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<document id="9F20E98FB3A2CBFD606B131873943FE2" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C40CE27FE89C6D39FC8FD80" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Takedaplax Ng, Ahyong &amp; T., 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="348" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="347" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EEE6ABBB0C7FDFB8F62ACD6D4A5AE0F9">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
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<mods:namePart id="8A775F6E5652383D4A07D9BCE2B7F877">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="050E9025A1BBE17694D882CC9F0D8F64">Ahyong,</mods:namePart>
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<mods:namePart id="0CBFC432AD5244B209A0DAED00538A8C">Shane T.</mods:namePart>
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<mods:title id="866883D7F4D160B943258CD62C2EE2DB">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="21D48AAF2722F1B7E80C11D76ECE14F7">2024</mods:date>
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<subSubSection id="DDB00AC92C40CE26FE89C6D39C14F903" box="[323,587,1757,1781]" pageId="12" pageNumber="347" type="nomenclature">
<paragraph id="951559422C40CE26FE89C6D39C14F903" blockId="12.[323,587,1757,1781]" box="[323,587,1757,1781]" pageId="12" pageNumber="347">
<heading id="CE5DEE2E2C40CE26FE89C6D39C14F903" bold="true" box="[323,587,1757,1781]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="347" reason="2">
<emphasis id="A7DE85502C40CE26FE89C6D39C14F903" bold="true" box="[323,587,1757,1781]" pageId="12" pageNumber="347">
<taxonomicName id="52AA22C12C40CE26FE89C6D39F9BF903" ID-CoL="CL3BY" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[323,452,1757,1781]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C40CE26FE89C6D39F9BF903" bold="true" box="[323,452,1757,1781]" italics="true" pageId="12" pageNumber="347">Takedaplax</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C40CE26FE1EC6D39C14F903" box="[468,587,1757,1781]" pageId="12" pageNumber="347" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="DDB00AC92C40CE26FF42C7139F65F843" pageId="12" pageNumber="347" type="type_taxon">
<paragraph id="951559422C40CE26FF42C7139CACF8A3" blockId="12.[136,775,1821,1878]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FF42C7139F40F8C3" bold="true" box="[136,287,1821,1845]" pageId="12" pageNumber="347">
<typeStatus id="4A11E7E02C40CE26FF42C7139E9DF8C3" box="[136,194,1821,1845]" pageId="12" pageNumber="347">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="52AA22C12C40CE26FEECC7139E9EF8A3" ID-CoL="6QMFQ" authority="Takeda &amp; Kurata, 1984" authorityName="Takeda &amp; Kurata" authorityYear="1984" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C40CE26FEECC7139C62F8C3" box="[294,573,1821,1845]" italics="true" pageId="12" pageNumber="347">Lophoplax sextuberculata</emphasis>
<bibRefCitation id="F13B24B32C40CE26FD8FC7139E9EF8A3" author="Takeda M &amp; Kurata Y" pageId="12" pageNumber="347" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
</taxonomicName>
, by present designation. Gender of genus feminine.
</paragraph>
<paragraph id="951559422C40CE26FF42C7739F65F843" blockId="12.[136,774,1917,1973]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FF42C7739F13F863" bold="true" box="[136,332,1917,1941]" pageId="12" pageNumber="347">Included species.</emphasis>
Only
<taxonomicName id="52AA22C12C40CE26FE5FC7739F69F843" ID-CoL="6QMFQ" authority="Takeda &amp; Kurata, 1984" authorityName="Takeda &amp; Kurata" authorityYear="1984" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C40CE26FE5FC7739CF1F863" box="[405,686,1917,1941]" italics="true" pageId="12" pageNumber="347">Lophoplax sextuberculata</emphasis>
<bibRefCitation id="F13B24B32C40CE26FD72C7739F69F843" author="Takeda M &amp; Kurata Y" pageId="12" pageNumber="347" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C40CE26FF42C7D39B36FBA3" pageId="12" pageNumber="347" type="diagnosis">
<paragraph id="951559422C40CE26FF42C7D39B36FBA3" blockId="12.[136,775,2013,2069]" lastBlockId="12.[835,1475,189,1109]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FF42C7D39EA2F803" bold="true" box="[136,253,2013,2037]" pageId="12" pageNumber="347">Diagnosis.</emphasis>
Carapace rectangular; epigastric, postorbital and hepatic areoles well developed with anterior margin cristate to subcristate; epigastric region gently convex, smooth, setose, not areolate; protogastric, cardiac, mesobranchial and cardiobranchial regions may be raised to form areole in larger specimens, barely or not visible in smaller specimens; intestinal region not prominently raised, no visible intestino-branchial areole; surface adjacent to posterior carapace margin almost flat, not forming ridge, no transverse channel visible; orbit clearly visible in dorsal view, supraorbital margin breaking general carapace outline, forming distinct concavity; eyes freely movable, ocular peduncle relatively long; lateral lobe of frontal margin low, indistinct; anterolateral margin with 4 visible teeth or spines (including external orbital tooth), last one often small; posterolateral margins sinuous, gently converging towards posterior carapace margin. Endostome not obliquely recessed posteriorly into buccal cavern, subparallel to adjacent pterygostomial surface, low, short endostomial ridges. Posterior margin of epistome entire, without lateral fissures, median part triangular, lateral parts distinctly concave. Third maxilliped ischium subquadrate, length about 1.3× width. Adult male chelipeds almost symmetrical; dorsal surface of carpus with prominent longitudinal areole, separated from swollen lateral fringe by groove; chela relatively slender with external surface covered with tubercles and granules. Ambulatory legs elongate, slender, P4 longest, about 2.0× carapace width; P25 merus without crest on extensor margin, distally unarmed, outer surface smooth, without medio-longitudinal swelling; P4 merus length about 4.3× height, distinctly longer than half carapace width. G1 strongly sinuous, tip recurved.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C40CE26FC89C4739A44FAA3" pageId="12" pageNumber="347" type="etymology">
<paragraph id="951559422C40CE26FC89C4739A44FAA3" blockId="12.[835,1474,1149,1365]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FC89C4739D9AFB63" bold="true" box="[835,965,1149,1173]" pageId="12" pageNumber="347">Etymology.</emphasis>
The genus is named after Masatsune Takeda, whose work on the genus has been invaluable in the present revision, and we thank him for encouraging us to “finish the job” by revising the generic position of all species previously placed in
<taxonomicName id="52AA22C12C40CE26FBEDC4F39AC6FAE3" authorityName="Tesch" authorityYear="1918" box="[1063,1177,1277,1301]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FBEDC4F39AC6FAE3" box="[1063,1177,1277,1301]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
. The name is derived from his family name, in arbitrary combination with
<taxonomicName id="52AA22C12C40CE26FA83C5139BE4FAC3" authorityName="Tesch" authorityYear="1918" box="[1353,1467,1309,1333]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FA83C5139BE4FAC3" box="[1353,1467,1309,1333]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
. Gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C40CE27FC89C5739FC8FD80" lastPageId="13" lastPageNumber="348" pageId="12" pageNumber="347" type="discussion">
<paragraph id="951559422C40CE26FC89C5739A55F803" blockId="12.[835,1475,1405,2038]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FC89C5739DEDFA63" bold="true" box="[835,946,1405,1429]" pageId="12" pageNumber="347">Remarks.</emphasis>
The six prominent areolets on the anterior half of the carapace of
<taxonomicName id="52AA22C12C40CE26FBD6C5939B61FA43" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[1052,1342,1437,1461]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C40CE26FBD6C5939B61FA43" box="[1052,1342,1437,1461]" italics="true" pageId="12" pageNumber="347">Lophoplax sextuberculata</emphasis>
</taxonomicName>
is the only character that links the species to
<taxonomicName id="52AA22C12C40CE26FB73C5B39B74FA23" authorityName="Tesch" authorityYear="1918" box="[1209,1323,1469,1493]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FB73C5B39B74FA23" box="[1209,1323,1469,1493]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
, although the position of the areolets are different. Compared to
<taxonomicName id="52AA22C12C40CE26FA85C5D39B9FFA03" authorityName="Tesch" authorityYear="1918" box="[1359,1472,1501,1525]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="A7DE85502C40CE26FA85C5D39B9FFA03" box="[1359,1472,1501,1525]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCED382B2C40CE26FC89C5F09D8EF9E3" box="[835,977,1533,1557]" pageId="12" pageNumber="347" sensu="stricto">sensu stricto</taxonomicNameLabel>
, however,
<taxonomicName id="52AA22C12C40CE26FB84C5F09B51F9E3" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[1102,1294,1533,1557]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C40CE26FB84C5F09B51F9E3" box="[1102,1294,1533,1557]" italics="true" pageId="12" pageNumber="347">L. sextuberculata</emphasis>
</taxonomicName>
has a distinctly wider carapace which appears rectangular in form, in sharp contrast to the more quadrate form. Most markedly, it differs in having the posterior margin of the epistome entire, without any lateral fissures (
<figureCitation id="0D9145C72C40CE26FBFAC6739ACBF963" box="[1072,1172,1661,1685]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="12" pageNumber="347">Figs. 8E</figureCitation>
,
<figureCitation id="0D9145C72C40CE26FB6EC6739A96F963" box="[1188,1225,1661,1685]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="12" pageNumber="347">9D</figureCitation>
). In
<taxonomicName id="52AA22C12C40CE26FAC9C6739B25F963" authorityName="Tesch" authorityYear="1918" box="[1283,1402,1661,1685]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="A7DE85502C40CE26FAC9C6739B25F963" box="[1283,1402,1661,1685]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCED382B2C40CE26FA4EC6709DD7F943" pageId="12" pageNumber="347" sensu="stricto">sensu stricto</taxonomicNameLabel>
, as well as
<taxonomicName id="52AA22C12C40CE26FBC4C6939AC0F943" authorityName="Deb" authorityYear="1989" box="[1038,1183,1693,1717]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FBC4C6939AC0F943" box="[1038,1183,1693,1717]" italics="true" pageId="12" pageNumber="347">Myopilumnus</emphasis>
</taxonomicName>
,
<taxonomicName id="52AA22C12C40CE26FB65C6939B59F943" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[1199,1286,1693,1717]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C40CE26FB65C6939B59F943" box="[1199,1286,1693,1717]" italics="true" pageId="12" pageNumber="347">Hosekia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C40CE26FADDC6909BD1F940" box="[1303,1422,1694,1718]" pageId="12" pageNumber="347" rank="genus">new genus</taxonomicNameLabel>
, and
<taxonomicName id="52AA22C12C40CE26FC89C6B39D8DF923" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[835,978,1725,1749]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C40CE26FC89C6B39D8DF923" box="[835,978,1725,1749]" italics="true" pageId="12" pageNumber="347">Striacoeloma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C40CE26FC29C6B09A05F920" box="[995,1114,1726,1750]" pageId="12" pageNumber="347" rank="genus">new genus</taxonomicNameLabel>
, the posterior epistomial margin is divided into three distinct lobes by well-defined fissures (e.g.,
<figureCitation id="0D9145C72C40CE26FC49C6F39DBDF8E3" box="[899,994,1789,1813]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="12" pageNumber="347">Figs. 3B</figureCitation>
,
<figureCitation id="0D9145C72C40CE26FC3AC6F39A4FF8E3" box="[1008,1040,1789,1813]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="12" pageNumber="347">4E</figureCitation>
,
<figureCitation id="0D9145C72C40CE26FBD4C6F39A61F8E3" box="[1054,1086,1789,1813]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="12" pageNumber="347">5E</figureCitation>
,
<figureCitation id="0D9145C72C40CE26FB86C6F39ACEF8E3" box="[1100,1169,1789,1813]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="12" pageNumber="347">6C, D</figureCitation>
). The form of the epistome is the same as in the pilumnid genus
<taxonomicName id="52AA22C12C40CE26FB05C7139D27F8A3" authority="Ward, 1942" authorityName="Ward" authorityYear="1942" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FB05C7139B26F8C3" box="[1231,1401,1821,1845]" italics="true" pageId="12" pageNumber="347">Pseudolitochira</emphasis>
<bibRefCitation id="F13B24B32C40CE26FA4AC7139D27F8A3" author="Ward M" pageId="12" pageNumber="347" pagination="49 - 113" refId="ref15262" refString="Ward M (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2): 49 - 113." type="journal article" year="1942">Ward, 1942</bibRefCitation>
</taxonomicName>
(sensu
<bibRefCitation id="F13B24B32C40CE26FC1BC7309A21F8A3" author="Ng PKL &amp; Clark PF &amp; Clark B &amp; Kamanli SA" box="[977,1150,1853,1878]" pageId="12" pageNumber="347" pagination="377 - 391" refId="ref14159" refString="Ng PKL, Clark PF, Clark B &amp; Kamanli SA (2021) Pseudolitochira integra (Miers, 1884) (Crustacea: Brachyura: Pilumnidae): redescribed and illustrated from micro-CT scanning the type female. Zootaxa, 4969 (2): 377 - 391." type="journal article" year="2021">Ng et al., 2021</bibRefCitation>
). Species of
<taxonomicName id="52AA22C12C40CE26FADAC7339BE4F8A3" authorityName="Ward" authorityYear="1942" box="[1296,1467,1853,1877]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FADAC7339BE4F8A3" box="[1296,1467,1853,1877]" italics="true" pageId="12" pageNumber="347">Pseudolitochira</emphasis>
</taxonomicName>
, however, are very setose, with long silky setae that partially or completely obscure the margins and surfaces, and lack any trace of areoles on the carapace, with the regions smooth and hardly demarcated (
<bibRefCitation id="F13B24B32C40CE26FB86C7B39B7BF823" author="Ng PKL &amp; Clark PF" box="[1100,1316,1981,2005]" pageId="12" pageNumber="347" pagination="349 - 364" refId="ref14076" refString="Ng PKL &amp; Clark PF (2022 a) Descriptions of three new Pseudolitochira Ward, 1942 species (Crustacea: Brachyura: Pilumnidae) from the West Pacific. Journal of Natural History, 56 (5 - 8): 349 - 364." type="journal article" year="2022">Ng &amp; Clark, 2022a</bibRefCitation>
, b;
<bibRefCitation id="F13B24B32C40CE26FA85C7B09D23F803" author="Ng PKL &amp; Lin C-W" pageId="12" pageNumber="347" pagination="101 - 114" refId="ref14285" refString="Ng PKL &amp; Lin C-W (2023) Two new species of hairy crabs (Pilumnidae) and the first record of Crinitocinus alcocki (Borradaile, 1900) (Acidopsidae) (Crustacea: Brachyura) from Taiwan. Zootaxa, 5297 (1): 101 - 114." type="journal article" year="2023">Ng &amp; Lin, 2023</bibRefCitation>
;
<bibRefCitation id="F13B24B32C40CE26FC43C7D09DA1F803" author="Ng PKL" box="[905,1022,2013,2038]" pageId="12" pageNumber="347" pagination="120 - 126" refId="ref13924" refString="Ng PKL (2024 a) Pseudolitochira spinosa, a new species of pilumnid crab from Panglao, Philippines, with a note on P. setosa (A. Milne-Edwards, 1873) (Crustacea, Brachyura, Pilumnoidea). Zootaxa, 5512 (1): 120 - 126." type="journal article" year="2024">Ng, 2024a</bibRefCitation>
).
</paragraph>
<paragraph id="951559422C41CE27FFBBC0B09C1FFDE3" blockId="13.[113,753,189,534]" pageId="13" pageNumber="348">
In the general appearance of the carapace, especially in the overall shape and the reduced size of the fourth anterolateral tooth/spine,
<taxonomicName id="52AA22C12C41CE27FE69C0F39C8CFEE3" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[419,723,253,277]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C41CE27FE69C0F39C8CFEE3" box="[419,723,253,277]" italics="true" pageId="13" pageNumber="348">Lophoplax sextuberculata</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="52AA22C12C41CE27FEBBC1139C9DFEC3" authority="Ward, 1942" authorityName="Ward" authorityYear="1942" box="[369,706,285,309]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="A7DE85502C41CE27FEBBC1139C6BFEC3" box="[369,564,285,309]" italics="true" pageId="13" pageNumber="348">Viaderiana typica</emphasis>
<bibRefCitation id="F13B24B32C41CE27FDF4C1139C9DFEC3" author="Ward M" box="[574,706,285,309]" pageId="13" pageNumber="348" pagination="49 - 113" refId="ref15262" refString="Ward M (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2): 49 - 113." type="journal article" year="1942">Ward, 1942</bibRefCitation>
</taxonomicName>
, the
<typeStatus id="4A11E7E02C41CE27FFBBC1309EC0FEA0" box="[113,159,318,342]" pageId="13" pageNumber="348">type</typeStatus>
species of
<taxonomicName id="52AA22C12C41CE27FED7C1339C7DFEA3" authority="Ward, 1942" authorityName="Ward" authorityYear="1942" box="[285,546,317,341]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C41CE27FED7C1339FCAFEA3" box="[285,405,317,341]" italics="true" pageId="13" pageNumber="348">Viaderiana</emphasis>
<bibRefCitation id="F13B24B32C41CE27FE54C1339C7DFEA3" author="Ward M" box="[414,546,317,341]" pageId="13" pageNumber="348" pagination="49 - 113" refId="ref15262" refString="Ward M (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2): 49 - 113." type="journal article" year="1942">Ward, 1942</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="52AA22C12C41CE27FDE4C1339CAFFEA3" authorityName="Ward" authorityYear="1942" box="[558,752,317,341]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="A7DE85502C41CE27FDE4C1339CAFFEA3" box="[558,752,317,341]" italics="true" pageId="13" pageNumber="348">Viaderiana typica</emphasis>
</taxonomicName>
(and congeners), however, have the extensor margin of the ambulatory merus armed with at least one spine, the regions are never prominently raised to form granules, the ischium of the third maxilliped is distinctly more elongate, and the posterior epistomal margin is divided into three distinct lobes by distinct fissures (see Ng, 2023, 2024b).
