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<mods:title id="B6EA024F6E4BFE6129C693E77199C974">New specimens of Saturnalia tupiniquim (Dinosauria: Sauropodomorpha): insights into intraspecific variation, rostral anatomy, and skull size</mods:title>
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<mods:title id="2BA588EA33B8EC746941D198F3ADF93C">New specimens of Saturnalia tupiniquim (Dinosauria: Sauropodomorpha): insights into intraspecific variation, rostral anatomy, and skull size</mods:title>
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@ -173,13 +176,13 @@ Garcia
<emphasis id="B902116DFFE6A868FC47FC31FE46F952" box="[943,1326,1654,1681]" italics="true" pageId="1" pageNumber="2">Referred specimen UFSM 11660 (AN)</emphasis>
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<paragraph id="8BC9CD7FFFE6A868FCC2FCDAFEDBF80D" blockId="1.[809,1460,1654,1999]" pageId="1" pageNumber="2">An association of at least three individuals with different degrees of articulation. It comprises a partial skull with braincase, lower jaws, vertebrae from distinct portions of the column, ribs, chevrons, pelvic girdle, femora, tibiae, metatarsals, and phalanges. Because of its gracile morphology, the specimen was affectionately nicknamed as Gracinha, meaning graceful in Portuguese. The specific preserved elements are: UFSM 11660A, skull, right maxilla, braincase elements; UFSM 11660B, dorsal vertebrae; UFSM 11660C, sacrum articulated against the ilium, ischium, pubis; UFSM 11660D, partial sacrum; UFSM 11660E, damaged sacrum;</paragraph>
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<emphasis id="B902116DFFE5A86BFF69FD68FBB0F884" bold="true" box="[129,216,1839,1863]" pageId="2" pageNumber="3">Figure 1.</emphasis>
Study area and geological context of UFSM 11660: (A) location and geological context of Cerro da Alemoa outcrop, (B) general view of the Cerro da Alemoa outcrop, (C) plaster jacket containing the elements of UFSM 11660 partially exposed and reconstruction of preserved elements of UFSM 11660 (artwork by Maurício S. Garcia). The sites indicating where the type-specimens were found are based on the recollection of one of the authors (M.C.L.), who excavated the three specimens back in 1998; henceforth, they are not totally precise.
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<emphasis id="B902116DFFE4A86AFF99FAC8FBA8FF65" bold="true" box="[113,192,142,167]" pageId="3" pageNumber="4">Table 1.</emphasis>
Measurements employed in order to estimate skull length of UFSM 11660 and
@ -387,7 +390,7 @@ S
W
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), in the municipality of Santa Maria (
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<figureCitation id="134DD1FAFFE4A86AFEF3FE00FA3DFB9C" box="[283,341,1095,1119]" captionStart="Figure 1" captionStartId="2.[129,194,1839,1863]" captionTargetBox="[179,1423,145,1810]" captionTargetId="figure-9@2.[178,1424,144,1811]" captionTargetPageId="2" captionText="Figure 1. Study area and geological context of UFSM 11660: (A) location and geological context of Cerro da Alemoa outcrop, (B) general view of the Cerro da Alemoa outcrop, (C) plaster jacket containing the elements of UFSM 11660 partially exposed and reconstruction of preserved elements of UFSM 11660 (artwork by Maurício S. Garcia). The sites indicating where the type-specimens were found are based on the recollection of one of the authors (M.C.L.), who excavated the three specimens back in 1998; henceforth, they are not totally precise." figureDoi="http://doi.org/10.5281/zenodo.14827319" httpUri="https://zenodo.org/record/14827319/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
), which exposes rocks of the Candelária Sequence (
<bibRefCitation id="EFE7B08EFFE4A86AFF0DFE20FAE3FBBD" author="Horn BLD &amp; Melo TM &amp; Schultz CL" box="[229,395,1126,1150]" pageId="3" pageNumber="4" pagination="123 - 32" refId="ref14227" refString="Horn BLD, Melo TM, Schultz CL et al. A new third-order sequence stratigraphic frame-work applied to the Triassic of the Parana Basin, Rio Grande do Sul, Brazil, based on structural, stratigraphic and paleontological data. Journal of South American Earth Sciences 2014; 55: 123 - 32. https: // doi. org / 10.1016 / j. jsames. 2014.07.007" type="journal article" year="2014">
Horn
@ -504,10 +507,10 @@ Griffin and Nesbitt 2016
</paragraph>
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The description followed the veterinarian anatomical terms (i.e. cranial/rostral and caudal are used rather than anterior and posterior), the complete list of specimens used for comparison is in
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<tableCitation id="C6F4F8C4FFE4A86AFCB1F94AF8CCFCE5" box="[857,932,781,806]" captionStart="Table 2" captionStartId="4.[129,184,143,167]" captionTargetPageId="4" captionText="Table 2. Sources of anatomical data used for comparison" httpUri="http://table.plazi.org/id/DF099DF7FFE3A86DFF69FAC8F9FCFF64" pageId="3" pageNumber="4" tableUuid="DF099DF7FFE3A86DFF69FAC8F9FCFF64">Table 2</tableCitation>
and measurements of UFSM
<quantity id="4C8E609AFFE4A86AFB3BF94AFE45FCE6" box="[1235,1325,781,805]" metricMagnitude="2" metricUnit="m" metricValue="2.96164" pageId="3" pageNumber="4" unit="in" value="11660.0">11660 in</quantity>
<tableCitation id="C6F4F8C4FFE4A86AFADAF94AFE15FCE5" box="[1330,1405,781,806]" captionStart="Table 3" captionStartId="5.[113,168,143,167]" captionTargetPageId="5" captionText="Table 3. Measurements of the elements of UFSM 11600 (in mm)." pageId="3" pageNumber="4">Table 3</tableCitation>
<tableCitation id="C6F4F8C4FFE4A86AFADAF94AFE15FCE5" box="[1330,1405,781,806]" captionStart="Table 3" captionStartId="5.[113,168,143,167]" captionTargetPageId="5" captionText="Table 3. Measurements of the elements of UFSM 11600 (in mm)." httpUri="http://table.plazi.org/id/DF099DF7FFE2A86CFF99FAC8F9B6FF64" pageId="3" pageNumber="4" tableUuid="DF099DF7FFE2A86CFF99FAC8F9B6FF64">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8BC9CD7FFFE4A86AFBF2F902FFAAFC9C" blockId="3.[810,1461,837,1056]" box="[1050,1218,837,863]" pageId="3" pageNumber="4">
@ -515,13 +518,13 @@ and measurements of UFSM
</paragraph>
<paragraph id="8BC9CD7FFFE4A86AFCC2F92CF8A7FBE3" blockId="3.[810,1461,837,1056]" pageId="3" pageNumber="4">
The association includes one skull with articulated to partially articulated elements (
<figureCitation id="134DD1FAFFE4A86AFBE4F9CCFF2FFC60" box="[1036,1095,907,931]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." pageId="3" pageNumber="4">Fig. 2</figureCitation>
<figureCitation id="134DD1FAFFE4A86AFBE4F9CCFF2FFC60" box="[1036,1095,907,931]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." figureDoi="http://doi.org/10.5281/zenodo.14827321" httpUri="https://zenodo.org/record/14827321/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
). Hence, the following description of the rostral region, braincase, and partial lower jaws refers to a single individual. The bone surface of the elements is usually well-preserved, but the extremities of some bones are not entirely preserved.
</paragraph>
<paragraph id="8BC9CD7FFFE4A86AFCC2FE06FEDBF8C9" blockId="3.[810,1462,1089,1803]" pageId="3" pageNumber="4">
<emphasis id="B902116DFFE4A86AFCC2FE06F8F5FB9A" box="[810,925,1089,1113]" italics="true" pageId="3" pageNumber="4">Premaxilla:</emphasis>
Both premaxillae are preserved, although slightly displaced from their original position (
<figureCitation id="134DD1FAFFE4A86AFB29FE26FF93FBBA" box="[1217,1275,1121,1145]" captionStart="Figure 3" captionStartId="8.[129,194,1522,1546]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-187@8.[129,1473,144,1495]" captionTargetPageId="8" captionText="Figure 3. Premaxilla and maxilla of UFSM 11660A:(A, B) lateral and (C, D) medial views. Abbreviations: clp, caudolateral process; dp, dorsal process; f, foramen; idp, interdental plate; l.mx, left maxilla; mk, Meckelian groove; mp, medial process; n, nasal; r.d, right dentary; pm, premaxilla; pmf, premaxillary foramen; t, tooth." pageId="3" pageNumber="4">Fig. 3</figureCitation>
<figureCitation id="134DD1FAFFE4A86AFB29FE26FF93FBBA" box="[1217,1275,1121,1145]" captionStart="Figure 3" captionStartId="8.[129,194,1522,1546]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-187@8.[129,1473,144,1495]" captionTargetPageId="8" captionText="Figure 3. Premaxilla and maxilla of UFSM 11660A:(A, B) lateral and (C, D) medial views. Abbreviations: clp, caudolateral process; dp, dorsal process; f, foramen; idp, interdental plate; l.mx, left maxilla; mk, Meckelian groove; mp, medial process; n, nasal; r.d, right dentary; pm, premaxilla; pmf, premaxillary foramen; t, tooth." figureDoi="http://doi.org/10.5281/zenodo.14827323" httpUri="https://zenodo.org/record/14827323/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
). The bone has a convex rostroventral margin and the narial fossa occupies the dorsal half of its main body. The right premaxilla preserved the caudolateral and medial processes, whereas only the dorsal process is present in the left bone. The anterior premaxillary foramen (
<emphasis id="B902116DFFE4A86AFC9BFEB9F8CEFAD6" box="[883,934,1278,1301]" italics="true" pageId="3" pageNumber="4">sensu</emphasis>
<bibRefCitation id="EFE7B08EFFE4A86AFC44FEB9FF09FAD5" author="Sereno PC &amp; Martinez RN &amp; Alcober OA" box="[940,1121,1277,1302]" pageId="3" pageNumber="4" pagination="83 - 179" refId="ref16388" refString="Sereno PC, Martinez RN, Alcober OA. Osteology of Eoraptor lunensis (Dinosauria, Sauropodomorpha). Journal of Vertebrate Paleontology 2013; 32: 83 - 179." type="journal article" year="2013">
@ -575,9 +578,9 @@ Müller
<paragraph id="8BC9CD7FFFE4A86DFCC2FD6BFA86FB18" blockId="3.[810,1459,1836,1985]" lastBlockId="4.[127,779,843,1243]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B902116DFFE4A86AFCC2FD6BF8E8F887" box="[810,896,1836,1860]" italics="true" pageId="3" pageNumber="4">Maxilla:</emphasis>
Both maxillae are preserved, but only the left one in its full length (
<figureCitation id="134DD1FAFFE4A86AFC2AFD0CF897F8A7" box="[962,1023,1867,1892]" captionStart="Figure 3" captionStartId="8.[129,194,1522,1546]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-187@8.[129,1473,144,1495]" captionTargetPageId="8" captionText="Figure 3. Premaxilla and maxilla of UFSM 11660A:(A, B) lateral and (C, D) medial views. Abbreviations: clp, caudolateral process; dp, dorsal process; f, foramen; idp, interdental plate; l.mx, left maxilla; mk, Meckelian groove; mp, medial process; n, nasal; r.d, right dentary; pm, premaxilla; pmf, premaxillary foramen; t, tooth." pageId="3" pageNumber="4">Figs 3</figureCitation>
<figureCitation id="134DD1FAFFE4A86AFC2AFD0CF897F8A7" box="[962,1023,1867,1892]" captionStart="Figure 3" captionStartId="8.[129,194,1522,1546]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-187@8.[129,1473,144,1495]" captionTargetPageId="8" captionText="Figure 3. Premaxilla and maxilla of UFSM 11660A:(A, B) lateral and (C, D) medial views. Abbreviations: clp, caudolateral process; dp, dorsal process; f, foramen; idp, interdental plate; l.mx, left maxilla; mk, Meckelian groove; mp, medial process; n, nasal; r.d, right dentary; pm, premaxilla; pmf, premaxillary foramen; t, tooth." figureDoi="http://doi.org/10.5281/zenodo.14827323" httpUri="https://zenodo.org/record/14827323/files/figure.png" pageId="3" pageNumber="4">Figs 3</figureCitation>
,
<figureCitation id="134DD1FAFFE4A86AFBE3FD0BFF70F8A7" box="[1035,1048,1868,1892]" captionStart="Figure 4" captionStartId="9.[113,178,1764,1788]" captionTargetBox="[114,1458,144,1737]" captionTargetId="figure-6@9.[114,1458,144,1737]" captionTargetPageId="9" captionText="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." pageId="3" pageNumber="4">4</figureCitation>
<figureCitation id="134DD1FAFFE4A86AFBE3FD0BFF70F8A7" box="[1035,1048,1868,1892]" captionStart="Figure 4" captionStartId="9.[113,178,1764,1788]" captionTargetBox="[114,1458,144,1737]" captionTargetId="figure-6@9.[114,1458,144,1737]" captionTargetPageId="9" captionText="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." figureDoi="http://doi.org/10.5281/zenodo.14827326" httpUri="https://zenodo.org/record/14827326/files/figure.png" pageId="3" pageNumber="4">4</figureCitation>
). The rostral process is short and its lateral surface bears some foramina. As in
<taxonomicName id="4C76B6FCFFE4A86AFB54FD2BFE32F840" box="[1212,1370,1899,1923]" genus="Pampadromaeus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="barberenai">
<emphasis id="B902116DFFE4A86AFB54FD2BFE32F840" box="[1212,1370,1899,1923]" italics="true" pageId="3" pageNumber="4">Pam. barberenai</emphasis>
@ -589,10 +592,10 @@ Müller
2018c
</bibRefCitation>
), the rostralmost portion of the antorbital fossa lacks any evidence of a promaxillary fenestra, or of any other excavation. Both internal and external antorbital fenestrae are rostrocaudally elongated. The rounded rostroventral corner of the internal antorbital fenestra forms a straight to slightly obtuse angle (
<figureCitation id="134DD1FAFFE3A86DFE39FE02F975FB9E" box="[465,541,1093,1117]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." pageId="4" pageNumber="5">Fig. 2C</figureCitation>
<figureCitation id="134DD1FAFFE3A86DFE39FE02F975FB9E" box="[465,541,1093,1117]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." figureDoi="http://doi.org/10.5281/zenodo.14827321" httpUri="https://zenodo.org/record/14827321/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
). The maxillary caudal process is elongated and tapers at the distalmost portion, which has a longitudinally elongated slot on its dorsolateral margin to receive the jugal. The tooth margin is straight along its entire length and bears 19 tooth positions.
</paragraph>
<caption id="DF099DF7FFE3A86DFF69FAC8F9FCFF64" box="[129,660,142,167]" pageId="4" pageNumber="5" startId="4.[129,184,143,167]" targetBox="[129,1474,188,775]" targetIsTable="true" targetPageId="4" targetType="table">
<caption id="DF099DF7FFE3A86DFF69FAC8F9FCFF64" ID-Table-UUID="DF099DF7FFE3A86DFF69FAC8F9FCFF64" box="[129,660,142,167]" httpUri="http://table.plazi.org/id/DF099DF7FFE3A86DFF69FAC8F9FCFF64" pageId="4" pageNumber="5" startId="4.[129,184,143,167]" targetBox="[129,1474,188,775]" targetIsTable="true" targetPageId="4" targetType="table">
<paragraph id="8BC9CD7FFFE3A86DFF69FAC8F9FCFF64" blockId="4.[129,660,142,167]" box="[129,660,142,167]" pageId="4" pageNumber="5">
<emphasis id="B902116DFFE3A86DFF69FAC8FBA6FF65" bold="true" box="[129,206,142,167]" pageId="4" pageNumber="5">Table 2.</emphasis>
Sources of anatomical data used for comparison
@ -819,7 +822,7 @@ Leal
<paragraph id="8BC9CD7FFFE3A86DFF69FEBBF97FF9ED" blockId="4.[129,778,1276,1582]" pageId="4" pageNumber="5">
<emphasis id="B902116DFFE3A86DFF69FEBBFBA9FAD7" box="[129,193,1276,1300]" italics="true" pageId="4" pageNumber="5">Nasal:</emphasis>
The specimen preserves the left nasal, which is not completely preserved (
<figureCitation id="134DD1FAFFE3A86DFEA1FF5CFAEFFAF0" box="[329,391,1307,1331]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." pageId="4" pageNumber="5">Fig. 2</figureCitation>
<figureCitation id="134DD1FAFFE3A86DFEA1FF5CFAEFFAF0" box="[329,391,1307,1331]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." figureDoi="http://doi.org/10.5281/zenodo.14827321" httpUri="https://zenodo.org/record/14827321/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
). It corresponds to a thin blade of bone, with a dorsal recess in its lateral side. In dorsal view, the rostral process tapers medially. In lateral view, it projects ventrally to form the dorsal rim of the external nares. The lateral margin of the bone is poorly preserved. As a consequence, it is uncertain if it forms a well-projected shelf, as in other early sauropodomorphs (
<bibRefCitation id="EFE7B08EFFE3A86DFEBAFF90F91FFA2C" author="Martinez RN &amp; Alcober OA" box="[338,631,1495,1519]" pageId="4" pageNumber="5" refId="ref15072" refString="Martinez RN, Alcober OA. A basal sauropodomorph (Dinosauria: Saurischia) from the Ischigualasto formation (Triassic, Carnian) and the early evolution of Sauropodomorpha. PLoS One 2009; 4: e 4397 - 12. https: // doi. org / 10.1371 / journal. pone. 0004397" type="journal volume" year="2009">Martínez and Alcober 2009</bibRefCitation>
,
@ -839,7 +842,7 @@ Müller
<paragraph id="8BC9CD7FFFE3A86DFF69FC08F9E3F803" blockId="4.[128,780,1615,1984]" pageId="4" pageNumber="5">
<emphasis id="B902116DFFE3A86DFF69FC08FB89F9A4" box="[129,225,1615,1639]" italics="true" pageId="4" pageNumber="5">Lacrimal:</emphasis>
Only the left lacrimal is preserved. It has an inverted L-shape and contributes to the dorsal and caudal margins of the antorbital fenestra (
<figureCitation id="134DD1FAFFE3A86DFE99FCC9FAC1F965" box="[369,425,1678,1702]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." pageId="4" pageNumber="5">Fig. 2</figureCitation>
<figureCitation id="134DD1FAFFE3A86DFE99FCC9FAC1F965" box="[369,425,1678,1702]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." figureDoi="http://doi.org/10.5281/zenodo.14827321" httpUri="https://zenodo.org/record/14827321/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
). The bone also contributes to the caudal margin of the antorbital fossa, which invades the ventral and anterior processes. The ventral process is slightly longer than the rostral. The lateral margin of the rostral process folds ventrally, forming a ventral crest that extends longitudinally. The lateral surface of the boundary between the rostral and ventral processes is slightly striated. Yet, this region lacks the flange present in
<taxonomicName id="4C76B6FCFFE3A86DFF3DFD2DFA23F842" box="[213,331,1897,1921]" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="schultzi">
<emphasis id="B902116DFFE3A86DFF3DFD2DFA23F842" box="[213,331,1897,1921]" italics="true" pageId="4" pageNumber="5">Bu. schultzi</emphasis>
@ -851,7 +854,7 @@ Only the left lacrimal is preserved. It has an inverted L-shape and contributes
<paragraph id="8BC9CD7FFFE3A86DFCD1F90CF8FCFBDC" blockId="4.[825,1476,843,1055]" pageId="4" pageNumber="5">
<emphasis id="B902116DFFE3A86DFCD1F90CF8CAFCA0" box="[825,930,843,867]" italics="true" pageId="4" pageNumber="5">Prefrontal:</emphasis>
The left prefrontal is disarticulated and preserves the caudal process (
<figureCitation id="134DD1FAFFE3A86DFC35F92DFF7EFC41" box="[989,1046,874,898]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." pageId="4" pageNumber="5">Fig. 2</figureCitation>
<figureCitation id="134DD1FAFFE3A86DFC35F92DFF7EFC41" box="[989,1046,874,898]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." figureDoi="http://doi.org/10.5281/zenodo.14827321" httpUri="https://zenodo.org/record/14827321/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
). The preserved dorsal portion is flat. The corner between the dorsal and lateral surfaces is convex in lateral view. The caudal process tappers caudally and is slightly concave in lateral view. Its general morphology resembles that of coeval sauropodomorphs (e.g.
