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;
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+ textfigs 15; accepted after
<treatmentCitation id="0AC41098F9429D5CFBE027EA88F5AF91" author="Liao, Y." box="[1044,1246,975,1000]" page="264" pageId="16" pageNumber="17" year="1997">
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+ textfig. 157ae.
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<mods:identifier id="4F2A6421E6EDD70FBE840A519F91DE85" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0425C3BAD16D5E403B177B04476216FB" type="Zenodo-Dep">14703528</mods:identifier>
<mods:identifier id="9D06F078CCE0B28BF9AA4BD22FC0D921" type="ZooBank">96194340-054D-49C0-BD89-D47899BCAF14</mods:identifier>
</mods:mods>
<treatment id="03CC879FF9449D57FF63217B8936AB73" ID-DOI="http://doi.org/10.5281/zenodo.14704968" ID-Zenodo-Dep="14704968" LSID="urn:lsid:plazi:treatment:03CC879FF9449D57FF63217B8936AB73" httpUri="http://treatment.plazi.org/id/03CC879FF9449D57FF63217B8936AB73" lastPageId="27" lastPageNumber="28" pageId="22" pageNumber="23">
<subSubSection id="C37F6502F9449D5AFF63217B8DC8A92D" box="[151,483,1344,1371]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BDA3689F9449D5AFF63217B8DC8A92D" blockId="22.[151,705,1344,1443]" box="[151,483,1344,1371]" pageId="22" pageNumber="23">
<heading id="D09281E5F9449D5AFF63217B8DC8A92D" bold="true" box="[151,483,1344,1371]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B911EA9BF9449D5AFF63217B8DC8A92D" bold="true" box="[151,483,1344,1371]" pageId="22" pageNumber="23">
<taxonomicName id="4C654D0AF9449D5AFF63217B8D55A92D" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[151,382,1344,1371]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="22" pageNumber="23" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF9449D5AFF63217B8D55A92D" bold="true" box="[151,382,1344,1371]" italics="true" pageId="22" pageNumber="23">Patinapta deformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257E0F9449D5AFE71217B8DC8A92D" box="[389,483,1345,1371]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5AFF63215C8EEAA909" box="[151,705,1381,1408]" pageId="22" pageNumber="23" type="vernacular_names">
<paragraph id="8BDA3689F9449D5AFF63215C8EEAA909" blockId="22.[151,705,1344,1443]" box="[151,705,1381,1408]" pageId="22" pageNumber="23">[New Japanese name: Yoron-himo-ikari-namako]</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5AFF6321B38DA4A98B" pageId="22" pageNumber="23" type="description">
<paragraph id="8BDA3689F9449D5AFF6321B38DE0A9D5" blockId="22.[151,705,1344,1443]" box="[151,459,1417,1443]" pageId="22" pageNumber="23">
(
<figureCitation id="135E2A0CF9449D5AFF6B21B38D0AA9D5" box="[159,289,1417,1443]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStartId-0="23.[151,250,1446,1470]" captionStartId-1="25.[151,250,1699,1723]" captionStartId-2="26.[151,250,516,540]" captionTargetBox-0="[151,1436,309,1423]" captionTargetBox-1="[151,1436,181,1667]" captionTargetBox-2="[164,1423,188,485]" captionTargetId-0="figure-69@23.[151,1436,309,1423]" captionTargetId-1="figure-20@25.[151,1436,181,1675]" captionTargetId-2="figure-29@26.[151,1435,181,493]" captionTargetPageId-0="23" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="FIGURE 12. External view of holotype specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (specimen WMNH-2015-INV-106): A, lateral view, anterior left, dorsal top; B, oral-side view of the tentacles; C, out-side view of the tentacles." captionText-1="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." captionText-2="FIGURE 14. Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima (holotype specimen, WMNH-2015-INV-106), described as Patinapta deformis sp. nov., in the present study. The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stand for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi-0="http://doi.org/10.5281/zenodo.14703556" figureDoi-1="http://doi.org/10.5281/zenodo.14703560" figureDoi-2="http://doi.org/10.5281/zenodo.14703562" httpUri-0="https://zenodo.org/record/14703556/files/figure.png" httpUri-1="https://zenodo.org/record/14703560/files/figure.png" httpUri-2="https://zenodo.org/record/14703562/files/figure.png" pageId="22" pageNumber="23">Figs 1214</figureCitation>
;
<tableCitation id="C6E70332F9449D5AFED821B38DA1A9D5" box="[300,394,1417,1443]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="22" pageNumber="23">Tables 1</tableCitation>
,
<tableCitation id="C6E70332F9449D5AFE6C21B38D83A9D5" box="[408,424,1417,1443]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="22" pageNumber="23">6</tableCitation>
,
<tableCitation id="C6E70332F9449D5AFE4021B38DE9A9D5" box="[436,450,1417,1443]" captionStart="TABLE 8" captionStartId="31.[152,244,152,176]" captionTargetPageId="31" captionText="TABLE 8. List of WMNH-INV specimens of Patinapta deformis sp. nov. as of 2024." pageId="22" pageNumber="23">8</tableCitation>
)
</paragraph>
<paragraph id="8BDA3689F9449D5AFF6321FD8DA4A98B" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">
<specimenCount id="9D63FD00F9449D5AFF6321FD8D68A9A9" box="[151,323,1479,1503]" count="3" pageId="22" pageNumber="23" type="generic">Three specimens</specimenCount>
from
<location id="8EBA6052F9449D5AFE7521FD8E12A9A9" LSID="urn:lsid:plazi:treatment:03CC879FF9449D57FF63217B8936AB73:8EBA6052F9449D5AFE7521FD8E12A9A9" box="[385,569,1479,1503]" municipality="We" name="Yoron-jima Island" pageId="22" pageNumber="23">Yoron-jima Island</location>
were identified as new to science.
<collectingMunicipality id="6BBEACF3F9449D5AFC6B21FD8FEBA9A8" box="[927,960,1479,1502]" pageId="22" pageNumber="23">We</collectingMunicipality>
herein described the morphological characters of this new species.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5AFF63223D8DC3AAF8" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BDA3689F9449D5AFF63223D8DC3AAF8" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">
<emphasis id="B911EA9BF9449D5AFF63223D8D39AA57" bold="true" box="[151,274,1543,1569]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
Total shapes of anchor plates strongly distorted, their large perforations (mostly 23) fewer than those of the two congener nominal species (mostly 47), around these few perforations, marginal area broad and flat. Basal T-shaped end of anchor ossicles large, semicircular-shape. Ciliated funnels attached at inner side body wall of right dorso-lateral interradial (
<emphasis id="B911EA9BF9449D5AFE5B224E8DF3AAF8" box="[431,472,1652,1678]" italics="true" pageId="22" pageNumber="23">IR4</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5AFF3322A288F2AAC4" box="[199,1241,1687,1714]" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BDA3689F9449D5AFF3322A288F2AAC4" blockId="22.[151,1437,1479,2038]" box="[199,1241,1687,1714]" pageId="22" pageNumber="23">
<emphasis id="B911EA9BF9449D5AFF3322A28D78AAC7" bold="true" box="[199,339,1687,1714]" pageId="22" pageNumber="23">Type series.</emphasis>
<materialsCitation id="3B0D3CD4F9449D5AFEAE22A28EF7AAC4" box="[346,732,1687,1714]" pageId="22" pageNumber="23">
<typeStatus id="54DE882BF9449D5AFEAE22A28DEEAAC4" box="[346,453,1688,1714]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
, WMNH-2015-INV-106
</materialsCitation>
;
<materialsCitation id="3B0D3CD4F9449D5AFD1322A288FEAAC4" box="[743,1237,1687,1714]" pageId="22" pageNumber="23">
<specimenCount id="9D63FD00F9449D5AFD1322A28FADAAC4" box="[743,902,1688,1714]" count="2" pageId="22" pageNumber="23" type="generic" typeStatus="paratype">two paratypes</specimenCount>
, WMNH-2015-INV-105, 108
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5BFF3322818D56AC8F" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="8BDA3689F9449D5AFF3322818E71ABD8" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">
<emphasis id="B911EA9BF9449D5AFF3322818880AAA3" bold="true" box="[199,1195,1723,1750]" pageId="22" pageNumber="23">
Description of
<typeStatus id="54DE882BF9449D5AFE8322868DF5AAA0" box="[375,478,1724,1750]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(of WMNH-2015-INV-106: middle-size complete specimen).
</emphasis>
External and internal morphologies as follows (
<tableCitation id="C6E70332F9449D5AFE4122DA8E22AA8C" box="[437,521,1760,1786]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="22" pageNumber="23">Table 1</tableCitation>
): 30 years-preserved specimen pale peanut color (
<figureCitation id="135E2A0CF9449D5AFBC122DA88ACAA8C" box="[1077,1159,1760,1786]" captionStart="FIGURE 12" captionStartId="23.[151,250,1446,1470]" captionTargetBox="[151,1436,309,1423]" captionTargetId="figure-69@23.[151,1436,309,1423]" captionTargetPageId="23" captionText="FIGURE 12. External view of holotype specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (specimen WMNH-2015-INV-106): A, lateral view, anterior left, dorsal top; B, oral-side view of the tentacles; C, out-side view of the tentacles." figureDoi="http://doi.org/10.5281/zenodo.14703556" httpUri="https://zenodo.org/record/14703556/files/figure.png" pageId="22" pageNumber="23">Fig. 12</figureCitation>
). Tentacles 12, each with 10 digits of 5 pairs. On the oral-side of tentacle stem, about 1020 sensory cups sporadically arranged in U- shape. Polian vesicle in medio-ventral position, fusiform, double (
<tableCitation id="C6E70332F9449D5AFCB323128FB5AB34" box="[839,926,1832,1858]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="22" pageNumber="23">Table 1</tableCitation>
), stone canal single thread-like, attached to narrow space between basal parts of tentacle ampullae at mediodorsal (
<emphasis id="B911EA9BF9449D5AFC4323768FF4AB10" box="[951,991,1868,1894]" italics="true" pageId="22" pageNumber="23">IR5</emphasis>
), suddenly turned upward and forward, resulted in vesicle facing forward, distally with mold-like madreporite. Ciliated funnels in one crowded band along midline of right dorsal interradius (
<emphasis id="B911EA9BF9449D5AFDD523AE8E61ABD8" box="[545,586,1940,1966]" italics="true" pageId="22" pageNumber="23">IR4</emphasis>
).
</paragraph>
<paragraph id="8BDA3689F9449D5AFF332382889CAB80" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">Inside body, two tufts of gonad tubules attached with both sides of anterior dorsal mesentery, also two whitish tubule organs adhered to upper and under sides of intestine canal. Intestine canal lacking loop.</paragraph>
<paragraph id="8BDA3689F9459D5BFF3324AD8D56AC8F" blockId="23.[151,1436,151,285]" pageId="23" pageNumber="24">Body wall coarse surface, hard and thick, sparsely scattered with ossicles, arranging latitudinal layout of all as sets of anchor and plate, their directions somewhat ordered but without symmetrical design against each axial of longitudinal muscle.</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9459D56FF3325398824A9B3" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BDA3689F9459D5BFF33253988B3AD6B" blockId="23.[151,1436,151,285]" box="[199,1176,259,285]" pageId="23" pageNumber="24">
<emphasis id="B911EA9BF9459D5BFF3325398D65AD6B" bold="true" box="[199,334,259,285]" pageId="23" pageNumber="24">Etymology.</emphasis>
The species name
<taxonomicName id="4C654D0AF9459D5BFDD625398EAFAD6B" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[546,644,259,285]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="deformis">
<emphasis id="B911EA9BF9459D5BFDD625398EAFAD6B" box="[546,644,259,285]" italics="true" pageId="23" pageNumber="24">deformis</emphasis>
</taxonomicName>
, alluding to the strongly distorted anchor plate.
</paragraph>
<caption id="DF1A6601F9459D5BFF63219C8EE8AA70" ID-DOI="http://doi.org/10.5281/zenodo.14703556" ID-Zenodo-Dep="14703556" httpUri="https://zenodo.org/record/14703556/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1446,1470]" targetBox="[151,1436,309,1423]" targetPageId="23" targetType="figure">
<paragraph id="8BDA3689F9459D5BFF63219C8EE8AA70" blockId="23.[151,1437,1446,1543]" pageId="23" pageNumber="24">
<emphasis id="B911EA9BF9459D5BFF63219C8D0FA9C8" bold="true" box="[151,292,1446,1470]" pageId="23" pageNumber="24">FIGURE 12.</emphasis>
External view of holotype specimen collected from Yoron-jima Island, Kagoshima, described as
<taxonomicName id="4C654D0AF9459D5BFACC219D8CDBA994" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF9459D5BFACC219D8CDBA994" italics="true" pageId="23" pageNumber="24">Patinapta deformis</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9459D5BFF0221F18D6DA995" bold="true" box="[246,326,1483,1507]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="A22257E0F9459D5BFF0221F18D6DA995" box="[246,326,1483,1507]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study (specimen WMNH-2015-INV-106): A, lateral view, anterior left, dorsal top; B, oral-side view of the tentacles; C, out-side view of the tentacles.
</paragraph>
</caption>
<paragraph id="8BDA3689F9459D5BFF6322638E86AA04" blockId="23.[151,685,1624,1651]" box="[151,685,1624,1651]" pageId="23" pageNumber="24">
<heading id="D09281E5F9459D5BFF6322638E86AA04" bold="true" box="[151,685,1624,1651]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B911EA9BF9459D5BFF6322638E86AA04" bold="true" box="[151,685,1624,1651]" pageId="23" pageNumber="24">The present results of ossicle morphologies</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9459D5BFF63229B8D4BAB19" blockId="23.[151,1437,1697,1903]" pageId="23" pageNumber="24">
Body wall ossicles anchors and anchor plates (
<figureCitation id="135E2A0CF9459D5BFD59229B8F13AACD" box="[685,824,1697,1723]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="23" pageNumber="24">Fig. 13B, D</figureCitation>
;
<tableCitation id="C6E70332F9459D5BFCB1229B8FB7AACD" box="[837,924,1697,1723]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="23" pageNumber="24">Table 6</tableCitation>
). Anchor plates ranged within 98128 μm in length. Size of perforations noticeably variable between central area and marginal area. Largest perforation situated noticeably biased from the center, surrounded by scattering smaller perforations. Few numbers of largest perforations and several small perforations equipped with 418 spaced sharp-teeth arraign along with inner rim. Two calculated indicators for anchor plates and a parameter were variated within
<emphasis id="B911EA9BF9459D5BFC82230B8F8DAB3D" box="[886,934,1841,1867]" italics="true" pageId="23" pageNumber="24">APS</emphasis>
=0.622.4;
<emphasis id="B911EA9BF9459D5BFBD3230B8864AB3C" box="[1063,1103,1841,1866]" italics="true" pageId="23" pageNumber="24">Ntp</emphasis>
=27;
<emphasis id="B911EA9BF9459D5BFB6C230B88E1AB3D" box="[1176,1226,1841,1867]" italics="true" pageId="23" pageNumber="24">APT</emphasis>
=11.820.8% (
<figureCitation id="135E2A0CF9459D5BFA9B230B8CD9AB19" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="23" pageNumber="24">Fig. 13 B, D</figureCitation>
;
<tableCitation id="C6E70332F9459D5BFF09236F8D79AB19" box="[253,338,1877,1903]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="23" pageNumber="24">Table 6</tableCitation>
).
</paragraph>
<caption id="DF1A6601F94A9D54FF6C24AD8942ACC6" box="[152,1385,151,176]" pageId="24" pageNumber="25" startId="24.[152,244,151,175]" targetBox="[163,1425,192,1866]" targetIsTable="true" targetPageId="24" targetType="table">
<paragraph id="8BDA3689F94A9D54FF6C24AD8942ACC6" blockId="24.[152,1385,151,176]" box="[152,1385,151,176]" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF6C24AD8D25ACD9" bold="true" box="[152,270,151,175]" pageId="24" pageNumber="25">TABLE 6.</emphasis>
Measurements (mean±SD μm) and counts of ossicles of three specimens of
<taxonomicName id="4C654D0AF94A9D54FBB424A28926ACD9" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[1088,1293,151,175]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="24" pageNumber="25" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF94A9D54FBB424A28926ACD9" box="[1088,1293,151,175]" italics="true" pageId="24" pageNumber="25">Patinapta deformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257E0F94A9D54FAE024A28942ACC6" box="[1300,1385,152,176]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BDA3689F94A9D54FF5724FA89BAAB3C" pageId="24" pageNumber="25">
<table id="F965C429F94A62B3FF5724FA89BAAB3C" box="[163,1425,192,1866]" gridcols="8" gridrows="42" pageId="24" pageNumber="25">
<tr id="355534CBF94A62B3FF5724FA89BAACAE" box="[163,1425,192,216]" gridrow="0" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5724FA8D32ACAE" box="[163,281,192,216]" gridcol="0" gridrow="0" pageId="24" pageNumber="25">Body parts</th>
<th id="76845DB7F94A62B3FEB824FA8DD6ACAE" box="[332,509,192,216]" gridcol="1" gridrow="0" pageId="24" pageNumber="25">(Ossicle-type)</th>
<th id="76845DB7F94A62B3FDEE24FA8F0BACAE" box="[538,800,192,216]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="24" pageNumber="25">2015-INV-105</th>
<th id="76845DB7F94A62B3FCC824FA887FACAE" box="[828,1108,192,216]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="24" pageNumber="25">2015-INV-106</th>
<th id="76845DB7F94A62B3FB8F24FA89BAACAE" box="[1147,1425,192,216]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="24" pageNumber="25">2015-INV-108</th>
</tr>
<tr id="355534CBF94A62B3FF5724D289BAAD76" box="[163,1425,232,256]" gridrow="1" pageId="24" pageNumber="25" rowspan-7="1">
<th id="76845DB7F94A62B3FF5724D28D32AD76" box="[163,281,232,256]" gridcol="0" gridrow="1" pageId="24" pageNumber="25">Tentacle</th>
<td id="76845DB7F94A62B3FEB824D28DD6AD76" box="[332,509,232,256]" gridcol="1" gridrow="1" pageId="24" pageNumber="25">
Rods (
<emphasis id="B911EA9BF94A9D54FE4F24D38DE3AD76" box="[443,456,233,256]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE24D28F0BAD76" box="[538,800,232,256]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="24" pageNumber="25">13</td>
<td id="76845DB7F94A62B3FCC824D2887FAD76" box="[828,1108,232,256]" colspan="2" colspanRight="1" gridcol="4" gridrow="1" pageId="24" pageNumber="25">11</td>
<td id="76845DB7F94A62B3FB8F24D28922AD76" box="[1147,1289,232,256]" gridcol="6" gridrow="1" pageId="24" pageNumber="25">5</td>
</tr>
<tr id="355534CBF94A62B3FF57252A89BAAD5E" box="[163,1425,272,296]" gridrow="2" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57252A8D32AD5E" box="[163,281,272,296]" gridcol="0" gridrow="2" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB8252A8DD6AD5E" box="[332,509,272,296]" gridcol="1" gridrow="2" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE252A8EA5AD5E" box="[538,654,272,296]" gridcol="2" gridrow="2" pageId="24" pageNumber="25">53.7±14.3</td>
<td id="76845DB7F94A62B3FD5E252A8F0BAD5E" box="[682,800,272,296]" gridcol="3" gridrow="2" pageId="24" pageNumber="25">(3577)</td>
<td id="76845DB7F94A62B3FCC8252A8F99AD5E" box="[828,946,272,296]" gridcol="4" gridrow="2" pageId="24" pageNumber="25">63.8± 11.9</td>
<td id="76845DB7F94A62B3FC30252A887FAD5E" box="[964,1108,272,296]" gridcol="5" gridrow="2" pageId="24" pageNumber="25">(5285)</td>
<td id="76845DB7F94A62B3FB8F252A8922AD5E" box="[1147,1289,272,296]" gridcol="6" gridrow="2" pageId="24" pageNumber="25">67.4± 7.6</td>
<td id="76845DB7F94A62B3FAEF252A89BAAD5E" box="[1307,1425,272,296]" gridcol="7" gridrow="2" pageId="24" pageNumber="25">(6080)</td>
</tr>
<tr id="355534CBF94A62B3FF57250289BAAD27" box="[163,1425,312,337]" gridrow="3" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5725028D32AD27" box="[163,281,312,337]" gridcol="0" gridrow="3" pageId="24" pageNumber="25">Breadth</th>
<td id="76845DB7F94A62B3FEB825028DD6AD27" box="[332,509,312,337]" gridcol="1" gridrow="3" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE25028EA5AD27" box="[538,654,312,337]" gridcol="2" gridrow="3" pageId="24" pageNumber="25">8.5± 2.5</td>
<td id="76845DB7F94A62B3FD5E25028F0BAD27" box="[682,800,312,337]" gridcol="3" gridrow="3" pageId="24" pageNumber="25">(514)</td>
<td id="76845DB7F94A62B3FCC825028F99AD27" box="[828,946,312,337]" gridcol="4" gridrow="3" pageId="24" pageNumber="25">9.4± 2.7</td>
<td id="76845DB7F94A62B3FC302502887FAD27" box="[964,1108,312,337]" gridcol="5" gridrow="3" pageId="24" pageNumber="25">(715)</td>
<td id="76845DB7F94A62B3FB8F25028922AD27" box="[1147,1289,312,337]" gridcol="6" gridrow="3" pageId="24" pageNumber="25">10.6± 1.1</td>
<td id="76845DB7F94A62B3FAEF250289BAAD27" box="[1307,1425,312,337]" gridcol="7" gridrow="3" pageId="24" pageNumber="25">(912)</td>
</tr>
<tr id="355534CBF94A62B3FF57255A89BAAD0F" box="[163,1425,352,377]" gridrow="4" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57255A8D32AD0F" box="[163,281,352,377]" gridcol="0" gridrow="4" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4425588CE4AD0F" box="[176,207,354,377]" italics="true" pageId="24" pageNumber="25">RP</emphasis>
a (%)
</th>
<td id="76845DB7F94A62B3FEB8255A8DD6AD0F" box="[332,509,352,377]" gridcol="1" gridrow="4" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE255A8EA5AD0F" box="[538,654,352,377]" gridcol="2" gridrow="4" pageId="24" pageNumber="25">16.3± 5.0</td>
<td id="76845DB7F94A62B3FD5E255A8F0BAD0F" box="[682,800,352,377]" gridcol="3" gridrow="4" pageId="24" pageNumber="25">(10.126.3)</td>
