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<document id="827D7B2DF6BA88DC76210F618065B61B" ID-ISSN="1732-2421" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1723062957633" checkinUser="felipe" docAuthor="Valiukevičius, Juozas" docDate="2004" docId="782E87AD0D42FF8CFCBC74F68385A3FA" docLanguage="en" docName="ActaPalaeontolPol.49.1.147-160.pdf" docOrigin="Acta Palaeontologica Polonica 49 (1)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Rohonilepis breviornatus Valiukevičius, 2004, gen. et sp. nov." docType="treatment" docVersion="2" lastPageNumber="158" masterDocId="8417FFD50D4AFF87FF9071468168A706" masterDocTitle="New Wenlock-Pridoli (Silurian) acanthodian fishes from Lithuania" masterLastPageNumber="160" masterPageNumber="147" pageNumber="155" updateTime="1724888488487" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="77920DD895C0663E288B099F79681983" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B96CAA3CCA4B8B4FE066D008B0F83CC1">
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<mods:title id="FC07EDF2E6D9E457098F5910F95A845C">New Wenlock-Pridoli (Silurian) acanthodian fishes from Lithuania</mods:title>
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</mods:titleInfo>
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<mods:name id="7D0F87D42319D05BA7319E6AD09F951C" type="personal">
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<mods:roleTerm id="80C37F08E6F1A87304C2632679F271A4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="310C2868D2E9B796E8B4DBC92CBB6FF3">Valiukevičius, Juozas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="09146E53A99220C9D78AD53D45CBDA81">text</mods:typeOfResource>
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<mods:titleInfo id="B60C8B4BCCD91F26C55E603D8AB70FCC">
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<mods:title id="754CBEB3249A06F98BD5D8F63E8D1274">Acta Palaeontologica Polonica</mods:title>
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</mods:titleInfo>
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<mods:part id="E9801AC4B4D241BD822BF20239333F21">
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<mods:date id="D99D05482DF5481999EE18BB88815294">2004</mods:date>
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<mods:detail id="A7BDB6B570488C5A6236EE6F7F9AF8CB" type="pubDate">
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<mods:number id="74E257CF76E651D48B8D84563E75F178">2004-12-31</mods:number>
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</mods:detail>
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<mods:detail id="2F7D1AE0DABE03F344F6D2348ACD959E" type="volume">
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<mods:number id="CAC51DA76A23DD5637CFB3CD57BDF2C2">49</mods:number>
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</mods:detail>
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<mods:detail id="C63CB1487CED99A725814D98F889FE73" type="issue">
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<mods:number id="FE043DDB0047DF02CF4A9575426D3B41">1</mods:number>
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<mods:extent id="75D3575DB09F0618DD87E041D0748640" unit="page">
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<mods:start id="71A2C21B9B52EA65F13DC195339AAAC1">147</mods:start>
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<mods:end id="B87714F3CD13762BE7EB3F0418DBC7DE">160</mods:end>
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</mods:relatedItem>
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<mods:classification id="4EE25FEC0521B0049EC554A355D589FE">journal article</mods:classification>
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<mods:identifier id="4FFEBE0160C92A322FE90CBC5C2CB871" type="ISSN">1732-2421</mods:identifier>
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</mods:mods>
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<treatment id="782E87AD0D42FF8CFCBC74F68385A3FA" LSID="urn:lsid:plazi:treatment:782E87AD0D42FF8CFCBC74F68385A3FA" httpUri="http://treatment.plazi.org/id/782E87AD0D42FF8CFCBC74F68385A3FA" lastPageId="11" lastPageNumber="158" pageId="8" pageNumber="155">
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<subSubSection id="B89D65300D42FF8FFCBC74F68452A2CB" box="[812,1338,1456,1485]" pageId="8" pageNumber="155" type="nomenclature">
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<paragraph id="F03836BB0D42FF8FFCBC74F68452A2CB" blockId="8.[812,1338,1456,1518]" box="[812,1338,1456,1485]" pageId="8" pageNumber="155">
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<heading id="AB7081D70D42FF8FFCBC74F68452A2CB" box="[812,1338,1456,1485]" fontSize="12" level="2" pageId="8" pageNumber="155" reason="9">
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||||
<taxonomicName id="37874D380D42FF8FFCBC74F68518A2CB" authorityName="Valiukevičius" authorityYear="2004" box="[812,1136,1456,1485]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="8" pageNumber="155" phylum="Chordata" rank="species" species="breviornatus" status="gen. et sp. nov.">
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||||
<emphasis id="C2F3EAA90D42FF8FFCBC74F68518A2CB" box="[812,1136,1456,1485]" italics="true" pageId="8" pageNumber="155">Rohonilepis breviornatus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D42FF8FFBE874F78452A2CB" box="[1144,1338,1457,1485]" pageId="8" pageNumber="155" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="B89D65300D42FF8CFCBC749E8385A3FA" lastPageId="11" lastPageNumber="158" pageId="8" pageNumber="155" type="description">
|
||||
<paragraph id="F03836BB0D42FF8FFCBC749E8202A2E8" blockId="8.[812,1338,1456,1518]" box="[812,874,1496,1518]" pageId="8" pageNumber="155">
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||||
<figureCitation id="68BC2A3E0D42FF8FFCBC749E820EA2E8" box="[812,870,1496,1518]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. A–D. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25−A−2457. B. LIGG 25−A−2458. C. Holotype, LIGG 25−A−2459. D. LIGG 25−A−2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="8" pageNumber="155">Fig. 8</figureCitation>
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||||
.
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||||
</paragraph>
|
||||
<paragraph id="F03836BB0D42FF8FFCBC74B9847AA113" blockId="8.[812,1485,1535,1654]" box="[812,1298,1535,1557]" pageId="8" pageNumber="155">
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||||
<materialsCitation id="40EF3CE60D42FF8FFCBC74B9847AA113" box="[812,1298,1535,1557]" collectionCode="LIGG" pageId="8" pageNumber="155" specimenCode="LIGG 25" specimenCount="1" typeStatus="holotype">
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||||
<emphasis id="C2F3EAA90D42FF8FFCBC74B982ECA113" box="[812,900,1535,1557]" italics="true" pageId="8" pageNumber="155">
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||||
<typeStatus id="2F3C88190D42FF8FFCBC74B982ECA113" box="[812,900,1535,1557]" pageId="8" pageNumber="155" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
:
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||||
<specimenCode id="A0219EC00D42FF8FFC0174B9828DA113" box="[913,997,1535,1557]" collectionCode="LIGG" pageId="8" pageNumber="155">LIGG 25</specimenCode>
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−A−2459, trunk scale (
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<figureCitation id="68BC2A3E0D42FF8FFB2B74B9846FA113" box="[1211,1287,1535,1557]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. A–D. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25−A−2457. B. LIGG 25−A−2458. C. Holotype, LIGG 25−A−2459. D. LIGG 25−A−2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="8" pageNumber="155">Fig. 8C</figureCitation>
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).
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</materialsCitation>
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||||
</paragraph>
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||||
<paragraph id="F03836BB0D42FF8FFCBC776484F6A131" blockId="8.[812,1485,1535,1654]" box="[812,1438,1569,1591]" pageId="8" pageNumber="155">
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||||
<emphasis id="C2F3EAA90D42FF8FFCBC776482C1A131" box="[812,937,1569,1591]" italics="true" pageId="8" pageNumber="155">Type locality</emphasis>
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||||
: Kurtuvėnai−162 borehole, depth
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<quantity id="377F9B5E0D42FF8FFB7E776784F1A131" box="[1262,1433,1569,1591]" metricMagnitude="3" metricUnit="m" metricValue="1.10455" metricValueMax="1.1178" metricValueMin="1.0913" pageId="8" pageNumber="155" unit="m" value="1104.55" valueMax="1117.8" valueMin="1091.3">1091.3–1117.8 m</quantity>
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||||
.
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||||
</paragraph>
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||||
<paragraph id="F03836BB0D42FF8FFCBC770282A9A170" blockId="8.[812,1485,1535,1654]" pageId="8" pageNumber="155">
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||||
<emphasis id="C2F3EAA90D42FF8FFCBC770282C4A15C" box="[812,940,1604,1626]" italics="true" pageId="8" pageNumber="155">Type horizon</emphasis>
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||||
: Ventspils and Minija formations, Ludlow and Pridoli, Upper Silurian.
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</paragraph>
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||||
<paragraph id="F03836BB0D42FF8FFCBC77CE851DA1BF" blockId="8.[812,1485,1671,1721]" pageId="8" pageNumber="155">
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||||
<emphasis id="C2F3EAA90D42FF8FFCBC77CE8201A19B" box="[812,873,1672,1693]" italics="true" pageId="8" pageNumber="155">Range</emphasis>
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:UpperSilurian,Ludlow,PagėgiaiRegionalStage,toPridoli,the lower part of Jūra Regional Stage.
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</paragraph>
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||||
<paragraph id="F03836BB0D42FF8FFCBC778D82CDA1FA" blockId="8.[812,1486,1738,1788]" pageId="8" pageNumber="155">
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||||
<emphasis id="C2F3EAA90D42FF8FFCBC778D8280A1E6" box="[812,1000,1738,1760]" italics="true" pageId="8" pageNumber="155">Derivation of name</emphasis>
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||||
: From Latin
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<emphasis id="C2F3EAA90D42FF8FFBF8778C85CAA1E6" box="[1128,1186,1738,1760]" italics="true" pageId="8" pageNumber="155">brevis</emphasis>
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, short, and
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<emphasis id="C2F3EAA90D42FF8FFA84778D8435A1E6" box="[1300,1373,1739,1760]" italics="true" pageId="8" pageNumber="155">ornatus</emphasis>
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decorated, ornamented.
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</paragraph>
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<paragraph id="F03836BB0D42FF8FFCBC764A85BFA023" blockId="8.[812,1485,1804,2026]" box="[812,1239,1804,1829]" pageId="8" pageNumber="155">
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<emphasis id="C2F3EAA90D42FF8FFCBC764A82E4A023" box="[812,908,1804,1829]" italics="true" pageId="8" pageNumber="155">Material</emphasis>
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. Total more than 2100 scales.
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</paragraph>
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<paragraph id="F03836BB0D42FF8EFCBC76738089A060" blockId="8.[812,1485,1804,2026]" lastBlockId="9.[102,775,1838,2026]" lastPageId="9" lastPageNumber="156" pageId="8" pageNumber="155">
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<emphasis id="C2F3EAA90D42FF8FFCBC767382F3A04A" box="[812,923,1845,1868]" italics="true" pageId="8" pageNumber="155">Diagnosis</emphasis>
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.—
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<taxonomicName id="37874D380D42FF8FFC2E76728529A04B" authorityName="Valiukevicius" authorityYear="2004" box="[958,1089,1844,1869]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="8" pageNumber="155" phylum="Chordata" rank="genus">
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<emphasis id="C2F3EAA90D42FF8FFC2E76728529A04B" box="[958,1089,1844,1869]" italics="true" pageId="8" pageNumber="155">Rohonilepis</emphasis>
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</taxonomicName>
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with tiny scales sculptured by 6 to 10 short, sharp, symmetrical sub−parallel anterior ridges turning anteriorly toward the neck−base junction. The two longest posteriorly pointing ridges can form an elevated medial area separating narrow, lowered lateral folds. Crown dentine contains medium−branched linear ascending and simple widened radial vascular canals over the flat−pyramidal base, that is composed of dense acellular bone.
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</paragraph>
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<caption id="A4F866330D43FF8EFFF6773282BDA1F1" pageId="9" pageNumber="156" startId="9.[102,135,1652,1671]" targetBox="[104,1482,229,1625]" targetPageId="9" targetType="figure">
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||||
<paragraph id="F03836BB0D43FF8EFFF6773282BDA1F1" blockId="9.[102,1486,1652,1784]" pageId="9" pageNumber="156">
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||||
Fig. 7.
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<taxonomicName id="37874D380D43FF8EFF34773280FFA181" authorityName="Valiukevičius" authorityYear="2004" box="[164,407,1652,1671]" class="Acanthodii" family="Ischnacanthidae" genus="Bracteatacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthiformes" pageId="9" pageNumber="156" phylum="Chordata" rank="species" species="assiduus" status="gen. et sp. nov.">
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||||
<emphasis id="C2F3EAA90D43FF8EFF34773280FFA181" box="[164,407,1652,1671]" italics="true" pageId="9" pageNumber="156">Bracteatacanthus assiduus</emphasis>
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</taxonomicName>
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||||
<taxonomicNameLabel id="D9C057D20D43FF8EFE0C7733834CA18E" box="[412,548,1653,1672]" pageId="9" pageNumber="156" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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||||
<emphasis id="C2F3EAA90D43FF8EFDB77732833AA18E" bold="true" box="[551,594,1652,1672]" pageId="9" pageNumber="156">A–E</emphasis>
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. SEM micrographs of trunk scales; crown views, anterior upwards, except for
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<emphasis id="C2F3EAA90D43FF8EFA8277338449A18E" bold="true" box="[1298,1313,1653,1672]" pageId="9" pageNumber="156">B</emphasis>
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, lateral view, anterior to the left.
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<emphasis id="C2F3EAA90D43FF8EFF7977D68191A1A5" bold="true" box="[233,249,1680,1699]" pageId="9" pageNumber="156">A</emphasis>
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||||
. LIGG 25−A−2450.
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<emphasis id="C2F3EAA90D43FF8EFE3C77D780D3A1A2" bold="true" box="[428,443,1681,1700]" pageId="9" pageNumber="156">B</emphasis>
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||||
. LIGG 25−A−2451.
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<emphasis id="C2F3EAA90D43FF8EFDFD77D68315A1A5" bold="true" box="[621,637,1680,1699]" pageId="9" pageNumber="156">C</emphasis>
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||||
. LIGG 25−A−2452.
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<emphasis id="C2F3EAA90D43FF8EFCA077D78228A1A2" bold="true" box="[816,832,1681,1700]" pageId="9" pageNumber="156">D</emphasis>
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||||
. Holotype, LIGG 25−A−2453.
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<emphasis id="C2F3EAA90D43FF8EFBC177D78508A1A2" bold="true" box="[1105,1120,1681,1700]" pageId="9" pageNumber="156">E</emphasis>
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||||
. LIGG 25−A−2454.
