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Etymology:
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, from the Shoshone
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·tum (pronounced ah-vay-tum), meaning long ago in reference to the ancient nature of UWGM 1975. Pronunciation guide: the sound ah is as in autumn, vay as in vague, and tum as in autumn. See Supporting Information, Audio S1 (discussion of name); recordings by Eastern Shoshone Elder (RT).
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species:
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.
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<emphasis id="B95CF77B0872FFA11435C9F9FC311A46" box="[825,927,1784,1807]" italics="true" pageId="4">Diagnosis:</emphasis>
As for
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species, by monotypy.
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<mods:role id="CAEB7E04BA95ACBB357D15C18FF769A9">
<mods:roleTerm id="918F27C3A81EEEC6CE01700E1B256BB6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E81C634B4DF68135A42F1478379F78E6">Mann, Joshua</mods:namePart>
<mods:affiliation id="3862598167F63AB5F3A8A77E7D6FAAFD">Eastern Shoshone Tribal Historic Preservation Office, Bldg 17 A, North Fork Rd., Fort Washakie, WY 82514, United States</mods:affiliation>
</mods:name>
<mods:name id="AE1AFCEC7443776C494490F396EE4638" type="personal">
<mods:role id="B8CF2213B403FAC65434D987677E0DFE">
<mods:roleTerm id="B090A55C8D4A81082C6574D48AC7BB48">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="790E2B2DED246C4046FF7993C77D4F45">Teran, Reba</mods:namePart>
<mods:affiliation id="44F987F594D322AF9F98642515DF6CD2">Shoshone Language Consultant, Wind River Reservation, Fort Washakie, WY 82514, United States</mods:affiliation>
</mods:name>
<mods:typeOfResource id="89BF7B333EA14B77A57A0E984154A427">text</mods:typeOfResource>
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<mods:titleInfo id="452B9B98B46866AE3E0A7F5D94695502">
<mods:title id="B8C89D1FAB9ACA7C90F0E89ECEA7137F">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="B546DABC66BC30DD71AFB7FA8D1104B9">
<mods:date id="EC39FF3FB412E688E292BB95C49681E1">2025</mods:date>
<mods:detail id="E27C1BD367E10B360B2300001AA1208B" type="volume">
<mods:number id="C301B5FEC47860DD2FE3E4A5E5A44CD3">203</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="09A719D7B616AB0D64210562DAF30BDD">journal volume</mods:classification>
<mods:identifier id="6A0FB5C378A3505E65C5CA5BF29301F2" type="CLB-Dataset">308019</mods:identifier>
<mods:identifier id="CC3502BBCBF186549030B7C4BA737C93" type="DOI">10.1093/zoolinnean/zlae153</mods:identifier>
<mods:identifier id="280A500A56906C93DCF044E2AB3C4A52" type="GBIF-Dataset">744b4603-204d-448c-a32f-fd0be17f047b</mods:identifier>
<mods:identifier id="FC806F45A70CCC3ABFF0EE12C95434D2" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="D40B23877230E3542E6AF5C2735E933C" type="Zenodo-Dep">14827184</mods:identifier>
<mods:identifier id="375B28E731BB5316422A4614D9914F8A" type="ZooBank">38C156C-5CC1-4706-8D8A-FFA6757A6590</mods:identifier>
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<treatment id="03819A7F0872FFAD14B3C833FE7819A5" ID-DOI="http://doi.org/10.5281/zenodo.14827192" ID-Zenodo-Dep="14827192" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5" httpUri="http://treatment.plazi.org/id/03819A7F0872FFAD14B3C833FE7819A5" lastPageId="8" pageId="4">
<subSubSection id="C33278E20872FFA114B3C833FA921A04" box="[959,1340,1842,1869]" pageId="4" type="nomenclature">
<paragraph id="8B972B690872FFA114B3C833FA921A04" blockId="4.[959,1340,1842,1869]" box="[959,1340,1842,1869]" pageId="4">
<heading id="D0DF9C050872FFA114B3C833FA921A04" bold="true" box="[959,1340,1842,1869]" centered="true" fontSize="11" level="2" pageId="4" reason="2">
<emphasis id="B95CF77B0872FFA114B3C833FA921A04" bold="true" box="[959,1340,1842,1869]" pageId="4">
<taxonomicName id="4C2850EA0872FFA114B3C833FB4E1A04" authorityName="Lovelace &amp; Kufner &amp; Fitch &amp; Rogers &amp; Schmitz &amp; Schwartz &amp; LeClair-Diaz &amp; St.Clair &amp; Mann &amp; Teran" authorityYear="2025" box="[959,1248,1842,1869]" genus="Ahvaytum" kingdom="Animalia" order="Dinosauria" pageId="4" rank="species" species="bahndooiveche" status="sp. nov.">
<emphasis id="B95CF77B0872FFA114B3C833FB4E1A04" bold="true" box="[959,1248,1842,1869]" italics="true" pageId="4">Ahvaytum bahndooiveche</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F4A000872FFA113E9C832FA921A04" box="[1253,1340,1843,1869]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33278E20872FFAD1358C870FE7819A5" lastPageId="8" pageId="4" type="description">
<paragraph id="8B972B690872FFA11358C870FB091AC2" blockId="4.[1108,1191,1905,1931]" box="[1108,1191,1905,1931]" pageId="4">
<emphasis id="B95CF77B0872FFA11358C870FB091AC2" bold="true" box="[1108,1191,1905,1931]" pageId="4">
(
<figureCitation id="131337EC0872FFA11352C870FB361AC2" box="[1118,1176,1905,1931]" captionStart="Figure 2" captionStartId="4.[129,194,885,909]" captionTargetBox="[131,1471,146,855]" captionTargetId="figure-419@4.[129,1473,144,857]" captionTargetPageId="4" captionText="Figure 2. Holotype left astragalus of Ahvaytum bahndooiveche gen.et sp. nov.(UWGM 1975). 3D model in (A) medial, (B) lateral, (C) lateral transparent, (D) posterior transparent, (E) distal, (F) proximal, (I) anterior, and (J) posterior orthographic views. Photographs in (G) proximal and (H) distal views. Abbreviations: amc, anteromedial corner; ap, ascending process of the astragalus; g, groove; f, foramen; ff, fibular facet; ldn, laterodistal notch (= lateroventral depression); mf, medial fossa; nf, non-articular fossa (= dorsal basin, = semi-elliptical fossa); p, platform; plp, posterolateral process; plr, posterolateral ridge; tf, tibial facet. Diagonal lines indicate broken surfaces.Arrows indicate anterior direction. Scale bar = 1 cm. STL models archived online (morphosource.org ID: 000607569)." figureDoi="http://doi.org/10.5281/zenodo.14827190" httpUri="https://zenodo.org/record/14827190/files/figure.png" pageId="4">Fig. 2</figureCitation>
).
</emphasis>
</paragraph>
<paragraph id="8B972B690872FFA01435C8B0FF721C6C" blockId="4.[825,1391,1969,1994]" lastBlockId="5.[113,764,144,293]" lastPageId="5" pageId="4">
<materialsCitation id="3B4021340872FFA11435C8B0FAC11A83" box="[825,1391,1969,1994]" collectionCode="UWGM" pageId="4" specimenCode="UWGM 1975, UWGM 7549" specimenCount="1" typeStatus="holotype">
<emphasis id="B95CF77B0872FFA11435C8B0FC371A80" box="[825,921,1969,1993]" italics="true" pageId="4">
<typeStatus id="549395CB0872FFA11435C8B0FC3A1A80" box="[825,916,1969,1993]" pageId="4" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DB8E83120872FFA11493C8B3FB9E1A80" box="[927,1072,1969,1994]" collectionCode="UWGM" pageId="4">UWGM 1975</specimenCode>
, isolated left astragalus (
<figureCitation id="131337EC0872FFA1122AC8B0FAF11A83" box="[1318,1375,1969,1994]" captionStart="Figure 2" captionStartId="4.[129,194,885,909]" captionTargetBox="[131,1471,146,855]" captionTargetId="figure-419@4.[129,1473,144,857]" captionTargetPageId="4" captionText="Figure 2. Holotype left astragalus of Ahvaytum bahndooiveche gen.et sp. nov.(UWGM 1975). 3D model in (A) medial, (B) lateral, (C) lateral transparent, (D) posterior transparent, (E) distal, (F) proximal, (I) anterior, and (J) posterior orthographic views. Photographs in (G) proximal and (H) distal views. Abbreviations: amc, anteromedial corner; ap, ascending process of the astragalus; g, groove; f, foramen; ff, fibular facet; ldn, laterodistal notch (= lateroventral depression); mf, medial fossa; nf, non-articular fossa (= dorsal basin, = semi-elliptical fossa); p, platform; plp, posterolateral process; plr, posterolateral ridge; tf, tibial facet. Diagonal lines indicate broken surfaces.Arrows indicate anterior direction. Scale bar = 1 cm. STL models archived online (morphosource.org ID: 000607569)." figureDoi="http://doi.org/10.5281/zenodo.14827190" httpUri="https://zenodo.org/record/14827190/files/figure.png" pageId="4">Fig. 2</figureCitation>
).
</materialsCitation>
<emphasis id="B95CF77B0873FFA0177DCF91FE881DE1" box="[113,294,144,168]" italics="true" pageId="5">Referred material:</emphasis>
<materialsCitation id="3B4021340873FFA0163CCF91FF721C6C" collectionCode="UWGM" pageId="5" specimenCode="UWGM 1975, UWGM 7549" specimenCount="1" typeStatus="holotype">
The proximal end of a left femur (
<specimenCode id="DB8E83120873FFA015ACCF91FF061D8E" collectionCode="UWGM" pageId="5">UWGM 7549</specimenCode>
;
<figureCitation id="131337EC0873FFA017BFCFAEFF451D8E" box="[179,235,175,199]" captionStart="Figure 3" captionStartId="5.[113,178,1842,1866]" captionTargetBox="[149,1426,449,1812]" captionTargetId="figure-133@5.[145,1428,447,1814]" captionTargetPageId="5" captionText="Figure 3. Proximal end of a left femur UWGM 7549 (AE) referred to Ahvaytum bahndooiveche gen. et sp. nov.). 3D model in (A) anteromedial, (B) posterolateral, (C) proximal, (D) anterolateral, and (E) posteromedial orthographic views. Abbreviations: alt, anterolateral tuber; amt, anteromedial tuber; ce, concave emargination; dlt, dorsolateral trochanter; ft, fossa trochanterica (= facies articularis antitrochanterica); gt, greater trochanter; pmt, posteromedial tuber; ve, ventral emargination.Arrows indicate anterior direction. Scale bar = 1 cm. STL models archived online (morphosource.org ID: 000607572)." figureDoi="http://doi.org/10.5281/zenodo.14827194" httpUri="https://zenodo.org/record/14827194/files/figure.png" pageId="5">Fig. 3</figureCitation>
) is referred to
<taxonomicName id="4C2850EA0873FFA01688CFAEFDD01D8E" authorityName="Lovelace &amp; Kufner &amp; Fitch &amp; Rogers &amp; Schmitz &amp; Schwartz &amp; LeClair-Diaz &amp; St.Clair &amp; Mann &amp; Teran" authorityYear="2025" box="[388,638,175,199]" genus="Ahvaytum" pageId="5" rank="species" species="bahndooiveche">
<emphasis id="B95CF77B0873FFA01688CFAEFDD01D8E" box="[388,638,175,199]" italics="true" pageId="5">Ahvaytum bahndooiveche</emphasis>
</taxonomicName>
due to clear saurischian affinities. This femur was found
<emphasis id="B95CF77B0873FFA0153FCFCEFDDC1DAF" box="[563,626,207,230]" italics="true" pageId="5">ex situ</emphasis>
within a 5-m radius of the type specimen and encrusted with a similar micritic carbonate.
</materialsCitation>
</paragraph>
<paragraph id="8B972B690873FFA3177DCE4EFDD11C4F" blockId="5.[113,762,335,391]" lastBlockId="6.[129,778,144,262]" lastPageId="6" pageId="5">
<emphasis id="B95CF77B0873FFA0177DCE4EFED41C2E" box="[113,378,335,359]" italics="true" pageId="5">
Etymology:
<taxonomicName id="4C2850EA0873FFA017E6CE4EFED41C2E" authorityName="Lovelace &amp; Kufner &amp; Fitch &amp; Rogers &amp; Schmitz &amp; Schwartz &amp; LeClair-Diaz &amp; St.Clair &amp; Mann &amp; Teran" authorityYear="2025" box="[234,378,335,359]" genus="Ahvaytum" pageId="5" rank="species" species="bahndooiveche">bahndooiveche</taxonomicName>
</emphasis>
, from the Shoshone ban·döi·ve·che (pronounced bon-do-ee-vee-chee), meaning waters young handsome man in reference to the colourful salamanders found in the region, and also means dinosaur (Supporting Information, Audio S1), which is the meaning used here. The name
<taxonomicName id="4C2850EA0873FFA014AECFEFFB0A1C4F" authorityName="Lovelace &amp; Kufner &amp; Fitch &amp; Rogers &amp; Schmitz &amp; Schwartz &amp; LeClair-Diaz &amp; St.Clair &amp; Mann &amp; Teran" authorityYear="2025" box="[930,1188,238,262]" genus="Ahvaytum" pageId="5" rank="species" species="bahndooiveche">
<emphasis id="B95CF77B0873FFA014AECFEFFB0A1C4F" box="[930,1188,238,262]" italics="true" pageId="5">Ahvaytum bahndooiveche</emphasis>
</taxonomicName>
(long ago dinosaur) is the product of a multigenerational collaboration between the Fort Washakie Schools 7th grade cohort (2022), educators, Eastern Shoshone Tribal Historic Preservation Office, and Tribal Elders. Pronunciation guide: bahn as in bonnet, do-ee as in dewy, vee as in ivy, and chee as in cheese. See the Supporting Information, Audio S1 for recorded pronunciation (recording by RT. LSID
<uri id="FFB9276B0870FFA316F4CFCEFDDD1C4F" pageId="6">
urn:lsid:zoobank.org:pub:
<uuid id="FF8E11BC0870FFA3178DCFEFFDDD1C4F" box="[129,627,238,262]" pageId="6">538C156C-5CC1-4706-8D8A-FFA6757A6590</uuid>
</uri>
).
</paragraph>
<caption id="DF577BE10873FFA0177DC833FCFC1AF2" ID-DOI="http://doi.org/10.5281/zenodo.14827194" ID-Zenodo-Dep="14827194" httpUri="https://zenodo.org/record/14827194/files/figure.png" pageId="5" startId="5.[113,178,1842,1866]" targetBox="[149,1426,449,1812]" targetPageId="5" targetType="figure">
<paragraph id="8B972B690873FFA0177DC833FCFC1AF2" blockId="5.[113,1439,1842,1979]" pageId="5">
<emphasis id="B95CF77B0873FFA0177DC833FF671A02" bold="true" box="[113,201,1842,1867]" pageId="5">Figure 3.</emphasis>
Proximal end of a left femur UWGM 7549 (AE) referred to
<taxonomicName id="4C2850EA0873FFA01400C832FC5D1A02" authorityName="Lovelace &amp; Kufner &amp; Fitch &amp; Rogers &amp; Schmitz &amp; Schwartz &amp; LeClair-Diaz &amp; St.Clair &amp; Mann &amp; Teran" authorityYear="2025" box="[780,1011,1843,1867]" genus="Ahvaytum" pageId="5" rank="species" species="bahndooiveche" status="gen. et sp. nov.">
<emphasis id="B95CF77B0873FFA01400C832FC5D1A02" box="[780,1011,1843,1867]" italics="true" pageId="5">Ahvaytum bahndooiveche</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F4A000873FFA014FBC832FB2B1A02" box="[1015,1157,1843,1867]" pageId="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
). 3D model in (A) anteromedial, (B) posterolateral, (C) proximal, (D) anterolateral, and (E) posteromedial orthographic views. Abbreviations: alt, anterolateral tuber; amt, anteromedial tuber; ce, concave emargination; dlt, dorsolateral trochanter; ft, fossa trochanterica (= facies articularis antitrochanterica); gt, greater trochanter; pmt, posteromedial tuber; ve, ventral emargination. Arrows indicate anterior direction. Scale bar = 1 cm. STL models archived online (morphosource.org ID: 000607572).
</paragraph>
</caption>
<paragraph id="8B972B690870FFA3178DCE26FD531FBF" blockId="6.[127,780,295,758]" pageId="6">
<emphasis id="B95CF77B0870FFA3178DCE26FEF11C76" box="[129,351,295,319]" italics="true" pageId="6">Locality and horizon:</emphasis>
<collectingCounty id="62F653E50870FFA31661CE26FD8B1C76" box="[365,549,295,319]" pageId="6">Garretts Surprise</collectingCounty>
(after undergraduate field assistant
<collectingMunicipality id="6BF3B1130870FFA3161CCE46FE1B1C16" box="[272,437,327,351]" pageId="6">Garrett Johnson</collectingMunicipality>
who discovered the locality) is a small exposure of the
<location id="8EF77DB20870FFA31656CE67FD9A1C37" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA31656CE67FD9A1C37" box="[346,564,358,382]" county="Garrett's Surprise" municipality="Garrett Johnson" name="Popo Agie Formation" pageId="6" stateProvince="Wyoming">Popo Agie Formation</location>
found in an erosional window within the overlying
<location id="8EF77DB20870FFA316BDCE84FD321CD4" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA316BDCE84FD321CD4" box="[433,668,389,413]" county="Garrett's Surprise" municipality="Garrett Johnson" name="Wind River Formation" pageId="6" stateProvince="Wyoming">
<location id="8EF77DB20870FFA316BDCE84FD871CD4" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA316BDCE84FD871CD4" box="[433,553,389,413]" county="Garrett's Surprise" municipality="Garrett Johnson" name="Wind River" pageId="6" stateProvince="Wyoming">
Wind
<location id="8EF77DB20870FFA316FECE84FD871CD4" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA316FECE84FD871CD4" box="[498,553,389,413]" county="Garrett's Surprise" municipality="Garrett Johnson" name="River Formation" pageId="6" stateProvince="Wyoming">River</location>
</location>
Formation
</location>
(
<location id="8EF77DB20870FFA315A7CE84FD501CD4" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA315A7CE84FD501CD4" box="[683,766,389,413]" county="Garrett's Surprise" municipality="Garrett Johnson" name="Eocene" pageId="6" stateProvince="Wyoming">Eocene</location>
), on
<collectingRegion id="49ECE58B0870FFA317AFCEA4FEA91CF4" box="[163,263,421,445]" country="United States of America" name="Wyoming" pageId="6">Wyoming</collectingRegion>
<location id="8EF77DB20870FFA31602CEA4FEE21CF4" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA31602CEA4FEE21CF4" box="[270,332,421,445]" county="Garrett's Surprise" municipality="Garrett Johnson" name="Game" pageId="6" stateProvince="Wyoming">Game</location>
and
<location id="8EF77DB20870FFA31673CEA4FE041CF4" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA31673CEA4FE041CF4" box="[383,426,421,445]" county="Garrett's Surprise" municipality="Garrett Johnson" name="Fish" pageId="6" stateProvince="Wyoming">Fish</location>
administered lands,
<emphasis id="B95CF77B0870FFA3158DCEA4FD261CF5" box="[641,648,421,444]" italics="true" pageId="6">c</emphasis>
.
<quantity id="4CD0868C0870FFA31598CEA4FD671CF4" box="[660,713,421,445]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" unit="km" value="1.0">1 km</quantity>
south of the confluence of the
<location id="8EF77DB20870FFA31672CEC5FE711C95" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA31672CEC5FE711C95" box="[382,479,452,476]" county="Garrett's Surprise" municipality="Garrett Johnson" name="East Fork" pageId="6" stateProvince="Wyoming">East Fork</location>
of the
<location id="8EF77DB20870FFA31525CEC5FD0E1C95" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA31525CEC5FD0E1C95" box="[553,672,452,476]" county="Garrett's Surprise" municipality="Garrett Johnson" name="Wind River" pageId="6" stateProvince="Wyoming">Wind River</location>
and
<location id="8EF77DB20870FFA315DFCEC5FF111CB2" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA315DFCEC5FF111CB2" county="Garrett's Surprise" municipality="Garrett Johnson" name="Spear Creek" pageId="6" stateProvince="Wyoming">Spear Creek</location>
(
<figureCitation id="131337EC0870FFA317D8CEE2FE8D1CB2" box="[212,291,483,507]" captionStart="Figure 1" captionStartId="2.[129,194,1602,1626]" captionTargetBox="[134,1470,147,1571]" captionTargetId="figure-6@2.[129,1473,144,1574]" captionTargetPageId="2" captionText="Figure 1. A palaeogeographic map (A) of Triassic outcrops shows the position of key localities (1 to 9); Chugwater Group, Wyoming, USA (inset), local geology surrounding Garretts Surprise (star). Chronostratigraphic context of early dinosaur-bearing units (B) shows the relative stratigraphic positions for first occurrences of dinosaurian body and trace fossils tied to a generalized stratigraphic section (C) of the upper Chugwater Group [after Lovelace et al. (2024)]. The shaded box highlights the stratigraphic position of UWGM 1975 and UWGM 7407 within the Popo Agie Formation. Localities:1, Argentina; 2, Brazil; 3, Zimbabwe; 4, India; 5, Germany; 6, Britain; 7, Morocco; 8, Texas, USA (Dockum Group); 9, south-western USA (Chinle/Moenkopi formations); 10, Wyoming, USA (Chugwater Group). Silhouettes (not to scale) on outcrop demonstrate faunal distribution within the Popo Agie and Jelm formations.At, Atreipus sp.; AtGr, AtreipusGrallator plexus; An, Anaschisma browni; Be, Beesiiwo cooowuse; Bu, Buettnererpeton bakeri; Br, Brachychirotherium sp.; Eu, Eubrachiosaurus browni; He, Heptasuchus clarki; Pa, Parasuchus bransoni; Po, Poposaurus gracilis; Ni, Ninumbeehan dookoodukah. Artist credits for silhouettes from Phylopic.org and abbreviated formation names provided in the Supporting Information, Text S1. Previously published dinosaur-bearing localities recovered from PBDB (PaleobioDB.org; accessed Feb, 2024; Supporting Information, Data S1)." figureDoi="http://doi.org/10.5281/zenodo.14827188" httpUri="https://zenodo.org/record/14827188/files/figure.png" pageId="6">Fig. 1A</figureCitation>
). GPS coordinates for
<location id="8EF77DB20870FFA3152BCEE2FD711CB2" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA3152BCEE2FD711CB2" box="[551,735,483,507]" county="Garrett's Surprise" municipality="Garrett Johnson" name="Garrett's Surprise" pageId="6" stateProvince="Wyoming">Garretts Surprise</location>
are reposited with the
<typeStatus id="549395CB0870FFA3164ACD02FEDD1F52" box="[326,371,515,539]" pageId="6">type</typeStatus>
specimen.
<collectorName id="26DD4EBF0870FFA316E5CD02FDFC1F52" box="[489,594,515,539]" pageId="6">Vertebrate</collectorName>
material was surface collected and no
<emphasis id="B95CF77B0870FFA3166DCD22FE301F73" box="[353,414,547,570]" italics="true" pageId="6">in situ</emphasis>
specimens were recovered during a screening effort of the area; all material is constrained to the mid to upper part of the purple unit of the
<location id="8EF77DB20870FFA316F9CD60FCA41F30" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA316F9CD60FCA41F30" box="[501,778,609,633]" county="Garrett's Surprise" municipality="Garrett Johnson" name="lower Popo Agie Formation" pageId="6" stateProvince="Wyoming">lower Popo Agie Formation</location>
(
<bibRefCitation id="EFB956980870FFA31787CD81FEE01FD1" author="Lovelace DM &amp; Fitch AJ &amp; Schwartz D" box="[139,334,640,664]" pageId="6" pagination="2305 - 24" refId="ref20218" refString="Lovelace DM, Fitch AJ, Schwartz D, et al. Concurrence of Late Triassic lithostratigraphic, radioisotopic, and biostratigraphic data support a Carnian age for the Popo Agie Formation (Chugwater Group), Wyoming, USA. GSA Bulletin 2024; 136: 2305 - 24. https: // doi. org / 10.1130 / B 36807.1" type="journal article" year="2024">
Lovelace
<emphasis id="B95CF77B0870FFA317E5CD80FEB91FD1" box="[233,279,640,664]" italics="true" pageId="6">et al.</emphasis>
2024
</bibRefCitation>
).
<collectorName id="26DD4EBF0870FFA3166ECD81FE271FD1" box="[354,393,640,664]" pageId="6">The</collectorName>
ochre unit is poorly exposed along the fall line where specimens were collected and is unconformably overlain by
<collectorName id="26DD4EBF0870FFA317F6CDBEFEEB1F9F" box="[250,325,703,726]" pageId="6">Eocene</collectorName>
deposits of the
<collectorName id="26DD4EBF0870FFA316E1CDBEFD761F9E" box="[493,728,703,727]" pageId="6">
<location id="8EF77DB20870FFA316E1CDBEFDCB1F9E" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA316E1CDBEFDCB1F9E" box="[493,613,703,727]" county="Garrett's Surprise" municipality="Garrett Johnson" name="Wind River" pageId="6" stateProvince="Wyoming">
Wind
<location id="8EF77DB20870FFA31522CDBEFDCB1F9E" LSID="urn:lsid:plazi:treatment:03819A7F0872FFAD14B3C833FE7819A5:8EF77DB20870FFA31522CDBEFDCB1F9E" box="[558,613,703,727]" county="Garrett's Surprise" municipality="Garrett Johnson" name="River Formation" pageId="6" stateProvince="Wyoming">River</location>
</location>
Formation
</collectorName>
[see
<collectorName id="26DD4EBF0870FFA3178DCDDFFEDB1FBF" box="[129,373,734,758]" pageId="6">Supporting Information</collectorName>
,
<collectorName id="26DD4EBF0870FFA31672CDDFFE6E1FBF" box="[382,448,734,758]" pageId="6">Text S</collectorName>
1 (
<collectorName id="26DD4EBF0870FFA316D9CDDFFD911FBF" box="[469,575,734,758]" pageId="6">Fieldwork</collectorName>
and
<collectorName id="26DD4EBF0870FFA31563CDDFFD441FBF" box="[623,746,734,758]" pageId="6">Site Details</collectorName>
)].
</paragraph>
<paragraph id="8B972B690870FFA3178DCC19FC471CCA" blockId="6.[126,781,791,1975]" lastBlockId="6.[825,1477,144,387]" pageId="6">
<emphasis id="B95CF77B0870FFA3178DCC19FE5A1E66" box="[129,500,791,815]" italics="true" pageId="6">
Diagnosis:
<taxonomicName id="4C2850EA0870FFA317F8CC16FE5A1E66" authorityName="Lovelace &amp; Kufner &amp; Fitch &amp; Rogers &amp; Schmitz &amp; Schwartz &amp; LeClair-Diaz &amp; St.Clair &amp; Mann &amp; Teran" authorityYear="2025" box="[244,500,791,815]" genus="Ahvaytum" pageId="6" rank="species" species="bahndooiveche">Ahvaytum bahndooiveche</taxonomicName>
</emphasis>
shares a combination of features present in sauropodomorph dinosaurs to the exclusion of theropods, herrerasaurs, and ornithischians, including: a flat, roller-shaped distal surface of the astragalus shared with sauropodomorphs but not herrerasaurs and neotheropods (
<bibRefCitation id="EFB956980870FFA31787CCB5FEE31E85" author="Marsh AD &amp; Parker WG &amp; Langer MC" box="[139,333,948,972]" pageId="6" refId="ref20319" refString="Marsh AD, Parker WG, Langer MC et al. Redescription of the holotype specimen of Chindesaurus bryansmalli Long and Murry, 1995 (Dinosauria, Theropoda), from Petrified Forest National Park, Arizona. Journal of Vertebrate Paleontology 2019; 39: e 1645682. https: // doi. org / 10.1080 / 02724634.2019.1645682" type="journal volume" year="2019">
Marsh
<emphasis id="B95CF77B0870FFA317D5CCB4FEA01E85" box="[217,270,948,972]" italics="true" pageId="6">et al.</emphasis>
2019
</bibRefCitation>
); a tibial facet that does not extend on to the anteromedial corner of the astragalus shared with all sauropodomorphs in our study, the silesaurid
<taxonomicName id="4C2850EA0870FFA31597CCF2FEDA1963" authority="Romer, 1972" authorityName="Romer" authorityYear="1972" genus="Lewisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="admixtus">
<emphasis id="B95CF77B0870FFA31597CCF2FF721963" italics="true" pageId="6">Lewisuchus admixtus</emphasis>
<bibRefCitation id="EFB956980870FFA317E4CB13FEDA1963" author="Romer AS" box="[232,372,1042,1066]" pageId="6" pagination="1 - 7" refId="ref22568" refString="Romer AS. The Chanares (Argentina) Triassic reptile fauna. XIV. Lewisuchus admixtus gen. et sp. nov., a further thecodont from the Chanares Beds. Breviora 1972; 390: 1 - 7." type="journal article" year="1972">Romer, 1972</bibRefCitation>
</taxonomicName>
, the theropods
<taxonomicName id="4C2850EA0870FFA31525CB13FD781963" authorityName="Nesbitt and Ezcurra" authorityYear="2015" box="[553,726,1042,1066]" family="Coelophysidae" genus="Lepidus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="praecisio">
<emphasis id="B95CF77B0870FFA31525CB13FD781963" box="[553,726,1042,1066]" italics="true" pageId="6">Lepidus praecisio</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2850EA0870FFA3178DCB30FD861900" authority="Martinez et al., 2011" authorityName="Martinez" authorityYear="2011" box="[129,552,1073,1097]" genus="Eodromaeus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="murphi">
<emphasis id="B95CF77B0870FFA3178DCB30FEE71900" box="[129,329,1073,1097]" italics="true" pageId="6">Eodromaeus murphi</emphasis>
<bibRefCitation id="EFB956980870FFA3165CCB30FD861900" author="Martinez RN &amp; Sereno PC &amp; Alcober OA" box="[336,552,1073,1097]" pageId="6" pagination="206 - 10" refId="ref20570" refString="Martinez RN, Sereno PC, Alcober OA et al. A basal dinosaur from the dawn of the dinosaur era in southwestern Pangaea. Science 2011; 331: 206 - 10. https: // doi. org / 10.1126 / science. 1198467" type="journal article" year="2011">
Martinez
<emphasis id="B95CF77B0870FFA316BACB33FE4F1900" box="[438,481,1073,1097]" italics="true" pageId="6">et al</emphasis>
., 2011
</bibRefCitation>
</taxonomicName>
, and the herrerasaurs
<taxonomicName id="4C2850EA0870FFA3178DCB50FE601920" authority="Nesbitt et al., 2009" authorityName="Nesbitt" authorityYear="2009" box="[129,462,1105,1129]" class="Reptilia" genus="Tawa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="hallae">
<emphasis id="B95CF77B0870FFA3178DCB50FF551920" box="[129,251,1105,1129]" italics="true" pageId="6">Tawa hallae</emphasis>
Nesbitt
<emphasis id="B95CF77B0870FFA31655CB50FE221920" box="[345,396,1105,1129]" italics="true" pageId="6">et al.</emphasis>
, 2009
</taxonomicName>
and
<taxonomicName id="4C2850EA0870FFA3150BCB50FEDE19C1" authority="Long and Murry, 1995" authorityName="Long and Murry" authorityYear="1995" family="Staurikosauridae" genus="Chindesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="bryansmalli">
<emphasis id="B95CF77B0870FFA3150BCB50FCA71920" box="[519,777,1105,1129]" italics="true" pageId="6">Chindesaurus bryansmalli</emphasis>
<bibRefCitation id="EFB956980870FFA3178DCB71FEDE19C1" author="Long R &amp; Murry P" box="[129,368,1136,1160]" pageId="6" pagination="1 - 254" refId="ref19965" refString="Long R, Murry P. Late Triassic (Carnian and Norian) Tetrapods from the Southwestern United States cover. New Mexico Museum of Natural History and Science Bulletin 1995; 4: 1 - 254." type="journal article" year="1995">Long and Murry, 1995</bibRefCitation>
</taxonomicName>
; a shallow laterodistal notch of the astragalus shared with non-massopodan sauropodomorphs, except
<taxonomicName id="4C2850EA0870FFA317DECBAEFD7D198E" authority="Martinez and Alcober, 2009" authorityName="Martinez and Alcober" authorityYear="2009" box="[210,723,1199,1223]" class="Reptilia" family="Guaibasauridae" genus="Panphagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="protos">
<emphasis id="B95CF77B0870FFA317DECBAEFE28198F" box="[210,390,1199,1223]" italics="true" pageId="6">Panphagia protos</emphasis>
<bibRefCitation id="EFB956980870FFA3169ACBAEFD7D198E" author="Martinez RN &amp; Alcober OA" box="[406,723,1199,1223]" pageId="6" refId="ref20517" refString="Martinez RN, Alcober OA. A basal sauropodomorph (Dinosauria: Saurischia) from the Ischigualasto Formation (Triassic, Carnian) and the early evolution of Sauropodomorpha. PLoS One 2009; 4: e 4397. https: // doi. org / 10.1371 / journal. pone. 0004397" type="journal volume" year="2009">Martinez and Alcober, 2009</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2850EA0870FFA3178DCBCFFE7D19AF" authority="von Huene, 1875" authorityName="von Huene" authorityYear="1875" box="[129,467,1230,1254]" class="Reptilia" family="Anchisauridae" genus="Efraasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="minor">
<emphasis id="B95CF77B0870FFA3178DCBCFFEBA19AF" box="[129,276,1230,1254]" italics="true" pageId="6">Efraasia minor</emphasis>
von Huene, 1875
</taxonomicName>
(
<emphasis id="B95CF77B0870FFA316EBCBCEFDB419AF" box="[487,538,1231,1254]" italics="true" pageId="6">sensu</emphasis>
<bibRefCitation id="EFB956980870FFA3152ECBCFFD0819AF" author="Galton PM" box="[546,678,1230,1254]" pageId="6" pagination="229 - 55" refId="ref17563" refString="Galton PM. On the anatomy and relationships of Efraasia diagnostica (Huene) n. gen., a prosauropod dinosaur (Reptilia: Saurischia) from the Upper Triassic of Germany. Palaontologische Zeitschrift 1973; 47: 229 - 55. https: // doi. org / 10.1007 / bf 02985709" type="journal article" year="1973">Galton 1973</bibRefCitation>
) and the herrerasaur
<taxonomicName id="4C2850EA0870FFA317F3CBEFFEF7184C" box="[255,345,1261,1285]" class="Reptilia" genus="Tawa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="hallae">
<emphasis id="B95CF77B0870FFA317F3CBEFFEF7184C" box="[255,345,1261,1285]" italics="true" pageId="6">T. hallae</emphasis>
</taxonomicName>
; a rounded anterior and medial margin on the proximomedial surface of the astragalus shared with the theropod
<taxonomicName id="4C2850EA0870FFA317E0CA2DFE17180D" box="[236,441,1324,1348]" genus="Eodromaeus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="murphi">
<emphasis id="B95CF77B0870FFA317E0CA2DFE17180D" box="[236,441,1324,1348]" italics="true" pageId="6">Eodromaeus murphi</emphasis>
</taxonomicName>
and sauropodomorphs, except
<taxonomicName id="4C2850EA0870FFA3178DCA4AFD9D182A" authority="Langer et al., 1999" authorityName="Langer" authorityYear="1999" box="[129,563,1355,1380]" class="Reptilia" genus="Saturnalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="tupiniquim">
<emphasis id="B95CF77B0870FFA3178DCA4AFEF1182A" box="[129,351,1355,1379]" italics="true" pageId="6">Saturnalia tupiniquim</emphasis>
<bibRefCitation id="EFB956980870FFA31665CA4DFD9D182A" author="Langer MC &amp; Abdala F &amp; Richter M" box="[361,563,1355,1380]" pageId="6" pagination="511 - 7" refId="ref19428" refString="Langer MC, Abdala F, Richter M et al. A sauropodomorph dinosaur from the Upper Triassic (Carnian) of southern Brazil. Comptes Rendus de l'Academie des Sciences - Series IIA - Earth and Planetary Science 1999; 329: 511 - 7. https: // doi. org / 10.1016 / s 1251 - 8050 (00) 80025 - 7" type="journal article" year="1999">
Langer
<emphasis id="B95CF77B0870FFA316B7CA4DFE47182A" box="[443,489,1355,1379]" italics="true" pageId="6">et al</emphasis>
., 1999
</bibRefCitation>
</taxonomicName>
and a referred specimen of
<taxonomicName id="4C2850EA0870FFA317EECA6AFDC318CA" authority="Cabreira et al., 2016" authorityName="Cabreira" authorityYear="2016" box="[226,621,1387,1411]" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="schultzi">
<emphasis id="B95CF77B0870FFA317EECA6AFE3418CA" box="[226,410,1387,1411]" italics="true" pageId="6">Buriolestes schultzi</emphasis>
<bibRefCitation id="EFB956980870FFA31693CA6AFDC318CA" author="Cabreira SF &amp; Kellner AWA &amp; Dias-da-Silva S" box="[415,621,1387,1411]" pageId="6" pagination="3090 - 5" refId="ref16346" refString="Cabreira SF, Kellner AWA, Dias-da-Silva S et al. A unique Late Triassic Dinosauromorph assemblage reveals dinosaur ancestral anatomy and diet. Current Biology 2016; 26: 3090 - 5." type="journal article" year="2016">
Cabreira
<emphasis id="B95CF77B0870FFA316F2CA6AFD8918CA" box="[510,551,1387,1411]" italics="true" pageId="6">et al</emphasis>
., 2016
</bibRefCitation>
</taxonomicName>
(
<emphasis id="B95CF77B0870FFA31572CA6AFD1F18CB" box="[638,689,1387,1410]" italics="true" pageId="6">sensu</emphasis>
<bibRefCitation id="EFB956980870FFA315B4CA6AFF7C18EB" author="Moro D &amp; Damke LVS &amp; Muller RT" pageId="6" pagination="1025 - 59" refId="ref21003" refString="Moro D, Damke LVS, Muller RT et al. An unusually robust specimen attributed to Buriolestes schultzi (Dinosauria: Sauropodomorpha) from the Late Triassic of Southern Brazil. Anatomical Record (Hoboken, N. J.: 2007) 2024; 307: 1025 - 59. https: // doi. org / 10.1002 / ar. 25319" type="journal article" year="2024">
Moro
<emphasis id="B95CF77B0870FFA315F4CA6AFF3718EB" italics="true" pageId="6">et al.</emphasis>
2024
</bibRefCitation>
); a posterolateral ridge between the anterior ascending process and the posterolateral process that is always taller than the posterolateral process shared with the herrerasaurids
<taxonomicName id="4C2850EA0870FFA3178DCAE8FD981B49" authority="Reig, 1963" authorityName="Reig" authorityYear="1963" box="[129,566,1512,1536]" class="Reptilia" family="Herrerasauridae" genus="Herrerasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="ischgualastensis">
<emphasis id="B95CF77B0870FFA3178DCAE8FE1E1B49" box="[129,432,1512,1536]" italics="true" pageId="6">Herrerasaurus ischgualastensis</emphasis>
<bibRefCitation id="EFB956980870FFA316B3CAE9FD981B49" author="Reig OA" box="[447,566,1512,1536]" pageId="6" pagination="3 - 20" refId="ref22496" refString="Reig OA. La presencia de dinosaurios saurisquios en los ' Estratos de Ischigualasto' (Mesotriasico Superior) de las provincias de San Juan y La Rioja (Republica Argentina). Ameghiniana 1963; 3: 3 - 20." type="journal article" year="1963">Reig, 1963</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2850EA0870FFA31575CAE9FDA51B56" authority="Alcober and Martinez, 2010" authorityName="Alcober and Martinez" authorityYear="2010" family="Herrerasauridae" genus="Sanjuansaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="gordilloi">
<emphasis id="B95CF77B0870FFA31575CAE9FF7F1B56" italics="true" pageId="6">Sanjuansaurus gordilloi</emphasis>
<bibRefCitation id="EFB956980870FFA317D7C906FDA51B56" author="Alcober O &amp; Martinez R" box="[219,523,1543,1567]" pageId="6" pagination="55 - 81" refId="ref15691" refString="Alcober O, Martinez R. A new herrerasaurid (Dinosauria, Saurischia) from the Upper Triassic Ischigualasto Formation of Northwestern Argentina. ZooKeys 2010; 63: 55 - 81." type="journal article" year="2010">Alcober and Martínez, 2010</bibRefCitation>
</taxonomicName>
and sauropodomorphs, except
<taxonomicName id="4C2850EA0870FFA317D9C926FD221B76" authority="Leal et al., 2004" authorityName="Leal" authorityYear="2004" box="[213,652,1575,1599]" class="Reptilia" family="Plateosauridae" genus="Unaysaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="tolentinoi">
<emphasis id="B95CF77B0870FFA317D9C926FE181B76" box="[213,438,1575,1599]" italics="true" pageId="6">Unaysaurus tolentinoi</emphasis>
<bibRefCitation id="EFB956980870FFA316C4C926FD221B76" author="Leal LA &amp; Azevedo SK &amp; Kellner AWA" box="[456,652,1575,1599]" pageId="6" pagination="1 - 24" refId="ref19715" refString="Leal LA, Azevedo SK, Kellner AWA et al. A new early dinosaur (Sauropodomorpha) from the Caturrita Formation (Late Triassic), Paran Basin, Brazil. Zootaxa 2004; 690: 1 - 24." type="journal article" year="2004">
Leal
<emphasis id="B95CF77B0870FFA31509C926FD951B76" box="[517,571,1575,1599]" italics="true" pageId="6">et al</emphasis>
., 2004
</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C2850EA0870FFA315A8C926FEA11B17" authorityName="Lovelace &amp; Kufner &amp; Fitch &amp; Rogers &amp; Schmitz &amp; Schwartz &amp; LeClair-Diaz &amp; St.Clair &amp; Mann &amp; Teran" authorityYear="2025" genus="Ahvaytum" pageId="6" rank="species" species="bahndooiveche">
<emphasis id="B95CF77B0870FFA315A8C926FEA11B17" italics="true" pageId="6">Ahvaytum bahndooiveche</emphasis>
</taxonomicName>
is differentiated from all other sauropodomorphs with comparable material by the following autapomorphy: fossa on the medial surface of the astragalus that is open distally.
<taxonomicName id="4C2850EA0870FFA3178DC9A5FED51BF5" authorityName="Lovelace &amp; Kufner &amp; Fitch &amp; Rogers &amp; Schmitz &amp; Schwartz &amp; LeClair-Diaz &amp; St.Clair &amp; Mann &amp; Teran" authorityYear="2025" box="[129,379,1700,1724]" genus="Ahvaytum" pageId="6" rank="species" species="bahndooiveche">
<emphasis id="B95CF77B0870FFA3178DC9A5FED51BF5" box="[129,379,1700,1724]" italics="true" pageId="6">Ahvaytum bahndooiveche</emphasis>
</taxonomicName>
is differentiated from
<taxonomicName id="4C2850EA0870FFA3156DC9A4FEE71B92" authority="Sereno et al., 1993" authorityName="Sereno" authorityYear="1993" class="Reptilia" genus="Eoraptor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="lunensis">
<emphasis id="B95CF77B0870FFA3156DC9A4FCA41BF5" box="[609,778,1700,1724]" italics="true" pageId="6">Eoraptor lunensis</emphasis>
<bibRefCitation id="EFB956980870FFA3178DC9C2FEE71B92" author="Sereno PC &amp; Forster CA &amp; Rogers RR" box="[129,329,1731,1755]" pageId="6" pagination="64 - 6" refId="ref22876" refString="Sereno PC, Forster CA, Rogers RR et al. Primitive dinosaur skeleton from Argentina and the early evolution of Dinosauria. Nature 1993; 361: 64 - 6. https: // doi. org / 10.1038 / 361064 a 0" type="journal article" year="1993">
Sereno
<emphasis id="B95CF77B0870FFA317DEC9C5FF511B92" box="[210,255,1731,1755]" italics="true" pageId="6">et al</emphasis>
., 1993
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2850EA0870FFA31655C9C2FD471B92" authority="Griffin et al., 2022" authorityName="Griffin" authorityYear="2022" box="[345,745,1731,1755]" genus="Mbiresaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="raathi">
<emphasis id="B95CF77B0870FFA31655C9C2FDB61B92" box="[345,536,1731,1755]" italics="true" pageId="6">Mbiresaurus raathi</emphasis>
<bibRefCitation id="EFB956980870FFA3152EC9C2FD471B92" author="Griffin CT &amp; Wynd BM &amp; Munyikwa D" box="[546,745,1731,1755]" pageId="6" pagination="313 - 9" refId="ref17996" refString="Griffin CT, Wynd BM, Munyikwa D et al. Africa's oldest dinosaurs reveal early suppression of dinosaur distribution. Nature 2022; 609: 313 - 9. https: // doi. org / 10.1038 / s 41586 - 022 - 05133 - x" type="journal article" year="2022">
Griffin
<emphasis id="B95CF77B0870FFA3157EC9C5FD311B92" box="[626,671,1731,1755]" italics="true" pageId="6">et al</emphasis>
., 2022
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2850EA0870FFA315F5C9C5FF121BB3" authorityName="Martinez and Alcober" authorityYear="2009" class="Reptilia" family="Guaibasauridae" genus="Panphagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="protos">
<emphasis id="B95CF77B0870FFA315F5C9C5FF121BB3" italics="true" pageId="6">P. protos</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2850EA0870FFA317C4C9E2FEE31BB3" box="[200,333,1763,1786]" class="Reptilia" genus="Saturnalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="tupiniquim">
<emphasis id="B95CF77B0870FFA317C4C9E2FEE31BB3" box="[200,333,1763,1786]" italics="true" pageId="6">S. tupiniquim</emphasis>
</taxonomicName>
, and a referred specimen of
<emphasis id="B95CF77B0870FFA3157AC9E2FD571BB2" box="[630,761,1763,1787]" italics="true" pageId="6">
<taxonomicName id="4C2850EA0870FFA3157AC9E2FD751BB2" box="[630,731,1763,1787]" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="schultzi">B. schultzi</taxonomicName>
by
</emphasis>
a proportionally wider astragalus (max. anteroposterior length/ max. transverse width:
<taxonomicName id="4C2850EA0870FFA31687C820FE5F1A70" authorityName="Lovelace &amp; Kufner &amp; Fitch &amp; Rogers &amp; Schmitz &amp; Schwartz &amp; LeClair-Diaz &amp; St.Clair &amp; Mann &amp; Teran" authorityYear="2025" box="[395,497,1825,1849]" genus="Ahvaytum" pageId="6" rank="genus">
<emphasis id="B95CF77B0870FFA31687C820FE5F1A70" box="[395,497,1825,1849]" italics="true" pageId="6">Ahvaytum</emphasis>
</taxonomicName>
= 0.53,
<taxonomicName id="4C2850EA0870FFA3155DC820FD161A70" box="[593,696,1825,1849]" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="genus">
<emphasis id="B95CF77B0870FFA3155DC820FD161A70" box="[593,696,1825,1849]" italics="true" pageId="6">Buriolestes</emphasis>
</taxonomicName>
<emphasis id="B95CF77B0870FFA315B2C820FD7F1A70" bold="true" box="[702,721,1825,1849]" pageId="6"></emphasis>
0.71,
<taxonomicName id="4C2850EA0870FFA3178EC840FF771A11" box="[130,217,1857,1880]" class="Reptilia" genus="Eoraptor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="genus">
<emphasis id="B95CF77B0870FFA3178EC840FF771A11" box="[130,217,1857,1880]" italics="true" pageId="6">Eoraptor</emphasis>
</taxonomicName>
<emphasis id="B95CF77B0870FFA317D2C840FF5F1A10" bold="true" box="[222,241,1857,1881]" pageId="6"></emphasis>
0.63,
<taxonomicName id="4C2850EA0870FFA31630C840FE181A10" box="[316,438,1857,1881]" genus="Mbiresaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="genus">
<emphasis id="B95CF77B0870FFA31630C840FE181A10" box="[316,438,1857,1881]" italics="true" pageId="6">Mbiresaurus</emphasis>
</taxonomicName>
<emphasis id="B95CF77B0870FFA316B7C840FE601A10" bold="true" box="[443,462,1857,1881]" pageId="6"></emphasis>
0.64,
<taxonomicName id="4C2850EA0870FFA31515C840FD2C1A10" authorityName="Martinez &amp; Alcober" authorityYear="2009" box="[537,642,1857,1881]" class="Reptilia" family="Guaibasauridae" genus="Panphagia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="genus">
<emphasis id="B95CF77B0870FFA31515C840FD2C1A10" box="[537,642,1857,1881]" italics="true" pageId="6">Panphagia</emphasis>
</taxonomicName>
<emphasis id="B95CF77B0870FFA3158BC840FD341A10" bold="true" box="[647,666,1857,1881]" pageId="6"></emphasis>
0.88, and
<taxonomicName id="4C2850EA0870FFA3178EC861FF471A31" authorityName="Langer, Abdala, Richter &amp; Benton" authorityYear="1999" box="[130,233,1888,1912]" class="Reptilia" genus="Saturnalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="genus">
<emphasis id="B95CF77B0870FFA3178EC861FF471A31" box="[130,233,1888,1912]" italics="true" pageId="6">Saturnalia</emphasis>
</taxonomicName>
<emphasis id="B95CF77B0870FFA317E3C861FEAC1A31" bold="true" box="[239,258,1888,1912]" pageId="6"></emphasis>
0.73) andbyarelativelywidefibularfacetoccupying more than a quarter of the transverse width of the astragalus. Further differentiated from
<taxonomicName id="4C2850EA0870FFA31696C89EFDEC1AFE" box="[410,578,1951,1975]" class="Reptilia" genus="Eoraptor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="lunensis">
<emphasis id="B95CF77B0870FFA31696C89EFDEC1AFE" box="[410,578,1951,1975]" italics="true" pageId="6">Eoraptor lunensis</emphasis>
</taxonomicName>
by a posteromedial angle on the astragalus. Further differentiated from a referred specimen of
<taxonomicName id="4C2850EA0870FFA314CECFB1FAAE1D8E" authority="(Moro et al., 2024)" baseAuthorityName="Moro" baseAuthorityYear="2024" box="[962,1280,175,199]" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="schultzi">
<emphasis id="B95CF77B0870FFA314CECFB1FB871D8E" box="[962,1065,175,199]" italics="true" pageId="6">B. schultzi</emphasis>
(
<bibRefCitation id="EFB956980870FFA31336CFAEFB571D8E" author="Moro D &amp; Damke LVS &amp; Muller RT" box="[1082,1273,175,199]" pageId="6" pagination="1025 - 59" refId="ref21003" refString="Moro D, Damke LVS, Muller RT et al. An unusually robust specimen attributed to Buriolestes schultzi (Dinosauria: Sauropodomorpha) from the Late Triassic of Southern Brazil. Anatomical Record (Hoboken, N. J.: 2007) 2024; 307: 1025 - 59. https: // doi. org / 10.1002 / ar. 25319" type="journal article" year="2024">
Moro
<emphasis id="B95CF77B0870FFA3138DCFB1FB1D1D8E" box="[1153,1203,175,199]" italics="true" pageId="6">et al.</emphasis>
, 2024
</bibRefCitation>
)
</taxonomicName>
by a poorly proximally expanded posteromedial process (= pyramidal process of some authors) and by a relatively tall posterolateral process (= posterior ascending process of some authors). Further differentiated from both
<taxonomicName id="4C2850EA0870FFA3130BCE2CFBC21C0C" box="[1031,1132,301,325]" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="schultzi">
<emphasis id="B95CF77B0870FFA3130BCE2CFBC21C0C" box="[1031,1132,301,325]" italics="true" pageId="6">B. schultzi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2850EA0870FFA313ADCE2CFB511C0C" box="[1185,1279,301,325]" genus="Mbiresaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="raathi">
<emphasis id="B95CF77B0870FFA313ADCE2CFB511C0C" box="[1185,1279,301,325]" italics="true" pageId="6">M. raathi</emphasis>
</taxonomicName>
by a posterolateral process that is level with the posterior margin of the astragalus in proximal view.
</paragraph>
<paragraph id="8B972B690870FFA31435CEA4FB101FD1" blockId="6.[825,1475,421,664]" pageId="6">
<emphasis id="B95CF77B0870FFA31435CEA4FC391CF4" box="[825,919,421,445]" italics="true" pageId="6">Remarks:</emphasis>
The proximal end of a silesaurid femur (UWGM 7407) and the distal end of a probable silesaurid humerus (UWGM 7550; see below) were recovered from the study area precluding the referral of 15 non-diagnostic dinosauromorph elements (UWGM 7434; Supporting Information,
<figureCitation id="131337EC0870FFA31256CD23FA0A1F73" box="[1370,1444,546,570]" captionStart="Figure 1" captionStartId="2.[129,194,1602,1626]" captionTargetBox="[134,1470,147,1571]" captionTargetId="figure-6@2.[129,1473,144,1574]" captionTargetPageId="2" captionText="Figure 1. A palaeogeographic map (A) of Triassic outcrops shows the position of key localities (1 to 9); Chugwater Group, Wyoming, USA (inset), local geology surrounding Garretts Surprise (star). Chronostratigraphic context of early dinosaur-bearing units (B) shows the relative stratigraphic positions for first occurrences of dinosaurian body and trace fossils tied to a generalized stratigraphic section (C) of the upper Chugwater Group [after Lovelace et al. (2024)]. The shaded box highlights the stratigraphic position of UWGM 1975 and UWGM 7407 within the Popo Agie Formation. Localities:1, Argentina; 2, Brazil; 3, Zimbabwe; 4, India; 5, Germany; 6, Britain; 7, Morocco; 8, Texas, USA (Dockum Group); 9, south-western USA (Chinle/Moenkopi formations); 10, Wyoming, USA (Chugwater Group). Silhouettes (not to scale) on outcrop demonstrate faunal distribution within the Popo Agie and Jelm formations.At, Atreipus sp.; AtGr, AtreipusGrallator plexus; An, Anaschisma browni; Be, Beesiiwo cooowuse; Bu, Buettnererpeton bakeri; Br, Brachychirotherium sp.; Eu, Eubrachiosaurus browni; He, Heptasuchus clarki; Pa, Parasuchus bransoni; Po, Poposaurus gracilis; Ni, Ninumbeehan dookoodukah. Artist credits for silhouettes from Phylopic.org and abbreviated formation names provided in the Supporting Information, Text S1. Previously published dinosaur-bearing localities recovered from PBDB (PaleobioDB.org; accessed Feb, 2024; Supporting Information, Data S1)." figureDoi="http://doi.org/10.5281/zenodo.14827188" httpUri="https://zenodo.org/record/14827188/files/figure.png" pageId="6">
Fig. S
<quantity id="4CD0868C0870FFA31296CD23FA0A1F73" box="[1434,1444,546,570]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" unit="in" value="1.0">1</quantity>
</figureCitation>
in Text S1) to higher taxa. Additional vertebrate material was surface collected from the Garretts Surprise locality and identified as non-ornithodiran or indeterminate.
</paragraph>
<paragraph id="8B972B690870FFA21435CDBBFEB91D8E" blockId="6.[824,1476,697,1975]" lastBlockId="7.[113,762,144,199]" lastPageId="7" pageId="6">
<emphasis id="B95CF77B0870FFA31435CDBBFC1E1F98" box="[825,944,698,721]" italics="true" pageId="6">Description:</emphasis>
The
<typeStatus id="549395CB0870FFA314F7CDBBFB871F9B" box="[1019,1065,698,722]" pageId="6">type</typeStatus>
specimen of
<taxonomicName id="4C2850EA0870FFA313CECDB8FC4C1FB9" authority="(UWGM 1975)" baseAuthorityName="UWGM" baseAuthorityYear="1975" genus="Ahvaytum" pageId="6" rank="species" species="bahndooiveche">
<emphasis id="B95CF77B0870FFA313CECDB8FA6C1F98" box="[1218,1474,697,721]" italics="true" pageId="6">Ahvaytum bahndooiveche</emphasis>
(UWGM 1975)
</taxonomicName>
is an isolated left astragalus (
<figureCitation id="131337EC0870FFA31216CDD8FAF81FB8" box="[1306,1366,729,753]" captionStart="Figure 2" captionStartId="4.[129,194,885,909]" captionTargetBox="[131,1471,146,855]" captionTargetId="figure-419@4.[129,1473,144,857]" captionTargetPageId="4" captionText="Figure 2. Holotype left astragalus of Ahvaytum bahndooiveche gen.et sp. nov.(UWGM 1975). 3D model in (A) medial, (B) lateral, (C) lateral transparent, (D) posterior transparent, (E) distal, (F) proximal, (I) anterior, and (J) posterior orthographic views. Photographs in (G) proximal and (H) distal views. Abbreviations: amc, anteromedial corner; ap, ascending process of the astragalus; g, groove; f, foramen; ff, fibular facet; ldn, laterodistal notch (= lateroventral depression); mf, medial fossa; nf, non-articular fossa (= dorsal basin, = semi-elliptical fossa); p, platform; plp, posterolateral process; plr, posterolateral ridge; tf, tibial facet. Diagonal lines indicate broken surfaces.Arrows indicate anterior direction. Scale bar = 1 cm. STL models archived online (morphosource.org ID: 000607569)." figureDoi="http://doi.org/10.5281/zenodo.14827190" httpUri="https://zenodo.org/record/14827190/files/figure.png" pageId="6">Fig. 2</figureCitation>
). A calcaneum was not recovered and there is no evidence of fusion of these elements. The transverse width of the astragalus is nearly twice its maximum anteroposterior length. The proximal surface is slightly weathered, but the rest of the surface of the element is essentially intact. There is a transverse groove on the anterior surface of the body of the astragalus, which is widespread among dinosauromorphs. The ascending process is separated from the anterior surface by a platform and broken at approximately mid-height. A foramen on the anterior surface of the ascending process extends posterodistally into the body of the astragalus. A posterolateral ridge extends from the ascending process to meet a posterolateral process [= posterior ascending process of
<bibRefCitation id="EFB956980870FFA31458CB71FBC319C1" author="Sereno PC &amp; Arcucci AB" box="[852,1133,1136,1160]" pageId="6" pagination="385 - 99" refId="ref22836" refString="Sereno PC, Arcucci AB. Dinosaurian precursors from the Middle Triassic of Argentina: Lagerpeton chanarensis. Journal of Vertebrate Paleontology 1994; 13: 385 - 99. https: // doi. org / 10.1080 / 02724634.1994.10011522" type="journal article" year="1994">Sereno and Arcucci (1994)</bibRefCitation>
] along the posterior margin. The anteromedial corner is acute as in other early dinosaurs (e.g.
<bibRefCitation id="EFB956980870FFA31435CBAEFC15198E" author="Muller RT" box="[825,955,1199,1223]" pageId="6" pagination="2534 - 41" refId="ref21067" refString="Muller RT. Astragalar anatomy of an early dinosaur from the Upper Triassic of southern Brazil. Historical Biology 2021; 33: 2534 - 41. https: // doi. org / 10.1080 / 08912963.2020.1814276" type="journal article" year="2021">Müller 2021</bibRefCitation>
) and projects well beyond the anterolateral condyle. The tibial articular facet is separated from the anterior and medial margins by a wide, rounded lip and does not extend on to the anteromedial corner, which is identical to the condition in several sauropodomorphs (e.g.
<bibRefCitation id="EFB956980870FFA3137ECA2DFB41180D" author="Muller RT" box="[1138,1263,1324,1348]" pageId="6" pagination="2534 - 41" refId="ref21067" refString="Muller RT. Astragalar anatomy of an early dinosaur from the Upper Triassic of southern Brazil. Historical Biology 2021; 33: 2534 - 41. https: // doi. org / 10.1080 / 08912963.2020.1814276" type="journal article" year="2021">Müller 2021</bibRefCitation>
). The medial surface of the astragalus bears a fossa that opens toward the distal surface, which appears to be autapomorphic among dinosaurs. The posteromedial margin forms a corner, and the proximal margin here is not raised into a pyramidal process such as that found in theropods (e.g.
<bibRefCitation id="EFB956980870FFA314D0CAC8FB4C18A9" author="Nesbitt SJ &amp; Ezcurra MD" box="[988,1250,1480,1505]" pageId="6" pagination="513 - 26" refId="ref21377" refString="Nesbitt SJ, Ezcurra MD. The early fossil record of dinosaurs in North America: a new neotheropod from the base of the Upper Triassic Dockum group of Texas. Acta Palaeontologica Polonica 2015; 60: 513 - 26." type="journal article" year="2015">Nesbitt and Ezcurra 2015</bibRefCitation>
) or the elevated crest in unaysaurids (e.g.
<bibRefCitation id="EFB956980870FFA3130ACAE9FB2A1B49" author="Muller RT" box="[1030,1156,1512,1536]" pageId="6" pagination="2534 - 41" refId="ref21067" refString="Muller RT. Astragalar anatomy of an early dinosaur from the Upper Triassic of southern Brazil. Historical Biology 2021; 33: 2534 - 41. https: // doi. org / 10.1080 / 08912963.2020.1814276" type="journal article" year="2021">Müller 2021</bibRefCitation>
,
<bibRefCitation id="EFB956980870FFA3139CCAE9FAE01B49" author="Ezcurra MD &amp; Muller RT &amp; Novas FE" box="[1168,1358,1512,1536]" pageId="6" pagination="1093 - 112" refId="ref17193" refString="Ezcurra MD, Muller RT, Novas FE et al. Osteology of the sauropodomorph dinosaur Jaklapallisaurus asymmetricus from the Late Triassic of central India. The Anatomical Record 2023; 307: 1093 - 112. https: // doi. org / 10.1002 / ar. 25359" type="journal article" year="2023">
Ezcurra
<emphasis id="B95CF77B0870FFA313E9CAE8FABA1B49" box="[1253,1300,1512,1536]" italics="true" pageId="6">et al.</emphasis>
2023
</bibRefCitation>
). Posterior to the ascending process there is a non-articular fossa (= dorsal basin, = semi-elliptical fossa) with a weathered but raised rim along its medial margin. Within this fossa there is a single oblong vascular foramen whose canal is short and anteriorly directed at the base of the ascending process (
<figureCitation id="131337EC0870FFA31206C984FA2F1BD4" box="[1290,1409,1669,1693]" captionStart="Figure 2" captionStartId="4.[129,194,885,909]" captionTargetBox="[131,1471,146,855]" captionTargetId="figure-419@4.[129,1473,144,857]" captionTargetPageId="4" captionText="Figure 2. Holotype left astragalus of Ahvaytum bahndooiveche gen.et sp. nov.(UWGM 1975). 3D model in (A) medial, (B) lateral, (C) lateral transparent, (D) posterior transparent, (E) distal, (F) proximal, (I) anterior, and (J) posterior orthographic views. Photographs in (G) proximal and (H) distal views. Abbreviations: amc, anteromedial corner; ap, ascending process of the astragalus; g, groove; f, foramen; ff, fibular facet; ldn, laterodistal notch (= lateroventral depression); mf, medial fossa; nf, non-articular fossa (= dorsal basin, = semi-elliptical fossa); p, platform; plp, posterolateral process; plr, posterolateral ridge; tf, tibial facet. Diagonal lines indicate broken surfaces.Arrows indicate anterior direction. Scale bar = 1 cm. STL models archived online (morphosource.org ID: 000607569)." figureDoi="http://doi.org/10.5281/zenodo.14827190" httpUri="https://zenodo.org/record/14827190/files/figure.png" pageId="6">Figs. 2CD</figureCitation>
). The fibular facet occupies just over a quarter of the transverse width of the astragalus and is delineated by a sharp medial margin. A laterodistal notch (= lateroventral depression) is present on the distal surface, presumably for reception of a medial process of the calcaneum, as is the case in
<taxonomicName id="4C2850EA0870FFA31388C823FCD71A10" authority="(Sereno et al., 2012)" baseAuthorityName="Sereno" baseAuthorityYear="2012" class="Reptilia" genus="Eoraptor" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" phylum="Chordata" rank="species" species="lunensis">
<emphasis id="B95CF77B0870FFA31388C823FA801A70" box="[1156,1326,1825,1849]" italics="true" pageId="6">Eoraptor lunensis</emphasis>
(
<bibRefCitation id="EFB956980870FFA31232C820FCC31A10" author="Sereno PC &amp; Martinez RN &amp; Alcober OA" pageId="6" pagination="83 - 179" refId="ref22920" refString="Sereno PC, Martinez RN, Alcober OA. Osteology of Eoraptor lunensis (Dinosauria, Sauropodomorpha). Journal of Vertebrate Paleontology 2012; 32: 83 - 179. https: // doi. org / 10.1080 / 02724634.2013.820113" type="journal article" year="2012">
Sereno
<emphasis id="B95CF77B0870FFA31280C823FA191A70" box="[1420,1463,1825,1849]" italics="true" pageId="6">et al</emphasis>
., 2012
</bibRefCitation>
)
</taxonomicName>
. An incipient protuberance is present on the distolateral surface of the astragalus just anterior to the laterodistal notch; however, unlike the condition in the unaysaurids
<taxonomicName id="4C2850EA0870FFA31229C87EFBD21AFE" authority="Novas et al., 2010" authorityName="Novas" authorityYear="2010" genus="Jaklapallisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" phylum="Chordata" rank="species" species="asymmetricus">
<emphasis id="B95CF77B0870FFA31229C87EFC131AFF" italics="true" pageId="6">Jaklapallisaurus asymmetricus</emphasis>
<bibRefCitation id="EFB956980870FFA314C8C89EFBD21AFE" author="Novas FE &amp; Ezcurra MD &amp; Chatterjee S" box="[964,1148,1951,1975]" pageId="6" pagination="333 - 49" refId="ref21760" refString="Novas FE, Ezcurra MD, Chatterjee S et al. New dinosaur species from the Upper Triassic Upper Maleri and Lower Dharmaram formations of Central India. Earth and Environmental Science Transactions of The Royal Society of Edinburgh 2010; 101: 333 - 49. https: // doi. org / 10.1017 / s 1755691011020093" type="journal article" year="2010">
Novas
<emphasis id="B95CF77B0870FFA31307C89EFB9B1AFE" box="[1035,1077,1951,1975]" italics="true" pageId="6">et al</emphasis>
., 2010
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2850EA0870FFA213BDC89EFF701DE1" authority="Muller et al., 2018" authorityName="Muller" authorityYear="2018" genus="Macrocollum" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="7" pageId="6" phylum="Chordata" rank="species" species="itaquii">
<emphasis id="B95CF77B0870FFA313BDC89EFAD61AFF" box="[1201,1400,1951,1975]" italics="true" pageId="6">Macrocollum itaquii</emphasis>
<bibRefCitation id="EFB956980870FFA21271C89EFF701DE1" author="Muller RT &amp; Langer MC &amp; Dias-da-Silva S" lastPageId="7" pageId="6" refId="ref21291" refString="Muller RT, Langer MC, Dias-da-Silva S. An exceptionally preserved association of complete dinosaur skeletons reveals the oldest long-necked sauropodomorphs. Biology Letters 2018; 14: 20180633. https: // doi. org / 10.1098 / rsbl. 2018.0633" type="journal volume" year="2018">
Müller
<emphasis id="B95CF77B0871FFA2177DCF90FF341DE1" box="[113,154,144,168]" italics="true" pageId="7">et al</emphasis>
., 2018
</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFB956980871FFA217E2CF91FE061DE1" author="Ezcurra MD &amp; Muller RT &amp; Novas FE" box="[238,424,144,168]" pageId="7" pagination="1093 - 112" refId="ref17193" refString="Ezcurra MD, Muller RT, Novas FE et al. Osteology of the sauropodomorph dinosaur Jaklapallisaurus asymmetricus from the Late Triassic of central India. The Anatomical Record 2023; 307: 1093 - 112. https: // doi. org / 10.1002 / ar. 25359" type="journal article" year="2023">
Ezcurra
<emphasis id="B95CF77B0871FFA2164ECF90FEDE1DE1" box="[322,368,144,168]" italics="true" pageId="7">et al.</emphasis>
2023
</bibRefCitation>
: fig. 3eg) it does not form a distinct projection.
</paragraph>
<paragraph id="8B972B690871FFAD177DCFEEFD351F48" blockId="7.[113,764,239,545]" lastBlockId="8.[127,778,144,513]" lastPageId="8" pageId="7">
<emphasis id="B95CF77B0871FFA2177DCFEEFE891C4E" box="[113,295,239,263]" italics="true" pageId="7">Referred material:</emphasis>
The proximal end of a left femur (UWGM 7549) possesses several dinosaurian and saurischian features but is too incomplete for a referral beyond
<taxonomicName id="4C2850EA0871FFA21546CE2CFD1A1C0C" authorityName="Seeley" authorityYear="1887" box="[586,692,301,325]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="7" phylum="Chordata" rank="order">Saurischia</taxonomicName>
on its own (
<figureCitation id="131337EC0871FFA217B8CE4CFF5F1C2C" box="[180,241,333,357]" captionStart="Figure 3" captionStartId="5.[113,178,1842,1866]" captionTargetBox="[149,1426,449,1812]" captionTargetId="figure-133@5.[145,1428,447,1814]" captionTargetPageId="5" captionText="Figure 3. Proximal end of a left femur UWGM 7549 (AE) referred to Ahvaytum bahndooiveche gen. et sp. nov.). 3D model in (A) anteromedial, (B) posterolateral, (C) proximal, (D) anterolateral, and (E) posteromedial orthographic views. Abbreviations: alt, anterolateral tuber; amt, anteromedial tuber; ce, concave emargination; dlt, dorsolateral trochanter; ft, fossa trochanterica (= facies articularis antitrochanterica); gt, greater trochanter; pmt, posteromedial tuber; ve, ventral emargination.Arrows indicate anterior direction. Scale bar = 1 cm. STL models archived online (morphosource.org ID: 000607572)." figureDoi="http://doi.org/10.5281/zenodo.14827194" httpUri="https://zenodo.org/record/14827194/files/figure.png" pageId="7">Fig. 3</figureCitation>
). The proximal surface of the femur appears to be slightly abraded revealing trabecular bone and hindering the identification of a transverse groove, such as that seen in UWGM 7407 (
<figureCitation id="131337EC0871FFA2161FCEAAFEF51C8A" box="[275,347,427,451]" captionStart="Figure 4" captionStartId="7.[113,178,1814,1838]" captionTargetBox="[116,1456,592,1784]" captionTargetId="figure-257@7.[114,1458,590,1786]" captionTargetPageId="7" captionText="Figure 4. Photographs of sulcimentisaurian silesaurid elements from the Garretts Surprise locality.Distal end of a left humerus UWGM 7550 (AE) in (A) anterior, (B) posterior, (C) medial, (D) lateral, (E) and distal views. Proximal end of a right femur UWGM 7407 (FJ) in (F) proximal, (G) anterolateral, (H) posteromedial, (I) posterolateral, and (J) anteromedial views.Abbreviations:alt, anterolateral tuber; amt, anteromedial tuber; at, anterior trochanter; dlt, dorsolateral trochanter; ect, ectepicondyle; ent, entepicondyle; g, groove; gt, greater trochanter; ipmt, incipient posteromedial tuber; n, notch; rc, radial condyle; uc, ulnar condyle.Arrows point in the anterior direction.Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14827200" httpUri="https://zenodo.org/record/14827200/files/figure.png" pageId="7">Fig. 4F</figureCitation>
) and widespread among early-diverging sauropodomorphs and dinosauromorphs more generally. There are also several cracks present in this element, one of which passes through the position that would be occupied by a ligament sulcus between the anteromedial and posteromedial tubera. The posteromedial tuber is small and rounded, and the larger anteromedial tuber is also rounded. The anterolateral tuber forms a broad, rounded profile along the anterolateral surface of the femur in proximal view. The head of the femur is offset from the shaft resulting in a concave emargination just ventral to the head, common among all dinosaurs. On the anterolateral surface of the femur, a ridge extends from the ventral margin of the head then merges distally into the shaft forming a ventral emargination, such as that seen in
<taxonomicName id="4C2850EA0871FFA213DECEABFA031C8B" box="[1234,1453,426,450]" class="Reptilia" genus="Saturnalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="7" phylum="Chordata" rank="species" species="tupiniquim">
<emphasis id="B95CF77B0871FFA213DECEABFA031C8B" box="[1234,1453,426,450]" italics="true" pageId="7">Saturnalia tupiniquim</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2850EA0871FFA21426CEC8FA9E1CA8" authority="Marsola et al., 2018" authorityName="Marsola" authorityYear="2018" box="[810,1328,457,481]" genus="Nhandumirim" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" phylum="Chordata" rank="species" species="waldsangae">
<emphasis id="B95CF77B0871FFA21426CEC8FB941CA8" box="[810,1082,457,481]" italics="true" pageId="7">Nhandumirim waldsangae</emphasis>
<bibRefCitation id="EFB956980871FFA21346CEC8FA9E1CA8" author="Marsola JCA &amp; Bittencourt JS &amp; Butler RJ" box="[1098,1328,457,481]" pageId="7" refId="ref20468" refString="Marsola JCA, Bittencourt JS, Butler RJ et al. A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, south Brazil. Journal of Vertebrate Paleontology 2018; 38: e 1531878. https: // doi. org / 10.1080 / 02724634.2018.1531878" type="journal volume" year="2018">
Marsola
<emphasis id="B95CF77B0871FFA213A1CECBFB4E1CA8" box="[1197,1248,457,481]" italics="true" pageId="7">et al</emphasis>
., 2018
</bibRefCitation>
</taxonomicName>
, and some coelophysoids (
<bibRefCitation id="EFB956980871FFA214DCCEE8FB231F48" author="Kirmse JPS &amp; Benton MJ &amp; Hildebrandt C" box="[976,1165,489,513]" pageId="7" pagination="562 - 72" refId="ref18962" refString="Kirmse JPS, Benton MJ, Hildebrandt C et al. A Coelophysoidea (Dinosauria, Theropoda) femur from the Tytherington fissures (Rhaetian, Late Triassic), Bristol, UK. Proceedings of the Geologists' Association 2023; 134: 562 - 72." type="journal article" year="2023">
Kirmse
<emphasis id="B95CF77B0871FFA2132ECEE8FBFD1F48" box="[1058,1107,489,513]" italics="true" pageId="7">et al.</emphasis>
2023
</bibRefCitation>
), but unknown among core ornithischians, exclusive of silesaurids. The shaft of the femur is broken approximately just above the position of the anterior trochanter. The dorsolateral trochanter is present as a rounded rugosity along the lateral side of the femoral shaft and probably missing its distal extent due to the break. The greater trochanter is prominent and squared off with a straight margin between it and the head of the femur in anterior/posterior view similar to
<taxonomicName id="4C2850EA087EFFAD178DCE4CFDF81C2D" authority="(Novas, 1994)" baseAuthorityName="Novas" baseAuthorityYear="1994" box="[129,598,332,356]" class="Reptilia" family="Herrerasauridae" genus="Herrerasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="8" phylum="Chordata" rank="species" species="ischigualastensis">
<emphasis id="B95CF77B087EFFAD178DCE4CFE181C2D" box="[129,438,332,356]" italics="true" pageId="8">Herrerasaurus ischigualastensis</emphasis>
(
<bibRefCitation id="EFB95698087EFFAD16C4CE4DFDE51C2D" author="Novas FE" box="[456,587,332,356]" pageId="8" pagination="400 - 23" refId="ref21705" refString="Novas FE. New information on the systematics and postcranial skeleton of Herrerasaurus ischigualastensis (Theropoda: Herrerasauridae) from the Ischigualasto Formation (Upper Triassic) of Argentina. Journal of Vertebrate Paleontology 1994; 13: 400 - 23. https: // doi. org / 10.1080 / 0 2724634.1994. 10011523" type="journal article" year="1994">Novas, 1994</bibRefCitation>
)
</taxonomicName>
and some specimens of
<taxonomicName id="4C2850EA087EFFAD17D3CE6AFD771CCA" authority="Cabreira et al., 2011" authorityName="Cabreira" authorityYear="2011" box="[223,729,363,387]" genus="Pampadromaeus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" phylum="Chordata" rank="species" species="barberenai">
<emphasis id="B95CF77B087EFFAD17D3CE6AFE591CCA" box="[223,503,363,387]" italics="true" pageId="8">Pampadromaeus barberenai</emphasis>
<bibRefCitation id="EFB95698087EFFAD150CCE6AFD771CCA" author="Cabreira S &amp; Schultz C &amp; Bittencourt J" box="[512,729,363,387]" pageId="8" pagination="1035 - 40" refId="ref16312" refString="Cabreira S, Schultz C, Bittencourt J et al. New stem-sauropodomorph (Dinosauria, Saurischia) from the Triassic of Brazil. Die Naturwissenschaften 2011; 98: 1035 - 40." type="journal article" year="2011">
Cabreira
<emphasis id="B95CF77B087EFFAD156FCE6DFD3E1CCA" box="[611,656,363,387]" italics="true" pageId="8">et al</emphasis>
., 2011
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2850EA087EFFAD178DCE8AFE581CEA" authority="(Muller, 2022)" baseAuthorityName="Muller" baseAuthorityYear="2022" box="[129,502,395,419]" genus="Buriolestes" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" phylum="Chordata" rank="species" species="schultzi">
<emphasis id="B95CF77B087EFFAD178DCE8AFEED1CEA" box="[129,323,395,419]" italics="true" pageId="8">Buriolestes schultzi</emphasis>
(Müller, 2022)
</taxonomicName>
. The fossa trochanterica (= facies articularis antitrochanterica) along the posterior side of the greater trochanter is ventrally (= distally) descended, which is common among dinosaurs (
<bibRefCitation id="EFB95698087EFFAD16F2CEE8FD241F48" author="Nesbitt SJ" box="[510,650,489,513]" pageId="8" pagination="1 - 292" refId="ref21334" refString="Nesbitt SJ. The early evolution of archosaurs: relationships and the origin of major clades. Bulletin of the American Museum of Natural History 2011; 352: 1 - 292. https: // doi. org / 10.1206 / 352.1" type="journal article" year="2011">Nesbitt, 2011</bibRefCitation>
).
</paragraph>
<caption id="DF577BE10871FFA2177DC817FEA41AF3" ID-DOI="http://doi.org/10.5281/zenodo.14827200" ID-Zenodo-Dep="14827200" httpUri="https://zenodo.org/record/14827200/files/figure.png" pageId="7" startId="7.[113,178,1814,1838]" targetBox="[116,1456,592,1784]" targetPageId="7" targetType="figure">
<paragraph id="8B972B690871FFA2177DC817FAC11A66" blockId="7.[113,1429,1814,1979]" box="[113,1391,1814,1839]" pageId="7">
<emphasis id="B95CF77B0871FFA2177DC817FF671A66" bold="true" box="[113,201,1814,1839]" pageId="7">Figure 4.</emphasis>
Photographs of sulcimentisaurian silesaurid elements from the Garretts Surprise locality. Distal end of a left humerus UWGM
</paragraph>
<paragraph id="8B972B690871FFA2177DC832FA2F1A02" blockId="7.[113,1429,1814,1979]" box="[113,1409,1843,1867]" pageId="7">7550 (AE) in (A) anterior, (B) posterior, (C) medial, (D) lateral, (E) and distal views. Proximal end of a right femur UWGM 7407 (FJ)</paragraph>
<paragraph id="8B972B690871FFA2177DC84EFEA41AF3" blockId="7.[113,1429,1814,1979]" pageId="7">in (F) proximal, (G) anterolateral, (H) posteromedial, (I) posterolateral, and (J) anteromedial views. Abbreviations: alt, anterolateral tuber; amt, anteromedial tuber; at, anterior trochanter; dlt, dorsolateral trochanter; ect, ectepicondyle; ent, entepicondyle; g, groove; gt, greater trochanter; ipmt, incipient posteromedial tuber; n, notch; rc, radial condyle; uc, ulnar condyle. Arrows point in the anterior direction. Scale bar equals 1 cm.</paragraph>
</caption>
<paragraph id="8B972B69087EFFAD178DCD23FE7819A5" blockId="8.[128,780,546,1260]" pageId="8">
<emphasis id="B95CF77B087EFFAD178DCD23FEC51F73" box="[129,363,546,570]" italics="true" pageId="8">Ontogenetic assessment:</emphasis>
Both the
<typeStatus id="549395CB087EFFAD16D4CD23FD9A1F73" box="[472,564,546,570]" pageId="8" type="holotype">holotype</typeStatus>
astragalus (
<specimenCode id="DB8E8312087EFFAD15A3CD23FF161F10" collectionCode="UWGM" pageId="8">UWGM 1975</specimenCode>
) and referred femur (
<specimenCode id="DB8E8312087EFFAD169DCD40FD861F10" box="[401,552,577,601]" collectionCode="UWGM" pageId="8">UWGM 7549</specimenCode>
) exhibit smooth bone textures and prominent condyles that are consistent with relatively older ontogenetic status in this specimen (
<bibRefCitation id="EFB95698087EFFAD1585CD81FF1B1FFE" author="Griffin CT &amp; Bano LS &amp; Turner AH" pageId="8" refId="ref17941" refString="Griffin CT, Bano LS, Turner AH et al. Integrating gross morphology and bone histology to assess skeletal maturity in early dinosauromorphs: new insights from Dromomeron (Archosauria: Dinosauromorpha). PeerJ 2019; 7: e 6331. https: // doi. org / 10.7717 / peerj. 6331" type="journal volume" year="2019">
Griffin
<emphasis id="B95CF77B087EFFAD15DBCD80FCA41FD1" box="[727,778,640,664]" italics="true" pageId="8">et al.</emphasis>
2019
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). Similarly, the referred femur exhibits muscle scars that are robustly developed and also support the notion of an older ontogenetic status for
<specimenCode id="DB8E8312087EFFAD1667CDDFFE511FBF" box="[363,511,734,758]" collectionCode="UWGM" pageId="8">UWGM 7549</specimenCode>
. An external fundamental system (
<collectionCode id="ED39B3AC087EFFAD17D6CDFCFEA51E5C" box="[218,267,765,789]" pageId="8">EFS</collectionCode>
) is absent from the femoral cross-section (see below); its presence would indicate the attainment of skeletal maturity. That said, other osteohistological signatures are consistent with a more advanced ontogenetic status for
<specimenCode id="DB8E8312087EFFAD15BFCC5AFF161EDB" collectionCode="UWGM" pageId="8">UWGM 7549</specimenCode>
. These data include the presence of endosteal remodelling around the medullary cavity forming an internal fundamental system (
<collectionCode id="ED39B3AC087EFFAD17D4CCB8FEAD1E98" box="[216,259,953,977]" pageId="8">IFS</collectionCode>
), cortical remodelling that extends into the midcortex alongside a transition to more highly organized and less vascularized primary bone tissue at the periosteal margin, which signals a decrease in primary bone apposition in later ontogeny. A lack of an
<collectionCode id="ED39B3AC087EFFAD1606CB36FE9B1906" box="[266,309,1079,1103]" pageId="8">EFS</collectionCode>
indicates that
<specimenCode id="DB8E8312087EFFAD16DDCB36FDCA1907" box="[465,612,1078,1103]" collectionCode="UWGM" pageId="8">UWGM 7549</specimenCode>
was still slowly growing at the time of death. In light of these histological details, the absence of lines of arrested growth (
<collectionCode id="ED39B3AC087EFFAD155DCB74FD2519C4" box="[593,651,1141,1165]" pageId="8">LAG</collectionCode>
) in
<specimenCode id="DB8E8312087EFFAD15BFCB74FF1819E5" collectionCode="UWGM" pageId="8">UWGM 7549</specimenCode>
should not be presumed to indicate an individual less than a single year of age. Instead, this individual had probably progressed beyond earliest ontogeny.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:date id="206D15638AAB1D9D0E640FB530FE240C">2025</mods:date>
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<mods:number id="A703BCAC377597727AF39BD61AD0E05F">203</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="5B30914D5FD52DEF4DF1AF1967D982D7">journal volume</mods:classification>
<mods:identifier id="0B8C6314EB96066767C74154929A06FB" type="CLB-Dataset">308019</mods:identifier>
<mods:identifier id="CCE54397D8F82FDB5E870F0E660B7417" type="DOI">10.1093/zoolinnean/zlae153</mods:identifier>
<mods:identifier id="F57D55EE650CBE9AD9B0FC8CD69ED575" type="GBIF-Dataset">744b4603-204d-448c-a32f-fd0be17f047b</mods:identifier>
<mods:identifier id="905E4CF1A6B2774AD9C81DB51D907E54" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="DD746C11E27478C8C6DDF3ABAB6F6283" type="Zenodo-Dep">14827184</mods:identifier>
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</mods:mods>
<treatment id="03819A7F087EFFAD1617CAD3FA281AE2" ID-DOI="http://doi.org/10.5281/zenodo.14827198" ID-Zenodo-Dep="14827198" LSID="urn:lsid:plazi:treatment:03819A7F087EFFAD1617CAD3FA281AE2" httpUri="http://treatment.plazi.org/id/03819A7F087EFFAD1617CAD3FA281AE2" pageId="8">
<subSubSection id="C33278E2087EFFAD1617CAD3FDC118A4" box="[283,623,1490,1517]" pageId="8" type="nomenclature">
<paragraph id="8B972B69087EFFAD1617CAD3FDC118A4" blockId="8.[283,623,1490,1517]" box="[283,623,1490,1517]" pageId="8">
<heading id="D0DF9C05087EFFAD1617CAD3FDC118A4" bold="true" box="[283,623,1490,1517]" centered="true" fontSize="11" level="2" pageId="8" reason="2">
<emphasis id="B95CF77B087EFFAD1617CAD3FDC118A4" bold="true" box="[283,623,1490,1517]" pageId="8">
<taxonomicName id="4C2850EA087EFFAD1617CAD3FE6D18A4" box="[283,451,1490,1517]" family="Silesauridae" genus="Sulcimentisauria" kingdom="Animalia" order="Dinosauria" pageId="8" phylum="Chordata" rank="genus">
<emphasis id="B95CF77B087EFFAD1617CAD3FE6D18A4" bold="true" box="[283,451,1490,1517]" italics="true" pageId="8">Sulcimentisauria</emphasis>
</taxonomicName>
gen. et sp. indet.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33278E2087EFFAD1692C905FE421B57" box="[414,492,1540,1567]" pageId="8" type="description">
<paragraph id="8B972B69087EFFAD1692C905FE421B57" blockId="8.[414,492,1540,1567]" box="[414,492,1540,1567]" pageId="8">
<emphasis id="B95CF77B087EFFAD1692C905FE421B57" bold="true" box="[414,492,1540,1567]" pageId="8">
(
<figureCitation id="131337EC087EFFAD16A4C904FE4C1B56" box="[424,482,1541,1567]" captionStart="Figure 4" captionStartId="7.[113,178,1814,1838]" captionTargetBox="[116,1456,592,1784]" captionTargetId="figure-257@7.[114,1458,590,1786]" captionTargetPageId="7" captionText="Figure 4. Photographs of sulcimentisaurian silesaurid elements from the Garretts Surprise locality.Distal end of a left humerus UWGM 7550 (AE) in (A) anterior, (B) posterior, (C) medial, (D) lateral, (E) and distal views. Proximal end of a right femur UWGM 7407 (FJ) in (F) proximal, (G) anterolateral, (H) posteromedial, (I) posterolateral, and (J) anteromedial views.Abbreviations:alt, anterolateral tuber; amt, anteromedial tuber; at, anterior trochanter; dlt, dorsolateral trochanter; ect, ectepicondyle; ent, entepicondyle; g, groove; gt, greater trochanter; ipmt, incipient posteromedial tuber; n, notch; rc, radial condyle; uc, ulnar condyle.Arrows point in the anterior direction.Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14827200" httpUri="https://zenodo.org/record/14827200/files/figure.png" pageId="8">Fig. 4</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33278E2087EFFAD178DC944FF631BBC" pageId="8" type="materials_examined">
<paragraph id="8B972B69087EFFAD178DC944FD351B34" blockId="8.[129,777,1605,1661]" pageId="8">
<emphasis id="B95CF77B087EFFAD178DC944FE9E1B14" box="[129,304,1605,1629]" italics="true" pageId="8">Referred material:</emphasis>
<collectionCode id="ED39B3AC087EFFAD163AC944FE221B14" box="[310,396,1605,1629]" pageId="8">UWGM</collectionCode>
7550, the distal end of a left humerus and the proximal end of a right femur,
<collectionCode id="ED39B3AC087EFFAD1509C964FDF51B34" box="[517,603,1637,1661]" pageId="8">UWGM</collectionCode>
7407.
</paragraph>
<paragraph id="8B972B69087EFFAD178DC99FFF631BBC" blockId="8.[129,777,1694,1781]" pageId="8">
<emphasis id="B95CF77B087EFFAD178DC99FFEF21BFF" box="[129,348,1694,1718]" italics="true" pageId="8">Locality and horizon:</emphasis>
The Garretts Surprise locality from the purple unit of the Popo Agie Formation (see further details above).
</paragraph>
</subSubSection>
<subSubSection id="C33278E2087EFFAD178DC817FB8F1F69" pageId="8" type="discussion">
<paragraph id="8B972B69087EFFAD178DC817FB8F1F69" blockId="8.[129,777,1814,1964]" lastBlockId="8.[824,1476,144,544]" pageId="8">
<emphasis id="B95CF77B087EFFAD178DC817FF701A67" box="[129,222,1814,1838]" italics="true" pageId="8">Remarks:</emphasis>
The silesaurid affinities of
<collectionCode id="ED39B3AC087EFFAD150CC817FDF81A67" box="[512,598,1814,1838]" pageId="8">UWGM</collectionCode>
7407 are readily determined by the following combination of features: a notch ventral to the anteromedial tuber; a subtriangular proximal outline due to a prominent anterolateral tuber and incipient posteromedial tuber; and a transverse groove on the proximal surface of the femur (
<bibRefCitation id="EFB95698087EFFAD1314CF91FB521DE1" author="Martz JW &amp; Small BJ" box="[1048,1276,144,168]" pageId="8" refId="ref20705" refString="Martz JW, Small BJ. Non-dinosaurian dinosauromorphs from the Chinle Formation (Upper Triassic) of the Eagle Basin, northern Colorado: Dromomeron romeri (Lagerpetidae) and a new taxon, Kwanasaurus williamparkeri (Silesauridae). PeerJ 2019; 7: e 7551. https: // doi. org / 10.7717 / peerj. 7551" type="journal volume" year="2019">Martz and Small 2019</bibRefCitation>
). The silesaurid affinities of the distal humerus
<collectionCode id="ED39B3AC087EFFAD1369CFAEFB151D8E" box="[1125,1211,175,199]" pageId="8">UWGM</collectionCode>
7550 are less certain, but it compares favourably with the morphology of the Moroccan silesaurid
<taxonomicName id="4C2850EA087EFFAD1493CFEFFAC41C4F" authority="Kammerer et al., 2012" authorityName="Kammerer" authorityYear="2012" box="[927,1386,238,262]" family="Silesauridae" genus="Diodorus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" phylum="Chordata" rank="species" species="scytobrachion">
<emphasis id="B95CF77B087EFFAD1493CFEFFB2D1C4F" box="[927,1155,238,262]" italics="true" pageId="8">Diodorus scytobrachion</emphasis>
<bibRefCitation id="EFB95698087EFFAD1385CFEFFAC41C4F" author="Kammerer CF &amp; Nesbitt SJ &amp; Shubin NH" box="[1161,1386,238,262]" pageId="8" pagination="277 - 84" refId="ref18789" refString="Kammerer CF, Nesbitt SJ, Shubin NH. The first silesaurid dinosauriform from the Late Triassic of Morocco. Acta Palaeontologica Polonica 2012; 57: 277 - 84. https: // doi. org / 10.4202 / app. 2011.0015" type="journal article" year="2012">
Kammerer
<emphasis id="B95CF77B087EFFAD13F1CFEEFA8B1C4F" box="[1277,1317,238,262]" italics="true" pageId="8">et al</emphasis>
., 2012
</bibRefCitation>
</taxonomicName>
. We will refer to the combined OTU of
<collectionCode id="ED39B3AC087EFFAD137FCE0CFB671C6C" box="[1139,1225,269,293]" pageId="8">UWGM</collectionCode>
7407 and
<collectionCode id="ED39B3AC087EFFAD123FCE0CFA271C6C" box="[1331,1417,269,293]" pageId="8">UWGM</collectionCode>
7550 simply as
<collectionCode id="ED39B3AC087EFFAD1496CE2CFC5E1C0C" box="[922,1008,301,325]" pageId="8">UWGM</collectionCode>
7407, unless specifically referring to the femur or humerus, then the element will be included and the catalogue number in parentheses. The combined character states found in the femur (
<collectionCode id="ED39B3AC087EFFAD14A1CE8AFBA91CEA" box="[941,1031,395,419]" pageId="8">UWGM</collectionCode>
7407) are also present in
<taxonomicName id="4C2850EA087EFFAD1219CE8AFA131CEA" box="[1301,1469,395,419]" family="Silesauridae" genus="Diodorus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" phylum="Chordata" rank="species" species="scytobrachion">
<emphasis id="B95CF77B087EFFAD1219CE8AFA131CEA" box="[1301,1469,395,419]" italics="true" pageId="8">Di. scytobrachion</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2850EA087EFFAD1435CEABFA891C8B" authority="Ferigolo and Langer, 2007" authorityName="Ferigolo and Langer" authorityYear="2007" box="[825,1319,426,450]" family="Silesauridae" genus="Sacisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" phylum="Chordata" rank="species" species="agudoensis">
<emphasis id="B95CF77B087EFFAD1435CEABFBBF1C8B" box="[825,1041,426,450]" italics="true" pageId="8">Sacisaurus agudoensis</emphasis>
<bibRefCitation id="EFB95698087EFFAD131BCEABFA891C8B" author="Ferigolo J &amp; Langer M" box="[1047,1319,426,450]" pageId="8" pagination="23 - 33" refId="ref17300" refString="Ferigolo J, Langer M. A Late Triassic dinosauriform from south Brazil and the origin of the ornithischian predentary bone. Historical Biology 2007; 19: 23 - 33. https: // doi. org / 10.1080 / 08912960600845767" type="journal article" year="2007">Ferigolo and Langer, 2007</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C2850EA087EFFAD126ECEABFBA61CA8" authority="Dzik, 2003" authorityName="Dzik" authorityYear="2003" class="Reptilia" genus="Silesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" phylum="Chordata" rank="species" species="opolensis">
<emphasis id="B95CF77B087EFFAD126ECEABFC211CA8" italics="true" pageId="8">Silesaurus opolensis</emphasis>
<bibRefCitation id="EFB95698087EFFAD149ACEC8FBA61CA8" author="Dzik J" box="[918,1032,457,481]" pageId="8" pagination="556 - 74" refId="ref16887" refString="Dzik J. A beaked herbivorous archosaur with dinosaur affinities from the early Late Triassic of Poland. Journal of Vertebrate Paleontology 2003; 23: 556 - 74. https: // doi. org / 10.1671 / a 1097" type="journal article" year="2003">Dzik, 2003</bibRefCitation>
</taxonomicName>
; unfortunately the characters that differentiate these taxa are damaged, obscured, or entirely missing in
<collectionCode id="ED39B3AC087EFFAD1435CD09FC211F69" box="[825,911,520,544]" pageId="8">UWGM</collectionCode>
7407 (
<figureCitation id="131337EC087EFFAD14D4CD09FBBE1F69" box="[984,1040,520,544]" captionStart="Figure 4" captionStartId="7.[113,178,1814,1838]" captionTargetBox="[116,1456,592,1784]" captionTargetId="figure-257@7.[114,1458,590,1786]" captionTargetPageId="7" captionText="Figure 4. Photographs of sulcimentisaurian silesaurid elements from the Garretts Surprise locality.Distal end of a left humerus UWGM 7550 (AE) in (A) anterior, (B) posterior, (C) medial, (D) lateral, (E) and distal views. Proximal end of a right femur UWGM 7407 (FJ) in (F) proximal, (G) anterolateral, (H) posteromedial, (I) posterolateral, and (J) anteromedial views.Abbreviations:alt, anterolateral tuber; amt, anteromedial tuber; at, anterior trochanter; dlt, dorsolateral trochanter; ect, ectepicondyle; ent, entepicondyle; g, groove; gt, greater trochanter; ipmt, incipient posteromedial tuber; n, notch; rc, radial condyle; uc, ulnar condyle.Arrows point in the anterior direction.Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14827200" httpUri="https://zenodo.org/record/14827200/files/figure.png" pageId="8">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33278E2087EFFAD1435CD51FA281AE2" pageId="8" type="description">
<paragraph id="8B972B69087EFFAD1435CD51FBA21871" blockId="8.[825,1477,591,1963]" pageId="8">
<emphasis id="B95CF77B087EFFAD1435CD51FC1E1F2E" box="[825,944,592,615]" italics="true" pageId="8">Description:</emphasis>
The humerus (
<collectionCode id="ED39B3AC087EFFAD1367CD4EFB6B1F2E" box="[1131,1221,591,615]" pageId="8">UWGM</collectionCode>
7550) is fragmentary with only the distal end preserved and significant fracturing through the shaft. The anatomical orientation of the distal end of this silesaurid humerus is complicated by the presence of torsion in the humerus of
<taxonomicName id="4C2850EA087EFFAD1350CDCCFC461E4D" authority="Martz and Small, 2019" authorityName="Martz and Small" authorityYear="2019" genus="Kwanasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" phylum="Chordata" rank="species" species="williamparkeri">
<emphasis id="B95CF77B087EFFAD1350CDCCFAD41FAC" box="[1116,1402,717,741]" italics="true" pageId="8">Kwanasaurus williamparkeri</emphasis>
<bibRefCitation id="EFB95698087EFFAD1288CDCCFC461E4D" author="Martz JW &amp; Small BJ" pageId="8" refId="ref20705" refString="Martz JW, Small BJ. Non-dinosaurian dinosauromorphs from the Chinle Formation (Upper Triassic) of the Eagle Basin, northern Colorado: Dromomeron romeri (Lagerpetidae) and a new taxon, Kwanasaurus williamparkeri (Silesauridae). PeerJ 2019; 7: e 7551. https: // doi. org / 10.7717 / peerj. 7551" type="journal volume" year="2019">Martz and Small, 2019</bibRefCitation>
</taxonomicName>
and lack thereof in other sulcimentisaurian silesaurids (
<bibRefCitation id="EFB95698087EFFAD14B4CC0AFB861E6A" author="Dzik J" box="[952,1064,779,803]" pageId="8" pagination="556 - 74" refId="ref16887" refString="Dzik J. A beaked herbivorous archosaur with dinosaur affinities from the early Late Triassic of Poland. Journal of Vertebrate Paleontology 2003; 23: 556 - 74. https: // doi. org / 10.1671 / a 1097" type="journal article" year="2003">Dzik 2003</bibRefCitation>
,
<bibRefCitation id="EFB95698087EFFAD133BCC0DFA8F1E6A" author="Kammerer CF &amp; Nesbitt SJ &amp; Shubin NH" box="[1079,1313,779,803]" pageId="8" pagination="277 - 84" refId="ref18789" refString="Kammerer CF, Nesbitt SJ, Shubin NH. The first silesaurid dinosauriform from the Late Triassic of Morocco. Acta Palaeontologica Polonica 2012; 57: 277 - 84. https: // doi. org / 10.4202 / app. 2011.0015" type="journal article" year="2012">
Kammerer
<emphasis id="B95CF77B087EFFAD13BCCC0DFB4D1E6A" box="[1200,1251,779,803]" italics="true" pageId="8">et al.</emphasis>
2012
</bibRefCitation>
) and the non-sulcimentisaurian silesaurid
<taxonomicName id="4C2850EA087EFFAD136ACC2AFCC11E2B" authority="Nesbitt et al., 2010" authorityName="Nesbitt" authorityYear="2010" family="Silesauridae" genus="Asilisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" phylum="Chordata" rank="species" species="kongwe">
<emphasis id="B95CF77B087EFFAD136ACC2AFA881E0A" box="[1126,1318,811,835]" italics="true" pageId="8">Asilisaurus kongwe</emphasis>
<bibRefCitation id="EFB95698087EFFAD123DCC2AFCC11E2B" author="Nesbitt SJ &amp; Sidor CA &amp; Irmis RB" pageId="8" pagination="95 - 8" refId="ref21419" refString="Nesbitt SJ, Sidor CA, Irmis RB et al. Ecologically distinct dinosaurian sister group shows early diversification of Ornithodira. Nature 2010; 464: 95 - 8. https: // doi. org / 10.1038 / nature 08718" type="journal article" year="2010">
Nesbitt
<emphasis id="B95CF77B087EFFAD1284CC2AFA191E0A" box="[1416,1463,811,835]" italics="true" pageId="8">et al</emphasis>
., 2010
</bibRefCitation>
</taxonomicName>
, so all interpretations of orientation assume no humeral torsion as is plesiomorphic for silesaurids. The only character relevant to the character matrix determined from the humerus is that the distal condyles are relatively narrow relative to the humeral shaft, as is typical for sulcimentisaurian silesaurids (e.g.
