="10.11646/zootaxa.5636.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="15561733" ID-ZooBank="ED905457-C12F-45D2-B3F4-77082279E26B" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1748690707546" checkinUser="plazi" docAuthor="Schwarzhans, Werner W., Psomadakis, Peter N. &amp; Nielsen, Jørgen G." docDate="2025" docId="03DC87D1901BFFC3FF77FF6F8C16AB03" docLanguage="en" docName="zootaxa.5636.3.2.pdf" docOrigin="Zootaxa 5636 (3)" docSource="https://doi.org/10.11646/zootaxa.5636.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pycnocraspedum africanum Schwarzhans, Psomadakis &amp; Nielsen, 2025, sp. nov." docType="treatment" docVersion="5" lastPageNumber="455" masterDocId="FFE5FFA99011FFCFFFE0FFF88946A83A" masterDocTitle="Revision of the bathydemersal fish genus Pycnocraspedum (Ophidiidae; Teleostei), with description of two new species" masterLastPageNumber="472" masterPageNumber="443" pageNumber="453" updateTime="1749063193116" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5F55E05ED4C5409655960F2AA46BCD91">Revision of the bathydemersal fish genus Pycnocraspedum (Ophidiidae; Teleostei), with description of two new species</mods:title>
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<mods:namePart id="8F16C6BF0E5F30F05FC8CD5DAEE06FBD">Schwarzhans, Werner W.</mods:namePart>
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<mods:namePart id="CBF8E999B4DB15577F5E715C619E5CBD">Psomadakis, Peter N.</mods:namePart>
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<mods:namePart id="2EB273FBFD2B1FE7E62A3B186CE424EF">Nielsen, Jørgen G.</mods:namePart>
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<mods:date id="64640DA974A5B2B6D5CE8AFA2BCDB405">2025</mods:date>
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<paragraph id="8BCA36C7901BFFC5FF77FF6F8B17A88B" blockId="10.[151,593,151,214]" box="[151,593,151,178]" pageId="10" pageNumber="453">
<heading id="D08281AB901BFFC5FF77FF6F8B17A88B" bold="true" box="[151,593,151,178]" fontSize="11" level="1" pageId="10" pageNumber="453" reason="1">
<emphasis id="B901EAD5901BFFC5FF77FF6F8B17A88B" bold="true" box="[151,593,151,178]" pageId="10" pageNumber="453">
<taxonomicName id="4C754D44901BFFC5FF77FF6F88AAA88B" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[151,492,151,178]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="africanum" status="sp. nov.">
<emphasis id="B901EAD5901BFFC5FF77FF6F88AAA88B" bold="true" box="[151,492,151,178]" italics="true" pageId="10" pageNumber="453">Pycnocraspedum africanum</emphasis>
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<subSubSection id="C36F654C901BFFC5FF77FF4388ACA8EF" box="[151,490,187,214]" pageId="10" pageNumber="453" type="description">
<paragraph id="8BCA36C7901BFFC5FF77FF4388ACA8EF" blockId="10.[151,593,151,214]" box="[151,490,187,214]" pageId="10" pageNumber="453">
<figureCitation id="134E2A42901BFFC5FF77FF438847A8EF" box="[151,257,187,213]" captionStart="FIGURE 3" captionStartId="11.[151,250,1786,1810]" captionTargetBox="[256,1328,181,1748]" captionTargetId="figure-17@11.[256,1331,181,1762]" captionTargetPageId="11" captionText="FIGURE 3. Detail figures and drawings of preopercular spines and lower preopercular pores in Pycnocraspedum; preopercular spines are numbered from 1 through 4 from the lower angle of the preoperculum upwards; abbreviation lpp = lower preopercular pore. 3A3B) Pycnocraspedum armatum, USNM 227412, 3A1-3A2 specimen 1 left side, 3A3 specimen 1 right side (mirror imaged), 3B specimen 2; 3C3D) Pycnocraspedum phyllosoma, 3C USNM 421587, 3D USNM 421586; 3E) Pycnocraspedum fulvum (holotype), BSKU 27497; 3F3G) Pycnocraspedum africanum sp. nov., 3F holotype, MNHN 2000-5691, 3G paratype, MNHN 2000-5692; 3H) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021; 3I3K) Pycnocraspedum squamipinne, 3I lectotype, BMNH 1890.7.31.1, 3J ZMUC P771871, 3K ZMUC P771737." figureDoi="http://doi.org/10.5281/zenodo.15561739" httpUri="https://zenodo.org/record/15561739/files/figure.png" pageId="10" pageNumber="453">Figures 3</figureCitation>
,
<figureCitation id="134E2A42901BFFC5FEEEFF438858A8EC" box="[270,286,187,214]" captionStart="FIGURE 5" captionStartId="14.[151,250,1750,1774]" captionTargetBox="[245,1358,193,1715]" captionTargetId="figure-23@14.[222,1366,181,1726]" captionTargetPageId="14" captionText="FIGURE 5. Otoliths, inner faces (if not noted otherwise) shown from right side. 5A5C) Pycnocraspedum africanum sp. nov., 5A holotype, MNHN 2000-5691 (5A2 dorsal view), 5B paratype MNHN 2000-5692, 5C tentatively assigned specimen, IRSNB (5C2 dorsal view, 5C3 anterior view); 5D5E) Pycnocraspedum armatum, 5D USNM 227412 (5D2 ventral view), 5E USNM 227390 (5E2 ventral view); 5F) Pycnocraspedum fulvum (holotype), BSKU 27497 (5F2 ventral view, 5F3 anterior view); 5G) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021 (5G2 ventral view); 5H) Pycnocraspedum phyllosoma, USNM 421587 (5H2 ventral view, 5H3 anterior view); 5I5K) Pycnocraspedum squamipinne, 5I lectotype, BMNH 1890.7.31.1 (5I2 ventral view), 5J ZMUC P771871, 5L ZMUC P771739 (5L2 ventral view), 5K ZMUC P771738 (5K2 ventral view); 5M) †Pycnocraspedum cetonaense (holotype) SMF P.5694 (5M2 dorsal view, 5M3 anterior view)." figureDoi="http://doi.org/10.5281/zenodo.15561743" httpUri="https://zenodo.org/record/15561743/files/figure.png" pageId="10" pageNumber="453">5</figureCitation>
,
<figureCitation id="134E2A42901BFFC5FECAFF43887CA8EF" box="[298,314,187,213]" captionStart="FIGURE 6" captionStartId="16.[151,250,1563,1587]" captionTargetBox="[151,1435,181,1539]" captionTargetId="figure-23@16.[151,1435,181,1539]" captionTargetPageId="16" captionText="FIGURE 6. Pycnocraspedum africanum sp. nov., holotype, MNHN 2000-5691, 420 mm SL, 22°21'7.2''S, 43°4'4.8''E, 450 m." figureDoi="http://doi.org/10.5281/zenodo.15561745" httpUri="https://zenodo.org/record/15561745/files/figure.png" pageId="10" pageNumber="453">6</figureCitation>
,
<figureCitation id="134E2A42901BFFC5FEA6FF438823A8EF" box="[326,357,187,213]" captionStart="FIGURE 12" captionStartId="24.[151,250,899,923]" captionTargetBox="[153,1433,186,871]" captionTargetId="figure-23@24.[151,1436,181,875]" captionTargetPageId="24" captionText="FIGURE 12. Geographic distribution of the species of Pycnocraspedum. One symbol can represent more than one location." figureDoi="http://doi.org/10.5281/zenodo.15561755" httpUri="https://zenodo.org/record/15561755/files/figure.png" pageId="10" pageNumber="453">12</figureCitation>
;
<tableCitation id="C6F7037C901BFFC5FE90FF438888A8EF" box="[368,462,187,213]" captionStart="TABLE 1" captionStartId="2.[152,244,151,175]" captionTargetBox="[163,1411,227,1782]" captionTargetId="graphics-48@2.[151,1437,256,1787]" captionText="TABLE 1. Selected keyparameters ofthecurrently 37recognizedophidiidgenera.Characterssharedwith Pycnocraspedum are highlighted. D/V = first dorsal-fin ray above vertebra." httpUri="http://table.plazi.org/id/DF0A664F9013FFCDFF78FF6F8BBBA8E9" pageId="10" pageNumber="453" tableUuid="DF0A664F9013FFCDFF78FF6F8BBBA8E9">Tables 1</tableCitation>
,
<tableCitation id="C6F7037C901BFFC5FE3CFF4388ACA8EF" box="[476,490,187,213]" captionStart="TABLE 2" captionStartId="8.[152,244,720,744]" captionTargetPageId="8" captionText="TABLE 2. Meristic, morphometric and otolith characters of the species of the genus Pycnocraspedum. For abbreviations" httpUri="http://table.plazi.org/id/DF0A664F9019FFC7FF78FD288B4AAB36" pageId="10" pageNumber="453" tableUuid="DF0A664F9019FFC7FF78FD288B4AAB36">2</tableCitation>
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<taxonomicName id="4C754D44901BFFC5FF77FEF88889A92D" ID-CoL="4QM83" authorityName="Alcock" authorityYear="1889" box="[151,463,256,280]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901BFFC5FF77FEF88889A92D" box="[151,463,256,280]" italics="true" pageId="10" pageNumber="453">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
(non
<bibRefCitation id="EFE44B36901BFFC5FDECFEF88BDDA922" author="Alcock, A." box="[524,667,256,280]" pageId="10" pageNumber="453" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
):
<bibRefCitation id="EFE44B36901BFFC5FD4EFEF88A29A922" author="Schwarzhans, W." box="[686,879,256,280]" pageId="10" pageNumber="453" pagination="71 - 98" refId="ref19961" refString="Schwarzhans, W. (1994) Sexual and ontogenetic dimorphism in otoliths of the family Ophidiidae. Cybium, 18, 71-98." type="journal article" year="1994">Schwarzhans 1994</bibRefCitation>
(including
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from MNHN),
<bibRefCitation id="EFE44B36901BFFC5FB04FEF88CDBA922" author="Fricke, R. &amp; Mahafina, J. &amp; Behivoke, F. &amp; Jaonalison, H. &amp; Leopold, M. &amp; Ponton, D." box="[1252,1437,256,280]" pageId="10" pageNumber="453" pagination="1 - 432" refId="ref18190" refString="Fricke, R., Mahafina, J., Behivoke, F., Jaonalison, H., Leopold, M. &amp; Ponton, D. (2018) Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa, 3 (1), 1-432." type="journal article" year="2018">
Fricke
<emphasis id="B901EAD5901BFFC5FACDFEF88C24A922" box="[1325,1378,256,280]" italics="true" pageId="10" pageNumber="453">et al.</emphasis>
2018
</bibRefCitation>
(same specimens from MNHN including
<typeStatus id="54CE8865901BFFC5FD8BFEE68B82A90C" box="[619,708,286,310]" pageId="10" pageNumber="453" type="holotype">holotype</typeStatus>
and
<typeStatus id="54CE8865901BFFC5FD15FEE68A16A90C" box="[757,848,286,310]" pageId="10" pageNumber="453" type="paratype">paratype</typeStatus>
), Nielsen &amp; Uiblein (2022).
</paragraph>
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<treatmentCitationGroup id="AB6511E9901BFFC5FF77FEC58868A94E" pageId="10" pageNumber="453">
<taxonomicName id="4C754D44901BFFC5FF77FEC58B52A96F" ID-CoL="4QM83" authorityName="Alcock" authorityYear="1889" box="[151,532,317,341]" family="Ophidiidae" genus="Pycnocraspedum" isUncertain="true" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B901EAD5901BFFC5FF77FEC58803A96F" box="[151,325,317,341]" italics="true" pageId="10" pageNumber="453">Pycnocraspedum</emphasis>
sp. cf.
<emphasis id="B901EAD5901BFFC5FE73FEC68B52A96F" box="[403,532,318,341]" italics="true" pageId="10" pageNumber="453">squamipinne</emphasis>
</taxonomicName>
? (non
<bibRefCitation id="EFE44B36901BFFC5FDBCFEC58BACA96F" author="Alcock, A." box="[604,746,317,341]" pageId="10" pageNumber="453" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
):
<treatmentCitation id="0AD410D6901BFFC5FD1CFEC58AA7A96F" author="Prokofiev, A. M." box="[764,993,317,341]" page="118" pageId="10" pageNumber="453" year="2005">
<bibRefCitation id="EFE44B36901BFFC5FD1CFEC58AF6A96F" author="Prokofiev, A. M." box="[764,944,317,341]" pageId="10" pageNumber="453" pagination="111 - 128" refId="ref19532" refString="Prokofiev, A. M. (2005) On some rare ophidiiform fishes from the South Atlantic and Indo-West Pacific, with erection of a new genus, Megacataetyx gen. novum (Teleostei: Ophidiiformes). Estestvennye i Tekhnicheskie Nauki, 2, 111-128." type="journal article" year="2005">Prokofiev (2005)</bibRefCitation>
: 118
</treatmentCitation>
, figs. 910 (same specimen from ZIN now
<typeStatus id="54CE8865901BFFC5FF27FEA48865A94E" box="[199,291,348,372]" pageId="10" pageNumber="453" type="paratype">paratype</typeStatus>
).
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</paragraph>
</subSubSection>
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<emphasis id="B901EAD5901BFFC5FF77FE59883BA981" bold="true" box="[151,381,417,443]" pageId="10" pageNumber="453">Material examined</emphasis>
(
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,
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SL):
<materialsCitation id="3B1D3C9A901BFFC5FCE0FE5A8D8EA9DA" collectingDate="1973-11-27" collectingDateMax="2007-11-24" collectingDateMin="1972-01-19" collectingMethod="beam trawl" collectionCode="MNHN" country="Madagascar" county="Vityaz" elevation="450" latitude="-21.266666" location="Crosnier st. ch 095" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." pageId="10" pageNumber="453" specimenCode="2000-5691" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE8865901BFFC5FCE0FE5A8A2BA986" box="[768,877,418,444]" pageId="10" pageNumber="453" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED64AE02901BFFC5FC9AFE5A8A96A986" box="[890,976,418,444]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="453">MNHN</collectionCode>
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SL,
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,
<geoCoordinate id="EE415000901BFFC5FF77FE3E8849A9DA" box="[151,271,454,480]" degrees="43" direction="east" minutes="4" orientation="longitude" pageId="10" pageNumber="453" precision="1" seconds="4.8" value="43.068">43°4'4.8''E</geoCoordinate>
(off western
<collectingCountry id="F3627657901BFFC5FE42FE3E8B68A9DA" box="[418,558,454,480]" name="Madagascar" pageId="10" pageNumber="453">Madagascar</collectingCountry>
),
<collectingMethod id="52344ED0901BFFC5FDDFFE3E8BFAA9DA" box="[575,700,454,480]" pageId="10" pageNumber="453">beam trawl</collectingMethod>
<quantity id="4C8D9B22901BFFC5FD21FE3E8A4FA9DA" box="[705,777,454,481]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="453" unit="m" value="450.0">
<elevation id="0058D1F4901BFFC5FD21FE3E8A4FA9DA" box="[705,777,454,481]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="453" unit="m" value="450.0">450 m</elevation>
</quantity>
,
<location id="8EAA601C901BFFC5FCF3FE3E8AA6A9DB" LSID="urn:lsid:plazi:treatment:03DC87D1901BFFC3FF77FF6F8C16AB03:8EAA601C901BFFC5FCF3FE3E8AA6A9DB" box="[787,992,454,481]" country="Madagascar" county="Vityaz" latitude="-21.266666" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." name="Crosnier st. ch 095" pageId="10" pageNumber="453">Crosnier st. ch095</location>
,
<date id="FFCB1007901BFFC5FC0AFE3E8D85A9DA" box="[1002,1219,454,480]" pageId="10" pageNumber="453" value="1973-11-27">
<collectingDate id="EF8FE9EF901BFFC5FC0AFE3E8D85A9DA" box="[1002,1219,454,480]" pageId="10" pageNumber="453" value="1973-11-27">27 November 1973</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="3B1D3C9A901BFFC5FB2EFE3E881FAA12" collectingDate="1972-01-19" collectingDateMax="2007-11-24" collectingDateMin="1972-01-19" collectingMethod="beam trawl" collectionCode="MNHN" country="Madagascar" county="Vityaz" elevation="603" latitude="-21.266666" location="Channel" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." pageId="10" pageNumber="453" specimenCode="2000-5692" specimenCount="1" typeStatus="paratype">
<typeStatus id="54CE8865901BFFC5FB2EFE3E8C78A9DA" box="[1230,1342,454,480]" pageId="10" pageNumber="453" type="paratype">paratypes</typeStatus>
:
<collectionCode id="ED64AE02901BFFC5FAA7FE3E8CDBA9DA" box="[1351,1437,454,480]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="453">MNHN</collectionCode>
<quantity id="4C8D9B22901BFFC5FF77FE12883DAA3E" box="[151,379,490,517]" metricMagnitude="3" metricUnit="m" metricValue="2.847165" metricValueMax="5.69233" metricValueMin="0.002" pageId="10" pageNumber="453" unit="mm" value="2847165.0" valueMax="5692330.0" valueMin="2000.0">
<specimenCode id="DBD39EBC901BFFC5FF77FE12885EAA3F" box="[151,280,490,517]" pageId="10" pageNumber="453">20005692</specimenCode>
, 330 mm
</quantity>
SL,
<geoCoordinate id="EE415000901BFFC5FE4BFE128B77AA3E" box="[427,561,490,516]" degrees="12" direction="south" minutes="27" orientation="latitude" pageId="10" pageNumber="453" precision="1" seconds="7.2" value="-12.452">12°27'7.2''S</geoCoordinate>
<geoCoordinate id="EE415000901BFFC5FDD5FE128BFAAA3E" box="[565,700,490,516]" degrees="48" direction="east" minutes="12" orientation="longitude" pageId="10" pageNumber="453" precision="1" seconds="3.6" value="48.201">48°12'3.6''E</geoCoordinate>
(
<collectingCountry id="F3627657901BFFC5FD27FE128A16AA3E" box="[711,848,490,516]" name="Madagascar" pageId="10" pageNumber="453">Madagascar</collectingCountry>
<location id="8EAA601C901BFFC5FCB5FE128AF1AA3E" LSID="urn:lsid:plazi:treatment:03DC87D1901BFFC3FF77FF6F8C16AB03:8EAA601C901BFFC5FCB5FE128AF1AA3E" box="[853,951,490,516]" country="Madagascar" county="Vityaz" latitude="-21.266666" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." name="Channel" pageId="10" pageNumber="453">Channel</location>
),
<collectingMethod id="52344ED0901BFFC5FC27FE128D04AA3E" box="[967,1090,490,516]" pageId="10" pageNumber="453">beam trawl</collectingMethod>
<quantity id="4C8D9B22901BFFC5FBA6FE128D82AA3E" box="[1094,1220,490,517]" metricMagnitude="2" metricUnit="m" metricValue="6.025" metricValueMax="6.05" metricValueMin="6.0" pageId="10" pageNumber="453" unit="m" value="602.5" valueMax="605.0" valueMin="600.0">
<elevation id="0058D1F4901BFFC5FBA6FE128D82AA3E" box="[1094,1220,490,517]" metricMagnitude="2" metricUnit="m" metricValue="6.025" metricValueMax="6.05" metricValueMin="6.0" pageId="10" pageNumber="453" unit="m" value="602.5" valueMax="605.0" valueMin="600.0">600605 m</elevation>
</quantity>
,
<location id="8EAA601C901BFFC5FB2EFE128CDEAA3E" LSID="urn:lsid:plazi:treatment:03DC87D1901BFFC3FF77FF6F8C16AB03:8EAA601C901BFFC5FB2EFE128CDEAA3E" box="[1230,1432,490,516]" country="Madagascar" county="Vityaz" latitude="-21.266666" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." name="Crosnier st. ch 021" pageId="10" pageNumber="453">Crosnier st. ch021</location>
,
<date id="FFCB1007901BFFC5FF77FDF68813AA12" box="[151,341,526,552]" pageId="10" pageNumber="453" value="1972-01-19">
<collectingDate id="EF8FE9EF901BFFC5FF77FDF68813AA12" box="[151,341,526,552]" pageId="10" pageNumber="453" value="1972-01-19">19 January 1972</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="3B1D3C9A901BFFC5FE82FDF68B84AA76" collectingDate="2007-11-23" collectingDateMax="2007-11-24" collectingDateMin="1972-01-19" collectingMethod="beam trawl" collectionCode="SAIAB" country="Mozambique" county="Vityaz" elevation="292" latitude="-21.266666" location="Mozambique" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." pageId="10" pageNumber="453" specimenCode="98892" specimenCount="2" typeStatus="holotype">
<specimenCount id="9D73FD4E901BFFC5FE82FDF688ABAA12" box="[354,493,526,552]" count="2" pageId="10" pageNumber="453" type="generic">2 specimens</specimenCount>
<collectionCode id="ED64AE02901BFFC5FE14FDF68B0EAA12" box="[500,584,526,552]" pageId="10" pageNumber="453">SAIAB</collectionCode>
<specimenCode id="DBD39EBC901BFFC5FDB0FDF68BDFAA12" box="[592,665,526,552]" pageId="10" pageNumber="453">98892</specimenCode>
,
<quantity id="4C8D9B22901BFFC5FD45FDF68A7CAA12" box="[677,826,526,552]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="10" pageNumber="453" unit="mm" value="105.0" valueMax="110.0" valueMin="100.0">100110 mm</quantity>
SL,
<geoCoordinate id="EE415000901BFFC5FC91FDF68AA1AA13" box="[881,999,526,553]" degrees="12.6953" direction="south" orientation="latitude" pageId="10" pageNumber="453" precision="5" value="-12.6953">12.6953°S</geoCoordinate>
<geoCoordinate id="EE415000901BFFC5FC0FFDF68D1EAA12" box="[1007,1112,526,552]" degrees="40.667" direction="east" orientation="longitude" pageId="10" pageNumber="453" precision="55" value="40.667">40.667°E</geoCoordinate>
(off northern
<collectingCountry id="F3627657901BFFC5FB18FDF68CD6AA12" box="[1272,1424,526,552]" name="Mozambique" pageId="10" pageNumber="453">Mozambique</collectingCountry>
), haul 135,
<collectingMethod id="52344ED0901BFFC5FEE7FDCA88C3AA76" box="[263,389,562,588]" pageId="10" pageNumber="453">beam trawl</collectingMethod>
<quantity id="4C8D9B22901BFFC5FE6CFDCA8893AA76" box="[396,469,562,588]" metricMagnitude="2" metricUnit="m" metricValue="2.92" pageId="10" pageNumber="453" unit="m" value="292.0">
<elevation id="0058D1F4901BFFC5FE6CFDCA8893AA76" box="[396,469,562,588]" metricMagnitude="2" metricUnit="m" metricValue="2.92" pageId="10" pageNumber="453" unit="m" value="292.0">292 m</elevation>
</quantity>
,
<date id="FFCB1007901BFFC5FE01FDCA8BFBAA76" box="[481,701,562,588]" pageId="10" pageNumber="453" value="2007-11-23">
<collectingDate id="EF8FE9EF901BFFC5FE01FDCA8BFBAA76" box="[481,701,562,588]" pageId="10" pageNumber="453" value="2007-11-23">23 November 2007</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="3B1D3C9A901BFFC5FD29FDCA8A9DAA4A" collectingDate="2007-11-23" collectingDateMax="2007-11-24" collectingDateMin="1972-01-19" collectingMethod="beam trawl" collectionCode="SAIAB" country="Mozambique" county="Vityaz" elevation="391" latitude="-21.266666" location="Mozambique" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." pageId="10" pageNumber="453" specimenCode="98894" specimenCount="1" typeStatus="holotype">
<specimenCount id="9D73FD4E901BFFC5FD29FDCA8A0FAA76" box="[713,841,562,588]" count="1" pageId="10" pageNumber="453" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE02901BFFC5FCB0FDCA8AE2AA76" box="[848,932,562,588]" pageId="10" pageNumber="453">SAIAB</collectionCode>
<specimenCode id="DBD39EBC901BFFC5FC4BFDCA8AB2AA76" box="[939,1012,562,588]" pageId="10" pageNumber="453">98894</specimenCode>
,
<quantity id="4C8D9B22901BFFC5FBE0FDCA8D09AA76" box="[1024,1103,562,589]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="10" pageNumber="453" unit="mm" value="85.0">85 mm</quantity>
SL,
<geoCoordinate id="EE415000901BFFC5FB65FDCA8DBDAA77" box="[1157,1275,562,589]" degrees="12.6953" direction="south" orientation="latitude" pageId="10" pageNumber="453" precision="5" value="-12.6953">12.6953°S</geoCoordinate>
<geoCoordinate id="EE415000901BFFC5FAE2FDCA8C2DAA76" box="[1282,1387,562,588]" degrees="40.667" direction="east" orientation="longitude" pageId="10" pageNumber="453" precision="55" value="40.667">40.667°E</geoCoordinate>
(off northern
<collectingCountry id="F3627657901BFFC5FF1EFDAE88D0AA4A" box="[254,406,598,624]" name="Mozambique" pageId="10" pageNumber="453">Mozambique</collectingCountry>
), haul 136,
<collectingMethod id="52344ED0901BFFC5FDFCFDAE8BDDAA4A" box="[540,667,598,624]" pageId="10" pageNumber="453">beam trawl</collectingMethod>
<quantity id="4C8D9B22901BFFC5FD42FDAE8BAAAA4A" box="[674,748,598,624]" metricMagnitude="2" metricUnit="m" metricValue="3.91" pageId="10" pageNumber="453" unit="m" value="391.0">
<elevation id="0058D1F4901BFFC5FD42FDAE8BAAAA4A" box="[674,748,598,624]" metricMagnitude="2" metricUnit="m" metricValue="3.91" pageId="10" pageNumber="453" unit="m" value="391.0">391 m</elevation>
</quantity>
,
<date id="FFCB1007901BFFC5FD19FDAE8A91AA4A" box="[761,983,598,624]" pageId="10" pageNumber="453" value="2007-11-23">
<collectingDate id="EF8FE9EF901BFFC5FD19FDAE8A91AA4A" box="[761,983,598,624]" pageId="10" pageNumber="453" value="2007-11-23">23 November 2007</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="3B1D3C9A901BFFC5FC04FDAE8AEAAAAE" collectingDate="2007-11-24" collectingDateMax="2007-11-24" collectingDateMin="1972-01-19" collectionCode="SAIAB" country="Mozambique" county="Vityaz" elevation="450" latitude="-21.266666" location="Mozambique" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." pageId="10" pageNumber="453" specimenCode="98897" specimenCount="1" typeStatus="holotype">
<specimenCount id="9D73FD4E901BFFC5FC04FDAE8D22AA4A" box="[996,1124,598,624]" count="1" pageId="10" pageNumber="453" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE02901BFFC5FB8BFDAE8DF9AA4A" box="[1131,1215,598,624]" pageId="10" pageNumber="453">SAIAB</collectionCode>
<specimenCode id="DBD39EBC901BFFC5FB28FDAE8C57AA4A" box="[1224,1297,598,624]" pageId="10" pageNumber="453">98897</specimenCode>
,
<quantity id="4C8D9B22901BFFC5FAFDFDAE8C2BAA4A" box="[1309,1389,598,625]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="10" pageNumber="453" unit="mm" value="95.0">95 mm</quantity>
SL,
<geoCoordinate id="EE415000901BFFC5FF77FD82884AAAAE" box="[151,268,634,660]" degrees="11.1341" direction="south" orientation="latitude" pageId="10" pageNumber="453" precision="5" value="-11.1341">11.1341°S</geoCoordinate>
<geoCoordinate id="EE415000901BFFC5FEF2FD8288CFAAAE" box="[274,393,634,660]" degrees="40.7696" direction="east" orientation="longitude" pageId="10" pageNumber="453" precision="5" value="40.7696">40.7696°E</geoCoordinate>
(off northern
<collectingCountry id="F3627657901BFFC5FDC4FD828BFAAAAE" box="[548,700,634,660]" name="Mozambique" pageId="10" pageNumber="453">Mozambique</collectingCountry>
),
<date id="FFCB1007901BFFC5FD2EFD828AE1AAAE" box="[718,935,634,660]" pageId="10" pageNumber="453" value="2007-11-24">
<collectingDate id="EF8FE9EF901BFFC5FD2EFD828AE1AAAE" box="[718,935,634,660]" pageId="10" pageNumber="453" value="2007-11-24">24 November 2007</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="3B1D3C9A901BFFC5FC52FD828D8CAA82" collectingDate="1988-11-21" collectingDateMax="2007-11-24" collectingDateMin="1972-01-19" collectionCode="ZIN" country="South Africa" county="Vityaz" elevation="490" latitude="-21.266666" location="st. 2522" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." pageId="10" pageNumber="453" specimenCode="51521" specimenCount="1" typeStatus="holotype">
<specimenCount id="9D73FD4E901BFFC5FC52FD828D75AAAE" box="[946,1075,634,660]" count="1" pageId="10" pageNumber="453" type="generic">1 specimen</specimenCount>
,
<collectionCode id="ED64AE02901BFFC5FBDDFD828D2AAAAE" box="[1085,1132,634,660]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="453" type="Museum">ZIN</collectionCode>
<quantity id="4C8D9B22901BFFC5FB91FD828C66AAAE" box="[1137,1312,634,661]" metricMagnitude="4" metricUnit="m" metricValue="5.152120999999999" pageId="10" pageNumber="453" unit="mm" value="5.152121E7">
<specimenCode id="DBD39EBC901BFFC5FB91FD828DFCAAAF" box="[1137,1210,634,661]" pageId="10" pageNumber="453">51521</specimenCode>
, 210 mm
</quantity>
SL, Vityaz cruise 17,
<location id="8EAA601C901BFFC5FEEBFD668821AA83" LSID="urn:lsid:plazi:treatment:03DC87D1901BFFC3FF77FF6F8C16AB03:8EAA601C901BFFC5FEEBFD668821AA83" box="[267,359,670,697]" country="South Africa" county="Vityaz" latitude="-21.266666" longLatPrecision="1264" longitude="35.683334" municipality="Crosnier st." name="st. 2522" pageId="10" pageNumber="453">st. 2522</location>
,
<geoCoordinate id="EE415000901BFFC5FE92FD66888CAA82" box="[370,458,670,696]" degrees="21" direction="south" minutes="16" orientation="latitude" pageId="10" pageNumber="453" precision="925" value="-21.266666">21°16'S</geoCoordinate>
<geoCoordinate id="EE415000901BFFC5FE31FD668B6CAA83" box="[465,554,670,697]" degrees="35" direction="east" minutes="41" orientation="longitude" pageId="10" pageNumber="453" precision="925" value="35.683334">35°41'E</geoCoordinate>
(off eastern
<collectingCountry id="F3627657901BFFC5FD58FD668A0CAA82" box="[696,842,670,696]" name="South Africa" pageId="10" pageNumber="453">South Africa</collectingCountry>
),
<quantity id="4C8D9B22901BFFC5FCBDFD668A98AA82" box="[861,990,670,697]" metricMagnitude="2" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="10" pageNumber="453" unit="m" value="490.0" valueMax="500.0" valueMin="480.0">
<elevation id="0058D1F4901BFFC5FCBDFD668A98AA82" box="[861,990,670,697]" metricMagnitude="2" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="10" pageNumber="453" unit="m" value="490.0" valueMax="500.0" valueMin="480.0">480500 m</elevation>
</quantity>
,
<date id="FFCB1007901BFFC5FC0AFD668D83AA82" box="[1002,1221,670,696]" pageId="10" pageNumber="453" value="1988-11-21">
<collectingDate id="EF8FE9EF901BFFC5FC0AFD668D83AA82" box="[1002,1221,670,696]" pageId="10" pageNumber="453" value="1988-11-21">21 November 1988</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901BFFC5FF27FD398BF1ABAA" pageId="10" pageNumber="453" type="diagnosis">
<paragraph id="8BCA36C7901BFFC5FF27FD398BF1ABAA" blockId="10.[151,1437,417,2028]" pageId="10" pageNumber="453">
<emphasis id="B901EAD5901BFFC5FF27FD398804AAE1" bold="true" box="[199,322,705,731]" pageId="10" pageNumber="453">Diagnosis.</emphasis>
Precaudal vertebrae 1213, total vertebrae 5154; dorsal-fin rays 8892; anal-fin rays 6871; pectoral-fin rays 25; long gill rakers on first gill arch 4; pseudobranchials 45; 13 dorsal-fin rays in front of first vertebra; HL 27.127.6% of SL; body depth at vent 2122.5% of SL; predorsal length 2324.5% of SL; transversal scale rows above origin of anal fin 8590, and above lateral line 2223; broad expansion on preopercular angle with two spines, fleshy flap on hind margin of preopercle above spine 2; otoliths&gt;
<quantity id="4C8D9B22901BFFC5FBE4FCAA8D15AB56" box="[1028,1107,850,877]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="453" unit="mm" value="15.0">15 mm</quantity>
in length: OL:OH = 2.52.7, dorsal rim with rounded obtuse middorsal angle.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901BFFC5FF27FC618D12AEF6" pageId="10" pageNumber="453" type="description">
<paragraph id="8BCA36C7901BFFC5FF27FC6188EDABC6" blockId="10.[151,1437,417,2028]" pageId="10" pageNumber="453">
<emphasis id="B901EAD5901BFFC5FF27FC61881FAB89" bold="true" box="[199,345,921,947]" pageId="10" pageNumber="453">Description.</emphasis>
Relatively robust fish with stout tapering tail and rounded snout.Maximal size of fishes investigated
<quantity id="4C8D9B22901BFFC5FF77FC4689B3ABE2" box="[151,245,958,984]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="453" unit="mm" value="420.0">420 mm</quantity>
SL (
<typeStatus id="54CE8865901BFFC5FECBFC4688D2ABE2" box="[299,404,958,984]" pageId="10" pageNumber="453" type="holotype">holotype</typeStatus>
). Preopercular rim with two blunt spines positioned at bend, fleshy flap above 2
<superScript id="7C009B8F901BFFC5FAF9FC458C6FABF1" attach="left" box="[1305,1321,957,971]" fontSize="6" pageId="10" pageNumber="453">nd</superScript>
spine and no spine further upward.