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<emphasis id="C2F3EAA90D43FF8EFA8377D78449A1A2" bold="true" box="[1299,1313,1681,1700]" pageId="9" pageNumber="156">F</emphasis>
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||||
,
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<emphasis id="C2F3EAA90D43FF8EFABB77D68455A1A5" bold="true" box="[1323,1341,1680,1699]" pageId="9" pageNumber="156">G</emphasis>
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||||
. Microstructure of scales in vertical longitudinal sections. F
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||||
<subScript id="6C0334FE0D43FF8EFE7777F38087A1C5" attach="left" box="[487,495,1717,1731]" fontSize="6" pageId="9" pageNumber="156">1</subScript>
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||||
. Thin section LIGG 3705, general view. F
|
||||
<subScript id="6C0334FE0D43FF8EFCF977F38219A1C5" attach="left" box="[873,881,1717,1731]" fontSize="6" pageId="9" pageNumber="156">2</subScript>
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||||
. Detail of the left part of crown and base apex of the same scale. G
|
||||
<subScript id="6C0334FE0D43FF8EFA2F77F384AFA1C5" attach="left" box="[1471,1479,1717,1731]" fontSize="6" pageId="9" pageNumber="156">1</subScript>
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||||
. Thin section LIGG 3708. G
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||||
<subScript id="6C0334FE0D43FF8EFECC7797800CA1D9" attach="left" box="[348,356,1745,1759]" fontSize="6" pageId="9" pageNumber="156">2</subScript>
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||||
. Detail of the left part of crown and base apex of the same scale. G
|
||||
<subScript id="6C0334FE0D43FF8EFC24779782D4A1D9" attach="left" box="[948,956,1745,1759]" fontSize="6" pageId="9" pageNumber="156">3</subScript>
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. The same scale, detail of the right part. Scale bars 0.1 mm. Nida−44 borehole, depth 1213.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation.
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</paragraph>
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</caption>
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<paragraph id="F03836BB0D43FF8DFFF6763285F0A08E" blockId="9.[102,775,1838,2026]" lastBlockId="10.[812,1485,1841,2026]" lastPageId="10" lastPageNumber="157" pageId="9" pageNumber="156">
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<emphasis id="C2F3EAA90D43FF8EFFF676328180A08D" box="[102,232,1908,1931]" italics="true" pageId="9" pageNumber="156">Description</emphasis>
|
||||
.—Tiny scales with crowns that only rarely reach
|
||||
<quantity id="377F9B5E0D43FF8EFFF676D481D0A0AC" box="[102,184,1938,1963]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="156" unit="mm" value="0.5">0.5 mm</quantity>
|
||||
in length and scale width varies from
|
||||
<quantity id="377F9B5E0D43FF8EFDC076D5826BA0AC" box="[592,771,1939,1963]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" metricValueMax="3.8" metricValueMin="2.2" pageId="9" pageNumber="156" unit="mm" value="0.3" valueMax="0.38" valueMin="0.22">0.22 to 0.38 mm</quantity>
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||||
. The crown plate is horizontal, with up to one−third of the posterior unsculptured part overhanging the base. A quarter to one−third of the crown length has 6 to 10 short, sharp anterior ridges with a symmetrical profile, which fade posteriorly. Crown ridges may be straight (
|
||||
<figureCitation id="68BC2A3E0D43FF8EFB10762985BFA08E" box="[1152,1239,1903,1928]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. A–D. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25−A−2457. B. LIGG 25−A−2458. C. Holotype, LIGG 25−A−2459. D. LIGG 25−A−2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="9" pageNumber="156">Fig. 8D</figureCitation>
|
||||
) or slightly curved towards mid−scale (
|
||||
<figureCitation id="68BC2A3E0D43FF8EFC6376D68506A0AE" box="[1011,1134,1936,1961]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. A–D. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25−A−2457. B. LIGG 25−A−2458. C. Holotype, LIGG 25−A−2459. D. LIGG 25−A−2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="9" pageNumber="156">Fig. 8B, C</figureCitation>
|
||||
). The holotype−like specimens, with sub−parallel ridges are more abundant in samples studied, however, scales with almost parallel ridges also are known. Ridges are characteristically downwardly sloped anteriorly, sometimes reaching the base of the neck. Grooves between ridges are moderately deep and wide. Some scales have narrow lowered lateral ledges (
|
||||
<figureCitation id="68BC2A3E0D40FF8DFE0176D7836EA0AC" box="[401,518,1937,1962]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. A–D. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25−A−2457. B. LIGG 25−A−2458. C. Holotype, LIGG 25−A−2459. D. LIGG 25−A−2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="10" pageNumber="157">Fig. 8A, C</figureCitation>
|
||||
), separated from the remainder of the crown by the two longest, posteriorly pointing ridges. Moderately high to low neck usually has linear rows of pores. The scale base slightly protrudes beyond the crown antero−laterally, but may be also displaced far in advance of the anterior margin of the crown.
|
||||
</paragraph>
|
||||
<caption id="A4F866330D40FF8DFFF677E383CAA00A" pageId="10" pageNumber="157" startId="10.[102,135,1701,1720]" targetBox="[107,1482,225,1674]" targetPageId="10" targetType="figure">
|
||||
<paragraph id="F03836BB0D40FF8DFFF677E383CAA00A" blockId="10.[102,1486,1701,1805]" pageId="10" pageNumber="157">
|
||||
Fig. 8.
|
||||
<taxonomicName id="37874D380D40FF8DFF3B77E380FBA1BE" authorityName="Valiukevičius" authorityYear="2004" box="[171,403,1701,1720]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="10" pageNumber="157" phylum="Chordata" rank="species" species="breviornatus" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D40FF8DFF3B77E380FBA1BE" box="[171,403,1701,1720]" italics="true" pageId="10" pageNumber="157">Rohonilepis breviornatus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D40FF8DFE0B77E08345A1BF" box="[411,557,1702,1721]" pageId="10" pageNumber="157" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="C2F3EAA90D40FF8DFDA577E38308A1BE" bold="true" box="[565,608,1701,1721]" pageId="10" pageNumber="157">A–D</emphasis>
|
||||
. SEM micrographs of scales, crown views, anterior upwards.
|
||||
<emphasis id="C2F3EAA90D40FF8DFB3B77E385D3A1BE" bold="true" box="[1195,1211,1701,1720]" pageId="10" pageNumber="157">A</emphasis>
|
||||
. LIGG 25−A−2457.
|
||||
<emphasis id="C2F3EAA90D40FF8DFAE977E384E0A1BE" bold="true" box="[1401,1416,1701,1720]" pageId="10" pageNumber="157">B</emphasis>
|
||||
. LIGG 25−A−2458.
|
||||
<emphasis id="C2F3EAA90D40FF8DFF457787818DA1D2" bold="true" box="[213,229,1729,1748]" pageId="10" pageNumber="157">C</emphasis>
|
||||
. Holotype, LIGG 25−A−2459.
|
||||
<emphasis id="C2F3EAA90D40FF8DFE6B77878363A1D2" bold="true" box="[507,523,1729,1748]" pageId="10" pageNumber="157">D</emphasis>
|
||||
. LIGG 25−A−2461.
|
||||
<emphasis id="C2F3EAA90D40FF8DFD52778783B9A1D2" bold="true" box="[706,721,1729,1748]" pageId="10" pageNumber="157">E</emphasis>
|
||||
,
|
||||
<emphasis id="C2F3EAA90D40FF8DFD4D77878383A1D2" bold="true" box="[733,747,1729,1748]" pageId="10" pageNumber="157">F</emphasis>
|
||||
. Microstructure of scales in vertical longitudinal sections. E
|
||||
<subScript id="6C0334FE0D40FF8DFA9D778C847DA1DE" attach="left" box="[1293,1301,1738,1752]" fontSize="6" pageId="10" pageNumber="157">1</subScript>
|
||||
. Thin section LIGG 3726, general view. E
|
||||
<subScript id="6C0334FE0D40FF8DFEBC77A0805CA1F2" attach="left" box="[300,308,1766,1780]" fontSize="6" pageId="10" pageNumber="157">2</subScript>
|
||||
. Detail of the upper right crown part of the same scale.
|
||||
<emphasis id="C2F3EAA90D40FF8DFCBD779B8253A1F6" bold="true" box="[813,827,1757,1776]" pageId="10" pageNumber="157">F</emphasis>
|
||||
. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="F03836BB0D40FF8CFCC276D480DCA51C" blockId="10.[812,1485,1841,2026]" lastBlockId="11.[102,775,231,1277]" lastPageId="11" lastPageNumber="158" pageId="10" pageNumber="157">
|
||||
Histologic microstructure has not been sufficiently studied yet because of the preservation of the scales, but they are certainly of the
|
||||
<taxonomicName id="37874D380D40FF8DFC2B76978553A0EC" authorityName="Baly" authorityYear="1864" box="[955,1083,2001,2026]" class="Insecta" family="Chrysomelidae" genus="Acanthodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="157" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="C2F3EAA90D40FF8DFC2B76978553A0EC" box="[955,1083,2001,2026]" italics="true" pageId="10" pageNumber="157">Acanthodes</emphasis>
|
||||
</taxonomicName>
|
||||
−
|
||||
<typeStatus id="2F3C88190D40FF8DFBD97694851FA0EC" box="[1097,1143,2002,2026]" pageId="10" pageNumber="157">type</typeStatus>
|
||||
. At least, 6 growth lamellae in crowns composed of dentine can be counted (
|
||||
<figureCitation id="68BC2A3E0D41FF8CFDFA71A183D7A606" box="[618,703,231,256]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. A–D. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25−A−2457. B. LIGG 25−A−2458. C. Holotype, LIGG 25−A−2459. D. LIGG 25−A−2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="11" pageNumber="158">Fig. 8F</figureCitation>
|
||||
), with the oldest lamella unusually large. The vascular system is poorly defined due to re−mineralization of tissues and only short segments (
|
||||
<figureCitation id="68BC2A3E0D41FF8CFE8B70038006A658" box="[283,366,325,350]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. A–D. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25−A−2457. B. LIGG 25−A−2458. C. Holotype, LIGG 25−A−2459. D. LIGG 25−A−2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="11" pageNumber="158">Fig. 8E</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D41FF8CFEFE7014801FA665" box="[366,375,338,355]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="11" pageNumber="158">
|
||||
<subScript id="6C0334FE0D41FF8CFEFE7014801FA665" attach="left" box="[366,375,338,355]" fontSize="7" pageId="11" pageNumber="158">2</subScript>
|
||||
</figureCitation>
|
||||
, F) of ascending and other vascular canals are visible. Ascending canals are moderately branched and mostly linear along growth lamellae. Radial vascular canals are simple and located just over the basal surface (
|
||||
<figureCitation id="68BC2A3E0D41FF8CFD2070E28269A6BA" box="[688,769,420,445]" captionStart="Fig" captionStartId="10.[102,135,1701,1720]" captionTargetBox="[107,1482,225,1674]" captionTargetId="figure-122@10.[104,1483,225,1673]" captionTargetPageId="10" captionText="Fig. 8. Rohonilepis breviornatus gen. et sp. nov. A–D. SEM micrographs of scales, crown views, anterior upwards. A. LIGG 25−A−2457. B. LIGG 25−A−2458. C. Holotype, LIGG 25−A−2459. D. LIGG 25−A−2461. E, F. Microstructure of scales in vertical longitudinal sections. E1. Thin section LIGG 3726, general view. E2. Detail of the upper right crown part of the same scale. F. Thin section LIGG 3727. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1103.0 m. Upper Silurian, Ludlow, Ventspils Formation." pageId="11" pageNumber="158">Fig. 8F</figureCitation>
|
||||
, right side). Durodentine has not been definitely recognized, but may be present. Acellular basal bone is dense and forms a flat−pyramid below the crown.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D41FF8CFFF6736D8096A49D" blockId="11.[102,775,231,1277]" pageId="11" pageNumber="158">
|
||||
<emphasis id="C2F3EAA90D41FF8CFFF6736D81B6A544" box="[102,222,555,578]" italics="true" pageId="11" pageNumber="158">Discussion</emphasis>
|
||||
.—
|
||||
<taxonomicName id="37874D380D41FF8CFE91736F8371A544" authorityName="Valiukevičius" authorityYear="2004" box="[257,537,553,578]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="11" pageNumber="158" phylum="Chordata" rank="species" species="breviornatus">
|
||||
<emphasis id="C2F3EAA90D41FF8CFE91736F8371A544" box="[257,537,553,578]" italics="true" pageId="11" pageNumber="158">Rohonilepis breviornatus</emphasis>
|
||||
</taxonomicName>
|
||||
is comparable to the above described
|
||||
<taxonomicName id="37874D380D41FF8CFE8E730F832EA564" authorityName="Valiukevičius" authorityYear="2004" box="[286,582,585,610]" class="Acanthodii" family="Ischnacanthidae" genus="Bracteatacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthiformes" pageId="11" pageNumber="158" phylum="Chordata" rank="species" species="assiduus">
|
||||
<emphasis id="C2F3EAA90D41FF8CFE8E730F832EA564" box="[286,582,585,610]" italics="true" pageId="11" pageNumber="158">Bracteatacanthus assiduus</emphasis>
|
||||
</taxonomicName>
|
||||
, as histologically they both have many characters in common. The latter is distinguished by the peculiar arrangement and position of radial vascular canals in the dentine of the crown, that is definitely not present in
|
||||
<emphasis id="C2F3EAA90D41FF8CFE92738E80DCA5D9" box="[258,436,710,735]" italics="true" pageId="11" pageNumber="158">
|
||||
<taxonomicName id="37874D380D41FF8CFE92738E80D8A5D9" authorityName="Valiukevičius" authorityYear="2004" box="[258,432,710,735]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="11" pageNumber="158" phylum="Chordata" rank="species" species="breviornatus">R. breviornatus</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
Morphological differences are more substantial: asymmetrically sloped (in profile) rounded ridges do not extend baseward in
|
||||
<taxonomicName id="37874D380D41FF8CFE40724383FAA418" authorityName="Valiukevicius" authorityYear="2004" box="[464,658,773,798]" class="Acanthodii" family="Ischnacanthidae" genus="Bracteatacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthiformes" pageId="11" pageNumber="158" phylum="Chordata" rank="genus">
|
||||
<emphasis id="C2F3EAA90D41FF8CFE40724383FAA418" box="[464,658,773,798]" italics="true" pageId="11" pageNumber="158">Bracteatacanthus</emphasis>
|
||||
</taxonomicName>
|
||||
scales and sharper, symmetric, frequently centrally curved ridges sometimes reach the neck base in
|
||||
<taxonomicName id="37874D380D41FF8CFE227202835DA45B" authorityName="Valiukevicius" authorityYear="2004" box="[434,565,836,861]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="11" pageNumber="158" phylum="Chordata" rank="genus">
|
||||
<emphasis id="C2F3EAA90D41FF8CFE227202835DA45B" box="[434,565,836,861]" italics="true" pageId="11" pageNumber="158">Rohonilepis</emphasis>
|
||||
</taxonomicName>
|
||||
. The lateral lower step−like margins of
|
||||
<taxonomicName id="37874D380D41FF8CFED87223809BA47A" authorityName="Valiukevičius" authorityYear="2004" box="[328,499,867,892]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="11" pageNumber="158" phylum="Chordata" rank="species" species="breviornatus">
|
||||
<emphasis id="C2F3EAA90D41FF8CFED87223809BA47A" box="[328,499,867,892]" italics="true" pageId="11" pageNumber="158">R. breviornatus</emphasis>
|
||||
</taxonomicName>
|
||||
scales have not been observed on those of
|
||||
<taxonomicName id="37874D380D41FF8CFEA572C5809FA49A" authorityName="Valiukevicius" authorityYear="2004" box="[309,503,899,924]" class="Acanthodii" family="Ischnacanthidae" genus="Bracteatacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthiformes" pageId="11" pageNumber="158" phylum="Chordata" rank="genus">
|
||||
<emphasis id="C2F3EAA90D41FF8CFEA572C5809FA49A" box="[309,503,899,924]" italics="true" pageId="11" pageNumber="158">Bracteatacanthus</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D41FF8CFF1B72E48317A371" blockId="11.[102,775,231,1277]" pageId="11" pageNumber="158">
|
||||
The sculpture of some scales of
|
||||
<taxonomicName id="37874D380D41FF8CFE7872E583FAA4BD" authorityName="Valiukevičius" authorityYear="2004" box="[488,658,930,955]" class="Acanthodii" family="Ischnacanthidae" genus="Rohonilepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ischnacanthida" pageId="11" pageNumber="158" phylum="Chordata" rank="species" species="breviornatus">
|
||||
<emphasis id="C2F3EAA90D41FF8CFE7872E583FAA4BD" box="[488,658,930,955]" italics="true" pageId="11" pageNumber="158">R. breviornatus</emphasis>
|
||||
</taxonomicName>
|
||||
resembles several specimens of
|
||||
<taxonomicName id="37874D380D41FF8CFEDC72848083A4DD" authorityName="Gross" authorityYear="1971" box="[332,491,962,987]" class="Acanthodii" family="Ischnacanthidae" genus="Gomphonchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="11" pageNumber="158" phylum="Chordata" rank="genus">
|
||||
<emphasis id="C2F3EAA90D41FF8CFEDC72848083A4DD" box="[332,491,962,987]" italics="true" pageId="11" pageNumber="158">Gomphonchus</emphasis>
|
||||
</taxonomicName>
|
||||
from the Hviždalka locality,
|
||||
<collectingCountry id="8890762B0D41FF8CFF0372A7802CA4FC" box="[147,324,993,1018]" name="Czech Republic" pageId="11" pageNumber="158">Czech Republic</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="94164B4A0D41FF8CFEC672A480B7A4FC" author="Marss, T." box="[342,479,994,1018]" pageId="11" pageNumber="158" pagination="17 - 41" refId="ref9224" refString="Marss, T. 1997. Vertebrates of the Pridoli and Silurian - Devonian boundary beds in Europe. Modern Geology 21: 17 - 41." type="journal article" year="1997">Märss 1997</bibRefCitation>
|
||||
: pl. 7: 3, 10).