<bibRefCitation id="EFB95698087EFFAD1435CCE6FC091EB6" author="Dzik J" box="[825,935,999,1023]" pageId="8" pagination="556 - 74" refId="ref16887" refString="Dzik J. A beaked herbivorous archosaur with dinosaur affinities from the early Late Triassic of Poland. Journal of Vertebrate Paleontology 2003; 23: 556 - 74. https: // doi. org / 10.1671 / a 1097" type="journal article" year="2003">Dzik 2003</bibRefCitation>
,
<bibRefCitation id="EFB95698087EFFAD14B9CCE6FB341EB6" author="Kammerer CF &amp; Nesbitt SJ &amp; Shubin NH" box="[949,1178,999,1023]" pageId="8" pagination="277 - 84" refId="ref18789" refString="Kammerer CF, Nesbitt SJ, Shubin NH. The first silesaurid dinosauriform from the Late Triassic of Morocco. Acta Palaeontologica Polonica 2012; 57: 277 - 84. https: // doi. org / 10.4202 / app. 2011.0015" type="journal article" year="2012">
Kammerer
<emphasis id="B95CF77B087EFFAD1320CCE6FBF01EB6" box="[1068,1118,999,1023]" italics="true" pageId="8">et al.</emphasis>
2012
</bibRefCitation>
), in contrast to the broader distal humerus of non-sulcimentisaurian silesaurids (
<bibRefCitation id="EFB95698087EFFAD1250CB07FC371974" author="Ezcurra MD &amp; Nesbitt SJ &amp; Fiorelli LE" pageId="8" pagination="1393 - 438" refId="ref17126" refString="Ezcurra MD, Nesbitt SJ, Fiorelli LE et al. New specimen sheds light on the anatomy and taxonomy of the early Late Triassic Dinosauriforms from the Chanares formation, NW Argentina. Anatomical Record (Hoboken, N. J.: 2007) 2020 b; 303: 1393 - 438. https: // doi. org / 10.1002 / ar. 24243" type="journal article" year="2020">
Ezcurra
<emphasis id="B95CF77B087EFFAD12BDCB06FCFC1974" italics="true" pageId="8">et al.</emphasis>
2020b
</bibRefCitation>
,
<bibRefCitation id="EFB95698087EFFAD14ABCB24FBDC1974" author="Nesbitt SJ &amp; Langer MC &amp; Ezcurra MD" box="[935,1138,1061,1085]" pageId="8" pagination="813 - 73" refId="ref21461" refString="Nesbitt SJ, Langer MC, Ezcurra MD. The anatomy of Asilisaurus kongwe, a Dinosauriform from the Lifua member of the Manda beds (Middle Triassic) of Africa. The Anatomical Record 2020 a; 303: 813 - 73." type="journal article" year="2020">
Nesbitt
<emphasis id="B95CF77B087EFFAD14F7CB27FB821974" box="[1019,1068,1061,1085]" italics="true" pageId="8">et al.</emphasis>
2020a
</bibRefCitation>
). The radial and ulnar condyles both form rounded, subtriangular projections with a notch between them, as opposed to the more well-developed condyles of non-sulcimentisaurian silesaurids (
<bibRefCitation id="EFB95698087EFFAD13ACCB85FAC419D2" author="Ezcurra MD &amp; Nesbitt SJ &amp; Fiorelli LE" box="[1184,1386,1155,1179]" pageId="8" pagination="1393 - 438" refId="ref17126" refString="Ezcurra MD, Nesbitt SJ, Fiorelli LE et al. New specimen sheds light on the anatomy and taxonomy of the early Late Triassic Dinosauriforms from the Chanares formation, NW Argentina. Anatomical Record (Hoboken, N. J.: 2007) 2020 b; 303: 1393 - 438. https: // doi. org / 10.1002 / ar. 24243" type="journal article" year="2020">
Ezcurra
<emphasis id="B95CF77B087EFFAD13F9CB85FA8B19D2" box="[1269,1317,1155,1179]" italics="true" pageId="8">et al.</emphasis>
2020b
</bibRefCitation>
,
<bibRefCitation id="EFB95698087EFFAD127BCB82FC0319F2" author="Nesbitt SJ &amp; Langer MC &amp; Ezcurra MD" pageId="8" pagination="813 - 73" refId="ref21461" refString="Nesbitt SJ, Langer MC, Ezcurra MD. The anatomy of Asilisaurus kongwe, a Dinosauriform from the Lifua member of the Manda beds (Middle Triassic) of Africa. The Anatomical Record 2020 a; 303: 813 - 73." type="journal article" year="2020">
Nesbitt
<emphasis id="B95CF77B087EFFAD1435CBA2FCC719F2" box="[825,873,1187,1211]" italics="true" pageId="8">et al.</emphasis>
2020a
</bibRefCitation>
). The ulnar condyle is slightly larger than the radial condyle and has a medial margin that is more posteriorly projected. On both the anterior and posterior face of the humerus above the condyles, there is a fossa that extends about
<quantity id="4CD0868C087EFFAD127CCA00FA0F1851" box="[1392,1441,1281,1305]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" unit="cm" value="1.0">1 cm</quantity>
on to the humeral shaft.
</paragraph>
<paragraph id="8B972B69087EFFAD1459CA3EFA281AE2" blockId="8.[825,1477,591,1963]" pageId="8">
The proximal outline of the femur (
<collectionCode id="ED39B3AC087EFFAD13F5CA3EFAFD181E" box="[1273,1363,1343,1367]" pageId="8">UWGM</collectionCode>
7407) is subtriangular due to the presence of a prominent anterolateral and anteromedial tuber and significant reduction of the posteromedial tuber and straight margins between these features. A straight, transverse groove is present along the proximal surface. A notch is present distal to the head of the femur, which is ubiquitous among silesaurids (e.g.
<bibRefCitation id="EFB95698087EFFAD13CFCAFDFA1F1B5A" author="Martz JW &amp; Small BJ" box="[1219,1457,1531,1555]" pageId="8" refId="ref20705" refString="Martz JW, Small BJ. Non-dinosaurian dinosauromorphs from the Chinle Formation (Upper Triassic) of the Eagle Basin, northern Colorado: Dromomeron romeri (Lagerpetidae) and a new taxon, Kwanasaurus williamparkeri (Silesauridae). PeerJ 2019; 7: e 7551. https: // doi. org / 10.7717 / peerj. 7551" type="journal volume" year="2019">Martz and Small 2019</bibRefCitation>
). The dorsolateral trochanter is broken and displaced across the anterolateral face, but what is preserved is ridge-like. The broken dorsolateral trochanter and surrounding matrix obscure a clear view of the anterior trochanter (= lesser trochanter), but what is visible is mound-like. However, it cannot be determined if the anterior trochanter was small and confluent with the femoral shaft, as in non-sulcimentisaurian silesaurids, or if it was large and offset from the shaft by a cleft, as in sulcimentisaurs, some early core ornithischians, and some neotheropods. The greater trochanter is prominent and rounded, and the fossa trochanterica (= facies articularis antitrochanterica) is level with the greater trochanter. The femoral shaft curves slightly posteriorly, and the femur is broken above the fourth trochanter.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="67F8891951BDD11BE8A7E2E95698FEA1">Rethinking dinosaur origins: oldest known equatorial dinosaur-bearing assemblage (mid-late Carnian Popo Agie FM, Wyoming, USA)</mods:title>
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<mods:namePart id="E9BF63B3F25F8470542DB063B8C8FA8A">Schmitz, Mark</mods:namePart>
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<mods:date id="C7EB27172D08A6DAB9C7ED7B43D8BB45">2025</mods:date>
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<emphasis id="B95CF77B087FFFAC17E7CE0FFD2E1C61" bold="true" box="[235,640,269,296]" pageId="9">
Ichnogenus
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<emphasis id="B95CF77B087FFFAC1660CE0CFE651C61" bold="true" box="[364,459,269,296]" italics="true" pageId="9">Grallator</emphasis>
Hitchcock, 1858
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<emphasis id="B95CF77B087FFFAC177DCE4FFE9E1C2F" box="[113,304,334,358]" italics="true" pageId="9">Referred specimens:</emphasis>
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7435, a partial right pes trace preserved in concave epirelief was collected in the upper Jelm Formation along the Red Wall, Natrona County,
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;
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Jelm Dino Track site
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(
<figureCitation id="131337EC087FFFAC16B4CEADFE5E1C8D" box="[440,496,428,452]" captionStart="Figure 5" captionStartId="9.[115,180,1842,1866]" captionTargetBox="[116,1456,1198,1813]" captionTargetId="figure-569@9.[114,1458,1190,1815]" captionTargetPageId="9" captionText="Figure 5.UWGM 7435 (left) is an isolated slab containing a single tridactyl pes and possible manus impression attributed to an Atreipus Grallator plexus tracemaker from the upper Jelm Formation, Natrona County, WY, USA.A digital surface-depth map (right) produced in METASHAPE (v.2.0.3; Agisoft) from surface light-scans demonstrates the depth and toe pad delineations of pes (p) digits IIIV.The manus impression may be present (m?); other than a very slight depression there are no morphological features to confidently identify it as such. Scale bar is in 1-cm increments. Data reposited on Morphosource.org, ID: 000607578." figureDoi="http://doi.org/10.5281/zenodo.14827204" httpUri="https://zenodo.org/record/14827204/files/figure.png" pageId="9">Fig. 5</figureCitation>
).
</paragraph>
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<subSubSection id="C33278E2087FFFAC177DCEE7FAD81D8E" pageId="9" type="description">
<paragraph id="8B972B69087FFFAC177DCEE7FAD81D8E" blockId="9.[113,764,485,1105]" lastBlockId="9.[810,1459,144,199]" pageId="9">
<emphasis id="B95CF77B087FFFAC177DCEE7FF461CB4" box="[113,232,486,509]" italics="true" pageId="9">Description:</emphasis>
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7435 is a small (9.0 ×
<quantity id="4CD0868C087FFFAC1535CEE4FD2E1CB7" box="[569,640,485,510]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="9" unit="cm" value="5.6">5.6 cm</quantity>
) partial pes impression preserving digits IIIV and, potentially, a very faint manus impression. The isolated slab was found
<emphasis id="B95CF77B087FFFAC156ECD24FD0D1F75" box="[610,675,549,572]" italics="true" pageId="9">ex situ</emphasis>
, though confidentlyconstrainedto 12 mofstratigraphicthickness (topographic constraint) ~
<quantity id="4CD0868C087FFFAC1644CD62FED31F33" box="[328,381,611,635]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="9" unit="m" value="15.0">15 m</quantity>
below the top of the Jelm Formation. The track is located on a 21.5 × 12-cm slab of fluvial sandstone with broad-wavelength, ripple-marked bedding; the preserved trace is digitigrade and tridactyl, mesaxonic where digit III is longest and potentially quadrupedal (
<figureCitation id="131337EC087FFFAC16F9CDE1FD811FB1" box="[501,559,736,760]" captionStart="Figure 5" captionStartId="9.[115,180,1842,1866]" captionTargetBox="[116,1456,1198,1813]" captionTargetId="figure-569@9.[114,1458,1190,1815]" captionTargetPageId="9" captionText="Figure 5.UWGM 7435 (left) is an isolated slab containing a single tridactyl pes and possible manus impression attributed to an Atreipus Grallator plexus tracemaker from the upper Jelm Formation, Natrona County, WY, USA.A digital surface-depth map (right) produced in METASHAPE (v.2.0.3; Agisoft) from surface light-scans demonstrates the depth and toe pad delineations of pes (p) digits IIIV.The manus impression may be present (m?); other than a very slight depression there are no morphological features to confidently identify it as such. Scale bar is in 1-cm increments. Data reposited on Morphosource.org, ID: 000607578." figureDoi="http://doi.org/10.5281/zenodo.14827204" httpUri="https://zenodo.org/record/14827204/files/figure.png" pageId="9">Fig. 5</figureCitation>
). The pes digits are relatively straight, long, and slender with small acuminate claw impressions. Pads are observable, but not sharply defined. Digit IV is closely aligned to III with a 7.5° divarication. Digit III to II has just over double the divarication (17°). The impression of digit IV is
<emphasis id="B95CF77B087FFFAC17D3CC7FFF491EDC" box="[223,231,894,917]" italics="true" pageId="9">c</emphasis>
.
<quantity id="4CD0868C087FFFAC17FECC7CFE801EDD" box="[242,302,893,917]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" unit="mm" value="8.0">8 mm</quantity>
wide with little variance, while digits III and II are
<emphasis id="B95CF77B087FFFAC17BCCC9CFF161EFD" box="[176,184,925,948]" italics="true" pageId="9">c</emphasis>
. 13 and
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, respectively. Although lacking morphological details, a faint impression anterolateral to pes digits IIIII is interpreted to be a manus with three anterolaterally projected digits. Because of this ambiguity in manus identification, we consider this trace within the
<emphasis id="B95CF77B087FFFAC1672CB1BFD99197B" box="[382,567,1050,1074]" italics="true" pageId="9">
<taxonomicName id="4C2850EA087FFFAC1672CB1BFE611978" authorityName="Olsen and Baird" authorityYear="1986" box="[382,463,1050,1073]" class="Reptilia" genus="Atreipus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" phylum="Chordata" rank="genus">Atreipus</taxonomicName>
<taxonomicName id="4C2850EA087FFFAC16D0CB1BFD99197B" authorityName="Hitchcock" authorityYear="1858" box="[476,567,1050,1074]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="9" phylum="Chordata" rank="genus">Grallator</taxonomicName>
</emphasis>
plexus, but cannot confidently assign it to either
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<emphasis id="B95CF77B087FFFAC16ADCB38FE5D1919" box="[417,499,1081,1104]" italics="true" pageId="9">Atreipus</emphasis>
</taxonomicName>
or
<taxonomicName id="4C2850EA087FFFAC151ACB38FDDE1918" authorityName="Hitchcock" authorityYear="1858" box="[534,624,1081,1105]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="9" phylum="Chordata" rank="genus">
<emphasis id="B95CF77B087FFFAC151ACB38FDDE1918" box="[534,624,1081,1105]" italics="true" pageId="9">Grallator</emphasis>
</taxonomicName>
. Considering the ecological implications of a
<taxonomicName id="4C2850EA087FFFAC1398CF91FB411DE1" authorityName="Hitchcock" authorityYear="1858" box="[1172,1263,144,168]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="9" phylum="Chordata" rank="genus">
<emphasis id="B95CF77B087FFFAC1398CF91FB411DE1" box="[1172,1263,144,168]" italics="true" pageId="9">Grallator</emphasis>
</taxonomicName>
(i.e. dinosaurian) tracemaker, we conservatively assign this trace to
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<emphasis id="B95CF77B087FFFAC1212CFB1FADE1D8E" box="[1310,1392,176,199]" italics="true" pageId="9">Atreipus</emphasis>
</taxonomicName>
.
</paragraph>
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</treatment>
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@ -1,75 +1,78 @@
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<mods:title id="5B24118FB30BFB994EE718112A891D13">New insights into the origin and phylogeny of Niphargidae (Crustacea: Amphipoda), with description of a new species and synonymization of the genus Niphargellus with Niphargus</mods:title>
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@ -259,7 +262,7 @@ Small niphargid (body length up to
). First article of mandibulary palp without setae; second article with 2 setae; distal article bearing a single A seta located on outer margin and 57 E setae; comb-like D setae absent; a notsclerotized pouch bearing several small setules is present instead of A and B setae. Gnathopod propodites with convex, slightly inclined palm; palmar corner with a single elongated strong spine. Dactylopodite bearing 1 seta along anterior margin. Telson lobes bearing 4 apical spines; lateral margin with 2 plumose setae; dorsal surface smooth. Epimeral plate I with 1 ventral and 2 posterior spiniform setae; plate III with postero-ventral corner not produced. One short spine present near insertion of uropod I. Uropod I rami subequal in female, exopodite length 85% than endopodite length in male. Uropod III short in both sexes; endopodite stout, as long as wide; exopodite 2-articulated, distal article very short.
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<emphasis id="B91A4549FF99FF8D38B42A48B5FDF8AD" bold="true" box="[129,216,1842,1866]" pageId="8" pageNumber="9">Figure 5.</emphasis>
Chronogram of
@ -269,7 +272,7 @@ and
inferred in BEAST2. Blue bars represent 95% Highest Posterior Density interval for node ages; red dots indicate posterior probabilities (pp)&gt; 0.95, while other dots represent nodes with lower pp; dot area is proportional to pp; blue circles mark calibration points (see text).
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<emphasis id="B91A4549FF98FF8C384429DAB5ECFB5F" bold="true" box="[113,201,1184,1208]" pageId="9" pageNumber="10">Figure 6.</emphasis>
Map showing the distribution of the three species in the
@ -309,7 +312,7 @@ and female
:
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Habitus in
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<figureCitation id="135585DEFF98FF8C38442B4FB5EDF9AB" box="[113,200,1589,1613]" captionStart="Figure 7" captionStartId="10.[129,194,770,794]" captionTargetBox="[323,1278,146,739]" captionTargetId="figure-781@10.[321,1281,144,742]" captionTargetPageId="10" captionText="Figure 7. Niphargus quimperensis, habitus.A, Male holotype. B, Ovigerous female, paratype (photo J.Citoleaux)." figureDoi="http://doi.org/10.5281/zenodo.14827105" httpUri="https://zenodo.org/record/14827105/files/figure.png" pageId="9" pageNumber="10">Figure 7</figureCitation>
&gt; (for comparison habitus of
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<emphasis id="B91A4549FF98FF8C39CA2B4FB7E8F9AA" box="[511,717,1589,1613]" italics="true" pageId="9" pageNumber="10">Niphargellus glenniei</emphasis>
@ -319,52 +322,52 @@ and
<emphasis id="B91A4549FF98FF8C38442B2EB419F98B" box="[113,316,1620,1644]" italics="true" pageId="9" pageNumber="10">Niphargus irlandicus</emphasis>
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reported in
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<figureCitation id="135585DEFF98FF8C398D2B2EB70CF98C" box="[440,553,1620,1644]" captionStart="Figure 12" captionStartId="15.[113,178,926,950]" captionTargetBox="[306,1266,144,898]" captionTargetId="figure-591@15.[306,1266,144,898]" captionTargetPageId="15" captionText="Figure 12. Habitus of male specimens of: (A) Niphargus glenniei; (B) Niphargus irlandicus. Distal article of mandibular palp of: (C) Niphargus glenniei (outer surface); (D) Niphargus irlandicus (inner surface). Field-emission SEM photographs." figureDoi="http://doi.org/10.5281/zenodo.14827117" httpUri="https://zenodo.org/record/14827117/files/figure.png" pageId="9" pageNumber="10">Fig. 12A, B</figureCitation>
, field emission scanning electron microscope photographs). Body length
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<quantity id="4C9634BEFF98FF8C38DF2BE8B41DF94D" box="[234,312,1682,1706]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="10" unit="mm" value="2.6">2.6 mm</quantity>
(female). Head length 10% of body length; rostrum absent. Pereonites IVI with 23 small dorso-medial sensorial setae, pereonite VII with 4 small dorso-medial sensorial setae. Pleonites IIII with 13 small dorsal sensorial setae (
<figureCitation id="135585DEFF98FF8C38492A6AB5F7F8CF" box="[124,210,1808,1832]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." pageId="9" pageNumber="10">Fig. 11A</figureCitation>
<figureCitation id="135585DEFF98FF8C38492A6AB5F7F8CF" box="[124,210,1808,1832]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="9" pageNumber="10">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="8BD1995BFF98FF8C38442A55B082FA55" blockId="9.[113,763,1557,1989]" lastBlockId="9.[809,1460,1340,1991]" pageId="9" pageNumber="10">
Antenna I (
<figureCitation id="135585DEFF98FF8C38C02A55B461F8A0" box="[245,324,1839,1863]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="9" pageNumber="10">Fig. 8A</figureCitation>
<figureCitation id="135585DEFF98FF8C38C02A55B461F8A0" box="[245,324,1839,1863]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="9" pageNumber="10">Fig. 8A</figureCitation>
). 46% of body length. Flagellum of 12 articles; articles bearing 1 aesthetasc, its length more than one half of article length; aesthetascs accompanied by 46 setae. Distal article shorter than the aesthetasc of penultimate article, bearing 6 distal setae and a short aesthetasc. Peduncle article triarticulated (
<figureCitation id="135585DEFF98FF8C3BF72846B128FAB3" box="[962,1037,1340,1364]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="9" pageNumber="10">Fig. 8A</figureCitation>
<figureCitation id="135585DEFF98FF8C3BF72846B128FAB3" box="[962,1037,1340,1364]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="9" pageNumber="10">Fig. 8A</figureCitation>
), proportions of articles length 1:2:3 as 1.0:0.8:0.5. Accessory flagellum biarticulated (
<figureCitation id="135585DEFF98FF8C3D282821B04CFA94" box="[1309,1385,1371,1395]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="9" pageNumber="10">Fig. 8B</figureCitation>
<figureCitation id="135585DEFF98FF8C3D282821B04CFA94" box="[1309,1385,1371,1395]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="9" pageNumber="10">Fig. 8B</figureCitation>
); proximal article bearing 3 distal setae; distal article as long as one third of proximal article, bearing 2 setae and a short aesthetasc.
</paragraph>
<paragraph id="8BD1995BFF98FF8C3B1F28C3B656F989" blockId="9.[809,1460,1340,1991]" pageId="9" pageNumber="10">
Antenna II (
<figureCitation id="135585DEFF98FF8C3B9D28C3B6D7FA36" box="[936,1010,1465,1489]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="9" pageNumber="10">Fig. 8C</figureCitation>
<figureCitation id="135585DEFF98FF8C3B9D28C3B6D7FA36" box="[936,1010,1465,1489]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="9" pageNumber="10">Fig. 8C</figureCitation>
) as long as one third of antenna I. Flagellum of antenna II with 5 articles; each article with 57 setae and one sensillum; distal article bearing 4 distal setae accompanied by 1 aesthetasc. Peduncle with proportions of articles length 1:2:3 as 1.0:4.0:4.0; articles 2 and 3 slender; flagellum 70% of peduncle length.
</paragraph>
<paragraph id="8BD1995BFF98FF8C3B702B0FB0A1F94A" blockId="9.[809,1460,1340,1991]" pageId="9" pageNumber="10">
Labium (
<figureCitation id="135585DEFF98FF8C3B912B0FB6CBF96A" box="[932,1006,1653,1677]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="9" pageNumber="10">Fig. 8D</figureCitation>
<figureCitation id="135585DEFF98FF8C3B912B0FB6CBF96A" box="[932,1006,1653,1677]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="9" pageNumber="10">Fig. 8D</figureCitation>
) with short inner lobes, without setae. Outer lobes bearing one subapical row of fine setae on inner sides.
</paragraph>
<paragraph id="8BD1995BFF98FF8C3B702BCEB0B0F92B" blockId="9.[809,1460,1340,1991]" box="[837,1429,1716,1740]" pageId="9" pageNumber="10">
Labrum (
<figureCitation id="135585DEFF98FF8C3B9D2BCEB6CBF92B" box="[936,1006,1716,1740]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="9" pageNumber="10">Fig. 8E</figureCitation>
<figureCitation id="135585DEFF98FF8C3B9D2BCEB6CBF92B" box="[936,1006,1716,1740]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="9" pageNumber="10">Fig. 8E</figureCitation>
) typical of the genus, sub-ovoid in shape.
</paragraph>
<paragraph id="8BD1995BFF98FF8F3B702BA9B4EFFB02" blockId="9.[809,1460,1340,1991]" lastBlockId="10.[127,779,853,2005]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Mandible (
<figureCitation id="135585DEFF98FF8C3B8D2BA9B13EF90C" box="[952,1051,1747,1771]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="9" pageNumber="10">Fig. 8F, G</figureCitation>
<figureCitation id="135585DEFF98FF8C3B8D2BA9B13EF90C" box="[952,1051,1747,1771]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="9" pageNumber="10">Fig. 8F, G</figureCitation>
). Right mandible (
<figureCitation id="135585DEFF98FF8C3CEA2BA9B000F90C" box="[1247,1317,1747,1771]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="9" pageNumber="10">Fig. 8F</figureCitation>
<figureCitation id="135585DEFF98FF8C3CEA2BA9B000F90C" box="[1247,1317,1747,1771]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="9" pageNumber="10">Fig. 8F</figureCitation>
): incisor process with 4 teeth, lacinia mobilis with 2 teeth; between lacinia mobilis and pars molaris a row of 5 setae is present; long seta accompanying pars molaris present (truncated in figure); 3 small spines accompanying pars molaris present. Left mandible (
<figureCitation id="135585DEFF98FF8C3DB82A2BB66FF86F" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="9" pageNumber="10">Fig. 8G</figureCitation>
<figureCitation id="135585DEFF98FF8C3DB82A2BB66FF86F" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="9" pageNumber="10">Fig. 8G</figureCitation>
): incisor process with 4 teeth, lacinia mobilis with 2 teeth, one of them pluritoothed; between lacinia mobilis and pars molaris a row of 3 setae is present. Proportions of mandibular palp articles (
<figureCitation id="135585DEFF9BFF8F39392E2FB472FC8A" box="[268,343,853,877]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="10" pageNumber="11">Fig. 8G</figureCitation>
<figureCitation id="135585DEFF9BFF8F39392E2FB472FC8A" box="[268,343,853,877]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="10" pageNumber="11">Fig. 8G</figureCitation>
) 1:2:3 as 1.0:3.0:2.4. Proximal palp article without setae; second article with 2 setae; distal article with 57 E setae (
<figureCitation id="135585DEFF9BFF8F393F2EEEB452FC4C" box="[266,375,916,940]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="10" pageNumber="11">Fig. 8G, H</figureCitation>
<figureCitation id="135585DEFF9BFF8F393F2EEEB452FC4C" box="[266,375,916,940]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="10" pageNumber="11">Fig. 8G, H</figureCitation>
); comb-like D setae absent; a single A seta is present, shifted to the outer margin. An unsclerotized pouch bearing several small setules is present, instead of A and B setae (
<figureCitation id="135585DEFF9BFF8F38EC2E88B400FBED" box="[217,293,1010,1034]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="10" pageNumber="11">Fig. 8H</figureCitation>
<figureCitation id="135585DEFF9BFF8F38EC2E88B400FBED" box="[217,293,1010,1034]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="10" pageNumber="11">Fig. 8H</figureCitation>
). This unusual feature is present in
<taxonomicName id="4C6EE2D8FF9BFF8F3AB82E88B5EEFBCE" baseAuthorityName="Spooner" baseAuthorityYear="1952" class="Malacostraca" family="Niphargidae" genus="Niphargellus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="glenniei">
<emphasis id="B91A4549FF9BFF8F3AB82E88B5EEFBCE" italics="true" pageId="10" pageNumber="11">Niphargellus glenniei</emphasis>
@ -374,14 +377,14 @@ and
<emphasis id="B91A4549FF9BFF8F3935296BB4EBFBCE" box="[256,462,1041,1065]" italics="true" pageId="10" pageNumber="11">Niphargus irlandicus</emphasis>
</taxonomicName>
as well (
<figureCitation id="135585DEFF9BFF8F3A1B296BB78DFBCF" box="[558,680,1041,1065]" captionStart="Figure 12" captionStartId="15.[113,178,926,950]" captionTargetBox="[306,1266,144,898]" captionTargetId="figure-591@15.[306,1266,144,898]" captionTargetPageId="15" captionText="Figure 12. Habitus of male specimens of: (A) Niphargus glenniei; (B) Niphargus irlandicus. Distal article of mandibular palp of: (C) Niphargus glenniei (outer surface); (D) Niphargus irlandicus (inner surface). Field-emission SEM photographs." pageId="10" pageNumber="11">Fig. 12C, D</figureCitation>
<figureCitation id="135585DEFF9BFF8F3A1B296BB78DFBCF" box="[558,680,1041,1065]" captionStart="Figure 12" captionStartId="15.[113,178,926,950]" captionTargetBox="[306,1266,144,898]" captionTargetId="figure-591@15.[306,1266,144,898]" captionTargetPageId="15" captionText="Figure 12. Habitus of male specimens of: (A) Niphargus glenniei; (B) Niphargus irlandicus. Distal article of mandibular palp of: (C) Niphargus glenniei (outer surface); (D) Niphargus irlandicus (inner surface). Field-emission SEM photographs." figureDoi="http://doi.org/10.5281/zenodo.14827117" httpUri="https://zenodo.org/record/14827117/files/figure.png" pageId="10" pageNumber="11">Fig. 12C, D</figureCitation>
; FESEM photographs), and it was never described in other niphargids. Unfortunately, the material at our disposition did not allow the examination of the setule pouch of
<taxonomicName id="4C6EE2D8FF9BFF8F39D12915B7EBFB60" authorityName="Citoleux, Flot &amp; Stoch" authorityYear="2024" box="[484,718,1135,1159]" class="Malacostraca" family="Niphargidae" genus="Niphargus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="quimperensis">
<emphasis id="B91A4549FF9BFF8F39D12915B7EBFB60" box="[484,718,1135,1159]" italics="true" pageId="10" pageNumber="11">Niphargus quimperensis</emphasis>
</taxonomicName>
using FESEM because the cuticle shrank during dehydration, but the structure is well evident in the observation using the compound microscope equipped with DIC.
</paragraph>
<caption id="DF11C9D3FF9BFF8F38B42E78B187FCFD" box="[129,1186,770,795]" pageId="10" pageNumber="11" startId="10.[129,194,770,794]" targetBox="[323,1278,146,739]" targetPageId="10" targetType="figure">
<caption id="DF11C9D3FF9BFF8F38B42E78B187FCFD" ID-DOI="http://doi.org/10.5281/zenodo.14827105" ID-Zenodo-Dep="14827105" box="[129,1186,770,795]" httpUri="https://zenodo.org/record/14827105/files/figure.png" pageId="10" pageNumber="11" startId="10.[129,194,770,794]" targetBox="[323,1278,146,739]" targetPageId="10" targetType="figure">
<paragraph id="8BD1995BFF9BFF8F38B42E78B187FCFD" blockId="10.[129,1186,770,795]" box="[129,1186,770,795]" pageId="10" pageNumber="11">
<emphasis id="B91A4549FF9BFF8F38B42E78B5FDFCFD" bold="true" box="[129,216,770,794]" pageId="10" pageNumber="11">Figure 7.</emphasis>
<taxonomicName id="4C6EE2D8FF9BFF8F38E82E78B490FCFD" authorityName="Citoleux, Flot &amp; Stoch" authorityYear="2024" box="[221,437,770,794]" class="Malacostraca" family="Niphargidae" genus="Niphargus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="quimperensis">
@ -392,55 +395,55 @@ using FESEM because the cuticle shrank during dehydration, but the structure is
</caption>
<paragraph id="8BD1995BFF9BFF8F38A92996B737FA47" blockId="10.[127,779,853,2005]" pageId="10" pageNumber="11">
Maxilla I (
<figureCitation id="135585DEFF9BFF8F393F2996B46EFAE3" box="[266,331,1260,1284]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="10" pageNumber="11">Fig. 8I</figureCitation>
<figureCitation id="135585DEFF9BFF8F393F2996B46EFAE3" box="[266,331,1260,1284]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="10" pageNumber="11">Fig. 8I</figureCitation>
). Palp articles 1 and 2 ratio 1:3, with distal article bearing 6 thicker apical setae and several outer setae on the surface. Outer lobe with 6 spines with 12 small teeth and one spine with several, small teeth; a group of 6 facial setae is present. Inner lobe elongated (3 times longer than wide), with 4 inner setae and a single, long distal seta.
</paragraph>
<paragraph id="8BD1995BFF9BFF8F38A928D2B75EFA19" blockId="10.[127,779,853,2005]" pageId="10" pageNumber="11">
Maxilla II (
<figureCitation id="135585DEFF9BFF8F392528D2B474FA27" box="[272,337,1448,1472]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="10" pageNumber="11">Fig. 8J</figureCitation>
<figureCitation id="135585DEFF9BFF8F392528D2B474FA27" box="[272,337,1448,1472]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="10" pageNumber="11">Fig. 8J</figureCitation>
) inner lobe 0.8 times as long as outer lobe; both of them with the apical and subapical setae as usual in the genus, accompanied by several, sparse facial setae.
</paragraph>
<paragraph id="8BD1995BFF9BFF8F38A92B7CB422F97C" blockId="10.[127,779,853,2005]" pageId="10" pageNumber="11">
Maxilliped (
<figureCitation id="135585DEFF9BFF8F39152B7CB455F9F9" box="[288,368,1542,1566]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="10" pageNumber="11">Fig. 8K</figureCitation>
<figureCitation id="135585DEFF9BFF8F39152B7CB455F9F9" box="[288,368,1542,1566]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="10" pageNumber="11">Fig. 8K</figureCitation>
). Palp distal article with 2 very welldeveloped setae close to the insertion of nail. Outer lobe with 2 strong setae and 7 flattened, thick spines. Inner lobe bearing 5 apical setae. Numerous sparse setae are present on the lobes, as in
<figureCitation id="135585DEFF9BFF8F38AF2BFEB425F97B" box="[154,256,1668,1692]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." pageId="10" pageNumber="11">Figure 8K</figureCitation>
<figureCitation id="135585DEFF9BFF8F38AF2BFEB425F97B" box="[154,256,1668,1692]" captionStart="Figure 8" captionStartId="11.[114,179,1836,1860]" captionTargetBox="[193,1381,144,1808]" captionTargetId="graphics-8@11.[193,1380,144,1760]" captionTargetPageId="11" captionText="Figure 8. Niphargus quimperensis, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium.E, Labrum. F, Right mandible.G, Left mandible.H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="10" pageNumber="11">Figure 8K</figureCitation>
.
</paragraph>
<paragraph id="8BD1995BFF9BFF8F38A92BD9B004FC0D" blockId="10.[127,779,853,2005]" lastBlockId="10.[823,1477,853,2005]" pageId="10" pageNumber="11">
Gnathopod I (
<figureCitation id="135585DEFF9BFF8F390F2BD9B4A2F95C" box="[314,391,1699,1723]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." pageId="10" pageNumber="11">Fig. 9A</figureCitation>
<figureCitation id="135585DEFF9BFF8F390F2BD9B4A2F95C" box="[314,391,1699,1723]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827111" httpUri="https://zenodo.org/record/14827111/files/figure.png" pageId="10" pageNumber="11">Fig. 9A</figureCitation>
). Coxa 1.75 times wider than long, with 1 long marginal seta. Basipodite 2.3 times longer than wide, bearing 3 long setae on posterior margin; ischiopodite with 2 posterior setae, meropodite bearing a postero-distal group of 7 setae; carpopodite with a continuous row of setae of variable length on posterior margin, and only one seta on anterior margin. Propodite smaller and narrower than in gnathopod 2, with convex, only slightly inclined palm, bearing along anterior margin 3 small groups of 23 setae each; posterior margin usually with a single seta; a row of 5 facial setae close to the insertion of palmar spine present both on inner and outer face; antero-distal group of 4 setae close to the insertion of dactylopodite. Palmar corner with a single strong elongated spine. Dactylopodite bearing 1 seta along anterior margin; distal nail as long as one half of dactylopodite, bearing a subungueal seta.
</paragraph>
<paragraph id="8BD1995BFF9BFF8F3B602E88B09DFA66" blockId="10.[823,1477,853,2005]" pageId="10" pageNumber="11">
Gnathopod II (
<figureCitation id="135585DEFF9BFF8F3BC02E88B17EFBED" box="[1013,1115,1010,1034]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." pageId="10" pageNumber="11">Fig. 9B, C</figureCitation>
<figureCitation id="135585DEFF9BFF8F3BC02E88B17EFBED" box="[1013,1115,1010,1034]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827111" httpUri="https://zenodo.org/record/14827111/files/figure.png" pageId="10" pageNumber="11">Fig. 9B, C</figureCitation>
). Coxa suboval, with three slender posterior setae. Gill (epipodite) as long as basipodite. Oostegite in females as long as gill. Basipodite elongated, 2.5 times longer than wide; ischiopodite, meropodite, and carpopodite shape and setation similar to gnathopod 1 (
<figureCitation id="135585DEFF9BFF8F3CA62915B1F9FB60" box="[1171,1244,1135,1159]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." pageId="10" pageNumber="11">Fig. 9B</figureCitation>
<figureCitation id="135585DEFF9BFF8F3CA62915B1F9FB60" box="[1171,1244,1135,1159]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827111" httpUri="https://zenodo.org/record/14827111/files/figure.png" pageId="10" pageNumber="11">Fig. 9B</figureCitation>
). Propodite 1.2 times longer than wide; palm convex (
<figureCitation id="135585DEFF9BFF8F3CA129F4B1DAFB41" box="[1172,1279,1166,1190]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." pageId="10" pageNumber="11">Fig. 9B, C</figureCitation>
<figureCitation id="135585DEFF9BFF8F3CA129F4B1DAFB41" box="[1172,1279,1166,1190]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827111" httpUri="https://zenodo.org/record/14827111/files/figure.png" pageId="10" pageNumber="11">Fig. 9B, C</figureCitation>
) identical in both sexes, bearing along posterior margin 4 dense rows of setae; anterior margin with a single row of 3 setae, and a row of 4 setae close to the insertion of palmar spine; antero-distal group of 34 setae close to the insertion of dactylopodite. Palmar corner with a strong spine not accompanied by smaller spines. Dactylopodite bearing 1 seta along dorsal margin; distal nail as long as one third of dactylopodite; distal nail accompanied by a subungueal seta.
</paragraph>
<paragraph id="8BD1995BFF9BFF8F3B6028F3B12DF9F9" blockId="10.[823,1477,853,2005]" pageId="10" pageNumber="11">
Pereopods IIIIV (
<figureCitation id="135585DEFF9BFF8F3C2D28F3B15EFA47" box="[1048,1147,1417,1441]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." pageId="10" pageNumber="11">Fig. 9D, E</figureCitation>
<figureCitation id="135585DEFF9BFF8F3C2D28F3B15EFA47" box="[1048,1147,1417,1441]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827111" httpUri="https://zenodo.org/record/14827111/files/figure.png" pageId="10" pageNumber="11">Fig. 9D, E</figureCitation>
). Pereopods III and IV approximately subequal, shape and setation as in
<figureCitation id="135585DEFF9BFF8F3CD128D2B01DFA27" box="[1252,1336,1448,1472]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." pageId="10" pageNumber="11">Figure 9</figureCitation>
<figureCitation id="135585DEFF9BFF8F3CD128D2B01DFA27" box="[1252,1336,1448,1472]" captionStart="Figure 9" captionStartId="12.[129,194,1832,1856]" captionTargetBox="[235,1367,140,1804]" captionTargetId="graphics-7@12.[235,1367,140,1804]" captionTargetPageId="12" captionText="Figure 9. Niphargus quimperensis, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV.Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827111" httpUri="https://zenodo.org/record/14827111/files/figure.png" pageId="10" pageNumber="11">Figure 9</figureCitation>
. Gills as long as basipodite, elongated. Dactylopodites IIIIV with a single dorsal plumose seta, and one short seta on ventral side, close to the insertion of nail.
</paragraph>
<paragraph id="8BD1995BFF9BFF8F3B602B5CB174F890" blockId="10.[823,1477,853,2005]" pageId="10" pageNumber="11">
Pereopods VVII (
<figureCitation id="135585DEFF9BFF8F3C1B2B5CB15AF9D9" box="[1070,1151,1574,1598]" captionStart="Figure 10" captionStartId="13.[115,180,1834,1858]" captionTargetBox="[197,1379,143,1806]" captionTargetId="graphics-6@13.[197,1379,143,1807]" captionTargetPageId="13" captionText="Figure 10. Niphargus quimperensis, male, holotype (C, D, E) and female, paratype (A, B). A, Pereopod V. B, Dactylopodite of pereopod V.C, Pereopod VI. D, Pereopod VII. E, Meropodite of pereopod 7, lateral view. Scale bars 0.30 mm." pageId="10" pageNumber="11">Fig. 10</figureCitation>
<figureCitation id="135585DEFF9BFF8F3C1B2B5CB15AF9D9" box="[1070,1151,1574,1598]" captionStart="Figure 10" captionStartId="13.[115,180,1834,1858]" captionTargetBox="[197,1379,143,1806]" captionTargetId="graphics-6@13.[197,1379,143,1807]" captionTargetPageId="13" captionText="Figure 10. Niphargus quimperensis, male, holotype (C, D, E) and female, paratype (A, B). A, Pereopod V. B, Dactylopodite of pereopod V.C, Pereopod VI. D, Pereopod VII. E, Meropodite of pereopod 7, lateral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.14827113" httpUri="https://zenodo.org/record/14827113/files/figure.png" pageId="10" pageNumber="11">Fig. 10</figureCitation>
). Proportions of pereopods V:VI:VII as 1.0:1.2:1.2, shape and setation as in
<figureCitation id="135585DEFF9BFF8F3D232B3FB053F9BA" box="[1302,1398,1605,1629]" captionStart="Figure 10" captionStartId="13.[115,180,1834,1858]" captionTargetBox="[197,1379,143,1806]" captionTargetId="graphics-6@13.[197,1379,143,1807]" captionTargetPageId="13" captionText="Figure 10. Niphargus quimperensis, male, holotype (C, D, E) and female, paratype (A, B). A, Pereopod V. B, Dactylopodite of pereopod V.C, Pereopod VI. D, Pereopod VII. E, Meropodite of pereopod 7, lateral view. Scale bars 0.30 mm." pageId="10" pageNumber="11">Figure 10</figureCitation>
<figureCitation id="135585DEFF9BFF8F3D232B3FB053F9BA" box="[1302,1398,1605,1629]" captionStart="Figure 10" captionStartId="13.[115,180,1834,1858]" captionTargetBox="[197,1379,143,1806]" captionTargetId="graphics-6@13.[197,1379,143,1807]" captionTargetPageId="13" captionText="Figure 10. Niphargus quimperensis, male, holotype (C, D, E) and female, paratype (A, B). A, Pereopod V. B, Dactylopodite of pereopod V.C, Pereopod VI. D, Pereopod VII. E, Meropodite of pereopod 7, lateral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.14827113" httpUri="https://zenodo.org/record/14827113/files/figure.png" pageId="10" pageNumber="11">Figure 10</figureCitation>
. Coxae narrow and elongate; coxa VII small, 3 times wider than long, with 2 marginal setae. Gills (epipodites) on pereopod V and VI as long as coxal width. Basipodites VVII, with slightly concave posterior margins, without distal lobes; posterior margins with 47 short setae; anterior margins with setae only. Ischiopodites VVII with a marked notch. Dactylopodites VVII with nails length about one third of total dactylus length; dorsal margins with a single plumose seta; ventral margin bearing a distal spine inserted near nail insertion.
</paragraph>
<paragraph id="8BD1995BFF9BFF8F3B602A04B148F832" blockId="10.[823,1477,853,2005]" pageId="10" pageNumber="11">
Pleopods (
<figureCitation id="135585DEFF9BFF8F3BF72A04B136F871" box="[962,1043,1918,1942]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." pageId="10" pageNumber="11">Fig.11B</figureCitation>
<figureCitation id="135585DEFF9BFF8F3BF72A04B136F871" box="[962,1043,1918,1942]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="10" pageNumber="11">Fig.11B</figureCitation>
). PleopodsIIIIprotopodites (peduncles) with 2 hooked retinacles. Rami (exopods and endopods) of 6 articles with 2 long setae each.
</paragraph>
<caption id="DF11C9D3FF9AFF8E38472A56B01AF8B8" pageId="11" pageNumber="12" startId="11.[114,179,1836,1860]" targetBox="[193,1381,144,1808]" targetPageId="11" targetType="figure">
<caption id="DF11C9D3FF9AFF8E38472A56B01AF8B8" ID-DOI="http://doi.org/10.5281/zenodo.14827107" ID-Zenodo-Dep="14827107" httpUri="https://zenodo.org/record/14827107/files/figure.png" pageId="11" pageNumber="12" startId="11.[114,179,1836,1860]" targetBox="[193,1381,144,1808]" targetPageId="11" targetType="figure">
<paragraph id="8BD1995BFF9AFF8E38472A56B01AF8B8" blockId="11.[114,1431,1836,1888]" pageId="11" pageNumber="12">
<emphasis id="B91A4549FF9AFF8E38472A56B5ECF8A3" bold="true" box="[114,201,1836,1860]" pageId="11" pageNumber="12">Figure 8.</emphasis>
<taxonomicName id="4C6EE2D8FF9AFF8E38FB2A56B483F8A3" authorityName="Citoleux, Flot &amp; Stoch" authorityYear="2024" box="[206,422,1836,1860]" class="Malacostraca" family="Niphargidae" genus="Niphargus" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="quimperensis">
@ -449,7 +452,7 @@ Pleopods (
, male, holotype. A, Antenna I. B, Accessory flagellum of antenna I. C, Antenna II. D, Labium. E, Labrum. F, Right mandible. G, Left mandible. H, Distal article of mandibular palp. I, Maxilla I. J, Maxilla II. K, Maxilliped. Scale bars 0.10 mm.
</paragraph>
</caption>
<caption id="DF11C9D3FF9DFF8938B42A52B6BFF8BB" pageId="12" pageNumber="13" startId="12.[129,194,1832,1856]" targetBox="[235,1367,140,1804]" targetPageId="12" targetType="figure">
<caption id="DF11C9D3FF9DFF8938B42A52B6BFF8BB" ID-DOI="http://doi.org/10.5281/zenodo.14827111" ID-Zenodo-Dep="14827111" httpUri="https://zenodo.org/record/14827111/files/figure.png" pageId="12" pageNumber="13" startId="12.[129,194,1832,1856]" targetBox="[235,1367,140,1804]" targetPageId="12" targetType="figure">
<paragraph id="8BD1995BFF9DFF8938B42A52B6BFF8BB" blockId="12.[129,1442,1832,1884]" pageId="12" pageNumber="13">
<emphasis id="B91A4549FF9DFF8938B42A52B5FDF8A7" bold="true" box="[129,216,1832,1856]" pageId="12" pageNumber="13">Figure 9.</emphasis>
<taxonomicName id="4C6EE2D8FF9DFF8938E82A52B490F8A7" authorityName="Citoleux, Flot &amp; Stoch" authorityYear="2024" box="[221,437,1832,1856]" class="Malacostraca" family="Niphargidae" genus="Niphargus" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quimperensis">
@ -458,7 +461,7 @@ Pleopods (
, male, holotype (A, C) and female, paratype (B, D, E). A, Gnathopod I. B, Gnathopod II. C, Propodite and dactylopodite of gnathopod II. D, Pereopod III. E, Pereopod IV. Scale bars 0.20 mm.
</paragraph>
</caption>
<caption id="DF11C9D3FF9CFF8838462A50B6C3F8B9" pageId="13" pageNumber="14" startId="13.[115,180,1834,1858]" targetBox="[197,1379,143,1806]" targetPageId="13" targetType="figure">
<caption id="DF11C9D3FF9CFF8838462A50B6C3F8B9" ID-DOI="http://doi.org/10.5281/zenodo.14827113" ID-Zenodo-Dep="14827113" httpUri="https://zenodo.org/record/14827113/files/figure.png" pageId="13" pageNumber="14" startId="13.[115,180,1834,1858]" targetBox="[197,1379,143,1806]" targetPageId="13" targetType="figure">
<paragraph id="8BD1995BFF9CFF8838462A50B6C3F8B9" blockId="13.[115,1434,1834,1887]" pageId="13" pageNumber="14">
<emphasis id="B91A4549FF9CFF8838462A50B5F4F8A4" bold="true" box="[115,209,1834,1859]" pageId="13" pageNumber="14">Figure 10</emphasis>
.
@ -468,7 +471,7 @@ Pleopods (
, male, holotype (C, D, E) and female, paratype (A, B). A, Pereopod V. B, Dactylopodite of pereopod V. C, Pereopod VI. D, Pereopod VII. E, Meropodite of pereopod 7, lateral view. Scale bars 0.30 mm.
</paragraph>
</caption>
<caption id="DF11C9D3FF9FFF8B38B42A50B66FF871" pageId="14" pageNumber="15" startId="14.[129,194,1834,1858]" targetBox="[199,1405,142,1807]" targetPageId="14" targetType="figure">
<caption id="DF11C9D3FF9FFF8B38B42A50B66FF871" ID-DOI="http://doi.org/10.5281/zenodo.14827115" ID-Zenodo-Dep="14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="14" pageNumber="15" startId="14.[129,194,1834,1858]" targetBox="[199,1405,142,1807]" targetPageId="14" targetType="figure">
<paragraph id="8BD1995BFF9FFF8B38B42A50B66FF871" blockId="14.[129,1474,1834,1943]" pageId="14" pageNumber="15">
<emphasis id="B91A4549FF9FFF8B38B42A50B5C0F8A4" bold="true" box="[129,229,1834,1859]" pageId="14" pageNumber="15">Figure 11.</emphasis>
<taxonomicName id="4C6EE2D8FF9FFF8B38DF2A51B4E7F8A5" authorityName="Citoleux, Flot &amp; Stoch" authorityYear="2024" box="[234,450,1835,1859]" class="Malacostraca" family="Niphargidae" genus="Niphargus" kingdom="Animalia" order="Amphipoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="quimperensis">
@ -477,7 +480,7 @@ Pleopods (
, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson). B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm.
</paragraph>
</caption>
<caption id="DF11C9D3FF9EFF8A38442EE4B13DFC35" pageId="15" pageNumber="16" startId="15.[113,178,926,950]" targetBox="[306,1266,144,898]" targetPageId="15" targetType="figure">
<caption id="DF11C9D3FF9EFF8A38442EE4B13DFC35" ID-DOI="http://doi.org/10.5281/zenodo.14827117" ID-Zenodo-Dep="14827117" httpUri="https://zenodo.org/record/14827117/files/figure.png" pageId="15" pageNumber="16" startId="15.[113,178,926,950]" targetBox="[306,1266,144,898]" targetPageId="15" targetType="figure">
<paragraph id="8BD1995BFF9EFF8A38442EE4B13DFC35" blockId="15.[113,1441,926,978]" pageId="15" pageNumber="16">
<emphasis id="B91A4549FF9EFF8A38442EE4B5F3FC51" bold="true" box="[113,214,926,950]" pageId="15" pageNumber="16">Figure 12.</emphasis>
Habitus of male specimens of: (A)
@ -501,35 +504,35 @@ Habitus of male specimens of: (A)
</caption>
<paragraph id="8BD1995BFF9EFF8A38B82978B4C6FA75" blockId="15.[112,765,1026,1771]" pageId="15" pageNumber="16">
Uropods (
<figureCitation id="135585DEFF9EFF8A38C92978B45FFBFD" box="[252,378,1026,1050]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." pageId="15" pageNumber="16">Fig. 11DG</figureCitation>
<figureCitation id="135585DEFF9EFF8A38C92978B45FFBFD" box="[252,378,1026,1050]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="15" pageNumber="16">Fig. 11DG</figureCitation>
). Uropod I (
<figureCitation id="135585DEFF9EFF8A3A3B2978B74EFBFD" box="[526,619,1026,1050]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." pageId="15" pageNumber="16">Fig. 11D</figureCitation>
<figureCitation id="135585DEFF9EFF8A3A3B2978B74EFBFD" box="[526,619,1026,1050]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="15" pageNumber="16">Fig. 11D</figureCitation>
) protopodite with no dorsal spines and 1 distal spine close to the insertion of exopodite; length ratio endopodite:exopodite as 1.00:
<quantity id="4C9634BEFF9EFF8A3A84293BB7DFFBBE" box="[689,762,1089,1113]" metricMagnitude="-2" metricUnit="m" metricValue="2.159" pageId="15" pageNumber="16" unit="in" value="0.85">0.85 in</quantity>
male, subequal in female; rami quite straight; endopodite with 5 terminal spines, exopodite with 5 terminal spines. Uropod II (
<figureCitation id="135585DEFF9EFF8A384929E5B5F5FB50" box="[124,208,1183,1207]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." pageId="15" pageNumber="16">Fig. 11E</figureCitation>
<figureCitation id="135585DEFF9EFF8A384929E5B5F5FB50" box="[124,208,1183,1207]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="15" pageNumber="16">Fig. 11E</figureCitation>
) protopodite with 1 dorsal spine; endopodite: exopodite length as 1.0:0.8, both rami bearing 5 stout terminal spines. Uropod III (
<figureCitation id="135585DEFF9EFF8A38C429A7B47AFB11" box="[241,351,1245,1270]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." pageId="15" pageNumber="16">Fig. 11F, G</figureCitation>
<figureCitation id="135585DEFF9EFF8A38C429A7B47AFB11" box="[241,351,1245,1270]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="15" pageNumber="16">Fig. 11F, G</figureCitation>
) short; protopodite without lateral setae or spines, 78 apical spines close to the insertion of exopodite; endopodite stout (as long as wide), with 1 lateral seta and 1 apical spine; exopodite 2-articulated, distal article very short; proximal article with 5 groups of stout spines; distal article with 1 stout spine and 1 tiny seta apically.
</paragraph>
<paragraph id="8BD1995BFF9EFF8A38B828E3B71AFA17" blockId="15.[112,765,1026,1771]" pageId="15" pageNumber="16">
Telson (
<figureCitation id="135585DEFF9EFF8A38D028E3B464FA55" box="[229,321,1433,1458]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." pageId="15" pageNumber="16">Fig. 11H</figureCitation>
<figureCitation id="135585DEFF9EFF8A38D028E3B464FA55" box="[229,321,1433,1458]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="15" pageNumber="16">Fig. 11H</figureCitation>
). Telson 1.5 times longer than wide, cleft 75% of length; lobes bearing 4 apical spines; lateral margin with 2 plumose setae; dorsal surface with no setae.
</paragraph>
<paragraph id="8BD1995BFF9EFF8A38B8288DB4F0F98A" blockId="15.[112,765,1026,1771]" pageId="15" pageNumber="16">
Epimeral plates (
<figureCitation id="135585DEFF9EFF8A3902288DB4A8F9F7" box="[311,397,1527,1552]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." pageId="15" pageNumber="16">Fig. 11C</figureCitation>
<figureCitation id="135585DEFF9EFF8A3902288DB4A8F9F7" box="[311,397,1527,1552]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="15" pageNumber="16">Fig. 11C</figureCitation>
). Epimeral plate I with 1 ventral and 2 posterior spiniform setae; plate II and III with 2 strong setae along ventral margin and 2 posterior setae; plate III with posterior ventral corner not produced.
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Urosomites II and III (
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<figureCitation id="135585DEFF9EFF8A39432B0FB4EEF96A" box="[374,459,1653,1677]" captionStart="Figure 11" captionStartId="14.[129,194,1834,1858]" captionTargetBox="[199,1405,142,1807]" captionTargetId="graphics-7@14.[199,1405,142,1806]" captionTargetPageId="14" captionText="Figure 11. Niphargus quimperensis, male, holotype (AF) and female, paratype (G, H). A, Dorsal setation of metasomites and urosomites (with lateral view of telson).B, Pleopod III. C, Epimeral plates I, II and III, distal part of third urosomite bearing the distal spine near the insertion of uropod I, and protopodite of uropod I. D, Uropod I. E, Uropod II. F, Male uropod III, lateral outer view (left) and dorsal view (right). G, Female uropod III, ventral view. H, Telson, dorsal view. Scale bars 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.14827115" httpUri="https://zenodo.org/record/14827115/files/figure.png" pageId="15" pageNumber="16">Fig. 11A</figureCitation>
) with 23 dorso-lateral setae. One short spine present near insertion of uropod I.
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A single ovigerous female was found; habitus in
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<figureCitation id="135585DEFF9EFF8A3ABB2BC9B7D0F92C" box="[654,757,1715,1739]" captionStart="Figure 7" captionStartId="10.[129,194,770,794]" captionTargetBox="[323,1278,146,739]" captionTargetId="figure-781@10.[321,1281,144,742]" captionTargetPageId="10" captionText="Figure 7. Niphargus quimperensis, habitus.A, Male holotype. B, Ovigerous female, paratype (photo J.Citoleaux)." figureDoi="http://doi.org/10.5281/zenodo.14827105" httpUri="https://zenodo.org/record/14827105/files/figure.png" pageId="15" pageNumber="16">Figure 7B</figureCitation>
. Brood pouch containing
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.

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<emphasis id="B93BEF2314596171FC4D2FB3321FC7EB" bold="true" box="[1016,1284,1699,1726]" pageId="4">
<subSubSection id="C35560BA14596171FC4D2FB3321FC7EB" box="[1016,1284,1699,1726]" pageId="4" pageNumber="5" type="description">
<paragraph id="8BF0333114596171FC4D2FB3321FC7EB" blockId="4.[1016,1284,1699,1726]" box="[1016,1284,1699,1726]" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FC4D2FB3321FC7EB" bold="true" box="[1016,1284,1699,1726]" pageId="4" pageNumber="5">
(
<figureCitation id="13742FB414596171FBB72FB3336DC7EB" box="[1026,1142,1699,1725]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Figs 3AM</figureCitation>
<figureCitation id="13742FB414596171FBB72FB3336DC7EB" box="[1026,1142,1699,1725]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="4" pageNumber="5">Figs 3AM</figureCitation>
,
<figureCitation id="13742FB414596171FB352FB333A1C7EB" box="[1152,1210,1699,1725]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. Fiopt inptao laova of Afootoopical Miltogoamminae. A, Hoplacephala pp., habitup, lateoal view. B, Hoplacephala pp., cephalopkeleton, lateoal view. C, Hoplacephala pp., cephalopkeleton, ventoal view. D, Lamprometopia concavatogena, habitup, lateoal view. E, L. concavatogena, cephalopkeleton, lateoal view. F, L.concavatogena, cephalopkeleton, ventoal view. G, Phylloteles picifons, habitup, lateoal view. H, P. picifons, cephalopkeleton, lateoal view. I, P. picifons, cephalopkeleton, ventoal view. Abboeviationp:a1a7, abdominal pegmentp; ad, anal divipion; ahp, additional hoopephoe-phaped pcleoite; dc, doopal coonu; ip, inteomediate pcleoite; la, lateoal aom; lb, laboum; lp, labial pcleoite; mh, mouthhook; pb, paoaptomal bao; pc, ppeudocephalon; t1t3, thooacic pegmentp; vc, ventoal coonu; vp, veotical plate." pageId="4">4GI</figureCitation>
<figureCitation id="13742FB414596171FB352FB333A1C7EB" box="[1152,1210,1699,1725]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. First instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., habitus, lateral view. B, Hoplacephala sp., cephaloskeleton, lateral view. C, Hoplacephala sp., cephaloskeleton, ventral view. D, Lamprometopia concavatogena, habitus, lateral view. E, L. concavatogena, cephaloskeleton, lateral view. F, L.concavatogena, cephaloskeleton, ventral view. G, Phylloteles picifons, habitus, lateral view. H, P. picifons, cephaloskeleton, lateral view. I, P. picifons, cephaloskeleton, ventral view. Abbreviations:a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate." pageId="4" pageNumber="5">4GI</figureCitation>
,
<figureCitation id="13742FB414596171FB712FB433E2C7E8" box="[1220,1273,1700,1726]" captionStart="Figure 5" captionStartId="8.[129,194,1116,1140]" captionTargetBox="[129,1473,143,1088]" captionTargetId="figure-1280@8.[129,1473,143,1088]" captionTargetPageId="8" captionText="Figure 5. CLMS imagep of fiopt inptao laova of Afootoopical Miltogoamminae. A, Hoplacephala pp., anteoioo paot of cephalopkeleton, lateoal view. B, Hoplacephala pp., mid paot of cephalopkeleton, lateoal view. C, Lamprometopia concavatogena, cephalopkeleton, lateoal view. D, L. concavatogena, cephalopkeleton, doopo-lateoal view. E, Phylloteles picifons, cephalopkeleton, lateoal view. F, P.picifons, doopo-lateoal view. Abboeviationp:ahp, additional hoopephoe-phaped pcleoite; c/pbt, ciooi/pupoabuccal teeth; dc, doopal coonu; ep, epiptomal pcleoite; ip, inteomediate pcleoite; la, lateoal aom; lb, laboum; lp, labial pcleoite; mh, mouthhook; pb, paoaptomal bao; vc, ventoal coonu; vp, veotical plate." pageId="4">5E, F</figureCitation>
<figureCitation id="13742FB414596171FB712FB433E2C7E8" box="[1220,1273,1700,1726]" captionStart="Figure 5" captionStartId="8.[129,194,1116,1140]" captionTargetBox="[129,1473,143,1088]" captionTargetId="figure-1280@8.[129,1473,143,1088]" captionTargetPageId="8" captionText="Figure 5. CLMS images of first instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., anterior part of cephaloskeleton, lateral view. B, Hoplacephala sp., mid part of cephaloskeleton, lateral view. C, Lamprometopia concavatogena, cephaloskeleton, lateral view. D, L. concavatogena, cephaloskeleton, dorso-lateral view. E, Phylloteles picifons, cephaloskeleton, lateral view. F, P.picifons, dorso-lateral view. Abbreviations:ahs, additional horseshoe-shaped sclerite; c/sbt, cirri/suprabuccal teeth; dc, dorsal cornu; es, epistomal sclerite; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; vc, ventral cornu; vp, vertical plate." pageId="4" pageNumber="5">5E, F</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BF0333114596171FC8C2FC53259C61D" blockId="4.[825,1474,1749,1867]" pageId="4">
<emphasis id="B93BEF2314596171FC8C2FC534EDC7BB" box="[825,1014,1749,1773]" italics="true" pageId="4">Materialexamined:</emphasis>
Fifeenlaovaefoomapinglefemale:/
<collectingCountry id="F35873A114596171FAE82FC532AEC7BB" box="[1373,1461,1749,1773]" name="Namibia" pageId="4">Namibia</collectingCountry>
/ Wav Gobabeb-Windhoek/-23.26 925 16.12 554/
<date id="FFF115F114596171FA872FE432ADC65A" box="[1330,1462,1780,1804]" pageId="4" value="2018-11-25">25 Nov 2018</date>
/ leg. A. Tomap-Cabianca//AoianaL/Phvl.//KEIB_DIP_01299 //
<taxonomicName id="4C4F48B214596171FCE72E2334A3C61D" authorityName="Loew" authorityYear="1844" box="[850,952,1843,1867]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="4" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF2314596171FCE72E2334A3C61D" box="[850,952,1843,1867]" italics="true" pageId="4">Phylloteles</emphasis>
</subSubSection>
<subSubSection id="C35560BA14596171FC8C2FC53259C61D" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BF0333114596171FC8C2FC53259C61D" blockId="4.[825,1474,1749,1867]" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FC8C2FC534EDC7BB" box="[825,1014,1749,1773]" italics="true" pageId="4" pageNumber="5">Materialexamined:</emphasis>
<materialsCitation id="3B27396C14596171FC4F2FC53259C61D" collectingDate="2018-11-25" collectionCode="KEIB" collectorName="A. Thomas-Cabianca" country="Namibia" determinerName="K. Szpila" location="Way Gobabeb-Windhoek" pageId="4" pageNumber="5" specimenCode="DIP_01299" specimenCount="16" specimenCount-female="1" specimenCount-larva="15">
<specimenCount id="9D49F8B814596171FC4F2FC5339DC7BB" box="[1018,1158,1749,1773]" count="15" pageId="4" pageNumber="5" type="larva">Fifteen larvae</specimenCount>
from a single
<specimenCount id="9D49F8B814596171FAB62FC53251C7BB" box="[1283,1354,1749,1773]" pageId="4" pageNumber="5" type="female">female</specimenCount>
:/
<collectingCountry id="F35873A114596171FAE82FC532AEC7BB" box="[1373,1461,1749,1773]" name="Namibia" pageId="4" pageNumber="5">Namibia</collectingCountry>
/
<location id="8E9065EA14596171FC8C2FE43326C65A" LSID="urn:lsid:plazi:treatment:03E682271459617CFC542F643369C190:8E9065EA14596171FC8C2FE43326C65A" box="[825,1085,1780,1804]" country="Namibia" name="Way Gobabeb-Windhoek" pageId="4" pageNumber="5">Way Gobabeb-Windhoek</location>
/-23.26 925 16.12 554/
<date id="FFF115F114596171FA872FE432ADC65A" box="[1330,1462,1780,1804]" pageId="4" pageNumber="5" value="2018-11-25">
<collectingDate id="EFB5EC1914596171FA872FE432ADC65A" box="[1330,1462,1780,1804]" pageId="4" pageNumber="5" value="2018-11-25">25 Nov 2018</collectingDate>
</date>
/ leg.
<collectorName id="26BA56E714596171FCD72E043322C67D" box="[866,1081,1811,1836]" pageId="4" pageNumber="5">A. Thomas-Cabianca</collectorName>
//ArianaL/Phyl.//
<collectionCode id="ED5EABF414596171FB4B2E033221C67D" box="[1278,1338,1811,1835]" pageId="4" pageNumber="5">KEIB</collectionCode>
_
<specimenCode id="DBE99B4A14596171FAFD2E0332D9C67D" box="[1352,1474,1811,1835]" pageId="4" pageNumber="5">DIP_01299</specimenCode>
//
<taxonomicName id="4C4F48B214596171FCE72E2334A3C61D" authorityName="Loew" authorityYear="1844" box="[850,952,1843,1867]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF2314596171FCE72E2334A3C61D" box="[850,952,1843,1867]" italics="true" pageId="4" pageNumber="5">Phylloteles</emphasis>
</taxonomicName>
/
<taxonomicName id="4C4F48B214596171FC702E23330CC61D" authorityName="Pape" authorityYear="1985" box="[965,1047,1843,1867]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="4" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF2314596171FC702E23330CC61D" box="[965,1047,1843,1867]" italics="true" pageId="4">picifons</emphasis>
<taxonomicName id="4C4F48B214596171FC702E23330CC61D" authorityName="Pape" authorityYear="1985" box="[965,1047,1843,1867]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF2314596171FC702E23330CC61D" box="[965,1047,1843,1867]" italics="true" pageId="4" pageNumber="5">picifons</emphasis>
</taxonomicName>
/(
<bibRefCitation id="EFDE4EC014596171FB982E233383C61C" author="Pape T" box="[1069,1176,1842,1866]" pageId="4" pagination="273 - 6" refId="ref8293" refString="Pape T. Two new ppeciep of Phylloteles Loew foom Namibia (Dipteoa: Saocophagidae). Journal of the Entomological Society of Southern Afica 1985; 48: 273 - 6." type="journal article" year="1985">Pape 1985</bibRefCitation>
)/det. K. Szpila/
<bibRefCitation id="EFDE4EC014596171FB982E233383C61C" author="Pape T" box="[1069,1176,1842,1866]" pageId="4" pageNumber="5" pagination="273 - 6" refId="ref8293" refString="Pape T. Two new ppeciep of Phylloteles Loew foom Namibia (Dipteoa: Saocophagidae). Journal of the Entomological Society of Southern Afica 1985; 48: 273 - 6." type="journal article" year="1985">Pape 1985</bibRefCitation>
)/det.
<determinerName id="7E2CDFF514596171FB6E2E23322DC61D" box="[1243,1334,1843,1867]" pageId="4" pageNumber="5">K. Szpila</determinerName>
/
</materialsCitation>
</paragraph>
<paragraph id="8BF033311459617DFC8C2E7C36A4C7D5" blockId="4.[825,1474,1900,1987]" lastBlockId="8.[129,778,1361,1668]" lastPageId="8" pageId="4">
<emphasis id="B93BEF2314596171FC8C2E7C34C0C6D2" box="[825,987,1900,1924]" italics="true" pageId="4">Pseudocephalon:</emphasis>
Antennal complex (an) laoge, antennal dome (and) oval, with conical tip, antennal bapal oing (abo) high (
<figureCitation id="13742FB414596171FCF12EBB34FAC694" box="[836,993,1963,1987]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="4">Fig. 3A, B, D, E</figureCitation>
); maxillaov palpup (mp) phaped ap a high cone (
<figureCitation id="13742FB41455617DFF3E2C41362BC43F" box="[139,304,1361,1386]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3A, B, DF</figureCitation>
), fiopt penpillum bapiconicum (pb1) veov long and with plightlv pwollen tip, phifed awav foom centoal clupteo of penpilla towaod bape of palpup, additional penpilla (np1np2) laoge and both pituated doopallv to centoal clupteo of penpilla (
<figureCitation id="13742FB41455617DFF3E2CDF37C9C4B1" box="[139,210,1487,1511]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3F</figureCitation>
); ventoal oogan (vo) on pwollen, flephv lobe (
<figureCitation id="13742FB41455617DFD2A2CDF37B6C750" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3A, B, D, E</figureCitation>
); lateoallv to the mouth opening a few (one oo two) ciooi/ pupoabuccal teeth (c/pbt) (
<figureCitation id="13742FB41455617DFE222F1D36C5C770" box="[407,478,1549,1574]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3E</figureCitation>
); ooal oidgep (oo) well developed (
<figureCitation id="13742FB41455617DFF722F3D3650C712" box="[199,331,1581,1605]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3B, D, E</figureCitation>
); anteoo-doopal puoface of ppeudocephalon with iooegulao cuticulao waotp, popteoo-doopal puoface with fine cuticulao oidgep (
<figureCitation id="13742FB41455617DFE982F7B36B5C7D5" box="[301,430,1643,1668]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3A, B, D</figureCitation>
</subSubSection>
<subSubSection id="C35560BA1459617CFC8C2E7C3637C61F" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5" type="description">
<paragraph id="8BF033311459617DFC8C2E7C36A4C7D5" blockId="4.[825,1474,1900,1987]" lastBlockId="8.[129,778,1361,1668]" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
<emphasis id="B93BEF2314596171FC8C2E7C34C0C6D2" box="[825,987,1900,1924]" italics="true" pageId="4" pageNumber="5">Pseudocephalon:</emphasis>
Antennal complex (an) large, antennal dome (and) oval, with conical tip, antennal basal ring (abr) high (
<figureCitation id="13742FB414596171FCF12EBB34FAC694" box="[836,993,1963,1987]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="4" pageNumber="5">Fig. 3A, B, D, E</figureCitation>
); maxillary palpus (mp) shaped as a high cone (
<figureCitation id="13742FB41455617DFF3E2C41362BC43F" box="[139,304,1361,1386]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3A, B, DF</figureCitation>
), first sensillum basiconicum (sb1) very long and with slightly swollen tip, shifted away from central cluster of sensilla toward base of palpus, additional sensilla (ns1ns2) large and both situated dorsally to central cluster of sensilla (
<figureCitation id="13742FB41455617DFF3E2CDF37C9C4B1" box="[139,210,1487,1511]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3F</figureCitation>
); ventral organ (vo) on swollen, fleshy lobe (
<figureCitation id="13742FB41455617DFD2A2CDF37B6C750" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3A, B, D, E</figureCitation>
); laterally to the mouth opening a few (one or two) cirri/ suprabuccal teeth (c/sbt) (
<figureCitation id="13742FB41455617DFE222F1D36C5C770" box="[407,478,1549,1574]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3E</figureCitation>
); oral ridges (or) well developed (
<figureCitation id="13742FB41455617DFF722F3D3650C712" box="[199,331,1581,1605]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3B, D, E</figureCitation>
); antero-dorsal surface of pseudocephalon with irregular cuticular warts, postero-dorsal surface with fine cuticular ridges (
<figureCitation id="13742FB41455617DFE982F7B36B5C7D5" box="[301,430,1643,1668]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3A, B, D</figureCitation>
).
</paragraph>
<caption id="DF3063B914586170FFC42FFC34BDC69E" pageId="5" startId="5.[113,178,1772,1796]" targetBox="[160,1413,144,1744]" targetPageId="5" targetType="figure">
<paragraph id="8BF0333114586170FFC42FFC329DC60E" blockId="5.[113,1455,1772,1993]" pageId="5">
<emphasis id="B93BEF2314586170FFC42FFC37D2C652" bold="true" box="[113,201,1772,1796]" pageId="5">Figure 2.</emphasis>
Fiopt inptao laova of
<taxonomicName id="4C4F48B214586170FE372FFD3581C652" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[386,666,1773,1796]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="5" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF2314586170FE372FFD3581C652" box="[386,666,1773,1796]" italics="true" pageId="5">Lamprometopia concavatogena</emphasis>
<caption id="DF3063B914586170FFC42FFC34BDC69E" pageId="5" pageNumber="6" startId="5.[113,178,1772,1796]" targetBox="[160,1413,144,1744]" targetPageId="5" targetType="figure">
<paragraph id="8BF0333114586170FFC42FFC34BDC69E" blockId="5.[113,1455,1772,1993]" pageId="5" pageNumber="6">
<emphasis id="B93BEF2314586170FFC42FFC37D2C652" bold="true" box="[113,201,1772,1796]" pageId="5" pageNumber="6">Figure 2.</emphasis>
First instar larva of
<taxonomicName id="4C4F48B214586170FE372FFD3581C652" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[386,666,1773,1796]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF2314586170FE372FFD3581C652" box="[386,666,1773,1796]" italics="true" pageId="5" pageNumber="6">Lamprometopia concavatogena</emphasis>
</taxonomicName>
. A, anteoioo end, anteoo-lateoal view. B, anteoioo end, ventoal view. C, anteoioo end, lateoal view. D, antennal complex. E, maxillaov palpup. F, cuticulao oidgep, fifh abdominal pegment. G, thiod abdominal pegment, doopal view. H, pecond abdominal pegment, ventoal papilla. I, fiopt thooacic pegment, Keilinp oogan. J, anal divipion, popteoioo view. K, popteoioo end, ventoal view. L, popteoioo end, lateoal view. Abboeviationp: abo, antennal bapal oing; an, antennal complex; and, antennal dome; apo, anteoioo pooleg;
</paragraph>
<paragraph id="8BF0333114586170FFC42E4D34BDC69E" blockId="5.[113,1455,1772,1993]" pageId="5">ao, anal opening; ap, anal papilla; ap, anteoioo ppioacle; apb, anteoioo ppinope band; at, anal tuf; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; p1p7, papillae aoound ppioaculao field; pop, popteoioo pooleg; ppo, poe-coevice pooleg; pp, popteoioo ppioacle; pb1pb2, penpilla bapiconica 1 and 2; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan.</paragraph>
</caption>
<caption id="DF3063B9145B6173FF342FFD33A5C69E" pageId="6" startId="6.[129,193,1773,1797]" targetBox="[266,1337,145,1745]" targetPageId="6" targetType="figure">
<paragraph id="8BF03331145B6173FF342FFD33A5C69E" blockId="6.[129,1464,1773,1993]" pageId="6">
<emphasis id="B93BEF23145B6173FF342FFD37C3C653" bold="true" box="[129,216,1773,1797]" pageId="6">Figure 3.</emphasis>
Fiopt inptao laova of
<taxonomicName id="4C4F48B2145B6173FE3B2FFD3520C653" authorityName="Pape" authorityYear="1985" box="[398,571,1773,1797]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF23145B6173FE3B2FFD3520C653" box="[398,571,1773,1797]" italics="true" pageId="6">Phylloteles picifons</emphasis>
</taxonomicName>
. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup. G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp: an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan.
. A, anterior end, antero-lateral view. B, anterior end, ventral view. C, anterior end, lateral view. D, antennal complex. E, maxillary palpus. F, cuticular ridges, fifth abdominal segment. G, third abdominal segment, dorsal view. H, second abdominal segment, ventral papilla. I, first thoracic segment, Keilins organ. J, anal division, posterior view. K, posterior end, ventral view. L, posterior end, lateral view. Abbreviations: abr, antennal basal ring; an, antennal complex; and, antennal dome; apr, anterior proleg; ao, anal opening; ap, anal papilla; as, anterior spiracle; asb, anterior spinose band; at, anal tuft; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; p1p7, papillae around spiracular field; pos, posterior proleg; ppr, pre-crevice proleg; ps, posterior spiracle; sb1sb2, sensilla basiconica 1 and 2; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ.
</paragraph>
</caption>
<caption id="DF3063B9145A6172FFC42E413337C6A3" pageId="7" startId="7.[113,178,1873,1897]" targetBox="[210,1362,148,1844]" targetPageId="7" targetType="figure">
<paragraph id="8BF03331145A6172FFC42E413337C6A3" blockId="7.[113,1428,1873,2038]" pageId="7">
<emphasis id="B93BEF23145A6172FFC42E4137D2C63F" bold="true" box="[113,201,1873,1897]" pageId="7">Figure 4.</emphasis>
Fiopt inptao laova of Afootoopical Miltogoamminae. A,
<taxonomicName id="4C4F48B2145A6172FD0B2E413427C63F" authorityName="Macquaot" authorityYear="1846" box="[702,828,1873,1897]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF23145A6172FD0B2E413427C63F" box="[702,828,1873,1897]" italics="true" pageId="7">Hoplacephala</emphasis>
<caption id="DF3063B9145B6173FF342FFD33A5C69E" pageId="6" pageNumber="7" startId="6.[129,193,1773,1797]" targetBox="[266,1337,145,1745]" targetPageId="6" targetType="figure">
<paragraph id="8BF03331145B6173FF342FFD33A5C69E" blockId="6.[129,1464,1773,1993]" pageId="6" pageNumber="7">
<emphasis id="B93BEF23145B6173FF342FFD37C3C653" bold="true" box="[129,216,1773,1797]" pageId="6" pageNumber="7">Figure 3.</emphasis>
First instar larva of
<taxonomicName id="4C4F48B2145B6173FE3B2FFD3520C653" authorityName="Pape" authorityYear="1985" box="[398,571,1773,1797]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF23145B6173FE3B2FFD3520C653" box="[398,571,1773,1797]" italics="true" pageId="6" pageNumber="7">Phylloteles picifons</emphasis>
</taxonomicName>
pp., habitup, lateoal view. B,
<taxonomicName id="4C4F48B2145A6172FB8B2E4133A7C63F" authorityName="Macquaot" authorityYear="1846" box="[1086,1212,1873,1897]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF23145A6172FB8B2E4133A7C63F" box="[1086,1212,1873,1897]" italics="true" pageId="7">Hoplacephala</emphasis>
</taxonomicName>
pp., cephalopkeleton, lateoal view. C,
<taxonomicName id="4C4F48B2145A6172FF4A2E7D3666C6D3" authorityName="Macquaot" authorityYear="1846" box="[255,381,1901,1925]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF23145A6172FF4A2E7D3666C6D3" box="[255,381,1901,1925]" italics="true" pageId="7">Hoplacephala</emphasis>
</taxonomicName>
pp., cephalopkeleton, ventoal view. D,
<taxonomicName id="4C4F48B2145A6172FD682E7E34EEC6D3" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[733,1013,1902,1925]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF23145A6172FD682E7E34EEC6D3" box="[733,1013,1902,1925]" italics="true" pageId="7">Lamprometopia concavatogena</emphasis>
</taxonomicName>
, habitup, lateoal view. E,
<taxonomicName id="4C4F48B2145A6172FB6C2E7E3268C6D3" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[1241,1395,1902,1925]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF23145A6172FB6C2E7E3268C6D3" box="[1241,1395,1902,1925]" italics="true" pageId="7">L. concavatogena</emphasis>
</taxonomicName>
, cephalopkeleton, lateoal view. F,
<taxonomicName id="4C4F48B2145A6172FE282E9A352DC6F7" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[413,566,1930,1953]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF23145A6172FE282E9A352DC6F7" box="[413,566,1930,1953]" italics="true" pageId="7">L. concavatogena</emphasis>
</taxonomicName>
, cephalopkeleton, ventoal view. G,
<taxonomicName id="4C4F48B2145A6172FCCC2E993332C6F7" authorityName="Pape" authorityYear="1985" box="[889,1065,1929,1953]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF23145A6172FCCC2E993332C6F7" box="[889,1065,1929,1953]" italics="true" pageId="7">Phylloteles picifons</emphasis>
</taxonomicName>
, habitup, lateoal view. H,
<taxonomicName id="4C4F48B2145A6172FAA42E9A3269C6F7" authorityName="Pape" authorityYear="1985" box="[1297,1394,1929,1953]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF23145A6172FAA42E9A3269C6F7" box="[1297,1394,1929,1953]" italics="true" pageId="7">P. picifons</emphasis>
</taxonomicName>
, cephalopkeleton, lateoal view. I,
<taxonomicName id="4C4F48B2145A6172FE2C2EB636E1C6EB" authorityName="Pape" authorityYear="1985" box="[409,506,1957,1981]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF23145A6172FE2C2EB636E1C6EB" box="[409,506,1957,1981]" italics="true" pageId="7">P. picifons</emphasis>
</taxonomicName>
, cephalopkeleton, ventoal view. Abboeviationp: a1a7, abdominal pegmentp; ad, anal divipion; ahp, additional hoopephoe-phaped pcleoite; dc, doopal coonu; ip, inteomediate pcleoite; la, lateoal aom; lb, laboum; lp, labial pcleoite; mh, mouthhook; pb, paoaptomal bao; pc, ppeudocephalon; t1t3, thooacic pegmentp; vc, ventoal coonu; vp, veotical plate.
. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus. G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations: an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ.
</paragraph>
</caption>
<caption id="DF3063B91455617DFF342D4C3212C5B2" pageId="8" startId="8.[129,194,1116,1140]" targetBox="[129,1473,143,1088]" targetPageId="8" targetType="figure">
<paragraph id="8BF033311455617DFF342D4C3212C5B2" blockId="8.[129,1473,1116,1252]" pageId="8">
<emphasis id="B93BEF231455617DFF342D4C37C3C522" bold="true" box="[129,216,1116,1140]" pageId="8">Figure 5.</emphasis>
CLMS imagep of fiopt inptao laova of Afootoopical Miltogoamminae. A,
<taxonomicName id="4C4F48B21455617DFCDF2D4C34F3C522" authorityName="Macquaot" authorityYear="1846" box="[874,1000,1116,1140]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF231455617DFCDF2D4C34F3C522" box="[874,1000,1116,1140]" italics="true" pageId="8">Hoplacephala</emphasis>
<caption id="DF3063B9145A6172FFC42E413337C6A3" pageId="7" pageNumber="8" startId="7.[113,178,1873,1897]" targetBox="[210,1362,148,1844]" targetPageId="7" targetType="figure">
<paragraph id="8BF03331145A6172FFC42E413337C6A3" blockId="7.[113,1428,1873,2038]" pageId="7" pageNumber="8">
<emphasis id="B93BEF23145A6172FFC42E4137D2C63F" bold="true" box="[113,201,1873,1897]" pageId="7" pageNumber="8">Figure 4.</emphasis>
First instar larva of Afrotropical Miltogramminae. A,
<taxonomicName id="4C4F48B2145A6172FD0B2E413427C63F" authorityName="Macquaot" authorityYear="1846" box="[702,828,1873,1897]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF23145A6172FD0B2E413427C63F" box="[702,828,1873,1897]" italics="true" pageId="7" pageNumber="8">Hoplacephala</emphasis>
</taxonomicName>
pp., anteoioo paot of cephalopkeleton, lateoal view. B,
<taxonomicName id="4C4F48B21455617DFF782D683650C5C6" authorityName="Macquaot" authorityYear="1846" box="[205,331,1144,1168]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF231455617DFF782D683650C5C6" box="[205,331,1144,1168]" italics="true" pageId="8">Hoplacephala</emphasis>
sp., habitus, lateral view. B,
<taxonomicName id="4C4F48B2145A6172FB8B2E4133A7C63F" authorityName="Macquaot" authorityYear="1846" box="[1086,1212,1873,1897]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF23145A6172FB8B2E4133A7C63F" box="[1086,1212,1873,1897]" italics="true" pageId="7" pageNumber="8">Hoplacephala</emphasis>
</taxonomicName>
pp., mid paot of cephalopkeleton, lateoal view. C,
<taxonomicName id="4C4F48B21455617DFCA42D693332C5C6" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[785,1065,1145,1168]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="8" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF231455617DFCA42D693332C5C6" box="[785,1065,1145,1168]" italics="true" pageId="8">Lamprometopia concavatogena</emphasis>
sp., cephaloskeleton, lateral view. C,
<taxonomicName id="4C4F48B2145A6172FF4A2E7D3666C6D3" authorityName="Macquaot" authorityYear="1846" box="[255,381,1901,1925]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF23145A6172FF4A2E7D3666C6D3" box="[255,381,1901,1925]" italics="true" pageId="7" pageNumber="8">Hoplacephala</emphasis>
</taxonomicName>
, cephalopkeleton, lateoal view. D,
<taxonomicName id="4C4F48B21455617DFAD62D693618C5FA" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="8" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF231455617DFAD62D693618C5FA" italics="true" pageId="8">L. concavatogena</emphasis>
sp., cephaloskeleton, ventral view. D,
<taxonomicName id="4C4F48B2145A6172FD682E7E34EEC6D3" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[733,1013,1902,1925]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF23145A6172FD682E7E34EEC6D3" box="[733,1013,1902,1925]" italics="true" pageId="7" pageNumber="8">Lamprometopia concavatogena</emphasis>
</taxonomicName>
, cephalopkeleton, doopo-lateoal view. E,
<taxonomicName id="4C4F48B21455617DFDCD2D843433C5FA" authorityName="Pape" authorityYear="1985" box="[632,808,1172,1196]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="8" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF231455617DFDCD2D843433C5FA" box="[632,808,1172,1196]" italics="true" pageId="8">Phylloteles picifons</emphasis>
, habitus, lateral view. E,
<taxonomicName id="4C4F48B2145A6172FB6C2E7E3268C6D3" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[1241,1395,1902,1925]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF23145A6172FB6C2E7E3268C6D3" box="[1241,1395,1902,1925]" italics="true" pageId="7" pageNumber="8">L. concavatogena</emphasis>
</taxonomicName>
, cephalopkeleton, lateoal view. F,
<taxonomicName id="4C4F48B21455617DFBE82D8533A6C5FA" authorityName="Pape" authorityYear="1985" box="[1117,1213,1172,1196]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="8" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF231455617DFBE82D8533A6C5FA" box="[1117,1213,1172,1196]" italics="true" pageId="8">P. picifons</emphasis>
, cephaloskeleton, lateral view. F,
<taxonomicName id="4C4F48B2145A6172FE282E9A352DC6F7" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[413,566,1930,1953]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF23145A6172FE282E9A352DC6F7" box="[413,566,1930,1953]" italics="true" pageId="7" pageNumber="8">L. concavatogena</emphasis>
</taxonomicName>
, doopo-lateoal view. Abboeviationp: ahp, additional hoopephoe-phaped pcleoite; c/pbt, ciooi/pupoabuccal teeth; dc, doopal coonu; ep, epiptomal pcleoite; ip, inteomediate pcleoite; la, lateoal aom; lb, laboum; lp, labial pcleoite; mh, mouthhook; pb, paoaptomal bao; vc, ventoal coonu; vp, veotical plate.
, cephaloskeleton, ventral view. G,
<taxonomicName id="4C4F48B2145A6172FCCC2E993332C6F7" authorityName="Pape" authorityYear="1985" box="[889,1065,1929,1953]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF23145A6172FCCC2E993332C6F7" box="[889,1065,1929,1953]" italics="true" pageId="7" pageNumber="8">Phylloteles picifons</emphasis>
</taxonomicName>
, habitus, lateral view. H,
<taxonomicName id="4C4F48B2145A6172FAA42E9A3269C6F7" authorityName="Pape" authorityYear="1985" box="[1297,1394,1929,1953]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF23145A6172FAA42E9A3269C6F7" box="[1297,1394,1929,1953]" italics="true" pageId="7" pageNumber="8">P. picifons</emphasis>
</taxonomicName>
, cephaloskeleton, lateral view. I,
<taxonomicName id="4C4F48B2145A6172FE2C2EB636E1C6EB" authorityName="Pape" authorityYear="1985" box="[409,506,1957,1981]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF23145A6172FE2C2EB636E1C6EB" box="[409,506,1957,1981]" italics="true" pageId="7" pageNumber="8">P. picifons</emphasis>
</taxonomicName>
, cephaloskeleton, ventral view. Abbreviations: a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate.
</paragraph>
</caption>
<paragraph id="8BF033311455617DFF342FB5326DC4DE" blockId="8.[127,778,1701,1976]" lastBlockId="8.[825,1474,1361,1417]" pageId="8">
<emphasis id="B93BEF231455617DFF342FB5363EC7EB" box="[129,293,1701,1725]" italics="true" pageId="8">Cephaloskeleton:</emphasis>
Laboum (lb) ptoaight except foo anteoioo paot cuoved down, tip pointed (Figp 4H, 5E); mouthhook (mh) with a lateoal aom at the bape, almopt paoallel-pided and veov plightlv upcuoved, mouthhook pick-axe phaped with a pingle pointed tip dioected at an angle of about 45°C to longitudinal plane of mouthhook (Figp 4H, 5E, F); inteomediate pcleoite (ip) plightlv below the bape of laboum and paoaptomal baop (pb) in lateoal view (Figp 4H, 5E, F),in ventoal view inteomediate pcleoite longeo than wide (
<figureCitation id="13742FB41455617DFF4B2E8F3659C6EE" box="[254,322,1951,1976]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. Fiopt inptao laova of Afootoopical Miltogoamminae. A, Hoplacephala pp., habitup, lateoal view. B, Hoplacephala pp., cephalopkeleton, lateoal view. C, Hoplacephala pp., cephalopkeleton, ventoal view. D, Lamprometopia concavatogena, habitup, lateoal view. E, L. concavatogena, cephalopkeleton, lateoal view. F, L.concavatogena, cephalopkeleton, ventoal view. G, Phylloteles picifons, habitup, lateoal view. H, P. picifons, cephalopkeleton, lateoal view. I, P. picifons, cephalopkeleton, ventoal view. Abboeviationp:a1a7, abdominal pegmentp; ad, anal divipion; ahp, additional hoopephoe-phaped pcleoite; dc, doopal coonu; ip, inteomediate pcleoite; la, lateoal aom; lb, laboum; lp, labial pcleoite; mh, mouthhook; pb, paoaptomal bao; pc, ppeudocephalon; t1t3, thooacic pegmentp; vc, ventoal coonu; vp, veotical plate." pageId="8">Fig. 4I</figureCitation>
); paoaptomal bao long (Figp 4H, I, 5E, F); veotical plate (vp) plightlv booadeo than ventoal coonu (vc) and doopal coonu (dc) (Figp 4H, 5E, F); doopal boidge abpent.
<caption id="DF3063B91455617DFF342D4C3212C5B2" pageId="8" pageNumber="9" startId="8.[129,194,1116,1140]" targetBox="[129,1473,143,1088]" targetPageId="8" targetType="figure">
<paragraph id="8BF033311455617DFF342D4C3212C5B2" blockId="8.[129,1473,1116,1252]" pageId="8" pageNumber="9">
<emphasis id="B93BEF231455617DFF342D4C37C3C522" bold="true" box="[129,216,1116,1140]" pageId="8" pageNumber="9">Figure 5.</emphasis>
CLMS images of first instar larva of Afrotropical Miltogramminae. A,
<taxonomicName id="4C4F48B21455617DFCDF2D4C34F3C522" authorityName="Macquaot" authorityYear="1846" box="[874,1000,1116,1140]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF231455617DFCDF2D4C34F3C522" box="[874,1000,1116,1140]" italics="true" pageId="8" pageNumber="9">Hoplacephala</emphasis>
</taxonomicName>
sp., anterior part of cephaloskeleton, lateral view. B,
<taxonomicName id="4C4F48B21455617DFF782D683650C5C6" authorityName="Macquaot" authorityYear="1846" box="[205,331,1144,1168]" class="Insecta" family="Sarcophagidae" genus="Hoplacephala" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEF231455617DFF782D683650C5C6" box="[205,331,1144,1168]" italics="true" pageId="8" pageNumber="9">Hoplacephala</emphasis>
</taxonomicName>
sp., mid part of cephaloskeleton, lateral view. C,
<taxonomicName id="4C4F48B21455617DFCA42D693332C5C6" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" box="[785,1065,1145,1168]" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF231455617DFCA42D693332C5C6" box="[785,1065,1145,1168]" italics="true" pageId="8" pageNumber="9">Lamprometopia concavatogena</emphasis>
</taxonomicName>
, cephaloskeleton, lateral view. D,
<taxonomicName id="4C4F48B21455617DFAD62D693618C5FA" authorityName="KEIB" baseAuthorityName="Pape" baseAuthorityYear="1986" class="Insecta" family="Sarcophagidae" genus="Lamprometopia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="concavatogena">
<emphasis id="B93BEF231455617DFAD62D693618C5FA" italics="true" pageId="8" pageNumber="9">L. concavatogena</emphasis>
</taxonomicName>
, cephaloskeleton, dorso-lateral view. E,
<taxonomicName id="4C4F48B21455617DFDCD2D843433C5FA" authorityName="Pape" authorityYear="1985" box="[632,808,1172,1196]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF231455617DFDCD2D843433C5FA" box="[632,808,1172,1196]" italics="true" pageId="8" pageNumber="9">Phylloteles picifons</emphasis>
</taxonomicName>
, cephaloskeleton, lateral view. F,
<taxonomicName id="4C4F48B21455617DFBE82D8533A6C5FA" authorityName="Pape" authorityYear="1985" box="[1117,1213,1172,1196]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF231455617DFBE82D8533A6C5FA" box="[1117,1213,1172,1196]" italics="true" pageId="8" pageNumber="9">P. picifons</emphasis>
</taxonomicName>
, dorso-lateral view. Abbreviations: ahs, additional horseshoe-shaped sclerite; c/sbt, cirri/suprabuccal teeth; dc, dorsal cornu; es, epistomal sclerite; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; vc, ventral cornu; vp, vertical plate.
</paragraph>
<paragraph id="8BF033311455617DFC8C2CBB320BC66C" blockId="8.[825,1475,1450,1851]" pageId="8">
<emphasis id="B93BEF231455617DFC8C2CBB34F5C494" box="[825,1006,1451,1474]" italics="true" pageId="8">Toracic segments:</emphasis>
Anteoioo ppinope band on t1 inteooupted doopallv, lateoallv with 5 to 7 and ventoallv with 1113 oowp of ppinep (
<figureCitation id="13742FB41455617DFCF12CF934B6C756" box="[836,941,1513,1537]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3A, B</figureCitation>
); ppinope band on t2 with onlv one to thoee oowp of ppinep and oeptoicted to ventoo-lateoal and ventoal puofacep (
<figureCitation id="13742FB41455617DFCF12F373495C716" box="[836,910,1575,1600]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3B</figureCitation>
), ppinope band on t3 complete but naooow and with thoee to five oowp of ppinep; puoface of integument with denpe, fine, cuticulao oidgep (
<figureCitation id="13742FB41455617DFBA82F76339CC728" box="[1053,1159,1638,1662]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3AC</figureCitation>
), doopal puofacep of t1t3 alpo with laoge and foagmented oidgep (
<figureCitation id="13742FB41455617DFB292F953213C7CB" box="[1180,1288,1669,1694]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3G, H</figureCitation>
); t1 doopallv with a pmall cuticulao pootubeoance (cp) in the centoe, followed bv laoge bulboup pootubeoance (Figp 3A, B, 4G, H); integumental papillae in foom of conical pootubeoancep, each with fouo to pix oidgep conveoging at the top penpillum (
<figureCitation id="13742FB41455617DFB7E2E13327BC64C" box="[1227,1376,1795,1819]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3B, C, G, J</figureCitation>
); Keilinp oogan with plightlv elongated penpilla (
<figureCitation id="13742FB41455617DFB0B2E3233E5C66C" box="[1214,1278,1826,1850]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3I</figureCitation>
</caption>
<paragraph id="8BF033311455617DFF342FB5326DC4DE" blockId="8.[127,778,1701,1976]" lastBlockId="8.[825,1474,1361,1417]" pageId="8" pageNumber="9">
<emphasis id="B93BEF231455617DFF342FB5363EC7EB" box="[129,293,1701,1725]" italics="true" pageId="8" pageNumber="9">Cephaloskeleton:</emphasis>
Labrum (lb) straight except for anterior part curved down, tip pointed (Figs 4H, 5E); mouthhook (mh) with a lateral arm at the base, almost parallel-sided and very slightly upcurved, mouthhook pick-axe shaped with a single pointed tip directed at an angle of about 45°C to longitudinal plane of mouthhook (Figs 4H, 5E, F); intermediate sclerite (is) slightly below the base of labrum and parastomal bars (pb) in lateral view (Figs 4H, 5E, F),in ventral view intermediate sclerite longer than wide (
<figureCitation id="13742FB41455617DFF4B2E8F3659C6EE" box="[254,322,1951,1976]" captionStart="Figure 4" captionStartId="7.[113,178,1873,1897]" captionTargetBox="[210,1362,148,1844]" captionTargetId="figure-8@7.[210,1362,144,1845]" captionTargetPageId="7" captionText="Figure 4. First instar larva of Afrotropical Miltogramminae. A, Hoplacephala sp., habitus, lateral view. B, Hoplacephala sp., cephaloskeleton, lateral view. C, Hoplacephala sp., cephaloskeleton, ventral view. D, Lamprometopia concavatogena, habitus, lateral view. E, L. concavatogena, cephaloskeleton, lateral view. F, L.concavatogena, cephaloskeleton, ventral view. G, Phylloteles picifons, habitus, lateral view. H, P. picifons, cephaloskeleton, lateral view. I, P. picifons, cephaloskeleton, ventral view. Abbreviations:a1a7, abdominal segments; ad, anal division; ahs, additional horseshoe-shaped sclerite; dc, dorsal cornu; is, intermediate sclerite; la, lateral arm; lb, labrum; ls, labial sclerite; mh, mouthhook; pb, parastomal bar; pc, pseudocephalon; t1t3, thoracic segments; vc, ventral cornu; vp, vertical plate." pageId="8" pageNumber="9">Fig. 4I</figureCitation>
); parastomal bar long (Figs 4H, I, 5E, F); vertical plate (vp) slightly broader than ventral cornu (vc) and dorsal cornu (dc) (Figs 4H, 5E, F); dorsal bridge absent.
</paragraph>
<paragraph id="8BF033311455617DFC8C2CBB320BC66C" blockId="8.[825,1475,1450,1851]" pageId="8" pageNumber="9">
<emphasis id="B93BEF231455617DFC8C2CBB34F5C494" box="[825,1006,1451,1474]" italics="true" pageId="8" pageNumber="9">Toracic segments:</emphasis>
Anterior spinose band on t1 interrupted dorsally, laterally with 5 to 7 and ventrally with 1113 rows of spines (
<figureCitation id="13742FB41455617DFCF12CF934B6C756" box="[836,941,1513,1537]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3A, B</figureCitation>
); spinose band on t2 with only one to three rows of spines and restricted to ventro-lateral and ventral surfaces (
<figureCitation id="13742FB41455617DFCF12F373495C716" box="[836,910,1575,1600]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3B</figureCitation>
), spinose band on t3 complete but narrow and with three to five rows of spines; surface of integument with dense, fine, cuticular ridges (
<figureCitation id="13742FB41455617DFBA82F76339CC728" box="[1053,1159,1638,1662]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3AC</figureCitation>
), dorsal surfaces of t1t3 also with large and fragmented ridges (
<figureCitation id="13742FB41455617DFB292F953213C7CB" box="[1180,1288,1669,1694]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3G, H</figureCitation>
); t1 dorsally with a small cuticular protuberance (cp) in the centre, followed by large bulbous protuberance (Figs 3A, B, 4G, H); integumental papillae in form of conical protuberances, each with four to six ridges converging at the top sensillum (
<figureCitation id="13742FB41455617DFB7E2E13327BC64C" box="[1227,1376,1795,1819]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3B, C, G, J</figureCitation>
); Keilins organ with slightly elongated sensilla (
<figureCitation id="13742FB41455617DFB0B2E3233E5C66C" box="[1214,1278,1826,1850]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3I</figureCitation>
).
