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<document id="C4098F8F35384A7A3EA089C6A3CF373A" ID-DOI="10.11646/zootaxa.3974.3.3" ID-GBIF-Dataset="53679d76-ae41-4651-a3fc-b6823993d5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="231780" ID-ZooBank="408C29FE-AF3B-4D4D-950C-7DFED83B2159" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461219790961" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Anker, Arthur" docDate="2015" docId="F911475EFFE81114EBF8FF53FD3E6460" docLanguage="en" docName="zt03974p360.pdf" docOrigin="Zootaxa 3974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Naushonia draconis Anker 2014" docType="treatment" docVersion="16" lastPageNumber="358" masterDocId="05283F26FFF91105EB6FFFC4FFF16112" masterDocTitle="Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key" masterLastPageNumber="360" masterPageNumber="341" pageNumber="358" updateTime="1698603758967" updateUser="plazi">
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<mods:title id="E427B34E243F0C6A3BE850787534AC41">Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key</mods:title>
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<heading id="2A4F4124FFE81114EBF8FF53FDDB61A0" bold="true" box="[151,554,151,178]" fontSize="11" level="1" pageId="17" pageNumber="358" reason="1">
<taxonomicName id="B6B88DCBFFE81114EBF8FF53FDDB61A0" ID-CoL="16176e27-ddea-456e-ac3c-e51e8ebff7ac" authority="Anker, 2014" authorityName="Anker" authorityYear="2014" box="[151,554,151,178]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114EBF8FF53FDDB61A0" bold="true" box="[151,554,151,178]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBF8FF53FE7D61A3" bold="true" box="[151,396,151,177]" italics="true" pageId="17" pageNumber="358">Naushonia draconis</emphasis>
<bibRefCitation id="15298BB9FFE81114EAFCFF5CFDDB61A0" author="Anker" box="[403,554,152,178]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker, 2014</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<figureCitation id="E983EACDFFE81114EBF8FF7EFF2861C1" box="[151,217,186,211]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
</paragraph>
</subSubSection>
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<emphasis id="43CC2A5AFFE81114EBF8FEC6FE92600A" box="[151,355,258,280]" italics="true" pageId="17" pageNumber="358">Naushonia draconis</emphasis>
<treatmentCitation id="F019D059FFE81114EA06FEC6FDE6600A" author="Anker" box="[361,535,258,280]" page="761" pageId="17" pageNumber="358" year="2014">
<bibRefCitation id="15298BB9FFE81114EA06FEC6FE17600A" author="Anker" box="[361,486,258,280]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
: 761
</treatmentCitation>
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, figs. 15 (
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locality:
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, French Antilles).
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</paragraph>
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<subSubSection id="39A2A5C3FFE81114EBF8FE81FCBA60B4" pageId="17" pageNumber="358" type="materials_examined">
<paragraph id="7107F648FFE81114EBF8FE81FCBA60B4" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBF8FE81FE8F604C" bold="true" box="[151,382,325,350]" pageId="17" pageNumber="358">Material examined</emphasis>
.
<collectingCountry id="09AFB6D8FFE81114EAFDFE81FE20604C" box="[402,465,325,350]" name="Cuba" pageId="17" pageNumber="358">Cuba</collectingCountry>
:
<specimenCount id="67BE3DC1FFE81114EA8EFE82FDC3604C" box="[481,562,325,350]" pageId="17" pageNumber="358" type="male">1 male</specimenCount>
(pocl
<quantity id="B6405BADFFE81114E9EBFE81FD12604C" box="[644,739,325,350]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="17" pageNumber="358" unit="mm" value="5.1">5.1 mm</quantity>
),
<specimenCount id="67BE3DC1FFE81114E994FE82FC90604C" box="[763,865,325,350]" pageId="17" pageNumber="358" type="female">1 female</specimenCount>
(pocl
<quantity id="B6405BADFFE81114E8DBFE81FBE2604C" box="[948,1043,325,350]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="17" pageNumber="358" unit="mm" value="5.6">5.6 mm</quantity>
),
<collectingCountry id="09AFB6D8FFE81114EF44FE81FBA2604C" box="[1067,1107,325,350]" name="Iceland" pageId="17" pageNumber="358">Isla</collectingCountry>
de la Juventud,
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,
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,
<quantity id="B6405BADFFE81114EA4EFEADFEAC6093" box="[289,349,361,385]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="358" unit="m" value="11.0">11 m</quantity>
, coral reef, rubble, coll. M. Johnson
<emphasis id="43CC2A5AFFE81114E864FEAEFCB76093" box="[779,838,361,385]" italics="true" pageId="17" pageNumber="358">et al.</emphasis>
,
<date id="0506D088FFE81114E839FEADFC056093" box="[854,1012,361,385]" pageId="17" pageNumber="358" value="2014-06-28">28 June 2014</date>
,
<collectionCode id="17A96E8DFFE81114EF6DFEADFBAE6093" box="[1026,1119,361,385]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="358">MZUSP</collectionCode>
33201; 1 ovigerous female (pocl
<quantity id="B6405BADFFE81114EBB8FE49FEC060B4" box="[215,305,397,422]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="17" pageNumber="358" unit="mm" value="5.2">5.2 mm</quantity>
), same collection data,
<collectionCode id="17A96E8DFFE81114E958FE49FD6060B4" LSID="urn:lsid:biocol.org:col:34992" box="[567,657,397,422]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="358">RMNH</collectionCode>
.
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.D.57224.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFE81114EBA8FE75FDE66324" pageId="17" pageNumber="358" type="description">
<paragraph id="7107F648FFE81114EBA8FE75FDDF60DB" blockId="17.[151,1437,325,1394]" box="[199,558,433,458]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBA8FE75FEA060D8" bold="true" box="[199,337,433,458]" pageId="17" pageNumber="358">Description</emphasis>
. See
<bibRefCitation id="15298BB9FFE81114EAE0FE75FDD860DB" author="Anker" box="[399,553,433,458]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker (2014)</bibRefCitation>
.
</paragraph>
<paragraph id="7107F648FFE81114EBA8FE11FDE66324" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBA8FE11FE5C60FC" bold="true" box="[199,429,469,494]" pageId="17" pageNumber="358">Colouration in life</emphasis>
. Body and appendages whitish with pale orange-yellow tinge and some iridescence, especially on chelipeds and pleon (see
<bibRefCitation id="15298BB9FFE81114E921FE3DFD266303" author="Anker" box="[590,727,505,530]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
, fig. 5); eggs in ovigerous females bright orange-red (based on low-resolution field photographs).
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFE81114EBA8FD85FECD636C" pageId="17" pageNumber="358" type="distribution">
<paragraph id="7107F648FFE81114EBA8FD85FECD636C" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBA8FD85FEA86348" bold="true" box="[199,345,577,602]" pageId="17" pageNumber="358">Distribution</emphasis>
. Western Atlantic:
<collectingCountry id="09AFB6D8FFE81114E955FD85FD306348" box="[570,705,577,602]" name="Guadeloupe" pageId="17" pageNumber="358">Guadeloupe</collectingCountry>
(French Antilles) and
<collectingCountry id="09AFB6D8FFE81114E8A6FD85FBF56348" box="[969,1028,577,602]" name="Cuba" pageId="17" pageNumber="358">Cuba</collectingCountry>
(
<collectingCountry id="09AFB6D8FFE81114EF79FD85FBB16348" box="[1046,1088,577,602]" name="Iceland" pageId="17" pageNumber="358">Isla</collectingCountry>
de la Juventud) (
<bibRefCitation id="15298BB9FFE81114EE64FD85FEC1636C" author="Anker" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014; present study</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFE81114EBA8FD4DFD3E6460" pageId="17" pageNumber="358" type="discussion">
<paragraph id="7107F648FFE81114EBA8FD4DFC726220" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<emphasis id="43CC2A5AFFE81114EBA8FD4DFEC563B0" bold="true" box="[199,308,649,674]" pageId="17" pageNumber="358">Remarks</emphasis>
.
<taxonomicName id="B6B88DCBFFE81114EA27FD4DFDC063B3" box="[328,561,649,673]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114EA27FD4DFDC063B3" box="[328,561,649,673]" italics="true" pageId="17" pageNumber="358">Naushonia draconis</emphasis>
</taxonomicName>
was previously known from a single specimen, the male
<typeStatus id="AE0348EAFFE81114EF99FD4DFAA963B0" box="[1270,1368,649,674]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
from
<collectingCountry id="09AFB6D8FFE81114EBF8FD69FED063D4" box="[151,289,685,710]" name="Guadeloupe" pageId="17" pageNumber="358">Guadeloupe</collectingCountry>
, French Antilles (
<bibRefCitation id="15298BB9FFE81114EA85FD69FD8663D4" author="Anker" box="[490,631,685,710]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
). The Cuban material agrees almost perfectly with the
<typeStatus id="AE0348EAFFE81114EF8DFD69FAB663D4" box="[1250,1351,685,710]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
, except for some minor details, which all appear to be within an expected intraspecific variation in species of
<taxonomicName id="B6B88DCBFFE81114EE73FD15FA6463FB" box="[1308,1429,721,745]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFE81114EE73FD15FA6463FB" box="[1308,1429,721,745]" italics="true" pageId="17" pageNumber="358">Naushonia</emphasis>
</taxonomicName>
. This variation includes the shape and armature of the mid-dorsal crests on the pleon (
<figureCitation id="E983EACDFFE81114EF34FD31FB6E621C" box="[1115,1183,757,782]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
A, I), ventral armature of the pleomeres (
<figureCitation id="E983EACDFFE81114EA0EFCDDFE526220" box="[353,419,793,818]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
B) and dentition on the chelipeds (
<figureCitation id="E983EACDFFE81114E849FCDDFC986220" box="[806,873,793,818]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
J).
</paragraph>
<paragraph id="7107F648FFE81114EBA8FCF9FECC65AC" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
<bibRefCitation id="15298BB9FFE81114EBA8FCF9FEAE6244" author="Anker" box="[199,351,829,854]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker (2014)</bibRefCitation>
dissected and illustrated the third and second maxillipeds of the
<typeStatus id="AE0348EAFFE81114EF5AFCF9FB666244" box="[1077,1175,829,854]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B6B88DCBFFE81114EFD3FCFAFACE6247" box="[1212,1343,829,853]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114EFD3FCFAFACE6247" box="[1212,1343,829,853]" italics="true" pageId="17" pageNumber="358">N. draconis</emphasis>
</taxonomicName>
, but not the rest of the mouthparts in order not to further damage the unique specimen at that time. The new material enables us to illustrate the previously non-figured mandible, maxillule, maxilla and first maxilliped (
<figureCitation id="E983EACDFFE81114EE7CFC41FAA6628C" box="[1299,1367,901,926]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
CG), which are all fairly typical for
<taxonomicName id="B6B88DCBFFE81114EA83FC6DFD9462D3" box="[492,613,937,961]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFE81114EA83FC6DFD9462D3" box="[492,613,937,961]" italics="true" pageId="17" pageNumber="358">Naushonia</emphasis>
</taxonomicName>
. The posterolateral angles of the sixth pleomere are indeed produced into strong sharp spines, i.e. one on each side of the telson; both spines were broken in the
<typeStatus id="AE0348EAFFE81114EF00FC09FB2062F4" box="[1135,1233,973,998]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
(see
<bibRefCitation id="15298BB9FFE81114EE61FC09FA6962F4" author="Anker" box="[1294,1432,973,998]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
: fig. 2E, F). An interesting feature not specified in the original description of
<taxonomicName id="B6B88DCBFFE81114E890FC36FB72651B" box="[1023,1155,1009,1033]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114E890FC36FB72651B" box="[1023,1155,1009,1033]" italics="true" pageId="17" pageNumber="358">N. draconis</emphasis>
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is the presence a strong, sharp, posteriorly directed, median tooth on the sternum of the sixth pleomere; this tooth is present in both males and females and can be seen in the ventral view of the
<typeStatus id="AE0348EAFFE81114E870FBFDFC706540" box="[799,897,1081,1106]" pageId="17" pageNumber="358" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="15298BB9FFE81114E8FCFBFDFB716540" author="Anker" box="[915,1152,1081,1106]" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014: fig. 1B</bibRefCitation>
). The mesial side of the cheliped coxa is armed with a very stout, anteriorly directed, sharp spine; although not specifically mentioned in the original description, this spine can be seen even in the lateral view of the basal articles of the cheliped (
<bibRefCitation id="15298BB9FFE81114EE3BFB45FEC165AC" author="Anker" pageId="17" pageNumber="358" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014: fig. 3K</bibRefCitation>
).
</paragraph>
<paragraph id="7107F648FFE81114EBA8FB0DFD3E6460" blockId="17.[151,1437,325,1394]" pageId="17" pageNumber="358">
The previously incompletely known gill formula of
<taxonomicName id="B6B88DCBFFE81114E87EFB0EFC6565F3" box="[785,916,1225,1249]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="358" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFE81114E87EFB0EFC6565F3" box="[785,916,1225,1249]" italics="true" pageId="17" pageNumber="358">N. draconis</emphasis>
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was examined in a male specimen from
<collectingCountry id="09AFB6D8FFE81114EE0EFB0DFA6D65F0" box="[1377,1436,1225,1250]" name="Cuba" pageId="17" pageNumber="358">Cuba</collectingCountry>
(MZUSP 33201) and generally conforms to that of the other five species of the genus reported herein (see Table I). The arthrobranch at the base of the first maxilliped is well developed, with multiple lamellae (
<figureCitation id="E983EACDFFE81114EFDFFAD5FB026438" box="[1200,1267,1297,1322]" captionStart="FIGURE 9" captionStartId="16.[151,250,1782,1804]" captionTargetBox="[229,1346,222,1722]" captionTargetId="figure@16.[206,1365,195,1748]" captionTargetPageId="16" captionText="FIGURE 9. Naushonia draconis Anker, 2014, male (A H; pocl 5.1 mm) and female (I, J; pocl 5.6 mm) from Isla de la Juventud, Cuba (MZUSP 33201), A, pleon, mid-dorsal crests of first to sixth pleomeres, lateral view; B, same, ventral margins of pleomeres, lateral view; C, right mandible, lateral view; D, same, mesial view; E, right maxillule, lateral view; F, right maxilla, lateral view; G, right first maxilliped, lateral view; H, epipod complex of right third pereopod, lateral view; I, pleon, mid-dorsal crests of first to fourth pleomeres, lateral view; J, left cheliped, chela and distal portion of carpus, ventral view. Scale bars: 1 mm for A, B, E-G, I, J; 0.5 mm for C, D." httpUri="https://zenodo.org/record/231789/files/figure.png" pageId="17" pageNumber="358">Fig. 9</figureCitation>
G). The epipod of the fourth pereopod is reduced to a simple lobe; there is apparently no setobranch on the fourth pereopod, corresponding to the epipod on the third pereopod.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,279 @@
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<emphasis id="43CC2A5AFFF41108EBF8F998FE4C6764" bold="true" box="[151,445,1628,1654]" italics="true" pageId="13" pageNumber="354">Naushonia serratipalma</emphasis>
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,
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, figs. 1C, 57 (
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locality: Moorea, Society Islands,
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).—
<treatmentCitation id="F019D059FFF41108EBA8F921FE8367E9" author="Anker" box="[199,370,1765,1787]" page="760" pageId="13" pageNumber="354" year="2014">
<bibRefCitation id="15298BB9FFF41108EBA8F921FEB267E9" author="Anker" box="[199,323,1765,1787]" pageId="13" pageNumber="354" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
: 760
</treatmentCitation>
(key), 767, fig. 6 (Moorea).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFF41108EBF8F8E3FCAF66C3" pageId="13" pageNumber="354" type="materials_examined">
<paragraph id="7107F648FFF41108EBF8F8E3FCAF66C3" blockId="13.[151,1436,1831,2037]" pageId="13" pageNumber="354">
<emphasis id="43CC2A5AFFF41108EBF8F8E3FE8B6652" bold="true" box="[151,378,1831,1856]" pageId="13" pageNumber="354">Material examined</emphasis>
.
<collectingCountry id="09AFB6D8FFF41108EAE5F8E3FDD26652" box="[394,547,1831,1856]" name="Saudi Arabia" pageId="13" pageNumber="354">Saudi Arabia</collectingCountry>
:
<specimenCount id="67BE3DC1FFF41108E941F8ECFD616652" box="[558,656,1831,1856]" pageId="13" pageNumber="354" type="female">1 female</specimenCount>
(pocl 3.0 mm), Red Sea coast, Al Lith, sta. SAFA-039,
<geoCoordinate id="148C908FFFF41108EE7DF8E3FA666652" box="[1298,1431,1831,1856]" direction="north" orientation="latitude" pageId="13" pageNumber="354" precision="9" value="20.278667">20°16.72N</geoCoordinate>
,
<geoCoordinate id="148C908FFFF41108EBF8F888FEEB6676" box="[151,282,1868,1892]" direction="east" orientation="longitude" pageId="13" pageNumber="354" precision="9" value="40.372166">40°22.33E</geoCoordinate>
, silt-sand flat with coral rubble, near small young mangrove,
<quantity id="B6405BADFFF41108E8B1F888FBD86676" box="[990,1065,1868,1892]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="354" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">12 m</quantity>
, coll. A. Anker
<emphasis id="43CC2A5AFFF41108EF8BF889FAEF6676" box="[1252,1310,1868,1892]" italics="true" pageId="13" pageNumber="354">et al.</emphasis>
,
<date id="0506D088FFF41108EE41F888FF23669A" pageId="13" pageNumber="354" value="2013-03-21">21 March 2013</date>
, BDJRS-3079,
<collectionCode id="17A96E8DFFF41108EAE9F8B4FE196695" box="[390,488,1904,1927]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="13" pageNumber="354">FLMNH</collectionCode>
UF Crustacea 35970.
<collectingCountry id="09AFB6D8FFF41108E981F8ABFC46669A" box="[750,951,1903,1928]" name="French Polynesia" pageId="13" pageNumber="354">French Polynesia</collectingCountry>
:
<specimenCount id="67BE3DC1FFF41108E8ABF8B4FBD9669A" box="[964,1064,1903,1928]" pageId="13" pageNumber="354" type="female">1 female</specimenCount>
(pocl
<quantity id="B6405BADFFF41108EF1AF8B4FB23669A" box="[1141,1234,1904,1928]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="13" pageNumber="354" unit="mm" value="4.4">4.4 mm</quantity>
), Society Islands, Moorea Island, near Papetoai,
<quantity id="B6405BADFFF41108E96EF850FDBE66BE" box="[513,591,1940,1964]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="13" pageNumber="354" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">01 m</quantity>
, sand flat with mounds, coral rubble and living corals, under coral,
<date id="0506D088FFF41108EEEFF850FEBE66C2" pageId="13" pageNumber="354" value="2008-11-23">23 November 2008</date>
, coll. A. Anker,
<collectionCode id="17A96E8DFFF41108E968F87DFD9866C2" box="[519,617,1977,2000]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="13" pageNumber="354">FLMNH</collectionCode>
UF Crustacea 16537.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFF4110BEBA8F818FB466617" lastPageId="14" lastPageNumber="355" pageId="13" pageNumber="354" type="description">
<paragraph id="7107F648FFF41108EBA8F818FD6F66E6" blockId="13.[151,1436,1831,2037]" box="[199,670,2012,2037]" pageId="13" pageNumber="354">
<emphasis id="43CC2A5AFFF41108EBA8F818FEA066E7" bold="true" box="[199,337,2012,2037]" pageId="13" pageNumber="354">Description</emphasis>
. See
<bibRefCitation id="15298BB9FFF41108EAE0F818FD6B66E6" author="Komai" box="[399,666,2012,2037]" pageId="13" pageNumber="354" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504, 31 - 46." type="journal article" year="2010">Komai &amp; Anker (2010)</bibRefCitation>
.
</paragraph>
<caption id="25C7A6C0FFF7110BEBF8F9B6FAAC67D7" httpUri="https://zenodo.org/record/231787/files/figure.png" pageId="14" pageNumber="355" targetBox="[169,1408,185,1628]" targetPageId="14">
<paragraph id="7107F648FFF7110BEBF8F9B6FAAC67D7" blockId="14.[151,1436,1650,1733]" pageId="14" pageNumber="355">
<emphasis id="43CC2A5AFFF7110BEBF8F9B6FEE16795" bold="true" box="[151,272,1650,1672]" pageId="14" pageNumber="355">FIGURE 7</emphasis>
.
<taxonomicName id="B6B88DCBFFF7110BEA70F9B6FCE3679A" authority="Komai &amp; Anker, 2010" authorityName="Komai &amp; Anker" authorityYear="2010" box="[287,786,1650,1672]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="43CC2A5AFFF7110BEA70F9B6FDEB679A" box="[287,538,1650,1672]" italics="true" pageId="14" pageNumber="355">Naushonia serratipalma</emphasis>
Komai &amp; Anker, 2010
</taxonomicName>
, female (pocl 3.0 mm) from Al Lith, Red Sea coast of Saudi Arabia, FLMNH UF Crustacea 35970. A, anterior part of carapace and cephalic appendages, dorsal view; B, right cheliped, dorsal view (damaged on mesial side); C, same, ventral view; D, telson, dorsal view (setae omitted). Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="7107F648FFF7110BEBA8F928FB466617" blockId="14.[151,1436,1772,2031]" box="[199,1207,1772,1797]" pageId="14" pageNumber="355">
<emphasis id="43CC2A5AFFF7110BEBA8F928FE536617" bold="true" box="[199,418,1772,1797]" pageId="14" pageNumber="355">Colouration in life</emphasis>
. Body and appendages whitish with pale orange-yellow tinge (
<figureCitation id="E983EACDFFF7110BEF0AF928FB586617" box="[1125,1193,1772,1797]" captionStart="FIGURE 8" captionStartId="15.[151,250,1287,1309]" captionTargetBox="[242,1344,176,1265]" captionTargetId="figure@15.[242,1345,169,1265]" captionTargetPageId="15" captionText="FIGURE 8. Naushonia serratipalma Komai &amp; Anker, 2010, female (pocl 3.0 mm) from Al Lith, Red Sea coast of Saudi Arabia, FLMNH UF Crustacea 35970, living individual in dorsal and lateral views." httpUri="https://zenodo.org/record/231788/files/figure.png" pageId="14" pageNumber="355">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFF7110BEBA8F8CAFC9B665A" pageId="14" pageNumber="355" type="distribution">
<paragraph id="7107F648FFF7110BEBA8F8CAFC9B665A" blockId="14.[151,1436,1772,2031]" pageId="14" pageNumber="355">
<emphasis id="43CC2A5AFFF7110BEBA8F8CAFEA86635" bold="true" box="[199,345,1806,1831]" pageId="14" pageNumber="355">Distribution</emphasis>
. Indo-West Pacific:
<collectingCountry id="09AFB6D8FFF7110BE923F8CAFCE36635" box="[588,786,1806,1831]" name="French Polynesia" pageId="14" pageNumber="355">French Polynesia</collectingCountry>
(Society Islands) (
<bibRefCitation id="15298BB9FFF7110BE89CF8CAFB0A6635" author="Komai" box="[1011,1275,1806,1831]" pageId="14" pageNumber="355" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504, 31 - 46." type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
;
<bibRefCitation id="15298BB9FFF7110BEE64F8CAFA666635" author="Anker" box="[1291,1431,1806,1831]" pageId="14" pageNumber="355" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
; present study) and
<collectingCountry id="09AFB6D8FFF7110BEA04F8EBFE0F665A" box="[363,510,1839,1864]" name="Saudi Arabia" pageId="14" pageNumber="355">Saudi Arabia</collectingCountry>
(Red Sea coast) (present study).
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFF7110AEBA8F894FB2B66A8" lastPageId="15" lastPageNumber="356" pageId="14" pageNumber="355" type="discussion">
<paragraph id="7107F648FFF7110BEBA8F894FC8266FD" blockId="14.[151,1436,1772,2031]" pageId="14" pageNumber="355">
<emphasis id="43CC2A5AFFF7110BEBA8F894FEC5667B" bold="true" box="[199,308,1872,1897]" pageId="14" pageNumber="355">Remarks</emphasis>
.
<taxonomicName id="B6B88DCBFFF7110BEA2DF895FDA2667B" box="[322,595,1873,1897]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="43CC2A5AFFF7110BEA2DF895FDA2667B" box="[322,595,1873,1897]" italics="true" pageId="14" pageNumber="355">Naushonia serratipalma</emphasis>
</taxonomicName>
was originally described based on a female
<typeStatus id="AE0348EAFFF7110BEF2CF894FB54667B" box="[1091,1189,1872,1897]" pageId="14" pageNumber="355" type="holotype">holotype</typeStatus>
from Moorea,
<collectingCountry id="09AFB6D8FFF7110BEE21F894FEF56699" name="French Polynesia" pageId="14" pageNumber="355">French Polynesia</collectingCountry>
(
<bibRefCitation id="15298BB9FFF7110BEA7CF8B6FDFE6699" author="Komai" box="[275,527,1906,1931]" pageId="14" pageNumber="355" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504, 31 - 46." type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
), with a subsequent record of another female specimen from the same locality in
<bibRefCitation id="15298BB9FFF7110BEBF8F857FEC766BE" author="Anker" box="[151,310,1939,1964]" pageId="14" pageNumber="355" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker (2014)</bibRefCitation>
. The Saudi Arabian specimen, if confirmed to be conspecific with
<taxonomicName id="B6B88DCBFFF7110BEF20F850FAF566BE" box="[1103,1284,1940,1964]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="355" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="43CC2A5AFFF7110BEF20F850FAF566BE" box="[1103,1284,1940,1964]" italics="true" pageId="14" pageNumber="355">N. serratipalma</emphasis>
</taxonomicName>
(see below), would represent a significant range extension of the species, with the geographic range possibly spanning the entire tropical Indo-West Pacific from the Red Sea to
<collectingCountry id="09AFB6D8FFF7110BE9C5F812FC8166FD" box="[682,880,2006,2031]" name="French Polynesia" pageId="14" pageNumber="355">French Polynesia</collectingCountry>
.
</paragraph>
<caption id="25C7A6C0FFF6110AEBF8FAC3FC126429" httpUri="https://zenodo.org/record/231788/files/figure.png" pageId="15" pageNumber="356" targetBox="[242,1344,176,1265]" targetPageId="15">
<paragraph id="7107F648FFF6110AEBF8FAC3FC126429" blockId="15.[151,1436,1287,1339]" pageId="15" pageNumber="356">
<emphasis id="43CC2A5AFFF6110AEBF8FAC3FEE1640E" bold="true" box="[151,272,1287,1309]" pageId="15" pageNumber="356">FIGURE 8</emphasis>
.
<taxonomicName id="B6B88DCBFFF6110AEA70FAC3FCE3640F" authority="Komai &amp; Anker, 2010" authorityName="Komai &amp; Anker" authorityYear="2010" box="[287,786,1287,1309]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="43CC2A5AFFF6110AEA70FAC3FDEB640F" box="[287,538,1287,1309]" italics="true" pageId="15" pageNumber="356">Naushonia serratipalma</emphasis>
Komai &amp; Anker, 2010
</taxonomicName>
, female (pocl 3.0 mm) from Al Lith, Red Sea coast of Saudi Arabia, FLMNH UF Crustacea 35970, living individual in dorsal and lateral views.
</paragraph>
</caption>
<paragraph id="7107F648FFF6110AEBA8FA49FBAA6759" blockId="15.[151,1437,1420,1978]" pageId="15" pageNumber="356">
Among the species characterized by the non-carinate pleomeres and marginally unarmed pleura,
<taxonomicName id="B6B88DCBFFF6110AEEEDFA49FED964D7" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="43CC2A5AFFF6110AEEEDFA49FED964D7" italics="true" pageId="15" pageNumber="356">N. serratipalma</emphasis>
</taxonomicName>
is morphologically closest to
<taxonomicName id="B6B88DCBFFF6110AE911FA6AFCF564D7" box="[638,772,1453,1477]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="latimana">
<emphasis id="43CC2A5AFFF6110AE911FA6AFCF564D7" box="[638,772,1453,1477]" italics="true" pageId="15" pageNumber="356">N. latimana</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFF6110AE87DFA6AFC4364D7" box="[786,946,1454,1477]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="macginitiei">
<emphasis id="43CC2A5AFFF6110AE87DFA6AFC4364D7" box="[786,946,1454,1477]" italics="true" pageId="15" pageNumber="356">N. macginitiei</emphasis>
</taxonomicName>
and
<taxonomicName id="B6B88DCBFFF6110AE884FA6AFB9564D7" box="[1003,1124,1454,1477]" class="Malacostraca" family="Laomediidae" genus="Coralliocrangon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="43CC2A5AFFF6110AE884FA6AFB9564D7" box="[1003,1124,1454,1477]" italics="true" pageId="15" pageNumber="356">N. perrieri</emphasis>
</taxonomicName>
. The four species share the presence of less than 10 teeth on the lateral margin of the antennal scaphocerite, with the terminal tooth pointing straight-forwards, not incurved. However,
<taxonomicName id="B6B88DCBFFF6110AE91BFA35FCD7671A" box="[628,806,1520,1544]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="43CC2A5AFFF6110AE91BFA35FCD7671A" box="[628,806,1520,1544]" italics="true" pageId="15" pageNumber="356">N. serratipalma</emphasis>
</taxonomicName>
can be distinguished from the other three species by the distinctly serrated proximal part of the occlusal margin of the cheliped palm vs. non-serrated in
<taxonomicName id="B6B88DCBFFF6110AEF84F9D6FA806738" box="[1259,1393,1554,1578]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="latimana">
<emphasis id="43CC2A5AFFF6110AEF84F9D6FA806738" box="[1259,1393,1554,1578]" italics="true" pageId="15" pageNumber="356">N. latimana</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFF6110AEEEDF9D6FEE46759" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="macginitiei">
<emphasis id="43CC2A5AFFF6110AEEEDF9D6FEE46759" italics="true" pageId="15" pageNumber="356">N. macginitiei</emphasis>
</taxonomicName>
and
<taxonomicName id="B6B88DCBFFF6110AEA24F9F0FE326759" box="[331,451,1588,1611]" class="Malacostraca" family="Laomediidae" genus="Coralliocrangon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="43CC2A5AFFF6110AEA24F9F0FE326759" box="[331,451,1588,1611]" italics="true" pageId="15" pageNumber="356">N. perrieri</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="15298BB9FFF6110AEA98F9F7FD626759" author="Glassell" box="[503,659,1587,1612]" pageId="15" pageNumber="356" refString="Glassell, S. A. (1938) New and obscure decapod Crustacea from the West American coasts. Transactions of the San Diego Society of Natural History, 3, 411 - 454." type="journal article" year="1938">Glassell 1938</bibRefCitation>
;
<bibRefCitation id="15298BB9FFF6110AE9F0F9F0FCBB675E" author="Ngoc-Ho" box="[671,842,1587,1612]" pageId="15" pageNumber="356" refString="Ngoc-Ho, N. (1996) Redescription des types de Naushonia perrieri (Nobili, 1904) (Crustacea, Decapoda, Laomediidae). Bulletin du Museum National d'Histoire Naturelle, 18, 565 - 70." type="journal article" year="1996">Ngoc-Ho 1996</bibRefCitation>
;
<bibRefCitation id="15298BB9FFF6110AE83AF9F7FBBE6759" author="Komai" box="[853,1103,1587,1612]" pageId="15" pageNumber="356" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504, 31 - 46." type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
).
</paragraph>
<paragraph id="7107F648FFF6110AEBA8F990FB2B66A8" blockId="15.[151,1437,1420,1978]" pageId="15" pageNumber="356">
The topotypic specimen from Moorea generally agrees with the
<typeStatus id="AE0348EAFFF6110AE8FAF990FC0B677F" box="[917,1018,1620,1645]" pageId="15" pageNumber="356" type="holotype">holotype</typeStatus>
, differing only in the slightly broader rostrum and weaker sculpture on the pleonal tergites (cf.
<bibRefCitation id="15298BB9FFF6110AE84EF9B1FBEF679C" author="Komai" box="[801,1054,1653,1678]" pageId="15" pageNumber="356" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504, 31 - 46." type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
). The apparently young specimen (cl 3.0 mm) from
<collectingCountry id="09AFB6D8FFF6110AEA08F952FE0D67BD" box="[359,508,1686,1711]" name="Saudi Arabia" pageId="15" pageNumber="356">Saudi Arabia</collectingCountry>
differs from the French Polynesian specimens in the relatively narrower rostrum with stronger marginal teeth (
<figureCitation id="E983EACDFFF6110AEAB0F97CFDD367C2" box="[479,546,1720,1745]" captionStart="FIGURE 7" captionStartId="14.[151,250,1650,1672]" captionTargetBox="[169,1408,185,1628]" captionTargetId="figure@14.[151,1436,161,1658]" captionTargetPageId="14" captionText="FIGURE 7. Naushonia serratipalma Komai &amp; Anker, 2010, female (pocl 3.0 mm) from Al Lith, Red Sea coast of Saudi Arabia, FLMNH UF Crustacea 35970. A, anterior part of carapace and cephalic appendages, dorsal view; B, right cheliped, dorsal view (damaged on mesial side); C, same, ventral view; D, telson, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231787/files/figure.png" pageId="15" pageNumber="356">Fig. 7</figureCitation>
A); the possession of two or three minute denticles, instead of only one denticle, on the lateral margin of the telson (
<figureCitation id="E983EACDFFF6110AE94DF91DFD9767E0" box="[546,614,1753,1778]" captionStart="FIGURE 7" captionStartId="14.[151,250,1650,1672]" captionTargetBox="[169,1408,185,1628]" captionTargetId="figure@14.[151,1436,161,1658]" captionTargetPageId="14" captionText="FIGURE 7. Naushonia serratipalma Komai &amp; Anker, 2010, female (pocl 3.0 mm) from Al Lith, Red Sea coast of Saudi Arabia, FLMNH UF Crustacea 35970. A, anterior part of carapace and cephalic appendages, dorsal view; B, right cheliped, dorsal view (damaged on mesial side); C, same, ventral view; D, telson, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231787/files/figure.png" pageId="15" pageNumber="356">Fig. 7</figureCitation>
B); having no trace of sculpture on the pleonal tergites; and the relatively more slender cheliped palm (cf.
<figureCitation id="E983EACDFFF6110AE969F93FFDB76601" box="[518,582,1787,1812]" captionStart="FIGURE 8" captionStartId="15.[151,250,1287,1309]" captionTargetBox="[242,1344,176,1265]" captionTargetId="figure@15.[242,1345,169,1265]" captionTargetPageId="15" captionText="FIGURE 8. Naushonia serratipalma Komai &amp; Anker, 2010, female (pocl 3.0 mm) from Al Lith, Red Sea coast of Saudi Arabia, FLMNH UF Crustacea 35970, living individual in dorsal and lateral views." httpUri="https://zenodo.org/record/231788/files/figure.png" pageId="15" pageNumber="356">Fig. 8</figureCitation>
A and
<bibRefCitation id="15298BB9FFF6110AE9FBF93FFC616601" author="Komai" box="[660,912,1787,1812]" pageId="15" pageNumber="356" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504, 31 - 46." type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
: fig. 1C). In all other diagnostic features, such as the proximal part of the occlusal margin of the cheliped palm serrated with small teeth (
<figureCitation id="E983EACDFFF6110AEFEDF8D8FB326627" box="[1154,1219,1820,1845]" captionStart="FIGURE 7" captionStartId="14.[151,250,1650,1672]" captionTargetBox="[169,1408,185,1628]" captionTargetId="figure@14.[151,1436,161,1658]" captionTargetPageId="14" captionText="FIGURE 7. Naushonia serratipalma Komai &amp; Anker, 2010, female (pocl 3.0 mm) from Al Lith, Red Sea coast of Saudi Arabia, FLMNH UF Crustacea 35970. A, anterior part of carapace and cephalic appendages, dorsal view; B, right cheliped, dorsal view (damaged on mesial side); C, same, ventral view; D, telson, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231787/files/figure.png" pageId="15" pageNumber="356">Fig. 7</figureCitation>
C) and the antennal scaphocerite bearing five or six teeth on the lateral margin, including one straight terminal tooth (
<figureCitation id="E983EACDFFF6110AEE69F8F9FABA6644" box="[1286,1355,1853,1878]" captionStart="FIGURE 7" captionStartId="14.[151,250,1650,1672]" captionTargetBox="[169,1408,185,1628]" captionTargetId="figure@14.[151,1436,161,1658]" captionTargetPageId="14" captionText="FIGURE 7. Naushonia serratipalma Komai &amp; Anker, 2010, female (pocl 3.0 mm) from Al Lith, Red Sea coast of Saudi Arabia, FLMNH UF Crustacea 35970. A, anterior part of carapace and cephalic appendages, dorsal view; B, right cheliped, dorsal view (damaged on mesial side); C, same, ventral view; D, telson, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231787/files/figure.png" pageId="15" pageNumber="356">Fig. 7</figureCitation>
A), the Saudi Arabian specimen agrees well with
<taxonomicName id="B6B88DCBFFF6110AE904F8A4FCED6665" box="[619,796,1887,1911]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="43CC2A5AFFF6110AE904F8A4FCED6665" box="[619,796,1887,1911]" italics="true" pageId="15" pageNumber="356">N. serratipalma</emphasis>
</taxonomicName>
. Similar intraspecific variation in the shape and armature of the rostrum is known in
<taxonomicName id="B6B88DCBFFF6110AEA8FF845FC7C668B" authority="Dworschak et al. 2006" authorityName="Dworschak et al." authorityYear="2006" box="[480,909,1920,1945]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFF6110AEA8FF845FD95668A" box="[480,612,1921,1944]" italics="true" pageId="15" pageNumber="356">N. carinata</emphasis>
(
<bibRefCitation id="15298BB9FFF6110AE918F844FC74668B" author="Dworschak" box="[631,901,1920,1945]" pageId="15" pageNumber="356" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak
<emphasis id="43CC2A5AFFF6110AE86BF845FCB1668A" box="[772,832,1920,1944]" italics="true" pageId="15" pageNumber="356">et al.</emphasis>
2006
</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="B6B88DCBFFF6110AE8A3F845FB81668A" box="[972,1136,1920,1944]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFF6110AE8A3F845FB81668A" box="[972,1136,1920,1944]" italics="true" pageId="15" pageNumber="356">N. lactoalbida</emphasis>
</taxonomicName>
(see above), whereas the number of denticles on the lateral margin of the telson is also variable in
<taxonomicName id="B6B88DCBFFF6110AE8A7F866FBBA66AB" box="[968,1099,1954,1977]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="356" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFF6110AE8A7F866FBBA66AB" box="[968,1099,1954,1977]" italics="true" pageId="15" pageNumber="356">N. japonica</emphasis>
</taxonomicName>
(see above).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,309 @@
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<treatment id="F911475EFFF81106EBF8FA9CFBC161C7" ID-DOI="http://doi.org/10.5281/zenodo.5073329" ID-GBIF-Taxon="119631468" ID-Zenodo-Dep="5073329" LSID="urn:lsid:plazi:treatment:F911475EFFF81106EBF8FA9CFBC161C7" httpUri="http://treatment.plazi.org/id/F911475EFFF81106EBF8FA9CFBC161C7" lastPageId="3" lastPageNumber="343" pageId="1" pageNumber="342">
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<paragraph id="7107F648FFF81104EBF8FA9CFDC16460" blockId="1.[151,560,1367,1394]" box="[151,560,1367,1394]" pageId="1" pageNumber="342">
<heading id="2A4F4124FFF81104EBF8FA9CFDC16460" bold="true" box="[151,560,1367,1394]" fontSize="11" level="1" pageId="1" pageNumber="342" reason="1">
<emphasis id="43CC2A5AFFF81104EBF8FA9CFDC16460" bold="true" box="[151,560,1367,1394]" pageId="1" pageNumber="342">
Genus
<taxonomicName id="B6B88DCBFFF81104EB81FA93FDC16460" ID-CoL="b0d1bf87-999b-436b-86a4-3e91990de14e" authority="Kingsley, 1897" authorityName="Kingsley" authorityYear="1897" box="[238,560,1367,1394]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="342" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFF81104EB81FA93FE856463" bold="true" box="[238,372,1367,1393]" italics="true" pageId="1" pageNumber="342">Naushonia</emphasis>
<bibRefCitation id="15298BB9FFF81104EA14FA9CFDC16460" author="Kingsley" box="[379,560,1368,1394]" pageId="1" pageNumber="342" refString="Kingsley, J. S. (1897) On a new genus and two new species of macrurous Crustacea. Bulletin of the Essex Institute, 27, 95 - 99." type="journal article" year="1897">Kingsley, 1897</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="7107F648FFF81107EBF8FA53FBBD61C7" blockId="1.[151,1437,1431,2033]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="343" pageId="1" pageNumber="342">
<emphasis id="43CC2A5AFFF81104EBF8FA53FE7064A2" bold="true" box="[151,385,1431,1456]" pageId="1" pageNumber="342">Emended diagnosis</emphasis>
. Laomediid mud shrimps with firm, well-calcified integument. Carapace with well-marked linea thalassinica, running entire or almost entire length of carapace; rostrum well developed, rounded or pointed distally, with variable armature; postorbital spine simple; branchiostegal spine usually strong; carapace armature extremely variable, ranging from low carinae with small granules to prominent crests and rows of strong spines. Pleon with first somite short, second to sixth somites subequal in length; dorsolateral surface of pleomeres typically smooth or with faint carinae, sometimes with stronger armature consisting of crests, spines and tubercles; pleural margins entire or variously toothed. Telson with posterior margin convex, with or without spines on lateral margin; dorsal surface usually armed with spines or tubercles. Eye relatively small, cornea with normal or reduced pigmentation. Antennular peduncle with second and third segments not particularly elongate. Antennal peduncle with well-developed, laterally toothed scaphocerite. Mandible with sharp teeth along entire distal edge of incisor process; palp with three articles. First maxilliped with large endopod and well-developed epipod. Second maxilliped with scaphognathite furnished with long posterior setae. Third maxilliped pediform, with welldeveloped crista dentata on ischium; exopod well developed. First pereopods subchelate, symmetrical, carried with dactylus in distolateral position relative to dorsoventrally flattened palm; occlusal (= cutting) margin of palm more or less denticulate, often with strong teeth. Second pereopod non-chelate; dactylus with dense setal brush on dorsal to lateral surfaces. Dactyli of third to fifth pereopods each with thin, comb-like, toothed lamina on flexor margin. First pleopods absent in male, uniramous in female; second to fifth pleopods biramous, without appendices internae, male second pleopod without appendix masculina. Uropods with complete, finely toothed diaeresis on both exopod and endopod. For branchial formula, see
<tableCitation id="3C3AC3F3FFFB1107E99AFF78FCB961C6" box="[757,840,188,213]" captionStart="TABLE 1" captionStartId="2.[151,239,254,277]" captionText="TABLE 1. Gill and exopod formula of the six species of Naushonia treated in this study. [r—reduced; * setobranchs present on the fourth pereopod in N. carinata, N. japonica, N. lactoalbida, N. portoricensis, N. serratipalma, apparently absent in N. draconis]." pageId="2" pageNumber="343">Table 1</tableCitation>
and “Remarks” below.
</paragraph>
<caption id="25C7A6C0FFFB1107EBF8FF3AFE786045" pageId="2" pageNumber="343">
<paragraph id="7107F648FFFB1107EBF8FF3AFE786045" blockId="2.[151,1436,253,343]" pageId="2" pageNumber="343">
<emphasis id="43CC2A5AFFFB1107EBF8FF3AFEF46007" bold="true" box="[151,261,254,277]" pageId="2" pageNumber="343">TABLE 1</emphasis>
. Gill and exopod formula of the six species of
<taxonomicName id="B6B88DCBFFFB1107E87CFF39FC776007" box="[787,902,253,277]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFB1107E87CFF39FC776007" box="[787,902,253,277]" italics="true" pageId="2" pageNumber="343">Naushonia</emphasis>
</taxonomicName>
treated in this study. [r—reduced; * setobranchs present on the fourth pereopod in
<taxonomicName id="B6B88DCBFFFB1107E96DFEDAFD8D6027" box="[514,636,286,309]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFB1107E96DFEDAFD8D6027" box="[514,636,286,309]" italics="true" pageId="2" pageNumber="343">N. carinata</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107E9E5FEDAFCF76027" box="[650,774,286,309]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFB1107E9E5FEDAFCF76027" box="[650,774,286,309]" italics="true" pageId="2" pageNumber="343">N. japonica</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107E87BFEDAFC5C6027" box="[788,941,285,309]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFFB1107E87BFEDAFC5C6027" box="[788,941,285,309]" italics="true" pageId="2" pageNumber="343">N. lactoalbida</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107E8D4FEDAFB976027" box="[955,1126,286,309]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFFB1107E8D4FEDAFB976027" box="[955,1126,286,309]" italics="true" pageId="2" pageNumber="343">N. portoricensis</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107EF1BFEDAFAEF6027" box="[1140,1310,285,309]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="43CC2A5AFFFB1107EF1BFEDAFAEF6027" box="[1140,1310,285,309]" italics="true" pageId="2" pageNumber="343">N. serratipalma</emphasis>
</taxonomicName>
, apparently absent in
<taxonomicName id="B6B88DCBFFFB1107EB92FE84FE886045" box="[253,377,319,343]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFFB1107EB92FE84FE886045" box="[253,377,319,343]" italics="true" pageId="2" pageNumber="343">N. draconis</emphasis>
</taxonomicName>
].