<td id="76845DB7F94A62B3FCC8255A8F99AD0F" box="[828,946,352,377]" gridcol="4" gridrow="4" pageId="24" pageNumber="25">14.7± 3.0</td>
<td id="76845DB7F94A62B3FC30255A887FAD0F" box="[964,1108,352,377]" gridcol="5" gridrow="4" pageId="24" pageNumber="25">(10.720.3)</td>
<td id="76845DB7F94A62B3FB8F255A8922AD0F" box="[1147,1289,352,377]" gridcol="6" gridrow="4" pageId="24" pageNumber="25">15.8± 2.1</td>
<td id="76845DB7F94A62B3FAEF255A89BAAD0F" box="[1307,1425,352,377]" gridcol="7" gridrow="4" pageId="24" pageNumber="25">(13.818.3)</td>
</tr>
<tr id="355534CBF94A62B3FF5725B389BAADD7" box="[163,1425,393,417]" gridrow="5" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5725B38D32ADD7" box="[163,281,393,417]" gridcol="0" gridrow="5" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4425B38CFBADD6" box="[176,208,393,416]" italics="true" pageId="24" pageNumber="25">RC</emphasis>
b
</th>
<td id="76845DB7F94A62B3FEB825B38DD6ADD7" box="[332,509,393,417]" gridcol="1" gridrow="5" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE25B38EA5ADD7" box="[538,654,393,417]" gridcol="2" gridrow="5" pageId="24" pageNumber="25">11.1± 1.7</td>
<td id="76845DB7F94A62B3FD5E25B38F0BADD7" box="[682,800,393,417]" gridcol="3" gridrow="5" pageId="24" pageNumber="25">(5.317.3)</td>
<td id="76845DB7F94A62B3FCC825B38F99ADD7" box="[828,946,393,417]" gridcol="4" gridrow="5" pageId="24" pageNumber="25">7.6± 1.7</td>
<td id="76845DB7F94A62B3FC3025B3887FADD7" box="[964,1108,393,417]" gridcol="5" gridrow="5" pageId="24" pageNumber="25">(5.412.2)</td>
<td id="76845DB7F94A62B3FB8F25B38922ADD7" box="[1147,1289,393,417]" gridcol="6" gridrow="5" pageId="24" pageNumber="25">7.9± 2.7</td>
<td id="76845DB7F94A62B3FAEF25B389BAADD7" box="[1307,1425,393,417]" gridcol="7" gridrow="5" pageId="24" pageNumber="25">(5.011.7)</td>
</tr>
<tr id="355534CBF94A62B3FF57258B89BAADBF" box="[163,1425,433,457]" gridrow="6" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57258B89BAADBF" box="[163,1425,433,457]" colspan="8" colspanRight="7" gridcol="0" gridrow="6" pageId="24" pageNumber="25">Ventral side</th>
</tr>
<tr id="355534CBF94A62B3FF5725E389BAAD84" box="[163,1425,473,498]" gridrow="7" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF5725E38D32AD84" box="[163,281,473,498]" gridcol="0" gridrow="7" pageId="24" pageNumber="25">L-Muscle</th>
<td id="76845DB7F94A62B3FEB825E38DD6AD84" box="[332,509,473,498]" gridcol="1" gridrow="7" pageId="24" pageNumber="25">
Granules (
<emphasis id="B911EA9BF94A9D54FE3B25E08DF7AD87" box="[463,476,474,497]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE25E38F0BAD84" box="[538,800,473,498]" colspan="2" colspanRight="1" gridcol="2" gridrow="7" pageId="24" pageNumber="25">7</td>
<td id="76845DB7F94A62B3FC3025E3887FAD84" box="[964,1108,473,498]" gridcol="5" gridrow="7" pageId="24" pageNumber="25">9</td>
<td id="76845DB7F94A62B3FB8F25E38922AD84" box="[1147,1289,473,498]" gridcol="6" gridrow="7" pageId="24" pageNumber="25">9</td>
</tr>
<tr id="355534CBF94A62B3FF57263889BAAE6C" box="[163,1425,514,538]" gridrow="8" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726388D32AE6C" box="[163,281,514,538]" gridcol="0" gridrow="8" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB826388DD6AE6C" box="[332,509,514,538]" gridcol="1" gridrow="8" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE26388EA5AE6C" box="[538,654,514,538]" gridcol="2" gridrow="8" pageId="24" pageNumber="25">30.3 ± 3.9</td>
<td id="76845DB7F94A62B3FD5E26388F0BAE6C" box="[682,800,514,538]" gridcol="3" gridrow="8" pageId="24" pageNumber="25">(2436)</td>
<td id="76845DB7F94A62B3FCC826388F99AE6C" box="[828,946,514,538]" gridcol="4" gridrow="8" pageId="24" pageNumber="25">29.3 ± 4.4</td>
<td id="76845DB7F94A62B3FC302638887FAE6C" box="[964,1108,514,538]" gridcol="5" gridrow="8" pageId="24" pageNumber="25">(2336)</td>
<td id="76845DB7F94A62B3FB8F26388922AE6C" box="[1147,1289,514,538]" gridcol="6" gridrow="8" pageId="24" pageNumber="25">28.8 ± 3.9</td>
<td id="76845DB7F94A62B3FAEF263889BAAE6C" box="[1307,1425,514,538]" gridcol="7" gridrow="8" pageId="24" pageNumber="25">(2434)</td>
</tr>
<tr id="355534CBF94A62B3FF57261089BAAE34" box="[163,1425,554,578]" gridrow="9" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726108D32AE34" box="[163,281,554,578]" gridcol="0" gridrow="9" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB826108DD6AE34" box="[332,509,554,578]" gridcol="1" gridrow="9" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE26108EA5AE34" box="[538,654,554,578]" gridcol="2" gridrow="9" pageId="24" pageNumber="25">14.3 ±1.5</td>
<td id="76845DB7F94A62B3FD5E26108F0BAE34" box="[682,800,554,578]" gridcol="3" gridrow="9" pageId="24" pageNumber="25">(1216)</td>
<td id="76845DB7F94A62B3FCC826108F99AE34" box="[828,946,554,578]" gridcol="4" gridrow="9" pageId="24" pageNumber="25">16.8 ± 2.3</td>
<td id="76845DB7F94A62B3FC302610887FAE34" box="[964,1108,554,578]" gridcol="5" gridrow="9" pageId="24" pageNumber="25">(1321)</td>
<td id="76845DB7F94A62B3FB8F26108922AE34" box="[1147,1289,554,578]" gridcol="6" gridrow="9" pageId="24" pageNumber="25">13.8 ± 1.9</td>
<td id="76845DB7F94A62B3FAEF261089BAAE34" box="[1307,1425,554,578]" gridcol="7" gridrow="9" pageId="24" pageNumber="25">(1017)</td>
</tr>
<tr id="355534CBF94A62B3FF57266B89BAAE1C" box="[163,1425,593,618]" gridrow="10" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57266B8D32AE1C" box="[163,281,593,618]" gridcol="0" gridrow="10" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4426698CF9AE1C" box="[176,210,595,618]" italics="true" pageId="24" pageNumber="25">GP</emphasis>
c
</th>
<td id="76845DB7F94A62B3FEB8266B8DD6AE1C" box="[332,509,593,618]" gridcol="1" gridrow="10" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE266B8EA5AE1C" box="[538,654,593,618]" gridcol="2" gridrow="10" pageId="24" pageNumber="25">48.4 ± 11.2</td>
<td id="76845DB7F94A62B3FD5E266B8F0BAE1C" box="[682,800,593,618]" gridcol="3" gridrow="10" pageId="24" pageNumber="25">(36.166.7)</td>
<td id="76845DB7F94A62B3FCC8266B887FAE1C" box="[828,1108,593,618]" colspan="2" colspanRight="1" gridcol="4" gridrow="10" pageId="24" pageNumber="25">59.5 ± 15.2 (36.182.6)</td>
<td id="76845DB7F94A62B3FB8F266B8922AE1C" box="[1147,1289,593,618]" gridcol="6" gridrow="10" pageId="24" pageNumber="25">48.2 ± 5.9</td>
<td id="76845DB7F94A62B3FAEF266B89BAAE1C" box="[1307,1425,593,618]" gridcol="7" gridrow="10" pageId="24" pageNumber="25">(39.455.6)</td>
</tr>
<tr id="355534CBF94A62B3FF57264089BAAEE5" box="[163,1425,634,659]" gridrow="11" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726408D32AEE5" box="[163,281,634,659]" gridcol="0" gridrow="11" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4426418CCEAEE4" box="[176,229,635,658]" italics="true" pageId="24" pageNumber="25">GCO</emphasis>
d (%)
</th>
<td id="76845DB7F94A62B3FEB826408DD6AEE5" box="[332,509,634,659]" gridcol="1" gridrow="11" pageId="24" pageNumber="25">(Ratio)</td>
<td id="76845DB7F94A62B3FDEE26408EA5AEE5" box="[538,654,634,659]" gridcol="2" gridrow="11" pageId="24" pageNumber="25">14.3</td>
<td id="76845DB7F94A62B3FD5E26408F0BAEE5" box="[682,800,634,659]" gridcol="3" gridrow="11" pageId="24" pageNumber="25">(O1: C6)</td>
<td id="76845DB7F94A62B3FCC826408F99AEE5" box="[828,946,634,659]" gridcol="4" gridrow="11" pageId="24" pageNumber="25">44.4</td>
<td id="76845DB7F94A62B3FC302640887FAEE5" box="[964,1108,634,659]" gridcol="5" gridrow="11" pageId="24" pageNumber="25">(O4: C5)</td>
<td id="76845DB7F94A62B3FB8F26408922AEE5" box="[1147,1289,634,659]" gridcol="6" gridrow="11" pageId="24" pageNumber="25">22.2</td>
<td id="76845DB7F94A62B3FAEF264089BAAEE5" box="[1307,1425,634,659]" gridcol="7" gridrow="11" pageId="24" pageNumber="25">(O2: C7)</td>
</tr>
<tr id="355534CBF94A62B3FF57269989BAAECD" box="[163,1425,675,699]" gridrow="12" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF5726998D32AECD" box="[163,281,675,699]" gridcol="0" gridrow="12" pageId="24" pageNumber="25">Body wall</th>
<td id="76845DB7F94A62B3FEB826998DD6AECD" box="[332,509,675,699]" gridcol="1" gridrow="12" pageId="24" pageNumber="25">
Anchor plates (
<emphasis id="B911EA9BF94A9D54FE1C269E8DDEAECD" box="[488,501,676,699]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE26998F0BAECD" box="[538,800,675,699]" colspan="2" colspanRight="1" gridcol="2" gridrow="12" pageId="24" pageNumber="25">7</td>
<td id="76845DB7F94A62B3FC302699887FAECD" box="[964,1108,675,699]" gridcol="5" gridrow="12" pageId="24" pageNumber="25">4</td>
<td id="76845DB7F94A62B3FB8F26998922AECD" box="[1147,1289,675,699]" gridcol="6" gridrow="12" pageId="24" pageNumber="25">7</td>
</tr>
<tr id="355534CBF94A62B3FF5726F189BAAE95" box="[163,1425,715,739]" gridrow="13" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726F18D32AE95" box="[163,281,715,739]" gridcol="0" gridrow="13" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB826F18DD6AE95" box="[332,509,715,739]" gridcol="1" gridrow="13" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE26F18EA5AE95" box="[538,654,715,739]" gridcol="2" gridrow="13" pageId="24" pageNumber="25">127.7 ± 6.3</td>
<td id="76845DB7F94A62B3FD5E26F18F0BAE95" box="[682,800,715,739]" gridcol="3" gridrow="13" pageId="24" pageNumber="25">(119138)</td>
<td id="76845DB7F94A62B3FCC826F1887FAE95" box="[828,1108,715,739]" colspan="2" colspanRight="1" gridcol="4" gridrow="13" pageId="24" pageNumber="25">110.5 ± 6.2 (104119)</td>
<td id="76845DB7F94A62B3FB8F26F18922AE95" box="[1147,1289,715,739]" gridcol="6" gridrow="13" pageId="24" pageNumber="25">95.7 ± 8.5</td>
<td id="76845DB7F94A62B3FAEF26F189BAAE95" box="[1307,1425,715,739]" gridcol="7" gridrow="13" pageId="24" pageNumber="25">(83108)</td>
</tr>
<tr id="355534CBF94A62B3FF5726C989BAAF7D" box="[163,1425,755,779]" gridrow="14" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726C98D32AF7D" box="[163,281,755,779]" gridcol="0" gridrow="14" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB826C98DD6AF7D" box="[332,509,755,779]" gridcol="1" gridrow="14" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE26C98EA5AF7D" box="[538,654,755,779]" gridcol="2" gridrow="14" pageId="24" pageNumber="25">99.0 ± 5.1</td>
<td id="76845DB7F94A62B3FD5E26C98F0BAF7D" box="[682,800,755,779]" gridcol="3" gridrow="14" pageId="24" pageNumber="25">(94107)</td>
<td id="76845DB7F94A62B3FCC826C98F99AF7D" box="[828,946,755,779]" gridcol="4" gridrow="14" pageId="24" pageNumber="25">70.5 ± 3.9</td>
<td id="76845DB7F94A62B3FC3026C9887FAF7D" box="[964,1108,755,779]" gridcol="5" gridrow="14" pageId="24" pageNumber="25">(6776)</td>
<td id="76845DB7F94A62B3FB8F26C98922AF7D" box="[1147,1289,755,779]" gridcol="6" gridrow="14" pageId="24" pageNumber="25">64.1 ± 6.4</td>
<td id="76845DB7F94A62B3FAEF26C989BAAF7D" box="[1307,1425,755,779]" gridcol="7" gridrow="14" pageId="24" pageNumber="25">(5469)</td>
</tr>
<tr id="355534CBF94A62B3FF57272189BAAF42" box="[163,1425,795,820]" gridrow="15" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5727218D32AF42" box="[163,281,795,820]" gridcol="0" gridrow="15" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4427268CF7AF45" box="[176,220,796,819]" italics="true" pageId="24" pageNumber="25">APS</emphasis>
e
</th>
<td id="76845DB7F94A62B3FEB827218DD6AF42" box="[332,509,795,820]" gridcol="1" gridrow="15" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE27218EA5AF42" box="[538,654,795,820]" gridcol="2" gridrow="15" pageId="24" pageNumber="25">9.9 ± 5.7</td>
<td id="76845DB7F94A62B3FD5E27218F0BAF42" box="[682,800,795,820]" gridcol="3" gridrow="15" pageId="24" pageNumber="25">(2.313.5)</td>
<td id="76845DB7F94A62B3FCC827218F99AF42" box="[828,946,795,820]" gridcol="4" gridrow="15" pageId="24" pageNumber="25">5.1 ± 4.8</td>
<td id="76845DB7F94A62B3FC302721887FAF42" box="[964,1108,795,820]" gridcol="5" gridrow="15" pageId="24" pageNumber="25">(0.611.7)</td>
<td id="76845DB7F94A62B3FB8F27218922AF42" box="[1147,1289,795,820]" gridcol="6" gridrow="15" pageId="24" pageNumber="25">2.5 ± 2.5</td>
<td id="76845DB7F94A62B3FAEF272189BAAF42" box="[1307,1425,795,820]" gridcol="7" gridrow="15" pageId="24" pageNumber="25">(0.66.1)</td>
</tr>
<tr id="355534CBF94A62B3FF57277E89BAAF2A" box="[163,1425,836,860]" gridrow="16" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57277E8D32AF2A" box="[163,281,836,860]" gridcol="0" gridrow="16" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44277F8CFEAF2A" box="[176,213,837,860]" italics="true" pageId="24" pageNumber="25">Ntp</emphasis>
f
</th>
<td id="76845DB7F94A62B3FEB8277E8DD6AF2A" box="[332,509,836,860]" gridcol="1" gridrow="16" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE277E8EA5AF2A" box="[538,654,836,860]" gridcol="2" gridrow="16" pageId="24" pageNumber="25">3.6 ± 1.4</td>
<td id="76845DB7F94A62B3FD5E277E8F0BAF2A" box="[682,800,836,860]" gridcol="3" gridrow="16" pageId="24" pageNumber="25">(15)</td>
<td id="76845DB7F94A62B3FCC8277E8F99AF2A" box="[828,946,836,860]" gridcol="4" gridrow="16" pageId="24" pageNumber="25">4.5 ± 1.7</td>
<td id="76845DB7F94A62B3FC30277E887FAF2A" box="[964,1108,836,860]" gridcol="5" gridrow="16" pageId="24" pageNumber="25">(37)</td>
<td id="76845DB7F94A62B3FB8F277E8922AF2A" box="[1147,1289,836,860]" gridcol="6" gridrow="16" pageId="24" pageNumber="25">2.3 ± 0.8</td>
<td id="76845DB7F94A62B3FAEF277E89BAAF2A" box="[1307,1425,836,860]" gridcol="7" gridrow="16" pageId="24" pageNumber="25">(13)</td>
</tr>
<tr id="355534CBF94A62B3FF57275189BAAFF2" box="[163,1425,875,900]" gridrow="17" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5727518D32AFF2" box="[163,281,875,900]" gridcol="0" gridrow="17" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4427568CF6AFF5" box="[176,221,876,899]" italics="true" pageId="24" pageNumber="25">APT</emphasis>
g (%)
</th>
<td id="76845DB7F94A62B3FEB827518DD6AFF2" box="[332,509,875,900]" gridcol="1" gridrow="17" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE27518EA5AFF2" box="[538,654,875,900]" gridcol="2" gridrow="17" pageId="24" pageNumber="25">9.9 ± 3.7</td>
<td id="76845DB7F94A62B3FD5E27518F0BAFF2" box="[682,800,875,900]" gridcol="3" gridrow="17" pageId="24" pageNumber="25">(2.313.5)</td>
<td id="76845DB7F94A62B3FCC827518F99AFF2" box="[828,946,875,900]" gridcol="4" gridrow="17" pageId="24" pageNumber="25">14.6 ± 2.1</td>
<td id="76845DB7F94A62B3FC302751887FAFF2" box="[964,1108,875,900]" gridcol="5" gridrow="17" pageId="24" pageNumber="25">(11.816.7)</td>
<td id="76845DB7F94A62B3FB8F27518922AFF2" box="[1147,1289,875,900]" gridcol="6" gridrow="17" pageId="24" pageNumber="25">11.8 ± 3.8</td>
<td id="76845DB7F94A62B3FAEF275189BAAFF2" box="[1307,1425,875,900]" gridcol="7" gridrow="17" pageId="24" pageNumber="25">(8.018.8)</td>
</tr>
<tr id="355534CBF94A62B3FF5727AE89BAAFDB" box="[163,1425,916,941]" gridrow="18" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF5727AE8D32AFDB" box="[163,281,916,941]" gridcol="0" gridrow="18" pageId="24" pageNumber="25">Body wall</th>
<td id="76845DB7F94A62B3FEB827AE8DD6AFDB" box="[332,509,916,941]" gridcol="1" gridrow="18" pageId="24" pageNumber="25">
Anchor (
<emphasis id="B911EA9BF94A9D54FE3327AF8DFFAFDA" box="[455,468,917,940]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE27AE8F0BAFDB" box="[538,800,916,941]" colspan="2" colspanRight="1" gridcol="2" gridrow="18" pageId="24" pageNumber="25">7</td>
<td id="76845DB7F94A62B3FC3027AE887FAFDB" box="[964,1108,916,941]" gridcol="5" gridrow="18" pageId="24" pageNumber="25">4</td>
<td id="76845DB7F94A62B3FB8F27AE8922AFDB" box="[1147,1289,916,941]" gridcol="6" gridrow="18" pageId="24" pageNumber="25">8</td>
</tr>
<tr id="355534CBF94A62B3FF57278689BAAFA3" box="[163,1425,956,981]" gridrow="19" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5727868D32AFA3" box="[163,281,956,981]" gridcol="0" gridrow="19" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB827868DD6AFA3" box="[332,509,956,981]" gridcol="1" gridrow="19" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE27868EA5AFA3" box="[538,654,956,981]" gridcol="2" gridrow="19" pageId="24" pageNumber="25">166.3±9.5</td>
<td id="76845DB7F94A62B3FD5E27868F0BAFA3" box="[682,800,956,981]" gridcol="3" gridrow="19" pageId="24" pageNumber="25">(157181)</td>
<td id="76845DB7F94A62B3FCC82786887FAFA3" box="[828,1108,956,981]" colspan="2" colspanRight="1" gridcol="4" gridrow="19" pageId="24" pageNumber="25">157.8 ± 12.5 (144174)</td>
<td id="76845DB7F94A62B3FB8F27868922AFA3" box="[1147,1289,956,981]" gridcol="6" gridrow="19" pageId="24" pageNumber="25">158.1 ± 7.1</td>
<td id="76845DB7F94A62B3FAEF278689BAAFA3" box="[1307,1425,956,981]" gridcol="7" gridrow="19" pageId="24" pageNumber="25">(147169)</td>
</tr>
<tr id="355534CBF94A62B3FF5727DF89BAAF8B" box="[163,1425,997,1021]" gridrow="20" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5727DF8D32AF8B" box="[163,281,997,1021]" gridcol="0" gridrow="20" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB827DF8DD6AF8B" box="[332,509,997,1021]" gridcol="1" gridrow="20" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE27DF8EA5AF8B" box="[538,654,997,1021]" gridcol="2" gridrow="20" pageId="24" pageNumber="25">85.0 ± 5.9</td>
<td id="76845DB7F94A62B3FD5E27DF8F0BAF8B" box="[682,800,997,1021]" gridcol="3" gridrow="20" pageId="24" pageNumber="25">(7693)</td>
<td id="76845DB7F94A62B3FCC827DF8F99AF8B" box="[828,946,997,1021]" gridcol="4" gridrow="20" pageId="24" pageNumber="25">85.3 ± 5.2</td>
<td id="76845DB7F94A62B3FC3027DF887FAF8B" box="[964,1108,997,1021]" gridcol="5" gridrow="20" pageId="24" pageNumber="25">(7990)</td>
<td id="76845DB7F94A62B3FB8F27DF8922AF8B" box="[1147,1289,997,1021]" gridcol="6" gridrow="20" pageId="24" pageNumber="25">68.5 ± 9.4</td>
<td id="76845DB7F94A62B3FAEF27DF89BAAF8B" box="[1307,1425,997,1021]" gridcol="7" gridrow="20" pageId="24" pageNumber="25">(5884)</td>
</tr>
<tr id="355534CBF94A62B3FF57203789BAA853" box="[163,1425,1037,1061]" gridrow="21" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720378D32A853" box="[163,281,1037,1061]" gridcol="0" gridrow="21" pageId="24" pageNumber="25">Teeth</th>
<td id="76845DB7F94A62B3FEB820378DD6A853" box="[332,509,1037,1061]" gridcol="1" gridrow="21" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE20378EA5A853" box="[538,654,1037,1061]" gridcol="2" gridrow="21" pageId="24" pageNumber="25">1.6 ± 0.9</td>
<td id="76845DB7F94A62B3FD5E20378F0BA853" box="[682,800,1037,1061]" gridcol="3" gridrow="21" pageId="24" pageNumber="25">(03)</td>
<td id="76845DB7F94A62B3FCC820378F99A853" box="[828,946,1037,1061]" gridcol="4" gridrow="21" pageId="24" pageNumber="25">3.3 ± 1.5</td>
<td id="76845DB7F94A62B3FC302037887FA853" box="[964,1108,1037,1061]" gridcol="5" gridrow="21" pageId="24" pageNumber="25">(25)</td>
<td id="76845DB7F94A62B3FB8F20378922A853" box="[1147,1289,1037,1061]" gridcol="6" gridrow="21" pageId="24" pageNumber="25">3.7 ± 0.7</td>
<td id="76845DB7F94A62B3FAEF203789BAA853" box="[1307,1425,1037,1061]" gridcol="7" gridrow="21" pageId="24" pageNumber="25">(35)</td>
</tr>
<tr id="355534CBF94A62B3FF57200F89BAA838" box="[163,1425,1077,1102]" gridrow="22" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57200F8D32A838" box="[163,281,1077,1102]" gridcol="0" gridrow="22" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44200C8CCAA83B" box="[176,225,1078,1101]" italics="true" pageId="24" pageNumber="25">ASW</emphasis>
h(%)
</th>
<td id="76845DB7F94A62B3FEB8200F8DD6A838" box="[332,509,1077,1102]" gridcol="1" gridrow="22" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE200F8EA5A838" box="[538,654,1077,1102]" gridcol="2" gridrow="22" pageId="24" pageNumber="25">23.1 ± 1.2</td>
<td id="76845DB7F94A62B3FD5E200F8F0BA838" box="[682,800,1077,1102]" gridcol="3" gridrow="22" pageId="24" pageNumber="25">(21.525.0)</td>
<td id="76845DB7F94A62B3FCC8200F8F99A838" box="[828,946,1077,1102]" gridcol="4" gridrow="22" pageId="24" pageNumber="25">20.1 ± 0.7</td>
<td id="76845DB7F94A62B3FC30200F887FA838" box="[964,1108,1077,1102]" gridcol="5" gridrow="22" pageId="24" pageNumber="25">(19.421.1)</td>
<td id="76845DB7F94A62B3FB8F200F8922A838" box="[1147,1289,1077,1102]" gridcol="6" gridrow="22" pageId="24" pageNumber="25">25.0 ± 2.1</td>
<td id="76845DB7F94A62B3FAEF200F89BAA838" box="[1307,1425,1077,1102]" gridcol="7" gridrow="22" pageId="24" pageNumber="25">(21.428.4)</td>
</tr>
<tr id="355534CBF94A62B3FF57206789BAA800" box="[163,1425,1117,1142]" gridrow="23" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720678D32A800" box="[163,281,1117,1142]" gridcol="0" gridrow="23" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4420648CCFA803" box="[176,228,1118,1141]" italics="true" pageId="24" pageNumber="25">AEW</emphasis>
i (%)
</th>
<td id="76845DB7F94A62B3FEB820678DD6A800" box="[332,509,1117,1142]" gridcol="1" gridrow="23" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE20678EA5A800" box="[538,654,1117,1142]" gridcol="2" gridrow="23" pageId="24" pageNumber="25">43.0 ± 3.0</td>
<td id="76845DB7F94A62B3FD5E20678F0BA800" box="[682,800,1117,1142]" gridcol="3" gridrow="23" pageId="24" pageNumber="25">(40.247.4)</td>
<td id="76845DB7F94A62B3FCC820678F99A800" box="[828,946,1117,1142]" gridcol="4" gridrow="23" pageId="24" pageNumber="25">48.1 ± 3.1</td>