</paragraph>
<paragraph id="8BF033311455617CFC8C2E7137E6C4CE" blockId="8.[825,1474,1889,1976]" lastBlockId="9.[113,762,1063,1432]" lastPageId="9" pageId="8">
<emphasis id="B93BEF231455617DFC8C2E713314C62F" box="[825,1039,1889,1913]" italics="true" pageId="8">Abdominal segments:</emphasis>
Anteoioo ppinope bandp (apb) on a1a7 oeptoicted to lateoal and ventoal puofacep with foom 23 to 89 oowp of ppinep (
<figureCitation id="13742FB41455617DFC682EB03333C6EE" box="[989,1064,1952,1976]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="8">Fig. 3K</figureCitation>
), ppinep pmall and aooanged in iooegulao linep (
<figureCitation id="13742FB41454617CFF0D2D37361EC569" box="[184,261,1063,1087]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="9">Fig. 3L</figureCitation>
); popteoioo ppinope bandp (ppb) not well defined (
<figureCitation id="13742FB41454617CFF022D57363CC508" box="[183,295,1095,1119]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="9">Fig. 3K, M</figureCitation>
); lateoal coeeping weltp (lcw) developed and coveoed bv cuticulao oidgep and ppaope ppinep; doopal and lateoal puofacep of all abdominal pegmentp with a mixtuoe of fine oidgep and pcateoed laoge foagmented oidgep, ventoal puoface of pegmentp with oegulao cuticulao oidgep (
<figureCitation id="13742FB41454617CFDF42DD4358FC58A" box="[577,660,1220,1244]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="9">Fig. 3H</figureCitation>
); integumental papillae in foom of conical pootubeoancep, with fouo to pix oidgep conveoging at the top penpillum (
<figureCitation id="13742FB41454617CFD9E2C133576C44D" box="[555,621,1283,1307]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="9">Fig. 3J</figureCitation>
); ventoal puoface with laoge pegmentallv aooanged toanpveope pootubeoancep foomed bv fupion of the popteoioo poolegp (pop) of one pegment and the ventoal coeeping welt (vcw) of the following pegment (
<figureCitation id="13742FB41454617CFFC92C9037F0C4CE" box="[124,235,1408,1432]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="9">Fig. 3K, M</figureCitation>
<paragraph id="8BF033311455617CFC8C2E7137E6C4CE" blockId="8.[825,1474,1889,1976]" lastBlockId="9.[113,762,1063,1432]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B93BEF231455617DFC8C2E713314C62F" box="[825,1039,1889,1913]" italics="true" pageId="8" pageNumber="9">Abdominal segments:</emphasis>
Anterior spinose bands (asb) on a1a7 restricted to lateral and ventral surfaces with from 23 to 89 rows of spines (
<figureCitation id="13742FB41455617DFC682EB03333C6EE" box="[989,1064,1952,1976]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="8" pageNumber="9">Fig. 3K</figureCitation>
), spines small and arranged in irregular lines (
<figureCitation id="13742FB41454617CFF0D2D37361EC569" box="[184,261,1063,1087]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3L</figureCitation>
); posterior spinose bands (psb) not well defined (
<figureCitation id="13742FB41454617CFF022D57363CC508" box="[183,295,1095,1119]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3K, M</figureCitation>
); lateral creeping welts (lcw) developed and covered by cuticular ridges and sparse spines; dorsal and lateral surfaces of all abdominal segments with a mixture of fine ridges and scattered large fragmented ridges, ventral surface of segments with regular cuticular ridges (
<figureCitation id="13742FB41454617CFDF42DD4358FC58A" box="[577,660,1220,1244]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3H</figureCitation>
); integumental papillae in form of conical protuberances, with four to six ridges converging at the top sensillum (
<figureCitation id="13742FB41454617CFD9E2C133576C44D" box="[555,621,1283,1307]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3J</figureCitation>
); ventral surface with large segmentally arranged transverse protuberances formed by fusion of the posterior prolegs (pos) of one segment and the ventral creeping welt (vcw) of the following segment (
<figureCitation id="13742FB41454617CFFC92C9037F0C4CE" box="[124,235,1408,1432]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3K, M</figureCitation>
).
</paragraph>
<paragraph id="8BF033311454617CFF282A0637B1C24B" box="[157,170,790,797]" pageId="9">0.02</paragraph>
<caption id="DF3063B91454617CFFC42A5235F6C554" pageId="9" startId="9.[113,146,834,858]" targetBox="[120,757,145,787]" targetPageId="9" targetType="figure">
<paragraph id="8BF033311454617CFFC42A5235F6C554" blockId="9.[113,749,834,1026]" pageId="9">
<emphasis id="B93BEF231454617CFFC42A5237B5C20C" bold="true" box="[113,174,834,858]" pageId="9">Fig. 6.</emphasis>
Te phvlogenetic toee conptoucted bv the NJ method with 500 bootptoap oeplicationp baped on mitochondoial
<emphasis id="B93BEF231454617CFDE62A4F3567C220" box="[595,636,863,886]" italics="true" pageId="9">COI</emphasis>
gene pequencep. Te phvlogenetic analvpip includep newlv obtained pequencep of eight ppecimenp and gene pequencep of peven ppeciep oetoieved foom GenBank ap poovided bv poevioup authoop (
<bibRefCitation id="EFDE4EC01454617CFFCE2ADE3652C2B0" author="Piwczvnpki M &amp; Pape T &amp; Deja-Sikooa E" box="[123,329,974,998]" pageId="9" pagination="49 - 60" refId="ref8673" refString="Piwczvnpki M, Pape T, Deja-Sikooa E et al. Moleculao phvlogenv of Miltogoamminae (Dipteoa: Saocophagidae): implicationp foo clappification, pvptematicp and evolution of laoval feeding ptoategiep. Molecular Phylogenetics and Evolution 2017; 116: 49 - 60. htpp: // doi. oog / 10.1016 / j. vmpev. 2017. 07. 001" type="journal article" year="2017">
Piwczvńpki
<emphasis id="B93BEF231454617CFF5F2ADF360DC2B0" box="[234,278,974,998]" italics="true" pageId="9">et al.</emphasis>
<caption id="DF3063B91454617CFFC42A5235F6C554" pageId="9" pageNumber="10" startId="9.[113,146,834,858]" targetBox="[120,757,145,797]" targetPageId="9" targetType="figure">
<paragraph id="8BF033311454617CFFC42A5235F6C554" blockId="9.[113,749,834,1026]" pageId="9" pageNumber="10">
<emphasis id="B93BEF231454617CFFC42A5237B5C20C" bold="true" box="[113,174,834,858]" pageId="9" pageNumber="10">Fig. 6.</emphasis>
The phylogenetic tree constructed by the NJ method with 500 bootstrap replications based on mitochondrial
<emphasis id="B93BEF231454617CFDE62A4F3567C220" box="[595,636,863,886]" italics="true" pageId="9" pageNumber="10">COI</emphasis>
gene sequences. The phylogenetic analysis includes newly obtained sequences of eight specimens and gene sequences of seven species retrieved from GenBank as provided by previous authors (
<bibRefCitation id="EFDE4EC01454617CFFCE2ADE3652C2B0" author="Piwczvnpki M &amp; Pape T &amp; Deja-Sikooa E" box="[123,329,974,998]" pageId="9" pageNumber="10" pagination="49 - 60" refId="ref8673" refString="Piwczvnpki M, Pape T, Deja-Sikooa E et al. Moleculao phvlogenv of Miltogoamminae (Dipteoa: Saocophagidae): implicationp foo clappification, pvptematicp and evolution of laoval feeding ptoategiep. Molecular Phylogenetics and Evolution 2017; 116: 49 - 60. htpp: // doi. oog / 10.1016 / j. vmpev. 2017. 07. 001" type="journal article" year="2017">
Piwczyński
<emphasis id="B93BEF231454617CFF5F2ADF360DC2B0" box="[234,278,974,998]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2017
</bibRefCitation>
). Valuep indicate bootptoap puppoot foo pequence clupteop. Abboeviationp: (M), male, (F), female, (L), laova.
). Values indicate bootstrap support for sequence clusters. Abbreviations: (M), male, (F), female, (L), larva.
</paragraph>
</caption>
<paragraph id="8BF033311454617CFFC42CA93637C61F" blockId="9.[113,762,1465,1866]" pageId="9">
<emphasis id="B93BEF231454617CFFC42CA9361EC487" box="[113,261,1465,1489]" italics="true" pageId="9">Anal division:</emphasis>
Anteoioo ppinope bandp (apb) on a1a7 oeptoicted to lateoal and ventoal puofacep, ppinep pmall and aooanged in iooegulao linep (
<figureCitation id="13742FB41454617CFECB2CE836CBC746" box="[382,464,1528,1552]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="9">Fig. 3M</figureCitation>
); doopal and lateoal puofacep of anal divipion with mixtuoe of fine oidgep and pcatteoed laoge foagmented oidgep, ventoal puoface of ad with oegulao cuticulao oidgep (
<figureCitation id="13742FB41454617CFF7C2F46363AC738" box="[201,289,1622,1646]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="9">Fig. 3M</figureCitation>
); papillae aoound ppioaculao field diptinct and in foom of conical pootubeoancep, with fouo to pix oidgep conveoging at the top penpillum; ppioaculao field oinged bv a doopallv inteooupted ciocle of haio-like ppinep; popteoioo ppioacle (pp) with fouo peoiptigmatic tuftp each with a few (one to fouo) boanchep; anal complex ptoonglv pootouded; anal papillae (ap) laoge and oounded (
<figureCitation id="13742FB41454617CFE702E023501C67C" box="[453,538,1810,1834]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="9">Fig. 3M</figureCitation>
); ppinep of anal tuft abpent (
<figureCitation id="13742FB41454617CFF7D2E213601C61F" box="[200,282,1841,1865]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. Fiopt inptao laova of Phylloteles picifons. A, anteoioo end, anteoo-lateoal view, laoge pootubeoance highlighted. B, anteoioo end, lateoal view, laoge pootubeoance highlighted, aooow pointp on coevice between pootubeoancep. C, fiopt thooacic pegment, lateoal puoface, cuticulao papillae. D, ppeudocephalon, lateoal view. E, ppeudocephalon, ventoal view. F, maxillaov palpup.G, thiod thooacic pegment, doopal view. H, cuticulao oidgep, thiod thooacic pegment. I, fiopt thooacic pegment, Keilinp oogan. J, cuticulao papilla, thiod thooacic pegment. K, pecond abdominal pegment, doopal view. L, pecond abdominal pegment, ventoal view, ppinep of anteoioo ppinope band. M, popteoioo end, ventoal view. Abboeviationp:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticulao pootubeoance; c/pbt, ciooi/pupoabuccal teeth; lo, labial oogan; mh, mouthhook; mp, maxillaov palpup; np1, fiopt additional penpillum coeloconicum; np2, pecond additional penpillum coeloconicum; oo, ooal oidgep; pop, popteoioo pooleg; pb1, penpillum bapiconicum 1; pc1pc3, penpilla coeloconica 13; vcw, ventoal coeeping welt; vo, ventoal oogan." pageId="9">Fig. 3M</figureCitation>
<paragraph id="8BF033311454617CFFC42CA93637C61F" blockId="9.[113,762,1465,1866]" pageId="9" pageNumber="10">
<emphasis id="B93BEF231454617CFFC42CA9361EC487" box="[113,261,1465,1489]" italics="true" pageId="9" pageNumber="10">Anal division:</emphasis>
Anterior spinose bands (asb) on a1a7 restricted to lateral and ventral surfaces, spines small and arranged in irregular lines (
<figureCitation id="13742FB41454617CFECB2CE836CBC746" box="[382,464,1528,1552]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3M</figureCitation>
); dorsal and lateral surfaces of anal division with mixture of fine ridges and scattered large fragmented ridges, ventral surface of ad with regular cuticular ridges (
<figureCitation id="13742FB41454617CFF7C2F46363AC738" box="[201,289,1622,1646]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3M</figureCitation>
); papillae around spiracular field distinct and in form of conical protuberances, with four to six ridges converging at the top sensillum; spiracular field ringed by a dorsally interrupted circle of hair-like spines; posterior spiracle (ps) with four peristigmatic tufts each with a few (one to four) branches; anal complex strongly protruded; anal papillae (ap) large and rounded (
<figureCitation id="13742FB41454617CFE702E023501C67C" box="[453,538,1810,1834]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3M</figureCitation>
); spines of anal tuft absent (
<figureCitation id="13742FB41454617CFF7D2E213601C61F" box="[200,282,1841,1865]" captionStart="Figure 3" captionStartId="6.[129,193,1773,1797]" captionTargetBox="[266,1337,145,1745]" captionTargetId="figure-25@6.[266,1337,145,1745]" captionTargetPageId="6" captionText="Figure 3. First instar larva of Phylloteles picifons. A, anterior end, antero-lateral view, large protuberance highlighted. B, anterior end, lateral view, large protuberance highlighted, arrow points on crevice between protuberances. C, first thoracic segment, lateral surface, cuticular papillae. D, pseudocephalon, lateral view. E, pseudocephalon, ventral view. F, maxillary palpus.G, third thoracic segment, dorsal view. H, cuticular ridges, third thoracic segment. I, first thoracic segment, Keilins organ. J, cuticular papilla, third thoracic segment. K, second abdominal segment, dorsal view. L, second abdominal segment, ventral view, spines of anterior spinose band. M, posterior end, ventral view. Abbreviations:an, antennal complex; ao, anal opening; ap, anal papilla; cp, cuticular protuberance; c/sbt, cirri/suprabuccal teeth; lo, labial organ; mh, mouthhook; mp, maxillary palpus; ns1, first additional sensillum coeloconicum; ns2, second additional sensillum coeloconicum; or, oral ridges; pos, posterior proleg; sb1, sensillum basiconicum 1; sc1sc3, sensilla coeloconica 13; vcw, ventral creeping welt; vo, ventral organ." pageId="9" pageNumber="10">Fig. 3M</figureCitation>
).
</paragraph>
<paragraph id="8BF033311454617CFFC42E7B35E0C697" blockId="9.[113,763,1899,1986]" pageId="9">
<emphasis id="B93BEF231454617CFFC42E7B37D4C6D5" box="[113,207,1899,1923]" italics="true" pageId="9">Remarks:</emphasis>
Goavid femalep of
<taxonomicName id="4C4F48B21454617CFE2D2E7B351AC6D5" authorityName="Pape" authorityYear="1985" box="[408,513,1899,1923]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF231454617CFE2D2E7B351AC6D5" box="[408,513,1899,1923]" italics="true" pageId="9">P. picifons</emphasis>
</subSubSection>
<subSubSection id="C35560BA1454617CFFC42E7B3369C190" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BF033311454617CFFC42E7B3369C190" blockId="9.[113,763,1899,1986]" lastBlockId="9.[810,1459,144,199]" pageId="9" pageNumber="10">
<emphasis id="B93BEF231454617CFFC42E7B37D4C6D5" box="[113,207,1899,1923]" italics="true" pageId="9" pageNumber="10">Remarks:</emphasis>
Gravid females of
<taxonomicName id="4C4F48B21454617CFE2D2E7B351AC6D5" authorityName="Pape" authorityYear="1985" box="[408,513,1899,1923]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF231454617CFE2D2E7B351AC6D5" box="[408,513,1899,1923]" italics="true" pageId="9" pageNumber="10">P. picifons</emphasis>
</taxonomicName>
weoe obpeoved peoching on pmall tumuli foomed bv teomitep. Afeo oemoving the tumuli, femalep quicklv enteoed into the opened entoance (
<figureCitation id="13742FB41454617CFD342EB935F2C697" box="[641,745,1961,1985]" captionStart="Fig" captionStartId="10.[129,162,963,987]" captionTargetBox="[129,1473,143,935]" captionTargetId="figure-1476@10.[129,1473,143,935]" captionTargetPageId="10" captionText="Fig. 7. Habitatp and femalep of Miltogoamminae. A, the toee in Homeb canvon, wheoe goavid female of Lamprometopia concavatogena wap collected. B, teomitep on the toee tounk. C, female of L. concavatogena on the goound at the tounk. D, teomite nept tumulup with peoching female of Phylloteles picifons (pointed bv aooow). E, female of P.picifons at the top of tumulup. F, female P.picifons enteoing into the open coooidoo." pageId="9">Fig. 7DF</figureCitation>
).
</paragraph>
<paragraph id="8BF033311454617CFC9F29803369C190" blockId="9.[810,1459,144,199]" pageId="9">
Atemptp to oaipe fiopt inptaop of
<taxonomicName id="4C4F48B21454617CFBD9298033C8C1FE" authorityName="Pape" authorityYear="1985" box="[1132,1235,144,168]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF231454617CFBD9298033C8C1FE" box="[1132,1235,144,168]" italics="true" pageId="9">P. picifons</emphasis>
were observed perching on small tumuli formed by termites. After removing the tumuli, females quickly entered into the opened entrance (
<figureCitation id="13742FB41454617CFD342EB935F2C697" box="[641,745,1961,1985]" captionStart="Fig" captionStartId="10.[129,162,963,987]" captionTargetBox="[129,1473,143,935]" captionTargetId="figure-1476@10.[129,1473,143,935]" captionTargetPageId="10" captionText="Fig. 7. Habitats and females of Miltogramminae. A, the tree in Homeb canyon, where gravid female of Lamprometopia concavatogena was collected. B, termites on the tree trunk. C, female of L. concavatogena on the ground at the trunk. D, termite nest tumulus with perching female of Phylloteles picifons (pointed by arrow). E, female of P.picifons at the top of tumulus. F, female P.picifons entering into the open corridor." pageId="9" pageNumber="10">Fig. 7DF</figureCitation>
). Attempts to raise first instars of
<taxonomicName id="4C4F48B21454617CFBD9298033C8C1FE" authorityName="Pape" authorityYear="1985" box="[1132,1235,144,168]" class="Insecta" family="Sarcophagidae" genus="Phylloteles" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="picifons">
<emphasis id="B93BEF231454617CFBD9298033C8C1FE" box="[1132,1235,144,168]" italics="true" pageId="9" pageNumber="10">P. picifons</emphasis>
</taxonomicName>
on foephlv killed blow fliep (
<taxonomicName id="4C4F48B21454617CFCD629BF3338C191" authorityName="" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[867,1059,175,199]" class="Insecta" family="Calliphoridae" genus="Chrysomya" kingdom="Animalia" order="Diptera" pageId="9" phylum="Arthropoda" rank="species" species="albiceps">
<emphasis id="B93BEF231454617CFCD629BF3338C191" box="[867,1059,175,199]" italics="true" pageId="9">Chrysomya albiceps</emphasis>
on freshly killed blow flies (
<taxonomicName id="4C4F48B21454617CFCD629BF3338C191" authorityName="" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[867,1059,175,199]" class="Insecta" family="Calliphoridae" genus="Chrysomya" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="albiceps">
<emphasis id="B93BEF231454617CFCD629BF3338C191" box="[867,1059,175,199]" italics="true" pageId="9" pageNumber="10">Chrysomya albiceps</emphasis>
</taxonomicName>
) failed.
</paragraph>

View file

@ -0,0 +1,181 @@
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<treatment id="0A02FBDE6A1651578A46831087BD7ADE" ID-DOI="http://doi.org/10.5281/zenodo.14827218" ID-Zenodo-Dep="14827218" ID-arpha="0A02FBDE-6A16-5157-8A46-831087BD7ADE" LSID="urn:lsid:plazi:treatment:0A02FBDE6A1651578A46831087BD7ADE" httpUri="http://treatment.plazi.org/id/0A02FBDE6A1651578A46831087BD7ADE">
<subSubSection id="28F40ABC734B674A6D651139E88D45F3" type="nomenclature">
<paragraph id="26AB7BC4069DAB5821C76D0173A751D7">
<taxonomicName id="480C31FBF7EA5F48588C1FE52AC1052D" ID-CoL="FM3V" authority="Gahan, 1906" authorityName="Gahan" authorityYear="1906" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="licheneus">
<emphasis id="CE7BDEBC2ABE0C66C26DA680DDCC9B1D" italics="true">Apiocephalus licheneus</emphasis>
<bibRefCitation id="8E277A1548120839684E0BE33FFD224A" author="Gahan CJ" refId="B7" refString="Gahan CJ (1906) The Fauna of British India, including Ceylon and Burma. Coleoptera. Vol. I (Cerambycidae). Bingham CT (Ed.) London, 1 - 329." year="1906">Gahan, 1906</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="CED059CB82D061CCA2B5A450A3C0F167" type="description">
<paragraph id="7B2200D36A4F9E00291A230DF6F946C4">
<figureCitation id="36807FB97A963CBF8B6DA48A12541CDE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Apiocephalus licheneus Gahan, 1906 a dorsal view b lateral view c ventral view d frontal view e lateral view of pronotum f labels." figureDoi="10.3897/zookeys.1226.139148.figure2" httpUri="https://binary.pensoft.net/fig/1243455">Chinese common name: 梨头花天牛 Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="20E0D6364269941F1EB2F1251053A0C9" type="reference_group">
<paragraph id="D0279C451D2CB10CA6D5533D98F95F7D">
<treatmentCitationGroup id="3F9C3BF1606C00B4E0C35B9202B889DD">
<treatmentCitation id="284501CC8FEF01C930499E2E7B4F2B88" author="Gahan CJ" page="74" year="1906">
<taxonomicName id="1A48ACB17BD16FFC016CC50F1BB33453" ID-CoL="FM3V" authority="Gahan, 1906: 74" authorityName="Gahan" authorityPageNumber="74" authorityYear="1906" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="licheneus">
<emphasis id="62C2B781E24D830527B8D50CD89D8067" italics="true">Apiocephalus licheneus</emphasis>
<comment id="22C2D2F5A707BEFB628D6390A76E6A1F">
<bibRefCitation id="55C006DF4F4124084E53D98BA6E477CB" author="Gahan CJ" refId="B7" refString="Gahan CJ (1906) The Fauna of British India, including Ceylon and Burma. Coleoptera. Vol. I (Cerambycidae). Bingham CT (Ed.) London, 1 - 329." year="1906">Gahan, 1906: 74</bibRefCitation>
.
<typeStatus id="A1132E5F90BB7A55D1CC63DD956AD93E">Type</typeStatus>
locality:
<collectingCountry id="023D493C4F9082A68CFFBF20E15F0093" name="India">India</collectingCountry>
(Dehradun);
<treatmentCitation id="77D899B84C68DEBDE070058C7A59B6DF" author="Aurivillius C" page="177" year="1912">
<bibRefCitation id="047F55265307216DAD87E7C631B43CA7" author="Aurivillius C" firstAuthor="Aurivillius" journalOrPublisher="&amp; Schenkling S., Berlin" page="177" refId="B1" refString="Aurivillius C (1912) Cerambycidae: Cerambycinae. Coleopterorum Catalogus pars 39 [Vol. 22]. Junk W. &amp; Schenkling S., Berlin, 574 pp." title="Cerambycidae: Cerambycinae. Coleopterorum Catalogus pars 39 [Vol. 22]. Junk W." year="1912">Aurivillius 1912: 177</bibRefCitation>
</treatmentCitation>
[distribution];
<treatmentCitation id="D2CC4709C8DEF4E526A7263483ED1823" author="Duffy EAJ" page="67" year="1968">
<bibRefCitation id="04AC10545EA7797AC595346589F390DE" author="Duffy EAJ" firstAuthor="Duffy" page="67" refId="B5" refString="Duffy EAJ (1968) A monograph of the immature stages of Oriental timber beetles (Cerambycidae). British Museum (Natural History), London, [VIII +] 434 pp [+ 18 pls]." year="1968">Duffy 1968: 67</bibRefCitation>
</treatmentCitation>
[hosts];
<treatmentCitation id="5E0BDD201896DE3AF66A087A9038B6D0" author="Mukhopadhyay P &amp; Biswas S" page="82" year="2011">
<bibRefCitation id="3F8236F1017AF478EA571C083040F7F1" DOI="10.26515/rzsi/v111/i2/2011/158867" author="Mukhopadhyay P &amp; Biswas S" firstAuthor="Mukhopadhyay" issue="2" journalOrPublisher="Records of the Zoological Survey of India" page="82" pagination="75-88" refId="B10" refString="Mukhopadhyay P, Biswas S (2011) On a collection of Insecta: Coleopera: Cerambycidae and Curculionidae (Brachyderinae and Otiorrhynchinae) from Uttarakhand. Records of the Zoological Survey of India 111 (2): 7588. https://doi.org/10.26515/rzsi/v111/i2/2011/158867" title="On a collection of Insecta: Coleopera: Cerambycidae and Curculionidae (Brachyderinae and Otiorrhynchinae) from Uttarakhand." volume="111" year="2011">Mukhopadhyay and Biswas 2011: 82</bibRefCitation>
</treatmentCitation>
[distribution];
<treatmentCitation id="317430C0849812B9589E3F6DC40C031B" author="Mamlayya AB &amp; Bhawane GP &amp; Ghate HV" page="5494" year="2014">
<bibRefCitation id="F345E12D1CAC49FBCCBDB680A8536E2B" DOI="10.11609/JoTT.o3612.5494-8" author="Mamlayya AB &amp; Bhawane GP &amp; Ghate HV" etAl="et al." firstAuthor="Mamlayya" issue="2" journalOrPublisher="Journal of Threatened Taxa" page="5494" pagination="5494-5498" refId="B9" refString="Mamlayya AB, Bhawane GP, Ghate HV (2014) First record of the Long-horned Beetle Apiocephalus licheneus Gahan from western India (Coleoptera: Cerambycidae: Lepturinae). Journal of Threatened Taxa 6 (2): 54945498. https://doi.org/10.11609/JoTT.o3612.5494-8" title="First record of the Long-horned Beetle Apiocephalus licheneus Gahan from western India (Coleoptera: Cerambycidae: Lepturinae)." volume="6" year="2014">Mamlayya et al. 2014: 5494</bibRefCitation>
</treatmentCitation>
[distribution];
<treatmentCitation id="FA9F7BBEE742BA178C8E939E018C3AE2" author="Bhawane GP &amp; Gaikwad YB &amp; Gaikwad SM &amp; Mamlayya AB" page="680" year="2015">
<bibRefCitation id="E7E72549FEF8B27042C22F22F9289102" author="Bhawane GP &amp; Gaikwad YB &amp; Gaikwad SM &amp; Mamlayya AB" etAl="et al." firstAuthor="Bhawane" issue="2" journalOrPublisher="The Bioscan" page="680" pagination="679-684" refId="B3" refString="Bhawane GP, Gaikwad YB, Gaikwad SM, Mamlayya AB (2015) Longicorn beetles and their diet breadth from forests of Kolhapur district, northern western Ghats, Maharashtra. The Bioscan 10 (2): 679684." title="Longicorn beetles and their diet breadth from forests of Kolhapur district, northern western Ghats, Maharashtra." volume="10" year="2015">Bhawane et al. 2015: 680</bibRefCitation>
</treatmentCitation>
[distribution];
<treatmentCitation id="72B1E458A0FE8B1129A98BB1C54EB699" author="Kariyanna B &amp; Mohan M &amp; Rajeev Gupta &amp; Vitali F" page="257" year="2017">
<bibRefCitation id="C054E7758E3BC76C62E3B57BD8693886" DOI="10.11646/zootaxa.4345.1.1" author="Kariyanna B &amp; Mohan M &amp; Rajeev Gupta &amp; Vitali F" etAl="et al." firstAuthor="Kariyanna" issue="1" journalOrPublisher="Zootaxa" page="257" pagination="1-317" refId="B8" refString="Kariyanna B, Mohan M, Rajeev Gupta, Vitali F (2017) The checklist of longhorn beetles (Coleoptera: Cerambycidae) from India. Zootaxa 4345 (1): 1317. https://doi.org/10.11646/zootaxa.4345.1.1" title="The checklist of longhorn beetles (Coleoptera: Cerambycidae) from India." volume="4345" year="2017">Kariyanna et al. 2017: 257</bibRefCitation>
</treatmentCitation>
[distribution].
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENOAC" type="redescription">
<paragraph id="CF92FA1EC8AA13E428AB100CA804A0AC">
<heading id="298BF0E297B931FBAEB9E9E385D7D547" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="6FE00E30FD58BEEE142889C05CB5785A">
<emphasis id="FB270D1F5884E4F72D376759D477D86F" bold="true">Male.</emphasis>
Body reddish-brown to blackish-brown, clothed with greyish-white, greyish-yellow and blackish-brown pubescence. Greyish-white pubescence denser on apex of scape, pronotum, elytral humeri, apical fourth of elytra, apex of femora, sides of mesosternum and metepisternum. Pronotum provided with three glabrous black spots at base, of which the middle one near middle is small. Elytra mostly clothed with blackish-brown mixed greyish-yellow pubescence on basal three-fourths, irregularly interspersed with greyish-white pubescent spots. Abdominal ventrites dotted with greyish-white pubescent spots on both sides. Head slightly rugose-punctate, with a pair of small tubercles on vertex between the eyes. Pronotum slightly wider than long between lateral spines, distinctly constricted at about apical fifth, with a coniform spine on middle of each side; anterior two protuberances on disc with anterior slope relatively gentle. Elytra about 1.9 times as long as humeral width, with sides subparallel in basal two-thirds, thence converging more strongly towards apices; surface strongly and closely punctate on basal three-fourths, inconspicuously on apical fourth. Legs long, femora slightly clavate, somewhat constricted apically.
</paragraph>
<caption id="9F993FB7384B92A1B67554AB003097B6" ID-DOI="10.3897/zookeys.1226.139148.figure2" ID-arpha="E918976E-89C0-529D-B7CA-5450586FFF15" httpUri="https://binary.pensoft.net/fig/1243455" startId="F2">
<paragraph id="9E7EF8C2F0E6952191DC52713F9444D0">
<label id="C3D27446B0EAC4FB19D0DE825F03807B">Figure 2.</label>
</paragraph>
<paragraph id="B7AF90DA90848608BD1558F53B3C2476">
Holotype of
<taxonomicName id="095FA15AF5D2248507BD096F40839393" authority="Gahan, 1906 a" authorityName="Gahan" authorityYear="1906" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="licheneus">
<emphasis id="DFAB6BFF3A40E479229FC8C7148DB5D9" italics="true">Apiocephalus licheneus</emphasis>
<bibRefCitation id="EAAC26BBD44638370DB4935BF066A385" author="Gahan CJ" refId="B7" refString="Gahan CJ (1906) The Fauna of British India, including Ceylon and Burma. Coleoptera. Vol. I (Cerambycidae). Bingham CT (Ed.) London, 1 - 329." year="1906">Gahan, 1906</bibRefCitation>
<emphasis id="FDD622C1E6E764D8347984152D92E44D" bold="true">a</emphasis>
</taxonomicName>
dorsal view
<emphasis id="9F66D4CE64FABC9702CE6F9DC1DB9FB8" bold="true">b</emphasis>
lateral view
<emphasis id="FCC4C1CFE148C8D1892D73906BAE8C94" bold="true">c</emphasis>
ventral view
<emphasis id="4740ED5B4EA7C9223EE3E6B663FCCA9D" bold="true">d</emphasis>
frontal view
<emphasis id="410200A6F5EAD63CA2101CDB49371914" bold="true">e</emphasis>
lateral view of pronotum
<emphasis id="5C96D38FC6B4E13926E5299F887CCD1D" bold="true">f</emphasis>
labels.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EZPAC" type="type material">
<paragraph id="30186220A1C6A4454071F9BDADA9F905">
<heading id="AC4A1558F323D7DFAE3102125B5BB72D" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="2828C5E5F4B7BF7DE7C9C68207FA878F">
<materialsCitation id="92D8150FC833AC2D243F96A2EE8C2299" collectionCode="NHMUK, DEHRA" location="Type Gahan" municipality="Type" specimenCode="NHMUK 014596631" specimenCount="1" stateProvince="Down" typeStatus="holotype">
<emphasis id="58253CD0E4598CD8880791D2A3B396C7" bold="true" italics="true">
<typeStatus id="7790A41983EDBED10968BB63604ACD2D" type="holotype">Holotype</typeStatus>
</emphasis>
• male (
<named-content id="AE8ECAA7732F80B193E44717F3393F10" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London, UK">
<collectionCode id="F49A3E7D88ED9050A693629A925BB80F" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
):
<collectingMunicipality id="8857A6F657EAC3171A82A8D8471588A4">Type</collectingMunicipality>
[p, label circular, red framed];
<collectionCode id="B3096058FC30952A3B3CEF03AE608DB0">DEHRA</collectionCode>
<collectingRegion id="C8D0054441F67D4FD30D2548E3F166FA" country="United Kingdom" name="Down">DUN</collectingRegion>
[p]; 1900 / 14 [p]; apiocephalus /
<taxonomicName id="CB33435129C1AC355C1DD1DFD64D76D2" authorityName="Gahan" authorityYear="1906" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="licheneus">
<emphasis id="8C94A30185BFD6960DDC87C55B53E572" italics="true">licheneus</emphasis>
</taxonomicName>
, /
<location id="17F327066189A3E87DF9207CA1052E94" LSID="urn:lsid:plazi:treatment:0A02FBDE6A1651578A46831087BD7ADE:17F327066189A3E87DF9207CA1052E94" municipality="Type" name="Type Gahan" stateProvince="Down">Type Gahan</location>
[h];
<specimenCode id="222D0C7A584A8CD4C94D91B8D7AABD42" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK 014596631</specimenCode>
[p] .
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETAAE" type="distribution">
<paragraph id="9A0EDFCA2A3982B227CF3B950644FE92">
<heading id="52A39B393FA002876AA4B2540421E35B" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="8B24DF01EF7919FAC705F12A1328C2D2">
<collectingCountry id="680FC965AE9B713E89F945C9621FF4E9" name="India">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYAAE" type="remarks">
<paragraph id="ED0E045B69F8612AE2FD303E725227FC">
<heading id="45B56F3EBFC7991FB2CAC20F1CBE3602" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="D6EDC499B22104437F65F2CFEC548CCC">
This species differs from
<taxonomicName id="68AD8B0DEE97739D9E218AA64193557A" authorityName="Gahan" authorityYear="1898" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="D01ED3BEA764B1C6B00067F4E7460159" italics="true">A. punctipennis</emphasis>
</taxonomicName>
primarily by the denser greyish-white pubescence on the pronotum, the less steep anterior slope of the two anterior protuberances on the disc of the pronotum, and the presence of pubescent black spots on the elytra instead of glabrous black spots.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,530 @@
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<subSubSection id="BAFEF0DAA4834C9913F21DA751A37CE3" type="nomenclature">
<paragraph id="70F9C12C83BE3E27B040FA096B121D90">
<taxonomicName id="350F65FFAD3177D5BBB998B184A9BA56" authority="(Chen et al., 1976)" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" baseAuthorityName="Feng &amp; Yang &amp; Ge" baseAuthorityYear="1976" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="quadricavata" status="comb. nov.">
<emphasis id="7473A6CF31DCCF8AB6AAA8E9B7BB21BC" italics="true">Paleosepharia quadricavata</emphasis>
(
<bibRefCitation id="4AEF6F0797BC36E8B3979A2FA08CC14C" author="Chen SH &amp; Yu PY &amp; Wang SY &amp; Jiang SQ" journalOrPublisher="Acta Entomologica Sinica" pagination="205-224" refId="B9" refString="Chen SH, Yu PY, Wang SY, Jiang SQ (1976) New leaf beetles from west China. Acta Entomologica Sinica 19: 205-224." title="New leaf beetles from west China." volume="19" year="1976">Chen et al., 1976</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="23058BBDD4E3C9C1A8B8E3E4A928D184" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9F5792B16E708DFFC1D3EBFB74E9C8FD" type="description">
<paragraph id="3F0F799DD06DE2F4B93A153A74202E66">
<figureCitation id="6DD18770B806ABF5F5EB0163D6392514" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Paleosepharia quadricavata (Chen et al., 1976), comb. nov. (holotype, male) A C habitus. A dorsal view B ventral view C lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1226.140239.figure8" httpUri="https://binary.pensoft.net/fig/1243039">Figs 8</figureCitation>
,
<figureCitation id="245687B6E293AB05FDA95000E490DFA5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paleosepharia quadricavata (Chen et al., 1976), comb. nov. (male) A C habitus D F aedeagus G depression of the elytra of the male. A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1 mm (A C); 0.5 mm (D F)." figureDoi="10.3897/zookeys.1226.140239.figure9" httpUri="https://binary.pensoft.net/fig/1243040">9</figureCitation>
,
<figureCitation id="B0AA876336B79A2635761F5DBCCAB670" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Paleosepharia quadricavata (Chen et al., 1976), comb. nov. (female) A C habitus D vaginal palps E spermatheca F apex of abdomen of the female. A dorsal view B ventral view C lateral view. Scale bars: 1 mm (A C); 0.5 mm (D, E)." figureDoi="10.3897/zookeys.1226.140239.figure10" httpUri="https://binary.pensoft.net/fig/1243041">10</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="69F5A567DC3730729A23C7937268E289">
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<treatmentCitation id="39B5B1C6D00CD1FDA72363E09E507C77" author="Chen SH &amp; Yu PY &amp; Wang SY &amp; Jiang SQ" page="205" year="1976">
<taxonomicName id="16B5268DDD2AD560581C0708A1723189" authority="Chen, 1976: 205" authorityName="Chen" authorityPageNumber="205" authorityYear="1976" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="quadricavata">
<emphasis id="407D0D004947AE2035FDA6861D705D96" italics="true">Monolepta quadricavata</emphasis>
<comment id="2B590B455C75EF2FE3AFD757DCD909D5">
<bibRefCitation id="0F0F26C4FEC51EDB4248470B8D5BE997" author="Chen SH &amp; Yu PY &amp; Wang SY &amp; Jiang SQ" journalOrPublisher="Acta Entomologica Sinica" pagination="205-224" refId="B9" refString="Chen SH, Yu PY, Wang SY, Jiang SQ (1976) New leaf beetles from west China. Acta Entomologica Sinica 19: 205-224." title="New leaf beetles from west China." volume="19" year="1976">Chen, 1976: 205</bibRefCitation>
.
<abbrev id="ABBRID0ECYAE" xlink_title="type locality">TL</abbrev>
:
<collectingCountry id="CA1D4C7E827A6AEE5F042314C7C0AD2F" name="China">China</collectingCountry>
,
<collectingRegion id="B52418097B8513CE66B5CBF60D5E2616" country="China" name="Xizang">Xizang</collectingRegion>
;
<abbrev id="ABBRID0EGYAE" xlink_title="type deposition">TD</abbrev>
:
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<paragraph id="34ED2596832F3290CADFC0A6B38641E1">
<heading id="52DCE1030EFF7B68DC96904523B95AC0" reason="title">Type material.</heading>
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<paragraph id="3FDBD53A0CEC49837D83C123996B694F">
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<emphasis id="C2E3FC6D4A5698A1954A369F20CAA59F" bold="true" italics="true">
<typeStatus id="ECE8B1A917F089621F91D7A34FEBF94C" type="holotype">Holotype</typeStatus>
</emphasis>
: •
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<collectingCountry id="53659C0A60E876F64673EEAB33C70A03" name="China">China</collectingCountry>
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<location id="D1336E7BAF88E8D2B3EBC75C57023F13" LSID="urn:lsid:plazi:treatment:0D570786B52851CDB3289AEFAF984C0B:D1336E7BAF88E8D2B3EBC75C57023F13" country="China" municipality="Chayu" name="Shama" stateProvince="Xizang">Shama</location>
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<caption id="AC82AF4176E883AE2A9ECA84F65147EA" ID-DOI="10.3897/zookeys.1226.140239.figure8" ID-arpha="4131C5A1-ABE8-5535-9B2E-3DD1CBE1F3F0" httpUri="https://binary.pensoft.net/fig/1243039" startId="F8">
<paragraph id="A76369A0FB4F2D20A78AFD9424ABC09D">
<label id="4C6B789BE8B48814DDC94B665EFC52A5">Figure 8.</label>
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<paragraph id="0F7CFFF50B14DC2915D269E99CB49365">
<taxonomicName id="DFC73A807D4638A1AFC4A7A2CF37A2E4" authority="(Chen et al., 1976)" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" baseAuthorityName="Feng &amp; Yang &amp; Ge" baseAuthorityYear="1976" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="quadricavata" status="comb. nov.">
<emphasis id="AA037D43F09524FE782DA0489FF75D1A" italics="true">Paleosepharia quadricavata</emphasis>
(
<bibRefCitation id="49DA645C5E4C1828DD6AE14AE17FABA3" author="Chen" etAl="et al." firstAuthor="Chen" journalOrPublisher="Acta Entomologica Sinica" pagination="205-224" refId="B9" refString="Chen SH, Yu PY, Wang SY, Jiang SQ (1976) New leaf beetles from west China. Acta Entomologica Sinica 19: 205224." title="New leaf beetles from west China." volume="19" year="1976">Chen et al., 1976</bibRefCitation>
)
</taxonomicName>
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(holotype, male)
<emphasis id="24A10F1EB212096AC23532A2093A6CA1" bold="true">A C</emphasis>
habitus.
<emphasis id="58BCDF112E8FC74ABA0E296CCA22C02B" bold="true">A</emphasis>
dorsal view
<emphasis id="BB3F0882335788CEE74C25461E6C6A43" bold="true">B</emphasis>
ventral view
<emphasis id="174C87574EAAD8A8F007CC2D6D939659" bold="true">C</emphasis>
lateral view. Scale bar: 1 mm.
</paragraph>
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<paragraph id="B9E675261B1D66EEA4A9C01129D31F86">
<heading id="7E80CCEF5AE1AE8300EF3ACA8F3E82A3" reason="title">Other specimens examined.</heading>
</paragraph>
<paragraph id="386E2AE30291D987526CDE89CF3609E6">
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<quantity id="AAABA4A17759DB1784100806C13470CA" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
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<collectingDate id="520AAF75443FDD0C374F5A38F7A8A50A" value="2024-08-11">11 Aug. 2024</collectingDate>
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<collectorName id="AC7838F09599DBEF37B43BD0655241A9">Chuan Feng</collectorName>
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<collectionCode id="6F9274A3FCA6957D560D6861C3CFF033" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
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<materialsCitation id="A263BA9305707F2153554AE3BC40E5AA" collectingDate="2005-08-26" collectionCode="IZCAS" collectorName="Hongbin Liang" country="China" county="Fugong County" elevation="1505" location="Majimi" municipality="Maji" specimenCount="2" specimenCount-female="2" stateProvince="Yunan Province">
<specimenCount id="2EA106F8E714DD6ABAC71E268D775524" count="2" type="female">2 ♀♀</specimenCount>
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<location id="04C7133850758108176CBC34D5C9F8CC" LSID="urn:lsid:plazi:treatment:0D570786B52851CDB3289AEFAF984C0B:04C7133850758108176CBC34D5C9F8CC" country="China" county="Fugong County" municipality="Maji" name="Majimi" stateProvince="Yunan Province">Majimi</location>
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<quantity id="FAA403039F57D08D8A6154B27906D082" metricMagnitude="3" metricUnit="m" metricValue="1.505" unit="m" value="1505.0">1505 m</quantity>
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<collectionCode id="86BEE56783E449509799CE430AAD4B2C" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
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<materialsCitation id="3E1269A4EE02ACB0CB448780E61F1DD9" collectingDate="2024-07-24" collectionCode="IZCAS" collectorName="Chuan Feng" country="China" county="Motuo County" elevation="1410" location="Badeng" municipality="Beibeng" specimenCount="1" specimenCount-male="1" stateProvince="Xizang">
<specimenCount id="89CAB9BD484BD4ADD5F217E9984E2235" count="1" type="male">1 ♂</specimenCount>
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<collectingCounty id="9DE74C16297617D7F7CE65CCE492A044">Motuo County</collectingCounty>
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<location id="7D4C12B0166330191C4178D2B2DFC7D8" LSID="urn:lsid:plazi:treatment:0D570786B52851CDB3289AEFAF984C0B:7D4C12B0166330191C4178D2B2DFC7D8" country="China" county="Motuo County" municipality="Beibeng" name="Badeng" stateProvince="Xizang">Badeng</location>
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<collectingDate id="EA7A828CAC6DF465D121D99B1B4328F8" value="2024-07-24">24 Jul. 2024</collectingDate>
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<materialsCitation id="F310514411F8AF990F07818A14427CA6" collectingDate="2024-08-08" collectionCode="IZCAS" collectorName="Chuan Feng" country="China" county="Bomi County" elevation="2050" location="Yigong" specimenCount="2" specimenCount-male="2" stateProvince="Xizang">
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<quantity id="6E006A103E37E021E0E9D1F0D48E66AE" metricMagnitude="3" metricUnit="m" metricValue="2.05" unit="m" value="2050.0">2050 m</quantity>
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<collectingDate id="E663EFB52C021C9B16FE9969B99F9E86" value="2024-08-08">8 Aug. 2024</collectingDate>
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</named-content>
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<materialsCitation id="12F49496D1DA4B90DA1AF396363CFF73" collectingDate="2015-08-19" collectionCode="IZCAS" collectorName="Jian Yao" country="China" county="Motuo County" elevation="799" location="Beibeng" specimenCount="4" specimenCount-male="4" stateProvince="Xizang">
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<quantity id="D57655A42BB5B588125ADBC047F87A2C" metricMagnitude="2" metricUnit="m" metricValue="7.99" unit="m" value="799.0">799 m</quantity>
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<collectingDate id="B363916DC55DD4363D5EA897D4916057" value="2015-08-19">19 Aug. 2015</collectingDate>
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<collectorName id="3440BFD4945179D6BBA2338CA01DC2BD">Jian Yao</collectorName>
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; •
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,
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,
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<quantity id="00BBA127CA553868CFF29B0CD16A309F" metricMagnitude="2" metricUnit="m" metricValue="7.99" unit="m" value="799.0">799 m</quantity>
a. s. l.
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,
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<collectingDate id="DFD9F92AB555E97EFF2C902AEEC5E246" value="2015-08-19">19 Aug. 2015</collectingDate>
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,
<collectorName id="77B2E71F1C4F19DC33F81871363B9D5B">Hongbin Liang</collectorName>
&amp;
<collectorName id="920F41D6817619B948486B0A9B13D335">Zhengzhong Huang</collectorName>
leg.
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; •
<materialsCitation id="491F968CED47D1C6BA8050FE40A8C607" collectingDate="2015-08-17" collectionCode="IZCAS" collectorName="Jian Yao" country="China" county="Motuo County" elevation="707" location="Ximohe" specimenCount="5" specimenCount-male="5" stateProvince="Xizang">
<specimenCount id="AF3CE575847FA916C748755400D4183E" count="5" type="male">5 ♂♂</specimenCount>
,
<collectingCountry id="B9A787BE439C26211CBC201FDD318AFD" name="China">China</collectingCountry>
,
<collectingRegion id="3ECA65E47243E739A4257CCF3B93C7B6" country="China" name="Xizang">Xizang Province</collectingRegion>
,
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,
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<quantity id="3D3E15AE50078F1941C2FACA44CF6795" metricMagnitude="2" metricUnit="m" metricValue="7.07" unit="m" value="707.0">707 m</quantity>
a. s. l.
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,
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<collectingDate id="3931782B2E8A5CC8C59B195B5B22F303" value="2015-08-17">17 Aug. 2015</collectingDate>
</date>
,
<collectorName id="C596442B32E2EB30ECC2CF1ABEF40083">Jian Yao</collectorName>
leg.
<named-content id="B64AAADCE289FF29B615274820AEE2B4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
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</named-content>
</materialsCitation>
; •
<materialsCitation id="A5443A39051B5AF7A4416755FF924842" collectingDate="1982-08-27" collectionCode="IZCAS" collectorName="Yinheng Han" country="China" county="Motuo County" elevation="900" location="Motuo County" specimenCount="1" specimenCount-female="1" stateProvince="Xizang">
<specimenCount id="ACE9ED38025944FF540805748F1FA9A9" count="1" type="female">1 ♀</specimenCount>
,
<collectingCountry id="B2C5ABFE6EE5C7A2AEFAA8A8C0221309" name="China">China</collectingCountry>
,
<collectingRegion id="96281500360CED4216FFD9B112C0C566" country="China" name="Xizang">Xizang Province</collectingRegion>
,
<collectingCounty id="BC502192D0A5AAB5E48C8B87F874CBF0">Motuo County</collectingCounty>
,
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<quantity id="132156B8F2022BBF441A615E22D35A23" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</quantity>
a. s. l.
</elevation>
,
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<collectingDate id="B2BEE0AF7C21A8F1C27BAEAC372D7084" value="1982-08-27">27 Aug. 1982</collectingDate>
</date>
,
<collectorName id="2605260AC84B8FC6E88E25EB087BE88A">Yinheng Han</collectorName>
leg.
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<collectionCode id="9D82C15786FD5B640D0A66D6A5282B1D" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<caption id="DA77BA374EA413DDAB067309B01F24E4" ID-DOI="10.3897/zookeys.1226.140239.figure9" ID-arpha="EE0BBF1F-318A-571A-8774-68234DFF1147" httpUri="https://binary.pensoft.net/fig/1243040" startId="F9">
<paragraph id="1DB3AB66E4BBEBAF70EB24BEFB0660A9">
<label id="A4BF860290971ECE7C78A511FBC9D46C">Figure 9.</label>
</paragraph>
<paragraph id="F020AC8CBBF1172502445ED0B130CA8C">
<taxonomicName id="0A0E2469DD657C2A576EA3A22BEDB8CC" authority="(Chen et al., 1976)" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" baseAuthorityName="Feng &amp; Yang &amp; Ge" baseAuthorityYear="1976" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="quadricavata" status="comb. nov.">
<emphasis id="C2B4E46BEC5BC38242CDA064444543A3" italics="true">Paleosepharia quadricavata</emphasis>
(
<bibRefCitation id="F986145C799566ECEFF313C46ABC6FC1" author="Chen" etAl="et al." firstAuthor="Chen" journalOrPublisher="Acta Entomologica Sinica" pagination="205-224" refId="B9" refString="Chen SH, Yu PY, Wang SY, Jiang SQ (1976) New leaf beetles from west China. Acta Entomologica Sinica 19: 205224." title="New leaf beetles from west China." volume="19" year="1976">Chen et al., 1976</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="2C83C386212700571E4483C645A9D0F2" rank="species">comb. nov.</taxonomicNameLabel>
(male)
<emphasis id="727206EE980C73336BF4EC2EF5094C9E" bold="true">A C</emphasis>
habitus
<emphasis id="5016FBAEC4568E70E57BF632B036F0AC" bold="true">D F</emphasis>
aedeagus
<emphasis id="A9C7617EDE3B538F1D23CFA676470F09" bold="true">G</emphasis>
depression of the elytra of the male.
<emphasis id="0A8EE54B008519AE8B8382D90664EBD0" bold="true">A, D</emphasis>
dorsal view
<emphasis id="5611ABF5995C2F52C4FBB99AB77D82B1" bold="true">B, E</emphasis>
ventral view
<emphasis id="AF2FD16D6302CD0C1D1AB39158E83921" bold="true">C, F</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="B0A8B8CD5BC5AD3CC8122433442D662F" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="2FE1EFD32CE82FE8275F951E941B32C7" bold="true">D F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ET3AE" type="notes">
<paragraph id="18C9E6381BE3E4BED92C219C26AEFFF5">
<heading id="6FD4D60D8E2C689C20790969820EFF10" reason="title">Notes.</heading>
</paragraph>
<paragraph id="2DECAB1CAC28558438A0A8122E40D8D5">
The species is transferred from
<taxonomicName id="570BC2FE60A7C6D1706BEC6BD047B63B" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="05A0705765A609C1A474B398F310AB1E" italics="true">Monolepta</emphasis>
</taxonomicName>
to
<taxonomicName id="48151777B527CB251F21D939C385168B" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="B911F8E3DEEBC3A88426A813C1B554E9" italics="true">Paleosepharia</emphasis>
</taxonomicName>
based on the following characters: the third antennomere longer than the second; depression in the elytra; aedeagus parallel-sided in the basal two-thirds, apical third strongly narrowed; tectum long, rounded at apex.
</paragraph>
<caption id="0085AB0245F4A63EC171119B40E92E57" ID-DOI="10.3897/zookeys.1226.140239.figure10" ID-arpha="2DB6E103-044B-5D9E-80D3-88F4B47C14FD" httpUri="https://binary.pensoft.net/fig/1243041" startId="F10">
<paragraph id="CFC249C4BFC25ABBA3428A2E18491DF1">
<label id="39E4CB022427407D3C5A197D66A5A174">Figure 10.</label>
</paragraph>
<paragraph id="92882353B9D8BE45D432F95B6BDDC660">
<taxonomicName id="622E103134A6E9A8E6277EE7B694D214" authority="(Chen et al., 1976)" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" baseAuthorityName="Feng &amp; Yang &amp; Ge" baseAuthorityYear="1976" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="quadricavata" status="comb. nov.">
<emphasis id="1CC46683305200C59B491DFF4493FCB9" italics="true">Paleosepharia quadricavata</emphasis>
(
<bibRefCitation id="A02C0FBD6AE5DAB4A3D6A854E8A32CD6" author="Chen" etAl="et al." firstAuthor="Chen" journalOrPublisher="Acta Entomologica Sinica" pagination="205-224" refId="B9" refString="Chen SH, Yu PY, Wang SY, Jiang SQ (1976) New leaf beetles from west China. Acta Entomologica Sinica 19: 205224." title="New leaf beetles from west China." volume="19" year="1976">Chen et al., 1976</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="CDAC7DF227978BFAF0B1DBA026F733AB" rank="species">comb. nov.</taxonomicNameLabel>
(female)
<emphasis id="53DEB53085B4BC9414B80D48DD7952F8" bold="true">A C</emphasis>
habitus
<emphasis id="EB1698A4E7AF4505FF3A671C9C2F8C2B" bold="true">D</emphasis>
vaginal palps
<emphasis id="B6FF580D3E198ABBB6B3DA9F99561016" bold="true">E</emphasis>
spermatheca
<emphasis id="AEA964C4A4E860B8071C6F96ECA33E5B" bold="true">F</emphasis>
apex of abdomen of the female.
<emphasis id="775328C87B820F33091385312991FED1" bold="true">A</emphasis>
dorsal view
<emphasis id="2634273542E1AE36A8582C2D8BEC97DC" bold="true">B</emphasis>
ventral view
<emphasis id="FA08D79EDE6E9D6389557C52785C2D73" bold="true">C</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="1A17B516CCE8CC6BAA9C0E3B25CAA116" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="E2E9FEC0E450B099D6DA774E06D47132" bold="true">D, E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EV5AE" type="diagnosis">
<paragraph id="783E53D77FEEC5044DCF656528B62C91">
<heading id="99DC4956426E7DB63260ACF36AAD6797" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="F1F3070E803EC0206F1755D2836C6530">This species is easy to distinguish from other species based on the elytra red or reddish-brown, with a large, deep concavity near suture in male.</paragraph>
</subSubSection>
<subSubSection id="SECID0E15AE" type="redescription">
<paragraph id="B11DB26B431970616D0DFF4099DEF068">
<heading id="EA4989D476CCA265A984710075498242" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="945264A6A384F029BFD2B4DD6632B628">
<emphasis id="DD87E2207097967A33E6E5DAC247A222" bold="true">Male</emphasis>
: Length: 4.25.0 mm, width: 2.0
<quantity id="A1E55C350E36C04E620EBF06EC1F59B4" metricMagnitude="-3" metricUnit="m" metricValue="2.4" unit="mm" value="2.4">2.4 mm</quantity>
(mean length
<quantity id="C17A38B768E2AAF3C0975A87778D2DAE" metricMagnitude="-3" metricUnit="m" metricValue="4.6" unit="mm" value="4.6">4.6 mm</quantity>
, mean width
<quantity id="D25E90EC548C8B87212B54248512EBA1" metricMagnitude="-3" metricUnit="m" metricValue="2.1" unit="mm" value="2.1">2.1 mm</quantity>
,
<emphasis id="1A240F4DCA19D720D6B871AF1A8EB559" italics="true">N</emphasis>
= 19).
</paragraph>
<paragraph id="631ECC4C441B313A2121592535A4FCEA">General color red or reddish-brown, head black, pronotum yellow, antennae and legs brown.</paragraph>
<paragraph id="18CBD1982D044745FBF18627F5968B17">Vertex with sparse punctures. Frontal tubercles transverse, extending downward between antennal bases. Antennae slender, equal to length of body. First antennomere shiny bare, rod-shaped, second to eleventh antennomeres with short hairs, second antennomere shortest, third antennomere about 1.3 × as long as second; fourth antennomere about twice as long as third, fifth antennomere about 1.2 × as long as fourth, sixth to eleventh antennomeres equal in length to fourth. Pronotum about 1.4 × as wide as long, lateral margins straight, basal border slightly concaved in middle, apical border straight, disc with lateral fovea and dense minute punctures. Scutellum triangular, smooth, impunctate. Elytra wider than pronotum, humeri convex, with large, deep concavity near suture at basal 1 / 3, disc with dense, minute punctures. Elytral epipleuron broad at base, gradually narrowed from middle to apex. Each tibia with distinct spur at apex, segment 1 of hind tarsi longer than remaining segments combined. Aedeagus robust, close to outsole shape in dorsal and ventral views, moderately recurved at apex in lateral view. Lateral spiculae short and curved, ventral spiculae comb-shaped, and median spiculae long.</paragraph>
<paragraph id="1C95058ED841FF38CBEE6EA82081EC9C">
<emphasis id="C0B093ACE7743FF61B24371995B0C6E2" bold="true">Female</emphasis>
: Length: 4.25.0 mm, width:
<quantity id="F4BE5CABF346FA30EE10D38A1078DCA0" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" unit="mm" value="2.35" valueMax="2.6" valueMin="2.1">2.12.6 mm</quantity>
(mean length
<quantity id="8EC5F70F6564D5263A57C20605639F4D" metricMagnitude="-3" metricUnit="m" metricValue="4.7" unit="mm" value="4.7">4.7 mm</quantity>
, mean width
<quantity id="C38FD539AE57DC58BAF3D3CFD851BA44" metricMagnitude="-3" metricUnit="m" metricValue="2.3" unit="mm" value="2.3">2.3 mm</quantity>
,
<emphasis id="F60B81EA4EFD274D91CE9D9232EF2355" bold="true" italics="true">N</emphasis>
= 3).
</paragraph>
<paragraph id="D5D54F84BED4B69440E495E70C9CC8D5">Third antennomere about 1.6 × as long as second; fourth antennomere about 1.6 × as long as third. Elytra only with one slight depression at basal 1 / 3. Spermatheca with big and rounded nodulus, middle part short, cornu short and slightly curved. Vaginal palps with wide base and rounded apex, each palp slightly narrowing posteriorly, with 5 setae placed at apex, additional 2 setae subapically.</paragraph>
</subSubSection>
<subSubSection id="SECID0EO6AE" type="distribution">
<paragraph id="B74029D379F45CC25C213914195062D3">
<heading id="E921386D0FAAD19501D17E74DC0A6772" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="C995F53504CF9AFAFC0472C1EF91D3CA">
<collectingCountry id="1E268F695543807DE96124A17740B8CF" name="China">China</collectingCountry>
:
<collectingRegion id="B0D0CF8D7FA4ED1AEB6C81C3DD4B5D52" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingRegion id="B61EA88E8BFA2D004CE8DAA56E0C46AF" country="China" name="Xizang">Xizang</collectingRegion>
;
<collectingCountry id="166803F35E8D0872E9957824CE89AED1" name="Nepal">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="0D7113211E9A7729C31A88B4EE8EAFCB">
<title id="D10FF5EFC6F4FD57D029694DAB0B96CB">New record</title>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,175 @@
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<mods:title id="E8B099BEFE11F82C9ED9AC0A15493A80">Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province</mods:title>
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<mods:affiliation id="6864D5D39E06ADAB12A188B533E5E755">Department of Biology, San Diego State University, San Diego, California 92182 4614, USA &amp; Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, California 90007, USA</mods:affiliation>
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<subSubSection id="AFA16B587C31F17F9EA9022750B53054" type="nomenclature">
<paragraph id="C7370542A9ACE8119A44A4F8A7D806D1">
<taxonomicName id="B8D6F134355BAA90CC403C031BEB738A" ID-CoL="5BZFP" authority="Roth, 1981" authorityName="Roth" authorityYear="1981" class="Arachnida" family="Cybaeidae" genus="Willisus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="gertschi">
<emphasis id="F0DD5A7F0634A745F2EDD047A328BCDC" italics="true">Willisus gertschi</emphasis>
<bibRefCitation id="5B431F79AD708A7B35301BFC658E4030" author="Roth VD" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="101-105" refId="B54" refString="Roth VD (1981) A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy. Bulletin of the American Museum of Natural History 170: 101-105. https://wsc.nmbe.ch/species/12952" title="A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy." url="https://wsc.nmbe.ch/species/12952" volume="170" year="1981">Roth, 1981</bibRefCitation>
</taxonomicName>
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<paragraph id="81258B0C1EE04BC4921328D2742F14A1">
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<treatmentCitation id="667F5A31F2CF675DF03A05739F512BA4" author="Roth VD" page="103" year="1981">
<taxonomicName id="64D285A51E07CA85588074F4ABA6B0BF" ID-CoL="5BZFP" authority="Roth, 1981: 103" authorityName="Roth" authorityPageNumber="103" authorityYear="1981" class="Arachnida" family="Cybaeidae" genus="Willisus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="gertschi">
<emphasis id="B9A84A0A1320CC220FC1664918CB24D5" italics="true">Willisus gertschi</emphasis>
<comment id="FE5E70616355F7BDF84D8EFCDAADCE19">
<bibRefCitation id="69B8A0DF215BA97EE45DBE91DB3480C5" author="Roth VD" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="101-105" refId="B54" refString="Roth VD (1981) A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy. Bulletin of the American Museum of Natural History 170: 101-105. https://wsc.nmbe.ch/species/12952" title="A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy." url="https://wsc.nmbe.ch/species/12952" volume="170" year="1981">Roth, 1981: 103</bibRefCitation>
, figs 18 (
<specimenCount id="0A7F6002C5BBD88ACBA049D3CFFD68E3" count="1" type="male"></specimenCount>
<specimenCount id="90EFD81C4A2F00D1B59FFD8517B7A24D" count="1" type="female"></specimenCount>
).
</comment>
</taxonomicName>
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</paragraph>
<paragraph id="EE319CE604D74141C4C86BE509660F4F">
<treatmentCitationGroup id="74852AAFE196BE327A9A247B8EF66E4C">
<treatmentCitation id="7BD918B4A6677D2F5B6AD6475884D86F">
<taxonomicName id="1A6557ADD331874527261C84911969CD" ID-CoL="5BZFP" authorityName="Roth" authorityYear="1981" class="Arachnida" family="Cybaeidae" genus="Willisus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="gertschi">
<emphasis id="250E4E7294703253A62A43DCFB640071" italics="true">Willisus gertschi</emphasis>
</taxonomicName>
: Roth 1982: 78, figs 1, 7 (
<specimenCount id="B4021DB4802B69217055D221E9BB3743" count="1" type="male"></specimenCount>
<specimenCount id="CA727F1057D498E6C12752439C6E9F96" count="1" type="female"></specimenCount>
); Roth 1985: B 16, figs 1, 7 (
<specimenCount id="478D24E887EDCFEB522A6AC88A6B4FBF" count="1" type="male"></specimenCount>
<specimenCount id="DACA1AD0E1E654AC4E3B22DD3699D347" count="1" type="female"></specimenCount>
);
<bibRefCitation id="2B9958BCEB86BC26C3474643F0E440F3" author="Roth VD" firstAuthor="Roth" journalOrPublisher="American Arachnological Society, Gainesville" page="53" refId="B55" refString="Roth VD (1994) Spider Genera of North America, with Keys to Families and Genera, and a Guide to Literature, third edition. American Arachnological Society, Gainesville, 203 pp." title="Spider Genera of North America, with Keys to Families and Genera, and a Guide to Literature, third edition." year="1994">Roth 1994: 53</bibRefCitation>
</treatmentCitation>
, figs 1, 7 (
<specimenCount id="09A50CF3B47EB1DDF7F228B7C2905D1F" count="1" type="male"></specimenCount>
<specimenCount id="8F42E6E51924C9D6CAB75B594341EB81" count="1" type="female"></specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVSAI" type="LSID">
<paragraph id="B16D5F97711A8909FD583C102D7AA272">
<heading id="4E1FE14DC5CCEBF0130B326C59ED28A2" reason="title">
<collectionCode id="7883EBDD3A59AF02E1BA6614F22E7C8F">LSID</collectionCode>
.
</heading>
</paragraph>
<paragraph id="81A848F7C15FCE585C756F457179EDDB">[urn:lsid:nmbe.ch:spidersp:022044].</paragraph>
</subSubSection>
<subSubSection id="SECID0E1SAI" type="material">
<paragraph id="4699427B8F75FC84B5579623474A5E17">
<heading id="908960346D10D6C7C80B76FA32C8EF2F" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="7B2419B80E917127BF6430278EC04686">
<emphasis id="6AA5C30159150CDE1A6177ECA000F8B6" bold="true">
<collectingRegion id="41D01DE48680C7C6D54E57D6194E6A70" country="United States of America" name="California">California</collectingRegion>
, San Bernardino Co.
</emphasis>
<materialsCitation id="F3D74693C4B83D62F42824E82EF19ACB" collectingDate="2023-09-09" collectionCode="SDSU" collectorName="M. Hedin" location="East Fork Mountain Home Creek" specimenCode="G 4110" specimenCount="1" specimenCount-male="1">
<specimenCount id="69B32A137A515F482251808C50D62F20" count="1" type="male"></specimenCount>
; San Bernardino Mountains, Hwy 38, crossing of tributary of
<location id="2E2BECE8315885584CFF4EFE14404F8E" LSID="urn:lsid:plazi:treatment:0FBC7844D0EC561E9A2DDE1778ED9CF3:2E2BECE8315885584CFF4EFE14404F8E" name="East Fork Mountain Home Creek">East Fork Mountain Home Creek</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFTAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-116.980000,34.120000]}">34.12, - 116.98</named-content>
;
<date id="B3CAD6AD0AE70BF0F08119D46174E602" value="2023-09-09">
<collectingDate id="3644DD2778F98B2717FC73FC75B9798C" value="2023-09-09">9 September 2023</collectingDate>
</date>
; coll.
<collectorName id="81A377165356D93004022785537F39AD">M. Hedin</collectorName>
; MCH 23 _ 048;
<collectionCode id="F957E7E6876BDE2E1E739AAA71495E6D" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="E7FF135D36BA8EC589437A5D1C310216" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">G 4110</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKTAI" type="variation">
<paragraph id="4D7242D8136B9EB848AE64156F3E5299">
<heading id="D2A7D4FAFBC370E81216744A3D4D14B8" reason="title">Variation.</heading>
</paragraph>
<paragraph id="D417D4607876FA1F5F047D7DE2CAB9CB">
We have not seen type specimens of
<taxonomicName id="5D5103ACA28CC9D74F7B6CE3B32E75A4" authorityName="Roth" authorityYear="1981" class="Arachnida" family="Cybaeidae" genus="Willisus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="gertschi">
<emphasis id="C8FEDD49212D90D15363918C59046A84" italics="true">W. gertschi</emphasis>
</taxonomicName>
, but descriptions and illustrations of the
<typeStatus id="1CC691D1AE63CAB095C7C43D662269AD" type="holotype">holotype</typeStatus>
male (
<bibRefCitation id="BB71BAFD203EC9D758A9CCF2D7D9BE59" author="Roth" firstAuthor="Roth" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="101-105" refId="B54" refString="Roth VD (1981) A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy. Bulletin of the American Museum of Natural History 170: 101105. https://wsc.nmbe.ch/species/12952" title="A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy." url="https://wsc.nmbe.ch/species/12952" volume="170" year="1981">Roth 1981</bibRefCitation>
) mostly match characters seen in the San Bernardino Mtns male specimen, including the globular conductor and robustness of the dorsal tegular process. The San Bernardino Mtns male embolus appears less sinuate than
<typeStatus id="9DD5F49FCFE93DC725D886C7181AE81E" type="holotype">holotype</typeStatus>
illustrations (
<bibRefCitation id="85C3517D703966BFD02E1C8972C81D4F" author="Roth" firstAuthor="Roth" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="101-105" refId="B54" refString="Roth VD (1981) A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy. Bulletin of the American Museum of Natural History 170: 101105. https://wsc.nmbe.ch/species/12952" title="A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy." url="https://wsc.nmbe.ch/species/12952" volume="170" year="1981">Roth 1981</bibRefCitation>
: figs 4, 5).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDUAI" type="comments">
<paragraph id="00C81FB58588AC035A5FF95F35587768">
<heading id="1F5C8DB5C5CF96B4874F2FDB8BEE7B91" reason="title">Comments.</heading>
</paragraph>
<paragraph id="8561BB8B7D4A19CFE4E3543564FCD108">
Previously known only from the
<typeStatus id="68EFFA4229314EC62B18E95B33889FFD">type</typeStatus>
locality in the San Gabriel Mountains of southern
<collectingRegion id="37EA79F1CD0431026EBF24BF24B3004D" country="United States of America" name="California">California</collectingRegion>
(Mt. Baldy, Manker Flats Campground;
<bibRefCitation id="352F0801CB28B3438BDB3668F3F77FE5" author="Roth" firstAuthor="Roth" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="101-105" refId="B54" refString="Roth VD (1981) A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy. Bulletin of the American Museum of Natural History 170: 101105. https://wsc.nmbe.ch/species/12952" title="A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy." url="https://wsc.nmbe.ch/species/12952" volume="170" year="1981">Roth 1981</bibRefCitation>
). Our new records from the San Bernardino Mountains extend the distribution of this species eastward in the Los Angeles Basin. We conservatively treat these populations as conspecific until additional collections can be made. We also urge additional collecting at high elevations in neighboring mountain ranges (e. g., San Jacinto Mountains, Santa Rosa Mountains) to perhaps uncover additional records of this rare genus.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,367 @@
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<taxonomicName id="09F18E79C879CFC66F41CEE977D6487A" authority="Feng, Yang &amp; Ge" authorityName="Feng, Yang &amp; Ge" authorityYear="2025" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="emeiensis" status="sp. nov.">
<emphasis id="CE11EEA0BEB68068C4559CDE20022A63" italics="true">Paleosepharia emeiensis</emphasis>
Feng, Yang &amp; Ge
</taxonomicName>
<taxonomicNameLabel id="6B978A9185B177BF157C505F5A08CDE8" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="AB4095B1803AC812A8DD06F9C165CF97">
<figureCitation id="F6AB83ACE290EEE76A70AD667ED14646" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paleosepharia emeiensis sp. nov. (holotype, male) A C habitus D F aedeagus G depression of the elytra of the male. A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1 mm (A C); 0.5 mm (D F)." figureDoi="10.3897/zookeys.1226.140239.figure3" httpUri="https://binary.pensoft.net/fig/1243034">Figs 3</figureCitation>
,
<figureCitation id="6180298052A28EF1D48E1D4413729C09" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Paleosepharia emeiensis sp. nov. (paratype, female) A C habitus D vaginal palps E spermatheca F apex of abdomen of the female. A dorsal view B ventral view C lateral view. Scale bars: 1 mm (A C); 0.5 mm (D, E)." figureDoi="10.3897/zookeys.1226.140239.figure4" httpUri="https://binary.pensoft.net/fig/1243035">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECHAE" type="type material">
<paragraph id="5C5A145D40CAA807B0C0D40E960C9F05">
<heading id="E625266695F566EED3EB24FDD1C6A381" reason="title">Type material.</heading>
</paragraph>
<paragraph id="641B8F6F5D11649DC2174C3035A85626">
<materialsCitation id="354911C9FAE363379F453ADEC6845A0A" collectingDate="1957-05-09" collectionCode="IZCAS" collectorName="Keren Huang" country="China" elevation="650" location="Baoguosi" municipality="Mt Emei" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="76E5624BE5D8EBA14DF613687E184136" bold="true" italics="true">
<typeStatus id="454A2239F4B3EBAD2BAC0391C353F002" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCount id="D823A4445E23C723CDD9E91038D9A85F" count="1" type="male"></specimenCount>
,
<collectingCountry id="1F51B7A2B35DB339B5C7514ACC9BD0BC" name="China">China</collectingCountry>
,
<collectingRegion id="36F938699ED2F1FCFDDDA18E9BE155CF" country="China" name="Sichuan">Sichuan Province</collectingRegion>
,
<collectingMunicipality id="E6B31B49884AFDA4AEF967301E505CCC">Mt Emei</collectingMunicipality>
,
<location id="CEA75F9EFB025479D4E454B8476D05FC" LSID="urn:lsid:plazi:treatment:1B94D1A8DB6F5996B821FCDC1B70A6FB:CEA75F9EFB025479D4E454B8476D05FC" country="China" municipality="Mt Emei" name="Baoguosi" stateProvince="Sichuan">Baoguosi</location>
,
<elevation id="74AAEE8CFE7A6E150D92DCBF192F3831" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">
<quantity id="86A5BF5C268C55E6359A7E479435EE8D" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">550750 m</quantity>
a. s. l.
</elevation>
,
<date id="6FB89B1D0E9596411FD271E83C2F57D0" value="1957-05-09">
<collectingDate id="834978DC90DCA8DD1B2FBA18ED89C371" value="1957-05-09">9 May 1957</collectingDate>
</date>
,
<collectorName id="7AECD05549F30E4F7A950ECCA6CC6600">Keren Huang</collectorName>
leg.,
<named-content id="84535FF047D94191FA2EECFC3AA0EF02" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="0C265A27713C260621D7182AC5D13D3F" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
.
<materialsCitation id="8A2FC31CA4B8104F78844E31093D260F" collectingDate="1957-05-09" collectionCode="IZCAS" collectorName="Keren Huang" country="China" elevation="650" location="Baoguosi" municipality="Mt Emei" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Sichuan" typeStatus="paratype">
<emphasis id="501F0A47B2A481DAAB76F9B1A84E20E5" bold="true" italics="true">
<typeStatus id="E8D3EF83B516E7DBF72F4EA8F08419A4" type="paratype">Paratype</typeStatus>
</emphasis>
: •
<specimenCount id="D83E772DC042F70552822129B1D7D21C" count="3" type="male">3 ♂♂</specimenCount>
<specimenCount id="757083C3954859699D7D3F3A7B8CAAE1" count="4" type="female">4 ♀♀</specimenCount>
, same data as for holotype
</materialsCitation>
; •
<materialsCitation id="517FF7891F04FACD1E4FB21339CC765F" collectingDate="1957-06-02" collectionCode="IZCAS" collectorName="Keren Huang" country="China" elevation="650" location="Baoguosi" municipality="Mt Emei" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="48CFC3FAA7548427AF99D9A5DED0E7DB" count="3" type="male">3 ♂♂</specimenCount>
<specimenCount id="8DE1FD9876E2D9DDEF7D271E6FAE0B9D" count="2" type="female">2 ♀♀</specimenCount>
,
<collectingCountry id="9F36215A48E64A09E489B12A87A85A52" name="China">China</collectingCountry>
,
<collectingRegion id="0C2AE999F85153900BE61BA7FD33DAA9" country="China" name="Sichuan">Sichuan Province</collectingRegion>
,
<collectingMunicipality id="5B4739F7C866464DF1259560BA704BB7">Mt Emei</collectingMunicipality>
,
<location id="3A0DB35B46BF14E69738E149D29B0A1A" LSID="urn:lsid:plazi:treatment:1B94D1A8DB6F5996B821FCDC1B70A6FB:3A0DB35B46BF14E69738E149D29B0A1A" country="China" municipality="Mt Emei" name="Baoguosi" stateProvince="Sichuan">Baoguosi</location>
,
<elevation id="1E5AC60577ACD39C48FB86F2A3EEAC73" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">
<quantity id="6582FAD9515914C0234F9658E0D31CD7" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">550750 m</quantity>
a. s. l.
</elevation>
,
<date id="61B77ED9FBD8F27559C542B034C66D0E" value="1957-06-02">
<collectingDate id="3743FE8BB02B34E50D18914BE1D32A5E" value="1957-06-02">2 Jun. 1957</collectingDate>
</date>
,
<collectorName id="EE81F3211CADF3ED746C8DCABB133F68">Keren Huang</collectorName>
leg.,
<named-content id="04560E678566F6957A497196FA72699B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="0D82C72D8EA3BC23A013C41CD5220945" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="FF56266B550D9C65FD3A80FF9BD4F5BB" collectingDate="1994-06-03" collectionCode="IZCAS" collectorName="Xingke Yang" country="China" county="Fengdu County" elevation="610" location="Shiping" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="4E817596A4BB07BEF6D64390BEED1381" count="3" type="male">3 ♂♂</specimenCount>
<specimenCount id="A74818E3573E5509CE864E7B5B715591" count="4" type="female">4 ♀♀</specimenCount>
,
<collectingCountry id="77F5933CA19033479E735ECDA9E6D114" name="China">China</collectingCountry>
,
<collectingRegion id="9C6BEF4605AB15C224F6285C2FEB1D48" country="China" name="Sichuan">Sichuan Province</collectingRegion>
,
<collectingCounty id="2EC5AF5CC67F398C110510D1E8FDE091">Fengdu County</collectingCounty>
,
<location id="B34C764F149447FB3CA02562157593E1" LSID="urn:lsid:plazi:treatment:1B94D1A8DB6F5996B821FCDC1B70A6FB:B34C764F149447FB3CA02562157593E1" country="China" county="Fengdu County" name="Shiping" stateProvince="Sichuan">Shiping</location>
,
<elevation id="99BE49A7E13A9B7614DA755D17B8B3BB" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">
<quantity id="CF3C2C34939E255EB3970DCC726D9B51" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">610 m</quantity>
a. s. l.
</elevation>
,
<date id="F9A3DEF5664D8959EC1A6ADB05D4A8E0" value="1994-06-03">
<collectingDate id="352B6F363ED2C1E8CE6AA65856747B49" value="1994-06-03">3 Jun. 1994</collectingDate>
</date>
,
<collectorName id="220CB63DAAA4C4E12A1760F8B6D40D33">Xingke Yang</collectorName>
leg.,
<named-content id="C20F2FC801C9456D4F777CD981C61D4A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="DFCB7A80A9328D670BB1B48553FB2A0C" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="CAD89E34ED82BDE9308DAB42FE4F669A" collectingDate="1994-06-03" collectionCode="IZCAS" collectorName="Youwei Zhang" country="China" county="Fengdu County" elevation="610" location="Shiping" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="A0651E3467B96450B3CD44B0BA2CB8B6" count="2" type="male">2 ♂♂</specimenCount>
<specimenCount id="B593F5BD344756717E370765B17E1103" count="5" type="female">5 ♀♀</specimenCount>
,
<collectingCountry id="44C6331A6852BD2A0911BDCADA1694D7" name="China">China</collectingCountry>
,
<collectingRegion id="FEE872F4A19E05F1EE889A7646D6FA85" country="China" name="Sichuan">Sichuan Province</collectingRegion>
,
<collectingCounty id="0034F4CCC1DCF66A391D4FA6ED70B389">Fengdu County</collectingCounty>
,
<location id="593E7437587F73F774E64D2FC164FBA3" LSID="urn:lsid:plazi:treatment:1B94D1A8DB6F5996B821FCDC1B70A6FB:593E7437587F73F774E64D2FC164FBA3" country="China" county="Fengdu County" name="Shiping" stateProvince="Sichuan">Shiping</location>
,
<elevation id="159363D9F44872E7A837A01C14DB7B2D" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">
<quantity id="F9FADBF0C44537E9BE5D705665F80889" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">610 m</quantity>
a. s. l.
</elevation>
,
<date id="3D7433ED357E5EC6DBB10944772D9FE8" value="1994-06-03">
<collectingDate id="2A4EA9DF720F1DF9251F0CEE51468BF6" value="1994-06-03">3 Jun. 1994</collectingDate>
</date>
,
<collectorName id="ADC490F87FE4AE148344FADA71C75E02">Youwei Zhang</collectorName>
leg.,
<named-content id="1262613AB399D9F6ED68588949C51B60" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="FDCB2F036781F71A1A51197EB5C3F067" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="8A576ACCD3DABC751FD9093B0C75C711" collectingDate="1994-06-03" collectionCode="IZCAS" collectorName="Wenzhu Li" country="China" county="Fengdu County" elevation="610" location="Shiping" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9283C48BD162DEE47F796821D838EFD3" count="2" type="male">2 ♂♂</specimenCount>
<specimenCount id="CE0F40261ABD72FE2216E3A728D2113D" count="6" type="female">6 ♀♀</specimenCount>
,
<collectingCountry id="493BF5B776EDF68C824D6E1D780EB739" name="China">China</collectingCountry>
,
<collectingRegion id="687AAB3EB2A17B6264B1744209AF3687" country="China" name="Sichuan">Sichuan Province</collectingRegion>
,
<collectingCounty id="89627711AF564A1CC51E77C5C8BB3A4F">Fengdu County</collectingCounty>
,
<location id="E2C9C8B0409FA070D18C7E89D8C86F1A" LSID="urn:lsid:plazi:treatment:1B94D1A8DB6F5996B821FCDC1B70A6FB:E2C9C8B0409FA070D18C7E89D8C86F1A" country="China" county="Fengdu County" name="Shiping" stateProvince="Sichuan">Shiping</location>
,
<elevation id="84780BC4ACE1905CC3C7CD4B7C4A812D" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">
<quantity id="E140F144553AC72FDA771ECA09641CE5" metricMagnitude="2" metricUnit="m" metricValue="6.1" unit="m" value="610.0">610 m</quantity>
a. s. l.
</elevation>
,
<date id="CDE6B5455D35BEDA6E307DF8CE3F85CF" value="1994-06-03">
<collectingDate id="5769150BE92A00448C0323CFBCC8C72F" value="1994-06-03">3 Jun. 1994</collectingDate>
</date>
,
<collectorName id="4402172C6B2FD71266D84FC4A288ABCB">Wenzhu Li</collectorName>
leg.,
<named-content id="B930E4668980D947CC3A52DF87CBEC37" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="545E9C55CE3E07B869AC168AE6439D22" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<caption id="2C7B9249E73220989818037854C1AF00" ID-DOI="10.3897/zookeys.1226.140239.figure3" ID-arpha="D3F675BD-0C5C-5B2E-A1FC-5D77478C987C" httpUri="https://binary.pensoft.net/fig/1243034" startId="F3">
<paragraph id="14C8E8D1A5094B233E3E1591B882B382">
<label id="DD05FC9D7A9BF3061C571B716BA24DCF">Figure 3.</label>
</paragraph>
<paragraph id="41F455B433351EB5896DDFF16C59F5CD">
<taxonomicName id="095707B548CDB7FBFC2E7970E70131AA" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="emeiensis" status="sp. nov.">
<emphasis id="6622184C7AC0503EF6CB3516B8F2BE6D" italics="true">Paleosepharia emeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9C8C3CB5340700AB8A12FF09007693E4" rank="species">sp. nov.</taxonomicNameLabel>
(holotype, male)
<emphasis id="A9BC0FD4687065F3063A146B5D07F49F" bold="true">A C</emphasis>
habitus
<emphasis id="BCB2F6FCCCC4E07F0A4701AB8179B677" bold="true">D F</emphasis>
aedeagus
<emphasis id="4D86D1E88445C256ED1D6F96B364AC4B" bold="true">G</emphasis>
depression of the elytra of the male.
<emphasis id="B6C915E92ABCC63DAF01559B7472153E" bold="true">A, D</emphasis>
dorsal view
<emphasis id="884C7F4F28749B4208047BD205408CF0" bold="true">B, E</emphasis>
ventral view
<emphasis id="5C349752780A51ED6EE62BA4904D6FAC" bold="true">C, F</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="82F498823680FB6D521EB1ED7C25D2CE" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="FD76C976C33D042811B9910CFA3D8EA0" bold="true">D F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EMJAE" type="etymology">
<paragraph id="38FEDFB6D665C9C45A738049683D40DD">
<heading id="07076EA43B160589320812108E89D39C" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A6E7A93E9782D10669F7206B9BB28AC0">
This species name refers to the
<typeStatus id="DF7CB07F18464F1CF116DFCE7E6F1859">type</typeStatus>
locality.
</paragraph>
<caption id="62C83A419B0B43CCDCA8C61AF3651E4E" ID-DOI="10.3897/zookeys.1226.140239.figure4" ID-arpha="3E43CBF4-C598-5D06-BA45-D861A50EF292" httpUri="https://binary.pensoft.net/fig/1243035" startId="F4">
<paragraph id="20E93438C3BF88D904D260AF64729900">
<label id="F1F9D3A6D258ED0FDB7202468D3A758D">Figure 4.</label>
</paragraph>
<paragraph id="C823785D33B8A3CB3E91D217B502C23A">
<taxonomicName id="057B08664C547EEB3E0A288F77972028" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="emeiensis" status="sp. nov.">
<emphasis id="110997339C776F1D1B9ECB7F78977F3B" italics="true">Paleosepharia emeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EA111AC9040A176B6783CEACDBD5F08" rank="species">sp. nov.</taxonomicNameLabel>
(paratype, female)
<emphasis id="1B347E3DC410FAAA6123E92D09ACC067" bold="true">A C</emphasis>
habitus
<emphasis id="81C39717D3043F1C38AF37C4E598D599" bold="true">D</emphasis>
vaginal palps
<emphasis id="AC3C21360BD0C7C1BEF2ED3E0C5FE5AD" bold="true">E</emphasis>
spermatheca
<emphasis id="DDC46C7A322EB5C5AE1F36B7736A06ED" bold="true">F</emphasis>
apex of abdomen of the female.
<emphasis id="A49DEA2F9AB39F18719EFB26D7742AE3" bold="true">A</emphasis>
dorsal view
<emphasis id="5DE0A91E3B504E7EE19DCCE8658FFDC7" bold="true">B</emphasis>
ventral view
<emphasis id="E405CEC5D276C2A5AF484A3C6FEE913B" bold="true">C</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="E1F30F61447214778FD17153A333C159" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="F22C1A7081EF80093366B52466931629" bold="true">D, E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E2KAE" type="diagnosis">
<paragraph id="D9A691986ACCF4CA2555EE0D9BBEDE0B">
<heading id="73AC0DDE23E93EBBE77D6264F6E125A9" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="01495BDD26B77F211E2C7F8A39B68E89">
The new species closely resembles
<taxonomicName id="F1F99127C385C4BA22E2539C3CF255A1" authority="Chen, 1942" authorityName="Chen" authorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fulvicornis">
<emphasis id="B35A781110D5C5B5D238199AA6A0CCD7" italics="true">P. fulvicornis</emphasis>
<bibRefCitation id="D3D2F8521827143C54D721876AF3C4B1" author="Chen SH" journalOrPublisher="Notes dEntomologie Chinoise" pagination="9-67" refId="B6" refString="Chen SH (1942) Galerucinae nouveaux de la faune chinoise. Notes d'Entomologie Chinoise 9: 9-67." title="Galerucinae nouveaux de la faune chinoise." volume="9" year="1942">Chen, 1942</bibRefCitation>
</taxonomicName>
. It differs from
<taxonomicName id="A6BC0C760246FF73F45E162F880162FA" authorityName="Chen" authorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fulvicornis">
<emphasis id="1F8404D65CA2FD415A0032405DADF84F" italics="true">P. fulvicornis</emphasis>
</taxonomicName>
in the elytra with a longitudinal depression at base, aedeagus slender. In
<taxonomicName id="DA5F7701A8D7D05A2DFE2A807D864705" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="emeiensis">
<emphasis id="2C8DD28F24D79B0F2A0D250E79E162F7" italics="true">P. emeiensis</emphasis>
</taxonomicName>
, elytra have a hook-shaped depression at base, and aedeagus robust, 3.1 times longer than wide, parallel-sided in the basal two-thirds, apical third strongly narrowed, apical protrusions nipple-shaped. Tectum long, not reaching the apex of the aedeagus, and truncate at apex.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECMAE" type="description">
<paragraph id="F2484A9897A56CE910D7FD402C2C21F1">
<heading id="029B055104DA974172FE0DA0914A531A" reason="title">Description.</heading>
</paragraph>
<paragraph id="C5CF772F64ACCBA1D8D2DC7F9433A9EC">
<emphasis id="886DA33F5875182AD8E6AA9A90C8FFD5" bold="true">Male</emphasis>
: Length:
<quantity id="C1485F88184FF1E02BF76B2F2C6CCCF1" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.6" metricValueMin="4.4" unit="mm" value="5.0" valueMax="5.6" valueMin="4.4">4.45.6 mm</quantity>
, width:
<quantity id="E694D35F63FF6D5B44E90F2C5B212D4A" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.22.8 mm</quantity>
(mean length
<quantity id="02DDE38E84792DE982DE2823A14B4197" metricMagnitude="-3" metricUnit="m" metricValue="5.2" unit="mm" value="5.2">5.2 mm</quantity>
, mean width
<quantity id="D604319379D9728C4D2520931990C33B" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.5 mm</quantity>
,
<emphasis id="9D6EC2877AB09CFB53AB063D5186E2ED" italics="true">N</emphasis>
= 7).
</paragraph>
<paragraph id="4B444E77D2FDEF35282498E8649D50C1">General color yellow or yellowish-brown, basal part of inner margin of epipleuron, mesothorax, except mesoepisternum, metathorax and base of first metatarsomere black.</paragraph>
<paragraph id="763AE818859BF8F80C3D847C5FD2A6B3">Vertex with sparse punctures. Frontal tubercles transverse, extending downward between antenna bases. Antennae slender, 0.9 × as long as body. First antennomere shiny bare, rod-shaped, second to eleventh antennomeres with short hairs, second antennomere shortest, third antennomere 1.8 × as long as second, fourth antennomere about 1.3 × as long as third, fifth to eleventh antennomeres shorter than fourth and gradually shortened. Pronotum about 1.6 × as wide as long, lateral margins slightly curved, basal border slightly convex, apical border slightly concave, disc strongly convex, with dense punctures. Scutellum triangular, smooth, impunctate. Elytra wider than pronotum, humeri convex, disc with dense, minute punctures; and hook-shaped depression behind scutellum. Elytral epipleuron broad at base, continues towards apex. Each tibia with distinct spur at apex, segment 1 of hind tarsi longer than remaining segments combined. Aedeagus robust, with sides slightly rounded, slightly curved at apex in lateral view. Apical protrusions nipple-shaped, small, well separated from each other. Median spiculae with two pairs of slender sclerites, lateral spiculae strongly curved.</paragraph>
<paragraph id="076A854914F9ED50BD277DBFEDDCCDF6">
<emphasis id="E6BA0518D16D6C8DB1787048F598C6C1" bold="true">Female</emphasis>
: Length:
<quantity id="9BB6A6B3E9083CC28BF5417EF7529FB1" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.8" metricValueMin="4.8" unit="mm" value="5.3" valueMax="5.8" valueMin="4.8">4.85.8 mm</quantity>
, width:
<quantity id="025D2559571AC97469767C90FAA336FF" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.2" metricValueMin="2.3" unit="mm" value="2.75" valueMax="3.2" valueMin="2.3">2.33.2 mm</quantity>
(mean length
<quantity id="576C41E52D15A649B4CE04A10556FDA7" metricMagnitude="-3" metricUnit="m" metricValue="5.4" unit="mm" value="5.4">5.4 mm</quantity>
, mean width
<quantity id="AE6FB8CDEA57F4B914ACC8F4C14C47FB" metricMagnitude="-3" metricUnit="m" metricValue="2.6" unit="mm" value="2.6">2.6 mm</quantity>
,
<emphasis id="A01B9EF3EDE9EF4C9F7BF55D5C94CDAA" italics="true">N</emphasis>
= 8).
</paragraph>
<paragraph id="24AA6BC04E932DCE3D9369B049D29D1C">General color yellow or yellowish-brown, scutellum, basal part of inner margin of epipleuron, mesothorax, except mesoepisternum, metathorax and base of first metatarsomere black. Third antennomere about twice as long as second. Elytra lacking longitudinal impression. Spermatheca with big and rounded nodulus, middle part short, and cornu strongly curved. Vaginal palps with wide base and rounded apex, each palp slightly narrowing posteriorly, with 5 setae placed at apex, additional 2 setae subapically.</paragraph>
</subSubSection>
<subSubSection id="SECID0EVMAE" type="distribution">
<paragraph id="B05A48588E12084E353BE01D9CDE6690">
<heading id="6C35A7B82886F1203174A74038CBB776" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="0035BE2036071AABD4F59FF70543594F">
<collectingCountry id="077B7DE03DD2A84546886864E36F1157" name="China">China</collectingCountry>
:
<collectingRegion id="536EBDADFD5928C498FEFDBF336499B3" country="China" name="Sichuan">Sichuan</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,642 @@
<document id="26D7540DF84182FF7FEB540A0C186446" ID-CLB-Dataset="308017" ID-DOI="10.3897/zookeys.1226.137752" ID-GBIF-Dataset="616e2c2f-f236-44da-b264-bb87eaf513a7" ID-ZooBank="6891FA95-5A4A-456F-8AC3-5A14939628E8" ID-publisher-id="137752" URI-arpha="5C3BF073-C8DB-59FD-9484-28BF518C616A" URI-zoobank="http://zoobank.org/6891FA95-5A4A-456F-8AC3-5A14939628E8" article-type="research-article" checkinTime="1738866260897" checkinUser="pensoft" docAuthor="Mu, Yannan, Wang, Luyu, Zhang, Feng &amp; Zhang, Zhisheng" docDate="2025" docId="2374E035C29E54C0A0423255513F03F7" docLanguage="en" docName="ZooKeys 1226: 121-137" docOrigin="ZooKeys 1226" docSource="https://zookeys.pensoft.net/article/137752/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Mutatus tujia Mu, Wang, Zhang &amp; Zhang, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="137" masterDocId="5C3BF073C8DB59FD948428BF518C616A" masterDocTitle="Mutatus gen. nov., a new genus of Phrurolithidae (Arachnida, Araneae) from China, with descriptions of four new species" masterLastPageNumber="137" masterPageNumber="121" pageNumber="121" updateTime="1738866914701" updateUser="ExternalLinkService">
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<mods:title id="9655953CF3F5F6C8C2DFDA50085ACFF1">Mutatus gen. nov., a new genus of Phrurolithidae (Arachnida, Araneae) from China, with descriptions of four new species</mods:title>
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<mods:name id="C70AF1190DF7AF4DF284A7F8FDFFCAE6" type="personal">
<mods:role id="DA8F430A007B788FA51A5170C3C351E5">
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<mods:namePart id="CB4FDB490345EC8F55B308CDE05077AB">Mu, Yannan</mods:namePart>
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<mods:affiliation id="975B8F54806B8B7C731902D78119612C">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China &amp; Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="54D35DF490DA3F407191F9FC2901E9BD">Wang, Luyu</mods:namePart>
<mods:nameIdentifier id="CC7949F3E191DFE116D1F21BF8DE67ED" type="ORCID">0000-0002-5250-3473</mods:nameIdentifier>
<mods:affiliation id="BE120D7CC5BBDB1A875DC50D4BCFDB28">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
</mods:name>
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<mods:namePart id="D6656CD0D8725B871F2F7A9F8F3AF6B9">Zhang, Feng</mods:namePart>
<mods:nameIdentifier id="C4180D874EA3EBF053C15A0EA7EEC82D" type="ORCID">0000-0002-3347-1031</mods:nameIdentifier>
<mods:affiliation id="85392FD4AF6E5DB91A82A0CFF1D3B72D">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="586DE88150657B87BDD8272108DACAFB">Zhang, Zhisheng</mods:namePart>
<mods:nameIdentifier id="7A08007030DC47AD9F2D49CFB0C6D8E0" type="ORCID">0000-0002-9304-1789</mods:nameIdentifier>
<mods:affiliation id="64DAE220AE8DE57B86737F73E3F32453">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:date id="52E2448971BC026CEF697C33C3C6A9D2">2025</mods:date>
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<mods:number id="C4F05D7FA220111BC86DA5A6B8366657">2025-02-06</mods:number>
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<treatment id="2374E035C29E54C0A0423255513F03F7" ID-DOI="http://doi.org/10.5281/zenodo.14827172" ID-Zenodo-Dep="14827172" ID-arpha="2374E035-C29E-54C0-A042-3255513F03F7" ID-zoobank="https://zoobank.org/995C189B-E908-4132-BD2C-5F62D4BB9794" LSID="urn:lsid:plazi:treatment:2374E035C29E54C0A0423255513F03F7" LSID-ZBK="urn:lsid:zoobank.org:act:995C189B-E908-4132-BD2C-5F62D4BB9794" httpUri="http://treatment.plazi.org/id/2374E035C29E54C0A0423255513F03F7">
<subSubSection id="AF8453A8E34292AC83732D88079D20C7" type="nomenclature">
<paragraph id="318AE3DAC78B32DD52E500A30789B4B9">
<taxonomicName id="0DCAE1BCF021341A277067D911083662" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Phrurolithidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tujia" status="sp. nov.">
<emphasis id="DF48B36FFFAD1DBB0ED2F0D2B76FBC2A" italics="true">Mutatus tujia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3FD068360AEA994F4B5C327F8FC01858" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="20994A88086901B06E5A68357D14AB76" type="description">
<paragraph id="788D661428750EACC19AB9FD948E0A97">
<figureCitation id="B3480026A2D70414EB4CFBF2B3D4C054" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Mutatus tujia sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure3" httpUri="https://binary.pensoft.net/fig/1243802">Chinese name: 土家幻蛛 Figs 3</figureCitation>
,
<figureCitation id="56E102D3CC44E6FD023E6BC05E877D54" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4</figureCitation>
,
<figureCitation id="D5B1E9114FF087DC8DEB770083C0B98B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution map of Mutatus gen. nov." figureDoi="10.3897/zookeys.1226.137752.figure8" httpUri="https://binary.pensoft.net/fig/1243807">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQ5AE" type="type material">
<paragraph id="2F0EBB449B90BBF07E2B32E888EC7DA0">
<heading id="928E5BF98B404AEC916783F2396CBD2F" reason="title">Type material.</heading>
</paragraph>
<paragraph id="3CA71F26F180C5F560C2F60A0A9477F6">
<materialsCitation id="2958830BF763FBDBAB920287D995DD20" collectingDate="2023-05-19" collectorName="Z. Y. Li &amp; Z. Y. Yang" country="China" county="Shennongjia Forest District" elevation="1059" latitude="31.36663" location="Tujiazu Township" longLatPrecision="1" longitude="110.18842" municipality="Xiaguping" specimenCount="1" specimenCount-male="1" stateProvince="Hubei" typeStatus="holotype">
<emphasis id="E4C3E8A62D7CA6806A9C7FCCD7533B05" bold="true" italics="true">
<typeStatus id="A570E986FECFA1A92F25AD789F34E14C" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="387CCF76F9C2EFFF4041066351C75547" name="China">China</collectingCountry>
<specimenCount id="B6FB70BD1C3784CF3303254AC2BD10C9" count="1" type="male"></specimenCount>
;
<collectingRegion id="EBF826B96571009159F7AE9B57B3E7E9" country="China" name="Hubei">Hubei province</collectingRegion>
,
<collectingCounty id="DE0BA8B94242CEB07DDD2753CAF1C634">Shennongjia Forest District</collectingCounty>
,
<collectingMunicipality id="8C32E07A603029991C1F600AB93FF0BF">Xiaguping</collectingMunicipality>
,
<location id="00EDA177918ABBAB4FDB1CC1D2BEDA17" LSID="urn:lsid:plazi:treatment:2374E035C29E54C0A0423255513F03F7:00EDA177918ABBAB4FDB1CC1D2BEDA17" country="China" county="Shennongjia Forest District" latitude="31.36663" longLatPrecision="1" longitude="110.18842" municipality="Xiaguping" name="Tujiazu Township" stateProvince="Hubei">Tujiazu Township</location>
,
<location id="678F1912821C63B3C2C5A85BA3C169AF" LSID="urn:lsid:plazi:treatment:2374E035C29E54C0A0423255513F03F7:678F1912821C63B3C2C5A85BA3C169AF" country="China" county="Shennongjia Forest District" latitude="31.36663" longLatPrecision="1" longitude="110.18842" municipality="Xiaguping" name="Huangxi River" stateProvince="Hubei">Huangxi River</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E55AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[110.188422,31.366631]}">
<geoCoordinate id="85F98AA2BA2086F6451125B4CDE7329A" degrees="31" direction="north" minutes="21" orientation="latitude" precision="1" seconds="59.87" value="31.36663">31 ° 21 ' 59.87 &quot; N</geoCoordinate>
,
<geoCoordinate id="CDD16743528FB791AB5F4E0F7FA4318A" degrees="110" direction="east" minutes="11" orientation="longitude" precision="1" seconds="18.32" value="110.18842">110 ° 11 ' 18.32 &quot; E</geoCoordinate>
</named-content>
, elev.
