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data/F9/5B/85/F95B85F806D6615429BDA656A302ACEB.xml
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data/F9/5B/85/F95B85F806D6615429BDA656A302ACEB.xml
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<mods:titleInfo>
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<mods:title>Cyanobacteria of Greece: an annotated checklist</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gkelis, Spyros</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ourailidis, Iordanis</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:title>Biodiversity Data Journal</mods:title>
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<mods:part>
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<mods:date>2016</mods:date>
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<taxonomicName authority="(Kuetzing) Naegeli, 1849" authorityName="(Kuetzing) Naegeli" authorityYear="1849" class="Cyanophyceae" family="Chroococcaceae" genus="Gloeothece" higherTaxonomySource="CoL" kingdom="Bacteria" lsidName="Gloeothece palea" order="Chroococcales" pageId="0" pageNumber="10084" phylum="Cyanobacteria" rank="species" species="palea">
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Gloeothece palea (
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)
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, 1849
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<subSubSection pageId="0" pageNumber="10084" type="notes">
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<paragraph pageId="0" pageNumber="10084">Notes</paragraph>
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<paragraph pageId="0" pageNumber="10084">
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<bibRefCitation author="Anagnostidis, K." journalOrPublisher="Nova Hedwigia" pageId="0" pageNumber="10084" pagination="309 - 335" title="U ̈ ber die Okologie einiger Spirulina Arten" volume="11" year="1966">Anagnostidis and Golubic 1966</bibRefCitation>
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316
data/F9/5B/87/F95B87D7FFA1950DFF49FC42D6275CAF.xml
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316
data/F9/5B/87/F95B87D7FFA1950DFF49FC42D6275CAF.xml
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<mods:title id="1DC4064BF6C4ECC3529E01EA9DD5A10A">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<taxonomicName id="B6F24D42FFA1950CFF49FC42D1DB58E2" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,951,969,995]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Vovidesa">
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<emphasis id="4386EAD3FFA1950CFF49FC42D34B58E2" bold="true" box="[151,295,969,995]" italics="true" pageId="20" pageNumber="171">Ceratophila</emphasis>
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(
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<emphasis id="4386EAD3FFA1950CFEE9FC42D3F258E2" bold="true" box="[311,414,969,995]" italics="true" pageId="20" pageNumber="171">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFA1950CFE6CFC42D04358E2" bold="true" box="[434,559,969,995]" italics="true" pageId="20" pageNumber="171">mixeorum</emphasis>
|
||||
Tang, Skelley & Pérez-Farrera
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFA1950CFC1DFC42D63F58E2" box="[963,1107,969,995]" pageId="20" pageNumber="171" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="39E8654AFFA1950CFF49FC67D3355F05" box="[151,345,1004,1029]" pageId="20" pageNumber="171" type="description">
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<paragraph id="714D36C1FFA1950CFF49FC67D3355F05" blockId="20.[151,1107,969,1029]" box="[151,345,1004,1029]" pageId="20" pageNumber="171">
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(
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<figureCitation id="E9C92A44FFA1950CFF41FC67D2935F05" box="[159,255,1004,1029]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="20" pageNumber="171">Figs. 1G</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFA1950CFED5FC67D3745F05" box="[267,280,1004,1028]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="20" pageNumber="171">7</figureCitation>
|
||||
A–H)
|
||||
</paragraph>
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<subSubSection id="39E8654AFFA1950CFF49FBBFD1E75FB6" pageId="20" pageNumber="171" type="diagnosis">
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<paragraph id="714D36C1FFA1950CFF49FBBFD1E75FB6" blockId="20.[151,1437,1076,2037]" pageId="20" pageNumber="171">
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||||
<emphasis id="4386EAD3FFA1950CFF49FBBFD3365F4C" bold="true" box="[151,346,1076,1101]" pageId="20" pageNumber="171">Adult diagnosis.</emphasis>
|
||||
Distinguished from other
|
||||
<taxonomicName id="B6F24D42FFA1950CFD58FBBFD1E15F4D" authority="(Vovidesa )" baseAuthorityName="Vovidesa" box="[646,909,1076,1100]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="155" phylum="Tracheophyta" rank="genus">
|
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<emphasis id="4386EAD3FFA1950CFD58FBBFD1615F4D" box="[646,781,1076,1100]" italics="true" pageId="20" pageNumber="171">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA1950CFCC1FBBFD1EC5F4D" box="[799,896,1076,1100]" italics="true" pageId="20" pageNumber="171">Vovidesa</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
species by male with subapical emarginations on tibiae, submentum of both sexes with sparse punctures and setae, female elytra glossy medially and strongly dulled laterally, male genitalia with median lobe nearly cylindrical not laterally flattened, and with a known distribution in
|
||||
<collectingCountry id="09E57651FFA1950CFEE2FB14D3F85FB9" box="[316,404,1183,1208]" name="Mexico" pageId="20" pageNumber="171">Mexico</collectingCountry>
|
||||
, state of
|
||||
<collectingRegion id="B336F823FFA1950CFE26FB2BD0235FB9" box="[504,591,1184,1208]" country="Mexico" name="Oaxaca" pageId="20" pageNumber="171">Oaxaca</collectingRegion>
|
||||
, on
|
||||
<taxonomicName id="B6F24D42FFA1950CFDA3FB2BD1E85FB6" box="[637,900,1184,1207]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA1950CFDA3FB2BD1E85FB6" box="[637,900,1184,1207]" italics="true" pageId="20" pageNumber="171">Ceratozamia mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
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|
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<emphasis id="4386EAD3FFA1950CFF19FB4FD38E5FDC" bold="true" box="[199,482,1220,1245]" pageId="20" pageNumber="171">Adult male description.</emphasis>
|
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Length
|
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, width
|
||||
<quantity id="B60A9B24FFA1950CFCE2FB4FD18C5FDD" box="[828,992,1220,1244]" metricMagnitude="-3" metricUnit="m" metricValue="1.825" metricValueMax="1.91" metricValueMin="1.74" pageId="20" pageNumber="171" unit="mm" value="1.825" valueMax="1.91" valueMin="1.74">1.74–1.91 mm</quantity>
|
||||
(n = 7). Body in dorsal view elongateoval, greatest width at middle of elytra; in lateral view convex dorsally. General body color brown (
|
||||
<figureCitation id="E9C92A44FFA1950CFAD4FB6CD7345E01" box="[1290,1368,1255,1280]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="20" pageNumber="171">Figs. 7</figureCitation>
|
||||
A–E), head and pronotum concolorous and slightly darker than the yellowish-brown elytra; dorsal surface punctate, shining, short procumbent hairs associated with punctation, ventrally shining and appearing glabrous except mesoventrite and abdomen mostly covered with long procumbent setae.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA1950CFF19FAFCD3905C1D" blockId="20.[151,1437,1076,2037]" pageId="20" pageNumber="171">
|
||||
<emphasis id="4386EAD3FFA1950CFF19FAFCD36B5E91" bold="true" box="[199,263,1399,1424]" pageId="20" pageNumber="171">Head</emphasis>
|
||||
in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured (
|
||||
<figureCitation id="E9C92A44FFA1950CFEC9FA17D3CC5EB5" box="[279,416,1436,1461]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="20" pageNumber="171">Figs. 7A, D</figureCitation>
|
||||
), average distance between closest punctures 2× width of puncture; width
|
||||
<quantity id="B60A9B24FFA1950CFB2FFA17D7F45EB5" box="[1265,1432,1436,1461]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" metricValueMax="9.2" metricValueMin="8.6" pageId="20" pageNumber="171" unit="mm" value="0.89" valueMax="0.92" valueMin="0.86">0.86–0.92 mm</quantity>
|
||||
; dorsal interocular distance
|
||||
<quantity id="B60A9B24FFA1950CFE3DFA34D0FE5ED6" box="[483,658,1471,1496]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" metricValueMax="5.8" metricValueMin="5.4" pageId="20" pageNumber="171" unit="mm" value="0.56" valueMax="0.58" valueMin="0.54">0.54–0.58 mm</quantity>
|
||||
, head width/dorsal interocular distance ratio 1.49–1.61, ventral interocular distance
|
||||
<quantity id="B60A9B24FFA1950CFEA3FA6FD0485EFD" box="[381,548,1508,1532]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.699999999999999" metricValueMin="4.0" pageId="20" pageNumber="171" unit="mm" value="0.435" valueMax="0.47" valueMin="0.4">0.40–0.47 mm</quantity>
|
||||
, head width/ventral interocular distance ratio 2.13–2.50. Eye with large black facets. Antennal length approximately equal to pronotal width, 1.5× head width; antennomere I (scape) fairly large, slightly elongate, antennomere II subequal to III; IV–VII small, width equals length; VIII slightly wider than long; club fairly large, IX and X similar in length, XI slightly longer, globular with acuminate apex. Clypeus truncate anteriorly, moderately punctate, with narrow margin. Mentum and submentum with moderate punctation, distance between nearest punctures ~ 1× own width, each puncture with a long seta ~ 2–3× width of eye facet and reclining anteriad (often abraded). Gular area smooth, without punctation or setae, border with submentum usually with a shallow depression without punctation, but usually with a shallowly impressed circular or semicircular border along anterior margin (
|
||||
<figureCitation id="E9C92A44FFA1950CFE46F88FD3835C1D" box="[408,495,1796,1821]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="20" pageNumber="171">Fig. 7E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA1950DFF19F8ACD13559F1" blockId="20.[151,1437,1076,2037]" lastBlockId="21.[151,1437,151,1967]" lastPageId="21" lastPageNumber="172" pageId="20" pageNumber="171">
|
||||
<emphasis id="4386EAD3FFA1950CFF19F8ACD34C5C41" bold="true" box="[199,288,1831,1856]" pageId="20" pageNumber="171">Thorax</emphasis>
|
||||
with pronotum transverse in dorsal view with marginal beads basally and laterally, anterior margin with weak bead, reduced to absent medially; length/width ratio (PL/PW) = 0.70–0.74, rectangular; anterior and posterior angles present, anterior angles projecting; lateral carinae expanding in basal 1/10 to parallel sides, sometimes slightly emarginated, parallel for 50% of length, then gradually converging to anterior angles; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/5 length of posterior margin from posterior corners and touching and just anterior to marginal bead, pores mark base of longitudinal furrows onto disc. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae ~ 1/3 length of eye. Hypomeron laterally apparently lacking punctures, some with minute setae visible; medially with longitudinal striations. Elytra in dorsal view elongate-oval, convex; length/width 1.66–1.78, greatest width near midlength; without marginal line basally; 10 complete striae of moderate puncture size, scutellary striole extending ¼ elytral length, with 9–16 punctures; intervals of striae with fine, shallow punctures. All punctures of elytral bearing a single fine, procumbent seta; seta often only visible in profile, extending posteriorly to, over or beyond next puncture (often abraded), interval puncture setae longer than strial puncture setae. Punctation on meso- and metaventrites moderate to dense, distance between nearest punctures 1–2× width of punctures, puncture depth shallow. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over 3/4 metaventrite length. Legs with procoxa oval; mesocoxa globular; metacoxa transversely elongate-oval; trochanters obliquely truncate apically; in males all femora swollen, stout, dorso-ventral width greater than eye length; all tibiae with shallow medial subapical emargination, obliquely truncate at apex, with complete fringe of fine spinules on anterior and posterior margins; protibia swollen, not flattened in cross section notably larger than other tibiae, triangularly dilated to apex; mesotibia weakly swollen, moderately triangularly dilated to apex; metatibia narrowed, weakly dilating to midlength then nearly parallel to apex; pro- and mesotarsi greatly enlarged, corresponding tarsomere I width ~ 3× own length, metatarsomere I width ~ own length (
|
||||
<figureCitation id="E9C92A44FFA0950DFD16FD5CD17959F1" box="[712,789,727,752]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="21" pageNumber="172">Figs. 7</figureCitation>
|
||||
B–C).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA0950DFF19FD77D1A85E0D" blockId="21.[151,1437,151,1967]" pageId="21" pageNumber="172">
|
||||
<emphasis id="4386EAD3FFA0950DFF19FD77D32F5814" bold="true" box="[199,323,764,789]" pageId="21" pageNumber="172">Abdomen.</emphasis>
|
||||
Ventrite I with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; all ventrites strongly alutaceous laterally, less so and glossy medially, bearing moderate, shallow punctation, distance to nearest puncture ~ 1× width of puncture, punctures bearing mostly reclining setae; ventrites I–V with setae length ~ 2–3× width of puncture; I–V each with 2 or more median pairs of longer, semi-erect sensory hairs (often abraded), V with additional 4–5 pairs of long semi-erect hairs located postero-laterally, but not at submargin. Male genitalia with genital capsule fringed with fine setae; tegmen sclerotized, triangular, not laterally compressed; anterior region ring-like, posterior region sheath-like, 2–3× wider than deep; lateral margins gradually converging posteriorly (
|
||||
<figureCitation id="E9C92A44FFA0950DFEFDFBEFD31F5F7D" box="[291,371,1124,1149]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="21" pageNumber="172">Figs. 7</figureCitation>
|
||||
F–G); apically with 2 elongate, somewhat bar-shaped parameres. Parameres dorso-ventrally compressed, height approximately half its own width, in dorso-ventral view length/width ratio = 1.7–2.2; apically with setae, length of longest setae> width of paramere in dorso-ventral view (
|
||||
<figureCitation id="E9C92A44FFA0950DFBCFFB27D6005FC5" box="[1041,1132,1196,1221]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="21" pageNumber="172">Fig. 7H</figureCitation>
|
||||
); median lobe not laterally compressed, somewhat cylindrical, sclerotized, in lateral view weakly curved, apex tapering to a point. Ratio of penile strut length to median lobe length 2.11–2.41 (n = 2) (
|
||||
<figureCitation id="E9C92A44FFA0950DFCEAFB7FD1EC5E0D" box="[820,896,1268,1293]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="21" pageNumber="172">Figs. 7</figureCitation>
|
||||
F–G).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA0950DFF19FA9CD1E15EBF" blockId="21.[151,1437,151,1967]" pageId="21" pageNumber="172">
|
||||
<emphasis id="4386EAD3FFA0950DFF19FA9CD3725E31" bold="true" box="[199,286,1303,1328]" pageId="21" pageNumber="172">Female</emphasis>
|
||||
generally similar to male, except for sexual dimorphism. Female pronotum slightly more transverse than male, PL/PW = 0.65–0.69; elytra dulled (alutaceous) laterally; legs unmodified, fore and middle legs not swollen, femora dorso-ventral width about equal to eye, all tibiae gradually dilating to narrowly triangular apex; tibiae lacking median subapical emarginations; all legs with tarsomere I width ~ own length; metaventrite flattened medially, not impressed; abdomen less densely punctate and setose.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA0950DFF19FA42D7FA5DB6" pageId="21" pageNumber="172" type="materials_examined">
|
||||
<paragraph id="714D36C1FFA0950DFF19FA42D63A5EE3" blockId="21.[151,1437,151,1967]" box="[199,1110,1481,1506]" pageId="21" pageNumber="172">
|
||||
<materialsCitation id="C19A3C9CFFA0950DFF19FA42D63F5EE3" ID-GBIF-Occurrence="2028138739" box="[199,1107,1481,1506]" country="Mexico" elevation="1918" location="Sierra Mixe" municipality="State" pageId="21" pageNumber="172" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
|
||||
<emphasis id="4386EAD3FFA0950DFF19FA42D3095EE3" bold="true" box="[199,357,1481,1506]" pageId="21" pageNumber="172">
|
||||
<typeStatus id="AE498863FFA0950DFF19FA42D36E5EE3" box="[199,258,1481,1506]" pageId="21" pageNumber="172">Type</typeStatus>
|
||||
locality.
|
||||
</emphasis>
|
||||
<collectingCountry id="09E57651FFA0950DFEB2FA42D3A85EE3" box="[364,452,1481,1506]" name="Mexico" pageId="21" pageNumber="172">Mexico</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFA0950DFE11FA41D04F5EE3" box="[463,547,1482,1506]" country="Mexico" name="Oaxaca" pageId="21" pageNumber="172">Oaxaca</collectingRegion>
|
||||
<collectingMunicipality id="9129ACBBFFA0950DFDF4FA41D0095EE3" box="[554,613,1482,1506]" pageId="21" pageNumber="172">State</collectingMunicipality>
|
||||
,
|
||||
<location id="742D601AFFA0950DFDAEFA42D0945EE3" LSID="urn:lsid:plazi:treatment:F95B87D7FFA1950DFF49FC42D6275CAF:742D601AFFA0950DFDAEFA42D0945EE3" box="[624,760,1481,1506]" country="Mexico" municipality="State" name="Sierra Mixe" pageId="21" pageNumber="172" stateProvince="Oaxaca">Sierra Mixe</location>
|
||||
, [GPS coord. omitted],
|
||||
<elevation id="FADFD1F2FFA0950DFBDFFA41D63F5EE3" box="[1025,1107,1482,1506]" metricMagnitude="3" metricUnit="m" metricValue="1.918" pageId="21" pageNumber="172" unit="m" value="1918.0">
|
||||
<quantity id="B60A9B24FFA0950DFBDFFA41D63F5EE3" box="[1025,1107,1482,1506]" metricMagnitude="3" metricUnit="m" metricValue="1.918" pageId="21" pageNumber="172" unit="m" value="1918.0">1918m</quantity>
|
||||
</elevation>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA0950DFF19FA67D70B5D05" blockId="21.[151,1437,151,1967]" box="[199,1383,1516,1541]" pageId="21" pageNumber="172">
|
||||
<emphasis id="4386EAD3FFA0950DFF19FA67D3765D04" bold="true" box="[199,282,1516,1541]" pageId="21" pageNumber="172">Range.</emphasis>
|
||||
<collectingRegion id="B336F823FFA0950DFEFFFA66D3145D04" box="[289,376,1517,1541]" country="Mexico" name="Oaxaca" pageId="21" pageNumber="172">Oaxaca</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="09E57651FFA0950DFE5DFA67D3B45D04" box="[387,472,1516,1541]" name="Mexico" pageId="21" pageNumber="172">Mexico</collectingCountry>
|
||||
in the Sierra Mixe, known to inhabit only male cones of
|
||||
<taxonomicName id="B6F24D42FFA0950DFB86FA66D70C5D05" box="[1112,1376,1517,1540]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA0950DFB86FA66D70C5D05" box="[1112,1376,1517,1540]" italics="true" pageId="21" pageNumber="172">Ceratozamia mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA0950DFF19F99BD7FA5DB6" blockId="21.[151,1437,151,1967]" pageId="21" pageNumber="172">
|
||||
<emphasis id="4386EAD3FFA0950DFF19F99BD04A5D28" bold="true" box="[199,550,1552,1577]" pageId="21" pageNumber="172">
|
||||
Material examined.
|
||||
<materialsCitation id="C19A3C9CFFA0950DFE67F99BD04A5D28" ID-GBIF-Occurrence="2028138749" box="[441,550,1552,1577]" collectingDate="2016-04-04" collectorName="W. Tang" country="Mexico" elevation="1918" location="Sierra Mixe" pageId="21" pageNumber="172" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="AE498863FFA0950DFE67F99BD04A5D28" box="[441,550,1552,1577]" pageId="21" pageNumber="172" type="holotype">Holotype</typeStatus>
|
||||
</materialsCitation>
|
||||
</emphasis>
|
||||
(by designation) male
|
||||
<taxonomicName id="B6F24D42FFA0950DFCF5F99AD1D05D29" box="[811,956,1553,1576]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="species" species="mixeorum">
|
||||
<emphasis id="4386EAD3FFA0950DFCF5F99AD1D05D29" box="[811,956,1553,1576]" italics="true" pageId="21" pageNumber="172">C. mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||||
<collectingCountry id="09E57651FFA0950DFE22F9BFD0035D4D" box="[508,623,1588,1612]" name="Mexico" pageId="21" pageNumber="172">MEXICO</collectingCountry>
|
||||
, Oax.,
|
||||
<location id="742D601AFFA0950DFD12F9B8D1375D4D" LSID="urn:lsid:plazi:treatment:F95B87D7FFA1950DFF49FC42D6275CAF:742D601AFFA0950DFD12F9B8D1375D4D" box="[716,859,1587,1612]" country="Mexico" name="Sierra Mixe" pageId="21" pageNumber="172">Sierra Mixe</location>
|
||||
, [GPS coord. omitted],
|
||||
<elevation id="FADFD1F2FFA0950DFB5EF9BFD6BD5D4D" box="[1152,1233,1588,1612]" metricMagnitude="3" metricUnit="m" metricValue="1.918" pageId="21" pageNumber="172" unit="m" value="1918.0">
|
||||
<quantity id="B60A9B24FFA0950DFB5EF9BFD6BD5D4D" box="[1152,1233,1588,1612]" metricMagnitude="3" metricUnit="m" metricValue="1.918" pageId="21" pageNumber="172" unit="m" value="1918.0">1918m</quantity>
|
||||
</elevation>
|
||||
, ex
|
||||
<taxonomicName id="B6F24D42FFA0950DFAD5F9BFD36B5D6E" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA0950DFAD5F9BFD36B5D6E" italics="true" pageId="21" pageNumber="172">Ceratozamia mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
<specimenCount id="67F4FD48FFA0950DFECEF9D3D3495D6E" box="[272,293,1624,1647]" pageId="21" pageNumber="172" type="male">♂</specimenCount>
|
||||
cone,
|
||||
<collectingDate id="1508E9E9FFA0950DFEAAF9DCD3865D71" box="[372,490,1623,1648]" pageId="21" pageNumber="172" value="2016-04-04">
|
||||
<date id="054C1001FFA0950DFEAAF9DCD3865D71" box="[372,490,1623,1648]" pageId="21" pageNumber="172" value="2016-04-04">4-IV-2016</date>
|
||||
</collectingDate>
|
||||
,
|
||||
<collectorName id="DC075317FFA0950DFE26F9D3D0315D6E" box="[504,605,1623,1647]" pageId="21" pageNumber="172">W. Tang</collectorName>
|
||||
”; 2) [rectangular; red; printed in black ink]
|
||||
<materialsCitation id="C19A3C9CFFA0950DFB87F9DCD1D05D92" ID-GBIF-Occurrence="2028138750" collectionCode="FSCA" country="Mexico" pageId="21" pageNumber="172" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="AE498863FFA0950DFB87F9DCD6815D6E" box="[1113,1261,1623,1647]" pageId="21" pageNumber="172" type="holotype">HOLOTYPE</typeStatus>
|
||||
<specimenCount id="67F4FD48FFA0950DFB28F9D3D7675D6E" box="[1270,1291,1624,1647]" pageId="21" pageNumber="172" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFA0950DFACAF9DCD0C45D92" authority="Tang, Skelley & Perez-Farrera 2018" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA0950DFACAF9DCD36B5D93" italics="true" pageId="21" pageNumber="172">Ceratophila mixeorum</emphasis>
|
||||
<bibRefCitation id="15634B30FFA0950DFED0F9F0D0C45D92" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." box="[270,680,1658,1683]" pageId="21" pageNumber="172" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">Tang, Skelley & Pérez-Farrera 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. Deposited in the
|
||||
<collectionCode id="17E3AE04FFA0950DFCACF9F0D1D05D92" box="[882,956,1659,1683]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="172" type="Museum">FSCA</collectionCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="C19A3C9CFFA0950DFC19F9F1D7FE5DB6" ID-GBIF-Occurrence="2028138743" collectionCode="FSCA, ANIC, BMNH, IEXA, INBio, MNHN, NZAC, UNAM, USNM" country="Mexico" pageId="21" pageNumber="172" specimenCount="43" specimenCount-adult="41" typeStatus="allotype">
|
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<typeStatus id="AE498863FFA0950DFC19F9F1D6405D92" box="[967,1068,1658,1683]" pageId="21" pageNumber="172" type="allotype">
|
||||
<emphasis id="4386EAD3FFA0950DFC19F9F1D6405D92" bold="true" box="[967,1068,1658,1683]" pageId="21" pageNumber="172">Allotype</emphasis>
|
||||
</typeStatus>
|
||||
(
|
||||
<collectionCode id="17E3AE04FFA0950DFBE5F9F0D6E85D92" box="[1083,1156,1659,1683]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="172" type="Museum">FSCA</collectionCode>
|
||||
) and
|
||||
<specimenCount id="67F4FD48FFA0950DFB1DF9F0D7725D92" box="[1219,1310,1658,1683]" pageId="21" pageNumber="172" type="adult">41 adult</specimenCount>
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<typeStatus id="AE498863FFA0950DFAF8F9F1D7F05D92" box="[1318,1436,1658,1683]" pageId="21" pageNumber="172" type="paratype">
|
||||
<emphasis id="4386EAD3FFA0950DFAF8F9F1D7F05D92" bold="true" box="[1318,1436,1658,1683]" pageId="21" pageNumber="172">paratypes</emphasis>
|
||||
</typeStatus>
|
||||
(same label data as
|
||||
<typeStatus id="AE498863FFA0950DFEB1F915D3BB5DB6" box="[367,471,1694,1719]" pageId="21" pageNumber="172" type="holotype">holotype</typeStatus>
|
||||
): deposited at
|
||||
<collectionCode id="17E3AE04FFA0950DFDABF914D0D05DB6" box="[629,700,1695,1719]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="172">ANIC</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFA0950DFD19F914D1725DB7" box="[711,798,1695,1718]" pageId="21" pageNumber="172">BMNH</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFA0950DFCF6F914D11D5DB6" box="[808,881,1695,1719]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="172" type="Museum">FSCA</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFA0950DFCA5F914D1AC5DB6" box="[891,960,1695,1719]" pageId="21" pageNumber="172">IEXA</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFA0950DFC15F915D6785DB6" box="[971,1044,1694,1719]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="21" pageNumber="172">INBio</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFA0950DFBC1F914D6145DB7" box="[1055,1144,1695,1718]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="172">MNHN</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFA0950DFB5CF914D6BD5DB6" box="[1154,1233,1695,1719]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="21" pageNumber="172" type="Other institutional type">NZAC</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFA0950DFB05F914D75F5DB6" box="[1243,1331,1695,1719]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="21" pageNumber="172">UNAM</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFA0950DFAE0F914D7FE5DB6" box="[1342,1426,1695,1719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="172" type="Museum">USNM</collectionCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA0950DFF19F94AD37F5DFF" pageId="21" pageNumber="172" type="etymology">
|
||||
<paragraph id="714D36C1FFA0950DFF19F94AD37F5DFF" blockId="21.[151,1437,151,1967]" pageId="21" pageNumber="172">
|
||||
<emphasis id="4386EAD3FFA0950DFF19F94AD32A5DDB" bold="true" box="[199,326,1729,1754]" pageId="21" pageNumber="172">Etymology</emphasis>
|
||||
. This species is named for the host species on which it was discovered, applied as a noun in apposition.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA0950DFF19F883D6275CAF" pageId="21" pageNumber="172" type="discussion">
|
||||
<paragraph id="714D36C1FFA0950DFF19F883D6275CAF" blockId="21.[151,1437,151,1967]" pageId="21" pageNumber="172">
|
||||
<emphasis id="4386EAD3FFA0950DFF19F883D3575C20" bold="true" box="[199,315,1800,1825]" pageId="21" pageNumber="172">Remarks.</emphasis>
|
||||
The only known host of this beetle is
|
||||
<taxonomicName id="B6F24D42FFA0950DFD22F882D6695C21" box="[764,1029,1801,1824]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA0950DFD22F882D6695C21" box="[764,1029,1801,1824]" italics="true" pageId="21" pageNumber="172">Ceratozamia mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
. This cycad inhabits isolated cloud forest in the eastern Sierra Mixe where it occurs together with
|
||||
<emphasis id="4386EAD3FFA0950DFC4FF8A7D1C75C42" box="[913,939,1836,1859]" italics="true" pageId="21" pageNumber="172">C.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFA0950DFC69F8A7D6235C45" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[951,1103,1836,1860]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="species" species="gregoryi" subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFA0950DFC1FF8A7D1B85C42" box="[961,980,1836,1859]" italics="true" pageId="21" pageNumber="172">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFA0950DFC2EF8A6D6235C45" box="[1008,1103,1837,1860]" italics="true" pageId="21" pageNumber="172">gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
(see remarks under
|
||||
<emphasis id="4386EAD3FFA0950DFA97F8A7D70F5C42" box="[1353,1379,1836,1859]" italics="true" pageId="21" pageNumber="172">C.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFA0950DFAB1F8A7D29A5C66" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="species" species="gregoryi" subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFA0950DFAA7F8A7D7E05C42" box="[1401,1420,1836,1859]" italics="true" pageId="21" pageNumber="172">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFA0950DFF49F8DBD29A5C66" box="[151,246,1872,1895]" italics="true" pageId="21" pageNumber="172">gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
). In two cones sampled
|
||||
<emphasis id="4386EAD3FFA0950DFDF4F8DBD0515C66" box="[554,573,1872,1895]" italics="true" pageId="21" pageNumber="172">C</emphasis>
|
||||
. (
|
||||
<taxonomicName id="B6F24D42FFA0950DFD84F8C4D0D75C66" box="[602,699,1871,1895]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFA0950DFD84F8C4D0D75C66" box="[602,699,1871,1895]" italics="true" pageId="21" pageNumber="172">Vovidesa</emphasis>
|
||||
</taxonomicName>
|
||||
) accounted for 16% of
|
||||
<specimenCount id="67F4FD48FFA0950DFC37F8DBD6345C69" box="[1001,1112,1871,1896]" pageId="21" pageNumber="172" type="adult">273 adult</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFA0950DFBB8F8C4D74E5C69" authorityName="Crowson" authorityYear="1952" box="[1126,1314,1871,1896]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="172" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||||
collected, compared to 84% for
|
||||
<emphasis id="4386EAD3FFA0950DFE4DF8F8D3CA5C8B" box="[403,422,1907,1930]" italics="true" pageId="21" pageNumber="172">C</emphasis>
|
||||
.
|
||||
<taxonomicName id="B6F24D42FFA0950DFE6BF8F8D0265C8A" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[437,586,1907,1931]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="species" species="gregoryi" subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFA0950DFE61F8F8D3BE5C8B" box="[447,466,1907,1930]" italics="true" pageId="21" pageNumber="172">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFA0950DFE35F8FFD0265C8A" box="[491,586,1908,1931]" italics="true" pageId="21" pageNumber="172">gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
that coinhabit cones. In other species of
|
||||
<taxonomicName id="B6F24D42FFA0950DFBFAF8F8D6D95C8B" box="[1060,1205,1907,1930]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA0950DFBFAF8F8D6D95C8B" box="[1060,1205,1907,1930]" italics="true" pageId="21" pageNumber="172">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
adult individuals of subgenus
|
||||
<taxonomicName id="B6F24D42FFA0950DFED8F81DD30A5CAF" box="[262,358,1942,1966]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="21" pageNumber="172" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFA0950DFED8F81DD30A5CAF" box="[262,358,1942,1966]" italics="true" pageId="21" pageNumber="172">Vovidesa</emphasis>
|
||||
</taxonomicName>
|
||||
typically make up a smaller fraction of the adult beetles in cones.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
386
data/F9/5B/87/F95B87D7FFA29501FF49FF1CD3BF5C05.xml
Normal file
386
data/F9/5B/87/F95B87D7FFA29501FF49FF1CD3BF5C05.xml
Normal file
|
|
@ -0,0 +1,386 @@
|
|||
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|
||||
<paragraph id="714D36C1FFA2950FFF49FF1CD6785BB1" blockId="23.[151,1044,151,213]" box="[151,1044,151,176]" pageId="23" pageNumber="174">
|
||||
<heading id="2A0581ADFFA2950FFF49FF1CD6785BB1" bold="true" box="[151,1044,151,176]" fontSize="10" level="2" pageId="23" pageNumber="174" reason="2">
|
||||
<emphasis id="4386EAD3FFA2950FFF49FF1CD6785BB1" bold="true" box="[151,1044,151,176]" pageId="23" pageNumber="174">
|
||||
<taxonomicName id="B6F24D42FFA2950FFF49FF1CD1135BB1" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,895,151,176]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="23" pageNumber="174" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFA2950FFF49FF1CD34C5BB1" bold="true" box="[151,288,151,176]" italics="true" pageId="23" pageNumber="174">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA2950FFEEEFF1CD3FF5BB1" bold="true" box="[304,403,151,176]" italics="true" pageId="23" pageNumber="174">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFA2950FFE7BFF1CD06B5BB1" bold="true" box="[421,519,151,176]" italics="true" pageId="23" pageNumber="174">vazquezi</emphasis>
|
||||
Tang, Skelley & Pérez-Farrera
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFA2950FFC55FF1CD6785BB1" box="[907,1044,151,176]" pageId="23" pageNumber="174" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA2950FFF49FF37D3405BD5" box="[151,300,188,213]" pageId="23" pageNumber="174" type="description">
|
||||
<paragraph id="714D36C1FFA2950FFF49FF37D3405BD5" blockId="23.[151,1044,151,213]" box="[151,300,188,213]" pageId="23" pageNumber="174">
|
||||
(
|
||||
<figureCitation id="E9C92A44FFA2950FFF41FF37D2865BD5" box="[159,234,188,213]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Figs. 8</figureCitation>
|
||||
A–H)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA2950FFF49FE8FD6FB5A86" pageId="23" pageNumber="174" type="diagnosis">
|
||||
<paragraph id="714D36C1FFA2950FFF49FE8FD6FB5A86" blockId="23.[151,1437,260,1869]" pageId="23" pageNumber="174">
|
||||
<emphasis id="4386EAD3FFA2950FFF49FE8FD3365A1C" bold="true" box="[151,346,260,285]" pageId="23" pageNumber="174">Adult diagnosis.</emphasis>
|
||||
Distinguished from other
|
||||
<taxonomicName id="B6F24D42FFA2950FFD58FE8FD1E15A1D" authority="(Vovidesa )" baseAuthorityName="Vovidesa" box="[646,909,260,284]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="23" pageNumber="155" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA2950FFD58FE8FD1615A1D" box="[646,781,260,284]" italics="true" pageId="23" pageNumber="174">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA2950FFCC1FE8FD1EC5A1D" box="[799,896,260,284]" italics="true" pageId="23" pageNumber="174">Vovidesa</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
species by male with subapical emarginations on tibiae, submentum of both sexes with dense punctures and setae, male submental setae projecting laterally, female elytra similarly dulled medially and laterally, male genitalia with median lobe nearly laterally flattened and curved, and with a known distribution in
|
||||
<collectingCountry id="09E57651FFA2950FFDBBFEE4D0D15A89" box="[613,701,367,392]" name="Mexico" pageId="23" pageNumber="174">Mexico</collectingCountry>
|
||||
, state of
|
||||
<collectingRegion id="B336F823FFA2950FFCFFFEFBD1EB5A86" box="[801,903,368,391]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="23" pageNumber="174">Veracruz</collectingRegion>
|
||||
, on
|
||||
<emphasis id="4386EAD3FFA2950FFC6BFEFBD6FB5A86" box="[949,1175,368,391]" italics="true" pageId="23" pageNumber="174">
|
||||
<taxonomicName id="B6F24D42FFA2950FFC6BFEFBD6FF5A86" box="[949,1171,368,391]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="23" pageNumber="174" phylum="Tracheophyta" rank="species" species="tenuis">Ceratozamia tenuis</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA29501FF19FE1FD28F587E" lastPageId="25" lastPageNumber="176" pageId="23" pageNumber="174" type="description">
|
||||
<paragraph id="714D36C1FFA2950FFF19FE1FD1D0593D" blockId="23.[151,1437,260,1869]" pageId="23" pageNumber="174">
|
||||
<emphasis id="4386EAD3FFA2950FFF19FE1FD38C5AAC" bold="true" box="[199,480,404,429]" pageId="23" pageNumber="174">Adult male description.</emphasis>
|
||||
Length
|
||||
<quantity id="B60A9B24FFA2950FFDE0FE1FD08F5AAD" box="[574,739,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="4.51" metricValueMax="4.9" metricValueMin="4.12" pageId="23" pageNumber="174" unit="mm" value="4.51" valueMax="4.9" valueMin="4.12">4.12–4.90 mm</quantity>
|
||||
, width
|
||||
<quantity id="B60A9B24FFA2950FFCE8FE1FD1B65AAD" box="[822,986,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.705" metricValueMax="1.84" metricValueMin="1.57" pageId="23" pageNumber="174" unit="mm" value="1.705" valueMax="1.84" valueMin="1.57">1.57–1.84 mm</quantity>
|
||||
(n = 10). Body in dorsal view elongateoval, greatest width at middle of elytra; in lateral view convex dorsally. General body color brown (
|
||||
<figureCitation id="E9C92A44FFA2950FFAD5FE3CD7345AD1" box="[1291,1368,439,464]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Figs. 8</figureCitation>
|
||||
A–F), head and pronotum concolorous and slightly darker than the yellowish-brown elytra; dorsal surface punctate, shining, short procumbent hairs associated with punctation, ventrally shining and appearing glabrous except mesoventrite and abdomen mostly covered with long procumbent setae.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA2950FFF19FDCCD3DB58ED" blockId="23.[151,1437,260,1869]" pageId="23" pageNumber="174">
|
||||
<emphasis id="4386EAD3FFA2950FFF19FDCCD36B5961" bold="true" box="[199,263,583,608]" pageId="23" pageNumber="174">Head</emphasis>
|
||||
in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured (
|
||||
<figureCitation id="E9C92A44FFA2950FFEC9FDE7D3CC5985" box="[279,416,620,645]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Figs. 8A, D</figureCitation>
|
||||
), average distance between closest punctures 2× width of puncture; width
|
||||
<quantity id="B60A9B24FFA2950FFB2FFDE7D7F45985" box="[1265,1432,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" metricValueMax="9.1" metricValueMin="8.1" pageId="23" pageNumber="174" unit="mm" value="0.86" valueMax="0.91" valueMin="0.81">0.81–0.91 mm</quantity>
|
||||
; dorsal interocular distance
|
||||
<quantity id="B60A9B24FFA2950FFE3DFD04D0FE59A6" box="[483,658,655,680]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.8999999999999995" metricValueMin="5.2" pageId="23" pageNumber="174" unit="mm" value="0.5549999999999999" valueMax="0.59" valueMin="0.52">0.52–0.59 mm</quantity>
|
||||
, head width/dorsal interocular distance ratio 1.45–1.56, ventral interocular distance
|
||||
<quantity id="B60A9B24FFA2950FFEA3FD3FD04859CD" box="[381,548,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="23" pageNumber="174" unit="mm" value="0.4" valueMax="0.42" valueMin="0.38">0.38–0.42 mm</quantity>
|
||||
, head width/ventral interocular distance ratio 2.04–2.17. Eye with large black facets. Antennal length approximately equal to pronotal width, 1.5× head width; antennomere I (scape) fairly large, slightly elongate, antennomere II subequal to III; IV–VIII small, width equals length; club fairly large, IX and X similar in length, XI slightly longer, globular with acuminate apex. Clypeus truncate anteriorly, moderately punctate, with narrow margin. Mentum with moderate punctation, distance between nearest punctures ~ 1× own width; submentum finely, densely punctate, each puncture with a long seta ~ 2–3× width of eye facet, setae projecting laterally. Gular area smooth, without punctation or setae, border with submentum usually with a shallow depression without punctation, but usually with a shallowly impressed circular or semicircular border along anterior margin (
|
||||
<figureCitation id="E9C92A44FFA2950FFE8DFC5FD3C658ED" box="[339,426,980,1005]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Fig. 8E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA2950FFF19FC7CD6665C4D" blockId="23.[151,1437,260,1869]" pageId="23" pageNumber="174">
|
||||
<emphasis id="4386EAD3FFA2950FFF19FC7CD34C5F11" bold="true" box="[199,288,1015,1040]" pageId="23" pageNumber="174">Thorax</emphasis>
|
||||
with pronotum transverse in dorsal view with marginal beads basally and laterally, anterior margin with weak bead, reduced to absent medially; length/width ratio (PL/PW) = 0.76–0.79, rectangular; anterior and posterior angles present, anterior angles projecting; lateral carinae expanding in basal 1/10 to parallel sides, sometimes slightly emarginated, parallel for 50% of length, then gradually converging to anterior angles; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/5 length of posterior margin from posterior corners and touching and just anterior to marginal bead, pores mark base of longitudinal furrows onto disc. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae ~ 1/3 length of eye. Hypomeron laterally apparently lacking punctures, some with minute setae visible; medially with longitudinal striations. Elytra in dorsal view elongate-oval, convex; length/width 1.75–1.85, greatest width near midlength; without marginal line basally; 10 complete striae of moderate puncture size, scutellary striole extending ¼ elytral length, with 9–16 punctures; intervals of striae with fine, shallow punctures of similar size to strial punctures. All punctures of elytral bearing a single fine, procumbent seta; seta often only visible in profile, extending posteriorly to, over or beyond next puncture (often abraded), interval puncture setae longer than strial puncture setae. Punctation on meso- and metaventrites moderate to dense, distance between nearest punctures 1–2× width of punctures, puncture depth shallow. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over 3/4 metaventrite length. Legs with procoxa oval; mesocoxa globular; metacoxa transversely elongateoval; trochanters obliquely truncate apically; in males all femora swollen, stout, dorso-ventral width greater than eye length; protibia with shallow medial subapical emargination, meso- and metatibia with a distinct medial subapical emargination; all tibiae obliquely truncate at apex, with complete fringe of fine spinules on anterior and posterior margins; protibia swollen, not flattened in cross section notably larger than other tibiae, triangularly dilated to apex; mesotibia weakly swollen, moderately triangularly dilated to apex; metatibia narrowed, weakly dilating to midlength then slightly narrowing to apex; pro- and mesotarsi greatly enlarged, corresponding tarsomere I width ~ 3× own length, metatarsomere I width ~ own length (
|
||||
<figureCitation id="E9C92A44FFA2950FFC84F8BFD1C95C4D" box="[858,933,1844,1869]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="23" pageNumber="174">Figs. 8</figureCitation>
|
||||
A–C, F).