<taxonomicName id="4C76B6FCFFE3A86DFBC7F9A0FFB0FC3C" authorityName="Martínez &amp; Alcober" authorityYear="2009" box="[1071,1240,999,1023]" class="Reptilia" family="Guaibasauridae" genus="Panphagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="protos">
<emphasis id="B902116DFFE3A86DFBC7F9A0FFB0FC3C" box="[1071,1240,999,1023]" italics="true" pageId="4" pageNumber="5">Panphagia protos</emphasis>
@ -903,7 +906,7 @@ Martínez
<paragraph id="8BC9CD7FFFE3A86DFCD1FF12F8C0F926" blockId="4.[825,1475,1365,1765]" pageId="4" pageNumber="5">
<emphasis id="B902116DFFE3A86DFCD1FF12F81BFAAE" box="[825,883,1365,1389]" italics="true" pageId="4" pageNumber="5">Jugal:</emphasis>
The jugal is not entirely preserved (
<figureCitation id="134DD1FAFFE3A86DFAF4FF12FE33FAAE" box="[1308,1371,1365,1389]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." pageId="4" pageNumber="5">Fig. 2</figureCitation>
<figureCitation id="134DD1FAFFE3A86DFAF4FF12FE33FAAE" box="[1308,1371,1365,1389]" captionStart="Figure 2" captionStartId="7.[113,178,1213,1237]" captionTargetBox="[114,1437,144,1185]" captionTargetId="figure-421@7.[114,1458,144,1185]" captionTargetPageId="7" captionText="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." figureDoi="http://doi.org/10.5281/zenodo.14827321" httpUri="https://zenodo.org/record/14827321/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
), lacking the caudal portion. The rostral portion of the bone is elongated and gracile, unlike the taller main body of the bone in herrerasaurs (
<bibRefCitation id="EFE7B08EFFE3A86DFC39FFF4FFB3FA08" author="Sereno PC &amp; Novas FE" box="[977,1243,1459,1483]" pageId="4" pageNumber="5" pagination="451 - 76" refId="ref16359" refString="Sereno PC, Novas FE. The skull and neck of the basal theropod Herrerasaurus ischigualastensis. Journal of Vertebrate Paleontology 1993; 13: 451 - 76." type="journal article" year="1993">Sereno and Novas 1993</bibRefCitation>
,
@ -919,16 +922,16 @@ Pacheco
2019
</bibRefCitation>
). The rostral tip is particularly thin dorsoventrally (
<figureCitation id="134DD1FAFFE3A86DFC4CFFB6F8B5F9CA" box="[932,989,1521,1545]" captionStart="Figure 4" captionStartId="9.[113,178,1764,1788]" captionTargetBox="[114,1458,144,1737]" captionTargetId="figure-6@9.[114,1458,144,1737]" captionTargetPageId="9" captionText="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." pageId="4" pageNumber="5">Fig. 4</figureCitation>
<figureCitation id="134DD1FAFFE3A86DFC4CFFB6F8B5F9CA" box="[932,989,1521,1545]" captionStart="Figure 4" captionStartId="9.[113,178,1764,1788]" captionTargetBox="[114,1458,144,1737]" captionTargetId="figure-6@9.[114,1458,144,1737]" captionTargetPageId="9" captionText="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." figureDoi="http://doi.org/10.5281/zenodo.14827326" httpUri="https://zenodo.org/record/14827326/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
), where it articulates with the maxilla ventrally and dorsally to the lacrimal. The preserved part of the jugal has a smooth lateral surface, a sharp ventrolateral corner, and almost parallel dorsal and ventral margins. The dorsal surface is gently convex on the region that forms the ventral margin of the orbit. On the preserved caudal portion of the bone, there is a dorsal expansion (= dorsal process), which is not entirely preserved.
</paragraph>
<paragraph id="8BC9CD7FFFE3A86DFCD1FD40FECEF878" blockId="4.[824,1474,1798,1979]" pageId="4" pageNumber="5">
<emphasis id="B902116DFFE3A86DFCD1FD40F8F6F8DD" box="[825,926,1799,1822]" italics="true" pageId="4" pageNumber="5">Braincase:</emphasis>
Of the braincase elements, only the basioccipital, parabasisphenoid and prootic (
<figureCitation id="134DD1FAFFE3A86DFB97FD62FFD2F8FE" box="[1151,1210,1829,1853]" captionStart="Figure 5" captionStartId="10.[129,194,1301,1325]" captionTargetBox="[129,1473,144,1273]" captionTargetId="figure-361@10.[129,1473,144,1273]" captionTargetPageId="10" captionText="Figure 5. Braincase elements of UFSM 11660A: (A) cranial, (B) dorsal, (C) ventral, and (D) caudal views.Abbreviations:bt, basal tubera; bp, basipterygoid process; cp, cultriform process; fr, floccular recess; icaf, foramina for the internal carotid artery; oc, occipital condyle; pf, pituitary fossa; ssr, subsellar recess." pageId="4" pageNumber="5">Fig. 5</figureCitation>
<figureCitation id="134DD1FAFFE3A86DFB97FD62FFD2F8FE" box="[1151,1210,1829,1853]" captionStart="Figure 5" captionStartId="10.[129,194,1301,1325]" captionTargetBox="[129,1473,144,1273]" captionTargetId="figure-361@10.[129,1473,144,1273]" captionTargetPageId="10" captionText="Figure 5. Braincase elements of UFSM 11660A: (A) cranial, (B) dorsal, (C) ventral, and (D) caudal views.Abbreviations:bt, basal tubera; bp, basipterygoid process; cp, cultriform process; fr, floccular recess; icaf, foramina for the internal carotid artery; oc, occipital condyle; pf, pituitary fossa; ssr, subsellar recess." figureDoi="http://doi.org/10.5281/zenodo.14827328" httpUri="https://zenodo.org/record/14827328/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
) are partially exposed. A future contribution will present additional details with the aid of CT-scan data. Forming the caudal portion of the basioccipital, the occipital condyle is kidney shaped, with a more expanded and concave dorsal portion, tapering ventrally in caudal view.
</paragraph>
<caption id="DF099DF7FFE2A86CFF99FAC8F9B6FF64" box="[113,734,142,167]" pageId="5" pageNumber="6" startId="5.[113,168,143,167]" targetBox="[109,1418,189,1983]" targetIsTable="true" targetPageId="5" targetType="table">
<caption id="DF099DF7FFE2A86CFF99FAC8F9B6FF64" ID-Table-UUID="DF099DF7FFE2A86CFF99FAC8F9B6FF64" box="[113,734,142,167]" httpUri="http://table.plazi.org/id/DF099DF7FFE2A86CFF99FAC8F9B6FF64" pageId="5" pageNumber="6" startId="5.[113,168,143,167]" targetBox="[109,1418,189,1983]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="8BC9CD7FFFE2A86CFF99FAC8F9B6FF64" blockId="5.[113,734,142,167]" box="[113,734,142,167]" pageId="5" pageNumber="6">
<emphasis id="B902116DFFE2A86CFF99FAC8FBD7FF65" bold="true" box="[113,191,142,167]" pageId="5" pageNumber="6">Table 3.</emphasis>
Measurements of the elements of UFSM 11600 (in mm).
@ -1375,7 +1378,7 @@ Bronzati
<paragraph id="8BC9CD7FFFE1A86EFCD1FF8BF916FA09" blockId="6.[823,1476,1483,1977]" lastBlockId="7.[113,762,1364,1482]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B902116DFFE1A86FFCD1FF8BF8F9FA20" box="[825,913,1484,1507]" italics="true" pageId="6" pageNumber="7">Dentary:</emphasis>
Both dentaries are partially preserved (
<figureCitation id="134DD1FAFFE1A86FFAC5FF8CFEECFA20" box="[1325,1412,1483,1507]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="7.[113,178,1213,1237]" captionStartId-1="8.[129,194,1522,1546]" captionStartId-2="9.[113,178,1764,1788]" captionTargetBox-0="[114,1437,144,1185]" captionTargetBox-1="[129,1473,144,1495]" captionTargetBox-2="[114,1458,144,1737]" captionTargetId-0="figure-421@7.[114,1458,144,1185]" captionTargetId-1="figure-187@8.[129,1473,144,1495]" captionTargetId-2="figure-6@9.[114,1458,144,1737]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." captionText-1="Figure 3. Premaxilla and maxilla of UFSM 11660A:(A, B) lateral and (C, D) medial views. Abbreviations: clp, caudolateral process; dp, dorsal process; f, foramen; idp, interdental plate; l.mx, left maxilla; mk, Meckelian groove; mp, medial process; n, nasal; r.d, right dentary; pm, premaxilla; pmf, premaxillary foramen; t, tooth." captionText-2="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." pageId="6" pageNumber="7">Figs 24</figureCitation>
<figureCitation id="134DD1FAFFE1A86FFAC5FF8CFEECFA20" box="[1325,1412,1483,1507]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="7.[113,178,1213,1237]" captionStartId-1="8.[129,194,1522,1546]" captionStartId-2="9.[113,178,1764,1788]" captionTargetBox-0="[114,1437,144,1185]" captionTargetBox-1="[129,1473,144,1495]" captionTargetBox-2="[114,1458,144,1737]" captionTargetId-0="figure-421@7.[114,1458,144,1185]" captionTargetId-1="figure-187@8.[129,1473,144,1495]" captionTargetId-2="figure-6@9.[114,1458,144,1737]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="Figure 2. Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views.Abbreviations:d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal.l. left and r. right." captionText-1="Figure 3. Premaxilla and maxilla of UFSM 11660A:(A, B) lateral and (C, D) medial views. Abbreviations: clp, caudolateral process; dp, dorsal process; f, foramen; idp, interdental plate; l.mx, left maxilla; mk, Meckelian groove; mp, medial process; n, nasal; r.d, right dentary; pm, premaxilla; pmf, premaxillary foramen; t, tooth." captionText-2="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." figureDoi-0="http://doi.org/10.5281/zenodo.14827321" figureDoi-1="http://doi.org/10.5281/zenodo.14827323" figureDoi-2="http://doi.org/10.5281/zenodo.14827326" httpUri-0="https://zenodo.org/record/14827321/files/figure.png" httpUri-1="https://zenodo.org/record/14827323/files/figure.png" httpUri-2="https://zenodo.org/record/14827326/files/figure.png" pageId="6" pageNumber="7">Figs 24</figureCitation>
). The bone is relatively slender, unlike the robust dentary of latter sauropodomorphs.Thedorsalmarginofitsrostralendisventrally deflected, such as in other early sauropodomorphs (Matínez and Alcober 2009,
<bibRefCitation id="EFE7B08EFFE1A86FFC3DFC0FFFFAF9A3" author="Sereno PC &amp; Martinez RN &amp; Alcober OA" box="[981,1170,1608,1632]" pageId="6" pageNumber="7" pagination="83 - 179" refId="ref16388" refString="Sereno PC, Martinez RN, Alcober OA. Osteology of Eoraptor lunensis (Dinosauria, Sauropodomorpha). Journal of Vertebrate Paleontology 2013; 32: 83 - 179." type="journal article" year="2013">
Sereno
@ -1435,7 +1438,7 @@ Leal
<quantity id="4C8E609AFFE1A86FFC0BFDC5FF59F85A" box="[995,1073,1922,1946]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="7" unit="mm" value="2.4">2.4 mm</quantity>
from the rostralmost end of the bone. The rostralmost teeth are slightly rostrally inclined relative to the alveolar margin. The mandibular symphysis is short, restricted to the rostral end of the dentary. In medial view, it is possible to identify the interdental plates, which are triangular. The posterior portion of both dentaries is poorly preserved.
</paragraph>
<caption id="DF099DF7FFE0A86EFF99FEFAF833FB32" pageId="7" pageNumber="8" startId="7.[113,178,1213,1237]" targetBox="[114,1437,144,1185]" targetPageId="7" targetType="figure">
<caption id="DF099DF7FFE0A86EFF99FEFAF833FB32" ID-DOI="http://doi.org/10.5281/zenodo.14827321" ID-Zenodo-Dep="14827321" httpUri="https://zenodo.org/record/14827321/files/figure.png" pageId="7" pageNumber="8" startId="7.[113,178,1213,1237]" targetBox="[114,1437,144,1185]" targetPageId="7" targetType="figure">
<paragraph id="8BC9CD7FFFE0A86EFF99FEFAF833FB32" blockId="7.[113,1448,1213,1265]" pageId="7" pageNumber="8">
<emphasis id="B902116DFFE0A86EFF99FEFAFBA1FB16" bold="true" box="[113,201,1213,1237]" pageId="7" pageNumber="8">Figure 2.</emphasis>
Skull of UFSM 11660A: (A, B) lateral and (C, D) medial views. Abbreviations: d, dentary; fob, fossa for the olfactory bulb; j, jugal; l, lacrimal; mx, maxilla; n, nasal; pm, premaxilla; prf, prefrontal. l. left and
@ -1479,7 +1482,7 @@ Moro
2024
</bibRefCitation>
). The right premaxilla preserves three teeth caudal to an empty tooth position (
<figureCitation id="134DD1FAFFE0A86EFD59FC2FF981F943" box="[689,745,1640,1664]" captionStart="Figure 3" captionStartId="8.[129,194,1522,1546]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-187@8.[129,1473,144,1495]" captionTargetPageId="8" captionText="Figure 3. Premaxilla and maxilla of UFSM 11660A:(A, B) lateral and (C, D) medial views. Abbreviations: clp, caudolateral process; dp, dorsal process; f, foramen; idp, interdental plate; l.mx, left maxilla; mk, Meckelian groove; mp, medial process; n, nasal; r.d, right dentary; pm, premaxilla; pmf, premaxillary foramen; t, tooth." pageId="7" pageNumber="8">Fig. 3</figureCitation>
<figureCitation id="134DD1FAFFE0A86EFD59FC2FF981F943" box="[689,745,1640,1664]" captionStart="Figure 3" captionStartId="8.[129,194,1522,1546]" captionTargetBox="[129,1473,144,1495]" captionTargetId="figure-187@8.[129,1473,144,1495]" captionTargetPageId="8" captionText="Figure 3. Premaxilla and maxilla of UFSM 11660A:(A, B) lateral and (C, D) medial views. Abbreviations: clp, caudolateral process; dp, dorsal process; f, foramen; idp, interdental plate; l.mx, left maxilla; mk, Meckelian groove; mp, medial process; n, nasal; r.d, right dentary; pm, premaxilla; pmf, premaxillary foramen; t, tooth." figureDoi="http://doi.org/10.5281/zenodo.14827323" httpUri="https://zenodo.org/record/14827323/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
). The apicobasal length of the first preserved crown (second premaxillary tooth) is
<quantity id="4C8E609AFFE0A86EFEDDFCE0FA1CF97D" box="[309,372,1703,1726]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="8" unit="mm" value="7.0">7 mm</quantity>
. All teeth lack basal constrictions. The second tooth has serrations (eight per millimetre) only in the mesial margin. These are also seen in the distal margin of the 2
@ -1521,12 +1524,12 @@ Langer
2019
</bibRefCitation>
). The premaxillary teeth are longer and thinner than those of the maxilla (
<tableCitation id="C6F4F8C4FFE0A86EFEE4FDE5FA3FF879" box="[268,343,1954,1978]" captionStart="Table 3" captionStartId="5.[113,168,143,167]" captionTargetPageId="5" captionText="Table 3. Measurements of the elements of UFSM 11600 (in mm)." pageId="7" pageNumber="8">Table 3</tableCitation>
<tableCitation id="C6F4F8C4FFE0A86EFEE4FDE5FA3FF879" box="[268,343,1954,1978]" captionStart="Table 3" captionStartId="5.[113,168,143,167]" captionTargetPageId="5" captionText="Table 3. Measurements of the elements of UFSM 11600 (in mm)." httpUri="http://table.plazi.org/id/DF099DF7FFE2A86CFF99FAC8F9B6FF64" pageId="7" pageNumber="8" tableUuid="DF099DF7FFE2A86CFF99FAC8F9B6FF64">Table 3</tableCitation>
).
