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<document id="040057C0428724F02ECD81B7428C60F3" ID-ISSN="1732-2421" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1723062243441" checkinUser="felipe" docAuthor="Salesa, Manuel J., Sánchez, Israel M. &amp; Morales, Jorge" docDate="2004" docId="038C1A2E72084256B7263D9BFAF1FDD0" docLanguage="en" docName="ActaPalaeontolPol.49.2.189-196.pdf" docOrigin="Acta Palaeontologica Polonica 49 (2)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Sinohippus sampelayoi" docType="treatment" docVersion="2" lastPageNumber="192" masterDocId="FFB5625672094255B40A3846FFE1FF9C" masterDocTitle="Presence of the Asian horse Sinohippus in the Miocene of Europe" masterLastPageNumber="196" masterPageNumber="189" pageNumber="190" updateTime="1724889021027" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="97CF1D3CB183CB0B9B0B21CDA4CCF3DA">Presence of the Asian horse Sinohippus in the Miocene of Europe</mods:title>
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<mods:namePart id="675D7104307D124C8E31EC9B48ED8DA6">Salesa, Manuel J.</mods:namePart>
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<mods:namePart id="14946CF60D6EB0EECCC9F4370BE48F09">Sánchez, Israel M.</mods:namePart>
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<mods:namePart id="80358BBEB96E9211B3073F79DF9CD662">Morales, Jorge</mods:namePart>
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<mods:date id="17BE97164AE0D6E8F76192C053821497">2004</mods:date>
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<treatment id="038C1A2E72084256B7263D9BFAF1FDD0" LSID="urn:lsid:plazi:treatment:038C1A2E72084256B7263D9BFAF1FDD0" httpUri="http://treatment.plazi.org/id/038C1A2E72084256B7263D9BFAF1FDD0" lastPageId="3" lastPageNumber="192" pageId="1" pageNumber="190">
<subSubSection id="C33FF8B372084254B7263D9BFC9EF980" pageId="1" pageNumber="190" type="nomenclature">
<paragraph id="8B9AAB3872084254B7263D9BFC9EF980" blockId="1.[812,1419,1319,1597]" pageId="1" pageNumber="190">
<heading id="D0D21C5472084254B7263D9BFC9EF980" fontSize="12" level="2" pageId="1" pageNumber="190" reason="9">
<taxonomicName id="4C25D0BB72084254B7263D9BFC9AF980" authority="(Villalta and Crusafont, 1945)" baseAuthorityName="Villalta and Crusafont" baseAuthorityYear="1945" class="Mammalia" family="Equidae" genus="Sinohippus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="190" phylum="Chordata" rank="species" species="sampelayoi">
<emphasis id="B951772A72084254B7263D9BFBB5FA65" box="[812,1108,1501,1529]" italics="true" pageId="1" pageNumber="190">Sinohippus sampelayoi</emphasis>
(
<bibRefCitation id="EFB4D6C972084254B06F3D9AFC90F980" author="Villalta, J. F. &amp; Crusafont, M." pageId="1" pageNumber="190" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont, 1945</bibRefCitation>
)
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33FF8B372084256B7263E6EFAF1FDD0" lastPageId="3" lastPageNumber="192" pageId="1" pageNumber="190" type="description">
<paragraph id="8B9AAB3872084254B7263E6EFC6DF9A1" blockId="1.[812,1419,1319,1597]" box="[812,908,1576,1597]" pageId="1" pageNumber="190">
<figureCitation id="131EB7BD72084254B7263E6EFC69F9A1" box="[812,904,1576,1597]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="1.[102,135,1363,1383]" captionStartId-1="2.[102,135,1977,1997]" captionStartId-2="3.[102,135,1975,1995]" captionStartId-3="4.[102,135,758,778]" captionTargetBox-0="[128,747,237,1321]" captionTargetBox-1="[123,1464,225,1950]" captionTargetBox-2="[102,774,223,1948]" captionTargetBox-3="[102,775,227,720]" captionTargetId-0="figure-526@1.[123,753,233,1325]" captionTargetId-1="figure-10@2.[123,1464,225,1950]" captionTargetId-2="figure-471@3.[102,775,223,1948]" captionTargetId-3="figure-640@4.[102,775,227,720]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="Fig. 1. Cladogram proposed by Sánchez et al. (1998) for the Iberian Anchitheriinae, showing the relationships between the two lineages of Anchitherium. Abbreviations: A., Anchitherium; S., Sinohippus." captionText-1="Fig. 2. Sinohippus sampelayoi, IGME 1175M, holotype. Fragment of right hemimandible with p2, d2, p3, d3, p4, d4, m1, m2, and m3. In occlusal (A), buccal (B), and lingual (C) views." captionText-2="Fig. 3. Sinohippus sampelayoi, IPS 2001. Fragment of left hemimandible with p4, m1 and m2. In occlusal (A), lingual (B), and buccal (C) views." captionText-3="Fig. 4. Sinohippus sampelayoi, MNCN 50110. Occlusal view of fragment of right maxilla with P3P4." pageId="1" pageNumber="190">Figs. 14</figureCitation>
.
</paragraph>
<paragraph id="8B9AAB3872084254B7263E09FA2FF9F8" blockId="1.[812,1486,1615,2025]" box="[812,1486,1615,1636]" pageId="1" pageNumber="190">
<emphasis id="B951772A72084254B7263E09FC6DF9F8" box="[812,908,1615,1636]" italics="true" pageId="1" pageNumber="190">Synonyms</emphasis>
:
<taxonomicName id="4C25D0BB72084254B7943E09FA2BF9F8" authority="Villalta and Crusafont, 1945" authorityName="Villalta and Crusafont" authorityYear="1945" box="[926,1482,1615,1636]" class="Mammalia" family="Equidae" genus="Anchitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="190" phylum="Chordata" rank="species" species="sampelayoi">
<emphasis id="B951772A72084254B7943E09FB78F9F8" box="[926,1177,1615,1636]" italics="true" pageId="1" pageNumber="190">Anchitherium sampelayoi</emphasis>
<bibRefCitation id="EFB4D6C972084254B0AF3E09FA2BF9F8" author="Villalta, J. F. &amp; Crusafont, M." box="[1189,1482,1615,1636]" pageId="1" pageNumber="190" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont, 1945</bibRefCitation>
</taxonomicName>
:
</paragraph>
<paragraph id="8B9AAB3872084254B7583E2DFA2FF91C" blockId="1.[812,1486,1615,2025]" box="[850,1486,1643,1664]" pageId="1" pageNumber="190">
figs. 14;
<taxonomicName id="4C25D0BB72084254B7B73E2DFA2FF91C" authority="Abusch-Siewert, 1983" authorityName="Abusch-Siewert" authorityYear="1983" box="[957,1486,1643,1664]" class="Mammalia" family="Equidae" genus="Anchitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="1" pageNumber="190" phylum="Chordata" rank="subSpecies" species="ezquerrae" subSpecies="sampelayoi">
<emphasis id="B951772A72084254B7B73E2DFAC3F91C" box="[957,1314,1643,1664]" italics="true" pageId="1" pageNumber="190">Anchitherium ezquerrae sampelayoi</emphasis>
<bibRefCitation id="EFB4D6C972084254B1273E2DFA2FF91C" author="Abusch-Siewert, S." box="[1325,1486,1643,1664]" pageId="1" pageNumber="190" pagination="1 - 361" refId="ref4018" refString="Abusch-Siewert, S. 1983. Gebissmorphologische Untersuchungen an eurasiatischen Anchitherien (Equidae, Mammalia) unter besonderer Berucksichtigung der Fundstelle Sandelzhausen. Courier Forschungsinstitut Senckenberg 62: 1 - 361." type="journal article" year="1983">AbuschSiewert,</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B9AAB3872084254B7583EC1FC39F900" blockId="1.[812,1486,1615,2025]" box="[850,984,1671,1692]" pageId="1" pageNumber="190">1983: pl. 3: 2.</paragraph>
<paragraph id="8B9AAB3872084254B7263EEDFA20F940" blockId="1.[812,1486,1615,2025]" pageId="1" pageNumber="190">
<materialsCitation id="3B4DA16572084254B7263EEDFA20F940" collectionCode="IGME" pageId="1" pageNumber="190" specimenCode="IGME 1175" specimenCount="1" typeStatus="holotype">
<emphasis id="B951772A72084254B7263EEDFC65F95C" box="[812,900,1707,1728]" italics="true" pageId="1" pageNumber="190">
<typeStatus id="549E159A72084254B7263EEDFC65F95C" box="[812,900,1707,1728]" pageId="1" pageNumber="190" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCode id="DB83034372084254B79A3EEDFBE3F95C" box="[912,1026,1707,1728]" collectionCode="IGME" pageId="1" pageNumber="190">IGME 1175</specimenCode>
M (
<figureCitation id="131EB7BD72084254B02E3EEDFBBFF95C" box="[1060,1118,1707,1728]" captionStart="Fig" captionStartId="2.[102,135,1977,1997]" captionTargetBox="[123,1464,225,1950]" captionTargetId="figure-10@2.[123,1464,225,1950]" captionTargetPageId="2" captionText="Fig. 2. Sinohippus sampelayoi, IGME 1175M, holotype. Fragment of right hemimandible with p2, d2, p3, d3, p4, d4, m1, m2, and m3. In occlusal (A), buccal (B), and lingual (C) views." pageId="1" pageNumber="190">Fig. 2</figureCitation>
), Right hemimandible with p2, d2, p3, d3, p4, d4, m1, m2, and m3 (see
<bibRefCitation id="EFB4D6C972084254B0593E81FABAF940" author="Villalta, J. F. &amp; Crusafont, M." box="[1107,1371,1735,1756]" pageId="1" pageNumber="190" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont 1945</bibRefCitation>
:figs. 13).