|
||||
</paragraph>
|
||||
<caption id="258D6649FFAD9500FF49F805D7F05CE8" httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="24" pageNumber="175" startId="24.[151,250,1934,1956]" targetBox="[175,1416,193,1915]" targetPageId="24">
|
||||
<paragraph id="714D36C1FFAD9500FF49F805D7F05CE8" blockId="24.[151,1436,1934,2025]" pageId="24" pageNumber="175">
|
||||
<emphasis id="4386EAD3FFAD9500FF49F805D3785CA5" bold="true" box="[151,276,1934,1956]" pageId="24" pageNumber="175">FIGURE 8.</emphasis>
|
||||
Photographs of
|
||||
<taxonomicName id="B6F24D42FFAD9500FE1FF805D1625CA4" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[449,782,1934,1957]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="24" pageNumber="175" phylum="Tracheophyta" rank="subGenus" subGenus="Uovidesa">
|
||||
<emphasis id="4386EAD3FFAD9500FE1FF805D0505CA5" box="[449,572,1934,1956]" italics="true" pageId="24" pageNumber="175">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFAD9500FD92F805D0C85CA5" box="[588,676,1934,1956]" italics="true" pageId="24" pageNumber="175">Uovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFAD9500FD69F81BD1625CA4" box="[695,782,1936,1957]" italics="true" pageId="24" pageNumber="175">vazquezi</emphasis>
|
||||
</taxonomicName>
|
||||
(A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="714D36C1FFAC9501FF19FF1CD0BB59A9" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||||
<emphasis id="4386EAD3FFAC9501FF19FF1CD32F5BB1" bold="true" box="[199,323,151,176]" pageId="25" pageNumber="176">Abdomen.</emphasis>
|
||||
Ventrite I with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; all ventrites strongly alutaceous laterally, less so and glossy medially, bearing dense, moderate, shallow punctation, distance to nearest puncture ~ 1× width of puncture, punctures bearing mostly reclining setae; ventrites I–V with setae length ~ 2–3× width of puncture; I–V each with 2 or more median pairs of longer, semi-erect sensory hairs obscured in dense setation, V with additional 4–5 pairs of long semi-erect hairs located postero-laterally, but not at submargin. Male genitalia with genital capsule fringed with fine setae; tegmen sclerotized, triangular, laterally compressed; anterior region ring-like, posterior region sheath-like; lateral margins gradually converging posteriorly (
|
||||
<figureCitation id="E9C92A44FFAC9501FEFDFE74D31E5919" box="[291,370,511,536]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="25" pageNumber="176">Figs. 8</figureCitation>
|
||||
F–G); apically with 2 elongate, somewhat bar-shaped parameres. Parameres dorso-ventrally compressed, height approximately half its own width, in dorso-ventral view length/width ratio = 2.1–2.8; apically with setae, length of longest setae> width of paramere in dorso-ventral view (
|
||||
<figureCitation id="E9C92A44FFAC9501FBECFDCCD6FC5961" box="[1074,1168,583,608]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="25" pageNumber="176">Fig. 8H</figureCitation>
|
||||
); median lobe laterally compressed, sclerotized, in lateral view strongly curved, apex tapering to a point. Ratio of penile strut length to median lobe length 1.82–1.83 (n = 2) (
|
||||
<figureCitation id="E9C92A44FFAC9501FD99FD04D0FE59A9" box="[583,658,655,680]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="25" pageNumber="176">Figs. 8</figureCitation>
|
||||
F–G).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFAC9501FF19FD3FD28F587E" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||||
<emphasis id="4386EAD3FFAC9501FF19FD3FD37259CC" bold="true" box="[199,286,692,717]" pageId="25" pageNumber="176">Female</emphasis>
|
||||
generally similar to male, except for sexual dimorphism. Female pronotum slightly wider than male, PL/PW = 0.70–0.76; submentum densely punctate, setae short, not projecting; elytral surface dulled medially and laterally; legs unmodified, fore and middle legs not swollen, femora dorso-ventral width about equal to eye length, all tibia gradually dilating to narrowly triangular apex; tibia lacking median subapical emarginations; all legs with tarsomere I width ~ own length; metaventrite flattened medially, not impressed; abdomen less densely punctate and setose.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFAC9501FF19FC07D2835E31" pageId="25" pageNumber="176" type="materials_examined">
|
||||
<paragraph id="714D36C1FFAC9501FF19FC07D15258A4" blockId="25.[151,1437,151,1796]" box="[199,830,908,933]" pageId="25" pageNumber="176">
|
||||
<materialsCitation id="C19A3C9CFFAC9501FF19FC07D15558A4" ID-GBIF-Occurrence="2028138738" box="[199,825,908,933]" collectorName="El Esquilon" country="Mexico" location="Veracruz" pageId="25" pageNumber="176" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
|
||||
<emphasis id="4386EAD3FFAC9501FF19FC07D30958A4" bold="true" box="[199,357,908,933]" pageId="25" pageNumber="176">
|
||||
<typeStatus id="AE498863FFAC9501FF19FC07D36E58A4" box="[199,258,908,933]" pageId="25" pageNumber="176">Type</typeStatus>
|
||||
locality.
|
||||
</emphasis>
|
||||
<collectingCountry id="09E57651FFAC9501FEB2FC07D3A858A4" box="[364,452,908,933]" name="Mexico" pageId="25" pageNumber="176">Mexico</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFAC9501FE11FC06D05A58A5" box="[463,566,909,932]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||||
, Jilotepec,
|
||||
<collectorName id="DC075317FFAC9501FD6DFC07D15558A4" box="[691,825,908,933]" pageId="25" pageNumber="176">El Esquilón</collectorName>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFAC9501FF19FC24D1A558EC" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||||
<emphasis id="4386EAD3FFAC9501FF19FC24D37658C9" bold="true" box="[199,282,943,968]" pageId="25" pageNumber="176">Range.</emphasis>
|
||||
As currently understood, this species is known to inhabit
|
||||
<taxonomicName id="B6F24D42FFAC9501FC15FC3BD29858EC" authority="(Dyer) D. W. Stev. & Vovides" authorityName="D. W. Stev. & Vovides" baseAuthorityName="Dyer" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFAC9501FC15FC3BD6C658C6" box="[971,1194,944,967]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||||
(Dyer) D.W.Stev. & Vovides
|
||||
</taxonomicName>
|
||||
, which occurs in the vicinity north of Xalapa,
|
||||
<collectingRegion id="B336F823FFAC9501FD24FC5ED10D58ED" box="[762,865,981,1004]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="09E57651FFAC9501FCB2FC5FD1A958EC" box="[876,965,980,1005]" name="Mexico" pageId="25" pageNumber="176">Mexico</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFAC9501FF19FC7CD2835E31" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||||
<emphasis id="4386EAD3FFAC9501FF19FC7CD04F5F11" bold="true" box="[199,547,1015,1040]" pageId="25" pageNumber="176">
|
||||
Material examined.
|
||||
<typeStatus id="AE498863FFAC9501FE68FC7CD04F5F11" box="[438,547,1015,1040]" pageId="25" pageNumber="176" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
(by designation) male of
|
||||
<taxonomicName id="B6F24D42FFAC9501FC9CFC73D1AF5F0E" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[834,963,1016,1039]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="vazquezi">
|
||||
<emphasis id="4386EAD3FFAC9501FC9CFC73D1AF5F0E" box="[834,963,1016,1039]" italics="true" pageId="25" pageNumber="176">C. vazquezi</emphasis>
|
||||
</taxonomicName>
|
||||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||||
<collectingCountry id="09E57651FFAC9501FE05FB97D0235F35" box="[475,591,1052,1076]" name="Mexico" pageId="25" pageNumber="176">MEXICO</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFAC9501FD84FB96D0AD5F35" box="[602,705,1053,1076]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||||
, Cerro Coacoatzintla,
|
||||
<specimenCount id="67F4FD48FFAC9501FC65FB96D1BC5F35" box="[955,976,1053,1076]" pageId="25" pageNumber="176" type="male">♂</specimenCount>
|
||||
cone
|
||||
<taxonomicName id="B6F24D42FFAC9501FBCDFB97D6825F35" box="[1043,1262,1052,1076]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFAC9501FBCDFB97D6825F35" box="[1043,1262,1052,1076]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFAC9501FB22FB97D7275F35" box="[1276,1355,1052,1076]" pageId="25" pageNumber="176" value="1928-02-01">28-II-1</date>
|
||||
995, Y. Sanchez Tinoco”; 2) [rectangular; red; printed in black ink]
|
||||
<typeStatus id="AE498863FFAC9501FCE6FBCBD1A05F59" box="[824,972,1088,1112]" pageId="25" pageNumber="176" type="holotype">HOLOTYPE</typeStatus>
|
||||
<specimenCount id="67F4FD48FFAC9501FC0DFBCBD1845F56" box="[979,1000,1088,1111]" pageId="25" pageNumber="176" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFAC9501FC31FBCBD3195F7D" authority="Tang, Skelley & Perez-Farrera 2018" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFAC9501FC31FBCBD6B15F56" box="[1007,1245,1088,1112]" italics="true" pageId="25" pageNumber="176">Ceratophila vazquezi</emphasis>
|
||||
<bibRefCitation id="15634B30FFAC9501FB3BFBCBD3195F7D" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." pageId="25" pageNumber="176" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">Tang, Skelley & Pérez-Farrera 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. Deposited in the FSCA.
|
||||
<typeStatus id="AE498863FFAC9501FD40FBEFD16F5F7C" box="[670,771,1124,1149]" pageId="25" pageNumber="176" type="allotype">
|
||||
<emphasis id="4386EAD3FFAC9501FD40FBEFD16F5F7C" bold="true" box="[670,771,1124,1149]" pageId="25" pageNumber="176">Allotype</emphasis>
|
||||
</typeStatus>
|
||||
(FSCA) same data as
|
||||
<typeStatus id="AE498863FFAC9501FBD5FBEFD6015F7C" box="[1035,1133,1124,1149]" pageId="25" pageNumber="176" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="67F4FD48FFAC9501FB77FBEFD7795F7C" box="[1193,1301,1124,1149]" pageId="25" pageNumber="176" type="adult">535 adult</specimenCount>
|
||||
<typeStatus id="AE498863FFAC9501FAC0FBEFD7F85F7C" box="[1310,1428,1124,1149]" pageId="25" pageNumber="176" type="paratype">
|
||||
<emphasis id="4386EAD3FFAC9501FAC0FBEFD7F85F7C" bold="true" box="[1310,1428,1124,1149]" pageId="25" pageNumber="176">paratypes</emphasis>
|
||||
</typeStatus>
|
||||
:
|
||||
<collectingCountry id="09E57651FFAC9501FF49FB0CD3615FA1" box="[151,269,1159,1184]" name="Mexico" pageId="25" pageNumber="176">
|
||||
<emphasis id="4386EAD3FFAC9501FF49FB0CD3615FA1" bold="true" box="[151,269,1159,1184]" pageId="25" pageNumber="176">MEXICO</emphasis>
|
||||
</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="B336F823FFAC9501FEC5FB0CD3E45FA1" box="[283,392,1159,1184]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">
|
||||
<emphasis id="4386EAD3FFAC9501FEC5FB0CD3E45FA1" bold="true" box="[283,392,1159,1184]" pageId="25" pageNumber="176">Veracruz</emphasis>
|
||||
</collectingRegion>
|
||||
, Jilotepec, El Esquilón, ex male cone of
|
||||
<taxonomicName id="B6F24D42FFAC9501FC91FB03D63D5F9E" box="[847,1105,1160,1183]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="mexicana">
|
||||
<emphasis id="4386EAD3FFAC9501FC91FB03D63D5F9E" box="[847,1105,1160,1183]" italics="true" pageId="25" pageNumber="176">Ceratozamia mexicana</emphasis>
|
||||
</taxonomicName>
|
||||
[now
|
||||
<taxonomicName id="B6F24D42FFAC9501FB46FB03D6905F9E" box="[1176,1276,1160,1183]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFAC9501FB46FB03D6905F9E" box="[1176,1276,1160,1183]" italics="true" pageId="25" pageNumber="176">C. tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
],
|
||||
<date id="054C1001FFAC9501FACCFB03D7FB5FA1" box="[1298,1431,1160,1184]" pageId="25" pageNumber="176" value="1989-03-10">10-III-1989</date>
|
||||
, A. P. Vovides
|
||||
<quantity id="B60A9B24FFAC9501FEE8FB27D3EA5FC5" box="[310,390,1196,1220]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="176" unit="m" value="1300.0">1300m</quantity>
|
||||
, (97); Cerro Coacoatzintla,
|
||||
<specimenCount id="67F4FD48FFAC9501FD1FFB26D0BA5FC5" box="[705,726,1197,1220]" pageId="25" pageNumber="176" type="male">♂</specimenCount>
|
||||
cone
|
||||
<taxonomicName id="B6F24D42FFAC9501FCC4FB27D1995FC5" box="[794,1013,1196,1220]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFAC9501FCC4FB27D1995FC5" box="[794,1013,1196,1220]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFAC9501FBDAFB27D6EC5FC4" box="[1028,1152,1196,1221]" pageId="25" pageNumber="176" value="1995-02-28">28-II-1995</date>
|
||||
, Y. Sanchez Tinoco (77); W. of Mafafas, [GPS coord. omitted],
|
||||
<quantity id="B60A9B24FFAC9501FD99FB5BD0CE5FE9" box="[583,674,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.321" pageId="25" pageNumber="176" unit="m" value="1321.0">1321 m</quantity>
|
||||
., ex
|
||||
<specimenCount id="67F4FD48FFAC9501FD0BFB5AD0865FE9" box="[725,746,1233,1256]" pageId="25" pageNumber="176" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFAC9501FD2FFB5BD1A15FE6" box="[753,973,1232,1255]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFAC9501FD2FFB5BD1A15FE6" box="[753,973,1232,1255]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
late shed cone,
|
||||
<date id="054C1001FFAC9501FB5DFB44D7635FE9" box="[1155,1295,1231,1256]" pageId="25" pageNumber="176" value="2018-04-15">15- IV-2018</date>
|
||||
, M.A. Perez Farrera & W. Tang (359):
|
||||
<typeStatus id="AE498863FFAC9501FE1DFB7ED05C5E0C" box="[451,560,1269,1293]" pageId="25" pageNumber="176" type="paratype">Paratypes</typeStatus>
|
||||
deposited at ANIC, BMNH, FSCA, IEXA, INBio, MNHN, NZAC, UNAM, USNM.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFAC9501FF19FAB7D0BE5E79" pageId="25" pageNumber="176" type="etymology">
|
||||
<paragraph id="714D36C1FFAC9501FF19FAB7D0BE5E79" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||||
<emphasis id="4386EAD3FFAC9501FF19FAB7D3225E54" bold="true" box="[199,334,1340,1365]" pageId="25" pageNumber="176">Etymology.</emphasis>
|
||||
The species is named in honor of Dr. Mario Vázquez Torres for his work on the taxonomy, ecology and conservation of the cycads of
|
||||
<collectingCountry id="09E57651FFAC9501FDABFAD4D0A35E79" box="[629,719,1375,1400]" name="Mexico" pageId="25" pageNumber="176">Mexico</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFAC9501FF19FA0FD3BF5C05" pageId="25" pageNumber="176" type="discussion">
|
||||
<paragraph id="714D36C1FFAC9501FF19FA0FD3BF5C05" blockId="25.[151,1437,151,1796]" pageId="25" pageNumber="176">
|
||||
<emphasis id="4386EAD3FFAC9501FF19FA0FD3575E9C" bold="true" box="[199,315,1412,1437]" pageId="25" pageNumber="176">Remarks.</emphasis>
|
||||
<typeStatus id="AE498863FFAC9501FE9BFA0ED3105E9C" box="[325,380,1413,1437]" pageId="25" pageNumber="176">Type</typeStatus>
|
||||
specimens assigned to this species are limited to those collected on
|
||||
<taxonomicName id="B6F24D42FFAC9501FB7EFA0FD7125E9D" box="[1184,1406,1412,1436]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFAC9501FB7EFA0FD7125E9D" box="[1184,1406,1412,1436]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
, a species recently described by
|
||||
<bibRefCitation id="15634B30FFAC9501FE32FA2CD0B25EC1" author="Vovides, A. P. & Stevenson, D. W. & Perez-Farrera, M. A. & Lopez, S. & Avendano, S." box="[492,734,1447,1472]" pageId="25" pageNumber="176" pagination="419 - 429" refId="ref20651" refString="Vovides, A. P., Stevenson, D. W., Perez-Farrera, M. A., Lopez, S. & Avendano, S. (2016) What is Ceratozamia mexicana (Zamiaceae)? Botanical Sciences, 94, 419 - 429. https: // doi. org / 10.17129 / botsci. 449" type="journal article" year="2016">
|
||||
Vovides
|
||||
<emphasis id="4386EAD3FFAC9501FD90FA22D0E45EC1" box="[590,648,1448,1472]" italics="true" pageId="25" pageNumber="176">et al.</emphasis>
|
||||
(2016)
|
||||
</bibRefCitation>
|
||||
. Prior to 2016 this host had been erroneously identified as
|
||||
<taxonomicName id="B6F24D42FFAC9501FA5CFA23D30B5EE4" authority="Brongn." authorityName="Brongn." class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="mexicana">
|
||||
<emphasis id="4386EAD3FFAC9501FA5CFA23D36E5EE5" italics="true" pageId="25" pageNumber="176">C. mexicana</emphasis>
|
||||
Brongn.
|
||||
</taxonomicName>
|
||||
, a species now understood to occur only in the vicinity of El Mirador, further south in
|
||||
<collectingRegion id="B336F823FFAC9501FAEFFA46D7F55EE5" box="[1329,1433,1485,1508]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||||
. Besides
|
||||
<taxonomicName id="B6F24D42FFAC9501FF2BFA7BD3BC5D06" box="[245,464,1520,1543]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFAC9501FF2BFA7BD3BC5D06" box="[245,464,1520,1543]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
, several other species of
|
||||
<taxonomicName id="B6F24D42FFAC9501FD32FA7BD1115D06" box="[748,893,1520,1543]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFAC9501FD32FA7BD1115D06" box="[748,893,1520,1543]" italics="true" pageId="25" pageNumber="176">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
occur in the rugged terrain of western
|
||||
<collectingRegion id="B336F823FFAC9501FAEFFA7AD7F45D09" box="[1329,1432,1521,1544]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="25" pageNumber="176">Veracruz</collectingRegion>
|
||||
, including
|
||||
<taxonomicName id="B6F24D42FFAC9501FED6F99FD3F65D2D" box="[264,410,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="brevifrons">
|
||||
<emphasis id="4386EAD3FFAC9501FED6F99FD3F65D2D" box="[264,410,1556,1580]" italics="true" pageId="25" pageNumber="176">C. brevifrons</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFAC9501FE77F99FD05D5D2D" box="[425,561,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="delucana">
|
||||
<emphasis id="4386EAD3FFAC9501FE77F99FD05D5D2D" box="[425,561,1556,1580]" italics="true" pageId="25" pageNumber="176">C. delucana</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFAC9501FDE1F99FD09D5D2D" box="[575,753,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="haustecorum">
|
||||
<emphasis id="4386EAD3FFAC9501FDE1F99FD09D5D2D" box="[575,753,1556,1580]" italics="true" pageId="25" pageNumber="176">C. haustecorum</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFAC9501FD21F99FD1E15D2D" box="[767,909,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="mexicana">
|
||||
<emphasis id="4386EAD3FFAC9501FD21F99FD1E15D2D" box="[767,909,1556,1580]" italics="true" pageId="25" pageNumber="176">C. mexicana</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="B6F24D42FFAC9501FC1DF99FD6505D2D" box="[963,1084,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="morettii">
|
||||
<emphasis id="4386EAD3FFAC9501FC1DF99FD6505D2D" box="[963,1084,1556,1580]" italics="true" pageId="25" pageNumber="176">C. morettii</emphasis>
|
||||
</taxonomicName>
|
||||
. The possible occurrence of
|
||||
<taxonomicName id="B6F24D42FFAC9501FA5CF99FD3105D4E" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="vazquezi" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFAC9501FA5CF99FD7F95D2A" box="[1410,1429,1556,1579]" italics="true" pageId="25" pageNumber="176">C</emphasis>
|
||||
. (
|
||||
<emphasis id="4386EAD3FFAC9501FF7EF9B3D36D5D51" box="[160,257,1592,1616]" italics="true" pageId="25" pageNumber="176">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFAC9501FEC2F9B3D3105D4E" box="[284,380,1592,1615]" italics="true" pageId="25" pageNumber="176">vazquezi</emphasis>
|
||||
</taxonomicName>
|
||||
in these species is currently being investigated by the authors. In one male cone of
|
||||
<taxonomicName id="B6F24D42FFAC9501FF49F9D7D31B5D75" box="[151,375,1628,1652]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFAC9501FF49F9D7D31B5D75" box="[151,375,1628,1652]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
sampled at the end of its pollen shed near the town of Mafafas,
|
||||
<taxonomicName id="B6F24D42FFAC9501FB4DF9D7D7F05D75" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[1171,1436,1628,1652]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="vazquezi" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFAC9501FB4DF9D7D6CA5D72" box="[1171,1190,1628,1651]" italics="true" pageId="25" pageNumber="176">C</emphasis>
|
||||
. (
|
||||
<emphasis id="4386EAD3FFAC9501FB1CF9D7D74F5D75" box="[1218,1315,1628,1652]" italics="true" pageId="25" pageNumber="176">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFAC9501FAE2F9D6D7F05D75" box="[1340,1436,1629,1652]" italics="true" pageId="25" pageNumber="176">vazquezi</emphasis>
|
||||
</taxonomicName>
|
||||
accounted for 86% of
|
||||
<specimenCount id="67F4FD48FFAC9501FE72F90BD0705D99" box="[428,540,1663,1688]" pageId="25" pageNumber="176" type="adult">417 adult</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFAC9501FDF4F9F4D08A5D99" authorityName="Crowson" authorityYear="1952" box="[554,742,1663,1688]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="176" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||||
collected, while an undescribed species of
|
||||
<taxonomicName id="B6F24D42FFAC9501FB22F90BD7F05D99" authorityName="Reitter" authorityYear="1875" box="[1276,1436,1664,1688]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFAC9501FB22F90BD7F05D99" box="[1276,1436,1664,1688]" italics="true" pageId="25" pageNumber="176">Pharaxonotha</emphasis>
|
||||
</taxonomicName>
|
||||
accounted for the remaining 14% of adult beetles in the cone. Adult
|
||||
<taxonomicName id="B6F24D42FFAC9501FC7AF92FD6475DBD" box="[932,1067,1700,1724]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFAC9501FC7AF92FD6475DBD" box="[932,1067,1700,1724]" italics="true" pageId="25" pageNumber="176">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicName id="B6F24D42FFAC9501FBEAF92FD6B05DBD" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[1076,1244,1700,1724]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="sanchezae" subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFAC9501FBE3F92FD63C5DBA" box="[1085,1104,1700,1723]" italics="true" pageId="25" pageNumber="176">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFAC9501FBB7F92FD6B05DBD" box="[1129,1244,1700,1724]" italics="true" pageId="25" pageNumber="176">sanchezae</emphasis>
|
||||
</taxonomicName>
|
||||
, which are often found with these other two
|
||||
<taxonomicName id="B6F24D42FFAC9501FE11F94CD0E75DE1" authorityName="Crowson" authorityYear="1952" box="[463,651,1735,1760]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="176" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||||
in cones of
|
||||
<taxonomicName id="B6F24D42FFAC9501FCC7F943D1995DDE" box="[793,1013,1736,1759]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFAC9501FCC7F943D1995DDE" box="[793,1013,1736,1759]" italics="true" pageId="25" pageNumber="176">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
, were absent, but see Remarks under
|
||||
<taxonomicName id="B6F24D42FFAC9501FF49F967D3735C05" box="[151,287,1772,1796]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFAC9501FF49F967D3735C05" box="[151,287,1772,1796]" italics="true" pageId="25" pageNumber="176">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicName id="B6F24D42FFAC9501FEFBF967D3A35C05" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[293,463,1772,1796]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="25" pageNumber="176" phylum="Tracheophyta" rank="species" species="sanchezae" subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFAC9501FEF0F967D32D5C02" box="[302,321,1772,1795]" italics="true" pageId="25" pageNumber="176">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFAC9501FE86F967D3A35C05" box="[344,463,1772,1796]" italics="true" pageId="25" pageNumber="176">sanchezae</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="4386EAD3FFAC9501FE11F967D3BF5C05" box="[463,467,1772,1796]" italics="true" pageId="25" pageNumber="176">.</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
126
data/F9/5B/87/F95B87D7FFA49509FF49FE3CD7505E2B.xml
Normal file
126
data/F9/5B/87/F95B87D7FFA49509FF49FE3CD7505E2B.xml
Normal file
|
|
@ -0,0 +1,126 @@
|
|||
<document id="F7F03383AC8B8F911D0EC17BBA6869C6" ID-CLB-Dataset="28054" ID-DOI="10.11646/zootaxa.4508.2.1" ID-GBIF-Dataset="da79b654-bd50-4715-8ed3-80fa7a00d469" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606934" ID-ZooBank="5E2BC894-1919-4F63-8EF5-BAAC91913388" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585631772" checkinUser="plazi" docAuthor="Tang, William, Skelley, Paul & Pérez-Farrera, Miguel Angel" docDate="2018" docId="F95B87D7FFA49509FF49FE3CD7505E2B" docLanguage="en" docName="zootaxa.4508.2.1.pdf" docOrigin="Zootaxa 4508 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratophila (Vovidesa) Tang & Skelley & Pérez-Farrera 2018, new subgenus" docType="treatment" docVersion="7" lastPageNumber="168" masterDocId="0562FFAFFFB59518FFDEFF8BD26C5B01" masterDocTitle="Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)" masterLastPageNumber="178" masterPageNumber="151" pageNumber="168" updateTime="1698712984586" updateUser="ExternalLinkService">
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<mods:mods id="D2B47433135E45510EA5B48200F2C672" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1B0FD7A4E545798F744D86199B37844D">
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<mods:title id="E0547E42FC10E8560E161BDEF17419F2">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
|
||||
</mods:titleInfo>
|
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<mods:name id="4A70D0AE54816B72CC488ABA4089F3E6" type="personal">
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<mods:role id="1512D4D5165AFEB22D9523DAED851BC9">
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<mods:namePart id="50CB58A1919430C24CD6EF44048046C2">Tang, William</mods:namePart>
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</mods:role>
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||||
<mods:namePart id="DBAEED564896FD55F69F6F198ED8AFF7">Skelley, Paul</mods:namePart>
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<mods:name id="1FC2D8990E083AAA4C276360A7A0F44D" type="personal">
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</mods:role>
|
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<mods:namePart id="BE6E72415F0C0358BAAFF4784303FA72">Pérez-Farrera, Miguel Angel</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A2F540AE710478B04EF65DFAF7324264">text</mods:typeOfResource>
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<mods:title id="77B7DDB17FCEC24738BE3DC0ED248FA8">Zootaxa</mods:title>
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</mods:titleInfo>
|
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<mods:part id="B44A153B3C0A18D4E11F9F41896DDC85">
|
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<mods:date id="86E9D1CA96E396BD60D235901CD8F441">2018</mods:date>
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<mods:number id="6154AB06078D7E440F919B5BCD8F8D32">2018-10-31</mods:number>
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|
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<mods:number id="11D0DE577E6F0B391750E771616A124A">4508</mods:number>
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|
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<mods:classification id="3ADFDD0678ED1AC66FF9FAD86A3B30C4">journal article</mods:classification>
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<mods:identifier id="AD7A06D027717E3FE9349DD09499558F" type="GBIF-Dataset">da79b654-bd50-4715-8ed3-80fa7a00d469</mods:identifier>
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<mods:identifier id="E549597D4801F6C56B1B8885C196B0D7" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="F95B87D7FFA49509FF49FE3CD7505E2B" ID-DOI="http://doi.org/10.5281/zenodo.6491684" ID-GBIF-Taxon="154495721" ID-Zenodo-Dep="6491684" LSID="urn:lsid:plazi:treatment:F95B87D7FFA49509FF49FE3CD7505E2B" httpUri="http://treatment.plazi.org/id/F95B87D7FFA49509FF49FE3CD7505E2B" lastPageNumber="168" pageId="17" pageNumber="168">
|
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<subSubSection id="39E8654AFFA49509FF49FE3CD1825AD3" box="[151,1006,439,466]" pageId="17" pageNumber="168" type="nomenclature">
|
||||
<paragraph id="714D36C1FFA49509FF49FE3CD1825AD3" blockId="17.[151,1006,439,466]" box="[151,1006,439,466]" pageId="17" pageNumber="168">
|
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<heading id="2A0581ADFFA49509FF49FE3CD1825AD3" bold="true" box="[151,1006,439,466]" fontSize="11" level="1" pageId="17" pageNumber="168" reason="1">
|
||||
<emphasis id="4386EAD3FFA49509FF49FE3CD1825AD3" bold="true" box="[151,1006,439,466]" pageId="17" pageNumber="168">
|
||||
<taxonomicName id="B6F24D42FFA49509FF49FE3CD1585AD3" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,820,439,466]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="subGenus" status="subgen. nov." subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFA49509FF49FE3CD34B5AD0" bold="true" box="[151,295,439,465]" italics="true" pageId="17" pageNumber="168">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA49509FEE9FE3CD3F25AD0" bold="true" box="[311,414,439,465]" italics="true" pageId="17" pageNumber="168">Vovidesa</emphasis>
|
||||
) Tang, Skelley & Pérez-Farrera
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFA49509FC9EFE33D1825AD3" box="[832,1006,440,466]" pageId="17" pageNumber="168" rank="subGenus">new subgenus</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA49509FF49FE75D1145915" box="[151,888,510,533]" pageId="17" pageNumber="168" type="materials_examined">
|
||||
<paragraph id="714D36C1FFA49509FF49FE75D1145915" blockId="17.[151,888,510,533]" box="[151,888,510,533]" pageId="17" pageNumber="168">
|
||||
<emphasis id="4386EAD3FFA49509FF49FE75D34E5915" bold="true" box="[151,290,510,532]" pageId="17" pageNumber="168">
|
||||
<typeStatus id="AE498863FFA49509FF49FE75D2A05915" box="[151,204,510,532]" pageId="17" pageNumber="168">Type</typeStatus>
|
||||
species.
|
||||
</emphasis>
|
||||
<taxonomicName id="B6F24D42FFA49509FEF7FE75D0EC5914" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[297,640,510,533]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Uovidesa">
|
||||
<emphasis id="4386EAD3FFA49509FEF7FE75D3C85915" box="[297,420,510,532]" italics="true" pageId="17" pageNumber="168">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA49509FE6CFE75D0665915" box="[434,522,510,532]" italics="true" pageId="17" pageNumber="168">Uovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFA49509FDC5FD8BD0EC5914" box="[539,640,512,533]" italics="true" pageId="17" pageNumber="168">mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFA49509FD53FE75D0AA5915" box="[653,710,510,532]" pageId="17" pageNumber="168" rank="species">
|
||||
<emphasis id="4386EAD3FFA49509FD53FE75D0AA5915" bold="true" box="[653,710,510,532]" pageId="17" pageNumber="168">n. sp.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, here designated.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA49509FF49FDCAD64B59A3" pageId="17" pageNumber="168" type="diagnosis">
|
||||
<paragraph id="714D36C1FFA49509FF49FDCAD64B59A3" blockId="17.[151,1437,577,1322]" pageId="17" pageNumber="168">
|
||||
<emphasis id="4386EAD3FFA49509FF49FDCAD330595B" bold="true" box="[151,348,577,602]" pageId="17" pageNumber="168">Adult diagnosis.</emphasis>
|
||||
The subgenus
|
||||
<emphasis id="4386EAD3FFA49509FDCFFDCAD0485959" box="[529,548,577,600]" italics="true" pageId="17" pageNumber="168">C</emphasis>
|
||||
. (
|
||||
<taxonomicName id="B6F24D42FFA49509FDE1FDCAD0CC5958" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[575,672,577,601]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="172" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFA49509FDE1FDCAD0CC5958" box="[575,672,577,601]" italics="true" pageId="17" pageNumber="168">Vovidesa</emphasis>
|
||||
</taxonomicName>
|
||||
) is distinguished from members of the nominal subgenus by the explanate pronotal surface near the lateral carina; lateral carina weakly thicker anteriorly, meso- and metatibia not triangularly dilated toward apex; and strong male dimorphism present on all legs.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA49509FF19FD26D6D45F2F" pageId="17" pageNumber="168" type="description">
|
||||
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|
||||
<emphasis id="4386EAD3FFA49509FF19FD26D3CE59C7" bold="true" box="[199,418,685,710]" pageId="17" pageNumber="168">Adult description.</emphasis>
|
||||
Length
|
||||
<quantity id="B60A9B24FFA49509FDDAFD26D0C759C4" box="[516,683,685,710]" metricMagnitude="-3" metricUnit="m" metricValue="4.22" metricValueMax="4.9" metricValueMin="3.54" pageId="17" pageNumber="168" unit="mm" value="4.22" valueMax="4.9" valueMin="3.54">3.54–4.90 mm</quantity>
|
||||
, width
|
||||
<quantity id="B60A9B24FFA49509FCDDFD26D1CE59C7" box="[771,930,685,710]" metricMagnitude="-3" metricUnit="m" metricValue="1.705" metricValueMax="1.91" metricValueMin="1.5" pageId="17" pageNumber="168" unit="mm" value="1.705" valueMax="1.91" valueMin="1.5">1.50–1.91mm</quantity>
|
||||
. Mouthparts: submentum with long erect or reclining setae. Pronotum transverse, length/width ratio = 0.70–0.83; anterior margin near eyes moderately emarginate, anterior angles moderately projecting; surface near lateral carinae explanate, carinae in dorsal view not evenly arcuate with medial region parallel sided or slightly emarginate; carinae in lateral view weakly thickening anteriorly, anterior thickness <1.5× that at base; pronotal base with longitudinal groove extending anteriorly from lateral basal pore in margin. Elytra: in all populations color uniform without darkened punctation; most specimens with distinct fine setae on surface. Legs with protibial bearing distal lateral tooth only moderately developed, apical spinules of tooth reduced; metatibiae not triangularly dilated at apex which has fringe of long fine spinules. Male genitalia: median lobe and tegmen may or may not be compressed laterally, penile struts 1.93–2.41× longer than median lobe. Sexual dimorphism present; males with thickened femora and modified tibia of all legs, female legs narrow and unmodified. Males with broad, shallowly concave, medial portion of metaventrite.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA49509FF19FBB2D1875F77" pageId="17" pageNumber="168" type="etymology">
|
||||
<paragraph id="714D36C1FFA49509FF19FBB2D1875F77" blockId="17.[151,1437,577,1322]" pageId="17" pageNumber="168">
|
||||
<emphasis id="4386EAD3FFA49509FF19FBB2D3225F53" bold="true" box="[199,334,1081,1106]" pageId="17" pageNumber="168">Etymology.</emphasis>
|
||||
The subgenus is named in honor of Dr. Andrew P. Vovides for his work on the taxonomy, ecology, physiology and conservation of cycads in the New World. Gender feminine.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA49509FF19FB0AD7505E2B" pageId="17" pageNumber="168" type="discussion">
|
||||
<paragraph id="714D36C1FFA49509FF19FB0AD7505E2B" blockId="17.[151,1437,577,1322]" pageId="17" pageNumber="168">
|
||||
<emphasis id="4386EAD3FFA49509FF19FB0AD3585F9B" bold="true" box="[199,308,1153,1178]" pageId="17" pageNumber="168">Remarks</emphasis>
|
||||
. This subgenus shows more striking sexual dimorphism than any other pharaxonothine known. Males have highly modified legs and other structures that differ from population to population. Females, lacking such modifications, are difficult to identify when not associated with males. Thus, males of this group are needed for species recognition and description. Females of this subgenus are known from localities not cited here, requiring more field work and male specimens before the identity of those populations can be determined.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
402
data/F9/5B/87/F95B87D7FFA4950CFF49FAF6D120587E.xml
Normal file
402
data/F9/5B/87/F95B87D7FFA4950CFF49FAF6D120587E.xml
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|
|
@ -0,0 +1,402 @@
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<mods:title id="E7CFA56EAE018C26010417D93BCF6BD3">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<mods:namePart id="DCA0249F5089D27335F7CEB8E2004B1A">Pérez-Farrera, Miguel Angel</mods:namePart>
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<paragraph id="714D36C1FFA49509FF49FAF6D63A5E96" blockId="17.[151,1110,1405,1464]" box="[151,1110,1405,1431]" pageId="17" pageNumber="168">
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<heading id="2A0581ADFFA49509FF49FAF6D63A5E96" bold="true" box="[151,1110,1405,1431]" fontSize="11" level="1" pageId="17" pageNumber="168" reason="1">
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<emphasis id="4386EAD3FFA49509FF49FAF6D63A5E96" bold="true" box="[151,1110,1405,1431]" pageId="17" pageNumber="168">
|
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<taxonomicName id="B6F24D42FFA49509FF49FAF6D1D65E96" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,954,1405,1431]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Vovidesa">
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||||
<emphasis id="4386EAD3FFA49509FF49FAF6D34B5E96" bold="true" box="[151,295,1405,1431]" italics="true" pageId="17" pageNumber="168">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA49509FEE9FAF6D3F25E96" bold="true" box="[311,414,1405,1431]" italics="true" pageId="17" pageNumber="168">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFA49509FE6CFAF6D05D5E96" bold="true" box="[434,561,1405,1431]" italics="true" pageId="17" pageNumber="168">chiapensis</emphasis>
|
||||
Tang, Skelley & Pérez-Farrera
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFA49509FC19FAF6D63A5E96" box="[967,1110,1405,1431]" pageId="17" pageNumber="168" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA49509FF49FA14D3235EB9" box="[151,335,1439,1464]" pageId="17" pageNumber="168" type="description">
|
||||
<paragraph id="714D36C1FFA49509FF49FA14D3235EB9" blockId="17.[151,1110,1405,1464]" box="[151,335,1439,1464]" pageId="17" pageNumber="168">
|
||||
(
|
||||
<figureCitation id="E9C92A44FFA49509FF41FA14D2935EB9" box="[159,255,1439,1464]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="17" pageNumber="168">Figs. 1E</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFA49509FED2FA14D3755EB9" box="[268,281,1439,1464]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="17" pageNumber="168">6</figureCitation>
|
||||
A–I)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA49509FF49FA6CD0325D8E" pageId="17" pageNumber="168" type="diagnosis">
|
||||
<paragraph id="714D36C1FFA49509FF49FA6CD0325D8E" blockId="17.[151,1437,1511,2005]" pageId="17" pageNumber="168">
|
||||
<emphasis id="4386EAD3FFA49509FF49FA6CD30D5D01" bold="true" box="[151,353,1511,1536]" pageId="17" pageNumber="168">Adult diagnosis.</emphasis>
|
||||
Distinguished from other
|
||||
<taxonomicName id="B6F24D42FFA49509FD74FA63D1DA5D01" authority="(Vovidesa)" baseAuthorityName="Vovidesa" box="[682,950,1512,1536]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="155" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA49509FD74FA63D15D5D01" box="[682,817,1512,1536]" italics="true" pageId="17" pageNumber="168">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA49509FC94FA63D1C75D01" box="[842,939,1512,1536]" italics="true" pageId="17" pageNumber="168">Vovidesa</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
spp. by male lacking subapical medial emarginations on tibiae; submentum of both sexes with sparse punctures and setae; female elytra dulled medially and laterally; female abdomen strongly dulled, obscuring punctures; male genitalia with median lobe laterally flattened and curved; and with a known distribution in
|
||||
<collectingCountry id="09E57651FFA49509FCC3F9DFD1195D6C" box="[797,885,1620,1645]" name="Mexico" pageId="17" pageNumber="168">Mexico</collectingCountry>
|
||||
, state of
|
||||
<collectingRegion id="B336F823FFA49509FC3DF9DFD62C5D6C" box="[995,1088,1620,1645]" country="Mexico" name="Chiapas" pageId="17" pageNumber="168">Chiapas</collectingRegion>
|
||||
, on
|
||||
<taxonomicName id="B6F24D42FFA49509FBABF9DFD71C5D6D" authorityName="Perez-Farrera, Vovides & Iglesias" authorityYear="1999" box="[1141,1392,1620,1644]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="alvarezii">
|
||||
<emphasis id="4386EAD3FFA49509FBABF9DFD71C5D6D" box="[1141,1392,1620,1644]" italics="true" pageId="17" pageNumber="168">Ceratozamia alvarezii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFA49509FA5CF9DFD3695D91" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFA49509FA5CF9DFD7F95D6A" box="[1410,1429,1620,1643]" italics="true" pageId="17" pageNumber="168">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFA49509FF49F9F3D3695D91" box="[151,261,1656,1680]" italics="true" pageId="17" pageNumber="168">mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="4386EAD3FFA49509FEDBF9F3D3FA5D8E" box="[261,406,1656,1680]" italics="true" pageId="17" pageNumber="168">
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFA49509FECEF9F3D3FA5D8E" authorityName="D. W. Stevenson" authorityYear="1982" box="[272,406,1656,1679]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="norstogii">C. norstogii</taxonomicName>
|
||||
</emphasis>
|
||||
, and
|
||||
<taxonomicName id="B6F24D42FFA49509FE0DF9F3D0345D91" box="[467,600,1656,1680]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFA49509FE0DF9F3D38A5D8E" box="[467,486,1656,1679]" italics="true" pageId="17" pageNumber="168">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFA49509FE2AF9F3D0345D91" box="[500,600,1656,1680]" italics="true" pageId="17" pageNumber="168">vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA4950AFF19F917D28F5CDD" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168" type="description">
|
||||
<paragraph id="714D36C1FFA49509FF19F917D6855C45" blockId="17.[151,1437,1511,2005]" pageId="17" pageNumber="168">
|
||||
<emphasis id="4386EAD3FFA49509FF19F917D38C5DB4" bold="true" box="[199,480,1692,1717]" pageId="17" pageNumber="168">Adult male description.</emphasis>
|
||||
Length
|
||||
<quantity id="B60A9B24FFA49509FDE0F917D08F5DB5" box="[574,739,1692,1717]" metricMagnitude="-3" metricUnit="m" metricValue="4.015" metricValueMax="4.49" metricValueMin="3.54" pageId="17" pageNumber="168" unit="mm" value="4.015" valueMax="4.49" valueMin="3.54">3.54–4.49 mm</quantity>
|
||||
, width
|
||||
<quantity id="B60A9B24FFA49509FCE8F917D1B65DB5" box="[822,986,1692,1717]" metricMagnitude="-3" metricUnit="m" metricValue="1.645" metricValueMax="1.79" metricValueMin="1.5" pageId="17" pageNumber="168" unit="mm" value="1.645" valueMax="1.79" valueMin="1.5">1.50–1.79 mm</quantity>
|
||||
(n = 20). Body in dorsal view elongateoval, greatest width at middle of elytra; in lateral view convex dorsally. General body color brown (
|
||||
<figureCitation id="E9C92A44FFA49509FAD4F934D7345DD9" box="[1290,1368,1727,1752]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="17" pageNumber="168">Figs. 6</figureCitation>
|
||||
A–E), head and pronotum concolorous and on some slightly darker than the yellowish-brown elytra; dorsal surface punctate, shining but weakly alutaceous, short procumbent hairs associated with punctation, ventrally shining and appearing glabrous except mesoventrite and abdomen mostly covered with long procumbent setae.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA4950AFF19F8C4D3DB5AAC" blockId="17.[151,1437,1511,2005]" lastBlockId="18.[151,1437,151,2012]" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">
|
||||
<emphasis id="4386EAD3FFA49509FF19F8C4D36B5C69" bold="true" box="[199,263,1871,1896]" pageId="17" pageNumber="168">Head</emphasis>
|
||||
in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured (
|
||||
<figureCitation id="E9C92A44FFA49509FEC9F8FFD3CC5C8D" box="[279,416,1908,1933]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="17" pageNumber="168">Figs. 6A, D</figureCitation>
|
||||
), average distance between closest punctures 2× width of puncture; width
|
||||
<quantity id="B60A9B24FFA49509FB2FF8FFD7F45C8D" box="[1265,1432,1908,1933]" metricMagnitude="-4" metricUnit="m" metricValue="8.3" metricValueMax="9.1" metricValueMin="7.5" pageId="17" pageNumber="168" unit="mm" value="0.83" valueMax="0.91" valueMin="0.75">0.75–0.91 mm</quantity>
|
||||
; dorsal interocular distance
|
||||
<quantity id="B60A9B24FFA49509FE3DF813D0FE5CB1" box="[483,658,1944,1969]" metricMagnitude="-4" metricUnit="m" metricValue="5.449999999999999" metricValueMax="6.0" metricValueMin="4.9" pageId="17" pageNumber="168" unit="mm" value="0.5449999999999999" valueMax="0.6" valueMin="0.49">0.49–0.60 mm</quantity>
|
||||
, head width/dorsal interocular distance ratio 1.45–1.56, ventral interocular distance
|
||||
<quantity id="B60A9B24FFA49509FEA3F837D0485CD5" box="[381,548,1980,2005]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.5" metricValueMin="3.5999999999999996" pageId="17" pageNumber="168" unit="mm" value="0.405" valueMax="0.45" valueMin="0.36">0.36–0.45 mm</quantity>
|
||||
, head width/ventral interocular distance ratio 1.96–2.22. Eye with large black facets. Antennal length approximately equal to pronotal width, 1.5× head width; antennomere I (scape) fairly large, slightly elongate, antennomere II subequal to III; IV–VIII small, width equals length; club fairly large, IX and X similar in length, XI slightly longer, globular with acuminate apex. Clypeus truncate anteriorly, moderately punctate, with narrow margin. Mentum and submentum with moderate punctation, distance between nearest punctures ~ 2× own width, each puncture with a long seta ~ 2–3× width of eye facet and reclining anteriad (often abraded). Gular area smooth, without punctation or setae, border with submentum usually with a shallow depression without punctation, but usually with a shallowly impressed circular or semicircular border along anterior margin (
|
||||
<figureCitation id="E9C92A44FFA7950AFE8DFE1FD3C65AAC" box="[339,426,404,429]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">Fig. 6E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA7950AFF19FE3CD6B15FE9" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="169">
|
||||
<emphasis id="4386EAD3FFA7950AFF19FE3CD34C5AD1" bold="true" box="[199,288,439,464]" pageId="18" pageNumber="169">Thorax</emphasis>
|
||||
with pronotum weakly transverse in dorsal view with marginal beads basally and laterally, anterior margin with weak bead, reduced to absent medially; length/width ratio (PL/PW) = 0.76–0.83, rectangular; anterior and posterior angles present, anterior angles projecting; lateral carinae expanding in basal 1/10 to parallel sides, sometimes slightly emarginated, parallel for 50% of length, then gradually converging to anterior angles; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/5 length of posterior margin from posterior corners and touching and just anterior to marginal bead, pores mark base of longitudinal furrows onto disc. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae ~ 1/3 length of eye. Hypomeron laterally apparently lacking punctures, some with minute setae visible; medially with longitudinal striations. Elytra in dorsal view elongate-oval, convex; length/width 1.67–1.74, greatest width near midlength; without marginal line basally; 10 complete striae of moderate puncture size, scutellary striole extending ¼ elytral length, with 9–16 punctures; intervals of striae with shallow punctures similar to those of striae, obscuring striae. All punctures of elytral bearing a single fine, procumbent seta; seta often only visible in profile, extending posteriorly to, over or beyond next puncture (often abraded), interval puncture setae longer than strial puncture setae. Punctation on meso- and metaventrites moderate to dense, distance between nearest punctures 1–2× width of punctures, puncture depth shallow. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over 3/4 metaventrite length. Legs with procoxa oval; mesocoxa globular; metacoxa transversely elongateoval; trochanters obliquely truncate apically; in males all femora swollen, stout, dorso-ventral width greater than eye length; all tibiae lacking shallow medial subapical emargination, obliquely truncate at apex, with complete fringe of fine spinules on anterior and posterior margins; protibia swollen, not flattened in cross section notably larger than other tibiae, triangularly dilated to apex; mesotibia weakly swollen, moderately triangularly dilated to apex; metatibia narrowed, weakly dilating to midlength then nearly parallel to apex; pro- and mesotarsi greatly enlarged, corresponding tarsomere I width ~ 3× own length, metatarsomere I width ~ own length.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA7950AFF19FB7FD2A65C04" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="169">
|
||||
<emphasis id="4386EAD3FFA7950AFF19FB7FD32F5E0C" bold="true" box="[199,323,1268,1293]" pageId="18" pageNumber="169">Abdomen.</emphasis>
|
||||
Ventrite I with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; all ventrites strongly alutaceous across entire surface, bearing indistinct punctures, distance to nearest puncture ~ 1× width of puncture, punctures bearing mostly reclining setae; ventrites I–V with setae length ~ 2–3× width of puncture; I–V each with 2 or more median pairs of longer, semi-erect sensory hairs (obscured with other setation), V with additional 4–5 pairs of long semi-erect hairs located postero-laterally, but not at submargin. Male genitalia with genital capsule fringed with fine setae; tegmen sclerotized, triangular, laterally compressed; anterior region ring-like, posterior region sheath-like; lateral margins gradually converging posteriorly (
|
||||
<figureCitation id="E9C92A44FFA7950AFBAEF9BCD6D15D51" box="[1136,1213,1591,1616]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">Figs. 6</figureCitation>
|
||||
G–H); apically with 2 elongate, somewhat bar-shaped parameres. Parameres dorso-ventrally compressed, height approximately half its own width, in dorso-ventral view length/width ratio = 2.14–2.67; apically with setae, length of longest setae> width of paramere in dorso-ventral view (
|
||||
<figureCitation id="E9C92A44FFA7950AFDA9F92FD0A45DBC" box="[631,712,1700,1725]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">
|
||||
Fig.