</paragraph>
<paragraph id="8BC9CD7FFFE0A86EFCADFF13FF7AF927" blockId="7.[809,1460,1364,1764]" pageId="7" pageNumber="8">
In the maxilla, UFSM 11660A has at least 19 tooth positions (
<figureCitation id="134DD1FAFFE0A86EFCDDFF34F805FA48" box="[821,877,1395,1419]" captionStart="Figure 4" captionStartId="9.[113,178,1764,1788]" captionTargetBox="[114,1458,144,1737]" captionTargetId="figure-6@9.[114,1458,144,1737]" captionTargetPageId="9" captionText="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." pageId="7" pageNumber="8">Fig. 4</figureCitation>
<figureCitation id="134DD1FAFFE0A86EFCDDFF34F805FA48" box="[821,877,1395,1419]" captionStart="Figure 4" captionStartId="9.[113,178,1764,1788]" captionTargetBox="[114,1458,144,1737]" captionTargetId="figure-6@9.[114,1458,144,1737]" captionTargetPageId="9" captionText="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." figureDoi="http://doi.org/10.5281/zenodo.14827326" httpUri="https://zenodo.org/record/14827326/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
), whereas
<taxonomicName id="4C76B6FCFFE0A86EFC33FF33FF23FA48" box="[987,1099,1395,1419]" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="schultzi">
<emphasis id="B902116DFFE0A86EFC33FF33FF23FA48" box="[987,1099,1395,1419]" italics="true" pageId="7" pageNumber="8">Bu. schultzi</emphasis>
@ -1548,7 +1551,7 @@ in apicobasal length and the smaller is
in mesiodistal width, whereas the smaller is
<quantity id="4C8E609AFFE0A86EFAB7FC68FED8F985" box="[1375,1456,1583,1607]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="8" unit="mm" value="1.5">1.5 mm</quantity>
. The teeth bear ~8 serrations per millimetre. The number of dentary teeth cannot be defined, as the bone is not complete (
<figureCitation id="134DD1FAFFE0A86EFA65FC29F85FF966" captionStart="Figure 4" captionStartId="9.[113,178,1764,1788]" captionTargetBox="[114,1458,144,1737]" captionTargetId="figure-6@9.[114,1458,144,1737]" captionTargetPageId="9" captionText="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." pageId="7" pageNumber="8">Fig. 4</figureCitation>
<figureCitation id="134DD1FAFFE0A86EFA65FC29F85FF966" captionStart="Figure 4" captionStartId="9.[113,178,1764,1788]" captionTargetBox="[114,1458,144,1737]" captionTargetId="figure-6@9.[114,1458,144,1737]" captionTargetPageId="9" captionText="Figure 4. Dentition of UFSM 11660A:(A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view.Abbreviations:idp, interdental plates." figureDoi="http://doi.org/10.5281/zenodo.14827326" httpUri="https://zenodo.org/record/14827326/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
). The apical parts of the rostral teeth are broken. The 5
<superScript id="7C036037FFE0A86EFA80FCCCFE1DF95A" attach="left" box="[1384,1397,1675,1689]" fontSize="6" pageId="7" pageNumber="8">th</superScript>
tooth has the mesial margin convex and the distal one almost straight in labial/lingual views.
@ -1557,7 +1560,7 @@ tooth has the mesial margin convex and the distal one almost straight in labial/
<emphasis id="B902116DFFE0A86EFBCFFCB9FFDEF8DB" bold="true" box="[1063,1206,1790,1816]" pageId="7" pageNumber="8">Axial remains</emphasis>
</paragraph>
<paragraph id="8BC9CD7FFFE0A861FCC2FD62FA74F90A" blockId="7.[810,1459,1829,1978]" lastBlockId="8.[129,779,1650,1737]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">The preserved axial elements of UFSM 11660 include at least three dorsal vertebrae, three sets of sacral vertebrae, and 18 caudal vertebrae, only some of them partially articulated. One of the sets of sacral vertebrae is quite damaged, hampering mechanical preparation and, consequently, reliable observations. Due to the impossibility to assign the vertebrae to specific individuals, they are described below simply according to their position on the axial series.</paragraph>
<caption id="DF099DF7FFEFA861FF69FFB5F92CF980" pageId="8" pageNumber="9" startId="8.[129,194,1522,1546]" targetBox="[129,1473,144,1495]" targetPageId="8" targetType="figure">
<caption id="DF099DF7FFEFA861FF69FFB5F92CF980" ID-DOI="http://doi.org/10.5281/zenodo.14827323" ID-Zenodo-Dep="14827323" httpUri="https://zenodo.org/record/14827323/files/figure.png" pageId="8" pageNumber="9" startId="8.[129,194,1522,1546]" targetBox="[129,1473,144,1495]" targetPageId="8" targetType="figure">
<paragraph id="8BC9CD7FFFEFA861FF69FFB5FE14F9C8" blockId="8.[129,1455,1522,1603]" box="[129,1404,1522,1547]" pageId="8" pageNumber="9">
<emphasis id="B902116DFFEFA861FF69FFB5FBB0F9C9" bold="true" box="[129,216,1522,1546]" pageId="8" pageNumber="9">Figure 3.</emphasis>
Premaxilla and maxilla of UFSM 11660A: (A, B) lateral and (C, D) medial views. Abbreviations: clp, caudolateral process; dp,
@ -1567,7 +1570,7 @@ Premaxilla and maxilla of UFSM 11660A: (A, B) lateral and (C, D) medial views. A
<paragraph id="8BC9CD7FFFEFA861FF69FCADFFF9F885" blockId="8.[129,778,1770,1951]" lastBlockId="8.[825,1476,1650,1862]" pageId="8" pageNumber="9">
<emphasis id="B902116DFFEFA861FF69FCADFA43F8C1" box="[129,299,1770,1794]" italics="true" pageId="8" pageNumber="9">Dorsal vertebrae:</emphasis>
UFSM 11660B includes three dorsal vertebrae, two of which in articulation (
<figureCitation id="134DD1FAFFEFA861FE0BFD4DF973F8E1" box="[483,539,1802,1826]" captionStart="Figure 6" captionStartId="11.[115,180,1533,1557]" captionTargetBox="[114,1458,144,1506]" captionTargetId="figure-210@11.[114,1458,144,1506]" captionTargetPageId="11" captionText="Figure 6.Dorsal vertebrae of UFSM 11660B: (A, D, E) Lateral, (B) cranial, and (C) caudal views.Abbreviations:c, centra; ded, dorsal extensor depression; dp, diapophysis; ncs, neurocentral; ns, neural spine; pa, parapophysis; ph, phalange; poz, postzygapophysis; prz, prezygapophysis." pageId="8" pageNumber="9">Fig. 6</figureCitation>
<figureCitation id="134DD1FAFFEFA861FE0BFD4DF973F8E1" box="[483,539,1802,1826]" captionStart="Figure 6" captionStartId="11.[115,180,1533,1557]" captionTargetBox="[114,1458,144,1506]" captionTargetId="figure-210@11.[114,1458,144,1506]" captionTargetPageId="11" captionText="Figure 6.Dorsal vertebrae of UFSM 11660B: (A, D, E) Lateral, (B) cranial, and (C) caudal views.Abbreviations:c, centra; ded, dorsal extensor depression; dp, diapophysis; ncs, neurocentral; ns, neural spine; pa, parapophysis; ph, phalange; poz, postzygapophysis; prz, prezygapophysis." figureDoi="http://doi.org/10.5281/zenodo.14827332" httpUri="https://zenodo.org/record/14827332/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
). The centra are longer (
<quantity id="4C8E609AFFEFA861FF61FD6EFBB3F882" box="[137,219,1833,1857]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="9" unit="mm" value="2.3">2.3 mm</quantity>
) than tall (
@ -1595,16 +1598,16 @@ Langer
<paragraph id="8BC9CD7FFFEFA863FCD1FD2FFA3DF9AA" blockId="8.[825,1474,1896,1951]" lastBlockId="10.[128,780,1429,1986]" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<emphasis id="B902116DFFEFA861FCD1FD2FF8B5F843" box="[825,989,1896,1920]" italics="true" pageId="8" pageNumber="9">Sacral vertebrae:</emphasis>
The two best-preserved sets are composed of two articulated primordial sacral vertebrae. UFSM 11660C is articulated with the ilium and UFSM 11660D is isolated (
<figureCitation id="134DD1FAFFEDA863FD2AFFD2F991FA6E" box="[706,761,1429,1453]" captionStart="Figure 7" captionStartId="12.[130,195,1276,1300]" captionTargetBox="[129,1473,144,1248]" captionTargetId="figure-397@12.[129,1473,144,1248]" captionTargetPageId="12" captionText="Figure 7. Sacral vertebrae of UFSM 11660C (A, B) and UFSM 11660D (CF): (A, C) dorsal, (B, D) ventral, (E) caudal, and (F) lateral views.Abbreviations:ns, neural spine; prz, prezygapophysis; sr, sacral rib; tp, transverse process; 1, first primordial sacral vertebra; 2, second primordial sacral vertebra." pageId="10" pageNumber="11">Fig. 7</figureCitation>
<figureCitation id="134DD1FAFFEDA863FD2AFFD2F991FA6E" box="[706,761,1429,1453]" captionStart="Figure 7" captionStartId="12.[130,195,1276,1300]" captionTargetBox="[129,1473,144,1248]" captionTargetId="figure-397@12.[129,1473,144,1248]" captionTargetPageId="12" captionText="Figure 7. Sacral vertebrae of UFSM 11660C (A, B) and UFSM 11660D (CF): (A, C) dorsal, (B, D) ventral, (E) caudal, and (F) lateral views.Abbreviations:ns, neural spine; prz, prezygapophysis; sr, sacral rib; tp, transverse process; 1, first primordial sacral vertebra; 2, second primordial sacral vertebra." figureDoi="http://doi.org/10.5281/zenodo.14827336" httpUri="https://zenodo.org/record/14827336/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
). The centra of UFSM 11660D are slightly wider lateromedially and shallower dorsoventrally than those of UFSM 11660C. In UFSM 11660C, the S1 cranial articular surface is concave. In both sets, the S2 caudal articular surface is concave and almost as wide as deep. The sacral centra are not co-osified in both sets and lack ventral keel.
</paragraph>
<caption id="DF099DF7FFEEA860FF99FCA3F817F8F7" pageId="9" pageNumber="10" startId="9.[113,178,1764,1788]" targetBox="[114,1458,144,1737]" targetPageId="9" targetType="figure">
<caption id="DF099DF7FFEEA860FF99FCA3F817F8F7" ID-DOI="http://doi.org/10.5281/zenodo.14827326" ID-Zenodo-Dep="14827326" httpUri="https://zenodo.org/record/14827326/files/figure.png" pageId="9" pageNumber="10" startId="9.[113,178,1764,1788]" targetBox="[114,1458,144,1737]" targetPageId="9" targetType="figure">
<paragraph id="8BC9CD7FFFEEA860FF99FCA3F817F8F7" blockId="9.[113,1456,1764,1845]" pageId="9" pageNumber="10">
<emphasis id="B902116DFFEEA860FF99FCA3FBA1F93F" bold="true" box="[113,201,1764,1788]" pageId="9" pageNumber="10">Figure 4.</emphasis>
Dentition of UFSM 11660A: (A) overview of the left maxilla in lateral view; (B, C, D) maxillary teeth in labial view, (E) right maxilla in lateral view, (F) anterior maxillary teeth in labial view, (G) magnification of the anterior maxillary teeth, (H) interdental plates of the dentary, and (I) right dentary in medial view. Abbreviations: idp, interdental plates.
</paragraph>
</caption>
<caption id="DF099DF7FFEDA863FF69FF52FAA2FAA6" pageId="10" pageNumber="11" startId="10.[129,194,1301,1325]" targetBox="[129,1473,144,1273]" targetPageId="10" targetType="figure">
<caption id="DF099DF7FFEDA863FF69FF52FAA2FAA6" ID-DOI="http://doi.org/10.5281/zenodo.14827328" ID-Zenodo-Dep="14827328" httpUri="https://zenodo.org/record/14827328/files/figure.png" pageId="10" pageNumber="11" startId="10.[129,194,1301,1325]" targetBox="[129,1473,144,1273]" targetPageId="10" targetType="figure">
<paragraph id="8BC9CD7FFFEDA863FF69FF52FEF0FAEE" blockId="10.[129,1432,1301,1381]" box="[129,1432,1301,1326]" pageId="10" pageNumber="11">
<emphasis id="B902116DFFEDA863FF69FF52FBB0FAED" bold="true" box="[129,216,1301,1326]" pageId="10" pageNumber="11">Figure 5.</emphasis>
Braincase elements of UFSM 11660A: (A) cranial, (B) dorsal, (C) ventral, and (D) caudal views. Abbreviations: bt, basal tubera;
@ -1615,10 +1618,10 @@ Braincase elements of UFSM 11660A: (A) cranial, (B) dorsal, (C) ventral, and (D)
<paragraph id="8BC9CD7FFFEDA863FCD1FFD2FE17F801" blockId="10.[825,1477,1429,1986]" pageId="10" pageNumber="11">
<emphasis id="B902116DFFEDA863FCD1FFD2F8D6FA6E" box="[825,958,1429,1453]" italics="true" pageId="10" pageNumber="11">Caudal series:</emphasis>
The caudal series is composed of 18 vertebrae (
<figureCitation id="134DD1FAFFEDA863FA75FFD2F82EFA0E" captionStart="Figure 8" captionStartId="13.[113,178,1836,1860]" captionTargetBox="[170,1403,144,1808]" captionTargetId="figure-6@13.[170,1403,144,1808]" captionTargetPageId="13" captionText="Figure 8. Caudal vertebrae of UFSM 11660F (AK), UFSM 11660G (LM), and UFSM 11660H (NP): (AB, GP) lateral, (C) cranial, (D) caudal, (E) dorsal, and (F) ventral views.Abbreviations:ch, chevron; ns, neural spine; poz, postzygapophysis; prz, prezygapophysis; tp, transverse process." pageId="10" pageNumber="11">Fig. 8</figureCitation>
<figureCitation id="134DD1FAFFEDA863FA75FFD2F82EFA0E" captionStart="Figure 8" captionStartId="13.[113,178,1836,1860]" captionTargetBox="[170,1403,144,1808]" captionTargetId="figure-6@13.[170,1403,144,1808]" captionTargetPageId="13" captionText="Figure 8. Caudal vertebrae of UFSM 11660F (AK), UFSM 11660G (LM), and UFSM 11660H (NP): (AB, GP) lateral, (C) cranial, (D) caudal, (E) dorsal, and (F) ventral views.Abbreviations:ch, chevron; ns, neural spine; poz, postzygapophysis; prz, prezygapophysis; tp, transverse process." figureDoi="http://doi.org/10.5281/zenodo.14827340" httpUri="https://zenodo.org/record/14827340/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
), including an articulated set of eight mid-caudal vertebrae with chevrons (UFSM 11660G), and ten disarticulated elements, six located proximal (UFSM 11660F) and four distal (UFSM 11660H) to the articulates segment. The centra become longer in the direction to the distal portion of the tail. None of the caudal centra has a ventral keel and both their articular facets are concave. The six proximalmost vertebrae (UFSM 11660F) are axially shortened, with their transverse processes dorsolaterally oriented. The prezygapophysis projects craniodorsally, whereas the postzigapophysis projects caudoventrally. The eight articulated vertebrae (UFSM 11660G) have centra longer than deep, the ventral surface of which lacks a keel. The transverse process is thin and the neural spine is low. The prezygapophysis is craniodorsally inclined, with a rounded and elongated articular facet. The postzygapophysis are rounded. The distalmost vertebrae (UFSM 11660H) are very elongated. These elements are proportionally smaller and do not preserve neural arches.
</paragraph>
<caption id="DF099DF7FFECA862FF9BFFBAFA7CF98E" pageId="11" pageNumber="12" startId="11.[115,180,1533,1557]" targetBox="[114,1458,144,1506]" targetPageId="11" targetType="figure">
<caption id="DF099DF7FFECA862FF9BFFBAFA7CF98E" ID-DOI="http://doi.org/10.5281/zenodo.14827332" ID-Zenodo-Dep="14827332" httpUri="https://zenodo.org/record/14827332/files/figure.png" pageId="11" pageNumber="12" startId="11.[115,180,1533,1557]" targetBox="[114,1458,144,1506]" targetPageId="11" targetType="figure">
<paragraph id="8BC9CD7FFFECA862FF9BFFBAFA7CF98E" blockId="11.[113,1376,1533,1613]" pageId="11" pageNumber="12">
<emphasis id="B902116DFFECA862FF9BFFBAFBA3F9D6" bold="true" box="[115,203,1533,1557]" pageId="11" pageNumber="12">Figure 6.</emphasis>
Dorsal vertebrae of UFSM 11660B: (A, D, E) Lateral, (B) cranial, and (C) caudal views. Abbreviations: c, centra; ded, dorsal extensor depression; dp, diapophysis; ncs, neurocentral; ns, neural spine; pa, parapophysis; ph, phalange; poz, postzygapophysis; prz, prezygapophysis.
@ -1630,7 +1633,7 @@ Dorsal vertebrae of UFSM 11660B: (A, D, E) Lateral, (B) cranial, and (C) caudal
<paragraph id="8BC9CD7FFFECA862FF99FCF1F9D7F8EF" blockId="11.[113,762,1718,1993]" pageId="11" pageNumber="12">
<emphasis id="B902116DFFECA862FF99FCF1FBC6F90D" box="[113,174,1718,1742]" italics="true" pageId="11" pageNumber="12">Ilium:</emphasis>
One partial ilium is preserved with sacral vertebrae in the association UFSM 11660C (
<figureCitation id="134DD1FAFFECA862FE2FFC91F969F92D" box="[455,513,1750,1774]" captionStart="Figure 9" captionStartId="14.[129,194,691,715]" captionTargetBox="[129,1473,144,663]" captionTargetId="figure-58@14.[129,1473,144,663]" captionTargetPageId="14" captionText="Figure 9. Ilium of UFSM 11660C: (A) lateral, (B) dorsal, and (C) ventral views. Abbreviations: av, attached vertebra; bf, fossa brevis; bs, brevis shelf; fteo, flexor tibialis externus origin; ip, ischiatic peduncle; poa, postacetabular ala; pp, pubic peduncle; prz, prezygapophysis; S1, first primordial sacral vertebra 1; S2, second primordial sacral vertebra; sac, supra-acetabular crest; tp, transverse process." pageId="11" pageNumber="12">Fig. 9</figureCitation>
<figureCitation id="134DD1FAFFECA862FE2FFC91F969F92D" box="[455,513,1750,1774]" captionStart="Figure 9" captionStartId="14.[129,194,691,715]" captionTargetBox="[129,1473,144,663]" captionTargetId="figure-58@14.[129,1473,144,663]" captionTargetPageId="14" captionText="Figure 9. Ilium of UFSM 11660C: (A) lateral, (B) dorsal, and (C) ventral views. Abbreviations: av, attached vertebra; bf, fossa brevis; bs, brevis shelf; fteo, flexor tibialis externus origin; ip, ischiatic peduncle; poa, postacetabular ala; pp, pubic peduncle; prz, prezygapophysis; S1, first primordial sacral vertebra 1; S2, second primordial sacral vertebra; sac, supra-acetabular crest; tp, transverse process." figureDoi="http://doi.org/10.5281/zenodo.14827342" httpUri="https://zenodo.org/record/14827342/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
). It is almost complete, lacking the preacetabular ala and part of the iliac blade. A disarticulated caudal vertebra is attached to the acetabular wall.
