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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.797.29510" ID-GBIF-Dataset="17a988bc-106e-4cd5-af88-8a606353b34f" ID-PMC="PMC6255853" ID-Pensoft-Pub="1313-2970-797-19" ID-PubMed="30505161" ID-ZBK="A5403588F43D40FF81EE2009E49823B4" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-797-19" ModsDocOrigin="ZooKeys 797" ModsDocTitle="A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1542677994046" checkinUser="pensoft" docAuthor="Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai &amp; Panha, Somsak" docDate="2018" docId="F9DF34622ABDAE14E4695A8643751306" docLanguage="en" docName="ZooKeys 797: 19-69" docOrigin="ZooKeys 797" docSource="http://dx.doi.org/10.3897/zookeys.797.29510" docTitle="Spinaxytes sutchariti Srisonchai, Enghoff &amp; Panha, sp. n." docType="treatment" docUuid="4E54ED20-66AB-4BF7-B40D-D6F6699CF282" docUuidSource="ZooBank" docVersion="5" lastPageNumber="19" masterDocId="7C43FFACB225AA2193482143FFF2FF98" masterDocTitle="A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="69" masterPageNumber="19" originalUpdateDomain="Boston" originalUpdateTime="1542677994046" originalUpdateUser="pensoft" pageNumber="19" updateTime="1668166441925" updateUser="ExternalLinkService">
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<mods:title>A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Srisonchai, Ruttapon</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:date>2018</mods:date>
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<mods:number>797</mods:number>
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<taxonomicName LSID="http://zoobank.org/4E54ED20-66AB-4BF7-B40D-D6F6699CF282" authority="Srisonchai, Enghoff &amp; Panha" class="Diplopoda" family="Paradoxosomatidae" genus="Spinaxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinaxytes sutchariti" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="sutchariti">Spinaxytes sutchariti Srisonchai, Enghoff &amp; Panha</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="19">sp. n.</taxonomicNameLabel>
Figs 4P, Q; 5G; 24; 25
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
<paragraph pageId="0" pageNumber="19">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="19">
Holotype. ♂, THAILAND, Krabi Province, Muang Krabi District, Tham Na Mee (Na Mee Cave),
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.136666">8°08'12&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="98.80639">98°48'23&quot;E</geoCoordinate>
, ca. 70 m a.s.l., 31 Aug. 2015, C. Sutcharit leg. (CUMZ-pxDGT00217). Paratypes. 7 ♂♂, 4 ♀♀, same data as for holotype (CUMZ-pxDGT00218).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="etymology">
<paragraph pageId="0" pageNumber="19">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="19">The name honours associate professor Dr. Chirasak Sutcharit, malacologist of ASRU (CUMZ), collector of this new species and numerous other dragon millipedes.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="19">
Male femora without modification, sternal lobe between male coxae 4 incompletely bilobed. Similar in these respects to
<taxonomicName lsidName="S. palmata" pageId="0" pageNumber="19" rank="species" species="palmata">S. palmata</taxonomicName>
sp. n., but differs by having: a large and round lamina lateralis; tip of lamina medialis terminating in two spines; distal part of solenomere circular in tranverse section.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="description">
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
<paragraph pageId="0" pageNumber="19">SIZE. Length 20-25 mm (male), 23-27 mm (female); width of midbody metazona 1.5-1.8 mm (male), 1.9-2.2 mm (female). Width of collum = 2 = 3 = 4 &lt;head = 5-16, thereafter body gradually tapering towards telson.</paragraph>
<paragraph pageId="0" pageNumber="19">
Colour (Figure 24
<normalizedToken originalValue="AC">A-C</normalizedToken>
). Specimens in life with body black; paraterga brownish white; head, antennae (except whitish distal part of antennomeres 7 and 8), collum, prozona, metaterga and epiproct black; surface below paraterga black/brownish black; sterna brown; legs brown/blackish brown; a few basal podomeres whitish brown.