</materialsCitation>
</paragraph>
<paragraph id="8B9AAB3872084254B7263EAFFB64F886" blockId="1.[812,1486,1615,2025]" pageId="1" pageNumber="190">
<materialsCitation id="3B4DA16572084254B7263EAFFB60F886" collectionCode="IPS-" pageId="1" pageNumber="190" specimenCode="IPS-70" specimenCount="1" typeStatus="paratype">
<emphasis id="B951772A72084254B7263EAFFC65F962" box="[812,900,1769,1790]" italics="true" pageId="1" pageNumber="190">
<typeStatus id="549E159A72084254B7263EAFFC65F962" box="[812,900,1769,1790]" pageId="1" pageNumber="190" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCode id="DB83034372084254B79B3EAFFC36F962" box="[913,983,1769,1790]" collectionCode="IPS-" pageId="1" pageNumber="190">IPS70</specimenCode>
, Fragment of left hemimandible with d2 and d3 (see
<bibRefCitation id="EFB4D6C972084254B7263F43FBDAF886" author="Villalta, J. F. &amp; Crusafont, M." box="[812,1083,1797,1818]" pageId="1" pageNumber="190" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont 1945</bibRefCitation>
: fig. 4)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9AAB3872084254B7263F61FBCFF837" blockId="1.[812,1486,1615,2025]" pageId="1" pageNumber="190">
<emphasis id="B951772A72084254B7263F61FBC0F8A0" box="[812,1057,1831,1852]" italics="true" pageId="1" pageNumber="190">Other material examined</emphasis>
: IPS2001, fragment of hemimandible with p4, m1 and m2 (
<figureCitation id="131EB7BD72084254B7C13F04FBE4F8CB" box="[971,1029,1858,1879]" captionStart="Fig" captionStartId="3.[102,135,1975,1995]" captionTargetBox="[102,774,223,1948]" captionTargetId="figure-471@3.[102,775,223,1948]" captionTargetPageId="3" captionText="Fig. 3. Sinohippus sampelayoi, IPS 2001. Fragment of left hemimandible with p4, m1 and m2. In occlusal (A), lingual (B), and buccal (C) views." pageId="1" pageNumber="190">Fig. 3</figureCitation>
). There is a cast of an unpublished fragment of maxilla with P3 and P4 (
<figureCitation id="131EB7BD72084254B01A3F18FBA8F8EF" box="[1040,1097,1886,1907]" captionStart="Fig" captionStartId="4.[102,135,758,778]" captionTargetBox="[102,775,227,720]" captionTargetId="figure-640@4.[102,775,227,720]" captionTargetPageId="4" captionText="Fig. 4. Sinohippus sampelayoi, MNCN 50110. Occlusal view of fragment of right maxilla with P3P4." pageId="1" pageNumber="190">Fig. 4</figureCitation>
), deposited in the MNCN, which give the only information about the upper dentition of this species; the original specimen belongs to MLS.
</paragraph>
<paragraph id="8B9AAB3872084254B7263FFEFC36F875" blockId="1.[812,1486,1615,2025]" pageId="1" pageNumber="190">
<emphasis id="B951772A72084254B7263FFEFC4CF851" box="[812,941,1976,1997]" italics="true" pageId="1" pageNumber="190">Type locality</emphasis>
: Nombrevilla1 (CalatayudDaroca Basin, province of Zaragoza,
<collectingCountry id="F332EBA872084254B79B3F92FC2DF875" box="[913,972,2004,2025]" name="Spain" pageId="1" pageNumber="190">Spain</collectingCountry>
).
</paragraph>
<caption id="DF5AFBB0720B4257B46C3FFFFE7DF875" pageId="2" pageNumber="191" startId="2.[102,135,1977,1997]" targetBox="[123,1464,225,1950]" targetPageId="2" targetType="figure">
<paragraph id="8B9AAB38720B4257B46C3FFFFE7DF875" blockId="2.[102,1485,1977,2025]" pageId="2" pageNumber="191">
Fig. 2.
<taxonomicName id="4C25D0BB720B4257B4AD3FFFFE15F851" authority=", IGME 1175" authorityName=", IGME" authorityYear="1175" box="[167,500,1977,1997]" class="Mammalia" family="Equidae" genus="Sinohippus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="2" pageNumber="191" phylum="Chordata" rank="species" species="sampelayoi">
<emphasis id="B951772A720B4257B4AD3FFFFE98F851" box="[167,377,1977,1997]" italics="true" pageId="2" pageNumber="191">Sinohippus sampelayoi</emphasis>
, IGME 1175
</taxonomicName>
M, holotype. Fragment of right hemimandible with p2, d2, p3, d3, p4, d4, m1, m2, and m3. In occlusal (
<emphasis id="B951772A720B4257B1A53FFFFA5EF851" bold="true" box="[1455,1471,1977,1997]" pageId="2" pageNumber="191">A</emphasis>
), buccal (
<emphasis id="B951772A720B4257B4A43F93FF5CF874" bold="true" box="[174,189,2005,2024]" pageId="2" pageNumber="191">B</emphasis>
), and lingual (
<emphasis id="B951772A720B4257B5493F93FEB2F875" bold="true" box="[323,339,2005,2025]" pageId="2" pageNumber="191">C</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8B9AAB38720A4256B72638A1FA5BFF60" blockId="3.[812,1466,231,252]" box="[812,1466,231,252]" pageId="3" pageNumber="192">
<emphasis id="B951772A720A4256B72638A1FCB0FF60" box="[812,849,231,252]" italics="true" pageId="3" pageNumber="192">Age</emphasis>
: Early Vallesian (basal Late Miocene) (zone MN 9 of
<bibRefCitation id="EFB4D6C9720A4256B14F38A1FA4EFF60" author="Mein, P." box="[1349,1455,231,252]" pageId="3" pageNumber="192" pagination="77 - 81" refId="ref5509" refString="Mein, P. 1975. Biozonation du neogene Mediterranee a partir des mammiferes. Proceedings of the VI Congress of the Regional Committee on Mediterranean Neogene Stratigraphy, Bratislava, 77 - 81." type="book chapter" year="1975">
<collectingRegion id="49E165DA720A4256B14F38A1FA99FF60" box="[1349,1400,231,252]" country="United States of America" name="Maine" pageId="3" pageNumber="192">Mein</collectingRegion>
1975
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9AAB38720A4256B726394AFC01FEA1" blockId="3.[812,1485,268,317]" pageId="3" pageNumber="192">
<emphasis id="B951772A720A4256B726394AFC27FEBD" box="[812,966,268,289]" italics="true" pageId="3" pageNumber="192">Other localities</emphasis>
: Soblay and DouélaFontaine (
<collectingCountry id="F332EBA8720A4256B102394AFAAFFEBD" box="[1288,1358,268,289]" name="France" pageId="3" pageNumber="192">France</collectingCountry>
), and EçmeAkçaköi (
<collectingCountry id="F332EBA8720A4256B780396EFC34FEA1" box="[906,981,296,317]" name="Turkey" pageId="3" pageNumber="192">Turkey</collectingCountry>
).
</paragraph>
<paragraph id="8B9AAB38720A4256B7263912FAF1FDD0" blockId="3.[812,1486,339,588]" pageId="3" pageNumber="192">
<emphasis id="B951772A720A4256B7263912FC7AFEF7" box="[812,923,340,363]" italics="true" pageId="3" pageNumber="192">Diagnosis</emphasis>
.—In
<bibRefCitation id="EFB4D6C9720A4256B7D73915FAB2FEF0" author="Villalta, J. F. &amp; Crusafont, M." box="[989,1363,339,364]" pageId="3" pageNumber="192" pagination="51 - 82" refId="ref6300" refString="Villalta, J. F. and Crusafont, M. 1945. Un Anchitherium en el Pontiense espanol. Anchitherium sampelayoi, nova sp. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 14: 51 - 82." type="journal article" year="1945">Villalta and Crusafont (1945: 78)</bibRefCitation>
: “terminal, derived
<taxonomicName id="4C25D0BB720A4256B78C3932FBFDFE10" authorityName="von Meyer" authorityYear="1844" box="[902,1052,372,396]" class="Mammalia" family="Equidae" genus="Anchitherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="3" pageNumber="192" phylum="Chordata" rank="genus">
<emphasis id="B951772A720A4256B78C3932FBFDFE10" box="[902,1052,372,396]" italics="true" pageId="3" pageNumber="192">Anchitherium</emphasis>
</taxonomicName>
of large size, with marked tendency to hypsodonty, basal cingulum almost obsolete, mainly in the juvenile teeth; more evident homeodonty than in the primitive forms, milk teeth highly reduced, special shape of d2”; additionally: upper premolars with marked buccal cingulum and absence of lingual cingulum, hypostyle without connection with the metalophe, and absence of crochet.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="827D7B2DF6BA88DC76210F618065B61B" ID-ISSN="1732-2421" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1723062957633" checkinUser="felipe" docAuthor="Valiukevičius, Juozas" docDate="2004" docId="782E87AD0D42FF8CFCBC74F68385A3FA" docLanguage="en" docName="ActaPalaeontolPol.49.1.147-160.pdf" docOrigin="Acta Palaeontologica Polonica 49 (1)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Rohonilepis breviornatus Valiukevičius, 2004, gen. et sp. nov." docType="treatment" docVersion="2" lastPageNumber="158" masterDocId="8417FFD50D4AFF87FF9071468168A706" masterDocTitle="New Wenlock-Pridoli (Silurian) acanthodian fishes from Lithuania" masterLastPageNumber="160" masterPageNumber="147" pageNumber="155" updateTime="1724888488487" updateUser="carolina" zenodo-license-document="CC-BY-4.0"> <document id="45AF5B7ADFA0F4BF98CF2214188F68FF" ID-DOI="10.5281/zenodo.13406145" ID-ISSN="1732-2421" ID-Zenodo-Dep="13406145" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1723062957633" checkinUser="felipe" docAuthor="Valiukevičius, Juozas" docDate="2004" docId="782E87AD0D42FF8CFCBC74F68385A3FA" docLanguage="en" docName="ActaPalaeontolPol.49.1.147-160.pdf" docOrigin="Acta Palaeontologica Polonica 49 (1)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Rohonilepis breviornatus Valiukevičius, 2004, gen. et sp. nov." docType="treatment" docVersion="3" lastPageNumber="158" masterDocId="8417FFD50D4AFF87FF9071468168A706" masterDocTitle="New Wenlock-Pridoli (Silurian) acanthodian fishes from Lithuania" masterLastPageNumber="160" masterPageNumber="147" pageNumber="155" updateTime="1724888823245" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FC07EDF2E6D9E457098F5910F95A845C">New Wenlock-Pridoli (Silurian) acanthodian fishes from Lithuania</mods:title> <mods:title id="0C7AAD220DA0BB354205CBAE95B11B57">New Wenlock-Pridoli (Silurian) acanthodian fishes from Lithuania</mods:title>
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<paragraph id="F03836BB0D42FF8FFCBC74F68452A2CB" blockId="8.[812,1338,1456,1518]" box="[812,1338,1456,1485]" pageId="8" pageNumber="155"> <paragraph id="F03836BB0D42FF8FFCBC74F68452A2CB" blockId="8.[812,1338,1456,1518]" box="[812,1338,1456,1485]" pageId="8" pageNumber="155">
<heading id="AB7081D70D42FF8FFCBC74F68452A2CB" box="[812,1338,1456,1485]" fontSize="12" level="2" pageId="8" pageNumber="155" reason="9"> <heading id="AB7081D70D42FF8FFCBC74F68452A2CB" box="[812,1338,1456,1485]" fontSize="12" level="2" pageId="8" pageNumber="155" reason="9">
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<paragraph id="F03836BB0D42FF8FFCBC749E8202A2E8" blockId="8.[812,1338,1456,1518]" box="[812,874,1496,1518]" pageId="8" pageNumber="155"> <paragraph id="F03836BB0D42FF8FFCBC749E8202A2E8" blockId="8.[812,1338,1456,1518]" box="[812,874,1496,1518]" pageId="8" pageNumber="155">
<figureCitation id="68BC2A3E0D42FF8FFCBC749E820EA2E8" box="[812,870,1496,1518]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="8" pageNumber="155">Fig. 8</figureCitation> <figureCitation id="68BC2A3E0D42FF8FFCBC749E820EA2E8" box="[812,870,1496,1518]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." figureDoi="http://doi.org/10.5281/zenodo.13406189" httpUri="https://zenodo.org/record/13406189/files/figure.png" pageId="8" pageNumber="155">Fig. 8</figureCitation>
. .