<quantity id="A005594BEE9F5C8452F45A0B00E654B6" metricMagnitude="3" metricUnit="m" metricValue="1.059" unit="m" value="1059.0">
<elevation id="FC81C89A2758F5FE5BF5B5085020FC56" metricMagnitude="3" metricUnit="m" metricValue="1.059" unit="m" value="1059.0">1059 m</elevation>
</quantity>
,
<date id="95F1DD424F3AC79C5EE0ABDB1595553A" value="2023-05-19">
<collectingDate id="7D0B4DA45428213DCE3158FBCD560E76" value="2023-05-19">19 May 2023</collectingDate>
</date>
;
<collectorName id="82F8AB4F528349BB2132F012705953A3">Z. Y. Li</collectorName>
,
<collectorName id="BDF473E67966632C8C3D48C194B96806">Z. Y. Yang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="A6980A6650345ABA2D04B94B78BC6C26" collectingDate="2023-05-19" collectorName="Z. Y. Li &amp; Z. Y. Yang" country="China" county="Shennongjia Forest District" elevation="1059" latitude="31.36663" location="Tujiazu Township" longLatPrecision="1" longitude="110.18842" municipality="Xiaguping" specimenCount="8" specimenCount-female="7" specimenCount-male="1" stateProvince="Hubei" typeStatus="paratype">
<emphasis id="E6430ED6ED62492FE1B880A0C8AE5EA5" bold="true" italics="true">
<typeStatus id="EBCA36680C57441D2F8F6484AE55C775" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCount id="F3393CD9A00E834511EB82768EA114BE" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="66117C6482182523B425983B8015404D" count="7" type="female">7 ♀</specimenCount>
, with same data as holotype
</materialsCitation>
; •
<materialsCitation id="8C347671D81EDEEA8ED6F8C2F2309887" collectingDate="2023-05-19" collectorName="Z. Y. Li &amp; Z. Y. Yang" country="China" county="Shennongjia Forest District" elevation="1053" latitude="31.381737" location="Dongkou Village" longLatPrecision="1" longitude="110.16348" municipality="Xiaguping" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Hubei" typeStatus="paratype">
<specimenCount id="DF22140C506DD03A349DE069E2E831A0" count="2" type="male">2 ♂</specimenCount>
<specimenCount id="51099FF7C6A0F91B959904EB45CF1B72" count="4" type="female">4 ♀</specimenCount>
same locality,
<location id="E064750428DBD361C14F8FB75201FD28" LSID="urn:lsid:plazi:treatment:2374E035C29E54C0A0423255513F03F7:E064750428DBD361C14F8FB75201FD28" latitude="31.381737" longLatPrecision="1" longitude="110.16348" name="Dongkou Village">Dongkou Village</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJ6AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[110.163481,31.381736]}">
<geoCoordinate id="0E355F44338313C3ED6A6B612D0B2CA2" degrees="31" direction="north" minutes="22" orientation="latitude" precision="1" seconds="54.25" value="31.381737">31 ° 22 ' 54.25 &quot; N</geoCoordinate>
,
<geoCoordinate id="6FD8FB5F863CB4E3FED671B68D7B394C" degrees="110" direction="east" minutes="9" orientation="longitude" precision="1" seconds="48.53" value="110.16348">110 ° 9 ' 48.53 &quot; E</geoCoordinate>
</named-content>
,
<elevation id="134E0880E15966BBE71D9DC1315F0290" metricMagnitude="3" metricUnit="m" metricValue="1.053" unit="m" value="1053.0">
<quantity id="D0235C288DE856958F1D18EA71143099" metricMagnitude="3" metricUnit="m" metricValue="1.053" unit="m" value="1053.0">1053 m</quantity>
a. s. l.
</elevation>
),
<date id="8D5BBE2328314DC9CC6692B791E45C31" value="2023-05-19">
<collectingDate id="B56B77A219B58ACA083F579DC9AB696C" value="2023-05-19">19 May 2023</collectingDate>
</date>
,
<collectorName id="E752244C043C3F1463C8FA2424B50F79">Z. Y. Li</collectorName>
,
<collectorName id="3981E5A8F9EF8A375FB7A3F1629FD0BB">Z. Y. Yang</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EO6AE" type="etymology">
<paragraph id="144F326D545E4BA9F0F80B437F18F263">
<heading id="21C58031592D40702A1D49FDBD2662BB" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="542FBEE8EFC8B82DCFAB33840279AEAA">The specific name comes from the word “ Tujia ”. The name is one of the Chinese ethnic minorities that live adjacent to the area inhabited by the new species; a noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="SECID0ET6AE" type="diagnosis">
<paragraph id="727C39808B65EE8757FE30C6266FC443">
<heading id="1E748CF5E92B008529503D9A0D8C6519" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="9D7E039EABD51E574897D133A49B5D41">
The new species resembles
<taxonomicName id="AD8A12398C90E2F28244F0651930C53B" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tianyan" status="sp. nov.">
<emphasis id="4BFB7FB31B455E9D47FD5FB4890666E3" italics="true">M. tianyan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BA1E1FCFBE4C76065A6D8232FACC1F13" rank="species">sp. nov.</taxonomicNameLabel>
in having similar truncated prolateral tegular apophysis and embolus, but it can be differentiated by the following: posterior of tegular sheet tail-shaped (vs tongue-shaped; compare Fig.
<figureCitation id="8BD53EB7C36F81FFC6591A489341A7A4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 D</figureCitation>
with Fig.
<figureCitation id="557C315E3F962F968EC0DB3D84F4FA0F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 C</figureCitation>
); median septum thin (vs thick; compare Fig.
<figureCitation id="F6D813F3DB1FEC4E0B14963835C79954" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 E</figureCitation>
with Fig.
<figureCitation id="255C9C99174E81BD354E68BE2313655C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 E</figureCitation>
); bursa large, balloon-shaped (vs small, rectangular-shaped; compare Fig.
<figureCitation id="4699D06A65D8DE964E7768FF32E36818" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 F</figureCitation>
with Fig.
<figureCitation id="B71228ED20194942162CDE3DED195B0A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 F</figureCitation>
); glandular appendages distinct, large (vs indistinct, small; compare Fig.
<figureCitation id="251914CC9B14739C6E8AF46282B5882D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 F</figureCitation>
with Fig.
<figureCitation id="BFBEBA9C0C9A9390525BE04FBFFA7887" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFBAG" type="description">
<paragraph id="906F5E7B5546889E5A5320284D8AAD02">
<heading id="D2D0940CA5AC81C30FBE274DA433AF07" reason="title">Description.</heading>
</paragraph>
<paragraph id="93CC2DF98ACDDCFDE35113231151CA43">
Male (
<typeStatus id="ECE3FC0174E9458D648361D0F10C9E09" type="holotype">holotype</typeStatus>
, Fig.
<figureCitation id="A82A22BE1F14A007AEC870DC4BE50089" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Mutatus tujia sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure3" httpUri="https://binary.pensoft.net/fig/1243802">3 A, B</figureCitation>
). Total length 3.34, carapace 1.85 long, 1.61 wide; abdomen 1.35 long, 1.22 wide. Eye sizes and interdistances:
<abbrev id="ABBRID0EPBAG" xlink_title="anterior median eye">AME</abbrev>
0.09,
<abbrev id="ABBRID0ETBAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.13,
<abbrev id="ABBRID0EXBAG" xlink_title="posterior median eye">PME</abbrev>
0.07,
<abbrev id="ABBRID0E2BAG" xlink_title="posterior lateral eye">PLE</abbrev>
0.08;
<abbrev id="ABBRID0E6BAG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0EDCAG" xlink_title="anterior median eye">AME</abbrev>
0.06,
<abbrev id="ABBRID0EHCAG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0ELCAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0EPCAG" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0ETCAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.27,
<abbrev id="ABBRID0EXCAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E2CAG" xlink_title="posterior median eye">PME</abbrev>
0.11,
<abbrev id="ABBRID0E6CAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EDDAG" xlink_title="posterior lateral eye">PLE</abbrev>
0.07,
<abbrev id="ABBRID0EHDAG" xlink_title="posterior lateral eye">PLE</abbrev>
<abbrev id="ABBRID0ELDAG" xlink_title="posterior lateral eye">PLE</abbrev>
0.45,
<abbrev id="ABBRID0EPDAG" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0ETDAG" xlink_title="posterior lateral eye">PLE</abbrev>
0.10.
<abbrev id="ABBRID0EXDAG" xlink_title="eye area width">EAW</abbrev>
0.58,
<abbrev id="ABBRID0E2DAG" xlink_title="cephalic region width">CRW</abbrev>
0.83,
<abbrev id="ABBRID0E6DAG" xlink_title="eye area width">EAW</abbrev>
/
<abbrev id="ABBRID0EDEAG" xlink_title="cephalic region width">CRW</abbrev>
0.70,
<abbrev id="ABBRID0EHEAG" xlink_title="cephalic region width">CRW</abbrev>
/
<abbrev id="ABBRID0ELEAG" xlink_title="carapace width">CW</abbrev>
0.52.
<abbrev id="ABBRID0EPEAG" xlink_title="median ocular area">MOA</abbrev>
0.25 long, anterior width 0.22, posterior width 0.29.
<abbrev id="ABBRID0ETEAG" xlink_title="clypeal height">
<collectingCountry id="A02B9D9FA11091D811ADB4CBF4CA92AA" name="Switzerland">CH</collectingCountry>
</abbrev>
0.11,
<abbrev id="ABBRID0EXEAG" xlink_title="clypeal height">
<collectingCountry id="FB20C0FB0F879B203D909E9AC6823368" name="Switzerland">CH</collectingCountry>
</abbrev>
/
<abbrev id="ABBRID0E2EAG" xlink_title="anterior median eye">AME</abbrev>
1.22. Labium 0.35 long, 0.30 wide. Sternum 1.21 long, 1.17 wide. Carapace oval, cervical groove inconspicuous. Sternum, labium, and endite brown, without pattern. Abdomen dark brown, smooth, with large dorsal scutum covering almost entire abdomen. Legs brown.
</paragraph>
<caption id="5E56AB2746449563D10625853F98DC72" ID-DOI="10.3897/zookeys.1226.137752.figure3" ID-arpha="059BB2CA-3A1A-5DCD-9507-568B7925456B" httpUri="https://binary.pensoft.net/fig/1243802" startId="F3">
<paragraph id="29B65ABA99320D6A5DB22748C67571EF">
<label id="A501960A9E4AF8C6A73891B653AE3951">Figure 3.</label>
</paragraph>
<paragraph id="9BC4757149DBA50D3E7257216F3F89C4">
Habitus of
<taxonomicName id="07903F285E6989C411EEA48076961EBA" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tujia" status="sp. nov.">
<emphasis id="EACD70DE6B5F648366DC82DD0A76B5EB" italics="true">Mutatus tujia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0E1BA8EAA5979696BC52A206F5371592" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1E62EBF4488ACAD1398ABD493E19C5AB" bold="true">A</emphasis>
male holotype, dorsal view
<emphasis id="489BEC415ED69C87209B74F53F0A3D46" bold="true">B</emphasis>
same, ventral view
<emphasis id="EF97F52015702C73FEFA0B2DD38449DB" bold="true">C</emphasis>
female paratype, dorsal view
<emphasis id="5FE51D5E65A68280EB7DB2F5C408758D" bold="true">D</emphasis>
same, ventral view.
</paragraph>
</caption>
<paragraph id="B4B1C359C357BB9CF73A9BDEB07A038F">
Palp (Fig.
<figureCitation id="020EBAC15AF12E2143A5FB2F0275D8C1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 A D</figureCitation>
). Femoral apophysis (
<abbrev id="ABBRID0EFGAG" xlink_title="femur apophysis">FA</abbrev>
) well developed, thumb-shaped, located at middle part. Patella as long as tibia, about 1 / 3 length of femur. Tibia longer than wide, with 2 apophyses, retroventral apophysis (
<abbrev id="ABBRID0EJGAG" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
) tail-shaped; retrolateral apophysis (
<abbrev id="ABBRID0ENGAG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
) strong, longer and wider than retroventral apophysis, tapering towards tip. Bulb pyriform, sperm duct (
<abbrev id="ABBRID0ERGAG" xlink_title="sperm duct">SD</abbrev>
) distinct, V-shaped. Prolateral tegular apophysis (
<abbrev id="ABBRID0EVGAG" xlink_title="Prolateral tegular apophysis">PTA</abbrev>
) coniform, slightly curved, originating at nearly 9 oclock position. Tegular sheet (
<abbrev id="ABBRID0EZGAG" xlink_title="tegular sheet">TS</abbrev>
) tobacco pipe-shaped, anterior part large and wide, posterior part small, tail-shaped. Embolus (E) needle-like, tapering towards tip, slightly curved.
</paragraph>
<caption id="E4623A7A0E0110D4C3A74D26EDAF3524" ID-DOI="10.3897/zookeys.1226.137752.figure4" ID-arpha="EF56FBCA-1886-5E03-BB67-80FE2270292C" httpUri="https://binary.pensoft.net/fig/1243803" startId="F4">
<paragraph id="26136CA6CC1E10D144A164898E3EE041">
<label id="BF422362CB1A74B16A4A6747411CA574">Figure 4.</label>
</paragraph>
<paragraph id="E534E2AE38581E701C9C40428E51A372">
Copulatory organs of
<taxonomicName id="B58B2BAFBD9E6E4FCCE3488E21887245" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tujia" status="sp. nov.">
<emphasis id="0449FAD53427E8EACB04CDDF09958B12" italics="true">Mutatus tujia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CFF3D5C15126F56E99BA73690772BA56" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="53B2C8391CE41300893B25070DE31055" bold="true">A</emphasis>
male left palp, prolateral view
<emphasis id="0E7428234C13E7B7FC4FCD42127AC006" bold="true">B</emphasis>
same, retrolateral view
<emphasis id="7BA86E1CBE2108255E48D57EAE91C03D" bold="true">C</emphasis>
same, dorsal view
<emphasis id="47246DB0805129ED3C2E1EE51519A621" bold="true">D</emphasis>
same, ventral view
<emphasis id="38B67DC46681592786466BA7CB66DFF1" bold="true">E</emphasis>
epigyne, ventral view
<emphasis id="90D11F863F4A0807031C16D03541E519" bold="true">F</emphasis>
same, dorsal view. Abbreviations: E — embolus;
<abbrev id="ABBRID0E3HAG" xlink_title="femur apophysis">FA</abbrev>
— femoral apophysis;
<abbrev id="ABBRID0EAIAG" xlink_title="Prolateral tegular apophysis">PTA</abbrev>
— prolateral tegular apophysis;
<abbrev id="ABBRID0EEIAG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
— retrolateral tibial apophysis;
<abbrev id="ABBRID0EIIAG" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
— retroventral tibial apophysis;
<abbrev id="ABBRID0EMIAG" xlink_title="sperm duct">SD</abbrev>
— sperm duct;
<abbrev id="ABBRID0EQIAG" xlink_title="tegular sheet">TS</abbrev>
— tegular sheet; B — bursa;
<abbrev id="ABBRID0EUIAG" xlink_title="Copulatory ducts">CD</abbrev>
— copulatory duct;
<abbrev id="ABBRID0EYIAG" xlink_title="Copulatory openings">CO</abbrev>
— copulatory opening;
<abbrev id="ABBRID0E3IAG" xlink_title="connecting tube">CT</abbrev>
— connecting tube;
<abbrev id="ABBRID0EAJAG" xlink_title="Fertilization ducts">FD</abbrev>
— fertilization duct;
<abbrev id="ABBRID0EEJAG" xlink_title="Glandular appendages">GA</abbrev>
— glandular appendage;
<abbrev id="ABBRID0EIJAG" xlink_title="median septum">MS</abbrev>
— median septum; S — spermathecae.
</paragraph>
</caption>
<paragraph id="3EEF25EE76447771EA8E9E93427C57C0">
Female (one of the
<typeStatus id="39D1A7139DEDBD4E080909AD76BBE281" type="paratype">paratypes</typeStatus>
; Fig.
<figureCitation id="982B4113050393A69D6E00836D11A672" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Mutatus tujia sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure3" httpUri="https://binary.pensoft.net/fig/1243802">3 C, D</figureCitation>
). Total length 3.50, carapace 1.66 long, 1.49 wide; abdomen 1.86 long, 1.25 wide. Eye sizes and interdistances:
<abbrev id="ABBRID0EXJAG" xlink_title="anterior median eye">AME</abbrev>
0.08,
<abbrev id="ABBRID0E2JAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.09,
<abbrev id="ABBRID0E6JAG" xlink_title="posterior median eye">PME</abbrev>
0.07,
<abbrev id="ABBRID0EDKAG" xlink_title="posterior lateral eye">PLE</abbrev>
0.08;
<abbrev id="ABBRID0EHKAG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0ELKAG" xlink_title="anterior median eye">AME</abbrev>
0.06,
<abbrev id="ABBRID0EPKAG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0ETKAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0EXKAG" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0E2KAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.23,
<abbrev id="ABBRID0E6KAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EDLAG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0EHLAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0ELLAG" xlink_title="posterior lateral eye">PLE</abbrev>
0.06,
<abbrev id="ABBRID0EPLAG" xlink_title="posterior lateral eye">PLE</abbrev>
<abbrev id="ABBRID0ETLAG" xlink_title="posterior lateral eye">PLE</abbrev>
0.35,
<abbrev id="ABBRID0EXLAG" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0E2LAG" xlink_title="posterior lateral eye">PLE</abbrev>
0.09.
<abbrev id="ABBRID0E6LAG" xlink_title="eye area width">EAW</abbrev>
0.48,
<abbrev id="ABBRID0EDMAG" xlink_title="cephalic region width">CRW</abbrev>
0.71,
<abbrev id="ABBRID0EHMAG" xlink_title="eye area width">EAW</abbrev>
/
<abbrev id="ABBRID0ELMAG" xlink_title="cephalic region width">CRW</abbrev>
0.68,
<abbrev id="ABBRID0EPMAG" xlink_title="cephalic region width">CRW</abbrev>
/
<abbrev id="ABBRID0ETMAG" xlink_title="carapace width">CW</abbrev>
0.47.
<abbrev id="ABBRID0EXMAG" xlink_title="median ocular area">MOA</abbrev>
0.23 long, anterior width 0.23, posterior width 0.18.
<abbrev id="ABBRID0E2MAG" xlink_title="clypeal height">
<collectingCountry id="16658015513D89E0DBE23529EDFD9886" name="Switzerland">CH</collectingCountry>
</abbrev>
0.10,
<abbrev id="ABBRID0E6MAG" xlink_title="clypeal height">
<collectingCountry id="ADE6871EB9B48A35AD5D7DE66691A209" name="Switzerland">CH</collectingCountry>
</abbrev>
/
<abbrev id="ABBRID0EDNAG" xlink_title="anterior median eye">AME</abbrev>
1.25. Labium 0.23 long, 0.20 wide. Sternum 0.87 long, 0.75 wide. Other characters as for male, except dorsal scutum absent, color black and body slightly larger.
</paragraph>
<paragraph id="D29DCD3B142F54991EE0D2AEBFAA4D46">
Epigyne (Fig.
<figureCitation id="505115D7E8C0BA2CE2394EF7D0F9D760" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 E, F</figureCitation>
). Epigynal plate slightly sclerotized, with a pair of atria, a subcircular tumor located at middle part of lateral margin of atrium. Median septum (
<abbrev id="ABBRID0ENNAG" xlink_title="median septum">MS</abbrev>
) dumbbell-shaped. Copulatory openings (
<abbrev id="ABBRID0ERNAG" xlink_title="Copulatory openings">CO</abbrev>
) slit-like, located at anterior of atrium. Copulatory ducts (
<abbrev id="ABBRID0EVNAG" xlink_title="Copulatory ducts">CD</abbrev>
) straight, short, indistinct. Connecting tubes (
<abbrev id="ABBRID0EZNAG" xlink_title="connecting tube">CT</abbrev>
) distinct. Bursae (B) large, nearly balloon-shaped, translucent. Glandular appendages (
<abbrev id="ABBRID0E4NAG" xlink_title="Glandular appendages">GA</abbrev>
) mastoid-shaped. Spermathecae (S) clavate, anterior part close but separated posteriorly. Fertilization (
<abbrev id="ABBRID0EBOAG" xlink_title="Fertilization ducts">FD</abbrev>
) ducts located at posterior end of spermathecae.
</paragraph>
<paragraph id="8FD5C966DD2F8B6F26CD1B697611E0B0">Measurement of legs:</paragraph>
<paragraph id="3A6F5EA1C0951F53C7B59B15F48D0F9F">
<table id="TID0EHLAG" rules="all">
<tbody id="E40225D4E77BB05394ECAE8543FB2F88">
<table-wrap id="T3" orientation="portrait" position="float">
<tr id="F4C1AAECB133CF0649CAD764B7AD93BC">
<th id="423B025C53B704721853087DFDFF7F18" colspan="1" rowspan="1">male / female</th>
<th id="64D6D61F31566C94EA9A69AC04ABB24C" colspan="1" rowspan="1">Fe</th>
<th id="8DF8D311D4978144161A798384FAAA3F" colspan="1" rowspan="1">Pa</th>
<th id="8A6FA3C42374505569C0DFA05DF29E7E" colspan="1" rowspan="1">Ti</th>
<th id="EBA7FC1DD5E58DA97BDC3CA2D00880FD" colspan="1" rowspan="1">Me</th>
<th id="3FE00CFEDA16E338AFFDC9486F110D51" colspan="1" rowspan="1">Ta</th>
<th id="7166BE8989D7230EB257380D57BCC4EA" colspan="1" rowspan="1">Total</th>
</tr>
<tr id="EF4C967ED1430FBC1BCFA398E76082F6">
<td id="AE370E0353F5EDF0CE5EA949DE02A069" colspan="1" rowspan="1">Leg I</td>
<td id="07C85B7314A385DBFB9D82A14C4C59A1" colspan="1" rowspan="1">1.38 / 1.44</td>
<td id="FB8C302FCA2083F994506C33FF0C5656" colspan="1" rowspan="1">0.43 / 0.46</td>
<td id="A24F5793C7FFB537C1A8CF06F674D12E" colspan="1" rowspan="1">1.42 / 1.58</td>
<td id="B29D393C4EE36B907ADE188CDB769549" colspan="1" rowspan="1">1.14 / 1.34</td>
<td id="0CC9A9A6A512669AD7844606DE429488" colspan="1" rowspan="1">0.61 / 0.69</td>
<td id="3D83C154F12A5C919BFF195C51FBE933" colspan="1" rowspan="1">4.98 / 5.51</td>
</tr>
<tr id="64A403CE36C15238B19048FE4E98A7E5">
<td id="B0CE88A73F12361B70B59DB9DFE4066E" colspan="1" rowspan="1">Leg II</td>
<td id="96BCEA5A98DD61DFBE340F884E9BD8CD" colspan="1" rowspan="1">1.20 / 1.19</td>
<td id="40AE1581888ABBC462D11E2449270FCB" colspan="1" rowspan="1">0.45 / 0.44</td>
<td id="20053404EE68D8D4CCD79E992D4D4323" colspan="1" rowspan="1">1.10 / 1.21</td>
<td id="8AB9C5F6338A2036BED9E7EB2EA4B918" colspan="1" rowspan="1">0.96 / 1.07</td>
<td id="1A7ECD3556EDE985B582FF7907F14AF6" colspan="1" rowspan="1">0.58 / 0.58</td>
<td id="435B22A54D12285AA2AD963314E2AC0A" colspan="1" rowspan="1">4.29 / 4.40</td>
</tr>
<tr id="FBB844F88D119F31BA253B734D77AACA">
<td id="921CECA14DC4D1102FB21105F60A3B73" colspan="1" rowspan="1">Leg III</td>
<td id="20EBF3897AEB7B51E97BDA3D469C1B80" colspan="1" rowspan="1">1.00 / 1.01</td>
<td id="A11E4D448B50D1A4363DC29FAA7D4F57" colspan="1" rowspan="1">0.39 / 0.39</td>
<td id="DE07E7B8C5CC00FB72ED4C37D2910934" colspan="1" rowspan="1">0.80 / 0.81</td>
<td id="1464BF97BE67584E17D68785E482E9CB" colspan="1" rowspan="1">0.91 / 1.01</td>
<td id="52EA8ED7E79449934ABADE90E54CFEEC" colspan="1" rowspan="1">0.57 / 0.58</td>
<td id="943DAAD9ABD9F37D30FF8633A8F26B2C" colspan="1" rowspan="1">3.67 / 3.80</td>
</tr>
<tr id="0A785297473878100FFA3F2C29FB3058">
<td id="A9CDF94D929FCA8B598575EC0EA9DBB0" colspan="1" rowspan="1">Leg IV</td>
<td id="61EB814732EF3C6F8924BE0A7E246BD7" colspan="1" rowspan="1">1.50 / 1.62</td>
<td id="AC4230ECC8E49041F79A17C17868BEFE" colspan="1" rowspan="1">0.44 / 0.46</td>
<td id="92E9464E0CDF024C6E6180A3CE225F00" colspan="1" rowspan="1">1.32 / 1.53</td>
<td id="1332D456BBE679D287C5C3FD06C4C8CE" colspan="1" rowspan="1">1.45 / 1.60</td>
<td id="EF0369A87B4D99846A677A48389DE8F5" colspan="1" rowspan="1">0.77 / 0.89</td>
<td id="AAFC59AFF007E254D08BFD04D73E67BA" colspan="1" rowspan="1">5.48 / 6.10</td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
<paragraph id="5E01189861CFC3CABB192CC2DFB95C5E">Spination of legs:</paragraph>
<paragraph id="6F1ACA0B7F22483EF4EEBD090293446E">
<table id="TID0EGRAG" rules="all">
<tbody id="6A130DD5FA762F39D8785FDED528C343">
<table-wrap id="T4" orientation="portrait" position="float">
<tr id="BFE124B126FF1DF131DBB9226A47830F">
<th id="5E3D1B10D1DB7462C8409080DCFB486F" colspan="2" isEmpty="true" rowspan="1">-</th>
<th id="71A762B61A96DCD04302E7F8E8524C75" colspan="1" rowspan="1">Fe</th>
<th id="30057F89E65026EE58D95BB0D4D28BE5" colspan="1" rowspan="1">Pa</th>
<th id="69651F474324B59C545300C2A6724D6C" colspan="1" rowspan="1">Ti</th>
<th id="5679BB4F2078265E879FCEA6F351B74D" colspan="1" rowspan="1">Me</th>
<th id="0DE77D12884A2BE7B21675AEFF7CC32E" colspan="1" rowspan="1">Ta</th>
</tr>
<tr id="EB8391B087546E2D048408E5CBC338A4">
<td id="F0101E6D58EA6C4EB703913A471AFD53" colspan="1" rowspan="4">male</td>
<td id="C38E2D5CFD8C363962F265B73D95F4FB" colspan="1" rowspan="1">Leg I</td>
<td id="2AF95C9F1ECCE7072A5F6695FBBEDDFA" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EHTAG" xlink_title="prolateral">pl</abbrev>
3
</td>
<td id="05D81059D3EEDA8AFF3A334828F409EB" colspan="1" rowspan="1"></td>
<td id="755EFCE62E4B9012B24BE6E08406CDA7" colspan="1" rowspan="1">
<abbrev id="ABBRID0ESTAG" xlink_title="proventral">pv</abbrev>
7
<abbrev id="ABBRID0EWTAG" xlink_title="retroventral">rv</abbrev>
7
</td>
<td id="A17F40A149AA4CCFBFFD987A995DD0C4" colspan="1" rowspan="1">
<abbrev id="ABBRID0E5TAG" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0ECUAG" xlink_title="retroventral">rv</abbrev>
4
</td>
<td id="B66F307B94640FAF239360FA9438EF6C" colspan="1" rowspan="1"></td>
</tr>
<tr id="09B2F865BFDC435876695642D3E27A63">
<td id="9EA6B90E565899763157E16376462259" colspan="1" rowspan="1">Leg II</td>
<td id="C5A62FB41DAE4EE2A0A3A18B672AB295" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0ERUAG" xlink_title="prolateral">pl</abbrev>
0
</td>
<td id="CFBEE178E219DE8DF22115F657B792F4" colspan="1" rowspan="1"></td>
<td id="6639A6B7CD88F892A9C527D6FC4C6C98" colspan="1" rowspan="1">
<abbrev id="ABBRID0E3UAG" xlink_title="proventral">pv</abbrev>
6
<abbrev id="ABBRID0EAVAG" xlink_title="retroventral">rv</abbrev>
6
</td>
<td id="169A9D444B2A4C8DEC058DF0FF43758C" colspan="1" rowspan="1">
<abbrev id="ABBRID0EIVAG" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0EMVAG" xlink_title="retroventral">rv</abbrev>
3
</td>
<td id="B4416F12D1AD1972669BAE3664C66003" colspan="1" rowspan="1"></td>
</tr>
<tr id="BC2A32D601029B693F99A5B1CA030899">
<td id="51FF383D1561D3A03D7BEA93E23A92C6" colspan="1" rowspan="1">Leg III</td>
<td id="97D89F80620F6EE85654A48AB80A6ED1" colspan="1" rowspan="1">d 1</td>
<td id="ED10C39D807164EAEBBBB3B3FB77B146" colspan="1" rowspan="1"></td>
<td id="98441B3D64751B02AF4DA2F923153A48" colspan="1" rowspan="1"></td>
<td id="49539BB377EDA9B7DF31643031BD916C" colspan="1" rowspan="1"></td>
<td id="420119B854E7B08689941E94A2EF6C55" colspan="1" rowspan="1"></td>
</tr>
<tr id="1A195B97A0D08AC82AE051BF068D3B9A">
<td id="49FCC2A0F85184EDA3DFD3886E59A439" colspan="1" rowspan="1">Leg IV</td>
<td id="05A0CEA9F8BCA097B5D072E89A981847" colspan="1" rowspan="1">d 1</td>
<td id="391797241DB3010175BF51DFBD5E667B" colspan="1" rowspan="1"></td>
<td id="113D0B1562BD26C0B90F328EEBFAA54D" colspan="1" rowspan="1"></td>
<td id="4BB725A13257DDD77733A43D4EFF4370" colspan="1" rowspan="1"></td>
<td id="81FCF0FC785E77443E9AA71F32C82781" colspan="1" rowspan="1"></td>
</tr>
<tr id="0D21EEBAC11A650E079E04B417B2B2B1">
<td id="D54B6077EF5C507D544A0A20E58473A7" colspan="1" rowspan="4">female</td>
<td id="AE498072D0DE83FBF41E93D6DB2F80A9" colspan="1" rowspan="1">Leg I</td>
<td id="E7D63B2DF619A7C7FB415EFFAD9C00DB" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EEXAG" xlink_title="prolateral">pl</abbrev>
3
</td>
<td id="8329AC69DF4E94C5F442A9398AD469F1" colspan="1" rowspan="1"></td>
<td id="27FB118305C897BD6592E4C71D599B0C" colspan="1" rowspan="1">
<abbrev id="ABBRID0EPXAG" xlink_title="proventral">pv</abbrev>
6
<abbrev id="ABBRID0ETXAG" xlink_title="retroventral">rv</abbrev>
6
</td>
<td id="429D868D426211555820FD989B65AB9A" colspan="1" rowspan="1">
<abbrev id="ABBRID0E2XAG" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0E6XAG" xlink_title="retroventral">rv</abbrev>
4
</td>
<td id="CFD0F6801BCB86975220ED0798209074" colspan="1" rowspan="1"></td>
</tr>
<tr id="A03D7458757875E7866C5AAE8E6941E4">
<td id="205EE789FB24AF2B632D7CFC54F2D220" colspan="1" rowspan="1">Leg II</td>
<td id="E507DC2C990B9E09D49A3C9EEF74C477" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EOYAG" xlink_title="prolateral">pl</abbrev>
0
</td>
<td id="5D14D0AAB7AB4A651A6F669DE8C0FDB4" colspan="1" rowspan="1"></td>
<td id="C705309957F6D7C5A87110E0D0186C3A" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZYAG" xlink_title="proventral">pv</abbrev>
6
<abbrev id="ABBRID0E4YAG" xlink_title="retroventral">rv</abbrev>
6
</td>
<td id="41F6C16736059DC44FCF869B261E91D7" colspan="1" rowspan="1">
<abbrev id="ABBRID0EFZAG" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0EJZAG" xlink_title="retroventral">rv</abbrev>
4
</td>
<td id="D17CBE1A14F3A1BCBC759782F4BE51AD" colspan="1" rowspan="1"></td>
</tr>
<tr id="C7653B96DC0166D710F7AD1FCBA6B797">
<td id="67DD0639CB39821524A4AE32B6E6DCE9" colspan="1" rowspan="1">Leg III</td>
<td id="5F3A060CA784D649FED58B8D28F3DD5C" colspan="1" rowspan="1">d 1</td>
<td id="5977BB4775F5594338F2BA89D6A3C39D" colspan="1" rowspan="1"></td>
<td id="3FC5BA301E1720E45CEEC9F06CAC90A3" colspan="1" rowspan="1"></td>
<td id="400FDA6A8418A80197C6C3BA69967B5B" colspan="1" rowspan="1"></td>
<td id="3468AA58CA24080F2D1EBC7EC9C71DD4" colspan="1" rowspan="1"></td>
</tr>
<tr id="7E87FC46135C172C696CD46546448486">
<td id="34C497ADF76FB6CF2355B7003C46F28D" colspan="1" rowspan="1">Leg IV</td>
<td id="1BF2954C564AEFC41CCC6BE2BAA47222" colspan="1" rowspan="1">d 1</td>
<td id="6E909C0CF159AEDF9820BBB05BB26E05" colspan="1" rowspan="1"></td>
<td id="9EA9736AAD183A10A5FA2F0E49D863CA" colspan="1" rowspan="1"></td>
<td id="B91FB92F31993B47009522B8298279D9" colspan="1" rowspan="1"></td>
<td id="6CB9B54C936ABDAB529B09C5224ABC95" colspan="1" rowspan="1"></td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EW1AG" type="distribution">
<paragraph id="C1BA97C347429DBF03E43FBB5945970C">
<heading id="BEBE1695230CECFE410DF6ECF21E2F77" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="42313AD5B6B8F9DC850F544EEA42F54B">
Known only from the
<typeStatus id="217A001EB0BD967574D7035B6FFA833B">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,443 @@
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<treatment id="2643064135E357ACBD984644659ADEAB" ID-DOI="http://doi.org/10.5281/zenodo.14827176" ID-Zenodo-Dep="14827176" ID-arpha="26430641-35E3-57AC-BD98-4644659ADEAB" ID-zoobank="https://zoobank.org/942E6A33-B2A8-4B4E-9A5C-9EFBEB2379AE" LSID="urn:lsid:plazi:treatment:2643064135E357ACBD984644659ADEAB" LSID-ZBK="urn:lsid:zoobank.org:act:942E6A33-B2A8-4B4E-9A5C-9EFBEB2379AE" httpUri="http://treatment.plazi.org/id/2643064135E357ACBD984644659ADEAB">
<subSubSection id="641EA315D6715E759198259B38C06878" type="nomenclature">
<paragraph id="DCA47704BDC9545CF2F8EC6956B5E2DE">
<taxonomicName id="5A04217540B057079DA54FB858391C8B" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Phrurolithidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yintiaoling" status="sp. nov.">
<emphasis id="EA6AF482573EC7F9E1F3311D4196DDA5" italics="true">Mutatus yintiaoling</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CF4CD4952DC3834777963029F231F6F" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F8F3FF6CB4E387A69E8A101CFF6DFD6B" type="description">
<paragraph id="87AFD4DD2E4F0E53BD3BCB2DEDB519AE">
<figureCitation id="AE96977414FB5DCF4054BDAC6252DEC8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus and copulatory organs of Mutatus yintiaoling sp. nov. A male holotype, dorsal view B male left palp, ventral view C same, dorsal view D same, prolateral view E same, retrolateral view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet." figureDoi="10.3897/zookeys.1226.137752.figure7" httpUri="https://binary.pensoft.net/fig/1243806">Chinese name: 阴条岭幻蛛 Figs 7</figureCitation>
,
<figureCitation id="F8E2BF9C04D8F08DDF03D31AB6A873BC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution map of Mutatus gen. nov." figureDoi="10.3897/zookeys.1226.137752.figure8" httpUri="https://binary.pensoft.net/fig/1243807">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2YBG" type="type material">
<paragraph id="C1BA05BB348DDCF1A8BA9A6AF7F3964F">
<heading id="CA9403AFF274B160B77D74DF6A7FE557" reason="title">Type material.</heading>
</paragraph>
<paragraph id="E6E0AC072E12DDDCDA35246C5D8B9D8B">
<materialsCitation id="34318CA01D8EDC1D0D1D054A418EB161" collectingDate="2022-05-23" collectorName="H. L. Zhu" country="China" county="Municipality" elevation="1752" latitude="31.495312" location="Zhuanping" longLatPrecision="1" longitude="109.92165" municipality="Yintiaoling National Nature Reserve" specimenCount="1" specimenCount-male="1" stateProvince="Chongqing" typeStatus="holotype">
<emphasis id="8CBAEE9712B5E432AA1FC3A4126CB563" bold="true" italics="true">
<typeStatus id="C2975B6C8E2C6274618565B40A2760C2" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="440E5861B7DC174B47971F6D7DB316A0" count="1" type="male"></specimenCount>
<collectingCountry id="EEBDED284DCC9E4BE3B2A963E308CA8B" name="China">China</collectingCountry>
<collectingRegion id="507A5E337AC8FE256F99B5889A4FB530" country="China" name="Chongqing">Chongqing</collectingRegion>
<collectingCounty id="5CB20E021D55BA7EA7D3331B6A63D577">Municipality</collectingCounty>
:
<collectingMunicipality id="141DFB982799349FE2B765B5526B0E59">Yintiaoling National Nature Reserve</collectingMunicipality>
,
<location id="A92FC3A6E8352510FC4D3E3A2920F0EB" LSID="urn:lsid:plazi:treatment:2643064135E357ACBD984644659ADEAB:A92FC3A6E8352510FC4D3E3A2920F0EB" country="China" county="Municipality" latitude="31.495312" longLatPrecision="1" longitude="109.92165" municipality="Yintiaoling National Nature Reserve" name="Zhuanping" stateProvince="Chongqing">Zhuanping</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJZBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[109.921656,31.495311]}">
<geoCoordinate id="7C55C36B7360FD4706A5DD354A395357" degrees="31" direction="north" minutes="29" orientation="latitude" precision="1" seconds="43.12" value="31.495312">31 ° 29 ' 43.12 &quot; N</geoCoordinate>
,
<geoCoordinate id="B71A791A337FCB974414493E846AC436" degrees="109" direction="east" minutes="55" orientation="longitude" precision="1" seconds="17.96" value="109.92165">109 ° 55 ' 17.96 &quot; E</geoCoordinate>
</named-content>
, elev.
<quantity id="3ECE57117789FF30D586C9676C68A386" metricMagnitude="3" metricUnit="m" metricValue="1.752" unit="m" value="1752.0">
<elevation id="80BDA230E7A89049463690637E11DFCB" metricMagnitude="3" metricUnit="m" metricValue="1.752" unit="m" value="1752.0">1752 m</elevation>
</quantity>
),
<date id="799ADFB39DCAA90EE57BD3F43A5A415D" value="2022-05-23">
<collectingDate id="1A8B43107C85B390263F22E0B8463E5C" value="2022-05-23">23 May 2022</collectingDate>
</date>
,
<collectorName id="23C74D2EEF3BFA50FD6B8FB68617730B">H. L. Zhu</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOZBG" type="etymology">
<paragraph id="3FAC3B0A5E33ED9F1259794EECFC39E7">
<heading id="D86967D5C0585F7FAA2A9E9614E58FC5" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="95590F6C84C689882898327032E7F8AA">
This specific name is derived from the
<typeStatus id="9292130BBFFB18CB0CCBFB5A7FA8A39F">type</typeStatus>
locality; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETZBG" type="diagnosis">
<paragraph id="03B7D796F09EAE9C95F0BF8E0FF68662">
<heading id="4B545585DAE7B60C50AEE0C870766272" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="E402E55EE510B3A067539D29B4B062DF">
This new species can be differentiated from
<taxonomicName id="FE75E3BEB78D7FC602D7157159445C9B" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gelao" status="sp. nov.">
<emphasis id="E136D510236F9EBA172EE255721C2E70" italics="true">M. gelao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="416C3B3688956EB79BFE83A8FBEC8FEA" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="51E7BDBCE3E405D68A7B0282609C36CE" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tujia" status="sp. nov.">
<emphasis id="1CDAE01D87C9FF18BFA6900452C5AF68" italics="true">M. tujia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3E9B3B2B8C74396EF2A126A3FAE1E6B0" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4115E01AC5323699DE5F6BAD88D19DE5" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tianyan" status="sp. nov.">
<emphasis id="5A81DAC042AE7E5E257127FE11E1092B" italics="true">M. tianyan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FBAAF027F7A8BE9967D0C8E7B46D51B" rank="species">sp. nov.</taxonomicNameLabel>
by the shorter prolateral tegular apophysis (vs long; compare Fig.
<figureCitation id="9CBE88EC34895636D93F487738D02F20" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus and copulatory organs of Mutatus yintiaoling sp. nov. A male holotype, dorsal view B male left palp, ventral view C same, dorsal view D same, prolateral view E same, retrolateral view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet." figureDoi="10.3897/zookeys.1226.137752.figure7" httpUri="https://binary.pensoft.net/fig/1243806">7 B</figureCitation>
with Figs
<figureCitation id="68EF820B95EB001CD8575F67C31F4EC1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 B</figureCitation>
,
<figureCitation id="D5A56CE1E62E46AE656F607B5332ACDC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 D</figureCitation>
,
<figureCitation id="06291BA2C0A36314BFA421BC23452058" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Mutatus tianyan sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure5" httpUri="https://binary.pensoft.net/fig/1243804">5 C</figureCitation>
) and the truncated and more curved tip of the long branch of retrolateral tibial apophysis (vs not truncated and curved tip; compare Fig.
<figureCitation id="B52B38952F8E8CD4905147ECEB71AB3A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus and copulatory organs of Mutatus yintiaoling sp. nov. A male holotype, dorsal view B male left palp, ventral view C same, dorsal view D same, prolateral view E same, retrolateral view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet." figureDoi="10.3897/zookeys.1226.137752.figure7" httpUri="https://binary.pensoft.net/fig/1243806">7 C, E</figureCitation>
with Figs
<figureCitation id="668E292BE3CF9FA269C93AF65945A3B7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 E</figureCitation>
,
<figureCitation id="95EE9A2D04B044D333ABDD1C8A39E7D0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 D</figureCitation>
,
<figureCitation id="E6FA4A652FEBC1727AD8A3D28D5EAAED" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Mutatus tianyan sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure5" httpUri="https://binary.pensoft.net/fig/1243804">5 B</figureCitation>
).
</paragraph>
<caption id="1627C24521FE5EB1EB0F5D27E6438F85" ID-DOI="10.3897/zookeys.1226.137752.figure7" ID-arpha="6BF760E2-649F-573E-8997-55F38A36A81B" httpUri="https://binary.pensoft.net/fig/1243806" startId="F7">
<paragraph id="6C77BA7F0ADC97A7445B905FCBFF498E">
<label id="B5053B2BDC21080E97F570441C0CDB84">Figure 7.</label>
</paragraph>
<paragraph id="BEFAA1C82C9CB1ED6D6414653AB05625">
Habitus and copulatory organs of
<taxonomicName id="20A612BF03EE23255D31D666C55F86C0" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="yintiaoling" status="sp. nov.">
<emphasis id="86430602E061D0372DD570350A725FC8" italics="true">Mutatus yintiaoling</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47C2362CEFB9933F62570D62146022D3" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="9BBC7923E53DBBA97CE0724E84CDCAA9" bold="true">A</emphasis>
male holotype, dorsal view
<emphasis id="B0EA87D5097047459E648438DE383BEB" bold="true">B</emphasis>
male left palp, ventral view
<emphasis id="BA126D123CBB77253E1BFB438CB0047E" bold="true">C</emphasis>
same, dorsal view
<emphasis id="8FB58ABA509FE7FC1B0BAB446A2CE0BF" bold="true">D</emphasis>
same, prolateral view
<emphasis id="8A90BB3A9BEA76A9D5657633D75B3ACC" bold="true">E</emphasis>
same, retrolateral view. Abbreviations: E — embolus;
<abbrev id="ABBRID0EX3BG" xlink_title="femur apophysis">FA</abbrev>
— femoral apophysis;
<abbrev id="ABBRID0E23BG" xlink_title="Prolateral tegular apophysis">PTA</abbrev>
— prolateral tegular apophysis;
<abbrev id="ABBRID0E63BG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
— retrolateral tibial apophysis;
<abbrev id="ABBRID0ED4BG" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
— retroventral tibial apophysis;
<abbrev id="ABBRID0EH4BG" xlink_title="sperm duct">SD</abbrev>
— sperm duct;
<abbrev id="ABBRID0EL4BG" xlink_title="tegular sheet">TS</abbrev>
— tegular sheet.
</paragraph>
</caption>
<caption id="53C288CD269BFD772F0172CB33A28F58" ID-DOI="10.3897/zookeys.1226.137752.figure8" ID-arpha="EABF924B-69AF-5D08-A240-A5F2675E9CA7" httpUri="https://binary.pensoft.net/fig/1243807" startId="F8">
<paragraph id="253EE79D6847220A27CA29A01300B03C">
<label id="8EA9080BD0AE1E93F4F92E9F2DE1F920">Figure 8.</label>
</paragraph>
<paragraph id="66AF8ADB4A41166A078483014FEA4CB1">
Distribution map of
<taxonomicName id="1C66D36CB5C8D55344CEA6DD0BD43E84" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D6102190BE9D7159B7D4AF5D1D90BE53" italics="true">Mutatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E596C97665A4CBF1BA8D79337ED3E33" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EI5BG" type="description">
<paragraph id="59912590290B0E996F9B1BC4ED8EE548">
<heading id="58ABEC1DB4568D5AF6F7006567162B32" reason="title">Description.</heading>
</paragraph>
<paragraph id="7B5B8F77496A06CAEDEADA5667EFF9F6">
Male (
<typeStatus id="A54B348C7A8181D33218295F0AFBC2FA" type="holotype">holotype</typeStatus>
, Fig.
<figureCitation id="E4BAE55A63678A03F9974897A0A888BC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus and copulatory organs of Mutatus yintiaoling sp. nov. A male holotype, dorsal view B male left palp, ventral view C same, dorsal view D same, prolateral view E same, retrolateral view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet." figureDoi="10.3897/zookeys.1226.137752.figure7" httpUri="https://binary.pensoft.net/fig/1243806">7 A</figureCitation>
). Total length 3.90, carapace 1.90 long, 1.63 wide; abdomen 2.00 long, 1.28 wide. Eye sizes and interdistances:
<abbrev id="ABBRID0ES5BG" xlink_title="anterior median eye">AME</abbrev>
0.10,
<abbrev id="ABBRID0EW5BG" xlink_title="anterior lateral eye">ALE</abbrev>
0.13,
<abbrev id="ABBRID0E15BG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0E55BG" xlink_title="posterior lateral eye">PLE</abbrev>
0.10;
<abbrev id="ABBRID0EC6BG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0EG6BG" xlink_title="anterior median eye">AME</abbrev>
0.04,
<abbrev id="ABBRID0EK6BG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0EO6BG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0ES6BG" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0EW6BG" xlink_title="anterior lateral eye">ALE</abbrev>
0.27,
<abbrev id="ABBRID0E16BG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E56BG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0EDAAI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EHAAI" xlink_title="posterior lateral eye">PLE</abbrev>
0.06,
<abbrev id="ABBRID0ELAAI" xlink_title="posterior lateral eye">PLE</abbrev>
<abbrev id="ABBRID0EPAAI" xlink_title="posterior lateral eye">PLE</abbrev>
0.42,
<abbrev id="ABBRID0ETAAI" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0EXAAI" xlink_title="posterior lateral eye">PLE</abbrev>
0.07.
<abbrev id="ABBRID0E2AAI" xlink_title="eye area width">EAW</abbrev>
0.58,
<abbrev id="ABBRID0E6AAI" xlink_title="cephalic region width">CRW</abbrev>
0.89,
<abbrev id="ABBRID0EDBAI" xlink_title="eye area width">EAW</abbrev>
/
<abbrev id="ABBRID0EHBAI" xlink_title="cephalic region width">CRW</abbrev>
0.65.
<abbrev id="ABBRID0ELBAI" xlink_title="cephalic region width">CRW</abbrev>
/
<abbrev id="ABBRID0EPBAI" xlink_title="carapace width">CW</abbrev>
0.54.
<abbrev id="ABBRID0ETBAI" xlink_title="median ocular area">MOA</abbrev>
0.26 long, anterior width 0.24, posterior width 0.30. Clypeal height 0.15,
<abbrev id="ABBRID0EXBAI" xlink_title="clypeal height">
<collectingCountry id="0E8089CA2E948087E55B9101B97FC2F4" name="Switzerland">CH</collectingCountry>
</abbrev>
/
<abbrev id="ABBRID0E2BAI" xlink_title="anterior median eye">AME</abbrev>
1.50. Labium 0.25 long, 0.27 wide. Sternum 0.96 long, 1.00 wide. Carapace oval, brown, cervical groove inconspicuous. Sternum brown, without pattern. Labium and endite as same color as sternum. Abdomen lighter-colored than carapace, smooth, with large dorsal scutum covering almost entire abdomen. Legs brown.
</paragraph>
<paragraph id="B6D5432BC50DF2ECF096B4A72B214DAA">
Palp (Fig.
<figureCitation id="94E6C8D3F29F00BED9623E9466689CD4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus and copulatory organs of Mutatus yintiaoling sp. nov. A male holotype, dorsal view B male left palp, ventral view C same, dorsal view D same, prolateral view E same, retrolateral view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet." figureDoi="10.3897/zookeys.1226.137752.figure7" httpUri="https://binary.pensoft.net/fig/1243806">7 B E</figureCitation>
). Femoral apophysis (
<abbrev id="ABBRID0EFCAI" xlink_title="femur apophysis">FA</abbrev>
) well developed, located at middle part. Patella longer than tibia, as half the length as femur. Tibia longer than wide, with 2 apophyses, retroventral apophysis (
<abbrev id="ABBRID0EJCAI" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
) tail-shaped, tip slightly curved, tapering towards tip; retrolateral apophysis (
<abbrev id="ABBRID0ENCAI" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
) strong, longer and wider than
<abbrev id="ABBRID0ERCAI" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
, abruptly narrowed at middle part in dorsal view, tip curved towards cymbium in dorsal view. Bulb pyriform, sperm duct (
<abbrev id="ABBRID0EVCAI" xlink_title="sperm duct">SD</abbrev>
) distinct, V-shaped. Prolateral tegular apophysis (
<abbrev id="ABBRID0EZCAI" xlink_title="Prolateral tegular apophysis">PTA</abbrev>
) short, originating at nearly 9 oclock position, tip truncated. Tegular sheet (
<abbrev id="ABBRID0E4CAI" xlink_title="tegular sheet">TS</abbrev>
) spade-shaped, anterior part large and wide, posterior part small, protuberance-shaped. Embolus (E) needle-like, slightly curved, tapering towards tip.
</paragraph>
<paragraph id="B8266643A7B2C49889C06190BBCA8BF1">Female: unknown.</paragraph>
<paragraph id="99591CF75DB9B4D5CD881C0B7F4F8878">Measurement of legs:</paragraph>
<paragraph id="E05CDCD0A2A99766527C5E5145CFC3B2">
<table id="TID0ENMBG" rules="all">
<tbody id="F99656D3CD3F9D5E7085375431947FE7">
<table-wrap id="T7" orientation="portrait" position="float">
<tr id="EF78A1810199C3A7EFE76A0AA1688B7C">
<th id="DF7A1C266E17D1EB9F67F87AF56A4C57" colspan="1" rowspan="1">male</th>
<th id="1009DD36F5EA299C9A26284FBE8968FE" colspan="1" rowspan="1">Fe</th>
<th id="619AE26FFA28B7D7378B055A5983068E" colspan="1" rowspan="1">Pa</th>
<th id="0FE9140D5540B4F19676B45915CE2001" colspan="1" rowspan="1">Ti</th>
<th id="082F9B21E780836A26E4F8732A38ABC4" colspan="1" rowspan="1">Me</th>
<th id="FB2F63CAB6D2A95AB6EC995767ED7557" colspan="1" rowspan="1">Ta</th>
<th id="F2445785AF639690C3ED9DD1BAD36DE5" colspan="1" rowspan="1">Total</th>
</tr>
<tr id="F581FBF5A43F552DECD5D636940405FA">
<td id="AA1B8D07ABB777E189EA8BF8AA4E287C" colspan="1" rowspan="1">Leg I</td>
<td id="A68022321C974937061A1C4BA6837410" colspan="1" rowspan="1">1.62</td>
<td id="CA51E3499EFF40301F360CA52778D43E" colspan="1" rowspan="1">0.62</td>
<td id="119A5AC315F1171E3EA183E2CFA021C7" colspan="1" rowspan="1">1.61</td>
<td id="F4BF184BE3F2AD5F155507A4F36A9870" colspan="1" rowspan="1">1.60</td>
<td id="4524B98CE9290D7781E43E169D49C38D" colspan="1" rowspan="1">0.74</td>
<td id="5AFA7544ABC57C2F79F6C994ECCBC46A" colspan="1" rowspan="1">6.19</td>
</tr>
<tr id="CF19C71F03E849F07270C43EB39353D7">
<td id="59C1AA6C756E9CB5B82646FA87B2FE41" colspan="1" rowspan="1">Leg II</td>
<td id="ACD69B69331297B504AA25AFD7679AF6" colspan="1" rowspan="1">1.41</td>
<td id="D8FFF54934CF0DA43B8E17CCFCD1B566" colspan="1" rowspan="1">0.61</td>
<td id="94C9861D26D09FBBD87CF383F0401BE9" colspan="1" rowspan="1">1.21</td>
<td id="E01E16E0A77D725CCB6A3EDE902A703B" colspan="1" rowspan="1">1.11</td>
<td id="9EE959697799EEC9D852F4A1AF76EAA0" colspan="1" rowspan="1">0.73</td>
<td id="EEC1A3A5115C07D65191F90968D12100" colspan="1" rowspan="1">5.07</td>
</tr>
<tr id="06ED5D5574C0E36E05001B33F29C960A">
<td id="9176FD57E667E04EFCE31D43AB71D000" colspan="1" rowspan="1">Leg III</td>
<td id="B1A68A097271184DE4796C3FF2DE53CB" colspan="1" rowspan="1">1.26</td>
<td id="FB0AFE665480ADD5320D83BC69292FD3" colspan="1" rowspan="1">0.52</td>
<td id="1099E7173402A3D9CB8A3DF7D399A9B9" colspan="1" rowspan="1">0.94</td>
<td id="05A2CF30C52CB501BC81C147D4B3BBF0" colspan="1" rowspan="1">1.19</td>
<td id="4B7C4D203CCEC991D61B9C59CFC3D2DE" colspan="1" rowspan="1">0.76</td>
<td id="2E77D1D49C5CC1D2472ADD8FD65E1873" colspan="1" rowspan="1">4.67</td>
</tr>
<tr id="DEBE0B3E113E87EE4E0981B41E79FB60">
<td id="6F917FEC249B259B21E1372BB168F6CA" colspan="1" rowspan="1">Leg IV</td>
<td id="C612BF1A130EA13B5F4B4B13725A2F72" colspan="1" rowspan="1">1.81</td>
<td id="D3B28002146E3825EA9DF554B6E51325" colspan="1" rowspan="1">0.67</td>
<td id="7DD4847044DC5C3021E677ECAE6DF7FF" colspan="1" rowspan="1">1.41</td>
<td id="8619417C99EC5F0B6B373F1E789B0891" colspan="1" rowspan="1">1.64</td>
<td id="E94DBA387B931FD6AF81ABB26781DE14" colspan="1" rowspan="1">0.86</td>
<td id="B16E8E8F6BDA32002122C42930B2CF1E" colspan="1" rowspan="1">6.39</td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
<paragraph id="54948B7C523155CC7777D1CCF88C3F38">Spination of legs:</paragraph>
<paragraph id="527F0DD9D539A46DE5F6BFEBBFFB3766">
<table id="TID0EMSBG" rules="all">
<tbody id="5AEA28C77EDC301896149372C033ABAE">
<table-wrap id="T8" orientation="portrait" position="float">
<tr id="F702BAEC21BDCB0EE97AF2E133DA25E5">
<th id="1E7F449C4CEF2565B03E3E6D21D05735" colspan="2" isEmpty="true" rowspan="1">-</th>
<th id="69BABA48286A4B07649B08804B19C6DA" colspan="1" rowspan="1">Fe</th>
<th id="3C88A1742F79C08A4274D38EEF1BAE05" colspan="1" rowspan="1">Pa</th>
<th id="69DE22F74349709EFB30B983B4A35D92" colspan="1" rowspan="1">Ti</th>
<th id="3069B5315E7CC2A3BBB3A62F3B16F5EB" colspan="1" rowspan="1">Me</th>
<th id="7C17F0AFE3DFA741D3290D29BBD8D3A5" colspan="1" rowspan="1">Ta</th>
</tr>
<tr id="511CE7FD6228A3C49EEEC338BCC0EA35">
<td id="B275D0FB0501A3FF9161638F5F9570A7" colspan="1" rowspan="4">male</td>
<td id="22965C073ECB0DAD5E5FAFE3B3AA7682" colspan="1" rowspan="1">Leg I</td>
<td id="45314A58FBAAAF910CBA485BDC5FBA4A" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EEIAI" xlink_title="prolateral">pl</abbrev>
3
</td>
<td id="86107FE1037BF9372D9F676649DAC6D3" colspan="1" rowspan="1"></td>
<td id="09FE51E1E05C52FFFD9456251074E724" colspan="1" rowspan="1">
<abbrev id="ABBRID0EPIAI" xlink_title="proventral">pv</abbrev>
6
<abbrev id="ABBRID0ETIAI" xlink_title="retroventral">rv</abbrev>
6
</td>
<td id="1B256B35EF6A264BD7405738A500ADC7" colspan="1" rowspan="1">
<abbrev id="ABBRID0E2IAI" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0E6IAI" xlink_title="retroventral">rv</abbrev>
3
</td>
<td id="6224D20D207486C3F49EED4FD3CB308D" colspan="1" rowspan="1"></td>
</tr>
<tr id="FDEC1FA102B1B5E03A52DBF75C0462F6">
<td id="5D00141C21437CBB7D030F7CA900F802" colspan="1" rowspan="1">Leg II</td>
<td id="8C37EE9889677D598AEED946286C7674" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EOJAI" xlink_title="prolateral">pl</abbrev>
2
</td>
<td id="95A3AA1286DBFFA1832BB02FEC051841" colspan="1" rowspan="1"></td>
<td id="41BD74539813EAE777984F4262C2A34D" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZJAI" xlink_title="proventral">pv</abbrev>
6
<abbrev id="ABBRID0E4JAI" xlink_title="retroventral">rv</abbrev>
5
</td>
<td id="7F6815080032EAABBAAF561256FBEC94" colspan="1" rowspan="1">
<abbrev id="ABBRID0EFKAI" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0EJKAI" xlink_title="retroventral">rv</abbrev>
3
</td>
<td id="574843F315BBB0983C32F480859411D2" colspan="1" rowspan="1"></td>
</tr>
<tr id="A2A83F0337C3F714A6CE93B272922307">
<td id="013059141C350C997AFE7546E1922110" colspan="1" rowspan="1">Leg III</td>
<td id="12658A48F32B8FF667709E8EA277FDEC" colspan="1" rowspan="1">d 1</td>
<td id="287D2DFC0901AC601713C507FCAD8F6F" colspan="1" rowspan="1"></td>
<td id="C1665C5EA869E7EE3873679449F2299D" colspan="1" rowspan="1"></td>
<td id="63014FD5012EF1D6CF54F58E58E87F34" colspan="1" rowspan="1"></td>
<td id="B9E085CA997495F17329DF4D820A320A" colspan="1" rowspan="1"></td>
</tr>
<tr id="D15E1BA39A4E0194404F4F6DDF7DBDAA">
<td id="C8BADAEA7F508B71986FD6A7A086B9E7" colspan="1" rowspan="1">Leg IV</td>
<td id="9C88D42B37B4EC136A9E2D1664490FC4" colspan="1" rowspan="1">d 1</td>
<td id="EE4C555FDA28ED4EB111339B8B1A27EB" colspan="1" rowspan="1"></td>
<td id="4060EB84BAF9D1A04BE0B2A085EC2DA5" colspan="1" rowspan="1"></td>
<td id="57195F174E2FAB734B31AA63B8D666D7" colspan="1" rowspan="1"></td>
<td id="4C84C097F869CE7E6A4AB966B5F92C36" colspan="1" rowspan="1"></td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWLAI" type="distribution">
<paragraph id="4821574497308784E48E1AA1B0F54769">
<heading id="91920B1B86DD156CDD2DF3DDDA761029" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="13C171282D084B45299CCC9A8A68DB19">
Known only from the
<typeStatus id="CA0E9E468A9F4CA4A75977222D373471">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,155 @@
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<subSubSection id="FE7D06B61E3699C18E28A7073C5CBCF3" type="nomenclature">
<paragraph id="D93487A281C6B0332F7434CDBBFCA7A8">
<treatmentCitationGroup id="2C01DDE65FEC0F69B2A7969AB4E23BC3">
<treatmentCitation id="CD67C1C6AFD647EF704C561FD04E1A66" author="Laboissière V" page="251" year="1936">
<taxonomicName id="F3D824B3FD03B720645E94857BFE2908" authority="Laboissiere, 1936: 251" authorityName="Laboissiere" authorityPageNumber="251" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="B3321F20A92CB1A2BDD50D3EA88381FE" italics="true">Paleosepharia</emphasis>
<bibRefCitation id="899028E9E9BD1880C34FEF7753733BC2" DOI="10.1080/21686351.1936.12278963" author="Laboissière V" journalOrPublisher="Annales de la Société Entomologique de France" pagination="239-262" refId="B19" refString="Laboissiere V (1936) Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et especes (5 e partie). Annales de la Societe Entomologique de France 105: 239-262. https://doi.org/10.1080/21686351.1936.12278963" title="Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et espèces (5 e partie)." volume="105" year="1936">Laboissière, 1936: 251</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWKAC" type="type species">
<paragraph id="0BB51E4EB89788200FA84F219E38FE72">
<heading id="BAA829FD5CA8BB067608251BDCEEEB9E" reason="title">
<typeStatus id="3D99F5C9F39C86D231FD9ABD78B22976">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="618FBCA372CD1361928CA0EE7247AD7A">
<taxonomicName id="4F5EAF5724A2399382744B298D3429BE" authority="Laboissiere, 1936" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="902702B4D7D9EA4A453C769811400A3C" italics="true">Paleosepharia truncata</emphasis>
<bibRefCitation id="C8DA88E36F0A75D552AADCA9A69A6732" DOI="10.1080/21686351.1936.12278963" author="Laboissière V" journalOrPublisher="Annales de la Société Entomologique de France" pagination="239-262" refId="B19" refString="Laboissiere V (1936) Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et especes (5 e partie). Annales de la Societe Entomologique de France 105: 239-262. https://doi.org/10.1080/21686351.1936.12278963" title="Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et espèces (5 e partie)." volume="105" year="1936">Laboissière, 1936</bibRefCitation>
</taxonomicName>
, by monotypy and original designation. Redescription in
<bibRefCitation id="3E85DCE41A1A8915D64109DF1FB908E8" DOI="10.1649/0010-065X-70.2.395" author="Rizki" etAl="et al." firstAuthor="Rizki" journalOrPublisher="Coleopterists Bulletin" pagination="395-398" refId="B31" refString="Rizki A, Hazmi IR, Wagner T, Idris AB (2016) Redescription of Paleosepharia truncata Laboissière, 1936, type species of the genus Paleosepharia Laboissière, 1936 (Coleoptera: Chrysomelidae: Galerucinae). Coleopterists Bulletin 70: 395398. https://doi.org/10.1649/0010-065X-70.2.395" title="Redescription of Paleosepharia truncata Laboissière, 1936, type species of the genus Paleosepharia Laboissière, 1936 (Coleoptera: Chrysomelidae: Galerucinae)." volume="70" year="2016">Rizki et al. 2016</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELLAC" type="distribution">
<paragraph id="B3B96E51F74F5BEB6A96EFD4128A9FC7">
<heading id="2C2970B444E76F7C237A57801948142E" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="879064357C28D03FC11357C863CD44FA">Oriental and Palaearctic regions.</paragraph>
</subSubSection>
<subSubSection id="SECID0EQLAC" type="diagnosis">
<paragraph id="30780B17B13E56FE4E8C645E548DC660">
<heading id="508A415D74596AABE7C7A63DF46EF379" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="842F4AA44330F4F2F59B05A40FF10C16">
This genus is similar to
<taxonomicName id="DCC0286E6B7F8E263BAE524088D584DF" authority="Chevrolat, 1836" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Atrachya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="2339A1A86324BA2A3B5C46375E9248D3" italics="true">Atrachya</emphasis>
Chevrolat, 1836
</taxonomicName>
and
<taxonomicName id="3D0A9071F64336C77CF70181C403466D" authority="Chevrolat, 1836" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="0B31F5AB66DF03DB7967258444FADC28" italics="true">Monolepta</emphasis>
Chevrolat, 1836
</taxonomicName>
. In
<taxonomicName id="FB012796EA8C1FC88D60448839BDF7A7" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="D3B2F8E2B82214E0B7EB7A4E4A99C1D0" italics="true">Paleosepharia</emphasis>
</taxonomicName>
, the third antennomere is longer than the second (both antennomeres are more or less equal in
<taxonomicName id="21722868CC30D6E357046A8CE4A1AC31" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="38FD58A3EA2D5D0AAC2E02C71451233D" italics="true">Monolepta</emphasis>
</taxonomicName>
), and the epipleuron continues towards the apex (suddenly narrowed before the middle in
<taxonomicName id="373D1FA07CCE4F473B1025D9D9A028DF" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="704E8C3CA6C998CB1B6AD9F72DBDE28D" italics="true">Monolepta</emphasis>
</taxonomicName>
), procoxal cavity is closed behind (opened in
<taxonomicName id="6D0273C9D165AA57E341F3F5D5433A71" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Atrachya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="E795C18519A6E53D09660B18E0794163" italics="true">Atrachya</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8E972ACF29FF2002EEF878BEDC6D5C55">
Male genitalia. In
<taxonomicName id="8FAD764193542D80933D9197C5834CE9" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="D9097511BD63CD77E5CFB358817A8CE5" italics="true">Paleosepharia</emphasis>
</taxonomicName>
and
<taxonomicName id="A8DA608DA2FE33BF4A509F92A658C79B" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Atrachya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="6224B3158555A7B132BA8A20C903349B" italics="true">Atrachya</emphasis>
</taxonomicName>
the aedeagus is parallel-sided in the basal two-thirds, apical third strongly narrowed (aedeagus slender, parallel-sided in
<taxonomicName id="BC6ACC821927B15C7AA38C7FC68174E2" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="44ADC49E37EEDC90A2D592D7E83D31E0" italics="true">Monolepta</emphasis>
</taxonomicName>
). In
<taxonomicName id="E39843D4EB17B414CE93318FE0739A9E" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="EA252D84D1FDAA40401423DCA6447656" italics="true">Paleosepharia</emphasis>
</taxonomicName>
, tectum long, rounded or truncate at apex (rounded at apex in
<taxonomicName id="8E97C140020A1867B7504FD891365632" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="2C19CBF88FBDB4F5DBBF591876FAD2D5" italics="true">Monolepta</emphasis>
</taxonomicName>
; incised at apex, the two apical tips forming strong hooks in
<taxonomicName id="D8D745C49B2EC2DCBDD257AB6185681B" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Atrachya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="6762AF78219A334D0CC979CDFF9BBF8F" italics="true">Atrachya</emphasis>
</taxonomicName>
).
<taxonomicName id="31994A525C5D925EF637173A5E3CAEA1" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="AD28941C9610D58FBDB4C2AEB24FA902" italics="true">Paleosepharia</emphasis>
</taxonomicName>
usually has two pairs of strongly sclerotized spiculae (lateral and median), while the ventral spiculae are weakly sclerotized. The median spiculae consist of long, slender spikes.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,690 @@
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<mods:title id="3B8F2CA4DDB07DFA84866A822EDA035F">A review of the intertidal Medon Stephens (Coleoptera, Staphylinidae, Paederinae) with description of a new species on the East Asian coasts</mods:title>
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<mods:namePart id="7D8E5FE5C560A921CCA29FC15B2E5459">Ahn, Kee-Jeong</mods:namePart>
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</mods:role>
<mods:namePart id="3BA5A13E9E14861A2DC8AF1F3B62C61F">Jeon, Gae-Nam</mods:namePart>
<mods:affiliation id="EB5B3FCB66C4186A67CDD0084621C144">Department of Biology, Chungnam National University, Daejeon 34134, Republic of Korea</mods:affiliation>
</mods:name>
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<subSubSection id="8023E27E6D60CEDB74ACDA94AAD89A7A" type="nomenclature">
<paragraph id="480774D1FBDA78688229B7F4761455AC">
<taxonomicName id="5AD870AA98064CD4887EEAE043CDD821" authorityName="Ahn &amp; Jeon" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="calculosus" status="sp. nov.">
<emphasis id="06262AE95635C324892F215B1C7C42EA" italics="true">Medon calculosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2462B1921554CA5E990DF8D38F921FD" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C9A70577FC65C39F23A8A0D377DC0D5A" type="description">
<paragraph id="34792958EDF1EE78778F8362FA690F4B">
<figureCitation id="D96FC18B77E1576DC80CBAE570856800" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male habitus A Medon calculosus sp. nov. 3.1 mm B paler form of M. calculosus sp. nov. C M. tomokoae, 3.2 mm." figureDoi="10.3897/zookeys.1226.137489.figure1" httpUri="https://binary.pensoft.net/fig/1243890">Figs 1 A, B</figureCitation>
,
<figureCitation id="6BA60FA4D00E96C4F5016E093E9926E5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Medon calculosus sp. nov., male A labrum, dorsal aspect B mandible, dorsal aspect (left and right) C maxilla, dorsal aspect D labium, ventral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure2" httpUri="https://binary.pensoft.net/fig/1243891">2</figureCitation>
,
<figureCitation id="BFE8F4A142266CEF436AC57DBF489DDC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVDAE" type="Type specimens">
<paragraph id="DEFCE81C27A194A59D3D96C9FB9D680E">
<heading id="8ADB4276C4137CAD2599B7D96690E057" reason="title">Type specimens.</heading>
</paragraph>
<paragraph id="189C45F2188273DF7C30533A6FB6344A">
<materialsCitation id="0E9F15B21488B1D6EC2AD292191C0D97" collectingDate="2023-10-31" collectionCode="CNUIC" country="South Korea" latitude="35.855904" location="Gyeongbuk Prov." longLatPrecision="1" longitude="129.52467" specimenCount="2" stateProvince="Gyeongbuk" typeStatus="holotype">
<emphasis id="1C51A4B1083962E8CBB3956682EDB88B" bold="true" italics="true">
<typeStatus id="2E1094A9B54FA9A17BD09F621F01B2C3" type="holotype">Holotype</typeStatus>
</emphasis>
• labeled as follows: “
<collectingCountry id="D587E148EE2EEC9600A50DD312765AA6" name="South Korea">Korea</collectingCountry>
:
<collectingRegion id="A78871610F4B5E4AA90428EBD09E430F" country="South Korea" name="Gyeongbuk">Gyeongbuk Prov.</collectingRegion>
, Pohang-si, Janggi-myeon, Gaewon-ri 445-12
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<geoCoordinate id="6337BA52AC7EEB4265A4BF891FC46F1E" degrees="35.855905" direction="north" orientation="latitude" precision="1" value="35.855904">35.855905 ° N</geoCoordinate>
,
<geoCoordinate id="F36F1969A3BEFFA0450814169D32A47F" degrees="129.524668" direction="east" orientation="longitude" precision="1" value="129.52467">129.524668 ° E</geoCoordinate>
</named-content>
,
<date id="43B390A1433086BD305004975144785A" value="2023-10-31">
<collectingDate id="509110B11C86A587C88EDE37CA736EB1" value="2023-10-31">31 X 2023</collectingDate>
</date>
, KJ Ahn, pebble beach in low-tide zone;
<typeStatus id="5AB95AEDECCA8215691CFA9736CCE739" type="holotype">Holotype</typeStatus>
,
<taxonomicName id="79F6E0947C4695DC05484AA3C95516E3" authority="Ahn and Jeon, Desig. K. - J. Ahn 2024" authorityName="Ahn and Jeon, Desig. K. - J. Ahn" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="calculosus">
<emphasis id="9DC1D7B2E0388B6054D28D0061C2F4CA" italics="true">Medon calculosus</emphasis>
Ahn and Jeon, Desig. K. - J. Ahn 2024
</taxonomicName>
, deposited in
<abbrev content-type="institution" id="ABBRID0ETEAE" xlink_title="Chungnam National University Insect Collection">
<collectionCode id="ADD5450211A9ACFFE6F19E3647998F8C">CNUIC</collectionCode>
</abbrev>
,
<collectingRegion id="C781DC5CFEC4CD5C1A7A886952930A96" country="South Korea" name="Daejon">Daejeon</collectingRegion>
,
<collectingCountry id="C1E170F278D8481FFF7D64B5A1758ED9" name="South Korea">Korea</collectingCountry>
</materialsCitation>
.
<materialsCitation id="E8BB28D9CEA883D33F2D53628C6CD164" collectingDate="2023-10-31" collectionCode="CNUIC" country="South Korea" latitude="35.855904" location="Gyeongbuk Prov." longLatPrecision="1" longitude="129.52467" specimenCount="1" stateProvince="Gyeongbuk" typeStatus="paratype">
<emphasis id="8C6166CABA26AEC2019D9295AA55F1B5" bold="true" italics="true">
<typeStatus id="61765BEF855044C4F555E1F598940991" type="paratype">Paratypes</typeStatus>
</emphasis>
• 3 exx., same data as holotype
</materialsCitation>
<materialsCitation id="DF9E83888A467223CC740E42CAABA969" collectingDate="2023-11-02" collectorName="KJ Ahn" country="South Korea" county="Pohang-si" latitude="35.855904" location="Gaewon-ri" longLatPrecision="1" longitude="129.52467" municipality="Janggi-myeon" specimenCount="1" stateProvince="Gyeongbuk" typeStatus="paratype">
2 exx.,
<collectingCountry id="717768163F23DB37EE4C71B02523E112" name="South Korea">Korea</collectingCountry>
:
<collectingRegion id="E62DF248C9EF87D031A58CD9C294F38F" country="South Korea" name="Gyeongbuk">Gyeongbuk Prov.</collectingRegion>
,
<collectingCounty id="64E0F1E42C1C7206C3C1A4C7C029A718">Pohang-si</collectingCounty>
,
<collectingMunicipality id="49C138F3EF6030D93BE72CA8AC3ECCE1">Janggi-myeon</collectingMunicipality>
,
<location id="3D787C330DEBCE0AD9490E6FBE3B44C6" LSID="urn:lsid:plazi:treatment:41D1757AD8D954BB968916BB9BC005BA:3D787C330DEBCE0AD9490E6FBE3B44C6" country="South Korea" county="Pohang-si" latitude="35.855904" longLatPrecision="1" longitude="129.52467" municipality="Janggi-myeon" name="Gaewon-ri" stateProvince="Gyeongbuk">Gaewon-ri</location>
445-12
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<geoCoordinate id="E41BC6B7DC2FBBFCAF3718B0142B1E58" degrees="35.855905" direction="north" orientation="latitude" precision="1" value="35.855904">35.855905 ° N</geoCoordinate>
,
<geoCoordinate id="282831783DEECD51C477460A14491684" degrees="129.524668" direction="east" orientation="longitude" precision="1" value="129.52467">129.524668 ° E</geoCoordinate>
</named-content>
,
<date id="192034AE5B257CCF4903B9275B657D16" value="2023-11-02">
<collectingDate id="8A9CB70F04697969A869DF17D73478E7" value="2023-11-02">2 XI 2023</collectingDate>
</date>
,
<collectorName id="9A390F61D6F0DF5FE48630E0923AD5D9">KJ Ahn</collectorName>
, pebble beach in mid-tide zone
</materialsCitation>
<materialsCitation id="55535BAAAE2AFE5F50F7C43172EF6365" collectingDate="2023-03-24" collectorName="KJ Ahn" country="South Korea" county="Pohang-si" latitude="35.855904" location="Gaewon-ri" longLatPrecision="1" longitude="129.52467" municipality="Janggi-myeon" specimenCount="1" stateProvince="Gyeongbuk" typeStatus="paratype">
2 exx., same locality as above,
<date id="83C1A366917944FA4261112D2742DFEC" value="2023-03-24">
<collectingDate id="B61A32D69FFC6A3859B77E38ADC3FC45" value="2023-03-24">24 III 2023</collectingDate>
</date>
,
<collectorName id="02E26EA5FF83C0C621BCE997B9220227">KJ Ahn</collectorName>
, pebble beach in low-tide zone
</materialsCitation>
<materialsCitation id="0F1BF6CF2C489924AA2FA79175B9053D" collectingDate="2023-11-02" collectorName="KJ Ahn" country="South Korea" county="Gyeongju-si" latitude="35.652813" location="Suryeom-ri" longLatPrecision="1" longitude="129.45004" municipality="Yangnam-myeon" specimenCount="1" stateProvince="Gyeongbuk" typeStatus="paratype">
2 exx.,
<collectingCounty id="BAEFC7F04D36156A4107DE7E6B00F694">Gyeongju-si</collectingCounty>
,
<collectingMunicipality id="B80F7F00ED73687AE8BA4E251D9384CE">Yangnam-myeon</collectingMunicipality>
,
<location id="4F2CCC3F9EB978AC44534CB1BABA6443" LSID="urn:lsid:plazi:treatment:41D1757AD8D954BB968916BB9BC005BA:4F2CCC3F9EB978AC44534CB1BABA6443" country="South Korea" county="Gyeongju-si" latitude="35.652813" longLatPrecision="1" longitude="129.45004" municipality="Yangnam-myeon" name="Suryeom-ri" stateProvince="Gyeongbuk">Suryeom-ri</location>
,
<location id="41C30FF3EFDFF60147B901769147F817" LSID="urn:lsid:plazi:treatment:41D1757AD8D954BB968916BB9BC005BA:41C30FF3EFDFF60147B901769147F817" country="South Korea" county="Gyeongju-si" latitude="35.652813" longLatPrecision="1" longitude="129.45004" municipality="Yangnam-myeon" name="Jigyeong-beach" stateProvince="Gyeongbuk">Jigyeong-beach</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGFAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[129.450041,35.652814]}">
<geoCoordinate id="DD4482AC2775F65DBE694F4529B88E21" degrees="35.652814" direction="north" orientation="latitude" precision="1" value="35.652813">35.652814 ° N</geoCoordinate>
,
<geoCoordinate id="CC398BAD3A362D996CB1C67E87E44D94" degrees="129.450041" direction="east" orientation="longitude" precision="1" value="129.45004">129.450041 ° E</geoCoordinate>
</named-content>
,
<date id="DE2EA53F184F25FC697F9DE16BB585B8" value="2023-11-02">
<collectingDate id="A96ED383F09474EEDE94CDE6B1BCABD7" value="2023-11-02">2 XI 2023</collectingDate>
</date>
,
<collectorName id="23C95E4BD8072B6725F1FAB45C7E35C8">KJ Ahn</collectorName>
, pebble beach in low-tide zone
</materialsCitation>
<materialsCitation id="DDD4EFC44C1CBF4D645D1BD0998E2F8D" collectingDate="2018-07-13" collectorName="IS Yoo &amp; JS Lee &amp; Jung" country="South Korea" elevation="7" latitude="35.699253" location="Naa-beach" longLatPrecision="1" longitude="129.47409" municipality="Naa-ri" specimenCount="1" stateProvince="Gyeongbuk" typeStatus="paratype">
2 exx.,
<collectingMunicipality id="4C419701532F4E9AB01DB6FC05E5B672">Naa-ri</collectingMunicipality>
,
<location id="ECF8BC62D534DA3BBE5E71D038D29EB0" LSID="urn:lsid:plazi:treatment:41D1757AD8D954BB968916BB9BC005BA:ECF8BC62D534DA3BBE5E71D038D29EB0" country="South Korea" latitude="35.699253" longLatPrecision="1" longitude="129.47409" municipality="Naa-ri" name="Naa-beach" stateProvince="Gyeongbuk">Naa-beach</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOFAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[129.474092,35.699253]}">
<geoCoordinate id="FA3D8EDD77A110A2834651D2F5F480CC" degrees="35" direction="north" minutes="41" orientation="latitude" precision="1" seconds="57.31" value="35.699253">35 ° 41 57.31 ″ N</geoCoordinate>
,
<geoCoordinate id="C35D12323223D6B3F5F777E2BD56E31D" degrees="129" direction="east" minutes="28" orientation="longitude" precision="1" seconds="26.73" value="129.47409">129 ° 28 26.73 ″ E</geoCoordinate>
</named-content>
,
<quantity id="6952FA3923855DE1D8352D199370CE44" metricMagnitude="0" metricUnit="m" metricValue="7.0" unit="m" value="7.0">
<elevation id="35B91DE0C05946DB6DAAAFD90037F9E9" metricMagnitude="0" metricUnit="m" metricValue="7.0" unit="m" value="7.0">7 m</elevation>
</quantity>
,
<date id="17DA2D4F408627FCA4623DC55BCE59EA" value="2018-07-13">
<collectingDate id="86173B530EEE098718573E0868FB7236" value="2018-07-13">13 VII 2018</collectingDate>
</date>
,
<collectorName id="8D27643F4FF786897385AF184ABD17BD">IS Yoo</collectorName>
,
<collectorName id="3E52B9AE0AE0C184538F558B12D4D012">JS Lee</collectorName>
, JG
<collectorName id="F5F5DDFD89A5D9B04BA785B85B6EF067">Jung</collectorName>
, in gravels covered with seaweed in high-supratidal zone, flotation
</materialsCitation>
<materialsCitation id="499BE89364D9AC9557BB63261209E374" collectingDate="2022-09-24" collectorName="KJ Ahn" country="South Korea" county="Yeongduk-gun" latitude="36.60258" location="Byonggok-ri" longLatPrecision="1" longitude="129.4157" municipality="Byonggok-myeon" specimenCount="1" stateProvince="Gyeongbuk" typeStatus="paratype">
2 exx.,
<collectingCounty id="B6DE0EAAC12810ABD34883E113621003">Yeongduk-gun</collectingCounty>
,
<collectingMunicipality id="DE60C6238A59DD802811762E0CB654C7">Byonggok-myeon</collectingMunicipality>
,
<location id="EF3EED068A88356E5514CAAD986B025E" LSID="urn:lsid:plazi:treatment:41D1757AD8D954BB968916BB9BC005BA:EF3EED068A88356E5514CAAD986B025E" country="South Korea" county="Yeongduk-gun" latitude="36.60258" longLatPrecision="1" longitude="129.4157" municipality="Byonggok-myeon" name="Byonggok-ri" stateProvince="Gyeongbuk">Byonggok-ri</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWFAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[129.415695,36.602581]}">
<geoCoordinate id="0C5976EC8BA9C93F26C08CB5C0437A1A" degrees="36.602581" direction="north" orientation="latitude" precision="1" value="36.60258">36.602581 ° N</geoCoordinate>
,
<geoCoordinate id="575BEF59C0A7E681C64A27E1E1DA9374" degrees="129.415695" direction="east" orientation="longitude" precision="1" value="129.4157">129.415695 ° E</geoCoordinate>
</named-content>
,
<date id="1FDE4F86593B3334B3D86E46E4796B8C" value="2022-09-24">
<collectingDate id="18104FD0905714096AB3D6220842721D" value="2022-09-24">24 IX 2022</collectingDate>
</date>
,
<collectorName id="20D4BA016E443F5A864D390B66216416">KJ Ahn</collectorName>
, flotation on pebble beach
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2FAE" type="description">
<paragraph id="A65F48E9344C512FF7902B87EBD12948">
<heading id="A8257897DF1868EC031D40464827F4C9" reason="title">Description.</heading>
</paragraph>
<paragraph id="CBA1074B463068306AA2C3E442F40C32">
<emphasis id="6F09BF707EC710890295BCCD08502937" bold="true">Male.</emphasis>
Body length 3.0
<quantity id="A575EE634F8CCE0D46756120078A1FE7" metricMagnitude="-3" metricUnit="m" metricValue="3.3" unit="mm" value="3.3">3.3 mm</quantity>
. Body slender, more or less parallel-sided, flattened, and densely pubescent. Head, pronotum, elytra, antennae and legs brown to reddish-brown; abdomen black (Fig.
<figureCitation id="5AB7148DCCF2D74E94519BF550366A32" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male habitus A Medon calculosus sp. nov. 3.1 mm B paler form of M. calculosus sp. nov. C M. tomokoae, 3.2 mm." figureDoi="10.3897/zookeys.1226.137489.figure1" httpUri="https://binary.pensoft.net/fig/1243890">1</figureCitation>
).
<emphasis id="F2ECE4C883EC6C8F04B2C42A01D5892B" bold="true" italics="true">Head.</emphasis>
Subquadrate, about 1.03 times as long as wide, widest basal 1 / 5 and about 1.15 times wider than pronotum, dorsal surface covered with dense pubescence; eye small, about 0.26 times as long as temple; antenna very long, extending to middle of elytron; inserted under side of front head, insertion invisible from dorsal view; all antennomeres elongate, scape longest and widest, pedicel to antennomere 7 distinctly longer than wide, antennomeres 810 subglobular, antennomere 11 water drop-shaped, relative length of 11 antennomeres 21: 14: 14: 13: 14: 13: 13: 11: 11: 11: 14; gular sutures narrowly separated and converged posteriorly. Neck moderately wide, about 1 / 3 as wide as head.
</paragraph>
<caption id="5EC946D362202786792ED2606A558E2A" ID-DOI="10.3897/zookeys.1226.137489.figure1" ID-arpha="FAA25FCC-0212-53DB-BDEA-AAC6380E5349" httpUri="https://binary.pensoft.net/fig/1243890" startId="F1">
<paragraph id="24B6CBFF028E862F31724910B0C5B10A">
<label id="5E9A9F159D48E74BE5EFACBBF9BFF53A">Figure 1.</label>
</paragraph>
<paragraph id="74CB873732BBCE66CC1AA5A62437CE3B">
Male habitus
<emphasis id="0E1A842F1960E4179A4B9FB1F77B2584" bold="true">A</emphasis>
<taxonomicName id="5E467D08AD7C3E2A72C1B00C1713C083" authorityName="Ahn &amp; Jeon" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="calculosus" status="sp. nov.">
<emphasis id="1C7478BB13DF8CA773395F2D3879D594" italics="true">Medon calculosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7ACE58194A164E681BD74D0D8E0EAD6" rank="species">sp. nov.</taxonomicNameLabel>
3.1 mm
<emphasis id="31D619AA15EF4C15CA1F2507C9902C29" bold="true">B</emphasis>
paler form of
<taxonomicName id="5A35030776BD80C48FADBEC52C06CF43" authorityName="Ahn &amp; Jeon" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="calculosus" status="sp. nov.">
<emphasis id="4916937A9523BD1D22EBA59BE15D4C02" italics="true">M. calculosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0B1782C87D41213CCB98502F55FE6438" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="4E0415988CD1129395FC879DB7BEA401" bold="true">C</emphasis>
<taxonomicName id="E6C121855B3D0F9D357427354F4D3C5F" authorityName="Shibata &amp; Fujimoto" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tomokoae">
<emphasis id="EDAB5CD4188152F97BA89781BF41BF9B" italics="true">M. tomokoae</emphasis>
</taxonomicName>
, 3.2 mm.
</paragraph>
</caption>
<paragraph id="D1DDCC228D738B6FC93EBCCFFE539274">
<emphasis id="3129F75F7F050DC159E99084B00805BC" bold="true" italics="true">Mouthparts.</emphasis>
Labrum (Fig.
<figureCitation id="E2594349F0D0C5802EECDFF94A7155A9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Medon calculosus sp. nov., male A labrum, dorsal aspect B mandible, dorsal aspect (left and right) C maxilla, dorsal aspect D labium, ventral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure2" httpUri="https://binary.pensoft.net/fig/1243891">2 A</figureCitation>
) broad, anterior margin with two pairs of tooth-like processes, medial one longer and slender; mandibles (Fig.
<figureCitation id="D7F710EC5C0E56BC633B418F6E2BA2A2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Medon calculosus sp. nov., male A labrum, dorsal aspect B mandible, dorsal aspect (left and right) C maxilla, dorsal aspect D labium, ventral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure2" httpUri="https://binary.pensoft.net/fig/1243891">2 B</figureCitation>
) asymmetrical, with 34 internal teeth; maxilla (Fig.
<figureCitation id="20B82278D0702C8411EB82D982CF82C3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Medon calculosus sp. nov., male A labrum, dorsal aspect B mandible, dorsal aspect (left and right) C maxilla, dorsal aspect D labium, ventral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure2" httpUri="https://binary.pensoft.net/fig/1243891">2 C</figureCitation>
) with galea and lacinia fringed with long setae; maxillary palpomere 1 small, about 3.18 times as long as 2; palpomere 2 expanded apically, little curved inwardly, about 0.97 times as long as 3; palpomere 3 droplet-shaped, widest apical third; palpomere 4 minute, about 0.88 times as long as 1; labium (Fig.
<figureCitation id="123E523C62B4BA3580B34120189B4F71" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Medon calculosus sp. nov., male A labrum, dorsal aspect B mandible, dorsal aspect (left and right) C maxilla, dorsal aspect D labium, ventral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure2" httpUri="https://binary.pensoft.net/fig/1243891">2 D</figureCitation>
) with palpomere 1 small, about 0.46 times as long as 2; palpomere 2 widest near middle, about 1.87 times as long as 3; palpomere 3 small and thin, about 1.13 times as long as 1.
<emphasis id="E21BC0521CB7584170C373D4F7FACBF2" bold="true" italics="true">Thorax.</emphasis>
Pronotum more or less rectangular, about 1.17 times as long as wide; narrower than head, almost as long as head, widest apical 1 / 5 and narrowed posteriorly, yellow short setae densely present, impunctate central region present; prosternum well developed, median area upheaved with transverse carina, prosternal process acute and very long, hypomeral projection well developed and triangular. Elytra about 1.09 times as wide as pronotum, almost as long as pronotum, elytron 2.37 times as long as wide, distinct and shallow punctures present, covered with yellow setae, lateral margin straight and parallel. Metendosternite Y-shaped. Hind wings short but longer than elytron. Tarsal formula 5-5 - 5, front, middle and hind tarsomeres 15 each strongly widened.
<emphasis id="8C1DEB01801D10669CB02CD9643048D4" bold="true" italics="true">Abdomen.</emphasis>
Almost parallel-sided and broadened posteriorly after segment
<collectionCode id="8F965104AE0C9A9624E77545C9E5A577" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
. Tergites III
<collectionCode id="2434CD4558BD5AE9F597A3B41C093D80" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
with shallow and transverse basal depression. Sternite III with basal transverse carina, medially pointed; apical margin of tergite VII subtruncate (Fig.
<figureCitation id="C3AB390B7CDA3CA4C4034256C2EFA99C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 A</figureCitation>
) and sternite VIII slightly emarginate (Fig.
<figureCitation id="A8ABC4233D89F6F635F253D66CEADE7F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 B</figureCitation>
); apical margin of sternite IX (Fig.
<figureCitation id="5CA5BBB75BE71E01426615F15B62730C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 C</figureCitation>
) slightly emarginate; tergites IX and X (Fig.
<figureCitation id="1E457841357F3E04D881B9CF5FDD2BAB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 D</figureCitation>
) covered with scattered setae, apical margin with numerous long and brown setae.
<emphasis id="7BD866445421D5A77C5FD896A3C52632" bold="true" italics="true">Aedeagus.</emphasis>
Median lobe symmetrical, comprising about half sclerotized part and membraneous part; more or less triangular lobe protruded in dorsal view (Fig.
<figureCitation id="18D63245061215130F96B39AB290056F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 E</figureCitation>
); apical process narrow and well sclerotized in lateral view (Fig.
<figureCitation id="932E568E5075F31484B98E1ACB66D809" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 F</figureCitation>
).
</paragraph>
<caption id="81104C18CECFFCB1ECDAA9A498484AE1" ID-DOI="10.3897/zookeys.1226.137489.figure2" ID-arpha="DBAC3D24-C14E-534B-A1F0-7E8883D11361" httpUri="https://binary.pensoft.net/fig/1243891" startId="F2">
<paragraph id="056801427E652DFDE568311B1F1D004B">
<label id="B99040DDC90CC4DCEAAF0DBF94F55336">Figure 2.</label>
</paragraph>
<paragraph id="DF6CE2ABD689E8438E2A06A5F770F05F">
<taxonomicName id="1B5B41229D17EBD9D1AE6700220F3695" authorityName="Ahn &amp; Jeon" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="calculosus" status="sp. nov.">
<emphasis id="D8FCC82018FC829A2158E8BE0E7EA663" italics="true">Medon calculosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="075237BF2FF133BA87F8690635AD3E0B" rank="species">sp. nov.</taxonomicNameLabel>
, male
<emphasis id="0992804006F75F72ECB1BF157A3AA807" bold="true">A</emphasis>
labrum, dorsal aspect
<emphasis id="193FEBD53F093F51D1789B1B2DA7E65E" bold="true">B</emphasis>
mandible, dorsal aspect (left and right)
<emphasis id="301824EE7C245A91D9010789915B6A4C" bold="true">C</emphasis>
maxilla, dorsal aspect
<emphasis id="A81691A626F01B3531D7C4AC54494D77" bold="true">D</emphasis>
labium, ventral aspect. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="54550FFC5C858D9B66EEF8EF8862BC86" ID-DOI="10.3897/zookeys.1226.137489.figure3" ID-arpha="9A1E959A-FBD7-559C-9927-985DF5A59CDD" httpUri="https://binary.pensoft.net/fig/1243892" startId="F3">
<paragraph id="61DD0A51E96CB2BCF1A2AA624B472131">
<label id="22390214AA0D1A16B595352DCD707AC8">Figure 3.</label>
</paragraph>
<paragraph id="EFA7587C7B9F19F8044431557C26C225">
<taxonomicName id="7C92276C523C353EF0EDB38BC417D952" authorityName="Ahn &amp; Jeon" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="calculosus" status="sp. nov.">
<emphasis id="616F5FC30922E18165289FE9321D465D" italics="true">Medon calculosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4247791758721669D5527206FCD67180" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="117BC0CCCBB4C2E2CB7BE223951EC938" bold="true">A</emphasis>
male sternite VII, ventral aspect
<emphasis id="CEB3A66BCB3A5A5DFF30CD72E880F27C" bold="true">B</emphasis>
male sternite VIII, ventral aspect
<emphasis id="618F8187C0511B97BE2A497793274632" bold="true">C</emphasis>
male sternite IX, ventral aspect
<emphasis id="6E197DAA900480709A50F3D26CE0DDDA" bold="true">D</emphasis>
male tergites IX and X, dorsal aspect
<emphasis id="A09885B26E809001EC56403CA536BB74" bold="true">E</emphasis>
aedeagus, dorsal aspect
<emphasis id="96EBEA301CD82CEED52AAC02E81B3253" bold="true">F</emphasis>
aedeagus, lateral aspect. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="12719DF990EDC8F7C875CBDBA483B2CE">
<emphasis id="26DC06596647EB3027DC9952D077E3E2" bold="true">Female.</emphasis>
Similar to male, but apical margin of sternite VIII rounded.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5LAE" type="distribution">
<paragraph id="0E6D418A70D18A362441B9E1FFEAEB85">
<heading id="DB9E0630CCA87D00BF43E64BA3559580" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="F0131F964E1B681518235B5E652A6A95">
<collectingCountry id="D572D3C6B017998E8CE239794C9AF97E" name="South Korea">Korea</collectingCountry>
(South).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDMAE" type="etymology">
<paragraph id="92B198167335A0046634270A96721800">
<heading id="808B3AB2B5AA9904DA5468D77A402C1E" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="66520B1001D6FE9D17E5499C4DF407CE">
The name is derived from the Latin
<emphasis id="24C89DCC50D16B5B94A8D1450334593D" italics="true">calculus</emphasis>
meaning pebble, referring to the species marine coastal habitat.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELMAE" type="remarks">
<paragraph id="70989727465DD739585C92E7C99CA2D8">
<heading id="951AE28FFB0CD9DD5DDFC1BA5B641A80" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="B96A17E77ED458457E14C9DF0FB1754E">
This species is extremely similar to
<taxonomicName id="AD9EC75785D723C738DA90D8CA1B12EF" authorityName="Shibata &amp; Fujimoto" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tomokoae">
<emphasis id="0D36C76709C29C19B49573ABDFEC12D1" italics="true">M. tomokoae</emphasis>
</taxonomicName>
in external morphological characters (Figs
<figureCitation id="B3A93BB7BB55C20EBB4F1BA3EC90A1C6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 A D</figureCitation>
,
<figureCitation id="D761DDF0E3A2B8D0E596604D560D74BE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Medon tomokoae A male tergite VIII, dorsal aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure4" httpUri="https://binary.pensoft.net/fig/1243893">4 A D</figureCitation>
) but can be distinguished by the different external form and internal structure of the aedeagus (Figs
<figureCitation id="7C9ACEFEB1E918C1BD3691272FAE9527" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 E, F</figureCitation>
,
<figureCitation id="F3A0A8610716707A42EDBE1FB52A6CD0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Medon tomokoae A male tergite VIII, dorsal aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure4" httpUri="https://binary.pensoft.net/fig/1243893">4 E, F</figureCitation>
) and by the characters listed in Table
<tableCitation id="84B61AC4507F35BF86FB9B52A02C0196" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of morphological characters between Medon calculosus sp. nov., M. tomokoae, and M. prolixus." httpUri="http://table.plazi.org/id/7844F66CEB3CAB4417B8624CE70CE839" tableUuid="7844F66CEB3CAB4417B8624CE70CE839">1</tableCitation>
.