<td id="76845DB7F94A62B3FC302067887FA800" box="[964,1108,1117,1142]" gridcol="5" gridrow="23" pageId="24" pageNumber="25">(43.851.1)</td>
<td id="76845DB7F94A62B3FB8F20678922A800" box="[1147,1289,1117,1142]" gridcol="6" gridrow="23" pageId="24" pageNumber="25">52.6 ± 5.6</td>
<td id="76845DB7F94A62B3FAEF206789BAA800" box="[1307,1425,1117,1142]" gridcol="7" gridrow="23" pageId="24" pageNumber="25">(45.262.1)</td>
</tr>
<tr id="355534CBF94A62B3FF5720BC89BAA8E8" box="[163,1425,1158,1182]" gridrow="24" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720BC89BAA8E8" box="[163,1425,1158,1182]" colspan="8" colspanRight="7" gridcol="0" gridrow="24" pageId="24" pageNumber="25">Dorsal side</th>
</tr>
<tr id="355534CBF94A62B3FF57209489BAA8B0" box="[163,1425,1198,1222]" gridrow="25" pageId="24" pageNumber="25" rowspan-4="1">
<th id="76845DB7F94A62B3FF5720948D32A8B0" box="[163,281,1198,1222]" gridcol="0" gridrow="25" pageId="24" pageNumber="25">L-Muscle</th>
<td id="76845DB7F94A62B3FEB820948DD6A8B0" box="[332,509,1198,1222]" gridcol="1" gridrow="25" pageId="24" pageNumber="25">
Granules (
<emphasis id="B911EA9BF94A9D54FE3B20958DF7A8B0" box="[463,476,1199,1222]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE20948F0BA8B0" box="[538,800,1198,1222]" colspan="2" colspanRight="1" gridcol="2" gridrow="25" pageId="24" pageNumber="25">9</td>
<td id="76845DB7F94A62B3FC302094887FA8B0" box="[964,1108,1198,1222]" gridcol="5" gridrow="25" pageId="24" pageNumber="25">9</td>
<td id="76845DB7F94A62B3FB8F209489BAA8B0" box="[1147,1425,1198,1222]" colspan="2" colspanRight="1" gridcol="6" gridrow="25" pageId="24" pageNumber="25">12</td>
</tr>
<tr id="355534CBF94A62B3FF5720EC89BAA899" box="[163,1425,1238,1263]" gridrow="26" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720EC8D32A899" box="[163,281,1238,1263]" gridcol="0" gridrow="26" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB820EC8DD6A899" box="[332,509,1238,1263]" gridcol="1" gridrow="26" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE20EC8EA5A899" box="[538,654,1238,1263]" gridcol="2" gridrow="26" pageId="24" pageNumber="25">30.6 ± 2.5</td>
<td id="76845DB7F94A62B3FD5E20EC8F0BA899" box="[682,800,1238,1263]" gridcol="3" gridrow="26" pageId="24" pageNumber="25">(2834)</td>
<td id="76845DB7F94A62B3FCC820EC8F99A899" box="[828,946,1238,1263]" gridcol="4" gridrow="26" pageId="24" pageNumber="25">29.3 ± 2.5</td>
<td id="76845DB7F94A62B3FC3020EC887FA899" box="[964,1108,1238,1263]" gridcol="5" gridrow="26" pageId="24" pageNumber="25">(2533)</td>
<td id="76845DB7F94A62B3FB8F20EC8922A899" box="[1147,1289,1238,1263]" gridcol="6" gridrow="26" pageId="24" pageNumber="25">32.8 ± 2.1</td>
<td id="76845DB7F94A62B3FAEF20EC89BAA899" box="[1307,1425,1238,1263]" gridcol="7" gridrow="26" pageId="24" pageNumber="25">(2936)</td>
</tr>
<tr id="355534CBF94A62B3FF5720C489BAA961" box="[163,1425,1278,1303]" gridrow="27" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720C48D32A961" box="[163,281,1278,1303]" gridcol="0" gridrow="27" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB820C48DD6A961" box="[332,509,1278,1303]" gridcol="1" gridrow="27" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE20C48EA5A961" box="[538,654,1278,1303]" gridcol="2" gridrow="27" pageId="24" pageNumber="25">13.4 ± 3.0</td>
<td id="76845DB7F94A62B3FD5E20C48F0BA961" box="[682,800,1278,1303]" gridcol="3" gridrow="27" pageId="24" pageNumber="25">(718)</td>
<td id="76845DB7F94A62B3FCC820C48F99A961" box="[828,946,1278,1303]" gridcol="4" gridrow="27" pageId="24" pageNumber="25">15.4 ± 3.5</td>
<td id="76845DB7F94A62B3FC3020C4887FA961" box="[964,1108,1278,1303]" gridcol="5" gridrow="27" pageId="24" pageNumber="25">(1019)</td>
<td id="76845DB7F94A62B3FB8F20C48922A961" box="[1147,1289,1278,1303]" gridcol="6" gridrow="27" pageId="24" pageNumber="25">15.0 ± 1.5</td>
<td id="76845DB7F94A62B3FAEF20C489BAA961" box="[1307,1425,1278,1303]" gridcol="7" gridrow="27" pageId="24" pageNumber="25">(1318)</td>
</tr>
<tr id="355534CBF94A62B3FF57211C89BAA949" box="[163,1425,1318,1343]" gridrow="28" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57211C8D32A949" box="[163,281,1318,1343]" gridcol="0" gridrow="28" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44211D8CF9A948" box="[176,210,1319,1342]" italics="true" pageId="24" pageNumber="25">GP</emphasis>
c
</th>
<td id="76845DB7F94A62B3FEB8211C8DD6A949" box="[332,509,1318,1343]" gridcol="1" gridrow="28" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE211C8EA5A949" box="[538,654,1318,1343]" gridcol="2" gridrow="28" pageId="24" pageNumber="25">44.2 ± 10.9</td>
<td id="76845DB7F94A62B3FD5E211C8F0BA949" box="[682,800,1318,1343]" gridcol="3" gridrow="28" pageId="24" pageNumber="25">(25.062.1)</td>
<td id="76845DB7F94A62B3FCC8211C887FA949" box="[828,1108,1318,1343]" colspan="2" colspanRight="1" gridcol="4" gridrow="28" pageId="24" pageNumber="25">52.9 ± 12.5 (31.266.7)</td>
<td id="76845DB7F94A62B3FB8F211C8922A949" box="[1147,1289,1318,1343]" gridcol="6" gridrow="28" pageId="24" pageNumber="25">45.9 ± 6.1</td>
<td id="76845DB7F94A62B3FAEF211C89BAA949" box="[1307,1425,1318,1343]" gridcol="7" gridrow="28" pageId="24" pageNumber="25">(37.156.3)</td>
</tr>
<tr id="355534CBF94A62B3FF57217589BAA911" box="[163,1425,1359,1383]" gridrow="29" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5721758D32A911" box="[163,281,1359,1383]" gridcol="0" gridrow="29" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44216A8CCEA911" box="[176,229,1360,1383]" italics="true" pageId="24" pageNumber="25">GCO</emphasis>
d (%)
</th>
<td id="76845DB7F94A62B3FEB821758DD6A911" box="[332,509,1359,1383]" gridcol="1" gridrow="29" pageId="24" pageNumber="25">(Ratio)</td>
<td id="76845DB7F94A62B3FDEE21758EA5A911" box="[538,654,1359,1383]" gridcol="2" gridrow="29" pageId="24" pageNumber="25">11.1</td>
<td id="76845DB7F94A62B3FD5E21758F0BA911" box="[682,800,1359,1383]" gridcol="3" gridrow="29" pageId="24" pageNumber="25">(O1: C8)</td>
<td id="76845DB7F94A62B3FCC821758F99A911" box="[828,946,1359,1383]" gridcol="4" gridrow="29" pageId="24" pageNumber="25">77.8</td>
<td id="76845DB7F94A62B3FC302175887FA911" box="[964,1108,1359,1383]" gridcol="5" gridrow="29" pageId="24" pageNumber="25">(O7: C2)</td>
<td id="76845DB7F94A62B3FB8F21758922A911" box="[1147,1289,1359,1383]" gridcol="6" gridrow="29" pageId="24" pageNumber="25">33.3</td>
<td id="76845DB7F94A62B3FAEF217589BAA911" box="[1307,1425,1359,1383]" gridcol="7" gridrow="29" pageId="24" pageNumber="25">(O4: C8)</td>
</tr>
<tr id="355534CBF94A62B3FF57214D89BAA9E6" box="[163,1425,1399,1424]" gridrow="30" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF57214D8D32A9E6" box="[163,281,1399,1424]" gridcol="0" gridrow="30" pageId="24" pageNumber="25">Body wall</th>
<td id="76845DB7F94A62B3FEB8214D8DD6A9E6" box="[332,509,1399,1424]" gridcol="1" gridrow="30" pageId="24" pageNumber="25">
Anchor plates (
<emphasis id="B911EA9BF94A9D54FE1C21428DDEA9F9" box="[488,501,1400,1423]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE214D8F0BA9E6" box="[538,800,1399,1424]" colspan="2" colspanRight="1" gridcol="2" gridrow="30" pageId="24" pageNumber="25">10</td>
<td id="76845DB7F94A62B3FC30214D887FA9E6" box="[964,1108,1399,1424]" gridcol="5" gridrow="30" pageId="24" pageNumber="25">8</td>
<td id="76845DB7F94A62B3FB8F214D8922A9E6" box="[1147,1289,1399,1424]" gridcol="6" gridrow="30" pageId="24" pageNumber="25">6</td>
</tr>
<tr id="355534CBF94A62B3FF57219A89BAA9CE" box="[163,1425,1440,1464]" gridrow="31" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57219A8D32A9CE" box="[163,281,1440,1464]" gridcol="0" gridrow="31" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB8219A8DD6A9CE" box="[332,509,1440,1464]" gridcol="1" gridrow="31" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE219A8EA5A9CE" box="[538,654,1440,1464]" gridcol="2" gridrow="31" pageId="24" pageNumber="25">126.1 ± 9.4</td>
<td id="76845DB7F94A62B3FD5E219A8F0BA9CE" box="[682,800,1440,1464]" gridcol="3" gridrow="31" pageId="24" pageNumber="25">(109135)</td>
<td id="76845DB7F94A62B3FCC8219A887FA9CE" box="[828,1108,1440,1464]" colspan="2" colspanRight="1" gridcol="4" gridrow="31" pageId="24" pageNumber="25">109.6 ± 8.6 (98128)</td>
<td id="76845DB7F94A62B3FB8F219A8922A9CE" box="[1147,1289,1440,1464]" gridcol="6" gridrow="31" pageId="24" pageNumber="25">99.1 ± 7.3</td>
<td id="76845DB7F94A62B3FAEF219A89BAA9CE" box="[1307,1425,1440,1464]" gridcol="7" gridrow="31" pageId="24" pageNumber="25">(91108)</td>
</tr>
<tr id="355534CBF94A62B3FF5721F289BAA996" box="[163,1425,1480,1504]" gridrow="32" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5721F28D32A996" box="[163,281,1480,1504]" gridcol="0" gridrow="32" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB821F28DD6A996" box="[332,509,1480,1504]" gridcol="1" gridrow="32" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE21F28EA5A996" box="[538,654,1480,1504]" gridcol="2" gridrow="32" pageId="24" pageNumber="25">94.1 ± 9.4</td>
<td id="76845DB7F94A62B3FD5E21F28F0BA996" box="[682,800,1480,1504]" gridcol="3" gridrow="32" pageId="24" pageNumber="25">(76106)</td>
<td id="76845DB7F94A62B3FCC821F28F99A996" box="[828,946,1480,1504]" gridcol="4" gridrow="32" pageId="24" pageNumber="25">67.5 ± 7.6</td>
<td id="76845DB7F94A62B3FC3021F2887FA996" box="[964,1108,1480,1504]" gridcol="5" gridrow="32" pageId="24" pageNumber="25">(5580)</td>
<td id="76845DB7F94A62B3FB8F21F28922A996" box="[1147,1289,1480,1504]" gridcol="6" gridrow="32" pageId="24" pageNumber="25">66.8 ±11.5</td>
<td id="76845DB7F94A62B3FAEF21F289BAA996" box="[1307,1425,1480,1504]" gridcol="7" gridrow="32" pageId="24" pageNumber="25">(5379)</td>
</tr>
<tr id="355534CBF94A62B3FF5721D589BAAA7E" box="[163,1425,1519,1544]" gridrow="33" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5721D58D32AA7E" box="[163,281,1519,1544]" gridcol="0" gridrow="33" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4421CB8CF7AA7E" box="[176,220,1521,1544]" italics="true" pageId="24" pageNumber="25">APS</emphasis>
e
</th>
<td id="76845DB7F94A62B3FEB821D58DD6AA7E" box="[332,509,1519,1544]" gridcol="1" gridrow="33" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE21D58EA5AA7E" box="[538,654,1519,1544]" gridcol="2" gridrow="33" pageId="24" pageNumber="25">14.9 ±15.1</td>
<td id="76845DB7F94A62B3FD5E21D58F0BAA7E" box="[682,800,1519,1544]" gridcol="3" gridrow="33" pageId="24" pageNumber="25">(2.743.0)</td>
<td id="76845DB7F94A62B3FCC821D58F99AA7E" box="[828,946,1519,1544]" gridcol="4" gridrow="33" pageId="24" pageNumber="25">8.9 ± 6.6</td>
<td id="76845DB7F94A62B3FC3021D5887FAA7E" box="[964,1108,1519,1544]" gridcol="5" gridrow="33" pageId="24" pageNumber="25">(4.522.4)</td>
<td id="76845DB7F94A62B3FB8F21D58922AA7E" box="[1147,1289,1519,1544]" gridcol="6" gridrow="33" pageId="24" pageNumber="25">4.1 ± 4.4</td>
<td id="76845DB7F94A62B3FAEF21D589BAAA7E" box="[1307,1425,1519,1544]" gridcol="7" gridrow="33" pageId="24" pageNumber="25">(0.612.5)</td>
</tr>
<tr id="355534CBF94A62B3FF57222289BAAA47" box="[163,1425,1560,1585]" gridrow="34" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5722228D32AA47" box="[163,281,1560,1585]" gridcol="0" gridrow="34" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4422238CFEAA46" box="[176,213,1561,1584]" italics="true" pageId="24" pageNumber="25">Ntp</emphasis>
f
</th>
<td id="76845DB7F94A62B3FEB822228DD6AA47" box="[332,509,1560,1585]" gridcol="1" gridrow="34" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE22228EA5AA47" box="[538,654,1560,1585]" gridcol="2" gridrow="34" pageId="24" pageNumber="25">3.5 ± 1.1</td>
<td id="76845DB7F94A62B3FD5E22228F0BAA47" box="[682,800,1560,1585]" gridcol="3" gridrow="34" pageId="24" pageNumber="25">(26)</td>
<td id="76845DB7F94A62B3FCC822228F99AA47" box="[828,946,1560,1585]" gridcol="4" gridrow="34" pageId="24" pageNumber="25">2.9 ± 1.2</td>
<td id="76845DB7F94A62B3FC302222887FAA47" box="[964,1108,1560,1585]" gridcol="5" gridrow="34" pageId="24" pageNumber="25">(15)</td>
<td id="76845DB7F94A62B3FB8F22228922AA47" box="[1147,1289,1560,1585]" gridcol="6" gridrow="34" pageId="24" pageNumber="25">3.2 ± 0.8</td>
<td id="76845DB7F94A62B3FAEF222289BAAA47" box="[1307,1425,1560,1585]" gridcol="7" gridrow="34" pageId="24" pageNumber="25">(24)</td>
</tr>
<tr id="355534CBF94A62B3FF57227A89BAAA2F" box="[163,1425,1600,1625]" gridrow="35" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57227A8D32AA2F" box="[163,281,1600,1625]" gridcol="0" gridrow="35" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44227B8CF6AA2E" box="[176,221,1601,1624]" italics="true" pageId="24" pageNumber="25">APT</emphasis>
g (%)
</th>
<td id="76845DB7F94A62B3FEB8227A8DD6AA2F" box="[332,509,1600,1625]" gridcol="1" gridrow="35" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE227A8EA5AA2F" box="[538,654,1600,1625]" gridcol="2" gridrow="35" pageId="24" pageNumber="25">10.5±2.4</td>
<td id="76845DB7F94A62B3FD5E227A8F0BAA2F" box="[682,800,1600,1625]" gridcol="3" gridrow="35" pageId="24" pageNumber="25">(6.914.6)</td>
<td id="76845DB7F94A62B3FCC8227A8F99AA2F" box="[828,946,1600,1625]" gridcol="4" gridrow="35" pageId="24" pageNumber="25">10.4 ± 5.7</td>
<td id="76845DB7F94A62B3FC30227A887FAA2F" box="[964,1108,1600,1625]" gridcol="5" gridrow="35" pageId="24" pageNumber="25">(2.820.8)</td>
<td id="76845DB7F94A62B3FB8F227A8922AA2F" box="[1147,1289,1600,1625]" gridcol="6" gridrow="35" pageId="24" pageNumber="25">17.4 ± 4.1</td>
<td id="76845DB7F94A62B3FAEF227A89BAAA2F" box="[1307,1425,1600,1625]" gridcol="7" gridrow="35" pageId="24" pageNumber="25">(14.325.0)</td>
</tr>
<tr id="355534CBF94A62B3FF57225389BAAAF7" box="[163,1425,1641,1665]" gridrow="36" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF5722538D32AAF7" box="[163,281,1641,1665]" gridcol="0" gridrow="36" pageId="24" pageNumber="25">Body wall</th>
<td id="76845DB7F94A62B3FEB822538DD6AAF7" box="[332,509,1641,1665]" gridcol="1" gridrow="36" pageId="24" pageNumber="25">
Anchor (
<emphasis id="B911EA9BF94A9D54FE3322508DFFAAF7" box="[455,468,1642,1665]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE22538F0BAAF7" box="[538,800,1641,1665]" colspan="2" colspanRight="1" gridcol="2" gridrow="36" pageId="24" pageNumber="25">9</td>
<td id="76845DB7F94A62B3FC302253887FAAF7" box="[964,1108,1641,1665]" gridcol="5" gridrow="36" pageId="24" pageNumber="25">8</td>
<td id="76845DB7F94A62B3FB8F22538922AAF7" box="[1147,1289,1641,1665]" gridcol="6" gridrow="36" pageId="24" pageNumber="25">6</td>
</tr>
<tr id="355534CBF94A62B3FF5722AB89BAAADF" box="[163,1425,1681,1705]" gridrow="37" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5722AB8D32AADF" box="[163,281,1681,1705]" gridcol="0" gridrow="37" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB822AB8DD6AADF" box="[332,509,1681,1705]" gridcol="1" gridrow="37" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE22AB8EA5AADF" box="[538,654,1681,1705]" gridcol="2" gridrow="37" pageId="24" pageNumber="25">165.6 ± 9.3</td>
<td id="76845DB7F94A62B3FD5E22AB8F0BAADF" box="[682,800,1681,1705]" gridcol="3" gridrow="37" pageId="24" pageNumber="25">(151178)</td>
<td id="76845DB7F94A62B3FCC822AB887FAADF" box="[828,1108,1681,1705]" colspan="2" colspanRight="1" gridcol="4" gridrow="37" pageId="24" pageNumber="25">168.5 ± 10.3 (153178)</td>
<td id="76845DB7F94A62B3FB8F22AB8922AADF" box="[1147,1289,1681,1705]" gridcol="6" gridrow="37" pageId="24" pageNumber="25">160.5 ± 14.6</td>
<td id="76845DB7F94A62B3FAEF22AB89BAAADF" box="[1307,1425,1681,1705]" gridcol="7" gridrow="37" pageId="24" pageNumber="25">(144181)</td>
</tr>
<tr id="355534CBF94A62B3FF57228389BAAAA4" box="[163,1425,1721,1746]" gridrow="38" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5722838D32AAA4" box="[163,281,1721,1746]" gridcol="0" gridrow="38" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB822838DD6AAA4" box="[332,509,1721,1746]" gridcol="1" gridrow="38" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE22838EA5AAA4" box="[538,654,1721,1746]" gridcol="2" gridrow="38" pageId="24" pageNumber="25">86.4 ± 6.7</td>
<td id="76845DB7F94A62B3FD5E22838F0BAAA4" box="[682,800,1721,1746]" gridcol="3" gridrow="38" pageId="24" pageNumber="25">(7495)</td>
<td id="76845DB7F94A62B3FCC822838F99AAA4" box="[828,946,1721,1746]" gridcol="4" gridrow="38" pageId="24" pageNumber="25">90.9 ± 6.9</td>
<td id="76845DB7F94A62B3FC302283887FAAA4" box="[964,1108,1721,1746]" gridcol="5" gridrow="38" pageId="24" pageNumber="25">(81103)</td>
<td id="76845DB7F94A62B3FB8F22838922AAA4" box="[1147,1289,1721,1746]" gridcol="6" gridrow="38" pageId="24" pageNumber="25">67.2 ± 11.2</td>
<td id="76845DB7F94A62B3FAEF228389BAAAA4" box="[1307,1425,1721,1746]" gridcol="7" gridrow="38" pageId="24" pageNumber="25">(5587)</td>
</tr>
<tr id="355534CBF94A62B3FF5722D889BAAA8C" box="[163,1425,1762,1786]" gridrow="39" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5722D88D32AA8C" box="[163,281,1762,1786]" gridcol="0" gridrow="39" pageId="24" pageNumber="25">Teeth</th>
<td id="76845DB7F94A62B3FEB822D88DD6AA8C" box="[332,509,1762,1786]" gridcol="1" gridrow="39" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE22D88EA5AA8C" box="[538,654,1762,1786]" gridcol="2" gridrow="39" pageId="24" pageNumber="25">2.6±1.9</td>
<td id="76845DB7F94A62B3FD5E22D88F0BAA8C" box="[682,800,1762,1786]" gridcol="3" gridrow="39" pageId="24" pageNumber="25">(24)</td>
<td id="76845DB7F94A62B3FCC822D88F99AA8C" box="[828,946,1762,1786]" gridcol="4" gridrow="39" pageId="24" pageNumber="25">3.4±0.7</td>
<td id="76845DB7F94A62B3FC3022D8887FAA8C" box="[964,1108,1762,1786]" gridcol="5" gridrow="39" pageId="24" pageNumber="25">(24)</td>
<td id="76845DB7F94A62B3FB8F22D88922AA8C" box="[1147,1289,1762,1786]" gridcol="6" gridrow="39" pageId="24" pageNumber="25">6.2 ± 1.6</td>
<td id="76845DB7F94A62B3FAEF22D889BAAA8C" box="[1307,1425,1762,1786]" gridcol="7" gridrow="39" pageId="24" pageNumber="25">(48)</td>
</tr>
<tr id="355534CBF94A62B3FF57233089BAAB54" box="[163,1425,1802,1826]" gridrow="40" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5723308D32AB54" box="[163,281,1802,1826]" gridcol="0" gridrow="40" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4423308CCAAB57" box="[176,225,1802,1825]" italics="true" pageId="24" pageNumber="25">ASW</emphasis>
h(%)
</th>
<td id="76845DB7F94A62B3FEB823308DD6AB54" box="[332,509,1802,1826]" gridcol="1" gridrow="40" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE23308EA5AB54" box="[538,654,1802,1826]" gridcol="2" gridrow="40" pageId="24" pageNumber="25">22.4 ±1.6</td>
<td id="76845DB7F94A62B3FD5E23308F0BAB54" box="[682,800,1802,1826]" gridcol="3" gridrow="40" pageId="24" pageNumber="25">(19.725.0)</td>
<td id="76845DB7F94A62B3FCC823308F99AB54" box="[828,946,1802,1826]" gridcol="4" gridrow="40" pageId="24" pageNumber="25">20.3 ± 0.8</td>
<td id="76845DB7F94A62B3FC302330887FAB54" box="[964,1108,1802,1826]" gridcol="5" gridrow="40" pageId="24" pageNumber="25">(19.121.3)</td>
<td id="76845DB7F94A62B3FB8F23308922AB54" box="[1147,1289,1802,1826]" gridcol="6" gridrow="40" pageId="24" pageNumber="25">25.8 ± 3.5</td>
<td id="76845DB7F94A62B3FAEF233089BAAB54" box="[1307,1425,1802,1826]" gridcol="7" gridrow="40" pageId="24" pageNumber="25">(20.730.9)</td>
</tr>
<tr id="355534CBF94A62B3FF57230889BAAB3C" box="[163,1425,1842,1866]" gridrow="41" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5723088D32AB3C" box="[163,281,1842,1866]" gridcol="0" gridrow="41" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4423098CCFAB3C" box="[176,228,1843,1866]" italics="true" pageId="24" pageNumber="25">AEW</emphasis>
i (%)
</th>
<td id="76845DB7F94A62B3FEB823088DD6AB3C" box="[332,509,1842,1866]" gridcol="1" gridrow="41" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE23088EA5AB3C" box="[538,654,1842,1866]" gridcol="2" gridrow="41" pageId="24" pageNumber="25">42.6 ± 3.1</td>
<td id="76845DB7F94A62B3FD5E23088F0BAB3C" box="[682,800,1842,1866]" gridcol="3" gridrow="41" pageId="24" pageNumber="25">(38.247.7)</td>
<td id="76845DB7F94A62B3FCC823088F99AB3C" box="[828,946,1842,1866]" gridcol="4" gridrow="41" pageId="24" pageNumber="25">44.9 ± 2.8</td>
<td id="76845DB7F94A62B3FC302308887FAB3C" box="[964,1108,1842,1866]" gridcol="5" gridrow="41" pageId="24" pageNumber="25">(40.848.3)</td>
<td id="76845DB7F94A62B3FB8F23088922AB3C" box="[1147,1289,1842,1866]" gridcol="6" gridrow="41" pageId="24" pageNumber="25">53.2 ± 7.4</td>
<td id="76845DB7F94A62B3FAEF230889BAAB3C" box="[1307,1425,1842,1866]" gridcol="7" gridrow="41" pageId="24" pageNumber="25">(43.763.8)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDA3689F94A9D54FF63236289A1AB06" blockId="24.[151,1418,1880,1904]" box="[151,1418,1880,1904]" pageId="24" pageNumber="25">
<tableNote id="76833707F94A9D54FF63236289A1AB06" box="[151,1418,1880,1904]" pageId="24" pageNumber="25" targetBox="[163,1425,192,1866]" targetPageId="24">
ai, (see, Materials and Methods) Indicators for ossicles shape, and one parameter (
<emphasis id="B911EA9BF94A9D54FBE623628812AB19" box="[1042,1081,1880,1903]" italics="true" pageId="24" pageNumber="25">Ntp</emphasis>
, number s teethed perforation).