</paragraph>
<paragraph pageId="0" pageNumber="19">Antennae. Reaching to body ring 8 or 9 (male) and 6 (female) when stretched dorsally.</paragraph>
<paragraph pageId="0" pageNumber="19">Collum. With three transverse rows of setiferous tubercles, 4+4 tubercles in anterior row, 1+1 tubercles in intermediate row and 2+2 tubercles in posterior row; with one inconspicuous setiferous notch at lateral margin; paraterga spiniform, quite short, tip sharp, elevated at ca. 15°-20° in both male and female, directed caudolaterad.</paragraph>
<paragraph pageId="0" pageNumber="19">Tegument. Quite dull; collum and metaterga (posterior part) coarsely microgranulate; metaterga (anterior part) and surface below paraterga smooth.</paragraph>
<paragraph pageId="0" pageNumber="19">Metaterga. With two transverse rows of setiferous tubercles and setiferous spines; metaterga 2-19 with 2+2 tubercles in anterior row and 2+2 spines in posterior row; lateral spines of posterior row bigger and longer than mesal ones, subequal in size and length on all body rings.</paragraph>
<paragraph pageId="0" pageNumber="19">Paraterga. Long; directed dorsolaterad on body rings 2-16, elevated at ca. 45°-50° (male) 40°-50° (female), directed dorsocaudad on ring 17, directed increasingly caudad on body rings 18 and 19. Ozopore visible in lateral view.</paragraph>
<paragraph pageId="0" pageNumber="19">Telson. Epiproct quite long; tip subtruncate; lateral setiferous tubercles conspicuous; apical tubercles inconspicuous. Hypoproct subtrapeziform (in some specimens subtriangular); caudal margin round (in some specimens angular), with inconspicuous setiferous tubercles.</paragraph>
<paragraph pageId="0" pageNumber="19">Sterna (Figure 4P). Sternal lobe between male coxae 4 incompletely bilobed; tips sharp, in situ directed laterad; posterior surface bearing 2 pores.</paragraph>
<paragraph pageId="0" pageNumber="19">Legs (Figure 4Q). Male femora without modification.</paragraph>
<paragraph pageId="0" pageNumber="19">Gonopods (Figs 5G, 25). Coxa subequal in length to femur. Prefemoral part ca. 2/3 as long as femur. Femur not enlarged distally. Postfemoral part narrow. Mesal sulcus and lateral sulcus wide. Solenophore bigger and longer than postfemoral part: lamina lateralis oval, large, long, tip round: lamina medialis long and slender; base enlarged, slightly attenuated near the tip; tip curving down, with two sharp spines (one smaller, one bigger). Solenomere curving up, circular in transverse section, tip directed anteriad.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="distribution">
<paragraph pageId="0" pageNumber="19">Distribution and habitat</paragraph>
<paragraph pageId="0" pageNumber="19">
(Figure 24D).
<taxonomicName lsidName="S. sutchariti" pageId="0" pageNumber="19" rank="species" species="sutchariti">S. sutchariti</taxonomicName>
sp. n. is known only from the type locality, and we regard it as endemic to Thailand. The new species can be found in the same area as
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Gigaxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gigaxytes gigas" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="gigas">Gigaxytes gigas</taxonomicName>
, but we assume that they live in different microhabitats:
<taxonomicName lsidName="G. gigas" pageId="0" pageNumber="19" rank="species" species="gigas">G. gigas</taxonomicName>
was collected from the ground in leaf litter, whereas the new species was found on humid rock walls.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="remarks">
<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="19">
We found variation in the hypoproct: in some specimens subtrapeziform, in others subtriangular; caudal margin in some individuals round, in others angular. Parasitic mite larvae, probably of the genus
<taxonomicName family="Erythraeidae" genus="Leptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptus" order="Trombidiformes" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">Leptus</taxonomicName>
Latreille, 1796, were found attached to the anterior body part of some female specimens. Larvae of?