</paragraph>
</caption>
<paragraph id="7107F648FFFB1107EBF0FEB0FC2460A8" blockId="2.[159,1344,372,826]" pageId="2" pageNumber="343">Thoracic somites 1 2 3 4 5 6 7 8 Appendages Maxillipeds Pereopods</paragraph>
<paragraph id="7107F648FFFB1107E97BFE10FEF8645C" blockId="2.[159,1344,372,826]" lastBlockId="2.[151,1437,901,2006]" pageId="2" pageNumber="343">
1 2 3 1 2 3 4 5 Pleurobranch 0 0 0 0 0 0 0 0 Arthrobranchs 1 2 2 2 2 2 2 0 Podobranchs 0 1 1 1 1 1 0/r 0 Epipods 1 1 1 1 1 1 1 0 Mastigobranchs 0 0 1 1 1 1 1 0 Exopod 1 1 1 0 0 0 0 0 Setobranchs 0 0 1 1 1 1 1/0* 0
<emphasis id="43CC2A5AFFFB1107EBA8FC41FEC5628C" bold="true" box="[199,308,901,926]" pageId="2" pageNumber="343">Remarks</emphasis>
. As noted by
<bibRefCitation id="15298BB9FFFB1107EAB9FC41FD80628C" author="Anker" box="[470,625,901,926]" pageId="2" pageNumber="343" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker (2014)</bibRefCitation>
, the branchial formula of
<taxonomicName id="B6B88DCBFFFB1107E8F5FC41FBE5628F" box="[922,1044,901,925]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFB1107E8F5FC41FBE5628F" box="[922,1044,901,925]" italics="true" pageId="2" pageNumber="343">Naushonia</emphasis>
</taxonomicName>
remains incompletely known, due partly to the rarity of specimens, particularly in species formerly assigned to
<taxonomicName id="B6B88DCBFFFB1107EF58FC6DFAF562D3" box="[1079,1284,937,961]" class="Malacostraca" family="Laomediidae" genus="Espeleonaushonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFB1107EF58FC6DFAF562D3" box="[1079,1284,937,961]" italics="true" pageId="2" pageNumber="343">Espeleonaushonia</emphasis>
</taxonomicName>
. In fact, the complete branchial formula remains unknown in
<taxonomicName id="B6B88DCBFFFB1107E9B5FC0AFC9862F7" box="[730,873,973,997]" class="Malacostraca" family="Laomediidae" genus="Espeleonaushonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="augudrea">
<emphasis id="43CC2A5AFFFB1107E9B5FC0AFC9862F7" box="[730,873,973,997]" italics="true" pageId="2" pageNumber="343">N. augudrea</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107E813FC0AFBEE62F7" box="[892,1055,974,997]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="macginitiei">
<emphasis id="43CC2A5AFFFB1107E813FC0AFBEE62F7" box="[892,1055,974,997]" italics="true" pageId="2" pageNumber="343">N. macginitiei</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107EF5EFC0AFB3162F7" box="[1073,1216,974,997]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="manningi">
<emphasis id="43CC2A5AFFFB1107EF5EFC0AFB3162F7" box="[1073,1216,974,997]" italics="true" pageId="2" pageNumber="343">N. manningi</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107EFBCFC0AFA8162F7" box="[1235,1392,973,997]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="palauensis">
<emphasis id="43CC2A5AFFFB1107EFBCFC0AFA8162F7" box="[1235,1392,973,997]" italics="true" pageId="2" pageNumber="343">N. palauensis</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107EEEDFC0AFEEA651B" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="43CC2A5AFFFB1107EEEDFC0AFEEA651B" italics="true" pageId="2" pageNumber="343">N. panamensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="B6B88DCBFFFB1107EA33FC36FE24651B" box="[348,469,1010,1033]" class="Malacostraca" family="Laomediidae" genus="Coralliocrangon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="43CC2A5AFFFB1107EA33FC36FE24651B" box="[348,469,1010,1033]" italics="true" pageId="2" pageNumber="343">N. perrieri</emphasis>
</taxonomicName>
, and was also unknown, until now, in
<taxonomicName id="B6B88DCBFFFB1107E8FEFC36FBE7651B" box="[913,1046,1009,1033]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="draconis">
<emphasis id="43CC2A5AFFFB1107E8FEFC36FBE7651B" box="[913,1046,1009,1033]" italics="true" pageId="2" pageNumber="343">N. draconis</emphasis>
</taxonomicName>
and
<taxonomicName id="B6B88DCBFFFB1107EF3FFC36FB25651B" box="[1104,1236,1010,1033]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFB1107EF3FFC36FB25651B" box="[1104,1236,1010,1033]" italics="true" pageId="2" pageNumber="343">N. japonica</emphasis>
</taxonomicName>
(see below). The presence/absence of an arthrobranch on the first maxilliped and of a podobranch on the fourth pereopod appear to be intragenerically variable characters in
<taxonomicName id="B6B88DCBFFFB1107E90CFBFDFC8F6540" authority="Anker 2014" authorityName="Anker" authorityYear="2014" box="[611,894,1081,1106]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFB1107E90CFBFDFD2C6543" box="[611,733,1081,1105]" italics="true" pageId="2" pageNumber="343">Naushonia</emphasis>
(
<bibRefCitation id="15298BB9FFFB1107E984FBFDFC876540" author="Anker" box="[747,886,1081,1106]" pageId="2" pageNumber="343" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
)
</taxonomicName>
. Considering the difficulties to observe the basal area of the anterior maxillipeds, we tried to verify the presence or absence of an arthrobranch at the base of the first maxilliped in the six species treated in this study, through a careful dissection of the upper-lying appendages. We here confirm that one arthrobranch is constantly present on the first maxilliped in all six species, although this gill was previously reported as absent in
<taxonomicName id="B6B88DCBFFFB1107E931FB0EFBEB65F0" authority="Dworschak et al. 2006" authorityName="Dworschak et al." authorityYear="2006" box="[606,1050,1225,1250]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFB1107E931FB0EFD1565F3" box="[606,740,1226,1249]" italics="true" pageId="2" pageNumber="343">N. carinata</emphasis>
(
<bibRefCitation id="15298BB9FFFB1107E995FB0DFBE365F0" author="Dworschak" box="[762,1042,1225,1250]" pageId="2" pageNumber="343" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak
<emphasis id="43CC2A5AFFFB1107E8E6FB0EFC3965F3" box="[905,968,1225,1249]" italics="true" pageId="2" pageNumber="343">et al.</emphasis>
2006
</bibRefCitation>
)
</taxonomicName>
, which was confirmed by a reexamination of the two
<typeStatus id="AE0348EAFFFB1107EAC9FB2AFDE06414" box="[422,529,1262,1286]" pageId="2" pageNumber="343" type="paratype">paratypes</typeStatus>
of this species by Peter C. Dworschak (see “Remarks” under
<taxonomicName id="B6B88DCBFFFB1107EFB8FB2AFAA96417" box="[1239,1368,1262,1285]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFB1107EFB8FB2AFAA96417" box="[1239,1368,1262,1285]" italics="true" pageId="2" pageNumber="343">N. carinata</emphasis>
</taxonomicName>
). The podobranch on the fourth pereopod is absent or greatly reduced to a rudimentary bud in all six species reported in this study.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFB1106EBA8FA9DFBC161C7" lastPageId="3" lastPageNumber="344" pageId="2" pageNumber="343" type="materials_examined">
<paragraph id="7107F648FFFB1106EBA8FA9DFBC161C7" blockId="2.[151,1437,901,2006]" lastBlockId="3.[151,1436,151,213]" lastPageId="3" lastPageNumber="344" pageId="2" pageNumber="343">
<bibRefCitation id="15298BB9FFFB1107EBA8FA9DFDCB6460" author="Martin" box="[199,570,1369,1394]" pageId="2" pageNumber="343" refString="Martin, J. W. &amp; Abele, L. G. (1982) Naushonia panamensis, new species (Decapoda: Thalassinidea: Laomediidae) from the Pacific coast of Panama, with notes on the genus. Proceedings of the Biological Society of Washington, 95, 478 - 483." type="journal article" year="1982">Martin &amp; Abele (1982: 482, key)</bibRefCitation>
were the first to use the shape of the postorbital spines of the carapace as one of the key characters differentiating species of
<taxonomicName id="B6B88DCBFFFB1107E9C6FAB9FCD36487" box="[681,802,1405,1429]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFB1107E9C6FAB9FCD36487" box="[681,802,1405,1429]" italics="true" pageId="2" pageNumber="343">Naushonia</emphasis>
</taxonomicName>
, i.e., “simple” in
<taxonomicName id="B6B88DCBFFFB1107E884FABAFB606487" box="[1003,1169,1406,1429]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="43CC2A5AFFFB1107E884FABAFB606487" box="[1003,1169,1406,1429]" italics="true" pageId="2" pageNumber="343">N. panamensis</emphasis>
</taxonomicName>
, versus “bifid or trifid” in
<taxonomicName id="B6B88DCBFFFB1107EBD1FA66FE7064AB" box="[190,385,1441,1465]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="crangonoides">
<emphasis id="43CC2A5AFFFB1107EBD1FA66FE7064AB" box="[190,385,1441,1465]" italics="true" pageId="2" pageNumber="343">N. crangonoides</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107EAF5FA66FDA964AB" box="[410,600,1442,1465]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFFB1107EAF5FA66FDA964AB" box="[410,600,1442,1465]" italics="true" pageId="2" pageNumber="343">N. portoricensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B6B88DCBFFFB1107E9CBFA66FCBD64AB" box="[676,844,1442,1465]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="macginitiei">
<emphasis id="43CC2A5AFFFB1107E9CBFA66FCBD64AB" box="[676,844,1442,1465]" italics="true" pageId="2" pageNumber="343">N. macginitiei</emphasis>
</taxonomicName>
. This distinction originates from the somewhat diagrammatic figures of the carapace in
<bibRefCitation id="15298BB9FFFB1107E939FA02FBF164CC" author="Goy" box="[598,1024,1477,1502]" pageId="2" pageNumber="343" refString="Goy, J. W. &amp; Provenzano, A. J. Jr. (1978) Larval development of the rare burrowing mud shrimp Naushonia crangonoides Kingsley (Decapoda: Thalassinidea: Laomediidae). Biological Bulletin, 154, 241 - 61. http: // dx. doi. org / 10.2307 / 1541125" type="journal article" year="1978">Goy &amp; Provenzano (1978: fig. 6AD)</bibRefCitation>
, in which the postorbital spines were illustrated as simple in
<taxonomicName id="B6B88DCBFFFB1107EAF5FA2EFDE06713" box="[410,529,1514,1537]" class="Malacostraca" family="Laomediidae" genus="Coralliocrangon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="43CC2A5AFFFB1107EAF5FA2EFDE06713" box="[410,529,1514,1537]" italics="true" pageId="2" pageNumber="343">N. perrieri</emphasis>
</taxonomicName>
, and as bifid or multifid in
<taxonomicName id="B6B88DCBFFFB1107E82AFA2EFC0C6713" box="[837,1021,1513,1537]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="crangonoides">
<emphasis id="43CC2A5AFFFB1107E82AFA2EFC0C6713" box="[837,1021,1513,1537]" italics="true" pageId="2" pageNumber="343">N. crangonoides</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107EF63FA2EFB5B6713" box="[1036,1194,1514,1537]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="macginitiei">
<emphasis id="43CC2A5AFFFB1107EF63FA2EFB5B6713" box="[1036,1194,1514,1537]" italics="true" pageId="2" pageNumber="343">N. macginitiei</emphasis>
</taxonomicName>
and
<taxonomicName id="B6B88DCBFFFB1107EF8EFA2EFA646713" box="[1249,1429,1514,1537]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFFB1107EF8EFA2EFA646713" box="[1249,1429,1514,1537]" italics="true" pageId="2" pageNumber="343">N. portoricensis</emphasis>
</taxonomicName>
. Martin &amp; Abeles (1982) key has been followed by all successive workers expanding their keys to include additional species, i.e.,
<bibRefCitation id="15298BB9FFFB1107EAC0F9F6FD806758" author="Berggren" box="[431,625,1586,1610]" pageId="2" pageNumber="343" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. http: // dx. doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren (1992)</bibRefCitation>
,
<bibRefCitation id="15298BB9FFFB1107E9ECF9F5FC706758" box="[643,897,1585,1610]" pageId="2" pageNumber="343" refString="Alvarez, F., Villalobos, J. L. &amp; Iliffe, T. M. (2000) Naushonia manningi, new species (Decapoda: Thalassinidea: Laomediidae), from Acklins Island, Bahamas. Journal of Crustacean Biology, 20, 192 - 198." type="journal article">
Alvarez
<emphasis id="43CC2A5AFFFB1107E986F9F6FCD6675B" box="[745,807,1585,1609]" italics="true" pageId="2" pageNumber="343">et al.</emphasis>
(2000)
</bibRefCitation>
,
<bibRefCitation id="15298BB9FFFB1107E8FCF9F5FB416758" author="Komai" box="[915,1200,1585,1610]" pageId="2" pageNumber="343" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504, 31 - 46." type="journal article" year="2010">Komai &amp; Anker (2010)</bibRefCitation>
, and
<bibRefCitation id="15298BB9FFFB1107EF98F9F5FA696758" author="Anker" box="[1271,1432,1585,1610]" pageId="2" pageNumber="343" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker (2014)</bibRefCitation>
. However, during this study, we examined a specimen of
<taxonomicName id="B6B88DCBFFFB1107E87EF991FC7A677F" box="[785,907,1621,1645]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFB1107E87EF991FC7A677F" box="[785,907,1621,1645]" italics="true" pageId="2" pageNumber="343">Naushonia</emphasis>
</taxonomicName>
from
<collectingCountry id="09AFB6D8FFFB1107E8A1F992FBB3677C" box="[974,1090,1621,1646]" name="Sint Martin" pageId="2" pageNumber="343">St. Martin</collectingCountry>
, French Antilles, which agrees well with
<taxonomicName id="B6B88DCBFFFB1107EA61F9BEFE366783" box="[270,455,1658,1681]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFFB1107EA61F9BEFE366783" box="[270,455,1658,1681]" italics="true" pageId="2" pageNumber="343">N. portoricensis</emphasis>
</taxonomicName>
, except for the presence of a simple, acuminate, postorbital spine. This observation raised the question whether the postorbital spine is bifid or multifid in the three above-listed American species of
<taxonomicName id="B6B88DCBFFFB1107EBF8F905FEE167CB" box="[151,272,1729,1753]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFB1107EBF8F905FEE167CB" box="[151,272,1729,1753]" italics="true" pageId="2" pageNumber="343">Naushonia</emphasis>
</taxonomicName>
. Original descriptions of
<taxonomicName id="B6B88DCBFFFB1107E92AF906FCF267CB" box="[581,771,1729,1753]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="crangonoides">
<emphasis id="43CC2A5AFFFB1107E92AF906FCF267CB" box="[581,771,1729,1753]" italics="true" pageId="2" pageNumber="343">N. crangonoides</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFFB1107E878F906FC4A67CB" box="[791,955,1730,1753]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="macginitiei">
<emphasis id="43CC2A5AFFFB1107E878F906FC4A67CB" box="[791,955,1730,1753]" italics="true" pageId="2" pageNumber="343">N. macginitiei</emphasis>
</taxonomicName>
and
<taxonomicName id="B6B88DCBFFFB1107E893F906FB4767CB" box="[1020,1206,1730,1753]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFFB1107E893F906FB4767CB" box="[1020,1206,1730,1753]" italics="true" pageId="2" pageNumber="343">N. portoricensis</emphasis>
</taxonomicName>
do not specifically describe the structure of the postorbital spine, however, all published figures actually appear to show simple postorbital spines (
<bibRefCitation id="15298BB9FFFB1107EA05F8CDFDE26630" author="Kingsley" box="[362,531,1801,1826]" pageId="2" pageNumber="343" refString="Kingsley, J. S. (1897) On a new genus and two new species of macrurous Crustacea. Bulletin of the Essex Institute, 27, 95 - 99." type="journal article" year="1897">Kingsley 1897</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFB1107E970F8CDFD316630" author="Rathbun" box="[543,704,1801,1826]" pageId="2" pageNumber="343" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Puerto Rico. United States Fish Commission Bulletin, 20, 1 - 137." type="journal article" year="1901">Rathbun 1901</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFB1107E9A3F8CDFC746630" author="Thompson" box="[716,901,1801,1826]" pageId="2" pageNumber="343" refString="Thompson, M. T. (1903) A rare thalassinid and its larva. Proceedings of the Boston Society of Natural History, 31, 1 - 21. http: // dx. doi. org / 10.1080 / 00222930308678827" type="journal article" year="1903">Thompson 1903</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFB1107E8FEF8CDFBDF6630" author="Glassell" box="[913,1070,1801,1826]" pageId="2" pageNumber="343" refString="Glassell, S. A. (1938) New and obscure decapod Crustacea from the West American coasts. Transactions of the San Diego Society of Natural History, 3, 411 - 454." type="journal article" year="1938">Glassell 1938</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFB1107EF55F8CDFB336630" author="Chace" box="[1082,1218,1801,1826]" pageId="2" pageNumber="343" refString="Chace, F. A. Jr. (1939) On the systematic status of the crustacean genera Naushonia, Homoriscus and Coralliocrangon. Annals and Magazine of Natural History, 3 (11), 524 - 530. http: // dx. doi. org / 10.1080 / 03745481.1939.9723632" type="journal article" year="1939">Chace 1939</bibRefCitation>
). In order to clarify this problem, we asked Dr. Rafael Lemaitre (
<collectionCode id="17A96E8DFFFB1107E9C5F8EAFD0F6654" LSID="urn:lsid:biocol.org:col:34871" box="[682,766,1838,1862]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="343">USNM</collectionCode>
) to re-examine material of
<taxonomicName id="B6B88DCBFFFB1107EF2BF8E9FB4F6657" box="[1092,1214,1837,1861]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFB1107EF2BF8E9FB4F6657" box="[1092,1214,1837,1861]" italics="true" pageId="2" pageNumber="343">Naushonia</emphasis>
</taxonomicName>
studied by
<bibRefCitation id="15298BB9FFFB1107EE24F8EAFDF86678" author="Goy" pageId="2" pageNumber="343" refString="Goy, J. W. &amp; Provenzano, A. J. Jr. (1979) Juvenile morphology of the rare burrowing mud shrimp Naushonia crangonoides Kingsley, with a review of the genus Naushonia (Decapoda: Thalassinidea: Laomediidae). Proceedings of the Biological Society of Washington, 92, 339 - 359." type="journal article" year="1979">Goy &amp; Provenzano (1979: pp. 348351)</bibRefCitation>
, viz.
<taxonomicName id="B6B88DCBFFFB1107E92BF896FD0C667B" box="[580,765,1873,1897]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="crangonoides">
<emphasis id="43CC2A5AFFFB1107E92BF896FD0C667B" box="[580,765,1873,1897]" italics="true" pageId="2" pageNumber="343">N. crangonoides</emphasis>
</taxonomicName>
(
<collectionCode id="17A96E8DFFFB1107E863F896FCAE6678" LSID="urn:lsid:biocol.org:col:34871" box="[780,863,1874,1898]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="343">USNM</collectionCode>
34143,
<date id="0506D088FFFB1107E8D6F896FBE16678" box="[953,1040,1874,1898]" pageId="2" pageNumber="343" value="77-10-22">102277</date>
,
<date id="0506D088FFFB1107EF74F896FB836678" box="[1051,1138,1874,1898]" pageId="2" pageNumber="343" value="79-10-22">102279</date>
,
<date id="0506D088FFFB1107EF12F896FB246678" box="[1149,1237,1874,1898]" pageId="2" pageNumber="343" value="80-10-22">102280</date>
),
<taxonomicName id="B6B88DCBFFFB1107EF87F896FA6D667B" box="[1256,1436,1874,1897]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFFB1107EF87F896FA6D667B" box="[1256,1436,1874,1897]" italics="true" pageId="2" pageNumber="343">N. portoricensis</emphasis>
</taxonomicName>
(
<typeStatus id="AE0348EAFFFB1107EBF0F8B1FEF6669C" box="[159,263,1909,1934]" pageId="2" pageNumber="343" type="holotype">holotype</typeStatus>
:
<collectionCode id="17A96E8DFFFB1107EA7BF8B2FE94669C" LSID="urn:lsid:biocol.org:col:34871" box="[276,357,1910,1934]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="343">USNM</collectionCode>
23782, 155101), and
<taxonomicName id="B6B88DCBFFFB1107E90AF8B2FCF6669F" box="[613,775,1910,1933]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="macginitiei">
<emphasis id="43CC2A5AFFFB1107E90AF8B2FCF6669F" box="[613,775,1910,1933]" italics="true" pageId="2" pageNumber="343">N. macginitiei</emphasis>
</taxonomicName>
(
<collectionCode id="17A96E8DFFFB1107E877F8B2FC9A669C" LSID="urn:lsid:biocol.org:col:34871" box="[792,875,1910,1934]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="343">USNM</collectionCode>
171604, 171605, 14449). Dr. R. Lemaitre (pers. comm.,
<date id="0506D088FFFB1107EB9DF85EFE5966A0" box="[242,424,1946,1970]" pageId="2" pageNumber="343" value="2014-08-12">12 August 2014</date>
) informed us that none of the
<collectionCode id="17A96E8DFFFB1107E994F85EFCBD66A0" LSID="urn:lsid:biocol.org:col:34871" box="[763,844,1946,1970]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="343">USNM</collectionCode>
specimens has bifid or multifid postorbital spines. It can be concluded that more generally, all species of
<taxonomicName id="B6B88DCBFFFB1107E982F879FC9766C7" box="[749,870,1981,2005]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFB1107E982F879FC9766C7" box="[749,870,1981,2005]" italics="true" pageId="2" pageNumber="343">Naushonia</emphasis>
</taxonomicName>
have simple postorbital spines and never bifid or multifid spines, thus making previous identification keys extremely ambiguous. Therefore, a new, fully revised identification key to all presently known species of
<taxonomicName id="B6B88DCBFFFA1106E9B6FF78FCA361C6" box="[729,850,188,212]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFA1106E9B6FF78FCA361C6" box="[729,850,188,212]" italics="true" pageId="3" pageNumber="344">Naushonia</emphasis>
</taxonomicName>
is presented below.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,295 @@
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<mods:title id="8761855ECE861055DBE7367E8A4E0811">Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key</mods:title>
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<treatment id="F911475EFFFA1100EBF8F96EFC356397" ID-DOI="http://doi.org/10.5281/zenodo.3511421" ID-GBIF-Taxon="119631472" ID-Zenodo-Dep="3511421" LSID="urn:lsid:plazi:treatment:F911475EFFFA1100EBF8F96EFC356397" httpUri="http://treatment.plazi.org/id/F911475EFFFA1100EBF8F96EFC356397" lastPageId="5" lastPageNumber="346" pageId="3" pageNumber="344">
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<paragraph id="7107F648FFFA1106EBF8F96EFCC867D7" blockId="3.[151,825,1706,1766]" box="[151,825,1706,1733]" pageId="3" pageNumber="344">
<heading id="2A4F4124FFFA1106EBF8F96EFCC867D7" bold="true" box="[151,825,1706,1733]" fontSize="11" level="1" pageId="3" pageNumber="344" reason="1">
<taxonomicName id="B6B88DCBFFFA1106EBF8F96EFCC867D7" ID-CoL="883855dd-769a-4f57-9694-b8c4f9ab6ff6" authority="Dworschak, Marin &amp; Anker, 2006" authorityName="Dworschak, Marin &amp; Anker" authorityYear="2006" box="[151,825,1706,1733]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFA1106EBF8F96EFCC867D7" bold="true" box="[151,825,1706,1733]" pageId="3" pageNumber="344">
<emphasis id="43CC2A5AFFFA1106EBF8F96EFE7867D6" bold="true" box="[151,393,1706,1732]" italics="true" pageId="3" pageNumber="344">Naushonia carinata</emphasis>
<bibRefCitation id="15298BB9FFFA1106EAE0F96FFCC867D7" author="Dworschak" box="[399,825,1707,1733]" pageId="3" pageNumber="344" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">Dworschak, Marin &amp; Anker, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7107F648FFFA1106EBF8F909FF1C67F4" blockId="3.[151,825,1706,1766]" box="[151,237,1741,1766]" pageId="3" pageNumber="344">
<figureCitation id="E983EACDFFFA1106EBF8F909FF1C67F4" box="[151,237,1741,1766]" captionStart="FIGURE 1. A" captionStartId="4.[151,250,1425,1447]" captionTargetBox="[258,1328,193,1403]" captionTargetId="figure@4.[258,1329,193,1403]" captionTargetPageId="4" captionText="FIGURE 1. A, Naushonia carinata Dworschak, Marin &amp; Anker, 2006, female (pocl 7.1 mm) from Iriomote, Japan, RUMF-ZC 3677, living individual, dorsal view; B, Naushonia japonica Komai, 2004, female (pocl 5.3 mm) from Katsuura, Japan, CBM- ZC 12594, living individual, dorsal view." httpUri="https://zenodo.org/record/231781/files/figure.png" pageId="3" pageNumber="344">Fig. 1A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFA1106EBF8F8D1FD5B6658" pageId="3" pageNumber="344" type="reference_group">
<paragraph id="7107F648FFFA1106EBF8F8D1FD5B6658" blockId="3.[151,1436,1813,1866]" pageId="3" pageNumber="344">
<treatmentCitationGroup id="51A8D166FFFA1106EBF8F8D1FD5B6658" infered="true" pageId="3" pageNumber="344">
<taxonomicName id="B6B88DCBFFFA1106EBF8F8D1FD9A6639" ID-CoL="883855dd-769a-4f57-9694-b8c4f9ab6ff6" authority="Dworschak et al. 2006: 3" authorityName="Dworschak et al." authorityPageNumber="3" authorityYear="2006" box="[151,619,1813,1835]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFA1106EBF8F8D1FE936639" box="[151,354,1813,1835]" italics="true" pageId="3" pageNumber="344">Naushonia carinata</emphasis>
<treatmentCitation id="F019D059FFFA1106EA07F8D1FD9A6639" author="Dworschak" box="[360,619,1813,1835]" httpUri="http://treatment.plazi.org/id/03BA87DFFF8BB33B1E636FCDFD23FDEA" page="3" pageId="3" pageNumber="344" year="2006">
<bibRefCitation id="15298BB9FFFA1106EA07F8D1FDA26639" author="Dworschak" box="[360,595,1813,1835]" pageId="3" pageNumber="344" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak
<emphasis id="43CC2A5AFFFA1106EA8CF8D2FDE76639" box="[483,534,1813,1835]" italics="true" pageId="3" pageNumber="344">et al.</emphasis>
2006
</bibRefCitation>
: 3
</treatmentCitation>
</taxonomicName>
, figs. 16 (
<typeStatus id="AE0348EAFFFA1106E9B0F8D1FCFD6639" box="[735,780,1813,1835]" pageId="3" pageNumber="344">type</typeStatus>
locality: Dâm Bay of Tré
<collectingCountry id="09AFB6D8FFFA1106EF75F8D1FBAB6639" box="[1050,1114,1813,1835]" name="Iceland" pageId="3" pageNumber="344">Island</collectingCountry>
, Nha Trang,
<collectingCountry id="09AFB6D8FFFA1106EFB0F8D1FAC66639" box="[1247,1335,1813,1835]" name="Vietnam" pageId="3" pageNumber="344">Vietnam</collectingCountry>
).—
<treatmentCitation id="F019D059FFFA1106EE34F8D1FE996658" ID-DOI="http://doi.org/10.5281/zenodo.3500313" ID-Zenodo-Dep="3500313" author="Anker" httpUri="http://treatment.plazi.org/id/AD437D0CF154AD5F94CE106BF4BD1ED3" page="333" pageId="3" pageNumber="344" year="2015">
<bibRefCitation id="15298BB9FFFA1106EE34F8D1FEC76658" author="Anker" pageId="3" pageNumber="344" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans from Lombok, Indonesia. Part I. Alpheidae, Laomediidae, Strahlaxiidae. Zootaxa, 3911 (3), 301 - 342. http: // dx. doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker
<emphasis id="43CC2A5AFFFA1106EBA8F8F1FF0B6658" box="[199,250,1844,1866]" italics="true" pageId="3" pageNumber="344">et al.</emphasis>
2015
</bibRefCitation>
: 333
</treatmentCitation>
, fig. 24A (Lombok,
<collectingCountry id="09AFB6D8FFFA1106E957F8F0FD6E6658" box="[568,671,1844,1866]" name="Indonesia" pageId="3" pageNumber="344">Indonesia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFA1101EBF8F8B2FDCA675F" lastPageId="4" lastPageNumber="345" pageId="3" pageNumber="344" type="materials_examined">
<paragraph id="7107F648FFFA1106EBF8F8B2FD5766C5" blockId="3.[151,1436,1910,2007]" pageId="3" pageNumber="344">
<emphasis id="43CC2A5AFFFA1106EBF8F8B2FE8C669D" bold="true" box="[151,381,1910,1935]" pageId="3" pageNumber="344">Material examined</emphasis>
.
<collectingCountry id="09AFB6D8FFFA1106EAE0F8B3FE22669C" box="[399,467,1911,1934]" name="Japan" pageId="3" pageNumber="344">Japan</collectingCountry>
: 1 ovigerous female (pocl
<quantity id="B6405BADFFFA1106E87EF8B3FC81669D" box="[785,880,1911,1935]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="3" pageNumber="344" unit="mm" value="7.1">7.1 mm</quantity>
), Yaeyama Islands, Iriomote Island, Shirahama, intertidal, soft sediment with sea grass, suction pump, coll. T. Komai,
<date id="0506D088FFFA1106E8C8F85FFBCE66A1" box="[935,1087,1947,1971]" pageId="3" pageNumber="344" value="2014-05-14">14 May 2014</date>
, RUMF-ZC 3677.
<emphasis id="43CC2A5AFFFA1106EBA8F87AFEA066C5" bold="true" box="[199,337,1982,2007]" pageId="3" pageNumber="344">Description</emphasis>
. See
<bibRefCitation id="15298BB9FFFA1106EAE0F87AFD5366C5" box="[399,674,1982,2007]" pageId="3" pageNumber="344" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article">
Dworschak
<emphasis id="43CC2A5AFFFA1106E97AF87BFDBC66C4" box="[533,589,1982,2006]" italics="true" pageId="3" pageNumber="344">et al.</emphasis>
(2006)
</bibRefCitation>
.
</paragraph>
<caption id="25C7A6C0FFFD1101EBF8FA55FDC864F6" httpUri="https://zenodo.org/record/231781/files/figure.png" pageId="4" pageNumber="345" targetBox="[258,1328,193,1403]" targetPageId="4">
<paragraph id="7107F648FFFD1101EBF8FA55FDC864F6" blockId="4.[151,1436,1425,1508]" pageId="4" pageNumber="345">
<emphasis id="43CC2A5AFFFD1101EBF8FA55FEFD64B4" bold="true" box="[151,268,1425,1447]" pageId="4" pageNumber="345">FIGURE 1</emphasis>
. A,
<taxonomicName id="B6B88DCBFFFD1101EA59FA55FC9064B5" authority="Dworschak, Marin &amp; Anker, 2006" authorityName="Dworschak, Marin &amp; Anker" authorityYear="2006" box="[310,865,1425,1447]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFD1101EA59FA55FDF164B5" box="[310,512,1425,1447]" italics="true" pageId="4" pageNumber="345">Naushonia carinata</emphasis>
Dworschak, Marin &amp; Anker, 2006
</taxonomicName>
, female (pocl 7.1 mm) from Iriomote, Japan, RUMF-ZC 3677, living individual, dorsal view; B,
<taxonomicName id="B6B88DCBFFFD1101E942FA6BFC7664D7" authority="Komai, 2004" authorityName="Komai" authorityYear="2004" box="[557,903,1455,1478]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFD1101E942FA6BFD0B64D4" box="[557,762,1455,1478]" italics="true" pageId="4" pageNumber="345">Naushonia japonica</emphasis>
Komai, 2004
</taxonomicName>
, female (pocl 5.3 mm) from Katsuura, Japan, CBM- ZC 12594, living individual, dorsal view.
</paragraph>
</caption>
<paragraph id="7107F648FFFD1101EBA8F9D4FDCA675F" blockId="4.[151,1436,1552,2009]" pageId="4" pageNumber="345">
<emphasis id="43CC2A5AFFFD1101EBA8F9D4FE56673B" bold="true" box="[199,423,1552,1577]" pageId="4" pageNumber="345">Colouration in life</emphasis>
. Body and appendages whitish; yellowish-brown hepatopancreas and pale orange ovary visible through integument (
<figureCitation id="E983EACDFFFD1101EABBF9F0FDDC675F" box="[468,557,1588,1613]" captionStart="FIGURE 1. A" captionStartId="4.[151,250,1425,1447]" captionTargetBox="[258,1328,193,1403]" captionTargetId="figure@4.[258,1329,193,1403]" captionTargetPageId="4" captionText="FIGURE 1. A, Naushonia carinata Dworschak, Marin &amp; Anker, 2006, female (pocl 7.1 mm) from Iriomote, Japan, RUMF-ZC 3677, living individual, dorsal view; B, Naushonia japonica Komai, 2004, female (pocl 5.3 mm) from Katsuura, Japan, CBM- ZC 12594, living individual, dorsal view." httpUri="https://zenodo.org/record/231781/files/figure.png" pageId="4" pageNumber="345">Fig. 1A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFD1101EBA8F99CFCB067CC" pageId="4" pageNumber="345" type="distribution">
<paragraph id="7107F648FFFD1101EBA8F99CFBB46784" blockId="4.[151,1436,1552,2009]" pageId="4" pageNumber="345">
<emphasis id="43CC2A5AFFFD1101EBA8F99CFEA86763" bold="true" box="[199,345,1624,1649]" pageId="4" pageNumber="345">Distribution</emphasis>
. Western Pacific:
<collectingCountry id="09AFB6D8FFFD1101E92DF99CFD506763" box="[578,673,1624,1649]" name="Vietnam" pageId="4" pageNumber="345">Vietnam</collectingCountry>
(Nha Trang Bay),
<collectingCountry id="09AFB6D8FFFD1101E8F5F99CFBE96763" box="[922,1048,1624,1649]" name="Philippines" pageId="4" pageNumber="345">Philippines</collectingCountry>
(Panglao),
<collectingCountry id="09AFB6D8FFFD1101EFC2F99CFAEB6763" box="[1197,1306,1624,1649]" name="Indonesia" pageId="4" pageNumber="345">Indonesia</collectingCountry>
(Lombok) (
<bibRefCitation id="15298BB9FFFD1101EBF1F9B9FE506784" author="Dworschak" box="[158,417,1661,1686]" pageId="4" pageNumber="345" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak
<emphasis id="43CC2A5AFFFD1101EA49F9BAFEAF6787" box="[294,350,1661,1685]" italics="true" pageId="4" pageNumber="345">et al.</emphasis>
2006
</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFD1101EAC3F9B9FD856784" author="Anker" box="[428,628,1661,1686]" pageId="4" pageNumber="345" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans from Lombok, Indonesia. Part I. Alpheidae, Laomediidae, Strahlaxiidae. Zootaxa, 3911 (3), 301 - 342. http: // dx. doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker
<emphasis id="43CC2A5AFFFD1101EA96F9BAFDC36787" box="[505,562,1661,1685]" italics="true" pageId="4" pageNumber="345">et al.</emphasis>
2015
</bibRefCitation>
);
<collectingCountry id="09AFB6D8FFFD1101E9E7F9BAFD396787" box="[648,712,1662,1685]" name="Japan" pageId="4" pageNumber="345">Japan</collectingCountry>
(Ryukyu Islands) (present study).
</paragraph>
<paragraph id="7107F648FFFD1101EBA8F964FCB067CC" blockId="4.[151,1436,1552,2009]" pageId="4" pageNumber="345">
<emphasis id="43CC2A5AFFFD1101EBA8F964FED267AB" bold="true" box="[199,291,1696,1721]" pageId="4" pageNumber="345">Habitat</emphasis>
. Intertidal and shallow subtidal, presumably in burrows of the callianassid ghost shrimp
<taxonomicName id="B6B88DCBFFFD1101EE43F965FCCC67CC" authority="(A. Milne-Edwards, 1870) (Dworschak et al. 2006)" authorityName="(A. Milne-Edwards, 1870) (Dworschak et al." authorityYear="2006" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="43CC2A5AFFFD1101EE43F965FF0267CF" italics="true" pageId="4" pageNumber="345">Glypturus armatus</emphasis>
(
<bibRefCitation id="15298BB9FFFD1101EA6DF901FDEB67CF" author="Milne-Edwards" box="[258,538,1733,1758]" pageId="4" pageNumber="345" refString="Milne-Edwards, A. (1870) Revision du genre Callianassa (Leach) et description de plusieurs especes nouvelles de ce groupe fausant partie de la collection du museum. Nouvelle Archives du Museum d'Histoire Naturelle, Paris, 6, 75 - 101." type="journal article" year="1870">A. Milne-Edwards, 1870</bibRefCitation>
) (
<bibRefCitation id="15298BB9FFFD1101E95EF901FCC467CC" author="Dworschak" box="[561,821,1733,1758]" pageId="4" pageNumber="345" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak
<emphasis id="43CC2A5AFFFD1101E9D6F902FD0367CF" box="[697,754,1733,1757]" italics="true" pageId="4" pageNumber="345">et al.</emphasis>
2006
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFD1100EBA8F92CFB216076" lastPageId="5" lastPageNumber="346" pageId="4" pageNumber="345" type="discussion">
<paragraph id="7107F648FFFD1101EBA8F92CFDB3667C" blockId="4.[151,1436,1552,2009]" pageId="4" pageNumber="345">
<emphasis id="43CC2A5AFFFD1101EBA8F92CFEC56613" bold="true" box="[199,308,1768,1793]" pageId="4" pageNumber="345">Remarks</emphasis>
.
<taxonomicName id="B6B88DCBFFFD1101EA28F92DFDDA6612" box="[327,555,1769,1793]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFD1101EA28F92DFDDA6612" box="[327,555,1769,1793]" italics="true" pageId="4" pageNumber="345">Naushonia carinata</emphasis>
</taxonomicName>
was originally described on the basis of four
<typeStatus id="AE0348EAFFFD1101EF38F92DFB766613" box="[1111,1159,1769,1793]" pageId="4" pageNumber="345">type</typeStatus>
specimens, viz. female
<typeStatus id="AE0348EAFFFD1101EBF8F8C9FF0D6634" box="[151,252,1805,1830]" pageId="4" pageNumber="345" type="holotype">holotype</typeStatus>
, male allotype and two female
<typeStatus id="AE0348EAFFFD1101E91FF8CAFD2F6634" box="[624,734,1806,1830]" pageId="4" pageNumber="345" type="paratype">paratypes</typeStatus>
, from Nha Trang,
<collectingCountry id="09AFB6D8FFFD1101E8D3F8C9FBEF6634" box="[956,1054,1805,1830]" name="Vietnam" pageId="4" pageNumber="345">Vietnam</collectingCountry>
, and one additional female from Panglao, near Bohol, in the
<collectingCountry id="09AFB6D8FFFD1101EABBF8F4FDA3665B" box="[468,594,1840,1865]" name="Philippines" pageId="4" pageNumber="345">Philippines</collectingCountry>
(
<bibRefCitation id="15298BB9FFFD1101E90EF8F4FC98665B" author="Dworschak" box="[609,873,1840,1865]" pageId="4" pageNumber="345" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak
<emphasis id="43CC2A5AFFFD1101E984F8F5FCD5665B" box="[747,804,1841,1865]" italics="true" pageId="4" pageNumber="345">et al.</emphasis>
2006
</bibRefCitation>
). More recently,
<bibRefCitation id="15298BB9FFFD1101EF47F8F4FAF3665B" box="[1064,1282,1840,1865]" pageId="4" pageNumber="345" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans from Lombok, Indonesia. Part I. Alpheidae, Laomediidae, Strahlaxiidae. Zootaxa, 3911 (3), 301 - 342. http: // dx. doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article">
Anker
<emphasis id="43CC2A5AFFFD1101EF19F8F5FB5E665B" box="[1142,1199,1841,1865]" italics="true" pageId="4" pageNumber="345">et al.</emphasis>
(2015)
</bibRefCitation>
recorded this species also from Lombok,
<collectingCountry id="09AFB6D8FFFD1101EAA2F891FDCF667C" box="[461,574,1877,1902]" name="Indonesia" pageId="4" pageNumber="345">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="7107F648FFFD1100EBA8F8BCFB216076" blockId="4.[151,1436,1552,2009]" lastBlockId="5.[151,1436,151,645]" lastPageId="5" lastPageNumber="346" pageId="4" pageNumber="345">
The present specimen agrees well with the description provided by
<bibRefCitation id="15298BB9FFFD1101E8D6F8BCFB3A6683" box="[953,1227,1912,1937]" pageId="4" pageNumber="345" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article">
Dworschak
<emphasis id="43CC2A5AFFFD1101EF50F8BDFB866683" box="[1087,1143,1913,1937]" italics="true" pageId="4" pageNumber="345">et al.</emphasis>
(2006)
</bibRefCitation>
, except for the gill formula. According to
<bibRefCitation id="15298BB9FFFD1101EAF3F859FD5A66A4" author="Dworschak" box="[412,683,1949,1974]" pageId="4" pageNumber="345" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak
<emphasis id="43CC2A5AFFFD1101E94BF85AFDAC66A7" box="[548,605,1949,1973]" italics="true" pageId="4" pageNumber="345">et al.</emphasis>
(2006
</bibRefCitation>
:
<tableCitation id="3C3AC3F3FFFD1101E9D8F859FCFA66A7" box="[695,779,1949,1974]" captionStart="TABLE 1" captionStartId="2.[151,239,254,277]" captionText="TABLE 1. Gill and exopod formula of the six species of Naushonia treated in this study. [r—reduced; * setobranchs present on the fourth pereopod in N. carinata, N. japonica, N. lactoalbida, N. portoricensis, N. serratipalma, apparently absent in N. draconis]." pageId="4" pageNumber="345">Table 1</tableCitation>
, figure 4f), in the
<typeStatus id="AE0348EAFFFD1101E8B1F85AFBFF66A4" box="[990,1038,1950,1974]" pageId="4" pageNumber="345">type</typeStatus>
specimens of
<taxonomicName id="B6B88DCBFFFD1101EFDBF85AFAC466A7" box="[1204,1333,1950,1973]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFD1101EFDBF85AFAC466A7" box="[1204,1333,1950,1973]" italics="true" pageId="4" pageNumber="345">N. carinata</emphasis>
</taxonomicName>
, the first maxilliped is devoid of an arthrobranch at its base. In contrast, our specimen from Iriomote Island has a single, well-developed, multi-lamellate arthrobranch near the base of the first maxilliped. At our request, Dr. Peter C. Dworschak re-examined two
<typeStatus id="AE0348EAFFFC1100EA8FFF79FDBD61C7" box="[480,588,189,213]" pageId="5" pageNumber="346" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="B6B88DCBFFFC1100E91FFF79FD1E61C6" box="[624,751,189,212]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFC1100E91FFF79FD1E61C6" box="[624,751,189,212]" italics="true" pageId="5" pageNumber="346">N. carinata</emphasis>
</taxonomicName>
deposited in the Naturhistorisches Museum in Wien, Vienna,
<collectingCountry id="09AFB6D8FFFC1100EBF8FF1BFF1B61EA" box="[151,234,223,248]" name="Austria" pageId="5" pageNumber="346">Austria</collectingCountry>
(NHMW), and confirmed that in these specimens there is indeed no arthrobranch at the base of the first maxilliped. Therefore, it seems most reasonable to assume that the development of this arthrobranch is a variable character in
<taxonomicName id="B6B88DCBFFFC1100EA4DFEEDFE50602D" box="[290,417,296,320]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFC1100EA4DFEEDFE50602D" box="[290,417,296,320]" italics="true" pageId="5" pageNumber="346">N. carinata</emphasis>
</taxonomicName>
. For instance,
<bibRefCitation id="15298BB9FFFC1100E928FEE3FCEA6052" box="[583,795,295,320]" pageId="5" pageNumber="346" refString="Astall, C. A., Anderson, S. J. &amp; Taylor, A. C. (1997) Comparative studies of the branchial morphology, gill area and gill ultrastructure of some thalassinidean mud-shrimps (Crustacea: Decapoda: Thalassinidea). Journal of Zoology, London, 241, 665 - 688. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1997. tb 05740. x" type="journal article">
Astall
<emphasis id="43CC2A5AFFFC1100E9FEFEEDFD386052" box="[657,713,296,320]" italics="true" pageId="5" pageNumber="346">et al.</emphasis>
(1997)
</bibRefCitation>
reported a variation in the number of gills present on the first and second maxillipeds in the northeastern Atlantic laomediid
<taxonomicName id="B6B88DCBFFFC1100E8E7FE89FB3A6076" authority="Nardo, 1847" authorityName="Nardo" authorityYear="1847" box="[904,1227,332,357]" class="Malacostraca" family="Laomediidae" genus="Jaxea" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="nocturna">
<emphasis id="43CC2A5AFFFC1100E8E7FE89FBC76076" box="[904,1078,333,356]" italics="true" pageId="5" pageNumber="346">Jaxea nocturna</emphasis>
<bibRefCitation id="15298BB9FFFC1100EF53FE88FB3A6076" author="Nardo" box="[1084,1227,332,357]" pageId="5" pageNumber="346" refString="Nardo, D. (1847) Sinonimia moderna delle specie registrate nell'opera intitolata: descrizione de'crostacei, de'testacei e de'pesci che abitano le Lagune e Golfo Veneto, rappresentati in figure, a chiaroscuro ed a colori dall' Abate Stefano Chiereghini Ven. Clodiense applicata per commissione governativa: i - xi, 1 - 127. Venice." type="book chapter" year="1847">Nardo, 1847</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFC1100EBA8FEB4FC356397" pageId="5" pageNumber="346" type="reference_group">
<paragraph id="7107F648FFFC1100EBA8FEB4FC356397" blockId="5.[151,1436,151,645]" pageId="5" pageNumber="346">
<taxonomicName id="B6B88DCBFFFC1100EBA8FEB4FE576095" ID-CoL="883855dd-769a-4f57-9694-b8c4f9ab6ff6" box="[199,422,368,392]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFC1100EBA8FEB4FE576095" box="[199,422,368,392]" italics="true" pageId="5" pageNumber="346">Naushonia carinata</emphasis>
</taxonomicName>
is one of the most distinctive species of the genus, recognizable by the generally stout and wide body, the presence of three sharply defined, longitudinal carinae (one median and two submedian) on the first to fifth pleomeres, and the relatively broad cheliped palm (
<figureCitation id="E983EACDFFFC1100E841FE73FC7960C2" box="[814,904,439,464]" captionStart="FIGURE 1. A" captionStartId="4.[151,250,1425,1447]" captionTargetBox="[258,1328,193,1403]" captionTargetId="figure@4.[258,1329,193,1403]" captionTargetPageId="4" captionText="FIGURE 1. A, Naushonia carinata Dworschak, Marin &amp; Anker, 2006, female (pocl 7.1 mm) from Iriomote, Japan, RUMF-ZC 3677, living individual, dorsal view; B, Naushonia japonica Komai, 2004, female (pocl 5.3 mm) from Katsuura, Japan, CBM- ZC 12594, living individual, dorsal view." httpUri="https://zenodo.org/record/231781/files/figure.png" pageId="5" pageNumber="346">Fig. 1A</figureCitation>
).
<bibRefCitation id="15298BB9FFFC1100E8F2FE73FB5E60C2" box="[925,1199,439,464]" pageId="5" pageNumber="346" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article">
Dworschak
<emphasis id="43CC2A5AFFFC1100EF4BFE7DFBAC60C2" box="[1060,1117,440,464]" italics="true" pageId="5" pageNumber="346">et al.</emphasis>
(2006)
</bibRefCitation>
suggested a possible association of
<taxonomicName id="B6B88DCBFFFC1100EA53FE19FE4F60E6" ID-CoL="883855dd-769a-4f57-9694-b8c4f9ab6ff6" box="[316,446,477,500]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="43CC2A5AFFFC1100EA53FE19FE4F60E6" box="[316,446,477,500]" italics="true" pageId="5" pageNumber="346">N. carinata</emphasis>
</taxonomicName>
with spacious burrows of the large callianassid ghost shrimp,
<taxonomicName id="B6B88DCBFFFC1100EFE3FE18FA9060E6" box="[1164,1377,476,500]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="43CC2A5AFFFC1100EFE3FE18FA9060E6" box="[1164,1377,476,500]" italics="true" pageId="5" pageNumber="346">Glypturus armatus</emphasis>
</taxonomicName>
. The Indonesian specimen recently reported by
<bibRefCitation id="15298BB9FFFC1100E91AFE3BFCBF630A" box="[629,846,511,536]" pageId="5" pageNumber="346" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans from Lombok, Indonesia. Part I. Alpheidae, Laomediidae, Strahlaxiidae. Zootaxa, 3911 (3), 301 - 342. http: // dx. doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article">
Anker
<emphasis id="43CC2A5AFFFC1100E9ACFDC5FD0D630A" box="[707,764,512,536]" italics="true" pageId="5" pageNumber="346">et al.</emphasis>
(2015)
</bibRefCitation>
was collected from a burrow of unknown host, on a shallow sea grass flat with abundance of mounds and burrow entrance holes of
<taxonomicName id="B6B88DCBFFFC1100EF73FDE0FB69632E" box="[1052,1176,548,572]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="43CC2A5AFFFC1100EF73FDE0FB69632E" box="[1052,1176,548,572]" italics="true" pageId="5" pageNumber="346">G. armatus</emphasis>
</taxonomicName>
(A. Anker, pers. obs.). Similarly, the specimen from Iriomote Island was collected from a burrow on an intertidal sand-mud flat with some sea grass, however, without a clear evidence for callianassid association.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,204 @@
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<mods:identifier id="0B288BE79ACFD41FB11F5EC342E64B1E" type="DOI">10.11646/zootaxa.3974.3.3</mods:identifier>
<mods:identifier id="E376181195B6D3F571AED2827E4D2B9E" type="GBIF-Dataset">53679d76-ae41-4651-a3fc-b6823993d5e9</mods:identifier>
<mods:identifier id="A089B5B75F1BCB0E7FA08BE26D1F11DE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="09376BA844A5DF0989E7FE439BD1489D" type="Zenodo-Dep">231780</mods:identifier>
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</mods:mods>
<treatment id="F911475EFFFC1100EBF8FD09FC6C67FC" ID-DOI="http://doi.org/10.5281/zenodo.5073331" ID-GBIF-Taxon="119631471" ID-Zenodo-Dep="5073331" LSID="urn:lsid:plazi:treatment:F911475EFFFC1100EBF8FD09FC6C67FC" httpUri="http://treatment.plazi.org/id/F911475EFFFC1100EBF8FD09FC6C67FC" lastPageNumber="346" pageId="5" pageNumber="346">
<subSubSection id="39A2A5C3FFFC1100EBF8FD09FF1A621A" pageId="5" pageNumber="346" type="nomenclature">
<paragraph id="7107F648FFFC1100EBF8FD09FDC063F5" blockId="5.[151,561,717,776]" box="[151,561,717,743]" pageId="5" pageNumber="346">
<heading id="2A4F4124FFFC1100EBF8FD09FDC063F5" bold="true" box="[151,561,717,743]" fontSize="11" level="1" pageId="5" pageNumber="346" reason="1">
<taxonomicName id="B6B88DCBFFFC1100EBF8FD09FDC063F5" ID-CoL="1866053f-8438-4dee-9c58-05214f34c5eb" authority="Komai, 2004" authorityName="Komai" authorityYear="2004" box="[151,561,717,743]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFC1100EBF8FD09FDC063F5" bold="true" box="[151,561,717,743]" pageId="5" pageNumber="346">
<emphasis id="43CC2A5AFFFC1100EBF8FD09FE7D63F5" bold="true" box="[151,396,717,743]" italics="true" pageId="5" pageNumber="346">Naushonia japonica</emphasis>
<bibRefCitation id="15298BB9FFFC1100EAFCFD09FDC063F5" author="Komai" box="[403,561,717,743]" pageId="5" pageNumber="346" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai, 2004</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7107F648FFFC1100EBF8FD2BFF1A621A" blockId="5.[151,561,717,776]" box="[151,235,751,776]" pageId="5" pageNumber="346">
<figureCitation id="E983EACDFFFC1100EBF8FD2BFF24621A" box="[151,213,751,776]" captionStart="FIGURE 1. A" captionStartId="4.[151,250,1425,1447]" captionTargetBox="[258,1328,193,1403]" captionTargetId="figure@4.[258,1329,193,1403]" captionTargetPageId="4" captionText="FIGURE 1. A, Naushonia carinata Dworschak, Marin &amp; Anker, 2006, female (pocl 7.1 mm) from Iriomote, Japan, RUMF-ZC 3677, living individual, dorsal view; B, Naushonia japonica Komai, 2004, female (pocl 5.3 mm) from Katsuura, Japan, CBM- ZC 12594, living individual, dorsal view." httpUri="https://zenodo.org/record/231781/files/figure.png" pageId="5" pageNumber="346">Fig. 1</figureCitation>
B
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFC1100EBF8FCFCFEE1627E" pageId="5" pageNumber="346" type="reference_group">
<paragraph id="7107F648FFFC1100EBF8FCFCFEE1627E" blockId="5.[151,1436,824,876]" pageId="5" pageNumber="346">
<treatmentCitationGroup id="51A8D166FFFC1100EBF8FCFCFEE1627E" pageId="5" pageNumber="346">
<taxonomicName id="B6B88DCBFFFC1100EBF8FCFCFDD2625C" ID-CoL="1866053f-8438-4dee-9c58-05214f34c5eb" authority="Komai 2004: 17" authorityName="Komai" authorityPageNumber="17" authorityYear="2004" box="[151,547,824,846]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFC1100EBF8FCFCFE98625C" box="[151,361,824,846]" italics="true" pageId="5" pageNumber="346">Naushonia japonica</emphasis>
<treatmentCitation id="F019D059FFFC1100EA1BFCFCFDD2625C" author="Komai" box="[372,547,824,846]" page="17" pageId="5" pageNumber="346" year="2004">
<bibRefCitation id="15298BB9FFFC1100EA1BFCFCFE0B625C" author="Komai" box="[372,506,824,846]" pageId="5" pageNumber="346" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai 2004</bibRefCitation>
: 17
</treatmentCitation>
</taxonomicName>
, figs. 1A, 24 (
<typeStatus id="AE0348EAFFFC1100E9BCFCFCFCF1625C" box="[723,768,824,846]" pageId="5" pageNumber="346">type</typeStatus>
locality: Yoshimi Beach, Shimonoseki, Yamaguchi Prefecture,
<collectingCountry id="09AFB6D8FFFC1100EBA8FC93FEF4627E" box="[199,261,855,876]" name="Japan" pageId="5" pageNumber="346">Japan</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFC1100EBF8FC5DFBB362C4" pageId="5" pageNumber="346" type="materials_examined">
<paragraph id="7107F648FFFC1100EBF8FC5DFBB362C4" blockId="5.[151,1437,921,1774]" pageId="5" pageNumber="346">
<emphasis id="43CC2A5AFFFC1100EBF8FC5DFE8B62A0" bold="true" box="[151,378,921,946]" pageId="5" pageNumber="346">Material examined</emphasis>
.
<collectingCountry id="09AFB6D8FFFC1100EAE6FC5EFE3C62A3" box="[393,461,922,945]" name="Japan" pageId="5" pageNumber="346">Japan</collectingCountry>
:
<specimenCount id="67BE3DC1FFFC1100EAB7FC5EFDCB62A0" box="[472,570,921,946]" pageId="5" pageNumber="346" type="female">1 female</specimenCount>
(pocl
<quantity id="B6405BADFFFC1100E9EBFC5DFD2E62A0" box="[644,735,921,946]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="346" unit="mm" value="5.3">5.3 mm</quantity>
), Chiba Prefecture, Katsuura, Ubara, scuba diving,
<quantity id="B6405BADFFFC1100EE48FC5DFAA462A3" box="[1319,1365,921,946]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="346" unit="m" value="5.0">5 m</quantity>
, sand, under rock, coll. S. Komai and N. Nakamura,
<date id="0506D088FFFC1100E9F5FC7AFC8762C4" box="[666,886,957,982]" pageId="5" pageNumber="346" value="2013-09-30">30 September 2013</date>
, CBM-ZC 12594.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFC1100EBA8FC25FBCF650C" pageId="5" pageNumber="346" type="description">
<paragraph id="7107F648FFFC1100EBA8FC25FDC562E8" blockId="5.[151,1437,921,1774]" box="[199,564,993,1018]" pageId="5" pageNumber="346">
<emphasis id="43CC2A5AFFFC1100EBA8FC25FEA062E8" bold="true" box="[199,337,993,1018]" pageId="5" pageNumber="346">Description</emphasis>
. See
<bibRefCitation id="15298BB9FFFC1100EAE0FC25FDC162E8" author="Komai" box="[399,560,993,1018]" pageId="5" pageNumber="346" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai (2004)</bibRefCitation>
.