</paragraph>
<paragraph id="951559422C41CE27FFBBC2309FC8FD80" blockId="13.[113,752,573,630]" pageId="13" pageNumber="348">
As such,
<taxonomicName id="52AA22C12C41CE27FF19C2339FB6FDA3" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[211,489,573,597]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C41CE27FF19C2339FB6FDA3" box="[211,489,573,597]" italics="true" pageId="13" pageNumber="348">Lophoplax sextuberculata</emphasis>
</taxonomicName>
is made the
<typeStatus id="4A11E7E02C41CE27FDA5C2309CC2FDA0" box="[623,669,574,598]" pageId="13" pageNumber="348">type</typeStatus>
species of
<taxonomicName id="52AA22C12C41CE27FF5AC2539F53FD83" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[144,268,605,629]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C41CE27FF5AC2539F53FD83" box="[144,268,605,629]" italics="true" pageId="13" pageNumber="348">Takedaplax</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C41CE27FED6C2509FCCFD80" box="[284,403,606,630]" pageId="13" pageNumber="348" rank="genus">new genus</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,317 @@
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<mods:title id="7C1342A2A7A84FC79AAF1E86EAAF0D25">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
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<paragraph id="951559422C41CE27FF4AC2939C4CFD23" blockId="13.[128,737,669,757]" pageId="13" pageNumber="348">
<heading id="CE5DEE2E2C41CE27FF4AC2939C4CFD23" bold="true" centered="true" fontSize="10" level="2" pageId="13" pageNumber="348" reason="2">
<emphasis id="A7DE85502C41CE27FF4AC2939C4CFD23" bold="true" pageId="13" pageNumber="348">
<taxonomicName id="52AA22C12C41CE27FF4AC2939C83FD43" ID-CoL="CL2XM" authority="(Takeda &amp; Kurata, 1984)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Takeda &amp; Kurata" baseAuthorityYear="1984" box="[128,732,669,693]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="sextuberculata" status="comb. nov.">
<emphasis id="A7DE85502C41CE27FF4AC2939FF4FD43" bold="true" box="[128,427,669,693]" italics="true" pageId="13" pageNumber="348">Takedaplax sextuberculata</emphasis>
(
<bibRefCitation id="F13B24B32C41CE27FE71C2939C8BFD43" author="Takeda M &amp; Kurata Y" box="[443,724,669,693]" pageId="13" pageNumber="348" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C41CE27FE84C2B39C4CFD23" box="[334,531,701,725]" pageId="13" pageNumber="348" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE27FEBEC2D39FB2FD03" box="[372,493,733,757]" pageId="13" pageNumber="348" type="description">
<paragraph id="951559422C41CE27FEBEC2D39FB2FD03" blockId="13.[128,737,669,757]" box="[372,493,733,757]" pageId="13" pageNumber="348">
(
<figureCitation id="0D9145C72C41CE27FEB6C2D39F96FD03" box="[380,457,733,757]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Figs. 8</figureCitation>
,
<figureCitation id="0D9145C72C41CE27FE1DC2D39FBAFD03" box="[471,485,733,757]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE27FFBBC3129CCEFC9D" pageId="13" pageNumber="348" type="reference_group">
<paragraph id="951559422C41CE27FFBBC3129CCEFC9D" blockId="13.[113,752,796,875]" pageId="13" pageNumber="348">
<treatmentCitationGroup id="B5BA7E6C2C41CE27FFBBC3129CCEFC9D" pageId="13" pageNumber="348">
<treatmentCitation id="140B7F532C41CE27FFBBC3129C26FCC7" author="Takeda M &amp; Kurata Y" box="[113,633,796,818]" page="200" pageId="13" pageNumber="348" year="1984">
<taxonomicName id="52AA22C12C41CE27FFBBC3129C26FCC7" ID-CoL="6QMFQ" authority="Takeda &amp; Kurata, 1984: 200" authorityName="Takeda &amp; Kurata" authorityPageNumber="200" authorityYear="1984" box="[113,633,796,818]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C41CE27FFBBC3129F37FCC7" box="[113,360,796,817]" italics="true" pageId="13" pageNumber="348">Lophoplax sextuberculata</emphasis>
<bibRefCitation id="F13B24B32C41CE27FEA6C3129C26FCC7" author="Takeda M &amp; Kurata Y" box="[364,633,796,818]" pageId="13" pageNumber="348" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984: 200</bibRefCitation>
</taxonomicName>
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, figs. 1416;
<treatmentCitation id="140B7F532C41CE27FF5DC3379F19FCB8" author="Takeda M" box="[151,326,825,847]" page="40" pageId="13" pageNumber="348" year="1993">
<bibRefCitation id="F13B24B32C41CE27FF5DC3379F19FCB8" author="Takeda M" box="[151,326,825,847]" pageId="13" pageNumber="348" refId="ref14889" refString="Takeda M (1993) Catalogue of crab specimens (Crustacea: Decapoda: Brachyura). Part 1. National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda, 1993: 40</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C41CE27FE98C3379C42FCB8" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[338,541,825,847]" page="143" pageId="13" pageNumber="348" year="2008">
<bibRefCitation id="F13B24B32C41CE27FE98C3379C42FCB8" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[338,541,825,847]" pageId="13" pageNumber="348" pagination="1 - 286" refId="ref14245" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 143</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C41CE27FDE3C3379EA3FC9D" author="Takeda M &amp; Komatsu H" page="176" pageId="13" pageNumber="348" year="2018">
<bibRefCitation id="F13B24B32C41CE27FDE3C3379EA3FC9D" author="Takeda M &amp; Komatsu H" pageId="13" pageNumber="348" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018: 176</bibRefCitation>
</treatmentCitation>
, fig. 6H;
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<bibRefCitation id="F13B24B32C41CE27FE90C35B9CD2FC9D" author="Takeda M &amp; Komatsu H" box="[346,653,853,875]" pageId="13" pageNumber="348" pagination="161 - 178" refId="ref14998" refString="Takeda M &amp; Komatsu H (2023) Updated checklist of the crabs (Crustacea: Decapoda: Brachyura) from the sea around the Ogasawara Islands, Japan. Bulletin of the National Museum of Nature and Science, Series A, 49 (4): 161 - 178." type="journal article" year="2023">Takeda &amp; Komatsu, 2023: 167</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE27FFBBC39D9F52FBDA" pageId="13" pageNumber="348" type="materials_examined">
<paragraph id="951559422C41CE27FFBBC39D9F52FBDA" blockId="13.[113,752,915,1068]" pageId="13" pageNumber="348">
<emphasis id="A7DE85502C41CE27FFBBC39D9F0CFC5D" bold="true" box="[113,339,915,939]" pageId="13" pageNumber="348">Material examined.</emphasis>
<materialsCitation id="25C2531F2C41CE27FE96C39A9F56FBDA" collectingDate="2008-10-28" collectionCode="R, V" country="Japan" elevation="81" latitude="27.078833" location="Chichi-jima" longLatPrecision="12" longitude="142.15517" municipality="Ogasawara Islands" pageId="13" pageNumber="348" specimenCount="1" specimenCount-male="1">
<specimenCount id="83AC92CB2C41CE27FE96C39A9FF9FC5A" box="[348,422,916,940]" count="1" pageId="13" pageNumber="348" type="male">1 male</specimenCount>
(4.0 ×
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) (NSMT-Cr S 1253), west of
<location id="90750F992C41CE27FED0C3BA9FC6FC3A" LSID="urn:lsid:plazi:treatment:1D03E8542C41CE3AFF4AC2939D57FE63:90750F992C41CE27FED0C3BA9FC6FC3A" box="[282,409,948,972]" country="Japan" latitude="27.078833" longLatPrecision="12" longitude="142.15517" municipality="Ogasawara Islands" name="Chichi-jima" pageId="13" pageNumber="348">Chichi-jima</location>
[= Island],
<collectingMunicipality id="7571C3382C41CE27FDD4C3BA9CB3FC3A" box="[542,748,948,972]" pageId="13" pageNumber="348">Ogasawara Islands</collectingMunicipality>
,
<collectingCountry id="EDBD19D22C41CE27FFBBC3DA9EEEFC1D" box="[113,177,980,1003]" name="Japan" pageId="13" pageNumber="348">Japan</collectingCountry>
,
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<geoCoordinate id="F09E3F852C41CE27FE8DC3DC9F91FC1D" box="[327,462,978,1003]" degrees="142" direction="east" minutes="09.16" orientation="longitude" pageId="13" pageNumber="348" precision="9" value="142.15266">142°09.16E</geoCoordinate>
<geoCoordinate id="F09E3F852C41CE27FE16C3DC9C07FC1D" box="[476,600,978,1003]" degrees="27" direction="north" minutes="04.73" orientation="latitude" pageId="13" pageNumber="348" precision="9" value="27.078833">27°04.73N</geoCoordinate>
<geoCoordinate id="F09E3F852C41CE27FDA8C3DC9CB3FC1D" box="[610,748,978,1003]" degrees="142" direction="east" minutes="09.31" orientation="longitude" pageId="13" pageNumber="348" precision="9" value="142.15517">142°09.31E</geoCoordinate>
, 83
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<elevation id="1E87BE712C41CE27FF56C3FA9E88FBFD" box="[156,215,1012,1036]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="348" unit="m" value="81.0">81 m</elevation>
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,
<collectionCode id="F3BBC1872C41CE27FF2DC3FA9EA6FBFA" box="[231,249,1012,1036]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="348">R</collectionCode>
/
<collectionCode id="F3BBC1872C41CE27FECBC3FA9F4BFBFA" box="[257,276,1012,1036]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="348" type="Museum">V</collectionCode>
Koyo, 2008 cruise, station KY-08-15,
<date id="E1147F822C41CE27FD1FC3FA9F56FBDA" pageId="13" pageNumber="348" value="2008-10-28">
<collectingDate id="F150866A2C41CE27FD1FC3FA9F56FBDA" pageId="13" pageNumber="348" value="2008-10-28">28 October 2008</collectingDate>
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.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE27FFBBC45D9FDDFB9A" box="[113,386,1107,1132]" pageId="13" pageNumber="348" type="diagnosis">
<paragraph id="951559422C41CE27FFBBC45D9FDDFB9A" blockId="13.[113,386,1107,1132]" box="[113,386,1107,1132]" pageId="13" pageNumber="348">
<emphasis id="A7DE85502C41CE27FFBBC45D9EB9FB9D" bold="true" box="[113,230,1107,1131]" pageId="13" pageNumber="348">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE25FFBBC49D9F6BF9AE" lastPageId="15" lastPageNumber="350" pageId="13" pageNumber="348" type="description">
<paragraph id="951559422C41CE27FFBBC49D9ACBFE23" blockId="13.[113,753,1171,2060]" lastBlockId="13.[812,1452,189,469]" pageId="13" pageNumber="348">
<emphasis id="A7DE85502C41CE27FFBBC49D9EA3FB5D" bold="true" box="[113,252,1171,1195]" pageId="13" pageNumber="348">Description.</emphasis>
Carapace rectangular; frontal and anterolateral margins with scattered tufts of short and medium-length setae, not brush-like, not concealing margins; regions well demarcated with 6 anterior areoles always large, distinct: epigastric areole with cristate anterior margin; postorbital areole transversely wide, anterior margin subcristate; hepatic areole rounded with subcristate anterior margin; protogastric, cardiac, mesobranchial and cardiobranchial regions gently convex to barely discernible in smaller specimens, gently raised to low areoles in larger specimens; gastro-cardiac with grooves shallow (
<figureCitation id="0D9145C72C41CE27FEBCC5DA9FAAFA1A" box="[374,501,1492,1516]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, B</figureCitation>
); intestinal region not prominently raised, no visible intestino-branchial areole; surface adjacent to posterior carapace margin almost flat, not forming ridge, no transverse channel visible (
<figureCitation id="0D9145C72C41CE27FD47C63A9CBDF9BA" box="[653,738,1588,1612]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8B</figureCitation>
); mesobranchial regions smooth; metabranchial regions covered with granules (
<figureCitation id="0D9145C72C41CE27FEBAC67A9F9AF97A" box="[368,453,1652,1676]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8B</figureCitation>
). Subhepatic and suborbital regions almost smooth; pterygostomial region with scattered small granules (
<figureCitation id="0D9145C72C41CE27FED7C6BA9F2CF93A" box="[285,371,1716,1740]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8D</figureCitation>
). Frontal margin distinctly bilobed, convex, with deep median cleft, lateral lobe very low, just visible in dorsal view (
<figureCitation id="0D9145C72C41CE27FE94C6FA9F90F8FA" box="[350,463,1780,1804]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, B</figureCitation>
). Supraorbital margin long, sinuous, with distinct submedian lobe, without fissure or cleft (
<figureCitation id="0D9145C72C41CE27FFB3C73A9E8DF8BA" box="[121,210,1844,1868]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8B</figureCitation>
). Orbit dorsal, supraorbital margin, eyes clearly visible in dorsal view; eyes freely movable, ocular peduncle relatively long, filling orbit (
<figureCitation id="0D9145C72C41CE27FE08C77A9C1BF87A" box="[450,580,1908,1932]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8B, D</figureCitation>
). Anterolateral margin with 4 visible teeth or spines (including external orbital tooth), external orbital tooth triangular, next 3 teeth acutely triangular to spiniform, last tooth smallest (
<figureCitation id="0D9145C72C41CE27FD09C7DA9EE6F7FA" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, B</figureCitation>
). Posterolateral margin sinuous, unarmed, margins gently converging towards almost straight posterior carapace margin (
<figureCitation id="0D9145C72C41CE27FC4EC0D39DABFF03" box="[900,1012,221,245]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, B</figureCitation>
). Antennules rectangular, relatively large; antennules folding laterally (
<figureCitation id="0D9145C72C41CE27FB94C0F39AEBFEE3" box="[1118,1204,253,277]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8D</figureCitation>
). Basal antennal article subquadrate; flagellum entering orbital hiatus. Posterior margin of epistome entire, without lateral fissures, median part triangular, lateral parts distinctly concave (8E, 9D). Endostome not obliquely recessed posteriorly into buccal cavern, subparallel to adjacent pterygostomial surface, low, short endostomial ridges present.
</paragraph>
<paragraph id="951559422C41CE27FCE7C1F39A2BFD63" blockId="13.[813,1453,509,661]" pageId="13" pageNumber="348">
Third maxilliped with ischium relatively short, subquadrate, length 1.3× width, with shallow median oblique sulcus; merus subquadrate, anterolateral angle slightly auriculiform. Exopod relatively stout, tip just reaching to before distal edge of merus (
<figureCitation id="0D9145C72C41CE27FC12C2739A67FD63" box="[984,1080,637,661]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Figs. 8C</figureCitation>
,
<figureCitation id="0D9145C72C41CE27FB8FC2739A38FD63" box="[1093,1127,637,661]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">9C</figureCitation>
).
</paragraph>
<paragraph id="951559422C41CE27FCE7C2B39B45FC43" blockId="13.[813,1452,701,949]" pageId="13" pageNumber="348">
Male thoracic sternum surface smooth; sternites 1 and 2 completely fused, wide, lateral margins gently concave to sinuous; sternites 3 and 4 fused with only very shallow median groove and lateral notches visible; suture between sternites 2 and 3 gently sinuous (
<figureCitation id="0D9145C72C41CE27FB44C3339ABFFCA3" box="[1166,1248,829,853]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8F</figureCitation>
). Sternite 8 clearly visible as subovate plate adjacent to pleonal somite 2 when pleon closed (
<figureCitation id="0D9145C72C41CE27FC03C3739A7CFC63" box="[969,1059,893,917]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">Fig. 9G</figureCitation>
). Tubercle of male pleonal locking mechanism on anterior one-third of somite 5.
</paragraph>
<paragraph id="951559422C41CE27FCE7C3D39BF3FA43" blockId="13.[812,1452,989,1462]" pageId="13" pageNumber="348">
Adult male chelipeds almost symmetrical, relatively slender (
<figureCitation id="0D9145C72C41CE27FCFFC3F39D84FBE3" box="[821,987,1021,1045]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, G, H</figureCitation>
). Basis-ischium fused but suture visible; extensor margins granulated (
<figureCitation id="0D9145C72C41CE27FBA0C4139AE2FBC3" box="[1130,1213,1053,1077]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="13" pageNumber="348">Fig. 1C</figureCitation>
). Merus short, surface slightly rugose; flexor and extensor margins with tubercles and granules; inner distal angle dentiform with low subdistal angle (
<figureCitation id="0D9145C72C41CE27FCBFC4739DB3FB63" box="[885,1004,1149,1173]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, H</figureCitation>
). Carpus surface granulate; dorsal surface of carpus with prominent longitudinal areole, separated from swollen lateral fringe by groove; inner distal angle with sharp tooth (
<figureCitation id="0D9145C72C41CE27FC76C4D39A1AFB03" box="[956,1093,1245,1269]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Figs. 8A, H</figureCitation>
,
<figureCitation id="0D9145C72C41CE27FB9EC4D39A29FB03" box="[1108,1142,1245,1269]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">9B</figureCitation>
). Chela with dorsal surface covered with granules; inner surface almost smooth; median and ventral surfaces covered with tubercles and granules; margins of chela with scattered long and short setae, not dense or brush-like; fingers slender, slightly shorter than palm, pigmented brown along distal half; occlusal margins with low teeth, forming distinct gape when closed (
<figureCitation id="0D9145C72C41CE27FAECC5939BFFFA43" box="[1318,1440,1437,1461]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8G, H</figureCitation>
).
</paragraph>
<paragraph id="951559422C41CE27FCE7C5D39A61F8A3" blockId="13.[813,1452,1501,1877]" pageId="13" pageNumber="348">
Ambulatory legs long, P4 longest, about 2.0× carapace width; P4 merus length about 4.3× height, distinctly longer than half carapace width; outer surface almost smooth; coxa rounded; merus elongate, P25 extensor margin not armed, entire, lined with short and long setae that does not obscure surface; surface of P25 gently convex, without median swelling or ridge; P25 carpus relatively smooth, with very low submarginal ridge, lowest or barely visible in P5; propodus laterally flattened, unarmed, relatively long in P24, shortest in P5; P24 dactylus falciform, unarmed, P5 dactylus shorter, slightly upturned; dactylo-propodal lock visible (
<figureCitation id="0D9145C72C41CE27FC4EC7339DBAF8A3" box="[900,997,1853,1877]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Figs. 8A</figureCitation>
,
<figureCitation id="0D9145C72C41CE27FC38C7339A6EF8A3" box="[1010,1073,1853,1877]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">9E, F</figureCitation>
).
</paragraph>
<paragraph id="951559422C41CE27FCE7C7739BC2F7E3" blockId="13.[813,1451,1917,2069]" pageId="13" pageNumber="348">
Male pleon triangular, all somites and telson free; somite 1 broadly rectangular, reaching base of P5 coxae; somite 2 transversely narrower than somite 1, subtrapezoidal; somite 3 broadly subtrapezoidal, as wide as somite 1, somites 4 and 5 trapezoidal, somite 6 and telson missing (
<figureCitation id="0D9145C72C41CE27FAF3C7F39BCEF7E3" box="[1337,1425,2045,2069]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">Fig. 9G</figureCitation>
).
</paragraph>
<caption id="C1D509CA2C42CE24FF42C7FD9AFDF7B5" ID-DOI="http://doi.org/10.5281/zenodo.14682783" ID-Zenodo-Dep="14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="14" pageNumber="349" startId="14.[136,172,2035,2057]" targetBox="[146,1464,189,2020]" targetPageId="14" targetType="figure">
<paragraph id="951559422C42CE24FF42C7FD9AFDF7B5" blockId="14.[136,1475,2035,2115]" pageId="14" pageNumber="349">
Fig. 8.
<taxonomicName id="52AA22C12C42CE24FF19C7FD9CB6F7FE" authority="(Takeda &amp; Kurata, 1984)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Takeda &amp; Kurata" baseAuthorityYear="1984" box="[211,745,2035,2057]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="349" phylum="Arthropoda" rank="species" species="sextuberculata" status="comb. nov.">
<emphasis id="A7DE85502C42CE24FF19C7FD9F81F7FE" box="[211,478,2035,2056]" italics="true" pageId="14" pageNumber="349">Takedaplax sextuberculata</emphasis>
(
<bibRefCitation id="F13B24B32C42CE24FE24C7FD9CBDF7FE" author="Takeda M &amp; Kurata Y" box="[494,738,2035,2057]" pageId="14" pageNumber="349" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C42CE24FD3DC7FA9DFDF7FF" box="[759,930,2035,2057]" pageId="14" pageNumber="349" rank="species">new combination</taxonomicNameLabel>
, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped.
</paragraph>
</caption>
<caption id="C1D509CA2C43CE25FFBBC52E9F58FA5E" ID-DOI="http://doi.org/10.5281/zenodo.14682785" ID-Zenodo-Dep="14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="15" pageNumber="350" startId="15.[113,149,1312,1334]" targetBox="[114,1452,188,1301]" targetPageId="15" targetType="figure">
<paragraph id="951559422C43CE25FFBBC52E9F58FA5E" blockId="15.[113,1452,1312,1448]" pageId="15" pageNumber="350">
Fig. 9.
<taxonomicName id="52AA22C12C43CE25FF77C52E9C8CFAC3" authority="(Takeda &amp; Kurata, 1984)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Takeda &amp; Kurata" baseAuthorityYear="1984" box="[189,723,1312,1334]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="350" phylum="Arthropoda" rank="species" species="sextuberculata" status="comb. nov.">
<emphasis id="A7DE85502C43CE25FF77C52E9F97FAC3" box="[189,456,1312,1333]" italics="true" pageId="15" pageNumber="350">Takedaplax sextuberculata</emphasis>
(
<bibRefCitation id="F13B24B32C43CE25FE1DC52E9C93FAC3" author="Takeda M &amp; Kurata Y" box="[471,716,1312,1334]" pageId="15" pageNumber="350" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C43CE25FD2AC52E9DD3FAC0" box="[736,908,1312,1334]" pageId="15" pageNumber="350" rank="species">new combination</taxonomicNameLabel>
. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after
<bibRefCitation id="F13B24B32C43CE25FF27C5339F87FAA4" author="Takeda M &amp; Kurata Y" box="[237,472,1341,1363]" pageId="15" pageNumber="350" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm.
</paragraph>
</caption>
<paragraph id="951559422C43CE25FFBBC5EE9F6BF9AE" blockId="15.[113,752,1504,1624]" pageId="15" pageNumber="350">
G1strongly sinuous, distally recurved, tapering to slightly rounded tip, hooked inwards; subdistal surface with 1 long stout seta (
<figureCitation id="0D9145C72C43CE25FF25C62E9F3BF9CE" box="[239,356,1568,1592]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="15" pageNumber="350">Fig. 9H, I</figureCitation>
). G2 short, about ¼ length of G1, sigmoid (
<figureCitation id="0D9145C72C43CE25FF12C64E9F78F9AE" box="[216,295,1600,1624]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="15" pageNumber="350">Fig. 9J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C43CE3AFFBBC68E9D57FE63" lastPageId="16" lastPageNumber="351" pageId="15" pageNumber="350" type="discussion">
<paragraph id="951559422C43CE25FFBBC68E9C57F88E" blockId="15.[113,753,1664,1912]" pageId="15" pageNumber="350">
<emphasis id="A7DE85502C43CE25FFBBC68E9EBFF96E" bold="true" box="[113,224,1664,1688]" pageId="15" pageNumber="350">Remarks.</emphasis>
<taxonomicName id="52AA22C12C43CE25FF2FC68E9FA7F96E" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[229,504,1664,1688]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="350" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C43CE25FF2FC68E9FA7F96E" box="[229,504,1664,1688]" italics="true" pageId="15" pageNumber="350">Lophoplax sextuberculata</emphasis>
</taxonomicName>
was described from the female
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(8.2 ×
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) and
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(both 5.2 ×
<quantity id="5252F4A72C43CE25FF7CC6CE9F52F92E" box="[182,269,1728,1752]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="350" unit="mm" value="3.8">3.8 mm</quantity>
) collected from inside a fish stomach from Takinoura Bay, Ani-jima [=
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], Ogasawara Islands, and a
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male (5.5 ×
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) from Tanegashima [=
<collectingCountry id="EDBD19D22C43CE25FD6CC70E9CB5F8EE" box="[678,746,1792,1816]" name="Iceland" pageId="15" pageNumber="350">Island</collectingCountry>
] (southern Kyushu) (
<bibRefCitation id="F13B24B32C43CE25FE8CC72E9C19F8CE" author="Takeda M &amp; Kurata Y" box="[326,582,1824,1848]" pageId="15" pageNumber="350" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
;
<bibRefCitation id="F13B24B32C43CE25FD9AC72E9CBBF8CE" author="Takeda M" box="[592,740,1824,1848]" pageId="15" pageNumber="350" refId="ref14889" refString="Takeda M (1993) Catalogue of crab specimens (Crustacea: Decapoda: Brachyura). Part 1. National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda, 1993</bibRefCitation>
).
<bibRefCitation id="F13B24B32C43CE25FFBBC74E9FFAF8AE" author="Takeda M &amp; Marumura M" box="[113,421,1856,1880]" pageId="15" pageNumber="350" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura (1995)</bibRefCitation>
reported additional specimens from the Kii Peninsula and Ryukyus.
</paragraph>
<paragraph id="951559422C43CE3AFFBBC7AE9D57FE63" blockId="15.[113,753,1952,2104]" lastBlockId="16.[136,776,189,406]" lastPageId="16" lastPageNumber="351" pageId="15" pageNumber="350">
They noted that while the median and posterior surfaces of the carapace were smooth in the
<typeStatus id="4A11E7E02C43CE25FE2BC7CF9C50F82F" box="[481,527,1985,2009]" pageId="15" pageNumber="350">type</typeStatus>
specimens from the Ogasawara Islands (as in the case of the present specimen; see also
<bibRefCitation id="F13B24B32C43CE25FF04C80E9FB6F7EE" author="Takeda M &amp; Komatsu H" box="[206,489,2048,2072]" pageId="15" pageNumber="350" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018</bibRefCitation>
: fig. 6H); the specimens from Tanegashima [=
<collectingCountry id="EDBD19D22C43CE25FEACC82E9FF5F7CE" box="[358,426,2080,2104]" name="Iceland" pageId="15" pageNumber="350">Island</collectingCountry>
] and Kii Peninsula had seven additional areoles in two transverse rows on the following regions: two on the protogastric, one each on anterior parts of the mesobranchials, one cardiac and one each on the cardiobranchials (see
<bibRefCitation id="F13B24B32C43CE25FBD4C64E9B23F9AE" author="Takeda M &amp; Marumura M" box="[1054,1404,1600,1624]" pageId="15" pageNumber="350" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura, 1995: 89</bibRefCitation>
, 91, fig. 1). They noted that the only specimen they had from the Ryukyus was small (4.8 × 4.0 mm) and had the median and posterior parts of the carapace smooth (
<bibRefCitation id="F13B24B32C43CE25FB0CC6AE9DD6F92E" author="Takeda M &amp; Marumura M" pageId="15" pageNumber="350" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura, 1995: 91</bibRefCitation>
). As such,
<bibRefCitation id="F13B24B32C43CE25FC33C6CE9B19F92E" author="Takeda M &amp; Marumura M" box="[1017,1350,1728,1752]" pageId="15" pageNumber="350" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura (1995: 91)</bibRefCitation>
observed that “… there may be some individual differences as for the development of the areolets on the posterior part, but it is concluded that
<taxonomicName id="52AA22C12C43CE25FC13C72E9AA9F8CE" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[985,1270,1824,1848]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="350" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C43CE25FC13C72E9AA9F8CE" box="[985,1270,1824,1848]" italics="true" pageId="15" pageNumber="350">Lophoplax sextuberculata</emphasis>
</taxonomicName>
is characterised by having 13 areolets, not 6 mentioned in the original description and scientific name”. Examining the present specimen from the Ogasawara Islands, we note that its protogastric, mesobranchial, cardiac, and cardiobranchial regions are only gently convex and smooth, and barely discernible as areolets in dorsal view (
<figureCitation id="0D9145C72C43CE25FB0CC7EE9B04F80E" box="[1222,1371,2016,2040]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="15" pageNumber="350">Fig. 8A, B, D</figureCitation>
). In the colour figure of this specimen (
<bibRefCitation id="F13B24B32C43CE25FB4CC80E9BF7F7EE" author="Takeda M &amp; Komatsu H" box="[1158,1448,2048,2072]" pageId="15" pageNumber="350" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018</bibRefCitation>
: fig. 6H), we note that these regions are also slightly more orangish than the surrounding beige surface as described by
<bibRefCitation id="F13B24B32C5CCE3AFF42C0D39FE8FF03" author="Takeda M &amp; Marumura M" box="[136,439,221,245]" pageId="16" pageNumber="351" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura (1995)</bibRefCitation>
for their specimens. As such, it seems likely that these areolets are present but not always discernible, probably becoming more prominent in larger individuals.