</paragraph> </paragraph>
<paragraph id="F03836BB0D42FF8FFCBC74B9847AA113" blockId="8.[812,1485,1535,1654]" box="[812,1298,1535,1557]" pageId="8" pageNumber="155"> <paragraph id="F03836BB0D42FF8FFCBC74B9847AA113" blockId="8.[812,1485,1535,1654]" box="[812,1298,1535,1557]" pageId="8" pageNumber="155">
@ -58,7 +60,7 @@
: :
<specimenCode id="A0219EC00D42FF8FFC0174B9828DA113" box="[913,997,1535,1557]" collectionCode="LIGG" pageId="8" pageNumber="155">LIGG 25</specimenCode> <specimenCode id="A0219EC00D42FF8FFC0174B9828DA113" box="[913,997,1535,1557]" collectionCode="LIGG" pageId="8" pageNumber="155">LIGG 25</specimenCode>
A2459, trunk scale ( A2459, trunk scale (
<figureCitation id="68BC2A3E0D42FF8FFB2B74B9846FA113" box="[1211,1287,1535,1557]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="8" pageNumber="155">Fig. 8C</figureCitation> <figureCitation id="68BC2A3E0D42FF8FFB2B74B9846FA113" box="[1211,1287,1535,1557]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." figureDoi="http://doi.org/10.5281/zenodo.13406189" httpUri="https://zenodo.org/record/13406189/files/figure.png" pageId="8" pageNumber="155">Fig. 8C</figureCitation>
). ).
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@ -96,7 +98,7 @@ decorated, ornamented.
</taxonomicName> </taxonomicName>
with tiny scales sculptured by 6 to 10 short, sharp, symmetrical subparallel anterior ridges turning anteriorly toward the neckbase junction. The two longest posteriorly pointing ridges can form an elevated medial area separating narrow, lowered lateral folds. Crown dentine contains mediumbranched linear ascending and simple widened radial vascular canals over the flatpyramidal base, that is composed of dense acellular bone. with tiny scales sculptured by 6 to 10 short, sharp, symmetrical subparallel anterior ridges turning anteriorly toward the neckbase junction. The two longest posteriorly pointing ridges can form an elevated medial area separating narrow, lowered lateral folds. Crown dentine contains mediumbranched linear ascending and simple widened radial vascular canals over the flatpyramidal base, that is composed of dense acellular bone.
</paragraph> </paragraph>
<caption id="A4F866330D43FF8EFFF6773282BDA1F1" pageId="9" pageNumber="156" startId="9.[102,135,1652,1671]" targetBox="[104,1482,229,1625]" targetPageId="9" targetType="figure"> <caption id="A4F866330D43FF8EFFF6773282BDA1F1" ID-DOI="http://doi.org/10.5281/zenodo.13406183" ID-Zenodo-Dep="13406183" httpUri="https://zenodo.org/record/13406183/files/figure.png" pageId="9" pageNumber="156" startId="9.[102,135,1652,1671]" targetBox="[104,1482,229,1625]" targetPageId="9" targetType="figure">
<paragraph id="F03836BB0D43FF8EFFF6773282BDA1F1" blockId="9.[102,1486,1652,1784]" pageId="9" pageNumber="156"> <paragraph id="F03836BB0D43FF8EFFF6773282BDA1F1" blockId="9.[102,1486,1652,1784]" pageId="9" pageNumber="156">
Fig. 7. Fig. 7.
<taxonomicName id="37874D380D43FF8EFF34773280FFA181" authorityName="Valiukevičius" authorityYear="2004" box="[164,407,1652,1671]" class="Acanthodii" family="Ischnacanthidae" genus="Bracteatacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthiformes" pageId="9" pageNumber="156" phylum="Chordata" rank="species" species="assiduus" status="gen. et sp. nov."> <taxonomicName id="37874D380D43FF8EFF34773280FFA181" authorityName="Valiukevičius" authorityYear="2004" box="[164,407,1652,1671]" class="Acanthodii" family="Ischnacanthidae" genus="Bracteatacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthiformes" pageId="9" pageNumber="156" phylum="Chordata" rank="species" species="assiduus" status="gen. et sp. nov.">
@ -140,14 +142,14 @@ Fig. 7.
in length and scale width varies from in length and scale width varies from
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. The crown plate is horizontal, with up to onethird of the posterior unsculptured part overhanging the base. A quarter to onethird of the crown length has 6 to 10 short, sharp anterior ridges with a symmetrical profile, which fade posteriorly. Crown ridges may be straight ( . The crown plate is horizontal, with up to onethird of the posterior unsculptured part overhanging the base. A quarter to onethird of the crown length has 6 to 10 short, sharp anterior ridges with a symmetrical profile, which fade posteriorly. Crown ridges may be straight (
<figureCitation id="68BC2A3E0D43FF8EFB10762985BFA08E" box="[1152,1239,1903,1928]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="9" pageNumber="156">Fig. 8D</figureCitation> <figureCitation id="68BC2A3E0D43FF8EFB10762985BFA08E" box="[1152,1239,1903,1928]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." figureDoi="http://doi.org/10.5281/zenodo.13406189" httpUri="https://zenodo.org/record/13406189/files/figure.png" pageId="9" pageNumber="156">Fig. 8D</figureCitation>
) or slightly curved towards midscale ( ) or slightly curved towards midscale (
<figureCitation id="68BC2A3E0D43FF8EFC6376D68506A0AE" box="[1011,1134,1936,1961]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="9" pageNumber="156">Fig. 8B, C</figureCitation> <figureCitation id="68BC2A3E0D43FF8EFC6376D68506A0AE" box="[1011,1134,1936,1961]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." figureDoi="http://doi.org/10.5281/zenodo.13406189" httpUri="https://zenodo.org/record/13406189/files/figure.png" pageId="9" pageNumber="156">Fig. 8B, C</figureCitation>
). The holotypelike specimens, with subparallel ridges are more abundant in samples studied, however, scales with almost parallel ridges also are known. Ridges are characteristically downwardly sloped anteriorly, sometimes reaching the base of the neck. Grooves between ridges are moderately deep and wide. Some scales have narrow lowered lateral ledges ( ). The holotypelike specimens, with subparallel ridges are more abundant in samples studied, however, scales with almost parallel ridges also are known. Ridges are characteristically downwardly sloped anteriorly, sometimes reaching the base of the neck. Grooves between ridges are moderately deep and wide. Some scales have narrow lowered lateral ledges (
<figureCitation id="68BC2A3E0D40FF8DFE0176D7836EA0AC" box="[401,518,1937,1962]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="10" pageNumber="157">Fig. 8A, C</figureCitation> <figureCitation id="68BC2A3E0D40FF8DFE0176D7836EA0AC" box="[401,518,1937,1962]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." figureDoi="http://doi.org/10.5281/zenodo.13406189" httpUri="https://zenodo.org/record/13406189/files/figure.png" pageId="10" pageNumber="157">Fig. 8A, C</figureCitation>
), separated from the remainder of the crown by the two longest, posteriorly pointing ridges. Moderately high to low neck usually has linear rows of pores. The scale base slightly protrudes beyond the crown anterolaterally, but may be also displaced far in advance of the anterior margin of the crown. ), separated from the remainder of the crown by the two longest, posteriorly pointing ridges. Moderately high to low neck usually has linear rows of pores. The scale base slightly protrudes beyond the crown anterolaterally, but may be also displaced far in advance of the anterior margin of the crown.
</paragraph> </paragraph>
<caption id="A4F866330D40FF8DFFF677E383CAA00A" pageId="10" pageNumber="157" startId="10.[102,135,1701,1720]" targetBox="[107,1482,225,1674]" targetPageId="10" targetType="figure"> <caption id="A4F866330D40FF8DFFF677E383CAA00A" ID-DOI="http://doi.org/10.5281/zenodo.13406189" ID-Zenodo-Dep="13406189" httpUri="https://zenodo.org/record/13406189/files/figure.png" pageId="10" pageNumber="157" startId="10.[102,135,1701,1720]" targetBox="[107,1482,225,1674]" targetPageId="10" targetType="figure">
<paragraph id="F03836BB0D40FF8DFFF677E383CAA00A" blockId="10.[102,1486,1701,1805]" pageId="10" pageNumber="157"> <paragraph id="F03836BB0D40FF8DFFF677E383CAA00A" blockId="10.[102,1486,1701,1805]" pageId="10" pageNumber="157">
Fig. 8. Fig. 8.