</paragraph>
<paragraph id="7107F648FFFC1100EBA8FBC1FBCF650C" blockId="5.[151,1437,921,1774]" box="[199,1086,1029,1054]" pageId="5" pageNumber="346">
<emphasis id="43CC2A5AFFFC1100EBA8FBC1FE53650C" bold="true" box="[199,418,1029,1054]" pageId="5" pageNumber="346">Colouration in life</emphasis>
. Body and appendages whitish; cornea dark grey (
<figureCitation id="E983EACDFFFC1100E8B6FBC1FBE8650C" box="[985,1049,1029,1054]" captionStart="FIGURE 1. A" captionStartId="4.[151,250,1425,1447]" captionTargetBox="[258,1328,193,1403]" captionTargetId="figure@4.[258,1329,193,1403]" captionTargetPageId="4" captionText="FIGURE 1. A, Naushonia carinata Dworschak, Marin &amp; Anker, 2006, female (pocl 7.1 mm) from Iriomote, Japan, RUMF-ZC 3677, living individual, dorsal view; B, Naushonia japonica Komai, 2004, female (pocl 5.3 mm) from Katsuura, Japan, CBM- ZC 12594, living individual, dorsal view." httpUri="https://zenodo.org/record/231781/files/figure.png" pageId="5" pageNumber="346">Fig. 1</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFC1100EBA8FBEDFD4B6577" pageId="5" pageNumber="346" type="distribution">
<paragraph id="7107F648FFFC1100EBA8FBEDFB116550" blockId="5.[151,1437,921,1774]" box="[199,1248,1065,1090]" pageId="5" pageNumber="346">
<emphasis id="43CC2A5AFFFC1100EBA8FBEDFEA86550" bold="true" box="[199,345,1065,1090]" pageId="5" pageNumber="346">Distribution</emphasis>
. Northwestern Pacific:
<collectingCountry id="09AFB6D8FFFC1100E90CFBEEFD526553" box="[611,675,1066,1089]" name="Japan" pageId="5" pageNumber="346">Japan</collectingCountry>
(Yamaguchi, Chiba and possibly Mie Prefectures).
</paragraph>
<paragraph id="7107F648FFFC1100EBA8FB89FD4B6577" blockId="5.[151,1437,921,1774]" box="[199,698,1101,1126]" pageId="5" pageNumber="346">
<emphasis id="43CC2A5AFFFC1100EBA8FB89FED26574" bold="true" box="[199,291,1101,1126]" pageId="5" pageNumber="346">Habitat</emphasis>
. Shallow subtidal sand flats,
<quantity id="B6405BADFFFC1100E903FB89FD476577" box="[620,694,1101,1126]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="346" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">
5
<specimenCount id="67BE3DC1FFFC1100E9E7FB89FD476577" box="[648,694,1101,1126]" pageId="5" pageNumber="346" type="generic">7 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFC1100EBA8FBB5FC6C67FC" pageId="5" pageNumber="346" type="discussion">
<paragraph id="7107F648FFFC1100EBA8FBB5FD7B6494" blockId="5.[151,1437,921,1774]" pageId="5" pageNumber="346">
<emphasis id="43CC2A5AFFFC1100EBA8FBB5FEC56598" bold="true" box="[199,308,1137,1162]" pageId="5" pageNumber="346">Remarks</emphasis>
.
<taxonomicName id="B6B88DCBFFFC1100EA2CFBB5FDD7659B" box="[323,550,1137,1161]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFC1100EA2CFBB5FDD7659B" box="[323,550,1137,1161]" italics="true" pageId="5" pageNumber="346">Naushonia japonica</emphasis>
</taxonomicName>
is presently known only from
<collectingCountry id="09AFB6D8FFFC1100E8ECFBB6FC37659B" box="[899,966,1138,1161]" name="Japan" pageId="5" pageNumber="346">Japan</collectingCountry>
. This species was originally described on the basis of a single adult male specimen from Yoshimi, Yamaguchi Prefecture, although prior to the collection of the
<typeStatus id="AE0348EAFFFC1100EBABFB7DFED865C0" box="[196,297,1209,1234]" pageId="5" pageNumber="346" type="holotype">holotype</typeStatus>
,
<bibRefCitation id="15298BB9FFFC1100EA58FB7DFE1465C0" author="Konishi" box="[311,485,1209,1234]" pageId="5" pageNumber="346" refString="Konishi, K. (2001) First record of larvae of the rare mud shrimp Naushonia Kingsley (Crustacea: Decapoda: Laomediidae) from Asian waters. Proceedings of the Biological Society of Washington, 114, 611 - 617." type="journal article" year="2001">Konishi (2001)</bibRefCitation>
reported larvae of an unidentified species of
<taxonomicName id="B6B88DCBFFFC1100E891FB7DFB8665C3" box="[1022,1143,1209,1233]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFFC1100E891FB7DFB8665C3" box="[1022,1143,1209,1233]" italics="true" pageId="5" pageNumber="346">Naushonia</emphasis>
</taxonomicName>
from Gokasyo Bay, Mie Prefecture, which possibly also represent
<taxonomicName id="B6B88DCBFFFC1100E914FB1AFCF065E7" box="[635,769,1246,1269]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFC1100E914FB1AFCF065E7" box="[635,769,1246,1269]" italics="true" pageId="5" pageNumber="346">N. japonica</emphasis>
</taxonomicName>
(see
<bibRefCitation id="15298BB9FFFC1100E82BFB19FC2665E4" author="Komai" box="[836,983,1245,1270]" pageId="5" pageNumber="346" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai 2004</bibRefCitation>
). The herein reported female specimen from Chiba Prefecture agrees well with the male
<typeStatus id="AE0348EAFFFC1100E9D2FAC5FCD36408" box="[701,802,1281,1306]" pageId="5" pageNumber="346" type="holotype">holotype</typeStatus>
, except for some sexual characters and the possession of minute denticles on the lateral margins of the telson (three on the left side and two on the right side); no such denticles are present in the
<typeStatus id="AE0348EAFFFC1100EAB8FA8DFDCC6470" box="[471,573,1353,1378]" pageId="5" pageNumber="346" type="holotype">holotype</typeStatus>
. The first pair of pleopods of the female is uniramous and the articulation between the protopod and ramus is obsolete.
</paragraph>
<paragraph id="7107F648FFFC1100EBA8FA56FDEF674C" blockId="5.[151,1437,921,1774]" pageId="5" pageNumber="346">
Among the species characterized by the non-carinate pleomeres and rounded, unarmed pleura,
<taxonomicName id="B6B88DCBFFFC1100EF92FA56FA7064BB" box="[1277,1409,1426,1449]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFC1100EF92FA56FA7064BB" box="[1277,1409,1426,1449]" italics="true" pageId="5" pageNumber="346">N. japonica</emphasis>
</taxonomicName>
is most similar to the western Atlantic
<taxonomicName id="B6B88DCBFFFC1100E95EFA72FD1A64DF" box="[561,747,1461,1485]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="crangonoides">
<emphasis id="43CC2A5AFFFC1100E95EFA72FD1A64DF" box="[561,747,1461,1485]" italics="true" pageId="5" pageNumber="346">N. crangonoides</emphasis>
</taxonomicName>
, for instance, sharing the presence of numerous (10 or more) teeth on the lateral margin of the antennal scaphocerite. However,
<taxonomicName id="B6B88DCBFFFC1100E8E4FA1EFBFF64E3" box="[907,1038,1498,1521]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFC1100E8E4FA1EFBFF64E3" box="[907,1038,1498,1521]" italics="true" pageId="5" pageNumber="346">N. japonica</emphasis>
</taxonomicName>
may be readily distinguished from
<taxonomicName id="B6B88DCBFFFC1100EBF8FA3AFEA76707" box="[151,342,1533,1557]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="crangonoides">
<emphasis id="43CC2A5AFFFC1100EBF8FA3AFEA76707" box="[151,342,1533,1557]" italics="true" pageId="5" pageNumber="346">N. crangonoides</emphasis>
</taxonomicName>
by the cheliped dactylus distinctly broadened proximally to mid-length vs. not particularly broadened in
<taxonomicName id="B6B88DCBFFFC1100EA55F9E6FE08672B" box="[314,505,1569,1593]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="crangonoides">
<emphasis id="43CC2A5AFFFC1100EA55F9E6FE08672B" box="[314,505,1569,1593]" italics="true" pageId="5" pageNumber="346">N. crangonoides</emphasis>
</taxonomicName>
, as well as by the better-defined gastric carinae (cf.
<bibRefCitation id="15298BB9FFFC1100EF16F9E5FAC96728" author="Thompson" box="[1145,1336,1569,1594]" pageId="5" pageNumber="346" refString="Thompson, M. T. (1903) A rare thalassinid and its larva. Proceedings of the Boston Society of Natural History, 31, 1 - 21. http: // dx. doi. org / 10.1080 / 00222930308678827" type="journal article" year="1903">Thompson 1903</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFC1100EE26F9E6FEAC674C" author="Goy" pageId="5" pageNumber="346" refString="Goy, J. W. &amp; Provenzano, A. J. Jr. (1979) Juvenile morphology of the rare burrowing mud shrimp Naushonia crangonoides Kingsley, with a review of the genus Naushonia (Decapoda: Thalassinidea: Laomediidae). Proceedings of the Biological Society of Washington, 92, 339 - 359." type="journal article" year="1979">Goy &amp; Provenzano 1979</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFC1100EA06F981FDE3674C" author="Williams" box="[361,530,1605,1630]" pageId="5" pageNumber="346" refString="Williams, A. B. (1984) Shrimps, lobsters, and crabs of the Atlantic coast of the eastern United States, Maine to Florida. Smithsonian Institution Press, Washington, D. C., 550 pp." type="book" year="1984">Williams 1984</bibRefCitation>
).
</paragraph>
<paragraph id="7107F648FFFC1100EBA8F9ADFC6C67FC" blockId="5.[151,1437,921,1774]" pageId="5" pageNumber="346">
<bibRefCitation id="15298BB9FFFC1100EBA8F9ADFE946790" author="Komai" box="[199,357,1641,1666]" pageId="5" pageNumber="346" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai (2004)</bibRefCitation>
did not fully examine the gill formula of the
<typeStatus id="AE0348EAFFFC1100E830F9ADFC306790" box="[863,961,1641,1666]" pageId="5" pageNumber="346" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B6B88DCBFFFC1100E88AF9AEFB996793" box="[997,1128,1642,1665]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFC1100E88AF9AEFB996793" box="[997,1128,1642,1665]" italics="true" pageId="5" pageNumber="346">N. japonica</emphasis>
</taxonomicName>
, in order not to damage the
<typeStatus id="AE0348EAFFFC1100EBF8F949FF0D67B4" box="[151,252,1677,1702]" pageId="5" pageNumber="346" type="holotype">holotype</typeStatus>
, the only specimen available at that time. The more detailed examination of the present specimen revealed that
<taxonomicName id="B6B88DCBFFFC1100EBA5F976FEBF67DB" box="[202,334,1714,1737]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFFC1100EBA5F976FEBF67DB" box="[202,334,1714,1737]" italics="true" pageId="5" pageNumber="346">N. japonica</emphasis>
</taxonomicName>
has a gill formula typical for the genus, as summarized in
<tableCitation id="3C3AC3F3FFFC1100E898F975FBBF67D8" box="[1015,1102,1713,1738]" captionStart="TABLE 1" captionStartId="2.[151,239,254,277]" captionText="TABLE 1. Gill and exopod formula of the six species of Naushonia treated in this study. [r—reduced; * setobranchs present on the fourth pereopod in N. carinata, N. japonica, N. lactoalbida, N. portoricensis, N. serratipalma, apparently absent in N. draconis]." pageId="5" pageNumber="346">Table 1</tableCitation>
. The arthrobranch associated with the first maxilliped is well developed and has multiple lamellae.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="9694713C07A642568D2FB96C0F05D329">Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key</mods:title>
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<mods:namePart id="CBE35C2C142E6885D3E27169078F408B">Komai, Tomoyuki</mods:namePart>
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<heading id="2A4F4124FFFC1100EBF8F8F2FD9E6642" bold="true" box="[151,623,1846,1872]" fontSize="11" level="1" pageId="5" pageNumber="346" reason="1">
<taxonomicName id="B6B88DCBFFFC1100EBF8F8F2FD9E6642" ID-CoL="a744ff5b-c6dc-4783-a46e-5f8a7b90c42e" authority="Berggren, 1992" authorityName="Berggren" authorityYear="1992" box="[151,623,1846,1872]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="lactoalbida">
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<emphasis id="43CC2A5AFFFC1100EBF8F8F2FE5B6642" bold="true" box="[151,426,1846,1872]" italics="true" pageId="5" pageNumber="346">Naushonia lactoalbida</emphasis>
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</emphasis>
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</paragraph>
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<figureCitation id="E983EACDFFFC1100EBF8F89DFF286660" box="[151,217,1881,1906]" captionStart="FIGURE 2" captionStartId="6.[151,250,1884,1906]" captionTargetBox="[150,1436,429,1810]" captionTargetId="figure@6.[132,1456,401,1863]" captionTargetPageId="6" captionText="FIGURE 2. Naushonia lactoalbida Berggren, 1992. A, female (pocl 5.3 mm) from Madang, Papua New Guinea, MNHN-IU- 2013 - 0627, living individual, dorsal view; B, female (pocl 9.3 mm) from Okinawa, Japan, CBM-ZC 12595, carapace, dorsal view. Scale bar for B: 2 mm." httpUri="https://zenodo.org/record/231782/files/figure.png" pageId="5" pageNumber="346">Fig. 2</figureCitation>
</paragraph>
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<taxonomicName id="B6B88DCBFFFC1100EBF8F865FDA566A5" ID-CoL="a744ff5b-c6dc-4783-a46e-5f8a7b90c42e" authority="Berggren 1992: 514" authorityName="Berggren" authorityPageNumber="514" authorityYear="1992" box="[151,596,1953,1975]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFFC1100EBF8F865FE7166A5" box="[151,384,1953,1975]" italics="true" pageId="5" pageNumber="346">Naushonia lactoalbida</emphasis>
<treatmentCitation id="F019D059FFFC1100EAE8F866FDA566A5" author="Berggren" box="[391,596,1953,1975]" page="514" pageId="5" pageNumber="346" year="1992">
<bibRefCitation id="15298BB9FFFC1100EAE8F866FDD266A5" author="Berggren" box="[391,547,1953,1975]" pageId="5" pageNumber="346" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. http: // dx. doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren 1992</bibRefCitation>
: 514
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</taxonomicName>
(
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locality: Inhaca Island,
<collectingCountry id="09AFB6D8FFFC1100E8E6F865FBE266A5" box="[905,1043,1953,1975]" name="Mozambique" pageId="5" pageNumber="346">Mozambique</collectingCountry>
), 522 (key), figs. 16.—
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<bibRefCitation id="15298BB9FFFC1100EE64F865FF0F66C4" author="Alvarez" pageId="5" pageNumber="346" refString="Alvarez, F., Villalobos, J. L. &amp; Iliffe, T. M. (2000) Naushonia manningi, new species (Decapoda: Thalassinidea: Laomediidae), from Acklins Island, Bahamas. Journal of Crustacean Biology, 20, 192 - 198." type="journal article" year="2000">
Alvarez
<emphasis id="43CC2A5AFFFC1100EE07F866FA6D66A5" box="[1384,1436,1953,1975]" italics="true" pageId="5" pageNumber="346">et al.</emphasis>
2000
</bibRefCitation>
: 198
</treatmentCitation>
(key).—
<treatmentCitation id="F019D059FFFC1100EAE9F804FDC066C4" author="Komai" box="[390,561,1984,2006]" page="22" pageId="5" pageNumber="346" year="2004">
<bibRefCitation id="15298BB9FFFC1100EAE9F804FDFD66C4" author="Komai" box="[390,524,1984,2006]" pageId="5" pageNumber="346" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai 2004</bibRefCitation>
: 22
</treatmentCitation>
, figs. 1B, 5, 6 (Iriomote Island, Ryukyu Islands,
<collectingCountry id="09AFB6D8FFFC1100EF4BF804FBAE66C4" box="[1060,1119,1984,2006]" name="Japan" pageId="5" pageNumber="346">Japan</collectingCountry>
).—
<treatmentCitation id="F019D059FFFC1100EFECF804FA9F66C4" ID-DOI="http://doi.org/10.5281/zenodo.3508258" ID-Zenodo-Dep="3508258" author="Anker" box="[1155,1390,1984,2006]" httpUri="http://treatment.plazi.org/id/AD437D0CF154AD5D94CE12E8F70419B4" page="333" pageId="5" pageNumber="346" year="2015">
<bibRefCitation id="15298BB9FFFC1100EFECF804FACD66C4" author="Anker" box="[1155,1340,1984,2006]" pageId="5" pageNumber="346" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans from Lombok, Indonesia. Part I. Alpheidae, Laomediidae, Strahlaxiidae. Zootaxa, 3911 (3), 301 - 342. http: // dx. doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker
<emphasis id="43CC2A5AFFFC1100EFA3F805FAF166C4" box="[1228,1280,1984,2006]" italics="true" pageId="5" pageNumber="346">et al.</emphasis>
2015
</bibRefCitation>
: 333
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, fig. 24B (Lombok,
<collectingCountry id="09AFB6D8FFFC1100EA0DF81AFE3866E6" box="[354,457,2014,2036]" name="Indonesia" pageId="5" pageNumber="346">Indonesia</collectingCountry>
).
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</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFF1103EBF8FF53FC0C600F" pageId="6" pageNumber="347" type="materials_examined">
<paragraph id="7107F648FFFF1103EBF8FF53FC0C600F" blockId="6.[151,1437,151,392]" pageId="6" pageNumber="347">
<emphasis id="43CC2A5AFFFF1103EBF8FF53FE8861A2" bold="true" box="[151,377,151,176]" pageId="6" pageNumber="347">Material examined</emphasis>
.
<collectingCountry id="09AFB6D8FFFF1103EAE8FF5CFE3A61BD" box="[391,459,152,175]" name="Japan" pageId="6" pageNumber="347">Japan</collectingCountry>
:
<specimenCount id="67BE3DC1FFFF1103EABAFF5CFDC761A2" box="[469,566,151,176]" pageId="6" pageNumber="347" type="female">1 female</specimenCount>
(pocl
<quantity id="B6405BADFFFF1103E911FF5CFD2661A2" box="[638,727,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="6" pageNumber="347" unit="mm" value="9.3">9.3 mm</quantity>
), Ryukyu Islands, Okinawa Island, Nago, Kyoda, scuba diving,
<quantity id="B6405BADFFFF1103EBF8FF78FF3961C6" box="[151,200,188,213]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="347" unit="m" value="5.0">5 m</quantity>
, coral rubble, coll. N. Shirakawa,
<date id="0506D088FFFF1103E90FFF78FCB261C6" box="[608,835,188,213]" pageId="6" pageNumber="347" value="2012-11-10">10 November 2012</date>
, CBM-ZC 12595.
<collectingCountry id="09AFB6D8FFFF1103EF4EFF79FAF761C7" box="[1057,1286,188,213]" name="Papua New Guinea" pageId="6" pageNumber="347">Papua New Guinea</collectingCountry>
: 1 ovigerous female (pocl
<quantity id="B6405BADFFFF1103EA45FF1BFE7261EA" box="[298,387,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="347" unit="mm" value="5.3">5.3 mm</quantity>
), Madang lagoon, scuba diving, subtidal flat with coral rubble,
<quantity id="B6405BADFFFF1103EF2AFF1BFB7C61E5" box="[1093,1165,223,248]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="6" pageNumber="347" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">35 m</quantity>
, on silt sand under coral rocks, coll. A. Anker,
<date id="0506D088FFFF1103EAE1FEC0FD96600E" box="[398,615,260,285]" pageId="6" pageNumber="347" value="2012-11-16">16 November 2012</date>
[PZD-653], MNHN-IU-2013-0627.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFF1103EBA8FEE3FD53609A" pageId="6" pageNumber="347" type="description">
<paragraph id="7107F648FFFF1103EBA8FEE3FCD56052" blockId="6.[151,1437,151,392]" box="[199,804,295,320]" pageId="6" pageNumber="347">
<emphasis id="43CC2A5AFFFF1103EBA8FEE3FEA06052" bold="true" box="[199,337,295,320]" pageId="6" pageNumber="347">Description</emphasis>
. See
<bibRefCitation id="15298BB9FFFF1103EAE0FEECFDB96052" author="Berggren" box="[399,584,296,320]" pageId="6" pageNumber="347" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. http: // dx. doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren (1992)</bibRefCitation>
and
<bibRefCitation id="15298BB9FFFF1103E911FEE3FCD16052" author="Komai" box="[638,800,295,320]" pageId="6" pageNumber="347" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai (2004)</bibRefCitation>
.
</paragraph>
<paragraph id="7107F648FFFF1103EBA8FE88FD53609A" blockId="6.[151,1437,151,392]" pageId="6" pageNumber="347">
<emphasis id="43CC2A5AFFFF1103EBA8FE88FE576077" bold="true" box="[199,422,332,357]" pageId="6" pageNumber="347">Colouration in life</emphasis>
. Body and appendages whitish with some pale yellow tinge; orange ovary partly visible through semitransparent integument (
<figureCitation id="E983EACDFFFF1103E954FEABFD8F609A" box="[571,638,367,392]" captionStart="FIGURE 2" captionStartId="6.[151,250,1884,1906]" captionTargetBox="[150,1436,429,1810]" captionTargetId="figure@6.[132,1456,401,1863]" captionTargetPageId="6" captionText="FIGURE 2. Naushonia lactoalbida Berggren, 1992. A, female (pocl 5.3 mm) from Madang, Papua New Guinea, MNHN-IU- 2013 - 0627, living individual, dorsal view; B, female (pocl 9.3 mm) from Okinawa, Japan, CBM-ZC 12595, carapace, dorsal view. Scale bar for B: 2 mm." httpUri="https://zenodo.org/record/231782/files/figure.png" pageId="6" pageNumber="347">Fig. 2</figureCitation>
A).
</paragraph>
</subSubSection>
<caption id="25C7A6C0FFFF1103EBF8F898FE4B66BD" httpUri="https://zenodo.org/record/231782/files/figure.png" pageId="6" pageNumber="347" targetBox="[150,1436,429,1810]" targetPageId="6">
<paragraph id="7107F648FFFF1103EBF8F898FE4B66BD" blockId="6.[151,1436,1884,1967]" pageId="6" pageNumber="347">
<emphasis id="43CC2A5AFFFF1103EBF8F898FEFC6663" bold="true" box="[151,269,1884,1906]" pageId="6" pageNumber="347">FIGURE 2</emphasis>
.
<taxonomicName id="B6B88DCBFFFF1103EA75F898FD5B6660" authority="Berggren, 1992" authorityName="Berggren" authorityYear="1992" box="[282,682,1884,1906]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFFF1103EA75F898FDF26660" box="[282,515,1884,1906]" italics="true" pageId="6" pageNumber="347">Naushonia lactoalbida</emphasis>
Berggren, 1992
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. A, female (pocl 5.3 mm) from Madang, Papua New Guinea, MNHN-IU- 2013-0627, living individual, dorsal view; B, female (pocl 9.3 mm) from Okinawa, Japan, CBM-ZC 12595, carapace, dorsal view. Scale bar for B: 2 mm.
</paragraph>
</caption>
<subSubSection id="39A2A5C3FFFE1102EBA8FF53FBA861E1" pageId="7" pageNumber="348" type="distribution">
<paragraph id="7107F648FFFE1102EBA8FF53FB3F61C0" blockId="7.[151,1437,151,976]" pageId="7" pageNumber="348">
<emphasis id="43CC2A5AFFFE1102EBA8FF53FEA861A2" bold="true" box="[199,345,151,176]" pageId="7" pageNumber="348">Distribution</emphasis>
. Indo-West Pacific:
<collectingCountry id="09AFB6D8FFFE1102E92BFF53FD2961A2" box="[580,728,151,176]" name="Mozambique" pageId="7" pageNumber="348">Mozambique</collectingCountry>
(Inhaca Island),
<collectingCountry id="09AFB6D8FFFE1102E8F5FF5CFC2B61BD" box="[922,986,152,175]" name="Japan" pageId="7" pageNumber="348">Japan</collectingCountry>
(Ryukyu Islands),
<collectingCountry id="09AFB6D8FFFE1102EFDAFF53FAD361A2" box="[1205,1314,151,176]" name="Indonesia" pageId="7" pageNumber="348">Indonesia</collectingCountry>
(Lombok) (
<bibRefCitation id="15298BB9FFFE1102EBF1FF7DFEBB61C3" author="Berggren" box="[158,330,185,209]" pageId="7" pageNumber="348" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. http: // dx. doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren 1992</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFE1102EA3AFF7DFE1461C3" author="Komai" box="[341,485,185,210]" pageId="7" pageNumber="348" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai 2004</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFE1102EA9FFF7DFD4961C0" author="Anker" box="[496,696,185,210]" pageId="7" pageNumber="348" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans from Lombok, Indonesia. Part I. Alpheidae, Laomediidae, Strahlaxiidae. Zootaxa, 3911 (3), 301 - 342. http: // dx. doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker
<emphasis id="43CC2A5AFFFE1102E953FF7EFD8461C3" box="[572,629,185,209]" italics="true" pageId="7" pageNumber="348">et al.</emphasis>
2015
</bibRefCitation>
),
<collectingCountry id="09AFB6D8FFFE1102E9A4FF7EFC5261C0" box="[715,931,185,210]" name="Papua New Guinea" pageId="7" pageNumber="348">Papua New Guinea</collectingCountry>
(Madang) (present study).
</paragraph>
<paragraph id="7107F648FFFE1102EBA8FF1EFBA861E1" blockId="7.[151,1437,151,976]" box="[199,1113,218,243]" pageId="7" pageNumber="348">
<emphasis id="43CC2A5AFFFE1102EBA8FF1EFED261E1" bold="true" box="[199,291,218,243]" pageId="7" pageNumber="348">Habitat</emphasis>
. Intertidal and shallow subtidal to about
<quantity id="B6405BADFFFE1102E981FF1EFCEA61E0" box="[750,795,218,243]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="348" unit="m" value="5.0">5 m</quantity>
; on sand under coral rubble.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFE1102EBA8FF38FA7362C2" pageId="7" pageNumber="348" type="discussion">
<paragraph id="7107F648FFFE1102EBA8FF38FF2F60EF" blockId="7.[151,1437,151,976]" pageId="7" pageNumber="348">
<emphasis id="43CC2A5AFFFE1102EBA8FF38FEC56007" bold="true" box="[199,308,252,277]" pageId="7" pageNumber="348">Remarks</emphasis>
.
<taxonomicName id="B6B88DCBFFFE1102EA26FF38FDA06006" box="[329,593,252,276]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFFE1102EA26FF38FDA06006" box="[329,593,252,276]" italics="true" pageId="7" pageNumber="348">Naushonia lactoalbida</emphasis>
</taxonomicName>
was originally described based on three specimens, including female
<typeStatus id="AE0348EAFFFE1102EBF8FED9FF0D6024" box="[151,252,285,310]" pageId="7" pageNumber="348" type="holotype">holotype</typeStatus>
, male allotype and female
<typeStatus id="AE0348EAFFFE1102E940FEDAFD636024" box="[559,658,286,310]" pageId="7" pageNumber="348" type="paratype">paratype</typeStatus>
, all collected on tidal flats of Inhaca Island,
<collectingCountry id="09AFB6D8FFFE1102EFE0FED9FAD26024" box="[1167,1315,285,310]" name="Mozambique" pageId="7" pageNumber="348">Mozambique</collectingCountry>
(
<bibRefCitation id="15298BB9FFFE1102EE5CFEDAFF226045" author="Berggren" pageId="7" pageNumber="348" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. http: // dx. doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren 1992</bibRefCitation>
). Subsequently,
<bibRefCitation id="15298BB9FFFE1102EAE2FEFAFDDC6045" author="Komai" box="[397,557,318,343]" pageId="7" pageNumber="348" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai (2004)</bibRefCitation>
reported a single female specimen referable to this species, collected off the coast of Iriomote Island, Ryukyu Islands,
<collectingCountry id="09AFB6D8FFFE1102E9FFFEA4FD226065" box="[656,723,352,375]" name="Japan" pageId="7" pageNumber="348">Japan</collectingCountry>
, thereby considerably extending its geographic range. More recently,
<bibRefCitation id="15298BB9FFFE1102EA6EFE45FE106088" box="[257,481,385,410]" pageId="7" pageNumber="348" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans from Lombok, Indonesia. Part I. Alpheidae, Laomediidae, Strahlaxiidae. Zootaxa, 3911 (3), 301 - 342. http: // dx. doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article">
Anker
<emphasis id="43CC2A5AFFFE1102EA3EFE46FE7D608B" box="[337,396,385,409]" italics="true" pageId="7" pageNumber="348">et al.</emphasis>
(2015)
</bibRefCitation>
reported two specimens of
<taxonomicName id="B6B88DCBFFFE1102E847FE46FC3D608B" box="[808,972,385,409]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFFE1102E847FE46FC3D608B" box="[808,972,385,409]" italics="true" pageId="7" pageNumber="348">N. lactoalbida</emphasis>
</taxonomicName>
from shallow sea grass flats in western Lombok,
<collectingCountry id="09AFB6D8FFFE1102EA67FE66FE8860A9" box="[264,377,418,443]" name="Indonesia" pageId="7" pageNumber="348">Indonesia</collectingCountry>
. The ovigerous female from Madang extends the geographic range of this species further eastward, to the eastern coast of New
<collectingCountry id="09AFB6D8FFFE1102E92EFE00FD6460CF" box="[577,661,452,477]" name="Guinea-Bissau" pageId="7" pageNumber="348">Guinea</collectingCountry>
, whereas the female from Okinawa confirms its presence in southern
<collectingCountry id="09AFB6D8FFFE1102EBF8FE22FF2B60EF" box="[151,218,486,509]" name="Japan" pageId="7" pageNumber="348">Japan</collectingCountry>
.
</paragraph>
<paragraph id="7107F648FFFE1102EBA8FDC3FA89627E" blockId="7.[151,1437,151,976]" pageId="7" pageNumber="348">
Among the species characterized by the non-carinate pleomeres and rounded pleura,
<taxonomicName id="B6B88DCBFFFE1102EFE5FDC3FADD630C" box="[1162,1324,518,542]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFFE1102EFE5FDC3FADD630C" box="[1162,1324,518,542]" italics="true" pageId="7" pageNumber="348">N. lactoalbida</emphasis>
</taxonomicName>
is readily recognizable by the presence of spines on the gastric carinae of the carapace and also by the presence of several small spines on the anterolateral areas of the carapace (
<bibRefCitation id="15298BB9FFFE1102E991FD8DFC5B6373" author="Berggren" box="[766,938,585,609]" pageId="7" pageNumber="348" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. http: // dx. doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren 1992</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFE1102E8DAFD8DFBB46373" author="Komai" box="[949,1093,585,610]" pageId="7" pageNumber="348" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai 2004</bibRefCitation>
). The specimen from Okinawa differs from the previous descriptions of
<taxonomicName id="B6B88DCBFFFE1102E91DFDAFFCE46391" box="[626,789,619,643]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFFE1102E91DFDAFFCE46391" box="[626,789,619,643]" italics="true" pageId="7" pageNumber="348">N. lactoalbida</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="15298BB9FFFE1102E820FDAFFC0D6391" author="Berggren" box="[847,1020,619,643]" pageId="7" pageNumber="348" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. http: // dx. doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren 1992</bibRefCitation>
;
<bibRefCitation id="15298BB9FFFE1102EF65FDAEFB6B6391" author="Komai" box="[1034,1178,618,643]" pageId="7" pageNumber="348" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai 2004</bibRefCitation>
) in possessing several small spines or tubercles on the postcervical area of the carapace and having noticeably stronger teeth on the middorsal carina of the carapace (
<figureCitation id="E983EACDFFFE1102EA95FD69FDB363D4" box="[506,578,685,710]" captionStart="FIGURE 2" captionStartId="6.[151,250,1884,1906]" captionTargetBox="[150,1436,429,1810]" captionTargetId="figure@6.[132,1456,401,1863]" captionTargetPageId="6" captionText="FIGURE 2. Naushonia lactoalbida Berggren, 1992. A, female (pocl 5.3 mm) from Madang, Papua New Guinea, MNHN-IU- 2013 - 0627, living individual, dorsal view; B, female (pocl 9.3 mm) from Okinawa, Japan, CBM-ZC 12595, carapace, dorsal view. Scale bar for B: 2 mm." httpUri="https://zenodo.org/record/231782/files/figure.png" pageId="7" pageNumber="348">Fig. 2</figureCitation>
B). These two differences can also be seen in one of the specimens from Lombok illustrated by
<bibRefCitation id="15298BB9FFFE1102EAF7FD0AFD8263F5" box="[408,627,718,743]" pageId="7" pageNumber="348" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans from Lombok, Indonesia. Part I. Alpheidae, Laomediidae, Strahlaxiidae. Zootaxa, 3911 (3), 301 - 342. http: // dx. doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article">
Anker
<emphasis id="43CC2A5AFFFE1102EA89FD0BFDEF63F4" box="[486,542,718,742]" italics="true" pageId="7" pageNumber="348">et al.</emphasis>
(2015)
</bibRefCitation>
, but are much less marked in the specimen from Madang (
<figureCitation id="E983EACDFFFE1102EE67FD0AFABD63F5" box="[1288,1356,718,743]" captionStart="FIGURE 2" captionStartId="6.[151,250,1884,1906]" captionTargetBox="[150,1436,429,1810]" captionTargetId="figure@6.[132,1456,401,1863]" captionTargetPageId="6" captionText="FIGURE 2. Naushonia lactoalbida Berggren, 1992. A, female (pocl 5.3 mm) from Madang, Papua New Guinea, MNHN-IU- 2013 - 0627, living individual, dorsal view; B, female (pocl 9.3 mm) from Okinawa, Japan, CBM-ZC 12595, carapace, dorsal view. Scale bar for B: 2 mm." httpUri="https://zenodo.org/record/231782/files/figure.png" pageId="7" pageNumber="348">Fig. 2</figureCitation>
A). We tentatively consider these differences as a size-related intraspecific variation, because both the Okinawa and Lombok specimens are larger than the specimens reported by
<bibRefCitation id="15298BB9FFFE1102E839FCD6FBE16238" author="Berggren" box="[854,1040,786,810]" pageId="7" pageNumber="348" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. http: // dx. doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren (1992)</bibRefCitation>
and
<bibRefCitation id="15298BB9FFFE1102EF26FCD5FB1B6238" author="Komai" box="[1097,1258,785,810]" pageId="7" pageNumber="348" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai (2004)</bibRefCitation>
, or the Madang specimen. In addition, there appear to be some differences in the shape of the rostrum, which is broader and shorter in the Madang specimen (
<figureCitation id="E983EACDFFFE1102EAD6FC90FE0A627E" box="[441,507,852,877]" captionStart="FIGURE 2" captionStartId="6.[151,250,1884,1906]" captionTargetBox="[150,1436,429,1810]" captionTargetId="figure@6.[132,1456,401,1863]" captionTargetPageId="6" captionText="FIGURE 2. Naushonia lactoalbida Berggren, 1992. A, female (pocl 5.3 mm) from Madang, Papua New Guinea, MNHN-IU- 2013 - 0627, living individual, dorsal view; B, female (pocl 9.3 mm) from Okinawa, Japan, CBM-ZC 12595, carapace, dorsal view. Scale bar for B: 2 mm." httpUri="https://zenodo.org/record/231782/files/figure.png" pageId="7" pageNumber="348">Fig. 2</figureCitation>
A) compared to that of the Japanese specimens (
<figureCitation id="E983EACDFFFE1102EF76FC90FBAC627E" box="[1049,1117,852,877]" captionStart="FIGURE 2" captionStartId="6.[151,250,1884,1906]" captionTargetBox="[150,1436,429,1810]" captionTargetId="figure@6.[132,1456,401,1863]" captionTargetPageId="6" captionText="FIGURE 2. Naushonia lactoalbida Berggren, 1992. A, female (pocl 5.3 mm) from Madang, Papua New Guinea, MNHN-IU- 2013 - 0627, living individual, dorsal view; B, female (pocl 9.3 mm) from Okinawa, Japan, CBM-ZC 12595, carapace, dorsal view. Scale bar for B: 2 mm." httpUri="https://zenodo.org/record/231782/files/figure.png" pageId="7" pageNumber="348">Fig. 2</figureCitation>
B; see also
<bibRefCitation id="15298BB9FFFE1102EFB3FC90FA9D627E" author="Komai" box="[1244,1388,852,877]" pageId="7" pageNumber="348" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai 2004</bibRefCitation>
).
</paragraph>
<paragraph id="7107F648FFFE1102EBA8FCB1FA7362C2" blockId="7.[151,1437,151,976]" pageId="7" pageNumber="348">
Neither
<bibRefCitation id="15298BB9FFFE1102EA4CFCB2FE2C629C" author="Berggren" box="[291,477,886,910]" pageId="7" pageNumber="348" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. http: // dx. doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren (1992)</bibRefCitation>
nor
<bibRefCitation id="15298BB9FFFE1102E97CFCB1FD40629C" author="Komai" box="[531,689,885,910]" pageId="7" pageNumber="348" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26." type="journal article" year="2004">Komai (2004)</bibRefCitation>
mentioned the presence of an arthrobranch at the base of the first maxilliped in
<taxonomicName id="B6B88DCBFFFE1102EA52FC53FE1262BC" box="[317,483,918,942]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFFE1102EA52FC53FE1262BC" box="[317,483,918,942]" italics="true" pageId="7" pageNumber="348">N. lactoalbida</emphasis>
</taxonomicName>
. Our examination of the Okinawa specimen confirmed the presence of a single arthrobranch at the first maxilliped in
<taxonomicName id="B6B88DCBFFFE1102E92EFC7DFD1062C2" box="[577,737,952,976]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="43CC2A5AFFFE1102E92EFC7DFD1062C2" box="[577,737,952,976]" italics="true" pageId="7" pageNumber="348">N. lactoalbida</emphasis>
</taxonomicName>
, as in the other four species reported in this study (
<tableCitation id="3C3AC3F3FFFE1102EE72FC73FA8062C2" box="[1309,1393,951,976]" captionStart="TABLE 1" captionStartId="2.[151,239,254,277]" captionText="TABLE 1. Gill and exopod formula of the six species of Naushonia treated in this study. [r—reduced; * setobranchs present on the fourth pereopod in N. carinata, N. japonica, N. lactoalbida, N. portoricensis, N. serratipalma, apparently absent in N. draconis]." pageId="7" pageNumber="348">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="F911475EFFFE1108EBF8FBD8FF1B671A" ID-DOI="http://doi.org/10.5281/zenodo.5073333" ID-GBIF-Taxon="119631469" ID-Zenodo-Dep="5073333" LSID="urn:lsid:plazi:treatment:F911475EFFFE1108EBF8FBD8FF1B671A" httpUri="http://treatment.plazi.org/id/F911475EFFFE1108EBF8FBD8FF1B671A" lastPageId="13" lastPageNumber="354" pageId="7" pageNumber="348">
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<paragraph id="7107F648FFFE1102EBF8FBD8FD636524" blockId="7.[151,658,1052,1111]" box="[151,658,1052,1078]" pageId="7" pageNumber="348">
<heading id="2A4F4124FFFE1102EBF8FBD8FD636524" bold="true" box="[151,658,1052,1078]" fontSize="11" level="1" pageId="7" pageNumber="348" reason="1">
<taxonomicName id="B6B88DCBFFFE1102EBF8FBD8FD636524" ID-CoL="74fb034f-130b-4850-be89-ccf35361ac46" authority="Rathbun, 1901" authorityName="Rathbun" authorityYear="1901" box="[151,658,1052,1078]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFFE1102EBF8FBD8FD636524" bold="true" box="[151,658,1052,1078]" pageId="7" pageNumber="348">
<emphasis id="43CC2A5AFFFE1102EBF8FBD8FE316524" bold="true" box="[151,448,1052,1078]" italics="true" pageId="7" pageNumber="348">Naushonia portoricensis</emphasis>
(
<bibRefCitation id="15298BB9FFFE1102EAA0FBD8FD7B6524" author="Rathbun" box="[463,650,1052,1078]" pageId="7" pageNumber="348" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Puerto Rico. United States Fish Commission Bulletin, 20, 1 - 137." type="journal article" year="1901">Rathbun, 1901</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7107F648FFFE1102EBF8FBFAFEF16545" blockId="7.[151,658,1052,1111]" box="[151,256,1086,1111]" pageId="7" pageNumber="348">
<figureCitation id="E983EACDFFFE1102EBF8FBFAFEF16545" box="[151,256,1086,1111]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="9.[151,250,1921,1943]" captionStartId-1="10.[151,250,1877,1899]" captionStartId-2="11.[151,250,1883,1905]" captionStartId-3="12.[151,250,1374,1396]" captionTargetBox-0="[184,1400,207,1888]" captionTargetBox-1="[231,1344,208,1858]" captionTargetBox-2="[204,1339,196,1841]" captionTargetBox-3="[242,1344,193,1351]" captionTargetId-0="figure@9.[171,1412,193,1903]" captionTargetId-1="figure@10.[201,1387,193,1860]" captionTargetId-2="figure@11.[187,1373,180,1866]" captionTargetId-3="figure@12.[242,1345,193,1353]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." captionText-1="FIGURE 4. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, anterior part of carapace and cephalic appendages, dorsal view; B, left antennal scale, dorsal view (setae omitted); C, left antennal peduncle, ventral view (setae omitted); D, left cheliped, dorsal view; E, same, ischium and merus, ventral view; F, same, carpus and chela, ventral view (only ventral setae shown); G, right uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm." captionText-2="FIGURE 5. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, left thoracic appendages. A, third maxilliped, lateral view; B, same, ischium, dorsal view; C, same, exopod, dorsal view; D, second pereopod, lateral view; E, same, dactylus, mesial view; F, third pereopod, lateral view; G, same, dactylus, lateral view; H, fourth pereopod, lateral view; I, same, dactylus, lateral view; J, fifth pereopod, lateral view; K, same, propodus and dactylus, lateral view. Scale bars: 0.5 mm." captionText-3="FIGURE 6. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, living individual in dorsal and lateral views." httpUri-0="https://zenodo.org/record/231783/files/figure.png" httpUri-1="https://zenodo.org/record/231784/files/figure.png" httpUri-2="https://zenodo.org/record/231785/files/figure.png" httpUri-3="https://zenodo.org/record/231786/files/figure.png" pageId="7" pageNumber="348">Figs. 36</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFE1102EBF8FB46FB4B6400" pageId="7" pageNumber="348" type="reference_group">
<paragraph id="7107F648FFFE1102EBF8FB46FB816585" blockId="7.[151,1436,1153,1298]" box="[151,1136,1153,1175]" pageId="7" pageNumber="348">
<treatmentCitationGroup id="51A8D166FFFE1102EBF8FB46FB816585" box="[151,1136,1153,1175]" pageId="7" pageNumber="348">
<emphasis id="43CC2A5AFFFE1102EBF8FB46FEE06585" box="[151,273,1154,1175]" italics="true" pageId="7" pageNumber="348">Homariscus</emphasis>
<taxonomicName id="B6B88DCBFFFE1102EA78FB46FDAA6585" ID-CoL="74fb034f-130b-4850-be89-ccf35361ac46" authority="Rathbun 1901: 98" authorityName="Rathbun" authorityPageNumber="98" authorityYear="1901" box="[279,603,1153,1175]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFFE1102EA78FB46FE6C6585" box="[279,413,1154,1175]" italics="true" pageId="7" pageNumber="348">portoricensis</emphasis>
<treatmentCitation id="F019D059FFFE1102EACCFB45FDAA6585" author="Rathbun" box="[419,603,1153,1175]" page="98" pageId="7" pageNumber="348" year="1901">
<bibRefCitation id="15298BB9FFFE1102EACCFB45FDC46585" author="Rathbun" box="[419,565,1153,1175]" pageId="7" pageNumber="348" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Puerto Rico. United States Fish Commission Bulletin, 20, 1 - 137." type="journal article" year="1901">Rathbun 1901</bibRefCitation>
: 98
</treatmentCitation>
</taxonomicName>
, fig. 19 (
<typeStatus id="AE0348EAFFFE1102E9DAFB45FD136585" box="[693,738,1153,1175]" pageId="7" pageNumber="348">type</typeStatus>
locality: Playa de Ponce,
<collectingCountry id="09AFB6D8FFFE1102E886FB46FB946585" box="[1001,1125,1153,1175]" name="Puerto Rico" pageId="7" pageNumber="348">Puerto Rico</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="7107F648FFFE1102EBF8FB64FB4B6400" blockId="7.[151,1436,1153,1298]" pageId="7" pageNumber="348">
<treatmentCitationGroup id="51A8D166FFFE1102EBF8FB64FB4B6400" pageId="7" pageNumber="348">
<taxonomicName id="B6B88DCBFFFE1102EBF8FB64FD9365A4" ID-CoL="74fb034f-130b-4850-be89-ccf35361ac46" authority="Chace 1939: 524" authorityName="Chace" authorityPageNumber="524" authorityYear="1939" box="[151,610,1184,1206]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFFE1102EBF8FB64FE6765A4" box="[151,406,1184,1206]" italics="true" pageId="7" pageNumber="348">Naushonia portoricensis</emphasis>
:
<treatmentCitation id="F019D059FFFE1102EAC5FB64FD9365A4" author="Chace" box="[426,610,1184,1206]" page="524" pageId="7" pageNumber="348" year="1939">
<bibRefCitation id="15298BB9FFFE1102EAC5FB64FDDA65A4" author="Chace" box="[426,555,1184,1206]" pageId="7" pageNumber="348" refString="Chace, F. A. Jr. (1939) On the systematic status of the crustacean genera Naushonia, Homoriscus and Coralliocrangon. Annals and Magazine of Natural History, 3 (11), 524 - 530. http: // dx. doi. org / 10.1080 / 03745481.1939.9723632" type="journal article" year="1939">Chace 1939</bibRefCitation>
: 524
</treatmentCitation>
</taxonomicName>
530, figs. 114 (Havana,
<collectingCountry id="09AFB6D8FFFE1102E8E8FB64FC4F65A4" box="[903,958,1184,1206]" name="Cuba" pageId="7" pageNumber="348">Cuba</collectingCountry>
).—
<treatmentCitation id="F019D059FFFE1102E88DFB64FAD165A4" author="Gurney" box="[994,1312,1184,1206]" page="610" pageId="7" pageNumber="348" year="1939">
<bibRefCitation id="15298BB9FFFE1102E88DFB64FB1965A4" author="Gurney" box="[994,1256,1184,1206]" pageId="7" pageNumber="348" refString="Gurney, R. &amp; Lebour, M. V. (1939) The larvae of the decapod genus Naushonia. Annals and Magazine of Natural History, Series 11 (3), 609 - 614." type="journal article" year="1939">Gurney &amp; Lebour 1939</bibRefCitation>
: 610
</treatmentCitation>
, figs. 114 (
<collectingCountry id="09AFB6D8FFFE1102EBA1FB7AFEDE65C6" box="[206,303,1214,1236]" name="Bermuda" pageId="7" pageNumber="348">Bermuda</collectingCountry>
, larvae).—
<treatmentCitation id="F019D059FFFE1102EAF0FB7AFD2665C6" author="Goy" box="[415,727,1214,1236]" page="352" pageId="7" pageNumber="348" year="1979">
<bibRefCitation id="15298BB9FFFE1102EAF0FB7AFD5465C6" author="Goy" box="[415,677,1214,1236]" pageId="7" pageNumber="348" refString="Goy, J. W. &amp; Provenzano, A. J. Jr. (1979) Juvenile morphology of the rare burrowing mud shrimp Naushonia crangonoides Kingsley, with a review of the genus Naushonia (Decapoda: Thalassinidea: Laomediidae). Proceedings of the Biological Society of Washington, 92, 339 - 359." type="journal article" year="1979">Goy &amp; Provenzano 1979</bibRefCitation>
: 352
</treatmentCitation>
357, figs. 6B, 7B, 8B (Playa de Ponce,
<collectingCountry id="09AFB6D8FFFE1102EF12FB7BFB0B65C6" box="[1149,1274,1214,1236]" name="Puerto Rico" pageId="7" pageNumber="348">Puerto Rico</collectingCountry>
; Quintana Roo,
<collectingCountry id="09AFB6D8FFFE1102EBA8FB19FEE765E1" box="[199,278,1245,1267]" name="Mexico" pageId="7" pageNumber="348">Mexico</collectingCountry>
).—
<treatmentCitation id="F019D059FFFE1102EA55FB19FDA965E1" author="Martin" box="[314,600,1245,1267]" page="482" pageId="7" pageNumber="348" year="1982">
<bibRefCitation id="15298BB9FFFE1102EA55FB19FDD465E1" author="Martin" box="[314,549,1245,1267]" pageId="7" pageNumber="348" refString="Martin, J. W. &amp; Abele, L. G. (1982) Naushonia panamensis, new species (Decapoda: Thalassinidea: Laomediidae) from the Pacific coast of Panama, with notes on the genus. Proceedings of the Biological Society of Washington, 95, 478 - 483." type="journal article" year="1982">Martin &amp; Abele 1982</bibRefCitation>
: 482
</treatmentCitation>
(key).—
<treatmentCitation id="F019D059FFFE1102E9DDFB19FC3265E1" author="Dworschak" box="[690,963,1245,1267]" page="2" pageId="7" pageNumber="348" year="2006">
<bibRefCitation id="15298BB9FFFE1102E9DDFB19FC5865E1" author="Dworschak" box="[690,937,1245,1267]" pageId="7" pageNumber="348" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak
<emphasis id="43CC2A5AFFFE1102E85CFB1AFC9B65E1" box="[819,874,1245,1267]" italics="true" pageId="7" pageNumber="348">et al.</emphasis>
2006
</bibRefCitation>
: 2
</treatmentCitation>
(Boca Chica,
<collectingCountry id="09AFB6D8FFFE1102EF32FB19FAC465E1" box="[1117,1333,1245,1267]" name="Dominican Republic" pageId="7" pageNumber="348">Dominican Republic</collectingCountry>
).—Perez-
<treatmentCitation id="F019D059FFFE1102EBA8FB38FE8C6400" author="Gelabert" box="[199,381,1276,1298]" page="36" pageId="7" pageNumber="348" year="2008">
<bibRefCitation id="15298BB9FFFE1102EBA8FB38FEAB6400" author="Gelabert" box="[199,346,1276,1298]" pageId="7" pageNumber="348" refString="Perez-Gelabert, D. E. (2008) Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography. Zootaxa, 1831, 1 - 530." type="journal article" year="2008">Gelabert 2008</bibRefCitation>
: 36
</treatmentCitation>
(list).—
<treatmentCitation id="F019D059FFFE1102EABFFB38FD266400" author="Komai" box="[464,727,1276,1298]" page="45" pageId="7" pageNumber="348" year="2010">
<bibRefCitation id="15298BB9FFFE1102EABFFB38FD446400" author="Komai" box="[464,693,1276,1298]" pageId="7" pageNumber="348" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504, 31 - 46." type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
: 45
</treatmentCitation>
(key).—
<treatmentCitation id="F019D059FFFE1102E840FB38FC2F6400" author="Anker" box="[815,990,1276,1298]" page="758" pageId="7" pageNumber="348" year="2014">
<bibRefCitation id="15298BB9FFFE1102E840FB38FC5E6400" author="Anker" box="[815,943,1276,1298]" pageId="7" pageNumber="348" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. http: // dx. doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
: 758
</treatmentCitation>
(Table II), 760 (key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFE1102EBF8FAF8FF146485" pageId="7" pageNumber="348" type="materials_examined">
<paragraph id="7107F648FFFE1102EBF8FAF8FF146485" blockId="7.[151,1437,1340,2031]" pageId="7" pageNumber="348">
<emphasis id="43CC2A5AFFFE1102EBF8FAF8FE8B6447" bold="true" box="[151,378,1340,1365]" pageId="7" pageNumber="348">Material examined</emphasis>
. French Antilles:
<specimenCount id="67BE3DC1FFFE1102E929FAF8FD586447" box="[582,681,1340,1365]" pageId="7" pageNumber="348" type="female">1 female</specimenCount>
(pocl
<quantity id="B6405BADFFFE1102E99DFAF8FCBC6446" box="[754,845,1340,1365]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="348" unit="mm" value="3.5">3.5 mm</quantity>
),
<collectingCountry id="09AFB6D8FFFE1102E80FFAF8FC246447" box="[864,981,1340,1365]" name="Sint Martin" pageId="7" pageNumber="348">St. Martin</collectingCountry>
, Pinel Island, sta. STM-060,
<geoCoordinate id="148C908FFFFE1102EE70FAF8FA696446" box="[1311,1432,1340,1364]" direction="north" orientation="latitude" pageId="7" pageNumber="348" precision="92" value="18.171667">18°10.3N</geoCoordinate>
,
<geoCoordinate id="148C908FFFFE1102EBF8FA99FEE26464" box="[151,275,1373,1398]" direction="west" orientation="longitude" pageId="7" pageNumber="348" precision="92" value="-63.02667">63°01.6W</geoCoordinate>
,
<quantity id="B6405BADFFFE1102EA4DFA99FE686467" box="[290,409,1373,1398]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="7.5" metricValueMin="1.5" pageId="7" pageNumber="348" unit="m" value="4.5" valueMax="7.5" valueMin="1.5">1.57.5 m</quantity>
, reef, under rocks, coll. A. Anker,
<date id="0506D088FFFE1102E855FA9AFC126464" box="[826,995,1373,1398]" pageId="7" pageNumber="348" value="2012-04-24">24 April 2012</date>
, BSTM-1708,
<collectionCode id="17A96E8DFFFE1102EFFEFA9AFB026467" box="[1169,1267,1374,1397]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="7" pageNumber="348">FLMNH</collectionCode>
UF Crustacea 32500.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFFE1108EBA8FA5BFACF63DF" lastPageId="13" lastPageNumber="354" pageId="7" pageNumber="348" type="description">
<paragraph id="7107F648FFFE1102EBA8FA5BFAA5670E" blockId="7.[151,1437,1340,2031]" pageId="7" pageNumber="348">
<emphasis id="43CC2A5AFFFE1102EBA8FA5BFEA064AA" bold="true" box="[199,337,1439,1464]" pageId="7" pageNumber="348">Description</emphasis>
. Body (
<figureCitation id="E983EACDFFFE1102EAC2FA5BFE1F64AA" box="[429,494,1439,1464]" captionStart="FIGURE 3" captionStartId="9.[151,250,1921,1943]" captionTargetBox="[184,1400,207,1888]" captionTargetId="figure@9.[171,1412,193,1903]" captionTargetPageId="9" captionText="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="7" pageNumber="348">Fig. 3</figureCitation>
AD) moderately robust, not particularly broadened or slender. Rostrum (
<figureCitation id="E983EACDFFFE1102EE5CFA5BFA7164AA" box="[1331,1408,1439,1464]" captionStart="FIGURE 3" captionStartId="9.[151,250,1921,1943]" captionTargetBox="[184,1400,207,1888]" captionTargetId="figure@9.[171,1412,193,1903]" captionTargetPageId="9" captionText="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="7" pageNumber="348">Figs. 3</figureCitation>
A, 4A) strongly flattened dorsoventrally, broadly triangular in dorsal view (approximately as long as basal width), terminating in subacute apex, slightly overreaching distal margin of second article of antennular peduncle; lateral margins feebly convex, minutely denticulate, without conspicuous spines; dorsal surface shallowly concave.