|
||||
<bibRefCitation id="94164B4A0D41FF8CFD1572A481C7A31F" author="Vergoossen, J. M. J." pageId="11" pageNumber="158" pagination="93 - 158" refId="ref9479" refString="Vergoossen, J. M. J. 2002. Late Silurian fish microfossils from Ramsasa (sites D and " south of church "), Skane, south Sweden. Scripta Geologica 123: 93 - 158." type="journal article" year="2002">Vergoossen (2002)</bibRefCitation>
|
||||
illustrated specimens of
|
||||
<taxonomicName id="37874D380D41FF8CFE597546832BA31F" box="[457,579,1024,1049]" class="Acanthodii" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="158" phylum="Chordata" rank="class">Acanthodii</taxonomicName>
|
||||
gen. et sp. indet. that are similar but not as well preserved as the material figured here, but suggest, that some comparable varieties of scales might also be present in southern
|
||||
<collectingCountry id="8890762B0D41FF8CFDB075188313A371" box="[544,635,1118,1143]" name="Sweden" pageId="11" pageNumber="158">Sweden</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D41FF8CFFF675C18385A3FA" blockId="11.[102,775,231,1277]" pageId="11" pageNumber="158">
|
||||
<emphasis id="C2F3EAA90D41FF8CFFF675C1818DA398" box="[102,229,1159,1182]" italics="true" pageId="11" pageNumber="158">Occurrence</emphasis>
|
||||
.—Kurtuvėnai−162borehole,depth
|
||||
<quantity id="377F9B5E0D41FF8CFDF175C08116A3B8" metricMagnitude="3" metricUnit="m" metricValue="1.1144" metricValueMax="1.1375" metricValueMin="1.0913" pageId="11" pageNumber="158" unit="m" value="1114.4" valueMax="1137.5" valueMin="1091.3">1091.3–1137.5 m</quantity>
|
||||
; Sutkai−87:
|
||||
<quantity id="377F9B5E0D41FF8CFE9A75E080DDA3B8" box="[266,437,1190,1215]" metricMagnitude="2" metricUnit="m" metricValue="6.1825" metricValueMax="6.236" metricValueMin="6.129" pageId="11" pageNumber="158" unit="m" value="618.25" valueMax="623.6" valueMin="612.9">612.9–623.6 m</quantity>
|
||||
; Gėluva−99:
|
||||
<quantity id="377F9B5E0D41FF8CFDDB75E0826AA3B8" box="[587,770,1190,1215]" metricMagnitude="2" metricUnit="m" metricValue="6.697" metricValueMax="6.853" metricValueMin="6.5409999999999995" pageId="11" pageNumber="158" unit="m" value="669.7" valueMax="685.3" valueMin="654.1">654.1– 685.3 m</quantity>
|
||||
; Stoniškiai−1: 1224.0–
|
||||
<quantity id="377F9B5E0D41FF8CFEDD758080DCA3DB" box="[333,436,1222,1246]" metricMagnitude="3" metricUnit="m" metricValue="1.3089" pageId="11" pageNumber="158" unit="m" value="1308.9">1308.9 m</quantity>
|
||||
; Šešuvis−11: 1047.0–
|
||||
<quantity id="377F9B5E0D41FF8CFD0B7583826AA3DB" box="[667,770,1221,1246]" metricMagnitude="3" metricUnit="m" metricValue="1.0495" pageId="11" pageNumber="158" unit="m" value="1049.5">1049.5 m</quantity>
|
||||
; Ledai−179:
|
||||
<quantity id="377F9B5E0D41FF8CFF7275A280EFA3FA" box="[226,391,1252,1277]" metricMagnitude="2" metricUnit="m" metricValue="5.349000000000001" metricValueMax="5.441000000000001" metricValueMin="5.257000000000001" pageId="11" pageNumber="158" unit="m" value="534.9" valueMax="544.1" valueMin="525.7">525.7–544.1 m</quantity>
|
||||
; Liepkalnis−137:
|
||||
<quantity id="377F9B5E0D41FF8CFDD375A38381A3FA" box="[579,745,1253,1277]" metricMagnitude="2" metricUnit="m" metricValue="9.0465" metricValueMax="9.092" metricValueMin="9.001" pageId="11" pageNumber="158" unit="m" value="904.65" valueMax="909.2" valueMin="900.1">900.1–909.2 m</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
338
data/78/2E/87/782E87AD0D4BFF84FFF67508841DA1B5.xml
Normal file
338
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<mods:title id="C0AE0736FCDBEDED88174F3223245BE0">New Wenlock-Pridoli (Silurian) acanthodian fishes from Lithuania</mods:title>
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<mods:namePart id="019F8B68A3A44D4DB35FC300230F871F">Valiukevičius, Juozas</mods:namePart>
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<mods:title id="A182FB4CAE5C551D0750CE66B78980CD">Acta Palaeontologica Polonica</mods:title>
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<mods:part id="F5FDE2A8C098AED8D48DF7C3EBCA0DE5">
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<mods:date id="7375EF324A237121D9914BF6876AA60F">2004</mods:date>
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<mods:number id="9D1E8DC89CE4463237F0A7266B01653A">2004-12-31</mods:number>
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<treatment id="782E87AD0D4BFF84FFF67508841DA1B5" LSID="urn:lsid:plazi:treatment:782E87AD0D4BFF84FFF67508841DA1B5" httpUri="http://treatment.plazi.org/id/782E87AD0D4BFF84FFF67508841DA1B5" lastPageId="3" lastPageNumber="150" pageId="1" pageNumber="148">
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<subSubSection id="B89D65300D4BFF86FFF67508835AA36D" box="[102,562,1102,1131]" pageId="1" pageNumber="148" type="nomenclature">
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<paragraph id="F03836BB0D4BFF86FFF67508835AA36D" blockId="1.[102,562,1102,1163]" box="[102,562,1102,1131]" pageId="1" pageNumber="148">
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<heading id="AB7081D70D4BFF86FFF67508835AA36D" box="[102,562,1102,1131]" fontSize="12" level="2" pageId="1" pageNumber="148" reason="9">
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<taxonomicName id="37874D380D4BFF86FFF675088000A36D" authorityName="Valiukevičius" authorityYear="2004" box="[102,360,1102,1131]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="1" pageNumber="148" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D4BFF86FFF675088000A36D" box="[102,360,1102,1131]" italics="true" pageId="1" pageNumber="148">Vesperalia perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D4BFF86FEE07509835AA36D" box="[368,562,1103,1131]" pageId="1" pageNumber="148" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
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</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="B89D65300D4BFF84FFF67533841DA1B5" lastPageId="3" lastPageNumber="150" pageId="1" pageNumber="148" type="description">
|
||||
<paragraph id="F03836BB0D4BFF86FFF6753381ADA38D" blockId="1.[102,562,1102,1163]" box="[102,197,1141,1163]" pageId="1" pageNumber="148">
|
||||
<figureCitation id="68BC2A3E0D4BFF86FFF6753381C1A38D" box="[102,169,1141,1163]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="1" pageNumber="148">Figs. 2</figureCitation>
|
||||
,
|
||||
<figureCitation id="68BC2A3E0D4BFF86FF23753381A9A38D" box="[179,193,1141,1163]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="1" pageNumber="148">3</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4BFF86FCBC71A1847CA7FB" blockId="1.[812,1486,231,1164]" box="[812,1300,231,253]" pageId="1" pageNumber="148">
|
||||
<materialsCitation id="40EF3CE60D4BFF86FCBC71A1847CA7FB" box="[812,1300,231,253]" collectionCode="LIGG" pageId="1" pageNumber="148" specimenCode="LIGG 25" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="C2F3EAA90D4BFF86FCBC71A182ECA7FB" box="[812,900,231,253]" italics="true" pageId="1" pageNumber="148">
|
||||
<typeStatus id="2F3C88190D4BFF86FCBC71A182ECA7FB" box="[812,900,231,253]" pageId="1" pageNumber="148" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCode id="A0219EC00D4BFF86FC0171A1828EA7FB" box="[913,998,231,253]" collectionCode="LIGG" pageId="1" pageNumber="148">LIGG 25</specimenCode>
|
||||
−A−2413, trunk scale (
|
||||
<figureCitation id="68BC2A3E0D4BFF86FB2B71A18461A7FB" box="[1211,1289,231,253]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="1" pageNumber="148">Fig. 2A</figureCitation>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4BFF86FCBC704F8412A618" blockId="1.[812,1486,231,1164]" box="[812,1402,264,286]" pageId="1" pageNumber="148">
|
||||
<emphasis id="C2F3EAA90D4BFF86FCBC704F82C1A618" box="[812,937,264,286]" italics="true" pageId="1" pageNumber="148">Type locality</emphasis>
|
||||
: Stoniškiai−1 borehole, depth 1211.0–1217.0 m.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4BFF86FCBC706C8546A667" blockId="1.[812,1486,231,1164]" pageId="1" pageNumber="148">
|
||||
<emphasis id="C2F3EAA90D4BFF86FCBC706C82CAA639" box="[812,930,297,319]" italics="true" pageId="1" pageNumber="148">Type horizon</emphasis>
|
||||
:RietavasBedsoftheJūraFormation,Pridoli,UpperSilurian.