<bibRefCitation id="F13B24B32C5CCE3AFEC0C1309FB7FEA3" author="Ng PKL &amp; Rahayu DL" box="[266,488,317,342]" pageId="16" pageNumber="351" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu (2023)</bibRefCitation>
reported a similar situation for
<taxonomicName id="52AA22C12C5CCE3AFF7AC1539FF9FE83" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Ng &amp; Rahayu" baseAuthorityYear="2023" box="[176,422,349,373]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="sordidus">
<emphasis id="A7DE85502C5CCE3AFF7AC1539FF9FE83" box="[176,422,349,373]" italics="true" pageId="16" pageNumber="351">Myopilumnus sordidus</emphasis>
</taxonomicName>
and
<taxonomicName id="52AA22C12C5CCE3AFE2AC1509CD2FE83" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[480,653,349,373]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="vermiculatus">
<emphasis id="A7DE85502C5CCE3AFE2AC1509CD2FE83" box="[480,653,349,373]" italics="true" pageId="16" pageNumber="351">M. vermiculatus</emphasis>
</taxonomicName>
, where the areoles are relatively poorly developed in young specimens.
</paragraph>
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</treatment>
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View file

@ -0,0 +1,337 @@
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<mods:title id="567D2FE8C49EDA20FFAB691F99807578">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
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<emphasis id="A7DE85502C45CE23FC93C3F39AEDFBC3" bold="true" pageId="9" pageNumber="344">
<taxonomicName id="52AA22C12C45CE23FC93C3F39B1DFBE3" ID-CoL="CKZYG" authority="(Edmondson, 1951)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[857,1346,1021,1045]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="symmetrinuda" status="comb. nov.">
<emphasis id="A7DE85502C45CE23FC93C3F39A06FBE3" bold="true" box="[857,1113,1021,1045]" italics="true" pageId="9" pageNumber="344">Hosekia symmetrinuda</emphasis>
(
<bibRefCitation id="F13B24B32C45CE23FBA3C3F39B64FBE3" author="Edmondson CH" box="[1129,1339,1021,1045]" pageId="9" pageNumber="344" pagination="183 - 243" refId="ref13456" refString="Edmondson CH (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13): 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
)
</taxonomicName>
,
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</emphasis>
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</subSubSection>
<subSubSection id="DDB00AC92C45CE23FBE5C4339AF7FBA3" box="[1071,1192,1085,1109]" pageId="9" pageNumber="344" type="description">
<paragraph id="951559422C45CE23FBE5C4339AF7FBA3" blockId="9.[857,1406,1021,1109]" box="[1071,1192,1085,1109]" pageId="9" pageNumber="344">
(
<figureCitation id="0D9145C72C45CE23FBFDC4339ADBFBA3" box="[1079,1156,1085,1109]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="9" pageNumber="344">Figs. 6</figureCitation>
,
<figureCitation id="0D9145C72C45CE23FB58C4339AFFFBA3" box="[1170,1184,1085,1109]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="9" pageNumber="344">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C45CE23FCE6C4729A00FAC8" pageId="9" pageNumber="344" type="reference_group">
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<treatmentCitationGroup id="B5BA7E6C2C45CE23FCE6C4729AE6FB1E" pageId="9" pageNumber="344">
<treatmentCitation id="140B7F532C45CE23FCE6C4729BDEFB67" author="Edmondson CH" box="[812,1409,1148,1170]" page="233" pageId="9" pageNumber="344" year="1951">
<taxonomicName id="52AA22C12C45CE23FCE6C4729BDEFB67" ID-CoL="4NV9Q" authority="Edmondson, 1951: 233" authorityName="Edmondson" authorityPageNumber="233" authorityYear="1951" box="[812,1409,1148,1170]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="symmetrinudus">
<emphasis id="A7DE85502C45CE23FCE6C4729AC6FB67" box="[812,1177,1148,1169]" italics="true" pageId="9" pageNumber="344">Pseudocryptocoeloma symmetrinudus</emphasis>
<bibRefCitation id="F13B24B32C45CE23FB55C4729BDEFB67" author="Edmondson CH" box="[1183,1409,1148,1170]" pageId="9" pageNumber="344" pagination="183 - 243" refId="ref13456" refString="Edmondson CH (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13): 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951: 233</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 34;
<treatmentCitation id="140B7F532C45CE23FCB2C4979A7FFB59" author="Serene R" box="[888,1056,1177,1199]" page="86" pageId="9" pageNumber="344" year="1968">
<bibRefCitation id="F13B24B32C45CE23FCB2C4979A7FFB59" author="Serene R" box="[888,1056,1177,1199]" pageId="9" pageNumber="344" pagination="33 - 120" refId="ref14694" refString="Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science No. 1: 33 - 120." type="book chapter" year="1968">Serène, 1968: 86</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FBE1C4979AF1FB58" author="Ng PKL" box="[1067,1198,1177,1198]" page="79" pageId="9" pageNumber="344" year="1987">
<bibRefCitation id="F13B24B32C45CE23FBE1C4979AF1FB58" author="Ng PKL" box="[1067,1198,1177,1198]" pageId="9" pageNumber="344" pagination="69 - 111" refId="ref13788" refString="Ng PKL (1987) The IndoPacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). IndoMalayan Zoology, 1987, 4 (1): 69 - 111, pl. 1." type="journal article" year="1987">Ng, 1987: 79</bibRefCitation>
</treatmentCitation>
, 97;
<treatmentCitation id="140B7F532C45CE23FB2AC4979BF8FB58" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1248,1447,1177,1199]" page="144" pageId="9" pageNumber="344" year="2008">
<bibRefCitation id="F13B24B32C45CE23FB2AC4979BF8FB58" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1248,1447,1177,1199]" pageId="9" pageNumber="344" pagination="1 - 286" refId="ref14245" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 144</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FC98C4B89AC9FB3D" author="Marumura M &amp; Takeda M" box="[850,1174,1205,1227]" page="192" pageId="9" pageNumber="344" year="2012">
<bibRefCitation id="F13B24B32C45CE23FC98C4B89AC9FB3D" author="Marumura M &amp; Takeda M" box="[850,1174,1205,1227]" pageId="9" pageNumber="344" pagination="189 - 197" refId="ref13535" refString="Marumura M &amp; Takeda M (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1): 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda, 2012: 192</bibRefCitation>
</treatmentCitation>
, figs. 2A, 3, 4;
<treatmentCitation id="140B7F532C45CE23FAFFC4B89DF5FB11" author="Maenosono T" page="32" pageId="9" pageNumber="344" year="2019">
<bibRefCitation id="F13B24B32C45CE23FAFFC4B89DF5FB11" author="Maenosono T" pageId="9" pageNumber="344" pagination="19 - 44" refId="ref13491" refString="Maenosono T (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48: 19 - 44. [In Japanese with English abstract]" type="journal article" year="2019">Maenosono, 2019: 32</bibRefCitation>
</treatmentCitation>
, figs. 8B, 10, 13B, 14K, L.
</treatmentCitationGroup>
</paragraph>
<paragraph id="951559422C45CE23FCE6C4E19A00FAC8" blockId="9.[812,1452,1148,1343]" pageId="9" pageNumber="344">
<treatmentCitationGroup id="B5BA7E6C2C45CE23FCE6C4E19A00FAC8" pageId="9" pageNumber="344">
<taxonomicName id="52AA22C12C45CE23FCE6C4E19A76FAF2" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[812,1065,1263,1284]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="symmetrinudus">
<emphasis id="A7DE85502C45CE23FCE6C4E19A76FAF2" box="[812,1065,1263,1284]" italics="true" pageId="9" pageNumber="344">Lophoplax symmetrinudus</emphasis>
</taxonomicName>
<treatmentCitation id="140B7F532C45CE23FB81C4E19B03FAF2" author="Takeda M &amp; Kurata Y" box="[1099,1372,1263,1285]" page="201" pageId="9" pageNumber="344" year="1984">
<bibRefCitation id="F13B24B32C45CE23FB81C4E19B03FAF2" author="Takeda M &amp; Kurata Y" box="[1099,1372,1263,1285]" pageId="9" pageNumber="344" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984: 201</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FAAEC4E19A63FAD7" author="Takeda M &amp; Marumura M" page="90" pageId="9" pageNumber="344" year="1995">
<bibRefCitation id="F13B24B32C45CE23FAAEC4E19A63FAD7" author="Takeda M &amp; Marumura M" pageId="9" pageNumber="344" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura, 1995: 90</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FB82C5029B39FAD4" author="Poore GCB &amp; Ahyong ST" box="[1096,1382,1292,1314]" page="680" pageId="9" pageNumber="344" year="2023">
<bibRefCitation id="F13B24B32C45CE23FB82C5029B39FAD4" author="Poore GCB &amp; Ahyong ST" box="[1096,1382,1292,1314]" pageId="9" pageNumber="344" refId="ref14446" refString="Poore GCB &amp; Ahyong ST (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Collingwood, Australia, xii + 890 pp." type="book" year="2023">Poore &amp; Ahyong, 2023: 680</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FAB8C5029A51FAC8" author="Ng PKL &amp; Rahayu DL" page="444" pageId="9" pageNumber="344" year="2023">
<bibRefCitation id="F13B24B32C45CE23FAB8C5029A51FAC8" author="Ng PKL &amp; Rahayu DL" pageId="9" pageNumber="344" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu, 2023: 444</bibRefCitation>
</treatmentCitation>
, fig. 12.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C45CE23FCE6C5699AB0F9C9" pageId="9" pageNumber="344" type="materials_examined">
<paragraph id="951559422C45CE23FCE6C5699AB0F9C9" blockId="9.[812,1451,1383,1599]" pageId="9" pageNumber="344">
<emphasis id="A7DE85502C45CE23FCE6C5699A50FA89" bold="true" box="[812,1039,1383,1407]" pageId="9" pageNumber="344">Material examined.</emphasis>
<materialsCitation id="25C2531F2C45CE23FBD3C5699AB0F9C9" collectingDate="2016-02-10" collectingDateMax="2020-04-07" collectingDateMin="2016-02-10" collectionCode="T" collectorName="Siufaga &amp; Tau &amp; W. Harris &amp; Others &amp; Bise &amp; Motobu &amp; Ryukyus &amp; Maenosono" country="Samoa" county="Odo Beach" location="Ryukyus" municipality="Itoman" pageId="9" pageNumber="344" specimenCode="BPBM 5109" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<typeStatus id="4A11E7E02C45CE23FBD3C5699ADEFA89" box="[1049,1153,1383,1407]" pageId="9" pageNumber="344" type="holotype">Holotype</typeStatus>
:
<specimenCode id="C50CF1392C45CE23FB44C5699B44FA89" box="[1166,1307,1383,1407]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="9" pageNumber="344" type="Museum">BPBM 5109</specimenCode>
, male (6.5 × 5.0 mm),
<collectorName id="385F3C942C45CE23FC56C5899DAEFA69" box="[924,1009,1415,1439]" pageId="9" pageNumber="344">Siufaga</collectorName>
,
<collectorName id="385F3C942C45CE23FBCAC5899A72FA68" box="[1024,1069,1415,1438]" pageId="9" pageNumber="344">Tau</collectorName>
,
<collectingCountry id="EDBD19D22C45CE23FBF1C5899AD8FA69" box="[1083,1159,1415,1439]" name="Samoa" pageId="9" pageNumber="344">Samoa</collectingCountry>
, low tide, table reef, coll.
<collectorName id="385F3C942C45CE23FCE6C5A99DC2FA49" box="[812,925,1447,1471]" pageId="9" pageNumber="344">W. Harris</collectorName>
, 1937.
<collectorName id="385F3C942C45CE23FC3BC5A99A62FA49" box="[1009,1085,1447,1471]" pageId="9" pageNumber="344">Others</collectorName>
: RUMF-ZC-5199,
<specimenCount id="83AC92CB2C45CE23FAD8C5A99B02FA49" box="[1298,1373,1447,1471]" count="1" pageId="9" pageNumber="344" type="male">1 male</specimenCount>
(7.6 ×
<quantity id="5252F4A72C45CE23FCE6C5C99DDDFA29" box="[812,898,1479,1503]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="9" pageNumber="344" unit="mm" value="4.2">4.2 mm</quantity>
),
<collectorName id="385F3C942C45CE23FC5CC5C99D97FA29" box="[918,968,1479,1503]" pageId="9" pageNumber="344">Bise</collectorName>
,
<collectorName id="385F3C942C45CE23FC1EC5C99A74FA29" box="[980,1067,1479,1503]" pageId="9" pageNumber="344">Motobu</collectorName>
,
<collectingRegion id="576E97A02C45CE23FBFDC5C99AC5FA29" box="[1079,1178,1479,1503]" country="Japan" name="Okinawa" pageId="9" pageNumber="344">Okinawa</collectingRegion>
,
<collectorName id="385F3C942C45CE23FB6FC5C99B58FA29" box="[1189,1287,1479,1503]" pageId="9" pageNumber="344">Ryukyus</collectorName>
,
<collectingCountry id="EDBD19D22C45CE23FAD9C5C99B0DFA28" box="[1299,1362,1479,1502]" name="Japan" pageId="9" pageNumber="344">Japan</collectingCountry>
, coll.
<collectionCode id="F3BBC1872C45CE23FA5EC5C99BF8FA28" box="[1428,1447,1479,1502]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="344">T</collectionCode>
.
<collectorName id="385F3C942C45CE23FCE6C5E99DF4FA08" box="[812,939,1511,1534]" pageId="9" pageNumber="344">Maenosono</collectorName>
,
<date id="E1147F822C45CE23FC72C5E99A22FA09" box="[952,1149,1511,1535]" pageId="9" pageNumber="344" value="2016-02-10">
<collectingDate id="F150866A2C45CE23FC72C5E99A22FA09" box="[952,1149,1511,1535]" pageId="9" pageNumber="344" value="2016-02-10">10 February 2016</collectingDate>
</date>
; RUMF-ZC-7536,
<specimenCount id="83AC92CB2C45CE23FA87C5E99BF4FA09" box="[1357,1451,1511,1535]" count="1" pageId="9" pageNumber="344" type="female">1 female</specimenCount>
(9.7 ×
<quantity id="5252F4A72C45CE23FCB0C6099D8BF9E9" box="[890,980,1543,1567]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="9" pageNumber="344" unit="mm" value="6.9">6.9 mm</quantity>
),
<collectingCounty id="7C7421CE2C45CE23FC20C6099A37F9E9" box="[1002,1128,1543,1567]" pageId="9" pageNumber="344">Odo Beach</collectingCounty>
,
<collectingMunicipality id="7571C3382C45CE23FBBCC6099A9AF9E8" box="[1142,1221,1543,1566]" pageId="9" pageNumber="344">Itoman</collectingMunicipality>
,
<collectingRegion id="576E97A02C45CE23FB19C6099B68F9E9" box="[1235,1335,1543,1567]" country="Japan" name="Okinawa" pageId="9" pageNumber="344">Okinawa</collectingRegion>
,
<location id="90750F992C45CE23FA8FC6099BF8F9E9" LSID="urn:lsid:plazi:treatment:1D03E8542C45CE26FC93C3F39D57F943:90750F992C45CE23FA8FC6099BF8F9E9" box="[1349,1447,1543,1567]" country="Samoa" county="Odo Beach" municipality="Itoman" name="Ryukyus" pageId="9" pageNumber="344" stateProvince="Okinawa">Ryukyus</location>
,
<collectingCountry id="EDBD19D22C45CE23FCE6C6299D34F9C8" box="[812,875,1575,1598]" name="Japan" pageId="9" pageNumber="344">Japan</collectingCountry>
, coll.
<collectionCode id="F3BBC1872C45CE23FC7AC6299D9CF9C8" box="[944,963,1575,1598]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="344">T</collectionCode>
.
<location id="90750F992C45CE23FC05C6299A11F9C8" LSID="urn:lsid:plazi:treatment:1D03E8542C45CE26FC93C3F39D57F943:90750F992C45CE23FC05C6299A11F9C8" box="[975,1102,1575,1598]" country="Samoa" county="Odo Beach" municipality="Itoman" name="Maenosono" pageId="9" pageNumber="344" stateProvince="Okinawa">Maenosono</location>
,
<date id="E1147F822C45CE23FB91C6299AB4F9C9" box="[1115,1259,1575,1599]" pageId="9" pageNumber="344" value="2020-04-07">
<collectingDate id="F150866A2C45CE23FB91C6299AB4F9C9" box="[1115,1259,1575,1599]" pageId="9" pageNumber="344" value="2020-04-07">7 April 2020</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C45CE23FCE6C6699A62F989" box="[812,1085,1639,1663]" pageId="9" pageNumber="344" type="diagnosis">
<paragraph id="951559422C45CE23FCE6C6699A62F989" blockId="9.[812,1085,1639,1663]" box="[812,1085,1639,1663]" pageId="9" pageNumber="344">
<emphasis id="A7DE85502C45CE23FCE6C6699DFEF989" bold="true" box="[812,929,1639,1663]" pageId="9" pageNumber="344">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C45CE26FCE6C6A99F30FA03" lastPageId="12" lastPageNumber="347" pageId="9" pageNumber="344" type="description">
<paragraph id="951559422C45CE20FCE6C6A99B27F7FD" blockId="9.[812,1452,1703,2111]" lastBlockId="10.[835,1475,1682,2059]" lastPageId="10" lastPageNumber="345" pageId="9" pageNumber="344">
<emphasis id="A7DE85502C45CE23FCE6C6A99DE4F949" bold="true" box="[812,955,1703,1727]" pageId="9" pageNumber="344">Description.</emphasis>
Carapace transversely ovate-subhexagonal; frontal and anterolateral margins with scattered tufts of short and long setae, not brush-like, not concealing margins; regions well demarcated with areoles smooth, prominent: epigastric areoles distinct; protogastric areole well developed, extending obliquely to branchial region, approximately separated by median constriction; cardio-branchial areole transversely ovate, well-developed; gastro-cardiac region depressed, with grooves barely visible (
<figureCitation id="0D9145C72C45CE23FB12C7A99B07F848" box="[1240,1368,1959,1983]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="9" pageNumber="344">Figs. 6A, B</figureCitation>
,
<figureCitation id="0D9145C72C45CE23FAAEC7A99D1DF829" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="9" pageNumber="344">7A, C, D</figureCitation>
); intestino-branchial areoles prominent, transversely ovate, one on each side of low intestinal region; posterior carapace margin cristate, adjacent to prominent intestino-branchial areole, forming narrow channel extending to beginning of sub-branchial region; no ridges on sub-branchial region (
<figureCitation id="0D9145C72C46CE20FF5AC6BC9F4EF93C" box="[144,273,1714,1738]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6A, B</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FED4C6BD9FDBF93D" box="[286,388,1715,1739]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7A, C, D</figureCitation>
). Subhepatic and suborbital regions slightly rugose to almost smooth; pterygostomial region with numerous closely packed small granules (
<figureCitation id="0D9145C72C46CE20FD4DC6FC9CA3F8FC" box="[647,764,1778,1802]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6CE</figureCitation>
). Frontal margin bilobed, gently convex, with narrow, small median notch, without lateral lobe (
<figureCitation id="0D9145C72C46CE20FDD4C73C9CFBF8BC" box="[542,676,1842,1866]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6AD</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FD79C73D9CA4F8BD" box="[691,763,1843,1867]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7C, D</figureCitation>
). Supraorbital margin short, sinuous, with low lobe on posterior half (formed by anterior margin of hepatic areole), without fissure or cleft (
<figureCitation id="0D9145C72C46CE20FEF9C79C9FCCF85C" box="[307,403,1938,1962]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FE55C79D9FBCF85D" box="[415,483,1939,1963]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7C, D</figureCitation>
). Orbit dorsal, supraorbital margin appears confluent with general carapace outline, eyes clearly visible in dorsal view; eyes mobile, ocular peduncle short, filling orbit (
<figureCitation id="0D9145C72C46CE20FE96C7FC9F8AF7FC" box="[348,469,2034,2058]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6BD</figureCitation>
). Anterolateral margin with 3 distinct teeth (including external orbital tooth), first 2 broadly triangular with cristate margin, last tooth lower, more rounded; distinct groove visible adjacent to margin, formed with outer margin of hepatic lobe (
<figureCitation id="0D9145C72C46CE20FAE3C6DC9BD2F91C" box="[1321,1421,1746,1770]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FA56C6DD9D06F8FD" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7C, D</figureCitation>
). Posterolateral margin gently convex, smooth, unarmed; margins distinctly converging towards posterior carapace margin (
<figureCitation id="0D9145C72C46CE20FC57C73C9DA4F8BC" box="[925,1019,1842,1866]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FBCFC73D9A18F8BD" box="[1029,1095,1843,1867]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7C, D</figureCitation>
). Antennules rectangular, relatively large; antennules folding laterally (
<figureCitation id="0D9145C72C46CE20FB60C75C9B46F89D" box="[1194,1305,1874,1899]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6C, D</figureCitation>
). Basal antennal article subquadrate; flagellum enters orbital hiatus (
<figureCitation id="0D9145C72C46CE20FA5FC77C9DD8F85D" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6C, D</figureCitation>
). Epistome with biconcave posterior margin, median lobe triangular with very short median fissue (
<figureCitation id="0D9145C72C46CE20FAF7C7BC9BEAF83D" box="[1341,1461,1970,1995]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6C, D</figureCitation>
). Endostome distinct, sloping gradually posteriorly into buccal cavern, endostomial ridges, short, low (
<figureCitation id="0D9145C72C46CE20FB38C7FC9B34F7FD" box="[1266,1387,2034,2059]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6C, D</figureCitation>
).
</paragraph>
<caption id="C1D509CA2C46CE20FF42C6029B9CF9AD" ID-DOI="http://doi.org/10.5281/zenodo.14682775" ID-Zenodo-Dep="14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345" startId="10.[136,171,1548,1570]" targetBox="[135,1473,189,1532]" targetPageId="10" targetType="figure">
<paragraph id="951559422C46CE20FF42C6029B9CF9AD" blockId="10.[136,1475,1548,1627]" pageId="10" pageNumber="345">
Fig. 6.
<taxonomicName id="52AA22C12C46CE20FF18C6029C20F9D4" authority="(Edmondson, 1951)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[210,639,1548,1570]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="345" phylum="Arthropoda" rank="species" species="symmetrinuda" status="comb. nov.">
<emphasis id="A7DE85502C46CE20FF18C6029FEDF9D7" box="[210,434,1548,1569]" italics="true" pageId="10" pageNumber="345">Hosekia symmetrinuda</emphasis>
(
<bibRefCitation id="F13B24B32C46CE20FE0BC6029C27F9D4" author="Edmondson CH" box="[449,632,1548,1570]" pageId="10" pageNumber="345" pagination="183 - 243" refId="ref13456" refString="Edmondson CH (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13): 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C46CE20FD46C6029D67F9D4" box="[652,824,1548,1570]" pageId="10" pageNumber="345" rank="species">new combination</taxonomicNameLabel>
, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela.
</paragraph>
</caption>
<caption id="C1D509CA2C47CE21FFBBC6949FA8F8F0" ID-DOI="http://doi.org/10.5281/zenodo.14682777" ID-Zenodo-Dep="14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346" startId="11.[113,148,1690,1712]" targetBox="[113,1451,189,1674]" targetPageId="11" targetType="figure">
<paragraph id="951559422C47CE21FFBBC6949FA8F8F0" blockId="11.[113,1452,1690,1798]" pageId="11" pageNumber="346">
Fig. 7.
