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<document ID-CLB-Dataset="291603" ID-GBIF-Dataset="9972c1a8-94d2-4800-bd5b-2e485f3409a8" ID-ISBN="978-3-258-08047-5" checkinTime="1706569717550" checkinUser="plazi" docAuthor="Konrad Lauber, Gerhart Wagner &amp; Andreas Gygax" docDate="2018" docId="F91ED8E749499D03560F461804EA8D8F" docLanguage="de" docName="FloraHelv.Lamiaceae.xml" docOrigin="Flora Helvetica, Bern: Haupt Verlag" docTitle="Stachys byzantina K. Koch" docType="treatment" docVersion="7" id="D7E2CFFFA139FA594D69F062A234698C" lastPageNumber="864" masterDocId="3DD339DDD9CC8C7489449EAD6D2ADFC8" masterDocTitle="Flora Helvetica - Lamiaceae" masterLastPageNumber="882" masterPageNumber="836" pageNumber="864" updateTime="1711164571443" updateUser="ExternalLinkService">
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<mods:titleInfo id="B8DBFFC9280337874D3844B1FA288070">
<mods:title id="406B48F8E0D88F029FE03F5F4DAAE4D9">Flora Helvetica - Lamiaceae</mods:title>
</mods:titleInfo>
<mods:name id="94DB20C44B9F35979F4DE898860F3A35" type="personal">
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<mods:namePart id="D40BA2779A2A9BF4F9378463E984EB51">Konrad Lauber</mods:namePart>
</mods:name>
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<mods:namePart id="836D8B29CEB4DADB41EC33B08A97D3B9">Gerhart Wagner</mods:namePart>
</mods:name>
<mods:name id="EDF7456327564FB3F228A7B0AC1B1745" type="personal">
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<mods:roleTerm id="F10FF82DBFDFE0D59610BDAF05C00228">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E9D4583D4E7E5E7A90EB922DE29236EB">Andreas Gygax</mods:namePart>
</mods:name>
<mods:typeOfResource id="6586825E42E82921E2CFA79D086ECF4D">text</mods:typeOfResource>
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<mods:dateIssued id="0D84816F3FB795C552C21A76CAC943E6">2018</mods:dateIssued>
<mods:publisher id="69568D549B227A3475BD7F22D41A7695">Haupt Verlag</mods:publisher>
<mods:place id="4C5A1E209FCC1C044260488583553E86">
<mods:placeTerm id="7B647FBB6587519B53231F99AFB67576">Bern</mods:placeTerm>
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<mods:titleInfo id="3A6FAADBC5B533B8DD7D1945195B0B4A">
<mods:title id="AD3FD8324CBA8E4D6DA1B689479CF975">Flora Helvetica</mods:title>
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<mods:classification id="BFF8AE71CBB990CFB68771378450A2FB">book chapter</mods:classification>
<mods:identifier id="E4B14C54DE128ECFE749BE29C37B475B" type="ISBN">978-3-258-08047-5</mods:identifier>
</mods:mods>
<treatment id="F91ED8E749499D03560F461804EA8D8F" ID-DOI="http://doi.org/10.5281/zenodo.10859633" ID-GBIF-Taxon="224054713" ID-Zenodo-Dep="10859633" LSID="urn:lsid:plazi:treatment:F91ED8E749499D03560F461804EA8D8F" httpUri="http://treatment.plazi.org/id/F91ED8E749499D03560F461804EA8D8F" lastPageNumber="864" pageNumber="864">
<subSubSection id="6E26721A82672E631BAA0D533DA46DF3" pageNumber="864" type="nomenclature">
<paragraph id="815CC1A2615E4C5749AA8990CD38C60E" pageNumber="864">
<taxonomicName id="2056C54175832416B0CE9FB2AC0C1F6B" ID-CoL="6ZGPC" authority="K. Koch" authorityName="K. Koch" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageNumber="856" phylum="Tracheophyta" rank="species" species="byzantina">
<emphasis id="A4E14E10E6973C2682FC233241766A36" italics="true" pageNumber="864">Stachys byzantina</emphasis>
K. Koch
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F77978E4C22C4E3D61EA4FCE4B830649" pageNumber="864" type="description">
<paragraph id="6302AC110F3E3C7E27683A9F4C1C5CD5" pageNumber="864">
Artbeschreibung:
<normalizedToken id="F27AD52AA58909E233F3DC31F6856630" originalValue="Ähnlich" pageNumber="856">Aehnlich</normalizedToken>
wie
<taxonomicName id="B92433F2AD11B2CF6B86ED7CF756FFFF" class="Magnoliopsida" family="Lamiaceae" genus="Stachys" kingdom="Plantae" order="Lamiales" pageNumber="856" phylum="Tracheophyta" rank="species" species="germanica">
<emphasis id="655669CDD51BAF633B03205C0E488221" italics="true" pageNumber="864">S. germanica</emphasis>
</taxonomicName>
, aber viel dichter behaart (Epidermis nicht sichtbar),
<emphasis id="65766EAB19D24F63EB8A1BB357034BE5" bold="true" pageNumber="864">
<normalizedToken id="A0B180E6483AC05207D3F4AAF51045A9" originalValue="Blätter" pageNumber="856">Blaetter</normalizedToken>
weiss, am Grund in den Stiel
<normalizedToken id="68F98F46383A423FBED04FBD3D5FF999" originalValue="verschmälert" pageNumber="856">verschmaelert</normalizedToken>
</emphasis>
, nicht
<normalizedToken id="AD68F38937C67D30D1F29F99F8909704" originalValue="herzförmig" pageNumber="856">herzfoermig</normalizedToken>
, fein
<normalizedToken id="EF430EDD51708324DBF9EB75C1E76062" originalValue="gezähnt" pageNumber="856">gezaehnt</normalizedToken>
bis fast ganzrandig, stark runzelig.
</paragraph>
</subSubSection>
<subSubSection id="EB3247C4049123686025FACCF814CFE5" pageNumber="864" type="biology_ecology">
<paragraph id="CA2A56DCF4E81C8FA171DFBCCE97AAC0" pageNumber="864">
<normalizedToken id="AF1F031E0D710302E643F420128BBC4A" originalValue="Blütezeit" pageNumber="856">Bluetezeit</normalizedToken>
: 7-8
</paragraph>
<paragraph id="6DBD862AAE415D27A04E714D3B580602" pageNumber="864">Standort und Verbreitung in der Schweiz: Angepflanzt und gelegentlich verwildert /</paragraph>
</subSubSection>
<subSubSection id="80321105957A7C6BDB8055026AD1C6CF" pageNumber="864" type="distribution">
<paragraph id="FF57CB43DFED4EB282AB0D2CD1E5A5C2" pageNumber="864">Verbreitung global: Ostmediterran</paragraph>
</subSubSection>
<subSubSection id="CCD33ADF5DC0BC1483288A284BBD6177" pageNumber="864" type="biology_ecology">
<paragraph id="2660554EBBCF16EF95256C6604D7FE5F" pageNumber="864">
<normalizedToken id="6D3F9B6EE9C8B45D115CE7A656797664" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="DCBAEC1EB8B70F441808B4F33766F241" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." pageNumber="864" publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="8C618F2D2A2ACAF9778E0376FBB6C6CF" pageNumber="864">
<table id="7106AFF96CB88E8562582AFAEAF8F1DC" class="table" inLine="true" pageNumber="864">
<tr id="A0EC19AC8C9562468BAAA273A694E777" pageNumber="864">
<th id="23B425A58CE18AD7652C9AABCEE5DE6A" colspan="2" pageNumber="864">
<emphasis id="A7C8E287C1F8C5F6E43BB27898EAA299" bold="true" pageNumber="864">Bodenfaktoren</emphasis>
</th>
<th id="399A0B2E5AD4CDFD90C3340C713C196F" colspan="2" pageNumber="864">
<emphasis id="6A5F8694544A123E73F62A9BF6693638" bold="true" pageNumber="864">Klimafaktoren</emphasis>
</th>
<th id="0C53D28B2495CD8877038E1CDF5B927E" colspan="2" pageNumber="864">
<emphasis id="E4938B7E4EBFB87F6176A1A20AE0D15B" bold="true" pageNumber="864">Salztoleranz</emphasis>
</th>
</tr>
<tr id="1A836BF400DFD84093CEA710D109A372" pageNumber="864">
<td id="7FEB109C8566C5F7E214E1346824A149" pageNumber="864">Feuchtezahl F</td>
<td id="4826C8195CEFF062E1743B73D0745A1C" pageNumber="864" style="text-align:left;width:100px">--</td>
<td id="681CB7E14B59C4709331D4AA4307E586" pageNumber="864">Lichtzahl L</td>
<td id="A3EEA9E080175E6D967118E49131FE7B" pageNumber="864" style="text-align:left;width:60px">--</td>
<td id="4278E8AF46F86171F1AE0C013BA02E87" pageNumber="864">Salzzeichen</td>
<td id="7A16AA905BE5D6F913E2E580490ED839" pageNumber="864" style="text-align:left;width:60px">--</td>
</tr>
<tr id="ED5AAD8C113C84F2E49C23F30DEF0037" pageNumber="864">
<td id="F99CD153F5E0FC8B99AFCCE827C28C6C" pageNumber="864">Reaktionszahl R</td>
<td id="2C84ADC08D1AC09A901CA434E7CA5134" pageNumber="864" style="text-align:left">--</td>
<td id="F1045F29F7D846759565C3F64A61AC98" pageNumber="864">Temperaturzahl T</td>
<td id="676D03D323FF4079E57724CF61279FDA" pageNumber="864" style="text-align:left">--</td>
</tr>
<tr id="6779B03733C895B1A0C85406C7B64FFD" pageNumber="864">
<td id="DE8BF1EF5461A64AA569CBAE53B59609" pageNumber="864">
<normalizedToken id="727131F78ABD3B65E5FE8CC2401180EF" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="1774537A285100C985171EF5B25FD01F" pageNumber="864" style="text-align:left">--</td>
<td id="FC87B5283F587839C99D733F0ED074B7" pageNumber="864">
<normalizedToken id="87F01AF96DD6CDD3549BD58F9C1B1602" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="0FC9B9E3D2C19FBBD8038CB22DCB0196" pageNumber="864" style="text-align:left">--</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="A871F8A6225E0C2D8A610B43A4BAC043" pageNumber="864" type="vernacular_names">
<paragraph id="00331DD35BB860E5075C5386F3088227" pageNumber="864">
Volksname Deutscher Name:
<vernacularName id="BDDBA9B8EA121896758D04BC4E3CE689" language="deu" pageNumber="864">Wolliger Ziest</vernacularName>
Nom
<normalizedToken id="76FF2686010B62EF5BC684A842C91FD3" originalValue="français">francais</normalizedToken>
:
<vernacularName id="F64D83FB6B582FF7E873CBECE25FF518" language="fra" pageNumber="864">Epiaire laineuse</vernacularName>
Nome italiano:
<vernacularName id="0FF02DC04B1753BC4B9CD9FF218CD7BC" language="ita" pageNumber="864">Stregona candida</vernacularName>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE0081028FFA0FE3ADEB69FF8698F" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus gilleti Janssens" docType="treatment" docVersion="8" lastPageNumber="147" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="146" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
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<mods:namePart>Frolov, Andrey V.</mods:namePart>
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<mods:date>2014</mods:date>
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<mods:number>2014-06-30</mods:number>
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<mods:detail type="volume">
<mods:number>49</mods:number>
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<mods:number>3</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331151" ID-GBIF-Taxon="167368591" ID-Zenodo-Dep="4331151" LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F" httpUri="http://treatment.plazi.org/id/F91EE0081028FFA0FE3ADEB69FF8698F" lastPageId="18" lastPageNumber="147" pageId="17" pageNumber="146">
<subSubSection box="[470,777,1253,1278]" pageId="17" pageNumber="146" type="nomenclature">
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<heading box="[470,777,1253,1278]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="146" reason="2">
<taxonomicName authority="Janssens" authorityName="Janssens" box="[470,777,1253,1278]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="146" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis bold="true" box="[470,672,1253,1277]" italics="true" pageId="17" pageNumber="146">Delopleurus gilleti</emphasis>
Janssens
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</heading>
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(
<figureCitation box="[550,698,1286,1310]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="17" pageNumber="146">Figure 5A–F</figureCitation>
)
</paragraph>
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<paragraph blockId="17.[143,777,1253,1353]" box="[143,511,1328,1353]" pageId="17" pageNumber="146">
<taxonomicName authority="Janssens 1939" authorityName="Janssens" authorityYear="1939" box="[143,511,1328,1353]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="146" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis box="[143,345,1329,1353]" italics="true" pageId="17" pageNumber="146">Delopleurus gilleti</emphasis>
<bibRefCitation author="Janssens A" box="[354,511,1328,1352]" pageId="17" pageNumber="146" pagination="1 - 104" refId="ref10112" refString="Janssens A. 1939. Coprini. Exploration du Parc National Albert, Mission GF de Witte. Institut des parcs nationaux du Congo Belge. 29: 1 - 104." type="journal article" year="1939">Janssens 1939</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="146" type="diagnosis">
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<emphasis box="[144,252,1395,1418]" italics="true" pageId="17" pageNumber="146">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[144,1104,1395,1553]" pageId="17" pageNumber="146">
This species is most similar to male
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[556,656,1434,1458]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="146" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[556,656,1434,1458]" italics="true" pageId="17" pageNumber="146">D. pullus</emphasis>
</taxonomicName>
but can be separated from it in having the pygidium with smaller disc which is punctate with bigger punctures (
<figureCitation box="[951,1095,1466,1490]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="17" pageNumber="146">Figure 5B, E</figureCitation>
) and in having relatively densely punctate disc of pronotum (
<figureCitation box="[864,1026,1498,1522]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="17" pageNumber="146">Figure 5A, D</figureCitation>
). This species does not show sexual dimorphism in the sculpture of the pygidium.
</paragraph>
</subSubSection>
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<heading box="[144,271,1628,1651]" fontSize="10" level="3" pageId="17" pageNumber="146" reason="5">
<emphasis box="[144,271,1628,1651]" italics="true" pageId="17" pageNumber="146">Description</emphasis>
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<paragraph blockId="17.[144,1104,1628,1691]" lastBlockId="18.[212,1172,168,670]" lastPageId="18" lastPageNumber="147" pageId="17" pageNumber="146">
Beetles are small-sized (3.5–5.0 mm), strongly convex, black or dark brown, glabrous.
<emphasis box="[212,270,168,192]" italics="true" pageId="18" pageNumber="147">Male</emphasis>
(
<figureCitation box="[291,412,168,192]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="18" pageNumber="147">Figure 5A</figureCitation>
). Clypeus quadridentate, medial teeth acute, lateral ones acuteangled. Clypeal margin laterad of lateral teeth somewhat angulate so clypeus looks hexadentate. Clypeal surface rugose in anterior part and laterally, frons densely punctate (punctures separated by less than puncture diameters).
</paragraph>
<paragraph blockId="18.[212,1172,168,670]" pageId="18" pageNumber="147">Pronotum about two times wider than long. Anterior and lateral margins distinctly bordered, base not bordered. Pronotum densely almost uniformly punctate with punctures separated by less than a puncture diameter.</paragraph>
<paragraph blockId="18.[212,1172,168,670]" pageId="18" pageNumber="147">Elytra trapezoidal, as wide as long, shiny. Stria relatively deep, with punctures slightly larger than striae width. Elytral intervals convex, with minute punctuation.</paragraph>
<paragraph blockId="18.[212,1172,168,670]" pageId="18" pageNumber="147">Anterior tibiae with three outer teeth, without a small acute tooth between first outer tooth and apical spur. Anterior tibial spur bifurcated apically.</paragraph>
<paragraph blockId="18.[212,1172,168,670]" pageId="18" pageNumber="147">
Pygidium with relatively narrow borders and feebly convex disc. Disc glabrous, coarsely punctate with punctures separated by about one puncture diameter (
<figureCitation box="[220,334,583,607]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="18" pageNumber="147">Figure 5B</figureCitation>
).
</paragraph>
<paragraph blockId="18.[212,1172,168,670]" pageId="18" pageNumber="147">
Parameres in lateral view somewhat abruptly truncate, with distinct processes (
<figureCitation box="[220,335,646,670]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="18" pageNumber="147">Figure 5C</figureCitation>
).
</paragraph>
<paragraph blockId="18.[212,1172,710,766]" pageId="18" pageNumber="147">
<emphasis box="[212,292,711,735]" italics="true" pageId="18" pageNumber="147">Female</emphasis>
(
<figureCitation box="[308,425,710,734]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="18" pageNumber="147">Figure 5D</figureCitation>
) can be separated from male in having acute, not bifurcated spurs of anterior tibiae. Sculpture of pygidium is similar to that of male (
<figureCitation box="[982,1096,742,766]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="18" pageNumber="147">Figure 5E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="147" type="materials_examined">
<paragraph blockId="18.[212,1172,830,1053]" box="[212,368,830,854]" pageId="18" pageNumber="147">
<heading box="[212,368,830,854]" fontSize="10" level="3" pageId="18" pageNumber="147" reason="5">
<emphasis box="[212,368,830,854]" italics="true" pageId="18" pageNumber="147">Type material</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[212,1172,830,1053]" pageId="18" pageNumber="147">
<materialsCitation ID-GBIF-Occurrence="2849122341" collectingDate="1933" collectingDateMax="1935" collectingDateMin="1933" collectionCode="IRSNB" determinerName="A. Janssens" pageId="18" pageNumber="147" specimenCount="1" typeStatus="holotype">
<typeStatus box="[212,321,870,894]" pageId="18" pageNumber="147">Holotype</typeStatus>
, female with the labels. ‘
<collectorName box="[609,1120,870,895]" pageId="18" pageNumber="147">Coll. R.I.Sc.N.B Togo Togo Coll. J.J.Gillet’</collectorName>
, ‘
<determinerName pageId="18" pageNumber="147">A. Janssens</determinerName>
det., 1939
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[439,648,902,926]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="147" phylum="Arthropoda" rank="species" species="gilleti" status="n. sp.">Delopleurus gilleti</taxonomicName>
<taxonomicNameLabel box="[659,722,902,925]" pageId="18" pageNumber="147" rank="species">n. sp.</taxonomicNameLabel>
type’, ‘cf.
<collectorName box="[849,1171,902,926]" pageId="18" pageNumber="147">Expl. P.N.A. G.F. de Witte</collectorName>
(
<date box="[219,341,934,958]" pageId="18" pageNumber="147" value="1933" valueMax="1935">
<collectingDate box="[219,341,934,958]" pageId="18" pageNumber="147" value="1933" valueMax="1935">1933–1935</collectingDate>
</date>
) fasc 29, 1939, p. 130’, ‘TYPE’, and ‘
<determinerName box="[798,932,934,958]" pageId="18" pageNumber="147">A. Janssens</determinerName>
det., 1941 gilleti
<collectorName pageId="18" pageNumber="147">A. Janssens’</collectorName>
(
<collectionCode box="[333,422,965,989]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="18" pageNumber="147">IRSNB</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122314" collectingDate="1933" collectingDateMax="1935" collectingDateMin="1933" collectionCode="IRSNB" collectorName="de Luebo, G" determinerName="A. Janssens" pageId="18" pageNumber="147" specimenCount="2" stateProvince="Bas-Congo" typeStatus="paratype">
<typeStatus box="[445,559,966,990]" pageId="18" pageNumber="147">Paratypes</typeStatus>
: male and female with the same locality data as the
<typeStatus box="[212,310,997,1021]" pageId="18" pageNumber="147">holotype</typeStatus>
(
<collectionCode box="[327,415,997,1021]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="18" pageNumber="147">IRSNB</collectionCode>
); female,
<collectingRegion box="[527,631,997,1021]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="18" pageNumber="147">CONGO</collectingRegion>
(
<collectingRegion box="[648,757,997,1021]" country="Democratic Republic of the Congo" name="Kinshasa" pageId="18" pageNumber="147">Kinshasa</collectingRegion>
),
<collectingRegion box="[777,845,997,1021]" country="Democratic Republic of the Congo" name="Kasai-Oriental" pageId="18" pageNumber="147">Kasai</collectingRegion>
, forêt
<collectorName box="[921,1070,997,1021]" pageId="18" pageNumber="147">de Luebo, G</collectorName>
. Babault (
<collectionCode box="[220,309,1029,1053]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="18" pageNumber="147">IRSNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="18.[212,1172,1117,1532]" box="[212,427,1117,1141]" pageId="18" pageNumber="147">
<heading box="[212,427,1117,1141]" fontSize="10" level="3" pageId="18" pageNumber="147" reason="5">
<emphasis box="[212,427,1117,1141]" italics="true" pageId="18" pageNumber="147">Additional material</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[212,1172,1117,1532]" pageId="18" pageNumber="147">
<materialsCitation ID-GBIF-Occurrence="2849122321" collectingDate="1980-09" collectionCode="MNHN" collectorName="Y. Cambefort" county="Fitini" location="National Park De La Comoe" municipality="Quango" pageId="18" pageNumber="147" specimenCount="1" specimenCount-female="1">
CÔTE D’ IVOIRE:
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FE2FDED69D686B89" box="[451,814,1157,1181]" county="Fitini" municipality="Quango" name="National Park De La Comoe" pageId="18" pageNumber="147">National Park De La Comoe</location>
,
<collectingMunicipality box="[833,926,1157,1181]" pageId="18" pageNumber="147">Quango</collectingMunicipality>
,
<collectingCounty box="[946,1011,1157,1181]" pageId="18" pageNumber="147">Fitini</collectingCounty>
,
<date box="[1031,1124,1157,1181]" pageId="18" pageNumber="147" value="1980-09">
<collectingDate box="[1031,1124,1157,1181]" pageId="18" pageNumber="147" value="1980-09">IX.1980</collectingDate>
</date>
,
<collectorName pageId="18" pageNumber="147">Y. Cambefort</collectorName>
leg.,
<specimenCount box="[410,512,1189,1213]" pageId="18" pageNumber="147" type="female">1 female</specimenCount>
(
<collectionCode box="[535,631,1189,1213]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="147">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122304" collectingDate="1968-01" collectionCode="MNHN" collectorName="Girard" location="Lamto-Savane" pageId="18" pageNumber="147" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FD7EDEF69D7C6BA9" box="[658,826,1189,1213]" name="Lamto-Savane" pageId="18" pageNumber="147">Lamto-Savane</location>
,
<collectorName box="[845,922,1189,1213]" pageId="18" pageNumber="147">Girard</collectorName>
leg.,
<date box="[998,1071,1189,1213]" pageId="18" pageNumber="147" value="1968-01">
<collectingDate box="[998,1071,1189,1213]" pageId="18" pageNumber="147" value="1968-01">I.1968</collectingDate>
</date>
, 1 spm. (
<collectionCode box="[220,316,1221,1245]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="147">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122376" box="[336,854,1221,1245]" collectingDate="1968-10" collectionCode="MNHN" collectorName="Girard" location="Lamto" pageId="18" pageNumber="147" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FEBCDE969FD96BC8" box="[336,415,1221,1244]" name="Lamto" pageId="18" pageNumber="147">Lamto</location>
,
<collectorName box="[426,503,1221,1245]" pageId="18" pageNumber="147">Girard</collectorName>
leg.,
<date box="[565,648,1221,1245]" pageId="18" pageNumber="147" value="1968-10">
<collectingDate box="[565,648,1221,1245]" pageId="18" pageNumber="147" value="1968-10">X.1968</collectingDate>
</date>
,
<specimenCount box="[660,735,1221,1245]" pageId="18" pageNumber="147" type="male">1 male</specimenCount>
(
<collectionCode box="[751,846,1221,1245]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="147">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122310" collectionCode="UPSA" location="Touba" municipality="Biemasso" pageId="18" pageNumber="147" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FC8FDE969DF66BC9" box="[867,944,1221,1245]" municipality="Biemasso" name="Touba" pageId="18" pageNumber="147">Touba</location>
,
<collectingMunicipality box="[956,1065,1221,1245]" pageId="18" pageNumber="147">Biemasso</collectingMunicipality>
,
<specimenCount box="[1076,1171,1221,1245]" pageId="18" pageNumber="147" type="female">1 female</specimenCount>
(
<collectionCode box="[220,298,1253,1277]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="18" pageNumber="147">UPSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122362" box="[319,701,1253,1277]" collectionCode="MNHN" country="Serbia" location="Kita" pageId="18" pageNumber="147" specimenCount="1" specimenCount-female="1">
<collectingCountry box="[319,397,1253,1277]" name="Serbia" pageId="18" pageNumber="147">MALI</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FE77DEB69F976BE9" box="[411,465,1253,1277]" country="Serbia" name="Kita" pageId="18" pageNumber="147">Kita</location>
,
<specimenCount box="[480,578,1253,1277]" pageId="18" pageNumber="147" type="female">1 female</specimenCount>
(
<collectionCode box="[597,693,1253,1277]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="147">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122354" collectingDate="1974-07-25" collectionCode="SANC" collectorName="G. F. Bornemissza" country="Nigeria" location="Yankari" pageId="18" pageNumber="147" specimenCount="1" specimenCount-female="1">
<collectingCountry box="[715,841,1253,1277]" name="Nigeria" pageId="18" pageNumber="147">NIGERIA</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FCB4DEB69DFE6BE9" box="[856,952,1253,1277]" country="Nigeria" name="Yankari" pageId="18" pageNumber="147">Yankari</location>
,
<date box="[966,1103,1253,1277]" pageId="18" pageNumber="147" value="1974-07-25">
<collectingDate box="[966,1103,1253,1277]" pageId="18" pageNumber="147" value="1974-07-25">25.VII.1974</collectingDate>
</date>
,
<collectorName pageId="18" pageNumber="147">G.F. Bornemissza</collectorName>
leg.,
<specimenCount box="[428,530,1284,1308]" pageId="18" pageNumber="147" type="female">1 female</specimenCount>
(
<collectionCode box="[552,632,1284,1308]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="18" pageNumber="147" type="Museum">SANC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122357" collectingDate="1974-07-19" collectionCode="SANC" collectorName="G. F. Bornemissza" country="Nigeria" location="Kontagora" pageId="18" pageNumber="147" specimenCount="14" specimenCount-female="14">
<quantity box="[658,735,1284,1308]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="147" unit="km" value="10.0">10 km</quantity>
E of
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FCDEDF569DF76A09" box="[818,945,1285,1309]" country="Nigeria" name="Kontagora" pageId="18" pageNumber="147">Kontagora</location>
,
<date box="[963,1099,1284,1308]" pageId="18" pageNumber="147" value="1974-07-19">
<collectingDate box="[963,1099,1284,1308]" pageId="18" pageNumber="147" value="1974-07-19">19.VII.1974</collectingDate>
</date>
,
<collectorName pageId="18" pageNumber="147">G.F. Bornemissza</collectorName>
leg.,
<specimenCount box="[423,545,1316,1340]" pageId="18" pageNumber="147" type="female">14 females</specimenCount>
(
<collectionCode box="[565,645,1316,1340]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="18" pageNumber="147" type="Museum">SANC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122345" collectionCode="MNHN" collectorName="G. F. Bornemissza" country="Nigeria" location="Samaru" pageId="18" pageNumber="147" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FD71DF779CB16A28" box="[669,759,1316,1340]" country="Nigeria" name="Samaru" pageId="18" pageNumber="147">Samaru</location>
, 20.VII.74
<collectorName box="[893,1089,1316,1340]" pageId="18" pageNumber="147">G.F.Bornemissza</collectorName>
leg.,
<specimenCount pageId="18" pageNumber="147" type="male">1 male</specimenCount>
(
<collectionCode box="[287,383,1348,1372]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="147">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122367" collectingDate="1912-12-27" collectingDateMax="1913-02-06" collectingDateMin="1912-12-27" collectionCode="BMNH" collectorName="S. A. Neave" country="Malawi" location="Mlanje" pageId="18" pageNumber="147" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectingCountry box="[408,533,1348,1372]" name="Malawi" pageId="18" pageNumber="147">MALAWI</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FDCADF179C3F6A48" box="[550,633,1348,1372]" country="Malawi" name="Mlanje" pageId="18" pageNumber="147">Mlanje</location>
,
<collectorName box="[650,775,1348,1372]" pageId="18" pageNumber="147">S.A.Neave</collectorName>
:
<date box="[792,899,1348,1372]" pageId="18" pageNumber="147" value="1913-01-21">
<collectingDate box="[792,899,1348,1372]" pageId="18" pageNumber="147" value="1913-01-21">21.I.1913</collectingDate>
</date>
,
<specimenCount box="[916,1020,1348,1372]" pageId="18" pageNumber="147" type="female">1 female</specimenCount>
,
<date box="[1037,1141,1348,1372]" pageId="18" pageNumber="147" value="1913-02-06">
<collectingDate box="[1037,1141,1348,1372]" pageId="18" pageNumber="147" value="1913-02-06">6.II.1913</collectingDate>
</date>
,
<specimenCount pageId="18" pageNumber="147" type="female">1 female</specimenCount>
,
<date box="[304,441,1380,1404]" pageId="18" pageNumber="147" value="1912-12-27">
<collectingDate box="[304,441,1380,1404]" pageId="18" pageNumber="147" value="1912-12-27">27.XII.1912</collectingDate>
</date>
,
<specimenCount box="[456,534,1380,1404]" pageId="18" pageNumber="147" type="male">1 male</specimenCount>
(
<collectionCode box="[553,645,1380,1404]" pageId="18" pageNumber="147">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122377" collectingDate="1910-03-20" collectingDateMax="1910-03-25" collectingDateMin="1910-03-20" collectionCode="BMNH" collectorName="S. A. Neave" country="Malawi" location="Ft. Mangoche" municipality="Chikala Boma" pageId="18" pageNumber="147" specimenCount="1" specimenCount-female="1">
between
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FCEEDF369DE36A68" box="[770,933,1380,1404]" country="Malawi" municipality="Chikala Boma" name="Ft. Mangoche" pageId="18" pageNumber="147">Ft. Mangoche</location>
and
<collectingMunicipality box="[997,1167,1380,1404]" pageId="18" pageNumber="147">Chikala Boma</collectingMunicipality>
,
<date box="[212,378,1412,1436]" pageId="18" pageNumber="147" value="1910-03-20" valueMax="1910-03-25" valueMin="1910-03-20">
<collectingDate box="[212,378,1412,1436]" pageId="18" pageNumber="147" value="1910-03-20" valueMax="1910-03-25" valueMin="1910-03-20">20–25.III.1910</collectingDate>
</date>
,
<collectorName box="[392,516,1412,1436]" pageId="18" pageNumber="147">S.A.Neave</collectorName>
,
<specimenCount box="[530,627,1412,1436]" pageId="18" pageNumber="147" type="female">1 female</specimenCount>
(
<collectionCode box="[645,737,1412,1436]" pageId="18" pageNumber="147">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122315" collectingDate="1925-02" collectionCode="SAMC" country="Namibia" location="Kaross" pageId="18" pageNumber="147" specimenCount="1">
<collectingCountry box="[758,889,1412,1436]" name="Namibia" pageId="18" pageNumber="147">NAMIBIA</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FC6ADFD69D9F6A88" box="[902,985,1413,1436]" country="Namibia" name="Kaross" pageId="18" pageNumber="147">Kaross</location>
,
<date box="[998,1081,1412,1436]" pageId="18" pageNumber="147" value="1925-02">
<collectingDate box="[998,1081,1412,1436]" pageId="18" pageNumber="147" value="1925-02">II.1925</collectingDate>
</date>
, 1 spm. (
<collectionCode box="[220,305,1444,1468]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="18" pageNumber="147" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122339" box="[325,838,1444,1468]" collectionCode="MNHN" country="Tanzania" location="Kigonsera" pageId="18" pageNumber="147" specimenCount="1" specimenCount-female="1">
<collectingCountry box="[325,479,1444,1468]" name="Tanzania" pageId="18" pageNumber="147">TANZANIA</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FE07DFF79C246AA8" box="[491,610,1444,1468]" country="Tanzania" name="Kigonsera" pageId="18" pageNumber="147">Kigonsera</location>
,
<specimenCount box="[622,718,1444,1468]" pageId="18" pageNumber="147" type="female">1 female</specimenCount>
(
<collectionCode box="[734,830,1444,1468]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="147">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122322" collectionCode="BMNH" collectorName="H. C. Dollman" country="Zambia" location="Mwengwa" pageId="18" pageNumber="147" specimenCount="3" specimenCount-female="3">
<collectingCountry box="[850,966,1444,1468]" name="Zambia" pageId="18" pageNumber="147">ZAMBIA</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FC3FDFF69A0D6AA9" box="[979,1099,1445,1469]" country="Zambia" name="Mwengwa" pageId="18" pageNumber="147">Mwengwa</location>
, 1919,
<collectorName box="[212,367,1476,1500]" pageId="18" pageNumber="147">H.C.Dollman</collectorName>
leg.,
<specimenCount box="[431,536,1476,1500]" pageId="18" pageNumber="147" type="female">3 females</specimenCount>
(
<collectionCode box="[553,645,1476,1500]" pageId="18" pageNumber="147">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122307" collectionCode="MNHN" collectorName="J. Surcouf" country="Mozambique" location="Canxixe" pageId="18" pageNumber="147" specimenCount="1" specimenCount-male="1">
<collectingCountry box="[665,868,1476,1500]" name="Mozambique" pageId="18" pageNumber="147">MOZAMBIQUE</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081028FFA0FE3ADEB69FF8698F:746807C5102BFFA0FC9CDF979D896AC8" box="[880,975,1476,1500]" country="Mozambique" name="Canxixe" pageId="18" pageNumber="147">Canxixe</location>
, 1926,
<collectorName box="[1057,1172,1476,1500]" pageId="18" pageNumber="147">J. Surcouf</collectorName>
leg.,
<specimenCount box="[267,343,1508,1532]" pageId="18" pageNumber="147" type="male">1 male</specimenCount>
(
<collectionCode box="[360,456,1508,1532]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="147">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="147" type="distribution">
<paragraph blockId="18.[212,1172,1596,1691]" box="[212,344,1596,1620]" pageId="18" pageNumber="147">
<heading box="[212,344,1596,1620]" fontSize="10" level="3" pageId="18" pageNumber="147" reason="5">
<emphasis box="[212,344,1596,1620]" italics="true" pageId="18" pageNumber="147">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[212,1172,1596,1691]" pageId="18" pageNumber="147">
The species is widely distributed in the savannas both south and north from the equator (
<figureCitation box="[316,430,1667,1691]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="18" pageNumber="147">Figure 5F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

259
data/F9/1E/E0/F91EE0081029FFA3FE24D90F9FCD6BBA.xml Normal file
View file

@ -0,0 +1,259 @@
<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE0081029FFA3FE24D90F9FCD6BBA" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus darrenmanni Frolov 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="146" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="145" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
</mods:titleInfo>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>49</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>129</mods:start>
<mods:end>154</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2014.909072</mods:identifier>
<mods:identifier type="GBIF-Dataset">3bad6762-1788-4f94-9cfd-1281b8619c55</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">4002597</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331155" ID-GBIF-Taxon="167368590" ID-Zenodo-Dep="4331155" LSID="urn:lsid:plazi:treatment:F91EE0081029FFA3FE24D90F9FCD6BBA" httpUri="http://treatment.plazi.org/id/F91EE0081029FFA3FE24D90F9FCD6BBA" lastPageId="17" lastPageNumber="146" pageId="16" pageNumber="145">
<subSubSection box="[456,927,860,885]" pageId="16" pageNumber="145" type="nomenclature">
<paragraph blockId="16.[456,927,860,917]" box="[456,927,860,885]" pageId="16" pageNumber="145">
<heading box="[456,927,860,885]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="145" reason="2">
<taxonomicName ID-CoL="34H55" authority="Frolov" authorityName="Frolov" authorityYear="2014" box="[456,824,860,885]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="145" phylum="Arthropoda" rank="species" species="darrenmanni" status="sp. nov.">
<emphasis bold="true" box="[456,735,860,884]" italics="true" pageId="16" pageNumber="145">Delopleurus darrenmanni</emphasis>
Frolov
</taxonomicName>
,
<taxonomicNameLabel box="[837,927,862,885]" pageId="16" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[609,775,893,917]" pageId="16" pageNumber="145" type="description">
<paragraph blockId="16.[456,927,860,917]" box="[609,775,893,917]" pageId="16" pageNumber="145">
(
<figureCitation box="[618,766,893,917]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="16" pageNumber="145">Figure 4A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="145" type="diagnosis">
<paragraph blockId="16.[212,1172,952,1110]" box="[212,320,952,975]" pageId="16" pageNumber="145">
<heading box="[212,320,952,975]" fontSize="10" level="3" pageId="16" pageNumber="145" reason="5">
<emphasis box="[212,320,952,975]" italics="true" pageId="16" pageNumber="145">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="16.[212,1172,952,1110]" pageId="16" pageNumber="145">
This species can be separated from other
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[692,823,991,1015]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis box="[692,823,991,1015]" italics="true" pageId="16" pageNumber="145">Delopleurus</emphasis>
</taxonomicName>
species by the sculpture of its pygidium. Males have punctate, glabrous, narrow disc of pygidium which is almost as long as apical and basal border in the middle (
<figureCitation box="[728,840,1054,1078]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="16" pageNumber="145">Figure 4B</figureCitation>
). Females have characteristic slit-shaped disc with a transverse carina in the middle (
<figureCitation box="[841,954,1086,1110]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="16" pageNumber="145">Figure 4E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="146" pageId="16" pageNumber="145" type="description">
<paragraph blockId="16.[212,1172,1182,1691]" box="[212,339,1182,1205]" pageId="16" pageNumber="145">
<heading box="[212,339,1182,1205]" fontSize="10" level="3" pageId="16" pageNumber="145" reason="5">
<emphasis box="[212,339,1182,1205]" italics="true" pageId="16" pageNumber="145">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="16.[212,1172,1182,1691]" box="[212,1172,1221,1245]" pageId="16" pageNumber="145">
<emphasis box="[212,277,1221,1245]" italics="true" pageId="16" pageNumber="145">Male,</emphasis>
<typeStatus box="[283,381,1221,1245]" pageId="16" pageNumber="145">holotype</typeStatus>
. Body strongly convex, black, glabrous, its length 6.0 mm (
<figureCitation box="[1042,1156,1221,1245]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="16" pageNumber="145">Figure 4A</figureCitation>
).