</paragraph>
<paragraph id="7107F648FFFE1102EBA8F9E2FEB866BF" blockId="7.[151,1437,1340,2031]" pageId="7" pageNumber="348">
Carapace (
<figureCitation id="E983EACDFFFE1102EA2DF9E1FE63672C" box="[322,402,1573,1598]" captionStart="FIGURE 3" captionStartId="9.[151,250,1921,1943]" captionTargetBox="[184,1400,207,1888]" captionTargetId="figure@9.[171,1412,193,1903]" captionTargetPageId="9" captionText="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="7" pageNumber="348">Figs. 3</figureCitation>
A, B, E, 4A) subcylindrical, with distinct linea thalassinica extending almost over entire length; surface glabrous except for sparse short setae on dorsal surface. Postorbital spine simple, acuminate, with smooth mesial margin. Anterolateral margin deeply notched just ventral to small branchiostegal spine, margin superior to branchiostegal spine unarmed; pterygostomial angle extending as far as branchiostegal spine, acutely pointed. Gastric region with five low longitudinal carinae, median part flanked by submedian carinae forming plateau-like elevation; mid-dorsal carina, interrupted by cervical groove, extending from anterior 0.2 of carapace length to posterodorsal margin, minutely granulate; submedian carinae diverging posteriorly and minutely granulate in anterior 0.8 of carapace, curved mesially and becoming blunter at posterior 0.2 of carapace, reaching to cervical groove; lateral carinae minutely spinulose, diverging posteriorly, gradually blunter, non-spinulose posteriorly, continuous to posterior border of cervical groove. Cervical groove conspicuous, shallow, crossing slightly posterior to mid-length of carapace, continuous with depressions delimiting posterior part of gastric median plateau.
</paragraph>
<paragraph id="7107F648FFFE110DEBA8F871FE046045" blockId="7.[151,1437,1340,2031]" lastBlockId="8.[151,1437,151,1943]" lastPageId="8" lastPageNumber="349" pageId="7" pageNumber="348">
Pleon (
<figureCitation id="E983EACDFFFE1102EA78F871FEAB66DC" box="[279,346,1973,1998]" captionStart="FIGURE 3" captionStartId="9.[151,250,1921,1943]" captionTargetBox="[184,1400,207,1888]" captionTargetId="figure@9.[171,1412,193,1903]" captionTargetPageId="9" captionText="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="7" pageNumber="348">Fig. 3</figureCitation>
C, D) smooth. First pleomere with tergum divided in two sections by shallow transverse groove, posterior section with low, short mid-dorsal carina and faint transverse carina extending to junction between tergum and pleuron; pleuron rounded posteroventrally and obtusely angled on anterior margin, without conspicuous knob-like elevation at articulation to second pleomere. Second pleuron marginally unarmed, with posterior lobe less expanded than anterior lobe; third to fifth pleura rounded, subrectangular in general outline, marginally unarmed. Sixth pleomere not particularly widened posteriorly, without conspicuous tubercles or other armature on surface; tergum with W-shaped area marking uncalcified integument; posteroventral angle rounded; posterolateral process rounded.
</paragraph>
<paragraph id="7107F648FFF1110DEBA8FE9BFECD60CE" blockId="8.[151,1437,151,1943]" pageId="8" pageNumber="349">
Telson (
<figureCitation id="E983EACDFFF1110DEA49FE9BFE9F606A" box="[294,366,351,376]" captionStart="FIGURE 3" captionStartId="9.[151,250,1921,1943]" captionTargetBox="[184,1400,207,1888]" captionTargetId="figure@9.[171,1412,193,1903]" captionTargetPageId="9" captionText="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="8" pageNumber="349">Fig. 3</figureCitation>
F) about 1.3 times longer than wide, rounded posteriorly; dorsal surface slightly elevated medially in anterior part, with many spinules or minute tubercles on either side of midline; lateral margins each with 1 minute tooth located at posterior 0.4, gradually converging into rounded posterior margin fringed with long plumose setae.
</paragraph>
<paragraph id="7107F648FFF1110DEBA8FE21FECA630D" blockId="8.[151,1437,151,1943]" pageId="8" pageNumber="349">
Eyestalks (
<figureCitation id="E983EACDFFF1110DEA2DFE21FE7F60EC" box="[322,398,485,510]" captionStart="FIGURE 3" captionStartId="9.[151,250,1921,1943]" captionTargetBox="[184,1400,207,1888]" captionTargetId="figure@9.[171,1412,193,1903]" captionTargetPageId="9" captionText="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="8" pageNumber="349">Figs. 3</figureCitation>
E, 4A) short, each with small tubercle at distomesial angle; cornea darkly pigmented, visible in dorsal view.
</paragraph>
<paragraph id="7107F648FFF1110DEBA8FDE3FC0563F5" blockId="8.[151,1437,151,1943]" pageId="8" pageNumber="349">
Antennular peduncle (
<figureCitation id="E983EACDFFF1110DEAA1FDE3FDD16352" box="[462,544,551,576]" captionStart="FIGURE 3" captionStartId="9.[151,250,1921,1943]" captionTargetBox="[184,1400,207,1888]" captionTargetId="figure@9.[171,1412,193,1903]" captionTargetPageId="9" captionText="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="8" pageNumber="349">Figs. 3</figureCitation>
E, 4A) slightly falling short of distal margin of fifth segment of antennal peduncle. First segment not visible in dorsal view, nearly as long as distal 2 segments combined; dorsal surface with statolith opening closed by stiff setae, distodorsal margin strongly raised; ventral surface with distinct median ridge in proximal half and small distomesial spine. Second and third segments unarmed, latter fairly flattened dorsoventrally. Lateral flagellum longer than peduncle, composed of 13 (left) or 14 (right) articles; mesial flagellum about 0.7 length of lateral flagellum, composed of about 8 articles.
</paragraph>
<paragraph id="7107F648FFF1110DEBA8FD2BFB8862E0" blockId="8.[151,1437,151,1943]" pageId="8" pageNumber="349">
Antennal peduncle (
<figureCitation id="E983EACDFFF1110DEAC2FD2BFE0A621A" box="[429,507,751,776]" captionStart="FIGURE 3" captionStartId="9.[151,250,1921,1943]" captionTargetBox="[184,1400,207,1888]" captionTargetId="figure@9.[171,1412,193,1903]" captionTargetPageId="9" captionText="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="8" pageNumber="349">Figs. 3</figureCitation>
E, 4AC) stout. First segment with few minute denticles laterally on distoventral margin. Second segment with 1 strong distodorsal spine laterally, distolateral margin otherwise unarmed. Third segment with 1 minute subdistal spine on ventromesial margin. Fourth and fifth segments each with very small distolateral spine; fifth segment slightly wider than long. Antennal scale moderately broad, short, almost reaching distal margin of fifth segment, terminating in strong, mesially curved spine; lateral margin slightly convex, armed with 4 small spines (except for terminal spine) in distal 0.7 of its length, spines increasing in size distally; mesial margin strongly convex, with row of stiff setae; dorsal surface with low longitudinal carina. Antennal flagellum slightly shorter than body; articles with numerous short and long setae on distal margins.
</paragraph>
<paragraph id="7107F648FFF1110DEBA8FC3EFAD66501" blockId="8.[151,1437,151,1943]" box="[199,1319,1018,1043]" pageId="8" pageNumber="349">Epistomial horn moderately slender, overreaching mid-length of first article of antennular peduncle.</paragraph>
<paragraph id="7107F648FFF1110DEBA8FBD8FE3464B1" blockId="8.[151,1437,151,1943]" pageId="8" pageNumber="349">
Mouthparts not removed. Third maxilliped (
<figureCitation id="E983EACDFFF1110DE9ADFBD8FCF76527" box="[706,774,1052,1077]" captionStart="FIGURE 5" captionStartId="11.[151,250,1883,1905]" captionTargetBox="[204,1339,196,1841]" captionTargetId="figure@11.[187,1373,180,1866]" captionTargetPageId="11" captionText="FIGURE 5. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, left thoracic appendages. A, third maxilliped, lateral view; B, same, ischium, dorsal view; C, same, exopod, dorsal view; D, second pereopod, lateral view; E, same, dactylus, mesial view; F, third pereopod, lateral view; G, same, dactylus, lateral view; H, fourth pereopod, lateral view; I, same, dactylus, lateral view; J, fifth pereopod, lateral view; K, same, propodus and dactylus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231785/files/figure.png" pageId="8" pageNumber="349">Fig. 5</figureCitation>
AC) with moderately slender endopod composed of five articles. Coxa with setobranch composed of several long setae arising from small tubercle located near base of epipod; distoventral margin unarmed. Basis short, with 1 spinule on ventromesial margin. Ischium flattened dorsoventrally, slightly widened distally, distodorsal angle slightly produced; ventrolateral margin carinate, spinulose; distomesial angle strongly produced; crista dentata well developed, with row of narrowly spaced, slender spinules. Merus shorter than ischium, with 1 small distodorsal spine; lateral surface with few spinules adjacent to dorsal margin; ventral margin spinulose, with conspicuous distal spine. Carpus short, cup-shaped. Propodus slightly longer than dactylus. Dactylus slightly curved, tapering to blunt tip, with thick cluster of stiff setae along flexor surface to tip. Exopod slender, overreaching distal margin of ischium; flagellum multiarticulated. Epipod large, tapering distally, margins spinulose (spinules on dorsal margin much larger than those on ventral margin); mastigobranch subrectangular with truncate distal margin, narrowing basally; podobranch consisting of numerous, slender lamella.
</paragraph>
<paragraph id="7107F648FFF11109EBA8FA68FB90679E" blockId="8.[151,1437,151,1943]" lastBlockId="12.[151,1437,1471,2036]" lastPageId="12" lastPageNumber="353" pageId="8" pageNumber="349">
First pereopods (= chelipeds) (
<figureCitation id="E983EACDFFF1110DE943FA68FD8064D6" box="[556,625,1452,1477]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[231,1344,208,1858]" captionTargetId="figure@10.[201,1387,193,1860]" captionTargetPageId="10" captionText="FIGURE 4. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, anterior part of carapace and cephalic appendages, dorsal view; B, left antennal scale, dorsal view (setae omitted); C, left antennal peduncle, ventral view (setae omitted); D, left cheliped, dorsal view; E, same, ischium and merus, ventral view; F, same, carpus and chela, ventral view (only ventral setae shown); G, right uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231784/files/figure.png" pageId="8" pageNumber="349">Fig. 4</figureCitation>
DF) strong, subchelate, equal in size, symmetrical in shape, strongly flattened dorsoventrally. Ischium widened distally; narrow mesial surface flanked by clearly delimited, microscopically spinulose, dorsomesial and ventromesial margins; lateral margin carinate, microscopically spinulose. Merus widened distally, cross-section subtriangular; dorsal surface almost smooth; distomesial angle produced as trispinose lobe; lateral margin distinctly carinate, microscopically spinulose (spinules increasing in size distally), terminating in minute spine; mesial face narrow, flanked by sharply delimited, microscopically spinulose dorsomesial and ventromesial margins, latter continuing to distomesial lobe; ventral surface elevated along midline, ventromesial face nearly smooth. Carpus short, slightly widened distally; surfaces smooth; distodorsal margin smooth; dorsolateral margin distinctly carinate, microscopically granulate; mesial margin distinctly carinate; ventral surface distinctly carinate along midline, distoventral margin smooth. Palm elongate, subovate in general outline, rhomboidal in cross-section, about 1.6 times as long as wide, subequal in length to ischium and merus combined; dorsal surface faintly elevated along midline, with numerous, scattered, short setae, otherwise smooth; lateral margin carinate, microscopically spinulose except for some distal spinules, with row of sparse short to long setae; mesial margin (proximal to fixed finger) diverging against lateral margin, sharply carinate, microscopically spinulose, with row of long setae; ventral surface weakly elevated along midline, with scattered low granules, latter particularly numerous lateral to midline; pollex triangular, relatively strong, arising at mid-length of palm; occlusal margin strongly oblique, its distal 0.4 with row of 4 sharp teeth decreasing in size distally, proximal-most tooth prominent, much stronger than others, broader than fixed finger, proximal 0.6 without conspicuous armature, with row of stiff setae. Dactylus moderately slender, gently curved, closing completely against cutting edge, tip overlapping dorsal side of palm when closed; extensor margin with double row of long setae, proximal part not particularly expanded; flexor (occlusal) margin sharply edged.
</paragraph>
<caption id="25C7A6C0FFF0110CEBF8F845FECF66E1" httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="9" pageNumber="350" targetBox="[184,1400,207,1888]" targetPageId="9">
<paragraph id="7107F648FFF0110CEBF8F845FECF66E1" blockId="9.[151,1436,1921,2035]" pageId="9" pageNumber="350">
<emphasis id="43CC2A5AFFF0110CEBF8F845FEFF6684" bold="true" box="[151,270,1921,1943]" pageId="9" pageNumber="350">FIGURE 3</emphasis>
.
<taxonomicName id="B6B88DCBFFF0110CEA73F845FD3B6685" authority="Rathbun, 1901" authorityName="Rathbun" authorityYear="1901" box="[284,714,1921,1943]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFF0110CEA73F845FDE66685" box="[284,535,1921,1943]" italics="true" pageId="9" pageNumber="350">Naushonia portoricensis</emphasis>
(Rathbun, 1901)
</taxonomicName>
, female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for AD; 0.5 mm for E, F.
</paragraph>
</caption>
<caption id="25C7A6C0FFF3110FEBF8F891FE9166F7" httpUri="https://zenodo.org/record/231784/files/figure.png" pageId="10" pageNumber="351" targetBox="[231,1344,208,1858]" targetPageId="10">
<paragraph id="7107F648FFF3110FEBF8F891FE9166F7" blockId="10.[151,1436,1877,2021]" pageId="10" pageNumber="351">
<emphasis id="43CC2A5AFFF3110FEBF8F891FEFF6678" bold="true" box="[151,270,1877,1899]" pageId="10" pageNumber="351">FIGURE 4</emphasis>
.
<taxonomicName id="B6B88DCBFFF3110FEA73F891FD3B6679" authority="Rathbun, 1901" authorityName="Rathbun" authorityYear="1901" box="[284,714,1877,1899]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="351" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFF3110FEA73F891FDE66679" box="[284,535,1877,1899]" italics="true" pageId="10" pageNumber="351">Naushonia portoricensis</emphasis>
(Rathbun, 1901)
</taxonomicName>
, female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, anterior part of carapace and cephalic appendages, dorsal view; B, left antennal scale, dorsal view (setae omitted); C, left antennal peduncle, ventral view (setae omitted); D, left cheliped, dorsal view; E, same, ischium and merus, ventral view; F, same, carpus and chela, ventral view (only ventral setae shown); G, right uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="25C7A6C0FFF2110EEBF8F89FFD2366FE" httpUri="https://zenodo.org/record/231785/files/figure.png" pageId="11" pageNumber="352" targetBox="[204,1339,196,1841]" targetPageId="11">
<paragraph id="7107F648FFF2110EEBF8F89FFD2366FE" blockId="11.[151,1436,1883,2028]" pageId="11" pageNumber="352">
<emphasis id="43CC2A5AFFF2110EEBF8F89FFEFF6663" bold="true" box="[151,270,1883,1905]" pageId="11" pageNumber="352">FIGURE 5</emphasis>
.
<taxonomicName id="B6B88DCBFFF2110EEA73F89FFD3B6663" authority="(Rathbun, 1901)" authorityName="Rathbun" authorityYear="1901" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[284,714,1883,1905]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="352" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFF2110EEA73F89FFDE66663" box="[284,535,1883,1905]" italics="true" pageId="11" pageNumber="352">Naushonia portoricensis</emphasis>
(Rathbun, 1901)
</taxonomicName>
, female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, left thoracic appendages. A, third maxilliped, lateral view; B, same, ischium, dorsal view; C, same, exopod, dorsal view; D, second pereopod, lateral view; E, same, dactylus, mesial view; F, third pereopod, lateral view; G, same, dactylus, lateral view; H, fourth pereopod, lateral view; I, same, dactylus, lateral view; J, fifth pereopod, lateral view; K, same, propodus and dactylus, lateral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="25C7A6C0FFF51109EBF8FA9AFCFD6481" httpUri="https://zenodo.org/record/231786/files/figure.png" pageId="12" pageNumber="353" targetBox="[242,1344,193,1351]" targetPageId="12">
<paragraph id="7107F648FFF51109EBF8FA9AFCFD6481" blockId="12.[151,1435,1374,1427]" pageId="12" pageNumber="353">
<emphasis id="43CC2A5AFFF51109EBF8FA9AFEFF6461" bold="true" box="[151,270,1374,1396]" pageId="12" pageNumber="353">FIGURE 6</emphasis>
.
<taxonomicName id="B6B88DCBFFF51109EA73FA9AFD3B6466" authority="(Rathbun, 1901)" authorityName="Rathbun" authorityYear="1901" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[284,714,1374,1396]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="353" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFF51109EA73FA9AFDE66466" box="[284,535,1374,1396]" italics="true" pageId="12" pageNumber="353">Naushonia portoricensis</emphasis>
(Rathbun, 1901)
</taxonomicName>
, female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, living individual in dorsal and lateral views.
</paragraph>
</caption>
<paragraph id="7107F648FFF51109EBA8F953FCEC660E" blockId="12.[151,1437,1471,2036]" pageId="12" pageNumber="353">
Second pereopod (
<figureCitation id="E983EACDFFF51109EAF3F953FE1167A2" box="[412,480,1687,1712]" captionStart="FIGURE 5" captionStartId="11.[151,250,1883,1905]" captionTargetBox="[204,1339,196,1841]" captionTargetId="figure@11.[187,1373,180,1866]" captionTargetPageId="11" captionText="FIGURE 5. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, left thoracic appendages. A, third maxilliped, lateral view; B, same, ischium, dorsal view; C, same, exopod, dorsal view; D, second pereopod, lateral view; E, same, dactylus, mesial view; F, third pereopod, lateral view; G, same, dactylus, lateral view; H, fourth pereopod, lateral view; I, same, dactylus, lateral view; J, fifth pereopod, lateral view; K, same, propodus and dactylus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231785/files/figure.png" pageId="12" pageNumber="353">Fig. 5</figureCitation>
D, E) moderately stout, simple. Merus with sparse row of elongate setae on ventral margin. Carpus about 0.4 times as long as merus. Propodus longer than wide, subequal in length to carpus. Dactylus feebly curved, terminating in small, acute unguis; dorsal surface with dense setae forming brush-like structure; ventral margin minutely pectinate in distal half.
</paragraph>
<paragraph id="7107F648FFF51109EBA8F8E3FA6C66E6" blockId="12.[151,1437,1471,2036]" pageId="12" pageNumber="353">
Third to fifth pereopods moderately slender, decreasing in length posteriorly. Third pereopod (
<figureCitation id="E983EACDFFF51109EF87F8E3FAD86652" box="[1256,1321,1831,1856]" captionStart="FIGURE 5" captionStartId="11.[151,250,1883,1905]" captionTargetBox="[204,1339,196,1841]" captionTargetId="figure@11.[187,1373,180,1866]" captionTargetPageId="11" captionText="FIGURE 5. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, left thoracic appendages. A, third maxilliped, lateral view; B, same, ischium, dorsal view; C, same, exopod, dorsal view; D, second pereopod, lateral view; E, same, dactylus, mesial view; F, third pereopod, lateral view; G, same, dactylus, lateral view; H, fourth pereopod, lateral view; I, same, dactylus, lateral view; J, fifth pereopod, lateral view; K, same, propodus and dactylus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231785/files/figure.png" pageId="12" pageNumber="353">Fig. 5</figureCitation>
F, G) with ischium, merus and carpus unarmed, with sparse short setae on dorsal and ventral margins; merus about 5 times longer than wide; carpus about half-length of propodus; propodus with row of short stiff setae on ventral margin; dactylus about 0.6 times as long propodus, terminating in slender, basally demarcated unguis, lateral surface with row of 4 small movable spines adjacent to dorsal margin, ventral margin forming thin lamella with slender comblike spinules, few spinules proximal to mid-length longer than others, tip of spinules blunt. Fourth pereopod (
<figureCitation id="E983EACDFFF51109EE00F81FFA6C66E6" box="[1391,1437,2011,2036]" captionStart="FIGURE 5" captionStartId="11.[151,250,1883,1905]" captionTargetBox="[204,1339,196,1841]" captionTargetId="figure@11.[187,1373,180,1866]" captionTargetPageId="11" captionText="FIGURE 5. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, left thoracic appendages. A, third maxilliped, lateral view; B, same, ischium, dorsal view; C, same, exopod, dorsal view; D, second pereopod, lateral view; E, same, dactylus, mesial view; F, third pereopod, lateral view; G, same, dactylus, lateral view; H, fourth pereopod, lateral view; I, same, dactylus, lateral view; J, fifth pereopod, lateral view; K, same, propodus and dactylus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231785/files/figure.png" pageId="12" pageNumber="353">Fig.</figureCitation>
</paragraph>
<paragraph id="7107F648FFF41108EBF8FF53FDFB61EA" blockId="13.[151,1437,151,1544]" pageId="13" pageNumber="354">
5H, I) similar to third, as illustrated; dactylus with 6 movable spines on lateral surface adjacent to dorsal margin. Fifth pereopod (
<figureCitation id="E983EACDFFF41108EA3FFF78FE6561C7" box="[336,404,188,213]" captionStart="FIGURE 5" captionStartId="11.[151,250,1883,1905]" captionTargetBox="[204,1339,196,1841]" captionTargetId="figure@11.[187,1373,180,1866]" captionTargetPageId="11" captionText="FIGURE 5. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, left thoracic appendages. A, third maxilliped, lateral view; B, same, ischium, dorsal view; C, same, exopod, dorsal view; D, second pereopod, lateral view; E, same, dactylus, mesial view; F, third pereopod, lateral view; G, same, dactylus, lateral view; H, fourth pereopod, lateral view; I, same, dactylus, lateral view; J, fifth pereopod, lateral view; K, same, propodus and dactylus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231785/files/figure.png" pageId="13" pageNumber="354">Fig. 5</figureCitation>
J, K) with grooming setae on distal part of propodus; dactylus unarmed on lateral surface, ventral lamina poorly developed.
</paragraph>
<paragraph id="7107F648FFF41108EBA8FEC0FEA360BF" blockId="13.[151,1437,151,1544]" pageId="13" pageNumber="354">
Gill formula as summarized in
<tableCitation id="3C3AC3F3FFF41108E944FEC0FD71600E" box="[555,640,260,285]" captionStart="TABLE 1" captionStartId="2.[151,239,254,277]" captionText="TABLE 1. Gill and exopod formula of the six species of Naushonia treated in this study. [r—reduced; * setobranchs present on the fourth pereopod in N. carinata, N. japonica, N. lactoalbida, N. portoricensis, N. serratipalma, apparently absent in N. draconis]." pageId="13" pageNumber="354">Table 1</tableCitation>
; single arthrobranch on first maxilliped small but distinctly lamellate; epipods on first to third pereopod marginally spinulose, each with sickle-shaped mastigobranch and multi-lamellate podobranch; epipod on fourth pereopod tapering distally to rod-like prolongation; podobranch greatly reduced to rudimentary bud; mastigobranch also reduced to tiny bud-like process; setobranchs present on third maxilliped to fourth pereopod.
</paragraph>
<paragraph id="7107F648FFF41108EBA8FE73FABC60E7" blockId="13.[151,1437,151,1544]" pageId="13" pageNumber="354">First pleopod uniramous; articulation between protopod and ramus obsolete. Second to fifth pleopods biramous, rami slender, slightly unequal, exopods longer than endopods, latter without appendices internae.</paragraph>
<paragraph id="7107F648FFF41108EBA8FE3BFC4363BA" blockId="13.[151,1437,151,1544]" pageId="13" pageNumber="354">
Uropod (
<figureCitation id="E983EACDFFF41108EA5EFE3BFE88630A" box="[305,377,511,536]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[231,1344,208,1858]" captionTargetId="figure@10.[201,1387,193,1860]" captionTargetPageId="10" captionText="FIGURE 4. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, anterior part of carapace and cephalic appendages, dorsal view; B, left antennal scale, dorsal view (setae omitted); C, left antennal peduncle, ventral view (setae omitted); D, left cheliped, dorsal view; E, same, ischium and merus, ventral view; F, same, carpus and chela, ventral view (only ventral setae shown); G, right uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231784/files/figure.png" pageId="13" pageNumber="354">Fig. 4</figureCitation>
G) overreaching posterior margin of telson. Protopod bilobed, with deep V-shaped notch, posterior margin microscopically spinulose. Endopod and exopod subovate, each with minutely spinulose transverse suture. Exopod with low mid-dorsal ridge bearing some spinules; lateral margin faintly sinuous, serrated with 4 spinules and 1 posterolateral movable spinule. Endopod with low mid-dorsal ridge bearing some minute spinules; lateral margin nearly straight, with small posterolateral spine.
</paragraph>
<paragraph id="7107F648FFF41108EBA8FD70FACF63DF" blockId="13.[151,1437,151,1544]" box="[199,1342,692,717]" pageId="13" pageNumber="354">
<emphasis id="43CC2A5AFFF41108EBA8FD70FE5363DF" bold="true" box="[199,418,692,717]" pageId="13" pageNumber="354">Colouration in life</emphasis>
. Body and appendages whitish; reddish ovary visible through integument (
<figureCitation id="E983EACDFFF41108EF84FD70FAC163DF" box="[1259,1328,692,717]" captionStart="FIGURE 6" captionStartId="12.[151,250,1374,1396]" captionTargetBox="[242,1344,193,1351]" captionTargetId="figure@12.[242,1345,193,1353]" captionTargetPageId="12" captionText="FIGURE 6. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500, living individual in dorsal and lateral views." httpUri="https://zenodo.org/record/231786/files/figure.png" pageId="13" pageNumber="354">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFF41108EBA8FD13FF176292" pageId="13" pageNumber="354" type="distribution">
<paragraph id="7107F648FFF41108EBA8FD13FAB2622A" blockId="13.[151,1437,151,1544]" pageId="13" pageNumber="354">
<emphasis id="43CC2A5AFFF41108EBA8FD13FEA863E2" bold="true" box="[199,345,727,752]" pageId="13" pageNumber="354">Distribution</emphasis>
. Western Atlantic:
<collectingCountry id="09AFB6D8FFF41108E92EFD1CFD3A63E2" box="[577,715,727,752]" name="Puerto Rico" pageId="13" pageNumber="354">Puerto Rico</collectingCountry>
(Ponce),
<collectingCountry id="09AFB6D8FFF41108E82FFD13FBDF63E2" box="[832,1070,727,752]" name="Dominican Republic" pageId="13" pageNumber="354">Dominican Republic</collectingCountry>
(Boca Chica),
<collectingCountry id="09AFB6D8FFF41108EF89FD13FAD063E2" box="[1254,1313,727,752]" name="Cuba" pageId="13" pageNumber="354">Cuba</collectingCountry>
(Havana),
<collectingCountry id="09AFB6D8FFF41108EBF8FD38FF1D6207" box="[151,236,764,789]" name="Mexico" pageId="13" pageNumber="354">Mexico</collectingCountry>
(Quintana Roo) (
<bibRefCitation id="15298BB9FFF41108EAD8FD38FDAD6206" author="Rathbun" box="[439,604,764,789]" pageId="13" pageNumber="354" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Puerto Rico. United States Fish Commission Bulletin, 20, 1 - 137." type="journal article" year="1901">Rathbun 1901</bibRefCitation>
;
<bibRefCitation id="15298BB9FFF41108E905FD38FD056206" author="Chace" box="[618,756,764,789]" pageId="13" pageNumber="354" refString="Chace, F. A. Jr. (1939) On the systematic status of the crustacean genera Naushonia, Homoriscus and Coralliocrangon. Annals and Magazine of Natural History, 3 (11), 524 - 530. http: // dx. doi. org / 10.1080 / 03745481.1939.9723632" type="journal article" year="1939">Chace 1939</bibRefCitation>
;
<bibRefCitation id="15298BB9FFF41108E86DFD38FBD56206" author="Goy" box="[770,1060,764,788]" pageId="13" pageNumber="354" refString="Goy, J. W. &amp; Provenzano, A. J. Jr. (1979) Juvenile morphology of the rare burrowing mud shrimp Naushonia crangonoides Kingsley, with a review of the genus Naushonia (Decapoda: Thalassinidea: Laomediidae). Proceedings of the Biological Society of Washington, 92, 339 - 359." type="journal article" year="1979">Goy &amp; Provenzano 1979</bibRefCitation>
;
<bibRefCitation id="15298BB9FFF41108EF5EFD38FAC86207" author="Dworschak" box="[1073,1337,764,789]" pageId="13" pageNumber="354" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak
<emphasis id="43CC2A5AFFF41108EFD6FD39FB056206" box="[1209,1268,764,788]" italics="true" pageId="13" pageNumber="354">et al.</emphasis>
2006
</bibRefCitation>
), French Antilles (
<collectingCountry id="09AFB6D8FFF41108EB90FCE4FE83622A" box="[255,370,799,824]" name="Sint Martin" pageId="13" pageNumber="354">St. Martin</collectingCountry>
) (present study), possibly also in
<collectingCountry id="09AFB6D8FFF41108E983FCDBFCA2622A" box="[748,851,799,824]" name="Bermuda" pageId="13" pageNumber="354">Bermuda</collectingCountry>
(larval material of
<bibRefCitation id="15298BB9FFF41108EF45FCE4FAC7622A" author="Gurney" box="[1066,1334,799,824]" pageId="13" pageNumber="354" refString="Gurney, R. &amp; Lebour, M. V. (1939) The larvae of the decapod genus Naushonia. Annals and Magazine of Natural History, Series 11 (3), 609 - 614." type="journal article" year="1939">Gurney &amp; Lebour 1939</bibRefCitation>
).
</paragraph>
<paragraph id="7107F648FFF41108EBA8FC80FF176292" blockId="13.[151,1437,151,1544]" pageId="13" pageNumber="354">
<emphasis id="43CC2A5AFFF41108EBA8FC80FED2624F" bold="true" box="[199,291,836,861]" pageId="13" pageNumber="354">Habitat</emphasis>
. Intertidal and shallow subtidal; on sand or mixed sand-rubble bottoms, under large rocks and coral rubble.
</paragraph>
</subSubSection>
<subSubSection id="39A2A5C3FFF41108EBA8FC48FF1B671A" pageId="13" pageNumber="354" type="discussion">
<paragraph id="7107F648FFF41108EBA8FC48FE7965D7" blockId="13.[151,1437,151,1544]" pageId="13" pageNumber="354">
<emphasis id="43CC2A5AFFF41108EBA8FC48FEC562B7" bold="true" box="[199,308,908,933]" pageId="13" pageNumber="354">Remarks</emphasis>
.
<taxonomicName id="B6B88DCBFFF41108EA27FC48FD9362B6" box="[328,610,908,932]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFF41108EA27FC48FD9362B6" box="[328,610,908,932]" italics="true" pageId="13" pageNumber="354">Naushonia portoricensis</emphasis>
</taxonomicName>
was originally described on the basis of a single incomplete female specimen, without pereopods, collected at Playa de Ponce,
<collectingCountry id="09AFB6D8FFF41108E8EFFC74FBE162DA" box="[896,1040,943,968]" name="Puerto Rico" pageId="13" pageNumber="354">Puerto Rico</collectingCountry>
(
<bibRefCitation id="15298BB9FFF41108EF46FC6BFE4E62FF" author="Rathbun" pageId="13" pageNumber="354" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Puerto Rico. United States Fish Commission Bulletin, 20, 1 - 137." type="journal article" year="1901">
Rathbun 1901; as
<emphasis id="43CC2A5AFFF41108EE7BFC74FEDA62FE" italics="true" pageId="13" pageNumber="354">
Homariscus
<taxonomicName id="B6B88DCBFFF41108EBF8FC11FEDA62FE" box="[151,299,981,1004]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="portoricensis">portoricensis</taxonomicName>
</emphasis>
,
<taxonomicName id="B6B88DCBFFF41108EA52FC10FE4E62FF" box="[317,447,980,1005]" class="Malacostraca" family="Nephropidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="family">Homaridae</taxonomicName>
</bibRefCitation>
).
<bibRefCitation id="15298BB9FFF41108EABAFC10FD8362FE" author="Chace" box="[469,626,980,1005]" pageId="13" pageNumber="354" refString="Chace, F. A. Jr. (1939) On the systematic status of the crustacean genera Naushonia, Homoriscus and Coralliocrangon. Annals and Magazine of Natural History, 3 (11), 524 - 530. http: // dx. doi. org / 10.1080 / 03745481.1939.9723632" type="journal article" year="1939">Chace (1939)</bibRefCitation>
reported a complete female of
<taxonomicName id="B6B88DCBFFF41108E884FC11FB5562FE" box="[1003,1188,981,1004]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFF41108E884FC11FB5562FE" box="[1003,1188,981,1004]" italics="true" pageId="13" pageNumber="354">N. portoricensis</emphasis>
</taxonomicName>
from Havana,
<collectingCountry id="09AFB6D8FFF41108EE35FC10FA6962FF" box="[1370,1432,980,1005]" name="Cuba" pageId="13" pageNumber="354">Cuba</collectingCountry>
, providing a rather superficial description and some illustrations.
<bibRefCitation id="15298BB9FFF41108E814FC3CFB6A6502" author="Gurney" box="[891,1179,1015,1040]" pageId="13" pageNumber="354" refString="Gurney, R. &amp; Lebour, M. V. (1939) The larvae of the decapod genus Naushonia. Annals and Magazine of Natural History, Series 11 (3), 609 - 614." type="journal article" year="1939">Gurney &amp; Lebour (1939)</bibRefCitation>
identified larvae from plankton samples taken off
<collectingCountry id="09AFB6D8FFF41108EA8EFBD8FDB96527" box="[481,584,1052,1077]" name="Bermuda" pageId="13" pageNumber="354">Bermuda</collectingCountry>
as
<taxonomicName id="B6B88DCBFFF41108E918FBD9FCC06526" box="[631,817,1053,1076]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFF41108E918FBD9FCC06526" box="[631,817,1053,1076]" italics="true" pageId="13" pageNumber="354">N. portoricensis</emphasis>
</taxonomicName>
; however, this record requires confirmation.
<bibRefCitation id="15298BB9FFF41108EE25FBD8FE9D654A" author="Goy" pageId="13" pageNumber="354" refString="Goy, J. W. &amp; Provenzano, A. J. Jr. (1979) Juvenile morphology of the rare burrowing mud shrimp Naushonia crangonoides Kingsley, with a review of the genus Naushonia (Decapoda: Thalassinidea: Laomediidae). Proceedings of the Biological Society of Washington, 92, 339 - 359." type="journal article" year="1979">Goy &amp; Provenzano (1979)</bibRefCitation>
examined the female
<typeStatus id="AE0348EAFFF41108E90DFBFBFD35654A" box="[610,708,1087,1112]" pageId="13" pageNumber="354" type="holotype">holotype</typeStatus>
and two additional male specimens from Quintana Roo,
<collectingCountry id="09AFB6D8FFF41108EE2FFBFBFA69654A" box="[1344,1432,1087,1112]" name="Mexico" pageId="13" pageNumber="354">Mexico</collectingCountry>
, and provided comparative remarks and illustrations of selected body parts to show some diagnostic characters. Most recently,
<bibRefCitation id="15298BB9FFF41108EA2DFB43FDA965B2" box="[322,600,1159,1184]" pageId="13" pageNumber="354" refString="Dworschak, P., Marin, I. &amp; Anker, A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article">
Dworschak
<emphasis id="43CC2A5AFFF41108EAA5FB4DFDF565B2" box="[458,516,1160,1184]" italics="true" pageId="13" pageNumber="354">et al.</emphasis>
(2006)
</bibRefCitation>
listed, under comparative material, a female specimen from Boca Chica,
<collectingCountry id="09AFB6D8FFF41108EBF8FB68FE7265D7" box="[151,387,1196,1221]" name="Dominican Republic" pageId="13" pageNumber="354">Dominican Republic</collectingCountry>
.
</paragraph>
<paragraph id="7107F648FFF41108EBA8FB0BFC026446" blockId="13.[151,1437,151,1544]" pageId="13" pageNumber="354">
The present specimen from
<collectingCountry id="09AFB6D8FFF41108E962FB14FD7365FA" box="[525,642,1231,1256]" name="Sint Martin" pageId="13" pageNumber="354">St. Martin</collectingCountry>
generally agrees with the original description of
<taxonomicName id="B6B88DCBFFF41108EFABFB15FA8D65F5" box="[1220,1404,1232,1256]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFF41108EFABFB15FA8D65F5" box="[1220,1404,1232,1256]" italics="true" pageId="13" pageNumber="354">N. portoricensis</emphasis>
</taxonomicName>
in Rathbun (1091) and taxonomic accounts in
<bibRefCitation id="15298BB9FFF41108E9FEFB30FCDA641E" author="Chace" box="[657,811,1268,1293]" pageId="13" pageNumber="354" refString="Chace, F. A. Jr. (1939) On the systematic status of the crustacean genera Naushonia, Homoriscus and Coralliocrangon. Annals and Magazine of Natural History, 3 (11), 524 - 530. http: // dx. doi. org / 10.1080 / 03745481.1939.9723632" type="journal article" year="1939">Chace (1939)</bibRefCitation>
and
<bibRefCitation id="15298BB9FFF41108E808FB30FB6D641E" author="Goy" box="[871,1180,1268,1292]" pageId="13" pageNumber="354" refString="Goy, J. W. &amp; Provenzano, A. J. Jr. (1979) Juvenile morphology of the rare burrowing mud shrimp Naushonia crangonoides Kingsley, with a review of the genus Naushonia (Decapoda: Thalassinidea: Laomediidae). Proceedings of the Biological Society of Washington, 92, 339 - 359." type="journal article" year="1979">Goy &amp; Provenzano (1979)</bibRefCitation>
. As mentioned above, however, the illustration of the carapace in
<bibRefCitation id="15298BB9FFF41108E9E4FADCFBEC6422" author="Goy" box="[651,1053,1303,1328]" pageId="13" pageNumber="354" refString="Goy, J. W. &amp; Provenzano, A. J. Jr. (1979) Juvenile morphology of the rare burrowing mud shrimp Naushonia crangonoides Kingsley, with a review of the genus Naushonia (Decapoda: Thalassinidea: Laomediidae). Proceedings of the Biological Society of Washington, 92, 339 - 359." type="journal article" year="1979">Goy &amp; Provenzano (1979: fig. 6B)</bibRefCitation>
is not accurate in depicting bifid postorbital spines, which, as we concluded above, do not exist in
<taxonomicName id="B6B88DCBFFF41108E81CFAF8FC1D6446" box="[883,1004,1340,1364]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="genus">
<emphasis id="43CC2A5AFFF41108E81CFAF8FC1D6446" box="[883,1004,1340,1364]" italics="true" pageId="13" pageNumber="354">Naushonia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="7107F648FFF41108EBA8FA9BFF1B671A" blockId="13.[151,1437,151,1544]" pageId="13" pageNumber="354">
Morphologically,
<taxonomicName id="B6B88DCBFFF41108EAE0FAA5FDB26465" box="[399,579,1376,1400]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="43CC2A5AFFF41108EAE0FAA5FDB26465" box="[399,579,1376,1400]" italics="true" pageId="13" pageNumber="354">N. portoricensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="B6B88DCBFFF41108E87FFAA5FC39646A" box="[784,968,1376,1400]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="crangonoides">
<emphasis id="43CC2A5AFFF41108E87FFAA5FC39646A" box="[784,968,1376,1400]" italics="true" pageId="13" pageNumber="354">N. crangonoides</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFF41108E8B9FAA5FBA96465" box="[982,1112,1376,1400]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="43CC2A5AFFF41108E8B9FAA5FBA96465" box="[982,1112,1376,1400]" italics="true" pageId="13" pageNumber="354">N. japonica</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFF41108EF0AFAA5FB1B646A" box="[1125,1258,1376,1400]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="latimana">
<emphasis id="43CC2A5AFFF41108EF0AFAA5FB1B646A" box="[1125,1258,1376,1400]" italics="true" pageId="13" pageNumber="354">N. latimana</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFF41108EF98FAA5FA646465" box="[1271,1429,1376,1400]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="macginitiei">
<emphasis id="43CC2A5AFFF41108EF98FAA5FA646465" box="[1271,1429,1376,1400]" italics="true" pageId="13" pageNumber="354">N. macginitiei</emphasis>
</taxonomicName>
,
<taxonomicName id="B6B88DCBFFF41108EBF8FA41FEFF648E" box="[151,270,1413,1436]" class="Malacostraca" family="Laomediidae" genus="Coralliocrangon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="43CC2A5AFFF41108EBF8FA41FEFF648E" box="[151,270,1413,1436]" italics="true" pageId="13" pageNumber="354">N. perrieri</emphasis>
</taxonomicName>
, and
<taxonomicName id="B6B88DCBFFF41108EA25FA41FE0A648E" box="[330,507,1412,1436]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="354" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="43CC2A5AFFF41108EA25FA41FE0A648E" box="[330,507,1412,1436]" italics="true" pageId="13" pageNumber="354">N. serratipalma</emphasis>
</taxonomicName>
, being distinguishable from all of them by a strongly incurved terminal tooth of the antennal scaphocerite (
<figureCitation id="E983EACDFFF41108EAF7FA63FE2D64D2" box="[408,476,1447,1472]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[231,1344,208,1858]" captionTargetId="figure@10.[201,1387,193,1860]" captionTargetPageId="10" captionText="FIGURE 4. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, anterior part of carapace and cephalic appendages, dorsal view; B, left antennal scale, dorsal view (setae omitted); C, left antennal peduncle, ventral view (setae omitted); D, left cheliped, dorsal view; E, same, ischium and merus, ventral view; F, same, carpus and chela, ventral view (only ventral setae shown); G, right uropod, dorsal (perpendicular) view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/231784/files/figure.png" pageId="13" pageNumber="354">Fig. 4</figureCitation>
B) and the plateau-like elevation on the gastric region of the carapace, flanked by two submedian carinae and posteriorly defined by conspicuous depressions continuing to the cervical groove (
<figureCitation id="E983EACDFFF41108EE5BFA08FA7164F6" box="[1332,1408,1484,1509]" captionStart="FIGURE 3" captionStartId="9.[151,250,1921,1943]" captionTargetBox="[184,1400,207,1888]" captionTargetId="figure@9.[171,1412,193,1903]" captionTargetPageId="9" captionText="FIGURE 3. Naushonia portoricensis (Rathbun, 1901), female (pocl 3.5 mm) from St. Martin, French Antilles, FLMNH UF Crustacea 32500. A, carapace, dorsal view; B, same, lateral view; C, pleon, dorsal view; D, pleon, telson and left uropod, lateral view; E, anterior part of carapace and left cephalic appendages, lateral view; F, telson, dorsal view. Scale bars: 1 mm for A D; 0.5 mm for E, F." httpUri="https://zenodo.org/record/231783/files/figure.png" pageId="13" pageNumber="354">Figs. 3</figureCitation>
A, B, 4A).
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="F9115369534575D2BC663602E4CB09D7" ID-DOI="http://doi.org/10.5281/zenodo.6290881" ID-GBIF-Taxon="100129428" ID-Zenodo-Dep="6290881" LSID="urn:lsid:plazi:treatment:F9115369534575D2BC663602E4CB09D7" httpUri="http://treatment.plazi.org/id/F9115369534575D2BC663602E4CB09D7" lastPageNumber="71" pageNumber="71">
<subSubSection id="DCD44B46348D40DDB3E7617CC237A4FF" type="nomenclature">
<paragraph id="4AF78F6F59C79EDFD30834C3B5FCF4D7" pageNumber="71">
<taxonomicName id="0E060C5C1DA48139A555FF62C86B4711" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145012" authority="Forel, 1890" authorityName="Forel" authorityYear="1890" class="Hexapoda" family="Formicidae" genus="Triglyphothrix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Triglyphothrix mucidus Forel" lsidName-HNS="Triglyphothrix mucidus Forel" order="Hymenoptera" pageNumber="71" phylum="Arthropoda" rank="species" species="mucidus">Triglyphothrix mucidus</taxonomicName>
<taxonomicNameLabel id="A339D7FC90DC0F8EAD1E34BEED49DBB1" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="13A96B852857E45DEB630E45D576E394" type="description">
<paragraph id="0A2F3137C1C2E54F370617B5FEF65EC9" pageNumber="71">
— [[ worker ]]. — Long. 3,4 mill. — Tres voisin de
<taxonomicName id="A6031E7C46EFE353F1579119FC8D6964" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145010" authority="Forel, 1890" authorityName="Forel" authorityYear="1890" class="Hexapoda" family="Formicidae" genus="Triglyphothrix" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Triglyphothrix gabonensis André" lsidName-HNS="Triglyphothrix gabonensis André" order="Hymenoptera" pageNumber="71" phylum="Arthropoda" rank="species" species="gabonensis">gabonensis</taxonomicName>
Andre, dont il differe comme suit: Mandibules assez densement striees-ridees, sauf a l'extremite. Epistome bien plus largement echancre au milieu de son bord anterieur. Tete bien plus distinctement concave derriere, avec les angles posterieurs moins arrondis. Le scrobe pour les scapes est plus profond, plus distinctement borde derriere, formant de cote une faible depression pour les funicules. Les yeux sont ovales et convexes, ne formant pas de pointe en avant et en bas. Thorax plus large, moins convexe transversalement, surtout le pronotum. Epines a peine aussi longues que leur intervalle (plus longue que lui chez le
<taxonomicName id="A7C46CF7FAD44DFC4442DB30AED9BC9D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145010" authority="Forel, 1890" authorityName="Forel" authorityYear="1890" class="Hexapoda" family="Formicidae" genus="Triglyphothrix" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Triglyphothrix gabonensis André" lsidName-HNS="Triglyphothrix gabonensis André" order="Hymenoptera" pageNumber="71" phylum="Arthropoda" rank="species" species="gabonensis">gabonensis</taxonomicName>
). Petiole anterieur du premier n oe ud un peu plus court que le n oe ud (plus long chez le
<taxonomicName id="AC647CCAA1391486AE5B9EC95B8E0D26" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145010" authority="Forel, 1890" authorityName="Forel" authorityYear="1890" class="Hexapoda" family="Formicidae" genus="Triglyphothrix" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Triglyphothrix gabonensis André" lsidName-HNS="Triglyphothrix gabonensis André" order="Hymenoptera" pageNumber="71" phylum="Arthropoda" rank="species" species="gabonensis">gabonensis</taxonomicName>
).