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have previously been found on species of
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(
<taxonomicName lsidName="D. cervina" pageId="0" pageNumber="19" rank="species" species="cervina">D. cervina</taxonomicName>
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagaxytes" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">Nagaxytes</taxonomicName>
(
<taxonomicName lsidName="N. acantherpestes" pageId="0" pageNumber="19" rank="species" species="acantherpestes">N. acantherpestes</taxonomicName>
(Golovatch &amp; Enghoff, 1994)) (
<bibRefCitation author="Srisonchai, R" journalOrPublisher="ZooKeys" pageId="0" pageNumber="19" pagination="1 - 177" title="A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae)." url="https://doi.org/10.3897/zookeys.761.24214" volume="761" year="2018 a">Srisonchai et al. 2018a</bibRefCitation>
,
<bibRefCitation author="Srisonchai, R" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="19" pagination="1 - 44" title="A revision of dragon millipedes II: the new genus Nagaxytes gen. nov., with the description of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)." url="https://doi.org/10.5852/ejt.2018.462" volume="462" year="2018 b">2018b</bibRefCitation>
, see also
<bibRefCitation author="Southcott, RV" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="19" pagination="1 - 153" title="Revision of the larvae of Leptus Latreille (Acarina: Erythraeidae) of Europe and North America, with descriptions of post-larval instars." url="https://doi.org/10.1111/j.1096-3642.1992.tb01228.x" volume="105" year="1992">Southcott 1992</bibRefCitation>
).
</paragraph>
<caption pageId="0" pageNumber="19">
<paragraph pageId="0" pageNumber="19">
Figure 15. Photographs of live
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Spinaxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinaxytes krabiensis" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="krabiensis">Spinaxytes krabiensis</taxonomicName>
sp. n. and habitat A ♂ paratype, CUMZ-pxDGT00212 B ♂, specimen from Tham Sa Yuan Thong (Sa Yuan Thong Cave) C ♀ paratype D mating couple E habitat.
</paragraph>
</caption>
<caption pageId="0" pageNumber="19">
<paragraph pageId="0" pageNumber="19">
Figure 16.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Spinaxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinaxytes krabiensis" order="Polydesmida" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="krabiensis">Spinaxytes krabiensis</taxonomicName>
sp. n., ♂ paratype, CUMZ-pxDGT00212 A, B anterior body part C, D body rings 9-11 E, H, I posteriormost body rings and telson F, G sternal lobe between coxae 4 J legs 4-8 K legs 1-3 L gonopods M right antenna N sculpture of body ring 10.
</paragraph>
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<emphasis italics="true">Antennae</emphasis>
subfiliformes, extrosum crassiores.
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<emphasis italics="true">Elytra</emphasis>
marginata.
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<emphasis italics="true">Caput</emphasis>
sub thoracis clypeo plano reconditum.
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.
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Buthus bonito
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&amp; Geniez, 2005
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</paragraph>
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<paragraph id="DC7E31DEEBA23C964F915A3A2499F166" pageId="11" pageNumber="26">
=
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<bibRefCitation id="2E94037F4B78F0B7ED578B692585563D" pageId="11" pageNumber="26">Touloun 2012</bibRefCitation>
: 46, 48-58, fig.10, 13, 14 (Syn. n.). 1 F holotype (MNHN), Khnifiss lagoon, Tan-Tan Province, Morocco.