</paragraph>
<paragraph id="8BCA36C7901BFFC5FF30FBFE8D0BACB6" blockId="10.[151,1437,417,2028]" pageId="10" pageNumber="453">
Meristics (
<specimenCount id="9D73FD4E901BFFC5FEA9FBFE889DAC1A" box="[329,475,1030,1056]" count="6" pageId="10" pageNumber="453" type="generic">6 specimens</specimenCount>
): precaudal vertebrae 13 (1213); total vertebrae 53 (5154); dorsal-fin rays 91 (88 92); anal-fin rays 69 (6871); pectoral-fin rays 25; caudal-fin rays 10; D/V = -2 (-1 -3); D/A = 28 (2328); V/A = 19 (1519); long gill rakers on first gill arch 4; pseudobranchial filaments 4 (45); transversal scale rows above origin of anal fin 8590, and above lateral line 2223; pyloric coeca unable to count.
</paragraph>
<paragraph id="8BCA36C7901BFFC5FF27FB6E8BB7AD27" blockId="10.[151,1437,417,2028]" pageId="10" pageNumber="453">
Morphometrics (
<specimenCount id="9D73FD4E901BFFC5FE64FB6E8B49AC8A" box="[388,527,1174,1200]" count="3" pageId="10" pageNumber="453" type="generic">3 specimens</specimenCount>
) in % of SL: HL 27.127.6; body depth at vent 21 (2122.5); diameter of eye 4.1 (4.14.8); upper jaw length 15 (1215); interorbital width 7.1 (7.17.5); pelvic-fin base to anal-fin base 24 (2426); predorsal length 24.5 (22.924.5); preanal length 52 (46.252); prepelvic length 27.5 (22.927.5); pectoral-fin length 13.5 (1215.7); pelvic-fin length 12 (1212.5).
</paragraph>
<paragraph id="8BCA36C7901BFFC5FF27FADE8B87AE92" blockId="10.[151,1437,417,2028]" pageId="10" pageNumber="453">
Otolith morphology (
<specimenCount id="9D73FD4E901BFFC5FE5BFADE8B08AD7A" box="[443,590,1318,1344]" count="4" pageId="10" pageNumber="453" type="generic">4 specimens</specimenCount>
). Size up to
<quantity id="4C8D9B22901BFFC5FD02FADE8A75AD7A" box="[738,819,1318,1344]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="453" unit="mm" value="22.0">22 mm</quantity>
in length (
<typeStatus id="54CE8865901BFFC5FC53FADE8D5DAD7A" box="[947,1051,1318,1344]" pageId="10" pageNumber="453" type="holotype">holotype</typeStatus>
); OL:OH =
<quantity id="4C8D9B22901BFFC5FB4AFADE8C58AD7A" box="[1194,1310,1318,1345]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" metricValueMax="6.858" metricValueMin="6.35" pageId="10" pageNumber="453" unit="in" value="2.6" valueMax="2.7" valueMin="2.5">2.52.7 in</quantity>
specimens larger than
<quantity id="4C8D9B22901BFFC5FEF8FAB28821AD5E" box="[280,359,1354,1381]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="453" unit="mm" value="15.0">15 mm</quantity>
in length,
<quantity id="4C8D9B22901BFFC5FE01FAB28B64AD5E" box="[481,546,1354,1380]" metricMagnitude="-2" metricUnit="m" metricValue="4.064" pageId="10" pageNumber="453" unit="in" value="1.6">1.6 in</quantity>
specimens smaller than
<quantity id="4C8D9B22901BFFC5FCD8FAB28A3FAD5E" box="[824,889,1354,1380]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="453" unit="mm" value="7.0">7 mm</quantity>
in length, and single intermediate form with 2.3 at about
<quantity id="4C8D9B22901BFFC5FF1BFA96880AADB2" box="[251,332,1390,1416]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="453" unit="mm" value="12.0">12 mm</quantity>
in length; OH:OT = 2.73.0. Thin, oval, elongated, anteriorly moderately projecting, posteriorly distinctly pointed, anterior and posterior tips inferior. Dorsal rim with broad, rounded, obtuse mediodorsal angle in large specimens and shallow ventral rim. All rims smooth in large specimens (&gt;
<quantity id="4C8D9B22901BFFC5FB80FA4E8DF5ADEA" box="[1120,1203,1462,1489]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="453" unit="mm" value="15.0">15 mm</quantity>
in length) intensely crenulated in smaller specimens (&lt;
<quantity id="4C8D9B22901BFFC5FDC1FA228B37ADCE" box="[545,625,1498,1524]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="453" unit="mm" value="12.0">12 mm</quantity>
in length). Inner face slightly bent in horizontal direction, nearly flat in vertical direction, with very long, undivided sulcus reaching close to anterior and posterior rims of otolith. Sulcus with shallow, undivided, uniform colliculum with flat to slightly concave dorsal rim and slightly convex ventral rim and rounded anterior and posterior tips. OL:CL = 1.31.35. Dorsal field with broad, indistinct depression; ventral field smooth, occasionally with faint central furrow. Outer face smooth in large specimens, with many radial furrows and central tubercles in small specimens.
</paragraph>
<paragraph id="8BCA36C7901BFFC5FF27F94A8D12AEF6" blockId="10.[151,1437,417,2028]" box="[199,1108,1714,1740]" pageId="10" pageNumber="453">Coloration. Live coloration unknown. Color of preserved specimens light brown.</paragraph>
</subSubSection>
<subSubSection id="C36F654C901BFFC3FF27F92D8DB0AABF" lastPageId="12" lastPageNumber="455" pageId="10" pageNumber="453" type="discussion">
<paragraph id="8BCA36C7901BFFC3FF27F92D8DFDA997" blockId="10.[151,1437,417,2028]" lastBlockId="12.[151,1437,150,825]" lastPageId="12" lastPageNumber="455" pageId="10" pageNumber="453">
<emphasis id="B901EAD5901BFFC5FF27F92D880AAED5" bold="true" box="[199,332,1749,1775]" pageId="10" pageNumber="453">Discussion.</emphasis>
<emphasis id="B901EAD5901BFFC5FEB3F92E8BDDAECA" box="[339,667,1750,1776]" italics="true" pageId="10" pageNumber="453">
Pyconocraspedum
<taxonomicName id="4C754D44901BFFC5FDC8F92E8BDDAECA" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[552,667,1750,1776]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="africanum">africanum</taxonomicName>
</emphasis>
is the largest species of the genus so far known, with the
<typeStatus id="54CE8865901BFFC5FAFCF92E8C38AECA" box="[1308,1406,1750,1776]" pageId="10" pageNumber="453" type="holotype">holotype</typeStatus>
of
<quantity id="4C8D9B22901BFFC5FF77F90289B4AF2E" box="[151,242,1786,1812]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="453" unit="mm" value="420.0">420 mm</quantity>
of SL. The species differs from the type species
<taxonomicName id="4C754D44901BFFC5FCE3F9028D1EAF2E" authorityName="Alcock" authorityYear="1889" box="[771,1112,1786,1812]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901BFFC5FCE3F9028D1EAF2E" box="[771,1112,1786,1812]" italics="true" pageId="10" pageNumber="453">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
in the higher number of anal-fin rays (6871 vs. 5765), the higher number of pseudobranchial filaments (45 vs. 12), the more slender shape (depth at vent in % of SL 2122 vs. 1718) and the shorter head (27.127.6 vs. 29.631.5 % of SL).
<taxonomicName id="4C754D44901BFFC5FB3BF8BA884CAFBA" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901BFFC5FB3BF8BA884CAFBA" italics="true" pageId="10" pageNumber="453">Pycnocraspedum africanum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C754D44901BFFC5FE43F89F8B4AAFBA" authorityName="Machida" authorityYear="1984" box="[419,524,1894,1920]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="fulvum">
<emphasis id="B901EAD5901BFFC5FE43F89F8B4AAFBA" box="[419,524,1894,1920]" italics="true" pageId="10" pageNumber="453">P. fulvum</emphasis>
</taxonomicName>
in the higher number of dorsal-fin rays (8892 vs. 78) and anal-fin rays (6871 vs. 63).
<taxonomicName id="4C754D44901BFFC5FF13F8728B69AF9E" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[243,559,1930,1956]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901BFFC5FF13F8728B69AF9E" box="[243,559,1930,1956]" italics="true" pageId="10" pageNumber="453">Pycnocraspedum africanum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C754D44901BFFC5FD25F8738A00AF9E" authorityName=", Gosline" authorityYear="1954" box="[709,838,1931,1956]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901BFFC5FD25F8738A00AF9E" box="[709,838,1931,1956]" italics="true" pageId="10" pageNumber="453">P. armatum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D44901BFFC5FC9FF8738D5AAF9E" baseAuthorityName="Parr" baseAuthorityYear="1933" box="[895,1052,1930,1956]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD5901BFFC5FC9FF8738D5AAF9E" box="[895,1052,1930,1956]" italics="true" pageId="10" pageNumber="453">P. phyllosoma</emphasis>
</taxonomicName>
in the lower number of dorsal-fin rays (8892 vs. 95103).
<taxonomicName id="4C754D44901BFFC5FE5AF8568BB0AFF2" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[442,758,1966,1992]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901BFFC5FE5AF8568BB0AFF2" box="[442,758,1966,1992]" italics="true" pageId="10" pageNumber="453">Pycnocraspedum africanum</emphasis>
</taxonomicName>
resembles closest
<taxonomicName id="4C754D44901BFFC5FC2BF8578D2DAFF2" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[971,1131,1966,1992]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="10" pageNumber="453" phylum="Chordata" rank="species" species="rowleyensis" status="sp. nov.">
<emphasis id="B901EAD5901BFFC5FC2BF8578D2DAFF2" box="[971,1131,1966,1992]" italics="true" pageId="10" pageNumber="453">P. rowleyensis</emphasis>
</taxonomicName>
<emphasis id="B901EAD5901BFFC5FB93F8568D88AFF2" bold="true" box="[1139,1230,1966,1992]" pageId="10" pageNumber="453">
<taxonomicNameLabel id="A23257AE901BFFC5FB93F8568D88AFF2" box="[1139,1230,1966,1992]" pageId="10" pageNumber="453" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from off northern
<collectingRegion id="49B1F825901BFFC5FF77F82A8819AFD6" box="[151,351,2002,2028]" country="Australia" name="Western Australia" pageId="10" pageNumber="453">Western Australia</collectingRegion>
in the number of dorsal-fin rays and the presence of only 2 spines on the hind margin of the preopercle, but differs in the lower number of transversal scale rows (8590 vs. 70), scale rows above the lateral line (2223 vs. 17), the presence of a flashy flap above the lower preopercular spines (vs. concave section) and the more elongate otoliths (OL:OH = 2.52.7 vs. 2.1).
<taxonomicName id="4C754D44901DFFC3FD7EFF278A9DA8C3" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[670,987,223,249]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901DFFC3FD7EFF278A9DA8C3" box="[670,987,223,249]" italics="true" pageId="12" pageNumber="455">Pycnocraspedum africanum</emphasis>
</taxonomicName>
differs from all other
<taxonomicName id="4C754D44901DFFC3FB3BFF2789AEA927" pageId="12" pageNumber="455">
<emphasis id="B901EAD5901DFFC3FB3BFF278CDAA8C3" box="[1243,1436,223,249]" italics="true" pageId="12" pageNumber="455">Pycnocraspedum</emphasis>
species
</taxonomicName>
in the high number of transversal scale rows above the origin of the anal fin of 8590 (vs. 5278). The otoliths of
<taxonomicName id="4C754D44901DFFC3FEF2FEDF88E4A97B" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[274,418,295,321]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901DFFC3FEF2FEDF88E4A97B" box="[274,418,295,321]" italics="true" pageId="12" pageNumber="455">P. africanum</emphasis>
</taxonomicName>
are slenderer than those of
<taxonomicName id="4C754D44901DFFC3FD3DFEDF8A19A97A" authorityName=", Gosline" authorityYear="1954" box="[733,863,295,320]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901DFFC3FD3DFEDF8A19A97A" box="[733,863,295,320]" italics="true" pageId="12" pageNumber="455">P. armatum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D44901DFFC3FC77FEDF8D72A97B" baseAuthorityName="Parr" baseAuthorityYear="1933" box="[919,1076,295,321]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD5901DFFC3FC77FEDF8D72A97B" box="[919,1076,295,321]" italics="true" pageId="12" pageNumber="455">P. phyllosoma</emphasis>
</taxonomicName>
(OL:OH = 2.52.7 vs. 2.02.3) and are less strongly bent; and they differ from those of
<taxonomicName id="4C754D44901DFFC3FCF6FEB38A84A95E" authorityName="Alcock" authorityYear="1889" box="[790,962,331,356]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901DFFC3FCF6FEB38A84A95E" box="[790,962,331,356]" italics="true" pageId="12" pageNumber="455">P. squamipinne</emphasis>
</taxonomicName>
in the long sulcus (OL:CL = 1.31.35 vs. 1.51.6). The otoliths of
<taxonomicName id="4C754D44901DFFC3FE4AFE978B71A9B3" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[426,567,367,393]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901DFFC3FE4AFE978B71A9B3" box="[426,567,367,393]" italics="true" pageId="12" pageNumber="455">P. africanum</emphasis>
</taxonomicName>
are similar to those of the fossil
<taxonomicName id="4C754D44901DFFC3FC7FFE978C73A9B3" authority="(Schwarzhans, 1978)" baseAuthorityName="Schwarzhans" baseAuthorityYear="1978" box="[927,1333,367,393]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="cetonanense">
<emphasis id="B901EAD5901DFFC3FC7FFE978D05A9B2" box="[927,1091,367,392]" italics="true" pageId="12" pageNumber="455">P. cetonanense</emphasis>
(
<bibRefCitation id="EFE44B36901DFFC3FBB1FE978C6AA9B3" author="Schwarzhans, W." box="[1105,1324,367,393]" pageId="12" pageNumber="455" pagination="1 - 52" refId="ref19902" refString="Schwarzhans, W. (1978) Otolithen aus dem Unter-Pliozan von Sud-Sizilien und aus der Toscana. Berliner Geowissenschaftliche Abhandlungen, 8, 1-52." type="journal article" year="1978">Schwarzhans, 1978</bibRefCitation>
)
</taxonomicName>
differing mainly in the narrower sulcus and the single, undivided colliculum (vs. two separate colliculi).
</paragraph>
<caption id="DF0A664F901AFFC4FF77F9028D6DAFD0" ID-DOI="http://doi.org/10.5281/zenodo.15561739" ID-Zenodo-Dep="15561739" httpUri="https://zenodo.org/record/15561739/files/figure.png" pageId="11" pageNumber="454" startId="11.[151,250,1786,1810]" targetBox="[256,1328,181,1748]" targetPageId="11" targetType="figure">
<paragraph id="8BCA36C7901AFFC4FF77F9028D6DAFD0" blockId="11.[151,1437,1786,2026]" pageId="11" pageNumber="454">
<emphasis id="B901EAD5901AFFC4FF77F9028854AF28" bold="true" box="[151,274,1786,1810]" pageId="11" pageNumber="454">FIGURE 3.</emphasis>
Detail figures and drawings of preopercular spines and lower preopercular pores in
<taxonomicName id="4C754D44901AFFC4FB81F9028C49AF28" authorityName="Alcock" authorityYear="1889" box="[1121,1295,1786,1810]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="454" phylum="Chordata" rank="genus">
<emphasis id="B901EAD5901AFFC4FB81F9028C49AF28" box="[1121,1295,1786,1810]" italics="true" pageId="11" pageNumber="454">Pycnocraspedum</emphasis>
</taxonomicName>
; preopercular spines are numbered from 1 through 4 from the lower angle of the preoperculum upwards; abbreviation lpp = lower preopercular pore.
<emphasis id="B901EAD5901AFFC4FF32F8BA885AAF60" bold="true" box="[210,284,1858,1882]" pageId="11" pageNumber="454">3A3B</emphasis>
)
<taxonomicName id="4C754D44901AFFC4FECCF8BA8BD3AF60" authority=", USNM" authorityName="USNM" box="[300,661,1858,1882]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="454" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901AFFC4FECCF8BA8B7BAF60" box="[300,573,1858,1882]" italics="true" pageId="11" pageNumber="454">Pycnocraspedum armatum</emphasis>
, USNM
</taxonomicName>
227412, 3A1-3A2 specimen 1 left side, 3A3 specimen 1 right side (mirror imaged), 3B specimen 2;
<emphasis id="B901EAD5901AFFC4FE7AF89E88A3AF44" bold="true" box="[410,485,1894,1918]" pageId="11" pageNumber="454">3C3D</emphasis>
)
<taxonomicName id="4C754D44901AFFC4FE13F89E8A5BAF44" baseAuthorityName="Parr" baseAuthorityYear="1933" box="[499,797,1894,1918]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="454" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD5901AFFC4FE13F89E8A5BAF44" box="[499,797,1894,1918]" italics="true" pageId="11" pageNumber="454">Pycnocraspedum phyllosoma</emphasis>
</taxonomicName>
, 3C USNM 421587, 3D USNM 421586;
<emphasis id="B901EAD5901AFFC4FB22F89E8D99AF44" bold="true" box="[1218,1247,1894,1918]" pageId="11" pageNumber="454">3E</emphasis>
)
<taxonomicName id="4C754D44901AFFC4FB0EF89E899AAF98" authorityName="Machida" authorityYear="1984" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="454" phylum="Chordata" rank="species" species="fulvum">
<emphasis id="B901EAD5901AFFC4FB0EF89E899AAF98" italics="true" pageId="11" pageNumber="454">Pycnocraspedum fulvum</emphasis>
</taxonomicName>
(holotype), BSKU 27497;
<emphasis id="B901EAD5901AFFC4FE0EF8728B7EAF98" bold="true" box="[494,568,1930,1954]" pageId="11" pageNumber="454">3F3G</emphasis>
)
<taxonomicName id="4C754D44901AFFC4FDA6F8728A25AF98" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[582,867,1930,1954]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="454" phylum="Chordata" rank="species" species="africanum" status="sp. nov.">
<emphasis id="B901EAD5901AFFC4FDA6F8728A25AF98" box="[582,867,1930,1954]" italics="true" pageId="11" pageNumber="454">Pycnocraspedum africanum</emphasis>
</taxonomicName>
<emphasis id="B901EAD5901AFFC4FC89F8728AFFAF98" bold="true" box="[873,953,1930,1954]" pageId="11" pageNumber="454">
<taxonomicNameLabel id="A23257AE901AFFC4FC89F8728AFFAF98" box="[873,953,1930,1954]" pageId="11" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 3F holotype, MNHN 2000-5691, 3G paratype, MNHN 2000-5692;
<emphasis id="B901EAD5901AFFC4FE93F85688D5AFFC" bold="true" box="[371,403,1966,1990]" pageId="11" pageNumber="454">3H</emphasis>
)
<taxonomicName id="4C754D44901AFFC4FE48F8568B9CAFFC" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[424,730,1966,1990]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="454" phylum="Chordata" rank="species" species="rowleyensis" status="sp. nov.">
<emphasis id="B901EAD5901AFFC4FE48F8568B9CAFFC" box="[424,730,1966,1990]" italics="true" pageId="11" pageNumber="454">Pycnocraspedum rowleyensis</emphasis>
</taxonomicName>
<emphasis id="B901EAD5901AFFC4FD07F8568A78AFFC" bold="true" box="[743,830,1966,1990]" pageId="11" pageNumber="454">
<taxonomicNameLabel id="A23257AE901AFFC4FD07F8568A78AFFC" box="[743,830,1966,1990]" pageId="11" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, WAM I.28059.021;
<emphasis id="B901EAD5901AFFC4FB75F8568D9FAFFC" bold="true" box="[1173,1241,1966,1990]" pageId="11" pageNumber="454">3I3K</emphasis>
)
<taxonomicName id="4C754D44901AFFC4FB0EF856885EAFD0" authorityName="Alcock" authorityYear="1889" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="11" pageNumber="454" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901AFFC4FB0EF856885EAFD0" italics="true" pageId="11" pageNumber="454">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
, 3I lectotype, BMNH 1890.7.31.1, 3J ZMUC P771871, 3K ZMUC P771737.
</paragraph>
</caption>
<paragraph id="8BCA36C7901DFFC3FF27FE4F8DB0AABF" blockId="12.[151,1437,150,825]" pageId="12" pageNumber="455">
The specimens here selected as
<typeStatus id="54CE8865901DFFC3FDD7FE4F8BDFA9EB" box="[567,665,439,465]" pageId="12" pageNumber="455" type="holotype">holotype</typeStatus>
and one of the
<typeStatus id="54CE8865901DFFC3FCB1FE4F8AFAA9EB" box="[849,956,439,465]" pageId="12" pageNumber="455" type="paratype">paratypes</typeStatus>
from MNHN have previously been listed by
<bibRefCitation id="EFE44B36901DFFC3FF5BFE2388D2A9CF" author="Fricke, R. &amp; Mahafina, J. &amp; Behivoke, F. &amp; Jaonalison, H. &amp; Leopold, M. &amp; Ponton, D." box="[187,404,475,501]" pageId="12" pageNumber="455" pagination="1 - 432" refId="ref18190" refString="Fricke, R., Mahafina, J., Behivoke, F., Jaonalison, H., Leopold, M. &amp; Ponton, D. (2018) Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa, 3 (1), 1-432." type="journal article" year="2018">
Fricke
<emphasis id="B901EAD5901DFFC3FEEAFE238804A9CF" box="[266,322,475,501]" italics="true" pageId="12" pageNumber="455">et al.</emphasis>
(2018)
</bibRefCitation>
as
<taxonomicName id="4C754D44901DFFC3FE5AFE238B23A9CE" authorityName="Alcock" authorityYear="1889" box="[442,613,475,500]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901DFFC3FE5AFE238B23A9CE" box="[442,613,475,500]" italics="true" pageId="12" pageNumber="455">P. squamipinne</emphasis>
</taxonomicName>
. Another specimen here listed as
<typeStatus id="54CE8865901DFFC3FC01FE238D07A9CF" box="[993,1089,475,501]" pageId="12" pageNumber="455" type="paratype">paratype</typeStatus>
from ZIN has been tentatively attributed to
<taxonomicName id="4C754D44901DFFC3FED0FE078899AA22" authorityName="Alcock" authorityYear="1889" box="[304,479,511,536]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901DFFC3FED0FE078899AA22" box="[304,479,511,536]" italics="true" pageId="12" pageNumber="455">P. squamipinne</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE44B36901DFFC3FDF2FE078B9CAA23" author="Prokofiev, A. M." box="[530,730,510,537]" pageId="12" pageNumber="455" pagination="111 - 128" refId="ref19532" refString="Prokofiev, A. M. (2005) On some rare ophidiiform fishes from the South Atlantic and Indo-West Pacific, with erection of a new genus, Megacataetyx gen. novum (Teleostei: Ophidiiformes). Estestvennye i Tekhnicheskie Nauki, 2, 111-128." type="journal article" year="2005">Prokofiev (2005)</bibRefCitation>
. He noticed certain differences from typical
<taxonomicName id="4C754D44901DFFC3FB0DFE078CDAAA22" authorityName="Alcock" authorityYear="1889" box="[1261,1436,511,536]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901DFFC3FB0DFE078CDAAA22" box="[1261,1436,511,536]" italics="true" pageId="12" pageNumber="455">P. squamipinne</emphasis>
</taxonomicName>
specimens such as the presence of 4 pseudobranchial filaments (vs. 12) and a more compressed, shorter head. It is possible that a large specimen recorded by
<bibRefCitation id="EFE44B36901DFFC3FD53FDBF8ACFAA5B" author="Teena, T. K. &amp; Murugan, A. &amp; Kumar, A. T. T. &amp; Lal, K. K." box="[691,905,583,609]" pageId="12" pageNumber="455" pagination="77 - 83" refId="ref20468" refString="Teena, T. K., Murugan, A., Kumar, A. T. T. &amp; Lal, K. K. (2021) Redescription of a rare cusk eel, Pycnocraspedum squamipinne (Actinopterygii, Ophidiiformes, Ophidiidae) from Bay of Bengal. Acta Ichthyologica et Piscatoria, 51 (1), 77-83. https://doi.org/10.3897/aiep.51.63469" type="journal article" year="2021">
Teena
<emphasis id="B901EAD5901DFFC3FD1EFDBF8A71AA5B" box="[766,823,583,609]" italics="true" pageId="12" pageNumber="455">et al.</emphasis>
(2021)
</bibRefCitation>
from off Chennai coast, southern
<collectingCountry id="F3627657901DFFC3FAEEFDBF8C0CAA5B" box="[1294,1354,583,609]" name="India" pageId="12" pageNumber="455">India</collectingCountry>
, with a suspiciously high number of transversal scale rows could potentially represent
<taxonomicName id="4C754D44901DFFC3FBEBFD938DDFAABF" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[1035,1177,619,645]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901DFFC3FBEBFD938DDFAABF" box="[1035,1177,619,645]" italics="true" pageId="12" pageNumber="455">P. africanum</emphasis>
</taxonomicName>
as well.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901DFFC3FF27FD768B5EAB2F" pageId="12" pageNumber="455" type="distribution">
<paragraph id="8BCA36C7901DFFC3FF27FD768B5EAB2F" blockId="12.[151,1437,150,825]" pageId="12" pageNumber="455">
<emphasis id="B901EAD5901DFFC3FF27FD768827AA92" bold="true" box="[199,353,654,680]" pageId="12" pageNumber="455">Distribution.</emphasis>
<taxonomicName id="4C754D44901DFFC3FE88FD778BE4AA93" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[360,674,655,681]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901DFFC3FE88FD778BE4AA93" box="[360,674,655,681]" italics="true" pageId="12" pageNumber="455">Pycnocraspedum africanum</emphasis>
</taxonomicName>
is a relatively deep dwelling species at
<quantity id="4C8D9B22901DFFC3FB80FD768C4DAA93" box="[1120,1291,654,681]" metricMagnitude="-1" metricUnit="m" metricValue="5.275" metricValueMax="6.05" metricValueMin="4.5" pageId="12" pageNumber="455" unit="mm" value="527.5" valueMax="605.0" valueMin="450.0">450 to 605 mm</quantity>
depth, while all other species mostly occur between 300 and
<quantity id="4C8D9B22901DFFC3FD4CFD4A8BB5AAF7" box="[684,755,690,717]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="455" unit="m" value="500.0">500 m</quantity>
(
<bibRefCitation id="EFE44B36901DFFC3FCE1FD4B8A9FAAF7" author="Nielsen, J. G. &amp; Cohen, D. M. &amp; Markle, D. F. &amp; Robins, C. R." box="[769,985,691,717]" pageId="12" pageNumber="455" refId="ref19264" refString="Nielsen, J. G., Cohen, D. M., Markle, D. F. &amp; Robins, C. R. (1999) An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiiform fishes known to date. FAO Fisheries Synopsis No. 125. Vol. 18. FAO, Rome, 178 pp." type="book" year="1999">
Nielsen
<emphasis id="B901EAD5901DFFC3FCBEFD4B8AD0AAF7" box="[862,918,691,717]" italics="true" pageId="12" pageNumber="455">et al.</emphasis>
1999
</bibRefCitation>
).