</paragraph>
<paragraph id="8BC9CD7FFFECA862FF65FD73FE73F8B2" blockId="11.[113,762,1718,1993]" lastBlockId="11.[810,1461,1661,1905]" pageId="11" pageNumber="12">
@ -1641,14 +1644,14 @@ The postacetabular ala is elongated, unlike that of herrerasaurids. There is a c
<paragraph id="8BC9CD7FFFECA865FCC2FDD5FAECF993" blockId="11.[810,1459,1938,1994]" lastBlockId="12.[128,778,1404,1616]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B902116DFFECA862FCC2FDD5F80FF869" box="[810,871,1938,1962]" italics="true" pageId="11" pageNumber="12">Pubis:</emphasis>
The pair of partially articulated pubis (UFSM 11660C;
<figureCitation id="134DD1FAFFECA862FCC4FDF6F819F809" box="[812,881,1969,1994]" captionStart="Figure 10" captionStartId="14.[129,194,1765,1789]" captionTargetBox="[129,1473,818,1736]" captionTargetId="figure-116@14.[129,1473,818,1736]" captionTargetPageId="14" captionText="Figure 10. Pubis of UFSM 11660C: (A) cranial, (B) lateral, (C) caudal, and (D) distal views.Abbreviations: ap, ambiens process; ed, expanded distal; ml, medial lamina; r, rib; s, striated surface." pageId="11" pageNumber="12">Fig. 10</figureCitation>
<figureCitation id="134DD1FAFFECA862FCC4FDF6F819F809" box="[812,881,1969,1994]" captionStart="Figure 10" captionStartId="14.[129,194,1765,1789]" captionTargetBox="[129,1473,818,1736]" captionTargetId="figure-116@14.[129,1473,818,1736]" captionTargetPageId="14" captionText="Figure 10. Pubis of UFSM 11660C: (A) cranial, (B) lateral, (C) caudal, and (D) distal views.Abbreviations: ap, ambiens process; ed, expanded distal; ml, medial lamina; r, rib; s, striated surface." figureDoi="http://doi.org/10.5281/zenodo.14827344" httpUri="https://zenodo.org/record/14827344/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
) are assigned to a single individual. The most complete left bone is
<quantity id="4C8E609AFFEBA865FF1BFF3BFA0FFA57" box="[243,359,1404,1428]" metricMagnitude="-1" metricUnit="m" metricValue="1.1738" pageId="12" pageNumber="13" unit="mm" value="117.38">117.38 mm</quantity>
long, with the main axis caudodorsally to cranioventrally oriented. The proximal outline of the iliac peduncle is rounded and the ambiens process forms a raised protuberance on the lateral surface of the main body of the bone. The shaft encompasses a thin medial lamina and its distal end expands dorsoventrally, but not as much as in herrerasaurids or theropods (
<bibRefCitation id="EFE7B08EFFEBA865FF11FC7EFA1BF993" author="Novas FE" box="[249,371,1592,1616]" pageId="12" pageNumber="13" pagination="400 - 23" refId="ref15821" refString="Novas FE. New information on the systematics and postcranial skeleton of Herrerasaurus ischigualastensis (Theropoda: Herrerasauridae) from the Ischigualasto Formation (Upper Triassic) of Argentina. Journal of Vertebrate Paleontology 1994; 13: 400 - 23. https: // doi. org / 10.1080 / 0 2724634.1994. 10011523" type="journal article" year="1994">Novas 1994</bibRefCitation>
).
</paragraph>
<caption id="DF099DF7FFEBA865FF6AFEBBFA11FA8F" pageId="12" pageNumber="13" startId="12.[130,195,1276,1300]" targetBox="[129,1473,144,1248]" targetPageId="12" targetType="figure">
<caption id="DF099DF7FFEBA865FF6AFEBBFA11FA8F" ID-DOI="http://doi.org/10.5281/zenodo.14827336" ID-Zenodo-Dep="14827336" httpUri="https://zenodo.org/record/14827336/files/figure.png" pageId="12" pageNumber="13" startId="12.[130,195,1276,1300]" targetBox="[129,1473,144,1248]" targetPageId="12" targetType="figure">
<paragraph id="8BC9CD7FFFEBA865FF6AFEBBFA11FA8F" blockId="12.[129,1444,1276,1356]" pageId="12" pageNumber="13">
<emphasis id="B902116DFFEBA865FF6AFEBBFBB1FAD7" bold="true" box="[130,217,1276,1300]" pageId="12" pageNumber="13">Figure 7.</emphasis>
Sacral vertebrae of UFSM 11660C (A, B) and UFSM 11660D (CF): (A, C) dorsal, (B, D) ventral, (E) caudal, and (F) lateral views. Abbreviations: ns, neural spine; prz, prezygapophysis; sr, sacral rib; tp, transverse process; 1, first primordial sacral vertebra; 2, second primordial sacral vertebra.
@ -1661,7 +1664,7 @@ A single left ischium (UFSM 11660C) was preserved. It is
.
<quantity id="4C8E609AFFEBA865FF2FFCD6FA74F96B" box="[199,284,1681,1705]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="mm" value="100.0">100 mm</quantity>
long and almost complete, lacking only part of the obturator plate. Its main body is subtriangular in lateral/ medial views and the shaft is elongated, expanding dorsoventrally at the distal end (
<figureCitation id="134DD1FAFFEBA865FE87FCA8FADDF8C4" box="[367,437,1775,1799]" captionStart="Figure 11" captionStartId="15.[113,178,1257,1281]" captionTargetBox="[114,1458,144,1230]" captionTargetId="figure-417@15.[114,1458,144,1230]" captionTargetPageId="15" captionText="Figure 11. Left ischium of UFSM 11660C: (A) lateral, (B) medial, (C) dorsal, (D) proximal, and (E) distal views. Abbreviations: at, antitrochanter; bp, bump; ls, lateral surface; ms, medial surface; obp, obturator plate area; rd, ridge." pageId="12" pageNumber="13">Fig. 11</figureCitation>
<figureCitation id="134DD1FAFFEBA865FE87FCA8FADDF8C4" box="[367,437,1775,1799]" captionStart="Figure 11" captionStartId="15.[113,178,1257,1281]" captionTargetBox="[114,1458,144,1230]" captionTargetId="figure-417@15.[114,1458,144,1230]" captionTargetPageId="15" captionText="Figure 11. Left ischium of UFSM 11660C: (A) lateral, (B) medial, (C) dorsal, (D) proximal, and (E) distal views. Abbreviations: at, antitrochanter; bp, bump; ls, lateral surface; ms, medial surface; obp, obturator plate area; rd, ridge." figureDoi="http://doi.org/10.5281/zenodo.14827346" httpUri="https://zenodo.org/record/14827346/files/figure.png" pageId="12" pageNumber="13">Fig. 11</figureCitation>
). The main proximal articulation (for the ilium and femur) has an ovoid outline, and the lateral (antitrochanteric) margin is convex, whereas the medial is straight to convex. The angle formed between the dorsal margin of the ischial body and the ischial shaft is more acute in UFSM 11660C,
<taxonomicName id="4C76B6FCFFEBA865FF37FDCBFA27F860" box="[223,335,1931,1955]" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="schultzi">
<emphasis id="B902116DFFEBA865FF37FDCBFA27F860" box="[223,335,1931,1955]" italics="true" pageId="12" pageNumber="13">Bu. schultzi</emphasis>
@ -1701,7 +1704,7 @@ The antitrochanter is well-expanded laterally, so that its transition to the mai
<paragraph id="8BC9CD7FFFEBA867FCD1FD48FBD4F879" blockId="12.[825,1474,1806,1987]" lastBlockId="14.[129,777,1891,1978]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<emphasis id="B902116DFFEBA865FCD1FD48F817F8E5" box="[825,895,1807,1830]" italics="true" pageId="12" pageNumber="13">Femur:</emphasis>
There are two right femora in the association (
<figureCitation id="134DD1FAFFEBA865FA82FD49FED8F8E5" box="[1386,1456,1806,1830]" captionStart="Figure 12" captionStartId="16.[129,194,1809,1833]" captionTargetBox="[129,1473,144,1781]" captionTargetId="figure-9@16.[129,1473,144,1781]" captionTargetPageId="16" captionText="Figure 12. Right femora of UFSM 11660I (AE) and UFSM 11660J (FK): (A, F) cranial, (B, G) medial, (C, H) caudal, (D, I, K) lateral, (E) distal, and (J) proximal views. Abbreviations: 4th, fourth trochanter; cmt, caudomedial tuber; crlt, craniolateral tuber; crmt, craniomedial tuber; cil, cranial intermuscular line; clil, caudolateral intermuscular line; cls, craniolateral scar; cmc, craniomedial crest; ct, cranial trochanter; ctf, crista tibiofibularis; dlt, dorsolateral trochanter; lc, lateral condyle; mc, medial condyle; pf, popliteal fossa; ts, trochanteric shelf." pageId="12" pageNumber="13">Fig. 12</figureCitation>
<figureCitation id="134DD1FAFFEBA865FA82FD49FED8F8E5" box="[1386,1456,1806,1830]" captionStart="Figure 12" captionStartId="16.[129,194,1809,1833]" captionTargetBox="[129,1473,144,1781]" captionTargetId="figure-9@16.[129,1473,144,1781]" captionTargetPageId="16" captionText="Figure 12. Right femora of UFSM 11660I (AE) and UFSM 11660J (FK): (A, F) cranial, (B, G) medial, (C, H) caudal, (D, I, K) lateral, (E) distal, and (J) proximal views. Abbreviations: 4th, fourth trochanter; cmt, caudomedial tuber; crlt, craniolateral tuber; crmt, craniomedial tuber; cil, cranial intermuscular line; clil, caudolateral intermuscular line; cls, craniolateral scar; cmc, craniomedial crest; ct, cranial trochanter; ctf, crista tibiofibularis; dlt, dorsolateral trochanter; lc, lateral condyle; mc, medial condyle; pf, popliteal fossa; ts, trochanteric shelf." figureDoi="http://doi.org/10.5281/zenodo.14827348" httpUri="https://zenodo.org/record/14827348/files/figure.png" pageId="12" pageNumber="13">Fig. 12</figureCitation>
): one almost complete (UFSM 11660I),
<quantity id="4C8E609AFFEBA865FB3DFD6AFE42F886" box="[1237,1322,1837,1861]" metricMagnitude="-1" metricUnit="m" metricValue="1.5699999999999998" pageId="12" pageNumber="13" unit="mm" value="157.0">157 mm</quantity>
long, and one preserving only the proximal part (UFSM 11660J),
@ -1709,22 +1712,22 @@ long, and one preserving only the proximal part (UFSM 11660J),
.
<quantity id="4C8E609AFFEBA865FA92FD0AFEAAF8A7" box="[1402,1474,1869,1893]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="13" unit="mm" value="55.0">55 mm</quantity>
long as preserved. UFSM 11660I is sigmoid in cranial/caudal and lateral/medial views. Its proximal portion is poorly preserved. The craniomedial tuber is reduced and the craniolateral is connected to a well-developed craniomedial crest (cmc in
<figureCitation id="134DD1FAFFE9A867FF69FDC4FBAFF858" box="[129,199,1923,1947]" captionStart="Figure 12" captionStartId="16.[129,194,1809,1833]" captionTargetBox="[129,1473,144,1781]" captionTargetId="figure-9@16.[129,1473,144,1781]" captionTargetPageId="16" captionText="Figure 12. Right femora of UFSM 11660I (AE) and UFSM 11660J (FK): (A, F) cranial, (B, G) medial, (C, H) caudal, (D, I, K) lateral, (E) distal, and (J) proximal views. Abbreviations: 4th, fourth trochanter; cmt, caudomedial tuber; crlt, craniolateral tuber; crmt, craniomedial tuber; cil, cranial intermuscular line; clil, caudolateral intermuscular line; cls, craniolateral scar; cmc, craniomedial crest; ct, cranial trochanter; ctf, crista tibiofibularis; dlt, dorsolateral trochanter; lc, lateral condyle; mc, medial condyle; pf, popliteal fossa; ts, trochanteric shelf." pageId="14" pageNumber="15">Fig. 12</figureCitation>
<figureCitation id="134DD1FAFFE9A867FF69FDC4FBAFF858" box="[129,199,1923,1947]" captionStart="Figure 12" captionStartId="16.[129,194,1809,1833]" captionTargetBox="[129,1473,144,1781]" captionTargetId="figure-9@16.[129,1473,144,1781]" captionTargetPageId="16" captionText="Figure 12. Right femora of UFSM 11660I (AE) and UFSM 11660J (FK): (A, F) cranial, (B, G) medial, (C, H) caudal, (D, I, K) lateral, (E) distal, and (J) proximal views. Abbreviations: 4th, fourth trochanter; cmt, caudomedial tuber; crlt, craniolateral tuber; crmt, craniomedial tuber; cil, cranial intermuscular line; clil, caudolateral intermuscular line; cls, craniolateral scar; cmc, craniomedial crest; ct, cranial trochanter; ctf, crista tibiofibularis; dlt, dorsolateral trochanter; lc, lateral condyle; mc, medial condyle; pf, popliteal fossa; ts, trochanteric shelf." figureDoi="http://doi.org/10.5281/zenodo.14827348" httpUri="https://zenodo.org/record/14827348/files/figure.png" pageId="14" pageNumber="15">Fig. 12</figureCitation>
) that extends distally along the proximal portion of the bone.
</paragraph>
<caption id="DF099DF7FFEAA864FF99FD6BFA4BF8B8" pageId="13" pageNumber="14" startId="13.[113,178,1836,1860]" targetBox="[170,1403,144,1808]" targetPageId="13" targetType="figure">
<caption id="DF099DF7FFEAA864FF99FD6BFA4BF8B8" ID-DOI="http://doi.org/10.5281/zenodo.14827340" ID-Zenodo-Dep="14827340" httpUri="https://zenodo.org/record/14827340/files/figure.png" pageId="13" pageNumber="14" startId="13.[113,178,1836,1860]" targetBox="[170,1403,144,1808]" targetPageId="13" targetType="figure">
<paragraph id="8BC9CD7FFFEAA864FF99FD6BFA4BF8B8" blockId="13.[113,1426,1836,1915]" pageId="13" pageNumber="14">
<emphasis id="B902116DFFEAA864FF99FD6BFBA1F887" bold="true" box="[113,201,1836,1860]" pageId="13" pageNumber="14">Figure 8.</emphasis>
Caudal vertebrae of UFSM 11660F (AK), UFSM 11660G (LM), and UFSM 11660H (NP): (AB, GP) lateral, (C) cranial, (D) caudal, (E) dorsal, and (F) ventral views. Abbreviations: ch, chevron; ns, neural spine; poz, postzygapophysis; prz, prezygapophysis; tp, transverse process.
</paragraph>
</caption>
<caption id="DF099DF7FFE9A867FF69F8F4FFADFCC1" pageId="14" pageNumber="15" startId="14.[129,194,691,715]" targetBox="[129,1473,144,663]" targetPageId="14" targetType="figure">
<caption id="DF099DF7FFE9A867FF69F8F4FFADFCC1" ID-DOI="http://doi.org/10.5281/zenodo.14827342" ID-Zenodo-Dep="14827342" httpUri="https://zenodo.org/record/14827342/files/figure.png" pageId="14" pageNumber="15" startId="14.[129,194,691,715]" targetBox="[129,1473,144,663]" targetPageId="14" targetType="figure">
<paragraph id="8BC9CD7FFFE9A867FF69F8F4FFADFCC1" blockId="14.[129,1474,691,771]" pageId="14" pageNumber="15">
<emphasis id="B902116DFFE9A867FF69F8F4FBB0FD08" bold="true" box="[129,216,691,715]" pageId="14" pageNumber="15">Figure 9.</emphasis>
Ilium of UFSM 11660C: (A) lateral, (B) dorsal, and (C) ventral views. Abbreviations: av, attached vertebra; bf, fossa brevis; bs, brevis shelf; fteo, flexor tibialis externus origin; ip, ischiatic peduncle; poa, postacetabular ala; pp, pubic peduncle; prz, prezygapophysis; S1, first primordial sacral vertebra 1; S2, second primordial sacral vertebra; sac, supra-acetabular crest; tp, transverse process.
</paragraph>
</caption>
<caption id="DF099DF7FFE9A867FF69FCA2F9DBF8DA" pageId="14" pageNumber="15" startId="14.[129,194,1765,1789]" targetBox="[129,1473,818,1736]" targetPageId="14" targetType="figure">
<caption id="DF099DF7FFE9A867FF69FCA2F9DBF8DA" ID-DOI="http://doi.org/10.5281/zenodo.14827344" ID-Zenodo-Dep="14827344" httpUri="https://zenodo.org/record/14827344/files/figure.png" pageId="14" pageNumber="15" startId="14.[129,194,1765,1789]" targetBox="[129,1473,818,1736]" targetPageId="14" targetType="figure">
<paragraph id="8BC9CD7FFFE9A867FF69FCA2F9DBF8DA" blockId="14.[129,1379,1765,1817]" pageId="14" pageNumber="15">
<emphasis id="B902116DFFE9A867FF69FCA2FB8DF93E" bold="true" box="[129,229,1765,1789]" pageId="14" pageNumber="15">Figure 10.</emphasis>
Pubis of UFSM 11660C: (A) cranial, (B) lateral, (C) caudal, and (D) distal views. Abbreviations: ap, ambiens process; ed, expanded distal; ml, medial lamina; r, rib; s, striated surface.
@ -1736,7 +1739,7 @@ The far better-preserved femoral head of UFSM 11660J has a transverse groove on
<bibRefCitation id="EFE7B08EFFE8A866FF07FF0AF948FAA5" author="Bittencourt JS &amp; Kellner AWA" box="[239,544,1357,1382]" pageId="15" pageNumber="16" pagination="1 - 56" refId="ref13142" refString="Bittencourt JS, Kellner AWA. The anatomy and phylogenetic position of the Triassic dinosaur Staurikosaurus pricei Colbert, 1970. Zootaxa 2009; 2079: 1 - 56." type="journal article" year="2009">Bittencourt and Kellner 2009</bibRefCitation>
), which extends distally along the cranial surface of the bone. The craniomedial tuber is the largest of the femoral head tubers and is separated from the caudomedial tuber by the sulcus for the capitis femoris ligament. Lateral to the caudomedial tuber, the facies articularis antitrochanterica slopes distally.
</paragraph>
<caption id="DF099DF7FFE8A866FF99FEAEFF78FADE" pageId="15" pageNumber="16" startId="15.[113,178,1257,1281]" targetBox="[114,1458,144,1230]" targetPageId="15" targetType="figure">
<caption id="DF099DF7FFE8A866FF99FEAEFF78FADE" ID-DOI="http://doi.org/10.5281/zenodo.14827346" ID-Zenodo-Dep="14827346" httpUri="https://zenodo.org/record/14827346/files/figure.png" pageId="15" pageNumber="16" startId="15.[113,178,1257,1281]" targetBox="[114,1458,144,1230]" targetPageId="15" targetType="figure">
<paragraph id="8BC9CD7FFFE8A866FF99FEAEFF78FADE" blockId="15.[113,1356,1257,1309]" pageId="15" pageNumber="16">
<emphasis id="B902116DFFE8A866FF99FEAEFBBEFAC2" bold="true" box="[113,214,1257,1281]" pageId="15" pageNumber="16">Figure 11.</emphasis>
Left ischium of UFSM 11660C: (A) lateral, (B) medial, (C) dorsal, (D) proximal, and (E) distal views. Abbreviations: at, antitrochanter; bp, bump; ls, lateral surface; ms, medial surface; obp, obturator plate area; rd, ridge.