</paragraph>
<paragraph id="1593A2C3D8465107404C4108BFDD46DF" pageId="11" pageNumber="26">
<taxonomicName id="ED9B0CAD0B8BDFB9C70FECA212E369A5" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus bonito" order="Scorpiones" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="bonito">Buthus bonito</taxonomicName>
:
<bibRefCitation id="7742169324AA5AD4FB5CFAAA4982C52B" author="Lourenco, WR" journalOrPublisher="Euscorpius" pageId="31" pageNumber="46" pagination="1 - 6" title="A new scorpion species of the genus Buthus Leach, 1815 (Scorpiones, Buthidae) from Morocco." url="http://www.science.marshall.edu/fet/euscorpius/p2005_19.pdf" volume="19" year="2005">
<normalizedToken id="640283696D83117A4CB421237AB7C0FD" originalValue="Lourenço">Lourenco</normalizedToken>
and Geniez 2005
</bibRefCitation>
: 1-5, fig. 1-8, 10;
<bibRefCitation id="54944E3C0CAAF0BDD5EB0AED969FE70F" author="Touloun, O" journalOrPublisher="Poiretia" pageId="35" pageNumber="50" url="http://s1.e-monsite.com/2009/01/24/458084arachnides-54-pdf" year="2008">Touloun et al. 2008</bibRefCitation>
: 3-4, fig.1;
<bibRefCitation id="A8C404A432DD5D6A272699E600A542EE" author="Stockmann, R" journalOrPublisher="P Editions, Rodez" pageId="35" pageNumber="50" title="Scorpions of the World. N. A." url="http://www.napeditions.com/en/scorpions/20-scorpions-du-monde.htmlidTab2" year="2010">Stockmann and Ythier 2010</bibRefCitation>
: 362-363;
<bibRefCitation id="B518ED6C07903B31E411B39906FCC933" author="Pedroso, D" journalOrPublisher="African Invertebrates" pageId="33" pageNumber="48" pagination="298 - 308" title="Phylogeography of Buthus Leach, 1815 (Scorpiones: Buthidae): a multigene molecular approach reveals a further complex evolutionary history in the Maghreb." url="https://doi.org/10.3377/004.048.0216" volume="48" year="2013">Pedroso et al. 2013</bibRefCitation>
: 300;
<bibRefCitation id="658582D77577D6C613FEC1F7CBCDF4A6" author="Aboumaad, B" journalOrPublisher="Bulletin de la Societe de pathologie exotique" pageId="22" pageNumber="37" pagination="1 - 9" title="L'envenimation scorpionique au Maroc: scorpions du genre Androctonus, Buthus et Hottentota." url="https://doi.org/10.1007/s13149-014-0332-7" volume="107" year="2014">
<normalizedToken id="F778B0C0CE7879F6154B9FF79ECD9D9E" originalValue="Aboumaâd">Aboumaad</normalizedToken>
et al. 2014
</bibRefCitation>
: 6;
<bibRefCitation id="2464C40E656227EC634E5D8AF5E45998" author="Touloun, O" journalOrPublisher="Asian Pacific Journal of Tropical Disease" pageId="35" pageNumber="50" pagination="878 - 881" title="Species composition and geographical distribution of Saharan scorpion fauna, Morocco." url="https://doi.org/10.1016/S2222-1808(16)61150-7" volume="6" year="2016">Touloun et al. 2016</bibRefCitation>
: 880, fig. 2D.
</paragraph>
</subSubSection>
<subSubSection id="3C854260D29585592D19C99513B494CE" pageId="11" pageNumber="26" type="type material">
<paragraph id="2AA8468AE0BE6CEB7CBA04C0108C1339" pageId="11" pageNumber="26">Type material.</paragraph>
<paragraph id="7F5A1705A498F91DC83E97438CDE6B68" pageId="11" pageNumber="26">1 M holotype (MNHN N° RS8669), Khnifiss lagoon (approx. 27.93°, -12.34°), Tarfaya, Morocco. Paratypes: 2 F (MNHN N° RS8670), from the same locality.</paragraph>
</subSubSection>
<subSubSection id="87904052B3A7FDEFF56E1EBB1F5DF08F" pageId="11" pageNumber="26" type="distribution">
<paragraph id="E971D59944090960570964CC9AFB6CA8" pageId="11" pageNumber="26">Distribution.</paragraph>
<paragraph id="550F33CD182F96D9C7827F65D793200E" pageId="11" pageNumber="26">
known from the Atlantic coast of Morocco south of Tan-Tan extending almost to Dakhla in the Western Sahara (
<bibRefCitation id="3D9B2E020DB5B8004C085A4FDFF6B32E" author="Touloun, O" journalOrPublisher="Asian Pacific Journal of Tropical Disease" pageId="35" pageNumber="50" pagination="878 - 881" title="Species composition and geographical distribution of Saharan scorpion fauna, Morocco." url="https://doi.org/10.1016/S2222-1808(16)61150-7" volume="6" year="2016">Touloun et al. 2016</bibRefCitation>
).