</paragraph>
<paragraph blockId="16.[212,1172,1182,1691]" pageId="16" pageNumber="145">Clypeus quadridentate, medial teeth acute (right tooth malformed), lateral ones acute-angled. Clypeal margin laterad of lateral teeth somewhat angulate so clypeus looks hexadentate. Clypeal surface rugose in anterior part and laterally, disc of clypeus and frons densely punctate (punctures separated by less than a puncture diameter).</paragraph>
<paragraph blockId="16.[212,1172,1182,1691]" pageId="16" pageNumber="145">Pronotum more or less trapezoidal, about two times wider than long. Anterior and lateral margins distinctly bordered, base feebly bordered. Surface relatively sparsely punctate on disc (punctures separated by two to three puncture diameters) and denser laterally.</paragraph>
<paragraph blockId="16.[212,1172,1182,1691]" pageId="16" pageNumber="145">Elytra trapezoidal, as wide as long, shiny. Stria relatively deep, with punctures slightly larger than striae width. Elytral intervals feebly convex, with minute punctation.</paragraph>
<paragraph blockId="16.[212,1172,1182,1691]" pageId="16" pageNumber="145">Anterior tibiae with three outer teeth, without a small acute tooth between first outer tooth and apical spur. Anterior tibial spur bifurcated apically.</paragraph>
<paragraph blockId="17.[144,1104,168,288]" pageId="17" pageNumber="146">
Pygidium with relatively thick borders and convex disc. Apical border almost as long as disc in the middle. Disc glabrous, punctate with punctures separated by two to four puncture diameters (
<figureCitation box="[434,548,232,256]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="17" pageNumber="146">Figure 4B</figureCitation>
).
</paragraph>
<paragraph blockId="17.[144,1104,168,288]" box="[186,725,264,288]" pageId="17" pageNumber="146">
Parameres rounded in lateral view (
<figureCitation box="[594,709,264,288]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="17" pageNumber="146">Figure 4C</figureCitation>
).
</paragraph>
<paragraph blockId="17.[144,1103,337,425]" pageId="17" pageNumber="146">
<emphasis box="[144,224,338,362]" italics="true" pageId="17" pageNumber="146">Female</emphasis>
can be separated from male in having acute, not bifurcated spurs of anterior tibiae and in sculpture of pygidium, which has very wide borders and very narrow disc with transverse keel (
<figureCitation box="[438,551,401,425]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="17" pageNumber="146">Figure 4E</figureCitation>
).
</paragraph>
<paragraph blockId="17.[144,760,475,500]" box="[144,760,475,500]" pageId="17" pageNumber="146">
<heading box="[144,760,475,500]" fontSize="10" level="3" pageId="17" pageNumber="146" reason="5">
<emphasis box="[144,271,475,499]" italics="true" pageId="17" pageNumber="146">Variability.</emphasis>
Body length of the
<typeStatus box="[502,611,476,500]" pageId="17" pageNumber="146">paratypes</typeStatus>
4.0–
<quantity box="[666,756,475,499]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="17" pageNumber="146" unit="mm" value="6.2">6.2 mm</quantity>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="146" type="distribution">
<paragraph blockId="17.[144,1104,573,668]" box="[144,276,573,597]" pageId="17" pageNumber="146">
<heading box="[144,276,573,597]" fontSize="10" level="3" pageId="17" pageNumber="146" reason="5">
<emphasis box="[144,276,573,597]" italics="true" pageId="17" pageNumber="146">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[144,1104,573,668]" pageId="17" pageNumber="146">
The species is known from three localities in a relatively small area between the
<collectingRegion box="[144,418,644,668]" isEnumeration="true" pageId="17" pageNumber="146">Zambezi and Okavango</collectingRegion>
rivers in Zambezian woodland region (
<figureCitation box="[871,986,644,668]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="17" pageNumber="146">Figure 4F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[144,1098,742,806]" pageId="17" pageNumber="146" type="etymology">
<paragraph blockId="17.[144,1098,742,806]" box="[144,265,742,766]" pageId="17" pageNumber="146">
<heading box="[144,265,742,766]" fontSize="10" level="3" pageId="17" pageNumber="146" reason="5">
<emphasis box="[144,265,742,766]" italics="true" pageId="17" pageNumber="146">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[144,1098,742,806]" box="[144,1098,782,806]" pageId="17" pageNumber="146">
The species is named after Darren Mann (OUMNH), who collected the
<typeStatus box="[973,1021,782,806]" pageId="17" pageNumber="146">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="146" type="materials_examined">
<paragraph blockId="17.[144,1104,880,1198]" box="[144,300,880,904]" pageId="17" pageNumber="146">
<heading box="[144,300,880,904]" fontSize="10" level="3" pageId="17" pageNumber="146" reason="5">
<emphasis box="[144,300,880,904]" italics="true" pageId="17" pageNumber="146">Type material</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[144,1104,880,1198]" pageId="17" pageNumber="146">
<materialsCitation ID-GBIF-Occurrence="2849122379" collectingDate="1999-12-19" collectionCode="OUMNH" country="Namibia" location="W. Caprivi Park Nova" longitude="21.741945" pageId="17" pageNumber="146" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[144,253,919,943]" pageId="17" pageNumber="146">Holotype</typeStatus>
,
<specimenCount box="[269,323,919,943]" pageId="17" pageNumber="146" type="male">male</specimenCount>
with the labels ‘
<collectingCountry box="[528,627,919,943]" name="Namibia" pageId="17" pageNumber="146">Namibia</collectingCountry>
<location LSID="urn:lsid:plazi:treatment:F91EE0081029FFA3FE24D90F9FCD6BBA:746807C51028FFA3FD6CD9CB9DD56CBB" box="[640,915,919,943]" country="Namibia" longitude="21.741945" name="W. Caprivi Park Nova" pageId="17" pageNumber="146">W. Caprivi Park Nova</location>
,
<locationDeviation pageId="17" pageNumber="146">
<quantity box="[932,993,919,943]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="146" unit="km" value="5.0">5 km</quantity>
north of Okavango River
</locationDeviation>
18°09
<emphasis bold="true" box="[405,411,951,975]" italics="true" pageId="17" pageNumber="146">ʹ</emphasis>
56
<emphasis bold="true" box="[437,443,951,975]" italics="true" pageId="17" pageNumber="146">ʹ</emphasis>
S, 21°44
<emphasis bold="true" box="[546,552,951,975]" italics="true" pageId="17" pageNumber="146">ʹ</emphasis>
31
<emphasis bold="true" box="[578,588,951,975]" italics="true" pageId="17" pageNumber="146">ʺ</emphasis>
E,
<date box="[630,756,951,975]" pageId="17" pageNumber="146" value="1999-12-19">
<collectingDate box="[630,756,951,975]" pageId="17" pageNumber="146" value="1999-12-19">19.xii.1999</collectingDate>
</date>
. In fungi. Coll.
<collectorName box="[939,1010,952,975]" pageId="17" pageNumber="146">Mann</collectorName>
,
<collectorName box="[1023,1103,951,975]" pageId="17" pageNumber="146">Marais</collectorName>
&amp;
<collectorName box="[177,286,983,1007]" pageId="17" pageNumber="146">Newman’</collectorName>
, ‘D.J.Mann colln. OX.UNI.MUS.NAT.HIST (
<collectionCode box="[848,962,983,1007]" collectionName="United Kingdom, Oxford, University Museum of Natural History" pageId="17" pageNumber="146">OUMNH</collectionCode>
) OUMNH- 2006-093
</materialsCitation>
’.
<materialsCitation ID-GBIF-Occurrence="2849122336" collectingDate="1999-12-19" collectionCode="OUMNH" collectorName="Botswana. Namibia" location="Namibia" longitude="21.741945" municipality="Zambia" pageId="17" pageNumber="146" specimenCount="23" specimenCount-female="6" specimenCount-male="17" typeStatus="paratype">
<typeStatus box="[267,380,1016,1040]" pageId="17" pageNumber="146">Paratypes</typeStatus>
,
<specimenCount box="[392,490,1015,1039]" pageId="17" pageNumber="146" type="male">17 males</specimenCount>
and
<specimenCount box="[546,651,1015,1039]" pageId="17" pageNumber="146" type="female">6 females</specimenCount>
from
<location LSID="urn:lsid:plazi:treatment:F91EE0081029FFA3FE24D90F9FCD6BBA:746807C51028FFA3FD3CD9A49D706B1B" box="[720,822,1015,1039]" municipality="Zambia" name="Namibia" pageId="17" pageNumber="146">Namibia</location>
,
<collectingMunicipality box="[834,925,1015,1039]" pageId="17" pageNumber="146">Zambia</collectingMunicipality>
, and
<collectorName pageId="17" pageNumber="146">Botswana. Namibia</collectorName>
: same data as the holotype
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122380" box="[561,959,1047,1071]" collectionCode="OUMNH" pageId="17" pageNumber="146" specimenCount="18" specimenCount-female="5" specimenCount-male="13" typeStatus="paratype">
<specimenCount box="[561,659,1047,1071]" pageId="17" pageNumber="146" type="male">13 males</specimenCount>
and
<specimenCount box="[715,819,1047,1071]" pageId="17" pageNumber="146" type="female">5 females</specimenCount>
(
<collectionCode box="[834,950,1047,1071]" pageId="17" pageNumber="146">OUMNH</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122301" collectionCode="ZIN" country="Botswana" location="BOTSWANA" pageId="17" pageNumber="146" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount box="[970,1054,1047,1071]" pageId="17" pageNumber="146" type="male">2 males</specimenCount>
and
<specimenCount box="[144,243,1079,1103]" pageId="17" pageNumber="146" type="female">1 female</specimenCount>
(
<collectionCode box="[264,318,1079,1103]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="17" pageNumber="146" type="Museum">ZIN</collectionCode>
).
<collectingCountry box="[342,506,1079,1103]" name="Botswana" pageId="17" pageNumber="146">BOTSWANA</collectingCountry>
</materialsCitation>
:
<materialsCitation ID-GBIF-Occurrence="2849122347" collectionCode="UPSA" collectorName="Savuti Botswana S &amp; Deschodt Tshikai" country="Zambia" location="ZAMBIA" longitude="18.5651" pageId="17" pageNumber="146" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[521,601,1079,1103]" pageId="17" pageNumber="146" type="male">1 male</specimenCount>
with the label ‘
<collectorName box="[795,1015,1079,1103]" pageId="17" pageNumber="146">Savuti Botswana S</collectorName>
18.5651 E24.0629
<collectorName box="[262,468,1110,1134]" pageId="17" pageNumber="146">Deschodt Tshikai</collectorName>
07.12.05’ (
<collectionCode box="[605,682,1110,1134]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="17" pageNumber="146">UPSA</collectionCode>
).
<collectingCountry box="[708,825,1110,1134]" name="Zambia" pageId="17" pageNumber="146">ZAMBIA</collectingCountry>
</materialsCitation>
:
<materialsCitation ID-GBIF-Occurrence="2849122359" ID-GBIF-Specimen="2592561513" collectingDate="2003-11-26" collectingDateMax="2003-11-31" collectingDateMin="2003-11-26" collectionCode="UPSA" collectorName="Deschodt &amp; Groenewald'" country="Zambia" httpUri="https://www.gbif.org/occurrence/2592561513" latitude="-17.041313" location="Kabula Loge" longLatPrecision="1" longitude="24.013018" municipality="Zambia" pageId="17" pageNumber="146" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[842,924,1110,1134]" pageId="17" pageNumber="146" type="male">1 male</specimenCount>
with the label ‘
<location LSID="urn:lsid:plazi:treatment:F91EE0081029FFA3FE24D90F9FCD6BBA:746807C51028FFA3FF7BDE259F7E6B9B" box="[151,312,1142,1167]" country="Zambia" latitude="-17.041313" longLatPrecision="1" longitude="24.013018" municipality="Zambia" name="Kabula Loge" pageId="17" pageNumber="146">Kabula Loge</location>
,
<collectingMunicipality box="[334,422,1142,1166]" pageId="17" pageNumber="146">Zambia</collectingMunicipality>
<geoCoordinate box="[439,586,1142,1166]" degrees="17.041314" direction="south" orientation="latitude" pageId="17" pageNumber="146" precision="1" value="-17.041313">17.041314 S</geoCoordinate>
,
<geoCoordinate box="[608,770,1142,1166]" degrees="024.013017" direction="east" orientation="longitude" pageId="17" pageNumber="146" precision="1" value="24.013018">024.013017 E</geoCoordinate>
<date box="[787,942,1142,1166]" pageId="17" pageNumber="146" value="2003-11-26" valueMax="2003-11-31" valueMin="2003-11-26">
<collectingDate box="[787,942,1142,1166]" pageId="17" pageNumber="146" value="2003-11-26" valueMax="2003-11-31" valueMin="2003-11-26">26–31.xi.2003</collectingDate>
</date>
<collectorName box="[959,1066,1142,1166]" pageId="17" pageNumber="146">Deschodt</collectorName>
&amp;
<collectorName box="[144,289,1174,1198]" pageId="17" pageNumber="146">Groenewald’</collectorName>
(
<collectionCode box="[306,384,1174,1198]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="17" pageNumber="146">UPSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

278
data/F9/1E/E0/F91EE008102AFFA6FE70DAFB9C836DBD.xml Normal file
View file

@ -0,0 +1,278 @@
<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE008102AFFA6FE70DAFB9C836DBD" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus krikkeni Frolov 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="149" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="148" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>49</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>129</mods:start>
<mods:end>154</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2014.909072</mods:identifier>
<mods:identifier type="GBIF-Dataset">3bad6762-1788-4f94-9cfd-1281b8619c55</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">4002597</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331159" ID-GBIF-Taxon="167368589" ID-Zenodo-Dep="4331159" LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD" httpUri="http://treatment.plazi.org/id/F91EE008102AFFA6FE70DAFB9C836DBD" lastPageId="20" lastPageNumber="149" pageId="19" pageNumber="148">
<subSubSection box="[412,836,168,192]" pageId="19" pageNumber="148" type="nomenclature">
<paragraph blockId="19.[412,836,168,224]" box="[412,836,168,192]" pageId="19" pageNumber="148">
<heading box="[412,836,168,192]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="148" reason="2">
<taxonomicName ID-CoL="34H59" authority="Frolov" authorityName="Frolov" authorityYear="2014" box="[412,733,168,192]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="148" phylum="Arthropoda" rank="species" species="krikkeni" status="sp. nov.">
<emphasis bold="true" box="[412,645,168,192]" italics="true" pageId="19" pageNumber="148">Delopleurus krikkeni</emphasis>
Frolov
</taxonomicName>
,
<taxonomicNameLabel box="[746,836,169,192]" pageId="19" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[539,709,200,224]" pageId="19" pageNumber="148" type="description">
<paragraph blockId="19.[412,836,168,224]" box="[539,709,200,224]" pageId="19" pageNumber="148">
(
<figureCitation box="[548,700,200,224]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="148" type="diagnosis">
<paragraph blockId="19.[144,1104,273,463]" box="[144,252,273,296]" pageId="19" pageNumber="148">
<heading box="[144,252,273,296]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
<emphasis box="[144,252,273,296]" italics="true" pageId="19" pageNumber="148">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[144,1104,273,463]" pageId="19" pageNumber="148">
This species can be easily separated from other
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[735,866,312,336]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis box="[735,866,312,336]" italics="true" pageId="19" pageNumber="148">Delopleurus</emphasis>
</taxonomicName>
species by sparsely punctate to impunctate disc and base of pronotum. Its pygidium is similar to that of
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[177,301,376,400]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="148" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
<emphasis box="[177,301,376,400]" italics="true" pageId="19" pageNumber="148">D. fossatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[311,402,376,399]" pageId="19" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
but differs in having convex basal border distinctly widened medially (
<figureCitation box="[255,368,407,431]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6B</figureCitation>
). From
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[461,583,408,432]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="148" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
<emphasis box="[461,583,408,432]" italics="true" pageId="19" pageNumber="148">D. fossatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[590,678,408,431]" pageId="19" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
it also differs in having larger eyes (in dorsal view).
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="148" type="description">
<paragraph blockId="19.[144,1104,544,1053]" box="[144,271,544,567]" pageId="19" pageNumber="148">
<heading box="[144,271,544,567]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
<emphasis box="[144,271,544,567]" italics="true" pageId="19" pageNumber="148">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[144,1104,544,1053]" box="[144,1099,583,607]" pageId="19" pageNumber="148">
<emphasis box="[144,323,583,607]" italics="true" pageId="19" pageNumber="148">
<typeStatus box="[144,249,583,607]" pageId="19" pageNumber="148">Holotype</typeStatus>
, male.
</emphasis>
Body strongly convex, black, glabrous, length
<quantity box="[863,949,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="19" pageNumber="148" unit="mm" value="4.6">4.6 mm</quantity>
(
<figureCitation box="[966,1083,583,607]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6A</figureCitation>
).
</paragraph>
<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">Clypeus quadridentate. Two medial teeth acute, lateral ones right-angled. Clypeal surface coarsely punctate in anterior part and laterally, frons densely punctate (punctures separated by one to two puncture diameters).</paragraph>
<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">Pronotum about two times wider than long. Anterior and lateral margins distinctly bordered, base feebly bordered laterally. Disc and base of pronotum impunctate, sides relatively densely punctate.</paragraph>
<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">Elytra trapezoidal, as wide as long, shiny. Stria relatively deep, with punctures 1.5–2 times larger than striae width. Elytral intervals convex, with minute punctation.</paragraph>
<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">Anterior tibiae with three outer teeth, without a small acute tooth between first outer tooth and apical spur. Anterior tibial spur bifurcated apically.</paragraph>
<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">
Pygidium with relatively strongly convex disc separate by deep circular fossa from border. The apical border wider medially and somewhat tapering laterally; apical border somewhat tuberculate in the middle (
<figureCitation box="[652,766,997,1021]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6B</figureCitation>
).
</paragraph>
<paragraph blockId="19.[144,1104,544,1053]" box="[186,862,1029,1053]" pageId="19" pageNumber="148">
Parameres in lateral view with small processes (
<figureCitation box="[730,846,1029,1053]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6C</figureCitation>
).
</paragraph>
<paragraph blockId="19.[144,1103,1109,1165]" pageId="19" pageNumber="148">
<emphasis box="[144,319,1109,1133]" italics="true" pageId="19" pageNumber="148">
<typeStatus box="[144,248,1110,1133]" pageId="19" pageNumber="148">Paratype</typeStatus>
, male.
</emphasis>
Differs from the
<typeStatus box="[514,612,1109,1133]" pageId="19" pageNumber="148">holotype</typeStatus>
in being slightly larger (length
<quantity box="[961,1047,1109,1133]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="19" pageNumber="148" unit="mm" value="4.8">4.8 mm</quantity>
) and having only anteromedial part of pronotal disc impunctate.
</paragraph>
<paragraph blockId="19.[144,346,1221,1245]" box="[144,346,1221,1245]" pageId="19" pageNumber="148">
<heading box="[144,346,1221,1245]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
<emphasis box="[144,224,1221,1245]" italics="true" pageId="19" pageNumber="148">Female</emphasis>
unknown.
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="148" type="distribution">
<paragraph blockId="19.[144,1104,1325,1548]" box="[144,276,1325,1349]" pageId="19" pageNumber="148">
<heading box="[144,276,1325,1349]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
<emphasis box="[144,276,1325,1349]" italics="true" pageId="19" pageNumber="148">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[144,1104,1325,1548]" pageId="19" pageNumber="148">
<collectingCounty box="[144,188,1364,1388]" pageId="19" pageNumber="148">The</collectingCounty>
locality where the
<typeStatus box="[428,526,1364,1388]" pageId="19" pageNumber="148">holotype</typeStatus>
was collected is uncertain (Adrian Davis, pers. comm.).
<collectingMunicipality box="[246,270,1397,1420]" pageId="19" pageNumber="148">In</collectingMunicipality>
1896, ‘
<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FE8ADF279F926A98" box="[358,468,1396,1420]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="Samburu'" pageId="19" pageNumber="148" stateProvince="Turkana">Samburu’</location>
might refer to a large area to the northwest of
<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FBEDDF269A096A98" box="[1025,1103,1397,1420]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="Mount" pageId="19" pageNumber="148" stateProvince="Turkana">Mount</location>
<collectingCountry box="[144,218,1429,1453]" name="Kenya" pageId="19" pageNumber="148">Kenya</collectingCountry>
and to the south of
<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FE2CDFC79FBF6AB8" box="[448,505,1428,1452]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="Lake" pageId="19" pageNumber="148" stateProvince="Turkana">Lake</location>
<collectingRegion box="[513,614,1428,1452]" country="Kenya" name="Turkana" pageId="19" pageNumber="148">Turkana</collectingRegion>
, that was occupied by the
<collectingRegion box="[912,1015,1428,1452]" country="Kenya" name="Samburu" pageId="19" pageNumber="148">Samburu</collectingRegion>
people.
<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FF7CDFE79EFF6ADF" box="[144,185,1460,1483]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="But" pageId="19" pageNumber="148" stateProvince="Turkana">But</location>
it might also be the railway station (
<geoCoordinate box="[596,676,1460,1484]" degrees="3" direction="south" minutes="46" orientation="latitude" pageId="19" pageNumber="148" precision="925" value="-3.7666667">
3°46
<emphasis bold="true" box="[647,653,1460,1484]" italics="true" pageId="19" pageNumber="148">ʹ</emphasis>
S
</geoCoordinate>
,
<geoCoordinate box="[688,787,1460,1484]" pageId="19" pageNumber="148">
39°16
<emphasis bold="true" box="[752,762,1460,1484]" italics="true" pageId="19" pageNumber="148">ʺ</emphasis>
E
</geoCoordinate>
) on the railroad Mombasa–Nairobi. This latter locality is some
<quantity box="[562,647,1492,1516]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="148" unit="km" value="100.0">100 km</quantity>
east-northeast of the locality where the
<typeStatus box="[144,243,1524,1548]" pageId="19" pageNumber="148">paratype</typeStatus>
was collected,
<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FE4DDFA79CFB691F" box="[417,701,1524,1548]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="Mkomazi Game Reserve" pageId="19" pageNumber="148" stateProvince="Turkana">Mkomazi Game Reserve</location>
(
<figureCitation box="[719,837,1524,1548]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[144,828,1628,1691]" pageId="19" pageNumber="148" type="etymology">
<paragraph blockId="19.[144,828,1628,1691]" box="[144,265,1628,1652]" pageId="19" pageNumber="148">
<heading box="[144,265,1628,1652]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
<emphasis box="[144,265,1628,1652]" italics="true" pageId="19" pageNumber="148">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[144,828,1628,1691]" box="[144,828,1667,1691]" pageId="19" pageNumber="148">The species is named after Jan Krikken (Naturalis, Leiden).</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="149" type="materials_examined">
<paragraph blockId="20.[212,1172,168,328]" box="[212,368,168,192]" pageId="20" pageNumber="149">
<heading box="[212,368,168,192]" fontSize="10" level="3" pageId="20" pageNumber="149" reason="5">
<emphasis box="[212,368,168,192]" italics="true" pageId="20" pageNumber="149">Type material</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[212,1172,168,328]" pageId="20" pageNumber="149">
<materialsCitation ID-GBIF-Occurrence="2849122352" collectingDateMax="1836-11-20" collectingDateMin="1896-10-31" collectionCode="BMNH" collectorName="B. E. Africa &amp; C. S. Bretton" country="Kenya" location="Samburu" pageId="20" pageNumber="149" specimenCount="1" stateProvince="Samburu" typeStatus="holotype">
<typeStatus box="[212,321,208,232]" pageId="20" pageNumber="149">Holotype</typeStatus>
,
<specimenCount box="[333,387,208,232]" pageId="20" pageNumber="149" type="male">male</specimenCount>
with labels ‘
<collectingRegion box="[533,636,208,232]" country="Kenya" name="Samburu" pageId="20" pageNumber="149">Samburu</collectingRegion>
[
<collectingCountry box="[651,755,208,232]" name="Kenya" pageId="20" pageNumber="149">KENYA</collectingCountry>
]
<date box="[770,934,208,232]" pageId="20" pageNumber="149">30.X to 20.XI.</date>
[18]96’, ‘
<collectorName box="[1034,1171,208,232]" pageId="20" pageNumber="149">B. E. Africa</collectorName>
<collectorName box="[212,362,240,264]" pageId="20" pageNumber="149">C. S. Bretton</collectorName>
98-12’, ‘diff. from the others Krikken 1975’ (
<collectionCode box="[882,974,240,264]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="20" pageNumber="149">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122306" collectingDate="1995-04-07" collectingDateMax="1995-04-08" collectingDateMin="1995-04-07" collectionCode="BMNH" collectorName="Mkomazi GR &amp; J. G. Davies" country="Tanzania" location="Ibaya Camp" pageId="20" pageNumber="149" specimenCount="1" typeStatus="paratype">
<typeStatus box="[994,1097,241,265]" pageId="20" pageNumber="149">Paratype</typeStatus>
, male:
<collectingCountry box="[212,367,272,296]" name="Tanzania" pageId="20" pageNumber="149">TANZANIA</collectingCountry>
:
<collectorName box="[383,551,272,296]" pageId="20" pageNumber="149">Mkomazi GR</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102DFFA6FDD5DB439C896E3C" box="[569,719,272,296]" country="Tanzania" name="Ibaya Camp" pageId="20" pageNumber="149">Ibaya Camp</location>
, unburnt grassland,
<collectingMethod box="[967,1107,272,296]" pageId="20" pageNumber="149">pitfall traps</collectingMethod>
,
<date pageId="20" pageNumber="149" value="1995-04-07" valueMax="1995-04-08" valueMin="1995-04-07">
<collectingDate pageId="20" pageNumber="149" value="1995-04-07" valueMax="1995-04-08" valueMin="1995-04-07">7–8. IV.1995</collectingDate>
</date>
,
<collectorName box="[317,451,304,328]" pageId="20" pageNumber="149">J.G. Davies</collectorName>
leg. (
<collectionCode box="[516,609,304,328]" pageId="20" pageNumber="149">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="149" type="discussion">
<paragraph blockId="20.[212,1172,394,681]" box="[212,310,394,418]" pageId="20" pageNumber="149">
<heading box="[212,310,394,418]" fontSize="10" level="3" pageId="20" pageNumber="149" reason="5">
<emphasis box="[212,310,394,418]" italics="true" pageId="20" pageNumber="149">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[212,1172,394,681]" pageId="20" pageNumber="149">
Two new species are described in this work from the Eastern Arc Region,
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[1046,1172,434,458]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="krikkeni" status="sp. nov.">
<emphasis box="[1046,1066,435,458]" italics="true" pageId="20" pageNumber="149">D</emphasis>
.