@ -1789,7 +1792,7 @@ and some specimens of both
<collectingRegion id="49B2039DFFF6A878FF03FB4AFA28FEE6" box="[235,320,269,293]" country="South Africa" name="Western Cape" pageId="17" pageNumber="18">CAPPA</collectingRegion>
/UFSM 0035), the distal end of the femur is more mediolaterally expanded in UFSM 11660I, as in other specimens of the latter taxon (ULBRA-PVT 280).
</paragraph>
<caption id="DF099DF7FFF7A879FF69FD56FE38F8BE" pageId="16" pageNumber="17" startId="16.[129,194,1809,1833]" targetBox="[129,1473,144,1781]" targetPageId="16" targetType="figure">
<caption id="DF099DF7FFF7A879FF69FD56FE38F8BE" ID-DOI="http://doi.org/10.5281/zenodo.14827348" ID-Zenodo-Dep="14827348" httpUri="https://zenodo.org/record/14827348/files/figure.png" pageId="16" pageNumber="17" startId="16.[129,194,1809,1833]" targetBox="[129,1473,144,1781]" targetPageId="16" targetType="figure">
<paragraph id="8BC9CD7FFFF7A879FF69FD56FE38F8BE" blockId="16.[129,1459,1809,1917]" pageId="16" pageNumber="17">
<emphasis id="B902116DFFF7A879FF69FD56FB8DF8EA" bold="true" box="[129,229,1809,1833]" pageId="16" pageNumber="17">Figure 12.</emphasis>
Right femora of UFSM 11660I (AE) and UFSM 11660J (FK): (A, F) cranial, (B, G) medial, (C, H) caudal, (D, I, K) lateral, (E) distal, and (J) proximal views. Abbreviations: 4th, fourth trochanter; cmt, caudomedial tuber; crlt, craniolateral tuber; crmt, craniomedial tuber; cil, cranial intermuscular line; clil, caudolateral intermuscular line; cls, craniolateral scar; cmc, craniomedial crest; ct, cranial trochanter; ctf, crista tibiofibularis; dlt, dorsolateral trochanter; lc, lateral condyle; mc, medial condyle; pf, popliteal fossa; ts, trochanteric shelf.
@ -1807,7 +1810,7 @@ thick. Both caudolateral and cranial intermuscular lines are present and well de
<paragraph id="8BC9CD7FFFF6A878FCC2FAD7FFCDFE40" blockId="17.[810,1461,144,544]" pageId="17" pageNumber="18">
<emphasis id="B902116DFFF6A878FCC2FAD7F80FFF6B" box="[810,871,144,168]" italics="true" pageId="17" pageNumber="18">Tibia:</emphasis>
UFSM 11660 includes a pair of left/right partial tibiae with their proximal portions preserved, but it is not possible to assume that they belong to the same individual (
<figureCitation id="134DD1FAFFF6A878FAC3FA88FE1BFF24" box="[1323,1395,207,231]" captionStart="Figure 13" captionStartId="17.[113,178,1938,1962]" captionTargetBox="[114,1458,603,1910]" captionTargetId="figure-267@17.[114,1458,599,1910]" captionTargetPageId="17" captionText="Figure 13. Tibiae of UFSM 11660K (AC, G; left) and UFSM 11660L (DF; right): (A, D) lateral, (B, E) medial, (C) caudal, and (FG) proximal views.Abbreviations:cn, cnemial crest; fcr, fibular crest; lc, lateral condyle; mc, medial condyle; rs, rugose surface." pageId="17" pageNumber="18">Fig. 13</figureCitation>
<figureCitation id="134DD1FAFFF6A878FAC3FA88FE1BFF24" box="[1323,1395,207,231]" captionStart="Figure 13" captionStartId="17.[113,178,1938,1962]" captionTargetBox="[114,1458,603,1910]" captionTargetId="figure-267@17.[114,1458,599,1910]" captionTargetPageId="17" captionText="Figure 13. Tibiae of UFSM 11660K (AC, G; left) and UFSM 11660L (DF; right): (A, D) lateral, (B, E) medial, (C) caudal, and (FG) proximal views.Abbreviations:cn, cnemial crest; fcr, fibular crest; lc, lateral condyle; mc, medial condyle; rs, rugose surface." figureDoi="http://doi.org/10.5281/zenodo.14827350" httpUri="https://zenodo.org/record/14827350/files/figure.png" pageId="17" pageNumber="18">Fig. 13</figureCitation>
). The left bone (UFSM 11660K) is better preserved and
<quantity id="4C8E609AFFF6A878FAA3FAA9FEDBFEC6" box="[1355,1459,238,262]" metricMagnitude="-2" metricUnit="m" metricValue="7.194" pageId="17" pageNumber="18" unit="mm" value="71.94">71.94 mm</quantity>
long, whereas the right one (UFSM 11660L) is swollen and
@ -1841,7 +1844,7 @@ and other specimens of
</taxonomicName>
(ULBRA-PVT 280), the medial condyle lacks a caudal projection in UFSM 11660K. The condyles and the cnemial crest are poorly preserved in UFSM 11660L, but the latter is less arched than in the left tibia.
</paragraph>
<caption id="DF099DF7FFF6A878FF99FDD5FF94F805" pageId="17" pageNumber="18" startId="17.[113,178,1938,1962]" targetBox="[114,1458,603,1910]" targetPageId="17" targetType="figure">
<caption id="DF099DF7FFF6A878FF99FDD5FF94F805" ID-DOI="http://doi.org/10.5281/zenodo.14827350" ID-Zenodo-Dep="14827350" httpUri="https://zenodo.org/record/14827350/files/figure.png" pageId="17" pageNumber="18" startId="17.[113,178,1938,1962]" targetBox="[114,1458,603,1910]" targetPageId="17" targetType="figure">
<paragraph id="8BC9CD7FFFF6A878FF99FDD5FF94F805" blockId="17.[113,1408,1938,1991]" pageId="17" pageNumber="18">
<emphasis id="B902116DFFF6A878FF99FDD5FBBDF869" bold="true" box="[113,213,1938,1962]" pageId="17" pageNumber="18">Figure 13.</emphasis>
Tibiae of UFSM 11660K (AC, G; left) and UFSM 11660L (DF; right): (A, D) lateral, (B, E) medial, (C) caudal, and (FG) proximal views. Abbreviations: cn, cnemial crest; fcr, fibular crest; lc, lateral condyle; mc, medial condyle; rs, rugose surface.
@ -1875,7 +1878,7 @@ distally. A fibular crest extends caudally to its depression, as seen in
<paragraph id="8BC9CD7FFFF5A87BFF69F86FFF71FF25" blockId="18.[129,777,552,607]" lastBlockId="18.[825,1476,144,607]" pageId="18" pageNumber="19">
<emphasis id="B902116DFFF5A87BFF69F86FFAFCFD83" box="[129,404,552,576]" italics="true" pageId="18" pageNumber="19">Metatarsals and phalanges:</emphasis>
The preserved pedal parts include two sets of right metatarsals with similar morphology (
<figureCitation id="134DD1FAFFF5A87BFD5BF800F991FD9C" box="[691,761,583,607]" captionStart="Figure 14" captionStartId="18.[129,194,1868,1892]" captionTargetBox="[129,1473,681,1840]" captionTargetId="figure-308@18.[129,1473,681,1840]" captionTargetPageId="18" captionText="Figure 14. Right pes of UFSM 11660M (AD, GH) and UFSM 11660N (EF): (A, E) metatarsal III in proximal (top), anterior (middle), distal (bottom) views; (B, F) metatarsal II in proximal (top), anterior (middle), distal (bottom) views; (C) metatarsal I in lateral and (D) medial views; (GH) phalanx in lateral view. Abbreviations: a.mt, articulates with metatarsal; clp, collateral ligament; ed, extensor depressions; rdg, ridge." pageId="18" pageNumber="19">Fig. 14</figureCitation>
<figureCitation id="134DD1FAFFF5A87BFD5BF800F991FD9C" box="[691,761,583,607]" captionStart="Figure 14" captionStartId="18.[129,194,1868,1892]" captionTargetBox="[129,1473,681,1840]" captionTargetId="figure-308@18.[129,1473,681,1840]" captionTargetPageId="18" captionText="Figure 14. Right pes of UFSM 11660M (AD, GH) and UFSM 11660N (EF): (A, E) metatarsal III in proximal (top), anterior (middle), distal (bottom) views; (B, F) metatarsal II in proximal (top), anterior (middle), distal (bottom) views; (C) metatarsal I in lateral and (D) medial views; (GH) phalanx in lateral view. Abbreviations: a.mt, articulates with metatarsal; clp, collateral ligament; ed, extensor depressions; rdg, ridge." figureDoi="http://doi.org/10.5281/zenodo.14827352" httpUri="https://zenodo.org/record/14827352/files/figure.png" pageId="18" pageNumber="19">Fig. 14</figureCitation>
). UFSM 11660M is slightly smaller, comprising metatarsals I, II, and III. UFSM 11660N comprises longer metatarsals II (lacking the diaphysis) and III.
</paragraph>
<paragraph id="8BC9CD7FFFF5A87BFCBDFAA9FE38FE40" blockId="18.[825,1476,144,607]" pageId="18" pageNumber="19">
@ -1933,7 +1936,7 @@ Sereno
</bibRefCitation>
). The distal articulation has the medial condyle slightly displaced medially.
</paragraph>
<caption id="DF099DF7FFF5A87BFF69FD0BFB89F87B" pageId="18" pageNumber="19" startId="18.[129,194,1868,1892]" targetBox="[129,1473,681,1840]" targetPageId="18" targetType="figure">
<caption id="DF099DF7FFF5A87BFF69FD0BFB89F87B" ID-DOI="http://doi.org/10.5281/zenodo.14827352" ID-Zenodo-Dep="14827352" httpUri="https://zenodo.org/record/14827352/files/figure.png" pageId="18" pageNumber="19" startId="18.[129,194,1868,1892]" targetBox="[129,1473,681,1840]" targetPageId="18" targetType="figure">
<paragraph id="8BC9CD7FFFF5A87BFF69FD0BFB89F87B" blockId="18.[129,1469,1868,1976]" pageId="18" pageNumber="19">
<emphasis id="B902116DFFF5A87BFF69FD0BFB8DF8A7" bold="true" box="[129,229,1868,1892]" pageId="18" pageNumber="19">Figure 14.</emphasis>
Right pes of UFSM 11660M (AD, GH) and UFSM 11660N (EF): (A, E) metatarsal III in proximal (top), anterior (middle), distal (bottom) views; (B, F) metatarsal II in proximal (top), anterior (middle), distal (bottom) views; (C) metatarsal I in lateral and (D) medial views; (GH) phalanx in lateral view. Abbreviations: a.mt, articulates with metatarsal; clp, collateral ligament; ed, extensor depressions; rdg, ridge.
@ -1941,7 +1944,7 @@ Right pes of UFSM 11660M (AD, GH) and UFSM 11660N (EF): (A, E) metatars
</caption>
<paragraph id="8BC9CD7FFFF4A87AFF65FB6AFADCFDBD" blockId="19.[113,763,144,638]" pageId="19" pageNumber="20">
Some phalanges are preserved in the association (UFSM 11660M), but it is not possible to assign their position in the pes. A non-terminal phalanx was preserved articulated with an ungual (
<figureCitation id="134DD1FAFFF4A87AFF2EFBCCFA62FE60" box="[198,266,395,419]" captionStart="Figure 14" captionStartId="18.[129,194,1868,1892]" captionTargetBox="[129,1473,681,1840]" captionTargetId="figure-308@18.[129,1473,681,1840]" captionTargetPageId="18" captionText="Figure 14. Right pes of UFSM 11660M (AD, GH) and UFSM 11660N (EF): (A, E) metatarsal III in proximal (top), anterior (middle), distal (bottom) views; (B, F) metatarsal II in proximal (top), anterior (middle), distal (bottom) views; (C) metatarsal I in lateral and (D) medial views; (GH) phalanx in lateral view. Abbreviations: a.mt, articulates with metatarsal; clp, collateral ligament; ed, extensor depressions; rdg, ridge." pageId="19" pageNumber="20">Fig. 14</figureCitation>
<figureCitation id="134DD1FAFFF4A87AFF2EFBCCFA62FE60" box="[198,266,395,419]" captionStart="Figure 14" captionStartId="18.[129,194,1868,1892]" captionTargetBox="[129,1473,681,1840]" captionTargetId="figure-308@18.[129,1473,681,1840]" captionTargetPageId="18" captionText="Figure 14. Right pes of UFSM 11660M (AD, GH) and UFSM 11660N (EF): (A, E) metatarsal III in proximal (top), anterior (middle), distal (bottom) views; (B, F) metatarsal II in proximal (top), anterior (middle), distal (bottom) views; (C) metatarsal I in lateral and (D) medial views; (GH) phalanx in lateral view. Abbreviations: a.mt, articulates with metatarsal; clp, collateral ligament; ed, extensor depressions; rdg, ridge." figureDoi="http://doi.org/10.5281/zenodo.14827352" httpUri="https://zenodo.org/record/14827352/files/figure.png" pageId="19" pageNumber="20">Fig. 14</figureCitation>
). Their morphology resembles that of other early sauropodomorph phalanges. The non-terminal phalanx is longer than wide and presents a constriction at the mid-length. This condition differs from the stouter phalanx of sauropodiforms. Collateral ligament pits excavate the distal condyles. The ungual phalanx is ventrally curved and presents a conspicuous ridge on the lateral surface. It lacks a well-developed tuber on the ventral surface of the proximal portion.