</paragraph>
<paragraph id="78ADEEB7EB9DD3196EFDF39E033E7640" pageNumber="71">Tete, thorax et pedicule mats, assez grossierement et densement rides-reticules longitudinalement et en outre finement reticules. Abdomen et membres lisses et luisants avec une ponctuation fine et espacee.</paragraph>
<paragraph id="1079628BD92CC8B5DDD061157C403DA5" pageNumber="71">
Tout le corps, les pattes et les antennes densement recouverts d'une longe pilosite blanchatre, laineuse et polyfide qui leur donne l'air d'etre couverts de moisissure. Cette pilosite est bien plus longue, bien plus dense et bien plus polyfide que chez le
<taxonomicName id="4DC0EB813FCD6E60E8A0DF7F451CAB27" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145010" authority="Forel, 1890" authorityName="Forel" authorityYear="1890" class="Hexapoda" family="Formicidae" genus="Triglyphothrix" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Triglyphothrix gabonensis André" lsidName-HNS="Triglyphothrix gabonensis André" order="Hymenoptera" pageNumber="71" phylum="Arthropoda" rank="species" species="gabonensis">gabonensis</taxonomicName>
. Un seul poil peut se diviser en six, huit et meme plus de branches, partout, meme sur les tarses; sur le corps, chaque poil forme une meche polyfide.
</paragraph>
<paragraph id="94789F97EB3A647AF488DEC41B984902" pageNumber="71">D'un brun noiratre; pedicule plus clair. Pattes, antennes, mandibules, epistome et aretes frontales roussatres.</paragraph>
<paragraph id="804DE6D6FF36AB4C9F6F978BA55121EC" pageNumber="71">
Sankuru, Kassai, Congo belge (Luja). Cette curieuse espece est bien distincte de
<taxonomicName id="A9236F9289D15BCF1EDE2BCDB6EED1C9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145010" authority="Forel, 1890" authorityName="Forel" authorityYear="1890" class="Hexapoda" family="Formicidae" genus="Triglyphothrix" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Triglyphothrix gabonensis André" lsidName-HNS="Triglyphothrix gabonensis André" order="Hymenoptera" pageNumber="71" phylum="Arthropoda" rank="species" species="gabonensis">gabonensis</taxonomicName>
Andre, sa voisine, par sa forte sculpture mate, par sa pilosite dense et polyfide qui surencherit sur celle de toutes les especes du genre, enfin par la forme de la tete, du scrobe, etc.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="92D6A34146E08C253C619E647D126DCF" ID-CoL="63CYY" authority="Lesson 1842" authorityName="Lesson" authorityYear="1842" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="genus">
<emphasis id="BC6DEB7B9674DF89630282060946C173" italics="true" pageNumber="441">Mops</emphasis>
Lesson 1842
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="81A64F0DB8888444ADAE3A1E586CB4E1" pageNumber="441" type="reference_group">
<paragraph id="B70BDCF78C611FDD25A7C1E378ECA5A5" pageNumber="441">
<treatmentCitation id="BEDF9DA783CD2856BBAF307B7994E302" author="Lesson" authority="Lesson 1842" authorityName="Lesson" authorityYear="1842" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" page="18" pageNumber="441" phylum="Chordata" rank="genus" title="Mops" volumeTitle="Nouv. Tabl. Regn. Anim. Mammiferes" year="1842">
<taxonomicName id="E112F77CA2F9A6E88522E4FD2D3E4813" ID-CoL="63CYY" authority="Lesson 1842" authorityName="Lesson" authorityYear="1842" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="genus">
<emphasis id="52E7816E3026CE1A9242242F8A10F4EF" italics="true" pageNumber="441">Mops</emphasis>
Lesson 1842
</taxonomicName>
,
<bibCitation id="4C51421E6B4CD14828EFF5323DBB5F56" author="Lesson" pageNumber="441" pagination="18" title="Mops" volumeTitle="Nouv. Tabl. Regn. Anim. Mammiferes" year="1842">Nouv. Tabl. Regn. Anim. Mammiferes: 18</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F00E740AAE380EB2CA73911E0DFB37DA" pageNumber="441" type="type_taxon">
<paragraph id="75D0DCC9F99015A9345B8D5C9F58B880" pageNumber="441">
<emphasis id="0FA416D570FAF5205E203F64087A45BA" inLineHeading="true" pageNumber="441">Type Species:</emphasis>
<taxonomicName id="2B7981D3F9621057EEFD3EF0AFD167CB" authority="Lesson 1842" authorityName="Lesson" authorityYear="1842" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="indicus">
<emphasis id="838E9BE3448770322CCC71AACEFB128D" italics="true" pageNumber="441">Mops indicus</emphasis>
Lesson 1842
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="363B072ABD6C3FF04C0FC660CFE3A714" pageNumber="441" type="synonym_list">
<paragraph id="E829C400C8DD648CA0FF83A25A8B06E3" pageNumber="441">
<emphasis id="854664E3E6E1D92F5B7558BAB8CA61EA" inLineHeading="true" pageNumber="441">Synonyms:</emphasis>
<taxonomicName id="5489426C8769585052E16A28A635E4FF" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Allomops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="genus">
<emphasis id="60CF497CD893A3F9BFF2538F5B3E64BA" italics="true" pageNumber="441">Allomops</emphasis>
J. A. Allen 1917
</taxonomicName>
;
<taxonomicName id="D0C1CC079DE947A244D0DC37DAA47F3C" authority="Taylor 1934" authorityName="Taylor" authorityYear="1934" class="Mammalia" family="Molossidae" genus="Philippinopterus" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="genus">
<emphasis id="67291365A171BD2F458CBAFE80466678" italics="true" pageNumber="441">Philippinopterus</emphasis>
<bibRefCitation id="71F33E69389C52EF2E3FBD82B388460A" author="Taylor, E. H." refId="ref348092" refString="Taylor, E. H. 1934. Philippine land mammals. Monograph of the Bureau of Science (Manila), 30: 1 - 548." year="1934">Taylor 1934</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="55A7880386EFFFCB4FC8FECC091132D1" pageNumber="441" type="child_taxa">
<paragraph id="383BA76ABC6F5C69FF859F420197F4D8" pageNumber="441">
<emphasis id="3E9E5DCE91563B6D8B7AB4AD777B8735" inLineHeading="true" pageNumber="441">Species and subspecies:</emphasis>
15 species with 10 subspecies in 2 subgenera:
</paragraph>
<paragraph id="B9ED42E29D238F341764680228F751B7" pageNumber="441">
Subgenus
<taxonomicName id="88D5056E315F78C3647D16FB5FDFC4B1" authority="Lesson 1842" authorityName="Lesson" authorityYear="1842" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subGenus" subGenus="Mops">
<emphasis id="9E646645DDC283EAB606CB4DE336C55E" italics="true" pageNumber="441">Mops (Mops)</emphasis>
Lesson 1842
</taxonomicName>
</paragraph>
<paragraph id="C98D8E591924ADBEB032EA42E23EEF59" pageNumber="441">
Subgenus
<taxonomicName id="6D4DCC2233EFE6AF74BB1ED609A390FF" authority="Dollman 1911" authorityName="Dollman" authorityYear="1911" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subGenus" subGenus="Xiphonycteris">
<emphasis id="AD83CCFD8774B7893BA32A597E37AFF4" italics="true" pageNumber="441">Mops (Xiphonycteris)</emphasis>
<bibRefCitation id="7CFE4B2E0D826317F51C329D2F9A01DE" author="Dollman, G." refId="ref87523" refString="Dollman, G. 1911. On Arvicanthis abyssinicus and allied East-African species, with descriptions of four new forms. Annals and Magazine of Natural History, ser. 8, 8: 334 - 353." year="1911">Dollman 1911</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="4EFD29356E836667DC2125C0B674799A" pageNumber="441">
Species
<taxonomicName id="3B5690BB5A67994EA49E077B8EFCC8D7" authority="Peters 1852" authorityName="Peters" authorityYear="1852" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="brachypterus" subGenus="Xiphonycteris">
<emphasis id="0130D32F79BB945AC3BF87F6CA6FB6BD" italics="true" pageNumber="441">Mops (Xiphonycteris) brachypterus</emphasis>
<bibRefCitation id="CE82F3128B4B008E130F2D38E3991D4E" author="Peters, W. C. H." refId="ref282132" refString="Peters, W. C. H. 1852. Reise nach Mossambique. I. Saugethiere. Druck &amp; Verlag, Berlin, 202 pp." year="1852">Peters 1852</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="EF8D0F5FFAA94D7D4AC2D250DA93AE59" pageNumber="441">
Subspecies
<taxonomicName id="5AE7E8972FB34C402193639CCE1600C0" authority="Peters 1852" authorityName="Peters" authorityYear="1852" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="brachypterus" subGenus="Xiphonycteris" subSpecies="brachypterus">
<emphasis id="394008505E361C0BFC0C3893C82A632B" italics="true" pageNumber="441">Mops (Xiphonycteris) brachypterus</emphasis>
subsp.
<emphasis id="4CEF33CD37BF931002C85CD53A1F74CE" italics="true" pageNumber="441">brachypterus</emphasis>
<bibRefCitation id="DB0544EB85C26EA48BA99AC3FF4E39CF" author="Peters, W. C. H." refId="ref282132" refString="Peters, W. C. H. 1852. Reise nach Mossambique. I. Saugethiere. Druck &amp; Verlag, Berlin, 202 pp." year="1852">Peters 1852</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="845F2EBF20F9C763CE046C3568ACA93B" pageNumber="441">
Subspecies
<taxonomicName id="EE801BD0A45709E7FDC5E30D57456E9F" authority="Thomas 1908" authorityName="Thomas" authorityYear="1908" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="brachypterus" subGenus="Xiphonycteris" subSpecies="leonis">
<emphasis id="0B032B3E13AC8553A933641CAAF62D5C" italics="true" pageNumber="441">Mops (Xiphonycteris) brachypterus</emphasis>
subsp.
<emphasis id="11326154E8E6AD6A3A067DE7A8D8ABD5" italics="true" pageNumber="441">leonis</emphasis>
Thomas 1908
</taxonomicName>
</paragraph>
<paragraph id="815110AE9F768DE7149D5254D1001249" pageNumber="441">
Species
<taxonomicName id="2511E99399DB39FEB2A792129B371ED6" authority="(A. Smith 1833)" baseAuthorityName="A. Smith" baseAuthorityYear="1833" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="condylurus" subGenus="Mops">
<emphasis id="4C146C98201E2384334E2269202F4ACF" italics="true" pageNumber="441">Mops (Mops) condylurus</emphasis>
(A.
<bibRefCitation id="07CD7F3D830E247362D9C663C9722AB4" author="Smith, A." refId="ref330294" refString="Smith, A. 1833 - 1834. An epitome of African Zoology; or, a concise description of the objects of the animal kingdom inhabiting Africa, its islands and seas. South African Quarterly Journal, 2: 16 - 32, 49 - 64, 81 - 96, 113 - 128, 145 - 160, 169 - 192, 209 - 224, 233 - 248." year="1833">Smith 1833</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="B79DE2C4D1316C8959F2B48CB683C798" pageNumber="441">
Subspecies
<taxonomicName id="9EEF18514443902E0B6DFBFAD6AAE721" authority="A. Smith 1833" authorityName="A. Smith" authorityYear="1833" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="condylurus" subGenus="Mops" subSpecies="condylurus">
<emphasis id="4E309D85AF68D0D00723D64F7C4B01AF" italics="true" pageNumber="441">Mops (Mops) condylurus</emphasis>
subsp.
<emphasis id="28BC69EB2EB1E66A71F7AF67FFB34E82" italics="true" pageNumber="441">condylurus</emphasis>
A.
<bibRefCitation id="277BA1ABDD5AA16D2073856F6AFAB6D9" author="Smith, A." refId="ref330294" refString="Smith, A. 1833 - 1834. An epitome of African Zoology; or, a concise description of the objects of the animal kingdom inhabiting Africa, its islands and seas. South African Quarterly Journal, 2: 16 - 32, 49 - 64, 81 - 96, 113 - 128, 145 - 160, 169 - 192, 209 - 224, 233 - 248." year="1833">Smith 1833</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="D190528B53B32A313C7548D09F8A79F1" pageNumber="441">
Subspecies
<taxonomicName id="3ECEEA6E9DFD9C5C0FC5EA3B1C18FA75" authority="G. M. Allen and Loveridge 1942" authorityName="G. M. Allen and Loveridge" authorityYear="1942" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="condylurus" subGenus="Mops" subSpecies="orientis">
<emphasis id="5AF251395B5E3472582ECE1F2E9D13E6" italics="true" pageNumber="441">Mops (Mops) condylurus</emphasis>
subsp.
<emphasis id="2174533ADCD0F388B487FDB022AA0457" italics="true" pageNumber="441">orientis</emphasis>
G. M.
<bibRefCitation id="CB50FB53652A880EF7546A432C6EC023" author="Allen, G. M. &amp; A. Loveridge" refId="ref5059" refString="Allen, G. M., and A. Loveridge. 1942. Scientific results of a fourth expedition to forested areas in East and Central Africa. I. Mammals. Bulletin of the Museum of Comparative Zoology at Harvard College, 89: 147 - 214." year="1942">Allen and Loveridge 1942</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="D036EA3225F0FD868C1C5DB80457792B" pageNumber="441">
Subspecies
<taxonomicName id="102D3B5594DA7BB09B3298A7AA215296" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="condylurus" subGenus="Mops" subSpecies="osborni">
<emphasis id="E3D77D4E1FA21F6E23E41C5A66F93448" italics="true" pageNumber="441">Mops (Mops) condylurus</emphasis>
subsp.
<emphasis id="ABBB106BE44722955E068897CDC7FB29" italics="true" pageNumber="441">osborni</emphasis>
J. A. Allen 1917
</taxonomicName>
</paragraph>
<paragraph id="2D9C2B04DC428CC32C453A8F99D7735B" pageNumber="441">
Subspecies
<taxonomicName id="04B2DC1F09094E87A9C0FE8413B35D2A" authority="Sanbron 1936" authorityName="Sanbron" authorityYear="1936" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="condylurus" subGenus="Mops" subSpecies="wonderi">
<emphasis id="1511EE40E36EBCC1C470CEF452900F9F" italics="true" pageNumber="441">Mops (Mops) condylurus</emphasis>
subsp.
<emphasis id="A3E063ACFE6D3EF2621BF330648A23E7" italics="true" pageNumber="441">wonderi</emphasis>
Sanbron 1936
</taxonomicName>
</paragraph>
<paragraph id="86C023B84F3C43F5A256012842FDB6B9" pageNumber="441">
Species
<taxonomicName id="2A4DC234311CA4CC07003B829D2F7B65" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="congicus" subGenus="Mops">
<emphasis id="AD2EC3AD9E43267792FF1A7428B652E6" italics="true" pageNumber="441">Mops (Mops) congicus</emphasis>
J. A. Allen 1917
</taxonomicName>
</paragraph>
<paragraph id="0FDD88E2003D8E262304F45DE58005B2" pageNumber="441">
Species
<taxonomicName id="A43CBD9878960778F8834EEE9818ECB7" authority="Thomas 1903" authorityName="Thomas" authorityYear="1903" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="demonstrator" subGenus="Mops">
<emphasis id="F8D5B17435BD82AB4FC75B803DFD8706" italics="true" pageNumber="441">Mops (Mops) demonstrator</emphasis>
<bibRefCitation id="5625D2D274C56461FC5DFAC379E9982F" author="Thomas, O." refId="ref352582" refString="Thomas, O. 1903. Notes on Neotropical mammals of the genera Felis, Hapale, Oryzomys, Akodon, and Ctenomys, with descriptions of new species. Annals and Magazine of Natural History, ser. 7, 12: 234 - 243." year="1903">Thomas 1903</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="70A8D17F72CDE3E7C26A63BDD1EC9325" pageNumber="441">
Species
<taxonomicName id="2139B7BE78822327B140FE9F7501AE02" authority="G. M. Allen 1918" authorityName="G. M. Allen" authorityYear="1918" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="leucostigma" subGenus="Mops">
<emphasis id="825B27C971CAC2EACE09586CBA2E57A9" italics="true" pageNumber="441">Mops (Mops) leucostigma</emphasis>
G. M. Allen 1918
</taxonomicName>
</paragraph>
<paragraph id="AAFB34AE33D16D4EB02614EE2A030CB7" pageNumber="441">
Species
<taxonomicName id="C168EF1F3C1ADD0CD79D3A66B9C2B03E" authority="Sundevall 1842" authorityName="Sundevall" authorityYear="1842" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="midas" subGenus="Mops">
<emphasis id="F370C0BBD0EA47CF67CC9C2B10174B74" italics="true" pageNumber="441">Mops (Mops) midas</emphasis>
Sundevall 1842
</taxonomicName>
</paragraph>
<paragraph id="26281CB39608B26D7955D01AD7CD4387" pageNumber="441">
Subspecies
<taxonomicName id="5ED43EEC9D7EC8AF2C0A581308388AA6" authority="Sundevall 1842" authorityName="Sundevall" authorityYear="1842" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="midas" subGenus="Mops" subSpecies="midas">
<emphasis id="35A43D7C9BCF9215B6E5EAB6BE5788EF" italics="true" pageNumber="441">Mops (Mops) midas</emphasis>
subsp.
<emphasis id="8E9D1C68EF3E2058F48695E21A71A77F" italics="true" pageNumber="441">midas</emphasis>
Sundevall 1842
</taxonomicName>
</paragraph>
<paragraph id="938172ED1A0249DCBF3D6721BA3CD8DE" pageNumber="441">
Subspecies
<taxonomicName id="F391459628FB0A386A5C3B53E80E75BD" authority="A. Grandidier 1869" authorityName="A. Grandidier" authorityYear="1869" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="midas" subGenus="Mops" subSpecies="miarensis">
<emphasis id="9BD949725D6329F86061B20E65657D65" italics="true" pageNumber="441">Mops (Mops) midas</emphasis>
subsp.
<emphasis id="A17285EF5B6FF7DAB2ACBF6F47F527F4" italics="true" pageNumber="441">miarensis</emphasis>
A. Grandidier 1869
</taxonomicName>
</paragraph>
<paragraph id="90298847C71BD6479C32196B042722DE" pageNumber="441">
Species
<taxonomicName id="68D328059F68AE3CD461DBF41451EDD5" authority="(de Blainville 1840)" baseAuthorityName="de Blainville" baseAuthorityYear="1840" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="mops" subGenus="Mops">
<emphasis id="B4F736BD77AA63522FF7D911143AEF5C" italics="true" pageNumber="441">Mops (Mops) mops</emphasis>
(de Blainville 1840)
</taxonomicName>
</paragraph>
<paragraph id="7F7A07F2E5C5C6BF71898E436159AF64" pageNumber="441">
Species
<taxonomicName id="728CC2B9E290E54E7061A457A3D8D4E2" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="nanulus" subGenus="Xiphonycteris">
<emphasis id="C29808CE58BF08DB6D56ADF28B5BE46C" italics="true" pageNumber="441">Mops (Xiphonycteris) nanulus</emphasis>
J. A. Allen 1917
</taxonomicName>
</paragraph>
<paragraph id="5BA5C739635F5C4ECC6470418D3C90D4" pageNumber="441">
Species
<taxonomicName id="3B635532D0C75EBFEA6BD11C81B34CBD" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="niangarae" subGenus="Mops">
<emphasis id="95594A584E2CAFED642CC71E45FBB52C" italics="true" pageNumber="441">Mops (Mops) niangarae</emphasis>
J. A. Allen 1917
</taxonomicName>
</paragraph>
<paragraph id="4C186F01D51A54D4F95E9B9BDA7F89EE" pageNumber="441">
Species
<taxonomicName id="8D13174E56FC335A33C2524B5138CC31" authority="Cabrera and Ruxton 1926" authorityName="Cabrera and Ruxton" authorityYear="1926" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="niveiventer" subGenus="Mops">
<emphasis id="C2FADE4E785F07DFA7BF99BE7DE9CB89" italics="true" pageNumber="441">Mops (Mops) niveiventer</emphasis>
Cabrera and Ruxton 1926
</taxonomicName>
</paragraph>
<paragraph id="7484E4651D019716F0D242EBC29A4E80" pageNumber="441">
Species
<taxonomicName id="D18DB38ACB066E036FB2AD96075F2264" authority="El Rayah 1981" authorityName="El Rayah" authorityYear="1981" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="petersoni" subGenus="Xiphonycteris">
<emphasis id="4894D2D09B3BA14732BA026C866CF5E6" italics="true" pageNumber="441">Mops (Xiphonycteris) petersoni</emphasis>
El Rayah 1981
</taxonomicName>
</paragraph>
<paragraph id="E5AC42853E00C7E026B1CB98D090FDC0" pageNumber="441">
Species
<taxonomicName id="C167544B71F1458979AA0863396E3581" authority="A. Meyer 1899" authorityName="A. Meyer" authorityYear="1899" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="sarasinorum" subGenus="Mops">
<emphasis id="66393F21507C4889E4603CCB1A60C799" italics="true" pageNumber="441">Mops (Mops) sarasinorum</emphasis>
A. Meyer 1899
</taxonomicName>
</paragraph>
<paragraph id="6124ECB958B06DBD9C36EB07A575AEAA" pageNumber="441">
Subspecies
<taxonomicName id="D465CD96F08688075467CFC45F4B4AB1" authority="A. Meyer 1899" authorityName="A. Meyer" authorityYear="1899" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="sarasinorum" subGenus="Mops" subSpecies="sarasinorum">
<emphasis id="384C47685551FB88928D0C99F8B9CD4F" italics="true" pageNumber="441">Mops (Mops) sarasinorum</emphasis>
subsp.
<emphasis id="4193D3297CD5DEFF573CF172CC0987FC" italics="true" pageNumber="441">sarasinorum</emphasis>
A. Meyer 1899
</taxonomicName>
</paragraph>
<paragraph id="D40656C0A8F9EC381811AFA3536795F9" pageNumber="441">
Subspecies
<taxonomicName id="91426FF0465B8DB11E7F6B900DDF8ECA" authority="Taylor 1934" authorityName="Taylor" authorityYear="1934" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="subSpecies" species="sarasinorum" subGenus="Mops" subSpecies="lanei">
<emphasis id="E136071EDDC70B17AF8826A3D702C6AF" italics="true" pageNumber="441">Mops (Mops) sarasinorum</emphasis>
subsp.
<emphasis id="DA1EB602341FD468424330FD68310C5B" italics="true" pageNumber="441">lanei</emphasis>
<bibRefCitation id="8E646FC44F4EC1616DB2870D506C1ACD" author="Taylor, E. H." refId="ref348092" refString="Taylor, E. H. 1934. Philippine land mammals. Monograph of the Bureau of Science (Manila), 30: 1 - 548." year="1934">Taylor 1934</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="114F3BD220F45361B24CDF1B01C12D18" pageNumber="441">
Species
<taxonomicName id="A9F23C3CDE1B45E9BCAA09367C596841" authority="Dollman 1911" authorityName="Dollman" authorityYear="1911" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="spurrelli" subGenus="Xiphonycteris">
<emphasis id="9DF1755946DBF2CE28FFB96B93D2F7A9" italics="true" pageNumber="441">Mops (Xiphonycteris) spurrelli</emphasis>
<bibRefCitation id="049C20FF9879D560E6F684328D94E019" author="Dollman, G." refId="ref87523" refString="Dollman, G. 1911. On Arvicanthis abyssinicus and allied East-African species, with descriptions of four new forms. Annals and Magazine of Natural History, ser. 8, 8: 334 - 353." year="1911">Dollman 1911</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="D0F821C1AE68D8B0800E7AF64588FAF2" pageNumber="441">
Species
<taxonomicName id="6A0483D22FE596E64689D20F7D4BD446" authority="Thomas 1903" authorityName="Thomas" authorityYear="1903" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="thersites" subGenus="Xiphonycteris">
<emphasis id="F7BF50C89A4C246844195BC943B51AA0" italics="true" pageNumber="441">Mops (Xiphonycteris) thersites</emphasis>
<bibRefCitation id="047955E8ABF01AFF83683DEFF8F17958" author="Thomas, O." refId="ref352582" refString="Thomas, O. 1903. Notes on Neotropical mammals of the genera Felis, Hapale, Oryzomys, Akodon, and Ctenomys, with descriptions of new species. Annals and Magazine of Natural History, ser. 7, 12: 234 - 243." year="1903">Thomas 1903</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="6D259DCA98C12186EA69873518985B42" pageNumber="441">
Species
<taxonomicName id="5CE77B2D6CE3556766D5D03572F3B215" authority="J. A. Allen 1917" authorityName="J. A. Allen" authorityYear="1917" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" pageNumber="441" phylum="Chordata" rank="species" species="trevori" subGenus="Mops">
<emphasis id="86F7276EFAC0E3825F4DEE02E9EFFDC1" italics="true" pageNumber="441">Mops (Mops) trevori</emphasis>
J. A. Allen 1917
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="65E88B200C38C0539F4E86DDAE955CCF" pageNumber="441" type="discussion">
<paragraph id="B6664DDEAC403719AAFDD9DFF2E4FE8E" pageNumber="441">
<emphasis id="1AD47577F93D7C69642927AAC697A9E9" inLineHeading="true" pageNumber="441">Discussion:</emphasis>
Formerly included in
<taxonomicName id="3D51799E52CF1AEB34355088D571CD9D" authorityName="Rafinesque" authorityYear="1814" class="Mammalia" family="Molossidae" genus="Tadarida" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="genus">
<emphasis id="EAC59CF99339FA986C8E3D7D2D03E84B" italics="true" pageNumber="441">Tadarida</emphasis>
</taxonomicName>
, often as a subgenus, but apparently distinct; see
<bibRefCitation id="EE3E848A9DD9A1615B2849C3C47834E9" author="Freeman, P. W." refId="ref108769" refString="Freeman, P. W. 1981. A multivariate study of the family Molossidae (Mammalia: Chiroptera): Morphology, ecology, evolution. Fieldiana: Zoology, n. s., 7: 1 - 173." year="1981">Freeman (1981)</bibRefCitation>
, also see
<bibRefCitation id="C706615FB80FBBA07CFE64D7AFE856E4" author="Legendre, S." refId="ref209353" refString="Legendre, S. 1984. Etude odotologique des representants actuels du groupe Tadarida (Chiroptera, Molossidae). Implications phylogeniques, systematiques et zoogeographiques. Revue Suisse de Zoologie, 91: 399 - 442." year="1984">Legendre (1984)</bibRefCitation>
.
<bibRefCitation id="8668E0F85A05FCB3826B4D92F1050D4B" author="Dunlop, J." refId="ref89931" refString="Dunlop, J. 1999. Mops midas. Mammalian Species, 615: 1 - 4." year="1999">Dunlop (1999)</bibRefCitation>
provided a key to subgenera and species in this genus. Two subgenera are recognized,
<taxonomicName id="B3C4D0ABC6B31CA457271CE57EDC24EE" authorityName="Lesson" authorityYear="1842" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="genus">
<emphasis id="E8649CDFDA311CE3BE67995404EC89F3" italics="true" pageNumber="441">Mops</emphasis>
</taxonomicName>
and
<taxonomicName id="AAEB2A96C4A3B0C6FBA12DA409AC9D07" authorityName="Dollman" authorityYear="1911" class="Mammalia" family="Molossidae" genus="Mops" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="subGenus" subGenus="Xiphonycteris">
<emphasis id="83498D634716261765E69C2794EA440B" italics="true" pageNumber="441">Xiphonycteris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart id="614A5E45FA2111B22BD8FC3D2EB3ABBF">Vu, Dang Quoc</mods:namePart>
<mods:affiliation id="E3CF021BF2E5DA67407F5F5269DF205C">Forest Protection Department, Vietnam Administration of Forestry, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:name id="0C7ACACFB6D471B57B743B2088C48FDA" type="personal">
<mods:role id="9B1FD678974313986960BD30392E884F">
<mods:roleTerm id="97159DD37B4F48B204727A58741C924C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E9A0120B4BB23133EF17CC6F29B2AD68">Ma, Jin-Shuang</mods:namePart>
<mods:affiliation id="347CF5A726967F5A1DE46ED480F73D51">Shanghai Chenshan Plant Science Research Center, Chinese Academy of Sciences, Shanghai Chenshan Botanical Garden, 3888 Chenhua Road, Songjiang, Shanghai 201602, China; E-mail: majinshuang @ sibs. ac. cn</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5D435A9FCA28306DEBEEA203EDAE514A">text</mods:typeOfResource>
<mods:relatedItem id="D5D6ADDBE6175B30D85C407B7429BC51" type="host">
<mods:titleInfo id="E0EDBE9A2D4F7B34970E59B7EFFA485C">
<mods:title id="F17BBE881290788BB1851A0317BF6B58">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="CA984D7F13B64BB7B075005C459248EA">
<mods:date id="350B423C58CF85DEA9B5A5B6C0BB7815">2014</mods:date>
<mods:detail id="F6AC0788355835A81962A35762BDF844" type="pubDate">
<mods:number id="3743138ECB40993C20777811F6E1FD5E">2014-12-12</mods:number>
</mods:detail>
<mods:detail id="A5893571A87ACE4A9E32A699A0C3BEE0" type="volume">
<mods:number id="7E7F3351B95439251CFDA949EB9B422F">188</mods:number>
</mods:detail>
<mods:detail id="1041795852D2AC3D36B7C1F6E1382C83" type="issue">
<mods:number id="70F4687169B97199E7951EBDACEC053A">4</mods:number>
</mods:detail>
<mods:extent id="E99129D14414381241FC46E4DBDEF56A" unit="page">
<mods:start id="039BCA6F14658E6C1903453E417CFDF3">176</mods:start>
<mods:end id="4003C810F6C006A082828F9BE414D17D">180</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3CD02784B97C53DE84DD3D461AB5FC1E">
<mods:url id="7648A240B452B622DFEFB89832E7C682">http://dx.doi.org/10.11646/phytotaxa.188.3.7</mods:url>
</mods:location>
<mods:classification id="DFAA7768B1FECE75A3CDBA1089F24EE9">journal article</mods:classification>
<mods:identifier id="46C8F2FB99BF87123332058B02A2A651" type="DOI">10.11646/phytotaxa.188.3.7</mods:identifier>
<mods:identifier id="D500CBBA1754132362D8148D25CB5C85" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="07660EE6F94829CCA21892A00908CA13" type="Zenodo-Dep">5147403</mods:identifier>
</mods:mods>
<treatment id="F9118781F12BFFFE2E850104FB9D2CAC" ID-DOI="http://doi.org/10.5281/zenodo.5152323" ID-Zenodo-Dep="5152323" LSID="urn:lsid:plazi:treatment:F9118781F12BFFFE2E850104FB9D2CAC" httpUri="http://treatment.plazi.org/id/F9118781F12BFFFE2E850104FB9D2CAC" lastPageId="3" lastPageNumber="179" pageId="0" pageNumber="176">
<subSubSection id="39A2651CF12BFFFD2E850104FBE42DEB" box="[136,1106,1148,1175]" pageId="0" pageNumber="176" type="nomenclature">
<paragraph id="71073697F12BFFFD2E850104FBE42DEB" blockId="0.[136,1106,1148,1175]" box="[136,1106,1148,1175]" pageId="0" pageNumber="176">
<heading id="2A4F81FBF12BFFFD2E850104FBE42DEB" box="[136,1106,1148,1175]" fontSize="11" level="2" pageId="0" pageNumber="176" reason="3">
<taxonomicName id="B6B84D14F12BFFFD2E850104FCC62DEA" ID-CoL="7RTSL" authority="N.T.T Huong, B. H. Quang &amp; J. S Ma" authorityName="N. T. T Huong, B. H. Quang &amp; J. S Ma" authorityYear="2014" box="[136,880,1148,1175]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="xuanlienensis" status="sp. nov.">
<emphasis id="43CCEA85F12BFFFD2E850104FE0B2DEA" bold="true" box="[136,445,1148,1175]" italics="true" pageId="0" pageNumber="176">Aristolochia xuanlienensis</emphasis>
N.T.T Huong, B. H. Quang &amp; J. S Ma
</taxonomicName>
,
<emphasis id="43CCEA85F12BFFFD2D760105FC662DEB" box="[891,976,1149,1174]" italics="true" pageId="0" pageNumber="176">
<taxonomicNameLabel id="58FF57FEF12BFFFD2D760105FC662DEB" box="[891,976,1149,1174]" pageId="0" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="E9832A12F12BFFFD2DD20104FB9A2DEB" box="[991,1068,1148,1174]" captionStart="FIGURE 1" captionStartId="1.[136,229,1954,1976]" captionTargetBox="[164,1420,226,1922]" captionTargetId="figure-25@1.[151,1436,224,1932]" captionTargetPageId="1" captionText="FIGURE 1. Aristolochia xuanlienensis. A. Habit. B. Young branchlets. C. Old branchlets. D. Cross section of branchlets. E. Flowers (front view). F. Gynostemium with three lobes of stigma. G. Anthers magnified. H. Capsules. I. Cross section of capsules (illustrated by Le Kim Chi, AC &amp; EG from XL 129, D &amp; HI from DVH 105)." figureDoi="http://doi.org/10.5281/zenodo.5147405" httpUri="https://zenodo.org/record/5147405/files/figure.png" pageId="0" pageNumber="176">Figs. 1</figureCitation>
,
<figureCitation id="E9832A12F12BFFFD2A370104FBFF2DEB" box="[1082,1097,1148,1174]" captionStart="FIGURE 2" captionStartId="2.[136,229,1999,2021]" captionTargetBox="[158,1431,181,1985]" captionTargetId="figure-18@2.[152,1435,175,1989]" captionTargetPageId="2" captionText="FIGURE 2. Aristolochia xuanlienensis. AB. Habit. C. Young branchlets. D. Old branchlets. E. Cross section of branchlets. F. Leaf, adaxial view. G. Leaf, abaxial view. H. Racemes. I. Flowers (front view). J. Flowers (side view). K. Anthers. L. Gymnostemium. M. Capsules. N. Cross section of capsules. O. Young seed. (Photographs by Hai D.V (AJ) and Quang B.H (KO).)" figureDoi="http://doi.org/10.5281/zenodo.5147407" httpUri="https://zenodo.org/record/5147407/files/figure.png" pageId="0" pageNumber="176">2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39A2651CF12BFFFD2E8501B0FDFC2C5B" pageId="0" pageNumber="176" type="reference_group">
<paragraph id="71073697F12BFFFD2E8501B0FDFC2C5B" blockId="0.[136,1451,1224,1318]" pageId="0" pageNumber="176">
<taxonomicName id="B6B84D14F12BFFFD2E8501B0FE3E2DA0" ID-CoL="7RTSL" authorityName="N. T. T Huong, B. H. Quang &amp; J. S Ma" authorityYear="2014" box="[136,392,1224,1245]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="xuanlienensis">
<emphasis id="43CCEA85F12BFFFD2E8501B0FE3E2DA0" box="[136,392,1224,1245]" italics="true" pageId="0" pageNumber="176">Aristolochia xuanlienensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="B6B84D14F12BFFFD2C0A01B0FCEF2DA3" ID-CoL="7XD6L" authority="Merrill (1922: 341)" authorityName="Merrill" authorityPageNumber="341" authorityYear="1922" box="[519,857,1224,1246]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="hainanensis">
<emphasis id="43CCEA85F12BFFFD2C0A01B0FD232DA0" box="[519,661,1224,1245]" italics="true" pageId="0" pageNumber="176">A. hainanensis</emphasis>
Merrill (1922: 341)
</taxonomicName>
,
<taxonomicName id="B6B84D14F12BFFFD2D6E01B0FB102DA3" ID-CoL="7RTLH" authority="Franchet (1898: 311)" authorityName="Franchet" authorityPageNumber="311" authorityYear="1898" box="[867,1190,1224,1246]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="balansae">
<emphasis id="43CCEA85F12BFFFD2D6E01B0FC632DA0" box="[867,981,1224,1245]" italics="true" pageId="0" pageNumber="176">A. balansae</emphasis>
Franchet (1898: 311)
</taxonomicName>
and
<taxonomicName id="B6B84D14F12BFFFD2ADB01B0FE952C7F" ID-CoL="7RTQ2" authority="Do (2014: 52)" authorityName="Do" authorityPageNumber="52" authorityYear="2014" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="quangbinhensis">
<emphasis id="43CCEA85F12BFFFD2ADB01B0FA3E2DA0" box="[1238,1416,1224,1245]" italics="true" pageId="0" pageNumber="176">A. quangbinhensis</emphasis>
<bibRefCitation id="15294B66F12BFFFD2B8301B0FE952C7F" author="Do, T. V &amp; Nghiem, T. D. &amp; Wanke, S. &amp; Neinhuis, C." pageId="0" pageNumber="176" pagination="51 - 59" refId="ref1797" refString="Do, T. V, Nghiem, T. D., Wanke, S. &amp; Neinhuis, C. (2014) Arisotolochia quangbinhensis (Aristolochiaceae), a new species from central Vietnam. PhytoKeys 33: 51 - 59. http: // dx. doi. org / 10.3897 / phytokeys. 33.6094" type="journal article" year="2014">Do (2014: 52)</bibRefCitation>
</taxonomicName>
, but its white perianth with light purple veins on the outside and dark purple spots both in tube and on limb has never been reported before among the Asian species.
</paragraph>
</subSubSection>
<subSubSection id="39A2651CF12BFFFD2E850020FC832EF6" pageId="0" pageNumber="176" type="materials_examined">
<paragraph id="71073697F12BFFFD2E850020FBD92CEF" blockId="0.[136,1451,1368,1426]" pageId="0" pageNumber="176">
<materialsCitation id="C1D03CCAF12BFFFD2E850020FBDD2CEF" collectingDate="2011-11-07" collectionCode="HN, CSH" collectorName="Hai &amp; Quang &amp; Cuong &amp; Vu &amp; Thanh" country="Vietnam" elevation="870" latitude="19.995695" location="Xuan Lien Nature Reserve" longLatPrecision="1" longitude="105.00242" municipality="Type" pageId="0" pageNumber="176" specimenCount="1" stateProvince="Thanh Hoa" typeStatus="holotype">
<collectingMunicipality id="9163ACEDF12BFFFD2E850020FF0C2C10" box="[136,186,1368,1389]" pageId="0" pageNumber="176">Type</collectingMunicipality>
:—
<collectingCountry id="09AF7607F12BFFFD2EDB0020FEFE2C10" box="[214,328,1368,1389]" name="Vietnam" pageId="0" pageNumber="176">VIETNAM</collectingCountry>
.
<collectingRegion id="B37CF875F12BFFFD2F420020FDA02C13" box="[335,534,1368,1390]" country="Vietnam" name="Thanh Hoa" pageId="0" pageNumber="176">Thanh Hoa Province</collectingRegion>
:
<location id="7467604CF12BFFFD2C130020FCAB2C10" LSID="urn:lsid:plazi:treatment:F9118781F12BFFFE2E850104FB9D2CAC:7467604CF12BFFFD2C130020FCAB2C10" box="[542,797,1368,1390]" country="Vietnam" latitude="19.995695" longLatPrecision="1" longitude="105.00242" municipality="Type" name="Xuan Lien Nature Reserve" pageId="0" pageNumber="176" stateProvince="Thanh Hoa">Xuan Lien Nature Reserve</location>
,
<geoCoordinate id="148C5050F12BFFFD2D280020FC042C13" box="[805,946,1368,1390]" degrees="19" direction="north" minutes="59" orientation="latitude" pageId="0" pageNumber="176" precision="1" seconds="44.5" value="19.995695">19°5944.5N</geoCoordinate>
,
<geoCoordinate id="148C5050F12BFFFD2DB70020FBE72C10" box="[954,1105,1368,1390]" degrees="105" direction="east" minutes="00" orientation="longitude" pageId="0" pageNumber="176" precision="1" seconds="08.7" value="105.00242">105°0008.7E</geoCoordinate>
, elevation about
<quantity id="B6409B72F12BFFFD2AFD0020FA9B2C10" box="[1264,1325,1368,1389]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="0" pageNumber="176" unit="m" value="870.0">
<elevation id="FA95D1A4F12BFFFD2AFD0020FA9B2C10" box="[1264,1325,1368,1389]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="0" pageNumber="176" unit="m" value="870.0">870 m</elevation>
</quantity>
,
<date id="05061057F12BFFFD2B3B0020FF582CEC" pageId="0" pageNumber="176" value="2011-11-07">
<collectingDate id="1542E9BFF12BFFFD2B3B0020FF582CEC" pageId="0" pageNumber="176" value="2011-11-07">7 November 2011</collectingDate>
</date>
,
<emphasis id="43CCEA85F12BFFFD2EF40004FD1C2CEC" box="[249,682,1404,1425]" italics="true" pageId="0" pageNumber="176">
<collectorName id="DC4D5341F12BFFFD2EF40004FEA92CEC" box="[249,287,1404,1425]" pageId="0" pageNumber="176">Hai</collectorName>
,
<collectorName id="DC4D5341F12BFFFD2F240004FEDB2CEC" box="[297,365,1404,1425]" pageId="0" pageNumber="176">Quang</collectorName>
,
<collectorName id="DC4D5341F12BFFFD2F7B0004FE0E2CEC" box="[374,440,1404,1425]" pageId="0" pageNumber="176">Cuong</collectorName>
,
<collectorName id="DC4D5341F12BFFFD2FCF0004FE6B2CEC" box="[450,477,1404,1425]" pageId="0" pageNumber="176">Vu</collectorName>
,
<collectorName id="DC4D5341F12BFFFD2FEA0004FD922CEC" box="[487,548,1404,1425]" pageId="0" pageNumber="176">Thanh</collectorName>
&amp;
<collectorName id="DC4D5341F12BFFFD2C4E0004FD362CEC" box="[579,640,1404,1425]" pageId="0" pageNumber="176">Thanh</collectorName>
129
</emphasis>
(
<typeStatus id="AE038835F12BFFFD2CBA0004FCBB2CEF" box="[695,781,1404,1426]" pageId="0" pageNumber="176" type="holotype">holotype</typeStatus>
<collectionCode id="17A9AE52F12BFFFD2D1F0004FC832CEC" box="[786,821,1404,1425]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="0" pageNumber="176" type="Herbarium">HN</collectionCode>
00000 68875; isotypes:
<collectionCode id="17A9AE52F12BFFFD2A3C0004FBD22CEF" box="[1073,1124,1404,1426]" country="China" httpUri="http://grbio.org/cool/vr0i-2neq" name="Chenshan Botanical Garden" pageId="0" pageNumber="176" type="Herbarium">CSH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="71073697F12BFFFD2E8500BCFC192C80" blockId="0.[136,1451,1476,1534]" pageId="0" pageNumber="176">
Other specimens examined:—
<materialsCitation id="C1D03CCAF12BFFFD2FA300BCFC1D2C80" collectingDate="2014-04-17" collectionCode="HN, VNMN" collectorName="Huong &amp; Hai &amp; Quang &amp; Khang" country="Vietnam" elevation="200" latitude="19.847584" location="Xuan Lien Nature Reserve" longLatPrecision="1" longitude="105.244606" pageId="0" pageNumber="176" specimenCount="1" stateProvince="Thanh Hoa">
<collectingCountry id="09AF7607F12BFFFD2FA300BCFD962CA7" box="[430,544,1476,1498]" name="Vietnam" pageId="0" pageNumber="176">VIETNAM</collectingCountry>
.
<collectingRegion id="B37CF875F12BFFFD2C2600BCFD4F2CA7" box="[555,761,1476,1498]" country="Vietnam" name="Thanh Hoa" pageId="0" pageNumber="176">Thanh Hoa Province</collectingRegion>
:
<location id="7467604CF12BFFFD2D0900BCFBBB2CA4" LSID="urn:lsid:plazi:treatment:F9118781F12BFFFE2E850104FB9D2CAC:7467604CF12BFFFD2D0900BCFBBB2CA4" box="[772,1037,1476,1498]" country="Vietnam" latitude="19.847584" longLatPrecision="1" longitude="105.244606" name="Xuan Lien Nature Reserve" pageId="0" pageNumber="176" stateProvince="Thanh Hoa">Xuan Lien Nature Reserve</location>
,
<geoCoordinate id="148C5050F12BFFFD2A1500BCFB132CA7" box="[1048,1189,1476,1498]" degrees="19" direction="north" minutes="50" orientation="latitude" pageId="0" pageNumber="176" precision="1" seconds="51.3" value="19.847584">19°5051.3N</geoCoordinate>
,
<geoCoordinate id="148C5050F12BFFFD2ABC00BCFAFE2CA7" box="[1201,1352,1476,1498]" degrees="105" direction="east" minutes="14" orientation="longitude" pageId="0" pageNumber="176" precision="1" seconds="40.6" value="105.244606">105°1440.6E</geoCoordinate>
, elevation about
<quantity id="B6409B72F12BFFFD2EF40090FE8E2C80" box="[249,312,1512,1533]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="176" unit="m" value="200.0">
<elevation id="FA95D1A4F12BFFFD2EF40090FE8E2C80" box="[249,312,1512,1533]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="176" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="05061057F12BFFFD2F4C0090FE7F2C80" box="[321,457,1512,1534]" pageId="0" pageNumber="176" value="2014-04-17">
<collectingDate id="1542E9BFF12BFFFD2F4C0090FE7F2C80" box="[321,457,1512,1534]" pageId="0" pageNumber="176" value="2014-04-17">17 April 2014</collectingDate>
</date>
,
<emphasis id="43CCEA85F12BFFFD2FDE0090FCAC2C80" box="[467,794,1512,1533]" italics="true" pageId="0" pageNumber="176">
<collectorName id="DC4D5341F12BFFFD2FDE0090FDA12C80" box="[467,535,1512,1533]" pageId="0" pageNumber="176">Huong</collectorName>
,
<collectorName id="DC4D5341F12BFFFD2C2D0090FDF02C80" box="[544,582,1512,1533]" pageId="0" pageNumber="176">Hai</collectorName>
,
<collectorName id="DC4D5341F12BFFFD2C5D0090FD272C80" box="[592,657,1512,1533]" pageId="0" pageNumber="176">Quang</collectorName>
&amp;
<collectorName id="DC4D5341F12BFFFD2CBD0090FD462C80" box="[688,752,1512,1533]" pageId="0" pageNumber="176">Khang</collectorName>
105
</emphasis>
(
<collectionCode id="17A9AE52F12BFFFD2D2A0090FCFB2C80" box="[807,845,1512,1533]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="0" pageNumber="176" type="Herbarium">HN</collectionCode>
,
<collectionCode id="17A9AE52F12BFFFD2D5B0090FC122C80" box="[854,932,1512,1533]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="0" pageNumber="176">VNMN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="71073697F12BFFFD2E850354FD142E1A" blockId="0.[136,1452,1580,2075]" pageId="0" pageNumber="176">
Liana with a woody stem, terete, striate, yellow, hairs brown, old branchlets glabrous, up to more than
<quantity id="B6409B72F12BFFFD2AE00354FA952F3A" box="[1261,1315,1580,1607]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="176" unit="cm" value="1.0">1 cm</quantity>
in diameter. Leaves with the petioles
<quantity id="B6409B72F12BFFFD2F920328FE442F16" box="[415,498,1616,1643]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="0" pageNumber="176" unit="cm" value="5.5" valueMax="8.0" valueMin="3.0">38 cm</quantity>
long, densely yellow-brown villous; leaf blade ovate or ovate-lanceolate to narrowly ovate, 1225 x
<quantity id="B6409B72F12BFFFD2F39030CFE2E2FF2" box="[308,408,1652,1679]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="0" pageNumber="176" unit="cm" value="11.0" valueMax="15.0" valueMin="7.0">715 cm</quantity>
, both surface densely villous when young, becoming glabrous when old; margin entire; veins palmate, 45 pairs from base; base rounded, apex acuminate. Racemes usually in axils of leafy shoots or on old woody stems, 34(5) flowers, ca
<quantity id="B6409B72F12BFFFD2FB603C4FDBE2FAA" box="[443,520,1724,1751]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="0" pageNumber="176" unit="cm" value="3.5">3.5 cm</quantity>
long; peduncle often pendulous, ca.
<quantity id="B6409B72F12BFFFD2DA703C4FC542FAA" box="[938,994,1724,1751]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="176" unit="cm" value="3.0">3 cm</quantity>
long, densely pubescent. Perianth white with light purple veins outside and dark purple spots both in tube and on limb; limb obliquely trumpet-shaped, ca
<quantity id="B6409B72F12BFFFD2B850398FF1C2E62" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="0" pageNumber="176" unit="cm" value="2.5">2.5 cm</quantity>
in diameter, trilobed; lobes subequal, densely papillate, upper 2 oblong, lower one broadly deltoid. Gynostemium is with trilobed stigma on top, anthers
<quantity id="B6409B72F12BFFFD2C100250FDFE2E3F" box="[541,584,1832,1858]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="0" pageNumber="176" unit="in" value="6.0">6 in</quantity>
3 pairs, oblong, ca.
<quantity id="B6409B72F12BFFFD2D260250FCDA2E3E" box="[811,876,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="176" unit="mm" value="2.0">2 mm</quantity>
long. Capsule narrowly cylindric, yellow, 8.010.0 x 2.0
<quantity id="B6409B72F12BFFFD2EC20234FEA82E1A" box="[207,286,1868,1895]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="0" pageNumber="176" unit="cm" value="2.5">2.5 cm</quantity>
. Seeds cordate, ca. 46 x
<quantity id="B6409B72F12BFFFD2C330234FD282E1A" box="[574,670,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="0" pageNumber="176" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
.