<emphasis box="[1081,1172,434,458]" italics="true" pageId="20" pageNumber="149">krikkeni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[212,302,467,490]" pageId="20" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[363,494,467,490]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
<emphasis box="[363,383,467,490]" italics="true" pageId="20" pageNumber="149">D</emphasis>
.
<emphasis box="[400,494,467,490]" italics="true" pageId="20" pageNumber="149">naviauxi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[503,594,467,490]" pageId="20" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
The former species is known from two males and the latter from two females only. I believe that these specimens are not conspecific because the male specimens lack a characteristic small tooth apicad of the first major outer tooth on the anterior tibae, which is found in the
<taxonomicName authorityName="Frolov et Cambefort" authorityYear="2014" box="[866,998,562,586]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis box="[866,886,563,586]" italics="true" pageId="20" pageNumber="149">D</emphasis>
.
<emphasis box="[904,998,562,585]" italics="true" pageId="20" pageNumber="149">naviauxi</emphasis>
</taxonomicName>
<typeStatus box="[1008,1110,561,585]" pageId="20" pageNumber="149">holotype</typeStatus>
. Sexdependent modifications of the legs are common in the
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[835,983,593,617]" class="Insecta" family="Scarabaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="153" phylum="Arthropoda" rank="subFamily" subFamily="Scarabaeinae">Scarabaeinae</taxonomicName>
but found in the males only. I think that this additional tibial tooth is sex-independent and should be present in the males of
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[480,610,658,681]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
<emphasis box="[480,500,658,681]" italics="true" pageId="20" pageNumber="149">D</emphasis>
.
<emphasis box="[516,610,658,681]" italics="true" pageId="20" pageNumber="149">naviauxi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[618,709,658,681]" pageId="20" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

303
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<treatment id="F91EE008102CFFA4FE34D9D09A1A6A6D" ID-DOI="http://doi.org/10.5281/zenodo.4331153" ID-GBIF-Taxon="167368588" ID-Zenodo-Dep="4331153" LSID="urn:lsid:plazi:treatment:F91EE008102CFFA4FE34D9D09A1A6A6D" httpUri="http://treatment.plazi.org/id/F91EE008102CFFA4FE34D9D09A1A6A6D" lastPageId="22" lastPageNumber="151" pageId="21" pageNumber="150">
<subSubSection id="39AD0295102CFFA7FE34D9D09D4E6C8F" box="[472,776,899,923]" pageId="21" pageNumber="150" type="nomenclature">
<paragraph id="7108511E102CFFA7FE34D9D09D4E6C8F" blockId="21.[472,776,899,955]" box="[472,776,899,923]" pageId="21" pageNumber="150">
<heading id="2A40E672102CFFA7FE34D9D09D4E6C8F" box="[472,776,899,923]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="150" reason="2">
<taxonomicName id="B6B72A9D102CFFA7FE34D9D09D4E6C8F" ID-CoL="34H5C" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[472,776,899,923]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="150" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="43C38D0C102CFFA7FE34D9D09CED6C8E" bold="true" box="[472,683,899,923]" italics="true" pageId="21" pageNumber="150">Delopleurus parvus</emphasis>
(Sharp)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102CFFA7FDF1D9F09C856CAF" box="[541,707,931,955]" pageId="21" pageNumber="150" type="description">
<paragraph id="7108511E102CFFA7FDF1D9F09C856CAF" blockId="21.[472,776,899,955]" box="[541,707,931,955]" pageId="21" pageNumber="150">
(
<figureCitation id="E98C4D9B102CFFA7FDCAD9F09CFC6CAF" box="[550,698,931,955]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="21" pageNumber="150">Figure 8A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102CFFA7FF7ED98B9FA36CE4" box="[146,485,984,1008]" pageId="21" pageNumber="150" type="reference_group">
<paragraph id="7108511E102CFFA7FF7ED98B9FA36CE4" blockId="21.[145,643,984,1041]" box="[146,485,984,1008]" pageId="21" pageNumber="150">
<heading id="2A40E672102CFFA7FF7ED98B9FA36CE4" box="[146,485,984,1008]" fontSize="10" level="3" pageId="21" pageNumber="150" reason="5">
<taxonomicName id="B6B72A9D102CFFA7FF7ED98B9F986CE4" ID-CoL="Y889" authority="Sharp 1876" authorityName="Sharp" authorityYear="1876" box="[146,478,984,1008]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="150" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="43C38D0C102CFFA7FF7ED98B9F126CE4" box="[146,340,984,1008]" italics="true" pageId="21" pageNumber="150">Coptorhina parva</emphasis>
<bibRefCitation id="15262CEF102CFFA7FEB2D98B9F986CE4" author="Sharp D" box="[350,478,984,1008]" pageId="21" pageNumber="150" pagination="33 - 54" refId="ref10290" refString="Sharp D. 1876. Descriptions of some new genera and species of Scarabaeidae from tropical Asia and Malasia. Col Hefte. 13: 33 - 54." type="journal article" year="1876">Sharp 1876</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102CFFA7FF7DD9AB9CC56B04" box="[145,643,1016,1041]" pageId="21" pageNumber="150" type="reference_group">
<paragraph id="7108511E102CFFA7FF7DD9AB9CC56B04" blockId="21.[145,643,984,1041]" box="[145,643,1016,1041]" pageId="21" pageNumber="150">
<taxonomicName id="B6B72A9D102CFFA7FF7DD9AB9C4A6B04" ID-CoL="34H54" authority="Paulian 1934" authorityName="Paulian" authorityYear="1934" box="[145,524,1016,1040]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="150" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="43C38D0C102CFFA7FF7DD9AB9F296B04" box="[145,367,1016,1040]" italics="true" pageId="21" pageNumber="150">Delopleurus cardoni</emphasis>
<bibRefCitation id="15262CEF102CFFA7FE94D9AB9C4A6B04" author="Paulian R" box="[376,524,1016,1040]" pageId="21" pageNumber="150" pagination="162 - 164" refId="ref10236" refString="Paulian R. 1934. Quelques Panelini asiatiques nouveaux ou peu connus. Bull Soc Ent Fr. 39: 162 - 164." type="journal article" year="1934">Paulian 1934</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="58F03077102CFFA7FDF0D9AA9CC56B04" box="[540,643,1017,1041]" pageId="21" pageNumber="150">syn. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102CFFA7FF7CDE069C8F6BC0" pageId="21" pageNumber="150" type="diagnosis">
<paragraph id="7108511E102CFFA7FF7CDE069EBA6B78" blockId="21.[144,1104,1109,1236]" box="[144,252,1109,1132]" pageId="21" pageNumber="150">
<heading id="2A40E672102CFFA7FF7CDE069EBA6B78" box="[144,252,1109,1132]" fontSize="10" level="3" pageId="21" pageNumber="150" reason="5">
<emphasis id="43C38D0C102CFFA7FF7CDE069EBA6B78" box="[144,252,1109,1132]" italics="true" pageId="21" pageNumber="150">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102CFFA7FF7CDE2F9C8F6BC0" blockId="21.[144,1104,1109,1236]" pageId="21" pageNumber="150">
This species differs from the other
<taxonomicName id="B6B72A9D102CFFA7FDC7DE2F9CE86B80" authorityName="Erichson" authorityYear="1847" box="[555,686,1148,1172]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="43C38D0C102CFFA7FDC7DE2F9CE86B80" box="[555,686,1148,1172]" italics="true" pageId="21" pageNumber="150">Delopleurus</emphasis>
</taxonomicName>
species in having bilobate glabrous disc of pygidium with very thick borders similar in both sexes (
<figureCitation id="E98C4D9B102CFFA7FC98DECF9A4A6BA0" box="[884,1036,1180,1204]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="21" pageNumber="150">Figure 8B, E</figureCitation>
), and elytral striae with large punctures (
<figureCitation id="E98C4D9B102CFFA7FDF3DEEF9CFF6BC7" box="[543,697,1212,1236]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="21" pageNumber="150">Figure 8A, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102CFFA4FF7CDF4A9A1F6EB1" lastPageId="22" lastPageNumber="151" pageId="21" pageNumber="150" type="description">
<paragraph id="7108511E102CFFA7FF7CDF4A9F496A24" blockId="21.[144,1061,1305,1368]" box="[144,271,1305,1328]" pageId="21" pageNumber="150">
<heading id="2A40E672102CFFA7FF7CDF4A9F496A24" box="[144,271,1305,1328]" fontSize="10" level="3" pageId="21" pageNumber="150" reason="5">
<emphasis id="43C38D0C102CFFA7FF7CDF4A9F496A24" box="[144,271,1305,1328]" italics="true" pageId="21" pageNumber="150">Description</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102CFFA7FF7CDF139A636A4C" blockId="21.[144,1061,1305,1368]" box="[144,1061,1344,1368]" pageId="21" pageNumber="150">
Beetles are small (
<quantity id="B64FFCFB102CFFA7FEB3DF139FAE6A4C" box="[351,488,1344,1368]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.2" metricValueMin="4.5" pageId="21" pageNumber="150" unit="mm" value="4.85" valueMax="5.2" valueMin="4.5">4.5–5.2 mm</quantity>
), strongly convex, black or dark brown, glabrous.
</paragraph>
<paragraph id="7108511E102CFFA7FF7CDFD79CB46AC8" blockId="21.[144,1104,1412,1691]" pageId="21" pageNumber="150">
<emphasis id="43C38D0C102CFFA7FF7CDFD79E8C6A88" box="[144,202,1412,1436]" italics="true" pageId="21" pageNumber="150">Male</emphasis>
(
<figureCitation id="E98C4D9B102CFFA7FF0CDFD79F1C6A88" box="[224,346,1412,1436]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="21" pageNumber="150">Figure 8A</figureCitation>
). Clypeus quadridentate, medial teeth acute, lateral ones rightangled. Clypeal surface rugose in anterior part and laterally, frons densely punctate (punctures separated by less than puncture diameter).
</paragraph>
<paragraph id="7108511E102CFFA7FF56DFB79F06694F" blockId="21.[144,1104,1412,1691]" pageId="21" pageNumber="150">Pronotum about two times wider than long. Anterior and lateral margins distinctly bordered, base not or feebly bordered laterally. Surface of pronotum densely punctate anterolaterally, sparsely punctate with smaller punctures basally and anteromedially.</paragraph>
<paragraph id="7108511E102CFFA7FF56DC309A7C698F" blockId="21.[144,1104,1412,1691]" pageId="21" pageNumber="150">Elytra trapezoidal, as wide as long, shiny. Stria fine, with punctures much larger than striae width. Elytral intervals slightly convex to almost flat, with minute punctuation.</paragraph>
<paragraph id="7108511E102FFFA4FF12DAFB9D9A6FF4" blockId="22.[212,1172,168,320]" pageId="22" pageNumber="151">Anterior tibiae with three outer teeth, without a small acute tooth between first outer tooth and apical spur. Anterior tibial spur bifurcated apically.</paragraph>
<paragraph id="7108511E102FFFA4FF12DABB9FBD6E34" blockId="22.[212,1172,168,320]" pageId="22" pageNumber="151">
Pygidium with wide borders. Disc relatively small, somewhat bilobate, smooth and glabrous (
<figureCitation id="E98C4D9B102FFFA4FE95DB5B9FAD6E34" box="[377,491,264,288]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="22" pageNumber="151">Figure 8E</figureCitation>
).
</paragraph>
<paragraph id="7108511E102FFFA4FF12DB7B9DB76E54" blockId="22.[212,1172,168,320]" box="[254,1009,296,320]" pageId="22" pageNumber="151">
Parameres rounded in lateral view, without processes (
<figureCitation id="E98C4D9B102FFFA4FC81DB7B9DA76E54" box="[877,993,296,320]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="22" pageNumber="151">Figure 8C</figureCitation>
).
</paragraph>
<paragraph id="7108511E102FFFA4FF38DB3D9A1F6EB1" blockId="22.[212,1172,365,421]" pageId="22" pageNumber="151">
<emphasis id="43C38D0C102FFFA4FF38DB3D9F626E92" box="[212,292,366,390]" italics="true" pageId="22" pageNumber="151">Female</emphasis>
(
<figureCitation id="E98C4D9B102FFFA4FED8DB3E9FEF6E91" box="[308,425,365,389]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="22" pageNumber="151">Figure 8D</figureCitation>
) can be separated from male in having acute, not bifurcated spurs of anterior tibiae. Sculpture of pygidium is similar to that of male (
<figureCitation id="E98C4D9B102FFFA4FC3ADBDE9A0F6EB1" box="[982,1097,397,421]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="22" pageNumber="151">Figure 8E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102FFFA4FF38DB8F9A6B6CBB" pageId="22" pageNumber="151" type="materials_examined">
<paragraph id="7108511E102FFFA4FF38DB8F9F366EE7" blockId="22.[212,1172,475,666]" box="[212,368,475,499]" pageId="22" pageNumber="151">
<heading id="2A40E672102FFFA4FF38DB8F9F366EE7" box="[212,368,475,499]" fontSize="10" level="3" pageId="22" pageNumber="151" reason="5">
<emphasis id="43C38D0C102FFFA4FF38DB8F9F366EE7" box="[212,368,475,499]" italics="true" pageId="22" pageNumber="151">
<typeStatus id="AE0CEFBC102FFFA4FF38DB8F9F4A6EE7" box="[212,268,476,499]" pageId="22" pageNumber="151">Type</typeStatus>
material
</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102FFFA4FF38D8509F8B6D4E" blockId="22.[212,1172,475,666]" pageId="22" pageNumber="151">
<taxonomicName id="B6B72A9D102FFFA4FF38D8509FE16D0F" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[212,423,515,539]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="151" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="43C38D0C102FFFA4FF38D8509FE16D0F" box="[212,423,515,539]" italics="true" pageId="22" pageNumber="151">Delopleurus parvus</emphasis>
</taxonomicName>
–
<materialsCitation id="C1DF5B43102FFFA4FE2BD8509F8F6D4E" ID-GBIF-Occurrence="2849122332" collectionCode="MNHN" location="Bor" pageId="22" pageNumber="151" specimenCount="1" stateProvince="Bor" typeStatus="holotype">
<typeStatus id="AE0CEFBC102FFFA4FE2BD8509C6A6D0F" box="[455,556,515,539]" pageId="22" pageNumber="151" type="holotype">holotype</typeStatus>
,
<specimenCount id="67B19A97102FFFA4FDD5D8509CC56D0F" box="[569,643,515,539]" pageId="22" pageNumber="151" type="female">female</specimenCount>
with the labels ‘Ind. Bor.’, ‘Laforte 5821’, ‘ex Mus. D.Sharp 1890’, ‘Museum Paris 1952 Coll. R. Oberthür’, ‘
<taxonomicName id="B6B72A9D102FFFA4FC29D8719AD56D2E" authorityName="Sharp" authorityYear="1876" box="[965,1171,546,570]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="151" phylum="Arthropoda" rank="species" species="parva">Coptorhina parva</taxonomicName>
Type D.S.’ (
<collectionCode id="17A6C9DB102FFFA4FE8CD8119F866D4E" box="[352,448,578,602]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="151">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="7108511E102FFFA4FF12D8309C136D8E" blockId="22.[212,1172,475,666]" pageId="22" pageNumber="151">
<taxonomicName id="B6B72A9D102FFFA4FF12D8309FA76D6F" authorityName="Paulian" authorityYear="1934" box="[254,481,611,635]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="151" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="43C38D0C102FFFA4FF12D8309FA76D6F" box="[254,481,611,635]" italics="true" pageId="22" pageNumber="151">Delopleurus cardoni</emphasis>
</taxonomicName>
–
<materialsCitation id="C1DF5B43102FFFA4FDE5D8319C176D8E" ID-GBIF-Occurrence="2849122346" collectionCode="MNHN" collectorName="Chota-Nagpore Palkot R. P. Cardon" pageId="22" pageNumber="151" specimenCount="3" typeStatus="syntype">
<specimenCount id="67B19A97102FFFA4FDE5D8319CC26D6F" box="[521,644,610,635]" pageId="22" pageNumber="151" type="generic" typeStatus="syntypes">3 syntypes</specimenCount>
with the label ‘
<collectorName id="DC4234C8102FFFA4FCA7D8319F6F6D8E" pageId="22" pageNumber="151">Chota-Nagpore Palkot R.P. Cardon</collectorName>
VII-VIII 1897’ (
<collectionCode id="17A6C9DB102FFFA4FE05D8D19C0F6D8E" box="[489,585,642,666]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="151">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="7108511E102FFFA4FF38D8839FED6DFC" blockId="22.[212,1172,720,943]" box="[212,427,720,744]" pageId="22" pageNumber="151">
<heading id="2A40E672102FFFA4FF38D8839FED6DFC" box="[212,427,720,744]" fontSize="10" level="3" pageId="22" pageNumber="151" reason="5">
<emphasis id="43C38D0C102FFFA4FF38D8839FED6DFC" box="[212,427,720,744]" italics="true" pageId="22" pageNumber="151">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102FFFA4FF38D8A49A6B6CBB" blockId="22.[212,1172,720,943]" pageId="22" pageNumber="151">
<materialsCitation id="C1DF5B43102FFFA4FF38D8A49F7A6C3B" ID-GBIF-Occurrence="2849122302" collectionCode="IRSNB" collectorName="Chota Nagpore Nowatoli &amp; Cardon" country="India" location="INDIA" pageId="22" pageNumber="151" specimenCount="3" specimenCount-female="3">
<collectingCountry id="09A0118E102FFFA4FF38D8A49F6A6C1B" box="[212,300,759,783]" name="India" pageId="22" pageNumber="151">INDIA</collectingCountry>
:
<collectorName id="DC4234C8102FFFA4FED1D8A49C2D6C1B" box="[317,619,759,784]" pageId="22" pageNumber="151">Chota Nagpore Nowatoli</collectorName>
R.P.
<collectorName id="DC4234C8102FFFA4FD49D8A49D476C1B" box="[677,769,759,783]" pageId="22" pageNumber="151">Cardon</collectorName>
VIII-IX 1896,
<specimenCount id="67B19A97102FFFA4FC53D8A49A776C1B" box="[959,1073,759,783]" pageId="22" pageNumber="151" type="female">3 females</specimenCount>
, 6 spm. (
<collectionCode id="17A6C9DB102FFFA4FF30D9449F726C3B" box="[220,308,791,815]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="151">IRSNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C1DF5B43102FFFA4FEA7D9449A706C3B" ID-GBIF-Occurrence="2849122382" box="[331,1078,791,815]" collectingDate="1897-10" collectionCode="MNHN" collectorName="Chota-Nagpore Nawatoli &amp; Cardon" country="India" pageId="22" pageNumber="151" specimenCount="1">
<collectorName id="DC4234C8102FFFA4FEA7D9449C296C3B" box="[331,623,791,815]" pageId="22" pageNumber="151">Chota-Nagpore Nawatoli</collectorName>
R.P.
<collectorName id="DC4234C8102FFFA4FD4AD9449D446C3B" box="[678,770,791,815]" pageId="22" pageNumber="151">Cardon</collectorName>
<date id="050977DE102FFFA4FCE1D9449D266C3B" box="[781,864,791,815]" pageId="22" pageNumber="151" value="1897-10">
<collectingDate id="154D8E36102FFFA4FCE1D9449D266C3B" box="[781,864,791,815]" pageId="22" pageNumber="151" value="1897-10">X.1897</collectingDate>
</date>
, 2 spm. (
<collectionCode id="17A6C9DB102FFFA4FC23D9449A696C3B" box="[975,1071,791,815]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="151">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C1DF5B43102FFFA4FBA9D9449D066C5B" ID-GBIF-Occurrence="2849122369" collectionCode="MNHN" collectorName="Nagpore Nawatoli &amp; Cardon" country="India" pageId="22" pageNumber="151" specimenCount="1">
<collectorName id="DC4234C8102FFFA4FBA9D9449FE06C5B" pageId="22" pageNumber="151">Chota-Nagpore Nawatoli</collectorName>
R.P.
<collectorName id="DC4234C8102FFFA4FE35D9649C736C5B" box="[473,565,823,847]" pageId="22" pageNumber="151">Cardon</collectorName>
1898, 3 spm. (
<collectionCode id="17A6C9DB102FFFA4FD34D9649D7E6C5B" box="[728,824,823,847]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="151">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C1DF5B43102FFFA4FCA7D9649CDD6C7B" ID-GBIF-Occurrence="2849122348" collectionCode="MNHN" collectorName="Chota-Nagpore Barway &amp; Cardon" country="India" pageId="22" pageNumber="151" specimenCount="1">
<collectorName id="DC4234C8102FFFA4FCA7D9649A1C6C44" box="[843,1114,823,848]" pageId="22" pageNumber="151">Chota-Nagpore Barway</collectorName>
R.P.
<collectorName id="DC4234C8102FFFA4FF38D9049F6F6C7B" box="[212,297,855,879]" pageId="22" pageNumber="151">Cardon</collectorName>
VI-VII.1897, 1 spm. (
<collectionCode id="17A6C9DB102FFFA4FDD8D9049CD26C7B" box="[564,660,855,879]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="151">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C1DF5B43102FFFA4FD40D9049F396C9B" ID-GBIF-Occurrence="2849122385" collectionCode="MNHN" collectorName="Bangalore Chikmagalur Tabourel" country="India" pageId="22" pageNumber="151" specimenCount="1">
<collectorName id="DC4234C8102FFFA4FD40D9049A746C7B" box="[684,1074,855,879]" pageId="22" pageNumber="151">Bangalore Chikmagalur Tabourel</collectorName>
1900, 1 spm. (
<collectionCode id="17A6C9DB102FFFA4FEF4D9249F316C9B" box="[280,375,887,911]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="151">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C1DF5B43102FFFA4FE67D9249DEC6C9B" ID-GBIF-Occurrence="2849122338" box="[395,938,887,911]" collectionCode="MNHN" collectorName="Pulney Hills &amp; Castets" country="India" pageId="22" pageNumber="151" specimenCount="1">
<collectorName id="DC4234C8102FFFA4FE67D9249C526C9B" box="[395,532,887,911]" pageId="22" pageNumber="151">Pulney Hills</collectorName>
R.P.
<collectorName id="DC4234C8102FFFA4FDABD9249CD96C9B" box="[583,671,887,911]" pageId="22" pageNumber="151">Castets</collectorName>
1898, 3 spm. (
<collectionCode id="17A6C9DB102FFFA4FCADD9249DE76C9B" box="[833,929,887,911]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="151">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C1DF5B43102FFFA4FC58D9249A6F6CBB" ID-GBIF-Occurrence="2849122317" collectionCode="IRSNB" collectorName="Coll. R. I. Sc. N. B Ceylan &amp; Doherty Collection E. Candeze" country="Sri Lanka" location="SRI LANKA" pageId="22" pageNumber="151" specimenCount="1" specimenCount-female="1">
<collectingCountry id="09A0118E102FFFA4FC58D9249A146C9B" box="[948,1106,887,911]" name="Sri Lanka" pageId="22" pageNumber="151">SRI LANKA</collectingCountry>
:
<collectorName id="DC4234C8102FFFA4FBB0D9249FEE6CBB" pageId="22" pageNumber="151">Coll. R.I.Sc.N.B Ceylan</collectorName>
coll.
<collectorName id="DC4234C8102FFFA4FE06D9C49D0C6CBB" box="[490,842,919,943]" pageId="22" pageNumber="151">Doherty Collection E.Candeze</collectorName>
,
<specimenCount id="67B19A97102FFFA4FCBBD9C49DF16CBB" box="[855,951,919,943]" pageId="22" pageNumber="151" type="female">1 female</specimenCount>
(
<collectionCode id="17A6C9DB102FFFA4FC24D9C49A676CBB" box="[968,1057,919,943]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="151">IRSNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102FFFA4FF38D9B69AC46B70" pageId="22" pageNumber="151" type="distribution">
<paragraph id="7108511E102FFFA4FF38D9B69F1E6CE9" blockId="22.[212,1172,997,1124]" box="[212,344,997,1021]" pageId="22" pageNumber="151">
<heading id="2A40E672102FFFA4FF38D9B69F1E6CE9" box="[212,344,997,1021]" fontSize="10" level="3" pageId="22" pageNumber="151" reason="5">
<emphasis id="43C38D0C102FFFA4FF38D9B69F1E6CE9" box="[212,344,997,1021]" italics="true" pageId="22" pageNumber="151">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102FFFA4FF38DE5F9AC46B70" blockId="22.[212,1172,997,1124]" pageId="22" pageNumber="151">
The species is known from a few localities in the Indian Peninsula – southeastern Deccan
<collectingRegion id="B3739FFC102FFFA4FF38DE7F9F616B50" box="[212,295,1068,1092]" country="Nigeria" name="Plateau" pageId="22" pageNumber="151">Plateau</collectingRegion>
and Chota Nagpur
<collectingRegion id="B3739FFC102FFFA4FDE2DE7F9C236B50" box="[526,613,1068,1092]" country="Nigeria" name="Plateau" pageId="22" pageNumber="151">Plateau</collectingRegion>
. Locality map (
<figureCitation id="E98C4D9B102FFFA4FCF4DE7F9DCF6B50" box="[792,905,1068,1092]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="22" pageNumber="151">Figure 8F</figureCitation>
) suggests that it can be associated with transitional zones between moist and dry deciduous forest ecoregions.
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102FFFA4FF38DEC99A1A6A6D" pageId="22" pageNumber="151" type="discussion">
<paragraph id="7108511E102FFFA4FF38DEC99F706BA6" blockId="22.[212,1172,1178,1402]" box="[212,310,1178,1202]" pageId="22" pageNumber="151">
<heading id="2A40E672102FFFA4FF38DEC99F706BA6" box="[212,310,1178,1202]" fontSize="10" level="3" pageId="22" pageNumber="151" reason="5">
<emphasis id="43C38D0C102FFFA4FF38DEC99F706BA6" box="[212,310,1178,1202]" italics="true" pageId="22" pageNumber="151">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102FFFA4FF38DE929A1A6A6D" blockId="22.[212,1172,1178,1402]" pageId="22" pageNumber="151">
<bibRefCitation id="15262CEF102FFFA4FF38DE929F336BCD" author="Paulian R" box="[212,373,1217,1241]" pageId="22" pageNumber="151" pagination="162 - 164" refId="ref10236" refString="Paulian R. 1934. Quelques Panelini asiatiques nouveaux ou peu connus. Bull Soc Ent Fr. 39: 162 - 164." type="journal article" year="1934">Paulian (1934)</bibRefCitation>
described
<taxonomicName id="B6B72A9D102FFFA4FE06DE919C846BCE" authorityName="Paulian" authorityYear="1934" box="[490,706,1218,1242]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="151" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="43C38D0C102FFFA4FE06DE919C846BCE" box="[490,706,1218,1242]" italics="true" pageId="22" pageNumber="151">Delopleurus cardoni</emphasis>
</taxonomicName>
from nine specimens from northern
<collectingCountry id="09A0118E102FFFA4FBB5DE929AD56BCD" box="[1113,1171,1217,1241]" name="India" pageId="22" pageNumber="151">India</collectingCountry>
(‘Chota-Nagpore Nawatoli Palkot, R.P.Cardon VII-VIII.1897’). He separated
<taxonomicName id="B6B72A9D102FFFA4FBA2DEB19F456A0E" authorityName="Paulian" authorityYear="1934" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="151" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="43C38D0C102FFFA4FBA2DEB19F456A0E" italics="true" pageId="22" pageNumber="151">D.cardoni</emphasis>
</taxonomicName>
from
<taxonomicName id="B6B72A9D102FFFA4FEABDF519FEE6A0D" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[327,424,1282,1305]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="151" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="43C38D0C102FFFA4FEABDF519FEE6A0D" box="[327,424,1282,1305]" italics="true" pageId="22" pageNumber="151">D.parvus</emphasis>
</taxonomicName>
on the basis of the punctate disk of pronotum in the former species and smooth in the latter. Comparison of the
<typeStatus id="AE0CEFBC102FFFA4FD28DF719CBB6A2E" box="[708,765,1314,1338]" pageId="22" pageNumber="151">types</typeStatus>
of the both nominal species showed, however, that the both have relatively densely punctate disk of pronotum and do not differ significantly in other characters. The new synonymy is therefore established.
</paragraph>
</subSubSection>
</treatment>
</document>

312
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@ -0,0 +1,312 @@
<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE008102DFFA7FE0ED8B39DD36C5F" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus fossatus Frolov 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="150" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="149" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
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<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>49</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
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<mods:start>129</mods:start>
<mods:end>154</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2014.909072</mods:identifier>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">4002597</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331147" ID-GBIF-Taxon="167368587" ID-Zenodo-Dep="4331147" LSID="urn:lsid:plazi:treatment:F91EE008102DFFA7FE0ED8B39DD36C5F" httpUri="http://treatment.plazi.org/id/F91EE008102DFFA7FE0ED8B39DD36C5F" lastPageId="21" lastPageNumber="150" pageId="20" pageNumber="149">
<subSubSection box="[482,901,736,761]" pageId="20" pageNumber="149" type="nomenclature">
<paragraph blockId="20.[482,901,736,793]" box="[482,901,736,761]" pageId="20" pageNumber="149">
<heading box="[482,901,736,761]" centered="true" fontSize="10" level="2" pageId="20" pageNumber="149" reason="2">
<taxonomicName ID-CoL="34H56" authority="Frolov" authorityName="Frolov" authorityYear="2014" box="[482,798,736,761]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
<emphasis bold="true" box="[482,709,736,760]" italics="true" pageId="20" pageNumber="149">Delopleurus fossatus</emphasis>
Frolov
</taxonomicName>
,
<taxonomicNameLabel box="[811,901,738,761]" pageId="20" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[609,775,769,793]" pageId="20" pageNumber="149" type="description">
<paragraph blockId="20.[482,901,736,793]" box="[609,775,769,793]" pageId="20" pageNumber="149">
(
<figureCitation box="[618,766,769,793]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="20" pageNumber="149">Figure 7A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="149" type="diagnosis">
<paragraph blockId="20.[212,1172,828,988]" box="[212,320,828,851]" pageId="20" pageNumber="149">
<heading box="[212,320,828,851]" fontSize="10" level="3" pageId="20" pageNumber="149" reason="5">
<emphasis box="[212,320,828,851]" italics="true" pageId="20" pageNumber="149">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[212,1172,828,988]" pageId="20" pageNumber="149">
This species is similar to
<taxonomicName authorityName="Frolov" authorityYear="2014" box="[493,619,867,891]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="krikkeni">
<emphasis box="[493,619,867,891]" italics="true" pageId="20" pageNumber="149">D. krikkeni</emphasis>
</taxonomicName>
in having pygidium with deep circular fossa, but can be separated from it by having basal border of pygidium not or indistinctly widened medially, punctate disc of pronotum and smaller, slit-shaped dorsal parts of eyes.
</paragraph>
</subSubSection>
<subSubSection lastPageId="21" lastPageNumber="150" pageId="20" pageNumber="149" type="description">
<paragraph blockId="20.[212,1172,1054,1691]" box="[212,339,1054,1077]" pageId="20" pageNumber="149">
<heading box="[212,339,1054,1077]" fontSize="10" level="3" pageId="20" pageNumber="149" reason="5">
<emphasis box="[212,339,1054,1077]" italics="true" pageId="20" pageNumber="149">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[212,1172,1054,1691]" pageId="20" pageNumber="149">
<emphasis box="[212,403,1094,1118]" italics="true" pageId="20" pageNumber="149">
<typeStatus box="[212,317,1094,1118]" pageId="20" pageNumber="149">Holotype</typeStatus>
, male.