</paragraph>
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<subSubSection id="0576682286690A6695CF43C95FF3FDED" pageId="11" pageNumber="26" type="remarks">
<paragraph id="6ECA80516F67A5DAE6B265540E9F9982" pageId="11" pageNumber="26">Remarks.</paragraph>
<paragraph id="A485A15B82D67FF8B6C941FD83C7CC04" pageId="11" pageNumber="26">
Although the type material of
<taxonomicName id="BCBFC722B54A1E081BD6A3328981C6CB" lsidName="B. sabulicola" pageId="11" pageNumber="26" rank="species" species="sabulicola">B. sabulicola</taxonomicName>
was collected in 2002 by Touloun, Stockmann and Slimani, the species was not formally described until the publication of the PhD thesis of Oulaid Touloun in 2012. The type specimens of
<taxonomicName id="D5720599E3DC19BCF4EF67A9BE948DB8" lsidName="B. bonito" pageId="11" pageNumber="26" rank="species" species="bonito">B. bonito</taxonomicName>
and
<taxonomicName id="B69448B677AF29B3A714A5B12D08800C" lsidName="B. sabulicola" pageId="11" pageNumber="26" rank="species" species="sabulicola">B. sabulicola</taxonomicName>
are from the exact same locality, the Khnifiss lagoon, and both descriptions are almost identical.
<bibRefCitation id="04DE4FE7D277269CE55714C8D4DF5D2B" author="Touloun, O" journalOrPublisher="Asian Pacific Journal of Tropical Disease" pageId="35" pageNumber="50" pagination="878 - 881" title="Species composition and geographical distribution of Saharan scorpion fauna, Morocco." url="https://doi.org/10.1016/S2222-1808(16)61150-7" volume="6" year="2016">Touloun et al. (2016)</bibRefCitation>
, probably by mistake, indicated that the fifth metasomal segment and telson are darkened in
<taxonomicName id="2C7EEDAF82C2D0170E28EAF72B617B16" lsidName="B. bonito" pageId="11" pageNumber="26" rank="species" species="bonito">B. bonito</taxonomicName>
, but the trait does not appear in the accompanying figure.
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<document id="CEF65B9E8E25320E6A731BC0DC4D8D60" ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e7964" ID-PMC="PMC4911545" ID-Pensoft-Pub="1314-2828-4-7964" ID-PubMed="27350764" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e7964" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="Guide to the littoral zone vascular flora of Carolina bay lakes (U.S.A.)" checkinTime="1459887716076" checkinUser="pensoft" docAuthor="Howell, Nathan, Krings, Alexander &amp; Braham, Richard R" docDate="2016" docId="F9DFCA90F9FEBC750CCAF42C120722E2" docLanguage="en" docName="BiodivDatJour 4: e7964" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e7964" docTitle="Ilex coriacea Chapm." docType="treatment" docVersion="5" lastPageNumber="7964" masterDocId="FF542A1F5274FF9AAF1CF606FF9DFF9E" masterDocTitle="Guide to the littoral zone vascular flora of Carolina bay lakes (U. S. A.)" masterLastPageNumber="7964" masterPageNumber="7964" pageNumber="7964" updateTime="1668126861817" updateUser="ExternalLinkService">
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<taxonomicName id="C08CBFC97984CA5AFD5F033524DBFF0D" ID-CoL="6MV99" ID-ENA="185501" authority="(Pursh) Chapm." class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex coriacea" order="Aquifoliales" pageId="0" pageNumber="7964" phylum="Tracheophyta" rank="species" species="coriacea">Ilex coriacea (Pursh) Chapm.</taxonomicName>
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<taxonomicName id="D3C8AED07974E9ED0B2339446AA1759C" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex coriacea" order="Aquifoliales" pageId="0" pageNumber="7964" phylum="Tracheophyta" rank="species" species="coriacea">Ilex coriacea</taxonomicName>
Basionym:
<taxonomicName id="CE9FD7CCE33CE5F0C006E02B627D72F3" class="Magnoliopsida" family="Aquifoliaceae" genus="Prinos" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Prinos coriaceus" order="Aquifoliales" pageId="0" pageNumber="7964" phylum="Tracheophyta" rank="species" species="coriaceus">Prinos coriaceus</taxonomicName>
Pursh