<taxonomicName id="4C754D44901DFFC3FC0BFD4B8C60AAF7" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[1003,1318,691,717]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901DFFC3FC0BFD4B8C60AAF7" box="[1003,1318,691,717]" italics="true" pageId="12" pageNumber="455">Pycnocraspedum africanum</emphasis>
</taxonomicName>
occurs off the East African coast from
<geoCoordinate id="EE415000901DFFC3FE36FD2F8B4AAACB" box="[470,524,727,753]" degrees="11" direction="south" orientation="latitude" pageId="12" pageNumber="455" precision="55555" value="-11.0">11°S</geoCoordinate>
to
<geoCoordinate id="EE415000901DFFC3FDD2FD2F8B2DAACB" box="[562,619,727,753]" degrees="22" direction="south" orientation="latitude" pageId="12" pageNumber="455" precision="55555" value="-22.0">22°S</geoCoordinate>
, but could be more widely spread in the Indian Ocean subject to review of previously recorded specimens.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901DFFC3FF27FCE78C16AB03" box="[199,1360,799,825]" pageId="12" pageNumber="455" type="etymology">
<paragraph id="8BCA36C7901DFFC3FF27FCE78C16AB03" blockId="12.[151,1437,150,825]" box="[199,1360,799,825]" pageId="12" pageNumber="455">
<emphasis id="B901EAD5901DFFC3FF27FCE78808AB03" bold="true" box="[199,334,799,825]" pageId="12" pageNumber="455">Etymology.</emphasis>
Named after the continent of Africa from where the studied specimens have been obtained.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,711 @@
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<taxonomicName id="4C754D44901DFFC3FF77FC728BCAAB9E" ID-CoL="4QM7X" authority="Gosline, 1954" authorityName=", Gosline" authorityYear="1954" box="[151,652,906,933]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="armatum">
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<emphasis id="B901EAD5901DFFC3FF77FC72889AAB9F" bold="true" box="[151,476,906,933]" italics="true" pageId="12" pageNumber="455">Pycnocraspedum armatum</emphasis>
<bibRefCitation id="EFE44B36901DFFC3FE04FC728BCAAB9E" author="Gosline, W. A." box="[484,652,906,932]" pageId="12" pageNumber="455" pagination="68 - 83" refId="ref18451" refString="Gosline, W. A. (1954) Fishes killed by the 1950 eruption of Mauna Loa. Brotulidae. Pacific Science, 8, 68-83." type="journal article" year="1954">Gosline, 1954</bibRefCitation>
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<figureCitation id="134E2A42901DFFC3FF77FC578846ABF3" box="[151,256,943,969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1527,1551]" captionTargetBox="[168,1411,202,1477]" captionTargetId="figure-17@3.[151,1436,181,1504]" captionTargetPageId="3" captionText="FIGURE 1. Radiograph figures of selected specimens of the genus Pycnocraspedum to visualize position of dorsal-fin rays in relation to first vertebra; “1” denotes the first dorsal-fin ray above the first vertebra; numbers with negative denotation represent dorsal-fin rays in front of the first vertebra. 1A) Pycnocraspedum armatum, USNM 227390; 1B) Pycnocraspedum phyllosoma USNM 227413; 1C) Pycnocraspedum squamipinne, ZMUC P771871; 1D) Pycnocraspedum fulvum (holotype), BSKU 27497." figureDoi="http://doi.org/10.5281/zenodo.15561735" httpUri="https://zenodo.org/record/15561735/files/figure.png" pageId="12" pageNumber="455">Figures 1</figureCitation>
,
<figureCitation id="134E2A42901DFFC3FEEEFC578858ABF3" box="[270,286,943,969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1457,1481]" captionTargetBox="[151,1436,181,1432]" captionTargetId="figure-23@4.[151,1436,181,1433]" captionTargetPageId="4" captionText="FIGURE 2. Gill rakers, pseudobranchial filaments and dentition details. 2A) Extracted first gill arch from Pycnocraspedum squamipinne, ZMUC P771871; 2B, 2E) Dentition of premaxilla, vomer and palatines of Pycnocraspedum armatum, USNM 227412, 2E is close-up to show sandpaper like texture of granular teeth; 2C2D) Pseudobranchial filaments in Pycnocraspedum armatum, USNM 227412." figureDoi="http://doi.org/10.5281/zenodo.15561737" httpUri="https://zenodo.org/record/15561737/files/figure.png" pageId="12" pageNumber="455">2</figureCitation>
,
<figureCitation id="134E2A42901DFFC3FECAFC57887FABF3" box="[298,313,943,969]" captionStart="FIGURE 3" captionStartId="11.[151,250,1786,1810]" captionTargetBox="[256,1328,181,1748]" captionTargetId="figure-17@11.[256,1331,181,1762]" captionTargetPageId="11" captionText="FIGURE 3. Detail figures and drawings of preopercular spines and lower preopercular pores in Pycnocraspedum; preopercular spines are numbered from 1 through 4 from the lower angle of the preoperculum upwards; abbreviation lpp = lower preopercular pore. 3A3B) Pycnocraspedum armatum, USNM 227412, 3A1-3A2 specimen 1 left side, 3A3 specimen 1 right side (mirror imaged), 3B specimen 2; 3C3D) Pycnocraspedum phyllosoma, 3C USNM 421587, 3D USNM 421586; 3E) Pycnocraspedum fulvum (holotype), BSKU 27497; 3F3G) Pycnocraspedum africanum sp. nov., 3F holotype, MNHN 2000-5691, 3G paratype, MNHN 2000-5692; 3H) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021; 3I3K) Pycnocraspedum squamipinne, 3I lectotype, BMNH 1890.7.31.1, 3J ZMUC P771871, 3K ZMUC P771737." figureDoi="http://doi.org/10.5281/zenodo.15561739" httpUri="https://zenodo.org/record/15561739/files/figure.png" pageId="12" pageNumber="455">3</figureCitation>
,
<figureCitation id="134E2A42901DFFC3FEA6FC568810ABF3" box="[326,342,942,969]" captionStart="FIGURE 5" captionStartId="14.[151,250,1750,1774]" captionTargetBox="[245,1358,193,1715]" captionTargetId="figure-23@14.[222,1366,181,1726]" captionTargetPageId="14" captionText="FIGURE 5. Otoliths, inner faces (if not noted otherwise) shown from right side. 5A5C) Pycnocraspedum africanum sp. nov., 5A holotype, MNHN 2000-5691 (5A2 dorsal view), 5B paratype MNHN 2000-5692, 5C tentatively assigned specimen, IRSNB (5C2 dorsal view, 5C3 anterior view); 5D5E) Pycnocraspedum armatum, 5D USNM 227412 (5D2 ventral view), 5E USNM 227390 (5E2 ventral view); 5F) Pycnocraspedum fulvum (holotype), BSKU 27497 (5F2 ventral view, 5F3 anterior view); 5G) Pycnocraspedum rowleyensis sp. nov., holotype, WAM I.28059.021 (5G2 ventral view); 5H) Pycnocraspedum phyllosoma, USNM 421587 (5H2 ventral view, 5H3 anterior view); 5I5K) Pycnocraspedum squamipinne, 5I lectotype, BMNH 1890.7.31.1 (5I2 ventral view), 5J ZMUC P771871, 5L ZMUC P771739 (5L2 ventral view), 5K ZMUC P771738 (5K2 ventral view); 5M) †Pycnocraspedum cetonaense (holotype) SMF P.5694 (5M2 dorsal view, 5M3 anterior view)." figureDoi="http://doi.org/10.5281/zenodo.15561743" httpUri="https://zenodo.org/record/15561743/files/figure.png" pageId="12" pageNumber="455">5</figureCitation>
,
<figureCitation id="134E2A42901DFFC3FE82FC578834ABF3" box="[354,370,943,969]" captionStart="FIGURE 7" captionStartId="17.[151,250,1753,1777]" captionTargetBox="[151,1435,181,1730]" captionTargetId="figure-17@17.[151,1435,181,1730]" captionTargetPageId="17" captionText="FIGURE 7. Pycnocraspedum armatum Gosline, 1954; 7A, 7C) USNM 227412, 225 mm SL, 21.53°N 158.38°W, 229247 m; 7B) radiograph of holotype, USNM 162717, 310 mm SL, off Kona coast, Hawaii." figureDoi="http://doi.org/10.5281/zenodo.15561747" httpUri="https://zenodo.org/record/15561747/files/figure.png" pageId="12" pageNumber="455">7</figureCitation>
,
<figureCitation id="134E2A42901DFFC3FE9EFC5788DBABF3" box="[382,413,943,969]" captionStart="FIGURE 12" captionStartId="24.[151,250,899,923]" captionTargetBox="[153,1433,186,871]" captionTargetId="figure-23@24.[151,1436,181,875]" captionTargetPageId="24" captionText="FIGURE 12. Geographic distribution of the species of Pycnocraspedum. One symbol can represent more than one location." figureDoi="http://doi.org/10.5281/zenodo.15561755" httpUri="https://zenodo.org/record/15561755/files/figure.png" pageId="12" pageNumber="455">12</figureCitation>
;
<tableCitation id="C6F7037C901DFFC3FE48FC578B40ABF3" box="[424,518,943,969]" captionStart="TABLE 1" captionStartId="2.[152,244,151,175]" captionTargetBox="[163,1411,227,1782]" captionTargetId="graphics-48@2.[151,1437,256,1787]" captionText="TABLE 1. Selected keyparameters ofthecurrently 37recognizedophidiidgenera.Characterssharedwith Pycnocraspedum are highlighted. D/V = first dorsal-fin ray above vertebra." httpUri="http://table.plazi.org/id/DF0A664F9013FFCDFF78FF6F8BBBA8E9" pageId="12" pageNumber="455" tableUuid="DF0A664F9013FFCDFF78FF6F8BBBA8E9">Tables 1</tableCitation>
,
<tableCitation id="C6F7037C901DFFC3FDF4FC578B64ABF3" box="[532,546,943,969]" captionStart="TABLE 2" captionStartId="8.[152,244,720,744]" captionTargetPageId="8" captionText="TABLE 2. Meristic, morphometric and otolith characters of the species of the genus Pycnocraspedum. For abbreviations" httpUri="http://table.plazi.org/id/DF0A664F9019FFC7FF78FD288B4AAB36" pageId="12" pageNumber="455" tableUuid="DF0A664F9019FFC7FF78FD288B4AAB36">2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901DFFC3FF77FC0B8D55ACBC" pageId="12" pageNumber="455" type="reference_group">
<paragraph id="8BCA36C7901DFFC3FF77FC0B8CC7AC31" blockId="12.[151,1437,1011,1158]" box="[151,1409,1011,1035]" pageId="12" pageNumber="455">
<treatmentCitationGroup id="AB6511E9901DFFC3FF77FC0B8CC7AC31" box="[151,1409,1011,1035]" pageId="12" pageNumber="455">
<treatmentCitation id="0AD410D6901DFFC3FF77FC0B8BCDAC31" author="Gosline, W. A." box="[151,651,1011,1035]" page="81" pageId="12" pageNumber="455" year="1954">
<taxonomicName id="4C754D44901DFFC3FF77FC0B8BCDAC31" ID-CoL="4QM7X" authority="Gosline, 1954: 80 - 81" authorityName="Gosline" authorityPageNumber="80 - 81" authorityYear="1954" box="[151,651,1011,1035]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901DFFC3FF77FC0B88E1AC31" box="[151,423,1011,1035]" italics="true" pageId="12" pageNumber="455">Pycnocraspedum armatum</emphasis>
<bibRefCitation id="EFE44B36901DFFC3FE4EFC0B8BCDAC31" author="Gosline, W. A." box="[430,651,1011,1035]" pageId="12" pageNumber="455" pagination="68 - 83" refId="ref18451" refString="Gosline, W. A. (1954) Fishes killed by the 1950 eruption of Mauna Loa. Brotulidae. Pacific Science, 8, 68-83." type="journal article" year="1954">Gosline, 1954: 8081</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, tables 12, figs. 2e, 3e, off the Mauna Loa lava flow, Kona coast,
<collectingRegion id="49B1F825901DFFC3FAD0FC0B8C3AAC31" box="[1328,1404,1011,1035]" country="United States of America" name="Hawaii" pageId="12" pageNumber="455">Hawaii</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA36C7901DFFC3FF77FBEA8B08AC5D" blockId="12.[151,1437,1011,1158]" pageId="12" pageNumber="455">
<treatmentCitationGroup id="AB6511E9901DFFC3FF77FBEA8B08AC5D" pageId="12" pageNumber="455">
<taxonomicName id="4C754D44901DFFC3FF77FBEA8B78AC10" ID-CoL="4QM7X" authority="Gosline, 1954" authorityName="Gosline" authorityYear="1954" box="[151,574,1042,1066]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901DFFC3FF77FBEA88E1AC10" box="[151,423,1042,1066]" italics="true" pageId="12" pageNumber="455">Pycnocraspedum armatum</emphasis>
<bibRefCitation id="EFE44B36901DFFC3FE4DFBEA8B78AC10" author="Gosline, W. A." box="[429,574,1042,1066]" pageId="12" pageNumber="455" pagination="68 - 83" refId="ref18451" refString="Gosline, W. A. (1954) Fishes killed by the 1950 eruption of Mauna Loa. Brotulidae. Pacific Science, 8, 68-83." type="journal article" year="1954">Gosline, 1954</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="EFE44B36901DFFC3FDA7FBEA8A06AC10" author="Gosline, W. A. &amp; Brock, V. E." box="[583,832,1042,1066]" pageId="12" pageNumber="455" refId="ref18481" refString="Gosline, W. A. &amp; Brock, V. E. (1960) Handbook of Hawaiian fishes. University of Hawaii Press, Honolulu, 372 pp. https://doi.org/10.1515/9780824885090" type="book" year="1960">Gosline &amp; Brock (1960)</bibRefCitation>
,
<bibRefCitation id="EFE44B36901DFFC3FCA9FBEA8D02AC10" author="Cohen, D. M. &amp; Nielsen, J. G." box="[841,1092,1042,1066]" pageId="12" pageNumber="455" pagination="1 - 72" refId="ref17901" refString="Cohen, D. M. &amp; Nielsen, J. G. (1978) Guide to the identification of genera of the fish order Ophidiiformes with a tentative classification of the order. NOAA Technical Report NMFS Circular, 417, 1-72. https://doi.org/10.5962/bhl.title.63242" type="journal article" year="1978">Cohen &amp; Nielsen (1978)</bibRefCitation>
, Machida
<emphasis id="B901EAD5901DFFC3FB4CFBEA8D86AC13" box="[1196,1216,1042,1065]" italics="true" pageId="12" pageNumber="455">in</emphasis>
Okamura &amp; Kitajima (1984),
<bibRefCitation id="EFE44B36901DFFC3FEF8FBC88B55AC73" author="Chave, E. H. &amp; Mundy, B. C." box="[280,531,1072,1097]" pageId="12" pageNumber="455" pagination="367 - 409" refId="ref17780" refString="Chave, E. H. &amp; Mundy, B. C. (1994) Deep-sea benthic fish of the Hawaiian archipelago, Cross Seamount, and Johnson Atoll. Pacific Science, 48 (4), 367-409." type="journal article" year="1994">Chave &amp; Mundy (1994)</bibRefCitation>
,
<bibRefCitation id="EFE44B36901DFFC3FDFEFBC88BB2AC73" author="Nielsen, J. G. &amp; Cohen, D. M. &amp; Markle, D. F. &amp; Robins, C. R." box="[542,756,1072,1097]" pageId="12" pageNumber="455" refId="ref19264" refString="Nielsen, J. G., Cohen, D. M., Markle, D. F. &amp; Robins, C. R. (1999) An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiiform fishes known to date. FAO Fisheries Synopsis No. 125. Vol. 18. FAO, Rome, 178 pp." type="book" year="1999">
Nielsen
<emphasis id="B901EAD5901DFFC3FD92FBC98BE1AC72" box="[626,679,1072,1096]" italics="true" pageId="12" pageNumber="455">et al.</emphasis>
(1999)
</bibRefCitation>
,
<bibRefCitation id="EFE44B36901DFFC3FD1FFBC88AD3AC72" author="Mundy, B. C." box="[767,917,1072,1096]" pageId="12" pageNumber="455" pagination="1 - 704" refId="ref19115" refString="Mundy, B. C. (2005) Checklist of the fishes from the Hawaiian Archipelago. Bishop Museum Bulletin Zoology, (6), 1-704." type="journal article" year="2005">Mundy (2005)</bibRefCitation>
,
<bibRefCitation id="EFE44B36901DFFC3FC41FBC88D14AC72" author="Prokofiev, A. M." box="[929,1106,1072,1096]" pageId="12" pageNumber="455" pagination="111 - 128" refId="ref19532" refString="Prokofiev, A. M. (2005) On some rare ophidiiform fishes from the South Atlantic and Indo-West Pacific, with erection of a new genus, Megacataetyx gen. novum (Teleostei: Ophidiiformes). Estestvennye i Tekhnicheskie Nauki, 2, 111-128." type="journal article" year="2005">Prokofiev (2005)</bibRefCitation>
, Evseenko &amp; Okyama (2006),?
<bibRefCitation id="EFE44B36901DFFC3FF27FBB78831AC5D" author="Prokofiev, A. M." box="[199,375,1103,1127]" pageId="12" pageNumber="455" pagination="1019 - 1024" refId="ref19635" refString="Prokofiev, A. M. (2022) New data on the morphology and distribution of two rare species of Ophidiiformes: Diplacanthopoma japonicum (Bythitidae) and Pycnocraspedum microlepis (Neobythitidae). Voprosy Ikhtiologii, 62 (6), 680 - 685. [in Russian, English translation appeared in Journal of Ichthyology, 62 (6), 1019-1024] https://doi.org/10.1134/S0032945222060224" type="journal article" year="2022">Prokofiev (2022)</bibRefCitation>
,
<bibRefCitation id="EFE44B36901DFFC3FE62FBB78B0FAC5D" author="Girard, M. G. &amp; Mundy, B. C. &amp; Nonaka, A. &amp; Johnson, G. D." box="[386,585,1103,1127]" pageId="12" pageNumber="455" pagination="474 - 489" refId="ref18250" refString="Girard, M. G., Mundy, B. C., Nonaka, A. &amp; Johnson, G. D. (2023) Cusk-eel confusion: revisions of larval Luciobrotula and Pycnocraspedum and re-descriptions of two bythitid larvae (Ophidiiformes). Ichthyological Research, 70 (4), 474-489. https://doi.org/10.1007/s10228-023-00906-4" type="journal article" year="2023">
Girard
<emphasis id="B901EAD5901DFFC3FE2AFBA888BBAC5D" box="[458,509,1103,1127]" italics="true" pageId="12" pageNumber="455">et al.</emphasis>
(2023)
</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA36C7901DFFC3FF77FB968D55ACBC" blockId="12.[151,1437,1011,1158]" box="[151,1043,1134,1158]" pageId="12" pageNumber="455">
<taxonomicName id="4C754D44901DFFC3FF77FB96888BACBF" authorityName="Alcock" authorityYear="1889" box="[151,461,1134,1158]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901DFFC3FF77FB96888BACBF" box="[151,461,1134,1158]" italics="true" pageId="12" pageNumber="455">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
(non
<bibRefCitation id="EFE44B36901DFFC3FDE7FB968BD2ACBC" author="Alcock, A." box="[519,660,1134,1158]" pageId="12" pageNumber="455" pagination="376 - 399" refId="ref17472" refString="Alcock, A. (1889) Natural history notes from H. M. Indian Marine Survey Steamer &quot; Investigator &quot;. No. 13. On the bathybial fishes from the Bay of Bengal and neighbouring waters obtained during the seasons 1885 - 1889. Annals Magazine Natural History, Series 6, 4 (23), 376-399. https://doi.org/10.1080/00222938909460547" type="journal article" year="1889">Alcock, 1889</bibRefCitation>
):
<bibRefCitation id="EFE44B36901DFFC3FD45FB968A79ACBC" author="Nielsen, J. G." box="[677,831,1134,1158]" pageId="12" pageNumber="455" pagination="51 - 82" refId="ref19211" refString="Nielsen, J. G. (1997) Deepwater ophidiiform fishes from off New Caledonia with six new species. In: Seret, B. (Ed.), Resultats des Campagnes MUSORSTOM. Vol. 17. Memoires Museum national Histoire naturelle, 174, 51-82." type="journal article" year="1997">Nielsen (1997)</bibRefCitation>
,
<bibRefCitation id="EFE44B36901DFFC3FCA9FB968D56ACBC" author="Fricke, R. &amp; Kulbicki, M. &amp; Wantiez, L." box="[841,1040,1134,1158]" pageId="12" pageNumber="455" pagination="341 - 463" refId="ref18138" refString="Fricke, R., Kulbicki, M. &amp; Wantiez, L. (2011) Checklist of the fishes of New Caledonia, and their distribution in the Southwest Pacific Ocean (Pisces). Stuttgarter Beitrage zur Naturkunde A, Neue Serie, 4, 341-463." type="journal article" year="2011">
Fricke
<emphasis id="B901EAD5901DFFC3FC70FB978A82ACBC" box="[912,964,1134,1158]" italics="true" pageId="12" pageNumber="455">et al.</emphasis>
(2011)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901DFFC3FF77FB4C8B2CAF6D" pageId="12" pageNumber="455" type="materials_examined">
<paragraph id="8BCA36C7901DFFC3FF77FB4C8B2CAF6D" blockId="12.[151,1437,1204,2023]" pageId="12" pageNumber="455">
<emphasis id="B901EAD5901DFFC3FF77FB4C883EACF4" bold="true" box="[151,376,1204,1230]" pageId="12" pageNumber="455">Material examined</emphasis>
(
<specimenCount id="9D73FD4E901DFFC3FE67FB4C8B64ACF4" box="[391,546,1204,1230]" count="14" pageId="12" pageNumber="455" type="generic">14 specimens</specimenCount>
, 77.5334 mm SL): specimens originally identified as
<taxonomicName id="4C754D44901DFFC3FB61FB4D8C56ACF4" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[1153,1296,1204,1230]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901DFFC3FB61FB4D8C56ACF4" box="[1153,1296,1204,1230]" italics="true" pageId="12" pageNumber="455">P. microlepis</emphasis>
</taxonomicName>
:
<materialsCitation id="3B1D3C9A901DFFC3FAFEFB4C8835AD2D" collectingDate="1955-12-27" collectionCode="BSKU" collectorName="Mimase Fish Market" country="Japan" location="City" municipality="Shikoku Island" pageId="12" pageNumber="455" specimenCount="1" stateProvince="Kochi Prefecture">
<specimenCount id="9D73FD4E901DFFC3FAFEFB4C8CDAACF4" box="[1310,1436,1204,1230]" count="1" pageId="12" pageNumber="455" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE02901DFFC3FF77FB2089A4ACC8" box="[151,226,1240,1266]" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="12" pageNumber="455">BSKU</collectionCode>
5055, 234 mm SL,
<collectorName id="26805311901DFFC3FE5DFB208BEEACC8" box="[445,680,1240,1266]" pageId="12" pageNumber="455">Mimase Fish Market</collectorName>
, Kochi
<location id="8EAA601C901DFFC3FD1DFB208A68ACC8" LSID="urn:lsid:plazi:treatment:03DC87D1901DFFC0FF77FC728D63AFAF:8EAA601C901DFFC3FD1DFB208A68ACC8" box="[765,814,1240,1266]" country="Japan" municipality="Shikoku Island" name="City" pageId="12" pageNumber="455" stateProvince="Kochi Prefecture">City</location>
,
<collectingRegion id="49B1F825901DFFC3FCD9FB208ABFACC8" box="[825,1017,1240,1266]" country="Japan" name="Kochi" pageId="12" pageNumber="455">Kochi Prefecture</collectingRegion>
,
<collectingMunicipality id="6BAEACBD901DFFC3FBE4FB208DE8ACC8" box="[1028,1198,1240,1266]" pageId="12" pageNumber="455">Shikoku Island</collectingMunicipality>
,
<collectingCountry id="F3627657901DFFC3FB59FB218DBAACC9" box="[1209,1276,1241,1267]" name="Japan" pageId="12" pageNumber="455">Japan</collectingCountry>
, bottom trawl,
<date id="FFCB1007901DFFC3FF77FB048835AD2D" box="[151,371,1276,1303]" pageId="12" pageNumber="455" value="1955-12-27">
<collectingDate id="EF8FE9EF901DFFC3FF77FB048835AD2D" box="[151,371,1276,1303]" pageId="12" pageNumber="455" value="1955-12-27">27 December 1955</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A901DFFC3FE61FB048BA9AD00" collectingDate="1983-03-14" collectionCode="BSKU" collectorName="Mimase Fish Market" country="Japan" location="City" municipality="Shikoku Island" pageId="12" pageNumber="455" specimenCount="1" stateProvince="Kochi Prefecture">
<specimenCount id="9D73FD4E901DFFC3FE61FB048B44AD2C" box="[385,514,1276,1302]" count="1" pageId="12" pageNumber="455" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE02901DFFC3FDEAFB048B13AD2C" box="[522,597,1276,1302]" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="12" pageNumber="455">BSKU</collectionCode>
38643, 245 mm SL,
<collectorName id="26805311901DFFC3FCACFB048D7AAD2C" box="[844,1084,1276,1302]" pageId="12" pageNumber="455">Mimase Fish Market</collectorName>
, Kochi
<location id="8EAA601C901DFFC3FB76FB048D81AD2C" LSID="urn:lsid:plazi:treatment:03DC87D1901DFFC0FF77FC728D63AFAF:8EAA601C901DFFC3FB76FB048D81AD2C" box="[1174,1223,1276,1302]" country="Japan" municipality="Shikoku Island" name="City" pageId="12" pageNumber="455" stateProvince="Kochi Prefecture">City</location>
,
<collectingRegion id="49B1F825901DFFC3FB35FB048CDEAD2C" box="[1237,1432,1276,1302]" country="Japan" name="Kochi" pageId="12" pageNumber="455">Kochi Prefecture</collectingRegion>
,
<collectingMunicipality id="6BAEACBD901DFFC3FF77FAD88802AD00" box="[151,324,1312,1338]" pageId="12" pageNumber="455">Shikoku Island</collectingMunicipality>
,
<collectingCountry id="F3627657901DFFC3FEB0FAD988D5AD01" box="[336,403,1313,1339]" name="Japan" pageId="12" pageNumber="455">Japan</collectingCountry>
, bottom trawl,
<date id="FFCB1007901DFFC3FDDFFAD88BA9AD00" box="[575,751,1312,1338]" pageId="12" pageNumber="455" value="1983-03-14">
<collectingDate id="EF8FE9EF901DFFC3FDDFFAD88BA9AD00" box="[575,751,1312,1338]" pageId="12" pageNumber="455" value="1983-03-14">14 March 1983</collectingDate>
</date>
</materialsCitation>
; specimens originally identified as
<taxonomicName id="4C754D44901DFFC3FB60FAD98C44AD00" authorityName=", Gosline" authorityYear="1954" box="[1152,1282,1313,1338]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901DFFC3FB60FAD98C44AD00" box="[1152,1282,1313,1338]" italics="true" pageId="12" pageNumber="455">P. armatum</emphasis>
</taxonomicName>
:
<materialsCitation id="3B1D3C9A901DFFC3FAF2FAD88B86AD9D" collectingDate="1950-06-02" collectingDateMax="1989-11-16" collectingDateMin="1950-06-02" collectionCode="RV" country="Japan" location="Townsend Cromwell" municipality="Hawaii" pageId="12" pageNumber="455" specimenCode="LACM 44837-1, USNM 162717" specimenCount="2" stateProvince="Kochi" typeStatus="holotype">
<specimenCount id="9D73FD4E901DFFC3FAF2FAD88CDBAD00" box="[1298,1437,1312,1338]" count="2" pageId="12" pageNumber="455" type="generic">2 specimens</specimenCount>
<specimenCode id="DBD39EBC901DFFC3FF77FABC8809AD64" box="[151,335,1348,1374]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="12" pageNumber="455" type="Museum">LACM 44837-1</specimenCode>
, 213334 mm SL,
<collectionCode id="ED64AE02901DFFC3FDC7FABD8B0DAD65" box="[551,587,1349,1375]" country="Italy" name="Collection of Leptospira Strains" pageId="12" pageNumber="455">RV</collectionCode>
<location id="8EAA601C901DFFC3FDB2FABC8A79AD64" LSID="urn:lsid:plazi:treatment:03DC87D1901DFFC0FF77FC728D63AFAF:8EAA601C901DFFC3FDB2FABC8A79AD64" box="[594,831,1348,1374]" country="Japan" municipality="Hawaii" name="Townsend Cromwell" pageId="12" pageNumber="455" stateProvince="Kochi">Townsend Cromwell</location>
, st. 5264 (off
<collectingMunicipality id="6BAEACBD901DFFC3FC09FABC8D78AD64" box="[1001,1086,1348,1374]" pageId="12" pageNumber="455">Hawaii</collectingMunicipality>
),
<date id="FFCB1007901DFFC3FBB2FABC8C68AD64" box="[1106,1326,1348,1374]" pageId="12" pageNumber="455" value="1989-11-16">
<collectingDate id="EF8FE9EF901DFFC3FBB2FABC8C68AD64" box="[1106,1326,1348,1374]" pageId="12" pageNumber="455" value="1989-11-16">16 November 1989</collectingDate>
</date>
;
<typeStatus id="54CE8865901DFFC3FADAFABC8CDAAD64" box="[1338,1436,1348,1374]" pageId="12" pageNumber="455" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C754D44901DFFC3FF58FA9188D1ADB8" authority="USNM" authorityName="USNM" box="[184,407,1384,1410]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901DFFC3FF58FA91887AADB8" box="[184,316,1385,1410]" italics="true" pageId="12" pageNumber="455">P. armatum</emphasis>
<specimenCode id="DBD39EBC901DFFC3FEA6FA9088BEADB8" box="[326,504,1384,1410]" pageId="12" pageNumber="455">USNM 162717</specimenCode>
</taxonomicName>
, 310 mm SL, off
<location id="8EAA601C901DFFC3FD2BFA918A97ADB8" LSID="urn:lsid:plazi:treatment:03DC87D1901DFFC0FF77FC728D63AFAF:8EAA601C901DFFC3FD2BFA918A97ADB8" box="[715,977,1384,1411]" country="Japan" municipality="Hawaii" name="Kona Coast of Hawaii" pageId="12" pageNumber="455" stateProvince="Kochi">Kona Coast of Hawaii</location>
, about 1/
<locationDeviation id="9A782FE5901DFFC3FBDBFA908DC1ADB8" box="[1083,1159,1384,1410]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="12" pageNumber="455" unit="mi" value="2.0">2 mile</locationDeviation>
off shore from point of entry into sea of the 1950 lava flow,
<date id="FFCB1007901DFFC3FDD5FA748B86AD9D" box="[565,704,1420,1447]" pageId="12" pageNumber="455" value="1950-06-02">
<collectingDate id="EF8FE9EF901DFFC3FDD5FA748B86AD9D" box="[565,704,1420,1447]" pageId="12" pageNumber="455" value="1950-06-02">2 June 1950</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A901DFFC3FD2CFA748CC7ADF0" collectingDate="1968-11-19" collectionCode="RV" country="Japan" elevation="12" latitude="21.035" longLatPrecision="75" longitude="-156.797" pageId="12" pageNumber="455" specimenCode="USNM 227389" specimenCount="1" stateProvince="Kochi" typeStatus="holotype">
<specimenCount id="9D73FD4E901DFFC3FD2CFA748A0AAD9C" box="[716,844,1420,1446]" count="1" pageId="12" pageNumber="455" type="generic">1 specimen</specimenCount>
<specimenCode id="DBD39EBC901DFFC3FCB3FA748D44AD9C" box="[851,1026,1420,1446]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="455" type="Museum">USNM 227389</specimenCode>
, 77.5 mm SL,
<geoCoordinate id="EE415000901DFFC3FB48FA748C53AD9D" box="[1192,1301,1420,1447]" degrees="21.035" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="55" value="21.035">21.035°N</geoCoordinate>
<geoCoordinate id="EE415000901DFFC3FAFCFA748CDBAD9D" box="[1308,1437,1420,1447]" degrees="156.797" direction="west" orientation="longitude" pageId="12" pageNumber="455" precision="55" value="-156.797">156.797°W</geoCoordinate>
(off
<collectorName id="26805311901DFFC3FF29FA48886DADF0" box="[201,299,1456,1482]" pageId="12" pageNumber="455">Molokai</collectorName>
,
<collectorName id="26805311901DFFC3FED6FA4888CDADF0" box="[310,395,1456,1482]" pageId="12" pageNumber="455">Hawaii</collectorName>
),
<collectionCode id="ED64AE02901DFFC3FE7FFA498885ADF1" box="[415,451,1457,1483]" country="Italy" name="Collection of Leptospira Strains" pageId="12" pageNumber="455">RV</collectionCode>
<collectorName id="26805311901DFFC3FE2BFA488BFFADF0" box="[459,697,1456,1482]" pageId="12" pageNumber="455">Townsend Cromwell</collectorName>
, st. 4067, 41 ft shrimp trawl, 199233 m,
<date id="FFCB1007901DFFC3FB44FA488CC7ADF0" box="[1188,1409,1456,1482]" pageId="12" pageNumber="455" value="1968-11-19">
<collectingDate id="EF8FE9EF901DFFC3FB44FA488CC7ADF0" box="[1188,1409,1456,1482]" pageId="12" pageNumber="455" value="1968-11-19">19 November 1968</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A901DFFC3FA6EFA488BF5AE28" collectingDate="1968-03-27" collectionCode="RV" country="Japan" elevation="12" latitude="20.98" longLatPrecision="557" longitude="-156.758" pageId="12" pageNumber="455" specimenCode="USNM 227390" specimenCount="1" stateProvince="Kochi" typeStatus="holotype">
<specimenCount id="9D73FD4E901DFFC3FA6EFA488847ADD4" count="1" pageId="12" pageNumber="455" type="generic">1 specimen</specimenCount>
<specimenCode id="DBD39EBC901DFFC3FEE8FA2C88F0ADD4" box="[264,438,1492,1518]" pageId="12" pageNumber="455">USNM 227390</specimenCode>
, 134 mm SL,