</paragraph>
<paragraph id="8BC9CD7FFFF4A87AFE90F8F2FA9BFD0C" blockId="19.[376,499,693,719]" box="[376,499,693,719]" pageId="19" pageNumber="20">
@ -1966,7 +1969,7 @@ The analysis employing the data matrix ofNorman
<emphasis id="B902116DFFF4A87AFF77F9CFFACCFC63" box="[159,420,904,928]" italics="true" pageId="19" pageNumber="20">Nhandumirim waldsangae</emphasis>
</taxonomicName>
(
<figureCitation id="134DD1FAFFF4A87AFE5EF9CFFA95FC63" box="[438,509,904,928]" captionStart="Figure 15" captionStartId="19.[113,178,1954,1978]" captionTargetBox="[116,1450,1184,1923]" captionTargetId="figure-540@19.[114,1458,1182,1926]" captionTargetPageId="19" captionText="Figure 15. Abbreviated consensus tree of the first phylogenetic analysis." pageId="19" pageNumber="20">Fig. 15</figureCitation>
<figureCitation id="134DD1FAFFF4A87AFE5EF9CFFA95FC63" box="[438,509,904,928]" captionStart="Figure 15" captionStartId="19.[113,178,1954,1978]" captionTargetBox="[116,1450,1184,1923]" captionTargetId="figure-540@19.[114,1458,1182,1926]" captionTargetPageId="19" captionText="Figure 15. Abbreviated consensus tree of the first phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.14827354" httpUri="https://zenodo.org/record/14827354/files/figure.png" pageId="19" pageNumber="20">Fig. 15</figureCitation>
). The IterPCR (
<bibRefCitation id="EFE7B08EFFF4A87AFD43F9CFFB9CFC7C" author="Pol D &amp; Escapa H" pageId="19" pageNumber="20" pagination="515 - 27" refId="ref16105" refString="Pol D, Escapa H. Unstable taxa in cladistic analysis: identification and the assessment of relevant characters. Cladistics 2009; 25: 515 - 27." type="journal article" year="2009">Pol and Escapa 2009</bibRefCitation>
) method was used in order to detect any floating OTUs. The protocol identified
@ -1999,7 +2002,7 @@ Langer
<emphasis id="B902116DFFF4A87AFB24FAF7FE32FF04" box="[1228,1370,176,199]" italics="true" pageId="19" pageNumber="20">S. tupiniquim</emphasis>
</taxonomicName>
in both scenarios and the general topology was the same as recovered in the former study (
<figureCitation id="134DD1FAFFF4A87AFBE0FAA9FF38FEC5" box="[1032,1104,238,262]" captionStart="Figure 16" captionStartId="20.[129,194,1678,1702]" captionTargetBox="[130,1471,145,1648]" captionTargetId="figure-107@20.[129,1473,144,1650]" captionTargetPageId="20" captionText="Figure 16. Phylogenetic relations and morphospace position of UFSM 11660 among Carnian sauropodomorphs:(A) per specimen analysis; (B) combined analysis; (C) morphospace represented by PCoA1 (21.45%) and PCoA2 (16.71%). Sauropodomorph silhouette by Maurício S. Garcia." pageId="19" pageNumber="20">Fig. 16</figureCitation>
<figureCitation id="134DD1FAFFF4A87AFBE0FAA9FF38FEC5" box="[1032,1104,238,262]" captionStart="Figure 16" captionStartId="20.[129,194,1678,1702]" captionTargetBox="[130,1471,145,1648]" captionTargetId="figure-107@20.[129,1473,144,1650]" captionTargetPageId="20" captionText="Figure 16. Phylogenetic relations and morphospace position of UFSM 11660 among Carnian sauropodomorphs:(A) per specimen analysis; (B) combined analysis; (C) morphospace represented by PCoA1 (21.45%) and PCoA2 (16.71%). Sauropodomorph silhouette by Maurício S. Garcia." figureDoi="http://doi.org/10.5281/zenodo.14827356" httpUri="https://zenodo.org/record/14827356/files/figure.png" pageId="19" pageNumber="20">Fig. 16</figureCitation>
). In the per specimen analysis 35 MPTs of 1554 steps (CI = 0.449, RI = 0.449), with UFSM 11660 nested amid the type-series of
<taxonomicName id="4C76B6FCFFF4A87AFB5BFB6AFE5EFE87" baseAuthorityName="Langer" baseAuthorityYear="2003" box="[1203,1334,301,324]" class="Reptilia" genus="Saturnalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tupiniquim">
<emphasis id="B902116DFFF4A87AFB5BFB6AFE5EFE87" box="[1203,1334,301,324]" italics="true" pageId="19" pageNumber="20">S. tupiniquim</emphasis>
@ -2023,7 +2026,7 @@ Langer
andMoro
<emphasis id="B902116DFFF4A87AFBBEF84EFFEAFDE3" box="[1110,1154,520,544]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
(2024). PCoA1 corresponded to 21.45% of the total variance, whereas PCoA2 corresponded to 16.71% (
<figureCitation id="134DD1FAFFF4A87AFC4DF800F896FD9C" box="[933,1022,583,607]" captionStart="Figure 16" captionStartId="20.[129,194,1678,1702]" captionTargetBox="[130,1471,145,1648]" captionTargetId="figure-107@20.[129,1473,144,1650]" captionTargetPageId="20" captionText="Figure 16. Phylogenetic relations and morphospace position of UFSM 11660 among Carnian sauropodomorphs:(A) per specimen analysis; (B) combined analysis; (C) morphospace represented by PCoA1 (21.45%) and PCoA2 (16.71%). Sauropodomorph silhouette by Maurício S. Garcia." pageId="19" pageNumber="20">Fig. 16C</figureCitation>
<figureCitation id="134DD1FAFFF4A87AFC4DF800F896FD9C" box="[933,1022,583,607]" captionStart="Figure 16" captionStartId="20.[129,194,1678,1702]" captionTargetBox="[130,1471,145,1648]" captionTargetId="figure-107@20.[129,1473,144,1650]" captionTargetPageId="20" captionText="Figure 16. Phylogenetic relations and morphospace position of UFSM 11660 among Carnian sauropodomorphs:(A) per specimen analysis; (B) combined analysis; (C) morphospace represented by PCoA1 (21.45%) and PCoA2 (16.71%). Sauropodomorph silhouette by Maurício S. Garcia." figureDoi="http://doi.org/10.5281/zenodo.14827356" httpUri="https://zenodo.org/record/14827356/files/figure.png" pageId="19" pageNumber="20">Fig. 16C</figureCitation>
). The morphospaces of the hypodigms of
<taxonomicName id="4C76B6FCFFF4A87AFCC4F820F8F9FDBD" box="[812,913,614,638]" class="Reptilia" genus="Eoraptor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="lunensis">
<emphasis id="B902116DFFF4A87AFCC4F820F8F9FDBD" box="[812,913,614,638]" italics="true" pageId="19" pageNumber="20">E. lunensis</emphasis>

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@ -1,168 +1,190 @@
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<figureCitation id="13742FB414596171FEF22C9836A9C4F4" box="[327,434,1416,1442]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Figs 2AL</figureCitation>
<figureCitation id="13742FB414596171FEF22C9836A9C4F4" box="[327,434,1416,1442]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Figs 2AL</figureCitation>
,
<figureCitation id="13742FB414596171FE082C9836E0C4F4" box="[445,507,1416,1442]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. Fiopt inptao laova of Afootoopical Miltogoamminae. A, Hoplacephala pp., habitup, lateoal view. B, Hoplacephala pp., cephalopkeleton, lateoal view. C, Hoplacephala pp., cephalopkeleton, ventoal view. D, Lamprometopia concavatogena, habitup, lateoal view. E, L. concavatogena, cephalopkeleton, lateoal view. F, L.concavatogena, cephalopkeleton, ventoal view. G, Phylloteles picifons, habitup, lateoal view. H, P. picifons, cephalopkeleton, lateoal view. I, P. picifons, cephalopkeleton, ventoal view. Abboeviationp:a1a7, abdominal pegmentp; ad, anal divipion; ahp, additional hoopephoe-phaped pcleoite; dc, doopal coonu; ip, inteomediate pcleoite; la, lateoal aom; lb, laboum; lp, labial pcleoite; mh, mouthhook; pb, paoaptomal bao; pc, ppeudocephalon; t1t3, thooacic pegmentp; vc, ventoal coonu; vp, veotical plate." pageId="4">4DF</figureCitation>
<figureCitation id="13742FB414596171FE082C9836E0C4F4" box="[445,507,1416,1442]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. First instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., habitus, lateral view. B, Hoplacephala sp., cephaloskeleton, lateral view. C, Hoplacephala sp., cephaloskeleton, ventral view. D, Lamprometopia concavatogena, habitus, lateral view. E, L. concavatogena, cephaloskeleton, lateral view. F, L.concavatogena, cephaloskeleton, ventral view. G, Phylloteles picifons, habitus, lateral view. H, P. picifons, cephaloskeleton, lateral view. I, P. picifons, cephaloskeleton, ventral view. Abbreviations:a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate." figureDoi="http://doi.org/10.5281/zenodo.14827243" httpUri="https://zenodo.org/record/14827243/files/figure.png" pageId="4" pageNumber="5">4DF</figureCitation>
,
<figureCitation id="13742FB414596171FDB12C983559C4F4" box="[516,578,1416,1442]" captionStart="Figure 5" captionStartId="8.[129,194,1116,1140]" captionTargetBox="[129,1473,143,1088]" captionTargetId="figure-1280@8.[129,1473,143,1088]" captionTargetPageId="8" captionText="Figure 5. CLMS imagep of fiopt inptao laova of Afootoopical Miltogoamminae. A, Hoplacephala pp., anteoioo paot of cephalopkeleton, lateoal view. B, Hoplacephala pp., mid paot of cephalopkeleton, lateoal view. C, Lamprometopia concavatogena, cephalopkeleton, lateoal view. D, L. concavatogena, cephalopkeleton, doopo-lateoal view. E, Phylloteles picifons, cephalopkeleton, lateoal view. F, P.picifons, doopo-lateoal view. Abboeviationp:ahp, additional hoopephoe-phaped pcleoite; c/pbt, ciooi/pupoabuccal teeth; dc, doopal coonu; ep, epiptomal pcleoite; ip, inteomediate pcleoite; la, lateoal aom; lb, laboum; lp, labial pcleoite; mh, mouthhook; pb, paoaptomal bao; vc, ventoal coonu; vp, veotical plate." pageId="4">5C, D</figureCitation>
<figureCitation id="13742FB414596171FDB12C983559C4F4" box="[516,578,1416,1442]" captionStart="Figure 5" captionStartId="8.[129,194,1116,1140]" captionTargetBox="[129,1473,143,1088]" captionTargetId="figure-1280@8.[129,1473,143,1088]" captionTargetPageId="8" captionText="Figure 5. CLMS images of first instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., anterior part of cephaloskeleton, lateral view. B, Hoplacephala sp., mid part of cephaloskeleton, lateral view. C, Lamprometopia concavatogena, cephaloskeleton, lateral view. D, L. concavatogena, cephaloskeleton, dorso-lateral view. E, Phylloteles picifons, cephaloskeleton, lateral view. F, P.picifons, dorso-lateral view. Abbreviations:ahs, additional horseshoe-shaped sclerite; c/sbt, cirri/suprabuccal teeth; dc, dorsal cornu; es, epistomal sclerite; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; vc, ventral cornu; vp, vertical plate." figureDoi="http://doi.org/10.5281/zenodo.14827247" httpUri="https://zenodo.org/record/14827247/files/figure.png" pageId="4" pageNumber="5">5C, D</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BF0333114596171FF342CAC359BC764" blockId="4.[126,778,1468,1586]" pageId="4">
<emphasis id="B93BEF2314596171FF342CAC3652C482" box="[129,329,1468,1492]" italics="true" pageId="4">Material examined:</emphasis>
Eighteen laovae foom a pingle female:/ NamibiaWelw/Namib-NauklufNP/-23.54 126 15.29 55 9/
<date id="FFF115F114596171FD5A2CCB37FCC745" pageId="4" value="2018-11-22">22 Nov 2018</date>
/leg. KEiB Expedition//Lampo1/Namib.//
<emphasis id="B93BEF2314596171FD1D2CEB37A7C764" italics="true" pageId="4">Lamprom etopia</emphasis>
</subSubSection>
<subSubSection id="C35560BA14596171FF342CAC359BC764" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BF0333114596171FF342CAC359BC764" blockId="4.[126,778,1468,1586]" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FF342CAC3652C482" box="[129,329,1468,1492]" italics="true" pageId="4" pageNumber="5">Material examined:</emphasis>
<materialsCitation id="3B27396C14596171FEEC2CAC359BC764" determinerName="K. Szpila" location="Namib-NaukluftNP" pageId="4" pageNumber="5" specimenCount="19" specimenCount-female="1" specimenCount-larva="18">
<specimenCount id="9D49F8B814596171FEEC2CAC36E4C482" box="[345,511,1468,1492]" count="18" pageId="4" pageNumber="5" type="larva">Eighteen larvae</specimenCount>
from a single
<specimenCount id="9D49F8B814596171FD072CAC35EDC482" box="[690,758,1468,1492]" pageId="4" pageNumber="5" type="female">female</specimenCount>
:/ NamibiaWelw/
<location id="8E9065EA14596171FE9A2CCB36E6C4A5" LSID="urn:lsid:plazi:treatment:03E6822714596171FF572C493358C702:8E9065EA14596171FE9A2CCB36E6C4A5" box="[303,509,1499,1523]" name="Namib-NaukluftNP" pageId="4" pageNumber="5">Namib-NaukluftNP</location>
/-23.54 126 15.29
<date id="FFF115F114596171FD0C2CCB3412C4A5" box="[697,777,1499,1523]" pageId="4" pageNumber="5" value="1955-09-22">55 9/22</date>
Nov 2018/leg. KEiB Expedition//Lampr1/Namib.//
<emphasis id="B93BEF2314596171FD1D2CEB37A7C764" italics="true" pageId="4" pageNumber="5">Lamprom etopia</emphasis>
/
<taxonomicName id="4C4F48B214596171FF7D2F0B364FC764" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[200,340,1563,1586]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="4" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF2314596171FF7D2F0B364FC764" box="[200,340,1563,1586]" italics="true" pageId="4">concavatogena</emphasis>
<taxonomicName id="4C4F48B214596171FF7D2F0B364FC764" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[200,340,1563,1586]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF2314596171FF7D2F0B364FC764" box="[200,340,1563,1586]" italics="true" pageId="4" pageNumber="5">concavatogena</emphasis>
</taxonomicName>
/(
<bibRefCitation id="EFDE4EC014596171FEDD2F0A36C1C764" author="Pape T" box="[360,474,1562,1586]" pageId="4" pagination="1 - 6" refId="ref8326" refString="Pape T. Two new ppeciep of Hoplatainia Zumpt (Dipteoa: Saocophagidae). Durban Museum Novitates 1986; 14: 1 - 6." type="journal article" year="1986">Pape 1986</bibRefCitation>
)/det. K. Szpila/
<bibRefCitation id="EFDE4EC014596171FEDD2F0A36C1C764" author="Pape T" box="[360,474,1562,1586]" pageId="4" pageNumber="5" pagination="1 - 6" refId="ref8326" refString="Pape T. Two new ppeciep of Hoplatainia Zumpt (Dipteoa: Saocophagidae). Durban Museum Novitates 1986; 14: 1 - 6." type="journal article" year="1986">Pape 1986</bibRefCitation>
)/det.
<determinerName id="7E2CDFF514596171FDAF2F0A356EC764" box="[538,629,1562,1586]" pageId="4" pageNumber="5">K. Szpila</determinerName>
/
</materialsCitation>
</paragraph>
<paragraph id="8BF0333114596171FF342F4333A5C191" blockId="4.[128,778,1619,1988]" lastBlockId="4.[825,1474,144,199]" pageId="4">
<emphasis id="B93BEF2314596171FF342F433638C73D" box="[129,291,1619,1643]" italics="true" pageId="4">Pseudocephalon:</emphasis>
Antennal complex (an) laoge, antennal dome (and) oval, with oounded tip, antennal bapal oing (abo) high (
<figureCitation id="13742FB414596171FF3E2F8237E2C7FC" box="[139,249,1682,1706]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2AD</figureCitation>
); maxillaov palpup (mp) phaped ap a high, cvlindoical pootubeoance (
<figureCitation id="13742FB414596171FED72FA136CBC79F" box="[354,464,1713,1737]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2AC</figureCitation>
), fiopt penpillum bapiconicum (pb1) long and with plightlv pwollen tip, phifed awav foom centoal clupteo of penpilla towaod medio-doopal boodeo of palpup, additional penpilla (np1np2) laoge and both pituated doopallv to centoal clupteo of penpilla (
<figureCitation id="13742FB414596171FE322E3F36D6C611" box="[391,461,1839,1863]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2E</figureCitation>
); ventoal oogan (vo) on plightlv pwollen, flephv lobe (
<figureCitation id="13742FB414596171FEE22E5E36DAC630" box="[343,449,1870,1894]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2AC</figureCitation>
), edge of penpillao pit with a few conical pootubeoancep (
<figureCitation id="13742FB414596171FEC62E7D36A2C6D3" box="[371,441,1901,1925]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2B</figureCitation>
); lateoallv to the mouth opening a few (one to two) ciooi/pupoabuccal teeth (c/pbt) (Figp 2B, 5C); ooal oidgep (oo) well developed (
<figureCitation id="13742FB414596171FE782EBC352CC692" box="[461,567,1964,1988]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2AC</figureCitation>
); anteoioo puoface of ppeudocephalon with iooegulao cuticulao waotp, popteoioo puoface with fine cuticulao oidgep (
<figureCitation id="13742FB414596171FBF129BF33B6C191" box="[1092,1197,175,199]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2AC</figureCitation>
</subSubSection>
<subSubSection id="C35560BA14596171FF342F43336FC4CC" pageId="4" pageNumber="5" type="description">
<paragraph id="8BF0333114596171FF342F4333A5C191" blockId="4.[128,778,1619,1988]" lastBlockId="4.[825,1474,144,199]" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FF342F433638C73D" box="[129,291,1619,1643]" italics="true" pageId="4" pageNumber="5">Pseudocephalon:</emphasis>
Antennal complex (an) large, antennal dome (and) oval, with rounded tip, antennal basal ring (abr) high (
<figureCitation id="13742FB414596171FF3E2F8237E2C7FC" box="[139,249,1682,1706]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2AD</figureCitation>
); maxillary palpus (mp) shaped as a high, cylindrical protuberance (
<figureCitation id="13742FB414596171FED72FA136CBC79F" box="[354,464,1713,1737]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2AC</figureCitation>
), first sensillum basiconicum (sb1) long and with slightly swollen tip, shifted away from central cluster of sensilla toward medio-dorsal border of palpus, additional sensilla (ns1ns2) large and both situated dorsally to central cluster of sensilla (
<figureCitation id="13742FB414596171FE322E3F36D6C611" box="[391,461,1839,1863]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2E</figureCitation>
); ventral organ (vo) on slightly swollen, fleshy lobe (
<figureCitation id="13742FB414596171FEE22E5E36DAC630" box="[343,449,1870,1894]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2AC</figureCitation>
), edge of sensillar pit with a few conical protuberances (
<figureCitation id="13742FB414596171FEC62E7D36A2C6D3" box="[371,441,1901,1925]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2B</figureCitation>
); laterally to the mouth opening a few (one to two) cirri/suprabuccal teeth (c/sbt) (Figs 2B, 5C); oral ridges (or) well developed (
<figureCitation id="13742FB414596171FE782EBC352CC692" box="[461,567,1964,1988]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2AC</figureCitation>
); anterior surface of pseudocephalon with irregular cuticular warts, posterior surface with fine cuticular ridges (
<figureCitation id="13742FB414596171FBF129BF33B6C191" box="[1092,1197,175,199]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2AC</figureCitation>
).
</paragraph>
<paragraph id="8BF0333114596171FC8C29F932B9C34C" blockId="4.[824,1474,233,539]" pageId="4">
<emphasis id="B93BEF2314596171FC8C29F934C5C057" box="[825,990,233,257]" italics="true" pageId="4">Cephaloskeleton:</emphasis>
Laboum (lb) ptoaight but with anteoioopaot bent down, tip pointed (Figp 4E, 5C, D); mouthhook (mh) with lateoalaomat bape, almoptpaoallel-pided and veovplightlv upcuoved, pick-axe phaped with a pingle pointed tip dioected about 45°C to longitudinal axip of mouthhook (Figp 4E, 5C, D); inteomediate pcleoite (ip) plightlv below bape of laboum and paoaptomal baop (pb) in lateoal view (Figp 4E, 5C, D), in ventoal view inteomediate pcleoite longeo than wide (
<figureCitation id="13742FB414596171FBF528D4339EC08A" box="[1088,1157,452,476]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. Fiopt inptao laova of Afootoopical Miltogoamminae. A, Hoplacephala pp., habitup, lateoal view. B, Hoplacephala pp., cephalopkeleton, lateoal view. C, Hoplacephala pp., cephalopkeleton, ventoal view. D, Lamprometopia concavatogena, habitup, lateoal view. E, L. concavatogena, cephalopkeleton, lateoal view. F, L.concavatogena, cephalopkeleton, ventoal view. G, Phylloteles picifons, habitup, lateoal view. H, P. picifons, cephalopkeleton, lateoal view. I, P. picifons, cephalopkeleton, ventoal view. Abboeviationp:a1a7, abdominal pegmentp; ad, anal divipion; ahp, additional hoopephoe-phaped pcleoite; dc, doopal coonu; ip, inteomediate pcleoite; la, lateoal aom; lb, laboum; lp, labial pcleoite; mh, mouthhook; pb, paoaptomal bao; pc, ppeudocephalon; t1t3, thooacic pegmentp; vc, ventoal coonu; vp, veotical plate." pageId="4">Fig. 4F</figureCitation>
); paoaptomal bao long (Figp 4E, F, 5C, D); veotical plate (vp) plightlv booadeo than ventoal coonu (vc) and doopal coonu (Figp 4E, 5C, D); doopal boidge abpent.
<paragraph id="8BF0333114596171FC8C29F932B9C34C" blockId="4.[824,1474,233,539]" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FC8C29F934C5C057" box="[825,990,233,257]" italics="true" pageId="4" pageNumber="5">Cephaloskeleton:</emphasis>
Labrum (lb) straight but with anteriorpart bent down, tip pointed (Figs 4E, 5C, D); mouthhook (mh) with lateralarmat base, almostparallel-sided and veryslightly upcurved, pick-axe shaped with a single pointed tip directed about 45°C to longitudinal axis of mouthhook (Figs 4E, 5C, D); intermediate sclerite (is) slightly below base of labrum and parastomal bars (pb) in lateral view (Figs 4E, 5C, D), in ventral view intermediate sclerite longer than wide (
<figureCitation id="13742FB414596171FBF528D4339EC08A" box="[1088,1157,452,476]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. First instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., habitus, lateral view. B, Hoplacephala sp., cephaloskeleton, lateral view. C, Hoplacephala sp., cephaloskeleton, ventral view. D, Lamprometopia concavatogena, habitus, lateral view. E, L. concavatogena, cephaloskeleton, lateral view. F, L.concavatogena, cephaloskeleton, ventral view. G, Phylloteles picifons, habitus, lateral view. H, P. picifons, cephaloskeleton, lateral view. I, P. picifons, cephaloskeleton, ventral view. Abbreviations:a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate." figureDoi="http://doi.org/10.5281/zenodo.14827243" httpUri="https://zenodo.org/record/14827243/files/figure.png" pageId="4" pageNumber="5">Fig. 4F</figureCitation>
); parastomal bar long (Figs 4E, F, 5C, D); vertical plate (vp) slightly broader than ventral cornu (vc) and dorsal cornu (Figs 4E, 5C, D); dorsal bridge absent.