<taxonomicName id="37874D380D40FF8DFF3B77E380FBA1BE" authorityName="Valiukevičius" authorityYear="2004" box="[171,403,1701,1720]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="10" pageNumber="157" phylum="Chordata" rank="species" species="breviornatus" status="gen. et sp. nov."> <taxonomicName id="37874D380D40FF8DFF3B77E380FBA1BE" authorityName="Valiukevičius" authorityYear="2004" box="[171,403,1701,1720]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="10" pageNumber="157" phylum="Chordata" rank="species" species="breviornatus" status="gen. et sp. nov.">
@ -184,14 +186,14 @@ Histologic microstructure has not been sufficiently studied yet because of the p
<typeStatus id="2F3C88190D40FF8DFBD97694851FA0EC" box="[1097,1143,2002,2026]" pageId="10" pageNumber="157">type</typeStatus> <typeStatus id="2F3C88190D40FF8DFBD97694851FA0EC" box="[1097,1143,2002,2026]" pageId="10" pageNumber="157">type</typeStatus>
. At least, 6 growth lamellae in crowns composed of dentine can be counted ( . At least, 6 growth lamellae in crowns composed of dentine can be counted (
<figureCitation id="68BC2A3E0D41FF8CFDFA71A183D7A606" box="[618,703,231,256]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="11" pageNumber="158">Fig. 8F</figureCitation> <figureCitation id="68BC2A3E0D41FF8CFDFA71A183D7A606" box="[618,703,231,256]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." figureDoi="http://doi.org/10.5281/zenodo.13406189" httpUri="https://zenodo.org/record/13406189/files/figure.png" pageId="11" pageNumber="158">Fig. 8F</figureCitation>
), with the oldest lamella unusually large. The vascular system is poorly defined due to remineralization of tissues and only short segments ( ), with the oldest lamella unusually large. The vascular system is poorly defined due to remineralization of tissues and only short segments (
<figureCitation id="68BC2A3E0D41FF8CFE8B70038006A658" box="[283,366,325,350]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="11" pageNumber="158">Fig. 8E</figureCitation> <figureCitation id="68BC2A3E0D41FF8CFE8B70038006A658" box="[283,366,325,350]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." figureDoi="http://doi.org/10.5281/zenodo.13406189" httpUri="https://zenodo.org/record/13406189/files/figure.png" pageId="11" pageNumber="158">Fig. 8E</figureCitation>
<figureCitation id="68BC2A3E0D41FF8CFEFE7014801FA665" box="[366,375,338,355]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="11" pageNumber="158"> <figureCitation id="68BC2A3E0D41FF8CFEFE7014801FA665" box="[366,375,338,355]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="11" pageNumber="158">
<subScript id="6C0334FE0D41FF8CFEFE7014801FA665" attach="left" box="[366,375,338,355]" fontSize="7" pageId="11" pageNumber="158">2</subScript> <subScript id="6C0334FE0D41FF8CFEFE7014801FA665" attach="left" box="[366,375,338,355]" fontSize="7" pageId="11" pageNumber="158">2</subScript>
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, F) of ascending and other vascular canals are visible. Ascending canals are moderately branched and mostly linear along growth lamellae. Radial vascular canals are simple and located just over the basal surface ( , F) of ascending and other vascular canals are visible. Ascending canals are moderately branched and mostly linear along growth lamellae. Radial vascular canals are simple and located just over the basal surface (
<figureCitation id="68BC2A3E0D41FF8CFD2070E28269A6BA" box="[688,769,420,445]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="11" pageNumber="158">Fig. 8F</figureCitation> <figureCitation id="68BC2A3E0D41FF8CFD2070E28269A6BA" box="[688,769,420,445]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. AD. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25A2457. B. LIGG 25A2458. C. Holotype, LIGG 25A2459. D. LIGG 25A2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." figureDoi="http://doi.org/10.5281/zenodo.13406189" httpUri="https://zenodo.org/record/13406189/files/figure.png" pageId="11" pageNumber="158">Fig. 8F</figureCitation>
, right side). Durodentine has not been definitely recognized, but may be present. Acellular basal bone is dense and forms a flatpyramid below the crown. , right side). Durodentine has not been definitely recognized, but may be present. Acellular basal bone is dense and forms a flatpyramid below the crown.
</paragraph> </paragraph>
<paragraph id="F03836BB0D41FF8CFFF6736D8096A49D" blockId="11.[102,775,231,1277]" pageId="11" pageNumber="158"> <paragraph id="F03836BB0D41FF8CFFF6736D8096A49D" blockId="11.[102,775,231,1277]" pageId="11" pageNumber="158">

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<mods:identifier id="8C0BF240364F5CEC417164718A3155B1" type="ISSN">1732-2421</mods:identifier>
<mods:identifier id="484984096FC4F2F875D268A59BB4B601" type="Zenodo-Dep">13406145</mods:identifier>
</mods:mods> </mods:mods>
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<subSubSection id="B89D65300D4BFF86FFF67508835AA36D" box="[102,562,1102,1131]" pageId="1" pageNumber="148" type="nomenclature"> <subSubSection id="B89D65300D4BFF86FFF67508835AA36D" box="[102,562,1102,1131]" pageId="1" pageNumber="148" type="nomenclature">
<paragraph id="F03836BB0D4BFF86FFF67508835AA36D" blockId="1.[102,562,1102,1163]" box="[102,562,1102,1131]" pageId="1" pageNumber="148"> <paragraph id="F03836BB0D4BFF86FFF67508835AA36D" blockId="1.[102,562,1102,1163]" box="[102,562,1102,1131]" pageId="1" pageNumber="148">
<heading id="AB7081D70D4BFF86FFF67508835AA36D" box="[102,562,1102,1131]" fontSize="12" level="2" pageId="1" pageNumber="148" reason="9"> <heading id="AB7081D70D4BFF86FFF67508835AA36D" box="[102,562,1102,1131]" fontSize="12" level="2" pageId="1" pageNumber="148" reason="9">
@ -47,9 +49,9 @@
</subSubSection> </subSubSection>
<subSubSection id="B89D65300D4BFF84FFF67533841DA1B5" lastPageId="3" lastPageNumber="150" pageId="1" pageNumber="148" type="description"> <subSubSection id="B89D65300D4BFF84FFF67533841DA1B5" lastPageId="3" lastPageNumber="150" pageId="1" pageNumber="148" type="description">
<paragraph id="F03836BB0D4BFF86FFF6753381ADA38D" blockId="1.[102,562,1102,1163]" box="[102,197,1141,1163]" pageId="1" pageNumber="148"> <paragraph id="F03836BB0D4BFF86FFF6753381ADA38D" blockId="1.[102,562,1102,1163]" box="[102,197,1141,1163]" pageId="1" pageNumber="148">
<figureCitation id="68BC2A3E0D4BFF86FFF6753381C1A38D" box="[102,169,1141,1163]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="1" pageNumber="148">Figs. 2</figureCitation> <figureCitation id="68BC2A3E0D4BFF86FFF6753381C1A38D" box="[102,169,1141,1163]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="1" pageNumber="148">Figs. 2</figureCitation>
, ,
<figureCitation id="68BC2A3E0D4BFF86FF23753381A9A38D" box="[179,193,1141,1163]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="1" pageNumber="148">3</figureCitation> <figureCitation id="68BC2A3E0D4BFF86FF23753381A9A38D" box="[179,193,1141,1163]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="1" pageNumber="148">3</figureCitation>
. .
</paragraph> </paragraph>
<paragraph id="F03836BB0D4BFF86FCBC71A1847CA7FB" blockId="1.[812,1486,231,1164]" box="[812,1300,231,253]" pageId="1" pageNumber="148"> <paragraph id="F03836BB0D4BFF86FCBC71A1847CA7FB" blockId="1.[812,1486,231,1164]" box="[812,1300,231,253]" pageId="1" pageNumber="148">
@ -60,7 +62,7 @@
: :
<specimenCode id="A0219EC00D4BFF86FC0171A1828EA7FB" box="[913,998,231,253]" collectionCode="LIGG" pageId="1" pageNumber="148">LIGG 25</specimenCode> <specimenCode id="A0219EC00D4BFF86FC0171A1828EA7FB" box="[913,998,231,253]" collectionCode="LIGG" pageId="1" pageNumber="148">LIGG 25</specimenCode>
A2413, trunk scale ( A2413, trunk scale (
<figureCitation id="68BC2A3E0D4BFF86FB2B71A18461A7FB" box="[1211,1289,231,253]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="1" pageNumber="148">Fig. 2A</figureCitation> <figureCitation id="68BC2A3E0D4BFF86FB2B71A18461A7FB" box="[1211,1289,231,253]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="1" pageNumber="148">Fig. 2A</figureCitation>
). ).
