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<document id="04C93FB4B1A05D5D34501C3D831F9E5B" ID-CLB-Dataset="46773" ID-DOI="10.11646/zootaxa.4078.1.6" ID-GBIF-Dataset="598cb955-c4b7-4c2b-8c74-6ee0348c7f7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="270853" ID-ZooBank="3D0D79C2-2583-47F4-BC01-5CF347614B44" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460362976381" checkinUser="plazi" docAuthor="Pinto, Ângelo Parise &amp; Almeida, Marcus Vinícius Oliveira De" docDate="2016" docId="495E274CFFC4FF97FF43FAD0ED4AFCCC" docLanguage="en" docName="zootaxa.4078.1.6.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyanogomphus comparabilis Belle 1994" docType="treatment" docVersion="8" masterDocId="B5675F34FFC9FF87FFD4FFFCE97EFD39" masterDocTitle="A taxonomic synopsis of South American Cyanogomphini Carle with description of Cyanogomphus angelomachadoi sp. nov. from the Cerrado of Brazil (Odonata: Gomphidae)" updateTime="1698294756334" updateUser="ExternalLinkService">
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<mods:title id="2EE8B7F7BD001A2D08D046EA2D8FEF63">A taxonomic synopsis of South American Cyanogomphini Carle with description of Cyanogomphus angelomachadoi sp. nov. from the Cerrado of Brazil (Odonata: Gomphidae)</mods:title>
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<mods:namePart id="20C98FED812AE1301EAD4EDABAF08382">Pinto, Ângelo Parise</mods:namePart>
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<mods:namePart id="2DFBF1C2D346BF5A8B70EFA233353CC4">Almeida, Marcus Vinícius Oliveira De</mods:namePart>
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<mods:title id="032F775BBAEE3B33123487AE4C4C2E08">Zootaxa</mods:title>
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<mods:date id="9114F403CA748EC60F5C2D789D2BAA5A">2016</mods:date>
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<mods:number id="B5D9E328321B6006F2B80E12CC05050B">4078</mods:number>
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<mods:identifier id="62E29446399D00C9A66C8C791E598B1A" type="DOI">10.11646/zootaxa.4078.1.6</mods:identifier>
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<treatment id="495E274CFFC4FF97FF43FAD0ED4AFCCC" ID-DOI="http://doi.org/10.5281/zenodo.6077751" ID-GBIF-Taxon="119410164" ID-Zenodo-Dep="6077751" LSID="urn:lsid:plazi:treatment:495E274CFFC4FF97FF43FAD0ED4AFCCC" httpUri="http://treatment.plazi.org/id/495E274CFFC4FF97FF43FAD0ED4AFCCC" lastPageId="16" pageId="13">
<subSubSection id="89EDC5D1FFC4FF8AFF43FAD0EB02F8BE" pageId="13" type="nomenclature">
<paragraph id="C148965AFFC4FF8AFF43FAD0EBF8F87F" blockId="13.[151,991,1324,1415]" box="[151,646,1324,1350]" pageId="13">
<heading id="9A002136FFC4FF8AFF43FAD0EBF8F87F" bold="true" box="[151,646,1324,1350]" fontSize="11" level="1" pageId="13" reason="1">
<taxonomicName id="06F7EDD9FFC4FF8AFF43FAD0EBF8F87F" ID-CoL="32NML" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[151,646,1324,1350]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFC4FF8AFF43FAD0EBF8F87F" bold="true" box="[151,646,1324,1350]" pageId="13">
<emphasis id="F3834A48FFC4FF8AFF43FAD0E889F87F" bold="true" box="[151,503,1324,1350]" italics="true" pageId="13">Cyanogomphus comparabilis</emphasis>
<bibRefCitation id="A566EBABFFC4FF8AFE2AFAD0EBF8F87F" author="Belle" box="[510,646,1324,1350]" pageId="13" refString="Belle, J. (1994) Three new Neotropical Gomphidae from the genera Archaeogomphus Williamson, Cyanogomphus Selys and Epigomphus Hagen (Anisoptera). Odonatologica, 23 (1), 45 - 50." type="journal article" year="1994">Belle, 1994</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="C148965AFFC4FF8AFF43FAB2EB02F8BE" blockId="13.[151,991,1324,1415]" pageId="13">
LSID urn:lsid:zoobank.org:act:3C2DF075-F675-4A30-B8E1-A9628E7367DB (
<figureCitation id="59CC8ADFFFC4FF8AFF4AFA92E874F8BE" box="[158,266,1390,1415]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="13">Figures 2</figureCitation>
C, 4C, 5E, 7D–F, 8E–F, 10C, 12)
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFC4FF8AFF43FA4AE88BFB5F" pageId="13" type="reference_group">
<paragraph id="C148965AFFC4FF8AFF43FA4AE88BFB5F" blockId="13.[151,1437,1462,1638]" pageId="13">
<treatmentCitationGroup id="E1E7B174FFC4FF8AFF43FA4AE88BFB5F" infered="true" pageId="13">
<taxonomicName id="06F7EDD9FFC4FF8AFF43FA4AEBFFF8F5" ID-CoL="32NML" authority="Belle, 1994: 47" authorityName="Belle" authorityPageNumber="47" authorityYear="1994" box="[151,641,1462,1484]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFC4FF8AFF43FA4AE8B6F8F5" box="[151,456,1462,1484]" italics="true" pageId="13">Cyanogomphus comparabilis</emphasis>
<treatmentCitation id="4056B04BFFC4FF8AFE02FA4AEBFFF8F5" author="Belle" box="[470,641,1462,1484]" page="47" pageId="13" year="1994">
<bibRefCitation id="A566EBABFFC4FF8AFE02FA4AEB2BF8F5" author="Belle" box="[470,597,1462,1484]" pageId="13" refString="Belle, J. (1994) Three new Neotropical Gomphidae from the genera Archaeogomphus Williamson, Cyanogomphus Selys and Epigomphus Hagen (Anisoptera). Odonatologica, 23 (1), 45 - 50." type="journal article" year="1994">Belle, 1994</bibRefCitation>
: 47
</treatmentCitation>
</taxonomicName>
–49, figs 5–8 (description of ♂
<typeStatus id="1E4C28F8FFC4FF8AFC39FA4AED36F8F5" box="[1005,1096,1462,1484]" pageId="13" type="holotype">holotype</typeStatus>
,
<collectingCountry id="B9E0D6CAFFC4FF8AFB8EFA4BEDC7F8F5" box="[1114,1209,1463,1484]" name="Brazil" pageId="13">BRAZIL</collectingCountry>
. Mato Grosso State: Diamantino municipality,
<date id="B549B09AFFC4FF8AFE03FA29EB57F8D2" box="[471,553,1493,1515]" pageId="13" value="1988-04">IV.1988</date>
, E. Furtado leg. in ABMM, illustrations of color of mesepisternum in dorsal view, secondary genitalia in lateral view, caudal appendages in dorsal and lateral views of
<typeStatus id="1E4C28F8FFC4FF8AFBE5FA08EDF2FB33" box="[1073,1164,1524,1546]" pageId="13" type="holotype">holotype</typeStatus>
);—
<bibRefCitation id="A566EBABFFC4FF8AFB65FA08E803FB11" author="Bridges" pageId="13" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Odonata of the world (Third Edition). Privately published, Urbana, Illinois, 905 pp." type="book" year="1994">
Bridges (1994:
<date id="B549B09AFFC4FF8AFA87FA08ECE9FB33" box="[1363,1431,1524,1546]" pageId="13" value="1954-07">VII.54</date>
, VIII.18, mention)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFC4FF8AFE4FF9EEED1CFB11" author="Belle" box="[411,1122,1554,1576]" pageId="13" refString="Belle, J. (1996) Higher classification of the South-American Gomphidae (Odonata). Zoologische Mededelingen, 70, 297 - 324." type="journal article" year="1996">
Belle (1996: 298, comparison with
<taxonomicName id="06F7EDD9FFC4FF8AFCD0F9EFEA09FB11" ID-CoL="32NMT" box="[772,887,1554,1576]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC4FF8AFCD0F9EFEA09FB11" box="[772,887,1554,1576]" italics="true" pageId="13">C. waltheri</emphasis>
</taxonomicName>
and
<taxonomicName id="06F7EDD9FFC4FF8AFC7DF9EEED52FB11" ID-CoL="9WVZC" box="[937,1068,1554,1576]" class="Insecta" family="Gomphidae" genus="Ebegomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFC4FF8AFC7DF9EEED52FB11" box="[937,1068,1554,1576]" italics="true" pageId="13">Ebegomphus</emphasis>
</taxonomicName>
spp.)
</bibRefCitation>
;—
<treatmentCitation id="4056B04BFFC4FF8AFB56F9EEEC2BFB11" author="Costa" box="[1154,1365,1554,1576]" page="8" pageId="13" year="2000">
<bibRefCitation id="A566EBABFFC4FF8AFB56F9EEEC41FB11" author="Costa" box="[1154,1343,1554,1576]" pageId="13" refString="Costa, J. M., Machado, A. B. M., Lencioni, F. A. A. &amp; Santos, T. C. (2000) Diversidade e distribuicao dos Odonata (Insecta) no estado de Sao Paulo, Brasil: Parte I - Lista das especies e registros bibliograficos. Publicacoes Avulsas do Museu Nacional, 80, 3 - 28." type="journal article" year="2000">
Costa
<emphasis id="F3834A48FFC4FF8AFB12F9EFED84FB11" box="[1222,1274,1554,1576]" italics="true" pageId="13">et al.</emphasis>
(2000
</bibRefCitation>
: 8
</treatmentCitation>
–9, 14, mention);—
<treatmentCitation id="4056B04BFFC4FF8AFE95F9CDEB3DFB7E" author="Garrison" box="[321,579,1585,1607]" page="96" pageId="13" year="2006">
<bibRefCitation id="A566EBABFFC4FF8AFE95F9CDEB62FB7E" author="Garrison" box="[321,540,1585,1607]" pageId="13" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
Garrison
<emphasis id="F3834A48FFC4FF8AFE76F9CEE8A8FB7E" box="[418,470,1585,1607]" italics="true" pageId="13">et al.</emphasis>
(2006
</bibRefCitation>
: 96
</treatmentCitation>
, mention);—
<treatmentCitation id="4056B04BFFC4FF8AFD13F9CDEADAFB7E" author="Heckman" box="[711,932,1585,1607]" page="644" pageId="13" year="2006">
<bibRefCitation id="A566EBABFFC4FF8AFD13F9CDEA0EFB7E" author="Heckman" box="[711,880,1585,1607]" pageId="13" refString="Heckman, C. W. (2006) Encyclopedia of South American aquatic insects: Odonata Anisoptera. Illustrated keys to known families, genera, and species in South America. Springer, Dordrecht, viii + 725 pp." type="book" year="2006">Heckman (2006</bibRefCitation>
: 644
</treatmentCitation>
, fig. 3.2.771, key, reproduction of illustrations of ♂
<typeStatus id="1E4C28F8FFC4FF8AFF34F9ACE847FB5F" box="[224,313,1616,1638]" pageId="13" type="holotype">holotype</typeStatus>
from
<bibRefCitation id="A566EBABFFC4FF8AFEA2F9ACE897FB5F" author="Belle" box="[374,489,1616,1638]" pageId="13" refString="Belle, J. (1994) Three new Neotropical Gomphidae from the genera Archaeogomphus Williamson, Cyanogomphus Selys and Epigomphus Hagen (Anisoptera). Odonatologica, 23 (1), 45 - 50." type="journal article" year="1994">Belle 1994</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFC4FF8AFF43F96EEDFDFA02" pageId="13" type="materials_examined">
<paragraph id="C148965AFFC4FF8AFF43F96EEAB9FA2E" blockId="13.[151,1436,1682,2031]" pageId="13">
<emphasis id="F3834A48FFC4FF8AFF43F96EE8B6FB92" bold="true" box="[151,456,1682,1707]" pageId="13">
Material examined (
<specimenCount id="D7F15DD3FFC4FF8AFE41F96EE8C7FB92" box="[405,441,1682,1707]" pageId="13" type="generic">2♂</specimenCount>
).
</emphasis>
<collectingCountry id="B9E0D6CAFFC4FF8AFE01F96FEB40FB92" box="[469,574,1683,1707]" name="Brazil" pageId="13">BRAZIL</collectingCountry>
. M[a]t[o] Grosso State:
<emphasis id="F3834A48FFC4FF8AFCB7F96EEA8FFB92" bold="true" box="[867,1009,1682,1707]" pageId="13">
<typeStatus id="1E4C28F8FFC4FF8AFCB7F96EEAAEFB92" box="[867,976,1682,1707]" pageId="13" type="holotype">Holotype</typeStatus>
♂
</emphasis>
, Diamantino municipality, Alto Rio Arinós [
<geoCoordinate id="A4C3F09DFFC4FF8AFF21F94AE800FBF6" box="[245,382,1718,1743]" direction="south" orientation="latitude" pageId="13" precision="15" value="-14.16">14°09’36”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFC4FF8AFE59F94AEB63FBF6" box="[397,541,1718,1743]" direction="west" orientation="longitude" pageId="13" precision="15" value="-56.2">56°12’00”W</geoCoordinate>
,
<quantity id="060F3BBFFFC4FF8AFDF8F94BEB09FBF7" box="[556,631,1719,1743]" metricMagnitude="2" metricUnit="m" metricValue="3.37" pageId="13" unit="m" value="337.0">337 m</quantity>
a.s.l.], pond at Cerrado near gallery forest,
<date id="B549B09AFFC4FF8AFBA8F94BEDA9FBF6" box="[1148,1239,1719,1743]" pageId="13" value="1988-04">IV.1988</date>
, E. Furtado leg.; S[ão] P[aulo] State:
<emphasis id="F3834A48FFC4FF8AFEAFF926EB6FFBCA" bold="true" box="[379,529,1754,1779]" pageId="13">
<specimenCount id="D7F15DD3FFC4FF8AFEAFF926E8E0FBCA" box="[379,414,1754,1779]" pageId="13" type="generic">1♂</specimenCount>
<typeStatus id="1E4C28F8FFC4FF8AFE70F926EB6FFBCA" box="[420,529,1754,1779]" pageId="13" type="paratype">Paratype</typeStatus>
</emphasis>
, Igarapava municipality [
<geoCoordinate id="A4C3F09DFFC4FF8AFCE4F927EAA3FBCA" box="[816,989,1755,1779]" direction="south" orientation="latitude" pageId="13" precision="1" value="-20.0389">20°02’20.04”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFC4FF8AFC3CF927EDE5FBCA" box="[1000,1179,1755,1779]" direction="west" orientation="longitude" pageId="13" precision="1" value="-47.747">47°44’49.20”W</geoCoordinate>
,
<quantity id="060F3BBFFFC4FF8AFB73F926ED90FBCB" box="[1191,1262,1754,1779]" metricMagnitude="2" metricUnit="m" metricValue="5.779999999999999" pageId="13" unit="m" value="578.0">578 m</quantity>
a.s.l.], Córrego Canabrava, [0]
<date id="B549B09AFFC4FF8AFEE8F902E814FA2E" box="[316,362,1790,1815]" pageId="13">6.II.</date>
[19]89, P.A. Machado &amp; A. Costa leg. All in
<collectionCode id="A7E60E9FFFC4FF8AFCB3F903EABDFA2E" box="[871,963,1791,1815]" pageId="13">ABMM</collectionCode>
.
</paragraph>
<paragraph id="C148965AFFC4FF8AFF13F8DEEDFDFA02" blockId="13.[151,1436,1682,2031]" box="[199,1155,1826,1851]" pageId="13">
<emphasis id="F3834A48FFC4FF8AFF13F8DEE8F7FA02" bold="true" box="[199,393,1826,1851]" pageId="13">
<typeStatus id="1E4C28F8FFC4FF8AFF13F8DEE87CFA02" box="[199,258,1826,1851]" pageId="13">Type</typeStatus>
repository.
</emphasis>
<typeStatus id="1E4C28F8FFC4FF8AFE5BF8DEE889FA02" box="[399,503,1826,1851]" pageId="13" type="holotype">Holotype</typeStatus>
♂ and one
<typeStatus id="1E4C28F8FFC4FF8AFDADF8DFEBA7FA02" box="[633,729,1827,1851]" pageId="13" type="paratype">paratype</typeStatus>
♂ by original designation in
<collectionCode id="A7E60E9FFFC4FF8AFBF6F8DFED00FA02" box="[1058,1150,1827,1851]" pageId="13">ABMM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFC4FF89FF13F8BAEB58F8CE" lastPageId="14" pageId="13" type="description">
<paragraph id="C148965AFFC4FF8AFF13F8BAE896FAD6" blockId="13.[151,1436,1682,2031]" pageId="13">
<emphasis id="F3834A48FFC4FF8AFF13F8BAE803FA66" bold="true" box="[199,381,1862,1887]" pageId="13">Measurements.</emphasis>
Males (n = 2). Total length (including caudal appendages) 42.7–47.0; abdomen length (excluding caudal appendages) 31.8–34.5; head maximum width 5.6–5.9; Fw length 24.9–25.8; Hw length 23.8–24.8; Fw maximum width 5.3–5.7, in Hw 6.4–7.1; pt length
<quantity id="060F3BBFFFC4FF8AFC63F873ED50FA9E" box="[951,1070,1934,1959]" metricMagnitude="-2" metricUnit="m" metricValue="8.255" metricValueMax="8.636" metricValueMin="7.874" pageId="13" unit="in" value="3.25" valueMax="3.4" valueMin="3.1">3.1–3.4 in</quantity>
Fw,
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Hw; length of metathoracic femur 6.1–7.2; metathoracic tibia 4.3–5.8; length of S9+
<quantity id="060F3BBFFFC4FF8AFC6AF84FEA84FAF2" box="[958,1018,1970,1995]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="13" unit="in" value="10.0">10 in</quantity>
lateral view 3.1–3.8; total length of cercus in lateral view 1.0–1.3.
</paragraph>
<caption id="9588C6D2FFC7FF89FF43FAA7EC76F88D" httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="14" targetBox="[155,1434,153,1337]" targetPageId="14">
<paragraph id="C148965AFFC7FF89FF43FAA7EC76F88D" blockId="14.[151,1437,1371,1460]" pageId="14">
<emphasis id="F3834A48FFC7FF89FF43FAA7E873F849" bold="true" box="[151,269,1371,1393]" pageId="14">FIGURE 6</emphasis>
. Male vesica spermalis (VS) of Cyanogomphini. A–C.
<taxonomicName id="06F7EDD9FFC7FF89FC94FAA7EDF3F848" box="[832,1165,1371,1393]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC7FF89FC94FAA7EDF3F848" box="[832,1165,1371,1393]" italics="true" pageId="14">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC7FF89FB47FAA7ED9DF849" bold="true" box="[1171,1251,1371,1392]" pageId="14">
<taxonomicNameLabel id="E8B0F733FFC7FF89FB47FAA7ED9DF849" box="[1171,1251,1371,1392]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Paratype (Brazil. MG: Lagoa Santa, MNRJ); D–E.
<taxonomicName id="06F7EDD9FFC7FF89FE3DFA81EA22F8AA" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" box="[489,860,1405,1427]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC7FF89FE3DFA81EB9EF8AA" box="[489,736,1405,1427]" italics="true" pageId="14">Cyanogomphus waltheri</emphasis>
Selys, 1873
</taxonomicName>
(Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F–G.
<taxonomicName id="06F7EDD9FFC7FF89FEC2FA62EA7EF88D" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[278,768,1438,1460]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFC7FF89FEC2FA62E89AF88D" box="[278,484,1438,1460]" italics="true" pageId="14">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</taxonomicName>
(Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="C148965AFFC7FF89FF13FA22EB58F8CE" blockId="14.[151,1436,1502,2031]" box="[199,550,1502,1527]" pageId="14">
<emphasis id="F3834A48FFC7FF89FF13FA22E8D7F8CE" bold="true" box="[199,425,1502,1527]" pageId="14">Female and Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFC7FF89FF13F9FEEC06FAD6" pageId="14" type="diagnosis">
<paragraph id="C148965AFFC7FF89FF13F9FEEB4DFBBE" blockId="14.[151,1436,1502,2031]" pageId="14">
<emphasis id="F3834A48FFC7FF89FF13F9FEE83FFB22" bold="true" box="[199,321,1538,1563]" pageId="14">Diagnosis.</emphasis>
Males of
<taxonomicName id="06F7EDD9FFC7FF89FE60F9FEEB80FB22" box="[436,766,1538,1563]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFC7FF89FE60F9FEEB80FB22" box="[436,766,1538,1563]" italics="true" pageId="14">Cyanogomphus comparabilis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="06F7EDD9FFC7FF89FBE6F9FEEDAFFB22" box="[1074,1233,1538,1563]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFC7FF89FBE6F9FEEDAFFB22" box="[1074,1233,1538,1563]" italics="true" pageId="14">Tibiagomphus</emphasis>
</taxonomicName>
spp. by its cercus cylindrical, slightly curved toward to the upturned tapered apex (
<figureCitation id="59CC8ADFFFC7FF89FCB8F9DAEACCFB06" box="[876,946,1574,1599]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="14">Figs 7</figureCitation>
A–I, 8A–F) and branches of epiproct with a distal concavity at posterodorsal apex (
<figureCitation id="59CC8ADFFFC7FF89FD9FF9B6EBECFB5A" box="[587,658,1610,1635]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="14">Figs 8</figureCitation>
B, D, F, apex of cercus directed posteriorly, epiproct dorsally smooth, without concavity in
<taxonomicName id="06F7EDD9FFC7FF89FE50F992EB5DFBBE" box="[388,547,1646,1671]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFC7FF89FE50F992EB5DFBBE" box="[388,547,1646,1671]" italics="true" pageId="14">Tibiagomphus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="C148965AFFC7FF89FF13F96EEC06FAD6" blockId="14.[151,1436,1502,2031]" pageId="14">
This distinctive species is distinguished from congeners by the largely pale mesepisternum close to the dorsal carina (dark in
<taxonomicName id="06F7EDD9FFC7FF89FE9DF94BEB59FBF6" box="[329,551,1718,1743]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC7FF89FE9DF94BEB59FBF6" box="[329,551,1718,1743]" italics="true" pageId="14">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC7FF89FDE4F94AEBF3FBF6" bold="true" box="[560,653,1718,1743]" pageId="14">
<taxonomicNameLabel id="E8B0F733FFC7FF89FDE4F94AEBF3FBF6" box="[560,653,1718,1743]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="06F7EDD9FFC7FF89FD1CF94BEA36FBF6" box="[712,840,1718,1743]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC7FF89FD1CF94BEA36FBF6" box="[712,840,1718,1743]" italics="true" pageId="14">C. waltheri</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFC7FF89FC8DF94AEADFFBF6" box="[857,929,1718,1743]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="14">Figs 1</figureCitation>
,
<figureCitation id="59CC8ADFFFC7FF89FC66F94BEAB3FBF6" box="[946,973,1719,1743]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="14">10</figureCitation>
A–B, D, 11A); light brownish-yellow pt (
<figureCitation id="59CC8ADFFFC7FF89FF4BF926E994FBCA" box="[159,234,1754,1779]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="14">Figs 2</figureCitation>
C, 10C, darker, brown to dark brown in
<taxonomicName id="06F7EDD9FFC7FF89FD12F927EADBFBCA" box="[710,933,1754,1779]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC7FF89FD12F927EADBFBCA" box="[710,933,1754,1779]" italics="true" pageId="14">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC7FF89FC7BF926ED73FBCA" bold="true" box="[943,1037,1754,1779]" pageId="14">
<taxonomicNameLabel id="E8B0F733FFC7FF89FC7BF926ED73FBCA" box="[943,1037,1754,1779]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="06F7EDD9FFC7FF89FB9EF927EDB5FBCA" box="[1098,1227,1754,1779]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC7FF89FB9EF927EDB5FBCA" box="[1098,1227,1754,1779]" italics="true" pageId="14">C. waltheri</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFC7FF89FB08F926EC5BFBCA" box="[1244,1317,1754,1779]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="14">Figs 2</figureCitation>
A–B, D); almost uniformly colored abdomen, lacking distinct spots (
<figureCitation id="59CC8ADFFFC7FF89FC9CF902EAE3FA2E" box="[840,925,1790,1815]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="14">Fig. 10</figureCitation>
C; patterned with dark and pale spots in
<taxonomicName id="06F7EDD9FFC7FF89FA56F903E82CFA02" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC7FF89FA56F903E82CFA02" italics="true" pageId="14">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC7FF89FE8EF8DEE8C8FA02" bold="true" box="[346,438,1826,1851]" pageId="14">
<taxonomicNameLabel id="E8B0F733FFC7FF89FE8EF8DEE8C8FA02" box="[346,438,1826,1851]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="06F7EDD9FFC7FF89FE3BF8DFEB11FA02" box="[495,623,1826,1851]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC7FF89FE3BF8DFEB11FA02" box="[495,623,1826,1851]" italics="true" pageId="14">C. waltheri</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFC7FF89FDABF8DEEBADFA02" box="[639,723,1826,1851]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="14">Figs 10</figureCitation>
A–B, D, 11A); sharply pointed apex of posterior hamule (
<figureCitation id="59CC8ADFFFC7FF89FABEF8DEE9D8FA66" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="14">Figs 4</figureCitation>
C, distinctly less sharp in
<taxonomicName id="06F7EDD9FFC7FF89FE09F8BBEBC3FA66" box="[477,701,1862,1887]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC7FF89FE09F8BBEBC3FA66" box="[477,701,1862,1887]" italics="true" pageId="14">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC7FF89FD1CF8BAEA58FA66" bold="true" box="[712,806,1862,1887]" pageId="14">
<taxonomicNameLabel id="E8B0F733FFC7FF89FD1CF8BAEA58FA66" box="[712,806,1862,1887]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="06F7EDD9FFC7FF89FCB2F8BBEA96FA66" box="[870,1000,1862,1887]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC7FF89FCB2F8BBEA96FA66" box="[870,1000,1862,1887]" italics="true" pageId="14">C. waltheri</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFC7FF89FC2FF8BAED38FA66" box="[1019,1094,1862,1887]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="14">Figs 4</figureCitation>
B–D); a constriction at about anterior 0.66 to the enlarged apex of epiproct in lateral view (
<figureCitation id="59CC8ADFFFC7FF89FC86F896EAE4FABB" box="[850,922,1898,1923]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="14">Figs 7</figureCitation>
D, 8E–F, not markedly constricted anterior to the enlarged apex in
<taxonomicName id="06F7EDD9FFC7FF89FE56F873EB21FA9E" box="[386,607,1934,1959]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC7FF89FE56F873EB21FA9E" box="[386,607,1934,1959]" italics="true" pageId="14">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC7FF89FDB2F872EBBEFA9E" bold="true" box="[614,704,1934,1959]" pageId="14">
<taxonomicNameLabel id="E8B0F733FFC7FF89FDB2F872EBBEFA9E" box="[614,704,1934,1959]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="06F7EDD9FFC7FF89FD2CF873EA08FA9E" box="[760,886,1934,1959]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC7FF89FD2CF873EA08FA9E" box="[760,886,1934,1959]" italics="true" pageId="14">C. waltheri</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFC7FF89FC51F872EAB5FA9E" box="[901,971,1934,1959]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="14">Figs 7</figureCitation>
A, G, 8A, D) and by the very small distal concavity on the epiproct, almost restricted to the apex (
<figureCitation id="59CC8ADFFFC7FF89FC9EF84EEAE8FAF3" box="[842,918,1970,1995]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="14">Figs 7</figureCitation>
D–F, 8E–F, distal concavity larger, roughly quadrangular, and extending more proximal in
<taxonomicName id="06F7EDD9FFC7FF89FD71F82BEAFFFAD6" box="[677,897,2006,2031]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC7FF89FD71F82BEAFFFAD6" box="[677,897,2006,2031]" italics="true" pageId="14">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC7FF89FC5CF82AEA9FFAD6" bold="true" box="[904,993,2006,2031]" pageId="14">
<taxonomicNameLabel id="E8B0F733FFC7FF89FC5CF82AEA9FFAD6" box="[904,993,2006,2031]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="06F7EDD9FFC7FF89FBCCF82BEDE8FAD6" box="[1048,1174,2006,2031]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="14" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC7FF89FBCCF82BEDE8FAD6" box="[1048,1174,2006,2031]" italics="true" pageId="14">C. waltheri</emphasis>
</taxonomicName>
<figureCitation id="59CC8ADFFFC7FF89FB49F82AED9CFAD6" box="[1181,1250,2006,2031]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="14">Figs 7</figureCitation>
A, G, 8A–D).
</paragraph>
</subSubSection>
<caption id="9588C6D2FFC6FF88FF43F914E8CCFAAE" httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="15" targetBox="[151,1436,197,1742]" targetPageId="15">
<paragraph id="C148965AFFC6FF88FF43F914E8CCFAAE" blockId="15.[151,1436,1768,1943]" pageId="15">
<emphasis id="F3834A48FFC6FF88FF43F914E873FBC4" bold="true" box="[151,269,1768,1790]" pageId="15">FIGURE 7</emphasis>
. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A–C.
<taxonomicName id="06F7EDD9FFC6FF88FE8DF8FAEBD8FA25" box="[345,678,1798,1820]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="15" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC6FF88FE8DF8FAEBD8FA25" box="[345,678,1798,1820]" italics="true" pageId="15">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC6FF88FD78F8FBEB83FA25" bold="true" box="[684,765,1799,1820]" pageId="15">
<taxonomicNameLabel id="E8B0F733FFC6FF88FD78F8FBEB83FA25" box="[684,765,1799,1820]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D–F.
<taxonomicName id="06F7EDD9FFC6FF88FECBF8D9EBAFFA02" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[287,721,1829,1851]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="15" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFC6FF88FECBF8D9EB32FA02" box="[287,588,1829,1851]" italics="true" pageId="15">Cyanogomphus comparabilis</emphasis>
Belle, 1994
</taxonomicName>
, Holotype (Brazil. MT: Diamantino, ABMM); G–I.
<taxonomicName id="06F7EDD9FFC6FF88FB2AF8D9E813FA63" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="15" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC6FF88FB2AF8D9E995FA63" italics="true" pageId="15">Cyanogomphus waltheri</emphasis>
Selys, 1873
</taxonomicName>
(Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J–L.
<taxonomicName id="06F7EDD9FFC6FF88FB86F8B8E844FA40" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="15" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFC6FF88FB86F8B8EC5CFA63" box="[1106,1314,1860,1882]" italics="true" pageId="15">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</taxonomicName>
(Brazil. RS: Jaguarão, MNRJ); M–O.
<taxonomicName id="06F7EDD9FFC6FF88FD0BF89FED19FA40" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[735,1127,1891,1913]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="15" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFC6FF88FD0BF89FEAB7FA40" box="[735,969,1891,1913]" italics="true" pageId="15">Tibiagomphus uncatus</emphasis>
(Fraser, 1947)
</taxonomicName>
(Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="89EDC5D1FFD9FF97FF13FF6BEAB4FDC1" pageId="16" type="distribution">
<paragraph id="C148965AFFD9FF97FF13FF6BEAB4FDC1" blockId="16.[151,1436,151,501]" pageId="16">
<emphasis id="F3834A48FFD9FF97FF13FF6BE81EFD89" bold="true" box="[199,352,151,176]" pageId="16">Distribution.</emphasis>
Mato Grosso and São Paulo States in
<collectingCountry id="B9E0D6CAFFD9FF97FCD9FF6BEA2AFD89" box="[781,852,151,176]" name="Brazil" pageId="16">Brazil</collectingCountry>
, in the Cerrado biogeographical province (
<figureCitation id="59CC8ADFFFD9FF97FAE7FF6BECF9FD89" box="[1331,1415,151,176]" captionStart="FIGURE 12" captionStartId="24.[151,250,1240,1262]" captionTargetBox="[151,1436,215,1198]" captionTargetId="figure@24.[151,1436,195,1215]" captionTargetPageId="24" captionText="FIGURE 12. Map of southeastern South America with distribution records of Cyanogomphus Selys, 1873, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: MI, Misiones Province, Argentina; BA, Bahia; GO, Goiás; MG, Minas Gerais; MT, Mato Grosso; RJ, Rio de Janeiro; RS, Rio Grande do Sul; SC, Santa Catarina, and SP, São Paulo for Brazilian States." httpUri="https://zenodo.org/record/270865/files/figure.png" pageId="16">Fig. 12</figureCitation>
).
<emphasis id="F3834A48FFD9FF97FF13FF40EB4BFDEC" bold="true" box="[199,565,188,213]" pageId="16">Biological and ecological data.</emphasis>
Known only from two males collected close to streams or brooklets in lotic systems surrounded by forests (riparian), at 337–578 meters of elevation.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFD9FF97FF13FEF8ED4AFCCC" pageId="16" type="discussion">
<paragraph id="C148965AFFD9FF97FF13FEF8ED4AFCCC" blockId="16.[151,1436,151,501]" pageId="16">
<emphasis id="F3834A48FFD9FF97FF13FEF8E845FC24" bold="true" box="[199,315,260,285]" pageId="16">Remarks.</emphasis>
<bibRefCitation id="A566EBABFFD9FF97FE97FEF8EB66FC25" author="Belle" box="[323,536,260,285]" pageId="16" refString="Belle, J. (1994) Three new Neotropical Gomphidae from the genera Archaeogomphus Williamson, Cyanogomphus Selys and Epigomphus Hagen (Anisoptera). Odonatologica, 23 (1), 45 - 50." type="journal article" year="1994" yearSuffix="p">Belle (1994, p. 47)</bibRefCitation>
and
<bibRefCitation id="A566EBABFFD9FF97FD84FEF8EA62FC25" author="Costa" box="[592,796,260,285]" pageId="16" refString="Costa, J. M., Machado, A. B. M., Lencioni, F. A. A. &amp; Santos, T. C. (2000) Diversidade e distribuicao dos Odonata (Insecta) no estado de Sao Paulo, Brasil: Parte I - Lista das especies e registros bibliograficos. Publicacoes Avulsas do Museu Nacional, 80, 3 - 28." type="journal article" year="2000">
Costa
<emphasis id="F3834A48FFD9FF97FD43FEF9EBAEFC24" box="[663,720,260,285]" italics="true" pageId="16">et al.</emphasis>
(2000
</bibRefCitation>
, p. 8) have stated that the male
<typeStatus id="1E4C28F8FFD9FF97FB50FEF9ED9AFC24" box="[1156,1252,261,285]" pageId="16" type="paratype">paratype</typeStatus>
from São Paulo was deposited in Naturalis Biodiversity Center, Leiden,
<collectingCountry id="B9E0D6CAFFD9FF97FCC6FEDBEAB3FC79" box="[786,973,295,320]" name="Netherlands" pageId="16">The Netherlands</collectingCountry>
(RMNH). However, it was never sent to that collection and is still deposited in ABMM, whose collection was donated to and will be transferred to the Universidade Federal de Minas Gerais, Belo Horizonte,
<collectingCountry id="B9E0D6CAFFD9FF97FCEFFE93EA01FCB1" box="[827,895,367,392]" name="Brazil" pageId="16">Brazil</collectingCountry>
(UFMG). We suspect that the illustration of secondary genitalia by
<bibRefCitation id="A566EBABFFD9FF97FE4FFE68EB0FFC94" author="Belle" box="[411,625,404,429]" pageId="16" refString="Belle, J. (1994) Three new Neotropical Gomphidae from the genera Archaeogomphus Williamson, Cyanogomphus Selys and Epigomphus Hagen (Anisoptera). Odonatologica, 23 (1), 45 - 50." type="journal article" year="1994">Belle (1994, fig. 6)</bibRefCitation>
was based on the
<typeStatus id="1E4C28F8FFD9FF97FCEBFE69EADCFC94" box="[831,930,405,429]" pageId="16" type="paratype">paratype</typeStatus>
, as it differs from the
<typeStatus id="1E4C28F8FFD9FF97FB4EFE68ED82FC94" box="[1178,1276,404,429]" pageId="16" type="holotype">holotype</typeStatus>
that shows an unusual shape (
<figureCitation id="59CC8ADFFFD9FF97FE9CFE4BE8F0FCE9" box="[328,398,439,464]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="16">Fig. 4</figureCitation>
C). We illustrate the posterior hamule of the
<typeStatus id="1E4C28F8FFD9FF97FC5BFE44EA91FCE9" box="[911,1007,440,464]" pageId="16" type="paratype">paratype</typeStatus>
(
<figureCitation id="59CC8ADFFFD9FF97FBD5FE4BED3AFCE9" box="[1025,1092,439,464]" captionStart="FIGURE 5" captionStartId="11.[151,250,1535,1557]" captionTargetBox="[151,1436,195,1507]" captionTargetId="figure@11.[151,1436,195,1511]" captionTargetPageId="11" captionText="FIGURE 5. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A – D) and posterior hamule in lateral view (E – L). A – B, F – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); C – D, I – L. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). E. Cyanogomphus comparabilis Belle, 1994, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="16">Fig. 5</figureCitation>
E) which resembles that of
<taxonomicName id="06F7EDD9FFD9FF97FA56FE44E98AFCCC" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFD9FF97FA56FE44E98AFCCC" italics="true" pageId="16">C. waltheri</emphasis>
</taxonomicName>
, showing a more usual shape, similar to the other
<taxonomicName id="06F7EDD9FFD9FF97FCF3FE20EAABFCCC" box="[807,981,476,501]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFD9FF97FCF3FE20EAABFCCC" box="[807,981,476,501]" italics="true" pageId="16">Cyanogomphus</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="495E274CFFCEFF8AFF43FB81EAB8F9D1" ID-DOI="http://doi.org/10.5281/zenodo.6077749" ID-GBIF-Taxon="119410156" ID-Zenodo-Dep="6077749" LSID="urn:lsid:zoobank.org:act:61BDDFBA-202E-41CD-AA6D-2E65CD171CF4" httpUri="http://treatment.plazi.org/id/495E274CFFCEFF8AFF43FB81EAB8F9D1" lastPageId="13" pageId="7">
<subSubSection id="89EDC5D1FFCEFF80FF43FB81E810F805" pageId="7" type="nomenclature">
<paragraph id="C148965AFFCEFF80FF43FB81EBF6F9AE" blockId="7.[151,1024,1149,1240]" box="[151,648,1149,1175]" pageId="7">
<heading id="9A002136FFCEFF80FF43FB81EBF6F9AE" bold="true" box="[151,648,1149,1175]" fontSize="11" level="1" pageId="7" reason="1">
<emphasis id="F3834A48FFCEFF80FF43FB81EBF6F9AE" bold="true" box="[151,648,1149,1175]" pageId="7">
<taxonomicName id="06F7EDD9FFCEFF80FF43FB81EB5DF9AE" ID-CoL="9WYQQ" box="[151,547,1149,1175]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="7" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFCEFF80FF43FB81EB5DF9AE" bold="true" box="[151,547,1149,1175]" italics="true" pageId="7">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8B0F733FFCEFF80FDFFFB81EBF6F9AE" box="[555,648,1149,1175]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C148965AFFCEFF80FF43FB63EA93F9E1" blockId="7.[151,1024,1149,1240]" pageId="7">
LSID urn:lsid:zoobank.org:act:61BDDFBA-202E-41CD-AA6D-2E65CD171CF4 (
<figureCitation id="59CC8ADFFFCEFF80FF4AFB43E877F9E1" box="[158,265,1215,1240]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="7">Figures 1</figureCitation>
,
<figureCitation id="59CC8ADFFFCEFF80FEC3FB3CE85BF9E1" box="[279,293,1216,1240]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="7">2</figureCitation>
A–B, 3A, 4A–B, 6A–C, 7A–C, 8A–B, 9C, 10A–B, 11A–B, 12)
</paragraph>
<paragraph id="C148965AFFCEFF80FF43FAFBE810F805" blockId="7.[151,1435,1287,1340]" pageId="7">
<treatmentCitationGroup id="E1E7B174FFCEFF80FF43FAFBE810F805" pageId="7">
<taxonomicName id="06F7EDD9FFCEFF80FF43FAFBE84BF824" ID-CoL="3XNW" box="[151,309,1287,1309]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="7" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="F3834A48FFCEFF80FF43FAFBE84BF824" box="[151,309,1287,1309]" italics="true" pageId="7">Cyanogomphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8B0F733FFCEFF80FEE9FAF4E8F2F824" box="[317,396,1288,1309]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
—
<treatmentCitation id="4056B04BFFCEFF80FE72FAF4EB9CF824" author="" box="[422,738,1287,1310]" page="418" pageId="7">
<bibRefCitation id="A566EBABFFCEFF80FE72FAF4EBCEF824" box="[422,688,1287,1310]" pageId="7" refString="De Almeida, M. V. O., Pinto, A. P., Carvalho, A. L. &amp; Takiya, D. M. (2013) When rare is just a matter of sampling: Unexpected dominance of clubtail dragonflies (Odonata, Gomphidae) through different collecting methods at Parque Nacional da Serra do Cipo, Minas Gerais State, Brazil. Revista Brasileira de Entomologia, 57 (4), 417 - 423. http: // dx. doi. org / 10.1590 / S 0085 - 56262013005000042" type="journal article">
<collectingCountry id="B9E0D6CAFFCEFF80FE72FAF4E8BAF824" box="[422,452,1288,1309]" name="South Georgia and the South Sandwich Islands" pageId="7">De</collectingCountry>
Almeida
<emphasis id="F3834A48FFCEFF80FDF8FAF5EB1FF824" box="[556,609,1287,1310]" italics="true" pageId="7">et al.</emphasis>
(2013)
</bibRefCitation>
: 418
</treatmentCitation>
(mention of the material described here from “Parque Nacional da Serra do Cipó”).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFCEFF80FF43FA95EC19FA37" pageId="7" type="materials_examined">
<paragraph id="C148965AFFCEFF80FF43FA95EC19FA37" blockId="7.[151,1437,1385,2022]" pageId="7">
<emphasis id="F3834A48FFCEFF80FF43FA95EBEDF8BB" bold="true" box="[151,659,1385,1410]" pageId="7">
Material examined (
<specimenCount id="D7F15DD3FFCEFF80FE46FA95E8CBF8BB" box="[402,437,1385,1410]" pageId="7" type="generic">4♂</specimenCount>
, 2♀).