<taxonomicName id="52AA22C12C47CE21FF76C6949C33F946" authority="(Edmondson, 1951)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[188,620,1690,1712]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="symmetrinuda" status="comb. nov.">
<emphasis id="A7DE85502C47CE21FF76C6949FC1F959" box="[188,414,1690,1711]" italics="true" pageId="11" pageNumber="346">Hosekia symmetrinuda</emphasis>
(
<bibRefCitation id="F13B24B32C47CE21FE64C6949C3AF946" author="Edmondson CH" box="[430,613,1690,1712]" pageId="11" pageNumber="346" pagination="183 - 243" refId="ref13456" refString="Edmondson CH (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13): 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C47CE21FDB0C6949D77F946" box="[634,808,1690,1712]" pageId="11" pageNumber="346" rank="species">new combination</taxonomicNameLabel>
. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae.
</paragraph>
</caption>
<paragraph id="951559422C47CE21FFBBC7339C36F843" blockId="11.[113,753,1853,1973]" pageId="11" pageNumber="346">
Third maxilliped with ischium relatively short, subquadrate, length 1.2× width; merus quadrate, anterolateral angle gently auriculiform. Exopod relatively stout, tip reaching to just before distal edge of merus (
<figureCitation id="0D9145C72C47CE21FE66C7939C71F843" box="[428,558,1949,1973]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="11" pageNumber="346">Figs. 6D, E</figureCitation>
,
<figureCitation id="0D9145C72C47CE21FDF0C7939C02F843" box="[570,605,1949,1973]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346">7B</figureCitation>
).
</paragraph>
<paragraph id="951559422C47CE21FFBBC7D39A09F843" blockId="11.[113,752,2013,2102]" lastBlockId="11.[813,1452,1853,1973]" pageId="11" pageNumber="346">
Male thoracic sternum surface relatively smooth; sternites 1 and 2 completely fused, lateral margins gently concave; sternites 3 and 4 fused with only very shallow median groove and lateral notches visible; suture between sternites 2 and 3 gently concave towards buccal cavity (
<figureCitation id="0D9145C72C47CE21FB25C7539B2EF883" box="[1263,1393,1885,1909]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="11" pageNumber="346">Figs. 6D, E</figureCitation>
,
<figureCitation id="0D9145C72C47CE21FAB4C7539BC0F883" box="[1406,1439,1885,1909]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346">7E</figureCitation>
). Tubercle of male pleonal locking mechanism on anterior third of somite 5 (
<figureCitation id="0D9145C72C47CE21FC3EC7939A16F843" box="[1012,1097,1949,1973]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="11" pageNumber="346">Fig. 6E</figureCitation>
).
</paragraph>
<paragraph id="951559422C47CE26FCE7C7D39CC1FE23" blockId="11.[813,1452,2013,2102]" lastBlockId="12.[136,775,189,470]" lastPageId="12" lastPageNumber="347" pageId="11" pageNumber="346">
Adult male chelipeds symmetrical, stout (
<figureCitation id="0D9145C72C47CE21FB30C7D39B0CF803" box="[1274,1363,2013,2037]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346">Fig. 7A</figureCitation>
). Basis-ischium fused but suture visible, flexor margin with low granules (
<figureCitation id="0D9145C72C47CE21FC56C8139DAAF7C3" box="[924,1013,2077,2101]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346">Fig. 7E</figureCitation>
). Merus short, surface slightly rugose; flexor and extensor margins with small granules; inner distal angle sharp visible but not with subdistal angle not elongate (
<figureCitation id="0D9145C72C40CE26FF5AC0F39EBBFEE3" box="[144,228,253,277]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7E</figureCitation>
). Carpus surface granulate; inner distal angle with low sharp tooth (
<figureCitation id="0D9145C72C40CE26FEF0C1139FD0FEC3" box="[314,399,285,309]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7A</figureCitation>
). Chela with dorsal surface densely covered with granules; inner surface almost smooth; margins of chela with scattered long and short setae, not dense or brush-like; fingers relatively slender, shorter than palm, pigmented brown along distal ⅔; occlusal margins with low teeth, forming small gape when closed (
<figureCitation id="0D9145C72C40CE26FDF4C1B39CCEFE23" box="[574,657,445,469]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="12" pageNumber="347">Fig. 6F</figureCitation>
).
</paragraph>
<paragraph id="951559422C40CE26FF42C1F39D58FCE3" blockId="12.[136,775,509,789]" pageId="12" pageNumber="347">
Ambulatory legs short, P4 longest, about 1.2× carapace width; P4 merus length about 2.22.4× height, less than half carapace width; outer surface slightly rugose to almost smooth; coxa rounded; merus short, extensor margin not cristate, surface of P25 gently convex, without ridge; P25 carpus smooth, with low submarginal ridge, lowest in P5; propodus laterally flattened, unarmed, relatively short in P24, shortest in P5; P24 dactylus falciform, unarmed, P5 dactylus shorter, tip slightly upturned; dactylo-propodal lock distinct (
<figureCitation id="0D9145C72C40CE26FD69C2F39CA4FCE3" box="[675,763,765,789]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="951559422C40CE26FF42C3339F84FC03" blockId="12.[136,775,829,1013]" pageId="12" pageNumber="347">
Male pleon triangular, all somites and telson free; somite 1 broadly rectangular, reaching base of P5 coxae; somite 2 transversely narrower than somite 1, trapezoidal; somites 36 trapezoidal, gradually decreasing in width. Telson wider than long, rounded-subtriangular, lateral margins weakly convex, longer than somite 6 (
<figureCitation id="0D9145C72C40CE26FEB3C3D39F91FC03" box="[377,462,989,1013]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="951559422C40CE26FF42C4139F25FB83" blockId="12.[136,775,1053,1141]" pageId="12" pageNumber="347">G1 strongly sinuous, distally gently curved outwards, tip mucronate; with short row of setae subdistally. G2 sigmoid, about ¼ length of G1.</paragraph>
<paragraph id="951559422C40CE26FF42C4939CF2FA83" blockId="12.[136,775,1181,1397]" pageId="12" pageNumber="347">Female pleon longitudinally ovate; with 6 somites and telson, all free; not completely covering thoracic sternum; somite 1 broadly subrectangular, just reaching base of P5 coxae; somite 2 transversely narrower than somite 1, trapezoidal; somite 3 widest, somite 4 as wide as somite 2; somites 36 trapezoidal, gradually decreasing in width; telson triangular with convex lateral margins, longer than somite 6.</paragraph>
<paragraph id="951559422C40CE26FF42C5939F30FA03" blockId="12.[136,775,1437,1525]" pageId="12" pageNumber="347">
Sternopleonal cavity shallow (
<figureCitation id="0D9145C72C40CE26FE0DC5939C48FA43" box="[455,535,1437,1461]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7F</figureCitation>
; vulva large, obliquely ovate, covering anterior ⅔ on sternite 6, without vulvar projections (
<figureCitation id="0D9145C72C40CE26FEDAC5D39F3BFA03" box="[272,356,1501,1525]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C40CE26FF42C6139D57F943" pageId="12" pageNumber="347" type="discussion">
<paragraph id="951559422C40CE26FF42C6139D57F943" blockId="12.[136,776,1565,1717]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FF42C6139EA8F9C3" bold="true" box="[136,247,1565,1589]" pageId="12" pageNumber="347">Remarks.</emphasis>
<taxonomicName id="52AA22C12C40CE26FF37C6139FADF9C3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[253,498,1565,1589]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="symmetrinuda">
<emphasis id="A7DE85502C40CE26FF37C6139FADF9C3" box="[253,498,1565,1589]" italics="true" pageId="12" pageNumber="347">Hosekia symmetrinuda</emphasis>
</taxonomicName>
is a reef species currently known only from
<collectingCountry id="EDBD19D22C40CE26FE8EC6339FD2F9A3" box="[324,397,1597,1621]" name="Samoa" pageId="12" pageNumber="347">Samoa</collectingCountry>
and southern
<collectingCountry id="EDBD19D22C40CE26FDEAC6309C3FF9A3" box="[544,608,1598,1621]" name="Japan" pageId="12" pageNumber="347">Japan</collectingCountry>
. The specimens from both locations are nearly identical and we are confident they are conspecific. Nothing else is known about its biology except that it occurs in the intertidal zone (
<bibRefCitation id="F13B24B32C40CE26FDF1C6939CA3F943" author="Maenosono T" box="[571,764,1693,1717]" pageId="12" pageNumber="347" pagination="19 - 44" refId="ref13491" refString="Maenosono T (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48: 19 - 44. [In Japanese with English abstract]" type="journal article" year="2019">Maenosono, 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,274 @@
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<mods:title id="A36049B5263BF5E77EEA304C07437F3C">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
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<mods:namePart id="7D81C47583789D4AF1E01F477F73B8CC">Shane T.</mods:namePart>
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<taxonomicName id="52AA22C12C4ACE2CFC0AC1739B1AFE63" ID-CoL="3W3WV" authority="Sakai, 1935 a" authorityName="Sakai" authorityYear="1935" box="[960,1349,381,405]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFC0AC1739B1AFE63" bold="true" box="[960,1349,381,405]" pageId="6" pageNumber="341">
<emphasis id="A7DE85502C4ACE2CFC0AC1739AF4FE63" bold="true" box="[960,1195,381,405]" italics="true" pageId="6" pageNumber="341">Lophoplax takakurai</emphasis>
<bibRefCitation id="F13B24B32C4ACE2CFB79C1739B1AFE63" author="Sakai T" box="[1203,1349,381,405]" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
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</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE2CFB8CC1939A9FFE43" box="[1094,1216,413,437]" pageId="6" pageNumber="341" type="description">
<paragraph id="951559422C4ACE2CFB8CC1939A9FFE43" blockId="6.[960,1349,381,437]" box="[1094,1216,413,437]" pageId="6" pageNumber="341">
(
<figureCitation id="0D9145C72C4ACE2CFB84C1939AC4FE43" box="[1102,1179,413,437]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Figs. 4</figureCitation>
,
<figureCitation id="0D9145C72C4ACE2CFB63C1939AE8FE43" box="[1193,1207,413,437]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE2CFC89C1D29B9CFD93" pageId="6" pageNumber="341" type="reference_group">
<paragraph id="951559422C4ACE2CFC89C1D29B9CFD93" blockId="6.[835,1475,476,613]" pageId="6" pageNumber="341">
<treatmentCitationGroup id="B5BA7E6C2C4ACE2CFC89C1D29B9CFD93" pageId="6" pageNumber="341">
<treatmentCitation id="140B7F532C4ACE2CFC89C1D29A9EFE07" author="Sakai T" box="[835,1217,476,498]" page="82" pageId="6" pageNumber="341" year="1935">
<taxonomicName id="52AA22C12C4ACE2CFC89C1D29A9EFE07" ID-CoL="3W3WV" authority="Sakai, 1935 a: 82" authorityName="Sakai" authorityPageNumber="82" authorityYear="1935" box="[835,1217,476,498]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFC89C1D29A50FE07" box="[835,1039,476,497]" italics="true" pageId="6" pageNumber="341">Lophoplax takakurai</emphasis>
<bibRefCitation id="F13B24B32C4ACE2CFBD2C1D29A9EFE07" author="Sakai T" box="[1048,1217,476,498]" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a: 82</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, text-fig. 15, pl. 7, fig. 2;
<treatmentCitation id="140B7F532C4ACE2CFCA3C1F79A40FDF8" author="Sakai T" box="[873,1055,505,527]" page="188" pageId="6" pageNumber="341" year="1935">
<bibRefCitation id="F13B24B32C4ACE2CFCA3C1F79A40FDF8" author="Sakai T" box="[873,1055,505,527]" pageId="6" pageNumber="341" refId="ref14541" refString="Sakai T (1935 b) Crabs of Japan. 66 Plates in Life colours with descriptions. Sanseido, Tokyo, 239 pp." type="book" year="1935">Sakai, 1935b: 188</bibRefCitation>
</treatmentCitation>
, frontispiece, fig. 2;
<treatmentCitation id="140B7F532C4ACE2CFB3BC1F79BC3FDF9" author="Sakai T" box="[1265,1436,505,527]" page="567" pageId="6" pageNumber="341" year="1939">
<bibRefCitation id="F13B24B32C4ACE2CFB3BC1F79BC3FDF9" author="Sakai T" box="[1265,1436,505,527]" pageId="6" pageNumber="341" pagination="42 - 111" refId="ref14566" refString="Sakai T (1939) Studies on the Crabs of Japan. Brachygnatha, Brachyrhyncha, IV. Yokendo, Tokyo, pp. 365 - 741 + 1 - 23 [index], pls. 42 - 111." type="book chapter" year="1939">Sakai, 1939: 567</bibRefCitation>
</treatmentCitation>
, pl. 67, fig. l;
<treatmentCitation id="140B7F532C4ACE2CFC05C21B9A29FDDD" author="Sakai T" box="[975,1142,533,555]" page="541" pageId="6" pageNumber="341" year="1976">
<bibRefCitation id="F13B24B32C4ACE2CFC05C21B9A29FDDD" author="Sakai T" box="[975,1142,533,555]" pageId="6" pageNumber="341" pagination="1 - 16" refId="ref14607" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. In three volumes; English Text, pp. xxix + 773 pp., Japanese Text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251. Kodansha Ltd., Tokyo." type="book chapter" year="1976">Sakai, 1976: 541</bibRefCitation>
</treatmentCitation>
pl. 191, fig. 4, text-fig. 288;
<treatmentCitation id="140B7F532C4ACE2CFA55C2189D9FFDB1" author="Ng PKL" page="79" pageId="6" pageNumber="341" year="1987">
<bibRefCitation id="F13B24B32C4ACE2CFA55C2189D9FFDB1" author="Ng PKL" pageId="6" pageNumber="341" pagination="69 - 111" refId="ref13788" refString="Ng PKL (1987) The IndoPacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). IndoMalayan Zoology, 1987, 4 (1): 69 - 111, pl. 1." type="journal article" year="1987">Ng, 1987: 79</bibRefCitation>
</treatmentCitation>
, 100;
<treatmentCitation id="140B7F532C4ACE2CFC31C23C9AFBFDB1" author="Takeda M" box="[1019,1188,562,584]" page="41" pageId="6" pageNumber="341" year="1993">
<bibRefCitation id="F13B24B32C4ACE2CFC31C23C9AFBFDB1" author="Takeda M" box="[1019,1188,562,584]" pageId="6" pageNumber="341" refId="ref14889" refString="Takeda M (1993) Catalogue of crab specimens (Crustacea: Decapoda: Brachyura). Part 1. National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda, 1993: 41</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C4ACE2CFB65C23D9B2DFDB1" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1199,1394,562,584]" page="143" pageId="6" pageNumber="341" year="2008">
<bibRefCitation id="F13B24B32C4ACE2CFB65C23D9B2DFDB1" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1199,1394,562,584]" pageId="6" pageNumber="341" pagination="1 - 286" refId="ref14245" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 143</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C4ACE2CFAB1C23C9A17FD93" author="Takeda M &amp; Komatsu H" page="176" pageId="6" pageNumber="341" year="2018">
<bibRefCitation id="F13B24B32C4ACE2CFAB1C23C9A17FD93" author="Takeda M &amp; Komatsu H" pageId="6" pageNumber="341" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018: 176</bibRefCitation>
</treatmentCitation>
, fig. 7;
<treatmentCitation id="140B7F532C4ACE2CFB58C2419BE0FD93" author="Takeda M &amp; Komatsu H" box="[1170,1471,591,613]" page="167" pageId="6" pageNumber="341" year="2023">
<bibRefCitation id="F13B24B32C4ACE2CFB58C2419BE0FD93" author="Takeda M &amp; Komatsu H" box="[1170,1471,591,613]" pageId="6" pageNumber="341" pagination="161 - 178" refId="ref14998" refString="Takeda M &amp; Komatsu H (2023) Updated checklist of the crabs (Crustacea: Decapoda: Brachyura) from the sea around the Ogasawara Islands, Japan. Bulletin of the National Museum of Nature and Science, Series A, 49 (4): 161 - 178." type="journal article" year="2023">Takeda &amp; Komatsu, 2023: 167</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE2CFC89C2839A17FCD3" pageId="6" pageNumber="341" type="materials_examined">
<paragraph id="951559422C4ACE2CFC89C2839A17FCD3" blockId="6.[835,1475,653,805]" pageId="6" pageNumber="341">
<emphasis id="A7DE85502C4ACE2CFC89C2839A73FD53" bold="true" box="[835,1068,653,677]" pageId="6" pageNumber="341">Material examined.</emphasis>
<materialsCitation id="25C2531F2C4ACE2CFBFCC2839A1BFCD3" collectingDate="2010-07-09" collectionCode="R, V" country="Japan" county="Ogasawara Islands" elevation="42" latitude="27.079334" location="Futami Bay" longLatPrecision="12" longitude="142.1955" municipality="Chichi-jima" pageId="6" pageNumber="341" specimenCount="1" specimenCount-male="1">
<specimenCount id="83AC92CB2C4ACE2CFBFCC2839ADCFD53" box="[1078,1155,653,677]" count="1" pageId="6" pageNumber="341" type="male">1 male</specimenCount>
(5.0 ×
<quantity id="5252F4A72C4ACE2CFB14C2839B68FD53" box="[1246,1335,653,677]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="341" unit="mm" value="4.2">4.2 mm</quantity>
) (NSMT-Cr S 1254),
<location id="90750F992C4ACE2CFC67C2A39A6DFD30" LSID="urn:lsid:plazi:treatment:1D03E8542C4ACE22FC0AC1739CF3F98B:90750F992C4ACE2CFC67C2A39A6DFD30" box="[941,1074,685,710]" country="Japan" county="Ogasawara Islands" latitude="27.079334" longLatPrecision="12" longitude="142.1955" municipality="Chichi-jima" name="Futami Bay" pageId="6" pageNumber="341">Futami Bay</location>
,
<collectingMunicipality id="7571C3382C4ACE2CFB8AC2A39A9EFD33" box="[1088,1217,685,709]" pageId="6" pageNumber="341">Chichi-jima</collectingMunicipality>
[= Island],
<collectingCounty id="7C7421CE2C4ACE2CFA81C2A39DC9FD13" pageId="6" pageNumber="341">Ogasawara Islands</collectingCounty>
,
<collectingCountry id="EDBD19D22C4ACE2CFC6DC2C09DB5FD13" box="[935,1002,718,741]" name="Japan" pageId="6" pageNumber="341">Japan</collectingCountry>
,
<geoCoordinate id="F09E3F852C4ACE2CFC30C2C29A24FD13" box="[1018,1147,716,741]" degrees="27" direction="north" minutes="04.77" orientation="latitude" pageId="6" pageNumber="341" precision="9" value="27.0795">27°04.77N</geoCoordinate>
<geoCoordinate id="F09E3F852C4ACE2CFB4DC2C29B4BFD13" box="[1159,1300,716,741]" degrees="142" direction="east" minutes="11.68" orientation="longitude" pageId="6" pageNumber="341" precision="9" value="142.19467">142°11.68E</geoCoordinate>
to
<geoCoordinate id="F09E3F852C4ACE2CFA8BC2C29B9DFD13" box="[1345,1474,716,741]" degrees="27" direction="north" minutes="04.76" orientation="latitude" pageId="6" pageNumber="341" precision="9" value="27.079334">27°04.76N</geoCoordinate>
<geoCoordinate id="F09E3F852C4ACE2CFC8EC2E29D93FCF3" box="[836,972,748,773]" degrees="142" direction="east" minutes="11.73" orientation="longitude" pageId="6" pageNumber="341" precision="9" value="142.1955">142°11.73E</geoCoordinate>
, 42.2
<quantity id="5252F4A72C4ACE2CFBDEC2E39A3DFCF3" box="[1044,1122,749,773]" metricMagnitude="1" metricUnit="m" metricValue="4.16" pageId="6" pageNumber="341" unit="m" value="41.6">
<elevation id="1E87BE712C4ACE2CFBDEC2E39A3DFCF3" box="[1044,1122,749,773]" metricMagnitude="1" metricUnit="m" metricValue="4.16" pageId="6" pageNumber="341" unit="m" value="41.6">41.6 m</elevation>
</quantity>
,
<collectionCode id="F3BBC1872C4ACE2CFBA5C2E39ADEFCF3" box="[1135,1153,749,773]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="341">R</collectionCode>
/
<collectionCode id="F3BBC1872C4ACE2CFB43C2E39AC3FCF3" box="[1161,1180,749,773]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="341" type="Museum">V</collectionCode>
Koyo, 2010 cruise, station KY-10-25,
<date id="E1147F822C4ACE2CFC0AC3039A1BFCD3" box="[960,1092,781,805]" pageId="6" pageNumber="341" value="2010-07-09">
<collectingDate id="F150866A2C4ACE2CFC0AC3039A1BFCD3" box="[960,1092,781,805]" pageId="6" pageNumber="341" value="2010-07-09">9 July 2010</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE2CFC89C3439AB9FAD3" pageId="6" pageNumber="341" type="diagnosis">
<paragraph id="951559422C4ACE2CFC89C3439AB9FAD3" blockId="6.[835,1475,845,1317]" pageId="6" pageNumber="341">
<emphasis id="A7DE85502C4ACE2CFC89C3439DE7FC93" bold="true" box="[835,952,845,869]" pageId="6" pageNumber="341">Diagnosis.</emphasis>
Carapace frontal margin gently convex, smooth, produced anteriorly, lateral lobe not visible (
<figureCitation id="0D9145C72C4ACE2CFAF2C3639BEBFC73" box="[1336,1460,877,901]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4A, B</figureCitation>
); epigastric areole barely visible; protogastric lobe low, with granular surface (
<figureCitation id="0D9145C72C4ACE2CFBCEC3A39A03FC33" box="[1028,1116,941,965]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4B</figureCitation>
); metabranchial region with low granules (
<figureCitation id="0D9145C72C4ACE2CFC7EC3C39A4FFC13" box="[948,1040,973,997]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4B</figureCitation>
); median lobe on posterior margin of epistome subtruncate in form (
<figureCitation id="0D9145C72C4ACE2CFB5AC3E39AB1FBF3" box="[1168,1262,1005,1029]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Figs. 4E</figureCitation>
,
<figureCitation id="0D9145C72C4ACE2CFB30C3E39B45FBF3" box="[1274,1306,1005,1029]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">5E</figureCitation>
). Distal margin of ischium of third maxilliped with row of closely set low granules (
<figureCitation id="0D9145C72C4ACE2CFC67C4239A5CFBB3" box="[941,1027,1069,1093]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4C</figureCitation>
). P25 merus extensor margin P25 with low proximal tooth (
<figureCitation id="0D9145C72C4ACE2CFBE4C4439AF5FB93" box="[1070,1194,1101,1125]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">Fig. 5F, G</figureCitation>
). Cheliped carpus dorsal surface with longitudinal areole separated by groove from swollen lateral margin (
<figureCitation id="0D9145C72C4ACE2CFB82C4839AE7FB53" box="[1096,1208,1165,1189]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4A, I</figureCitation>
); proximal basal part of male chela covered with strong tubercles (
<figureCitation id="0D9145C72C4ACE2CFAD1C4A39B2AFB33" box="[1307,1397,1197,1221]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4G</figureCitation>
). Male thoracic sternum anteriorly relatively narrower (
<figureCitation id="0D9145C72C4ACE2CFAF5C4C39BCFFB13" box="[1343,1424,1229,1253]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4F</figureCitation>
). G1 strongly sinuous, distal part of G1 gently recurved (
<figureCitation id="0D9145C72C4ACE2CFAB8C4E39D0FFAD3" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">Fig. 5I, J</figureCitation>
). G2 sigmoid, about ¼ length of G1.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE22FC89C5439CF3F98B" lastPageId="8" lastPageNumber="343" pageId="6" pageNumber="341" type="discussion">
<paragraph id="951559422C4ACE2CFC89C5439A1EF850" blockId="6.[835,1475,1357,1958]" pageId="6" pageNumber="341">
<emphasis id="A7DE85502C4ACE2CFC89C5439DEDFA93" bold="true" box="[835,946,1357,1381]" pageId="6" pageNumber="341">Remarks.</emphasis>
<taxonomicName id="52AA22C12C4ACE2CFC70C5439AC5FA93" authorityName="Sakai" authorityYear="1935" box="[954,1178,1357,1381]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFC70C5439AC5FA93" box="[954,1178,1357,1381]" italics="true" pageId="6" pageNumber="341">Lophoplax takakurai</emphasis>
</taxonomicName>
was described on the basis of a female (11.0 × 9.0 mm) collected off Yoshihama, west of Sagami Bay, and later reported from off Mikawa Bay (south of Nagoya) (
<bibRefCitation id="F13B24B32C4ACE2CFBD1C5A39AF4FA33" author="Sakai T" box="[1051,1195,1453,1477]" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
, b, 1976).