</paragraph>
<caption id="6E31BC713750EF7F553CF07BE1BD8A1E" ID-DOI="10.3897/zookeys.1226.137489.figure4" ID-arpha="C5F37798-86DC-5A57-AF73-49EC36382A1F" httpUri="https://binary.pensoft.net/fig/1243893" startId="F4">
<paragraph id="A7D9546105C608C75088D5AC471ED4A4">
<label id="398ECF879C7476271FA34784F4512278">Figure 4.</label>
</paragraph>
<paragraph id="9191E416FFE76FBA687CADDC77AF09C1">
<taxonomicName id="8F6562A2954776D1AC7723B4B952550F" authorityName="Shibata &amp; Fujimoto" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tomokoae">
<emphasis id="9D4825C46F6037562924F4F8854F4A32" italics="true">Medon tomokoae</emphasis>
</taxonomicName>
<emphasis id="5C7350B4D1145C8975862243499345F6" bold="true">A</emphasis>
male tergite VIII, dorsal aspect
<emphasis id="D39D0B410D432C29B83E535E1D50A3F8" bold="true">B</emphasis>
male sternite VIII, ventral aspect
<emphasis id="965944E82299BE9D9F5C1B4912B90917" bold="true">C</emphasis>
male sternite IX, ventral aspect
<emphasis id="3E1676D4199D5FD57B3C7E6F9F7C5EC4" bold="true">D</emphasis>
male tergites IX and X, dorsal aspect
<emphasis id="75C72980F53996A051FAF0860F816928" bold="true">E</emphasis>
aedeagus, dorsal aspect
<emphasis id="1716D06425C07D82BD686F6EAF4E33CF" bold="true">F</emphasis>
aedeagus, lateral aspect. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="7844F66CEB3CAB4417B8624CE70CE839" ID-Table-UUID="7844F66CEB3CAB4417B8624CE70CE839" httpUri="http://table.plazi.org/id/7844F66CEB3CAB4417B8624CE70CE839" startId="T1" targetIsTable="true">
<paragraph id="07D9F9DD58534B82662BF67B25894B69">
<label id="6193F8B39AAC321360ACFD167979E94E">Table 1.</label>
</paragraph>
<paragraph id="285F2CD1114C8229CAD67C621B711A56">
Comparison of morphological characters between
<taxonomicName id="6231FE08B29A46017483638174EE97A2" authorityName="Ahn &amp; Jeon" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="calculosus" status="sp. nov.">
<emphasis id="355003D483D24998BD2C755D82DD3AEB" italics="true">Medon calculosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="01097A3A3143D5235D0023868B4123EB" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="AF9BD9B2D26AFF0A62F28D5A3BFD505E" authorityName="Shibata &amp; Fujimoto" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tomokoae">
<emphasis id="757A6CCA6A9C4C0FE0DCA55AC8153DF9" italics="true">M. tomokoae</emphasis>
</taxonomicName>
, and
<taxonomicName id="1A100D4B4A67C6539F392A6BF4A0271A" baseAuthorityName="Sharp" baseAuthorityYear="1874" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="2B4F13C9ED17D4E324342DE5CCCA5569" italics="true">M. prolixus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="04247A677906302B760F1B8793C377A4">
<table id="TID0E4TAE" rules="all">
<tbody id="0A74DE3F8CDE55B271725E67A81604CA">
<tr id="A83D88FC5E7890B19614B7FC079D95FD">
<th id="DDFFBF3C9214BF33BB01C1BE62505C4A" colspan="1" isEmpty="true" rowspan="1">-</th>
<th id="75521131BF2B754C00FF1291C796BC14" colspan="1" rowspan="1">
<taxonomicName id="FFE78F39B616132849B5A1D1B308DD32" authorityName="Ahn &amp; Jeon" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="calculosus" status="sp. nov.">
<emphasis id="B9E1C9F298E5D0DAC05CF01CAF92793B" italics="true">M. calculosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B634150E15F3191C55F107E382A8D677" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<emphasis id="7B274788E956BB1239EAAB917A331398" italics="true">
<th id="497BBC0F6ACC382DC24258FA4FED3FC4" colspan="1" rowspan="1">
<taxonomicName id="6190773F8DB8AB764D1EAABA4322E7C8" authorityName="Shibata &amp; Fujimoto" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tomokoae">M. tomokoae</taxonomicName>
</th>
<taxonomicName id="420C52235C43449C3192BA4623EFF920" authority="Body" authorityName="Body" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="prolixus">
<th id="4DD6F7A00D7C123C8CA304C4AE69C6DA" colspan="1" rowspan="1">M. prolixus</th>
<tr id="2F378423DB83D6513FDC72D0C34358FC">
<td id="230457C97E5BB16CE6BF0F1DF6D13324" colspan="1" rowspan="1">Body length</td>
<td id="A830437CFDFA8DA45B15A56BC42F6E7D" colspan="1" rowspan="1">3.03.3 mm</td>
<td id="A0EDADD5138653D7A9DFC02038BE47BF" colspan="1" rowspan="1">2.83.5 mm</td>
<td id="EB490707DDAA4C5E9AF477A3A3D452BC" colspan="1" rowspan="1">4.04.3 mm</td>
</tr>
</taxonomicName>
</emphasis>
</tr>
<tr id="A35F2A6E053013D196A9B3681AC7E87F">
<td id="B3BB8F613E76BF99AD994947596B182F" colspan="1" rowspan="1">Body color</td>
<td id="DE3C6073D448904C92BE4B98591A3217" colspan="1" rowspan="1">brown to reddish-brown</td>
<td id="4346F5C53477746D950598BDDED86B00" colspan="1" rowspan="1">brown to reddish-brown</td>
<td id="AAFA5C76B00EF92F47D80D6962E353FD" colspan="1" rowspan="1">black with brown elytra</td>
</tr>
<tr id="D7185EADF4220AC1130FB0F9F1B66AA6">
<td id="8982A04FC7A9036E7E57F5BC017DB847" colspan="1" rowspan="1">Antenna length</td>
<td id="D400B26B459BED28DB3530BD455974A7" colspan="1" rowspan="1">very long, extending to the middle of elytron</td>
<td id="DDBD733353930302C65DBE14BD5D1721" colspan="1" rowspan="1">very long, reaching to the middle of elytron</td>
<td id="9EA29D2B2902C9334EA6EB104BF3F998" colspan="1" rowspan="1">long, reaching to just behind the pronotum</td>
</tr>
<tr id="4B8577080FFE727746964C9520AE0EC6">
<td id="4C2005402430D5A2E4EE0DD21879F04D" colspan="1" rowspan="1">Antennomeres</td>
<td id="3559258BA2A73E0486BC6539EF686C65" colspan="1" rowspan="1">all elongate</td>
<td id="C453625DCC43E65BB1D743A71AE561F7" colspan="1" rowspan="1">all elongate</td>
<td id="520AB640FA8097BD026108F12F912F44" colspan="1" rowspan="1">not all elongate, 810 subquadrate</td>
</tr>
<tr id="148E3E9B67954766741529CFA917A6FA">
<td id="FA0235DF2BF831DB1597D59A7C56A5F5" colspan="1" rowspan="1">Eye size</td>
<td id="73A02DA0AC67797BCFAB67801B306607" colspan="1" rowspan="1">very small, 0.26 times as long as temple</td>
<td id="67C31DEEE6FB745252708F5CB7D3680B" colspan="1" rowspan="1">very small, 0.28 times as long as temple</td>
<td id="92381816834A75048A65B806366E109B" colspan="1" rowspan="1">small, 0.4 times as long as temple</td>
</tr>
<tr id="14F9C0D9A689311B08058309D466EF61">
<td id="806C63E9C9E466A0C9404A3BA47ACB95" colspan="1" rowspan="1">Black tentorial spots on vertex</td>
<td id="93D7C1BA63E85727F2F6A27EA2F55BF1" colspan="1" rowspan="1">present</td>
<td id="B4A14D49EF59C9A8D6ED8A87BDCA6B2D" colspan="1" rowspan="1">present</td>
<td id="F5C09DB0C9DA91B24570C7B7157118F0" colspan="1" rowspan="1">absent</td>
</tr>
<tr id="E2F95F6AA424AC4F997E54C986472F57">
<td id="10716DBA58AE4B1126C773B8913E35A6" colspan="1" rowspan="1">Pronotum</td>
<td id="4FBF294D29A6AE2853402C257D647D2D" colspan="1" rowspan="1">slightly narrowed posteriorly</td>
<td id="A46B1E1BB761D19409B3C3B656D05D3A" colspan="1" rowspan="1">slightly narrowed posteriorly</td>
<td id="88D56F05703785AC73AA5349501736D3" colspan="1" rowspan="1">parallel-sided</td>
</tr>
<tr id="EB847F13A23A1CACE9172E8188E61528">
<td id="157C7DCDB7018FBE21429718EE0D7EDB" colspan="1" rowspan="1">Hind wings</td>
<td id="0F03B25E13286F7BE739CAB0510A454B" colspan="1" rowspan="1">short, less than 2.0 times as long as elytron</td>
<td id="C1D14128FE895FB4F8A9AE89C4A43FDF" colspan="1" rowspan="1">short, less than 2.0 times as long as elytron</td>
<td id="79E4FFB78071F7F737F9DD4BF92F7A3D" colspan="1" rowspan="1">long, more than 2.0 times as long as elytron</td>
</tr>
<tr id="A7DB4E1CA7768D338607D42632132AF3">
<td id="541C7B8893A52A0B36F06D4E864A41A8" colspan="1" rowspan="1">Male sternite VII</td>
<td id="D341EF6A5448F4F773B3D43D4881E63A" colspan="1" rowspan="1">
subtruncate (Fig.
<figureCitation id="07D015390D8EA367AE37974C0B134FD6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 A</figureCitation>
)
</td>
<td id="0AC61922A503F3C90F71565303E9E1FF" colspan="1" rowspan="1">subtruncate</td>
<td id="D927A502300A3B69FBC88697189E1AC5" colspan="1" rowspan="1">
protruded posteriorly (Fig.
<figureCitation id="F79728B57093513DC4ED9D7F7F9D821A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Medon prolixus A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C aedeagus, dorsal aspect D aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure5" httpUri="https://binary.pensoft.net/fig/1243894">5 A</figureCitation>
)
</td>
</tr>
<tr id="0E8C06884CDEE091AB21673FD6CC699E">
<td id="7B5EB74631034EF8354A6EC39D6B694A" colspan="1" rowspan="1">Male sternite VIII</td>
<td id="72633CB1CD65272AFAD0D81754A0ADD2" colspan="1" rowspan="1">
slightly emarginate (Fig.
<figureCitation id="AC16996C0CAE8F2F1BDA0A02D0383B66" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 B</figureCitation>
)
</td>
<td id="FD5E6BA445110BAA4D22D276EC348CCE" colspan="1" rowspan="1">
subtruncate posteriorly (Fig.
<figureCitation id="6A9FBD10A8F8EF2D8C9256AF49679418" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Medon tomokoae A male tergite VIII, dorsal aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure4" httpUri="https://binary.pensoft.net/fig/1243893">4 B</figureCitation>
)
</td>
<td id="07E51FF803C55395E93F3023D1F9943F" colspan="1" rowspan="1">
deeply emarginate (Fig.
<figureCitation id="8A4DDDEF2D15139EAF9B3520C8DF74BE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Medon prolixus A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C aedeagus, dorsal aspect D aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure5" httpUri="https://binary.pensoft.net/fig/1243894">5 B</figureCitation>
)
</td>
</tr>
<tr id="CA6067B714BCE3F133C2A19781535531">
<td id="812B50DDBB43CBE63A1897F49E6CF819" colspan="1" rowspan="1">Median lobe in dorsal aspect</td>
<td id="1C7BB43BB25E6AF39375279507F2353E" colspan="1" rowspan="1">
broader (Fig.
<figureCitation id="F35D937CA173D81CE01F833D441DBBE2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 E</figureCitation>
)
</td>
<td id="5B11DB027EBA5BBD6354E6CCA9F7484C" colspan="1" rowspan="1">
narrower (Fig.
<figureCitation id="A949CA30B12A71A4FCDDB4E56580E86F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Medon tomokoae A male tergite VIII, dorsal aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure4" httpUri="https://binary.pensoft.net/fig/1243893">4 E</figureCitation>
)
</td>
<td id="773B60B87F3A5AF072318157572333E2" colspan="1" rowspan="1">
narrower (Fig.
<figureCitation id="6EDD37D0E59C135E5A3D5802E3428FC2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Medon prolixus A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C aedeagus, dorsal aspect D aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure5" httpUri="https://binary.pensoft.net/fig/1243894">5 C</figureCitation>
)
</td>
</tr>
<tr id="56B5A3589065C7903713F8BE545B1831">
<td id="09C730651BCAFCF2F4FF43A047FE6ADC" colspan="1" rowspan="1">Median lobe in lateral aspect</td>
<td id="889D8EC78791FA1BC14ADC19CF701362" colspan="1" rowspan="1">
apical process shorter (Fig.
<figureCitation id="8126B72369230DEB7860FC10C5339776" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Medon calculosus sp. nov. A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure3" httpUri="https://binary.pensoft.net/fig/1243892">3 F</figureCitation>
)
</td>
<td id="B41950F83C683C43A37DC5E8DB39D88B" colspan="1" rowspan="1">
apical process longer (Fig.
<figureCitation id="074BAABB4539B7E49EAFA426F9D7DE22" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Medon tomokoae A male tergite VIII, dorsal aspect B male sternite VIII, ventral aspect C male sternite IX, ventral aspect D male tergites IX and X, dorsal aspect E aedeagus, dorsal aspect F aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure4" httpUri="https://binary.pensoft.net/fig/1243893">4 F</figureCitation>
)
</td>
<td id="6B29414D8D99536407158A244C3CEA2E" colspan="1" rowspan="1">
apical process shorter (Fig.
<figureCitation id="DCA09D602BF8C32BD22EE8182FD22C71" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Medon prolixus A male sternite VII, ventral aspect B male sternite VIII, ventral aspect C aedeagus, dorsal aspect D aedeagus, lateral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1226.137489.figure5" httpUri="https://binary.pensoft.net/fig/1243894">5 D</figureCitation>
)
</td>
</tr>
<tr id="710DDD7F521CC98D60DF6AE7D8A5CB00">
<td id="2AF23D32FCAF6BA9DE6B3487AE7F858D" colspan="1" rowspan="1">Microhabitats</td>
<td id="3258351BB2F4F33D7E67925A71AAB699" colspan="1" rowspan="1">pebble / gravel beach</td>
<td id="6D1C66F5CEDF826D6A0260AAFEBFBEBC" colspan="1" rowspan="1">pebble / gravel beach</td>
<td id="673BD2ACFA3B2124EBA2D09B241541DB" colspan="1" rowspan="1">sandy beach</td>
</tr>
</tbody>
</table>
</paragraph>
<caption id="E854B7B1E0828DCACFB55F353B5CE630" ID-Table-UUID="E854B7B1E0828DCACFB55F353B5CE630" httpUri="http://table.plazi.org/id/E854B7B1E0828DCACFB55F353B5CE630" startId="T2" targetIsTable="true">
<paragraph id="C5484611F4B5197E9E72D08A9E900795">
<label id="D1DBFC5539BABFDB61087FEAD078959C">Table 2.</label>
</paragraph>
<paragraph id="FA660A8E95CF2DE0901A2C69ED445596">List of study species with their locality data and GenBank accession numbers.</paragraph>
</caption>
<paragraph id="E790986FACE8064127FF5C281DEC1912">
<table id="TID0EG5AE" rules="all">
<tbody id="23C07153A87F2188421724018B067C5D">
<tr id="0F11D17321C488CD89FD032B4FEF57AF">
<th id="175D8604EEFD9BA67639C7926BCF8820" colspan="1" rowspan="1">Species</th>
<th id="1F56D576AAE4069BEBC49AF7DA847D36" colspan="1" rowspan="1">Collection locality</th>
<emphasis id="518D5A065C09F761AF403815F620D55C" italics="true">
<th id="35E0EC7B57337D110308D110DC6DA73B" colspan="1" rowspan="1">
<abbrev id="ABBRID0ELZAE" xlink_title="morphological and molecular characters">COI</abbrev>
</th>
<tr id="964AF0EC519E6266BABDA55C0FF2B4E6">
<td id="7DADD05839C9AE3C5CE26AE30652E370" colspan="1" rowspan="1">
<taxonomicName id="462590A9C05A88D941876DA9206F7D1C" authorityName="Ahn &amp; Jeon" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="calculosus" status="sp. nov.">Medon calculosus</taxonomicName>
<taxonomicNameLabel id="7913869516C60959C4D02EF523D57724" rank="species">sp. nov.</taxonomicNameLabel>
</td>
<td id="197156A74FD3281CED4B114B3F76B7C4" colspan="1" rowspan="1">KOREA: Gyeongbuk, Gyeongju-si</td>
<td id="928A5E8AD452ABC7005A0D4910C17451" colspan="1" rowspan="1" style="color: #396ba6">
<ext-link id="2992DE1ED405B72F9F8B57FEEF95B20E" ext-link-type="gen" xlink_href="PP578214" xlink_type="simple">PP 578214</ext-link>
</td>
</tr>
</emphasis>
</tr>
<tr id="A1EB637F40AA8A555E1197186493DBAD">
<td id="131670B2A7CA74E158B5429C07A8F293" colspan="1" rowspan="1">
<taxonomicName id="E2BA5DC48E76A4F15EFDEB8743FEE2C0" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="EE41B5D6C08AEF39A51BD869D60B28E6" italics="true">Medon prolixus</emphasis>
</taxonomicName>
1
</td>
<td id="DD3888D6655C13D6CEF778C32D644558" colspan="1" rowspan="1">KOREA: Gyeongbuk, Uljin-gun</td>
<td id="A48B51FDEB6325D78693535798715D01" colspan="1" rowspan="1" style="color: #396ba6">
<ext-link id="B71CDC90EC20B37930D83FDE9EB3CD99" ext-link-type="gen" xlink_href="PP578207" xlink_type="simple">PP 578207</ext-link>
</td>
</tr>
<tr id="2E142104F479FF201D246BE884E90470">
<td id="A757C41E6081D842BAD6488B7785C55B" colspan="1" rowspan="1">
<taxonomicName id="F0CA660577D9EF5E7B696B93FC0B6D20" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="88B0DA0A22E8963DC0EB3330F28580F5" italics="true">Medon prolixus</emphasis>
</taxonomicName>
2
</td>
<td id="F26DCE085C581E97E9AFB166EC3091FE" colspan="1" rowspan="1">KOREA: Gyeongbuk, Uljin-gun</td>
<td id="BF8F3077D0FE1A436F2C6A3162A79703" colspan="1" rowspan="1" style="color: #396ba6">
<ext-link id="5DF8CF615DD03A29ED91B334212F9915" ext-link-type="gen" xlink_href="PP578210" xlink_type="simple">PP 578210</ext-link>
</td>
</tr>
<tr id="BFF524B31688CF4A9F651CBCC0F5A376">
<td id="051941D5720D035EC55A890F164F0D38" colspan="1" rowspan="1">
<taxonomicName id="62C50C50A9B84674264B8F86487185A7" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="DAB497941FB0D505D2BC931796F5634A" italics="true">Medon prolixus</emphasis>
</taxonomicName>
3
</td>
<td id="1C935294993BC3471AE07FF37B3D5B2D" colspan="1" rowspan="1">KOREA: Gyeongnam, Geoje-si</td>
<td id="5CCA1674DA7CBCB1EDC194FCFB5AA061" colspan="1" rowspan="1" style="color: #396ba6">
<ext-link id="7AF91FD906107DB402FA8A9E6232D2A3" ext-link-type="gen" xlink_href="PP578211" xlink_type="simple">PP 578211</ext-link>
</td>
</tr>
<tr id="5EF85060183C9CDFD1945F625C68DB57">
<td id="049C1DE21C3076AC68721CB0C804AF15" colspan="1" rowspan="1">
<taxonomicName id="BD7A43F1A4D260A4350BCE94C6D55500" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="8BB982340F112F7E16B7738873654353" italics="true">Medon prolixus</emphasis>
</taxonomicName>
4
</td>
<td id="F7B0369B1C2EF9BC630BA414AA81E9E5" colspan="1" rowspan="1">KOREA: Gyeongnam, Tongyeong-si</td>
<td id="5C1E3F6E242D291786EA57F9A0FBC2D7" colspan="1" rowspan="1" style="color: #396ba6">
<ext-link id="D8662C825F18BDACFA3CA330D7C8F632" ext-link-type="gen" xlink_href="PP578212" xlink_type="simple">PP 578212</ext-link>
</td>
</tr>
<tr id="00033C583EA11CCAB0092DC1D5620909">
<td id="FEAB0925D042C7AA6E1E9C7CEDF8170B" colspan="1" rowspan="1">
<taxonomicName id="9C201E203AF3941B91A38D544FACCFA1" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="DC59FEA407DAF120B3ED9CBBED1ED062" italics="true">Medon prolixus</emphasis>
</taxonomicName>
5
</td>
<td id="0507F413D865E82AB91C6FC336F8364B" colspan="1" rowspan="1">KOREA: Jeju Prov., Jeju-si</td>
<td id="C46461F49366A1D1CA5BEA93AEEB5093" colspan="1" rowspan="1" style="color: #396ba6">
<ext-link id="B1B58718E019853F448DE7D3626E7BEE" ext-link-type="gen" xlink_href="PP578213" xlink_type="simple">PP 578213</ext-link>
</td>
</tr>
<tr id="FA4290DEEAECF5BCF3D14876AFE7F267">
<td id="BCC2C8E00EDE793030F55E8B89F34786" colspan="1" rowspan="1">
<taxonomicName id="E791CE3D40A87E303A70D22934ACBF56" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="83EE68C1A5768535CDEBD5B0F1DCBF05" italics="true">Medon</emphasis>
sp. 1
</taxonomicName>
A
</td>
<td id="602053DD7AAE3FA7C6EB8E39F8A4D2F2" colspan="1" rowspan="1">KOREA: Gangwon, Sokcho-si</td>
<td id="25CEAB6D9C5D95EA054614B488582E30" colspan="1" rowspan="1" style="color: #396ba6">
<ext-link id="0A036E622563ECBFF076B48BE1894779" ext-link-type="gen" xlink_href="PP578208" xlink_type="simple">PP 578208</ext-link>
</td>
</tr>
<tr id="614F555C361D1CD25C46516EE49D1D45">
<td id="7A1E79001FEF5D3B681A0AB8C633FA9C" colspan="1" rowspan="1">
<taxonomicName id="C2457DC7A8BD3D2A257F8AA3A5352BD5" class="Insecta" family="Staphylinidae" genus="Medon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="A3341A2975630C322E0214EAF096B931" italics="true">Medon</emphasis>
sp. 1
</taxonomicName>
B
</td>
<td id="CBA485D57773032BC957190374419CB7" colspan="1" rowspan="1">KOREA: Jeonnam, Goheung-gun</td>
<td id="471F47354402598ADBC363CCD205EEB1" colspan="1" rowspan="1" style="color: #396ba6">
<ext-link id="1213F8B95FFCF87F22BC9E04E1ADB1BB" ext-link-type="gen" xlink_href="PP578209" xlink_type="simple">PP 578209</ext-link>
</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,232 @@
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<mods:title id="95A50B60B6CB76BC8897F53E35BB3655">Mutatus gen. nov., a new genus of Phrurolithidae (Arachnida, Araneae) from China, with descriptions of four new species</mods:title>
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<mods:namePart id="46092497EB6C2A3D2D8DE115AB2E50E4">Mu, Yannan</mods:namePart>
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<mods:affiliation id="7B24462BDE798A65C0DA3310660A6C15">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China &amp; Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="1898FAE5B3FB3D2351863479678388ED">Wang, Luyu</mods:namePart>
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<mods:affiliation id="A38C39BF929A727BD588447B02B7BA8A">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:affiliation id="D63BC8F0259858A69272F8D4B9CEA7D6">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:affiliation id="B25B378308A6DAC2D576B94B535416EB">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<subSubSection id="4F4E87654397A7F95BD94B93A357F29C" type="nomenclature">
<paragraph id="6EFFAE38F49CDE3EB5489F1B70C3A23B">
<taxonomicName id="379C1CD4AECA953BA9E280B68E8EB62A" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Phrurolithidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="61FC929EA524E4CA4CCCDA1B47D7CDEE" italics="true">Mutatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2C2A3F2531DDCBC7FDE92477A283848" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDHAC" type="etymology">
<paragraph id="529C540FEFC3D90796B5468C286DD495">
<heading id="309DDE48AC1393A9591720C03B693BF0" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="8AD89A0CEB44453D8F3A06486CB0DCA7">
The genus name is derived from a Latin word “
<emphasis id="D2083518BCF840FC6B17280140D48725" italics="true">mutatio</emphasis>
”, meaning “ variable ”, referring to the unique combination of characters in this new genus. Unlike the similar genera
<emphasis id="D63B7841A193C8DDB6D5275E2A43B1EF" italics="true">Lingulatus</emphasis>
, which possesses only a tegular sheet, and
<emphasis id="8B144BD8D0C93A49ADB8E65078158D92" italics="true">Xilithus</emphasis>
, which features only prolateral tegular apophysis, the new genus exhibits both characteristics in the male palp. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPHAC" type="diagnosis">
<paragraph id="D1934204C9938BAA983F0A02A066DEE7">
<heading id="1CC1804C27B6C9B815D38D95663D2F07" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B89AB8EFA0E487E9B078630EC8402406">
The male of new genus resembles those of the genera
<taxonomicName id="D8C43A3CC12CFFAACCC4AECF472F8F6C" authorityName="Mu &amp; Zhang" authorityYear="2022" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="A9D8850D68308A7BFF2F17AFCFDA7C8A" italics="true">Lingulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="CFFA598B3EBDBEE00FDF2A1AB9F389A7" authorityName="Liu &amp; Li" authorityYear="2023" class="Arachnida" family="Phrurolithidae" genus="Xilithus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="0A4D87E8C4AB1B3E8BA6388CFEE8F795" italics="true">Xilithus</emphasis>
</taxonomicName>
in having both tegular sheet and prolateral tegular apophysis, but
<taxonomicName id="D16A5267DB19B104A36DD1E065CB4DDE" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="1699762992B1453CBC1121D344C8A518" italics="true">Mutatus</emphasis>
</taxonomicName>
can be differentiated by the absence of a ventral tibial apophysis (present both in
<taxonomicName id="6FC3468D4B9F136A87828984395599AF" authorityName="Mu &amp; Zhang" authorityYear="2022" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="5A275823D923A65566B11AF018200138" italics="true">Lingulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="A7154ACEDABF458671EA236B81A54D69" authorityName="Liu &amp; Li" authorityYear="2023" class="Arachnida" family="Phrurolithidae" genus="Xilithus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="2493142AC8410924C6AD51F4C6DAC30B" italics="true">Xilithus</emphasis>
</taxonomicName>
), tegular sheet present in both
<taxonomicName id="62F288777AB163B4B525A2EB0CEB3C4E" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="9E351113E979507585241CE8C739A1DD" italics="true">Mutatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4A157C46406955012BF544CB603F136" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="920AAD8A97B94CD91693E94A176D68E5" authorityName="Mu &amp; Zhang" authorityYear="2022" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="011994E0995657BF972B30F551307F7E" italics="true">Lingulatus</emphasis>
</taxonomicName>
(absent in
<taxonomicName id="5293B3E7FE232A5BB35DE96E4652B5F5" authorityName="Liu &amp; Li" authorityYear="2023" class="Arachnida" family="Phrurolithidae" genus="Xilithus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="7114A9EB839FB8EC271C1DC1B352B572" italics="true">Xilithus</emphasis>
</taxonomicName>
), and sperm duct V-shaped in both
<taxonomicName id="A060031F0156CFE55836367A0713A9AB" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39C13430EEA30076436583647541FC3D" italics="true">Mutatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6C1509338ABCCBF59D281E033020F87" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="65D5CA58047DE2CE6923022F0226C001" authorityName="Mu &amp; Zhang" authorityYear="2022" class="Arachnida" family="Phrurolithidae" genus="Lingulatus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="1B1C6343180A85DE5C7B10FBC74453A5" italics="true">Lingulatus</emphasis>
</taxonomicName>
(U-shaped in
<taxonomicName id="F96F325DA2EA3E02AFA46D4464569ED0" authorityName="Liu &amp; Li" authorityYear="2023" class="Arachnida" family="Phrurolithidae" genus="Xilithus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="8DB654225DC9E19E19BF1B878E828934" italics="true">Xilithus</emphasis>
</taxonomicName>
). The female of new genus resembles
<taxonomicName id="70D793769266881147CBDA7305753B76" authorityName="Liu &amp; Li" authorityYear="2023" class="Arachnida" family="Phrurolithidae" genus="Xilithus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="7C4B240B92DE2D6F4C30114BDD8FCEB4" italics="true">Xilithus</emphasis>
</taxonomicName>
in having a pair of large concavities (atria), a pair of small bursae, and clavate spermathecae, but it can be differentiated by the presence of a protuberance at the middle part of the lateral margin of the atrium (absent in
<taxonomicName id="68739DD66542103B6FBF394366D865EE" authorityName="Liu &amp; Li" authorityYear="2023" class="Arachnida" family="Phrurolithidae" genus="Xilithus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="23A5CCF8D0F717612A3BD083B9085E84" italics="true">Xilithus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQKAC" type="description">
<paragraph id="091340DE61FD0FC8DDFBD08E361E7E24">
<heading id="E6ECF41DEF4A8775CE29413506D7F2E5" reason="title">Description.</heading>
</paragraph>
<paragraph id="E6DAE4FF4C92157CB81EBEDF2FA5BA21">
Small, total length
<quantity id="E46F6A693CA056F243C5167B594B8B60" metricMagnitude="-3" metricUnit="m" metricValue="3.98" metricValueMax="4.78" metricValueMin="3.18" unit="mm" value="3.98" valueMax="4.78" valueMin="3.18">3.184.78 mm</quantity>
. Carapace oval, smooth, brownish to black, widest at coxae II and III, highest near fovea, with several markings resembling flowing water droplets beside fovea. Fovea dark-red.
<abbrev id="ABBRID0EWKAC" xlink_title="posterior eye row">
<collectionCode id="1FD465978337B5E32E9D6213B2C4A527" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" type="Herbarium">PER</collectionCode>
</abbrev>
slightly wider than
<abbrev id="ABBRID0E1KAC" xlink_title="anterior eye row">
<collectionCode id="30A06C24718CF510A4E7C02722CFC9CB">AER</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0E5KAC" xlink_title="posterior eye row">
<collectionCode id="97B0989CF83382F7A36897E3F8CFD652" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" type="Herbarium">PER</collectionCode>
</abbrev>
recurved in dorsal view,
<abbrev id="ABBRID0ECLAC" xlink_title="anterior eye row">
<collectionCode id="E0441F50AD3B8830EE79928AC5CD3278">AER</collectionCode>
</abbrev>
recurved in dorsal view. Chelicerae brown, with 1 long and 1 short spine anteriorly. Endites longer than wide, labium wider than long. Sternum brown, shield-shaped, smooth and without pattern, longer than wide. Legs dark brown, all femora with 1 dorsal spine; femur I with 35 prolateral spines distally, femur II with 02 prolateral spines distally; tibiae I II usually with 67 pairs of ventral spines, tibiae III IV without ventral spines; metatarsus I with 45 pairs of ventral spines, metatarsus II usually with 4 proventral spines and 3 retroventral spines, metatarsi III IV without ventral spines but with distal preening brush. Leg formula 4123. Abdomen oval, gray to black in dorsal view, with dorsal scutum covering almost entire abdomen in males, scutum in females absent.
</paragraph>
<paragraph id="BCE290C66BE4E9E6D573FF976049CB74">Male palp: femur with well-developed apophysis located at middle part; patella short, as long as tibia; tibia slightly bulged ventrally, with retroventral and retrolateral apophyses; cymbium longer than femur; bulb pyriform; prolateral tegular apophysis and tegular sheet (spade-shaped) originating near base of embolus; embolus acicular; conductor absent.</paragraph>
<paragraph id="E34D63C0F0C8D66EEB23CA29C8BA1D2E">Epigyne: epigynal plate weakly sclerotized, with pair of atria; septum dumbbell-shaped; copulatory openings small, separated; copulatory ducts thin; connecting tubes thick, shorter than copulatory ducts; bursae small; spermathecae clavate, located posteriorly; glandular appendages mastoid-shaped.</paragraph>
</subSubSection>
<subSubSection id="SECID0EILAC" type="type species">
<paragraph id="BA8501403A97E9E8F5DEA4EAA6AE6739">
<heading id="E4F40BDDCA20122FBC46304EC39FE0FB" reason="title">
<typeStatus id="35296EED5E3CABA4E807D3FFBB3A556A">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="A71F42F2A1E2A609E08AC3BF663D2FE7">
<taxonomicName id="D1FDFD91AB18E40E5AF02C4A13E5AA41" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gelao" status="sp. nov.">
<emphasis id="3BECCFFC2329571ED686BC39F4D4EAB5" italics="true">Mutatus gelao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10CAF2068D3DB2B6B3A1367333076D32" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZLAC" type="Composition">
<paragraph id="4904BE74F2770599CE95DD947C73EFEB">
<heading id="8F8A52ADADA28ACF6AC76898D4E436AB" reason="title">Composition.</heading>
</paragraph>
<paragraph id="A4F5FF27045965B6B329215E0B2BA87B">
<taxonomicName id="00747045CD0465550D6F6136B3BDBED2" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gelao" status="sp. nov.">
<emphasis id="8D2D68B7E1E6FDF7B950DF843DB361B5" italics="true">Mutatus gelao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67814A6E6E9C77F80971B90609493F44" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="B0A1151AD7EF45B97783FC75194B7392" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tujia" status="sp. nov.">
<emphasis id="16D42F37F88D4CE4DAFCF730ED0BB5B4" italics="true">M. tujia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DE33118102DB719E6CF97BCCA6847BE2" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="07B6BF2EAA84A6B76292A6E4FE2E9B6E" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tianyan" status="sp. nov.">
<emphasis id="279D7E820E3999E1999A306B928D68D8" italics="true">M. tianyan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABEEC41448825C6F3FCEDF3005032DB2" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="67C2D5B9B9CA286086DBCDF42BB00753" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="yintiaoling" status="sp. nov.">
<emphasis id="3510481AD38D5DBD127732EF48AC7DFB" italics="true">M. yintiaoling</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44EBE27420B913ED663DE3F9D53BE6BB" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELNAC" type="distribution">
<paragraph id="5D7462E8FB83F73C5ECF83EFD1FEC24F">
<heading id="AAD812B6E1AD3EE7D1ED9DFCB778A5B6" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="9778852ABC49B3339A1ADB931FE83747">
<collectingCountry id="C052D38E157D1BC4A32B5AA9937EBCF6" name="China">China</collectingCountry>
(
<collectingRegion id="10979BA1333720CB5F0AC8CB56B6B8EC" country="China" name="Hubei">Hubei Province</collectingRegion>
,
<collectingRegion id="B88532A57A99C03FBCD6D10D9E4154D3" country="China" name="Guizhou">Guizhou Province</collectingRegion>
,
<collectingRegion id="1C691A24BA400CD5D9ADC7443C7D5413" country="China" name="Chongqing">Chongqing</collectingRegion>
Municipality).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,597 @@
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<mods:mods id="A264F79F165C9F69723A2BDA8AF8C4B7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="66E621F2ECB84A69D01685938923BC45">
<mods:title id="A796E1059045A1CC790ECC09D307BBE8">Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province</mods:title>
</mods:titleInfo>
<mods:name id="925750875E7F2D27DDB352BC3EF47C94" type="personal">
<mods:role id="3D55F682DDD28CDE2557D3B4D460680E">
<mods:roleTerm id="8980327187D86FF9F68E44135B1CECE7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="06244868F478AB12E884175ED34EF4E3">Hedin, Marshal</mods:namePart>
<mods:nameIdentifier id="D2C38F18E029970402D6C46F7A321D98" type="ORCID">0000-0001-7683-9572</mods:nameIdentifier>
<mods:affiliation id="379481E1D4C94259E107FD58E09FBB52">Department of Biology, San Diego State University, San Diego, California 92182 4614, USA</mods:affiliation>
</mods:name>
<mods:name id="9064BA4BF1C7ACA6E5DD31BB61B1DE9F" type="personal">
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</mods:role>
<mods:namePart id="C1CA665CCFB4F709807740936B883315">Ramírez, Martín J.</mods:namePart>
<mods:nameIdentifier id="88611C619AB8B8E0C1A5F5E089BE1BA6" type="ORCID">0000-0002-0358-0130</mods:nameIdentifier>
<mods:affiliation id="6AFD9113FF2C71EA902C5FD19F9C7955">Division of Arachnology, Museo Argentino de Ciencias Naturales - CONICET, Buenos Aires, Argentina</mods:affiliation>
</mods:name>
<mods:name id="02307A2061F27B7E09A50282E60CF3DA" type="personal">
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</mods:role>
<mods:namePart id="0F3E4C90759FBD371B7C54DFFB27428A">Monjaraz-Ruedas, Rodrigo</mods:namePart>
<mods:nameIdentifier id="53A9FDDE7D42BBD849E18E857AC3462E" type="ORCID">0000-0002-6462-3739</mods:nameIdentifier>
<mods:affiliation id="47FFB50C42D71C1F0913A8EDE8EE2D40">Department of Biology, San Diego State University, San Diego, California 92182 4614, USA &amp; Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, California 90007, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="08D6AFE9CE676C6E28F8C3D056C29296">text</mods:typeOfResource>
<mods:relatedItem id="1E0C9A472B8AC77E245E68A2B95C83A2" type="host">
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<mods:title id="36845D8B50CC8CE2CE447D150163C7A7">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="C89AF829A273BCAEA937F560E0557FB9">
<mods:date id="2BA91726F3E661FE704E6C5F146139D4">2025</mods:date>
<mods:detail id="6C833A3C75C8E22DE2AA1566AA357B59" type="pubDate">
<mods:number id="1461698102A3CAF37AD1EEEDAC28762C">2025-02-06</mods:number>
</mods:detail>
<mods:detail id="D8E64A835712C434A96BAF7EDECEAEC6" type="volume">
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<mods:classification id="908A2C7BE2158A638703478BF2AF7B3D">journal article</mods:classification>
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<mods:identifier id="A4F0D9DE3865C47308DC5CD863C6BD11" type="ZooBank">AC08832F-748C-44E2-ADFD-514F258779C8</mods:identifier>
</mods:mods>
<treatment id="514AFCE05B2A5ABE8B0BD2F27531210A" ID-DOI="http://doi.org/10.5281/zenodo.14827257" ID-Zenodo-Dep="14827257" ID-arpha="514AFCE0-5B2A-5ABE-8B0B-D2F27531210A" ID-zoobank="https://zoobank.org/76BC699E-09AE-4ABA-B8AF-7AFCE8676EFA" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A" LSID-ZBK="urn:lsid:zoobank.org:act:76BC699E-09AE-4ABA-B8AF-7AFCE8676EFA" httpUri="http://treatment.plazi.org/id/514AFCE05B2A5ABE8B0BD2F27531210A">
<subSubSection id="FB9F7639B3A953EA31F796AB6A50FB51" type="nomenclature">
<paragraph id="9FFEDA68B3FE60634BB53170F733EF57">
<taxonomicName id="C84408F64B34394A480EB76773E36A28" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="furtiva" status="sp. nov.">
<emphasis id="53DE53CE6A2CFD64275D8C8D9D2660CF" italics="true">Cybaeozyga furtiva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AC60797C4489F66CB17F6A96E7AE9997" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0D505FC0D09F2C33013DC8F10701D44E" type="description">
<paragraph id="10104DAF93788B9D296092F904F45AFB">
<figureCitation id="CF4F494167DB673C0711A125D6B46694" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cybaeozyga furtiva sp. nov. ♀ (A MACN - Ar 38880, B H holotype) A live specimen MACN - Ar 38880 B dorsal view C ventral view D prosoma dorsal view E prosoma lateral view F prosoma anterior view G eyes dorsal view H eyes anterior view." figureDoi="10.3897/zookeys.1226.140204.figure8" httpUri="https://binary.pensoft.net/fig/1243097">Figs 8</figureCitation>
,
<figureCitation id="3FFE51192D1E10A9EA032CA8F78577B6" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cybaeozyga furtiva sp. nov. ♀ (A, J M holotype B I paratype MACN - Ar 46970) A patellae and tibiae I, arrows to fracture lines B patellae III IV fractured, arrows to fracture lines C left leg I, prolateral view D same, retrolateral view E tarsus I prolateral view F tarsus IV prolateral view G tibia I ventral view H metatarsus I ventral view I left leg IV prolateral view J spinnerets ventral view K epigyne ventral view L cleared epigyne ventral view M cleared epigyne dorsal view. Abbreviations: BG = Bennetts gland, FD = fertilization duct, PCD = proximal copulatory duct, S = spermatheca." figureDoi="10.3897/zookeys.1226.140204.figure9" httpUri="https://binary.pensoft.net/fig/1243098">9</figureCitation>
,
<figureCitation id="4EE7DAA6745914D561864F714CC7C2DF" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Species distributions of Siskiyu armilla sp. nov. and Cybaeozyga, including C. furtiva sp. nov. and C. heterops. Locations approximate, see text and Suppl. material 1 for precise location data." figureDoi="10.3897/zookeys.1226.140204.figure10" httpUri="https://binary.pensoft.net/fig/1243099">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETBAI" type="material">
<paragraph id="5536C7F7BA5474CF2EEF6EA366CAD0FF">
<heading id="233D37E430D5F7F34D34A38D6125A5FA" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="4E0ECA2B759724E2D4E10BFB4910BC52">
<materialsCitation id="119F78203FAEA2929A6F37F9BB48ED9B" collectingDate="2017-06-25" collectingDateMax="2017-06-29" collectingDateMin="2017-06-25" collectionCode="GPS, SDSU" collectorName="E Crescent City &amp; M. Ramirez &amp; P. Michalik" county="Del Norte Co." elevation="150" location="Del Norte Co." specimenCode="TAC 000889" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="holotype">
<emphasis id="E7DBDC4FA59598835C1CF4C5ECEE3311" bold="true">
<emphasis id="BFBB52AACA99C5FEBEB5686AAC73CE78" bold="true" italics="true">
<typeStatus id="D3B001FC9D42974563FEB3500B9F3196" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingRegion id="4124492CBB152A5B0D05759377C1A7D1" country="United States of America" name="California">California</collectingRegion>
,
<collectingCounty id="194CD6124300837E906399734CF9677B">Del Norte Co.</collectingCounty>
</emphasis>
<specimenCount id="E90119CF332F6B0289FDA68B4DE770EA" count="1" type="female"></specimenCount>
;
<collectorName id="8E557C2AAACF305329538459BD916AB2">E Crescent City</collectorName>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBCAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-124.000000,41.800000]}">41.8, - 124.0</named-content>
(
<collectionCode id="D89AA60E5A87E0ACB3B6CD956698CB62">GPS</collectionCode>
, ±
<quantity id="DB1231C4076D1B25CB591689261CACF6" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
); elev.
<quantity id="080DB7A3D678E797659F9E4B82CF707E" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">
<elevation id="185A400D74318DFFC75BA14EDE6D0AD1" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="27D26958AD1605807F7127B5F0613B9B" value="2017-06-25" valueMax="2017-06-29" valueMin="2017-06-25">
<collectingDate id="DDA052A0C6F25ACEE4208EFD978B33E2" value="2017-06-25" valueMax="2017-06-29" valueMin="2017-06-25">2529 Jun. 2017</collectingDate>
</date>
; wet mixed forest with redwood; coll.
<collectorName id="7B57F90EB6A116F053C400A28FC3D437">M. Ramírez</collectorName>
&amp;
<collectorName id="E485AF6EB6A489139380FB83472BE293">P. Michalik</collectorName>
;
<collectionCode id="4B91F279FD6806FB26BB32DB8D078468" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="4E75AB68D164171BFA5EC2175C4EA10B" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">TAC 000889</specimenCode>
</materialsCitation>
;
<materialsCitation id="C217476CBBD4D930CFD74D76349C816A" collectingDate="2017-06-25" collectingDateMax="2017-06-29" collectingDateMin="2017-06-25" collectionCode="GPS, SDSU" collectorName="E Crescent City &amp; M. Ramirez &amp; P. Michalik" county="Del Norte Co." elevation="150" location="Del Norte Co." specimenCount="1" stateProvince="California" typeStatus="paratype">
<emphasis id="C61121314FA2D7545BCBCF32D2AC329E" bold="true" italics="true">
<typeStatus id="F110EEC01039441D9EEADD59FBA13023" type="paratype">Paratypes</typeStatus>
</emphasis>
: same data as holotype
</materialsCitation>
; •
<materialsCitation id="FF79368696523B19C1354428DC38FC13" collectionCode="SDSU" specimenCode="TAC 000890" specimenCount="2" specimenCount-female="2" stateProvince="California" typeStatus="paratype">
<specimenCount id="89357FE0B43C02D4FE69BC23BDE57D99" count="2" type="female">2 ♀</specimenCount>
;
<collectionCode id="937A17AF38DB1E524AFC0C232D270E70" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="7EC97370C5039397FF2079D04827C34C" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">TAC 000890</specimenCode>
</materialsCitation>
; •
<materialsCitation id="F6D17CFC3ED739098F7A07EC67362CC8" collectionCode="MACN" specimenCount="3" specimenCount-female="3" stateProvince="California" typeStatus="paratype">
<specimenCount id="E4B597D7CCBD28795DC1914AEA71FE3C" count="3" type="female">3 ♀</specimenCount>
;
<abbrev id="ABBRID0EJCAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="6F161DFB4C496B273A5A15C50D453E85" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
- Ar 46970
</materialsCitation>
; •
<materialsCitation id="CE2E8047796981D2D7D3E49FB37C1B4B" collectionCode="MACN" specimenCount="2" specimenCount-female="2" stateProvince="California" typeStatus="paratype">
<specimenCount id="7E3EB4E57EE61B38F0CA6A22BEE563EB" count="2" type="female">2 ♀</specimenCount>
;
<abbrev id="ABBRID0ENCAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="7A5939441DF6F9D0E48AD02BE0A3F3C2" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
- Ar 38631
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERCAI" type="material">
<paragraph id="0DAD9A68032C10AF04BB2A1B1989F1BB">
<heading id="071C2E47F50DFE8E343B785FB9F0F62B" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="8A964E67B0690ACD0967357A8482313E">
<emphasis id="11E8C1E1022DB9D5D11F854D8639834E" bold="true">
<materialsCitation id="20EAAE496FD5A1AAAC8B37B97B01EA70" collectorName="Del Norte Co" location="California" specimenCount="1" stateProvince="California">
<collectingRegion id="C317DCE6116744482532CCF491579B9E" country="United States of America" name="California">California</collectingRegion>
,
<collectorName id="189030DE8F596E77A66F76F77C6A9593">Del Norte Co</collectorName>
</materialsCitation>
.
</emphasis>
<materialsCitation id="C8EB1E4CAF83DF8A273833DD1157DC1F" collectingDate="2017-06-25" collectingDateMax="2017-06-29" collectingDateMin="2017-06-25" collectionCode="MACN" collectorName="E Crescent City &amp; M. Ramirez &amp; P. Michalik" county="Del Norte Co." elevation="150" location="Ar" specimenCount="9" specimenCount-female="9" stateProvince="California">
<specimenCount id="5AAEECFA7401BE4B4B91F8D5D8752CDA" count="9" type="female">9 ♀</specimenCount>
(together with several immatures); same data as holotype;
<abbrev id="ABBRID0EZCAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="7ED1D8F289558D67CE54C6AE3558B7CD" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
-
<location id="44657B9230D09444BE81543968C3F973" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A:44657B9230D09444BE81543968C3F973" name="Ar">Ar</location>
38888
</materialsCitation>
; •
<materialsCitation id="D6997EC330FC3BC65E20254A64EBFAB2" collectingDate="2017-06-25" collectingDateMax="2017-06-29" collectingDateMin="2017-06-25" collectionCode="MACN" collectorName="E Crescent City &amp; M. Ramirez &amp; P. Michalik" county="Del Norte Co." elevation="150" location="Ar" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="4E660D3CAA6921EFD47F56C47C36C617" count="1" type="female">1 ♀</specimenCount>
; same data as holotype;
<abbrev id="ABBRID0E4CAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="A34FE59D476BC9B1863EB0F0C4056399" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
-
<location id="07BA55CF4B1230D57F156BF734FE42EA" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A:07BA55CF4B1230D57F156BF734FE42EA" name="Ar">Ar</location>
38862; sample MJR- 2128
</materialsCitation>
; •
<materialsCitation id="8336CF3CF364B73F2354B37167CEEBBE" collectingDate="2017-06-25" collectingDateMax="2017-06-29" collectingDateMin="2017-06-25" collectionCode="MACN" collectorName="E Crescent City &amp; M. Ramirez &amp; P. Michalik" county="Del Norte Co." elevation="150" location="Ar" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="49235608B58E0ECF371F78695BCBF4B4" count="1" type="female">1 ♀</specimenCount>
; same data as holotype;
<abbrev id="ABBRID0EBDAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="AF26B54844F640D3FE9A851B5CE57A35" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
-
<location id="2D5715C347570881CA0FB0850691A88F" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A:2D5715C347570881CA0FB0850691A88F" name="Ar">Ar</location>
38880; samples MJR- 1985, 1986, 1987, 1989; photos 79327934
</materialsCitation>
; •
<materialsCitation id="37566F4EB164D443AECA550B83CD6D20" collectingDate="2017-06-25" collectingDateMax="2017-06-29" collectingDateMin="2017-06-25" collectionCode="MACN" collectorName="E Crescent City &amp; M. Ramirez &amp; P. Michalik" county="Del Norte Co." elevation="150" location="Ar" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="DE936B726ADF2AFF530E2624959291E2" count="1" type="female">1 ♀</specimenCount>
; same data as holotype;
<abbrev id="ABBRID0EFDAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="37F721ACFBEB8F180FE46639FF17CD8B" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
-
<location id="0B331FE0836415C1D2C83D725F848525" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A:0B331FE0836415C1D2C83D725F848525" name="Ar">Ar</location>
38948; sample MJR- 2036 attachment disks
</materialsCitation>
; •
<materialsCitation id="A0190E3976372AB9FF7581070B5CB8C5" collectingDate="2017-06-25" collectingDateMax="2017-06-29" collectingDateMin="2017-06-25" collectionCode="MACN" collectorName="E Crescent City &amp; M. Ramirez &amp; P. Michalik" county="Del Norte Co." elevation="150" location="Ar" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="4F2F13BE481775578DEF69090E477387" count="1" type="female">1 ♀</specimenCount>
; same data as holotype;
<abbrev id="ABBRID0EJDAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="5CF9C130D06BD511C9979644D90EA264" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
-
<location id="50AA15A46AB63B44E5BEE103678CB91E" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A:50AA15A46AB63B44E5BEE103678CB91E" name="Ar">Ar</location>
38936; samples MJR- 1985, 1987, photos 79157930
</materialsCitation>
; •
<materialsCitation id="706A0D75FD2548E897BBFF885BA412C8" collectingDate="2017-06-25" collectingDateMax="2017-06-29" collectingDateMin="2017-06-25" collectionCode="MACN" collectorName="E Crescent City &amp; M. Ramirez &amp; P. Michalik" county="Del Norte Co." elevation="150" location="Ar" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="562AEB3FDF8797885A0828885864FAF3" count="1" type="female">1 ♀</specimenCount>
; same data as holotype;
<abbrev id="ABBRID0ENDAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="C9E61EC5F702E3160B42CA9733790160" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
-
<location id="171FF2D517453FAEB411D89AC2261C5F" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A:171FF2D517453FAEB411D89AC2261C5F" name="Ar">Ar</location>
38958; sample MJR- 2130
</materialsCitation>
; •
<materialsCitation id="0B81726BF252155CD0E1C08BCE61C770" collectingDate="2017-06-25" collectingDateMax="2017-06-29" collectingDateMin="2017-06-25" collectionCode="MACN" collectorName="E Crescent City &amp; M. Ramirez &amp; P. Michalik" county="Del Norte Co." elevation="150" location="Ar" specimenCount="3" specimenCount-female="2" specimenCount-immature="1" stateProvince="California">
<specimenCount id="C7C1DE9A4EB4AA306ED71B3551879CCE" count="2" type="female">2 ♀</specimenCount>
(together with
<specimenCount id="216F0B4DAC04379E828143A0EA534FE9" count="1" type="immature">1 immature</specimenCount>
); same data as holotype;
<abbrev id="ABBRID0ERDAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="189B88398A4126652F9951A2FEDB9A4C" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
-
<location id="7C6E5ACFA776FAAE54CE11EDB8284EEB" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A:7C6E5ACFA776FAAE54CE11EDB8284EEB" name="Ar">Ar</location>
38386
</materialsCitation>
; •
<materialsCitation id="4E3814E66A1170D84DA0CDCFF1FAF610" collectingDate="2024-07-24" collectionCode="GPS" collectorName="M. Hedin &amp; O. Hedin" location="South Fork Smith River" municipality="Hiouchi" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="8977873C08EBF9C8E30FA435E3CF6DAD" count="1" type="female">1 ♀</specimenCount>
, several immatures; southeast of
<collectingMunicipality id="63FDCFA1FFA71746C8C73F9F986F6072">Hiouchi</collectingMunicipality>
, along
<location id="5CB6A7F7A3F9574D07AF60B11A410556" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A:5CB6A7F7A3F9574D07AF60B11A410556" municipality="Hiouchi" name="South Fork Smith River" stateProvince="California">South Fork Smith River</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYDAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-124.010000,41.760000]}">41.76, - 124.01</named-content>
(
<collectionCode id="65339ADB78E0852CEDDEC863442638C7">GPS</collectionCode>
, ±
<quantity id="F788015FE55A16F1A26BCD924E7BA5F9" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
);
<date id="1643F566CC75043DE13FBEA88E779C05" value="2024-07-24">
<collectingDate id="3B7B1044A41C6CDED9DCD571B35ADBEF" value="2024-07-24">24 Jul. 2024</collectingDate>
</date>
; N-facing rockpile, mixed redwood forest; coll.
<collectorName id="74B10C7F013D87087B51FB5756E5B553">M. Hedin</collectorName>
&amp;
<collectorName id="FA5EB170FEAFD0BBFF17D509D518930D">O. Hedin</collectorName>
;
<emphasis id="1E227F1A1EF3E373157B19D766FF1481" bold="true">
<collectingRegion id="B21ED5FACDDC54705F178A572DDC4DB9" country="United States of America" name="California">California</collectingRegion>
, Humboldt Co.
</emphasis>
</materialsCitation>
<materialsCitation id="9F1F20A2B5656230C7FC9A58AF7BE985" collectingDate="2017-06-26" collectionCode="MACN" collectorName="M. Ramirez &amp; P. Michalik &amp; Ar" elevation="120" latitude="41.01914" location="Tish Tang Campground" longLatPrecision="1" longitude="-123.63594" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="657C404BB994FB54F742871AEFBD1005" count="1" type="female">1 ♀</specimenCount>
; nr.
<location id="651641B89FEB1B688EFA4C1953DF8312" LSID="urn:lsid:plazi:treatment:514AFCE05B2A5ABE8B0BD2F27531210A:651641B89FEB1B688EFA4C1953DF8312" latitude="41.01914" longLatPrecision="1" longitude="-123.63594" name="Tish Tang Campground" stateProvince="California">Tish Tang Campground</location>
,
<locationDeviation id="69A5D088A18A5E5BA20B84EFB74ED850" location="Hoopa">SE of Hoopa</locationDeviation>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECEAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-123.635940,41.019140]}">
<geoCoordinate id="39AD0BB2B32ACE4F89BBFF6B42E3F2A7" degrees="41.01914" direction="north" orientation="latitude" precision="1" value="41.01914">41.01914</geoCoordinate>
,
<geoCoordinate id="38CBB90586BD100EAC2A02B4A0057A6C" degrees="123.63594" direction="west" orientation="longitude" precision="1" value="-123.63594">- 123.63594</geoCoordinate>
</named-content>
; elev.
<quantity id="8E8735935581E976A96D8D6F562D3A17" metricMagnitude="2" metricUnit="m" metricValue="1.2" unit="m" value="120.0">
<elevation id="D997A1BF2D06B095770BB8A59A0958EC" metricMagnitude="2" metricUnit="m" metricValue="1.2" unit="m" value="120.0">120 m</elevation>
</quantity>
;
<date id="A67D69CC517556601FADAC74C626D328" value="2017-06-26">
<collectingDate id="634DFD60E3B2EA28AE210C8B4E03D1B6" value="2017-06-26">26 Jun. 2017</collectingDate>
</date>
; mixed broadleaf forest; coll.
<collectorName id="38AAB6FE59CDC2A8786BE2F2DD2A10C6">M. Ramírez</collectorName>
&amp;
<collectorName id="C9810A00D15E75A3077B997E157E250F">P. Michalik</collectorName>
;
<abbrev id="ABBRID0EHEAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="4992F5DB9B4B48DC8744C5D78FD1DE53" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
-
<collectorName id="AB1AFECEA4B220C1476E95BB7E407118">Ar</collectorName>
38673
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELEAI" type="etymology">
<paragraph id="7E4E98F674B3530A479CA08128517503">
<heading id="CBC156EC100F0B5BC5B0496C9FAC4EDA" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="7839E22586C616CC3BC40C88D0B773D7">
<emphasis id="88B83FAA31B4B03BA6ECD9EA4D9E0FF0" italics="true">furtiva</emphasis>
(L., hidden, concealed), from the rarity, and microhabitat preference, of this species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2EAI" type="diagnosis">
<paragraph id="DBE389B5AB616BE3F464EBBB58244F08">
<heading id="4446678A1EB8579FCBB262A94F33864F" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="DF1BCF192650E33B51EA9A1F374D5A21">
The epigynum of
<taxonomicName id="FC2D37C8F9A85F48C8894E855601F259" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="furtiva" status="sp. nov.">
<emphasis id="B80B34F0875F64E41EA0BA37D9D0782F" italics="true">C. furtiva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C097E9193E70A0D99EA94681F729E70C" rank="species">sp. nov.</taxonomicNameLabel>
differs from that of
<taxonomicName id="CBA8E0970FA8CAF5F0719201E48FF666" authority="Chamberlin &amp; Ivie (1937)" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="heterops">
<emphasis id="527AA701BF36E3AD02BC425F667757B0" italics="true">C. heterops</emphasis>
<bibRefCitation id="2550C563E093331AA9CB265CC60608D0" DOI="10.1093/aesa/30.2.211" author="Chamberlin RV &amp; Ivie W" issue="2" journalOrPublisher="Annals of the Entomological Society of America" pagination="211-241" refId="B16" refString="Chamberlin RV, Ivie W (1937) New spiders of the family Agelenidae from western North America. Annals of the Entomological Society of America 30 (2): 211-241. https://doi.org/10.1093/aesa/30.2.211" title="New spiders of the family Agelenidae from western North America." volume="30" year="1937">Chamberlin &amp; Ivie (1937)</bibRefCitation>
</taxonomicName>
in possessing a slightly more sclerotized and rugous epigynal plate anterior to the epigastric furrow, with white (glandular?) material lying anterior to the spermathecae, and longer spermathecae that nearly meet at the midline.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXFAI" type="description">
<paragraph id="88EA9CA748CE9BDB93F65CCADE073985">
<heading id="C8BAF8DC46BE3E6CC39E496206E29E7A" reason="title">
Description of
<specimenCount id="FC0494C97604EC0E36E3BB3619F42643" count="1" type="female"></specimenCount>
<typeStatus id="BC2DBCEA3C1C16E2DD45BB20220D38C1" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="6285706ACD1A1BE3807C3A38E660D51C">
(
<collectionCode id="094618F18A6B2021976DE3066FBE28EF" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_ TAC 000889; Figs
<figureCitation id="91FFB37548E9DCFA8878A590A5E6C872" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cybaeozyga furtiva sp. nov. ♀ (A MACN - Ar 38880, B H holotype) A live specimen MACN - Ar 38880 B dorsal view C ventral view D prosoma dorsal view E prosoma lateral view F prosoma anterior view G eyes dorsal view H eyes anterior view." figureDoi="10.3897/zookeys.1226.140204.figure8" httpUri="https://binary.pensoft.net/fig/1243097">8 B H</figureCitation>
,
<figureCitation id="D864021482AC81F9DFDC4E343382E5DA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cybaeozyga furtiva sp. nov. ♀ (A, J M holotype B I paratype MACN - Ar 46970) A patellae and tibiae I, arrows to fracture lines B patellae III IV fractured, arrows to fracture lines C left leg I, prolateral view D same, retrolateral view E tarsus I prolateral view F tarsus IV prolateral view G tibia I ventral view H metatarsus I ventral view I left leg IV prolateral view J spinnerets ventral view K epigyne ventral view L cleared epigyne ventral view M cleared epigyne dorsal view. Abbreviations: BG = Bennetts gland, FD = fertilization duct, PCD = proximal copulatory duct, S = spermatheca." figureDoi="10.3897/zookeys.1226.140204.figure9" httpUri="https://binary.pensoft.net/fig/1243098">9 A, J M</figureCitation>
). Color in alcohol greenish brown with dark gray dorsal pattern. Carapace with dark markings extending from eyes to cephalic area, and sides of cephalic area; abdomen grayish with dark pattern dorsally, laterally and around spinnerets. Sternum unmarked. Legs darker at distal femora, basal tibiae, and basal metatarsi. Cheliceral promargin with three teeth (two basal together, one distal parted), retromargin with seven very small teeth.
</paragraph>
<caption id="E361B3C0B262FE226B0F0225449D8B28" ID-DOI="10.3897/zookeys.1226.140204.figure8" ID-arpha="76347A6C-9156-5E2C-B09D-B81985BBA865" httpUri="https://binary.pensoft.net/fig/1243097" startId="F8">
<paragraph id="E1D4B6D007590F894F90EE36A429B31B">
<label id="931D6EE7CD50CF704DAE4C7B632EE157">Figure 8.</label>
</paragraph>
<paragraph id="F79F08095096C6C7DD653C9E623A31AC">
<taxonomicName id="17BB1F0A68FB4F4CA5427E842412B5A7" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="furtiva" status="sp. nov.">
<emphasis id="10401A8EF721F872B2763C4C1EE1848C" italics="true">Cybaeozyga furtiva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC015A98421B6E65E0454B49EA73D08" rank="species">sp. nov.</taxonomicNameLabel>
♀ (A
<abbrev id="ABBRID0EYGAI" xlink_title="Museo Argentino de Ciencias Naturales">MACN</abbrev>
- Ar 38880, B H holotype)
<emphasis id="3BB7AE5F636BC502441E3642488739BE" bold="true">A</emphasis>
live specimen
<abbrev id="ABBRID0E5GAI" xlink_title="Museo Argentino de Ciencias Naturales">MACN</abbrev>
- Ar 38880
<emphasis id="9F2091E73ED62C9A8275C5320BE57F22" bold="true">B</emphasis>
dorsal view
<emphasis id="C2576D81E0373D9E04E028F28A675A6B" bold="true">C</emphasis>
ventral view
<emphasis id="07C2BAB006A69920F56BBE7C5874A3D1" bold="true">D</emphasis>
prosoma dorsal view
<emphasis id="492AA67A2A5C8FE2D6CCC1FECAAF0E37" bold="true">E</emphasis>
prosoma lateral view
<emphasis id="13C2E13D30F8FEC421DF553DA6EA5908" bold="true">F</emphasis>
prosoma anterior view
<emphasis id="901AF0D2469E179C4771D02D66F92C68" bold="true">G</emphasis>
eyes dorsal view
<emphasis id="312E697472112E227E6AC6F56210D359" bold="true">H</emphasis>
eyes anterior view.
</paragraph>
</caption>
<caption id="00408D9FC1269CC06F7C8617A9ED7D68" ID-DOI="10.3897/zookeys.1226.140204.figure9" ID-arpha="3E263464-B8C0-5B49-B198-2F43986311DA" httpUri="https://binary.pensoft.net/fig/1243098" startId="F9">
<paragraph id="C037D0E53F314EE93D894C16E0675458">
<label id="A99EADA1F62CB8EB192A49E65D0A8432">Figure 9.</label>
</paragraph>
<paragraph id="F5A08224813C3DBD3AA349F24E27DA94">
<taxonomicName id="DE55C0EAC85C5C3C27B0245924CCAAFB" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="furtiva" status="sp. nov.">
<emphasis id="8E4265E05BC6F4C87D0728ACC38A28AD" italics="true">Cybaeozyga furtiva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F427AFB7992558D755AA448CEACDD24" rank="species">sp. nov.</taxonomicNameLabel>
♀ (
<emphasis id="DCDEF03F7359BDB81252CCCBEAB42EFA" bold="true">A, J M</emphasis>
holotype
<emphasis id="2052971F19103B0C8671202FFC771C6B" bold="true">B I</emphasis>
paratype
<abbrev id="ABBRID0EMIAI" xlink_title="Museo Argentino de Ciencias Naturales">MACN</abbrev>
- Ar 46970)
<emphasis id="78E5C6A123FFD6B5D564C69507C48E31" bold="true">A</emphasis>
patellae and tibiae I, arrows to fracture lines
<emphasis id="3C2FB8F67BE5696E2B4F3C2B8E25F3A8" bold="true">B</emphasis>
patellae III IV fractured, arrows to fracture lines
<emphasis id="52A97711AB9F6758188F067287219E54" bold="true">C</emphasis>
left leg I, prolateral view
<emphasis id="FF328DF02BB05C3A797C40BB03FDA3D4" bold="true">D</emphasis>
same, retrolateral view
<emphasis id="29FF38F1B3F10E472099E18A317D6492" bold="true">E</emphasis>
tarsus I prolateral view
<emphasis id="A7D228218318F0827B50707461204D6C" bold="true">F</emphasis>
tarsus IV prolateral view
<emphasis id="72064D334E74A36C692A7B9782575128" bold="true">G</emphasis>
tibia I ventral view
<emphasis id="A8FB1A908CEABB087564713AF3B52432" bold="true">H</emphasis>
metatarsus I ventral view
<emphasis id="6F7EFDEC9DEE5D1BED18D8043FAA96DF" bold="true">I</emphasis>
left leg IV prolateral view
<emphasis id="11C0B0E0F1FAB368BE10C75AA64CE7A5" bold="true">J</emphasis>
spinnerets ventral view
<emphasis id="5DD615846F9C94DC64EB01FA10831F89" bold="true">K</emphasis>
epigyne ventral view
<emphasis id="C6EB80DA7F0BA24354EA3B2AFFEEE26C" bold="true">L</emphasis>
cleared epigyne ventral view
<emphasis id="FC8320F7E5EF083BD9AD6E562CDCD122" bold="true">M</emphasis>
cleared epigyne dorsal view. Abbreviations: BG = Bennetts gland, FD = fertilization duct, PCD = proximal copulatory duct, S = spermatheca.
</paragraph>
</caption>
<paragraph id="0BFB7753180611E13FB56458181A22B9">
Total length 3.53, carapace length 1.40, carapace width 1.05, cephalic region width 0.60, posterior eye row width 0.45. Anterior median eyes missing, represented by small dark blotches of black pigment. Eye diameters ALE: PLE: PME = 0.09: 0.08: 0.08. Sternum length 0.70, sternum width 0.67. Leg formula 1423. Leg I article lengths (1.27, 0.49, 1.08, 1.05, 0.75 = 4.64) leg
<collectionCode id="207376F8B2377DAF18447FD9ECCA3789">IV</collectionCode>
article lengths (1.37, 0.45, 1.25, 2.67, 0.90 = 6.64). Abdomen 2.17 long.
</paragraph>
<paragraph id="1DDBA9D4F4C6210716B020708634413E">
Leg macrosetae (Fig.
<figureCitation id="35D59880B8F4F0939995BAEAB8B476B3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cybaeozyga furtiva sp. nov. ♀ (A, J M holotype B I paratype MACN - Ar 46970) A patellae and tibiae I, arrows to fracture lines B patellae III IV fractured, arrows to fracture lines C left leg I, prolateral view D same, retrolateral view E tarsus I prolateral view F tarsus IV prolateral view G tibia I ventral view H metatarsus I ventral view I left leg IV prolateral view J spinnerets ventral view K epigyne ventral view L cleared epigyne ventral view M cleared epigyne dorsal view. Abbreviations: BG = Bennetts gland, FD = fertilization duct, PCD = proximal copulatory duct, S = spermatheca." figureDoi="10.3897/zookeys.1226.140204.figure9" httpUri="https://binary.pensoft.net/fig/1243098">9 C I</figureCitation>
,
<typeStatus id="E567F11FF4A5FADEA086CA00ADDEF22C" type="paratype">paratype</typeStatus>
<abbrev id="ABBRID0EWJAI" xlink_title="Museo Argentino de Ciencias Naturales">
<collectionCode id="BB1502C20A336D64933A74636449E6D1" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</abbrev>
- Ar 46970) absent on femora, 1-1 on patellae (dorsal), thick on legs I
<collectionCode id="3910EBFFCD63C98BF6852828F55D5D63">III</collectionCode>
, weaker on
<collectionCode id="28CE173083C1BF0F5388AAFD254BC867">IV</collectionCode>
. Tibia I ventral 2-2 - 0, prolateral 1-1 - 0, dorsal 1-0 weaker; metatarsus I prolateral 1-1 - 1 - 1, ventral 2-2 - 2 - 0 not well paired. Tibia
<collectionCode id="77ADFA6BBD0F97B8EA815C4A227DB8D1">IV</collectionCode>
prolateral 1-1, ventral 1-1, retrolateral 0-1, dorsal 1-0 - 1; metatarsus
<collectionCode id="4EE9ED7D2D7E21BBDCE8BB5BC09F8B55">IV</collectionCode>
prolateral 1-1, ventral 0-1 - 1, retrolateral 0-0 - 2. Patellae with basal fracture region seen as darker indentations at the sides (Fig.
<figureCitation id="A7FB18154DFEF8694C78FDC91E6FE356" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cybaeozyga furtiva sp. nov. ♀ (A, J M holotype B I paratype MACN - Ar 46970) A patellae and tibiae I, arrows to fracture lines B patellae III IV fractured, arrows to fracture lines C left leg I, prolateral view D same, retrolateral view E tarsus I prolateral view F tarsus IV prolateral view G tibia I ventral view H metatarsus I ventral view I left leg IV prolateral view J spinnerets ventral view K epigyne ventral view L cleared epigyne ventral view M cleared epigyne dorsal view. Abbreviations: BG = Bennetts gland, FD = fertilization duct, PCD = proximal copulatory duct, S = spermatheca." figureDoi="10.3897/zookeys.1226.140204.figure9" httpUri="https://binary.pensoft.net/fig/1243098">9 A, B</figureCitation>
).
</paragraph>
<paragraph id="09A0EAB7ED7F23ED8571EDE2B40BEB54">
Spinnerets short, colulus a hairy patch (Fig.
<figureCitation id="4DD7E04AC2D5DB4B8F765B61F1677607" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cybaeozyga furtiva sp. nov. ♀ (A, J M holotype B I paratype MACN - Ar 46970) A patellae and tibiae I, arrows to fracture lines B patellae III IV fractured, arrows to fracture lines C left leg I, prolateral view D same, retrolateral view E tarsus I prolateral view F tarsus IV prolateral view G tibia I ventral view H metatarsus I ventral view I left leg IV prolateral view J spinnerets ventral view K epigyne ventral view L cleared epigyne ventral view M cleared epigyne dorsal view. Abbreviations: BG = Bennetts gland, FD = fertilization duct, PCD = proximal copulatory duct, S = spermatheca." figureDoi="10.3897/zookeys.1226.140204.figure9" httpUri="https://binary.pensoft.net/fig/1243098">9 J</figureCitation>
). Epigynal plate oval with sinuous transversal ridge and rugous median area (Fig.
<figureCitation id="965638BA8CC518E6E89D99498E2EBCD4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cybaeozyga furtiva sp. nov. ♀ (A, J M holotype B I paratype MACN - Ar 46970) A patellae and tibiae I, arrows to fracture lines B patellae III IV fractured, arrows to fracture lines C left leg I, prolateral view D same, retrolateral view E tarsus I prolateral view F tarsus IV prolateral view G tibia I ventral view H metatarsus I ventral view I left leg IV prolateral view J spinnerets ventral view K epigyne ventral view L cleared epigyne ventral view M cleared epigyne dorsal view. Abbreviations: BG = Bennetts gland, FD = fertilization duct, PCD = proximal copulatory duct, S = spermatheca." figureDoi="10.3897/zookeys.1226.140204.figure9" httpUri="https://binary.pensoft.net/fig/1243098">9 K, L</figureCitation>
). Wide proximal copulatory ducts filled with whitish material (Fig.
<figureCitation id="CCD3592B9097BDE75C1BD05043DE4736" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cybaeozyga furtiva sp. nov. ♀ (A, J M holotype B I paratype MACN - Ar 46970) A patellae and tibiae I, arrows to fracture lines B patellae III IV fractured, arrows to fracture lines C left leg I, prolateral view D same, retrolateral view E tarsus I prolateral view F tarsus IV prolateral view G tibia I ventral view H metatarsus I ventral view I left leg IV prolateral view J spinnerets ventral view K epigyne ventral view L cleared epigyne ventral view M cleared epigyne dorsal view. Abbreviations: BG = Bennetts gland, FD = fertilization duct, PCD = proximal copulatory duct, S = spermatheca." figureDoi="10.3897/zookeys.1226.140204.figure9" httpUri="https://binary.pensoft.net/fig/1243098">9 K</figureCitation>
), spermathecae posterior, transversal. Copulatory openings not seen, probably in the anterior ridges and leading to the wide, soft copulatory ducts and from there to posterior spermathecae. Bennetts gland large, on ectal side of spermatheca (Fig.
<figureCitation id="8CA872CDF455AB9E229E0CE87467AC15" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cybaeozyga furtiva sp. nov. ♀ (A, J M holotype B I paratype MACN - Ar 46970) A patellae and tibiae I, arrows to fracture lines B patellae III IV fractured, arrows to fracture lines C left leg I, prolateral view D same, retrolateral view E tarsus I prolateral view F tarsus IV prolateral view G tibia I ventral view H metatarsus I ventral view I left leg IV prolateral view J spinnerets ventral view K epigyne ventral view L cleared epigyne ventral view M cleared epigyne dorsal view. Abbreviations: BG = Bennetts gland, FD = fertilization duct, PCD = proximal copulatory duct, S = spermatheca." figureDoi="10.3897/zookeys.1226.140204.figure9" httpUri="https://binary.pensoft.net/fig/1243098">9 M</figureCitation>
), fertilization ducts posteriorly placed.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQKAI" type="variation">
<paragraph id="CF7D72BD1037AB63EA2F090680ECD8D9">
<heading id="52332FAE91D872028CC6AD0963B9CC11" reason="title">Variation.</heading>
</paragraph>
<paragraph id="5604F115C413028E2C1E3B2D43D4D231">
The black pigment replacing the missing anterior eyes is variable in location between and below the ALEs, and often asymmetrical (Fig.
<figureCitation id="5E1EE31CDD3444E4DC4FB2985F8484C4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cybaeozyga furtiva sp. nov. ♀ (A MACN - Ar 38880, B H holotype) A live specimen MACN - Ar 38880 B dorsal view C ventral view D prosoma dorsal view E prosoma lateral view F prosoma anterior view G eyes dorsal view H eyes anterior view." figureDoi="10.3897/zookeys.1226.140204.figure8" httpUri="https://binary.pensoft.net/fig/1243097">8 H</figureCitation>
). Epigyna from the two geographically distant locations (Fig.
<figureCitation id="D3179CE788710BDE99D80FAB04D3D1AD" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Species distributions of Siskiyu armilla sp. nov. and Cybaeozyga, including C. furtiva sp. nov. and C. heterops. Locations approximate, see text and Suppl. material 1 for precise location data." figureDoi="10.3897/zookeys.1226.140204.figure10" httpUri="https://binary.pensoft.net/fig/1243099">10</figureCitation>
) are very similar in detail.
</paragraph>
<caption id="00881F514A64958319B06DF8276D85B4" ID-DOI="10.3897/zookeys.1226.140204.figure10" ID-arpha="5E3615BB-7F77-5751-B15E-E7EA13716ACA" httpUri="https://binary.pensoft.net/fig/1243099" startId="F10">
<paragraph id="DDC1E0B00FC791BE2D85D9A558CF4A52">
<label id="BA85F3AF73C3EDF46B2CCB2D3B59E7EE">Figure 10.</label>
</paragraph>
<paragraph id="9436661B3C6C9D555F676396CBAB6EDC">
Species distributions of
<taxonomicName id="0E39D7BCEA202F69DC81DE616237186F" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Siskiyu" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="armilla" status="sp. nov.">
<emphasis id="AFD26860B6FC4370F94A006AF8C3134F" italics="true">Siskiyu armilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EA19CF3C4BC3690052078791B4F10A6" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="95E9BEFDE7B3AE3A1B1B8A721EEE4888" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="EA30746B95055AA2401ED284439B8230" italics="true">Cybaeozyga</emphasis>
</taxonomicName>
, including
<taxonomicName id="B4A3F787B302A035F55C80913A6E34A5" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="furtiva" status="sp. nov.">
<emphasis id="B94096776C17771DB147C09A940C7235" italics="true">C. furtiva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7C347A2CD36C434F4AE370DA773239CA" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="095E6C09BF9265EEC385FC032C57496D" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="heterops">
<emphasis id="3A825D4D7F24C1050E23E1A9F596C384" italics="true">C. heterops</emphasis>
</taxonomicName>
. Locations approximate, see text and Suppl. material
<xref id="0158605DA8230EEA1BD7DDF331F02379" ref-type="supplementary-material" rid="S1">1</xref>
for precise location data.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EXMAI" type="distribution">
<paragraph id="A312B1834725581C43A48467E787DC42">
<heading id="875B062EE0FB2A9B118BDB7AA88782D1" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="CD45F437FCF518E441FB3D2EB9A3F6C2">
Confirmed specimens from Del Norte and Humboldt counties,
<collectingRegion id="85607BEADC7E1D313B0A1C594C3A5D8A" country="United States of America" name="California">California</collectingRegion>
(Fig.
<figureCitation id="BB232F74ED2EA57B396614559285FB4B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Species distributions of Siskiyu armilla sp. nov. and Cybaeozyga, including C. furtiva sp. nov. and C. heterops. Locations approximate, see text and Suppl. material 1 for precise location data." figureDoi="10.3897/zookeys.1226.140204.figure10" httpUri="https://binary.pensoft.net/fig/1243099">10</figureCitation>
), at lower elevations (
<quantity id="78AE6FA6B5E3DE9936E2A4410A86D209" metricMagnitude="2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" unit="m" value="135.0" valueMax="150.0" valueMin="120.0">120150 m</quantity>
). Topotypic specimens were abundantly collected under rock piles or logs, and in leaf litter, in wet mixed conifer forest. Males were only represented as penultimates. Robb Bennett (pers. comm.) has examined female specimens from Siskiyou County (southeast of Seiad Valley, Fig.
<figureCitation id="7B38849D0BD9C24A46DB58183150BEB6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Species distributions of Siskiyu armilla sp. nov. and Cybaeozyga, including C. furtiva sp. nov. and C. heterops. Locations approximate, see text and Suppl. material 1 for precise location data." figureDoi="10.3897/zookeys.1226.140204.figure10" httpUri="https://binary.pensoft.net/fig/1243099">10</figureCitation>
), currently housed in the American Museum of Natural History, that appear to match this species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFNAI" type="comments">
<paragraph id="AD0EFF4D90109B7049AEB3E3EE8248D7">
<heading id="91CDE112A7913520AF6D1CA6F5F96E68" reason="title">Comments.</heading>
</paragraph>
<paragraph id="EB6240A9BA9BC82C53823E3C8E04D64D">
This new species adds to the described species diversity of
<taxonomicName id="B552293C9ECD11EA7B48A626358654B7" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="E5EE4F3CD627BE528B71BE5EE4984149" italics="true">Cybaeozyga</emphasis>
</taxonomicName>
, a notoriously poorly known genus.
<bibRefCitation id="1D0825C8142F8B35E8DE0422CF68E797" DOI="10.1093/aesa/30.2.211" author="Chamberlin RV &amp; Ivie W" firstAuthor="Chamberlin" issue="2" journalOrPublisher="Annals of the Entomological Society of America" pagination="211-241" refId="B16" refString="Chamberlin RV, Ivie W (1937) New spiders of the family Agelenidae from western North America. Annals of the Entomological Society of America 30 (2): 211241. https://doi.org/10.1093/aesa/30.2.211" title="New spiders of the family Agelenidae from western North America." volume="30" year="1937">Chamberlin and Ivie (1937)</bibRefCitation>
described
<taxonomicName id="097BC305DF09CF7F34D7A42D68C0EA85" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="F0D2338BCF6A8504BCCCF95356BC770F" italics="true">Cybaeozyga</emphasis>
</taxonomicName>
and the then only known species (
<taxonomicName id="23C98CB7B13C8F679C3E685AC24CE990" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="heterops">
<emphasis id="20D1F799000A0A7339DD0C29678859E2" italics="true">C. heterops</emphasis>
</taxonomicName>
) from a single male specimen, citing the type locality as “ Grave Creek,
<collectingRegion id="657931F9553437EBC619F3CB0C8BEC5B" country="United States of America" name="Oregon">Oregon</collectingRegion>
(
<emphasis id="1D17EAFD3BC08B3591468A56A71FEDA9" italics="true">near Klamath Falls</emphasis>
). ” The authors did not include county information in their locality data. Later,
<bibRefCitation id="563CB693A4B75DC02AD7ED38E281F44D" author="Roth VD &amp; Brame PL" firstAuthor="Roth" refId="B56" refString="Roth VD, Brame PL (1972) Nearctic genera of the spider family Agelenidae (Arachnida, Araneida). American Museum Novitates: 2505. http://hdl.handle.net/2246/2712" url="http://hdl.handle.net/2246/2712" year="1972">Roth and Brame (1972)</bibRefCitation>
provided illustrations for a male
<taxonomicName id="18BAD15515376BD25B6ABB10740C6D8F" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="heterops">
<emphasis id="C1C1E89EFB8495CFB5D0CAAEBADA4C6D" italics="true">C. heterops</emphasis>
</taxonomicName>
, matching the
<typeStatus id="374AF0F5F99D0BC5E0D9A39767CCBB93" type="holotype">holotype</typeStatus>
description, and the previously undescribed female, presumably from the same location. Although precise locality data were not provided in this publication, we have seen specimens of
<taxonomicName id="016F86AEC2C7B24AAD4E8C3D38007C16" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="464F51A76A7B1AD6EC6D44D0994F9F4A" italics="true">Cybaeozyga</emphasis>
</taxonomicName>
from Grave Creek, Josephine County, OR collected by Roth (M. Hedin, pers. obs.). Our female specimens of
<taxonomicName id="EFDE12625155D2DD54A63C32CE73CCF6" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="heterops">
<emphasis id="1EF7CFF84A6950A8CF56581E648562D5" italics="true">C. heterops</emphasis>
</taxonomicName>
match closely the epigynal drawings of
<bibRefCitation id="60ED65A7B78A0C24EBFD4B872CECDD56" author="Roth and Brame" firstAuthor="Roth" refId="B56" refString="Roth VD, Brame PL (1972) Nearctic genera of the spider family Agelenidae (Arachnida, Araneida). American Museum Novitates: 2505. http://hdl.handle.net/2246/2712" url="http://hdl.handle.net/2246/2712" year="1972">Roth and Brame (1972</bibRefCitation>
: fig. 25), and originate from Grave Creek, near confluence with Butte Creek, ~
<quantity id="31C0E9233E6567546D974F77150B3140" metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
NNW of Grants Pass, Josephine County, OR (Suppl. material
<xref id="BC91721FD985FD4FD4AAD2E59AF7423F" ref-type="supplementary-material" rid="S1">1</xref>
). We view this as near the probable type locality for this species (Fig.
<figureCitation id="A12B77314671CA8FEF25598816A0E631" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Species distributions of Siskiyu armilla sp. nov. and Cybaeozyga, including C. furtiva sp. nov. and C. heterops. Locations approximate, see text and Suppl. material 1 for precise location data." figureDoi="10.3897/zookeys.1226.140204.figure10" httpUri="https://binary.pensoft.net/fig/1243099">10</figureCitation>
); this location is approximately
<quantity id="4C507AA9F557BD40AB0B893514433B08" metricMagnitude="5" metricUnit="m" metricValue="2.0" unit="km" value="200.0">200 km</quantity>
NW of Klamath Falls,
<collectingRegion id="0EB320D88139E49779B338474F4E9D42" country="United States of America" name="Oregon">Oregon</collectingRegion>
.
</paragraph>
<paragraph id="E91C2C7F4506499848710D707169C4C7">
Additional known, but still undescribed species of
<taxonomicName id="6E547DCBC72FA940E40A05963D2C3AC6" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="05C3FC95226AC8E7A31254E9CEEE6288" italics="true">Cybaeozyga</emphasis>
</taxonomicName>
have been mentioned in the literature.
<bibRefCitation id="48618E8AB830CE48CF3D992E169E999B" author="Roth VD &amp; Brame PL" firstAuthor="Roth" refId="B56" refString="Roth VD, Brame PL (1972) Nearctic genera of the spider family Agelenidae (Arachnida, Araneida). American Museum Novitates: 2505. http://hdl.handle.net/2246/2712" url="http://hdl.handle.net/2246/2712" year="1972">Roth and Brame (1972)</bibRefCitation>
note three undescribed species from caves in northwestern
<collectingRegion id="2524480E3B7F7A3993FB830A410F7CB5" country="United States of America" name="California">California</collectingRegion>
(see also
<bibRefCitation id="3FF163BD1B8F6F27FBCE8FDA69FCB67A" author="Bennett" etAl="et al." firstAuthor="Bennett" refId="B5" refString="Bennett RG, Ubick D, Paquin P, Cushing PE, Roth V (2017) Cybaeidae. Spiders of North America: an identification manual, Second Edition. American Arachnological Society, 96101." year="2017">Bennett et al. 2017</bibRefCitation>
). Overall, the distribution of
<taxonomicName id="3423FCD95D00D944D9D86137CB243CFF" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="908B9C6AFCED879122671C7058E8C8AE" italics="true">Cybaeozyga</emphasis>
</taxonomicName>
appears to include forests and caves of the Klamath Mountains ecoregion of northwestern
<collectingRegion id="156BAFDAF84FD1EE1119CE98CF0BB608" country="United States of America" name="California">California</collectingRegion>
and southwestern
<collectingRegion id="FDA263063388B0ACC1B769F31506D8C2" country="United States of America" name="Oregon">Oregon</collectingRegion>
(Fig.
<figureCitation id="2AA9B5F6C92A5765336E15C8785CB113" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Species distributions of Siskiyu armilla sp. nov. and Cybaeozyga, including C. furtiva sp. nov. and C. heterops. Locations approximate, see text and Suppl. material 1 for precise location data." figureDoi="10.3897/zookeys.1226.140204.figure10" httpUri="https://binary.pensoft.net/fig/1243099">10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,589 @@
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<mods:title id="FDF2107E4F5AE99D588DE9329A420F68">Mutatus gen. nov., a new genus of Phrurolithidae (Arachnida, Araneae) from China, with descriptions of four new species</mods:title>
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<mods:namePart id="EE2110E3F3388F69B50EB54796582ECF">Mu, Yannan</mods:namePart>
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<mods:affiliation id="C33048B57D15230B5340F77B0FC53B22">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China &amp; Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="E63FE27CB558EB9924B8DC8AA2310DE1">Wang, Luyu</mods:namePart>
<mods:nameIdentifier id="46387B6BCD506170E4714F529ABE8021" type="ORCID">0000-0002-5250-3473</mods:nameIdentifier>
<mods:affiliation id="543F2ACCAEA440CF7BEE8461F94A29AB">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:namePart id="380C45E5EBA5CCE259E36D9BE5D0E6B5">Zhang, Feng</mods:namePart>
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<mods:affiliation id="3A74AC295D77845F1FC026E4F1EA3046">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="189C1ABFB7457252C59E0EC1814CACA6">Zhang, Zhisheng</mods:namePart>
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<mods:affiliation id="66D9E0ED3B345FE2220B663603DEFB00">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<subSubSection id="A437871B6B99BAAE7D8798865341C6EC" type="nomenclature">
<paragraph id="9BAE9ED46DD0BF4EDB20359661D42812">
<taxonomicName id="CD270DA590D247E8945E57AAEFF481FA" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Phrurolithidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianyan" status="sp. nov.">
<emphasis id="E99B65CC1A8B1652D0BDCFA356C9F129" italics="true">Mutatus tianyan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35B7A741FD3A63EBC002BD6C00400066" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2DDA5806DE5AF297420E2F03BB9D402C" type="description">
<paragraph id="28DFCA9B915EBB4D8441860FA3366FAF">
<figureCitation id="0F98B6F3527E66A5E659729D139BD97F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Mutatus tianyan sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure5" httpUri="https://binary.pensoft.net/fig/1243804">Chinese name: 天燕幻蛛 Figs 5</figureCitation>
,
<figureCitation id="3D81E6597B3A751055EA59A6EAD2E452" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6</figureCitation>
,
<figureCitation id="1D250405F861F47435E967335054A1AF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution map of Mutatus gen. nov." figureDoi="10.3897/zookeys.1226.137752.figure8" httpUri="https://binary.pensoft.net/fig/1243807">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQ3AG" type="type material">
<paragraph id="7CE6D3AF424B0139F1FAD5229094B513">
<heading id="5614D2DEF671006D0D1ADB0BB755249E" reason="title">Type material.</heading>
</paragraph>
<paragraph id="E1B403AF59E417FCEC1AF4A11CECBA05">
<materialsCitation id="695915B19032CA3A5868AC1EDD7C856C" collectingDate="2023-06-14" collectorName="Z. S. Zhang &amp; Y. J. Cai" country="China" elevation="1871" latitude="31.721111" location="Tianyan Primitive Ecotourism Area" longLatPrecision="20" longitude="110.47528" municipality="Shennongjia Forest District" specimenCount="1" specimenCount-male="1" stateProvince="Hubei" typeStatus="holotype">
<emphasis id="21A0AA217E39A136E80B92BF680EA1FB" bold="true" italics="true">
<typeStatus id="556D6A8BF4ED557D23E4F0697D39D033" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="BDF2E4603FDE3EF1EC49C382244F3640" name="China">China</collectingCountry>
<specimenCount id="81E5E1D1392CA4CE19AC6A2A2AA931F2" count="1" type="male"></specimenCount>
<collectingRegion id="3FA6286038321E44A148FADFF476FFFE" country="China" name="Hubei">Hubei province</collectingRegion>
:
<collectingMunicipality id="19A12B186136AE4D784C5DDD3E522CD6">Shennongjia Forest District</collectingMunicipality>
,
<location id="DD88AE1B79CDFB855CC89398931DC7C5" LSID="urn:lsid:plazi:treatment:5B085250E23D5468B9FF2085D7785F85:DD88AE1B79CDFB855CC89398931DC7C5" country="China" latitude="31.721111" longLatPrecision="20" longitude="110.47528" municipality="Shennongjia Forest District" name="Tianyan Primitive Ecotourism Area" stateProvince="Hubei">Tianyan Primitive Ecotourism Area</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E53AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[110.475278,31.721111]}">
<geoCoordinate id="35CED03D7AA36A467539FD30AF032A97" degrees="31" direction="north" minutes="43" orientation="latitude" precision="15" seconds="16" value="31.721111">31 ° 43 ' 16 &quot; N</geoCoordinate>
,
<geoCoordinate id="05454960E4F31F3595B2EA58D90FC65C" degrees="110" direction="east" minutes="28" orientation="longitude" precision="15" seconds="31" value="110.47528">110 ° 28 ' 31 &quot; E</geoCoordinate>
</named-content>
, elev.
<quantity id="6E814794997E6A2D1CCAD4E003E268F3" metricMagnitude="3" metricUnit="m" metricValue="1.871" unit="m" value="1871.0">
<elevation id="5FA2675233124346CF0EE182CDA5B7BC" metricMagnitude="3" metricUnit="m" metricValue="1.871" unit="m" value="1871.0">1871 m</elevation>
</quantity>
,
<date id="D37BCA2B99E09818C5218EFB469F5D9C" value="2023-06-14">
<collectingDate id="83B52947ABC2FC6550E5AEEC976A713D" value="2023-06-14">14 June 2023</collectingDate>
</date>
,
<collectorName id="46AD654404874DCFA875DA6611FB6010">Z. S. Zhang</collectorName>
,
<collectorName id="7F65ED900588C7CD2B5EB0E3083ECAF2">Y. J. Cai</collectorName>
leg.
</materialsCitation>
<materialsCitation id="CF67F1E20E5DFF81D9D1D75F3C02252F" collectingDate="2023-06-14" collectorName="Z. S. Zhang &amp; Y. J. Cai" country="China" elevation="1871" latitude="31.721111" location="Tianyan Primitive Ecotourism Area" longLatPrecision="20" longitude="110.47528" municipality="Shennongjia Forest District" specimenCount="7" specimenCount-female="7" stateProvince="Hubei" typeStatus="paratype">
<emphasis id="7C9C53B3B2CB6ACAC19E58BCF50138F3" bold="true" italics="true">
<typeStatus id="69856FA1BCAED952316FDBEBAC008014" type="paratype">Paratype</typeStatus>
</emphasis>
: •
<specimenCount id="D8139CA6F8C2C94B7D4E7D641BEAB91D" count="7" type="female">7 ♀</specimenCount>
, with same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EG4AG" type="etymology">
<paragraph id="37D159E08969E3C61E0DC1B051E661FF">
<heading id="7423A1E681BF22320D193846CA4A71DE" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="31C8746B3D87048C8DB558D9C2798A90">
This specific name is derived from the
<typeStatus id="A217EB39C006599C7767C0146C0E5842">type</typeStatus>
locality; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EL4AG" type="diagnosis">
<paragraph id="B955AABCC200BD7DFCD074C46C274BAD">
<heading id="9DDE269BBE94A1007F446C5AF3A3F560" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="FC918DB250C25F91B65AEA215C6B286F">
See the diagnosis of
<taxonomicName id="F00E549E032B5901103FEC99717607D6" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tujia" status="sp. nov.">
<emphasis id="514F38B003636C465F0A9309B41CBB7E" italics="true">M. tujia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D5F02F7D365037FB8A54E30D52C8493" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E34AG" type="description">
<paragraph id="D5413C612590A0C541AD30A20D808390">
<heading id="8485C2CA7FCF133956DE67CC7DE52774" reason="title">Description.</heading>
</paragraph>
<paragraph id="0F2702A2005A19AE27FA479315182B75">
Male (
<typeStatus id="64C535FB2426F0E3C703259C2A1C8D8A" type="holotype">holotype</typeStatus>
, Fig.
<figureCitation id="B8718F665E76F5A01834BB66E7078970" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Mutatus tianyan sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure5" httpUri="https://binary.pensoft.net/fig/1243804">5 A, B</figureCitation>
). Total length 3.18, carapace 1.59 long, 1.35 wide; abdomen 1.59 long, 1.05 wide. Eye sizes and interdistances:
<abbrev id="ABBRID0EG5AG" xlink_title="anterior median eye">AME</abbrev>
0.07,
<abbrev id="ABBRID0EK5AG" xlink_title="anterior lateral eye">ALE</abbrev>
0.09,
<abbrev id="ABBRID0EO5AG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0ES5AG" xlink_title="posterior lateral eye">PLE</abbrev>
0.09;
<abbrev id="ABBRID0EW5AG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0E15AG" xlink_title="anterior median eye">AME</abbrev>
0.06,
<abbrev id="ABBRID0E55AG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0EC6AG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0EG6AG" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0EK6AG" xlink_title="anterior lateral eye">ALE</abbrev>
0.24,
<abbrev id="ABBRID0EO6AG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0ES6AG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EW6AG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E16AG" xlink_title="posterior lateral eye">PLE</abbrev>
0.08,
<abbrev id="ABBRID0E56AG" xlink_title="posterior lateral eye">PLE</abbrev>
<abbrev id="ABBRID0ECABG" xlink_title="posterior lateral eye">PLE</abbrev>
0.40,
<abbrev id="ABBRID0EGABG" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0EKABG" xlink_title="posterior lateral eye">PLE</abbrev>
0.06.
<abbrev id="ABBRID0EOABG" xlink_title="eye area width">EAW</abbrev>
0.52,
<abbrev id="ABBRID0ESABG" xlink_title="cephalic region width">CRW</abbrev>
0.74,
<abbrev id="ABBRID0EWABG" xlink_title="eye area width">EAW</abbrev>
/
<abbrev id="ABBRID0E1ABG" xlink_title="cephalic region width">CRW</abbrev>
0.70.
<abbrev id="ABBRID0E5ABG" xlink_title="cephalic region width">CRW</abbrev>
/
<abbrev id="ABBRID0ECBBG" xlink_title="carapace width">CW</abbrev>
0.55.
<abbrev id="ABBRID0EGBBG" xlink_title="median ocular area">MOA</abbrev>
0.22 long, anterior width 0.19, posterior width 0.25. Clypeal height 0.15,
<abbrev id="ABBRID0EKBBG" xlink_title="clypeal height">
<collectingCountry id="3613FEE2ACA3ED5326986D51ACFD3740" name="Switzerland">CH</collectingCountry>
</abbrev>
/
<abbrev id="ABBRID0EOBBG" xlink_title="anterior median eye">AME</abbrev>
2.14. Labium 0.21 long, 0.25 wide. Sternum 0.96 long, 0.91 wide. Carapace oval, cervical groove inconspicuous. Sternum, labium, and endite brown, without pattern. Abdomen brown, smooth, with large dorsal scutum covering almost entire abdomen. Legs brown.
</paragraph>
<caption id="1E7E8D67A07163390481871FCF2BA75F" ID-DOI="10.3897/zookeys.1226.137752.figure5" ID-arpha="650FAD39-E5A2-5F03-9275-96F6F28BE617" httpUri="https://binary.pensoft.net/fig/1243804" startId="F5">
<paragraph id="0BCC6275AE19E960E1864BA61983C109">
<label id="9BD453F99493462B5B592BF850BB93EF">Figure 5.</label>
</paragraph>
<paragraph id="C953B3517EEDD767E4740D4657137F46">
Habitus of
<taxonomicName id="54FB2BE20CCE125AEFF62B0E0C7AF32B" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tianyan" status="sp. nov.">
<emphasis id="1ED8FF1233A41A8AA42CCF5F5D5103D6" italics="true">Mutatus tianyan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4D924701B47044716962CF6E2C08B68" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1452B8606E448E3E2EE6BA80DD567F85" bold="true">A</emphasis>
male holotype, dorsal view
<emphasis id="EB7DF692FD589549FFE22AB78B084FF2" bold="true">B</emphasis>
same, ventral view
<emphasis id="443EC9E4112D22F2AB53EB21A0A10F29" bold="true">C</emphasis>
female paratype, dorsal view
<emphasis id="D279B64B56CEE56584BF0D97B700B3AC" bold="true">D</emphasis>
same, ventral view.
</paragraph>
</caption>
<paragraph id="D987FD10836FF4B330A874CFCE9D8B21">
Palp (Fig.
<figureCitation id="5960A2EFE082E7F0855E8921EA2F663F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 A D</figureCitation>
). Femoral apophysis (
<abbrev id="ABBRID0EYCBG" xlink_title="femur apophysis">FA</abbrev>
) well developed, thumb-shaped, located at middle part. Patella longer than tibia, as half the length of femur. Tibia longer than wide, with 2 apophyses, retroventral apophysis (
<abbrev id="ABBRID0E3CBG" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
) tail-shaped, tapering towards tip; retrolateral apophysis (
<abbrev id="ABBRID0EADBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
) strong, longer and wider than retroventral apophysis, tip curved ventrally in dorsal view. Bulb pyriform, sperm duct (
<abbrev id="ABBRID0EEDBG" xlink_title="sperm duct">SD</abbrev>
) distinct, V-shaped. Prolateral tegular apophysis (
<abbrev id="ABBRID0EIDBG" xlink_title="Prolateral tegular apophysis">PTA</abbrev>
) originating at nearly 9 oclock position, slightly curved, tip truncated. Tegular sheet (
<abbrev id="ABBRID0EMDBG" xlink_title="tegular sheet">TS</abbrev>
) spade-shaped, anterior part large and wide, posterior part small, tongue-shaped. Embolus (E) needle-like, tapering towards tip.
</paragraph>
<caption id="73A2F2D28251AE5A249F6B0B210DDFEE" ID-DOI="10.3897/zookeys.1226.137752.figure6" ID-arpha="36B089EF-2012-59AF-851E-9A8D694A5AD4" httpUri="https://binary.pensoft.net/fig/1243805" startId="F6">
<paragraph id="931CD527AEE65F14367B1610154C65AA">
<label id="BACCECCB5BF65A89B836A727F553EACF">Figure 6.</label>
</paragraph>
<paragraph id="79F3E9ADD893EB340221A27267932345">
Copulatory organs of
<taxonomicName id="BECB2536C938B11C9F0580CD0C43E7A6" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tianyan" status="sp. nov.">
<emphasis id="93D3997CA5843087114CA54C8D5961CC" italics="true">Mutatus tianyan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="68B9FFBE0EA47357E85DA3AE51613760" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="241D17F3BD72C96524C0E40A4B0E041D" bold="true">A</emphasis>
male left palp, prolateral view
<emphasis id="ADB3A6CC61CF66E63F194D568F294E79" bold="true">B</emphasis>
same, retrolateral view
<emphasis id="8BB154AE631F46EF02268B61EE835685" bold="true">C</emphasis>
same, ventral view
<emphasis id="1B5112B661CB80B1AA83A414677D9555" bold="true">D</emphasis>
same, dorsal view
<emphasis id="936E5AE7265EC5913A2152857CC41E37" bold="true">E</emphasis>
epigyne, ventral view
<emphasis id="9A035AD7996ECB1BE4D6CA399F91F418" bold="true">F</emphasis>
same, dorsal view. Abbreviations: E — embolus;
<abbrev id="ABBRID0EPEBG" xlink_title="femur apophysis">FA</abbrev>
— femoral apophysis;
<abbrev id="ABBRID0ETEBG" xlink_title="Prolateral tegular apophysis">PTA</abbrev>
— prolateral tegular apophysis;
<abbrev id="ABBRID0EXEBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
— retrolateral tibial apophysis;
<abbrev id="ABBRID0E2EBG" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
— retroventral tibial apophysis;
<abbrev id="ABBRID0E6EBG" xlink_title="sperm duct">SD</abbrev>
— sperm duct;
<abbrev id="ABBRID0EDFBG" xlink_title="tegular sheet">TS</abbrev>
— tegular sheet; B — bursa;
<abbrev id="ABBRID0EHFBG" xlink_title="Copulatory ducts">CD</abbrev>
— copulatory duct;
<abbrev id="ABBRID0ELFBG" xlink_title="Copulatory openings">CO</abbrev>
— copulatory opening;
<abbrev id="ABBRID0EPFBG" xlink_title="connecting tube">CT</abbrev>
— connecting tube;
<abbrev id="ABBRID0ETFBG" xlink_title="Fertilization ducts">FD</abbrev>
— fertilization duct;
<abbrev id="ABBRID0EXFBG" xlink_title="Glandular appendages">GA</abbrev>
— glandular appendage;
<abbrev id="ABBRID0E2FBG" xlink_title="median septum">MS</abbrev>
— median septum; S — spermathecae.
</paragraph>
</caption>
<paragraph id="FA0F82CB79A18DF088E23CA4D74A9D1C">
Female (one of
<typeStatus id="75F8639E17B38B9EBEA28EBB28BF3725" type="paratype">paratype</typeStatus>
, Fig.