</tableNote>
</paragraph>
<caption id="DF1A6601F94B9D55FF6322998EA7AB51" ID-DOI="http://doi.org/10.5281/zenodo.14703560" ID-Zenodo-Dep="14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1699,1723]" targetBox="[151,1436,181,1667]" targetPageId="25" targetType="figure">
<paragraph id="8BDA3689F94B9D55FF6322998EA7AB51" blockId="25.[151,1437,1699,1831]" pageId="25" pageNumber="26">
<emphasis id="B911EA9BF94B9D55FF6322998D0BAACD" bold="true" box="[151,288,1699,1723]" pageId="25" pageNumber="26">FIGURE 13.</emphasis>
SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as
<taxonomicName id="4C654D0AF94B9D55FF4722F28D5DAAA9" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[179,374,1735,1759]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="25" pageNumber="26" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF94B9D55FF4722F28D5DAAA9" box="[179,374,1735,1759]" italics="true" pageId="25" pageNumber="26">Patinapta deformis</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF94B9D55FE8822FD8DE7AAA9" bold="true" box="[380,460,1735,1759]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A22257E0F94B9D55FE8822FD8DE7AAA9" box="[380,460,1735,1759]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle.
</paragraph>
</caption>
<caption id="DF1A6601F9489D56FF63263E895FAEDA" ID-DOI="http://doi.org/10.5281/zenodo.14703562" ID-Zenodo-Dep="14703562" httpUri="https://zenodo.org/record/14703562/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,516,540]" targetBox="[164,1423,188,485]" targetPageId="26" targetType="figure">
<paragraph id="8BDA3689F9489D56FF63263E895FAEDA" blockId="26.[151,1437,516,685]" pageId="26" pageNumber="27">
<emphasis id="B911EA9BF9489D56FF63263E8D35AE6B" bold="true" box="[151,286,516,541]" pageId="26" pageNumber="27">FIGURE 14.</emphasis>
Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima (holotype specimen, WMNH-2015-INV-106), described as
<taxonomicName id="4C654D0AF9489D56FD7A26138F78AE37" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[654,851,553,577]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="26" pageNumber="27" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF9489D56FD7A26138F78AE37" box="[654,851,553,577]" italics="true" pageId="26" pageNumber="27">Patinapta deformis</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9489D56FCAE26138F80AE37" bold="true" box="[858,939,553,577]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A22257E0F9489D56FCAE26138F80AE37" box="[858,939,553,577]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study. The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system:
<emphasis id="B911EA9BF9489D56FCF626778F6FAE12" box="[770,836,589,612]" italics="true" pageId="26" pageNumber="27">RIRV</emphasis>
and
<emphasis id="B911EA9BF9489D56FC8226778FE7AE12" box="[886,972,589,613]" italics="true" pageId="26" pageNumber="27">IR1IR5</emphasis>
where
<emphasis id="B911EA9BF9489D56FBE32677880DAE12" box="[1047,1062,589,612]" italics="true" pageId="26" pageNumber="27">R</emphasis>
and
<emphasis id="B911EA9BF9489D56FBAD2677885AAE12" box="[1113,1137,589,612]" italics="true" pageId="26" pageNumber="27">IR</emphasis>
stand for positions for radius and interradius, respectively. Hence
<emphasis id="B911EA9BF9489D56FDFD264B8E0AAEFE" box="[521,545,625,648]" italics="true" pageId="26" pageNumber="27">RI</emphasis>
, mid-ventral;
<emphasis id="B911EA9BF9489D56FD45264B8EFAAEFE" box="[689,721,625,648]" italics="true" pageId="26" pageNumber="27">RII</emphasis>
, left ventral;
<emphasis id="B911EA9BF9489D56FCAF264B8FAFAEFE" box="[859,900,625,648]" italics="true" pageId="26" pageNumber="27">RIII</emphasis>
, right ventral;
<emphasis id="B911EA9BF9489D56FBEF264B8869AEFE" box="[1051,1090,625,648]" italics="true" pageId="26" pageNumber="27">RIV</emphasis>
, left dorsal;
<emphasis id="B911EA9BF9489D56FB37264B88CAAEFE" box="[1219,1249,625,648]" italics="true" pageId="26" pageNumber="27">RV</emphasis>
, right dorsal; and:
<emphasis id="B911EA9BF9489D56FF6326AF8C90AEDB" box="[151,187,661,685]" italics="true" pageId="26" pageNumber="27">IR1</emphasis>
, left ventral;
<emphasis id="B911EA9BF9489D56FEB026AF8D42AEDB" box="[324,361,661,685]" italics="true" pageId="26" pageNumber="27">IR2</emphasis>
, right ventral;
<emphasis id="B911EA9BF9489D56FE0B26AF8E72AEDB" box="[511,601,661,685]" italics="true" pageId="26" pageNumber="27">IR3-IR3</emphasis>
, left dorso-lateral;
<emphasis id="B911EA9BF9489D56FCE826AF8F5DAEDB" box="[796,886,661,685]" italics="true" pageId="26" pageNumber="27">IR4-IR4</emphasis>
, right dorso-lateral;
<emphasis id="B911EA9BF9489D56FBB226AF8841AEDA" box="[1094,1130,661,684]" italics="true" pageId="26" pageNumber="27">IR5</emphasis>
, mid-dorsal. Anterior top.
</paragraph>
</caption>
<paragraph id="8BDA3689F9489D56FF3326C58F4BAFBB" blockId="26.[151,1437,767,1190]" pageId="26" pageNumber="27">
Anchor ossicles ranged within 144178 μm in length (
<figureCitation id="135E2A0CF9489D56FCED26C58F88AF6F" box="[793,931,767,793]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13 B, D</figureCitation>
;
<tableCitation id="C6E70332F9489D56FC5826C58FD4AF6F" box="[940,1023,767,793]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="26" pageNumber="27">Table 6</tableCitation>
). All the anchor arms (bills) equipped with minute teeth on their outer tips, numbers of teeth variable within 27 (mostly
<quantity id="4C9D9B6CF9489D56FBBB271988BCAF4B" box="[1103,1175,803,829]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="26" pageNumber="27" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">13 in</quantity>
one arm of the anchor ossicle). Anchor ossicles of this specimen had broad stem and relatively wide anchor arms, with
<emphasis id="B911EA9BF9489D56FB33277D88D6AF17" box="[1223,1277,839,865]" italics="true" pageId="26" pageNumber="27">ASW</emphasis>
=19.121.3%;
<emphasis id="B911EA9BF9489D56FF6327518CFAAFF3" box="[151,209,875,901]" italics="true" pageId="26" pageNumber="27">AEW</emphasis>
=40.851.1%, and T-shaped basal end of the anchor ossicle of this species strongly widened and semicircular in shape (
<figureCitation id="135E2A0CF9489D56FEF527B58DA3AFDF" box="[257,392,911,937]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13B, D</figureCitation>
). Such features were have not been reported in both previously described congeneric Japanese species (both species possess crescent-like T-shaped basal end).
</paragraph>
<paragraph id="8BDA3689F9489D56FF3327ED88A9A82B" blockId="26.[151,1437,767,1190]" pageId="26" pageNumber="27">
Body longitudinal muscle, granule ossicles of this specimen ranged within 2336 μm in length (
<figureCitation id="135E2A0CF9489D56FAFB27ED89BCAF87" box="[1295,1431,983,1009]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13C, E</figureCitation>
;
<tableCitation id="C6E70332F9489D56FF6327C18CC0A863" box="[151,235,1019,1045]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="26" pageNumber="27">Table 6</tableCitation>
). Calculated indicator for granule proportions was variated within
<emphasis id="B911EA9BF9489D56FC2527C18FDDA863" box="[977,1014,1019,1045]" italics="true" pageId="26" pageNumber="27">GP</emphasis>
=31.282.6%;
<emphasis id="B911EA9BF9489D56FB6F27C188FDA863" box="[1179,1238,1019,1045]" italics="true" pageId="26" pageNumber="27">GCO</emphasis>
=44.477.8% and were not noticeably unbalanced between O-shaped or C-shaped granules, however some granule ossicles had one or two minute perforations, resulting in turtle-face-like characteristic shape (
<figureCitation id="135E2A0CF9489D56FC1B2079885EA82B" box="[1007,1141,1091,1117]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13C, E</figureCitation>
).
</paragraph>
<paragraph id="8BDA3689F9489D56FF33205D8815A8D0" blockId="26.[151,1437,767,1190]" pageId="26" pageNumber="27">
Tentacle ossicles rods, ranged within 5285 μm in length (
<figureCitation id="135E2A0CF9489D56FCA5205D8F9CA8F7" box="[849,951,1127,1153]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13A</figureCitation>
;
<tableCitation id="C6E70332F9489D56FC37205D883CA8F7" box="[963,1047,1127,1153]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="26" pageNumber="27">Table 6</tableCitation>
). Two calculated indicators for rod proportions and complexities were variated within
<emphasis id="B911EA9BF9489D56FD2620B68EDFA8D3" box="[722,756,1164,1189]" italics="true" pageId="26" pageNumber="27">RP</emphasis>
=10.720.3%;
<emphasis id="B911EA9BF9489D56FC5520B18FEEA8D3" box="[929,965,1163,1189]" italics="true" pageId="26" pageNumber="27">RC</emphasis>
=5.112.2.
</paragraph>
<paragraph id="8BDA3689F9489D56FF6320CD8F20A967" blockId="26.[151,779,1271,1297]" box="[151,779,1271,1297]" pageId="26" pageNumber="27">
<heading id="D09281E5F9489D56FF6320CD8F20A967" bold="true" box="[151,779,1271,1297]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B911EA9BF9489D56FF6320CD8F20A967" bold="true" box="[151,779,1271,1297]" pageId="26" pageNumber="27">Observation on the morphology of calcareous ring</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9489D56FF6321058824A9B3" blockId="26.[151,1436,1343,1477]" pageId="26" pageNumber="27">
Calcareous plates 12, firmly bound (
<figureCitation id="135E2A0CF9489D56FDC921058EB9A92F" box="[573,658,1343,1369]" captionStart="FIGURE 14" captionStartId="26.[151,250,516,540]" captionTargetBox="[164,1423,188,485]" captionTargetId="figure-29@26.[151,1435,181,493]" captionTargetPageId="26" captionText="FIGURE 14. Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima (holotype specimen, WMNH-2015-INV-106), described as Patinapta deformis sp. nov., in the present study. The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stand for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi="http://doi.org/10.5281/zenodo.14703562" httpUri="https://zenodo.org/record/14703562/files/figure.png" pageId="26" pageNumber="27">Fig. 14</figureCitation>
), with slight anterior projection and slight posterior depression. All radial plates perforated in each anterior half. Two ventro-lateral plates (
<emphasis id="B911EA9BF9489D56FC36215E880EA90B" box="[962,1061,1380,1405]" italics="true" pageId="26" pageNumber="27">RII; RIII</emphasis>
) protruding strongly, and exhibit pentagonal shape, resulting in both suddenly turning sharply on these ventro-lateral plates. There is no space between the additional interradial plate (
<emphasis id="B911EA9BF9489D56FE0C21918E47A9B3" box="[504,620,1451,1477]" italics="true" pageId="26" pageNumber="27">IR4; IR3</emphasis>
) and adjacent radial plate (
<emphasis id="B911EA9BF9489D56FC6921968FD4A9B3" box="[925,1023,1452,1477]" italics="true" pageId="26" pageNumber="27">RIII; RII</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9489D57FF63222D8878AFBF" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="8BDA3689F9489D56FF63222D8EC6AA47" blockId="26.[151,749,1559,1585]" box="[151,749,1559,1585]" pageId="26" pageNumber="27">
<heading id="D09281E5F9489D56FF63222D8EC6AA47" bold="true" box="[151,749,1559,1585]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B911EA9BF9489D56FF63222D8EC6AA47" bold="true" box="[151,749,1559,1585]" pageId="26" pageNumber="27">Variation of external and internal morphologies</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9489D56FF6322658E37AAB7" blockId="26.[151,1436,1631,1729]" pageId="26" pageNumber="27">
<specimenCount id="9D63FD00F9489D56FF6322658D7DAA0F" box="[151,342,1631,1657]" count="3" pageId="26" pageNumber="27" type="generic">Three specimens</specimenCount>
were all large-size specimens of about
<quantity id="4C9D9B6CF9489D56FCE822658F8EAA0F" box="[796,933,1631,1657]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="26" pageNumber="27" unit="mm" value="105.0" valueMax="120.0" valueMin="90.0">90120 mm</quantity>
(
<tableCitation id="C6E70332F9489D56FC4122658821AA0F" box="[949,1034,1631,1657]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="26" pageNumber="27">Table 1</tableCitation>
), all
<specimenCount id="9D63FD00F9489D56FBBD226588D4AA0F" box="[1097,1279,1631,1657]" count="3" pageId="26" pageNumber="27" type="generic">three specimens</specimenCount>
well-matured possessing fully equipped gonad tubules in their body cavities.Their external and internal morphologies approximately agreed with each others (
<tableCitation id="C6E70332F9489D56FE43229D8E20AAB7" box="[439,523,1703,1729]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="26" pageNumber="27">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BDA3689F9489D56FF6323298E55AB5B" blockId="26.[151,638,1811,1837]" box="[151,638,1811,1837]" pageId="26" pageNumber="27">
<heading id="D09281E5F9489D56FF6323298E55AB5B" bold="true" box="[151,638,1811,1837]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B911EA9BF9489D56FF6323298E55AB5B" bold="true" box="[151,638,1811,1837]" pageId="26" pageNumber="27">
Ossicle variation among the
<typeStatus id="54DE882BF9489D56FE0E23298E1BAB5B" box="[506,560,1811,1837]" pageId="26" pageNumber="27">type</typeStatus>
series
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9489D57FF6323618E3AAD6B" blockId="26.[151,1436,1883,2018]" lastBlockId="27.[151,1437,151,969]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Body wall ossicles anchors and anchor plates. Anchor plate ossicle length and calculated indicators of
<specimenCount id="9D63FD00F9489D56FA9023618D27ABEF" count="3" pageId="26" pageNumber="27" type="generic">three specimens</specimenCount>
ranged within: 83138 μm in length,
<emphasis id="B911EA9BF9489D56FD3523458EDAABEF" box="[705,753,1919,1945]" italics="true" pageId="26" pageNumber="27">APS</emphasis>
=0.643.0,
<emphasis id="B911EA9BF9489D56FC8723BA8FB0ABEF" box="[883,923,1920,1945]" italics="true" pageId="26" pageNumber="27">Ntp</emphasis>
=16,
<emphasis id="B911EA9BF9489D56FC112345883CABEF" box="[997,1047,1919,1945]" italics="true" pageId="26" pageNumber="27">APT</emphasis>
=2.325.0%. Among these values, length, frequency of teethed perforations
<emphasis id="B911EA9BF9489D56FD9D23998EB0ABCB" box="[617,667,1955,1981]" italics="true" pageId="26" pageNumber="27">APT</emphasis>
and their parameter
<emphasis id="B911EA9BF9489D56FC70239E8F87ABCB" box="[900,940,1956,1981]" italics="true" pageId="26" pageNumber="27">Ntp</emphasis>
, and non-size dependance indicator of plate skewness
<emphasis id="B911EA9BF9489D56FEF823FD8D17AB97" box="[268,316,1991,2017]" italics="true" pageId="26" pageNumber="27">APS</emphasis>
of anchor plate ossicles (both ventrally and dorsally) was not significantly different (
<emphasis id="B911EA9BF9489D56FAD923F28962AB97" box="[1325,1353,1992,2017]" italics="true" pageId="26" pageNumber="27">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test). From these results, the anchor plate ossicles in the body wall of the present
<specimenCount id="9D63FD00F9499D57FB1C24AD89B6ACC7" box="[1256,1437,151,177]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
were not different in ossicle length or the skewness of the layout for perforations. Therefore, the length (about 80140 μm), numbers of teethed perforations (16), and strongly distorted (up to 43) anchor plate ossicles can be appropriate diagnostic characters.
</paragraph>
<paragraph id="8BDA3689F9499D57FF33251D8820AD83" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
Anchor ossicle length and calculated indicators ranged within: 144181 μm in length,
<emphasis id="B911EA9BF9499D57FB3C251D88D5AD37" box="[1224,1278,295,321]" italics="true" pageId="27" pageNumber="28">ASW</emphasis>
=19.130.9%,
<emphasis id="B911EA9BF9499D57FF6325718CFAAD13" box="[151,209,331,357]" italics="true" pageId="27" pageNumber="28">AEW</emphasis>
=38.263.8% (
<tableCitation id="C6E70332F9499D57FE8D25718DF9AD13" box="[377,466,331,357]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="27" pageNumber="28">Table 6</tableCitation>
). Among these values, lengths of anchor ossicles (both ventrally and dorsally) were not significantly different (
<emphasis id="B911EA9BF9499D57FE3025558DCBADFE" box="[452,480,367,392]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test), while the indicator for anchor stem breadth
<emphasis id="B911EA9BF9499D57FAE42555896DADFF" box="[1296,1350,367,393]" italics="true" pageId="27" pageNumber="28">ASW</emphasis>
and for basal / distal ends width
<emphasis id="B911EA9BF9499D57FE5C25A98DC9ADDB" box="[424,482,403,429]" italics="true" pageId="27" pageNumber="28">AEW</emphasis>
were significantly different (
<emphasis id="B911EA9BF9499D57FCD325A98F68ADDA" box="[807,835,403,428]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test). From these results, the anchor ossicles in the body wall of the present
<specimenCount id="9D63FD00F9499D57FD51258D8F71ADA7" box="[677,858,439,465]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
were not different in ossicle length, but different in the shape. Therefore, proportion of the anchor ossicles can be the accurate key.
</paragraph>
<paragraph id="8BDA3689F9499D57FF3325C58D67AE87" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
Longitudinal muscle involved both O-shaped and C-shaped granule ossicles, length and calculated indicators ranged within: 2336 μm length,
<emphasis id="B911EA9BF9499D57FDDD26198E65AE4B" box="[553,590,547,573]" italics="true" pageId="27" pageNumber="28">GP</emphasis>
=31.282.6%,
<emphasis id="B911EA9BF9499D57FD0E26198F1EAE4B" box="[762,821,547,573]" italics="true" pageId="27" pageNumber="28">GCO</emphasis>
=11.177.8% (
<tableCitation id="C6E70332F9499D57FC2B26198810AE4B" box="[991,1083,547,573]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="27" pageNumber="28">Table 6</tableCitation>
). Among these values, length of granule ossicles was not different significantly (
<emphasis id="B911EA9BF9499D57FD12267D8F29AE16" box="[742,770,583,608]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test), also calculated indicator for proportions
<emphasis id="B911EA9BF9499D57FEEB26518D6FAEF3" box="[287,324,619,645]" italics="true" pageId="27" pageNumber="28">GP</emphasis>
was not different significantly in
<emphasis id="B911EA9BF9499D57FD4F26518ECBAEF3" box="[699,736,619,645]" italics="true" pageId="27" pageNumber="28">GP</emphasis>
(
<emphasis id="B911EA9BF9499D57FD0426518F27AEF2" box="[752,780,619,644]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test, both in ventrally and dorsally), but different significantly in
<emphasis id="B911EA9BF9499D57FE2A26B58E32AEDF" box="[478,537,655,681]" italics="true" pageId="27" pageNumber="28">GCO</emphasis>
(
<emphasis id="B911EA9BF9499D57FDDE26B58E6DAEDE" box="[554,582,655,680]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&lt;0.05, χ
<superScript id="7C109BC1F9499D57FD5726B48E80AEEA" attach="right" box="[675,683,654,668]" fontSize="6" pageId="27" pageNumber="28">2</superScript>
-multiple test). Therefore, the length and proportion of the granule ossicles can be the appropriate keys, while the ratio C- or O-shaped granule ossicles of this species cannot be the appropriate key.
</paragraph>
<paragraph id="8BDA3689F9499D57FF3326C18878AFBF" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
Tentacle ossicles rods, length and calculated indicators of
<specimenCount id="9D63FD00F9499D57FCA026C18823AF63" box="[852,1032,763,789]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
ranged within: 3585 μm in length,
<emphasis id="B911EA9BF9499D57FF6327258C92AF4E" box="[151,185,799,824]" italics="true" pageId="27" pageNumber="28">RP</emphasis>
=10.126.3%, and
<emphasis id="B911EA9BF9499D57FE6227258D91AF4F" box="[406,442,799,825]" italics="true" pageId="27" pageNumber="28">RC</emphasis>
=5.017.3. Among these values, length of rod ossicles was not significantly different (
<emphasis id="B911EA9BF9499D57FF5527798C99AF2A" box="[161,178,835,860]" italics="true" pageId="27" pageNumber="28">P</emphasis>
&gt;0.05, Kruskal-Wallis test), also non-size dependence indicator for proportion (
<emphasis id="B911EA9BF9499D57FBBD27798840AF2A" box="[1097,1131,835,860]" italics="true" pageId="27" pageNumber="28">RP</emphasis>
) and non-size dependence complexities (
<emphasis id="B911EA9BF9499D57FEC0275D8D73AFF7" box="[308,344,871,897]" italics="true" pageId="27" pageNumber="28">RC</emphasis>
) were not significantly different (
<emphasis id="B911EA9BF9499D57FD3C275D8ECFAFF6" box="[712,740,871,896]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test). From these results, the rod ossicles in the tentacles of the present
<specimenCount id="9D63FD00F9499D57FE0427B18E8DAFD3" box="[496,678,907,933]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
have similar lengths, proportions and complexities. Therefore, the morphologies of the rod ossicles of this species are appropriate diagnostic characters.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9499D57FF6320218F8CA9EB" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BDA3689F9499D57FF6320218D22A843" blockId="27.[151,265,1051,1077]" box="[151,265,1051,1077]" pageId="27" pageNumber="28">
<emphasis id="B911EA9BF9499D57FF6320218D22A843" bold="true" box="[151,265,1051,1077]" pageId="27" pageNumber="28">Remarks</emphasis>
</paragraph>
<paragraph id="8BDA3689F9499D57FF6320598F8CA9EB" blockId="27.[151,1437,1123,1437]" pageId="27" pageNumber="28">
Prior specimens of this species have been misidentified as
<taxonomicName id="4C654D0AF9499D57FCBC20598FD8A80A" authorityName="Chao, Rowe &amp; Chang" authorityYear="1988" box="[840,1011,1123,1148]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="taiwaniensis">
<emphasis id="B911EA9BF9499D57FCBC20598FD8A80A" box="[840,1011,1123,1148]" italics="true" pageId="27" pageNumber="28">P. taiwaniensis</emphasis>
</taxonomicName>
, based on the distorted anchor plate ossicles. However, observations herein showed that all
<specimenCount id="9D63FD00F9499D57FD0720BD8F8EA8D7" box="[755,933,1159,1185]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
possessed ciliated funnels in their right dorsal interradius (
<emphasis id="B911EA9BF9499D57FED420918D62A8B3" box="[288,329,1195,1221]" italics="true" pageId="27" pageNumber="28">IR4</emphasis>
), while
<taxonomicName id="4C654D0AF9499D57FE5320918E64A8B2" authorityName="Chao, Rowe &amp; Chang" authorityYear="1988" box="[423,591,1195,1220]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="taiwaniensis">
<emphasis id="B911EA9BF9499D57FE5320918E64A8B2" box="[423,591,1195,1220]" italics="true" pageId="27" pageNumber="28">P. taiwaniensis</emphasis>
</taxonomicName>
possesses ciliated funnels in left dorsal interradius (
<emphasis id="B911EA9BF9499D57FB54209188E3A8B3" box="[1184,1224,1195,1221]" italics="true" pageId="27" pageNumber="28">IR3</emphasis>
). Such differences are a critical morphological character (Table 7-1). Presently, genus
<taxonomicName id="4C654D0AF9499D57FC5E20F588EBA89F" authority="Heding, 1928" authorityName="Heding" authorityYear="1928" box="[938,1216,1231,1257]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="genus">
<emphasis id="B911EA9BF9499D57FC5E20F58833A89E" box="[938,1048,1231,1256]" italics="true" pageId="27" pageNumber="28">Patinapta</emphasis>
<bibRefCitation id="EFF44B78F9499D57FBD720F588EBA89F" author="Heding, S. G." box="[1059,1216,1231,1257]" pageId="27" pageNumber="28" pagination="105 - 323" refId="ref31634" refString="Heding, S. G. (1928) Synaptidae. Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. No. 66. Videnskabelige Meddeleser fra Dansk naturhistorisk Forening i KObenhavn, 85, 105 - 323." type="journal article" year="1928">Heding, 1928</bibRefCitation>
</taxonomicName>
accommodates six species from Indo-West Pacific waters (WoRMS 2024, Table 7-1; 7-2). Four other species had been described in this genus before:
<taxonomicName id="4C654D0AF9499D57FEC3212D8E59A947" authority="Heding, 1929" authorityName="Heding" authorityYear="1929" box="[311,626,1303,1329]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="crosslandii">
<emphasis id="B911EA9BF9499D57FEC3212D8DE5A947" box="[311,462,1303,1329]" italics="true" pageId="27" pageNumber="28">P. crosslandii</emphasis>
<bibRefCitation id="EFF44B78F9499D57FE22212D8E59A947" author="Heding, S. G." box="[470,626,1303,1329]" pageId="27" pageNumber="28" pagination="139 - 154" refId="ref31677" refString="Heding, S. G. (1929 [dated 1930]) Contributions to the Kowledge of the Synaputidae. I. Videnskabelige Meddeleser fra Dansk naturhistorisk Forening i KObenhavn, 88, 139 - 154." type="journal article" year="1929">Heding, 1929</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C654D0AF9499D57FD8B212D8FE3A947" authority="Cherbonnier, 1954" authorityName="Cherbonnier" authorityYear="1954" box="[639,968,1302,1329]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="dumasi">
<emphasis id="B911EA9BF9499D57FD8B212D8EC7A947" box="[639,748,1303,1329]" italics="true" pageId="27" pageNumber="28">P. dumasi</emphasis>
<bibRefCitation id="EFF44B78F9499D57FD07212D8FE3A947" author="Cherbonnier, G." box="[755,968,1302,1329]" pageId="27" pageNumber="28" pagination="252 - 260" refId="ref31597" refString="Cherbonnier, G. (1954) Note preliminarie sur les holothuries de la Mer Rouge. Bulletin Museum National Histoire Naturelle, Paris, Serie 2, 26 (2), 252 - 260." type="journal article" year="1954">Cherbonnier, 1954</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C654D0AF9499D57FC20212D88DAA947" authority="(Bedford, 1899)" baseAuthorityName="Bedford" baseAuthorityYear="1899" box="[980,1265,1303,1329]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="laevis">
<emphasis id="B911EA9BF9499D57FC20212D881AA947" box="[980,1073,1303,1329]" italics="true" pageId="27" pageNumber="28">P. laevis</emphasis>
(
<bibRefCitation id="EFF44B78F9499D57FBB5212D88C2A947" author="Bedford, F. P." box="[1089,1257,1303,1329]" pageId="27" pageNumber="28" pagination="141 - 150" refId="ref31264" refString="Bedford, F. P. (1899) Holothurians. In: Willey, A. (Ed.), Zoological Results based on material from New Britain, New Guinea, Loyalty Islands and elsewhere, collected during the years 1895, 1896 and 1897. Part II. University Press, Cambridge, pp. 141 - 150." type="book chapter" year="1899">Bedford, 1899</bibRefCitation>
)
</taxonomicName>
; and
<taxonomicName id="4C654D0AF9499D57FADA212D8DF1A923" authority="Cherbonnier, 1953" authorityName="Cherbonnier" authorityYear="1953" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="vaughani">
<emphasis id="B911EA9BF9499D57FADA212D8CD4A923" italics="true" pageId="27" pageNumber="28">Patinapta vaughani</emphasis>
<bibRefCitation id="EFF44B78F9499D57FEF221018DF1A923" author="Cherbonnier, G." box="[262,474,1338,1365]" pageId="27" pageNumber="28" pagination="501 - 504" refId="ref31556" refString="Cherbonnier, G. (1953) Note sur une nouvelle espece de Synapte de l'Ie Maurice: Patinapta vaughani n. sp. Bulletin Museum National Histoire Naturelle, Paris, Serie 2, 25, 501 - 504." type="journal article" year="1953">Cherbonnier, 1953</bibRefCitation>
</taxonomicName>
. Among these species, only
<taxonomicName id="4C654D0AF9499D57FCE221018FB2A923" authorityName="Cherbonnier" authorityYear="1953" box="[790,921,1339,1365]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="vaughani">
<emphasis id="B911EA9BF9499D57FCE221018FB2A923" box="[790,921,1339,1365]" italics="true" pageId="27" pageNumber="28">P. vaughani</emphasis>
</taxonomicName>
was reported as possessing ciliated funnels in the right dorsal interradius (
<emphasis id="B911EA9BF9499D57FE2021658DD6A90F" box="[468,509,1375,1401]" italics="true" pageId="27" pageNumber="28">IR4</emphasis>
). However, each species is distinguishable by the differences of the shapes of the longitudinal muscle granules and body wall anchor plates (Table 7-2).