|
||||
<date id="054C1001FFA7950AFD70F92FD0A45DBC" box="[686,712,1700,1725]" pageId="18" pageNumber="169">6I</date>
|
||||
</figureCitation>
|
||||
); median lobe laterally compressed, sclerotized, in lateral view strongly curved, apex tapering to a point. Ratio of penile strut length to median lobe length 1.93–2.19 (n = 2) (
|
||||
<figureCitation id="E9C92A44FFA7950AFAB1F94CD2D15C04" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">Fig. 6H</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA7950AFF19F884D28F5CDD" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="169">
|
||||
<emphasis id="4386EAD3FFA7950AFF19F884D3725C29" bold="true" box="[199,286,1807,1832]" pageId="18" pageNumber="169">Female</emphasis>
|
||||
generally similar to male, except for sexually dimorphisms. Female pronotum slightly more transverse than male, PL/PW = 0.75–0.81; elytra more strongly dulled and alutaceous laterally and over most of disc; legs unmodified, fore and middle legs not swollen, femora dorso-ventral width about equal to eye, all tibia gradually dilating to narrowly triangular apex, all legs with tarsomere I width ~ own length (
|
||||
<figureCitation id="E9C92A44FFA7950AFBB3F8F7D6D35C94" box="[1133,1215,1916,1941]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="18" pageNumber="169">Figs. 6</figureCitation>
|
||||
A–C); metaventrite flattened medially, not impressed; abdomen strongly alutaceous, obscuring punctures, less densely punctate and setose.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="258D6649FFA6950BFF49F808D2A25CF4" httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="19" pageNumber="170" startId="19.[151,250,1923,1945]" targetBox="[203,1389,193,1908]" targetPageId="19">
|
||||
<paragraph id="714D36C1FFA6950BFF49F808D2A25CF4" blockId="19.[151,1437,1923,2037]" pageId="19" pageNumber="170">
|
||||
<emphasis id="4386EAD3FFA6950BFF49F808D3785C98" bold="true" box="[151,276,1923,1945]" pageId="19" pageNumber="170">FIGURE 6.</emphasis>
|
||||
Photographs of
|
||||
<taxonomicName id="B6F24D42FFA6950BFE63F808D1705C98" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[445,796,1923,1945]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="19" pageNumber="170" phylum="Tracheophyta" rank="subGenus" subGenus="Uovidesa">
|
||||
<emphasis id="4386EAD3FFA6950BFE63F808D0545C98" box="[445,568,1923,1945]" italics="true" pageId="19" pageNumber="170">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA6950BFD96F808D0CE5C98" box="[584,674,1923,1945]" italics="true" pageId="19" pageNumber="170">Uovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFA6950BFD6FF808D1705C98" box="[689,796,1923,1945]" italics="true" pageId="19" pageNumber="170">chiapensis</emphasis>
|
||||
</taxonomicName>
|
||||
collected from
|
||||
<taxonomicName id="B6F24D42FFA6950BFC61F80FD6CF5C98" box="[959,1187,1923,1945]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="19" pageNumber="170" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFA6950BFC61F80FD6CF5C98" box="[959,1187,1923,1945]" italics="true" pageId="19" pageNumber="170">Ceratozamia vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
(A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="39E8654AFFA1950CFF19FF1CD7E959CD" pageId="20" pageNumber="171" type="materials_examined">
|
||||
<paragraph id="714D36C1FFA1950CFF19FF1CD66F5BB1" blockId="20.[151,1437,151,896]" box="[199,1027,151,176]" pageId="20" pageNumber="171">
|
||||
<materialsCitation id="C19A3C9CFFA1950CFF19FF1CD66F5BB1" ID-GBIF-Occurrence="2028138746" box="[199,1027,151,176]" country="Mexico" elevation="957" location="Jaltenango" pageId="20" pageNumber="171" specimenCount="957" stateProvince="Chiapas" typeStatus="holotype">
|
||||
<emphasis id="4386EAD3FFA1950CFF19FF1CD3095BB1" bold="true" box="[199,357,151,176]" pageId="20" pageNumber="171">
|
||||
<typeStatus id="AE498863FFA1950CFF19FF1CD36E5BB1" box="[199,258,151,176]" pageId="20" pageNumber="171">Type</typeStatus>
|
||||
locality.
|
||||
</emphasis>
|
||||
<collectingCountry id="09E57651FFA1950CFEB2FF1CD3A85BB1" box="[364,452,151,176]" name="Mexico" pageId="20" pageNumber="171">Mexico</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFA1950CFE11FF1CD0405BB1" box="[463,556,151,176]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||||
,
|
||||
<location id="742D601AFFA1950CFDE6FF1CD0DF5BB1" LSID="urn:lsid:plazi:treatment:F95B87D7FFA4950CFF49FAF6D120587E:742D601AFFA1950CFDE6FF1CD0DF5BB1" box="[568,691,151,176]" country="Mexico" name="Jaltenango" pageId="20" pageNumber="171" stateProvince="Chiapas">Jaltenango</location>
|
||||
, [GPS coord. omitted],
|
||||
<specimenCount id="67F4FD48FFA1950CFC62FF1CD66F5BB1" box="[956,1027,151,176]" pageId="20" pageNumber="171" type="generic">
|
||||
<elevation id="FADFD1F2FFA1950CFC62FF1CD1935BB1" box="[956,1023,151,176]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="20" pageNumber="171" unit="m" value="957.0">
|
||||
<quantity id="B60A9B24FFA1950CFC62FF1CD1935BB1" box="[956,1023,151,176]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="20" pageNumber="171" unit="m" value="957.0">957m</quantity>
|
||||
</elevation>
|
||||
.
|
||||
</specimenCount>
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA1950CFF19FF37D76A5BD5" blockId="20.[151,1437,151,896]" box="[199,1286,188,213]" pageId="20" pageNumber="171">
|
||||
<emphasis id="4386EAD3FFA1950CFF19FF37D3765BD4" bold="true" box="[199,282,188,213]" pageId="20" pageNumber="171">Range.</emphasis>
|
||||
Sierra Madre de
|
||||
<collectingRegion id="B336F823FFA1950CFE03FF37D0565BD4" box="[477,570,188,213]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="09E57651FFA1950CFD9BFF37D0F15BD4" box="[581,669,188,213]" name="Mexico" pageId="20" pageNumber="171">Mexico</collectingCountry>
|
||||
, inhabiting male cones of four species of
|
||||
<taxonomicName id="B6F24D42FFA1950CFBB1FF37D76C5BD2" box="[1135,1280,188,211]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA1950CFBB1FF37D76C5BD2" box="[1135,1280,188,211]" italics="true" pageId="20" pageNumber="171">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFA1950CFF19FF54D7E959CD" blockId="20.[151,1437,151,896]" pageId="20" pageNumber="171">
|
||||
<emphasis id="4386EAD3FFA1950CFF19FF54D05B5BF9" bold="true" box="[199,567,223,248]" pageId="20" pageNumber="171">
|
||||
Material examined.
|
||||
<typeStatus id="AE498863FFA1950CFE14FF54D05B5BF9" box="[458,567,223,248]" pageId="20" pageNumber="171" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
(by designation) male of
|
||||
<taxonomicName id="B6F24D42FFA1950CFC59FF6BD6445BF9" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[903,1064,224,248]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="chiapensis">
|
||||
<emphasis id="4386EAD3FFA1950CFC59FF6BD6445BF9" box="[903,1064,224,248]" italics="true" pageId="20" pageNumber="171">C. chiapensis</emphasis>
|
||||
</taxonomicName>
|
||||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||||
<collectingCountry id="09E57651FFA1950CFD57FE8FD0915A1D" box="[649,765,260,284]" name="Mexico" pageId="20" pageNumber="171">MEXICO</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFA1950CFCD3FE8FD1065A1C" box="[781,874,260,285]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||||
, Jaltenango, [GPS coord. omitted],
|
||||
<quantity id="B60A9B24FFA1950CFAD1FE8FD73D5A1C" box="[1295,1361,260,285]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="20" pageNumber="171" unit="m" value="957.0">957m</quantity>
|
||||
, ex
|
||||
<specimenCount id="67F4FD48FFA1950CFA59FE8ED7F05A1D" box="[1415,1436,261,284]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFA1950CFF49FEA3D3F85A41" box="[151,404,296,320]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFA1950CFF49FEA3D3F85A41" box="[151,404,296,320]" italics="true" pageId="20" pageNumber="171">Ceratozamia vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
cone,
|
||||
<date id="054C1001FFA1950CFE01FEA3D0375A41" box="[479,603,296,320]" pageId="20" pageNumber="171" value="2017-05-21">
|
||||
21-
|
||||
<bibRefCitation id="15634B30FFA1950CFDDCFEA3D0375A41" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[514,603,296,320]" pageId="20" pageNumber="171" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">V-2017</bibRefCitation>
|
||||
</date>
|
||||
, H. Gomez Dominguez & W.Tang”; 2) [rectangular; red; printed in black ink]
|
||||
<typeStatus id="AE498863FFA1950CFF0CFEC7D30A5A65" box="[210,358,332,356]" pageId="20" pageNumber="171" type="holotype">HOLOTYPE</typeStatus>
|
||||
<specimenCount id="67F4FD48FFA1950CFEABFEC6D3E65A65" box="[373,394,333,356]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFA1950CFE46FEC7D6045A65" authority="Tang, Skelley & Perez-Farrera 2018" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[408,1128,332,357]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA1950CFE46FEC7D0C85A65" box="[408,676,332,356]" italics="true" pageId="20" pageNumber="171">Ceratophila chiapensis</emphasis>
|
||||
<bibRefCitation id="15634B30FFA1950CFD6CFEC6D6045A65" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." box="[690,1128,332,357]" pageId="20" pageNumber="171" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">Tang, Skelley & Pérez-Farrera 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. Deposited in the FSCA.
|
||||
<typeStatus id="AE498863FFA1950CFF49FEE4D2905A89" box="[151,252,367,392]" pageId="20" pageNumber="171" type="allotype">
|
||||
<emphasis id="4386EAD3FFA1950CFF49FEE4D2905A89" bold="true" box="[151,252,367,392]" pageId="20" pageNumber="171">Allotype</emphasis>
|
||||
</typeStatus>
|
||||
(FSCA) same data as
|
||||
<typeStatus id="AE498863FFA1950CFE2AFEE4D03A5A89" box="[500,598,367,392]" pageId="20" pageNumber="171" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="67F4FD48FFA1950CFD55FEE4D0985A89" box="[651,756,367,392]" pageId="20" pageNumber="171" type="adult">175 adult</specimenCount>
|
||||
<typeStatus id="AE498863FFA1950CFD24FEE4D11C5A89" box="[762,880,367,392]" pageId="20" pageNumber="171" type="paratype">
|
||||
<emphasis id="4386EAD3FFA1950CFD24FEE4D11C5A89" bold="true" box="[762,880,367,392]" pageId="20" pageNumber="171">paratypes</emphasis>
|
||||
</typeStatus>
|
||||
:
|
||||
<emphasis id="4386EAD3FFA1950CFCA0FEE4D60A5A89" bold="true" box="[894,1126,367,392]" pageId="20" pageNumber="171">
|
||||
<collectingCountry id="09E57651FFA1950CFCA0FEE4D1945A89" box="[894,1016,367,392]" name="Mexico" pageId="20" pageNumber="171">MEXICO</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="B336F823FFA1950CFBDAFEE4D60A5A89" box="[1028,1126,367,392]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||||
</emphasis>
|
||||
, Ejido La Sombra, Mun. de Villa Flores,
|
||||
<taxonomicName id="B6F24D42FFA1950CFEF3FE1FD05E5AAD" box="[301,562,404,428]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFA1950CFEF3FE1FD05E5AAD" box="[301,562,404,428]" italics="true" pageId="20" pageNumber="171">Ceratozamia mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFA1950CFD9DFE1FD0C35AAD" box="[579,687,404,428]" pageId="20" pageNumber="171" value="1994-05-01">1-V-1994</date>
|
||||
, M. A. Perez-Farrera (3); Mpio. Villa Flores,
|
||||
<quantity id="B60A9B24FFA1950CFB1CFE1FD76D5AAC" box="[1218,1281,404,429]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="171" unit="km" value="2.0">2 km</quantity>
|
||||
. Oeste Ejido: “La Sombra”,
|
||||
<quantity id="B60A9B24FFA1950CFEE0FE3CD3ED5AD1" box="[318,385,439,464]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="20" pageNumber="171" unit="m" value="850.0">850m</quantity>
|
||||
, oak forest on
|
||||
<taxonomicName id="B6F24D42FFA1950CFDEEFE33D1435ACE" authorityName="D. W. Stevenson" authorityYear="1982" box="[560,815,440,463]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="norstogii">
|
||||
<emphasis id="4386EAD3FFA1950CFDEEFE33D1435ACE" box="[560,815,440,463]" italics="true" pageId="20" pageNumber="171">Ceratozamia norstogii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFA1950CFC9EFE3CD1C15AD1" box="[832,941,439,464]" pageId="20" pageNumber="171" value="1995-05-01">1-V-1995</date>
|
||||
, M. A. Perez, (4); Mpio. Cintalpa, Rancho Cafetal,
|
||||
<taxonomicName id="B6F24D42FFA1950CFF22FE57D3975AF5" box="[252,507,476,500]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="alvaresii">
|
||||
<emphasis id="4386EAD3FFA1950CFF22FE57D3975AF5" box="[252,507,476,500]" italics="true" pageId="20" pageNumber="171">Ceratozamia alvaresii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFA1950CFDD1FE57D0EA5AF4" box="[527,646,476,501]" pageId="20" pageNumber="171" value="1996-03-04">4-III-1996</date>
|
||||
, A. Vovides, 1A, (1); Mpo. Cintalapa, Ejido Flor de
|
||||
<collectingRegion id="B336F823FFA1950CFACDFE57D71C5AF4" box="[1299,1392,476,501]" country="Mexico" name="Chiapas" pageId="20" pageNumber="171">Chiapas</collectingRegion>
|
||||
, ex
|
||||
<taxonomicName id="B6F24D42FFA1950CFF49FD8BD3F05916" authorityName="D. W. Stevenson" authorityYear="1982" box="[151,412,512,535]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="norstogii">
|
||||
<emphasis id="4386EAD3FFA1950CFF49FD8BD3F05916" box="[151,412,512,535]" italics="true" pageId="20" pageNumber="171">Ceratozamia norstogii</emphasis>
|
||||
</taxonomicName>
|
||||
<specimenCount id="67F4FD48FFA1950CFE72FD8BD3AD5916" box="[428,449,512,535]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||||
cone;
|
||||
<date id="054C1001FFA1950CFDC3FE74D0FD5919" box="[541,657,511,536]" pageId="20" pageNumber="171" value="2010-02">Feb 2010</date>
|
||||
, M.A. Perez Farrera (1); Mpo. Villa Flores, Ejido La Sombra,
|
||||
<taxonomicName id="B6F24D42FFA1950CFF49FDAFD3C9593D" box="[151,421,548,572]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFA1950CFF49FDAFD3C9593D" box="[151,421,548,572]" italics="true" pageId="20" pageNumber="171">Ceratozamia mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
<specimenCount id="67F4FD48FFA1950CFE69FDAED3A0593D" box="[439,460,549,572]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||||
cone,
|
||||
<date id="054C1001FFA1950CFDF2FDAFD0E1593D" box="[556,653,548,572]" pageId="20" pageNumber="171" value="2010-11">XI-2010</date>
|
||||
, M.A. Perez Farrera (25); Champerico, [GPS coord. omitted],
|
||||
<taxonomicName id="B6F24D42FFA1950CFF49FDC3D3F35961" box="[151,415,584,608]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFA1950CFF49FDC3D3F35961" box="[151,415,584,608]" italics="true" pageId="20" pageNumber="171">Ceratozamia mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
<specimenCount id="67F4FD48FFA1950CFE72FDC3D3AD595E" box="[428,449,584,607]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||||
cone,
|
||||
<date id="054C1001FFA1950CFDC8FDC3D0FE5961" box="[534,658,584,608]" pageId="20" pageNumber="171" value="2017-05-20">
|
||||
20-
|
||||
<bibRefCitation id="15634B30FFA1950CFDE4FDC3D0FE5961" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[570,658,584,608]" pageId="20" pageNumber="171" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">V-2017</bibRefCitation>
|
||||
</date>
|
||||
, H. Gomez Dominguez & W.Tang (46); Jaltenango, [GPS coord. omitted],
|
||||
<quantity id="B60A9B24FFA1950CFED5FDE7D3225984" box="[267,334,620,645]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="20" pageNumber="171" unit="m" value="957.0">957m</quantity>
|
||||
, ex
|
||||
<specimenCount id="67F4FD48FFA1950CFE57FDE6D3F25985" box="[393,414,621,644]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFA1950CFE73FDE7D0DC5985" box="[429,688,620,644]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFA1950CFE73FDE7D0DC5985" box="[429,688,620,644]" italics="true" pageId="20" pageNumber="171">Ceratozamia vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
cone,
|
||||
<date id="054C1001FFA1950CFCD6FDE7D1EF5985" box="[776,899,620,644]" pageId="20" pageNumber="171" value="2017-05-21">
|
||||
21-
|
||||
<bibRefCitation id="15634B30FFA1950CFCF4FDE7D1EF5985" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[810,899,620,644]" pageId="20" pageNumber="171" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">V-2017</bibRefCitation>
|
||||
</date>
|
||||
, H. Gomez Dominguez & W.Tang (71); La Concordia, Rancho Los Pinos, ex
|
||||
<specimenCount id="67F4FD48FFA1950CFDF8FD1BD05759A6" box="[550,571,656,679]" pageId="20" pageNumber="171" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFA1950CFD9BFD1BD12859A9" box="[581,836,656,680]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFA1950CFD9BFD1BD12859A9" box="[581,836,656,680]" italics="true" pageId="20" pageNumber="171">Ceratozamia vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
cone,
|
||||
<date id="054C1001FFA1950CFC4AFD1BD66159A9" box="[916,1037,656,680]" pageId="20" pageNumber="171" value="2017-06-01">
|
||||
1-
|
||||
<bibRefCitation id="15634B30FFA1950CFC7BFD1BD66159A9" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[933,1037,656,680]" pageId="20" pageNumber="171" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">VI-2017</bibRefCitation>
|
||||
</date>
|
||||
, M.A. Pérez Farrera & H. Gómez Domínguez (22).
|
||||
<typeStatus id="AE498863FFA1950CFE83FD3ED3A659CC" box="[349,458,693,717]" pageId="20" pageNumber="171" type="paratype">Paratypes</typeStatus>
|
||||
deposited at ANIC, BMNH, FSCA, IEXA, INBio, MNHN, NZAC, UNAM, USNM.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA1950CFF19FD5CD63659F1" box="[199,1114,727,752]" pageId="20" pageNumber="171" type="etymology">
|
||||
<paragraph id="714D36C1FFA1950CFF19FD5CD63659F1" blockId="20.[151,1437,151,896]" box="[199,1114,727,752]" pageId="20" pageNumber="171">
|
||||
<emphasis id="4386EAD3FFA1950CFF19FD5CD32259F1" bold="true" box="[199,334,727,752]" pageId="20" pageNumber="171">Etymology.</emphasis>
|
||||
This species is named for the Mexican state where it was discovered.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFA1950CFF19FD77D120587E" pageId="20" pageNumber="171" type="discussion">
|
||||
<paragraph id="714D36C1FFA1950CFF19FD77D120587E" blockId="20.[151,1437,151,896]" pageId="20" pageNumber="171">
|
||||
<emphasis id="4386EAD3FFA1950CFF19FD77D3575814" bold="true" box="[199,315,764,789]" pageId="20" pageNumber="171">Remarks.</emphasis>
|
||||
This species coinhabits male cones of at least four species of
|
||||
<taxonomicName id="B6F24D42FFA1950CFBE0FD77D6A35812" box="[1086,1231,764,787]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA1950CFBE0FD77D6A35812" box="[1086,1231,764,787]" italics="true" pageId="20" pageNumber="171">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="B6F24D42FFA1950CFB3BFD77D7025815" authorityName="Perez-Farrera, Vovides & Iglesias" authorityYear="1999" box="[1253,1390,764,788]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="alvarezii">
|
||||
<emphasis id="4386EAD3FFA1950CFB3BFD77D6945812" box="[1253,1272,764,787]" italics="true" pageId="20" pageNumber="171">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFA1950CFAD3FD77D7025815" box="[1293,1390,764,788]" italics="true" pageId="20" pageNumber="171">alvarezii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFA1950CFA5CFD77D3695839" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFA1950CFA5CFD77D7F95812" box="[1410,1429,764,787]" italics="true" pageId="20" pageNumber="171">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFA1950CFF49FCABD3695839" box="[151,261,800,824]" italics="true" pageId="20" pageNumber="171">mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="4386EAD3FFA1950CFEDBFCABD3D95836" box="[261,437,800,824]" italics="true" pageId="20" pageNumber="171">
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFA1950CFEC3FCABD3DE5836" authorityName="D. W. Stevenson" authorityYear="1982" box="[285,434,800,823]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="norstogii">C. norstogii</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
<taxonomicName id="B6F24D42FFA1950CFE17FCABD0355839" box="[457,601,800,824]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFA1950CFE17FCABD3B05836" box="[457,476,800,823]" italics="true" pageId="20" pageNumber="171">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFA1950CFE28FCABD0355839" box="[502,601,800,824]" italics="true" pageId="20" pageNumber="171">vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
) with
|
||||
<taxonomicName id="B6F24D42FFA1950CFD65FCABD6655836" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[699,1033,800,824]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA1950CFD65FCABD12F5839" box="[699,835,800,824]" italics="true" pageId="20" pageNumber="171">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA1950CFC81FCABD1155836" box="[863,889,800,823]" italics="true" pageId="20" pageNumber="171">C.</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFA1950CFC48FCABD6655836" box="[918,1033,800,823]" italics="true" pageId="20" pageNumber="171">picipennis</emphasis>
|
||||
</taxonomicName>
|
||||
and an undescribed species of
|
||||
<taxonomicName id="B6F24D42FFA1950CFF49FCCFD35B585D" authorityName="Reitter" authorityYear="1875" box="[151,311,836,860]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFA1950CFF49FCCFD35B585D" box="[151,311,836,860]" italics="true" pageId="20" pageNumber="171">Pharaxonotha</emphasis>
|
||||
</taxonomicName>
|
||||
. In nine collections from four host species listed above,
|
||||
<taxonomicName id="B6F24D42FFA1950CFBD6FCCFD7F0585D" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[1032,1436,836,860]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="20" pageNumber="171" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFA1950CFBD6FCCFD6FC585D" box="[1032,1168,836,860]" italics="true" pageId="20" pageNumber="171">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA1950CFB77FCCFD766585D" box="[1193,1290,836,860]" italics="true" pageId="20" pageNumber="171">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFA1950CFAF8FCCFD7F0585D" box="[1318,1436,836,860]" italics="true" pageId="20" pageNumber="171">chiapensis</emphasis>
|
||||
</taxonomicName>
|
||||
accounted for 6% of the adult pharaxonothine beetles present.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
155
data/F9/5B/87/F95B87D7FFB0951EFF49FA02D7175A1C.xml
Normal file
155
data/F9/5B/87/F95B87D7FFB0951EFF49FA02D7175A1C.xml
Normal file
|
|
@ -0,0 +1,155 @@
|
|||
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<mods:mods id="0D2AA722D9EA1CCF672B69DF9A7AD0A2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B30BEBB1355B62F1975C72B3AEC296D6">
|
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<mods:title id="A3A266323AAA39048A64788CEB002DCD">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
|
||||
</mods:titleInfo>
|
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<mods:name id="0F23FA7E9B5776C1BA153F5F02C9245C" type="personal">
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<mods:role id="6C67BBC6A77C8496BDD85DC992224BED">
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<mods:namePart id="38820C88A30DBCB641D3D53BB52F2195">Tang, William</mods:namePart>
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</mods:role>
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<mods:namePart id="26F353470367F54D5F4127B2F6B3CD05">Skelley, Paul</mods:namePart>
|
||||
</mods:name>
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<mods:name id="87B351EFBD4D4B9B6B8FEE8C89EFA476" type="personal">
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<mods:roleTerm id="A1099B9A7FEDBA5167349838EC1906BA">Author</mods:roleTerm>
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</mods:role>
|
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<mods:namePart id="430D408A4E08DC6A9B07D332C177FBB8">Pérez-Farrera, Miguel Angel</mods:namePart>
|
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</mods:name>
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<mods:typeOfResource id="C5A1977F6A15DC67611284F942348BC9">text</mods:typeOfResource>
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<mods:titleInfo id="6C8848C02828FB0CEB9154F6AA2419C3">
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<mods:title id="26349C3BEB2771748D71997F1464E44A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="76727DAF6BD7830BE3ADD8825EA93E27">
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<mods:date id="291C17A5990A01986286528568339B6E">2018</mods:date>
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<mods:number id="8C04BBB2F8E092FFB04D3EA3E0FDE6E9">2018-10-31</mods:number>
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<mods:number id="E2190A0974474310B2BE1AAC13D7102D">4508</mods:number>
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<mods:detail id="EB8BDE6CAFE980518D1004B6DB938B01" type="issue">
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<mods:number id="6C3CC65B10247946ED7035B0E5FAFCE7">2</mods:number>
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<mods:classification id="51A6F3530EBD51993A43D682DEE0594F">journal article</mods:classification>
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<mods:identifier id="6E00400E66437AD3C04B9C80889DBA71" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="F95B87D7FFB0951EFF49FA02D7175A1C" ID-DOI="http://doi.org/10.5281/zenodo.5958001" ID-GBIF-Taxon="154495719" ID-Zenodo-Dep="5958001" LSID="urn:lsid:plazi:treatment:F95B87D7FFB0951EFF49FA02D7175A1C" httpUri="http://treatment.plazi.org/id/F95B87D7FFB0951EFF49FA02D7175A1C" lastPageId="6" lastPageNumber="157" pageId="5" pageNumber="156">
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<subSubSection id="39E8654AFFB0951DFF49FA02D63A5EA2" box="[151,1110,1417,1443]" pageId="5" pageNumber="156" type="nomenclature">
|
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<paragraph id="714D36C1FFB0951DFF49FA02D63A5EA2" blockId="5.[151,1110,1417,1443]" box="[151,1110,1417,1443]" pageId="5" pageNumber="156">
|
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<heading id="2A0581ADFFB0951DFF49FA02D63A5EA2" bold="true" box="[151,1110,1417,1443]" fontSize="11" level="1" pageId="5" pageNumber="156" reason="1">
|
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<emphasis id="4386EAD3FFB0951DFF49FA02D63A5EA2" bold="true" box="[151,1110,1417,1443]" pageId="5" pageNumber="156">
|
||||
<taxonomicName id="B6F24D42FFB0951DFF49FA02D3BC5EA2" box="[151,464,1417,1443]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB0951DFF49FA02D34B5EA2" bold="true" box="[151,295,1417,1443]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFEE9FA02D3AA5EA2" bold="true" box="[311,454,1417,1443]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
Tang, Skelley & Pérez-Farrera, nominate subgenus
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB0951DFF49FA5BD1F75EE7" box="[151,923,1488,1510]" pageId="5" pageNumber="156" type="materials_examined">
|
||||
<paragraph id="714D36C1FFB0951DFF49FA5BD1F75EE7" blockId="5.[151,923,1488,1510]" box="[151,923,1488,1510]" pageId="5" pageNumber="156">
|
||||
<emphasis id="4386EAD3FFB0951DFF49FA5BD34E5EE7" bold="true" box="[151,290,1488,1510]" pageId="5" pageNumber="156">
|
||||
<typeStatus id="AE498863FFB0951DFF49FA5BD2A05EE7" box="[151,204,1488,1510]" pageId="5" pageNumber="156">Type</typeStatus>
|
||||
species.
|
||||
</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB0951DFEF7FA5BD0C85EE7" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[297,676,1488,1510]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB0951DFEF7FA5BD3C85EE7" box="[297,420,1488,1510]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFE6CFA5BD0415EE7" box="[434,557,1488,1510]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFB0951DFDE2FA5AD0C85EE7" box="[572,676,1489,1510]" italics="true" pageId="5" pageNumber="156">picipennis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB0951DFD6EFA5BD0855EE7" box="[688,745,1488,1510]" pageId="5" pageNumber="156" rank="species">
|
||||
<emphasis id="4386EAD3FFB0951DFD6EFA5BD0855EE7" bold="true" box="[688,745,1488,1510]" pageId="5" pageNumber="156">n. sp.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, here designated.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB0951DFF49F999D0D25D72" pageId="5" pageNumber="156" type="diagnosis">
|
||||
<paragraph id="714D36C1FFB0951DFF49F999D0D25D72" blockId="5.[151,1437,1554,2011]" pageId="5" pageNumber="156">
|
||||
<emphasis id="4386EAD3FFB0951DFF49F999D3355D2A" bold="true" box="[151,345,1554,1579]" pageId="5" pageNumber="156">Adult diagnosis.</emphasis>
|
||||
The subgenus
|
||||
<emphasis id="4386EAD3FFB0951DFDDBF998D0745D2B" box="[517,536,1555,1578]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
. (
|
||||
<taxonomicName id="B6F24D42FFB0951DFDEFF998D0D45D2A" box="[561,696,1555,1579]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB0951DFDEFF998D0D45D2A" box="[561,696,1555,1579]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
) is distinguished from the subgenus
|
||||
<emphasis id="4386EAD3FFB0951DFB82F998D6035D2B" box="[1116,1135,1555,1578]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
. (
|
||||
<taxonomicName id="B6F24D42FFB0951DFB59F998D6845D2A" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1159,1256,1555,1579]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="172" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFB0951DFB59F998D6845D2A" box="[1159,1256,1555,1579]" italics="true" pageId="5" pageNumber="156">Vovidesa</emphasis>
|
||||
</taxonomicName>
|
||||
) by the convex pronotal surface near lateral carina; lateral carina distinctly thicker anteriorly; metatibia triangularly dilated toward apex; and apparently lacking sexual dimorphism.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB0951DFF19F9F5D7F05CDB" pageId="5" pageNumber="156" type="description">
|
||||
<paragraph id="714D36C1FFB0951DFF19F9F5D7F05CDB" blockId="5.[151,1437,1554,2011]" pageId="5" pageNumber="156">
|
||||
<emphasis id="4386EAD3FFB0951DFF19F9F5D3CD5D96" bold="true" box="[199,417,1662,1687]" pageId="5" pageNumber="156">Adult description.</emphasis>
|
||||
Length
|
||||
<quantity id="B60A9B24FFB0951DFDDFF9F5D0CB5D97" box="[513,679,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="4.07" metricValueMax="5.11" metricValueMin="3.03" pageId="5" pageNumber="156" unit="mm" value="4.07" valueMax="5.11" valueMin="3.03">3.03–5.11 mm</quantity>
|
||||
, width
|
||||
<quantity id="B60A9B24FFB0951DFD23F9F5D1C85D97" box="[765,932,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="2.04" metricValueMin="1.26" pageId="5" pageNumber="156" unit="mm" value="1.65" valueMax="2.04" valueMin="1.26">1.26–2.04 mm</quantity>
|
||||
. Mouthparts: submentum without long erect or reclining setae. Pronotum: nearly quadrate, length/width = 0.78–0.90; anterior margin near eyes weakly emarginate, anterior angles slightly projecting, anterior angles rounded; surface near lateral carinae not explanate, carinae in dorsal view evenly arcuate; in lateral view thickening anteriorly, 2× thicker than posterior edge; pronotal disc lacking longitudinal groove extending anteriorly from lateral basal pore in margin. Elytra: in most populations often laterally with dark coloration that may extend to disc, maculation merges gradually with lighter ground color anteriad and posteriad; lacking visible setae on surface. Legs: protibia with distal lateral tooth strongly developed in both sexes, apex of tooth with cluster of 3 short stout spinules; meso- and metatibiae moderately angularly dilated at apex which is fringed with short stout spinules. Male genitalia: median lobe and tegmen compressed laterally, penile struts 1.15–1.90× longer than median lobe. Sexual dimorphism absent, except in
|
||||
<emphasis id="4386EAD3FFB0951DFB0EF848D68F5CDB" box="[1232,1251,1987,2010]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.
|
||||
<taxonomicName id="B6F24D42FFB0951DFB2FF848D7F85CDA" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1265,1428,1987,2011]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="chemnicki" subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFB24F848D7615CDB" box="[1274,1293,1987,2010]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFB0951DFAFDF848D7F85CDA" box="[1315,1428,1987,2011]" italics="true" pageId="5" pageNumber="156">chemnicki</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB3951EFF19FF1CD7175A1C" pageId="6" pageNumber="157" type="discussion">
|
||||
<paragraph id="714D36C1FFB3951EFF19FF1CD7175A1C" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="157">
|
||||
<emphasis id="4386EAD3FFB3951EFF19FF1CD3585BB1" bold="true" box="[199,308,151,176]" pageId="6" pageNumber="157">Remarks</emphasis>
|
||||
.
|
||||
<taxonomicName id="B6F24D42FFB3951EFE9AFF13D0035BB1" box="[324,623,152,176]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB3951EFE9AFF13D3A05BB1" box="[324,460,152,176]" italics="true" pageId="6" pageNumber="157">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFB3951EFE00FF13D0095BB1" box="[478,613,152,176]" italics="true" pageId="6" pageNumber="157">Ceratophila</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
is notably different from
|
||||
<emphasis id="4386EAD3FFB3951EFC44FF13D1C15BAE" box="[922,941,152,175]" italics="true" pageId="6" pageNumber="157">C</emphasis>
|
||||
. (
|
||||
<taxonomicName id="B6F24D42FFB3951EFC18FF13D64B5BB1" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[966,1063,152,176]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="172" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFB3951EFC18FF13D64B5BB1" box="[966,1063,152,176]" italics="true" pageId="6" pageNumber="157">Vovidesa</emphasis>
|
||||
</taxonomicName>
|
||||
) in an apparent lack of external sexual dimorphism (exception, see remarks under
|
||||
<taxonomicName id="B6F24D42FFB3951EFD3AFF37D1165BD5" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[740,890,188,212]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="157" phylum="Tracheophyta" rank="species" species="chemnicki">
|
||||
<emphasis id="4386EAD3FFB3951EFD3AFF37D1165BD5" box="[740,890,188,212]" italics="true" pageId="6" pageNumber="157">C. chemnicki</emphasis>
|
||||
</taxonomicName>
|
||||
) and in other noted external characters. Many populations of
|
||||
<taxonomicName id="B6F24D42FFB3951EFE9BFF6BD3BA5BF6" box="[325,470,224,247]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="157" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFB3951EFE9BFF6BD3BA5BF6" box="[325,470,224,247]" italics="true" pageId="6" pageNumber="157">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
appear to have a member of each subgenus on the male cones. The morphological differences between these beetles imply they occupy different niches on the cones, fulfilling different functions.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
349
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Normal file
349
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|
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@ -0,0 +1,349 @@
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<document id="078AD73D381A5F3B5AA2BF48775E28FC" ID-CLB-Dataset="28054" ID-DOI="10.11646/zootaxa.4508.2.1" ID-GBIF-Dataset="da79b654-bd50-4715-8ed3-80fa7a00d469" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606934" ID-ZooBank="5E2BC894-1919-4F63-8EF5-BAAC91913388" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585631772" checkinUser="plazi" docAuthor="Tang, William, Skelley, Paul & Pérez-Farrera, Miguel Angel" docDate="2018" docId="F95B87D7FFB1951DFF49F8E7D7FA5E38" docLanguage="en" docName="zootaxa.4508.2.1.pdf" docOrigin="Zootaxa 4508 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratophila (Ceratophila)" docType="key" docVersion="7" lastPageNumber="156" masterDocId="0562FFAFFFB59518FFDEFF8BD26C5B01" masterDocTitle="Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)" masterLastPageNumber="178" masterPageNumber="151" pageNumber="155" updateTime="1698712984586" updateUser="ExternalLinkService">
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<mods:title id="B34E4A54A6A7F38675741EEDE31B3446">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<mods:namePart id="9450ED148F148036C19D8DB3BD747634">Tang, William</mods:namePart>
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<mods:namePart id="E6C4676ACC6123B54FB5ECB9C5488D04">Skelley, Paul</mods:namePart>
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<mods:namePart id="03A18ACA4F49E118F5D5F3907E2437D5">Pérez-Farrera, Miguel Angel</mods:namePart>
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<mods:title id="7855219676FE8349F7EB53C61A9CFAF9">Zootaxa</mods:title>
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<mods:date id="86F571E016C86E71630318B738AE9085">2018</mods:date>
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<treatment id="F95B87D7FFB1951DFF49F8E7D7FA5E38" ID-DOI="http://doi.org/10.5281/zenodo.5957999" ID-GBIF-Taxon="154495718" ID-Zenodo-Dep="5957999" LSID="urn:lsid:plazi:treatment:F95B87D7FFB1951DFF49F8E7D7FA5E38" httpUri="http://treatment.plazi.org/id/F95B87D7FFB1951DFF49F8E7D7FA5E38" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155">
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<subSubSection id="39E8654AFFB1951DFF49F8E7D0415BD5" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155" type="nomenclature">
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<paragraph id="714D36C1FFB1951CFF49F8E7D3A55C87" blockId="4.[151,457,1900,1926]" box="[151,457,1900,1926]" pageId="4" pageNumber="155">
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<heading id="2A0581ADFFB1951CFF49F8E7D3A55C87" bold="true" box="[151,457,1900,1926]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
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<emphasis id="4386EAD3FFB1951CFF49F8E7D3A55C87" bold="true" box="[151,457,1900,1926]" pageId="4" pageNumber="155">
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||||
Key to adult
|
||||
<taxonomicName id="B6F24D42FFB1951CFEE7F8E7D3A55C87" box="[313,457,1900,1926]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB1951CFEE7F8E7D3A55C87" bold="true" box="[313,457,1900,1926]" italics="true" pageId="4" pageNumber="155">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB1951CFF49F83FD7F05CEE" blockId="4.[151,1436,1971,2032]" pageId="4" pageNumber="155">Because of variable and overlapping morphological character states with some species, it is desirable, and in some cases necessary, to have a series of specimens representing color variations and both sexes to properly identify a</paragraph>
|
||||
<paragraph id="714D36C1FFB0951DFF49FF1CD0415BD5" blockId="5.[151,1436,151,213]" pageId="5" pageNumber="156">
|
||||
species. In the following key, text in brackets before the species name are the Mexican states of occurrence and the known host species of
|
||||
<taxonomicName id="B6F24D42FFB0951DFE48FF37D04B5BD2" box="[406,551,188,211]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFB0951DFE48FF37D04B5BD2" box="[406,551,188,211]" italics="true" pageId="5" pageNumber="156">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB0951DFF49FE88D7FA5E38" pageId="5" pageNumber="156" type="key">
|
||||
<keyStep id="CA062E64FFB0951DFF49FE88D7F05AC9" pageId="5" pageNumber="156">
|
||||
<paragraph id="714D36C1FFB0951DFF49FE88D7F05A71" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FE88D7F05A71" pageId="5" pageNumber="156">
|
||||
1. Pronotum not explanate laterally, surface convexly curved to lateral carinae; lateral carinae in lateral view distinctly thicker anteriorly, anterior thickness 2× posterior thickness (
|
||||
<figureCitation id="E9C92A44FFB0951DFD3EFEABD1415A37" box="[736,813,288,310]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="5" pageNumber="156">Fig. 1D</figureCitation>
|
||||
); pronotal disc lacking longitudinal groove extending anteriorly from lateral basal pore in margin (
|
||||
<figureCitation id="E9C92A44FFB0951DFDF6FEB6D01F5A52" box="[552,627,317,339]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="5" pageNumber="156">Fig. 1B</figureCitation>
|
||||
); metatibia triangularly dilated toward apex (
|
||||
<figureCitation id="E9C92A44FFB0951DFBFAFEB6D6075A52" box="[1060,1131,317,339]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="5" pageNumber="156">Fig. 1F</figureCitation>
|
||||
)......................................................................................... [subgenus
|
||||
<taxonomicName id="B6F24D42FFB0951DFB85FED0D7055A71" box="[1115,1385,347,369]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="subGenus">
|
||||
<emphasis id="4386EAD3FFB0951DFB85FED0D7055A71" bold="true" box="[1115,1385,347,369]" pageId="5" pageNumber="156">
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<emphasis id="4386EAD3FFB0951DFB85FED0D6BC5A70" bold="true" box="[1115,1232,347,369]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
|
||||
, new subgenus
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
] ...