<geoCoordinate id="EE415000901DFFC3FDB3FA2C8BF7ADD4" box="[595,689,1492,1518]" degrees="20.98" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="555" value="20.98">20.98°N</geoCoordinate>
<geoCoordinate id="EE415000901DFFC3FD58FA2C8A7EADD5" box="[696,824,1492,1519]" degrees="156.758" direction="west" orientation="longitude" pageId="12" pageNumber="455" precision="55" value="-156.758">156.758°W</geoCoordinate>
(off
<collectorName id="26805311901DFFC3FC8FFA2C8AEBADD4" box="[879,941,1492,1518]" pageId="12" pageNumber="455">Maui</collectorName>
,
<collectorName id="26805311901DFFC3FC58FA2C8D4BADD4" box="[952,1037,1492,1518]" pageId="12" pageNumber="455">Hawaii</collectorName>
),
<collectionCode id="ED64AE02901DFFC3FBC0FA2D8D02ADD5" box="[1056,1092,1493,1519]" country="Italy" name="Collection of Leptospira Strains" pageId="12" pageNumber="455">RV</collectionCode>
<collectorName id="26805311901DFFC3FBAAFA2C8C71ADD4" box="[1098,1335,1492,1518]" pageId="12" pageNumber="455">Townsend Cromwell</collectorName>
, st. 35-1, 41 ft shrimp trawl, 205209 m,
<date id="FFCB1007901DFFC3FDE2FA008BF5AE28" box="[514,691,1528,1554]" pageId="12" pageNumber="455" value="1968-03-27">
<collectingDate id="EF8FE9EF901DFFC3FDE2FA008BF5AE28" box="[514,691,1528,1554]" pageId="12" pageNumber="455" value="1968-03-27">27 March 1968</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A901DFFC3FD20FA008C39AE0C" collectingDate="1968-11-25" collectionCode="RV" country="Japan" elevation="12" latitude="21.0633" longLatPrecision="51" longitude="-156.518" pageId="12" pageNumber="455" specimenCode="USNM 227406" specimenCount="1" stateProvince="Kochi" typeStatus="holotype">
<specimenCount id="9D73FD4E901DFFC3FD20FA008A06AE28" box="[704,832,1528,1554]" count="1" pageId="12" pageNumber="455" type="generic">1 specimen</specimenCount>
<specimenCode id="DBD39EBC901DFFC3FCA8FA008ABEAE28" box="[840,1016,1528,1554]" pageId="12" pageNumber="455">USNM 227406</specimenCode>
, 243 mm SL,
<geoCoordinate id="EE415000901DFFC3FB7AFA008C52AE28" box="[1178,1300,1528,1554]" degrees="21.0633" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="5" value="21.0633">21.0633°N</geoCoordinate>
<geoCoordinate id="EE415000901DFFC3FAFCFA008CDBAE29" box="[1308,1437,1528,1555]" degrees="156.518" direction="west" orientation="longitude" pageId="12" pageNumber="455" precision="55" value="-156.518">156.518°W</geoCoordinate>
(off
<collectorName id="26805311901DFFC3FF2BF9E4884FAE0C" box="[203,265,1564,1590]" pageId="12" pageNumber="455">Maui</collectorName>
,
<collectorName id="26805311901DFFC3FEF7F9E4882AAE0C" box="[279,364,1564,1590]" pageId="12" pageNumber="455">Hawaii</collectorName>
),
<collectionCode id="ED64AE02901DFFC3FE62F9E588E0AE0D" box="[386,422,1565,1591]" country="Italy" name="Collection of Leptospira Strains" pageId="12" pageNumber="455">RV</collectionCode>
<collectorName id="26805311901DFFC3FE50F9E48BD9AE0C" box="[432,671,1564,1590]" pageId="12" pageNumber="455">Townsend Cromwell</collectorName>
, st. 4092, 41 ft shrimp trawl, 274318 m,
<date id="FFCB1007901DFFC3FB7EF9E48C39AE0C" box="[1182,1407,1564,1591]" pageId="12" pageNumber="455" value="1968-11-25">
<collectingDate id="EF8FE9EF901DFFC3FB7EF9E48C39AE0C" box="[1182,1407,1564,1591]" pageId="12" pageNumber="455" value="1968-11-25">25 November 1968</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A901DFFC3FA6EF9E48BAAAE44" collectingDate="1967-11-09" collectionCode="RV" country="Japan" elevation="21" latitude="20.6617" longLatPrecision="51" longitude="-156.684" pageId="12" pageNumber="455" specimenCode="USNM 227408" specimenCount="1" stateProvince="Kochi" typeStatus="holotype">
<pagination id="B2BCC833901DFFC3FA6EF9E48BAAAE44" pageId="12" pageNumber="455">
<specimenCount id="9D73FD4E901DFFC3FA6EF9E48847AE60" count="1" pageId="12" pageNumber="455" type="generic">1 specimen</specimenCount>
<specimenCode id="DBD39EBC901DFFC3FEE7F9B888F2AE60" box="[263,436,1600,1626]" pageId="12" pageNumber="455">USNM 227408</specimenCode>
, 191 mm SL,
<geoCoordinate id="EE415000901DFFC3FDACF9B88B80AE60" box="[588,710,1600,1626]" degrees="20.6617" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="5" value="20.6617">20.6617°N</geoCoordinate>
<geoCoordinate id="EE415000901DFFC3FD2CF9B88A0AAE61" box="[716,844,1600,1627]" degrees="156.684" direction="west" orientation="longitude" pageId="12" pageNumber="455" precision="55" value="-156.684">156.684°W</geoCoordinate>
(off
<collectorName id="26805311901DFFC3FC61F9B88A82AE60" box="[897,964,1600,1626]" pageId="12" pageNumber="455">Lanai</collectorName>
,
<collectorName id="26805311901DFFC3FC2EF9B88D65AE60" box="[974,1059,1600,1626]" pageId="12" pageNumber="455">Hawaii</collectorName>
),
<collectionCode id="ED64AE02901DFFC3FBD5F9B98D1FAE61" box="[1077,1113,1601,1627]" country="Italy" name="Collection of Leptospira Strains" pageId="12" pageNumber="455">RV</collectionCode>
<collectorName id="26805311901DFFC3FBBEF9B88C0FAE60" box="[1118,1353,1600,1626]" pageId="12" pageNumber="455">Townsend Cromwell</collectorName>
, st. 33 38, 70 ft shrimp trawl, 289296 m,
<date id="FFCB1007901DFFC3FDC0F99C8BAAAE44" box="[544,748,1636,1662]" pageId="12" pageNumber="455" value="1967-11-09">
<collectingDate id="EF8FE9EF901DFFC3FDC0F99C8BAAAE44" box="[544,748,1636,1662]" pageId="12" pageNumber="455" value="1967-11-09">9 November 1967</collectingDate>
</date>
</pagination>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A901DFFC3FD17F99C8C4AAE98" collectingDate="1972-09-24" collectionCode="RV" collectorName="Hawaii" country="Japan" elevation="247" latitude="21.53" location="Townsend Cromwell" longLatPrecision="758" longitude="-158.38" pageId="12" pageNumber="455" specimenCode="USNM 227412, 225" specimenCount="2" stateProvince="Kochi" typeStatus="holotype">
<specimenCount id="9D73FD4E901DFFC3FD17F99C8AC6AE44" box="[759,896,1636,1662]" count="2" pageId="12" pageNumber="455" type="generic">2 specimens</specimenCount>
<specimenCode id="DBD39EBC901DFFC3FC66F99C8D73AE44" box="[902,1077,1636,1662]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="455" type="Museum">USNM 227412</specimenCode>
,
<specimenCode id="DBD39EBC901DFFC3FBDFF99C8D2FAE45" box="[1087,1129,1636,1663]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="455" type="Museum">225</specimenCode>
and 265+ mm SL,
<geoCoordinate id="EE415000901DFFC3FADEF99C8CDAAE45" box="[1342,1436,1636,1663]" degrees="21.53" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="555" value="21.53">21.53°N</geoCoordinate>
<geoCoordinate id="EE415000901DFFC3FF77F970884FAE99" box="[151,265,1672,1699]" degrees="158.38" direction="west" orientation="longitude" pageId="12" pageNumber="455" precision="555" value="-158.38">158.38°W</geoCoordinate>
(off
<collectorName id="26805311901DFFC3FEA6F97088DDAE98" box="[326,411,1672,1698]" pageId="12" pageNumber="455">Hawaii</collectorName>
),
<collectionCode id="ED64AE02901DFFC3FE51F9718893AE99" box="[433,469,1673,1699]" country="Italy" name="Collection of Leptospira Strains" pageId="12" pageNumber="455">RV</collectionCode>
<location id="8EAA601C901DFFC3FE3FF9708B88AE98" LSID="urn:lsid:plazi:treatment:03DC87D1901DFFC0FF77FC728D63AFAF:8EAA601C901DFFC3FE3FF9708B88AE98" box="[479,718,1672,1698]" country="Japan" latitude="21.53" longLatPrecision="758" longitude="-158.38" name="Townsend Cromwell" pageId="12" pageNumber="455" stateProvince="Kochi">Townsend Cromwell</location>
, cruise 54, st. 8, 229247 m,
<date id="FFCB1007901DFFC3FBCAF9708C4AAE98" box="[1066,1292,1672,1698]" pageId="12" pageNumber="455" value="1972-09-24">
<collectingDate id="EF8FE9EF901DFFC3FBCAF9708C4AAE98" box="[1066,1292,1672,1698]" pageId="12" pageNumber="455" value="1972-09-24">24 September 1972</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A901DFFC3FAFBF9708B24AED0" collectingDate="1968-11-17" collectionCode="USNM, RV" collectorName="Maui &amp; Hawaii" country="Japan" elevation="218" latitude="21.0108" location="Townsend Cromwell" longLatPrecision="51" longitude="-156.759" pageId="12" pageNumber="455" specimenCount="1" stateProvince="Kochi" typeStatus="holotype">
<specimenCount id="9D73FD4E901DFFC3FAFBF9708CDAAE98" box="[1307,1436,1672,1698]" count="1" pageId="12" pageNumber="455" type="generic">1 specimen</specimenCount>
<collectionCode id="ED64AE02901DFFC3FF77F95489AEAEFC" box="[151,232,1708,1734]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="455" type="Museum">USNM</collectionCode>
455317, 165 mm SL,
<geoCoordinate id="EE415000901DFFC3FE04F9548B19AEFC" box="[484,607,1708,1734]" degrees="21.0108" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="5" value="21.0108">21.0108°N</geoCoordinate>
<geoCoordinate id="EE415000901DFFC3FD86F9548BA1AEFD" box="[614,743,1708,1735]" degrees="156.759" direction="west" orientation="longitude" pageId="12" pageNumber="455" precision="55" value="-156.759">156.759°W</geoCoordinate>
(off
<collectorName id="26805311901DFFC3FCFEF9548A1AAEFC" box="[798,860,1708,1734]" pageId="12" pageNumber="455">Maui</collectorName>
,
<collectorName id="26805311901DFFC3FC87F9548AFAAEFC" box="[871,956,1708,1734]" pageId="12" pageNumber="455">Hawaii</collectorName>
),
<collectionCode id="ED64AE02901DFFC3FC30F9558AB2AEFD" box="[976,1012,1709,1735]" country="Italy" name="Collection of Leptospira Strains" pageId="12" pageNumber="455">RV</collectionCode>
<location id="8EAA601C901DFFC3FC1BF9548DAEAEFC" LSID="urn:lsid:plazi:treatment:03DC87D1901DFFC0FF77FC728D63AFAF:8EAA601C901DFFC3FC1BF9548DAEAEFC" box="[1019,1256,1708,1734]" country="Japan" latitude="21.0108" longLatPrecision="51" longitude="-156.759" name="Townsend Cromwell" pageId="12" pageNumber="455" stateProvince="Kochi">Townsend Cromwell</location>
, st. 4049, 41 ft shrimp trawl, 218 m,
<date id="FFCB1007901DFFC3FE67F9288B24AED0" box="[391,610,1744,1770]" pageId="12" pageNumber="455" value="1968-11-17">
<collectingDate id="EF8FE9EF901DFFC3FE67F9288B24AED0" box="[391,610,1744,1770]" pageId="12" pageNumber="455" value="1968-11-17">17 November 1968</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1D3C9A901DFFC3FD8EF9288DCBAF34" collectingDate="1968-03-28" collectionCode="RV" country="Japan" elevation="12" latitude="20.99" longLatPrecision="557" longitude="-156.751" pageId="12" pageNumber="455" specimenCode="USNM 455319" specimenCount="1" stateProvince="Kochi" typeStatus="holotype">
<specimenCount id="9D73FD4E901DFFC3FD8EF9288BAAAED0" box="[622,748,1744,1770]" count="1" pageId="12" pageNumber="455" type="generic">1 specimen</specimenCount>
<specimenCode id="DBD39EBC901DFFC3FD13F9288AE4AED1" box="[755,930,1744,1771]" pageId="12" pageNumber="455">USNM 455319</specimenCode>
, 276 mm SL,
<geoCoordinate id="EE415000901DFFC3FBDEF9288DDAAED0" box="[1086,1180,1744,1770]" degrees="20.99" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="555" value="20.99">20.99°N</geoCoordinate>
<geoCoordinate id="EE415000901DFFC3FB43F9288C65AED1" box="[1187,1315,1744,1771]" degrees="156.751" direction="west" orientation="longitude" pageId="12" pageNumber="455" precision="55" value="-156.751">156.751°W</geoCoordinate>
(off
<collectorName id="26805311901DFFC3FABAF9288CDEAED0" box="[1370,1432,1744,1770]" pageId="12" pageNumber="455">Maui</collectorName>
,
<collectorName id="26805311901DFFC3FF77F90C89AAAF34" box="[151,236,1780,1806]" pageId="12" pageNumber="455">Hawaii</collectorName>
),
<collectionCode id="ED64AE02901DFFC3FF1FF90D8865AF35" box="[255,291,1781,1807]" country="Italy" name="Collection of Leptospira Strains" pageId="12" pageNumber="455">RV</collectionCode>
<collectorName id="26805311901DFFC3FEC9F90C8B53AF34" box="[297,533,1780,1806]" pageId="12" pageNumber="455">Townsend Cromwell</collectorName>
, st. 35-2, 41 ft shrimp trawl, 201216 m,
<date id="FFCB1007901DFFC3FC00F90C8DCBAF34" box="[992,1165,1780,1806]" pageId="12" pageNumber="455" value="1968-03-28">
<collectingDate id="EF8FE9EF901DFFC3FC00F90C8DCBAF34" box="[992,1165,1780,1806]" pageId="12" pageNumber="455" value="1968-03-28">28 March 1968</collectingDate>
</date>
</materialsCitation>
; specimens identified as
<taxonomicName id="4C754D44901DFFC3FF77F8E18807AF08" authorityName="Alcock" authorityYear="1889" box="[151,321,1817,1842]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="12" pageNumber="455" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901DFFC3FF77F8E18807AF08" box="[151,321,1817,1842]" italics="true" pageId="12" pageNumber="455">P. squamipinne</emphasis>
</taxonomicName>
:
<materialsCitation id="3B1D3C9A901DFFC3FEAFF8E08B20AF6D" collectingDate="1985-09-09" collectionCode="MNHN" country="New Caledonia" elevation="515" latitude="-22.182999" location="Biocal" longLatPrecision="14" longitude="167.25" pageId="12" pageNumber="455" specimenCode="MNHN 1994-0762" specimenCount="1" typeStatus="holotype">
<specimenCount id="9D73FD4E901DFFC3FEAFF8E08888AF08" box="[335,462,1816,1842]" count="1" pageId="12" pageNumber="455" type="generic">1 specimen</specimenCount>
<specimenCode id="DBD39EBC901DFFC3FE34F8E18BEBAF08" box="[468,685,1816,1843]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="455">MNHN 1994-0762</specimenCode>
, 99 mm SL,
<geoCoordinate id="EE415000901DFFC3FCD9F8E08A8AAF09" box="[825,972,1816,1843]" degrees="22" direction="south" minutes="10" orientation="latitude" pageId="12" pageNumber="455" precision="1" seconds="58.8" value="-22.182999">22°10'58.8''S</geoCoordinate>
<geoCoordinate id="EE415000901DFFC3FC33F8E08D14AF09" box="[979,1106,1816,1843]" degrees="167" direction="east" minutes="15" orientation="longitude" pageId="12" pageNumber="455" precision="15" seconds="0" value="167.25">167°15'0''E</geoCoordinate>
(off
<collectingCountry id="F3627657901DFFC3FB69F8E18C7CAF08" box="[1161,1338,1816,1843]" name="New Caledonia" pageId="12" pageNumber="455">New Caledonia</collectingCountry>
),
<location id="8EAA601C901DFFC3FAACF8E08CDEAF08" LSID="urn:lsid:plazi:treatment:03DC87D1901DFFC0FF77FC728D63AFAF:8EAA601C901DFFC3FAACF8E08CDEAF08" box="[1356,1432,1816,1842]" country="New Caledonia" latitude="-22.182999" longLatPrecision="14" longitude="167.25" name="Biocal" pageId="12" pageNumber="455">Biocal</location>
, st. cp109, 495
<quantity id="4C8D9B22901DFFC3FEA3F8C488CDAF6D" box="[323,395,1852,1879]" metricMagnitude="2" metricUnit="m" metricValue="5.15" pageId="12" pageNumber="455" unit="m" value="515.0">
<elevation id="0058D1F4901DFFC3FEA3F8C488CDAF6D" box="[323,395,1852,1879]" metricMagnitude="2" metricUnit="m" metricValue="5.15" pageId="12" pageNumber="455" unit="m" value="515.0">515 m</elevation>
</quantity>
,
<date id="FFCB1007901DFFC3FE77F8C48B20AF6D" box="[407,614,1852,1879]" pageId="12" pageNumber="455" value="1985-09-09">
<collectingDate id="EF8FE9EF901DFFC3FE77F8C48B20AF6D" box="[407,614,1852,1879]" pageId="12" pageNumber="455" value="1985-09-09">9 September 1985</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901DFFC2FF27F8988873A8EF" lastPageId="13" lastPageNumber="456" pageId="12" pageNumber="455" type="diagnosis">
<paragraph id="8BCA36C7901DFFC2FF27F8988873A8EF" blockId="12.[151,1437,1204,2023]" lastBlockId="13.[151,1437,150,321]" lastPageId="13" lastPageNumber="456" pageId="12" pageNumber="455">
<emphasis id="B901EAD5901DFFC3FF27F8988804AF40" bold="true" box="[199,322,1888,1914]" pageId="12" pageNumber="455">Diagnosis.</emphasis>
Precaudal vertebrae 1114, total vertebrae 5259; dorsal-fin rays 95103; anal-fin rays 7179; pectoral-fin rays 2526; long gill rakers on first gill arch 45; pseudobranchials 47; 25 dorsal-fin rays in front of first vertebra; HL 23.826.9% of SL; body depth at vent 19.423.3% of SL; predorsal length 1722.4% of SL; rear margin of preopercle with 4 spines and fleshy flap between spines 2 and 3; transversal scale rows above origin of anal fin 6068, and above lateral line 1520; otoliths&gt;
<quantity id="4C8D9B22901CFFC2FCE2FF6E8A16A88B" box="[770,848,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="456" unit="mm" value="15.0">15 mm</quantity>
in length: OL:OH = 1.952.2, dorsal rim crenulated or undulating.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901CFFC0FF27FF268867ABBB" lastPageId="15" lastPageNumber="458" pageId="13" pageNumber="456" type="description">
<paragraph id="8BCA36C7901CFFC2FF27FF26880AA97B" blockId="13.[151,1437,150,321]" pageId="13" pageNumber="456">
<emphasis id="B901EAD5901CFFC2FF27FF26881FA8C2" bold="true" box="[199,345,222,248]" pageId="13" pageNumber="456">Description.</emphasis>
Relatively robust fish with stout tapering tail and rounded snout. Maximal size of fishes investigated
<quantity id="4C8D9B22901CFFC2FEC6FEFB88C5A927" box="[294,387,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="3.34" pageId="13" pageNumber="456" unit="mm" value="334.0">334 mm</quantity>
SL. Preopercular rim with 4 spines, 2 spines positioned at bend and 2 further up, with fleshy flap in between.
</paragraph>
<caption id="DF0A664F901CFFC2FF77F8EB8A3CAFAD" ID-DOI="http://doi.org/10.5281/zenodo.15561741" ID-Zenodo-Dep="15561741" httpUri="https://zenodo.org/record/15561741/files/figure.png" pageId="13" pageNumber="456" startId="13.[151,250,1811,1835]" targetBox="[151,1436,352,1787]" targetPageId="13" targetType="figure">
<paragraph id="8BCA36C7901CFFC2FF77F8EB8A3CAFAD" blockId="13.[151,1436,1811,1943]" pageId="13" pageNumber="456">
<emphasis id="B901EAD5901CFFC2FF77F8EB8854AF11" bold="true" box="[151,274,1811,1835]" pageId="13" pageNumber="456">FIGURE 4.</emphasis>
Head and body squamation details in
<taxonomicName id="4C754D44901CFFC2FD70F8EB8D4BAF11" authority=", USNM" authorityName="USNM" box="[656,1037,1811,1835]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="13" pageNumber="456" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD5901CFFC2FD70F8EB8AFFAF11" box="[656,953,1811,1835]" italics="true" pageId="13" pageNumber="456">Pycnocraspedum phyllosoma</emphasis>
, USNM
</taxonomicName>
421587; abbreviations used: aip-1aip- 3 = anterior infraorbital pores 13, asp = anterior supraorbital pore, pip-1 = posterior infraorbital pore 1; preopercular spines numbered.
<emphasis id="B901EAD5901CFFC2FEE9F8A3886EAF49" bold="true" box="[265,296,1883,1907]" pageId="13" pageNumber="456">4A</emphasis>
) head squamation in lateral view;
<emphasis id="B901EAD5901CFFC2FD66F8A38BE2AF49" bold="true" box="[646,676,1883,1907]" pageId="13" pageNumber="456">4B</emphasis>
) trunk squamation and every 10 scale row count marked by white dot;
<emphasis id="B901EAD5901CFFC2FA95F8A38CD2AF49" bold="true" box="[1397,1428,1883,1907]" pageId="13" pageNumber="456">4C</emphasis>
) snout squamation and pores detail;
<emphasis id="B901EAD5901CFFC2FE1DF8878B5AAFAD" bold="true" box="[509,540,1919,1943]" pageId="13" pageNumber="456">4D</emphasis>
) head squamation in ventral view.
</paragraph>
</caption>
<caption id="DF0A664F901FFFC1FF77F92E8D08AFD0" ID-DOI="http://doi.org/10.5281/zenodo.15561743" ID-Zenodo-Dep="15561743" httpUri="https://zenodo.org/record/15561743/files/figure.png" pageId="14" pageNumber="457" startId="14.[151,250,1750,1774]" targetBox="[245,1358,193,1715]" targetPageId="14" targetType="figure">
<paragraph id="8BCA36C7901FFFC1FF77F92E8D08AFD0" blockId="14.[151,1437,1750,2026]" pageId="14" pageNumber="457">
<emphasis id="B901EAD5901FFFC1FF77F92E8855AED4" bold="true" box="[151,275,1750,1774]" pageId="14" pageNumber="457">FIGURE 5.</emphasis>
Otoliths, inner faces (if not noted otherwise) shown from right side.
<emphasis id="B901EAD5901FFFC1FC29F92E8D52AED4" bold="true" box="[969,1044,1750,1774]" pageId="14" pageNumber="457">5A5C</emphasis>
)
<taxonomicName id="4C754D44901FFFC1FBC2F92E8C79AED4" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[1058,1343,1750,1774]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="14" pageNumber="457" phylum="Chordata" rank="species" species="africanum" status="sp. nov.">
<emphasis id="B901EAD5901FFFC1FBC2F92E8C79AED4" box="[1058,1343,1750,1774]" italics="true" pageId="14" pageNumber="457">Pycnocraspedum africanum</emphasis>
</taxonomicName>
<emphasis id="B901EAD5901FFFC1FAA5F92E8CD3AED4" bold="true" box="[1349,1429,1750,1774]" pageId="14" pageNumber="457">
<taxonomicNameLabel id="A23257AE901FFFC1FAA5F92E8CD3AED4" box="[1349,1429,1750,1774]" pageId="14" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 5A holotype, MNHN 2000-5691 (5A2 dorsal view), 5B paratype MNHN 2000-5692, 5C tentatively assigned specimen, IRSNB (5C2 dorsal view, 5C3 anterior view);
<emphasis id="B901EAD5901FFFC1FDC1F8E68B2CAF0C" bold="true" box="[545,618,1822,1846]" pageId="14" pageNumber="457">5D5E</emphasis>
)
<taxonomicName id="4C754D44901FFFC1FD9AF8E68ACCAF0C" box="[634,906,1822,1846]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="14" pageNumber="457" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901FFFC1FD9AF8E68ACCAF0C" box="[634,906,1822,1846]" italics="true" pageId="14" pageNumber="457">Pycnocraspedum armatum</emphasis>
</taxonomicName>
, 5D USNM 227412 (5D2 ventral view), 5E USNM 227390 (5E2 ventral view);
<emphasis id="B901EAD5901FFFC1FE56F8BA8894AF60" bold="true" box="[438,466,1858,1882]" pageId="14" pageNumber="457">5F</emphasis>
)
<taxonomicName id="4C754D44901FFFC1FE01F8BA8B9DAF60" authorityName="Machida" authorityYear="1984" box="[481,731,1858,1882]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="14" pageNumber="457" phylum="Chordata" rank="species" species="fulvum">
<emphasis id="B901EAD5901FFFC1FE01F8BA8B9DAF60" box="[481,731,1858,1882]" italics="true" pageId="14" pageNumber="457">Pycnocraspedum fulvum</emphasis>
</taxonomicName>
(holotype), BSKU 27497 (5F2 ventral view, 5F3 anterior view);
<emphasis id="B901EAD5901FFFC1FA93F8BA8CD2AF60" bold="true" box="[1395,1428,1858,1882]" pageId="14" pageNumber="457">5G</emphasis>
)
<taxonomicName id="4C754D44901FFFC1FF77F89E8880AF44" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[151,454,1894,1918]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="14" pageNumber="457" phylum="Chordata" rank="species" species="rowleyensis" status="sp. nov.">
<emphasis id="B901EAD5901FFFC1FF77F89E8880AF44" box="[151,454,1894,1918]" italics="true" pageId="14" pageNumber="457">Pycnocraspedum rowleyensis</emphasis>
</taxonomicName>
<emphasis id="B901EAD5901FFFC1FE2EF89E8B67AF44" bold="true" box="[462,545,1894,1918]" pageId="14" pageNumber="457">
<taxonomicNameLabel id="A23257AE901FFFC1FE2EF89E8B67AF44" box="[462,545,1894,1918]" pageId="14" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, WAM I.28059.021 (5G2 ventral view);
<emphasis id="B901EAD5901FFFC1FBD8F89E8D1EAF44" bold="true" box="[1080,1112,1894,1918]" pageId="14" pageNumber="457">5H</emphasis>
)
<taxonomicName id="4C754D44901FFFC1FB8AF89E89A6AF98" authority=", USNM" authorityName="USNM" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="14" pageNumber="457" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD5901FFFC1FB8AF89E8CD0AF44" box="[1130,1430,1894,1918]" italics="true" pageId="14" pageNumber="457">Pycnocraspedum phyllosoma</emphasis>
, USNM
</taxonomicName>
421587 (5H2 ventral view, 5H3 anterior view);
<emphasis id="B901EAD5901FFFC1FD20F8728A45AF98" bold="true" box="[704,771,1930,1954]" pageId="14" pageNumber="457">5I5K</emphasis>
)
<taxonomicName id="4C754D44901FFFC1FCF1F8728D03AF98" authorityName="Alcock" authorityYear="1889" box="[785,1093,1930,1954]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="14" pageNumber="457" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901FFFC1FCF1F8728D03AF98" box="[785,1093,1930,1954]" italics="true" pageId="14" pageNumber="457">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
, 5I lectotype, BMNH 1890.7.31.1 (5I2 ventral view), 5J ZMUC P771871, 5L ZMUC P771739 (5L2 ventral view), 5K ZMUC P771738 (5K2 ventral view);
<emphasis id="B901EAD5901FFFC1FA8FF8568CD5AFFC" bold="true" box="[1391,1427,1966,1990]" pageId="14" pageNumber="457">5M</emphasis>
)
<emphasis id="B901EAD5901FFFC1FF77F82A888CAFD0" box="[151,458,2002,2026]" italics="true" pageId="14" pageNumber="457">
<taxonomicName id="4C754D44901FFFC1FF42F82A888CAFD0" baseAuthorityName="Schwarzhans" baseAuthorityYear="1978" box="[162,458,2002,2026]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="14" pageNumber="457" phylum="Chordata" rank="species" species="cetonaense">Pycnocraspedum cetonaense</taxonomicName>
</emphasis>
(holotype) SMF P.5694 (5M2 dorsal view, 5M3 anterior view).
</paragraph>
</caption>
<paragraph id="8BCA36C7901EFFC0FF27FF6F8827A97B" blockId="15.[151,1437,151,1941]" pageId="15" pageNumber="458">
Meristics (
<specimenCount id="9D73FD4E901EFFC0FEA1FF6F8899A88B" box="[321,479,151,177]" count="14" pageId="15" pageNumber="458" type="generic">14 specimens</specimenCount>
, including
<typeStatus id="54CE8865901EFFC0FDBEFF6F8B86A88B" box="[606,704,151,177]" pageId="15" pageNumber="458" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C754D44901EFFC0FD09FF6F8A2DA88A" box="[745,875,151,176]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FD09FF6F8A2DA88A" box="[745,875,151,176]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
): precaudal vertebrae 13 (1114); total vertebrae 56 (5259); dorsal-fin rays 98 (95103); anal-fin rays 71 (7179); pectoral-fin rays 26 (2526); caudal-fin rays 10; D/V = -3 (-2 -5); D/A = 29 (2331); V/A = 17 (1517); long gill rakers on first gill arch 5 (45); pseudobranchial filaments 5 (47); transversal scale rows above origin of anal fin 6068, and above lateral line 20 (1520); pyloric coeca 12 (1216).
</paragraph>
<paragraph id="8BCA36C7901EFFC0FF27FEB38ADEA9EB" blockId="15.[151,1437,151,1941]" pageId="15" pageNumber="458">
Morphometrics (4
<specimenCount id="9D73FD4E901EFFC0FE45FEB38B07A95F" box="[421,577,331,357]" count="14" pageId="15" pageNumber="458" type="generic">14 specimens</specimenCount>
) in % of SL: HL 24.6 (23.826.9); body depth at vent 22.7 (19.423.3); diameter of eye 4.4 (3.94.8); upper jaw length 13.4 (11.815); interorbital width 7.7 (6.17.7); pelvic-fin base to anal-fin base 21.2 (21.128); predorsal length 18.4 (1722.4); preanal length 46.3 (4048); prepelvic length 21.2 (19.323.3); pectoral-fin length 16 (1618.1); pelvic-fin length 17.5.
</paragraph>
<paragraph id="8BCA36C7901EFFC0FF27FE238D4EAB03" blockId="15.[151,1437,151,1941]" pageId="15" pageNumber="458">
Otolith morphology (
<specimenCount id="9D73FD4E901EFFC0FE55FE238B02A9CF" box="[437,580,475,501]" count="4" pageId="15" pageNumber="458" type="generic">4 specimens</specimenCount>
). Size up to
<quantity id="4C8D9B22901EFFC0FD2BFE238A5CA9CF" box="[715,794,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="458" unit="mm" value="18.0">18 mm</quantity>
in length; OL:OH =
<quantity id="4C8D9B22901EFFC0FBE2FE228DC7A9CF" box="[1026,1153,474,501]" metricMagnitude="-2" metricUnit="m" metricValue="5.2705" metricValueMax="5.588" metricValueMin="4.953" pageId="15" pageNumber="458" unit="in" value="2.075" valueMax="2.2" valueMin="1.95">1.952.2 in</quantity>
specimens larger than
<quantity id="4C8D9B22901EFFC0FA60FE228985AA23" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="458" unit="mm" value="15.0">15 mm</quantity>
in length,
<quantity id="4C8D9B22901EFFC0FEDDFE07883BAA23" box="[317,381,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="4.064" pageId="15" pageNumber="458" unit="in" value="1.6">1.6 in</quantity>
specimens smaller than
<quantity id="4C8D9B22901EFFC0FD73FE078B93AA23" box="[659,725,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="458" unit="mm" value="7.0">7 mm</quantity>
in length; OH:OT = 2.32.9. Moderately thin, oval, elongated, anteriorly moderately projecting, posteriorly distinctly pointed, anterior and posterior tips inferior. Dorsal rim regularly curved, more or less strongly crenulated, undulating in largest specimen; ventral rim shallower and smoother than dorsal rim. Ornamentation of rims decreasing with size. Inner face moderately bent in horizontal direction, and slightly bent in vertical direction, with long, undivided sulcus reaching moderately close to anterior and posterior rims of otolith. Sulcus with shallow, undivided, uniform colliculum with flat to slightly concave dorsal rim and slightly convex ventral rim and rounded anterior and posterior tips. OL:CL = 1.31.45. Dorsal field with narrow, indistinct depression; ventral field smooth. Outer face less convex than inner face, smooth in large specimens, with many radial furrows and central tubercles in small specimens.
</paragraph>
<paragraph id="8BCA36C7901EFFC0FF27FCBB8867ABBB" blockId="15.[151,1437,151,1941]" pageId="15" pageNumber="458">
Coloration. Live coloration dark brown, sometimes light, speckled (
<bibRefCitation id="EFE44B36901EFFC0FC53FCBB8D04AB67" author="HURL &amp; Hawaii Undersea Research Laboratory Archive" box="[947,1090,835,861]" pageId="15" pageNumber="458" refId="ref18574" refString="HURL (Hawaii Undersea Research Laboratory Archive) (2019) HURL Archive. Available from: https://www.soest.hawaii.edu/HURL/HURLarchive/index.php (accessed March 2019)" type="url" year="2019">HURL 2019</bibRefCitation>
). Color of preserved specimens light brown.