</emphasis>
Body strongly convex, black, glabrous, its length
<quantity box="[1073,1172,1093,1117]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="20" pageNumber="149" unit="mm" value="5.3">5.3 mm</quantity>
(
<figureCitation box="[220,337,1125,1149]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="20" pageNumber="149">Figure 7A</figureCitation>
).
</paragraph>
<paragraph blockId="20.[212,1172,1054,1691]" pageId="20" pageNumber="149">Clypeus quadridentate. Two medial teeth acute, lateral ones obtuse. Clypeus rugose in anterior part and laterally, frons densely punctate (punctures separated by one to three puncture diameters).</paragraph>
<paragraph blockId="20.[212,1172,1054,1691]" pageId="20" pageNumber="149">Pronotum about two times wider than long. Anterior and lateral margins distinctly bordered, base feebly bordered. Pronotum densely punctate (punctures separated by one to two puncture diameters on disc, becoming denser laterally).</paragraph>
<paragraph blockId="20.[212,1172,1054,1691]" pageId="20" pageNumber="149">Elytra trapezoidal, as wide as long, shiny. Stria relatively deep, with punctures 1.5–2 times larger than striae width. Elytral intervals convex, with minute punctation.</paragraph>
<paragraph blockId="20.[212,1172,1054,1691]" pageId="20" pageNumber="149">Anterior tibiae with three outer teeth, without a small acute tooth between first outer tooth and apical spur. Anterior tibial spur bifurcated apically.</paragraph>
<paragraph blockId="20.[212,1172,1054,1691]" pageId="20" pageNumber="149">
Pygidium with strongly convex disc separated by deep circular fossa from borders. Apical and basal borders about the same thickness, not significantly thicker in the middle. Disc glabrous and coarsely punctate (
<figureCitation box="[779,893,1540,1564]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="20" pageNumber="149">Figure 7B</figureCitation>
).
</paragraph>
<paragraph blockId="20.[212,1172,1054,1691]" box="[254,930,1571,1595]" pageId="20" pageNumber="149">
Parameres with small processes in lateral view (
<figureCitation box="[799,914,1571,1595]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="20" pageNumber="149">Figure 7C</figureCitation>
).
</paragraph>
<paragraph blockId="20.[212,1172,1054,1691]" pageId="20" pageNumber="149">
<emphasis box="[254,334,1604,1628]" italics="true" pageId="20" pageNumber="149">Female</emphasis>
can be separated from male in having acute, not bifurcated spurs of anterior tibiae. Sculpture of pygidium is similar to that of male but the pygidium is one-eighth to one-ninth shorter (
<figureCitation box="[584,698,1667,1691]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="20" pageNumber="149">Figure 7E</figureCitation>
).
</paragraph>
<paragraph blockId="21.[144,1104,168,224]" pageId="21" pageNumber="150">
<emphasis box="[144,262,169,192]" italics="true" pageId="21" pageNumber="150">
<typeStatus box="[144,259,169,192]" pageId="21" pageNumber="150">Paratypes</typeStatus>
.
</emphasis>
Body length varies from
<quantity box="[558,645,168,192]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="150" unit="mm" value="4.5">4.5 mm</quantity>
to
<quantity box="[687,777,168,192]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="21" pageNumber="150" unit="mm" value="5.5">5.5 mm</quantity>
, otherwise the
<typeStatus box="[950,1059,169,193]" pageId="21" pageNumber="150">paratypes</typeStatus>
are similar to the
<typeStatus box="[304,406,200,224]" pageId="21" pageNumber="150">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="150" type="materials_examined">
<paragraph blockId="21.[144,1104,292,579]" box="[144,300,292,316]" pageId="21" pageNumber="150">
<heading box="[144,300,292,316]" fontSize="10" level="3" pageId="21" pageNumber="150" reason="5">
<emphasis box="[144,300,292,316]" italics="true" pageId="21" pageNumber="150">Type material</emphasis>
</heading>
</paragraph>
<paragraph blockId="21.[144,1104,292,579]" pageId="21" pageNumber="150">
<materialsCitation ID-GBIF-Occurrence="2849122335" collectingDate="1913-02-11" collectionCode="BMNH" collectorName="A. Neave" country="Malawi" location="Nyassaland Mlanje" pageId="21" pageNumber="150" specimenCount="1" stateProvince="Mulanje" typeStatus="holotype">
<typeStatus box="[144,253,332,356]" pageId="21" pageNumber="150">Holotype</typeStatus>
,
<specimenCount box="[264,318,332,356]" pageId="21" pageNumber="150" type="male">male</specimenCount>
with label ‘
<location LSID="urn:lsid:plazi:treatment:F91EE008102DFFA7FE0ED8B39DD36C5F:746807C5102CFFA7FE2CDB1F9CDC6E70" box="[448,666,332,356]" country="Malawi" name="Nyassaland Mlanje" pageId="21" pageNumber="150" stateProvince="Mulanje">Nyassaland Mlanje</location>
[
<collectingCountry box="[679,804,332,356]" name="Malawi" pageId="21" pageNumber="150">MALAWI</collectingCountry>
,
<collectingRegion box="[815,911,332,356]" country="Malawi" name="Mulanje" pageId="21" pageNumber="150">Mulanje</collectingRegion>
]
<date box="[924,1075,332,356]" pageId="21" pageNumber="150" value="1913-02-11">
<collectingDate box="[924,1075,332,356]" pageId="21" pageNumber="150" value="1913-02-11">Feb. 11, 1913</collectingDate>
</date>
S.
<collectorName box="[144,243,364,388]" pageId="21" pageNumber="150">A.Neave</collectorName>
1913-140’ (
<collectionCode box="[377,469,364,388]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="21" pageNumber="150">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122386" box="[490,1099,364,389]" collectingDate="1913-02-11" collectionCode="BMNH" collectorName="A. Neave" country="Malawi" location="Nyassaland Mlanje" pageId="21" pageNumber="150" specimenCount="1" specimenCount-male="1" stateProvince="Mulanje" typeStatus="paratype">
<typeStatus box="[490,604,365,389]" pageId="21" pageNumber="150">Paratypes</typeStatus>
:
<specimenCount box="[615,691,364,388]" pageId="21" pageNumber="150" type="male">1 male</specimenCount>
with the same data as the holotype
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122351" collectingDate="1913-02-27" collectionCode="BMNH" collectorName="A. Neave" country="Malawi" location="Nyassaland Mlanje" pageId="21" pageNumber="150" specimenCount="13" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[144,291,396,420]" pageId="21" pageNumber="150" type="generic">13 specimens</specimenCount>
with the same data as the holotype except for the date of collecting:
<date pageId="21" pageNumber="150" value="1913-02-27">
<collectingDate pageId="21" pageNumber="150" value="1913-02-27">27. II.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122372" box="[245,605,428,452]" collectingDate="1913-02-24" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-female="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[245,347,428,452]" pageId="21" pageNumber="150" type="female">1 female</specimenCount>
(
<collectionCode box="[370,462,428,452]" pageId="21" pageNumber="150">BMNH</collectionCode>
),
<date box="[488,605,428,452]" pageId="21" pageNumber="150" value="1913-02-24">
<collectingDate box="[488,605,428,452]" pageId="21" pageNumber="150" value="1913-02-24">24.II.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122358" box="[624,983,428,452]" collectingDate="1913-02-14" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-female="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[624,725,428,452]" pageId="21" pageNumber="150" type="female">1 female</specimenCount>
(
<collectionCode box="[748,840,428,452]" pageId="21" pageNumber="150">BMNH</collectionCode>
),
<date box="[866,983,428,452]" pageId="21" pageNumber="150" value="1913-02-14">
<collectingDate box="[866,983,428,452]" pageId="21" pageNumber="150" value="1913-02-14">14.II.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122361" collectingDate="1913-03-07" collectionCode="ZIN" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-female="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[1002,1103,428,452]" pageId="21" pageNumber="150" type="female">1 female</specimenCount>
(
<collectionCode box="[152,205,460,484]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="21" pageNumber="150" type="Museum">ZIN</collectionCode>
),
<date box="[232,345,460,484]" pageId="21" pageNumber="150" value="1913-03-07">
<collectingDate box="[232,345,460,484]" pageId="21" pageNumber="150" value="1913-03-07">7.III.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122312" box="[365,726,460,484]" collectingDate="1912-12-19" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-male="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[365,447,460,484]" pageId="21" pageNumber="150" type="male">1 male</specimenCount>
(
<collectionCode box="[469,561,460,484]" pageId="21" pageNumber="150">BMNH</collectionCode>
),
<date box="[588,726,460,484]" pageId="21" pageNumber="150" value="1912-12-19">
<collectingDate box="[588,726,460,484]" pageId="21" pageNumber="150" value="1912-12-19">19.XII.1912</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122363" box="[744,1071,460,484]" collectingDate="1913-01-07" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="2" specimenCount-male="2" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[744,836,460,484]" pageId="21" pageNumber="150" type="male">2 males</specimenCount>
(
<collectionCode box="[859,951,460,484]" pageId="21" pageNumber="150">BMNH</collectionCode>
),
<date box="[978,1071,460,484]" pageId="21" pageNumber="150" value="1913-01-07">
<collectingDate box="[978,1071,460,484]" pageId="21" pageNumber="150" value="1913-01-07">7.I.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122353" collectingDate="1913-02-20" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-male="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount pageId="21" pageNumber="150" type="male">1 male</specimenCount>
(
<collectionCode box="[222,314,491,515]" pageId="21" pageNumber="150">BMNH</collectionCode>
),
<date box="[342,459,491,515]" pageId="21" pageNumber="150" value="1913-02-20">
<collectingDate box="[342,459,491,515]" pageId="21" pageNumber="150" value="1913-02-20">20.II.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122329" box="[480,821,491,515]" collectingDate="1913-03-05" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-male="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[480,563,491,515]" pageId="21" pageNumber="150" type="male">1 male</specimenCount>
(
<collectionCode box="[587,679,491,515]" pageId="21" pageNumber="150">BMNH</collectionCode>
),
<date box="[708,821,491,515]" pageId="21" pageNumber="150" value="1913-03-05">
<collectingDate box="[708,821,491,515]" pageId="21" pageNumber="150" value="1913-03-05">5.III.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122356" collectingDate="1913-01-12" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-male="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[842,925,491,515]" pageId="21" pageNumber="150" type="male">1 male</specimenCount>
(
<collectionCode box="[949,1041,491,515]" pageId="21" pageNumber="150">BMNH</collectionCode>
),
<date pageId="21" pageNumber="150" value="1913-01-12">
<collectingDate pageId="21" pageNumber="150" value="1913-01-12">12. I.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122342" box="[233,512,523,547]" collectingDate="1913-02-01" collectionCode="ZIN" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-male="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[233,311,523,547]" pageId="21" pageNumber="150" type="male">1 male</specimenCount>
(
<collectionCode box="[331,384,523,547]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="21" pageNumber="150" type="Museum">ZIN</collectionCode>
),
<date box="[408,512,523,547]" pageId="21" pageNumber="150" value="1913-02-01">
<collectingDate box="[408,512,523,547]" pageId="21" pageNumber="150" value="1913-02-01">1.II.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122364" box="[528,865,523,547]" collectingDate="1913-02-08" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-female="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[528,626,523,547]" pageId="21" pageNumber="150" type="female">1 female</specimenCount>
(
<collectionCode box="[646,738,523,547]" pageId="21" pageNumber="150">BMNH</collectionCode>
),
<date box="[762,865,523,547]" pageId="21" pageNumber="150" value="1913-02-08">
<collectingDate box="[762,865,523,547]" pageId="21" pageNumber="150" value="1913-02-08">8.II.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122387" collectingDate="1913-01-21" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-female="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[881,980,523,547]" pageId="21" pageNumber="150" type="female">1 female</specimenCount>
(
<collectionCode box="[999,1091,523,547]" pageId="21" pageNumber="150">BMNH</collectionCode>
),
<date box="[144,251,555,579]" pageId="21" pageNumber="150" value="1913-01-21">
<collectingDate box="[144,251,555,579]" pageId="21" pageNumber="150" value="1913-01-21">21.I.1913</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2849122318" box="[264,478,555,579]" collectionCode="BMNH" country="Malawi" pageId="21" pageNumber="150" specimenCount="1" specimenCount-female="1" stateProvince="Mulanje" typeStatus="paratype">
<specimenCount box="[264,360,555,579]" pageId="21" pageNumber="150" type="female">1 female</specimenCount>
(
<collectionCode box="[377,469,555,579]" pageId="21" pageNumber="150">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[144,1102,648,711]" pageId="21" pageNumber="150" type="distribution">
<paragraph blockId="21.[144,1102,648,711]" box="[144,276,648,672]" pageId="21" pageNumber="150">
<heading box="[144,276,648,672]" fontSize="10" level="3" pageId="21" pageNumber="150" reason="5">
<emphasis box="[144,276,648,672]" italics="true" pageId="21" pageNumber="150">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="21.[144,1102,648,711]" box="[144,1102,687,711]" pageId="21" pageNumber="150">
The species is known from Mlanje,
<collectingCountry box="[553,639,687,711]" name="Malawi" pageId="21" pageNumber="150">Malawi</collectingCountry>
(
<figureCitation box="[657,771,687,711]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="21" pageNumber="150">Figure 7F</figureCitation>
) in the southern Rift Valley.
</paragraph>
</subSubSection>
<subSubSection box="[144,917,780,843]" pageId="21" pageNumber="150" type="etymology">
<paragraph blockId="21.[144,917,780,843]" box="[144,265,780,804]" pageId="21" pageNumber="150">
<heading box="[144,265,780,804]" fontSize="10" level="3" pageId="21" pageNumber="150" reason="5">
<emphasis box="[144,265,780,804]" italics="true" pageId="21" pageNumber="150">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="21.[144,917,780,843]" box="[144,917,819,843]" pageId="21" pageNumber="150">The epithet is a Latin word denoting the sculpture of the pygidium.</paragraph>
</subSubSection>
</treatment>
</document>

182
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<mods:title id="A282ADA996B918BDA207C4406DEE6B49">Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
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<treatment id="F91EE008102EFFAAFE9BDE049DB96E5C" ID-DOI="http://doi.org/10.5281/zenodo.4331163" ID-GBIF-Taxon="167368581" ID-Zenodo-Dep="4331163" LSID="urn:lsid:plazi:treatment:F91EE008102EFFAAFE9BDE049DB96E5C" httpUri="http://treatment.plazi.org/id/F91EE008102EFFAAFE9BDE049DB96E5C" lastPageId="24" lastPageNumber="153" pageId="23" pageNumber="152">
<subSubSection id="39AD0295102EFFA5FE9BDE049D2F6B64" box="[375,873,1111,1136]" pageId="23" pageNumber="152" type="nomenclature">
<paragraph id="7108511E102EFFA5FE9BDE049D2F6B64" blockId="23.[375,873,1111,1168]" box="[375,873,1111,1136]" pageId="23" pageNumber="152">
<heading id="2A40E672102EFFA5FE9BDE049D2F6B64" box="[375,873,1111,1136]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="152" reason="2">
<taxonomicName id="B6B72A9D102EFFA5FE9BDE049C9D6B64" ID-CoL="428Y3" authority="(Frey)" authorityName="Frolov" authorityYear="2014" baseAuthorityName="Frolov" baseAuthorityYear="1963" box="[375,731,1111,1136]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi" status="comb. nov.">
<emphasis id="43C38D0C102EFFA5FE9BDE049CCC6B7B" bold="true" box="[375,650,1111,1135]" italics="true" pageId="23" pageNumber="152">Metacatharsius janssensi</emphasis>
(Frey)
</taxonomicName>
,
<taxonomicNameLabel id="58F03077102EFFA5FD05DE0B9D2F6B64" box="[745,873,1112,1136]" pageId="23" pageNumber="152" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102EFFA5FDC2DE2B9CF46B84" box="[558,690,1144,1168]" pageId="23" pageNumber="152" type="description">
<paragraph id="7108511E102EFFA5FDC2DE2B9CF46B84" blockId="23.[375,873,1111,1168]" box="[558,690,1144,1168]" pageId="23" pageNumber="152">
(
<figureCitation id="E98C4D9B102EFFA5FDDBDE2B9CEF6B84" box="[567,681,1144,1168]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="23" pageNumber="152">Figure 9E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102EFFA5FF7CDEF69D966AE8" pageId="23" pageNumber="152" type="reference_group">
<paragraph id="7108511E102EFFA5FF7CDEF69FB96BA9" blockId="23.[144,511,1189,1214]" box="[144,511,1189,1214]" pageId="23" pageNumber="152">
<heading id="2A40E672102EFFA5FF7CDEF69FB96BA9" box="[144,511,1189,1214]" fontSize="10" level="3" pageId="23" pageNumber="152" reason="5">
<taxonomicName id="B6B72A9D102EFFA5FF7CDEF69FB96BA9" ID-CoL="34H58" authority="Frey, 1963" authorityName="Frey" authorityYear="1963" box="[144,511,1189,1214]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FF7CDEF69F3A6BA9" box="[144,380,1189,1213]" italics="true" pageId="23" pageNumber="152">Delopleurus janssensi</emphasis>
<bibRefCitation id="15262CEF102EFFA5FE69DEF59FB96BA9" author="Frey G" box="[389,511,1189,1214]" pageId="23" pageNumber="152" pagination="538 - 549" refId="ref9887" refString="Frey G. 1963. Neue Onthophagen (Col. Scarab.) aus Australien (Queenslandausbeute von H. Demarz) und aus Afrika. Ent Arb Mus Frey. 14: 538 - 549." type="journal article" year="1963">Frey, 1963</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7108511E102EFFA5FF7CDEB69D966AE8" blockId="23.[144,1104,1253,1691]" pageId="23" pageNumber="152">
<taxonomicName id="B6B72A9D102EFFA5FF7CDEB69F3D6BE9" ID-CoL="34H58" authorityName="Frey" authorityYear="1963" box="[144,379,1253,1277]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FF7CDEB69F3D6BE9" box="[144,379,1253,1277]" italics="true" pageId="23" pageNumber="152">Delopleurus janssensi</emphasis>
</taxonomicName>
was described from two specimens from Warder,
<collectingRegion id="B3739FFC102EFFA5FC5EDEB69A166BE9" box="[946,1104,1253,1277]" country="Ethiopia" name="Sumale" pageId="23" pageNumber="152">Somali region</collectingRegion>
of
<collectingCountry id="09A0118E102EFFA5FF5FDF579F5E6A08" box="[179,280,1284,1308]" name="Ethiopia" pageId="23" pageNumber="152">Ethiopia</collectingCountry>
. The
<typeStatus id="AE0CEFBC102EFFA5FE8DDF579F806A08" box="[353,454,1284,1308]" pageId="23" pageNumber="152" type="holotype">holotype</typeStatus>
, female, has all the diagnostic characters of the genus
<taxonomicName id="B6B72A9D102EFFA5FF7CDF769FDD6A28" ID-CoL="5Q6D" authority="Paulian" authorityName="Paulian" authorityYear="1939" box="[144,411,1316,1341]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="43C38D0C102EFFA5FF7CDF769F7D6A29" box="[144,315,1317,1341]" italics="true" pageId="23" pageNumber="152">Metacatharsius</emphasis>
Paulian
</taxonomicName>
(large dorsal eye parts can be seen on the habitual photo,
<figureCitation id="E98C4D9B102EFFA5FF7CDF179F446A48" box="[144,258,1348,1372]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="23" pageNumber="152">Figure 9E</figureCitation>
). The
<typeStatus id="AE0CEFBC102EFFA5FEA3DF169FF46A49" box="[335,434,1349,1373]" pageId="23" pageNumber="152" type="paratype">paratype</typeStatus>
is similar to and apparently conspecific with the
<typeStatus id="AE0CEFBC102EFFA5FC0ADF179A0A6A48" box="[998,1100,1348,1372]" pageId="23" pageNumber="152" type="holotype">holotype</typeStatus>
. In the original description, the author compared this species with
<taxonomicName id="B6B72A9D102EFFA5FC63DF369DB36A68" ID-CoL="6CHFF" authorityName="Boheman" authorityYear="1857" box="[911,1013,1380,1404]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="pullus">
<emphasis id="43C38D0C102EFFA5FC63DF369DB36A68" box="[911,1013,1380,1404]" italics="true" pageId="23" pageNumber="152">D. pullus</emphasis>
</taxonomicName>
and
<taxonomicName id="B6B72A9D102EFFA5FBD8DF369E886A88" authorityName="Frolov" authorityYear="2014" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis id="43C38D0C102EFFA5FBD8DF369E886A88" italics="true" pageId="23" pageNumber="152">D. gilleti</emphasis>
</taxonomicName>
but did not provide characters specific for the genus
<emphasis id="43C38D0C102EFFA5FCCADFD79DF76A88" box="[806,945,1412,1436]" italics="true" pageId="23" pageNumber="152">
<taxonomicName id="B6B72A9D102EFFA5FCCADFD79DEB6A88" ID-CoL="42PQ" authorityName="Erichson" authorityYear="1847" box="[806,941,1412,1436]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">Delopleurus</taxonomicName>
.
</emphasis>
However, the original description and locality data agree well with the type specimens. There is no doubt that
<taxonomicName id="B6B72A9D102EFFA5FEF5DF979C4F6AC8" ID-CoL="34H58" authorityName="Frey" authorityYear="1963" box="[281,521,1476,1500]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FEF5DF979C4F6AC8" box="[281,521,1476,1500]" italics="true" pageId="23" pageNumber="152">Delopleurus janssensi</emphasis>
</taxonomicName>
was described from misidentified
<taxonomicName id="B6B72A9D102EFFA5FC49DF979A166AC8" ID-CoL="5Q6D" authorityName="Paulian" authorityYear="1939" box="[933,1104,1476,1500]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="43C38D0C102EFFA5FC49DF979A166AC8" box="[933,1104,1476,1500]" italics="true" pageId="23" pageNumber="152">Metacatharsius</emphasis>
</taxonomicName>
specimens and the new generic combination is therefore established here.
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102EFFA5FF56DC579CE0698F" pageId="23" pageNumber="152" type="description">
<paragraph id="7108511E102EFFA5FF56DC579CE0698F" blockId="23.[144,1104,1253,1691]" pageId="23" pageNumber="152">
It is possible that
<taxonomicName id="B6B72A9D102EFFA5FE68DC569C4B6908" authorityName="Frey" authorityYear="1963" box="[388,525,1541,1564]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FE68DC569C4B6908" box="[388,525,1541,1564]" italics="true" pageId="23" pageNumber="152">M. janssensi</emphasis>
</taxonomicName>
is conspecific with another described species of the genus. There are up to 18 species of this genus known from adjacent
<collectingCountry id="09A0118E102EFFA5FC18DC709A09692F" box="[1012,1103,1571,1595]" name="Somalia" pageId="23" pageNumber="152">Somalia</collectingCountry>
(
<bibRefCitation id="15262CEF102EFFA5FF7BDC109F74694F" author="Ferreira M" box="[151,306,1603,1627]" pageId="23" pageNumber="152" pagination="5 - 1088" refId="ref9860" refString="Ferreira M. 1972. Os escarabideos de Africa (Sul do Saara), I. Rev Ent Mocambique. 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972</bibRefCitation>
) and the genus is currently a subject of taxonomic revision (François Génier, pers. comm.). It is out of the scope of the present work to specify the taxonomic status of
<taxonomicName id="B6B72A9D102EFFA5FE95DCD79C45698F" authorityName="Frey" authorityYear="1963" box="[377,515,1668,1691]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FE95DCD79C45698F" box="[377,515,1668,1691]" italics="true" pageId="23" pageNumber="152">M. janssensi</emphasis>
</taxonomicName>
more exactly.
</paragraph>
</subSubSection>
<subSubSection id="39AD02951021FFAAFF38DAFA9DB96E5C" pageId="24" pageNumber="153" type="materials_examined">
<paragraph id="7108511E1021FFAAFF38DAFA9F366FD4" blockId="24.[212,1172,168,328]" box="[212,368,168,192]" pageId="24" pageNumber="153">
<heading id="2A40E6721021FFAAFF38DAFA9F366FD4" box="[212,368,168,192]" fontSize="10" level="3" pageId="24" pageNumber="153" reason="5">
<emphasis id="43C38D0C1021FFAAFF38DAFA9F366FD4" box="[212,368,168,192]" italics="true" pageId="24" pageNumber="153">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E1021FFAAFF38DA839DB96E5C" blockId="24.[212,1172,168,328]" pageId="24" pageNumber="153">
<materialsCitation id="C1DF5B431021FFAAFF38DA839A1D6E1C" ID-GBIF-Occurrence="2849122365" collectionCode="BMNH" determinerName="G. Frey" location="Wardere Somalia'" municipality="Wardere" pageId="24" pageNumber="153" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="AE0CEFBC1021FFAAFF38DA839F076FFC" box="[212,321,208,232]" pageId="24" pageNumber="153" type="holotype">Holotype</typeStatus>
,
<specimenCount id="67B19A971021FFAAFEA0DA839FD06FFC" box="[332,406,208,232]" pageId="24" pageNumber="153" type="female">female</specimenCount>
with the labels: ‘
<location id="746807C51021FFAAFDB4DA839D626FFC" LSID="urn:lsid:plazi:treatment:F91EE008102EFFAAFE9BDE049DB96E5C:746807C51021FFAAFDB4DA839D626FFC" box="[600,804,208,232]" municipality="Wardere" name="Wardere Somalia'" pageId="24" pageNumber="153">Wardere Somalia’</location>
, ‘Holotype’, ‘TYPE’, ‘Brit. Mus 1963-277’, ‘
<taxonomicName id="B6B72A9D1021FFAAFEB1DAA39C106E1C" authorityName="Frey" authorityYear="1963" box="[349,598,240,264]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="153" phylum="Arthropoda" rank="species" species="janssensi">Delopleurus janssensi</taxonomicName>
det.
<determinerName id="84D4BDDA1021FFAAFD70DAA39CA86E1C" box="[668,750,240,264]" pageId="24" pageNumber="153">
G.
<bibRefCitation id="15262CEF1021FFAAFD59DAA39CA86E1C" author="Frey G" box="[693,750,240,264]" pageId="24" pageNumber="153" pagination="538 - 549" refId="ref9887" refString="Frey G. 1963. Neue Onthophagen (Col. Scarab.) aus Australien (Queenslandausbeute von H. Demarz) und aus Afrika. Ent Arb Mus Frey. 14: 538 - 549." type="journal article" year="1963">Frey</bibRefCitation>
</determinerName>
1963, n. sp. Typus’ (
<collectionCode id="17A6C9DB1021FFAAFC16DAA39A156E1C" box="[1018,1107,240,264]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="24" pageNumber="153">BMNH</collectionCode>
)
</materialsCitation>
and
<materialsCitation id="C1DF5B431021FFAAFF38DB429DB96E5C" ID-GBIF-Occurrence="3757906302" collectingDate="1943-10" collectionCode="NHMB" country="Somalia" determinerName="G. Frey" location="Delopleurus" pageId="24" pageNumber="153" specimenCount="1" typeStatus="paratype">
<typeStatus id="AE0CEFBC1021FFAAFF38DB429F716E3D" box="[212,311,273,297]" pageId="24" pageNumber="153" type="paratype">paratype</typeStatus>
with the labels: ‘Wardere
<collectingCountry id="09A0118E1021FFAAFD2EDB439D5B6E3C" box="[706,797,272,296]" name="Somalia" pageId="24" pageNumber="153">Somalia</collectingCountry>
<date id="050977DE1021FFAAFCD5DB439DFB6E3C" box="[825,957,272,296]" pageId="24" pageNumber="153" value="1943-10">
<collectingDate id="154D8E361021FFAAFCD5DB439DFB6E3C" box="[825,957,272,296]" pageId="24" pageNumber="153" value="1943-10">Oct. 1943</collectingDate>
</date>
, T.H.E.Jackson’, ‘
<taxonomicName id="B6B72A9D1021FFAAFF37DB639F896E5C" authorityName="Frey" authorityYear="1963" box="[219,463,304,328]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="153" phylum="Arthropoda" rank="species" species="janssensi">
<location id="746807C51021FFAAFF37DB639F226E5C" LSID="urn:lsid:plazi:treatment:F91EE008102EFFAAFE9BDE049DB96E5C:746807C51021FFAAFF37DB639F226E5C" box="[219,356,304,328]" country="Somalia" name="Delopleurus" pageId="24" pageNumber="153">Delopleurus</location>
janssensi
</taxonomicName>
det.
<determinerName id="84D4BDDA1021FFAAFDE6DB639C1A6E5C" box="[522,604,304,328]" pageId="24" pageNumber="153">
G.
<bibRefCitation id="15262CEF1021FFAAFDCFDB639C1A6E5C" author="Frey G" box="[547,604,304,328]" pageId="24" pageNumber="153" pagination="538 - 549" refId="ref9887" refString="Frey G. 1963. Neue Onthophagen (Col. Scarab.) aus Australien (Queenslandausbeute von H. Demarz) und aus Afrika. Ent Arb Mus Frey. 14: 538 - 549." type="journal article" year="1963">Frey</bibRefCitation>
</determinerName>
1963, p. Typus’, ‘Cotype’ (
<collectionCode id="17A6C9DB1021FFAAFC7BDB639DB56E5C" box="[919,1011,304,328]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="24" pageNumber="153">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

165
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<mods:title id="1A6A3A25FB598F70F3798143B643290A">Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
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<treatment id="F91EE008102FFFA5FDF0DFF39F966B34" ID-DOI="http://doi.org/10.5281/zenodo.4331145" ID-GBIF-Taxon="167368586" ID-Zenodo-Dep="4331145" LSID="urn:lsid:plazi:treatment:F91EE008102FFFA5FDF0DFF39F966B34" httpUri="http://treatment.plazi.org/id/F91EE008102FFFA5FDF0DFF39F966B34" lastPageId="23" lastPageNumber="152" pageId="22" pageNumber="151">
<subSubSection id="39AD0295102FFFA4FDF0DFF39D0A6AAD" box="[540,844,1440,1465]" pageId="22" pageNumber="151" type="nomenclature">
<paragraph id="7108511E102FFFA4FDF0DFF39D0A6AAD" blockId="22.[540,844,1440,1497]" box="[540,844,1440,1465]" pageId="22" pageNumber="151">
<heading id="2A40E672102FFFA4FDF0DFF39D0A6AAD" box="[540,844,1440,1465]" centered="true" fontSize="10" level="2" pageId="22" pageNumber="151" reason="2">
<taxonomicName id="B6B72A9D102FFFA4FDF0DFF39D0A6AAD" ID-CoL="34H5F" authority="Arrow" authorityName="Arrow" box="[540,844,1440,1465]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="151" phylum="Arthropoda" rank="species" species="striatus">
<emphasis id="43C38D0C102FFFA4FDF0DFF39CBF6AAC" bold="true" box="[540,761,1440,1464]" italics="true" pageId="22" pageNumber="151">Delopleurus striatus</emphasis>
Arrow
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102FFFA4FDB3DF939D4F6ACC" box="[607,777,1472,1497]" pageId="22" pageNumber="151" type="description">
<paragraph id="7108511E102FFFA4FDB3DF939D4F6ACC" blockId="22.[540,844,1440,1497]" box="[607,777,1472,1497]" pageId="22" pageNumber="151">
(
<figureCitation id="E98C4D9B102FFFA4FD84DF939D466ACC" box="[616,768,1472,1497]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="22" pageNumber="151">Figure 9A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102FFFA4FF38DFBD9C046912" box="[212,578,1518,1542]" pageId="22" pageNumber="151" type="reference_group">
<paragraph id="7108511E102FFFA4FF38DFBD9C046912" blockId="22.[212,578,1518,1542]" box="[212,578,1518,1542]" pageId="22" pageNumber="151">
<heading id="2A40E672102FFFA4FF38DFBD9C046912" box="[212,578,1518,1542]" fontSize="10" level="3" pageId="22" pageNumber="151" reason="5">
<taxonomicName id="B6B72A9D102FFFA4FF38DFBD9C046912" ID-CoL="34H5F" authority="Arrow 1931" authorityName="Arrow" authorityYear="1931" box="[212,578,1518,1542]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="151" phylum="Arthropoda" rank="species" species="striatus">
<emphasis id="43C38D0C102FFFA4FF38DFBD9FF46912" box="[212,434,1518,1542]" italics="true" pageId="22" pageNumber="151">Delopleurus striatus</emphasis>
<bibRefCitation id="15262CEF102FFFA4FE56DFBD9C046912" author="Arrow GJ" box="[442,578,1518,1542]" pageId="22" pageNumber="151" refId="ref9736" refString="Arrow GJ. 1931. The Fauna of British India III, Coprinae. London: Taylor &amp; Francis." type="book" year="1931">Arrow 1931</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102FFFA4FF38DC6F9DCA698F" pageId="22" pageNumber="151" type="diagnosis">
<paragraph id="7108511E102FFFA4FF38DC6F9F066947" blockId="22.[212,1172,1596,1691]" box="[212,320,1596,1619]" pageId="22" pageNumber="151">
<heading id="2A40E672102FFFA4FF38DC6F9F066947" box="[212,320,1596,1619]" fontSize="10" level="3" pageId="22" pageNumber="151" reason="5">
<emphasis id="43C38D0C102FFFA4FF38DC6F9F066947" box="[212,320,1596,1619]" italics="true" pageId="22" pageNumber="151">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102FFFA4FF38DC309DCA698F" blockId="22.[212,1172,1596,1691]" pageId="22" pageNumber="151">
This species (known from a single female) differs from other
<taxonomicName id="B6B72A9D102FFFA4FC76DC309A5B696F" authorityName="Erichson" authorityYear="1847" box="[922,1053,1635,1659]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="43C38D0C102FFFA4FC76DC309A5B696F" box="[922,1053,1635,1659]" italics="true" pageId="22" pageNumber="151">Delopleurus</emphasis>
</taxonomicName>
species in having characteristic granulate disc of pygidium (
<figureCitation id="E98C4D9B102FFFA4FCE5DCD09D3A698F" box="[777,892,1667,1691]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="22" pageNumber="151">Figure 9C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102EFFA5FF7CDAFA9F7B6D9B" pageId="23" pageNumber="152" type="description">
<paragraph id="7108511E102EFFA5FF7CDAFA9F496FD4" blockId="23.[144,1104,169,583]" box="[144,271,169,192]" pageId="23" pageNumber="152">
<heading id="2A40E672102EFFA5FF7CDAFA9F496FD4" box="[144,271,169,192]" fontSize="10" level="3" pageId="23" pageNumber="152" reason="5">
<emphasis id="43C38D0C102EFFA5FF7CDAFA9F496FD4" box="[144,271,169,192]" italics="true" pageId="23" pageNumber="152">Description</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102EFFA5FF7CDA839DB66FFC" blockId="23.[144,1104,169,583]" box="[144,1008,208,232]" pageId="23" pageNumber="152">
<emphasis id="43C38D0C102EFFA5FF7CDA839EA16FFC" box="[144,231,208,232]" italics="true" pageId="23" pageNumber="152">Female.</emphasis>
Body strongly convex, black, glabrous, length
<quantity id="B64FFCFB102EFFA5FCEFDA839D1C6FFC" box="[771,858,208,232]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="23" pageNumber="152" unit="mm" value="5.1">5.1 mm</quantity>
(
<figureCitation id="E98C4D9B102EFFA5FC87DA839DA66FFC" box="[875,992,208,232]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="23" pageNumber="152">Figure 9A</figureCitation>
).