|
||||
<emphasis id="C2F3EAA90D4BFF86FCBC700A8201A667" box="[812,873,332,353]" italics="true" pageId="1" pageNumber="148">Range</emphasis>
|
||||
:
|
||||
<typeStatus id="2F3C88190D4BFF86FCE6700D82CFA667" box="[886,935,331,353]" pageId="1" pageNumber="148">Type</typeStatus>
|
||||
horizon only.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4BFF86FCBC702B841FA684" blockId="1.[812,1486,231,1164]" box="[812,1399,364,386]" pageId="1" pageNumber="148">
|
||||
<emphasis id="C2F3EAA90D4BFF86FCBC702B828FA684" box="[812,999,364,386]" italics="true" pageId="1" pageNumber="148">Derivation of name</emphasis>
|
||||
: From Latin
|
||||
<emphasis id="C2F3EAA90D4BFF86FBF6702A85ABA684" box="[1126,1219,364,386]" italics="true" pageId="1" pageNumber="148">perplexus</emphasis>
|
||||
, tangled, intricate.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4BFF86FCBC70D785C4A6AC" blockId="1.[812,1486,231,1164]" box="[812,1196,401,426]" pageId="1" pageNumber="148">
|
||||
<emphasis id="C2F3EAA90D4BFF86FCBC70D782E4A6AC" box="[812,908,401,426]" italics="true" pageId="1" pageNumber="148">Material</emphasis>
|
||||
.—Total about 600 scales.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4BFF86FCBC70FC8528A40D" blockId="1.[812,1486,231,1164]" pageId="1" pageNumber="148">
|
||||
<emphasis id="C2F3EAA90D4BFF86FCBC70FC82F3A6D7" box="[812,923,442,465]" italics="true" pageId="1" pageNumber="148">Diagnosis</emphasis>
|
||||
.—
|
||||
<taxonomicName id="37874D380D4BFF86FC2E70FF855DA6D4" authorityName="Valiukevicius" authorityYear="2004" box="[958,1077,441,466]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="1" pageNumber="148" phylum="Chordata" rank="genus">
|
||||
<emphasis id="C2F3EAA90D4BFF86FC2E70FF855DA6D4" box="[958,1077,441,466]" italics="true" pageId="1" pageNumber="148">Vesperalia</emphasis>
|
||||
</taxonomicName>
|
||||
with high−crowned scales, porous neck and thick crown with a sculpture of four to six linear or wavy longitudinal ridges, of which two converging ones may form a raised medial area. Ridges usually cross the entire length of the crown or fade out mid−length, continuing after a smooth area on the posterior edge of the crown. Scales grow superpositionally or both superpositionally and areally. Stranggewebe with unusually large oriented lacunae and simple mesodentine form both posterior and anterior parts of the crown. The crown contains a system of complicated and enlarged vascular canals.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4BFF85FCBC725D8409A08F" blockId="1.[812,1486,231,1164]" lastBlockId="2.[812,1486,1807,2026]" lastPageId="2" lastPageNumber="149" pageId="1" pageNumber="148">
|
||||
<emphasis id="C2F3EAA90D4BFF86FCBC725D82C6A434" box="[812,942,795,818]" italics="true" pageId="1" pageNumber="148">Description</emphasis>
|
||||
.— Most trunk scales have rounded rhomboidal crowns varying from isometric to elongate, with length of
|
||||
<quantity id="377F9B5E0D4BFF86FCBC721F82D4A474" box="[812,956,857,882]" metricMagnitude="-4" metricUnit="m" metricValue="6.85" metricValueMax="8.7" metricValueMin="5.0" pageId="1" pageNumber="148" unit="mm" value="0.685" valueMax="0.87" valueMin="0.5">0.5–0.87mm</quantity>
|
||||
andwidth of
|
||||
<quantity id="377F9B5E0D4BFF86FBDB721F8588A474" box="[1099,1248,857,882]" metricMagnitude="-4" metricUnit="m" metricValue="7.55" metricValueMax="9.0" metricValueMin="6.1" pageId="1" pageNumber="148" unit="mm" value="0.755" valueMax="0.9" valueMin="0.61">0.61–0.9mm</quantity>
|
||||
. Thelength andwidth of the larger scale specimens is
|
||||
<quantity id="377F9B5E0D4BFF86FB3A723F8427A496" box="[1194,1359,889,913]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.47" metricValueMin="0.93" pageId="1" pageNumber="148" unit="mm" value="1.2" valueMax="1.47" valueMin="0.93">0.93–1.47 mm</quantity>
|
||||
and
|
||||
<quantity id="377F9B5E0D4BFF86FA1F723E82EEA4A9" metricMagnitude="-3" metricUnit="m" metricValue="1.1749999999999998" metricValueMax="1.4" metricValueMin="0.95" pageId="1" pageNumber="148" unit="mm" value="1.1749999999999998" valueMax="1.4" valueMin="0.95">0.95– 1.4 mm</quantity>
|
||||
, respectively. The crown surface is broad and flat, with slightly downsloped lateral and anterior margins. Four to six sharp sub−parallel straight or wavy ridges extend the whole crown length. The ridges become more robust towards the scale base.They are usually of the same height across the crown but may be more pronounced and higher posteriorly (
|
||||
<figureCitation id="68BC2A3E0D4BFF86FCA4751282A3A36A" box="[820,971,1108,1133]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="1" pageNumber="148">Fig. 2C, D, G</figureCitation>
|
||||
). The two most conspicuous ridges converge at the posterior tip of the crown and may form a slightly raised triangular medial plateau (
|
||||
<figureCitation id="68BC2A3E0D48FF85FE197649836BA02E" box="[393,515,1807,1832]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2B, D</figureCitation>
|
||||
), which has a sculpture of weak anterior ridgelets (
|
||||
<figureCitation id="68BC2A3E0D48FF85FE017676808EA04E" box="[401,486,1840,1865]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2B</figureCitation>
|
||||
) or has an anteriorly widened shallow groove (
|
||||
<figureCitation id="68BC2A3E0D48FF85FEC6761680C6A06F" box="[342,430,1872,1897]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2D</figureCitation>
|
||||
). The
|
||||
<typeStatus id="2F3C88190D48FF85FE627616833AA06F" box="[498,594,1872,1897]" pageId="2" pageNumber="149" type="holotype">holotype</typeStatus>
|
||||
and other specimens have ridges that fade out at the mid−length of the crown andafteracentralsmooth area,continueandincreaseinheight posteriorly (
|
||||
<figureCitation id="68BC2A3E0D48FF85FF7E76F78001A0CC" box="[238,361,1969,1994]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2A, E</figureCitation>
|
||||
). Of particular interest are the specimens with crown ridges and grooves on the anterior and posterior areas that do not join, but are shifted laterally relative to the long axis of the scale (
|
||||
<figureCitation id="68BC2A3E0D48FF85FBF7767685A9A04E" box="[1127,1217,1840,1865]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2E</figureCitation>
|
||||
). The grooves between ridges are of irregular depth and width, and sometimes form deeper troughs at their anterior end (
|
||||
<figureCitation id="68BC2A3E0D48FF85FB2C7636843CA08F" box="[1212,1364,1904,1929]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2E, G, H</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="A4F866330D4BFF86FFF676398328A0E0" pageId="1" pageNumber="148" startId="1.[102,135,1919,1938]" targetBox="[110,1473,1208,1887]" targetPageId="1" targetType="figure">
|
||||
<paragraph id="F03836BB0D4BFF86FFF676398328A0E0" blockId="1.[102,1485,1919,2022]" pageId="1" pageNumber="148">
|
||||
Fig. 2.
|
||||
<taxonomicName id="37874D380D4BFF86FF3876398008A094" authorityName="Valiukevičius" authorityYear="2004" box="[168,352,1919,1938]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="1" pageNumber="148" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D4BFF86FF3876398008A094" box="[168,352,1919,1938]" italics="true" pageId="1" pageNumber="148">Vesperalia perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D4BFF86FEF77639809DA094" box="[359,501,1919,1938]" pageId="1" pageNumber="148" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (
|
||||
<emphasis id="C2F3EAA90D4BFF86FF4676DD818EA0A8" bold="true" box="[214,230,1947,1966]" pageId="1" pageNumber="148">A</emphasis>
|
||||
), LIGG 25−A−2416 (
|
||||
<emphasis id="C2F3EAA90D4BFF86FE3676DD80DDA0A8" bold="true" box="[422,437,1947,1966]" pageId="1" pageNumber="148">B</emphasis>
|
||||
), LIGG 25−A−2417 (
|
||||
<emphasis id="C2F3EAA90D4BFF86FDE476DD83ECA0A8" bold="true" box="[628,644,1947,1966]" pageId="1" pageNumber="148">C</emphasis>
|
||||
), LIGG 25−A−2418 (
|
||||
<emphasis id="C2F3EAA90D4BFF86FCD376DD823BA0A8" bold="true" box="[835,851,1947,1966]" pageId="1" pageNumber="148">D</emphasis>
|
||||
), LIGG 25−A−2420 (
|
||||
<emphasis id="C2F3EAA90D4BFF86FB8376DD854AA0A8" bold="true" box="[1043,1058,1947,1966]" pageId="1" pageNumber="148">E</emphasis>
|
||||
), LIGG 25−A−2422 (
|
||||
<emphasis id="C2F3EAA90D4BFF86FB7176DD8587A0A8" bold="true" box="[1249,1263,1947,1966]" pageId="1" pageNumber="148">F</emphasis>
|
||||
), LIGG 25−A−2421 (
|
||||
<emphasis id="C2F3EAA90D4BFF86FA3E76DD84A8A0A8" bold="true" box="[1454,1472,1947,1966]" pageId="1" pageNumber="148">G</emphasis>
|
||||
), LIGG 25−A−2414(
|
||||
<emphasis id="C2F3EAA90D4BFF86FE9F76F18049A0CC" bold="true" box="[271,289,1975,1994]" pageId="1" pageNumber="148">H</emphasis>
|
||||
). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="A4F866330D48FF85FFF6770B8254A1D6" pageId="2" pageNumber="149" startId="2.[102,135,1613,1632]" targetBox="[102,1485,227,1583]" targetPageId="2" targetType="figure">
|
||||
<paragraph id="F03836BB0D48FF85FFF6770B8254A1D6" blockId="2.[102,1485,1613,1744]" pageId="2" pageNumber="149">
|
||||
Fig. 3.
|
||||
<taxonomicName id="37874D380D48FF85FF36770B8035A166" authorityName="Valiukevičius" authorityYear="2004" box="[166,349,1613,1632]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="2" pageNumber="149" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D48FF85FF36770B8035A166" box="[166,349,1613,1632]" italics="true" pageId="2" pageNumber="149">Vesperalia perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D48FF85FEF3770B8085A166" box="[355,493,1613,1632]" pageId="2" pageNumber="149" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
Microstructure of scales in vertical longitudinal (
|
||||
<emphasis id="C2F3EAA90D48FF85FC3D770B82D5A166" bold="true" box="[941,957,1613,1632]" pageId="2" pageNumber="149">A</emphasis>
|
||||
) and transverse (
|
||||
<emphasis id="C2F3EAA90D48FF85FBCA770B8501A166" bold="true" box="[1114,1129,1613,1632]" pageId="2" pageNumber="149">B</emphasis>
|
||||
) sections. A
|
||||
<subScript id="6C0334FE0D48FF85FB4B7710858BA162" attach="left" box="[1243,1251,1622,1636]" fontSize="6" pageId="2" pageNumber="149">1</subScript>
|
||||
. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A
|
||||
<subScript id="6C0334FE0D48FF85FF3E77C881DEA19A" attach="right" box="[174,182,1678,1692]" fontSize="6" pageId="2" pageNumber="149">2</subScript>
|
||||
. Detail of posterior edge of crown and a small portion of base of the same scale. B
|
||||
<subScript id="6C0334FE0D48FF85FC0077C882F0A19A" attach="right" box="[912,920,1678,1692]" fontSize="6" pageId="2" pageNumber="149">1</subScript>
|
||||
. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B
|
||||
<subScript id="6C0334FE0D48FF85FEBB77EC805BA1BE" attach="left" box="[299,307,1706,1720]" fontSize="6" pageId="2" pageNumber="149">2</subScript>
|
||||
. Detail of the leftpart of the same scale. B
|
||||
<subScript id="6C0334FE0D48FF85FD3877EC83D8A1BE" attach="left" box="[680,688,1706,1720]" fontSize="6" pageId="2" pageNumber="149">3</subScript>
|
||||
. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="F03836BB0D48FF84FCC276D483B9A626" blockId="2.[812,1486,1807,2026]" lastBlockId="3.[102,775,231,2026]" lastPageId="3" lastPageNumber="150" pageId="2" pageNumber="149">
|
||||
Scale necks are tall and contain large linearly opening basal pores on all the walls, with up to four on each side (
|
||||
<figureCitation id="68BC2A3E0D48FF85FADE76F484A9A0CC" box="[1358,1473,1970,1995]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="2" pageNumber="149">Fig. 2A, E</figureCitation>
|
||||
). The scale base is rhombic in outline, moderately to greatly convex, usually protruding slightly beyond the crown antero−laterally. The deepest point of the base is centrally positioned.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FF1B70608088A55C" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
|
||||
Putative headscales(veryrareinthestudiedsamples)have isometric rhomboid, thickened crowns (up to
|
||||
<quantity id="377F9B5E0D49FF84FDF57000826FA658" box="[613,775,326,351]" metricMagnitude="-4" metricUnit="m" metricValue="7.35" metricValueMax="8.2" metricValueMin="6.5" pageId="3" pageNumber="150" unit="mm" value="0.735" valueMax="0.82" valueMin="0.65">0.65–0.82 mm</quantity>
|
||||
long and wide) and low porous necks as compared to trunk scales. Head scale crowns have three to four prominent, robust and rounded ridges of even height extending over the entire crown length. These robust ridges bend medially towards the posterior edge (
|
||||
<figureCitation id="68BC2A3E0D49FF84FE9E70A58008A6FA" box="[270,352,483,508]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 2F</figureCitation>
|
||||
), but do not join each other. Additional lateral oblique riblets occur on the side of each ridge. The postero−lateral crown edge is multicuspid with several (up to four on each side) rough spinelets.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FF1B73278363A492" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
|
||||
The histology of all
|
||||
<taxonomicName id="37874D380D49FF84FEF97324808FA57C" authorityName="Valiukevičius" authorityYear="2004" box="[361,487,609,634]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="3" pageNumber="150" phylum="Chordata" rank="species" species="perplexa">
|
||||
<emphasis id="C2F3EAA90D49FF84FEF97324808FA57C" box="[361,487,609,634]" italics="true" pageId="3" pageNumber="150">V. perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
scales (
|
||||
<figureCitation id="68BC2A3E0D49FF84FDAE73278317A57C" box="[574,639,609,634]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3</figureCitation>
|
||||
) is uniform. Crowns are composed of stranggewebe (mesodentinal tissue containing elongate, linearly oriented lacunae and incorporating osteocytes; for histologic terminology see
|
||||
<bibRefCitation id="94164B4A0D49FF84FDCC738683B1A5DE" author="Gross, W." box="[604,729,704,728]" pageId="3" pageNumber="150" pagination="1 - 82" refId="ref9073" refString="Gross, W. 1971. Downtonische und Dittonische Acanthodier-Reste des Ostseegebietes. Palaeontographica A 136: 1 - 82." type="journal article" year="1971">Gross 1971</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation id="94164B4A0D49FF84FFF673988197A5F1" author="Denison, R." box="[102,255,734,759]" pageId="3" pageNumber="150" pagination="1 - 62" refId="ref9033" refString="Denison, R. 1979. Acanthodii. In: H. - P. Schultze (ed.), Handbook of Paleoichthyology, 5. VI + 1 - 62 pp. Gustav Fischer Verlag, Stuttgart." type="book chapter" year="1979">Denison 1979</bibRefCitation>
|
||||
) and simple mesodentine arranged in four thick growth lamellae. The growth
|
||||
<typeStatus id="2F3C88190D49FF84FE5873B9809FA411" box="[456,503,767,791]" pageId="3" pageNumber="150">type</typeStatus>
|
||||
differs slightly among scales. Scales with interrupted ridges (
|
||||
<figureCitation id="68BC2A3E0D49FF84FD85725B8303A430" box="[533,619,797,822]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3A</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D49FF84FDFC726C83C8A43D" box="[620,672,798,827]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1−Stoniškiai, 7−Krekenava, 8−Stačiūnai, 11−Šešuvis, 12−Kunkojai, 44−Nida, 61−Viduklė, 87−Sutkai, 99−Gėluva, 128−Vilkaviškis, 137−Liepkalnis, 162−Kurtuvėnai, 179−Ledai, 241−Butkūnai, 252−Svėdasai." pageId="3" pageNumber="150">
|
||||
<subScript id="6C0334FE0D49FF84FDFC726C831DA43D" attach="right" box="[620,629,810,827]" fontSize="7" pageId="3" pageNumber="150">1</subScript>
|
||||
, B
|
||||