<figureCitation id="2350F4CFD75BA8724C347B4317965822" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Mutatus tianyan sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure5" httpUri="https://binary.pensoft.net/fig/1243804">5 C, D</figureCitation>
). Total length 4.02, carapace 1.55 long, 1.44 wide; abdomen 2.47 long, 1.56 wide. Eye sizes and interdistances:
<abbrev id="ABBRID0EKGBG" xlink_title="anterior median eye">AME</abbrev>
0.07,
<abbrev id="ABBRID0EOGBG" xlink_title="anterior lateral eye">ALE</abbrev>
0.10,
<abbrev id="ABBRID0ESGBG" xlink_title="posterior median eye">PME</abbrev>
0.07,
<abbrev id="ABBRID0EWGBG" xlink_title="posterior lateral eye">PLE</abbrev>
0.10;
<abbrev id="ABBRID0E1GBG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0E5GBG" xlink_title="anterior median eye">AME</abbrev>
0.05,
<abbrev id="ABBRID0ECHBG" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0EGHBG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0EKHBG" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0EOHBG" xlink_title="anterior lateral eye">ALE</abbrev>
0.24,
<abbrev id="ABBRID0ESHBG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EWHBG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0E1HBG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E5HBG" xlink_title="posterior lateral eye">PLE</abbrev>
0.06,
<abbrev id="ABBRID0ECIBG" xlink_title="posterior lateral eye">PLE</abbrev>
<abbrev id="ABBRID0EGIBG" xlink_title="posterior lateral eye">PLE</abbrev>
0.37,
<abbrev id="ABBRID0EKIBG" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0EOIBG" xlink_title="posterior lateral eye">PLE</abbrev>
0.06.
<abbrev id="ABBRID0ESIBG" xlink_title="eye area width">EAW</abbrev>
0.52,
<abbrev id="ABBRID0EWIBG" xlink_title="cephalic region width">CRW</abbrev>
0.75,
<abbrev id="ABBRID0E1IBG" xlink_title="eye area width">EAW</abbrev>
/
<abbrev id="ABBRID0E5IBG" xlink_title="cephalic region width">CRW</abbrev>
0.69.
<abbrev id="ABBRID0ECJBG" xlink_title="cephalic region width">CRW</abbrev>
/
<abbrev id="ABBRID0EGJBG" xlink_title="carapace width">CW</abbrev>
0.52.
<abbrev id="ABBRID0EKJBG" xlink_title="median ocular area">MOA</abbrev>
0.22 long, anterior width 0.19, posterior width 0.25. Clypeal height 0.14,
<abbrev id="ABBRID0EOJBG" xlink_title="clypeal height">
<collectingCountry id="B0EB771A452D5DF58053F6160BA68862" name="Switzerland">CH</collectingCountry>
</abbrev>
/
<abbrev id="ABBRID0ESJBG" xlink_title="anterior median eye">AME</abbrev>
2.00. Labium 0.21 long, 0.25 wide. Sternum 0.96 long, 0.91 wide. Other characters as for male, except dorsal scutum absent, color lighter and body slightly larger.
</paragraph>
<paragraph id="52337C66A919982F7C819F9710979F15">
Epigyne (Fig.
<figureCitation id="DB9B094CF2A13FB1CD6BF9C4DA91119E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 E, F</figureCitation>
). Epigynal plate slightly sclerotized, with a pair of atria, a clavate protuberance located at middle part of lateral margin of atrium. Median septum (
<abbrev id="ABBRID0E3JBG" xlink_title="median septum">MS</abbrev>
) dumbbell-shaped. Copulatory openings (
<abbrev id="ABBRID0EAKBG" xlink_title="Copulatory openings">CO</abbrev>
) slit-like, inconspicuous, located at anterior of atrium. Copulatory ducts (
<abbrev id="ABBRID0EEKBG" xlink_title="Copulatory ducts">CD</abbrev>
) straight, indistinct. Connecting tubes (
<abbrev id="ABBRID0EIKBG" xlink_title="connecting tube">CT</abbrev>
) shorter than copulatory ducts. Bursae (B) small, almost rectangular, translucent. Glandular appendages (
<abbrev id="ABBRID0EMKBG" xlink_title="Glandular appendages">GA</abbrev>
) mastoid-shaped. Spermathecae (S) clavate, separated by median septum. Fertilization ducts (
<abbrev id="ABBRID0EQKBG" xlink_title="Fertilization ducts">FD</abbrev>
) located at posterior of spermathecae.
</paragraph>
<paragraph id="69CDA4388287B11027BCD88D8FC58EF4">Measurement of legs:</paragraph>
<paragraph id="5247A1CC45BB2744FC5F345F2AA3C975">
<table id="TID0E62AG" rules="all">
<tbody id="2E4E6471E0AB61439D2BA2BD6F992B38">
<table-wrap id="T5" orientation="portrait" position="float">
<tr id="FE5D7BB9C878B0A631F6B419687B4D9E">
<th id="2E9A665EE210DE6AADD704B3CBE9A19F" colspan="1" rowspan="1">male / female</th>
<th id="68C8FE6C4EC5E82564A9CE75E31D9889" colspan="1" rowspan="1">Fe</th>
<th id="02069E39C4551E988EB04AC4C2DEB275" colspan="1" rowspan="1">Pa</th>
<th id="773AA52C8FEF05AA34B3482EBD2F791B" colspan="1" rowspan="1">Ti</th>
<th id="1FB6056B5A25C940594260780C98559A" colspan="1" rowspan="1">Me</th>
<th id="31237E8142005516DE3C508E1FF2487F" colspan="1" rowspan="1">Ta</th>
<th id="2816216759425791700C756C92549676" colspan="1" rowspan="1">Total</th>
</tr>
<tr id="078B426D1C16B7C2FC7BA106EF8C3E6F">
<td id="2CAF1778BA2380BCF5C82E79E9C053CA" colspan="1" rowspan="1">Leg I</td>
<td id="C30AD152D8FD5B005147EEF1942E2000" colspan="1" rowspan="1">1.49 / 1.43</td>
<td id="96DDE89A5D564B6CDDB7690F2784560F" colspan="1" rowspan="1">0.53 / 0.57</td>
<td id="054C91CF1F7F866984ED7B16B421EEB0" colspan="1" rowspan="1">1.45 / 1.55</td>
<td id="4D0A9B3811C0ED60EC9D1C3909BF0700" colspan="1" rowspan="1">1.30 / 1.37</td>
<td id="FC0B521B6A858935FA3C51ACF2668C0B" colspan="1" rowspan="1">0.73 / 0.77</td>
<td id="9554FC8030302A6CEDA7BA5434A0DA44" colspan="1" rowspan="1">5.50 / 5.69</td>
</tr>
<tr id="0F134CFEFA79DB855D8A5D3E4135D8DC">
<td id="2460474FE331E5E96EE45BE4B208C2AF" colspan="1" rowspan="1">Leg II</td>
<td id="FD58663F11A32EDF7FC448C2523A3CD3" colspan="1" rowspan="1">1.28 / 1.27</td>
<td id="13F7914910DEF2D0362A97B682D5F26E" colspan="1" rowspan="1">0.49 / 0.54</td>
<td id="03E496C2C422FA9A273252FAC2A6FB50" colspan="1" rowspan="1">1.09 / 1.13</td>
<td id="8DDFE33A79D2D7E278D2C65A1B638B9A" colspan="1" rowspan="1">1.07 / 1.10</td>
<td id="1671C20CC4EFA5949C2E7A37A02DE3CF" colspan="1" rowspan="1">0.70 / 0.75</td>
<td id="458A6524401B189097E637CE32052C16" colspan="1" rowspan="1">4.63 / 4.79</td>
</tr>
<tr id="D5474C1180C08C4D07FBE687F06ABD3D">
<td id="A5C3098C32DCA662AC10D5AD80DBD5E5" colspan="1" rowspan="1">Leg III</td>
<td id="085BD9D67B90A658F7CF7426B400932B" colspan="1" rowspan="1">1.10 / 1.10</td>
<td id="E5AC253DAC8936CE9F454E85AC33EA6B" colspan="1" rowspan="1">0.46 / 0.43</td>
<td id="283C5BABC39A3C0F15027CB953062124" colspan="1" rowspan="1">0.84 / 0.82</td>
<td id="35225926DF293F46FA837CB57DA47927" colspan="1" rowspan="1">1.02 / 1.10</td>
<td id="AB4BDDE293B59E403BFF53076FDEB716" colspan="1" rowspan="1">0.61 / 0.68</td>
<td id="4286DE9CB9F4512BDA580AF067020794" colspan="1" rowspan="1">4.03 / 4.13</td>
</tr>
<tr id="1C75F91A636821308E0789BA13D9B370">
<td id="7FD0B442D11804002A06DE2AA3230173" colspan="1" rowspan="1">Leg IV</td>
<td id="D3C0D3C3139456AF252A0A638846E707" colspan="1" rowspan="1">1.59 / 1.68</td>
<td id="1695638F8F2C1CF86B8C7159BF0EDB5D" colspan="1" rowspan="1">0.53 / 0.57</td>
<td id="EE4173EBB441EE7DF688998DB1A3E564" colspan="1" rowspan="1">1.34 / 1.45</td>
<td id="031D77D2901AD7BA4F1937D20314022A" colspan="1" rowspan="1">1.64 / 1.69</td>
<td id="F2996C05FAE12C7B4E3C98A43B89205C" colspan="1" rowspan="1">0.84 / 0.93</td>
<td id="EBB14109ED07DC0DD62F35D45A048370" colspan="1" rowspan="1">5.94 / 6.32</td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
<paragraph id="D6E63DB041BDCDB2401BB41A70E974DA">Spination of legs:</paragraph>
<paragraph id="188D556BDE92CBBE31B58D61A634BDA1">
<table id="TID0E5BBG" rules="all">
<tbody id="925E2A684DEC0A56CAD53F838B4BE03D">
<table-wrap id="T6" orientation="portrait" position="float">
<tr id="CDD09917EFB5FB7375AE78D3B2463528">
<th id="020F5E57246DDC7258839687B077ED6A" colspan="2" isEmpty="true" rowspan="1">-</th>
<th id="D927018ADA706C6F1534240E969AB24D" colspan="1" rowspan="1">Fe</th>
<th id="1545E09B13869E70F2F3D0D5934F6C3B" colspan="1" rowspan="1">Pa</th>
<th id="ABFC90A97236A9140A90A3C9EDE6DE2B" colspan="1" rowspan="1">Ti</th>
<th id="04094C0A3AE01D1B282C828E3DA64673" colspan="1" rowspan="1">Me</th>
<th id="CAF69DA2A08DE5BF5E63217D5638DFC1" colspan="1" rowspan="1">Ta</th>
</tr>
<tr id="CB557617DDBB1B5817E867C66BA904C2">
<td id="B0AAA09B4D10074406BB350402EC55E1" colspan="1" rowspan="4">male</td>
<td id="84CBDDE1A73BF757D297C8028E281E6B" colspan="1" rowspan="1">Leg I</td>
<td id="EC5C7BB863374F1C7D668BD87C8EF0FC" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EWPBG" xlink_title="prolateral">pl</abbrev>
4
</td>
<td id="79DDC34EC272872B5DF00FB0B31160F4" colspan="1" rowspan="1"></td>
<td id="237A158563C6AACFE1A5A779A7459D4E" colspan="1" rowspan="1">
<abbrev id="ABBRID0EBQBG" xlink_title="proventral">pv</abbrev>
6
<abbrev id="ABBRID0EFQBG" xlink_title="retroventral">rv</abbrev>
6
</td>
<td id="EB8E917E1B3A4A9398BAF917369BBB55" colspan="1" rowspan="1">
<abbrev id="ABBRID0ENQBG" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0ERQBG" xlink_title="retroventral">rv</abbrev>
4
</td>
<td id="B212D015DF473AE40801C0C0AEF99FF9" colspan="1" rowspan="1"></td>
</tr>
<tr id="44A552D879BD1E6C33671F575D32F891">
<td id="437A84E6863A25DE24456595A1EE6F21" colspan="1" rowspan="1">Leg II</td>
<td id="D24C9DB209C068055810E8EFF29D67EB" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EARBG" xlink_title="prolateral">pl</abbrev>
1
</td>
<td id="7F44B92DAF955E5AF5D260B2CDB8C562" colspan="1" rowspan="1"></td>
<td id="33A88924C17CA5F12BA8FBB18EFD24B0" colspan="1" rowspan="1">
<abbrev id="ABBRID0ELRBG" xlink_title="proventral">pv</abbrev>
6
<abbrev id="ABBRID0EPRBG" xlink_title="retroventral">rv</abbrev>
4
</td>
<td id="9D6DE673877C05D9F649D46EA2E9B8CB" colspan="1" rowspan="1">
<abbrev id="ABBRID0EXRBG" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0E2RBG" xlink_title="retroventral">rv</abbrev>
3
</td>
<td id="6662811C35494EAB25357172C80A12C8" colspan="1" rowspan="1"></td>
</tr>
<tr id="A892D15D6EA30F95B24E4B1B0C8F4077">
<td id="2A052E06F0C0E615343EE3D2C3506FFE" colspan="1" rowspan="1">Leg III</td>
<td id="AA72B333908FE6533E8EC364AE37595A" colspan="1" rowspan="1">d 1</td>
<td id="9FBAD50D2BAE4219CBCF16582E2E0EB5" colspan="1" rowspan="1"></td>
<td id="4ABDB23CE1EEF8D8341F06E1E1317514" colspan="1" rowspan="1"></td>
<td id="D883C53FC11CC2693A0CAD5056179195" colspan="1" rowspan="1"></td>
<td id="29B4B89B2EAB3BB7737623AEDEA5F58B" colspan="1" rowspan="1"></td>
</tr>
<tr id="8A204D7F5F475F36D97FB699DDDE1C70">
<td id="B29F39D83C24AEC9AC0350B085BDCFC4" colspan="1" rowspan="1">Leg IV</td>
<td id="602DCDFAC4D9B45423ECCD47BFA0249D" colspan="1" rowspan="1">d 1</td>
<td id="E24EAAB2669BB42A44CD741D2912098D" colspan="1" rowspan="1"></td>
<td id="167C00194BB2C18A810547CFC366FE12" colspan="1" rowspan="1"></td>
<td id="48C17D52D54F4F44941524D421158A6F" colspan="1" rowspan="1"></td>
<td id="0F33882EC5E6020069055F6277814669" colspan="1" rowspan="1"></td>
</tr>
<tr id="6FC8F1F5D554B492337385A97BB5F664">
<td id="76F98A9547DEC04AC769A526253BF329" colspan="1" rowspan="4">female</td>
<td id="495F48BD76CE54BCEEB9542C432B96AE" colspan="1" rowspan="1">Leg I</td>
<td id="FE0977534867148C8E5C90A38BA96E51" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0ETTBG" xlink_title="prolateral">pl</abbrev>
3
</td>
<td id="1318B954DFE95F70D3BAB190A8F4B016" colspan="1" rowspan="1"></td>
<td id="2E25F46CF43B7605AD71279F73B0F5F3" colspan="1" rowspan="1">
<abbrev id="ABBRID0E5TBG" xlink_title="proventral">pv</abbrev>
7
<abbrev id="ABBRID0ECUBG" xlink_title="retroventral">rv</abbrev>
7
</td>
<td id="7447F4806DBD888352F802390D7C011E" colspan="1" rowspan="1">
<abbrev id="ABBRID0EKUBG" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0EOUBG" xlink_title="retroventral">rv</abbrev>
4
</td>
<td id="8621B37754748246B3B7E969C9736B4A" colspan="1" rowspan="1"></td>
</tr>
<tr id="FFD02B82C020A2AE8969F145FC571C13">
<td id="D040E0B59F5D9926ABA37C25DA16ADE0" colspan="1" rowspan="1">Leg II</td>
<td id="475CFC9F17184D7B7D73E24CE2C3D945" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0E4UBG" xlink_title="prolateral">pl</abbrev>
2
</td>
<td id="ED961A06393642CF4060AABBA6C1EE9E" colspan="1" rowspan="1"></td>
<td id="D2494A3E1D3FEE5664B82CC68DB5A621" colspan="1" rowspan="1">
<abbrev id="ABBRID0EIVBG" xlink_title="proventral">pv</abbrev>
6
<abbrev id="ABBRID0EMVBG" xlink_title="retroventral">rv</abbrev>
6
</td>
<td id="90B7C37C86725597177F354D9098293B" colspan="1" rowspan="1">
<abbrev id="ABBRID0EUVBG" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0EYVBG" xlink_title="retroventral">rv</abbrev>
3
</td>
<td id="D6FB27216D6B4FEC3C512E918AAF9DB7" colspan="1" rowspan="1"></td>
</tr>
<tr id="04A44AC00F025F16448C8DB878B66B9D">
<td id="0F6182C4BD3FFAF069E4321F55AF7E21" colspan="1" rowspan="1">Leg III</td>
<td id="FB610F8F2DC970A84B66A7516290D665" colspan="1" rowspan="1">d 1</td>
<td id="ADD5CB165276AD24D98BB0DF441DFA81" colspan="1" rowspan="1"></td>
<td id="3B29FA682B34F0B53472F25494C1EF17" colspan="1" rowspan="1"></td>
<td id="7D51A77CBCB32A3C3CEB3663C8A474E8" colspan="1" rowspan="1"></td>
<td id="E8857F45CD998268493FFED5E4BAF1CD" colspan="1" rowspan="1"></td>
</tr>
<tr id="8B15E71363B97D13F0C859C4BF4313CE">
<td id="A38D4729A68203C832E4F71974A364A2" colspan="1" rowspan="1">Leg IV</td>
<td id="795C2D9456A6E89C40FA96949159AAA1" colspan="1" rowspan="1">d 1</td>
<td id="3BC83CC40B156EDC66BD1D1BBEFDEBA0" colspan="1" rowspan="1"></td>
<td id="CCF9A83B609131D83CE646C9B4E97EC2" colspan="1" rowspan="1"></td>
<td id="B621E148BFA857332684D3F11BE8C117" colspan="1" rowspan="1"></td>
<td id="01F70C74DEAE9F2B5BBD1E0C5C71D6F0" colspan="1" rowspan="1"></td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFXBG" type="distribution">
<paragraph id="CB6BB1DB6F46938B7D52ACFDA3F22C61">
<heading id="54D5CE31580AB9902E4DB0B1FED16038" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="819EBD38DB4D839E17C411CE4FD12FD8">
Known only from the
<typeStatus id="0B9A67159E67EE029F2211F378F920E5">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,253 @@
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<mods:title id="4451A6A6C3A098D9C6A8C2F6674E14CA">Two new species and two new combinations within Paleosepharia (Coleoptera, Chrysomelidae, Galerucinae) from China</mods:title>
</mods:titleInfo>
<mods:name id="87891034D38D75F541AAE89907802496" type="personal">
<mods:role id="39B8085AE5CB4166329E7D6FC3CCCACA">
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</mods:role>
<mods:namePart id="D411A397B4A3E43F6390E570B8EC21F8">Feng, Chuan</mods:namePart>
<mods:nameIdentifier id="8968058D39A46E1ED06C4025ABC90910" type="ORCID">https://orcid.org/0009-0001-9299-3526</mods:nameIdentifier>
<mods:affiliation id="F57BC1BF8BE87ABD48F0E8A45BA278C1">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Road, Chaoyang District, Beijing 100101, China &amp; University of Chinese Academy of Sciences, No. 19 (A) Yuquan Road, Shijingshan District, Beijing, 100049, China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="B3FE4CE599DCF0F158749EE3CE9A2FDF">Yang, Xingke</mods:namePart>
<mods:nameIdentifier id="4E4EE3FC2FC2C9E2BD0959709F97909F" type="ORCID">0000-0002-0870-4657</mods:nameIdentifier>
<mods:affiliation id="DE4DE0B2C4980E79AB6E34F5329E0D30">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Road, Chaoyang District, Beijing 100101, China</mods:affiliation>
</mods:name>
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<mods:namePart id="D59B35B9ECD055F4A2FE9AB548A4DC12">Ge, Siqin</mods:namePart>
<mods:nameIdentifier id="9AAF6C8A39294B3EC16D8E018B3C18AE" type="ORCID">0000-0001-5924-3400</mods:nameIdentifier>
<mods:affiliation id="38AC6D636D47CD052FEB18126BAC048B">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Road, Chaoyang District, Beijing 100101, China &amp; University of Chinese Academy of Sciences, No. 19 (A) Yuquan Road, Shijingshan District, Beijing, 100049, China</mods:affiliation>
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</mods:titleInfo>
<mods:part id="A8E8112944CFFD5F887634370ADA8847">
<mods:date id="131F082EC029B24BAC61213A508D447F">2025</mods:date>
<mods:detail id="2C35F180270CC3341E1D0026B0788F7E" type="pubDate">
<mods:number id="9FAF470BA422DCE0A1C50A829BDED32C">2025-02-06</mods:number>
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<subSubSection id="04B0B84956FBC7A4CD671140CE66DD46" type="nomenclature">
<paragraph id="FBAC9AED61DFBBC12BF69DE95A75A4F4">
<taxonomicName id="EDF483A8C5ECC06144C0CCAFE481ED4E" authority="Feng, Yang &amp; Ge" authorityName="Feng, Yang &amp; Ge" authorityYear="2025" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="subrubra" status="sp. nov.">
<emphasis id="9354F8F45A96CFD3B85BFE7D9D953726" italics="true">Paleosepharia subrubra</emphasis>
Feng, Yang &amp; Ge
</taxonomicName>
<taxonomicNameLabel id="8AFC922CEC22C6F78872ACABFF95945E" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DDD9D67F7497B6085C7F1B398FBDE053" type="description">
<paragraph id="4145EF63BE319A1EC5ADF4CE20A00BA7">
<figureCitation id="0D1838B0B735F6A491795BA5308AD7A3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paleosepharia subrubra sp. nov. (holotype, male) A C habitus D F aedeagus G depression of the elytra of the male. A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1 mm (A C); 0.5 mm (D F)." figureDoi="10.3897/zookeys.1226.140239.figure1" httpUri="https://binary.pensoft.net/fig/1243032">Figs 1</figureCitation>
,
<figureCitation id="9F2C425E290CDB76DF52C5DB17CCCC1A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paleosepharia subrubra sp. nov. (paratype, female) A C habitus D vaginal palps E spermatheca F apex of abdomen of the female. A dorsal view B ventral view C lateral view. Scale bars: 1 mm (A C); 0.5 mm (D)." figureDoi="10.3897/zookeys.1226.140239.figure2" httpUri="https://binary.pensoft.net/fig/1243033">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGAAE" type="type material">
<paragraph id="3AEDD5C4014D58B04C6DC9B9957948D8">
<heading id="9B81BE2185755F97B15FD5F5B540735E" reason="title">Type material.</heading>
</paragraph>
<paragraph id="CF785C3EF868BAE11597CBEAB4E48EBC">
<materialsCitation id="5963BC7448B2D38D45B6AD2F08174B78" collectingDate="2022-07-26" collectionCode="IZGAS" country="China" county="Chebaling National National Nature Reserve" location="Meiying Lin" municipality="Xiba" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="925E0295CEBCEC36EC7E753905180C7D" bold="true" italics="true">
<typeStatus id="37ED6646C547BB3A6BED6E1B57906C4C" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCount id="EFB2AE7EA643783D6F7AAE554D769E51" count="1" type="male"></specimenCount>
,
<collectingCountry id="F4002F096A7C40F7EA327CDD99057CC7" name="China">China</collectingCountry>
,
<collectingRegion id="62FC11BB3D11A54AB6129F7ABBABB1FE" country="China" name="Guangdong">Guangdong Province</collectingRegion>
,
<collectingCounty id="00927FC29795F6861991F90A1E70610E">Chebaling National National Nature Reserve</collectingCounty>
,
<collectingMunicipality id="624A8890C3A03805B4CF68EC45F0E441">Xiba</collectingMunicipality>
,
<date id="AAB2C77574A59A8C0761A5EFE53B8133" value="2022-07-26">
<collectingDate id="7F8BB42C7D51D6B8E5F5875400AAC5FE" value="2022-07-26">26 Jul. 2022</collectingDate>
</date>
,
<location id="FFBF3A576921A38A69615C1138ACBDF4" LSID="urn:lsid:plazi:treatment:67D6D9A77AEF557F9863D8152F60573C:FFBF3A576921A38A69615C1138ACBDF4" country="China" county="Chebaling National National Nature Reserve" municipality="Xiba" name="Meiying Lin" stateProvince="Guangdong">Meiying Lin</location>
et al. leg.,
<abbrev id="ABBRID0ERAAE" xlink_title="Institute of Zoology, Guangdong Academy of Sciences, Guangzhou">
<collectionCode id="10CDBC0EB884D4F4D1C73D5AAA36E5D2" country="Georgia" httpUri="http://grbio.org/cool/ahe1-tsbj" name="Georgian Academy of Sciences, Insititute of Zoology" type="">IZGAS</collectionCode>
</abbrev>
.
</materialsCitation>
</paragraph>
<caption id="4AB18F6357E8669F2C58BD880275A2DB" ID-DOI="10.3897/zookeys.1226.140239.figure1" ID-arpha="DE9AD48C-4C2B-5FD2-8C2A-65B32921839D" httpUri="https://binary.pensoft.net/fig/1243032" startId="F1">
<paragraph id="9CA4BDC0A1568AE544AB7698B8928927">
<label id="600B53EF1415ACB4D8C0E25F4E3CE781">Figure 1.</label>
</paragraph>
<paragraph id="DBD69B010E6A014E487E692A6273FA99">
<taxonomicName id="0195B585361ED58B80009A71EE941BB5" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="subrubra" status="sp. nov.">
<emphasis id="3EE76DE0187246FB7DB6062ED7175031" italics="true">Paleosepharia subrubra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A55E75DEB86F6EBC29F73A2942A2A505" rank="species">sp. nov.</taxonomicNameLabel>
(holotype, male)
<emphasis id="07364065040B0D447BDA5F008B58717D" bold="true">A C</emphasis>
habitus
<emphasis id="E224FB5CBF8C1E24CDF035506E7E8F29" bold="true">D F</emphasis>
aedeagus
<emphasis id="3BF49DEA48FF6F120FB6F4A7538D5AB2" bold="true">G</emphasis>
depression of the elytra of the male.
<emphasis id="A62C23A33AA442D81364172DB98C5143" bold="true">A, D</emphasis>
dorsal view
<emphasis id="20622704A10AEE2E4B4D4E11A8854BE6" bold="true">B, E</emphasis>
ventral view
<emphasis id="9349928C8667669134754F4D4D937694" bold="true">C, F</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="BDB0D81A85F77D3188E0DC55A3008D55" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="CFAACCEC82B4D3D93C11665ED87D915F" bold="true">D F</emphasis>
).
</paragraph>
</caption>
<paragraph id="3A1C1B5C12DF09D204BF7E9C628633BC">
<emphasis id="0D707E7FA3155066140959FA85F09A04" bold="true" italics="true">
<typeStatus id="5DFE6CF48AFAD645C79D45BADAA70D2F" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCount id="8032D8EE687A6F6163D3BCE487C77124" count="1" type="female">1 ♀</specimenCount>
, •
<materialsCitation id="4F0C2816B2289EF4CAC89E1C56E029E5" collectingDate="1997-08-01" collectionCode="IZCAS" collectorName="Youwei Zhang" country="China" location="Huangxizhou" municipality="Mount Wuyi" specimenCount="1" stateProvince="Fujian" typeStatus="paratype">
<collectingCountry id="2C61D6103B9CB37CEF49FC042C890491" name="China">China</collectingCountry>
,
<collectingRegion id="CF4B06383C0BBAC4F900192037AC6F0E" country="China" name="Fujian">Fujian Province</collectingRegion>
,
<collectingMunicipality id="5FE98E480486BA71F339F2147DB77257">Mount Wuyi</collectingMunicipality>
,
<location id="391CC2D43D1FFBDF0BBBAD70387ACB10" LSID="urn:lsid:plazi:treatment:67D6D9A77AEF557F9863D8152F60573C:391CC2D43D1FFBDF0BBBAD70387ACB10" country="China" municipality="Mount Wuyi" name="Huangxizhou" stateProvince="Fujian">Huangxizhou</location>
,
<date id="AB9E1B3444245645E2292A0E0A0F83AE" value="1997-08-01">
<collectingDate id="92A25D5EE402A271C598211C8D5E668C" value="1997-08-01">1 Aug. 1997</collectingDate>
</date>
,
<collectorName id="E4D72451176BD64B99F2EAC3F94B1ACF">Youwei Zhang</collectorName>
leg.,
<named-content id="58DA2B8D3B16239C5206F4E471048412" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="E85ED74C31F5EF6524A12D10056D5AF3" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<caption id="02EB4CB4C8B65E4AFB4D20DCED5F5407" ID-DOI="10.3897/zookeys.1226.140239.figure2" ID-arpha="EEFD0A14-2CFF-5C80-B13A-5E2B67DD78A0" httpUri="https://binary.pensoft.net/fig/1243033" startId="F2">
<paragraph id="905C75193FBCEB0B67725125DEB342BA">
<label id="C73FC884F86CF0A277AA78BC3EF317E3">Figure 2.</label>
</paragraph>
<paragraph id="CD0DE09469642E95D5385D6879112887">
<taxonomicName id="7D38CAB5E136F6C46707F680BF0EA575" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="subrubra" status="sp. nov.">
<emphasis id="A5657F401772F504B11E10A60716BE01" italics="true">Paleosepharia subrubra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5D03DC55CA169326ADF7C7E4E711372C" rank="species">sp. nov.</taxonomicNameLabel>
(paratype, female)
<emphasis id="E623641A4448939494E0A680A1AF276A" bold="true">A C</emphasis>
habitus
<emphasis id="96D3CDF86DEEE5B1B6014CE284566A01" bold="true">D</emphasis>
vaginal palps
<emphasis id="10CA5C8FA75632B8356C5C449B6B104A" bold="true">E</emphasis>
spermatheca
<emphasis id="1631E571D5AC035DD9F4D384705C6483" bold="true">F</emphasis>
apex of abdomen of the female.
<emphasis id="6F70B8369B0D1D51BCF09734F21D303D" bold="true">A</emphasis>
dorsal view
<emphasis id="71A1AFE2D8265E2CF5A8E011E7807053" bold="true">B</emphasis>
ventral view
<emphasis id="69C81E71AF63E71F9A6C426F1C31875E" bold="true">C</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="200869DAC0ABA3E7418E5B88E512FAB5" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="82E5527705B0A5F13836CE6D6AEADC76" bold="true">D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESDAE" type="etymology">
<paragraph id="A9DAB86DD7507985B8D8289693003BB2">
<heading id="14D88B7A10F37FFB90EABF02389E6AD3" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="B354992AA5202A21C6D0D8E2A7598804">This species name refers to the red base of the elytra.</paragraph>
</subSubSection>
<subSubSection id="SECID0EXDAE" type="diagnosis">
<paragraph id="CB8E4F3F81CD16B0F5715CD0CEEA0CEA">
<heading id="4008C958E99C1ABC2C21EF0F3A9DEE64" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="706F6173660CE0714B04BF1C0ABCCEEB">
The new species closely resembles
<taxonomicName id="03AED64C2B3A047B57E4F05219C2DE1D" authority="Medvedev, 2004" authorityName="Medvedev" authorityYear="2004" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="kubani">
<emphasis id="3A62AA31552011ECCDDC0C18F03D5E41" italics="true">P. kubani</emphasis>
Medvedev, 2004
</taxonomicName>
with both species having black elytra and each elytron with two yellow spots. It differs from
<taxonomicName id="EB64DC2677450A3C288D4C8BFCC65AA4" authorityName="Medvedev" authorityYear="2004" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="kubani">
<emphasis id="7650265CA83ECE691E74BB24D646EFD6" italics="true">P. kubani</emphasis>
</taxonomicName>
in the elytra with a longitudinal depression at the base and connected to a semicircular depression at the end. In
<taxonomicName id="24E936F74B79C010577CDB4D969EAD64" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="subrubra" status="sp. nov.">
<emphasis id="7AC1F552B327DF5F21896759DE366156" italics="true">P. subrubra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4B5368215C983ED2F7C43A58E563FB0D" rank="species">sp. nov.</taxonomicNameLabel>
, the elytra have a rectangular depression at the base. Aedeagus slender, rounded at apex, tectum long, truncate at apex, not reaching apex of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5EAE" type="description">
<paragraph id="2EE54723FDC763050576CBDD058C48E0">
<heading id="9341F748184155B98100CF20C515B79D" reason="title">Description.</heading>
</paragraph>
<paragraph id="313042278AE079F51257C3CC47A3D841">
<emphasis id="823D0F7A8CC9A56FEA1901A35A231FE6" bold="true">Male</emphasis>
: Length:
<quantity id="9172B58C1D133571D10A71A88EEEE764" metricMagnitude="-3" metricUnit="m" metricValue="5.2" unit="mm" value="5.2">5.2 mm</quantity>
, width:
<quantity id="1D202EBDE5FBAF62489A30780EF1B9CC" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.5 mm</quantity>
. Head, pronotum, scutellum, ventral surface of body and femur reddish-brown, antennae black-brown, with first antennomere reddish-brown. Elytra black and reddish-brown at base, each elytron has two white spots, tibiae and tarsi black. Vertex with sparse punctures. Frontal tubercles transverse, extending downward between antennal bases. Antennae slender, longer than body. First antennomere shiny, bare, rod-shaped, second to eleventh antennomeres with short hairs, second antennomere shortest, third antennomere about 1.55 × as long as second; fourth antennomere about 1.78 × as long as third, fifth antennomere about 1.1 × as long as fourth, sixth to eleventh antennomeres equal in length to fifth. Pronotum about 1.29 × as wide as long, lateral margins straight, basal border slightly convex, apical border slightly concave, disc strongly convex, with lateral fovea and dense minute punctures. Scutellum triangular, smooth, impunctate. Elytra wider than pronotum, humeri convex, with pair of weak depressions and pair of rounded protrusions near suture at basal 1 / 3; disc with dense, minute punctures. Elytral epipleuron broad at base, gradually narrowed from middle to apex. Each tibia with distinct spur at apex, segment 1 of hind tarsi longer than remaining segments combined. Aedeagus slender, with parallel-sided and rounded tip, moderately recurved at apex in lateral view. Lateral spiculae short and slightly curved, ventral spiculae comb-shaped, and median spiculae long.
</paragraph>
<paragraph id="F82B804D1C2BB756615B4F0F2BA42BF5">
<emphasis id="83710F7B17F8C2F053E50B7C96A87C16" bold="true">Female</emphasis>
: Length:
<quantity id="4A1549DF1491E4EF8AACC33AFF8DC382" metricMagnitude="-3" metricUnit="m" metricValue="4.6" unit="mm" value="4.6">4.6 mm</quantity>
, width:
<quantity id="7E8F8DF8CD443C467DA8834397D4B767" metricMagnitude="-3" metricUnit="m" metricValue="2.2" unit="mm" value="2.2">2.2 mm</quantity>
.
</paragraph>
<paragraph id="E3A2583C33ED01C0CEA4C953140BF25F">Third antennomere about 1.8 × as long as second; fourth antennomere about 1.6 × as long as third. Elytra without protrusions and with slight depressions at base. Spermatheca with big and rounded nodulus, middle part short, and cornu straight. Vaginal palps with wide base and rounded apex, each palp slightly narrowing posteriorly, with 5 setae placed at apex, additional 2 setae subapically.</paragraph>
</subSubSection>
<subSubSection id="SECID0ELFAE" type="distribution">
<paragraph id="1A6914C7005C7A12B749E9886EA38347">
<heading id="4FAB4E4AB0A3B7F653305D64D7A60957" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="49429ADE0F57D962728F90902D37BCB2">
<collectingCountry id="40CEB35FDA108ADFB2F8C529061F1D4B" name="China">China</collectingCountry>
:
<collectingRegion id="BFCEDD13E59AF2FB87D6C6F3DD0FBF49" country="China" name="Fujian">Fujian</collectingRegion>
,
<collectingRegion id="0AAB6554F15C75CABEE5FED75EDC80FC" country="China" name="Guangdong">Guangdong</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,643 @@
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<mods:title id="179E40D2D110756F0E961EEBF2276207">Reniscymnus gen. nov., a new genus of Scymnini (Coleoptera, Coccinellidae) from the Oriental region</mods:title>
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<mods:namePart id="0FFE3D4F827ECBA9C7F6097031D64497">Peng, Feng</mods:namePart>
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<mods:namePart id="270B90F8F90E445A798097BF9A6AC7B0">Tang, Mingjie</mods:namePart>
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<subSubSection id="935FB180BF76ACBF675C3D61B1404C76" type="nomenclature">
<paragraph id="3E51D4409C263FB0541EFCC8664E79EB">
<taxonomicName id="F916886CF991853FEC673CB96A8D7012" authority="Peng &amp; Chen" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cordatus" status="sp. nov.">
<emphasis id="6321AA8495AF3C318A34B9ED6B3A1C82" italics="true">Reniscymnus cordatus</emphasis>
Peng &amp; Chen
</taxonomicName>
<taxonomicNameLabel id="311E8CE085D1B2AFC5B31E4CBB1039D6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="B9E15C16C515491FB12CF7D34291D174" type="description">
<paragraph id="2CC31A97B533A73E877859D0DCE2A7A5">
<figureCitation id="90C1B60F6973196284B68A0372E25616" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EF3AE" type="diagnosis">
<paragraph id="D484CCAA3BED3C28C3A35927EF525D1D">
<heading id="CDEC0CD9898839257F6674BD0F0E50BD" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B2E8FF7B14CCEFEBDBF7F773C9850B86">
This species resembles
<taxonomicName id="FBCD0AF7F1CE5AD4EFDE85EA0124E293" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="explanatus" status="sp. nov.">
<emphasis id="307494385F5692BF50894FD036F60FD9" italics="true">R. explanatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1F8C3443B5D308889200C1DCB0E6735" rank="species">sp. nov.</taxonomicNameLabel>
in external appearance but differs from the latter in the well-developed and elongate inner arm of penis capsule, and the subtriangular tegmen with penis guide in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EW3AE" type="description">
<paragraph id="13F1D6D2AFE707F6A1A0CCB74F3FBEEC">
<heading id="2D04030548DFDE0D3018E6A44693024B" reason="title">Description.</heading>
</paragraph>
<paragraph id="1462D7B7C4AB9CC1A848B0CA3B037C4D">
<abbrev id="ABBRID0E33AE" xlink_title="total length">TL</abbrev>
:
<quantity id="736ED056A3A1BE132872DF11374A5146" metricMagnitude="-3" metricUnit="m" metricValue="1.505" metricValueMax="1.63" metricValueMin="1.38" unit="mm" value="1.505" valueMax="1.63" valueMin="1.38">1.381.63 mm</quantity>
,
<abbrev id="ABBRID0EA4AE" xlink_title="total width">TW</abbrev>
:
<quantity id="460F3287CBB452F5517BADA21BD537C0" metricMagnitude="-3" metricUnit="m" metricValue="1.165" metricValueMax="1.25" metricValueMin="1.08" unit="mm" value="1.165" valueMax="1.25" valueMin="1.08">1.081.25 mm</quantity>
,
<abbrev id="ABBRID0EE4AE" xlink_title="total height">TH</abbrev>
:
<quantity id="45D0E040109AF84D8A85CE57C12B7B67" metricMagnitude="-4" metricUnit="m" metricValue="7.4" metricValueMax="7.8" metricValueMin="7.0" unit="mm" value="0.74" valueMax="0.78" valueMin="0.7">0.700.78 mm</quantity>
,
<abbrev id="ABBRID0EI4AE" xlink_title="total length">TL</abbrev>
/
<abbrev id="ABBRID0EM4AE" xlink_title="total width">TW</abbrev>
: 1.261.41,
<abbrev id="ABBRID0EQ4AE" xlink_title="elytral length">EL</abbrev>
/
<abbrev id="ABBRID0EU4AE" xlink_title="elytral width">EW</abbrev>
: 0.991.09,
<abbrev id="ABBRID0EY4AE" xlink_title="pronotal length">
<collectionCode id="C7842554FD19E73406290F77B4C20128" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0E34AE" xlink_title="pronotal width">
<collectionCode id="540E0F2F7D53E918F116B390E3964E6E" country="Thailand" name="Paleontological Collections">PW</collectionCode>
</abbrev>
: 0.490.52,
<abbrev id="ABBRID0EA5AE" xlink_title="head width">HW</abbrev>
/
<abbrev id="ABBRID0EE5AE" xlink_title="pronotal width">
<collectionCode id="D052B19102585BF442D93F515DCD3F30" country="Thailand" name="Paleontological Collections">PW</collectionCode>
</abbrev>
: 0.640.67,
<abbrev id="ABBRID0EI5AE" xlink_title="pronotal width">
<collectionCode id="3EF079AD841365B0490C00DD3CA7CC7C" country="Thailand" name="Paleontological Collections">PW</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EM5AE" xlink_title="elytral width">EW</abbrev>
: 0.660.69.
</paragraph>
<paragraph id="0F32B1028244F377F5C3B2111258D7EA">
Body elongate oval, dorsum with white pubescence. Area of frons near clypeus with suberect and downward setae, distinctly longer and denser than those on top (Fig.
<figureCitation id="1E6FA93362A4F10879F4E1F38182E9CD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 B</figureCitation>
). Body uniformly pale yellow, except tip of mandibles dark brown (Fig.
<figureCitation id="BD1DD1DD695FE01D7F036160347050EE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 A C</figureCitation>
). Eyes finely faceted, large, interocular distance about 0.3 times head width. Head and pronotal punctures fine and dense. Surface of elytra with coarse and shallowly impressed punctures.
</paragraph>
<paragraph id="460333E35AD97C2F8A2AE0B0982B5275">
Abdominal postcoxal lines incomplete and roundly recurved, reaching 3 / 4 of length of ventrite 1, area enclosed by the lines sparely punctate, widely smooth along lines; ventrite 1 with fine and sparse punctures in middle, irregularly distributed (Fig.
<figureCitation id="944D68F3DC4312854B1B9FB69455CA25" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 D</figureCitation>
).
</paragraph>
<paragraph id="6405880419DDD0E7F223416AFFA18643">
Male genitalia. Penis moderately stout and rather long, longer than abdomen; penis capsule with long inner arm and short outer arm (Fig.
<figureCitation id="0B9156E282E2FB38A5FB3DA2FCBDD201" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 E, G</figureCitation>
), apex of penis swollen with cordate membranous appendages (Fig.
<figureCitation id="2CFA66DE345475D28C739FDCF23518A2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 F, H</figureCitation>
); tegminal strut stout with a finger-like protrusion at apex (Fig.
<figureCitation id="D7A06C10ED47EDE8672A60D949E9CFD8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 I J</figureCitation>
); phallobase subtrapezoid in lateral view (Fig.
<figureCitation id="C1A3DDF17F572864636FC17F3E54A606" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 I</figureCitation>
); tegmen with penis guide triangular, widest at base, apex pointed in lateral view, 3.5 times as long as parameres (Fig.
<figureCitation id="294295E49DA320F091D91BCF93E517B1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 I</figureCitation>
); in inner view, penis guide triangular, gradually converging apically; parameres short with few long setae at apex (Fig.
<figureCitation id="F53E8D5E5C1F6D1AB314DB6CB3306DE8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E16AE" type="type material">
<paragraph id="A93ABA934C6BFFF43C144B1C1B0062C7">
<heading id="15F330F9FBE8C293E481CACD85299D23" reason="title">Type material.</heading>
</paragraph>
<paragraph id="66DA8E9AC861F8E8CEB77E2E0999C9A6">
<materialsCitation id="7D0984F34BA7F3CE097E39F7403403CC" collectingDate="2007-06-02" collectingDateMax="2007-06-04" collectingDateMin="2007-06-02" collectionCode="NBCA, SCAU" collectorName="X. M. Wang &amp; No." country="Laos" elevation="625" location="Phou Kbouay" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B651CDE71C7CA7A206F10F71F8C46364" bold="true" italics="true">
<typeStatus id="25C81AF050D083EF8693E2892ED40E1E" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="A1DC8292283B94BF2920D3A12C204026" name="Laos">Laos</collectingCountry>
<specimenCount id="F506C691807C00489D716281DAF214C5" count="1" type="male">1 ♂</specimenCount>
;
<location id="39DCDB0D89D09A4F7BADA7A354AA5975" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:39DCDB0D89D09A4F7BADA7A354AA5975" country="Laos" name="Phou Kbouay">Phou Kbouay</location>
<collectionCode id="AD504F3A93C6647C3B50BFD6FE19C538">NBCA</collectionCode>
; alt.
<quantity id="D323CAA83C20D209766F72AD810E4901" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" unit="m" value="625.0" valueMax="750.0" valueMin="500.0">
<elevation id="1C3D7EAB72A3651B85DCC7C9002D89A8" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" unit="m" value="625.0" valueMax="750.0" valueMin="500.0">500750 m</elevation>
</quantity>
;
<date id="813EEF3171317DD69242109987BD2DA3" value="2007-06-02" valueMax="2007-06-04" valueMin="2007-06-02">
<collectingDate id="A21D6D0EBE1F248D7167D15F139A5412" value="2007-06-02" valueMax="2007-06-04" valueMin="2007-06-02">24 Jun. 2007</collectingDate>
</date>
;
<collectorName id="528B7DB698F24ADEB78E44A9430EFC83">X. M. Wang</collectorName>
et al. leg.;
<collectorName id="359175B08B9E5A4FCC92C192C4EF696F">No.</collectorName>
<named-content id="0499B67ED7164D43FFFD26BB5D021180" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Department of Entomology, South China Agricultural University">
<collectionCode id="188CC26AD78FD7826DD33396DB4ED702">SCAU</collectionCode>
</named-content>
(E) 17549;
<named-content id="9DE4FBCDB3D50E3D98B51E03076AE4F3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Department of Entomology, South China Agricultural University">
<collectionCode id="BA9BB16DE9C5FB2FBAEA915BF6DAF285">SCAU</collectionCode>
</named-content>
</materialsCitation>
.
<materialsCitation id="7C70EE9C0B1290BB3A25066A2ED42D68" collectingDate="2007-06-02" collectingDateMax="2007-06-04" collectingDateMin="2007-06-02" collectionCode="NBCA, SCAU" collectorName="X. M. Wang &amp; No." country="Laos" elevation="625" location="Phou Kbouay" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="6BD55B88E99D746025A69D4D866EE665" bold="true" italics="true">
<typeStatus id="58C3753210CAD4F0185761EBA6EBEE28" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="553E2C74949371BBD457E43FFB7A581D" count="1" type="male">1 ♂</specimenCount>
; same data as for holotype
</materialsCitation>
<materialsCitation id="C33CAA78986A35EADA7396ADC73630DD" collectingDate="2007-06-11" collectingDateMax="2007-06-12" collectingDateMin="2007-06-11" collectorName="X. M. Wang" country="Laos" elevation="1000" location="Xam Nua" municipality="Vieng Xai" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="6EC983D3E972C1A8DD23F2B5A6529030" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="DE84A0CEE24E1EE6FB3BE17EB4E7DA8D" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="3880C3C1ED527E95A4F57BFC4669D176">Vieng Xai</collectingMunicipality>
,
<location id="F5740E85473B84EDBDFF386285F4B539" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:F5740E85473B84EDBDFF386285F4B539" country="Laos" municipality="Vieng Xai" name="Xam Nua">Xam Nua</location>
; alt.
<quantity id="F764C10ACC550B25F64364B1C1DDAD64" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<elevation id="6A896F27377375F045D6393ED020F9B5" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</elevation>
</quantity>
;
<date id="B42965E15A16D39585B7BDC393FB402C" value="2007-06-11" valueMax="2007-06-12" valueMin="2007-06-11">
<collectingDate id="C7DB387184F73DF5B5C4F27B16597541" value="2007-06-11" valueMax="2007-06-12" valueMin="2007-06-11">1112 Jun. 2007</collectingDate>
</date>
;
<collectorName id="F22E9F36DF492752EADD479E21826D58">X. M. Wang</collectorName>
et al. leg
</materialsCitation>
. •
<materialsCitation id="F68A49552D2ECD11EBA39448F9A572BC" collectingDate="2007-05-25" collectorName="X. M. Wang" country="Laos" elevation="380" location="Heu Gnommolat" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A4F1CE52CFBBFC2E018544CEC7B45086" count="1" type="female">1 ♀</specimenCount>
;
<location id="FB199E0C827958CA8CE39DEF9412FB24" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:FB199E0C827958CA8CE39DEF9412FB24" country="Laos" name="Heu Gnommolat">Heu Gnommolat</location>
; alt.
<quantity id="362C09336AD72E2BF95701A767E8B415" metricMagnitude="2" metricUnit="m" metricValue="3.8" unit="m" value="380.0">
<elevation id="D0AA808F8D94324B3DCB702405C858B9" metricMagnitude="2" metricUnit="m" metricValue="3.8" unit="m" value="380.0">380 m</elevation>
</quantity>
;
<date id="DE72C7B33B3C82F3B0896B78A1E9EDC4" value="2007-05-25">
<collectingDate id="E8D9AA6A70F38035CD6FB6E7E975E87F" value="2007-05-25">25 May 2007</collectingDate>
</date>
;
<collectorName id="2390A8283E0C907F8CFBB206601B370D">X. M. Wang</collectorName>
et al. leg
</materialsCitation>
. •
<materialsCitation id="5B2C7A16F0A241D7B15CF30DCB6E9621" collectingDate="2006-06-11" collectorName="C. Telakang" country="Laos" location="Pakxong" municipality="Tad Fane" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="13F7057D0F727DD02B49713A4285388F" count="2" type="female">2 ♀</specimenCount>
;
<collectingMunicipality id="5CF5DD114F939588697D67A8C02CE5F9">Tad Fane</collectingMunicipality>
,
<location id="80299FD599824A960386F7E88EC16BD4" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:80299FD599824A960386F7E88EC16BD4" country="Laos" municipality="Tad Fane" name="Pakxong">Pakxong</location>
;
<date id="80BB69C10E01D9BFE1B652B266711709" value="2006-06-11">
<collectingDate id="267F96D70647C08DA9264EFEF0D3D6CB" value="2006-06-11">11 Jun. 2006</collectingDate>
</date>
;
<collectorName id="ECFA094B446FD382E3B2F8E57D3F9CD9">C. Telakang</collectorName>
leg.
</materialsCitation>
<specimenCount id="BE8F501FA732B78DA909275281B8ED1F" count="1" type="female">1 ♀</specimenCount>
; LiPHi, VoeunKHom;
<date id="33EEEF47E717A95E3922254C7E0B6E12" value="2006-06-09">9 Jun. 2006</date>
; C. Telakang leg. •
<materialsCitation id="25A8727985C64AC57F6161E33A4BB136" collectingDate="2005-12-16" collectorName="C. Telakang" country="China" location="Lakxao" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="D5DD75C8C37F4F3DC69E2090C274249F" count="1" type="female">1 ♀</specimenCount>
; Khammouane
<location id="3BB14EF364A995BEB2F10F8AA4E1A55E" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:3BB14EF364A995BEB2F10F8AA4E1A55E" country="China" name="Lakxao" stateProvince="Hainan">Lakxao</location>
;
<date id="91B891BC065CECBF4223E256D2F01BEF" value="2005-12-16">
<collectingDate id="FF14D3E9960D4901EA54FCD8AD8D7E2D" value="2005-12-16">16 Dec. 2005</collectingDate>
</date>
;
<collectorName id="B2F60ADE16893C35819753E35A923545">C. Telakang</collectorName>
leg.
<collectingCountry id="06DD0B5E4DD90352136011E42D3FFB2A" name="China">China</collectingCountry>
<emphasis id="1D24616BC243745996E433727968AE3D" bold="true">
<collectingRegion id="D57A6B0E7C8F7A8E06A4EFA600468F3E" country="China" name="Hainan">Hainan Prov.</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="94E9A8F4E808E9C7EA7F0DF298168A10" collectingDate="1995-09-18" collectorName="Z. Q. Peng" country="China" location="Diaoluoshan Mountains" municipality="Xin'an" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="9A1C2C1437FF67B22C83309483F85BC3" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="2C39F69BF2616A47670DD54D029EA4E1" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="4B9EDFD6B1E0FEFBDAE59D403C553FD3">Xinan</collectingMunicipality>
,
<location id="C195D64FE68A51A93134471F3B3D5306" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:C195D64FE68A51A93134471F3B3D5306" country="China" municipality="Xin'an" name="Diaoluoshan Mountains" stateProvince="Hainan">Diaoluoshan Mountains</location>
;
<date id="29895EAD0ACC8EC639BDA0275E05DF8F" value="1995-09-18">
<collectingDate id="21F3973D552E4837D5DF00BF2D69DEC0" value="1995-09-18">18 Sep. 1995</collectingDate>
</date>
;
<collectorName id="8AC04927E5807520C73A8E13CC6025DE">Z. Q. Peng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="68378EAF8364C308913A790EF3D69E80" collectingDate="1996-04-01" collectorName="Z. Q. Peng" country="China" location="Jianfengling" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="BDA45470F37FC2ED34562BE3158714D0" count="1" type="male">1 ♂</specimenCount>
;
<location id="C7692936F062BB5DCB2E81C0E27B321A" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:C7692936F062BB5DCB2E81C0E27B321A" country="China" name="Jianfengling" stateProvince="Hainan">Jianfengling</location>
;
<date id="6AE0DDF0D17D8164D56DA3DD09D081BB" value="1996-04-01">
<collectingDate id="2D4E0E25DCA284EFE06C2081AAF1DB17" value="1996-04-01">1 Apr. 1996</collectingDate>
</date>
;
<collectorName id="A85FDD60395B6B5F9B946A0982340240">Z. Q. Peng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="7138A43D29DF55B27803A8D78A88F367" collectingDate="1995-09" collectorName="Z. Q. Peng" country="China" location="Jianfengling" municipality="Tianchi" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="50E1D5E80496FCC6197E551BD41175BD" count="2" type="male">2 ♂</specimenCount>
<specimenCount id="9EFA51463088CE0162B16B891CC8D8F2" count="4" type="female">4 ♀</specimenCount>
;
<collectingMunicipality id="2068107420D0E3415D117C9D25D8F6B5">Tianchi</collectingMunicipality>
,
<location id="0416FDCA5B5EF5412B7595BC63C7D4F0" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:0416FDCA5B5EF5412B7595BC63C7D4F0" country="China" municipality="Tianchi" name="Jianfengling" stateProvince="Hainan">Jianfengling</location>
;
<date id="168AF52DFA3903CE61C3BEC193E61B63" value="1995-09">
<collectingDate id="F5964841764824848CFC3AACBAD6EB0E" value="1995-09">Sep. 1995</collectingDate>
</date>
;
<collectorName id="914024E5A4923F5EB280535B4F9D6B34">Z. Q. Peng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="56223000B77F54A9EA68667B6B8C060D" collectingDate="1996-05-03" collectorName="Z. Q. Peng" country="China" location="Wuzhishan Mountains" specimenCount="10" specimenCount-female="8" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="FC56BFAFD76EED92ACDD27E164FB2BDB" count="2" type="male">2 ♂</specimenCount>
<specimenCount id="053741C1454230D5525341B1B62EC055" count="8" type="female">8 ♀</specimenCount>
;
<location id="B28D38E2801D0659F177DF65DAD321CB" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:B28D38E2801D0659F177DF65DAD321CB" country="China" name="Wuzhishan Mountains" stateProvince="Hainan">Wuzhishan Mountains</location>
;
<date id="3125D8B0C5859357146504A3923FAC43" value="1996-05-03">
<collectingDate id="66430C556EA6A9B5B099F691E5CE1CC5" value="1996-05-03">3 May 1996</collectingDate>
</date>
;
<collectorName id="19F55EF7935B01CD0EE7B2E2FA7E687A">Z. Q. Peng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="603FB7D150F7A1C4DAC18D825DB42BB3" collectingDate="1995-08" collectorName="Z. Q. Peng" country="China" location="Danzhou City" municipality="Shamaoling" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C6C1C92F0B2B99395A9F54AAD47087E6" count="1" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="74AE756BC1B4AF83746A4F8BC5F780CF">Shamaoling</collectingMunicipality>
,
<location id="3DC7EEACE03A71945CFCE4376F79B01F" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:3DC7EEACE03A71945CFCE4376F79B01F" country="China" municipality="Shamaoling" name="Danzhou City" stateProvince="Hainan">Danzhou City</location>
;
<date id="2B83EB3CA16796601EE6CA41B90B69F5" value="1995-08">
<collectingDate id="EAE8995787DEC2C6FC424F892D33A8AE" value="1995-08">Aug. 1995</collectingDate>
</date>
;
<collectorName id="35C6CF7756899495802E2EF6EC48883C">Z. Q. Peng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="E8ABCEC485F55F6F8A11327BEFD63E78" collectingDate="1995-09" collectorName="Z. Q. Peng" country="China" location="Diaoluoshan Mountains" municipality="Shijing" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="2F5020B581B740A427EFE8EB0B80D068" count="1" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="F6A986CB61CA1E921F1A8B7F5AF2C7B0">Shijing</collectingMunicipality>
,
<location id="B09897785DC646C6CB48D94C56B2A8A9" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:B09897785DC646C6CB48D94C56B2A8A9" country="China" municipality="Shijing" name="Diaoluoshan Mountains" stateProvince="Hainan">Diaoluoshan Mountains</location>
;
<date id="257B0AC7CBA97BBD5D2B1C7C8CC5D9B8" value="1995-09">
<collectingDate id="DF3EA8E3BF7A9146A4DF22554FFF827D" value="1995-09">Sep. 1995</collectingDate>
</date>
;
<collectorName id="0818EB403C13FB1FBCDC808D7F2AAD14">Z. Q. Peng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="8565110746A1DDE0136F398BE3F155C6" collectingDate="1995-09" collectorName="Z. Q. Peng" country="China" location="Limushan Mountains" municipality="Wufenchang" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="2CA9F2280A4108C6AA9AFA928B3E405A" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="75DA73CA7C48683C96193670EC0089EE">Wufenchang</collectingMunicipality>
,
<location id="27578C5F34ED324EBD5695AA84EA47F1" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:27578C5F34ED324EBD5695AA84EA47F1" country="China" municipality="Wufenchang" name="Limushan Mountains" stateProvince="Hainan">Limushan Mountains</location>
;
<date id="8ED61DBD1551805D300104CDE90BC03C" value="1995-09">
<collectingDate id="B434074A8765D50C51343D0238330F60" value="1995-09">Sep. 1995</collectingDate>
</date>
;
<collectorName id="AF79A594B06F299093131148B265C593">Z. Q. Peng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="AFC9C5B0CE78523B928A3D03F44336D6" collectingDate="1996-04-21" collectorName="Z. Q. Peng" country="China" location="Limushan Mountains" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="B34D0A832E7CF8D40056513EF791F073" count="1" type="female">1 ♀</specimenCount>
;
<location id="55D995E647281C7C570DA0792AE4C1F0" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:55D995E647281C7C570DA0792AE4C1F0" country="China" name="Limushan Mountains" stateProvince="Hainan">Limushan Mountains</location>
;
<date id="523EBC805B90E673085CCAA19ACC1DA8" value="1996-04-21">
<collectingDate id="50CEC9713D16E2861E14083B9E54DE26" value="1996-04-21">21 Apr. 1996</collectingDate>
</date>
;
<collectorName id="3DF3FDC1AD0D4D7FA34923E164B919A4">Z. Q. Peng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="834438C779711C1534395160D344C6FB" collectingDate="2019-04-16" collectorName="X. M. Wang" country="China" location="Yinggeling" municipality="Yinggezui" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="5E0007625DB690CD17085997D5166C7B" count="1" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="8BC364BE94F1389F7149EA4C01B856DD">Yinggezui</collectingMunicipality>
,
<location id="A828AA8026A2384287DDEE522D29ACB6" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:A828AA8026A2384287DDEE522D29ACB6" country="China" municipality="Yinggezui" name="Yinggeling" stateProvince="Hainan">Yinggeling</location>
;
<date id="A251721B0A539F6423127CA3F5E3EFA2" value="2019-04-16">
<collectingDate id="BD2E834FB1E8E6E59708C1C386D5C597" value="2019-04-16">16 Apr. 2019</collectingDate>
</date>
;
<collectorName id="8A14C5389F4252FF2EE0A7CA904AA1EB">X. M. Wang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="A0D2F859F15165A69F674BECB4E0307D" collectingDate="2019-04-16" collectorName="X. M. Wang" country="China" elevation="670" latitude="18.907223" location="Wuzhishan Forest" longLatPrecision="21" longitude="109.67972" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="B3EA07949D8201E825FAA0B29B50B12A" count="1" type="female">1 ♀</specimenCount>
;
<location id="4AE5A82E125BEF56F2561E878DEE9B94" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:4AE5A82E125BEF56F2561E878DEE9B94" country="China" latitude="18.907223" longLatPrecision="21" longitude="109.67972" name="Wuzhishan Forest" stateProvince="Hainan">Wuzhishan Forest</location>
Park;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYAAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[109.679722,18.907222]}">
<geoCoordinate id="38E9719149AE3A8DFE97959ACEBDF0F7" degrees="18" direction="north" minutes="54" orientation="latitude" precision="15" seconds="26" value="18.907223">18 ° 54 ' 26 &quot; N</geoCoordinate>
,
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</named-content>
; alt.
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</quantity>
;
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<collectingDate id="CF6A7EF776A6DAA801FEB3505F6AF9ED" value="2019-04-16">16 Apr. 2019</collectingDate>
</date>
;
<collectorName id="FF6E6AF1BBF128D19C1068E141B11A48">X. M. Wang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="9C9F1C6913E968EFC2A2FC570C78019D" collectingDate="1997-10-16" collectorName="Z. Q. Peng" country="China" location="Bawangling" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="FBF7A73145793A1B5BD5D00092A5D996" count="1" type="female">1 ♀</specimenCount>
;
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;
<date id="CB6F2343BDB2DF6D1D9AE0905A935D4C" value="1997-10-16">
<collectingDate id="3ED4444945FC8D5ADE3DEE27370B780A" value="1997-10-16">16 Oct. 1997</collectingDate>
</date>
;
<collectorName id="F73B97169EE8BC7A7975DAD735198479">Z. Q. Peng</collectorName>
leg.
</materialsCitation>
<materialsCitation id="EA4E8CB355D96977C53D9F8C9F120DD9" collectingDate="2006-07-22" collectorName="Z. Q. Peng" country="China" location="Limushan Mountains" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="0265D5BF59B8FC9D29360C50E5A8799E" count="1" type="female">1 ♀</specimenCount>
;
<location id="F04455B7F82DB68778B401AF8AD73E73" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:F04455B7F82DB68778B401AF8AD73E73" country="China" name="Limushan Mountains" stateProvince="Guangxi">Limushan Mountains</location>
;
<date id="AEB5836749695347F4977C608D846876" value="2006-07-22">
<collectingDate id="F2989D794975AAD6E47E44D3115CFEF1" value="2006-07-22">22 Jul. 2006</collectingDate>
</date>
;
<collectorName id="88BCA1CF52AE4ABDD990435E3BF5238A">Z. Q. Peng</collectorName>
leg.
<emphasis id="D76791A074927F7E729C4C0C5960D953" bold="true">
<collectingRegion id="7EC0DE29D63A1AA2072CEE9EF4B935B6" country="China" name="Guangxi">Guangxi Prov.</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="10FEEAD2F9301B6732D330F713F2725E" collectingDate="2004-11-07" collectorName="C. W. Zhang" country="China" location="Shiwandashan Mountains" municipality="Fulongshan Mountains" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="C421F8CF872C17C8564834F93BCCDCE1" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="7B333898ECF405E2A4E839C8DE0E146A" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="8613A08801BBCBAFBBD701B4A8D3CCD6">Fulongshan Mountains</collectingMunicipality>
,
<location id="E5830210A49ABDC4C4DC69903E8AA144" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:E5830210A49ABDC4C4DC69903E8AA144" country="China" municipality="Fulongshan Mountains" name="Shiwandashan Mountains" stateProvince="Guangxi">Shiwandashan Mountains</location>
;
<date id="621EACC4195924566AFA21B9A062FC94" value="2004-11-07">
<collectingDate id="A21D2678AFC2E18CA048E1A115CE682A" value="2004-11-07">7 Nov. 2004</collectingDate>
</date>
;
<collectorName id="E26331A06BFFD6E939C6FC846584725F">C. W. Zhang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="8F96A9C255649E68DCEC6AB5BB0133E8" collectingDate="2005-08-02" collectorName="X. M. Wang" country="China" location="Pingxiang" municipality="Daqingshan Mountains" specimenCount="2" specimenCount-female="2" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="2B21C1CE25D724B9CE8F0EAC9F58AAFF" count="2" type="female">2 ♀</specimenCount>
;
<collectingMunicipality id="148E57A799A439FA7326D9A98B0D9F73">Daqingshan Mountains</collectingMunicipality>
,
<location id="D1929F9D5CCE56D15B5DDFB74DE202D7" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:D1929F9D5CCE56D15B5DDFB74DE202D7" country="China" municipality="Daqingshan Mountains" name="Pingxiang" stateProvince="Guangxi">Pingxiang</location>
;
<date id="B1B636ACBC0407BFEAA822E7EE67EB47" value="2005-08-02">
<collectingDate id="F3B4FAA316564C77218653609A7F749F" value="2005-08-02">2 Aug. 2005</collectingDate>
</date>
;
<collectorName id="D9756357FDAB8121C670E8E77DBB8214">X. M. Wang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="4F99CABBE9B64422D3A82026F731AF15" collectingDate="2004-10-16" collectorName="X. M. Wang" country="China" location="Mao'ershan Mountains" municipality="Banshanyao" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong" typeStatus="paratype">
<specimenCount id="DA1233CEE54935712F28048B88825BA1" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="F78D2A16B3DF6AE43FB6C4F697CD4817">Banshanyao</collectingMunicipality>
,
<location id="D1CDCDBEDFCCA81E4411740764E4F700" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:D1CDCDBEDFCCA81E4411740764E4F700" country="China" municipality="Banshanyao" name="Mao'ershan Mountains" stateProvince="Guangdong">Maoershan Mountains</location>
;
<date id="315EDC4FCA239BAA3BBB4571644B2E46" value="2004-10-16">
<collectingDate id="5B2AE98C517D54CEB377C050E84F3578" value="2004-10-16">16 Oct. 2004</collectingDate>
</date>
;
<collectorName id="D0266E46F0F196CC101B32CB6DD5F274">X. M. Wang</collectorName>
leg.
<emphasis id="9A3592F8AE6460CBDD3DA4008F2E1F3C" bold="true">
<collectingRegion id="7E53A36B8D61A8D6527FD5A4F1E12FFF" country="China" name="Guangdong">Guangdong Prov.</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="4F42DB7BC9D679711CB5EB5F29ABA083" collectingDate="2004-12-21" collectorName="Collector" country="China" location="Huizhou" municipality="Nankunshan Mountains" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Guangdong" typeStatus="paratype">
<specimenCount id="4283F16C5932A5F049671B9C22D9FC5F" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="02ED5A011268B3357B8C80244345F897" count="3" type="female">3 ♀</specimenCount>
;
<collectingMunicipality id="C2E096BEFAAA0E3E97BBEFAACFA5E81C">Nankunshan Mountains</collectingMunicipality>
,
<location id="5E2083D0B2729022F536B5F9565E3683" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:5E2083D0B2729022F536B5F9565E3683" country="China" municipality="Nankunshan Mountains" name="Huizhou" stateProvince="Guangdong">Huizhou</location>
;
<date id="FC17DB28EFD1536B1D6F0A4EB69B769A" value="2004-12-21">
<collectingDate id="7B36CE3FAB0723804B5B912BEC6743D8" value="2004-12-21">21 Dec. 2004</collectingDate>
</date>
;
<collectorName id="430B020C527ECC1B7B826C1B9861B4C9">Collector</collectorName>
unknown
</materialsCitation>
. •
<materialsCitation id="7C06515E70B9004BD2E10EDB9B4D192D" collectingDate="2024-03-16" collectorName="Z. D. Huang" country="China" county="Wuzhishan Scenic Spot" elevation="433" latitude="23.720556" location="Conghua District" longLatPrecision="21" longitude="113.814445" municipality="Liangkou Town" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="CEB42B14750FE7DED512E5659EC566E1" count="1" type="male">1 ♂</specimenCount>
;
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,
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,
<location id="C8043F6E7AC5D639F47161791C35ACE3" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:C8043F6E7AC5D639F47161791C35ACE3" country="China" county="Wuzhishan Scenic Spot" latitude="23.720556" longLatPrecision="21" longitude="113.814445" municipality="Liangkou Town" name="Conghua District" stateProvince="Yunnan">Conghua District</location>
,
<location id="1D91515E72385A3A311F25A121C0495A" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:1D91515E72385A3A311F25A121C0495A" country="China" county="Wuzhishan Scenic Spot" latitude="23.720556" longLatPrecision="21" longitude="113.814445" municipality="Liangkou Town" name="Guangzhou City" stateProvince="Yunnan">Guangzhou City</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EEBAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[113.814444,23.720556]}">
<geoCoordinate id="21B4F07E895B624CD444E0ECD496F1DA" degrees="23" direction="north" minutes="43" orientation="latitude" precision="15" seconds="14" value="23.720556">23 ° 43 ' 14 &quot; N</geoCoordinate>
,
<geoCoordinate id="B49A7054A7919923AE8EFE83638A789F" degrees="113" direction="east" minutes="48" orientation="longitude" precision="15" seconds="52" value="113.814445">113 ° 48 ' 52 &quot; E</geoCoordinate>
</named-content>
; alt.
<quantity id="E5985B27B4D097E435D6EFAFCA57215C" metricMagnitude="2" metricUnit="m" metricValue="4.33" unit="m" value="433.0">
<elevation id="973829A5D48D8C427D6D4F836C4ACA64" metricMagnitude="2" metricUnit="m" metricValue="4.33" unit="m" value="433.0">433 m</elevation>
</quantity>
;
<date id="365021CBB75A595FD2DDBF8C287EBF7A" value="2024-03-16">
<collectingDate id="5CF908B292970F43CD0EDD3412C4257D" value="2024-03-16">16 Mar. 2024</collectingDate>
</date>
;
<collectorName id="9F5CE9BBC56C29ED3FBDEE155C51B2A0">Z. D. Huang</collectorName>
et al. leg.
<emphasis id="9C529B863D8BBB0C76F63B4A171D3435" bold="true">
<collectingRegion id="10F7E5D159367FFDE24540588C63D2A4" country="China" name="Yunnan">Yunnan Prov.</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="E5A0DAC9FD3323C0B5BF95B3B2C35D03" collectingDate="2008-05-22" collectingDateMax="2008-05-23" collectingDateMin="2008-05-22" collectorName="X. M. Wang" country="China" county="Tongbiguan" elevation="1000" location="Yingjiang" municipality="Nabang" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="074B895F26A36FA9EF2D63A6BDDE7577" count="2" type="female">2 ♀</specimenCount>
;
<collectingCounty id="285C48A24C45B9C160EC2487753689EF">Tongbiguan</collectingCounty>
,
<collectingMunicipality id="8CE920B8C652F431355FA082D677969B">Nabang</collectingMunicipality>
,
<location id="CC8858D42F4F38215E35CF0BF009E22A" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:CC8858D42F4F38215E35CF0BF009E22A" country="China" county="Tongbiguan" municipality="Nabang" name="Yingjiang" stateProvince="Yunnan">Yingjiang</location>
; alt.
<quantity id="C5F532230E50A9DDD593EA298362810A" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<elevation id="AAA1E30DA3B4C0AE7F65E1D2EB744474" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</elevation>
</quantity>
;
<date id="B0C144ACB9B506C5D8A88CF59CE799ED" value="2008-05-22" valueMax="2008-05-23" valueMin="2008-05-22">
<collectingDate id="FB2CE21C31503950EF14F6A0F278AE35" value="2008-05-22" valueMax="2008-05-23" valueMin="2008-05-22">2223 May 2008</collectingDate>
</date>
;
<collectorName id="07DB998DC8A5298A031215CC20EAA9DC">X. M. Wang</collectorName>
et al. leg
</materialsCitation>
. •
<materialsCitation id="0258E6F6CCAF80181FC58FF7DC23B199" collectingDate="2008-04-23" collectorName="X. M. Wang" country="China" location="Hekou" municipality="Xiaoweishan Mountains" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="0BC2742C0F1B20801325543D713E0CE6" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="AC42E1FE84B8E612B1A1CCD08B4EA277">Xiaoweishan Mountains</collectingMunicipality>
,
<location id="B3B5DB036A470E27CDD3A2AFD6E00B28" LSID="urn:lsid:plazi:treatment:743646CE001E5F378DBBCAE2B8CE5FBD:B3B5DB036A470E27CDD3A2AFD6E00B28" country="China" municipality="Xiaoweishan Mountains" name="Hekou" stateProvince="Yunnan">Hekou</location>
;
<date id="20E02ECDCC59D355AADF9193C572B43D" value="2008-04-23">
<collectingDate id="7156BBC78D7A6B2826DB062D55ECEFC7" value="2008-04-23">23 Apr. 2008</collectingDate>
</date>
;
<collectorName id="55489A86C5759BD3EE113BFA40AAD33D">X. M. Wang</collectorName>
et al. leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELBAG" type="distribution">
<paragraph id="D1D2DCED7305DEFC98CB3091A9C0C5D3">
<heading id="E37D27567F76B8124A5258DC3E0DF26D" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="ACAD1EFCEDA1D3A780E147C873A0D9DF">
<collectingCountry id="4049F7D18F36FB63899E7F0FE4AE60A8" name="China">China</collectingCountry>
(
<collectingRegion id="6B6658FD90A623AA7679EB4339D93ECC" country="China" name="Hainan">Hainan</collectingRegion>
,
<collectingRegion id="C922780155155DDFE023F4912A6F96E8" country="China" name="Guangdong">Guangdong</collectingRegion>
,
<collectingRegion id="9E61E0363823B35B655E8B6140796534" country="China" name="Guangxi">Guangxi</collectingRegion>
,
<collectingRegion id="4597FADF5D3F0F03A3AD57EC6A24028B" country="China" name="Yunnan">Yunnan</collectingRegion>
);
<collectingCountry id="7CCA215CAB6843E0FAAF6066CF8B24C5" name="Laos">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQBAG" type="etymology">
<paragraph id="CC26A0D2565BE1F0832A164640565462">
<heading id="8EFE8DEA9CC38A2BD707DB96C20547F9" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="CC3D508C0F8CFE1A5E69AF4FC0EB628A">
The species epithet is derived from the Latin adjective “
<emphasis id="B5F628831FBF5ABC79AAAABF6F8CF78D" italics="true">cordatus</emphasis>
”, referring to the nearly cardioid apex of the penis.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EACAG" type="remarks">
<paragraph id="77C3B5EDD6021F0667E77F8DE510B1E3">
<heading id="BF68B98F0F83EEF0512B93DF8B4ACC30" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="113CD24F54BE8F492E37992A7586123E">
The apex of the penis of this species is spatially swollen; consequently, different shapes were observed when the apex was rotated or viewed from different angles (Fig.
<figureCitation id="15FC691F7526A150210B02588F7EF11F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 E H</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,176 @@
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<subSubSection id="346A3063C0EA7F325AEDFADA0401492D" type="nomenclature">
<paragraph id="235821B6A11C2702486D44576D1A2B64">
<taxonomicName id="1BD8961F303A543080CC54F52E975750" ID-CoL="FM3W" authority="Gahan, 1898" authorityName="Gahan" authorityYear="1898" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="A1A38782864990E7590811E88B59CE65" italics="true">Apiocephalus punctipennis</emphasis>
<bibRefCitation id="6F59F3DC3AD6226F1C0B0CBCC591CB8F" DOI="10.1080/00222939808678011" author="Gahan CJ" refId="B6" refString="Gahan CJ (1898) Descriptions of new Longicorn Coleoptera from East Africa. The Annals and Magazine of Natural History Series 7, 2 (7): 40 - 59. https://doi.org/10.1080/00222939808678011" year="1898">Gahan, 1898</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="2E4898135E32381EF5E68E26F30C72D7" type="description">
<paragraph id="8BF91CFBAC49B72803510B204C949F26">
<figureCitation id="4CC0CE8B90D32B110337ECDB4918768A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype of Apiocephalus punctipennis Gahan, 1898 a dorsal view b lateral view c ventral view d frontal view e lateral view of pronotum f labels." figureDoi="10.3897/zookeys.1226.139148.figure1" httpUri="https://binary.pensoft.net/fig/1243454">Chinese common name: 头花天牛 Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DD7FACE7790DACEA72116E9D3F0B7A94" type="reference_group">
<paragraph id="0125C51A9E92B07A95E49C4C60863EFD">
<treatmentCitationGroup id="3BD46EB48ADA9FF69664A5375745E642">
<treatmentCitation id="F08861F4E3B0A951785C457ADBD30C55" author="Gahan CJ" page="43" year="1898">
<taxonomicName id="483664967BB1BF34F5A5D17AE00674EE" ID-CoL="FM3W" authority="Gahan, 1898: 43" authorityName="Gahan" authorityPageNumber="43" authorityYear="1898" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="0B6603AF512ED65263282BA538D8B8A2" italics="true">Apiocephalus punctipennis</emphasis>
<comment id="C78C4B92BCB65510365C597851743A0B">
<bibRefCitation id="C35CBA39939639D95A783E8A7905A669" DOI="10.1080/00222939808678011" author="Gahan CJ" refId="B6" refString="Gahan CJ (1898) Descriptions of new Longicorn Coleoptera from East Africa. The Annals and Magazine of Natural History Series 7, 2 (7): 40 - 59. https://doi.org/10.1080/00222939808678011" year="1898">Gahan, 1898: 43</bibRefCitation>
.
<typeStatus id="E035426A64E619BC8E5AAC5AD81E8E30">Type</typeStatus>
locality:
<collectingCountry id="32BB322272157A05F847E598923C97AF" name="Kenya">Kenya</collectingCountry>
(
<collectingCountry id="E3E06B8B84E143B0FF2F07A1F661516A" name="Kenya">British East Africa</collectingCountry>
);
<treatmentCitation id="453D9D9A02BB259666134B5C84552916" author="Aurivillius C" page="177" year="1912">
<bibRefCitation id="2AB91925A8C26EA295CAEFFAECC73D46" author="Aurivillius C" firstAuthor="Aurivillius" journalOrPublisher="&amp; Schenkling S., Berlin" page="177" refId="B1" refString="Aurivillius C (1912) Cerambycidae: Cerambycinae. Coleopterorum Catalogus pars 39 [Vol. 22]. Junk W. &amp; Schenkling S., Berlin, 574 pp." title="Cerambycidae: Cerambycinae. Coleopterorum Catalogus pars 39 [Vol. 22]. Junk W." year="1912">Aurivillius 1912: 177</bibRefCitation>
</treatmentCitation>
[distribution];
<treatmentCitation id="4A79832FC571F9F9FDBAA5E758F6CCD4">
<bibRefCitation id="41217CEC425BB8FF29E7213A66CD2FE4" author="Corinta Ferreira and Veiga Ferreira" firstAuthor="Corinta Ferreira" journalOrPublisher="Memórias do Instituto de Investigação científica de Moçambique" pagination="77-398" refId="B4" refString="Corinta Ferreira M, Veiga Ferreira G (1959) Catálogo dos Cerambicídeos da Região Etiópica. II. Parte - Supertribos Disteniina, Asemina, Cerambycina, Auxesina &amp; Lepturina. Memórias do Instituto de Investigação científica de Moçambique 1: 77398." title="Catálogo dos Cerambicídeos da Região Etiópica. II. Parte - Supertribos Disteniina, Asemina, Cerambycina, Auxesina &amp; Lepturina." volume="1" year="1959">Corinta Ferreira and Veiga Ferreira 1959</bibRefCitation>
</treatmentCitation>
[catalogue];
<treatmentCitation id="45FF959659EC52E6794EC167B22855DD" author="Duffy EAJ" page="67" year="1968">
<bibRefCitation id="80E71FC996C09932448514C206C96345" author="Duffy EAJ" firstAuthor="Duffy" page="67" refId="B5" refString="Duffy EAJ (1968) A monograph of the immature stages of Oriental timber beetles (Cerambycidae). British Museum (Natural History), London, [VIII +] 434 pp [+ 18 pls]." year="1968">Duffy 1968: 67</bibRefCitation>
</treatmentCitation>
[larva].
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2IAC" type="redescription">
<paragraph id="AD501238DC4523FF8C13036DCFBBD8F2">
<heading id="55E37C3DC746D156DC428648E7344CC0" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="8300827D1B7A4C5486F8D3747A0DD26B">
<emphasis id="BB3E2E101CC4C0CB40E495570C02A5B7" bold="true">Female.</emphasis>
Body testaceous to blackish-brown, clothed with off-white, yellowish-brown to greyish-yellow pubescence, with glabrous areas forming black spots. Pronotum clothed with greyish-white pubescence on middle of disc. Elytra clothed with pale yellowish-brown mixed with greyish-white pubescence and decorated with glabrous irregular black patches. Head rugosely punctate, with a median sulcus on vertex and occiput. Pronotum slightly wider than long between lateral spines, at about apical third distinctly constricted, with a coniform spine on middle of each side; anterior two protuberances on disc with rather steep anterior slope. Elytra about 1.8 times as long as humeral width, with sides slightly constricted at the basal third; disc coarsely punctate, with the punctures gradually becoming shallower towards apex. Legs moderately long, with femur slightly clavate.
</paragraph>
<caption id="18E4E54CB39DB7515B79D00411085413" ID-DOI="10.3897/zookeys.1226.139148.figure1" ID-arpha="07147D92-E71D-5A16-8DB2-120F64D46C80" httpUri="https://binary.pensoft.net/fig/1243454" startId="F1">
<paragraph id="BDDF79952EB532096FC372B848CDCC97">
<label id="CC1F36EBB3890CB79B20F5C322CFD0EB">Figure 1.</label>
</paragraph>
<paragraph id="D759F641DDB64DFD91F71A929FACF840">
Holotype of
<taxonomicName id="B9C5AD7099B42643E865D8CF4394F1A0" authority="Gahan, 1898 a" authorityName="Gahan" authorityYear="1898" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="A808748B922ED360B8158CD7EDA65721" italics="true">Apiocephalus punctipennis</emphasis>
<bibRefCitation id="8FD58FD76FB74E1F9B327B6537EFDF3C" DOI="10.1080/00222939808678011" author="Gahan CJ" refId="B6" refString="Gahan CJ (1898) Descriptions of new Longicorn Coleoptera from East Africa. The Annals and Magazine of Natural History Series 7, 2 (7): 40 - 59. https://doi.org/10.1080/00222939808678011" year="1898">Gahan, 1898</bibRefCitation>
<emphasis id="B89D62DD1DBA478EC129A17BBABCAEFB" bold="true">a</emphasis>
</taxonomicName>
dorsal view
<emphasis id="304F6BEC0D76CBF3278299AEC748AAB5" bold="true">b</emphasis>
lateral view
<emphasis id="6EB1B2D2EEFC253E1BD46D4E3EB95F3F" bold="true">c</emphasis>
ventral view
<emphasis id="290CCAA9B460533D4617D7A5DD539BBA" bold="true">d</emphasis>
frontal view
<emphasis id="E177F6581C6A42D0BD26B1800A1CD49C" bold="true">e</emphasis>
lateral view of pronotum
<emphasis id="45460AC0BA3D911B871640DB326F6B6D" bold="true">f</emphasis>
labels.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EHKAC" type="type material">
<paragraph id="D50DE26D431DC8484DF478AE12AF9CDF">
<heading id="21AB6774223AE920BA43A16BCC3FD8E6" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="53F35DFABA68560C9914753B4F2BEB31">
<emphasis id="7087875578C516F9D36961762E5DB83C" bold="true" italics="true">
<typeStatus id="C24AA35B1719781765B78FB0208B6A9A" type="holotype">Holotype</typeStatus>
</emphasis>
• female (
<named-content id="ECF23C46051CD945E985B9181203FCB9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London, UK">
<collectionCode id="B90726E34BD3F25BE2508B854E3F7204" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
): Type [p, label circular, red framed]; B. E. Africa. / C. S. Betton. / 9812. [p];
<taxonomicName id="53AFBE74865F1F7334A506EFF85D5AAF" authorityName="Gahan" authorityYear="1898" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="49CEF30CA216446433E0C84FFA6CCB8A" italics="true">Apiocephalus</emphasis>
</taxonomicName>
/
<taxonomicName id="20F63D501AA8634D3B98AFB12C83F7BF" authorityName="Gahan" authorityYear="1898" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="2CBBE711D13B17FDAC1031E719F2B592" italics="true">punctipennis</emphasis>
</taxonomicName>
/ Type / Gahan [h];
<collectionCode id="DC42E780B08DE47C85B677DF69D83D3F" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
014596632 [p] .
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGLAC" type="distribution">
<paragraph id="88BE8D249948D51FC48AB67287806ADE">
<heading id="E84492008FBDB83E85518A9F7B19FD41" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="0A91CD6DE9784B3ABA873E73DAB071CC">
<collectingCountry id="9D22794E9C4A88B3921F7255AFADE27C" name="Kenya">Kenya</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELLAC" type="remarks">
<paragraph id="A89EC08DCBCF3B883DDE604E9D5E0F69">
<heading id="2EFC05AF3141C54B35E213EC4B2064B3" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="1BE0983A1713F4A40CAD3415DF86642B">
Although
<bibRefCitation id="D9AC9B118F64B2F46DAE522E83291422" DOI="10.1080/00222939808678011" author="Gahan CJ" firstAuthor="Gahan" refId="B6" refString="Gahan CJ (1898) Descriptions of new Longicorn Coleoptera from East Africa. The Annals and Magazine of Natural History Series 7, 2 (7) : 4059. https://doi.org/10.1080/00222939808678011" year="1898">Gahan (1898)</bibRefCitation>
did not specify the gender of the
<typeStatus id="994F702EA56566CBE58442D7BB0600F5">type</typeStatus>
specimen in the original description, its antennal length and abdominal shape indicate that it is female. Additionally, based on the data uploaded by
<bibRefCitation id="B6BFD4478628A5F99280D8F091333EA9" author="Bezark LG" firstAuthor="Bezark" refId="B2" refString="Bezark LG (2013) Topic: Tanzania: Apiocephalus punctipennis. In Cerambycoidea Forum. https://www.cerambycoidea.com/forum/topic.asp?TOPIC_ID=16191 [accessed 29 November 2024]" url="https://www.cerambycoidea.com/forum/topic.asp?TOPIC_ID=16191" year="2013">Bezark (2013)</bibRefCitation>
on the website “
<tp_taxon-name id="B658412ECA745816D4CE5A39CB08FF38">
<tp_taxon-name-part id="78001C84985E805862659985A9FF2E7E" taxon-name-part-type="superfamily">Cerambycoidea</tp_taxon-name-part>
</tp_taxon-name>
”, this species occurs in
<collectingRegion id="17508D532C95C9F66F6D1CFFF5CA6E4F" country="Tanzania" name="Manyara">Manyara</collectingRegion>
,
<collectingCountry id="7F8C717AB611DD83848AAE4293C8D16D" name="Tanzania">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,508 @@
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<mods:title id="CF2CA7E05F511CBC7BFF11F7CB241F8E">Two new species and two new combinations within Paleosepharia (Coleoptera, Chrysomelidae, Galerucinae) from China</mods:title>
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<mods:namePart id="BD2D0633B4D02A4E153EA81E742E6E97">Feng, Chuan</mods:namePart>
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<mods:namePart id="A9DCCCA56C1524BF05104D07BA6E16E7">Yang, Xingke</mods:namePart>
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<mods:affiliation id="5BE6F73ACC29A1C82710FF0EEF98B92D">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Road, Chaoyang District, Beijing 100101, China</mods:affiliation>
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<mods:date id="42D995F64973315E930766B9AB2B687D">2025</mods:date>
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<treatment id="7F96DC36E5D05B83872FF61079C08F33" ID-arpha="7F96DC36-E5D0-5B83-872F-F61079C08F33" LSID="urn:lsid:plazi:treatment:7F96DC36E5D05B83872FF61079C08F33" httpUri="http://treatment.plazi.org/id/7F96DC36E5D05B83872FF61079C08F33">
<subSubSection id="5DE47528CD9ECA8E7574F916502989DB" type="nomenclature">
<paragraph id="F07E1BABBE614E07300CE334B0A1BB7B">
<taxonomicName id="D2158BE6D7B3FC8412AD99D314E0ED18" authority="(Chen, 1942)" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" baseAuthorityName="Chen" baseAuthorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bicavipennis" status="comb. nov.">
<emphasis id="396C5022A453671F7F2EC029C251A61A" italics="true">Paleosepharia bicavipennis</emphasis>
(
<bibRefCitation id="65280A88FBECA1697172F54E2A4741FB" author="Chen SH" journalOrPublisher="Notes dEntomologie Chinoise" pagination="9-67" refId="B6" refString="Chen SH (1942) Galerucinae nouveaux de la faune chinoise. Notes d'Entomologie Chinoise 9: 9-67." title="Galerucinae nouveaux de la faune chinoise." volume="9" year="1942">Chen, 1942</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0632EB834F43C52A0E6FC77663AB4CFB" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D7859E1A24B364B1A408347F79A311E0" type="description">
<paragraph id="0088C884A69FDBBA8CB2E638B6A392AD">
<figureCitation id="6FE62F5545C92087D8F4F5202CDB97BD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paleosepharia bicavipennis (Chen, 1942), comb. nov. (holotype; male) A C habitus. A dorsal view B ventral view C lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1226.140239.figure5" httpUri="https://binary.pensoft.net/fig/1243036">Figs 5</figureCitation>
,
<figureCitation id="CAFA5D89EDC8C323B861F16D27922E92" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paleosepharia bicavipennis (Chen, 1942), comb. nov. (male) A C habitus D F aedeagus G depression of the elytra of the male. A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1 mm (A C); 0.5 mm (D F)." figureDoi="10.3897/zookeys.1226.140239.figure6" httpUri="https://binary.pensoft.net/fig/1243037">6</figureCitation>
,
<figureCitation id="0F63F61D8755B4F8882F0E2044DB9AC8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paleosepharia bicavipennis (Chen, 1942), comb. nov. (female) A C habitus D vaginal palps E spermatheca F apex of abdomen of the female. A dorsal view B ventral view C lateral view. Scale bars: 1 mm (A C); 0.5 mm (D, E)." figureDoi="10.3897/zookeys.1226.140239.figure7" httpUri="https://binary.pensoft.net/fig/1243038">7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7841B3AC267B82AA1863C0154DC1A74" type="reference_group">
<paragraph id="07127AFBB70A73A68CF89288FD10A221">
<treatmentCitationGroup id="FA4AB82FF0C43B7209634D99B5978050">
<treatmentCitation id="C410E3664F2DF66C8125DD7583970D0A" author="Chen SH" page="55" year="1942">
<taxonomicName id="B82B9D729E56826956E959A38C5C7D47" authority="Chen, 1942: 55" authorityName="Chen" authorityPageNumber="55" authorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bicavipennis">
<emphasis id="08A3E91A6467B5D1D2A8C3FDBECEBA99" italics="true">Monolepta bicavipennis</emphasis>
<comment id="E7C3CD8CB6F61657B862F51A678243A7">
<bibRefCitation id="E38C91889518BB688F37F3C4A8859AF6" author="Chen SH" journalOrPublisher="Notes dEntomologie Chinoise" pagination="9-67" refId="B6" refString="Chen SH (1942) Galerucinae nouveaux de la faune chinoise. Notes d'Entomologie Chinoise 9: 9-67." title="Galerucinae nouveaux de la faune chinoise." volume="9" year="1942">Chen, 1942: 55</bibRefCitation>
.
<abbrev id="ABBRID0E1OAE" xlink_title="type locality">TL</abbrev>
<collectingCountry id="62D2A73E018E896E189A8CB52D40B820" name="China">China</collectingCountry>
:
<collectingRegion id="7D6C6AF6EF4D439259970B9689C01A67" country="China" name="Shaanxi">Shaanxi</collectingRegion>
.
<abbrev id="ABBRID0E5OAE" xlink_title="type deposition">TD</abbrev>
<named-content id="764345DE0269C4D39EAAA4DF3F6659D2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="E42CDE5492B16A1C78D830B2EBA11BD3" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFPAE" type="material">
<paragraph id="BD2B0D7E284E50683BECFC4C3EFE9902">
<heading id="2958EC8BE151D9B9F0E652B625BED2F4" reason="title">Type specimens examined.</heading>
</paragraph>
<paragraph id="F8A96EED9FDB17C95A2714D10BD04514">
<emphasis id="1FAFE82152E4BB8EA640DF28BF0B76D4" bold="true" italics="true">
<typeStatus id="C838502100030295E7913CBE73C92841" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCount id="F004C43B7C147CF53AEF8E5BB448129F" count="1" type="male"></specimenCount>
Shensi, Wei-cze-ping,
<date id="D9B5775595CAB933C4FD20BDE351B281" value="1916-08-18">18 Aug. 1916</date>
, P. Licent.
<materialsCitation id="BE06CE55A59574F200F05055C8EC83BD" collectingDate="1947-08-26" collectionCode="IZCAS" country="China" location="Mount Tianmu" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Zhejiang" typeStatus="paratype">
<emphasis id="A7FCD1137AE39B9F85A52E128445A559" bold="true" italics="true">
<typeStatus id="F1E82A41E2FB7E287EC9C59704CB2D65" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCount id="CFE1D33E09831F88CB7F70944F1CB016" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="118FF824A99F76CCD812506DECC21DAD" count="2" type="female">2 ♀♀</specimenCount>
,
<collectingCountry id="C8C2DFC1A59FE7AAAC9B6F5B05074CC4" name="China">China</collectingCountry>
,
<collectingRegion id="DCA3835591BE7C1FC3C74A113EACF79C" country="China" name="Zhejiang">Zhejiang Province</collectingRegion>
,
<location id="C8CAD89C8F52E967E39840EF80A5C883" LSID="urn:lsid:plazi:treatment:7F96DC36E5D05B83872FF61079C08F33:C8CAD89C8F52E967E39840EF80A5C883" country="China" name="Mount Tianmu" stateProvince="Zhejiang">Mount Tianmu</location>
,
<date id="61E24BB5EF03C13DCF687EC520FA4044" value="1947-08-26">
<collectingDate id="C6AEFAAEFA07A061B79BC0055C0C8A5C" value="1947-08-26">26 Aug. 1947</collectingDate>
</date>
,
<named-content id="044C9C0E281C3E5F00B9FB322F77BAEE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="55989A56657183F99533D566B6F9C2F1" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="DE126B12C2B030B04A03CE34045ABE38" collectingDate="1947-08-31" collectionCode="IZCAS" country="China" location="Mount Tianmu" specimenCount="2" specimenCount-female="2" stateProvince="Zhejiang" typeStatus="paratype">
<specimenCount id="3C4D6AA651EE2422D51B4E2633261CA7" count="2" type="female">2 ♀♀</specimenCount>
,
<collectingCountry id="371076C1694DF381765BEBF8D5B2ED58" name="China">China</collectingCountry>
,
<collectingRegion id="B9155BA07C00C60C223D129DFEB5B649" country="China" name="Zhejiang">Zhejiang Province</collectingRegion>
,
<location id="00EA1D6E94E1A562BBE0B0AD266B7FE2" LSID="urn:lsid:plazi:treatment:7F96DC36E5D05B83872FF61079C08F33:00EA1D6E94E1A562BBE0B0AD266B7FE2" country="China" name="Mount Tianmu" stateProvince="Zhejiang">Mount Tianmu</location>
,
<date id="44F62146AE64069CE41AD8450DBC9CFC" value="1947-08-31">
<collectingDate id="DEED3D732EB89108A0602038E3B00BF7" value="1947-08-31">31 Aug. 1947</collectingDate>
</date>
,
<named-content id="E991502C52155777692C7DBD82FF26DB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="35E528467387745151BF9EC5B756B0A1" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<caption id="2ADFEFB4B19935574055D1356F25495E" ID-DOI="10.3897/zookeys.1226.140239.figure5" ID-arpha="6C8BA7B5-516D-5892-A53C-5EC9384DB948" httpUri="https://binary.pensoft.net/fig/1243036" startId="F5">
<paragraph id="F14ECC7AC9A61B1A0A6A370EAE0E5DDF">
<label id="AEFDF7CFBDC8CB38FF5513E13EF79D94">Figure 5.</label>
</paragraph>
<paragraph id="8815AD389D13686C1AEB9EB1D9E1CDB1">
<taxonomicName id="D9AB624D7A40955B5E3E5354A6D1F062" authority="(Chen, 1942)" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" baseAuthorityName="Chen" baseAuthorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bicavipennis" status="comb. nov.">
<emphasis id="8D9A76C05030E944A0FAC49EFEF3CB0E" italics="true">Paleosepharia bicavipennis</emphasis>
(
<bibRefCitation id="59EE42CE508A9B837AFBD94C8BC68080" author="Chen SH" journalOrPublisher="Notes dEntomologie Chinoise" pagination="9-67" refId="B6" refString="Chen SH (1942) Galerucinae nouveaux de la faune chinoise. Notes d'Entomologie Chinoise 9: 9-67." title="Galerucinae nouveaux de la faune chinoise." volume="9" year="1942">Chen, 1942</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="E69773FF4D78E99E9BF987B2F9FF8B7B" rank="species">comb. nov.</taxonomicNameLabel>
(holotype; male)
<emphasis id="58C8A62AC011AAD05F324AC8ACFF20DF" bold="true">A C</emphasis>
habitus.
<emphasis id="2CAB9CB5B1E772E18E05F697EBEB2568" bold="true">A</emphasis>
dorsal view
<emphasis id="B2C4DC03BFEDE2309DCBA678C888FC6F" bold="true">B</emphasis>
ventral view
<emphasis id="AA8745FF06D6EFEFD685A3B73981FB5F" bold="true">C</emphasis>
lateral view. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E2QAE" type="material">
<paragraph id="A26F940AB054169F2D3C220D402EAC60">
<heading id="59291D1C835835490E89E16BA4B8DBCD" reason="title">Other specimens examined.</heading>
</paragraph>
<paragraph id="7DC18D12F48FED734D21DD14ADEC7113">
<specimenCount id="DF9229E77DF7EE1E4724424DD9045707" count="7" type="male">7 ♂♂</specimenCount>
, •
<materialsCitation id="86F35074BFBE27E0B8A4FBFB6F41FA06" collectingDate="2002-07-19" collectionCode="IZCAS" collectorName="Lijie Zhang" country="China" county="Song County" elevation="1600" location="Mount Baiyun" specimenCount="1" stateProvince="Henan">
<collectingCountry id="EBB20E0E53A05D6B60213F51003EF0AB" name="China">China</collectingCountry>
,
<collectingRegion id="F2D9B11798492310393B9CEFE6644CBD" country="China" name="Henan">Henan Province</collectingRegion>
,
<collectingCounty id="A85C8DC4A499342CDD6FE56220A509B7">Song County</collectingCounty>
,
<location id="DC2B47245BB91FAE3447C12308A4738F" LSID="urn:lsid:plazi:treatment:7F96DC36E5D05B83872FF61079C08F33:DC2B47245BB91FAE3447C12308A4738F" country="China" county="Song County" name="Mount Baiyun" stateProvince="Henan">Mount Baiyun</location>
,
<elevation id="3BF5051B141A8AC5F094D84A3FE255B4" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<quantity id="5A066737BDF8E0BFF11F5BE446974B32" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</quantity>
a. s. l.
</elevation>
,
<date id="CFEDEFB9C40EE40B86DEFA1C319EBD7B" value="2002-07-19">
<collectingDate id="8D1EFB82A0D71A163AE2E4E07D7E82D3" value="2002-07-19">19 Jul. 2002</collectingDate>
</date>
,
<collectorName id="837553B71B93871AE9A8175F00022C6A">Lijie Zhang</collectorName>
leg.,
<named-content id="B6F7FF3B54C22C6991FEE861B647F39D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="DF313BF8D996BD2705535EC7E8F75F28" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="BE6DFB828CB65650FAC31C6A88FD2422" collectingDate="1998-07-25" collectionCode="IZCAS" collectorName="Jun Chen" country="China" elevation="935" location="Yaohe" municipality="Foping" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Shaanxi">
<specimenCount id="9511219A98923FF4F125351A948F2C64" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="E3567303A66AE212D9268EEDAA6514F5" count="5" type="female">5 ♀♀</specimenCount>
,
<collectingCountry id="37C9D45E87E09A8B839F0BE57150C9B9" name="China">China</collectingCountry>
,
<collectingRegion id="4BFC3AA3C4D782E0FE68540004E4BD35" country="China" name="Shaanxi">Shaanxi Province</collectingRegion>
,
<collectingMunicipality id="B5138B8E2E8A6F0C1859FFF0AECB8C61">Foping</collectingMunicipality>
,
<location id="287080C253DFA55E79198F7A2ACB8F15" LSID="urn:lsid:plazi:treatment:7F96DC36E5D05B83872FF61079C08F33:287080C253DFA55E79198F7A2ACB8F15" country="China" municipality="Foping" name="Yaohe" stateProvince="Shaanxi">Yaohe</location>
,
<elevation id="7C45511BA2DA3F41617D82CA2450221A" metricMagnitude="2" metricUnit="m" metricValue="9.35" metricValueMax="10.0" metricValueMin="8.7" unit="m" value="935.0" valueMax="1000.0" valueMin="870.0">
<quantity id="86FE41FD28973C58E14DCF0BFCF2A445" metricMagnitude="2" metricUnit="m" metricValue="9.35" metricValueMax="10.0" metricValueMin="8.7" unit="m" value="935.0" valueMax="1000.0" valueMin="870.0">8701000 m</quantity>
a. s. l.