</paragraph>
<paragraph id="7108511E102EFFA5FF56DAA39C4D6E5C" blockId="23.[144,1104,169,583]" pageId="23" pageNumber="152">
Clypeus appear bidentate with two medial teeth acute (
<figureCitation id="E98C4D9B102EFFA5FCB3DAA39D916E1C" box="[863,983,240,264]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="23" pageNumber="152">Figure 9B</figureCitation>
). Clypeus rugose in anterior part and laterally, frons densely punctate (punctures separated by one to three puncture diameters).
</paragraph>
<paragraph id="7108511E102EFFA5FF56DB039D966EB3" blockId="23.[144,1104,169,583]" pageId="23" pageNumber="152">Pronotum about two times wider than long. Anterior and lateral margins distinctly bordered, base bordered except in the middle. Pronotum densely punctate anterolaterally, sparsely punctate on the base and anterior part medially.</paragraph>
<paragraph id="7108511E102EFFA5FF56DBFC9A0F6EF3" blockId="23.[144,1104,169,583]" pageId="23" pageNumber="152">Elytra trapezoidal, as wide as long, shiny. Stria relatively deep, with punctures 1.5–2 times larger than striae width. Elytral intervals flat, with minute punctuation.</paragraph>
<paragraph id="7108511E102EFFA5FF56DBBC9DC26D33" blockId="23.[144,1104,169,583]" pageId="23" pageNumber="152">Anterior tibiae with three outer teeth, without a small acute tooth between first outer tooth and apical spur. Anterior tibial spur rounded apically.</paragraph>
<paragraph id="7108511E102EFFA5FF56D87C9A166D53" blockId="23.[144,1104,169,583]" box="[186,1104,559,583]" pageId="23" pageNumber="152">
Pygidium with relatively thick borders. Disc granulate, without setae (
<figureCitation id="E98C4D9B102EFFA5FC22D87C9A066D53" box="[974,1088,559,583]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="23" pageNumber="152">Figure 9C</figureCitation>
).
</paragraph>
<paragraph id="7108511E102EFFA5FF7CD82B9F7B6D9B" blockId="23.[144,317,631,656]" box="[144,317,631,656]" pageId="23" pageNumber="152">
<heading id="2A40E672102EFFA5FF7CD82B9F7B6D9B" box="[144,317,631,656]" fontSize="10" level="3" pageId="23" pageNumber="152" reason="5">
<emphasis id="43C38D0C102EFFA5FF7CD82B9E8C6D84" box="[144,202,632,656]" italics="true" pageId="23" pageNumber="152">Male</emphasis>
unknown
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102EFFA5FF7CD88A9FD76C6C" pageId="23" pageNumber="152" type="materials_examined">
<paragraph id="7108511E102EFFA5FF7CD88A9F6A6DE5" blockId="23.[144,1104,729,888]" box="[144,300,729,753]" pageId="23" pageNumber="152">
<heading id="2A40E672102EFFA5FF7CD88A9F6A6DE5" box="[144,300,729,753]" fontSize="10" level="3" pageId="23" pageNumber="152" reason="5">
<emphasis id="43C38D0C102EFFA5FF7CD88A9F6A6DE5" box="[144,300,729,753]" italics="true" pageId="23" pageNumber="152">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102EFFA5FF7CD9539FD76C6C" blockId="23.[144,1104,729,888]" pageId="23" pageNumber="152">
<materialsCitation id="C1DF5B43102EFFA5FF7CD9539FD76C6C" ID-GBIF-Occurrence="3757906303" collectingDate="1927-08" collectionCode="BMNH" collectorName="H. G. Champion'" country="India" elevation="610" location="United Provs. Dehra Dun." municipality="United Provs. Dehra Dun." pageId="23" pageNumber="152" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="AE0CEFBC102EFFA5FF7CD9539EBB6C0C" box="[144,253,768,792]" pageId="23" pageNumber="152" type="holotype">Holotype</typeStatus>
,
<specimenCount id="67B19A97102EFFA5FEE4D9539F146C0C" box="[264,338,768,792]" pageId="23" pageNumber="152" type="female">female</specimenCount>
with the labels ‘
<collectingCountry id="09A0118E102EFFA5FDE1D9539C0A6C0C" box="[525,588,768,792]" name="India" pageId="23" pageNumber="152">India</collectingCountry>
.
<collectingMunicipality id="916CCB64102EFFA5FDBBD9539DC36C0C" box="[599,901,768,792]" pageId="23" pageNumber="152">United Provs. Dehra Dun.</collectingMunicipality>
<quantity id="B64FFCFB102EFFA5FC60D9539DAF6C0C" box="[908,1001,768,792]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="23" pageNumber="152" unit="ft" value="2000.0">
<elevation id="FA9AB62D102EFFA5FC60D9539DAF6C0C" box="[908,1001,768,792]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="23" pageNumber="152" unit="ft" value="2000.0">2,000 ft.</elevation>
</quantity>
<date id="050977DE102EFFA5FC1CD9539A096C0C" box="[1008,1103,768,792]" pageId="23" pageNumber="152" value="1927-08">
<collectingDate id="154D8E36102EFFA5FC1CD9539A096C0C" box="[1008,1103,768,792]" pageId="23" pageNumber="152" value="1927-08">viii.1927</collectingDate>
</date>
<collectorName id="DC4234C8102EFFA5FF7CD9739F0E6C2C" box="[144,328,800,824]" pageId="23" pageNumber="152">H.G.Champion’</collectorName>
, ‘India H.G.Champion B.M. 1931-8’, ‘
<taxonomicName id="B6B72A9D102EFFA5FCCBD9739A576C2C" box="[807,1041,800,824]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="striatus">Delopleurus striatus</taxonomicName>
type Arrow’, ‘Type’, ‘SYNTYPE’, ‘
<taxonomicName id="B6B72A9D102EFFA5FE17D9139D986C4C" authority="Arrow M. E. Bacchus" box="[507,990,832,856]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="striatus">Delopleurus striatus Arrow M.E.Bacchus</taxonomicName>
det 1975 SYNTYPE’ (
<collectionCode id="17A6C9DB102EFFA5FEC4D9339FC26C6C" box="[296,388,864,888]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="23" pageNumber="152">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102EFFA5FF7CD9929F966B34" pageId="23" pageNumber="152" type="distribution">
<paragraph id="7108511E102EFFA5FF7CD9929F526CCD" blockId="23.[144,1104,961,1056]" box="[144,276,961,985]" pageId="23" pageNumber="152">
<heading id="2A40E672102EFFA5FF7CD9929F526CCD" box="[144,276,961,985]" fontSize="10" level="3" pageId="23" pageNumber="152" reason="5">
<emphasis id="43C38D0C102EFFA5FF7CD9929F526CCD" box="[144,276,961,985]" italics="true" pageId="23" pageNumber="152">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E102EFFA5FF7CD9BB9F966B34" blockId="23.[144,1104,961,1056]" pageId="23" pageNumber="152">
The species is known from the single locality in northern periphery of the Indo-Gangetic Plain (
<figureCitation id="E98C4D9B102EFFA5FEA6DE5B9F866B34" box="[330,448,1032,1056]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="23" pageNumber="152">Figure 9D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

248
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@ -0,0 +1,248 @@
<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE0081034FFBCFE78DEE59FCE6AD8" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus pubescens Frolov 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="143" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="142" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
</mods:titleInfo>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
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<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>49</mods:number>
</mods:detail>
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<mods:number>3</mods:number>
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<mods:identifier type="DOI">10.1080/00222933.2014.909072</mods:identifier>
<mods:identifier type="GBIF-Dataset">3bad6762-1788-4f94-9cfd-1281b8619c55</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">4002597</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331157" ID-GBIF-Taxon="167368585" ID-Zenodo-Dep="4331157" LSID="urn:lsid:plazi:treatment:F91EE0081034FFBCFE78DEE59FCE6AD8" httpUri="http://treatment.plazi.org/id/F91EE0081034FFBCFE78DEE59FCE6AD8" lastPageId="14" lastPageNumber="143" pageId="13" pageNumber="142">
<subSubSection box="[404,844,1206,1231]" pageId="13" pageNumber="142" type="nomenclature">
<paragraph blockId="13.[404,844,1206,1263]" box="[404,844,1206,1231]" pageId="13" pageNumber="142">
<heading box="[404,844,1206,1231]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="142" reason="2">
<taxonomicName ID-CoL="34H5D" authority="Frolov" authorityName="Frolov" authorityYear="2014" box="[404,741,1206,1231]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="species" species="pubescens" status="sp. nov.">
<emphasis bold="true" box="[404,652,1206,1230]" italics="true" pageId="13" pageNumber="142">Delopleurus pubescens</emphasis>
Frolov
</taxonomicName>
,
<taxonomicNameLabel box="[754,844,1208,1231]" pageId="13" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[538,709,1239,1263]" pageId="13" pageNumber="142" type="description">
<paragraph blockId="13.[404,844,1206,1263]" box="[538,709,1239,1263]" pageId="13" pageNumber="142">
(
<figureCitation box="[547,700,1239,1263]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="13" pageNumber="142">Figure 2D–G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="142" type="diagnosis">
<paragraph blockId="13.[144,1104,1334,1494]" box="[144,252,1334,1357]" pageId="13" pageNumber="142">
<heading box="[144,252,1334,1357]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
<emphasis box="[144,252,1334,1357]" italics="true" pageId="13" pageNumber="142">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[144,1104,1334,1494]" pageId="13" pageNumber="142">
Males of this species can be separated from other
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[721,852,1373,1397]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis box="[721,852,1373,1397]" italics="true" pageId="13" pageNumber="142">Delopleurus</emphasis>
</taxonomicName>
species by the disc of pygidium being rugose, somewhat granulate and pubescent with sparse brown setae (
<figureCitation box="[152,265,1437,1461]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="13" pageNumber="142">Figure 2E</figureCitation>
). Such a sculpture is similar to that of females of
<taxonomicName box="[823,922,1437,1461]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[823,922,1437,1461]" italics="true" pageId="13" pageNumber="142">D. pullus</emphasis>
</taxonomicName>
(
<figureCitation box="[938,1051,1437,1461]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="13" pageNumber="142">Figure 3E</figureCitation>
) but it is not rugose in the latter.
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="143" pageId="13" pageNumber="142" type="description">
<paragraph blockId="13.[144,1103,1564,1691]" box="[144,271,1564,1587]" pageId="13" pageNumber="142">
<heading box="[144,271,1564,1587]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
<emphasis box="[144,271,1564,1587]" italics="true" pageId="13" pageNumber="142">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[144,1103,1564,1691]" box="[144,1100,1603,1628]" pageId="13" pageNumber="142">
<emphasis box="[144,323,1604,1628]" italics="true" pageId="13" pageNumber="142">
<typeStatus box="[144,249,1604,1628]" pageId="13" pageNumber="142">Holotype</typeStatus>
, male.
</emphasis>
Body strongly convex, black, glabrous, length 5.0 mm (
<figureCitation box="[966,1084,1603,1627]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="13" pageNumber="142">Figure 2D</figureCitation>
).
</paragraph>
<paragraph blockId="13.[144,1103,1564,1691]" lastBlockId="14.[212,1172,168,670]" lastPageId="14" lastPageNumber="143" pageId="13" pageNumber="142">Clypeus quadridentate, two medial teeth are acute, lateral ones are somewhat angulate. Genae right-angled, indistinctly separated from clypeus. Frontoclypeal and genal suturae almost indistinct. Clypeus rugose in anterior part and laterally, frons densely punctate (punctures separated by about one puncture diameter).</paragraph>
<paragraph blockId="14.[212,1172,168,670]" pageId="14" pageNumber="143">Pronotum more or less trapezoidal, about two times wider than long. All margins with distinct border. Disc and base punctate with punctures separated by two to three puncture diameters, sides and especially anterior angles more densely punctate.</paragraph>
<paragraph blockId="14.[212,1172,168,670]" pageId="14" pageNumber="143">Elytra trapezoidal, as wide as long, somewhat opaque. Stria deep, with punctures slightly larger than striae width. Elytral intervals slightly convex (more distinctly on disc).</paragraph>
<paragraph blockId="14.[212,1172,168,670]" pageId="14" pageNumber="143">Anterior tibiae with three outer teeth, without a small acute tooth between first outer tooth and apical spur.</paragraph>
<paragraph blockId="14.[212,1172,168,670]" pageId="14" pageNumber="143">
Pygidium with relatively slender borders and convex disc. Basal border slender and almost parallel-sided except in the middle. Apical border about twice as thick in the middle as the basal border, becoming more slender laterally. Disc with irregular, rugose and granulate sculpture, pubescent with short yellowish setae (
<figureCitation box="[1008,1123,583,607]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="14" pageNumber="143">Figure 2E</figureCitation>
).
</paragraph>
<paragraph blockId="14.[212,1172,168,670]" pageId="14" pageNumber="143">
Parameres with a small protuberance ventroapically (visible in lateral view,
<figureCitation box="[212,326,646,670]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="14" pageNumber="143">Figure 2F</figureCitation>
).
</paragraph>
<paragraph blockId="14.[212,414,720,744]" box="[212,414,720,744]" pageId="14" pageNumber="143">
<heading box="[212,414,720,744]" fontSize="10" level="3" pageId="14" pageNumber="143" reason="5">
<emphasis box="[212,292,720,744]" italics="true" pageId="14" pageNumber="143">Female</emphasis>
unknown.
</heading>
</paragraph>
<paragraph blockId="14.[212,627,793,817]" box="[212,627,793,817]" pageId="14" pageNumber="143">
<heading box="[212,627,793,817]" fontSize="10" level="3" pageId="14" pageNumber="143" reason="5">
<emphasis box="[212,330,794,817]" italics="true" pageId="14" pageNumber="143">
<typeStatus box="[212,327,794,817]" pageId="14" pageNumber="143">Paratypes</typeStatus>
.
</emphasis>
Body length
<quantity box="[486,623,793,817]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.5" metricValueMin="4.3" pageId="14" pageNumber="143" unit="mm" value="4.9" valueMax="5.5" valueMin="4.3">4.3–5.5 mm</quantity>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="143" type="distribution">
<paragraph blockId="14.[212,1171,891,986]" box="[212,344,891,915]" pageId="14" pageNumber="143">
<heading box="[212,344,891,915]" fontSize="10" level="3" pageId="14" pageNumber="143" reason="5">
<emphasis box="[212,344,891,915]" italics="true" pageId="14" pageNumber="143">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="14.[212,1171,891,986]" pageId="14" pageNumber="143">
The species is known from a few localities in the Congolese forest-savanna and Miombo woodlands (
<figureCitation box="[458,576,962,986]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="14" pageNumber="143">Figure 2G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[212,1154,1060,1123]" pageId="14" pageNumber="143" type="etymology">
<paragraph blockId="14.[212,1154,1060,1123]" box="[212,333,1060,1084]" pageId="14" pageNumber="143">
<heading box="[212,333,1060,1084]" fontSize="10" level="3" pageId="14" pageNumber="143" reason="5">
<emphasis box="[212,333,1060,1084]" italics="true" pageId="14" pageNumber="143">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="14.[212,1154,1060,1123]" box="[212,1154,1099,1123]" pageId="14" pageNumber="143">The species name is a Latin word denoting the peculiar sculpture of the pygidium.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="143" type="materials_examined">
<paragraph blockId="14.[212,1172,1197,1484]" box="[212,368,1197,1221]" pageId="14" pageNumber="143">
<heading box="[212,368,1197,1221]" fontSize="10" level="3" pageId="14" pageNumber="143" reason="5">
<emphasis box="[212,368,1197,1221]" italics="true" pageId="14" pageNumber="143">Type material</emphasis>
</heading>
</paragraph>
<paragraph blockId="14.[212,1172,1197,1484]" pageId="14" pageNumber="143">
<materialsCitation ID-GBIF-Occurrence="2849122344" collectingDate="1906-02" collectionCode="BMNH" collectorName="G. A. Marshall'" location="Salisbury Mashonaland" pageId="14" pageNumber="143" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[212,321,1237,1261]" pageId="14" pageNumber="143">Holotype</typeStatus>
,
<specimenCount box="[338,392,1237,1261]" pageId="14" pageNumber="143" type="male">male</specimenCount>
with the label ‘
<location LSID="urn:lsid:plazi:treatment:F91EE0081034FFBCFE78DEE59FCE6AD8:746807C51037FFBCFDA1DE869D186BF9" box="[589,862,1237,1261]" name="Salisbury Mashonaland" pageId="14" pageNumber="143">Salisbury Mashonaland</location>
<date box="[875,992,1237,1261]" pageId="14" pageNumber="143" value="1906-02">
<collectingDate box="[875,992,1237,1261]" pageId="14" pageNumber="143" value="1906-02">Feb. 1906</collectingDate>
</date>
<collectorName box="[1006,1171,1237,1261]" pageId="14" pageNumber="143">G.A.Marshall’</collectorName>
(
<collectionCode box="[220,312,1269,1293]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="14" pageNumber="143">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122355" box="[333,1081,1269,1293]" collectionCode="BMNH" pageId="14" pageNumber="143" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<typeStatus box="[333,447,1269,1293]" pageId="14" pageNumber="143">Paratypes</typeStatus>
:
<specimenCount box="[460,547,1269,1293]" pageId="14" pageNumber="143" type="male">3 males</specimenCount>
with the same label as the
<typeStatus box="[865,963,1269,1293]" pageId="14" pageNumber="143">holotype</typeStatus>
(
<collectionCode box="[981,1073,1269,1293]" pageId="14" pageNumber="143">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122375" collectionCode="SAMC" collectorName="Marshall" location="Salisbury Mashonaland" pageId="14" pageNumber="143" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[1095,1172,1269,1293]" pageId="14" pageNumber="143" type="male">1 male</specimenCount>
with the label ‘
<location LSID="urn:lsid:plazi:treatment:F91EE0081034FFBCFE78DEE59FCE6AD8:746807C51037FFBCFE6CDF479CCB6A38" box="[384,653,1300,1324]" name="Salisbury Mashonaland" pageId="14" pageNumber="143">Salisbury Mashonaland</location>
G.A.
<collectorName box="[712,821,1300,1324]" pageId="14" pageNumber="143">Marshall</collectorName>
189[4]’ (
<collectionCode box="[921,1005,1300,1324]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="14" pageNumber="143" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122349" collectionCode="MRAC" collectorName="R. P. Van Eyen'" country="Democratic Republic of the Congo" location="CONGO" pageId="14" pageNumber="143" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount box="[1027,1113,1300,1324]" pageId="14" pageNumber="143" type="male">2 males</specimenCount>
with the labels ‘COLL. MUS.
<collectingCountry box="[503,607,1332,1356]" name="Democratic Republic of the Congo" pageId="14" pageNumber="143">CONGO</collectingCountry>
Mayidi – 1945
<collectorName box="[789,968,1332,1357]" pageId="14" pageNumber="143">R. P.Van Eyen’</collectorName>
(
<collectionCode box="[985,1075,1332,1356]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="14" pageNumber="143" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122316" collectingDate="1933-11-17" collectionCode="MRAC" collectorName="G. F. Overlaet'" country="Democratic Republic of the Congo" county="Musee" location="Kapelekese" municipality="Lulua" pageId="14" pageNumber="143" specimenCount="1" specimenCount-male="1" stateProvince="Bas-Congo" typeStatus="paratype">
<specimenCount box="[1096,1172,1332,1356]" pageId="14" pageNumber="143" type="male">1 male</specimenCount>
with the label ‘
<collectingCounty box="[397,471,1365,1388]" pageId="14" pageNumber="143">Musee</collectingCounty>
du
<collectingRegion box="[527,602,1364,1388]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="14" pageNumber="143">Congo</collectingRegion>
<collectingMunicipality box="[616,687,1364,1388]" pageId="14" pageNumber="143">Lulua</collectingMunicipality>
: r.
<location LSID="urn:lsid:plazi:treatment:F91EE0081034FFBCFE78DEE59FCE6AD8:746807C51037FFBCFD33DF079D196A78" box="[735,863,1364,1388]" country="Democratic Republic of the Congo" county="Musee" municipality="Lulua" name="Kapelekese" pageId="14" pageNumber="143" stateProvince="Bas-Congo">Kapelekese</location>
<date box="[876,1000,1364,1388]" pageId="14" pageNumber="143" value="1933-11-17">
<collectingDate box="[876,1000,1364,1388]" pageId="14" pageNumber="143" value="1933-11-17">17.XI.1933</collectingDate>
</date>
<collectorName box="[1014,1171,1364,1388]" pageId="14" pageNumber="143">G.F.Overlaet’</collectorName>
(
<collectionCode box="[220,310,1396,1420]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="14" pageNumber="143" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122383" collectingDate="1900-12" collectionCode="IRSNB" collectorName="Coll. I. R. Sc. N. B. Rhodesie du Sud Salisbury &amp; Marshall Coll." country="Democratic Republic of the Congo" pageId="14" pageNumber="143" specimenCount="1" specimenCount-male="1" stateProvince="Bas-Congo" typeStatus="paratype">
<specimenCount box="[332,409,1396,1420]" pageId="14" pageNumber="143" type="male">1 male</specimenCount>
with the label ‘
<collectorName box="[593,1109,1396,1420]" pageId="14" pageNumber="143">Coll. I.R.Sc.N.B. Rhodesie du Sud Salisbury</collectorName>
<date pageId="14" pageNumber="143" value="1900-12">
<collectingDate pageId="14" pageNumber="143" value="1900-12">Dec. 1900</collectingDate>
</date>
G.A.K.
<collectorName box="[349,522,1428,1452]" pageId="14" pageNumber="143">Marshall Coll.</collectorName>
J.J.Gillet’ (
<collectionCode box="[652,740,1428,1452]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="14" pageNumber="143">IRSNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122360" collectionCode="DMAGD" country="Democratic Republic of the Congo" location="Salisbury" pageId="14" pageNumber="143" specimenCount="1" specimenCount-male="1" stateProvince="Bas-Congo" typeStatus="paratype">
<specimenCount box="[760,834,1428,1452]" pageId="14" pageNumber="143" type="male">1 male</specimenCount>
with the label ‘
<location LSID="urn:lsid:plazi:treatment:F91EE0081034FFBCFE78DEE59FCE6AD8:746807C51037FFBCFC02DFC79A106AB8" box="[1006,1110,1428,1452]" country="Democratic Republic of the Congo" name="Salisbury" pageId="14" pageNumber="143" stateProvince="Bas-Congo">Salisbury</location>
14/1/ 15’ (
<collectionCode box="[262,379,1460,1484]" pageId="14" pageNumber="143">DMAGD</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

197
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@ -0,0 +1,197 @@
<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE0081035FFBFFE63D9269C486B6B" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus naviauxi Frolov et Cambefort 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="142" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="141" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>49</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>129</mods:start>
<mods:end>154</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2014.909072</mods:identifier>
<mods:identifier type="GBIF-Dataset">3bad6762-1788-4f94-9cfd-1281b8619c55</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">4002597</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331161" ID-GBIF-Taxon="167368584" ID-Zenodo-Dep="4331161" LSID="urn:lsid:plazi:treatment:F91EE0081035FFBFFE63D9269C486B6B" httpUri="http://treatment.plazi.org/id/F91EE0081035FFBFFE63D9269C486B6B" lastPageId="13" lastPageNumber="142" pageId="12" pageNumber="141">
<subSubSection box="[399,984,885,909]" pageId="12" pageNumber="141" type="nomenclature">
<paragraph blockId="12.[399,984,885,941]" box="[399,984,885,909]" pageId="12" pageNumber="141">
<heading box="[399,984,885,909]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="141" reason="2">
<taxonomicName ID-CoL="34H5B" authority="Frolov et Cambefort" authorityName="Frolov et Cambefort" authorityYear="2014" box="[399,881,885,909]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
<emphasis bold="true" box="[399,633,885,909]" italics="true" pageId="12" pageNumber="141">Delopleurus naviauxi</emphasis>
Frolov et Cambefort
</taxonomicName>
,
<taxonomicNameLabel box="[894,984,886,909]" pageId="12" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[578,806,917,941]" pageId="12" pageNumber="141" type="description">
<paragraph blockId="12.[399,984,885,941]" box="[578,806,917,941]" pageId="12" pageNumber="141">
(
<figureCitation box="[586,715,917,941]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="12" pageNumber="141">Figures 1D</figureCitation>
,
<figureCitation box="[731,797,917,941]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="12" pageNumber="141">2A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="141" type="diagnosis">
<paragraph blockId="12.[212,1172,982,1077]" box="[212,320,982,1005]" pageId="12" pageNumber="141">
<heading box="[212,320,982,1005]" fontSize="10" level="3" pageId="12" pageNumber="141" reason="5">
<emphasis box="[212,320,982,1005]" italics="true" pageId="12" pageNumber="141">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[212,1172,982,1077]" pageId="12" pageNumber="141">
This species can be separated from other
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[665,797,1022,1046]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis box="[665,797,1022,1046]" italics="true" pageId="12" pageNumber="141">Delopleurus</emphasis>
</taxonomicName>
species by having deep transverse fossa on the pygidium and by the fore tibia with a small acute tooth near apical spur.
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="142" pageId="12" pageNumber="141" type="description">
<paragraph blockId="12.[212,1172,1150,1691]" box="[212,339,1150,1173]" pageId="12" pageNumber="141">
<heading box="[212,339,1150,1173]" fontSize="10" level="3" pageId="12" pageNumber="141" reason="5">
<emphasis box="[212,339,1150,1173]" italics="true" pageId="12" pageNumber="141">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[212,1172,1150,1691]" box="[212,1172,1189,1213]" pageId="12" pageNumber="141">
<emphasis box="[212,410,1189,1213]" italics="true" pageId="12" pageNumber="141">
<typeStatus box="[212,317,1189,1213]" pageId="12" pageNumber="141">Holotype</typeStatus>
, female.
</emphasis>
Body strongly convex, black, glabrous, length
<quantity box="[939,1024,1189,1213]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="12" pageNumber="141" unit="mm" value="4.9">4.9 mm</quantity>
(
<figureCitation box="[1040,1155,1189,1213]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="12" pageNumber="141">Figure 2A</figureCitation>
).
</paragraph>
<paragraph blockId="12.[212,1172,1150,1691]" pageId="12" pageNumber="141">Clypeus quadridentate. All four clypeal teeth acute, two medial ones relatively slender. Head without carinae on disc, small carinae present near inner margin of eyes. Genae right-angled, indistinctly separated from clypeus. Frontoclypeal and genal suturae indistinct. Eyes small, their dorsal parts slit-shaped, ventral parts subrectangular. Clypeus rugose in anterior part and laterally, frons densely punctate with elongate punctures.</paragraph>
<paragraph blockId="12.[212,1172,1150,1691]" pageId="12" pageNumber="141">Pronotum trapezoidal, about two times wider than long. Anterior and lateral margins bordered, base without border. Pronotum relatively densely punctate on disc (punctures separated by one to two puncture diameters), punctation becoming denser laterally.</paragraph>
<paragraph blockId="12.[212,1172,1150,1691]" pageId="12" pageNumber="141">Elytra trapezoidal, as wide as long, shiny. Stria distinct, with punctures larger than striae. Elytral intervals slightly convex on disc, with minute punctation.</paragraph>
<paragraph blockId="12.[212,1172,1150,1691]" pageId="12" pageNumber="141">
Anterior tibiae with three outer teeth and a small acute tooth between first outer tooth and apical spur (
<figureCitation box="[474,592,1635,1659]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="12" pageNumber="141">Figure 1D</figureCitation>
).
</paragraph>
<paragraph blockId="12.[212,1172,1150,1691]" lastBlockId="13.[144,1104,168,256]" lastPageId="13" lastPageNumber="142" pageId="12" pageNumber="141">
Pygidium with deep transversal slit-shaped fossa with yellowish setae (
<figureCitation box="[1043,1155,1667,1691]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="12" pageNumber="141">Figure 2B</figureCitation>
).
<emphasis box="[144,252,169,192]" italics="true" pageId="13" pageNumber="142">
<typeStatus box="[144,249,169,192]" pageId="13" pageNumber="142">Paratype</typeStatus>
.
</emphasis>
female, slightly larger than the
<typeStatus box="[649,747,168,192]" pageId="13" pageNumber="142">holotype</typeStatus>
(6.0 mm). The specimen has malformed or worn, reduced anterior tibiae without outer teeth and tarsi, and reduced clypeal teeth. It also differs from the
<typeStatus box="[650,748,232,256]" pageId="13" pageNumber="142">holotype</typeStatus>
in having less shiny dorsal side.
</paragraph>
<paragraph blockId="13.[144,324,303,327]" box="[144,324,303,327]" pageId="13" pageNumber="142">
<heading box="[144,324,303,327]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
<emphasis box="[144,202,303,327]" italics="true" pageId="13" pageNumber="142">Male</emphasis>
unknown.