</materialsCitation> </materialsCitation>
</paragraph> </paragraph>
@ -105,24 +107,24 @@ andwidth of
and and
<quantity id="377F9B5E0D4BFF86FA1F723E82EEA4A9" metricMagnitude="-3" metricUnit="m" metricValue="1.1749999999999998" metricValueMax="1.4" metricValueMin="0.95" pageId="1" pageNumber="148" unit="mm" value="1.1749999999999998" valueMax="1.4" valueMin="0.95">0.95 1.4 mm</quantity> <quantity id="377F9B5E0D4BFF86FA1F723E82EEA4A9" metricMagnitude="-3" metricUnit="m" metricValue="1.1749999999999998" metricValueMax="1.4" metricValueMin="0.95" pageId="1" pageNumber="148" unit="mm" value="1.1749999999999998" valueMax="1.4" valueMin="0.95">0.95 1.4 mm</quantity>
, respectively. The crown surface is broad and flat, with slightly downsloped lateral and anterior margins. Four to six sharp subparallel straight or wavy ridges extend the whole crown length. The ridges become more robust towards the scale base.They are usually of the same height across the crown but may be more pronounced and higher posteriorly ( , respectively. The crown surface is broad and flat, with slightly downsloped lateral and anterior margins. Four to six sharp subparallel straight or wavy ridges extend the whole crown length. The ridges become more robust towards the scale base.They are usually of the same height across the crown but may be more pronounced and higher posteriorly (
<figureCitation id="68BC2A3E0D4BFF86FCA4751282A3A36A" box="[820,971,1108,1133]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="1" pageNumber="148">Fig. 2C, D, G</figureCitation> <figureCitation id="68BC2A3E0D4BFF86FCA4751282A3A36A" box="[820,971,1108,1133]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="1" pageNumber="148">Fig. 2C, D, G</figureCitation>
). The two most conspicuous ridges converge at the posterior tip of the crown and may form a slightly raised triangular medial plateau ( ). The two most conspicuous ridges converge at the posterior tip of the crown and may form a slightly raised triangular medial plateau (
<figureCitation id="68BC2A3E0D48FF85FE197649836BA02E" box="[393,515,1807,1832]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2B, D</figureCitation> <figureCitation id="68BC2A3E0D48FF85FE197649836BA02E" box="[393,515,1807,1832]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="2" pageNumber="149">Fig. 2B, D</figureCitation>
), which has a sculpture of weak anterior ridgelets ( ), which has a sculpture of weak anterior ridgelets (
<figureCitation id="68BC2A3E0D48FF85FE017676808EA04E" box="[401,486,1840,1865]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2B</figureCitation> <figureCitation id="68BC2A3E0D48FF85FE017676808EA04E" box="[401,486,1840,1865]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="2" pageNumber="149">Fig. 2B</figureCitation>
) or has an anteriorly widened shallow groove ( ) or has an anteriorly widened shallow groove (
<figureCitation id="68BC2A3E0D48FF85FEC6761680C6A06F" box="[342,430,1872,1897]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2D</figureCitation> <figureCitation id="68BC2A3E0D48FF85FEC6761680C6A06F" box="[342,430,1872,1897]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="2" pageNumber="149">Fig. 2D</figureCitation>
). The ). The
<typeStatus id="2F3C88190D48FF85FE627616833AA06F" box="[498,594,1872,1897]" pageId="2" pageNumber="149" type="holotype">holotype</typeStatus> <typeStatus id="2F3C88190D48FF85FE627616833AA06F" box="[498,594,1872,1897]" pageId="2" pageNumber="149" type="holotype">holotype</typeStatus>
and other specimens have ridges that fade out at the midlength of the crown andafteracentralsmooth area,continueandincreaseinheight posteriorly ( and other specimens have ridges that fade out at the midlength of the crown andafteracentralsmooth area,continueandincreaseinheight posteriorly (
<figureCitation id="68BC2A3E0D48FF85FF7E76F78001A0CC" box="[238,361,1969,1994]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2A, E</figureCitation> <figureCitation id="68BC2A3E0D48FF85FF7E76F78001A0CC" box="[238,361,1969,1994]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="2" pageNumber="149">Fig. 2A, E</figureCitation>
). Of particular interest are the specimens with crown ridges and grooves on the anterior and posterior areas that do not join, but are shifted laterally relative to the long axis of the scale ( ). Of particular interest are the specimens with crown ridges and grooves on the anterior and posterior areas that do not join, but are shifted laterally relative to the long axis of the scale (
<figureCitation id="68BC2A3E0D48FF85FBF7767685A9A04E" box="[1127,1217,1840,1865]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2E</figureCitation> <figureCitation id="68BC2A3E0D48FF85FBF7767685A9A04E" box="[1127,1217,1840,1865]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="2" pageNumber="149">Fig. 2E</figureCitation>
). The grooves between ridges are of irregular depth and width, and sometimes form deeper troughs at their anterior end ( ). The grooves between ridges are of irregular depth and width, and sometimes form deeper troughs at their anterior end (
<figureCitation id="68BC2A3E0D48FF85FB2C7636843CA08F" box="[1212,1364,1904,1929]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2E, G, H</figureCitation> <figureCitation id="68BC2A3E0D48FF85FB2C7636843CA08F" box="[1212,1364,1904,1929]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="2" pageNumber="149">Fig. 2E, G, H</figureCitation>
). ).
</paragraph> </paragraph>
<caption id="A4F866330D4BFF86FFF676398328A0E0" pageId="1" pageNumber="148" startId="1.[102,135,1919,1938]" targetBox="[110,1473,1208,1887]" targetPageId="1" targetType="figure"> <caption id="A4F866330D4BFF86FFF676398328A0E0" ID-DOI="http://doi.org/10.5281/zenodo.13406158" ID-Zenodo-Dep="13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="1" pageNumber="148" startId="1.[102,135,1919,1938]" targetBox="[110,1473,1208,1887]" targetPageId="1" targetType="figure">
<paragraph id="F03836BB0D4BFF86FFF676398328A0E0" blockId="1.[102,1485,1919,2022]" pageId="1" pageNumber="148"> <paragraph id="F03836BB0D4BFF86FFF676398328A0E0" blockId="1.[102,1485,1919,2022]" pageId="1" pageNumber="148">
Fig. 2. Fig. 2.
<taxonomicName id="37874D380D4BFF86FF3876398008A094" authorityName="Valiukevičius" authorityYear="2004" box="[168,352,1919,1938]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="1" pageNumber="148" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov."> <taxonomicName id="37874D380D4BFF86FF3876398008A094" authorityName="Valiukevičius" authorityYear="2004" box="[168,352,1919,1938]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="1" pageNumber="148" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov.">
@ -148,7 +150,7 @@ SEM micrographs, trunk scales except for F, head? scale, crown views, anterior u
). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation. ). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation.
</paragraph> </paragraph>
</caption> </caption>
<caption id="A4F866330D48FF85FFF6770B8254A1D6" pageId="2" pageNumber="149" startId="2.[102,135,1613,1632]" targetBox="[102,1485,227,1583]" targetPageId="2" targetType="figure"> <caption id="A4F866330D48FF85FFF6770B8254A1D6" ID-DOI="http://doi.org/10.5281/zenodo.13406163" ID-Zenodo-Dep="13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="2" pageNumber="149" startId="2.[102,135,1613,1632]" targetBox="[102,1485,227,1583]" targetPageId="2" targetType="figure">
<paragraph id="F03836BB0D48FF85FFF6770B8254A1D6" blockId="2.[102,1485,1613,1744]" pageId="2" pageNumber="149"> <paragraph id="F03836BB0D48FF85FFF6770B8254A1D6" blockId="2.[102,1485,1613,1744]" pageId="2" pageNumber="149">
Fig. 3. Fig. 3.
<taxonomicName id="37874D380D48FF85FF36770B8035A166" authorityName="Valiukevičius" authorityYear="2004" box="[166,349,1613,1632]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="2" pageNumber="149" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov."> <taxonomicName id="37874D380D48FF85FF36770B8035A166" authorityName="Valiukevičius" authorityYear="2004" box="[166,349,1613,1632]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="2" pageNumber="149" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov.">
@ -174,14 +176,14 @@ Microstructure of scales in vertical longitudinal (
</caption> </caption>
<paragraph id="F03836BB0D48FF84FCC276D483B9A626" blockId="2.[812,1486,1807,2026]" lastBlockId="3.[102,775,231,2026]" lastPageId="3" lastPageNumber="150" pageId="2" pageNumber="149"> <paragraph id="F03836BB0D48FF84FCC276D483B9A626" blockId="2.[812,1486,1807,2026]" lastBlockId="3.[102,775,231,2026]" lastPageId="3" lastPageNumber="150" pageId="2" pageNumber="149">
Scale necks are tall and contain large linearly opening basal pores on all the walls, with up to four on each side ( Scale necks are tall and contain large linearly opening basal pores on all the walls, with up to four on each side (
<figureCitation id="68BC2A3E0D48FF85FADE76F484A9A0CC" box="[1358,1473,1970,1995]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2A, E</figureCitation> <figureCitation id="68BC2A3E0D48FF85FADE76F484A9A0CC" box="[1358,1473,1970,1995]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="2" pageNumber="149">Fig. 2A, E</figureCitation>
). The scale base is rhombic in outline, moderately to greatly convex, usually protruding slightly beyond the crown anterolaterally. The deepest point of the base is centrally positioned. ). The scale base is rhombic in outline, moderately to greatly convex, usually protruding slightly beyond the crown anterolaterally. The deepest point of the base is centrally positioned.
</paragraph> </paragraph>
<paragraph id="F03836BB0D49FF84FF1B70608088A55C" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150"> <paragraph id="F03836BB0D49FF84FF1B70608088A55C" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
Putative headscales(veryrareinthestudiedsamples)have isometric rhomboid, thickened crowns (up to Putative headscales(veryrareinthestudiedsamples)have isometric rhomboid, thickened crowns (up to
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long and wide) and low porous necks as compared to trunk scales. Head scale crowns have three to four prominent, robust and rounded ridges of even height extending over the entire crown length. These robust ridges bend medially towards the posterior edge ( long and wide) and low porous necks as compared to trunk scales. Head scale crowns have three to four prominent, robust and rounded ridges of even height extending over the entire crown length. These robust ridges bend medially towards the posterior edge (
<figureCitation id="68BC2A3E0D49FF84FE9E70A58008A6FA" box="[270,352,483,508]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 2F</figureCitation> <figureCitation id="68BC2A3E0D49FF84FE9E70A58008A6FA" box="[270,352,483,508]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="3" pageNumber="150">Fig. 2F</figureCitation>
), but do not join each other. Additional lateral oblique riblets occur on the side of each ridge. The posterolateral crown edge is multicuspid with several (up to four on each side) rough spinelets. ), but do not join each other. Additional lateral oblique riblets occur on the side of each ridge. The posterolateral crown edge is multicuspid with several (up to four on each side) rough spinelets.