<typeStatus id="1E4C28F8FFCEFF80FDD2FA95EB0DF8BB" box="[518,627,1385,1410]" pageId="7" type="holotype">Holotype</typeStatus>
♂
</emphasis>
.
<collectingCountry id="B9E0D6CAFFCEFF80FD72FA95EA71F8B8" box="[678,783,1385,1409]" name="Brazil" pageId="7">BRAZIL</collectingCountry>
. Minas Gerais State: Jaboticatubas municipality, Parque Nacional da Serra do Cipó (
<collectionCode id="A7E60E9FFFCEFF80FE0AFA71EB57F89C" box="[478,553,1421,1445]" pageId="7">PNSC</collectionCode>
),, light trap using a white sheet with mixed mercury vapor lights (250W) at Córrego das Pedras stream (
<geoCoordinate id="A4C3F09DFFCEFF80FE0EFA4DEB1FF8F0" box="[474,609,1457,1481]" direction="south" orientation="latitude" pageId="7" precision="15" value="-19.37139">19°22’17”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFCEFF80FDBBFA4DEB80F8F3" box="[623,766,1457,1482]" direction="west" orientation="longitude" pageId="7" precision="15" value="-43.60083">43°36’03”W</geoCoordinate>
,
<quantity id="060F3BBFFFCEFF80FCD8FA4DEA2AF8F0" box="[780,852,1457,1482]" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" pageId="7" unit="m" value="766.0">766 m</quantity>
a.s.l.),
<date id="B549B09AFFCEFF80FC7FFA4DED4BF8F0" box="[939,1077,1457,1481]" pageId="7" value="2011-12-12">12.XII.2011</date>
, A.P.M. Santos &amp; D.M. Takiya leg. (
<collectionCode id="A7E60E9FFFCEFF80FF1BFA29E86AF8D4" box="[207,276,1493,1517]" pageId="7">DZRJ</collectionCode>
1449);
<emphasis id="F3834A48FFCEFF80FEBEFA29EB7FF8D7" bold="true" box="[362,513,1493,1518]" pageId="7">
1♀
<typeStatus id="1E4C28F8FFCEFF80FE40FA29EB7FF8D7" box="[404,513,1493,1518]" pageId="7" type="paratype">Paratype</typeStatus>
</emphasis>
, same data as
<typeStatus id="1E4C28F8FFCEFF80FD77FA29EA7BF8D7" box="[675,773,1493,1518]" pageId="7" type="holotype">holotype</typeStatus>
but 6 meter-long Malaise trap crossing Córrego das Pedras stream,
<date id="B549B09AFFCEFF80FF21FA05E8E2FB28" box="[245,412,1529,1553]" pageId="7" value="2011-12-09" valueMax="2011-12-13" valueMin="2011-12-09">9–13.XII.2011</date>
, A.P.M. Santos, D.M. Takiya, M.L. Monné &amp; R.R. Cavichioli leg. (
<collectionCode id="A7E60E9FFFCEFF80FB06FA05EC69FB28" box="[1234,1303,1529,1553]" pageId="7">DZRJ</collectionCode>
1448);
<emphasis id="F3834A48FFCEFF80FAADFA05E87AFB0F" bold="true" pageId="7">
<specimenCount id="D7F15DD3FFCEFF80FAADFA05ECE2FB2B" box="[1401,1436,1529,1554]" pageId="7" type="generic">1♂</specimenCount>
<typeStatus id="1E4C28F8FFCEFF80FF43F9E1E87AFB0F" box="[151,260,1565,1590]" pageId="7" type="paratype">Paratype</typeStatus>
</emphasis>
, [Lagoa Santa municipality], Estr[ada] B[elo] Horizonte, Serra do Cipó, Km 46–47 [
<geoCoordinate id="A4C3F09DFFCEFF80FB3FF9E1ECE9FB0F" box="[1259,1431,1565,1590]" direction="south" orientation="latitude" pageId="7" precision="1" value="-19.561266">19°33’40.56”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFCEFF80FF43F9BDE837FB63" box="[151,329,1601,1626]" direction="west" orientation="longitude" pageId="7" precision="1" value="-43.913116">43°54’47.21”W</geoCoordinate>
,
<quantity id="060F3BBFFFCEFF80FE81F9BDE8E0FB60" box="[341,414,1601,1626]" metricMagnitude="2" metricUnit="m" metricValue="6.64" pageId="7" unit="m" value="664.0">664 m</quantity>
a.s.l.], Exc[ursão] 221, Col[eta] 30,
<date id="B549B09AFFCEFF80FCE9F9BDEAB9FB63" box="[829,967,1601,1626]" pageId="7" value="1963-12-01">01.XII.1963</date>
, N.[D.] Santos, [J.P.] Machado &amp; Borges leg. (
<collectionCode id="A7E60E9FFFCEFF80FF1BF999E85EFB44" box="[207,288,1637,1661]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="7">MNRJ</collectionCode>
);
<emphasis id="F3834A48FFCEFF80FEE6F999E8B7FB47" bold="true" box="[306,457,1637,1662]" pageId="7">
<specimenCount id="D7F15DD3FFCEFF80FEE6F999E828FB47" box="[306,342,1637,1662]" pageId="7" type="generic">1♂</specimenCount>
<typeStatus id="1E4C28F8FFCEFF80FE88F999E8B7FB47" box="[348,457,1637,1662]" pageId="7" type="paratype">Paratype</typeStatus>
</emphasis>
, Serra do Salitre municipality, river at riparian Forest,
<date id="B549B09AFFCEFF80FBFFF999EDE7FB44" box="[1067,1177,1637,1661]" pageId="7" value="2014-01-13">13.I.2014</date>
, J.E. Santos Junior leg. (
<collectionCode id="A7E60E9FFFCEFF80FF4BF975E980FB98" box="[159,254,1673,1697]" pageId="7">ABMM</collectionCode>
); São Paulo State:
<emphasis id="F3834A48FFCEFF80FE1BF975EB18FB9B" bold="true" box="[463,614,1673,1698]" pageId="7">
<specimenCount id="D7F15DD3FFCEFF80FE1BF975E88CFB9B" box="[463,498,1673,1698]" pageId="7" type="generic">1♂</specimenCount>
<typeStatus id="1E4C28F8FFCEFF80FE2DF975EB18FB9B" box="[505,614,1673,1698]" pageId="7" type="paratype">Paratype</typeStatus>
</emphasis>
, Brotas municipality, Fazenda
<collectingCountry id="B9E0D6CAFFCEFF80FC16F975ED39FB9B" box="[962,1095,1673,1698]" name="Saint Lucia" pageId="7">Santa Lúcia</collectingCountry>
da Boa Vista (
<geoCoordinate id="A4C3F09DFFCEFF80FB39F975ECE6FB98" box="[1261,1432,1673,1697]" direction="south" orientation="latitude" pageId="7" precision="1" value="-22.2108">22°12’38.88”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFCEFF80FF43F951E837FBFF" box="[151,329,1709,1734]" direction="west" orientation="longitude" pageId="7" precision="1" value="-48.2433">48°14’35.88”W</geoCoordinate>
,
<quantity id="060F3BBFFFCEFF80FE8DF951E8DAFBFC" box="[345,420,1709,1734]" metricMagnitude="2" metricUnit="m" metricValue="4.85" pageId="7" unit="m" value="485.0">485 m</quantity>
a.s.l.),
<date id="B549B09AFFCEFF80FDD4F951EBFCFBFC" box="[512,642,1709,1733]" pageId="7" value="2001-11-02">02.XI.2001</date>
, F.A.A. Lencioni leg. (RWG, not examined).
<emphasis id="F3834A48FFCEFF80FB48F951ECE2FBFF" bold="true" box="[1180,1436,1709,1734]" pageId="7">Additional specimen.</emphasis>
<collectingCountry id="B9E0D6CAFFCEFF80FF43F92DE87EFBD0" box="[151,256,1745,1769]" name="Brazil" pageId="7">BRAZIL</collectingCountry>
. Minas Gerais State: 1♀, Serra do Salitre municipality, river at riparian forest,
<bibRefCitation id="A566EBABFFCEFF80FBA7F92DEDB9FBD0" author="II" box="[1139,1223,1745,1769]" pageId="7" refString="GEOnet Names Server (GNS), National Geospatial-Intelligence Agency (2014) U. S. National Imagery and Mapping Agency's (NIMA) database of foreign geographic feature names. Available from: http: // geonames. nga. mil / gns / html / (accessed October, 2014). [Argentina gazetteer available from: http: // biogeo. ucdavis. edu / data / diva / gaz / ARG _ gaz. zip; Uruguay gazetteer available from: http: // biogeo. ucdavis. edu / data / diva / gaz / URY _ gaz. zip]" type="book chapter" year="2014">
<date id="B549B09AFFCEFF80FBA7F92DEDB9FBD0" box="[1139,1223,1745,1769]" pageId="7" value="2014-02">II.2014</date>
</bibRefCitation>
, J.E. Santos Junior leg. (
<collectionCode id="A7E60E9FFFCEFF80FF1BF909E850FA34" box="[207,302,1781,1805]" pageId="7">ABMM</collectionCode>
). Specimens in
<collectionCode id="A7E60E9FFFCEFF80FE0AF90AEB5FFA34" box="[478,545,1782,1805]" pageId="7">DZRJ</collectionCode>
collected under ICMBIO/SISBIO licenses numbers 14591-8 and 23620-6.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFCEFF8BFF13F8E5E8EAF8DE" lastPageId="12" pageId="7" type="description">
<paragraph id="C148965AFFCEFF8FFF13F8E5ED1EFCCC" blockId="7.[151,1437,1385,2022]" lastBlockId="8.[151,1437,151,2016]" lastPageId="8" pageId="7">
<emphasis id="F3834A48FFCEFF80FF13F8E5EBF4FA0B" bold="true" box="[199,650,1817,1842]" pageId="7">
Description of male
<typeStatus id="1E4C28F8FFCEFF80FE1AF8E5EB49FA0B" box="[462,567,1817,1842]" pageId="7" type="holotype">holotype</typeStatus>
. Head
</emphasis>
(
<figureCitation id="59CC8ADFFFCEFF80FD75F8E5EB92FA08" box="[673,748,1817,1842]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="7">Fig. 4</figureCitation>
A). Labium greenish-yellow, visible parts of maxilla and mandible greenish-yellow, except dark-brown to glossy black movable hook, extreme apex of maxillary outer lobe and mandibular incisor lobe. Genae brown. Labrum greenish-yellow, with a narrow straight dark brown to black stripe at ventral margin, one lateral margin with a narrow dark brown line from ventral margin to level of clypeolabral suture, clypeolabral suture light brown connected to a small mesal rounded spot over labrum occupying about half of its total height. Anteclypeus brownish-yellow, clypeus greenish-yellow. Frons yellowishgreen, darker than remainder of face, with dark brown bar at posterior 0.4, extended laterally. Antenna, vertex and occiput dark brown to black, except yellowish membranous part of scape and area close to postocellar suture noticeably lighter than adjacent areas of cranium. Posterior region of cranium (“postgena” plus “occiput”) gradually lightening ventrally with irregular dark areas, especially close to postoccipital suture. Ventral margin of labrum with a wide concavity almost mesal, giving to sclerite a slight asymmetric shape; ante- and postfrons division carinated, dorsal margin of frons sinuous in anterior view with a small mesal concavity, anterior margin regularly convex and with a shallow mesal furrow in dorsal view. Posterior portion of vertex with a ridge (postocellar ridge
<emphasis id="F3834A48FFC1FF8FFEBCFE69E8D8FC95" box="[360,422,405,428]" italics="true" pageId="8">sensu</emphasis>
Belle 1973) slightly projecting anteriorly over lateral ocelli, occiput (occipital plate) in dorsal view with a distinct ridge along its entire width (
<figureCitation id="59CC8ADFFFC1FF8FFCF2FE4BEA13FCE9" box="[806,877,439,464]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="8">Fig. 4</figureCitation>
A), area posterior to ridge distinctly higher than anterior, posterior margin of occiput with scattered setae, not forming a distinct fringe.
</paragraph>
<paragraph id="C148965AFFC1FF8FFF13FE03ED79F86D" blockId="8.[151,1437,151,2016]" pageId="8">
<emphasis id="F3834A48FFC1FF8FFF13FE03E858FF21" bold="true" box="[199,294,511,536]" pageId="8">Thorax.</emphasis>
Prothorax greenish-yellow, lightening ventrally to basisternum with dark brown spots or stripes on anterior margin of anterior lobe, an ill-defined spot on dorsum of median lobe and distinct stripes surrounding central pale area of posterior lobe. Synthorax (
<figureCitation id="59CC8ADFFFC1FF8FFD6AFDBBEA7AFF59" box="[702,772,583,608]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="8">Figs 1</figureCitation>
A–B) largely pale, patterned with dark brown spots and greenish-yellow stripes dorsally, almost entirely greenish-yellow laterally; dorsal carina dark on anterior third, then with narrow yellowish line; mesepisternum with first and second pale antehumeral stripes distinct, first antehumeral stripe joined to pale mesothoracic collar, to second antehumeral stripe at about half length, and at level of antealar sinus (
<figureCitation id="59CC8ADFFFC1FF8FFE89FD2BE8DEFFC9" box="[349,416,727,752]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="8">Figs 1</figureCitation>
A–B); pale areas enclosing three large dark spots, a triangular dark brown mesal spot from collar to antealar sinus, at maximum half pleurite width, a barely defined rounded brown spot from mesinfraepisternum to about of posterior 0.4, and an ellipsoid dark brown spot posteriorly; a narrow brown stripe parallel to the pale mesopleural suture (humeral) beginning at 0.2 height of epimeron and extending to near antealar sinus, enlarged posteriorly; mesepisternum with a dark brown stripe parallel to mesopleural suture darkened and wider ventrally close to mesinfraepisternum (
<figureCitation id="59CC8ADFFFC1FF8FFD7AFC70EB88FE9D" box="[686,758,908,933]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="8">Figs 1</figureCitation>
A–B); metepisternum with a barely defined brown stripe dorsally from metapleural antealar process to intersegmental suture, remainder of synthorax greenish-yellow lightening to light yellow ventrally, with irregular ill-defined dark areas laterally (
<figureCitation id="59CC8ADFFFC1FF8FFBBAFC28EDCAFED5" box="[1134,1204,980,1005]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="8">Fig. 1</figureCitation>
B). Legs, coxa and trochanter yellowish-brown, ventral and posteroventral surfaces of femur light yellow, anteroventral and dorsal surfaces dark brown to black in pro- and mesothoracic legs, in metathoracic leg yellowish-brown with irregular dark areas and distal 0.1 black; tibia black with dorsal surface yellow to brownish-yellow (
<figureCitation id="59CC8ADFFFC1FF8FFB4AFBC3ED9FF961" box="[1182,1249,1087,1112]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="8">Fig. 3</figureCitation>
A); tarsus black, pretarsal claws dark brown with distinct supplementary inferior tooth. Proximal portion of trochanter (1
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<emphasis id="F3834A48FFC1FF8FFA8AFB99ECE2F945" box="[1374,1436,1125,1148]" italics="true" pageId="8">sensu</emphasis>
<bibRefCitation id="A566EBABFFC1FF8FFF43FB7BE835F999" author="Snodgrass" box="[151,331,1159,1184]" pageId="8" refString="Snodgrass, R. E. (1935) Principles of insect morphology. McGraw-Hill Book Company, New York, 667 pp." type="book" year="1935">Snodgrass 1935</bibRefCitation>
) armed with a row of small black spurs only in the mesothoracic leg, distal portion (2
<collectingCountry id="B9E0D6CAFFC1FF8FFADCFB74EC5BF999" box="[1288,1317,1160,1184]" name="Chile" pageId="8">Tr</collectingCountry>
) armed on all legs, most numerous on mesothoracic leg. Femur with rows of numerous small black spurs, metathoracic femur slight convex anteriorly in lateral view, anteroventral surface of prothoracic tibia armed with 10–12 spurs (including scale-like spurs of tibial comb), 11–12 on mesothoracic, 14–15 on metathoracic, posteroventral surface with 14–15 on prothoracic, 14–15 on mesothoracic, 13–14 on metathoracic, usually at least two times longer than intervening space. Tibial and tarsal spurs modified, basally enlarged (
<figureCitation id="59CC8ADFFFC1FF8FFC74FAC0EA9DF86D" box="[928,995,1340,1365]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="8">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="C148965AFFC1FF8FFF13FAA3EBF5FA3E" blockId="8.[151,1437,151,2016]" pageId="8">
Wings (
<figureCitation id="59CC8ADFFFC1FF8FFEF0FAA3E812F841" box="[292,364,1375,1400]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="8">Fig. 2</figureCitation>
A). Membrane hyaline; venation dark-brown to black; pt brown, contrasting with black of marginal veins; one basal subcostal crossvein in all wings, membranula strongly reduced. Venation as follows: 12–13 Ax in Fw, 9–10 Ax in Hw, Ax2 the 5th in all wings; 9 Px all wings; 8–9 postsubnodals (including brace vein) in Fw,
<quantity id="060F3BBFFFC1FF8FFF3CFA33E868F8DE" box="[232,278,1486,1511]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" unit="in" value="8.0">8 in</quantity>
Hw; 5–6 bridge crossveins in Fw,
<quantity id="060F3BBFFFC1FF8FFD79FA33EBA4F8DE" box="[685,730,1486,1511]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" unit="in" value="4.0">4 in</quantity>
Hw; arc at or slightly distal to second Ax; sectors of arc not stalked, origin located 0.5 basal of arc in all wings; 3–4 crossveins in the space between arc and RP-midfork in Fw,
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Hw; Rspl indistinct in all wings; discoidal triangles, supratriangles and subtriangles not crossed in four wings; space between CuP-crossing and PsA with one crossvein in all wings; Fw discoidal field with two rows of cells up to level of nodus, beyond that three and increasing to 9–11 cells at wing margin; Hw discoidal field divergent, with 4–5 rows of two cells, followed by 1–2 rows of three cells, then broadening to 9–11 cells towards wing margin; Mspl indistinct in the four wings; anal loop rectangular with four cells; seven paranal cells from wing base to apex of subtriangle in Fw, five in Hw; anal triangle distinct, three-celled, with seven strong and short spines along the dorso-proximal margin (AP&amp;AA).
</paragraph>
<paragraph id="C148965AFFC1FF8EFF13F8EEE81FF841" blockId="8.[151,1437,151,2016]" lastBlockId="9.[151,1437,151,1977]" lastPageId="9" pageId="8">
<emphasis id="F3834A48FFC1FF8FFF13F8EEE842FA12" bold="true" box="[199,316,1810,1835]" pageId="8">Abdomen</emphasis>
(
<figureCitation id="59CC8ADFFFC1FF8FFE99F8EEE8DFFA12" box="[333,417,1810,1835]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="8">Fig. 10</figureCitation>
A). Dark brown with pale greenish-yellow spots. Tergite of S1 almost entirely greenishyellow expect the dark brown rounded process (swelling of abdominal articulation
<emphasis id="F3834A48FFC1FF8FFB9AF8C4EDF2FA76" box="[1102,1164,1848,1871]" italics="true" pageId="8">sensu</emphasis>
<bibRefCitation id="A566EBABFFC1FF8FFB41F8CAEC4DFA76" author="Asahina" box="[1173,1331,1846,1871]" pageId="8" refString="Asahina, S. (1954) A morphological study of a relic dragonfly Epiophlebia superstes. The Japan Society for the promotion of science publ., Tokyo, iv + 153 pp., 71 pls." type="book" year="1954">Asahina 1954</bibRefCitation>
) close to posterior carina; S2 with a large pale spot on anterior 0.6, narrowing posteriad to 0.33 of segment height in lateral view, a small pale spot close to posterior carina and a middorsal pale line along entire segment; S3–7 with a pale proximal ring up to transverse carina, darkened in its middle, remainder of segment posteriorly dark brown to black; S8 almost entirely dark brown with an irregular lateroventral pale spot, enlarged at both ends of segment; S9 dark brown with ill-defined lateral lighter areas; S10 dark brown above, with pale lateral spot extended dorsally in a narrow ring at 0.66 length separating a rounded basal spot from the dark posterior end (
<figureCitation id="59CC8ADFFFC0FF8EFB1CFF40EC6CFDED" box="[1224,1298,188,213]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
A–C, 10A); sternites dark brown to black; ventral carina of S2–8 and anterior 0.5 of S9 greenish-yellow; S1–7 cylindrical, S8–10 distinctly wider than previous segments, maximum width at S8. Auricles rounded, positioned obliquely to S2 longitudinal axis, length about one-third height of S2; posterior margin with two patches of 5–6 minute dark spines; posterior carina of S9 armed with small spines dorsally and with V-shaped concavity. Secondary genitalia typical of the
<taxonomicName id="06F7EDD9FFC0FF8EFEE0FE93E8A7FCB1" box="[308,473,367,392]" class="Insecta" family="Gomphidae" genus="Agriogomphus" kingdom="Animalia" order="Odonata" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFC0FF8EFEE0FE93E8A7FCB1" box="[308,473,367,392]" italics="true" pageId="9">Agriogomphus</emphasis>
</taxonomicName>
-complex, anterior lamina not visible in lateral view, in ventral view posterior margin with a wide U-shaped ridge, anterior hamule (
<figureCitation id="59CC8ADFFFC0FF8EFD60FE68EB84FC95" box="[692,762,404,429]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="9">Fig. 4</figureCitation>
B) yellowish-brown, quadrangular in lateral view, anterior margin strongly ridged, sickle-shaped, posteriorly prolonged into a concave laminar projection with scattered hairlike setae denser posteriorly, a small ventral V-shaped concavity between anterior ridge and posterior laminar projection; in ventral view posterior margin convergent, the limit between anterior ridge and posterior laminar projection concave in a wide V-shape; posterior hamule (
<figureCitation id="59CC8ADFFFC0FF8EFCCAFDD8EA1DFF05" box="[798,867,548,573]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="9">Fig. 4</figureCitation>
B) light brown, except black stout acute anteriorly directed apex, distinctly longer than other structures of secondary genitalia in lateral view, anterior margin slightly sinuous, slightly convex at proximal 0.4 then gently concave, with small deep concavity before extreme distal; two distinct sets of pale hair-like long setae, a basal line with 8–10 setae at anterior margin and a distal tuft with about a dozen setae close to acute apex (
<figureCitation id="59CC8ADFFFC0FF8EFDF5FD48EB14FFF5" box="[545,618,692,717]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="9">Fig. 4</figureCitation>
B); in ventral view regularly curved toward convergent black apexes, somewhat flattened; without a distinct genital lobe. Vesica spermalis (
<figureCitation id="59CC8ADFFFC0FF8EFC18FD2BED6AFFC9" box="[972,1044,727,752]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="9">Fig. 4</figureCitation>
B) with
<date id="B549B09AFFC0FF8EFBA3FD24EDE7FFC9" box="[1143,1177,728,752]" pageId="9">V1</date>
black, in lateral view slightly higher than level of anterior hamule, scoop-like process on
<date id="B549B09AFFC0FF8EFC5AFD00EACEFE2D" box="[910,944,764,788]" pageId="9">V1</date>
(median cleft
<emphasis id="F3834A48FFC0FF8EFB87FD01EDEFFE2D" box="[1107,1169,765,788]" italics="true" pageId="9">sensu</emphasis>
<bibRefCitation id="A566EBABFFC0FF8EFB4CFD00EC6AFE2D" author="Belle" box="[1176,1300,764,789]" pageId="9" refString="Belle, J. (1988) A synopsis of the species of Phyllocycla Calvert, with descriptions of four new taxa and a key to the genera of Neotropical Gomphidae (Odonata, Gomphidae). Tijdschrift voor Entomologie, 131, 73 - 102." type="journal article" year="1988">Belle 1988</bibRefCitation>
) rounded in front view, strongly concave and covered with hair-like yellow setae internally; see
<typeStatus id="1E4C28F8FFC0FF8EFB82FCDCEDC8FE01" box="[1110,1206,800,824]" pageId="9" type="paratype">paratype</typeStatus>
for full description. Cercus (
<figureCitation id="59CC8ADFFFC0FF8EFF21FCB8E842FE65" box="[245,316,836,861]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
A–C, 8A–B) greenish-yellow spotted with brown by numerous tubercular setal alveoli proximally, darkening to black apex, ventral branch dark brown (
<figureCitation id="59CC8ADFFFC0FF8EFD3EFC9BEA50FEB9" box="[746,814,871,896]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
A); in lateral view cercus strongly curved at half-length at level of epiproct posteriorly, directed obliquely dorsad (
<figureCitation id="59CC8ADFFFC0FF8EFCB4FC70EAD0FE9D" box="[864,942,908,933]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
A, 8A); apex acute and slightly directed posteriorly; posterior margin flattened, forming a distinct rounded keel (
<figureCitation id="59CC8ADFFFC0FF8EFC6FFC53EA83FEF1" box="[955,1021,943,968]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
A); ventral branch strongly curved in lateral view with apex directed anteriorly (
<figureCitation id="59CC8ADFFFC0FF8EFDAEFC28EBBCFED5" box="[634,706,980,1005]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
A, 8A); cercus in dorsal view cylindrical, tapering posteriorly to acute apex, slightly divergent (
<figureCitation id="59CC8ADFFFC0FF8EFE2EFC0BEB3EF929" box="[506,576,1015,1040]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
B); mesal margin with a rounded basal projection, then gradually tapering, external margin almost straight (
<figureCitation id="59CC8ADFFFC0FF8EFDC0FBE0EB22F90D" box="[532,604,1052,1077]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
B); cerci in ventral view touching each other proximally and gradually diverging distally, apexes almost parallel, ventral branch strongly directing parallel toward anteriorly, apex rounded, slightly flattened (
<figureCitation id="59CC8ADFFFC0FF8EFE1AFB98EB6CF945" box="[462,530,1124,1149]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
C). Epiproct dark brown, largely bifid, posteriorly at level of cercus (
<figureCitation id="59CC8ADFFFC0FF8EFACDFB98EC1EF945" box="[1305,1376,1124,1149]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
A, C, 8A–B); almost straight in lateral view, with a dorsal and ventral ridge at proximal 0.25, a small tubercle-like rounded process on dorsal margin at ca. distal 0.70 (
<figureCitation id="59CC8ADFFFC0FF8EFD39FB50EA4BF9FD" box="[749,821,1196,1221]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
A, 8A–B); laterodistal projection forefinger-like, apex extending obliquely upward, posterior margin slightly concave (
<figureCitation id="59CC8ADFFFC0FF8EFC72FB33EA88F9D1" box="[934,1014,1231,1256]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
A, 8A–B); in dorsal view slightly convergent, dilated at about posterior 0.70 arms surrounding a deep quadrangular concavity (hereafter distal concavity,
<figureCitation id="59CC8ADFFFC0FF8EFEC5FAEBE82DF809" box="[273,339,1303,1328]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="9">Fig. 8</figureCitation>
B); in dorsolateral view with a strong tubercle-like process anterior to the concavity (
<figureCitation id="59CC8ADFFFC0FF8EFACCFAEBEC25F809" box="[1304,1371,1303,1328]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="9">Fig. 8</figureCitation>
B); in ventral view almost straight, laterodistal projection distinctly posterior to internal margin, posterior margin slightly sinuous (
<figureCitation id="59CC8ADFFFC0FF8EFF28FAA3E841F841" box="[252,319,1375,1400]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
C).
</paragraph>
<paragraph id="C148965AFFC0FF8EFF13FA78EBB4FB31" blockId="9.[151,1437,151,1977]" pageId="9">
<emphasis id="F3834A48FFC0FF8EFF13FA78E803F8A4" bold="true" box="[199,381,1412,1437]" pageId="9">Measurements.</emphasis>
Total length (including caudal appendages) 40.5; abdomen length (excluding caudal appendages) 29.2; head maximum width 5.7; Fw length 25; Hw length 23.5; Fw maximum width 5.6, in Hw 7.2; pt length
<quantity id="060F3BBFFFC0FF8EFF32FA30E856F8DC" box="[230,296,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="7.111999999999999" pageId="9" unit="in" value="2.8">2.8 in</quantity>
Fw,
<quantity id="060F3BBFFFC0FF8EFEB6FA30E8DAF8DC" box="[354,420,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="7.366" pageId="9" unit="in" value="2.9">2.9 in</quantity>
Hw; length of metathoracic femur 6.8; metathoracic tibia 5.4; length of S9+
<quantity id="060F3BBFFFC0FF8EFAC7FA30EC30F8DC" box="[1299,1358,1484,1509]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="9" unit="in" value="10.0">10 in</quantity>
lateral view 3.3; total length of cercus in lateral view 1.4.
</paragraph>
<paragraph id="C148965AFFC0FF8EFF13F9E8E82CFBD9" blockId="9.[151,1437,151,1977]" pageId="9">
<emphasis id="F3834A48FFC0FF8EFF13F9E8EB31FB14" bold="true" box="[199,591,1556,1581]" pageId="9">
Description of female
<typeStatus id="1E4C28F8FFC0FF8EFE09F9E8EB34FB14" box="[477,586,1556,1581]" pageId="9" type="paratype">paratype</typeStatus>
.
</emphasis>
Similar to
<typeStatus id="1E4C28F8FFC0FF8EFD08F9E8EA3FFB14" box="[732,833,1556,1581]" pageId="9" type="holotype">holotype</typeStatus>
, only the differences described here.
<emphasis id="F3834A48FFC0FF8EFAD5F9E8EC36FB14" bold="true" box="[1281,1352,1556,1581]" pageId="9">Head.</emphasis>
Genae greenish-yellow. Dark brown to black stripe at ventral margin of labrum prolonged laterally to dorsal 0.5. Frons uniformly light brown, darker than remainder of face. Vertex and occiput light brown, the latter slightly paler. Posterior region of cranium (“postgena” plus “occiput”) light brown with irregular pale areas. Ante- and postfrons division slightly carinated. Occiput (occipital plate) behind carina clearly swollen, posterior margin sinuous with a mesal concavity.
</paragraph>
<paragraph id="C148965AFFC0FF8EFF13F910E81DFA81" blockId="9.[151,1437,151,1977]" pageId="9">
<emphasis id="F3834A48FFC0FF8EFF13F910E858FA3C" bold="true" box="[199,294,1772,1797]" pageId="9">Thorax.</emphasis>
Prothorax with posterior lobe entirely greenish-yellow. Synthorax with dark areas slightly paler than in
<typeStatus id="1E4C28F8FFC0FF8EFF60F8F3E868FA11" box="[180,278,1807,1832]" pageId="9" type="holotype">holotype</typeStatus>
(
<figureCitation id="59CC8ADFFFC0FF8EFEF1F8F3E819FA11" box="[293,359,1807,1832]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="9">Figs 1</figureCitation>
E–F, 11A); pale area over dorsal carina larger. Legs with dark areas on anteroventral and dorsal surfaces more extensive; tibia dark brown to black with dorsal surface yellow (
<figureCitation id="59CC8ADFFFC0FF8EFBDAF8C8ED20FA75" box="[1038,1118,1844,1869]" captionStart="FIGURE 11" captionStartId="23.[151,250,1163,1185]" captionTargetBox="[151,1436,193,1140]" captionTargetId="figure@23.[151,1436,193,1141]" captionTargetPageId="23" captionText="FIGURE 11. Female habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Paratype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448), B. Additional specimen (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); D. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270864/files/figure.png" pageId="9">Fig. 11</figureCitation>
A). Anteroventral surface of prothoracic tibia armed with 11 spurs, 12–13 on mesothoracic, 16 on metathoracic, posteroventral surface of prothoracic tibia with 14, 16 on mesothoracic, 13–15 on metathoracic, usually more than two times longer than intervening space.
</paragraph>
<caption id="9588C6D2FFC3FF8DFF43F9C2EC55FBE3" httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="10" targetBox="[154,1436,151,1575]" targetPageId="10">
<paragraph id="C148965AFFC3FF8DFF43F9C2EC55FBE3" blockId="10.[151,1436,1598,1754]" pageId="10">
<emphasis id="F3834A48FFC3FF8DFF43F9C2E86DFB6A" bold="true" box="[151,275,1598,1620]" pageId="10">FIGURE 4</emphasis>
. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B–F). A–B.
<taxonomicName id="06F7EDD9FFC3FF8DFF43F99CE894FB4F" box="[151,490,1632,1654]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC3FF8DFF43F99CE894FB4F" box="[151,490,1632,1654]" italics="true" pageId="10">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC3FF8DFE22F99CEB32FB4C" bold="true" box="[502,588,1632,1653]" pageId="10">
<taxonomicNameLabel id="E8B0F733FFC3FF8DFE22F99CEB32FB4C" box="[502,588,1632,1653]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C.
<taxonomicName id="06F7EDD9FFC3FF8DFF43F97DEB39FBAE" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[151,583,1665,1687]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFC3FF8DFF43F97DE8BAFBAE" box="[151,452,1665,1687]" italics="true" pageId="10">Cyanogomphus comparabilis</emphasis>
Belle, 1994
</taxonomicName>
, Holotype (Brazil. MT: Diamantino, ABMM); D.
<taxonomicName id="06F7EDD9FFC3FF8DFB8EF97DE9B4FB80" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC3FF8DFB8EF97DEC2BFBAE" box="[1114,1365,1665,1687]" italics="true" pageId="10">Cyanogomphus waltheri</emphasis>
Selys, 1873
</taxonomicName>
(Brazil. RJ: Santana de Japuíba district, MNRJ); E.
<taxonomicName id="06F7EDD9FFC3FF8DFCC4F95FEC6FFB80" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[784,1297,1699,1721]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFC3FF8DFCC4F95FEA9DFB80" box="[784,995,1699,1721]" italics="true" pageId="10">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</taxonomicName>
(Brazil. RS: Jaguarão, MNRJ); F.
<taxonomicName id="06F7EDD9FFC3FF8DFEBBF938EB96FBE3" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[367,744,1732,1754]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFC3FF8DFEBBF938EB2AFBE3" box="[367,596,1732,1754]" italics="true" pageId="10">Tibiagomphus uncatus</emphasis>
(Fraser, 1947)
</taxonomicName>
(Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="C148965AFFC3FF8DFF13F8F8EB62FACC" blockId="10.[151,1436,1796,2037]" pageId="10">
Wings (
<figureCitation id="59CC8ADFFFC3FF8DFECAF8F8E81DFA24" box="[286,355,1796,1821]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="10">Fig. 2</figureCitation>
B). Venation as follows: 14 Ax in Fw,
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Hw, 9 Px in Fw,
<quantity id="060F3BBFFFC3FF8DFBC4F8F9ED0CFA24" box="[1040,1138,1796,1821]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="10" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10–11 in</quantity>
Hw; 10–11 postsubnodals (including brace vein) in Fw,
<quantity id="060F3BBFFFC3FF8DFE30F8D4EB44FA78" box="[484,570,1832,1857]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="10" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9–10 in</quantity>
Hw; 7 bridge crossveins in Fw,
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Hw; arc at or slightly distal to second Ax; 4 crossveins in space between arc and RP-midfork in Fw,
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Hw; Fw discoidal field increasing to 10–11 cells at wing margin; Hw discoidal field with 4–6 rows of two cells, followed by 1–2 rows of three cells, then broadening to 11 cells toward wing margin; anal loop barely defined with 6 cells; 6–7 paranal cells from wing base to apex of subtriangle in Fw; anal wing base rounded without distinct anal triangle and strong spines along of the dorso-proximal margin (AP&amp;AA).
</paragraph>
<caption id="9588C6D2FFC2FF8CFF43FA03EA75FB48" httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="11" targetBox="[151,1436,195,1507]" targetPageId="11">
<paragraph id="C148965AFFC2FF8CFF43FA03EA75FB48" blockId="11.[151,1436,1535,1650]" pageId="11">
<emphasis id="F3834A48FFC2FF8CFF43FA03E870FB2D" bold="true" box="[151,270,1535,1557]" pageId="11">FIGURE 5</emphasis>
. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A–D) and posterior hamule in lateral view (E–L). A–B, F–H.
<taxonomicName id="06F7EDD9FFC2FF8CFE64F9E1EAC2FB0A" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[432,956,1565,1587]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="11" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFC2FF8CFE64F9E1EBF8FB0A" box="[432,646,1565,1587]" italics="true" pageId="11">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</taxonomicName>
(Brazil. RS: Jaguarão, MNRJ); C–D, I–L.
<taxonomicName id="06F7EDD9FFC2FF8CFF43F9C0EB5EFB6B" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[151,544,1596,1618]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="11" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFC2FF8CFF43F9C0E8FFFB6B" box="[151,385,1596,1618]" italics="true" pageId="11">Tibiagomphus uncatus</emphasis>
(Fraser, 1947)
</taxonomicName>
(Brazil. SC: Nova Teutônia, MZSP). E.
<taxonomicName id="06F7EDD9FFC2FF8CFC35F9C0ECE6FB6B" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[993,1432,1596,1618]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="11" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFC2FF8CFC35F9C0EC71FB6B" box="[993,1295,1596,1618]" italics="true" pageId="11">Cyanogomphus comparabilis</emphasis>
Belle, 1994
</taxonomicName>
, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="C148965AFFC2FF8CFF13F964E8C1FA5C" blockId="11.[151,1436,1688,2037]" pageId="11">
<emphasis id="F3834A48FFC2FF8CFF13F964E842FB88" bold="true" box="[199,316,1688,1713]" pageId="11">Abdomen</emphasis>
(only S1–4). Largely pale, greenish-yellow with irregular dark brown transverse stripes and irregular spots, transverse carina ill defined (
<figureCitation id="59CC8ADFFFC2FF8CFD5BF940EBA1FBEC" box="[655,735,1724,1749]" captionStart="FIGURE 11" captionStartId="23.[151,250,1163,1185]" captionTargetBox="[151,1436,193,1140]" captionTargetId="figure@23.[151,1436,193,1141]" captionTargetPageId="23" captionText="FIGURE 11. Female habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Paratype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448), B. Additional specimen (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); D. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270864/files/figure.png" pageId="11">Fig. 11</figureCitation>
A). S2 yellowish lateroventrally in anterior 0.5, light brown at level of transverse carina (not carinated) and dark brown at posterior field, S3–4 with a dark brown stripe on posterior 0.8 adjacent to ventral carina, enlarging and darkening posteriorly, a transverse dark brown stripe at level of transverse carina and at posterior field, and a dorsolateral oval dark brown spot not connected dorsally, similar to PD spot of
<bibRefCitation id="A566EBABFFC2FF8CFEC1F8B0E8C4FA5C" author="Walker" box="[277,442,1868,1893]" pageId="11" refString="Walker, E. M. (1912) The North American dragonflies of the genus Aeshna. University of Toronto Studies, Biological Series, 11, 1 - 213, pls. 1 - 28." type="journal article" year="1912">Walker (1912)</bibRefCitation>
.
</paragraph>
<paragraph id="C148965AFFC2FF8CFF13F88CEC77FA94" blockId="11.[151,1436,1688,2037]" pageId="11">
<emphasis id="F3834A48FFC2FF8CFF13F88CE803FAB0" bold="true" box="[199,381,1904,1929]" pageId="11">Measurements.</emphasis>
Head maximum width 5.7; Fw length 25.8–26.0; Hw length 24.4–24.5; Fw maximum width 5.7, in Hw 7.2; pt length
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Fw, 3.0 in Hw; length of metathoracic femur 6.3; metathoracic tibia 5.
</paragraph>
<paragraph id="C148965AFFC2FF8BFF13F844EC43F836" blockId="11.[151,1436,1688,2037]" lastBlockId="12.[151,1437,151,2016]" lastPageId="12" pageId="11">
<emphasis id="F3834A48FFC2FF8CFF13F844EB2CFAE8" bold="true" box="[199,594,1976,2001]" pageId="11">
Variation in the male
<typeStatus id="1E4C28F8FFC2FF8CFE01F844EB30FAE8" box="[469,590,1976,2001]" pageId="11" type="paratype">paratypes</typeStatus>
.
</emphasis>
The general appearance in size and color of the male from Lagoa Santa (MNRJ) is very similar to that of the
<typeStatus id="1E4C28F8FFC2FF8CFDEBF820EBDAFACC" box="[575,676,2012,2037]" pageId="11" type="holotype">holotype</typeStatus>
, though slightly paler, while that from Serra do Salitre (ABMM) is smaller and distinctly darker with first and second humeral stripes clearly defined and limited by dark ground color (
<figureCitation id="59CC8ADFFFC5FF8BFF4BFF40E99FFDED" box="[159,225,188,213]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="12">Figs 1</figureCitation>
C–D, 10B). They agree with the
<typeStatus id="1E4C28F8FFC5FF8BFD80FF40EBC8FDEC" box="[596,694,188,213]" pageId="12" type="holotype">holotype</typeStatus>
except as follows (characters observed in one or both specimens):
<emphasis id="F3834A48FFC5FF8BFF43FF23E9AAFDC1" box="[151,212,223,248]" italics="true" pageId="12">Head</emphasis>
. Labrum with the black distal stripe almost restricted to ventral margin, mesal dark spot longer, occupying 0.66 of its total length (height). Anteclypeus greenish-yellow. Frontoclypeal suture with a distinct brownish stripe widening dorsolaterally over frons.