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9499D57FF6321D48936AB73" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BDA3689F9499D57FF6321D48D1BAA7E" blockId="27.[151,304,1518,1544]" box="[151,304,1518,1544]" pageId="27" pageNumber="28">
<emphasis id="B911EA9BF9499D57FF6321D48D1BAA7E" bold="true" box="[151,304,1518,1544]" pageId="27" pageNumber="28">Distribution</emphasis>
</paragraph>
<paragraph id="8BDA3689F9499D57FF63220D8936AB73" blockId="27.[151,1437,1591,1797]" pageId="27" pageNumber="28">
Southern
<collectingCountry id="F3727619F9499D57FEF3220D8D6CAA27" box="[263,327,1591,1617]" name="Japan" pageId="27" pageNumber="28">Japan</collectingCountry>
(Kyushu Island and the Ryukyu Islands). The present
<specimenCount id="9D63FD00F9499D57FC32220D8856AA27" box="[966,1149,1591,1617]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
collected from intertidal sandy beach with boulders originated from dead corals, in the
<typeStatus id="54DE882BF9499D57FC9F22618FB0AA03" box="[875,923,1627,1653]" pageId="27" pageNumber="28">type</typeStatus>
locality, Yoron-jima Island, Ryukyu Islands,
<collectingRegion id="49A1F86BF9499D57FF6322458D32AAEF" box="[151,281,1663,1689]" country="Japan" name="Kagoshima" pageId="27" pageNumber="28">Kagoshima</collectingRegion>
,
<collectingCountry id="F3727619F9499D57FEDE22458D46AAEF" box="[298,365,1663,1689]" name="Japan" pageId="27" pageNumber="28">Japan</collectingCountry>
. Among the specimens housed in WMNH-INV, including the present specimens, 4 bottles including
<specimenCount id="9D63FD00F9499D57FEFC22998DB9AACB" box="[264,402,1699,1725]" count="9" pageId="27" pageNumber="28" type="generic">9 specimens</specimenCount>
were identified as this species (
<tableCitation id="C6E70332F9499D57FD0222998F67AACB" box="[758,844,1699,1725]" captionStart="TABLE 8" captionStartId="31.[152,244,152,176]" captionTargetPageId="31" captionText="TABLE 8. List of WMNH-INV specimens of Patinapta deformis sp. nov. as of 2024." pageId="27" pageNumber="28">Table 8</tableCitation>
). Other than the
<typeStatus id="54DE882BF9499D57FBFD22998812AACB" box="[1033,1081,1699,1725]" pageId="27" pageNumber="28">type</typeStatus>
locality,
<specimenCount id="9D63FD00F9499D57FB542299890BAACB" box="[1184,1312,1699,1725]" count="6" pageId="27" pageNumber="28" type="immature">6 immature</specimenCount>
specimens collected from Yokaku bay and Hirose-gawa river mouth Amakusa City,
<collectingRegion id="49A1F86BF9499D57FC3B22FD887BAA97" box="[975,1104,1735,1761]" country="Japan" name="Kumamoto" pageId="27" pageNumber="28">Kumamoto</collectingRegion>
,
<collectingCountry id="F3727619F9499D57FBA822FD88B7AA97" box="[1116,1180,1735,1761]" name="Japan" pageId="27" pageNumber="28">Japan</collectingCountry>
were identified as this species. In both localities in Amakusa City, the specimens were collected from estuarine intertidal flats.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,153 +0,0 @@
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<mods:title id="9A307E0C853D23E0E166F101707B56A7">First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan</mods:title>
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<subSubSection id="C37F6502F9449D5AFF63217B8EEAA909" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BDA3689F9449D5AFF63217B8DC8A92D" blockId="22.[151,705,1344,1443]" box="[151,483,1344,1371]" pageId="22" pageNumber="23">
<heading id="D09281E5F9449D5AFF63217B8DC8A92D" bold="true" box="[151,483,1344,1371]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B911EA9BF9449D5AFF63217B8DC8A92D" bold="true" box="[151,483,1344,1371]" pageId="22" pageNumber="23">
<taxonomicName id="4C654D0AF9449D5AFF63217B8D55A92D" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[151,382,1344,1371]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="22" pageNumber="23" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF9449D5AFF63217B8D55A92D" bold="true" box="[151,382,1344,1371]" italics="true" pageId="22" pageNumber="23">Patinapta deformis</emphasis>
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<taxonomicNameLabel id="A22257E0F9449D5AFE71217B8DC8A92D" box="[389,483,1345,1371]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9449D5AFF63215C8EEAA909" blockId="22.[151,705,1344,1443]" box="[151,705,1381,1408]" pageId="22" pageNumber="23">[New Japanese name: Yoron-himo-ikari-namako]</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5AFF6321B38DA4A98B" pageId="22" pageNumber="23" type="description">
<paragraph id="8BDA3689F9449D5AFF6321B38DE0A9D5" blockId="22.[151,705,1344,1443]" box="[151,459,1417,1443]" pageId="22" pageNumber="23">
(
<figureCitation id="135E2A0CF9449D5AFF6B21B38D0AA9D5" box="[159,289,1417,1443]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStartId-0="23.[151,250,1446,1470]" captionStartId-1="25.[151,250,1699,1723]" captionStartId-2="26.[151,250,516,540]" captionTargetBox-0="[151,1436,309,1423]" captionTargetBox-1="[151,1436,181,1667]" captionTargetBox-2="[164,1423,188,485]" captionTargetId-0="figure-69@23.[151,1436,309,1423]" captionTargetId-1="figure-20@25.[151,1436,181,1675]" captionTargetId-2="figure-29@26.[151,1435,181,493]" captionTargetPageId-0="23" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="FIGURE 12. External view of holotype specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (specimen WMNH-2015-INV-106): A, lateral view, anterior left, dorsal top; B, oral-side view of the tentacles; C, out-side view of the tentacles." captionText-1="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." captionText-2="FIGURE 14. Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima (holotype specimen, WMNH-2015-INV-106), described as Patinapta deformis sp. nov., in the present study. The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stand for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi-0="http://doi.org/10.5281/zenodo.14703556" figureDoi-1="http://doi.org/10.5281/zenodo.14703560" figureDoi-2="http://doi.org/10.5281/zenodo.14703562" httpUri-0="https://zenodo.org/record/14703556/files/figure.png" httpUri-1="https://zenodo.org/record/14703560/files/figure.png" httpUri-2="https://zenodo.org/record/14703562/files/figure.png" pageId="22" pageNumber="23">Figs 1214</figureCitation>
;
<tableCitation id="C6E70332F9449D5AFED821B38DA1A9D5" box="[300,394,1417,1443]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="22" pageNumber="23">Tables 1</tableCitation>
,
<tableCitation id="C6E70332F9449D5AFE6C21B38D83A9D5" box="[408,424,1417,1443]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="22" pageNumber="23">6</tableCitation>
,
<tableCitation id="C6E70332F9449D5AFE4021B38DE9A9D5" box="[436,450,1417,1443]" captionStart="TABLE 8" captionStartId="31.[152,244,152,176]" captionText="TABLE 8. List of WMNH-INV specimens of Patinapta deformis sp. nov. as of 2024." pageId="22" pageNumber="23">8</tableCitation>
)
</paragraph>
<paragraph id="8BDA3689F9449D5AFF6321FD8DA4A98B" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">
<materialsCitation id="3B0D3CD4F9449D5AFF6321FD8DA1A98B" location="Yoron-jima Island" municipality="We" pageId="22" pageNumber="23" specimenCount="3">
<specimenCount id="9D63FD00F9449D5AFF6321FD8D68A9A9" box="[151,323,1479,1503]" count="3" pageId="22" pageNumber="23" type="generic">Three specimens</specimenCount>
from
<location id="8EBA6052F9449D5AFE7521FD8E12A9A9" LSID="urn:lsid:plazi:treatment:03CC879FF9449D5BFF63217B88B3AD6B:8EBA6052F9449D5AFE7521FD8E12A9A9" box="[385,569,1479,1503]" municipality="We" name="Yoron-jima Island" pageId="22" pageNumber="23">Yoron-jima Island</location>
were identified as new to science.
<collectingMunicipality id="6BBEACF3F9449D5AFC6B21FD8FEBA9A8" box="[927,960,1479,1502]" pageId="22" pageNumber="23">We</collectingMunicipality>
herein described the morphological characters of this new species
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5AFF63223D8DC3AAF8" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BDA3689F9449D5AFF63223D8DC3AAF8" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">
<emphasis id="B911EA9BF9449D5AFF63223D8D39AA57" bold="true" box="[151,274,1543,1569]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
Total shapes of anchor plates strongly distorted, their large perforations (mostly 23) fewer than those of the two congener nominal species (mostly 47), around these few perforations, marginal area broad and flat. Basal T-shaped end of anchor ossicles large, semicircular-shape. Ciliated funnels attached at inner side body wall of right dorso-lateral interradial (
<emphasis id="B911EA9BF9449D5AFE5B224E8DF3AAF8" box="[431,472,1652,1678]" italics="true" pageId="22" pageNumber="23">IR4</emphasis>
).
</paragraph>
</subSubSection>
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<paragraph id="8BDA3689F9449D5AFF3322A288F2AAC4" blockId="22.[151,1437,1479,2038]" box="[199,1241,1687,1714]" pageId="22" pageNumber="23">
<emphasis id="B911EA9BF9449D5AFF3322A28D78AAC7" bold="true" box="[199,339,1687,1714]" pageId="22" pageNumber="23">Type series.</emphasis>
<typeStatus id="54DE882BF9449D5AFEAE22A28DEEAAC4" box="[346,453,1688,1714]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
, WMNH-2015-INV-106;
<specimenCount id="9D63FD00F9449D5AFD1322A28FADAAC4" box="[743,902,1688,1714]" count="2" pageId="22" pageNumber="23" type="generic" typeStatus="paratype">two paratypes</specimenCount>
, WMNH-2015-INV-105, 108.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5BFF3322818D56AC8F" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="8BDA3689F9449D5AFF3322818E71ABD8" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">
<emphasis id="B911EA9BF9449D5AFF3322818880AAA3" bold="true" box="[199,1195,1723,1750]" pageId="22" pageNumber="23">
Description of
<typeStatus id="54DE882BF9449D5AFE8322868DF5AAA0" box="[375,478,1724,1750]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(of WMNH-2015-INV-106: middle-size complete specimen).
</emphasis>
External and internal morphologies as follows (
<tableCitation id="C6E70332F9449D5AFE4122DA8E22AA8C" box="[437,521,1760,1786]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="22" pageNumber="23">Table 1</tableCitation>
): 30 years-preserved specimen pale peanut color (
<figureCitation id="135E2A0CF9449D5AFBC122DA88ACAA8C" box="[1077,1159,1760,1786]" captionStart="FIGURE 12" captionStartId="23.[151,250,1446,1470]" captionTargetBox="[151,1436,309,1423]" captionTargetId="figure-69@23.[151,1436,309,1423]" captionTargetPageId="23" captionText="FIGURE 12. External view of holotype specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (specimen WMNH-2015-INV-106): A, lateral view, anterior left, dorsal top; B, oral-side view of the tentacles; C, out-side view of the tentacles." figureDoi="http://doi.org/10.5281/zenodo.14703556" httpUri="https://zenodo.org/record/14703556/files/figure.png" pageId="22" pageNumber="23">Fig. 12</figureCitation>
). Tentacles 12, each with 10 digits of 5 pairs. On the oral-side of tentacle stem, about 1020 sensory cups sporadically arranged in U- shape. Polian vesicle in medio-ventral position, fusiform, double (
<tableCitation id="C6E70332F9449D5AFCB323128FB5AB34" box="[839,926,1832,1858]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="22" pageNumber="23">Table 1</tableCitation>
), stone canal single thread-like, attached to narrow space between basal parts of tentacle ampullae at mediodorsal (
<emphasis id="B911EA9BF9449D5AFC4323768FF4AB10" box="[951,991,1868,1894]" italics="true" pageId="22" pageNumber="23">IR5</emphasis>
), suddenly turned upward and forward, resulted in vesicle facing forward, distally with mold-like madreporite. Ciliated funnels in one crowded band along midline of right dorsal interradius (
<emphasis id="B911EA9BF9449D5AFDD523AE8E61ABD8" box="[545,586,1940,1966]" italics="true" pageId="22" pageNumber="23">IR4</emphasis>
).
</paragraph>
<paragraph id="8BDA3689F9449D5AFF332382889CAB80" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">Inside body, two tufts of gonad tubules attached with both sides of anterior dorsal mesentery, also two whitish tubule organs adhered to upper and under sides of intestine canal. Intestine canal lacking loop.</paragraph>
<paragraph id="8BDA3689F9459D5BFF3324AD8D56AC8F" blockId="23.[151,1436,151,285]" pageId="23" pageNumber="24">Body wall coarse surface, hard and thick, sparsely scattered with ossicles, arranging latitudinal layout of all as sets of anchor and plate, their directions somewhat ordered but without symmetrical design against each axial of longitudinal muscle.</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9459D5BFF33253988B3AD6B" box="[199,1176,259,285]" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BDA3689F9459D5BFF33253988B3AD6B" blockId="23.[151,1436,151,285]" box="[199,1176,259,285]" pageId="23" pageNumber="24">
<emphasis id="B911EA9BF9459D5BFF3325398D65AD6B" bold="true" box="[199,334,259,285]" pageId="23" pageNumber="24">Etymology.</emphasis>
The species name
<taxonomicName id="4C654D0AF9459D5BFDD625398EAFAD6B" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[546,644,259,285]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="deformis">
<emphasis id="B911EA9BF9459D5BFDD625398EAFAD6B" box="[546,644,259,285]" italics="true" pageId="23" pageNumber="24">deformis</emphasis>
</taxonomicName>
, alluding to the strongly distorted anchor plate.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,164 +0,0 @@
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<mods:title id="6FB93982A8E49CD2BEB337DF35DCB769">First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan</mods:title>
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<heading id="D09281E5F94D9D53FF6326248E0AAE4E" bold="true" box="[151,545,541,568]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B911EA9BF94D9D53FF6326248E0AAE4E" bold="true" box="[151,545,541,568]" pageId="31" pageNumber="32">
<taxonomicName id="4C654D0AF94D9D53FF6326248D97AE4E" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[151,444,541,568]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="parvaspiculus" status="sp. nov.">
<emphasis id="B911EA9BF94D9D53FF6326248D97AE4E" bold="true" box="[151,444,541,568]" italics="true" pageId="31" pageNumber="32">Patinapta parvaspiculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257E0F94D9D53FE3726248E0AAE4E" box="[451,545,542,568]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F94D9D53FF6326788EF7AE2A" blockId="31.[151,732,541,640]" box="[151,732,578,604]" pageId="31" pageNumber="32">[New Japanese name: Kokotsu-himo-ikari-namako]</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF63265C89A9AE97" pageId="31" pageNumber="32" type="description">
<paragraph id="8BDA3689F94D9D53FF63265C8DF1AEF6" blockId="31.[151,732,541,640]" box="[151,474,613,640]" pageId="31" pageNumber="32">
(
<figureCitation id="135E2A0CF94D9D53FF6B265C8D0AAEF6" box="[159,289,613,640]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStart-2="FIGURE 17" captionStartId-0="33.[151,250,1258,1282]" captionStartId-1="34.[151,250,1674,1698]" captionStartId-2="35.[151,250,628,652]" captionTargetBox-0="[151,1436,181,1234]" captionTargetBox-1="[158,1430,181,1643]" captionTargetBox-2="[164,1422,281,600]" captionTargetId-0="figure-20@33.[151,1436,178,1237]" captionTargetId-1="figure-29@34.[158,1430,181,1650]" captionTargetId-2="figure-65@35.[151,1435,273,605]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="35" captionText-0="FIGURE 15. External view of a damaged specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, lateral view, anterior left, dorsal top with damage on anterior body wall (from sampling shovel; gonad tubules also damaged); B, oral-side view of the tentacles; C, outer-side view of the tentacles." captionText-1="FIGURE 16. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." captionText-2="FIGURE 17. Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157). The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stands for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi-0="http://doi.org/10.5281/zenodo.14703564" figureDoi-1="http://doi.org/10.5281/zenodo.14703566" figureDoi-2="http://doi.org/10.5281/zenodo.14703568" httpUri-0="https://zenodo.org/record/14703564/files/figure.png" httpUri-1="https://zenodo.org/record/14703566/files/figure.png" httpUri-2="https://zenodo.org/record/14703568/files/figure.png" pageId="31" pageNumber="32">Figs 1517</figureCitation>
;
<tableCitation id="C6E70332F94D9D53FED8265C8DA1AEF6" box="[300,394,614,640]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="31" pageNumber="32">Tables 1</tableCitation>
,
<tableCitation id="C6E70332F94D9D53FE6C265C8D83AEF6" box="[408,424,614,640]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="31" pageNumber="32">9</tableCitation>
,
<tableCitation id="C6E70332F94D9D53FE40265C8DFAAEF6" box="[436,465,614,640]" captionStart="TABLE 10" captionStartId="36.[152,244,872,896]" captionText="TABLE 10. List of WMNH-INV specimens of Patinapta parvaspiculus sp. nov. as of 2024." pageId="31" pageNumber="32">10</tableCitation>
)
</paragraph>
<paragraph id="8BDA3689F94D9D53FF63269189A9AE97" blockId="31.[151,1436,682,737]" pageId="31" pageNumber="32">
<materialsCitation id="3B0D3CD4F94D9D53FF6326918FD6AEB4" box="[151,1021,682,706]" location="Yoron-jima Island" pageId="31" pageNumber="32" specimenCount="2">
<specimenCount id="9D63FD00F94D9D53FF6326918D1FAEB4" box="[151,308,682,706]" count="2" pageId="31" pageNumber="32" type="generic">Two specimens</specimenCount>
from
<location id="8EBA6052F94D9D53FE8526908E02AEB4" LSID="urn:lsid:plazi:treatment:03CC879FF94D9D53FF6326248879AA37:8EBA6052F94D9D53FE8526908E02AEB4" box="[369,553,682,706]" name="Yoron-jima Island" pageId="31" pageNumber="32">Yoron-jima Island</location>
(WMNH-2015-INV-107; 2023-INV-157) and
</materialsCitation>
<materialsCitation id="3B0D3CD4F94D9D53FBF726918956AE97" location="Imari Bay" municipality="We" pageId="31" pageNumber="32" specimenCount="1">
<specimenCount id="9D63FD00F94D9D53FBF7269188A5AEB4" box="[1027,1166,682,706]" count="1" pageId="31" pageNumber="32" type="generic">one specimen</specimenCount>
from
<location id="8EBA6052F94D9D53FB3826908919AEB5" LSID="urn:lsid:plazi:treatment:03CC879FF94D9D53FF6326248879AA37:8EBA6052F94D9D53FB3826908919AEB5" box="[1228,1330,682,707]" municipality="We" name="Imari Bay" pageId="31" pageNumber="32">Imari Bay</location>
(WMNH-2024-INV-156) were identified as new to science.
<collectingMunicipality id="6BBEACF3F94D9D53FD3226F38ECCAE96" box="[710,743,713,736]" pageId="31" pageNumber="32">We</collectingMunicipality>
herein describe the morphological characters of this new species
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF6327358EBFAFAB" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BDA3689F94D9D53FF6327358EBFAFAB" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">
<emphasis id="B911EA9BF94D9D53FF6327358D39AF5F" bold="true" box="[151,274,783,809]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
Anchor and anchor plate ossicles small. Anchor plate ossicles smaller than 100 μm, while anchor ossicles up to 150 μm, resulting in unbalanced size combinations of bounded ossicles. Total shapes of anchor plates strongly distorted (because of small total numbers of perforations), basally with a concave shallow or deep, resulting in a two-tipped ovoid shape. Perforations of anchor plate ossicles mostly without teeth, while outer edge of anchor plate ossicles equipped with minute teeth in small numbers. Ciliated funnels attached to inner side body wall along midline of left dorso-lateral interradius (
<emphasis id="B911EA9BF94D9D53FDAF27F98EA8AFAB" box="[603,643,963,989]" italics="true" pageId="31" pageNumber="32">IR3</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF3327DD8CE3A850" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BDA3689F94D9D53FF3327DD8CE3A850" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">
<emphasis id="B911EA9BF94D9D53FF3327DD8D7DA877" bold="true" box="[199,342,999,1025]" pageId="31" pageNumber="32">Type series.</emphasis>
<typeStatus id="54DE882BF94D9D53FE9427DD8DE0A877" box="[352,459,999,1025]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
, WMNH-2023-INV-157;
<specimenCount id="9D63FD00F94D9D53FD0727D28FBEA874" box="[755,917,1000,1026]" count="2" pageId="31" pageNumber="32" type="generic" typeStatus="paratype">two paratypes</specimenCount>
, WMNH-2015-INV-107; WMNH-2024-INV-156.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF3320158E22AA6B" pageId="31" pageNumber="32" type="description">
<paragraph id="8BDA3689F94D9D53FF3320158F22A8C3" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">
<emphasis id="B911EA9BF94D9D53FF332015888AA83F" bold="true" box="[199,1185,1071,1097]" pageId="31" pageNumber="32">
Description of
<typeStatus id="54DE882BF94D9D53FE8920158DCFA83F" box="[381,484,1071,1097]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
(WMNH-2023-INV-157: middle-size damaged specimen).