</paragraph>
<paragraph id="71073697F12BFFFD2EB00208FC832EF6" blockId="0.[136,1452,1580,2075]" box="[189,821,1904,1931]" pageId="0" pageNumber="176">
<emphasis id="43CCEA85F12BFFFD2EB00208FE072EF7" bold="true" box="[189,433,1904,1930]" pageId="0" pageNumber="176">Vernacular name:—</emphasis>
Phòng kỷ Xuân Liên (Vietnamese)
</paragraph>
</subSubSection>
<subSubSection id="39A2651CF12BFFFE2EB002ECFC6E2DCC" lastPageId="3" lastPageNumber="179" pageId="0" pageNumber="176" type="distribution">
<paragraph id="71073697F12BFFFD2EB002ECFBB42167" blockId="0.[136,1452,1580,2075]" pageId="0" pageNumber="176">
<emphasis id="43CCEA85F12BFFFD2EB002ECFDB52ED3" bold="true" box="[189,515,1940,1966]" pageId="0" pageNumber="176">Distribution and habitat:—</emphasis>
<taxonomicName id="B6B84D14F12BFFFD2C0902ECFC9A2ED3" authorityName="N. T. T Huong, B. H. Quang &amp; J. S Ma" authorityYear="2014" box="[516,812,1940,1966]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="xuanlienensis">
<emphasis id="43CCEA85F12BFFFD2C0902ECFC9A2ED3" box="[516,812,1940,1966]" italics="true" pageId="0" pageNumber="176">Aristolochia xuanlienensis</emphasis>
</taxonomicName>
is currently known only from Xuan Lien Nature Reserve,
<collectingRegion id="B37CF875F12BFFFD2E8502C0FEC42EAF" box="[136,370,1976,2003]" country="Vietnam" name="Thanh Hoa" pageId="0" pageNumber="176">Thanh Hoa Province</collectingRegion>
,
<collectingCountry id="09AF7607F12BFFFD2F7002C0FE692EAF" box="[381,479,1976,2002]" name="Vietnam" pageId="0" pageNumber="176">Vietnam</collectingCountry>
, at elevations between
<quantity id="B6409B72F12BFFFD2CEC02C0FCD52EAE" box="[737,867,1976,2003]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="0" pageNumber="176" unit="m" value="550.0" valueMax="900.0" valueMin="200.0">
200
<specimenCount id="67BEFD1EF12BFFFD2D1402C0FCD52EAE" box="[793,867,1976,2003]" pageId="0" pageNumber="176" type="generic">900 m</specimenCount>
</quantity>
. The new species usually grows in the understory of primary subtropical evergreen forests, along with tree species including
<taxonomicName id="B6B84D14F12BFFFD2DCD02A4FB302E8A" box="[960,1158,2012,2039]" class="Liliopsida" family="Araceae" genus="Raphidophora" kingdom="Plantae" order="Alismatales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="43CCEA85F12BFFFD2DCD02A4FBD42E8B" box="[960,1122,2012,2038]" italics="true" pageId="0" pageNumber="176">Raphidophora</emphasis>
sp
</taxonomicName>
,
<taxonomicName id="B6B84D14F12BFFFD2A9E02A4FA962E8A" box="[1171,1312,2012,2039]" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="43CCEA85F12BFFFD2A9E02A4FB432E8B" box="[1171,1269,2012,2038]" italics="true" pageId="0" pageNumber="176">Alocasia</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="B6B84D14F12BFFFD2B2302A4FA102E8A" box="[1326,1446,2012,2039]" class="Liliopsida" family="Araceae" genus="Pothos" kingdom="Plantae" order="Alismatales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="43CCEA85F12BFFFD2B2302A4FACA2E8B" box="[1326,1404,2012,2038]" italics="true" pageId="0" pageNumber="176">Pothos</emphasis>
sp.
</taxonomicName>
,
<emphasis id="43CCEA85F12BFFFD2E850D78FEB62167" box="[136,256,2048,2074]" italics="true" pageId="0" pageNumber="176">Paraborea</emphasis>
sp.,
<taxonomicName id="B6B84D14F12BFFFD2F380D79FD962167" authority="Pit." authorityName="Pit." box="[309,544,2048,2074]" class="Magnoliopsida" family="Rubiaceae" genus="Tarenna" kingdom="Plantae" order="Gentianales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="43CCEA85F12BFFFD2F380D79FE472167" box="[309,497,2048,2074]" italics="true" pageId="0" pageNumber="176">Tarenna latifolia</emphasis>
Pit.
</taxonomicName>
,
<taxonomicName id="B6B84D14F12BFFFD2C260D78FD792166" box="[555,719,2048,2075]" class="Magnoliopsida" family="Urticaceae" genus="Boehmeria" kingdom="Plantae" order="Rosales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="43CCEA85F12BFFFD2C260D78FD102167" box="[555,678,2048,2074]" italics="true" pageId="0" pageNumber="176">Boehmeria</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="B6B84D14F12BFFFD2CD60D79FC8D2166" box="[731,827,2049,2075]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="0" pageNumber="176" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="43CCEA85F12BFFFD2CD60D79FCAF2167" box="[731,793,2049,2074]" italics="true" pageId="0" pageNumber="176">Ficus</emphasis>
sp
</taxonomicName>
, ferns and others.
</paragraph>
<caption id="25C7661FF12AFFFC2E8502DAFCB5217D" ID-DOI="http://doi.org/10.5281/zenodo.5147405" ID-Zenodo-Dep="5147405" httpUri="https://zenodo.org/record/5147405/files/figure.png" pageId="1" pageNumber="177" startId="1.[136,229,1954,1976]" targetBox="[164,1420,226,1922]" targetPageId="1">
<paragraph id="71073697F12AFFFC2E8502DAFCB5217D" blockId="1.[136,1451,1954,2049]" pageId="1" pageNumber="177">
<emphasis id="43CCEA85F12AFFFC2E8502DAFF4F2ECA" bold="true" box="[136,249,1954,1976]" pageId="1" pageNumber="177">FIGURE 1</emphasis>
.
<emphasis id="43CCEA85F12AFFFC2F0B02DAFD9B2ECA" bold="true" box="[262,557,1954,1976]" pageId="1" pageNumber="177">
<emphasis id="43CCEA85F12AFFFC2F0B02DAFDA02EC5" bold="true" box="[262,534,1954,1976]" italics="true" pageId="1" pageNumber="177">
<taxonomicName id="B6B84D14F12AFFFC2F0B02DAFDA42EC5" authorityName="N. T. T Huong, B. H. Quang &amp; J. S Ma" authorityYear="2014" box="[262,530,1954,1976]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="xuanlienensis">Aristolochia xuanlienensis</taxonomicName>
.
</emphasis>
A
</emphasis>
. Habit.
<emphasis id="43CCEA85F12AFFFC2C8E02DBFD2C2EC5" bold="true" box="[643,666,1955,1976]" pageId="1" pageNumber="177">B.</emphasis>
Young branchlets.
<emphasis id="43CCEA85F12AFFFC2D5802DAFCDB2EC5" bold="true" box="[853,877,1954,1976]" pageId="1" pageNumber="177">C.</emphasis>
Old branchlets.
<emphasis id="43CCEA85F12AFFFC2A0202DBFB902EC5" bold="true" box="[1039,1062,1955,1976]" pageId="1" pageNumber="177">D.</emphasis>
Cross section of branchlets.
<emphasis id="43CCEA85F12AFFFC2B4C02DBFAE12EC5" bold="true" box="[1345,1367,1955,1976]" pageId="1" pageNumber="177">E.</emphasis>
Flowers (front view).
<emphasis id="43CCEA85F12AFFFC2F0A02BFFEAC2EA1" bold="true" box="[263,282,1991,2012]" pageId="1" pageNumber="177">F.</emphasis>
Gynostemium with three lobes of stigma.
<emphasis id="43CCEA85F12AFFFC2CB902BEFD7A2EA1" bold="true" box="[692,716,1990,2012]" pageId="1" pageNumber="177">G.</emphasis>
Anthers magnified.
<emphasis id="43CCEA85F12AFFFC2D9B02BFFC192EA1" bold="true" box="[918,943,1991,2012]" pageId="1" pageNumber="177">H.</emphasis>
Capsules.
<emphasis id="43CCEA85F12AFFFC2A1502BFFB912EA1" bold="true" box="[1048,1063,1991,2012]" pageId="1" pageNumber="177">I.</emphasis>
Cross section of capsules (illustrated by Le Kim Chi, AC &amp; EG from
<emphasis id="43CCEA85F12AFFFC2FB50293FE48217D" box="[440,510,2027,2048]" italics="true" pageId="1" pageNumber="177">XL 129</emphasis>
, D &amp; HI from
<emphasis id="43CCEA85F12AFFFC2C970293FD43217D" box="[666,757,2027,2048]" italics="true" pageId="1" pageNumber="177">DVH 105</emphasis>
).
</paragraph>
</caption>
<caption id="25C7661FF129FFFF2E8502B7FB73215A" ID-DOI="http://doi.org/10.5281/zenodo.5147407" ID-Zenodo-Dep="5147407" httpUri="https://zenodo.org/record/5147407/files/figure.png" pageId="2" pageNumber="178" startId="2.[136,229,1999,2021]" targetBox="[158,1431,181,1985]" targetPageId="2">
<paragraph id="71073697F129FFFF2E8502B7FA1D217F" blockId="2.[136,1452,1999,2087]" pageId="2" pageNumber="178">
<emphasis id="43CCEA85F129FFFF2E8502B7FF4F2E99" bold="true" box="[136,249,1999,2021]" pageId="2" pageNumber="178">FIGURE 2</emphasis>
.
<emphasis id="43CCEA85F129FFFF2F0A02B7FDFA2E99" bold="true" box="[263,588,1999,2021]" pageId="2" pageNumber="178">
<emphasis id="43CCEA85F129FFFF2F0A02B7FDA12E98" bold="true" box="[263,535,1999,2021]" italics="true" pageId="2" pageNumber="178">
<taxonomicName id="B6B84D14F129FFFF2F0A02B7FDA22E98" authorityName="N. T. T Huong, B. H. Quang &amp; J. S Ma" authorityYear="2014" box="[263,532,1999,2021]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="2" pageNumber="178" phylum="Tracheophyta" rank="species" species="xuanlienensis">Aristolochia xuanlienensis</taxonomicName>
.
</emphasis>
AB
</emphasis>
. Habit.
<emphasis id="43CCEA85F129FFFF2C9102B7FD022E98" bold="true" box="[668,692,1999,2021]" pageId="2" pageNumber="178">C.</emphasis>
Young branchlets.
<emphasis id="43CCEA85F129FFFF2D7C02A8FC3F2E98" bold="true" box="[881,905,2000,2021]" pageId="2" pageNumber="178">D.</emphasis>
Old branchlets.
<emphasis id="43CCEA85F129FFFF2A2102A8FBF42E98" bold="true" box="[1068,1090,2000,2021]" pageId="2" pageNumber="178">E.</emphasis>
Cross section of branchlets.
<emphasis id="43CCEA85F129FFFF2B5202A8FAC42E98" bold="true" box="[1375,1394,2000,2021]" pageId="2" pageNumber="178">F.</emphasis>
Leaf, adaxial view.
<emphasis id="43CCEA85F129FFFF2F1D0295FE9F217F" bold="true" box="[272,297,2029,2051]" pageId="2" pageNumber="178">G.</emphasis>
Leaf, abaxial view.
<emphasis id="43CCEA85F129FFFF2FFF0295FDBD217F" bold="true" box="[498,523,2029,2050]" pageId="2" pageNumber="178">H.</emphasis>
Racemes.
<emphasis id="43CCEA85F129FFFF2C7A0295FD30217F" bold="true" box="[631,646,2029,2050]" pageId="2" pageNumber="178">I.</emphasis>
Flowers (front view).
<emphasis id="43CCEA85F129FFFF2D6B0295FCCE217F" bold="true" box="[870,888,2029,2050]" pageId="2" pageNumber="178">J.</emphasis>
Flowers (side view).
<emphasis id="43CCEA85F129FFFF2A5D0295FBDF217F" bold="true" box="[1104,1129,2029,2050]" pageId="2" pageNumber="178">K.</emphasis>
Anthers.
<emphasis id="43CCEA85F129FFFF2AC50295FB69217F" bold="true" box="[1224,1247,2029,2050]" pageId="2" pageNumber="178">L.</emphasis>
Gymnostemium.
<emphasis id="43CCEA85F129FFFF2B820295FA1D217F" bold="true" box="[1423,1451,2029,2050]" pageId="2" pageNumber="178">M.</emphasis>
</paragraph>
<paragraph id="71073697F129FFFF2E850D69FB73215A" blockId="2.[136,1452,1999,2087]" box="[136,1221,2065,2087]" pageId="2" pageNumber="178">
Capsules.
<emphasis id="43CCEA85F129FFFF2EE60D69FEB4215B" bold="true" box="[235,258,2065,2086]" pageId="2" pageNumber="178">N.</emphasis>
Cross section of capsules.
<emphasis id="43CCEA85F129FFFF2C0B0D69FDA9215A" bold="true" box="[518,543,2065,2087]" pageId="2" pageNumber="178">O.</emphasis>
Young seed. (Photographs by Hai D.V (AJ) and Quang B.H (KO).)
</paragraph>
</caption>
<caption id="25C7661FF128FFFE2E8505D8FC7729CA" box="[136,961,160,184]" pageId="3" pageNumber="179" startId="3.[136,224,160,184]">
<paragraph id="71073697F128FFFE2E8505D8FC7729CA" blockId="3.[136,961,160,184]" box="[136,961,160,184]" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2E8505D8FF4D29C5" bold="true" box="[136,251,160,184]" pageId="3" pageNumber="179">TABLE 1.</emphasis>
Comparison of key characters in
<taxonomicName id="B6B84D14F128FFFE2C6E05D8FCA429C5" authorityName="N. T. T Huong, B. H. Quang &amp; J. S Ma" authorityYear="2014" box="[611,786,160,184]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="179" phylum="Tracheophyta" rank="species" species="xuanlienensis">
<emphasis id="43CCEA85F128FFFE2C6E05D8FCA429C5" box="[611,786,160,184]" italics="true" pageId="3" pageNumber="179">A. xuanlienensis</emphasis>
</taxonomicName>
and related taxa
</paragraph>
</caption>
<paragraph id="71073697F128FFFE2E9E05B0FAF42D38" pageId="3" pageNumber="179">
<table id="03B8C437F12800022E9E05B0FA152D38" box="[147,1443,200,1093]" gridcols="5" gridrows="6" pageId="3" pageNumber="179">
<tr id="CF8834D5F12800022E9E05B0FA1529A3" box="[147,1443,200,222]" gridrow="0" pageId="3" pageNumber="179">
<th id="8C595DA9F12800022E9E05B0FE8529A3" box="[147,307,200,222]" gridcol="0" gridrow="0" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2E9E05B0FEB029A3" bold="true" box="[147,262,200,222]" pageId="3" pageNumber="179">Characters</emphasis>
</th>
<th id="8C595DA9F12800022F6C05B0FDC229A3" box="[353,628,200,222]" gridcol="1" gridrow="0" pageId="3" pageNumber="179">
<taxonomicName id="B6B84D14F128FFFE2F6C05B1FE4829A3" authorityName="N. T. T Huong, B. H. Quang &amp; J. S Ma" authorityYear="2014" box="[353,510,201,222]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="179" phylum="Tracheophyta" rank="species" species="xuanlienensis">
<emphasis id="43CCEA85F128FFFE2F6C05B1FE4829A3" box="[353,510,201,222]" italics="true" pageId="3" pageNumber="179">A. xuanlienensis</emphasis>
</taxonomicName>
</th>
<th id="8C595DA9F12800022C8005B0FCD629A3" box="[653,864,200,222]" gridcol="2" gridrow="0" pageId="3" pageNumber="179">
<taxonomicName id="B6B84D14F128FFFE2C8005B1FD4829A3" authorityName="Franchet" authorityYear="1898" box="[653,766,201,222]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="179" phylum="Tracheophyta" rank="species" species="balansae">
<emphasis id="43CCEA85F128FFFE2C8005B1FD4829A3" box="[653,766,201,222]" italics="true" pageId="3" pageNumber="179">A. balansae</emphasis>
</taxonomicName>
</th>
<th id="8C595DA9F12800022D7405B0FBC229A3" box="[889,1140,200,222]" gridcol="3" gridrow="0" pageId="3" pageNumber="179">
<taxonomicName id="B6B84D14F128FFFE2D7405B1FBB129A3" authorityName="Merrill" authorityYear="1922" box="[889,1031,201,222]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="179" phylum="Tracheophyta" rank="species" species="hainanensis">
<emphasis id="43CCEA85F128FFFE2D7405B1FBB129A3" box="[889,1031,201,222]" italics="true" pageId="3" pageNumber="179">A. hainanensis</emphasis>
</taxonomicName>
</th>
<th id="8C595DA9F12800022A9805B0FA1529A3" box="[1173,1443,200,222]" gridcol="4" gridrow="0" pageId="3" pageNumber="179">
<taxonomicName id="B6B84D14F128FFFE2A9805B1FAF129A3" authorityName="Do" authorityYear="2014" box="[1173,1351,201,222]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="179" phylum="Tracheophyta" rank="species" species="quangbinhensis">
<emphasis id="43CCEA85F128FFFE2A9805B1FAF129A3" box="[1173,1351,201,222]" italics="true" pageId="3" pageNumber="179">A. quangbinhensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="CF8834D5F12800022E9E0588FA1528A3" box="[147,1443,240,478]" gridrow="1" pageId="3" pageNumber="179">
<th id="8C595DA9F12800022E9E0588FE8528A3" box="[147,307,240,478]" gridcol="0" gridrow="1" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2E9E0588FF582878" bold="true" box="[147,238,240,261]" pageId="3" pageNumber="179">Racemes</emphasis>
</th>
<td id="8C595DA9F12800022F6C0588FDC228A3" box="[353,628,240,478]" gridcol="1" gridrow="1" pageId="3" pageNumber="179">In axils of leafy shoots or on old woody stems, 34 (5) flowers, ca 3.5 cm long; peduncle ca. 3 cm long, densely pubescent, bracteole subulate, ca. 2 × 1 mm</td>
<td id="8C595DA9F12800022C800588FCD628A3" box="[653,864,240,478]" gridcol="2" gridrow="1" pageId="3" pageNumber="179">In axils of leafy shoots, 23 flowers, ca. 5 cm long; peduncle 35 cm long, pubescent, bracteole small at base</td>
<td id="8C595DA9F12800022D740588FBC228A3" box="[889,1140,240,478]" gridcol="3" gridrow="1" pageId="3" pageNumber="179">In axils of leafy shoots or on old woody stems, 36flowered, ca. 3 cm long; peduncle 3.54.0 cm long, densely villous; bracteole subulate, ca. 2 × 1 mm</td>
<td id="8C595DA9F12800022A980588FA1528A3" box="[1173,1443,240,478]" gridcol="4" gridrow="1" pageId="3" pageNumber="179">Terminal, solitary, accompanied by a lanceolate scale-like bracteole, (2.0)3.0 mm long, (1.5)2.0 mm wide; peduncle 12 cm long</td>
</tr>
<tr id="CF8834D5F12800022E9E076BFA152BC7" box="[147,1443,531,698]" gridrow="2" pageId="3" pageNumber="179">
<th id="8C595DA9F12800022E9E076BFE852BC7" box="[147,307,531,698]" gridcol="0" gridrow="2" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2E9E076BFE942B54" bold="true" box="[147,290,531,553]" pageId="3" pageNumber="179">Perianth limb</emphasis>
</th>
<td id="8C595DA9F12800022F6C076BFDC22BC7" box="[353,628,531,698]" gridcol="1" gridrow="2" pageId="3" pageNumber="179">Limb obliquely trumpet- shaped, ca 2.5 cm in diameter; trilobed, lobes white, subequal, with purple spots</td>
<td id="8C595DA9F12800022C80076BFCD62BC7" box="[653,864,531,698]" gridcol="2" gridrow="2" pageId="3" pageNumber="179">Limb two-lobed, bell- shaped, ca 1.8 cm in diameter, lobes red, papillose</td>
<td id="8C595DA9F12800022D74076BFBC22BC7" box="[889,1140,531,698]" gridcol="3" gridrow="2" pageId="3" pageNumber="179">Limb obliquely trumpet- shaped, ca. 3 cm in diameter; trilobed, lobes purple, subequal, densely papillate</td>
<td id="8C595DA9F12800022A98076BFA152BC7" box="[1173,1443,531,698]" gridcol="4" gridrow="2" pageId="3" pageNumber="179">Limb trilobed, bell-shaped, 1.02.5(3) cm diameter, purplish-pink on both sides, without spots and papillae</td>
</tr>
<tr id="CF8834D5F12800022E9E078AFA152AE5" box="[147,1443,754,920]" gridrow="3" pageId="3" pageNumber="179">
<th id="8C595DA9F12800022E9E078AFE852AE5" box="[147,307,754,920]" gridcol="0" gridrow="3" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2E9E078AFE852A7A" bold="true" box="[147,307,754,776]" pageId="3" pageNumber="179">Perianth throat</emphasis>
</th>
<td id="8C595DA9F12800022F6C078AFDC22AE5" box="[353,628,754,920]" gridcol="1" gridrow="3" pageId="3" pageNumber="179">White with dark purple spots both in tube and on limb, tube geniculate curved, without villous</td>
<td id="8C595DA9F12800022C80078AFCD62AE5" box="[653,864,754,920]" gridcol="2" gridrow="3" pageId="3" pageNumber="179">Red with dirty spots within the tube but outside light red with hair, geniculate curved</td>
<td id="8C595DA9F12800022D74078AFBC22AE5" box="[889,1140,754,920]" gridcol="3" gridrow="3" pageId="3" pageNumber="179">Yellow without spots; tube geniculate curved, abaxially sparsely villous</td>
<td id="8C595DA9F12800022A98078AFA152AE5" box="[1173,1443,754,920]" gridcol="4" gridrow="3" pageId="3" pageNumber="179">Dark purple, without spots; densely papillose, annulus present, formed by an ellipsoid flange</td>
</tr>
<tr id="CF8834D5F12800022E9E06B9FA152A81" box="[147,1443,961,1020]" gridrow="4" pageId="3" pageNumber="179">
<th id="8C595DA9F12800022E9E06B9FE852A81" box="[147,307,961,1020]" gridcol="0" gridrow="4" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2E9E06B9FF502AAA" bold="true" box="[147,230,961,983]" pageId="3" pageNumber="179">Capsule</emphasis>
</th>
<td id="8C595DA9F12800022F6C06B9FDC22A81" box="[353,628,961,1020]" gridcol="1" gridrow="4" pageId="3" pageNumber="179">Cylindric, yellow, 8.010.0 × 2.02.5 cm.</td>
<td id="8C595DA9F12800022C8006B9FCD62A81" box="[653,864,961,1020]" gridcol="2" gridrow="4" pageId="3" pageNumber="179">Cylindric, 10.012.0 × 3.04.0 cm</td>
<td id="8C595DA9F12800022D7406B9FBC22A81" box="[889,1140,961,1020]" gridcol="3" gridrow="4" pageId="3" pageNumber="179">Cylindric, 7.010.0 cm × 2.53.0 cm</td>
<td id="8C595DA9F12800022A9806B9FA152A81" box="[1173,1443,961,1020]" gridcol="4" gridrow="4" pageId="3" pageNumber="179">Unknown</td>
</tr>
<tr id="CF8834D5F12800022E9E0156FA152D38" box="[147,1443,1070,1093]" gridrow="5" pageId="3" pageNumber="179">
<th id="8C595DA9F12800022E9E0156FE852D38" box="[147,307,1070,1093]" gridcol="0" gridrow="5" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2E9E0156FEA62D39" bold="true" box="[147,272,1070,1092]" pageId="3" pageNumber="179">Distribution</emphasis>
</th>
<td id="8C595DA9F12800022F6C0156FDC22D38" box="[353,628,1070,1093]" gridcol="1" gridrow="5" pageId="3" pageNumber="179">Southern Vietnam</td>
<td id="8C595DA9F12800022C800156FCD62D38" box="[653,864,1070,1093]" gridcol="2" gridrow="5" pageId="3" pageNumber="179">Northern Vietnam</td>
<td id="8C595DA9F12800022D740156FBC22D38" box="[889,1140,1070,1093]" gridcol="3" gridrow="5" pageId="3" pageNumber="179">China, Hainan</td>
<td id="8C595DA9F12800022A980156FA152D38" box="[1173,1443,1070,1093]" gridcol="4" gridrow="5" pageId="3" pageNumber="179">Southern Vietnam</td>
</tr>
</table>
</paragraph>
<paragraph id="71073697F128FFFE2EB001E0FC6E2DCC" blockId="3.[136,1452,1175,1490]" box="[189,984,1175,1202]" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2EB001E0FEF52DCF" bold="true" box="[189,323,1176,1202]" pageId="3" pageNumber="179">Phenology:</emphasis>
—Flowering SeptemberJanuary and fruiting MarchMay.
</paragraph>
</subSubSection>
<subSubSection id="39A2651CF128FFFE2EB001C4FB9D2CAC" pageId="3" pageNumber="179" type="etymology">
<paragraph id="71073697F128FFFE2EB001C4FF592D84" blockId="3.[136,1452,1175,1490]" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2EB001C4FED22DAB" bold="true" box="[189,356,1212,1238]" pageId="3" pageNumber="179">Etymology:—</emphasis>
Named after the location of Xuan Lien,
<collectingRegion id="B37CF875F128FFFE2D3A01C3FB912DA8" box="[823,1063,1211,1238]" country="Vietnam" name="Thanh Hoa" pageId="3" pageNumber="179">Thanh Hoa Province</collectingRegion>
,
<collectingCountry id="09AF7607F128FFFE2A3801C3FB212DA8" box="[1077,1175,1211,1237]" name="Vietnam" pageId="3" pageNumber="179">Vietnam</collectingCountry>
, to which the species is endemic.
</paragraph>
<paragraph id="71073697F128FFFE2EB0007BFB9D2CAC" blockId="3.[136,1452,1175,1490]" pageId="3" pageNumber="179">
<emphasis id="43CCEA85F128FFFE2EB0007BFE652C63" bold="true" box="[189,467,1283,1310]" pageId="3" pageNumber="179">Conservation status:—</emphasis>
Within Xuan Lien Nature Reserve,
<taxonomicName id="B6B84D14F128FFFE2D79007BFB172C60" authorityName="N. T. T Huong, B. H. Quang &amp; J. S Ma" authorityYear="2014" box="[884,1185,1283,1309]" class="Magnoliopsida" family="Aristolochiaceae" genus="Aristolochia" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="179" phylum="Tracheophyta" rank="species" species="xuanlienensis">
<emphasis id="43CCEA85F128FFFE2D79007BFB172C60" box="[884,1185,1283,1309]" italics="true" pageId="3" pageNumber="179">Aristolochia xuanlienensis</emphasis>
</taxonomicName>
is known only from a single population, with fewer than five individuals observed among our two expeditions in the past few years. It is therefore assigned a preliminary status of Vulnerable (VU) according to IUCN Red List criteria (
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), which indicates a population with a very restricted area (typically less than
<quantity id="B6409B72F128FFFE2D8B0008FC7B2CF4" box="[902,973,1391,1418]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="179" unit="km" value="20.0">20 km</quantity>
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). More populations of this species remain to be found in similar habitats in the area. However, an extensive field survey is needed since the species is small liana that is inconspicuous in the forests and thus usually difficult to observe in the field.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,428 @@
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<emphasis id="43CCEAE9FFE92E3E75C6F91EFE2AF962" bold="true" box="[151,483,1703,1729]" italics="true" pageId="15" pageNumber="16">Eurycercus pompholygodes</emphasis>
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</paragraph>
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<emphasis id="43CCEAE9FFE92E3E75C6F8AAFE67F88B" box="[151,430,1810,1832]" italics="true" pageId="15" pageNumber="16">Eurycercus pompholygodes</emphasis>
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284,
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31;
<treatmentCitation id="F01910EAFFE92E3E764FF8AAFBEDF88B" author="Kotov, A. A. &amp; Sinev, A. Y. &amp; Glagolev, S. M. &amp; Smirnov, N. N." box="[798,1060,1810,1832]" page="234" pageId="15" pageNumber="16" year="2010">
<bibRefCitation id="15294B0AFFE92E3E764FF8AAFC12F88B" author="Kotov, A. A. &amp; Sinev, A. Y. &amp; Glagolev, S. M. &amp; Smirnov, N. N." box="[798,987,1810,1832]" pageId="15" pageNumber="16" pagination="151 - 276" refId="ref27377" refString="Kotov, A. A., Sinev, A. Y., Glagolev, S. M. &amp; Smirnov, N. N. (2010 b) Water fleas (Cladocera). In: Alexeev V. R., Tsalolokhin S. Y. (eds.), Key book for zooplankton and zoobenthos of fresh waters of European Russia. KMK, Moscow, pp. 151 - 276." type="book chapter" year="2010">
Kotov
<emphasis id="43CCEAE9FFE92E3E7633F8AAFC5DF88B" box="[866,916,1810,1832]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2010b
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, p. 234
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235, Fig. 137(1214).
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<emphasis id="43CCEAE9FFE92E3E75C6F8ECFEFCF8CD" bold="true" box="[151,309,1877,1902]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Type locality.</emphasis>
“Bythotrephes Pond, located about
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northwest of the limnological station at Kuokkel and about
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north of Björkstugan at the west end of Tome Träsk in Swedish Lapland&quot; (
<bibRefCitation id="15294B0AFFE92E3E716CF8C0FB7EF832" author="Frey, D. G." box="[1085,1207,1913,1937]" pageId="15" pageNumber="16" pagination="263 - 300" refId="ref25854" refString="Frey, D. G. (1975) Subgeneric differentiation within Eurycercus (Cladocera, Chydoridae) and a new species from Northern Sweden. Hydrobiologia, 46, 263 - 300." type="journal article" year="1975">Frey 1975</bibRefCitation>
). Approximate geographic coordinates: 68.4, 18.4
</paragraph>
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<emphasis id="43CCEAE9FFF62E2175C6F82BFEC4F80B" bold="true" box="[151,269,1938,1961]" pageId="16" pageNumber="17">FIGURE 5</emphasis>
.
<taxonomicName id="B6B84D78FFF62E21744DF82DFDFCF80A" authorityName="Frey" authorityYear="1975" box="[284,565,1939,1961]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pompholygodes">
<emphasis id="43CCEAE9FFF62E21744DF82DFDFCF80A" box="[284,565,1939,1961]" italics="true" pageId="16" pageNumber="17">Eurycercus pompholygodes</emphasis>
</taxonomicName>
, parthenogenetic female from Lake №6 near River Tareya, a tributary of River Pyasina, Taimyr Autonomous Area, Russia. A, lateral view. B, ventral view. C, anterior view. D, head. E, head pores. F, postabdomen. G, its distal portion. H, antenna I. IJ, inner distal lobe of limb I. Scale bar denotes 1 mm (AC); 0.1 mm (DJ).
</paragraph>
</caption>
<paragraph id="710736FBFFF72E207598FF21FB33FF0C" blockId="17.[151,1436,151,2012]" box="[201,1274,151,176]" pageId="17" pageNumber="18">
Type material.
<materialsCitation id="C1D03CA6FFF72E20742CFF21FB33FF0C" box="[381,1274,151,176]" collectionCode="NHM" pageId="17" pageNumber="18" specimenCode="NHM 1973.845" specimenCount="1" typeStatus="holotype">
<emphasis id="43CCEAE9FFF72E20742CFF21FE38FF13" bold="true" box="[381,497,152,176]" pageId="17" pageNumber="18">
<typeStatus id="AE038859FFF72E20742CFF21FE25FF13" box="[381,492,152,176]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
.
</emphasis>
A parthenogenetic female
<quantity id="B6409B1EFFF72E207670FF2EFC4DFF0C" box="[801,900,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="17" pageNumber="18" unit="mm" value="1.92">1.92 mm</quantity>
long in alcohol,
<specimenCode id="211E9E80FFF72E207110FF21FB3FFF0C" box="[1089,1270,151,175]" collectionCode="NHM" pageId="17" pageNumber="18">NHM 1973.845</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="710736FBFFF72E207598FF05FD27FE6C" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<materialsCitation id="C1D03CA6FFF72E207598FF05FB34FF54" collectionCode="NHM" pageId="17" pageNumber="18" specimenCode="NHM 1973.846" specimenCount="1" typeStatus="paratype">
<emphasis id="43CCEAE9FFF72E207598FF05FE81FF77" bold="true" box="[201,328,188,212]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">
<typeStatus id="AE038859FFF72E207598FF05FE8AFF77" box="[201,323,188,212]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Two parthenogenetic females in alcohol, one parthenogenetic female on a slide in glycerine jelly, and three exuviae mounted individually in polyvinyl lactophenol on the same slide,
<specimenCode id="211E9E80FFF72E207117FF59FB34FF54" box="[1094,1277,223,247]" collectionCode="NHM" pageId="17" pageNumber="18">NHM 1973.846</specimenCode>
</materialsCitation>
-
<materialsCitation id="C1D03CA6FFF72E207055FF66FAE6FF54" box="[1284,1327,223,247]" collectionCode="NHM" isTailEnumeration="true" pageId="17" pageNumber="18" specimenCode="849" specimenCount="1" typeStatus="paratype">
<specimenCode id="211E9E80FFF72E207055FF66FAE6FF54" box="[1284,1327,223,247]" collectionCode="NHM" pageId="17" pageNumber="18">849</specimenCode>
</materialsCitation>
.
<materialsCitation id="C1D03CA6FFF72E20706BFF66FC11FE9C" collectionCode="USNM" pageId="17" pageNumber="18" specimenCode="USNM 143768" specimenCount="1" typeStatus="paratype">
One parthenogenetic female in alcohol, one parthenogenetic female on a slide in glycerine jelly, and three exuviae mounted individually in polyvinyl lactophenol on the same slide,
<specimenCode id="211E9E80FFF72E207677FE9EFC11FE9C" box="[806,984,295,319]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM 143768</specimenCode>
</materialsCitation>
.
<materialsCitation id="C1D03CA6FFF72E2076B7FE9EFD23FE6C" collectionCode="ZIN, USNM, DGF" pageId="17" pageNumber="18" specimenCode="USNM 403774" specimenCount="1" typeStatus="paratype">
One parthenogenetic female and three exuviae in polyvinyl lactophenol on the same slide (
<collectionCode id="17A9AE3EFFF72E2077B0FEF5FCDDFEC7" box="[737,788,332,356]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="17" pageNumber="18" type="Museum">ZIN</collectionCode>
, accession numbers 2656 and 2667), one parthenogenetic female and three exuviae in polyvinyl lactophenol on the same slide,
<collectionCode id="17A9AE3EFFF72E20769AFEC9FC30FE24" box="[971,1017,368,391]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="17" pageNumber="18" type="Museum">ZIN</collectionCode>
(
<collectionCode id="17A9AE3EFFF72E20715CFED6FBF6FE24" box="[1037,1087,367,391]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="17" pageNumber="18" type="Museum">ZIN</collectionCode>
, accession numbers 2658 and 2659). Some other specimens were kept in the D. G. FREY collection in Bloomington, now they are in
<collectionCode id="17A9AE3EFFF72E20701AFE2DFA55FE0F" box="[1355,1436,404,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM</collectionCode>
(accession number of whole
<collectionCode id="17A9AE3EFFF72E207488FE0EFDD8FE6C" box="[473,529,439,463]" pageId="17" pageNumber="18">DGF</collectionCode>
is
<specimenCode id="211E9E80FFF72E207760FE0EFD2BFE6C" box="[561,738,439,463]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM 403774</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="710736FBFFF72E207598FE62FD09FD9F" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598FE62FE32FE57" bold="true" box="[201,507,475,500]" pageId="17" pageNumber="18">Material examined here:</emphasis>
<materialsCitation id="C1D03CA6FFF72E207758FE65FDFFFD9F" collectingDate="1972-07-20" collectingDateMax="1972-07-27" collectingDateMin="1972-07-20" collectionCode="DGF" collectorName="Abisko Turiststation &amp; D. G. Frey &amp; Bythotrephes &amp; Abisko" country="Sweden" elevation="8" latitude="68.358" location="Small" longLatPrecision="59" longitude="18.783" municipality="Kuokkel" pageId="17" pageNumber="18" specimenCode="DGF 3260, DGF 3268" specimenCount="1">
<emphasis id="43CCEAE9FFF72E207758FE65FDAFFE57" box="[521,614,476,500]" italics="true" pageId="17" pageNumber="18">
<collectingCountry id="09AF766BFFF72E207758FE65FDABFE57" box="[521,610,476,500]" name="Sweden" pageId="17" pageNumber="18">Sweden</collectingCountry>
.
</emphasis>
<location id="74676020FFF72E207723FE65FD7AFE57" LSID="urn:lsid:plazi:treatment:F91187EDFFE92E2375C6F91EFB9FFDB4:74676020FFF72E207723FE65FD7AFE57" box="[626,691,476,500]" country="Sweden" latitude="68.358" longLatPrecision="59" longitude="18.783" municipality="Kuokkel" name="Small" pageId="17" pageNumber="18">Small</location>
pond ca.
<quantity id="B6409B1EFFF72E207663FE65FCAAFE50" box="[818,867,476,500]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="18" unit="m" value="8.0">
<elevation id="FA95D1C8FFF72E207663FE65FCAAFE50" box="[818,867,476,500]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="18" unit="m" value="8.0">8 m</elevation>
</quantity>
within
<quantity id="B6409B1EFFF72E207694FE65FBD8FE50" box="[965,1041,476,500]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="100.0">
<elevation id="FA95D1C8FFF72E207694FE65FBD8FE50" box="[965,1041,476,500]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="100.0">100 m</elevation>
</quantity>
of
<collectorName id="DC4D532DFFF72E207113FE65FAF9FE57" box="[1090,1328,476,500]" pageId="17" pageNumber="18">Abisko Turiststation</collectorName>
, coll. in.
<date id="0506103BFFF72E2075C6FE46FEDCFDB4" box="[151,277,511,535]" pageId="17" pageNumber="18" value="1972-07-20">
<collectingDate id="1542E9D3FFF72E2075C6FE46FEDCFDB4" box="[151,277,511,535]" pageId="17" pageNumber="18" value="1972-07-20">20.07.1972</collectingDate>
</date>
by
<collectorName id="DC4D532DFFF72E207417FDB9FE0BFDBB" box="[326,450,511,536]" pageId="17" pageNumber="18">D. G. Frey</collectorName>
,
<specimenCode id="211E9E80FFF72E207480FE46FD82FDB4" box="[465,587,511,535]" collectionCode="DGF" pageId="17" pageNumber="18">DGF 3260</specimenCode>
(
<geoCoordinate id="148C503CFFF72E20770CFE46FD67FDB4" box="[605,686,511,535]" degrees="68.358" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="55" value="68.358">68.358</geoCoordinate>
,
<geoCoordinate id="148C503CFFF72E2077EFFE46FCC6FDB4" box="[702,783,511,535]" degrees="18.783" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="55" value="18.783">18.783</geoCoordinate>
);
<collectorName id="DC4D532DFFF72E207677FE46FC72FDB4" box="[806,955,511,535]" pageId="17" pageNumber="18">Bythotrephes</collectorName>
pond, north of
<collectingMunicipality id="9163AC81FFF72E207127FE46FB10FDB4" box="[1142,1241,511,535]" pageId="17" pageNumber="18">Kuokkel</collectingMunicipality>
,
<collectorName id="DC4D532DFFF72E2071B9FE46FAF5FDB4" box="[1256,1340,511,535]" pageId="17" pageNumber="18">Abisko</collectorName>
, coll. in
<date id="0506103BFFF72E2075C6FD9DFEDCFD9F" box="[151,277,548,572]" pageId="17" pageNumber="18" value="1972-07-27">
<collectingDate id="1542E9D3FFF72E2075C6FD9DFEDCFD9F" box="[151,277,548,572]" pageId="17" pageNumber="18" value="1972-07-27">27.07.1972</collectingDate>
</date>
by
<collectorName id="DC4D532DFFF72E20746EFD9DFE7DFD9F" box="[319,436,548,572]" pageId="17" pageNumber="18">D. G. Frey</collectorName>
,
<specimenCode id="211E9E80FFF72E2074EEFD9DFDFFFD9F" box="[447,566,548,572]" collectionCode="DGF" pageId="17" pageNumber="18">DGF 3268</specimenCode>
</materialsCitation>
(68.4, 18.4)
</paragraph>
<paragraph id="710736FBFFF72E207598FDF1FCBCFC94" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<materialsCitation id="C1D03CA6FFF72E207598FDF1FE57FD04" collectingDate="1982-10-08" collectingDateMax="2010-07-20" collectingDateMin="1982-10-08" collectorName="Autonomous Republic. A &amp; Pechora-Ilych National &amp; A. A. Kalinin &amp; Scheljajur" country="Russia" latitude="63.1514" location="Pechora-Ilych National Reserve" longLatPrecision="5" longitude="59.0225" municipality="River" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Komi">
<emphasis id="43CCEAE9FFF72E207598FDF1FDACFDC3" box="[201,613,584,608]" italics="true" pageId="17" pageNumber="18">
<collectingCountry id="09AF766BFFF72E207598FDF1FEDEFDFC" box="[201,279,584,607]" name="Russia" pageId="17" pageNumber="18">Russia</collectingCountry>
.
<collectingRegion id="B37CF819FFF72E207473FDF1FE97FDFC" box="[290,350,584,607]" country="Russia" name="Komi" pageId="17" pageNumber="18">Komi</collectingRegion>
<collectorName id="DC4D532DFFF72E207437FDF1FDACFDC3" box="[358,613,584,608]" pageId="17" pageNumber="18">Autonomous Republic.</collectorName>
</emphasis>
A bay of the Kozhim-Yu
<collectingMunicipality id="9163AC81FFF72E2076DDFDFEFC05FDFC" box="[908,972,583,607]" pageId="17" pageNumber="18">River</collectingMunicipality>
,
<location id="74676020FFF72E207689FDFEFA8BFDFC" LSID="urn:lsid:plazi:treatment:F91187EDFFE92E2375C6F91EFB9FFDB4:74676020FFF72E207689FDFEFA8BFDFC" box="[984,1346,583,607]" country="Russia" latitude="63.1514" longLatPrecision="5" longitude="59.0225" municipality="River" name="Pechora-Ilych National Reserve" pageId="17" pageNumber="18" stateProvince="Komi">
<collectorName id="DC4D532DFFF72E207689FDFEFB15FDFC" box="[984,1244,583,607]" pageId="17" pageNumber="18">Pechora-Ilych National</collectorName>
Reserve
</location>
, coll. in
<date id="0506103BFFF72E2075C6FDD5FEDCFD27" box="[151,277,620,644]" pageId="17" pageNumber="18" value="2010-07-20">
<collectingDate id="1542E9D3FFF72E2075C6FDD5FEDCFD27" box="[151,277,620,644]" pageId="17" pageNumber="18" value="2010-07-20">20.07.2010</collectingDate>
</date>
by
<collectorName id="DC4D532DFFF72E207410FDD5FE16FD27" box="[321,479,620,644]" pageId="17" pageNumber="18">A. A. Kalinin</collectorName>
, AAK M-1619 (
<geoCoordinate id="148C503CFFF72E2077CAFDD5FD33FD27" box="[667,762,620,644]" degrees="63.1514" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="5" value="63.1514">63.1514</geoCoordinate>
,
<geoCoordinate id="148C503CFFF72E207657FDD5FCAFFD27" box="[774,870,620,644]" degrees="59.0225" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="5" value="59.0225">59.0225</geoCoordinate>
); Kurja near
<collectorName id="DC4D532DFFF72E2076AAFDD5FBA6FD27" box="[1019,1135,620,644]" pageId="17" pageNumber="18">Scheljajur</collectorName>
, coll. in
<date id="0506103BFFF72E207183FDD5FA9AFD27" box="[1234,1363,620,644]" pageId="17" pageNumber="18" value="1982-10-08">
<collectingDate id="1542E9D3FFF72E207183FDD5FA9AFD27" box="[1234,1363,620,644]" pageId="17" pageNumber="18" value="1982-10-08">10.08.1982</collectingDate>
</date>
, AAK 1999-043 (65.33, 53.4)
</materialsCitation>
.
<materialsCitation id="C1D03CA6FFF72E2074F8FD29FB3CFD04" box="[425,1269,655,680]" collectorName="Area. A" country="Russia" location="Belomorskaya Biological Station" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Murmansk">
<emphasis id="43CCEAE9FFF72E2074F8FD29FD94FD0B" box="[425,605,656,680]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="B37CF819FFF72E2074F8FD29FDE8FD0B" box="[425,545,656,680]" country="Russia" name="Murmansk" pageId="17" pageNumber="18">Murmansk</collectingRegion>
<collectorName id="DC4D532DFFF72E207779FD29FD94FD0B" box="[552,605,656,680]" pageId="17" pageNumber="18">Area</collectorName>
</emphasis>
. A small lake near the
<location id="74676020FFF72E207632FD36FB1EFD04" LSID="urn:lsid:plazi:treatment:F91187EDFFE92E2375C6F91EFB9FFDB4:74676020FFF72E207632FD36FB1EFD04" box="[867,1239,655,679]" country="Russia" name="Belomorskaya Biological Station" pageId="17" pageNumber="18" stateProvince="Murmansk">Belomorskaya Biological Station</location>
of
</materialsCitation>
<materialsCitation id="C1D03CA6FFF72E2071ACFD29FCB8FC94" collectingDate="1974-11-08" collectingDateMax="2009-01-08" collectingDateMin="1974-11-08" collectorName="University &amp; A. Yu. Sinev &amp; Taimyr Autonomous Area. A &amp; Talnah &amp; Y. I. Chernov &amp; River" country="Russia" county="Lake Bolshoy Vudjavr" latitude="67.6306" location="River Tareya" longLatPrecision="22" longitude="33.673" municipality="Taimyr Autonomous Area" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Moscow">
<collectingRegion id="B37CF819FFF72E2071ACFD29FA55FD04" box="[1277,1436,655,679]" country="Russia" name="Moscow" pageId="17" pageNumber="18">Moscow State</collectingRegion>
<collectorName id="DC4D532DFFF72E2075C6FD0DFEC7FD6F" box="[151,270,692,716]" pageId="17" pageNumber="18">University</collectorName>
, coll. in 08. 1995 by
<collectorName id="DC4D532DFFF72E20774FFD0DFD72FD6F" box="[542,699,692,716]" pageId="17" pageNumber="18">A. Yu. Sinev</collectorName>
, AAK 2004-020 (66.5, 33.1);
<collectingCounty id="98664E77FFF72E20717DFD0DFAFEFD6F" box="[1068,1335,692,716]" pageId="17" pageNumber="18">Lake Bolshoy Vudjavr</collectingCounty>
, coll. in
<date id="0506103BFFF72E2075C6FD6EFEDEFD4C" box="[151,279,727,751]" pageId="17" pageNumber="18" value="2009-01-08">
<collectingDate id="1542E9D3FFF72E2075C6FD6EFEDEFD4C" box="[151,279,727,751]" pageId="17" pageNumber="18" value="2009-01-08">01.08.2009</collectingDate>
</date>
, AAK M-1404 (
<geoCoordinate id="148C503CFFF72E20749EFD6EFDE6FD4C" box="[463,559,727,751]" degrees="67.6306" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="5" value="67.6306">67.6306</geoCoordinate>
,
<geoCoordinate id="148C503CFFF72E207768FD6EFD42FD4C" box="[569,651,727,751]" degrees="33.673" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="55" value="33.673">33.673</geoCoordinate>
).
<collectorName id="DC4D532DFFF72E2077CCFD61FC15FD4C" box="[669,988,727,752]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E2077CCFD61FC0BFD53" box="[669,962,728,752]" italics="true" pageId="17" pageNumber="18">
<collectingMunicipality id="9163AC81FFF72E2077CCFD61FC77FD53" box="[669,958,728,752]" pageId="17" pageNumber="18">Taimyr Autonomous Area</collectingMunicipality>
.
</emphasis>
A
</collectorName>
tarn at
<collectorName id="DC4D532DFFF72E207161FD6EFBB6FD4C" box="[1072,1151,727,751]" pageId="17" pageNumber="18">Talnah</collectorName>
, coll. in
<date id="0506103BFFF72E20718FFD6EFA93FD4C" box="[1246,1370,727,751]" pageId="17" pageNumber="18" value="1974-11-08">
<collectingDate id="1542E9D3FFF72E20718FFD6EFA93FD4C" box="[1246,1370,727,751]" pageId="17" pageNumber="18" value="1974-11-08">11.08.1974</collectingDate>
</date>
by
<collectorName id="DC4D532DFFF72E2070D5FD61FED9FCB7" pageId="17" pageNumber="18">Y. I. Chernov</collectorName>
, AAK 1999-075 and AAK 1999-049 (69.5, 88.4); Lake №6,
<location id="74676020FFF72E2076EEFD45FB9DFCB7" LSID="urn:lsid:plazi:treatment:F91187EDFFE92E2375C6F91EFB9FFDB4:74676020FFF72E2076EEFD45FB9DFCB7" box="[959,1108,764,788]" country="Russia" county="Lake Bolshoy Vudjavr" latitude="67.6306" longLatPrecision="22" longitude="33.673" municipality="Taimyr Autonomous Area" name="River Tareya" pageId="17" pageNumber="18" stateProvince="Moscow">River Tareya</location>
, a tributary of
<location id="74676020FFF72E2071A9FD45FA51FCB7" LSID="urn:lsid:plazi:treatment:F91187EDFFE92E2375C6F91EFB9FFDB4:74676020FFF72E2071A9FD45FA51FCB7" box="[1272,1432,764,788]" country="Russia" county="Lake Bolshoy Vudjavr" latitude="67.6306" longLatPrecision="22" longitude="33.673" municipality="Taimyr Autonomous Area" name="River Pyasina" pageId="17" pageNumber="18" stateProvince="Moscow">
<collectorName id="DC4D532DFFF72E2071A9FD45FAFEFCB7" box="[1272,1335,764,788]" pageId="17" pageNumber="18">River</collectorName>
Pyasina
</location>
, coll. in
<date id="0506103BFFF72E2075BAFCA6FEA1FC94" box="[235,360,799,823]" pageId="17" pageNumber="18" value="1974-11-08">
<collectingDate id="1542E9D3FFF72E2075BAFCA6FEA1FC94" box="[235,360,799,823]" pageId="17" pageNumber="18" value="1974-11-08">11.08.1974</collectingDate>
</date>
by
<collectorName id="DC4D532DFFF72E2074C3FC99FDE3FC94" box="[402,554,799,823]" pageId="17" pageNumber="18">Y. I. Chernov</collectorName>
, AAK 1999-074 (73.3, 90.8)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF72E237598FCFAFC1DFF54" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="710736FBFFF72E207598FCFAFE0AFB3C" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598FCFAFDA7FCFF" bold="true" box="[201,622,835,860]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598FCFAFE8AFCFF" box="[201,323,835,860]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">Diagnosis.</emphasis>
Parthenogenetic female.
</emphasis>
Dorsal head pores on a bubble-like projection located immediately on head shield. In anterior view, body wide, not compressed laterally, median dorsal keel absent. Rostrum relatively long. Ocellus minute. Lateral head pore elongated. Labrum with a relatively small median keel, broadly rounded, reaching only half of antenna I length. Postabdomen with sub-parallel dorsal and ventral margins, preanal teeth pointed. Spines at base of pre-claw portion predominantly clustered. Antenna I with antennular sensory seta arising in middle. Denticles in rows encircling antennular surface especially small. On antenna II, spine situated on proximal segment of exopod somewhat shorter than second segment. Limb I IDL with a remarkably strong hook-like seta, the smallest IDL seta especially fine and short. IDL with 64 distal spinules, 610 proximal spinules, 36 marginal spinules and 12 basal spinules. Eight setae in filter plate II, 8 setae in filter plate III; 9 setae in filter plate IV, 8 setae in filter plate V.
</paragraph>
<paragraph id="710736FBFFF72E207598FB15FB61FB67" blockId="17.[151,1436,151,2012]" box="[201,1192,1195,1220]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598FB15FC2CFB67" bold="true" box="[201,997,1195,1220]" pageId="17" pageNumber="18">Full description of parthenogenetic female, ephippial female, male.</emphasis>
See
<bibRefCitation id="15294B0AFFF72E20714DFB15FB6DFB67" author="Frey, D. G." box="[1052,1188,1196,1220]" pageId="17" pageNumber="18" pagination="263 - 300" refId="ref25854" refString="Frey, D. G. (1975) Subgeneric differentiation within Eurycercus (Cladocera, Chydoridae) and a new species from Northern Sweden. Hydrobiologia, 46, 263 - 300." type="journal article" year="1975">Frey (1975)</bibRefCitation>
.