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901EFFC0FF27FC728CDFAF77" pageId="15" pageNumber="458" type="discussion">
<paragraph id="8BCA36C7901EFFC0FF27FC728C3EAC47" blockId="15.[151,1437,151,1941]" pageId="15" pageNumber="458">
<emphasis id="B901EAD5901EFFC0FF27FC72880AAB9E" bold="true" box="[199,332,906,932]" pageId="15" pageNumber="458">Discussion.</emphasis>
Specimens studied from BSKU (5055 and 38643) originally identified as
<taxonomicName id="4C754D44901EFFC0FB40FC738C74AB9F" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[1184,1330,907,933]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901EFFC0FB40FC738C74AB9F" box="[1184,1330,907,933]" italics="true" pageId="15" pageNumber="458">P. microlepis</emphasis>
</taxonomicName>
from the Mimase fish market in
<collectingCountry id="F3627657901EFFC0FE46FC5788ABABF3" box="[422,493,943,969]" name="India" pageId="15" pageNumber="458">Kochi</collectingCountry>
, were probably trawled in Tosa Bay, which is not far from the type locality of
<taxonomicName id="4C754D44901EFFC0FA68FC57884AABD7" baseAuthorityName="Matsubara" baseAuthorityYear="1943" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901EFFC0FA68FC57884AABD7" italics="true" pageId="15" pageNumber="458">P. microlepis</emphasis>
</taxonomicName>
given as east of Kii Peninsula. The
<specimenCount id="9D73FD4E901EFFC0FD7CFC2B8A04ABD7" box="[668,834,979,1005]" count="2" pageId="15" pageNumber="458" type="generic">two specimens</specimenCount>
from BSKU match Matsubaras description (1943) of the unique
<typeStatus id="54CE8865901EFFC0FEF6FC0F883EAC2B" box="[278,376,1015,1041]" pageId="15" pageNumber="458" type="holotype">holotype</typeStatus>
reasonably well in many aspects except for the number of pseudobranchial filaments (absent in the
<typeStatus id="54CE8865901EFFC0FF3DFBE38879AC0F" box="[221,319,1051,1077]" pageId="15" pageNumber="458" type="holotype">holotype</typeStatus>
according to Matsubara vs.
<quantity id="4C8D9B22901EFFC0FD9CFBE28B85AC0F" box="[636,707,1050,1077]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="15" pageNumber="458" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">56 in</quantity>
the BSKU specimens) and pyloric coeca (20 vs.
<quantity id="4C8D9B22901EFFC0FB09FBE38C64AC0F" box="[1257,1314,1051,1077]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="15" pageNumber="458" unit="in" value="13.0">13 in</quantity>
the BSKU 38643 specimen). Matsubaras description is very detailed, and therefore we consider the
<specimenCount id="9D73FD4E901EFFC0FB97FBC78C5AAC63" box="[1143,1308,1087,1113]" count="2" pageId="15" pageNumber="458" type="generic">two specimens</specimenCount>
in question from BSKU to represent
<taxonomicName id="4C754D44901EFFC0FE51FB9B8B77AC46" box="[433,561,1123,1148]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FE51FB9B8B77AC46" box="[433,561,1123,1148]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
rather than
<taxonomicName id="4C754D44901EFFC0FD58FB9B8A0EAC47" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[696,840,1123,1149]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901EFFC0FD58FB9B8A0EAC47" box="[696,840,1123,1149]" italics="true" pageId="15" pageNumber="458">P. microlepis</emphasis>
</taxonomicName>
. The nature of
<taxonomicName id="4C754D44901EFFC0FC14FB9B8DC2AC47" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[1012,1156,1123,1149]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901EFFC0FC14FB9B8DC2AC47" box="[1012,1156,1123,1149]" italics="true" pageId="15" pageNumber="458">P. microlepis</emphasis>
</taxonomicName>
thus remains elusive.
</paragraph>
<paragraph id="8BCA36C7901EFFC0FF27FB7F8A68AE17" blockId="15.[151,1437,151,1941]" pageId="15" pageNumber="458">
<bibRefCitation id="EFE44B36901EFFC0FF27FB7F8836AC9B" author="Gosline, W. A." box="[199,368,1158,1185]" pageId="15" pageNumber="458" pagination="68 - 83" refId="ref18451" refString="Gosline, W. A. (1954) Fishes killed by the 1950 eruption of Mauna Loa. Brotulidae. Pacific Science, 8, 68-83." type="journal article" year="1954">Gosline (1954)</bibRefCitation>
described
<taxonomicName id="4C754D44901EFFC0FE0BFB7F8B2AAC9A" box="[491,620,1159,1184]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FE0BFB7F8B2AAC9A" box="[491,620,1159,1184]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
from off Hawaii based on fishes killed by the eruption of Mauna Loa in 1950. He compared
<taxonomicName id="4C754D44901EFFC0FE9EFB538B46ACFE" box="[382,512,1195,1220]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FE9EFB538B46ACFE" box="[382,512,1195,1220]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
to the type species
<taxonomicName id="4C754D44901EFFC0FD03FB538AC9ACFE" authorityName="Alcock" authorityYear="1889" box="[739,911,1195,1220]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901EFFC0FD03FB538AC9ACFE" box="[739,911,1195,1220]" italics="true" pageId="15" pageNumber="458">P. squamipinne</emphasis>
</taxonomicName>
but not to
<taxonomicName id="4C754D44901EFFC0FBEDFB538DD9ACFF" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[1037,1183,1195,1221]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901EFFC0FBEDFB538DD9ACFF" box="[1037,1183,1195,1221]" italics="true" pageId="15" pageNumber="458">P. microlepis</emphasis>
</taxonomicName>
, which was originally described as
<taxonomicName id="4C754D44901EFFC0FECAFB3788A9ACD3" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[298,495,1231,1257]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901EFFC0FECAFB3788A9ACD3" box="[298,495,1231,1257]" italics="true" pageId="15" pageNumber="458">Itatius microlepis</emphasis>
</taxonomicName>
. The description of the unique
<typeStatus id="54CE8865901EFFC0FCB5FB378AF1ACD3" box="[853,951,1231,1257]" pageId="15" pageNumber="458" type="holotype">holotype</typeStatus>
did not contain meristics and it is difficult to assess morphometric data based on the drawing. Pseudobranchial filaments were not mentioned for
<taxonomicName id="4C754D44901EFFC0FAFBFB0B8CDAAD36" box="[1307,1436,1267,1292]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FAFBFB0B8CDAAD36" box="[1307,1436,1267,1292]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
but were mentioned in the description of the genus as rudimentary. A restudy of the
<typeStatus id="54CE8865901EFFC0FB80FAEF8D84AD0B" box="[1120,1218,1303,1329]" pageId="15" pageNumber="458" type="holotype">holotype</typeStatus>
as well as multiple subsequently collected specimens of
<taxonomicName id="4C754D44901EFFC0FDD9FAC38BBCAD6F" authorityName="Alcock" authorityYear="1889" box="[569,762,1339,1365]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="genus">
<emphasis id="B901EAD5901EFFC0FDD9FAC38BBCAD6F" box="[569,762,1339,1365]" italics="true" pageId="15" pageNumber="458">Pycnocraspedum</emphasis>
</taxonomicName>
from the Hawaiian Islands in the collections of LACM and USNM revealed a good match with the specimens from
<collectingCountry id="F3627657901EFFC0FCE0FAA78A06AD43" box="[768,832,1375,1401]" name="Japan" pageId="15" pageNumber="458">Japan</collectingCountry>
(BSKU).
<bibRefCitation id="EFE44B36901EFFC0FC4EFAA78D26AD43" author="Machida, Y." box="[942,1120,1375,1401]" pageId="15" pageNumber="458" pagination="371 - 375" refId="ref18614" refString="Machida, Y. (1984) Ophidiidae. In: Okamura, O. &amp; Kitajima, T. (Eds.), Fishes of the Okinawa Trough and the adjacent waters. Vol. I. The intensive research of unexploited fishery resources on continental slopes. Japan Fisheries Resource Conservation Association, Tokyo, pp. 244 - 261 + 371-375." type="book chapter" year="1984">Machida (1984)</bibRefCitation>
listed a difference of pyloric caeca of 12 for
<taxonomicName id="4C754D44901EFFC0FEA4FA7B8882ADA6" box="[324,452,1411,1436]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FEA4FA7B8882ADA6" box="[324,452,1411,1436]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
and 20 for
<taxonomicName id="4C754D44901EFFC0FDA3FA7B8B92ADA7" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[579,724,1411,1437]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901EFFC0FDA3FA7B8B92ADA7" box="[579,724,1411,1437]" italics="true" pageId="15" pageNumber="458">P. microlepis</emphasis>
</taxonomicName>
and 4 preopercular spines in
<taxonomicName id="4C754D44901EFFC0FBFBFA7B8DDDADA6" box="[1051,1179,1411,1436]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FBFBFA7B8DDDADA6" box="[1051,1179,1411,1436]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
(vs.
<quantity id="4C8D9B22901EFFC0FB32FA7B8DBAADA7" box="[1234,1276,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="458" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C754D44901EFFC0FAE3FA7B8CD5ADA7" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[1283,1427,1411,1437]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901EFFC0FAE3FA7B8CD5ADA7" box="[1283,1427,1411,1437]" italics="true" pageId="15" pageNumber="458">P. microlepis</emphasis>
</taxonomicName>
) based on the original descriptions. The specimens BSKU 5055 and 38643 from
<collectingCountry id="F3627657901EFFC0FBFAFA5F8D1CADFB" box="[1050,1114,1447,1473]" name="Japan" pageId="15" pageNumber="458">Japan</collectingCountry>
have 5 to 6 pseudobranchial filaments and the specimen BSKU 38643 has 13 pyloric coeca (information provided by N. Nakayama). Therefore, we consider
<taxonomicName id="4C754D44901EFFC0FEC6FA178BC9AE33" authority="(Matsubara, 1943)" baseAuthorityName="Matsubara" baseAuthorityYear="1943" box="[294,655,1519,1545]" family="Ophidiidae" genus="Itatius" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B901EAD5901EFFC0FEC6FA1788FEAE33" box="[294,440,1519,1545]" italics="true" pageId="15" pageNumber="458">P. microlepis</emphasis>
(
<bibRefCitation id="EFE44B36901EFFC0FE27FA178BC1AE33" author="Matsubara, K." box="[455,647,1519,1545]" pageId="15" pageNumber="458" pagination="37 - 81" refId="ref18682" refString="Matsubara, K. (1943) Ichthyological annotations from the depths of the sea Japan, I - VII. Journal of Sigenkagaku Kenkyusyo, 1, 37-81." type="journal article" year="1943">Matsubara, 1943</bibRefCitation>
)
</taxonomicName>
as a nominally valid species distinct from
<taxonomicName id="4C754D44901EFFC0FB95FA178CDAAE33" authority="Gosline, 1954" authorityName=", Gosline" authorityYear="1954" box="[1141,1436,1518,1545]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FB95FA178DB0AE32" box="[1141,1270,1519,1544]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
<bibRefCitation id="EFE44B36901EFFC0FB1EFA178CDAAE33" author="Gosline, W. A." box="[1278,1436,1518,1545]" pageId="15" pageNumber="458" pagination="68 - 83" refId="ref18451" refString="Gosline, W. A. (1954) Fishes killed by the 1950 eruption of Mauna Loa. Brotulidae. Pacific Science, 8, 68-83." type="journal article" year="1954">Gosline, 1954</bibRefCitation>
</taxonomicName>
and place the
<specimenCount id="9D73FD4E901EFFC0FED2F9EB889EAE17" box="[306,472,1555,1581]" count="2" pageId="15" pageNumber="458" type="generic">two specimens</specimenCount>
from BSKU with
<taxonomicName id="4C754D44901EFFC0FD46F9EB8A61AE16" authorityName=", Gosline" authorityYear="1954" box="[678,807,1555,1580]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FD46F9EB8A61AE16" box="[678,807,1555,1580]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCA36C7901EFFC0FF27F9CF8CDFAF77" blockId="15.[151,1437,151,1941]" pageId="15" pageNumber="458">
<taxonomicName id="4C754D44901EFFC0FF27F9CF88B5AE6A" authorityName=", Gosline" authorityYear="1954" box="[199,499,1591,1617]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FF27F9CF88B5AE6A" box="[199,499,1591,1617]" italics="true" pageId="15" pageNumber="458">Pycnocraspedum armatum</emphasis>
</taxonomicName>
is distinguished from its congeners by the higher number of dorsal-fin rays (95103 vs. &lt;93) except
<taxonomicName id="4C754D44901EFFC0FEB1F9A388B6AE4F" box="[337,496,1627,1653]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD5901EFFC0FEB1F9A388B6AE4F" box="[337,496,1627,1653]" italics="true" pageId="15" pageNumber="458">P. phyllosoma</emphasis>
</taxonomicName>
. Anal-fin rays also tend to be higher in number in
<taxonomicName id="4C754D44901EFFC0FBDFF9A38D84AE4E" authorityName=", Gosline" authorityYear="1954" box="[1087,1218,1627,1652]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FBDFF9A38D84AE4E" box="[1087,1218,1627,1652]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D44901EFFC0FB1DF9A38CDAAE4F" box="[1277,1436,1627,1653]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD5901EFFC0FB1DF9A38CDAAE4F" box="[1277,1436,1627,1653]" italics="true" pageId="15" pageNumber="458">P. phyllosoma</emphasis>
</taxonomicName>
(7079 vs. 5771). The dorsal-fin origin has a more anterior position in
<taxonomicName id="4C754D44901EFFC0FC25F9878D00AEA2" authorityName=", Gosline" authorityYear="1954" box="[965,1094,1663,1688]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FC25F9878D00AEA2" box="[965,1094,1663,1688]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
compared to the other species with 2 to 5 fin rays before of the first vertebra (vs.
<quantity id="4C8D9B22901EFFC0FD2AF95B8A63AE87" box="[714,805,1699,1725]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="7.62" metricValueMin="0.0" pageId="15" pageNumber="458" unit="in" value="1.5" valueMax="3.0" valueMin="0.0">0 to 3 in</quantity>
other species) and a predorsal length of 1722.4, mostly &lt;21. Otoliths&gt;
<quantity id="4C8D9B22901EFFC0FEA3F93E88D7AEDB" box="[323,401,1734,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="458" unit="mm" value="15.0">15 mm</quantity>
in length of
<taxonomicName id="4C754D44901EFFC0FDFFF93F8BE6AEDA" authorityName=", Gosline" authorityYear="1954" box="[543,672,1735,1760]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FDFFF93F8BE6AEDA" box="[543,672,1735,1760]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D44901EFFC0FD36F93F8A35AEDB" box="[726,883,1735,1761]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD5901EFFC0FD36F93F8A35AEDB" box="[726,883,1735,1761]" italics="true" pageId="15" pageNumber="458">P. phyllosoma</emphasis>
</taxonomicName>
are more compressed than in
<taxonomicName id="4C754D44901EFFC0FB23F93F8C17AEDB" authorityName="Schwarzhans &amp; Psomadakis &amp; Nielsen" authorityYear="2025" box="[1219,1361,1735,1761]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="africanum">
<emphasis id="B901EAD5901EFFC0FB23F93F8C17AEDB" box="[1219,1361,1735,1761]" italics="true" pageId="15" pageNumber="458">P. africanum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D44901EFFC0FA68F93F8860AF3E" authorityName="Alcock" authorityYear="1889" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="squamipinne">
<emphasis id="B901EAD5901EFFC0FA68F93F8860AF3E" italics="true" pageId="15" pageNumber="458">P. squamipinne</emphasis>
</taxonomicName>
(OL:OH = 1.952.2 vs. 2.42.7). The distinction of
<taxonomicName id="4C754D44901EFFC0FC8CF9138AADAF3E" authorityName=", Gosline" authorityYear="1954" box="[876,1003,1771,1796]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FC8CF9138AADAF3E" box="[876,1003,1771,1796]" italics="true" pageId="15" pageNumber="458">P. armatum</emphasis>
</taxonomicName>
from
<taxonomicName id="4C754D44901EFFC0FBCEF9138D8CAF3F" box="[1070,1226,1771,1797]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="phyllosoma">
<emphasis id="B901EAD5901EFFC0FBCEF9138D8CAF3F" box="[1070,1226,1771,1797]" italics="true" pageId="15" pageNumber="458">P. phyllosoma</emphasis>
</taxonomicName>
is relatively subtle despite their wide geographic separation (see below) and confined to differences in the number of transversal scale rows (6068 vs. 7275), scale rows above the lateral line (1520 vs. 2021) and spines on the preopercle (4 vs. 3).
</paragraph>
</subSubSection>
<subSubSection id="C36F654C901EFFC0FF27F8AE8D63AFAF" pageId="15" pageNumber="458" type="distribution">
<paragraph id="8BCA36C7901EFFC0FF27F8AE8D63AFAF" blockId="15.[151,1437,151,1941]" pageId="15" pageNumber="458">
<emphasis id="B901EAD5901EFFC0FF27F8AE8827AF4A" bold="true" box="[199,353,1878,1904]" pageId="15" pageNumber="458">Distribution.</emphasis>
<taxonomicName id="4C754D44901EFFC0FE87F8AF8BD5AF4A" authorityName=", Gosline" authorityYear="1954" box="[359,659,1879,1905]" family="Ophidiidae" genus="Pycnocraspedum" kingdom="Animalia" order="Ophidiiformes" pageId="15" pageNumber="458" phylum="Chordata" rank="species" species="armatum">
<emphasis id="B901EAD5901EFFC0FE87F8AF8BD5AF4A" box="[359,659,1879,1905]" italics="true" pageId="15" pageNumber="458">Pycnocraspedum armatum</emphasis>
</taxonomicName>
appears to be widely distributed from
<collectingCountry id="F3627657901EFFC0FBDFF8AF8D39AF4B" box="[1087,1151,1879,1905]" name="Japan" pageId="15" pageNumber="458">Japan</collectingCountry>
to Hawaii and southward to
<collectingCountry id="F3627657901EFFC0FF54F8838822AFAF" box="[180,356,1915,1941]" name="New Caledonia" pageId="15" pageNumber="458">New Caledonia</collectingCountry>
, and possibly Central West Pacific according to
<bibRefCitation id="EFE44B36901EFFC0FC61F8838D67AFAF" author="HURL &amp; Hawaii Undersea Research Laboratory Archive" box="[897,1057,1915,1941]" pageId="15" pageNumber="458" refId="ref18574" refString="HURL (Hawaii Undersea Research Laboratory Archive) (2019) HURL Archive. Available from: https://www.soest.hawaii.edu/HURL/HURLarchive/index.php (accessed March 2019)" type="url" year="2019">HURL (2019)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,56 +1,56 @@
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<mods:namePart id="76D3B31569E1156395744F9A57596336">Mirghaffari, Seyyed Ahmad</mods:namePart>
<mods:affiliation id="70E34503EB56F1DE60C6009D4FBCCE90">Department of Biology, Faculty of Science, Razi University, Baghabrisham 6714967346, Kermanshah, Iran.</mods:affiliation>
<mods:namePart id="F4C4E38723020C0CB13C0178B7214B75">Mirghaffari, Seyyed Ahmad</mods:namePart>
<mods:affiliation id="BC1B967230D0D8954642800089D11FAE">Department of Biology, Faculty of Science, Razi University, Baghabrisham 6714967346, Kermanshah, Iran.</mods:affiliation>
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<mods:namePart id="EE8E8A962A075194F2D02C4BCB67E959">Esmaeili-Rineh, Somayeh</mods:namePart>
<mods:affiliation id="E3F8A1E321E64295C840DEE457B03C69">Department of Biology, Faculty of Science, Razi University, Baghabrisham 6714967346, Kermanshah, Iran.</mods:affiliation>
<mods:namePart id="8BAFE952122333EFF16A4BBB3A297A3B">Esmaeili-Rineh, Somayeh</mods:namePart>
<mods:affiliation id="8EFA6B5D5DDB70B70D08F7F74C946F15">Department of Biology, Faculty of Science, Razi University, Baghabrisham 6714967346, Kermanshah, Iran.</mods:affiliation>
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<treatment id="FB2287EBBD17491048952E4CFC018981" ID-DOI="http://doi.org/10.5281/zenodo.15561671" ID-GBIF-Taxon="262796255" ID-Zenodo-Dep="15561671" LSID="urn:lsid:plazi:treatment:FB2287EBBD17491048952E4CFC018981" httpUri="http://treatment.plazi.org/id/FB2287EBBD17491048952E4CFC018981" lastPageId="7" lastPageNumber="480" pageId="4" pageNumber="477" scope_class="Malacostraca" scope_family="Niphargidae" scope_order="Amphipoda" scope_subOrder="Senticaudata">
<subSubSection id="3B916576BD17491348952E4CFD928B43" box="[151,527,1303,1330]" pageId="4" pageNumber="477" type="nomenclature">
<paragraph id="733436FDBD17491348952E4CFD928B43" blockId="4.[151,527,1303,1365]" box="[151,527,1303,1330]" pageId="4" pageNumber="477">
<heading id="287C8191BD17491348952E4CFD928B43" bold="true" box="[151,527,1303,1330]" fontSize="11" level="1" pageId="4" pageNumber="477" reason="1">
@ -71,12 +71,15 @@
<subSubSection id="3B916576BD17491348952ED8FB918B97" pageId="4" pageNumber="477" type="materials_examined">
<paragraph id="733436FDBD17491348952ED8FB918B97" blockId="4.[151,1437,1411,2013]" pageId="4" pageNumber="477">
<emphasis id="41FFEAEFBD17491348952ED8FE198BEF" bold="true" box="[151,388,1411,1437]" pageId="4" pageNumber="477">Material examined.</emphasis>
<materialsCitation id="C3E33CA0BD174913498C2ED8FF4E8B97" collectingDate="2022-06-22" collectionCode="ZCRU" collectorName="S. A. Mirghaffari" country="Iran" latitude="36.190556" location="Samaq Abad Spring" longLatPrecision="19" longitude="50.614998" municipality="Abyek City" pageId="4" pageNumber="477" specimenCount="1" stateProvince="Qazvin" typeStatus="holotype">
<materialsCitation id="C3E33CA0BD174913498C2ED8FF4E8B97" ID-GBIF-Occurrence="5169430302" collectingDate="2022-06-22" collectionCode="ZCRU" collectorName="S. A. Mirghaffari" country="Iran" latitude="36.190556" location="Samaq Abad Spring" longLatPrecision="19" longitude="50.614998" municipality="Abyek City" pageId="4" pageNumber="477" specimenCode="Amph.1504" specimenCount="1" stateProvince="Qazvin Province" typeStatus="holotype">
<typeStatus id="AC30885FBD174913498C2ED8FE648BEF" box="[398,505,1411,1437]" pageId="4" pageNumber="477" type="holotype">Holotype</typeStatus>
,
<collectionCode id="159AAE38BD1749134A052ED8FDCF8BEF" box="[519,594,1411,1437]" pageId="4" pageNumber="477">ZCRU</collectionCode>
Amph.1504, male specimen (8.0 mm) from
<location id="76546026BD1749134C5E2ED8FAD98BEF" LSID="urn:lsid:plazi:treatment:FB2287EBBD17491048952E4CFC018981:76546026BD1749134C5E2ED8FAD98BEF" box="[1116,1348,1411,1437]" country="Iran" latitude="36.190556" longLatPrecision="19" longitude="50.614998" municipality="Abyek City" name="Samaq Abad Spring" pageId="4" pageNumber="477" stateProvince="Qazvin">Samaq Abad Spring</location>
<specimenCode id="232D9E86BD1749134A592ED8FD7F8BEC" box="[603,738,1411,1438]" pageId="4" pageNumber="477">Amph.1504</specimenCode>
,
<specimenCount id="658DFD74BD1749134AF32ED8FC068BEF" box="[753,923,1411,1437]" pageId="4" pageNumber="477" type="male">male specimen</specimenCount>
(8.0 mm) from
<location id="76546026BD1749134C5E2ED8FAD98BEF" LSID="urn:lsid:plazi:treatment:FB2287EBBD17491048952E4CFC018981:76546026BD1749134C5E2ED8FAD98BEF" box="[1116,1348,1411,1437]" country="Iran" latitude="36.190556" longLatPrecision="19" longitude="50.614998" municipality="Abyek City" name="Samaq Abad Spring" pageId="4" pageNumber="477" stateProvince="Qazvin Province">Samaq Abad Spring</location>
,
<collectingMunicipality id="9350AC87BD1749134D502ED8FF558BB3" pageId="4" pageNumber="477">Abyek City</collectingMunicipality>
,
@ -95,11 +98,12 @@ on
</date>
</materialsCitation>
.
<materialsCitation id="C3E33CA0BD17491348DF2E97FB958B97" box="[221,1032,1483,1510]" collectingDate="2022-06-22" collectionCode="ZCRU" collectorName="S. A. Mirghaffari" country="Iran" latitude="36.190556" location="Samaq Abad Spring" longLatPrecision="19" longitude="50.614998" municipality="Abyek City" pageId="4" pageNumber="477" specimenCount="6" stateProvince="Qazvin" typeStatus="paratype">
<materialsCitation id="C3E33CA0BD17491348DF2E97FB958B97" ID-GBIF-Occurrence="5169430301" box="[221,1032,1483,1510]" collectingDate="2022-06-22" collectionCode="ZCRU" collectorName="S. A. Mirghaffari" country="Iran" latitude="36.190556" location="Samaq Abad Spring" longLatPrecision="19" longitude="50.614998" municipality="Abyek City" pageId="4" pageNumber="477" specimenCode="Amph.1504" specimenCount="6" stateProvince="Qazvin" typeStatus="paratype">
<typeStatus id="AC30885FBD17491348DF2E97FED38B94" box="[221,334,1484,1510]" pageId="4" pageNumber="477" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="159AAE38BD174913495B2E90FE398B97" box="[345,420,1483,1509]" pageId="4" pageNumber="477">ZCRU</collectionCode>
Amph.1504,
<specimenCode id="232D9E86BD17491349AB2E90FDAC8B94" box="[425,561,1483,1510]" pageId="4" pageNumber="477">Amph.1504</specimenCode>
,
<specimenCount id="658DFD74BD1749134A3E2E90FD418B97" box="[572,732,1483,1509]" count="6" pageId="4" pageNumber="477" type="generic">six specimens</specimenCount>
, data same as for holotype
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@ -135,6 +139,720 @@ Antennae I. Antennae I 0.47 of body length. Peduncular articles 13 progressiv
<figureCitation id="EBB02A78BD174913489D2C98FE8889AF" box="[159,277,1987,2013]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[222,1365,191,1909]" captionTargetId="figure-26@8.[212,1375,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Niphargus qazvinensis sp. nov., Samaq Abad Spring, male 8 mm (holotype). A, Head; B, Antenna I; C, Antenna II; D, Mandibular palp; E-F, Maxilla I; G, Right mandible; H, Left mandible. Scale bars: 1= 0.25 mm (GH); 2= 0.5 mm (A, DF); 3= 1mm (BC)." figureDoi="http://doi.org/10.5281/zenodo.15561700" httpUri="https://zenodo.org/record/15561700/files/figure.png" pageId="4" pageNumber="477">Figure 4B</figureCitation>
).
</paragraph>
<caption id="27F46675BD16491248952A90F9038C7B" ID-Table-UUID="27F46675BD16491248952A90F9038C7B" httpUri="http://table.plazi.org/id/27F46675BD16491248952A90F9038C7B" pageId="5" pageNumber="478" startId="5.[151,243,459,485]" targetBox="[162,2008,536,1729]" targetIsTable="true" targetPageId="5">
<paragraph id="733436FDBD16491248952A90F9038C7B" blockId="5.[151,2037,459,521]" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD16491248952A90FE938F97" bold="true" box="[151,270,459,485]" pageId="5" pageNumber="478">TABLE 1.</emphasis>
Kimura 2-parameter-distances (%) between Iranian species and new collected populations of the genus
<emphasis id="41FFEAEFBD1649124D5F2A90FA508F97" box="[1373,1485,459,485]" italics="true" pageId="5" pageNumber="478">Niphargus</emphasis>
Schiödte, 1849 (based on 28S ribosomal DNA gene (below diagonal) and mtDNA (COI) gene (above diagonal). Symbol:
<emphasis id="41FFEAEFBD1649124B7F2AB4FC178C7B" bold="true" box="[893,906,495,521]" pageId="5" pageNumber="478">*</emphasis>
, indicates that the sequencing has failed and comparison cannot be made.
</paragraph>
</caption>
<paragraph id="733436FDBD16491249542943F84588B3" pageId="5" pageNumber="478">
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<th id="8E6A5DC3BD16B6E849542943FE158C5C" box="[342,392,536,558]" gridcol="1" gridrow="0" pageId="5" pageNumber="478">1</th>
<th id="8E6A5DC3BD16B6E849A12943FE4B8C5C" box="[419,470,536,558]" gridcol="2" gridrow="0" pageId="5" pageNumber="478">2</th>
<th id="8E6A5DC3BD16B6E849F32943FDB98C5C" box="[497,548,536,558]" gridcol="3" gridrow="0" pageId="5" pageNumber="478">3</th>
<th id="8E6A5DC3BD16B6E84A3C2943FDEC8C5C" box="[574,625,536,558]" gridcol="4" gridrow="0" pageId="5" pageNumber="478">4</th>
<th id="8E6A5DC3BD16B6E84A8E2943FD228C5C" box="[652,703,536,558]" gridcol="5" gridrow="0" pageId="5" pageNumber="478">5</th>
<th id="8E6A5DC3BD16B6E84ADB2943FC918C5C" box="[729,780,536,558]" gridcol="6" gridrow="0" pageId="5" pageNumber="478">6</th>
<th id="8E6A5DC3BD16B6E84B242943FCC48C5C" box="[806,857,536,558]" gridcol="7" gridrow="0" pageId="5" pageNumber="478">7</th>
<th id="8E6A5DC3BD16B6E84B762943FC3A8C5C" box="[884,935,536,558]" gridcol="8" gridrow="0" pageId="5" pageNumber="478">8</th>
<th id="8E6A5DC3BD16B6E84BC32943FC698C5C" box="[961,1012,536,558]" gridcol="9" gridrow="0" pageId="5" pageNumber="478">9</th>
<th id="8E6A5DC3BD16B6E84C0D2943FBDF8C5C" box="[1039,1090,536,558]" gridcol="10" gridrow="0" pageId="5" pageNumber="478">10</th>
<th id="8E6A5DC3BD16B6E84C5E2943FB128C5C" box="[1116,1167,536,558]" gridcol="11" gridrow="0" pageId="5" pageNumber="478">11</th>
<th id="8E6A5DC3BD16B6E84CA82943FB408C5C" box="[1194,1245,536,558]" gridcol="12" gridrow="0" pageId="5" pageNumber="478">12</th>
<th id="8E6A5DC3BD16B6E84CF52943FAB78C5C" box="[1271,1322,536,558]" gridcol="13" gridrow="0" pageId="5" pageNumber="478">13</th>
<th id="8E6A5DC3BD16B6E84D472943FAE58C5C" box="[1349,1400,536,558]" gridcol="14" gridrow="0" pageId="5" pageNumber="478">14</th>
<th id="8E6A5DC3BD16B6E84D902943FA588C5C" box="[1426,1477,536,558]" gridcol="15" gridrow="0" pageId="5" pageNumber="478">15</th>
<th id="8E6A5DC3BD16B6E84DDD2943F98F8C5C" box="[1503,1554,536,558]" gridcol="16" gridrow="0" pageId="5" pageNumber="478">16</th>
<th id="8E6A5DC3BD16B6E84E2F2943F9FD8C5C" box="[1581,1632,536,558]" gridcol="17" gridrow="0" pageId="5" pageNumber="478">17</th>
<th id="8E6A5DC3BD16B6E84E782943F9308C5C" box="[1658,1709,536,558]" gridcol="18" gridrow="0" pageId="5" pageNumber="478">18</th>
<th id="8E6A5DC3BD16B6E84ECA2943F96D8C5C" box="[1736,1776,536,558]" gridcol="19" gridrow="0" pageId="5" pageNumber="478">19</th>
<th id="8E6A5DC3BD16B6E84F172943F8A08C5C" box="[1813,1853,536,558]" gridcol="20" gridrow="0" pageId="5" pageNumber="478">20</th>
<th id="8E6A5DC3BD16B6E84F612943F8168C5C" box="[1891,1931,536,558]" gridcol="21" gridrow="0" pageId="5" pageNumber="478">21</th>
<th id="8E6A5DC3BD16B6E84FB22943F8458C5C" box="[1968,2008,536,558]" gridcol="22" gridrow="0" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124FB22943F85A8C5C" bold="true" box="[1968,1991,536,558]" pageId="5" pageNumber="478">22</emphasis>
</th>
</tr>
<tr id="CDBB34BFBD16B6E848A02965F8458C0A" box="[162,2008,574,632]" gridrow="1" pageId="5" pageNumber="478" rowspan-1="1">
<th id="8E6A5DC3BD16B6E848A02965FEA08C0A" box="[162,317,574,632]" gridcol="0" gridrow="1" pageId="5" pageNumber="478">
1:
<emphasis id="41FFEAEFBD16491248B82965FEA08C26" box="[186,317,574,596]" italics="true" pageId="5" pageNumber="478">N. qazvinensis</emphasis>
sp.nov.