</emphasis>
External and internal morphologies as follows (
<tableCitation id="C6E70332F94D9D53FE3620698E31A81B" box="[450,538,1107,1133]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="31" pageNumber="32">Table 1</tableCitation>
): 30 year-old preserved well-matured specimen pale peanut color, body large about
<quantity id="4C9D9B6CF94D9D53FF2A204D8D16A8E4" box="[222,317,1143,1170]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="mm" value="100.0">100 mm</quantity>
length (
<figureCitation id="135E2A0CF94D9D53FE68204D8DDAA8E4" box="[412,497,1143,1170]" captionStart="FIGURE 15" captionStartId="33.[151,250,1258,1282]" captionTargetBox="[151,1436,181,1234]" captionTargetId="figure-20@33.[151,1436,178,1237]" captionTargetPageId="33" captionText="FIGURE 15. External view of a damaged specimen collected from Yoron-jima Island, Kagoshima, erected as Patinapta parvaspiculus sp. nov., in the present study (holotype specimen, WMNH-2023-INV-157): A, lateral view, anterior left, dorsal top with damage on anterior body wall (from sampling shovel; gonad tubules also damaged); B, oral-side view of the tentacles; C, outer-side view of the tentacles." figureDoi="http://doi.org/10.5281/zenodo.14703564" httpUri="https://zenodo.org/record/14703564/files/figure.png" pageId="31" pageNumber="32">Fig. 15</figureCitation>
). Tentacles 12, each with 10 digits of 5 pairs. On oral-side of tentacle stem, about 1016 sensory cups sporadically arranged in a U-shape.
</paragraph>
<paragraph id="8BDA3689F94D9D53FF3320858DFBA933" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">
Polian vesicle in medio-ventral position, fusiform, triple (
<tableCitation id="C6E70332F94D9D53FCB920858F8AA8AF" box="[845,929,1215,1241]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="31" pageNumber="32">Table 1</tableCitation>
), stone canal single and thread-like, attached to narrow space between basal parts of tentacle ampullae at mediodorsal (
<emphasis id="B911EA9BF94D9D53FC3420D98FC2A88B" box="[960,1001,1251,1277]" italics="true" pageId="31" pageNumber="32">IR5</emphasis>
), suddenly turned upward and forward, resulted in it facing forward, distally with moldy surface madreporite. Ciliated funnels in one crowded band along left dorsal interradius (
<emphasis id="B911EA9BF94D9D53FE6321118D94A933" box="[407,447,1323,1349]" italics="true" pageId="31" pageNumber="32">IR3</emphasis>
).
</paragraph>
<paragraph id="8BDA3689F94D9D53FF332175889CA9FB" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">Inside body, two tufts of gonad tubules attached to both sides of anterior dorsal mesentery, also two whitish tubule organs adhered to upper and under sides of intestine canal. Intestine canal lacking loop.</paragraph>
<paragraph id="8BDA3689F94D9D53FF3321AD8E22AA6B" blockId="31.[151,1437,783,1602]" pageId="31" pageNumber="32">Body wall smooth surface, hard but thin, with ossicle arranging latitudinal layout of all as sets of anchor and plate. Both sides along five longitudinal muscles with distal ends of anchor ossicles facing towards interradial area, however ossicles on radial body walls upon longitudinal muscles and interradial body walls have distal ends randomly facing either direction.</paragraph>
</subSubSection>
<subSubSection id="C37F6502F94D9D53FF33221D8879AA37" box="[199,1106,1575,1602]" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BDA3689F94D9D53FF33221D8879AA37" blockId="31.[151,1437,783,1602]" box="[199,1106,1575,1602]" pageId="31" pageNumber="32">
<emphasis id="B911EA9BF94D9D53FF33221D8D65AA37" bold="true" box="[199,334,1575,1601]" pageId="31" pageNumber="32">Etymology.</emphasis>
The species name
<taxonomicName id="4C654D0AF94D9D53FDD6221D8E94AA37" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[546,703,1575,1601]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="parvaspiculus">
<emphasis id="B911EA9BF94D9D53FDD6221D8E94AA37" box="[546,703,1575,1601]" italics="true" pageId="31" pageNumber="32">parvaspiculus</emphasis>
</taxonomicName>
, alluding to the ossicles small size.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="826FBEE58D871F2A17EBAC3B36CD322F">First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan</mods:title>
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<mods:namePart id="43F9F9D64A839E30D41FB972E8BF05B5">Yamana, Yusuke</mods:namePart>
<mods:affiliation id="9EF68ECE01F5005B14202BE7A6CE6AF1">Wakayama Prefectural Museum of Natural History (WMNH), Invertebrate Specimen House, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan</mods:affiliation>
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<emphasis id="B911EA9BF9629D7CFF6327E98FC5AF9B" bold="true" box="[151,1006,979,1005]" pageId="48" pageNumber="49">
Key to Japanese species of
<taxonomicName id="4C654D0AF9629D7CFE1C27E98E76AF9B" ID-CoL="7PQHQ" authorityName="Heding" authorityYear="1928" box="[488,605,979,1005]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="genus">
<emphasis id="B911EA9BF9629D7CFE1C27E98E76AF9B" bold="true" box="[488,605,979,1005]" italics="true" pageId="48" pageNumber="49">Patinapta</emphasis>
</taxonomicName>
(with present four new species).
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9629D7CFF63202289B7A9F8" pageId="48" pageNumber="49" type="key">
<keyStep id="30912E2CF9629D7CFF63202289B7A83C" pageId="48" pageNumber="49">
<paragraph id="8BDA3689F9629D7CFF63202289B7A85B" blockId="48.[151,1437,1048,1422]" box="[151,1436,1048,1070]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF63202289B7A85B" box="[151,1436,1048,1070]" pageId="48" pageNumber="49">1. In tentacles, rod-shaped ossicles present................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BDA3689F9629D7CFF63200C89B7A83C" blockId="48.[151,1437,1048,1422]" box="[151,1436,1077,1099]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF63200C89B7A83C" box="[151,1436,1077,1099]" pageId="48" pageNumber="49">
- In tentacles, ossicles mostly elongated granules present.......................................
<taxonomicName id="4C654D0AF9629D7CFB22200C8961A83C" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[1238,1354,1077,1099]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="edentatus" status="sp. nov.">
<emphasis id="B911EA9BF9629D7CFB22200C8961A83C" box="[1238,1354,1077,1099]" italics="true" pageId="48" pageNumber="49">P. edentatus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9629D7CFAA4200F89B7A83C" bold="true" box="[1360,1436,1077,1098]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A22257E0F9629D7CFAA4200F89B7A83C" box="[1360,1436,1077,1098]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E2CF9629D7CFF63206989B7A8D4" pageId="48" pageNumber="49">
<paragraph id="8BDA3689F9629D7CFF63206989B7A81E" blockId="48.[151,1437,1048,1422]" box="[151,1436,1107,1129]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF63206989B7A81E" box="[151,1436,1107,1129]" pageId="48" pageNumber="49">2. In mid-body wall, more or less symmetrical anchor plate ossicles present......................................... 3</keyLead>
</paragraph>
<paragraph id="8BDA3689F9629D7CFF63204A89B7A8D4" blockId="48.[151,1437,1048,1422]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF63204A89B7A8D4" pageId="48" pageNumber="49">
- Anchor plate ossicles possess strongly distorted layout of few large perforations and small marginal perforations, with small numbers of teethed perforations. Anchor ossicles with semicircular T-shaped basal end...............
<taxonomicName id="4C654D0AF9629D7CFB2A20B48961A8D4" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[1246,1354,1165,1187]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF9629D7CFB2A20B48961A8D4" box="[1246,1354,1165,1187]" italics="true" pageId="48" pageNumber="49">P. deformis</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9629D7CFAA420B789B7A8D4" bold="true" box="[1360,1436,1165,1186]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A22257E0F9629D7CFAA420B789B7A8D4" box="[1360,1436,1165,1186]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E2CF9629D7CFF63209189B7A88C" pageId="48" pageNumber="49">
<paragraph id="8BDA3689F9629D7CFF63209189B7A8B6" blockId="48.[151,1437,1048,1422]" box="[151,1436,1195,1217]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF63209189B7A8B6" box="[151,1436,1195,1217]" pageId="48" pageNumber="49">3. Anchor plate ossicles in mid-body wall mostly larger than 100 μm.............................................. 4</keyLead>
</paragraph>
<paragraph id="8BDA3689F9629D7CFF6320F289B7A88C" blockId="48.[151,1437,1048,1422]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF6320F289B7A88C" pageId="48" pageNumber="49">
- Anchor plate ossicles with few numbers of teethed perforations and few numbers of teeth on peripheral plate edge, basal tip of plate depressed, resulting in double tips.................................................
<taxonomicName id="4C654D0AF9629D7CFB5F20DC8961A88C" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[1195,1354,1253,1275]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="parvaspiculus" status="sp. nov.">
<emphasis id="B911EA9BF9629D7CFB5F20DC8961A88C" box="[1195,1354,1253,1275]" italics="true" pageId="48" pageNumber="49">P. parvaspiculus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9629D7CFAA420DF89B7A88C" bold="true" box="[1360,1436,1253,1274]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A22257E0F9629D7CFAA420DF89B7A88C" box="[1360,1436,1253,1274]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E2CF9629D7CFF63213989B7A924" pageId="48" pageNumber="49">
<paragraph id="8BDA3689F9629D7CFF63213989B7A940" blockId="48.[151,1437,1048,1422]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF63213989B7A940" pageId="48" pageNumber="49">4. Anchor plate ossicles approximately 90110 μm in length, without minute teeth on peripheral plate edge. Anchor ossicles possess teeth (barb).................................................................................... 5</keyLead>
</paragraph>
<paragraph id="8BDA3689F9629D7CFF63210489B7A924" blockId="48.[151,1437,1048,1422]" box="[151,1436,1341,1363]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF63210489B7A924" box="[151,1436,1341,1363]" pageId="48" pageNumber="49">
- Anchor ossicles mostly lacking teeth (barbless) in both immature and mature animals................
<taxonomicName id="4C654D0AF9629D7CFB2321048961A924" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[1239,1354,1341,1363]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="B911EA9BF9629D7CFB2321048961A924" box="[1239,1354,1341,1363]" italics="true" pageId="48" pageNumber="49">P. neglectus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9629D7CFAA4210789B7A924" bold="true" box="[1360,1436,1341,1362]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A22257E0F9629D7CFAA4210789B7A924" box="[1360,1436,1341,1362]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30912E2CF9629D7CFF63216189B7A9F8" pageId="48" pageNumber="49">
<paragraph id="8BDA3689F9629D7CFF63216189B7A906" blockId="48.[151,1437,1048,1422]" box="[151,1436,1371,1393]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF63216189B7A906" box="[151,1436,1371,1393]" pageId="48" pageNumber="49">
5. Most of perforations of anchor plate ossicles with teeth, which are sharp and closely and continuously standing....
<taxonomicName id="4C654D0AF9629D7CFAB7216189B7A906" ID-CoL="765ZB" box="[1347,1436,1371,1392]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="ooplax">
<emphasis id="B911EA9BF9629D7CFAB7216189B7A906" box="[1347,1436,1371,1392]" italics="true" pageId="48" pageNumber="49">P. ooplax</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDA3689F9629D7CFF63214289B7A9F8" blockId="48.[151,1437,1048,1422]" box="[151,1436,1400,1422]" pageId="48" pageNumber="49">
<keyLead id="309495BCF9629D7CFF63214289B7A9F8" box="[151,1436,1400,1422]" pageId="48" pageNumber="49">
- small numbers of perforations (14) of anchor plates with teeth, teeth blunt and discontinuously standing....
<taxonomicName id="4C654D0AF9629D7CFAF9214389B7A9F8" ID-CoL="6TTZC" box="[1293,1436,1401,1422]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="taiwaniensis">
<emphasis id="B911EA9BF9629D7CFAF9214389B7A9F8" box="[1293,1436,1401,1422]" italics="true" pageId="48" pageNumber="49">P. taiwaniensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

View file

@ -1,69 +1,69 @@
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<mods:affiliation id="4EE5A2FA84D5681D9729366617E4948E">Wakayama Prefectural Museum of Natural History (WMNH), Invertebrate Specimen House, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan</mods:affiliation>
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<treatment id="03CC879FF9769D64FF63229C8FC6ADFF" LSID="urn:lsid:plazi:treatment:03CC879FF9769D64FF63229C8FC6ADFF" httpUri="http://treatment.plazi.org/id/03CC879FF9769D64FF63229C8FC6ADFF" lastPageId="40" lastPageNumber="41" pageId="36" pageNumber="37">
<subSubSection id="C37F6502F9769D68FF63229C8EF7AA92" pageId="36" pageNumber="37" type="nomenclature">
<treatment id="03CC879FF9769D66FF63229C8E96A923" ID-DOI="http://doi.org/10.5281/zenodo.14704984" ID-Zenodo-Dep="14704984" LSID="urn:lsid:plazi:treatment:03CC879FF9769D66FF63229C8E96A923" httpUri="http://treatment.plazi.org/id/03CC879FF9769D66FF63229C8E96A923" lastPageId="42" lastPageNumber="43" pageId="36" pageNumber="37">
<subSubSection id="C37F6502F9769D68FF63229C8DC0AAB6" box="[151,491,1702,1729]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="8BDA3689F9769D68FF63229C8DC0AAB6" blockId="36.[151,732,1702,1800]" box="[151,491,1702,1729]" pageId="36" pageNumber="37">
<heading id="D09281E5F9769D68FF63229C8DC0AAB6" bold="true" box="[151,491,1702,1729]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<emphasis id="B911EA9BF9769D68FF63229C8DC0AAB6" bold="true" box="[151,491,1702,1729]" pageId="36" pageNumber="37">
@ -74,32 +74,36 @@
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9769D68FF6322F18EF7AA92" box="[151,732,1738,1765]" pageId="36" pageNumber="37" type="vernacular_names">
<paragraph id="8BDA3689F9769D68FF6322F18EF7AA92" blockId="36.[151,732,1702,1800]" box="[151,732,1738,1765]" pageId="36" pageNumber="37">[New Japanese name: Satsuma-himo-ikari-namako]</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9769D68FF6322D48EADAB1F" pageId="36" pageNumber="37" type="description">
<subSubSection id="C37F6502F9769D68FF6322D48DCDAB7E" box="[151,486,1774,1800]" pageId="36" pageNumber="37" type="description">
<paragraph id="8BDA3689F9769D68FF6322D48DCDAB7E" blockId="36.[151,732,1702,1800]" box="[151,486,1774,1800]" pageId="36" pageNumber="37">
(
<figureCitation id="135E2A0CF9769D68FF6B22D48D0AAB7E" box="[159,289,1774,1800]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStart-2="FIGURE 20" captionStartId-0="37.[151,250,1319,1343]" captionStartId-1="38.[151,250,1613,1637]" captionStartId-2="41.[151,250,585,609]" captionTargetBox-0="[151,1436,181,1295]" captionTargetBox-1="[155,1433,181,1581]" captionTargetBox-2="[163,1422,188,552]" captionTargetId-0="figure-20@37.[151,1436,181,1295]" captionTargetId-1="figure-29@38.[155,1433,181,1589]" captionTargetId-2="figure-20@41.[151,1435,181,561]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="41" captionText-0="FIGURE 18. External view of a depressed specimen collected from Iso beach inner coast of Kagoshima Bay, Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, lateral view, anterior left; B, oral-side view of the tentacles; C, outer-side view of the tentacles." captionText-1="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." captionText-2="FIGURE 20. Outer-surface view of calcareous ring of the calcareous ring plates of a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022- INV-154): Positions of the plates denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stands for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi-0="http://doi.org/10.5281/zenodo.14703570" figureDoi-1="http://doi.org/10.5281/zenodo.14703572" figureDoi-2="http://doi.org/10.5281/zenodo.14703574" httpUri-0="https://zenodo.org/record/14703570/files/figure.png" httpUri-1="https://zenodo.org/record/14703572/files/figure.png" httpUri-2="https://zenodo.org/record/14703574/files/figure.png" pageId="36" pageNumber="37">Figs 1820</figureCitation>
;
<tableCitation id="C6E70332F9769D68FED822D48DA1AB7E" box="[300,394,1774,1800]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="36" pageNumber="37">Tables 1</tableCitation>
<tableCitation id="C6E70332F9769D68FED822D48DA1AB7E" box="[300,394,1774,1800]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="36" pageNumber="37">Tables 1</tableCitation>
,
<tableCitation id="C6E70332F9769D68FE6C22D48D9EAB7E" box="[408,437,1774,1800]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="36" pageNumber="37">11</tableCitation>
<tableCitation id="C6E70332F9769D68FE6C22D48D9EAB7E" box="[408,437,1774,1800]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="36" pageNumber="37">11</tableCitation>
,
<tableCitation id="C6E70332F9769D68FE3522D48DF5AB7E" box="[449,478,1774,1800]" captionStart="TABLE 12" captionStartId="42.[152,244,1412,1436]" captionText="TABLE 12. List of WMNH-INV specimens of Patinapta neglectus sp. nov., as of 2024." pageId="36" pageNumber="37">12</tableCitation>
<tableCitation id="C6E70332F9769D68FE3522D48DF5AB7E" box="[449,478,1774,1800]" captionStart="TABLE 12" captionStartId="42.[152,244,1412,1436]" captionTargetPageId="42" captionText="TABLE 12. List of WMNH-INV specimens of Patinapta neglectus sp. nov., as of 2024." pageId="36" pageNumber="37">12</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9769D68FF6323098EADAB1F" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BDA3689F9769D68FF6323098EADAB1F" blockId="36.[151,1437,1843,1897]" pageId="36" pageNumber="37">
<materialsCitation id="3B0D3CD4F9769D68FF6323098F57AB3D" box="[151,892,1843,1867]" location="Nagaura-higata" pageId="36" pageNumber="37" specimenCount="2">
<specimenCount id="9D63FD00F9769D68FF6323098D1DAB3D" box="[151,310,1843,1867]" count="2" pageId="36" pageNumber="37" type="generic">Two specimens</specimenCount>
from
<location id="8EBA6052F9769D68FE8223098E39AB3D" LSID="urn:lsid:plazi:treatment:03CC879FF9769D64FF63229C8FC6ADFF:8EBA6052F9769D68FE8223098E39AB3D" box="[374,530,1843,1867]" name="Nagaura-higata" pageId="36" pageNumber="37">Nagaura-higata</location>
<location id="8EBA6052F9769D68FE8223098E39AB3D" LSID="urn:lsid:plazi:treatment:03CC879FF9769D66FF63229C8E96A923:8EBA6052F9769D68FE8223098E39AB3D" box="[374,530,1843,1867]" name="Nagaura-higata" pageId="36" pageNumber="37">Nagaura-higata</location>
(WMNH-2024-INV-193, 194) and
</materialsCitation>
<materialsCitation id="3B0D3CD4F9769D68FC7023098EA9AB1F" location="Bay" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Kagoshima">
<specimenCount id="9D63FD00F9769D68FC702309883BAB3D" box="[900,1040,1843,1867]" count="1" pageId="36" pageNumber="37" type="generic">one specimen</specimenCount>
from
<collectingRegion id="49A1F86BF9769D68FBA5230988EFAB3D" box="[1105,1220,1843,1867]" country="Japan" name="Kagoshima" pageId="36" pageNumber="37">Kagoshima</collectingRegion>
<location id="8EBA6052F9769D68FB38230988DEAB3D" LSID="urn:lsid:plazi:treatment:03CC879FF9769D64FF63229C8FC6ADFF:8EBA6052F9769D68FB38230988DEAB3D" box="[1228,1269,1843,1867]" name="Bay" pageId="36" pageNumber="37" stateProvince="Kagoshima">Bay</location>
<location id="8EBA6052F9769D68FB38230988DEAB3D" LSID="urn:lsid:plazi:treatment:03CC879FF9769D66FF63229C8E96A923:8EBA6052F9769D68FB38230988DEAB3D" box="[1228,1269,1843,1867]" name="Bay" pageId="36" pageNumber="37" stateProvince="Kagoshima">Bay</location>
(WMNH-2022- INV-154) were identified as new to science
</materialsCitation>
.
@ -135,7 +139,7 @@ Tentacles possessing sensory cups, situated on basal part of stem, not only on o
, WMNH-2024-INV-193, 194.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9779D64FF3322C38FC6ADFF" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38" type="description">
<subSubSection id="C37F6502F9779D64FF3322C38CE9A89F" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38" type="description">
<paragraph id="8BDA3689F9779D69FF3322C388A2AB09" blockId="37.[151,1437,1497,1920]" pageId="37" pageNumber="38">
<emphasis id="B911EA9BF9779D69FF3322C3888FAB65" bold="true" box="[199,1188,1785,1811]" pageId="37" pageNumber="38">
Description of
@ -143,7 +147,7 @@ Description of
(WMNH-2022-INV-154: middle-size depressed specimen).
</emphasis>
External and internal morphologies as follows (
<tableCitation id="C6E70332F9779D69FE4923278E38AB41" box="[445,531,1821,1847]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="37" pageNumber="38">Table 1</tableCitation>
<tableCitation id="C6E70332F9779D69FE4923278E38AB41" box="[445,531,1821,1847]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="37" pageNumber="38">Table 1</tableCitation>
): 5 year-old preserved specimen pale peanut color (
<figureCitation id="135E2A0CF9779D69FB962327889CAB41" box="[1122,1207,1821,1847]" captionStart="FIGURE 18" captionStartId="37.[151,250,1319,1343]" captionTargetBox="[151,1436,181,1295]" captionTargetId="figure-20@37.[151,1436,181,1295]" captionTargetPageId="37" captionText="FIGURE 18. External view of a depressed specimen collected from Iso beach inner coast of Kagoshima Bay, Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, lateral view, anterior left; B, oral-side view of the tentacles; C, outer-side view of the tentacles." figureDoi="http://doi.org/10.5281/zenodo.14703570" httpUri="https://zenodo.org/record/14703570/files/figure.png" pageId="37" pageNumber="38">Fig. 18</figureCitation>
). Tentacles 12, each with 9 digits of 4 pairs and unpaired 1. On oral-side of tentacle stem, about 3040 sensory cups, also on outer oral-side tentacle stem, about 2040 sensory cups present densely arranged in a V- shape band.
@ -163,14 +167,14 @@ SEM images of the ossicles extracted from a specimen collected from Iso Beach, K
</caption>
<paragraph id="8BDA3689F9749D6AFF3323198DFBABDF" blockId="38.[151,1436,1827,1961]" pageId="38" pageNumber="39">
Polian vesicle in medio-ventral position, fusiform, single (
<tableCitation id="C6E70332F9749D6AFCA523198F8FAB4B" box="[849,932,1827,1853]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="38" pageNumber="39">Table 1</tableCitation>
<tableCitation id="C6E70332F9749D6AFCA523198F8FAB4B" box="[849,932,1827,1853]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="38" pageNumber="39">Table 1</tableCitation>
), stone canal single and thread-like, attached to narrow space between basal parts of tentacle ampullae at mediodorsal (
<emphasis id="B911EA9BF9749D6AFC3423728FC2AB14" box="[960,1001,1864,1890]" italics="true" pageId="38" pageNumber="39">IR5</emphasis>
), suddenly turned upward and forward, resulted in it facing forward, distally with moldy surface madreporite. Ciliated funnels in one crowded band along left dorsal interradius (
<emphasis id="B911EA9BF9749D6AFE6323B58D94ABDF" box="[407,447,1935,1961]" italics="true" pageId="38" pageNumber="39">IR3</emphasis>
).