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<paragraph id="ECE8DDC4251B905D0DC6640E9BD5567B" pageId="0" pageNumber="7964">
<taxonomicName id="D269F53F163B8BBA1DF125C39CB1D83C" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex coriacea" order="Aquifoliales" pageId="0" pageNumber="7964" phylum="Tracheophyta" rank="species" species="coriacea">Ilex coriacea</taxonomicName>
Taxon concept: [= RAB, GW, Weakley]
</paragraph>
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<paragraph id="0EE9C2931111F7C1B6558EB29F88E068" pageId="0" pageNumber="7964">Distribution</paragraph>
<paragraph id="53723F807A8D9377B84C7AEA01D12A58" pageId="0" pageNumber="7964">Bay Tree Lake (Rare): Howell BATR16 (NCSC!)</paragraph>
<paragraph id="1C570B5374CAA9593F48F92B8E012C59" pageId="0" pageNumber="7964">Jones Lake (Occasional): Howell JOLA10, 32 (NCSC!)</paragraph>
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<paragraph id="8A859050D6FB412A61764AFF392ED884" pageId="0" pageNumber="7964">Shrubs. Juncture of eulittoral and supralittoral zones (NLSSC). AprMay; SepOct. Fig. 109</paragraph>
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Figs 22, 28 31
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:9, type locality &quot;juxta fluvium Svakop,&quot; holotype in Naturhistoriska Riksmuseet, Stockholm)
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(Boheman)
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:367)
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</subSubSection>
<subSubSection pageId="11" pageNumber="34" type="diagnosis">
<paragraph pageId="11" pageNumber="34">Diagnosis.</paragraph>
<paragraph pageId="11" pageNumber="34">Apparent body length (ABL) 18-24 mm; easily recognized by the golden-yellow pubescence on the head, pronotum, and base of elytra adjacent to the suture (Fig. 22). The elytral surface sculpturing is also diagnostic, with small narrow costae separated by flat intervals with a single row of minute punctures in each interval (Fig. 28).</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="34" type="materials examined">
<paragraph pageId="11" pageNumber="34">Materials examined.</paragraph>
<paragraph pageId="11" pageNumber="34">45 specimens from the following localities: Eastern Cape Province: Nduma, Willowmore. KwaZulu/Natal Province: Maritzburg, Melmoth, Mkuze, St. Lucia, &quot;E. Zululand.&quot; North West Province: Lichtenburg, 30 km W Vryberg.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,263 @@
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<mods:namePart>Kahanpaeae, Jere</mods:namePart>
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<taxonomicName authority="Zetterstedt, 1846" authorityName="Zetterstedt" authorityYear="1846" class="Insecta" family="Ephydridae" genus="Hydrellia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrellia mutata" order="Diptera" pageId="0" pageNumber="4701" phylum="Arthropoda" rank="species" species="mutata">Hydrellia mutata (Zetterstedt, 1846)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4701" type="materials_examined">
<paragraph pageId="0" pageNumber="4701">Materials</paragraph>
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Type status:
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. Occurrence: catalogNumber:
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<materialsCitation collectingDate="1936-7-1" collectionCode="MZH" collectorName="Tiensuu, Lauri" country="Russia" latitude="61.7" location="Russia" longitude="30.7" pageId="0" pageNumber="4701" specimenCode="http: // id. luomus. fi / GV. 16612" specimenCount="1" typeStatus="Other material">
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Hydrelliamutata (Zetterstedt, 1846); order: Diptera; family: Ephydridae; genus: Hydrellia; specificEpithet: mutata; scientificNameAuthorship: (Zetterstedt, 1846); Location: country:
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, Jere
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Coll.