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="142" type="materials_examined">
<paragraph blockId="13.[144,1104,397,557]" box="[144,300,397,421]" pageId="13" pageNumber="142">
<heading box="[144,300,397,421]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
<emphasis box="[144,300,397,421]" italics="true" pageId="13" pageNumber="142">Type material</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[144,1104,397,557]" pageId="13" pageNumber="142">
<materialsCitation ID-GBIF-Occurrence="3757906304" collectingDateMax="1937-05-31" collectingDateMin="1937-03-31" collectionCode="BMNH" collectorName="C. D. Knight'" country="Kenya" location="Colony Makuyu" pageId="13" pageNumber="142" specimenCount="1" typeStatus="holotype">
<typeStatus box="[144,253,437,461]" pageId="13" pageNumber="142">Holotype</typeStatus>
,
<specimenCount box="[268,342,437,461]" pageId="13" pageNumber="142" type="female">female</specimenCount>
with the labels ‘
<collectingCountry box="[545,620,438,462]" name="Kenya" pageId="13" pageNumber="142">Kenya</collectingCountry>
<location LSID="urn:lsid:plazi:treatment:F91EE0081035FFBFFE63D9269C486B6B:746807C51034FFBFFD9BDBE69D706ED9" box="[631,822,437,461]" country="Kenya" name="Colony Makuyu" pageId="13" pageNumber="142">Colony Makuyu</location>
iii-v 1937
<collectorName box="[954,1096,437,461]" pageId="13" pageNumber="142">C.D.Knight’</collectorName>
, ‘Imp. Inst. Entom.’ (
<collectionCode box="[384,473,469,493]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="13" pageNumber="142">BMNH</collectionCode>
) and
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3757906301" collectingDate="1978-04-30" collectionCode="MNHN" collectorName="R. Naviaux'" country="Kenya" location="Maji ya Chumvi" pageId="13" pageNumber="142" specimenCount="1" typeStatus="paratype">
<typeStatus box="[543,645,470,494]" pageId="13" pageNumber="142">paratype</typeStatus>
,
<specimenCount box="[659,733,469,493]" pageId="13" pageNumber="142" type="female">female</specimenCount>
with the labels ‘
<collectingCountry box="[928,1002,470,494]" name="Kenya" pageId="13" pageNumber="142">Kenya</collectingCountry>
<location LSID="urn:lsid:plazi:treatment:F91EE0081035FFBFFE63D9269C486B6B:746807C51034FFBFFC19DB869EAC6D19" country="Kenya" name="Maji ya Chumvi" pageId="13" pageNumber="142">Maji ya Chumvi</location>
<date box="[247,328,501,525]" pageId="13" pageNumber="142" value="78-04-30">
<collectingDate box="[247,328,501,525]" pageId="13" pageNumber="142" value="78-04-30">30.4.78</collectingDate>
</date>
R.N.’, ‘coll.
<collectorName box="[494,638,501,525]" pageId="13" pageNumber="142">R. Naviaux’</collectorName>
, ‘
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[666,803,501,525]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="152" phylum="Arthropoda" rank="genus">Delopleurus</taxonomicName>
n. sp? Y.Cambefort det’ (
<collectionCode box="[152,248,533,557]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="142">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="142" type="distribution">
<paragraph blockId="13.[144,1104,627,723]" box="[144,276,627,651]" pageId="13" pageNumber="142">
<heading box="[144,276,627,651]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
<emphasis box="[144,276,627,651]" italics="true" pageId="13" pageNumber="142">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[144,1104,627,723]" pageId="13" pageNumber="142">
The species is known from two localities in Northern Acacia-Commiphora bushlands and thickets ecoregion in East Africa (
<figureCitation box="[585,700,699,723]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="13" pageNumber="142">Figure 2C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[144,859,794,858]" pageId="13" pageNumber="142" type="etymology">
<paragraph blockId="13.[144,859,794,858]" box="[144,265,794,818]" pageId="13" pageNumber="142">
<heading box="[144,265,794,818]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
<emphasis box="[144,265,794,818]" italics="true" pageId="13" pageNumber="142">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[144,859,794,858]" box="[144,859,833,858]" pageId="13" pageNumber="142">The species is named after Roger Naviaux, a French collector.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="142" type="discussion">
<paragraph blockId="13.[144,1104,928,1151]" box="[144,242,928,952]" pageId="13" pageNumber="142">
<heading box="[144,242,928,952]" fontSize="10" level="3" pageId="13" pageNumber="142" reason="5">
<emphasis box="[144,242,928,952]" italics="true" pageId="13" pageNumber="142">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[144,1104,928,1151]" pageId="13" pageNumber="142">
Yves Cambefort (MNHN) examined the
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[608,739,968,992]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis box="[608,739,968,992]" italics="true" pageId="13" pageNumber="142">Delopleurus</emphasis>
</taxonomicName>
specimen from Maji ya Chumvi in the 1980s and noted that it was distinct from other described species of this genus. However, he did not describe it at that time. In the course of the revision of the genus, I examined the specimen from Makuyu which is housed in the BMNH. After comparison of the two specimens we concluded that they are conspecific and belong to a new species described above.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
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<heading box="[531,852,1539,1564]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="143" reason="2">
<taxonomicName ID-CoL="6CHFF" ID-ENA="794866" authority="Boheman" authorityName="Boheman" authorityYear="1857" box="[531,852,1539,1564]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="species" species="pullus">
<emphasis bold="true" box="[531,735,1539,1563]" italics="true" pageId="14" pageNumber="143">Delopleurus pullus</emphasis>
Boheman
</taxonomicName>
</heading>
</paragraph>
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<subSubSection box="[609,775,1571,1596]" pageId="14" pageNumber="143" type="description">
<paragraph blockId="14.[531,852,1539,1596]" box="[609,775,1571,1596]" pageId="14" pageNumber="143">
(
<figureCitation box="[618,766,1571,1596]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="14" pageNumber="143">Figure 3A–F</figureCitation>
)
</paragraph>
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<subSubSection pageId="14" pageNumber="143" type="reference_group">
<paragraph blockId="14.[212,1171,1635,1691]" pageId="14" pageNumber="143">
<treatmentCitationGroup pageId="14" pageNumber="143">
<treatmentCitation author="Boheman" box="[212,618,1635,1660]" pageId="14" pageNumber="143" year="1857">
<taxonomicName ID-CoL="6CHFF" authority="Boheman 1857" authorityName="Boheman" authorityYear="1857" box="[212,618,1635,1660]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="143" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[212,424,1636,1660]" italics="true" pageId="14" pageNumber="143">Delopleurus pullus</emphasis>
<bibRefCitation author="Boheman CH" box="[440,618,1635,1659]" pageId="14" pageNumber="143" refId="ref9783" refString="Boheman CH. 1857. Insecta Caffrariae annis 1838 - 1845 a J. A. Wahlberg collecta amici auxilio sussultus descripsit. Vol. 2. Coleoptera. Holmiae: Ex officina Norstedtiana." type="book" year="1857">Boheman 1857</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation author="Péringuey" box="[641,830,1635,1659]" pageId="14" pageNumber="143" year="1901">
<bibRefCitation author="Peringuey L" box="[641,823,1635,1659]" pageId="14" pageNumber="143" pagination="187 - 265" refId="ref10260" refString="Peringuey L. 1901. Tribe: COPRINI (Part 2). Trans South Afr Phil Soc. 12: 187 - 265. doi: 10.1080 / 21560382.1901.9525981" type="journal article" year="1901">Péringuey 1901</bibRefCitation>
;
</treatmentCitation>
<treatmentCitation author="Ferreira" box="[846,1014,1635,1659]" pageId="14" pageNumber="143" year="1972">
<bibRefCitation author="Ferreira M" box="[846,1007,1635,1659]" pageId="14" pageNumber="143" pagination="5 - 1088" refId="ref9860" refString="Ferreira M. 1972. Os escarabideos de Africa (Sul do Saara), I. Rev Ent Mocambique. 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972</bibRefCitation>
;
</treatmentCitation>
<treatmentCitation author="Gillet" box="[1031,1171,1635,1659]" pageId="14" pageNumber="143" year="1911">
<bibRefCitation author="Gillet JE" box="[1031,1164,1635,1659]" pageId="14" pageNumber="143" pagination="1 - 100" refId="ref10076" refString="Gillet JE. 1911. Scarabaeidae: Coprinae, I. In: Junk W, editor. Coleopterorum Catalogus auspicis et auxilio. Berlin: A. Schenkling; 38: 1 - 100." type="journal article" year="1911">Gillet 1911</bibRefCitation>
;
</treatmentCitation>
<treatmentCitation author="Janssens" box="[212,376,1667,1691]" pageId="14" pageNumber="143" year="1939">
<bibRefCitation author="Janssens A" box="[212,369,1667,1691]" pageId="14" pageNumber="143" pagination="1 - 104" refId="ref10112" refString="Janssens A. 1939. Coprini. Exploration du Parc National Albert, Mission GF de Witte. Institut des parcs nationaux du Congo Belge. 29: 1 - 104." type="journal article" year="1939">Janssens 1939</bibRefCitation>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="144" type="diagnosis">
<paragraph blockId="15.[144,1104,169,328]" box="[144,252,169,192]" pageId="15" pageNumber="144">
<heading box="[144,252,169,192]" fontSize="10" level="3" pageId="15" pageNumber="144" reason="5">
<emphasis box="[144,252,169,192]" italics="true" pageId="15" pageNumber="144">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="15.[144,1104,169,328]" pageId="15" pageNumber="144">
This species is most similar to
<taxonomicName box="[501,601,208,232]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="144" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis box="[501,601,208,232]" italics="true" pageId="15" pageNumber="144">D. gilleti</emphasis>
</taxonomicName>
but can be separated from it in having the pygidium with relatively larger disc punctate with smaller and sparser punctures in males and granulate and pubescent in females, and in having relatively sparsely punctate disc of pronotum.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="144" type="description">
<paragraph blockId="15.[144,1103,397,492]" box="[144,271,397,420]" pageId="15" pageNumber="144">
<heading box="[144,271,397,420]" fontSize="10" level="3" pageId="15" pageNumber="144" reason="5">
<emphasis box="[144,271,397,420]" italics="true" pageId="15" pageNumber="144">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="15.[144,1103,397,492]" pageId="15" pageNumber="144">
Beetles are small (
<quantity box="[349,486,436,460]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="6.1" metricValueMin="4.5" pageId="15" pageNumber="144" unit="mm" value="5.3" valueMax="6.1" valueMin="4.5">4.5–6.1 mm</quantity>
), strongly convex, black or dark brown, upper surface glabrous.
</paragraph>
<paragraph blockId="15.[144,1104,536,1071]" pageId="15" pageNumber="144">
<emphasis box="[144,202,537,561]" italics="true" pageId="15" pageNumber="144">Male</emphasis>
(
<figureCitation box="[219,336,536,560]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="15" pageNumber="144">Figure 3A</figureCitation>
). Clypeus quadridentate, two medial teeth acute, lateral ones rightangled to acute-angled. Clypeus rugose in anterior part and laterally, frons densely punctate (punctures separated by less than a puncture diameter).
</paragraph>
<paragraph blockId="15.[144,1104,536,1071]" pageId="15" pageNumber="144">Pronotum more or less trapezoidal, about two times wider than long. Anterior and lateral margins bordered, base not bordered. Surface relatively sparsely punctate on disc (punctures separated by two to three puncture diameters) and denser laterally.</paragraph>
<paragraph blockId="15.[144,1104,536,1071]" pageId="15" pageNumber="144">Elytra trapezoidal, as wide as long, shiny. Stria relatively deep, with punctures slightly larger than striae width. Elytral intervals feebly convex to almost flat, with minute punctation.</paragraph>
<paragraph blockId="15.[144,1104,536,1071]" pageId="15" pageNumber="144">Anterior tibiae with three outer teeth, without a small acute tooth between first outer tooth and apical spur.</paragraph>
<paragraph blockId="15.[144,1104,536,1071]" pageId="15" pageNumber="144">
Pygidium with relatively slender borders and convex disc. Basal border slender and almost parallel-sided except in the middle. Apical border about twice as thick in the middle as the basal border, becoming more slender laterally. Disc glabrous, punctate with punctures separated by one to two puncture diameters (
<figureCitation box="[945,1059,983,1007]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="15" pageNumber="144">Figure 3B</figureCitation>
).
</paragraph>
<paragraph blockId="15.[144,1104,536,1071]" pageId="15" pageNumber="144">
Parameres with a small protrusion ventroapically (visible in lateral view,
<figureCitation box="[144,259,1047,1071]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="15" pageNumber="144">Figure 3C</figureCitation>
).
</paragraph>
<paragraph blockId="15.[144,1103,1115,1203]" pageId="15" pageNumber="144">
<emphasis box="[144,224,1116,1140]" italics="true" pageId="15" pageNumber="144">Female</emphasis>
(
<figureCitation box="[244,365,1115,1139]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="15" pageNumber="144">Figure 3D</figureCitation>
) can be separated from male in having acute, not bifurcated spurs of anterior tibiae and disc of pygidium with small granules and setae (
<figureCitation box="[152,268,1179,1203]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="15" pageNumber="144">Figure 3E</figureCitation>
).
</paragraph>
</subSubSection>
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<paragraph blockId="15.[144,1104,1272,1399]" box="[144,300,1272,1296]" pageId="15" pageNumber="144">
<heading box="[144,300,1272,1296]" fontSize="10" level="3" pageId="15" pageNumber="144" reason="5">
<emphasis box="[144,300,1272,1296]" italics="true" pageId="15" pageNumber="144">Type material</emphasis>
</heading>
</paragraph>
<paragraph blockId="15.[144,1104,1272,1399]" pageId="15" pageNumber="144">
<materialsCitation ID-GBIF-Occurrence="2849122333" collectionCode="NHRS" pageId="15" pageNumber="144" specimenCount="1" typeStatus="lectotype">
<typeStatus box="[144,256,1312,1336]" pageId="15" pageNumber="144">Lectotype</typeStatus>
(here designated), male with the labels ‘
<collectorName box="[711,821,1312,1336]" pageId="15" pageNumber="144">Caffraria’</collectorName>
, ‘
<collectorName box="[844,949,1312,1336]" pageId="15" pageNumber="144">J. Wahlb</collectorName>
[erg].’, ‘
<collectorName box="[1032,1096,1312,1336]" pageId="15" pageNumber="144">Type’</collectorName>
, ‘
<taxonomicName authority="Boh." authorityName="Boheman" authorityYear="1857" box="[151,282,1343,1367]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="144" phylum="Arthropoda" rank="species" species="pullus">pullus Boh.</taxonomicName>
det
<collectorName box="[336,433,1344,1368]" pageId="15" pageNumber="144">J.Ferrer’</collectorName>
(
<collectionCode box="[451,534,1343,1367]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="15" pageNumber="144">NHRS</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122327" collectionCode="NHRS" collectorName="Caffraria' &amp; J. Wahlb" pageId="15" pageNumber="144" specimenCount="1" typeStatus="paralectotype">
<typeStatus box="[556,723,1343,1367]" pageId="15" pageNumber="144">Paralectotypes</typeStatus>
, male and female with the labels ‘
<collectorName box="[151,261,1375,1399]" pageId="15" pageNumber="144">Caffraria’</collectorName>
, ‘
<collectorName box="[285,391,1375,1399]" pageId="15" pageNumber="144">J. Wahlb</collectorName>
[erg].’ (
<collectionCode box="[469,552,1375,1399]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="15" pageNumber="144">NHRS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="15.[144,1104,1468,1691]" box="[144,359,1468,1492]" pageId="15" pageNumber="144">
<heading box="[144,359,1468,1492]" fontSize="10" level="3" pageId="15" pageNumber="144" reason="5">
<emphasis box="[144,359,1468,1492]" italics="true" pageId="15" pageNumber="144">Additional material</emphasis>
</heading>
</paragraph>
<paragraph blockId="15.[144,1104,1468,1691]" lastBlockId="16.[212,1172,168,671]" lastPageId="16" lastPageNumber="145" pageId="15" pageNumber="144">
<materialsCitation ID-GBIF-Occurrence="2849122305" box="[144,838,1508,1532]" collectingDate="1923-02" collectionCode="SAMC" country="Angola" location="Namakunde" pageId="15" pageNumber="144" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectingCountry box="[144,271,1508,1532]" name="Angola" pageId="15" pageNumber="144">ANGOLA</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51036FFBDFEF6DFB79FEF6AE8" box="[282,425,1508,1532]" country="Angola" name="Namakunde" pageId="15" pageNumber="144">Namakunde</location>
,
<date box="[437,520,1508,1532]" pageId="15" pageNumber="144" value="1923-02">
<collectingDate box="[437,520,1508,1532]" pageId="15" pageNumber="144" value="1923-02">II.1923</collectingDate>
</date>
,
<specimenCount box="[533,611,1508,1532]" pageId="15" pageNumber="144" type="male">1 male</specimenCount>
,
<specimenCount box="[623,729,1508,1532]" pageId="15" pageNumber="144" type="female">2 females</specimenCount>
(
<collectionCode box="[745,830,1508,1532]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="15" pageNumber="144" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122331" collectionCode="SAMC" collectorName="A. Bodong" country="Zimbabwe" location="Mutare" pageId="15" pageNumber="144" specimenCount="1" specimenCount-male="1">
<collectingCountry box="[850,1008,1508,1532]" name="Zimbabwe" pageId="15" pageNumber="144">ZIMBABWE</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51036FFBDFC17DFB79A096AEF" box="[1019,1103,1508,1531]" country="Zimbabwe" name="Mutare" pageId="15" pageNumber="144">Mutare</location>
[Umtali], 1903,
<collectorName box="[326,442,1540,1564]" pageId="15" pageNumber="144">A.Bodong</collectorName>
leg.,
<specimenCount box="[509,587,1540,1564]" pageId="15" pageNumber="144" type="male">1 male</specimenCount>
(
<collectionCode box="[605,690,1540,1564]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="15" pageNumber="144" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122330" collectionCode="SAMC" collectorName="Neil" country="Zimbabwe" location="Empandeni" pageId="15" pageNumber="144" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51036FFBDFD24DC579D0A6908" box="[712,844,1540,1564]" country="Zimbabwe" name="Empandeni" pageId="15" pageNumber="144">Empandeni</location>
, 1911, J.O’
<collectorName box="[976,1024,1540,1564]" pageId="15" pageNumber="144">Neil</collectorName>
leg.,
<specimenCount pageId="15" pageNumber="144" type="male">2 males</specimenCount>
,
<specimenCount box="[227,338,1571,1595]" pageId="15" pageNumber="144" type="female">2 females</specimenCount>
(
<collectionCode box="[358,443,1571,1595]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="15" pageNumber="144" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122323" box="[468,1099,1571,1595]" collectionCode="SAMC" collectorName="G. A. Marshall" country="Zimbabwe" location="Salisbury" pageId="15" pageNumber="144" specimenCount="3" specimenCount-male="3">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51036FFBDFE38DC709C79692F" box="[468,575,1571,1595]" country="Zimbabwe" name="Salisbury" pageId="15" pageNumber="144">Salisbury</location>
, 1894,
<collectorName box="[665,823,1571,1595]" pageId="15" pageNumber="144">G.A.Marshall</collectorName>
leg.,
<specimenCount box="[896,986,1571,1595]" pageId="15" pageNumber="144" type="male">3 males</specimenCount>
(
<collectionCode box="[1007,1091,1571,1595]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="15" pageNumber="144" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122368" box="[144,794,1603,1627]" collectingDate="1897-12" collectionCode="BMNH" collectorName="G. A. Marshall" country="Zimbabwe" location="Salisbury" pageId="15" pageNumber="144" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51036FFBDFF7CDC109EBD694F" box="[144,251,1603,1627]" country="Zimbabwe" name="Salisbury" pageId="15" pageNumber="144">Salisbury</location>
,
<date box="[265,368,1603,1627]" pageId="15" pageNumber="144" value="1897-12">
<collectingDate box="[265,368,1603,1627]" pageId="15" pageNumber="144" value="1897-12">XII.1897</collectingDate>
</date>
,
<collectorName box="[382,541,1603,1627]" pageId="15" pageNumber="144">G.A.Marshall</collectorName>
leg,
<specimenCount box="[600,676,1603,1627]" pageId="15" pageNumber="144" type="male">1 male</specimenCount>
(
<collectionCode box="[694,786,1603,1627]" pageId="15" pageNumber="144">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122319" collectingDate="1900-12" collectionCode="IRSNB" collectorName="G. A. K. Marshall" country="Zimbabwe" location="Salisbury" pageId="15" pageNumber="144" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51036FFBDFCC4DC109DD5694F" box="[808,915,1603,1627]" country="Zimbabwe" name="Salisbury" pageId="15" pageNumber="144">Salisbury</location>
,
<date box="[929,1032,1603,1627]" pageId="15" pageNumber="144" value="1900-12">
<collectingDate box="[929,1032,1603,1627]" pageId="15" pageNumber="144" value="1900-12">XII.1900</collectingDate>
</date>
,
<collectorName pageId="15" pageNumber="144">G.A. K.Marshall</collectorName>
leg.,
<specimenCount box="[344,423,1635,1659]" pageId="15" pageNumber="144" type="male">1 male</specimenCount>
(
<collectionCode box="[443,531,1635,1659]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="15" pageNumber="144">IRSNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122311" collectingDate="1972-01-05" collectionCode="SANC" collectorName="Bornemissza &amp; Kirk" country="Zimbabwe" location="Matimba" pageId="15" pageNumber="144" specimenCount="1" specimenCount-male="1">
<quantity box="[555,638,1635,1659]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="15" pageNumber="144" unit="mi" value="7.0">7 miles</quantity>
(
<quantity box="[658,754,1635,1659]" metricMagnitude="4" metricUnit="m" metricValue="1.13" pageId="15" pageNumber="144" unit="km" value="11.3">11.3 km</quantity>
) south of
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51036FFBDFC9EDC309D98696F" box="[882,990,1635,1659]" country="Zimbabwe" name="Matimba" pageId="15" pageNumber="144">Matimba</location>
,
<date box="[1005,1098,1635,1659]" pageId="15" pageNumber="144" value="1972-01-05">
<collectingDate box="[1005,1098,1635,1659]" pageId="15" pageNumber="144" value="1972-01-05">5.I.1972</collectingDate>
</date>
,
<collectorName box="[144,286,1667,1691]" pageId="15" pageNumber="144">Bornemissza</collectorName>
&amp;
<collectorName box="[320,373,1667,1691]" pageId="15" pageNumber="144">Kirk</collectorName>
leg.,
<specimenCount box="[433,507,1667,1691]" pageId="15" pageNumber="144" type="male">1 male</specimenCount>
(
<collectionCode box="[522,602,1667,1691]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="15" pageNumber="144" type="Museum">SANC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122350" collectingDate="1903-12" collectionCode="BMNH" collectorName="G. A. K. Marshall" country="Zimbabwe" lastPageId="16" lastPageNumber="145" location="Bulawayo" pageId="15" pageNumber="144" specimenCount="1" specimenCount-female="1" stateProvince="Bulawayo">
<collectingRegion box="[621,737,1667,1691]" country="Zimbabwe" name="Bulawayo" pageId="15" pageNumber="144">Bulawayo</collectingRegion>
,
<date box="[748,852,1667,1691]" pageId="15" pageNumber="144" value="1903-12">
<collectingDate box="[748,852,1667,1691]" pageId="15" pageNumber="144" value="1903-12">XII.1903</collectingDate>
</date>
,
<collectorName box="[863,1050,1667,1691]" pageId="15" pageNumber="144">G.A.K.Marshall</collectorName>
leg.,
<specimenCount box="[212,317,168,192]" pageId="16" pageNumber="145" type="female">1 female</specimenCount>
(
<collectionCode box="[343,436,168,192]" pageId="16" pageNumber="145">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122370" collectingDate="1951-12" collectionCode="TMSA" collectorName="G. Hobohm" country="Namibia" location="Abachaus" municipality="Otjiwarongo district" pageId="16" pageNumber="145" specimenCount="1" specimenCount-male="1">
<collectingCountry box="[465,596,168,192]" name="Namibia" pageId="16" pageNumber="145">NAMIBIA</collectingCountry>
:
<collectingMunicipality box="[617,858,168,192]" pageId="16" pageNumber="145">Otjiwarongo district</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FC9CDAFB9DA56FD4" box="[880,995,168,192]" country="Namibia" municipality="Otjiwarongo district" name="Abachaus" pageId="16" pageNumber="145">Abachaus</location>
,
<date box="[1017,1120,168,192]" pageId="16" pageNumber="145" value="1951-12">
<collectingDate box="[1017,1120,168,192]" pageId="16" pageNumber="145" value="1951-12">XII.1951</collectingDate>
</date>
,
<collectorName pageId="16" pageNumber="145">G. Hobohm</collectorName>
leg.,
<specimenCount box="[394,478,200,224]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
(
<collectionCode box="[503,586,200,224]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="16" pageNumber="145">TMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122326" collectingDate="1956-01-16" collectionCode="NHMB" collectorName="G. Hobohm" country="Namibia" location="Abachaus" municipality="Otjiwarongo district" pageId="16" pageNumber="145" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectingMunicipality box="[616,856,200,224]" pageId="16" pageNumber="145">Otjiwarongo district</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FC82DA9B9DA76FF4" box="[878,993,200,224]" country="Namibia" municipality="Otjiwarongo district" name="Abachaus" pageId="16" pageNumber="145">Abachaus</location>
,
<date box="[1014,1120,200,224]" pageId="16" pageNumber="145" value="1956-01-16">
<collectingDate box="[1014,1120,200,224]" pageId="16" pageNumber="145" value="1956-01-16">16.I.1956</collectingDate>
</date>
,
<collectorName pageId="16" pageNumber="145">G. Hobohm</collectorName>
leg.,
<specimenCount box="[374,452,232,256]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
,
<specimenCount box="[464,568,232,256]" pageId="16" pageNumber="145" type="female">2 females</specimenCount>
(
<collectionCode box="[584,676,232,256]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="145">NHMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122366" collectionCode="SAMC" country="Namibia" location="Kamanyab" pageId="16" pageNumber="145" specimenCount="5" specimenCount-female="4" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FD54DABB9D706E14" box="[696,822,232,256]" country="Namibia" name="Kamanyab" pageId="16" pageNumber="145">Kamanyab</location>
[Kamanjab],
<specimenCount box="[978,1055,232,256]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
,
<specimenCount box="[1067,1171,232,256]" pageId="16" pageNumber="145" type="female">4 females</specimenCount>
(
<collectionCode box="[220,305,264,288]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="16" pageNumber="145" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122324" box="[323,706,264,288]" collectingDate="1923-02" collectionCode="SAMC" country="Namibia" location="Mafa" pageId="16" pageNumber="145" specimenCount="2" specimenCount-female="2">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FEAFDB5B9FC56E34" box="[323,387,264,288]" country="Namibia" name="Mafa" pageId="16" pageNumber="145">Mafa</location>
,
<date box="[399,482,264,288]" pageId="16" pageNumber="145" value="1923-02">
<collectingDate box="[399,482,264,288]" pageId="16" pageNumber="145" value="1923-02">II.1923</collectingDate>
</date>
,
<specimenCount box="[494,598,264,288]" pageId="16" pageNumber="145" type="female">2 females</specimenCount>
(
<collectionCode box="[613,698,264,288]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="16" pageNumber="145" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122371" collectingDate="1922-01" collectionCode="SAMC" country="Namibia" location="Nagusib" pageId="16" pageNumber="145" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FD21DB5B9D6D6E34" box="[717,811,264,288]" country="Namibia" name="Nagusib" pageId="16" pageNumber="145">Nagusib</location>
[Farm 25 southeast Namutoni],
<date box="[212,285,296,320]" pageId="16" pageNumber="145" value="1922-01">
<collectingDate box="[212,285,296,320]" pageId="16" pageNumber="145" value="1922-01">I.1922</collectingDate>
</date>
, 1 spm. (
<collectionCode box="[402,486,296,320]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="16" pageNumber="145" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122334" collectingDate="1975-02-07" collectionCode="TMSA" collectorName="Endrody-Younga" country="Namibia" location="Ohopoho" municipality="Kaokoveld" pageId="16" pageNumber="145" specimenCount="2" specimenCount-female="2">
<collectingMunicipality box="[512,640,296,320]" pageId="16" pageNumber="145">Kaokoveld</collectingMunicipality>
,
<quantity box="[657,732,296,320]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="16" pageNumber="145" unit="km" value="44.0">44 km</quantity>
northwest
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FC89DB7B9D956E54" box="[869,979,296,320]" country="Namibia" municipality="Kaokoveld" name="Ohopoho" pageId="16" pageNumber="145">Ohopoho</location>
,
<date box="[996,1099,296,320]" pageId="16" pageNumber="145" value="1975-02-07">
<collectingDate box="[996,1099,296,320]" pageId="16" pageNumber="145" value="1975-02-07">7.II.1975</collectingDate>
</date>
, from under stones, leg.