<subScript id="6C0334FE0D49FF84FD07726C83C8A43D" attach="left" box="[663,672,810,827]" fontSize="7" pageId="3" pageNumber="150">1</subScript>
|
||||
</figureCitation>
|
||||
) demonstrate a superpositional growth in only the two oldest lamellae, whereas the two youngest are added areally. Scales with continuous ridges grow superpositionally.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FF1B72DD83BDA376" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
|
||||
Thestranggewebeischaracterizedbyenlargedandelongate stranglacunae with interconnecting short processes. An unevenly thick mantle of simple mesodentine is superficial to the stranggewebe in each growth lamella (
|
||||
<figureCitation id="68BC2A3E0D49FF84FD8172BF830EA314" box="[529,614,1017,1042]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3B</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D49FF84FDF575408306A311" box="[613,622,1030,1047]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">
|
||||
<subScript id="6C0334FE0D49FF84FDF575408306A311" attach="left" box="[613,622,1030,1047]" fontSize="7" pageId="3" pageNumber="150">2</subScript>
|
||||
</figureCitation>
|
||||
). In scales of this species, the stranggewebe develops with a similar intensity not only in the posterior part of the crown, as it is characteristic of most climatiids, but also in its anterior parts (
|
||||
<figureCitation id="68BC2A3E0D49FF84FDF9751E83D4A376" box="[617,700,1112,1137]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3A</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D49FF84FD2C752283ADA373" box="[700,709,1124,1141]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1−Stoniškiai, 7−Krekenava, 8−Stačiūnai, 11−Šešuvis, 12−Kunkojai, 44−Nida, 61−Viduklė, 87−Sutkai, 99−Gėluva, 128−Vilkaviškis, 137−Liepkalnis, 162−Kurtuvėnai, 179−Ledai, 241−Butkūnai, 252−Svėdasai." pageId="3" pageNumber="150">
|
||||
<subScript id="6C0334FE0D49FF84FD2C752283ADA373" attach="left" box="[700,709,1124,1141]" fontSize="7" pageId="3" pageNumber="150">1</subScript>
|
||||
</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FF1B75318268A22B" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
|
||||
The basal part of the neck has rare, typical mesodentine osteocyte cavities. The principal vascular system is formed of enlarged radial vascular canals positioned over the base, and wide ascending and slightly branched main canals supplying each growth lamella (
|
||||
<figureCitation id="68BC2A3E0D49FF84FEDC75B380F6A208" box="[332,414,1269,1294]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3A</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D49FF84FE0E744480CFA215" box="[414,423,1282,1299]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">
|
||||
<subScript id="6C0334FE0D49FF84FE0E744480CFA215" attach="left" box="[414,423,1282,1299]" fontSize="7" pageId="3" pageNumber="150">2</subScript>
|
||||
</figureCitation>
|
||||
, B
|
||||
<figureCitation id="68BC2A3E0D49FF84FE56744480A7A215" box="[454,463,1282,1299]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">
|
||||
<subScript id="6C0334FE0D49FF84FE56744480A7A215" attach="left" box="[454,463,1282,1299]" fontSize="7" pageId="3" pageNumber="150">3</subScript>
|
||||
</figureCitation>
|
||||
). This system is complicated intheearliest/oldestlamellaebecauseofadditionalbranchings.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FF1B747283C6A2AD" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
|
||||
The simple mesodentine enclosing stranggewebe in the outer parts of lamellae contains a characteristic net of winding dentine tubules with many osteocyte cavities even in the superficial layers (
|
||||
<figureCitation id="68BC2A3E0D49FF84FE8874D48006A2AD" box="[280,366,1426,1451]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig. 3A</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D49FF84FEFD74D980C8A2B6" box="[365,416,1428,1456]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">
|
||||
<subScript id="6C0334FE0D49FF84FEFD74D9801EA2B6" attach="left" box="[365,374,1439,1456]" fontSize="7" pageId="3" pageNumber="150">2</subScript>
|
||||
, B
|
||||
<subScript id="6C0334FE0D49FF84FE0774D980C8A2B6" attach="right" box="[407,416,1439,1456]" fontSize="7" pageId="3" pageNumber="150">2</subScript>
|
||||
</figureCitation>
|
||||
). Durodentine is absent.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FF1B74F48393A16E" blockId="3.[102,775,231,2026]" pageId="3" pageNumber="150">
|
||||
Scale bases are composed of cellular bone arranged in very thin and dense growth lamellae with plenty of osteocyte cavities and medium to long Sharpey’s fibre traces. The osteocyte density increases towards the tip of pyramidal base, where they become oriented along the growth lines, and approximate thestructureofbothbonetissueandstranggewebe(
|
||||
<figureCitation id="68BC2A3E0D49FF84FD097709838AA16E" box="[665,738,1615,1640]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">Fig.3B</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D49FF84FD71771A8382A16B" box="[737,746,1628,1645]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="3" pageNumber="150">
|
||||
<subScript id="6C0334FE0D49FF84FD71771A8382A16B" attach="left" box="[737,746,1628,1645]" fontSize="7" pageId="3" pageNumber="150">2</subScript>
|
||||
</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FFF6773F8429A639" blockId="3.[102,775,231,2026]" lastBlockId="3.[812,1486,231,1641]" pageId="3" pageNumber="150">
|
||||
<emphasis id="C2F3EAA90D49FF84FFF6773F81B6A196" box="[102,222,1657,1680]" italics="true" pageId="3" pageNumber="150">Discussion</emphasis>
|
||||
.—The trunk scales of
|
||||
<taxonomicName id="37874D380D49FF84FE43773E8322A197" authorityName="Valiukevicius" authorityYear="2004" box="[467,586,1656,1681]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="3" pageNumber="150" phylum="Chordata" rank="genus">
|
||||
<emphasis id="C2F3EAA90D49FF84FE43773E8322A197" box="[467,586,1656,1681]" italics="true" pageId="3" pageNumber="150">Vesperalia</emphasis>
|
||||
</taxonomicName>
|
||||
differ from those of all known climatiids in having a high and linearly porous neckdistinct fromtheconvexbasebyasharprim andirregular ridges ornamenting the horizontal plate of the crown. They also have wide, pronounced, often wavy subparallel ridges; the two central ridges may form an elevated medial area and point towards the posterior end of the crown. Ridges of some scale specimens aredeveloped along the anterior and posterior edges of the crown, leaving the median part of the crown unsculptured a character not previously observed in acanthodians. Histological microstructure corresponds to the
|
||||
<emphasis id="C2F3EAA90D49FF84FD2D76F581CBA0EC" italics="true" pageId="3" pageNumber="150">Nostolepis−</emphasis>
|
||||
<typeStatus id="2F3C88190D49FF84FF33769481BDA0EC" box="[163,213,2002,2026]" pageId="3" pageNumber="150">type</typeStatus>
|
||||
, distinguished by very large ascending and radial vascular canals, unusually elongate lacunae of stranggewebe covered by a mantle of osteocyte−rich mesodentine in scale crowns (the tissue has bone−like characteristics).
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FCC270018420A493" blockId="3.[812,1486,231,1641]" pageId="3" pageNumber="150">
|
||||
The
|
||||
<emphasis id="C2F3EAA90D49FF84FC1B7001856DA666" box="[907,1029,327,352]" italics="true" pageId="3" pageNumber="150">Nostolepis−</emphasis>
|
||||
<typeStatus id="2F3C88190D49FF84FB96700E855CA666" box="[1030,1076,328,352]" pageId="3" pageNumber="150">type</typeStatus>
|
||||
histology suggests that
|
||||
<taxonomicName id="37874D380D49FF84FAC9700182E2A679" authorityName="Valiukevičius" authorityYear="2004" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="3" pageNumber="150" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D49FF84FAC9700182E2A679" italics="true" pageId="3" pageNumber="150">Vesperalia perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D49FF84FC04702E8529A679" box="[916,1089,360,384]" pageId="3" pageNumber="150" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
can be asigned to the climatiids as presently classified. Based on morphologic characters, particularly the distribution of crown ridges, the scales of
|
||||
<taxonomicName id="37874D380D49FF84FAC170E184A5A6B8" authorityName="Valiukevičius" authorityYear="2004" box="[1361,1485,421,446]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="3" pageNumber="150" phylum="Chordata" rank="species" species="perplexa">
|
||||
<emphasis id="C2F3EAA90D49FF84FAC170E184A5A6B8" box="[1361,1485,421,446]" italics="true" pageId="3" pageNumber="150">V. perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
differfromknownnostolepids,andshouldbeconsideredgenerically distinct. Other morphological features, that are worth mentioning are the high and porous necks (not characteristic of most climatiids), and the crown sculpture of interrupted ridges. In addition, both superpositional and areal growth
|
||||
<typeStatus id="2F3C88190D49FF84FAD573058416A55D" box="[1349,1406,579,603]" pageId="3" pageNumber="150">types</typeStatus>
|
||||
are developed in scales of
|
||||
<taxonomicName id="37874D380D49FF84FB96732585EAA57D" authorityName="Valiukevičius" authorityYear="2004" box="[1030,1154,610,635]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="3" pageNumber="150" phylum="Chordata" rank="species" species="perplexa">
|
||||
<emphasis id="C2F3EAA90D49FF84FB96732585EAA57D" box="[1030,1154,610,635]" italics="true" pageId="3" pageNumber="150">V. perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
. It is obvious, that the replacement of the superpositional growth by the areal one takes always place after the formation of two the oldest lamellae (of four), and can not be attached to the ontogenetic development stage(youngerorolder)ofscales.Histologically,scalesaredistinguished from those of other
|
||||
<taxonomicName id="37874D380D49FF84FB1173B9844CA41E" authorityName="Berg" authorityYear="1940" box="[1153,1316,767,792]" class="Acanthodii" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="3" pageNumber="150" phylum="Chordata" rank="order">Climatiiformes</taxonomicName>
|
||||
not only by an exhuberant development of stranggewebe, a unique character among climatiids, but also by the particularly large oriented lacunae, clearly separated strips of stranggewebe and thick mantle of simple mesodentine in growth lamellae.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FCC272DB8587A16F" blockId="3.[812,1486,231,1641]" pageId="3" pageNumber="150">
|
||||
There are no closely related climatiids to be compared. According to several characters,
|
||||
<taxonomicName id="37874D380D49FF84FBE772FB843DA4D0" authorityName="Valiukevičius" authorityYear="2004" box="[1143,1365,957,982]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="3" pageNumber="150" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D49FF84FBE772FB843DA4D0" box="[1143,1365,957,982]" italics="true" pageId="3" pageNumber="150">Vesperalia perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D49FF84FACC72F88235A4F2" pageId="3" pageNumber="150" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
resembles representatives of
|
||||
<taxonomicName id="37874D380D49FF84FB3D729A8453A4F3" authorityName="Valiukevicius" authorityYear="1998" box="[1197,1339,988,1013]" genus="Endemolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="150" phylum="Chordata" rank="genus">
|
||||
<emphasis id="C2F3EAA90D49FF84FB3D729A8453A4F3" box="[1197,1339,988,1013]" italics="true" pageId="3" pageNumber="150">Endemolepis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<emphasis id="C2F3EAA90D49FF84FAE6729B82E9A312" italics="true" pageId="3" pageNumber="150">Tareyacanthus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="94164B4A0D49FF84FC0072BD8503A312" author="Valiukevicius, J." box="[912,1131,1019,1044]" pageId="3" pageNumber="150" pagination="236 - 243" refId="ref9291" refString="Valiukevicius, J. 1994. Acanthodiansandtheirstratigraphicsignificance [in Russian]. In: S. Cherkesova, V. Karatajute- Talimaa, and R. Matukhin (eds.), Stratigrafia i fauna niznedevonskih otlozenij Tareiskogo opornogo razreza, 131 - 197 and 236 - 243. Vserossijskij naucno-issledovatel'skij institut geologii i mineral'nyh ressursov Mirovogo okeana, Sankt Petersburg." type="book chapter" year="1994">Valiukevičius 1994</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="94164B4A0D49FF84FBE672BA85D9A312" author="Valiukevicius, J." box="[1142,1201,1020,1044]" pageId="3" pageNumber="150" pagination="1 - 53" refId="ref9353" refString="Valiukevicius, J. 1998. Acanthodians and zonal stratigraphy of Lower and Middle Devonian in East Baltic and Byelorussia. Palaeontographica A 248: 1 - 53." type="journal article" year="1998">1998</bibRefCitation>
|
||||
). The former, represented by a single species
|
||||
<taxonomicName id="37874D380D49FF84FB83755D820DA355" authority="Valiukevicius, 1998" authorityName="Valiukevicius" authorityYear="1998" genus="Endemolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="150" phylum="Chordata" rank="species" species="inconstans">
|
||||
<emphasis id="C2F3EAA90D49FF84FB83755D8449A335" box="[1043,1313,1051,1076]" italics="true" pageId="3" pageNumber="150">Endemolepis inconstans</emphasis>
|
||||
<bibRefCitation id="94164B4A0D49FF84FABE755D820DA355" author="Valiukevicius, J." pageId="3" pageNumber="150" pagination="1 - 53" refId="ref9353" refString="Valiukevicius, J. 1998. Acanthodians and zonal stratigraphy of Lower and Middle Devonian in East Baltic and Byelorussia. Palaeontographica A 248: 1 - 53." type="journal article" year="1998">Valiukevičius, 1998</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, ranges across the Silurian–Devonian boundary in the Baltic, is distinguished by crown ridges of changeable form and length (sharp narrow, broad rounded, single or meeting in pairs, frequently with additional oblique branchings) extending the entire crown length, except for a diagnostic, narrow, unornamented anterior strip (
|
||||
<bibRefCitation id="94164B4A0D49FF84FBFC75918421A3F6" author="Valiukevicius, J." box="[1132,1353,1239,1264]" pageId="3" pageNumber="150" pagination="1 - 53" refId="ref9353" refString="Valiukevicius, J. 1998. Acanthodians and zonal stratigraphy of Lower and Middle Devonian in East Baltic and Byelorussia. Palaeontographica A 248: 1 - 53." type="journal article" year="1998">Valiukevičius 1998</bibRefCitation>
|
||||
: pl. 6: 3–7). Two known Devonian representatives of
|
||||
<emphasis id="C2F3EAA90D49FF84FA9275B084CDA209" box="[1282,1445,1270,1295]" italics="true" pageId="3" pageNumber="150">Tareyacanthus</emphasis>
|
||||
,
|
||||
<emphasis id="C2F3EAA90D49FF84FA2775BE82CCA229" italics="true" pageId="3" pageNumber="150">T. magnificus</emphasis>
|
||||
<bibRefCitation id="94164B4A0D49FF84FC3E745085E6A229" author="Valiukevicius, J." box="[942,1166,1302,1327]" pageId="3" pageNumber="150" pagination="236 - 243" refId="ref9291" refString="Valiukevicius, J. 1994. Acanthodiansandtheirstratigraphicsignificance [in Russian]. In: S. Cherkesova, V. Karatajute- Talimaa, and R. Matukhin (eds.), Stratigrafia i fauna niznedevonskih otlozenij Tareiskogo opornogo razreza, 131 - 197 and 236 - 243. Vserossijskij naucno-issledovatel'skij institut geologii i mineral'nyh ressursov Mirovogo okeana, Sankt Petersburg." type="book chapter" year="1994">Valiukevičius, 1994</bibRefCitation>
|
||||
(pl. 23: 9–14) from Taimyr Peninsula (
|
||||