</paragraph>
<paragraph id="710736FBFFF72E207598FB76FA42FAF7" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598FB76FD2CFB4B" bold="true" box="[201,741,1231,1256]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598FB76FE73FB4B" box="[201,442,1231,1256]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">Short redescription.</emphasis>
Parthenogenetic female.
</emphasis>
In lateral view (
<figureCitation id="E9832A7EFFF72E2076F4FB69FC36FB4B" box="[933,1023,1232,1256]" captionStart="FIGURE 5" captionStartId="16.[151,250,1938,1961]" captionTargetBox="[196,1396,197,1858]" captionTargetId="figure-86@16.[188,1398,193,1918]" captionTargetPageId="16" captionText="FIGURE 5. Eurycercus pompholygodes, parthenogenetic female from Lake №6 near River Tareya, a tributary of River Pyasina, Taimyr Autonomous Area, Russia. A, lateral view. B, ventral view. C, anterior view. D, head. E, head pores. F, postabdomen. G, its distal portion. H, antenna I. IJ, inner distal lobe of limb I. Scale bar denotes 1 mm (AC); 0.1 mm (DJ)." figureDoi="http://doi.org/10.5281/zenodo.5248221" httpUri="https://zenodo.org/record/5248221/files/figure.png" pageId="17" pageNumber="18">Fig. 5A</figureCitation>
) body in larger females ovoid (BH/ BL=0.600.66), but in juveniles more sub-rectangular, than in other species. Dorsal margin interrupted by a dorsal bubble bearing head pores. In anterior of ventral view, body wide, not compressed laterally, median dorsal keel is absent (
<figureCitation id="E9832A7EFFF72E2075BFFA85FEAEFAF7" box="[238,359,1340,1364]" captionStart="FIGURE 5" captionStartId="16.[151,250,1938,1961]" captionTargetBox="[196,1396,197,1858]" captionTargetId="figure-86@16.[188,1398,193,1918]" captionTargetPageId="16" captionText="FIGURE 5. Eurycercus pompholygodes, parthenogenetic female from Lake №6 near River Tareya, a tributary of River Pyasina, Taimyr Autonomous Area, Russia. A, lateral view. B, ventral view. C, anterior view. D, head. E, head pores. F, postabdomen. G, its distal portion. H, antenna I. IJ, inner distal lobe of limb I. Scale bar denotes 1 mm (AC); 0.1 mm (DJ)." figureDoi="http://doi.org/10.5281/zenodo.5248221" httpUri="https://zenodo.org/record/5248221/files/figure.png" pageId="17" pageNumber="18">Fig. 5BC</figureCitation>
). Intestine has a single loop, posterior intestinal caecum present. Few eggs in the brood pouch.
</paragraph>
<paragraph id="710736FBFFF72E207598FAD9FBA3FA3F" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598FAD9FECFFADB" box="[201,262,1376,1400]" italics="true" pageId="17" pageNumber="18">Head</emphasis>
with short rostrum (
<figureCitation id="E9832A7EFFF72E2074BDFAD9FD8FFADB" box="[492,582,1376,1400]" captionStart="FIGURE 5" captionStartId="16.[151,250,1938,1961]" captionTargetBox="[196,1396,197,1858]" captionTargetId="figure-86@16.[188,1398,193,1918]" captionTargetPageId="16" captionText="FIGURE 5. Eurycercus pompholygodes, parthenogenetic female from Lake №6 near River Tareya, a tributary of River Pyasina, Taimyr Autonomous Area, Russia. A, lateral view. B, ventral view. C, anterior view. D, head. E, head pores. F, postabdomen. G, its distal portion. H, antenna I. IJ, inner distal lobe of limb I. Scale bar denotes 1 mm (AC); 0.1 mm (DJ)." figureDoi="http://doi.org/10.5281/zenodo.5248221" httpUri="https://zenodo.org/record/5248221/files/figure.png" pageId="17" pageNumber="18">Fig. 5D</figureCitation>
). A single major “head pore” as a ringed, sub-oval field of special cuticle; a minute, elongated lateral pore located at either side of major pore, closer to it (
<figureCitation id="E9832A7EFFF72E207157FA3DFB95FA3F" box="[1030,1116,1412,1436]" captionStart="FIGURE 5" captionStartId="16.[151,250,1938,1961]" captionTargetBox="[196,1396,197,1858]" captionTargetId="figure-86@16.[188,1398,193,1918]" captionTargetPageId="16" captionText="FIGURE 5. Eurycercus pompholygodes, parthenogenetic female from Lake №6 near River Tareya, a tributary of River Pyasina, Taimyr Autonomous Area, Russia. A, lateral view. B, ventral view. C, anterior view. D, head. E, head pores. F, postabdomen. G, its distal portion. H, antenna I. IJ, inner distal lobe of limb I. Scale bar denotes 1 mm (AC); 0.1 mm (DJ)." figureDoi="http://doi.org/10.5281/zenodo.5248221" httpUri="https://zenodo.org/record/5248221/files/figure.png" pageId="17" pageNumber="18">Fig. 5E</figureCitation>
).
</paragraph>
<paragraph id="710736FBFFF72E207598FA11FAD9FA63" blockId="17.[151,1436,151,2012]" box="[201,1296,1448,1472]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598FA11FEEBFA63" box="[201,290,1448,1472]" italics="true" pageId="17" pageNumber="18">Labrum</emphasis>
with a small, rounded median keel, reaching only about middle of antenna I (
<figureCitation id="E9832A7EFFF72E2071D6FA11FACAFA63" box="[1159,1283,1448,1472]" captionStart="FIGURE 5" captionStartId="16.[151,250,1938,1961]" captionTargetBox="[196,1396,197,1858]" captionTargetId="figure-86@16.[188,1398,193,1918]" captionTargetPageId="16" captionText="FIGURE 5. Eurycercus pompholygodes, parthenogenetic female from Lake №6 near River Tareya, a tributary of River Pyasina, Taimyr Autonomous Area, Russia. A, lateral view. B, ventral view. C, anterior view. D, head. E, head pores. F, postabdomen. G, its distal portion. H, antenna I. IJ, inner distal lobe of limb I. Scale bar denotes 1 mm (AC); 0.1 mm (DJ)." figureDoi="http://doi.org/10.5281/zenodo.5248221" httpUri="https://zenodo.org/record/5248221/files/figure.png" pageId="17" pageNumber="18">Fig. 5A, D</figureCitation>
).
</paragraph>
<paragraph id="710736FBFFF72E207598FA75FEABF9AB" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598FA75FE5EFA47" box="[201,407,1484,1508]" italics="true" pageId="17" pageNumber="18">Armature of valve</emphasis>
margin as in previous species (VL/BL = 0.810.86), with sub-parallel dorsal and ventral margins (
<figureCitation id="E9832A7EFFF72E207451FA49FE9CF9AB" box="[256,341,1520,1544]" captionStart="FIGURE 5" captionStartId="16.[151,250,1938,1961]" captionTargetBox="[196,1396,197,1858]" captionTargetId="figure-86@16.[188,1398,193,1918]" captionTargetPageId="16" captionText="FIGURE 5. Eurycercus pompholygodes, parthenogenetic female from Lake №6 near River Tareya, a tributary of River Pyasina, Taimyr Autonomous Area, Russia. A, lateral view. B, ventral view. C, anterior view. D, head. E, head pores. F, postabdomen. G, its distal portion. H, antenna I. IJ, inner distal lobe of limb I. Scale bar denotes 1 mm (AC); 0.1 mm (DJ)." figureDoi="http://doi.org/10.5281/zenodo.5248221" httpUri="https://zenodo.org/record/5248221/files/figure.png" pageId="17" pageNumber="18">Fig. 5F</figureCitation>
).
</paragraph>
<paragraph id="710736FBFFF72E207598F9ADFCAAF943" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598F9ADFEA0F98F" box="[201,361,1556,1580]" italics="true" pageId="17" pageNumber="18">Postabdomen.</emphasis>
Distal anal embayment shallow. Armature of the preanal margin as a series of preanal teeth (NT about 83), distalmost tooth remarkably larger than others; small gap lacking any teeth at base of postabdominal setae; teeth in middle of preanal margin with sharp tips. Distalmost postanal teeth on pre-claw portion long, thin, predominantly double, teeth on pre-claw portion shorter, generally clustered (doubled or tripled) (
<figureCitation id="E9832A7EFFF72E20718FF939FAFDF93B" box="[1246,1332,1664,1688]" captionStart="FIGURE 5" captionStartId="16.[151,250,1938,1961]" captionTargetBox="[196,1396,197,1858]" captionTargetId="figure-86@16.[188,1398,193,1918]" captionTargetPageId="16" captionText="FIGURE 5. Eurycercus pompholygodes, parthenogenetic female from Lake №6 near River Tareya, a tributary of River Pyasina, Taimyr Autonomous Area, Russia. A, lateral view. B, ventral view. C, anterior view. D, head. E, head pores. F, postabdomen. G, its distal portion. H, antenna I. IJ, inner distal lobe of limb I. Scale bar denotes 1 mm (AC); 0.1 mm (DJ)." figureDoi="http://doi.org/10.5281/zenodo.5248221" httpUri="https://zenodo.org/record/5248221/files/figure.png" pageId="17" pageNumber="18">Fig. 5G</figureCitation>
, arrows). Postabdominal claw robust (CL/PL = 0.260.30); first (distalmost) basal spine long (DS/CL = 0.360.50), second (basalmost) spine short (BS/CL = 0.180.25; BS/DS about 0.5).
</paragraph>
<paragraph id="710736FBFFF72E207598F955FE78F8D3" blockId="17.[151,1436,151,2012]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598F955FEF1F8A7" box="[201,312,1772,1796]" italics="true" pageId="17" pageNumber="18">Antenna I</emphasis>
(
<figureCitation id="E9832A7EFFF72E20741BF955FE6AF8A7" box="[330,419,1772,1796]" captionStart="FIGURE 5" captionStartId="16.[151,250,1938,1961]" captionTargetBox="[196,1396,197,1858]" captionTargetId="figure-86@16.[188,1398,193,1918]" captionTargetPageId="16" captionText="FIGURE 5. Eurycercus pompholygodes, parthenogenetic female from Lake №6 near River Tareya, a tributary of River Pyasina, Taimyr Autonomous Area, Russia. A, lateral view. B, ventral view. C, anterior view. D, head. E, head pores. F, postabdomen. G, its distal portion. H, antenna I. IJ, inner distal lobe of limb I. Scale bar denotes 1 mm (AC); 0.1 mm (DJ)." figureDoi="http://doi.org/10.5281/zenodo.5248221" httpUri="https://zenodo.org/record/5248221/files/figure.png" pageId="17" pageNumber="18">Fig. 5H</figureCitation>
) short relatively body length (AL/BL = 0.110.13), but protruding greatly beyond labral keel and tip of rostrum; elongated (AL/DA = 2.33.3), triangular in cross section. Antennular sensory seta arising near the middle of the antenna I. Denticles in rows encircling antennular surface especially small, ridges on the body of antenna I absent.
</paragraph>
<paragraph id="710736FBFFF72E207598F8C5FA4EF837" blockId="17.[151,1436,151,2012]" box="[201,1415,1916,1940]" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598F8C5FE89F837" box="[201,320,1916,1940]" italics="true" pageId="17" pageNumber="18">Antenna II</emphasis>
. Spine situated on proximal segment of exopod somewhat shorter than length of second segment.
</paragraph>
<paragraph id="710736FBFFF72E237598F819FBFCFF70" blockId="17.[151,1436,151,2012]" lastBlockId="18.[151,1436,151,536]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="43CCEAE9FFF72E207598F819FEDDF81B" box="[201,276,1952,1976]" italics="true" pageId="17" pageNumber="18">Limb I</emphasis>
with IDL supplied with three bisegmented setae generally decreasing in size towards endites, among them, a strong hook-like seta, but not too large like one in
<taxonomicName id="B6B84D78FFF72E207615F87CFBCDF87F" authorityName="Frey" authorityYear="1973" box="[836,1028,1988,2012]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="macracanthus">
<emphasis id="43CCEAE9FFF72E207615F87CFBCDF87F" box="[836,1028,1988,2012]" italics="true" pageId="17" pageNumber="18">E. macracanthus</emphasis>
</taxonomicName>
. IDL supplied with 46 long distal spinules, 610 long proximal spinules, 36 short marginal spinules and 12 very short basal spinules. Other limbs as in previous species, filter plate III with 910 setae, IV and V limbs with 8 setae.
</paragraph>
<paragraph id="710736FBFFF42E237598FF66FC1DFF54" blockId="18.[151,1436,151,536]" box="[201,980,223,248]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598FF66FEC9FF5B" bold="true" box="[201,256,223,248]" pageId="18" pageNumber="19">Size.</emphasis>
0.6901.67 in our material, 0.622.36 according to
<bibRefCitation id="15294B0AFFF42E237619FF59FC18FF54" author="Frey, D. G." box="[840,977,223,248]" pageId="18" pageNumber="19" pagination="263 - 300" refId="ref25854" refString="Frey, D. G. (1975) Subgeneric differentiation within Eurycercus (Cladocera, Chydoridae) and a new species from Northern Sweden. Hydrobiologia, 46, 263 - 300." type="journal article" year="1975">Frey (1975)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF42E237598FEBAFB12FEBF" box="[201,1243,259,284]" pageId="18" pageNumber="19" type="discussion">
<paragraph id="710736FBFFF42E237598FEBAFB12FEBF" blockId="18.[151,1436,151,536]" box="[201,1243,259,284]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598FEBAFE98FEBF" bold="true" box="[201,337,259,284]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Comments.</emphasis>
It is a well-defined Arctic-Subarctic taxon with a characteristic short labral keel.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF42E237598FE9EFB4CFE24" pageId="18" pageNumber="19" type="distribution">
<paragraph id="710736FBFFF42E237598FE9EFB4CFE24" blockId="18.[151,1436,151,536]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598FE9EFEABFEE3" bold="true" box="[201,354,295,320]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Distribution.</emphasis>
This taxon was known only from Swedish Lapland (
<bibRefCitation id="15294B0AFFF42E2376EAFE9EFBF8FE9C" author="Frey, D. G." box="[955,1073,295,319]" pageId="18" pageNumber="19" pagination="263 - 300" refId="ref25854" refString="Frey, D. G. (1975) Subgeneric differentiation within Eurycercus (Cladocera, Chydoridae) and a new species from Northern Sweden. Hydrobiologia, 46, 263 - 300." type="journal article" year="1975">Frey 1975</bibRefCitation>
) and a few localities in
<collectingCountry id="09AF766BFFF42E237013FE91FA55FEE3" box="[1346,1436,296,320]" name="Norway" pageId="18" pageNumber="19">Norway</collectingCountry>
(
<bibRefCitation id="15294B0AFFF42E2375CFFEF5FE70FEC7" author="Hessen, D. O. &amp; Walseng, B." box="[158,441,332,356]" pageId="18" pageNumber="19" pagination="2026 - 2035" refId="ref26487" refString="Hessen, D. O. &amp; Walseng, B. (2008) The rarity concept and the commonness of rarity in freshwater zooplankton. Freshwater Biology, 53, 2026 - 2035." type="journal article" year="2008">Hessen &amp; Walseng 2008</bibRefCitation>
). Now it is clear that
<taxonomicName id="B6B84D78FFF42E2377F6FEF4FCBFFEC7" authorityName="Frey" authorityYear="1975" box="[679,886,332,356]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pompholygodes">
<emphasis id="43CCEAE9FFF42E2377F6FEF4FCBFFEC7" box="[679,886,332,356]" italics="true" pageId="18" pageNumber="19">E. pompholygodes</emphasis>
</taxonomicName>
occurs in the north of European
<collectingCountry id="09AF766BFFF42E2371B7FEF5FAF8FEC7" box="[1254,1329,332,356]" name="Russia" pageId="18" pageNumber="19">Russia</collectingCountry>
and even on the Taimyr Peninsula. The range could reach the eastern part of the Siberian Subarctic.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF42E237598FE2DFB9FFDB4" pageId="18" pageNumber="19" type="biology_ecology">
<paragraph id="710736FBFFF42E237598FE2DFB9FFDB4" blockId="18.[151,1436,151,536]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598FE2DFEE4FE0F" bold="true" box="[201,301,404,428]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Ecology.</emphasis>
<bibRefCitation id="15294B0AFFF42E237465FE2DFE78FE0F" author="Frey, D. G." box="[308,433,404,428]" pageId="18" pageNumber="19" pagination="263 - 300" refId="ref25854" refString="Frey, D. G. (1975) Subgeneric differentiation within Eurycercus (Cladocera, Chydoridae) and a new species from Northern Sweden. Hydrobiologia, 46, 263 - 300." type="journal article" year="1975">Frey (1975</bibRefCitation>
, p. 265266) wrote that the species is present in the waters bodies where &quot;water has a low concentration of electrolytes, tends to be acidic, and quite frequently has a perceptible humic color&quot;, and that it sometimes co-occurs with
<taxonomicName id="B6B84D78FFF42E237497FE64FD95FE57" baseAuthorityName="O. F. Muller" baseAuthorityYear="1776" box="[454,604,476,500]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lamellatus">
<emphasis id="43CCEAE9FFF42E237497FE64FD95FE57" box="[454,604,476,500]" italics="true" pageId="18" pageNumber="19">E. lamellatus</emphasis>
</taxonomicName>
. We found that
<taxonomicName id="B6B84D78FFF42E237645FE64FC2DFE57" authorityName="Frey" authorityYear="1975" box="[788,996,476,500]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pompholygodes">
<emphasis id="43CCEAE9FFF42E237645FE64FC2DFE57" box="[788,996,476,500]" italics="true" pageId="18" pageNumber="19">E. pompholygodes</emphasis>
</taxonomicName>
is present in subarctic water bodies of different
<typeStatus id="AE038859FFF42E2375AFFDB9FEF4FDBB" box="[254,317,512,536]" pageId="18" pageNumber="19">types</typeStatus>
: tundra ponds of different size, small or large lakes, and bays of rivers.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="E1A1DD53626ED504DBBBE1C16457ACB6">A revision of the subgenus Eurycercus (Eurycercus) Baird, 1843 emend. nov. (Cladocera: Eurycercidae) in the Holarctic with the description of a new species from Alaska</mods:title>
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<mods:title id="4CAA3696967699F1F27CC0246540B2D9">Zootaxa</mods:title>
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<mods:date id="64C2D94FBCA879AD606282DD104F61AC">2012</mods:date>
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<mods:number id="5B148EA172DEC9530F3F0F958B19E21B">2012-02-24</mods:number>
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<mods:number id="C7D2604A37E162C74EE0878323297B1E">3206</mods:number>
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<mods:number id="30FCB4A8DA89E3AD623FEAB2C82903C6">1</mods:number>
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<mods:url id="ACEED97E66DA4A8C8DDDA9231F8B30C6">https://biotaxa.org/Zootaxa/article/view/zootaxa.3206.1.1</mods:url>
</mods:location>
<mods:classification id="86ABF5BE921021C61BA23B08EF777410">journal article</mods:classification>
<mods:identifier id="AABD160665F28AE2B9C144403C7B46D5" type="DOI">10.11646/zootaxa.3206.1.1</mods:identifier>
<mods:identifier id="95D6256BD5D207D2A3EA39DE5F1BD888" type="ISSN">1175-5326</mods:identifier>
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<treatment id="F91187EDFFF12E2875C6FACFFB90FE77" ID-DOI="http://doi.org/10.5281/zenodo.5249600" ID-Zenodo-Dep="5249600" LSID="urn:lsid:plazi:treatment:F91187EDFFF12E2875C6FACFFB90FE77" httpUri="http://treatment.plazi.org/id/F91187EDFFF12E2875C6FACFFB90FE77" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection id="39A26570FFF12E2675C6FACFFD0AFA33" box="[151,707,1398,1424]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="710736FBFFF12E2675C6FACFFD0AFA33" blockId="23.[151,707,1398,1457]" box="[151,707,1398,1424]" pageId="23" pageNumber="24">
<heading id="2A4F8197FFF12E2675C6FACFFD0AFA33" bold="true" box="[151,707,1398,1424]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<taxonomicName id="B6B84D78FFF12E2675C6FACFFD0AFA33" ID-CoL="3D3KK" authority="Tanaka &amp; Fujita, 2002" authorityName="Tanaka &amp; Fujita" authorityYear="2002" box="[151,707,1398,1424]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nipponica">
<emphasis id="43CCEAE9FFF12E2675C6FACFFD0AFA33" bold="true" box="[151,707,1398,1424]" pageId="23" pageNumber="24">
<emphasis id="43CCEAE9FFF12E2675C6FACFFE68FA33" bold="true" box="[151,417,1398,1424]" italics="true" pageId="23" pageNumber="24">Eurycercus nipponica</emphasis>
<bibRefCitation id="15294B0AFFF12E2674F9FACFFD0AFA33" author="Tanaka, M. &amp; Fujita, S." box="[424,707,1398,1424]" pageId="23" pageNumber="24" pagination="1 - 25" refId="ref28361" refString="Tanaka, M. &amp; Fujita, S. (2002) Study on the plankton from Lake Utonai-numa, Hokkaido. Yokkaichi University Journal of Environmental and Information Sciences, 6, 1 - 25 (in Japanese)." type="journal article" year="2002">Tanaka &amp; Fujita, 2002</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF12E2675C6FA20FF31FA12" box="[151,248,1433,1457]" pageId="23" pageNumber="24" type="description">
<paragraph id="710736FBFFF12E2675C6FA20FF31FA12" blockId="23.[151,707,1398,1457]" box="[151,248,1433,1457]" pageId="23" pageNumber="24">
<figureCitation id="E9832A7EFFF12E2675C6FA20FF31FA12" box="[151,248,1433,1457]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="25.[151,250,1901,1924]" captionStartId-1="26.[151,250,1863,1886]" captionTargetBox-0="[155,1424,527,1873]" captionTargetBox-1="[158,1425,250,1794]" captionTargetId-0="figure-249@25.[151,1436,517,1889]" captionTargetId-1="figure-101@26.[151,1436,241,1803]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." captionText-1="FIGURE 9. Eurycercus nipponica, limbs of parthenogenetic female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, limb I. B, inner distal lobe of limb I. C, limb II. D, its distal armature. E, limb III. F, distal armature of its gnathobase. G, limb IV. H, distal armature of its gnathobase. I, limb V. J, limb VI. Scale bar denotes 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5248229" figureDoi-1="http://doi.org/10.5281/zenodo.5248231" httpUri-0="https://zenodo.org/record/5248229/files/figure.png" httpUri-1="https://zenodo.org/record/5248231/files/figure.png" pageId="23" pageNumber="24">Figs 89</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF12E2675C6FA5BFA4EFA54" box="[151,1415,1505,1527]" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="710736FBFFF12E2675C6FA5BFA4EFA54" blockId="23.[151,1415,1505,1527]" box="[151,1415,1505,1527]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="51A811D5FFF12E2675C6FA5BFA4EFA54" box="[151,1415,1505,1527]" pageId="23" pageNumber="24">
<treatmentCitation id="F01910EAFFF12E2675C6FA5BFD69FA54" author="Tanaka, M. &amp; Fujita, S." box="[151,672,1505,1527]" page="13" pageId="23" pageNumber="24" year="2002">
<taxonomicName id="B6B84D78FFF12E2675C6FA5BFD69FA54" ID-CoL="3D3KK" authority="Tanaka &amp; Fujita, 2002, p. 13" authorityName="Tanaka &amp; Fujita" authorityPageNumber="13" authorityYear="2002" box="[151,672,1505,1527]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nipponica">
<emphasis id="43CCEAE9FFF12E2675C6FA5BFEBDFA54" box="[151,372,1506,1527]" italics="true" pageId="23" pageNumber="24">Eurycercus nipponica</emphasis>
<bibRefCitation id="15294B0AFFF12E26742AFA58FDA8FA54" author="Tanaka, M. &amp; Fujita, S." box="[379,609,1505,1527]" pageId="23" pageNumber="24" pagination="1 - 25" refId="ref28361" refString="Tanaka, M. &amp; Fujita, S. (2002) Study on the plankton from Lake Utonai-numa, Hokkaido. Yokkaichi University Journal of Environmental and Information Sciences, 6, 1 - 25 (in Japanese)." type="journal article" year="2002">Tanaka &amp; Fujita, 2002</bibRefCitation>
, p. 13
</taxonomicName>
</treatmentCitation>
, Pl. 5: figs 12, Pl. 6: figs 12, Pl. 7: figs 16;
<treatmentCitation id="F01910EAFFF12E26712DFA58FA4DFA54" author="Tanaka, S. &amp; Ohtaka, A. &amp; Nishino, M." box="[1148,1412,1505,1527]" page="172" pageId="23" pageNumber="24" year="2004">
<bibRefCitation id="15294B0AFFF12E26712DFA58FAFEFA54" author="Tanaka, S. &amp; Ohtaka, A. &amp; Nishino, M." box="[1148,1335,1505,1527]" pageId="23" pageNumber="24" pagination="167 - 179" refId="ref28402" refString="Tanaka, S., Ohtaka, A. &amp; Nishino, M. (2004) Cladoceran fauna in littoral zones and Naikos (attached lakes or lagoons) of Lake Biwa, central Japan. Japanese Journal of Limnology, 65, 167 - 179." type="journal article" year="2004">
Tanaka
<emphasis id="43CCEAE9FFF12E26719BFA5BFB34FA54" box="[1226,1277,1505,1527]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2004
</bibRefCitation>
, p. 172
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF12E2975C6F99DFEB5FE9C" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="710736FBFFF12E2675C6F99DFDEAF9FC" blockId="23.[151,1436,1571,2028]" pageId="23" pageNumber="24">
<emphasis id="43CCEAE9FFF12E2675C6F99DFEFEF99F" bold="true" box="[151,311,1571,1596]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">Type locality.</emphasis>
&quot;Lake Utonai-numa&quot; (
<bibRefCitation id="15294B0AFFF12E267762F99DFCE3F99F" author="Tanaka, M. &amp; Fujita, S." box="[563,810,1572,1596]" pageId="23" pageNumber="24" pagination="1 - 25" refId="ref28361" refString="Tanaka, M. &amp; Fujita, S. (2002) Study on the plankton from Lake Utonai-numa, Hokkaido. Yokkaichi University Journal of Environmental and Information Sciences, 6, 1 - 25 (in Japanese)." type="journal article" year="2002">Tanaka &amp; Fujita 2002</bibRefCitation>
) =Unonaito-numa,
<collectingRegion id="B37CF819FFF12E267157F99DFBBEF99F" box="[1030,1143,1572,1596]" country="Japan" name="Hokkaido" pageId="23" pageNumber="24">Hokkaido</collectingRegion>
,
<collectingCountry id="09AF766BFFF12E2671D3F99DFB0CF998" box="[1154,1221,1572,1595]" name="Japan" pageId="23" pageNumber="24">Japan</collectingCountry>
. Approximate geographic coordinates: 42.70, 141.71.
</paragraph>
<paragraph id="710736FBFFF12E267598F9D5FDE0F904" blockId="23.[151,1436,1571,2028]" pageId="23" pageNumber="24">
<emphasis id="43CCEAE9FFF12E267598F9D5FEF4F927" bold="true" box="[201,317,1644,1668]" inLineHeading="true" pageId="23" pageNumber="24" reason="1">
<typeStatus id="AE038859FFF12E267598F9D5FEF1F927" box="[201,312,1644,1668]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
.
</emphasis>
A female with number Y7860825, but the place if its deposition unclear, and absent in the first description (
<bibRefCitation id="15294B0AFFF12E267473F936FDD4F904" author="Tanaka, M. &amp; Fujita, S." box="[290,541,1679,1703]" pageId="23" pageNumber="24" pagination="1 - 25" refId="ref28361" refString="Tanaka, M. &amp; Fujita, S. (2002) Study on the plankton from Lake Utonai-numa, Hokkaido. Yokkaichi University Journal of Environmental and Information Sciences, 6, 1 - 25 (in Japanese)." type="journal article" year="2002">Tanaka &amp; Fujita 2002</bibRefCitation>
).
</paragraph>
<paragraph id="710736FBFFF12E267598F90AFB39F94C" blockId="23.[151,1436,1571,2028]" pageId="23" pageNumber="24">
<emphasis id="43CCEAE9FFF12E267598F90AFE39F96F" bold="true" box="[201,496,1715,1740]" pageId="23" pageNumber="24">Material examined here:</emphasis>
<materialsCitation id="C1D03CA6FFF12E2674A9F90CFEB6F94C" collectingDate="2006-06-12" collectorName="Aoki Ko &amp; S. Tanaka" country="Japan" latitude="36.6089" location="Nagano" longLatPrecision="6" longitude="137.8542" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Nagano">
<emphasis id="43CCEAE9FFF12E2674A9F90CFD8DF96F" box="[504,580,1717,1740]" italics="true" pageId="23" pageNumber="24">
<collectingCountry id="09AF766BFFF12E2674A9F90CFD89F96F" box="[504,576,1717,1740]" name="Japan" pageId="23" pageNumber="24">Japan</collectingCountry>
.
</emphasis>
<collectingRegion id="B37CF819FFF12E26771AF90DFD6FF96F" box="[587,678,1716,1740]" country="Japan" name="Nagano" pageId="23" pageNumber="24">Nagano</collectingRegion>
.
<collectorName id="DC4D532DFFF12E2677E1F90DFCDCF968" box="[688,789,1716,1740]" pageId="23" pageNumber="24">Aoki Ko</collectorName>
, coll. in
<date id="0506103BFFF12E267627F90DFC3AF96F" box="[886,1011,1716,1740]" pageId="23" pageNumber="24" value="2006-06-12">
<collectingDate id="1542E9D3FFF12E267627F90DFC3AF96F" box="[886,1011,1716,1740]" pageId="23" pageNumber="24" value="2006-06-12">06.12.2006</collectingDate>
</date>
by
<collectorName id="DC4D532DFFF12E26714EF90DFB59F96F" box="[1055,1168,1716,1740]" pageId="23" pageNumber="24">S. Tanaka</collectorName>
, DJT 4-235, DJT 4-236 (
<geoCoordinate id="148C503CFFF12E2675CFF96EFF34F94C" box="[158,253,1751,1775]" degrees="36.6089" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="36.6089">36.6089</geoCoordinate>
,
<geoCoordinate id="148C503CFFF12E267458F96EFEBEF94C" box="[265,375,1751,1775]" degrees="137.8542" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="137.8542">137.8542</geoCoordinate>
)
</materialsCitation>
; Kizaki Ko, coll. in
<date id="0506103BFFF12E267730F96EFD17F94C" box="[609,734,1751,1775]" pageId="23" pageNumber="24" value="2006-06-12">06.12.2006</date>
by S. Tanaka, DJT 4-237 (
<geoCoordinate id="148C503CFFF12E26715DF96EFBA2F94C" box="[1036,1131,1751,1775]" degrees="36.5557" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="36.5557">36.5557</geoCoordinate>
,
<geoCoordinate id="148C503CFFF12E267126F96EFB2CF94C" box="[1143,1253,1751,1775]" degrees="137.8408" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="137.8408">137.8408</geoCoordinate>
).
</paragraph>
<paragraph id="710736FBFFF12E297598F944FEB5FE9C" blockId="23.[151,1436,1571,2028]" lastBlockId="24.[151,1437,151,2012]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<materialsCitation id="C1D03CA6FFF12E267598F944FE59F894" collectionCode="AAK" collectorName="A. A. Kotov" country="Russia" latitude="51.41754" location="Area." longLatPrecision="3" longitude="157.0459" pageId="23" pageNumber="24" specimenCode="AAK 2009-104" specimenCount="1" stateProvince="Kamchatka">
<emphasis id="43CCEAE9FFF12E267598F944FE3AF8B7" box="[201,499,1788,1812]" italics="true" pageId="23" pageNumber="24">
<collectingCountry id="09AF766BFFF12E267598F944FEDEF8B7" box="[201,279,1789,1812]" name="Russia" pageId="23" pageNumber="24">Russia</collectingCountry>
.
<collectingRegion id="B37CF819FFF12E267479F945FE6EF8B7" box="[296,423,1788,1812]" country="Russia" name="Kamchatka" pageId="23" pageNumber="24">Kamchatka</collectingRegion>
<location id="74676020FFF12E2674E7F945FE3AF8B7" LSID="urn:lsid:plazi:treatment:F91187EDFFF12E2875C6FACFFB90FE77:74676020FFF12E2674E7F945FE3AF8B7" box="[438,499,1788,1812]" country="Russia" latitude="51.41754" longLatPrecision="3" longitude="157.0459" name="Area." pageId="23" pageNumber="24" stateProvince="Kamchatka">Area.</location>
</emphasis>
A bay of Kurilskoe Lake, coll. in 08.2009 by
<collectorName id="DC4D532DFFF12E26716AF945FB1DF8B0" box="[1083,1236,1788,1812]" pageId="23" pageNumber="24">A. A. Kotov</collectorName>
,
<specimenCode id="211E9E80FFF12E2671B7F945FA55F8B7" box="[1254,1436,1788,1812]" collectionCode="AAK" pageId="23" pageNumber="24">AAK 2009-104</specimenCode>
(
<geoCoordinate id="148C503CFFF12E2675CFF8A6FEC2F894" box="[158,267,1823,1847]" degrees="51.41754" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="51.41754">51.41754</geoCoordinate>
,
<geoCoordinate id="148C503CFFF12E26744BF8A6FE41F894" box="[282,392,1823,1847]" degrees="157.0459" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="157.0459">157.0459</geoCoordinate>
)
</materialsCitation>
;
<materialsCitation id="C1D03CA6FFF12E2674CFF8A6FDF1F8FF" collectionCode="AAK" collectorName="Kurilskoe &amp; A. A. Kotov" country="Russia" latitude="51.4172" location="Travianoy Cape" longLatPrecision="6" longitude="157.0454" pageId="23" pageNumber="24" specimenCode="AAK 2009-103" specimenCount="1" stateProvince="Kamchatka">
A swampy area near
<location id="74676020FFF12E2677C6F8A6FC86F894" LSID="urn:lsid:plazi:treatment:F91187EDFFF12E2875C6FACFFB90FE77:74676020FFF12E2677C6F8A6FC86F894" box="[663,847,1823,1847]" country="Russia" latitude="51.4172" longLatPrecision="6" longitude="157.0454" name="Travianoy Cape" pageId="23" pageNumber="24" stateProvince="Kamchatka">Travianoy Cape</location>
,
<collectorName id="DC4D532DFFF12E26760FF8A6FC05F894" box="[862,972,1823,1847]" pageId="23" pageNumber="24">Kurilskoe</collectorName>
Lake, coll. in 08.2009 by
<collectorName id="DC4D532DFFF12E267057F8A6FA51F894" box="[1286,1432,1823,1847]" pageId="23" pageNumber="24">A. A. Kotov</collectorName>
,
<specimenCode id="211E9E80FFF12E2675C6F8FDFE8FF8FF" box="[151,326,1860,1884]" collectionCode="AAK" pageId="23" pageNumber="24">AAK 2009-103</specimenCode>
(
<geoCoordinate id="148C503CFFF12E267406F8FDFE7FF8FF" box="[343,438,1860,1884]" degrees="51.4172" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="51.4172">51.4172</geoCoordinate>
,
<geoCoordinate id="148C503CFFF12E267493F8FDFDF9F8FF" box="[450,560,1860,1884]" degrees="157.0454" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="157.0454">157.0454</geoCoordinate>
)
</materialsCitation>
;
<materialsCitation id="C1D03CA6FFF12E267715F8FDFB77F8DC" collectingDate="2009-08-16" collectorName="Kurilskoe &amp; A. A. Kotov" country="Russia" latitude="51.4165" location="Travianoy Cape" longLatPrecision="6" longitude="157.045" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Kamchatka">
A swampy area (with a fresh bear trail in time of sampling) near
<location id="74676020FFF12E26707BF8FDFF1AF8DC" LSID="urn:lsid:plazi:treatment:F91187EDFFF12E2875C6FACFFB90FE77:74676020FFF12E26707BF8FDFF1AF8DC" country="Russia" latitude="51.4165" longLatPrecision="6" longitude="157.045" name="Travianoy Cape" pageId="23" pageNumber="24" stateProvince="Kamchatka">Travianoy Cape</location>
,
<collectorName id="DC4D532DFFF12E26758EF8DEFE84F8DC" box="[223,333,1895,1919]" pageId="23" pageNumber="24">Kurilskoe</collectorName>
Lake, coll. in
<date id="0506103BFFF12E2674A1F8DEFDA7F8DC" box="[496,622,1895,1919]" pageId="23" pageNumber="24" value="2009-08-16">
<collectingDate id="1542E9D3FFF12E2674A1F8DEFDA7F8DC" box="[496,622,1895,1919]" pageId="23" pageNumber="24" value="2009-08-16">16.08.2009</collectingDate>
</date>
by
<collectorName id="DC4D532DFFF12E2677C9F8DEFCEDF8DC" box="[664,804,1895,1919]" pageId="23" pageNumber="24">A. A. Kotov</collectorName>
, AAK M-1352 (
<geoCoordinate id="148C503CFFF12E26768CF8DEFBF5F8DC" box="[989,1084,1895,1919]" degrees="51.4165" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="51.4165">51.4165</geoCoordinate>
,
<geoCoordinate id="148C503CFFF12E267119F8DEFB7FF8DC" box="[1096,1206,1895,1919]" degrees="157.0450" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="157.045">157.0450</geoCoordinate>
)
</materialsCitation>
;
<materialsCitation id="C1D03CA6FFF12E267198F8DEFBBAF807" collectingDate="2009-08-16" collectorName="A. A. Kotov" country="Russia" latitude="51.4101" location="Kurilskoe Lake" longLatPrecision="6" longitude="157.049" municipality="Tundra lake" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Kamchatka">
<collectingMunicipality id="9163AC81FFF12E267198F8DEFA86F8DC" box="[1225,1359,1895,1919]" pageId="23" pageNumber="24">Tundra lake</collectingMunicipality>
1 near
<location id="74676020FFF12E2675C6F835FE8EF807" LSID="urn:lsid:plazi:treatment:F91187EDFFF12E2875C6FACFFB90FE77:74676020FFF12E2675C6F835FE8EF807" box="[151,327,1932,1956]" country="Russia" latitude="51.4101" longLatPrecision="6" longitude="157.049" municipality="Tundra lake" name="Kurilskoe Lake" pageId="23" pageNumber="24" stateProvince="Kamchatka">Kurilskoe Lake</location>
, coll. in
<date id="0506103BFFF12E2674F6F835FDECF807" box="[423,549,1932,1956]" pageId="23" pageNumber="24" value="2009-08-16">
<collectingDate id="1542E9D3FFF12E2674F6F835FDECF807" box="[423,549,1932,1956]" pageId="23" pageNumber="24" value="2009-08-16">16.08.2009</collectingDate>
</date>
by
<collectorName id="DC4D532DFFF12E26771EF835FD12F800" box="[591,731,1932,1956]" pageId="23" pageNumber="24">A. A. Kotov</collectorName>
, AAK M-1353 (
<geoCoordinate id="148C503CFFF12E2676C2F835FC3BF807" box="[915,1010,1932,1956]" degrees="51.4101" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="51.4101">51.4101</geoCoordinate>
,
<geoCoordinate id="148C503CFFF12E2676ACF835FBA2F807" box="[1021,1131,1932,1956]" degrees="157.0490" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="157.049">157.0490</geoCoordinate>
)
</materialsCitation>
;
<materialsCitation id="C1D03CA6FFF12E26712FF835FBFAF86B" collectingDate="2009-08-16" collectorName="A. A. Kotov" country="Russia" latitude="51.409" location="Kurilskoe Lake" longLatPrecision="6" longitude="157.0517" municipality="Tundra lake" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Kamchatka">
<collectingMunicipality id="9163AC81FFF12E26712FF835FACDF807" box="[1150,1284,1932,1956]" pageId="23" pageNumber="24">Tundra lake</collectingMunicipality>
4 near
<location id="74676020FFF12E267006F835FEC5F86B" LSID="urn:lsid:plazi:treatment:F91187EDFFF12E2875C6FACFFB90FE77:74676020FFF12E267006F835FEC5F86B" country="Russia" latitude="51.409" longLatPrecision="6" longitude="157.0517" municipality="Tundra lake" name="Kurilskoe Lake" pageId="23" pageNumber="24" stateProvince="Kamchatka">Kurilskoe Lake</location>
, coll. in
<date id="0506103BFFF12E26743BF809FE21F86B" box="[362,488,1968,1992]" pageId="23" pageNumber="24" value="2009-08-16">
<collectingDate id="1542E9D3FFF12E26743BF809FE21F86B" box="[362,488,1968,1992]" pageId="23" pageNumber="24" value="2009-08-16">16.08.2009</collectingDate>
</date>
by
<collectorName id="DC4D532DFFF12E267740F809FD55F86B" box="[529,668,1968,1992]" pageId="23" pageNumber="24">A. A. Kotov</collectorName>
, AAK M-1357 (
<geoCoordinate id="148C503CFFF12E267602F809FC7BF86B" box="[851,946,1968,1992]" degrees="51.4090" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="51.409">51.4090</geoCoordinate>
,
<geoCoordinate id="148C503CFFF12E2676ECF809FBE2F86B" box="[957,1067,1968,1992]" degrees="157.0517" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="157.0517">157.0517</geoCoordinate>
)
</materialsCitation>
; A forest puddle 1 near the Plotnikova River, coll. in
<date id="0506103BFFF12E2674DEF86DFDC4F84F" box="[399,525,2004,2028]" pageId="23" pageNumber="24" value="2009-08-13">13.08.2009</date>
by A. A. Kotov, AAK M-1344 (
<geoCoordinate id="148C503CFFF12E2676D5F86DFC2AF84F" box="[900,995,2004,2028]" degrees="52.9208" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="52.9208">52.9208</geoCoordinate>
,
<geoCoordinate id="148C503CFFF12E2676BEF86DFB94F84F" box="[1007,1117,2004,2028]" degrees="157.1527" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="157.1527">157.1527</geoCoordinate>
);
<materialsCitation id="C1D03CA6FFF12E297123F86DFB97FF0C" collectionCode="AAK" collectorName="Bystraya &amp; Kotov" country="Russia" lastPageId="24" lastPageNumber="25" latitude="52.9264" longLatPrecision="6" longitude="156.6012" pageId="23" pageNumber="24" specimenCode="AAK 2009-093" specimenCount="1" stateProvince="Kamchatka">
A forest puddle 2 near the
<collectorName id="DC4D532DFFFE2E2975C6FF21FF33FF13" box="[151,250,152,176]" pageId="24" pageNumber="25">Bystraya</collectorName>
River, coll. in 08.2009 by A. A.
<collectorName id="DC4D532DFFFE2E29773CFF21FD7DFF0C" box="[621,692,152,175]" pageId="24" pageNumber="25">Kotov</collectorName>
,
<specimenCode id="211E9E80FFFE2E2977EEFF2EFCA7FF0C" box="[703,878,151,175]" collectionCode="AAK" pageId="24" pageNumber="25">AAK 2009-093</specimenCode>
(
<geoCoordinate id="148C503CFFFE2E29762CFF2EFC14FF0C" box="[893,989,151,175]" degrees="52.9264" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="52.9264">52.9264</geoCoordinate>
,
<geoCoordinate id="148C503CFFFE2E2976B9FF2EFB9FFF0C" box="[1000,1110,151,175]" degrees="156.6012" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="156.6012">156.6012</geoCoordinate>
)
</materialsCitation>
;
<materialsCitation id="C1D03CA6FFFE2E297138FF2EFB16FF77" collectionCode="AAK" collectorName="Bystraya &amp; A. A. Kotov" country="Russia" latitude="52.9304" location="An" longLatPrecision="6" longitude="156.6031" pageId="24" pageNumber="25" specimenCode="AAK 2009-094" specimenCount="1" stateProvince="Kamchatka">
<location id="74676020FFFE2E297138FF2EFB42FF0C" LSID="urn:lsid:plazi:treatment:F91187EDFFF12E2875C6FACFFB90FE77:74676020FFFE2E297138FF2EFB42FF0C" box="[1129,1163,151,175]" country="Russia" latitude="52.9304" longLatPrecision="6" longitude="156.6031" name="An" pageId="24" pageNumber="25" stateProvince="Kamchatka">An</location>
affluent of the
<collectorName id="DC4D532DFFFE2E297069FF21FA52FF13" box="[1336,1435,152,176]" pageId="24" pageNumber="25">Bystraya</collectorName>
River, coll. in 08.2009 by
<collectorName id="DC4D532DFFFE2E297494FF05FD9DFF70" box="[453,596,188,212]" pageId="24" pageNumber="25">A. A. Kotov</collectorName>
,
<specimenCode id="211E9E80FFFE2E297733FF05FCDAFF77" box="[610,787,188,212]" collectionCode="AAK" pageId="24" pageNumber="25">AAK 2009-094</specimenCode>
and AAK M-1376 (
<geoCoordinate id="148C503CFFFE2E2976ACFF05FB95FF77" box="[1021,1116,188,212]" degrees="52.9304" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="52.9304">52.9304</geoCoordinate>
,
<geoCoordinate id="148C503CFFFE2E29713BFF05FB1EFF77" box="[1130,1239,188,212]" degrees="156.6031" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="156.6031">156.6031</geoCoordinate>
)
</materialsCitation>
;
<materialsCitation id="C1D03CA6FFFE2E2971BDFF05FA87FF54" collectingDate="2009-08-13" collectionCode="AAK" collectorName="T. N. Travina" country="Russia" latitude="52.7612" location="Bolshaya River" longLatPrecision="6" longitude="156.2647" pageId="24" pageNumber="25" specimenCode="AAK 2009-079" specimenCount="1" stateProvince="Kamchatka">
<location id="74676020FFFE2E2971BDFF05FA55FF77" LSID="urn:lsid:plazi:treatment:F91187EDFFF12E2875C6FACFFB90FE77:74676020FFFE2E2971BDFF05FA55FF77" box="[1260,1436,188,212]" country="Russia" latitude="52.7612" longLatPrecision="6" longitude="156.2647" name="Bolshaya River" pageId="24" pageNumber="25" stateProvince="Kamchatka">Bolshaya River</location>
near the KamchatNIRO station, coll. in
<date id="0506103BFFFE2E29770DFF66FD13FF54" box="[604,730,223,247]" pageId="24" pageNumber="25" value="2009-08-13">
<collectingDate id="1542E9D3FFFE2E29770DFF66FD13FF54" box="[604,730,223,247]" pageId="24" pageNumber="25" value="2009-08-13">13.08.2009</collectingDate>
</date>
by
<collectorName id="DC4D532DFFFE2E297657FF59FC69FF54" box="[774,928,223,247]" pageId="24" pageNumber="25">T. N. Travina</collectorName>
,
<specimenCode id="211E9E80FFFE2E2976FCFF66FB94FF54" box="[941,1117,223,247]" collectionCode="AAK" pageId="24" pageNumber="25">AAK 2009-079</specimenCode>
(
<geoCoordinate id="148C503CFFFE2E29713CFF66FB04FF54" box="[1133,1229,223,247]" degrees="52.7612" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="52.7612">52.7612</geoCoordinate>
,
<geoCoordinate id="148C503CFFFE2E297188FF66FA8EFF54" box="[1241,1351,223,247]" degrees="156.2647" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="156.2647">156.2647</geoCoordinate>
)
</materialsCitation>
;
<materialsCitation id="C1D03CA6FFFE2E29700AFF66FEB1FE9C" collectingDate="1985-02-10" collectionCode="NNS" country="Russia" location="Lake Azabachje on Cape Ivashka" municipality="Small lake" pageId="24" pageNumber="25" specimenCode="NNS 1999-008, NNS 1999-017" specimenCount="1" stateProvince="Kamchatka">
<collectingMunicipality id="9163AC81FFFE2E29700AFF66FF0FFEBF" pageId="24" pageNumber="25">Small lake</collectingMunicipality>
near
<location id="74676020FFFE2E297457FEBDFD4CFEBF" LSID="urn:lsid:plazi:treatment:F91187EDFFF12E2875C6FACFFB90FE77:74676020FFFE2E297457FEBDFD4CFEBF" box="[262,645,260,284]" country="Russia" municipality="Small lake" name="Lake Azabachje on Cape Ivashka" pageId="24" pageNumber="25" stateProvince="Kamchatka">Lake Azabachje on Cape Ivashka</location>
, coll. in
<date id="0506103BFFFE2E2977B8FEBDFCA3FEBF" box="[745,874,260,284]" pageId="24" pageNumber="25" value="1985-02-10">
<collectingDate id="1542E9D3FFFE2E2977B8FEBDFCA3FEBF" box="[745,874,260,284]" pageId="24" pageNumber="25" value="1985-02-10">02.10.1985</collectingDate>
</date>
,
<specimenCode id="211E9E80FFFE2E297627FEBDFBEDFEBF" box="[886,1060,260,284]" collectionCode="NNS" pageId="24" pageNumber="25">NNS 1999-008</specimenCode>
,
<specimenCode id="211E9E80FFFE2E297160FEBDFB15FEBF" box="[1073,1244,260,284]" collectionCode="NNS" pageId="24" pageNumber="25">NNS 1999-017</specimenCode>
and AAK 1999- 077 (56.16, 161.85)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFFE2E297598FEF2FB60F88B" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="710736FBFFFE2E297598FEF2FD10FD04" blockId="24.[151,1437,151,2012]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FEF2FDA7FEC7" bold="true" box="[201,622,331,356]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FEF2FE8AFEC7" box="[201,323,331,356]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">Diagnosis.</emphasis>
Parthenogenetic female.
</emphasis>
Dorsal head pores on a bubble-like projection located immediately on head shield. In anterior view, body wide, not compressed laterally, median dorsal keel absent. Rostrum relatively long. Ocellus small. Lateral head pore minute, circular. Labrum with a large median keel, terminating in an angled apex, reaching distal end of antenna I. Postabdomen with sub-parallel dorsal and ventral margins, preanal teeth pointed. Spines at base of pre-claw portion predominantly single. Antenna I with antennular sensory seta arising somewhat basally to middle. Denticles in rows encircling antennular surface relatively large. On antenna II, spine situated on proximal segment of exopod equal to or somewhat shorter than second segment. Limb I IDL with a strong hook-like seta, the smallest IDL seta especially short. IDL with about 11 distal spinules, about 8 proximal spinules, about 6 marginal spinules and about 10 basal spinules. Eight setae in filter plate II, 8 setae in filter plate III; 8 setae in filter plate IV, 8 setae in filter plate V.
</paragraph>
<paragraph id="710736FBFFFE2E297598FD0AFD57FCFF" blockId="24.[151,1437,151,2012]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FD0AFD2EFD6F" bold="true" box="[201,743,691,716]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FD0AFE72FD6F" box="[201,443,691,716]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">Short redescription.</emphasis>
Parthenogenetic female.