</paragraph>
<paragraph id="8BF0333114596171FC8C2B2D3258C218" blockId="4.[825,1475,572,847]" pageId="4">
<emphasis id="B93BEF2314596171FC8C2B2D34F4C302" box="[825,1007,573,596]" italics="true" pageId="4">Toracic segments:</emphasis>
Anteoioo ppinope bandp (apb) with foom 56 (doopal puofacep) to 1112 (ventoal puoface of fiopt pegment) oowp ofppinep, ppinep aooanged pepaoatelv foom each otheo (
<figureCitation id="13742FB414596171FAFF2B6B32B4C3C5" box="[1354,1455,635,659]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig.2AC</figureCitation>
); lateoal puoface of fiopt thooacic pegment with apeotuoe of anteoioo ppioacle (ap) (
<figureCitation id="13742FB414596171FC712BA93316C387" box="[964,1037,697,721]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2C</figureCitation>
); oemaining aoea of thooacic pegmentp with denpelv pet cuticulao oidgep (
<figureCitation id="13742FB414596171FBD32BC933CAC3A7" box="[1126,1233,729,753]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2AC</figureCitation>
), oidgep on doopal and doopo-lateoal puofacep foagmented, oidgep on ventoal and ventoo-lateoal puofacep continued along the entioe length of pegment (
<figureCitation id="13742FB414596171FA282A073494C218" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2AC, F</figureCitation>
); Kelinp oogan with phoot penpilla (
<figureCitation id="13742FB414596171FB472A273229C219" box="[1266,1330,823,847]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2I</figureCitation>
<paragraph id="8BF0333114596171FC8C2B2D3258C218" blockId="4.[825,1475,572,847]" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FC8C2B2D34F4C302" box="[825,1007,573,596]" italics="true" pageId="4" pageNumber="5">Toracic segments:</emphasis>
Anterior spinose bands (asb) with from 56 (dorsal surfaces) to 1112 (ventral surface of first segment) rows ofspines, spines arranged separately from each other (
<figureCitation id="13742FB414596171FAFF2B6B32B4C3C5" box="[1354,1455,635,659]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig.2AC</figureCitation>
); lateral surface of first thoracic segment with aperture of anterior spiracle (as) (
<figureCitation id="13742FB414596171FC712BA93316C387" box="[964,1037,697,721]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
); remaining area of thoracic segments with densely set cuticular ridges (
<figureCitation id="13742FB414596171FBD32BC933CAC3A7" box="[1126,1233,729,753]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2AC</figureCitation>
), ridges on dorsal and dorso-lateral surfaces fragmented, ridges on ventral and ventro-lateral surfaces continued along the entire length of segment (
<figureCitation id="13742FB414596171FA282A073494C218" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2AC, F</figureCitation>
); Kelins organ with short sensilla (
<figureCitation id="13742FB414596171FB472A273229C219" box="[1266,1330,823,847]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2I</figureCitation>
).
</paragraph>
<paragraph id="8BF0333114596171FC8C2A6032D9C535" blockId="4.[824,1475,880,1123]" pageId="4">
<emphasis id="B93BEF2314596171FC8C2A6034E4C2DE" box="[825,1023,880,904]" italics="true" pageId="4">Abdominal segments:</emphasis>
Spinep of ppinope bandp on doopal and lateoal puofacep goaduallv changing to foagmented cuticulao oidgep along each pegment (
<figureCitation id="13742FB414596171FC292ABF34E7C291" box="[924,1020,943,967]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2F, G</figureCitation>
), ventoal ppinep pmalleo than lateoal and doopal onep, ventoal puoface with continued cuticulao oidgep (
<figureCitation id="13742FB414596171FAF72ADE3297C2B0" box="[1346,1420,974,998]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2H</figureCitation>
); lateoal coeeping weltp (lcw) developed and denpelv coveoed bv ppinep/ oidgep; ventoal puoface of a1a7 with well-defined fupifoom coeeping weltp (vcw) at the anteoioo maogin of pegmentp and anteoioo pooleg (apo) poe-coevice pooleg (pop) and popteoioo poolegp (pop) (
<figureCitation id="13742FB414596171FADE2D5B32A9C535" box="[1387,1458,1099,1123]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig.2H</figureCitation>
<paragraph id="8BF0333114596171FC8C2A6032D9C535" blockId="4.[824,1475,880,1123]" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FC8C2A6034E4C2DE" box="[825,1023,880,904]" italics="true" pageId="4" pageNumber="5">Abdominal segments:</emphasis>
Spines of spinose bands on dorsal and lateral surfaces gradually changing to fragmented cuticular ridges along each segment (
<figureCitation id="13742FB414596171FC292ABF34E7C291" box="[924,1020,943,967]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2F, G</figureCitation>
), ventral spines smaller than lateral and dorsal ones, ventral surface with continued cuticular ridges (
<figureCitation id="13742FB414596171FAF72ADE3297C2B0" box="[1346,1420,974,998]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2H</figureCitation>
); lateral creeping welts (lcw) developed and densely covered by spines/ ridges; ventral surface of a1a7 with well-defined fusiform creeping welts (vcw) at the anterior margin of segments and anterior proleg (apr) pre-crevice proleg (prp) and posterior prolegs (pos) (
<figureCitation id="13742FB414596171FADE2D5B32A9C535" box="[1387,1458,1099,1123]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig.2H</figureCitation>
).
</paragraph>
<paragraph id="8BF0333114596171FC8C2D98336FC4CC" blockId="4.[824,1476,1160,1435]" pageId="4">
<emphasis id="B93BEF2314596171FC8C2D9834DAC5F6" box="[825,961,1160,1184]" italics="true" pageId="4">Anal division:</emphasis>
Spinep of ppinope bandp on doopal and lateoal puofacep goaduallv changing to foagmented cuticulao oidgep along each pegment, on ventoal puoface ppinep well defined, ventoal puoface popteoioo to the ppinep with continued cuticulao oidgep (
<figureCitation id="13742FB414596171FCF12C153492C44B" box="[836,905,1285,1309]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2L</figureCitation>
); ppioaculao field with plightlv pootouded papillae p1p7 and iooegulao oow of pmall ppinep (
<figureCitation id="13742FB414596171FB392C3533D7C46B" box="[1164,1228,1317,1341]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2J</figureCitation>
); popteoioo ppioacle (pp) with fouo peoiptigmatic tufp, each with five to peven boanchep (
<figureCitation id="13742FB414596171FCF12C73349CC42D" box="[836,903,1379,1403]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2J</figureCitation>
); anal papillae (ap) oounded (
<figureCitation id="13742FB414596171FB712C73323EC42D" box="[1220,1317,1379,1403]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2J, K</figureCitation>
); anal tuf (at) with peveoal ppinep (
<figureCitation id="13742FB414596171FBB22C93337FC4CC" box="[1031,1124,1411,1435]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. Fiopt inptao laova of Lamprometopia concavatogena. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup.F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg; ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 2J, K</figureCitation>
<paragraph id="8BF0333114596171FC8C2D98336FC4CC" blockId="4.[824,1476,1160,1435]" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FC8C2D9834DAC5F6" box="[825,961,1160,1184]" italics="true" pageId="4" pageNumber="5">Anal division:</emphasis>
Spines of spinose bands on dorsal and lateral surfaces gradually changing to fragmented cuticular ridges along each segment, on ventral surface spines well defined, ventral surface posterior to the spines with continued cuticular ridges (
<figureCitation id="13742FB414596171FCF12C153492C44B" box="[836,905,1285,1309]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2L</figureCitation>
); spiracular field with slightly protruded papillae p1p7 and irregular row of small spines (
<figureCitation id="13742FB414596171FB392C3533D7C46B" box="[1164,1228,1317,1341]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2J</figureCitation>
); posterior spiracle (ps) with four peristigmatic tufts, each with five to seven branches (
<figureCitation id="13742FB414596171FCF12C73349CC42D" box="[836,903,1379,1403]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2J</figureCitation>
); anal papillae (ap) rounded (
<figureCitation id="13742FB414596171FB712C73323EC42D" box="[1220,1317,1379,1403]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2J, K</figureCitation>
); anal tuft (at) with several spines (
<figureCitation id="13742FB414596171FBB22C93337FC4CC" box="[1031,1124,1411,1435]" captionStart="Figure 2" captionStartId="5.[113,178,1772,1796]" captionTargetBox="[160,1413,144,1744]" captionTargetId="figure-9@5.[160,1413,144,1744]" captionTargetPageId="5" captionText="Figure 2. First instar larva of Lamprometopia concavatogena. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus.F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="4" pageNumber="5">Fig. 2J, K</figureCitation>
).
</paragraph>
<paragraph id="8BF0333114596171FC8C2CAF3358C702" blockId="4.[825,1474,1471,1621]" pageId="4">
<emphasis id="B93BEF2314596171FC8C2CAF348CC481" box="[825,919,1471,1495]" italics="true" pageId="4">Remarks:</emphasis>
Goavid femalep weoe atoacted to damaged teomite commuteo tunnelp on a toee tounk and on the goound puoface below the toee (
<figureCitation id="13742FB414596171FC552CEE3350C740" box="[992,1099,1534,1558]" captionStart="Fig" captionStartId="10.[129,162,963,987]" captionTargetBox="[129,1473,143,935]" captionTargetId="figure-1476@10.[129,1473,143,935]" captionTargetPageId="10" captionText="Fig. 7. Habitatp and femalep of Miltogoamminae. A, the toee in Homeb canvon, wheoe goavid female of Lamprometopia concavatogena wap collected. B, teomitep on the toee tounk. C, female of L. concavatogena on the goound at the tounk. D, teomite nept tumulup with peoching female of Phylloteles picifons (pointed bv aooow). E, female of P.picifons at the top of tumulup. F, female P.picifons enteoing into the open coooidoo." pageId="4">Fig. 7AC</figureCitation>
). Some fiopt inptao laovae developed pucceppfullv to the thiod inptao bv feeding on foephlv killed blow fliep (
<taxonomicName id="4C4F48B214596171FCC72F2D3329C703" authorityName="" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[882,1074,1597,1621]" class="Insecta" family="Calliphoridae" genus="Chrysomya" kingdom="Animalia" order="Diptera" pageId="4" phylum="Arthropoda" rank="species" species="albiceps">
<emphasis id="B93BEF2314596171FCC72F2D3329C703" box="[882,1074,1597,1621]" italics="true" pageId="4">Chrysomya albiceps</emphasis>
</subSubSection>
<subSubSection id="C35560BA14596171FC8C2CAF3358C702" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BF0333114596171FC8C2CAF3358C702" blockId="4.[825,1474,1471,1621]" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FC8C2CAF348CC481" box="[825,919,1471,1495]" italics="true" pageId="4" pageNumber="5">Remarks:</emphasis>
Gravid females were attracted to damaged termite commuter tunnels on a tree trunk and on the ground surface below the tree (
<figureCitation id="13742FB414596171FC552CEE3350C740" box="[992,1099,1534,1558]" captionStart="Fig" captionStartId="10.[129,162,963,987]" captionTargetBox="[129,1473,143,935]" captionTargetId="figure-1476@10.[129,1473,143,935]" captionTargetPageId="10" captionText="Fig. 7. Habitats and females of Miltogramminae. A, the tree in Homeb canyon, where gravid female of Lamprometopia concavatogena was collected. B, termites on the tree trunk. C, female of L. concavatogena on the ground at the trunk. D, termite nest tumulus with perching female of Phylloteles picifons (pointed by arrow). E, female of P.picifons at the top of tumulus. F, female P.picifons entering into the open corridor." figureDoi="http://doi.org/10.5281/zenodo.14827253" httpUri="https://zenodo.org/record/14827253/files/figure.png" pageId="4" pageNumber="5">Fig. 7AC</figureCitation>
). Some first instar larvae developed successfully to the third instar by feeding on freshly killed blow flies (
<taxonomicName id="4C4F48B214596171FCC72F2D3329C703" authorityName="" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[882,1074,1597,1621]" class="Insecta" family="Calliphoridae" genus="Chrysomya" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="albiceps">
<emphasis id="B93BEF2314596171FCC72F2D3329C703" box="[882,1074,1597,1621]" italics="true" pageId="4" pageNumber="5">Chrysomya albiceps</emphasis>
</taxonomicName>
).
</paragraph>

View file

@ -1,63 +1,66 @@
<document id="F2F131E647E662023A224AC59AE1B2C1" ID-DOI="10.1093/zoolinnean/zlae030" ID-ISSN="0024-4082" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1738154820953" checkinUser="plazi" docAuthor="Szpila, Kozvpztof, Walczak, Kinga, Sopzvńpka, Agniepzka &amp; Pape, Tomap" docDate="2025" docId="03E682271459617CFC542F643369C190" docLanguage="en" docName="zlae030.pdf" docOrigin="Zoological Journal of the Linnean Society 203" docSource="https://doi.org/10.1093/zoolinnean/zlae030" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Phylloteles picifons Pape 1985" docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="FFDFFA5F145D6175FFB52910371BC156" masterDocTitle="Fiopt data on eaolv inptao Afootoopical Miltogoamminae (Dipteoa: Saocophagidae) oeveal unexpected moophological diveopitv" masterLastPageNumber="13" masterPageNumber="1" pageNumber="5" updateTime="1738866455774" updateUser="jonas">
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<mods:title id="547A3070C970C3AA9CA086AAA936BFA8">Fiopt data on eaolv inptao Afootoopical Miltogoamminae (Dipteoa: Saocophagidae) oeveal unexpected moophological diveopitv</mods:title>
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<figureCitation id="13742FB414596171FBB72FB3336DC7EB" box="[1026,1142,1699,1725]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="4" pageNumber="5">Figs 3AM</figureCitation>
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<figureCitation id="13742FB414596171FB352FB333A1C7EB" box="[1152,1210,1699,1725]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. First instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., habitus, lateral view. B, Hoplacephala sp., cephaloskeleton, lateral view. C, Hoplacephala sp., cephaloskeleton, ventral view. D, Lamprometopia concavatogena, habitus, lateral view. E, L. concavatogena, cephaloskeleton, lateral view. F, L.concavatogena, cephaloskeleton, ventral view. G, Phylloteles picifons, habitus, lateral view. H, P. picifons, cephaloskeleton, lateral view. I, P. picifons, cephaloskeleton, ventral view. Abbreviations:a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate." pageId="4" pageNumber="5">4GI</figureCitation>
<figureCitation id="13742FB414596171FB352FB333A1C7EB" box="[1152,1210,1699,1725]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. First instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., habitus, lateral view. B, Hoplacephala sp., cephaloskeleton, lateral view. C, Hoplacephala sp., cephaloskeleton, ventral view. D, Lamprometopia concavatogena, habitus, lateral view. E, L. concavatogena, cephaloskeleton, lateral view. F, L.concavatogena, cephaloskeleton, ventral view. G, Phylloteles picifons, habitus, lateral view. H, P. picifons, cephaloskeleton, lateral view. I, P. picifons, cephaloskeleton, ventral view. Abbreviations:a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate." figureDoi="http://doi.org/10.5281/zenodo.14827243" httpUri="https://zenodo.org/record/14827243/files/figure.png" pageId="4" pageNumber="5">4GI</figureCitation>
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<figureCitation id="13742FB414596171FB712FB433E2C7E8" box="[1220,1273,1700,1726]" captionStart="Figure 5" captionStartId="8.[129,194,1116,1140]" captionTargetBox="[129,1473,143,1088]" captionTargetId="figure-1280@8.[129,1473,143,1088]" captionTargetPageId="8" captionText="Figure 5. CLMS images of first instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., anterior part of cephaloskeleton, lateral view. B, Hoplacephala sp., mid part of cephaloskeleton, lateral view. C, Lamprometopia concavatogena, cephaloskeleton, lateral view. D, L. concavatogena, cephaloskeleton, dorso-lateral view. E, Phylloteles picifons, cephaloskeleton, lateral view. F, P.picifons, dorso-lateral view. Abbreviations:ahs, additional horseshoe-shaped sclerite; c/sbt, cirri/suprabuccal teeth; dc, dorsal cornu; es, epistomal sclerite; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; vc, ventral cornu; vp, vertical plate." pageId="4" pageNumber="5">5E, F</figureCitation>
<figureCitation id="13742FB414596171FB712FB433E2C7E8" box="[1220,1273,1700,1726]" captionStart="Figure 5" captionStartId="8.[129,194,1116,1140]" captionTargetBox="[129,1473,143,1088]" captionTargetId="figure-1280@8.[129,1473,143,1088]" captionTargetPageId="8" captionText="Figure 5. CLMS images of first instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., anterior part of cephaloskeleton, lateral view. B, Hoplacephala sp., mid part of cephaloskeleton, lateral view. C, Lamprometopia concavatogena, cephaloskeleton, lateral view. D, L. concavatogena, cephaloskeleton, dorso-lateral view. E, Phylloteles picifons, cephaloskeleton, lateral view. F, P.picifons, dorso-lateral view. Abbreviations:ahs, additional horseshoe-shaped sclerite; c/sbt, cirri/suprabuccal teeth; dc, dorsal cornu; es, epistomal sclerite; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; vc, ventral cornu; vp, vertical plate." figureDoi="http://doi.org/10.5281/zenodo.14827247" httpUri="https://zenodo.org/record/14827247/files/figure.png" pageId="4" pageNumber="5">5E, F</figureCitation>
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<emphasis id="B93BEF2314596171FC8C2E7C34C0C6D2" box="[825,987,1900,1924]" italics="true" pageId="4" pageNumber="5">Pseudocephalon:</emphasis>
Antennal complex (an) large, antennal dome (and) oval, with conical tip, antennal basal ring (abr) high (
<figureCitation id="13742FB414596171FCF12EBB34FAC694" box="[836,993,1963,1987]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="4" pageNumber="5">Fig. 3A, B, D, E</figureCitation>
<figureCitation id="13742FB414596171FCF12EBB34FAC694" box="[836,993,1963,1987]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="4" pageNumber="5">Fig. 3A, B, D, E</figureCitation>
); maxillary palpus (mp) shaped as a high cone (
<figureCitation id="13742FB41455617DFF3E2C41362BC43F" box="[139,304,1361,1386]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3A, B, DF</figureCitation>
<figureCitation id="13742FB41455617DFF3E2C41362BC43F" box="[139,304,1361,1386]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3A, B, DF</figureCitation>
), first sensillum basiconicum (sb1) very long and with slightly swollen tip, shifted away from central cluster of sensilla toward base of palpus, additional sensilla (ns1ns2) large and both situated dorsally to central cluster of sensilla (
<figureCitation id="13742FB41455617DFF3E2CDF37C9C4B1" box="[139,210,1487,1511]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3F</figureCitation>
<figureCitation id="13742FB41455617DFF3E2CDF37C9C4B1" box="[139,210,1487,1511]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3F</figureCitation>
); ventral organ (vo) on swollen, fleshy lobe (
<figureCitation id="13742FB41455617DFD2A2CDF37B6C750" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3A, B, D, E</figureCitation>
<figureCitation id="13742FB41455617DFD2A2CDF37B6C750" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3A, B, D, E</figureCitation>
); laterally to the mouth opening a few (one or two) cirri/ suprabuccal teeth (c/sbt) (
<figureCitation id="13742FB41455617DFE222F1D36C5C770" box="[407,478,1549,1574]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3E</figureCitation>
<figureCitation id="13742FB41455617DFE222F1D36C5C770" box="[407,478,1549,1574]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3E</figureCitation>
); oral ridges (or) well developed (
<figureCitation id="13742FB41455617DFF722F3D3650C712" box="[199,331,1581,1605]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3B, D, E</figureCitation>
<figureCitation id="13742FB41455617DFF722F3D3650C712" box="[199,331,1581,1605]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3B, D, E</figureCitation>
); antero-dorsal surface of pseudocephalon with irregular cuticular warts, postero-dorsal surface with fine cuticular ridges (
<figureCitation id="13742FB41455617DFE982F7B36B5C7D5" box="[301,430,1643,1668]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3A, B, D</figureCitation>
<figureCitation id="13742FB41455617DFE982F7B36B5C7D5" box="[301,430,1643,1668]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3A, B, D</figureCitation>
).