|
||||
<emphasis id="4386EAD3FFB0951DFA4EFED0D7F05A71" bold="true" box="[1424,1436,347,368]" pageId="5" pageNumber="156">2</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB0951DFF49FEF2D7F05AC9" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FEF2D7F05AC9" pageId="5" pageNumber="156">
|
||||
- Pronotum explanate laterally, surface broadly concave near lateral carinae; lateral carina weakly thickening anteriorly, anterior thickness <1.5× that at base (
|
||||
<figureCitation id="E9C92A44FFB0951DFDDAFE1ED0225AAA" box="[516,590,405,427]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="5" pageNumber="156">Fig. 1E</figureCitation>
|
||||
); pronotal disc with longitudinal groove extending anteriorly from lateral basal pore in margin (
|
||||
<figureCitation id="E9C92A44FFB0951DFEEFFE38D3135AC8" box="[305,383,435,457]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="5" pageNumber="156">Fig. 6D</figureCitation>
|
||||
); metatibia not triangularly dilated toward apex (
|
||||
<figureCitation id="E9C92A44FFB0951DFC8AFE38D1F25AC8" box="[852,926,435,457]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="5" pageNumber="156">Fig. 1G</figureCitation>
|
||||
)............... [
|
||||
<emphasis id="4386EAD3FFB0951DFBADFE38D7085AC9" bold="true" box="[1139,1380,435,457]" pageId="5" pageNumber="156">
|
||||
<taxonomicName id="B6F24D42FFB0951DFBADFE38D6AB5AC8" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1139,1223,435,457]" pageId="5" pageNumber="156" rank="subGenus" status="subgen. nov." subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFB0951DFBADFE38D6AB5AC8" bold="true" box="[1139,1223,435,457]" italics="true" pageId="5" pageNumber="156">Vovidesa</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB0951DFB08FE38D7085AC9" box="[1238,1380,435,456]" pageId="5" pageNumber="156" rank="subGenus">new subgenus</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
] …
|
||||
<emphasis id="4386EAD3FFB0951DFA4EFE38D7F05AC9" bold="true" box="[1424,1436,435,456]" pageId="5" pageNumber="156">5</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB0951DFF49FE5BD7F05997" pageId="5" pageNumber="156">
|
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<paragraph id="714D36C1FFB0951DFF49FE5BD7F7593F" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FE5BD7F7593F" pageId="5" pageNumber="156">
|
||||
2(1). Head with broad medial transverse ridge anterior of a distinct transverse basal groove in males (
|
||||
<figureCitation id="E9C92A44FFB0951DFB4DFE5BD68E5AE7" box="[1171,1250,464,486]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="5" pageNumber="156">Fig. 2D</figureCitation>
|
||||
), ridge and groove weak in females, ridge interrupting supraocular line in both male and female; head broad, width = 0.74–0.75× pronotal width; anterior clypeal margin emarginate; pronotal hypomeron densely punctate (
|
||||
<figureCitation id="E9C92A44FFB0951DFC6DFD80D1925920" box="[947,1022,523,545]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="5" pageNumber="156">Fig. 2E</figureCitation>
|
||||
); [Veracruz—
|
||||
<taxonomicName id="B6F24D42FFB0951DFB5AFD80D74C5921" ID-CoL="SQKV" box="[1156,1312,523,544]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="euryphyllidia">
|
||||
<emphasis id="4386EAD3FFB0951DFB5AFD80D6F85921" box="[1156,1172,523,544]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFB0951DFB7EFD80D74C5921" box="[1184,1312,523,544]" italics="true" pageId="5" pageNumber="156">euryphyllidia</emphasis>
|
||||
</taxonomicName>
|
||||
]....................................................................................
|
||||
<emphasis id="4386EAD3FFB0951DFBB2FDA3D7F7593F" bold="true" box="[1132,1435,552,574]" pageId="5" pageNumber="156">
|
||||
<emphasis id="4386EAD3FFB0951DFBB2FDA3D6ED593F" bold="true" box="[1132,1153,552,574]" italics="true" pageId="5" pageNumber="156">C.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB0951DFB59FDA2D776593F" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1159,1306,552,574]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="chemnicki" status="sp. nov." subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFB51FDA3D6F3593F" bold="true" box="[1167,1183,552,574]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFB0951DFB6DFDA3D776593F" bold="true" box="[1203,1306,552,574]" italics="true" pageId="5" pageNumber="156">chemnicki</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB0951DFAF8FDA2D7F7593F" box="[1318,1435,553,574]" pageId="5" pageNumber="156" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB0951DFF49FDCDD7F05997" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FDCDD7F05997" pageId="5" pageNumber="156">
|
||||
- Base of head at most with indistinct shallow transverse groove, supraocular line complete (
|
||||
<figureCitation id="E9C92A44FFB0951DFB86FDCED6CA595A" box="[1112,1190,581,603]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="5" pageNumber="156">Fig. 3D</figureCitation>
|
||||
); head narrower, width = 0.66–0.68× pronotal width; anterior clypeal margin truncate or weakly convex; pronotal hypomeron with punctures sparse and minute or lacking (
|
||||
<figureCitation id="E9C92A44FFB0951DFE49FD0BD3B35997" box="[407,479,640,662]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="5" pageNumber="156">Fig. 3E</figureCitation>
|
||||
)..............................................................................
|
||||
<emphasis id="4386EAD3FFB0951DFA4EFD0AD7F05997" bold="true" box="[1424,1436,641,662]" pageId="5" pageNumber="156">3</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB0951DFF49FD16D7F05828" pageId="5" pageNumber="156">
|
||||
<paragraph id="714D36C1FFB0951DFF49FD16D7F759EF" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FD16D7F759EF" pageId="5" pageNumber="156">
|
||||
3(2). Dark elytral markings (when present) laterally covering most of disc (
|
||||
<figureCitation id="E9C92A44FFB0951DFC53FD16D1B159B2" box="[909,989,669,691]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="5" pageNumber="156">Fig. 4A</figureCitation>
|
||||
); pronotum generally more elongate, pronotal length/pronotal width (PL/PW) = 0.83–0.90; [Chiapas—
|
||||
<taxonomicName id="B6F24D42FFB0951DFCDEFD37D10359D0" ID-CoL="SQKM" authorityName="Perez-Farrera, Vovides & Iglesias" authorityYear="1999" box="[768,879,700,721]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="alvarezii">
|
||||
<emphasis id="4386EAD3FFB0951DFCDEFD37D17C59D0" box="[768,784,700,721]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFB0951DFCC2FD37D10359D0" box="[796,879,700,721]" italics="true" pageId="5" pageNumber="156">alvarezii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFB0951DFCA5FD37D19859D0" ID-CoL="SQLB" box="[891,1012,700,721]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFB0951DFCA5FD37D1E759D0" box="[891,907,700,721]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFB0951DFC48FD37D19859D0" box="[918,1012,700,721]" italics="true" pageId="5" pageNumber="156">mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="4386EAD3FFB0951DFC2BFD37D61B59D0" box="[1013,1143,700,721]" italics="true" pageId="5" pageNumber="156">
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFB0951DFC20FD37D61F59D0" ID-CoL="69L9T" authorityName="D. W. Stevenson" authorityYear="1982" box="[1022,1139,700,721]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="norstogii">C. norstogii</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB0951DFBA3FD37D68259D0" ID-CoL="SQLM" box="[1149,1262,700,721]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFB0951DFBA3FD37D6E159D0" box="[1149,1165,700,721]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFB0951DFB47FD37D68259D0" box="[1177,1262,700,721]" italics="true" pageId="5" pageNumber="156">vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
].........................................................................................
|
||||
<emphasis id="4386EAD3FFB0951DFBB3FD53D7F759EF" bold="true" box="[1133,1435,728,750]" pageId="5" pageNumber="156">
|
||||
<emphasis id="4386EAD3FFB0951DFBB3FD53D6EF59EF" bold="true" box="[1133,1155,728,750]" italics="true" pageId="5" pageNumber="156">C.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB0951DFB57FD52D77559EF" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1161,1305,728,750]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="picipennis" status="sp. nov." subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFB4FFD53D6CD59EF" bold="true" box="[1169,1185,728,750]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFB0951DFB6AFD53D77559EF" bold="true" box="[1204,1305,728,750]" italics="true" pageId="5" pageNumber="156">picipennis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB0951DFAF8FD52D7F759EF" box="[1318,1435,729,750]" pageId="5" pageNumber="156" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB0951DFF49FD7DD7F05828" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FD7DD7F05828" pageId="5" pageNumber="156">
|
||||
- Dark elytral markings (when present) only along suture and laterally (
|
||||
<figureCitation id="E9C92A44FFB0951DFC53FD7ED1B7580A" box="[909,987,757,779]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="5" pageNumber="156">Fig. 5A</figureCitation>
|
||||
); pronotum generally shorter, PL/PW = 0.80– 0.84................................................................................................
|
||||
<emphasis id="4386EAD3FFB0951DFA4EFC9FD7F05828" bold="true" box="[1424,1436,788,809]" pageId="5" pageNumber="156">4</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB0951DFF49FCBBD7F758F7" pageId="5" pageNumber="156">
|
||||
<paragraph id="714D36C1FFB0951DFF49FCBBD7F75880" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FCBBD7F75880" pageId="5" pageNumber="156">
|
||||
4(3). Color of pronotum and elytra similar (
|
||||
<figureCitation id="E9C92A44FFB0951DFD8EFCBBD0F15847" box="[592,669,816,838]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="5" pageNumber="156">Fig. 5A</figureCitation>
|
||||
); pronotum generally more quadrate; male genitalia with penile struts relatively long, length of penile strut relative to median lobe 2:1; setae at apex of parameres with greatest length <dorsal width of parameres (
|
||||
<figureCitation id="E9C92A44FFB0951DFE8EFCE0D3F55880" box="[336,409,875,897]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="5" pageNumber="156">Fig. 5F</figureCitation>
|
||||
); [Veracruz—
|
||||
<taxonomicName id="B6F24D42FFB0951DFDFEFCE7D0195880" ID-CoL="SQLK" box="[544,629,876,897]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFB0951DFDFEFCE7D0195880" box="[544,629,876,897]" italics="true" pageId="5" pageNumber="156">C. tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
].........................................
|
||||
<emphasis id="4386EAD3FFB0951DFBB3FCE0D7F75880" bold="true" box="[1133,1435,875,897]" pageId="5" pageNumber="156">
|
||||
<emphasis id="4386EAD3FFB0951DFBB3FCE0D6EF5880" bold="true" box="[1133,1155,875,897]" italics="true" pageId="5" pageNumber="156">C.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB0951DFB56FCE7D7755880" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1160,1305,875,897]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="sanchezae" status="sp. nov." subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFB4FFCE0D6CD5880" bold="true" box="[1169,1185,875,897]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFB0951DFB6AFCE0D7755880" bold="true" box="[1204,1305,875,897]" italics="true" pageId="5" pageNumber="156">sanchezae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB0951DFAF8FCE7D7F75880" box="[1318,1435,876,897]" pageId="5" pageNumber="156" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB0951DFF49FC02D7F758F7" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FC02D7F758F7" pageId="5" pageNumber="156">
|
||||
- Pronotum reddish brown contrasting with light brown of elytra (
|
||||
<figureCitation id="E9C92A44FFB0951DFC83FC03D1C0589F" box="[861,940,904,926]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="5" pageNumber="156">Fig. 3A</figureCitation>
|
||||
); pronotum generally more trapezoidal, narrowing anteriorly; male genitalia with penile struts shorter, length of penile struts relative to median lobe 3:2; setae at apex of parameres with greatest length ± dorsal width of parameres (
|
||||
<figureCitation id="E9C92A44FFB0951DFCF8FC48D11F58D8" box="[806,883,963,985]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="5" pageNumber="156">Fig. 3H</figureCitation>
|
||||
); [Oaxaca—
|
||||
<taxonomicName id="B6F24D42FFB0951DFC33FC4FD60558D8" box="[1005,1129,964,985]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="mixeorum">
|
||||
<emphasis id="4386EAD3FFB0951DFC33FC4FD19158D8" box="[1005,1021,964,985]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFB0951DFBD7FC4FD60558D8" box="[1033,1129,964,985]" italics="true" pageId="5" pageNumber="156">mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
]......................................................................................................
|
||||
<emphasis id="4386EAD3FFB0951DFBA1FC6BD7F758F7" bold="true" box="[1151,1435,992,1014]" pageId="5" pageNumber="156">
|
||||
<emphasis id="4386EAD3FFB0951DFBA1FC6BD6F958F7" bold="true" box="[1151,1173,992,1014]" italics="true" pageId="5" pageNumber="156">C.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB0951DFB42FC6AD77558F7" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1180,1305,992,1014]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="gregoryi" status="sp. nov." subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFB7DFC6BD6DF58F7" bold="true" box="[1187,1203,992,1014]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFB0951DFB19FC6BD77558F7" bold="true" box="[1223,1305,992,1014]" italics="true" pageId="5" pageNumber="156">gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB0951DFAF8FC6AD7F758F7" box="[1318,1435,993,1014]" pageId="5" pageNumber="156" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB0951DFF49FC76D7F75F88" pageId="5" pageNumber="156">
|
||||
<paragraph id="714D36C1FFB0951DFF49FC76D7F05F30" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FC76D7F05F30" pageId="5" pageNumber="156">
|
||||
5(1). Male meso- and metatibiae each with medial subapical emargination (
|
||||
<figureCitation id="E9C92A44FFB0951DFC47FC76D1845F12" box="[921,1000,1021,1043]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="5" pageNumber="156">Fig. 1G</figureCitation>
|
||||
); female abdominal segments weakly alutaceous, punctures and setae visible except on lateral quarters....................................................
|
||||
<emphasis id="4386EAD3FFB0951DFA4EFB97D7F05F30" bold="true" box="[1424,1436,1052,1073]" pageId="5" pageNumber="156">6</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB0951DFF49FBB2D7F75F88" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FBB2D7F75F88" pageId="5" pageNumber="156">
|
||||
- Male tibiae lacking medial subapical emargination; female abdominal segments strongly alutaceous (
|
||||
<figureCitation id="E9C92A44FFB0951DFB07FBB3D7445F4F" box="[1241,1320,1080,1102]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="5" pageNumber="156">Fig. 6B</figureCitation>
|
||||
), punctures obscured across middle, setae reduced [Chiapas—
|
||||
<taxonomicName id="B6F24D42FFB0951DFD1DFBDDD15E5F6A" ID-CoL="SQKM" authorityName="Perez-Farrera, Vovides & Iglesias" authorityYear="1999" box="[707,818,1110,1131]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="alvarezii">
|
||||
<emphasis id="4386EAD3FFB0951DFD1DFBDDD0BF5F6A" box="[707,723,1110,1131]" italics="true" pageId="5" pageNumber="156">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFB0951DFD01FBDDD15E5F6A" box="[735,818,1110,1131]" italics="true" pageId="5" pageNumber="156">alvarezii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFB0951DFCE3FBDDD1D95F6A" ID-CoL="SQLB" box="[829,949,1110,1131]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFB0951DFCE3FBDDD1D95F6A" box="[829,949,1110,1131]" italics="true" pageId="5" pageNumber="156">C. mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="4386EAD3FFB0951DFC1CFBDDD6C65F6A" box="[962,1194,1110,1131]" italics="true" pageId="5" pageNumber="156">
|
||||
<taxonomicName id="B6F24D42FFB0951DFC1CFBDDD6435F6A" box="[962,1071,1110,1131]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="norstogi">C. norstogi</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFB0951DFBE7FBDDD6C65F6A" ID-CoL="SQLM" box="[1081,1194,1110,1131]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="vovidesii">C. vovidesii</taxonomicName>
|
||||
</emphasis>
|
||||
]...............................................................................................
|
||||
<emphasis id="4386EAD3FFB0951DFBB3FBF8D7F75F88" bold="true" box="[1133,1435,1139,1161]" pageId="5" pageNumber="156">
|
||||
<emphasis id="4386EAD3FFB0951DFBB3FBF8D6EF5F88" bold="true" box="[1133,1155,1139,1161]" italics="true" pageId="5" pageNumber="156">C.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB0951DFB57FBFFD7765F88" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1161,1306,1139,1161]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="chiapensis" status="sp. nov." subGenus="Vovidesa">
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFB4FFBF8D6F25F88" bold="true" box="[1169,1182,1139,1161]" italics="true" pageId="5" pageNumber="156">V</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFB0951DFB6CFBF8D7765F88" bold="true" box="[1202,1306,1139,1161]" italics="true" pageId="5" pageNumber="156">chiapensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB0951DFAF8FBFFD7F75F88" box="[1318,1435,1140,1161]" pageId="5" pageNumber="156" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB0951DFF49FB1BD7FA5E38" pageId="5" pageNumber="156">
|
||||
<paragraph id="714D36C1FFB0951DFF49FB1BD7F75FE0" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FB1BD7F75FE0" pageId="5" pageNumber="156">
|
||||
6(5). Submentum of male and female with sparse patch of setae-bearing punctures, male with long setae projecting anteriorly (
|
||||
<figureCitation id="E9C92A44FFB0951DFAABFB1BD2925FC2" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="5" pageNumber="156">Fig. 7E</figureCitation>
|
||||
); male genitalia more dorsoventrally compressed, median lobe nearly cylindrical; [Veracruz—
|
||||
<taxonomicName id="B6F24D42FFB0951DFB56FB25D7685FC2" box="[1160,1284,1198,1219]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="mixeorum">
|
||||
<emphasis id="4386EAD3FFB0951DFB56FB25D7685FC2" box="[1160,1284,1198,1219]" italics="true" pageId="5" pageNumber="156">C. mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
]........................................................................................
|
||||
<emphasis id="4386EAD3FFB0951DFBAEFB40D7F75FE0" bold="true" box="[1136,1435,1227,1249]" pageId="5" pageNumber="156">
|
||||
<taxonomicName id="B6F24D42FFB0951DFBAEFB40D7755FE0" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1136,1305,1227,1249]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="mixeorum" status="sp. nov." subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFB0951DFBAEFB40D6EA5FE0" bold="true" box="[1136,1158,1227,1249]" italics="true" pageId="5" pageNumber="156">C.</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFB4AFB40D6C95FE0" bold="true" box="[1172,1189,1227,1249]" italics="true" pageId="5" pageNumber="156">V.</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFB0951DFB6AFB40D7755FE0" bold="true" box="[1204,1305,1227,1249]" italics="true" pageId="5" pageNumber="156">mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB0951DFAF8FB47D7F75FE0" box="[1318,1435,1228,1249]" pageId="5" pageNumber="156" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB0951DFF49FB62D7FA5E38" blockId="5.[151,1437,259,1337]" pageId="5" pageNumber="156">
|
||||
<keyLead id="CA0395F4FFB0951DFF49FB62D7FA5E38" pageId="5" pageNumber="156">
|
||||
- Submentum of male and female with dense patch of setae-bearing punctures, male with long setae projecting laterally (
|
||||
<figureCitation id="E9C92A44FFB0951DFAB5FB63D2805E1A" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="5" pageNumber="156">Figs. 8</figureCitation>
|
||||
E–F); male genitalia laterally compressed, median lobe laterally flattened and distinctly curved in lateral view; [Veracruz—
|
||||
<taxonomicName id="B6F24D42FFB0951DFA58FA8DD3765E38" ID-CoL="SQLK" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFB0951DFA58FA8DD3765E38" italics="true" pageId="5" pageNumber="156">C. tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
].......................................................................
|
||||
<emphasis id="4386EAD3FFB0951DFBA5FAA8D7FA5E38" bold="true" box="[1147,1430,1315,1337]" pageId="5" pageNumber="156">
|
||||
<taxonomicName id="B6F24D42FFB0951DFBA5FAA8D7785E38" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1147,1300,1315,1337]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="vazquezi" status="sp. nov." subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFB0951DFBA5FAA8D6FD5E38" bold="true" box="[1147,1169,1315,1337]" italics="true" pageId="5" pageNumber="156">C.</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFB0951DFB41FAA8D6DC5E38" bold="true" box="[1183,1200,1315,1337]" italics="true" pageId="5" pageNumber="156">V.</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFB0951DFB1EFAA8D7785E38" bold="true" box="[1216,1300,1315,1337]" italics="true" pageId="5" pageNumber="156">vazquezi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB0951DFAFEFAAFD7FA5E38" box="[1312,1430,1316,1337]" pageId="5" pageNumber="156" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
292
data/F9/5B/87/F95B87D7FFB39510FF49FEE4D3265D74.xml
Normal file
292
data/F9/5B/87/F95B87D7FFB39510FF49FEE4D3265D74.xml
Normal file
|
|
@ -0,0 +1,292 @@
|
|||
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<mods:title id="16A16531761867027332643B62F67E43">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<taxonomicName id="B6F24D42FFB3951EFF49FEE4D1B25A8B" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,990,367,394]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="157" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Ceratophila">
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<emphasis id="4386EAD3FFB3951EFF49FEE4D34B5A88" bold="true" box="[151,295,367,393]" italics="true" pageId="6" pageNumber="157">Ceratophila</emphasis>
|
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(
|
||||
<emphasis id="4386EAD3FFB3951EFEE9FEE4D3AA5A88" bold="true" box="[311,454,367,393]" italics="true" pageId="6" pageNumber="157">Ceratophila</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFB3951EFE09FEE4D0395A88" bold="true" box="[471,597,367,393]" italics="true" pageId="6" pageNumber="157">chemnicki</emphasis>
|
||||
Tang, Skelley & Pérez-Farrera
|
||||
</taxonomicName>
|
||||
,
|
||||
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||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB3951EFF49FE19D33E5AAA" box="[151,338,402,427]" pageId="6" pageNumber="157" type="description">
|
||||
<paragraph id="714D36C1FFB3951EFF49FE19D33E5AAA" blockId="6.[151,1146,367,427]" box="[151,338,402,427]" pageId="6" pageNumber="157">
|
||||
(
|
||||
<figureCitation id="E9C92A44FFB3951EFF41FE19D36D5AAA" box="[159,257,402,427]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="6" pageNumber="157">Figs. 1B</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFB3951EFED3FE18D3765AAA" box="[269,282,403,427]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="6" pageNumber="157">2</figureCitation>
|
||||
A–J)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB3951EFF49FE51D35859EF" pageId="6" pageNumber="157" type="diagnosis">
|
||||
<paragraph id="714D36C1FFB3951EFF49FE51D35859EF" blockId="6.[151,1437,474,2011]" pageId="6" pageNumber="157">
|
||||
<emphasis id="4386EAD3FFB3951EFF49FE51D3345AF2" bold="true" box="[151,344,474,499]" pageId="6" pageNumber="157">Adult diagnosis.</emphasis>
|
||||
Distinguished from all other
|
||||
<taxonomicName id="B6F24D42FFB3951EFD7CFE50D1465AF2" box="[674,810,475,499]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB3951EFD7CFE50D1465AF2" box="[674,810,475,499]" italics="true" pageId="6" pageNumber="157">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
spp. by several characters including males with a broad transverse ridge and distinct groove at base of head, but weak in females, ridge interrupting supraocular line in both sexes; head broad, width = 0.74× pronotal width; anterior clypeal margin emarginate; pronotal hypomeron densely punctate; pronotum in dorsal view with middle third of lateral carinae parallel (
|
||||
<figureCitation id="E9C92A44FFB3951EFBEFFDCDD6FA595E" box="[1073,1174,582,607]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="6" pageNumber="157">Figs. 1B</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFB3951EFB7BFDCCD6A7595E" box="[1189,1227,583,607]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="6" pageNumber="157">2D</figureCitation>
|
||||
); elytra uniformly brown without darkened punctation or dark markings; male genitalia with length of median lobe relative to penile struts ~ 1:1; and female terminal abdominal ventrite with apex flexed inward, giving the appearance of a swollen and concave margin (
|
||||
<figureCitation id="E9C92A44FFB3951EFE46FD39D38359CA" box="[408,495,690,715]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="6" pageNumber="157">Fig. 2J</figureCitation>
|
||||
); and with a known distribution in
|
||||
<collectingCountry id="09E57651FFB3951EFC41FD39D19B59CA" box="[927,1015,690,715]" name="Mexico" pageId="6" pageNumber="157">Mexico</collectingCountry>
|
||||
, state of
|
||||
<collectingRegion id="B336F823FFB3951EFBB2FD38D6BE59CB" box="[1132,1234,691,714]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="157">Veracruz</collectingRegion>
|
||||
, on
|
||||
<emphasis id="4386EAD3FFB3951EFAD5FD38D35859EF" italics="true" pageId="6" pageNumber="157">
|
||||
<taxonomicName id="B6F24D42FFB3951EFAD5FD38D35C59EF" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="157" phylum="Tracheophyta" rank="species" species="euryphyllidia">Ceratozamia euryphyllidia</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB39510FF19FD71D0C45F34" lastPageId="8" lastPageNumber="159" pageId="6" pageNumber="157" type="description">
|
||||
<paragraph id="714D36C1FFB3951EFF19FD71D09958A3" blockId="6.[151,1437,474,2011]" pageId="6" pageNumber="157">
|
||||
<emphasis id="4386EAD3FFB3951EFF19FD71D38F5812" bold="true" box="[199,483,762,787]" pageId="6" pageNumber="157">Adult male description.</emphasis>
|
||||
Length
|
||||
<quantity id="B60A9B24FFB3951EFD9CFD71D0845813" box="[578,744,762,787]" metricMagnitude="-3" metricUnit="m" metricValue="5.154999999999999" metricValueMax="5.25" metricValueMin="5.06" pageId="6" pageNumber="157" unit="mm" value="5.154999999999999" valueMax="5.25" valueMin="5.06">5.06–5.25 mm</quantity>
|
||||
, width
|
||||
<quantity id="B60A9B24FFB3951EFCE3FD70D18D5813" box="[829,993,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="2.045" metricValueMax="2.12" metricValueMin="1.97" pageId="6" pageNumber="157" unit="mm" value="2.045" valueMax="2.12" valueMin="1.97">1.97–2.12 mm</quantity>
|
||||
(n = 2). Body in dorsal view elongate, greatest width at middle of elytra; in lateral view slightly flattened dorsally. General body color uniformly redbrown (
|
||||
<figureCitation id="E9C92A44FFB3951EFF2AFCC9D325585A" box="[244,329,834,859]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="6" pageNumber="157">Figs. 2</figureCitation>
|
||||
A–F); elytra uniformly brown without darkened punctation or dark markings; dorsal surface punctate, shining and appearing glabrous, ventrally shining and appearing glabrous except mesoventrite and abdomen mostly covered with long procumbent setae.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB3951EFF19FC25D6B65E9A" blockId="6.[151,1437,474,2011]" pageId="6" pageNumber="157">
|
||||
<emphasis id="4386EAD3FFB3951EFF19FC25D36B58C6" bold="true" box="[199,263,942,967]" pageId="6" pageNumber="157">Head</emphasis>
|
||||
broad, width = 0.74–0.75× pronotal width; dorsal view conical, narrowed anteriorly, surface irregularly convex with distinct depressions on each side along fronto-clypeal suture and a broad transverse groove at base of head just behind a broad weak transverse ridge between the eyes, this ridge interrupts the supraocular line; distinctly punctured (
|
||||
<figureCitation id="E9C92A44FFB3951EFE59FB91D07C5F32" box="[391,528,1050,1075]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="6" pageNumber="157">Figs. 2A, D</figureCitation>
|
||||
), average distance between closest punctures 2× width of puncture; width
|
||||
<quantity id="B60A9B24FFB3951EFA83FB90D2915F57" metricMagnitude="-3" metricUnit="m" metricValue="1.24" metricValueMax="1.26" metricValueMin="1.22" pageId="6" pageNumber="157" unit="mm" value="1.24" valueMax="1.26" valueMin="1.22">1.22– 1.26 mm</quantity>
|
||||
; dorsal interocular distance
|
||||
<quantity id="B60A9B24FFB3951EFDE8FBB5D0B75F57" box="[566,731,1086,1111]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.6" metricValueMin="6.5" pageId="6" pageNumber="157" unit="mm" value="0.655" valueMax="0.66" valueMin="0.65">0.65–0.66 mm</quantity>
|
||||
; head width/dorsal interocular distance ratio 1.85–1.94; ventral interocular distance
|
||||
<quantity id="B60A9B24FFB3951EFEA7FBE8D0715F7B" box="[377,541,1123,1147]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.699999999999999" metricValueMin="4.3" pageId="6" pageNumber="157" unit="mm" value="0.44999999999999996" valueMax="0.47" valueMin="0.43">0.43–0.47 mm</quantity>
|
||||
, head width/ventral interocular distance ratio 2.68–2.86, anterior clypeal margin shallowly concave, base of head with broad transverse groove distinct in males. Eye with large black facets. Antennal length 3/4 pronotal width, 1× head width, antennomere I (scape) fairly large, slightly elongate; antennomere II slightly smaller than III; IV–VII small, width equals length; VIII width slightly more than length; club fairly large, IX and X similar in length; XI slightly longer than X, globular with angulate apex. Clypeus emarginate anteriorly and laterally, distinctly punctate. Mentum and submentum with moderate punctation, distance between nearest punctures approximately 1× own diameter on mentum, 2–3× own diameter on submentum, each puncture with a short seta. Gular area smooth, without punctation or setae, border with submentum flattened without punctation, medially with a small impressed circular area (
|
||||
<figureCitation id="E9C92A44FFB3951EFBA8FA09D6A25E9A" box="[1142,1230,1410,1435]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="6" pageNumber="157">Fig. 2E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB39510FF19FA2DD30F5A64" blockId="6.[151,1437,474,2011]" lastBlockId="8.[151,1437,151,1653]" lastPageId="8" lastPageNumber="159" pageId="6" pageNumber="157">
|
||||
<emphasis id="4386EAD3FFB3951EFF19FA2DD34C5EBE" bold="true" box="[199,288,1446,1471]" pageId="6" pageNumber="157">Thorax</emphasis>
|
||||
with pronotum quadrate in dorsal view, middle third of lateral margins parallel (
|
||||
<figureCitation id="E9C92A44FFB3951EFB79FA2DD7645EBE" box="[1191,1288,1446,1471]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="6" pageNumber="157">Figs. 1B</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFB3951EFACBFA2CD7565EBE" box="[1301,1338,1447,1471]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="6" pageNumber="157">2D</figureCitation>
|
||||
), length/ width ratio 0.79–0.82; with distinct marginal beads laterally and basally, anterior margin with weak bead medially; pronotal disc weakly flattened; anterior and posterior angles weakly developed; lateral carinae in dorsal view with basal ¼ expanding to parallel medial third, then gradually converging in apical ½; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/7 length of posterior margin from posterior corners just anterior to marginal bead, longitudinal furrows of disc lacking. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae approximately ¼ length of eye. Hypomeron with lateral ½ distinctly punctate, medially with few weak striations. Elytra together in dorsal view elongate-oval, convex; length/width 1.50–1.60, greatest width near midlength; without marginal line basally; 8 complete striae of moderate puncture size, stria 9 vague on apical 1/3; scutellary striole extending ¼ elytral length, with 10–15 punctures; intervals of striae with fine, shallow punctures; punctures of elytral striae apparently lacking seta. Mesoventrite with moderate to strong punctation, distance between nearest punctures approximately equal to diameter of punctures, puncture depth moderate. Metaventrite entirely glossy, with strong lateral punctation separated by 1–2× own diameter; medial surface smooth, finely punctured, separated by 5–6× own diameter; convex laterally, slightly flattened medially, metathoracic discrimen extending slightly over half metaventrite length. Legs stout, relatively similar in length and shape. Procoxa oval; mesocoxa globular; metacoxa transversely elongate-oval; trochanters obliquely truncate apically; femora robust, moderately compressed laterally; triangular tibiae shorter than femora, dilated to obliquely truncate apices; protibia with disto-lateral tooth (
|
||||
<figureCitation id="E9C92A44FFBD9510FD68FF54D16E5BF9" box="[694,770,223,248]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="8" pageNumber="159">Figs. 2</figureCitation>
|
||||
A–B), with apical fringe of short, stout spinules of posterior margin narrowly interrupted along concave medial margin, long stout setae on anterior margin medially, anterior margin bare laterally; meso- and metatibia with apical fringe of short, stout spinules on anterior margin, reduced on posterior margins.
|
||||
</paragraph>
|
||||
<caption id="258D6649FFB2951FFF49F81DD7145CE8" httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="7" pageNumber="158" startId="7.[151,250,1942,1964]" targetBox="[195,1399,193,1925]" targetPageId="7">
|
||||
<paragraph id="714D36C1FFB2951FFF49F81DD7145CE8" blockId="7.[151,1435,1942,2025]" pageId="7" pageNumber="158">
|
||||
<emphasis id="4386EAD3FFB2951FFF49F81DD37B5CAD" bold="true" box="[151,279,1942,1964]" pageId="7" pageNumber="158">FIGURE 2.</emphasis>
|
||||
Photographs of
|
||||
<taxonomicName id="B6F24D42FFB2951FFE12F81DD1225CAD" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[460,846,1942,1964]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="7" pageNumber="158" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB2951FFE12F81DD02B5CAD" box="[460,583,1942,1964]" italics="true" pageId="7" pageNumber="158">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFB2951FFD87F81DD0BF5CAD" box="[601,723,1942,1964]" italics="true" pageId="7" pageNumber="158">Ceratophila</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFB2951FFD39F81DD1225CAD" box="[743,846,1942,1964]" italics="true" pageId="7" pageNumber="158">chemnicki</emphasis>
|
||||
</taxonomicName>
|
||||
(A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="714D36C1FFBD9510FF19FEE4D67B5881" blockId="8.[151,1437,151,1653]" pageId="8" pageNumber="159">
|
||||
<emphasis id="4386EAD3FFBD9510FF19FEE4D32F5A89" bold="true" box="[199,323,367,392]" pageId="8" pageNumber="159">Abdomen.</emphasis>
|
||||
Ventrite I with intercoxal process narrow, with triangular point anteromedially; lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; ventrite surface with punctures bearing reclining setae; nearly coalescing apical punctures of ventrite V with setae length approximately 4× diameter of puncture; ventrites I–IV bearing evenly distributed punctation from side to side, coarse punctures laterally separated by 1× diameter, finer medially separated by 3-4× diameter, I–IV each with 2 or more median pairs of longer, semi-erect sensory hairs (often abraded). Male genital capsule fringed with fine setae. Male genitalia (tegmen and median lobe) oriented upside down (displayed right side up in
|
||||
<figureCitation id="E9C92A44FFBD9510FB66FD04D77D59A9" box="[1208,1297,655,680]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="8" pageNumber="159">Fig. 2G</figureCitation>
|
||||
to facilitate comparisons). Tegmen sclerotized, triangular, laterally compressed; with anterior region ring-like, posterior region sheath-like; lateral margins gradually converging posteriorly; apically with 2 elongate, somewhat bar-shaped parameres. Parameres dorso-ventrally compressed, height approximately 1/2 its own width, in dorso-ventral view length/width ratio = 2.24–2.53; apically with setae, length of longest setae> width of paramere in dorso-ventral view. Aedeagus with median lobe laterally compressed, sclerotized, in lateral view strongly curved, apex tapering to rounded point. Ratio of penile strut length to median lobe length 1.15 (n = 1).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBD9510FF19FC07D0C45F34" blockId="8.[151,1437,151,1653]" pageId="8" pageNumber="159">
|
||||
<emphasis id="4386EAD3FFBD9510FF19FC07D37258A4" bold="true" box="[199,286,908,933]" pageId="8" pageNumber="159">Female</emphasis>
|
||||
similar to male, legs not sexually dimorphic. Base of head with broad transverse ridge and groove weak. The female notably differs from the male, and other species in the genus, by the modified terminal abdominal ventrite with the medial apex strongly flexed inward, giving the appearance of a swollen and concave margin (
|
||||
<figureCitation id="E9C92A44FFBD9510FF29FC7CD3265F11" box="[247,330,1015,1040]" captionStart="FIGURE 2" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[195,1399,193,1925]" captionTargetId="figure@7.[195,1399,193,1925]" captionTargetPageId="7" captionText="FIGURE 2. Photographs of Ceratophila (Ceratophila) chemnicki (A–I: male holotype): A–C) dorsal, ventral and lateral habitus; D–F) head, dorsal, ventral and lateral views; G) median lobe still attached with tegmen, lateral and ventral views; H) male genital capsule, dorsal view; I) close up of parameres of tegmen, ventral view; J) terminal abdominal ventrite, female." httpUri="https://zenodo.org/record/2606938/files/figure.png" pageId="8" pageNumber="159">Fig. 2J</figureCitation>
|
||||
); surface of apical margin and inflexion densely punctate and setose; terminal abdominal tergite sclerotized, with small medial tooth on margin.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBD9510FF19FBB4D0DE5E31" pageId="8" pageNumber="159" type="materials_examined">
|
||||
<paragraph id="714D36C1FFBD9510FF19FBB4D0BE5F59" blockId="8.[151,1437,151,1653]" box="[199,722,1087,1112]" pageId="8" pageNumber="159">
|
||||
<materialsCitation id="C19A3C9CFFBD9510FF19FBB4D0A25F59" ID-GBIF-Occurrence="2028138741" box="[199,718,1087,1112]" collectorName="El Luchador" country="Mexico" location="Veracruz" pageId="8" pageNumber="159" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
|
||||
<emphasis id="4386EAD3FFBD9510FF19FBB4D3095F59" bold="true" box="[199,357,1087,1112]" pageId="8" pageNumber="159">
|
||||
<typeStatus id="AE498863FFBD9510FF19FBB4D36E5F59" box="[199,258,1087,1112]" pageId="8" pageNumber="159">Type</typeStatus>
|
||||
locality.
|
||||
</emphasis>
|
||||
<collectingCountry id="09E57651FFBD9510FEB2FBB4D3A85F59" box="[364,452,1087,1112]" name="Mexico" pageId="8" pageNumber="159">Mexico</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFBD9510FE11FBCBD05A5F56" box="[463,566,1088,1111]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="8" pageNumber="159">Veracruz</collectingRegion>
|
||||
,
|
||||
<collectorName id="DC075317FFBD9510FD9FFBB4D0A25F59" box="[577,718,1087,1112]" pageId="8" pageNumber="159">El Luchador</collectorName>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBD9510FF19FBEFD6E25F7D" blockId="8.[151,1437,151,1653]" box="[199,1166,1124,1149]" pageId="8" pageNumber="159">
|
||||
<emphasis id="4386EAD3FFBD9510FF19FBEFD3765F7C" bold="true" box="[199,282,1124,1149]" pageId="8" pageNumber="159">Range.</emphasis>
|
||||
Only known from El Luchador,
|
||||
<collectingRegion id="B336F823FFBD9510FD57FBEED0815F7D" box="[649,749,1125,1148]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="8" pageNumber="159">Veracruz</collectingRegion>
|
||||
inhabiting cones of
|
||||
<taxonomicName id="B6F24D42FFBD9510FC0FFBEFD6E65F7D" box="[977,1162,1124,1148]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="159" phylum="Tracheophyta" rank="species" species="euryphyllidia">
|
||||
<emphasis id="4386EAD3FFBD9510FC0FFBEFD1885F7A" box="[977,996,1124,1147]" italics="true" pageId="8" pageNumber="159">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFBD9510FC2FFBEFD6E65F7D" box="[1009,1162,1124,1148]" italics="true" pageId="8" pageNumber="159">euryphyllidia</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="4386EAD3FFBD9510FB54FBEFD6E25F7D" box="[1162,1166,1124,1148]" italics="true" pageId="8" pageNumber="159">.</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBD9510FF19FB0CD0DE5E31" blockId="8.[151,1437,151,1653]" pageId="8" pageNumber="159">
|
||||
<emphasis id="4386EAD3FFBD9510FF19FB0CD0545FA1" bold="true" box="[199,568,1159,1184]" pageId="8" pageNumber="159">
|
||||
Material examined.