</th>
<td id="8E6A5DC3BD16B6E849A12965FE4B8C0A" box="[419,470,574,632]" gridcol="2" gridrow="1" pageId="5" pageNumber="478">12.79</td>
<td id="8E6A5DC3BD16B6E849F32965FDB98C0A" box="[497,548,574,632]" gridcol="3" gridrow="1" pageId="5" pageNumber="478">10.24</td>
<td id="8E6A5DC3BD16B6E84A3C2965FDEC8C0A" box="[574,625,574,632]" gridcol="4" gridrow="1" pageId="5" pageNumber="478">12.51</td>
<td id="8E6A5DC3BD16B6E84A8E2965FD228C0A" box="[652,703,574,632]" gridcol="5" gridrow="1" pageId="5" pageNumber="478">20.11</td>
<td id="8E6A5DC3BD16B6E84ADB2965FC918C0A" box="[729,780,574,632]" gridcol="6" gridrow="1" pageId="5" pageNumber="478">11.82</td>
<td id="8E6A5DC3BD16B6E84B242965FCC48C0A" box="[806,857,574,632]" gridcol="7" gridrow="1" pageId="5" pageNumber="478">9.57</td>
<td id="8E6A5DC3BD16B6E84B762965FC3A8C0A" box="[884,935,574,632]" gridcol="8" gridrow="1" pageId="5" pageNumber="478">14.63</td>
<td id="8E6A5DC3BD16B6E84BC32965FC698C0A" box="[961,1012,574,632]" gridcol="9" gridrow="1" pageId="5" pageNumber="478">10.9</td>
<td id="8E6A5DC3BD16B6E84C0D2965FBDF8C0A" box="[1039,1090,574,632]" gridcol="10" gridrow="1" pageId="5" pageNumber="478">11.59</td>
<td id="8E6A5DC3BD16B6E84C5E2965FB128C0A" box="[1116,1167,574,632]" gridcol="11" gridrow="1" pageId="5" pageNumber="478">11.14</td>
<td id="8E6A5DC3BD16B6E84CA82965FB408C0A" box="[1194,1245,574,632]" gridcol="12" gridrow="1" pageId="5" pageNumber="478">11.37</td>
<td id="8E6A5DC3BD16B6E84CF52965FAB78C0A" box="[1271,1322,574,632]" gridcol="13" gridrow="1" pageId="5" pageNumber="478">13</td>
<td id="8E6A5DC3BD16B6E84D472965FAE58C0A" box="[1349,1400,574,632]" gridcol="14" gridrow="1" pageId="5" pageNumber="478">10.9</td>
<td id="8E6A5DC3BD16B6E84D902965FA588C0A" box="[1426,1477,574,632]" gridcol="15" gridrow="1" pageId="5" pageNumber="478">13.92</td>
<td id="8E6A5DC3BD16B6E84DDD2965F98F8C0A" box="[1503,1554,574,632]" gridcol="16" gridrow="1" pageId="5" pageNumber="478">15.91</td>
<td id="8E6A5DC3BD16B6E84E2F2965F9FD8C0A" box="[1581,1632,574,632]" gridcol="17" gridrow="1" pageId="5" pageNumber="478">8.26</td>
<td id="8E6A5DC3BD16B6E84E782965F9308C0A" box="[1658,1709,574,632]" gridcol="18" gridrow="1" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E78290BF9308C14" bold="true" box="[1658,1709,592,614]" pageId="5" pageNumber="478">14.22</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2965F96D8C0A" box="[1736,1776,574,632]" gridcol="19" gridrow="1" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172965F8A08C0A" box="[1813,1853,574,632]" gridcol="20" gridrow="1" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612965F8168C0A" box="[1891,1931,574,632]" gridcol="21" gridrow="1" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22965F8458C0A" box="[1968,2008,574,632]" gridcol="22" gridrow="1" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A029D3F8458CED" box="[162,2008,648,671]" gridrow="2" pageId="5" pageNumber="478" rowspan-2="1">
<th id="8E6A5DC3BD16B6E848A029D3FEA08CED" box="[162,317,648,671]" gridcol="0" gridrow="2" pageId="5" pageNumber="478">
2:
<emphasis id="41FFEAEFBD16491248B829D2FE868CED" box="[186,283,649,671]" italics="true" pageId="5" pageNumber="478">N. alisadri</emphasis>
</th>
<td id="8E6A5DC3BD16B6E8495429D3FE158CED" box="[342,392,648,671]" gridcol="1" gridrow="2" pageId="5" pageNumber="478">2.2</td>
<td id="8E6A5DC3BD16B6E849F329D3FDB98CED" box="[497,548,648,671]" gridcol="3" gridrow="2" pageId="5" pageNumber="478">11.03</td>
<td id="8E6A5DC3BD16B6E84A3C29D3FDEC8CED" box="[574,625,648,671]" gridcol="4" gridrow="2" pageId="5" pageNumber="478">16.88</td>
<td id="8E6A5DC3BD16B6E84A8E29D3FD228CED" box="[652,703,648,671]" gridcol="5" gridrow="2" pageId="5" pageNumber="478">20.91</td>
<td id="8E6A5DC3BD16B6E84ADB29D3FC918CED" box="[729,780,648,671]" gridcol="6" gridrow="2" pageId="5" pageNumber="478">12.35</td>
<td id="8E6A5DC3BD16B6E84B2429D3FCC48CED" box="[806,857,648,671]" gridcol="7" gridrow="2" pageId="5" pageNumber="478">12.16</td>
<td id="8E6A5DC3BD16B6E84B7629D3FC3A8CED" box="[884,935,648,671]" gridcol="8" gridrow="2" pageId="5" pageNumber="478">11.9</td>
<td id="8E6A5DC3BD16B6E84BC329D3FC698CED" box="[961,1012,648,671]" gridcol="9" gridrow="2" pageId="5" pageNumber="478">13.08</td>
<td id="8E6A5DC3BD16B6E84C0D29D3FBDF8CED" box="[1039,1090,648,671]" gridcol="10" gridrow="2" pageId="5" pageNumber="478">12.38</td>
<td id="8E6A5DC3BD16B6E84C5E29D3FB128CED" box="[1116,1167,648,671]" gridcol="11" gridrow="2" pageId="5" pageNumber="478">12.1</td>
<td id="8E6A5DC3BD16B6E84CA829D3FB408CED" box="[1194,1245,648,671]" gridcol="12" gridrow="2" pageId="5" pageNumber="478">12.36</td>
<td id="8E6A5DC3BD16B6E84CF529D3FAB78CED" box="[1271,1322,648,671]" gridcol="13" gridrow="2" pageId="5" pageNumber="478">15.41</td>
<td id="8E6A5DC3BD16B6E84D4729D3FAE58CED" box="[1349,1400,648,671]" gridcol="14" gridrow="2" pageId="5" pageNumber="478">12.36</td>
<td id="8E6A5DC3BD16B6E84D9029D3FA588CED" box="[1426,1477,648,671]" gridcol="15" gridrow="2" pageId="5" pageNumber="478">15.76</td>
<td id="8E6A5DC3BD16B6E84DDD29D3F98F8CED" box="[1503,1554,648,671]" gridcol="16" gridrow="2" pageId="5" pageNumber="478">14.15</td>
<td id="8E6A5DC3BD16B6E84E2F29D3F9FD8CED" box="[1581,1632,648,671]" gridcol="17" gridrow="2" pageId="5" pageNumber="478">12.15</td>
<td id="8E6A5DC3BD16B6E84E7829D3F9308CED" box="[1658,1709,648,671]" gridcol="18" gridrow="2" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E7829D3F9308CEC" bold="true" box="[1658,1709,648,670]" pageId="5" pageNumber="478">15.93</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA29D3F96D8CED" box="[1736,1776,648,671]" gridcol="19" gridrow="2" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F1729D3F8A08CED" box="[1813,1853,648,671]" gridcol="20" gridrow="2" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F6129D3F8168CED" box="[1891,1931,648,671]" gridcol="21" gridrow="2" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB229D3F8458CED" box="[1968,2008,648,671]" gridcol="22" gridrow="2" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A029F4F8458CB7" box="[162,2008,687,709]" gridrow="3" pageId="5" pageNumber="478" rowspan-3="1">
<th id="8E6A5DC3BD16B6E848A029F4FEA08CB7" box="[162,317,687,709]" gridcol="0" gridrow="3" pageId="5" pageNumber="478">
3:
<emphasis id="41FFEAEFBD16491248B829F4FEAB8CB7" box="[186,310,687,709]" italics="true" pageId="5" pageNumber="478">N. bisitunicus</emphasis>
</th>
<td id="8E6A5DC3BD16B6E8495429F4FE158CB7" box="[342,392,687,709]" gridcol="1" gridrow="3" pageId="5" pageNumber="478">2.07</td>
<td id="8E6A5DC3BD16B6E849A129F4FE4B8CB7" box="[419,470,687,709]" gridcol="2" gridrow="3" pageId="5" pageNumber="478">1.15</td>
<td id="8E6A5DC3BD16B6E84A3C29F4FDEC8CB7" box="[574,625,687,709]" gridcol="4" gridrow="3" pageId="5" pageNumber="478">14.7</td>
<td id="8E6A5DC3BD16B6E84A8E29F4FD228CB7" box="[652,703,687,709]" gridcol="5" gridrow="3" pageId="5" pageNumber="478">21.67</td>
<td id="8E6A5DC3BD16B6E84ADB29F4FC918CB7" box="[729,780,687,709]" gridcol="6" gridrow="3" pageId="5" pageNumber="478">10.49</td>
<td id="8E6A5DC3BD16B6E84B2429F4FCC48CB7" box="[806,857,687,709]" gridcol="7" gridrow="3" pageId="5" pageNumber="478">10.72</td>
<td id="8E6A5DC3BD16B6E84B7629F4FC3A8CB7" box="[884,935,687,709]" gridcol="8" gridrow="3" pageId="5" pageNumber="478">10.51</td>
<td id="8E6A5DC3BD16B6E84BC329F4FC698CB7" box="[961,1012,687,709]" gridcol="9" gridrow="3" pageId="5" pageNumber="478">11.16</td>
<td id="8E6A5DC3BD16B6E84C0D29F4FBDF8CB7" box="[1039,1090,687,709]" gridcol="10" gridrow="3" pageId="5" pageNumber="478">11.16</td>
<td id="8E6A5DC3BD16B6E84C5E29F4FB128CB7" box="[1116,1167,687,709]" gridcol="11" gridrow="3" pageId="5" pageNumber="478">12.07</td>
<td id="8E6A5DC3BD16B6E84CA829F4FB408CB7" box="[1194,1245,687,709]" gridcol="12" gridrow="3" pageId="5" pageNumber="478">12.8</td>
<td id="8E6A5DC3BD16B6E84CF529F4FAB78CB7" box="[1271,1322,687,709]" gridcol="13" gridrow="3" pageId="5" pageNumber="478">14.9</td>
<td id="8E6A5DC3BD16B6E84D4729F4FAE58CB7" box="[1349,1400,687,709]" gridcol="14" gridrow="3" pageId="5" pageNumber="478">8.7</td>
<td id="8E6A5DC3BD16B6E84D9029F4FA588CB7" box="[1426,1477,687,709]" gridcol="15" gridrow="3" pageId="5" pageNumber="478">14.71</td>
<td id="8E6A5DC3BD16B6E84DDD29F4F98F8CB7" box="[1503,1554,687,709]" gridcol="16" gridrow="3" pageId="5" pageNumber="478">13.6</td>
<td id="8E6A5DC3BD16B6E84E2F29F4F9FD8CB7" box="[1581,1632,687,709]" gridcol="17" gridrow="3" pageId="5" pageNumber="478">9.8</td>
<td id="8E6A5DC3BD16B6E84E7829F4F9308CB7" box="[1658,1709,687,709]" gridcol="18" gridrow="3" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E7829F4F93F8CB7" bold="true" box="[1658,1698,687,709]" pageId="5" pageNumber="478">13.5</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA29F4F96D8CB7" box="[1736,1776,687,709]" gridcol="19" gridrow="3" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F1729F4F8A08CB7" box="[1813,1853,687,709]" gridcol="20" gridrow="3" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F6129F4F8168CB7" box="[1891,1931,687,709]" gridcol="21" gridrow="3" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB229F4F8458CB7" box="[1968,2008,687,709]" gridcol="22" gridrow="3" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A0298EF8458C99" box="[162,2008,725,747]" gridrow="4" pageId="5" pageNumber="478" rowspan-4="1">
<th id="8E6A5DC3BD16B6E848A0298EFEA08C99" box="[162,317,725,747]" gridcol="0" gridrow="4" pageId="5" pageNumber="478">
4:
<emphasis id="41FFEAEFBD16491248B8298EFE948C99" box="[186,265,725,747]" italics="true" pageId="5" pageNumber="478">N. borisi</emphasis>
</th>
<td id="8E6A5DC3BD16B6E84954298EFE158C99" box="[342,392,725,747]" gridcol="1" gridrow="4" pageId="5" pageNumber="478">3.52</td>
<td id="8E6A5DC3BD16B6E849A1298EFE4B8C99" box="[419,470,725,747]" gridcol="2" gridrow="4" pageId="5" pageNumber="478">2.17</td>
<td id="8E6A5DC3BD16B6E849F3298EFDB98C99" box="[497,548,725,747]" gridcol="3" gridrow="4" pageId="5" pageNumber="478">2.7</td>
<td id="8E6A5DC3BD16B6E84A8E298EFD228C99" box="[652,703,725,747]" gridcol="5" gridrow="4" pageId="5" pageNumber="478">23.23</td>
<td id="8E6A5DC3BD16B6E84ADB298EFC918C99" box="[729,780,725,747]" gridcol="6" gridrow="4" pageId="5" pageNumber="478">13.19</td>
<td id="8E6A5DC3BD16B6E84B24298EFCC48C99" box="[806,857,725,747]" gridcol="7" gridrow="4" pageId="5" pageNumber="478">14.12</td>
<td id="8E6A5DC3BD16B6E84B76298EFC3A8C99" box="[884,935,725,747]" gridcol="8" gridrow="4" pageId="5" pageNumber="478">17.07</td>
<td id="8E6A5DC3BD16B6E84BC3298EFC698C99" box="[961,1012,725,747]" gridcol="9" gridrow="4" pageId="5" pageNumber="478">15.34</td>
<td id="8E6A5DC3BD16B6E84C0D298EFBDF8C99" box="[1039,1090,725,747]" gridcol="10" gridrow="4" pageId="5" pageNumber="478">15.82</td>
<td id="8E6A5DC3BD16B6E84C5E298EFB128C99" box="[1116,1167,725,747]" gridcol="11" gridrow="4" pageId="5" pageNumber="478">14.88</td>
<td id="8E6A5DC3BD16B6E84CA8298EFB408C99" box="[1194,1245,725,747]" gridcol="12" gridrow="4" pageId="5" pageNumber="478">14.37</td>
<td id="8E6A5DC3BD16B6E84CF5298EFAB78C99" box="[1271,1322,725,747]" gridcol="13" gridrow="4" pageId="5" pageNumber="478">10.04</td>
<td id="8E6A5DC3BD16B6E84D47298EFAE58C99" box="[1349,1400,725,747]" gridcol="14" gridrow="4" pageId="5" pageNumber="478">13.9</td>
<td id="8E6A5DC3BD16B6E84D90298EFA588C99" box="[1426,1477,725,747]" gridcol="15" gridrow="4" pageId="5" pageNumber="478">12.76</td>
<td id="8E6A5DC3BD16B6E84DDD298EF98F8C99" box="[1503,1554,725,747]" gridcol="16" gridrow="4" pageId="5" pageNumber="478">17.86</td>
<td id="8E6A5DC3BD16B6E84E2F298EF9FD8C99" box="[1581,1632,725,747]" gridcol="17" gridrow="4" pageId="5" pageNumber="478">13.91</td>
<td id="8E6A5DC3BD16B6E84E78298EF9308C99" box="[1658,1709,725,747]" gridcol="18" gridrow="4" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E78298EF93F8C99" bold="true" box="[1658,1698,725,747]" pageId="5" pageNumber="478">7.38</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA298EF96D8C99" box="[1736,1776,725,747]" gridcol="19" gridrow="4" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F17298EF8A08C99" box="[1813,1853,725,747]" gridcol="20" gridrow="4" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F61298EF8168C99" box="[1891,1931,725,747]" gridcol="21" gridrow="4" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB2298EF8458C99" box="[1968,2008,725,747]" gridcol="22" gridrow="4" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A029A7F8458D60" box="[162,2008,764,786]" gridrow="5" pageId="5" pageNumber="478" rowspan-5="1">
<th id="8E6A5DC3BD16B6E848A029A7FEA08D60" box="[162,317,764,786]" gridcol="0" gridrow="5" pageId="5" pageNumber="478">
5:
<emphasis id="41FFEAEFBD16491248B829A7FE898D60" box="[186,276,764,786]" italics="true" pageId="5" pageNumber="478">N. daniali</emphasis>
</th>
<td id="8E6A5DC3BD16B6E8495429A7FE158D60" box="[342,392,764,786]" gridcol="1" gridrow="5" pageId="5" pageNumber="478">11.91</td>
<td id="8E6A5DC3BD16B6E849A129A7FE4B8D60" box="[419,470,764,786]" gridcol="2" gridrow="5" pageId="5" pageNumber="478">11.2</td>
<td id="8E6A5DC3BD16B6E849F329A7FDB98D60" box="[497,548,764,786]" gridcol="3" gridrow="5" pageId="5" pageNumber="478">11.67</td>
<td id="8E6A5DC3BD16B6E84A3C29A7FDEC8D60" box="[574,625,764,786]" gridcol="4" gridrow="5" pageId="5" pageNumber="478">11.49</td>
<td id="8E6A5DC3BD16B6E84ADB29A7FC918D60" box="[729,780,764,786]" gridcol="6" gridrow="5" pageId="5" pageNumber="478">17.12</td>
<td id="8E6A5DC3BD16B6E84B2429A7FCC48D60" box="[806,857,764,786]" gridcol="7" gridrow="5" pageId="5" pageNumber="478">19.62</td>
<td id="8E6A5DC3BD16B6E84B7629A7FC3A8D60" box="[884,935,764,786]" gridcol="8" gridrow="5" pageId="5" pageNumber="478">22.18</td>
<td id="8E6A5DC3BD16B6E84BC329A7FC698D60" box="[961,1012,764,786]" gridcol="9" gridrow="5" pageId="5" pageNumber="478">21.96</td>
<td id="8E6A5DC3BD16B6E84C0D29A7FBDF8D60" box="[1039,1090,764,786]" gridcol="10" gridrow="5" pageId="5" pageNumber="478">19.89</td>
<td id="8E6A5DC3BD16B6E84C5E29A7FB128D60" box="[1116,1167,764,786]" gridcol="11" gridrow="5" pageId="5" pageNumber="478">19.87</td>
<td id="8E6A5DC3BD16B6E84CA829A7FB408D60" box="[1194,1245,764,786]" gridcol="12" gridrow="5" pageId="5" pageNumber="478">17.06</td>
<td id="8E6A5DC3BD16B6E84CF529A7FAB78D60" box="[1271,1322,764,786]" gridcol="13" gridrow="5" pageId="5" pageNumber="478">23.05</td>
<td id="8E6A5DC3BD16B6E84D4729A7FAE58D60" box="[1349,1400,764,786]" gridcol="14" gridrow="5" pageId="5" pageNumber="478">17.82</td>
<td id="8E6A5DC3BD16B6E84D9029A7FA588D60" box="[1426,1477,764,786]" gridcol="15" gridrow="5" pageId="5" pageNumber="478">22.18</td>
<td id="8E6A5DC3BD16B6E84DDD29A7F98F8D60" box="[1503,1554,764,786]" gridcol="16" gridrow="5" pageId="5" pageNumber="478">21.7</td>
<td id="8E6A5DC3BD16B6E84E2F29A7F9FD8D60" box="[1581,1632,764,786]" gridcol="17" gridrow="5" pageId="5" pageNumber="478">18.59</td>
<td id="8E6A5DC3BD16B6E84E7829A7F9308D60" box="[1658,1709,764,786]" gridcol="18" gridrow="5" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E7829A7F9308D60" bold="true" box="[1658,1709,764,786]" pageId="5" pageNumber="478">21.93</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA29A7F96D8D60" box="[1736,1776,764,786]" gridcol="19" gridrow="5" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F1729A7F8A08D60" box="[1813,1853,764,786]" gridcol="20" gridrow="5" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F6129A7F8168D60" box="[1891,1931,764,786]" gridcol="21" gridrow="5" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB229A7F8458D60" box="[1968,2008,764,786]" gridcol="22" gridrow="5" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02879F8458D4A" box="[162,2008,802,824]" gridrow="6" pageId="5" pageNumber="478" rowspan-6="1">
<th id="8E6A5DC3BD16B6E848A02879FEA08D4A" box="[162,317,802,824]" gridcol="0" gridrow="6" pageId="5" pageNumber="478">
6:
<emphasis id="41FFEAEFBD16491248B82879FE838D4A" box="[186,286,802,824]" italics="true" pageId="5" pageNumber="478">N. darvishi</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542879FE158D4A" box="[342,392,802,824]" gridcol="1" gridrow="6" pageId="5" pageNumber="478">2.46</td>
<td id="8E6A5DC3BD16B6E849A12879FE4B8D4A" box="[419,470,802,824]" gridcol="2" gridrow="6" pageId="5" pageNumber="478">0.76</td>
<td id="8E6A5DC3BD16B6E849F32879FDB98D4A" box="[497,548,802,824]" gridcol="3" gridrow="6" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84A3C2879FDEC8D4A" box="[574,625,802,824]" gridcol="4" gridrow="6" pageId="5" pageNumber="478">2.69</td>
<td id="8E6A5DC3BD16B6E84A8E2879FD228D4A" box="[652,703,802,824]" gridcol="5" gridrow="6" pageId="5" pageNumber="478">11.51</td>
<td id="8E6A5DC3BD16B6E84B242879FCC48D4A" box="[806,857,802,824]" gridcol="7" gridrow="6" pageId="5" pageNumber="478">9.58</td>
<td id="8E6A5DC3BD16B6E84B762879FC3A8D4A" box="[884,935,802,824]" gridcol="8" gridrow="6" pageId="5" pageNumber="478">12.08</td>
<td id="8E6A5DC3BD16B6E84BC32879FC698D4A" box="[961,1012,802,824]" gridcol="9" gridrow="6" pageId="5" pageNumber="478">10.27</td>
<td id="8E6A5DC3BD16B6E84C0D2879FBDF8D4A" box="[1039,1090,802,824]" gridcol="10" gridrow="6" pageId="5" pageNumber="478">9.82</td>
<td id="8E6A5DC3BD16B6E84C5E2879FB128D4A" box="[1116,1167,802,824]" gridcol="11" gridrow="6" pageId="5" pageNumber="478">12.27</td>
<td id="8E6A5DC3BD16B6E84CA82879FB408D4A" box="[1194,1245,802,824]" gridcol="12" gridrow="6" pageId="5" pageNumber="478">10.01</td>
<td id="8E6A5DC3BD16B6E84CF52879FAB78D4A" box="[1271,1322,802,824]" gridcol="13" gridrow="6" pageId="5" pageNumber="478">13.65</td>
<td id="8E6A5DC3BD16B6E84D472879FAE58D4A" box="[1349,1400,802,824]" gridcol="14" gridrow="6" pageId="5" pageNumber="478">3.82</td>
<td id="8E6A5DC3BD16B6E84D902879FA588D4A" box="[1426,1477,802,824]" gridcol="15" gridrow="6" pageId="5" pageNumber="478">14.86</td>
<td id="8E6A5DC3BD16B6E84DDD2879F98F8D4A" box="[1503,1554,802,824]" gridcol="16" gridrow="6" pageId="5" pageNumber="478">13.33</td>
<td id="8E6A5DC3BD16B6E84E2F2879F9FD8D4A" box="[1581,1632,802,824]" gridcol="17" gridrow="6" pageId="5" pageNumber="478">8.69</td>
<td id="8E6A5DC3BD16B6E84E782879F9308D4A" box="[1658,1709,802,824]" gridcol="18" gridrow="6" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782879F93D8D4A" bold="true" box="[1658,1696,802,824]" pageId="5" pageNumber="478">11.8</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2879F96D8D4A" box="[1736,1776,802,824]" gridcol="19" gridrow="6" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172879F8A08D4A" box="[1813,1853,802,824]" gridcol="20" gridrow="6" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612879F8168D4A" box="[1891,1931,802,824]" gridcol="21" gridrow="6" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22879F8458D4A" box="[1968,2008,802,824]" gridcol="22" gridrow="6" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02812F8458D2D" box="[162,2008,841,863]" gridrow="7" pageId="5" pageNumber="478" rowspan-7="1">
<th id="8E6A5DC3BD16B6E848A02812FEA08D2D" box="[162,317,841,863]" gridcol="0" gridrow="7" pageId="5" pageNumber="478">
7:
<emphasis id="41FFEAEFBD16491248B82812FE9E8D2D" box="[186,259,841,863]" italics="true" pageId="5" pageNumber="478">N. fiseri</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542812FE158D2D" box="[342,392,841,863]" gridcol="1" gridrow="7" pageId="5" pageNumber="478">1.8</td>
<td id="8E6A5DC3BD16B6E849A12812FE4B8D2D" box="[419,470,841,863]" gridcol="2" gridrow="7" pageId="5" pageNumber="478">2.57</td>
<td id="8E6A5DC3BD16B6E849F32812FDB98D2D" box="[497,548,841,863]" gridcol="3" gridrow="7" pageId="5" pageNumber="478">2.96</td>
<td id="8E6A5DC3BD16B6E84A3C2812FDEC8D2D" box="[574,625,841,863]" gridcol="4" gridrow="7" pageId="5" pageNumber="478">4.29</td>
<td id="8E6A5DC3BD16B6E84A8E2812FD228D2D" box="[652,703,841,863]" gridcol="5" gridrow="7" pageId="5" pageNumber="478">11.47</td>
<td id="8E6A5DC3BD16B6E84ADB2812FC918D2D" box="[729,780,841,863]" gridcol="6" gridrow="7" pageId="5" pageNumber="478">3.09</td>
<td id="8E6A5DC3BD16B6E84B762812FC3A8D2D" box="[884,935,841,863]" gridcol="8" gridrow="7" pageId="5" pageNumber="478">12.52</td>
<td id="8E6A5DC3BD16B6E84BC32812FC698D2D" box="[961,1012,841,863]" gridcol="9" gridrow="7" pageId="5" pageNumber="478">12.98</td>
<td id="8E6A5DC3BD16B6E84C0D2812FBDF8D2D" box="[1039,1090,841,863]" gridcol="10" gridrow="7" pageId="5" pageNumber="478">11.37</td>
<td id="8E6A5DC3BD16B6E84C5E2812FB128D2D" box="[1116,1167,841,863]" gridcol="11" gridrow="7" pageId="5" pageNumber="478">11.59</td>
<td id="8E6A5DC3BD16B6E84CA82812FB408D2D" box="[1194,1245,841,863]" gridcol="12" gridrow="7" pageId="5" pageNumber="478">11.37</td>
<td id="8E6A5DC3BD16B6E84CF52812FAB78D2D" box="[1271,1322,841,863]" gridcol="13" gridrow="7" pageId="5" pageNumber="478">12.98</td>
<td id="8E6A5DC3BD16B6E84D472812FAE58D2D" box="[1349,1400,841,863]" gridcol="14" gridrow="7" pageId="5" pageNumber="478">10.02</td>
<td id="8E6A5DC3BD16B6E84D902812FA588D2D" box="[1426,1477,841,863]" gridcol="15" gridrow="7" pageId="5" pageNumber="478">13.95</td>
<td id="8E6A5DC3BD16B6E84DDD2812F98F8D2D" box="[1503,1554,841,863]" gridcol="16" gridrow="7" pageId="5" pageNumber="478">13.78</td>
<td id="8E6A5DC3BD16B6E84E2F2812F9FD8D2D" box="[1581,1632,841,863]" gridcol="17" gridrow="7" pageId="5" pageNumber="478">7.62</td>
<td id="8E6A5DC3BD16B6E84E782812F9308D2D" box="[1658,1709,841,863]" gridcol="18" gridrow="7" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782812F9308D2D" bold="true" box="[1658,1709,841,863]" pageId="5" pageNumber="478">15.14</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2812F96D8D2D" box="[1736,1776,841,863]" gridcol="19" gridrow="7" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172812F8A08D2D" box="[1813,1853,841,863]" gridcol="20" gridrow="7" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612812F8168D2D" box="[1891,1931,841,863]" gridcol="21" gridrow="7" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22812F8458D2D" box="[1968,2008,841,863]" gridcol="22" gridrow="7" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02834F8458DF7" box="[162,2008,879,901]" gridrow="8" pageId="5" pageNumber="478" rowspan-8="1">
<th id="8E6A5DC3BD16B6E848A02834FEA08DF7" box="[162,317,879,901]" gridcol="0" gridrow="8" pageId="5" pageNumber="478">
8:
<emphasis id="41FFEAEFBD16491248B82834FEB28DF7" box="[186,303,879,901]" italics="true" pageId="5" pageNumber="478">N. hosseiniei</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542834FE158DF7" box="[342,392,879,901]" gridcol="1" gridrow="8" pageId="5" pageNumber="478">1.94</td>
<td id="8E6A5DC3BD16B6E849A12834FE4B8DF7" box="[419,470,879,901]" gridcol="2" gridrow="8" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E849F32834FDB98DF7" box="[497,548,879,901]" gridcol="3" gridrow="8" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84A3C2834FDEC8DF7" box="[574,625,879,901]" gridcol="4" gridrow="8" pageId="5" pageNumber="478">2.82</td>
<td id="8E6A5DC3BD16B6E84A8E2834FD228DF7" box="[652,703,879,901]" gridcol="5" gridrow="8" pageId="5" pageNumber="478">12.28</td>
<td id="8E6A5DC3BD16B6E84ADB2834FC918DF7" box="[729,780,879,901]" gridcol="6" gridrow="8" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84B242834FCC48DF7" box="[806,857,879,901]" gridcol="7" gridrow="8" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84BC32834FC698DF7" box="[961,1012,879,901]" gridcol="9" gridrow="8" pageId="5" pageNumber="478">13.99</td>
<td id="8E6A5DC3BD16B6E84C0D2834FBDF8DF7" box="[1039,1090,879,901]" gridcol="10" gridrow="8" pageId="5" pageNumber="478">12.1</td>
<td id="8E6A5DC3BD16B6E84C5E2834FB128DF7" box="[1116,1167,879,901]" gridcol="11" gridrow="8" pageId="5" pageNumber="478">13.22</td>
<td id="8E6A5DC3BD16B6E84CA82834FB408DF7" box="[1194,1245,879,901]" gridcol="12" gridrow="8" pageId="5" pageNumber="478">14.18</td>
<td id="8E6A5DC3BD16B6E84CF52834FAB78DF7" box="[1271,1322,879,901]" gridcol="13" gridrow="8" pageId="5" pageNumber="478">13.44</td>
<td id="8E6A5DC3BD16B6E84D472834FAE58DF7" box="[1349,1400,879,901]" gridcol="14" gridrow="8" pageId="5" pageNumber="478">12.09</td>
<td id="8E6A5DC3BD16B6E84D902834FA588DF7" box="[1426,1477,879,901]" gridcol="15" gridrow="8" pageId="5" pageNumber="478">15.65</td>
<td id="8E6A5DC3BD16B6E84DDD2834F98F8DF7" box="[1503,1554,879,901]" gridcol="16" gridrow="8" pageId="5" pageNumber="478">9.65</td>
<td id="8E6A5DC3BD16B6E84E2F2834F9FD8DF7" box="[1581,1632,879,901]" gridcol="17" gridrow="8" pageId="5" pageNumber="478">13.7</td>
<td id="8E6A5DC3BD16B6E84E782834F9308DF7" box="[1658,1709,879,901]" gridcol="18" gridrow="8" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782834F9308DF7" bold="true" box="[1658,1709,879,901]" pageId="5" pageNumber="478">14.16</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2834F96D8DF7" box="[1736,1776,879,901]" gridcol="19" gridrow="8" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172834F8A08DF7" box="[1813,1853,879,901]" gridcol="20" gridrow="8" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612834F8168DF7" box="[1891,1931,879,901]" gridcol="21" gridrow="8" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22834F8458DF7" box="[1968,2008,879,901]" gridcol="22" gridrow="8" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A028CEF8458DDE" box="[162,2008,917,940]" gridrow="9" pageId="5" pageNumber="478" rowspan-9="1">