<collectorName box="[440,638,328,352]" pageId="16" pageNumber="145">Endrody-Younga</collectorName>
leg.,
<specimenCount box="[716,829,328,352]" pageId="16" pageNumber="145" type="female">2 females</specimenCount>
(
<collectionCode box="[854,938,328,352]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="16" pageNumber="145">TMSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122343" collectingDate="1929-12" collectionCode="MNHN" collectorName="P. Lesne" country="Mozambique" location="Chimoio" pageId="16" pageNumber="145" specimenCount="2" specimenCount-female="2">
<collectingCountry box="[965,1168,328,352]" name="Mozambique" pageId="16" pageNumber="145">MOZAMBIQUE</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FF38DB349F7C6E6B" box="[212,314,359,383]" country="Mozambique" name="Chimoio" pageId="16" pageNumber="145">Chimoio</location>
,
<date box="[327,431,359,383]" pageId="16" pageNumber="145" value="1929-12">
<collectingDate box="[327,431,359,383]" pageId="16" pageNumber="145" value="1929-12">XII.1929</collectingDate>
</date>
,
<collectorName box="[445,533,360,383]" pageId="16" pageNumber="145">P.Lesne</collectorName>
leg.,
<specimenCount box="[599,706,359,383]" pageId="16" pageNumber="145" type="female">2 females</specimenCount>
(
<collectionCode box="[724,820,359,383]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="145">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2849122373" collectingDate="1987-12-18" collectingDateMax="1987-12-20" collectingDateMin="1987-12-18" collectionCode="SANC" collectorName="Grobbelaar" country="South Africa" location="Ellisras District" municipality="Nyala Nature Reserve" pageId="16" pageNumber="145" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry box="[842,1060,359,383]" name="South Africa" pageId="16" pageNumber="145">SOUTH AFRICA</collectingCountry>
: D’
<collectingMunicipality pageId="16" pageNumber="145">Nyala Nature Reserve</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FE79DBD49C0E6E8B" box="[405,584,391,415]" country="South Africa" municipality="Nyala Nature Reserve" name="Ellisras District" pageId="16" pageNumber="145">Ellisras District</location>
,
<date box="[598,775,391,415]" pageId="16" pageNumber="145" value="1987-12-18" valueMax="1987-12-20" valueMin="1987-12-18">
<collectingDate box="[598,775,391,415]" pageId="16" pageNumber="145" value="1987-12-18" valueMax="1987-12-20" valueMin="1987-12-18">18–20.XII.1987</collectingDate>
</date>
,
<collectorName box="[788,918,391,415]" pageId="16" pageNumber="145">Grobbelaar</collectorName>
leg.,
<specimenCount box="[983,1062,391,415]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
,
<specimenCount box="[1075,1171,391,415]" pageId="16" pageNumber="145" type="female">1 female</specimenCount>
(
<collectionCode box="[220,300,423,447]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="16" pageNumber="145" type="Museum">SANC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122309" box="[325,1053,423,447]" collectionCode="TMSA" collectorName="P. A. Krantz" country="South Africa" location="Lydenburg District" pageId="16" pageNumber="145" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FEA9DBF49C636EAB" box="[325,549,423,447]" country="South Africa" name="Lydenburg District" pageId="16" pageNumber="145">Lydenburg District</location>
, 1896,
<collectorName box="[639,770,423,447]" pageId="16" pageNumber="145">P.A.Krantz</collectorName>
leg.,
<specimenCount box="[841,941,423,447]" pageId="16" pageNumber="145" type="female">1 female</specimenCount>
(
<collectionCode box="[962,1045,423,447]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="16" pageNumber="145">TMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122325" collectionCode="SAMC" collectorName="J. Brown" country="South Africa" location="Vryburg" pageId="16" pageNumber="145" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FBC2DBF49AC96EAB" box="[1070,1167,423,447]" country="South Africa" name="Vryburg" pageId="16" pageNumber="145">Vryburg</location>
, 1918,
<collectorName box="[281,375,456,479]" pageId="16" pageNumber="145">J.Brown</collectorName>
leg.,
<specimenCount box="[439,515,455,479]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
(
<collectionCode box="[531,615,455,479]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="16" pageNumber="145" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122384" collectionCode="MNHN" collectorName="E. Simoni" country="South Africa" location="Vryburg" pageId="16" pageNumber="145" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FD91DB949C996ECB" box="[637,735,455,479]" country="South Africa" name="Vryburg" pageId="16" pageNumber="145">Vryburg</location>
, 1893,
<collectorName box="[816,921,455,479]" pageId="16" pageNumber="145">E.Simoni</collectorName>
leg.,
<specimenCount box="[984,1063,455,479]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
,
<specimenCount box="[1076,1171,455,479]" pageId="16" pageNumber="145" type="female">1 female</specimenCount>
(
<collectionCode box="[220,316,487,511]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="145">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122320" box="[337,868,487,511]" collectionCode="TMSA" collectorName="G. van Son" country="South Africa" location="Malta" pageId="16" pageNumber="145" specimenCount="2" specimenCount-male="2">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FEBDDBB49FDF6EEB" box="[337,409,487,511]" country="South Africa" name="Malta" pageId="16" pageNumber="145">Malta</location>
, 1928,
<collectorName box="[490,611,487,511]" pageId="16" pageNumber="145">G.van Son</collectorName>
leg.,
<specimenCount box="[675,761,487,511]" pageId="16" pageNumber="145" type="male">2 males</specimenCount>
(
<collectionCode box="[777,860,487,511]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="16" pageNumber="145">TMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122328" collectionCode="SAMC" country="South Africa" location="Shiluvane" pageId="16" pageNumber="145" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FC9DDBB49DA46EEB" box="[881,994,487,511]" country="South Africa" name="Shiluvane" pageId="16" pageNumber="145">Shiluvane</location>
, 1902,
<specimenCount box="[1076,1171,487,511]" pageId="16" pageNumber="145" type="female">1 female</specimenCount>
(
<collectionCode box="[220,305,519,543]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="16" pageNumber="145" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122303" box="[323,941,519,543]" collectionCode="MNHN" collectorName="H. A. Janod" country="South Africa" location="Shiluvane" pageId="16" pageNumber="145" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FEAFD8549FF26D0B" box="[323,436,519,543]" country="South Africa" name="Shiluvane" pageId="16" pageNumber="145">Shiluvane</location>
,
<collectorName box="[448,579,519,543]" pageId="16" pageNumber="145">H.A. Janod</collectorName>
leg.,
<specimenCount box="[640,738,519,543]" pageId="16" pageNumber="145" type="female">1 female</specimenCount>
, 2 spm. (
<collectionCode box="[837,933,519,543]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="145">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122337" collectingDate="1980-11-24" collectingDateMax="1980-11-28" collectingDateMin="1980-11-24" collectionCode="SANC" collectorName="S. J. van Tonder" country="South Africa" location="Thabazimbi" municipality="Ben Alberts Nature Reserve" pageId="16" pageNumber="145" specimenCount="1" specimenCount-male="1">
<collectingMunicipality pageId="16" pageNumber="145">Ben Alberts Nature Reserve</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FEABD8749F976D2B" box="[327,465,551,575]" country="South Africa" municipality="Ben Alberts Nature Reserve" name="Thabazimbi" pageId="16" pageNumber="145">Thabazimbi</location>
,
<date box="[488,656,551,575]" pageId="16" pageNumber="145" value="1980-11-24" valueMax="1980-11-28" valueMin="1980-11-24">
<collectingDate box="[488,656,551,575]" pageId="16" pageNumber="145" value="1980-11-24" valueMax="1980-11-28" valueMin="1980-11-24">24–28.XI.1980</collectingDate>
</date>
,
<collectorName box="[679,882,551,575]" pageId="16" pageNumber="145">S.J. van Tonder</collectorName>
leg.,
<specimenCount box="[966,1052,551,575]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
(
<collectionCode box="[1079,1159,551,575]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="16" pageNumber="145" type="Museum">SANC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122381" box="[212,964,583,607]" collectingDate="1914-12" collectionCode="IRSNB" collectorName="C. J. Swierstra" country="South Africa" location="Moorddrift" pageId="16" pageNumber="145" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FF38D8149F116D4B" box="[212,343,583,607]" country="South Africa" name="Moorddrift" pageId="16" pageNumber="145">Moorddrift</location>
,
<date box="[371,474,583,607]" pageId="16" pageNumber="145" value="1914-12">
<collectingDate box="[371,474,583,607]" pageId="16" pageNumber="145" value="1914-12">XII.1914</collectingDate>
</date>
,
<collectorName box="[502,652,583,607]" pageId="16" pageNumber="145">C.J.Swierstra</collectorName>
leg.,
<specimenCount box="[745,836,583,607]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
(
<collectionCode box="[868,956,583,607]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="16" pageNumber="145">IRSNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122308" collectionCode="MNHN" country="South Africa" location="Boheman" municipality="Durban" pageId="16" pageNumber="145" specimenCount="1" specimenCount-female="1">
<collectingMunicipality box="[992,1083,583,607]" pageId="16" pageNumber="145">Durban</collectingMunicipality>
, 1846,
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FF38D8349F056D6B" box="[212,323,615,639]" country="South Africa" municipality="Durban" name="Boheman" pageId="16" pageNumber="145">Boheman</location>
,
<specimenCount box="[336,432,615,639]" pageId="16" pageNumber="145" type="female">1 female</specimenCount>
(
<collectionCode box="[449,545,615,639]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="145">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2849122378" collectingDate="2011-01-11" collectionCode="UPSA" collectorName="C. Deschodt" country="South Africa" location="Legonyane" pageId="16" pageNumber="145" specimenCount="5" specimenCount-female="4" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FDDBD8349CF06D6B" box="[567,694,615,639]" country="South Africa" name="Legonyane" pageId="16" pageNumber="145">Legonyane</location>
,
<date box="[707,814,615,639]" pageId="16" pageNumber="145" value="2011-01-11">
<collectingDate box="[707,814,615,639]" pageId="16" pageNumber="145" value="2011-01-11">11.I.2011</collectingDate>
</date>
, from mushroom,
<collectorName box="[1029,1172,615,639]" pageId="16" pageNumber="145">C. Deschodt</collectorName>
leg.,
<specimenCount box="[267,346,646,670]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
,
<specimenCount box="[359,465,646,670]" pageId="16" pageNumber="145" type="female">4 females</specimenCount>
(
<collectionCode box="[482,559,646,670]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="16" pageNumber="145">UPSA</collectionCode>
)
</materialsCitation>
; ‘
<materialsCitation ID-GBIF-Occurrence="2849122340" box="[587,872,646,671]" collectionCode="MNHN" country="South Africa" location="Natal'" pageId="16" pageNumber="145" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:F91EE0081037FFA2FDFFDC509AD26C31:746807C51029FFA2FDA7D8D59CD56D8B" box="[587,659,646,671]" country="South Africa" name="Natal'" pageId="16" pageNumber="145">Natal’</location>
,
<specimenCount box="[675,751,646,670]" pageId="16" pageNumber="145" type="male">1 male</specimenCount>
(
<collectionCode box="[768,864,646,670]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="145">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[212,1172,742,805]" pageId="16" pageNumber="145" type="distribution">
<paragraph blockId="16.[212,1172,742,805]" box="[212,344,742,766]" pageId="16" pageNumber="145">
<heading box="[212,344,742,766]" fontSize="10" level="3" pageId="16" pageNumber="145" reason="5">
<emphasis box="[212,344,742,766]" italics="true" pageId="16" pageNumber="145">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="16.[212,1172,742,805]" box="[212,1172,781,805]" pageId="16" pageNumber="145">
The species is widely distributed in the northern part of Southern Africa (
<figureCitation box="[1043,1156,781,805]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="16" pageNumber="145">Figure 3F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

813
data/F9/1E/E0/F91EE008103BFFBEFDF2DAFB9AD56C0A.xml Normal file
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@ -0,0 +1,813 @@
<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE008103BFFBEFDF2DAFB9AD56C0A" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus Erichson 1847" docType="treatment" docVersion="8" lastPageNumber="141" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="131" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
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<mods:namePart>Frolov, Andrey V.</mods:namePart>
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<mods:date>2014</mods:date>
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<mods:number>2014-06-30</mods:number>
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<mods:number>49</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331143" ID-GBIF-Taxon="167368579" ID-Zenodo-Dep="4331143" LSID="urn:lsid:plazi:treatment:F91EE008103BFFBEFDF2DAFB9AD56C0A" httpUri="http://treatment.plazi.org/id/F91EE008103BFFBEFDF2DAFB9AD56C0A" lastPageId="12" lastPageNumber="141" pageId="2" pageNumber="131">
<subSubSection box="[542,841,168,192]" pageId="2" pageNumber="131" type="nomenclature">
<paragraph blockId="2.[542,841,168,192]" box="[542,841,168,192]" pageId="2" pageNumber="131">
<heading box="[542,841,168,192]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="131" reason="2">
<taxonomicName ID-CoL="42PQ" ID-ENA="518467" authority="Erichson 1847" authorityName="Erichson" authorityYear="1847" box="[542,841,168,192]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[542,778,168,192]" italics="true" pageId="2" pageNumber="131">
<emphasis bold="true" box="[542,673,168,192]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
<bibRefCitation author="Erichson WF" box="[682,778,168,192]" pageId="2" pageNumber="131" refId="ref9841" refString="Erichson WF. 1847. Naturgeschichte des Insecten Deutschland. I. Coleoptera. Berlin: Nicolaischen Buchhandlung." type="book" year="1847">Erichson</bibRefCitation>
</emphasis>
1847
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="131" type="type_taxon">
<paragraph blockId="2.[212,1172,232,607]" box="[212,658,232,257]" pageId="2" pageNumber="131">
<heading box="[212,658,232,257]" fontSize="10" level="3" pageId="2" pageNumber="131" reason="5">
<typeStatus box="[212,269,233,257]" pageId="2" pageNumber="131">Type</typeStatus>
species: D.
<taxonomicName authority="Boheman 1857" authorityName="Boheman" authorityYear="1857" box="[408,651,232,256]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[408,472,232,256]" italics="true" pageId="2" pageNumber="131">pullus</emphasis>
<bibRefCitation author="Boheman CH" box="[481,651,232,256]" pageId="2" pageNumber="131" refId="ref9783" refString="Boheman CH. 1857. Insecta Caffrariae annis 1838 - 1845 a J. A. Wahlberg collecta amici auxilio sussultus descripsit. Vol. 2. Coleoptera. Holmiae: Ex officina Norstedtiana." type="book" year="1857">Boheman 1857</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
<paragraph blockId="2.[212,1172,232,607]" pageId="2" pageNumber="131">
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,343,264,288]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[212,343,264,288]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
</taxonomicName>
was established by
<bibRefCitation author="Erichson WF" box="[570,749,264,288]" pageId="2" pageNumber="131" refId="ref9841" refString="Erichson WF. 1847. Naturgeschichte des Insecten Deutschland. I. Coleoptera. Berlin: Nicolaischen Buchhandlung." type="book" year="1847">Erichson (1847)</bibRefCitation>
for ‘one small south African species, which differs from
<taxonomicName authorityName="Hope" authorityYear="1833" box="[424,549,296,320]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[424,549,296,320]" italics="true" pageId="2" pageNumber="131">Coptorhina</emphasis>
</taxonomicName>
in narrower hind legs’. Ten years later
<bibRefCitation author="Boheman CH" box="[987,1172,296,320]" pageId="2" pageNumber="131" refId="ref9783" refString="Boheman CH. 1857. Insecta Caffrariae annis 1838 - 1845 a J. A. Wahlberg collecta amici auxilio sussultus descripsit. Vol. 2. Coleoptera. Holmiae: Ex officina Norstedtiana." type="book" year="1857">Boheman (1857)</bibRefCitation>
gave the name to this species,
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[565,666,328,352]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[565,666,328,352]" italics="true" pageId="2" pageNumber="131">D. pullus</emphasis>
</taxonomicName>
, and so fixed the
<typeStatus box="[879,927,328,352]" pageId="2" pageNumber="131">type</typeStatus>
species of the genus.
<bibRefCitation author="Sharp D" box="[212,358,359,383]" pageId="2" pageNumber="131" pagination="33 - 54" refId="ref10290" refString="Sharp D. 1876. Descriptions of some new genera and species of Scarabaeidae from tropical Asia and Malasia. Col Hefte. 13: 33 - 54." type="journal article" year="1876">Sharp (1876)</bibRefCitation>
described
<taxonomicName authorityName="Sharp" authorityYear="1876" box="[481,676,360,384]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
<emphasis box="[481,676,360,384]" italics="true" pageId="2" pageNumber="131">Coptorhina parva</emphasis>
</taxonomicName>
from northern
<collectingCountry box="[857,920,359,383]" name="India" pageId="2" pageNumber="131">India</collectingCountry>
.
<bibRefCitation author="Arrow GJ" box="[933,1087,359,383]" pageId="2" pageNumber="131" refId="ref9736" refString="Arrow GJ. 1931. The Fauna of British India III, Coprinae. London: Taylor &amp; Francis." type="book" year="1931">Arrow (1931)</bibRefCitation>
moved
<taxonomicName authorityName="Sharp" authorityYear="1876" box="[212,300,392,416]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
<emphasis box="[212,300,392,416]" italics="true" pageId="2" pageNumber="131">C.parva</emphasis>
</taxonomicName>
to
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[352,483,392,416]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[352,483,392,416]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
</taxonomicName>
and described the second species from
<collectingCountry box="[975,1037,391,415]" name="India" pageId="2" pageNumber="131">India</collectingCountry>
,
<taxonomicName box="[1056,1164,392,415]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="striatus">
<emphasis box="[1056,1164,392,415]" italics="true" pageId="2" pageNumber="131">D.striatus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Paulian R" box="[212,380,423,447]" pageId="2" pageNumber="131" pagination="162 - 164" refId="ref10236" refString="Paulian R. 1934. Quelques Panelini asiatiques nouveaux ou peu connus. Bull Soc Ent Fr. 39: 162 - 164." type="journal article" year="1934">Paulian (1934)</bibRefCitation>
described the third species from
<collectingCountry box="[775,837,423,447]" name="India" pageId="2" pageNumber="131">India</collectingCountry>
,
<taxonomicName authorityName="Paulian" authorityYear="1934" box="[854,975,424,448]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis box="[854,975,424,448]" italics="true" pageId="2" pageNumber="131">D. cardoni</emphasis>
</taxonomicName>
.
<bibRefCitation author="Janssens A" box="[994,1172,423,447]" pageId="2" pageNumber="131" pagination="1 - 104" refId="ref10112" refString="Janssens A. 1939. Coprini. Exploration du Parc National Albert, Mission GF de Witte. Institut des parcs nationaux du Congo Belge. 29: 1 - 104." type="journal article" year="1939">Janssens (1939)</bibRefCitation>
described the second species from Africa,
<taxonomicName box="[691,790,455,479]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis box="[691,790,455,479]" italics="true" pageId="2" pageNumber="131">D. gilleti</emphasis>
</taxonomicName>
, and gave a diagnostic key to all species known by then.
<bibRefCitation author="Frey G" box="[482,614,487,512]" pageId="2" pageNumber="131" pagination="538 - 549" refId="ref9887" refString="Frey G. 1963. Neue Onthophagen (Col. Scarab.) aus Australien (Queenslandausbeute von H. Demarz) und aus Afrika. Ent Arb Mus Frey. 14: 538 - 549." type="journal article" year="1963">Frey (1963)</bibRefCitation>
described
<taxonomicName authorityName="Frey" authorityYear="1963" box="[736,868,488,511]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis box="[736,868,488,511]" italics="true" pageId="2" pageNumber="131">D. janssensi</emphasis>
</taxonomicName>
from
<collectingCountry box="[940,1041,487,511]" name="Ethiopia" pageId="2" pageNumber="131">Ethiopia</collectingCountry>
. Except for this primary taxonomic literature, members of the genus were listed in the catalogues and monographs (
<bibRefCitation author="Peringuey L" box="[421,597,551,575]" pageId="2" pageNumber="131" pagination="187 - 265" refId="ref10260" refString="Peringuey L. 1901. Tribe: COPRINI (Part 2). Trans South Afr Phil Soc. 12: 187 - 265. doi: 10.1080 / 21560382.1901.9525981" type="journal article" year="1901">Péringuey 1901</bibRefCitation>
;
<bibRefCitation author="Gillet JE" box="[615,742,551,575]" pageId="2" pageNumber="131" pagination="1 - 100" refId="ref10076" refString="Gillet JE. 1911. Scarabaeidae: Coprinae, I. In: Junk W, editor. Coleopterorum Catalogus auspicis et auxilio. Berlin: A. Schenkling; 38: 1 - 100." type="journal article" year="1911">Gillet 1911</bibRefCitation>
;
<bibRefCitation author="Balthasar V" box="[759,930,551,575]" pageId="2" pageNumber="131" refId="ref9756" refString="Balthasar V. 1963. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region; Coleoptera: Lamellicornia. Prague: Tschechoslowakische Akademie der Wissenschaften." type="book" year="1963">Balthasar 1963</bibRefCitation>
;
<bibRefCitation author="Ferreira M" box="[947,1102,551,575]" pageId="2" pageNumber="131" pagination="5 - 1088" refId="ref9860" refString="Ferreira M. 1972. Os escarabideos de Africa (Sul do Saara), I. Rev Ent Mocambique. 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972</bibRefCitation>
;
<bibRefCitation author="Kral D &amp; Lobl I" pageId="2" pageNumber="131" refId="ref10144" refString="Kral D, Lobl I. 2006. Ateuchini. In: Lobl I, Smetana A, editor, Catalogue of Palaearctic Coleoptera, V. 3. Stensrup: Apollo Books." type="journal volume" year="2006">Král and Löbl 2006</bibRefCitation>
;
<bibRefCitation author="Davis ALV &amp; Frolov AV &amp; Scholtz CH" box="[394,587,583,607]" pageId="2" pageNumber="131" refId="ref9818" refString="Davis ALV, Frolov AV, Scholtz CH. 2008. The African dung beetle genera. Pretoria: Protea Book House." type="book" year="2008">Davis et al. 2008</bibRefCitation>
) but few additional data were given.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="131" type="diagnosis">
<paragraph blockId="2.[212,1171,682,872]" box="[212,320,682,705]" pageId="2" pageNumber="131">
<heading box="[212,320,682,705]" fontSize="10" level="3" pageId="2" pageNumber="131" reason="5">
<emphasis box="[212,320,682,705]" italics="true" pageId="2" pageNumber="131">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="2.[212,1171,682,872]" pageId="2" pageNumber="131">
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,343,721,745]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[212,343,721,745]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
</taxonomicName>
and closely related
<taxonomicName authorityName="Hope" authorityYear="1833" box="[580,705,721,745]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[580,705,721,745]" italics="true" pageId="2" pageNumber="131">Coptorhina</emphasis>
</taxonomicName>
share somewhat rectangular metepisternum, widest in its hind part, very convex epipleural margin (
<figureCitation box="[917,1035,752,776]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="2" pageNumber="131">Figure 1A</figureCitation>
), and other taxonomic characters of the
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[540,666,785,809]" class="Insecta" family="Scarabaeidae" genus="Sarophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[540,666,785,809]" italics="true" pageId="2" pageNumber="131">Sarophorus</emphasis>
</taxonomicName>
group of genera (
<bibRefCitation author="Frolov AV &amp; Akhmetova LA &amp; Scholtz CH" box="[875,1082,784,808]" pageId="2" pageNumber="131" pagination="1477 - 1508" refId="ref9962" refString="Frolov AV, Akhmetova LA, Scholtz CH. 2008. Revision of the obligate mushroom- feeding African &quot; dung beetle &quot; genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae). J Nat Hist. 42: 1477 - 1508." type="journal article" year="2008">Frolov et al. 2008</bibRefCitation>
). From
<taxonomicName authorityName="Hope" authorityYear="1833" box="[212,337,817,841]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[212,337,817,841]" italics="true" pageId="2" pageNumber="131">Coptorhina</emphasis>
</taxonomicName>
it can easily be separated by quadridentate clypeus and smaller body size (
<quantity box="[219,354,848,872]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="6.1" metricValueMin="3.5" pageId="2" pageNumber="131" unit="mm" value="4.8" valueMax="6.1" valueMin="3.5">3.5–6.1 mm</quantity>
in
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[393,525,848,872]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[393,525,848,872]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
</taxonomicName>
and 8.0–20.0 mm in
<taxonomicName authorityName="Hope" authorityYear="1833" box="[770,895,848,872]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[770,895,848,872]" italics="true" pageId="2" pageNumber="131">Coptorhina</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="135" pageId="2" pageNumber="131" type="description">
<paragraph blockId="2.[212,1172,947,1137]" box="[212,339,947,970]" pageId="2" pageNumber="131">
<heading box="[212,339,947,970]" fontSize="10" level="3" pageId="2" pageNumber="131" reason="5">
<emphasis box="[212,339,947,970]" italics="true" pageId="2" pageNumber="131">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="2.[212,1172,947,1137]" box="[212,1129,986,1010]" pageId="2" pageNumber="131">
Beetles are small (
<quantity box="[419,556,986,1010]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="6.1" metricValueMin="3.5" pageId="2" pageNumber="131" unit="mm" value="4.8" valueMax="6.1" valueMin="3.5">3.5–6.1 mm</quantity>
), strongly convex, black or dark brown, glabrous.
</paragraph>
<paragraph blockId="2.[212,1172,947,1137]" lastBlockId="3.[144,1104,1313,1688]" lastPageId="3" lastPageNumber="132" pageId="2" pageNumber="131">
Clypeus distinctly quadridentate in most species. Two medial teeth are almost always acute, lateral ones are acute to right-angled. In some species, lateral clypeal teeth are almost not separated from the lateral margin (
<figureCitation box="[840,993,1082,1106]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="2" pageNumber="131">Figure 7A, D</figureCitation>
). Head without carinae on disc, and lacking horns or tubercles. Small carina present near inner margin of eye. Genae right-angled, indistinctly separated from clypeus. Frontoclypeal and genal suturae almost indistinct. Eyes small, their dorsal parts small, slit-shaped in some species, ventral parts larger, sub-rectangular. Distance between eye and gula approximately twice the width of eye in ventral view. Gula with longitudinal groove (
<figureCitation box="[152,265,1440,1464]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="3" pageNumber="132">Figure 9F</figureCitation>
). Dorsal surface of clypeus rugose in most species, frons densely punctate to rugose.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002599" ID-Zenodo-Dep="4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="2" pageNumber="131" startId="2.[212,279,1608,1630]" targetBox="[212,1172,1197,1575]" targetPageId="2">
<paragraph blockId="2.[212,1171,1608,1688]" pageId="2" pageNumber="131">
Figure 1.
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[316,434,1608,1629]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis box="[316,434,1608,1629]" italics="true" pageId="2" pageNumber="131">Delopleurus</emphasis>
</taxonomicName>
spp. (A)
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[532,642,1608,1630]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
<emphasis box="[532,642,1608,1630]" italics="true" pageId="2" pageNumber="131">D. fossatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[649,732,1608,1629]" pageId="2" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
, lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B)
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[599,692,1637,1659]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[599,692,1637,1659]" italics="true" pageId="2" pageNumber="131">D. pullus</emphasis>
</taxonomicName>
, male; (C)
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[817,910,1637,1659]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[817,910,1637,1659]" italics="true" pageId="2" pageNumber="131">D. pullus</emphasis>
</taxonomicName>
, female; (D)
<taxonomicName authorityName="Frolov et Cambefort" authorityYear="2014" box="[1056,1166,1637,1658]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis box="[1056,1166,1637,1658]" italics="true" pageId="2" pageNumber="131">D.naviauxi</emphasis>
</taxonomicName>
, female, holotype).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002601" ID-Zenodo-Dep="4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="3" pageNumber="132" startId="3.[144,211,1201,1223]" targetBox="[148,1104,172,1169]" targetPageId="3">
<paragraph blockId="3.[144,1104,1201,1252]" pageId="3" pageNumber="132">
Figure 2.
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[249,367,1201,1222]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis box="[249,367,1201,1222]" italics="true" pageId="3" pageNumber="132">Delopleurus</emphasis>
</taxonomicName>
spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C)
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[571,688,1230,1252]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
<emphasis box="[571,688,1230,1252]" italics="true" pageId="3" pageNumber="132">D. naviauxi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[696,779,1231,1252]" pageId="3" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
; (D–G)
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[865,995,1230,1252]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="pubescens" status="sp. nov.">
<emphasis box="[865,995,1230,1252]" italics="true" pageId="3" pageNumber="132">D. pubescens</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1003,1084,1231,1252]" pageId="3" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph blockId="3.[144,1104,1313,1688]" pageId="3" pageNumber="132">Pronotum more or less trapezoidal, about two times wider than long. Anterior and lateral margins with distinct border, base with or without border. Pronotum is not excavated, without horns or ridges but with fine short longitudinal groove basally along the midline.</paragraph>
<paragraph blockId="3.[144,1104,1313,1688]" lastBlockId="4.[212,1172,1565,1685]" lastPageId="4" lastPageNumber="133" pageId="3" pageNumber="132">
Elytra trapezoidal, as wide as long, shiny to opaque, with deep sinuation on lateral margins receiving convex metepisternum (
<figureCitation box="[703,823,1664,1688]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="3" pageNumber="132">Figure 1A</figureCitation>
). Striae fine but distinct, punctate; striae 1–7 reach base of elytron, stria 8 reaches the sinuation but not the base; striae 9–10 are very close to epipleura and mostly inseparable from each other except apically. Elytral intervals flat to convex, with minute punctation. Elytra fused along suture. Scutellum not visible from above.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002603" ID-Zenodo-Dep="4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="4" pageNumber="133" startId="4.[212,279,1464,1486]" targetBox="[212,1172,168,1432]" targetPageId="4">
<paragraph blockId="4.[212,1171,1464,1515]" pageId="4" pageNumber="133">
Figure 3.
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[315,497,1464,1485]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[315,497,1464,1485]" italics="true" pageId="4" pageNumber="133">Delopleurus pullus</emphasis>
</taxonomicName>
. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002605" ID-Zenodo-Dep="4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="5" pageNumber="134" startId="5.[144,211,1307,1329]" targetBox="[165,1079,176,1275]" targetPageId="5">
<paragraph blockId="5.[144,1104,1307,1359]" pageId="5" pageNumber="134">
Figure 4.
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[253,509,1308,1329]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="darrenmanni" status="sp. nov.">
<emphasis box="[253,509,1308,1329]" italics="true" pageId="5" pageNumber="134">Delopleurus darrenmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[522,608,1308,1329]" pageId="5" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
(A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
</paragraph>
</caption>
<paragraph blockId="5.[144,1104,1419,1667]" pageId="5" pageNumber="134">
Wings well developed and feature a number of characters that
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[904,1035,1420,1444]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis box="[904,1035,1420,1444]" italics="true" pageId="5" pageNumber="134">Delopleurus</emphasis>
</taxonomicName>
share with species of the
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[370,496,1452,1476]" class="Insecta" family="Scarabaeidae" genus="Sarophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis box="[370,496,1452,1476]" italics="true" pageId="5" pageNumber="134">Sarophorus</emphasis>
</taxonomicName>
group of genera (
<bibRefCitation author="Frolov AV &amp; Akhmetova LA &amp; Scholtz CH" box="[710,922,1451,1475]" pageId="5" pageNumber="134" pagination="1477 - 1508" refId="ref9962" refString="Frolov AV, Akhmetova LA, Scholtz CH. 2008. Revision of the obligate mushroom- feeding African &quot; dung beetle &quot; genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae). J Nat Hist. 42: 1477 - 1508." type="journal article" year="2008">Frolov et al. 2008</bibRefCitation>
): brown colour except for very basal part, reduced anal area (veins J and AP
<subScript attach="left" box="[862,895,1494,1510]" fontSize="7" pageId="5" pageNumber="134">3+4</subScript>
are absent), CuA widened apically along the wing margin, RA
<subScript attach="left" box="[664,674,1525,1541]" fontSize="7" pageId="5" pageNumber="134">4</subScript>
does not reach the wing margin and becomes wide and indistinct apically.
</paragraph>
<paragraph blockId="5.[144,1104,1419,1667]" lastBlockId="6.[212,1172,168,607]" lastPageId="6" lastPageNumber="135" pageId="5" pageNumber="134">
Anterior tibiae have three outer teeth. Margin basad of third tooth feebly or not serrate. Spur of anterior tibia more or less bifurcated in males and apically acute to rounded and curved inwards in females. In one species (
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[768,897,1644,1667]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
<emphasis box="[768,897,1644,1667]" italics="true" pageId="5" pageNumber="134">D. naviauxi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[904,994,1643,1667]" pageId="5" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
), anterior tibia with one small acute tooth between first outer tooth and spur (
<figureCitation box="[968,1084,168,192]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="6" pageNumber="135">Figure 1D</figureCitation>
). Outer margins of middle and posterior tibiae without transverse carinae, serrate. Tarsi of all legs well developed, shorter than tibiae. Claws about half the length of fifth tarsal segment.
</paragraph>
<paragraph blockId="6.[212,1172,168,607]" pageId="6" pageNumber="135">
Pygidium about twice as wide as long, its sculpture is species-specific and, in some species, differs strongly from that in the other scarabaeine taxa. In some
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[1040,1171,328,352]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis box="[1040,1171,328,352]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
</taxonomicName>
species basal and apical borders are relatively narrow and the disc occupies most of pygidium (
<figureCitation box="[334,447,391,415]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="6" pageNumber="135">Figure 2E</figureCitation>
). In others, however, the borders are much wider and the disc is almost slit-shaped (
<figureCitation box="[444,563,423,447]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="6" pageNumber="135">Figure 2B</figureCitation>
). Sculpture of pygidium is sex-dependent in some species.
</paragraph>
<paragraph blockId="6.[212,1172,168,607]" pageId="6" pageNumber="135">
Aedeagus of typical scarabaeine shape (
<figureCitation box="[722,849,487,511]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="6" pageNumber="135">Figures 2F</figureCitation>
,
<figureCitation box="[868,901,487,511]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="6" pageNumber="135">3C</figureCitation>
,
<figureCitation box="[919,951,487,511]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="6" pageNumber="135">4C</figureCitation>
,
<figureCitation box="[970,1002,487,511]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="6" pageNumber="135">5C</figureCitation>
,
<figureCitation box="[1021,1054,487,511]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="6" pageNumber="135">6C</figureCitation>
,
<figureCitation box="[1072,1105,487,511]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="6" pageNumber="135">7C</figureCitation>
,
<figureCitation box="[1123,1156,487,511]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="6" pageNumber="135">8C</figureCitation>
). Phallobase with two symmetrical tubercles. Parameres symmetrical, without setae apically, slightly distinctive in different species. Internal sac with armature consisting of a few sclerites of complex shape.
</paragraph>
<paragraph blockId="6.[212,1172,671,798]" box="[212,425,671,695]" pageId="6" pageNumber="135">
<heading box="[212,425,671,695]" fontSize="10" level="3" pageId="6" pageNumber="135" reason="5">
<emphasis box="[212,425,671,695]" italics="true" pageId="6" pageNumber="135">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[212,1172,671,798]" pageId="6" pageNumber="135">
Males can be easily separated from females by having more or less bifurcated spurs of anterior tibiae (
<figureCitation box="[394,548,742,766]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="6" pageNumber="135">Figure 1B, C</figureCitation>
). Some species differ strongly in the sculpture of the pygidium.