</paragraph> </paragraph>
<paragraph id="F03836BB0D49FF84FF1B73278363A492" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150"> <paragraph id="F03836BB0D49FF84FF1B73278363A492" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
@ -190,7 +192,7 @@ The histology of all
<emphasis id="C2F3EAA90D49FF84FEF97324808FA57C" box="[361,487,609,634]" italics="true" pageId="3" pageNumber="150">V. perplexa</emphasis> <emphasis id="C2F3EAA90D49FF84FEF97324808FA57C" box="[361,487,609,634]" italics="true" pageId="3" pageNumber="150">V. perplexa</emphasis>
</taxonomicName> </taxonomicName>
scales ( scales (
<figureCitation id="68BC2A3E0D49FF84FDAE73278317A57C" box="[574,639,609,634]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3</figureCitation> <figureCitation id="68BC2A3E0D49FF84FDAE73278317A57C" box="[574,639,609,634]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="3" pageNumber="150">Fig. 3</figureCitation>
) is uniform. Crowns are composed of stranggewebe (mesodentinal tissue containing elongate, linearly oriented lacunae and incorporating osteocytes; for histologic terminology see ) is uniform. Crowns are composed of stranggewebe (mesodentinal tissue containing elongate, linearly oriented lacunae and incorporating osteocytes; for histologic terminology see
<bibRefCitation id="94164B4A0D49FF84FDCC738683B1A5DE" author="Gross, W." box="[604,729,704,728]" pageId="3" pageNumber="150" pagination="1 - 82" refId="ref9073" refString="Gross, W. 1971. Downtonische und Dittonische Acanthodier-Reste des Ostseegebietes. Palaeontographica A 136: 1 - 82." type="journal article" year="1971">Gross 1971</bibRefCitation> <bibRefCitation id="94164B4A0D49FF84FDCC738683B1A5DE" author="Gross, W." box="[604,729,704,728]" pageId="3" pageNumber="150" pagination="1 - 82" refId="ref9073" refString="Gross, W. 1971. Downtonische und Dittonische Acanthodier-Reste des Ostseegebietes. Palaeontographica A 136: 1 - 82." type="journal article" year="1971">Gross 1971</bibRefCitation>
and and
@ -198,8 +200,8 @@ and
) and simple mesodentine arranged in four thick growth lamellae. The growth ) and simple mesodentine arranged in four thick growth lamellae. The growth
<typeStatus id="2F3C88190D49FF84FE5873B9809FA411" box="[456,503,767,791]" pageId="3" pageNumber="150">type</typeStatus> <typeStatus id="2F3C88190D49FF84FE5873B9809FA411" box="[456,503,767,791]" pageId="3" pageNumber="150">type</typeStatus>
differs slightly among scales. Scales with interrupted ridges ( differs slightly among scales. Scales with interrupted ridges (
<figureCitation id="68BC2A3E0D49FF84FD85725B8303A430" box="[533,619,797,822]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3A</figureCitation> <figureCitation id="68BC2A3E0D49FF84FD85725B8303A430" box="[533,619,797,822]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="3" pageNumber="150">Fig. 3A</figureCitation>
<figureCitation id="68BC2A3E0D49FF84FDFC726C83C8A43D" box="[620,672,798,827]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." pageId="3" pageNumber="150"> <figureCitation id="68BC2A3E0D49FF84FDFC726C83C8A43D" box="[620,672,798,827]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." figureDoi="http://doi.org/10.5281/zenodo.13406151" httpUri="https://zenodo.org/record/13406151/files/figure.png" pageId="3" pageNumber="150">
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, B , B
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@ -208,33 +210,33 @@ differs slightly among scales. Scales with interrupted ridges (
</paragraph> </paragraph>
<paragraph id="F03836BB0D49FF84FF1B72DD83BDA376" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150"> <paragraph id="F03836BB0D49FF84FF1B72DD83BDA376" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
Thestranggewebeischaracterizedbyenlargedandelongate stranglacunae with interconnecting short processes. An unevenly thick mantle of simple mesodentine is superficial to the stranggewebe in each growth lamella ( Thestranggewebeischaracterizedbyenlargedandelongate stranglacunae with interconnecting short processes. An unevenly thick mantle of simple mesodentine is superficial to the stranggewebe in each growth lamella (
<figureCitation id="68BC2A3E0D49FF84FD8172BF830EA314" box="[529,614,1017,1042]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3B</figureCitation> <figureCitation id="68BC2A3E0D49FF84FD8172BF830EA314" box="[529,614,1017,1042]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="3" pageNumber="150">Fig. 3B</figureCitation>
<figureCitation id="68BC2A3E0D49FF84FDF575408306A311" box="[613,622,1030,1047]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150"> <figureCitation id="68BC2A3E0D49FF84FDF575408306A311" box="[613,622,1030,1047]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="3" pageNumber="150">
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). In scales of this species, the stranggewebe develops with a similar intensity not only in the posterior part of the crown, as it is characteristic of most climatiids, but also in its anterior parts ( ). In scales of this species, the stranggewebe develops with a similar intensity not only in the posterior part of the crown, as it is characteristic of most climatiids, but also in its anterior parts (
<figureCitation id="68BC2A3E0D49FF84FDF9751E83D4A376" box="[617,700,1112,1137]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3A</figureCitation> <figureCitation id="68BC2A3E0D49FF84FDF9751E83D4A376" box="[617,700,1112,1137]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="3" pageNumber="150">Fig. 3A</figureCitation>
<figureCitation id="68BC2A3E0D49FF84FD2C752283ADA373" box="[700,709,1124,1141]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." pageId="3" pageNumber="150"> <figureCitation id="68BC2A3E0D49FF84FD2C752283ADA373" box="[700,709,1124,1141]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." figureDoi="http://doi.org/10.5281/zenodo.13406151" httpUri="https://zenodo.org/record/13406151/files/figure.png" pageId="3" pageNumber="150">
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). ).
</paragraph> </paragraph>
<paragraph id="F03836BB0D49FF84FF1B75318268A22B" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150"> <paragraph id="F03836BB0D49FF84FF1B75318268A22B" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
The basal part of the neck has rare, typical mesodentine osteocyte cavities. The principal vascular system is formed of enlarged radial vascular canals positioned over the base, and wide ascending and slightly branched main canals supplying each growth lamella ( The basal part of the neck has rare, typical mesodentine osteocyte cavities. The principal vascular system is formed of enlarged radial vascular canals positioned over the base, and wide ascending and slightly branched main canals supplying each growth lamella (
<figureCitation id="68BC2A3E0D49FF84FEDC75B380F6A208" box="[332,414,1269,1294]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3A</figureCitation> <figureCitation id="68BC2A3E0D49FF84FEDC75B380F6A208" box="[332,414,1269,1294]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="3" pageNumber="150">Fig. 3A</figureCitation>
<figureCitation id="68BC2A3E0D49FF84FE0E744480CFA215" box="[414,423,1282,1299]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150"> <figureCitation id="68BC2A3E0D49FF84FE0E744480CFA215" box="[414,423,1282,1299]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="3" pageNumber="150">
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, B , B
<figureCitation id="68BC2A3E0D49FF84FE56744480A7A215" box="[454,463,1282,1299]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150"> <figureCitation id="68BC2A3E0D49FF84FE56744480A7A215" box="[454,463,1282,1299]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="3" pageNumber="150">
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). This system is complicated intheearliest/oldestlamellaebecauseofadditionalbranchings. ). This system is complicated intheearliest/oldestlamellaebecauseofadditionalbranchings.
</paragraph> </paragraph>
<paragraph id="F03836BB0D49FF84FF1B747283C6A2AD" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150"> <paragraph id="F03836BB0D49FF84FF1B747283C6A2AD" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
The simple mesodentine enclosing stranggewebe in the outer parts of lamellae contains a characteristic net of winding dentine tubules with many osteocyte cavities even in the superficial layers ( The simple mesodentine enclosing stranggewebe in the outer parts of lamellae contains a characteristic net of winding dentine tubules with many osteocyte cavities even in the superficial layers (
<figureCitation id="68BC2A3E0D49FF84FE8874D48006A2AD" box="[280,366,1426,1451]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3A</figureCitation> <figureCitation id="68BC2A3E0D49FF84FE8874D48006A2AD" box="[280,366,1426,1451]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="3" pageNumber="150">Fig. 3A</figureCitation>
<figureCitation id="68BC2A3E0D49FF84FEFD74D980C8A2B6" box="[365,416,1428,1456]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150"> <figureCitation id="68BC2A3E0D49FF84FEFD74D980C8A2B6" box="[365,416,1428,1456]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="3" pageNumber="150">
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, B , B
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@ -243,8 +245,8 @@ The simple mesodentine enclosing stranggewebe in the outer parts of lamellae con
</paragraph> </paragraph>
<paragraph id="F03836BB0D49FF84FF1B74F48393A16E" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150"> <paragraph id="F03836BB0D49FF84FF1B74F48393A16E" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
Scale bases are composed of cellular bone arranged in very thin and dense growth lamellae with plenty of osteocyte cavities and medium to long Sharpeys fibre traces. The osteocyte density increases towards the tip of pyramidal base, where they become oriented along the growth lines, and approximate thestructureofbothbonetissueandstranggewebe( Scale bases are composed of cellular bone arranged in very thin and dense growth lamellae with plenty of osteocyte cavities and medium to long Sharpeys fibre traces. The osteocyte density increases towards the tip of pyramidal base, where they become oriented along the growth lines, and approximate thestructureofbothbonetissueandstranggewebe(
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<figureCitation id="68BC2A3E0D49FF84FD71771A8382A16B" box="[737,746,1628,1645]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150"> <figureCitation id="68BC2A3E0D49FF84FD71771A8382A16B" box="[737,746,1628,1645]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="3" pageNumber="150">
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<figureCitation id="68BC2A3E0D4EFF83FFF6705481C1A62E" box="[102,169,274,296]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Figs. 4</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FFF6705481C1A62E" box="[102,169,274,296]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." figureDoi="http://doi.org/10.5281/zenodo.13406169" httpUri="https://zenodo.org/record/13406169/files/figure.png" pageId="4" pageNumber="151">Figs. 4</figureCitation>
, ,
<figureCitation id="68BC2A3E0D4EFF83FF23705481A9A62E" box="[179,193,274,296]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">5</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FF23705481A9A62E" box="[179,193,274,296]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." figureDoi="http://doi.org/10.5281/zenodo.13406173" httpUri="https://zenodo.org/record/13406173/files/figure.png" pageId="4" pageNumber="151">5</figureCitation>
. .