<bibRefCitation id="F13B24B32C4ACE2CFAE9C5A39B9DFA33" author="Takeda M" box="[1315,1474,1453,1477]" pageId="6" pageNumber="341" refId="ref14889" refString="Takeda M (1993) Catalogue of crab specimens (Crustacea: Decapoda: Brachyura). Part 1. National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda (1993)</bibRefCitation>
subsequently recorded the species from Shibushi Bay in
<collectingRegion id="576E97A02C4ACE2CFC8EC5E39D9FF9F3" box="[836,960,1517,1541]" country="Japan" name="Kagoshima" pageId="6" pageNumber="341">Kagoshima</collectingRegion>
, with
<bibRefCitation id="F13B24B32C4ACE2CFC37C5E39B4FF9F3" author="Takeda M &amp; Komatsu H" box="[1021,1296,1517,1541]" pageId="6" pageNumber="341" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu (2018</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4ACE2CFAD3C5E39B0FF9F3" author="Takeda M &amp; Komatsu H" box="[1305,1360,1517,1541]" pageId="6" pageNumber="341" pagination="161 - 178" refId="ref14998" refString="Takeda M &amp; Komatsu H (2023) Updated checklist of the crabs (Crustacea: Decapoda: Brachyura) from the sea around the Ogasawara Islands, Japan. Bulletin of the National Museum of Nature and Science, Series A, 49 (4): 161 - 178." type="journal article" year="2023">2023</bibRefCitation>
) recording it from the Ogasawara Islands. The whereabouts of the type specimen is not known but appears to be lost. The present specimen matches the type description and figures of
<bibRefCitation id="F13B24B32C4ACE2CFA4DC6439DD0F973" author="Sakai T" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai (1935a</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4ACE2CFC50C6639D8DF973" author="Sakai T" box="[922,978,1645,1669]" pageId="6" pageNumber="341" pagination="42 - 111" refId="ref14566" refString="Sakai T (1939) Studies on the Crabs of Japan. Brachygnatha, Brachyrhyncha, IV. Yokendo, Tokyo, pp. 365 - 741 + 1 - 23 [index], pls. 42 - 111." type="book chapter" year="1939">1939</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4ACE2CFC17C6639A4BF973" author="Sakai T" box="[989,1044,1645,1669]" pageId="6" pageNumber="341" pagination="1 - 16" refId="ref14607" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. In three volumes; English Text, pp. xxix + 773 pp., Japanese Text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251. Kodansha Ltd., Tokyo." type="book chapter" year="1976">1976</bibRefCitation>
) well, especially in the general carapace features and proportions of the ambulatory legs. Significantly,
<bibRefCitation id="F13B24B32C4ACE2CFC89C6A39D84F933" author="Sakai T" box="[835,987,1709,1733]" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai (1935a</bibRefCitation>
: text-fig. 15c; present
<figureCitation id="0D9145C72C4ACE2CFB15C6A39B6AF933" box="[1247,1333,1709,1733]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">Fig. 5B</figureCitation>
) figured the third maxilliped showing a row of closely set granules on the distal margin of the ischium, a character also distinct in the present male specimen (
<figureCitation id="0D9145C72C4ACE2CFB57C7039AA9F8D3" box="[1181,1270,1805,1829]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4C</figureCitation>
). As such, we are confident of its identity. For the moment,
<taxonomicName id="52AA22C12C4ACE2CFADCC7209BC4F8B3" authorityName="Sakai" authorityYear="1935" box="[1302,1435,1837,1861]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFADCC7209BC4F8B3" box="[1302,1435,1837,1861]" italics="true" pageId="6" pageNumber="341">L. takakurai</emphasis>
</taxonomicName>
, as defined by
<bibRefCitation id="F13B24B32C4ACE2CFC73C7439A89F893" author="Takeda M &amp; Komatsu H" box="[953,1238,1869,1893]" pageId="6" pageNumber="341" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu (2018)</bibRefCitation>
and the present study, is clearly recognisable, so a
<typeStatus id="4A11E7E02C4ACE2CFB4BC7609A89F870" box="[1153,1238,1902,1926]" pageId="6" pageNumber="341" type="neotype">neotype</typeStatus>
designation does not appear to be necessary.
</paragraph>
<paragraph id="951559422C4ACE22FC89C7C39C82F9EB" blockId="6.[835,1475,1997,2085]" lastBlockId="8.[136,775,1477,1565]" lastPageId="8" lastPageNumber="343" pageId="6" pageNumber="341">
Although
<taxonomicName id="52AA22C12C4ACE2CFC78C7C09A68F813" authorityName="Sakai" authorityYear="1935" box="[946,1079,1997,2021]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFC78C7C09A68F813" box="[946,1079,1997,2021]" italics="true" pageId="6" pageNumber="341">L. takakurai</emphasis>
</taxonomicName>
differs markedly from
<taxonomicName id="52AA22C12C4ACE2CFAF7C7C09B9DF813" authorityName="Tesch" authorityYear="1918" box="[1341,1474,1997,2021]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4ACE2CFAF7C7C09B9DF813" box="[1341,1474,1997,2021]" italics="true" pageId="6" pageNumber="341">L. bicristata</emphasis>
</taxonomicName>
in many characters of the carapace and pereopods, both species have generally similar carapace shape and features. In addition, both species have a stouter G1 (
<figureCitation id="0D9145C72C44CE22FD9DC5CB9C98FA2B" box="[599,711,1477,1501]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="8" pageNumber="343">Figs. 3FI</figureCitation>
,
<figureCitation id="0D9145C72C44CE22FD1BC5CB9CA0FA2B" box="[721,767,1477,1501]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="8" pageNumber="343">5I, J</figureCitation>
) compared to more typical pilumnids. It is therefore reasonable to retain both species in
<taxonomicName id="52AA22C12C44CE22FE5CC60B9C57F9EB" authorityName="Tesch" authorityYear="1918" box="[406,520,1541,1565]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C44CE22FE5CC60B9C57F9EB" box="[406,520,1541,1565]" italics="true" pageId="8" pageNumber="343">Lophoplax</emphasis>
</taxonomicName>
for the time being.
</paragraph>
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<paragraph id="951559422C4BCE2DFFBBC7FD9D55F7B5" blockId="7.[113,1452,2035,2115]" pageId="7" pageNumber="342">
Fig. 4.
<taxonomicName id="52AA22C12C4BCE2DFF7CC7FD9C5BF7FF" authority="Sakai, 1935 a" authorityName="Sakai" authorityYear="1935" box="[182,516,2035,2057]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="342" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4BCE2DFF7CC7FD9F20F7FE" box="[182,383,2035,2056]" italics="true" pageId="7" pageNumber="342">Lophoplax takakurai</emphasis>
<bibRefCitation id="F13B24B32C4BCE2DFE4FC7FD9C5BF7FF" author="Sakai T" box="[389,516,2035,2057]" pageId="7" pageNumber="342" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
</taxonomicName>
, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped.
</paragraph>
</caption>
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<paragraph id="951559422C44CE22FF42C52C9D4FFA78" blockId="8.[136,1475,1314,1422]" pageId="8" pageNumber="343">
Fig. 5.
<taxonomicName id="52AA22C12C44CE22FF05C52C9C41FACE" authority="Sakai, 1935 a" authorityName="Sakai" authorityYear="1935" box="[207,542,1314,1336]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="343" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C44CE22FF05C52C9FC6FAC1" box="[207,409,1314,1335]" italics="true" pageId="8" pageNumber="343">Lophoplax takakurai</emphasis>
<bibRefCitation id="F13B24B32C44CE22FE55C52C9C41FACE" author="Sakai T" box="[415,542,1314,1336]" pageId="8" pageNumber="343" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
</taxonomicName>
. AD, holotype male (11.5 × 9.0 mm) (after
<bibRefCitation id="F13B24B32C44CE22FC1DC52C9A08FACE" author="Sakai T" box="[983,1111,1314,1336]" pageId="8" pageNumber="343" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm.
</paragraph>
</caption>
<paragraph id="951559422C44CE22FF42C64B9CF3F98B" blockId="8.[136,775,1605,1661]" pageId="8" pageNumber="343">
<taxonomicName id="52AA22C12C44CE22FF42C64B9F38F9AB" authorityName="Sakai" authorityYear="1935" box="[136,359,1605,1629]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="343" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C44CE22FF42C64B9F38F9AB" box="[136,359,1605,1629]" italics="true" pageId="8" pageNumber="343">Lophoplax takakurai</emphasis>
</taxonomicName>
occurs on muddy bottoms
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in depth (
<bibRefCitation id="F13B24B32C44CE22FF3BC66B9F29F98B" author="Sakai T" box="[241,374,1637,1661]" pageId="8" pageNumber="343" pagination="1 - 16" refId="ref14607" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. In three volumes; English Text, pp. xxix + 773 pp., Japanese Text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251. Kodansha Ltd., Tokyo." type="book chapter" year="1976">Sakai, 1976</bibRefCitation>
;
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).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,269 @@
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<mods:title id="0DC9D6634EDD27AC954062712655510B">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="26057BDD41DF3D6BC7EF301066A3EB6D">
<mods:date id="1D349A53DC6085D681CB838130CF0A04">2024</mods:date>
<mods:detail id="4C697793858B780D239A1EEB48627863" type="pubDate">
<mods:number id="CF01A903DFD37A3C82EA4B30E5D12062">2024-09-25</mods:number>
</mods:detail>
<mods:detail id="0C88E66C3918E4BA4381B274498D8555" type="volume">
<mods:number id="43834B4D50887F44BDA24E7B56803CBC">72</mods:number>
</mods:detail>
<mods:extent id="E3086DE17CEC8FAD3094064BAAF9A2A8" unit="page">
<mods:start id="A7D32E90FFC4D29E6DF3A2F8054B09C1">335</mods:start>
<mods:end id="98923269127A74E2D0C6613225AD3403">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="02DAA15E0D75961A78D6B8AFF4F5E84F">journal article</mods:classification>
<mods:identifier id="F29428A5BE11A5CF605C42BB05D756B7" type="DOI">10.26107/RBZ-2024-0027</mods:identifier>
<mods:identifier id="0370172153DDAD4AF64DC54359324013" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E6052FB4476BDC771F1003E02EB12527" type="Zenodo-Dep">14682761</mods:identifier>
<mods:identifier id="DAB7FC3E68B57E4B50938A8FC35F0084" type="ZooBank">4BFA36EF-2F2A-4162-8810-5E0414E5DE55</mods:identifier>
</mods:mods>
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<subSubSection id="DDB00AC92C4DCE2BFC23C5939AB0FA43" box="[1001,1263,1437,1461]" pageId="1" pageNumber="336" type="nomenclature">
<paragraph id="951559422C4DCE2BFC23C5939AB0FA43" blockId="1.[1001,1263,1437,1461]" box="[1001,1263,1437,1461]" pageId="1" pageNumber="336">
<heading id="CE5DEE2E2C4DCE2BFC23C5939AB0FA43" bold="true" box="[1001,1263,1437,1461]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="336" reason="2">
<taxonomicName id="52AA22C12C4DCE2BFC23C5939AB0FA43" ID-CoL="7P7TN" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[1001,1263,1437,1461]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4DCE2BFC23C5939AB0FA43" bold="true" box="[1001,1263,1437,1461]" pageId="1" pageNumber="336">
<emphasis id="A7DE85502C4DCE2BFC23C5939A3FFA43" bold="true" box="[1001,1120,1437,1461]" italics="true" pageId="1" pageNumber="336">Lophoplax</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFBA2C5939AB0FA43" author="Tesch JJ" box="[1128,1263,1437,1461]" pageId="1" pageNumber="336" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4DCE2BFCE7C5D29B09F9DD" pageId="1" pageNumber="336" type="reference_group">
<paragraph id="951559422C4DCE2BFCE7C5D29B09F9DD" blockId="1.[813,1452,1500,1579]" pageId="1" pageNumber="336">
<treatmentCitationGroup id="B5BA7E6C2C4DCE2BFCE7C5D29B09F9DD" pageId="1" pageNumber="336">
<treatmentCitation id="140B7F532C4DCE2BFCE7C5D29A1CFA04" author="Tesch JJ" box="[813,1091,1500,1522]" page="196" pageId="1" pageNumber="336" year="1918">
<taxonomicName id="52AA22C12C4DCE2BFCE7C5D29A1CFA04" ID-CoL="7P7TN" authority="Tesch, 1918: 196" authorityName="Tesch" authorityPageNumber="196" authorityYear="1918" box="[813,1091,1500,1522]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4DCE2BFCE7C5D29DCBFA07" box="[813,916,1500,1521]" italics="true" pageId="1" pageNumber="336">Lophoplax</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFC51C5D29A1CFA04" author="Tesch JJ" box="[923,1091,1500,1522]" pageId="1" pageNumber="336" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918: 196</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(part);
<treatmentCitation id="140B7F532C4DCE2BFB44C5D29B45FA07" author="Ng PKL" box="[1166,1306,1500,1521]" page="100" pageId="1" pageNumber="336" year="1987">
<bibRefCitation id="F13B24B32C4DCE2BFB44C5D29B45FA07" author="Ng PKL" box="[1166,1306,1500,1521]" pageId="1" pageNumber="336" pagination="69 - 111" refId="ref13788" refString="Ng PKL (1987) The IndoPacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). IndoMalayan Zoology, 1987, 4 (1): 69 - 111, pl. 1." type="journal article" year="1987">Ng, 1987: 100</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation id="140B7F532C4DCE2BFAAFC5D29DAEF9F8" author="Trivedi J &amp; Patel K &amp; Mitra S &amp; Ng PKL" page="595" pageId="1" pageNumber="336" year="2022">
<bibRefCitation id="F13B24B32C4DCE2BFAAFC5D29DAEF9F8" author="Trivedi J &amp; Patel K &amp; Mitra S &amp; Ng PKL" pageId="1" pageNumber="336" pagination="595 - 600" refId="ref15186" refString="Trivedi J, Patel K, Mitra S &amp; Ng PKL (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4): 595 - 600." type="journal article" year="2022">Trivedi et al., 2022: 595</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation id="140B7F532C4DCE2BFBF1C5F79B14F9F9" author="Poore GCB &amp; Ahyong ST" box="[1083,1355,1529,1551]" page="667" pageId="1" pageNumber="336" year="2023">
<bibRefCitation id="F13B24B32C4DCE2BFBF1C5F79B14F9F9" author="Poore GCB &amp; Ahyong ST" box="[1083,1355,1529,1551]" pageId="1" pageNumber="336" refId="ref14446" refString="Poore GCB &amp; Ahyong ST (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Collingwood, Australia, xii + 890 pp." type="book" year="2023">Poore &amp; Ahyong 2023: 667</bibRefCitation>
</treatmentCitation>
, 670, 680 (part), fig. 14.112u;
<treatmentCitation id="140B7F532C4DCE2BFBD3C6189B4DF9DD" author="Ng PKL &amp; Rahayu DL" box="[1049,1298,1557,1579]" page="429" pageId="1" pageNumber="336" year="2023">
<bibRefCitation id="F13B24B32C4DCE2BFBD3C6189B4DF9DD" author="Ng PKL &amp; Rahayu DL" box="[1049,1298,1557,1579]" pageId="1" pageNumber="336" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu, 2023: 429</bibRefCitation>
</treatmentCitation>
(part).
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</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4DCE2BFCE7C65A9A70F91D" pageId="1" pageNumber="336" type="type_taxon">
<paragraph id="951559422C4DCE2BFCE7C65A9A85F97A" blockId="1.[813,1451,1619,1676]" pageId="1" pageNumber="336">
<emphasis id="A7DE85502C4DCE2BFCE7C65A9D9DF99D" bold="true" box="[813,962,1619,1644]" pageId="1" pageNumber="336">
<typeStatus id="4A11E7E02C4DCE2BFCE7C65A9D38F99A" box="[813,871,1620,1644]" pageId="1" pageNumber="336">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="52AA22C12C4DCE2BFC0DC65A9B71F99A" ID-CoL="3W3WR" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[967,1326,1620,1644]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4DCE2BFC0DC65A9AFAF99A" box="[967,1189,1620,1644]" italics="true" pageId="1" pageNumber="336">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFB60C65A9B71F99A" author="Tesch JJ" box="[1194,1326,1620,1644]" pageId="1" pageNumber="336" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, by original designation. Gender of genus feminine.
</paragraph>
<paragraph id="951559422C4DCE2BFCE7C6BA9A70F91D" blockId="1.[813,1452,1715,1772]" pageId="1" pageNumber="336">
<emphasis id="A7DE85502C4DCE2BFCE7C6BA9DA7F93D" bold="true" box="[813,1016,1715,1740]" pageId="1" pageNumber="336">Included species.</emphasis>
<taxonomicName id="52AA22C12C4DCE2BFBC9C6BA9BD9F93A" ID-CoL="3W3WR" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[1027,1414,1716,1740]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4DCE2BFBC9C6BA9AB2F93A" box="[1027,1261,1716,1740]" italics="true" pageId="1" pageNumber="336">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFB32C6BA9BD9F93A" author="Tesch JJ" box="[1272,1414,1716,1740]" pageId="1" pageNumber="336" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="52AA22C12C4DCE2BFA5CC6BA9A75F91D" ID-CoL="3W3WV" authority="Sakai, 1935 a" authorityName="Sakai" authorityYear="1935" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4DCE2BFA5CC6BA9DCCF91A" italics="true" pageId="1" pageNumber="336">L. takakurai</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFC56C6DA9A75F91D" author="Sakai T" box="[924,1066,1747,1772]" pageId="1" pageNumber="336" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4DCE29FCE7C71D9F69FC63" lastPageId="3" lastPageNumber="338" pageId="1" pageNumber="336" type="diagnosis">
<paragraph id="951559422C4DCE29FCE7C71D9F69FC63" blockId="1.[812,1452,1811,2092]" lastBlockId="3.[113,754,189,917]" lastPageId="3" lastPageNumber="338" pageId="1" pageNumber="336">
<emphasis id="A7DE85502C4DCE2BFCE7C71D9DFAF8DD" bold="true" box="[813,933,1811,1835]" pageId="1" pageNumber="336">Diagnosis.</emphasis>
Carapace subquadrate; only protogastric and intestinal areoles visible; epigastric region gently convex, smooth but setose, not areolate; cardiac region raised but not areolate, granulate and setose; hepatic areole absent, with only tubercle present; intestinal region forming small areole, adjacent areas slightly raised, granulate and setose, no visible intestino-branchial areole; surface adjacent to posterior carapace margin gently concave, not raised, not forming ridge, no transverse channel visible; orbit clearly visible in dorsal view, supraorbital margin breaking general carapace outline, forming distinct concavity; eyes freely movable, ocular peduncle relatively long; lateral lobe of frontal margin low, indistinct; anterolateral margin with 4 teeth (including external orbital tooth), with 2 tubercles on posterolateral margin; posterolateral margins gently converging towards posterior carapace margin. Endostome not obliquely recessed posteriorly into buccal cavern, subparallel to adjacent pterygostomial surface, low, short endostomial ridges present. Posterior margin of epistome with distinct median lobe, separated from lateral lobes by distinct fissure. Third maxilliped ischium subquadrate, length about 1.2× width. Adult male chelipeds slightly asymmetrical; dorsal surface of carpus with quadrate areole with raised margins; chela relatively enlarged with external median surface smooth. Ambulatory legs elongate, slender, P4 longest, about 1.8× carapace width; merus without obvious crest on extensor margin, instead P24 lined with low, broad well-spaced setose prominences or low tooth; P5 extensor margin smooth, outer surface smooth, without medio-longitudinal swelling; P4 merus length about 5× height, distinctly longer than half carapace width. G1 sinuous, relatively stout, tip strongly recurved.
</paragraph>
</subSubSection>
<caption id="C1D509CA2C4ECE28FF4DC7939FA0F7B4" ID-DOI="http://doi.org/10.5281/zenodo.14682763" ID-Zenodo-Dep="14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="2" pageNumber="337" startId="2.[135,170,1949,1971]" targetBox="[176,1433,189,1933]" targetPageId="2" targetType="figure">
<paragraph id="951559422C4ECE28FF4DC7939FA0F7B4" blockId="2.[135,1474,1948,2115]" pageId="2" pageNumber="337">
Fig. 1. Terminology of carapace structures (based on
<taxonomicName id="52AA22C12C4ECE28FD5AC7939A47F844" authority="(Stimpson, 1858))" authorityName="Stimpson" authorityYear="1858" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[656,1048,1948,1971]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="sculpta">
<emphasis id="A7DE85502C4ECE28FD5AC7939D00F844" box="[656,863,1949,1970]" italics="true" pageId="2" pageNumber="337">Myopilumnus sculpta</emphasis>
(
<bibRefCitation id="F13B24B32C4ECE28FCA6C7939A56F844" author="Stimpson W" box="[876,1033,1948,1971]" pageId="2" pageNumber="337" pagination="93 - 110" refId="ref14833" refString="Stimpson W (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10: 93 - 110 [39 - 56]." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
))
</taxonomicName>
. A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14.
</paragraph>
</caption>
<subSubSection id="DDB00AC92C4FCE29FFBBC3B39A6CFD63" pageId="3" pageNumber="338" type="discussion">
<paragraph id="951559422C4FCE29FFBBC3B39F27F8A3" blockId="3.[113,754,957,1877]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FFBBC3B39EBFFC23" bold="true" box="[113,224,957,981]" pageId="3" pageNumber="338">Remarks.</emphasis>
<taxonomicName id="52AA22C12C4FCE29FF2CC3B39F07FC23" authorityName="Tesch" authorityYear="1918" box="[230,344,957,981]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="A7DE85502C4FCE29FF2CC3B39F07FC23" box="[230,344,957,981]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCED382B2C4FCE29FE95C3B09FBBFC23" box="[351,484,957,981]" pageId="3" pageNumber="338" sensu="stricto">sensu stricto</taxonomicNameLabel>
is distinct in its carapace ornamentation and the length and slenderness of the ambulatory legs.