<emphasis id="F3834A48FFC5FF8BFDFCFEDBEB07FC79" box="[552,633,295,320]" italics="true" pageId="12">Thorax</emphasis>
. Prothorax with dark brown areas better defined. Synthorax with larger and clearly better defined dark areas; mesepisternum at level of junction between first and second pale antehumeral stripes only slightly paler than anterior and posterior dark areas, pale areas not enclosing dark spots, showing a regular gomphid pattern with defined mesepisternal stripes (
<figureCitation id="59CC8ADFFFC5FF8BFC97FE68EAF9FC95" box="[835,903,404,429]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="12">Figs 1</figureCitation>
C–D); metepisternal dark stripe better defined. Legs, coxa and trochanter greenish-yellow; tibia dark brown to black with dorsal surface yellow. Anteroventral surface of prothoracic tibia armed with 10–11 spurs, 11–12 on mesothoracic, 14–18 on metathoracic, posteroventral surface 13–14 on prothoracic, 13 on mesothoracic, 14–16 on metathoracic. Wings (
<figureCitation id="59CC8ADFFFC5FF8BFAF0FE03EC05FF21" box="[1316,1403,511,536]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="12">Fig. 10</figureCitation>
B). Membrane hyaline, very slightly smoked with light brown. Venation as follows: one Hw lacking basal subcostal crossvein; 11–13 Ax in Fw, 10 Ax in Hw, Ax2 the 4th in one Fw and one Hw; 7–10 Px in Fw,
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Hw; 8–9 postsubnodals (including brace vein) in Fw,
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Hw; 5–7 bridge crossveins in Fw,
<quantity id="060F3BBFFFC5FF8BFB43FD92EDB8FFBE" box="[1175,1222,622,647]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="12" unit="in" value="5.0">5 in</quantity>
Hw; arc at or 0.3 proximal to second Ax; 3–5 crossveins in the space between arc and RP-midfork in Fw,
<quantity id="060F3BBFFFC5FF8BFB57FD6FEDB5FF92" box="[1155,1227,658,683]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="12" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
Hw; Fw discoidal field 9–12 cells at wing margin; Hw discoidal field with 2–5 rows of two cells, followed by 1–2 rows of three cells, then broadening to 9–13 cells towards wing margin; anal loop barely defined with five cells; 6 paranal cells from wing base to apex of subtriangle in Fw,
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Hw; 6–9 spines along of the dorso-proximal margin (AP&amp;AA).
<emphasis id="F3834A48FFC5FF8BFF43FCDEE87FFE02" box="[151,257,802,827]" italics="true" pageId="12">Abdomen</emphasis>
. S2 with the large pale spot with irregular margin; S7 with a pale spot entirely greenish-yellow; S8 with a round pale spot at anterior 0.33; S10 distinctly paler than former segments with ill-defined paler areas. Posterior margin of auricles with irregular rows containing 8–14 minute spines, 5 setae at the line of basal hair-like setae on anterior margin of posterior hamule. Vesica spermalis (based on
<typeStatus id="1E4C28F8FFC5FF8BFC7AFC6CED71FE91" box="[942,1039,912,936]" pageId="12" type="paratype">paratype</typeStatus>
from Lagoa Santa,
<figureCitation id="59CC8ADFFFC5FF8BFADEFC72EC29FE9E" box="[1290,1367,910,935]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
A–C). Sclerotized parts dark brown to black; ventral surface of
<date id="B549B09AFFC5FF8BFCC9FC4FEA41FEF2" box="[797,831,947,971]" pageId="12">V1</date>
with two processes, a small rounded one at proximal third and a large strong scoop-like process at distal third (median cleft
<emphasis id="F3834A48FFC5FF8BFC01FC24ED6DFED6" box="[981,1043,984,1007]" italics="true" pageId="12">sensu</emphasis>
<bibRefCitation id="A566EBABFFC5FF8BFBCAFC2AEDE0FED6" author="Belle" box="[1054,1182,982,1007]" pageId="12" refString="Belle, J. (1988) A synopsis of the species of Phyllocycla Calvert, with descriptions of four new taxa and a key to the genera of Neotropical Gomphidae (Odonata, Gomphidae). Tijdschrift voor Entomologie, 131, 73 - 102." type="journal article" year="1988">Belle 1988</bibRefCitation>
), the latter laminar in lateral view, in anteroposterior view with converging rounded apexes and covered with hair-like yellow setae internally (
<figureCitation id="59CC8ADFFFC5FF8BFEC5FBE2E827F90E" box="[273,345,1054,1079]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
A, C);
<date id="B549B09AFFC5FF8BFE72FBE3E8B7F90E" box="[422,457,1055,1079]" pageId="12">V2</date>
tubular, lacking a defined H;
<date id="B549B09AFFC5FF8BFCCDFBE3EA45F90E" box="[793,827,1055,1079]" pageId="12">V3</date>
globose in ventral view, dorsal surface (process ‘f’ of
<bibRefCitation id="A566EBABFFC5FF8BFF43FBBEE875F962" author="Pfau" box="[151,267,1090,1115]" pageId="12" refString="Pfau, H. K. (2011). Functional morphology and evolution of the male secondary copulatory apparatus of the Anisoptera (Insecta: Odonata). Zoologica, 156 (1), 1 - 103." type="journal article" year="2011">Pfau 2011</bibRefCitation>
) with a rounded, slightly sclerotized distal process (
<figureCitation id="59CC8ADFFFC5FF8BFC8EFBBEEADCF962" box="[858,930,1090,1115]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
B, h-f) prolonged over joint between
<date id="B549B09AFFC5FF8BFA9DFBBFE9C7F946" pageId="12">V3 and V4</date>
(J
<date id="B549B09AFFC5FF8BFF08FB9BE85EF946" box="[220,288,1127,1151]" pageId="12">v3-v4</date>
), ventrally with a distinct proximal sclerite (
<figureCitation id="59CC8ADFFFC5FF8BFCC6FB9AEA27F946" box="[786,857,1126,1151]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
A, p-ps), and a membranous-like hood distally over
<date id="B549B09AFFC5FF8BFF43FB77E9C2F99A" box="[151,188,1163,1187]" pageId="12">V4</date>
, tapering into a tongue-like apex ventrally (
<figureCitation id="59CC8ADFFFC5FF8BFD10FB76EA71F99A" box="[708,783,1162,1187]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
A, “mh”);
<date id="B549B09AFFC5FF8BFC5BFB77EACFF99A" box="[911,945,1163,1187]" pageId="12">V4</date>
cornet-like, with a pair of lateroproximal sclerotized projections (
<figureCitation id="59CC8ADFFFC5FF8BFE72FB52E891F9FE" box="[422,495,1198,1223]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
A-B, “lsp”), curved anteriorly in lateral view (
<figureCitation id="59CC8ADFFFC5FF8BFBD5FB52ED38F9FE" box="[1025,1094,1198,1223]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Fig. 6</figureCitation>
A), pair of long distal flagella somewhat forcipate in ventral view (
<figureCitation id="59CC8ADFFFC5FF8BFDFBFB2EEB0CF9D2" box="[559,626,1234,1259]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Fig. 6</figureCitation>
B, d-fl), in lateral view as long as
<date id="B549B09AFFC5FF8BFC3FFB2FED70F9D2" box="[1003,1038,1235,1259]" pageId="12">V4</date>
length (
<figureCitation id="59CC8ADFFFC5FF8BFBBDFB2EEDD2F9D2" box="[1129,1196,1234,1259]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Fig. 6</figureCitation>
A). Cercus brownishyellow to dark brown to black, ventral branch light brown to dark-brown. Epiproct brown to almost black.
</paragraph>
<paragraph id="C148965AFFC5FF8BFF13FAE6E98CF8FA" blockId="12.[151,1437,151,2016]" pageId="12">
<emphasis id="F3834A48FFC5FF8BFF13FAE6E803F80A" bold="true" box="[199,381,1306,1331]" pageId="12">Measurements.</emphasis>
Abdomen length (excluding caudal appendages) 27.5–29.2; head maximum width 6.0 (specimen from Serra do Salitre not measured, eyes smashed); Fw length 23.4–25.0; Hw length 22.2–23.8; Fw maximum width 5.1–5.2, in Hw 6.4–6.7; pt length
<quantity id="060F3BBFFFC5FF8BFD33FA9EEA25F842" box="[743,859,1378,1403]" metricMagnitude="-2" metricUnit="m" metricValue="6.731" metricValueMax="6.858" metricValueMin="6.604" pageId="12" unit="in" value="2.65" valueMax="2.7" valueMin="2.6">2.6–2.7 in</quantity>
Fw,
<quantity id="060F3BBFFFC5FF8BFC43FA9EED75F842" box="[919,1035,1378,1403]" metricMagnitude="-2" metricUnit="m" metricValue="6.858" metricValueMax="7.111999999999999" metricValueMin="6.604" pageId="12" unit="in" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 in</quantity>
Hw; length of metathoracic femur 6.0–6.3; metathoracic tibia 4.6–4.8; length of S9+
<quantity id="060F3BBFFFC5FF8BFD0AFA7BEA62F8A6" box="[734,796,1414,1439]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="12" unit="in" value="10.0">10 in</quantity>
lateral view 2.8; total length of cercus in lateral view 1.2–1.3.
</paragraph>
<paragraph id="C148965AFFC5FF8BFF13FA32E8EAF8DE" blockId="12.[151,1437,151,2016]" box="[199,404,1486,1511]" pageId="12">
<emphasis id="F3834A48FFC5FF8BFF13FA32E869F8DE" bold="true" box="[199,279,1486,1511]" pageId="12">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFC5FF8AFF13FA0EED66FCE9" lastPageId="13" pageId="12" type="diagnosis">
<paragraph id="C148965AFFC5FF8BFF13FA0EEB3FFB86" blockId="12.[151,1437,151,2016]" pageId="12">
<emphasis id="F3834A48FFC5FF8BFF13FA0EE83FFB32" bold="true" box="[199,321,1522,1547]" pageId="12">Diagnosis.</emphasis>
The wide cornet-like distal portion of
<date id="B549B09AFFC5FF8BFD27FA0FEA6BFB32" box="[755,789,1523,1547]" pageId="12">V4</date>
(
<figureCitation id="59CC8ADFFFC5FF8BFCF0FA0EEA15FB32" box="[804,875,1522,1547]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
A–B), upturned tapered apex of cylindrical cercus (
<figureCitation id="59CC8ADFFFC5FF8BFF4BF9EAE998FB16" box="[159,230,1558,1583]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="12">Figs 7</figureCitation>
A–C, 8A–B), distal concavity from posterior 0.7 of epiproct of males (
<figureCitation id="59CC8ADFFFC5FF8BFBDBF9EAED2DFB16" box="[1039,1107,1558,1583]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="12">Fig. 8</figureCitation>
B) and short subgenital plate (ratio between length of plate and S9 &lt;0.4) with a wide concavity of females (
<figureCitation id="59CC8ADFFFC5FF8BFB50F9C6EDAAFB6A" box="[1156,1236,1594,1619]" captionStart="FIGURE 9" captionStartId="18.[151,250,1160,1182]" captionTargetBox="[151,1434,193,1138]" captionTargetId="figure@18.[151,1436,193,1139]" captionTargetPageId="18" captionText="FIGURE 9. Female S 8 – 10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A – B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C. Cyanogomphus angelomachadoi sp. nov. additional specimen (Brazil. MG: Serra do Salitre, ABMM); D. Cyanogomphus sp. (Brazil. GO: Jataí, ABMM); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm." httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="12">Figs 9</figureCitation>
C), will separate
<taxonomicName id="06F7EDD9FFC5FF8BFF43F9A2EB79FB4E" box="[151,519,1630,1655]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC5FF8BFF43F9A2EB79FB4E" box="[151,519,1630,1655]" italics="true" pageId="12">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC5FF8BFDD9F9A2EB19FB4E" bold="true" box="[525,615,1630,1655]" pageId="12">
<taxonomicNameLabel id="E8B0F733FFC5FF8BFDD9F9A2EB19FB4E" box="[525,615,1630,1655]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="06F7EDD9FFC5FF8BFD7FF9A2EA34FB4E" box="[683,842,1630,1655]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFC5FF8BFD7FF9A2EA34FB4E" box="[683,842,1630,1655]" italics="true" pageId="12">Tibiagomphus</emphasis>
</taxonomicName>
(
<date id="B549B09AFFC5FF8BFC8DF9A3EA02FB4E" box="[857,892,1631,1655]" pageId="12">V4</date>
distal portion narrow, apex of cercus posteriorly directed, epiproct dorsally smooth, lacking concavity, and female subgenital plate large and with a narrow mesal incision,
<figureCitation id="59CC8ADFFFC5FF8BFF29F95AE83CFB86" box="[253,322,1702,1727]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
F–G, 7J–
<collectingCountry id="B9E0D6CAFFC5FF8BFE7CF95BE8C3FB86" box="[424,445,1703,1727]" name="Iceland" pageId="12">O</collectingCountry>
, 8G–J, 9E).
</paragraph>
<paragraph id="C148965AFFC5FF8AFF13F936ED66FCE9" blockId="12.[151,1437,151,2016]" lastBlockId="13.[151,1437,151,1257]" lastPageId="13" pageId="12">
The dark mesal part of the mesepisternum close to the dorsal carina (
<figureCitation id="59CC8ADFFFC5FF8BFC2CF936ED40FBDA" box="[1016,1086,1738,1763]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="12">Fig. 1</figureCitation>
), epiproct with a larger distal concavity at posterior 0.70 (
<figureCitation id="59CC8ADFFFC5FF8BFE0BF912EB5DFA3E" box="[479,547,1774,1799]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="12">Fig. 8</figureCitation>
B) and dark abdomen with pale spots (
<figureCitation id="59CC8ADFFFC5FF8BFC38F912ED3DFA3E" box="[1004,1091,1774,1799]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="12">Figs 10</figureCitation>
A–B, 11A–B) distinguish this species from
<taxonomicName id="06F7EDD9FFC5FF8BFEF9F8EFE89CFA12" box="[301,482,1810,1835]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFC5FF8BFEF9F8EFE89CFA12" box="[301,482,1810,1835]" italics="true" pageId="12">C. comparabilis</emphasis>
</taxonomicName>
(mesal part of mesepisternum pale, epiproct with distal concavity reduced to a small posterodorsal area, abdomen almost uniform in color,
<figureCitation id="59CC8ADFFFC5FF8BFD23F8CAEA42FA76" box="[759,828,1846,1871]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="12">Figs 7</figureCitation>
E, 8F, 10C).
<taxonomicName id="06F7EDD9FFC5FF8BFC1FF8CAEC45FA76" box="[971,1339,1846,1871]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC5FF8BFC1FF8CAEC45FA76" box="[971,1339,1846,1871]" italics="true" pageId="12">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC5FF8BFA96F8CAECE2FA76" bold="true" box="[1346,1436,1846,1871]" pageId="12">
<taxonomicNameLabel id="E8B0F733FFC5FF8BFA96F8CAECE2FA76" box="[1346,1436,1846,1871]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar in general appearance to
<taxonomicName id="06F7EDD9FFC5FF8BFD83F8A7EBA1FA4A" box="[599,735,1882,1907]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC5FF8BFD83F8A7EBA1FA4A" box="[599,735,1882,1907]" italics="true" pageId="12">C. waltheri</emphasis>
</taxonomicName>
, with which the smallest specimens might be confused.
<taxonomicName id="06F7EDD9FFC5FF8BFF43F882EB75FAAE" box="[151,523,1918,1943]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC5FF8BFF43F882EB75FAAE" box="[151,523,1918,1943]" italics="true" pageId="12">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC5FF8BFDC1F882EB0CFAAE" bold="true" box="[533,626,1918,1943]" pageId="12">
<taxonomicNameLabel id="E8B0F733FFC5FF8BFDC1F882EB0CFAAE" box="[533,626,1918,1943]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by its smaller size (length of Hw 22.2–24.5 vs. 25.8–31.6 for
<taxonomicName id="06F7EDD9FFC5FF8BFE90F85FE8B7FA82" box="[324,457,1954,1979]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC5FF8BFE90F85FE8B7FA82" box="[324,457,1954,1979]" italics="true" pageId="12">C. waltheri</emphasis>
</taxonomicName>
); general paler color (
<figureCitation id="59CC8ADFFFC5FF8BFD34F85EEA45FA82" box="[736,827,1954,1979]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="12">Figs 10</figureCitation>
A–B, 11A–B), with large pale areas connected on mesepisternum (
<figureCitation id="59CC8ADFFFC5FF8BFE85F83AE8E8FAE6" box="[337,406,1990,2015]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="12">Figs 1</figureCitation>
A–B, E–F), except the
<typeStatus id="1E4C28F8FFC5FF8BFD77F834EA7AFAD9" box="[675,772,1992,2016]" pageId="12" type="paratype">paratype</typeStatus>
male from Serra do Salitre (
<figureCitation id="59CC8ADFFFC5FF8BFB99F83AEDDFFAE6" box="[1101,1185,1990,2015]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="12">Fig. 10</figureCitation>
B), which is distinctly darker with pale areas only partially connected and showing well-defined mesepisternal stripes limited by dark ground color (
<figureCitation id="59CC8ADFFFC4FF8AFEE2FF40E806FDED" box="[310,376,188,213]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="13">Figs 1</figureCitation>
C–D), much like some
<taxonomicName id="06F7EDD9FFC4FF8AFD55FF41EB81FDEC" box="[641,767,188,213]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC4FF8AFD55FF41EB81FDEC" box="[641,767,188,213]" italics="true" pageId="13">C. waltheri</emphasis>
</taxonomicName>
specimens (
<taxonomicName id="06F7EDD9FFC4FF8AFC58FF41ED74FDEC" box="[908,1034,188,213]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC4FF8AFC58FF41ED74FDEC" box="[908,1034,188,213]" italics="true" pageId="13">C. waltheri</emphasis>
</taxonomicName>
pale areas less broad, with first and second antehumeral stripes well defined); dorsal surface of metathoracic tibia yellow to brownish-yellow (
<figureCitation id="59CC8ADFFFC4FF8AFAEBFF23ECFEFDC1" box="[1343,1408,223,248]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="13">Fig. 3</figureCitation>
A, black in
<taxonomicName id="06F7EDD9FFC4FF8AFF2DFEF9E809FC24" box="[249,375,260,285]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC4FF8AFF2DFEF9E809FC24" box="[249,375,260,285]" italics="true" pageId="13">C. waltheri</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFC4FF8AFE52FEF8E8B6FC25" box="[390,456,260,285]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="13">Fig. 3</figureCitation>
B), distal concavity on epiproct larger (beginning at &lt;0.76 length of epiproct,
<figureCitation id="59CC8ADFFFC4FF8AFA94FEF8ECFCFC25" box="[1344,1410,260,285]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="13">Fig. 8</figureCitation>
B; while in
<taxonomicName id="06F7EDD9FFC4FF8AFED5FED4E8FDFC79" box="[257,387,295,320]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC4FF8AFED5FED4E8FDFC79" box="[257,387,295,320]" italics="true" pageId="13">C. waltheri</emphasis>
</taxonomicName>
at&gt; 0.80 length,
<figureCitation id="59CC8ADFFFC4FF8AFD88FEDBEBDFFC79" box="[604,673,295,320]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="13">Fig. 8</figureCitation>
D) with laterodistal projection in lateral view longer and slender, forefinger-shaped (
<figureCitation id="59CC8ADFFFC4FF8AFEBBFEB0E8C6FC5D" box="[367,440,332,357]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="13">Figs 7</figureCitation>
A, 8A; in
<taxonomicName id="06F7EDD9FFC4FF8AFDE4FEB1EBCFFC5C" box="[560,689,332,357]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC4FF8AFDE4FEB1EBCFFC5C" box="[560,689,332,357]" italics="true" pageId="13">C. waltheri</emphasis>
</taxonomicName>
epiproct laterodistal projection smaller and stout, thumb-shaped,
<figureCitation id="59CC8ADFFFC4FF8AFF43FE93E9A4FCB1" box="[151,218,367,392]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="13">Figs 7</figureCitation>
G, 8C); and cercus upright (
<figureCitation id="59CC8ADFFFC4FF8AFDDAFE93EB2AFCB1" box="[526,596,367,392]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="13">Figs 7</figureCitation>
A, 8A) will distinguish it from
<taxonomicName id="06F7EDD9FFC4FF8AFC7BFE8CED52FCB1" box="[943,1068,367,392]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC4FF8AFC7BFE8CED52FCB1" box="[943,1068,367,392]" italics="true" pageId="13">C. waltheri</emphasis>
</taxonomicName>
(cercus less curved upright,
<figureCitation id="59CC8ADFFFC4FF8AFABFFE93E9DAFC95" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="13">Figs 7</figureCitation>
G, 8C). Both species appear to be allopatric with
<taxonomicName id="06F7EDD9FFC4FF8AFD1BFE69EAD2FC94" box="[719,940,404,429]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFC4FF8AFD1BFE69EAD2FC94" box="[719,940,404,429]" italics="true" pageId="13">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFC4FF8AFC67FE68ED73FC94" bold="true" box="[947,1037,404,429]" pageId="13">
<taxonomicNameLabel id="E8B0F733FFC4FF8AFC67FE68ED73FC94" box="[947,1037,404,429]" pageId="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, thus far known as a representative of the Cerrado, and
<taxonomicName id="06F7EDD9FFC4FF8AFEA2FE44E88AFCE9" box="[374,500,439,464]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFC4FF8AFEA2FE44E88AFCE9" box="[374,500,439,464]" italics="true" pageId="13">C. waltheri</emphasis>
</taxonomicName>
, a representative of the Atlantic Forest (
<figureCitation id="59CC8ADFFFC4FF8AFC6CFE4BED75FCE9" box="[952,1035,439,464]" captionStart="FIGURE 12" captionStartId="24.[151,250,1240,1262]" captionTargetBox="[151,1436,215,1198]" captionTargetId="figure@24.[151,1436,195,1215]" captionTargetPageId="24" captionText="FIGURE 12. Map of southeastern South America with distribution records of Cyanogomphus Selys, 1873, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: MI, Misiones Province, Argentina; BA, Bahia; GO, Goiás; MG, Minas Gerais; MT, Mato Grosso; RJ, Rio de Janeiro; RS, Rio Grande do Sul; SC, Santa Catarina, and SP, São Paulo for Brazilian States." httpUri="https://zenodo.org/record/270865/files/figure.png" pageId="13">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFC4FF8AFF13FE20EC02FE64" pageId="13" type="distribution">
<paragraph id="C148965AFFC4FF8AFF13FE20ECE2FCCD" blockId="13.[151,1437,151,1257]" box="[199,1436,476,501]" pageId="13">
<emphasis id="F3834A48FFC4FF8AFF13FE20E81EFCCC" bold="true" box="[199,352,476,501]" pageId="13">Distribution.</emphasis>
States of Minas Gerais and São Paulo,
<collectingCountry id="B9E0D6CAFFC4FF8AFCC3FE20EA20FCCC" box="[791,862,476,501]" name="Brazil" pageId="13">Brazil</collectingCountry>
, in the Cerrado biogeographical province (
<figureCitation id="59CC8ADFFFC4FF8AFAE8FE20ECF1FCCD" box="[1340,1423,476,501]" captionStart="FIGURE 12" captionStartId="24.[151,250,1240,1262]" captionTargetBox="[151,1436,215,1198]" captionTargetId="figure@24.[151,1436,195,1215]" captionTargetPageId="24" captionText="FIGURE 12. Map of southeastern South America with distribution records of Cyanogomphus Selys, 1873, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: MI, Misiones Province, Argentina; BA, Bahia; GO, Goiás; MG, Minas Gerais; MT, Mato Grosso; RJ, Rio de Janeiro; RS, Rio Grande do Sul; SC, Santa Catarina, and SP, São Paulo for Brazilian States." httpUri="https://zenodo.org/record/270865/files/figure.png" pageId="13">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="C148965AFFC4FF8AFF13FE03EC02FE64" blockId="13.[151,1437,151,1257]" pageId="13">
<emphasis id="F3834A48FFC4FF8AFF13FE03EB41FF21" bold="true" box="[199,575,511,536]" pageId="13">Biological and ecological data.</emphasis>
Adults were collected at lotic systems, near rivers, streams, and creeks, surrounded by forests (riparian) in rupestrian formations over ironstone outcrops known as “canga”, at 664–766 meters in the core of Serra do Espinhaço mountain range (PNSC and Lagoa Santa sites) and at 1205 meters of elevation in the Serra da Canastra formation (Serra do Salitre site) in the Brazilian Cerrado domain. The pair from PNSC was caught using a Malaise trap and light sheet, non-traditional methods to capture dragonflies (
<bibRefCitation id="A566EBABFFC4FF8AFAC7FD6CE86DFFF5" author="De" pageId="13" refString="De Almeida, M. V. O., Pinto, A. P., Carvalho, A. L. &amp; Takiya, D. M. (2013) When rare is just a matter of sampling: Unexpected dominance of clubtail dragonflies (Odonata, Gomphidae) through different collecting methods at Parque Nacional da Serra do Cipo, Minas Gerais State, Brazil. Revista Brasileira de Entomologia, 57 (4), 417 - 423. http: // dx. doi. org / 10.1590 / S 0085 - 56262013005000042" type="journal article" year="2013">
<collectingCountry id="B9E0D6CAFFC4FF8AFAC7FD6CEC4BFF91" box="[1299,1333,656,680]" name="South Georgia and the South Sandwich Islands" pageId="13">De</collectingCountry>
Almeida
<emphasis id="F3834A48FFC4FF8AFF43FD49E9B1FFF4" box="[151,207,692,717]" italics="true" pageId="13">et al.</emphasis>
2013
</bibRefCitation>
). Despite more than
<quantity id="060F3BBFFFC4FF8AFE2FFD48EB2EFFF4" box="[507,592,692,717]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="13" unit="km" value="300.0">300 km</quantity>
between sites in PNSC and Lagoa Santa (Espinhaço mountain range) from that in Serra do Salitre (Canastra mountain range), they are similar environmentally, typical of the Brazilian Cerrado in that region; temperature, altitude, vegetation and rock outcrops are common to both regions. However, the
<typeStatus id="1E4C28F8FFC4FF8AFF17FCDCE85DFE01" box="[195,291,800,824]" pageId="13" type="paratype">paratype</typeStatus>
from São Paulo was collected at 485 meters in an apparently different environment, a sedimentary basin located in the north of Serra Geral rather than rupestrian formation where other specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFC4FF8AFF13FC9BE984FEF1" pageId="13" type="etymology">
<paragraph id="C148965AFFC4FF8AFF13FC9BE984FEF1" blockId="13.[151,1437,151,1257]" pageId="13">
<emphasis id="F3834A48FFC4FF8AFF13FC9BE830FEB9" bold="true" box="[199,334,871,896]" pageId="13">Etymology.</emphasis>
<taxonomicName id="06F7EDD9FFC4FF8AFE80FC9BEB71FEB9" box="[340,527,871,896]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="angelomachadoi">
<emphasis id="F3834A48FFC4FF8AFE80FC9BEB71FEB9" box="[340,527,871,896]" italics="true" pageId="13">angelomachadoi</emphasis>
</taxonomicName>
(noun phrase in genitive singular formed by a given and surname) specific name in honor of our friend and eminent Brazilian odonatologist Angelo B. M. Machado on the occasion of his 80th birthday.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFC4FF8AFF13FC28EAB8F9D1" pageId="13" type="discussion">
<paragraph id="C148965AFFC4FF8AFF13FC28EAB8F9D1" blockId="13.[151,1437,151,1257]" pageId="13">
<emphasis id="F3834A48FFC4FF8AFF13FC28E84AFED4" bold="true" box="[199,308,980,1005]" pageId="13">Remarks</emphasis>
. The only bona fide female from PNSC (DZRJ) is damaged, lacking S5–10 (
<figureCitation id="59CC8ADFFFC4FF8AFB42FC28ED9BFED5" box="[1174,1253,980,1005]" captionStart="FIGURE 11" captionStartId="23.[151,250,1163,1185]" captionTargetBox="[151,1436,193,1140]" captionTargetId="figure@23.[151,1436,193,1141]" captionTargetPageId="23" captionText="FIGURE 11. Female habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Paratype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448), B. Additional specimen (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); D. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270864/files/figure.png" pageId="13">Fig. 11</figureCitation>
A), precluding a full description, while the complete female collected at same site of
<typeStatus id="1E4C28F8FFC4FF8AFC43FC04EA89F929" box="[919,1015,1016,1040]" pageId="13" type="paratype">paratype</typeStatus>
male from Serra do Salitre (ABMM) is poorly preserved as to general color pattern (
<figureCitation id="59CC8ADFFFC4FF8AFD72FBE0EB89F90D" box="[678,759,1052,1077]" captionStart="FIGURE 11" captionStartId="23.[151,250,1163,1185]" captionTargetBox="[151,1436,193,1140]" captionTargetId="figure@23.[151,1436,193,1141]" captionTargetPageId="23" captionText="FIGURE 11. Female habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Paratype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448), B. Additional specimen (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); D. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270864/files/figure.png" pageId="13">Fig. 11</figureCitation>
B), leaving in some doubt an association with this species. It was for this reason excluded from the
<typeStatus id="1E4C28F8FFC4FF8AFDB5FBBCEBEFF961" box="[609,657,1088,1112]" pageId="13">type</typeStatus>
series. The Lagoa Santa
<typeStatus id="1E4C28F8FFC4FF8AFC14FBBCED5EF961" box="[960,1056,1088,1112]" pageId="13" type="paratype">paratype</typeStatus>
(MNRJ) is damaged, with head detached, synthorax crushed, and S1–6 varying from slightly to strongly damaged, with the left auricle broken. The male
<typeStatus id="1E4C28F8FFC4FF8AFF00FB75E84BF998" box="[212,309,1161,1185]" pageId="13" type="paratype">paratype</typeStatus>
from São Paulo State in RWG collection was not examined; however, it agrees fairly well with other specimens, showing the mesepisternal stripes divided as in the
<typeStatus id="1E4C28F8FFC4FF8AFCB9FB50EAB1F9FC" box="[877,975,1196,1221]" pageId="13" type="holotype">holotype</typeStatus>
(RWG pers. comm.). The females from Goiás (ABMM) may also be this species (see under
<taxonomicName id="06F7EDD9FFC4FF8AFD34FB33EAF0F9D1" box="[736,910,1231,1256]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFC4FF8AFD34FB33EAF0F9D1" box="[736,910,1231,1256]" italics="true" pageId="13">Cyanogomphus</emphasis>
</taxonomicName>
sp.).
</paragraph>
</subSubSection>
</treatment>
</document>

166
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<mods:title id="0F2233FAA217F066422CCEFCA24525CA">A taxonomic synopsis of South American Cyanogomphini Carle with description of Cyanogomphus angelomachadoi sp. nov. from the Cerrado of Brazil (Odonata: Gomphidae)</mods:title>
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<treatment id="495E274CFFD0FF9DFF43F96DED92FCCC" ID-DOI="http://doi.org/10.5281/zenodo.6077759" ID-GBIF-Taxon="119410162" ID-Zenodo-Dep="6077759" LSID="urn:lsid:plazi:treatment:495E274CFFD0FF9DFF43F96DED92FCCC" httpUri="http://treatment.plazi.org/id/495E274CFFD0FF9DFF43F96DED92FCCC" lastPageId="26" pageId="25">
<subSubSection id="89EDC5D1FFD0FF9EFF43F96DE862FBF5" pageId="25" type="nomenclature">
<paragraph id="C148965AFFD0FF9EFF43F96DEBB1FB92" blockId="25.[151,719,1681,1741]" box="[151,719,1681,1707]" pageId="25">
<heading id="9A002136FFD0FF9EFF43F96DEBB1FB92" bold="true" box="[151,719,1681,1707]" fontSize="11" level="1" pageId="25" reason="1">
<emphasis id="F3834A48FFD0FF9EFF43F96DEBB1FB92" bold="true" box="[151,719,1681,1707]" pageId="25">
<taxonomicName id="06F7EDD9FFD0FF9EFF43F96DE82AFB92" ID-CoL="3XNW" box="[151,340,1681,1707]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFD0FF9EFF43F96DE82AFB92" bold="true" box="[151,340,1681,1707]" italics="true" pageId="25">Cyanogomphus</emphasis>
</taxonomicName>
sp. (undetermined specimens)
</emphasis>
</heading>
</paragraph>
<paragraph id="C148965AFFD0FF9EFF43F948E862FBF5" blockId="25.[151,719,1681,1741]" box="[151,284,1716,1741]" pageId="25">
(
<figureCitation id="59CC8ADFFFD0FF9EFF4AF948E980FBF5" box="[158,254,1716,1741]" captionStart="FIGURE 9" captionStartId="18.[151,250,1160,1182]" captionTargetBox="[151,1434,193,1138]" captionTargetId="figure@18.[151,1436,193,1139]" captionTargetPageId="18" captionText="FIGURE 9. Female S 8 – 10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A – B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C. Cyanogomphus angelomachadoi sp. nov. additional specimen (Brazil. MG: Serra do Salitre, ABMM); D. Cyanogomphus sp. (Brazil. GO: Jataí, ABMM); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm." httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="25">Figure 9</figureCitation>
D)
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFD0FF9EFF43F900EDDBFA08" pageId="25" type="reference_group">
<paragraph id="C148965AFFD0FF9EFF43F900EDDBFA08" blockId="25.[151,1436,1788,1841]" pageId="25">
<emphasis id="F3834A48FFD0FF9EFF43F900E8C0FA28" box="[151,446,1788,1810]" italics="true" pageId="25">
<taxonomicName id="06F7EDD9FFD0FF9EFF43F900E8EFFA2B" ID-CoL="32NMT" box="[151,401,1788,1810]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="waltheri">Cyanogomphus waltheri</taxonomicName>
<emphasis id="F3834A48FFD0FF9EFE4EF900E8C0FA28" bold="true" box="[410,446,1788,1809]" italics="true" pageId="25">nec</emphasis>
</emphasis>
<bibRefCitation id="A566EBABFFD0FF9EFE13F900EB41FA2B" author="Selys" box="[455,575,1788,1810]" pageId="25" refString="Selys Longchamps, M. E. de (1873) Troisiemes additions au synopsis des Gomphines. Bulletins de l'Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, Serie 2, 35, 732 - 774." type="journal article" year="1873">Selys, 1873</bibRefCitation>
:—
<bibRefCitation id="A566EBABFFD0FF9EFD8AF900EC7FFA2B" author="Belle" box="[606,1281,1788,1810]" pageId="25" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">
Belle (1980: 151, in part, records of ♀ from Goiás State,
<collectingCountry id="B9E0D6CAFFD0FF9EFB68F900ED85FA2B" box="[1212,1275,1788,1810]" name="Brazil" pageId="25">Brazil</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD0FF9EFAF4F900EDDEFA08" author="Carvalho" pageId="25" refString="Carvalho, A. L. &amp; Nessimian, J. L. (1998) Odonata do Estado do Rio de Janeiro, Brasil: habitats e habitos das larvas. In: Nessimian, J. L. &amp; Carvalho, A. L. (Eds.), Ecologia de Insetos Aquaticos. Series Oecologia Brasiliensis. Vol. V. PPGE- UFRJ, Rio de Janeiro, pp. 3 - 28." type="book chapter" year="1998">
Carvalho &amp; Nessimian (1998: 8, 17, in part, notes on larval habitat and record to Mucuri, Bahia State,
<collectingCountry id="B9E0D6CAFFD0FF9EFB83F8E7EDE7FA08" box="[1111,1177,1819,1841]" name="Brazil" pageId="25">Brazil</collectingCountry>
)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFD0FF9EFF43F8A1EBC7FAA3" pageId="25" type="materials_examined">
<paragraph id="C148965AFFD0FF9EFF43F8A1EBC7FAA3" blockId="25.[151,1436,1885,2018]" pageId="25">
<emphasis id="F3834A48FFD0FF9EFF43F8A1E807FA4F" bold="true" box="[151,377,1885,1910]" pageId="25">Material examined</emphasis>
(2♀).
<collectingCountry id="B9E0D6CAFFD0FF9EFE10F8A2EB53FA4F" box="[452,557,1886,1910]" name="Brazil" pageId="25">BRAZIL</collectingCountry>
. Go[iás] State: 2♀, Jataí municipality [
<geoCoordinate id="A4C3F09DFFD0FF9EFC0BF8A1EDF2FA4F" box="[991,1164,1885,1910]" direction="south" orientation="latitude" pageId="25" precision="1" value="-17.88085">17°52’51.06”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFD0FF9EFB4CF8A1EC34FA4F" box="[1176,1354,1885,1910]" direction="west" orientation="longitude" pageId="25" precision="1" value="-51.714">51°42’50.40”W</geoCoordinate>
,
<quantity id="060F3BBFFFD0FF9EFA81F8A1ECE2FA4C" box="[1365,1436,1885,1910]" metricMagnitude="2" metricUnit="m" metricValue="7.05" pageId="25" unit="m" value="705.0">705 m</quantity>
a.s.l.],
<date id="B549B09AFFD0FF9EFF36F87DE853FAA3" box="[226,301,1921,1946]" pageId="25" value="1955-01">I.1955</date>
, [A.B.M. Machado leg.] (
<collectionCode id="A7E60E9FFFD0FF9EFD9AF87EEBD3FAA3" box="[590,685,1922,1946]" pageId="25">ABMM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFD0FF9DFF13F859ED92FCCC" lastPageId="26" pageId="25" type="discussion">
<paragraph id="C148965AFFD0FF9DFF13F859EDCDFC5D" blockId="25.[151,1436,1885,2018]" lastBlockId="26.[151,1436,151,501]" lastPageId="26" pageId="25">
<emphasis id="F3834A48FFD0FF9EFF13F859E84AFA87" bold="true" box="[199,308,1957,1982]" pageId="25">Remarks</emphasis>
. The females from the Brazilian Cerrado from the State of Goiás, cited by
<bibRefCitation id="A566EBABFFD0FF9EFB15F859EC2DFA87" author="Belle" box="[1217,1363,1957,1982]" pageId="25" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle (1980)</bibRefCitation>
as
<taxonomicName id="06F7EDD9FFD0FF9EFA56F85AE98AFADB" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFD0FF9EFA56F85AE98AFADB" italics="true" pageId="25">C. waltheri</emphasis>
</taxonomicName>
, are badly preserved, partially broken and moldy. We cannot identify these specimens, even although they are similar in size and color to
<taxonomicName id="06F7EDD9FFD3FF9DFDC0FF64EB8BFD89" box="[532,757,151,176]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="26" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFD3FF9DFDC0FF64EB8BFD89" box="[532,757,151,176]" italics="true" pageId="26">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFD3FF9DFCD5FF6BEA21FD89" bold="true" box="[769,863,151,176]" pageId="26">
<taxonomicNameLabel id="E8B0F733FFD3FF9DFCD5FF6BEA21FD89" box="[769,863,151,176]" pageId="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However, the female of
<taxonomicName id="06F7EDD9FFD3FF9DFB59FF64EC39FD89" box="[1165,1351,151,176]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="26" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFD3FF9DFB59FF64EC39FD89" box="[1165,1351,151,176]" italics="true" pageId="26">C. comparabilis</emphasis>
</taxonomicName>
is still unknown, and the only bona fide female of
<taxonomicName id="06F7EDD9FFD3FF9DFD56FF41EA20FDEC" box="[642,862,188,213]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="26" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFD3FF9DFD56FF41EA20FDEC" box="[642,862,188,213]" italics="true" pageId="26">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFD3FF9DFCB1FF40EAC0FDEC" bold="true" box="[869,958,188,213]" pageId="26">
<taxonomicNameLabel id="E8B0F733FFD3FF9DFCB1FF40EAC0FDEC" box="[869,958,188,213]" pageId="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is missing S5–10 (see under that species); both species also are known only from the Brazilian Cerrado. We are confident that they do not represent the Atlantic Forest
<taxonomicName id="06F7EDD9FFD3FF9DFE93FEF9E8BBFC24" box="[327,453,260,285]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="26" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFD3FF9DFE93FEF9E8BBFC24" box="[327,453,260,285]" italics="true" pageId="26">C. waltheri</emphasis>
</taxonomicName>
based on locality data, since they are also smaller than specimens from Rio de Janeiro State collected close to the
<typeStatus id="1E4C28F8FFD3FF9DFE05FED4EB7FFC79" box="[465,513,296,320]" pageId="26">type</typeStatus>
locality, as well as paler (pale dorsal surface of metathoracic tibia is black in
<taxonomicName id="06F7EDD9FFD3FF9DFA56FED4E98AFC5C" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="26" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFD3FF9DFA56FED4E98AFC5C" italics="true" pageId="26">C. waltheri</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFD3FF9DFED6FEB0E83DFC5D" box="[258,323,332,357]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="26">Fig. 3</figureCitation>
B). Images of the subgenital plates are given for future comparisons (
<figureCitation id="59CC8ADFFFD3FF9DFB98FEB0EDF1FC5D" box="[1100,1167,332,357]" captionStart="FIGURE 9" captionStartId="18.[151,250,1160,1182]" captionTargetBox="[151,1434,193,1138]" captionTargetId="figure@18.[151,1436,193,1139]" captionTargetPageId="18" captionText="FIGURE 9. Female S 8 – 10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A – B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C. Cyanogomphus angelomachadoi sp. nov. additional specimen (Brazil. MG: Serra do Salitre, ABMM); D. Cyanogomphus sp. (Brazil. GO: Jataí, ABMM); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm." httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="26">Fig. 9</figureCitation>
D).