</paragraph>
<caption id="DF3063B914586170FFC42FFC34BDC69E" pageId="5" pageNumber="6" startId="5.[113,178,1772,1796]" targetBox="[160,1413,144,1744]" targetPageId="5" targetType="figure">
<caption id="DF3063B914586170FFC42FFC34BDC69E" ID-DOI="http://doi.org/10.5281/zenodo.14827237" ID-Zenodo-Dep="14827237" httpUri="https://zenodo.org/record/14827237/files/figure.png" pageId="5" pageNumber="6" startId="5.[113,178,1772,1796]" targetBox="[160,1413,144,1744]" targetPageId="5" targetType="figure">
<paragraph id="8BF0333114586170FFC42FFC34BDC69E" blockId="5.[113,1455,1772,1993]" pageId="5" pageNumber="6">
<emphasis id="B93BEF2314586170FFC42FFC37D2C652" bold="true" box="[113,201,1772,1796]" pageId="5" pageNumber="6">Figure 2.</emphasis>
First instar larva of
@ -149,7 +152,7 @@ First instar larva of
. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus. F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ.
</paragraph>
</caption>
<caption id="DF3063B9145B6173FF342FFD33A5C69E" pageId="6" pageNumber="7" startId="6.[129,193,1773,1797]" targetBox="[266,1337,145,1745]" targetPageId="6" targetType="figure">
<caption id="DF3063B9145B6173FF342FFD33A5C69E" ID-DOI="http://doi.org/10.5281/zenodo.14827241" ID-Zenodo-Dep="14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="6" pageNumber="7" startId="6.[129,193,1773,1797]" targetBox="[266,1337,145,1745]" targetPageId="6" targetType="figure">
<paragraph id="8BF03331145B6173FF342FFD33A5C69E" blockId="6.[129,1464,1773,1993]" pageId="6" pageNumber="7">
<emphasis id="B93BEF23145B6173FF342FFD37C3C653" bold="true" box="[129,216,1773,1797]" pageId="6" pageNumber="7">Figure 3.</emphasis>
First instar larva of
@ -159,7 +162,7 @@ First instar larva of
. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus. G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations: an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ.
</paragraph>
</caption>
<caption id="DF3063B9145A6172FFC42E413337C6A3" pageId="7" pageNumber="8" startId="7.[113,178,1873,1897]" targetBox="[210,1362,148,1844]" targetPageId="7" targetType="figure">
<caption id="DF3063B9145A6172FFC42E413337C6A3" ID-DOI="http://doi.org/10.5281/zenodo.14827243" ID-Zenodo-Dep="14827243" httpUri="https://zenodo.org/record/14827243/files/figure.png" pageId="7" pageNumber="8" startId="7.[113,178,1873,1897]" targetBox="[210,1362,148,1844]" targetPageId="7" targetType="figure">
<paragraph id="8BF03331145A6172FFC42E413337C6A3" blockId="7.[113,1428,1873,2038]" pageId="7" pageNumber="8">
<emphasis id="B93BEF23145A6172FFC42E4137D2C63F" bold="true" box="[113,201,1873,1897]" pageId="7" pageNumber="8">Figure 4.</emphasis>
First instar larva of Afrotropical Miltogramminae. A,
@ -201,7 +204,7 @@ sp., cephaloskeleton, ventral view. D,
, cephaloskeleton, ventral view. Abbreviations: a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate.
</paragraph>
</caption>
<caption id="DF3063B91455617DFF342D4C3212C5B2" pageId="8" pageNumber="9" startId="8.[129,194,1116,1140]" targetBox="[129,1473,143,1088]" targetPageId="8" targetType="figure">
<caption id="DF3063B91455617DFF342D4C3212C5B2" ID-DOI="http://doi.org/10.5281/zenodo.14827247" ID-Zenodo-Dep="14827247" httpUri="https://zenodo.org/record/14827247/files/figure.png" pageId="8" pageNumber="9" startId="8.[129,194,1116,1140]" targetBox="[129,1473,143,1088]" targetPageId="8" targetType="figure">
<paragraph id="8BF033311455617DFF342D4C3212C5B2" blockId="8.[129,1473,1116,1252]" pageId="8" pageNumber="9">
<emphasis id="B93BEF231455617DFF342D4C37C3C522" bold="true" box="[129,216,1116,1140]" pageId="8" pageNumber="9">Figure 5.</emphasis>
CLMS images of first instar larva of Afrotropical Miltogramminae. A,
@ -234,42 +237,42 @@ sp., mid part of cephaloskeleton, lateral view. C,
<paragraph id="8BF033311455617DFF342FB5326DC4DE" blockId="8.[127,778,1701,1976]" lastBlockId="8.[825,1474,1361,1417]" pageId="8" pageNumber="9">
<emphasis id="B93BEF231455617DFF342FB5363EC7EB" box="[129,293,1701,1725]" italics="true" pageId="8" pageNumber="9">Cephaloskeleton:</emphasis>
Labrum (lb) straight except for anterior part curved down, tip pointed (Figs 4H, 5E); mouthhook (mh) with a lateral arm at the base, almost parallel-sided and very slightly upcurved, mouthhook pick-axe shaped with a single pointed tip directed at an angle of about 45°C to longitudinal plane of mouthhook (Figs 4H, 5E, F); intermediate sclerite (is) slightly below the base of labrum and parastomal bars (pb) in lateral view (Figs 4H, 5E, F),in ventral view intermediate sclerite longer than wide (
<figureCitation id="13742FB41455617DFF4B2E8F3659C6EE" box="[254,322,1951,1976]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. First instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., habitus, lateral view. B, Hoplacephala sp., cephaloskeleton, lateral view. C, Hoplacephala sp., cephaloskeleton, ventral view. D, Lamprometopia concavatogena, habitus, lateral view. E, L. concavatogena, cephaloskeleton, lateral view. F, L.concavatogena, cephaloskeleton, ventral view. G, Phylloteles picifons, habitus, lateral view. H, P. picifons, cephaloskeleton, lateral view. I, P. picifons, cephaloskeleton, ventral view. Abbreviations:a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate." pageId="8" pageNumber="9">Fig. 4I</figureCitation>
<figureCitation id="13742FB41455617DFF4B2E8F3659C6EE" box="[254,322,1951,1976]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. First instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., habitus, lateral view. B, Hoplacephala sp., cephaloskeleton, lateral view. C, Hoplacephala sp., cephaloskeleton, ventral view. D, Lamprometopia concavatogena, habitus, lateral view. E, L. concavatogena, cephaloskeleton, lateral view. F, L.concavatogena, cephaloskeleton, ventral view. G, Phylloteles picifons, habitus, lateral view. H, P. picifons, cephaloskeleton, lateral view. I, P. picifons, cephaloskeleton, ventral view. Abbreviations:a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate." figureDoi="http://doi.org/10.5281/zenodo.14827243" httpUri="https://zenodo.org/record/14827243/files/figure.png" pageId="8" pageNumber="9">Fig. 4I</figureCitation>
); parastomal bar long (Figs 4H, I, 5E, F); vertical plate (vp) slightly broader than ventral cornu (vc) and dorsal cornu (dc) (Figs 4H, 5E, F); dorsal bridge absent.
</paragraph>
<paragraph id="8BF033311455617DFC8C2CBB320BC66C" blockId="8.[825,1475,1450,1851]" pageId="8" pageNumber="9">
<emphasis id="B93BEF231455617DFC8C2CBB34F5C494" box="[825,1006,1451,1474]" italics="true" pageId="8" pageNumber="9">Toracic segments:</emphasis>
Anterior spinose band on t1 interrupted dorsally, laterally with 5 to 7 and ventrally with 1113 rows of spines (
<figureCitation id="13742FB41455617DFCF12CF934B6C756" box="[836,941,1513,1537]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3A, B</figureCitation>
<figureCitation id="13742FB41455617DFCF12CF934B6C756" box="[836,941,1513,1537]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3A, B</figureCitation>
); spinose band on t2 with only one to three rows of spines and restricted to ventro-lateral and ventral surfaces (
<figureCitation id="13742FB41455617DFCF12F373495C716" box="[836,910,1575,1600]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3B</figureCitation>
<figureCitation id="13742FB41455617DFCF12F373495C716" box="[836,910,1575,1600]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3B</figureCitation>
), spinose band on t3 complete but narrow and with three to five rows of spines; surface of integument with dense, fine, cuticular ridges (
<figureCitation id="13742FB41455617DFBA82F76339CC728" box="[1053,1159,1638,1662]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3AC</figureCitation>
<figureCitation id="13742FB41455617DFBA82F76339CC728" box="[1053,1159,1638,1662]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3AC</figureCitation>
), dorsal surfaces of t1t3 also with large and fragmented ridges (
<figureCitation id="13742FB41455617DFB292F953213C7CB" box="[1180,1288,1669,1694]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3G, H</figureCitation>
<figureCitation id="13742FB41455617DFB292F953213C7CB" box="[1180,1288,1669,1694]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3G, H</figureCitation>
); t1 dorsally with a small cuticular protuberance (cp) in the centre, followed by large bulbous protuberance (Figs 3A, B, 4G, H); integumental papillae in form of conical protuberances, each with four to six ridges converging at the top sensillum (
<figureCitation id="13742FB41455617DFB7E2E13327BC64C" box="[1227,1376,1795,1819]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3B, C, G, J</figureCitation>
<figureCitation id="13742FB41455617DFB7E2E13327BC64C" box="[1227,1376,1795,1819]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3B, C, G, J</figureCitation>
); Keilins organ with slightly elongated sensilla (
<figureCitation id="13742FB41455617DFB0B2E3233E5C66C" box="[1214,1278,1826,1850]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3I</figureCitation>
<figureCitation id="13742FB41455617DFB0B2E3233E5C66C" box="[1214,1278,1826,1850]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3I</figureCitation>
).
</paragraph>
<paragraph id="8BF033311455617CFC8C2E7137E6C4CE" blockId="8.[825,1474,1889,1976]" lastBlockId="9.[113,762,1063,1432]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B93BEF231455617DFC8C2E713314C62F" box="[825,1039,1889,1913]" italics="true" pageId="8" pageNumber="9">Abdominal segments:</emphasis>
Anterior spinose bands (asb) on a1a7 restricted to lateral and ventral surfaces with from 23 to 89 rows of spines (
<figureCitation id="13742FB41455617DFC682EB03333C6EE" box="[989,1064,1952,1976]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3K</figureCitation>
<figureCitation id="13742FB41455617DFC682EB03333C6EE" box="[989,1064,1952,1976]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="8" pageNumber="9">Fig. 3K</figureCitation>
), spines small and arranged in irregular lines (
<figureCitation id="13742FB41454617CFF0D2D37361EC569" box="[184,261,1063,1087]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3L</figureCitation>
<figureCitation id="13742FB41454617CFF0D2D37361EC569" box="[184,261,1063,1087]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="9" pageNumber="10">Fig. 3L</figureCitation>
); posterior spinose bands (psb) not well defined (
<figureCitation id="13742FB41454617CFF022D57363CC508" box="[183,295,1095,1119]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3K, M</figureCitation>
<figureCitation id="13742FB41454617CFF022D57363CC508" box="[183,295,1095,1119]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="9" pageNumber="10">Fig. 3K, M</figureCitation>
); lateral creeping welts (lcw) developed and covered by cuticular ridges and sparse spines; dorsal and lateral surfaces of all abdominal segments with a mixture of fine ridges and scattered large fragmented ridges, ventral surface of segments with regular cuticular ridges (
<figureCitation id="13742FB41454617CFDF42DD4358FC58A" box="[577,660,1220,1244]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3H</figureCitation>
<figureCitation id="13742FB41454617CFDF42DD4358FC58A" box="[577,660,1220,1244]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="9" pageNumber="10">Fig. 3H</figureCitation>
); integumental papillae in form of conical protuberances, with four to six ridges converging at the top sensillum (
<figureCitation id="13742FB41454617CFD9E2C133576C44D" box="[555,621,1283,1307]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3J</figureCitation>
<figureCitation id="13742FB41454617CFD9E2C133576C44D" box="[555,621,1283,1307]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="9" pageNumber="10">Fig. 3J</figureCitation>
); ventral surface with large segmentally arranged transverse protuberances formed by fusion of the posterior prolegs (pos) of one segment and the ventral creeping welt (vcw) of the following segment (
<figureCitation id="13742FB41454617CFFC92C9037F0C4CE" box="[124,235,1408,1432]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3K, M</figureCitation>
<figureCitation id="13742FB41454617CFFC92C9037F0C4CE" box="[124,235,1408,1432]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="9" pageNumber="10">Fig. 3K, M</figureCitation>
).
</paragraph>
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<caption id="DF3063B91454617CFFC42A5235F6C554" ID-DOI="http://doi.org/10.5281/zenodo.14827251" ID-Zenodo-Dep="14827251" httpUri="https://zenodo.org/record/14827251/files/figure.png" pageId="9" pageNumber="10" startId="9.[113,146,834,858]" targetBox="[120,757,145,797]" targetPageId="9" targetType="figure">
<paragraph id="8BF033311454617CFFC42A5235F6C554" blockId="9.[113,749,834,1026]" pageId="9" pageNumber="10">
<emphasis id="B93BEF231454617CFFC42A5237B5C20C" bold="true" box="[113,174,834,858]" pageId="9" pageNumber="10">Fig. 6.</emphasis>
The phylogenetic tree constructed by the NJ method with 500 bootstrap replications based on mitochondrial
@ -286,13 +289,13 @@ Piwczyński
<paragraph id="8BF033311454617CFFC42CA93637C61F" blockId="9.[113,762,1465,1866]" pageId="9" pageNumber="10">
<emphasis id="B93BEF231454617CFFC42CA9361EC487" box="[113,261,1465,1489]" italics="true" pageId="9" pageNumber="10">Anal division:</emphasis>
Anterior spinose bands (asb) on a1a7 restricted to lateral and ventral surfaces, spines small and arranged in irregular lines (
<figureCitation id="13742FB41454617CFECB2CE836CBC746" box="[382,464,1528,1552]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3M</figureCitation>
<figureCitation id="13742FB41454617CFECB2CE836CBC746" box="[382,464,1528,1552]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="9" pageNumber="10">Fig. 3M</figureCitation>
); dorsal and lateral surfaces of anal division with mixture of fine ridges and scattered large fragmented ridges, ventral surface of ad with regular cuticular ridges (
<figureCitation id="13742FB41454617CFF7C2F46363AC738" box="[201,289,1622,1646]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3M</figureCitation>
<figureCitation id="13742FB41454617CFF7C2F46363AC738" box="[201,289,1622,1646]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="9" pageNumber="10">Fig. 3M</figureCitation>
); papillae around spiracular field distinct and in form of conical protuberances, with four to six ridges converging at the top sensillum; spiracular field ringed by a dorsally interrupted circle of hair-like spines; posterior spiracle (ps) with four peristigmatic tufts each with a few (one to four) branches; anal complex strongly protruded; anal papillae (ap) large and rounded (
<figureCitation id="13742FB41454617CFE702E023501C67C" box="[453,538,1810,1834]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3M</figureCitation>
<figureCitation id="13742FB41454617CFE702E023501C67C" box="[453,538,1810,1834]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="9" pageNumber="10">Fig. 3M</figureCitation>
); spines of anal tuft absent (
<figureCitation id="13742FB41454617CFF7D2E213601C61F" box="[200,282,1841,1865]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3M</figureCitation>
<figureCitation id="13742FB41454617CFF7D2E213601C61F" box="[200,282,1841,1865]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." figureDoi="http://doi.org/10.5281/zenodo.14827241" httpUri="https://zenodo.org/record/14827241/files/figure.png" pageId="9" pageNumber="10">Fig. 3M</figureCitation>
).
</paragraph>
</subSubSection>
@ -304,7 +307,7 @@ Gravid females of
<emphasis id="B93BEF231454617CFE2D2E7B351AC6D5" box="[408,513,1899,1923]" italics="true" pageId="9" pageNumber="10">P. picifons</emphasis>
</taxonomicName>
were observed perching on small tumuli formed by termites. After removing the tumuli, females quickly entered into the opened entrance (
<figureCitation id="13742FB41454617CFD342EB935F2C697" box="[641,745,1961,1985]" captionStart="Fig" captionStartId="10.[129,162,963,987]" captionTargetBox="[129,1473,143,935]" captionTargetId="figure-1476@10.[129,1473,143,935]" captionTargetPageId="10" captionText="Fig. 7. Habitats and females of Miltogramminae. A, the tree in Homeb canyon, where gravid female of Lamprometopia concavatogena was collected. B, termites on the tree trunk. C, female of L. concavatogena on the ground at the trunk. D, termite nest tumulus with perching female of Phylloteles picifons (pointed by arrow). E, female of P.picifons at the top of tumulus. F, female P.picifons entering into the open corridor." pageId="9" pageNumber="10">Fig. 7DF</figureCitation>
<figureCitation id="13742FB41454617CFD342EB935F2C697" box="[641,745,1961,1985]" captionStart="Fig" captionStartId="10.[129,162,963,987]" captionTargetBox="[129,1473,143,935]" captionTargetId="figure-1476@10.[129,1473,143,935]" captionTargetPageId="10" captionText="Fig. 7. Habitats and females of Miltogramminae. A, the tree in Homeb canyon, where gravid female of Lamprometopia concavatogena was collected. B, termites on the tree trunk. C, female of L. concavatogena on the ground at the trunk. D, termite nest tumulus with perching female of Phylloteles picifons (pointed by arrow). E, female of P.picifons at the top of tumulus. F, female P.picifons entering into the open corridor." figureDoi="http://doi.org/10.5281/zenodo.14827253" httpUri="https://zenodo.org/record/14827253/files/figure.png" pageId="9" pageNumber="10">Fig. 7DF</figureCitation>
). Attempts to raise first instars of
<taxonomicName id="4C4F48B21454617CFBD9298033C8C1FE" authorityName="Pape" authorityYear="1985" box="[1132,1235,144,168]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF231454617CFBD9298033C8C1FE" box="[1132,1235,144,168]" italics="true" pageId="9" pageNumber="10">P. picifons</emphasis>