</paragraph> </paragraph>
<paragraph id="F03836BB0D4EFF83FFF6707D8345A657" blockId="4.[102,775,315,603]" box="[102,557,315,337]" pageId="4" pageNumber="151"> <paragraph id="F03836BB0D4EFF83FFF6707D8345A657" blockId="4.[102,775,315,603]" box="[102,557,315,337]" pageId="4" pageNumber="151">
@ -69,7 +71,7 @@ sp. A
: :
<specimenCode id="A0219EC00D4EFF83FF5A70198076A673" box="[202,286,351,373]" collectionCode="LIGG" pageId="4" pageNumber="151">LIGG 25</specimenCode> <specimenCode id="A0219EC00D4EFF83FF5A70198076A673" box="[202,286,351,373]" collectionCode="LIGG" pageId="4" pageNumber="151">LIGG 25</specimenCode>
A2405, trunk scale ( A2405, trunk scale (
<figureCitation id="68BC2A3E0D4EFF83FE647019832AA673" box="[500,578,351,373]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4A</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FE647019832AA673" box="[500,578,351,373]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." figureDoi="http://doi.org/10.5281/zenodo.13406169" httpUri="https://zenodo.org/record/13406169/files/figure.png" pageId="4" pageNumber="151">Fig. 4A</figureCitation>
). ).
</materialsCitation> </materialsCitation>
</paragraph> </paragraph>
@ -107,7 +109,7 @@ sp. A
</taxonomicName> </taxonomicName>
scales having low horizontal crowns with crenulated sculpture anteriorly or around the entire crown margin, lacking a neck; deep convex base present. Scale crownscomposed of simple bonelike mesodentine with abundant osteocyte cavities and enlarged vascular canals present in the oldest growth lamellae only. Crown mesodentine merges gradually into highly cellular thinlamellar bone in the base that is penetrated by long traces of Sharpeys fibres. scales having low horizontal crowns with crenulated sculpture anteriorly or around the entire crown margin, lacking a neck; deep convex base present. Scale crownscomposed of simple bonelike mesodentine with abundant osteocyte cavities and enlarged vascular canals present in the oldest growth lamellae only. Crown mesodentine merges gradually into highly cellular thinlamellar bone in the base that is penetrated by long traces of Sharpeys fibres.
</paragraph> </paragraph>
<caption id="A4F866330D4EFF83FCBC73A18506A4A4" pageId="4" pageNumber="151" startId="4.[812,845,743,762]" targetBox="[813,1484,240,691]" targetPageId="4" targetType="figure"> <caption id="A4F866330D4EFF83FCBC73A18506A4A4" ID-DOI="http://doi.org/10.5281/zenodo.13406169" ID-Zenodo-Dep="13406169" httpUri="https://zenodo.org/record/13406169/files/figure.png" pageId="4" pageNumber="151" startId="4.[812,845,743,762]" targetBox="[813,1484,240,691]" targetPageId="4" targetType="figure">
<paragraph id="F03836BB0D4EFF83FCBC73A18506A4A4" blockId="4.[812,1485,743,930]" pageId="4" pageNumber="151"> <paragraph id="F03836BB0D4EFF83FCBC73A18506A4A4" blockId="4.[812,1485,743,930]" pageId="4" pageNumber="151">
Fig. 4. Fig. 4.
<taxonomicName id="37874D380D4EFF83FCFE73A18510A5FC" authorityName="Valiukevičius" authorityYear="2004" box="[878,1144,743,762]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov."> <taxonomicName id="37874D380D4EFF83FCFE73A18510A5FC" authorityName="Valiukevičius" authorityYear="2004" box="[878,1144,743,762]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov.">
@ -144,55 +146,55 @@ SEM micrographs of scales.
, and can be , and can be
<quantity id="377F9B5E0D4EFF83FDF172A5826FA4FD" box="[609,775,995,1019]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="7.7" metricValueMin="3.0" pageId="4" pageNumber="151" unit="mm" value="0.535" valueMax="0.77" valueMin="0.3">0.3 to 0.77 mm</quantity> <quantity id="377F9B5E0D4EFF83FDF172A5826FA4FD" box="[609,775,995,1019]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="7.7" metricValueMin="3.0" pageId="4" pageNumber="151" unit="mm" value="0.535" valueMax="0.77" valueMin="0.3">0.3 to 0.77 mm</quantity>
wide. The isometric rhomboid crown plate of trunk scales ( wide. The isometric rhomboid crown plate of trunk scales (
<figureCitation id="68BC2A3E0D4EFF83FFFE75678068A33C" box="[110,256,1057,1082]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4A, B, E</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FFFE75678068A33C" box="[110,256,1057,1082]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." figureDoi="http://doi.org/10.5281/zenodo.13406169" httpUri="https://zenodo.org/record/13406169/files/figure.png" pageId="4" pageNumber="151">Fig. 4A, B, E</figureCitation>
) is almost flat, very slightly curved downwards at the anterior margins, and posteriorly the crowns are elongated and rhomboidal to elipsoidal. The anterior margin of the crown has short but comparatively wide notches which extend to the surface of the scale base. The number of notches varies from 5 to 6 ( ) is almost flat, very slightly curved downwards at the anterior margins, and posteriorly the crowns are elongated and rhomboidal to elipsoidal. The anterior margin of the crown has short but comparatively wide notches which extend to the surface of the scale base. The number of notches varies from 5 to 6 (
<figureCitation id="68BC2A3E0D4EFF83FF7D75F88057A3D1" box="[237,319,1214,1239]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4E</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FF7D75F88057A3D1" box="[237,319,1214,1239]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." figureDoi="http://doi.org/10.5281/zenodo.13406169" httpUri="https://zenodo.org/record/13406169/files/figure.png" pageId="4" pageNumber="151">Fig. 4E</figureCitation>
) to 10 ( ) to 10 (
<figureCitation id="68BC2A3E0D4EFF83FE0375F88081A3D1" box="[403,489,1214,1239]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4B</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FE0375F88081A3D1" box="[403,489,1214,1239]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." figureDoi="http://doi.org/10.5281/zenodo.13406169" httpUri="https://zenodo.org/record/13406169/files/figure.png" pageId="4" pageNumber="151">Fig. 4B</figureCitation>
). The putative head scales ( ). The putative head scales (
<figureCitation id="68BC2A3E0D4EFF83FFFE7598818FA3F0" box="[110,231,1246,1271]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4C, D</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FFFE7598818FA3F0" box="[110,231,1246,1271]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." figureDoi="http://doi.org/10.5281/zenodo.13406169" httpUri="https://zenodo.org/record/13406169/files/figure.png" pageId="4" pageNumber="151">Fig. 4C, D</figureCitation>
) differ from the trunk scales in having isometric rhomboid to round crowns with notches on all margins. The deep,convexscale baseis rhomboid to round in outline, with a prominent lateral rim at the deepest point projecting anterior to the crown. In rare cases the deepest point of the base is situated posteriorly, so the base protrudes beyond the posterior margin of the crown ( ) differ from the trunk scales in having isometric rhomboid to round crowns with notches on all margins. The deep,convexscale baseis rhomboid to round in outline, with a prominent lateral rim at the deepest point projecting anterior to the crown. In rare cases the deepest point of the base is situated posteriorly, so the base protrudes beyond the posterior margin of the crown (
<figureCitation id="68BC2A3E0D4EFF83FEC474DC80C1A2B5" box="[340,425,1434,1459]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4E</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FEC474DC80C1A2B5" box="[340,425,1434,1459]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25A2405, anterolateral view, anterior to the left. B. LIGG 25A2407, crown view, anterior upwards. E. LIGG 25A2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25A2409. D. LIGG 25A2408. Scale bars 0.1 mm. Gėluva99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." figureDoi="http://doi.org/10.5281/zenodo.13406169" httpUri="https://zenodo.org/record/13406169/files/figure.png" pageId="4" pageNumber="151">Fig. 4E</figureCitation>
). Scales lack necks. ). Scales lack necks.