</paragraph>
<paragraph id="C148965AFFD3FF9DFF13FE93ED92FCCC" blockId="26.[151,1436,151,501]" pageId="26">
<bibRefCitation id="A566EBABFFD3FF9DFF13FE93EB60FCB1" author="Carvalho" box="[199,542,367,392]" pageId="26" refString="Carvalho, A. L. &amp; Nessimian, J. L. (1998) Odonata do Estado do Rio de Janeiro, Brasil: habitats e habitos das larvas. In: Nessimian, J. L. &amp; Carvalho, A. L. (Eds.), Ecologia de Insetos Aquaticos. Series Oecologia Brasiliensis. Vol. V. PPGE- UFRJ, Rio de Janeiro, pp. 3 - 28." type="book chapter" year="1998">Carvalho &amp; Nessimian (1998)</bibRefCitation>
cited
<taxonomicName id="06F7EDD9FFD3FF9DFDB1FE8CEB9BFCB1" box="[613,741,367,392]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="26" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFD3FF9DFDB1FE8CEB9BFCB1" box="[613,741,367,392]" italics="true" pageId="26">C. waltheri</emphasis>
</taxonomicName>
from Bahia State,
<collectingCountry id="B9E0D6CAFFD3FF9DFC69FE93ED7AFCB1" box="[957,1028,367,392]" name="Brazil" pageId="26">Brazil</collectingCountry>
, based on collected larvae (see also remarks under
<taxonomicName id="06F7EDD9FFD3FF9DFE95FE69E8BEFC94" box="[321,448,404,429]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="26" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFD3FF9DFE95FE69E8BEFC94" box="[321,448,404,429]" italics="true" pageId="26">C. waltheri</emphasis>
</taxonomicName>
). The larva that may be
<taxonomicName id="06F7EDD9FFD3FF9DFD0DFE69EA27FC94" box="[729,857,404,429]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="26" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFD3FF9DFD0DFE69EA27FC94" box="[729,857,404,429]" italics="true" pageId="26">C. waltheri</emphasis>
</taxonomicName>
is relative common in southeastern
<collectingCountry id="B9E0D6CAFFD3FF9DFB27FE68EC49FC94" box="[1267,1335,404,429]" name="Brazil" pageId="26">Brazil</collectingCountry>
and was recently collected in another site in Bahia State (APP unpublished). However, since the F-0 larva of this species was described by supposition (see
<bibRefCitation id="A566EBABFFD3FF9DFDCFFE20EBE7FCCC" author="Belle" box="[539,665,476,501]" pageId="26" refString="Belle, J. (1996) Higher classification of the South-American Gomphidae (Odonata). Zoologische Mededelingen, 70, 297 - 324." type="journal article" year="1996">Belle 1996</bibRefCitation>
), the identity of these specimens needs confirmation.
</paragraph>
</subSubSection>
</treatment>
</document>

675
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<emphasis id="F3834A48FFD9FF97FF43FDB4E8C0FF5B" bold="true" box="[151,446,584,610]" italics="true" pageId="16">Cyanogomphus waltheri</emphasis>
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LSID urn:lsid:zoobank.org:act:7C774C12-B047-4CB0-9A29-5333E67F9CB8 (
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D, 3B, 4D, 6D–E, 7G–I, 8C–D, 9A–B, 10D, 11C, 12)
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</subSubSection>
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<emphasis id="F3834A48FFD9FF97FF43FD2EE8ECFFD1" box="[151,402,722,744]" italics="true" pageId="16">Cyanogomphus waltheri</emphasis>
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–756 (description of ♂
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,
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. Rio de Janeiro State: Rio de Janeiro municipality in IRSNB);—
<bibRefCitation id="A566EBABFFD9FF97FE3FFD0DEB04FE3E" author="Hagen" box="[491,634,753,775]" pageId="16" refString="Hagen, H. A. (1875) [1877] Synopsis of the Odonata of America. Proceedings of the Boston Society of Natural History, 18, 20 - 96." type="journal article" year="1875">Hagen (1875</bibRefCitation>
[1877]: 51, mention);—
<bibRefCitation id="A566EBABFFD9FF97FC5AFD0DEDC7FE3E" author="Kirby" box="[910,1209,753,775]" pageId="16" refString="Kirby, W. F. (1890) A synonymic catalogue of Neuroptera Odonata, or dragonflies, with an appendix of fossil species. Gurney &amp; Jackson, London, 202 pp." type="book" year="1890">Kirby (1890: 71, mention)</bibRefCitation>
;—
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Selys (1894: 173, comparison with
<taxonomicName id="06F7EDD9FFD9FF97FEA2FCEDEB7FFE1F" ID-CoL="32NMN" box="[374,513,784,806]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="species" species="demerarae">
<emphasis id="F3834A48FFD9FF97FEA2FCEDEB7FFE1F" box="[374,513,784,806]" italics="true" pageId="16">E. demerarae</emphasis>
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)
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;—
<treatmentCitation id="4056B04BFFD9FF97FDF1FCECEA6BFE1F" author="Williamson" box="[549,789,784,806]" page="167" pageId="16" year="1916">
<bibRefCitation id="A566EBABFFD9FF97FDF1FCECEB9DFE1F" author="Williamson" box="[549,739,784,806]" pageId="16" refString="Williamson, E. B. (1916) A new Cyanogomphus (Odonata). Entomological News, 27, 167 - 172." type="journal article" year="1916">Williamson (1916</bibRefCitation>
: 167
</treatmentCitation>
, 170, 172, pl. VIII, figs 2–5, comparison with
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<emphasis id="F3834A48FFD9FF97FB38FCECEC11FE1F" box="[1260,1391,784,806]" italics="true" pageId="16">Ebegomphus</emphasis>
</taxonomicName>
ssp. and
<taxonomicName id="06F7EDD9FFD9FF97FF26FCD3E828FE7C" ID-CoL="B7QB" box="[242,342,815,837]" class="Insecta" family="Gomphidae" genus="Agriogomphus" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="species" species="tumens">
<emphasis id="F3834A48FFD9FF97FF26FCD3E828FE7C" box="[242,342,815,837]" italics="true" pageId="16">A. tumens</emphasis>
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, photo of wings of ♂ from Teresópolis [likely in Rio de Janeiro State,
<collectingCountry id="B9E0D6CAFFD9FF97FBF0FCD2ED19FE7D" box="[1060,1127,814,836]" name="Brazil" pageId="16">Brazil</collectingCountry>
], secondary genitalia in lateral view and caudal appendages in dorsal and lateral views probably from the same specimen
<emphasis id="F3834A48FFD9FF97FB77FCB1EDC3FE5A" box="[1187,1213,845,867]" italics="true" pageId="16">cf.</emphasis>
<treatmentCitation id="4056B04BFFD9FF97FB13FCB1EC0DFE5A" author="Belle" box="[1223,1395,845,867]" page="151" pageId="16" year="1980">
<bibRefCitation id="A566EBABFFD9FF97FB13FCB1EC40FE5A" author="Belle" box="[1223,1342,845,867]" pageId="16" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle 1980</bibRefCitation>
: 151
</treatmentCitation>
);—
<treatmentCitation id="4056B04BFFD9FF97FF13FC90E8E1FEBB" author="Williamson" box="[199,415,876,898]" page="9" pageId="16" year="1918">
<bibRefCitation id="A566EBABFFD9FF97FF13FC90E8FAFEBB" author="Williamson" box="[199,388,876,898]" pageId="16" refString="Williamson, E. B. (1918) Results of the University of Michigan-Williamson Expedition to Colombia, 1916 - 1917. I. Two interesting new Colombian gomphines (Odonata). Occasional Papers of the Museum of Zoology University of Michigan, 52, 1 - 14, pls. I - II." type="journal article" year="1918">Williamson (1918</bibRefCitation>
: 9
</treatmentCitation>
, comparison with
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<emphasis id="F3834A48FFD9FF97FDB0FC90EBC5FEBB" box="[612,699,876,898]" italics="true" pageId="16">A. jessei</emphasis>
<bibRefCitation id="A566EBABFFD9FF97FD11FC90EA03FEBB" author="Williamson" box="[709,893,876,898]" pageId="16" refString="Williamson, E. B. (1918) Results of the University of Michigan-Williamson Expedition to Colombia, 1916 - 1917. I. Two interesting new Colombian gomphines (Odonata). Occasional Papers of the Museum of Zoology University of Michigan, 52, 1 - 14, pls. I - II." type="journal article" year="1918">Williamson, 1918</bibRefCitation>
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and
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<emphasis id="F3834A48FFD9FF97FC60FC91ED44FEBB" box="[948,1082,876,898]" italics="true" pageId="16">E. conchinus</emphasis>
</taxonomicName>
);—
<bibRefCitation id="A566EBABFFD9FF97FB8BFC90ED64FE86" author="Needham" pageId="16" refString="Needham, J. G. (1940) Studies on Neotropical Gomphine Dragonflies. Transactions of the American Entomological Society, 65 (4), 363 - 394." type="journal article" year="1940">
Needham (1940: 382–384, pl. XXI, figs 27–29, description of F-0 larva by supposition, illustrations of habitus and labium in dorsal view, and dorsal margin of abdomen in lateral view, records from Santa Catarina State,
<collectingCountry id="B9E0D6CAFFD9FF97FC01FC55ED6DFE86" box="[981,1043,937,959]" name="Brazil" pageId="16">Brazil</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FBECFC55E87BFEE7" author="Needham" pageId="16" refString="Needham, J. G. (1944) Further Studies on Neotropical Gomphine Dragonflies (Odonata). Transactions of the American Entomological Society, 69 (4), 171 - 224." type="journal article" year="1944">Needham (1944: 184, taxonomic notes)</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FEF0FC34EC37FEE7" author="Belle" box="[292,1353,968,990]" pageId="16" refString="Belle, J. (1966) Surinam dragon-flies of the Agriogomphus complex of genera. Studies on the fauna of Suriname and other Guyanas, 8 (29), 29 - 60." type="journal article" year="1966">
Belle (1966: 29, 31, 45, 47, 59, comparison with
<taxonomicName id="06F7EDD9FFD9FF97FCF2FC35EAF1FEE7" ID-CoL="7BYMN" box="[806,911,969,990]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD9FF97FCF2FC35EAF1FEE7" box="[806,911,969,990]" italics="true" pageId="16">T. uncatus</emphasis>
</taxonomicName>
, notes on larva, venation and distribution)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FAB3FC34E83BF922" author="Belle" pageId="16" refString="Belle, J. (1970) Studies on South American Gomphidae (Odonata) with special reference to the species from Surinam. Studies on the fauna of Suriname and other Guyanas, 11 (43), 1 - 158, pls. XIX." type="journal article" year="1970" yearSuffix="a">
Belle (1970: 22–23, pl. IIIa, comparison with
<taxonomicName id="06F7EDD9FFD9FF97FD8AFC1AEB9FFEC5" ID-CoL="9WVZC" box="[606,737,998,1020]" class="Insecta" family="Gomphidae" genus="Ebegomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFD9FF97FD8AFC1AEB9FFEC5" box="[606,737,998,1020]" italics="true" pageId="16">Ebegomphus</emphasis>
</taxonomicName>
spp., mention of a second male in the IRSNB, photo of the wings of ♂
<typeStatus id="1E4C28F8FFD9FF97FF35FBF9E841F922" box="[225,319,1029,1051]" pageId="16" type="holotype">holotype</typeStatus>
)
</bibRefCitation>
;—St.
<bibRefCitation id="A566EBABFFD9FF97FE50FBF9EBC8F922" author="Quentin" box="[388,694,1029,1051]" pageId="16" refString="St. Quentin, D. (1973) Die Gomphidenfauna Sudamerikas (Ordn.: Odonata). Annalen des Naturhistorischen Museums in Wien, 77, 335 - 363." type="journal article" year="1973">Quentin (1973: 358, mention)</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FD00FBF9EB15F94E" author="Belle" pageId="16" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">
Belle (1980: 151–152, 155, figs 1–4, in part, notes on ♂
<typeStatus id="1E4C28F8FFD9FF97FACCFBF9EC0EF922" box="[1304,1392,1029,1051]" pageId="16" type="holotype">holotype</typeStatus>
and illustrations of synthorax, metathoracic tibia and caudal appendages, description and illustration of subgenital plate in ventral view of ♀ from Santa Catarina State,
<collectingCountry id="B9E0D6CAFFD9FF97FD4DFBBEEBA7F961" box="[665,729,1090,1112]" name="Brazil" pageId="16">Brazil</collectingCountry>
, records from
<collectingCountry id="B9E0D6CAFFD9FF97FCBBFBBEEAA9F961" box="[879,983,1090,1112]" name="Argentina" pageId="16">Argentina</collectingCountry>
, Misiones Province, and
<collectingCountry id="B9E0D6CAFFD9FF97FB0EFBBEEC64F961" box="[1242,1306,1090,1112]" name="Brazil" pageId="16">Brazil</collectingCountry>
, Rio Grande do Sul State, comparison with
<taxonomicName id="06F7EDD9FFD9FF97FE28FB9EEB1AF94E" ID-CoL="7BYMN" box="[508,612,1122,1143]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD9FF97FE28FB9EEB1AF94E" box="[508,612,1122,1143]" italics="true" pageId="16">T. uncatus</emphasis>
</taxonomicName>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FD5DFB9DE848F9AF" author="Jurzitza" pageId="16" refString="Jurzitza, G. (1981) Lista provicional de los odonatos del Parque Nacional Iguazu, Provincia de Misiones, Republica Argentina. Notulae Odonatologicae, 1 (7), 117 - 118." type="journal article" year="1981">
Jurzitza (1981: 117, record from Parque Nacional Iguazú, Misiones Province,
<collectingCountry id="B9E0D6CAFFD9FF97FF13FB7CE851F9AF" box="[199,303,1152,1174]" name="Argentina" pageId="16">Argentina</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FE80FB7CEBB4F9AF" author="Davies" box="[340,714,1152,1174]" pageId="16" refString="Davies, D. A. L. &amp; Tobin, P. (1985) The dragonflies of the world: a systematic list of the extant species of Odonata. Vol. 2. Anisoptera. Societas Internationalis Odonatologica, Rapid Communications, 5 (Supplement), 1 - 151." type="journal article" year="1985">Davies &amp; Tobin (1985: 26, mention)</bibRefCitation>
;—
<treatmentCitation id="4056B04BFFD9FF97FD3CFB7CED50F9AF" author="Watson" box="[744,1070,1152,1174]" page="501" pageId="16" year="1985">
<bibRefCitation id="A566EBABFFD9FF97FD3CFB7CEA82F9AF" author="Watson" box="[744,1020,1152,1174]" pageId="16" refString="Watson, J. A. L. &amp; O'Farrell, A. F. (1985) The affinities of the Australian Gomphidae (Odonata). Australian Journal Of Zoology, 33 (4), 499 - 508. http: // dx. doi. org / 10.1071 / ZO 9850499" type="journal article" year="1985">Watson &amp; O’Farrell (1985</bibRefCitation>
: 501
</treatmentCitation>
, mention);—
<bibRefCitation id="A566EBABFFD9FF97FB61FB7CEBD4F98D" author="Carle" pageId="16" refString="Carle, F. L. (1986) The classification, phylogeny and biogeography of the Gomphidae (Anisoptera). I. Classification. Odonatologica, 15 (3), 275 - 326." type="journal article" year="1986">
Carle (1986: 307–308, classification and comparison with
<taxonomicName id="06F7EDD9FFD9FF97FDEDFB5CEBDDF98C" ID-CoL="7BYMN" box="[569,675,1184,1205]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD9FF97FDEDFB5CEBDDF98C" box="[569,675,1184,1205]" italics="true" pageId="16">T. uncatus</emphasis>
</taxonomicName>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FD1DFB62ED54F98D" author="Santos" box="[713,1066,1182,1204]" pageId="16" refString="Santos, N. D. (1988) Catalogo Bibliografico de Ninfas de odonatos neotropicais. (Acompanhado de relacao alfabetica de autores e seus trabalhos). Acta Amazonica, 18 (1 - 2), 265 - 350." type="journal article" year="1988">Santos (1988: 267, 286, mention)</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FB9CFB62EC03F98D" author="Garrison" box="[1096,1405,1182,1204]" pageId="16" refString="Garrison, R. W. (1991) A synonymic list of the New World Odonata. Argia, 3 (2), 1 - 30." type="journal article" year="1991">Garrison (1991: 17, mention)</bibRefCitation>
;—
<treatmentCitation id="4056B04BFFD9FF97FF13FB41E882F9EA" author="Rodrigues" box="[199,508,1213,1235]" page="58" pageId="16" year="1992">
<bibRefCitation id="A566EBABFFD9FF97FF13FB41E8ABF9EA" author="Rodrigues" box="[199,469,1213,1235]" pageId="16" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">
Rodrigues
<bibRefCitation id="A566EBABFFD9FF97FEE3FB41E8ABF9EA" author="Capitulo" box="[311,469,1213,1235]" pageId="16" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">Capítulo (1992</bibRefCitation>
</bibRefCitation>
: 58
</treatmentCitation>
; record from Misiones Province,
<collectingCountry id="B9E0D6CAFFD9FF97FC83FB41EABEF9EA" box="[855,960,1213,1235]" name="Argentina" pageId="16">Argentina</collectingCountry>
);—
<bibRefCitation id="A566EBABFFD9FF97FC31FB41ED80F9EA" author="Belle" box="[997,1278,1213,1235]" pageId="16" refString="Belle, J. (1993) Annotated checklist and bibliography of the immature stages of Neotropical Gomphidae, published up to September 1993 (Anisoptera). Odonatologica, 22 (4), 399 - 409." type="journal article" year="1993">Belle (1993: 401, mention)</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FAC8FB41EB0DF9CB" author="Belle" pageId="16" refString="Belle, J. (1994) Three new Neotropical Gomphidae from the genera Archaeogomphus Williamson, Cyanogomphus Selys and Epigomphus Hagen (Anisoptera). Odonatologica, 23 (1), 45 - 50." type="journal article" year="1994">
Belle (1994: 47, 49, comparison with
<taxonomicName id="06F7EDD9FFD9FF97FE13FB20EB12F9CB" ID-CoL="32NML" box="[455,620,1244,1266]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFD9FF97FE13FB20EB12F9CB" box="[455,620,1244,1266]" italics="true" pageId="16">C. comparabilis</emphasis>
</taxonomicName>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FD46FB20EDB4F9CB" author="Bridges" box="[658,1226,1244,1266]" pageId="16" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Odonata of the world (Third Edition). Privately published, Urbana, Illinois, 905 pp." type="book" year="1994">
Bridges (1994:
<date id="B549B09AFFD9FF97FCE0FB20EA0EF9CB" box="[820,880,1244,1266]" pageId="16" value="2013-03">III.13</date>
, VI.9, VII.251,
<date id="B549B09AFFD9FF97FBC0FB20ED1CF9CB" box="[1044,1122,1244,1266]" pageId="16" value="1918-08">VIII.18</date>
, mention)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FB3DFB20EA3FF829" author="Belle" pageId="16" refString="Belle, J. (1996) Higher classification of the South-American Gomphidae (Odonata). Zoologische Mededelingen, 70, 297 - 324." type="journal article" year="1996">
Belle (1996: 298, 314, comparison with
<taxonomicName id="06F7EDD9FFD9FF97FE7BFB07EB2DF829" ID-CoL="32NML" box="[431,595,1274,1296]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFD9FF97FE7BFB07EB2DF829" box="[431,595,1274,1296]" italics="true" pageId="16">C. comparabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="06F7EDD9FFD9FF97FD5CFB06EA75F829" ID-CoL="9WVZC" box="[648,779,1274,1296]" class="Insecta" family="Gomphidae" genus="Ebegomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="16" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFD9FF97FD5CFB06EA75F829" box="[648,779,1274,1296]" italics="true" pageId="16">Ebegomphus</emphasis>
</taxonomicName>
spp.)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFD9FF97FCB4FB06ED99F829" author="Steinmann" box="[864,1255,1274,1296]" pageId="16" refString="Steinmann, H. (1997) World catalogue of Odonata, Volume II Anisoptera. Walter de Gruyter, Berlin, 636 pp." type="book" year="1997">Steinmann (1997: 103–104, mention)</bibRefCitation>
;—
<treatmentCitation id="4056B04BFFD9FF97FAD1FB06E854F816" author="Garrison" page="68" pageId="16" year="2006">
<bibRefCitation id="A566EBABFFD9FF97FAD1FB06E878F816" author="Garrison" pageId="16" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
Garrison
<emphasis id="F3834A48FFD9FF97FAB3FB07ECE3F829" box="[1383,1437,1274,1296]" italics="true" pageId="16">et al.</emphasis>
(2006
</bibRefCitation>
: 68
</treatmentCitation>
–71, 77, 79, 81, 83–84, 96–97, 328–331, 333; figs 365, 379, 370, 383, 392, 454, 474, 485, 510, 519, 585–590, illustrations of head in dorsal view, wings, metathoracic tibia and anterior and posterior hamules in lateral view of a male from São Paulo State, caudal appendages in lateral view from a male of Rio Grande do Sul State, subgenital plate from Santa Catarina State, all from
<collectingCountry id="B9E0D6CAFFD9FF97FDD0FA89EB3CF8B2" box="[516,578,1397,1419]" name="Brazil" pageId="16">Brazil</collectingCountry>
);—
<treatmentCitation id="4056B04BFFD9FF97FDB3FA89EA38F8B2" author="Heckman" box="[615,838,1397,1419]" page="643" pageId="16" year="2006">
<bibRefCitation id="A566EBABFFD9FF97FDB3FA89EA6DF8B2" author="Heckman" box="[615,787,1397,1419]" pageId="16" refString="Heckman, C. W. (2006) Encyclopedia of South American aquatic insects: Odonata Anisoptera. Illustrated keys to known families, genera, and species in South America. Springer, Dordrecht, viii + 725 pp." type="book" year="2006">Heckman (2006</bibRefCitation>
: 643
</treatmentCitation>
–644, 647, figs 3.2.770, 3.2.774, key, reproduction of the illustrations from
<bibRefCitation id="A566EBABFFD9FF97FEA9FA68EB69F893" author="Needham" box="[381,535,1428,1450]" pageId="16" refString="Needham, J. G. (1940) Studies on Neotropical Gomphine Dragonflies. Transactions of the American Entomological Society, 65 (4), 363 - 394." type="journal article" year="1940">Needham 1940</bibRefCitation>
and
<bibRefCitation id="A566EBABFFD9FF97FD9CFA68EBC9F893" author="Belle" box="[584,695,1428,1450]" pageId="16" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle 1980</bibRefCitation>
);—von
<bibRefCitation id="A566EBABFFD9FF97FCD8FA68E848F8F1" author="Ellenrieder" pageId="16" refString="von Ellenrieder, N. &amp; Muzon, J. (2008) An updated checklist of the Odonata from Argentina. Odonatologica, 37 (1), 55 - 68." type="journal article" year="2008">
Ellenrieder &amp; Muzón (2008: 61, record from Misiones Province,
<collectingCountry id="B9E0D6CAFFD9FF97FF13FA4EE851F8F1" box="[199,303,1458,1480]" name="Argentina" pageId="16">Argentina</collectingCountry>
)
</bibRefCitation>
;—von
<treatmentCitation id="4056B04BFFD9FF97FE55FA4EEB9EF8F1" author="Ellenrieder" box="[385,736,1458,1480]" page="123" pageId="16" year="2009">
<bibRefCitation id="A566EBABFFD9FF97FE55FA4EEBD1F8F1" author="Ellenrieder" box="[385,687,1458,1480]" pageId="16" refString="von Ellenrieder, N. &amp; Garrison, R. W. (2009) Odonata. In: Dominguez, E. &amp; Fernandez, H. R. (Eds.), Macroinvertebrados bentonicos sudamericanos. Sistematica y biologia. Fundacion Miguel Lillo, Tucuman, pp. 95 - 143." type="book chapter" year="2009">Ellenrieder &amp; Garrison (2009</bibRefCitation>
: 123
</treatmentCitation>
, figs 276, 282, reproduction of illustrations of ♀ subgenital plate and caudal appendages of ♂ from
<bibRefCitation id="A566EBABFFD9FF97FE2DFA2DEBBAF8DE" author="Garrison" box="[505,708,1489,1511]" pageId="16" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
Garrison
<emphasis id="F3834A48FFD9FF97FD8CFA2EEBF2F8DE" box="[600,652,1489,1511]" italics="true" pageId="16">et al.</emphasis>
2006
</bibRefCitation>
);—Hämäläinen (2013: 30–31, fig. 2, mention and photo of habitus of a male in the field from
<collectingCountry id="B9E0D6CAFFD9FF97FE7EFA0CEB6FFB3F" box="[426,529,1520,1542]" name="Argentina" pageId="16">Argentina</collectingCountry>
);—
<bibRefCitation id="A566EBABFFD9FF97FDE1FA0CEAD5FB3F" author="Hamalainen" box="[565,939,1520,1542]" pageId="16" refString="Hamalainen, M. (2015) Catalogue of individuals commemorated in the scientific names of extant dragonflies, including lists of all available eponymous species-group and genus-group names. International Dragonfly Fund Report, 80, 1 - 168." type="journal article" year="2015">Hämäläinen (2015: 158, etymology)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFD9FF95FF43F9CEEB00F844" lastPageId="18" pageId="16" type="materials_examined">
<paragraph id="C148965AFFD9FF97FF43F9CEEC4DFAEE" blockId="16.[151,1437,1586,2007]" pageId="16">
<emphasis id="F3834A48FFD9FF97FF43F9CEE88BFB72" bold="true" box="[151,501,1586,1611]" pageId="16">
Material examined (
<specimenCount id="D7F15DD3FFD9FF97FE5AF9CEE8CFFB72" box="[398,433,1586,1611]" pageId="16" type="generic">7♂</specimenCount>
, 2♀).
</emphasis>
<collectingCountry id="B9E0D6CAFFD9FF97FE2AF9CFEB19FB72" box="[510,615,1587,1611]" name="Brazil" pageId="16">BRAZIL</collectingCountry>
. [Minas Gerais] State:
<specimenCount id="D7F15DD3FFD9FF97FCA6F9CFEAE8FB72" box="[882,918,1587,1611]" pageId="16" type="generic">1♂</specimenCount>
, [Santa Bárbara/São Gonçalo do Rio Abaixo municipalities, Unidade Ambiental] Peti [
<collectionCode id="A7E60E9FFFD9FF97FDA0F9ABEBAFFB56" box="[628,721,1623,1647]" pageId="16">CEMIG</collectionCode>
,
<geoCoordinate id="A4C3F09DFFD9FF97FD34F9AAEAF7FB56" box="[736,905,1622,1647]" direction="south" orientation="latitude" pageId="16" precision="1" value="-19.885975">19°53’09.51”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFD9FF97FC4CF9AAED34FB56" box="[920,1098,1622,1647]" direction="west" orientation="longitude" pageId="16" precision="1" value="-43.370907">43°22’15.27”W</geoCoordinate>
,
<quantity id="060F3BBFFFD9FF97FB8DF9ABEDDDFB57" box="[1113,1187,1623,1647]" metricMagnitude="2" metricUnit="m" metricValue="7.2299999999999995" pageId="16" unit="m" value="723.0">723 m</quantity>
a.s.l.], road from left margin, II.[19]88, [A.B.M. Machado leg.?] (
<collectionCode id="A7E60E9FFFD9FF97FD72F987EA7BFBAA" box="[678,773,1659,1683]" pageId="16">ABMM</collectionCode>
);
<specimenCount id="D7F15DD3FFD9FF97FCCBF987EA3DFBAA" box="[799,835,1659,1683]" pageId="16" type="generic">1♂</specimenCount>
, S[ão] João Del Rei municipality [
<geoCoordinate id="A4C3F09DFFD9FF97FB3FF986ECE9FBAA" box="[1259,1431,1658,1683]" direction="south" orientation="latitude" pageId="16" precision="1" value="-21.136">21°08’09.60”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFD9FF97FF43F962E837FB8E" box="[151,329,1694,1719]" direction="west" orientation="longitude" pageId="16" precision="1" value="-44.26195">44°15’43.02”W</geoCoordinate>
,
<quantity id="060F3BBFFFD9FF97FE80F963E8E5FB8F" box="[340,411,1695,1719]" metricMagnitude="2" metricUnit="m" metricValue="9.14" pageId="16" unit="m" value="914.0">914 m</quantity>
a.s.l.], IV[?].[19]61, Jandico leg. (
<collectionCode id="A7E60E9FFFD9FF97FCC9F963EA10FB8E" box="[797,878,1695,1719]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="16">MNRJ</collectionCode>
); Rio de Janeiro State:
<specimenCount id="D7F15DD3FFD9FF97FBBAF963EDEDFB8E" box="[1134,1171,1695,1719]" pageId="16" type="generic">1♂</specimenCount>
, Cachoeiras de Macacu municipality, [Boca do Mato district, Represa do] Valério [
<geoCoordinate id="A4C3F09DFFD9FF97FCFFF93EEAA9FBE2" box="[811,983,1730,1755]" direction="south" orientation="latitude" pageId="16" precision="1" value="-22.41811">22°25’05.20”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFD9FF97FC37F93FED0CFBE2" box="[995,1138,1731,1755]" direction="west" orientation="longitude" pageId="16" precision="15" value="-42.621387">42°37’17”W</geoCoordinate>
,
<quantity id="060F3BBFFFD9FF97FBAAF93EEDBBFBE3" box="[1150,1221,1730,1755]" metricMagnitude="2" metricUnit="m" metricValue="2.36" pageId="16" unit="m" value="236.0">236 m</quantity>
a.s.l.],
<date id="B549B09AFFD9FF97FACCF93FEC34FBE2" box="[1304,1354,1731,1755]" pageId="16">12.I.</date>
[20]11, N. Tangerini leg. (
<collectionCode id="A7E60E9FFFD9FF97FEB0F91BE8D7FBC6" box="[356,425,1767,1791]" pageId="16">DZRJ</collectionCode>
2493); 1♀, same data but
<date id="B549B09AFFD9FF97FD07F91AEA63FBC6" box="[723,797,1766,1791]" pageId="16">26.XI.</date>
[20]11 (
<collectionCode id="A7E60E9FFFD9FF97FCA7F91BEAC6FBC6" box="[883,952,1767,1791]" pageId="16">DZRJ</collectionCode>
2494); 1♀, same municipality but Reserva Ecológica de Guapiaçu (
<collectionCode id="A7E60E9FFFD9FF97FE68F8F7EB5CFA1A" box="[444,546,1803,1827]" pageId="16">REGUA</collectionCode>
), Forest Fragment (
<geoCoordinate id="A4C3F09DFFD9FF97FCC4F8F7EAD3FA1A" box="[784,941,1803,1827]" direction="south" orientation="latitude" pageId="16" precision="1" value="-22.467001">22°28’01.2”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFD9FF97FC6BF8F6ED0FFA1A" box="[959,1137,1802,1827]" direction="west" orientation="longitude" pageId="16" precision="1" value="-42.754">42°45’14.40”W</geoCoordinate>
,
<quantity id="060F3BBFFFD9FF97FB57F8F6EDBCFA1B" box="[1155,1218,1802,1827]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="16" unit="m" value="46.0">46 m</quantity>
a.s.l.),
<date id="B549B09AFFD9FF97FAF5F8F6ECE6FA1A" box="[1313,1432,1802,1827]" pageId="16" value="2012-02-15">15.II.2012</date>
, T.M.F. Kompier leg. (
<collectionCode id="A7E60E9FFFD9FF97FE5FF8D3E8AEFA7E" box="[395,464,1839,1863]" pageId="16">DZRJ</collectionCode>
2495);
<specimenCount id="D7F15DD3FFD9FF97FDF3F8D3EB35FA7E" box="[551,587,1839,1863]" pageId="16" type="generic">1♂</specimenCount>
, same data but
<date id="B549B09AFFD9FF97FD2EF8D2EA08FA7E" box="[762,886,1838,1863]" pageId="16" value="2012-04-26">26.IV.2012</date>
(
<collectionCode id="A7E60E9FFFD9FF97FC50F8D3EAB7FA7E" box="[900,969,1839,1863]" pageId="16">DZRJ</collectionCode>
2496);
<specimenCount id="D7F15DD3FFD9FF97FBF5F8D3ED3BFA7E" box="[1057,1093,1839,1863]" pageId="16" type="generic">1♂</specimenCount>
, district of Santana de Japuíba [
<geoCoordinate id="A4C3F09DFFD9FF97FF4AF8AEE858FA52" box="[158,294,1874,1899]" direction="south" orientation="latitude" pageId="16" precision="15" value="-22.562777">22°33’46”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFD9FF97FEE0F8AFE8BDFA52" box="[308,451,1875,1899]" direction="west" orientation="longitude" pageId="16" precision="15" value="-42.69167">42°41’30”W</geoCoordinate>
,
<quantity id="060F3BBFFFD9FF97FE04F8AFEB74FA53" box="[464,522,1875,1899]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="16" unit="m" value="27.0">27 m</quantity>
a.s.l.], Rio São João,
<date id="B549B09AFFD9FF97FCD0F8AFEAF1FA52" box="[772,911,1875,1899]" pageId="16" value="2000-12-08">08.XII.2000</date>
, J.M. Costa, R. Garrison, A.N. Lourenço &amp; P. Vieira leg. (
<collectionCode id="A7E60E9FFFD9FF97FEFCF88BE807FAB6" box="[296,377,1911,1935]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="16">MNRJ</collectionCode>
); Rio Grande do Sul State:
<specimenCount id="D7F15DD3FFD9FF97FD01F88BEB84FAB6" box="[725,762,1911,1935]" pageId="16" type="generic">1♂</specimenCount>
, S[an]to Antônio da Patrulha municipality [
<geoCoordinate id="A4C3F09DFFD9FF97FADBF88AECE6FAB6" box="[1295,1432,1910,1935]" direction="south" orientation="latitude" pageId="16" precision="15" value="-14.16">14°09’36”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFD9FF97FF43F866E858FA8A" box="[151,294,1946,1971]" direction="west" orientation="longitude" pageId="16" precision="15" value="-56.2">56°12’00”W</geoCoordinate>
,
<quantity id="060F3BBFFFD9FF97FEE1F867E801FA8B" box="[309,383,1947,1971]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="16" unit="m" value="123.0">123 m</quantity>
a.s.l.],
<date id="B549B09AFFD9FF97FE03F866EB61FA8A" box="[471,543,1946,1971]" pageId="16">26.III.</date>
[19]88, Gelson [Fiorentin] leg. (
<collectionCode id="A7E60E9FFFD9FF97FC44F867EA91FA8A" box="[912,1007,1947,1971]" pageId="16">ABMM</collectionCode>
);
<specimenCount id="D7F15DD3FFD9FF97FBD1F867ED54FA8A" box="[1029,1066,1947,1971]" pageId="16" type="generic">1♂</specimenCount>
, without locality [very likely in southeastern
<collectingCountry id="B9E0D6CAFFD9FF97FEFEF842E80AFAEE" box="[298,372,1982,2007]" name="Brazil" pageId="16">Brazil</collectingCountry>
], 4 h a.m., light trap, staring night,
<date id="B549B09AFFD9FF97FD28F843EA2FFAEE" box="[764,849,1983,2007]" pageId="16">02.XII.</date>
[19]78, [A.B.M. Machado leg.?] (
<collectionCode id="A7E60E9FFFD9FF97FB1CF843EC59FAEE" box="[1224,1319,1983,2007]" pageId="16">ABMM</collectionCode>
).
</paragraph>
<caption id="9588C6D2FFD8FF96FF43F8B8E825FACA" httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="17" targetBox="[151,1435,197,1827]" targetPageId="17">
<paragraph id="C148965AFFD8FF96FF43F8B8E825FACA" blockId="17.[151,1437,1860,2035]" pageId="17">
<emphasis id="F3834A48FFD8FF96FF43F8B8E870FA60" bold="true" box="[151,270,1860,1882]" pageId="17">FIGURE 8</emphasis>
. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A–B.
<taxonomicName id="06F7EDD9FFD8FF96FE77F89EEB8EFA41" box="[419,752,1890,1912]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="17" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFD8FF96FE77F89EEB8EFA41" box="[419,752,1890,1912]" italics="true" pageId="17">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFD8FF96FD21F89FEA3BFA41" bold="true" box="[757,837,1891,1912]" pageId="17">
<taxonomicNameLabel id="E8B0F733FFD8FF96FD21F89FEA3BFA41" box="[757,837,1891,1912]" pageId="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C–D.
<taxonomicName id="06F7EDD9FFD8FF96FE8AF87DEBADFAAE" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" box="[350,723,1921,1943]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="17" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFD8FF96FE8AF87DEB28FAAE" box="[350,598,1921,1943]" italics="true" pageId="17">Cyanogomphus waltheri</emphasis>
Selys, 1873
</taxonomicName>
(Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E–F.
<taxonomicName id="06F7EDD9FFD8FF96FF43F85CEB43FA8F" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[151,573,1952,1974]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="17" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFD8FF96FF43F85CE8BEFA8F" box="[151,448,1952,1974]" italics="true" pageId="17">Cyanogomphus comparabilis</emphasis>
Belle, 1994
</taxonomicName>
, Holotype (Brazil. MT: Diamantino, ABMM); G–H.
<taxonomicName id="06F7EDD9FFD8FF96FB83F85CE848FAED" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="17" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFD8FF96FB83F85CEC5BFA8F" box="[1111,1317,1952,1974]" italics="true" pageId="17">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</taxonomicName>
(Brazil. RS: Jaguarão, MNRJ); I–J.
<taxonomicName id="06F7EDD9FFD8FF96FD7CF842ED5CFAED" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[680,1058,1982,2005]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="17" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD8FF96FD7CF842EAF3FAEC" box="[680,909,1982,2005]" italics="true" pageId="17">Tibiagomphus uncatus</emphasis>
(Fraser, 1947)
</taxonomicName>
(Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="9588C6D2FFDBFF95FF43FB74EBBFF820" httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="18" targetBox="[151,1434,193,1138]" targetPageId="18">
<paragraph id="C148965AFFDBFF95FF43FB74EBBFF820" blockId="18.[151,1436,1160,1305]" pageId="18">
<emphasis id="F3834A48FFDBFF95FF43FB74E86FF9A4" bold="true" box="[151,273,1160,1182]" pageId="18">FIGURE 9</emphasis>
. Female S8–10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A–B.
<taxonomicName id="06F7EDD9FFDBFF95FF43FB5BEB72F984" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" box="[151,524,1191,1213]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="18" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDBFF95FF43FB5BE8F1F984" box="[151,399,1191,1213]" italics="true" pageId="18">Cyanogomphus waltheri</emphasis>
Selys, 1873
</taxonomicName>
(Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C.
<taxonomicName id="06F7EDD9FFDBFF95FE32FB39EA4AF9E2" box="[486,820,1221,1243]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="18" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFDBFF95FE32FB39EA4AF9E2" box="[486,820,1221,1243]" italics="true" pageId="18">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFDBFF95FCE8FB3AEAF3F9E2" bold="true" box="[828,909,1222,1243]" pageId="18">
<taxonomicNameLabel id="E8B0F733FFDBFF95FCE8FB3AEAF3F9E2" box="[828,909,1222,1243]" pageId="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
additional specimen (Brazil. MG: Serra do Salitre, ABMM); D.
<taxonomicName id="06F7EDD9FFDBFF95FEC9FB18E8C5F9C3" box="[285,443,1252,1274]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="18" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDBFF95FEC9FB18E8C5F9C3" box="[285,443,1252,1274]" italics="true" pageId="18">Cyanogomphus</emphasis>
</taxonomicName>
sp. (Brazil. GO: Jataí, ABMM); E.
<taxonomicName id="06F7EDD9FFDBFF95FCF9FB18EC65F9C3" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[813,1307,1252,1274]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="18" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDBFF95FCF9FB18EA82F9C3" box="[813,1020,1252,1274]" italics="true" pageId="18">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</taxonomicName>
(Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm.
</paragraph>
</caption>
<paragraph id="C148965AFFDBFF95FF13FABCEB00F844" blockId="18.[151,1437,1344,2017]" pageId="18">
<emphasis id="F3834A48FFDBFF95FF13FABCE8F7F860" bold="true" box="[199,393,1344,1369]" pageId="18">
<typeStatus id="1E4C28F8FFDBFF95FF13FABCE87CF860" box="[199,258,1344,1369]" pageId="18">Type</typeStatus>
repository.
</emphasis>
<typeStatus id="1E4C28F8FFDBFF95FE44FABCE886F860" box="[400,504,1344,1369]" pageId="18" type="holotype">Holotype</typeStatus>
♂ by monotypy in Selys collection at Royal Belgian Institute of Natural Sciences, Brussels,
<collectingCountry id="B9E0D6CAFFDBFF95FED0FA98E81BF844" box="[260,357,1380,1405]" name="Belgium" pageId="18">Belgium</collectingCountry>
(
<collectionCode id="A7E60E9FFFDBFF95FEA7FA99E8B7F844" box="[371,457,1381,1405]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="18">IRSNB</collectionCode>
, not examined).