<th id="8E6A5DC3BD16B6E848A028CEFEA08DDE" box="[162,317,917,940]" gridcol="0" gridrow="9" pageId="5" pageNumber="478">
9:
<emphasis id="41FFEAEFBD16491248B828CDFEB78DDE" box="[186,298,918,940]" italics="true" pageId="5" pageNumber="478">N. ilamensis</emphasis>
</th>
<td id="8E6A5DC3BD16B6E8495428CEFE158DDE" box="[342,392,917,940]" gridcol="1" gridrow="9" pageId="5" pageNumber="478">1.94</td>
<td id="8E6A5DC3BD16B6E849A128CEFE4B8DDE" box="[419,470,917,940]" gridcol="2" gridrow="9" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E849F328CEFDB98DDE" box="[497,548,917,940]" gridcol="3" gridrow="9" pageId="5" pageNumber="478">1.15</td>
<td id="8E6A5DC3BD16B6E84A3C28CEFDEC8DDE" box="[574,625,917,940]" gridcol="4" gridrow="9" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84A8E28CEFD228DDE" box="[652,703,917,940]" gridcol="5" gridrow="9" pageId="5" pageNumber="478">11.84</td>
<td id="8E6A5DC3BD16B6E84ADB28CEFC918DDE" box="[729,780,917,940]" gridcol="6" gridrow="9" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84B2428CEFCC48DDE" box="[806,857,917,940]" gridcol="7" gridrow="9" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84B7628CEFC3A8DDE" box="[884,935,917,940]" gridcol="8" gridrow="9" pageId="5" pageNumber="478">0.89</td>
<td id="8E6A5DC3BD16B6E84C0D28CEFBDF8DDE" box="[1039,1090,917,940]" gridcol="10" gridrow="9" pageId="5" pageNumber="478">8.98</td>
<td id="8E6A5DC3BD16B6E84C5E28CEFB128DDE" box="[1116,1167,917,940]" gridcol="11" gridrow="9" pageId="5" pageNumber="478">12.52</td>
<td id="8E6A5DC3BD16B6E84CA828CEFB408DDE" box="[1194,1245,917,940]" gridcol="12" gridrow="9" pageId="5" pageNumber="478">11.87</td>
<td id="8E6A5DC3BD16B6E84CF528CEFAB78DDE" box="[1271,1322,917,940]" gridcol="13" gridrow="9" pageId="5" pageNumber="478">15.61</td>
<td id="8E6A5DC3BD16B6E84D4728CEFAE58DDE" box="[1349,1400,917,940]" gridcol="14" gridrow="9" pageId="5" pageNumber="478">10.07</td>
<td id="8E6A5DC3BD16B6E84D9028CEFA588DDE" box="[1426,1477,917,940]" gridcol="15" gridrow="9" pageId="5" pageNumber="478">14.67</td>
<td id="8E6A5DC3BD16B6E84DDD28CEF98F8DDE" box="[1503,1554,917,940]" gridcol="16" gridrow="9" pageId="5" pageNumber="478">13.31</td>
<td id="8E6A5DC3BD16B6E84E2F28CEF9FD8DDE" box="[1581,1632,917,940]" gridcol="17" gridrow="9" pageId="5" pageNumber="478">10.02</td>
<td id="8E6A5DC3BD16B6E84E7828CEF9308DDE" box="[1658,1709,917,940]" gridcol="18" gridrow="9" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E7828CEF9308DD9" bold="true" box="[1658,1709,917,939]" pageId="5" pageNumber="478">14.42</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA28CEF96D8DDE" box="[1736,1776,917,940]" gridcol="19" gridrow="9" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F1728CEF8A08DDE" box="[1813,1853,917,940]" gridcol="20" gridrow="9" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F6128CEF8168DDE" box="[1891,1931,917,940]" gridcol="21" gridrow="9" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB228CEF8458DDE" box="[1968,2008,917,940]" gridcol="22" gridrow="9" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A028E7F8458DA0" box="[162,2008,956,978]" gridrow="10" pageId="5" pageNumber="478" rowspan-10="1">
<th id="8E6A5DC3BD16B6E848A028E7FEA08DA0" box="[162,317,956,978]" gridcol="0" gridrow="10" pageId="5" pageNumber="478">
10:
<emphasis id="41FFEAEFBD16491248C728E7FEA08DA0" box="[197,317,956,978]" italics="true" pageId="5" pageNumber="478">N. khwarizmi</emphasis>
</th>
<td id="8E6A5DC3BD16B6E8495428E7FE158DA0" box="[342,392,956,978]" gridcol="1" gridrow="10" pageId="5" pageNumber="478">2.07</td>
<td id="8E6A5DC3BD16B6E849A128E7FE4B8DA0" box="[419,470,956,978]" gridcol="2" gridrow="10" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E849F328E7FDB98DA0" box="[497,548,956,978]" gridcol="3" gridrow="10" pageId="5" pageNumber="478">1.15</td>
<td id="8E6A5DC3BD16B6E84A3C28E7FDEC8DA0" box="[574,625,956,978]" gridcol="4" gridrow="10" pageId="5" pageNumber="478">2.56</td>
<td id="8E6A5DC3BD16B6E84A8E28E7FD228DA0" box="[652,703,956,978]" gridcol="5" gridrow="10" pageId="5" pageNumber="478">11.99</td>
<td id="8E6A5DC3BD16B6E84ADB28E7FC918DA0" box="[729,780,956,978]" gridcol="6" gridrow="10" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84B2428E7FCC48DA0" box="[806,857,956,978]" gridcol="7" gridrow="10" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84B7628E7FC3A8DA0" box="[884,935,956,978]" gridcol="8" gridrow="10" pageId="5" pageNumber="478">1.14</td>
<td id="8E6A5DC3BD16B6E84BC328E7FC698DA0" box="[961,1012,956,978]" gridcol="9" gridrow="10" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E84C5E28E7FB128DA0" box="[1116,1167,956,978]" gridcol="11" gridrow="10" pageId="5" pageNumber="478">11.61</td>
<td id="8E6A5DC3BD16B6E84CA828E7FB408DA0" box="[1194,1245,956,978]" gridcol="12" gridrow="10" pageId="5" pageNumber="478">10.52</td>
<td id="8E6A5DC3BD16B6E84CF528E7FAB78DA0" box="[1271,1322,956,978]" gridcol="13" gridrow="10" pageId="5" pageNumber="478">13.43</td>
<td id="8E6A5DC3BD16B6E84D4728E7FAE58DA0" box="[1349,1400,956,978]" gridcol="14" gridrow="10" pageId="5" pageNumber="478">8.95</td>
<td id="8E6A5DC3BD16B6E84D9028E7FA588DA0" box="[1426,1477,956,978]" gridcol="15" gridrow="10" pageId="5" pageNumber="478">15.85</td>
<td id="8E6A5DC3BD16B6E84DDD28E7F98F8DA0" box="[1503,1554,956,978]" gridcol="16" gridrow="10" pageId="5" pageNumber="478">15.53</td>
<td id="8E6A5DC3BD16B6E84E2F28E7F9FD8DA0" box="[1581,1632,956,978]" gridcol="17" gridrow="10" pageId="5" pageNumber="478">9.13</td>
<td id="8E6A5DC3BD16B6E84E7828E7F9308DA0" box="[1658,1709,956,978]" gridcol="18" gridrow="10" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E7828E7F9308DA0" bold="true" box="[1658,1709,956,978]" pageId="5" pageNumber="478">16.09</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA28E7F96D8DA0" box="[1736,1776,956,978]" gridcol="19" gridrow="10" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F1728E7F8A08DA0" box="[1813,1853,956,978]" gridcol="20" gridrow="10" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F6128E7F8168DA0" box="[1891,1931,956,978]" gridcol="21" gridrow="10" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB228E7F8458DA0" box="[1968,2008,956,978]" gridcol="22" gridrow="10" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A028B9F8458A6E" box="[162,2008,994,1052]" gridrow="11" pageId="5" pageNumber="478" rowspan-11="1">
<th id="8E6A5DC3BD16B6E848A028B9FEA08A6E" box="[162,317,994,1052]" gridcol="0" gridrow="11" pageId="5" pageNumber="478">
11:
<emphasis id="41FFEAEFBD16491248C628B9FF478D8A" box="[196,218,994,1016]" italics="true" pageId="5" pageNumber="478">N.</emphasis>
<emphasis id="41FFEAEFBD16491248A02F5DFEB98A6E" box="[162,292,1030,1052]" italics="true" pageId="5" pageNumber="478">kurdistanensis</emphasis>
</th>
<td id="8E6A5DC3BD16B6E8495428B9FE158A6E" box="[342,392,994,1052]" gridcol="1" gridrow="11" pageId="5" pageNumber="478">2.33</td>
<td id="8E6A5DC3BD16B6E849A128B9FE4B8A6E" box="[419,470,994,1052]" gridcol="2" gridrow="11" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E849F328B9FDB98A6E" box="[497,548,994,1052]" gridcol="3" gridrow="11" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84A3C28B9FDEC8A6E" box="[574,625,994,1052]" gridcol="4" gridrow="11" pageId="5" pageNumber="478">2.04</td>
<td id="8E6A5DC3BD16B6E84A8E28B9FD228A6E" box="[652,703,994,1052]" gridcol="5" gridrow="11" pageId="5" pageNumber="478">11.05</td>
<td id="8E6A5DC3BD16B6E84ADB28B9FC918A6E" box="[729,780,994,1052]" gridcol="6" gridrow="11" pageId="5" pageNumber="478">1.14</td>
<td id="8E6A5DC3BD16B6E84B2428B9FCC48A6E" box="[806,857,994,1052]" gridcol="7" gridrow="11" pageId="5" pageNumber="478">2.96</td>
<td id="8E6A5DC3BD16B6E84B7628B9FC3A8A6E" box="[884,935,994,1052]" gridcol="8" gridrow="11" pageId="5" pageNumber="478">1.66</td>
<td id="8E6A5DC3BD16B6E84BC328B9FC698A6E" box="[961,1012,994,1052]" gridcol="9" gridrow="11" pageId="5" pageNumber="478">1.66</td>
<td id="8E6A5DC3BD16B6E84C0D28B9FBDF8A6E" box="[1039,1090,994,1052]" gridcol="10" gridrow="11" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84CA828B9FB408A6E" box="[1194,1245,994,1052]" gridcol="12" gridrow="11" pageId="5" pageNumber="478">9.78</td>
<td id="8E6A5DC3BD16B6E84CF528B9FAB78A6E" box="[1271,1322,994,1052]" gridcol="13" gridrow="11" pageId="5" pageNumber="478">13.7</td>
<td id="8E6A5DC3BD16B6E84D4728B9FAE58A6E" box="[1349,1400,994,1052]" gridcol="14" gridrow="11" pageId="5" pageNumber="478">11.58</td>
<td id="8E6A5DC3BD16B6E84D9028B9FA588A6E" box="[1426,1477,994,1052]" gridcol="15" gridrow="11" pageId="5" pageNumber="478">14.7</td>
<td id="8E6A5DC3BD16B6E84DDD28B9F98F8A6E" box="[1503,1554,994,1052]" gridcol="16" gridrow="11" pageId="5" pageNumber="478">15.44</td>
<td id="8E6A5DC3BD16B6E84E2F28B9F9FD8A6E" box="[1581,1632,994,1052]" gridcol="17" gridrow="11" pageId="5" pageNumber="478">12.05</td>
<td id="8E6A5DC3BD16B6E84E7828B9F9308A6E" box="[1658,1709,994,1052]" gridcol="18" gridrow="11" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E7828AFF9308A78" bold="true" box="[1658,1709,1012,1034]" pageId="5" pageNumber="478">14.42</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA28B9F96D8A6E" box="[1736,1776,994,1052]" gridcol="19" gridrow="11" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F1728B9F8A08A6E" box="[1813,1853,994,1052]" gridcol="20" gridrow="11" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F6128B9F8168A6E" box="[1891,1931,994,1052]" gridcol="21" gridrow="11" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB228B9F8458A6E" box="[1968,2008,994,1052]" gridcol="22" gridrow="11" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02F76F8458A15" box="[162,2008,1069,1127]" gridrow="12" pageId="5" pageNumber="478" rowspan-12="1">
<th id="8E6A5DC3BD16B6E848A02F76FEA08A15" box="[162,317,1069,1127]" gridcol="0" gridrow="12" pageId="5" pageNumber="478">
12:
<emphasis id="41FFEAEFBD16491248C72F76FEB98A31" box="[197,292,1069,1091]" italics="true" pageId="5" pageNumber="478">Niphargus</emphasis>
sp. (Lebanon)
</th>
<td id="8E6A5DC3BD16B6E849542F76FE158A15" box="[342,392,1069,1127]" gridcol="1" gridrow="12" pageId="5" pageNumber="478">2.34</td>
<td id="8E6A5DC3BD16B6E849A12F76FE4B8A15" box="[419,470,1069,1127]" gridcol="2" gridrow="12" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E849F32F76FDB98A15" box="[497,548,1069,1127]" gridcol="3" gridrow="12" pageId="5" pageNumber="478">1.53</td>
<td id="8E6A5DC3BD16B6E84A3C2F76FDEC8A15" box="[574,625,1069,1127]" gridcol="4" gridrow="12" pageId="5" pageNumber="478">2.04</td>
<td id="8E6A5DC3BD16B6E84A8E2F76FD228A15" box="[652,703,1069,1127]" gridcol="5" gridrow="12" pageId="5" pageNumber="478">11.37</td>
<td id="8E6A5DC3BD16B6E84ADB2F76FC918A15" box="[729,780,1069,1127]" gridcol="6" gridrow="12" pageId="5" pageNumber="478">1.14</td>
<td id="8E6A5DC3BD16B6E84B242F76FCC48A15" box="[806,857,1069,1127]" gridcol="7" gridrow="12" pageId="5" pageNumber="478">2.97</td>
<td id="8E6A5DC3BD16B6E84B762F76FC3A8A15" box="[884,935,1069,1127]" gridcol="8" gridrow="12" pageId="5" pageNumber="478">1.66</td>
<td id="8E6A5DC3BD16B6E84BC32F76FC698A15" box="[961,1012,1069,1127]" gridcol="9" gridrow="12" pageId="5" pageNumber="478">1.66</td>
<td id="8E6A5DC3BD16B6E84C0D2F76FBDF8A15" box="[1039,1090,1069,1127]" gridcol="10" gridrow="12" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84C5E2F76FB128A15" box="[1116,1167,1069,1127]" gridcol="11" gridrow="12" pageId="5" pageNumber="478">0.25</td>
<td id="8E6A5DC3BD16B6E84CF52F76FAB78A15" box="[1271,1322,1069,1127]" gridcol="13" gridrow="12" pageId="5" pageNumber="478">15.82</td>
<td id="8E6A5DC3BD16B6E84D472F76FAE58A15" box="[1349,1400,1069,1127]" gridcol="14" gridrow="12" pageId="5" pageNumber="478">9.57</td>
<td id="8E6A5DC3BD16B6E84D902F76FA588A15" box="[1426,1477,1069,1127]" gridcol="15" gridrow="12" pageId="5" pageNumber="478">15.61</td>
<td id="8E6A5DC3BD16B6E84DDD2F76F98F8A15" box="[1503,1554,1069,1127]" gridcol="16" gridrow="12" pageId="5" pageNumber="478">16.16</td>
<td id="8E6A5DC3BD16B6E84E2F2F76F9FD8A15" box="[1581,1632,1069,1127]" gridcol="17" gridrow="12" pageId="5" pageNumber="478">9.82</td>
<td id="8E6A5DC3BD16B6E84E782F76F9308A15" box="[1658,1709,1069,1127]" gridcol="18" gridrow="12" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782F64F9308A27" bold="true" box="[1658,1709,1087,1109]" pageId="5" pageNumber="478">14.62</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2F76F96D8A15" box="[1736,1776,1069,1127]" gridcol="19" gridrow="12" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172F76F8A08A15" box="[1813,1853,1069,1127]" gridcol="20" gridrow="12" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612F76F8168A15" box="[1891,1931,1069,1127]" gridcol="21" gridrow="12" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22F76F8458A15" box="[1968,2008,1069,1127]" gridcol="22" gridrow="12" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02F2CF8458AFF" box="[162,2008,1143,1165]" gridrow="13" pageId="5" pageNumber="478" rowspan-13="1">
<th id="8E6A5DC3BD16B6E848A02F2CFEA08AFF" box="[162,317,1143,1165]" gridcol="0" gridrow="13" pageId="5" pageNumber="478">
13:
<emphasis id="41FFEAEFBD16491248C72F2CFEA18AFF" box="[197,316,1143,1165]" italics="true" pageId="5" pageNumber="478">N. nasrullahi</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542F2CFE158AFF" box="[342,392,1143,1165]" gridcol="1" gridrow="13" pageId="5" pageNumber="478">2.6</td>
<td id="8E6A5DC3BD16B6E849A12F2CFE4B8AFF" box="[419,470,1143,1165]" gridcol="2" gridrow="13" pageId="5" pageNumber="478">0.89</td>
<td id="8E6A5DC3BD16B6E849F32F2CFDB98AFF" box="[497,548,1143,1165]" gridcol="3" gridrow="13" pageId="5" pageNumber="478">1.53</td>
<td id="8E6A5DC3BD16B6E84A3C2F2CFDEC8AFF" box="[574,625,1143,1165]" gridcol="4" gridrow="13" pageId="5" pageNumber="478">1.78</td>
<td id="8E6A5DC3BD16B6E84A8E2F2CFD228AFF" box="[652,703,1143,1165]" gridcol="5" gridrow="13" pageId="5" pageNumber="478">11.36</td>
<td id="8E6A5DC3BD16B6E84ADB2F2CFC918AFF" box="[729,780,1143,1165]" gridcol="6" gridrow="13" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84B242F2CFCC48AFF" box="[806,857,1143,1165]" gridcol="7" gridrow="13" pageId="5" pageNumber="478">3.23</td>
<td id="8E6A5DC3BD16B6E84B762F2CFC3A8AFF" box="[884,935,1143,1165]" gridcol="8" gridrow="13" pageId="5" pageNumber="478">1.79</td>
<td id="8E6A5DC3BD16B6E84BC32F2CFC698AFF" box="[961,1012,1143,1165]" gridcol="9" gridrow="13" pageId="5" pageNumber="478">1.66</td>
<td id="8E6A5DC3BD16B6E84C0D2F2CFBDF8AFF" box="[1039,1090,1143,1165]" gridcol="10" gridrow="13" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84C5E2F2CFB128AFF" box="[1116,1167,1143,1165]" gridcol="11" gridrow="13" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84CA82F2CFB408AFF" box="[1194,1245,1143,1165]" gridcol="12" gridrow="13" pageId="5" pageNumber="478">1.28</td>
<td id="8E6A5DC3BD16B6E84D472F2CFAE58AFF" box="[1349,1400,1143,1165]" gridcol="14" gridrow="13" pageId="5" pageNumber="478">13.9</td>
<td id="8E6A5DC3BD16B6E84D902F2CFA588AFF" box="[1426,1477,1143,1165]" gridcol="15" gridrow="13" pageId="5" pageNumber="478">14.99</td>
<td id="8E6A5DC3BD16B6E84DDD2F2CF98F8AFF" box="[1503,1554,1143,1165]" gridcol="16" gridrow="13" pageId="5" pageNumber="478">16.94</td>
<td id="8E6A5DC3BD16B6E84E2F2F2CF9FD8AFF" box="[1581,1632,1143,1165]" gridcol="17" gridrow="13" pageId="5" pageNumber="478">13.22</td>
<td id="8E6A5DC3BD16B6E84E782F2CF9308AFF" box="[1658,1709,1143,1165]" gridcol="18" gridrow="13" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782F2CF93F8AFF" bold="true" box="[1658,1698,1143,1165]" pageId="5" pageNumber="478">8.98</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2F2CF96D8AFF" box="[1736,1776,1143,1165]" gridcol="19" gridrow="13" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172F2CF8A08AFF" box="[1813,1853,1143,1165]" gridcol="20" gridrow="13" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612F2CF8168AFF" box="[1891,1931,1143,1165]" gridcol="21" gridrow="13" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22F2CF8458AFF" box="[1968,2008,1143,1165]" gridcol="22" gridrow="13" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02FC6F8458AC6" box="[162,2008,1181,1204]" gridrow="14" pageId="5" pageNumber="478" rowspan-14="1">
<th id="8E6A5DC3BD16B6E848A02FC6FEA08AC6" box="[162,317,1181,1204]" gridcol="0" gridrow="14" pageId="5" pageNumber="478">
14:
<emphasis id="41FFEAEFBD16491248C72FC5FEB68AC6" box="[197,299,1182,1204]" italics="true" pageId="5" pageNumber="478">N. persicus</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542FC6FE158AC6" box="[342,392,1181,1204]" gridcol="1" gridrow="14" pageId="5" pageNumber="478">2.6</td>
<td id="8E6A5DC3BD16B6E849A12FC6FE4B8AC6" box="[419,470,1181,1204]" gridcol="2" gridrow="14" pageId="5" pageNumber="478">0.89</td>
<td id="8E6A5DC3BD16B6E849F32FC6FDB98AC6" box="[497,548,1181,1204]" gridcol="3" gridrow="14" pageId="5" pageNumber="478">1.53</td>
<td id="8E6A5DC3BD16B6E84A3C2FC6FDEC8AC6" box="[574,625,1181,1204]" gridcol="4" gridrow="14" pageId="5" pageNumber="478">2.82</td>
<td id="8E6A5DC3BD16B6E84A8E2FC6FD228AC6" box="[652,703,1181,1204]" gridcol="5" gridrow="14" pageId="5" pageNumber="478">11.67</td>
<td id="8E6A5DC3BD16B6E84ADB2FC6FC918AC6" box="[729,780,1181,1204]" gridcol="6" gridrow="14" pageId="5" pageNumber="478">0.25</td>
<td id="8E6A5DC3BD16B6E84B242FC6FCC48AC6" box="[806,857,1181,1204]" gridcol="7" gridrow="14" pageId="5" pageNumber="478">3.23</td>
<td id="8E6A5DC3BD16B6E84B762FC6FC3A8AC6" box="[884,935,1181,1204]" gridcol="8" gridrow="14" pageId="5" pageNumber="478">1.53</td>
<td id="8E6A5DC3BD16B6E84BC32FC6FC698AC6" box="[961,1012,1181,1204]" gridcol="9" gridrow="14" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84C0D2FC6FBDF8AC6" box="[1039,1090,1181,1204]" gridcol="10" gridrow="14" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84C5E2FC6FB128AC6" box="[1116,1167,1181,1204]" gridcol="11" gridrow="14" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84CA82FC6FB408AC6" box="[1194,1245,1181,1204]" gridcol="12" gridrow="14" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84CF52FC6FAB78AC6" box="[1271,1322,1181,1204]" gridcol="13" gridrow="14" pageId="5" pageNumber="478">1.53</td>
<td id="8E6A5DC3BD16B6E84D902FC6FA588AC6" box="[1426,1477,1181,1204]" gridcol="15" gridrow="14" pageId="5" pageNumber="478">14.87</td>
<td id="8E6A5DC3BD16B6E84DDD2FC6F98F8AC6" box="[1503,1554,1181,1204]" gridcol="16" gridrow="14" pageId="5" pageNumber="478">14.33</td>
<td id="8E6A5DC3BD16B6E84E2F2FC6F9FD8AC6" box="[1581,1632,1181,1204]" gridcol="17" gridrow="14" pageId="5" pageNumber="478">8.9</td>
<td id="8E6A5DC3BD16B6E84E782FC6F9308AC6" box="[1658,1709,1181,1204]" gridcol="18" gridrow="14" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782FC6F9308AC1" bold="true" box="[1658,1709,1181,1203]" pageId="5" pageNumber="478">12.98</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2FC6F96D8AC6" box="[1736,1776,1181,1204]" gridcol="19" gridrow="14" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172FC6F8A08AC6" box="[1813,1853,1181,1204]" gridcol="20" gridrow="14" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612FC6F8168AC6" box="[1891,1931,1181,1204]" gridcol="21" gridrow="14" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22FC6F8458AC6" box="[1968,2008,1181,1204]" gridcol="22" gridrow="14" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02F9FF8458AA8" box="[162,2008,1220,1242]" gridrow="15" pageId="5" pageNumber="478" rowspan-15="1">
<th id="8E6A5DC3BD16B6E848A02F9FFEA08AA8" box="[162,317,1220,1242]" gridcol="0" gridrow="15" pageId="5" pageNumber="478">
15:
<emphasis id="41FFEAEFBD16491248C72F9FFEBC8AA8" box="[197,289,1220,1242]" italics="true" pageId="5" pageNumber="478">N. sharifii</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542F9FFE158AA8" box="[342,392,1220,1242]" gridcol="1" gridrow="15" pageId="5" pageNumber="478">2.33</td>
<td id="8E6A5DC3BD16B6E849A12F9FFE4B8AA8" box="[419,470,1220,1242]" gridcol="2" gridrow="15" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E849F32F9FFDB98AA8" box="[497,548,1220,1242]" gridcol="3" gridrow="15" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E84A3C2F9FFDEC8AA8" box="[574,625,1220,1242]" gridcol="4" gridrow="15" pageId="5" pageNumber="478">1.78</td>
<td id="8E6A5DC3BD16B6E84A8E2F9FFD228AA8" box="[652,703,1220,1242]" gridcol="5" gridrow="15" pageId="5" pageNumber="478">11.05</td>
<td id="8E6A5DC3BD16B6E84ADB2F9FFC918AA8" box="[729,780,1220,1242]" gridcol="6" gridrow="15" pageId="5" pageNumber="478">1.14</td>
<td id="8E6A5DC3BD16B6E84B242F9FFCC48AA8" box="[806,857,1220,1242]" gridcol="7" gridrow="15" pageId="5" pageNumber="478">2.96</td>
<td id="8E6A5DC3BD16B6E84B762F9FFC3A8AA8" box="[884,935,1220,1242]" gridcol="8" gridrow="15" pageId="5" pageNumber="478">1.53</td>
<td id="8E6A5DC3BD16B6E84BC32F9FFC698AA8" box="[961,1012,1220,1242]" gridcol="9" gridrow="15" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84C0D2F9FFBDF8AA8" box="[1039,1090,1220,1242]" gridcol="10" gridrow="15" pageId="5" pageNumber="478">1.15</td>
<td id="8E6A5DC3BD16B6E84C5E2F9FFB128AA8" box="[1116,1167,1220,1242]" gridcol="11" gridrow="15" pageId="5" pageNumber="478">0.76</td>
<td id="8E6A5DC3BD16B6E84CA82F9FFB408AA8" box="[1194,1245,1220,1242]" gridcol="12" gridrow="15" pageId="5" pageNumber="478">0.76</td>
<td id="8E6A5DC3BD16B6E84CF52F9FFAB78AA8" box="[1271,1322,1220,1242]" gridcol="13" gridrow="15" pageId="5" pageNumber="478">0.76</td>
<td id="8E6A5DC3BD16B6E84D472F9FFAE58AA8" box="[1349,1400,1220,1242]" gridcol="14" gridrow="15" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84DDD2F9FF98F8AA8" box="[1503,1554,1220,1242]" gridcol="16" gridrow="15" pageId="5" pageNumber="478">15.71</td>
<td id="8E6A5DC3BD16B6E84E2F2F9FF9FD8AA8" box="[1581,1632,1220,1242]" gridcol="17" gridrow="15" pageId="5" pageNumber="478">13.7</td>
<td id="8E6A5DC3BD16B6E84E782F9FF9308AA8" box="[1658,1709,1220,1242]" gridcol="18" gridrow="15" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782F9FF9308AA8" bold="true" box="[1658,1709,1220,1242]" pageId="5" pageNumber="478">14.48</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2F9FF96D8AA8" box="[1736,1776,1220,1242]" gridcol="19" gridrow="15" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172F9FF8A08AA8" box="[1813,1853,1220,1242]" gridcol="20" gridrow="15" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612F9FF8168AA8" box="[1891,1931,1220,1242]" gridcol="21" gridrow="15" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22F9FF8458AA8" box="[1968,2008,1220,1242]" gridcol="22" gridrow="15" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02FB1F8458B56" box="[162,2008,1258,1316]" gridrow="16" pageId="5" pageNumber="478" rowspan-16="1">
<th id="8E6A5DC3BD16B6E848A02FB1FEA08B56" box="[162,317,1258,1316]" gridcol="0" gridrow="16" pageId="5" pageNumber="478">
16:
<emphasis id="41FFEAEFBD16491248C72FB1FF468B72" box="[197,219,1258,1280]" italics="true" pageId="5" pageNumber="478">N.</emphasis>
<emphasis id="41FFEAEFBD16491248A02E55FEB78B56" box="[162,298,1294,1316]" italics="true" pageId="5" pageNumber="478">sohrevardensis</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542FB1FE158B56" box="[342,392,1258,1316]" gridcol="1" gridrow="16" pageId="5" pageNumber="478">1.67</td>
<td id="8E6A5DC3BD16B6E849A12FB1FE4B8B56" box="[419,470,1258,1316]" gridcol="2" gridrow="16" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E849F32FB1FDB98B56" box="[497,548,1258,1316]" gridcol="3" gridrow="16" pageId="5" pageNumber="478">0.89</td>
<td id="8E6A5DC3BD16B6E84A3C2FB1FDEC8B56" box="[574,625,1258,1316]" gridcol="4" gridrow="16" pageId="5" pageNumber="478">2.56</td>
<td id="8E6A5DC3BD16B6E84A8E2FB1FD228B56" box="[652,703,1258,1316]" gridcol="5" gridrow="16" pageId="5" pageNumber="478">12.1</td>
<td id="8E6A5DC3BD16B6E84ADB2FB1FC918B56" box="[729,780,1258,1316]" gridcol="6" gridrow="16" pageId="5" pageNumber="478">1.01</td>
<td id="8E6A5DC3BD16B6E84B242FB1FCC48B56" box="[806,857,1258,1316]" gridcol="7" gridrow="16" pageId="5" pageNumber="478">2.56</td>
<td id="8E6A5DC3BD16B6E84B762FB1FC3A8B56" box="[884,935,1258,1316]" gridcol="8" gridrow="16" pageId="5" pageNumber="478">0.38</td>
<td id="8E6A5DC3BD16B6E84BC32FB1FC698B56" box="[961,1012,1258,1316]" gridcol="9" gridrow="16" pageId="5" pageNumber="478">0.51</td>
<td id="8E6A5DC3BD16B6E84C0D2FB1FBDF8B56" box="[1039,1090,1258,1316]" gridcol="10" gridrow="16" pageId="5" pageNumber="478">0.76</td>
<td id="8E6A5DC3BD16B6E84C5E2FB1FB128B56" box="[1116,1167,1258,1316]" gridcol="11" gridrow="16" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84CA82FB1FB408B56" box="[1194,1245,1258,1316]" gridcol="12" gridrow="16" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84CF52FB1FAB78B56" box="[1271,1322,1258,1316]" gridcol="13" gridrow="16" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84D472FB1FAE58B56" box="[1349,1400,1258,1316]" gridcol="14" gridrow="16" pageId="5" pageNumber="478">1.14</td>
<td id="8E6A5DC3BD16B6E84D902FB1FA588B56" box="[1426,1477,1258,1316]" gridcol="15" gridrow="16" pageId="5" pageNumber="478">1.15</td>
<td id="8E6A5DC3BD16B6E84E2F2FB1F9FD8B56" box="[1581,1632,1258,1316]" gridcol="17" gridrow="16" pageId="5" pageNumber="478">14.72</td>