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; basisOfRecord: PreservedSpecimen
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<materialsCitation collectionCode="Coll. Kahanpaeae" collectorName="Kahanpaeae, Jere" country="Finland" latitude="60.308" location="Vaanila" longitude="24.118" pageId="0" pageNumber="4701" specimenCount="1" typeStatus="Other material">
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<typeStatus pageId="0" pageNumber="4701">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="4701">
<normalizedToken originalValue="Kahanpää">Kahanpaeae</normalizedToken>
, Jere
</collectorName>
; individualCount:
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; sex:
<specimenType pageId="0" pageNumber="4701">M</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="4701">adult</specimenType>
; Taxon: scientificName: Hydrelliamutata (Zetterstedt, 1846); order: Diptera; family: Ephydridae; genus: Hydrellia; specificEpithet: mutata; scientificNameAuthorship: (Zetterstedt, 1846); Location: country:
<collectingCountry pageId="0" pageNumber="4701">Finland</collectingCountry>
; stateProvince: V; municipality: Lohja; locality:
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; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 50; georeferencedBy:
<normalizedToken originalValue="Kahanpää">Kahanpaeae</normalizedToken>
, Jere; Identification: identifiedBy:
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<normalizedToken originalValue="Kahanpää">Kahanpaeae</normalizedToken>
, Jere
</determinerName>
; dateIdentified: 2014-02-05; Event: year: 2004; month: 7; day: 16; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="4701">
Coll.
<normalizedToken originalValue="Kahanpää">Kahanpaeae</normalizedToken>
</collectionCode>
; basisOfRecord: PreservedSpecimen
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4701" type="diagnosis">
<paragraph pageId="0" pageNumber="4701">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="4701">
Best identified by structures of the male genitalia as illustrated by
<bibRefCitation author="Collin, J. E." journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Venezia" pageId="0" pageNumber="4701" pagination="7 - 18" title="A contribution towards the knowledge of the male genitalia of species of Hydrellia (Diptera, Ephydridae)" volume="16 [1963]" year="1966">Collin (1966)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4701" type="distribution">
<paragraph pageId="0" pageNumber="4701">Distribution</paragraph>
<paragraph pageId="0" pageNumber="4701">
First recorded from Finland by
<bibRefCitation author="Zatwarnicki, Tadeusz" journalOrPublisher="ZooKeys" pageId="0" pageNumber="4701" pagination="339 - 346" title="Checklist of the family Ephydridae of Finland (Insecta, Diptera)" volume="441" year="2014">
Zatwarnicki and
<normalizedToken originalValue="Kahanpää">Kahanpaeae</normalizedToken>
(2014)
</bibRefCitation>
. Northern, Western and Central Europe (
<bibRefCitation pageId="0" pageNumber="4701">Zatwarnicki 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4701" type="ecology">
<paragraph pageId="0" pageNumber="4701">Ecology</paragraph>
<paragraph pageId="0" pageNumber="4701">
Larvae have recently been found feeding on
<taxonomicName class="Liliopsida" family="Alismataceae" genus="Alisma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alisma" order="Alismatales" pageId="0" pageNumber="4701" phylum="Tracheophyta" rank="genus">Alisma</taxonomicName>
(
<bibRefCitation author="Robbins, R." journalOrPublisher="Warwickshire Museum Service" pageId="0" pageNumber="4701" title="A provisional atlas of the leaf miners of Warwickshire, with notes on others occurring in the Midlands" year="1990">Robbins 1990</bibRefCitation>
) and
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, possibly also
<taxonomicName class="Liliopsida" family="Araceae" genus="Lemna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lemna" order="Alismatales" pageId="0" pageNumber="4701" phylum="Tracheophyta" rank="genus">Lemna</taxonomicName>
and
<taxonomicName class="Liliopsida" family="Alismataceae" genus="Alisma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alisma" order="Alismatales" pageId="0" pageNumber="4701" phylum="Tracheophyta" rank="genus">Alisma</taxonomicName>
(see
<bibRefCitation author="Deonier, D. L." journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="4701" pagination="1 - 147" title="A systematic and ecological study of Nearctic Hydrellia (Diptera: Ephydridae)" volume="68" year="1971">Deonier 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4701" type="notes">
<paragraph pageId="0" pageNumber="4701">Notes</paragraph>
<paragraph pageId="0" pageNumber="4701">Correctly identified by Richard Frey and Lauri Tiensuu but the record was never published. Specimen http://id.luomus.fi/GV.16611 is illustrated in Fig. 1b.</paragraph>
</subSubSection>
</treatment>
</document>