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDBFF95FF13FA74EB9CFBDC" pageId="18" type="description">
<paragraph id="C148965AFFDBFF95FF13FA74E896FB08" blockId="18.[151,1437,1344,2017]" pageId="18">
<emphasis id="F3834A48FFDBFF95FF13FA74E803F898" bold="true" box="[199,381,1416,1441]" pageId="18">Measurements.</emphasis>
Males (n = 7). Total length (including caudal appendages) 43.6–50.1; abdomen length (excluding caudal appendages) 32.5–37.4; head maximum width 5.9–6.6; Fw length 26.4–29.5; Hw length 25.8–28.1; Fw maximum width 5.2–6.1, in Hw 7.0–7.6; pt length 3.0–
<quantity id="060F3BBFFFDBFF95FC3CFA2DED50F8D0" box="[1000,1070,1488,1513]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" pageId="18" unit="in" value="3.4">3.4 in</quantity>
Fw,
<quantity id="060F3BBFFFDBFF95FBA5FA2CED96F8D0" box="[1137,1256,1488,1513]" metricMagnitude="-2" metricUnit="m" metricValue="8.509" metricValueMax="9.144" metricValueMin="7.874" pageId="18" unit="in" value="3.35" valueMax="3.6" valueMin="3.1">3.1–3.6 in</quantity>
Hw; length of metathoracic femur 6.5–8.0; metathoracic tibia 4.7–5.7; length of S9+
<quantity id="060F3BBFFFDBFF95FC6AFA09EA84FB34" box="[958,1018,1524,1549]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="18" unit="in" value="10.0">10 in</quantity>
lateral view 2.7–3.5; total length of cercus in lateral view 1.1–1.4.
</paragraph>
<paragraph id="C148965AFFDBFF95FF13F9C0ECFBFBF8" blockId="18.[151,1437,1344,2017]" pageId="18">
Females (n = 2). Total length (including caudal appendages) 48.6–50.4; abdomen length (excluding caudal appendages) 35.8–35.9; head maximum width 6.8; Fw length 32.0–32.1; Hw length 30.9–31.6; Fw maximum width 6.2–6.4, in Hw 8.1–8.3; pt length
<quantity id="060F3BBFFFDBFF95FDACF978EB93FBA4" box="[632,749,1668,1693]" metricMagnitude="-2" metricUnit="m" metricValue="9.525" metricValueMax="9.906" metricValueMin="9.144" pageId="18" unit="in" value="3.75" valueMax="3.9" valueMin="3.6">3.6–3.9 in</quantity>
Fw,
<quantity id="060F3BBFFFDBFF95FCF9F979EADCFBA4" box="[813,930,1668,1693]" metricMagnitude="-2" metricUnit="m" metricValue="9.906" metricValueMax="10.414" metricValueMin="9.398" pageId="18" unit="in" value="3.9" valueMax="4.1" valueMin="3.7">3.7–4.1 in</quantity>
Hw; length of metathoracic femur 7.1–7.6 metathoracic tibia 5.4–6.3; length of S9+
<quantity id="060F3BBFFFDBFF95FDB7F955EBE2FBF8" box="[611,668,1704,1729]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="18" unit="in" value="10.0">10 in</quantity>
lateral view 2.2–2.2; total length of cercus in lateral view 1.0–1.2.
</paragraph>
<paragraph id="C148965AFFDBFF95FF13F930EB9CFBDC" blockId="18.[151,1437,1344,2017]" box="[199,738,1740,1765]" pageId="18">
<emphasis id="F3834A48FFDBFF95FF13F930E869FBDC" bold="true" box="[199,279,1740,1765]" pageId="18">Larva.</emphasis>
<bibRefCitation id="A566EBABFFDBFF95FECAF930EB51FBDC" author="Needham" box="[286,559,1740,1765]" pageId="18" refString="Needham, J. G. (1940) Studies on Neotropical Gomphine Dragonflies. Transactions of the American Entomological Society, 65 (4), 363 - 394." type="journal article" year="1940" yearSuffix="p">Needham (1940, p. 382)</bibRefCitation>
by supposition.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDBFF94FF13F90CEA30FCB1" lastPageId="19" pageId="18" type="diagnosis">
<paragraph id="C148965AFFDBFF94FF13F90CEA30FCB1" blockId="18.[151,1437,1344,2017]" lastBlockId="19.[151,1437,151,1293]" lastPageId="19" pageId="18">
<emphasis id="F3834A48FFDBFF95FF13F90CE83FFA30" bold="true" box="[199,321,1776,1801]" pageId="18">Diagnosis.</emphasis>
Both sexes of
<taxonomicName id="06F7EDD9FFDBFF95FE25F90DEB0FFA30" box="[497,625,1776,1801]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="18" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDBFF95FE25F90DEB0FFA30" box="[497,625,1776,1801]" italics="true" pageId="18">C. waltheri</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="06F7EDD9FFDBFF95FC7BF90CED30FA30" box="[943,1102,1776,1801]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="18" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDBFF95FC7BF90CED30FA30" box="[943,1102,1776,1801]" italics="true" pageId="18">Tibiagomphus</emphasis>
</taxonomicName>
spp. by defined antehumeral stripes (as
<figureCitation id="59CC8ADFFFDBFF95FEDAF8E8E832FA14" box="[270,332,1812,1837]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="18">Fig. 1</figureCitation>
B, stripes from undefined to a very narrow first antehumeral stripe in
<taxonomicName id="06F7EDD9FFDBFF95FB8DF8E8ED86FA14" box="[1113,1272,1812,1837]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="18" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDBFF95FB8DF8E8ED86FA14" box="[1113,1272,1812,1837]" italics="true" pageId="18">Tibiagomphus</emphasis>
</taxonomicName>
), males by
<date id="B549B09AFFDBFF95FAAEF8E9ECE2FA14" box="[1402,1436,1813,1837]" pageId="18">V4</date>
cornet-like distal portion wide (
<figureCitation id="59CC8ADFFFDBFF95FE23F8C4EB43FA68" box="[503,573,1848,1873]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="18">Figs 6</figureCitation>
C–D), cercus cylindrical, slightly curved toward to the upturned tapered apex (
<figureCitation id="59CC8ADFFFDBFF95FF4BF8A0E999FA4C" box="[159,231,1884,1909]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="18">Figs 7</figureCitation>
A–C, 8A–B), and branches of epiproct with a distal concavity (
<figureCitation id="59CC8ADFFFDBFF95FC17F8A0ED72FA4C" box="[963,1036,1884,1909]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="18">Figs 8</figureCitation>
B, D, F), and female by subgenital plate short (ratio between length of plate and S9 &lt;0.4), and with a wide concavity (
<figureCitation id="59CC8ADFFFDBFF95FB84F87CEDE6FAA0" box="[1104,1176,1920,1945]" captionStart="FIGURE 9" captionStartId="18.[151,250,1160,1182]" captionTargetBox="[151,1434,193,1138]" captionTargetId="figure@18.[151,1436,193,1139]" captionTargetPageId="18" captionText="FIGURE 9. Female S 8 – 10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A – B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C. Cyanogomphus angelomachadoi sp. nov. additional specimen (Brazil. MG: Serra do Salitre, ABMM); D. Cyanogomphus sp. (Brazil. GO: Jataí, ABMM); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm." httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="18">Figs 9</figureCitation>
A–B,
<date id="B549B09AFFDBFF95FB08F87DED81FAA0" box="[1244,1279,1921,1945]" pageId="18">V4</date>
distal portion narrow, apex of cercus toward posteriorly, epiproct dorsally smooth, without concavity, and female subgenital plate large and with a narrow mesal incision in
<taxonomicName id="06F7EDD9FFDBFF95FDB8F834EA75FAD8" box="[620,779,1992,2017]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="18" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDBFF95FDB8F834EA75FAD8" box="[620,779,1992,2017]" italics="true" pageId="18">Tibiagomphus</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDBFF95FCCDF834EA23FAD8" box="[793,861,1992,2017]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="18">Figs 6</figureCitation>
F–G, 7J–
<collectingCountry id="B9E0D6CAFFDBFF95FC17F835EAA6FAD8" box="[963,984,1993,2017]" name="Iceland" pageId="18">O</collectingCountry>
, 8G–J, 9E). This species is more similar to
<taxonomicName id="06F7EDD9FFDAFF94FF62FF64E8EAFD89" box="[182,404,151,176]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFDAFF94FF62FF64E8EAFD89" box="[182,404,151,176]" italics="true" pageId="19">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFDAFF94FE49FF6BE884FD89" bold="true" box="[413,506,151,176]" pageId="19">
<taxonomicNameLabel id="E8B0F733FFDAFF94FE49FF6BE884FD89" box="[413,506,151,176]" pageId="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, than to
<taxonomicName id="06F7EDD9FFDAFF94FDB7FF64EA64FD89" box="[611,794,151,176]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFDAFF94FDB7FF64EA64FD89" box="[611,794,151,176]" italics="true" pageId="19">C. comparabilis</emphasis>
</taxonomicName>
and is diagnosed under those species. The black dorsal surface of the tibia in
<taxonomicName id="06F7EDD9FFDAFF94FE41FF41EB6BFDEC" box="[405,533,188,213]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDAFF94FE41FF41EB6BFDEC" box="[405,533,188,213]" italics="true" pageId="19">C. waltheri</emphasis>
</taxonomicName>
and allopatric distribution in the Atlantic Forest further separates it from them (
<figureCitation id="59CC8ADFFFDAFF94FF4BFF23E992FDC1" box="[159,236,223,248]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="19">Figs 3</figureCitation>
B, 12, dorsal surface of tibia pale and Cerrado distribution in
<taxonomicName id="06F7EDD9FFDAFF94FC38FF1CEDB1FDC1" box="[1004,1231,223,248]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFDAFF94FC38FF1CEDB1FDC1" box="[1004,1231,223,248]" italics="true" pageId="19">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFDAFF94FB08FF23EC43FDC1" bold="true" box="[1244,1341,223,248]" pageId="19">
<taxonomicNameLabel id="E8B0F733FFDAFF94FB08FF23EC43FDC1" box="[1244,1341,223,248]" pageId="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="06F7EDD9FFDAFF94FA56FF1CE855FC24" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFDAFF94FA56FF1CE855FC24" italics="true" pageId="19">C. comparabilis</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDAFF94FEEDFEF8E803FC25" box="[313,381,260,285]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="19">Figs 3</figureCitation>
A, 12). The female differs from
<taxonomicName id="06F7EDD9FFDAFF94FD3CFEF9EABAFC24" box="[744,964,260,285]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFDAFF94FD3CFEF9EABAFC24" box="[744,964,260,285]" italics="true" pageId="19">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFDAFF94FC1FFEF8ED5BFC24" bold="true" box="[971,1061,260,285]" pageId="19">
<taxonomicNameLabel id="E8B0F733FFDAFF94FC1FFEF8ED5BFC24" box="[971,1061,260,285]" pageId="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the color pattern with defined mesepisternal stripes; however, the male from Serra do Salitre (ABMM) and possible females also can exhibit a mesepisternal color similar to that of the latter species (
<figureCitation id="59CC8ADFFFDAFF94FCC7FEB0EA28FC5D" box="[787,854,332,357]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="19">Figs 1</figureCitation>
A–B) but
<taxonomicName id="06F7EDD9FFDAFF94FC1BFEB1ED30FC5C" box="[975,1102,332,357]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDAFF94FC1BFEB1ED30FC5C" box="[975,1102,332,357]" italics="true" pageId="19">C. waltheri</emphasis>
</taxonomicName>
also differs by its larger size (Hw 30.9–31.6 vs.
<quantity id="060F3BBFFFDAFF94FEB9FE93E885FCB1" box="[365,507,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="6.2103" metricValueMax="6.223" metricValueMin="6.1975999999999996" pageId="19" unit="in" value="24.45" valueMax="24.5" valueMin="24.4">24.4–24.5 in</quantity>
<taxonomicName id="06F7EDD9FFDAFF94FDD5FE8CEBA3FCB1" box="[513,733,367,392]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFDAFF94FDD5FE8CEBA3FCB1" box="[513,733,367,392]" italics="true" pageId="19">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFDAFF94FD30FE93EA43FCB1" bold="true" box="[740,829,367,392]" pageId="19">
<taxonomicNameLabel id="E8B0F733FFDAFF94FD30FE93EA43FCB1" box="[740,829,367,392]" pageId="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDAFF94FF13FE68EDD8FFBC" pageId="19" type="distribution">
<paragraph id="C148965AFFDAFF94FF13FE68EC0BFCE9" blockId="19.[151,1437,151,1293]" pageId="19">
<emphasis id="F3834A48FFDAFF94FF13FE68E827FC94" bold="true" box="[199,345,404,429]" pageId="19">Distribution</emphasis>
. From Minas Gerais State south to Rio Grande do Sul State in
<collectingCountry id="B9E0D6CAFFDAFF94FBF9FE68ED0FFC94" box="[1069,1137,404,429]" name="Brazil" pageId="19">Brazil</collectingCountry>
and Misiones Province in
<collectingCountry id="B9E0D6CAFFDAFF94FF43FE4BE874FCE9" box="[151,266,439,464]" name="Argentina" pageId="19">Argentina</collectingCountry>
; along in the Parana dominion in the Atlantic and Parana Forest biogeographical provinces (
<figureCitation id="59CC8ADFFFDAFF94FAC0FE4BEC16FCE9" box="[1300,1384,439,464]" captionStart="FIGURE 12" captionStartId="24.[151,250,1240,1262]" captionTargetBox="[151,1436,215,1198]" captionTargetId="figure@24.[151,1436,195,1215]" captionTargetPageId="24" captionText="FIGURE 12. Map of southeastern South America with distribution records of Cyanogomphus Selys, 1873, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: MI, Misiones Province, Argentina; BA, Bahia; GO, Goiás; MG, Minas Gerais; MT, Mato Grosso; RJ, Rio de Janeiro; RS, Rio Grande do Sul; SC, Santa Catarina, and SP, São Paulo for Brazilian States." httpUri="https://zenodo.org/record/270865/files/figure.png" pageId="19">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="C148965AFFDAFF94FF13FE20EDD8FFBC" blockId="19.[151,1437,151,1293]" pageId="19">
<emphasis id="F3834A48FFDAFF94FF13FE20EB4FFCCC" bold="true" box="[199,561,476,501]" pageId="19">Biological and ecological data.</emphasis>
Supposed larvae of this species were found in lotic environments in silt, sand and gravel beds of rocky forested streams (
<bibRefCitation id="A566EBABFFDAFF94FDAEFE03EABEFF21" author="Carvalho" box="[634,960,511,536]" pageId="19" refString="Carvalho, A. L. &amp; Nessimian, J. L. (1998) Odonata do Estado do Rio de Janeiro, Brasil: habitats e habitos das larvas. In: Nessimian, J. L. &amp; Carvalho, A. L. (Eds.), Ecologia de Insetos Aquaticos. Series Oecologia Brasiliensis. Vol. V. PPGE- UFRJ, Rio de Janeiro, pp. 3 - 28." type="book chapter" year="1998">Carvalho &amp; Nessimian 1998</bibRefCitation>
;
<bibRefCitation id="A566EBABFFDAFF94FC18FDFCEDF1FF21" author="Costa" box="[972,1167,511,536]" pageId="19" refString="Costa, J. M., Irineu-de-Souza, L. O. &amp; Oldrini, B. B. (2004) Chave para identificacao das familias e generos das larvas conhecidas de Odonata do Brasil: comentarios e registros bibliograficos (Insecta, Odonata). Publicacoes Avulsas do Museu Nacional, 99, 1 - 44." type="journal article" year="2004">
Costa
<emphasis id="F3834A48FFDAFF94FBC7FDFDED32FF21" box="[1043,1100,511,536]" italics="true" pageId="19">et al.</emphasis>
2004
</bibRefCitation>
), where they apparently prey on smaller organisms found on the surface of the substrate (
<bibRefCitation id="A566EBABFFDAFF94FCB2FDD8EDD4FF05" author="Carvalho" box="[870,1194,548,573]" pageId="19" refString="Carvalho, A. L. &amp; Nessimian, J. L. (1998) Odonata do Estado do Rio de Janeiro, Brasil: habitats e habitos das larvas. In: Nessimian, J. L. &amp; Carvalho, A. L. (Eds.), Ecologia de Insetos Aquaticos. Series Oecologia Brasiliensis. Vol. V. PPGE- UFRJ, Rio de Janeiro, pp. 3 - 28." type="book chapter" year="1998">Carvalho &amp; Nessimian 1998</bibRefCitation>
). Larvae are common in rocky forested rivers in the Atlantic Forest domain of southeastern
<collectingCountry id="B9E0D6CAFFDAFF94FC49FDBBEA9FFF59" box="[925,993,583,608]" name="Brazil" pageId="19">Brazil</collectingCountry>
(APP, pers. obs.); adults were observed perched on low vegetation and shrubs at the shore, from sea level to 914 meters of elevation.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDAFF94FF13FD73EA6CF835" pageId="19" type="discussion">
<paragraph id="C148965AFFDAFF94FF13FD73EB65FEF1" blockId="19.[151,1437,151,1293]" pageId="19">
<emphasis id="F3834A48FFDAFF94FF13FD73E84AFF91" bold="true" box="[199,308,655,680]" pageId="19">Remarks</emphasis>
. Females from the Cerrado of Goiás State,
<collectingCountry id="B9E0D6CAFFDAFF94FCF4FD73EA19FF91" box="[800,871,655,680]" name="Brazil" pageId="19">Brazil</collectingCountry>
, determined by
<bibRefCitation id="A566EBABFFDAFF94FBC8FD73EDD4FF91" author="Belle" box="[1052,1194,655,680]" pageId="19" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle (1980)</bibRefCitation>
do not pertain to this species, and due to scarce data about females of other species, we are unable to identify them (see under
<taxonomicName id="06F7EDD9FFDAFF94FF43FD2BE83BFFC9" box="[151,325,727,752]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDAFF94FF43FD2BE83BFFC9" box="[151,325,727,752]" italics="true" pageId="19">Cyanogomphus</emphasis>
</taxonomicName>
sp.). The
<typeStatus id="1E4C28F8FFDAFF94FE63FD24E899FFC9" box="[439,487,728,752]" pageId="19">type</typeStatus>
locality at the Botanical Garden of Rio de Janeiro is contiguous to Parque Nacional da Tijuca; however, this species was not found in the Tijuca forest after 47 years of collections by
<bibRefCitation id="A566EBABFFDAFF94FB39FD00E99FFE01" author="Santos" pageId="19" refString="Santos, N. D. &amp; Costa, J. M. (1987) Preservacao dos odonatas do municipio do Rio de Janeiro. In: Resumos. XIV Congresso Brasileiro de Zoologia. Universidade Federal de Juiz de For a, Juiz de Fora, pp. 281." type="book chapter" year="1987">Santos &amp; Costa (1987)</bibRefCitation>
and recent collecting expeditions undertaken by APP (
<date id="B549B09AFFDAFF94FCB7FCDCEA9AFE01" box="[867,996,800,824]" pageId="19" value="2009" valueMax="2011">2009–2011</date>
) and MVA (since 2014), even though adults of other
<taxonomicName id="06F7EDD9FFDAFF94FE9BFCB8E8ACFE64" box="[335,466,836,861]" class="Insecta" family="Gomphidae" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="family">Gomphidae</taxonomicName>
including
<taxonomicName id="06F7EDD9FFDAFF94FD87FCB8EACCFE64" authority="Hagen" authorityName="Hagen" box="[595,946,836,861]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="paludosus">
<emphasis id="F3834A48FFDAFF94FD87FCB8EA23FE64" box="[595,861,836,861]" italics="true" pageId="19">Epigomphus paludosus</emphasis>
Hagen
</taxonomicName>
<emphasis id="F3834A48FFDAFF94FC6AFCB9EAAAFE65" box="[958,980,837,860]" italics="true" pageId="19">in</emphasis>
Selys, 1854,
<taxonomicName id="06F7EDD9FFDAFF94FBAFFCB8E861FEB9" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" class="Insecta" family="Gomphidae" genus="Progomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="complicatus">
<emphasis id="F3834A48FFDAFF94FBAFFCB8ECE5FE64" box="[1147,1435,836,861]" italics="true" pageId="19">Progomphus complicatus</emphasis>
Selys, 1854
</taxonomicName>
, and
<taxonomicName id="06F7EDD9FFDAFF94FE8AFC9BEA8CFEB9" authority="Burmeister, 1839" authorityName="Burmeister" authorityYear="1839" box="[350,1010,871,896]" class="Insecta" family="Gomphidae" genus="Zonophora" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="subSpecies" species="campanulata" subSpecies="campanulata">
<emphasis id="F3834A48FFDAFF94FE8AFC9BEA6CFEB9" box="[350,786,871,896]" italics="true" pageId="19">Zonophora campanulata campanulata</emphasis>
(Burmeister, 1839)
</taxonomicName>
were frequently collected during the summer season. Preferential habits for
<taxonomicName id="06F7EDD9FFDAFF94FD84FC71EBAEFE9C" box="[592,720,908,933]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDAFF94FD84FC71EBAEFE9C" box="[592,720,908,933]" italics="true" pageId="19">C. waltheri</emphasis>
</taxonomicName>
might be reduced, making it less abundant and thus difficult to detect, or it may be extinct locally.
</paragraph>
<paragraph id="C148965AFFDAFF94FF13FC28EDE8F944" blockId="19.[151,1437,151,1293]" pageId="19">
<collectingCountry id="B9E0D6CAFFDAFF94FF13FC28E998FED5" box="[199,230,980,1004]" name="American Samoa" pageId="19">As</collectingCountry>
in other species in the
<taxonomicName id="06F7EDD9FFDAFF94FE27FC28EBE6FED4" box="[499,664,980,1005]" class="Insecta" family="Gomphidae" genus="Agriogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDAFF94FE27FC28EBE6FED4" box="[499,664,980,1005]" italics="true" pageId="19">Agriogomphus</emphasis>
</taxonomicName>
-complex, synthoracic coloration in
<taxonomicName id="06F7EDD9FFDAFF94FBE0FC29EDCDFED4" box="[1076,1203,980,1005]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDAFF94FBE0FC29EDCDFED4" box="[1076,1203,980,1005]" italics="true" pageId="19">C. waltheri</emphasis>
</taxonomicName>
varies considerably. The majority have the pale areas similar to the
<typeStatus id="1E4C28F8FFDAFF94FD7DFC0BEA75F929" box="[681,779,1015,1040]" pageId="19" type="holotype">holotype</typeStatus>
illustrated by
<bibRefCitation id="A566EBABFFDAFF94FC7AFC0BEDFBF929" author="Belle" box="[942,1157,1015,1040]" pageId="19" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle (1980, fig. 1)</bibRefCitation>
, but these areas also can be larger, approaching the pattern observed in the
<typeStatus id="1E4C28F8FFDAFF94FD03FBE1EA49F90C" box="[727,823,1053,1077]" pageId="19" type="paratype">paratype</typeStatus>
of
<taxonomicName id="06F7EDD9FFDAFF94FCB4FBE1ED40F90C" box="[864,1086,1052,1077]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFDAFF94FCB4FBE1ED40F90C" box="[864,1086,1052,1077]" italics="true" pageId="19">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFDAFF94FB92FBE0EDDFF90C" bold="true" box="[1094,1185,1052,1077]" pageId="19">
<taxonomicNameLabel id="E8B0F733FFDAFF94FB92FBE0EDDFF90C" box="[1094,1185,1052,1077]" pageId="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Serra do Salitre (
<figureCitation id="59CC8ADFFFDAFF94FF4BFBC3E9A1F961" box="[159,223,1087,1112]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="19">Fig. 1</figureCitation>
C). In the darker specimens these areas are reduced, and the first antehumeral stripe is largely unconnected to other pale areas, thus not reaching the mesothoracic collar and base of the antealar sinus.
</paragraph>
<paragraph id="C148965AFFDAFF94FF13FB7BEA6CF835" blockId="19.[151,1437,151,1293]" pageId="19">
Probable larvae of this species are fairly common in sandy creeks in the Atlantic Forest of southeastern
<collectingCountry id="B9E0D6CAFFDAFF94FA8CFB7BECE2F999" box="[1368,1436,1159,1184]" name="Brazil" pageId="19">Brazil</collectingCountry>
based on previously published data (e.g.
<bibRefCitation id="A566EBABFFDAFF94FDA6FB50EAC2F9FD" author="Carvalho" box="[626,956,1196,1221]" pageId="19" refString="Carvalho, A. L. &amp; Nessimian, J. L. (1998) Odonata do Estado do Rio de Janeiro, Brasil: habitats e habitos das larvas. In: Nessimian, J. L. &amp; Carvalho, A. L. (Eds.), Ecologia de Insetos Aquaticos. Series Oecologia Brasiliensis. Vol. V. PPGE- UFRJ, Rio de Janeiro, pp. 3 - 28." type="book chapter" year="1998">Carvalho &amp; Nessimian 1998</bibRefCitation>
) and recent collections (APP, pers. obs.); however their identity needs confirmation, as the larva was described by supposition by
<bibRefCitation id="A566EBABFFDAFF94FBACFB33EC4BF9D1" author="Needham" box="[1144,1333,1231,1256]" pageId="19" refString="Needham, J. G. (1944) Further Studies on Neotropical Gomphine Dragonflies (Odonata). Transactions of the American Entomological Society, 69 (4), 171 - 224." type="journal article" year="1944">Needham (1944)</bibRefCitation>
and may be difficult to differentiate from larvae of
<taxonomicName id="06F7EDD9FFDAFF94FDB8FB08EA75F834" box="[620,779,1268,1293]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDAFF94FDB8FB08EA75F834" box="[620,779,1268,1293]" italics="true" pageId="19">Tibiagomphus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

681
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<mods:title id="0E5FF0B17505271923F1D80C0BD56A5B">A taxonomic synopsis of South American Cyanogomphini Carle with description of Cyanogomphus angelomachadoi sp. nov. from the Cerrado of Brazil (Odonata: Gomphidae)</mods:title>
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<heading id="9A002136FFDAFF94FF43FA9CEB99F843" bold="true" box="[151,743,1376,1402]" fontSize="11" level="1" pageId="19" reason="1">
<taxonomicName id="06F7EDD9FFDAFF94FF43FA9CEB99F843" ID-CoL="7BYMM" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[151,743,1376,1402]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDAFF94FF43FA9CEB99F843" bold="true" box="[151,743,1376,1402]" pageId="19">
<emphasis id="F3834A48FFDAFF94FF43FA9CE8F0F843" bold="true" box="[151,398,1376,1402]" italics="true" pageId="19">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</emphasis>
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LSID urn:lsid:zoobank.org:act:645A61F2-F59F-42A7-8B73-4270D67477AB (
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E, 3C, 4E, 5A–B, F–H, 6F–G, 7J–L, 8G–H, 9E, 10E, 11D, 13)
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<emphasis id="F3834A48FFDAFF94FF43FA16E8C3FB39" box="[151,445,1514,1536]" italics="true" pageId="19">
<taxonomicName id="06F7EDD9FFDAFF94FF43FA16E8F0FB39" ID-CoL="9WYQS" box="[151,398,1514,1536]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="uncatus">Cyanogomphus uncatus</taxonomicName>
<emphasis id="F3834A48FFDAFF94FE4DFA17E8C3FB39" bold="true" box="[409,445,1515,1536]" italics="true" pageId="19">nec</emphasis>
</emphasis>
<bibRefCitation id="A566EBABFFDAFF94FE1CFA17EB33FB39" author="Fraser" box="[456,589,1514,1536]" pageId="19" refString="Fraser, F. C. (1947) The Odonata of the Argentine Republic I. Acta Zoologica Lilloana del Instituto Miguel Lillo, 4, 427 - 462." type="journal article" year="1947">Fraser, 1947</bibRefCitation>
:—St.
<bibRefCitation id="A566EBABFFDAFF94FD45FA16EAFDFB26" author="Quentin" pageId="19" refString="St. Quentin, D. (1973) Die Gomphidenfauna Sudamerikas (Ordn.: Odonata). Annalen des Naturhistorischen Museums in Wien, 77, 335 - 363." type="journal article" year="1973">
Quentin (1973: 358, record from Rio Grande do Sul State,
<collectingCountry id="B9E0D6CAFFDAFF94FAC7FA16EC2DFB39" box="[1299,1363,1514,1536]" name="Brazil" pageId="19">Brazil</collectingCountry>
, likely pertains to this species; comparison with the
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of
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<emphasis id="F3834A48FFDAFF94FCC0F9F6EA03FB26" box="[788,893,1546,1567]" italics="true" pageId="19">T. uncatus</emphasis>
</taxonomicName>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDAFF94FC77F9F5EB64FBA3" author="Belle" pageId="19" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">
Belle (1980: 153–155, figs 5–13, in part, records from Entre Rios Province,
<collectingCountry id="B9E0D6CAFFDAFF94FE0DF9D4EB3FFB07" box="[473,577,1576,1598]" name="Argentina" pageId="19">Argentina</collectingCountry>
, and Florida and Durazno Departments,
<collectingCountry id="B9E0D6CAFFDAFF94FC34F9D5ED47FB07" box="[992,1081,1577,1598]" name="Uruguay" pageId="19">Uruguay</collectingCountry>
, illustrations of metathoracic tibia, tarsi and pretarsal claws in lateral view,
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V
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in lateral view,
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V
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in ventral view, ligula ‘penis guard’ in anteroposterior and lateral views, anterior hamules in ventral view and secondary genitalia in lateral view, comparison with male
<typeStatus id="1E4C28F8FFDAFF94FA97F999ECE2FB42" box="[1347,1436,1637,1659]" pageId="19" type="holotype">holotype</typeStatus>
and female and with
<taxonomicName id="06F7EDD9FFDAFF94FE74F978EB6CFBA3" ID-CoL="32NMT" box="[416,530,1668,1690]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDAFF94FE74F978EB6CFBA3" box="[416,530,1668,1690]" italics="true" pageId="19">C. waltheri</emphasis>
</taxonomicName>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDAFF94FDEEF978EAF8FBA3" author="Belle" box="[570,902,1668,1690]" pageId="19" refString="Belle, J. (1981) An annotated list of the Gomphidae of Uruguay (Odonata). Revista de la Facultad de Humanidades y Ciencias, Serie Ciencias Biologicas, 1 (16), 261 - 264." type="journal article" year="1981">Belle (1981: 261–262, mention)</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDAFF94FC70F978ED9AFBA3" author="De" box="[932,1252,1668,1690]" pageId="19" refString="De Abenante, Y. P. &amp; Philippi, M. E. (1982) Lista preliminar de los odonatos del Uruguay. Notulae Odonatologicae, 1 (9), 151." type="journal article" year="1982">
<collectingCountry id="B9E0D6CAFFDAFF94FC70F978EABCFBA3" box="[932,962,1668,1690]" name="South Georgia and the South Sandwich Islands" pageId="19">De</collectingCountry>
<bibRefCitation id="A566EBABFFDAFF94FC1EF978ED9AFBA3" author="Abenante" box="[970,1252,1668,1690]" pageId="19" refString="De Abenante, Y. P. &amp; Philippi, M. E. (1982) Lista preliminar de los odonatos del Uruguay. Notulae Odonatologicae, 1 (9), 151." type="journal article" year="1982">Abenante &amp; Philippi (1982</bibRefCitation>
</bibRefCitation>
: 151, record from
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);—
<bibRefCitation id="A566EBABFFDAFF94FE92F95EEB2DFB81" author="Rodrigues" box="[326,595,1698,1720]" pageId="19" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">
Rodrigues
<bibRefCitation id="A566EBABFFDAFF94FE62F95EEB2DFB81" author="Capitulo" box="[438,595,1698,1720]" pageId="19" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">Capítulo (1992</bibRefCitation>
</bibRefCitation>
: 58, in part, record from Entre Rios Province,
<collectingCountry id="B9E0D6CAFFDAFF94FBF3F95EEDEFFB81" box="[1063,1169,1698,1720]" name="Argentina" pageId="19">Argentina</collectingCountry>
).
</paragraph>
<paragraph id="C148965AFFDAFF94FF43F93DEAAAFA97" blockId="19.[151,1436,1514,2027]" pageId="19">
<taxonomicName id="06F7EDD9FFDAFF94FF43F93DEBC0FBEE" ID-CoL="9WYQR" authority="Rodrigues Capitulo, 1985: 329" authorityName="Rodrigues Capitulo" authorityPageNumber="329" authorityYear="1985" box="[151,702,1729,1751]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDAFF94FF43F93DE80BFBEE" box="[151,373,1729,1751]" italics="true" pageId="19">Cyanogomphus noval</emphasis>
Rodrigues Capítulo, 1985: 329
</taxonomicName>
–335, figs 2–14 (description of ♂
<typeStatus id="1E4C28F8FFDAFF94FBF0F93DED01FBEE" box="[1060,1151,1729,1751]" pageId="19" type="holotype">holotype</typeStatus>
,
<collectingCountry id="B9E0D6CAFFDAFF94FB5FF93DEC5CFBEE" box="[1163,1314,1729,1751]" name="Argentina" pageId="19">ARGENTINA</collectingCountry>
. Entre Rios Province: Colón Department, Parque Nacional El Palmar,
<date id="B549B09AFFDAFF94FCC6F91CEA16FBCF" box="[786,872,1760,1782]" pageId="19" value="1982-11">XI.1982</date>
, Rodrigues Capítulo leg. in ILPLA [sic MLP], ♀ and F- 0 larva, illustrations of postfrons in dorsal view, secondary genitalia in lateral and ventral views, caudal appendages in dorsal, lateral and ventral views, S
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dorsal view of ♂
<typeStatus id="1E4C28F8FFDAFF94FCD9F8E1EA16FA0A" box="[781,872,1821,1843]" pageId="19" type="holotype">holotype</typeStatus>
, subgenital plate and caudal appendages of ♀ in ventral view, habitus of F-0 larvae in dorsal view, head and labium in dorsal view, caudal appendages in dorsolateral view, dorsal margin of abdomen in lateral view comparison with
<taxonomicName id="06F7EDD9FFDAFF94FD27F8A7EA23FA49" ID-CoL="9WYQS" box="[755,861,1883,1904]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDAFF94FD27F8A7EA23FA49" box="[755,861,1883,1904]" italics="true" pageId="19">T. uncatus</emphasis>
</taxonomicName>
and
<taxonomicName id="06F7EDD9FFDAFF94FC41F8A7EA8DFA49" ID-CoL="B7Q8" box="[917,1011,1883,1904]" class="Insecta" family="Gomphidae" genus="Agriogomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="ericae">
<emphasis id="F3834A48FFDAFF94FC41F8A7EA8DFA49" box="[917,1011,1883,1904]" italics="true" pageId="19">A. ericae</emphasis>
</taxonomicName>
);—
<bibRefCitation id="A566EBABFFDAFF94FBCEF8A6EC03FA49" author="Santos" box="[1050,1405,1882,1904]" pageId="19" refString="Santos, N. D. (1988) Catalogo Bibliografico de Ninfas de odonatos neotropicais. (Acompanhado de relacao alfabetica de autores e seus trabalhos). Acta Amazonica, 18 (1 - 2), 265 - 350." type="journal article" year="1988">Santos (1988: 267, 286, mention)</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDAFF94FF13F885E8A5FAB6" author="Rodrigues" box="[199,475,1913,1935]" pageId="19" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">Rodrigues Capítulo (1992</bibRefCitation>
: 58, 67, 81, 83, 89, figs 1–2, 4–5, 110, 132, 134, 180–181, distribution, reproduction of illustrations from Rodrigues Capítulo 1985);—
<bibRefCitation id="A566EBABFFDAFF94FD4BF864EAAEFA97" author="Garrison" box="[671,976,1944,1966]" pageId="19" refString="Garrison, R. W. (1991) A synonymic list of the New World Odonata. Argia, 3 (2), 1 - 30." type="journal article" year="1991">Garrison (1991: 17, mention)</bibRefCitation>
.
</paragraph>
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<taxonomicName id="06F7EDD9FFDAFF94FF43F84AEB01FAF5" ID-CoL="7BYMM" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[151,639,1974,1996]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="19" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDAFF94FF43F84AE81AFAF5" box="[151,356,1974,1996]" italics="true" pageId="19">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</taxonomicName>
:—
<bibRefCitation id="A566EBABFFDAFF94FD4AF84AEA58FAF5" author="Belle" box="[670,806,1974,1996]" pageId="19" refString="Belle, J. (1992 a) Studies on ultimate instar larvae of Neotropical Gomphidae, with the description of Tibiagomphus gen. nov. (Anisoptera). Odonatologica, 21 (1), 1 - 24." type="journal article" year="1992" yearSuffix="a">Belle (1992a</bibRefCitation>
: 1, comb. nov., implicit by the context ICZN 1999, Art. 11.9.3 of the Code);—
<bibRefCitation id="A566EBABFFDAFF94FEB9F829EBF0FAD2" author="Belle" box="[365,654,2005,2027]" pageId="19" refString="Belle, J. (1993) Annotated checklist and bibliography of the immature stages of Neotropical Gomphidae, published up to September 1993 (Anisoptera). Odonatologica, 22 (4), 399 - 409." type="journal article" year="1993">Belle (1993: 406, mention)</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDAFF94FD78F829ED15FAD2" author="Bridges" box="[684,1131,2005,2027]" pageId="19" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Odonata of the world (Third Edition). Privately published, Urbana, Illinois, 905 pp." type="book" year="1994">
Bridges (1994: VII.168,
<date id="B549B09AFFDAFF94FC67F829ED7EFAD2" box="[947,1024,2005,2027]" pageId="19" value="1970-08">VIII.70</date>
, mention)
</bibRefCitation>
;—Muzón &amp; von
<bibRefCitation id="A566EBABFFDAFF93FAFFF829EB70FD97" author="Ellenrieder" lastPageId="20" pageId="19" refString="Muzon, J. &amp; von Ellenrieder, N. (1999) Status and distribution of Odonata (Insecta) within natural protected areas in Argentina. Biogeographica, 75 (3), 119 - 128." type="journal article" year="1999">Ellenrieder (1999: 125, distributional data)</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDDFF93FDF9FF64EA6DFD97" author="Garrison" box="[557,787,152,174]" pageId="20" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
Garrison
<emphasis id="F3834A48FFDDFF93FD46FF65EBB4FD97" box="[658,714,152,174]" italics="true" pageId="20">et al.</emphasis>
(2006
</bibRefCitation>
: 129–130, comparison, suggest synonym with
<taxonomicName id="06F7EDD9FFDDFF93FAD3FF65EC0DFD97" ID-CoL="9WYQS" box="[1287,1395,153,174]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDDFF93FAD3FF65EC0DFD97" box="[1287,1395,153,174]" italics="true" pageId="20">T. uncatus</emphasis>
</taxonomicName>
);—
<bibRefCitation id="A566EBABFFDDFF93FF13FF4AE813FDF5" author="Heckman" box="[199,365,182,204]" pageId="20" refString="Heckman, C. W. (2006) Encyclopedia of South American aquatic insects: Odonata Anisoptera. Illustrated keys to known families, genera, and species in South America. Springer, Dordrecht, viii + 725 pp." type="book" year="2006">Heckman (2006</bibRefCitation>
: 525, 534, 649, figs 3.2.589, 3.2.603, key, reproduction of illustrations from Rodrigues Capítulo 1985);— von
<bibRefCitation id="A566EBABFFDDFF93FF20FF29ED70FDD2" author="Ellenrieder" box="[244,1038,213,235]" pageId="20" refString="von Ellenrieder, N. &amp; Muzon, J. (2008) An updated checklist of the Odonata from Argentina. Odonatologica, 37 (1), 55 - 68." type="journal article" year="2008">
Ellenrieder &amp; Muzón (2008: 61, record from Entre Rios Province,
<collectingCountry id="B9E0D6CAFFDDFF93FC4BFF29ED79FDD2" box="[927,1031,213,235]" name="Argentina" pageId="20">Argentina</collectingCountry>
)
</bibRefCitation>
;—Muzón
<emphasis id="F3834A48FFDDFF93FBA8FF2AEDCEFDD2" box="[1148,1200,213,235]" italics="true" pageId="20">et al.</emphasis>
(2007: 302, 305, notes on the
<typeStatus id="1E4C28F8FFDDFF93FED8FF09E846FC33" box="[268,312,245,266]" pageId="20">type</typeStatus>
material);—
<bibRefCitation id="A566EBABFFDDFF93FE63FF08EA52FC33" author="Hamalainen" box="[439,812,244,266]" pageId="20" refString="Hamalainen, M. (2015) Catalogue of individuals commemorated in the scientific names of extant dragonflies, including lists of all available eponymous species-group and genus-group names. International Dragonfly Fund Report, 80, 1 - 168." type="journal article" year="2015">Hämäläinen (2015: 114, etymology)</bibRefCitation>
.