<taxonomicName id="52AA22C12C4FCE29FEE3C3F39FC4FBE3" authorityName="Tesch" authorityYear="1918" box="[297,411,1021,1045]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FEE3C3F39FC4FBE3" box="[297,411,1021,1045]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
is separable from
<taxonomicName id="52AA22C12C4FCE29FD95C3F39CAFFBE3" authorityName="Deb" authorityYear="1989" box="[607,752,1021,1045]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FD95C3F39CAFFBE3" box="[607,752,1021,1045]" italics="true" pageId="3" pageNumber="338">Myopilumnus</emphasis>
</taxonomicName>
and
<taxonomicName id="52AA22C12C4FCE29FF68C4139EA5FBC3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[162,250,1053,1077]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C4FCE29FF68C4139EA5FBC3" box="[162,250,1053,1077]" italics="true" pageId="3" pageNumber="338">Hosekia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C4FCE29FEC6C4109FDBFBC0" box="[268,388,1054,1078]" pageId="3" pageNumber="338" rank="genus">new genus</taxonomicNameLabel>
, in the following characters: the hepatic surface of the carapace is ornamented only with a tubercle (
<figureCitation id="0D9145C72C4FCE29FF13C4539F6EFB83" box="[217,305,1117,1141]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A</figureCitation>
) (versus with a large, prominent areole; cf.
<figureCitation id="0D9145C72C4FCE29FF5EC4739EADFB63" box="[148,242,1149,1173]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FF37C4739F1FFB63" box="[253,320,1149,1173]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="3" pageNumber="338">7C, D</figureCitation>
); the intestino-branchial areole is absent with only a low intestinal areole visible (
<figureCitation id="0D9145C72C4FCE29FDF1C4939CCDFB43" box="[571,658,1181,1205]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A</figureCitation>
) (versus intestino-branchial areole large and distinct; cf.
<figureCitation id="0D9145C72C4FCE29FD43C4B39CB3FB23" box="[649,748,1213,1237]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FFBBC4D39EECFB03" box="[113,179,1245,1269]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="3" pageNumber="338">7C, D</figureCitation>
); the surface adjacent to the posterior carapace margin is relatively wide, not forming a channel as no intestino-branchial areole is present (
<figureCitation id="0D9145C72C4FCE29FE6DC5139FA1FAC3" box="[423,510,1309,1333]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A</figureCitation>
) (versus with distinct channel between intestino-branchial areole and posterior carapace margin; cf.
<figureCitation id="0D9145C72C4FCE29FE93C5539FE6FA83" box="[345,441,1373,1397]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FE0DC5539C53FA83" box="[455,524,1373,1397]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="3" pageNumber="338">7C, D</figureCitation>
); and the absence of a dorsal crest on the ambulatory merus (
<figureCitation id="0D9145C72C4FCE29FDD4C5739CC3FA63" box="[542,668,1405,1429]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Figs. 2A, B</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FD6DC5739CB7FA63" box="[679,744,1405,1429]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">3C, D</figureCitation>
) (versus with prominent crest; cf.
<bibRefCitation id="F13B24B32C4FCE29FE14C5909C9FFA43" author="Ng PKL &amp; Rahayu DL" box="[478,704,1437,1462]" pageId="3" pageNumber="338" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu, 2023</bibRefCitation>
: fig. 7GL). The adult male chela is relatively more enlarged, with the external median surface almost smooth (
<figureCitation id="0D9145C72C4FCE29FD5FC5D39ED9F9E3" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2C, D</figureCitation>
) (versus chelae not enlarged and the outer surface of the chela densely covered with granules; cf.
<figureCitation id="0D9145C72C4FCE29FD94C6139CEEF9C3" box="[606,689,1565,1589]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Fig. 6F</figureCitation>
). The ambulatory legs are also comparatively slender (P4 merus length about 5× height) and more elongated (P4 length about 1.8× carapace width), with the merus surface lacking a medio-longitudinal swelling or ridge (
<figureCitation id="0D9145C72C4FCE29FDFBC6939CCAF943" box="[561,661,1693,1717]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Figs. 2A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FD6EC6939CB7F943" box="[676,744,1693,1717]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">3C, D</figureCitation>
) (versus median part swollen, P4 merus length 3.03.5× height; cf.
<bibRefCitation id="F13B24B32C4FCE29FF2FC6D09F9FF903" author="Ng PKL &amp; Rahayu DL" box="[229,448,1757,1782]" pageId="3" pageNumber="338" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu, 2023</bibRefCitation>
: fig. 7GL).
<taxonomicName id="52AA22C12C4FCE29FD83C6D39CE4F903" authorityName="Tesch" authorityYear="1918" box="[585,699,1757,1781]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FD83C6D39CE4F903" box="[585,699,1757,1781]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
, like
<taxonomicName id="52AA22C12C4FCE29FFB8C6F39F5CF8E3" authorityName="Deb" authorityYear="1989" box="[114,259,1789,1813]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FFB8C6F39F5CF8E3" box="[114,259,1789,1813]" italics="true" pageId="3" pageNumber="338">Myopilumnus</emphasis>
</taxonomicName>
, further differs from
<taxonomicName id="52AA22C12C4FCE29FE3AC6F39C1AF8E3" box="[496,581,1789,1813]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FE3AC6F39C1AF8E3" box="[496,581,1789,1813]" italics="true" pageId="3" pageNumber="338">Hosekia</emphasis>
</taxonomicName>
, in having four instead of three anterolateral carapace teeth (
<figureCitation id="0D9145C72C4FCE29FD94C7139CE0F8C3" box="[606,703,1821,1845]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FD06C7139EDBF8A3" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="3" pageNumber="338">7C, D</figureCitation>
versus
<figureCitation id="0D9145C72C4FCE29FF10C7339F66F8A3" box="[218,313,1853,1877]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="3" pageNumber="338">Figs. 1A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FE8DC7339F33F8A3" box="[327,364,1853,1877]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">2A</figureCitation>
).
</paragraph>
<paragraph id="951559422C4FCE29FFBBC7739D9FFEE3" blockId="3.[113,753,1917,2101]" lastBlockId="3.[813,1451,189,277]" pageId="3" pageNumber="338">
<taxonomicName id="52AA22C12C4FCE29FFBBC7739EB4F863" authorityName="Tesch" authorityYear="1918" box="[113,235,1917,1941]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FFBBC7739EB4F863" box="[113,235,1917,1941]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
superficially resembles
<taxonomicName id="52AA22C12C4FCE29FDDFC7739CF0F863" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[533,687,1917,1941]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C4FCE29FDDFC7739CF0F863" box="[533,687,1917,1941]" italics="true" pageId="3" pageNumber="338">Striacoeloma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C4FCE29FD08C7709EECF840" pageId="3" pageNumber="338" rank="genus">new genus</taxonomicNameLabel>
, in the slender ambulatory legs (albeit with P4 merus length about 5× height, versus 4× height) and in having blunt prominences along most of the lateral margins of the carapace, but is immediately distinguished by the position of the orbits: on the anterior margin and fully visible dorsally in
<taxonomicName id="52AA22C12C4FCE29FCE7C0B39DC0FF23" authorityName="Tesch" authorityYear="1918" box="[813,927,189,213]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FCE7C0B39DC0FF23" box="[813,927,189,213]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
, versus submarginal and scarcely visible dorsally in
<taxonomicName id="52AA22C12C4FCE29FC8DC0D39D8AFF03" box="[839,981,221,245]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FC8DC0D39D8AFF03" box="[839,981,221,245]" italics="true" pageId="3" pageNumber="338">Striacoeloma</emphasis>
</taxonomicName>
(resembling the condition in
<taxonomicName id="52AA22C12C4FCE29FACDC0D39DEBFEE3" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FACDC0D39BF4FF03" box="[1287,1451,221,245]" italics="true" pageId="3" pageNumber="338">Cryptocoeloma</emphasis>
<bibRefCitation id="F13B24B32C4FCE29FCE7C0F39DEBFEE3" author="Miers EJ" box="[813,948,253,277]" pageId="3" pageNumber="338" pagination="18 - 32" refId="ref13583" refString="Miers EJ (1884) Crustacea. In: Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 1882. Part I. The collections from Melanesia. London, British Museum (Natural History) pp. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="951559422C4FCE29FCE7C1339A6CFD63" blockId="3.[813,1452,317,661]" pageId="3" pageNumber="338">
<bibRefCitation id="F13B24B32C4FCE29FCE7C1339A7EFEA3" author="Serene R &amp; Soh CL" box="[813,1057,317,341]" pageId="3" pageNumber="338" pagination="1 - 37" refId="ref14789" refString="Serene R &amp; Soh CL (1976) Brachyura collected during the Thai- Danish Expedition (1966). Research Bulletin Phuket Marine Biological Center, 12: 1 - 37, figs. 1 - 28, pls. 1 - 7." type="journal article" year="1976">Serène &amp; Soh (1976: 4</bibRefCitation>
, 21) recorded a small male specimen of “?
<taxonomicName id="52AA22C12C4FCE29FCA7C1539A13FE83" authorityName="Sakai" authorityYear="1935" box="[877,1100,349,373]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4FCE29FCA7C1539A13FE83" box="[877,1100,349,373]" italics="true" pageId="3" pageNumber="338">Lophoplax takakurai</emphasis>
</taxonomicName>
” from the Andaman Sea stating only that “Like
<taxonomicName id="52AA22C12C4FCE29FC20C1739A09FE63" authorityName="Sakai" authorityYear="1935" box="[1002,1110,381,405]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4FCE29FC20C1739A09FE63" box="[1002,1110,381,405]" italics="true" pageId="3" pageNumber="338">takakurai</emphasis>
</taxonomicName>
the present specimen has no elevation, but it differs by its anterolateral teeth much less clearly separated one from the other”. On this basis, it seems unlikely to be
<taxonomicName id="52AA22C12C4FCE29FC06C1D39AABFE03" authorityName="Deb" authorityYear="1989" box="[972,1268,477,501]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="andamanicus">
<emphasis id="A7DE85502C4FCE29FC06C1D39AABFE03" box="[972,1268,477,501]" italics="true" pageId="3" pageNumber="338">Myopilumnus andamanicus</emphasis>
</taxonomicName>
, which was also described from the Andaman Sea (
<bibRefCitation id="F13B24B32C4FCE29FB09C1F39B1FFDE3" author="Deb M" box="[1219,1344,509,533]" pageId="3" pageNumber="338" pagination="113 - 116" refId="ref13417" refString="Deb M (1989) Myopilumnus adamanicus n. gen., n. sp., a xanthid crab from Andamans. Journal of the Andaman Science Association, 5 (2): 113 - 116." type="journal article" year="1989">Deb, 1989</bibRefCitation>
; see also
<bibRefCitation id="F13B24B32C4FCE29FCE7C2139A52FDC3" author="Trivedi J &amp; Patel K &amp; Mitra S &amp; Ng PKL" box="[813,1037,541,565]" pageId="3" pageNumber="338" pagination="595 - 600" refId="ref15186" refString="Trivedi J, Patel K, Mitra S &amp; Ng PKL (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4): 595 - 600." type="journal article" year="2022">Trivedi et al., 2022</bibRefCitation>
).
<bibRefCitation id="F13B24B32C4FCE29FBE9C2139B78FDC3" author="Serene R &amp; Soh CL" box="[1059,1319,541,565]" pageId="3" pageNumber="338" pagination="1 - 37" refId="ref14789" refString="Serene R &amp; Soh CL (1976) Brachyura collected during the Thai- Danish Expedition (1966). Research Bulletin Phuket Marine Biological Center, 12: 1 - 37, figs. 1 - 28, pls. 1 - 7." type="journal article" year="1976">Serène &amp; Sohs (1976)</bibRefCitation>
record was subsequently repeated in the checklists of the Thai fauna by
<bibRefCitation id="F13B24B32C4FCE29FC9BC2539A5BFD83" author="Naiyanetr P" box="[849,1028,605,629]" pageId="3" pageNumber="338" refId="ref13683" refString="Naiyanetr P (1980) Crustacean fauna of Thailand (Decapoda and Stomatopoda). Chulalongkorn University, Bangkok, Thailand, unnumbered pages + 73 pp." type="book" year="1980">Naiyanetr (1980</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4FCE29FBDBC2539A16FD83" author="Naiyanetr P" box="[1041,1097,605,629]" pageId="3" pageNumber="338" refId="ref13711" refString="Naiyanetr P (1998) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Environmental Policy and Planning, Bangkok, Thailand, 161 pp." type="book" year="1998">1998</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4FCE29FB9CC2539AD2FD83" author="Naiyanetr P" box="[1110,1165,605,629]" pageId="3" pageNumber="338" refId="ref13748" refString="Naiyanetr P (2007) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, Thailand, 196 pp." type="book" year="2007">2007</bibRefCitation>
) and
<bibRefCitation id="F13B24B32C4FCE29FB07C2509BF7FD83" author="Ng PKL &amp; Davie PJF" box="[1229,1448,605,630]" pageId="3" pageNumber="338" pagination="369 - 384" refId="ref14209" refString="Ng PKL &amp; Davie PJF (2002) A checklist of the brachyuran crabs of Phuket and western Thailand. Phuket Marine Biological Center Special Publication, 23 (2): 369 - 384." type="journal article" year="2002">Ng &amp; Davie (2002)</bibRefCitation>
; it requires confirmation.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,276 @@
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<emphasis id="A7DE85502C4FCE29FC7BC2B39B79FD23" bold="true" box="[945,1318,701,725]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FC7BC2B39AC8FD23" bold="true" box="[945,1175,701,725]" italics="true" pageId="3" pageNumber="338">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C4FCE29FB55C2B39B79FD23" author="Tesch JJ" box="[1183,1318,701,725]" pageId="3" pageNumber="338" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</emphasis>
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<paragraph id="951559422C4FCE29FBE5C2D39AF6FD03" blockId="3.[945,1318,701,757]" box="[1071,1193,733,757]" pageId="3" pageNumber="338">
(
<figureCitation id="0D9145C72C4FCE29FBFDC2D39ADAFD03" box="[1079,1157,733,757]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Figs. 2</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FB58C2D39AC0FD03" box="[1170,1183,733,757]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">3</figureCitation>
)
</paragraph>
</subSubSection>
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<emphasis id="A7DE85502C4FCE29FCE7C3139A52FCC3" bold="true" box="[813,1037,797,821]" pageId="3" pageNumber="338">Material examined.</emphasis>
<materialsCitation id="25C2531F2C4FCE29FBDFC3109A8CFC43" collectingDate="1899-12-16" collectingDateMax="1899-12-18" collectingDateMin="1899-12-16" collectionCode="RMNH-ZMA" collectorName="M. Weber" country="Indonesia" county="Siboga Expedition station" elevation="90" latitude="-5.6083336" location="Nuhu Jaan" longLatPrecision="130" longitude="132.92" municipality="Kei Islands" pageId="3" pageNumber="338" specimenCode="D103002" specimenCount="1" typeStatus="lectotype">
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(here designated):
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, male (5.8 ×
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from north point of
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, northwest of
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,
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260, coll.
<collectorName id="385F3C942C4FCE29FCA8C3909D8EFC43" box="[866,977,925,949]" pageId="3" pageNumber="338">M. Weber</collectorName>
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</date>
</materialsCitation>
.
<materialsCitation id="25C2531F2C4FCE29FB17C3939BF9FBE3" collectingDate="1899-06-10" collectionCode="RMNH-ZMA" collectorName="M. Weber" country="Indonesia" county="Siboga Expedition station" elevation="59" location="Borneo Bank" municipality="Strait of Makassar" pageId="3" pageNumber="338" specimenCode="D103001" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="4A11E7E02C4FCE29FB17C3939B31FC43" box="[1245,1390,925,949]" pageId="3" pageNumber="338" type="paralectotype">Paralectotype</typeStatus>
(here designated):
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, male (4.5 ×
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,
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,
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,
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77, coll. coll.
<collectorName id="385F3C942C4FCE29FB59C3F09B5BFBE3" box="[1171,1284,1021,1045]" pageId="3" pageNumber="338">M. Weber</collectorName>
,
<date id="E1147F822C4FCE29FADAC3F39BF9FBE3" box="[1296,1446,1021,1045]" pageId="3" pageNumber="338" value="1899-06-10">
<collectingDate id="F150866A2C4FCE29FADAC3F39BF9FBE3" box="[1296,1446,1021,1045]" pageId="3" pageNumber="338" value="1899-06-10">10 June 1899</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4FCE29FCE7C4339B6BF9E3" pageId="3" pageNumber="338" type="diagnosis">
<paragraph id="951559422C4FCE29FCE7C4339B6BF9E3" blockId="3.[811,1452,1085,1557]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FCE7C4339DC1FBA3" bold="true" box="[813,926,1085,1109]" pageId="3" pageNumber="338">Diagnosis.</emphasis>
Carapace frontal margin almost straight, granulate, not distinctly produced anteriorly, lateral lobe very low but just visible (
<figureCitation id="0D9145C72C4FCE29FC25C4739A2EFB63" box="[1007,1137,1149,1173]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A, B</figureCitation>
); epigastric areole low but visible; protogastric areolet distinct, smooth (
<figureCitation id="0D9145C72C4FCE29FAE9C4939BC1FB43" box="[1315,1438,1181,1205]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A, B</figureCitation>
); metabranchial region with prominent raised granules (
<figureCitation id="0D9145C72C4FCE29FA4AC4B39D0EFB03" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A</figureCitation>
); median lobe on posterior margin of epistome obtusely triangular (
<figureCitation id="0D9145C72C4FCE29FC65C4F39A54FAE3" box="[943,1035,1277,1301]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3B</figureCitation>
). Distal margin of ischium of third maxilliped smooth (
<figureCitation id="0D9145C72C4FCE29FBCBC5139A09FAC3" box="[1025,1110,1309,1333]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3A</figureCitation>
). Cheliped carpus dorsal surface with subquadrate areole with crenulate ridged margins (
<figureCitation id="0D9145C72C4FCE29FA4AC5339D2DFA83" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2AC</figureCitation>
); proximal basal part of male chela almost smooth or with very low tubercles (
<figureCitation id="0D9145C72C4FCE29FB96C5739AEBFA63" box="[1116,1204,1405,1429]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2D</figureCitation>
). P24 merus extensor margin distinctly crenulated, with low lobes. P5 merus unarmed (
<figureCitation id="0D9145C72C4FCE29FC6AC5B39A7FFA23" box="[928,1056,1469,1493]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3C, D</figureCitation>
). Male thoracic sternum anteriorly relatively wide (
<figureCitation id="0D9145C72C4FCE29FC16C5D39A6FFA03" box="[988,1072,1501,1525]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2E</figureCitation>
). G1 gently sinuous, distal strongly recurved (
<figureCitation id="0D9145C72C4FCE29FC51C5F39A5BF9E3" box="[923,1028,1533,1557]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3FI</figureCitation>
). G2 about ¼ length of G1.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4FCE2CFCE7C6339BEDFEA3" lastPageId="6" lastPageNumber="341" pageId="3" pageNumber="338" type="description">
<paragraph id="951559422C4FCE2FFCE7C6339A93F7E3" blockId="3.[813,1452,1597,2101]" lastBlockId="5.[813,1453,1725,2069]" lastPageId="5" lastPageNumber="340" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FCE7C6339DE8F9A3" bold="true" box="[813,951,1597,1621]" pageId="3" pageNumber="338">Description.</emphasis>
Carapace subquadrate; frontal and anterolateral margins with scattered tufts of short and long setae, not brush-like, not concealing margins; regions well demarcated with few areoles distinct: protogastric areole transversely ovate, distinct; epigastric region low, visible when denuded, not areolate; hepatic region with 1 tubercle, no areole visible; cardiac region raised, covered with granules, setose, not areolate, lateral parts with raised patches of granules, but no visible cardio-branchial areole; gastro-cardiac region depressed, with grooves visible but not deep (
<figureCitation id="0D9145C72C4FCE29FAEFC7539BC1F883" box="[1317,1438,1885,1909]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="3" pageNumber="338">Fig. 1A, B</figureCitation>
); intestinal region convex, forming rounded areole, adjacent surfaces gently raised, with small granules, setose, separated from cristate posterior carapace margin by wide space, no obvious channel (
<figureCitation id="0D9145C72C4FCE29FC3DC7D39A29F803" box="[1015,1142,2013,2037]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="3" pageNumber="338">Fig. 1A, B</figureCitation>
); meso- and metabranchial regions each with 1 gently raised area covered with granules; metabranchial region with tubercle just posterior to raised patch of granules (
<figureCitation id="0D9145C72C49CE2FFE84C6B39FF4F923" box="[334,427,1725,1749]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Subhepatic and suborbital regions slightly rugose to almost smooth; pterygostomial region with small granules. Frontal margin weakly bilobed, almost straight, with shallow median concavity, margins granulate; lateral lobe, small, low, just visible in dorsal view (
<figureCitation id="0D9145C72C49CE2FFFB3C7539E8FF883" box="[121,208,1885,1909]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Supraorbital margin short, sinuous, with distinct submedian lobe, without fissure or cleft (
<figureCitation id="0D9145C72C49CE2FFD8FC7739CC0F863" box="[581,671,1917,1941]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Orbit dorsal, supraorbital margin, eyes clearly visible in dorsal view; eyes freely movable, ocular peduncle relatively long, filling orbit (
<figureCitation id="0D9145C72C49CE2FFECAC7D39F06F803" box="[256,345,2013,2037]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Anterolateral margin with 4 small, sharp teeth (including external orbital tooth), external orbital tooth broader, next 3 teeth acutely triangular, tip slightly enlarged (
<figureCitation id="0D9145C72C49CE2FFC52C6D39A53F903" box="[920,1036,1757,1781]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="5" pageNumber="340">Fig. 2A, B</figureCitation>
). Posterolateral margin uneven, with 2 tubercles, each with smaller basal granules; margins gently converging towards posterior carapace margin (
<figureCitation id="0D9145C72C49CE2FFA8EC7139BC1F8C3" box="[1348,1438,1821,1845]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Antennules rectangular, relatively large; antennules folding laterally. Basal antennal article subquadrate; flagellum entering orbital hiatus. Epistome with median lobe of posterior margin obtusely triangular, no obvious fissure in frontal view; lateral parts sinuous with small cleft (
<figureCitation id="0D9145C72C49CE2FFA81C7B39BC1F823" box="[1355,1438,1981,2005]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="5" pageNumber="340">Fig. 3B</figureCitation>
). Endostome distinct, sloping gradually posteriorly into buccal cavern, endostomial ridges low, short.
</paragraph>
<caption id="C1D509CA2C48CE2EFF4DC7FD9B55F7B5" ID-DOI="http://doi.org/10.5281/zenodo.14682765" ID-Zenodo-Dep="14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="4" pageNumber="339" startId="4.[135,170,2035,2057]" targetBox="[162,1447,189,2020]" targetPageId="4" targetType="figure">
<paragraph id="951559422C48CE2EFF4DC7FD9B55F7B5" blockId="4.[135,1474,2035,2115]" pageId="4" pageNumber="339">
Fig. 2.