</paragraph>
<paragraph blockId="6.[212,1172,862,1054]" box="[212,452,862,886]" pageId="6" pageNumber="135">
<heading box="[212,452,862,886]" fontSize="10" level="3" pageId="6" pageNumber="135" reason="5">
<emphasis box="[212,452,862,886]" italics="true" pageId="6" pageNumber="135">Diagnostic characters</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[212,1172,862,1054]" pageId="6" pageNumber="135">
Unlike other scarabaeines,
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[519,650,902,926]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis box="[519,650,902,926]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
</taxonomicName>
species can be best separated by the sculpture of the pygidium. Although shape of the parameres and internal sac armature shows some species specific features, they are not considered reliable diagnostic characters. However, availability of the other characters, especially that of the pygidium, makes dissection unnecessary in routine identification.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="135" type="distribution">
<paragraph blockId="6.[212,1172,1117,1532]" box="[212,344,1117,1141]" pageId="6" pageNumber="135">
<heading box="[212,344,1117,1141]" fontSize="10" level="3" pageId="6" pageNumber="135" reason="5">
<emphasis box="[212,344,1117,1141]" italics="true" pageId="6" pageNumber="135">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[212,1172,1117,1532]" pageId="6" pageNumber="135">
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,343,1157,1181]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis box="[212,343,1157,1181]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
</taxonomicName>
has a disjunctive range covering most of the Afrotropical Region and Indian Peninsula. The ranges of species are rather limited (except for
<taxonomicName authority="Janssens" authorityName="Janssens" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis box="[1067,1171,1189,1213]" italics="true" pageId="6" pageNumber="135">D. gilleti</emphasis>
Janssens
</taxonomicName>
) although this may be a result of undersampling. Two regional faunas, Afrotropical and Indian, share no common species, which may indicate that they have long developed in isolation. The genus range disjunction can be best explained by the contraction of a previously very large range since the late Neogene. It is possible that in the late Miocene, when huge regions of Africa and Eurasia, including the territories of the present-day Sahara and Mediterranean, were covered by the African-type savannas,
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[497,628,1412,1436]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis box="[497,628,1412,1436]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
</taxonomicName>
was widely distributed in the Eurasia. No palaeontological data exist on the past occurrence of
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[855,986,1444,1468]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis box="[855,986,1444,1468]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
</taxonomicName>
in Europe and extratropical Asia, but considering very low taphonomic potential of these soil dwellers, discovery of any fossil
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[522,653,1508,1532]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis box="[522,653,1508,1532]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
</taxonomicName>
seems highly improbable.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="137" pageId="6" pageNumber="135" type="biology_ecology">
<paragraph blockId="6.[212,1172,1596,1692]" box="[212,297,1596,1620]" pageId="6" pageNumber="135">
<heading box="[212,297,1596,1620]" fontSize="10" level="3" pageId="6" pageNumber="135" reason="5">
<emphasis box="[212,297,1596,1620]" italics="true" pageId="6" pageNumber="135">Biology</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[212,1172,1596,1692]" lastBlockId="8.[212,1172,850,1384]" lastPageId="8" lastPageNumber="137" pageId="6" pageNumber="135">
Little information is available about the biology of
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[821,952,1636,1660]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis box="[821,952,1636,1660]" italics="true" pageId="6" pageNumber="135">Delopleurus</emphasis>
</taxonomicName>
and the immature stages of the genus are unknown. In
<collectingCountry box="[635,773,1667,1692]" name="Ivory Coast" pageId="6" pageNumber="135">Ivory Coast</collectingCountry>
, a few specimens of
<taxonomicName box="[1011,1110,1667,1691]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis box="[1011,1110,1667,1691]" italics="true" pageId="6" pageNumber="135">D. gilleti</emphasis>
</taxonomicName>
were collected on mushrooms of
<taxonomicName box="[469,625,1535,1558]" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="GBIF" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="136" phylum="Basidiomycota" rank="genus">
<emphasis box="[469,625,1535,1558]" italics="true" pageId="7" pageNumber="136">Termitomyces</emphasis>
</taxonomicName>
cf.
<emphasis box="[674,779,1534,1558]" italics="true" pageId="7" pageNumber="136">schimperi</emphasis>
along with more numerous
<taxonomicName authority="Boheman" authorityName="Boheman" authorityYear="1857" box="[144,537,1566,1590]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="nitidipennis">
<emphasis box="[144,412,1566,1590]" italics="true" pageId="7" pageNumber="136">Coptorhina nitidipennis</emphasis>
Boheman
</taxonomicName>
(Yves Cambefort, pers. comm.). In
<collectingCountry box="[1004,1103,1566,1590]" name="Namibia" pageId="7" pageNumber="136">Namibia</collectingCountry>
(
<collectingRegion box="[151,239,1598,1622]" country="Namibia" name="Zambezi" pageId="7" pageNumber="136">Caprivi</collectingRegion>
, Darren Mann, pers. comm.), beetles of
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[746,927,1598,1622]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="darrenmanni" status="sp. nov.">
<emphasis box="[746,927,1598,1622]" italics="true" pageId="7" pageNumber="136">D. darrenmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[942,1037,1599,1622]" pageId="7" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
were collected in mushrooms (
<taxonomicName box="[433,589,1631,1654]" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="GBIF" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="136" phylum="Basidiomycota" rank="genus">
<emphasis box="[433,589,1631,1654]" italics="true" pageId="7" pageNumber="136">Termitomyces</emphasis>
</taxonomicName>
sp.) on and around termite mounds, also in association with
<taxonomicName authorityName="Hope" authorityYear="1833" box="[342,467,1662,1686]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis box="[342,467,1662,1686]" italics="true" pageId="7" pageNumber="136">Coptorhina</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[488,653,1662,1686]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="nitidipennis">
<emphasis box="[488,653,1662,1686]" italics="true" pageId="7" pageNumber="136">C. nitidipennis</emphasis>
</taxonomicName>
and
<taxonomicName authority="Arrow" authorityName="Arrow" authorityYear="1906" box="[720,949,1662,1686]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="angolensis">
<emphasis box="[720,858,1662,1686]" italics="true" pageId="7" pageNumber="136">C.angolensis</emphasis>
Arrow
</taxonomicName>
). There were apparently
<taxonomicName authorityName="Hope" authorityYear="1833" box="[340,465,850,874]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[340,465,850,874]" italics="true" pageId="8" pageNumber="137">Coptorhina</emphasis>
</taxonomicName>
burrows under the fungal umbrella or a few centimetres away. No smaller burrows of
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[475,606,882,906]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[475,606,882,906]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
</taxonomicName>
were seen however. It is possible that
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[1040,1171,882,906]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[1040,1171,882,906]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
</taxonomicName>
can use mushroom pieces buried by
<taxonomicName authorityName="Hope" authorityYear="1833" box="[628,753,914,938]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[628,753,914,938]" italics="true" pageId="8" pageNumber="137">Coptorhina</emphasis>
</taxonomicName>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002607" ID-Zenodo-Dep="4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="7" pageNumber="136" startId="7.[144,211,1422,1444]" targetBox="[145,1103,172,1390]" targetPageId="7">
<paragraph blockId="7.[144,1103,1422,1473]" pageId="7" pageNumber="136">
Figure 5.
<taxonomicName box="[247,428,1422,1443]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis box="[247,428,1422,1443]" italics="true" pageId="7" pageNumber="136">Delopleurus gilleti</emphasis>
</taxonomicName>
. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002609" ID-Zenodo-Dep="4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="8" pageNumber="137" startId="8.[212,279,738,760]" targetBox="[214,1170,168,697]" targetPageId="8">
<paragraph blockId="8.[212,1171,738,789]" pageId="8" pageNumber="137">
Figure 6.
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[317,525,738,759]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="krikkeni" status="sp. nov.">
<emphasis box="[317,525,738,759]" italics="true" pageId="8" pageNumber="137">Delopleurus krikkeni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[533,617,738,759]" pageId="8" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map.
</paragraph>
</caption>
<paragraph blockId="8.[212,1172,850,1384]" pageId="8" pageNumber="137">
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[254,385,945,969]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[254,385,945,969]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
</taxonomicName>
beetles have never been found in or under dung pads, and they have never been collected by pitfalls baited with dung or carrion [Philippe Walter collected a few specimens in traps baited with fish (Yves Cambefort, pers. comm.) but these were most probably chance captures].
</paragraph>
<paragraph blockId="8.[212,1172,850,1384]" pageId="8" pageNumber="137">
Available data suggest that
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[589,721,1073,1097]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[589,721,1073,1097]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
</taxonomicName>
are specialist feeders of agaric mushrooms (
<taxonomicName authorityName="Underwood" authorityYear="1899" box="[306,430,1105,1129]" class="Agaricomycetes" higherTaxonomySource="GBIF" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="137" phylum="Basidiomycota" rank="order">Agaricales</taxonomicName>
). They seem to be associated with the termitophilous genus
<taxonomicName box="[212,368,1138,1161]" class="Agaricomycetes" family="Lyophyllaceae" genus="Termitomyces" higherTaxonomySource="GBIF" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="137" phylum="Basidiomycota" rank="genus">
<emphasis box="[212,368,1138,1161]" italics="true" pageId="8" pageNumber="137">Termitomyces</emphasis>
</taxonomicName>
, a widespread fungal genus with large fruit bodies. In the case of
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,343,1169,1193]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[212,343,1169,1193]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Hope" authorityYear="1833" box="[405,530,1169,1193]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[405,530,1169,1193]" italics="true" pageId="8" pageNumber="137">Coptorhina</emphasis>
</taxonomicName>
, feeding on mushrooms can be a secondary adaptation from the putatively generalist saprophagy of their ancestors. Feeding on mushrooms as an adaptation to arid biotopes in beetles was hypothesized by
<bibRefCitation author="Striganova BR" box="[957,1161,1232,1256]" pageId="8" pageNumber="137" refId="ref10319" refString="Striganova BR. 1980. Pitanie pochvennykh saprofagov [Feeding of soil saprophages]. Moscow: Nauka." type="book" year="1980">Striganova (1980)</bibRefCitation>
.
</paragraph>
<paragraph blockId="8.[212,1172,850,1384]" pageId="8" pageNumber="137">
All
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[299,430,1264,1288]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[299,430,1264,1288]" italics="true" pageId="8" pageNumber="137">Delopleurus</emphasis>
</taxonomicName>
specimens examined have well developed wings and apparently are good fliers. But no specimens have been collected at light, according to the labels and data from collectors. This, along with small eyes (typical for all members of the
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[212,338,1360,1384]" class="Insecta" family="Scarabaeidae" genus="Sarophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis box="[212,338,1360,1384]" italics="true" pageId="8" pageNumber="137">Sarophorus</emphasis>
</taxonomicName>
group genera), suggests that the beetles have diurnal flight activity.
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="141" pageId="8" pageNumber="137" type="key">
<paragraph blockId="8.[212,510,1459,1524]" box="[212,510,1459,1484]" pageId="8" pageNumber="137">
<heading box="[212,510,1459,1484]" fontSize="10" level="3" pageId="8" pageNumber="137" reason="5">
<emphasis bold="true" box="[212,510,1459,1484]" pageId="8" pageNumber="137">
Key to
<emphasis bold="true" box="[295,510,1459,1483]" italics="true" pageId="8" pageNumber="137">
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[295,426,1459,1483]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">Delopleurus</taxonomicName>
species
</emphasis>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="8.[212,510,1459,1524]" box="[212,280,1500,1524]" pageId="8" pageNumber="137">
<emphasis box="[212,280,1500,1524]" italics="true" pageId="8" pageNumber="137">Males</emphasis>
</paragraph>
<key lastPageId="11" lastPageNumber="140" pageId="8" pageNumber="137">
<paragraph blockId="8.[212,1172,1571,1691]" pageId="8" pageNumber="137">
<keyLead pageId="8" pageNumber="137">
<keyStep pageId="8" pageNumber="137">
1. Disc of pygidium more or less densely punctate (
<figureCitation box="[801,925,1571,1595]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="8" pageNumber="137">Figures 3B</figureCitation>
,
<figureCitation box="[939,970,1571,1595]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="8" pageNumber="137">4B</figureCitation>
,
<figureCitation box="[986,1016,1571,1595]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="8" pageNumber="137">5B</figureCitation>
) or granulate with short brown setae (
<figureCitation box="[526,641,1603,1627]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="8" pageNumber="137">Figure 2E</figureCitation>
). Species from Africa .................................. 2 Disc of pygidium almost smooth, without setae (
<figureCitation box="[801,914,1635,1659]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="8" pageNumber="137">Figure 8B</figureCitation>
). Species from Asia ... ...............................................................................................................
<taxonomicName baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[1064,1171,1668,1691]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="parvus">
<emphasis box="[1064,1171,1668,1691]" italics="true" pageId="8" pageNumber="137">D. parvus</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002611" ID-Zenodo-Dep="4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="9" pageNumber="138" startId="9.[144,211,1410,1432]" targetBox="[145,1103,169,1378]" targetPageId="9">
<paragraph blockId="9.[144,1103,1410,1461]" pageId="9" pageNumber="138">
Figure 7.
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[249,453,1410,1431]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
<emphasis box="[249,453,1410,1431]" italics="true" pageId="9" pageNumber="138">Delopleurus fossatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[462,544,1411,1432]" pageId="9" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
(A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
</paragraph>
</caption>
<paragraph blockId="9.[144,1109,1511,1599]" pageId="9" pageNumber="138">
<keyLead pageId="9" pageNumber="138">
<keyStep pageId="9" pageNumber="138">
2. Disc of pygidium rugose and with sparse brown setae (
<figureCitation box="[810,924,1511,1535]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="9" pageNumber="138">Figure 2E</figureCitation>
) ...................... .............................................................................................
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[861,1005,1543,1567]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="pubescens" status="sp. nov.">
<emphasis box="[861,1005,1543,1567]" italics="true" pageId="9" pageNumber="138">D. pubescens</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1014,1104,1544,1567]" pageId="9" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
Disc of pygidium with different sculpture, without setae. ................................. 3
</keyStep>
</keyLead>
</paragraph>
<keyStep lastPageId="10" lastPageNumber="139" pageId="9" pageNumber="138">
<paragraph blockId="9.[144,1103,1623,1679]" pageId="9" pageNumber="138">
<keyLead pageId="9" pageNumber="138">
3. Border of pygidium relatively narrow; punctation of pygidium disc sparser (
<figureCitation box="[189,303,1655,1679]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="9" pageNumber="138">Figure 3B</figureCitation>
) .............................................................................................
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[1003,1103,1655,1679]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[1003,1103,1655,1679]" italics="true" pageId="9" pageNumber="138">D. pullus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002613" ID-Zenodo-Dep="4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="10" pageNumber="139" startId="10.[212,279,1391,1413]" targetBox="[212,1171,171,1359]" targetPageId="10">
<paragraph blockId="10.[212,1172,1391,1442]" pageId="10" pageNumber="139">
Figure 8.
<taxonomicName baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[317,507,1391,1413]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="parvus">
<emphasis box="[317,507,1391,1413]" italics="true" pageId="10" pageNumber="139">Delopleurus parvus</emphasis>
</taxonomicName>
. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map.
</paragraph>
</caption>
<paragraph blockId="10.[248,1172,1503,1559]" pageId="10" pageNumber="139">
<keyLead pageId="10" pageNumber="139">
Border of pygidium thicker (
<figureCitation box="[593,720,1503,1527]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="10" pageNumber="139">Figures 4B</figureCitation>
,
<figureCitation box="[741,772,1503,1527]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="10" pageNumber="139">6B</figureCitation>
,
<figureCitation box="[792,823,1504,1527]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="10" pageNumber="139">7B</figureCitation>
), if similar to
<taxonomicName authorityName="Boheman" authorityYear="1857" box="[1005,1110,1503,1527]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[1005,1110,1503,1527]" italics="true" pageId="10" pageNumber="139">D. pullus</emphasis>
</taxonomicName>
then punctation of pygidium disc coarser (
<figureCitation box="[679,793,1535,1559]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="10" pageNumber="139">Figure 5B</figureCitation>
) .............................................. 4
</keyLead>
</paragraph>
</keyStep>
<paragraph blockId="10.[212,1178,1583,1671]" pageId="10" pageNumber="139">
<keyLead pageId="10" pageNumber="139">
<keyStep pageId="10" pageNumber="139">
4. Disc of pygidium almost as long as apical border (
<figureCitation box="[829,943,1583,1607]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="10" pageNumber="139">Figure 4B</figureCitation>
) ............................. .........................................................................................
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[897,1073,1615,1639]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="139" phylum="Arthropoda" rank="species" species="darrenmanni" status="sp. nov.">
<emphasis box="[897,1073,1615,1639]" italics="true" pageId="10" pageNumber="139">D. darrenmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1082,1172,1615,1638]" pageId="10" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
Disc of pygidium 1.5–2 times longer than apical border ................................... 5
</keyStep>
</keyLead>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4002615" ID-Zenodo-Dep="4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="11" pageNumber="140" startId="11.[144,211,939,961]" targetBox="[147,1104,168,906]" targetPageId="11">
<paragraph blockId="11.[144,1104,939,1049]" pageId="11" pageNumber="140">
Figure 9.
<taxonomicName authorityName="Erichson" authorityYear="1847" box="[252,370,939,960]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis box="[252,370,939,960]" italics="true" pageId="11" pageNumber="140">Delopleurus</emphasis>
</taxonomicName>
spp. and
<taxonomicName authorityName="Frolov" authorityYear="2014" baseAuthorityName="Frolov" baseAuthorityYear="1963" box="[482,732,939,960]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis box="[482,732,939,960]" italics="true" pageId="11" pageNumber="140">Metacatharsius janssensi</emphasis>
</taxonomicName>
. (A, E) habitus (A,
<taxonomicName box="[955,1064,939,961]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="striatus">
<emphasis box="[955,1064,939,961]" italics="true" pageId="11" pageNumber="140">D. striatus</emphasis>
</taxonomicName>
, E,
<taxonomicName authorityName="Frolov" authorityYear="2014" baseAuthorityName="Frolov" baseAuthorityYear="1963" box="[144,266,969,990]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis box="[144,266,969,990]" italics="true" pageId="11" pageNumber="140">M. janssensi</emphasis>
</taxonomicName>
); (B)
<taxonomicName box="[325,430,969,990]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="striatus">
<emphasis box="[325,430,969,990]" italics="true" pageId="11" pageNumber="140">D. striatus</emphasis>
</taxonomicName>
, head and pronotum in dorsal view; (C)
<taxonomicName box="[843,947,969,990]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="striatus">
<emphasis box="[843,947,969,990]" italics="true" pageId="11" pageNumber="140">D. striatus</emphasis>
</taxonomicName>
, pygidium; (D)
<taxonomicName box="[144,252,998,1020]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="striatus">
<emphasis box="[144,252,998,1020]" italics="true" pageId="11" pageNumber="140">D. striatus</emphasis>
</taxonomicName>
, locality map; (F)
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[458,572,998,1020]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
<emphasis box="[458,572,998,1020]" italics="true" pageId="11" pageNumber="140">D. fossatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[582,669,998,1019]" pageId="11" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
, ventral view of head showing gula with distinct longitudinal groove.
</paragraph>
</caption>
<paragraph blockId="11.[144,1103,1113,1233]" pageId="11" pageNumber="140">
<keyLead pageId="11" pageNumber="140">
<keyStep pageId="11" pageNumber="140">
5. Disc of pygidium feebly convex, almost in plane with basal border (in lateral view,
<figureCitation box="[180,294,1145,1169]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="11" pageNumber="140">Figure 5B</figureCitation>
) ..............................................................................................
<taxonomicName box="[1005,1103,1145,1169]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis box="[1005,1103,1145,1169]" italics="true" pageId="11" pageNumber="140">D. gilleti</emphasis>
</taxonomicName>
Disc of pygidium strongly convex, distinctly separated from basal border by a deep hollow (in lateral view,
<figureCitation box="[516,631,1209,1233]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="11" pageNumber="140">Figure 6B</figureCitation>
,
<figureCitation box="[646,677,1210,1233]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="11" pageNumber="140">7B</figureCitation>
) ..................................................... 6
</keyStep>
</keyLead>
</paragraph>
<paragraph blockId="11.[144,1104,1257,1440]" pageId="11" pageNumber="140">
<keyLead pageId="11" pageNumber="140">
<keyStep pageId="11" pageNumber="140">
6. Elytra with well-developed humeral umbones, sinuate behind umbones in dorsal view (
<figureCitation box="[251,370,1289,1313]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="11" pageNumber="140">Figure 6A</figureCitation>
). Basal border of pygidium strongly widened medially (
<figureCitation captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="11" pageNumber="140">Figure 6B</figureCitation>
) ........................................................................................
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[878,1005,1321,1345]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="krikkeni" status="sp. nov.">
<emphasis box="[878,1005,1321,1345]" italics="true" pageId="11" pageNumber="140">D. krikkeni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1014,1104,1321,1344]" pageId="11" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
Elytra with feebly developed humeral umbones, feebly or not sinuate behind umbones in dorsal view (
<figureCitation box="[473,591,1384,1408]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="11" pageNumber="140">Figure 7A</figureCitation>
). Basal border of pygidium slightly widened medially (
<figureCitation box="[297,411,1416,1440]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="11" pageNumber="140">Figure 7B</figureCitation>
) ..............................................................
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[881,1005,1416,1440]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="140" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
<emphasis box="[881,1005,1416,1440]" italics="true" pageId="11" pageNumber="140">D. fossatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1014,1104,1417,1440]" pageId="11" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</keyStep>
</keyLead>
</paragraph>
</key>
<paragraph blockId="11.[144,234,1500,1524]" box="[144,234,1500,1524]" pageId="11" pageNumber="140">
<heading box="[144,234,1500,1524]" fontSize="10" level="3" pageId="11" pageNumber="140" reason="5">
<emphasis box="[144,234,1500,1524]" italics="true" pageId="11" pageNumber="140">Females</emphasis>
</heading>
</paragraph>
<key lastPageId="12" lastPageNumber="141" pageId="11" pageNumber="140">
<paragraph blockId="11.[144,1103,1571,1691]" pageId="11" pageNumber="140">
<keyLead pageId="11" pageNumber="140">
<keyStep pageId="11" pageNumber="140">
1. Disc of pygidium more or less densely punctate, or slit-shaped, or granulate (with short brown setae). Species from Africa ............................................................ 2 Disc of pygidium almost smooth (
<figureCitation box="[619,742,1635,1659]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="11" pageNumber="140">Figure 8E</figureCitation>
) or granulate, without setae (
<figureCitation box="[192,308,1667,1691]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="11" pageNumber="140">Figure 9C</figureCitation>
). Species from Asia .......................................................................... 7
</keyStep>
</keyLead>
</paragraph>
<paragraph blockId="12.[212,1172,168,288]" pageId="12" pageNumber="141">
<keyLead pageId="12" pageNumber="141">
<keyStep pageId="12" pageNumber="141">
2. Fore tibia with small acute tooth near apical spur (
<figureCitation box="[819,936,168,192]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="12" pageNumber="141">Figure 1D</figureCitation>
). Pygidium with slitshaped pubescent disc (
<figureCitation box="[516,629,200,224]" captionStart="Figure 2" captionStartId="3.[144,211,1201,1223]" captionTargetBox="[148,1104,172,1169]" captionTargetId="figure@3.[144,1104,168,1169]" captionTargetPageId="3" captionText="Figure 2. Delopleurus spp. (A, D) habitus; (B, E) pygidium; (F) aedeagus in lateral view and internal sac; (C, G) locality maps. (A–C) D. naviauxi sp. nov.; (D–G) D. pubescens sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4002601" httpUri="https://zenodo.org/record/4002601/files/figure.png" pageId="12" pageNumber="141">Figure 2B</figureCitation>
) .......................................
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[943,1072,201,224]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
<emphasis box="[943,1072,201,224]" italics="true" pageId="12" pageNumber="141">D. naviauxi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1082,1172,201,224]" pageId="12" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
Fore tibia without tooth near apical spur (
<figureCitation box="[762,882,232,256]" captionStart="Figure 1" captionStartId="2.[212,279,1608,1630]" captionTargetBox="[212,1172,1197,1575]" captionTargetId="figure@2.[212,1172,1196,1576]" captionTargetPageId="2" captionText="Figure 1. Delopleurus spp. (A) D. fossatus sp. nov., lateral view of body (sinuate epipleuron is arrowed); (B–D) anterior tibiae: (B) D. pullus, male; (C) D. pullus, female; (D) D.naviauxi, female, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4002599" httpUri="https://zenodo.org/record/4002599/files/figure.png" pageId="12" pageNumber="141">Figure 1C</figureCitation>
). Sculpture of pygidium different; if disc of pygidium slit-shaped then it lacks setation (
<figureCitation box="[968,1081,264,288]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="12" pageNumber="141">Figure 4E</figureCitation>
) .... .. 3
</keyStep>
</keyLead>
</paragraph>
<paragraph blockId="12.[212,1172,312,399]" pageId="12" pageNumber="141">
<keyLead pageId="12" pageNumber="141">
<keyStep pageId="12" pageNumber="141">
3. Borders of pygidium relatively narrow; disc feebly convex, granulate and pubescent (
<figureCitation box="[333,448,344,368]" captionStart="Figure 3" captionStartId="4.[212,279,1464,1486]" captionTargetBox="[212,1172,168,1432]" captionTargetId="figure@4.[212,1172,168,1432]" captionTargetPageId="4" captionText="Figure 3. Delopleurus pullus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002603" httpUri="https://zenodo.org/record/4002603/files/figure.png" pageId="12" pageNumber="141">Figure 3E</figureCitation>
) ...................................................................................
<taxonomicName authority="Borders" authorityName="Borders" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="pullus">
<emphasis box="[1071,1171,344,368]" italics="true" pageId="12" pageNumber="141">D. pullus</emphasis>
Borders
</taxonomicName>
of pygidium relatively wide, disc without setation ............................... 4
</keyStep>
</keyLead>
</paragraph>
<paragraph blockId="12.[212,1177,423,511]" pageId="12" pageNumber="141">
<keyLead pageId="12" pageNumber="141">
<keyStep pageId="12" pageNumber="141">
4. Disc of pygidium slit-shaped, with transverse carina (
<figureCitation box="[857,971,423,447]" captionStart="Figure 4" captionStartId="5.[144,211,1307,1329]" captionTargetBox="[165,1079,176,1275]" captionTargetId="figure@5.[164,1083,168,1276]" captionTargetPageId="5" captionText="Figure 4. Delopleurus darrenmanni sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002605" httpUri="https://zenodo.org/record/4002605/files/figure.png" pageId="12" pageNumber="141">Figure 4E</figureCitation>
) ......................... .........................................................................................
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[897,1073,455,479]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="darrenmanni" status="sp. nov.">
<emphasis box="[897,1073,455,479]" italics="true" pageId="12" pageNumber="141">D. darrenmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1082,1172,456,479]" pageId="12" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
Disc of pygidium 1.5–2 times longer than apical border thickness ................... 5
</keyStep>
</keyLead>
</paragraph>
<paragraph blockId="12.[212,1172,535,655]" pageId="12" pageNumber="141">
<keyLead pageId="12" pageNumber="141">
<keyStep pageId="12" pageNumber="141">
5. Disc of pygidium feebly convex, almost in plane with basal border (in lateral view,
<figureCitation box="[248,362,567,591]" captionStart="Figure 5" captionStartId="7.[144,211,1422,1444]" captionTargetBox="[145,1103,172,1390]" captionTargetId="figure@7.[144,1104,168,1390]" captionTargetPageId="7" captionText="Figure 5. Delopleurus gilleti. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002607" httpUri="https://zenodo.org/record/4002607/files/figure.png" pageId="12" pageNumber="141">Figure 5E</figureCitation>
) ..............................................................................................
<taxonomicName box="[1073,1171,567,591]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis box="[1073,1171,567,591]" italics="true" pageId="12" pageNumber="141">D. gilleti</emphasis>
</taxonomicName>
Disc of pygidium strongly convex, distinctly separated from basal border by a deep hollow (in lateral view,
<figureCitation box="[584,698,630,654]" captionStart="Figure 7" captionStartId="9.[144,211,1410,1432]" captionTargetBox="[145,1103,169,1378]" captionTargetId="figure@9.[144,1104,168,1378]" captionTargetPageId="9" captionText="Figure 7. Delopleurus fossatus sp. nov. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002611" httpUri="https://zenodo.org/record/4002611/files/figure.png" pageId="12" pageNumber="141">Figure 7E</figureCitation>
) ...............................
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[949,1073,631,655]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
<emphasis box="[949,1073,631,655]" italics="true" pageId="12" pageNumber="141">D. fossatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1082,1172,631,654]" pageId="12" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</keyStep>
</keyLead>
</paragraph>
<paragraph blockId="12.[212,1172,678,798]" pageId="12" pageNumber="141">
<keyLead pageId="12" pageNumber="141">
<keyStep pageId="12" pageNumber="141">
6. Elytral striae with larger punctures (
<figureCitation box="[683,805,678,702]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="12" pageNumber="141">Figure 8D</figureCitation>
). Disc of pygidium smooth to indistinctly rugose (
<figureCitation box="[472,587,710,734]" captionStart="Figure 8" captionStartId="10.[212,279,1391,1413]" captionTargetBox="[212,1171,171,1359]" captionTargetId="figure@10.[212,1172,168,1359]" captionTargetPageId="10" captionText="Figure 8. Delopleurus parvus. (A, D) habitus (A, male, D, female); (B, E) pygidium (B, male, E, female); (C) aedeagus in lateral view and internal sac; (F) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002613" httpUri="https://zenodo.org/record/4002613/files/figure.png" pageId="12" pageNumber="141">Figure 8E</figureCitation>
) ..............................................................
<taxonomicName authority="Elytral" authorityName="Elytral" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="parvus">
<emphasis box="[1063,1171,711,734]" italics="true" pageId="12" pageNumber="141">D. parvus</emphasis>
Elytral
</taxonomicName>
striae with smaller punctures (
<figureCitation box="[698,819,742,766]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="12" pageNumber="141">Figure 9A</figureCitation>
). Disc of pygidium granulate (
<figureCitation box="[260,376,774,798]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="12" pageNumber="141">Figure 9C</figureCitation>
) ..........................................................................................
<taxonomicName box="[1054,1171,775,798]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="141" phylum="Arthropoda" rank="species" species="striatus">
<emphasis box="[1054,1171,775,798]" italics="true" pageId="12" pageNumber="141">D. striatus</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
</key>
</subSubSection>
</treatment>
</document>

63
data/F9/1E/EB/F91EEB8024A0647F542B0F63F044C038.xml Normal file
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3913712" ID-GBIF-Taxon="164811144" ID-Zenodo-Dep="3913712" LSID="urn:lsid:zoobank.org:act:A76FA440-DAFD-4BA6-965C-08EFC2EC726D" httpUri="http://treatment.plazi.org/id/F91EEB8024A0647F542B0F63F044C038" lastPageNumber="624" pageNumber="624" sourcePageUrl="https://www.biodiversitylibrary.org/page/727535">
<subSubSection type="nomenclature">
<paragraph pageNumber="624">
<taxonomicName LSID="urn:lsid:zoobank.org:act:A76FA440-DAFD-4BA6-965C-08EFC2EC726D" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Blaberidae" genus="Scorpio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" phylum="Arthropoda" rank="species" species="afer" status="spec. nov." zbkClass="Insecta" zbkKingdom="Animalia" zbkOrder="Aptera">Scorpio afer</taxonomicName>
[
<taxonomicNameLabel rank="species">spec. nov.</taxonomicNameLabel>
]
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="624">S. pectinibus 13-dentatis, manibus subcordatis pilosis.</paragraph>
<paragraph pageNumber="624">
<emphasis italics="true">Swamm. bibl. t.</emphasis>
3
<emphasis italics="true">f.</emphasis>
3. @/
<emphasis italics="true">Seb. mus.</emphasis>
1.
<emphasis italics="true">t.</emphasis>
70.
<emphasis italics="true">f.</emphasis>
4.
</paragraph>
<paragraph pageNumber="624">
<emphasis italics="true">quart. t.</emphasis>
6.
</paragraph>
</subSubSection>
<subSubSection type="distribution">
<paragraph pageNumber="624">
<emphasis italics="true">Habitat in</emphasis>
India.
</paragraph>
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