</paragraph>
<caption id="DF1A6601F9759D6BFF6C24A28957ACC7" box="[152,1404,152,177]" pageId="39" pageNumber="40" startId="39.[152,244,152,176]" targetType="table">
<caption id="DF1A6601F9759D6BFF6C24A28957ACC7" box="[152,1404,152,177]" pageId="39" pageNumber="40" startId="39.[152,244,152,176]" targetBox="[163,1418,192,1867]" targetIsTable="true" targetPageId="39" targetType="table">
<paragraph id="8BDA3689F9759D6BFF6C24A28957ACC7" blockId="39.[152,1404,152,177]" box="[152,1404,152,177]" pageId="39" pageNumber="40">
<emphasis id="B911EA9BF9759D6BFF6C24A28D30ACC6" bold="true" box="[152,283,152,176]" pageId="39" pageNumber="40">TABLE 11.</emphasis>
Measurements (mean±SD μm) and counts of ossicles of three specimens of
@ -618,6 +622,306 @@ ai, (see, Materials and Methods) Indicators for ossicle shapes, and one param
</paragraph>
<paragraph id="8BDA3689F97A9D64FF3324AD8CF1AC8F" blockId="40.[151,1436,151,393]" pageId="40" pageNumber="41">Inside body, two tufts of gonad tubules attached with both sides of anterior dorsal mesentery. Intestine canal lacking loop, upper and under sides of intestine canal left blank, no whitish tubule organs adhered to intestine canal.</paragraph>
<paragraph id="8BDA3689F97A9D64FF3325398FC6ADFF" blockId="40.[151,1436,151,393]" pageId="40" pageNumber="41">Body wall smooth surface, soft but thick, covered by a surface layer like &quot;intestinal villi,&quot; with ossicle arranging latitudinal layout of all as sets of anchor and plate. Both sides along five longitudinal muscles with distal ends of anchor ossicles facing towards interradial area, however ossicles on radial body walls upon longitudinal muscles and interradial body walls have distal ends randomly facing either direction.</paragraph>
<paragraph id="8BDA3689F97A9D64FF6325E18E86AD82" blockId="40.[151,685,474,501]" box="[151,685,474,501]" pageId="40" pageNumber="41">
<heading id="D09281E5F97A9D64FF6325E18E86AD82" bold="true" box="[151,685,474,501]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis id="B911EA9BF97A9D64FF6325E18E86AD82" bold="true" box="[151,685,474,501]" pageId="40" pageNumber="41">The present results of ossicle morphologies</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F97A9D64FF6326198E76AF63" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
Body wall ossicles anchors and anchor plates (
<figureCitation id="135E2A0CF97A9D64FD4026198F69AE4B" box="[692,834,547,573]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19B; D</figureCitation>
,
<tableCitation id="C6E70332F97A9D64FCBB26198F9FAE4B" box="[847,948,547,573]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
). Anchor plates ranged within 85123 μm in length. Size of perforations not noticeably variable between central area and marginal area, and instead almost constant between central area and marginal area, because of only few marginal perforations rarely equipped. Plates showed strong distortion, with approximate hexagonal arrangement perforations. Perforations of anchor plate ossicles mostly teethless, but seldomly possessing 12 minute teeth, while outer edge of all anchor plate ossicles lacking minute teeth. Two calculated indicators for anchor plates and a parameter varied;
<emphasis id="B911EA9BF97A9D64FB8926ED8886AE87" box="[1149,1197,727,753]" italics="true" pageId="40" pageNumber="41">APS</emphasis>
= 011.3;
<emphasis id="B911EA9BF97A9D64FAD126ED8966AE86" box="[1317,1357,727,752]" italics="true" pageId="40" pageNumber="41">Ntp</emphasis>
= 02;
<emphasis id="B911EA9BF97A9D64FF6326C18CE2AF63" box="[151,201,763,789]" italics="true" pageId="40" pageNumber="41">APT</emphasis>
= 011.8% (
<figureCitation id="135E2A0CF97A9D64FEAD26C18DC9AF63" box="[345,482,763,789]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19B; D</figureCitation>
,
<tableCitation id="C6E70332F97A9D64FE1926C18E64AF63" box="[493,591,763,789]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
).
</paragraph>
<paragraph id="8BDA3689F97A9D64FF3327258DA0AFBF" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
Anchor ossicles ranged within 140188 μm in length (
<figureCitation id="135E2A0CF97A9D64FCDE27258F98AF4F" box="[810,947,799,825]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19B; D</figureCitation>
,
<tableCitation id="C6E70332F97A9D64FC4927258834AF4F" box="[957,1055,799,825]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
). Most anchor arms (bills) devoid of minute teeth on their outer tips, numbers of teeth very small within 04 (mostly
<quantity id="4C9D9B6CF97A9D64FBBB277988B3AF2B" box="[1103,1176,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" metricValueMax="5.08" metricValueMin="0.0" pageId="40" pageNumber="41" unit="in" value="1.0" valueMax="2.0" valueMin="0.0">02 in</quantity>
one arm of the anchor ossicle), and anchor ossicles of this specimen possessed narrow stem and also relatively narrow anchor arm width, with
<emphasis id="B911EA9BF97A9D64FF2627B18D23AFD3" box="[210,264,907,933]" italics="true" pageId="40" pageNumber="41">ASW</emphasis>
= 22.728.3%;
<emphasis id="B911EA9BF97A9D64FE3627B18DD7AFD3" box="[450,508,907,933]" italics="true" pageId="40" pageNumber="41">AEW</emphasis>
= 38.555.1%. T-shaped basal end of the anchor ossicle of this species crescent-shaped (
<figureCitation id="135E2A0CF97A9D64FF0027958D55AFBF" box="[244,382,943,969]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19B; D</figureCitation>
).
</paragraph>
<paragraph id="8BDA3689F97A9D64FF3327E98D4BA80B" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
Body longitudinal muscle, granule ossicles of this specimen ranged within 2154 μm in length (
<figureCitation id="135E2A0CF97A9D64FAFB27E989BCAF9B" box="[1295,1431,979,1005]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19C; E</figureCitation>
,
<tableCitation id="C6E70332F97A9D64FF6327CD8CD0A867" box="[151,251,1015,1041]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
). Both C-shaped and O-shaped granule ossicles, mostly flattened, distorted and roundish triangle form, rarely not distorted and usual oval form. The calculated indicator for granule proportions were varied within
<emphasis id="B911EA9BF97A9D64FA83202189B7A843" box="[1399,1436,1051,1077]" italics="true" pageId="40" pageNumber="41">GP</emphasis>
= 29.5123.0%;
<emphasis id="B911EA9BF97A9D64FEA720058DA5A82F" box="[339,398,1087,1113]" italics="true" pageId="40" pageNumber="41">GCO</emphasis>
= 46.152.9% and did not have noticeable differences in amounts between the O-shape and C-shape granules.
</paragraph>
<paragraph id="8BDA3689F97A9D64FF3320BD8CE9A89F" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
Tentacle ossicles rods, often with complex shape, ranging within 6285 μm in length (
<figureCitation id="135E2A0CF97A9D64FB7B20BD88DDA8D7" box="[1167,1270,1159,1185]" captionStart="FIGURE 19" captionStartId="38.[151,250,1613,1637]" captionTargetBox="[155,1433,181,1581]" captionTargetId="figure-29@38.[155,1433,181,1589]" captionTargetPageId="38" captionText="FIGURE 19. SEM images of the ossicles extracted from a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022-INV-154): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703572" httpUri="https://zenodo.org/record/14703572/files/figure.png" pageId="40" pageNumber="41">Fig. 19A</figureCitation>
,
<tableCitation id="C6E70332F97A9D64FAF520BD897DA8D7" box="[1281,1366,1159,1185]" captionStart="TABLE 9" captionStartId="32.[152,243,152,176]" captionTargetPageId="32" captionText="TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov." pageId="40" pageNumber="41">Table 9</tableCitation>
). Two calculated indicators for rod proportions and complexities varied within
<emphasis id="B911EA9BF97A9D64FC2820918FD5A8B2" box="[988,1022,1195,1220]" italics="true" pageId="40" pageNumber="41">RP</emphasis>
= 9.621.5%;
<emphasis id="B911EA9BF97A9D64FB58209188FBA8B3" box="[1196,1232,1195,1221]" italics="true" pageId="40" pageNumber="41">RC</emphasis>
=2.717.8 (
<tableCitation id="C6E70332F97A9D64FAA820918C9FA89F" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="40" pageNumber="41">Table 11</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F97A9D66FF3320C98DA0ADA7" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="etymology">
<paragraph id="8BDA3689F97A9D64FF3320C989A0A90F" blockId="40.[151,1437,546,1401]" pageId="40" pageNumber="41">
<emphasis id="B911EA9BF97A9D64FF3320C98D65A97B" bold="true" box="[199,334,1267,1293]" pageId="40" pageNumber="41">Etymology.</emphasis>
The species name
<taxonomicName id="4C654D0AF97A9D64FDB520C98E80A97B" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[577,683,1267,1293]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="40" pageNumber="41" phylum="Echinodermata" rank="species" species="neglectus">
<emphasis id="B911EA9BF97A9D64FDB520C98E80A97B" box="[577,683,1267,1293]" italics="true" pageId="40" pageNumber="41">neglectus</emphasis>
</taxonomicName>
, alluding to Y.Y.s misidentification of the
<typeStatus id="54DE882BF97A9D64FB3620C9890FA97B" box="[1218,1316,1267,1293]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
specimen (WMNH-2022-INV-154). Previously, this specimen was reported as
<taxonomicName id="4C654D0AF97A9D64FC67212D8FD1A947" box="[915,1018,1303,1329]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="40" pageNumber="41" phylum="Echinodermata" rank="species" species="ooplax">
<emphasis id="B911EA9BF97A9D64FC67212D8FD1A947" box="[915,1018,1303,1329]" italics="true" pageId="40" pageNumber="41">P. ooplax</emphasis>
</taxonomicName>
with young and small ossicles, to Mr. Soichiro Tashima of Kusunoki-Shizenkan (N.P.O. Kusunoki Nature Study Museum), Aira City,
<collectingRegion id="49A1F86BF97A9D64FB2821018974A923" box="[1244,1375,1339,1365]" country="Japan" name="Kagoshima" pageId="40" pageNumber="41">Kagoshima</collectingRegion>
. This misjudgment was made due to Y.Y.s unsympathetic view on ossicle morphologies at that time (
<date id="FFDB1049F97A9D64FB3221658954A90F" box="[1222,1407,1375,1401]" pageId="40" pageNumber="41" value="2022-12">December 2022</date>
).
</paragraph>
<paragraph id="8BDA3689F97A9D64FF6321F08F36A992" blockId="40.[151,797,1482,1509]" box="[151,797,1482,1509]" pageId="40" pageNumber="41">
<heading id="D09281E5F97A9D64FF6321F08F36A992" bold="true" box="[151,797,1482,1509]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis id="B911EA9BF97A9D64FF6321F08F36A992" bold="true" box="[151,797,1482,1509]" pageId="40" pageNumber="41">Observation on the morphologies of calcareous ring</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F97A9D64FF632229884DAA97" blockId="40.[151,1436,1555,1761]" pageId="40" pageNumber="41">
Calcareous plates 12, mostly firmly bound (
<figureCitation id="135E2A0CF97A9D64FD8A22298EFBAA5B" box="[638,720,1555,1581]" captionStart="FIGURE 20" captionStartId="41.[151,250,585,609]" captionTargetBox="[163,1422,188,552]" captionTargetId="figure-20@41.[151,1435,181,561]" captionTargetPageId="41" captionText="FIGURE 20. Outer-surface view of calcareous ring of the calcareous ring plates of a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as Patinapta neglectus sp. nov., in the present study (holotype specimen, WMNH-2022- INV-154): Positions of the plates denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stands for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi="http://doi.org/10.5281/zenodo.14703574" httpUri="https://zenodo.org/record/14703574/files/figure.png" pageId="40" pageNumber="41">Fig. 20</figureCitation>
), with slight anterior projection and deep posterior depression in interradial plates, while only posterior depression in radial plates. Only medio-ventral radial plate perforated in its anterior half, relatively large perforation. Other four radial plates devoid of perforation, however, there were narrow “interradial perforations” situated in the connections between 4 plates situated in right and left dorso-lateral positions (
<emphasis id="B911EA9BF97A9D64FF5522998DACAACA" box="[161,391,1699,1725]" italics="true" pageId="40" pageNumber="41">IR4; RIII; IR3; RII</emphasis>
), where additional interradial plate (
<emphasis id="B911EA9BF97A9D64FCD022998FB2AACB" box="[804,921,1699,1725]" italics="true" pageId="40" pageNumber="41">IR4; IR3</emphasis>
) formed triangular shapes and adjacent radial plate (
<emphasis id="B911EA9BF97A9D64FF2A22FD8D14AA96" box="[222,319,1735,1760]" italics="true" pageId="40" pageNumber="41">RIII; RII</emphasis>
) form creviced squarish shapes, and weakly bound by their apical ends.
</paragraph>
<caption id="DF1A6601F97B9D65FF6326738DBFAF63" ID-DOI="http://doi.org/10.5281/zenodo.14703574" ID-Zenodo-Dep="14703574" httpUri="https://zenodo.org/record/14703574/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,585,609]" targetBox="[163,1422,188,552]" targetPageId="41" targetType="figure">
<paragraph id="8BDA3689F97B9D65FF6326738DBFAF63" blockId="41.[151,1436,585,789]" pageId="41" pageNumber="42">
<emphasis id="B911EA9BF97B9D65FF6326738D08AE17" bold="true" box="[151,291,585,609]" pageId="41" pageNumber="42">FIGURE 20.</emphasis>
Outer-surface view of calcareous ring of the calcareous ring plates of a specimen collected from Iso Beach, Kagoshima Bay Kagoshima, erected as
<taxonomicName id="4C654D0AF97B9D65FDDB26548ED0AEF3" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[559,763,621,645]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="B911EA9BF97B9D65FDDB26548ED0AEF3" box="[559,763,621,645]" italics="true" pageId="41" pageNumber="42">Patinapta neglectus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF97B9D65FCF626578F78AEF3" bold="true" box="[770,851,621,645]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A22257E0F97B9D65FCF626578F78AEF3" box="[770,851,621,645]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study (holotype specimen, WMNH-2022- INV-154): Positions of the plates denoted by Ludwigs [18891892 (1889), p156] system:
<emphasis id="B911EA9BF97B9D65FBDB26A8885AAEDF" box="[1071,1137,658,681]" italics="true" pageId="41" pageNumber="42">RIRV</emphasis>
and
<emphasis id="B911EA9BF97B9D65FB5726AB88D3AEDF" box="[1187,1272,657,681]" italics="true" pageId="41" pageNumber="42">IR1IR5</emphasis>
where
<emphasis id="B911EA9BF97B9D65FAB726A88979AEDF" box="[1347,1362,658,681]" italics="true" pageId="41" pageNumber="42">R</emphasis>
and
<emphasis id="B911EA9BF97B9D65FA7026A889B7AEDF" box="[1412,1436,658,681]" italics="true" pageId="41" pageNumber="42">IR</emphasis>
stands for positions for radius and interradius, respectively. Hence
<emphasis id="B911EA9BF97B9D65FCCD268C8F7AAEBB" box="[825,849,694,717]" italics="true" pageId="41" pageNumber="42">RI</emphasis>
, mid-ventral;
<emphasis id="B911EA9BF97B9D65FC14268C882BAEBB" box="[992,1024,694,717]" italics="true" pageId="41" pageNumber="42">RII</emphasis>
, left ventral;
<emphasis id="B911EA9BF97B9D65FB7C268C889AAEBB" box="[1160,1201,694,717]" italics="true" pageId="41" pageNumber="42">RIII</emphasis>
, right ventral;
<emphasis id="B911EA9BF97B9D65FAB3268C8945AEBB" box="[1351,1390,694,717]" italics="true" pageId="41" pageNumber="42">RIV</emphasis>
, left dorsal;
<emphasis id="B911EA9BF97B9D65FF1726E08D2AAE87" box="[227,257,730,753]" italics="true" pageId="41" pageNumber="42">RV</emphasis>
, right dorsal; and:
<emphasis id="B911EA9BF97B9D65FE3626E38DCDAE87" box="[450,486,729,753]" italics="true" pageId="41" pageNumber="42">IR1</emphasis>
, left ventral;
<emphasis id="B911EA9BF97B9D65FD9B26E38EBFAE87" box="[623,660,729,753]" italics="true" pageId="41" pageNumber="42">IR2</emphasis>
, right ventral;
<emphasis id="B911EA9BF97B9D65FCDE26E38FAFAE87" box="[810,900,729,753]" italics="true" pageId="41" pageNumber="42">IR3-IR3</emphasis>
, left dorso-lateral;
<emphasis id="B911EA9BF97B9D65FBB326E3888AAE87" box="[1095,1185,729,753]" italics="true" pageId="41" pageNumber="42">IR4-IR4</emphasis>
, right dorso-lateral;
<emphasis id="B911EA9BF97B9D65FA8526E089BEAE87" box="[1393,1429,730,753]" italics="true" pageId="41" pageNumber="42">IR5</emphasis>
, mid-dorsal. Anterior top.
</paragraph>
</caption>
<paragraph id="8BDA3689F97B9D65FF63275D8E9CAFF7" blockId="41.[151,695,871,898]" box="[151,695,871,898]" pageId="41" pageNumber="42">
<heading id="D09281E5F97B9D65FF63275D8E9CAFF7" bold="true" box="[151,695,871,898]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
<emphasis id="B911EA9BF97B9D65FF63275D8E9CAFF7" bold="true" box="[151,695,871,898]" pageId="41" pageNumber="42">
Ossicle variation among the
<typeStatus id="54DE882BF97B9D65FE0E27528E1BAFF4" box="[506,560,872,898]" pageId="41" pageNumber="42">type</typeStatus>
specimens
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F97B9D65FF63278A8E3AAA7F" blockId="41.[151,1437,943,2014]" pageId="41" pageNumber="42">
Among
<specimenCount id="9D63FD00F97B9D65FF0C27958D9FAFBF" box="[248,436,943,969]" count="3" pageId="41" pageNumber="42" type="generic">three specimens</specimenCount>
,
<specimenCount id="9D63FD00F97B9D65FE30278A8E5BAFBF" box="[452,624,943,970]" count="2" pageId="41" pageNumber="42" type="generic">two specimens</specimenCount>
(WMNH-2022-INV-154; 2024-INV-194) were mature and possessed gonad tubules, while
<specimenCount id="9D63FD00F97B9D65FE7C27EE8E0AAF9B" box="[392,545,979,1006]" count="1" pageId="41" pageNumber="42" type="generic">one specimen</specimenCount>
(WMNH-2024-INV-193) was immature and was the smallest specimen, which also possessed under-developed ossicles as below: short rod ossicles in tentacles, stout anchor and small anchor plate ossicles in body wall. In tentacles, simple short rod ossicles possess up to only two perforations and only four minute branches, anchor stem short and stout; the single basal tip of anchor plate ossicles with besides noticeable defections, resulting in the “table-tennis racket-form” those of congeners of the apodid genus
<taxonomicName id="4C654D0AF97B9D65FB56205989B3A8F6" authority="Ostergren" authorityName="Ostergren" authorityYear="1898" box="[1186,1432,1123,1152]" class="Holothuroidea" family="Synaptidae" genus="Labidoplax" kingdom="Animalia" order="Apodida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="genus">
<emphasis id="B911EA9BF97B9D65FB5620598909A80B" box="[1186,1314,1123,1149]" italics="true" pageId="41" pageNumber="42">Labidoplax</emphasis>
Ōstergren
</taxonomicName>
, as such feature was not observed from two mature specimens, which possessed normal ovoid anchor plates. Body wall ossicles anchors and anchor plates. Anchor plate ossicle length and calculated indicators of
<specimenCount id="9D63FD00F97B9D65FB13209189B6A8B3" box="[1255,1437,1195,1221]" count="3" pageId="41" pageNumber="42" type="generic">three specimens</specimenCount>
ranged within: 85123 μm in length,
<emphasis id="B911EA9BF97B9D65FDA320EA8EACA89C" box="[599,647,1232,1258]" italics="true" pageId="41" pageNumber="42">APS</emphasis>
=011.3,
<emphasis id="B911EA9BF97B9D65FD0220EA8F35A89F" box="[758,798,1232,1257]" italics="true" pageId="41" pageNumber="42">Ntp</emphasis>
=02,
<emphasis id="B911EA9BF97B9D65FC9F20EA8FB6A89C" box="[875,925,1232,1258]" italics="true" pageId="41" pageNumber="42">APT</emphasis>
=011.8%. Among these values, length and non—size dependance indicator of plate skewness
<emphasis id="B911EA9BF97B9D65FD2520CE8F2AA978" box="[721,769,1268,1294]" italics="true" pageId="41" pageNumber="42">APS</emphasis>
of anchor plate ossicles were not significantly different (
<emphasis id="B911EA9BF97B9D65FA7420CE89B7A97B" box="[1408,1436,1268,1293]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test), while frequency of teethed perforations
<emphasis id="B911EA9BF97B9D65FC6621228FEFA944" box="[914,964,1304,1330]" italics="true" pageId="41" pageNumber="42">APT</emphasis>
and their parameter
<emphasis id="B911EA9BF97B9D65FB52212288E5A947" box="[1190,1230,1304,1329]" italics="true" pageId="41" pageNumber="42">Ntp</emphasis>
were significantly different on the ventral side (
<emphasis id="B911EA9BF97B9D65FE2221068DD9A923" box="[470,498,1340,1365]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test) but not significantly different on dorsal side (
<emphasis id="B911EA9BF97B9D65FAC721068964A923" box="[1331,1359,1340,1365]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test). From these results, the anchor plate ossicles in the body wall of the present
<specimenCount id="9D63FD00F97B9D65FB1C216589B6A90F" box="[1256,1437,1375,1401]" count="3" pageId="41" pageNumber="42" type="generic">three specimens</specimenCount>
were not different in ossicle length or the skewness of the layout for perforations However, number of teethed perforation and their occupancies were different on the ventral body wall. Therefore, the length (about 85120 μm), numbers of teethed perforations (02) on dorsal side, and slightly distorted (up to 11.3) anchor plate ossicles can be appropriate diagnostic characters.
</paragraph>
<paragraph id="8BDA3689F97B9D65FF3322298CEEAB73" blockId="41.[151,1437,943,2014]" pageId="41" pageNumber="42">
Anchor ossicle length and calculated indicators ranged within: 108188 μm in length,
<emphasis id="B911EA9BF97B9D65FB3C222E88D5AA58" box="[1224,1278,1556,1582]" italics="true" pageId="41" pageNumber="42">ASW</emphasis>
=17.328.3%,
<emphasis id="B911EA9BF97B9D65FF6322028CFAAA24" box="[151,209,1592,1618]" italics="true" pageId="41" pageNumber="42">AEW</emphasis>
=29.655.4% (
<tableCitation id="C6E70332F97B9D65FE83220D8DF7AA24" box="[375,476,1591,1618]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="41" pageNumber="42">Table 11</tableCitation>
). Among these values, lengths, the indicator for anchor stem breadth
<emphasis id="B911EA9BF97B9D65FB0622028903AA24" box="[1266,1320,1592,1618]" italics="true" pageId="41" pageNumber="42">ASW</emphasis>
of anchor ossicles and for basal / distal ends width
<emphasis id="B911EA9BF97B9D65FD9F22668E8EAA00" box="[619,677,1628,1654]" italics="true" pageId="41" pageNumber="42">AEW</emphasis>
were significantly different (
<emphasis id="B911EA9BF97B9D65FC0522668826AA03" box="[1009,1037,1628,1653]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test, both in ventrally and dorsally), while number of the teeth was not significantly different (
<emphasis id="B911EA9BF97B9D65FBB222BA8849AAEF" box="[1094,1122,1664,1689]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test, both ventrally and dorsally). From these results, total shapes of anchor ossicles in the body wall of the present
<specimenCount id="9D63FD00F97B9D65FA9022998D27AA97" count="3" pageId="41" pageNumber="42" type="generic">three specimens</specimenCount>
cannot be appropriate keys, but the excessively low number of the counts for teeth can be the accurate key.
</paragraph>
<paragraph id="8BDA3689F97B9D66FF3323358859ACA3" blockId="41.[151,1437,943,2014]" lastBlockId="42.[151,1437,151,465]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
Longitudinal muscle involved both O-shaped and C-shaped granule ossicles, length and calculated indicators ranged within: 2854 μm length,
<emphasis id="B911EA9BF97B9D65FDEA230E8E68AB38" box="[542,579,1844,1870]" italics="true" pageId="41" pageNumber="42">GP</emphasis>
=28.9123.0%,
<emphasis id="B911EA9BF97B9D65FD0E230E8F1EAB38" box="[762,821,1844,1870]" italics="true" pageId="41" pageNumber="42">GCO</emphasis>
=33.366.7% (
<tableCitation id="C6E70332F97B9D65FC292309886FAB38" box="[989,1092,1843,1870]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="41" pageNumber="42">Table 11</tableCitation>
). Among these values, length of granule ossicles and calculated indicator for proportions
<emphasis id="B911EA9BF97B9D65FCB323628F47AB04" box="[839,876,1880,1906]" italics="true" pageId="41" pageNumber="42">GP</emphasis>
were different significantly (
<emphasis id="B911EA9BF97B9D65FB4B236288F0AB07" box="[1215,1243,1880,1905]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test, both in ventrally and dorsally), while the ratio C- or O-shaped granule ossicles
<emphasis id="B911EA9BF97B9D65FB67234688E5ABE0" box="[1171,1230,1916,1942]" italics="true" pageId="41" pageNumber="42">GCO</emphasis>
were not different significantly (
<emphasis id="B911EA9BF97B9D65FECE239A8D7DABCF" box="[314,342,1952,1977]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test, both ventrally and dorsally), but different significantly in
<emphasis id="B911EA9BF97B9D65FA95239A89B7ABCC" box="[1377,1436,1952,1978]" italics="true" pageId="41" pageNumber="42">GCO</emphasis>
(
<emphasis id="B911EA9BF97B9D65FF5523FE8C96ABAB" box="[161,189,1988,2013]" italics="true" pageId="41" pageNumber="42">Ps</emphasis>
&lt;0.05, χ
<superScript id="7C109BC1F97B9D65FEE823F98D0FABA7" attach="right" box="[284,292,1987,2001]" fontSize="6" pageId="41" pageNumber="42">2</superScript>
-multiple test, both in ventrally and dorsally). Therefore, the length and proportion of the granule ossicles cannot be appropriate keys, while ratio C- or O-shaped granule and the feature of granule ossicles being “flattened, distorted and roundish triangle form” of this species can be appropriate keys.