</paragraph>
<paragraph id="C148965AFFDDFF93FF43FEEEEB00FCC6" blockId="20.[151,1436,152,511]" pageId="20">
<emphasis id="F3834A48FFDDFF93FF43FEEEE8D7FC11" box="[151,425,274,296]" italics="true" pageId="20">
<taxonomicName id="06F7EDD9FFDDFF93FF43FEEEE803FC11" ID-CoL="7BYMN" box="[151,381,274,296]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">Tibiagomphus uncatus</taxonomicName>
<emphasis id="F3834A48FFDDFF93FE51FEEFE8D7FC11" bold="true" box="[389,425,275,296]" italics="true" pageId="20">nec</emphasis>
</emphasis>
(
<bibRefCitation id="A566EBABFFDDFF93FE6DFEEEEB44FC11" author="Fraser" box="[441,570,274,296]" pageId="20" refString="Fraser, F. C. (1947) The Odonata of the Argentine Republic I. Acta Zoologica Lilloana del Instituto Miguel Lillo, 4, 427 - 462." type="journal article" year="1947">Fraser, 1947</bibRefCitation>
):—
<bibRefCitation id="A566EBABFFDDFF93FDB4FEEEEC64FC11" author="Belle" box="[608,1306,274,296]" pageId="20" refString="Belle, J. (1996) Higher classification of the South-American Gomphidae (Odonata). Zoologische Mededelingen, 70, 297 - 324." type="journal article" year="1996">
Belle (1996: 323, fig. 8, in part, pair of wings of ♂ from
<collectingCountry id="B9E0D6CAFFDDFF93FB6DFEEEEC6DFC11" box="[1209,1299,274,296]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDDFF93FAECFEEEE878FC7E" author="Heckman" pageId="20" refString="Heckman, C. W. (2006) Encyclopedia of South American aquatic insects: Odonata Anisoptera. Illustrated keys to known families, genera, and species in South America. Springer, Dordrecht, viii + 725 pp." type="book" year="2006">Heckman (2006</bibRefCitation>
: 649–650, fig. 3.2.778, in part, reproduction of illustrations from
<bibRefCitation id="A566EBABFFDDFF93FC1DFECDED43FC7E" author="Belle" box="[969,1085,305,327]" pageId="20" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle 1980</bibRefCitation>
of ♂ from Entre Rios Province,
<collectingCountry id="B9E0D6CAFFDDFF93FF13FEACE851FC5F" box="[199,303,336,358]" name="Argentina" pageId="20">Argentina</collectingCountry>
, and Florida Department,
<collectingCountry id="B9E0D6CAFFDDFF93FD93FEACEBDFFC5F" box="[583,673,336,358]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
);—
<bibRefCitation id="A566EBABFFDDFF93FD12FEACEAD5FC5F" author="Garrison" box="[710,939,336,358]" pageId="20" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
Garrison
<emphasis id="F3834A48FFDDFF93FCFFFEADEA1CFC5F" box="[811,866,336,358]" italics="true" pageId="20">et al.</emphasis>
(2006
</bibRefCitation>
: 71, 81–83, 129–130, 329, 331, 337, figs 727, 729a,b, 730, 732, in part, illustrations of mesepisternum in dorsal view and subgenital plate in ventral view of ♀ from Entre Rios Province,
<collectingCountry id="B9E0D6CAFFDDFF93FE74FE71EB76FC9A" box="[416,520,397,419]" name="Argentina" pageId="20">Argentina</collectingCountry>
, metathoracic tibia, and secondary genitalia in lateral view from ♂ of Florida Department,
<collectingCountry id="B9E0D6CAFFDDFF93FF13FE51E85CFCFB" box="[199,290,429,450]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
);—von
<bibRefCitation id="A566EBABFFDDFF93FEA1FE50EDB8FCFB" author="Ellenrieder" box="[373,1222,428,450]" pageId="20" refString="von Ellenrieder, N. &amp; Muzon, J. (2008) An updated checklist of the Odonata from Argentina. Odonatologica, 37 (1), 55 - 68." type="journal article" year="2008">
Ellenrieder &amp; Muzón (2008: 61, in part, record to Entre Rios Province,
<collectingCountry id="B9E0D6CAFFDDFF93FB83FE50EDC1FCFB" box="[1111,1215,428,450]" name="Argentina" pageId="20">Argentina</collectingCountry>
)
</bibRefCitation>
;—von Ellenrieder
<emphasis id="F3834A48FFDDFF93FA5EFE51E99CFCD9" italics="true" pageId="20">et al.</emphasis>
(2009: 229, record from
<collectingCountry id="B9E0D6CAFFDDFF93FE35FE37EB42FCD9" box="[481,572,459,480]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
);—von
<bibRefCitation id="A566EBABFFDDFF93FD5BFE36EAC3FCD9" author="Ellenrieder" box="[655,957,458,480]" pageId="20" refString="von Ellenrieder, N. &amp; Garrison, R. W. (2009) Odonata. In: Dominguez, E. &amp; Fernandez, H. R. (Eds.), Macroinvertebrados bentonicos sudamericanos. Sistematica y biologia. Fundacion Miguel Lillo, Tucuman, pp. 95 - 143." type="book chapter" year="2009">Ellenrieder &amp; Garrison (2009</bibRefCitation>
: 123, fig. 275, reproduction of illustration of ♀ subgenital plate from
<bibRefCitation id="A566EBABFFDDFF93FE70FE15EB0DFCC6" author="Garrison" box="[420,627,489,511]" pageId="20" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
Garrison
<emphasis id="F3834A48FFDDFF93FDD7FE16EB48FCC6" box="[515,566,489,511]" italics="true" pageId="20">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDDFF93FF43FDD0ED35FE24" pageId="20" type="materials_examined">
<paragraph id="C148965AFFDDFF93FF43FDD0EA97FFED" blockId="20.[151,1437,556,2021]" pageId="20">
<emphasis id="F3834A48FFDDFF93FF43FDD0E88CFF7C" bold="true" box="[151,498,556,581]" pageId="20">
Material examined (
<specimenCount id="D7F15DD3FFDDFF93FE58FDD0E8CEFF7C" box="[396,432,556,581]" pageId="20" type="generic">8♂</specimenCount>
, 1♀).
</emphasis>
<collectingCountry id="B9E0D6CAFFDDFF93FE2EFDD0EB1DFF7D" box="[506,611,556,580]" name="Brazil" pageId="20">BRAZIL</collectingCountry>
. Rio Grande do Sul State:
<specimenCount id="D7F15DD3FFDDFF93FC40FDD0EAC6FF7D" box="[916,952,556,580]" pageId="20" type="generic">1♂</specimenCount>
, [Guaíba municipality], Arroio [stream] at BR 116 Km 310 [
<geoCoordinate id="A4C3F09DFFDDFF93FEB6FDB3EB73FF51" box="[354,525,591,616]" direction="south" orientation="latitude" pageId="20" precision="1" value="-30.195915">30°11’45.30”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFDDFF93FDCEFDB3EBB2FF51" box="[538,716,591,616]" direction="west" orientation="longitude" pageId="20" precision="1" value="-51.399048">51°23’56.56”W</geoCoordinate>
,
<quantity id="060F3BBFFFDDFF93FD0EFDACEA6AFF5E" box="[730,788,592,616]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="20" unit="m" value="20.0">20 m</quantity>
a.s.l.],
<date id="B549B09AFFDDFF93FCBDFDACEAE4FF51" box="[873,922,592,616]" pageId="20">17.I.</date>
[19]79, J.M. Costa leg. (
<collectionCode id="A7E60E9FFFDDFF93FB67FDACEC7AFF51" box="[1203,1284,592,616]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20">MNRJ</collectionCode>
);
<specimenCount id="D7F15DD3FFDDFF93FACDFDACEC43FF51" box="[1305,1341,592,616]" pageId="20" type="generic">1♂</specimenCount>
, Pelotas municipality [
<geoCoordinate id="A4C3F09DFFDDFF93FEE2FD88E89CFFB4" box="[310,482,628,653]" direction="south" orientation="latitude" pageId="20" precision="1" value="-31.772">31°46’19.20”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFDDFF93FE24FD88EBDDFFB4" box="[496,675,628,653]" direction="west" orientation="longitude" pageId="20" precision="1" value="-52.3428">52°20’34.08”W</geoCoordinate>
,
<quantity id="060F3BBFFFDDFF93FD65FD88EB93FFB5" box="[689,749,628,652]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="20" unit="m" value="14.0">14 m</quantity>
a.s.l.],
<date id="B549B09AFFDDFF93FC92FD88EAF1FFB4" box="[838,911,628,653]" pageId="20">09.III.</date>
[19]51, C. Biezanko leg. (
<collectionCode id="A7E60E9FFFDDFF93FB6EFD88EC78FFB5" box="[1210,1286,628,652]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20">MNRJ</collectionCode>
22082);
<specimenCount id="D7F15DD3FFDDFF93FAA6FD88ECE9FFB4" box="[1394,1431,628,653]" pageId="20" type="generic">6♂</specimenCount>
, Jaguarão municipality [
<geoCoordinate id="A4C3F09DFFDDFF93FE4AFD6BEB34FF89" box="[414,586,663,688]" direction="south" orientation="latitude" pageId="20" precision="1" value="-32.56585">32°33’57.06”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFDDFF93FD82FD6BEA77FF89" box="[598,777,663,688]" direction="west" orientation="longitude" pageId="20" precision="1" value="-53.376">53°22’33.60”W</geoCoordinate>
,
<quantity id="060F3BBFFFDDFF93FCC0FD64EA30FF96" box="[788,846,664,688]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="20" unit="m" value="19.0">19 m</quantity>
a.s.l.], Stream at
<quantity id="060F3BBFFFDDFF93FBC7FD6BED24FF89" box="[1043,1114,663,688]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="20" unit="km" value="15.0">15 km</quantity>
from highway,
<date id="B549B09AFFDDFF93FAD8FD64ECE6FF89" box="[1292,1432,664,688]" pageId="20" value="1978-12-12">12.XII.1978</date>
, J.M. Costa leg. (
<collectionCode id="A7E60E9FFFDDFF93FE9AFD40E8E1FFED" box="[334,415,700,724]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20">MNRJ</collectionCode>
); 1♀, same data but 12.XIII [sic XII].1978 (
<collectionCode id="A7E60E9FFFDDFF93FC58FD40EAA3FFED" box="[908,989,700,724]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20">MNRJ</collectionCode>
).
</paragraph>
<paragraph id="C148965AFFDDFF93FF13FD23ED35FE24" blockId="20.[151,1437,556,2021]" pageId="20">
<emphasis id="F3834A48FFDDFF93FF13FD23E8F1FFC1" bold="true" box="[199,399,735,760]" pageId="20">
<typeStatus id="1E4C28F8FFDDFF93FF13FD23E87CFFC1" box="[199,258,735,760]" pageId="20">Type</typeStatus>
repository.
</emphasis>
<typeStatus id="1E4C28F8FFDDFF93FE48FD23EB7AFFC1" box="[412,516,735,760]" pageId="20" type="holotype">Holotype</typeStatus>
♂ by original designation (see remarks) at Museo de La Plata, Universidad Nacional de La Plata, La Plata, Buenos Aires,
<collectingCountry id="B9E0D6CAFFDDFF93FD4BFCF8EA70FE24" box="[671,782,772,797]" name="Argentina" pageId="20">Argentina</collectingCountry>
(
<collectionCode id="A7E60E9FFFDDFF93FCC9FCF8EA26FE25" box="[797,856,772,796]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="20">MLP</collectionCode>
, examined by photo).
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDDFF93FF13FCDBED13F9BC" pageId="20" type="description">
<paragraph id="C148965AFFDDFF93FF13FCDBED6AFEE9" blockId="20.[151,1437,556,2021]" pageId="20">
<emphasis id="F3834A48FFDDFF93FF13FCDBE803FE79" bold="true" box="[199,381,807,832]" pageId="20">Measurements.</emphasis>
Males (n = 8). Total length (including caudal appendages) 45.4–47.0; abdomen length (excluding caudal appendages) 30.8–33.9; head maximum width 5.8–6.3; Fw length 25.8–30.5; Hw length 24.7–26.3; Fw maximum width 4.9–5.5, in Hw 6.1–7.0; pt length
<quantity id="060F3BBFFFDDFF93FC63FC93ED50FEB1" box="[951,1070,879,904]" metricMagnitude="-2" metricUnit="m" metricValue="7.493" metricValueMax="8.382" metricValueMin="6.604" pageId="20" unit="in" value="2.95" valueMax="3.3" valueMin="2.6">2.6–3.3 in</quantity>
Fw, 3.0–
<quantity id="060F3BBFFFDDFF93FB76FC93ED96FEB1" box="[1186,1256,879,904]" metricMagnitude="-2" metricUnit="m" metricValue="9.144" pageId="20" unit="in" value="3.6">3.6 in</quantity>
Hw; length of metathoracic femur 7.4–8.2; metathoracic tibia 5.7–6.2; length of S9+
<quantity id="060F3BBFFFDDFF93FC6AFC68EA84FE94" box="[958,1018,916,941]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" unit="in" value="10.0">10 in</quantity>
lateral view 3.2–3.8; total length of cercus in lateral view 1.6–1.9. Ratio cercus length / maximum width 0.30–0.36.
</paragraph>
<paragraph id="C148965AFFDDFF93FF13FC20EBB4F959" blockId="20.[151,1437,556,2021]" pageId="20">
Females (n = 1). Total length (including caudal appendages) 45.2; abdomen length (excluding caudal appendages) 31.8; head maximum width 6.3; Fw length 28.0; Hw length 27.1; Fw maximum width 5.6, in Hw 7.3; pt length
<quantity id="060F3BBFFFDDFF93FED5FBD8E83CF904" box="[257,322,1060,1085]" metricMagnitude="-2" metricUnit="m" metricValue="7.874" pageId="20" unit="in" value="3.1">3.1 in</quantity>
Fw,
<quantity id="060F3BBFFFDDFF93FEADFBD8E8C4F904" box="[377,442,1060,1085]" metricMagnitude="-2" metricUnit="m" metricValue="8.382" pageId="20" unit="in" value="3.3">3.3 in</quantity>
Hw; length of metathoracic femur 8.1; metathoracic tibia 6.2; length of S9+
<quantity id="060F3BBFFFDDFF93FAC1FBD8EC31F904" box="[1301,1359,1060,1085]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" unit="in" value="10.0">10 in</quantity>
lateral view 2.7; total length of cercus in lateral view 1.3.
</paragraph>
<paragraph id="C148965AFFDDFF93FF13FB90ED13F9BC" blockId="20.[151,1437,556,2021]" box="[199,1133,1132,1157]" pageId="20">
<emphasis id="F3834A48FFDDFF93FF13FB90E869F9BC" bold="true" box="[199,279,1132,1157]" pageId="20">Larva.</emphasis>
Rodrigues Capítulo (1985, p. 332) based on specimens reared in laboratory.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDDFF93FF13FB73EBD8FA09" pageId="20" type="diagnosis">
<paragraph id="C148965AFFDDFF93FF13FB73EA11F8D5" blockId="20.[151,1437,556,2021]" pageId="20">
<emphasis id="F3834A48FFDDFF93FF13FB73E844F991" bold="true" box="[199,314,1167,1192]" pageId="20">Diagnosis</emphasis>
. This pale species, as well as
<taxonomicName id="06F7EDD9FFDDFF93FD71FB6DEA64F991" box="[677,794,1169,1192]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDDFF93FD71FB6DEA64F991" box="[677,794,1169,1192]" italics="true" pageId="20">T. uncatus</emphasis>
</taxonomicName>
, can be separated from
<taxonomicName id="06F7EDD9FFDDFF93FBE8FB73ED94F991" box="[1084,1258,1167,1192]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDDFF93FBE8FB73ED94F991" box="[1084,1258,1167,1192]" italics="true" pageId="20">Cyanogomphus</emphasis>
</taxonomicName>
spp. except
<taxonomicName id="06F7EDD9FFDDFF93FA56FB6CE855F9F4" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFDDFF93FA56FB6CE855F9F4" italics="true" pageId="20">C. comparabilis</emphasis>
</taxonomicName>
by the mesepisternum close to the dorsal carina largely pale (dark in
<taxonomicName id="06F7EDD9FFDDFF93FBE6FB49EC73F9F4" box="[1074,1293,1204,1229]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFDDFF93FBE6FB49EC73F9F4" box="[1074,1293,1204,1229]" italics="true" pageId="20">C. angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFDDFF93FAC0FB48EC13F9F4" bold="true" box="[1300,1389,1204,1229]" pageId="20">
<taxonomicNameLabel id="E8B0F733FFDDFF93FAC0FB48EC13F9F4" box="[1300,1389,1204,1229]" pageId="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="06F7EDD9FFDDFF93FF43FB24E866F9C9" box="[151,280,1239,1264]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDDFF93FF43FB24E866F9C9" box="[151,280,1239,1264]" italics="true" pageId="20">C. waltheri</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDDFF93FEFDFB2BE80CF9C9" box="[297,370,1239,1264]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="20">Figs 1</figureCitation>
,
<figureCitation id="59CC8ADFFFDDFF93FE57FB24E8E0F9C9" box="[387,414,1240,1264]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="20">10</figureCitation>
A–B, D, 11A, C). The narrow cornet-like distal portion of
<date id="B549B09AFFDDFF93FB84FB24ED0DF9C9" box="[1104,1139,1240,1264]" pageId="20">
V
<quantity id="060F3BBFFFDDFF93FBB1FB24ED0DF9C9" box="[1125,1139,1240,1264]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="20" unit="in" value="4.0">4</quantity>
</date>
in ventral view (
<figureCitation id="59CC8ADFFFDDFF93FAEAFB2BECFDF9C9" box="[1342,1411,1239,1264]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="20">Fig. 6</figureCitation>
G, maximum width &lt;0.6 of
<date id="B549B09AFFDDFF93FE62FB00E8A6F82D" box="[438,472,1276,1300]" pageId="20">V3</date>
width vs.&gt;
<quantity id="060F3BBFFFDDFF93FDB1FB00EBCDF82C" box="[613,691,1276,1301]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" pageId="20" unit="in" value="0.75">0.75 in</quantity>
<taxonomicName id="06F7EDD9FFDDFF93FD6EFB00EA16F82C" box="[698,872,1276,1301]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDDFF93FD6EFB00EA16F82C" box="[698,872,1276,1301]" italics="true" pageId="20">Cyanogomphus</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDDFF93FCA2FB00EAC2F82C" box="[886,956,1276,1301]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="20">Figs 6</figureCitation>
B, E), blade-like cercus distinctly flattened with apex directed posteriorly (
<figureCitation id="59CC8ADFFFDDFF93FE20FAE3EB44F801" box="[500,570,1311,1336]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="20">Figs 7</figureCitation>
J–L, 8G–H, cercus cylindrical, curved toward to the upturned tapered apex in
<taxonomicName id="06F7EDD9FFDDFF93FF43FAB8E83BF864" box="[151,325,1348,1373]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDDFF93FF43FAB8E83BF864" box="[151,325,1348,1373]" italics="true" pageId="20">Cyanogomphus</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDDFF93FE80FAB8E8E4F865" box="[340,410,1348,1373]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="20">Figs 7</figureCitation>
A–I, 8A–F), the dorsally smooth epiproct, lacking a distal concavity on the epiproct (
<figureCitation id="59CC8ADFFFDDFF93FABEFAB8E9DAF8B9" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="20">Figs 8</figureCitation>
H, epiproct with a distal concavity in
<taxonomicName id="06F7EDD9FFDDFF93FD98FA9BEB84F8B9" box="[588,762,1383,1408]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDDFF93FD98FA9BEB84F8B9" box="[588,762,1383,1408]" italics="true" pageId="20">Cyanogomphus</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDDFF93FCDCFA9BEA30F8B9" box="[776,846,1383,1408]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="20">Figs 8</figureCitation>
B, D, F) of males, and subtriangular, long subgenital plate (length&gt; 0.5 of S9) with a narrow and deep mesal incision (
<figureCitation id="59CC8ADFFFDDFF93FCA4FA70EACCF89D" box="[880,946,1420,1445]" captionStart="FIGURE 9" captionStartId="18.[151,250,1160,1182]" captionTargetBox="[151,1434,193,1138]" captionTargetId="figure@18.[151,1436,193,1139]" captionTargetPageId="18" captionText="FIGURE 9. Female S 8 – 10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A – B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C. Cyanogomphus angelomachadoi sp. nov. additional specimen (Brazil. MG: Serra do Salitre, ABMM); D. Cyanogomphus sp. (Brazil. GO: Jataí, ABMM); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm." httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="20">Fig. 9</figureCitation>
E, subgenital plate short, length &lt;0.4 of S9, not incised, with a wide V- or U-shaped posteromesal concavity in
<taxonomicName id="06F7EDD9FFDDFF93FC16FA53ED0EF8F1" box="[962,1136,1455,1480]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDDFF93FC16FA53ED0EF8F1" box="[962,1136,1455,1480]" italics="true" pageId="20">Cyanogomphus</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDDFF93FB57FA53EDB3F8F1" box="[1155,1229,1455,1480]" captionStart="FIGURE 9" captionStartId="18.[151,250,1160,1182]" captionTargetBox="[151,1434,193,1138]" captionTargetId="figure@18.[151,1436,193,1139]" captionTargetPageId="18" captionText="FIGURE 9. Female S 8 – 10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A – B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C. Cyanogomphus angelomachadoi sp. nov. additional specimen (Brazil. MG: Serra do Salitre, ABMM); D. Cyanogomphus sp. (Brazil. GO: Jataí, ABMM); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm." httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="20">Figs 9</figureCitation>
A–D) of females, separates both species of
<taxonomicName id="06F7EDD9FFDDFF93FE67FA28EB2CF8D4" box="[435,594,1492,1517]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDDFF93FE67FA28EB2CF8D4" box="[435,594,1492,1517]" italics="true" pageId="20">Tibiagomphus</emphasis>
</taxonomicName>
from all
<taxonomicName id="06F7EDD9FFDDFF93FD6DFA28EA19F8D4" box="[697,871,1492,1517]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDDFF93FD6DFA28EA19F8D4" box="[697,871,1492,1517]" italics="true" pageId="20">Cyanogomphus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C148965AFFDDFF93FF13FA0BEBD8FA09" blockId="20.[151,1437,556,2021]" pageId="20">
Both sexes of
<taxonomicName id="06F7EDD9FFDDFF93FEB1FA05E8C1FB29" box="[357,447,1527,1552]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDDFF93FEB1FA05E8C1FB29" box="[357,447,1527,1552]" italics="true" pageId="20">T. noval</emphasis>
</taxonomicName>
can be distinguished from it sibling congener
<taxonomicName id="06F7EDD9FFDDFF93FC10FA05ED4AFB29" box="[964,1076,1529,1552]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDDFF93FC10FA05ED4AFB29" box="[964,1076,1529,1552]" italics="true" pageId="20">T. uncatus</emphasis>
</taxonomicName>
by its paler generally yellowish color (
<figureCitation id="59CC8ADFFFDDFF93FF34F9E0E848FB0D" box="[224,310,1564,1589]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="20">Figs 10</figureCitation>
E, 11D, distinctly darker in
<taxonomicName id="06F7EDD9FFDDFF93FDA5F9E1EB9AFB0D" box="[625,740,1565,1588]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDDFF93FDA5F9E1EB9AFB0D" box="[625,740,1565,1588]" italics="true" pageId="20">T. uncatus</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDDFF93FD27F9E0EA3AFB0D" box="[755,836,1564,1589]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="20">Fig. 10</figureCitation>
F), and by dorsally pale metathoracic tibia and tarsus (
<figureCitation id="59CC8ADFFFDDFF93FF4BF9C3E99FFB61" box="[159,225,1599,1624]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="20">Fig. 3</figureCitation>
C, dark in
<taxonomicName id="06F7EDD9FFDDFF93FE82F9BDE8B6FB61" box="[342,456,1601,1624]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDDFF93FE82F9BDE8B6FB61" box="[342,456,1601,1624]" italics="true" pageId="20">T. uncatus</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDDFF93FE02F9C3EB69FB61" box="[470,535,1599,1624]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="20">Fig. 3</figureCitation>
D). The posterior hamule varies greatly, being longer and slender with the “heel” distinctly less pronounced in
<taxonomicName id="06F7EDD9FFDDFF93FE32F999EB3CFB44" box="[486,578,1636,1661]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDDFF93FE32F999EB3CFB44" box="[486,578,1636,1661]" italics="true" pageId="20">T. noval</emphasis>
</taxonomicName>
(
<figureCitation id="59CC8ADFFFDDFF93FD87F998EBE2FB45" box="[595,668,1636,1661]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="20">Figs 4</figureCitation>
E, 5F–H, shorter and stouter in
<taxonomicName id="06F7EDD9FFDDFF93FBDCF999ED04FB45" box="[1032,1146,1637,1660]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDDFF93FBDCF999ED04FB45" box="[1032,1146,1637,1660]" italics="true" pageId="20">T. uncatus</emphasis>
</taxonomicName>
with a more pronounced “heel”,
<figureCitation id="59CC8ADFFFDDFF93FF3BF97BE849FB99" box="[239,311,1671,1696]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="20">Figs 4</figureCitation>
F,
<date id="B549B09AFFDDFF93FE81F97BE812FB99" box="[341,364,1671,1696]" pageId="20">5I</date>
–L), male cercus robust (
<figureCitation id="59CC8ADFFFDDFF93FD5AF97BEBA9FB99" box="[654,727,1671,1696]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="20">Figs 7</figureCitation>
G, 8G–H, ratio cercus length / maximum width 0.30–0.36 vs. more slender,
<quantity id="060F3BBFFFDDFF93FEEAF950E8B1FBFC" box="[318,463,1708,1733]" metricMagnitude="-3" metricUnit="m" metricValue="6.985" metricValueMax="7.62" metricValueMin="6.35" pageId="20" unit="in" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.30 in</quantity>
<taxonomicName id="06F7EDD9FFDDFF93FE0EF951EB2FFBFD" box="[474,593,1709,1732]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDDFF93FE0EF951EB2FFBFD" box="[474,593,1709,1732]" italics="true" pageId="20">T. uncatus</emphasis>
</taxonomicName>
,
<figureCitation id="59CC8ADFFFDDFF93FDB0F950EBD3FBFD" box="[612,685,1708,1733]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="20">Figs 7</figureCitation>
M,
<date id="B549B09AFFDDFF93FD0DF950EB8EFBFD" box="[729,752,1708,1732]" pageId="20">8I</date>
–J). Further, the smaller size of
<taxonomicName id="06F7EDD9FFDDFF93FBB9F951EDB7FBFC" box="[1133,1225,1708,1733]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDDFF93FBB9F951EDB7FBFC" box="[1133,1225,1708,1733]" italics="true" pageId="20">T. noval</emphasis>
</taxonomicName>
(male Hw length 24.7–26.3 vs. 26.8–29.7 of
<taxonomicName id="06F7EDD9FFDDFF93FE09F92DEB2CFBD1" box="[477,594,1745,1768]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDDFF93FE09F92DEB2CFBD1" box="[477,594,1745,1768]" italics="true" pageId="20">T. uncatus</emphasis>
</taxonomicName>
) and allopatric distribution, with
<taxonomicName id="06F7EDD9FFDDFF93FC30F92DED3CFBD1" box="[996,1090,1743,1768]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDDFF93FC30F92DED3CFBD1" box="[996,1090,1743,1768]" italics="true" pageId="20">T. noval</emphasis>
</taxonomicName>
occurring in biogeographical Pampean province of Chacoan dominion and
<taxonomicName id="06F7EDD9FFDDFF93FD61F909EA55FA35" box="[693,811,1781,1804]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="20" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDDFF93FD61F909EA55FA35" box="[693,811,1781,1804]" italics="true" pageId="20">T. uncatus</emphasis>
</taxonomicName>
in Araucaria and Parana Forest provinces in Parana dominion (
<figureCitation id="59CC8ADFFFDDFF93FEC6F8EBE81AFA09" box="[274,356,1815,1840]" captionStart="FIGURE 13" captionStartId="25.[151,250,1528,1550]" captionTargetBox="[159,1434,488,1498]" captionTargetId="figure@25.[151,1436,483,1503]" captionTargetPageId="25" captionText="FIGURE 13. Map of southeastern South America with distribution records of Tibiagomphus Belle, 1992, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: ER, Entre Rios and MI, Misiones for Argentine provinces; PR, Paraná, RS, Rio Grande do Sul, and SC, Santa Catarina for Brazilian States." httpUri="https://zenodo.org/record/270866/files/figure.png" pageId="20">Fig. 13</figureCitation>
), separate these two species.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDDFF93FF13F8C0EC00FADC" pageId="20" type="distribution">
<paragraph id="C148965AFFDDFF93FF13F8C0ED0BFA41" blockId="20.[151,1437,556,2021]" pageId="20">
<emphasis id="F3834A48FFDDFF93FF13F8C0E827FA6C" bold="true" box="[199,345,1852,1877]" pageId="20">Distribution</emphasis>
. From Rio Grande do Sul State,
<collectingCountry id="B9E0D6CAFFDDFF93FD30F8C0EA55FA6C" box="[740,811,1852,1877]" name="Brazil" pageId="20">Brazil</collectingCountry>
, and Entre Rios Province,
<collectingCountry id="B9E0D6CAFFDDFF93FBBCF8C0EDA6FA6C" box="[1128,1240,1852,1877]" name="Argentina" pageId="20">Argentina</collectingCountry>
south to Florida Department,
<collectingCountry id="B9E0D6CAFFDDFF93FEFDF89DE8F0FA40" box="[297,398,1889,1913]" name="Uruguay" pageId="20">Uruguay</collectingCountry>
; an inhabitant of the Pampean biogeographical province (
<figureCitation id="59CC8ADFFFDDFF93FBC0F8A3ED19FA41" box="[1044,1127,1887,1912]" captionStart="FIGURE 13" captionStartId="25.[151,250,1528,1550]" captionTargetBox="[159,1434,488,1498]" captionTargetId="figure@25.[151,1436,483,1503]" captionTargetPageId="25" captionText="FIGURE 13. Map of southeastern South America with distribution records of Tibiagomphus Belle, 1992, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: ER, Entre Rios and MI, Misiones for Argentine provinces; PR, Paraná, RS, Rio Grande do Sul, and SC, Santa Catarina for Brazilian States." httpUri="https://zenodo.org/record/270866/files/figure.png" pageId="20">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="C148965AFFDDFF93FF13F878EC00FADC" blockId="20.[151,1437,556,2021]" pageId="20">
<emphasis id="F3834A48FFDDFF93FF13F878EB4FFAA4" bold="true" box="[199,561,1924,1949]" pageId="20">Biological and ecological data.</emphasis>
Adults are found in lotic environments such as riffles and deep zones in slow streams surrounded by gallery forest, where adults perch in protected sites, from sea level to 107 meters of elevation. Larvae are shallow burrowers and were found with some adhered detritus (Rodrigues Capítulo 1985).
</paragraph>
</subSubSection>
<caption id="9588C6D2FFDCFF92FF43F992EC40FBC6" httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="21" targetBox="[151,1436,195,1620]" targetPageId="21">
<paragraph id="C148965AFFDCFF92FF43F992EC40FBC6" blockId="21.[151,1436,1646,1791]" pageId="21">
<emphasis id="F3834A48FFDCFF92FF43F992E865FBBA" bold="true" box="[151,283,1646,1668]" pageId="21">FIGURE 10</emphasis>
. Male habitus of Cyanogomphini in lateral view. A–B.
<taxonomicName id="06F7EDD9FFDCFF92FCB7F992EDCCFBBD" box="[867,1202,1646,1668]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFDCFF92FCB7F992EDCCFBBD" box="[867,1202,1646,1668]" italics="true" pageId="21">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFDCFF92FB6EF993EC73FBBD" bold="true" box="[1210,1293,1647,1668]" pageId="21">
<taxonomicNameLabel id="E8B0F733FFDCFF92FB6EF993EC73FBBD" box="[1210,1293,1647,1668]" pageId="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C.
<taxonomicName id="06F7EDD9FFDCFF92FF43F950EB39FBFB" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[151,583,1708,1730]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="comparabilis">
<emphasis id="F3834A48FFDCFF92FF43F950E8BAFBFB" box="[151,452,1708,1730]" italics="true" pageId="21">Cyanogomphus comparabilis</emphasis>
Belle, 1994
</taxonomicName>
, Holotype (Brazil. MT: Diamantino, ABMM); D.
<taxonomicName id="06F7EDD9FFDCFF92FB8EF950E9B2FBD9" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDCFF92FB8EF950EC2BFBFB" box="[1114,1365,1708,1730]" italics="true" pageId="21">Cyanogomphus waltheri</emphasis>
Selys, 1873
</taxonomicName>
, (Brazil. RJ: Santana de Japuíba district, MNRJ); E.
<taxonomicName id="06F7EDD9FFDCFF92FCDAF936EC71FBD9" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[782,1295,1738,1760]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDCFF92FCDAF936EA9FFBD9" box="[782,993,1738,1760]" italics="true" pageId="21">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</taxonomicName>
, (Brazil. RS: Jaguarão, MNRJ); F.
<taxonomicName id="06F7EDD9FFDCFF92FEBBF915EB95FBC6" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[367,747,1769,1791]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDCFF92FEBBF915EB2AFBC6" box="[367,596,1769,1791]" italics="true" pageId="21">Tibiagomphus uncatus</emphasis>
(Fraser, 1947)
</taxonomicName>
, (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm.
</paragraph>
</caption>
<subSubSection id="89EDC5D1FFDCFF91FF13F8DAECE2F945" lastPageId="22" pageId="21" type="discussion">
<paragraph id="C148965AFFDCFF91FF13F8DAE8DFFDC1" blockId="21.[151,1436,1830,2035]" lastBlockId="22.[151,1436,151,1149]" lastPageId="22" pageId="21">
<emphasis id="F3834A48FFDCFF92FF13F8DAE845FA06" bold="true" box="[199,315,1830,1855]" pageId="21">Remarks.</emphasis>
The typification by Rodrigues Capítulo (1985) is misleading and imprecise. In describing
<taxonomicName id="06F7EDD9FFDCFF92FAE8F8DBECE2FA06" box="[1340,1436,1830,1855]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="21" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDCFF92FAE8F8DBECE2FA06" box="[1340,1436,1830,1855]" italics="true" pageId="21">C. noval</emphasis>
</taxonomicName>
he designated an adult male emerged in the laboratory as
<typeStatus id="1E4C28F8FFDCFF92FC9DF8B6EAD5FA5A" box="[841,939,1866,1891]" pageId="21" type="holotype">holotype</typeStatus>
without mention of its exuvia (Rodrigues Capítulo 1985, p. 329). Furthermore, the other cited specimens, an adult female and six larvae, were not designated as
<typeStatus id="1E4C28F8FFDCFF92FF61F86FE85DFA92" box="[181,291,1939,1963]" pageId="21" type="paratype">paratypes</typeStatus>
, therefore this meets the requirement that “expressly excludes them from the
<typeStatus id="1E4C28F8FFDCFF92FB56F86FEDCCFA92" box="[1154,1202,1939,1963]" pageId="21">type</typeStatus>
series” (ICZN 1999, Art. 72.4.6), even though an allotype was mentioned in the abstract. The subsequent citation of the female as allotype by Muzón
<emphasis id="F3834A48FFDCFF92FEA1F820E8D0FACA" box="[373,430,2010,2035]" italics="true" pageId="21">et al.</emphasis>
(2007) should be considered an invalid nomenclatural act. The date of publication of all papers in that volume is stated to be 1984, but the cover states 1985. For this reason, without further investigation, we follow
<bibRefCitation id="A566EBABFFDFFF91FEC4FF40EB7BFDEC" box="[272,517,188,213]" pageId="22" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book">
Garrison
<emphasis id="F3834A48FFDFFF91FEAEFF41E8CDFDEC" box="[378,435,188,213]" italics="true" pageId="22">et al.</emphasis>
(2006)
</bibRefCitation>
considering 1985 as the correct year of publication, while the respective day and month still are pending.
</paragraph>
<paragraph id="C148965AFFDFFF91FF13FEF9E9A1F90D" blockId="22.[151,1436,151,1149]" pageId="22">
Except by the paler color (“pardo amarillenta”), all characters mentioned by Rodrigues Capítulo (1985, p. 329–332) are not diagnostic of
<taxonomicName id="06F7EDD9FFDFFF91FDDEFED5EB1BFC79" box="[522,613,295,320]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDFFF91FDDEFED5EB1BFC79" box="[522,613,295,320]" italics="true" pageId="22">T. noval</emphasis>
</taxonomicName>
in accordance with its new status recognized here, and his characters should be considered as examples of individual variation, as they are shared with some
<taxonomicName id="06F7EDD9FFDFFF91FB75FEB1EC6BFC5D" box="[1185,1301,333,356]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FB75FEB1EC6BFC5D" box="[1185,1301,333,356]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
. Rodrigues Capítulo (1985) provided no diagnosis, instead comparing a few selected characters with
<taxonomicName id="06F7EDD9FFDFFF91FB55FE8DED8DFCB1" box="[1153,1267,369,392]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FB55FE8DED8DFCB1" box="[1153,1267,369,392]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
as follows: (1) anterior margin of the postfrons in dorsal view sinuous with a small mesal convexity, while in
<taxonomicName id="06F7EDD9FFDFFF91FB3BFE69EC1DFC95" box="[1263,1379,405,428]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FB3BFE69EC1DFC95" box="[1263,1379,405,428]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
it is entirely convex; (2) male S9 with a V-shaped mesodorsal invagination vs. lacking invagination in
<taxonomicName id="06F7EDD9FFDFFF91FB23FE45EC15FCE9" box="[1271,1387,441,464]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FB23FE45EC15FCE9" box="[1271,1387,441,464]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
; (3) spines along posterior carina of S9 extend downward two-thirds of the segment height in the male vs. only over dorsal surface in
<taxonomicName id="06F7EDD9FFDFFF91FEB6FDFDE8A9FF21" box="[354,471,513,536]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FEB6FDFDE8A9FF21" box="[354,471,513,536]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
; (4) posterior margin of male epiproct almost straight vs. ovoid in
<taxonomicName id="06F7EDD9FFDFFF91FB26FDFDEC19FF21" box="[1266,1383,513,536]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FB26FDFDEC19FF21" box="[1266,1383,513,536]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
; (5) female distal margin of sternites of S9–10 distinct from
<taxonomicName id="06F7EDD9FFDFFF91FCECFDD9EAD0FF05" box="[824,942,549,572]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FCECFDD9EAD0FF05" box="[824,942,549,572]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
. The shape of the anterior margin of the postfrons (character 1) is variable, specimens of
<taxonomicName id="06F7EDD9FFDFFF91FD1CFDB5EA5CFF59" box="[712,802,583,608]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDFFF91FD1CFDB5EA5CFF59" box="[712,802,583,608]" italics="true" pageId="22">T. noval</emphasis>
</taxonomicName>
collected from the same site and day showing either a small mesal convexity (
<figureCitation id="59CC8ADFFFDFFF91FE71FD90E897FFBC" box="[421,489,620,645]" captionStart="FIGURE 5" captionStartId="11.[151,250,1535,1557]" captionTargetBox="[151,1436,195,1507]" captionTargetId="figure@11.[151,1436,195,1511]" captionTargetPageId="11" captionText="FIGURE 5. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A – D) and posterior hamule in lateral view (E – L). A – B, F – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); C – D, I – L. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). E. Cyanogomphus comparabilis Belle, 1994, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="22">Fig. 5</figureCitation>
B) or regularly convex postfrons (
<figureCitation id="59CC8ADFFFDFFF91FCB8FD90EAD1FFBC" box="[876,943,620,645]" captionStart="FIGURE 5" captionStartId="11.[151,250,1535,1557]" captionTargetBox="[151,1436,195,1507]" captionTargetId="figure@11.[151,1436,195,1511]" captionTargetPageId="11" captionText="FIGURE 5. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A – D) and posterior hamule in lateral view (E – L). A – B, F – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); C – D, I – L. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). E. Cyanogomphus comparabilis Belle, 1994, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="22">Fig. 5</figureCitation>
A) such as illustrated for the
<typeStatus id="1E4C28F8FFDFFF91FB2AFD90EC1EFFBC" box="[1278,1376,620,645]" pageId="22" type="holotype">holotype</typeStatus>
of
<taxonomicName id="06F7EDD9FFDFFF91FA53FD91E990FF91" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FA53FD91E990FF91" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
by Rodrigues Capítulo (1985, fig. 1). Additionally, variation in the postfrons in
<taxonomicName id="06F7EDD9FFDFFF91FB58FD6DEC7FFF91" box="[1164,1281,657,680]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FB58FD6DEC7FFF91" box="[1164,1281,657,680]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
also shows a slightly mesal projection along the anterior margin of postfrons (
<figureCitation id="59CC8ADFFFDFFF91FCA0FD48EAC5FFF4" box="[884,955,692,717]" captionStart="FIGURE 5" captionStartId="11.[151,250,1535,1557]" captionTargetBox="[151,1436,195,1507]" captionTargetId="figure@11.[151,1436,195,1511]" captionTargetPageId="11" captionText="FIGURE 5. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A – D) and posterior hamule in lateral view (E – L). A – B, F – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); C – D, I – L. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). E. Cyanogomphus comparabilis Belle, 1994, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="22">Figs 5</figureCitation>
C–D). A small mesodorsal invagination in S9 (character 2) is common to all Cyanogomphini including
<taxonomicName id="06F7EDD9FFDFFF91FC91FD25EAC6FFC9" box="[837,952,729,752]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FC91FD25EAC6FFC9" box="[837,952,729,752]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
(
<figureCitation id="59CC8ADFFFDFFF91FC1CFD2BED6EFFD6" box="[968,1040,727,752]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="22">Figs 7</figureCitation>
B, E, H, K, N). We did not find the shape of invagination to be a diagnostic character. The distribution of spines along S9 (character 3) also is variable among specimens, and in both
<taxonomicName id="06F7EDD9FFDFFF91FE27FCDDEB35FE01" box="[499,587,799,824]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDFFF91FE27FCDDEB35FE01" box="[499,587,799,824]" italics="true" pageId="22">T. noval</emphasis>
</taxonomicName>
and
<taxonomicName id="06F7EDD9FFDFFF91FD50FCDDEB88FE01" box="[644,758,801,824]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FD50FCDDEB88FE01" box="[644,758,801,824]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
they can extend down two-thirds of the segment height (see
<figureCitation id="59CC8ADFFFDFFF91FF43FCB8E9A3FE65" box="[151,221,836,861]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="22">Figs 7</figureCitation>
J, M). This variation might have lead J. Belle to misidentify some
<taxonomicName id="06F7EDD9FFDFFF91FC6AFCB9ED6BFE64" box="[958,1045,836,861]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDFFF91FC6AFCB9ED6BFE64" box="[958,1045,836,861]" italics="true" pageId="22">T. noval</emphasis>
</taxonomicName>
as
<taxonomicName id="06F7EDD9FFDFFF91FBE8FCB9EDD3FE65" box="[1084,1197,837,860]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FBE8FCB9EDD3FE65" box="[1084,1197,837,860]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
because they had the spines extending to the ventral half. Even though in
<taxonomicName id="06F7EDD9FFDFFF91FD35FC95EA47FEB9" box="[737,825,871,896]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDFFF91FD35FC95EA47FEB9" box="[737,825,871,896]" italics="true" pageId="22">T. noval</emphasis>
</taxonomicName>
the distal margin of the epiproct (character 4) appears be straighter than that of
<taxonomicName id="06F7EDD9FFDFFF91FE68FC71EB4EFE9D" box="[444,560,909,932]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FE68FC71EB4EFE9D" box="[444,560,909,932]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
(
<figureCitation id="59CC8ADFFFDFFF91FD96FC70EBF2FE9D" box="[578,652,908,933]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="22">Figs 7</figureCitation>
J, L, M,
<collectingCountry id="B9E0D6CAFFDFFF91FD25FC70EA79FE9D" box="[753,775,908,932]" name="Iceland" pageId="22">O</collectingCountry>
, 8G–J), it is not ovoid in
<taxonomicName id="06F7EDD9FFDFFF91FBE1FC71EDD4FE9D" box="[1077,1194,909,932]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FBE1FC71EDD4FE9D" box="[1077,1194,909,932]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
, and this structure is similar for both species. Finally, character 5 on female sternites of S9–10 cannot be verified, as differences were not depicted. Although Rodrigues Capítulo (1985) borrowed the
<typeStatus id="1E4C28F8FFDFFF91FCBDFC28EAB5FED4" box="[873,971,980,1005]" pageId="22" type="holotype">holotype</typeStatus>
of
<taxonomicName id="06F7EDD9FFDFFF91FC24FC29ED1EFED5" box="[1008,1120,981,1004]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FC24FC29ED1EFED5" box="[1008,1120,981,1004]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
, he adopted the illustrations by
<bibRefCitation id="A566EBABFFDFFF91FF69FC04E829F929" author="Fraser" box="[189,343,1016,1040]" pageId="22" refString="Fraser, F. C. (1947) The Odonata of the Argentine Republic I. Acta Zoologica Lilloana del Instituto Miguel Lillo, 4, 427 - 462." type="journal article" year="1947">Fraser (1947)</bibRefCitation>
for his comparisons, possibly due to the poor condition of the
<typeStatus id="1E4C28F8FFDFFF91FBE0FC0BEDE8F929" box="[1076,1174,1015,1040]" pageId="22" type="holotype">holotype</typeStatus>
(S4–10 lost, see
<bibRefCitation id="A566EBABFFDFFF91FAB5FC0BE9ADF90D" author="Belle" pageId="22" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle 1980</bibRefCitation>
).