|
||||
<materialsCitation id="C19A3C9CFFBD9510FE15FB0CD0545FA1" ID-GBIF-Occurrence="2028138748" box="[459,568,1159,1184]" collectingDate="2012-05-10" collectorName="M. A. Perez Farrera" country="Mexico" location="El Luchador" pageId="8" pageNumber="159" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="AE498863FFBD9510FE15FB0CD0545FA1" box="[459,568,1159,1184]" pageId="8" pageNumber="159" type="holotype">Holotype</typeStatus>
|
||||
</materialsCitation>
|
||||
</emphasis>
|
||||
(by designation) male of
|
||||
<taxonomicName id="B6F24D42FFBD9510FC54FB03D64A5FA1" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[906,1062,1160,1184]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="159" phylum="Tracheophyta" rank="species" species="chemnicki">
|
||||
<emphasis id="4386EAD3FFBD9510FC54FB03D64A5FA1" box="[906,1062,1160,1184]" italics="true" pageId="8" pageNumber="159">C. chemnicki</emphasis>
|
||||
</taxonomicName>
|
||||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||||
<collectingCountry id="09E57651FFBD9510FDAAFB27D0D65FC5" box="[628,698,1196,1220]" name="Mexico" pageId="8" pageNumber="159">MEX</collectingCountry>
|
||||
., V.C.,
|
||||
<location id="742D601AFFBD9510FCD4FB27D1FB5FC4" LSID="urn:lsid:plazi:treatment:F95B87D7FFB39510FF49FEE4D3265D74:742D601AFFBD9510FCD4FB27D1FB5FC4" box="[778,919,1196,1221]" country="Mexico" name="El Luchador" pageId="8" pageNumber="159">El Luchador</location>
|
||||
, ex
|
||||
<specimenCount id="67F4FD48FFBD9510FC18FB26D1B75FC5" box="[966,987,1197,1220]" pageId="8" pageNumber="159" type="male">♂</specimenCount>
|
||||
cone
|
||||
<taxonomicName id="B6F24D42FFBD9510FBC1FB27D7205FC5" box="[1055,1356,1196,1220]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="159" phylum="Tracheophyta" rank="species" species="euryphyllidia">
|
||||
<emphasis id="4386EAD3FFBD9510FBC1FB27D7205FC5" box="[1055,1356,1196,1220]" italics="true" pageId="8" pageNumber="159">Ceratozamia euryphyllidia</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<collectingDate id="1508E9E9FFBD9510FA82FB27D2BE5FE9" pageId="8" pageNumber="159" value="2012-05-10">
|
||||
<date id="054C1001FFBD9510FA82FB27D2BE5FE9" pageId="8" pageNumber="159" value="2012-05-10">10-V- 2012</date>
|
||||
</collectingDate>
|
||||
,
|
||||
<collectorName id="DC075317FFBD9510FF02FB5AD3AC5FE9" box="[220,448,1232,1256]" pageId="8" pageNumber="159">M. A. Perez Farrera</collectorName>
|
||||
”; 2) [rectangular; red; printed in black ink]
|
||||
<materialsCitation id="C19A3C9CFFBD9510FC7CFB5BD1675E0D" ID-GBIF-Occurrence="2028138751" collectionCode="FSCA" country="Mexico" pageId="8" pageNumber="159" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="AE498863FFBD9510FC7CFB5BD65A5FE9" box="[930,1078,1232,1256]" pageId="8" pageNumber="159" type="holotype">HOLOTYPE</typeStatus>
|
||||
<specimenCount id="67F4FD48FFBD9510FBE2FB5AD63D5FE9" box="[1084,1105,1233,1256]" pageId="8" pageNumber="159" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFBD9510FB86FB5BD39C5E0D" authority="Tang, Skelley & Perez-Farrera 2018" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="159" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBD9510FB86FB5BD73A5FE9" box="[1112,1366,1232,1256]" italics="true" pageId="8" pageNumber="159">Ceratophila chemnicki</emphasis>
|
||||
<bibRefCitation id="15634B30FFBD9510FA83FB5AD39C5E0D" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." pageId="8" pageNumber="159" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">Tang, Skelley & Pérez-Farrera 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. Deposited in the
|
||||
<collectionCode id="17E3AE04FFBD9510FD1FFB7FD1675E0D" box="[705,779,1268,1292]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="8" pageNumber="159" type="Museum">FSCA</collectionCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="C19A3C9CFFBD9510FCC9FB7FD0C25E31" ID-GBIF-Occurrence="2028138752" collectionCode="FSCA, IEXA" country="Mexico" pageId="8" pageNumber="159" specimenCount="3" typeStatus="allotype">
|
||||
<typeStatus id="AE498863FFBD9510FCC9FB7FD1105E0C" box="[791,892,1268,1293]" pageId="8" pageNumber="159" type="allotype">
|
||||
<emphasis id="4386EAD3FFBD9510FCC9FB7FD1105E0C" bold="true" box="[791,892,1268,1293]" pageId="8" pageNumber="159">Allotype</emphasis>
|
||||
</typeStatus>
|
||||
(
|
||||
<collectionCode id="17E3AE04FFBD9510FC53FB7FD1BA5E0D" box="[909,982,1268,1292]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="8" pageNumber="159" type="Museum">FSCA</collectionCode>
|
||||
) and three adult
|
||||
<typeStatus id="AE498863FFBD9510FB44FB7FD77C5E0C" box="[1178,1296,1268,1293]" pageId="8" pageNumber="159" type="paratype">
|
||||
<emphasis id="4386EAD3FFBD9510FB44FB7FD77C5E0C" bold="true" box="[1178,1296,1268,1293]" pageId="8" pageNumber="159">paratypes</emphasis>
|
||||
</typeStatus>
|
||||
(same label data as
|
||||
<typeStatus id="AE498863FFBD9510FF35FA9CD3235E31" box="[235,335,1303,1328]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
|
||||
) deposited at
|
||||
<collectionCode id="17E3AE04FFBD9510FE35FA93D05D5E31" box="[491,561,1304,1328]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="8" pageNumber="159" type="Museum">FSCA</collectionCode>
|
||||
and
|
||||
<collectionCode id="17E3AE04FFBD9510FDB9FA93D0C25E31" box="[615,686,1304,1328]" pageId="8" pageNumber="159">IEXA</collectionCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBD9510FF19FAB7D0645E9C" pageId="8" pageNumber="159" type="etymology">
|
||||
<paragraph id="714D36C1FFBD9510FF19FAB7D0645E9C" blockId="8.[151,1437,151,1653]" pageId="8" pageNumber="159">
|
||||
<emphasis id="4386EAD3FFBD9510FF19FAB7D3225E54" bold="true" box="[199,334,1340,1365]" pageId="8" pageNumber="159">Etymology.</emphasis>
|
||||
The species is named in honor of Jeff Chemnick, who assisted in the collection of the
|
||||
<typeStatus id="AE498863FFBD9510FAB2FAB6D7F05E54" box="[1388,1436,1341,1365]" pageId="8" pageNumber="159">type</typeStatus>
|
||||
specimens of this species and others studied in this paper and for his contributions to the taxonomy, ecology and conservation of Mexican cycads.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBD9510FF19FA2CD3265D74" pageId="8" pageNumber="159" type="discussion">
|
||||
<paragraph id="714D36C1FFBD9510FF19FA2CD3265D74" blockId="8.[151,1437,151,1653]" pageId="8" pageNumber="159">
|
||||
<emphasis id="4386EAD3FFBD9510FF19FA2CD3575EC1" bold="true" box="[199,315,1447,1472]" pageId="8" pageNumber="159">Remarks.</emphasis>
|
||||
While
|
||||
<taxonomicName id="B6F24D42FFBD9510FE4FFA23D04A5EC1" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[401,550,1448,1472]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="159" phylum="Tracheophyta" rank="species" species="chemnicki">
|
||||
<emphasis id="4386EAD3FFBD9510FE4FFA23D04A5EC1" box="[401,550,1448,1472]" italics="true" pageId="8" pageNumber="159">C. chemnicki</emphasis>
|
||||
</taxonomicName>
|
||||
possesses the primary characters which place it in
|
||||
<emphasis id="4386EAD3FFBD9510FBABFA23D6E45EBE" box="[1141,1160,1448,1471]" italics="true" pageId="8" pageNumber="159">C</emphasis>
|
||||
. (
|
||||
<taxonomicName id="B6F24D42FFBD9510FB7FFA23D7445EC1" box="[1185,1320,1448,1472]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFBD9510FB7FFA23D7445EC1" box="[1185,1320,1448,1472]" italics="true" pageId="8" pageNumber="159">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
), it is has many other unique characters. Placement in this subgenus is considered tentative pending further molecular and phylogenetic analysis. The structure of the female terminal abdominal ventrite and tergite are unique. The single collection of this species from one male cone of
|
||||
<emphasis id="4386EAD3FFBD9510FD6EF99FD1885D2D" box="[688,996,1556,1580]" italics="true" pageId="8" pageNumber="159">
|
||||
<taxonomicName id="B6F24D42FFBD9510FD6EF99FD1B35D2D" box="[688,991,1556,1580]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="159" phylum="Tracheophyta" rank="species" species="euryphyllidia">Ceratozamia euryphyllidia</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
contained only
|
||||
<specimenCount id="67F4FD48FFBD9510FB48F99FD69C5D2C" box="[1174,1264,1556,1581]" pageId="8" pageNumber="159" type="adult">6 adults</specimenCount>
|
||||
, accounting for 4% of the
|
||||
<specimenCount id="67F4FD48FFBD9510FECBF9BCD3ED5D51" box="[277,385,1591,1616]" pageId="8" pageNumber="159" type="adult">162 adult</specimenCount>
|
||||
beetles present in the sample. An undescribed species of
|
||||
<taxonomicName id="B6F24D42FFBD9510FBF6F9B3D6A45D51" authorityName="Reitter" authorityYear="1875" box="[1064,1224,1592,1616]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFBD9510FBF6F9B3D6A45D51" box="[1064,1224,1592,1616]" italics="true" pageId="8" pageNumber="159">Pharaxonotha</emphasis>
|
||||
</taxonomicName>
|
||||
accounted for the remaining 96%.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
246
data/F9/5B/87/F95B87D7FFB7951AFF49FC01D7F05DC2.xml
Normal file
246
data/F9/5B/87/F95B87D7FFB7951AFF49FC01D7F05DC2.xml
Normal file
|
|
@ -0,0 +1,246 @@
|
|||
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<mods:mods id="C97FF2C28B5AB21B97A77BDC576FB8CE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="66666307198A1BAA58144D63DFB2134E">
|
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<mods:title id="7417FA18EB37AE777B5336669ADD9B26">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
|
||||
</mods:titleInfo>
|
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||||
<mods:namePart id="A87299B1FD56CFCB553C58FB5C3C3A41">Tang, William</mods:namePart>
|
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|
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<mods:roleTerm id="EF366E3E0DBB16F970BC8E9A3DEF2FBC">Author</mods:roleTerm>
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</mods:role>
|
||||
<mods:namePart id="28DA45B46B1E4AC33BB2FF3FD6955D94">Skelley, Paul</mods:namePart>
|
||||
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|
||||
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|
||||
<mods:namePart id="C25B00F7DC85197ADBAE52FEB899AD17">Pérez-Farrera, Miguel Angel</mods:namePart>
|
||||
</mods:name>
|
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<mods:title id="A3E6CCD899C0533F90362CBF15D152B8">Zootaxa</mods:title>
|
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<mods:date id="CB421C7F98384EF86F5A51C3FBD6F5F3">2018</mods:date>
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<mods:detail id="5CCD8AAE2E2D5BA9048B7BE95E87F8FF" type="pubDate">
|
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<mods:number id="E74030DBD73F220F4FE93386383B94F4">2018-10-31</mods:number>
|
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<mods:detail id="DBEE305479A3BD005D965CBE7A8B44ED" type="volume">
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<mods:number id="155F1B8F09B4FDFD3D42A37DAB357008">4508</mods:number>
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||||
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|
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<mods:detail id="E9C9C6D6E97EBAF84715C37864D9F62C" type="issue">
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<mods:number id="D6E23B5186A159022783059946DC2261">2</mods:number>
|
||||
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|
||||
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<mods:start id="5A4F0F8DA6041AA5FE16A11059036B28">151</mods:start>
|
||||
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<mods:classification id="70ED7AF22CB52BBDC16FEA3F41648E45">journal article</mods:classification>
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<mods:identifier id="152CF5D5B45A52F8C0FA56BC512AC5F7" type="DOI">10.11646/zootaxa.4508.2.1</mods:identifier>
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<treatment id="F95B87D7FFB7951AFF49FC01D7F05DC2" ID-DOI="http://doi.org/10.5281/zenodo.5957995" ID-GBIF-Taxon="154495706" ID-Zenodo-Dep="5957995" LSID="urn:lsid:plazi:treatment:F95B87D7FFB7951AFF49FC01D7F05DC2" httpUri="http://treatment.plazi.org/id/F95B87D7FFB7951AFF49FC01D7F05DC2" lastPageNumber="153" pageId="2" pageNumber="153">
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<subSubSection id="39E8654AFFB7951AFF49FC01D6C458A5" box="[151,1192,906,932]" pageId="2" pageNumber="153" type="nomenclature">
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<paragraph id="714D36C1FFB7951AFF49FC01D6C458A5" blockId="2.[151,1192,906,932]" box="[151,1192,906,932]" pageId="2" pageNumber="153">
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<heading id="2A0581ADFFB7951AFF49FC01D6C458A5" bold="true" box="[151,1192,906,932]" fontSize="11" level="1" pageId="2" pageNumber="153" reason="1">
|
||||
<emphasis id="4386EAD3FFB7951AFF49FC01D6C458A5" bold="true" box="[151,1192,906,932]" pageId="2" pageNumber="153">
|
||||
Key to genera of
|
||||
<taxonomicName id="B6F24D42FFB7951AFEB2FC01D02F58A5" authorityName="Crowson" authorityYear="1952" box="[364,579,906,932]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||||
(modified from
|
||||
<bibRefCitation id="15634B30FFB7951AFCD1FC01D1D458A5" author="Leschen, R. A. B." box="[783,952,906,932]" pageId="2" pageNumber="153" refId="ref19416" refString="Leschen, R. A. B. (2003) Fauna of New Zealand 47. Erotylidae (Insecta: Coleoptera: Cucujoidea): Phylogeny and Review. Manaaki Whenua Press, Lincoln, 108 pp." type="book" year="2003">Leschen 2003</bibRefCitation>
|
||||
and
|
||||
<bibRefCitation id="15634B30FFB7951AFC2BFC01D6CC58A5" author="Xu, G. & Tang, W. & Skelley, P. & Liu, N. & Rich, S." box="[1013,1184,906,932]" pageId="2" pageNumber="153" pagination="251 - 278" refId="ref20834" refString="Xu, G., Tang, W., Skelley, P., Liu, N. & Rich, S. (2015) Cycadophila, a new genus (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas debaoensis (Cycadaceae) in Asia. Zootaxa, 3986 (3), 251 - 278. https: // doi. org / 10.11646 / zootaxa. 3986.3.1" type="journal article" year="2015">
|
||||
Xu
|
||||
<emphasis id="4386EAD3FFB7951AFBFCFC01D63158A5" bold="true" box="[1058,1117,906,932]" italics="true" pageId="2" pageNumber="153">et al.</emphasis>
|
||||
2015
|
||||
</bibRefCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB7951AFF49FC5BD7F05DC2" pageId="2" pageNumber="153" type="key">
|
||||
<keyStep id="CA062E64FFB7951AFF49FC5BD7F05F02" pageId="2" pageNumber="153">
|
||||
<paragraph id="714D36C1FFB7951AFF49FC5BD7F758E7" blockId="2.[151,1436,976,1732]" box="[151,1435,976,998]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FC5BD7F758E7" box="[151,1435,976,998]" pageId="2" pageNumber="153">
|
||||
1 Antennal club of 2 antennomeres..........................................................
|
||||
<taxonomicName id="B6F24D42FFB7951AFB00FC5BD7F758E7" authority="Jacobson" authorityName="Jacobson" authorityYear="1915" box="[1246,1435,976,998]" class="Insecta" family="Erotylidae" genus="Setariola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB7951AFB00FC5BD7F758E7" bold="true" box="[1246,1435,976,998]" pageId="2" pageNumber="153">
|
||||
<emphasis id="4386EAD3FFB7951AFB00FC5BD75B58E7" bold="true" box="[1246,1335,976,998]" italics="true" pageId="2" pageNumber="153">Setariola</emphasis>
|
||||
Jacobson
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB7951AFF49FC65D7F05F02" blockId="2.[151,1436,976,1732]" box="[151,1436,1005,1027]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FC65D7F05F02" box="[151,1436,1005,1027]" pageId="2" pageNumber="153">
|
||||
- Antennal club of 3 antennomeres.........................................................................
|
||||
<emphasis id="4386EAD3FFB7951AFA4EFC65D7F05F02" bold="true" box="[1424,1436,1006,1027]" pageId="2" pageNumber="153">2</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB7951AFF49FB80D7F05F3F" pageId="2" pageNumber="153">
|
||||
<paragraph id="714D36C1FFB7951AFF49FB80D7F75F20" blockId="2.[151,1436,976,1732]" box="[151,1435,1035,1057]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FB80D7F75F20" box="[151,1435,1035,1057]" pageId="2" pageNumber="153">
|
||||
2 Pronotal callosities present; vestiture of scale-like setae................................
|
||||
<taxonomicName id="B6F24D42FFB7951AFBA0FB80D7F75F20" authority="Rosenhauer" authorityName="Rosenhauer" authorityYear="1856" box="[1150,1435,1035,1057]" class="Insecta" family="Erotylidae" genus="Leucohimatium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB7951AFBA0FB80D7F75F20" bold="true" box="[1150,1435,1035,1057]" pageId="2" pageNumber="153">
|
||||
<emphasis id="4386EAD3FFB7951AFBA0FB80D7775F20" bold="true" box="[1150,1307,1035,1057]" italics="true" pageId="2" pageNumber="153">Leucohimatium</emphasis>
|
||||
Rosenhauer
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB7951AFF49FBA2D7F05F3F" blockId="2.[151,1436,976,1732]" box="[151,1436,1064,1086]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FBA2D7F05F3F" box="[151,1436,1064,1086]" pageId="2" pageNumber="153">
|
||||
- Pronotal callosities absent; vestiture of simple setae..........................................................
|
||||
<emphasis id="4386EAD3FFB7951AFA4EFBA2D7F05F3F" bold="true" box="[1424,1436,1065,1086]" pageId="2" pageNumber="153">3</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB7951AFF49FBCED7F05FB2" pageId="2" pageNumber="153">
|
||||
<paragraph id="714D36C1FFB7951AFF49FBCED7F05F78" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FBCED7F05F78" pageId="2" pageNumber="153">
|
||||
3 Eyes small, ventrally not encroaching upon gular area, distance between eye and lateral corner of mentum about equal to length of eye in ventral view; not associated with cycads......................................................
|
||||
<emphasis id="4386EAD3FFB7951AFA4EFBEFD7F05F78" bold="true" box="[1424,1436,1124,1145]" pageId="2" pageNumber="153">4</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB7951AFF49FB0AD7F05FB2" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FB0AD7F05FB2" pageId="2" pageNumber="153">
|
||||
- Eyes large, ventrally encroaching upon gular area, distance between eye and lateral corner of mentum <1/2 eye length in ventral view; most associated with cycads.....................................................................
|
||||
<emphasis id="4386EAD3FFB7951AFA4EFB15D7F05FB2" bold="true" box="[1424,1436,1182,1203]" pageId="2" pageNumber="153">5</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB7951AFF49FB30D7F05FEF" pageId="2" pageNumber="153">
|
||||
<paragraph id="714D36C1FFB7951AFF49FB30D7F05FD0" blockId="2.[151,1436,976,1732]" box="[151,1436,1211,1233]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FB30D7F05FD0" box="[151,1436,1211,1233]" pageId="2" pageNumber="153">
|
||||
4 Dorsal punctation strong and impressed....................................................
|
||||
<taxonomicName id="B6F24D42FFB7951AFB11FB30D7F05FD0" authority="Reitter" authorityName="Reitter" authorityYear="1878" box="[1231,1436,1211,1233]" class="Insecta" family="Erotylidae" genus="Henoticonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB7951AFB11FB30D7F05FD0" bold="true" box="[1231,1436,1211,1233]" pageId="2" pageNumber="153">
|
||||
<emphasis id="4386EAD3FFB7951AFB11FB30D7215FD0" bold="true" box="[1231,1357,1211,1233]" italics="true" pageId="2" pageNumber="153">Henoticonus</emphasis>
|
||||
Reitter
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB7951AFF49FB52D7F05FEF" blockId="2.[151,1436,976,1732]" box="[151,1436,1240,1262]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FB52D7F05FEF" box="[151,1436,1240,1262]" pageId="2" pageNumber="153">
|
||||
- Dorsal punctation weak and not strongly impressed.........................................
|
||||
<taxonomicName id="B6F24D42FFB7951AFB62FB53D7F05FEF" authority="Reitter" authorityName="Reitter" authorityYear="1876" box="[1212,1436,1240,1262]" class="Insecta" family="Erotylidae" genus="Loberogosmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB7951AFB62FB53D7F05FEF" bold="true" box="[1212,1436,1240,1262]" pageId="2" pageNumber="153">
|
||||
<emphasis id="4386EAD3FFB7951AFB62FB53D7215FEF" bold="true" box="[1212,1357,1240,1262]" italics="true" pageId="2" pageNumber="153">Loberogosmus</emphasis>
|
||||
Reitter
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB7951AFF49FB7ED7F05E9F" pageId="2" pageNumber="153">
|
||||
<paragraph id="714D36C1FFB7951AFF49FB7ED7F05E47" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FB7ED7F05E47" pageId="2" pageNumber="153">
|
||||
5 Submental-gular suture visible as depressed darkened area, often with row of punctures and setae; anterior pronotal margin of most with fine, distinct, complete marginal line; male genitalia with median lobe and tegmen laterally flattened and slightly to strongly twisted (at least basally); spiculum gastrale asymmetrical; Asia...............
|
||||
<taxonomicName id="B6F24D42FFB7951AFB8DFABBD7F05E47" authority="Xu, Tang & Skelley" authorityName="Xu, Tang & Skelley" box="[1107,1436,1328,1350]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB7951AFB8DFABBD7F05E47" bold="true" box="[1107,1436,1328,1350]" pageId="2" pageNumber="153">
|
||||
<emphasis id="4386EAD3FFB7951AFB8DFABBD6A35E47" bold="true" box="[1107,1231,1328,1350]" italics="true" pageId="2" pageNumber="153">Cycadophila</emphasis>
|
||||
Xu, Tang & Skelley
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB7951AFF49FAC5D7F05E9F" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FAC5D7F05E9F" pageId="2" pageNumber="153">
|
||||
- Submental-gular suture apparently lacking, surface flattened; anterior pronotal margin lacking marginal line, some with row of fine punctures; male genitalia with median lobe cylindrical and tegmen dorsoventrally or laterally flattened, but usually not twisted; spiculum gastrale symmetrical; Americas............................................................
|
||||
<emphasis id="4386EAD3FFB7951AFA4EFA02D7F05E9F" bold="true" box="[1424,1436,1417,1438]" pageId="2" pageNumber="153">6</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="CA062E64FFB7951AFF49FA2ED7F05DC2" pageId="2" pageNumber="153">
|
||||
<paragraph id="714D36C1FFB7951AFF49FA2ED7F05D30" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49FA2ED7F05D30" pageId="2" pageNumber="153">
|
||||
6 Lateral pronotal carina thicker anteriorly in lateral view (
|
||||
<figureCitation id="E9C92A44FFB7951AFCCBFA2ED1395EBA" box="[789,853,1445,1467]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">Figs. 1</figureCitation>
|
||||
D–E); head with supraocular lines and elytra lacking basal bead; stridulatory file at base of head separated by distance ~ 0.5× width of the scutellar shield (
|
||||
<figureCitation id="E9C92A44FFB7951AFBA4FA48D6A95ED8" box="[1146,1221,1475,1497]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">Fig. 1B</figureCitation>
|
||||
); tarsomeres I–III distinctly widened (
|
||||
<figureCitation id="E9C92A44FFB7951AFE5DFA6BD3A95EF7" box="[387,453,1504,1526]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="2" pageNumber="153">Figs. 4</figureCitation>
|
||||
D–E, 8F), metatarsomere I width ± length; male genitalia with median lobe arched in lateral view; flagellum length Ĺ length of median lobe and not coiled (
|
||||
<figureCitation id="E9C92A44FFB7951AFD32FA76D1535D12" box="[748,831,1533,1555]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="2" pageNumber="153">Figs. 4G</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFB7951AFC97FA76D10F5D12" box="[841,867,1533,1555]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="2" pageNumber="153">5F</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFB7951AFCB2FA76D1E05D12" box="[876,908,1533,1555]" captionStart="FIGURE 6" captionStartId="19.[151,250,1923,1945]" captionTargetBox="[203,1389,193,1908]" captionTargetId="figure@19.[203,1389,193,1908]" captionTargetPageId="19" captionText="FIGURE 6. Photographs of Ceratophila (Uovidesa) chiapensis collected from Ceratozamia vovidesii (A–F: female allotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal, ventral and lateral views; G) (left to right) male genital capsule, median lobe and tegmen, ventral views; H) median lobe and tegmen, lateral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606948/files/figure.png" pageId="2" pageNumber="153">6H</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFB7951AFC48FA76D1DF5D12" box="[918,947,1533,1555]" captionStart="FIGURE 7" captionStartId="22.[151,250,1950,1972]" captionTargetBox="[176,1417,195,1935]" captionTargetId="figure@22.[176,1417,195,1935]" captionTargetPageId="22" captionText="FIGURE 7. Photographs of Ceratophila (Uovidesa) mixeorum (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) (left to right) male genital capsule, median lobe and tegmen, ventral views; G) (top to bottom) median lobe and tegmen, lateral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606950/files/figure.png" pageId="2" pageNumber="153">7G</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFB7951AFC62FA76D1B05D12" box="[956,988,1533,1555]" captionStart="FIGURE 8" captionStartId="24.[151,250,1934,1956]" captionTargetBox="[175,1416,193,1915]" captionTargetId="figure@24.[175,1416,193,1915]" captionTargetPageId="24" captionText="FIGURE 8. Photographs of Ceratophila (Uovidesa) vazquezi (A–D, F: male holotype): A–C) dorsal, ventral and lateral and habitus; D) head, ventral view; E) head, female, ventral view; F) head lateral view; G) median lobe still attached to tegmen, ventral and lateral views; H) male genital capsule, ventral view; arrow indicates tuft of erect setae on mentum and submentum." httpUri="https://zenodo.org/record/2606952/files/figure.png" pageId="2" pageNumber="153">8G</figureCitation>
|
||||
);
|
||||
<collectingCountry id="09E57651FFB7951AFC30FA76D65B5D12" box="[1006,1079,1533,1555]" name="Mexico" pageId="2" pageNumber="153">Mexico</collectingCountry>
|
||||
and probably
|
||||
<collectingCountry id="09E57651FFB7951AFB1CFA76D76E5D12" box="[1218,1282,1533,1555]" name="Belize" pageId="2" pageNumber="153">Belize</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="09E57651FFB7951AFAD5FA76D71F5D12" box="[1291,1395,1533,1555]" name="Guatemala" pageId="2" pageNumber="153">Guatemala</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="09E57651FFB7951AFF3FF990D3525D30" box="[225,318,1563,1585]" name="Honduras" pageId="2" pageNumber="153">Honduras</collectingCountry>
|
||||
on
|
||||
<taxonomicName id="B6F24D42FFB7951AFEBCF997D3B25D30" ID-CoL="3KXP" box="[354,478,1564,1585]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFB7951AFEBCF997D3B25D30" box="[354,478,1564,1585]" italics="true" pageId="2" pageNumber="153">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
............................................................
|
||||
<emphasis id="4386EAD3FFB7951AFB6FF990D7F05D30" bold="true" box="[1201,1436,1563,1585]" pageId="2" pageNumber="153">
|
||||
<taxonomicName id="B6F24D42FFB7951AFB6FF990D74A5D30" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1201,1318,1563,1585]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus" status="gen. nov.">
|
||||
<emphasis id="4386EAD3FFB7951AFB6FF990D74A5D30" bold="true" box="[1201,1318,1563,1585]" italics="true" pageId="2" pageNumber="153">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB7951AFAECF997D7F05D30" box="[1330,1436,1564,1585]" pageId="2" pageNumber="153" rank="genus">new genus</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB7951AFF49F9B2D7F05DC2" blockId="2.[151,1436,976,1732]" pageId="2" pageNumber="153">
|
||||
<keyLead id="CA0395F4FFB7951AFF49F9B2D7F05DC2" pageId="2" pageNumber="153">
|
||||
- Lateral pronotal carina narrow, same thickness along entire length in lateral view (
|
||||
<figureCitation id="E9C92A44FFB7951AFC2CF9B3D6505D4F" box="[1010,1084,1592,1614]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">Fig. 1C</figureCitation>
|
||||
); head without supraocular lines and elytra with distinct basal bead (
|
||||
<figureCitation id="E9C92A44FFB7951AFDCEF9DED01B5D6A" box="[528,631,1621,1643]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">
|
||||
Fig. 1A;
|
||||
<taxonomicName id="B6F24D42FFB7951AFDB6F9DDD01B5D6A" authority="Chaves and Genaro" authorityName="Chaves and Genaro" box="[616,631,1622,1643]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="esperanzae">
|
||||
<emphasis id="4386EAD3FFB7951AFDB6F9DDD01B5D6A" box="[616,631,1622,1643]" italics="true" pageId="2" pageNumber="153">P</emphasis>
|
||||
</taxonomicName>
|
||||
</figureCitation>
|
||||
. esperanzae Chaves and Genaro is the exception); stridulatory files at base of head separated by distance ~ width of scutellar shield (
|
||||
<figureCitation id="E9C92A44FFB7951AFD1CF9F8D1635D88" box="[706,783,1651,1673]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="2" pageNumber="153">Fig. 1A</figureCitation>
|
||||
); tarsomeres I–III narrow, metatarsomere I length always distinctly greater than width; male genitalia with median lobe straight; flagellum length >> length of median lobe and coiled; widespread New World on
|
||||
<taxonomicName id="B6F24D42FFB7951AFEADF925D3835DC2" ID-CoL="3KXP" box="[371,495,1710,1731]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFB7951AFEADF925D3835DC2" box="[371,495,1710,1731]" italics="true" pageId="2" pageNumber="153">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="4386EAD3FFB7951AFE25F925D6DF5DC2" box="[507,1203,1710,1731]" italics="true" pageId="2" pageNumber="153">
|
||||
<taxonomicName id="B6F24D42FFB7951AFE25F925D0555DC2" ID-CoL="62R3L" box="[507,569,1710,1731]" class="Cycadopsida" family="Zamiaceae" genus="Dioon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">Dioon</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFB7951AFD9DF925D0D95DC2" ID-CoL="5RPV" box="[579,693,1710,1731]" class="Cycadopsida" family="Zamiaceae" genus="Microcycas" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">Microcycas</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFB7951AFD60F925D0975DC2" ID-CoL="63X6Q" box="[702,763,1710,1731]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus">Zamia</taxonomicName>
|
||||
.....................................
|
||||
</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB7951AFB61F925D7F05DC2" ID-CoL="6MVR" authority="Reitter" authorityName="Reitter" authorityYear="1875" box="[1215,1436,1710,1732]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB7951AFB61F925D7215DC5" bold="true" box="[1215,1357,1710,1732]" italics="true" pageId="2" pageNumber="153">Pharaxonotha</emphasis>
|
||||
Reitter
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
223
data/F9/5B/87/F95B87D7FFB7951CFF49F8C5D0C55AAD.xml
Normal file
223
data/F9/5B/87/F95B87D7FFB7951CFF49F8C5D0C55AAD.xml
Normal file
|
|
@ -0,0 +1,223 @@
|
|||
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<mods:mods id="A8230F070FD4ABE946B3188A6BEC4B11" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="8383325702E2558B21BD19F2B4560353">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<mods:namePart id="63CEECB36E96FC5F8AE9F94203B339CF">Pérez-Farrera, Miguel Angel</mods:namePart>
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<paragraph id="714D36C1FFB7951AFF49F8C5D1F85C69" blockId="2.[151,916,1870,1896]" box="[151,916,1870,1896]" pageId="2" pageNumber="153">
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<emphasis id="4386EAD3FFB7951AFF49F8C5D1F85C69" bold="true" box="[151,916,1870,1896]" pageId="2" pageNumber="153">
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Genus
|
||||
<taxonomicName id="B6F24D42FFB7951AFF30F8C5D16A5C69" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[238,774,1870,1896]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="153" phylum="Tracheophyta" rank="genus" status="gen. nov.">
|
||||
<emphasis id="4386EAD3FFB7951AFF30F8C5D3115C69" bold="true" box="[238,381,1870,1896]" italics="true" pageId="2" pageNumber="153">Ceratophila</emphasis>
|
||||
Tang, Skelley & Pérez-Farrera
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFB7951AFCCDF8C5D1F85C69" box="[787,916,1870,1896]" pageId="2" pageNumber="153" rank="genus">new genus</taxonomicNameLabel>
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||||
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|
||||
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|
||||
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|
||||
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|
||||
<subSubSection id="39E8654AFFB7951AFF49F81ED15E5CAA" box="[151,818,1941,1963]" pageId="2" pageNumber="153" type="materials_examined">
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<paragraph id="714D36C1FFB7951AFF49F81ED15E5CAA" blockId="2.[151,818,1941,1963]" box="[151,818,1941,1963]" pageId="2" pageNumber="153">
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<typeStatus id="AE498863FFB7951AFF49F81ED2A05CAA" box="[151,204,1941,1963]" pageId="2" pageNumber="153">Type</typeStatus>
|
||||
species.
|
||||
</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB7951AFEF7F81ED02C5CAA" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[297,576,1941,1963]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="genus" status="sp. nov.">
|
||||
<emphasis id="4386EAD3FFB7951AFEF7F81ED3C85CAA" box="[297,420,1941,1963]" italics="true" pageId="2" pageNumber="153">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFB7951AFE6CF81DD3AF5CAA" box="[434,451,1942,1963]" italics="true" pageId="2" pageNumber="153">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFB7951AFE06F81DD02C5CAA" box="[472,576,1942,1963]" italics="true" pageId="2" pageNumber="153">picipennis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="58B557A8FFB7951AFD99F81ED0EC5CAA" box="[583,640,1941,1963]" pageId="2" pageNumber="153" rank="species">
|
||||
<emphasis id="4386EAD3FFB7951AFD99F81ED0EC5CAA" bold="true" box="[583,640,1941,1963]" pageId="2" pageNumber="153">n. sp.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, here designated.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB7951BFF49F853D72A5BF9" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153" type="diagnosis">
|
||||
<paragraph id="714D36C1FFB7951BFF49F853D72A5BF9" blockId="2.[151,1436,2008,2033]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153">
|
||||
<emphasis id="4386EAD3FFB7951AFF49F853D37D5CF0" bold="true" box="[151,273,2008,2033]" pageId="2" pageNumber="153">Diagnosis.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFB7951AFEC7F853D3CC5CF1" box="[281,416,2008,2032]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB7951AFEC7F853D3CC5CF1" box="[281,416,2008,2032]" italics="true" pageId="2" pageNumber="153">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
is distinguished from other currently known cycad-associated genera of
|
||||
<taxonomicName id="B6F24D42FFB7951AFB07F853D7225CF0" authorityName="Latreille" authorityYear="1802" box="[1241,1358,2008,2033]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="family">Erotylidae</taxonomicName>
|
||||
by the presence of the supraocular line, the thickened lateral pronotal carina, lack of a basal marginal bead on the elytra, the narrowly separated stridulatory files on the head, and by the male genitalia with median lobe and tegmen held upside down in body, median lobe arched in lateral view, flagellum <length of penile struts and not coiled.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB6951BFF19FE8FD19A5C29" pageId="3" pageNumber="154" type="description">
|
||||
<paragraph id="714D36C1FFB6951BFF19FE8FD3F75A89" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="154">
|
||||
<emphasis id="4386EAD3FFB6951BFF19FE8FD3D25A1C" bold="true" box="[199,446,260,285]" pageId="3" pageNumber="154">Description of adult.</emphasis>
|
||||
Length
|
||||
<quantity id="B60A9B24FFB6951BFDC2FE8FD0AE5A1D" box="[540,706,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="4.14" metricValueMax="5.25" metricValueMin="3.03" pageId="3" pageNumber="154" unit="mm" value="4.14" valueMax="5.25" valueMin="3.03">3.03–5.25 mm</quantity>
|
||||
, width
|
||||
<quantity id="B60A9B24FFB6951BFCC8FE8FD1D15A1D" box="[790,957,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="2.04" metricValueMin="1.26" pageId="3" pageNumber="154" unit="mm" value="1.65" valueMax="2.04" valueMin="1.26">1.26–2.04 mm</quantity>
|
||||
. Body form elongate, not strongly parallel sided; dorsoventrally flattened, weakly convex; widest anterior to or at middle of elytra; color pale to dark reddishbrown with or without variously sized dark macula on elytra; dorsal punctation fine, evenly distributed; setae yellow to gold colored.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB6951BFF19FE1FD0155F34" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="154">
|
||||
<emphasis id="4386EAD3FFB6951BFF19FE1FD36B5AAC" bold="true" box="[199,263,404,429]" pageId="3" pageNumber="154">Head</emphasis>
|
||||
with dorsal surface flat or with a pair of shallow longitudinal furrows on the frons and/or a transverse interocular furrow; clypeus flat, apically truncate. Eye globular with large black facets, projecting laterally, bordered dorsally with long supraocular line reaching anterior angle of eye. Vertex with supraocular line basally meeting and continuous with transverse occipital ridge (vertexal line) situated where retracted head meets anterior pronotal margin; occipital surface immediately posterior to transverse occipital ridge medially smooth and without punctures; stridulatory files posterior to transverse occipital ridge on occipital surface present, one on each side of the midline, closely spaced, weakly divergent anteriorly, distance between files <½ width of scutellar shield. Frons with short, dark, subcuticular suture on each side, approximately 1/4 width of frons in length, extending obliquely to lateral margin anterior to antennal insertion. Eye ventrally encroaching upon gular area, interocular distance> width of single eye in ventral view; lacking lateral temple behind eye. Antennomeres I–VII length subequal to width, moniliform in most, slightly elongate in others; antennomere VIII transverse or similar to antennomere VII, antennal club with 3 antennomeres; with some species variation, antennomere relative lengths approximately 18:13:10:10:10:10:10:10:15:15:15. Mouthparts: maxillary palp 4-segmented, palpomere I very small; II, III and IV longer than wide; IV cylindrical, elongate oval, 2× longer than preceding palpomere, acuminate apically, apex densely papillate; relative lengths of II– IV approximately 4:3:7. Labial palp with 3 palpomeres; palpomere I small, elongate; II transverse; III large, elongate-oval; terminal labial palpomere flattened, roughly oval in shape, 2/3–3/4 length of terminal maxillary palpomere. Mentum pentagonal with large triangular base defined by carina, projecting medially; lateral pockets in front of carina not evident; juncture of submentum and gula with shallow depression weakly circular or semicircular.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB6951BFF19FBB4D7765D09" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="154">
|
||||
<emphasis id="4386EAD3FFB6951BFF19FBB4D34A5F59" bold="true" box="[199,294,1087,1112]" pageId="3" pageNumber="154">Thorax.</emphasis>
|
||||
Pronotum quadrate or transverse; anterior marginal bead distinct laterally, reduced to row of fine punctures medially (weakly present only in
|
||||
<taxonomicName id="B6F24D42FFB6951BFD68FBEFD13D5F7D" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[694,849,1124,1148]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="3" pageNumber="154" phylum="Tracheophyta" rank="species" species="chemnicki">
|
||||
<emphasis id="4386EAD3FFB6951BFD68FBEFD13D5F7D" box="[694,849,1124,1148]" italics="true" pageId="3" pageNumber="154">C. chemnicki</emphasis>
|
||||
</taxonomicName>
|
||||
); anterior pronotal margin near eyes weakly to moderately emarginate, anterior angles slightly to moderately projecting, narrowly rounded; pronotal lateral carina in dorsal view arcuate, lacking modifications, medially convex, straight or slightly emarginated; lateral pronotal carinae in lateral view thickened anteriorly. Scutellar shield transverse, posterior margin clearly projected, pentagonal. Elytra finely setose, setae indistinct; base of elytra without marginal bead. Prosternum mostly flat near margins, convex and arched in center anterior to coxae, depressed posterior of coxa; prosternal process expanded apically, truncate and convex at apex; procoxal cavities externally closed or narrowly open posteriorly; hypomeron apparently impunctate laterally (except
|
||||
<taxonomicName id="B6F24D42FFB6951BFDB2FAEBD1695E79" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[620,773,1376,1400]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="3" pageNumber="154" phylum="Tracheophyta" rank="species" species="chemnicki">
|
||||
<emphasis id="4386EAD3FFB6951BFDB2FAEBD1695E79" box="[620,773,1376,1400]" italics="true" pageId="3" pageNumber="154">C. chemnicki</emphasis>
|
||||
</taxonomicName>
|
||||
), with longitudinal striations medially. Protibia apically angularly truncate and moderately to strongly dilated distally, with disto-lateral tooth. Tarsi with 5 tarsomeres, tarsomeres I–III distinctly widened, length often less than width, never longer than wide; tarsomeres I–III lobed beneath, pubescent ventrally with long, fine, dense, distally directed setae; tarsomere IV narrow and inconspicuous, without dense ventral setae; tarsomere V elongate, approximately length of tarsomeres I–IV combined.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB6951BFF19F99FD6BF5D2C" blockId="3.[151,1437,151,2013]" box="[199,1235,1556,1581]" pageId="3" pageNumber="154">
|
||||
<emphasis id="4386EAD3FFB6951BFF19F99FD3505D2C" bold="true" box="[199,316,1556,1581]" pageId="3" pageNumber="154">Abdomen</emphasis>
|
||||
with 5 visible ventrites; lacking internal calli; visible ventrite I lacking coxal line.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB6951BFF19F9BCD3EB5DBC" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="154">
|
||||
<emphasis id="4386EAD3FFB6951BFF19F9BCD3145D51" bold="true" box="[199,376,1591,1616]" pageId="3" pageNumber="154">Male genitalia</emphasis>
|
||||
with median lobe and tegmen laterally compressed or not; held at rest upside down in abdomen; penile struts not fused, separate entire length, 1.15–2.41× longer than median lobe; struts not coiled; internal sac when fully retracted <length of penile struts, flagellum absent or minute and not visible; spiculum gastrale symmetrical.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB6951BFF19F94CD19A5C29" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="154">
|
||||
<emphasis id="4386EAD3FFB6951BFF19F94CD3E05DE1" bold="true" box="[199,396,1735,1760]" pageId="3" pageNumber="154">Female genitalia</emphasis>
|
||||
with gonocoxites triangular, gradually narrowing posteriorly; apices of gonocoxites laterally with concave impression and setae. Gonostylus inserted at the concave impression of the gonocoxite, with several short setae. Valvifers expanded posteriorly. Spermatheca elongate, C-shaped.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB6951BFF19F8BFD6305C94" pageId="3" pageNumber="154" type="etymology">
|
||||
<paragraph id="714D36C1FFB6951BFF19F8BFD6305C94" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="154">
|
||||
<emphasis id="4386EAD3FFB6951BFF19F8BFD3225C4C" bold="true" box="[199,334,1844,1869]" pageId="3" pageNumber="154">Etymology.</emphasis>
|
||||
The generic epithet reflect the genus name of the host. The name
|
||||
<taxonomicName id="B6F24D42FFB6951BFBE4F8BFD6AD5C4D" box="[1082,1217,1844,1868]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="3" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB6951BFBE4F8BFD6AD5C4D" box="[1082,1217,1844,1868]" italics="true" pageId="3" pageNumber="154">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
is formed from the words “kerato”, Greek for horn and referring to the pair of horn-shaped spines on the sporophylls of
|
||||
<taxonomicName id="B6F24D42FFB6951BFAD5F8D3D7F05C6E" box="[1291,1436,1880,1903]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="3" pageNumber="154" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFB6951BFAD5F8D3D7F05C6E" box="[1291,1436,1880,1903]" italics="true" pageId="3" pageNumber="154">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
cones in which this genus inhabits and “philia”, Greek for affection. Gender feminine.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB6951CFF19F82BD0C55AAD" lastPageId="4" lastPageNumber="155" pageId="3" pageNumber="154" type="discussion">
|
||||
<paragraph id="714D36C1FFB6951CFF19F82BD0C55AAD" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1436,151,429]" lastPageId="4" lastPageNumber="155" pageId="3" pageNumber="154">
|
||||
<emphasis id="4386EAD3FFB6951BFF19F82BD3575CB8" bold="true" box="[199,315,1952,1977]" pageId="3" pageNumber="154">Remarks.</emphasis>
|
||||
With the addition of
|
||||
<taxonomicName id="B6F24D42FFB6951BFDF1F82BD0DA5CB9" box="[559,694,1952,1976]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="3" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB6951BFDF1F82BD0DA5CB9" box="[559,694,1952,1976]" italics="true" pageId="3" pageNumber="154">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
there are now seven genera in
|
||||
<taxonomicName id="B6F24D42FFB6951BFBC7F82BD6B55CB8" authorityName="Crowson" authorityYear="1952" box="[1049,1241,1952,1977]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||||
. The relationship of
|
||||
<taxonomicName id="B6F24D42FFB6951BFF63F84FD3295CDD" box="[189,325,1988,2012]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="3" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB6951BFF63F84FD3295CDD" box="[189,325,1988,2012]" italics="true" pageId="3" pageNumber="154">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
to other pharaxonothines, particularly with the two other genera that inhabit cycad cones,
|
||||
<taxonomicName id="B6F24D42FFB1951CFF49FF13D34A5BB1" box="[151,294,152,176]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB1951CFF49FF13D34A5BB1" box="[151,294,152,176]" italics="true" pageId="4" pageNumber="155">Cycadophila</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="B6F24D42FFB1951CFE81FF13D3935BB1" authorityName="Reitter" authorityYear="1875" box="[351,511,152,176]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB1951CFE81FF13D3935BB1" box="[351,511,152,176]" italics="true" pageId="4" pageNumber="155">Pharaxonotha</emphasis>
|
||||
</taxonomicName>
|
||||
, has been explored with analysis of the 16S rRNA gene (
|
||||
<bibRefCitation id="15634B30FFB1951CFB4DFF1CD71C5BB1" author="Skelley, P. & Xu, G. & Tang, W. & Lindstrom, A. J. & Marler, T. & Khuraijam, J. S. & Singh, R. & Radha, P. & Rich, S." box="[1171,1392,151,176]" pageId="4" pageNumber="155" pagination="1 - 63" refId="ref19957" refString="Skelley, P., Xu, G., Tang, W., Lindstrom, A. J., Marler, T., Khuraijam, J. S., Singh, R., Radha, P. & Rich, S. (2017) Review of Cycadophila Xu, Tang & Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species. Zootaxa, 4267 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 4267.1.1" type="journal article" year="2017">
|
||||
Skelley
|
||||
<emphasis id="4386EAD3FFB1951CFB2FFF12D7475BB1" box="[1265,1323,152,176]" italics="true" pageId="4" pageNumber="155">et al.</emphasis>
|
||||
2017
|
||||
</bibRefCitation>
|
||||
). In that study samples D0257 and D0284 correspond with
|
||||
<taxonomicName id="B6F24D42FFB1951CFCCBFF37D1F05BD5" box="[789,924,188,212]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB1951CFCCBFF37D1F05BD5" box="[789,924,188,212]" italics="true" pageId="4" pageNumber="155">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
and the
|
||||
<taxonomicName id="B6F24D42FFB1951CFBDBFF37D6E15BD5" box="[1029,1165,188,212]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB1951CFBDBFF37D6E15BD5" box="[1029,1165,188,212]" italics="true" pageId="4" pageNumber="155">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
lineage is sister to that which includes
|
||||
<taxonomicName id="B6F24D42FFB1951CFE94FF6BD3B55BF9" box="[330,473,224,248]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB1951CFE94FF6BD3B55BF9" box="[330,473,224,248]" italics="true" pageId="4" pageNumber="155">Cycadophila</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="B6F24D42FFB1951CFDCDFF6BD0DF5BF9" authorityName="Reitter" authorityYear="1875" box="[531,691,224,248]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFB1951CFDCDFF6BD0DF5BF9" box="[531,691,224,248]" italics="true" pageId="4" pageNumber="155">Pharaxonotha</emphasis>
|
||||
</taxonomicName>
|
||||
. Morphological characters distinguishing
|
||||
<taxonomicName id="B6F24D42FFB1951CFB4DFF6BD7765BF9" box="[1171,1306,224,248]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB1951CFB4DFF6BD7765BF9" box="[1171,1306,224,248]" italics="true" pageId="4" pageNumber="155">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
from these other genera are discussed below.