<taxonomicName id="52AA22C12C48CE2EFF04C7FD9C49F7FE" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[206,534,2035,2057]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C48CE2EFF04C7FD9FC8F7FE" box="[206,407,2035,2056]" italics="true" pageId="4" pageNumber="339">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C48CE2EFE57C7FD9C49F7FE" author="Tesch JJ" box="[413,534,2035,2057]" pageId="4" pageNumber="339" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon.
</paragraph>
</caption>
<caption id="C1D509CA2C49CE2FFFBBC6179C11F973" ID-DOI="http://doi.org/10.5281/zenodo.14682767" ID-Zenodo-Dep="14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="5" pageNumber="340" startId="5.[113,148,1561,1583]" targetBox="[115,1452,189,1546]" targetPageId="5" targetType="figure">
<paragraph id="951559422C49CE2FFFBBC6179C11F973" blockId="5.[113,1452,1561,1670]" pageId="5" pageNumber="340">
Fig. 3.
<taxonomicName id="52AA22C12C49CE2FFF76C6179C52F9D8" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[188,525,1561,1583]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="340" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C49CE2FFF76C6179FD6F9D8" box="[188,393,1561,1582]" italics="true" pageId="5" pageNumber="340">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C49CE2FFE58C6179C52F9D8" author="Tesch JJ" box="[402,525,1561,1583]" pageId="5" pageNumber="340" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm.
</paragraph>
</caption>
<paragraph id="951559422C4ACE2CFF42C0B39FC7FEA3" blockId="6.[136,776,189,341]" pageId="6" pageNumber="341">
Third maxilliped with ischium relatively short, subquadrate, length 1.2× width, with shallow median oblique sulcus; merus subquadrate, anterolateral angle gently auriculiform. Exopod relatively stout, tip reaching to just before distal edge of merus (
<figureCitation id="0D9145C72C4ACE2CFEFEC1339FD3FEA3" box="[308,396,317,341]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C1739F15FD63" blockId="6.[136,776,381,661]" pageId="6" pageNumber="341">
Male thoracic sternum surface relatively smooth, slightly rugose; sternites 1 and 2 completely fused, wide, lateral margins gently concave; sternites 3 and 4 fused with only very shallow median groove and lateral notches visible; suture between sternites 2 and 3 almost straight (
<figureCitation id="0D9145C72C4ACE2CFD6CC1F39CA4FDE3" box="[678,763,509,533]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="6" pageNumber="341">Fig. 2E</figureCitation>
). Sternite 8 clearly visible as subrectangular plate adjacent to pleonal somite 2 when pleon closed (
<figureCitation id="0D9145C72C4ACE2CFDF4C2339CCCFDA3" box="[574,659,573,597]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3E</figureCitation>
). Tubercle of male pleonal locking mechanism on anterior third of thoracic somite 5.
</paragraph>
<paragraph id="951559422C4ACE2CFF42C2B39FBFFB63" blockId="6.[136,776,701,1174]" pageId="6" pageNumber="341">
Adult male chelipeds almost symmetrical, stout (
<figureCitation id="0D9145C72C4ACE2CFD60C2B39EC3FD03" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1A, B</figureCitation>
). Basis-ischium fused but suture visible; extensor margins uneven but unarmed (
<figureCitation id="0D9145C72C4ACE2CFEBAC2F39F9CFCE3" box="[368,451,765,789]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1C</figureCitation>
). Merus short, surface slightly rugose; flexor and extensor margins with tubercles and granules; inner distal angle dentiform with subdistal angle tuberculate (
<figureCitation id="0D9145C72C4ACE2CFEC2C3539F07FC83" box="[264,344,861,885]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1C</figureCitation>
). Carpus surface granulate; dorsal surface with subquadrate areole with crenulate ridged margins; inner distal angle with low sharp tooth (
<figureCitation id="0D9145C72C4ACE2CFDC9C3939C24FC43" box="[515,635,925,949]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1AC</figureCitation>
). Chela with dorsal surface covered with granules; inner surface almost smooth; median and ventral surfaces almost smooth or with very low tubercles; margins of chela with scattered long and short setae, not dense or brush-like; fingers relatively slender, slightly shorter than palm, pigmented brown along distal half; occlusal margins with low teeth, forming small gape when closed (
<figureCitation id="0D9145C72C4ACE2CFE90C4739F8CFB63" box="[346,467,1149,1173]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1C, D</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C4B39CCAF9C3" blockId="6.[136,776,1213,1589]" pageId="6" pageNumber="341">
Ambulatory legs long, P4 longest, about 1.8× carapace width; P4 merus length about 5× height, distinctly longer than half carapace width; outer surface slightly rugose to almost smooth; coxa rounded; merus elongate, P24 extensor margin not cristate with low, broad, well-spaced setose prominences, P5 merus extensor margin smooth, fully setose; surface of P25 gently convex, without median swelling or ridge; P25 carpus relatively smooth, with very low submarginal ridge, lowest in P5; propodus laterally flattened, unarmed, relatively long in P24, shortest in P5; P24 dactylus falciform, unarmed, P5 dactylus shorter, slightly upturned; dactylo-propodal lock visible (
<figureCitation id="0D9145C72C4ACE2CFE1FC6139C69F9C3" box="[469,566,1565,1589]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="6" pageNumber="341">Figs. 2A</figureCitation>
,
<figureCitation id="0D9145C72C4ACE2CFD89C6139CD7F9C3" box="[579,648,1565,1589]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">3C, D</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C6539FD1F8A3" blockId="6.[136,776,1629,1877]" pageId="6" pageNumber="341">
Male pleon triangular, all somites and telson free; somite 1 broadly rectangular, reaching base of P5 coxae; somite 2 transversely much narrower than somite 1, trapezoidal; somite 3 broadly subtrapezoidal, widest, somites 46 trapezoidal, gradually decreasing in width with somite 6 almost rectangular; telson slightly wider than long, rounded, slightly narrowly distally, lateral margins convex, longer than somite 6 (
<figureCitation id="0D9145C72C4ACE2CFEE7C7339FDDF8A3" box="[301,386,1853,1877]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C7739F80F803" blockId="6.[136,775,1917,2037]" pageId="6" pageNumber="341">
G1 relatively stout, gently sinuous, distally tapering to narrowly rounded tip, strongly hooked inwards; subdistal surface with 2 long stout setae (
<figureCitation id="0D9145C72C4ACE2CFE32C7B39C38F823" box="[504,615,1981,2005]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3FI</figureCitation>
). G2 sigmoid, about ¼ length of G1 (
<figureCitation id="0D9145C72C4ACE2CFE4EC7D39F8CF803" box="[388,467,2013,2037]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3J</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C8139BEDFEA3" blockId="6.[136,356,2077,2101]" lastBlockId="6.[835,1475,189,341]" pageId="6" pageNumber="341">
Females not known.
<emphasis id="A7DE85502C4ACE2CFC89C0B39DEBFF23" bold="true" box="[835,948,189,213]" pageId="6" pageNumber="341">Remarks.</emphasis>
<taxonomicName id="52AA22C12C4ACE2CFC74C0B39AFBFF23" authorityName="Tesch" authorityYear="1918" box="[958,1188,189,213]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4ACE2CFC74C0B39AFBFF23" box="[958,1188,189,213]" italics="true" pageId="6" pageNumber="341">Lophoplax bicristata</emphasis>
</taxonomicName>
is only known from
<specimenCount id="83AC92CB2C4ACE2CFA53C0B09DDDFF03" count="2" pageId="6" pageNumber="341" type="male">two males</specimenCount>
<materialsCitation id="25C2531F2C4ACE2CFC43C0D39BEDFEA3" country="Indonesia" county="They" elevation="75" location="Kei Islands" municipality="Both" pageId="6" pageNumber="341" specimenCount="1">
collected from relatively near each other (
<location id="90750F992C4ACE2CFA82C0D39B9CFF03" LSID="urn:lsid:plazi:treatment:1D03E8542C4FCE2CFC7BC2B39BEDFEA3:90750F992C4ACE2CFA82C0D39B9CFF03" box="[1352,1475,221,245]" country="Indonesia" county="They" municipality="Both" name="Kei Islands" pageId="6" pageNumber="341">Kei Islands</location>
and
<location id="90750F992C4ACE2CFCBFC0F09A54FEE3" LSID="urn:lsid:plazi:treatment:1D03E8542C4FCE2CFC7BC2B39BEDFEA3:90750F992C4ACE2CFCBFC0F09A54FEE3" box="[885,1035,253,277]" country="Indonesia" county="They" municipality="Both" name="Borneo Bank" pageId="6" pageNumber="341">Borneo Bank</location>
,
<collectingCountry id="EDBD19D22C4ACE2CFBD3C0F39AD7FEE3" box="[1049,1160,253,277]" name="Indonesia" pageId="6" pageNumber="341">Indonesia</collectingCountry>
).
<collectingMunicipality id="7571C3382C4ACE2CFB57C0F39A8DFEE3" box="[1181,1234,253,277]" pageId="6" pageNumber="341">Both</collectingMunicipality>
are mature and were covered with silt and mud, suggesting they live on soft substrates.
<collectingCounty id="7C7421CE2C4ACE2CFC76C1339DACFEA3" box="[956,1011,317,341]" pageId="6" pageNumber="341">They</collectingCounty>
were collected from depths of
<quantity id="5252F4A72C4ACE2CFA81C1339BF1FEA3" box="[1355,1454,317,341]" metricMagnitude="1" metricUnit="m" metricValue="7.45" metricValueMax="9.0" metricValueMin="5.9" pageId="6" pageNumber="341" unit="m" value="74.5" valueMax="90.0" valueMin="59.0">
<elevation id="1E87BE712C4ACE2CFA81C1339BF1FEA3" box="[1355,1454,317,341]" metricMagnitude="1" metricUnit="m" metricValue="7.45" metricValueMax="9.0" metricValueMin="5.9" pageId="6" pageNumber="341" unit="m" value="74.5" valueMax="90.0" valueMin="59.0">5990 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="A1B8F92611A03078A44D8C8E7B95B730" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C5CCE39FC07C4939E8AFD03" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Striacoeloma tubur Ng, Ahyong &amp; T., 2024, new species" docType="treatment" docVersion="2" lastPageNumber="354" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="351" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5145B194804FD586073FFB4D19F5F19C">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
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<mods:namePart id="D536352F4CB79EF086AE89023B6DE3B0">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="5B0DF55B2562BD6B60609450F02AC0C2">Ahyong,</mods:namePart>
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<mods:namePart id="6F7779CF19FDB4E6F494FCBDB03954BB">Shane T.</mods:namePart>
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<mods:typeOfResource id="568E31DFE84423585AC11458B018024D">text</mods:typeOfResource>
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<mods:title id="7AB9B264C0448CEABBB42022129E0038">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="CB99D10FC30B08B91D181B76E52A56EB">
<mods:date id="29D9C97020A70A37330042A7AFAE19D5">2024</mods:date>
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<mods:number id="D9FA3795775B59EB6A40614EAC4A44FE">2024-09-25</mods:number>
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<treatment id="1D03E8542C5CCE39FC07C4939E8AFD03" LSID="urn:lsid:plazi:treatment:1D03E8542C5CCE39FC07C4939E8AFD03" httpUri="http://treatment.plazi.org/id/1D03E8542C5CCE39FC07C4939E8AFD03" lastPageId="19" lastPageNumber="354" pageId="16" pageNumber="351">
<subSubSection id="DDB00AC92C5CCE3AFC07C4939B66FB43" box="[973,1337,1181,1205]" pageId="16" pageNumber="351" type="nomenclature">
<paragraph id="951559422C5CCE3AFC07C4939B66FB43" blockId="16.[973,1337,1181,1237]" box="[973,1337,1181,1205]" pageId="16" pageNumber="351">
<heading id="CE5DEE2E2C5CCE3AFC07C4939B66FB43" bold="true" box="[973,1337,1181,1205]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="351" reason="2">
<emphasis id="A7DE85502C5CCE3AFC07C4939B66FB43" bold="true" box="[973,1337,1181,1205]" pageId="16" pageNumber="351">
<taxonomicName id="52AA22C12C5CCE3AFC07C4939AFBFB43" ID-CoL="CL3GY" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[973,1188,1181,1205]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="tubur" status="sp. nov.">
<emphasis id="A7DE85502C5CCE3AFC07C4939AFBFB43" bold="true" box="[973,1188,1181,1205]" italics="true" pageId="16" pageNumber="351">Striacoeloma tubur</emphasis>
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,
<taxonomicNameLabel id="BCED382B2C5CCE3AFB79C4939B66FB43" box="[1203,1337,1181,1205]" pageId="16" pageNumber="351" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="DDB00AC92C5CCE3AFBF3C4B39A92FB23" box="[1081,1229,1213,1237]" pageId="16" pageNumber="351" type="description">
<paragraph id="951559422C5CCE3AFBF3C4B39A92FB23" blockId="16.[973,1337,1181,1237]" box="[1081,1229,1213,1237]" pageId="16" pageNumber="351">
(
<figureCitation id="0D9145C72C5CCE3AFB8BC4B39AC3FB23" box="[1089,1180,1213,1237]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Figs. 10</figureCitation>
,
<figureCitation id="0D9145C72C5CCE3AFB63C4B39A9BFB23" box="[1193,1220,1213,1237]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="16" pageNumber="351">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFC89C4F39DC3FA83" pageId="16" pageNumber="351" type="materials_examined">
<paragraph id="951559422C5CCE3AFC89C4F39DC3FA83" blockId="16.[835,1474,1277,1398]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFC89C4F39A7EFAE3" bold="true" box="[835,1057,1277,1301]" pageId="16" pageNumber="351">Material examined.</emphasis>
<materialsCitation id="25C2531F2C5CCE3AFBEDC4F39DC3FA83" collectionCode="AM" collectorName="Cardno Pty Ltd." country="Australia" elevation="6" latitude="-12.588056" location="Middle Arm" longLatPrecision="21" longitude="130.87138" municipality="Darwin Harbour" pageId="16" pageNumber="351" specimenCode="P107758" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<typeStatus id="4A11E7E02C5CCE3AFBEDC4F39AD1FAE3" box="[1063,1166,1277,1301]" pageId="16" pageNumber="351" type="holotype">Holotype</typeStatus>
:
<collectionCode id="F3BBC1872C5CCE3AFB52C4F09A9CFAE0" box="[1176,1219,1278,1302]" country="Australia" name="Australian Museum" pageId="16" pageNumber="351" type="Museum">AM</collectionCode>
<specimenCode id="C50CF1392C5CCE3AFB03C4F39B75FAE3" box="[1225,1322,1277,1301]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="16" pageNumber="351" type="Museum">P107758</specimenCode>
, female (8.6 ×
<quantity id="5252F4A72C5CCE3AFC89C5139DC8FAC3" box="[835,919,1309,1333]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="16" pageNumber="351" unit="mm" value="6.3">6.3 mm</quantity>
),
<location id="90750F992C5CCE3AFC63C5139A6FFAC0" LSID="urn:lsid:plazi:treatment:1D03E8542C5CCE39FC07C4939E8AFD03:90750F992C5CCE3AFC63C5139A6FFAC0" box="[937,1072,1309,1334]" country="Australia" latitude="-12.588056" longLatPrecision="21" longitude="130.87138" municipality="Darwin Harbour" name="Middle Arm" pageId="16" pageNumber="351" stateProvince="Northern Territory">Middle Arm</location>
,
<collectingMunicipality id="7571C3382C5CCE3AFBF0C5139AB3FAC3" box="[1082,1260,1309,1333]" pageId="16" pageNumber="351">Darwin Harbour</collectingMunicipality>
,
<collectingRegion id="576E97A02C5CCE3AFB3FC5139BE1FAC3" box="[1269,1470,1309,1333]" country="Australia" name="Northern Territory" pageId="16" pageNumber="351">Northern Territory</collectingRegion>
,
<collectingCountry id="EDBD19D22C5CCE3AFC89C5339DF5FAA3" box="[835,938,1341,1365]" name="Australia" pageId="16" pageNumber="351">Australia</collectingCountry>
,
<geoCoordinate id="F09E3F852C5CCE3AFC73C5329A69FAA3" box="[953,1078,1340,1365]" degrees="12" direction="south" minutes="35" orientation="latitude" pageId="16" pageNumber="351" precision="15" seconds="17" value="-12.588056">12°3517″S</geoCoordinate>
<geoCoordinate id="F09E3F852C5CCE3AFB8BC5329A90FAA3" box="[1089,1231,1340,1365]" degrees="130" direction="east" minutes="52" orientation="longitude" pageId="16" pageNumber="351" precision="15" seconds="17" value="130.87138">130°5217″E</geoCoordinate>
,
<quantity id="5252F4A72C5CCE3AFB14C5339B77FAA3" box="[1246,1320,1341,1365]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="16" pageNumber="351" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">
<elevation id="1E87BE712C5CCE3AFB14C5339B77FAA3" box="[1246,1320,1341,1365]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="16" pageNumber="351" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">57 m</elevation>
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, coll.
<collectorName id="385F3C942C5CCE3AFAB8C5339DC3FA83" pageId="16" pageNumber="351">Cardno Pty Ltd.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFC89C5939A4AFA03" pageId="16" pageNumber="351" type="etymology">
<paragraph id="951559422C5CCE3AFC89C5939A4AFA03" blockId="16.[835,1474,1437,1525]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFC89C5939D9AFA43" bold="true" box="[835,965,1437,1461]" pageId="16" pageNumber="351">Etymology.</emphasis>
Named
<taxonomicName id="52AA22C12C5CCE3AFBEFC5939A00FA43" box="[1061,1119,1437,1461]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="tubur">
<emphasis id="A7DE85502C5CCE3AFBEFC5939A00FA43" box="[1061,1119,1437,1461]" italics="true" pageId="16" pageNumber="351">tubur</emphasis>
</taxonomicName>
for the tubercle-like protrusions on the extensor margins of the ambulatory legs; used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFC89C6139A0BF9C0" box="[835,1108,1565,1590]" pageId="16" pageNumber="351" type="diagnosis">
<paragraph id="951559422C5CCE3AFC89C6139A0BF9C0" blockId="16.[835,1108,1565,1590]" box="[835,1108,1565,1590]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFC89C6139DE7F9C3" bold="true" box="[835,952,1565,1589]" pageId="16" pageNumber="351">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE39FC89C6539F64FE03" lastPageId="19" lastPageNumber="354" pageId="16" pageNumber="351" type="description">
<paragraph id="951559422C5CCE38FC89C6539F8EF917" blockId="16.[835,1475,1629,2101]" lastBlockId="18.[136,776,1129,1761]" lastPageId="18" lastPageNumber="353" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFC89C6539AEEF983" bold="true" box="[835,1201,1629,1653]" pageId="16" pageNumber="351">
Description of
<typeStatus id="4A11E7E02C5CCE3AFC38C6539A0BF983" box="[1010,1108,1629,1653]" pageId="16" pageNumber="351" type="holotype">holotype</typeStatus>
female.
</emphasis>
Carapace subhexagonal; frontal and anterolateral margins with scattered tufts of short and long setae, not brush-like; surface appearing irregularly corrugated, partly eroded; gastric regions prominently raised, forming 2 transverse swellings extending to branchial region, with posterior one reaching to lateral margin; gastro-cardiac region depressed, with grooves visible; cardiac region with transverse swelling, extends almost to posterolateral margin (
<figureCitation id="0D9145C72C5CCE3AFC81C7539D91F883" box="[843,974,1885,1909]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10AC</figureCitation>
); surface adjacent to posterior carapace margin raised, with prominent wide rounded transverse ridge adjacent to the posterior carapace margin, forming channel extending to sub-branchial region (
<figureCitation id="0D9145C72C5CCE3AFBACC7B39AA8F823" box="[1126,1271,1981,2005]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10A, B</figureCitation>
). Subhepatic and suborbital regions slightly rugose; pterygostomial region with numerous fine granules (
<figureCitation id="0D9145C72C5CCE3AFB87C7F39AEEF7E3" box="[1101,1201,2045,2069]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10D</figureCitation>
). Frontal margin bilobed, almost straight, lined with granules, with shallow median notch, lateral lobe small but distinct, not visible in dorsal view (
<figureCitation id="0D9145C72C5ECE38FF5AC4879F48FB54" box="[144,279,1161,1186]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10AD</figureCitation>
). Supraorbital margin sinuous, with low lobe on posterior third, with 2 short fissures (
<figureCitation id="0D9145C72C5ECE38FDF3C4A79CE4FB34" box="[569,699,1193,1218]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10B, C</figureCitation>
). Orbit submarginal, supraorbital margin appearing almost confluent with general carapace outline, eyes partially visible in dorsal view; eyes scarcely movable, ocular peduncle short, filling orbit, scarcely movable (
<figureCitation id="0D9145C72C5ECE38FE5BC5279C4BFAB4" box="[401,532,1321,1346]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10C, D</figureCitation>
). Anterolateral margin with 4 distinct teeth (including external orbital tooth), first 2 broad, lobiform, with surface adjacent to them eroded, each with median oblique depression; last 2 dentiform (
<figureCitation id="0D9145C72C5ECE38FD16C5879E82FA34" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10A, B</figureCitation>
). Posterolateral margin with 2 distinct tubercles, margins distinctly converging towards posterior carapace margin (
<figureCitation id="0D9145C72C5ECE38FF22C5E79F2BF9F4" box="[232,372,1513,1538]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10A, B</figureCitation>
). Antennules rectangular, relatively large; antennules folding laterally (
<figureCitation id="0D9145C72C5ECE38FE33C6079C04F9D4" box="[505,603,1545,1570]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10D</figureCitation>
). Basal antennal article subquadrate; flagellum entering orbital hiatus (
<figureCitation id="0D9145C72C5ECE38FD11C6279EE5F994" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10D</figureCitation>
). Epistome with shallow biconcave posterior margin, median lobe subtruncate with short median fissue (
<figureCitation id="0D9145C72C5ECE38FD13C6679EE4F954" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10D</figureCitation>
). Endostome short, recessed posteriorly into buccal cavern oblique to plane of adjacent pterygostomial surface, endostomial ridges low, short.
</paragraph>
<caption id="C1D509CA2C5DCE3BFFBBC7AA9DB5F802" ID-DOI="http://doi.org/10.5281/zenodo.14682787" ID-Zenodo-Dep="14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="17" pageNumber="352" startId="17.[113,148,1956,1978]" targetBox="[113,1451,268,1941]" targetPageId="17" targetType="figure">
<paragraph id="951559422C5DCE3BFFBBC7AA9DB5F802" blockId="17.[113,1452,1956,2036]" pageId="17" pageNumber="352">
Fig. 10.
<taxonomicName id="52AA22C12C5DCE3BFF0CC7AA9FDDF84F" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[198,386,1956,1977]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="352" phylum="Arthropoda" rank="species" species="tubur" status="new genus and species">
<emphasis id="A7DE85502C5DCE3BFF0CC7AA9FDDF84F" box="[198,386,1956,1977]" italics="true" pageId="17" pageNumber="352">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5DCE3BFE5BC7AB9C2AF84C" box="[401,629,1956,1978]" pageId="17" pageNumber="352" rank="species">new genus and species</taxonomicNameLabel>
, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped.
</paragraph>
</caption>
<caption id="C1D509CA2C5ECE38FF4DC40E9B68FBC4" ID-DOI="http://doi.org/10.5281/zenodo.14682789" ID-Zenodo-Dep="14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353" startId="18.[135,170,1024,1046]" targetBox="[135,1473,189,1008]" targetPageId="18" targetType="figure">
<paragraph id="951559422C5ECE38FF4DC40E9B68FBC4" blockId="18.[135,1474,1023,1074]" pageId="18" pageNumber="353">
Fig. 11.