</elevation>
,
<date id="9EAC3BB7EAB1EFEEA79C1FC76D920E81" value="1998-07-25">
<collectingDate id="BD4771341999DB73F716EA6E7796D75B" value="1998-07-25">25 Jul. 1998</collectingDate>
</date>
,
<collectorName id="357FFBB7D0C24E527C87CA8CC1F58838">Jun Chen</collectorName>
leg.,
<named-content id="954F5DCCFCDD13F14511F00F02DB80CC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="C5C2C166DDA0C2C4704D7E387CE67498" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="35DC86E23A72E4EE67C440AF1E7721B8" collectingDate="1999-07-28" collectionCode="IZCAS" collectorName="Jian Yao" country="China" county="Wen County" elevation="1220" location="Bifenggou" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Gansu">
<specimenCount id="05A611D95C5954189FE1197E66DFD274" count="5" type="male">5 ♂♂</specimenCount>
<specimenCount id="0D1ACD239D1E3121E762BCF200994CA1" count="1" type="female">1 ♀</specimenCount>
,
<collectingCountry id="071D5F2642642C983741FC1313362F34" name="China">China</collectingCountry>
,
<collectingRegion id="93A7A35ABAFB82A283B8CF5F90FB16B2" country="China" name="Gansu">Gansu Province</collectingRegion>
,
<collectingCounty id="BCC6B98CBA484C09688C08CC272032F1">Wen County</collectingCounty>
,
<location id="D39FBE9B9C6FF4E138B0CE5B95593E78" LSID="urn:lsid:plazi:treatment:7F96DC36E5D05B83872FF61079C08F33:D39FBE9B9C6FF4E138B0CE5B95593E78" country="China" county="Wen County" name="Bifenggou" stateProvince="Gansu">Bifenggou</location>
,
<elevation id="C6B009096E10469D30BE9072304AE9FE" metricMagnitude="3" metricUnit="m" metricValue="1.22" metricValueMax="1.5" metricValueMin="0.94" unit="m" value="1220.0" valueMax="1500.0" valueMin="940.0">
<quantity id="69DA0791544878EDB9B1ACE6212DD8EC" metricMagnitude="3" metricUnit="m" metricValue="1.22" metricValueMax="1.5" metricValueMin="0.94" unit="m" value="1220.0" valueMax="1500.0" valueMin="940.0">9401500 m</quantity>
a. s. l.
</elevation>
,
<date id="C312FD3F815578E41417930DF2410F14" value="1999-07-28">
<collectingDate id="2D128CE8FD2BA0A9C4B1CFDBEB52F1F1" value="1999-07-28">28 Jul. 1999</collectingDate>
</date>
,
<collectorName id="BDA1C2F254BC91340004D5C0109A869A">Jian Yao</collectorName>
leg.,
<named-content id="8D0D2ED5F475A57DEAE1327CECB6F902" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="41BCE613150CDE915806DE75B562120B" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="5D37F1CDE468B344A951CCE04F2BF5EB" collectingDate="1980-07-18" collectionCode="IZCAS" collectorName="Peiyu Yu" country="China" elevation="950" location="Songbai" municipality="Shennongjia" specimenCount="11" specimenCount-female="1" specimenCount-male="10" stateProvince="Hubei">
<specimenCount id="56E5CD0ED6316CCC57755040CA7BD14B" count="10" type="male">10 ♂♂</specimenCount>
<specimenCount id="9F1A953C53CBF3424E1F8C8EEEBA1980" count="1" type="female">1 ♀</specimenCount>
,
<collectingCountry id="24D50DE97BEF545E51AA30C515D4A8BF" name="China">China</collectingCountry>
,
<collectingRegion id="B7A7C1477C545A1281285BA8372D12D4" country="China" name="Hubei">Hubei Province</collectingRegion>
,
<collectingMunicipality id="A4AB7C54F31C20CCB388467A200D4463">Shennongjia</collectingMunicipality>
,
<location id="EF5BC80B44BA68703002E2E98E91A2EA" LSID="urn:lsid:plazi:treatment:7F96DC36E5D05B83872FF61079C08F33:EF5BC80B44BA68703002E2E98E91A2EA" country="China" municipality="Shennongjia" name="Songbai" stateProvince="Hubei">Songbai</location>
,
<elevation id="ED298B67CF824E0F7C1F02733601B2C8" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">
<quantity id="88FEF0599875707B54776394A5748B2E" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">950 m</quantity>
a. s. l.
</elevation>
,
<date id="A3EEE1F1110B5298B0827F688F5ECF82" value="1980-07-18">
<collectingDate id="657689BEB11E167833EF0DEAC94B58F3" value="1980-07-18">18 Jul. 1980</collectingDate>
</date>
,
<collectorName id="C54837A1CC23A61F819DD04815344BE5">Peiyu Yu</collectorName>
leg.,
<named-content id="151D740D4DD5AAC4DB5F7307E22D8220" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="7ACBC5421BFBDAC8D46711372B0D514D" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="F3FDF766BEE4170B027FDDA168E51886" collectingDate="1988-08-07" collectionCode="IZCAS" collectorName="Shuyong Wang" country="China" elevation="710" location="Yongshun" specimenCount="9" specimenCount-female="9" stateProvince="Hunan">
<specimenCount id="33244D235F83FE4E83C35391B1B012F6" count="9" type="female">9 ♀♀</specimenCount>
,
<collectingCountry id="BD0A8CBAE31D4C1E6D5823DCA22AAC6E" name="China">China</collectingCountry>
,
<collectingRegion id="6AA9BF69B9356D718A7BD639201BEDA1" country="China" name="Hunan">Hunan Province</collectingRegion>
,
<location id="2C134D950F83B699745AE126EA4E405C" LSID="urn:lsid:plazi:treatment:7F96DC36E5D05B83872FF61079C08F33:2C134D950F83B699745AE126EA4E405C" country="China" name="Yongshun" stateProvince="Hunan">Yongshun</location>
,
<elevation id="C857F40A315EB9AF668471E68B252AD1" metricMagnitude="2" metricUnit="m" metricValue="7.1" metricValueMax="8.2" metricValueMin="6.0" unit="m" value="710.0" valueMax="820.0" valueMin="600.0">
<quantity id="027BF3FE577069193A962A8A7A0BF70A" metricMagnitude="2" metricUnit="m" metricValue="7.1" metricValueMax="8.2" metricValueMin="6.0" unit="m" value="710.0" valueMax="820.0" valueMin="600.0">600820 m</quantity>
a. s. l.
</elevation>
,
<date id="0F9AC549E71AC822FC6EF042EB46F776" value="1988-08-07">
<collectingDate id="B10D950C7DA49A0802D51DFCA18A2BAC" value="1988-08-07">7 Aug. 1988</collectingDate>
</date>
,
<collectorName id="DCFC1274D8266ECF4B655E557E265551">Shuyong Wang</collectorName>
leg.,
<named-content id="26707EEC58508AEDE851918E6A7DE9AC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="3EC964BA97DCEF62643246F12D7DEB5A" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="5695D5A93CD6F27EC4A6FCC16FE3CE81" collectingDate="1981-07-10" collectionCode="IZCAS" collectorName="Shuyong Wang" country="China" county="Weixi County" elevation="1780" location="Baijixun" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Yunnan">
<specimenCount id="C58A82F84949000E80AE2434D9F3879A" count="5" type="male">5 ♂♂</specimenCount>
<specimenCount id="48088E390DB84973DF2F3B5F16827973" count="1" type="female">1 ♀</specimenCount>
,
<collectingCountry id="2D9E604050BD3B6B0D9351EC0385DED0" name="China">China</collectingCountry>
,
<collectingRegion id="BE2AFD0D2764CBAA1DA46C6EA2BF1573" country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingCounty id="F1D2A46DA5646CC220EEC3B9598A6D40">Weixi County</collectingCounty>
,
<location id="AC37F881C990F10D48143E86B336CDA7" LSID="urn:lsid:plazi:treatment:7F96DC36E5D05B83872FF61079C08F33:AC37F881C990F10D48143E86B336CDA7" country="China" county="Weixi County" name="Baijixun" stateProvince="Yunnan">Baijixun</location>
,
<elevation id="FBD4C192899853CD165A60FEC86F5098" metricMagnitude="3" metricUnit="m" metricValue="1.78" unit="m" value="1780.0">
<quantity id="29E31FC5906BCFDF6C02CCC03E005C5E" metricMagnitude="3" metricUnit="m" metricValue="1.78" unit="m" value="1780.0">1780 m</quantity>
a. s. l.
</elevation>
,
<date id="586C3CFC647099072252DBC89EC83621" value="1981-07-10">
<collectingDate id="4AB3A6FEAE1CF111DF8BB70490F24C76" value="1981-07-10">10 Jul. 1981</collectingDate>
</date>
,
<collectorName id="39A18AA579E2BDC7F3BE7FB46CD19F95">Shuyong Wang</collectorName>
leg.,
<named-content id="2AF3815505C351FD590E26DDB4BAC543" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="6E184DED9692E42F877F708FFE814AB3" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<caption id="FB3F61795BD7B8FEFBF3C00ABB120B3C" ID-DOI="10.3897/zookeys.1226.140239.figure6" ID-arpha="7DB8EFCA-D883-577E-B12E-ED321EAF7E57" httpUri="https://binary.pensoft.net/fig/1243037" startId="F6">
<paragraph id="899063F5CFFE5E5FCBE66FCE00EC31B2">
<label id="2D6AF1E3EAFFAE04AAE2CC42164DE6EB">Figure 6.</label>
</paragraph>
<paragraph id="6AA42C10FA3778F85340DBC1EE01EEDD">
<taxonomicName id="67A7A0E7BBECF41FC7DD18F809ACB43E" authority="(Chen, 1942)" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" baseAuthorityName="Chen" baseAuthorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bicavipennis" status="comb. nov.">
<emphasis id="CA70E4A9C6D9CE4280AA177587A9D8A3" italics="true">Paleosepharia bicavipennis</emphasis>
(
<bibRefCitation id="9D7377E08A707D51EEA6DBDEAFCDAF93" author="Chen SH" journalOrPublisher="Notes dEntomologie Chinoise" pagination="9-67" refId="B6" refString="Chen SH (1942) Galerucinae nouveaux de la faune chinoise. Notes d'Entomologie Chinoise 9: 9-67." title="Galerucinae nouveaux de la faune chinoise." volume="9" year="1942">Chen, 1942</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="B8CDC3103289DB2075D8757D91F70CAF" rank="species">comb. nov.</taxonomicNameLabel>
(male)
<emphasis id="81A7FCED3CD2EFFCD39D69BEFE02896A" bold="true">A C</emphasis>
habitus
<emphasis id="A5D888F2067087F021F2BAF3877AF41D" bold="true">D F</emphasis>
aedeagus
<emphasis id="D12740CBBF5D8DDAE67D9933EEE8F821" bold="true">G</emphasis>
depression of the elytra of the male.
<emphasis id="1586CD68D157C97A53C86AE7D0D0FE46" bold="true">A, D</emphasis>
dorsal view
<emphasis id="01827E59D1646D0BFF1B6A9954FB37D1" bold="true">B, E</emphasis>
ventral view
<emphasis id="99E65B4279E320D4B4EC1A4A12B491D7" bold="true">C, F</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="32CDEDAF811151149924E114B2A58A42" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="CA089FFC7405F0538F9022D316C2B51E" bold="true">D F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EBTAE" type="notes">
<paragraph id="749F56B4EEB1819C805F4D9122A76510">
<heading id="8954C6B27F8E080AD09DC825B4AFAD8B" reason="title">Notes.</heading>
</paragraph>
<paragraph id="2DC394003676D74EF3609576670563ED">
Based on the following characters of this species, which are characteristic for
<taxonomicName id="8D1A53E077F8D9C727A795AA6768A0CA" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="D527A16BB49EC67E4E5BDB53407A03BC" italics="true">Paleosepharia</emphasis>
</taxonomicName>
, it is transferred from
<taxonomicName id="D5882A39A4F867518617F8EFD596BBA9" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Monolepta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="18A090967C58B11AE04D031173C8F9C2" italics="true">Monolepta</emphasis>
</taxonomicName>
to
<taxonomicName id="C160B898D86C5B5310DDDD3B1D96B0FD" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="F03DF873F3C0111DD06E6276827BAC87" italics="true">Paleosepharia</emphasis>
</taxonomicName>
: the third antennomere is longer than the second; depression in the elytra; median lobe parallel-sided in the basal two-thirds, apical third strongly narrowed; and tectum long, truncate at apex.
</paragraph>
<caption id="13DB2260848878B88573EB9A004765B6" ID-DOI="10.3897/zookeys.1226.140239.figure7" ID-arpha="5CEF2F91-C600-55D2-9A6C-2B1D5B07A6B4" httpUri="https://binary.pensoft.net/fig/1243038" startId="F7">
<paragraph id="6C3D5E28684A220047E74E5C39FE5EFD">
<label id="2453E5D60A0F0C8D1B349A64E95CDC05">Figure 7.</label>
</paragraph>
<paragraph id="4ACCB77D541493993CFE1EB7C0D85EA7">
<taxonomicName id="C7721F74F6A15F8D0D4DC706010CE96D" authority="(Chen, 1942)" authorityName="Feng &amp; Yang &amp; Ge" authorityYear="2025" baseAuthorityName="Chen" baseAuthorityYear="1942" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bicavipennis" status="comb. nov.">
<emphasis id="187ED186D0480024F0BD48AC6E5B181D" italics="true">Paleosepharia bicavipennis</emphasis>
(
<bibRefCitation id="D3DDFBBACD5FDEBE51940FF89EAEE009" author="Chen SH" journalOrPublisher="Notes dEntomologie Chinoise" pagination="9-67" refId="B6" refString="Chen SH (1942) Galerucinae nouveaux de la faune chinoise. Notes d'Entomologie Chinoise 9: 9-67." title="Galerucinae nouveaux de la faune chinoise." volume="9" year="1942">Chen, 1942</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BEF46A3482BC4863FF3C4DB057907597" rank="species">comb. nov.</taxonomicNameLabel>
(female)
<emphasis id="4920E28D0D20465C2CD7CB0AF3F4BD29" bold="true">A C</emphasis>
habitus
<emphasis id="E7F14FBA551056D1C84011DAD12A531D" bold="true">D</emphasis>
vaginal palps
<emphasis id="8F2032BB8187D53C375408FD9E773A44" bold="true">E</emphasis>
spermatheca
<emphasis id="950C0EE079CA88F469F96E0CD9DDEA78" bold="true">F</emphasis>
apex of abdomen of the female.
<emphasis id="8F147C77ED3668D300F97764074080BB" bold="true">A</emphasis>
dorsal view
<emphasis id="AF44D8A2A8ADB7AFD2F3AAE339DB6400" bold="true">B</emphasis>
ventral view
<emphasis id="C4F2B6BBA732B92170D8B0FA34077A9C" bold="true">C</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="7DF57FCAF5E5212BBD417C1B6FB19074" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="7292CFF10F1C52D22B81E2C5620748C3" bold="true">D, E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EGVAE" type="diagnosis">
<paragraph id="0D75A2E00C15ED49DA4704CABF324B9E">
<heading id="59A69C55D00A34C4184F64B624115C69" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="40D968F52E1FBB0E5EF4C1D8CDBF4E77">This species is easy to distinguish from other species. The elytra is red or yellow-brown, with two comma-shaped deep concavities and two cylindrical protrusions in the middle, and cylindrical protrusions with short hairs.</paragraph>
</subSubSection>
<subSubSection id="SECID0ELVAE" type="redescription">
<paragraph id="AA336F0E5D11E4BDD3A8E64AF519B752">
<heading id="773AB6F3D82588D3A528716910DA6515" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="36468106D28CA555498AB31F6762E820">
<emphasis id="0AFC5CB02A39432B61C03BDFB8460A32" bold="true">Male</emphasis>
: Length:
<quantity id="B7257366313ED917C9C33583382935E1" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.9" metricValueMin="4.2" unit="mm" value="4.55" valueMax="4.9" valueMin="4.2">4.24.9 mm</quantity>
, width:
<quantity id="7007C1F05BB1A0582F90A3D76E472615" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" unit="mm" value="2.4" valueMax="2.6" valueMin="2.2">2.22.6 mm</quantity>
(mean length
<quantity id="550BE07A7300CE4726684545A0A5474B" metricMagnitude="-3" metricUnit="m" metricValue="4.6" unit="mm" value="4.6">4.6 mm</quantity>
, mean width
<quantity id="14446876C56F9AA7BB4446BB6DF12BC4" metricMagnitude="-3" metricUnit="m" metricValue="2.4" unit="mm" value="2.4">2.4 mm</quantity>
,
<emphasis id="B8C0D2B1B8E7A6449F507917581ED18D" italics="true">N</emphasis>
= 30). General color yellow-brown or red, head and pronotum black, antennae brown, legs dark brown, ventral side of hind femora yellow-brown, in some specimens, legs largely brown with most tibiae darkened. Vertex with sparse punctures. Frontal tubercles transverse, extending downward between antenna bases. Antennae slender, 0.95 × as long as body. First antennomere shiny bare, rod-shaped, second to eleventh antennomeres with short hairs, second antennomere shortest, third antennomere about 1.8 × as long as second; fourth antennomere about 1.5 × as long as third, fifth to eleventh antennomeres equal in length to fourth. Pronotum about 1.6 × as wide as long, lateral margins slightly curved, basal border slightly convex, apical border slightly concave, disc strongly convex, with lateral fovea and dense minute punctures. Scutellum triangular, smooth, impunctate. Elytra wider than pronotum, humeri convex. Each elytron with comma-shaped deep concavity and cylindrical protrusions in middle, cylindrical protrusions with short hairs. In some specimens, longitudinal depression end of concavity elongated or shortened, Disc with dense, minute punctures. Elytral epipleuron broad at base, gradually narrowed from middle to apex. Each tibia with distinct spur at apex, segment 1 of hind tarsi longer than remaining segments combined. Aedeagus robust, gradually widened towards subapex, triangle at apex, moderately recurved at apex in lateral view. Lateral spiculae short, ventral spiculae comb-shaped, and median spiculae long.
</paragraph>
<paragraph id="6ACF9D02383F9E3F6F21F478988BB55A">
<emphasis id="DB6A4912E5C065322BB7DF4B03F713D1" bold="true">Female</emphasis>
: Length: 4.25.0 mm, width: 2.0
<quantity id="A95BE4D332CA0379E261FDFDC4CD01E8" metricMagnitude="-3" metricUnit="m" metricValue="2.6" unit="mm" value="2.6">2.6 mm</quantity>
(mean length
<quantity id="D423A344B07A84650C740D1EDCF43DBF" metricMagnitude="-3" metricUnit="m" metricValue="4.7" unit="mm" value="4.7">4.7 mm</quantity>
, mean width
<quantity id="0E82CD682521B99042FDC14B044857D3" metricMagnitude="-3" metricUnit="m" metricValue="2.4" unit="mm" value="2.4">2.4 mm</quantity>
,
<emphasis id="93D504AC35610DCE750D2F9635A51A47" bold="true" italics="true">N</emphasis>
= 21).
</paragraph>
<paragraph id="E9959544A54AEA0CF7AE9085D5DF2971">Third antennomere about 1.5 × as long as second; fourth antennomere about 1.6 × as long as third. Elytra without short hairs and depressions. Spermatheca with big and rounded nodulus, middle part short, and cornu slightly curved. Vaginal palps with wide base, anteriorly with subtriangular tip, each palp slightly narrowing posteriorly, with pointed apex, 5 setae placed at apex, additional 2 setae subapically.</paragraph>
</subSubSection>
<subSubSection id="SECID0E4VAE" type="distribution">
<paragraph id="6BDF4C669C34D9925ED378B465791471">
<heading id="2C481ABFA3C03714C24EE660D23E0633" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="2D6CA968E0F378614C0E769EDC476EE3">
<collectingCountry id="114EDB8353CEEAB440CE902A31B211E6" name="China">China</collectingCountry>
:
<collectingRegion id="A28CA42592D01C391AD195B1CADDEA21" country="China" name="Gansu">Gansu</collectingRegion>
,
<collectingRegion id="8C77C4F75B11760363130E6214F31794" country="China" name="Shanxi">Shanxi</collectingRegion>
,
<collectingRegion id="0CE88BD06C6F47104AB3E6F565BE2E49" country="China" name="Shaanxi">Shaanxi</collectingRegion>
,
<collectingRegion id="6BE2CDE3EA5E48EE3CCF30820B2944DA" country="China" name="Henan">Henan</collectingRegion>
,
<collectingRegion id="FC157F824E9E27C5FBABB4AEED2F2C62" country="China" name="Anhui">Anhui</collectingRegion>
,
<collectingRegion id="06D6523204ADE8DFCF33314899FBE44B" country="China" name="Zhejiang">Zhejiang</collectingRegion>
,
<collectingRegion id="15537E2566ED774578C763A7A16D8AD0" country="China" name="Hubei">Hubei</collectingRegion>
,
<collectingRegion id="B2449315842AFBAF5B729C2D97621017" country="China" name="Jiangxi">Jiangxi</collectingRegion>
,
<collectingRegion id="2F3B107F1BA37CA8AB2A0170BBCC4EC3" country="China" name="Hunan">Hunan</collectingRegion>
,
<collectingRegion id="028E304A20A89C44746364609DF8FF5A" country="China" name="Guangxi">Guangxi</collectingRegion>
,
<collectingRegion id="A04300023F41C87CC2F294E2CFE67966" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingRegion id="6560D77B63922E2B3F58FB9C21D99149" country="China" name="Yunnan">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,308 @@
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<mods:title id="415D5C3BD2433110571F004555B0E3E1">Reniscymnus gen. nov., a new genus of Scymnini (Coleoptera, Coccinellidae) from the Oriental region</mods:title>
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<mods:namePart id="B3F40E484850E5CFC23160405B2B9F37">Peng, Feng</mods:namePart>
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<mods:classification id="B00ED44AF814BD6F48C3D2B07BE7A155">journal article</mods:classification>
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<mods:identifier id="D7AA65D9E3A93231077CAD27FDF69903" type="DOI">10.3897/zookeys.1226.130352</mods:identifier>
<mods:identifier id="A4C886FBE4273273B986DB7803B8BE5F" type="GBIF-Dataset">4de822ec-bfa2-4153-936f-41c90f741a75</mods:identifier>
<mods:identifier id="00646019BDF8711692602986E7A4A29C" type="ZooBank">91F3BBBF-E7D2-404B-B8DF-7210ECA9D76A</mods:identifier>
</mods:mods>
<treatment id="8EE0B488CC195C47AEFE070527DD1C64" ID-DOI="http://doi.org/10.5281/zenodo.14827182" ID-Zenodo-Dep="14827182" ID-arpha="8EE0B488-CC19-5C47-AEFE-070527DD1C64" ID-zoobank="https://zoobank.org/0BF770A6-FD04-4675-BDCC-271E0124A9FD" LSID="urn:lsid:plazi:treatment:8EE0B488CC195C47AEFE070527DD1C64" LSID-ZBK="urn:lsid:zoobank.org:act:0BF770A6-FD04-4675-BDCC-271E0124A9FD" httpUri="http://treatment.plazi.org/id/8EE0B488CC195C47AEFE070527DD1C64">
<subSubSection id="1852CFA9E2F64D87885030BFBDD8D1DD" type="nomenclature">
<paragraph id="F11D16F9FD6C09BCE1318FE91642E6A7">
<taxonomicName id="77154E680D81DBD0218F3385BAFA7543" authority="Peng &amp; Chen" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="explanatus" status="sp. nov.">
<emphasis id="09EF76E401D92ED2A34749D5F38A5313" italics="true">Reniscymnus explanatus</emphasis>
Peng &amp; Chen
</taxonomicName>
<taxonomicNameLabel id="28E86F171A89B14C73A5CF0BF08FF13B" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D246FA6980CD739FC206D33F93970649" type="description">
<paragraph id="16A396FFAABEBC539160C76C1EFF7A72">
<figureCitation id="91F04316AD00EE00D5B8910DD108803E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYDAG" type="diagnosis">
<paragraph id="97B504D94D9814835F9C64C2DF7C09DF">
<heading id="D950CB7AC8B8F0844A458D6B8289BF48" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="E3EA3A09B21823959D426C16BFC6B06E">
This species is similar to
<taxonomicName id="2629814D50E70B245F3E5ED2B7DF2D27" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cordatus" status="sp. nov.">
<emphasis id="AFC652BCE8C1DFFE0390DF4B7966FF00" italics="true">Reniscymnus cordatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7E8D0702428331C0FBF39F4F3B73B6A" rank="species">sp. nov.</taxonomicNameLabel>
in colour pattern, but can be distinguished from the latter by the relatively shortened inner arm of penis, flattened apex of penis, and the unique tegmen with penis guide recurved inwardly at apex in lateral view and constricted at base in inner view.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJEAG" type="description">
<paragraph id="AE1002E3F9EF31BCFBBFF25C6F540101">
<heading id="1D81163C9668A2C0C800FE38A0F20F41" reason="title">Description.</heading>
</paragraph>
<paragraph id="395ADE46D6160626B5CA9494546AB642">
<abbrev id="ABBRID0EPEAG" xlink_title="total length">TL</abbrev>
:
<quantity id="2F90523639F73D0144BBCCBA084FC486" metricMagnitude="-3" metricUnit="m" metricValue="1.58" metricValueMax="1.65" metricValueMin="1.51" unit="mm" value="1.58" valueMax="1.65" valueMin="1.51">1.511.65 mm</quantity>
,
<abbrev id="ABBRID0ETEAG" xlink_title="total width">TW</abbrev>
:
<quantity id="0A5268758685D248F9B8ACEAEACC8E29" metricMagnitude="-3" metricUnit="m" metricValue="1.17" metricValueMax="1.21" metricValueMin="1.13" unit="mm" value="1.17" valueMax="1.21" valueMin="1.13">1.131.21 mm</quantity>
,
<abbrev id="ABBRID0EXEAG" xlink_title="total height">TH</abbrev>
:
<quantity id="5EF08B4D9827270F42983DD0F62C8189" metricMagnitude="-4" metricUnit="m" metricValue="7.649999999999999" metricValueMax="8.299999999999999" metricValueMin="7.0" unit="mm" value="0.7649999999999999" valueMax="0.83" valueMin="0.7">0.700.83 mm</quantity>
,
<abbrev id="ABBRID0E2EAG" xlink_title="total length">TL</abbrev>
/
<abbrev id="ABBRID0E6EAG" xlink_title="total width">TW</abbrev>
: 1.331.36,
<abbrev id="ABBRID0EDFAG" xlink_title="elytral length">EL</abbrev>
/
<abbrev id="ABBRID0EHFAG" xlink_title="elytral width">EW</abbrev>
: 1.051.07,
<abbrev id="ABBRID0ELFAG" xlink_title="pronotal length">
<collectionCode id="C9597BBBAD3080D64E371AF1B564BC37" country="Czech Republic" name="Západoceské muzeum v Plzni" type="Museum">PL</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0EPFAG" xlink_title="pronotal width">
<collectionCode id="9125DAFC985941558DC3E7B98FA05028" country="Thailand" name="Paleontological Collections">PW</collectionCode>
</abbrev>
: 0.500.51,
<abbrev id="ABBRID0ETFAG" xlink_title="head width">HW</abbrev>
/
<abbrev id="ABBRID0EXFAG" xlink_title="pronotal width">
<collectionCode id="9E731F216AEE6EC557EDC42470923B92" country="Thailand" name="Paleontological Collections">PW</collectionCode>
</abbrev>
: 0.660.68,
<abbrev id="ABBRID0E2FAG" xlink_title="pronotal width">
<collectionCode id="E65BED6C2A81AFF19DA6210DBB8043C3" country="Thailand" name="Paleontological Collections">PW</collectionCode>
</abbrev>
/
<abbrev id="ABBRID0E6FAG" xlink_title="elytral width">EW</abbrev>
: 0.660.69.
</paragraph>
<paragraph id="DFB7F4CA153F69B27F6D4E755A4C9F36">
Body rounded oval, dorsum with white pubescence. Area of frons near clypeus with suberect and downward setae, distinctly longer and denser than those on top (Fig.
<figureCitation id="E5225FE12A5DACBF736CF4CC94D50EA8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 B</figureCitation>
). Body yellow, except tip of mandibles dark brown, metaventrite deep yellow in middle and elytral epipleurae brown (Fig.
<figureCitation id="10546E22B9B7627A5109DE84344996F7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 A C</figureCitation>
). Eyes finely faceted, large, interocular distance about 0.330.34 times head width. Head and pronotal punctures fine and dense. Surface of elytra with coarse, shallowly impressed punctures.
</paragraph>
<paragraph id="A8E57B81EC8A6D13CA515C0DB46709A1">
Abdominal postcoxal lines incomplete and roundly recurved, reaching 5 / 6 length of ventrite 1, area enclosed by the lines sparsely punctate, widely smooth along the lines; ventrite 1 with coarse and sparse punctures in middle, irregularly distributed (Fig.
<figureCitation id="C74CC0EE3DD849AADF3F67F4EFD2C36A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 D</figureCitation>
).
</paragraph>
<paragraph id="15D13C10D7656323432673141D8F9699">
Male genitalia. Penis stout and relatively short, shorter than abdomen; penis capsule with short inner arm, slightly longer than outer arm (Fig.
<figureCitation id="05C221D40FD24241018D98D188817DDC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 E</figureCitation>
); apex of penis wide, flattened with few membranous appendages (Fig.
<figureCitation id="FC02DB1B762A0F7B8CEEAE82D40AA153" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 F</figureCitation>
); tegminal strut stout with finger-like protrusion at apex (Fig.
<figureCitation id="A8EAB26E25DCF6EF1A4F8A76B4F884E6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 G</figureCitation>
); phallobase rounded in lateral view; tegmen with penis guide widest at base, gradually converging apically, slightly recurved at apex in lateral view, 3 times as long as parameres (Fig.
<figureCitation id="7C4A6A61CE4661A442A28276E2095A6F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 G</figureCitation>
); in inner view, penis guide slightly constricted at base, widest in basal 1 / 3, then gradually tapering apically, apex blunt (Fig.
<figureCitation id="1A7DD3BFF1381EB72DE5E1A4D82CCEAC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 H</figureCitation>
); parameres short with several long setae at apex (Fig.
<figureCitation id="65912DD8ECF6647C150F7241D65B1213" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENHAG" type="type material">
<paragraph id="2310EA3FB514C47E897E4DC886844158">
<heading id="2B70A01B8F53650B4C07F7DBBB08A0D5" reason="title">Type material.</heading>
</paragraph>
<paragraph id="F39F9EF900C1777D08B6CACE71EA6B90">
<materialsCitation id="851A008A3EFE15608EFBADC9BA2574FD" collectingDate="2007-05-24" collectionCode="SCAU" collectorName="X. M. Wang &amp; No." country="Laos" elevation="1280" location="Pakxong" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="70121B70B824D04D7A07FC8EE3573B9C" bold="true" italics="true">
<typeStatus id="E21D152FBF5D4BB7110AA144895B8AA9" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="36CE23EF0CFE651E444F230C3FFEF3FF" name="Laos">Laos</collectingCountry>
<specimenCount id="8DBE45046434D69CB310F57F40B9DABF" count="1" type="male"></specimenCount>
;
<location id="DBA2690482D4C3F67ADB42AB62B4604E" LSID="urn:lsid:plazi:treatment:8EE0B488CC195C47AEFE070527DD1C64:DBA2690482D4C3F67ADB42AB62B4604E" country="Laos" name="Pakxong">Pakxong</location>
; alt.
<quantity id="BDAC5D9AAB21A08910735AC309C48EED" metricMagnitude="3" metricUnit="m" metricValue="1.28" unit="m" value="1280.0">
<elevation id="4EC0D203CB109E284B66EAB152DF2BD6" metricMagnitude="3" metricUnit="m" metricValue="1.28" unit="m" value="1280.0">1280 m</elevation>
</quantity>
,
<date id="7DD6A91ECAA9D9E1A16EC37FEF9A9978" value="2007-05-24">
<collectingDate id="ABAAA914774D35D713D4ADC1E97360B2" value="2007-05-24">24 May 2007</collectingDate>
</date>
;
<collectorName id="4D5371F1ABBA2E59B94E62BA29BD1A28">X. M. Wang</collectorName>
et al. leg.
<collectorName id="DECD7C8EEEDA206BA7221794A45BF33D">No.</collectorName>
<named-content id="FF016D07C06283A1861C8BAC71D1ACC4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Department of Entomology, South China Agricultural University">
<collectionCode id="FAD42AB39E6F518345D130B39A36F73F">SCAU</collectionCode>
</named-content>
(E) 17561;
<named-content id="FE5C91840DBD98BCBC7F3B364E20FE6D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Department of Entomology, South China Agricultural University">
<collectionCode id="587D038D771FBCB91FD22E2AC3986D29">SCAU</collectionCode>
</named-content>
</materialsCitation>
.
<materialsCitation id="2C5C7BC707A5BC91333EF3EF008AC4FE" collectingDate="2007-05-24" collectionCode="SCAU" collectorName="X. M. Wang &amp; No." country="Laos" elevation="1280" location="Pakxong" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="240DB5F8A229E9A939BCAE59FED0A95A" bold="true" italics="true">
<typeStatus id="68A656D9A49F6FCD1D686512ACEE89CE" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCount id="B99F93A09FCCE82E22D947424007FF95" count="1" type="female">1 ♀</specimenCount>
; same data as for holotype
</materialsCitation>
<materialsCitation id="793CA671F9F4F1ED1F59F2FE789290F4" collectingDate="2007-05-25" collectorName="X. M. Wang" country="Laos" elevation="380" location="Heu Gnommolat" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="0DC1ED3A4530ECE15208328F7164B985" count="1" type="male">1 ♂</specimenCount>
;
<location id="C9DCDF6029D99C78575972E582D99EFE" LSID="urn:lsid:plazi:treatment:8EE0B488CC195C47AEFE070527DD1C64:C9DCDF6029D99C78575972E582D99EFE" country="Laos" name="Heu Gnommolat">Heu Gnommolat</location>
; alt.
<quantity id="7C2681479FEF77C06AB63A41E7841C9E" metricMagnitude="2" metricUnit="m" metricValue="3.8" unit="m" value="380.0">
<elevation id="0CA922A4BE977D111F06D22E7DAD690E" metricMagnitude="2" metricUnit="m" metricValue="3.8" unit="m" value="380.0">380 m</elevation>
</quantity>
;
<date id="7453153957C30F49190CA816ECC0727D" value="2007-05-25">
<collectingDate id="68DAABC8864F071A46AD82B46C46A46D" value="2007-05-25">25 May 2007</collectingDate>
</date>
;
<collectorName id="8B3768D6FCDDA78B09F0D2CA50E2E683">X. M. Wang</collectorName>
et al. leg
</materialsCitation>
. •
<materialsCitation id="AA42405D5C0944C3BC8A850C475F9D52" collectingDate="2007-06-02" collectingDateMax="2007-06-04" collectingDateMin="2007-06-02" collectionCode="NBCA" collectorName="X. M. Wang" country="Laos" elevation="625" location="Phou Khao Kbouay" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="813F85403005D8082AB3D8CD7C3EFEF3" count="1" type="female">1 ♀</specimenCount>
;
<location id="AA29A090C57F3D0667691C2203EAF6B0" LSID="urn:lsid:plazi:treatment:8EE0B488CC195C47AEFE070527DD1C64:AA29A090C57F3D0667691C2203EAF6B0" country="Laos" name="Phou Khao Kbouay">Phou Khao Kbouay</location>
<collectionCode id="EC2841CA8ADD4D39F00929D3DC52857E">NBCA</collectionCode>
; alt.
<quantity id="CC18E4545B3CC686501930FC35226EF7" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" unit="m" value="625.0" valueMax="750.0" valueMin="500.0">
<elevation id="29511A5E9B915FA01B09CF94B716DEAF" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" unit="m" value="625.0" valueMax="750.0" valueMin="500.0">500750 m</elevation>
</quantity>
;
<date id="1E529B1C7C3CB6B5362AEF8425435384" value="2007-06-02" valueMax="2007-06-04" valueMin="2007-06-02">
<collectingDate id="C581DC3C0974CB537E38CD2520A2C6CC" value="2007-06-02" valueMax="2007-06-04" valueMin="2007-06-02">24 Jun. 2007</collectingDate>
</date>
;
<collectorName id="6EADD9FCBAD7EEEBC1522F12F2E5C24B">X. M. Wang</collectorName>
et al. leg
</materialsCitation>
. •
<materialsCitation id="54897927796975FD3402B548005EB807" collectingDate="2005-12-21" collectorName="C. Telakang" country="Laos" location="Tnangend" specimenCount="14" specimenCount-female="9" specimenCount-male="5" stateProvince="Khammouan" typeStatus="paratype">
<specimenCount id="318C4F2B522BCFCB36B1F28C47E7C4AB" count="5" type="male">5 ♂</specimenCount>
<specimenCount id="7F359BE4D801762CB3983F531722478F" count="9" type="female">9 ♀</specimenCount>
;
<collectingRegion id="17D9B9A407A8BC2F77BB3E77C25C6AB6" country="Laos" name="Khammouan">Khammouane</collectingRegion>
<location id="1B8A77E48C03CA55DC33D370F7C45AC8" LSID="urn:lsid:plazi:treatment:8EE0B488CC195C47AEFE070527DD1C64:1B8A77E48C03CA55DC33D370F7C45AC8" country="Laos" name="Tnangend" stateProvince="Khammouan">Tnangend</location>
;
<date id="8BCD8F0DC92B7BB4E4072B792BA3EFCE" value="2005-12-21">
<collectingDate id="3DD8E4FD291A5084A710AAF08D067418" value="2005-12-21">21 Dec. 2005</collectingDate>
</date>
;
<collectorName id="CB0FD2CA3EC842601DB9F2D684FBFFDD">C. Telakang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="59A4F7658EDE60A154DE883E2380F681" collectingDate="2005-12-16" collectorName="C. Telakang" country="Laos" location="Lakxao" specimenCount="1" specimenCount-male="1" stateProvince="Khammouan" typeStatus="paratype">
<specimenCount id="3BC1697FB83D6F0B33458E2C18BAB908" count="1" type="male">1 ♂</specimenCount>
;
<collectingRegion id="636606E14509A08180E3952CF882C2D9" country="Laos" name="Khammouan">Khammouane</collectingRegion>
<location id="CEAA4F22028E92E0A6EE80D1108A5B6F" LSID="urn:lsid:plazi:treatment:8EE0B488CC195C47AEFE070527DD1C64:CEAA4F22028E92E0A6EE80D1108A5B6F" country="Laos" name="Lakxao" stateProvince="Khammouan">Lakxao</location>
;
<date id="E031C627D46B9A29EFA8D1E429A153BC" value="2005-12-16">
<collectingDate id="14F79DB8A5665748A415A0B77F929BED" value="2005-12-16">16 Dec. 2005</collectingDate>
</date>
;
<collectorName id="F17AE937928AE7D57B0E0B77E18E94FF">C. Telakang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="8DE3B4EA5479EC0C81B5395D1580D15C" collectingDate="2005-12-25" collectorName="C. Telakang" country="Laos" location="Thakhex" specimenCount="1" specimenCount-female="1" stateProvince="Khammouan" typeStatus="paratype">
<specimenCount id="631C91994611A1A0A4B1C39878070AB3" count="1" type="female">1 ♀</specimenCount>
;
<collectingRegion id="A2A6BD1BD8F7ED519C87066F5237277B" country="Laos" name="Khammouan">Khammouane</collectingRegion>
<location id="F65B6D33D163BC8B5EB6ACDED7A2151E" LSID="urn:lsid:plazi:treatment:8EE0B488CC195C47AEFE070527DD1C64:F65B6D33D163BC8B5EB6ACDED7A2151E" country="Laos" name="Thakhex" stateProvince="Khammouan">Thakhex</location>
;
<date id="0AF5E6C6BC4016E6DB144B1750304888" value="2005-12-25">
<collectingDate id="CC258F097187924AB858E1D951742146" value="2005-12-25">25 Dec. 2005</collectingDate>
</date>
;
<collectorName id="62AD4AE150967B4B270CAAEC6383902E">C. Telakang</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFIAG" type="distribution">
<paragraph id="1441E30F60D96A4E64336AA09D00138D">
<heading id="2118BFDB2F6E4D0B591D479E4A9E1C58" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="C283316229A25AD7147D12D03D5F62E3">
<collectingCountry id="7EF8814D367313120D96C3E0C4B031F4" name="Laos">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKIAG" type="etymology">
<paragraph id="0324385223D8159BD64BEE9AB034AE8C">
<heading id="4E6A1504FA60B24E3D0F9FB380D1F846" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="637703B25968ACF301139D1D9DC21C77">
The species epithet is derived from the Latin adjective “
<emphasis id="9A7B1FB4A54F780908BC9B36A93B2688" italics="true">explanatus</emphasis>
”, referring to the flattened apex of the penis.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,461 @@
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<mods:title id="342CCDB3494FC4B8840816E12CD52F5A">Reniscymnus gen. nov., a new genus of Scymnini (Coleoptera, Coccinellidae) from the Oriental region</mods:title>
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<mods:namePart id="5C513F4EE83B5FBF015C6EAD935F39B0">Peng, Feng</mods:namePart>
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<mods:affiliation id="25E603DDC6A1DC37B1B83F5632D637B0">Department of Forest Protection, College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510640, China &amp; Engineering Research Center of Biological Control, Ministry of Education, Guangzhou 510642, China</mods:affiliation>
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<mods:namePart id="FFF7B084AFB84BD479D3C3D9D3DED3A1">Tang, Mingjie</mods:namePart>
<mods:affiliation id="0AA04FC7AC2E2BD2C4395910BDCDEFA0">Department of Forest Protection, College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510640, China &amp; Engineering Research Center of Biological Control, Ministry of Education, Guangzhou 510642, China</mods:affiliation>
</mods:name>
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<mods:affiliation id="93CA3F3A1424944F4137C1E02A0E4D7F">Engineering Research Center of Biological Control, Ministry of Education, Guangzhou 510642, China &amp; Department of Forest Protection, College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510640, China</mods:affiliation>
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<subSubSection id="CBA589B86B6AD063D8DBC8E5B8BA0969" type="nomenclature">
<paragraph id="3579FC645DFA6117B09F3432977F8EC2">
<taxonomicName id="99FA634DE7ADC2E5F06CDAEB7BC19C15" authority="Peng &amp; Chen" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="9F2B890C9E98DDCEF78B7D3844BB1160" italics="true">Reniscymnus</emphasis>
Peng &amp; Chen
</taxonomicName>
<taxonomicNameLabel id="AB8B5076AFA95B3DFF8096B01ED84B59" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIHAE" type="type species">
<paragraph id="463ACC057561A52E14CDF42B83C64CDA">
<heading id="B6315128A3CE5BC86F18A7562D8AF8A8" reason="title">
<typeStatus id="4FCEABBB18F59A3BFC5F06BF924660A9">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="F2EB73716990FB4BDEA1594D9B3820EB">
<taxonomicName id="92B4EBE300E63D3F84F4B033BA3B9AEA" authority="Peng &amp; Chen" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cordatus" status="sp. nov.">
<emphasis id="DF22E5CA405134EDB3716AFD05E3E59B" italics="true">Reniscymnus cordatus</emphasis>
Peng &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="217F7029454F4C2338632D686FD2A5AE" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZHAE" type="etymology">
<paragraph id="944C2B3079DA605502259EBFB20941E5">
<heading id="44996F4C6FE8DE9DC9F8773FE51269D4" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="DBB76DD927D5FC77339F0D6EE3C17341">
The generic name is derived from the generic name
<emphasis id="39B82F5FA98C32846E6C15FA6EF9EC7E" italics="true">Scymnus</emphasis>
and the memory of the late Prof. Ren Shunxiang from South
<collectingCountry id="2A27C2498C2B28B9255A709AA5E9036E" name="China">China</collectingCountry>
Agricultural University, a well-known Chinese entomologist who devoted most of his life to the study of Coccinellidae and biological control. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELIAE" type="diagnosis">
<paragraph id="165336DF1622001FCD17C8D3BF72C519">
<heading id="9D69C44A2411E4EFE74A66FF7378DAD6" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="91190849F4DA923FF83569FEE0D7B55C">
<taxonomicName id="2A79A9B72A0C77D3912563587B10B385" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1C802E1AC23755B6C57269D1FFE5FFD6" italics="true">Reniscymnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0867CC83D23D36861C69E592917FC9ED" rank="genus">gen. nov.</taxonomicNameLabel>
presents some characters found in other genera of
<taxonomicName id="B9D3D27E6A8924A62F343EA0C359C24F" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="tribe" tribe="Scymnini">Scymnini</taxonomicName>
(especially
<taxonomicName id="925B5BD8109E9422FA9C7C1E95A00C5E" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="5701BB6E6535627F6F5F8451C0AE1FB7" italics="true">Sasajiscymnus</emphasis>
</taxonomicName>
,
<taxonomicName id="DC450BD5552E0BF198715214201F44D2" class="Insecta" family="Coccinellidae" genus="Slipinskiscymnus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="704C68AB9C3E0B73916C5A1FEFFEE499" italics="true">Slipinskiscymnus</emphasis>
</taxonomicName>
and
<taxonomicName id="2B856808C406321F6742C1A1A4431C7B" class="Insecta" family="Coccinellidae" genus="Axinoscymnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="E9EEAA8B4CBEB5843D24F51609F7FDC9" italics="true">Axinoscymnus</emphasis>
</taxonomicName>
). But the new genus can be easily distinguished from the latter by the following characters: frons narrow, less than the width of an eye (Fig.
<figureCitation id="F62F0923831625F3642FB72EC1F17EE0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 A</figureCitation>
) (vs. wide, more than the width of an eye in
<taxonomicName id="056205EF3290E87349731255DF8A5F51" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="AFBD0175480BC1BFE53B5A092153D1D5" italics="true">Sasajiscymnus</emphasis>
</taxonomicName>
and
<taxonomicName id="B7F90EA0AEC67A1D578EDBD14045D43A" class="Insecta" family="Coccinellidae" genus="Slipinskiscymnus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="ACB07B242A20186C1615604990D90D41" italics="true">Slipinskiscymnus</emphasis>
</taxonomicName>
); antenna composed of 8 antennomeres with a distinctly swollen scape and pedicel, antennomeres 58 forming a fusiform and inflated club and the terminal antennomere with two extremely long setae (Figs
<figureCitation id="E41A8EB3D1F52E03C1872DFFCDB5FA3F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 C</figureCitation>
,
<figureCitation id="B791CF5E37516FE5BEC6A3F21801D5BD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 I</figureCitation>
) (vs. 9 antennomeres in
<taxonomicName id="AABC25480AD6D5E02E7DB27834455AF9" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="A0E73212A2764BFFB204C417B5A240BA" italics="true">Sasajiscymnus</emphasis>
</taxonomicName>
; 10 antennomeres in
<taxonomicName id="4D3C48B36636DFE810A3AC3E48C576CA" class="Insecta" family="Coccinellidae" genus="Slipinskiscymnus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="ACE1A51CD84D48E154D31762A5A8B83B" italics="true">Slipinskiscymnus</emphasis>
</taxonomicName>
, and 11 antennomeres in
<taxonomicName id="81BC119F8DE76408050AA707FF8C9DCF" class="Insecta" family="Coccinellidae" genus="Axinoscymnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="1A2DB06F2C2FF83CE61F46B0A320E4D3" italics="true">Axinoscymnus</emphasis>
</taxonomicName>
); terminal maxillary palpomere slightly narrows apically (Figs
<figureCitation id="378D29A4E72A6E4E10997C4F5FF0640D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 D</figureCitation>
,
<figureCitation id="5CC35A38309F94E60AB84ED0FD135909" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 J</figureCitation>
) (vs. expanded apically in
<taxonomicName id="9E568189CF30C0E92C08E4AA943C489C" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="60C1FD7B18CB622BB595121CBE463651" italics="true">Sasajiscymnus</emphasis>
</taxonomicName>
;
<taxonomicName id="EA92CE13B906457F0DA9FC13212E7ECB" class="Insecta" family="Coccinellidae" genus="Slipinskiscymnus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="0B844E010ABC3586E30BC65C4324FAFD" italics="true">Slipinskiscymnus</emphasis>
</taxonomicName>
and
<taxonomicName id="BFA10D70AECEF2F3FE96AC2AB6A198D1" class="Insecta" family="Coccinellidae" genus="Axinoscymnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="0AB8A0A7D4906775E9FA2083D27B6BB6" italics="true">Axinoscymnus</emphasis>
</taxonomicName>
); prosternal process with carinae slightly convergent anteriorly (Fig.
<figureCitation id="C48F553C8635D5EC5D6BEB8D317A9C8C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 A</figureCitation>
) (vs. complete carinae parallel, joined anteriorly forming square in
<taxonomicName id="E02509B5B235FC5EB3121183C7FD7515" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="57D3EAC91211C8A83473EB09D32A7DB0" italics="true">Sasajiscymnus</emphasis>
</taxonomicName>
; narrow carinae in
<taxonomicName id="E28D256AB493A913542EBA08FDCBC9D5" class="Insecta" family="Coccinellidae" genus="Axinoscymnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="A0830BB32363D1C56F6C5D4886159D09" italics="true">Axinoscymnus</emphasis>
</taxonomicName>
); abdominal postcoxal lines incomplete, strongly recurved toward the base of ventrite 1 but not reaching it, the area enclosed by abdominal postcoxal lines distinctly wide (Figs
<figureCitation id="B4EE10CF6EC92B9A5023E9CAE2B1674C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 D</figureCitation>
,
<figureCitation id="B0D620BD5FEEACD5F094392B0B954A16" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 D</figureCitation>
) (vs. incomplete, generally parallel to the posterior margin of ventrite
<quantity id="DF27A1930557D429195452B6A1996097" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="D831F61F30ADBDF4595FE76BAA3D339B" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="9F37F15310FA5824717D54D557EE4056" italics="true">Sasajiscymnus</emphasis>
</taxonomicName>
; complete in
<taxonomicName id="ED9426E2AB6D91A7EDB8D653C180638D" class="Insecta" family="Coccinellidae" genus="Slipinskiscymnus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="FC7290C1B8F5751D5A72CCD0D18D3820" italics="true">Slipinskiscymnus</emphasis>
</taxonomicName>
and
<taxonomicName id="94CB350B23C03718C4D32BB5534A8393" class="Insecta" family="Coccinellidae" genus="Axinoscymnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="A44E05754C1C4F9F31FA6E27FB73B91C" italics="true">Axinoscymnus</emphasis>
</taxonomicName>
); penis guide subconical with a pointed apex (Fig.
<figureCitation id="8F2448A7829408639480493FD0C13FD5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 I J</figureCitation>
); parameres rather short with few long setae (Fig.
<figureCitation id="35DA102C4EBD7FD8171D11B924C1AE8F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 I J</figureCitation>
).
</paragraph>
<caption id="BE55F230FD6FCDCBE28835C7C04E05BE" ID-DOI="10.3897/zookeys.1226.130352.figure1" ID-arpha="B1DEC34B-DE60-55F4-83FA-31201C93A5BE" httpUri="https://binary.pensoft.net/fig/1243495" startId="F1">
<paragraph id="F106901574663299D823288F6B4C83AC">
<label id="80EB67D0A637DA1393398CB4B5CD263F">Figure 1.</label>
</paragraph>
<paragraph id="9A23ED9C3CCB18EB0678430B31284521">
Main characters of the genus
<emphasis id="7AB06F5EEE663CB1106690D973212634" italics="true">Reniscymnus</emphasis>
<taxonomicNameLabel id="85CCE1B60FF6BB15A8194D4D57994BD3" rank="genus">gen. nov.</taxonomicNameLabel>
<emphasis id="D30F47D8BEF9B09CB842C2B004586551" bold="true">A</emphasis>
<emphasis id="60911DF11849AB4063B3201F69214DD4" bold="true">I</emphasis>
<taxonomicName id="C99EB0B14F1E0DDE167FA0E67032396A" authority="Peng &amp; Chen" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cordatus" status="sp. nov.">
<emphasis id="44A700C3352FFE64F4BB17E554578785" italics="true">Reniscymnus cordatus</emphasis>
Peng &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="59CD2309456488A8BFD7698A5CE5D409" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="0B507B1DC5398F32560173033AD27C52" bold="true">A</emphasis>
ventral view
<emphasis id="E803FD7A58B6F5C2844974EE87E63850" bold="true">B</emphasis>
head
<emphasis id="39291DCF697168B42B095B547DFA55ED" bold="true">C</emphasis>
antenna
<emphasis id="E8FA4916458B8FBB003BFC6C3CE7AAEC" bold="true">D</emphasis>
maxilla
<emphasis id="5EB482E298093CFD62AF547CAB93C0B3" bold="true">E</emphasis>
mandible
<emphasis id="E4D319255AA9E8517C239291ABA5DDC0" bold="true">F</emphasis>
labium
<emphasis id="CD2986EA91C0E13928B6304E9A1E0743" bold="true">G</emphasis>
hind leg
<emphasis id="537AE4035DC8BAC82AE3C488DE33F90A" bold="true">H</emphasis>
spermatheca
<emphasis id="7E8DC009B81933203504F4DC824CC60A" bold="true">I</emphasis>
coxites. Scale bars: 0.5 mm (
<emphasis id="E7E75863789FE215AAE038F1474B898D" bold="true">A</emphasis>
); 0.1 mm (
<emphasis id="FF01D5D7FBDBF499A0E91A365B543708" bold="true">B</emphasis>
); 0.05 mm (
<emphasis id="159B797A7375D14C87AAF20CC1C56CF7" bold="true">C I</emphasis>
).
</paragraph>
</caption>
<caption id="FA398C25A917A8556D8D7D8C863C45DC" ID-DOI="10.3897/zookeys.1226.130352.figure2" ID-arpha="2F1832BC-CB26-5152-AC6F-2A8D01942177" httpUri="https://binary.pensoft.net/fig/1243496" startId="F2">
<paragraph id="6B4FBAD1A2E510A2FA884028FE3D9814">
<label id="3A3DEB719F2B727E3573BCC72F3EC8BC">Figure 2.</label>
</paragraph>
<paragraph id="D7C00018F6FFA90C36E25D9537ACFEE6">
<taxonomicName id="94A92D54B100E5713AD238A9149D88DC" authority="Peng &amp; Chen" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cordatus" status="sp. nov.">
<emphasis id="6F2294077E2E272E2F42DE21EF4E4EC3" italics="true">Reniscymnus cordatus</emphasis>
Peng &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="83845ECD0571D1D4ED71D45DE3948B54" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B7B341E6E50E9C1AC30C16A23A41F9AA" bold="true">A F, I, J</emphasis>
holotype
<emphasis id="6A441ABA908E8CA79363010B446D5702" bold="true">G, H</emphasis>
paratype)
<emphasis id="2624018EE1014E70A50DEB40ECEFC420" bold="true">A</emphasis>
dorsal view
<emphasis id="4A085E09BE54AA3B3155ACBC4E1C4A1A" bold="true">B</emphasis>
frontal view
<emphasis id="2525B2F6F1D3D47CDC3F174A94B0CB8F" bold="true">C</emphasis>
lateral view
<emphasis id="6205DEA6F344DDB56D41E1E05014D60D" bold="true">D</emphasis>
abdomen
<emphasis id="9C7494E8A8DE299EA155E54029D2248B" bold="true">E, G</emphasis>
penis
<emphasis id="1183BFBE0453D2BBAF9131E053787B25" bold="true">F, H</emphasis>
apex of penis
<emphasis id="A9709A58CDDC91B6CAEBC2DD82CAE713" bold="true">I</emphasis>
tegmen, lateral view
<emphasis id="BF0AB65E8AD79685EBA886DC0835F103" bold="true">J</emphasis>
tegmen, inner view. Scale bars: 0.5 mm (
<emphasis id="DA385FCFDDF74A01109D95D5CFC95271" bold="true">A C</emphasis>
); 0.1 mm (
<emphasis id="81CEBCAC59B4EFEF26D7C5F6C43F7179" bold="true">D J</emphasis>
).
</paragraph>
</caption>
<caption id="F5ED1D98724EB155C245CA4600FD8B6D" ID-DOI="10.3897/zookeys.1226.130352.figure3" ID-arpha="0169DD6D-7E82-54B4-9305-3C02BF54FA15" httpUri="https://binary.pensoft.net/fig/1243497" startId="F3">
<paragraph id="61802B6F289A0BB19872958915D926AE">
<label id="7F51F1D510AA1ECE18373834F2B6B0EC">Figure 3.</label>
</paragraph>
<paragraph id="F7C6F2B01FFAC3C1E9DA620E92C39EA3">
<taxonomicName id="590224E226A26BC8D4893EE2E0DE49CC" authority="Peng &amp; Chen" authorityName="Peng &amp; Tang &amp; Wang &amp; Chen" authorityYear="2025" class="Insecta" family="Coccinellidae" genus="Reniscymnus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="explanatus" status="sp. nov.">
<emphasis id="D4009D370979AFD61FF101D08798D042" italics="true">Reniscymnus explanatus</emphasis>
Peng &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="5E62EC1B86E09FFC4E83CBAD118A9EE4" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="0311BF04AA2EE44D6CEC3E768E10B047" bold="true">A H</emphasis>
holotype
<emphasis id="AB150B40BCC43181F0345E8647AEF91E" bold="true">I O</emphasis>
paratype)
<emphasis id="E2061A3A288304B94062C487E0DB1306" bold="true">A</emphasis>
dorsal view
<emphasis id="EE0FF6DCB51BC1C7993DC21815C4DE7E" bold="true">B</emphasis>
frontal view
<emphasis id="612EEA7E424CABEF1C4802F4C2286C77" bold="true">C</emphasis>
lateral view
<emphasis id="977BCC5F852A65B83B03173EE95538D4" bold="true">D</emphasis>
abdomen
<emphasis id="C4802F2B90D7147C6A970A31234E0D68" bold="true">E</emphasis>
penis
<emphasis id="056C341794D7D0BD26CCB10451ED39FE" bold="true">F</emphasis>
apex of penis
<emphasis id="9B7371F5E9EF495E27F5BFA454E7832F" bold="true">G</emphasis>
tegmen, lateral view
<emphasis id="2296713CCA0A6AA81C6AC22823E185FE" bold="true">H</emphasis>
tegmen, inner view
<emphasis id="BBCB4B66061EE216F644C81670C5743B" bold="true">I</emphasis>
antenna
<emphasis id="0D2F60C7BE8AE275EDA33423203EB6A7" bold="true">J</emphasis>
maxilla
<emphasis id="F96797F0566007FBA74D1F9B42B62C96" bold="true">K</emphasis>
mandible
<emphasis id="F24B68EDA843AC00170758F56800D6E4" bold="true">L</emphasis>
labium
<emphasis id="86A7A6222AF2095AACA186E771CC2191" bold="true">M</emphasis>
hind leg
<emphasis id="C59AAEAC87AA145347888E222C93BC72" bold="true">N</emphasis>
spermatheca
<emphasis id="2ACB44E78023079D03BA794A713E48C1" bold="true">O</emphasis>
coxites. Scale bars: 0.5 mm (
<emphasis id="F954A796E79482D9813F61ABC3197C1A" bold="true">A C</emphasis>
); 0.1 mm (
<emphasis id="B1DFC6EAF074311B749565F8F046FDAD" bold="true">D H</emphasis>
); 0.05 mm (
<emphasis id="B645E4A7361C186B54DE6E1C6C3A8638" bold="true">I O</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EATAE" type="description">
<paragraph id="6B93F4C93679B3FAF81D77B3504201DB">
<heading id="675CB918DDA664C322A44293A494646F" reason="title">Description.</heading>
</paragraph>
<paragraph id="4BD970DFD75B186E9CB252630BBE090B">
Body elongate oval, color lighter and yellowish, dorsum moderately convex, covered with uniform short setae (Fig.
<figureCitation id="36A068FF7ED7A39C27E642E115DB685C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 A C</figureCitation>
).
</paragraph>
<paragraph id="BD0392452DE8D4840AB82F31CD318973">
Head transverse; frons relatively narrow, less than the width of an eye (Fig.
<figureCitation id="E924CEC62B2BD2DD2EBC8C3BC8AC5ACD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 A</figureCitation>
). Eyes large, rounded without eye canthus, finely faceted with short interfacetal setae; inner ocular margin arcuate (Fig.
<figureCitation id="A388E78D139C191C991FD5C5551FFBA8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 B</figureCitation>
). Clypeus short, and expanded laterally, not covering antennal insertions, anterior margin slightly convex (Fig.
<figureCitation id="D512C3CDB7F236DCC46FC1A206CE3D5B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 B</figureCitation>
). Labrum exposed. Antennae short with 8 antennomeres (Figs
<figureCitation id="DBAC4095ECE01420EB1F4C162152B753" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 C</figureCitation>
,
<figureCitation id="351BC525EB53D5B800A03221FB6A010E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 I</figureCitation>
); scape stout, widened and curved apically with several long setae; pedicel longest, swollen and barrel-shaped, as wide as scape; scape and pedicel combined nearly as long as antennomeres 38; antennomere 3 distinctly narrowed at base, recurved and turgidly expanded apically; antennomere 4 recurved, expanded apically; antennomeres 58 forming fusiform and inflated club with many short setae, and terminal antennomere trochiform with two extremely long setae (Figs
<figureCitation id="DE7EE60D6A7C7A7B89F66AFF21022E97" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 C</figureCitation>
,
<figureCitation id="257885E2D4F694CC066AD35F931C1CA3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 I</figureCitation>
). Mandibles bifid apically, with well-developed molar tooth (Figs
<figureCitation id="6B4FCA5CCA8001ACB7091040601B6286" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 E</figureCitation>
,
<figureCitation id="9D48144FCFA6F0014EAC18DD8DC36F57" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 K</figureCitation>
). Maxillary palp 4 - segmented; palpomere 1 short, ring-like; palpomere 2 elongate, widened apically; palpomere 3 subtriagular and compact at inner margin; terminal palpomere slightly narrowing apically, apical margin slightly obliquely truncate (Figs
<figureCitation id="2FA8B159BE1B7831F3F9762598CB34E1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 D</figureCitation>
,
<figureCitation id="0A00F18289FD9B15D95AC0E6F99CDBF0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 J</figureCitation>
). Maxilla with cardo round and laterally not expanded. Mentum narrowed basally, parallel-sided in basal third, widened apically, anterior margin of mentum weakly arcuate (Figs
<figureCitation id="115884FE228AA2C5D9DB6CD2B699467B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 F</figureCitation>
,
<figureCitation id="39DB3EEA9FE7B8249F81E5C4D4650432" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 L</figureCitation>
). Labial palp 3 - segmented, terminal palpomere nearly as long as penultimate one (Figs
<figureCitation id="8514E56A2158F0EF661E8C8D04557645" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 F</figureCitation>
,
<figureCitation id="03EC5888B1FBC4019702B996BE7B4D1C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 L</figureCitation>
).
</paragraph>
<paragraph id="E7E867B75B25226F913D3C0DE460F429">
Pronotum transverse with anterior corners roundly produced. Scutellar shield triangular and distinctly tiny (Fig.
<figureCitation id="7E57765DEA1D71CF996A53ADA9E6B722" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 A</figureCitation>
). Elytra with humeral calli distinct, with apex sharply rounded. Prosternum with narrow procoxae anteriorly, prosternal process wide and subquadrate, carinae widely separated, slightly convergent anteriorly, reaching anterior prosternum margin (Fig.
<figureCitation id="5BB05639A6984DC4982507F7B313B0FA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 A</figureCitation>
). Mesoventrite transverse with a truncate anterior margin; Mesocoxal process wide, slightly wider than width of corresponding mesocoxal diameter. Meso-metaventral junction visible, forming straight line. Metaventrite weakly convex, metaventral postcoxal lines widely separate medially, roundly recurved and complete laterally (Fig.
<figureCitation id="B1995208243F501271BC5236880A1917" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 A</figureCitation>
). Discrimen visible. Elytral epipleuron incomplete, reaching posterior margin of abdominal ventrite 1, without foveae (Fig.
<figureCitation id="301B1F8772676BB9907D63C0C0DCD461" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 A</figureCitation>
).
</paragraph>
<paragraph id="DCA3D15B77547EA67DA60738B7EDB0E0">
Legs short, extending a little beyond lateral margins of elytra; trochanters wide, deviously and angulately produced externally; femora relatively slender; tibiae rather slender, about 1 / 2 as wide as femur, without apical spurs; tarsi with 3 tarsomeres, tarsomeres 1 and 2 lobed, tarsal claws with sharp basal tooth (Figs
<figureCitation id="7DDD60A850D28E79F28AC977046E9E51" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 G</figureCitation>
,
<figureCitation id="69048413DB9E2F1392FA5DA8151C3674" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 M</figureCitation>
).
</paragraph>
<paragraph id="6088E2979A331B235746633A18F04EBB">
Abdominal processes transverse, distinctly wide. Ventrite 1 distinctly longer than ventrite 2, in middle almost 2 times length of ventrite 2 (Figs
<figureCitation id="93433B415150295F1B8426DE0A18E238" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 D</figureCitation>
,
<figureCitation id="B241835D8FC4697062DBEE32939B2568" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 D</figureCitation>
). Abdomen with six ventrites in both sexes. Abdominal postcoxal lines incomplete, strongly recurved toward base of ventrite 1 but not reaching it, posteriorly extending more than half of length of ventrite 1 (Figs
<figureCitation id="E8B675F00BA617261B91245143977DB4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 D</figureCitation>
,
<figureCitation id="104C18717EF6984B38CC903DDF46C3BC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 D</figureCitation>
), area enclosed by abdominal postcoxal lines distinctly wide.
</paragraph>
<paragraph id="188C770E2D383F69844A197FC5B40DEE">
Male genitalia. Relatively long, penis slender (Fig.
<figureCitation id="B55DB915C03B50EEFD0777D7BC852F27" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 E</figureCitation>
); apex of penis modified with membranous appendages (Fig.
<figureCitation id="51E92D499B7599E76A1CAE60DA8CB86B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 F</figureCitation>
); tegminal strut stout with a finger-like protrusion at apex. Penis guide stout, subconical with a pointed apex in inner view, parameres rather short with few long setae (Fig.
<figureCitation id="F50DDDDFF60409F05BD7E3DE3805B774" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Reniscymnus cordatus Peng &amp; Chen, sp. nov. (A F, I, J holotype G, H paratype) A dorsal view B frontal view C lateral view D abdomen E, G penis F, H apex of penis I tegmen, lateral view J tegmen, inner view. Scale bars: 0.5 mm (A C); 0.1 mm (D J)." figureDoi="10.3897/zookeys.1226.130352.figure2" httpUri="https://binary.pensoft.net/fig/1243496">2 I J</figureCitation>
).
</paragraph>
<paragraph id="E9564FA6D091439A58EAB8F6B555DDA7">
Female genitalia. Coxites shortened without styli (Figs
<figureCitation id="0EEEB008429535EE15D5C68621FC3EE7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 I</figureCitation>
,
<figureCitation id="B05D2792B6C59795330C3E8171EDEBF1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 O</figureCitation>
); spermatheca short and recurved with shortened cornu and swollen ramus and nodulus (Figs
<figureCitation id="59ADAB4C0ACE53FB7FF51CB09E4DE7F5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Main characters of the genus Reniscymnus gen. nov. A I Reniscymnus cordatus Peng &amp; Chen, sp. nov. A ventral view B head C antenna D maxilla E mandible F labium G hind leg H spermatheca I coxites. Scale bars: 0.5 mm (A); 0.1 mm (B); 0.05 mm (C I)." figureDoi="10.3897/zookeys.1226.130352.figure1" httpUri="https://binary.pensoft.net/fig/1243495">1 H</figureCitation>
,
<figureCitation id="726FF0FB69E7510755DA77E63ECF0577" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reniscymnus explanatus Peng &amp; Chen, sp. nov. (A H holotype I O paratype) A dorsal view B frontal view C lateral view D abdomen E penis F apex of penis G tegmen, lateral view H tegmen, inner view I antenna J maxilla K mandible L labium M hind leg N spermatheca O coxites. Scale bars: 0.5 mm (A C); 0.1 mm (D H); 0.05 mm (I O)." figureDoi="10.3897/zookeys.1226.130352.figure3" httpUri="https://binary.pensoft.net/fig/1243497">3 N</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWXAE" type="distribution">
<paragraph id="3569C8490B4E8E15F7DC88E31F35DCB0">
<heading id="9EBEE5901A9530A3D463D64D67984E0D" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="EB5ABE7057A52EE69C61E3244280A420">
<collectingCountry id="DF795F28D4C65D4DF3BB0C10B0508926" name="China">China</collectingCountry>
(
<collectingRegion id="3DCC3C3DE232A4348D0F37078007415C" country="China" name="Hainan">Hainan</collectingRegion>
,
<collectingRegion id="1D23BA2E50C64F58AC03411729EE7C00" country="China" name="Guangdong">Guangdong</collectingRegion>
,
<collectingRegion id="F062D75739325612A4D2C5F320EC266E" country="China" name="Guangxi">Guangxi</collectingRegion>
,
<collectingRegion id="E208DEAC821316015381D9D7B1C875E7" country="China" name="Yunnan">Yunnan</collectingRegion>
);
<collectingCountry id="15C882CA4B59D2CBDB9258C937508868" name="Laos">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2XAE" type="remarks">
<paragraph id="ED487F8B4AE60B02908A472C7F86DD99">
<heading id="89F32608495CCFA8CDDAD177F1A2341F" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="355BA29D05CB603308CA4D3A526CD045">
The species described in this study are distributed in the Oriental region. Interestingly, both new species of this genus exhibit a lighter body coloration similar to that of
<taxonomicName id="E1BE14FDD81DA79E22498E084EE6B0B3" authority="Ren &amp; Pang, 1992" authorityName="Ren &amp; Pang" authorityYear="1992" class="Insecta" family="Coccinellidae" genus="Axinoscymnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cardilobus">
<emphasis id="D2C83AF94423F088A0728D141A58E3D7" italics="true">Axinoscymnus cardilobus</emphasis>
Ren &amp; Pang, 1992
</taxonomicName>
, which is in contrast to the darker or black body coloration among members of the tribe
<taxonomicName id="13F6A74E643AF56965A9B930A1C8CD1A" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="tribe" tribe="Scymnini">Scymnini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="502A2BD848AD7D6B7690BB437F2C6F58">
<taxonomicName id="19CC2C1E3C17B0D6D36760D0B622DE30" authority="Laboissiere, 1936" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="6970904C0D05EDBA46BAD06896CB9D36" italics="true">Paleosepharia truncata</emphasis>
<bibRefCitation id="2A28BA5E234CA21E8091D022D32CC470" DOI="10.1080/21686351.1936.12278963" author="Laboissière V" journalOrPublisher="Annales de la Société Entomologique de France" pagination="239-262" refId="B19" refString="Laboissiere V (1936) Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et especes (5 e partie). Annales de la Societe Entomologique de France 105: 239-262. https://doi.org/10.1080/21686351.1936.12278963" title="Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et espèces (5 e partie)." volume="105" year="1936">Laboissière, 1936</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="173CDED45349CA3907C4DC9C6571888F" type="description">
<paragraph id="9541BB30CBF55B401C44E77F5A985018">
<figureCitation id="999EC99BA43314DA2F10416404B514ED" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Paleosepharia truncata Laboissière, 1936 (paratype). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1226.140239.figure11" httpUri="https://binary.pensoft.net/fig/1243042">Figs 11</figureCitation>
,
<figureCitation id="D05DAEEE6A75875985D5E7F1BF7EEF47" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Paleosepharia truncata Laboissière, 1936 (male) A C habitus D F aedeagus G depression of the elytra of the male. A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1 mm (A C); 0.5 mm (D F)." figureDoi="10.3897/zookeys.1226.140239.figure12" httpUri="https://binary.pensoft.net/fig/1243043">12</figureCitation>
,
<figureCitation id="CA28A43F5472B84533E0B03953B4C9E8" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Paleosepharia truncata Laboissière, 1936 (female) A-C habitus D vaginal palps E spermatheca F apex of abdomen of the female. A dorsal view B ventral view C lateral view. Scale bars: 1 mm (A C); 0.5 mm (D, E)." figureDoi="10.3897/zookeys.1226.140239.figure13" httpUri="https://binary.pensoft.net/fig/1243044">13</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="8BB2589F9AD3AFE0FCDF423F15506A91">
<treatmentCitationGroup id="8E2EFD2C58A78DF758B7AA64EEB8E3DE">
<treatmentCitation id="7D0DD23AEFD3D310D8F23DE920D27159" author="Laboissière V" page="251" year="1936">
<taxonomicName id="E396434EABFBC4B6C865DE539BCEAD1B" authority="Laboissiere 1936: 251" authorityName="Laboissiere" authorityPageNumber="251" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="9B239DA77911A90D41CCA680EE862D9B" italics="true">Paleosepharia truncata</emphasis>
<comment id="153F9921A98542BD26DFB3EDB3F55E76">
<bibRefCitation id="233991C271437498A722BD0677E6AA29" DOI="10.1080/21686351.1936.12278963" author="Laboissière V" firstAuthor="Laboissiere" journalOrPublisher="Annales de la Société Entomologique de France" page="251" pagination="239-262" refId="B19" refString="Laboissière V (1936) Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et espèces (5 e partie). Annales de la Société Entomologique de France 105: 239262. https://doi.org/10.1080/21686351.1936.12278963" title="Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et espèces (5 e partie)." volume="105" year="1936">Laboissière 1936: 251</bibRefCitation>
.
<abbrev id="ABBRID0E1BAG" xlink_title="type locality">TL</abbrev>
<collectingCountry id="FCC3BAD608BFF927C9F43CA1811F70B2" name="Vietnam">Vietnam</collectingCountry>
.
<abbrev id="ABBRID0E5BAG" xlink_title="type deposition">TD</abbrev>
<named-content id="5FEB5980062EEA4D880E08C277D9C7B4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle, Paris">
<collectionCode id="ABEE609903100BA1EC3C4C6E2AECFCBF" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
,
<named-content id="3E18D5350D6E2F754A1310BCA436F825" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-museum-hamburg" xlink_title="Zoological Museum Hamburg">
<collectionCode id="AC5FDF2C2217AA9C488DF3F1C280FC6E" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" type="Museum">ZMH</collectionCode>
</named-content>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJCAG" type="type material">
<paragraph id="070D315FDD03ADE150D61990383F3DE2">
<heading id="6360B70C297391072740C44F1BC591CD" reason="title">
<typeStatus id="D307C6790C4D9D1179E9F5D9783526DB">Type</typeStatus>
material.
</heading>
</paragraph>
<paragraph id="8C8918D8A81C28E43195DE8F81035365">
<emphasis id="54626460D085CB9EF70F08554DAF9D6E" bold="true" italics="true">Cotype</emphasis>
, •
<specimenCount id="7EDA1A4F0C4918FED8C315FD8DED6230" count="1" type="male"></specimenCount>
, Le Moult Vend. via Reinbek Eing. Nr. 1, 1957.
<taxonomicName id="915C71E539AE6407698B8099E2CDBD90" authority="V. Laboissiere Det." authorityName="V. Laboissiere Det." class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="B6E7ACA8C86DC387EDD69915B89D547F" italics="true">Paleosepharia truncata</emphasis>
V. Laboissière Det.
</taxonomicName>
MUSEUM PARIS COCHINCHINE RIERRE 1878.
<emphasis id="02B5AD1A81A1E3F8D2A3567B709E14B4" italics="true">
<emphasis id="F5466250F920B57F96756857105AFB51" bold="true" italics="true">
<typeStatus id="268BB2E41CEC8FAF2C2EDD5301261DEA" type="paralectotype">Paralectotypus</typeStatus>
</emphasis>
<taxonomicName id="F132ECEA9BAB723DB90BFA745A3BEC0E" authority="Laboissiere, 1936 Hazmi et al." authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="truncata">
Paleosepharia truncata
<bibRefCitation id="0D040939374313C3DD47230A39BB401D" DOI="10.1080/21686351.1936.12278963" author="Laboissière V" journalOrPublisher="Annales de la Société Entomologique de France" pagination="239-262" refId="B19" refString="Laboissiere V (1936) Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et especes (5 e partie). Annales de la Societe Entomologique de France 105: 239-262. https://doi.org/10.1080/21686351.1936.12278963" title="Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et espèces (5 e partie)." volume="105" year="1936">Laboissière, 1936</bibRefCitation>
Hazmi et al.
</taxonomicName>
</emphasis>
desig.
<named-content id="93A8E3D2DA61C624240C1642E917BDE3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-museum-hamburg" xlink_title="Zoological Museum Hamburg">
<collectionCode id="6AA950A170A77FFDF4BA94261007FD68" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" type="Museum">ZMH</collectionCode>
</named-content>
844689.
</paragraph>
<caption id="4ADD82D1F37BF6D0616D065150162573" ID-DOI="10.3897/zookeys.1226.140239.figure11" ID-arpha="B2B076A9-5169-576E-9374-EA271296FC01" httpUri="https://binary.pensoft.net/fig/1243042" startId="F11">
<paragraph id="F2E0CAD5CFFF2980BFD901A82E4B26A3">
<label id="8ADA058F00590D4BC4F10C77B4432760">Figure 11.</label>
</paragraph>
<paragraph id="01F9B5D5D170DBD185C83730A8C77D11">
<taxonomicName id="A4FAC0053C66B375C0BD8FA0243C56D2" authority="Laboissiere, 1936" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="F8E9F4718F2257667264D1337865A5D8" italics="true">Paleosepharia truncata</emphasis>
<bibRefCitation id="EAEB5119915EED80F7316CF9745354FE" DOI="10.1080/21686351.1936.12278963" author="Laboissière V" journalOrPublisher="Annales de la Société Entomologique de France" pagination="239-262" refId="B19" refString="Laboissiere V (1936) Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et especes (5 e partie). Annales de la Societe Entomologique de France 105: 239-262. https://doi.org/10.1080/21686351.1936.12278963" title="Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et espèces (5 e partie)." volume="105" year="1936">Laboissière, 1936</bibRefCitation>
</taxonomicName>
(paratype). Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EOEAG" type="material">
<paragraph id="1093F0BD2DE9B1F82F10F88BE51F9CF8">
<heading id="43C701FD4B301CA69BB506ED37E7772A" reason="title">Other specimens examined.</heading>
</paragraph>
<paragraph id="F6EFCBFB8A29FD4665EFF7B53635A952">
<materialsCitation id="805C8A561C888719E8AD8D41D19E7C3A" collectingDate="1958-06-22" collectionCode="IZCAS" collectorName="Shuyong Wang" country="China" county="Mengzhe" elevation="1700" location="Yunan Province" municipality="Xishuangbanna" specimenCount="1" specimenCount-male="1">
<specimenCount id="C1274D67EC2E840A6A95467470506ADD" count="1" type="male">1 ♂</specimenCount>
,
<collectingCountry id="8F7CDA894AD5807CC44C419954B8859D" name="China">China</collectingCountry>
,
<location id="9D24951A6077664D8A8F13B998C40983" LSID="urn:lsid:plazi:treatment:B27B66F212A05BF4A53FCC4EAB8716F4:9D24951A6077664D8A8F13B998C40983" country="China" county="Mengzhe" municipality="Xishuangbanna" name="Yunan Province">Yunan Province</location>
,
<collectingMunicipality id="8DC3990C76ED099B66E0D226504D8F2E">Xishuangbanna</collectingMunicipality>
,
<collectingCounty id="F17491092A7578CBF776ACE95E6065AF">Mengzhe</collectingCounty>
,
<elevation id="2D30F3539B09F7D8572A4EBC46EC5070" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<quantity id="D25C5C55B618C77F3555D4C77C02BFC9" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</quantity>
a. s. l.
</elevation>
,
<date id="A640C2BEBD1AF568384EE4AF6F0ABDE8" value="1958-06-22">
<collectingDate id="DFA4F624FAA02360B66F4A4702ADBDE9" value="1958-06-22">22 Jun. 1958</collectingDate>
</date>
,
<collectorName id="8A0C0B458A8A2FCB5F764CC57ECA7F78">Shuyong Wang</collectorName>
leg.,
<named-content id="DCBBED419A8B4741AF0D9F1E217C2564" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="A9F1125E536F21AB3E1307A7F0B9EF82" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="8CCDFC4D9229132490D8408AC3C047AE" collectingDate="1958-06-04" collectionCode="IZCAS" collectorName="Shuyong Wang" country="China" county="Menghun" elevation="1200" location="Yunan Province" municipality="Xishuangbanna" specimenCount="1" specimenCount-female="1">
<specimenCount id="835D1AADC0CB4965961FB6C62F2492E3" count="1" type="female">1 ♀</specimenCount>
,
<collectingCountry id="7D65110852F175E420258C4330AC78A8" name="China">China</collectingCountry>
,
<location id="9D8C1653FB2808D5F952839081695BAC" LSID="urn:lsid:plazi:treatment:B27B66F212A05BF4A53FCC4EAB8716F4:9D8C1653FB2808D5F952839081695BAC" country="China" county="Menghun" municipality="Xishuangbanna" name="Yunan Province">Yunan Province</location>
,
<collectingMunicipality id="6C4086A91AEE7E6454419DE5ACDA4385">Xishuangbanna</collectingMunicipality>
,
<collectingCounty id="E44ECFCA47D29A0D8DA319054AA2A70C">Menghun</collectingCounty>
,
<elevation id="4F33F9A1022965E8FC6067B765AEBF06" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<quantity id="A855398FC6DBD8504027E51141B60479" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
a. s. l.
</elevation>
,
<date id="7FCC126D93BD1F50A4D5991B3290D7F5" value="1958-06-04">
<collectingDate id="76BDA2787D24BFE78E3DE1F545EBFB6F" value="1958-06-04">4 Jun. 1958</collectingDate>
</date>
,
<collectorName id="F7531F76BBDCFCD9423BF89D4034B960">Shuyong Wang</collectorName>
leg.,
<named-content id="30144B5914824288D0CA42EFD7E7E664" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="121D2340BE7070B85566851C35D25346" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="40D0F8F9074C31FF4620E68C6379E44E" collectingDate="1958-06-08" collectionCode="IZCAS" collectorName="Shuyong Wang" country="China" county="Menga" elevation="1065" location="Yunan Province" municipality="Xishuangbanna" specimenCount="1" specimenCount-female="1">
<specimenCount id="B103D991F13EB65616E04618A866E541" count="1" type="female">1 ♀</specimenCount>
,
<collectingCountry id="BC507B3A67D142674390DBEFDA5CB7DF" name="China">China</collectingCountry>
,
<location id="B21479704E117057895B077E2CB6EC2C" LSID="urn:lsid:plazi:treatment:B27B66F212A05BF4A53FCC4EAB8716F4:B21479704E117057895B077E2CB6EC2C" country="China" county="Menga" municipality="Xishuangbanna" name="Yunan Province">Yunan Province</location>
,
<collectingMunicipality id="09792F074B52056A9472ED18D84479FF">Xishuangbanna</collectingMunicipality>
,
<collectingCounty id="24C29289EA071E7FEA5AA3407F5E2396">Menga</collectingCounty>
,
<elevation id="8E4A7F98E66BA940D4088A1A7274E5AB" metricMagnitude="3" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" unit="m" value="1065.0" valueMax="1080.0" valueMin="1050.0">
<quantity id="046393AD4631F8D01824436C00D2FED4" metricMagnitude="3" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" unit="m" value="1065.0" valueMax="1080.0" valueMin="1050.0">10501080 m</quantity>
a. s. l.
</elevation>
,
<date id="18A1B182B33E7BD1BB386476CEC6AD2F" value="1958-06-08">
<collectingDate id="3A8D4068ADC918D9885016DF7DB37740" value="1958-06-08">8 Jun. 1958</collectingDate>
</date>
,
<collectorName id="41A6DB1BA3C5A9277795672FF1A2B6C2">Shuyong Wang</collectorName>
leg.,
<named-content id="7B3E24C5F9157DFE45E223AF137D8689" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="65D22E33EF03F0BC28E72D465EE2D4D4" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="CD68E0D8B70F508911E7CD66A3561C2D" collectingDate="1958-06-10" collectionCode="IZCAS" collectorName="Shuyong Wang" country="China" county="Mengzhe" elevation="1700" location="Yunan Province" municipality="Xishuangbanna" specimenCount="1" specimenCount-male="1">
<specimenCount id="ED587899AB49E61CDBCC968037AEB93E" count="1" type="male">1 ♂</specimenCount>
,
<collectingCountry id="DBB03F84156A340605B812ECFD36FA05" name="China">China</collectingCountry>
,
<location id="E8E87264FFB81365DF49A72C77AC7984" LSID="urn:lsid:plazi:treatment:B27B66F212A05BF4A53FCC4EAB8716F4:E8E87264FFB81365DF49A72C77AC7984" country="China" county="Mengzhe" municipality="Xishuangbanna" name="Yunan Province">Yunan Province</location>
,
<collectingMunicipality id="FA828FFFB8CC87EF01F3160D06B764B2">Xishuangbanna</collectingMunicipality>
,
<collectingCounty id="8D417223B11A22FF76F546615090763A">Mengzhe</collectingCounty>
,
<elevation id="A2EDBCAE7B75D401B191467C9A6CA4EA" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<quantity id="4AE752731412B973F0840D842BA2767A" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</quantity>
a. s. l.
</elevation>
,
<date id="E1082B120065D078EA31FF6BB432C751" value="1958-06-10">
<collectingDate id="63C4C294A4BC85F594741AF674EFC005" value="1958-06-10">10 Jun. 1958</collectingDate>
</date>
,
<collectorName id="D75F6199E822E3F6B70BEC2E8EF31086">Shuyong Wang</collectorName>
leg.,
<named-content id="696CB2E881A3E4923C7FFCC3E4C6E7C2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="8B00AA97B3D8ADC60AA5B1B59DE0ABFF" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
; •
<materialsCitation id="C054931448372361A5FB1991BC5EBCE5" collectingDate="1958-06-15" collectionCode="IZCAS" collectorName="Shuyong Wang" country="China" county="Mengzhe" elevation="1200" location="Yunan Province" municipality="Xishuangbanna" specimenCount="1" specimenCount-female="1">
<specimenCount id="959C32EF2A3713D67B4D067DFDF84CE7" count="1" type="female">1 ♀</specimenCount>
,
<collectingCountry id="2DFFD7263AA423B1DE3229796D84F44E" name="China">China</collectingCountry>
,
<location id="F49B2FADF0EC98DF9117DE956B877E20" LSID="urn:lsid:plazi:treatment:B27B66F212A05BF4A53FCC4EAB8716F4:F49B2FADF0EC98DF9117DE956B877E20" country="China" county="Mengzhe" municipality="Xishuangbanna" name="Yunan Province">Yunan Province</location>
,
<collectingMunicipality id="960253D500207BF189390230934F03CE">Xishuangbanna</collectingMunicipality>
,
<collectingCounty id="5D56559CA7925F2AE6217992E3BC2B3F">Mengzhe</collectingCounty>
,
<elevation id="31154192E763C754F1E3EC7DC91F2016" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<quantity id="1FCE1C6671BB3D55146E231A497F4BDF" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
a. s. l.
</elevation>
,
<date id="F849E62686D8F83082CFC3714CD37A6E" value="1958-06-15">
<collectingDate id="FF017979F3090695FB3FEA642774C813" value="1958-06-15">15 Jun. 1958</collectingDate>
</date>
,
<collectorName id="FB100A1636645AF9671F0DBEA388CD83">Shuyong Wang</collectorName>
leg.,
<named-content id="681FB8119A85D9C3E331388970F818F0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences, Beijing">
<collectionCode id="182658078C710CA91AB5989C1FA1DBEA" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<caption id="B890799941CD6A64E7D233331BD49BA9" ID-DOI="10.3897/zookeys.1226.140239.figure12" ID-arpha="26C7C624-74AE-5751-9B2A-E211827B4912" httpUri="https://binary.pensoft.net/fig/1243043" startId="F12">
<paragraph id="B2D922921D6B336EC9F757FA772360E2">
<label id="D3FF7478E832BA868541BD7C27D4D271">Figure 12.</label>
</paragraph>
<paragraph id="4AF711DCBFC53C3221BEF90B3173B4B4">
<taxonomicName id="7DFD5F788BF91412ACC4636DD77FCD90" authority="Laboissiere, 1936" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="327F602D3200716D2EF4F215226C1F17" italics="true">Paleosepharia truncata</emphasis>
<bibRefCitation id="CAA4CFB12F7D20388F3D88EAACB2BF0F" DOI="10.1080/21686351.1936.12278963" author="Laboissière V" journalOrPublisher="Annales de la Société Entomologique de France" pagination="239-262" refId="B19" refString="Laboissiere V (1936) Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et especes (5 e partie). Annales de la Societe Entomologique de France 105: 239-262. https://doi.org/10.1080/21686351.1936.12278963" title="Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et espèces (5 e partie)." volume="105" year="1936">Laboissière, 1936</bibRefCitation>
</taxonomicName>
(male)
<emphasis id="E87F45C706F601FC539C9D1AC0EAC535" bold="true">A C</emphasis>
habitus
<emphasis id="9C6E5289F5A7DAA8BDCC1F8304779CE6" bold="true">D F</emphasis>
aedeagus
<emphasis id="601504E2CA74DF9FB3C9C6B68268E1F8" bold="true">G</emphasis>
depression of the elytra of the male.
<emphasis id="EF83CF5788F8659F2D8DB3F249C23D6B" bold="true">A, D</emphasis>
dorsal view
<emphasis id="3229A4FBA59FAAE57F57C8D9EC36417E" bold="true">B, E</emphasis>
ventral view
<emphasis id="2DCCEB9B600C5414A6C21F283D24ABCD" bold="true">C, F</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="C09B2E4F75ACA8DB1B3698EFF77CA9EA" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="5EEED351AD44A05909B5D8633F6F228F" bold="true">D F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQGAG" type="description">
<paragraph id="0DF1152E1F8125B9C96315025737EC2F">
<heading id="E1B2D081DE6E41EE72F97A35DD3CD465" reason="title">Description.</heading>
</paragraph>
<paragraph id="B16B4F424A3DE28443B6942E71C52AEA">
<emphasis id="5D263246930A1BA9F5EC2FFCCD1CFA2C" bold="true">Male</emphasis>
: Length:
<quantity id="F1802AD26747BC2632FEED0294617256" metricMagnitude="-3" metricUnit="m" metricValue="6.8" unit="mm" value="6.8">6.8 mm</quantity>
, width:
<quantity id="4C4FF29A8A2C85717CE14AFB77936487" metricMagnitude="-3" metricUnit="m" metricValue="3.8" unit="mm" value="3.8">3.8 mm</quantity>
.
</paragraph>
<caption id="A9C47F158939C4AFE1AEAC0905C6F18C" ID-DOI="10.3897/zookeys.1226.140239.figure13" ID-arpha="1C46CFAF-0978-5EE1-BEA7-CB931F8AF63B" httpUri="https://binary.pensoft.net/fig/1243044" startId="F13">
<paragraph id="B8F0D597016E70D4CD1F0E2ADADEC7E1">
<label id="D3A65824FB072DE45A7C4E7ECDF1B37E">Figure 13.</label>
</paragraph>
<paragraph id="E9D5240273EAB4A50E1D31F858A50D0B">
<taxonomicName id="B17372628228CA483403B93EA00A986F" authority="Laboissiere, 1936" authorityName="Laboissiere" authorityYear="1936" class="Insecta" family="Chrysomelidae" genus="Paleosepharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="147251F76E852B526569E0EC4D68E59B" italics="true">Paleosepharia truncata</emphasis>
<bibRefCitation id="B7D4352FE2CC721DBD257175EB396E5F" DOI="10.1080/21686351.1936.12278963" author="Laboissière V" journalOrPublisher="Annales de la Société Entomologique de France" pagination="239-262" refId="B19" refString="Laboissiere V (1936) Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et especes (5 e partie). Annales de la Societe Entomologique de France 105: 239-262. https://doi.org/10.1080/21686351.1936.12278963" title="Observations sur les Galerucini asiatiques principalement du Tonkin et du Yunnan et descriptions de nouveaux genres et espèces (5 e partie)." volume="105" year="1936">Laboissière, 1936</bibRefCitation>
</taxonomicName>
(female)
<emphasis id="D570F5E3C473E5C7893EBFF6DF995F5D" bold="true">A-C</emphasis>
habitus
<emphasis id="F29721740B8D5F931C61F9CC9DD59355" bold="true">D</emphasis>
vaginal palps
<emphasis id="80396039BB720FFD8B92303ECA749D99" bold="true">E</emphasis>
spermatheca
<emphasis id="0F6C21076DB17A70322B213FFA56C283" bold="true">F</emphasis>
apex of abdomen of the female.
<emphasis id="9A9CE82DCE2BD9D146A627CAE38D700D" bold="true">A</emphasis>
dorsal view
<emphasis id="1BED34865BDB5343A3661DAA7F1C2193" bold="true">B</emphasis>
ventral view
<emphasis id="DBFA646446A8DA3518CEA85BB64BE03F" bold="true">C</emphasis>
lateral view. Scale bars: 1 mm (
<emphasis id="C1D04C25B2AC1A2B6CB250A56BE0D0A7" bold="true">A C</emphasis>
); 0.5 mm (
<emphasis id="D85AD99E60A41EA074F37423829C00A0" bold="true">D, E</emphasis>
).
</paragraph>
</caption>
<paragraph id="68E1A7D5C9E86246A70D5F12222DA5A5">General color yellow or yellowish-brown, metathorax reddish-brown, basal part of inner margin of epipleuron and base of first metatarsomere black.</paragraph>
<paragraph id="952D6CB8D396D2964869FD4045DC38DA">Vertex with sparse punctures. Frontal tubercles transverse, extending downward between antenna bases. Antennae slender, equal to length of body. First antennomere shiny bare, rod-shaped, second to eleventh antennomeres with short hairs, second antennomere shortest, third antennomere 1.9 × as long as second, fourth antennomere about 1.4 × as long as third, fifth to seventh antennomeres slightly longer than fourth, eighth to eleventh antennomeres equal in length to fourth. Pronotum about 1.55 × as wide as long, lateral margins slightly curved, basal border slightly convex, apical border slightly concave, disc strongly convex, with dense punctures. Scutellum triangular, smooth, impunctate. Elytra wider than pronotum, humeri convex, disc with dense, minute punctures; with elliptical postscutellar depression that extends obliquely backwards. Elytral epipleuron broad at base, continues towards apex. Each tibia with distinct spur at apex, segment 1 of hind tarsi longer than remaining segments combined. Aedeagus gradually narrowed towards apex, slightly curved at apex in lateral view. lateral and median spiculae clearly visible, ventral spiculae weakly. Median spiculae with two pairs of slender sclerites.</paragraph>
<paragraph id="BF8B92C4E976F9A289A19C1BC7D5C6BD">
<emphasis id="D57CD03D8FF4E2D246489D3DDF623833" bold="true">Female</emphasis>
: Length:
<quantity id="C81A555FA1347A1D3375AB908969D371" metricMagnitude="-3" metricUnit="m" metricValue="7.8" metricValueMax="8.2" metricValueMin="7.4" unit="mm" value="7.8" valueMax="8.2" valueMin="7.4">7.48.2 mm</quantity>
, width:
<quantity id="FF30F9C98816E9814321405E495B53CD" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" unit="mm" value="4.3" valueMax="4.4" valueMin="4.2">4.24.4 mm</quantity>
(mean length
<quantity id="D6E2453BDBEF176DC28BDE18A2DF7698" metricMagnitude="-3" metricUnit="m" metricValue="7.7" unit="mm" value="7.7">7.7 mm</quantity>
, mean width
<quantity id="5C829B6BE05BA247372AD46151B7F2FD" metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
,
<emphasis id="441D036A838B8B51FE8490F31005D3CF" bold="true" italics="true">N</emphasis>
= 4).
</paragraph>
<paragraph id="B0054406B0BB6CE79BF93F3D194553C5">Elytra lacking impression. Apex of abdomen with rounded concave. Spermatheca with big and rounded nodulus, middle part short, cornu long and strongly curved. Vaginal palps gradually widened towards apex, each palp with 5 setae placed at apex, additional 2 setae subapically.</paragraph>
</subSubSection>
<subSubSection id="SECID0EMIAG" type="distribution">
<paragraph id="84705A2B433D6BA3BD2902CA574A44A5">
<heading id="CCAD28BD6E59D6AAD079653060A79A71" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="2E05B03E9575E31ABBAA73C08D777946">
<collectingCountry id="2900AE1A75AD33968F70809E88756149" name="China">China</collectingCountry>
:
<collectingRegion id="3A25DC40C25481495FCD34F87A7529A6" country="China" name="Yunnan">Yunnan</collectingRegion>
;
<collectingCountry id="1596CDAA12A08799B288964FCF98DC55" name="Vietnam">Vietnam</collectingCountry>
,
<collectingCountry id="C5FF546D76D49158494F2129861152D7" name="Thailand">Thailand</collectingCountry>
,
<collectingCountry id="89265FB1DD195BD0624834DE054A3C68" name="Laos">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,632 @@
<document id="108E95A18B5DB933AE89CFE23DB7FA1A" ID-CLB-Dataset="308017" ID-DOI="10.3897/zookeys.1226.137752" ID-GBIF-Dataset="616e2c2f-f236-44da-b264-bb87eaf513a7" ID-ZooBank="6891FA95-5A4A-456F-8AC3-5A14939628E8" ID-publisher-id="137752" URI-arpha="5C3BF073-C8DB-59FD-9484-28BF518C616A" URI-zoobank="http://zoobank.org/6891FA95-5A4A-456F-8AC3-5A14939628E8" article-type="research-article" checkinTime="1738866260897" checkinUser="pensoft" docAuthor="Mu, Yannan, Wang, Luyu, Zhang, Feng &amp; Zhang, Zhisheng" docDate="2025" docId="CA91E2E8A5E25AFFA9482937CE5FBB61" docLanguage="en" docName="ZooKeys 1226: 121-137" docOrigin="ZooKeys 1226" docSource="https://zookeys.pensoft.net/article/137752/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Mutatus gelao Mu, Wang, Zhang &amp; Zhang, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="137" masterDocId="5C3BF073C8DB59FD948428BF518C616A" masterDocTitle="Mutatus gen. nov., a new genus of Phrurolithidae (Arachnida, Araneae) from China, with descriptions of four new species" masterLastPageNumber="137" masterPageNumber="121" pageNumber="121" updateTime="1738866914701" updateUser="ExternalLinkService">
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<mods:title id="E6B6A0E8B730D4DCACC0E0546FC670FF">Mutatus gen. nov., a new genus of Phrurolithidae (Arachnida, Araneae) from China, with descriptions of four new species</mods:title>
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<mods:namePart id="6CA5BC08502D14E5F45D37E393B3DE37">Mu, Yannan</mods:namePart>
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<mods:affiliation id="06276FED3DD5C1F7C5654B1A12B7A410">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China &amp; Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="B71783CF986F01BBE314A926B990F3BC">Wang, Luyu</mods:namePart>
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<mods:affiliation id="A4B7BE2FB19D47D33AEB958F2AEC20A7">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:namePart id="FDF9C6358AD3B83B4921B7C5C9ECCB54">Zhang, Feng</mods:namePart>
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<mods:affiliation id="A1EE5C9BC274127EE677B2D07507407A">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="E640A180E05BB7F7E0F434E6949EC0F7">Zhang, Zhisheng</mods:namePart>
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<mods:affiliation id="99C8E70DB927E001A969C09A8E790575">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<treatment id="CA91E2E8A5E25AFFA9482937CE5FBB61" ID-DOI="http://doi.org/10.5281/zenodo.14827168" ID-Zenodo-Dep="14827168" ID-arpha="CA91E2E8-A5E2-5AFF-A948-2937CE5FBB61" ID-zoobank="https://zoobank.org/C66AD91F-662D-45DE-9698-204FD511F7BD" LSID="urn:lsid:plazi:treatment:CA91E2E8A5E25AFFA9482937CE5FBB61" LSID-ZBK="urn:lsid:zoobank.org:act:C66AD91F-662D-45DE-9698-204FD511F7BD" httpUri="http://treatment.plazi.org/id/CA91E2E8A5E25AFFA9482937CE5FBB61">
<subSubSection id="EC8FB09EF9A11FC00A250F5BDABAF64D" type="nomenclature">
<paragraph id="65BD6D2A1FBA71BDB3031FDEB8391064">
<taxonomicName id="E2CBCC770B9351C8D1EBF93535C33C83" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Phrurolithidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="gelao" status="sp. nov.">
<emphasis id="7221D625467F3C4B57CF68A6689382CF" italics="true">Mutatus gelao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFD06D42EF99E8C317D48FB108525E15" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9DDADF2556446F044C234667078BF541" type="description">
<paragraph id="E2A4957423B19FB17043981AB24B2ED5">
<figureCitation id="5575DA1A5F481B50725B707FFED71912" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Mutatus gelao sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure1" httpUri="https://binary.pensoft.net/fig/1243800">Chinese name: 仡佬幻蛛 Figs 1</figureCitation>
,
<figureCitation id="C25081E2DF4A76792474B270E9B57759" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBPAC" type="type material">
<paragraph id="D44AB12027FE2EA88FDC2D34EDCCC410">
<heading id="42E2A780272E606E590724D898D3193F" reason="title">Type material.</heading>
</paragraph>
<paragraph id="FCD75AFAB6DACC8869B47D031F6FE5A0">
<materialsCitation id="546629CBF40BDDEC661087B9114C3E1F" collectingDate="2023-05-08" collectorName="Z. Y. Li &amp; W. H. Wang" country="China" county="Shiqian County" elevation="647" latitude="27.034199" location="Gelao Village" longLatPrecision="1" longitude="108.15193" municipality="Tongren City" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="FC5E28A36C0C28478166A4567FB374A7" bold="true" italics="true">
<typeStatus id="86FE4A4BC5BD6B4796EC106149A9CE0F" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="4C6C9CF3423B47E2656DA09DC06D46B4" name="China">China</collectingCountry>
<specimenCount id="95E2B918A6F5F7CCF3BBEEE4FD4491B7" count="1" type="male"></specimenCount>
;
<collectingRegion id="A7C3A0567393EF8BFC0932C8512D8889" country="China" name="Guizhou">Guizhou Province</collectingRegion>
,
<collectingMunicipality id="AF4B1B8893BCC65BE5ED1E8B9A3627B3">Tongren City</collectingMunicipality>
,
<collectingCounty id="BFA3F0EA2A766DC5701748B1E8F8E4B8">Shiqian County</collectingCounty>
,
<location id="DA7F9D4DD5C11D23AA610F709DF697FE" LSID="urn:lsid:plazi:treatment:CA91E2E8A5E25AFFA9482937CE5FBB61:DA7F9D4DD5C11D23AA610F709DF697FE" country="China" county="Shiqian County" latitude="27.034199" longLatPrecision="1" longitude="108.15193" municipality="Tongren City" name="Gelao Village" stateProvince="Guizhou">Gelao Village</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPPAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[108.151928,27.034200]}">
<geoCoordinate id="8DE2AA2AAA163B1FAE8CA7E1B0DFD2D6" degrees="27" direction="north" minutes="2" orientation="latitude" precision="1" seconds="3.12" value="27.034199">27 ° 2 ' 3.12 &quot; N</geoCoordinate>
,
<geoCoordinate id="561E7B47368063FBD56FB7F92181B742" degrees="108" direction="east" minutes="9" orientation="longitude" precision="1" seconds="6.94" value="108.15193">108 ° 9 ' 6.94 &quot; E</geoCoordinate>
</named-content>
, elev.