|
||||
<taxonomicName id="B6F24D42FFB1951CFDFEFE8FD0CB5A1D" box="[544,679,260,284]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB1951CFDFEFE8FD0CB5A1D" box="[544,679,260,284]" italics="true" pageId="4" pageNumber="155">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
is divided into two subgenera, the nominal subgenus
|
||||
<taxonomicName id="B6F24D42FFB1951CFACAFE8FD7F75A1D" box="[1300,1435,260,284]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB1951CFACAFE8FD7F75A1D" box="[1300,1435,260,284]" italics="true" pageId="4" pageNumber="155">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
and the subgenus
|
||||
<taxonomicName id="B6F24D42FFB1951CFE83FEA3D3D25A41" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[349,446,296,320]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFB1951CFE83FEA3D3D25A41" box="[349,446,296,320]" italics="true" pageId="4" pageNumber="155">Vovidesa</emphasis>
|
||||
</taxonomicName>
|
||||
and a total of seven species are recognized herein. All specimens of
|
||||
<taxonomicName id="B6F24D42FFB1951CFB63FEA3D7285A41" box="[1213,1348,296,320]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB1951CFB63FEA3D7285A41" box="[1213,1348,296,320]" italics="true" pageId="4" pageNumber="155">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
studied herein were collected from male cones of the New World cycad,
|
||||
<taxonomicName id="B6F24D42FFB1951CFCA9FEC7D6645A62" box="[887,1032,332,355]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFB1951CFCA9FEC7D6645A62" box="[887,1032,332,355]" italics="true" pageId="4" pageNumber="155">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
spp. Adults of two species and two larval morphotypes are typically found within one cone. The two larval morphotypes have been assigned to each subgenus and will be described in a later paper.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
369
data/F9/5B/87/F95B87D7FFBB9509FF49FB97D0C45A65.xml
Normal file
369
data/F9/5B/87/F95B87D7FFBB9509FF49FB97D0C45A65.xml
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<mods:title id="E1125611B5D2A98355881B34DD670A1F">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<mods:namePart id="E11FC93190E6408E5DE429AA61698779">Pérez-Farrera, Miguel Angel</mods:namePart>
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<heading id="2A0581ADFFBB9516FF49FB97D6155F37" bold="true" box="[151,1145,1052,1078]" fontSize="11" level="1" pageId="14" pageNumber="165" reason="1">
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<emphasis id="4386EAD3FFBB9516FF49FB97D6155F37" bold="true" box="[151,1145,1052,1078]" pageId="14" pageNumber="165">
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<taxonomicName id="B6F24D42FFBB9516FF49FB97D1B05F37" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,988,1052,1078]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFBB9516FF49FB97D34B5F37" bold="true" box="[151,295,1052,1078]" italics="true" pageId="14" pageNumber="165">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFBB9516FEE9FB97D3AA5F37" bold="true" box="[311,454,1052,1078]" italics="true" pageId="14" pageNumber="165">Ceratophila</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFBB9516FE09FB97D03F5F37" bold="true" box="[471,595,1052,1078]" italics="true" pageId="14" pageNumber="165">sanchezae</emphasis>
|
||||
Tang, Skelley & Pérez-Farrera
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFBB9516FC37FB97D6155F37" box="[1001,1145,1052,1078]" pageId="14" pageNumber="165" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBB9516FF49FBB5D0AB5F56" box="[151,711,1086,1111]" pageId="14" pageNumber="165" type="description">
|
||||
<paragraph id="714D36C1FFBB9516FF49FBB5D0AB5F56" blockId="14.[151,1145,1052,1111]" box="[151,711,1086,1111]" pageId="14" pageNumber="165">
|
||||
(
|
||||
<figureCitation id="E9C92A44FFBB9516FF41FBB5D36E5F56" box="[159,258,1086,1111]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="14" pageNumber="165">Figs. 1D</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFBB9516FED0FBB5D3775F56" box="[270,283,1086,1111]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="14" pageNumber="165">5</figureCitation>
|
||||
A–G; illustration 1E in
|
||||
<bibRefCitation id="15634B30FFBB9516FDFBFBB5D0D35F56" author="Vovides, A. P." box="[549,703,1086,1111]" pageId="14" pageNumber="165" pagination="102 - 104" refId="ref20613" refString="Vovides, A. P. (1991) Insect symbionts of some Mexican cycads in their natural habitat. Biotropica, 23, 102 - 104. https: // doi. org / 10.2307 / 2388697" type="journal article" year="1991">Vovides 1991</bibRefCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBB9516FF49FB0DD3155E2F" pageId="14" pageNumber="165" type="diagnosis">
|
||||
<paragraph id="714D36C1FFBB9516FF49FB0DD3155E2F" blockId="14.[151,1437,1158,2011]" pageId="14" pageNumber="165">
|
||||
<emphasis id="4386EAD3FFBB9516FF49FB0DD3345F9E" bold="true" box="[151,344,1158,1183]" pageId="14" pageNumber="165">Adult diagnosis.</emphasis>
|
||||
Distinguished from other
|
||||
<taxonomicName id="B6F24D42FFBB9516FD5EFB0DD1C45F9E" box="[640,936,1158,1183]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFBB9516FD5EFB0DD16B5F9F" box="[640,775,1158,1182]" italics="true" pageId="14" pageNumber="165">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFBB9516FCC9FB0DD1F25F9F" box="[791,926,1158,1182]" italics="true" pageId="14" pageNumber="165">Ceratophila</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
species by an unusually quadrate pronotum; dark maculation on elytra, when present, confined to suture and lateral margins, maculation well separated and not merging; male genitalia with length of penile strut relative to median lobe ~ 2:1; setae at apex of parameres with greatest length <dorsal width of parameres; and with a known distribution in
|
||||
<collectingCountry id="09E57651FFBB9516FBE1FB79D6FB5E0A" box="[1087,1175,1266,1291]" name="Mexico" pageId="14" pageNumber="165">Mexico</collectingCountry>
|
||||
, state of
|
||||
<collectingRegion id="B336F823FFBB9516FAD4FB78D71C5E0B" box="[1290,1392,1267,1290]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="14" pageNumber="165">Veracruz</collectingRegion>
|
||||
, on
|
||||
<emphasis id="4386EAD3FFBB9516FF49FA9CD3155E2F" box="[151,377,1303,1326]" italics="true" pageId="14" pageNumber="165">
|
||||
<taxonomicName id="B6F24D42FFBB9516FF49FA9CD3185E2F" box="[151,372,1303,1326]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="tenuis">Ceratozamia tenuis</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBB9517FF19FAB1D3B85EE4" lastPageId="15" lastPageNumber="166" pageId="14" pageNumber="165" type="description">
|
||||
<paragraph id="714D36C1FFBB9516FF19FAB1D6605D2B" blockId="14.[151,1437,1158,2011]" pageId="14" pageNumber="165">
|
||||
<emphasis id="4386EAD3FFBB9516FF19FAB1D38C5E52" bold="true" box="[199,480,1338,1363]" pageId="14" pageNumber="165">Adult male description.</emphasis>
|
||||
Length
|
||||
<quantity id="B60A9B24FFBB9516FDE0FAB1D08F5E53" box="[574,739,1338,1363]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.65" metricValueMin="3.25" pageId="14" pageNumber="165" unit="mm" value="3.95" valueMax="4.65" valueMin="3.25">3.25–4.65 mm</quantity>
|
||||
, width
|
||||
<quantity id="B60A9B24FFBB9516FCE8FAB0D1B65E53" box="[822,986,1339,1363]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" metricValueMax="1.91" metricValueMin="1.33" pageId="14" pageNumber="165" unit="mm" value="1.62" valueMax="1.91" valueMin="1.33">1.33–1.91 mm</quantity>
|
||||
(n = 10). Body in dorsal view elongateoval, greatest width at middle of elytra; in lateral view convex dorsally. General body color brown (
|
||||
<figureCitation id="E9C92A44FFBB9516FAD4FAD5D7345E76" box="[1290,1368,1374,1399]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="14" pageNumber="165">Figs. 5</figureCitation>
|
||||
A–E), head and pronotum concolorous and darker than base of elytra; central portion of elytra usually with a narrow black to dark brown coloration fading gradually to light brown at base and apex, lateral portion of elytra with more prominent dark coloration; dorsal surface punctate, shining and appearing glabrous, short procumbent hairs associated with punctation along lateral sides and declivity of elytra, ventrally shining and appearing glabrous except mesoventrite and abdomen mostly covered with long procumbent setae.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBB9516FF19F9BDD1B85CDA" blockId="14.[151,1437,1158,2011]" pageId="14" pageNumber="165">
|
||||
<emphasis id="4386EAD3FFBB9516FF19F9BDD36B5D4E" bold="true" box="[199,263,1590,1615]" pageId="14" pageNumber="165">Head</emphasis>
|
||||
not broad, width = 0.66–0.68× pronotal width; in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured (
|
||||
<figureCitation id="E9C92A44FFBB9516FCAFF9D1D66C5D72" box="[881,1024,1626,1651]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="14" pageNumber="165">Figs. 5A, E</figureCitation>
|
||||
), average distance between closest punctures 2× width of puncture; width
|
||||
<quantity id="B60A9B24FFBB9516FD96F9F4D0825D97" box="[584,750,1663,1687]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.2" metricValueMin="7.8" pageId="14" pageNumber="165" unit="mm" value="0.85" valueMax="0.92" valueMin="0.78">0.78–0.92 mm</quantity>
|
||||
; dorsal interocular distance
|
||||
<quantity id="B60A9B24FFBB9516FBFBF9F5D6A65D97" box="[1061,1226,1662,1687]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.8" metricValueMin="4.5" pageId="14" pageNumber="165" unit="mm" value="0.515" valueMax="0.58" valueMin="0.45">0.45–0.58 mm</quantity>
|
||||
, head width/dorsal interocular distance ratio 1.52–1.63, ventral interocular distance
|
||||
<quantity id="B60A9B24FFBB9516FC56F928D65E5DBB" box="[904,1074,1699,1723]" metricMagnitude="-4" metricUnit="m" metricValue="3.35" metricValueMax="3.9" metricValueMin="2.8" pageId="14" pageNumber="165" unit="mm" value="0.335" valueMax="0.39" valueMin="0.28">0.28–0.39 mm</quantity>
|
||||
, head width/ventral interocular distance ratio 2.38–2.78. Eye with large black facets. Antennal length slightly shorter than pronotal width, 1.2× head width; antennomere I (scape) fairly large, slightly elongate; antennomere II slightly smaller than III; IV–VII small, width equals length; VIII width 2× more than length; club fairly large, IX and similar in length, XI slightly longer, globular with angulate apex. Clypeus truncate anteriorly, moderately punctate; somewhat emarginate, with narrow margin. Mentum and submentum with moderate punctation, distance between nearest punctures approximately 1× own diameter, each puncture with a short seta. Gular area smooth, without punctation or setae, border with submentum usually with a shallow depression without punctation, but usually with a shallowly impressed circular or semicircular border along anterior margin (
|
||||
<figureCitation id="E9C92A44FFBB9516FCB3F849D1AB5CDA" box="[877,967,1986,2011]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="14" pageNumber="165">Fig. 5D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBA9517FF19FF1CD6AE5881" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="166">
|
||||
<emphasis id="4386EAD3FFBA9517FF19FF1CD34C5BB1" bold="true" box="[199,288,151,176]" pageId="15" pageNumber="166">Thorax</emphasis>
|
||||
with pronotum quadrate in dorsal view, length/width ratio 0.80–0.84; with marginal beads laterally and basally, anterior margin lacking distinct bead medially; convex; anterior and posterior angles weakly developed; basal ¼ expanding to parallel lateral carinae, then gradually converging in apical ½; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/7 length of posterior margin from posterior corners and touching and just anterior to marginal bead, longitudinal furrows lacking. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae approximately ½ length of eye. Hypomeron laterally with few minute punctures, medially with longitudinal striations. Elytra in dorsal view elongate-oval, convex; length/width 1.56– 1.66, greatest width near midlength; without marginal line basally; 10 complete striae of moderate puncture size; scutellary striole extending ¼ elytral length, with 10–15 punctures; intervals of striae with fine, shallow punctures. All punctures of elytral striae bearing a single short seta; seta only visible in profile, extending slightly out of puncture. Mesoventrite with moderate to strong punctation, distance between nearest punctures approximately equal to diameter of punctures, puncture depth moderate. Metaventrite dulled laterally, with strong lateral punctation separated by 2–3× own diameter; medial surface smooth, glossy, finely punctured, separated by 5–6× own diameter; convex laterally. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over half metaventrite length. Legs stout, relatively similar in length and shape. Procoxa oval; mesocoxa globular; metacoxa transversely elongate-oval; trochanters obliquely truncate apically; femora robust, moderately compressed laterally; tibiae shorter than femora, gradually dilated to obliquely truncate apices; protibia with disto-lateral tooth (
|
||||
<figureCitation id="E9C92A44FFBA9517FD87FC94D0C95839" box="[601,677,799,824]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="15" pageNumber="166">Figs. 5</figureCitation>
|
||||
A–C), with apical fringe of short, stout spinules of posterior margin lacking along concave medial margin (row broken) and long stout setae on anterior margin; meso- and metatibia with apical fringe of short, stout spinules on anterior margin, longer setae on posterior margins.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBA9517FF19FC07D71E5EC1" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="166">
|
||||
<emphasis id="4386EAD3FFBA9517FF19FC07D32F58A4" bold="true" box="[199,323,908,933]" pageId="15" pageNumber="166">Abdomen.</emphasis>
|
||||
Ventrite I with intercoxal process narrow, with triangular point anteromedially; lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; all ventrites bearing moderate, shallow punctation, distance to nearest puncture approximately 1× diameter of puncture, punctures bearing mostly reclining setae; ventrite V with setae length nearly uniformly approximately 2× diameter of puncture; ventrites I–IV with punctation and setae confined to median band, setae length approximately 1× diameter of puncture along anterior edge of band increasing to approximately 3× diameter along posterior edge; I–IV each with 2 or more median pairs of longer, semi-erect sensory hairs (often abraded), V with 2–3 pairs of long semi-erect hairs located postero-laterally. Male genital capsule fringed with fine setae. Male genitalia (tegmen and median lobe) oriented upside down (displayed right side up in
|
||||
<figureCitation id="E9C92A44FFBA9517FB71FB7FD7645E0C" box="[1199,1288,1268,1293]" captionStart="FIGURE 5" captionStartId="16.[151,250,1753,1775]" captionTargetBox="[151,1435,193,1732]" captionTargetId="figure@16.[151,1435,193,1732]" captionTargetPageId="16" captionText="FIGURE 5. Photographs of Ceratophila (Ceratophila) sanchezae (A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view." httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="15" pageNumber="166">Fig. 5F</figureCitation>
|
||||
to facilitate comparisons). Tegmen sclerotized, triangular, laterally compressed; with anterior region ring-like, posterior region sheath-like; lateral margins gradually converging posteriorly; apically with 2 elongate, slightly flattened parameres. Parameres in dorso-ventral view length/width ratio = 3.14–3.20; apically with setae, length of longest setae <width of paramere in dorso-ventral view. Aedeagus with median lobe laterally compressed, sclerotized, in lateral view strongly curved, apex tapering to rounded point. Ratio of penile strut length to median lobe length 1.90 (n = 1).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBA9517FF19FA47D3B85EE4" blockId="15.[151,1437,151,2013]" box="[199,468,1484,1509]" pageId="15" pageNumber="166">
|
||||
<emphasis id="4386EAD3FFBA9517FF19FA47D3725EE4" bold="true" box="[199,286,1484,1509]" pageId="15" pageNumber="166">Female</emphasis>
|
||||
similar to male.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBA9517FF19FA64D2835C71" pageId="15" pageNumber="166" type="materials_examined">
|
||||
<paragraph id="714D36C1FFBA9517FF19FA64D1455D09" blockId="15.[151,1437,151,2013]" box="[199,809,1519,1544]" pageId="15" pageNumber="166">
|
||||
<materialsCitation id="C19A3C9CFFBA9517FF19FA64D14A5D09" ID-GBIF-Occurrence="2028138745" box="[199,806,1519,1544]" country="Mexico" location="Cerro Coacoatzintla" pageId="15" pageNumber="166" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
|
||||
<emphasis id="4386EAD3FFBA9517FF19FA64D3095D09" bold="true" box="[199,357,1519,1544]" pageId="15" pageNumber="166">
|
||||
<typeStatus id="AE498863FFBA9517FF19FA64D36E5D09" box="[199,258,1519,1544]" pageId="15" pageNumber="166">Type</typeStatus>
|
||||
locality.
|
||||
</emphasis>
|
||||
<collectingCountry id="09E57651FFBA9517FEB2FA64D3A85D09" box="[364,452,1519,1544]" name="Mexico" pageId="15" pageNumber="166">Mexico</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFBA9517FE11FA7AD05A5D09" box="[463,566,1521,1544]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="15" pageNumber="166">Veracruz</collectingRegion>
|
||||
,
|
||||
<location id="742D601AFFBA9517FD9FFA7BD14A5D09" LSID="urn:lsid:plazi:treatment:F95B87D7FFBB9509FF49FB97D0C45A65:742D601AFFBA9517FD9FFA7BD14A5D09" box="[577,806,1519,1544]" country="Mexico" name="Cerro Coacoatzintla" pageId="15" pageNumber="166" stateProvince="Veracruz de Ignacio de la Llave">Cerro Coacoatzintla</location>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBA9517FF19F99FD1A55D51" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="166">
|
||||
<emphasis id="4386EAD3FFBA9517FF19F99FD3765D2C" bold="true" box="[199,282,1556,1581]" pageId="15" pageNumber="166">Range.</emphasis>
|
||||
As currently understood, this species is known to inhabit
|
||||
<taxonomicName id="B6F24D42FFBA9517FC15F99FD2985D51" authority="(Dyer) D. W. Stev. & Vovides" authorityName="D. W. Stev. & Vovides" baseAuthorityName="Dyer" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFBA9517FC15F99FD6C65D2D" box="[971,1194,1556,1580]" italics="true" pageId="15" pageNumber="166">Ceratozamia tenuis</emphasis>
|
||||
(Dyer) D.W.Stev. & Vovides
|
||||
</taxonomicName>
|
||||
, which occurs in the vicinity north of Xalapa,
|
||||
<collectingRegion id="B336F823FFBA9517FD24F9B2D10D5D51" box="[762,865,1593,1616]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="15" pageNumber="166">Veracruz</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="09E57651FFBA9517FCB2F9BCD1A95D51" box="[876,965,1591,1616]" name="Mexico" pageId="15" pageNumber="166">Mexico</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBA9517FF19F9D7D2835C71" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="166">
|
||||
<emphasis id="4386EAD3FFBA9517FF19F9D7D0545D74" bold="true" box="[199,568,1628,1653]" pageId="15" pageNumber="166">
|
||||
Material examined.
|
||||
<typeStatus id="AE498863FFBA9517FE15F9D7D0545D74" box="[459,568,1628,1653]" pageId="15" pageNumber="166" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
(by designation) male of
|
||||
<taxonomicName id="B6F24D42FFBA9517FC57F9D7D64B5D75" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[905,1063,1628,1652]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="sanchezae">
|
||||
<emphasis id="4386EAD3FFBA9517FC57F9D7D64B5D75" box="[905,1063,1628,1652]" italics="true" pageId="15" pageNumber="166">C. sanchezae</emphasis>
|
||||
</taxonomicName>
|
||||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||||
<collectingCountry id="09E57651FFBA9517FDA7F90BD0805D99" box="[633,748,1664,1688]" name="Mexico" pageId="15" pageNumber="166">MEXICO</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFBA9517FD27F90AD10C5D99" box="[761,864,1665,1688]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="15" pageNumber="166">Veracruz</collectingRegion>
|
||||
, Cerro Coacoatzintla,
|
||||
<specimenCount id="67F4FD48FFBA9517FB80F90AD61F5D99" box="[1118,1139,1665,1688]" pageId="15" pageNumber="166" type="male">♂</specimenCount>
|
||||
cone
|
||||
<taxonomicName id="B6F24D42FFBA9517FB67F90BD7F95D96" box="[1209,1429,1664,1687]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFBA9517FB67F90BD7F95D96" box="[1209,1429,1664,1687]" italics="true" pageId="15" pageNumber="166">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFBA9517FF49F92FD37F5DBC" box="[151,275,1700,1725]" pageId="15" pageNumber="166" value="1995-02-28">28-II-1995</date>
|
||||
, Y. Sanchez Tinoco”; 2) [rectangular; red; printed in black ink]
|
||||
<typeStatus id="AE498863FFBA9517FC3CF92FD61A5DBD" box="[994,1142,1700,1724]" pageId="15" pageNumber="166" type="holotype">HOLOTYPE</typeStatus>
|
||||
<specimenCount id="67F4FD48FFBA9517FBA0F92ED6FF5DBD" box="[1150,1171,1701,1724]" pageId="15" pageNumber="166" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFBA9517FB44F92FD0575DE1" authority="Tang, Skelley & Perez-Farrera 2018" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBA9517FB44F92FD7F05DBD" box="[1178,1436,1700,1724]" italics="true" pageId="15" pageNumber="166">Ceratophila sanchezae</emphasis>
|
||||
<bibRefCitation id="15634B30FFBA9517FF49F942D0575DE1" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." box="[151,571,1735,1761]" pageId="15" pageNumber="166" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">Tang, Skelley & Pérez-Farrera 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. Deposited in the FSCA.
|
||||
<typeStatus id="AE498863FFBA9517FCBBF94CD1A65DE1" box="[869,970,1735,1760]" pageId="15" pageNumber="166" type="allotype">
|
||||
<emphasis id="4386EAD3FFBA9517FCBBF94CD1A65DE1" bold="true" box="[869,970,1735,1760]" pageId="15" pageNumber="166">Allotype</emphasis>
|
||||
</typeStatus>
|
||||
(FSCA) same data as
|
||||
<typeStatus id="AE498863FFBA9517FB0AF94CD75A5DE1" box="[1236,1334,1735,1760]" pageId="15" pageNumber="166" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="67F4FD48FFBA9517FAACF943D2A35C04" pageId="15" pageNumber="166" type="adult">399 adult</specimenCount>
|
||||
<typeStatus id="AE498863FFBA9517FF07F967D3235C04" box="[217,335,1772,1797]" pageId="15" pageNumber="166" type="paratype">
|
||||
<emphasis id="4386EAD3FFBA9517FF07F967D3235C04" bold="true" box="[217,335,1772,1797]" pageId="15" pageNumber="166">paratypes</emphasis>
|
||||
</typeStatus>
|
||||
:
|
||||
<emphasis id="4386EAD3FFBA9517FEBEF967D03B5C04" bold="true" box="[352,599,1772,1797]" pageId="15" pageNumber="166">
|
||||
<collectingCountry id="09E57651FFBA9517FEBEF967D3B75C04" box="[352,475,1772,1797]" name="Mexico" pageId="15" pageNumber="166">MEXICO</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="B336F823FFBA9517FE34F967D03B5C04" box="[490,599,1772,1797]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="15" pageNumber="166">Veracruz</collectingRegion>
|
||||
</emphasis>
|
||||
, Jilotepec, El Esquilón,
|
||||
<quantity id="B60A9B24FFBA9517FCADF967D1A85C05" box="[883,964,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="166" unit="m" value="1300.0">1300m</quantity>
|
||||
, ex male cone of
|
||||
<taxonomicName id="B6F24D42FFBA9517FB48F967D7F05C05" box="[1174,1436,1772,1796]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="mexicana">
|
||||
<emphasis id="4386EAD3FFBA9517FB48F967D7F05C05" box="[1174,1436,1772,1796]" italics="true" pageId="15" pageNumber="166">Ceratozamia mexicana</emphasis>
|
||||
</taxonomicName>
|
||||
[now
|
||||
<taxonomicName id="B6F24D42FFBA9517FF09F89BD3565C26" box="[215,314,1808,1831]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFBA9517FF09F89BD3565C26" box="[215,314,1808,1831]" italics="true" pageId="15" pageNumber="166">C. tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
],
|
||||
<date id="054C1001FFBA9517FE8FF89BD3BA5C29" box="[337,470,1808,1832]" pageId="15" pageNumber="166" value="1989-03-10">10-III-1989</date>
|
||||
, A. P. Vovides, (133); Cerro Coacoatzintla,
|
||||
<specimenCount id="67F4FD48FFBA9517FC60F89AD1BF5C29" box="[958,979,1809,1832]" pageId="15" pageNumber="166" type="male">♂</specimenCount>
|
||||
cone
|
||||
<taxonomicName id="B6F24D42FFBA9517FBCBF89BD69C5C26" box="[1045,1264,1808,1831]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFBA9517FBCBF89BD69C5C26" box="[1045,1264,1808,1831]" italics="true" pageId="15" pageNumber="166">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFBA9517FB23F89BD7205C29" box="[1277,1356,1808,1832]" pageId="15" pageNumber="166" value="1928-02-01">28-II-1</date>
|
||||
995, Y. Sanchez Tinoco (266).
|
||||
<typeStatus id="AE498863FFBA9517FE76F8BED0795C4C" box="[424,533,1845,1869]" pageId="15" pageNumber="166" type="paratype">Paratypes</typeStatus>
|
||||
deposited at ANIC, BMNH, FSCA, IEXA, INBio, MNHN, NZAC, UNAM, USNM.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBA9517FF19F8F7D64E5CDC" pageId="15" pageNumber="166" type="etymology">
|
||||
<paragraph id="714D36C1FFBA9517FF19F8F7D64E5CDC" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="166">
|
||||
<emphasis id="4386EAD3FFBA9517FF19F8F7D3225C94" bold="true" box="[199,334,1916,1941]" pageId="15" pageNumber="166">Etymology.</emphasis>
|
||||
The species is named in honor of Maria Ydelia Sánchez Tinoco, who collected most of the
|
||||
<typeStatus id="AE498863FFBA9517FAB2F8F6D7F05C94" box="[1388,1436,1917,1941]" pageId="15" pageNumber="166">type</typeStatus>
|
||||
series of this species, as well as other species studied in this paper, and for her contributions to the reproductive biology of
|
||||
<taxonomicName id="B6F24D42FFBA9517FECCF84FD0835CDD" authority="(Sanchez-Tinoco et al. 2000)" authorityName="Sanchez-Tinoco" authorityYear="2000" baseAuthorityName="Sanchez-Tinoco" baseAuthorityYear="2000" box="[274,751,1988,2013]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="166" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBA9517FECCF84FD3CF5CDA" box="[274,419,1988,2011]" italics="true" pageId="15" pageNumber="166">Ceratozamia</emphasis>
|
||||
(
|
||||
<bibRefCitation id="15634B30FFBA9517FE6FF84FD08B5CDD" author="Sanchez-Tinoco, M. Y. & Engleman, E. M. & Vovides, A. P." box="[433,743,1988,2013]" pageId="15" pageNumber="166" pagination="15 - 23" refId="ref19911" refString="Sanchez-Tinoco, M. Y., Engleman, E. M. & Vovides, A. P. (2000) Cronologia reproductora de Ceratozamia mexicana (Cycadales). Boletin de la Sociedad Botanica de Mexico, 66, 15 - 23." type="journal article" year="2000">
|
||||
Sánchez-Tinoco
|
||||
<emphasis id="4386EAD3FFBA9517FDB0F84ED0CA5CDD" box="[622,678,1988,2012]" italics="true" pageId="15" pageNumber="166">et al.</emphasis>
|
||||
2000
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and other Mexican cycads.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="258D6649FFA59508FF49F952D3D25C2C" httpUri="https://zenodo.org/record/2606946/files/figure.png" pageId="16" pageNumber="167" startId="16.[151,250,1753,1775]" targetBox="[151,1435,193,1732]" targetPageId="16">
|
||||
<paragraph id="714D36C1FFA59508FF49F952D3D25C2C" blockId="16.[151,1436,1753,1837]" pageId="16" pageNumber="167">
|
||||
<emphasis id="4386EAD3FFA59508FF49F952D37E5DEE" bold="true" box="[151,274,1753,1775]" pageId="16" pageNumber="167">FIGURE 5.</emphasis>
|
||||
Photographs of
|
||||
<taxonomicName id="B6F24D42FFA59508FE67F952D1585DEE" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[441,820,1753,1775]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="167" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFA59508FE67F952D0585DEE" box="[441,564,1753,1775]" italics="true" pageId="16" pageNumber="167">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFA59508FD9DF952D0D15DEE" box="[579,701,1753,1775]" italics="true" pageId="16" pageNumber="167">Ceratophila</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFA59508FD12F952D1585DEE" box="[716,820,1753,1775]" italics="true" pageId="16" pageNumber="167">sanchezae</emphasis>
|
||||
</taxonomicName>
|
||||
(A–E: female allotype): A–C) dorsal, ventral and lateral and habitus; D–E) head, ventral and lateral views; F) median lobe still attached with tegmen, ventral and lateral views; G) male genital capsule, ventral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="39E8654AFFA59509FF19F8D2D0C45A65" lastPageId="17" lastPageNumber="168" pageId="16" pageNumber="167" type="discussion">
|
||||
<paragraph id="714D36C1FFA59509FF19F8D2D0C45A65" blockId="16.[151,1436,1881,2014]" lastBlockId="17.[151,1436,151,357]" lastPageId="17" lastPageNumber="168" pageId="16" pageNumber="167">
|
||||
<emphasis id="4386EAD3FFA59508FF19F8D2D3575C73" bold="true" box="[199,315,1881,1906]" pageId="16" pageNumber="167">Remarks.</emphasis>
|
||||
<typeStatus id="AE498863FFA59508FE98F8D1D3125C73" box="[326,382,1882,1906]" pageId="16" pageNumber="167">Type</typeStatus>
|
||||
specimens of this species are limited to those collected on
|
||||
<taxonomicName id="B6F24D42FFA59508FB9CF8D1D74D5C70" box="[1090,1313,1882,1905]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="167" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFA59508FB9CF8D1D74D5C70" box="[1090,1313,1882,1905]" italics="true" pageId="16" pageNumber="167">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
, a species recently described by
|
||||
<bibRefCitation id="15634B30FFA59508FE50F8F6D0105C97" author="Vovides, A. P. & Stevenson, D. W. & Perez-Farrera, M. A. & Lopez, S. & Avendano, S." box="[398,636,1917,1942]" pageId="16" pageNumber="167" pagination="419 - 429" refId="ref20651" refString="Vovides, A. P., Stevenson, D. W., Perez-Farrera, M. A., Lopez, S. & Avendano, S. (2016) What is Ceratozamia mexicana (Zamiaceae)? Botanical Sciences, 94, 419 - 429. https: // doi. org / 10.17129 / botsci. 449" type="journal article" year="2016">
|
||||
Vovides
|
||||
<emphasis id="4386EAD3FFA59508FE31F8F5D04B5C94" box="[495,551,1917,1941]" italics="true" pageId="16" pageNumber="167">et al.</emphasis>
|
||||
(2016)
|
||||
</bibRefCitation>
|
||||
. Prior to 2016 this host had been erroneously identified as
|
||||
<taxonomicName id="B6F24D42FFA59508FACEF8F5D2995CBB" authority="Brongn." authorityName="Brongn." class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="167" phylum="Tracheophyta" rank="species" species="mexicana">
|
||||
<emphasis id="4386EAD3FFA59508FACEF8F5D7F05C94" box="[1296,1436,1918,1941]" italics="true" pageId="16" pageNumber="167">C. mexicana</emphasis>
|
||||
Brongn.
|
||||
</taxonomicName>
|
||||
, a species now understood to occur only in the vicinity of El Mirador, further south in
|
||||
<collectingRegion id="B336F823FFA59508FB0CF829D7555CB8" box="[1234,1337,1954,1977]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="16" pageNumber="167">Veracruz</collectingRegion>
|
||||
. Besides
|
||||
<taxonomicName id="B6F24D42FFA59508FF49F84DD3145CDC" box="[151,376,1990,2013]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="167" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFA59508FF49F84DD3145CDC" box="[151,376,1990,2013]" italics="true" pageId="16" pageNumber="167">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
, several other species of
|
||||
<taxonomicName id="B6F24D42FFA59508FD68F84DD12B5CDC" box="[694,839,1990,2013]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="167" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA59508FD68F84DD12B5CDC" box="[694,839,1990,2013]" italics="true" pageId="16" pageNumber="167">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
occur in the rugged terrain of western
|
||||
<collectingRegion id="B336F823FFA59508FAEFF84DD7F45CDC" box="[1329,1432,1990,2013]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="16" pageNumber="167">Veracruz</collectingRegion>
|
||||
, including
|
||||
<taxonomicName id="B6F24D42FFA49509FED4FF13D3B35BB1" authority="Miq." authorityName="Miq." box="[266,479,151,176]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="brevifrons">
|
||||
<emphasis id="4386EAD3FFA49509FED4FF13D3F15BB1" box="[266,413,152,176]" italics="true" pageId="17" pageNumber="168">C. brevifrons</emphasis>
|
||||
Miq.
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFA49509FE32FF13D6375BB1" authority="Vazq. Torres, A. Moretti & Carvajal-Hern." authorityName="Vazq. Torres, A. Moretti & Carvajal-Hern." box="[492,1115,151,176]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="delucana">
|
||||
<emphasis id="4386EAD3FFA49509FE32FF13D01A5BB1" box="[492,630,152,176]" italics="true" pageId="17" pageNumber="168">C. delucana</emphasis>
|
||||
Vázq.Torres, A.Moretti & Carvajal-Hern.
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFA49509FBB9FF13D3215BD5" authority="Avendano, Vovides & Cast." authorityName="Avendano, Vovides & Cast." class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="haustecorum">
|
||||
<emphasis id="4386EAD3FFA49509FBB9FF13D7765BB1" box="[1127,1306,152,176]" italics="true" pageId="17" pageNumber="168">C. haustecorum</emphasis>
|
||||
Avendaño, Vovides & Cast.