<collectingCountry id="8890762B0D49FF84FC3474738298A248" box="[932,1008,1333,1358]" name="Russia" pageId="3" pageNumber="150">Russia</collectingCountry>
|
||||
) and
|
||||
<emphasis id="C2F3EAA90D49FF84FBB9747185CFA248" box="[1065,1191,1333,1358]" italics="true" pageId="3" pageNumber="150">T. dissectus</emphasis>
|
||||
<bibRefCitation id="94164B4A0D49FF84FB3D747384E0A248" author="Valiukevicius, J." box="[1197,1416,1333,1358]" pageId="3" pageNumber="150" pagination="1 - 53" refId="ref9353" refString="Valiukevicius, J. 1998. Acanthodians and zonal stratigraphy of Lower and Middle Devonian in East Baltic and Byelorussia. Palaeontographica A 248: 1 - 53." type="journal article" year="1998">Valiukevičius, 1998</bibRefCitation>
|
||||
(pl. 3: 22–24 and pl. 4: 7–9) from the Stoniškiai Formation of the Baltic coast, have scales ornamented with high, sharp parallel or slightly fan−shaped ridges (up to eight per scale), that may bifurcate anteriorly and are separated by deep grooves. The histologic microstructure of both taxa is only comparable with
|
||||
<taxonomicName id="37874D380D49FF84FCBC74B48561A10D" authorityName="Valiukevičius" authorityYear="2004" box="[812,1033,1522,1547]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="3" pageNumber="150" phylum="Chordata" rank="species" species="perplexa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D49FF84FCBC74B48561A10D" box="[812,1033,1522,1547]" italics="true" pageId="3" pageNumber="150">Vesperalia perplexa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D49FF84FB9F74B585DBA10C" box="[1039,1203,1523,1547]" pageId="3" pageNumber="150" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
as they all have a complicated system of principal vascular canals in crowns, but the details mentioned previously distinguish
|
||||
<taxonomicName id="37874D380D49FF84FB6F7776841EA14F" authorityName="Valiukevicius" authorityYear="2004" box="[1279,1398,1584,1609]" class="Acanthodii" family="Climatiidae" genus="Vesperalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="3" pageNumber="150" phylum="Chordata" rank="genus">
|
||||
<emphasis id="C2F3EAA90D49FF84FB6F7776841EA14F" box="[1279,1398,1584,1609]" italics="true" pageId="3" pageNumber="150">Vesperalia</emphasis>
|
||||
</taxonomicName>
|
||||
and its single species from all known climatiids.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D49FF84FCBC773A841DA1B5" blockId="3.[812,1485,1659,1716]" pageId="3" pageNumber="150">
|
||||
<emphasis id="C2F3EAA90D49FF84FCBC773A82C3A195" box="[812,939,1660,1683]" italics="true" pageId="3" pageNumber="150">Occurrence</emphasis>
|
||||
.—Stoniškiai−1 borehole, depth 1211.0–1217.0 m; Nida−44: 1213.0 m and Šešuvis−11: 1005.0–
|
||||
<quantity id="377F9B5E0D49FF84FA9977DD8419A1B5" box="[1289,1393,1691,1716]" metricMagnitude="3" metricUnit="m" metricValue="1.0152" pageId="3" pageNumber="150" unit="m" value="1015.2">1015.2 m</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
312
data/78/2E/87/782E87AD0D4EFF81FFF671AE83ECA5B9.xml
Normal file
312
data/78/2E/87/782E87AD0D4EFF81FFF671AE83ECA5B9.xml
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|
|
@ -0,0 +1,312 @@
|
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<mods:titleInfo id="1674D0059F29F734A46E389B8B386BDA">
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<mods:title id="AD43E928065D32C83241ED67AB739F8B">New Wenlock-Pridoli (Silurian) acanthodian fishes from Lithuania</mods:title>
|
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</mods:titleInfo>
|
||||
<mods:name id="D83D720D221230A94E3532907918DEDB" type="personal">
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<mods:roleTerm id="B0081D95FE17F34A8B1721E1FF7B50B6">Author</mods:roleTerm>
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</mods:role>
|
||||
<mods:namePart id="E538E4E2F166645B88581448A0C5975A">Valiukevičius, Juozas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4541E10343FC7872C04640FCA475228A">text</mods:typeOfResource>
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||||
<mods:title id="6774B80414EEA82297A6FFDF18CFC4B2">Acta Palaeontologica Polonica</mods:title>
|
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</mods:titleInfo>
|
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<mods:part id="94EFEF891C2258F733B06823AE7F502C">
|
||||
<mods:date id="6ADCBD126E9BEC5C219D88D344351183">2004</mods:date>
|
||||
<mods:detail id="521800C3940A69820CED0FC2B53FEA76" type="pubDate">
|
||||
<mods:number id="B3800B6D5E85110B21064A7B896836F5">2004-12-31</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="14B9646D0F4ED48E77DFDFCB9C6F1ACB" type="volume">
|
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<mods:number id="99A47CA04D4631B30E7ED12DA173F5FC">49</mods:number>
|
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</mods:detail>
|
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<mods:detail id="FA0C4F12575E60BA6E76FBC62514CBDC" type="issue">
|
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<mods:number id="947BB025384D209128EBD4E38056271C">1</mods:number>
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<mods:extent id="C838944F444ECCD894CFEA28A5884B51" unit="page">
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<mods:classification id="6739D4694DA23F1B4518FA8AC89A543C">journal article</mods:classification>
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<mods:identifier id="988566F393DF656556009C4EF7A46B4D" type="ISSN">1732-2421</mods:identifier>
|
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</mods:mods>
|
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<treatment id="782E87AD0D4EFF81FFF671AE83ECA5B9" LSID="urn:lsid:plazi:treatment:782E87AD0D4EFF81FFF671AE83ECA5B9" httpUri="http://treatment.plazi.org/id/782E87AD0D4EFF81FFF671AE83ECA5B9" lastPageId="6" lastPageNumber="153" pageId="4" pageNumber="151">
|
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<subSubSection id="B89D65300D4EFF83FFF671AE83CEA603" box="[102,678,232,261]" pageId="4" pageNumber="151" type="nomenclature">
|
||||
<paragraph id="F03836BB0D4EFF83FFF671AE83CEA603" blockId="4.[102,678,232,296]" box="[102,678,232,261]" pageId="4" pageNumber="151">
|
||||
<heading id="AB7081D70D4EFF83FFF671AE83CEA603" box="[102,678,232,261]" fontSize="12" level="2" pageId="4" pageNumber="151" reason="9">
|
||||
<taxonomicName id="37874D380D4EFF83FFF671AE80B4A603" authorityName="Valiukevičius" authorityYear="2004" box="[102,476,232,261]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF671AE80B4A603" box="[102,476,232,261]" italics="true" pageId="4" pageNumber="151">Fecundosquama basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D4EFF83FE7471AF83CEA603" box="[484,678,233,261]" pageId="4" pageNumber="151" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="B89D65300D4EFF81FFF6705483ECA5B9" lastPageId="6" lastPageNumber="153" pageId="4" pageNumber="151" type="description">
|
||||
<paragraph id="F03836BB0D4EFF83FFF6705481ADA62E" blockId="4.[102,678,232,296]" box="[102,197,274,296]" pageId="4" pageNumber="151">
|
||||
<figureCitation id="68BC2A3E0D4EFF83FFF6705481C1A62E" box="[102,169,274,296]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25−A−2405, anterolateral view, anterior to the left. B. LIGG 25−A−2407, crown view, anterior upwards. E. LIGG 25−A−2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25−A−2409. D. LIGG 25−A−2408. Scale bars 0.1 mm. Gėluva−99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Figs. 4</figureCitation>
|
||||
,
|
||||
<figureCitation id="68BC2A3E0D4EFF83FF23705481A9A62E" box="[179,193,274,296]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1–B3) and Ledai−179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">5</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FFF6707D8345A657" blockId="4.[102,775,315,603]" box="[102,557,315,337]" pageId="4" pageNumber="151">
|
||||
<taxonomicName id="37874D380D4EFF83FFF6707D8043A657" box="[102,299,315,337]" class="Acanthodii" family="Ischnacanthidae" genus="Gomphonchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="undefined-A">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF6707D8199A657" box="[102,241,315,337]" italics="true" pageId="4" pageNumber="151">Gomphonchus</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="94164B4A0D4EFF83FEA5707D80C2A657" author="Marss, T." box="[309,426,315,337]" pageId="4" pageNumber="151" pagination="17 - 41" refId="ref9224" refString="Marss, T. 1997. Vertebrates of the Pridoli and Silurian - Devonian boundary beds in Europe. Modern Geology 21: 17 - 41." type="journal article" year="1997">Märss 1997</bibRefCitation>
|
||||
: pl. 2: 13–15.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FFF670198325A673" blockId="4.[102,775,315,603]" box="[102,589,351,373]" pageId="4" pageNumber="151">
|
||||
<materialsCitation id="40EF3CE60D4EFF83FFF670198325A673" box="[102,589,351,373]" collectionCode="LIGG" pageId="4" pageNumber="151" specimenCode="LIGG 25" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF6701981D6A673" box="[102,190,351,373]" italics="true" pageId="4" pageNumber="151">
|
||||
<typeStatus id="2F3C88190D4EFF83FFF6701981D6A673" box="[102,190,351,373]" pageId="4" pageNumber="151" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCode id="A0219EC00D4EFF83FF5A70198076A673" box="[202,286,351,373]" collectionCode="LIGG" pageId="4" pageNumber="151">LIGG 25</specimenCode>
|
||||
−A−2405, trunk scale (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FE647019832AA673" box="[500,578,351,373]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25−A−2405, anterolateral view, anterior to the left. B. LIGG 25−A−2407, crown view, anterior upwards. E. LIGG 25−A−2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25−A−2409. D. LIGG 25−A−2408. Scale bars 0.1 mm. Gėluva−99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4A</figureCitation>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FFF670C283BFA69C" blockId="4.[102,775,315,603]" box="[102,727,388,410]" pageId="4" pageNumber="151">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF670C2818AA69C" box="[102,226,388,410]" italics="true" pageId="4" pageNumber="151">Type locality</emphasis>
|
||||
: Kurtuvėnai−162 borehole, depth
|
||||
<quantity id="377F9B5E0D4EFF83FDB870C283BBA69C" box="[552,723,388,410]" metricMagnitude="3" metricUnit="m" metricValue="1.0623" metricValueMax="1.0724" metricValueMin="1.0522" pageId="4" pageNumber="151" unit="m" value="1062.3" valueMax="1072.4" valueMin="1052.2">1052.2–1072.4 m</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FFF670EF80EFA6F0" blockId="4.[102,775,315,603]" pageId="4" pageNumber="151">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF670EF8189A6B8" box="[102,225,424,446]" italics="true" pageId="4" pageNumber="151">Type horizon</emphasis>
|
||||
:TheupperpartofMinijaRegionalStage(VarniaiBedsor theircorrelatives)tothelowerpartofJūraRegionalStage(lowerpartof the Jūra or Lapės formations).
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FFF67343839FA51D" blockId="4.[102,775,315,603]" box="[102,759,517,539]" pageId="4" pageNumber="151">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF6734381CBA51C" box="[102,163,517,538]" italics="true" pageId="4" pageNumber="151">Range</emphasis>
|
||||
:MinijaRegionalStagetothelowerpartofJūraRegionalStage.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FFF6736C809EA55D" blockId="4.[102,775,315,603]" pageId="4" pageNumber="151">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF6736C8043A539" box="[102,299,553,575]" italics="true" pageId="4" pageNumber="151">Derivation of name</emphasis>
|
||||
: From Latin
|
||||
<emphasis id="C2F3EAA90D4EFF83FE2B736F8084A539" box="[443,492,553,575]" italics="true" pageId="4" pageNumber="151">basis</emphasis>
|
||||
, base and
|
||||
<emphasis id="C2F3EAA90D4EFF83FDF2736F83DFA539" box="[610,695,553,575]" italics="true" pageId="4" pageNumber="151">globosus</emphasis>
|
||||
, round, globose, referring to the scale base shape.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FFF673298094A58E" blockId="4.[102,508,623,648]" box="[102,508,623,648]" pageId="4" pageNumber="151">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF6732981AEA58E" box="[102,198,623,648]" italics="true" pageId="4" pageNumber="151">Material</emphasis>
|
||||
.—More than 50 000 scales.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FFF673DB83B1A496" blockId="4.[102,775,668,912]" pageId="4" pageNumber="151">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF673DB81BDA5B2" box="[102,213,669,692]" italics="true" pageId="4" pageNumber="151">Diagnosis</emphasis>
|
||||
.—
|
||||
<taxonomicName id="37874D380D4EFF83FF6773DA80C5A5B3" authorityName="Valiukevicius" authorityYear="2004" box="[247,429,668,693]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="genus">
|
||||
<emphasis id="C2F3EAA90D4EFF83FF6773DA80C5A5B3" box="[247,429,668,693]" italics="true" pageId="4" pageNumber="151">Fecundosquama</emphasis>
|
||||
</taxonomicName>
|
||||
scales having low horizontal crowns with crenulated sculpture anteriorly or around the entire crown margin, lacking a neck; deep convex base present. Scale crownscomposed of simple bone−like mesodentine with abundant osteocyte cavities and enlarged vascular canals present in the oldest growth lamellae only. Crown mesodentine merges gradually into highly cellular thin−lamellar bone in the base that is penetrated by long traces of Sharpey’s fibres.
|
||||
</paragraph>
|
||||
<caption id="A4F866330D4EFF83FCBC73A18506A4A4" pageId="4" pageNumber="151" startId="4.[812,845,743,762]" targetBox="[813,1484,240,691]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="F03836BB0D4EFF83FCBC73A18506A4A4" blockId="4.[812,1485,743,930]" pageId="4" pageNumber="151">
|
||||
Fig. 4.
|
||||
<taxonomicName id="37874D380D4EFF83FCFE73A18510A5FC" authorityName="Valiukevičius" authorityYear="2004" box="[878,1144,743,762]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D4EFF83FCFE73A18510A5FC" box="[878,1144,743,762]" italics="true" pageId="4" pageNumber="151">Fecundosquama basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D4EFF83FBED73A18461A5FC" box="[1149,1289,743,762]" pageId="4" pageNumber="151" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
SEM micrographs of scales.
|
||||
<emphasis id="C2F3EAA90D4EFF83FCFE72458216A410" bold="true" box="[878,894,771,790]" pageId="4" pageNumber="151">A</emphasis>
|
||||
,
|
||||
<emphasis id="C2F3EAA90D4EFF83FC1A724582F1A410" bold="true" box="[906,921,771,790]" pageId="4" pageNumber="151">B</emphasis>
|
||||
, and
|
||||
<emphasis id="C2F3EAA90D4EFF83FC5C724582B3A410" bold="true" box="[972,987,771,790]" pageId="4" pageNumber="151">E</emphasis>
|
||||
. Trunk scales.
|
||||
<emphasis id="C2F3EAA90D4EFF83FBF57245851DA410" bold="true" box="[1125,1141,771,790]" pageId="4" pageNumber="151">A</emphasis>
|
||||
. Holotype, LIGG 25−A−2405, anterolateral view, anterior to the left.
|
||||
<emphasis id="C2F3EAA90D4EFF83FBDD72598534A434" bold="true" box="[1101,1116,799,818]" pageId="4" pageNumber="151">B</emphasis>
|
||||
. LIGG 25−A−2407, crown view, anterior upwards.
|
||||
<emphasis id="C2F3EAA90D4EFF83FC13727D82FAA448" bold="true" box="[899,914,827,846]" pageId="4" pageNumber="151">E</emphasis>
|
||||
. LIGG 25−A−2406, lateralview, anteriorto the left.
|
||||
<emphasis id="C2F3EAA90D4EFF83FACE727D8406A448" bold="true" box="[1374,1390,827,846]" pageId="4" pageNumber="151">C</emphasis>
|
||||
,
|
||||
<emphasis id="C2F3EAA90D4EFF83FAE8727D84E0A448" bold="true" box="[1400,1416,827,846]" pageId="4" pageNumber="151">D</emphasis>
|
||||
. Head? scales, crown views, anterior upwards.