</emphasis>
In lateral view body sub-ovoid in larger females (
<figureCitation id="E9832A7EFFFE2E297063FD0DFA46FD6F" box="[1330,1423,692,716]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8A</figureCitation>
), maximum height of the body in its middle portion (BH/BL= 0.500.66). Dorsal margin interrupted only by a bubble-like head pore. In anterior or ventral view, body wide, not compressed laterally (
<figureCitation id="E9832A7EFFFE2E297103FD45FB65FCB7" box="[1106,1196,764,788]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8D</figureCitation>
), maximum width of body at level of mandibular articulation, median dorsal keel absent. Intestine has a single loop, posterior intestinal caecum present. Few eggs in the brood pouch.
</paragraph>
<paragraph id="710736FBFFFE2E297598FCD1FA5CFC07" blockId="24.[151,1437,151,2012]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FCD1FECFFC23" box="[201,262,872,896]" italics="true" pageId="24" pageNumber="25">Head</emphasis>
with short rostrum (
<figureCitation id="E9832A7EFFFE2E2974A2FCDEFD85FCDC" box="[499,588,871,895]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8E</figureCitation>
). A single major “head pore” as a ringed, sub-oval field of special cuticle located on a dorsal bubble. A minute, circular lateral pore located at either side of major pore, closer to it (
<figureCitation id="E9832A7EFFFE2E297065FC35FA41FC07" box="[1332,1416,908,932]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8F</figureCitation>
).
</paragraph>
<paragraph id="710736FBFFFE2E297598FC09FEF6FC4F" blockId="24.[151,1437,151,2012]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FC09FEEBFC6B" box="[201,290,944,968]" italics="true" pageId="24" pageNumber="25">Labrum</emphasis>
fleshy body with paired lateral horn-like projections, a large median keel terminating in an angled apex (
<figureCitation id="E9832A7EFFFE2E29758AFC6DFEFBFC4F" box="[219,306,980,1004]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8E</figureCitation>
).
</paragraph>
<paragraph id="710736FBFFFE2E297598FC41FC09FBAC" blockId="24.[151,1437,151,2012]" box="[201,960,1015,1040]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FC41FED8FBB3" box="[201,273,1016,1040]" italics="true" pageId="24" pageNumber="25">Valves</emphasis>
generally ovoid (VL/BL= 0.840.96), as in previous species.
</paragraph>
<paragraph id="710736FBFFFE2E297598FBA5FB8EFAAF" blockId="24.[151,1437,151,2012]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FBA5FEA8FB97" box="[201,353,1052,1076]" italics="true" pageId="24" pageNumber="25">Postabdomen</emphasis>
large (PL/BL= 0.270.43), relatively broad (
<collectingCountry id="09AF766BFFFE2E29763BFBA5FC46FB97" box="[874,911,1052,1076]" name="Philippines" pageId="24" pageNumber="25">PH</collectingCountry>
/PL= 0.450.46), with sub-parallel dorsal and ventral margins (
<figureCitation id="E9832A7EFFFE2E297405FBF9FE67FBFB" box="[340,430,1088,1112]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8G</figureCitation>
). Distal anal embayment very shallow. Armature of the preanal margin as a series of preanal teeth (NT=101); small gap lacking any teeth at base of postabdominal setae (
<figureCitation id="E9832A7EFFFE2E29714BFBDDFBAEFBDF" box="[1050,1127,1124,1148]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8I</figureCitation>
); teeth in middle of preanal margin with sharp tips, distalmost tooth equal to or somewhat larger than others. Distalmost postanal teeth particularly large, predominantly clustered, teeth at base of pre-claw portion (=at distal part of anal margin) predominantly single (
<figureCitation id="E9832A7EFFFE2E2975B8FB69FE88FB4B" box="[233,321,1232,1256]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8H</figureCitation>
, arrows). Postabdominal claw relatively robust (CL/PL= 0.160.24); basal spines, first (distal) long (DS/CL= 0.250.38), second (basal) short (BS/CL= 0.130.25; BS/DS= 0.430.60).
</paragraph>
<paragraph id="710736FBFFFE2E297598FAA1FCAEFA47" blockId="24.[151,1437,151,2012]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FAA1FEF1FA93" box="[201,312,1304,1328]" italics="true" pageId="24" pageNumber="25">Antenna I</emphasis>
relatively short (AL/BL = 0.100.14; AL/DA=3.0
<quantity id="B6409B1EFFFE2E2976DAFAA1FC05FA93" box="[907,972,1304,1328]" metricMagnitude="-2" metricUnit="m" metricValue="9.144" pageId="24" pageNumber="25" unit="in" value="3.6">3.6 in</quantity>
adults), triangular in section; protruding greatly beyond tip of rostrum. Antennular sensory seta relatively short (about third of antenna I length), arising somewhat basally to antenna I middle (
<figureCitation id="E9832A7EFFFE2E29771CFAD9FD54FADB" box="[589,669,1376,1400]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8J</figureCitation>
). Nine bisegmented aesthetascs, with pointed teeth around them. No setules at anterior margin of antenna I. Numerous rows of minute denticles encircling antennular surface. Antenna II relatively short. On antenna II, spine situated on proximal segment of exopod somewhat shorter than length of second segment (
<figureCitation id="E9832A7EFFFE2E297408FA75FE65FA47" box="[345,428,1484,1508]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig.8K</figureCitation>
). Setae 0-0-3/1-1-3; spines 1-0-1/0-0-1.
</paragraph>
<paragraph id="710736FBFFFE2E297598FA49FCE7F918" blockId="24.[151,1437,151,2012]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598FA49FEC4F9AB" box="[201,269,1520,1544]" italics="true" pageId="24" pageNumber="25">Limbs</emphasis>
in general as in previous species (
<figureCitation id="E9832A7EFFFE2E2977DDFA49FCC9F9AB" box="[652,768,1520,1544]" captionStart="FIGURE 9" captionStartId="26.[151,250,1863,1886]" captionTargetBox="[158,1425,250,1794]" captionTargetId="figure-101@26.[151,1436,241,1803]" captionTargetPageId="26" captionText="FIGURE 9. Eurycercus nipponica, limbs of parthenogenetic female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, limb I. B, inner distal lobe of limb I. C, limb II. D, its distal armature. E, limb III. F, distal armature of its gnathobase. G, limb IV. H, distal armature of its gnathobase. I, limb V. J, limb VI. Scale bar denotes 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5248231" httpUri="https://zenodo.org/record/5248231/files/figure.png" pageId="24" pageNumber="25">Fig. 9AJ</figureCitation>
). IDL of limb I with a remarkable strong hook-like seta, but not so large as in
<taxonomicName id="B6B84D78FFFE2E29743AF9ACFDE2F98F" authorityName="Frey" authorityYear="1973" box="[363,555,1556,1580]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="macracanthus">
<emphasis id="43CCEAE9FFFE2E29743AF9ACFDE2F98F" box="[363,555,1556,1580]" italics="true" pageId="24" pageNumber="25">E. macracanthus</emphasis>
</taxonomicName>
, smallest seta very small, as in the latter (
<figureCitation id="E9832A7EFFFE2E29714AF9ADFB53F98F" box="[1051,1178,1556,1580]" captionStart="FIGURE 9" captionStartId="26.[151,250,1863,1886]" captionTargetBox="[158,1425,250,1794]" captionTargetId="figure-101@26.[151,1436,241,1803]" captionTargetPageId="26" captionText="FIGURE 9. Eurycercus nipponica, limbs of parthenogenetic female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, limb I. B, inner distal lobe of limb I. C, limb II. D, its distal armature. E, limb III. F, distal armature of its gnathobase. G, limb IV. H, distal armature of its gnathobase. I, limb V. J, limb VI. Scale bar denotes 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5248231" httpUri="https://zenodo.org/record/5248231/files/figure.png" pageId="24" pageNumber="25">Fig. 9AB</figureCitation>
). IDL with long distal spinules (about
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largest adults), long proximal spinules (
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largest adults), short marginal spinules (
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largest adults) and short basal spinules (
<quantity id="B6409B1EFFFE2E297778F9E5FDAAF9D7" box="[553,611,1628,1652]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="24" pageNumber="25" unit="in" value="10.0">10 in</quantity>
largest adults). On exopodite III seta 7 somewhat longer than seta 6 (
<figureCitation id="E9832A7EFFFE2E29703FF9E5FF70F93B" captionStart="FIGURE 9" captionStartId="26.[151,250,1863,1886]" captionTargetBox="[158,1425,250,1794]" captionTargetId="figure-101@26.[151,1436,241,1803]" captionTargetPageId="26" captionText="FIGURE 9. Eurycercus nipponica, limbs of parthenogenetic female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, limb I. B, inner distal lobe of limb I. C, limb II. D, its distal armature. E, limb III. F, distal armature of its gnathobase. G, limb IV. H, distal armature of its gnathobase. I, limb V. J, limb VI. Scale bar denotes 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5248231" httpUri="https://zenodo.org/record/5248231/files/figure.png" pageId="24" pageNumber="25">Fig. 9E</figureCitation>
). On exopodite IV both setae 1 and 2 short (
<figureCitation id="E9832A7EFFFE2E297799F939FCECF93B" box="[712,805,1664,1688]" captionStart="FIGURE 9" captionStartId="26.[151,250,1863,1886]" captionTargetBox="[158,1425,250,1794]" captionTargetId="figure-101@26.[151,1436,241,1803]" captionTargetPageId="26" captionText="FIGURE 9. Eurycercus nipponica, limbs of parthenogenetic female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, limb I. B, inner distal lobe of limb I. C, limb II. D, its distal armature. E, limb III. F, distal armature of its gnathobase. G, limb IV. H, distal armature of its gnathobase. I, limb V. J, limb VI. Scale bar denotes 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5248231" httpUri="https://zenodo.org/record/5248231/files/figure.png" pageId="24" pageNumber="25">Fig. 9G</figureCitation>
). On exopodite V setae 56 slightly increasing in size basally. Filter plate of gnathobase IIIV limbs with 8 setae.
</paragraph>
<paragraph id="710736FBFFFE2E297598F97EFB71F943" blockId="24.[151,1437,151,2012]" box="[201,1208,1735,1760]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598F97EFE52F97C" box="[201,411,1735,1760]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">
<emphasis id="43CCEAE9FFFE2E297598F97EFE5DF943" bold="true" box="[201,404,1735,1760]" pageId="24" pageNumber="25">Ephippial female</emphasis>
.
</emphasis>
Body more compressed laterally, with median dorsal keel (
<figureCitation id="E9832A7EFFFE2E297163F971FB62F943" box="[1074,1195,1736,1760]" captionStart="FIGURE 8" captionStartId="25.[151,250,1901,1924]" captionTargetBox="[155,1424,527,1873]" captionTargetId="figure-249@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 8. Eurycercus nipponica, female from a swampy area (with a fresh bear trail in time of sampling) near Travianoy Cape, Kurilskoe Lake, Kamchatka Area, Russia A, parthenogenetic female, lateral view. BC, ephippial female, lateral and anterior view. D, parthenogenetic female, ventral view. E, head. F, dorsal head pores. G, postabdomen. H, its distal portion. I, its basal portion. J, antenna I. K, antenna II. Scale bar denotes 1 mm (AD); 0.1 mm (EK)." figureDoi="http://doi.org/10.5281/zenodo.5248229" httpUri="https://zenodo.org/record/5248229/files/figure.png" pageId="24" pageNumber="25">Fig. 8BC</figureCitation>
).
</paragraph>
<paragraph id="710736FBFFFE2E297598F955FE4BF8A7" blockId="24.[151,1437,151,2012]" box="[201,386,1772,1796]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598F955FEC5F8A7" bold="true" box="[201,268,1772,1796]" pageId="24" pageNumber="25">Male.</emphasis>
Unknown
</paragraph>
<paragraph id="710736FBFFFE2E297598F8A9FB60F88B" blockId="24.[151,1437,151,2012]" box="[201,1193,1808,1832]" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598F8A9FEEDF88B" bold="true" box="[201,292,1808,1832]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">Length.</emphasis>
0.692.00 mm in our material, up to
<quantity id="B6409B1EFFFE2E297797F8A9FCCFF884" box="[710,774,1808,1832]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="25" unit="mm" value="3.0">3 mm</quantity>
according to
<bibRefCitation id="15294B0AFFFE2E2976CEF8A9FB6CF88B" author="Tanaka, M. &amp; Fujita, S." box="[927,1189,1808,1832]" pageId="24" pageNumber="25" pagination="1 - 25" refId="ref28361" refString="Tanaka, M. &amp; Fujita, S. (2002) Study on the plankton from Lake Utonai-numa, Hokkaido. Yokkaichi University Journal of Environmental and Information Sciences, 6, 1 - 25 (in Japanese)." type="journal article" year="2002">Tanaka &amp; Fujita (2002)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFFE2E287598F88AFAE3FF77" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
<paragraph id="710736FBFFFE2E287598F88AFAE3FF77" blockId="24.[151,1437,151,2012]" lastBlockId="25.[151,1436,151,468]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="43CCEAE9FFFE2E297598F88AFE98F8EF" bold="true" box="[201,337,1843,1868]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">Comments.</emphasis>
<bibRefCitation id="15294B0AFFFE2E297408F88DFDCAF8EF" author="Smirnov, N. N." box="[345,515,1844,1868]" pageId="24" pageNumber="25" pagination="63 - 83" refId="ref28130" refString="Smirnov, N. N. (1998) A revision of the genus Camptocercus (Anomopoda, Chydoridae, Aloninae). Hydrobiologia, 386, 63 - 83." type="journal article" year="1998">Smirnov (1998</bibRefCitation>
: p. 81) said that “Easternmost Siberia is also distinguished by the presence of ... an undescribed species of
<taxonomicName id="B6B84D78FFFE2E2974C9F8E0FDDDF8D3" authorityName="Baird" authorityYear="1843" box="[408,532,1881,1904]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="43CCEAE9FFFE2E2974C9F8E0FDDDF8D3" box="[408,532,1881,1904]" italics="true" pageId="24" pageNumber="25">Eurycercus</emphasis>
</taxonomicName>
”. As we found, this taxon has been formally described as
<taxonomicName id="B6B84D78FFFE2E2971CAF8E0FED4F837" authority="Tanaka &amp; Fujita, 2002" authorityName="Tanaka &amp; Fujita" authorityYear="2002" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nipponica">
<emphasis id="43CCEAE9FFFE2E2971CAF8E0FAE1F8CC" box="[1179,1320,1880,1904]" italics="true" pageId="24" pageNumber="25">E. nipponica</emphasis>
<bibRefCitation id="15294B0AFFFE2E297061F8E1FED4F837" author="Tanaka, M. &amp; Fujita, S." pageId="24" pageNumber="25" pagination="1 - 25" refId="ref28361" refString="Tanaka, M. &amp; Fujita, S. (2002) Study on the plankton from Lake Utonai-numa, Hokkaido. Yokkaichi University Journal of Environmental and Information Sciences, 6, 1 - 25 (in Japanese)." type="journal article" year="2002">Tanaka &amp; Fujita, 2002</bibRefCitation>
</taxonomicName>
based on samples from
<collectingCountry id="09AF766BFFFE2E297761F8C5FDA6F830" box="[560,623,1916,1939]" name="Japan" pageId="24" pageNumber="25">Japan</collectingCountry>
(
<bibRefCitation id="15294B0AFFFE2E29772FF8C5FCB0F837" author="Tanaka, M. &amp; Fujita, S." box="[638,889,1916,1940]" pageId="24" pageNumber="25" pagination="1 - 25" refId="ref28361" refString="Tanaka, M. &amp; Fujita, S. (2002) Study on the plankton from Lake Utonai-numa, Hokkaido. Yokkaichi University Journal of Environmental and Information Sciences, 6, 1 - 25 (in Japanese)." type="journal article" year="2002">Tanaka &amp; Fujita 2002</bibRefCitation>
). The authors diagnosis was lacking any helpful information on the taxon discrimination from other species, but it is obvious from the authors drawings and photos that: (1) the labral keel is very large and has an angled apex; (2) the head pores are located on a bubble-like projection (not represented by the authors in their drawings, but see in their photo in Plate 7 (fragment 1); (3) the preanal teeth on the pre-claw portion are mainly singular. After our redescription, it is clear that this is a valid species.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFFF2E287598FF5BFF28FEE7" pageId="25" pageNumber="26" type="distribution">
<paragraph id="710736FBFFFF2E287598FF5BFF28FEE7" blockId="25.[151,1436,151,468]" pageId="25" pageNumber="26">
<emphasis id="43CCEAE9FFFF2E287598FF5BFEABFF58" bold="true" box="[201,354,226,251]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Distribution.</emphasis>
<taxonomicName id="B6B84D78FFFF2E28743AFF5AFE32FF59" authorityName="Tanaka &amp; Fujita" authorityYear="2002" box="[363,507,227,250]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="nipponica">
<emphasis id="43CCEAE9FFFF2E28743AFF5AFE32FF59" box="[363,507,227,250]" italics="true" pageId="25" pageNumber="26">E. nipponica</emphasis>
</taxonomicName>
was known from
<collectingCountry id="09AF766BFFFF2E287799FF5AFCC1FF59" box="[712,776,227,250]" name="Japan" pageId="25" pageNumber="26">Japan</collectingCountry>
(
<bibRefCitation id="15294B0AFFFF2E287646FF5BFBDCFF59" author="Tanaka, M. &amp; Fujita, S." box="[791,1045,226,250]" pageId="25" pageNumber="26" pagination="1 - 25" refId="ref28361" refString="Tanaka, M. &amp; Fujita, S. (2002) Study on the plankton from Lake Utonai-numa, Hokkaido. Yokkaichi University Journal of Environmental and Information Sciences, 6, 1 - 25 (in Japanese)." type="journal article" year="2002">Tanaka &amp; Fujita 2002</bibRefCitation>
;
<bibRefCitation id="15294B0AFFFF2E287170FF5BFB3FFF59" author="Tanaka, S. &amp; Ohtaka, A. &amp; Nishino, M." box="[1057,1270,226,251]" pageId="25" pageNumber="26" pagination="167 - 179" refId="ref28402" refString="Tanaka, S., Ohtaka, A. &amp; Nishino, M. (2004) Cladoceran fauna in littoral zones and Naikos (attached lakes or lagoons) of Lake Biwa, central Japan. Japanese Journal of Limnology, 65, 167 - 179." type="journal article" year="2004">
Tanaka
<emphasis id="43CCEAE9FFFF2E28712AFF5AFB7DFF58" box="[1147,1204,227,251]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2004
</bibRefCitation>
) we found it also in Kamchatka (see
<figureCitation id="E9832A7EFFFF2E2874F3FEBEFE2BFEBC" box="[418,482,263,287]" captionStart="FIGURE 1" captionStartId="6.[151,250,1495,1518]" captionTargetBox="[188,1405,223,1440]" captionTargetId="figure-267@6.[151,1436,195,1471]" captionTargetPageId="6" captionText="FIGURE 1. Distribution of species from the subgenus Eurycercus (Eurycercus) emend. nov. in the Holarctic according to our original samples—no literature data included. E. lamellatus—grey circles. E. macracanthus—black circles. E. pompholygodes—solid quadrates (red in electronic version). E. microdontus—black stars. E. longirostris—grey quadrates. E. nipponica—solid triangles (green in electronic version). E. beringi sp. nov.—black crosses." figureDoi="http://doi.org/10.5281/zenodo.5248213" httpUri="https://zenodo.org/record/5248213/files/figure.png" pageId="25" pageNumber="26">Fig. 1</figureCitation>
). The species could be present in Sakhalin Island, Kurile Islands and, Chukotka Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFFF2E287598FEE9FB90FE77" pageId="25" pageNumber="26" type="biology_ecology">
<paragraph id="710736FBFFFF2E287598FEE9FB90FE77" blockId="25.[151,1436,151,468]" pageId="25" pageNumber="26">
<emphasis id="43CCEAE9FFFF2E287598FEE9FEE4FECB" bold="true" box="[201,301,336,360]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Ecology.</emphasis>
Mostly, it is present in relatively large lakes and smaller water bodies associated with them (i.e. connected during a spring flooding time). The species is found in very shallow flooded grasslands, but, as in case of
<taxonomicName id="B6B84D78FFFF2E2870D5FECCFEFFFE13" authorityName="Frey" authorityYear="1973" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="macracanthus">
<emphasis id="43CCEAE9FFFF2E2870D5FECCFEFFFE13" italics="true" pageId="25" pageNumber="26">E. macracanthus</emphasis>
</taxonomicName>
, we think that it could be found in temporary water bodies only as the remains of a larger water body formed by spring flooding. It is present in the vegetation patches in rivers themselves.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="4695ACD9BEDA31CD3F1DD4AE3BA0B0FA">A revision of the subgenus Eurycercus (Eurycercus) Baird, 1843 emend. nov. (Cladocera: Eurycercidae) in the Holarctic with the description of a new species from Alaska</mods:title>
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<mods:title id="A50743020563AEBDAE7E41A742E519DA">Zootaxa</mods:title>
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<mods:part id="4E4194B5860AFAAFC539BDF0D484BF4F">
<mods:date id="B0E49CC53B946E2FAA36B4BCCF606D8E">2012</mods:date>
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<mods:number id="CD8761B5C0E26AFFB82A1DD12B9B77BD">2012-02-24</mods:number>
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<mods:number id="446DAD75D574C777BF28A4DFA8E9B3A8">3206</mods:number>
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<mods:url id="B6B4C8C8EE3BF01A7B107D14B1049833">https://biotaxa.org/Zootaxa/article/view/zootaxa.3206.1.1</mods:url>
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<mods:identifier id="CA078DC23C1DDC39837AB4EEA19857C6" type="DOI">10.11646/zootaxa.3206.1.1</mods:identifier>
<mods:identifier id="37A84D4673A3D1B5BDB1404CF668B8AE" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="39A26570FFF42E2375C6FDD5FD81FD26" box="[151,584,619,646]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="710736FBFFF42E2375C6FDD5FD81FD26" blockId="18.[151,584,619,646]" box="[151,584,619,646]" pageId="18" pageNumber="19">
<heading id="2A4F8197FFF42E2375C6FDD5FD81FD26" bold="true" box="[151,584,619,646]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="B6B84D78FFF42E2375C6FDD5FD81FD26" ID-CoL="3D3KB" authority="Frey, 1978" authorityName="Frey" authorityYear="1978" box="[151,584,619,646]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="microdontus">
<emphasis id="43CCEAE9FFF42E2375C6FDD5FD81FD26" bold="true" box="[151,584,619,646]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E2375C6FDD5FE09FD25" bold="true" box="[151,448,620,646]" italics="true" pageId="18" pageNumber="19">Eurycercus microdontus</emphasis>
<bibRefCitation id="15294B0AFFF42E237496FDD2FD81FD26" author="Frey, D. G." box="[455,584,619,645]" pageId="18" pageNumber="19" pagination="1 - 25" refId="ref25888" refString="Frey, D. G. (1978) A new species of Eurycercus (Cladocera, Chydoridae) from the Southern United States. Tulane Studies in Zoology and Botany, 20, 1 - 25." type="journal article" year="1978">Frey, 1978</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="710736FBFFF42E2375C6FD0AFCA1FD6B" blockId="18.[151,872,690,712]" box="[151,872,690,712]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="51A811D5FFF42E2375C6FD0AFCA1FD6B" box="[151,872,690,712]" pageId="18" pageNumber="19">
<treatmentCitation id="F01910EAFFF42E2375C6FD0AFDF9FD6B" author="Frey, D. G." box="[151,560,690,712]" page="8" pageId="18" pageNumber="19" year="1978">
<taxonomicName id="B6B84D78FFF42E2375C6FD0AFDF9FD6B" ID-CoL="3D3KB" authority="Frey, 1978, p. 8" authorityName="Frey" authorityPageNumber="8" authorityYear="1978" box="[151,560,690,712]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="microdontus">
<emphasis id="43CCEAE9FFF42E2375C6FD0AFE44FD6B" box="[151,397,690,712]" italics="true" pageId="18" pageNumber="19">Eurycercus microdontus</emphasis>
<bibRefCitation id="15294B0AFFF42E2374C2FD0AFDC8FD6B" author="Frey, D. G." box="[403,513,690,712]" pageId="18" pageNumber="19" pagination="1 - 25" refId="ref25888" refString="Frey, D. G. (1978) A new species of Eurycercus (Cladocera, Chydoridae) from the Southern United States. Tulane Studies in Zoology and Botany, 20, 1 - 25." type="journal article" year="1978">Frey, 1978</bibRefCitation>
, p. 8
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22,
<figureCitation id="E9832A7EFFF42E237735FD0BFD6AFD6B" box="[612,675,690,712]" captionStart="FIGURE 2" captionStartId="8.[151,250,1721,1744]" captionTargetBox="[162,1395,434,1668]" captionTargetId="figure-263@8.[151,1436,405,1706]" captionTargetPageId="8" captionText="FIGURE 2. Characteristic traits of different species of Eurycercus (Eurycercus) emend. nov. AB, E. longirostris, dorsal view and dorsal head pores in lateral view. CD, E. lamellatus, dorsal view and dorsal head pores in lateral view. EF, E. beringi sp. nov., dorsal view and dorsal head pores in lateral view. Rather schematic, not to scale." figureDoi="http://doi.org/10.5281/zenodo.5248215" httpUri="https://zenodo.org/record/5248215/files/figure.png" pageId="18" pageNumber="19">Figs 2</figureCitation>
17, 1928, 3035.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF42E2375C6FD4CFAB4FBAA" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="710736FBFFF42E2375C6FD4CFEA9FC92" blockId="18.[151,1437,757,1753]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E2375C6FD4CFEFFFCAD" bold="true" box="[151,310,757,782]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Type locality.</emphasis>
&quot;Lake June in Winter, Highlands Co.,
<collectingRegion id="B37CF819FFF42E2377BDFD4CFCF6FCAE" box="[748,831,757,781]" country="United States of America" name="Florida" pageId="18" pageNumber="19">Florida</collectingRegion>
” (
<bibRefCitation id="15294B0AFFF42E237609FD4CFC19FCAE" author="Frey, D. G." box="[856,976,757,781]" pageId="18" pageNumber="19" pagination="1 - 25" refId="ref25888" refString="Frey, D. G. (1978) A new species of Eurycercus (Cladocera, Chydoridae) from the Southern United States. Tulane Studies in Zoology and Botany, 20, 1 - 25." type="journal article" year="1978">Frey 1978</bibRefCitation>
),
<collectingCountry id="09AF766BFFF42E2376B4FD4CFBE9FCAE" box="[997,1056,757,781]" name="United States of America" pageId="18" pageNumber="19">USA</collectingCountry>
. Approximate geographic coordinates: 27.3, -81.4.
</paragraph>
<paragraph id="710736FBFFF42E237598FC84FB89FCF6" blockId="18.[151,1437,757,1753]" box="[201,1088,829,853]" pageId="18" pageNumber="19">
<materialsCitation id="C1D03CA6FFF42E237598FC84FB89FCF6" box="[201,1088,829,853]" collectionCode="USNM" pageId="18" pageNumber="19" specimenCode="USNM 151210" specimenCount="1" typeStatus="holotype">
<emphasis id="43CCEAE9FFF42E237598FC84FEF5FCF6" bold="true" box="[201,316,829,853]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">
<typeStatus id="AE038859FFF42E237598FC84FEF1FCF6" box="[201,312,829,853]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
.
</emphasis>
A parthenogenetic female
<quantity id="B6409B1EFFF42E23773DFC84FD06FCF6" box="[620,719,829,853]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="18" pageNumber="19" unit="mm" value="1.67">1.67 mm</quantity>
long in alcohol,
<specimenCode id="211E9E80FFF42E2376DDFC84FBF5FCF6" box="[908,1084,829,853]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 151210</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="710736FBFFF42E237598FCD8FE50FC62" blockId="18.[151,1437,757,1753]" pageId="18" pageNumber="19">
<materialsCitation id="C1D03CA6FFF42E237598FCD8FA96FCDA" box="[201,1375,865,889]" collectionCode="USNM" pageId="18" pageNumber="19" specimenCode="USNM 151210" specimenCount="1" typeStatus="paratype">
<emphasis id="43CCEAE9FFF42E237598FCD8FE81FCDA" bold="true" box="[201,328,865,889]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">
<typeStatus id="AE038859FFF42E237598FCD8FE8AFCDA" box="[201,323,865,889]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Two parthenogenetic females in alcohol and one mounted in glycerine jelly,
<specimenCode id="211E9E80FFF42E2371FEFCD8FA96FCDA" box="[1199,1375,865,889]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 151210</specimenCode>
</materialsCitation>
.
<materialsCitation id="C1D03CA6FFF42E23703BFCDBFC10FC3E" collectionCode="NHM" pageId="18" pageNumber="19" specimenCode="NHM 1974" specimenCount="1" typeStatus="paratype">
Two parthenogenetic females in alcohol and one in glycerine jelly,
<specimenCode id="211E9E80FFF42E237604FC3FFC10FC3E" box="[853,985,901,925]" collectionCode="NHM" pageId="18" pageNumber="19">NHM 1974</specimenCode>
</materialsCitation>
.
<materialsCitation id="C1D03CA6FFF42E2376B5FC3CFE5CFC62" collectionCode="DGF" pageId="18" pageNumber="19" specimenCount="1" typeStatus="paratype">
716-718. All other specimens and exuvial fragments in
<collectionCode id="17A9AE3EFFF42E23740AFC10FE5CFC62" box="[347,405,937,961]" pageId="18" pageNumber="19">DGF</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="710736FBFFF42E237598FC74FAB4FBAA" blockId="18.[151,1437,757,1753]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598FC74FE39FC46" bold="true" box="[201,496,973,998]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Material examined here.</emphasis>
<materialsCitation id="C1D03CA6FFF42E2374A9FC77FAB1FBAA" country="United States of America" location="Florida" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Florida">
Many females from
<collectingCountry id="09AF766BFFF42E2377B1FC74FCE4FC46" box="[736,813,973,997]" name="United States of America" pageId="18" pageNumber="19">U.S.A.</collectingCountry>
(
<collectingRegion id="B37CF819FFF42E23766DFC74FC5BFC46" box="[828,914,973,997]" country="United States of America" name="Florida" pageId="18" pageNumber="19">Florida</collectingRegion>
,
<collectingRegion id="B37CF819FFF42E2376CEFC74FB81FC46" box="[927,1096,973,997]" country="United States of America" name="South Carolina" pageId="18" pageNumber="19">South Carolina</collectingRegion>
and
<collectingRegion id="B37CF819FFF42E2371D3FC74FAF9FC46" box="[1154,1328,973,997]" country="United States of America" name="North Carolina" pageId="18" pageNumber="19">North Carolina</collectingRegion>
). Full list will be represented in a special future publication concerning a revision of the
<taxonomicName id="B6B84D78FFF42E237157FC48FBB2FBAA" baseAuthorityName="O. F. Muller" baseAuthorityYear="1776" box="[1030,1147,1009,1033]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lamellatus">
<emphasis id="43CCEAE9FFF42E237157FC48FBB2FBAA" box="[1030,1147,1009,1033]" italics="true" pageId="18" pageNumber="19">lamellatus</emphasis>
</taxonomicName>
-like forms worldwide
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF42E237598FBACFD99FA1A" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="710736FBFFF42E237598FBACFD50FA36" blockId="18.[151,1437,757,1753]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598FBACFDA0FB8D" bold="true" box="[201,617,1045,1070]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598FBACFE8AFB8D" box="[201,323,1045,1070]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Diagnosis.</emphasis>
Parthenogenetic female.
</emphasis>
Dorsal head pores on a transverse fold in posterior portion of head shield with an indentation behind it. In anterior view, body strongly compressed laterally, median keel sharp, passes through whole carapace dorsum and actually begins on the head behind head pores. Dorsal margin of head flattened between compound eye and head pore region. Rostrum short. Ocellus minute. Lateral head pore small, rounded. Labrum with a median keel of moderate size, terminating in a widely rounded apex reaching distal end of antenna I. Postabdomen with sub-parallel dorsal and ventral margins, preanal teeth pointed. Spines at base of pre-claw portion predominantly clustered. Antenna I with antennular sensory seta arising in middle. Denticles in rows encircling antennular surface small. On antenna II, spine situated on proximal segment of exopod shorter than second segment. Limb I IDL with a hook-like seta markedly thicker than longest seta, the smallest IDL seta short. IDL with distal, proximal, marginal and basal group spinules. Eight setae in filter plate II, 9 setae in filter plate III; 9 setae in filter plate IV, 8 setae in filter plate V.
</paragraph>
<paragraph id="710736FBFFF42E237598FA18FD99FA1A" blockId="18.[151,1437,757,1753]" box="[201,592,1441,1466]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598FA18FE44FA19" bold="true" box="[201,397,1441,1466]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Full description.</emphasis>
See
<bibRefCitation id="15294B0AFFF42E237492FA1BFD85FA1A" author="Frey, D. G." box="[451,588,1441,1466]" pageId="18" pageNumber="19" pagination="1 - 25" refId="ref25888" refString="Frey, D. G. (1978) A new species of Eurycercus (Cladocera, Chydoridae) from the Southern United States. Tulane Studies in Zoology and Botany, 20, 1 - 25." type="journal article" year="1978">Frey (1978)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF42E237598FA7CFE3CF9A1" pageId="18" pageNumber="19" type="discussion">
<paragraph id="710736FBFFF42E237598FA7CFE3CF9A1" blockId="18.[151,1437,757,1753]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598FA7CFE98FA7D" bold="true" box="[201,337,1477,1502]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Comments.</emphasis>
It is a characteristic taxon from the south of the
<collectingCountry id="09AF766BFFF42E2376C4FA7CFC2DFA7E" box="[917,996,1477,1501]" name="United States of America" pageId="18" pageNumber="19">U.S.A.</collectingCountry>
, and is the most laterally compressed among all species of the genus.
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF42E237598F9B4FD1FF9EA" pageId="18" pageNumber="19" type="distribution">
<paragraph id="710736FBFFF42E237598F9B4FD1FF9EA" blockId="18.[151,1437,757,1753]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598F9B4FEABF985" bold="true" box="[201,354,1549,1574]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Distribution.</emphasis>
Southern states of the
<collectingCountry id="09AF766BFFF42E237724F9B4FD0BF986" box="[629,706,1549,1573]" name="United States of America" pageId="18" pageNumber="19">U.S.A.</collectingCountry>
The species is recorded from
<collectingRegion id="B37CF819FFF42E23717BF9B4FB10F986" box="[1066,1241,1549,1573]" country="United States of America" name="North Carolina" pageId="18" pageNumber="19">North Carolina</collectingRegion>
,
<collectingRegion id="B37CF819FFF42E2371B8F9B4FA51F986" box="[1257,1432,1549,1573]" country="United States of America" name="South Carolina" pageId="18" pageNumber="19">South Carolina</collectingRegion>
,
<collectingRegion id="B37CF819FFF42E2375C6F988FF3AF9EA" box="[151,243,1585,1609]" country="United States of America" name="Georgia" pageId="18" pageNumber="19">Georgia</collectingRegion>
,
<collectingRegion id="B37CF819FFF42E2375AFF988FE9BF9EA" box="[254,338,1585,1609]" country="United States of America" name="Florida" pageId="18" pageNumber="19">Florida</collectingRegion>
,
<collectingRegion id="B37CF819FFF42E23740CF988FE02F9EA" box="[349,459,1585,1609]" country="United States of America" name="Louisiana" pageId="18" pageNumber="19">Louisiana</collectingRegion>
and
<collectingRegion id="B37CF819FFF42E237753F98BFD8AF9EA" box="[514,579,1586,1609]" country="United States of America" name="Texas" pageId="18" pageNumber="19">Texas</collectingRegion>
(
<bibRefCitation id="15294B0AFFF42E237703F988FD03F9EA" author="Frey, D. G." box="[594,714,1585,1609]" pageId="18" pageNumber="19" pagination="1 - 25" refId="ref25888" refString="Frey, D. G. (1978) A new species of Eurycercus (Cladocera, Chydoridae) from the Southern United States. Tulane Studies in Zoology and Botany, 20, 1 - 25." type="journal article" year="1978">Frey 1978</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39A26570FFF42E237598F9ECFB59F97A" pageId="18" pageNumber="19" type="biology_ecology">
<paragraph id="710736FBFFF42E237598F9ECFB59F97A" blockId="18.[151,1437,757,1753]" pageId="18" pageNumber="19">
<emphasis id="43CCEAE9FFF42E237598F9ECFEE4F9CE" bold="true" box="[201,301,1621,1645]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">Ecology.</emphasis>
<bibRefCitation id="15294B0AFFF42E237465F9EFFE70F9CE" author="Frey, D. G." box="[308,441,1621,1646]" pageId="18" pageNumber="19" pagination="1 - 25" refId="ref25888" refString="Frey, D. G. (1978) A new species of Eurycercus (Cladocera, Chydoridae) from the Southern United States. Tulane Studies in Zoology and Botany, 20, 1 - 25." type="journal article" year="1978">Frey (1978)</bibRefCitation>
reported this taxon only from lakes, but he also said that &quot;the smaller water bodies, such as ponds, back-waters, slow streams, etc., particularly those with an abundance opf macrophytes, and hence some protection against predation by fishes, will yield many records of the species when they are studied intensively&quot; (
<bibRefCitation id="15294B0AFFF42E2375CFF978FEDCF97A" author="Frey, D. G." box="[158,277,1729,1753]" pageId="18" pageNumber="19" pagination="1 - 25" refId="ref25888" refString="Frey, D. G. (1978) A new species of Eurycercus (Cladocera, Chydoridae) from the Southern United States. Tulane Studies in Zoology and Botany, 20, 1 - 25." type="journal article" year="1978">Frey 1978</bibRefCitation>
, p. 24). Our study confirms this idea as we also found
<taxonomicName id="B6B84D78FFF42E237629F97BFBEBF97A" authorityName="Frey" authorityYear="1978" box="[888,1058,1729,1753]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="microdontus">
<emphasis id="43CCEAE9FFF42E237629F97BFBEBF97A" box="[888,1058,1729,1753]" italics="true" pageId="18" pageNumber="19">E. microdontus</emphasis>
</taxonomicName>
in ponds.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="7296C8190E7FF1AB15697EAA6A4AE71F">A revision of the subgenus Eurycercus (Eurycercus) Baird, 1843 emend. nov. (Cladocera: Eurycercidae) in the Holarctic with the description of a new species from Alaska</mods:title>
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Key for determination of adult parthenogenetic females of the Holarctic species of
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<emphasis id="43CCEAE9FFF92E2E71C4FA56FAD6F9AA" bold="true" box="[1173,1311,1519,1545]" italics="true" pageId="31" pageNumber="32">Eurycercus</emphasis>
(
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emend. nov.
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<keyLead id="CA4995CEFFF92E2E75C6F9E0FA55F928" pageId="31" pageNumber="32">1 Dorsal head pores lack a bubble-like projection and are located on a prominent transverse fold (indentation); a median keel passes through whole carapace dorsum.................................................................... 2</keyLead>
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<keyLead id="CA4995CEFFF92E2E75C6F92DFA55F965" pageId="31" pageNumber="32">- Dorsal head pores possess a bubble-like projection and are located on a prominent transverse fold (indentation) or on a flat head shield; a median keel is completely absent or expressed only in the posterior portion of the carapace dorsum......... 3</keyLead>
</paragraph>
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2 Small (seldom as large as
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), body very strongly compressed laterally, dorsal margin of head flattened or even slightly concave between compound eye and median head pore, the smallest seta IDL very short............
<taxonomicName id="B6B84D78FFF92E2E71CBF955FA55F8A2" ID-CoL="3D3KB" authority="Frey, 1978" authorityName="Frey" authorityYear="1978" box="[1178,1436,1772,1793]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="microdontus">
<emphasis id="43CCEAE9FFF92E2E71CBF955FAF9F8A2" box="[1178,1328,1772,1793]" italics="true" pageId="31" pageNumber="32">E. microdontus</emphasis>
<bibRefCitation id="15294B0AFFF92E2E7067F955FA55F8A2" author="Frey, D. G." box="[1334,1436,1772,1793]" pageId="31" pageNumber="32" pagination="1 - 25" refId="ref25888" refString="Frey, D. G. (1978) A new species of Eurycercus (Cladocera, Chydoridae) from the Southern United States. Tulane Studies in Zoology and Botany, 20, 1 - 25." type="journal article" year="1978">Frey, 1978</bibRefCitation>
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- Large species, body moderately compressed laterally, dorsal margin of head regularly convex, the smallest seta of IDL of moderate size.................................................................
<taxonomicName id="B6B84D78FFF92E2E7107F89FFA55F898" ID-CoL="6HF72" authority="(O. F. Muller, 1776)" baseAuthorityName="O. F. Muller" baseAuthorityYear="1776" box="[1110,1436,1830,1851]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="lamellatus">
<emphasis id="43CCEAE9FFF92E2E7107F89FFB1CF898" box="[1110,1237,1830,1851]" italics="true" pageId="31" pageNumber="32">E. lamellatus</emphasis>
(O. F.
<bibRefCitation id="15294B0AFFF92E2E7046F89FFA5CF898" author="Muller, O. F." box="[1303,1429,1830,1851]" pageId="31" pageNumber="32" pagination="1 - 282" refId="ref27693" refString="Muller, O. F. (1776) Zoologiae Danicae prodromus, seu animalium Daniae et Norvegiae indigenarum characteres, nomina, et synonyma imprimis popularium. (Zoologiae Danicae Prodromus). Typis Hallageriis, N. Mulleri aviae regiae typographie et filii, Havniae (Copenhagen), pp. i - xxxii + 1 - 282." type="book chapter" year="1776">Müller, 1776</bibRefCitation>
)
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3 Bubble-like projection on prominent transverse fold, median keel present in posterior portion of carapace dorsum....................................................................................................
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<emphasis id="43CCEAE9FFF92E2E71BCF8D8FA85F8D5" box="[1261,1356,1889,1910]" italics="true" pageId="31" pageNumber="32">E. beringi</emphasis>
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.
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<keyLead id="CA4995CEFFF92E2E75C6F8C7FA55F830" box="[151,1436,1918,1939]" pageId="31" pageNumber="32">- Bubble-like projection on flat head shield, median keel absent................................................. 4</keyLead>
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4 Median labral keel short, reaching approximately middle of antenna I......................
<taxonomicName id="B6B84D78FFF92E2E712EF825FA55F812" ID-CoL="3D3K8" authority="Frey, 1975" authorityName="Frey" authorityYear="1975" box="[1151,1436,1948,1969]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pompholygodes">
<emphasis id="43CCEAE9FFF92E2E712EF825FAE6F812" box="[1151,1327,1948,1969]" italics="true" pageId="31" pageNumber="32">E. pompholygodes</emphasis>
<bibRefCitation id="15294B0AFFF92E2E7064F825FA55F812" author="Frey, D. G." box="[1333,1436,1948,1969]" pageId="31" pageNumber="32" pagination="263 - 300" refId="ref25854" refString="Frey, D. G. (1975) Subgeneric differentiation within Eurycercus (Cladocera, Chydoridae) and a new species from Northern Sweden. Hydrobiologia, 46, 263 - 300." type="journal article" year="1975">Frey, 1975</bibRefCitation>
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<keyLead id="CA4995CEFFF92E2E75C6F800FA55F86D" box="[151,1436,1977,1998]" pageId="31" pageNumber="32">- Median labral keel large, approximately reaching tip of antenna I................................................. 5</keyLead>
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<paragraph id="710736FBFFF92E2E75C6F86FFA55F848" blockId="31.[151,1436,1625,2027]" box="[151,1436,2006,2027]" pageId="31" pageNumber="32">
<keyLead id="CA4995CEFFF92E2E75C6F86FFA55F848" box="[151,1436,2006,2027]" pageId="31" pageNumber="32">
5 Apex of median labral keel distinctly angled....................................
<taxonomicName id="B6B84D78FFF92E2E7115F86FFA55F848" ID-CoL="3D3KK" authority="Tanaka &amp; Fujita, 2002" authorityName="Tanaka &amp; Fujita" authorityYear="2002" box="[1092,1436,2006,2027]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nipponica">
<emphasis id="43CCEAE9FFF92E2E7115F86FFB76F848" box="[1092,1215,2006,2027]" italics="true" pageId="31" pageNumber="32">E. nipponica</emphasis>
<bibRefCitation id="15294B0AFFF92E2E7195F86FFA55F848" author="Tanaka, M. &amp; Fujita, S." box="[1220,1436,2006,2027]" pageId="31" pageNumber="32" pagination="1 - 25" refId="ref28361" refString="Tanaka, M. &amp; Fujita, S. (2002) Study on the plankton from Lake Utonai-numa, Hokkaido. Yokkaichi University Journal of Environmental and Information Sciences, 6, 1 - 25 (in Japanese)." type="journal article" year="2002">Tanaka &amp; Fujita, 2002</bibRefCitation>
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<keyLead id="CA4995CEFFC62E1175C6FF2EFA55FF0F" box="[151,1436,151,172]" pageId="32" pageNumber="33">- Apex of median labral keel rounded.......................................................................... 6</keyLead>
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</keyStep>
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<paragraph id="710736FBFFC62E1175C6FF0CFA55FF69" blockId="32.[151,1436,151,231]" box="[151,1436,181,202]" pageId="32" pageNumber="33">
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6 Hook-like seta on IDL especially strong, its base occupies the whole top of IDL...............
<taxonomicName id="B6B84D78FFC62E1171DCFF0CFA55FF69" ID-CoL="3D3K7" authority="Frey, 1973" authorityName="Frey" authorityYear="1973" box="[1165,1436,181,202]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="macracanthus">
<emphasis id="43CCEAE9FFC62E1171DCFF0CFAF9FF69" box="[1165,1328,181,202]" italics="true" pageId="32" pageNumber="33">E. macracanthus</emphasis>
Frey, 1973
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<paragraph id="710736FBFFC62E1175C6FF6BFA55FF44" blockId="32.[151,1436,151,231]" box="[151,1436,210,231]" pageId="32" pageNumber="33">
<keyLead id="CA4995CEFFC62E1175C6FF6BFA55FF44" box="[151,1436,210,231]" pageId="32" pageNumber="33">
- Hook-like seta on IDL not so strong, its base occupies only a part of top of IDL................
<taxonomicName id="B6B84D78FFC62E1171CDFF6BFA55FF44" ID-CoL="3D3K6" authority="Hann, 1982" authorityName="Hann" authorityYear="1982" box="[1180,1436,210,231]" class="Branchiopoda" family="Eurycercidae" genus="Eurycercus" kingdom="Animalia" order="Diplostraca" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="43CCEAE9FFC62E1171CDFF6BFAEFFF44" box="[1180,1318,210,231]" italics="true" pageId="32" pageNumber="33">E. longirostris</emphasis>
<bibRefCitation id="15294B0AFFC62E11707DFF6BFA55FF44" author="Hann, B. J." box="[1324,1436,210,231]" pageId="32" pageNumber="33" pagination="585 - 610" refId="ref26253" refString="Hann, B. J. (1982) Two new species of Eurycercus (Bullatifrons) from Eastern North America (Chydoridae, Cladocera). Taxonomy, ontogeny, and biology. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 67, 585 - 610." type="journal article" year="1982">Hann, 1982</bibRefCitation>
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<mods:namePart>Moore, Matthew R.</mods:namePart>
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<mods:nameIdentifier type="email">cyclocephala@gmail.com</mods:nameIdentifier>
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</mods:role>
<mods:namePart>Cave, Ronald D.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida, Indian River Research and Education Center, 2199 South Rock Road, Fort Pierce, FL 34945, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
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</mods:role>
<mods:namePart>Branham, Marc A.</mods:namePart>
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<mods:part>
<mods:date>2018</mods:date>
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<mods:number>2018-03-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>745</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>101</mods:start>
<mods:end>378</mods:end>
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</mods:mods>
<treatment ID-GBIF-Taxon="182229962" LSID="urn:lsid:plazi:treatment:F9118EF9AD00510EB0749C7A3D8B3977" httpUri="http://treatment.plazi.org/id/F9118EF9AD00510EB0749C7A3D8B3977" lastPageNumber="109" pageId="8" pageNumber="109">
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Tribe
<taxonomicName LSID="F9118EF9-AD00-510E-B074-9C7A3D8B3977" authority="Laporte, 1840" authorityName="Laporte" authorityYear="1840" family="Scarabaeidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="109" rank="tribe" tribe="Cyclocephalini">Cyclocephalini Laporte, 1840</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="109" type="reference_group">
<paragraph pageId="8" pageNumber="109">
<taxonomicName family="Scarabaeidae" genus="Cyclocephalites" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="109" rank="genus">Cyclocephalites</taxonomicName>
Laporte, 1840: 124 [original usage].
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<paragraph pageId="8" pageNumber="109">
<taxonomicName family="Cyclocephalidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="109" rank="family">Cyclocephalidae</taxonomicName>
[Burmeister, 1847: 21].
</paragraph>
<paragraph pageId="8" pageNumber="109">
<taxonomicName family="Scarabaeidae" genus="Cyclocephalides" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="109" rank="genus">Cyclocephalides</taxonomicName>
[Lacordaire, 1856: 393].
</paragraph>
<paragraph pageId="8" pageNumber="109">
<taxonomicName authorityName="Laporte de Castelnau" authorityYear="1840" family="Scarabaeidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="109" rank="tribe" tribe="Cyclocephalini">Cyclocephalini</taxonomicName>
[LeConte, 1862: 143].
</paragraph>
<paragraph pageId="8" pageNumber="109">
<taxonomicName family="Scarabaeidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="109" rank="subFamily" subFamily="Cyclocephalinae">Cyclocephalinae</taxonomicName>
[Bates, 1888: 296].
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="109" type="type genus">
<paragraph pageId="8" pageNumber="109">Type genus.</paragraph>
<paragraph pageId="8" pageNumber="109">
<taxonomicName authorityName="Dejean" authorityYear="1821" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala" order="Coleoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="109">Cyclocephala</emphasis>
</taxonomicName>
Dejean, 1821.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,207 @@
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<paragraph id="DBF56D4AB779E099A7952C559AC5ACFB" pageId="0" pageNumber="29443">Materials</paragraph>
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; verbatimElevation:
<elevation id="22268BE949FD612276029D8AC627F078" pageId="0" pageNumber="29443">943.48</elevation>
; decimalLatitude:
<geoCoordinate id="9E9C885DDAA02B3F34B5EC6DA8088C3E" orientation="latitude" pageId="0" pageNumber="29443" value="43.17227">43.17227</geoCoordinate>
; decimalLongitude:
<geoCoordinate id="91EA898221D8EA3E2C567D2D060DFF36" orientation="longitude" pageId="0" pageNumber="29443" value="-4.90857">-4.90857</geoCoordinate>
; geodeticDatum: WGS84; Event: eventID: E; samplingProtocol:
<collectingMethod id="8A4909FF8F69D326968298CE965FA030" pageId="0" pageNumber="29443">Pitfall</collectingMethod>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0E73183399CF2CB39D6E79A7560BA780" pageId="0" pageNumber="29443" type="distribution">
<paragraph id="F8DFCEFD25F7C961FAE07600671A5E87" pageId="0" pageNumber="29443">Distribution</paragraph>
<paragraph id="49C5D3807EFF61AD94825A28ED4381EA" pageId="0" pageNumber="29443">Europe to Central Asia</paragraph>
</subSubSection>
</treatment>
</document>