|
||||
</taxonomicName>
|
||||
-Campos,
|
||||
<taxonomicName id="B6F24D42FFA49509FE1EFF37D0275BD5" box="[448,587,188,212]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="mexicana">
|
||||
<emphasis id="4386EAD3FFA49509FE1EFF37D0275BD5" box="[448,587,188,212]" italics="true" pageId="17" pageNumber="168">C. mexicana</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="B6F24D42FFA49509FD5CFF37D6D45BD4" authority="Vazq. Torres & Vovides. The" authorityName="Vazq. Torres & Vovides. The" box="[642,1208,188,213]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="morettii">
|
||||
<emphasis id="4386EAD3FFA49509FD5CFF37D11C5BD5" box="[642,880,188,212]" italics="true" pageId="17" pageNumber="168">Ceratozamia morettii</emphasis>
|
||||
Vázq.Torres & Vovides. The
|
||||
</taxonomicName>
|
||||
possible occurrence of
|
||||
<taxonomicName id="B6F24D42FFA49509FF63FF6BD3AB5BF9" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[189,455,224,248]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFA49509FF63FF6BD3AB5BF9" box="[189,455,224,248]" italics="true" pageId="17" pageNumber="168">Ceratophila sanchezae</emphasis>
|
||||
</taxonomicName>
|
||||
in these species is currently being investigated by the authors. In one cone of
|
||||
<taxonomicName id="B6F24D42FFA49509FF49FE8FD3145A1D" box="[151,376,260,284]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="tenuis">
|
||||
<emphasis id="4386EAD3FFA49509FF49FE8FD3145A1D" box="[151,376,260,284]" italics="true" pageId="17" pageNumber="168">Ceratozamia tenuis</emphasis>
|
||||
</taxonomicName>
|
||||
sampled
|
||||
<taxonomicName id="B6F24D42FFA49509FE2FFE8FD0155A1D" box="[497,633,260,284]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFA49509FE2FFE8FD0155A1D" box="[497,633,260,284]" italics="true" pageId="17" pageNumber="168">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicName id="B6F24D42FFA49509FD58FE8FD15F5A1D" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[646,819,260,284]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="sanchezae" subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFA49509FD51FE8FD0CE5A1A" box="[655,674,260,283]" italics="true" pageId="17" pageNumber="168">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFA49509FD1EFE8FD15F5A1D" box="[704,819,260,284]" italics="true" pageId="17" pageNumber="168">sanchezae</emphasis>
|
||||
</taxonomicName>
|
||||
accounted for 22% of 1079 adult
|
||||
<taxonomicName id="B6F24D42FFA49509FB3EFE8FD7F05A1C" authorityName="Crowson" authorityYear="1952" box="[1248,1436,260,285]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||||
collected, while
|
||||
<taxonomicName id="B6F24D42FFA49509FE8EFEA3D0235A3E" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[336,591,296,320]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="17" pageNumber="168" phylum="Tracheophyta" rank="species" species="vazquezi" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFA49509FE8EFEA3D30F5A3E" box="[336,355,296,319]" italics="true" pageId="17" pageNumber="168">C</emphasis>
|
||||
. (
|
||||
<emphasis id="4386EAD3FFA49509FEA4FEA3D3B75A41" box="[378,475,296,320]" italics="true" pageId="17" pageNumber="168">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFA49509FE31FEA3D0235A3E" box="[495,591,296,319]" italics="true" pageId="17" pageNumber="168">vazquezi</emphasis>
|
||||
</taxonomicName>
|
||||
accounted for 7%. An undescribed species of
|
||||
<taxonomicName id="B6F24D42FFA49509FB85FEA3D6975A41" authorityName="Reitter" authorityYear="1875" box="[1115,1275,296,320]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFA49509FB85FEA3D6975A41" box="[1115,1275,296,320]" italics="true" pageId="17" pageNumber="168">Pharaxonotha</emphasis>
|
||||
</taxonomicName>
|
||||
accounted for the remaining 71% of adult beetles in the cone.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
365
data/F9/5B/87/F95B87D7FFBD9513FF49F94CD2955F7C.xml
Normal file
365
data/F9/5B/87/F95B87D7FFBD9513FF49F94CD2955F7C.xml
Normal file
|
|
@ -0,0 +1,365 @@
|
|||
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|
||||
<subSubSection id="39E8654AFFBD9510FF49F94CD60E5DE3" box="[151,1122,1735,1762]" pageId="8" pageNumber="159" type="nomenclature">
|
||||
<paragraph id="714D36C1FFBD9510FF49F94CD60E5DE3" blockId="8.[151,1122,1735,1795]" box="[151,1122,1735,1762]" pageId="8" pageNumber="159">
|
||||
<heading id="2A0581ADFFBD9510FF49F94CD60E5DE3" bold="true" box="[151,1122,1735,1762]" fontSize="11" level="1" pageId="8" pageNumber="159" reason="1">
|
||||
<emphasis id="4386EAD3FFBD9510FF49F94CD60E5DE3" bold="true" box="[151,1122,1735,1762]" pageId="8" pageNumber="159">
|
||||
<taxonomicName id="B6F24D42FFBD9510FF49F94CD1A95DE3" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,965,1735,1762]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="159" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFBD9510FF49F94CD34B5DE0" bold="true" box="[151,295,1735,1761]" italics="true" pageId="8" pageNumber="159">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFBD9510FEE9F94CD3AA5DE0" bold="true" box="[311,454,1735,1761]" italics="true" pageId="8" pageNumber="159">Ceratophila</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFBD9510FE09F94CD0505DE0" bold="true" box="[471,572,1735,1761]" italics="true" pageId="8" pageNumber="159">gregoryi</emphasis>
|
||||
Tang, Skelley & Pérez-Farrera
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFBD9510FC0CF943D60E5DE3" box="[978,1122,1736,1762]" pageId="8" pageNumber="159" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBD9510FF49F961D33A5C02" box="[151,342,1770,1795]" pageId="8" pageNumber="159" type="description">
|
||||
<paragraph id="714D36C1FFBD9510FF49F961D33A5C02" blockId="8.[151,1122,1735,1795]" box="[151,342,1770,1795]" pageId="8" pageNumber="159">
|
||||
(
|
||||
<figureCitation id="E9C92A44FFBD9510FF41F961D2905C02" box="[159,252,1770,1795]" captionStart="FIGURE 1" captionStartId="4.[151,250,1706,1728]" captionTargetBox="[151,1435,483,1681]" captionTargetId="figure@4.[151,1435,483,1681]" captionTargetPageId="4" captionText="FIGURE 1. Key characters of adults: base of head, pronotum and base of elytra, dorsal view: A) Pharaxonotha sp. collected from Ceratozamia tenuis; B) Ceratophila (C.) chemnicki; pronotal left lateral margins: C) Pharaxonotha sp. from Ceratozamia tenuis; D) Ceratophila (C.) sanchezae; E) Ceratophila (Uovidesa) chiapensis from Ceratozamia vovidesii; right metatibiae of male individuals, posterior view: F) Ceratophila (C.) gregoryi; G) C. (U.) mixeorum; b: basal bead of elytra; s: stridulatory files." httpUri="https://zenodo.org/record/2606936/files/figure.png" pageId="8" pageNumber="159">Figs. 1F</figureCitation>
|
||||
,
|
||||
<figureCitation id="E9C92A44FFBD9510FED6F960D3785C02" box="[264,276,1771,1795]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="8" pageNumber="159">3</figureCitation>
|
||||
A–H)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBD9510FF49F8B9D08B5CB7" pageId="8" pageNumber="159" type="diagnosis">
|
||||
<paragraph id="714D36C1FFBD9510FF49F8B9D08B5CB7" blockId="8.[151,1436,1842,1975]" pageId="8" pageNumber="159">
|
||||
<emphasis id="4386EAD3FFBD9510FF49F8B9D30D5C4A" bold="true" box="[151,353,1842,1867]" pageId="8" pageNumber="159">Adult diagnosis.</emphasis>
|
||||
Distinguished from other
|
||||
<taxonomicName id="B6F24D42FFBD9510FD72F8B8D1B15C4A" box="[684,989,1843,1867]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFBD9510FD72F8B8D1585C4A" box="[684,820,1843,1867]" italics="true" pageId="8" pageNumber="159">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFBD9510FC92F8B8D1BF5C4A" box="[844,979,1843,1867]" italics="true" pageId="8" pageNumber="159">Ceratophila</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
species by the pronotum narrowing anteriorly; dark marks on the elytra absent or faint; male genitalia with length of penile strut relative to median lobe ~ 3:2; setae at apex of parameres with greatest length ± dorsal width of parameres; and with a known distribution in
|
||||
<collectingCountry id="09E57651FFBD9510FF49F815D2835CB6" box="[151,239,1950,1975]" name="Mexico" pageId="8" pageNumber="159">Mexico</collectingCountry>
|
||||
, state of
|
||||
<collectingRegion id="B336F823FFBD9510FE8DF814D3C65CB6" box="[339,426,1951,1975]" country="Mexico" name="Oaxaca" pageId="8" pageNumber="159">Oaxaca</collectingRegion>
|
||||
, on
|
||||
<emphasis id="4386EAD3FFBD9510FE06F814D08B5CB7" box="[472,743,1951,1974]" italics="true" pageId="8" pageNumber="159">
|
||||
<taxonomicName id="B6F24D42FFBD9510FE06F814D08E5CB7" box="[472,738,1951,1974]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="8" pageNumber="159" phylum="Tracheophyta" rank="genus">Ceratozamia mixeorum</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="258D6649FFBC9511FF49F81BD6105CEB" httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="9" pageNumber="160" startId="9.[151,250,1936,1958]" targetBox="[203,1389,193,1919]" targetPageId="9">
|
||||
<paragraph id="714D36C1FFBC9511FF49F81BD6105CEB" blockId="9.[151,1436,1936,2026]" pageId="9" pageNumber="160">
|
||||
<emphasis id="4386EAD3FFBC9511FF49F81BD37B5CA7" bold="true" box="[151,279,1936,1958]" pageId="9" pageNumber="160">FIGURE 3.</emphasis>
|
||||
Photographs of
|
||||
<taxonomicName id="B6F24D42FFBC9511FE14F81BD1565CA7" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[458,826,1936,1958]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="9" pageNumber="160" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFBC9511FE14F81BD0295CA7" box="[458,581,1936,1958]" italics="true" pageId="9" pageNumber="160">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFBC9511FD89F81BD0BE5CA7" box="[599,722,1936,1958]" italics="true" pageId="9" pageNumber="160">Ceratophila</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFBC9511FD3AF81AD1565CA7" box="[740,826,1937,1958]" italics="true" pageId="9" pageNumber="160">gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
(B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="39E8654AFFBF9513FF10FF1CD3B85A1C" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161" type="description">
|
||||
<paragraph id="714D36C1FFBF9512FF10FF1CD6855A3E" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="161">
|
||||
<emphasis id="4386EAD3FFBF9512FF10FF1CD38B5BB1" bold="true" box="[206,487,151,176]" pageId="10" pageNumber="161">Adult male description.</emphasis>
|
||||
Length
|
||||
<quantity id="B60A9B24FFBF9512FD9AFF1CD08B5BAE" box="[580,743,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.57" metricValueMax="5.11" metricValueMin="4.03" pageId="10" pageNumber="161" unit="mm" value="4.57" valueMax="5.11" valueMin="4.03">4.03–5.11 mm</quantity>
|
||||
, width
|
||||
<quantity id="B60A9B24FFBF9512FCE7FF1CD1B05BAE" box="[825,988,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.815" metricValueMax="2.04" metricValueMin="1.59" pageId="10" pageNumber="161" unit="mm" value="1.815" valueMax="2.04" valueMin="1.59">1.59–2.04 mm</quantity>
|
||||
(n = 10). Body in dorsal view elongateoval, greatest width at middle of elytra; in lateral view convex dorsally. General body color red-brown (
|
||||
<figureCitation id="E9C92A44FFBF9512FAF2FF37D7145BD5" box="[1324,1400,188,213]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="10" pageNumber="161">Figs. 3</figureCitation>
|
||||
A– E), pronotum contrasting with light brown of elytra; dorsal surface punctate, shining and appearing glabrous, short procumbent hairs associated with punctation along lateral sides and declivity of elytra, ventrally shining and appearing glabrous except mesoventrite and abdomen mostly covered with long procumbent setae.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBF9512FF19FEC7D77459CD" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="161">
|
||||
<emphasis id="4386EAD3FFBF9512FF19FEC7D36B5A64" bold="true" box="[199,263,332,357]" pageId="10" pageNumber="161">Head</emphasis>
|
||||
not broad, width = 0.66–0.68× pronotal width; in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured (
|
||||
<figureCitation id="E9C92A44FFBF9512FCB1FEE4D66C5A89" box="[879,1024,367,392]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="10" pageNumber="161">Figs. 3A, D</figureCitation>
|
||||
), average distance between closest punctures 2× width of puncture; width
|
||||
<quantity id="B60A9B24FFBF9512FD96FE1FD0825AAD" box="[584,750,404,429]" metricMagnitude="-4" metricUnit="m" metricValue="9.55" metricValueMax="10.5" metricValueMin="8.6" pageId="10" pageNumber="161" unit="mm" value="0.955" valueMax="1.05" valueMin="0.86">0.86–1.05 mm</quantity>
|
||||
; dorsal interocular distance
|
||||
<quantity id="B60A9B24FFBF9512FBFBFE1FD6A65AAD" box="[1061,1226,404,429]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="6.3" metricValueMin="5.699999999999999" pageId="10" pageNumber="161" unit="mm" value="0.6" valueMax="0.63" valueMin="0.57">0.57–0.63 mm</quantity>
|
||||
, head width/dorsal interocular distance ratio 1.54–1.67, ventral interocular distance
|
||||
<quantity id="B60A9B24FFBF9512FC56FE33D65E5ACE" box="[904,1074,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="4.1" metricValueMin="3.4" pageId="10" pageNumber="161" unit="mm" value="0.375" valueMax="0.41" valueMin="0.34">0.34–0.41 mm</quantity>
|
||||
, head width/ventral interocular distance ratio 2.43–2.67. Eye with large black facets. Antennal length ¾ pronotal width, 1.2× head width, antennomere I (scape) fairly large, slightly elongate; antennomere II slightly smaller than III; IV–VII small, width equals length; VIII width slightly more than length; club fairly large, IX and X similar in length, XI slightly longer, globular with angulate apex. Clypeus truncate anteriorly, moderately punctate; somewhat emarginate, with narrow margin. Mentum and submentum with moderate punctation, distance between nearest punctures approximately 1× own diameter, each puncture with a short seta. Gular area smooth, without punctation or setae, border with submentum usually without punctation, with a shallowly impressed circular dimple medially (
|
||||
<figureCitation id="E9C92A44FFBF9512FB6AFD3FD76059CD" box="[1204,1292,692,717]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="10" pageNumber="161">Fig. 3E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBF9512FF19FD5CD01C5EE4" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="161">
|
||||
<emphasis id="4386EAD3FFBF9512FF19FD5CD34C59F1" bold="true" box="[199,288,727,752]" pageId="10" pageNumber="161">Thorax</emphasis>
|
||||
with pronotum somewhat quadrate in dorsal view narrowing anteriorly, length/width ratio 0.78–0.82; with marginal beads laterally and basally, anterior margin lacking distinct bead medially; convex; anterior angle weakly developed, posterior angles rounded; medial ½ of lateral carinae weakly converging anteriorly, then gradually converging in apical 1/4; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/7 length of posterior margin from posterior corners and touching and just anterior to marginal bead, longitudinal furrows lacking. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae approximately ½ length of eye. Hypomeron laterally with few minute punctures, medially with longitudinal striations. Elytra in dorsal view elongate-oval, convex; length/width 1.61–1.73, greatest width near midlength; without marginal line basally; 10 complete striae of moderate puncture size; scutellary striole extending ¼ elytral length, with 10–15 punctures; intervals of striae with fine, shallow punctures. All punctures of elytral striae bearing a single short seta; seta only visible in profile, extending slightly out of puncture. Mesoventrite with moderate to strong punctation, distance between nearest punctures approximately equal to diameter of punctures, puncture depth moderate. Metaventrite dulled laterally, with strong lateral punctation separated by 2–3× own diameter; medial surface smooth, glossy, finely punctured, punctation separated by 5–6× own diameter; convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over half metaventrite length. Legs stout, relatively similar in length and shape. Procoxa oval; mesocoxa globular; metacoxa transversely elongate-oval; trochanters obliquely truncate apically; femora robust, moderately compressed laterally; triangular tibiae shorter than femora, gradually dilated to obliquely truncate apices; protibia with disto-lateral tooth (
|
||||
<figureCitation id="E9C92A44FFBF9512FB15FAD4D7765E79" box="[1227,1306,1375,1400]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="10" pageNumber="161">Figs. 3</figureCitation>
|
||||
B–C), with apical fringe of short, stout spinules of posterior margin lacking along concave medial margin (row broken) and long stout setae on anterior margin; meso- and metatibia with apical fringe of short, stout spinules on anterior margin, longer setae on posterior margins.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBF9513FF19FA64D1415BF9" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1437,151,1149]" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161">
|
||||
<emphasis id="4386EAD3FFBF9512FF19FA64D32F5D09" bold="true" box="[199,323,1519,1544]" pageId="10" pageNumber="161">Abdomen.</emphasis>
|
||||
Ventrite I with intercoxal process narrow, with triangular point anteromedially; lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; all ventrites bearing moderate, shallow punctation, distance to nearest puncture approximately 1× diameter of puncture, punctures bearing mostly reclining setae; ventrite V with setae length nearly uniformly approximately 2× diameter of puncture; ventrites I–IV with punctation and setae confined to median band, setae length approximately 1× diameter of puncture along anterior edge of band increasing to approximately 3× diameter along posterior edge; I–IV each with 2 or more median pairs of longer, semi-erect sensory hairs (often abraded), V with 2–3 pairs of long semi-erect hairs located postero-laterally. Male genital capsule fringed with fine setae. Male genitalia (tegmen and median lobe) oriented upside down (displayed right side up in
|
||||
<figureCitation id="E9C92A44FFBF9512FB90F8DCD6CB5C71" box="[1102,1191,1879,1904]" captionStart="FIGURE 3" captionStartId="9.[151,250,1936,1958]" captionTargetBox="[203,1389,193,1919]" captionTargetId="figure@9.[203,1389,193,1919]" captionTargetPageId="9" captionText="FIGURE 3. Photographs of Ceratophila (Ceratophila) gregoryi (B–E: female allotype): A) dorsal habitus, female; B–C) ventral and lateral and habitus; D–E) head, dorsal and ventral views; F) male genital capsule, ventral view; G) median lobe still attached with tegmen, lateral and ventral views; H) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606940/files/figure.png" pageId="10" pageNumber="161">Fig. 3G</figureCitation>
|
||||
). Tegmen sclerotized, triangular, laterally compressed; with anterior region ring-like, posterior region sheath-like; lateral margins gradually converging posteriorly; apically with 2 elongate, somewhat bar-shaped parameres. Parameres dorsoventrally compressed, height approximately 1/2 its own width, in dorso-ventral view length/width ratio = 2.72– 3.30; apically with setae, length of longest setae ± width of paramere in dorso-ventral view. Aedeagus with median lobe laterally compressed, sclerotized, in lateral view strongly curved, apex tapering to rounded point. Ratio of penile strut length to median lobe length 1.54–1.66 (n = 2).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBE9513FF19FE8FD3B85A1C" blockId="11.[151,1437,151,1149]" box="[199,468,260,285]" pageId="11" pageNumber="162">
|
||||
<emphasis id="4386EAD3FFBE9513FF19FE8FD3725A1C" bold="true" box="[199,286,260,285]" pageId="11" pageNumber="162">Female</emphasis>
|
||||
similar to male.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBE9513FF19FEACD7235919" pageId="11" pageNumber="162" type="materials_examined">
|
||||
<paragraph id="714D36C1FFBE9513FF19FEACD63A5A41" blockId="11.[151,1437,151,1149]" box="[199,1110,295,320]" pageId="11" pageNumber="162">
|
||||
<materialsCitation id="C19A3C9CFFBE9513FF19FEACD63F5A41" ID-GBIF-Occurrence="2028138744" box="[199,1107,295,320]" country="Mexico" elevation="1918" location="Sierra Mixe" municipality="State" pageId="11" pageNumber="162" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
|
||||
<emphasis id="4386EAD3FFBE9513FF19FEACD3095A41" bold="true" box="[199,357,295,320]" pageId="11" pageNumber="162">
|
||||
<typeStatus id="AE498863FFBE9513FF19FEACD36E5A41" box="[199,258,295,320]" pageId="11" pageNumber="162">Type</typeStatus>
|
||||
locality.
|
||||
</emphasis>
|
||||
<collectingCountry id="09E57651FFBE9513FEB2FEACD3A85A41" box="[364,452,295,320]" name="Mexico" pageId="11" pageNumber="162">Mexico</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFBE9513FE11FEA3D04F5A41" box="[463,547,296,320]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="162">Oaxaca</collectingRegion>
|
||||
<collectingMunicipality id="9129ACBBFFBE9513FDF4FEA3D0095A41" box="[554,613,296,320]" pageId="11" pageNumber="162">State</collectingMunicipality>
|
||||
,
|
||||
<location id="742D601AFFBE9513FDAEFEACD0945A41" LSID="urn:lsid:plazi:treatment:F95B87D7FFBD9513FF49F94CD2955F7C:742D601AFFBE9513FDAEFEACD0945A41" box="[624,760,295,320]" country="Mexico" municipality="State" name="Sierra Mixe" pageId="11" pageNumber="162" stateProvince="Oaxaca">Sierra Mixe</location>
|
||||
, [GPS coord. omitted],
|
||||
<elevation id="FADFD1F2FFBE9513FBDFFEA3D63F5A41" box="[1025,1107,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.918" pageId="11" pageNumber="162" unit="m" value="1918.0">
|
||||
<quantity id="B60A9B24FFBE9513FBDFFEA3D63F5A41" box="[1025,1107,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.918" pageId="11" pageNumber="162" unit="m" value="1918.0">1918m</quantity>
|
||||
</elevation>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBE9513FF19FEC7D70B5A65" blockId="11.[151,1437,151,1149]" box="[199,1383,332,357]" pageId="11" pageNumber="162">
|
||||
<emphasis id="4386EAD3FFBE9513FF19FEC7D3765A64" bold="true" box="[199,282,332,357]" pageId="11" pageNumber="162">Range.</emphasis>
|
||||
<collectingRegion id="B336F823FFBE9513FEFFFEC7D3145A65" box="[289,376,332,356]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="162">Oaxaca</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="09E57651FFBE9513FE5DFEC7D3B45A64" box="[387,472,332,357]" name="Mexico" pageId="11" pageNumber="162">Mexico</collectingCountry>
|
||||
in the Sierra Mixe, known to inhabit only male cones of
|
||||
<taxonomicName id="B6F24D42FFBE9513FB86FEC7D70C5A65" box="[1112,1376,332,356]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBE9513FB86FEC7D70C5A65" box="[1112,1376,332,356]" italics="true" pageId="11" pageNumber="162">Ceratozamia mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBE9513FF19FEE4D7235919" blockId="11.[151,1437,151,1149]" pageId="11" pageNumber="162">
|
||||
<emphasis id="4386EAD3FFBE9513FF19FEE4D0485A89" bold="true" box="[199,548,367,392]" pageId="11" pageNumber="162">
|
||||
Material examined.
|
||||
<materialsCitation id="C19A3C9CFFBE9513FE69FEE4D0485A89" ID-GBIF-Occurrence="2028138747" box="[439,548,367,392]" collectingDate="2016-04-04" collectorName="W. Tang" country="Mexico" location="Sierra Mixe" pageId="11" pageNumber="162" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="AE498863FFBE9513FE69FEE4D0485A89" box="[439,548,367,392]" pageId="11" pageNumber="162" type="holotype">Holotype</typeStatus>
|
||||
</materialsCitation>
|
||||
</emphasis>
|
||||
(by designation) male of
|
||||
<taxonomicName id="B6F24D42FFBE9513FC9DFEFBD1AF5A86" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[835,963,368,391]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="gregoryi">
|
||||
<emphasis id="4386EAD3FFBE9513FC9DFEFBD1AF5A86" box="[835,963,368,391]" italics="true" pageId="11" pageNumber="162">C. gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||||
<collectingCountry id="09E57651FFBE9513FE2BFE1FD0045AAD" box="[501,616,404,428]" name="Mexico" pageId="11" pageNumber="162">MEXICO</collectingCountry>
|
||||
, Oax.,
|
||||
<location id="742D601AFFBE9513FD1DFE1FD13C5AAC" LSID="urn:lsid:plazi:treatment:F95B87D7FFBD9513FF49F94CD2955F7C:742D601AFFBE9513FD1DFE1FD13C5AAC" box="[707,848,404,429]" country="Mexico" name="Sierra Mixe" pageId="11" pageNumber="162">Sierra Mixe</location>
|
||||
, ex
|
||||
<taxonomicName id="B6F24D42FFBE9513FC56FE1FD6F95AAD" box="[904,1173,404,428]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBE9513FC56FE1FD6F95AAD" box="[904,1173,404,428]" italics="true" pageId="11" pageNumber="162">Ceratozamia mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
<specimenCount id="67F4FD48FFBE9513FB7CFE1ED6DB5AAD" box="[1186,1207,405,428]" pageId="11" pageNumber="162" type="male">♂</specimenCount>
|
||||
cone, [GPS coord. omitted],
|
||||
<collectingDate id="1508E9E9FFBE9513FED8FE3CD3115AD1" box="[262,381,439,464]" pageId="11" pageNumber="162" value="2016-04-04">
|
||||
<date id="054C1001FFBE9513FED8FE3CD3115AD1" box="[262,381,439,464]" pageId="11" pageNumber="162" value="2016-04-04">4-IV-2016</date>
|
||||
</collectingDate>
|
||||
,
|
||||
<collectorName id="DC075317FFBE9513FE55FE33D39D5AD1" box="[395,497,440,464]" pageId="11" pageNumber="162">W. Tang</collectorName>
|
||||
”; 2) [rectangular; red; printed in black ink]
|
||||
<materialsCitation id="C19A3C9CFFBE9513FC31FE33D13A5AF5" ID-GBIF-Occurrence="2028138737" collectionCode="FSCA" country="Mexico" pageId="11" pageNumber="162" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="AE498863FFBE9513FC31FE33D6EF5AD1" box="[1007,1155,440,464]" pageId="11" pageNumber="162" type="holotype">HOLOTYPE</typeStatus>
|
||||
<specimenCount id="67F4FD48FFBE9513FB53FE33D6CE5ACE" box="[1165,1186,440,463]" pageId="11" pageNumber="162" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFBE9513FB75FE33D0555AF5" authority="Tang, Skelley & Perez-Farrera 2018" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBE9513FB75FE33D7F05ACE" box="[1195,1436,440,464]" italics="true" pageId="11" pageNumber="162">Ceratophila gregoryi</emphasis>
|
||||
<bibRefCitation id="15634B30FFBE9513FF49FE56D0555AF5" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." box="[151,569,476,501]" pageId="11" pageNumber="162" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">Tang, Skelley & Pérez-Farrera 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. Deposited in the
|
||||
<collectionCode id="17E3AE04FFBE9513FCD2FE57D13A5AF5" box="[780,854,476,500]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="11" pageNumber="162" type="Museum">FSCA</collectionCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="C19A3C9CFFBE9513FCBCFE57D7275919" ID-GBIF-Occurrence="2028138740" collectionCode="FSCA, ANIC, BMNH, IEXA, INBio, MNHN, NZAC, UNAM, USNM" country="Mexico" pageId="11" pageNumber="162" specimenCount="229" specimenCount-adult="227" typeStatus="allotype">
|
||||
<typeStatus id="AE498863FFBE9513FCBCFE57D1AB5AF4" box="[866,967,476,501]" pageId="11" pageNumber="162" type="allotype">
|
||||
<emphasis id="4386EAD3FFBE9513FCBCFE57D1AB5AF4" bold="true" box="[866,967,476,501]" pageId="11" pageNumber="162">Allotype</emphasis>
|
||||
</typeStatus>
|
||||
(
|
||||
<collectionCode id="17E3AE04FFBE9513FC06FE57D64D5AF5" box="[984,1057,476,500]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="11" pageNumber="162" type="Museum">FSCA</collectionCode>
|
||||
) and
|
||||
<specimenCount id="67F4FD48FFBE9513FBBAFE57D6BC5AF4" box="[1124,1232,476,501]" pageId="11" pageNumber="162" type="adult">227 adult</specimenCount>
|
||||
<typeStatus id="AE498863FFBE9513FB07FE57D73C5AF4" box="[1241,1360,476,501]" pageId="11" pageNumber="162" type="paratype">
|
||||
<emphasis id="4386EAD3FFBE9513FB07FE57D73C5AF4" bold="true" box="[1241,1360,476,501]" pageId="11" pageNumber="162">paratypes</emphasis>
|
||||
</typeStatus>
|
||||
(same label data as
|
||||
<typeStatus id="AE498863FFBE9513FEF6FE74D3E05919" box="[296,396,511,536]" pageId="11" pageNumber="162" type="holotype">holotype</typeStatus>
|
||||
) deposited at
|
||||
<collectionCode id="17E3AE04FFBE9513FDF6FD8BD0035919" box="[552,623,512,536]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="162">ANIC</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFBE9513FDA5FD8BD0BE5916" box="[635,722,512,535]" pageId="11" pageNumber="162">BMNH</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFBE9513FD03FD8BD14A5919" box="[733,806,512,536]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="11" pageNumber="162" type="Museum">FSCA</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFBE9513FCEEFD8BD11A5919" box="[816,886,512,536]" pageId="11" pageNumber="162">IEXA</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFBE9513FC5FFE74D1A65919" box="[897,970,511,536]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="11" pageNumber="162">INBio</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFBE9513FC0BFD8BD6425916" box="[981,1070,512,535]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="162">MNHN</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFBE9513FBE7FD8BD6E45919" box="[1081,1160,512,536]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="11" pageNumber="162" type="Other institutional type">NZAC</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFBE9513FB4DFD8BD6875919" box="[1171,1259,512,536]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="11" pageNumber="162">UNAM</collectionCode>
|
||||
,
|
||||
<collectionCode id="17E3AE04FFBE9513FB28FD8BD7275919" box="[1270,1355,512,536]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="162" type="Museum">USNM</collectionCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBE9513FF19FDAFD0FB5961" pageId="11" pageNumber="162" type="etymology">
|
||||
<paragraph id="714D36C1FFBE9513FF19FDAFD0FB5961" blockId="11.[151,1437,151,1149]" pageId="11" pageNumber="162">
|
||||
<emphasis id="4386EAD3FFBE9513FF19FDAFD322593C" bold="true" box="[199,334,548,573]" pageId="11" pageNumber="162">Etymology.</emphasis>
|
||||
The species is named in honor of Dr. Timothy Gregory for his contributions to the taxonomy, ecology and conservation of Mexican cycads.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBE9513FF19FDE7D2955F7C" pageId="11" pageNumber="162" type="discussion">
|
||||
<paragraph id="714D36C1FFBE9513FF19FDE7D2955F7C" blockId="11.[151,1437,151,1149]" pageId="11" pageNumber="162">
|
||||
<emphasis id="4386EAD3FFBE9513FF19FDE7D3575984" bold="true" box="[199,315,620,645]" pageId="11" pageNumber="162">Remarks.</emphasis>
|
||||
Species lacking dark marks on the elytra superficially resemble
|
||||
<taxonomicName id="B6F24D42FFBE9513FBEAFDE7D6D45985" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[1076,1208,620,644]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="gregoryi">
|
||||
<emphasis id="4386EAD3FFBE9513FBEAFDE7D6D45985" box="[1076,1208,620,644]" italics="true" pageId="11" pageNumber="162">C. gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
. Male genitalia are diagnostic, but unassociated females will be difficult to assign. Pronotal shapes are characteristic also (
|
||||
<taxonomicName id="B6F24D42FFBE9513FA5CFD1BD36659CD" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="sanchezae">
|
||||
<emphasis id="4386EAD3FFBE9513FA5CFD1BD36659CD" italics="true" pageId="11" pageNumber="162">C. sanchezae</emphasis>
|
||||
</taxonomicName>
|
||||
more quadrate,
|
||||
<taxonomicName id="B6F24D42FFBE9513FE61FD3FD03E59CD" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[447,594,692,716]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="picipennis">
|
||||
<emphasis id="4386EAD3FFBE9513FE61FD3FD03E59CD" box="[447,594,692,716]" italics="true" pageId="11" pageNumber="162">C. picipennis</emphasis>
|
||||
</taxonomicName>
|
||||
more elongate,
|
||||
<taxonomicName id="B6F24D42FFBE9513FCD8FD3FD1E959CD" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[774,901,692,716]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="gregoryi">
|
||||
<emphasis id="4386EAD3FFBE9513FCD8FD3FD1E959CD" box="[774,901,692,716]" italics="true" pageId="11" pageNumber="162">C. gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
more anteriorly narrowing), but vary enough to be problematic on individual specimens. In all of these species, knowing where it was captured or the host cycad, will help identify them. Molecular work is currently underway to test these species hypotheses, but names need to be made available for current natural history and ecological work on both beetles and cycads. The only known host of this beetle is
|
||||
<taxonomicName id="B6F24D42FFBE9513FE86FCCFD008585D" box="[344,612,836,860]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBE9513FE86FCCFD008585D" box="[344,612,836,860]" italics="true" pageId="11" pageNumber="162">Ceratozamia mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
. This cycad is reported to inhabit tropical cloud forest in the extreme eastern Sierra Norte de
|
||||
<collectingRegion id="B336F823FFBE9513FE69FCE3D0675881" box="[439,523,872,896]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="162">Oaxaca</collectingRegion>
|
||||
(Sierra Mixe) at elevations from
|
||||
<quantity id="B60A9B24FFBE9513FC78FCECD6565881" box="[934,1082,871,896]" metricMagnitude="3" metricUnit="m" metricValue="1.6675" metricValueMax="1.895" metricValueMin="1.44" pageId="11" pageNumber="162" unit="m" value="1667.5" valueMax="1895.0" valueMin="1440.0">1440–1895m</quantity>
|
||||
(
|
||||
<bibRefCitation id="15634B30FFBE9513FB90FCECD73B5881" author="Chemnick, J. & Gregory, T. J. & Salas-Morales, S." box="[1102,1367,871,896]" pageId="11" pageNumber="162" pagination="47 - 52" refId="ref18885" refString="Chemnick, J., Gregory, T. J. & Salas-Morales, S. (1997) Ceratozamia mixeorum (Zamiaceae), a new species from Oaxaca, Mexico with comments on distribution, habitat, and species relationships. Phytologia, 83, 47 - 52." type="journal article" year="1997">
|
||||
Chemnick
|
||||
<emphasis id="4386EAD3FFBE9513FB0EFCE2D76B5881" box="[1232,1287,872,896]" italics="true" pageId="11" pageNumber="162">et al</emphasis>
|
||||
. 1997
|
||||
</bibRefCitation>
|
||||
). The specimens used in this study were collected at a slightly higher elevation of
|
||||
<quantity id="B60A9B24FFBE9513FC25FC07D62158A5" box="[1019,1101,908,932]" metricMagnitude="3" metricUnit="m" metricValue="1.918" pageId="11" pageNumber="162" unit="m" value="1918.0">1918m</quantity>
|
||||
. This habitat appears isolated and is wetter, higher and cooler than surrounding lowlands and relatively distant from other known
|
||||
<taxonomicName id="B6F24D42FFBE9513FAD5FC3BD7F058C6" box="[1291,1436,944,967]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBE9513FAD5FC3BD7F058C6" box="[1291,1436,944,967]" italics="true" pageId="11" pageNumber="162">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
populations. Unlike other
|
||||
<taxonomicName id="B6F24D42FFBE9513FE61FC5FD03C58EA" box="[447,592,980,1003]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBE9513FE61FC5FD03C58EA" box="[447,592,980,1003]" italics="true" pageId="11" pageNumber="162">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
in the region, our sample of beetles suggests
|
||||
<taxonomicName id="B6F24D42FFBE9513FB85FC5FD68158ED" box="[1115,1261,980,1004]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="mixeorum">
|
||||
<emphasis id="4386EAD3FFBE9513FB85FC5FD68158ED" box="[1115,1261,980,1004]" italics="true" pageId="11" pageNumber="162">C. mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
hosts only two species of
|
||||
<taxonomicName id="B6F24D42FFBE9513FEC2FC7CD3B75F11" authorityName="Crowson" authorityYear="1952" box="[284,475,1015,1040]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFBE9513FE33FC73D1145F0E" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[493,888,1016,1040]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFBE9513FE33FC73D0185F11" box="[493,628,1016,1040]" italics="true" pageId="11" pageNumber="162">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFBE9513FD53FC73D0825F11" box="[653,750,1016,1040]" italics="true" pageId="11" pageNumber="162">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFBE9513FCD6FC73D1145F0E" box="[776,888,1016,1039]" italics="true" pageId="11" pageNumber="162">mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<emphasis id="4386EAD3FFBE9513FC63FC73D1BB5F0E" box="[957,983,1016,1039]" italics="true" pageId="11" pageNumber="162">C.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFBE9513FC38FC73D6135F0E" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[998,1151,1016,1040]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="gregoryi" subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFBE9513FC31FC73D66E5F0E" box="[1007,1026,1016,1039]" italics="true" pageId="11" pageNumber="162">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFBE9513FBFEFC73D6135F0E" box="[1056,1151,1016,1039]" italics="true" pageId="11" pageNumber="162">gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="B6F24D42FFBE9513FB4BFC73D7595F11" authorityName="Reitter" authorityYear="1875" box="[1173,1333,1016,1040]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFBE9513FB4BFC73D7595F11" box="[1173,1333,1016,1040]" italics="true" pageId="11" pageNumber="162">Pharaxonotha</emphasis>
|
||||
</taxonomicName>
|
||||
species, typically found on male cones of other
|
||||
<taxonomicName id="B6F24D42FFBE9513FD8AFB97D0895F32" box="[596,741,1052,1075]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBE9513FD8AFB97D0895F32" box="[596,741,1052,1075]" italics="true" pageId="11" pageNumber="162">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
species, were absent in our sample. In two cones sampled
|
||||
<emphasis id="4386EAD3FFBE9513FA5CFB97D7F05F32" box="[1410,1436,1052,1075]" italics="true" pageId="11" pageNumber="162">C.</emphasis>
|
||||
<taxonomicName id="B6F24D42FFBE9513FF49FBCBD3455F56" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[151,297,1088,1112]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="gregoryi" subGenus="Ceratophila">
|
||||
(
|
||||
<emphasis id="4386EAD3FFBE9513FF7EFBCBD2DF5F56" box="[160,179,1088,1111]" italics="true" pageId="11" pageNumber="162">C</emphasis>
|
||||
.)
|
||||
<emphasis id="4386EAD3FFBE9513FF14FBCBD3455F56" box="[202,297,1088,1111]" italics="true" pageId="11" pageNumber="162">gregoryi</emphasis>
|
||||
</taxonomicName>
|
||||
accounted for 84% of
|
||||
<specimenCount id="67F4FD48FFBE9513FDF4FBCBD0FF5F59" box="[554,659,1087,1112]" pageId="11" pageNumber="162" type="adult">273 adult</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFBE9513FD44FBB4D13A5F59" authorityName="Crowson" authorityYear="1952" box="[666,854,1087,1112]" class="Insecta" family="Erotylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
|
||||
collected, while
|
||||
<taxonomicName id="B6F24D42FFBE9513FBCBFBCBD74F5F56" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1045,1315,1088,1112]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="mixeorum" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFBE9513FBCBFBCBD6445F56" box="[1045,1064,1088,1111]" italics="true" pageId="11" pageNumber="162">C</emphasis>
|
||||
. (
|
||||
<emphasis id="4386EAD3FFBE9513FBE1FBCBD6CC5F59" box="[1087,1184,1088,1112]" italics="true" pageId="11" pageNumber="162">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFBE9513FB6DFBCBD74F5F56" box="[1203,1315,1088,1111]" italics="true" pageId="11" pageNumber="162">mixeorum</emphasis>
|
||||
</taxonomicName>
|
||||
accounted for 16%.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
453
data/F9/5B/87/F95B87D7FFBE9516FF49FB4ED7E158C9.xml
Normal file
453
data/F9/5B/87/F95B87D7FFBE9516FF49FB4ED7E158C9.xml
Normal file
|
|
@ -0,0 +1,453 @@
|
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<mods:title id="7766B3DC366A41D91A3D11AD93DCB8EC">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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<mods:namePart id="5E3576E2D19B5AF99FB8A3991BE1442A">Skelley, Paul</mods:namePart>
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<mods:namePart id="904CF97732B798B7868E6D7BC5CFF71B">Pérez-Farrera, Miguel Angel</mods:namePart>
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<mods:title id="1CAEC2144367B0B8871680D5573F1B2E">Zootaxa</mods:title>
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<mods:date id="AB3109C67BC1278E15D8035F7D0450D5">2018</mods:date>
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<mods:number id="138015BE95AC5761F8594C65501409EE">2018-10-31</mods:number>
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<subSubSection id="39E8654AFFBE9513FF49FB4ED6155FDE" box="[151,1145,1221,1247]" pageId="11" pageNumber="162" type="nomenclature">
|
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<paragraph id="714D36C1FFBE9513FF49FB4ED6155FDE" blockId="11.[151,1145,1221,1280]" box="[151,1145,1221,1247]" pageId="11" pageNumber="162">
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<heading id="2A0581ADFFBE9513FF49FB4ED6155FDE" bold="true" box="[151,1145,1221,1247]" fontSize="11" level="1" pageId="11" pageNumber="162" reason="1">
|
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<emphasis id="4386EAD3FFBE9513FF49FB4ED6155FDE" bold="true" box="[151,1145,1221,1247]" pageId="11" pageNumber="162">
|
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<taxonomicName id="B6F24D42FFBE9513FF49FB4ED1B05FDE" authority="Tang, Skelley & Perez-Farrera" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[151,988,1221,1247]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFBE9513FF49FB4ED34B5FDE" bold="true" box="[151,295,1221,1247]" italics="true" pageId="11" pageNumber="162">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFBE9513FEE9FB4ED3AA5FDE" bold="true" box="[311,454,1221,1247]" italics="true" pageId="11" pageNumber="162">Ceratophila</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFBE9513FE09FB4ED03F5FDE" bold="true" box="[471,595,1221,1247]" italics="true" pageId="11" pageNumber="162">picipennis</emphasis>
|
||||
Tang, Skelley & Pérez-Farrera
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="58B557A8FFBE9513FC37FB4ED6155FDE" box="[1001,1145,1221,1247]" pageId="11" pageNumber="162" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBE9513FF49FB6CD0D55E01" box="[151,697,1255,1280]" pageId="11" pageNumber="162" type="description">
|
||||
<paragraph id="714D36C1FFBE9513FF49FB6CD0D55E01" blockId="11.[151,1145,1221,1280]" box="[151,697,1255,1280]" pageId="11" pageNumber="162">
|
||||
(
|
||||
<figureCitation id="E9C92A44FFBE9513FF41FB6CD2805E01" box="[159,236,1255,1280]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="11" pageNumber="162">Figs. 4</figureCitation>
|
||||
A–H; photograph 1F in
|
||||
<bibRefCitation id="15634B30FFBE9513FE27FB63D0DD5E01" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." box="[505,689,1256,1280]" pageId="11" pageNumber="162" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">
|
||||
Tang
|
||||
<emphasis id="4386EAD3FFBE9513FDE6FB62D01C5E01" box="[568,624,1256,1280]" italics="true" pageId="11" pageNumber="162">et al.</emphasis>
|
||||
2018
|
||||
</bibRefCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBE9513FF49FAA4D3A85EB5" pageId="11" pageNumber="162" type="diagnosis">
|
||||
<paragraph id="714D36C1FFBE9513FF49FAA4D3A85EB5" blockId="11.[151,1437,1327,2037]" pageId="11" pageNumber="162">
|
||||
<emphasis id="4386EAD3FFBE9513FF49FAA4D3305E49" bold="true" box="[151,348,1327,1352]" pageId="11" pageNumber="162">Adult diagnosis.</emphasis>
|
||||
Distinguished from other
|
||||
<taxonomicName id="B6F24D42FFBE9513FD4FFABBD1D15E49" box="[657,957,1328,1352]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFBE9513FD4FFABBD1745E49" box="[657,792,1328,1352]" italics="true" pageId="11" pageNumber="162">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFBE9513FCF2FABBD1DF5E49" box="[812,947,1328,1352]" italics="true" pageId="11" pageNumber="162">Ceratophila</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
species by the elongate pronotum; when present dark elytral markings covering most of disc; male genitalia with length of penile strut relative to median lobe ~ 3:2; and with a known distribution in
|
||||
<collectingCountry id="09E57651FFBE9513FD4CFAFCD0865E91" box="[658,746,1399,1424]" name="Mexico" pageId="11" pageNumber="162">Mexico</collectingCountry>
|
||||
, state of
|
||||
<collectingRegion id="B336F823FFBE9513FC8CFAFCD1C35E91" box="[850,943,1399,1424]" country="Mexico" name="Chiapas" pageId="11" pageNumber="162">Chiapas</collectingRegion>
|
||||
, on
|
||||
<taxonomicName id="B6F24D42FFBE9513FC00FAF3D6BB5E91" authorityName="Perez-Farrera, Vovides & Iglesias" authorityYear="1999" box="[990,1239,1400,1424]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="alvarezii">
|
||||
<emphasis id="4386EAD3FFBE9513FC00FAF3D6BB5E91" box="[990,1239,1400,1424]" italics="true" pageId="11" pageNumber="162">Ceratozamia alvarezii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFBE9513FB38FAF3D71A5E91" box="[1254,1398,1400,1424]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFBE9513FB38FAF3D6955E8E" box="[1254,1273,1400,1423]" italics="true" pageId="11" pageNumber="162">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFBE9513FAD6FAF3D71A5E91" box="[1288,1398,1400,1424]" italics="true" pageId="11" pageNumber="162">mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="4386EAD3FFBE9513FAA8FAF3D2905EB5" italics="true" pageId="11" pageNumber="162">
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFBE9513FA5CFAF3D2905EB5" authorityName="D. W. Stevenson" authorityYear="1982" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="norstogii">C. norstogii</taxonomicName>
|
||||
</emphasis>
|
||||
, and
|
||||
<taxonomicName id="B6F24D42FFBE9513FEE7FA17D3D25EB5" box="[313,446,1436,1460]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="11" pageNumber="162" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFBE9513FEE7FA17D3205EB2" box="[313,332,1436,1459]" italics="true" pageId="11" pageNumber="162">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFBE9513FE84FA17D3D25EB5" box="[346,446,1436,1460]" italics="true" pageId="11" pageNumber="162">vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBE9515FF10FA34D3B85D74" lastPageId="13" lastPageNumber="164" pageId="11" pageNumber="162" type="description">
|
||||
<paragraph id="714D36C1FFBE9513FF10FA34D6855DAE" blockId="11.[151,1437,1327,2037]" pageId="11" pageNumber="162">
|
||||
<emphasis id="4386EAD3FFBE9513FF10FA34D38B5ED9" bold="true" box="[206,487,1471,1496]" pageId="11" pageNumber="162">Adult male description.</emphasis>
|
||||
Length
|
||||
<quantity id="B60A9B24FFBE9513FD9DFA34D0845ED6" box="[579,744,1471,1496]" metricMagnitude="-3" metricUnit="m" metricValue="3.78" metricValueMax="4.53" metricValueMin="3.03" pageId="11" pageNumber="162" unit="mm" value="3.78" valueMax="4.53" valueMin="3.03">3.03–4.53 mm</quantity>
|
||||
, width
|
||||
<quantity id="B60A9B24FFBE9513FCE4FA34D1B15ED6" box="[826,989,1471,1496]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.79" metricValueMin="1.26" pageId="11" pageNumber="162" unit="mm" value="1.525" valueMax="1.79" valueMin="1.26">1.26–1.79 mm</quantity>
|
||||
(n = 20). Body in dorsal view elongateoval, greatest width at middle of elytra; in lateral view convex dorsally. General body color brown (
|
||||
<figureCitation id="E9C92A44FFBE9513FAD4FA6FD7355EFD" box="[1290,1369,1508,1533]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="11" pageNumber="162">Figs. 4</figureCitation>
|
||||
A–E), head and pronotum concolorous and darker than base of elytra; central portion of elytra usually with black to dark brown coloration fading gradually to light brown at base and apex, some individuals without black markings or with markings confined to elytral margins; dorsal surface punctate, shining and appearing glabrous, short procumbent hairs associated with punctation along lateral sides and declivity of elytra, ventrally shining and appearing glabrous except mesoventrite and abdomen mostly covered with long procumbent setae.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFBE9515FF19F937D6295BF9" blockId="11.[151,1437,1327,2037]" lastBlockId="13.[151,1437,151,2013]" lastPageId="13" lastPageNumber="164" pageId="11" pageNumber="162">
|
||||
<emphasis id="4386EAD3FFBE9513FF19F937D36B5DD4" bold="true" box="[199,263,1724,1749]" pageId="11" pageNumber="162">Head</emphasis>
|
||||
not broad, width = 0.66–0.68× pronotal width; in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured (
|
||||
<figureCitation id="E9C92A44FFBE9513FCB1F954D66C5DF9" box="[879,1024,1759,1784]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="11" pageNumber="162">Figs. 4A, D</figureCitation>
|
||||
), average distance between closest punctures 2× width of puncture; width
|
||||
<quantity id="B60A9B24FFBE9513FD96F88FD0825C1D" box="[584,750,1796,1821]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="9.3" metricValueMin="6.5" pageId="11" pageNumber="162" unit="mm" value="0.79" valueMax="0.93" valueMin="0.65">0.65–0.93 mm</quantity>
|
||||
; dorsal interocular distance
|
||||
<quantity id="B60A9B24FFBE9513FBFBF88FD6A65C1D" box="[1061,1226,1796,1821]" metricMagnitude="-4" metricUnit="m" metricValue="5.05" metricValueMax="5.8999999999999995" metricValueMin="4.2" pageId="11" pageNumber="162" unit="mm" value="0.505" valueMax="0.59" valueMin="0.42">0.42–0.59 mm</quantity>
|
||||
, head width/dorsal interocular distance ratio 1.52–1.69, ventral interocular distance
|
||||
<quantity id="B60A9B24FFBE9513FC56F8ACD65E5C3E" box="[904,1074,1831,1856]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="3.8" metricValueMin="2.6" pageId="11" pageNumber="162" unit="mm" value="0.32" valueMax="0.38" valueMin="0.26">0.26–0.38 mm</quantity>
|
||||
, head width/ventral interocular distance ratio 2.32–2.70. Eye with large black facets. Antennal length slightly shorter than pronotal width, 1.3× head width; antennomere I (scape) fairly large, slightly elongate; antennomere II slightly smaller than III; IV–VII small, width equals length; VIII width slightly more than length; club fairly large, IX and X similar in length, XI slightly longer, globular with angulate apex. Clypeus truncate anteriorly, moderately punctate; somewhat emarginate, with narrow margin. Mentum and submentum with moderate punctation, distance between nearest punctures approximately 1× own diameter, each puncture with a short seta. Gular area smooth, without punctation or setae, border with submentum usually with a shallow depression without punctation, but usually with a shallowly impressed circular or semicircular border along anterior margin (
|
||||
<figureCitation id="E9C92A44FFB89515FC3FFF54D6555BF9" box="[993,1081,223,248]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="13" pageNumber="164">Fig. 4E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="258D6649FFB99514FF49F8A4D7ED5C83" httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="12" pageNumber="163" startId="12.[151,250,1839,1861]" targetBox="[168,1428,199,1802]" targetPageId="12">
|
||||
<paragraph id="714D36C1FFB99514FF49F8A4D7ED5C83" blockId="12.[151,1436,1839,1922]" pageId="12" pageNumber="163">
|
||||
<emphasis id="4386EAD3FFB99514FF49F8A4D3775C44" bold="true" box="[151,283,1839,1861]" pageId="12" pageNumber="163">FIGURE 4.</emphasis>
|
||||
Photographs of
|
||||
<taxonomicName id="B6F24D42FFB99514FE04F8A4D1095C44" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[474,869,1839,1861]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="12" pageNumber="163" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFB99514FE04F8A4D0395C44" box="[474,597,1839,1861]" italics="true" pageId="12" pageNumber="163">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFB99514FDB2F8A4D08A5C44" box="[620,742,1839,1861]" italics="true" pageId="12" pageNumber="163">Ceratophila</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFB99514FD23F8BBD1095C44" box="[765,869,1840,1861]" italics="true" pageId="12" pageNumber="163">picipennis</emphasis>
|
||||
</taxonomicName>
|
||||
collected from
|
||||
<taxonomicName id="B6F24D42FFB99514FBC3F8BBD77C5C44" box="[1053,1296,1839,1861]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="12" pageNumber="163" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFB99514FBC3F8BBD77C5C44" box="[1053,1296,1839,1861]" italics="true" pageId="12" pageNumber="163">Ceratozamia mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
(A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="714D36C1FFB89515FF19FE8FD09D58EC" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
|
||||
<emphasis id="4386EAD3FFB89515FF19FE8FD34C5A1C" bold="true" box="[199,288,260,285]" pageId="13" pageNumber="164">Thorax</emphasis>
|
||||
with pronotum quadrate in dorsal view, length/width ratio 0.83–0.90; with marginal beads laterally and basally, anterior margin lacking distinct bead medially; convex; anterior and posterior angles weakly developed; basal ¼ expanding to parallel lateral carinae, then gradually converging in apical ½; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores located 1/7 length of posterior margin from posterior corners and touching and just anterior to marginal bead, longitudinal furrows lacking. Prosternum in ventral view convex; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae approximately ½ length of eye. Hypomeron laterally with few minute punctures, medially with longitudinal striations. Elytra in dorsal view elongate-oval, convex; length/width 1.63– 1.78, greatest width near midlength; without marginal line basally; 10 complete striae of moderate puncture size; scutellary striole extending ¼ elytral length, with 10–15 punctures; intervals of striae with fine, shallow punctures. All punctures of elytral striae bearing a single short seta; seta only visible in profile, extending slightly out of puncture. Mesoventrite with moderate to strong punctation, distance between nearest punctures approximately equal to diameter of punctures, puncture depth moderate. Metaventrite dulled laterally, with strong lateral punctation separated by 2–3× own diameter; medial surface smooth, glossy, finely punctured, separated by 5–6× own diameter; convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over half metaventrite length. Legs stout, relatively similar in length and shape. Procoxa oval; mesocoxa globular; metacoxa transversely elongate-oval; trochanters obliquely truncate apically; femora robust, moderately compressed laterally; tibiae shorter than femora, gradually dilated to obliquely truncate apices; protibia with disto-lateral tooth (
|
||||
<figureCitation id="E9C92A44FFB89515FF41FC07D28158A5" box="[159,237,908,933]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="13" pageNumber="164">Figs. 4</figureCitation>
|
||||
D–E), with apical fringe of short, stout spinules of posterior margin lacking along concave medial margin (row broken) and long stout setae on anterior margin; meso- and metatibia with apical fringe of short, stout spinules on anterior margin, longer setae on posterior margins.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB89515FF19FC7CD6555D51" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
|
||||
<emphasis id="4386EAD3FFB89515FF19FC7CD32F5F11" bold="true" box="[199,323,1015,1040]" pageId="13" pageNumber="164">Abdomen.</emphasis>
|
||||
Ventrite I with intercoxal process narrow, with triangular point anteromedially; lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, dense setae; all ventrites bearing moderate, shallow punctation, distance to nearest puncture approximately 1× diameter of puncture, punctures bearing mostly reclining setae; ventrite V with setae length nearly uniformly approximately 2× diameter of puncture; ventrites I–IV with punctation and setae confined to median band, setae length approximately 1× diameter of puncture along anterior edge of band increasing to approximately 3× diameter along posterior edge; I–IV each with 2 or more median pairs of longer, semi-erect sensory hairs (often abraded), V with 2–3 pairs of long semi-erect hairs located postero-laterally. Male genital capsule fringed with fine setae. Male genitalia (tegmen and median lobe) oriented upside down (displayed right side up in
|
||||
<figureCitation id="E9C92A44FFB89515FB75FAD4D76B5E79" box="[1195,1287,1375,1400]" captionStart="FIGURE 4" captionStartId="12.[151,250,1839,1861]" captionTargetBox="[168,1428,199,1802]" captionTargetId="figure@12.[168,1428,199,1802]" captionTargetPageId="12" captionText="FIGURE 4. Photographs of Ceratophila (Ceratophila) picipennis collected from Ceratozamia mirandae (A–E: male holotype): A–C) dorsal, ventral and lateral and habitus; D–F) head, dorsal and ventral views; G) male genital capsule, ventral view; H) median lobe still attached with tegmen, lateral and ventral views; I) close up of parameres of tegmen, ventral view." httpUri="https://zenodo.org/record/2606944/files/figure.png" pageId="13" pageNumber="164">Fig. 4G</figureCitation>
|
||||
to facilitate comparisons). Tegmen sclerotized, triangular, laterally compressed; with anterior region ring-like, posterior region sheath-like; lateral margins gradually converging posteriorly; apically with 2 elongate, somewhat bar-shaped parameres. Parameres dorso-ventrally compressed, height approximately 1/2 its own width, in dorso-ventral view length/width ratio = 2.3–2.7; apically with setae, length of longest setae <width of paramere in dorso-ventral view. Aedeagus with median lobe laterally compressed, sclerotized, in lateral view strongly curved, apex tapering to rounded point. Ratio of penile strut length to median lobe length 1.36–1.57 (n = 3).