</paragraph>
<paragraph id="C148965AFFDFFF91FF13FBC3ECE2F945" blockId="22.[151,1436,151,1149]" pageId="22">
The males from municipalities of Guaíba and Pelotas were determined as
<taxonomicName id="06F7EDD9FFDFFF91FBEEFBBDEDE9F961" box="[1082,1175,1087,1112]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDFFF91FBEEFBBDEDE9F961" box="[1082,1175,1087,1112]" italics="true" pageId="22">T. noval</emphasis>
</taxonomicName>
, while a couple from Jaguarão municipality, all from Rio Grande do Sul State,
<collectingCountry id="B9E0D6CAFFDFFF91FCCCFB98EA21F944" box="[792,863,1124,1149]" name="Brazil" pageId="22">Brazil</collectingCountry>
, were determined as
<taxonomicName id="06F7EDD9FFDFFF91FB9EFB99EDC2F945" box="[1098,1212,1125,1148]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FB9EFB99EDC2F945" box="[1098,1212,1125,1148]" italics="true" pageId="22">T. uncatus</emphasis>
</taxonomicName>
by J. Belle in 1992.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="06F7EDD9FFDFFF91FF43FB2CEB21F9D3" ID-CoL="7BYMN" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[151,607,1232,1258]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FF43FB2CEB21F9D3" bold="true" box="[151,607,1232,1258]" pageId="22">
<emphasis id="F3834A48FFDFFF91FF43FB2CE8D5F9D3" bold="true" box="[151,427,1232,1258]" italics="true" pageId="22">Tibiagomphus uncatus</emphasis>
(
<bibRefCitation id="A566EBABFFDFFF91FE6EFB2CEB29F9D3" author="Fraser" box="[442,599,1232,1258]" pageId="22" refString="Fraser, F. C. (1947) The Odonata of the Argentine Republic I. Acta Zoologica Lilloana del Instituto Miguel Lillo, 4, 427 - 462." type="journal article" year="1947">Fraser, 1947</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C148965AFFDFFF91FF43FB0EEB89F812" blockId="22.[151,978,1232,1323]" pageId="22">
LSID urn:lsid:zoobank.org:act:4A858689-3BB8-4F35-B74F-9313F7FF2E6E (
<figureCitation id="59CC8ADFFFDFFF91FF4AFAEEE877F812" box="[158,265,1298,1323]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="22">Figures 2</figureCitation>
F, 3D, 4F, 5C–D, I–L, 7M–
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,
<date id="B549B09AFFDFFF91FD89FAEFEB0DF812" box="[605,627,1299,1323]" pageId="22">8I</date>
–J, 10F, 13)
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDFFF91FF43FAA6ED35FAEF" pageId="22" type="reference_group">
<paragraph id="C148965AFFDFFF91FF43FAA6EC0BFBFB" blockId="22.[151,1436,1370,2006]" pageId="22">
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<taxonomicName id="06F7EDD9FFDFFF91FF43FAA6EB35F849" ID-CoL="9WYQS" authority="Fraser, 1947: 437" authorityName="Fraser" authorityPageNumber="437" authorityYear="1947" box="[151,587,1370,1392]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FF43FAA6E8F2F849" box="[151,396,1370,1392]" italics="true" pageId="22">Cyanogomphus uncatus</emphasis>
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: 437
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–439, figs 1a–c (description of ♂
<typeStatus id="1E4C28F8FFDFFF91FC79FAA6ED78F849" box="[941,1030,1370,1392]" pageId="22" type="holotype">holotype</typeStatus>
and allotype,
<collectingCountry id="B9E0D6CAFFDFFF91FB48FAA6EC4DF849" box="[1180,1331,1370,1392]" name="Argentina" pageId="22">ARGENTINA</collectingCountry>
. Misiones Province: Iguazú Department, Puerto Bemberg,
<date id="B549B09AFFDFFF91FD78FA85EA4BF8B6" box="[684,821,1401,1423]" pageId="22" value="1945-01-12" valueMax="1945-01-29" valueMin="1945-01-12">12–29.I.1945</date>
, Hayward, Willink &amp; Goldbach leg. in IFML, illustrations of caudal appendages in ventral, dorsal and lateral views of ♂
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;—
<bibRefCitation id="A566EBABFFDFFF91FC0AFA64E85CF8F5" author="Belle" pageId="22" refString="Belle, J. (1966) Surinam dragon-flies of the Agriogomphus complex of genera. Studies on the fauna of Suriname and other Guyanas, 8 (29), 29 - 60." type="journal article" year="1966">
Belle (1966: 29, 31, comparison with
<taxonomicName id="06F7EDD9FFDFFF91FA50FA65E865F8F5" ID-CoL="32NMT" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDFFF91FA50FA65E865F8F5" italics="true" pageId="22">C. waltheri</emphasis>
</taxonomicName>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDFFF91FE96FA4AED82F8D2" author="Belle" pageId="22" refString="Belle, J. (1970) Studies on South American Gomphidae (Odonata) with special reference to the species from Surinam. Studies on the fauna of Suriname and other Guyanas, 11 (43), 1 - 158, pls. XIX." type="journal article" year="1970">
Belle (1970: 14, 21, 29–30, figs 26, 52, pl. Va, comparison with
<taxonomicName id="06F7EDD9FFDFFF91FC04FA4BED4AF8F4" ID-CoL="B7QB" box="[976,1076,1463,1485]" class="Insecta" family="Gomphidae" genus="Agriogomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="tumens">
<emphasis id="F3834A48FFDFFF91FC04FA4BED4AF8F4" box="[976,1076,1463,1485]" italics="true" pageId="22">A. tumens</emphasis>
</taxonomicName>
and
<taxonomicName id="06F7EDD9FFDFFF91FBB1FA4AED96F8F5" ID-CoL="9WVZC" box="[1125,1256,1462,1484]" class="Insecta" family="Gomphidae" genus="Ebegomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDFFF91FBB1FA4AED96F8F5" box="[1125,1256,1462,1484]" italics="true" pageId="22">Ebegomphus</emphasis>
</taxonomicName>
spp., illustrations of synthorax, S
<quantity id="060F3BBFFFDFFF91FEB7FA29E8CCF8D2" box="[355,434,1493,1515]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="22" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">8–10 in</quantity>
ventral view, and photo of the wings of ♀ from Rio Grande do Sul State,
<collectingCountry id="B9E0D6CAFFDFFF91FB62FA29ED88F8D2" box="[1206,1270,1493,1515]" name="Brazil" pageId="22">Brazil</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDFFF91FACFFA29EBDDFB11" author="Belle" pageId="22" refString="Belle, J. (1972) Further studies on South American Gomphidae (Odonata). Tijdschrift voor Entomologie, 115 (5), 217 - 240." type="journal article" year="1972" yearSuffix="a">
Belle (1972: 219, 221, figs 5–6, illustrations of ♀ subgenital plates from Nova Teutônia village, Santa Catarina State and comparison with a ♀ from Rio Grande do Sul State,
<collectingCountry id="B9E0D6CAFFDFFF91FD89F9EEEBE2FB11" box="[605,668,1554,1576]" name="Brazil" pageId="22">Brazil</collectingCountry>
)
</bibRefCitation>
;—
<treatmentCitation id="4056B04BFFDFFF91FD15F9EEEAF4FB11" author="Paulson" box="[705,906,1554,1576]" page="175" pageId="22" year="1977">
<bibRefCitation id="A566EBABFFDFFF91FD15F9EEEA27FB11" author="Paulson" box="[705,857,1554,1576]" pageId="22" refString="Paulson, D. R. (1977) Odonata. In: Hurlbert, S. H. (Ed.), Biota Acuatica de Sudamerica Austral. San Diego state University, San Diego, pp. 170 - 184." type="book chapter" year="1977">Paulson (1977</bibRefCitation>
: 175
</treatmentCitation>
, mention to
<collectingCountry id="B9E0D6CAFFDFFF91FBDCF9EEED12FB11" box="[1032,1132,1554,1576]" name="Argentina" pageId="22">Argentina</collectingCountry>
and
<collectingCountry id="B9E0D6CAFFDFFF91FB4AF9EEEDA0FB11" box="[1182,1246,1554,1576]" name="Brazil" pageId="22">Brazil</collectingCountry>
, possibly based on
<bibRefCitation id="A566EBABFFDFFF91FF13F9CDE849FB7E" author="Belle" box="[199,311,1585,1607]" pageId="22" refString="Belle, J. (1970) Studies on South American Gomphidae (Odonata) with special reference to the species from Surinam. Studies on the fauna of Suriname and other Guyanas, 11 (43), 1 - 158, pls. XIX." type="journal article" year="1970">Belle 1970</bibRefCitation>
);—
<bibRefCitation id="A566EBABFFDFFF91FE8AF9CDED12FB5F" author="Belle" pageId="22" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">
Belle (1980: 153–155, in part, specimens from Misiones Province,
<collectingCountry id="B9E0D6CAFFDFFF91FBC7F9CDED05FB7E" box="[1043,1147,1585,1607]" name="Argentina" pageId="22">Argentina</collectingCountry>
, and Nova Teutônia village, Santa Catarina State,
<collectingCountry id="B9E0D6CAFFDFFF91FE4BF9ACE8A1FB5F" box="[415,479,1616,1638]" name="Brazil" pageId="22">Brazil</collectingCountry>
, comparison with ♂
<typeStatus id="1E4C28F8FFDFFF91FD64F9ACEA77FB5F" box="[688,777,1616,1638]" pageId="22" type="holotype">holotype</typeStatus>
and allotype, and with
<taxonomicName id="06F7EDD9FFDFFF91FC20F9ADED1BFB5F" ID-CoL="32NMT" box="[1012,1125,1616,1638]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDFFF91FC20F9ADED1BFB5F" box="[1012,1125,1616,1638]" italics="true" pageId="22">C. waltheri</emphasis>
</taxonomicName>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDFFF91FB5EF9ACEA21FBBD" author="Jurzitza" pageId="22" refString="Jurzitza, G. (1981) Lista provicional de los odonatos del Parque Nacional Iguazu, Provincia de Misiones, Republica Argentina. Notulae Odonatologicae, 1 (7), 117 - 118." type="journal article" year="1981">
Jurzitza (1981: 117, record from Parque Nacional Iguazú, Misiones Province,
<collectingCountry id="B9E0D6CAFFDFFF91FD24F992EA26FBBD" box="[752,856,1646,1668]" name="Argentina" pageId="22">Argentina</collectingCountry>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDFFF91FCA9F992EC71FBBD" author="Davies" box="[893,1295,1646,1668]" pageId="22" refString="Davies, D. A. L. &amp; Tobin, P. (1985) The dragonflies of the world: a systematic list of the extant species of Odonata. Vol. 2. Anisoptera. Societas Internationalis Odonatologica, Rapid Communications, 5 (Supplement), 1 - 151." type="journal article" year="1985">Davies &amp; Tobin (1985: 26, mention)</bibRefCitation>
;—
<treatmentCitation id="4056B04BFFDFFF91FAF9F992E8E5FB9A" author="Watson" page="501" pageId="22" year="1985">
<bibRefCitation id="A566EBABFFDFFF91FAF9F992E814FB9A" author="Watson" pageId="22" refString="Watson, J. A. L. &amp; O'Farrell, A. F. (1985) The affinities of the Australian Gomphidae (Odonata). Australian Journal Of Zoology, 33 (4), 499 - 508. http: // dx. doi. org / 10.1071 / ZO 9850499" type="journal article" year="1985">Watson &amp; O’Farrell (1985</bibRefCitation>
: 501
</treatmentCitation>
, mention);—
<bibRefCitation id="A566EBABFFDFFF91FDF4F971ED4CFB9A" author="Carle" box="[544,1074,1677,1699]" pageId="22" refString="Carle, F. L. (1986) The classification, phylogeny and biogeography of the Gomphidae (Anisoptera). I. Classification. Odonatologica, 15 (3), 275 - 326." type="journal article" year="1986">
Carle (1986: 307–308, comparison with
<taxonomicName id="06F7EDD9FFDFFF91FC6EF972ED55FB9A" ID-CoL="32NMT" box="[954,1067,1677,1699]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDFFF91FC6EF972ED55FB9A" box="[954,1067,1677,1699]" italics="true" pageId="22">C. waltheri</emphasis>
</taxonomicName>
)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDFFF91FB85F971EC03FB9A" author="Garrison" box="[1105,1405,1677,1699]" pageId="22" refString="Garrison, R. W. (1991) A synonymic list of the New World Odonata. Argia, 3 (2), 1 - 30." type="journal article" year="1991">Garrison (1991: 17, mention)</bibRefCitation>
;—
<treatmentCitation id="4056B04BFFDFFF91FF13F950E888FBFB" author="Rodrigues" box="[199,502,1708,1730]" page="58" pageId="22" year="1992">
<bibRefCitation id="A566EBABFFDFFF91FF13F950E8AFFBFB" author="Rodrigues" box="[199,465,1708,1730]" pageId="22" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">
Rodrigues
<bibRefCitation id="A566EBABFFDFFF91FEE1F950E8AFFBFB" author="Capitulo" box="[309,465,1708,1730]" pageId="22" refString="Rodrigues Capitulo, A. (1992) Los Odonata de la Republica Argentina (Insecta). Fauna de agua dulce de la Republica Argentina, 34 (1), 1 - 91." type="journal article" year="1992">Capítulo (1992</bibRefCitation>
</bibRefCitation>
: 58
</treatmentCitation>
, in part, record from Misiones Province,
<collectingCountry id="B9E0D6CAFFDFFF91FC43F950EA80FBFB" box="[919,1022,1708,1730]" name="Argentina" pageId="22">Argentina</collectingCountry>
);—
<bibRefCitation id="A566EBABFFDFFF91FBF7F950EC0FFBFB" author="Steinmann" box="[1059,1393,1708,1730]" pageId="22" refString="Steinmann, H. (1997) World catalogue of Odonata, Volume II Anisoptera. Walter de Gruyter, Berlin, 636 pp." type="book" year="1997">Steinmann (1997: 104, mention)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C148965AFFDFFF91FF43F936ED35FAEF" blockId="22.[151,1436,1370,2006]" pageId="22">
<taxonomicName id="06F7EDD9FFDFFF91FF43F936EB69FBD9" ID-CoL="7BYMN" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[151,535,1738,1760]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FF43F936E800FBD9" box="[151,382,1738,1760]" italics="true" pageId="22">Tibiagomphus uncatus</emphasis>
(
<bibRefCitation id="A566EBABFFDFFF91FE5AF936EB71FBD9" author="Fraser" box="[398,527,1738,1760]" pageId="22" refString="Fraser, F. C. (1947) The Odonata of the Argentine Republic I. Acta Zoologica Lilloana del Instituto Miguel Lillo, 4, 427 - 462." type="journal article" year="1947">Fraser, 1947</bibRefCitation>
)
</taxonomicName>
:—
<bibRefCitation id="A566EBABFFDFFF91FDE2F936EBBEFBD9" author="Belle" box="[566,704,1738,1760]" pageId="22" refString="Belle, J. (1992 a) Studies on ultimate instar larvae of Neotropical Gomphidae, with the description of Tibiagomphus gen. nov. (Anisoptera). Odonatologica, 21 (1), 1 - 24." type="journal article" year="1992" yearSuffix="a">Belle (1992a</bibRefCitation>
: 1–3, comb. nov., fixation of
<taxonomicName id="06F7EDD9FFDFFF91FC2DF936EC04FBD9" ID-CoL="9WYQS" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[1017,1402,1738,1760]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFDFFF91FC2DF936ED91FBD9" box="[1017,1263,1738,1760]" italics="true" pageId="22">Cyanogomphus uncatus</emphasis>
<bibRefCitation id="A566EBABFFDFFF91FB23F937EC04FBD9" author="Fraser" box="[1271,1402,1738,1760]" pageId="22" refString="Fraser, F. C. (1947) The Odonata of the Argentine Republic I. Acta Zoologica Lilloana del Instituto Miguel Lillo, 4, 427 - 462." type="journal article" year="1947">Fraser, 1947</bibRefCitation>
</taxonomicName>
, as
<typeStatus id="1E4C28F8FFDFFF91FF13F916E98DFBC6" box="[199,243,1770,1791]" pageId="22">type</typeStatus>
species of
<taxonomicName id="06F7EDD9FFDFFF91FEB3F915E888FBC6" ID-CoL="7WSM" box="[359,502,1769,1791]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDFFF91FEB3F915E888FBC6" box="[359,502,1769,1791]" italics="true" pageId="22">Tibiagomphus</emphasis>
</taxonomicName>
by original designation);—
<bibRefCitation id="A566EBABFFDFFF91FCC6F915EC36FBC6" author="Bridges" box="[786,1352,1769,1791]" pageId="22" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Odonata of the world (Third Edition). Privately published, Urbana, Illinois, 905 pp." type="book" year="1994">
Bridges (1994:
<date id="B549B09AFFDFFF91FC66F915EA93FBC6" box="[946,1005,1769,1791]" pageId="22" value="1951-03">III.51</date>
, VI.8, VII.241,
<date id="B549B09AFFDFFF91FB45F915EDA1FBC6" box="[1169,1247,1769,1791]" pageId="22" value="1970-08">VIII.70</date>
, mention)
</bibRefCitation>
;—
<bibRefCitation id="A566EBABFFDFFF91FAB2F915E87BFA27" author="Belle" pageId="22" refString="Belle, J. (1995) On the female sex of some elusive South-American Gomphidae with the descriptions of three new genera and four new species (Odonata). Zoologische Mededelingen, 69 (2), 19 - 36." type="journal article" year="1995">Belle (1995</bibRefCitation>
: 35–36, figs 15, 23, illustrations of ♀ dorsal detail of occipital plate, postocciput, postoccipital suture and occipital foramen in posterior view, occipital plate in dorsal view, comparison with
<taxonomicName id="06F7EDD9FFDFFF91FC60F8DAEC40FA05" ID-CoL="N7KR" authority="Belle, 1995" authorityName="Belle" authorityYear="1995" box="[948,1342,1830,1853]" class="Insecta" family="Gomphidae" genus="Brasiliogomphus" kingdom="Animalia" order="Odonata" pageId="22" phylum="Arthropoda" rank="species" species="uniseries">
<emphasis id="F3834A48FFDFFF91FC60F8DAEDBDFA04" box="[948,1219,1830,1853]" italics="true" pageId="22">Brasiliogomphus uniseries</emphasis>
<bibRefCitation id="A566EBABFFDFFF91FB1DF8DAEC40FA05" author="Belle" box="[1225,1342,1830,1852]" pageId="22" refString="Belle, J. (1995) On the female sex of some elusive South-American Gomphidae with the descriptions of three new genera and four new species (Odonata). Zoologische Mededelingen, 69 (2), 19 - 36." type="journal article" year="1995">Belle, 1995</bibRefCitation>
</taxonomicName>
);—
<bibRefCitation id="A566EBABFFDFFF91FAB0F8DAE87BFA62" author="Belle" pageId="22" refString="Belle, J. (1996) Higher classification of the South-American Gomphidae (Odonata). Zoologische Mededelingen, 70, 297 - 324." type="journal article" year="1996">Belle (1996</bibRefCitation>
: 316, in part, mention of ♀ cited in
<bibRefCitation id="A566EBABFFDFFF91FDB9F8B9EBA5FA62" author="Belle" box="[621,731,1861,1883]" pageId="22" refString="Belle, J. (1970) Studies on South American Gomphidae (Odonata) with special reference to the species from Surinam. Studies on the fauna of Suriname and other Guyanas, 11 (43), 1 - 158, pls. XIX." type="journal article" year="1970">Belle 1970</bibRefCitation>
);—
<bibRefCitation id="A566EBABFFDFFF91FCD5F8B9EAA5FA62" author="Garrison" box="[769,987,1861,1883]" pageId="22" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
Garrison
<emphasis id="F3834A48FFDFFF91FCB7F8BAEAE8FA62" box="[867,918,1861,1883]" italics="true" pageId="22">et al.</emphasis>
(2006
</bibRefCitation>
: 71, 129–130, 328, 337, figs 390, 728, 731a, b, in part, illustrations of wings and caudal appendages in lateral view from ♂ from Misiones Province,
<collectingCountry id="B9E0D6CAFFDFFF91FADAF898EC08FA43" box="[1294,1398,1892,1914]" name="Argentina" pageId="22">Argentina</collectingCountry>
);—
<bibRefCitation id="A566EBABFFDFFF91FF13F87EE811FAA1" author="Heckman" box="[199,367,1922,1944]" pageId="22" refString="Heckman, C. W. (2006) Encyclopedia of South American aquatic insects: Odonata Anisoptera. Illustrated keys to known families, genera, and species in South America. Springer, Dordrecht, viii + 725 pp." type="book" year="2006">Heckman (2006</bibRefCitation>
: 649–650, fig. 3.2.778, in part, key, reproduction of illustration of ♀ subgenital plate from
<bibRefCitation id="A566EBABFFDFFF91FAFFF87EECE2FAA1" author="Belle" box="[1323,1436,1922,1944]" pageId="22" refString="Belle, J. (1972) Further studies on South American Gomphidae (Odonata). Tijdschrift voor Entomologie, 115 (5), 217 - 240." type="journal article" year="1972">Belle 1972</bibRefCitation>
from Santa Catarina State,
<collectingCountry id="B9E0D6CAFFDFFF91FE0CF85DEB66FA8E" box="[472,536,1953,1975]" name="Brazil" pageId="22">Brazil</collectingCountry>
, reproduction of illustrations of ♂
<typeStatus id="1E4C28F8FFDFFF91FCADF85DEAACFA8E" box="[889,978,1953,1975]" pageId="22" type="holotype">holotype</typeStatus>
postfrons from Rodrigues Capítulo 1985);— von
<bibRefCitation id="A566EBABFFDFFF91FF27F83CED39FAEF" author="Ellenrieder" box="[243,1095,1984,2006]" pageId="22" refString="von Ellenrieder, N. &amp; Muzon, J. (2008) An updated checklist of the Odonata from Argentina. Odonatologica, 37 (1), 55 - 68." type="journal article" year="2008">
Ellenrieder &amp; Muzón (2008: 61, in part, record from Misiones Province,
<collectingCountry id="B9E0D6CAFFDFFF91FC03F83CED3FFAEF" box="[983,1089,1984,2006]" name="Argentina" pageId="22">Argentina</collectingCountry>
)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="9588C6D2FFDEFF90FF43FB77EABCF9C4" httpUri="https://zenodo.org/record/270864/files/figure.png" pageId="23" targetBox="[151,1436,193,1140]" targetPageId="23">
<paragraph id="C148965AFFDEFF90FF43FB77EABCF9C4" blockId="23.[151,1436,1163,1277]" pageId="23">
<emphasis id="F3834A48FFDEFF90FF43FB77E867F999" bold="true" box="[151,281,1163,1185]" pageId="23">FIGURE 11</emphasis>
. Female habitus of Cyanogomphini in lateral view. A–B.
<taxonomicName id="06F7EDD9FFDEFF90FCBAFB77EDC2F998" box="[878,1212,1163,1185]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="23" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
<emphasis id="F3834A48FFDEFF90FCBAFB77EDC2F998" box="[878,1212,1163,1185]" italics="true" pageId="23">Cyanogomphus angelomachadoi</emphasis>
</taxonomicName>
<emphasis id="F3834A48FFDEFF90FB17FB77EC6BF999" bold="true" box="[1219,1301,1163,1184]" pageId="23">
<taxonomicNameLabel id="E8B0F733FFDEFF90FB17FB77EC6BF999" box="[1219,1301,1163,1184]" pageId="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Paratype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448), B. Additional specimen (Brazil. MG: Serra do Salitre, ABMM); C.
<taxonomicName id="06F7EDD9FFDEFF90FF63FB34EB4BF9E7" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" box="[183,565,1224,1246]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="23" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="F3834A48FFDEFF90FF63FB34E8CCF9E7" box="[183,434,1224,1246]" italics="true" pageId="23">Cyanogomphus waltheri</emphasis>
Selys, 1873
</taxonomicName>
(Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); D.
<taxonomicName id="06F7EDD9FFDEFF90FB1FFB34E8D2F9C4" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="23" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDEFF90FB1FFB34ECE5F9E7" box="[1227,1435,1224,1246]" italics="true" pageId="23">Tibiagomphus noval</emphasis>
(Rodrigues Capítulo, 1985)
</taxonomicName>
(Brazil. RS: Jaguarão, MNRJ). Scale bars = 10 mm.
</paragraph>
</caption>
<subSubSection id="89EDC5D1FFDEFF90FF43FAD8EBF9FBB8" pageId="23" type="materials_examined">
<paragraph id="C148965AFFDEFF90FF43FAD8EB83FB00" blockId="23.[151,1437,1316,1989]" pageId="23">
<emphasis id="F3834A48FFDEFF90FF43FAD8E890F804" bold="true" box="[151,494,1316,1341]" pageId="23">
Material examined (
<specimenCount id="D7F15DD3FFDEFF90FE71FAD8E8A0F804" box="[421,478,1316,1341]" pageId="23" type="generic">7 ♂</specimenCount>
).
</emphasis>
<collectingCountry id="B9E0D6CAFFDEFF90FDD7FAD9EB12F804" box="[515,620,1317,1341]" name="Brazil" pageId="23">BRAZIL</collectingCountry>
. Paraná State:
<specimenCount id="D7F15DD3FFDEFF90FCEEFAD9EA20F804" box="[826,862,1317,1341]" pageId="23" type="generic">1♂</specimenCount>
, União da Vitória municipality [
<geoCoordinate id="A4C3F09DFFDEFF90FADBFAD8ECE9F804" box="[1295,1431,1316,1341]" direction="south" orientation="latitude" pageId="23" precision="15" value="-26.230001">26°13’48”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFDEFF90FF43FAB4E837F858" box="[151,329,1352,1377]" direction="west" orientation="longitude" pageId="23" precision="1" value="-51.086">51°05’09.60”W</geoCoordinate>
,
<quantity id="060F3BBFFFDEFF90FE88FAB4E8D5F859" box="[348,427,1352,1377]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="23" unit="m" value="761.0">761 m</quantity>
a.s.l.],
<date id="B549B09AFFDEFF90FDD8FAB4EB28F858" box="[524,598,1352,1377]" pageId="23" value="1952-01">I.1952</date>
, V. Stawiarski leg. (
<collectionCode id="A7E60E9FFFDEFF90FC87FAB5EAE1F858" box="[851,927,1353,1377]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="23">MNRJ</collectionCode>
22064); Santa Catarina State:
<specimenCount id="D7F15DD3FFDEFF90FAC8FAB5EC3EF858" box="[1308,1344,1353,1377]" pageId="23" type="generic">1♂</specimenCount>
, [Seara municipality], Nova Teutônia [village,
<geoCoordinate id="A4C3F09DFFDEFF90FD87FA91EB80F8BC" box="[595,766,1389,1413]" direction="south" orientation="latitude" pageId="23" precision="1" value="-27.160555">27°09’38.00”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFDEFF90FCDEFA90EAE4F8BC" box="[778,922,1388,1413]" direction="west" orientation="longitude" pageId="23" precision="15" value="-52.41611">52°24’58”W</geoCoordinate>
,
<quantity id="060F3BBFFFDEFF90FC73FA90EA8EF8BD" box="[935,1008,1388,1413]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="23" unit="m" value="355.0">355 m</quantity>
a.s.l.],
<date id="B549B09AFFDEFF90FB92FA91EDE4F8BC" box="[1094,1178,1389,1413]" pageId="23" value="1948-02">II.1948</date>
, B. Pohl leg. (
<collectionCode id="A7E60E9FFFDEFF90FA95FA91ECEEF8BC" box="[1345,1424,1389,1413]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23">MZSP</collectionCode>
);
<specimenCount id="D7F15DD3FFDEFF90FF43FA6DE9C5F890" box="[151,187,1425,1449]" pageId="23" type="generic">1♂</specimenCount>
, same data but
<date id="B549B09AFFDEFF90FEB9FA6DE891F890" box="[365,495,1425,1449]" pageId="23" value="1948-11-18">18.XI.1948</date>
, [Richard von Diringshofen leg.] (
<collectionCode id="A7E60E9FFFDEFF90FCA3FA6DEAB8F890" box="[887,966,1425,1449]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23">MZSP</collectionCode>
);
<specimenCount id="D7F15DD3FFDEFF90FC0EFA6DEA81F890" box="[986,1023,1425,1449]" pageId="23" type="generic">1♂</specimenCount>
, same data but
<date id="B549B09AFFDEFF90FB65FA6DEC4FF890" box="[1201,1329,1425,1449]" pageId="23" value="1948-11-19">19.XI.1948</date>
(
<collectionCode id="A7E60E9FFFDEFF90FA95FA6DECEEF890" box="[1345,1424,1425,1449]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23">MZSP</collectionCode>
);
<specimenCount id="D7F15DD3FFDEFF90FF43FA49E9C2F8F4" box="[151,188,1461,1485]" pageId="23" type="generic">2♂</specimenCount>
, same data but
<date id="B549B09AFFDEFF90FEB8FA49E892F8F4" box="[364,492,1461,1485]" pageId="23" value="1948-11-24">24.XI.1948</date>
(
<collectionCode id="A7E60E9FFFDEFF90FE28FA49EB35F8F4" box="[508,587,1461,1485]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23">MZSP</collectionCode>
);
<specimenCount id="D7F15DD3FFDEFF90FD8BFA49EBFDF8F4" box="[607,643,1461,1485]" pageId="23" type="generic">1♂</specimenCount>
, same data but
<date id="B549B09AFFDEFF90FCE1FA49EACBF8F4" box="[821,949,1461,1485]" pageId="23" value="1948-11-28">28.XI.1948</date>
(
<collectionCode id="A7E60E9FFFDEFF90FC10FA49ED6DF8F4" box="[964,1043,1461,1485]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23">MZSP</collectionCode>
).
<collectingCountry id="B9E0D6CAFFDEFF90FBF3FA49EDC0F8F4" box="[1063,1214,1461,1485]" name="Paraguay" pageId="23">PARAGUAY</collectingCountry>
. Itapúa department:
<specimenCount id="D7F15DD3FFDEFF90FF43FA25E9C5F8C8" box="[151,187,1497,1521]" pageId="23" type="generic">1♂</specimenCount>
, San Benito [Agricultural School,
<geoCoordinate id="A4C3F09DFFDEFF90FD98FA24EB8BF8C8" box="[588,757,1496,1521]" direction="south" orientation="latitude" pageId="23" precision="1" value="-26.831457">26°49’53.24”S</geoCoordinate>
,
<geoCoordinate id="A4C3F09DFFDEFF90FCD7FA24EACBF8C8" box="[771,949,1496,1521]" direction="west" orientation="longitude" pageId="23" precision="1" value="-55.725758">55°43’32.73”W</geoCoordinate>
,
<quantity id="060F3BBFFFDEFF90FC10FA25ED73F8C9" box="[964,1037,1497,1521]" metricMagnitude="2" metricUnit="m" metricValue="2.02" pageId="23" unit="m" value="202.0">202 m</quantity>
a.s.l.] / Pastoreo,
<date id="B549B09AFFDEFF90FB0BFA25EC33F8C8" box="[1247,1357,1497,1521]" pageId="23" value="1980-01-21">21.I.1980</date>
, Linda Strickman leg. (D. R. Paulson collection, Seattle, WA,
<collectingCountry id="B9E0D6CAFFDEFF90FCDAFA01EA37FB2C" box="[782,841,1533,1557]" name="United States of America" pageId="23">USA</collectingCountry>
, not examined). Specimens at
<collectionCode id="A7E60E9FFFDEFF90FB7DFA01ED8CFB2C" box="[1193,1266,1533,1557]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23">MZSP</collectionCode>
all ex-Richard von Diringshofen Collection under species number 83.
</paragraph>
<paragraph id="C148965AFFDEFF90FF13F9B8EBF9FBB8" blockId="23.[151,1437,1316,1989]" pageId="23">
<emphasis id="F3834A48FFDEFF90FF13F9B8E8F2FB64" bold="true" box="[199,396,1604,1629]" pageId="23">
<typeStatus id="1E4C28F8FFDEFF90FF13F9B8E87CFB64" box="[199,258,1604,1629]" pageId="23">Type</typeStatus>
repository.
</emphasis>
<typeStatus id="1E4C28F8FFDEFF90FE43F9B8E881FB64" box="[407,511,1604,1629]" pageId="23" type="holotype">Holotype</typeStatus>
♂ and allotype ♀ by original designation in Instituto Fundación Miguel Lillo, Tucumán,
<collectingCountry id="B9E0D6CAFFDEFF90FED9F994E803FBB8" box="[269,381,1640,1665]" name="Argentina" pageId="23">Argentina</collectingCountry>
(
<collectionCode id="A7E60E9FFFDEFF90FE58F995E8AAFBB8" box="[396,468,1641,1665]" pageId="23">IFML</collectionCode>
, not examined).
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDEFF90FF13F970E8EAFA60" pageId="23" type="description">
<paragraph id="C148965AFFDEFF90FF13F970EA60FA0C" blockId="23.[151,1437,1316,1989]" pageId="23">
<emphasis id="F3834A48FFDEFF90FF13F970E803FB9C" bold="true" box="[199,381,1676,1701]" pageId="23">Measurements.</emphasis>
Males (n = 7). Total length (incl. caudal appendages) 45.1–49.7; abdomen length (excluding caudal appendages) 32.3–36.4; head maximum width 6.2–6.5; Fw length 28.0–30.6; Hw length 26.8–29.7; Fw maximum width 5.7–6.8, in Hw 6.8–7.7; pt length
<quantity id="060F3BBFFFDEFF90FD33F928EA25FBD4" box="[743,859,1748,1773]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" metricValueMax="8.89" metricValueMin="7.366" pageId="23" unit="in" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 in</quantity>
Fw,
<quantity id="060F3BBFFFDEFF90FC43F929ED75FBD4" box="[919,1035,1748,1773]" metricMagnitude="-2" metricUnit="m" metricValue="9.017" metricValueMax="9.906" metricValueMin="8.128" pageId="23" unit="in" value="3.55" valueMax="3.9" valueMin="3.2">3.2–3.9 in</quantity>
Hw; length of metathoracic femur 7.3–8.7; metathoracic tibia 5.9–7.0; length of S9+
<quantity id="060F3BBFFFDEFF90FD19F905EA76FA28" box="[717,776,1784,1809]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="23" unit="in" value="10.0">10 in</quantity>
lateral view 3.4–4.0; total length of cercus in lateral view 1.8–2.1. Ratio cercus length / maximum width 0.25–0.30.
</paragraph>
<paragraph id="C148965AFFDEFF90FF13F8BCE8EAFA60" blockId="23.[151,1437,1316,1989]" box="[199,404,1856,1881]" pageId="23">
<emphasis id="F3834A48FFDEFF90FF13F8BCE869FA60" bold="true" box="[199,279,1856,1881]" pageId="23">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDEFF90FF13F898EC47FA45" box="[199,1337,1892,1917]" pageId="23" type="diagnosis">
<paragraph id="C148965AFFDEFF90FF13F898EC47FA45" blockId="23.[151,1437,1316,1989]" box="[199,1337,1892,1917]" pageId="23">
<emphasis id="F3834A48FFDEFF90FF13F898E844FA44" bold="true" box="[199,314,1892,1917]" pageId="23">Diagnosis</emphasis>
. This dark species is diagnosed from its congener and from
<taxonomicName id="06F7EDD9FFDEFF90FC09F898EDF5FA44" box="[989,1163,1892,1917]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="23" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFDEFF90FC09F898EDF5FA44" box="[989,1163,1892,1917]" italics="true" pageId="23">Cyanogomphus</emphasis>
</taxonomicName>
under
<taxonomicName id="06F7EDD9FFDEFF90FB0DF899EC4CFA44" box="[1241,1330,1892,1917]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="23" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFDEFF90FB0DF899EC4CFA44" box="[1241,1330,1892,1917]" italics="true" pageId="23">T. noval</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFDEFF9FFF13F874EBAAF897" lastPageId="24" pageId="23" type="distribution">
<paragraph id="C148965AFFDEFF90FF13F874EC52FAFC" blockId="23.[151,1437,1316,1989]" pageId="23">
<emphasis id="F3834A48FFDEFF90FF13F874E827FA98" bold="true" box="[199,345,1928,1953]" pageId="23">Distribution</emphasis>
. Misiones Province in
<collectingCountry id="B9E0D6CAFFDEFF90FDB0F874EBA8FA98" box="[612,726,1928,1953]" name="Argentina" pageId="23">Argentina</collectingCountry>
,
<collectingCountry id="B9E0D6CAFFDEFF90FD30F875EA32FA98" box="[740,844,1929,1953]" name="Paraguay" pageId="23">Paraguay</collectingCountry>
in Itapúa department, and Paraná to Santa Catarina States in
<collectingCountry id="B9E0D6CAFFDEFF90FF29F850E83AFAFC" box="[253,324,1964,1989]" name="Brazil" pageId="23">Brazil</collectingCountry>
, in Parana dominion in Araucaria and Parana Forest biogeographical provinces (
<figureCitation id="59CC8ADFFFDEFF90FB1FF850EC60FAFC" box="[1227,1310,1964,1989]" captionStart="FIGURE 13" captionStartId="25.[151,250,1528,1550]" captionTargetBox="[159,1434,488,1498]" captionTargetId="figure@25.[151,1436,483,1503]" captionTargetPageId="25" captionText="FIGURE 13. Map of southeastern South America with distribution records of Tibiagomphus Belle, 1992, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: ER, Entre Rios and MI, Misiones for Argentine provinces; PR, Paraná, RS, Rio Grande do Sul, and SC, Santa Catarina for Brazilian States." httpUri="https://zenodo.org/record/270866/files/figure.png" pageId="23">Fig. 13</figureCitation>
).
</paragraph>
<caption id="9588C6D2FFD1FF9FFF43FB24EB95F873" httpUri="https://zenodo.org/record/270865/files/figure.png" pageId="24" targetBox="[151,1436,215,1198]" targetPageId="24">
<paragraph id="C148965AFFD1FF9FFF43FB24EB95F873" blockId="24.[151,1436,1240,1354]" pageId="24">
<emphasis id="F3834A48FFD1FF9FFF43FB24E864F9D4" bold="true" box="[151,282,1240,1262]" pageId="24">FIGURE 12</emphasis>
. Map of southeastern South America with distribution records of
<taxonomicName id="06F7EDD9FFD1FF9FFC13FB24ED99F9D7" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" box="[967,1255,1240,1262]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="genus">
<emphasis id="F3834A48FFD1FF9FFC13FB24ED1AF9D7" box="[967,1124,1240,1262]" italics="true" pageId="24">Cyanogomphus</emphasis>
Selys, 1873
</taxonomicName>
, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: MI, Misiones Province, Argentina; BA, Bahia; GO, Goiás; MG, Minas Gerais; MT, Mato Grosso; RJ, Rio de Janeiro; RS, Rio Grande do Sul; SC, Santa Catarina, and SP, São Paulo for Brazilian States.