<taxonomicName id="52AA22C12C5ECE38FF16C40E9FC7FBE3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[220,408,1024,1045]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="353" phylum="Arthropoda" rank="species" species="tubur" status="new genus and species">
<emphasis id="A7DE85502C5ECE38FF16C40E9FC7FBE3" box="[220,408,1024,1045]" italics="true" pageId="18" pageNumber="353">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5ECE38FE6DC40E9CD4FBE0" box="[423,651,1024,1046]" pageId="18" pageNumber="353" rank="species">new genus and species</taxonomicNameLabel>
, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5.
</paragraph>
</caption>
<paragraph id="951559422C5ECE38FF42C7079D57F894" blockId="18.[136,776,1801,1890]" pageId="18" pageNumber="353">
Third maxilliped ischium short, quadrate, length 1.2× width; merus with anteroexternal angle distinctly auriculiform; exopod stout, tip reaching distal edge of merus (
<figureCitation id="0D9145C72C5ECE38FD53C7479CA3F894" box="[665,764,1865,1890]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10E</figureCitation>
).
</paragraph>
<paragraph id="951559422C5ECE38FF42C7879C38F7F4" blockId="18.[136,775,1929,2050]" pageId="18" pageNumber="353">
Thoracic sternum surface smooth; sternites 1 and 2 completely fused, lateral margins sinuous; sternites 3 and 4 fused with only lateral notches demarcating sternites; suture between sternites 2 and 3 almost straight (
<figureCitation id="0D9145C72C5ECE38FE3FC7E79C05F7F4" box="[501,602,2025,2050]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="951559422C5ECE38FC89C4679B4FF9D4" blockId="18.[835,1475,1129,1570]" pageId="18" pageNumber="353">
Adult chelipeds symmetrical or almost so, not enlarged (
<figureCitation id="0D9145C72C5ECE38FC86C4879DEBFB54" box="[844,948,1161,1186]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10A</figureCitation>
). Basis-ischium fused but suture visible, flexor margin with short sharp granules (
<figureCitation id="0D9145C72C5ECE38FB74C4A79B7CFB34" box="[1214,1315,1193,1218]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">Fig. 11A</figureCitation>
). Merus short, surface rugose; flexor and extensor margins with low sharp tubercles and granules; inner distal angle sharp but not elongate, with subdistal tooth (
<figureCitation id="0D9145C72C5ECE38FB6DC5079B44FAD4" box="[1191,1307,1289,1314]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Figs. 10A</figureCitation>
,
<figureCitation id="0D9145C72C5ECE38FAE0C5049B02FAD4" box="[1322,1373,1290,1314]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">11A</figureCitation>
). Carpus rugose; inner distal angle with strong sharp tooth (
<figureCitation id="0D9145C72C5ECE38FA9EC5279BE8FAB4" box="[1364,1463,1321,1346]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10A</figureCitation>
). Chela with dorsal surface rugose and covered with granules, median part smoother; ventral surface near pollex covered with small granules; inner surface smooth; margins of chela with scattered long and short setae, not dense or brush-like; fingers relatively slender, shorter than palm, pigmented brown along distal ⅔; occlusal margins with low teeth, forming small gape when closed (
<figureCitation id="0D9145C72C5ECE38FB92C6079A99F9D4" box="[1112,1222,1545,1570]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Figs. 10A</figureCitation>
,
<figureCitation id="0D9145C72C5ECE38FB1EC6049B5CF9D4" box="[1236,1283,1546,1570]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">11C</figureCitation>
).
</paragraph>
<paragraph id="951559422C5ECE38FC89C6479A0AF834" blockId="18.[835,1475,1609,1986]" pageId="18" pageNumber="353">
Ambulatory legs relatively short, P4 longest, about 1.3× carapace width; outer surface gently rugose; coxa with distal angle slightly raised, no visible serrated flange; P24 with outer surface slightly raised medio-longitudinally, indistinct on P5; merus relatively slender, extensor margin not cristate, with evenly, widely spaced row of blunt, prominent nonsetose nodules; P4 merus length about 4× height, length half carapace width; P24 carpus with outer submarginal ridge, absent on P5; propodus laterally flattened, elongate in P24, short in P5; P24 dactylus falciform, unarmed, P5 dactylus shorter, gently upturned; dactylo-propodal lock visible (
<figureCitation id="0D9145C72C5ECE38FC51C7A79A56F834" box="[923,1033,1961,1986]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Figs. 10A</figureCitation>
,
<figureCitation id="0D9145C72C5ECE38FBDDC7A49A17F834" box="[1047,1096,1962,1986]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">11D</figureCitation>
).
</paragraph>
<paragraph id="951559422C5ECE39FC89C7E79C1FFEC3" blockId="18.[835,1474,2025,2113]" lastBlockId="19.[113,752,189,309]" lastPageId="19" lastPageNumber="354" pageId="18" pageNumber="353">
Female pleon longitudinally ovate; with 6 somites and telson, all free; not completely covering thoracic sternum; somite 1 broadly rectangular, just reaches base of P5 coxae; somite 2 transversely narrower than somite 1, trapezoidal; somites 36 trapezoidal, gradually decreasing in width, somite 6 longest; telson semicircular with convex lateral margins, distinctly longer than somite 6 (
<figureCitation id="0D9145C72C5FCE39FE05C1139C6BFEC3" box="[463,564,285,309]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="19" pageNumber="354">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="951559422C5FCE39FFBBC1539FEAFE43" blockId="19.[113,753,349,437]" pageId="19" pageNumber="354">
Sternopleonal cavity shallow (
<figureCitation id="0D9145C72C5FCE39FE17C1539C16FE83" box="[477,585,349,373]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="19" pageNumber="354">Fig. 11B</figureCitation>
); vulva large, obliquely ovate, covering most of space on sternite 6, without vulvar projections (
<figureCitation id="0D9145C72C5FCE39FE8FC1939FF7FE43" box="[325,424,413,437]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="19" pageNumber="354">Fig. 11B</figureCitation>
).
</paragraph>
<paragraph id="951559422C5FCE39FFBBC1D39F64FE03" blockId="19.[113,315,477,501]" box="[113,315,477,501]" pageId="19" pageNumber="354">
<emphasis id="A7DE85502C5FCE39FFBBC1D39EEDFE03" bold="true" box="[113,178,477,501]" pageId="19" pageNumber="354">Male.</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5FCE39FFBBC2139E8AFD03" pageId="19" pageNumber="354" type="discussion">
<paragraph id="951559422C5FCE39FFBBC2139E8AFD03" blockId="19.[113,752,541,757]" pageId="19" pageNumber="354">
<emphasis id="A7DE85502C5FCE39FFBBC2139EBEFDC3" bold="true" box="[113,225,541,565]" pageId="19" pageNumber="354">Remarks.</emphasis>
<taxonomicName id="52AA22C12C5FCE39FF21C2139FE1FDC3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[235,446,541,565]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="tubur" status="new genus and species">
<emphasis id="A7DE85502C5FCE39FF21C2139FE1FDC3" box="[235,446,541,565]" italics="true" pageId="19" pageNumber="354">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5FCE39FE05C2109C8FFDC3" box="[463,720,541,566]" pageId="19" pageNumber="354" rank="species">new genus and species</taxonomicNameLabel>
, is currently known only from the female
<typeStatus id="4A11E7E02C5FCE39FDDEC2339C2AFDA3" box="[532,629,573,597]" pageId="19" pageNumber="354" type="holotype">holotype</typeStatus>
. Nothing is known about the ecology other than it was collected from soft, muddy substrates. The carapace and pereopods were partially encrusted with what appear to be light-brown, possibly ferric, precipitates, suggestive of reducing or anoxic environs.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="B800EC483DF96920B5C361CF91AE3D05" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C5CCE3AFEF1C1B39A31FB83" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Striacoeloma Ng, Ahyong &amp; T., 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="351" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="351" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B5F7177EDB16DC369E95FD191FD9BDDA">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
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<mods:namePart id="1C87A87F297C7971D0727F2F466C907E">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="1B2E86E8A5F71E94729AA924DA343FF9">Ahyong,</mods:namePart>
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<mods:namePart id="683F7A3B568B470A9F39A0EB3D5B9A2F">Shane T.</mods:namePart>
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<mods:title id="D7FCDB1648EF89994BC2B8EC14B0C6DF">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="630D1DB735CCBE8A78EFDA876D3FA589">2024</mods:date>
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<paragraph id="951559422C5CCE3AFEF1C1B39C0CFE23" blockId="16.[315,595,445,469]" box="[315,595,445,469]" pageId="16" pageNumber="351">
<heading id="CE5DEE2E2C5CCE3AFEF1C1B39C0CFE23" bold="true" box="[315,595,445,469]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="351" reason="2">
<emphasis id="A7DE85502C5CCE3AFEF1C1B39C0CFE23" bold="true" box="[315,595,445,469]" pageId="16" pageNumber="351">
<taxonomicName id="52AA22C12C5CCE3AFEF1C1B39F93FE23" ID-CoL="CL2RM" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[315,460,445,469]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C5CCE3AFEF1C1B39F93FE23" bold="true" box="[315,460,445,469]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
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,
<taxonomicNameLabel id="BCED382B2C5CCE3AFE16C1B39C0CFE23" box="[476,595,445,469]" pageId="16" pageNumber="351" rank="genus">new genus</taxonomicNameLabel>
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<subSubSection id="DDB00AC92C5CCE3AFF42C1F39CA4FD83" pageId="16" pageNumber="351" type="type_taxon">
<paragraph id="951559422C5CCE3AFF42C1F39C46FDC3" blockId="16.[136,775,509,566]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFF42C1F39F7FFDE3" bold="true" box="[136,288,509,533]" pageId="16" pageNumber="351">
<typeStatus id="4A11E7E02C5CCE3AFF42C1F39E9DFDE3" box="[136,194,509,533]" pageId="16" pageNumber="351">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="52AA22C12C5CCE3AFEEDC1F39FA7FDE3" ID-CoL="CL3GY" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[295,504,509,533]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="tubur" status="sp. nov.">
<emphasis id="A7DE85502C5CCE3AFEEDC1F39FA7FDE3" box="[295,504,509,533]" italics="true" pageId="16" pageNumber="351">Striacoeloma tubur</emphasis>
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,
<taxonomicNameLabel id="BCED382B2C5CCE3AFDCCC1F09CD7FDE3" box="[518,648,509,533]" pageId="16" pageNumber="351" rank="species">new species</taxonomicNameLabel>
, by present designation. Gender of genus neuter.
</paragraph>
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<emphasis id="A7DE85502C5CCE3AFF42C2539F14FD83" bold="true" box="[136,331,605,629]" pageId="16" pageNumber="351">Included species.</emphasis>
Only
<taxonomicName id="52AA22C12C5CCE3AFE5BC2539C3DFD83" ID-CoL="CL3GY" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[401,610,605,629]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="tubur" status="sp. nov.">
<emphasis id="A7DE85502C5CCE3AFE5BC2539C3DFD83" box="[401,610,605,629]" italics="true" pageId="16" pageNumber="351">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5CCE3AFDBBC2509CA9FD83" box="[625,758,605,629]" pageId="16" pageNumber="351" rank="species">new species</taxonomicNameLabel>
.
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<subSubSection id="DDB00AC92C5CCE3AFF42C2939D57F983" pageId="16" pageNumber="351" type="diagnosis">
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<emphasis id="A7DE85502C5CCE3AFF42C2939EA2FD43" bold="true" box="[136,253,669,693]" pageId="16" pageNumber="351">Diagnosis.</emphasis>
Carapace subhexagonal; frontal and anterolateral margins with scattered tufts of short and long setae, not brush-like; surface appearing irregularly transversely corrugated, partly eroded; gastric and cardiac regions prominently raised, forming transverse swellings extending to branchial region, almost to lateral margin; surface adjacent to posterior carapace margin raised, with prominent wide rounded transverse ridge adjacent to the posterior carapace margin, forming channel extending to sub-branchial region; orbit submarginal, supraorbital margin almost confluent with general carapace outline, eyes only partially visible in dorsal view; eyes scarcely movable, ocular peduncle short, filling orbit, scarcely movable; lateral lobe of frontal margin small but distinct; anterolateral margin with 4 distinct teeth (including external orbital tooth), first 2 broad, lobiform, with 2 additional tubercles on posterolateral margin; posterolateral margins distinctly converging towards posterior carapace margin. Epistome with shallow biconcave posterior margin, median lobe subtruncate. Endostome short, recessed posteriorly into buccal cavern oblique to plane of adjacent pterygostomial surface, endostomial ridges low, short. Third maxilliped ischium short, quadrate, length 1.2× width; merus with anterolateral angle distinctly auriculiform. Adult female chelipeds symmetrical or almost so, stout, outer surface of chela with margins rugose and covered with granules, median part smoother; margins not lined with dense brush-like setae. Ambulatory legs short, P4 longest, about 1.3× carapace width; merus slender, length about 4× height, extensor margin not cristate, with evenly, widely spaced row of blunt, prominent nodules, outer surface slightly raised medio-longitudinally; P4 merus length about 4× height, length half carapace width.
</paragraph>
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<subSubSection id="DDB00AC92C5CCE3AFF42C6939C48F8C3" pageId="16" pageNumber="351" type="etymology">
<paragraph id="951559422C5CCE3AFF42C6939C48F8C3" blockId="16.[136,775,1693,1846]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFF42C6939F54F943" bold="true" box="[136,267,1693,1717]" pageId="16" pageNumber="351">Etymology.</emphasis>
The name is an arbitrary combination of the Latin word “stria”, for ridge, in arbitrary combination with part of the genus name,
<taxonomicName id="52AA22C12C5CCE3AFE53C6D39C62F903" authorityName="Miers" authorityYear="1884" box="[409,573,1757,1781]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFE53C6D39C62F903" box="[409,573,1757,1781]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
</taxonomicName>
, in recognition of the ridged carapace surface and close relationship to the pilumnid genus. Gender is feminine.
</paragraph>
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<subSubSection id="DDB00AC92C5CCE3AFF42C7539A31FB83" pageId="16" pageNumber="351" type="discussion">
<paragraph id="951559422C5CCE3AFF42C7539A31FB83" blockId="16.[135,775,1885,2101]" lastBlockId="16.[835,1475,189,1142]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFF42C7539EA9F883" bold="true" box="[136,246,1885,1909]" pageId="16" pageNumber="351">Remarks.</emphasis>
Whereas
<taxonomicName id="52AA22C12C5CCE3AFE96C7539FB7F883" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[348,488,1885,1909]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C5CCE3AFE96C7539FB7F883" box="[348,488,1885,1909]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5CCE3AFE3EC7509C3AF880" box="[500,613,1886,1910]" pageId="16" pageNumber="351" rank="genus">new genus</taxonomicNameLabel>
, is superficially similar to various planopilumnids (now in the Pseudozioidea
<bibRefCitation id="F13B24B32C5CCE3AFF42C7939F45F843" author="Alcock A" box="[136,282,1949,1973]" pageId="16" pageNumber="351" pagination="67 - 233" refId="ref13260" refString="Alcock A (1898) Materials for a carcinological fauna of India. No. 3. The Brachyura Cyclometopa. Part I. The family Xanthidae. Journal of the Asiatic Society of Bengal, LVII (II): 67 - 233." type="book chapter" year="1898">Alcock, 1898</bibRefCitation>
) (cf.
<bibRefCitation id="F13B24B32C5CCE3AFE9FC7909FE3F843" author="Ng PKL" box="[341,444,1949,1974]" pageId="16" pageNumber="351" pagination="33 - 61" refId="ref13842" refString="Ng PKL (2010) On the Planopilumnidae Serene, 1984 (Crustacea: Brachyura: Pseudozioidea), with diagnoses of two new pilumnoid genera for species previously assigned to Planopilumnus Balss, 1933. Zootaxa, 2392: 33 - 61." type="journal article" year="2010">Ng, 2010</bibRefCitation>
), it is clearly a pilumnid based the form of its carapace and chelipeds. The new genus is morphologically more similar to
<taxonomicName id="52AA22C12C5CCE3AFE2AC7D39D58F803" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" box="[480,775,2013,2037]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFE2AC7D39CDCF803" box="[480,643,2013,2037]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
<bibRefCitation id="F13B24B32C5CCE3AFD42C7D39D58F803" author="Miers EJ" box="[648,775,2013,2037]" pageId="16" pageNumber="351" pagination="18 - 32" refId="ref13583" refString="Miers EJ (1884) Crustacea. In: Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 1882. Part I. The collections from Melanesia. London, British Museum (Natural History) pp. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
</taxonomicName>
than
<taxonomicName id="52AA22C12C5CCE3AFF0AC7F39F6DF7E3" authorityName="Tesch" authorityYear="1918" box="[192,306,2045,2069]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFF0AC7F39F6DF7E3" box="[192,306,2045,2069]" italics="true" pageId="16" pageNumber="351">Lophoplax</emphasis>
</taxonomicName>
,
<taxonomicName id="52AA22C12C5CCE3AFE88C7F39F8CF7E3" authorityName="Deb" authorityYear="1989" box="[322,467,2045,2069]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFE88C7F39F8CF7E3" box="[322,467,2045,2069]" italics="true" pageId="16" pageNumber="351">Myopilumnus</emphasis>
</taxonomicName>
, or
<taxonomicName id="52AA22C12C5CCE3AFDCEC7F39C04F7E3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[516,603,2045,2069]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C5CCE3AFDCEC7F39C04F7E3" box="[516,603,2045,2069]" italics="true" pageId="16" pageNumber="351">Hosekia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5CCE3AFDA6C7F09CBBF7E0" box="[620,740,2046,2070]" pageId="16" pageNumber="351" rank="genus">new genus</taxonomicNameLabel>
. In
<taxonomicName id="52AA22C12C5CCE3AFF42C8139F4CF7C3" box="[136,275,2077,2101]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFF42C8139F4CF7C3" box="[136,275,2077,2101]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
, like
<taxonomicName id="52AA22C12C5CCE3AFE80C8139FB4F7C3" authorityName="Miers" authorityYear="1884" box="[330,491,2077,2101]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFE80C8139FB4F7C3" box="[330,491,2077,2101]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
</taxonomicName>
, the orbits are submarginal, so the supraorbital margin does not form a strong concavity that noticeably breaks the general carapace outline. The eyes too, are relatively smaller in
<taxonomicName id="52AA22C12C5CCE3AFB63C0F39B68FEE3" box="[1193,1335,253,277]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFB63C0F39B68FEE3" box="[1193,1335,253,277]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
and scarcely movable, compared with the proportionally larger, freely movable eyes of
<taxonomicName id="52AA22C12C5CCE3AFBCDC1339A24FEA3" authorityName="Tesch" authorityYear="1918" box="[1031,1147,317,341]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFBCDC1339A24FEA3" box="[1031,1147,317,341]" italics="true" pageId="16" pageNumber="351">Lophoplax</emphasis>
</taxonomicName>
,
<taxonomicName id="52AA22C12C5CCE3AFB46C1339B7FFEA3" authorityName="Deb" authorityYear="1989" box="[1164,1312,317,341]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFB46C1339B7FFEA3" box="[1164,1312,317,341]" italics="true" pageId="16" pageNumber="351">Myopilumnus</emphasis>
</taxonomicName>
, and
<taxonomicName id="52AA22C12C5CCE3AFAA8C1339BE4FEA3" box="[1378,1467,317,341]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFAA8C1339BE4FEA3" box="[1378,1467,317,341]" italics="true" pageId="16" pageNumber="351">Hosekia</emphasis>
</taxonomicName>
. The chelae of
<taxonomicName id="52AA22C12C5CCE3AFC23C1539A24FE83" box="[1001,1147,349,373]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFC23C1539A24FE83" box="[1001,1147,349,373]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
and
<taxonomicName id="52AA22C12C5CCE3AFB7DC1539B00FE83" authorityName="Miers" authorityYear="1884" box="[1207,1375,349,373]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFB7DC1539B00FE83" box="[1207,1375,349,373]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
</taxonomicName>
are also stouter than in
<taxonomicName id="52AA22C12C5CCE3AFC17C1739A10FE63" authorityName="Tesch" authorityYear="1918" box="[989,1103,381,405]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFC17C1739A10FE63" box="[989,1103,381,405]" italics="true" pageId="16" pageNumber="351">Lophoplax</emphasis>
</taxonomicName>
and allies, and both have a shallow biconcave posterior margin of the epistome with the median lobe broader and subtruncate (
<figureCitation id="0D9145C72C5CCE3AFB4BC1B39ABFFE23" box="[1153,1248,445,469]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10D</figureCitation>
) (versus median lobe broadly triangular; Ng et al., 2022: fig. 3F).
<taxonomicName id="52AA22C12C5CCE3AFAE7C1D39BE4FE03" box="[1325,1467,477,501]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFAE7C1D39BE4FE03" box="[1325,1467,477,501]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
, however, differs from
<taxonomicName id="52AA22C12C5CCE3AFBF0C1F39A81FDE3" authorityName="Miers" authorityYear="1884" box="[1082,1246,509,533]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFBF0C1F39A81FDE3" box="[1082,1246,509,533]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
</taxonomicName>
in having the dorsal surface of carapace ornamented with transverse, corrugated swellings (
<figureCitation id="0D9145C72C5CCE3AFC7FC2339A6AFDA3" box="[949,1077,573,597]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10AC</figureCitation>
) (versus surface granulate and rugose but without swellings and regions well demarcated; Ng et al., 2022: fig. 2); the frontal and anterolateral margins have tufts of short and long setae but not dense or brush-like (
<figureCitation id="0D9145C72C5CCE3AFA5DC2939DCDFD23" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10A, B</figureCitation>
) (versus margins evenly lined with dense short and long setae, appearing brush-like; Ng et al., 2022: figs. 1A, B, 2A, B, 5A, B, 7A, B); the outer surface of the chelae are covered with scattered long and short setae, not dense or brush-like (
<figureCitation id="0D9145C72C5CCE3AFC2CC3339A13FCA3" box="[998,1100,829,853]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10A</figureCitation>
) (versus lined with dense brush-like setae; Ng et al., 2022: figs. 1A, B, 2A, B, 5A, B, 7A, B); the ischium of the third maxilliped is subquadrate and shorter (
<figureCitation id="0D9145C72C5CCE3AFC56C3939DA5FC43" box="[924,1018,925,949]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10E</figureCitation>
) (length 1.2× width versus 1.3×; Ng et al., 2022: fig. 4A); and the ambulatory merus is dentate along the extensor margin (
<figureCitation id="0D9145C72C5CCE3AFBE5C3D39ACDFC03" box="[1071,1170,989,1013]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="16" pageNumber="351">Fig. 11D</figureCitation>
) (versus smooth; Ng et al., 2022: figs. 2, 4B); and the vulva is proportionately larger, occupying most of the space of sternite 5 (
<figureCitation id="0D9145C72C5CCE3AFACDC4139B39FBC3" box="[1287,1382,1053,1077]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="16" pageNumber="351">Fig. 11B</figureCitation>
) (versus vulva smaller, occupying two-thirds the space of sternite 5; Ng et al., 2022: fig. 7E, F).
</paragraph>
</subSubSection>
</treatment>
</document>