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB89515FF19F9D7D3B85D74" blockId="13.[151,1437,151,2013]" box="[199,468,1628,1653]" pageId="13" pageNumber="164">
|
||||
<emphasis id="4386EAD3FFB89515FF19F9D7D3725D74" bold="true" box="[199,286,1628,1653]" pageId="13" pageNumber="164">Female</emphasis>
|
||||
similar to male.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFB89516FF19F9F4D07D5AD1" lastPageId="14" lastPageNumber="165" pageId="13" pageNumber="164" type="materials_examined">
|
||||
<paragraph id="714D36C1FFB89515FF19F9F4D2B25DBD" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
|
||||
<materialsCitation id="C19A3C9CFFB89515FF19F9F4D2B25DBD" ID-GBIF-Occurrence="2028138742" collectorName="de Villaflores" country="Mexico" elevation="980" location="Municipio de Villaflores" pageId="13" pageNumber="164" specimenCount="980" stateProvince="Chiapas" typeStatus="holotype">
|
||||
<emphasis id="4386EAD3FFB89515FF19F9F4D3065D99" bold="true" box="[199,362,1663,1688]" pageId="13" pageNumber="164">
|
||||
<typeStatus id="AE498863FFB89515FF19F9F4D36E5D99" box="[199,258,1663,1688]" pageId="13" pageNumber="164">Type</typeStatus>
|
||||
locality.
|
||||
</emphasis>
|
||||
<collectingCountry id="09E57651FFB89515FEA8F9F4D3A25D99" box="[374,462,1663,1688]" name="Mexico" pageId="13" pageNumber="164">Mexico</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFB89515FE01F9F4D0555D99" box="[479,569,1663,1688]" country="Mexico" name="Chiapas" pageId="13" pageNumber="164">Chiapas</collectingRegion>
|
||||
State,
|
||||
<location id="742D601AFFB89515FD4FF9F4D1DD5D99" LSID="urn:lsid:plazi:treatment:F95B87D7FFBE9516FF49FB4ED7E158C9:742D601AFFB89515FD4FF9F4D1DD5D99" box="[657,945,1663,1688]" country="Mexico" name="Municipio de Villaflores" pageId="13" pageNumber="164" stateProvince="Chiapas">
|
||||
Municipio
|
||||
<collectorName id="DC075317FFB89515FCCCF9F4D1DD5D99" box="[786,945,1663,1688]" pageId="13" pageNumber="164">de Villaflores</collectorName>
|
||||
</location>
|
||||
, near Champerico, [GPS coord. omitted],
|
||||
<specimenCount id="67F4FD48FFB89515FF49F92FD2B25DBD" box="[151,222,1700,1724]" pageId="13" pageNumber="164" type="generic">
|
||||
<elevation id="FADFD1F2FFB89515FF49F92FD2B65DBD" box="[151,218,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="13" pageNumber="164" unit="m" value="980.0">
|
||||
<quantity id="B60A9B24FFB89515FF49F92FD2B65DBD" box="[151,218,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="13" pageNumber="164" unit="m" value="980.0">980m</quantity>
|
||||
</elevation>
|
||||
.
|
||||
</specimenCount>
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB89515FF19F94CD76C5DE1" blockId="13.[151,1437,151,2013]" box="[199,1280,1735,1760]" pageId="13" pageNumber="164">
|
||||
<emphasis id="4386EAD3FFB89515FF19F94CD3765DE1" bold="true" box="[199,282,1735,1760]" pageId="13" pageNumber="164">Range.</emphasis>
|
||||
Sierra Madre de
|
||||
<collectingRegion id="B336F823FFB89515FE03F94CD0565DE1" box="[477,570,1735,1760]" country="Mexico" name="Chiapas" pageId="13" pageNumber="164">Chiapas</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="09E57651FFB89515FD9BF94CD0F65DE1" box="[581,666,1735,1760]" name="Mexico" pageId="13" pageNumber="164">Mexico</collectingCountry>
|
||||
inhabiting male cones of four species of
|
||||
<taxonomicName id="B6F24D42FFB89515FBB6F943D6955DDE" box="[1128,1273,1736,1759]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="13" pageNumber="164" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFB89515FBB6F943D6955DDE" box="[1128,1273,1736,1759]" italics="true" pageId="13" pageNumber="164">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="714D36C1FFB89516FF19F967D07D5AD1" blockId="13.[151,1437,151,2013]" lastBlockId="14.[151,1436,151,968]" lastPageId="14" lastPageNumber="165" pageId="13" pageNumber="164">
|
||||
<emphasis id="4386EAD3FFB89515FF19F967D0545C04" bold="true" box="[199,568,1772,1797]" pageId="13" pageNumber="164">
|
||||
Material examined.
|
||||
<typeStatus id="AE498863FFB89515FE15F967D0545C04" box="[459,568,1772,1797]" pageId="13" pageNumber="164" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
(by designation) male of
|
||||
<taxonomicName id="B6F24D42FFB89515FC57F967D64B5C05" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[905,1063,1772,1796]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="13" pageNumber="164" phylum="Tracheophyta" rank="species" species="picipennis">
|
||||
<emphasis id="4386EAD3FFB89515FC57F967D64B5C05" box="[905,1063,1772,1796]" italics="true" pageId="13" pageNumber="164">C. picipennis</emphasis>
|
||||
</taxonomicName>
|
||||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||||
<collectingCountry id="09E57651FFB89515FD79F89BD1775C29" box="[679,795,1808,1832]" name="Mexico" pageId="13" pageNumber="164">MEXICO</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="B336F823FFB89515FCF1F884D1E05C29" box="[815,908,1807,1832]" country="Mexico" name="Chiapas" pageId="13" pageNumber="164">Chiapas</collectingRegion>
|
||||
, Champerico, [GPS coord. omitted], ex
|
||||
<specimenCount id="67F4FD48FFB89515FA59F89AD7F05C29" box="[1415,1436,1809,1832]" pageId="13" pageNumber="164" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFB89515FF49F8BFD3F65C4D" box="[151,410,1844,1868]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="13" pageNumber="164" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFB89515FF49F8BFD3F65C4D" box="[151,410,1844,1868]" italics="true" pageId="13" pageNumber="164">Ceratozamia mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
cone,
|
||||
<date id="054C1001FFB89515FE3AF8BFD0335C4D" box="[484,607,1844,1868]" pageId="13" pageNumber="164" value="2017-05-20">
|
||||
20-
|
||||
<bibRefCitation id="15634B30FFB89515FDD6F8BFD0335C4D" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[520,607,1844,1868]" pageId="13" pageNumber="164" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">V-2017</bibRefCitation>
|
||||
</date>
|
||||
, H. Gomez Dominguez & W. Tang”; 2) [rectangular; red; printed in black ink]
|
||||
<typeStatus id="AE498863FFB89515FF14F8D3D3325C71" box="[202,350,1880,1904]" pageId="13" pageNumber="164" type="holotype">HOLOTYPE</typeStatus>
|
||||
<specimenCount id="67F4FD48FFB89515FEBBF8D2D3165C71" box="[357,378,1881,1904]" pageId="13" pageNumber="164" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFB89515FE5EF8D3D6715C71" authority="Tang, Skelley & Perez-Farrera 2018" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[384,1053,1879,1905]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="13" pageNumber="164" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFB89515FE5EF8D3D0EC5C6E" box="[384,640,1880,1904]" italics="true" pageId="13" pageNumber="164">Ceratophila picipennis</emphasis>
|
||||
<bibRefCitation id="15634B30FFB89515FD59F8D2D6715C71" author="Tang, W. & Xu, G. & O'Brien, C. W. & Calonje, M. & Franz, N. M. & Johnston, M. A. & Taylor, A. & Vovides, A. P. & Perez-Farrera, M. A. & Salas-Morales, S. H. & Lazcano-Lara, J. C. & Skelley, P. E. & Lopez-Gallego, C. & Lindstrom, A. & Rich, S." box="[647,1053,1879,1905]" pageId="13" pageNumber="164" pagination="1 - 26" refId="ref20329" refString="Tang, W., Xu, G., O'Brien, C. W., Calonje, M., Franz, N. M., Johnston, M. A., Taylor, A., Vovides, A. P., Perez-Farrera, M. A., Salas-Morales, S. H., Lazcano-Lara, J. C., Skelley, P. E., Lopez-Gallego, C., Lindstrom, A. & Rich, S. (2018) Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity, 10 (38), 1 - 26. https: // doi. org / 10.3390 / d 10020038" type="journal article" year="2018">Tang, Skelley & Pérez-Farrera 2018</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. Deposited in the FSCA.
|
||||
<typeStatus id="AE498863FFB89515FAE9F8DCD7F05C71" box="[1335,1436,1879,1904]" pageId="13" pageNumber="164" type="allotype">
|
||||
<emphasis id="4386EAD3FFB89515FAE9F8DCD7F05C71" bold="true" box="[1335,1436,1879,1904]" pageId="13" pageNumber="164">Allotype</emphasis>
|
||||
</typeStatus>
|
||||
(FSCA) same data as
|
||||
<typeStatus id="AE498863FFB89515FE73F8F7D0635C94" box="[429,527,1916,1941]" pageId="13" pageNumber="164" type="holotype">holotype</typeStatus>
|
||||
and
|
||||
<specimenCount id="67F4FD48FFB89515FD89F8F7D0A55C94" box="[599,713,1916,1941]" pageId="13" pageNumber="164" type="adult">644 adult</specimenCount>
|
||||
<typeStatus id="AE498863FFB89515FD07F8F7D1235C94" box="[729,847,1916,1941]" pageId="13" pageNumber="164" type="paratype">
|
||||
<emphasis id="4386EAD3FFB89515FD07F8F7D1235C94" bold="true" box="[729,847,1916,1941]" pageId="13" pageNumber="164">paratypes</emphasis>
|
||||
</typeStatus>
|
||||
:
|
||||
<emphasis id="4386EAD3FFB89515FCB8F8F7D63B5C94" bold="true" box="[870,1111,1916,1941]" pageId="13" pageNumber="164">
|
||||
<collectingCountry id="09E57651FFB89515FCB8F8F7D18D5C94" box="[870,993,1916,1941]" name="Mexico" pageId="13" pageNumber="164">MEXICO</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="B336F823FFB89515FC2BF8F7D63B5C94" box="[1013,1111,1916,1941]" country="Mexico" name="Chiapas" pageId="13" pageNumber="164">Chiapas</collectingRegion>
|
||||
</emphasis>
|
||||
, La Sombra, Villa Flores,
|
||||
<taxonomicName id="B6F24D42FFB89515FF49F82AD3F25CB9" box="[151,414,1952,1976]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="13" pageNumber="164" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFB89515FF49F82AD3F25CB9" box="[151,414,1952,1976]" italics="true" pageId="13" pageNumber="164">Ceratozamia mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFB89515FE6EF82BD0705CB9" box="[432,540,1952,1976]" pageId="13" pageNumber="164" value="1994-05-01">1-V-1994</date>
|
||||
, A. Vovides, 31A (3); Ejido La Sombra, Mun. de Villa Flores,
|
||||
<taxonomicName id="B6F24D42FFB89515FAD5F82AD36E5CDD" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="13" pageNumber="164" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFB89515FAD5F82AD36E5CDD" italics="true" pageId="13" pageNumber="164">Ceratozamia mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFB89515FECCF84FD3135CDD" box="[274,383,1988,2012]" pageId="13" pageNumber="164" value="1994-05-01">1-V-1994</date>
|
||||
, M.A. Perez-Farrera (4); Mpio Cintalpa, Reserva La Sepaltura,
|
||||
<specimenCount id="67F4FD48FFB89515FB83F84FD6C85CDC" box="[1117,1188,1988,2013]" pageId="13" pageNumber="164" type="generic">
|
||||
<quantity id="B60A9B24FFB89515FB83F84FD6CD5CDC" box="[1117,1185,1988,2013]" metricMagnitude="2" metricUnit="m" metricValue="9.35" pageId="13" pageNumber="164" unit="m" value="935.0">935m</quantity>
|
||||
.
|
||||
</specimenCount>
|
||||
alt., ex.
|
||||
<taxonomicName id="B6F24D42FFB89515FAD5F84FD7F05CDA" box="[1291,1436,1988,2011]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="13" pageNumber="164" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFB89515FAD5F84FD7F05CDA" box="[1291,1436,1988,2011]" italics="true" pageId="13" pageNumber="164">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
nsp.,
|
||||
<date id="054C1001FFBB9516FF09FF1CD35A5BB1" box="[215,310,151,176]" pageId="14" pageNumber="165" value="1996-03">III-1996</date>
|
||||
, M.A.Perez (15); Municipio Cintalpa, Rancho Cafetal,
|
||||
<taxonomicName id="B6F24D42FFBB9516FC1EFF13D6D15BB1" authorityName="Perez-Farrera, Vovides & Iglesias" authorityYear="1999" box="[960,1213,152,176]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="alvarezii">
|
||||
<emphasis id="4386EAD3FFBB9516FC1EFF13D6D15BB1" box="[960,1213,152,176]" italics="true" pageId="14" pageNumber="165">Ceratozamia alvarezii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<date id="054C1001FFBB9516FB11FF13D72B5BB1" box="[1231,1351,151,176]" pageId="14" pageNumber="165" value="1996-03-04">4-III-1996</date>
|
||||
, M. A. Perez-Farrera (6); Ocotones, Mpio. De Cintalapa,
|
||||
<taxonomicName id="B6F24D42FFBB9516FD09FF37D1BA5BD5" authorityName="D. W. Stevenson" authorityYear="1982" box="[727,982,188,212]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="norstogii">
|
||||
<emphasis id="4386EAD3FFBB9516FD09FF37D1BA5BD5" box="[727,982,188,212]" italics="true" pageId="14" pageNumber="165">Ceratozamia norstogii</emphasis>
|
||||
</taxonomicName>
|
||||
<specimenCount id="67F4FD48FFBB9516FC3EFF36D1995BD5" box="[992,1013,189,212]" pageId="14" pageNumber="165" type="male">♂</specimenCount>
|
||||
cone,
|
||||
<date id="054C1001FFBB9516FB9DFF37D6DE5BD5" box="[1091,1202,188,213]" pageId="14" pageNumber="165" value="2008-02">Feb 2008</date>
|
||||
, M.A. Pérez Farrera (1); Mpo. Cintalapa, Ejido Flor de
|
||||
<collectingRegion id="B336F823FFBB9516FDFAFF54D0ED5BF9" box="[548,641,223,248]" country="Mexico" name="Chiapas" pageId="14" pageNumber="165">Chiapas</collectingRegion>
|
||||
, ex
|
||||
<taxonomicName id="B6F24D42FFBB9516FD6EFF6BD1C25BF6" authorityName="D. W. Stevenson" authorityYear="1982" box="[688,942,224,247]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="norstogii">
|
||||
<emphasis id="4386EAD3FFBB9516FD6EFF6BD1C25BF6" box="[688,942,224,247]" italics="true" pageId="14" pageNumber="165">Ceratozamia norstogii</emphasis>
|
||||
</taxonomicName>
|
||||
<specimenCount id="67F4FD48FFBB9516FC68FF6BD1A75BF6" box="[950,971,224,247]" pageId="14" pageNumber="165" type="male">♂</specimenCount>
|
||||
cone;
|
||||
<date id="054C1001FFBB9516FBC6FF54D6E95BF9" box="[1048,1157,223,248]" pageId="14" pageNumber="165" value="2010-02">Feb 2010</date>
|
||||
, M.A. Perez Farrera (5); Mpo. Villa Flores, Ejido La Sombra,
|
||||
<taxonomicName id="B6F24D42FFBB9516FDB2FE8FD11B5A1D" box="[620,887,260,284]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFBB9516FDB2FE8FD11B5A1D" box="[620,887,260,284]" italics="true" pageId="14" pageNumber="165">Ceratozamia mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
<specimenCount id="67F4FD48FFBB9516FC59FE8ED1F05A1D" box="[903,924,261,284]" pageId="14" pageNumber="165" type="male">♂</specimenCount>
|
||||
cone,
|
||||
<date id="054C1001FFBB9516FC29FE8FD6345A1D" box="[1015,1112,260,284]" pageId="14" pageNumber="165" value="2010-11">XI-2010</date>
|
||||
, M.A. Perez Farrera (39); Champerico, [GPS coord. omitted],
|
||||
<quantity id="B60A9B24FFBB9516FDEAFEA3D01B5A41" box="[564,631,296,320]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="14" pageNumber="165" unit="m" value="980.0">980m</quantity>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFBB9516FD5AFEA3D1E55A41" box="[644,905,296,320]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFBB9516FD5AFEA3D1E55A41" box="[644,905,296,320]" italics="true" pageId="14" pageNumber="165">Ceratozamia mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
<specimenCount id="67F4FD48FFBB9516FC4FFEA3D1CA5A3E" box="[913,934,296,319]" pageId="14" pageNumber="165" type="male">♂</specimenCount>
|
||||
cone,
|
||||
<date id="054C1001FFBB9516FC2AFEA3D6035A41" box="[1012,1135,296,320]" pageId="14" pageNumber="165" value="2017-05-20">
|
||||
20-
|
||||
<bibRefCitation id="15634B30FFBB9516FBC6FEA3D6035A41" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[1048,1135,296,320]" pageId="14" pageNumber="165" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">V-2017</bibRefCitation>
|
||||
</date>
|
||||
, H. Gomez Dominguez & W. Tang (153); Jaltenango, [GPS coord. omitted],
|
||||
<quantity id="B60A9B24FFBB9516FD10FEC7D17D5A64" box="[718,785,332,357]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="14" pageNumber="165" unit="m" value="957.0">957m</quantity>
|
||||
, ex
|
||||
<specimenCount id="67F4FD48FFBB9516FCE1FEC6D1385A65" box="[831,852,333,356]" pageId="14" pageNumber="165" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFBB9516FC85FEC7D63A5A65" box="[859,1110,332,356]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFBB9516FC85FEC7D63A5A65" box="[859,1110,332,356]" italics="true" pageId="14" pageNumber="165">Ceratozamia vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
cone,
|
||||
<date id="054C1001FFBB9516FB7CFEC7D7715A65" box="[1186,1309,332,356]" pageId="14" pageNumber="165" value="2017-05-21">
|
||||
21-
|
||||
<bibRefCitation id="15634B30FFBB9516FB1AFEC7D7715A65" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[1220,1309,332,356]" pageId="14" pageNumber="165" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">V-2017</bibRefCitation>
|
||||
</date>
|
||||
, H. Gomez Dominguez & W. Tang (417); La Concordia, Rancho Los Pinos, ex
|
||||
<specimenCount id="67F4FD48FFBB9516FC6FFEFBD1AA5A86" box="[945,966,368,391]" pageId="14" pageNumber="165" type="male">♂</specimenCount>
|
||||
<taxonomicName id="B6F24D42FFBB9516FC0EFEFBD6A35A89" box="[976,1231,368,392]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFBB9516FC0EFEFBD6A35A89" box="[976,1231,368,392]" italics="true" pageId="14" pageNumber="165">Ceratozamia vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
cone,
|
||||
<date id="054C1001FFBB9516FAC1FEFBD7F45A89" box="[1311,1432,368,392]" pageId="14" pageNumber="165" value="2017-06-01">
|
||||
1-
|
||||
<bibRefCitation id="15634B30FFBB9516FAEEFEFBD7F45A89" author="Valencia-Montoya, W. A. & Tuberquia, D. & Guzman, P. A. & Cardona-Duque, J." box="[1328,1432,368,392]" pageId="14" pageNumber="165" pagination="717 - 729" refId="ref20525" refString="Valencia-Montoya, W. A., Tuberquia, D., Guzman, P. A. & Cardona-Duque, J. (2017) Pollination of the cycad Zamia incognita A. Lindstr. and Idarraga by Pharaxonotha beetles in the Magdalena Medio Valley, Colombia: A mutualism dependent on a specific pollinator and its significance for conservation. Arthropod-Plant Interactions, 11, 717 - 729. https: // doi. org / 10.1007 / s 11829 - 017 - 9511 - y" type="journal article" year="2017">VI-2017</bibRefCitation>
|
||||
</date>
|
||||
, M.A. Pérez Farrera & H. Gómez Domínguez (2).
|
||||
<typeStatus id="AE498863FFBB9516FD00FE1ED1275AAC" box="[734,843,405,429]" pageId="14" pageNumber="165" type="paratype">Paratypes</typeStatus>
|
||||
deposited at ANIC, BMNH, FSCA, IEXA, INBio, MNHN, NZAC, UNAM, USNM.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBB9516FF19FE57D7185919" pageId="14" pageNumber="165" type="etymology">
|
||||
<paragraph id="714D36C1FFBB9516FF19FE57D7185919" blockId="14.[151,1436,151,968]" pageId="14" pageNumber="165">
|
||||
<emphasis id="4386EAD3FFBB9516FF19FE57D3225AF4" bold="true" box="[199,334,476,501]" pageId="14" pageNumber="165">Etymology.</emphasis>
|
||||
The species epithet comes from the Latin ‘piceus’ for black or dark, and ‘pennis’ for wing, in combination referring to the relatively darkened elytra of this species compared to other members of the genus.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="39E8654AFFBB9516FF19FDAFD7E158C9" pageId="14" pageNumber="165" type="discussion">
|
||||
<paragraph id="714D36C1FFBB9516FF19FDAFD7E158C9" blockId="14.[151,1436,151,968]" pageId="14" pageNumber="165">
|
||||
<emphasis id="4386EAD3FFBB9516FF19FDAFD357593C" bold="true" box="[199,315,548,573]" pageId="14" pageNumber="165">Remarks.</emphasis>
|
||||
This beetle has been collected from 4 closely related species of
|
||||
<taxonomicName id="B6F24D42FFBB9516FBF7FDAFD6D6593A" box="[1065,1210,548,571]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBB9516FBF7FDAFD6D6593A" box="[1065,1210,548,571]" italics="true" pageId="14" pageNumber="165">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
found in the Sierra Madre de
|
||||
<collectingRegion id="B336F823FFBB9516FECEFDCCD3065961" box="[272,362,583,608]" country="Mexico" name="Chiapas" pageId="14" pageNumber="165">Chiapas</collectingRegion>
|
||||
in
|
||||
<collectingRegion id="B336F823FFBB9516FE4BFDCCD3835961" box="[405,495,583,608]" country="Mexico" name="Chiapas" pageId="14" pageNumber="165">Chiapas</collectingRegion>
|
||||
State, including
|
||||
<taxonomicName id="B6F24D42FFBB9516FD66FDC3D1515961" authorityName="Perez-Farrera, Vovides & Iglesias" authorityYear="1999" box="[696,829,584,608]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="alvarezii">
|
||||
<emphasis id="4386EAD3FFBB9516FD66FDC3D0A7595E" box="[696,715,584,607]" italics="true" pageId="14" pageNumber="165">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFBB9516FD02FDC3D1515961" box="[732,829,584,608]" italics="true" pageId="14" pageNumber="165">alvarezii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFBB9516FC91FDC3D1B35961" box="[847,991,584,608]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="mirandae">
|
||||
<emphasis id="4386EAD3FFBB9516FC91FDC3D10E595E" box="[847,866,584,607]" italics="true" pageId="14" pageNumber="165">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFBB9516FCAAFDC3D1B35961" box="[884,991,584,608]" italics="true" pageId="14" pageNumber="165">mirandae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B6F24D42FFBB9516FC2FFDC3D616595E" authorityName="D. W. Stevenson" authorityYear="1982" box="[1009,1146,584,607]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="norstogii">
|
||||
<emphasis id="4386EAD3FFBB9516FC2FFDC3D668595E" box="[1009,1028,584,607]" italics="true" pageId="14" pageNumber="165">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFBB9516FBCBFDC3D616595E" box="[1045,1146,584,607]" italics="true" pageId="14" pageNumber="165">norstogii</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="B6F24D42FFBB9516FB66FDC3D72D5961" box="[1208,1345,584,608]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="vovidesii">
|
||||
<emphasis id="4386EAD3FFBB9516FB66FDC3D6A7595E" box="[1208,1227,584,607]" italics="true" pageId="14" pageNumber="165">C</emphasis>
|
||||
.
|
||||
<emphasis id="4386EAD3FFBB9516FB03FDC3D72D5961" box="[1245,1345,584,608]" italics="true" pageId="14" pageNumber="165">vovidesii</emphasis>
|
||||
</taxonomicName>
|
||||
(Pérez- Farrera
|
||||
<emphasis id="4386EAD3FFBB9516FF30FDE6D34A5985" box="[238,294,620,644]" italics="true" pageId="14" pageNumber="165">et al.</emphasis>
|
||||
2004, 2007). One other possible host species in this relatively isolated mountain system,
|
||||
<taxonomicName id="B6F24D42FFBB9516FAD5FDE7D33759A9" authority="Lundell" authorityName="Lundell" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="matudae">
|
||||
<emphasis id="4386EAD3FFBB9516FAD5FDE7D29459A9" italics="true" pageId="14" pageNumber="165">Ceratozamia matudae</emphasis>
|
||||
Lundell
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="15634B30FFBB9516FEB0FD1BD0F759A9" author="Perez-Farrera, M. A. & Quintana-Ascencio, P. F. & Salvatierra-Izaba, B. & Vovides, A. P." box="[366,667,656,680]" pageId="14" pageNumber="165" pagination="291 - 299" refId="ref19757" refString="Perez-Farrera, M. A., Quintana-Ascencio, P. F., Salvatierra-Izaba, B. & Vovides, A. P. (2000) Population dynamic of Ceratozamia matudai in el Triunfo Biosphere Reserve, Chiapas, Mexico. Journal of the Torrey Botanical Society, 127, 291 - 299. https: // doi. org / 10.2307 / 3088647" type="journal article" year="2000">
|
||||
Pérez-Farrera
|
||||
<emphasis id="4386EAD3FFBB9516FDCBFD1AD02059A9" box="[533,588,656,680]" italics="true" pageId="14" pageNumber="165">et al</emphasis>
|
||||
. 2000
|
||||
</bibRefCitation>
|
||||
), has not been sampled. The ranges of these host species are in relatively close proximity and all are dwarf species that have small male cones compared with other
|
||||
<taxonomicName id="B6F24D42FFBB9516FAD5FD3FD7F059CA" box="[1291,1436,692,715]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBB9516FAD5FD3FD7F059CA" box="[1291,1436,692,715]" italics="true" pageId="14" pageNumber="165">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
species located in nearby, but disjunct regions of
|
||||
<collectingRegion id="B336F823FFBB9516FD3BFD5CD12E59F1" box="[741,834,727,752]" country="Mexico" name="Chiapas" pageId="14" pageNumber="165">Chiapas</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="B336F823FFBB9516FC8CFD53D1CA59F1" box="[850,934,728,752]" country="Mexico" name="Oaxaca" pageId="14" pageNumber="165">Oaxaca</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="B336F823FFBB9516FC38FD53D62159EE" box="[998,1101,728,751]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="14" pageNumber="165">Veracruz</collectingRegion>
|
||||
. The elongated pronotum of
|
||||
<taxonomicName id="B6F24D42FFBB9516FF49FD77D3F55815" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[151,409,764,788]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBB9516FF49FD77D3F55815" box="[151,409,764,788]" italics="true" pageId="14" pageNumber="165">Ceratophila picipennis</emphasis>
|
||||
</taxonomicName>
|
||||
may be an adaptation to the small cones in which they inhabit. The adults of
|
||||
<taxonomicName id="B6F24D42FFBB9516FAD9FD77D7F05815" authorityName="Tang, Skelley & Perez-Farrera" authorityYear="2018" box="[1287,1436,764,788]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="species" species="picipennis">
|
||||
<emphasis id="4386EAD3FFBB9516FAD9FD77D7F05815" box="[1287,1436,764,788]" italics="true" pageId="14" pageNumber="165">C. picipennis</emphasis>
|
||||
</taxonomicName>
|
||||
may occur in male cones by the hundreds, especially in male cones just beginning their elongation and pollen shedding. They typically account for the second highest percentage of adult pharaxonothine beetles on male cones sampled during cone elongation and pollen shed (range 29–42%, mean = 38%, n = 3 populations;
|
||||
<taxonomicName id="B6F24D42FFBB9516FB22FCE3D7F05881" authorityName="Reitter" authorityYear="1875" box="[1276,1436,872,896]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4386EAD3FFBB9516FB22FCE3D7F05881" box="[1276,1436,872,896]" italics="true" pageId="14" pageNumber="165">Pharaxonotha</emphasis>
|
||||
</taxonomicName>
|
||||
beetles usually outnumber them), and the most abundant of the two species of
|
||||
<taxonomicName id="B6F24D42FFBB9516FBD6FC07D6E358A5" box="[1032,1167,908,932]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
|
||||
<emphasis id="4386EAD3FFBB9516FBD6FC07D6E358A5" box="[1032,1167,908,932]" italics="true" pageId="14" pageNumber="165">Ceratophila</emphasis>
|
||||
</taxonomicName>
|
||||
typically found in male cones.
|
||||
<taxonomicName id="B6F24D42FFBB9516FF3AFC3BD00A58C9" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[228,614,944,968]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
|
||||
<emphasis id="4386EAD3FFBB9516FF3AFC3BD30758C9" box="[228,363,944,968]" italics="true" pageId="14" pageNumber="165">Ceratophila</emphasis>
|
||||
(
|
||||
<emphasis id="4386EAD3FFBB9516FEA2FC3BD3B158C9" box="[380,477,944,968]" italics="true" pageId="14" pageNumber="165">Vovidesa</emphasis>
|
||||
)
|
||||
<emphasis id="4386EAD3FFBB9516FE2EFC3BD00A58C9" box="[496,614,944,968]" italics="true" pageId="14" pageNumber="165">chiapensis</emphasis>
|
||||
</taxonomicName>
|
||||
coinhabit the same
|
||||
<taxonomicName id="B6F24D42FFBB9516FC98FC3BD1BB58C6" box="[838,983,944,967]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="165" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="4386EAD3FFBB9516FC98FC3BD1BB58C6" box="[838,983,944,967]" italics="true" pageId="14" pageNumber="165">Ceratozamia</emphasis>
|
||||
</taxonomicName>
|
||||
cones, but usually in smaller numbers.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
146
data/F9/5B/CA/F95BCAF6217839C4DAC1C7D2420D8F48.xml
Normal file
146
data/F9/5B/CA/F95BCAF6217839C4DAC1C7D2420D8F48.xml
Normal file
|
|
@ -0,0 +1,146 @@
|
|||
<document ID-DOI="http://dx.doi.org/10.3897/jhr.53.10217" ID-GBIF-Dataset="f3e2e8b3-5aef-4a00-9784-ec336edc9ebc" ID-Pensoft-Pub="1314-2607-53-1" ID-Pensoft-UUID="255935082C11B238FF99FFF1FFC4FF9E" ID-ZBK="13184D6320474F62A987B844E6386BCD" ID-Zenodo-Dep="575134" ID-ZooBank="13184D6320474F62A987B844E6386BCD" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2607-53-1" ModsDocOrigin="Journal of Hymenoptera Research 53" ModsDocTitle="Revision of New World Helava Masner & Huggert (Platygastridae, Sceliotrachelinae)" checkinTime="1482179232747" checkinUser="pensoft" docAuthor="Talamas, Elijah J. & Masner, Lubomir" docDate="2016" docId="F95BCAF6217839C4DAC1C7D2420D8F48" docLanguage="en" docName="JourHymenoptRes 53: 1-24" docOrigin="Journal of Hymenoptera Research 53" docPubDate="2016-12-19" docSource="http://dx.doi.org/10.3897/jhr.53.10217" docTitle="Helava reducta Masner & Talamas 2016, sp. n." docType="treatment" docUuid="908D245E-2CF2-495B-B711-87CB4D6AB3EE" docUuidSource="ZooBank" docVersion="5" id="255935082C11B238FF99FFF1FFC4FF9E" lastPageNumber="19" masterDocId="255935082C11B238FF99FFF1FFC4FF9E" masterDocTitle="Revision of New World Helava Masner & Huggert (Platygastridae, Sceliotrachelinae)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="16" updateTime="1678749293656" updateUser="pensoft">
|
||||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo>
|
||||
<mods:title>Revision of New World Helava Masner & Huggert (Platygastridae, Sceliotrachelinae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Talamas, Elijah J.</mods:namePart>
|
||||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1048-6345</mods:nameIdentifier>
|
||||
<mods:affiliation>Systematic Entomology Laboratory, USDA / ARS c / o USNM, Smithsonian Institution, Washington, D. C. 20560, U. S. A.</mods:affiliation>
|
||||
<mods:nameIdentifier type="email">billy.jenkins@GMAIL.COM</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Masner, Lubomir</mods:namePart>
|
||||
<mods:affiliation>Agriculture and Agri-Food Canada, K. W. Neatby Building, Ottawa, Ontario K 1 A 0 C 6, Canada</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:typeOfResource>text</mods:typeOfResource>
|
||||
<mods:relatedItem type="host">
|
||||
<mods:titleInfo>
|
||||
<mods:title>Journal of Hymenoptera Research</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part>
|
||||
<mods:date>2016</mods:date>
|
||||
<mods:detail type="pubDate">
|
||||
<mods:number>2016-12-19</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>53</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>1</mods:start>
|
||||
<mods:end>24</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location>
|
||||
<mods:url>http://dx.doi.org/10.3897/jhr.53.10217</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.53.10217</mods:identifier>
|
||||
<mods:identifier type="Pensoft-Pub">1314-2607-53-1</mods:identifier>
|
||||
<mods:identifier type="ZooBank">13184D6320474F62A987B844E6386BCD</mods:identifier>
|
||||
<mods:identifier type="Pensoft-UUID">255935082C11B238FF99FFF1FFC4FF9E</mods:identifier>
|
||||
<mods:identifier type="Zenodo-Dep">575134</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment ID-GBIF-Taxon="127889204" LSID="urn:lsid:zoobank.org:act:908D245E-2CF2-495B-B711-87CB4D6AB3EE" httpUri="http://treatment.plazi.org/id/F95BCAF6217839C4DAC1C7D2420D8F48" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16">
|
||||
<subSubSection pageId="15" pageNumber="16" type="nomenclature">
|
||||
<paragraph pageId="15" pageNumber="16">
|
||||
<taxonomicName LSID="http://zoobank.org/908D245E-2CF2-495B-B711-87CB4D6AB3EE" authority="Masner & Talamas" authorityName="Masner & Talamas" authorityYear="2016" class="Insecta" family="Platygastridae" genus="Helava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helava reducta" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="reducta" status="sp. n.">Helava reducta Masner & Talamas</taxonomicName>
|
||||
<taxonomicNameLabel pageId="15" pageNumber="16">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="16" pageNumber="17" type="description">
|
||||
<paragraph pageId="16" pageNumber="17">
|
||||
<figureCitation captionStart="Figures 45–49" captionStartId="F11" captionText="Figures 45 - 49. Helava reducta 45 female holotype (USNMENT 00989203), head, mesosoma, metasoma, lateral view 46 female holotype (USNMENT 00989203), head, mesosoma, metasoma, dorsal view 47 female holotype (USNMENT 00989203), head, anterior view 48 male paratype (USNMENT 00989204), antenna, dorsal view 49 female holotype (USNMENT 00989203), antenna dorsal view. Scale bars in millimeters." figureDoi="10.3897/jhr.53.10217.figures45-49" httpUri="https://binary.pensoft.net/fig/115464" pageId="16" pageNumber="17">
|
||||
<pageBreakToken pageId="16" pageNumber="17" start="start">Figures</pageBreakToken>
|
||||
45-49
|
||||
</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
|
||||
<paragraph pageId="16" pageNumber="17">
|
||||
<pageBreakToken pageId="16" pageNumber="17" start="start">Description</pageBreakToken>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
|
||||
Female body length: 1.01 mm (n=1). Male body length: 1.04 mm (n=1). Male antenna: apically clubbed. Number of antennomeres in male club: 3. Number of
|
||||
<pageBreakToken pageId="17" pageNumber="18" start="start">female</pageBreakToken>
|
||||
clavomeres: 4. Setation of frons anterior to ocellar triangle: sparse throughout. Setation of vertex posterior to lateral ocellus: very sparse or absent. Hyperoccipital carina: absent. Pronotum in dorsal view: large, collarlike. Dorsoventral band of dense setation on posterior part of lateral pronotum: absent. Setation of pronotal cervical sulcus: sparse to absent. Width of dorsal mesopleuron in lateral view: half as wide dorsally. Longitudinal striation on dorsal mesopleuron: absent. Transepisternal line: absent. Mesofurcal pit: present. Notaulus: absent. Wings: brachypterous. Color of legs: yellowish brown. Foamy structures of lateral propodeum: smaller than hairy metapleuron. Median tubercule on
|
||||
<pageBreakToken pageId="18" pageNumber="19" start="start">T</pageBreakToken>
|
||||
2: absent. Setation of anterior T2: interrupted medially. Lateral patch on T2: absent. Foamy structures on S1: absent. Transverse felt field on anterior S2: present as transverse strip. Shape of S2 in lateral view: broadly convex.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="18" pageNumber="19" type="diagnosis">
|
||||
<paragraph pageId="18" pageNumber="19">Diagnosis.</paragraph>
|
||||
<paragraph pageId="18" pageNumber="19">
|
||||
<taxonomicName authorityName="Masner & Talamas" authorityYear="2016" class="Insecta" family="Platygastridae" genus="Helava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helava reducta" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="reducta">
|
||||
<emphasis italics="true" pageId="18" pageNumber="19">Helava reducta</emphasis>
|
||||
</taxonomicName>
|
||||
can easily be identified by severe microptery, the absence of a scutoscutellar sulcus, and the absence of transepisternal line.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="18" pageNumber="19" type="etymology">
|
||||
<paragraph pageId="18" pageNumber="19">Etymology.</paragraph>
|
||||
<paragraph pageId="18" pageNumber="19">
|
||||
The Latin adjectival epithet "
|
||||
<taxonomicName authorityName="Masner & Talamas" authorityYear="2016" class="Insecta" family="Platygastridae" genus="Helava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helava reducta" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="reducta">Helava reducta</taxonomicName>
|
||||
" is applied to this species for the reduced segmentation of the mesosoma,
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="18" pageNumber="19" type="distribution">
|
||||
<paragraph pageId="18" pageNumber="19">Link to distribution map.</paragraph>
|
||||
<paragraph pageId="18" pageNumber="19">http://hol.osu.edu/map-large.html?id=354389</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="18" pageNumber="19" type="materials_examined">
|
||||
<paragraph pageId="18" pageNumber="19">Material examined.</paragraph>
|
||||
<paragraph pageId="18" pageNumber="19">
|
||||
<materialsCitation collectingDate="1981-04-29" collectorName="L. Masner" country="VENEZUELA" elevation="3500" location="Sierra Nevada National Park" municipality="Black Lagoon" specimenCount="1" stateProvince="Merida" typeStatus="Holotype">
|
||||
<typeStatus>Holotype</typeStatus>
|
||||
, female:
|
||||
<emphasis bold="true" pageId="18" pageNumber="19">
|
||||
<collectingCountry name="Venezuela">VENEZUELA</collectingCountry>
|
||||
</emphasis>
|
||||
:
|
||||
<normalizedToken originalValue="Mérida">
|
||||
<collectingRegion country="Venezuela" name="Merida">Merida</collectingRegion>
|
||||
</normalizedToken>
|
||||
St.,
|
||||
<collectingMunicipality>Black Lagoon</collectingMunicipality>
|
||||
,
|
||||
<location LSID="urn:lsid:plazi:treatment:F95BCAF6217839C4DAC1C7D2420D8F48:89F609B620C4DE5D6C679ECA4E1C44A9" country="VENEZUELA" municipality="Black Lagoon" name="Sierra Nevada National Park" stateProvince="Merida">Sierra Nevada National Park</location>
|
||||
,
|
||||
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.5" unit="m" value="3500.0">
|
||||
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.5" unit="m" value="3500.0">3500m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<collectingDate value="1981-04-29">29.IV.1981</collectingDate>
|
||||
, sweeping,
|
||||
<collectorName>L. Masner</collectorName>
|
||||
, USNMENT00989203 (deposited in CNCI)
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="VENEZUELA" location="VENEZUELA" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="Paratypes">
|
||||
<emphasis italics="true" pageId="18" pageNumber="19">
|
||||
<typeStatus>Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<emphasis bold="true" pageId="18" pageNumber="19">
|
||||
<collectingCountry name="Venezuela">VENEZUELA</collectingCountry>
|
||||
</emphasis>
|
||||
:
|
||||
<specimenCount type="female">1 female</specimenCount>
|
||||
,
|
||||
<specimenCount type="male">2 males</specimenCount>
|
||||
, CNC424744-424745, USNMENT00989204 (CNCI)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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