|
||||
<emphasis id="C2F3EAA90D4EFF83FB34721185DCA46C" bold="true" box="[1188,1204,855,874]" pageId="4" pageNumber="151">C</emphasis>
|
||||
. LIGG 25−A−2409.
|
||||
<emphasis id="C2F3EAA90D4EFF83FAE7721184EFA46C" bold="true" box="[1399,1415,855,874]" pageId="4" pageNumber="151">D</emphasis>
|
||||
. LIGG 25−A−2408. Scale bars 0.1 mm. Gėluva−99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="F03836BB0D4EFF83FFF672E383ECA2B5" blockId="4.[102,775,932,2026]" pageId="4" pageNumber="151">
|
||||
<emphasis id="C2F3EAA90D4EFF83FFF672E38180A4BA" box="[102,232,933,956]" italics="true" pageId="4" pageNumber="151">Description</emphasis>
|
||||
.—Scales have a very low crown and a large, convex base protruding beyond the crown on all sides. Crown length varies from
|
||||
<quantity id="377F9B5E0D4EFF83FEA472A580B4A4FD" box="[308,476,995,1020]" metricMagnitude="-4" metricUnit="m" metricValue="5.249999999999999" metricValueMax="7.0" metricValueMin="3.5" pageId="4" pageNumber="151" unit="mm" value="0.5249999999999999" valueMax="0.7" valueMin="0.35">0.35 to 0.7 mm</quantity>
|
||||
, and can be
|
||||
<quantity id="377F9B5E0D4EFF83FDF172A5826FA4FD" box="[609,775,995,1019]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="7.7" metricValueMin="3.0" pageId="4" pageNumber="151" unit="mm" value="0.535" valueMax="0.77" valueMin="0.3">0.3 to 0.77 mm</quantity>
|
||||
wide. The isometric rhomboid crown plate of trunk scales (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FFFE75678068A33C" box="[110,256,1057,1082]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25−A−2405, anterolateral view, anterior to the left. B. LIGG 25−A−2407, crown view, anterior upwards. E. LIGG 25−A−2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25−A−2409. D. LIGG 25−A−2408. Scale bars 0.1 mm. Gėluva−99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4A, B, E</figureCitation>
|
||||
) is almost flat, very slightly curved downwards at the anterior margins, and posteriorly the crowns are elongated and rhomboidal to elipsoidal. The anterior margin of the crown has short but comparatively wide notches which extend to the surface of the scale base. The number of notches varies from 5 to 6 (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FF7D75F88057A3D1" box="[237,319,1214,1239]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25−A−2405, anterolateral view, anterior to the left. B. LIGG 25−A−2407, crown view, anterior upwards. E. LIGG 25−A−2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25−A−2409. D. LIGG 25−A−2408. Scale bars 0.1 mm. Gėluva−99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4E</figureCitation>
|
||||
) to 10 (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FE0375F88081A3D1" box="[403,489,1214,1239]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25−A−2405, anterolateral view, anterior to the left. B. LIGG 25−A−2407, crown view, anterior upwards. E. LIGG 25−A−2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25−A−2409. D. LIGG 25−A−2408. Scale bars 0.1 mm. Gėluva−99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4B</figureCitation>
|
||||
). The putative head scales (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FFFE7598818FA3F0" box="[110,231,1246,1271]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25−A−2405, anterolateral view, anterior to the left. B. LIGG 25−A−2407, crown view, anterior upwards. E. LIGG 25−A−2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25−A−2409. D. LIGG 25−A−2408. Scale bars 0.1 mm. Gėluva−99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4C, D</figureCitation>
|
||||
) differ from the trunk scales in having isometric rhomboid to round crowns with notches on all margins. The deep,convexscale baseis rhomboid to round in outline, with a prominent lateral rim at the deepest point projecting anterior to the crown. In rare cases the deepest point of the base is situated posteriorly, so the base protrudes beyond the posterior margin of the crown (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FEC474DC80C1A2B5" box="[340,425,1434,1459]" captionStart="Fig" captionStartId="4.[812,845,743,762]" captionTargetBox="[813,1484,240,691]" captionTargetId="figure-840@4.[812,1485,237,693]" captionTargetPageId="4" captionText="Fig. 4. Fecundosquama basiglobosa gen. et sp. nov. SEM micrographs of scales. A, B, and E. Trunk scales. A. Holotype, LIGG 25−A−2405, anterolateral view, anterior to the left. B. LIGG 25−A−2407, crown view, anterior upwards. E. LIGG 25−A−2406, lateralview, anteriorto the left. C, D. Head? scales, crown views, anterior upwards. C. LIGG 25−A−2409. D. LIGG 25−A−2408. Scale bars 0.1 mm. Gėluva−99 borehole, depth 681.6 m. Upper Silurian, Pridoli, Vievis Formation." pageId="4" pageNumber="151">Fig. 4E</figureCitation>
|
||||
). Scales lack necks.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FF1B74FD83D1A0EC" blockId="4.[102,775,932,2026]" pageId="4" pageNumber="151">
|
||||
Simple mesodentine crown tissue (neither stranggewebe nor durodentine present) and cellular bone in scale bases can be seen in the sectioned scales. Up to six thin lamellae of superpositional growth in crowns are composed of bone−like mesodentine containing numerous multi−angular osteocyte spaces even in the latest, outer lamella (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FD82771E830EA177" box="[530,614,1624,1649]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1–B3) and Ledai−179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5A</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D4EFF83FDF677238307A170" box="[614,623,1637,1654]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151">
|
||||
<subScript id="6C0334FE0D4EFF83FDF677238307A170" attach="left" box="[614,623,1637,1654]" fontSize="7" pageId="4" pageNumber="151">2</subScript>
|
||||
</figureCitation>
|
||||
) and short radiating and winding dentine tubules form a fine network. The scale primordium does not differ from others in respect to tissue structure. Most scales sectioned have no enlarged vascular canal system in crowns (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FEFF779080ABA1E9" box="[367,451,1750,1775]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1–B3) and Ledai−179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5A</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D4EFF83FE5377A5809CA1F2" box="[451,500,1751,1780]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1−Stoniškiai, 7−Krekenava, 8−Stačiūnai, 11−Šešuvis, 12−Kunkojai, 44−Nida, 61−Viduklė, 87−Sutkai, 99−Gėluva, 128−Vilkaviškis, 137−Liepkalnis, 162−Kurtuvėnai, 179−Ledai, 241−Butkūnai, 252−Svėdasai." pageId="4" pageNumber="151">
|
||||
<subScript id="6C0334FE0D4EFF83FE5377A580A4A1F2" attach="left" box="[451,460,1763,1780]" fontSize="7" pageId="4" pageNumber="151">1</subScript>
|
||||
, B
|
||||
<subScript id="6C0334FE0D4EFF83FE7B77A5809CA1F2" attach="left" box="[491,500,1763,1780]" fontSize="7" pageId="4" pageNumber="151">1</subScript>
|
||||
</figureCitation>
|
||||
), and only one (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FD347790839EA1E9" box="[676,758,1750,1775]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1–B3) and Ledai−179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5C</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D4EFF83FD6677A58397A1F2" box="[758,767,1763,1780]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1−Stoniškiai, 7−Krekenava, 8−Stačiūnai, 11−Šešuvis, 12−Kunkojai, 44−Nida, 61−Viduklė, 87−Sutkai, 99−Gėluva, 128−Vilkaviškis, 137−Liepkalnis, 162−Kurtuvėnai, 179−Ledai, 241−Butkūnai, 252−Svėdasai." pageId="4" pageNumber="151">
|
||||
<subScript id="6C0334FE0D4EFF83FD6677A58397A1F2" attach="left" box="[758,767,1763,1780]" fontSize="7" pageId="4" pageNumber="151">1</subScript>
|
||||
</figureCitation>
|
||||
, C
|
||||
<figureCitation id="68BC2A3E0D4EFF83FFE8764481E9A015" box="[120,129,1794,1811]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151">
|
||||
<subScript id="6C0334FE0D4EFF83FFE8764481E9A015" attach="left" box="[120,129,1794,1811]" fontSize="7" pageId="4" pageNumber="151">2</subScript>
|
||||
</figureCitation>
|
||||
) demonstrates wide radial vascular canals in the two oldest growth lamellae. There is no sharp boundary between the crown mesodentine and bone of the base: the first tissue grades gradually into the second. The cellular bone of the base is composed of numerous very fine growth zones (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FD307635839EA08A" box="[672,758,1907,1932]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1–B3) and Ledai−179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5A</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D4EFF83FD6676C68397A097" box="[758,767,1920,1937]" captionStart="Fig" captionStartId="1.[102,135,838,857]" captionTargetBox="[102,773,222,811]" captionTargetId="graphics-280@1.[102,773,222,812]" captionTargetPageId="1" captionText="Fig. 1. Sketch map showing location of the boreholes yielding the majority of the material studied. Borehole and its original number: 1−Stoniškiai, 7−Krekenava, 8−Stačiūnai, 11−Šešuvis, 12−Kunkojai, 44−Nida, 61−Viduklė, 87−Sutkai, 99−Gėluva, 128−Vilkaviškis, 137−Liepkalnis, 162−Kurtuvėnai, 179−Ledai, 241−Butkūnai, 252−Svėdasai." pageId="4" pageNumber="151">
|
||||
<subScript id="6C0334FE0D4EFF83FD6676C68397A097" attach="left" box="[758,767,1920,1937]" fontSize="7" pageId="4" pageNumber="151">1</subScript>
|
||||
</figureCitation>
|
||||
, A
|
||||
<figureCitation id="68BC2A3E0D4EFF83FFE976D981EAA0B6" box="[121,130,1951,1968]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151">
|
||||
<subScript id="6C0334FE0D4EFF83FFE976D981EAA0B6" attach="left" box="[121,130,1951,1968]" fontSize="7" pageId="4" pageNumber="151">2</subScript>
|
||||
</figureCitation>
|
||||
) penetrated by long Sharpey’s fibre traces (
|
||||
<figureCitation id="68BC2A3E0D4EFF83FDE376D483A1A0AD" box="[627,713,1938,1963]" captionStart="Fig" captionStartId="5.[102,135,1892,1911]" captionTargetBox="[201,1386,225,1866]" captionTargetId="figure-8@5.[201,1387,274,1864]" captionTargetPageId="5" captionText="Fig. 5. Fecundosquama basiglobosa gen. et sp. nov. Microstructure of trunk scales in vertical transverse (A) and longitudinal (B, C) sections. A1. Thin section LIGG 3683. A2. Detail of the left part of the same scale. B1. Thin section LIGG 3686. B2. Detail of the right part of the base of the same scale. B3. Detail of the central part of crown and bordering base area of the same scale. C1. General view, thin section LIGG 3687. C2. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A1, A2 and B1–B3) and Ledai−179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation." pageId="4" pageNumber="151">Fig. 5B</figureCitation>
|
||||
<figureCitation id="68BC2A3E0D4EFF83FD5976D983BAA0B6" box="[713,722,1951,1968]" captionStart="Fig" captionStartId="1.[102,135,1919,1938]" captionTargetBox="[110,1473,1208,1887]" captionTargetId="figure-398@1.[105,1482,1203,1891]" captionTargetPageId="1" captionText="Fig. 2. Vesperalia perplexa gen. et sp. nov. SEM micrographs, trunk scales except for F, head? scale, crown views, anterior upwards: holotype, LIGG 25−A−2413 (A), LIGG 25−A−2416 (B), LIGG 25−A−2417 (C), LIGG 25−A−2418 (D), LIGG 25−A−2420 (E), LIGG 25−A−2422 (F), LIGG 25−A−2421 (G), LIGG 25−A−2414(H). Scalebars0.1 mm.Nida−44borehole, depth1213.0 m(A, E–H) andStoniškiai−1borehole, depth1211.0–1217.0 m(B–D). Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151">
|
||||
<subScript id="6C0334FE0D4EFF83FD5976D983BAA0B6" attach="left" box="[713,722,1951,1968]" fontSize="7" pageId="4" pageNumber="151">2</subScript>
|
||||
</figureCitation>
|
||||
, B
|
||||
<figureCitation id="68BC2A3E0D4EFF83FD6476D98395A0B6" box="[756,765,1951,1968]" captionStart="Fig" captionStartId="2.[102,135,1613,1632]" captionTargetBox="[102,1485,227,1583]" captionTargetId="figure-157@2.[102,1486,295,1583]" captionTargetPageId="2" captionText="Fig. 3. Vesperalia perplexa gen. et sp. nov. Microstructure of scales in vertical longitudinal (A) and transverse (B) sections. A1. Holotype−like scale with uniformbent, sub−parallelridgesprominentattheanteriorand posterioredges ofthecrown, and almostfadingoutnear thecrown center. ThinsectionLIGG 3678. A2. Detail of posterior edge of crown and a small portion of base of the same scale. B1. Widened scale with flattened ridges like those in Fig. 2E. Thin sectionLIGG 3679. B2. Detail of the leftpart of the same scale. B3. Detail of the rightpart of the same scale. Scale bars 0.1 mm. Stoniškiai−1borehole, depth 1211.0–1217.0 m. Upper Silurian, Pridoli, Rietavas Beds of the Jūra Formation." pageId="4" pageNumber="151">
|
||||
<subScript id="6C0334FE0D4EFF83FD6476D98395A0B6" attach="left" box="[756,765,1951,1968]" fontSize="7" pageId="4" pageNumber="151">3</subScript>
|
||||
</figureCitation>
|
||||
) and enclosed abundant osteocyte cavities with short cell processes. Osteocytes are oriented along growth lines.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FCBC72818448A23E" blockId="4.[812,1486,965,2026]" pageId="4" pageNumber="151">
|
||||
<emphasis id="C2F3EAA90D4EFF83FCBC728182C8A4D8" box="[812,928,967,990]" italics="true" pageId="4" pageNumber="151">Discussion</emphasis>
|
||||
.—This new genus and species is based on scales with clear morphologic (center of flat crown without ridge ornamentation, fine ridges confined to crown margins, no neck, deep convex base protruding beyond crown on all sides) and histologic characters (neither stranggewebe nor durodentine present in crowns, vascular canal system weakly developed, well−developed net of bone−like mesodentine with many osteocyte spaces, forming a gradual transition to cellular bone in scale bases). The comparatively short stratigraphic range of
|
||||
<taxonomicName id="37874D380D4EFF83FA8D758682C5A3FF" authorityName="Valiukevičius" authorityYear="2004" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D4EFF83FA8D758682C5A3FF" italics="true" pageId="4" pageNumber="151">Fecundosquama basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D4EFF83FC2675A78509A3FE" box="[950,1121,1249,1273]" pageId="4" pageNumber="151" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
together with diagnostic features makes it a potentially good biostratigraphic taxon for more detailed vertebrate zonations of the Late Silurian.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF83FCC27479846BA1D6" blockId="4.[812,1486,965,2026]" pageId="4" pageNumber="151">
|
||||
<taxonomicName id="37874D380D4EFF83FCC27479829EA251" authorityName="Valiukevičius" authorityYear="2004" box="[850,1014,1342,1367]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D4EFF83FCC27479829EA251" box="[850,1014,1342,1367]" italics="true" pageId="4" pageNumber="151">F. basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D4EFF83FC6C747985CBA250" box="[1020,1187,1343,1367]" pageId="4" pageNumber="151" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
has outstanding diagnostic features, and no related taxa are known to date. In terms of histology, there are some known climatiids similar
|
||||
<taxonomicName id="37874D380D4EFF83FAE0743882EBA2B3" authorityName="Valiukevičius" authorityYear="2004" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="basiglobosa">
|
||||
<emphasis id="C2F3EAA90D4EFF83FAE0743882EBA2B3" italics="true" pageId="4" pageNumber="151">F. basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
in some ways, for example, bone−like mesodentine, absence of stranggewebe or reduced principal vascular canal system. The presence of such scale characters was demonstrated in a study on phylogenetical development of nostolepids from the Timan−Pechora region spanning the Late Silurian–Early Devonian (
|
||||
<bibRefCitation id="94164B4A0D4EFF83FB8F777F8593A154" author="Valiukevicius, J." box="[1055,1275,1593,1618]" pageId="4" pageNumber="151" pagination="134 - 136" refId="ref9382" refString="Valiukevicius, J. 2000. PhylogeneticallineagesofEarlyDevoniannostolepids of Timan-Pechora region. In: A. Antoshkina, E. Malysheva, and M. V. H. Wilson (eds.), Pan-Arctic Tectonics, Evolution of Basins and Faunas. Ichthyolith Issues, Special Publication 6: 134 - 136. Syktyvkar." type="journal article" year="2000">Valiukevičius 2000</bibRefCitation>
|
||||
). All the characters together in
|
||||
<taxonomicName id="37874D380D4EFF83FC3B771C8527A177" authorityName="Valiukevičius" authorityYear="2004" box="[939,1103,1624,1649]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="basiglobosa">
|
||||
<emphasis id="C2F3EAA90D4EFF83FC3B771C8527A177" box="[939,1103,1624,1649]" italics="true" pageId="4" pageNumber="151">F. basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
, especially the distinctive features of bone tissue in scale bases (density and fine−lamellar cellular bone and gradual transition to the crown mesodentine), are observed in acanthodians for the first time.