</paragraph>
<paragraph id="8BDA3689F9789D66FF3324E58DA0ADA7" blockId="42.[151,1437,151,465]" pageId="42" pageNumber="43">
Tentacle ossicles rods, length and calculated indicators of
<specimenCount id="9D63FD00F9789D66FCA024E58823AC8F" box="[852,1032,223,249]" count="3" pageId="42" pageNumber="43" type="generic">three specimens</specimenCount>
ranged within: 4498 μm in length,
<emphasis id="B911EA9BF9789D66FF6325398C92AD6A" box="[151,185,259,284]" italics="true" pageId="42" pageNumber="43">RP</emphasis>
=9.621.5%, and
<emphasis id="B911EA9BF9789D66FE7025398D83AD6B" box="[388,424,259,285]" italics="true" pageId="42" pageNumber="43">RC</emphasis>
=2.717.8. Among these values, length of rod ossicles, nonsize dependence complexities (
<emphasis id="B911EA9BF9789D66FF55251D8CEEAD37" box="[161,197,295,321]" italics="true" pageId="42" pageNumber="43">RC</emphasis>
) were significantly different (
<emphasis id="B911EA9BF9789D66FDF8251D8E03AD36" box="[524,552,295,320]" italics="true" pageId="42" pageNumber="43">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test), while nonsize dependence indicator for proportion (
<emphasis id="B911EA9BF9789D66FF5525718CE8AD12" box="[161,195,331,356]" italics="true" pageId="42" pageNumber="43">RP</emphasis>
) was not different significantly (
<emphasis id="B911EA9BF9789D66FDB325718E48AD12" box="[583,611,331,356]" italics="true" pageId="42" pageNumber="43">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test). From these results, the rod ossicles in the tentacles of the present
<specimenCount id="9D63FD00F9789D66FE5425558E7FADFF" box="[416,596,367,393]" count="3" pageId="42" pageNumber="43" type="generic">three specimens</specimenCount>
had similar proportions, simultaneously complexities and length increased by body growth (
<tableCitation id="C6E70332F9789D66FEA825A98DEAADDB" box="[348,449,403,429]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="42" pageNumber="43">Table 11</tableCitation>
). Therefore, the morphologies of the rod ossicles of this species cannot be appropriate diagnostic characters.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9789D66FF6326198DD6AFBF" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BDA3689F9789D66FF6326198D22AE4B" blockId="42.[151,265,547,573]" box="[151,265,547,573]" pageId="42" pageNumber="43">
<emphasis id="B911EA9BF9789D66FF6326198D22AE4B" bold="true" box="[151,265,547,573]" pageId="42" pageNumber="43">Remarks</emphasis>
</paragraph>
<paragraph id="8BDA3689F9789D66FF6326518DD6AFBF" blockId="42.[151,1437,619,969]" pageId="42" pageNumber="43">
In the present
<specimenCount id="9D63FD00F9789D66FECE26518D81AEF3" box="[314,426,619,645]" count="3" pageId="42" pageNumber="43" type="generic">three type</specimenCount>
materials, ossicle morphologies were noticeably different between matured
<typeStatus id="54DE882BF9789D66FAFE26518947AEF3" box="[1290,1388,619,645]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
and two small
<typeStatus id="54DE882BF9789D66FEF826B58D51AEDF" box="[268,378,655,681]" pageId="42" pageNumber="43" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D63FD00F9789D66FE7126B58E34AEDF" box="[389,543,655,681]" count="1" pageId="42" pageNumber="43" type="immature">one immature</specimenCount>
and one in the development-stage of tubules (
<tableCitation id="C6E70332F9789D66FBD726B5885DAEDF" box="[1059,1142,655,681]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="42" pageNumber="43">Table 1</tableCitation>
). From the present results of measurements and counts on ossicles (
<tableCitation id="C6E70332F9789D66FD9C26898EE7AEBB" box="[616,716,691,717]" captionStart="TABLE 11" captionStartId="39.[152,244,152,176]" captionTargetPageId="39" captionText="TABLE 11. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta neglectus sp. nov." pageId="42" pageNumber="43">Table 11</tableCitation>
), it can be assumed that several morphological changes relating with individual growth: i) in anchor ossicles, minute teeth on outer edge of anchor-bills take numbers larger in small animals than in large animals; ii) in anchor plate ossicles, teethed perforationstake larger number in small animals than in large animals; however, iii) in tentacle rods, complexity of rod-shape (
<emphasis id="B911EA9BF9789D66FB8C272588B7AF4F" box="[1144,1180,799,825]" italics="true" pageId="42" pageNumber="43">RC</emphasis>
) take values higher in large animals than in small animals, and complexity may not be increasing or decreasing by individual growth. In the previous nominal species, none of the species possessed calcareous ring only perforated on medio-ventral plate (Table 7-3), and this must be largest difference of the present new species among all the congeners of genus
<taxonomicName id="4C654D0AF9789D66FF6327958D2EAFBE" authorityName="Heding" authorityYear="1928" box="[151,261,943,968]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="42" pageNumber="43" phylum="Echinodermata" rank="genus">
<emphasis id="B911EA9BF9789D66FF6327958D2EAFBE" box="[151,261,943,968]" italics="true" pageId="42" pageNumber="43">Patinapta</emphasis>
</taxonomicName>
previously described.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9789D66FF6320208E96A923" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BDA3689F9789D66FF6320208D1BA842" blockId="42.[151,304,1050,1076]" box="[151,304,1050,1076]" pageId="42" pageNumber="43">
<emphasis id="B911EA9BF9789D66FF6320208D1BA842" bold="true" box="[151,304,1050,1076]" pageId="42" pageNumber="43">Distribution</emphasis>
</paragraph>
<paragraph id="8BDA3689F9789D66FF6320598E96A923" blockId="42.[151,1437,1123,1365]" pageId="42" pageNumber="43">
Only known from Kyushu Island and adjacent islands, Mature specimen has been collected only from Iso Beach (the type locality) on the inner coast of
<collectingRegion id="49A1F86BF9789D66FDA220BD8EFDA8D7" box="[598,726,1159,1185]" country="Japan" name="Kagoshima" pageId="42" pageNumber="43">Kagoshima</collectingRegion>
Bay (
<tableCitation id="C6E70332F9789D66FCEE20BD8FABA8D7" box="[794,896,1159,1185]" captionStart="TABLE 12" captionStartId="42.[152,244,1412,1436]" captionTargetPageId="42" captionText="TABLE 12. List of WMNH-INV specimens of Patinapta neglectus sp. nov., as of 2024." pageId="42" pageNumber="43">Table 12</tableCitation>
). At the type locality, the
<typeStatus id="54DE882BF9789D66FB5020BD892DA8D7" box="[1188,1286,1159,1185]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
material was collected from intertidal sandy beach with boulders. Among the specimens housed in WMNH-INV, including the present specimens, 3 bottles including
<specimenCount id="9D63FD00F9789D66FDBB20F48EF1A89F" box="[591,730,1230,1257]" count="5" pageId="42" pageNumber="43" type="generic">5 specimens</specimenCount>
were identified as this species (
<tableCitation id="C6E70332F9789D66FBB120F58887A89F" box="[1093,1196,1231,1257]" captionStart="TABLE 12" captionStartId="42.[152,244,1412,1436]" captionTargetPageId="42" captionText="TABLE 12. List of WMNH-INV specimens of Patinapta neglectus sp. nov., as of 2024." pageId="42" pageNumber="43">Table 12</tableCitation>
). Other than the type locality,
<specimenCount id="9D63FD00F9789D66FF0E20C98D56A97B" box="[250,381,1267,1293]" count="2" pageId="42" pageNumber="43" type="immature">2 immature</specimenCount>
and mature specimens were collected from Nagaura-higata flat, Amakusa City,
<collectingRegion id="49A1F86BF9789D66FAEA20C989B7A97B" box="[1310,1436,1267,1293]" country="Japan" name="Kumamoto" pageId="42" pageNumber="43">Kumamoto</collectingRegion>
and
<specimenCount id="9D63FD00F9789D66FF33212D8D6CA947" box="[199,327,1303,1329]" count="2" pageId="42" pageNumber="43" type="immature">2 immature</specimenCount>
specimens collected from Uchiura Bay, Kamikoshiki-jima Island, adjacent to
<collectingRegion id="49A1F86BF9789D66FB43212D8912A947" box="[1207,1337,1303,1329]" country="Japan" name="Kagoshima" pageId="42" pageNumber="43">Kagoshima</collectingRegion>
, Kyushu Island,
<collectingCountry id="F3727619F9789D66FF1E21018D06A923" box="[234,301,1339,1365]" name="Japan" pageId="42" pageNumber="43">Japan</collectingCountry>
, and were identified as this species.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,175 +0,0 @@
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<mods:title id="EA8098E1155D938ED7123E6505B338D8">First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan</mods:title>
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<mods:namePart id="C57CB4607847B9021C7A1ABC6E2B4279">Yamana, Yusuke</mods:namePart>
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<mods:affiliation id="2877B55338A3F06C0B7B06F72AC2D153">Wakayama Prefectural Museum of Natural History (WMNH), 1 Room Curators Head Office, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan</mods:affiliation>
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<subSubSection id="C37F6502F9789D66FF6322838EF2AA81" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8BDA3689F9789D66FF6322838DC7AAA5" blockId="42.[151,729,1721,1819]" box="[151,492,1721,1748]" pageId="42" pageNumber="43">
<heading id="D09281E5F9789D66FF6322838DC7AAA5" bold="true" box="[151,492,1721,1748]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="B911EA9BF9789D66FF6322838DC7AAA5" bold="true" box="[151,492,1721,1748]" pageId="42" pageNumber="43">
<taxonomicName id="4C654D0AF9789D66FF6322838DA3AAA2" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[151,392,1721,1748]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="42" pageNumber="43" phylum="Echinodermata" rank="species" species="edentatus" status="sp. nov.">
<emphasis id="B911EA9BF9789D66FF6322838DA3AAA2" bold="true" box="[151,392,1721,1748]" italics="true" pageId="42" pageNumber="43">Patinapta edentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257E0F9789D66FE7B22838DC7AAA5" box="[399,492,1721,1747]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BDA3689F9789D66FF6322E48EF2AA81" blockId="42.[151,729,1721,1819]" box="[151,729,1757,1784]" pageId="42" pageNumber="43">[New Japanese name: Hanashi-himo-ikari-namako]</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9789D66FF63233B89BFAB28" pageId="42" pageNumber="43" type="description">
<paragraph id="8BDA3689F9789D66FF63233B8DC3AB6D" blockId="42.[151,729,1721,1819]" box="[151,488,1793,1819]" pageId="42" pageNumber="43">
(
<figureCitation id="135E2A0CF9789D66FF6B233B8D0BAB6D" box="[159,288,1793,1819]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStartId-0="43.[151,250,1410,1434]" captionStartId-1="44.[151,250,1844,1868]" captionStartId-2="46.[151,250,1409,1433]" captionTargetBox-0="[151,1436,273,1387]" captionTargetBox-1="[155,1433,345,1813]" captionTargetBox-2="[163,1423,1106,1381]" captionTargetId-0="figure-55@43.[151,1436,273,1387]" captionTargetId-1="figure-81@44.[155,1433,345,1821]" captionTargetId-2="figure-368@46.[151,1435,1101,1386]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionTargetPageId-2="46" captionText-0="FIGURE 21. External view of a specimen collected from Kabira Bay, Ishigaki-jima Island, Okinawa, erected as Patinapta edentatus sp. nov., in the present study (holotype specimen, WMNH-2021-INV-973): A, lateral view, anterior left, dorsal top; A, living specimen from the same lot (from collector Dr. Taeko Kimura), with co-existing bivalves Anisodevonia ohshimai infesting body surface; B, oral-side view of the tentacles; C, outer-side view of the tentacles." captionText-1="FIGURE 22. SEM images of the ossicles extracted from a specimen collected from Kabira Bay, Ishigaki-jima Island, Okinawa, erected as Patinapta edentatus sp. nov., in the present study (holotype specimen, WMNH-2021-INV-973): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." captionText-2="FIGURE 23. Outer-surface view of calcareous ring of the calcareous ring plates of a specimen collected from Kabira Bay, Ishigaki-jima Island, Okinawa, erected as Patinapta edentatus sp. nov., in the present study (holotype specimen, WMNH-2021- INV-973). The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stands for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorsolateral; IR5, mid-dorsal. Anterior top." figureDoi-0="http://doi.org/10.5281/zenodo.14703576" figureDoi-1="http://doi.org/10.5281/zenodo.14703578" figureDoi-2="http://doi.org/10.5281/zenodo.14703580" httpUri-0="https://zenodo.org/record/14703576/files/figure.png" httpUri-1="https://zenodo.org/record/14703578/files/figure.png" httpUri-2="https://zenodo.org/record/14703580/files/figure.png" pageId="42" pageNumber="43">Figs 2123</figureCitation>
,
<tableCitation id="C6E70332F9789D66FEDF233B8DA1AB6D" box="[299,394,1793,1819]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="42" pageNumber="43">Tables 1</tableCitation>
;
<tableCitation id="C6E70332F9789D66FE6C233B8D9CAB6D" box="[408,439,1793,1819]" captionStart="TABLE 13" captionStartId="45.[152,244,225,249]" captionText="TABLE 13. Measurements (mean±SD μm) and counts of ossicles of the present specimens of Patinapta edentatus sp. nov." pageId="42" pageNumber="43">13</tableCitation>
;
<tableCitation id="C6E70332F9789D66FE37233B8DCBAB6D" box="[451,480,1793,1819]" captionStart="TABLE 14" captionStartId="47.[152,244,1700,1724]" captionText="TABLE 14. List of the WMNH-INV specimens of Patinapta edentatus sp. nov. as of 2024." pageId="42" pageNumber="43">14</tableCitation>
)
</paragraph>
<paragraph id="8BDA3689F9789D66FF63237C89BFAB28" blockId="42.[151,1428,1862,1886]" box="[151,1428,1862,1886]" pageId="42" pageNumber="43">
<materialsCitation id="3B0D3CD4F9789D66FF63237C89BBAB28" box="[151,1424,1862,1886]" location="Kabira Bay" municipality="Ishigaki-jima Island" pageId="42" pageNumber="43" specimenCount="3">
<specimenCount id="9D63FD00F9789D66FF63237C8D68AB28" box="[151,323,1862,1886]" count="3" pageId="42" pageNumber="43" type="generic">Three specimens</specimenCount>
from
<location id="8EBA6052F9789D66FE74237C8DDEAB28" LSID="urn:lsid:plazi:treatment:03CC879FF9789D60FF63228388ACAD6A:8EBA6052F9789D66FE74237C8DDEAB28" box="[384,501,1862,1886]" municipality="Ishigaki-jima Island" name="Kabira Bay" pageId="42" pageNumber="43">Kabira Bay</location>
,
<collectingMunicipality id="6BBEACF3F9789D66FE0B237C8EE0AB28" box="[511,715,1862,1886]" pageId="42" pageNumber="43">Ishigaki-jima Island</collectingMunicipality>
(WMNH-2021-INV-972, 973, 974) were identified as new to science
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9789D67FF6323B68D73AC8F" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8BDA3689F9789D67FF6323B68D73AC8F" blockId="42.[151,1436,1932,2030]" lastBlockId="43.[151,1437,151,249]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="B911EA9BF9789D66FF6323B68D39ABD0" bold="true" box="[151,274,1932,1958]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
Outer edge of anchor plate ossicles and inner edge of their perforations lacking teeth anywhere, with large seven perforations arranged in symmetrical order in the distal part, while minute perforations sporadically scattered in basal part. Tentacles possess small numbers of simple rod ossicles and large numbers of simple elongated C-shaped granule ossicles. Mostly elongated C-shaped granules equipped in longitudinal muscles, rarely with Oshaped granules among them. Ciliated funnels attached to inner side body wall along midline of right dorso-lateral interradius (
<emphasis id="B911EA9BF9799D67FEEB24E58D63AC8F" box="[287,328,223,249]" italics="true" pageId="43" pageNumber="44">IR4</emphasis>
).
</paragraph>
</subSubSection>
<caption id="DF1A6601F9799D67FF6321B8886AAA70" ID-DOI="http://doi.org/10.5281/zenodo.14703576" ID-Zenodo-Dep="14703576" httpUri="https://zenodo.org/record/14703576/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,250,1410,1434]" targetBox="[151,1436,273,1387]" targetPageId="43" targetType="figure">
<paragraph id="8BDA3689F9799D67FF6321B8886AAA70" blockId="43.[151,1436,1410,1543]" pageId="43" pageNumber="44">
<emphasis id="B911EA9BF9799D67FF6321B88D09A9EC" bold="true" box="[151,290,1410,1434]" pageId="43" pageNumber="44">FIGURE 21.</emphasis>
External view of a specimen collected from Kabira Bay, Ishigaki-jima Island, Okinawa, erected as
<taxonomicName id="4C654D0AF9799D67FACC21B98CD3A9C8" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="43" pageNumber="44" phylum="Echinodermata" rank="species" species="edentatus" status="sp. nov.">
<emphasis id="B911EA9BF9799D67FACC21B98CD3A9C8" italics="true" pageId="43" pageNumber="44">Patinapta edentatus</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9799D67FF0B219D8D7AA9C9" bold="true" box="[255,337,1447,1471]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="A22257E0F9799D67FF0B219D8D7AA9C9" box="[255,337,1447,1471]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study (holotype specimen, WMNH-2021-INV-973): A, lateral view, anterior left, dorsal top; A, living specimen from the same lot (from collector Dr. Taeko Kimura), with co-existing bivalves
<taxonomicName id="4C654D0AF9799D67FB5921F089B7A994" baseAuthorityName="Kawahara" baseAuthorityYear="1942" box="[1197,1436,1482,1506]" class="Bivalvia" family="Lasaeidae" genus="Anisodevonia" kingdom="Animalia" order="Galeommatida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="ohshimai">
<emphasis id="B911EA9BF9799D67FB5921F089B7A994" box="[1197,1436,1482,1506]" italics="true" pageId="43" pageNumber="44">Anisodevonia ohshimai</emphasis>
</taxonomicName>
infesting body surface; B, oral-side view of the tentacles; C, outer-side view of the tentacles.
</paragraph>
</caption>
<subSubSection id="C37F6502F9799D67FF33226388F2AA05" box="[199,1241,1624,1651]" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BDA3689F9799D67FF33226388F2AA05" blockId="43.[151,1437,1624,2011]" box="[199,1241,1624,1651]" pageId="43" pageNumber="44">
<emphasis id="B911EA9BF9799D67FF3322638D78AA04" bold="true" box="[199,339,1624,1651]" pageId="43" pageNumber="44">Type series.</emphasis>
<typeStatus id="54DE882BF9799D67FEAE22638DEEAA05" box="[346,453,1625,1651]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
, WMNH-2021-INV-973;
<specimenCount id="9D63FD00F9799D67FD1322638FADAA05" box="[743,902,1625,1651]" count="2" pageId="43" pageNumber="44" type="generic" typeStatus="paratype">two paratypes</specimenCount>
, WMNH-2021-INV-972, 974.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9799D60FF3322468EACAC8F" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
<paragraph id="8BDA3689F9799D67FF3322468907AB51" blockId="43.[151,1437,1624,2011]" pageId="43" pageNumber="44">
<emphasis id="B911EA9BF9799D67FF332246888AAAE0" bold="true" box="[199,1185,1660,1687]" pageId="43" pageNumber="44">
Description of
<typeStatus id="54DE882BF9799D67FE8922478DCFAAE1" box="[381,484,1661,1687]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
(WMNH-2021-INV-973: middle-size complete specimen).
</emphasis>
External and internal morphologies as follows (
<tableCitation id="C6E70332F9799D67FE45229B8E28AACD" box="[433,515,1697,1723]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="43" pageNumber="44">Table 1</tableCitation>
): Specimen preserved for20 years; pale misty-rose (
<figureCitation id="135E2A0CF9799D67FBC9229B8888AACD" box="[1085,1187,1697,1723]" captionStart="FIGURE 21" captionStartId="43.[151,250,1410,1434]" captionTargetBox="[151,1436,273,1387]" captionTargetId="figure-55@43.[151,1436,273,1387]" captionTargetPageId="43" captionText="FIGURE 21. External view of a specimen collected from Kabira Bay, Ishigaki-jima Island, Okinawa, erected as Patinapta edentatus sp. nov., in the present study (holotype specimen, WMNH-2021-INV-973): A, lateral view, anterior left, dorsal top; A, living specimen from the same lot (from collector Dr. Taeko Kimura), with co-existing bivalves Anisodevonia ohshimai infesting body surface; B, oral-side view of the tentacles; C, outer-side view of the tentacles." figureDoi="http://doi.org/10.5281/zenodo.14703576" httpUri="https://zenodo.org/record/14703576/files/figure.png" pageId="43" pageNumber="44">Fig. 21A</figureCitation>
), based on information from the collector color was transparent rose in living state (
<figureCitation id="135E2A0CF9799D67FCCB22FF8FBAAAA9" box="[831,913,1733,1759]" captionStart="FIGURE 21" captionStartId="43.[151,250,1410,1434]" captionTargetBox="[151,1436,273,1387]" captionTargetId="figure-55@43.[151,1436,273,1387]" captionTargetPageId="43" captionText="FIGURE 21. External view of a specimen collected from Kabira Bay, Ishigaki-jima Island, Okinawa, erected as Patinapta edentatus sp. nov., in the present study (holotype specimen, WMNH-2021-INV-973): A, lateral view, anterior left, dorsal top; A, living specimen from the same lot (from collector Dr. Taeko Kimura), with co-existing bivalves Anisodevonia ohshimai infesting body surface; B, oral-side view of the tentacles; C, outer-side view of the tentacles." figureDoi="http://doi.org/10.5281/zenodo.14703576" httpUri="https://zenodo.org/record/14703576/files/figure.png" pageId="43" pageNumber="44">Fig. 21</figureCitation>
A). Tentacles 12, each with 9 digits, including four pairs of smaller digits equipped along lateral-stem of tentacle and large one digit equipped on distal end of tentacle. On the oral side of tentacle stem, about 1020 sensory cups sporadically arranged in a U-shape.
</paragraph>
<paragraph id="8BDA3689F9799D67FF33230B891FAB19" blockId="43.[151,1437,1624,2011]" pageId="43" pageNumber="44">
Polian vesicle single, fusiform with thread-like tip,
<quantity id="4C9D9B6CF9799D67FCE4230B8F5DAB3D" box="[784,886,1841,1867]" metricMagnitude="-3" metricUnit="m" metricValue="4.12" pageId="43" pageNumber="44" unit="mm" value="4.12">4.12 mm</quantity>
(
<tableCitation id="C6E70332F9799D67FC71230B8FF1AB3D" box="[901,986,1841,1867]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="43" pageNumber="44">Table 1</tableCitation>
). Ciliated funnels occur sporadically in one row along right dorso-lateral interradius (
<emphasis id="B911EA9BF9799D67FD63236F8E94AB19" box="[663,703,1877,1903]" italics="true" pageId="43" pageNumber="44">IR4</emphasis>
). Stone canal undetected, lacking or too small to detect.
</paragraph>
<paragraph id="8BDA3689F9799D67FF3323438FB6ABC1" blockId="43.[151,1437,1624,2011]" pageId="43" pageNumber="44">Inside body, two tufts of gonad tubules attached to both sides of anterior dorsal mesentery, none of the whitish tubule organs adhered to intestine canal. Intestine canal lacking loop.</paragraph>
<paragraph id="8BDA3689F9799D60FF3323FB8EACAC8F" blockId="43.[151,1437,1624,2011]" lastBlockId="44.[151,1437,151,285]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">Body wall coarse surface, rigid and thin, densely packed with ossicles arranging latitudinal layout of all as sets of anchor and plate. Both sides along five longitudinal muscles with distal ends of anchor ossicles facing towards interradial area, however ossicles on radial body walls upon longitudinal muscles and interradial body walls have distal ends randomly facing either direction.</paragraph>
</subSubSection>
<subSubSection id="C37F6502F97E9D60FF33253988ACAD6A" box="[199,1159,259,285]" pageId="44" pageNumber="45" type="etymology">
<paragraph id="8BDA3689F97E9D60FF33253988ACAD6A" blockId="44.[151,1437,151,285]" box="[199,1159,259,285]" pageId="44" pageNumber="45">
<emphasis id="B911EA9BF97E9D60FF3325398D65AD6B" bold="true" box="[199,334,259,285]" pageId="44" pageNumber="45">Etymology.</emphasis>
The species name
<taxonomicName id="4C654D0AF97E9D60FDD725398EA5AD6B" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[547,654,259,285]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="44" pageNumber="45" phylum="Echinodermata" rank="species" species="edentatus">
<emphasis id="B911EA9BF97E9D60FDD725398EA5AD6B" box="[547,654,259,285]" italics="true" pageId="44" pageNumber="45">edentatus</emphasis>
</taxonomicName>
alluding their teeth-less anchor plate ossicle.
</paragraph>
</subSubSection>
</treatment>
</document>