</paragraph>
</caption>
<paragraph id="C148965AFFD1FF9FFF13FA8DEBAAF897" blockId="24.[151,1437,1393,2030]" pageId="24">
<emphasis id="F3834A48FFD1FF9FFF13FA8DEB46F8B3" bold="true" box="[199,568,1393,1418]" pageId="24">Biological and ecological data.</emphasis>
Based on sites where adults have been collected, this species is associated with forested areas at 188–781 meters of elevation.
</paragraph>
</subSubSection>
<subSubSection id="89EDC5D1FFD1FF9EFF13FA45EC25FC94" lastPageId="25" pageId="24" type="discussion">
<paragraph id="C148965AFFD1FF9EFF13FA45E846FDED" blockId="24.[151,1437,1393,2030]" lastBlockId="25.[151,1436,151,429]" lastPageId="25" pageId="24">
<emphasis id="F3834A48FFD1FF9FFF13FA45E84AF8EB" bold="true" box="[199,308,1465,1490]" pageId="24">Remarks</emphasis>
. Belle’s (1980) paper contributes significantly to the identity of this species, even considering the poor condition of the
<typeStatus id="1E4C28F8FFD1FF9FFE40FA22E8BAF8CF" box="[404,452,1502,1526]" pageId="24">type</typeStatus>
series. Besides the male
<typeStatus id="1E4C28F8FFD1FF9FFD38FA21EA30F8CF" box="[748,846,1501,1526]" pageId="24" type="holotype">holotype</typeStatus>
and allotype, Belle examined three other Pampean males, one from Entre Rios Province,
<collectingCountry id="B9E0D6CAFFD1FF9FFD8FF9FDEBB4FB23" box="[603,714,1537,1562]" name="Argentina" pageId="24">Argentina</collectingCountry>
(
<typeStatus id="1E4C28F8FFD1FF9FFD0AF9FEEA6EFB23" box="[734,784,1538,1562]" pageId="24">type</typeStatus>
locality of
<taxonomicName id="06F7EDD9FFD1FF9FFC48F9FEEA86FB23" box="[924,1016,1537,1562]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFD1FF9FFC48F9FEEA86FB23" box="[924,1016,1537,1562]" italics="true" pageId="24">T. noval</emphasis>
</taxonomicName>
) and two from distinct localities in
<collectingCountry id="B9E0D6CAFFD1FF9FFF43F9DAE984FB07" box="[151,250,1574,1598]" name="Uruguay" pageId="24">Uruguay</collectingCountry>
, as well as two other males from Parana dominion (Atlantic Forest) of Misiones Province in
<collectingCountry id="B9E0D6CAFFD1FF9FFAF9F9D9ECE2FB07" box="[1325,1436,1573,1598]" name="Argentina" pageId="24">Argentina</collectingCountry>
(
<typeStatus id="1E4C28F8FFD1FF9FFF4AF9B6E9AEFB5B" box="[158,208,1610,1634]" pageId="24">type</typeStatus>
locality of
<taxonomicName id="06F7EDD9FFD1FF9FFE84F9B6E8BCFB58" box="[336,450,1610,1633]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD1FF9FFE84F9B6E8BCFB58" box="[336,450,1610,1633]" italics="true" pageId="24">T. uncatus</emphasis>
</taxonomicName>
), and Nova Teutônia village in Santa Catarina State,
<collectingCountry id="B9E0D6CAFFD1FF9FFBC1F9B5ED22FB5B" box="[1045,1116,1609,1634]" name="Brazil" pageId="24">Brazil</collectingCountry>
, making a total of six males. He recognized several differences among the Pampean southern specimens from the typical northern specimens of Argentinean and Brazilian Atlantic Forest (
<bibRefCitation id="A566EBABFFD1FF9FFD56F96DEAF1FB93" author="Belle" box="[642,911,1681,1706]" pageId="24" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980" yearSuffix="p">Belle 1980, p. 154–155</bibRefCitation>
) including the paler color, shape of postfrons, smaller size, and more slender hamule, all of which agree with
<taxonomicName id="06F7EDD9FFD1FF9FFC8FF94AEACBFBF7" box="[859,949,1717,1742]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFD1FF9FFC8FF94AEACBFBF7" box="[859,949,1717,1742]" italics="true" pageId="24">T. noval</emphasis>
</taxonomicName>
as described by Rodrigues Capítulo (1985) from Entre Rios Province. We examined seven males from the northern part of its range, including material from Nova Teutônia and eight males from southern parts of its range, including material as far south as Entre Rios Province. Based on Belle’s (1980) description of the
<typeStatus id="1E4C28F8FFD1FF9FFCD9F8DDEA11FA03" box="[781,879,1825,1850]" pageId="24" type="holotype">holotype</typeStatus>
and comparison with other specimens from its northern range in
<collectingCountry id="B9E0D6CAFFD1FF9FFE89F8B9E8B3FA67" box="[349,461,1861,1886]" name="Argentina" pageId="24">Argentina</collectingCountry>
and
<collectingCountry id="B9E0D6CAFFD1FF9FFDD7F8B9EB34FA67" box="[515,586,1861,1886]" name="Brazil" pageId="24">Brazil</collectingCountry>
, the specific characters cited for
<taxonomicName id="06F7EDD9FFD1FF9FFC6CF8BBED54FA67" box="[952,1066,1863,1886]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD1FF9FFC6CF8BBED54FA67" box="[952,1066,1863,1886]" italics="true" pageId="24">T. uncatus</emphasis>
</taxonomicName>
are consistent. Thus we consider the darker specimens from Parana dominion to be
<taxonomicName id="06F7EDD9FFD1FF9FFD1CF896EA46FAB8" box="[712,824,1898,1921]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD1FF9FFD1CF896EA46FAB8" box="[712,824,1898,1921]" italics="true" pageId="24">T. uncatus</emphasis>
</taxonomicName>
and the paler specimens from Pampean province to be
<taxonomicName id="06F7EDD9FFD1FF9FFF43F873E991FA9F" box="[151,239,1933,1958]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFD1FF9FFF43F873E991FA9F" box="[151,239,1933,1958]" italics="true" pageId="24">T. noval</emphasis>
</taxonomicName>
(
<figureCitation id="59CC8ADFFFD1FF9FFED4F871E826FA9F" box="[256,344,1933,1958]" captionStart="FIGURE 13" captionStartId="25.[151,250,1528,1550]" captionTargetBox="[159,1434,488,1498]" captionTargetId="figure@25.[151,1436,483,1503]" captionTargetPageId="25" captionText="FIGURE 13. Map of southeastern South America with distribution records of Tibiagomphus Belle, 1992, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: ER, Entre Rios and MI, Misiones for Argentine provinces; PR, Paraná, RS, Rio Grande do Sul, and SC, Santa Catarina for Brazilian States." httpUri="https://zenodo.org/record/270866/files/figure.png" pageId="24">Figs 13</figureCitation>
). Most males of
<taxonomicName id="06F7EDD9FFD1FF9FFDC3F873EBF6FA9F" box="[535,648,1935,1958]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD1FF9FFDC3F873EBF6FA9F" box="[535,648,1935,1958]" italics="true" pageId="24">T. uncatus</emphasis>
</taxonomicName>
lack a pale spot or stripe over the dark brown mesepisternum such as are shown by the
<typeStatus id="1E4C28F8FFD1FF9FFE8EF84DE8C2FAF3" box="[346,444,1969,1994]" pageId="24" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="A566EBABFFD1FF9FFE1EF84DEBE2FAF3" author="Belle" box="[458,668,1969,1994]" pageId="24" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980" yearSuffix="p">Belle 1980, p. 153</bibRefCitation>
). A few
<taxonomicName id="06F7EDD9FFD1FF9FFD2DF84EEA15FAF0" box="[761,875,1970,1993]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD1FF9FFD2DF84EEA15FAF0" box="[761,875,1970,1993]" italics="true" pageId="24">T. uncatus</emphasis>
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have a small elongated pale spot corresponding to the first antehumeral stripe, while males of
<taxonomicName id="06F7EDD9FFD1FF9FFD58F82BEB96FAD7" box="[652,744,2005,2030]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="24" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFD1FF9FFD58F82BEB96FAD7" box="[652,744,2005,2030]" italics="true" pageId="24">T. noval</emphasis>
</taxonomicName>
always have pale spots on the mesepisternum similar to the pattern illustrated for females of
<taxonomicName id="06F7EDD9FFD0FF9EFDDEFF65EB03FD89" box="[522,637,153,176]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD0FF9EFDDEFF65EB03FD89" box="[522,637,153,176]" italics="true" pageId="25">T. uncatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="A566EBABFFD0FF9EFD59FF6BEA1DFD89" author="Belle" box="[653,867,151,176]" pageId="25" refString="Belle, J. (1970) Studies on South American Gomphidae (Odonata) with special reference to the species from Surinam. Studies on the fauna of Suriname and other Guyanas, 11 (43), 1 - 158, pls. XIX." type="journal article" year="1970">Belle 1970, fig. 26</bibRefCitation>
) and
<taxonomicName id="06F7EDD9FFD0FF9EFC77FF65EA85FD89" box="[931,1019,151,176]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFD0FF9EFC77FF65EA85FD89" box="[931,1019,151,176]" italics="true" pageId="25">T. noval</emphasis>
</taxonomicName>
(
<bibRefCitation id="A566EBABFFD0FF9EFBD8FF6BED8DFD89" author="Garrison" box="[1036,1267,151,176]" pageId="25" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2006) Dragonfly genera of the New World, an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, xiv + 368 pp." type="book" year="2006">
Garrison
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. 2006
</bibRefCitation>
, fig. 727, cited as
<taxonomicName id="06F7EDD9FFD0FF9EFF61FF41E856FDED" box="[181,296,189,212]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD0FF9EFF61FF41E856FDED" box="[181,296,189,212]" italics="true" pageId="25">T. uncatus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="C148965AFFD0FF9EFF13FF23EDD5FC79" blockId="25.[151,1436,151,429]" pageId="25">
<bibRefCitation id="A566EBABFFD0FF9EFF13FF23E8D7FDC1" author="Belle" box="[199,425,223,248]" pageId="25" refString="Belle, J. (1980) Notes on three species of Cyanogomphus Selys, 1873 (Odonata: Gomphidae). Entomologische Berichten, 40 (10), 151 - 155." type="journal article" year="1980">Belle (1980, fig. 13)</bibRefCitation>
also illustrated an unusual posterior hamule of
<taxonomicName id="06F7EDD9FFD0FF9EFC6CFF1DED71FDC1" box="[952,1039,223,248]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFD0FF9EFC6CFF1DED71FDC1" box="[952,1039,223,248]" italics="true" pageId="25">T. noval</emphasis>
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from
<collectingCountry id="B9E0D6CAFFD0FF9EFB80FF1CEDC8FDC1" box="[1108,1206,224,248]" name="Uruguay" pageId="25">Uruguay</collectingCountry>
(cited as
<taxonomicName id="06F7EDD9FFD0FF9EFAF5FF1DECEFFDC1" box="[1313,1425,225,248]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD0FF9EFAF5FF1DECEFFDC1" box="[1313,1425,225,248]" italics="true" pageId="25">T. uncatus</emphasis>
</taxonomicName>
) with additional spines along the anterior margin, similar to one of the male
<taxonomicName id="06F7EDD9FFD0FF9EFC09FEF9ED33FC25" box="[989,1101,261,284]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD0FF9EFC09FEF9ED33FC25" box="[989,1101,261,284]" italics="true" pageId="25">T. uncatus</emphasis>
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we examined that has several additional differently sized spines, including minute ones, along the internal margin (
<figureCitation id="59CC8ADFFFD0FF9EFB9BFEDBEDECFC79" box="[1103,1170,295,320]" captionStart="FIGURE 5" captionStartId="11.[151,250,1535,1557]" captionTargetBox="[151,1436,195,1507]" captionTargetId="figure@11.[151,1436,195,1511]" captionTargetPageId="11" captionText="FIGURE 5. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A – D) and posterior hamule in lateral view (E – L). A – B, F – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); C – D, I – L. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). E. Cyanogomphus comparabilis Belle, 1994, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="25">
Fig.
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I).
</paragraph>
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Even though we examined no female of
<taxonomicName id="06F7EDD9FFD0FF9EFD5DFEB1EB85FC5D" box="[649,763,333,356]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD0FF9EFD5DFEB1EB85FC5D" box="[649,763,333,356]" italics="true" pageId="25">T. uncatus</emphasis>
</taxonomicName>
, the homogeneous darker general color diagnoses the sex of this species from
<taxonomicName id="06F7EDD9FFD0FF9EFE88FE8DE8CBFCB1" box="[348,437,367,392]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="noval">
<emphasis id="F3834A48FFD0FF9EFE88FE8DE8CBFCB1" box="[348,437,367,392]" italics="true" pageId="25">T. noval</emphasis>
</taxonomicName>
(a female of
<taxonomicName id="06F7EDD9FFD0FF9EFD99FE8DEBBEFCB1" box="[589,704,369,392]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="25" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="F3834A48FFD0FF9EFD99FE8DEBBEFCB1" box="[589,704,369,392]" italics="true" pageId="25">T. uncatus</emphasis>
</taxonomicName>
from Misiones in RWG collection agrees with the darker pattern described here, RWG pers. comm.) as well as by its more northern allopatric distribution as discussed above.
</paragraph>
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81
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<mods:classification id="F8CE7E9313A8ADA3EDCFB76CA4D5F8ED">journal article</mods:classification>
<mods:identifier id="4E25F9828D7B958339103B2919832019" type="DOI">http://dx.doi.org/10.3897/zookeys.1174.107314</mods:identifier>
<mods:identifier id="2B26E1CA943441625E1E84CEEA196CB5" type="Pensoft-Pub">1313-2970-1174-35</mods:identifier>
<mods:identifier id="BEFC051FC86CDA5DE474D243EF24D7F9" type="ZooBank">91E908240C45471A86BBA24ADC35E743</mods:identifier>
<mods:identifier id="47104CBE07544BB73A3B51D2AAD0C238" type="Pensoft-UUID">635A059AEF505E17B338F6484B5F4DBF</mods:identifier>
</mods:mods>
<treatment id="495EB4508B815496B0C4918B36F0C29C" LSID="urn:lsid:zoobank.org:act:BB6252E9-C6EC-4863-9596-7E1DA073C64F" httpUri="http://treatment.plazi.org/id/495EB4508B815496B0C4918B36F0C29C" lastPageNumber="35" pageId="0" pageNumber="35">
<subSubSection id="89608DC2603B6CD22DAD6B749A3C1080" pageId="0" pageNumber="35" type="nomenclature">
<paragraph id="883D376DE69DD683473D63619723F508" pageId="0" pageNumber="35">
<taxonomicName id="4D7ACCA6759C9CDFA444DBB25979B025" LSID="https://zoobank.org/BB6252E9-C6EC-4863-9596-7E1DA073C64F" authority="Harris &amp; Armitage, 2023" authorityName="Harris &amp; Armitage" authorityYear="2023" class="Insecta" family="Hydroptilidae" genus="Alisotrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alisotrichia eisbergae" order="Trichoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="eisbergae" status="sp. nov.">Alisotrichia eisbergae</taxonomicName>
<taxonomicNameLabel id="080D82232F152A60CD8E34C946884468" pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F22244BF721DDF4B27F239AFC18DCB73" pageId="0" pageNumber="35" type="description">
<paragraph id="A165FCBAFFFEAF6CF79206216F88BEC0" pageId="0" pageNumber="35">
<figureCitation id="CBA567F40A98E4FBD34C24BDF32AAF89" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Alisotrichia eisbergae sp. nov., male holotype, genitalia A left lateral B variation in apex of segment VIIl C dorsal D ventral E antenna, basal segments F maxillary palp G phallus, dorsal H phallus, left lateral." figureDoi="10.3897/zookeys.1174.107314.figure2" httpUri="https://binary.pensoft.net/fig/890127" pageId="0" pageNumber="35">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="98FDA98FA29C34F57DF8F70C0D40BA35" pageId="0" pageNumber="35" type="type locality">
<paragraph id="F40C7D6814DB3CB758778CD9E3E7E6DF" pageId="0" pageNumber="35">Type locality.</paragraph>
<paragraph id="088603C11D592169FC79B1C9F0C878D1" pageId="0" pageNumber="35">
<emphasis id="FB4CF09D6812DE1691E1E2FD79A98167" bold="true" pageId="0" pageNumber="35">Panama: Veraguas Province</emphasis>
: Cuenca 097; Santa Fe District; Santa Fe NP;
<normalizedToken id="3DED9684605B444924795A02D467175D" originalValue="Río">Rio</normalizedToken>
Piedra de Moler; PSPSCB-PNSF-C097-2017-012;
<geoCoordinate id="EFF5D90E9D869C81F88DBD74E73AEC5E" degrees="8.56553" direction="north" orientation="latitude" precision="1" value="8.56553">8.56553°N</geoCoordinate>
,
<geoCoordinate id="1747942A7C4DC961BEC0B80E0CD992DE" degrees="81.18817" direction="west" orientation="longitude" precision="1" value="-81.18817">81.18817°W</geoCoordinate>
; 340 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="0F50780BF0F9B6CD48B6ED8DF491936A" pageId="0" pageNumber="35" type="type material">
<paragraph id="65452F2241EBF0F9BA00DE8B05C1481B" pageId="0" pageNumber="35">Type material.</paragraph>
<paragraph id="C72620573EE7B4C722573F87DBB67025" pageId="0" pageNumber="35">
<emphasis id="576ED26721AAE18B6EDC7B332496125C" bold="true" pageId="0" pageNumber="35">
<emphasis id="E6AE798FC9818D2EE9F926391FC46FBB" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
: male, Panama: Veraguas Province
</emphasis>
: Cuenca 097; Santa Fe District; Santa Fe NP;
<normalizedToken id="47CB5170FC71EC52C18251EC30D6611A" originalValue="Río">Rio</normalizedToken>
Piedra de Moler; PSPSCB-PNSF-C097-2017-012;
<geoCoordinate id="2E535DF5C1E681DE2F91D02FD4DE7617" degrees="8.56553" direction="north" orientation="latitude" precision="1" value="8.56553">8.56553°N</geoCoordinate>
,
<geoCoordinate id="A53C7D55ABF6DE7A1A73E66B557050E5" degrees="81.18817" direction="west" orientation="longitude" precision="1" value="-81.18817">81.18817°W</geoCoordinate>
; 340 m a.s.l.; UV light trap; A. Cornejo, T.
<normalizedToken id="05D40B6A0C4F865B4AA72BCB21DD482A" originalValue="Ríos">Rios</normalizedToken>
, E.
<normalizedToken id="EE5DEE120A0D069C85E7266BA63BF81C" originalValue="Álvarez">Alvarez</normalizedToken>
, C. Nieto, leg.; 20.iv.2017; MIUP-001-T-2023 (in alcohol).
<emphasis id="B5C6C4C438F59474B0EDB9A45566526B" bold="true" italics="true" pageId="0" pageNumber="35">Paratypes</emphasis>
: same data as for holotype; 4 males; MIUP (in alcohol).
</paragraph>
</subSubSection>
<subSubSection id="AFE3030D7DCEF064DAA4007FA4861A07" pageId="0" pageNumber="35" type="materials_examined">
<paragraph id="65D617DB45275E4ECDF14210737868C6" pageId="0" pageNumber="35">Other material examined.</paragraph>
<paragraph id="912006ECC55EAD37695F58D7496372B1" pageId="0" pageNumber="35">
<materialsCitation id="BC33A5D117636DB422D84DD02D5653BD" collectingDate="2017-04-19" collectingDateMax="2017-04-27" collectingDateMin="2017-04-19" collectingMethod="Malaise trap, UV light trap" collectorName="Santa Fe NP, A. Cornejo, T. Rios, E. Alvarez, C. Nieto, Quebrada" country="Panama" county="Cuenca" elevation="623" latitude="8.55343" location="Quebrada Primer Brazo Mulaba" longLatPrecision="1" longitude="-81.17675" municipality="Santa Fe District" specimenCount="27" specimenCount-male="27" stateProvince="Veraguas Province">
<emphasis id="AF7B29F973C33A7A7BE3C3CFED984A8E" bold="true" pageId="0" pageNumber="35">
<collectingCountry id="E3207DE3F57EDB00D972E799A83272FA" name="Panama">Panama</collectingCountry>
:
<collectingRegion id="5F8453E9D52B609BCE06E388CCF773D5" country="Panama" name="Veraguas">Veraguas Province</collectingRegion>
</emphasis>
•
<specimenCount id="DF0A7BA8B6C490397C8027ABAA00E4FC" type="male">1 male</specimenCount>
;
<collectingCounty id="B30FE8E2D703B3C0ABA0311990EDF62F">Cuenca</collectingCounty>
132;
<collectingMunicipality id="4554CE9E26DAD4473A726475318E1478">Santa Fe District</collectingMunicipality>
;
<collectorName id="4C91A6FA153534716EFC359AF388C332">Santa Fe NP</collectorName>
;
<location id="2A8D791BB9D3021258F591026683DC82" LSID="urn:lsid:plazi:treatment:495EB4508B815496B0C4918B36F0C29C:2A8D791BB9D3021258F591026683DC82" country="Panama" county="Cuenca" latitude="8.55343" longLatPrecision="1" longitude="-81.17675" municipality="Santa Fe District" name="Quebrada Primer Brazo Mulaba" stateProvince="Veraguas Province">
Quebrada Primer Brazo
<normalizedToken id="3DCEE36FE5250E13A3C965A619551897" originalValue="Mulabá">Mulaba</normalizedToken>
</location>
; PSPSCB-PNSF-C132-2017-007;
<geoCoordinate id="86A0F36DD38DA1D915D453F2E2CBAAD4" degrees="8.52577" direction="north" orientation="latitude" precision="1" value="8.52577">8.52577°N</geoCoordinate>
,
<geoCoordinate id="D0442EAC29637093C661BBAE12BD0A0F" degrees="81.13045" direction="west" orientation="longitude" precision="1" value="-81.13045">81.13045°W</geoCoordinate>
;
<elevation id="25341172C67BF98C73FB2AAC350FD1B7" metricMagnitude="2" metricUnit="m" metricValue="6.2299999999999995" unit="m" value="623.0">
<quantity id="FA2E2FEB20AC24DC5DBE91566C16CC92" metricMagnitude="2" metricUnit="m" metricValue="6.2299999999999995" unit="m" value="623.0">623 m</quantity>
a.s.l.
</elevation>
;
<collectingMethod id="5C64D2DA9D6D44C152AACD8A3093D76D">Malaise trap</collectingMethod>
;
<collectorName id="072E04C871FCEA2A9B3EC74220EB43DC">A. Cornejo</collectorName>
,
<collectorName id="170804ED5A9C4CDD63CBD4F9E59DC5BF">
T.
<normalizedToken id="A9DE22110DE923AE1F8ED17F098AD89E" originalValue="Ríos">Rios</normalizedToken>
</collectorName>
,
<collectorName id="30CD3128ACB5880C8D7FDE6C96A439F9">
E.
<normalizedToken id="0D0FE7583F8EABF756E6C52645B0EDAF" originalValue="Álvarez">Alvarez</normalizedToken>
</collectorName>
,
<collectorName id="F989A38235153104453E320A3D8DA579">C. Nieto</collectorName>
, leg.;
<collectingDate id="A9B08D4BB593EC20205B174122BDB922" value="2017-04-19" valueMax="2017-04-23" valueMin="2017-04-19">19-23.iv.2017</collectingDate>
; MIUP • ibid.,
<specimenCount id="A7DD5D04791F37E0DA182E01C47E3E1C" type="male">23 males</specimenCount>
;
<location id="B3B2F36EBB0B5101EB82D5846EC8EA6B" LSID="urn:lsid:plazi:treatment:495EB4508B815496B0C4918B36F0C29C:B3B2F36EBB0B5101EB82D5846EC8EA6B" country="Panama" county="Cuenca" latitude="8.55343" longLatPrecision="1" longitude="-81.17675" municipality="Santa Fe District" name="Rio Calovebora" stateProvince="Veraguas Province">
<normalizedToken id="10B756DDCE3850189C24951D84C414EE" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="7F56788BB3D0F1901D4ED9F5943C948D" originalValue="Calovébora">Calovebora</normalizedToken>
</location>
; PSPSCB-NPSF-C-097-2017-005;
<geoCoordinate id="F42A5D7BD13FD1E955CE298E0594DE5A" degrees="8.54318" direction="north" orientation="latitude" precision="1" value="8.54318">8.54318°N</geoCoordinate>
,
<geoCoordinate id="D4057D3499820BD55F9A84BD4733ACC1" degrees="81.16398" direction="west" orientation="longitude" precision="1" value="-81.16398">81.16398°W</geoCoordinate>
;
<elevation id="C800890B81A8C83893727F16DEBAA7C2" metricMagnitude="2" metricUnit="m" metricValue="5.36" unit="m" value="536.0">
<quantity id="0ECFFC31E9869A25C953D67E3B6CEF47" metricMagnitude="2" metricUnit="m" metricValue="5.36" unit="m" value="536.0">536 m</quantity>
a.s.l.
</elevation>
;
<collectingMethod id="09B9F81D54014D31020B3832564CB589">Malaise trap</collectingMethod>
;
<collectorName id="E8EC1886A1C9B170442ECDAB23F43667">
T.
<normalizedToken id="B42258E3738EFBE33982DB7A6A82F420" originalValue="Ríos">Rios</normalizedToken>
</collectorName>
,
<collectorName id="E206BECEF9D4D2BDEB73271CAD9F2319">
E.
<normalizedToken id="8FDD61643184E21AC58B6E45D47DA297" originalValue="Álvarez">Alvarez</normalizedToken>
</collectorName>
,
<collectorName id="244C0726C771D5AC3DA4047F444F7AD8">C. Nieto</collectorName>
, leg.;
<collectingDate id="2C1671A093B777CC50031A2633DF154B" value="2017-04-19" valueMax="2017-04-23" valueMin="2017-04-19">19-23.iv.2017</collectingDate>
; MUPADI • ibid.,
<specimenCount id="B5B2449527C4699F34A903C72CD1B47C" type="male">2 males</specimenCount>
; Quebrada sin nombre; PSPSCB-NPSF-C-097-2017-006;
<geoCoordinate id="1C27743BB112091B21AB6FC69416A3AD" degrees="8.55038" direction="north" orientation="latitude" precision="1" value="8.55038">8.55038°N</geoCoordinate>
,
<geoCoordinate id="9E66C05F8B1BDC5BEDB84FEDAF5B279B" degrees="81.16486" direction="west" orientation="longitude" precision="1" value="-81.16486">81.16486°W</geoCoordinate>
;
<elevation id="AC49C2FFA172970F88862EA2AC019B6C" metricMagnitude="2" metricUnit="m" metricValue="5.15" unit="m" value="515.0">
<quantity id="D14CBD3DEE7EE283E30DB2610C2CB23D" metricMagnitude="2" metricUnit="m" metricValue="5.15" unit="m" value="515.0">515 m</quantity>
a.s.l.
</elevation>
;
<collectingMethod id="801E377BACDEB45A526229ECCF56DBF8">Malaise trap</collectingMethod>
;
<collectorName id="838823B41E1594A8B33B5A3D001517B4">A. Cornejo</collectorName>
,
<collectorName id="BB72DB77120342B677A9E4F6F301E56D">
T.
<normalizedToken id="767F1934088CD9E74F9E13ECB6438906" originalValue="Ríos">Rios</normalizedToken>
</collectorName>
,
<collectorName id="015E84CE0DAC9B721B02B187BF9E0A64">
E.
<normalizedToken id="1C43A068BE6C72B9A998346D0047997A" originalValue="Álvarez">Alvarez</normalizedToken>
</collectorName>
,
<collectorName id="85738C0C2C8E0191FE41E746D151718A">C. Nieto</collectorName>
, leg.;
<collectingDate id="EA5BD0984E9B5F290065469F9C1D0D7E" value="2017-04-23" valueMax="2017-04-27" valueMin="2017-04-23">23-27.iv.2017</collectingDate>
; MIUP • ibid.,
<specimenCount id="E5023C56963655EE880ADE6112FF651D" type="male">1 male</specimenCount>
;
<collectorName id="AC41C24809F0E510D7C2555164AC536A">Quebrada</collectorName>
sin nombre; PSPSCB-NPSF-C-097-2017-011;
<geoCoordinate id="308984E695641F4EAFE15B3240411897" degrees="8.55343" direction="north" orientation="latitude" precision="1" value="8.55343">8.55343°N</geoCoordinate>
,
<geoCoordinate id="9D3E0DE083336197DDEB3AD45E9E893F" degrees="81.17675" direction="west" orientation="longitude" precision="1" value="-81.17675">81.17675°W</geoCoordinate>
;
<elevation id="D27F6EF02DF9953C43D1BBFAD1F7F295" metricMagnitude="2" metricUnit="m" metricValue="3.95" unit="m" value="395.0">
<quantity id="B2AAB6CAEA35A1C5794FD2D6EBCF28AC" metricMagnitude="2" metricUnit="m" metricValue="3.95" unit="m" value="395.0">395 m</quantity>
a.s.l.
</elevation>
;
<collectingMethod id="0C49BC77986E17361668242826E29AD4">UV light trap</collectingMethod>
;
<collectorName id="AF853F765C550A9600239CF0174AC19A">A. Cornejo</collectorName>
,
<collectorName id="FF6E348FBD791A143F85207A915CE139">
T.
<normalizedToken id="5413628F73187B20E03AD5768B9C4176" originalValue="Ríos">Rios</normalizedToken>
</collectorName>
,
<collectorName id="2FF1CD66BA435899B9A970DAC75A1FCF">
E.
<normalizedToken id="A46FAF65CD54C3364D3D4AA5093747EF" originalValue="Álvarez">Alvarez</normalizedToken>
</collectorName>
,
<collectorName id="C247E2782431D21D733C22ADF6A9D971">C. Nieto</collectorName>
, leg.;
<collectingDate id="776DD012C2B67D7B5315991B6D846744" value="2017-04-20">20.iv.2017</collectingDate>
; MIUP
</materialsCitation>
.
<materialsCitation id="99FD27BFF1669F9A3FCA76B133225745" collectingDate="2018-05-23" collectingDateMax="2018-05-31" collectingDateMin="2018-05-23" collectingMethod="Malaise trap" collectorName="Alto Campana NP, T. Rios, Y. Aguirre, E. Perez" country="Panama" county="Cuenca" elevation="497" latitude="8.7165" location="Rio Cacaito" longLatPrecision="1" longitude="-80.0074" municipality="Capira District" specimenCount="1" specimenCount-male="1" stateProvince="Oeste Province">
<emphasis id="0150C69641D036956B12D128A3C29DF0" bold="true" pageId="0" pageNumber="35">
<collectingCountry id="5DCE9781E72C75A0DD77F9189B488354" name="Panama">Panama</collectingCountry>
<collectingRegion id="8D0EA4C5EDFCD52ED7966B85128B7261" inferredBySuffix="true">Oeste Province</collectingRegion>
</emphasis>
•
<specimenCount id="CA361DBE273D89F1BC2679F8CA4A44D2" type="male">1 male</specimenCount>
;
<collectingCounty id="03223CBC606F69D210C1889B6E07732B">Cuenca</collectingCounty>
115;
<collectingMunicipality id="E6C77A19CBEDD0119B1C2F908C560304">Capira District</collectingMunicipality>
;
<collectorName id="88D2EC97B8E896CDD9C8C795750DA3B1">Alto Campana NP</collectorName>
;
<location id="72AB58557C3443D99C639E20F4585099" LSID="urn:lsid:plazi:treatment:495EB4508B815496B0C4918B36F0C29C:72AB58557C3443D99C639E20F4585099" country="Panama" county="Cuenca" latitude="8.7165" longLatPrecision="1" longitude="-80.0074" municipality="Capira District" name="Rio Cacaito" stateProvince="Oeste Province">
<normalizedToken id="234422CB742FAF0BA138F7F5113006CC" originalValue="Río">Rio</normalizedToken>
Cacaito
</location>
; PSPSCB-PNAC-C115-2018-028;
<geoCoordinate id="69BD30AF73ED16F849F9063F0F325FF2" degrees="8.71650" direction="north" orientation="latitude" precision="1" value="8.7165">8.71650°N</geoCoordinate>
,
<geoCoordinate id="8D6F4BDA2A05F8B122C118BE71F2E35F" degrees="80.00740" direction="west" orientation="longitude" precision="1" value="-80.0074">80.00740°W</geoCoordinate>
;
<elevation id="AC10171D8D66F6406535EC21D98485FE" metricMagnitude="2" metricUnit="m" metricValue="4.97" unit="m" value="497.0">
<quantity id="741EE4B31E18104313595159B7F9044A" metricMagnitude="2" metricUnit="m" metricValue="4.97" unit="m" value="497.0">497 m</quantity>
a.s.l.
</elevation>
;
<collectingMethod id="AF80ABF6E537C418756DC62CCBCDAADC">Malaise trap</collectingMethod>
;
<collectorName id="0A06AA0CF99B1C49F7E42EFC863F1203">
T.
<normalizedToken id="FC5A076D5A0E383699A965E128782043" originalValue="Ríos">Rios</normalizedToken>
</collectorName>
,
<collectorName id="247BB695612E3BDE1095EC78F37E7596">Y. Aguirre</collectorName>
,
<collectorName id="4A5D536656497FC498D657B1704E0743">
E.
<normalizedToken id="113E29AB7B48F20696B194F34483C8B0" originalValue="Pérez">Perez</normalizedToken>
</collectorName>
;
<collectingDate id="EC6D613D34E56836070AA77A18E0B0D5" value="2018-05-23" valueMax="2018-05-31" valueMin="2018-05-23">23-31.v.2018</collectingDate>
; MUPADI
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C38F30E2076F702894EE5593A0E91F0E" pageId="0" pageNumber="35" type="diagnosis">
<paragraph id="D8E7546F92CD524D7A09A4A37CBE7D11" pageId="0" pageNumber="35">Diagnosis.</paragraph>
<paragraph id="300488B3E29CAAD5014D918C1DD2D4F3" pageId="0" pageNumber="35">
This species is placed in the
<taxonomicName id="8053BA36202F2ADBB38BA6BA13D1A5FA" authorityName="OS Flint" authorityYear="1968" class="Insecta" family="Hydroptilidae" genus="Alisotrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alisotrichia orophila" order="Trichoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="orophila">
<emphasis id="BC967294B51EC52B6FCE64647C43A3C9" italics="true" pageId="0" pageNumber="35">Alisotrichia orophila</emphasis>
</taxonomicName>
group of
<normalizedToken id="54D8E2BB3780E0026B3BB4D9EDDB7D9D" originalValue="Oláh">Olah</normalizedToken>
and Flint (2012) on the basis of the dorsolateral process from segment VIII which bears an elongate seta, with closest similarity to
<taxonomicName id="3B808E19B1F149142A95C63553781C38" lsidName="A. neblina" pageId="0" pageNumber="35" rank="species" species="neblina">
<emphasis id="F74B168C2B46CD6C6FDDB73D9E8B540F" italics="true" pageId="0" pageNumber="35">A. neblina</emphasis>
</taxonomicName>
Harris &amp; Flint and
<taxonomicName id="63D062551206C769008FE62EA7B26514" lsidName="A. coclensis" pageId="0" pageNumber="35" rank="species" species="coclensis">
<emphasis id="98C4CEC49B6B88CE7080FD5DE814DEF0" italics="true" pageId="0" pageNumber="35">A. coclensis</emphasis>
</taxonomicName>
Armitage &amp; Harris. Like these species,
<taxonomicName id="7E558567BC61C352C22F070F19FD0A91" lsidName="A. eisbergae" pageId="0" pageNumber="35" rank="species" species="eisbergae">
<emphasis id="EFBEDD0E0E90CA86A435C25C31A34ABC" italics="true" pageId="0" pageNumber="35">A. eisbergae</emphasis>
</taxonomicName>
sp. nov. has posterior processes and spines from the margin of segment VIII, but the new species is distinguished by the shape of these spines and by the phallic structure.
</paragraph>
<caption id="6F258AE2C54B035B6BB0C80608E6D6C2" doi="10.3897/zookeys.1174.107314.figure2" httpUri="https://binary.pensoft.net/fig/890127" pageId="0" pageNumber="35" start="Figure 2" startId="F2">
<paragraph id="72187DE1612921B9E9717F9F0C3A92E4" pageId="0" pageNumber="35">
<emphasis id="78EAE9D6BCD9E4F1F6EC7D3B532332BD" bold="true" pageId="0" pageNumber="35">Figure 2.</emphasis>
<taxonomicName id="7F3DEB7A2496766B09AC6F940031E20A" authorityName="Harris &amp; Armitage" authorityYear="2023" class="Insecta" family="Hydroptilidae" genus="Alisotrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alisotrichia eisbergae" order="Trichoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="eisbergae">
<emphasis id="B5E7FC99A46A476026E5D52271293E48" italics="true" pageId="0" pageNumber="35">Alisotrichia eisbergae</emphasis>
</taxonomicName>
sp. nov., male holotype, genitalia
<emphasis id="DCDA010A6620FF7974A04E3A963FD7B6" bold="true" pageId="0" pageNumber="35">A</emphasis>
left lateral
<emphasis id="9F8A99FDC59F0776456829FB614E3D15" bold="true" pageId="0" pageNumber="35">B</emphasis>
variation in apex of segment VIIl
<emphasis id="BE075E152B0A6F88878902CE6706DAAD" bold="true" pageId="0" pageNumber="35">C</emphasis>
dorsal
<emphasis id="2B4EDF31DE51ED6FF533B6243CB859AD" bold="true" pageId="0" pageNumber="35">D</emphasis>
ventral
<emphasis id="E005CCE8C3D2F113E7AF423D35147D49" bold="true" pageId="0" pageNumber="35">E</emphasis>
antenna, basal segments
<emphasis id="C0BD2C9C77E74977AD3F892337E01F3F" bold="true" pageId="0" pageNumber="35">F</emphasis>
maxillary palp
<emphasis id="E34F99CF2FB64DE9497DE75D50AD38A8" bold="true" pageId="0" pageNumber="35">G</emphasis>
phallus, dorsal
<emphasis id="73F0D33188FC6BA9018386DFBFD56873" bold="true" pageId="0" pageNumber="35">H</emphasis>
phallus, left lateral.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="25F443062FCA0C7A346A24F6AA688110" pageId="0" pageNumber="35" type="description">
<paragraph id="EF5BA3B4667026C4C7EA582FBED9872B" pageId="0" pageNumber="35">Description.</paragraph>
<paragraph id="C9A05BB809156C46E01E41844583337D" pageId="0" pageNumber="35">
<emphasis id="556BAE4A9BADF87FB0C0FA491D757A10" bold="true" pageId="0" pageNumber="35">Male.</emphasis>
Total length 1.6-1.8 mm, 17 antennal segments, scape enlarged, pedicel twice as long as proximal flagellomeres, maxillary palp with five segments, brown in alcohol with no obvious patterns on wings.
<emphasis id="8A1368604E4B944CF55BFE9A16FCD17B" bold="true" pageId="0" pageNumber="35">
G
<emphasis id="0D1C43EFA90C205A054A4A57F6E8F4F6" italics="true" pageId="0" pageNumber="35">enitalia</emphasis>
.
</emphasis>
Abdominal segment VII annular without ventromesal process. Segment VIII incomplete dorsally, setal-bearing process dorsally, pair of elongate processes posteriorly; in dorsal view, elongate seta emanating from apex of elongate, narrow, lateral lobes, posterior spines from mesal margins; in ventral view emarginated posteriorly with mesal spines. Segment IX elongate, anteriorly narrowing to short apodeme; in dorsal view narrowing to rounded distal margin; in ventral view narrow, mesal narrow process with crenulate outer margins. Segment X apparently fused with IX and indistinct laterally. Phallus tubular, narrowing at midlength, posteriorly with pair of lateral thin, lateral bands adjacent to and extending beyond the ejaculatory duct; in lateral view wide basally and distally, narrow, spinelike band mesally.
</paragraph>
</subSubSection>
<subSubSection id="6916D58CBE0F7C7B8789DCE22E8BA447" pageId="0" pageNumber="35" type="distribution">
<paragraph id="1E877ACB553F4DF08196DE2DFDE074F3" pageId="0" pageNumber="35">Distribution.</paragraph>
<paragraph id="74D86A0782D35B2FDC05ADCE46672027" pageId="0" pageNumber="35">Panama.</paragraph>
</subSubSection>
<subSubSection id="0457B66B7EDDCFD47E79BB28CE94D947" pageId="0" pageNumber="35" type="etymology">
<paragraph id="2DCBD56661439FA5D41923E216E92119" pageId="0" pageNumber="35">Etymology.</paragraph>
<paragraph id="1740F4E59AF76BECE0C9BC2D2B2239FF" pageId="0" pageNumber="35">This species is named for Ms. Deborah Eisberg of Boquete, Panama in recognition and thanks for her support of our research program. The species name is a female noun in the genitive case.</paragraph>
</subSubSection>
<subSubSection id="50ED3BFAA24FC2830EAE52A70AF5D9B2" pageId="0" pageNumber="35" type="remarks">
<paragraph id="F865E1D64DCC4A5782C9B7CB5F54F563" pageId="0" pageNumber="35">Remarks.</paragraph>
<paragraph id="0B85E125C5BD755216B24F4232448D52" pageId="0" pageNumber="35">
The pair of elongate processes on segment VIII differed slightly in the specimen from the
<normalizedToken id="E0177E115A4C72B191B4211DA39D8B2A" originalValue="Río">Rio</normalizedToken>
Cacaito (Panama Oeste Province) compared to the other specimens examined.
</paragraph>
</subSubSection>
</treatment>
</document>