</paragraph> </paragraph>
<paragraph id="F03836BB0D4EFF83FF1B74FD83D1A0EC" blockId="4.[102,775,932,2026]" pageId="4" pageNumber="151"> <paragraph id="F03836BB0D4EFF83FF1B74FD83D1A0EC" blockId="4.[102,775,932,2026]" pageId="4" pageNumber="151">
Simple mesodentine crown tissue (neither stranggewebe nor durodentine present) and cellular bone in scale bases can be seen in the sectioned scales. Up to six thin lamellae of superpositional growth in crowns are composed of bonelike mesodentine containing numerous multiangular osteocyte spaces even in the latest, outer lamella ( Simple mesodentine crown tissue (neither stranggewebe nor durodentine present) and cellular bone in scale bases can be seen in the sectioned scales. Up to six thin lamellae of superpositional growth in crowns are composed of bonelike mesodentine containing numerous multiangular osteocyte spaces even in the latest, outer lamella (
<figureCitation id="68BC2A3E0D4EFF83FD82771E830EA177" box="[530,614,1624,1649]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5A</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FD82771E830EA177" box="[530,614,1624,1649]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." figureDoi="http://doi.org/10.5281/zenodo.13406173" httpUri="https://zenodo.org/record/13406173/files/figure.png" pageId="4" pageNumber="151">Fig. 5A</figureCitation>
<figureCitation id="68BC2A3E0D4EFF83FDF677238307A170" box="[614,623,1637,1654]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151"> <figureCitation id="68BC2A3E0D4EFF83FDF677238307A170" box="[614,623,1637,1654]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="4" pageNumber="151">
<subScript id="6C0334FE0D4EFF83FDF677238307A170" attach="left" box="[614,623,1637,1654]" fontSize="7" pageId="4" pageNumber="151">2</subScript> <subScript id="6C0334FE0D4EFF83FDF677238307A170" attach="left" box="[614,623,1637,1654]" fontSize="7" pageId="4" pageNumber="151">2</subScript>
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) and short radiating and winding dentine tubules form a fine network. The scale primordium does not differ from others in respect to tissue structure. Most scales sectioned have no enlarged vascular canal system in crowns ( ) and short radiating and winding dentine tubules form a fine network. The scale primordium does not differ from others in respect to tissue structure. Most scales sectioned have no enlarged vascular canal system in crowns (
<figureCitation id="68BC2A3E0D4EFF83FEFF779080ABA1E9" box="[367,451,1750,1775]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5A</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FEFF779080ABA1E9" box="[367,451,1750,1775]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." figureDoi="http://doi.org/10.5281/zenodo.13406173" httpUri="https://zenodo.org/record/13406173/files/figure.png" pageId="4" pageNumber="151">Fig. 5A</figureCitation>
<figureCitation id="68BC2A3E0D4EFF83FE5377A5809CA1F2" box="[451,500,1751,1780]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." pageId="4" pageNumber="151"> <figureCitation id="68BC2A3E0D4EFF83FE5377A5809CA1F2" box="[451,500,1751,1780]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." figureDoi="http://doi.org/10.5281/zenodo.13406151" httpUri="https://zenodo.org/record/13406151/files/figure.png" pageId="4" pageNumber="151">
<subScript id="6C0334FE0D4EFF83FE5377A580A4A1F2" attach="left" box="[451,460,1763,1780]" fontSize="7" pageId="4" pageNumber="151">1</subScript> <subScript id="6C0334FE0D4EFF83FE5377A580A4A1F2" attach="left" box="[451,460,1763,1780]" fontSize="7" pageId="4" pageNumber="151">1</subScript>
, B , B
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), and only one ( ), and only one (
<figureCitation id="68BC2A3E0D4EFF83FD347790839EA1E9" box="[676,758,1750,1775]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5C</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FD347790839EA1E9" box="[676,758,1750,1775]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." figureDoi="http://doi.org/10.5281/zenodo.13406173" httpUri="https://zenodo.org/record/13406173/files/figure.png" pageId="4" pageNumber="151">Fig. 5C</figureCitation>
<figureCitation id="68BC2A3E0D4EFF83FD6677A58397A1F2" box="[758,767,1763,1780]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." pageId="4" pageNumber="151"> <figureCitation id="68BC2A3E0D4EFF83FD6677A58397A1F2" box="[758,767,1763,1780]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." figureDoi="http://doi.org/10.5281/zenodo.13406151" httpUri="https://zenodo.org/record/13406151/files/figure.png" pageId="4" pageNumber="151">
<subScript id="6C0334FE0D4EFF83FD6677A58397A1F2" attach="left" box="[758,767,1763,1780]" fontSize="7" pageId="4" pageNumber="151">1</subScript> <subScript id="6C0334FE0D4EFF83FD6677A58397A1F2" attach="left" box="[758,767,1763,1780]" fontSize="7" pageId="4" pageNumber="151">1</subScript>
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, C , C
<figureCitation id="68BC2A3E0D4EFF83FFE8764481E9A015" box="[120,129,1794,1811]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151"> <figureCitation id="68BC2A3E0D4EFF83FFE8764481E9A015" box="[120,129,1794,1811]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="4" pageNumber="151">
<subScript id="6C0334FE0D4EFF83FFE8764481E9A015" attach="left" box="[120,129,1794,1811]" fontSize="7" pageId="4" pageNumber="151">2</subScript> <subScript id="6C0334FE0D4EFF83FFE8764481E9A015" attach="left" box="[120,129,1794,1811]" fontSize="7" pageId="4" pageNumber="151">2</subScript>
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) demonstrates wide radial vascular canals in the two oldest growth lamellae. There is no sharp boundary between the crown mesodentine and bone of the base: the first tissue grades gradually into the second. The cellular bone of the base is composed of numerous very fine growth zones ( ) demonstrates wide radial vascular canals in the two oldest growth lamellae. There is no sharp boundary between the crown mesodentine and bone of the base: the first tissue grades gradually into the second. The cellular bone of the base is composed of numerous very fine growth zones (
<figureCitation id="68BC2A3E0D4EFF83FD307635839EA08A" box="[672,758,1907,1932]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5A</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FD307635839EA08A" box="[672,758,1907,1932]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." figureDoi="http://doi.org/10.5281/zenodo.13406173" httpUri="https://zenodo.org/record/13406173/files/figure.png" pageId="4" pageNumber="151">Fig. 5A</figureCitation>
<figureCitation id="68BC2A3E0D4EFF83FD6676C68397A097" box="[758,767,1920,1937]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." pageId="4" pageNumber="151"> <figureCitation id="68BC2A3E0D4EFF83FD6676C68397A097" box="[758,767,1920,1937]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1Stoniškiai, 7Krekenava, 8Stačiūnai, 11Šešuvis, 12Kunkojai, 44Nida, 61Viduklė, 87Sutkai, 99Gėluva, 128Vilkaviškis, 137Liepkalnis, 162Kurtuvėnai, 179Ledai, 241Butkūnai, 252Svėdasai." figureDoi="http://doi.org/10.5281/zenodo.13406151" httpUri="https://zenodo.org/record/13406151/files/figure.png" pageId="4" pageNumber="151">
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, A , A
<figureCitation id="68BC2A3E0D4EFF83FFE976D981EAA0B6" box="[121,130,1951,1968]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151"> <figureCitation id="68BC2A3E0D4EFF83FFE976D981EAA0B6" box="[121,130,1951,1968]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="4" pageNumber="151">
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) penetrated by long Sharpeys fibre traces ( ) penetrated by long Sharpeys fibre traces (
<figureCitation id="68BC2A3E0D4EFF83FDE376D483A1A0AD" box="[627,713,1938,1963]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5B</figureCitation> <figureCitation id="68BC2A3E0D4EFF83FDE376D483A1A0AD" box="[627,713,1938,1963]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1B3) and Ledai179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." figureDoi="http://doi.org/10.5281/zenodo.13406173" httpUri="https://zenodo.org/record/13406173/files/figure.png" pageId="4" pageNumber="151">Fig. 5B</figureCitation>
<figureCitation id="68BC2A3E0D4EFF83FD5976D983BAA0B6" box="[713,722,1951,1968]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151"> <figureCitation id="68BC2A3E0D4EFF83FD5976D983BAA0B6" box="[713,722,1951,1968]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25A2413 (A), LIGG 25A2416 (B), LIGG 25A2417 (C), LIGG 25A2418 (D), LIGG 25A2420 (E), LIGG 25A2422 (F), LIGG 25A2421 (G), LIGG 25A2414(H). Scalebars0.1 mm.Nida44borehole, depth1213.0 m(A, EH) andStoniškiai1borehole, depth1211.01217.0 m(BD). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406158" httpUri="https://zenodo.org/record/13406158/files/figure.png" pageId="4" pageNumber="151">
<subScript id="6C0334FE0D4EFF83FD5976D983BAA0B6" attach="left" box="[713,722,1951,1968]" fontSize="7" pageId="4" pageNumber="151">2</subScript> <subScript id="6C0334FE0D4EFF83FD5976D983BAA0B6" attach="left" box="[713,722,1951,1968]" fontSize="7" pageId="4" pageNumber="151">2</subScript>
</figureCitation> </figureCitation>
, B , B
<figureCitation id="68BC2A3E0D4EFF83FD6476D98395A0B6" box="[756,765,1951,1968]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151"> <figureCitation id="68BC2A3E0D4EFF83FD6476D98395A0B6" box="[756,765,1951,1968]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotypelike scale with uniformbent, subparallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai1borehole, depth 1211.01217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." figureDoi="http://doi.org/10.5281/zenodo.13406163" httpUri="https://zenodo.org/record/13406163/files/figure.png" pageId="4" pageNumber="151">
<subScript id="6C0334FE0D4EFF83FD6476D98395A0B6" attach="left" box="[756,765,1951,1968]" fontSize="7" pageId="4" pageNumber="151">3</subScript> <subScript id="6C0334FE0D4EFF83FD6476D98395A0B6" attach="left" box="[756,765,1951,1968]" fontSize="7" pageId="4" pageNumber="151">3</subScript>
</figureCitation> </figureCitation>
) and enclosed abundant osteocyte cavities with short cell processes. Osteocytes are oriented along growth lines. ) and enclosed abundant osteocyte cavities with short cell processes. Osteocytes are oriented along growth lines.
@ -253,7 +255,7 @@ and the Dubovskoye scales is the presenceofsmallbulbsorsometimes conelikespin
</taxonomicName> </taxonomicName>
, what suggests a climatiid rather than an ischnacanthid affinity for the Dubovskoye scales. , what suggests a climatiid rather than an ischnacanthid affinity for the Dubovskoye scales.
</paragraph> </paragraph>
<caption id="A4F866330D4FFF82FFF6762282F2A0E1" pageId="5" pageNumber="152" startId="5.[102,135,1892,1911]" targetBox="[201,1386,225,1866]" targetPageId="5" targetType="figure"> <caption id="A4F866330D4FFF82FFF6762282F2A0E1" ID-DOI="http://doi.org/10.5281/zenodo.13406173" ID-Zenodo-Dep="13406173" httpUri="https://zenodo.org/record/13406173/files/figure.png" pageId="5" pageNumber="152" startId="5.[102,135,1892,1911]" targetBox="[201,1386,225,1866]" targetPageId="5" targetType="figure">
<paragraph id="F03836BB0D4FFF82FFF6762282F2A0E1" blockId="5.[102,1485,1892,2027]" pageId="5" pageNumber="152"> <paragraph id="F03836BB0D4FFF82FFF6762282F2A0E1" blockId="5.[102,1485,1892,2027]" pageId="5" pageNumber="152">
Fig. 5. Fig. 5.
<taxonomicName id="37874D380D4FFF82FF34762280C4A071" authorityName="Valiukevičius" authorityYear="2004" box="[164,428,1892,1911]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="5" pageNumber="152" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov."> <taxonomicName id="37874D380D4FFF82FF34762280C4A071" authorityName="Valiukevičius" authorityYear="2004" box="[164,428,1892,1911]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="5" pageNumber="152" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov.">