|
||||
</paragraph>
|
||||
<paragraph id="F03836BB0D4EFF81FCC2779183EDA51D" blockId="4.[812,1486,965,2026]" lastBlockId="6.[102,775,231,704]" lastPageId="6" lastPageNumber="153" pageId="4" pageNumber="151">
|
||||
<bibRefCitation id="94164B4A0D4EFF83FCC277918284A1E9" author="Marss, T." box="[850,1004,1751,1775]" pageId="4" pageNumber="151" pagination="17 - 41" refId="ref9224" refString="Marss, T. 1997. Vertebrates of the Pridoli and Silurian - Devonian boundary beds in Europe. Modern Geology 21: 17 - 41." type="journal article" year="1997">Märss (1997)</bibRefCitation>
|
||||
presented morphologically similar specimens (histology not examined) from the same region of the Baltic, in the Dubovskoye borehole of the
|
||||
<collectingRegion id="3243F8590D4EFF83FB62765384A4A028" box="[1266,1484,1813,1838]" country="Russia" name="Kaliningrad" pageId="4" pageNumber="151">Kaliningrad District</collectingRegion>
|
||||
(former East Prussia). The rocks are from the Kaugatuma Stage, Pridoli, from depth
|
||||
<quantity id="377F9B5E0D4EFF83FBF07612844CA06A" box="[1120,1316,1876,1901]" metricMagnitude="3" metricUnit="m" metricValue="1.0751" metricValueMax="1.0855" metricValueMin="1.0647" pageId="4" pageNumber="151" unit="m" value="1075.1" valueMax="1085.5" valueMin="1064.7">1064.7–1085.5 m</quantity>
|
||||
and originally were identified as
|
||||
<taxonomicName id="37874D380D4EFF83FC6B763585B6A08A" box="[1019,1246,1907,1932]" class="Acanthodii" family="Ischnacanthidae" genus="Gomphonchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="undefined-A">
|
||||
<emphasis id="C2F3EAA90D4EFF83FC6B763585F2A08A" box="[1019,1178,1907,1932]" italics="true" pageId="4" pageNumber="151">Gomphonchus</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="94164B4A0D4EFF83FB7D7632841DA08A" author="Marss, T." box="[1261,1397,1908,1932]" pageId="4" pageNumber="151" pagination="17 - 41" refId="ref9224" refString="Marss, T. 1997. Vertebrates of the Pridoli and Silurian - Devonian boundary beds in Europe. Modern Geology 21: 17 - 41." type="journal article" year="1997">Märss 1997</bibRefCitation>
|
||||
: fig. 4). These scales are referred to
|
||||
<taxonomicName id="37874D380D4EFF83FBFE76D2847CA0AD" authorityName="Valiukevičius" authorityYear="2004" box="[1134,1300,1938,1963]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="151" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D4EFF83FBFE76D2847CA0AD" box="[1134,1300,1938,1963]" italics="true" pageId="4" pageNumber="151">F. basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D4EFF83FA8E76D584A5A0AC" box="[1310,1485,1939,1963]" pageId="4" pageNumber="151" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
Some characters (low horizontal crown, no neck, large convex base and proportions of crown/base development) demonstrate similarity between these scales, however, the scales are not identical. The Dubovskoye specimens (
|
||||
<bibRefCitation id="94164B4A0D4CFF81FDAE704183ACA619" author="Marss, T." box="[574,708,263,287]" pageId="6" pageNumber="153" pagination="17 - 41" refId="ref9224" refString="Marss, T. 1997. Vertebrates of the Pridoli and Silurian - Devonian boundary beds in Europe. Modern Geology 21: 17 - 41." type="journal article" year="1997">Märss 1997</bibRefCitation>
|
||||
: pl. 2: 13–15)not only havenotches onthe anterior part ofthe crown, but also have short surface ridgelets. The main difference between
|
||||
<taxonomicName id="37874D380D4CFF81FF3F7020803BA678" authorityName="Valiukevičius" authorityYear="2004" box="[175,339,357,382]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="6" pageNumber="153" phylum="Chordata" rank="species" species="basiglobosa">
|
||||
<emphasis id="C2F3EAA90D4CFF81FF3F7020803BA678" box="[175,339,357,382]" italics="true" pageId="6" pageNumber="153">F. basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
and the Dubovskoye scales is the presenceofsmallbulbsorsometimes cone−likespineletsdisplaced in lines on the postero−lateral crown walls in the latter. These specimens, in my opinion, are similar to and might belong to
|
||||
<taxonomicName id="37874D380D4CFF81FFF670A28079A6FD" authorityName="Valiukevičius" authorityYear="2004" box="[102,273,482,507]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="6" pageNumber="153" phylum="Chordata" rank="species" species="basiglobosa">
|
||||
<emphasis id="C2F3EAA90D4CFF81FFF670A28079A6FD" box="[102,273,482,507]" italics="true" pageId="6" pageNumber="153">F. basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
, what suggests a climatiid rather than an ischnacanthid affinity for the Dubovskoye scales.
|
||||
</paragraph>
|
||||
<caption id="A4F866330D4FFF82FFF6762282F2A0E1" pageId="5" pageNumber="152" startId="5.[102,135,1892,1911]" targetBox="[201,1386,225,1866]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="F03836BB0D4FFF82FFF6762282F2A0E1" blockId="5.[102,1485,1892,2027]" pageId="5" pageNumber="152">
|
||||
Fig. 5.
|
||||
<taxonomicName id="37874D380D4FFF82FF34762280C4A071" authorityName="Valiukevičius" authorityYear="2004" box="[164,428,1892,1911]" class="Acanthodii" family="Climatiidae" genus="Fecundosquama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="5" pageNumber="152" phylum="Chordata" rank="species" species="basiglobosa" status="gen. et sp. nov.">
|
||||
<emphasis id="C2F3EAA90D4FFF82FF34762280C4A071" box="[164,428,1892,1911]" italics="true" pageId="5" pageNumber="152">Fecundosquama basiglobosa</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C057D20D4FFF82FE207623835FA07E" box="[432,567,1893,1912]" pageId="5" pageNumber="152" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
Microstructure of trunk scales in vertical transverse (
|
||||
<emphasis id="C2F3EAA90D4FFF82FB827622854AA071" bold="true" box="[1042,1058,1892,1911]" pageId="5" pageNumber="152">A</emphasis>
|
||||
) and longitudinal (
|
||||
<emphasis id="C2F3EAA90D4FFF82FB5D762385B4A07E" bold="true" box="[1229,1244,1893,1912]" pageId="5" pageNumber="152">B</emphasis>
|
||||
,
|
||||
<emphasis id="C2F3EAA90D4FFF82FB777622859FA071" bold="true" box="[1255,1271,1892,1911]" pageId="5" pageNumber="152">C</emphasis>
|
||||
) sections. A
|
||||
<subScript id="6C0334FE0D4FFF82FAF7762B8407A07D" attach="right" box="[1383,1391,1901,1915]" fontSize="6" pageId="5" pageNumber="152">1</subScript>
|
||||
. Thin section LIGG 3683. A
|
||||
<subScript id="6C0334FE0D4FFF82FE9F76CF807FA091" attach="left" box="[271,279,1929,1943]" fontSize="6" pageId="5" pageNumber="152">2</subScript>
|
||||
. Detail of the left part of the same scale. B
|
||||
<subScript id="6C0334FE0D4FFF82FD1E76CF83FEA091" attach="left" box="[654,662,1929,1943]" fontSize="6" pageId="5" pageNumber="152">1</subScript>
|
||||
. Thin section LIGG 3686. B
|
||||
<subScript id="6C0334FE0D4FFF82FC0576CF82F5A091" attach="left" box="[917,925,1929,1943]" fontSize="6" pageId="5" pageNumber="152">2</subScript>
|
||||
. Detail of the right part of the base of the same scale. B
|
||||
<subScript id="6C0334FE0D4FFF82FA1476CF84E4A091" attach="right" box="[1412,1420,1929,1943]" fontSize="6" pageId="5" pageNumber="152">3</subScript>
|
||||
. Detail of the central part of crown and bordering base area of the same scale. C
|
||||
<subScript id="6C0334FE0D4FFF82FD7576E38385A0B5" attach="left" box="[741,749,1957,1971]" fontSize="6" pageId="5" pageNumber="152">1</subScript>
|
||||
. General view, thin section LIGG 3687. C
|
||||
<subScript id="6C0334FE0D4FFF82FBF876E38518A0B5" attach="left" box="[1128,1136,1957,1971]" fontSize="6" pageId="5" pageNumber="152">2</subScript>
|
||||
. The same scale, detail of the right part of crown and bordering base area. Scale bars 0.1 mm. Kurtuvėnai−162 borehole, depth 1052.2 m. Upper Silurian, Pridoli, Jūra Formation (A
|
||||
<subScript id="6C0334FE0D4FFF82FAE8768784E8A0C9" attach="right" box="[1400,1408,1985,1999]" fontSize="6" pageId="5" pageNumber="152">1</subScript>
|
||||
, A
|
||||
<subScript id="6C0334FE0D4FFF82FA0D768784CDA0C9" attach="left" box="[1437,1445,1985,1999]" fontSize="6" pageId="5" pageNumber="152">2</subScript>
|
||||
and B
|
||||
<subScript id="6C0334FE0D4FFF82FFE5769B8115A0ED" attach="left" box="[117,125,2013,2027]" fontSize="6" pageId="5" pageNumber="152">1</subScript>
|
||||
–B
|
||||
<subScript id="6C0334FE0D4FFF82FF07769B81F7A0ED" attach="left" box="[151,159,2013,2027]" fontSize="6" pageId="5" pageNumber="152">3</subScript>
|
||||
) and Ledai−179 borehole, depth 537.7 m. Upper Silurian, Pridoli, Lapės Formation.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="F03836BB0D4CFF81FFF6736D83ECA5B9" blockId="6.[102,775,231,704]" pageId="6" pageNumber="153">
|
||||
<emphasis id="C2F3EAA90D4CFF81FFF6736D818FA544" box="[102,231,555,578]" italics="true" pageId="6" pageNumber="153">Occurrence</emphasis>
|
||||
.—Ledai−179 borehole, depth
|
||||
<quantity id="377F9B5E0D4CFF81FDDF736C8269A544" box="[591,769,554,579]" metricMagnitude="2" metricUnit="m" metricValue="5.425" metricValueMax="5.476" metricValueMin="5.374" pageId="6" pageNumber="153" unit="m" value="542.5" valueMax="547.6" valueMin="537.4">537.4–547.6 m</quantity>
|
||||
; Gėluva−99:
|
||||
<quantity id="377F9B5E0D4CFF81FF75730F80E4A564" box="[229,396,585,610]" metricMagnitude="2" metricUnit="m" metricValue="6.8345" metricValueMax="6.853" metricValueMin="6.816" pageId="6" pageNumber="153" unit="m" value="683.45" valueMax="685.3" valueMin="681.6">681.6–685.3m</quantity>
|
||||
;Sutkai−87:
|
||||
<quantity id="377F9B5E0D4CFF81FD9D730F83C7A564" box="[525,687,585,610]" metricMagnitude="2" metricUnit="m" metricValue="6.122999999999999" metricValueMax="6.153" metricValueMin="6.092999999999999" pageId="6" pageNumber="153" unit="m" value="612.3" valueMax="615.3" valueMin="609.3">609.3–615.3m</quantity>
|
||||
;Kurtuvėnai−162:
|
||||
<quantity id="377F9B5E0D4CFF81FF76732E80C6A586" box="[230,430,616,641]" metricMagnitude="3" metricUnit="m" metricValue="1.0623" metricValueMax="1.0724" metricValueMin="1.0522" pageId="6" pageNumber="153" unit="m" value="1062.3" valueMax="1072.4" valueMin="1052.2">1052.2–1072.4 m</quantity>
|
||||
; Stoniškiai−1: 1311.0– 1359.0 m; Šešuvis−11:
|
||||
<quantity id="377F9B5E0D4CFF81FEDD73CE8346A5A6" box="[333,558,648,673]" metricMagnitude="3" metricUnit="m" metricValue="1.0962" metricValueMax="1.1195" metricValueMin="1.0729" pageId="6" pageNumber="153" unit="m" value="1096.2" valueMax="1119.5" valueMin="1072.9">1072.9–1119.5 m</quantity>
|
||||
; Liepkalnis−137:
|
||||
<quantity id="377F9B5E0D4CFF81FFF673EE8066A5B9" box="[102,270,680,704]" metricMagnitude="2" metricUnit="m" metricValue="8.93" metricValueMax="8.939" metricValueMin="8.921" pageId="6" pageNumber="153" unit="m" value="893.0" valueMax="893.9" valueMin="892.1">892.1–893.9 m</quantity>
|
||||
; Vilkaviškis−128: 706.2–714.0 m.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue