diff --git a/data/31/57/E0/3157E00AFFF2FFC6FDA40999FAA2DBF1.xml b/data/31/57/E0/3157E00AFFF2FFC6FDA40999FAA2DBF1.xml new file mode 100644 index 00000000000..b2bd4524748 --- /dev/null +++ b/data/31/57/E0/3157E00AFFF2FFC6FDA40999FAA2DBF1.xml @@ -0,0 +1,767 @@ + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +Author + +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be + + + +Author + +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org + + + +Author + +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be + +text + + +European Journal of Taxonomy + + +2024 + +2024-10-23 + + +966 + + +1 + + +1 +80 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 + + + + +Family + +Udubidae +Griswold & +Polotow, 2015 + + + + + +Udubidae Griswold & Polotow +in + +Polotow +et al. +, 2015: 151 + +. + + + + + + +Type +genus + + + + + +Uduba +Simon, 1880: 343 + +( +Type +species by monotypy: + +Olios madagascariensis +Vinson, 1863 + += + +Uduba madagascariensis +( +Vinson, 1863 +)) + +. + + + + + +Diagnosis + + + +Udubidae +are entelegyne RTA-clade spiders with three tarsal claws ( +Fig. 21A–B +), multiple rows of tarsal trichobothria of about equal length ( +Figs 13F +, +20E–F +), eight eyes in 2 nearly straight rows ( +Figs 3C, F +, 8A–B, 13C–D, 17A, 22A, C, 24A, C), indirect eyes with canoe-shaped tapeta and the cheliceral fang with interior serrula weak to absent. Males may or may not have a subbasal crack or line of autospasy on the leg tibiae ( +Fig. 24B +, arrow; +Griswold 1993 +: figs 3–4). All species have a hyaline conductor (C = the “Sierwald conductor” of + +Polotow +et al. +2015 + +), which is relatively small, transverse, hyaline and fan-shaped, prolaterally extending from narrow base to wide flattened tip ( +Figs 5A–C +, +6D–F +, +9A–C +, +11A–C +, +12E +, +14D, F +, +15D–F +). Most have an additional tegular process accompanying the embolus that is a narrow membranous or sclerotized extension inserted between the tegulum and the base of the median apophysis (MA), running backward to MA and ending with fattened fan-shaped tip accompanying the embolus: this is the paraconductor (Pc = the “MTP” and “STP” of +Griswold 1993 +) ( +Figs 5A–C +, +6D–F +, +9A–C +, +11A–C +, +12E +, +14D +, +15D–F +). The paraconductor is found in other members of the Oval Calamistrum (OC) clade (the “TA” in + +Griswold +et al. +2005 + +: figs 186B–C, 195B–C), and the Sierwald conductor (C) occurs widely among the OC clade and Dionycha +Petrunkevitch, 1928 +( + +Polotow +et al. +2015 + +: figs. 13E–F, 15A–B, E–F). + + +Udubidae +may be distinguished from most other araneomorph spider families found in +Madagascar +by the form of their tarsi. The presence of tarsal trichobothria distinguishes udubids from the + +Filistatidae +Ausserer, 1867 + +, Synspermiata +Michalik & Ramírez, 2014 +, +Palpimanoidea + +Forster & +Platnick +, 1984 + +, Araneoidea +Latreille, 1806 +, +Eresidae C.L. Koch, 1845 +(in +Berendt 1845 +), + +Hersiliidae +Thorell, 1869 + +and + +Oecobiidae +Blackwall, 1862 + +, + +Phyxelididae +Lehtinen, 1967 + +and + +Titanoecidae +Lehtinen, 1967 + +. Multiple rows of tarsal trichobothria distinguish +Udubidae +from taxa with only a single row of tarsal trichobothria including + +Agelenidae C.L. +Koch, 1837 + +, + +Dictynidae O. +Pickard-Cambridge, 1871 + +, + +Hahniidae +Bertkau, 1878 + +and most + +Zodariidae +Thorell, 1881 + +and + +Thomisidae +Sundevall, 1833 + +. Among taxa with multiple rows of tarsal trichobothria udubids may be distinguished from + +Lycosidae +Sundevall, 1833 + +, + +Pisauridae +Simon, 1890 + +and + +Oxyopidae + +Thorell, +1869 + + +in that these latter families have at least one eye row strongly curved, whereas +Udubidae +have both eye rows nearly straight. Udubids have at least a rudiment of the inferior tarsal claw and lack claw tufts, characters that distinguish them from + +Sparassidae +Bertkau, 1872 + +and the Dionycha ( + +Cheiracanthiidae +Wagner, 1887 + +, + +Clubionidae +Simon, 1878 + +, + +Corinnidae +Karsch, 1880 + +, + +Gallieniellidae +Millot, 1947 + +, + +Gnaphosidae +Banks, 1892 + +, + +Liocranidae +Simon, 1897 + +, + +Philodromidae +Thorell, 1869 + +, + +Salticidae +Blackwall, 1841 + +, + +Selenopidae +Simon, 1897 + +, + +Trachelidae +Simon, 1897 + +, + +Trochanteriidae +Karsch, 1879 + +and + +Viridasiidae +Lehtinen, 1967 + +). Finally, like +Udubidae +, the lone genus of + +Desidae +Pocock, 1895 + +from +Madagascar +, + +Desis +Walckenaer, 1837 + +, has numerous tarsal trichobothria and straight eye rows but the latter is quite distinct in having elongate chelicerae, a dense cover of fine hairs and the posterior tracheal spiracle on the abdomen advanced. + + + +Composition + + + + +Campostichomma +Karsch, 1892 + +( +Sri Lanka +), + +Raecius +Simon, 1892 + +(Africa), and + +Tabiboka + +gen. nov. +, + +Uduba +Simon 1880 + +, + +Zorascar + +gen. nov. +and + +Zorodictyna +Strand, 1907 + +( +Madagascar +). + + + + + + +Key to +Udubidae +of +Madagascar + + + + + + + +1. Male palp without process between tibia and patella; cymbium with retrobasal extension (CE), short to as long as rest of cymbium ( +Figs 5B–C +, +6A–C +, +9B–C +, +11B–C +, +12B–D +, +14A–C +, +15A–B +, +16A–B +, +23B–C +, +25C +); embolus broad and flat to slender, cylindrical, firmly attached to point of origin on retromargin of tegulum ( +Figs 5A–B +, +6D–E +, 8F, 9A–B, 11A–B, 12E, 14E–F, 15B, D–F, 23A–B, 25A–B); tegulum oval apically; conductor hyaline, fan-shaped crest arising from middle to apex of tegulum; female endogyne with spermathecal ducts short, stout, not making loops, at most twisted ( +Figs 5E +, +7C–E +, +9E +, +11E +, +12H +, +16D +, +18B–D +, +19C–F +, +23E +, +25E +) ..................................................... 2 + + + + +– Male palp with small ventral process at base of tibia between tibia and patella (“ +UTA +” in + +Griswold +et al. +2022 + +: figs 27A–B, 39A–C, 40A–C, 41B); cymbium rounded retrobasally; embolus flexibly attached to large, swollen base, embolus elongate and extending behind tegulum and subtegulum to emerge from alveolus near bulb apex ( + +Griswold +et al. +2022 + +: figs 27A–B, 32C, 41F, 51A); tegulum trilobed apically, with retrolateral (TA1), median (TA2) and prolateral (TA3) processes interacting apically ( + +Griswold +et al. +2022 + +: figs 27A–B, 29A–I, 51A–C); conductor rectangular hyaline crest arising along apical margin of prolateral tegular lobe near TA3; female endogyne with spermathecal ducts elongate, making at least one (to as many as five) loops ( + +Griswold +et al. +2022 + +: figs 68B–D, 80B–D, F–G, I–J) ............................................................................................................................... ......... + +Uduba +Simon, 1880 + +(the key to + +Uduba +species + +is amply detailed in + +Griswold +et al. +2022: 13 + +) + + + + + + +2. With entire cribellum or vestige; PLS and ALS equal in length; strong scopulae beneath leg tarsi and most of length of metatarsi; males with crack-suture on leg tibiae just anteriad of basalmost pair of ventral spines ( +Fig. 24B +) ........................................................................3. + +Zorodictyna +Strand, 1907 + + + + +– Ecribellate; PLS much smaller than ALS; scopulae absent or only with weak scopulae beneath leg tarsi; most males without crack-suture on leg tibiae ........................................................................ 5 + + + + + +3. Male palpal cymbium with rounded retrolateral paracymbial process (PP) ( +Figs 23B–C +, +25B–C +); cymbial retrobasal extension (CE) extending at least one-third length of tibia ( +Figs 23B–C +, +25B–C +); female epigyne lateral lobes (LL) lacking teeth ( +Figs 23D +, +25D +) .................................................... 4 + + + + +– Male palpal cymbium evenly convex retrolaterally, without paracymbial process ( +Griswold 1993 +: figs 20–21); cymbial retrobasal extension, short, extending only to tibial apex ( +Griswold 1993 +: figs 20–21); female epigyne lateral lobes with short teeth ( +Griswold 1993 +: fig. 24) ......................... ...................................................................................................... + +Zorodictyna oswaldi +( +Lenz, 1891 +) + + + + + + + +4. Male palp with basal embolic process (EBP) nearly triangular, embolus lacking crest, tegulum with retromedian longitudinal ridge that extends for most of tegulum length ( +Fig. 23A–C +); female epigyne lateral lobes (LL) parallel, median sector (MS) rectangular ( +Fig. 23D +); female endogyne with copulatory ducts (CD) larger than spermathecae ( +Fig. 23E +) .......... + +Zorodictyna almae + +sp. nov. + + + + +– Male palp with basal embolic process (EBP) slender, crescentic, embolus with subapical distal crest, and tegulum retromedian longitudinal ridge weak, short, extending for less than one-half tegulum length ( +Figs 25A–C +); female epigyne lateral lobes (LL) converging posteriorly, median sector (MS) triangular ( +Fig. 25D +); female endogyne with copulatory ducts (CD) and spermathecae equal in size ( +Fig. 25E +) ....................................................................................... + +Zorodictyna silvadavilae + +sp. nov. + + + + + + +5. Male palp with cymbial retrobasal extension (CE) elongate, extending most of length of tibia, palpal tibia with RTA only, lacking VTA ( +Figs 14A–B +, +15A–B +, +16A–B +); female epigyne median lobe (ML) short, concave medially, not extending to posterior margin, copulatory openings (CO) exposed ( +Figs 16C, E +, +18A +) ............................................................................................6. + +Zorascar + +gen. nov. + + + + +– Male palp with cymbial retrobasal extension (CE) shorter, extending to or just past tibial apex, palpal tibia with both VTA and RTA ( +Figs 5A–C +, +6A–C +, +9A–C +, +11A–C +, +12A–D +); female epigyne median lobe (ML) extends to posterior margin, separating lateral lobes (LL) and covering copulatory openings (CO) ( +Figs 4B +, +5D +, +7A–B +, +9D +, +11D +, +12G +) .......................................................7. + +Tabiboka + +gen. nov. + + + + + + +6. Female epigyne median lobe (ML) broad and shallowly concave medially, copulatory openings (CO) exposed anteromedially ( +Fig. 16C +), spermathecae elongate, curved, eggplant-shaped ( +Fig. 16D +); male palp as in figures ( +Figs 14A–F +, +15A–F +, +16A–B +) ..................... + +Zorascar pasunepipe + +gen. et sp. nov. + + + + +– Female epigyne median lobe (ML) narrow and deeply concave medially, copulatory openings (CO) exposed laterad of median lobe ( +Figs 16E +, +18A +, +19A–B +), spermathecae stout, rounded, reniform ( +Figs 18B–D +, +19C–F +); male unknown ................................ + +Zorascar pasunepomme + +gen. et sp. nov. + + + + + + +7. Male palp with conductor base originating anteriorly, hidden behind embolus or paraconductor (Pc), embolus short, arising retrolaterally near 270° ( +Figs 5A–C +; +6D–F +, 8F, 9A–C); female epigyne median lobe (ML) heart-shaped, sides rounded ( +Figs 4B +, +5D +, +7A +, +9D +) ....................................................... 8 + + + + +– Male palp conductor originating medially, near embolus base, not hidden, embolus elongate, arising retrobasally near 200° ( +Figs 11A–C +, +12A–C, E +); female epigyne median lobe (ML) rectangular, narrowest in middle or with sides straight ( +Figs 11D +, +12G +) ...... + +Tabiboka polotowae + +gen. et sp. nov. + + + + + + +8. Male palpal tibia with elongate retrobasal process (RTA), embolus broad, median apophysis (MA) short, apex far proximad of embolus apex ( +Figs 5A–C +, +6D–F +); female epigyne median lobe (ML) narrow anteriorly, anterior width less than one-half total width ( +Figs 4B +, +5D +, +7A +), endogyne with each spermatheca as broad as long ( +Figs 5E +, +7C–E +) ...................... + +Tabiboka milleri + +gen. et sp. nov. + + + + +– Male palpal tibia with short, blunt retromedian process (RTA), embolus slender; median apophysis (MA) elongate, cruciform, apex extends to embolus apex (Figs 8C–F, 9A–C); female epigyne median lobe (ML) broad anteriorly, anterior width greater than one-half total width ( +Fig. 9D +), endogyne with each spermatheca narrow, longer than broad ( +Fig. 9E +) ................... + +Tabiboka milloti + +gen. et sp. nov. + + + + + + + \ No newline at end of file diff --git a/data/31/57/E0/3157E00AFFF6FFDDFDF10CBEFD20D971.xml b/data/31/57/E0/3157E00AFFF6FFDDFDF10CBEFD20D971.xml index 39d7e1667a4..67c321e20b6 100644 --- a/data/31/57/E0/3157E00AFFF6FFDDFDF10CBEFD20D971.xml +++ b/data/31/57/E0/3157E00AFFF6FFDDFDF10CBEFD20D971.xml @@ -1,68 +1,68 @@ - - - -On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar + + + +On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar - - -Author + + +Author -Henrard, Arnaud -E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 -Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. -arnaud.henrard@africamuseum.be +Henrard, Arnaud +E1B02E6E-D91C-43FE-8D8C-CD102EFEE3B4 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +arnaud.henrard@africamuseum.be - - -Author + + +Author -Griswold, Charles -51E9CD73-DB08-49B6-9B2C-843B27B38B48 -Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. -cgriswold@calacademy.org +Griswold, Charles +51E9CD73-DB08-49B6-9B2C-843B27B38B48 +Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. +cgriswold@calacademy.org - - -Author + + +Author -Jocqué, Rudy -CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 -Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. -rudy.jocque@africamuseum.be +Jocqué, Rudy +CF15016C-8CD1-4C9D-9021-44CA7DC7A5D5 +Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium. +rudy.jocque@africamuseum.be -text - - -European Journal of Taxonomy +text + + +European Journal of Taxonomy - -2024 - -2024-10-23 + +2024 + +2024-10-23 - -966 + +966 - -1 + +1 - -1 -80 + +1 +80 - -https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453 -journal article -10.5852/ejt.2024.966.2697 -2118-9773 -14001790 -FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 +journal article +10.5852/ejt.2024.966.2697 +2118-9773 +14001790 +FFE41347-8B55-4CF1-B10B-53EBB4CF0E26 - + @@ -71,54 +71,68 @@ gen. et sp. nov. + + urn:lsid:zoobank.org:act: 33C8C4F1-A241-41E2-9B38-1D31AFBF4BE4 - - Figs 3–7 , 26 + + + Zorocratidae gen. Talatakely sp. Talatakely – + Morphbank 2023 + : images [476092–509952] SpiderAToL, March 3, 2008 , https://www.morphbank.net/?id=218853 [accessed 20 Oct. 2023 ]. + + Zorodictyna sp. 2 – + Polotow et al. 2015 + . + + Zorodictyna sp. CG300 – + Wheeler et al. 2017 + . + @@ -220,7 +234,7 @@ Miller - + MADAGASCAR @@ -231,7 +245,7 @@ Miller ; Ranomafana National Park , -Talatakely +Talatakely ; 21°15′ S , @@ -253,7 +267,9 @@ a.s.l. S. Larcher and R. Andriamasamanana -leg.; CASENT_9016094; +leg.; +CASENT_9016094 +; CAS . @@ -264,7 +280,7 @@ leg.; CASENT_9016094; - + MADAGASCAR @@ -272,17 +288,11 @@ leg.; CASENT_9016094; 1 ♀ -; same collection data as for holotype; [ -Standard -view preparations and images for AToL-Spiders project; SEM preparations -Box -79-8, 10, 11, 19, -Box -81, 1, 2]; CASENT_9016094; -CAS +; same collection data as for holotype; [Standard view preparations and images for AToL-Spiders project; SEM preparations Box 79-8, 10, 11, 19, Box 81, 1, 2]; CASENT_9016094; +CAS • - + 1 ♂ , 1 ♀ @@ -306,33 +316,36 @@ leg.; general collecting; [habitus photos, stacked images, Apr. 2005 -]; CASENT_9016098; -CAS +]; +CASENT_9016098 +; +CAS • - + 1♂ , 1 ♀ ; same data as for preceding; [ -palp drawn]; CASENT_9016086; -CAS +palp drawn]; +CASENT_9016086 +; +CAS • - + 1 ♂ , 1 ♀ -; Parc National -Ranomafana +; +Parc National Ranomafana , Talatekely forest , 42.3 km -58° NE of -Fianarantsoa +58° NE of Fianarantsoa ; 21°15′28.0″ S @@ -361,13 +374,17 @@ a.s.l. E. Rajeriarison , V. Andriamananony -leg.; montane rainforest; general collecting; [AToL DNA -Voucher -ARACG0300] [HW009/RNAlater: Mad-052]; CASENT_9024004; -CAS +leg.; +montane rainforest +; general collecting; [AToL DNA Voucher +ARACG0300 +] [HW009/RNAlater: Mad-052]; +CASENT_9024004 +; +CAS • - + 1 ♂ ; Ranomafana @@ -387,14 +404,16 @@ leg.; general collecting; [stacked images, HMW, April-2005 -]; CASENT_9016097; -CAS +]; +CASENT_9016097 +; +CAS • - + 1 ♀ -; Parc National -Ranomafana +; +Parc National Ranomafana , Vohiparara , @@ -427,13 +446,15 @@ a.s.l. N. Scharff and H. Wood -leg.; primary montane rainforest; general collecting day and night; [ -NS Photo -voucher 893-897; GH photos 5659-5666]; CASENT_9029871; -CAS +leg.; +primary montane rainforest +; general collecting day and night; [NS Photo voucher 893-897; GH photos 5659-5666]; +CASENT_9029871 +; +CAS . – - + Mahajanga Province @@ -443,8 +464,8 @@ voucher 893-897; GH photos 5659-5666]; CASENT_9029871; Ampijoroa , National Park Ankarafantsika -, near -Lake Ravelobe +, +near Lake Ravelobe ; 16°17′48″ S , @@ -459,10 +480,14 @@ voucher 893-897; GH photos 5659-5666]; CASENT_9029871; A. Saucedo , H. Wood -leg.; [SEM female epigyne + used for DNA voucher DP-2014c - “ +leg.; [SEM female epigyne + used for DNA voucher +DP-2014c +- “ Zorodictyna sp. 2 -”]; CASENT_9035866; -CAS +”]; +CASENT_9035866 +; +CAS . @@ -472,7 +497,7 @@ leg.; [SEM female epigyne + used for DNA voucher DP-2014c - “ - + MADAGASCAR @@ -483,7 +508,7 @@ leg.; [SEM female epigyne + used for DNA voucher DP-2014c - “ ; Tsinjoarivo Forest , -Mahatsinjo +Mahatsinjo ; 19°40′47.496″ S , @@ -499,37 +524,49 @@ a.s.l. ; B.L. Fisher -leg.; montane rainforest; BLF35410; CASENT_9065708; +leg.; +montane rainforest +; BLF35410; +CASENT_9065708 +; CAS • - + 1 ♂ -; same data as for preceding; CASENT_9065709; -CAS +; same data as for preceding; +CASENT_9065709 +; +CAS • - + 1 ♂ -; same data as for preceding; CASENT_9065678; -CAS +; same data as for preceding; +CASENT_9065678 +; +CAS • - + 1 ♂ -; same data as for preceding; CASENT_9065579; -CAS +; same data as for preceding; +CASENT_9065579 +; +CAS • - + 4 ♂♂ , 5 ♀♀ -; same data as for preceding; CASENT_9017817; -CAS +; same data as for preceding; +CASENT_9017817 +; +CAS • - + 2 ♀♀ ; Tsinjoarivo Forest @@ -551,11 +588,17 @@ a.s.l. ; B.L. Fisher et al. -leg.; montane rainforest, sifted litter; BLF33370; CASENT_9064661; -CAS +leg.; +montane rainforest +, +sifted litter +; BLF33370; +CASENT_9064661 +; +CAS . – - + Fianarantsoa Province @@ -569,9 +612,10 @@ leg.; montane rainforest, sifted litter; BLF33370; CASENT_9064661; SSW of Ambositra , + 4.7 km -SW of -Ambalamanakana +SW of Ambalamanakana + ; 20°47′ S , @@ -587,37 +631,49 @@ a.s.l. ; S. Goodman -, SG0001 leg.; disturbed upper montane forest; +, SG0001 leg.; +disturbed upper montane forest +; pitfall traps -#1–3; CASENT_9016064; -FMNH +#1–3; +CASENT_9016064 +; +FMNH • - + 9 ♀♀ -; same data as for preceding; CASENT_9016065; -FMNH +; same data as for preceding; +CASENT_9016065 +; +FMNH • - + 1 ♀ -; same data as for preceding; CASENT_9016066; -FMNH +; same data as for preceding; +CASENT_9016066 +; +FMNH • - + 9 ♀♀ -; same data as for preceding; CASENT_9016067; -FMNH +; same data as for preceding; +CASENT_9016067 +; +FMNH • - + 1 ♂ -; same data as for preceding; CASENT_9016068; -FMNH +; same data as for preceding; +CASENT_9016068 +; +FMNH • - + 1 ♂ ; Ankazomivady @@ -641,13 +697,17 @@ a.s.l. ; B.L. Fisher -leg.; forest; EBD10 +leg.; +forest +; EBD10 pitfall trap -; BLF1592b; CASENT_9016087; -CAS +; BLF1592b; +CASENT_9016087 +; +CAS • - + 1 ♀ ; Ambatofitorahana @@ -666,15 +726,15 @@ S of Ambositra ; A. Pauly -leg.; natural forest; BE_ -RMCA -_ARA. -Ara. -201626; -RMCA +leg.; +natural forest +; +BE_ RMCA_ARA.Ara.201626 +; +RMCA • - + 24 ♂♂ , 5 ♀♀ @@ -703,22 +763,28 @@ a.s.l. E. Elsom and F. Vargas -leg.; mixed tropical forest; +leg.; +mixed tropical forest +; pitfall traps -#4–16; COL-DHK-2001-001TN; CASENT_9003432; -CAS +#4–16; COL-DHK-2001-001TN; +CASENT_9003432 +; +CAS • - + 1 ♀ -; same data as for preceding; CASENT_9003534; -CAS +; same data as for preceding; +CASENT_9003534 +; +CAS • - + 1 ♀ -; Parc National -Ranomafana +; +Parc National Ranomafana , Talatakely ; @@ -745,20 +811,24 @@ and D. Ubick leg.; pitfall traps -; CASENT_9006115; -CAS +; +CASENT_9006115 +; +CAS • - + 1 ♂ -; same data as for preceding; CASENT_9016084; -CAS +; same data as for preceding; +CASENT_9016084 +; +CAS • - + 1 ♂ -; Parc National -Ranomafana +; +Parc National Ranomafana , Talatakely ; @@ -783,23 +853,31 @@ leg.; J. Schweikert , D. Ubick -leg.;general collecting; CASENT_9016082; -CAS +leg.; +general collecting +; +CASENT_9016082 +; +CAS • - + 2 ♀♀ -; same data as for preceding; CASENT_9016088; -CAS +; same data as for preceding; +CASENT_9016088 +; +CAS • - + 6 ♂♂ -; Parc National -Ranomafana +; +Parc National Ranomafana , Talatakely -, trail F; +, +trail F +; 21°16′ S , 47°25′ E @@ -818,26 +896,32 @@ and K.J. Ribardo leg.; pitfall trap -; CASENT_9016089; -CAS +; +CASENT_9016089 +; +CAS • - + 2 ♂♂ -; same data as for preceding; CASENT_9016090; -CAS +; same data as for preceding; +CASENT_9016090 +; +CAS • - + 1 ♂ -; same data as for preceding; CASENT_9016093; -CAS +; same data as for preceding; +CASENT_9016093 +; +CAS • - + 2 ♀♀ -; Parc National -Ranomafana +; +Parc National Ranomafana , Vatoharanana ; @@ -867,14 +951,20 @@ a.s.l. J. Schweikert , D. Ubick -leg.; primary forest; CEG-MA-1998-03; CASENT_9016080; -CAS +leg.; +primary forest +; +CEG-MA-1998-03 +; +CASENT_9016080 +; +CAS • - + 1 ♀ -; Parc National -Ranomafana +; +Parc National Ranomafana , Vatoharanana , trail K; @@ -896,20 +986,24 @@ and K.J. Ribardo leg.; pitfall trap -; VFL-KJR-02; CASENT_9016091; -CAS +; VFL-KJR-02; +CASENT_9016091 +; +CAS • - + 1 ♀ -; same data as for preceding; CASENT_9016092; -CAS +; same data as for preceding; +CASENT_9016092 +; +CAS • - + 1 ♀ -; Parc National -Ranomafana +; +Parc National Ranomafana , Vatoharanana River , @@ -932,17 +1026,23 @@ a.s.l. ; Fisher-Griswold Arthropod Team -leg.; montane rainforest; general day collecting; BLF8394; CASENT_9017837; -CAS +leg.; +montane rainforest +; +general day collecting +; BLF8394; +CASENT_9017837 +; +CAS • - + 1 ♀ -; Parc National -Ranomafana +; +Parc National Ranomafana , -Vohiparara -Piste Touristique; +Vohiparara Piste Touristique +; 21°14′ S , 47°24′ E @@ -971,14 +1071,20 @@ a.s.l. J. Schweikert , D. Ubick -leg.; running in litter; general collecting; CEG-MA-1998; CASENT_9016081; -CAS +leg.; +running in litter +; +general collecting +; CEG-MA-1998; +CASENT_9016081 +; +CAS • - + 6 ♂♂ ; -Ranomafana National Park +Ranomafana National Park , Vohiparara ; @@ -1002,14 +1108,18 @@ a.s.l. E. Elsom , F. Vargas -leg.; mixed tropical forest; general collecting; COL- DHK-2001-004; CASENT_9003479; -CAS +leg.; +mixed tropical forest +; general collecting; COL- DHK-2001-004; +CASENT_9003479 +; +CAS • - + 6 ♀♀ -; Parc National -Ranomafana +; +Parc National Ranomafana , 2.3 km @@ -1044,23 +1154,27 @@ a.s.l. J. Schweikert , D. Ubick -leg.; CASENT_9016083; -CAS +leg.; +CASENT_9016083 +; +CAS • - + 6♀♀ -; same data as for preceding; CASENT_9016085; -CAS +; same data as for preceding; +CASENT_9016085 +; +CAS • - + 2 ♀♀ ; Ranomafana N.P., -Trail -F; +Trail F +; 21°12′ S , 47°27′ E @@ -1074,17 +1188,23 @@ for Kariko , Roth -leg.; sifting #1; CASENT_9030889; -MCZ +leg.; +sifting +#1; +CASENT_9030889 +; +MCZ • - + 1 ♀ ; Ranomafana N.P., Maharira -, summit; +, +summit +; 21°12′ S , 47°27′ E @@ -1098,17 +1218,19 @@ for Kariko , Roth -leg.; cryptic #2; CASENT_9030890; -MCZ +leg.; cryptic #2; +CASENT_9030890 +; +MCZ • - + 1 ♂ ; Ranomafana N.P., -Trail -M; +Trail M +; 21°12′ S , 47°27′ E @@ -1122,17 +1244,21 @@ for Kariko , Roth -leg.; sifting 1hr. samples #2; CASENT_9030891; -MCZ +leg.; +sifting 1hr +. samples #2; +CASENT_9030891 +; +MCZ • - + 1 ♀ ; Ranomafana N.P., -Trail -M; +Trail M +; 21°12′ S , 47°27′ E @@ -1146,17 +1272,21 @@ for Kariko , Roth -leg.; sifting; CASENT_9030892; -MCZ +leg.; +sifting +; +CASENT_9030892 +; +MCZ • - + 3 ♀♀ ; Ranomafana N.P., -Trail -B; +Trail B +; 21°12′ S , 47°27′ E @@ -1175,17 +1305,23 @@ for Kariko , Roth -leg.; sifting, #1; CASENT_9030893; -MCZ +leg.; +sifting +, #1; +CASENT_9030893 +; +MCZ • - + 1 ♀ ; Ranomafana N.P., Maharira -, summit; +, +summit +; 21°12′ S , 47°27′ E @@ -1199,11 +1335,15 @@ for Kariko , Roth -leg.; sifting #3; CASENT_9030894; -MCZ +leg.; +sifting +#3; +CASENT_9030894 +; +MCZ • - + 2 ♀♀ ; Ranomafana @@ -1219,17 +1359,27 @@ N.P.; V. and B. Roth , S. Kariko -leg.; forest; general collecting from foliage; Ranomafana-1000-92; CASENT_9016099; -CAS +leg.; +forest +; +general collecting from foliage +; Ranomafana-1000-92; +CASENT_9016099 +; +CAS • - + 1 ♂ , 1 ♀ ; Ranomafana -N.P., trail F, secondary forest; +N.P., +trail F +, +secondary forest +; 21°12′ S , 47°27′ E @@ -1243,11 +1393,15 @@ for Kariko , Roth -leg.; sifting #3; MCZ-VBR-0116; CASENT_9016100; -MCZ +leg.; +sifting +#3; MCZ-VBR-0116; +CASENT_9016100 +; +MCZ • - + 4 ♂♂ , 7 ♀♀ @@ -1265,14 +1419,18 @@ N.P.; V. and B. Roth leg.; can traps -; MCZ-VBR-02; CASENT_9016101; -MCZ +; MCZ-VBR-02; +CASENT_9016101 +; +MCZ • - + 1 ♂ -; same data as for preceding; CASENT_9016102; -CAS +; same data as for preceding; +CASENT_9016102 +; +CAS . @@ -1376,7 +1534,7 @@ SEM of female genitalia, A–B, E–F = female paratype (CASENT_9016094; CAS); C . Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: A–E = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila. - + Description @@ -1687,6 +1845,8 @@ Wheeler (CASENT_9024004). + + Natural history