<td id="8E6A5DC3BD16B6E84E782FB1F9308B56" box="[1658,1709,1258,1316]" gridcol="18" gridrow="16" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782FA7F9308B60" bold="true" box="[1658,1709,1276,1298]" pageId="5" pageNumber="478">16.42</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2FB1F96D8B56" box="[1736,1776,1258,1316]" gridcol="19" gridrow="16" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172FB1F8A08B56" box="[1813,1853,1258,1316]" gridcol="20" gridrow="16" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612FB1F8168B56" box="[1891,1931,1258,1316]" gridcol="21" gridrow="16" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22FB1F8458B56" box="[1968,2008,1258,1316]" gridcol="22" gridrow="16" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02E6EF8458B1D" box="[162,2008,1333,1391]" gridrow="17" pageId="5" pageNumber="478" rowspan-17="1">
<th id="8E6A5DC3BD16B6E848A02E6EFEA08B1D" box="[162,317,1333,1391]" gridcol="0" gridrow="17" pageId="5" pageNumber="478">
17:
<emphasis id="41FFEAEFBD16491248C72E6EFF468B39" box="[197,219,1333,1355]" italics="true" pageId="5" pageNumber="478">N.</emphasis>
<emphasis id="41FFEAEFBD16491248A02E02FE988B1D" box="[162,261,1369,1391]" italics="true" pageId="5" pageNumber="478">urumiensis</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542E6EFE158B1D" box="[342,392,1333,1391]" gridcol="1" gridrow="17" pageId="5" pageNumber="478">1.15</td>
<td id="8E6A5DC3BD16B6E849A12E6EFE4B8B1D" box="[419,470,1333,1391]" gridcol="2" gridrow="17" pageId="5" pageNumber="478">2.33</td>
<td id="8E6A5DC3BD16B6E849F32E6EFDB98B1D" box="[497,548,1333,1391]" gridcol="3" gridrow="17" pageId="5" pageNumber="478">2.2</td>
<td id="8E6A5DC3BD16B6E84A3C2E6EFDEC8B1D" box="[574,625,1333,1391]" gridcol="4" gridrow="17" pageId="5" pageNumber="478">3.79</td>
<td id="8E6A5DC3BD16B6E84A8E2E6EFD228B1D" box="[652,703,1333,1391]" gridcol="5" gridrow="17" pageId="5" pageNumber="478">11.61</td>
<td id="8E6A5DC3BD16B6E84ADB2E6EFC918B1D" box="[729,780,1333,1391]" gridcol="6" gridrow="17" pageId="5" pageNumber="478">2.59</td>
<td id="8E6A5DC3BD16B6E84B242E6EFCC48B1D" box="[806,857,1333,1391]" gridcol="7" gridrow="17" pageId="5" pageNumber="478">2.06</td>
<td id="8E6A5DC3BD16B6E84B762E6EFC3A8B1D" box="[884,935,1333,1391]" gridcol="8" gridrow="17" pageId="5" pageNumber="478">2.59</td>
<td id="8E6A5DC3BD16B6E84BC32E6EFC698B1D" box="[961,1012,1333,1391]" gridcol="9" gridrow="17" pageId="5" pageNumber="478">2.6</td>
<td id="8E6A5DC3BD16B6E84C0D2E6EFBDF8B1D" box="[1039,1090,1333,1391]" gridcol="10" gridrow="17" pageId="5" pageNumber="478">2.46</td>
<td id="8E6A5DC3BD16B6E84C5E2E6EFB128B1D" box="[1116,1167,1333,1391]" gridcol="11" gridrow="17" pageId="5" pageNumber="478">2.46</td>
<td id="8E6A5DC3BD16B6E84CA82E6EFB408B1D" box="[1194,1245,1333,1391]" gridcol="12" gridrow="17" pageId="5" pageNumber="478">2.47</td>
<td id="8E6A5DC3BD16B6E84CF52E6EFAB78B1D" box="[1271,1322,1333,1391]" gridcol="13" gridrow="17" pageId="5" pageNumber="478">2.73</td>
<td id="8E6A5DC3BD16B6E84D472E6EFAE58B1D" box="[1349,1400,1333,1391]" gridcol="14" gridrow="17" pageId="5" pageNumber="478">2.73</td>
<td id="8E6A5DC3BD16B6E84D902E6EFA588B1D" box="[1426,1477,1333,1391]" gridcol="15" gridrow="17" pageId="5" pageNumber="478">2.2</td>
<td id="8E6A5DC3BD16B6E84DDD2E6EF98F8B1D" box="[1503,1554,1333,1391]" gridcol="16" gridrow="17" pageId="5" pageNumber="478">2.46</td>
<td id="8E6A5DC3BD16B6E84E782E6EF9308B1D" box="[1658,1709,1333,1391]" gridcol="18" gridrow="17" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782E1CF9308B2F" bold="true" box="[1658,1709,1351,1373]" pageId="5" pageNumber="478">13.47</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2E6EF96D8B1D" box="[1736,1776,1333,1391]" gridcol="19" gridrow="17" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172E6EF8A08B1D" box="[1813,1853,1333,1391]" gridcol="20" gridrow="17" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612E6EF8168B1D" box="[1891,1931,1333,1391]" gridcol="21" gridrow="17" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22E6EF8458B1D" box="[1968,2008,1333,1391]" gridcol="22" gridrow="17" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02E24F8458BE7" box="[162,2008,1407,1429]" gridrow="18" pageId="5" pageNumber="478" rowspan-18="1">
<th id="8E6A5DC3BD16B6E848A02E24FEA08BE7" box="[162,317,1407,1429]" gridcol="0" gridrow="18" pageId="5" pageNumber="478">
18:
<emphasis id="41FFEAEFBD16491248C72E24FEA18BE7" box="[197,316,1407,1429]" italics="true" pageId="5" pageNumber="478">N. yasujensis</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542E24FE158BE7" box="[342,392,1407,1429]" gridcol="1" gridrow="18" pageId="5" pageNumber="478">3.27</td>
<td id="8E6A5DC3BD16B6E849A12E24FE4B8BE7" box="[419,470,1407,1429]" gridcol="2" gridrow="18" pageId="5" pageNumber="478">1.66</td>
<td id="8E6A5DC3BD16B6E849F32E24FDB98BE7" box="[497,548,1407,1429]" gridcol="3" gridrow="18" pageId="5" pageNumber="478">2.31</td>
<td id="8E6A5DC3BD16B6E84A3C2E24FDEC8BE7" box="[574,625,1407,1429]" gridcol="4" gridrow="18" pageId="5" pageNumber="478">1.14</td>
<td id="8E6A5DC3BD16B6E84A8E2E24FD228BE7" box="[652,703,1407,1429]" gridcol="5" gridrow="18" pageId="5" pageNumber="478">11.21</td>
<td id="8E6A5DC3BD16B6E84ADB2E24FC918BE7" box="[729,780,1407,1429]" gridcol="6" gridrow="18" pageId="5" pageNumber="478">2.18</td>
<td id="8E6A5DC3BD16B6E84B242E24FCC48BE7" box="[806,857,1407,1429]" gridcol="7" gridrow="18" pageId="5" pageNumber="478">4.03</td>
<td id="8E6A5DC3BD16B6E84B762E24FC3A8BE7" box="[884,935,1407,1429]" gridcol="8" gridrow="18" pageId="5" pageNumber="478">2.57</td>
<td id="8E6A5DC3BD16B6E84BC32E24FC698BE7" box="[961,1012,1407,1429]" gridcol="9" gridrow="18" pageId="5" pageNumber="478">2.44</td>
<td id="8E6A5DC3BD16B6E84C0D2E24FBDF8BE7" box="[1039,1090,1407,1429]" gridcol="10" gridrow="18" pageId="5" pageNumber="478">2.18</td>
<td id="8E6A5DC3BD16B6E84C5E2E24FB128BE7" box="[1116,1167,1407,1429]" gridcol="11" gridrow="18" pageId="5" pageNumber="478">1.79</td>
<td id="8E6A5DC3BD16B6E84CA82E24FB408BE7" box="[1194,1245,1407,1429]" gridcol="12" gridrow="18" pageId="5" pageNumber="478">1.79</td>
<td id="8E6A5DC3BD16B6E84CF52E24FAB78BE7" box="[1271,1322,1407,1429]" gridcol="13" gridrow="18" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84D472E24FAE58BE7" box="[1349,1400,1407,1429]" gridcol="14" gridrow="18" pageId="5" pageNumber="478">2.31</td>
<td id="8E6A5DC3BD16B6E84D902E24FA588BE7" box="[1426,1477,1407,1429]" gridcol="15" gridrow="18" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84DDD2E24F98F8BE7" box="[1503,1554,1407,1429]" gridcol="16" gridrow="18" pageId="5" pageNumber="478">2.18</td>
<td id="8E6A5DC3BD16B6E84E2F2E24F9FD8BE7" box="[1581,1632,1407,1429]" gridcol="17" gridrow="18" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E2F2E24F9C88BE7" bold="true" box="[1581,1621,1407,1429]" pageId="5" pageNumber="478">3.26</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84ECA2E24F96D8BE7" box="[1736,1776,1407,1429]" gridcol="19" gridrow="18" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F172E24F8A08BE7" box="[1813,1853,1407,1429]" gridcol="20" gridrow="18" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612E24F8168BE7" box="[1891,1931,1407,1429]" gridcol="21" gridrow="18" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22E24F8458BE7" box="[1968,2008,1407,1429]" gridcol="22" gridrow="18" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02EFEF8458BCE" box="[162,2008,1445,1468]" gridrow="19" pageId="5" pageNumber="478" rowspan-19="1">
<th id="8E6A5DC3BD16B6E848A02EFEFEA08BCE" box="[162,317,1445,1468]" gridcol="0" gridrow="19" pageId="5" pageNumber="478">
19:
<emphasis id="41FFEAEFBD16491248C72EFDFE808BCE" box="[197,285,1446,1468]" italics="true" pageId="5" pageNumber="478">N. hakani</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542EFEFE158BCE" box="[342,392,1445,1468]" gridcol="1" gridrow="19" pageId="5" pageNumber="478">1.54</td>
<td id="8E6A5DC3BD16B6E849A12EFEFE4B8BCE" box="[419,470,1445,1468]" gridcol="2" gridrow="19" pageId="5" pageNumber="478">2.43</td>
<td id="8E6A5DC3BD16B6E849F32EFEFDB98BCE" box="[497,548,1445,1468]" gridcol="3" gridrow="19" pageId="5" pageNumber="478">2.56</td>
<td id="8E6A5DC3BD16B6E84A3C2EFEFDEC8BCE" box="[574,625,1445,1468]" gridcol="4" gridrow="19" pageId="5" pageNumber="478">3.88</td>
<td id="8E6A5DC3BD16B6E84A8E2EFEFD228BCE" box="[652,703,1445,1468]" gridcol="5" gridrow="19" pageId="5" pageNumber="478">11.76</td>
<td id="8E6A5DC3BD16B6E84ADB2EFEFC918BCE" box="[729,780,1445,1468]" gridcol="6" gridrow="19" pageId="5" pageNumber="478">2.95</td>
<td id="8E6A5DC3BD16B6E84B242EFEFCC48BCE" box="[806,857,1445,1468]" gridcol="7" gridrow="19" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84B762EFEFC3A8BCE" box="[884,935,1445,1468]" gridcol="8" gridrow="19" pageId="5" pageNumber="478">2.56</td>
<td id="8E6A5DC3BD16B6E84BC32EFEFC698BCE" box="[961,1012,1445,1468]" gridcol="9" gridrow="19" pageId="5" pageNumber="478">2.43</td>
<td id="8E6A5DC3BD16B6E84C0D2EFEFBDF8BCE" box="[1039,1090,1445,1468]" gridcol="10" gridrow="19" pageId="5" pageNumber="478">2.3</td>
<td id="8E6A5DC3BD16B6E84C5E2EFEFB128BCE" box="[1116,1167,1445,1468]" gridcol="11" gridrow="19" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84CA82EFEFB408BCE" box="[1194,1245,1445,1468]" gridcol="12" gridrow="19" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84CF52EFEFAB78BCE" box="[1271,1322,1445,1468]" gridcol="13" gridrow="19" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84D472EFEFAE58BCE" box="[1349,1400,1445,1468]" gridcol="14" gridrow="19" pageId="5" pageNumber="478">3.09</td>
<td id="8E6A5DC3BD16B6E84D902EFEFA588BCE" box="[1426,1477,1445,1468]" gridcol="15" gridrow="19" pageId="5" pageNumber="478">2.56</td>
<td id="8E6A5DC3BD16B6E84DDD2EFEF98F8BCE" box="[1503,1554,1445,1468]" gridcol="16" gridrow="19" pageId="5" pageNumber="478">2.17</td>
<td id="8E6A5DC3BD16B6E84E2F2EFEF9FD8BCE" box="[1581,1632,1445,1468]" gridcol="17" gridrow="19" pageId="5" pageNumber="478">2.2</td>
<td id="8E6A5DC3BD16B6E84E782EFEF9308BCE" box="[1658,1709,1445,1468]" gridcol="18" gridrow="19" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124E782EFEF93F8BC9" bold="true" box="[1658,1698,1445,1467]" pageId="5" pageNumber="478">3.35</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84F172EFEF8A08BCE" box="[1813,1853,1445,1468]" gridcol="20" gridrow="19" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84F612EFEF8168BCE" box="[1891,1931,1445,1468]" gridcol="21" gridrow="19" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22EFEF8458BCE" box="[1968,2008,1445,1468]" gridcol="22" gridrow="19" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02E97F8458874" box="[162,2008,1484,1542]" gridrow="20" pageId="5" pageNumber="478" rowspan-20="1">
<th id="8E6A5DC3BD16B6E848A02E97FEA08874" box="[162,317,1484,1542]" gridcol="0" gridrow="20" pageId="5" pageNumber="478">
20:
<emphasis id="41FFEAEFBD16491248C72E97FF468B90" box="[197,219,1484,1506]" italics="true" pageId="5" pageNumber="478">N.</emphasis>
<emphasis id="41FFEAEFBD16491248A02EABFEB48874" box="[162,297,1520,1542]" italics="true" pageId="5" pageNumber="478">hegmatanensis</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542E97FE158874" box="[342,392,1484,1542]" gridcol="1" gridrow="20" pageId="5" pageNumber="478">2.2</td>
<td id="8E6A5DC3BD16B6E849A12E97FE4B8874" box="[419,470,1484,1542]" gridcol="2" gridrow="20" pageId="5" pageNumber="478">0.25</td>
<td id="8E6A5DC3BD16B6E849F32E97FDB98874" box="[497,548,1484,1542]" gridcol="3" gridrow="20" pageId="5" pageNumber="478">1.15</td>
<td id="8E6A5DC3BD16B6E84A3C2E97FDEC8874" box="[574,625,1484,1542]" gridcol="4" gridrow="20" pageId="5" pageNumber="478">1.91</td>
<td id="8E6A5DC3BD16B6E84A8E2E97FD228874" box="[652,703,1484,1542]" gridcol="5" gridrow="20" pageId="5" pageNumber="478">11.51</td>
<td id="8E6A5DC3BD16B6E84ADB2E97FC918874" box="[729,780,1484,1542]" gridcol="6" gridrow="20" pageId="5" pageNumber="478">0.76</td>
<td id="8E6A5DC3BD16B6E84B242E97FCC48874" box="[806,857,1484,1542]" gridcol="7" gridrow="20" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84B762E97FC3A8874" box="[884,935,1484,1542]" gridcol="8" gridrow="20" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84BC32E97FC698874" box="[961,1012,1484,1542]" gridcol="9" gridrow="20" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84C0D2E97FBDF8874" box="[1039,1090,1484,1542]" gridcol="10" gridrow="20" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E84C5E2E97FB128874" box="[1116,1167,1484,1542]" gridcol="11" gridrow="20" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E84CA82E97FB408874" box="[1194,1245,1484,1542]" gridcol="12" gridrow="20" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E84CF52E97FAB78874" box="[1271,1322,1484,1542]" gridcol="13" gridrow="20" pageId="5" pageNumber="478">0.89</td>
<td id="8E6A5DC3BD16B6E84D472E97FAE58874" box="[1349,1400,1484,1542]" gridcol="14" gridrow="20" pageId="5" pageNumber="478">0.89</td>
<td id="8E6A5DC3BD16B6E84D902E97FA588874" box="[1426,1477,1484,1542]" gridcol="15" gridrow="20" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E84DDD2E97F98F8874" box="[1503,1554,1484,1542]" gridcol="16" gridrow="20" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E84E2F2E97F9FD8874" box="[1581,1632,1484,1542]" gridcol="17" gridrow="20" pageId="5" pageNumber="478">2.33</td>
<td id="8E6A5DC3BD16B6E84E782E97F9308874" box="[1658,1709,1484,1542]" gridcol="18" gridrow="20" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84ECA2E97F96D8874" box="[1736,1776,1484,1542]" gridcol="19" gridrow="20" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124ECA2E85F96D8B86" bold="true" box="[1736,1776,1502,1524]" pageId="5" pageNumber="478">2.43</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84F612E97F8168874" box="[1891,1931,1484,1542]" gridcol="21" gridrow="20" pageId="5" pageNumber="478">*</td>
<td id="8E6A5DC3BD16B6E84FB22E97F8458874" box="[1968,2008,1484,1542]" gridcol="22" gridrow="20" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02D4CF8458823" box="[162,2008,1559,1617]" gridrow="21" pageId="5" pageNumber="478" rowspan-21="1">
<th id="8E6A5DC3BD16B6E848A02D4CFEA08823" box="[162,317,1559,1617]" gridcol="0" gridrow="21" pageId="5" pageNumber="478">
21:
<emphasis id="41FFEAEFBD16491248C72D4CFF46885F" box="[197,219,1559,1581]" italics="true" pageId="5" pageNumber="478">N.</emphasis>
<emphasis id="41FFEAEFBD16491248A02D60FE8A8823" box="[162,279,1595,1617]" italics="true" pageId="5" pageNumber="478">kermanshahi</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542D4CFE158823" box="[342,392,1559,1617]" gridcol="1" gridrow="21" pageId="5" pageNumber="478">2.07</td>
<td id="8E6A5DC3BD16B6E849A12D4CFE4B8823" box="[419,470,1559,1617]" gridcol="2" gridrow="21" pageId="5" pageNumber="478">0.89</td>
<td id="8E6A5DC3BD16B6E849F32D4CFDB98823" box="[497,548,1559,1617]" gridcol="3" gridrow="21" pageId="5" pageNumber="478">0.51</td>
<td id="8E6A5DC3BD16B6E84A3C2D4CFDEC8823" box="[574,625,1559,1617]" gridcol="4" gridrow="21" pageId="5" pageNumber="478">2.43</td>
<td id="8E6A5DC3BD16B6E84A8E2D4CFD228823" box="[652,703,1559,1617]" gridcol="5" gridrow="21" pageId="5" pageNumber="478">11.67</td>
<td id="8E6A5DC3BD16B6E84ADB2D4CFC918823" box="[729,780,1559,1617]" gridcol="6" gridrow="21" pageId="5" pageNumber="478">1.14</td>
<td id="8E6A5DC3BD16B6E84B242D4CFCC48823" box="[806,857,1559,1617]" gridcol="7" gridrow="21" pageId="5" pageNumber="478">2.96</td>
<td id="8E6A5DC3BD16B6E84B762D4CFC3A8823" box="[884,935,1559,1617]" gridcol="8" gridrow="21" pageId="5" pageNumber="478">1.01</td>
<td id="8E6A5DC3BD16B6E84BC32D4CFC698823" box="[961,1012,1559,1617]" gridcol="9" gridrow="21" pageId="5" pageNumber="478">0.89</td>
<td id="8E6A5DC3BD16B6E84C0D2D4CFBDF8823" box="[1039,1090,1559,1617]" gridcol="10" gridrow="21" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E84C5E2D4CFB128823" box="[1116,1167,1559,1617]" gridcol="11" gridrow="21" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E84CA82D4CFB408823" box="[1194,1245,1559,1617]" gridcol="12" gridrow="21" pageId="5" pageNumber="478">1.28</td>
<td id="8E6A5DC3BD16B6E84CF52D4CFAB78823" box="[1271,1322,1559,1617]" gridcol="13" gridrow="21" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84D472D4CFAE58823" box="[1349,1400,1559,1617]" gridcol="14" gridrow="21" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84D902D4CFA588823" box="[1426,1477,1559,1617]" gridcol="15" gridrow="21" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E84DDD2D4CF98F8823" box="[1503,1554,1559,1617]" gridcol="16" gridrow="21" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E84E2F2D4CF9FD8823" box="[1581,1632,1559,1617]" gridcol="17" gridrow="21" pageId="5" pageNumber="478">2.46</td>
<td id="8E6A5DC3BD16B6E84E782D4CF9308823" box="[1658,1709,1559,1617]" gridcol="18" gridrow="21" pageId="5" pageNumber="478">2.05</td>
<td id="8E6A5DC3BD16B6E84ECA2D4CF96D8823" box="[1736,1776,1559,1617]" gridcol="19" gridrow="21" pageId="5" pageNumber="478">2.3</td>
<td id="8E6A5DC3BD16B6E84F172D4CF8A08823" box="[1813,1853,1559,1617]" gridcol="20" gridrow="21" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124F172D73F8A0884C" bold="true" box="[1813,1853,1576,1598]" pageId="5" pageNumber="478">0.89</emphasis>
</td>
<td id="8E6A5DC3BD16B6E84FB22D4CF8458823" box="[1968,2008,1559,1617]" gridcol="22" gridrow="21" pageId="5" pageNumber="478">*</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02D3AF84588E9" box="[162,2008,1633,1691]" gridrow="22" pageId="5" pageNumber="478" rowspan-22="1">
<th id="8E6A5DC3BD16B6E848A02D3AFEA088E9" box="[162,317,1633,1691]" gridcol="0" gridrow="22" pageId="5" pageNumber="478">
22:
<emphasis id="41FFEAEFBD16491248C72D3AFF468805" box="[197,219,1633,1655]" italics="true" pageId="5" pageNumber="478">N.</emphasis>
<emphasis id="41FFEAEFBD16491248A02DDEFE8488E9" box="[162,281,1669,1691]" italics="true" pageId="5" pageNumber="478">lorestanensis</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542D3AFE1588E9" box="[342,392,1633,1691]" gridcol="1" gridrow="22" pageId="5" pageNumber="478">2.33</td>
<td id="8E6A5DC3BD16B6E849A12D3AFE4B88E9" box="[419,470,1633,1691]" gridcol="2" gridrow="22" pageId="5" pageNumber="478">0.38</td>
<td id="8E6A5DC3BD16B6E849F32D3AFDB988E9" box="[497,548,1633,1691]" gridcol="3" gridrow="22" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84A3C2D3AFDEC88E9" box="[574,625,1633,1691]" gridcol="4" gridrow="22" pageId="5" pageNumber="478">2.56</td>
<td id="8E6A5DC3BD16B6E84A8E2D3AFD2288E9" box="[652,703,1633,1691]" gridcol="5" gridrow="22" pageId="5" pageNumber="478">11.36</td>
<td id="8E6A5DC3BD16B6E84ADB2D3AFC9188E9" box="[729,780,1633,1691]" gridcol="6" gridrow="22" pageId="5" pageNumber="478">0.38</td>
<td id="8E6A5DC3BD16B6E84B242D3AFCC488E9" box="[806,857,1633,1691]" gridcol="7" gridrow="22" pageId="5" pageNumber="478">2.7</td>
<td id="8E6A5DC3BD16B6E84B762D3AFC3A88E9" box="[884,935,1633,1691]" gridcol="8" gridrow="22" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84BC32D3AFC6988E9" box="[961,1012,1633,1691]" gridcol="9" gridrow="22" pageId="5" pageNumber="478">1.15</td>
<td id="8E6A5DC3BD16B6E84C0D2D3AFBDF88E9" box="[1039,1090,1633,1691]" gridcol="10" gridrow="22" pageId="5" pageNumber="478">1.15</td>
<td id="8E6A5DC3BD16B6E84C5E2D3AFB1288E9" box="[1116,1167,1633,1691]" gridcol="11" gridrow="22" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E84CA82D3AFB4088E9" box="[1194,1245,1633,1691]" gridcol="12" gridrow="22" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E84CF52D3AFAB788E9" box="[1271,1322,1633,1691]" gridcol="13" gridrow="22" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84D472D3AFAE588E9" box="[1349,1400,1633,1691]" gridcol="14" gridrow="22" pageId="5" pageNumber="478">0.5</td>
<td id="8E6A5DC3BD16B6E84D902D3AFA5888E9" box="[1426,1477,1633,1691]" gridcol="15" gridrow="22" pageId="5" pageNumber="478">1.02</td>
<td id="8E6A5DC3BD16B6E84DDD2D3AF98F88E9" box="[1503,1554,1633,1691]" gridcol="16" gridrow="22" pageId="5" pageNumber="478">0.89</td>
<td id="8E6A5DC3BD16B6E84E2F2D3AF9FD88E9" box="[1581,1632,1633,1691]" gridcol="17" gridrow="22" pageId="5" pageNumber="478">2.46</td>
<td id="8E6A5DC3BD16B6E84E782D3AF93088E9" box="[1658,1709,1633,1691]" gridcol="18" gridrow="22" pageId="5" pageNumber="478">2.05</td>
<td id="8E6A5DC3BD16B6E84ECA2D3AF96D88E9" box="[1736,1776,1633,1691]" gridcol="19" gridrow="22" pageId="5" pageNumber="478">2.83</td>
<td id="8E6A5DC3BD16B6E84F172D3AF8A088E9" box="[1813,1853,1633,1691]" gridcol="20" gridrow="22" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E84F612D3AF81688E9" box="[1891,1931,1633,1691]" gridcol="21" gridrow="22" pageId="5" pageNumber="478">
<emphasis id="41FFEAEFBD1649124F612D28F81688FB" bold="true" box="[1891,1931,1651,1673]" pageId="5" pageNumber="478">1.02</emphasis>
</td>
</tr>
<tr id="CDBB34BFBD16B6E848A02DF0F84588B3" box="[162,2008,1707,1729]" gridrow="23" pageId="5" pageNumber="478">
<th id="8E6A5DC3BD16B6E848A02DF0FEA088B3" box="[162,317,1707,1729]" gridcol="0" gridrow="23" pageId="5" pageNumber="478">
23:
<emphasis id="41FFEAEFBD16491248C72DF0FE9488B3" box="[197,265,1707,1729]" italics="true" pageId="5" pageNumber="478">N. sarii</emphasis>
</th>
<td id="8E6A5DC3BD16B6E849542DF0FE1588B3" box="[342,392,1707,1729]" gridcol="1" gridrow="23" pageId="5" pageNumber="478">2.33</td>
<td id="8E6A5DC3BD16B6E849A12DF0FE4B88B3" box="[419,470,1707,1729]" gridcol="2" gridrow="23" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E849F32DF0FDB988B3" box="[497,548,1707,1729]" gridcol="3" gridrow="23" pageId="5" pageNumber="478">1.27</td>
<td id="8E6A5DC3BD16B6E84A3C2DF0FDEC88B3" box="[574,625,1707,1729]" gridcol="4" gridrow="23" pageId="5" pageNumber="478">2.96</td>
<td id="8E6A5DC3BD16B6E84A8E2DF0FD2288B3" box="[652,703,1707,1729]" gridcol="5" gridrow="23" pageId="5" pageNumber="478">12.28</td>
<td id="8E6A5DC3BD16B6E84ADB2DF0FC9188B3" box="[729,780,1707,1729]" gridcol="6" gridrow="23" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84B242DF0FCC488B3" box="[806,857,1707,1729]" gridcol="7" gridrow="23" pageId="5" pageNumber="478">3.23</td>
<td id="8E6A5DC3BD16B6E84B762DF0FC3A88B3" box="[884,935,1707,1729]" gridcol="8" gridrow="23" pageId="5" pageNumber="478">0.76</td>
<td id="8E6A5DC3BD16B6E84BC32DF0FC6988B3" box="[961,1012,1707,1729]" gridcol="9" gridrow="23" pageId="5" pageNumber="478">1.14</td>
<td id="8E6A5DC3BD16B6E84C0D2DF0FBDF88B3" box="[1039,1090,1707,1729]" gridcol="10" gridrow="23" pageId="5" pageNumber="478">1.14</td>
<td id="8E6A5DC3BD16B6E84C5E2DF0FB1288B3" box="[1116,1167,1707,1729]" gridcol="11" gridrow="23" pageId="5" pageNumber="478">1.79</td>
<td id="8E6A5DC3BD16B6E84CA82DF0FB4088B3" box="[1194,1245,1707,1729]" gridcol="12" gridrow="23" pageId="5" pageNumber="478">1.79</td>
<td id="8E6A5DC3BD16B6E84CF52DF0FAB788B3" box="[1271,1322,1707,1729]" gridcol="13" gridrow="23" pageId="5" pageNumber="478">1.79</td>
<td id="8E6A5DC3BD16B6E84D472DF0FAE588B3" box="[1349,1400,1707,1729]" gridcol="14" gridrow="23" pageId="5" pageNumber="478">1.53</td>
<td id="8E6A5DC3BD16B6E84D902DF0FA5888B3" box="[1426,1477,1707,1729]" gridcol="15" gridrow="23" pageId="5" pageNumber="478">1.53</td>
<td id="8E6A5DC3BD16B6E84DDD2DF0F98F88B3" box="[1503,1554,1707,1729]" gridcol="16" gridrow="23" pageId="5" pageNumber="478">0.63</td>
<td id="8E6A5DC3BD16B6E84E2F2DF0F9FD88B3" box="[1581,1632,1707,1729]" gridcol="17" gridrow="23" pageId="5" pageNumber="478">2.73</td>
<td id="8E6A5DC3BD16B6E84E782DF0F93088B3" box="[1658,1709,1707,1729]" gridcol="18" gridrow="23" pageId="5" pageNumber="478">2.57</td>
<td id="8E6A5DC3BD16B6E84ECA2DF0F96D88B3" box="[1736,1776,1707,1729]" gridcol="19" gridrow="23" pageId="5" pageNumber="478">2.82</td>
<td id="8E6A5DC3BD16B6E84F172DF0F8A088B3" box="[1813,1853,1707,1729]" gridcol="20" gridrow="23" pageId="5" pageNumber="478">1.4</td>
<td id="8E6A5DC3BD16B6E84F612DF0F81688B3" box="[1891,1931,1707,1729]" gridcol="21" gridrow="23" pageId="5" pageNumber="478">1.01</td>
<td id="8E6A5DC3BD16B6E84FB22DF0F84588B3" box="[1968,2008,1707,1729]" gridcol="22" gridrow="23" pageId="5" pageNumber="478">1.27</td>
</tr>
</table>
</paragraph>
<paragraph id="733436FDBD15491148C52BCCFE5C8E8B" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="479">
Antennae II. Length ratio antenna I: II as 1: 0.74. Peduncular article 4 longer than article 5, with six and seven groups of simple setae, respectively; flagellum with eight articles. Length of flagellum: length of peduncle article 4 + 5 as 0.59: 1 (
<figureCitation id="EBB02A78BD154911493C2B84FE298E8B" box="[318,436,223,249]" captionStart="FIGURE 4" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[222,1365,191,1909]" captionTargetId="figure-26@8.[212,1375,181,1919]" captionTargetPageId="8" captionText="FIGURE 4. Niphargus qazvinensis sp. nov., Samaq Abad Spring, male 8 mm (holotype). A, Head; B, Antenna I; C, Antenna II; D, Mandibular palp; E-F, Maxilla I; G, Right mandible; H, Left mandible. Scale bars: 1= 0.25 mm (GH); 2= 0.5 mm (A, DF); 3= 1mm (BC)." figureDoi="http://doi.org/10.5281/zenodo.15561700" httpUri="https://zenodo.org/record/15561700/files/figure.png" pageId="6" pageNumber="479">Figure 4C</figureCitation>