<quantity id="A044FEB8B3263AEAE436BC86AE317E75" metricMagnitude="2" metricUnit="m" metricValue="6.47" unit="m" value="647.0">
<elevation id="D0ED68FE33C0A7DF5B1863CC6410187A" metricMagnitude="2" metricUnit="m" metricValue="6.47" unit="m" value="647.0">647 m</elevation>
</quantity>
,
<date id="57D013F3AD7D2C9E5D6D0C2735E98718" value="2023-05-08">
<collectingDate id="2C607F8494C8E6DC312D36113695EA07" value="2023-05-08">08 May 2023</collectingDate>
</date>
;
<collectorName id="46CDBAAEB6D1ED6C6657DB75FD58FB1F">Z. Y. Li</collectorName>
,
<collectorName id="15769FDFE4239884AB44767FC0E36A06">W. H. Wang</collectorName>
leg.
</materialsCitation>
<emphasis id="C7737143E7E8DEBD31FECDE8B64CB15D" bold="true" italics="true">
<typeStatus id="39A368AA32C044985087250773AD9576" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCount id="6D303C931C7543450CDF6AFE29709A2D" count="1" type="female">1 ♀</specimenCount>
, with same data as
<typeStatus id="FA43F8D69DBCE84DF4874CD40D18FC2F" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXPAC" type="etymology">
<paragraph id="CE241E96AC7326EE5B2C8D2AFC9A294C">
<heading id="6EFA02D22E405F5514C1977A43ACA74A" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="095CBA3953C2EE69A821363FDA775308">The specific name comes from the word “ Gelao ”. The name is one of the Chinese ethnic minorities that live adjacent to the area inhabited by the new species; a noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="SECID0E3PAC" type="diagnosis">
<paragraph id="601E655344AB93475471B7563BB43FF0">
<heading id="82BD25F83DBDE1AD85C4547BAB6A245B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="2FA785079AA815EBCD71A7599A9D12CD">
The new species can be differentiated from
<taxonomicName id="351D5908CFBEDCC2604C73BB8E24BE1A" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tujia" status="sp. nov.">
<emphasis id="8C56207C6F243FDDD54AC2F944F8634B" italics="true">Mutatus tujia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B30A7300A1B5579138405B7BBCFFE10B" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4F3BC8B2E58A6131FD2536143E09335D" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tianyan" status="sp. nov.">
<emphasis id="28552A010B17CC771A54FA7F87C94993" italics="true">M. tianyan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56F9A9FF68733E8DDF2B8B99348A210E" rank="species">sp. nov.</taxonomicNameLabel>
by the ventrally bulging tibia of the male palp (vs not inflated; compare Fig.
<figureCitation id="3E65F39835AFA8403579411548CC3247" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 C</figureCitation>
with Figs
<figureCitation id="44460BC5957EB194B4610C5302DD8FF8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 B</figureCitation>
,
<figureCitation id="C3ABA5E2BFCB14655EF5BCEF469DB92A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 B</figureCitation>
), almost bifurcate tip of the retrolateral tibial apophysis (vs smooth and tapering from base to tip; compare Fig.
<figureCitation id="5E9499AD3CCEAAE2D5B28419749C0520" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 C</figureCitation>
with Figs
<figureCitation id="315E1FA37AEDE115130B024942F2D9BE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 C</figureCitation>
,
<figureCitation id="ED5039CC3A90FAFA7E049C4DEEB5188B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 D</figureCitation>
), spade-shaped tegular sheet (vs tobacco-pipe-shaped, compare Fig.
<figureCitation id="CE3A7C252A148CE9B5F8C34E7592DC6D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 B</figureCitation>
with Figs
<figureCitation id="2C4F7C480FD5F4BD88C5F85113E2A220" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 D</figureCitation>
,
<figureCitation id="435C05A6EAE917308900CE63B36D5711" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 C</figureCitation>
), spiculate prolateral tegular apophysis (vs truncated, compare Fig.
<figureCitation id="A7D5355A638004C956BED02C2F563B56" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 B</figureCitation>
with Figs
<figureCitation id="0FCC0E8AA1D9A9DCB1F33371B49E5277" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 D</figureCitation>
,
<figureCitation id="E9EC605C23E0A048CB01336378404202" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 C</figureCitation>
), copulatory ducts which form a right angle with connecting tubes (vs forming a obtuse angle; compare Fig.
<figureCitation id="90BB1902887FEA2D35BD5AD500851C78" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 H</figureCitation>
with Figs
<figureCitation id="2FE3864D9F39E25AAA238161957F741B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 F</figureCitation>
,
<figureCitation id="96A03FBDBA23DDB7EBA821CDD7D00042" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 F</figureCitation>
), and larger and deeper, kidney-shaped atria (vs atria oval, small, and shallow; compare Fig.
<figureCitation id="6F2409019B9F53DA1DD8AE41B90997E2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 G</figureCitation>
with Figs
<figureCitation id="A19306EE9BBFD2570619641DC4126346" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Copulatory organs of Mutatus tujia sp. nov. A male left palp, prolateral view B same, retrolateral view C same, dorsal view D same, ventral view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure4" httpUri="https://binary.pensoft.net/fig/1243803">4 E</figureCitation>
,
<figureCitation id="13B9EDF2FE26C419816B60C296CF2999" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Copulatory organs of Mutatus tianyan sp. nov. A male left palp, prolateral view B same, retrolateral view C same, ventral view D same, dorsal view E epigyne, ventral view F same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure6" httpUri="https://binary.pensoft.net/fig/1243805">6 E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBDAE" type="description">
<paragraph id="E30343664DE8CC7F932C50174ABFC879">
<heading id="304461F2DEAE941ADFBE3423B9B62142" reason="title">Description.</heading>
</paragraph>
<paragraph id="EEAB9801CCF3AE5B4FA18BA3B6E48CCE">
Male (
<typeStatus id="DCB2B2B7F4578F59047813D689AADCCF" type="holotype">holotype</typeStatus>
, Fig.
<figureCitation id="BA7C2D1EC576472A5B0343981FE3F117" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Mutatus gelao sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure1" httpUri="https://binary.pensoft.net/fig/1243800">1 A, B</figureCitation>
). Total length 3.35, carapace 1.73 long, 1.55 wide; abdomen 1.62 long, 1.32 wide. Eye sizes and interdistances:
<abbrev id="ABBRID0ELDAE" xlink_title="anterior median eye">AME</abbrev>
0.11,
<abbrev id="ABBRID0EPDAE" xlink_title="anterior lateral eye">ALE</abbrev>
0.13,
<abbrev id="ABBRID0ETDAE" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0EXDAE" xlink_title="posterior lateral eye">PLE</abbrev>
0.10;
<abbrev id="ABBRID0E2DAE" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0E6DAE" xlink_title="anterior median eye">AME</abbrev>
0.04,
<abbrev id="ABBRID0EDEAE" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0EHEAE" xlink_title="anterior lateral eye">ALE</abbrev>
0.01,
<abbrev id="ABBRID0ELEAE" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0EPEAE" xlink_title="anterior lateral eye">ALE</abbrev>
0.27,
<abbrev id="ABBRID0ETEAE" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EXEAE" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0E2EAE" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E6EAE" xlink_title="posterior lateral eye">PLE</abbrev>
0.08,
<abbrev id="ABBRID0EDFAE" xlink_title="posterior lateral eye">PLE</abbrev>
<abbrev id="ABBRID0EHFAE" xlink_title="posterior lateral eye">PLE</abbrev>
0.41,
<abbrev id="ABBRID0ELFAE" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0EPFAE" xlink_title="posterior lateral eye">PLE</abbrev>
0.09.
<abbrev id="ABBRID0ETFAE" xlink_title="eye area width">EAW</abbrev>
0.56,
<abbrev id="ABBRID0EXFAE" xlink_title="cephalic region width">CRW</abbrev>
0.81,
<abbrev id="ABBRID0E2FAE" xlink_title="eye area width">EAW</abbrev>
/
<abbrev id="ABBRID0E6FAE" xlink_title="cephalic region width">CRW</abbrev>
0.69.
<abbrev id="ABBRID0EDGAE" xlink_title="cephalic region width">CRW</abbrev>
/
<abbrev id="ABBRID0EHGAE" xlink_title="carapace width">CW</abbrev>
0.52.
<abbrev id="ABBRID0ELGAE" xlink_title="median ocular area">MOA</abbrev>
0.27 long, anterior width 0.24, posterior width 0.26. Clypeal height 0.15,
<abbrev id="ABBRID0EPGAE" xlink_title="clypeal height">
<collectingCountry id="00FAD6798807194732B08C58D2716C12" name="Switzerland">CH</collectingCountry>
</abbrev>
/
<abbrev id="ABBRID0ETGAE" xlink_title="anterior median eye">AME</abbrev>
1.36. Labium 0.22 long, 0.24 wide. Sternum 0.96 long, 0.96 wide. Carapace oval, cervical groove inconspicuous. Sternum, labium, and endite brown. Abdomen dark brown, smooth, with large dorsal scutum covering almost entire abdomen. Legs brown.
</paragraph>
<caption id="74001878DA8D712E3DE38D695AC94DE1" ID-DOI="10.3897/zookeys.1226.137752.figure1" ID-arpha="1A3B43BD-5C2E-5BA2-8389-F878950C07F1" httpUri="https://binary.pensoft.net/fig/1243800" startId="F1">
<paragraph id="992EACE432F80D0DF0AAC7157200AD99">
<label id="9DC4BDE9DBD044065A0DCF6AF8B6035D">Figure 1.</label>
</paragraph>
<paragraph id="568CD6058A55196E3032801E177288AD">
Habitus of
<taxonomicName id="EDFE1F289C691998934A2F098C184402" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gelao" status="sp. nov.">
<emphasis id="0257C804EE79EFB90E0FBD70C11BC687" italics="true">Mutatus gelao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="060362DE2726BCB1D90A598F4C0D49C5" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B3372018863675C12320AAAB305B419D" bold="true">A</emphasis>
male holotype, dorsal view
<emphasis id="18FACB67C2D714B33E88C68126B8C092" bold="true">B</emphasis>
same, ventral view
<emphasis id="E2C1D0168EB2C5DFF392B27238016FB8" bold="true">C</emphasis>
female paratype, dorsal view
<emphasis id="9C6A533A2BD15C675442284515860804" bold="true">D</emphasis>
same, ventral view.
</paragraph>
</caption>
<paragraph id="880FB0DB0CADD320978087B5A5CA37DC">
Palp (Fig.
<figureCitation id="1A6F4BA7C95E220685F1832EDE3600D0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 A F</figureCitation>
). Femoral apophysis (
<abbrev id="ABBRID0E4HAE" xlink_title="femur apophysis">FA</abbrev>
) well developed, thumb-shaped, located at middle part. Patella about 1 / 3 length of femur, as long as tibia. Tibia longer than wide, slightly bulged ventrally, retroventral apophysis (
<abbrev id="ABBRID0EBIAE" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
) tail-shaped, tip slightly curved; retrolateral apophysis (
<abbrev id="ABBRID0EFIAE" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
) longer and wider than
<abbrev id="ABBRID0EJIAE" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
, tip bifurcated. Bulb pyriform, sperm duct (
<abbrev id="ABBRID0ENIAE" xlink_title="sperm duct">SD</abbrev>
) V-shaped. Prolateral tegular apophysis (
<abbrev id="ABBRID0ERIAE" xlink_title="Prolateral tegular apophysis">PTA</abbrev>
) coniform, slightly curved, originating at nearly 9 oclock position. Tegular sheet (
<abbrev id="ABBRID0EVIAE" xlink_title="tegular sheet">TS</abbrev>
) spade-shaped, anterior part with curved tip, posterior part tongue-shaped. Embolus (E) needle-like, tapering towards tip, slightly curved.
</paragraph>
<caption id="936F35D84C54FAAF63075253E50B094C" ID-DOI="10.3897/zookeys.1226.137752.figure2" ID-arpha="5AF88144-52C2-58D7-8DFF-63CFCFEAAF89" httpUri="https://binary.pensoft.net/fig/1243801" startId="F2">
<paragraph id="E4CF79DED1323D7B64E5DDBD112C483F">
<label id="4B4D6B7D220AFBC7A1305593C0CD1F96">Figure 2.</label>
</paragraph>
<paragraph id="D80183397333392322C0574A49D076A2">
Copulatory organs of
<taxonomicName id="14D146A045B89D60942C24524EB4E44D" authorityName="Mu &amp; Wang &amp; Zhang &amp; Zhang" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Mutatus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="gelao" status="sp. nov.">
<emphasis id="275B4366C4F999EDA7D8AF8756398C46" italics="true">Mutatus gelao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F4BF9F8AD984546049B2C23FD76D9A" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="F359F12BFB0262DF3FA212AF15F5EC23" bold="true">A</emphasis>
male left palp, prolateral view
<emphasis id="07374AB946C795086EBDB59F91ED9B73" bold="true">B</emphasis>
same, ventral view
<emphasis id="9405524BEC94EF6F213DEDEE46FD53F2" bold="true">C</emphasis>
same, retrolateral view
<emphasis id="589672DD1C38D57F3946BF79C0DE6F62" bold="true">D</emphasis>
same, dorsal view
<emphasis id="3A5A3D2CA97812C29C79220587F88755" bold="true">E</emphasis>
<abbrev id="ABBRID0EWJAE" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
, retrolateral view
<emphasis id="764B9407C95C0CBAA666CED1BC055855" bold="true">F</emphasis>
femur, prolateral view
<emphasis id="B5C8BC2FC31A5EDBEA9E56D843087E10" bold="true">G</emphasis>
epigyne, ventral view
<emphasis id="6721041F364097EC6F00DBD4F88771B2" bold="true">H</emphasis>
same, dorsal view. Abbreviations: E — embolus;
<abbrev id="ABBRID0EAKAE" xlink_title="femur apophysis">FA</abbrev>
— femoral apophysis;
<abbrev id="ABBRID0EEKAE" xlink_title="Prolateral tegular apophysis">PTA</abbrev>
— prolateral tegular apophysis;
<abbrev id="ABBRID0EIKAE" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
— retrolateral tibial apophysis;
<abbrev id="ABBRID0EMKAE" xlink_title="retroventral tibial apophysis">RvTA</abbrev>
— retroventral tibial apophysis;
<abbrev id="ABBRID0EQKAE" xlink_title="sperm duct">SD</abbrev>
— sperm duct;
<abbrev id="ABBRID0EUKAE" xlink_title="tegular sheet">TS</abbrev>
— tegular sheet; B — bursa;
<abbrev id="ABBRID0EYKAE" xlink_title="Copulatory ducts">CD</abbrev>
— copulatory duct;
<abbrev id="ABBRID0E3KAE" xlink_title="Copulatory openings">CO</abbrev>
— copulatory opening;
<abbrev id="ABBRID0EALAE" xlink_title="connecting tube">CT</abbrev>
— connecting tube;
<abbrev id="ABBRID0EELAE" xlink_title="Fertilization ducts">FD</abbrev>
— fertilization duct;
<abbrev id="ABBRID0EILAE" xlink_title="Glandular appendages">GA</abbrev>
— glandular appendage;
<abbrev id="ABBRID0EMLAE" xlink_title="median septum">MS</abbrev>
— median septum; S — spermathecae.
</paragraph>
</caption>
<paragraph id="68DB5AFDE51FE95DE2B0AC318EAE36DB">
Female (Fig.
<figureCitation id="C75F02D0F1A0DF1C3CB5D8AAE3529456" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Mutatus gelao sp. nov. A male holotype, dorsal view B same, ventral view C female paratype, dorsal view D same, ventral view." figureDoi="10.3897/zookeys.1226.137752.figure1" httpUri="https://binary.pensoft.net/fig/1243800">1 C, D</figureCitation>
). Total length 4.78, carapace 1.79 long, 1.64 wide; abdomen 3.08 long, 2.27 wide. Eye sizes and interdistances:
<abbrev id="ABBRID0E2LAE" xlink_title="anterior median eye">AME</abbrev>
0.11,
<abbrev id="ABBRID0E6LAE" xlink_title="anterior lateral eye">ALE</abbrev>
0.12,
<abbrev id="ABBRID0EDMAE" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EHMAE" xlink_title="posterior lateral eye">PLE</abbrev>
0.10;
<abbrev id="ABBRID0ELMAE" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0EPMAE" xlink_title="anterior median eye">AME</abbrev>
0.04,
<abbrev id="ABBRID0ETMAE" xlink_title="anterior median eye">AME</abbrev>
<abbrev id="ABBRID0EXMAE" xlink_title="anterior lateral eye">ALE</abbrev>
0.01,
<abbrev id="ABBRID0E2MAE" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0E6MAE" xlink_title="anterior lateral eye">ALE</abbrev>
0.24,
<abbrev id="ABBRID0EDNAE" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EHNAE" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0ELNAE" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EPNAE" xlink_title="posterior lateral eye">PLE</abbrev>
0.08,
<abbrev id="ABBRID0ETNAE" xlink_title="posterior lateral eye">PLE</abbrev>
<abbrev id="ABBRID0EXNAE" xlink_title="posterior lateral eye">PLE</abbrev>
0.39,
<abbrev id="ABBRID0E2NAE" xlink_title="anterior lateral eye">ALE</abbrev>
<abbrev id="ABBRID0E6NAE" xlink_title="posterior lateral eye">PLE</abbrev>
0.09.
<abbrev id="ABBRID0EDOAE" xlink_title="eye area width">EAW</abbrev>
0.54,
<abbrev id="ABBRID0EHOAE" xlink_title="cephalic region width">CRW</abbrev>
0.80,
<abbrev id="ABBRID0ELOAE" xlink_title="eye area width">EAW</abbrev>
/
<abbrev id="ABBRID0EPOAE" xlink_title="cephalic region width">CRW</abbrev>
0.68.
<abbrev id="ABBRID0ETOAE" xlink_title="cephalic region width">CRW</abbrev>
/
<abbrev id="ABBRID0EXOAE" xlink_title="carapace width">CW</abbrev>
0.49.
<abbrev id="ABBRID0E2OAE" xlink_title="median ocular area">MOA</abbrev>
0.28 long, anterior width 0.24, posterior width 0.24. Clypeal height 0.16,
<abbrev id="ABBRID0E6OAE" xlink_title="clypeal height">
<collectingCountry id="9C11CECB31C6708478E2A3C42470017A" name="Switzerland">CH</collectingCountry>
</abbrev>
/
<abbrev id="ABBRID0EDPAE" xlink_title="anterior median eye">AME</abbrev>
1.36. Labium 0.23 long, 0.25 wide. Sternum 1.07 long, 0.97 wide. Other characters as for male, except dorsal scutum absent, color darker and body slightly larger.
</paragraph>
<paragraph id="254B8AD69F926CBD35D29BD20400EFA9">
Epigyne (Fig.
<figureCitation id="A8BA187C13C91E1C3DA65746BF6EEF32" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Copulatory organs of Mutatus gelao sp. nov. A male left palp, prolateral view B same, ventral view C same, retrolateral view D same, dorsal view E RTA, retrolateral view F femur, prolateral view G epigyne, ventral view H same, dorsal view. Abbreviations: E — embolus; FA — femoral apophysis; PTA — prolateral tegular apophysis; RTA — retrolateral tibial apophysis; RvTA — retroventral tibial apophysis; SD — sperm duct; TS — tegular sheet; B — bursa; CD — copulatory duct; CO — copulatory opening; CT — connecting tube; FD — fertilization duct; GA — glandular appendage; MS — median septum; S — spermathecae." figureDoi="10.3897/zookeys.1226.137752.figure2" httpUri="https://binary.pensoft.net/fig/1243801">2 G, H</figureCitation>
). Epigynal plate slightly sclerotized, with a pair of atria. Median septum (
<abbrev id="ABBRID0ENPAE" xlink_title="median septum">MS</abbrev>
) dumbbell-shaped. Copulatory openings (
<abbrev id="ABBRID0ERPAE" xlink_title="Copulatory openings">CO</abbrev>
) small, located at anterior of atrium. Copulatory ducts (
<abbrev id="ABBRID0EVPAE" xlink_title="Copulatory ducts">CD</abbrev>
) slightly curved, X-shaped, longer and thinner than connecting tubes (
<abbrev id="ABBRID0EZPAE" xlink_title="connecting tube">CT</abbrev>
). Bursae (B) nearly balloon-shaped, translucent. Glandular appendages (
<abbrev id="ABBRID0E4PAE" xlink_title="Glandular appendages">GA</abbrev>
) mastoid-shaped. Spermathecae (S) clavate, separated by septum. Fertilization ducts (
<abbrev id="ABBRID0EBQAE" xlink_title="Fertilization ducts">FD</abbrev>
) located at median margin of spermathecae.
</paragraph>
<paragraph id="2404147460E41DD2E4E6242690E9B31C">Measurement of legs:</paragraph>
<paragraph id="0875BAC1C765C4CBEB4113DF286E7249">
<table id="TID0EO1AE" rules="all">
<tbody id="77211937ADE94EE09B989200513A2521">
<table-wrap id="T1" orientation="portrait" position="float">
<tr id="3604954D0B64B3FCA9E752720732413D">
<th id="533641C15E8D2A03B668246BA549DC38" colspan="1" rowspan="1">male / female</th>
<th id="375419AB3619F04470E816111246406C" colspan="1" rowspan="1">Fe</th>
<th id="1248A5C694CA51ECAEF2E16DA81DE888" colspan="1" rowspan="1">Pa</th>
<th id="A82C1E2BE0581A6CD406FA91757DC2E3" colspan="1" rowspan="1">Ti</th>
<th id="E82CFA6B76D9175371ED334D562769B8" colspan="1" rowspan="1">Me</th>
<th id="27E63B455ED32160B75E940B1E52FFB8" colspan="1" rowspan="1">Ta</th>
<th id="2B3CAD7EDFA907CB2432347552F8D5C3" colspan="1" rowspan="1">Total</th>
</tr>
<tr id="83F1FC36EB82F875C62F217EABFA5B6C">
<td id="E2F28BB81F7E669C81C5343FCD7C2AA2" colspan="1" rowspan="1">Leg I</td>
<td id="EBC363890AC20DB46403C41653D60247" colspan="1" rowspan="1">1.67 / 1.71</td>
<td id="C5C2ADDF025D7F5D5A8A8A52F72887DB" colspan="1" rowspan="1">0.60 / 0.56</td>
<td id="0D25AB6D52D9CF04D826B9FFD4108D50" colspan="1" rowspan="1">1.80 / 1.65</td>
<td id="62BD10299B80A640A38124BE7F84085B" colspan="1" rowspan="1">1.58 / 1.51</td>
<td id="87F25CEB13802CF1DA1604C36D917A79" colspan="1" rowspan="1">0.89 / 0.85</td>
<td id="3E85090AB210BAA7FACDDC7BE002271C" colspan="1" rowspan="1">6.54 / 6.27</td>
</tr>
<tr id="2BDC25439485469C530E6B9B1F2FA261">
<td id="C4C24A27BFCCF4E29466C1474F41FDC2" colspan="1" rowspan="1">Leg II</td>
<td id="49F5B57E2622F0F604C99A086F3706DC" colspan="1" rowspan="1">1.54 / 1.63</td>
<td id="8251A7006F83EF640FF058C77F53908D" colspan="1" rowspan="1">0.47 / 0.61</td>
<td id="14417D572636EBF0E5132CB91AE0C0FE" colspan="1" rowspan="1">1.44 / 1.49</td>
<td id="8825D2721C6072735E333B397A104D60" colspan="1" rowspan="1">1.20 / 1.38</td>
<td id="ECE46458ABA6E2ACA4252A445904DDC5" colspan="1" rowspan="1">0.86 / 0.93</td>
<td id="B14D234F33C37E7A6DA79BF347C7BF8A" colspan="1" rowspan="1">5.51 / 6.04</td>
</tr>
<tr id="B6914E5D3EEA99D2E84FD8DEFBAA1FB2">
<td id="C48598D6A11B4D035F5C1DD754632AE3" colspan="1" rowspan="1">Leg III</td>
<td id="95E4B8059A7B2A91922E122ED81A7AF2" colspan="1" rowspan="1">1.31 / 1.43</td>
<td id="AAADD988ED5AF7EBF0EFFC18E441CA53" colspan="1" rowspan="1">0.51 / 0.54</td>
<td id="88BD0C5B5955354026A5AA82636F5BB0" colspan="1" rowspan="1">1.07 / 1.07</td>
<td id="FE38F2DDB4601B7541990BA2DEF1799F" colspan="1" rowspan="1">1.22 / 1.33</td>
<td id="CC472D3181811CC9DF9E1AA6589B7C5E" colspan="1" rowspan="1">0.70 / 0.77</td>
<td id="156CC2335261A015FED2CC44C8BF004B" colspan="1" rowspan="1">4.81 / 5.14</td>
</tr>
<tr id="992090B5452BE02F67F808F513001446">
<td id="8D9BE266DCCAB9541ADCAC2A25C0DC17" colspan="1" rowspan="1">Leg IV</td>
<td id="1C70519FAB423110CE4F42AC82F21890" colspan="1" rowspan="1">1.96 / 2.10</td>
<td id="FBEA74E8259750ADFA1DC1C37B9E961D" colspan="1" rowspan="1">0.58 / 0.65</td>
<td id="C4B2204B68C60D8919A1CB76686963B4" colspan="1" rowspan="1">1.79 / 1.73</td>
<td id="C58F92D03E5A4CE542794A1E418DA8EA" colspan="1" rowspan="1">2.01 / 2.06</td>
<td id="5F6ED2687A3FFD811F818751CDD4D23F" colspan="1" rowspan="1">1.02 / 0.97</td>
<td id="4B820EC4DABDAC12E4E4ED2E97E559FF" colspan="1" rowspan="1">7.36 / 7.51</td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
<paragraph id="AB49F75DD9E3F0BAF87174080CDB9027">Spination of legs:</paragraph>
<paragraph id="29E7401FEADE75FD6CF4AE7BBD8E3C71">
<table id="TID0EOAAG" rules="all">
<tbody id="7852252A220B12D873E1D729CE1CC621">
<table-wrap id="T2" orientation="portrait" position="float">
<tr id="B283D5CF40A677D277028F08FC965C71">
<th id="133081963CC9010B6C86A4D4B2695B22" colspan="2" isEmpty="true" rowspan="1">-</th>
<th id="D3C9427317C72EDFC242B58356F21988" colspan="1" rowspan="1">Fe</th>
<th id="11CD8A3BBE92AF24680BC0230A9510A5" colspan="1" rowspan="1">Pa</th>
<th id="ADBA7D2769CD8E4914F3A580E33A876A" colspan="1" rowspan="1">Ti</th>
<th id="6E7F7AFFA887105E7588BBFD6DF7626D" colspan="1" rowspan="1">Me</th>
<th id="506242CCD1402188862242DED4C8ECB8" colspan="1" rowspan="1">Ta</th>
</tr>
<tr id="54363C74A424996DD2C0F895542E4FFA">
<td id="C3985A489D8F23D1B6AC3F7AF7F19DA1" colspan="1" rowspan="4">male</td>
<td id="A2852E699391F982E17A6C9EDDF71C3C" colspan="1" rowspan="1">Leg I</td>
<td id="74C13935B5ECC11E45EC3F9395A835CD" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EHVAE" xlink_title="prolateral">pl</abbrev>
4
</td>
<td id="89168BB7F83DDC4D69D78BB181D34F89" colspan="1" rowspan="1"></td>
<td id="669E41D50DD8C01937F112B233A79D1A" colspan="1" rowspan="1">
<abbrev id="ABBRID0ESVAE" xlink_title="proventral">pv</abbrev>
7
<abbrev id="ABBRID0EWVAE" xlink_title="retroventral">rv</abbrev>
7
</td>
<td id="49CD2BD619298702594297430ABCA978" colspan="1" rowspan="1">
<abbrev id="ABBRID0E5VAE" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0ECWAE" xlink_title="retroventral">rv</abbrev>
4
</td>
<td id="A382B8DB4A1873A91E66BE9569353205" colspan="1" rowspan="1"></td>
</tr>
<tr id="30956BB97335D6610418857D156FE34D">
<td id="ADE56038101042176AEAA1A1F6A1BDE9" colspan="1" rowspan="1">Leg II</td>
<td id="9AEAD6771B0446E4ED670BDFFF17C62F" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0ERWAE" xlink_title="prolateral">pl</abbrev>
0
</td>
<td id="E5263D882D252072712931D1B58630DC" colspan="1" rowspan="1"></td>
<td id="8539E924BF19E0B295DFC6CC4AD6285A" colspan="1" rowspan="1">
<abbrev id="ABBRID0E3WAE" xlink_title="proventral">pv</abbrev>
7
<abbrev id="ABBRID0EAXAE" xlink_title="retroventral">rv</abbrev>
5
</td>
<td id="B060C92EE8284FB6650C074464333ABC" colspan="1" rowspan="1">
<abbrev id="ABBRID0EIXAE" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0EMXAE" xlink_title="retroventral">rv</abbrev>
4
</td>
<td id="FA37B5CA8E5429105E446D940348FEC8" colspan="1" rowspan="1"></td>
</tr>
<tr id="0EC8E55E19CDF7B1FC8D1B8506372CFC">
<td id="D32451F500FCC098FE4EE29FAEE79B42" colspan="1" rowspan="1">Leg III</td>
<td id="8DD714AE993B33BC5862AE64F281AF68" colspan="1" rowspan="1">d 1</td>
<td id="8AF343B79FDD4D6485916D7029719151" colspan="1" rowspan="1"></td>
<td id="1C2DACAE0174833698B090A87581D2D8" colspan="1" rowspan="1"></td>
<td id="C21369C236D8846E462910149D7C07E4" colspan="1" rowspan="1"></td>
<td id="6D7F602582E23A05421B8C21606613E0" colspan="1" rowspan="1"></td>
</tr>
<tr id="B6FDD7FCD11210F1171714D00917BEDB">
<td id="CCF88A3E6DA3E38A304E879F7EF1AC58" colspan="1" rowspan="1">Leg IV</td>
<td id="94EC0364E124D60E3B49CAC651409736" colspan="1" rowspan="1">d 1</td>
<td id="12BBA8CB48AA21D1297D88AADBE4F14F" colspan="1" rowspan="1"></td>
<td id="66A058EB6EAD6543133FE2DE21CD79D0" colspan="1" rowspan="1"></td>
<td id="1D446D826FB1921937223C3CC581313E" colspan="1" rowspan="1"></td>
<td id="8637E8DAB3B78E94B6AB9C04A3160B2A" colspan="1" rowspan="1"></td>
</tr>
<tr id="9A22EC23C82F5DE41C909F44834E79F2">
<td id="57054DEDA382659421BF38B08F59F0E1" colspan="1" rowspan="4">female</td>
<td id="A956E3EAC471A0701760E45EF8DCF961" colspan="1" rowspan="1">Leg I</td>
<td id="907D9EF3414753B6E79CB140AAC8A486" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EEZAE" xlink_title="prolateral">pl</abbrev>
5
</td>
<td id="7926E366B18037A2DBEF07DF8CA8A217" colspan="1" rowspan="1"></td>
<td id="41C42CBD93BFC082D05C9E29B0049C12" colspan="1" rowspan="1">
<abbrev id="ABBRID0EPZAE" xlink_title="proventral">pv</abbrev>
7
<abbrev id="ABBRID0ETZAE" xlink_title="retroventral">rv</abbrev>
7
</td>
<td id="A325BB813EAE87774DA0E726B980E505" colspan="1" rowspan="1">
<abbrev id="ABBRID0E2ZAE" xlink_title="proventral">pv</abbrev>
5
<abbrev id="ABBRID0E6ZAE" xlink_title="retroventral">rv</abbrev>
5
</td>
<td id="CC59C571F5FDC183A42FD9BA5BD72182" colspan="1" rowspan="1"></td>
</tr>
<tr id="C910E2EFA134A8E6F943BCC0DBBAC2B2">
<td id="F9BC4EE9D4682F35FD663176D0709513" colspan="1" rowspan="1">Leg II</td>
<td id="A4225FA19526D189D293F0FC2EDE7D6B" colspan="1" rowspan="1">
d 1
<abbrev id="ABBRID0EO1AE" xlink_title="prolateral">pl</abbrev>
2
</td>
<td id="D0F440F160F6C0FDBE915EE411BBDD9F" colspan="1" rowspan="1"></td>
<td id="023CBA3FFCDE1AE61175F8B118B7D467" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZ1AE" xlink_title="proventral">pv</abbrev>
7
<abbrev id="ABBRID0E41AE" xlink_title="retroventral">rv</abbrev>
7
</td>
<td id="0374C72C7CBB013A095A7B2F3EF0BCFC" colspan="1" rowspan="1">
<abbrev id="ABBRID0EF2AE" xlink_title="proventral">pv</abbrev>
4
<abbrev id="ABBRID0EJ2AE" xlink_title="retroventral">rv</abbrev>
3
</td>
<td id="40834CCBBE2AF38B6F796BB47D932EC5" colspan="1" rowspan="1"></td>
</tr>
<tr id="930924F7AF8E9DFF1EE5D64A4FE4C551">
<td id="83A39DE47D8810B6FF543A766292F6CF" colspan="1" rowspan="1">Leg III</td>
<td id="9935B680820D249EB632C712D5553ACD" colspan="1" rowspan="1">d 1</td>
<td id="2C16CD016429891431E46257D40F9260" colspan="1" rowspan="1"></td>
<td id="0E2680D17DD860DD480C910190C2206F" colspan="1" rowspan="1"></td>
<td id="2F30FBF8928E6E3C83FD15ED0A390EEF" colspan="1" rowspan="1"></td>
<td id="9D49292EFB569171E5CE47A326FD397A" colspan="1" rowspan="1"></td>
</tr>
<tr id="56089CC263EDEBCD7392A9C6F7DB849C">
<td id="D8DB477E81313F22F50DE782C30DF281" colspan="1" rowspan="1">Leg IV</td>
<td id="B53177150AB5F42E2CD128C90A067F82" colspan="1" rowspan="1">d 1</td>
<td id="A3D415E3DE49AA584F0908297CC9547C" colspan="1" rowspan="1"></td>
<td id="62103AD7E116A5E183083A709F56198C" colspan="1" rowspan="1"></td>
<td id="AA14E2B4F8CC78E33F0F5BD98289B6ED" colspan="1" rowspan="1"></td>
<td id="8E54FDDFC7849FA7B7703063BFB0B2A0" colspan="1" rowspan="1"></td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EW3AE" type="distribution">
<paragraph id="78CF9135DC5CF10024AFFCBA7A214B2F">
<heading id="DB7E75F2F1D8715E52E147060434226E" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="EE1F055C14C7F87538837277A7725261">
Known only from the
<typeStatus id="9CFCC35324FC4412F4C257F32F587579">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="1BCC0D593AB7761AD4286032DFFC3D01" ID-DOI="10.3897/zookeys.1226.140204" ID-ZooBank="AC08832F-748C-44E2-ADFD-514F258779C8" ID-publisher-id="140204" URI-arpha="12D895A7-C86E-520B-9D0B-BCC564726837" URI-zoobank="http://zoobank.org/AC08832F-748C-44E2-ADFD-514F258779C8" article-type="research-article" checkinTime="1738866271465" checkinUser="pensoft" docAuthor="Hedin, Marshal, Ramírez, Martín J. &amp; Monjaraz-Ruedas, Rodrigo" docDate="2025" docId="D143EFD9B16158E1A385DBA7ACE1A587" docLanguage="en" docName="ZooKeys 1226: 47-75" docOrigin="ZooKeys 1226" docSource="https://zookeys.pensoft.net/article/140204/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Siskiyu Hedin, Ramírez &amp; Monjaraz-Ruedas, 2025, gen. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="75" masterDocId="12D895A7C86E520B9D0BBCC564726837" masterDocTitle="Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province" masterLastPageNumber="75" masterPageNumber="47" pageNumber="47" updateTime="1738866794866" updateUser="ExternalLinkService">
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<mods:title id="C082E900E0875E3BBCB553D59BC100FB">Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province</mods:title>
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<mods:namePart id="C21380646B2F90731DD36DEE5258E0AA">Hedin, Marshal</mods:namePart>
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<mods:affiliation id="D492D8D5A7A059F20081658022064CE4">Department of Biology, San Diego State University, San Diego, California 92182 4614, USA</mods:affiliation>
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<mods:namePart id="40C2599762834791710F7A52B8394A41">Ramírez, Martín J.</mods:namePart>
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<mods:affiliation id="15B13444B95FCF8A3878BFB3C103FCA7">Division of Arachnology, Museo Argentino de Ciencias Naturales - CONICET, Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="C2BFCA6779E0A4AE300A8730DA04116F">Monjaraz-Ruedas, Rodrigo</mods:namePart>
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<mods:affiliation id="F4473477D1403BF85DA6DF9BFAA3E4C7">Department of Biology, San Diego State University, San Diego, California 92182 4614, USA &amp; Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, California 90007, USA</mods:affiliation>
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<mods:date id="9D4807064E0DC8A1126B7D1E54A71884">2025</mods:date>
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<mods:number id="5C894D5D205E591524416CDD13420AE4">2025-02-06</mods:number>
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<subSubSection id="4904D299885B4C5B84D5C3FC02B31E52" type="nomenclature">
<paragraph id="67AEE6E2885CE04BC868785C500EB650">
<heading id="1A662443BBF2C6F2C1497355E90AF0BC" reason="label">Genus</heading>
<taxonomicName id="DCD30329CD4F666D7D3FF3018E75DF25" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Siskiyu" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B8EE785A4F6B3D0DD918737EFDF7B6F6" italics="true">Siskiyu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78A1AF370FBF94B57B5BBC16FD8EA94" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="A42A679B23A09387E54F2BE6623BF344" type="description">
<paragraph id="7211EF8915DBA07B65BFC8A5173ACB0B">
<figureCitation id="E684A6B28ED398F3890FE24F7FE086F7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Images of live cybaeid spiders. A Neocryphoeca beattyi ♀, Pinal County, AZ, RWM 22 _ 011 B Blabomma sp ♀, San Diego County, CA, MCH 22 _ 020 C Siskiyu armilla sp. nov. immature ♀, Siskiyou County CA, MCH 24 _ 057; patellar fracture line labeled D Siskiyu armilla sp. nov. immature ♂, Siskiyou County CA, MCH 24 _ 057; with silk webbing presumed to belong to the spider E Cybaeota shastae ♀ Chamberlin &amp; Ivie, 1937, East Rosebud Creek, MT F Cybaeus somesbar ♀ Bennett, 2009, Shasta County, CA, MCH 24 _ 012. All images taken by M Hedin, except for Neocryphoeca (RW Mendez)." figureDoi="10.3897/zookeys.1226.140204.figure1" httpUri="https://binary.pensoft.net/fig/1243090">Figs 1</figureCitation>
,
<figureCitation id="74CCD0756142C8FA7910ADBD19B55170" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siskiyu armilla sp. nov. holotype ♂ (SDSU _ G 1090) palp A ventral view B lateral view. Elongate cymbium, tegular process, embolus, conductor processes, patellar apophysis, and RTA basal and distal processes labeled." figureDoi="10.3897/zookeys.1226.140204.figure6" httpUri="https://binary.pensoft.net/fig/1243095">6</figureCitation>
,
<figureCitation id="E1837AC4AEA871C0A8FB30AEFBE89DB1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siskiyu armilla sp. nov. paratype ♀ (SDSU _ G 1086 A) A dorsal view B ventral view C leg IV patellar fracture line D ventral epigynum E dorsal epigynum. Patellar fracture lines and epigynal dorsal ridge, copulatory openings (co), copulatory ducts (cd), bilobed spermathecae and fertilization ducts (fd) labeled." figureDoi="10.3897/zookeys.1226.140204.figure7" httpUri="https://binary.pensoft.net/fig/1243096">7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEPBG" type="etymology">
<paragraph id="A7851F2C60458CE9797DFADB8FE262A8">
<heading id="A3D6F087EDFCD224BDF9D8A2F87C1DFA" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A4CF95BF69C35F7BCC6BDBD440FFD285">A modification of siskiyou, from the Klamath-Siskiyou Mountains that encompass much of the known distribution of this genus. The etymology of siskiyou, possibly indigenous, remains uncertain. Grammatical gender treated as feminine.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJPBG" type="diagnosis">
<paragraph id="F41D3A16462DEF8A4CCA15931B996EAA">
<heading id="5B8D36C21B0C718045ACAE308116283E" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="F9E0B1E8FC80A29EC1E11C9AE5BA4DAB">
With a conspicuous fracture line near the base of patellae I
<collectionCode id="E783FD1C18A31CC7C38D36FD0D673E91">IV</collectionCode>
, like close relatives
<taxonomicName id="43BDCA2FA3F0BCBBD53691B7FB95A395" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Calymmaria" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="439E46641A5CD682E8AB20CD73E71AF6" italics="true">Calymmaria</emphasis>
</taxonomicName>
and
<taxonomicName id="3D026BE4C77A6DE058EABD05DCBDF0A6" authorityName="Roth" authorityYear="1981" class="Arachnida" family="Cybaeidae" genus="Willisus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="0DB36361E807336CA3B6F3080953BE1D" italics="true">Willisus</emphasis>
</taxonomicName>
(
<bibRefCitation id="AB92507EDC1BE0F6E700ADC558669856" author="Roth" firstAuthor="Roth" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="101-105" refId="B54" refString="Roth VD (1981) A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy. Bulletin of the American Museum of Natural History 170: 101105. https://wsc.nmbe.ch/species/12952" title="A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy." url="https://wsc.nmbe.ch/species/12952" volume="170" year="1981">Roth 1981</bibRefCitation>
: fig. 1;
<bibRefCitation id="FE7C240FD49A84E3EEA33857BA12F268" DOI="10.1636/H02-12" author="Heiss and Draney" firstAuthor="Heiss" issue="3" journalOrPublisher="The Journal of Arachnology" pagination="457-525" refId="B28" refString="Heiss JS, Draney ML (2004) Revision of the Nearctic spider genus Calymmaria (Araneae, Hahniidae). The Journal of Arachnology 32 (3): 457525. https://doi.org/10.1636/H02-12" title="Revision of the Nearctic spider genus Calymmaria (Araneae, Hahniidae)." volume="32" year="2004">Heiss and Draney 2004</bibRefCitation>
: fig. 3). The male palp of
<taxonomicName id="58D2DD2BB1232666FB4E49B35A8EDAB4" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Siskiyu" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FA88C539550388519746037015D7C20D" italics="true">Siskiyu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8CB7C3A206DADCA8A92EAC159B0C108" rank="genus">gen. nov.</taxonomicNameLabel>
is diagnosed from
<taxonomicName id="51F3E2073B7A3EE7A080DD0A742F5716" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Calymmaria" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="B056FEC33759D5080ECE0CA5A6D68D77" italics="true">Calymmaria</emphasis>
</taxonomicName>
in possessing a dorsal tegular process, a thinner embolus with the distal tip projecting ventrally, associated with a trilobed conductor (Fig.
<figureCitation id="9B34094E14BE03C001F27D40D2D11BA3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siskiyu armilla sp. nov. holotype ♂ (SDSU _ G 1090) palp A ventral view B lateral view. Elongate cymbium, tegular process, embolus, conductor processes, patellar apophysis, and RTA basal and distal processes labeled." figureDoi="10.3897/zookeys.1226.140204.figure6" httpUri="https://binary.pensoft.net/fig/1243095">6 A, B</figureCitation>
).
<taxonomicName id="AFA06409509D9D0D45CE40066D4B4773" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Calymmaria" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="4A371ACDAB377B198D5108D2016DEDAD" italics="true">Calymmaria</emphasis>
</taxonomicName>
palps lack a tegular process, have generally thicker (or sometimes forked) emboli, with conductors that lack dorsal processes (
<bibRefCitation id="0C2863A71C7BED0CDFEFC006928EE1D8" DOI="10.1636/H02-12" author="Heiss and Draney" firstAuthor="Heiss" issue="3" journalOrPublisher="The Journal of Arachnology" pagination="457-525" refId="B28" refString="Heiss JS, Draney ML (2004) Revision of the Nearctic spider genus Calymmaria (Araneae, Hahniidae). The Journal of Arachnology 32 (3): 457525. https://doi.org/10.1636/H02-12" title="Revision of the Nearctic spider genus Calymmaria (Araneae, Hahniidae)." volume="32" year="2004">Heiss and Draney 2004</bibRefCitation>
: figs 6, 7).
<taxonomicName id="A858B56B9163CD03A3A09769A9200530" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Siskiyu" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AE139DB64296768578BE09B07A6A0C35" italics="true">Siskiyu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D19ED3B19B4305D169369DF61038937F" rank="genus">gen. nov.</taxonomicNameLabel>
male palps are most like those of
<taxonomicName id="7F5D4076055E1633A399FE8186E8FE6A" authorityName="Roth" authorityYear="1981" class="Arachnida" family="Cybaeidae" genus="Willisus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="E580A5268E205BAB108D57AC52F9F6A2" italics="true">Willisus</emphasis>
</taxonomicName>
but differ in the condition of the dorsal tegular process (more robust and toothed in
<taxonomicName id="681587435A5FB8C86974784DA6050BE2" authorityName="Roth" authorityYear="1981" class="Arachnida" family="Cybaeidae" genus="Willisus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="B7EFB39FAB80073AF5FD87E5D943EE86" italics="true">Willisus</emphasis>
</taxonomicName>
), shape of the embolus (sinuate in
<taxonomicName id="1E30F2F0415087E9B933735F344CCD60" authorityName="Roth" authorityYear="1981" class="Arachnida" family="Cybaeidae" genus="Willisus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="003B16E05C0C55CF8453E9C814B64961" italics="true">Willisus</emphasis>
</taxonomicName>
), and shapes of both the conductor and
<abbrev id="ABBRID0E5RBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
lobes (
<bibRefCitation id="5FC521A8FBA684E9090B8BFA1D68AD89" author="Roth" firstAuthor="Roth" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="101-105" refId="B54" refString="Roth VD (1981) A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy. Bulletin of the American Museum of Natural History 170: 101105. https://wsc.nmbe.ch/species/12952" title="A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy." url="https://wsc.nmbe.ch/species/12952" volume="170" year="1981">Roth 1981</bibRefCitation>
: figs 35).
<taxonomicName id="A9F68AAD374AE3C8CFB1561980CC796A" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Siskiyu" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="ED40956146C6A8D1178EB660D0C1CEB3" italics="true">Siskiyu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88E9FFA9AAF7EC2C83982A2E12FB4C85" rank="genus">gen. nov.</taxonomicNameLabel>
female genitalia differ from
<taxonomicName id="390EC7E5166A015D5FC8B9A1F7C5E2AF" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Calymmaria" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="8AD5244B11B95EA04D289AD129529830" italics="true">Calymmaria</emphasis>
</taxonomicName>
in possessing subequal, bilobed spherical spermathecae (Fig.
<figureCitation id="FC33CBD84DC42F33D06265DAE09A540C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siskiyu armilla sp. nov. paratype ♀ (SDSU _ G 1086 A) A dorsal view B ventral view C leg IV patellar fracture line D ventral epigynum E dorsal epigynum. Patellar fracture lines and epigynal dorsal ridge, copulatory openings (co), copulatory ducts (cd), bilobed spermathecae and fertilization ducts (fd) labeled." figureDoi="10.3897/zookeys.1226.140204.figure7" httpUri="https://binary.pensoft.net/fig/1243096">7 D, E</figureCitation>
), in contrast to the simple paired spherical spermathecae of most
<taxonomicName id="6D85866A5657D9886A9E47DA31656A99" authorityName="Chamberlin &amp; Ivie" authorityYear="1937" class="Arachnida" family="Cybaeidae" genus="Calymmaria" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="5E024F338D65CF40AB195C5F7C3432E8" italics="true">Calymmaria</emphasis>
</taxonomicName>
(
<bibRefCitation id="571857523C84ADF22A25B943659712A4" DOI="10.1636/H02-12" author="Heiss and Draney" firstAuthor="Heiss" issue="3" journalOrPublisher="The Journal of Arachnology" pagination="457-525" refId="B28" refString="Heiss JS, Draney ML (2004) Revision of the Nearctic spider genus Calymmaria (Araneae, Hahniidae). The Journal of Arachnology 32 (3): 457525. https://doi.org/10.1636/H02-12" title="Revision of the Nearctic spider genus Calymmaria (Araneae, Hahniidae)." volume="32" year="2004">Heiss and Draney 2004</bibRefCitation>
: figs 4, 5), and different from the distinctive
<taxonomicName id="E83E604B4885F3BE001030D0B7A52E12" authorityName="Heiss &amp; Draney" authorityYear="2004" class="Arachnida" family="Cybaeidae" genus="Calymmaria" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="alleni">
<emphasis id="83987FAB4C10DF103A82417DE2725549" italics="true">C. alleni</emphasis>
</taxonomicName>
epigynum;
<bibRefCitation id="11AC50543D93A6699DBD717642ADF804" DOI="10.1636/H02-12" author="Heiss and Draney" firstAuthor="Heiss" issue="3" journalOrPublisher="The Journal of Arachnology" pagination="457-525" refId="B28" refString="Heiss JS, Draney ML (2004) Revision of the Nearctic spider genus Calymmaria (Araneae, Hahniidae). The Journal of Arachnology 32 (3): 457525. https://doi.org/10.1636/H02-12" title="Revision of the Nearctic spider genus Calymmaria (Araneae, Hahniidae)." volume="32" year="2004">Heiss and Draney 2004</bibRefCitation>
: figs 8, 9).
<taxonomicName id="C13085839BCF951372A7CAF713559429" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Siskiyu" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="81F10EE5DFA44F6389E29C1597ED5631" italics="true">Siskiyu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AB7F1F69707D6B7906F9E3BBF6F67E4" rank="genus">gen. nov.</taxonomicNameLabel>
female genitalia are distinguished from those of
<taxonomicName id="D12C4DBF7D177E98FFA339293FFFD624" authorityName="Roth" authorityYear="1981" class="Arachnida" family="Cybaeidae" genus="Willisus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="54D87FD091CDF3509D05D7871D7F3262" italics="true">Willisus</emphasis>
</taxonomicName>
which include short copulatory ducts and oblique kidney-bean shaped spermathecae (
<bibRefCitation id="D773E9CF901FED7468DB265C15BEA7E3" author="Roth" firstAuthor="Roth" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="101-105" refId="B54" refString="Roth VD (1981) A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy. Bulletin of the American Museum of Natural History 170: 101105. https://wsc.nmbe.ch/species/12952" title="A new genus of spider (Agelenidae) from California exhibiting a third type of leg autospasy." url="https://wsc.nmbe.ch/species/12952" volume="170" year="1981">Roth 1981</bibRefCitation>
: figs 6, 7).
</paragraph>
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<paragraph id="3231D85749276DD5DD062712690EEFD6">
<label id="56FB632BE6DB98A5016FC17D394CCFCE">Figure 6.</label>
</paragraph>
<paragraph id="7ABD2C0313AC611EF9411E8E415E8C77">
<taxonomicName id="98E8F3D7678907635DFB6A65810F6C94" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Siskiyu" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="armilla" status="sp. nov.">
<emphasis id="7F6CC3C4218D430B339BB301D26CCFDB" italics="true">Siskiyu armilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67AA133521BB6B7E3C63D7B272DCFA7E" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♂ (SDSU _ G 1090) palp
<emphasis id="D8068315F03BF8FC6483E8D9277DA0D1" bold="true">A</emphasis>
ventral view
<emphasis id="397A4A12A530C226F70C4166F7A0E78B" bold="true">B</emphasis>
lateral view. Elongate cymbium, tegular process, embolus, conductor processes, patellar apophysis, and
<abbrev id="ABBRID0E2UBG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
basal and distal processes labeled.
</paragraph>
</caption>
<caption id="B1ED25FE7115B0E5489A9CB7D3F61AD6" ID-DOI="10.3897/zookeys.1226.140204.figure7" ID-arpha="4B6B7A22-A93C-52FC-94D6-0A3D5D6AA83F" httpUri="https://binary.pensoft.net/fig/1243096" startId="F7">
<paragraph id="DD5870C3359D3BAEB12AF6C56FF8C9EE">
<label id="CE46692666D7D7793B3C43F2FA0926DF">Figure 7.</label>
</paragraph>
<paragraph id="DC632D3B56B5DA35D15EB03BA98AC158">
<taxonomicName id="28A53F4FD57FD7B23ADFEB533F422EB8" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Siskiyu" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="armilla" status="sp. nov.">
<emphasis id="50B6A172007FD9784235F1FBDEC4911F" italics="true">Siskiyu armilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="033ACFA17731CCA1C144C1A3D04DAFD7" rank="species">sp. nov.</taxonomicNameLabel>
paratype ♀ (SDSU _ G 1086 A)
<emphasis id="3F04A8E4FD8CB1C1193FE658065C6B8A" bold="true">A</emphasis>
dorsal view
<emphasis id="E5AE415634BB282FF24CADE1C65E3397" bold="true">B</emphasis>
ventral view
<emphasis id="A659CB9FBC8140DBCBD5949417A59CA6" bold="true">C</emphasis>
leg IV patellar fracture line
<emphasis id="0755FB8A5129141203A0583A57812DB4" bold="true">D</emphasis>
ventral epigynum
<emphasis id="58EFC995C740373E40A9BE8A2B5D2C26" bold="true">E</emphasis>
dorsal epigynum. Patellar fracture lines and epigynal dorsal ridge, copulatory openings (co), copulatory ducts (cd), bilobed spermathecae and fertilization ducts (fd) labeled.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EGWBG" type="description">
<paragraph id="A21FD579A82D784F2F54507EAB3129C6">
<heading id="05753D3ADAE9C5A60AE7DAD5AF8ED6EF" reason="title">Description.</heading>
</paragraph>
<paragraph id="4DF42596CBEEA83F864FC2FBBB241CD4">
Adults
<quantity id="09B8C4C0756C016F517ECE319D9C7DDB" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
in length. Dorsal carapace mostly bare and lacking pigmentation, longitudinal fovea ~ 2 × length of PME diameter. Eight eyes, anterior eye row slightly recurved, posterior eye row slightly procurved. AMEs smallest, ~ ¼ diameter of ALEs, separated by diameter. Lateral eyes approximately contiguous, approximately equal in size; PMEs ½ diameter of PLEs, slightly closer to PLEs. Clypeus as high as ALEs. Chelicerae straight, longer than width of cephalic region, lateral boss present; anterior margin with three teeth, posterior margin with five or six teeth. Labium nearly square, narrowing slightly anteriorly. Endites slightly convergent, ~ 2 × as long as wide, with serrula. Sternum approximately heart-shaped but with truncated anterior edge, narrowed posteriorly, ending at posterior edge of hind coxae; sparsely covered with fine hairs. Trochanters without notches. Legs with fine hairs, mostly unmarked. Leg formula 1, 4, 2, 3. Patella-tibia I 1.62 × longer than carapace. Femurs lacking macrosetae; patellar, tibial, and metatarsal macrosetae restricted to posterior legs. Leg patellae with conspicuous cleavage plane basally at ⅕ of patella length (Fig.
<figureCitation id="CB3958A26DBDAA63AC60BA1E55E9C445" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siskiyu armilla sp. nov. paratype ♀ (SDSU _ G 1086 A) A dorsal view B ventral view C leg IV patellar fracture line D ventral epigynum E dorsal epigynum. Patellar fracture lines and epigynal dorsal ridge, copulatory openings (co), copulatory ducts (cd), bilobed spermathecae and fertilization ducts (fd) labeled." figureDoi="10.3897/zookeys.1226.140204.figure7" httpUri="https://binary.pensoft.net/fig/1243096">7 C</figureCitation>
). Tarsi and metatarsi with dorsal rows of 36 long trichobothria. Paired tarsal claws with ten teeth, median claw very small, semicircular hairs between claws not obvious. Abdomen with scattered simple hairs, more ventrally than dorsally. Colulus inconspicuous, with six setae. Anterior spinnerets are stout, conical, separated by basal width, distal segment hemispherical. Posterior spinnerets longer than anterior with wedge-shaped distal segments ~ 1.3 × longer than length of basal segment, pointed medially. Median spinnerets lying directly posterior and approximately equal in height to anterior spinnerets, closely adjacent.
</paragraph>
<paragraph id="2479C27E769064619D4771707A62AE22">Female palp with tibial and tarsal macrosetae, clothed with fine hairs. Female genitalia with paired lateral openings, elongate copulatory ducts, bilobed spermathecae. Male palp with distal patellar apophysis; tibia with basal and distal processes. Bulb with small tegular process, slender embolus protected by beak-like ventral process of triangular conductor.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,209 @@
<document id="21A87B5EEDC96B55C86EE4FB5A3656E2" ID-CLB-Dataset="308020" ID-DOI="10.3897/zookeys.1226.139148" ID-GBIF-Dataset="324061bd-6cf0-48f8-94db-83962d24ea9c" ID-ZooBank="413ACFE0-EA45-4033-8512-3ADF0AF2AEDD" ID-publisher-id="139148" URI-arpha="3105700B-0F4B-57E6-9BF5-3E4BD0688EFB" URI-zoobank="http://zoobank.org/413ACFE0-EA45-4033-8512-3ADF0AF2AEDD" article-type="rapid-communication" checkinTime="1738866268087" checkinUser="pensoft" docAuthor="Xie, Guanglin" docDate="2025" docId="D950883ACD8C5F7EBE494D9B1328B602" docLanguage="en" docName="ZooKeys 1226: 77-85" docOrigin="ZooKeys 1226" docSource="https://zookeys.pensoft.net/article/139148/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Apiocephalus yangmingae Xie, 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="85" masterDocId="3105700B0F4B57E69BF53E4BD0688EFB" masterDocTitle="Review of the genus Apiocephalus Gahan, 1898 (Coleoptera, Cerambycidae) with description of a new species" masterLastPageNumber="85" masterPageNumber="77" pageNumber="77" updateTime="1738866995953" updateUser="ExternalLinkService">
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<mods:title id="978E2BCF2C5D585F566F546489A2A0E2">Review of the genus Apiocephalus Gahan, 1898 (Coleoptera, Cerambycidae) with description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="1532D3B629BE0C44A32397F81FB55385" type="personal">
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</mods:role>
<mods:namePart id="52DA695BD11D6562A0D6601E848E7A5F">Xie, Guanglin</mods:namePart>
<mods:nameIdentifier id="11F24D54811D6278A80DC3FBE1B6E807" type="ORCID">0000-0003-3273-8985</mods:nameIdentifier>
<mods:affiliation id="A2D77057D02AB969929FF50DFFBCAA48">Institute of Entomology, College of Agriculture, Yangtze University, Jingzhou, Hubei, 434025, China</mods:affiliation>
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</mods:titleInfo>
<mods:part id="30B6B29C7303034EE3320E2BBC948E63">
<mods:date id="86FAD813FDC871B09913360D95A83F3D">2025</mods:date>
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<mods:number id="F082697B60676DA957DCF73B08B1BB13">2025-02-06</mods:number>
</mods:detail>
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<mods:identifier id="3922749F142585F014F7DBAA973816A2" type="DOI">10.3897/zookeys.1226.139148</mods:identifier>
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</mods:mods>
<treatment id="D950883ACD8C5F7EBE494D9B1328B602" ID-DOI="http://doi.org/10.5281/zenodo.14827222" ID-Zenodo-Dep="14827222" ID-arpha="D950883A-CD8C-5F7E-BE49-4D9B1328B602" ID-zoobank="https://zoobank.org/90F9D9C3-13A5-4195-83A3-E94B41D40BD4" LSID="urn:lsid:plazi:treatment:D950883ACD8C5F7EBE494D9B1328B602" LSID-ZBK="urn:lsid:zoobank.org:act:90F9D9C3-13A5-4195-83A3-E94B41D40BD4" httpUri="http://treatment.plazi.org/id/D950883ACD8C5F7EBE494D9B1328B602">
<subSubSection id="91264813A9E6D4F31E537282448482C2" type="nomenclature">
<paragraph id="89F7C40859022F7B5F89D48A847289C3">
<taxonomicName id="1CF0CFDCD20E501B81FA8D597929900F" authorityName="Xie" authorityYear="2025" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="yangmingae" status="sp. nov.">
<emphasis id="4C064084327A7F5CF194465C10D4CAA1" italics="true">Apiocephalus yangmingae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="714FE1639D62FB43F1F5952ABAD74870" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9166E455A550D9001B2846A3F150000A" type="description">
<paragraph id="68D905F45677796929CA30D3D639DC13">
<figureCitation id="AB5DE0C33B04846EDAB8397A59228788" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Apiocephalus yangmingae sp. nov., female a d habitus of adult e i terminalia a, e, i dorsal view b, f, h ventral view c lateral view d lateral view of the pronotum e, f sternite VIII g tignum h, i ovipositor c, d taken after remounted. scale: 1 mm (e i)." figureDoi="10.3897/zookeys.1226.139148.figure3" httpUri="https://binary.pensoft.net/fig/1243456">Chinese common name: 阳明头花天牛 Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWCAE" type="description">
<paragraph id="E7AE6CE70896A928DCC55AFD41DCFC3D">
<heading id="34CDF9F7BBCB05E20C5675678685B55E" reason="title">Description.</heading>
</paragraph>
<paragraph id="C2368B026F20560A79400116BB1BE5CE">
<emphasis id="D2448D332FF527BC9B1B57CB2F4053D3" bold="true">
<emphasis id="9E17BEA0F3F7A4C160C3784539590E14" bold="true" italics="true">
<typeStatus id="076E81682796E734766605ACFD52D0C6" type="holotype">Holotype</typeStatus>
</emphasis>
, female.
</emphasis>
Body length
<quantity id="3689EA789C932514B0DFE8ACCC3D0909" metricMagnitude="-2" metricUnit="m" metricValue="1.35" unit="mm" value="13.5">13.5 mm</quantity>
(from the front of mandible to elytral apex), humeral width
<quantity id="098B4C2AE6DFB6B60C7E9A296C1CD54F" metricMagnitude="-3" metricUnit="m" metricValue="4.5" unit="mm" value="4.5">4.5 mm</quantity>
. Body dull yellowish-brown to black, clothed with greyish-white, greyish-yellow to golden yellow pubescence.
<emphasis id="9598C3DE25A99410AC8F564FE90937DD" bold="true" italics="true">Head</emphasis>
black, clothed with greyish-yellow pubescence, which is thicker and denser on frons, upper half of genae and sides of the vertex; pubescence on sides of the vertex predominantly golden, forming two longitudinal stripes from antennal insertions to occiput.
<emphasis id="7DE1DB20C2543084E4A62B69681F3221" bold="true" italics="true">Pronotum</emphasis>
black, clothed with dense greyish-yellow pubescence on sides (including lateral spines) and provided with a golden yellow longitudinal pubescent stripe on either side of midline, connected anteriorly but not reaching basal margin posteriorly; the remainder clothed with rather sparse greyish-yellow pubescence.
<emphasis id="68B11D3DD2A7D66CDE6E99B5D20D8132" bold="true" italics="true">Scutellum</emphasis>
blackish-brown, clothed with greyish-white pubescence.
<emphasis id="038A88F5A625F5B3272D5E59378779A2" bold="true" italics="true">Elytra</emphasis>
dark brown, clothed with rather thin greyish-yellow to golden pubescence, and adorned with patches of relatively dense greyish-white pubescence; each elytron with a narrow transverse band at extreme base (golden near scutellum), extending inwards slightly behind humerus, then bending obliquely forward, forming an irregular ring; a broad transverse band behind middle, of which anterior boundary indistinct, posterior edge with five faintly visible spots of dense pubescence, each side containing a region of less greyish-white pubescence; an irregular transverse band before apex and a tuft of greyish-white setae at marginal angle.
<emphasis id="AFB5DF5502374C2B467B03395667870B" bold="true" italics="true">Underside</emphasis>
mostly clothed with dense and non-uniformly greyish-yellow pubescence, prosternum nearly glabrous except for intercoxal process; abdominal ventrites with a glabrous blackish-brown spot on each side. Legs mostly dull yellowish-brown, clothed with greyish-yellow pubescence.
</paragraph>
<caption id="29D5CC44817E2FAC92F92ED8A0B1A94D" ID-DOI="10.3897/zookeys.1226.139148.figure3" ID-arpha="054F88A7-D639-520A-B0A7-ED45E6DB9EEE" httpUri="https://binary.pensoft.net/fig/1243456" startId="F3">
<paragraph id="4C889FE6713D15992ED4CED1432921FB">
<label id="BDA15C50E4C6122034CBBB69895AA1B7">Figure 3.</label>
</paragraph>
<paragraph id="B86AACE93402B97054132C34E1397ABD">
Holotype of
<taxonomicName id="7DECEAA33C6FA363692CA538EF2769DF" authorityName="Xie" authorityYear="2025" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="yangmingae" status="sp. nov.">
<emphasis id="12F1D9F627CE578B8DC5B3AE72930AAD" italics="true">Apiocephalus yangmingae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95CEF96325A7EE2A0364A7DEF9377FCC" rank="species">sp. nov.</taxonomicNameLabel>
, female
<emphasis id="381521B6CCE09439623A7D1412005E18" bold="true">a d</emphasis>
habitus of adult
<emphasis id="002C741AB022B6D8FF1ED4182D972D18" bold="true">e i</emphasis>
terminalia
<emphasis id="B662B5872B3C093EBFE3BE157E5FE5DC" bold="true">a, e, i</emphasis>
dorsal view
<emphasis id="D592B880E10539CD6439958B5D4E246D" bold="true">b, f, h</emphasis>
ventral view
<emphasis id="4F8990C775214593BA1BC4387B2F0074" bold="true">c</emphasis>
lateral view
<emphasis id="A053D86A481D6875CCB216C2F3FFD497" bold="true">d</emphasis>
lateral view of the pronotum
<emphasis id="DC589E1962D5E6406BC63ECB2D3534C1" bold="true">e, f</emphasis>
sternite VIII
<emphasis id="5DE5FECE39C1CAB410C215FC65D06551" bold="true">g</emphasis>
tignum
<emphasis id="A9DECFA31C020B8045033E4DD138EEEC" bold="true">h, i</emphasis>
ovipositor
<emphasis id="76CA0B7C65E540EA4751BA5F78034253" bold="true">c, d</emphasis>
taken after remounted. scale: 1 mm (
<emphasis id="2CB0CD09202C2A632643A72B8112FE3A" bold="true">e i</emphasis>
).
</paragraph>
</caption>
<paragraph id="12050B9BF329B5E8EFD58DAACED387C9">
<emphasis id="8FE7DAA4C0DF71FAB3B8978C91F7FB0D" bold="true" italics="true">Head</emphasis>
prolonged and distinctly narrower than pronotum, featuring a median groove from antennal insertions to upper eye lobes; eyes obviously protruding outward; labrum twice as wide as long, emarginate on apical margin; genae slightly shorter than eye diameter; terminal segment of maxillary palpus prolonged spindle-shaped, with truncate apex. Antennae slightly longer than body, with the distal segment extending beyond the elytral apex; scape long, nearly reaching apical margin of pronotum, and strongly thickened apically; antennomere 3 about as long as antennomere 4 and shorter than antennomere 5.
<emphasis id="1394F935E2C1167C5202BA7BAAC7CDC2" bold="true" italics="true">Pronotum</emphasis>
slightly shorter than width between lateral spines; apical margin remarkably narrower than basal margin; sides conspicuously constricted at about apical fifth, with lower portion near apical edge distinctly expanded laterally; median spines well developed, blunt apically; disc coarsely and shallowly punctate, with four protuberances, of which the anterior pair before middle are rather low, the posterior pair behind middle are strongly raised, and also with several vague short longitudinal ridges near centre.
<emphasis id="DBF2109B8D305638D8FB549D4E9F9D7B" bold="true" italics="true">Scutellum</emphasis>
ligulate, slightly concave apically.
<emphasis id="43C747A2CE130DF53C467225FD53470E" bold="true" italics="true">Elytra</emphasis>
about two times as long as width across humeri, subparallel-sided, and arcuately convergent to apices on apical fifth; apices truncate, squarish at outer angles, rounded at inner angles; disc quite uneven, especially at base and apex; surface with extreme base granular-punctate, basal half deeply and closely punctate, apical half more shallowly and sparsely punctate; each elytron provided with an indistinct longitudinal costa.
<emphasis id="AC3777599D71B4923C75CF3EBB2D1965" bold="true" italics="true">Prosternal process</emphasis>
strongly arcuate, narrowed between procoxal cavities, then steeply sloped backwards and dilated apically.
<emphasis id="2E02802C2F1C0BA0D2B21A8A9361CEFB" bold="true" italics="true">Legs</emphasis>
long and slender; metafemora not reaching the elytral apex; metatarsomere 1 shorter than the remaining combined.
</paragraph>
<paragraph id="13CC220DEF17BEE0C26C8AD285E98E38">
<emphasis id="67029725941B273E40F3EFB4C1969E8C" bold="true">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYFAE" type="type material">
<paragraph id="8753300462248D5460565087A6CC6D18">
<heading id="0F8F9D2300E0C1473F2DBA1E5DB76A2B" reason="title">Type material.</heading>
</paragraph>
<paragraph id="E8B80F3854729FE7F6F7C9DB9B3A91E6">
<materialsCitation id="99445064986F62ADA397EE814DB14DB5" collectingDate="2018-05-08" collectionCode="YZU" collectorName="Guanglin Xie" country="China" county="Yangxian" elevation="1317" latitude="33.640003" location="Hongshiyao village" longLatPrecision="19" longitude="107.49" municipality="Huayang town" specimenCount="1" stateProvince="Shaanxi" typeStatus="holotype">
<emphasis id="4684022ED5548A4E0A4F612D6E00D071" bold="true" italics="true">
<typeStatus id="FCE29C9007BF9D8CA240F6944E7DCA2B" type="holotype">Holotype</typeStatus>
</emphasis>
,
<emphasis id="A7D5D5DDAF020A9759BC4460F5BE6B66" bold="true">
<collectingCountry id="6DB57F5721778EC6ABCC43A86A5924CD" name="China">China</collectingCountry>
</emphasis>
• female (
<abbrev content-type="institution" id="ABBRID0EFGAE" xlink_title="Insect Collection, College of Agriculture, Yangtze University, Jingzhou, China">
<collectionCode id="71970B9D1006FB4460243CD23A9DAAE1" country="China" lsid="urn:lsid:biocol.org:col:14820" name="Yuzhou University" type="Herbarium">YZU</collectionCode>
</abbrev>
);
<collectingRegion id="F111A494CC208882523F5BAEF04AB936" country="China" name="Shaanxi">Shaanxi</collectingRegion>
,
<collectingCounty id="66C6CA01127B1C547DB3374AF0563775">Yangxian</collectingCounty>
(洋县),
<collectingMunicipality id="0E1BAF0B2867467A62D0B8927F0216FF">Huayang town</collectingMunicipality>
(华阳镇),
<location id="D05EF33033B8F5FA93BF4B5ACAEB5677" LSID="urn:lsid:plazi:treatment:D950883ACD8C5F7EBE494D9B1328B602:D05EF33033B8F5FA93BF4B5ACAEB5677" country="China" county="Yangxian" latitude="33.640003" longLatPrecision="19" longitude="107.49" municipality="Huayang town" name="Hongshiyao village" stateProvince="Shaanxi">Hongshiyao village</location>
(红石窑村);
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENGAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.490000,33.640000]}">
<geoCoordinate id="5FB63FF8AE77CA47649BE2C2FCE37015" degrees="33" direction="north" minutes="38" orientation="latitude" precision="15" seconds="24" value="33.640003">33 ° 38 ' 24 &quot; N</geoCoordinate>
,
<geoCoordinate id="A017C4B738EA6D8426FE1902C37525BC" degrees="107" direction="east" minutes="29" orientation="longitude" precision="15" seconds="24" value="107.49">107 ° 29 ' 24 &quot; E</geoCoordinate>
</named-content>
; alt.
<quantity id="681D9777447E84C143FFAE22DD7326BE" metricMagnitude="3" metricUnit="m" metricValue="1.3169999999999997" unit="m" value="1317.0">
<elevation id="4A4E76DA9A8A2D76CCF011771A15565B" metricMagnitude="3" metricUnit="m" metricValue="1.3169999999999997" unit="m" value="1317.0">1317 m</elevation>
</quantity>
;
<date id="C0AC3B815A9D6587338874DC775C646B" value="2018-05-08">
<collectingDate id="6C3F59E14E303B5EECE47B25A6B52F41" value="2018-05-08">8 May 2018</collectingDate>
</date>
;
<collectorName id="C919EE10AC990459ECB8D183A144448F">Guanglin Xie</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESGAE" type="distribution">
<paragraph id="164350F9B323CF24FB384E182905FFB8">
<heading id="D952385263844AE6E9CCFD5A7B253C0F" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="CCC030BE5502FA58CF77C4903D5F582A">
<collectingCountry id="33B5AF2D1186021B6BBD72CFF50F4D1D" name="China">China</collectingCountry>
:
<collectingRegion id="6ED6FB13228197647F40ECD85E3196CE" country="China" name="Shaanxi">Shaanxi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXGAE" type="etymology">
<paragraph id="7A4A717BDED9B090AD4547421FB6499E">
<heading id="28F43FE4D79969A6BB3A0E705F7C0AA7" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="5B74D36B5E7889DF8C85C08C1BE440EB">The new species is named in honour of Ms Zhou Yangming, the authors mother, a traditional Chinese woman known for her gentle nature, intelligence, manual dexterity and family values. She passed away in 2022 while the author was visiting the Natural History Museum in London.</paragraph>
</subSubSection>
<subSubSection id="SECID0E3GAE" type="diagnosis">
<paragraph id="94D4CF03A1D2312913361C66EEEE6F1B">
<heading id="47DA1F2D96B8BE743F92FC899E81F7FF" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="09558641E14DD34B043C66BA529B7701">This new species can be distinguished from its congeners by the more exposed dorsum of the head and pronotum, both featuring golden yellow longitudinal stripes, more elongate pronotal lateral spines, and different elytral markings.</paragraph>
</subSubSection>
<subSubSection id="SECID0EBHAE" type="remarks">
<paragraph id="3F39282BEB1F14B2B54B9D9D47496A93">
<heading id="EAB270396BF4F157143D114150449BAB" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="CEFBCE2BBDEAAEFC45311ED67D9BFD1C">The new species was collected on an unknown flower, suggesting a flower-visiting habit.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,119 @@
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<mods:title id="255388D92A8E88302E41819597524DED">Review of the genus Apiocephalus Gahan, 1898 (Coleoptera, Cerambycidae) with description of a new species</mods:title>
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<paragraph id="7840A8B0EDC6618E8C981081D6803804">
<heading id="FB0079AE533AE991EF3C0D7B50F274A5" reason="label">Genus</heading>
<taxonomicName id="410BB92E74266E84325EAC869F46C446" ID-CoL="YM9" authority="Gahan, 1898" authorityName="Gahan" authorityYear="1898" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="154A0CCB0529548C1797F13F5048B955" italics="true">Apiocephalus</emphasis>
<bibRefCitation id="76DD988CB340165F0B19192AD277F6A4" DOI="10.1080/00222939808678011" author="Gahan CJ" refId="B6" refString="Gahan CJ (1898) Descriptions of new Longicorn Coleoptera from East Africa. The Annals and Magazine of Natural History Series 7, 2 (7): 40 - 59. https://doi.org/10.1080/00222939808678011" year="1898">Gahan, 1898</bibRefCitation>
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<taxonomicName id="EED861434B1DE3146A689E9D34349E5D" ID-CoL="YM9" authority="Gahan, 1898: 42" authorityName="Gahan" authorityPageNumber="42" authorityYear="1898" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="F32B5542D0E2591FF0B2328D8179320F" italics="true">Apiocephalus</emphasis>
<comment id="97A99FA1CF488893EA89AC050E7F45C9">
<bibRefCitation id="1B148720CD5578F5EF4548C0BAB1057C" DOI="10.1080/00222939808678011" author="Gahan CJ" refId="B6" refString="Gahan CJ (1898) Descriptions of new Longicorn Coleoptera from East Africa. The Annals and Magazine of Natural History Series 7, 2 (7): 40 - 59. https://doi.org/10.1080/00222939808678011" year="1898">Gahan, 1898: 42</bibRefCitation>
;
<treatmentCitation id="9E45730D1B5D4287EAA8613465F1E6A4" author="Gahan CJ" page="74" year="1906">
<bibRefCitation id="0C750E57E26043904A252CB3B7A0E81D" author="Gahan CJ" firstAuthor="Gahan" page="74" refId="B7" refString="Gahan CJ (1906) The Fauna of British India, including Ceylon and Burma. Coleoptera. Vol. I (Cerambycidæ). Bingham CT (Ed.) London, 1329." year="1906">Gahan 1906: 74</bibRefCitation>
</treatmentCitation>
[redescription].
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EYFAC" type="type species">
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<heading id="AE72648CFA4AD2A24078C9EB03E3D26C" reason="title">
<typeStatus id="904D4EC5EBEE5A7AF0492A878AC2AF45">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="6D9029F1428F6EFB80ABCEEB1A318778">
<taxonomicName id="AC888132C26C269912011613FD95F749" authority="Gahan, 1898" authorityName="Gahan" authorityYear="1898" class="Insecta" family="Cerambycidae" genus="Apiocephalus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="E2A0ECB6A28212F1828E326ABA79B8DD" italics="true">Apiocephalus punctipennis</emphasis>
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</taxonomicName>
.
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</subSubSection>
<subSubSection id="SECID0EJGAC" type="redescription">
<paragraph id="58CFD5F66419A5117EC4ABF9ACCC7081">
<heading id="26E23C321C04727A85639B2277DE4A04" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="5B325E6A6E005E8607FA527BB37FD306">Head prolonged, markedly narrowed behind eyes; eyes prominent, coarsely faceted. Antennae longer than body; scape long and curved, thickened apically; antennomere 3 approximately equal in length to antennomere 4 but distinctly shorter than antennomere 5; antennomeres 510 gradually decreasing in length. Pronotum strongly narrowed anteriorly, constricted before apex, with two developed lateral spines; disc provided with four protuberances, of which the anterior two are before middle, low, and the posterior two are after middle, strongly ridged. Elytra much broader than pronotum, with sides subparallel in basal two-thirds, thence converging more strongly towards apices which are truncate. Male metatarsomere 1 approximately as long as metatarsomere 2 and 3 combined.</paragraph>
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<subSubSection id="SECID0EOGAC" type="distribution">
<paragraph id="6F1DEDA4338EFFC93E5CBE02802C4A12">
<heading id="0E0DF5CB6E6990DC4618DBBA95DC8EEB" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="E053296C6EC8DD2966CD7BEA41717D65">
<collectingCountry id="69B42DC190F741EF25D2EEE4C5959553" name="Kenya">Kenya</collectingCountry>
,
<collectingCountry id="A39971417D608E4A5748AA0C111042C7" name="India">India</collectingCountry>
,
<collectingCountry id="866C43130826E8971B2873ACCCFB85B5" name="China">China</collectingCountry>
(new country record).
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<subSubSection id="SECID0ETGAC" type="remarks">
<paragraph id="4CD21087E43E871641E179154023DCCA">
<heading id="2854E18995C3B5132C274A47938381B2" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="884B3505EB4E167B19CA89719ABDD1A3">
This genus was first found in “
<collectingCountry id="6C973C1156055096428908055E4DDD75" name="Kenya">British East Africa</collectingCountry>
” (now part of
<collectingCountry id="6D3B163C64019ED390288507194F5AD9" name="Kenya">Kenya</collectingCountry>
). The second species was discovered in northern
<collectingCountry id="E90622DA97AAA06E3451A446B386717B" name="India">India</collectingCountry>
. The third species, to be described below, comes from central
<collectingCountry id="5F630A7CB0A9BEBFEFFFB8CF72CF450A" name="China">China</collectingCountry>
. This scattered distribution suggests that the genus exhibits a continental distribution pattern. It also indicates that with further research, additional new taxa may be discovered across the continents of Africa, Europe and Asia.
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<taxonomicName id="FA74F735F1D58AE9833CD698FD5AF5DF" authorityName="Hedin &amp; Ramírez &amp; Monjaraz-Ruedas" authorityYear="2025" class="Arachnida" family="Cybaeidae" genus="Siskiyu" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="armilla" status="sp. nov.">
<emphasis id="B2C490E83C060E6D0153A4D3ECF5CDC5" italics="true">Siskiyu armilla</emphasis>
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<taxonomicNameLabel id="770D8FE811929C3E5727EFB1F5341C45" rank="species">sp. nov.</taxonomicNameLabel>
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</subSubSection>
<subSubSection id="3F79C57A904EB4E5BAD72D79FC25B019" type="description">
<paragraph id="57821C9436CA1D7A47AFBF46595876D4">
<figureCitation id="A2BD3EE32AEECE99C7609B7F18939B55" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Images of live cybaeid spiders. A Neocryphoeca beattyi ♀, Pinal County, AZ, RWM 22 _ 011 B Blabomma sp ♀, San Diego County, CA, MCH 22 _ 020 C Siskiyu armilla sp. nov. immature ♀, Siskiyou County CA, MCH 24 _ 057; patellar fracture line labeled D Siskiyu armilla sp. nov. immature ♂, Siskiyou County CA, MCH 24 _ 057; with silk webbing presumed to belong to the spider E Cybaeota shastae ♀ Chamberlin &amp; Ivie, 1937, East Rosebud Creek, MT F Cybaeus somesbar ♀ Bennett, 2009, Shasta County, CA, MCH 24 _ 012. All images taken by M Hedin, except for Neocryphoeca (RW Mendez)." figureDoi="10.3897/zookeys.1226.140204.figure1" httpUri="https://binary.pensoft.net/fig/1243090">Figs 1</figureCitation>
,
<figureCitation id="0201B143425261BF2DE4EAF9E7A5DAE0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siskiyu armilla sp. nov. holotype ♂ (SDSU _ G 1090) palp A ventral view B lateral view. Elongate cymbium, tegular process, embolus, conductor processes, patellar apophysis, and RTA basal and distal processes labeled." figureDoi="10.3897/zookeys.1226.140204.figure6" httpUri="https://binary.pensoft.net/fig/1243095">6</figureCitation>
,
<figureCitation id="A329872F4B43D239251DD7EF4858C06F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siskiyu armilla sp. nov. paratype ♀ (SDSU _ G 1086 A) A dorsal view B ventral view C leg IV patellar fracture line D ventral epigynum E dorsal epigynum. Patellar fracture lines and epigynal dorsal ridge, copulatory openings (co), copulatory ducts (cd), bilobed spermathecae and fertilization ducts (fd) labeled." figureDoi="10.3897/zookeys.1226.140204.figure7" httpUri="https://binary.pensoft.net/fig/1243096">7</figureCitation>
,
<figureCitation id="641228116D4093AC14732C4FD663203B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Species distributions of Siskiyu armilla sp. nov. and Cybaeozyga, including C. furtiva sp. nov. and C. heterops. Locations approximate, see text and Suppl. material 1 for precise location data." figureDoi="10.3897/zookeys.1226.140204.figure10" httpUri="https://binary.pensoft.net/fig/1243099">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKYBG" type="material">
<paragraph id="48B03B9042E24A6737ECEBEAB60FE0A7">
<heading id="9D1AA0664106D2239999B49926CA60E9" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="54AEE4A42B7BBB64FE9BA5AC673F23E9">
<materialsCitation id="9E50C8ABB3026CD07CAC064A28247CF6" collectingDate="2006-08-15" collectionCode="R, SDSU" collectorName="M. Hedin &amp; Keith, M" county="Josephine Co." latitude="42.1643" location="Nelson Creek" longLatPrecision="6" longitude="-123.5008" municipality="Hwy" specimenCode="G 1090" specimenCount="1" specimenCount-male="1" stateProvince="Oregon" typeStatus="holotype">
<emphasis id="DE53EF9B65E51C2AEB2817B5E73A9783" bold="true">
<emphasis id="1798DC7050EF7FB0A1E8F9E7A0802452" bold="true" italics="true">
<typeStatus id="EE4BA2A9AF472F4E18D355378D5B4841" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingRegion id="6C9DBFE75AE410DE56B50385C53CFD6D" country="United States of America" name="Oregon">Oregon</collectingRegion>
,
<collectingCounty id="764EB1A441EAD73CF238D7CFC340DC8C">Josephine Co.</collectingCounty>
</emphasis>
<specimenCount id="249B29F475EA491305CD5D70DCA1F632" count="1" type="male"></specimenCount>
;
<collectingMunicipality id="F7FDBFBCB90BBA3E2BBC5261CF635E80">Hwy</collectingMunicipality>
46,
<locationDeviation id="9DB539319FC87FCAF73C7421AB2FC571" location="Cave Junction">
<quantity id="9FAC8F1E1215398BEC77539C8E1306B0" metricMagnitude="4" metricUnit="m" metricValue="1.33575552" unit="mi" value="8.3">8.3 mi.</quantity>
E Cave Junction
</locationDeviation>
, near
<location id="94D2B7440A2DA8082017026BA2693FC3" LSID="urn:lsid:plazi:treatment:F2A4040E5FF95633B2C103F53A888CFE:94D2B7440A2DA8082017026BA2693FC3" county="Josephine Co." latitude="42.1643" longLatPrecision="6" longitude="-123.5008" municipality="Hwy" name="Nelson Creek" stateProvince="Oregon">Nelson Creek</location>
, along
<location id="9DE4850D850C988349482AAFA3BDD071" LSID="urn:lsid:plazi:treatment:F2A4040E5FF95633B2C103F53A888CFE:9DE4850D850C988349482AAFA3BDD071" county="Josephine Co." latitude="42.1643" longLatPrecision="6" longitude="-123.5008" municipality="Hwy" name="Sucker Creek" stateProvince="Oregon">Sucker Creek</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYYBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-123.500800,42.164300]}">
<geoCoordinate id="87DB436DA52A3EF61AAEC1C27CBC9B9A" degrees="42.1643" direction="north" orientation="latitude" precision="5" value="42.1643">42.1643</geoCoordinate>
,
<geoCoordinate id="EE1996333C57B9C8603853D888BBE0B5" degrees="123.5008" direction="west" orientation="longitude" precision="5" value="-123.5008">- 123.5008</geoCoordinate>
</named-content>
;
<date id="8ECE803AF7C27DB7F6669793788889D8" value="2006-08-15">
<collectingDate id="1D1E8E88EEAA2039BB61F29F8D9485E3" value="2006-08-15">15 Aug. 2006</collectingDate>
</date>
; coll.
<collectorName id="9286D936BB9E0690928649233358510B">M. Hedin</collectorName>
,
<collectionCode id="E544A655FC56C147BFC7966AF65C3197" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectorName id="1E9B0450ADFCF883D9D0951499B997F3">Keith, M</collectorName>
. McCormack; MCH 06 _ 115;
<collectionCode id="1F3BAC4D6A2FFE9BFC912D96A4B83C97" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="E22A04C40B209A5B2190F0B63DB3DB18" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">G 1090</specimenCode>
</materialsCitation>
;
<emphasis id="56176B94A285C8738A996217B4B4181A" bold="true">
<materialsCitation id="06875B4DE3CDF9702ACCBBE8EDF2EA12" location="California" specimenCount="1" stateProvince="California" typeStatus="paratype">
<emphasis id="B67C3B37A23D9DEFAD582FACF949ACE5" bold="true" italics="true">
<typeStatus id="25934969474BAB87FAC0189F99163CE5" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingRegion id="C5939160FE5616164B2BAA07E1E4D975" country="United States of America" name="California">California</collectingRegion>
, Siskiyou Co
</materialsCitation>
.
</emphasis>
<materialsCitation id="21F55283ADF6C8ACE8F56B0D872C3BD3" collectingDate="2006-04-18" collectionCode="SDSU" collectorName="M. Hedin" county="West Fork Beaver Creek" latitude="41.9455" location="Forest Road" longLatPrecision="6" longitude="-122.8334" municipality="Little Soda Creek" specimenCode="G 1086" specimenCount="2" specimenCount-female="2" stateProvince="California" typeStatus="paratype">
<specimenCount id="8795CE48FEAC79AB1DF802AD20CF4E49" count="2" type="female">2 ♀</specimenCount>
;
<collectingCounty id="E6C998C84484D39B1083FC52BDD99B3D">West Fork Beaver Creek</collectingCounty>
, confluence with
<collectingMunicipality id="4DE183122D5424935687F6C1ADA074F3">Little Soda Creek</collectingMunicipality>
,
<location id="EEA84980C66F03170813DA7E57AC0CF0" LSID="urn:lsid:plazi:treatment:F2A4040E5FF95633B2C103F53A888CFE:EEA84980C66F03170813DA7E57AC0CF0" county="West Fork Beaver Creek" latitude="41.9455" longLatPrecision="6" longitude="-122.8334" municipality="Little Soda Creek" name="Forest Road" stateProvince="California">Forest Road</location>
47 N 01;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EEZBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-122.833400,41.945500]}">
<geoCoordinate id="223A5997E5F5C522968CD3996B0752B6" degrees="41.9455" direction="north" orientation="latitude" precision="5" value="41.9455">41.9455</geoCoordinate>
,
<geoCoordinate id="CD83B94972D4E3885D38428EE5695EDA" degrees="122.8334" direction="west" orientation="longitude" precision="5" value="-122.8334">- 122.8334</geoCoordinate>
</named-content>
;
<date id="F2F7EACC428A4631292D2017E985D63C" value="2006-04-18">
<collectingDate id="4C610893D1E751765149E970BBCF7708" value="2006-04-18">18 Apr. 2006</collectingDate>
</date>
; coll.
<collectorName id="1D8030C68502A0DE1AF85276F22DCAAD">M. Hedin</collectorName>
; MCH 06 _ 066;
<collectionCode id="0C43F020602EBC78CFFA87FA046C8CD3" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="2A78877B6F901D76568F357475B2DED4" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">G 1086</specimenCode>
,
<collectionCode id="602AE93DDFF90FAA3B839725E91327C0" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="01731CD7D7CBF1497E43428449E648C1" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">G 1086</specimenCode>
A
</materialsCitation>
; •
<materialsCitation id="2B4051A4692411C677A4BD2C0C1BAF9E" collectingDate="2005-07-10" collectingDateMax="2005-07-12" collectingDateMin="2005-07-10" collectorName="M. Hedin" latitude="41.1863" location="Salmon River" longLatPrecision="6" longitude="-123.2148" municipality="Matthews Creek" specimenCount="3" specimenCount-female="3" stateProvince="California" typeStatus="paratype">
<specimenCount id="18B55C0AAE621235FD785873D2A058A7" count="3" type="female">3 ♀</specimenCount>
;
<collectingMunicipality id="042A362B0D781E9360D867CB977A08E9">Matthews Creek</collectingMunicipality>
campground, on
<location id="953CA6D70DF1BDC01AB4B9EEFFEF1D6C" LSID="urn:lsid:plazi:treatment:F2A4040E5FF95633B2C103F53A888CFE:953CA6D70DF1BDC01AB4B9EEFFEF1D6C" latitude="41.1863" longLatPrecision="6" longitude="-123.2148" municipality="Matthews Creek" name="Salmon River" stateProvince="California">Salmon River</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMZBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-123.214800,41.186300]}">
<geoCoordinate id="EC03E8F59182EC111EECC4310C032FCE" degrees="41.1863" direction="north" orientation="latitude" precision="5" value="41.1863">41.1863</geoCoordinate>
,
<geoCoordinate id="6619225F69AEF10AF76319E67BBA0184" degrees="123.2148" direction="west" orientation="longitude" precision="5" value="-123.2148">- 123.2148</geoCoordinate>
</named-content>
;
<date id="DE64C74D06F2A285C4093D7C07995403" value="2005-07-10" valueMax="2005-07-12" valueMin="2005-07-10">
<collectingDate id="7F7D8EF6F524C9FEFACD8062BC358049" value="2005-07-10" valueMax="2005-07-12" valueMin="2005-07-10">1012 July 2005</collectingDate>
</date>
; coll.
<collectorName id="62F7AD0AE750B4170257946040AFCEA7">M. Hedin</collectorName>
; MCH 05 _ 029
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERZBG" type="material">
<paragraph id="85F9D1CE4C44A88CD1BD54A0C2CEE571">
<heading id="3CFB189D9F958D8F59ECD287BA2B2C43" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="C6102217938A06D4DCDF4D94D52CAED8">
<emphasis id="1228F2D6478B1649DA02C70A0E558A56" bold="true">
<materialsCitation id="13D56AD7783F4DCBD7A8C9C903514BAA" collectorName="Josephine Co" country="United States of America" location="Oregon" specimenCount="1" stateProvince="Oregon">
<collectingCountry id="58E1122931CB7110C618BA3AEDF0B706" name="United States of America">USA</collectingCountry>
<collectingRegion id="0285DE47709CA6FA790B4922BD9246E2" country="United States of America" name="Oregon">Oregon</collectingRegion>
,
<collectorName id="45F694CBC3C0C778D8AC178BC9023066">Josephine Co</collectorName>
</materialsCitation>
.
</emphasis>
<materialsCitation id="608198809E861C746F616C5E9B3737EF" collectingDate="2006-08-15" collectionCode="R, SDSU" collectorName="M. Hedin &amp; Keith, M &amp; McCormack" country="United States of America" county="Hwy" latitude="42.1643" location="Sucker Creek" longLatPrecision="6" longitude="-123.5008" municipality="Nelson Creek" specimenCode="G 1089" specimenCount="1" specimenCount-male="1" stateProvince="Oregon">
<specimenCount id="F49C441D8C9AADCAA077B3C7B9FFF65E" count="1" type="male"></specimenCount>
;
<collectingCounty id="5038EC42D156FEE5AEA17B75BB6D022F">Hwy</collectingCounty>
46,
<locationDeviation id="C61A9CD61C2A3B9A92AD1F487CC1DF7D" location="Cave Junction">
<quantity id="E224E9DE5845712DBD375DFD52332B98" metricMagnitude="4" metricUnit="m" metricValue="1.33575552" unit="mi" value="8.3">8.3 mi.</quantity>
E Cave Junction
</locationDeviation>
, near
<collectingMunicipality id="237F3B5C0A589855D98CDF872EA10631">Nelson Creek</collectingMunicipality>
, along
<location id="D642204B8029B60F096F4423159A5D5F" LSID="urn:lsid:plazi:treatment:F2A4040E5FF95633B2C103F53A888CFE:D642204B8029B60F096F4423159A5D5F" country="United States of America" county="Hwy" latitude="42.1643" longLatPrecision="6" longitude="-123.5008" municipality="Nelson Creek" name="Sucker Creek" stateProvince="Oregon">Sucker Creek</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3ZBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-123.500800,42.164300]}">
<geoCoordinate id="3E1D197151C1F06D379221E98EBCBC2F" degrees="42.1643" direction="north" orientation="latitude" precision="5" value="42.1643">42.1643</geoCoordinate>
,
<geoCoordinate id="A312DFF3435053F7CD44AE746D07D639" degrees="123.5008" direction="west" orientation="longitude" precision="5" value="-123.5008">- 123.5008</geoCoordinate>
</named-content>
;
<date id="D133B5AF23122E85D724A41562E9639A" value="2006-08-15">
<collectingDate id="7BAE0F5B5F9A65108FC94FA3F1082229" value="2006-08-15">15 Aug. 2006</collectingDate>
</date>
; coll.
<collectorName id="81BF85DEEFA16E6E2AFEAAADFA9A1241">M. Hedin</collectorName>
,
<collectionCode id="888DB9277EABEFFBA4E484F3704BFCA1" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectorName id="F08E99DE20FB520747AA1B32EBEFCFAF">Keith, M</collectorName>
.
<collectorName id="CC9C9F2A3B88573540CC2714F39D584F">McCormack</collectorName>
; MCH 06 _ 115;
<collectionCode id="C0A692E74B93D50238A4D073C5F474D3" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="8244747392FDC2DA1A1C9E165A23955A" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">G 1089</specimenCode>
</materialsCitation>
;
<emphasis id="F79474AAC25840D0631761AA06A50E0C" bold="true">
<collectingRegion id="1EF9D54AAEEB9B09ED467FA910D15CC9" country="United States of America" name="California">California</collectingRegion>
, Siskiyou Co.
</emphasis>
<materialsCitation id="B0C250F622BBD4D6053BCE1EA48BFEF4" collectingDate="2006-04-18" collectionCode="SDSU" collectorName="M. Hedin" country="United States of America" county="West Fork Beaver Creek" latitude="41.9455" location="Forest Road" longLatPrecision="6" longitude="-122.8334" municipality="Little Soda Creek" specimenCode="G 1087" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="56F386A38B1E5DFC76C90D1A48810AB3" count="1" type="female"></specimenCount>
;
<collectingCounty id="8873E85298C9FD668DD867876861E74B">West Fork Beaver Creek</collectingCounty>
, confluence with
<collectingMunicipality id="CF772E7D28B8A7DAFC6FF699C8E6B405">Little Soda Creek</collectingMunicipality>
,
<location id="C2B97D22009783387848D760DDA4D5DE" LSID="urn:lsid:plazi:treatment:F2A4040E5FF95633B2C103F53A888CFE:C2B97D22009783387848D760DDA4D5DE" country="United States of America" county="West Fork Beaver Creek" latitude="41.9455" longLatPrecision="6" longitude="-122.8334" municipality="Little Soda Creek" name="Forest Road" stateProvince="California">Forest Road</location>
47 N 01;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EG1BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-122.833400,41.945500]}">
<geoCoordinate id="CCC60427C270DF72CA10552CD640C68E" degrees="41.9455" direction="north" orientation="latitude" precision="5" value="41.9455">41.9455</geoCoordinate>
,
<geoCoordinate id="9FD03A2E188F02791D1F9236BE175570" degrees="122.8334" direction="west" orientation="longitude" precision="5" value="-122.8334">- 122.8334</geoCoordinate>
</named-content>
;
<date id="D75FE33B42B46AC621572DFA2C4A57DF" value="2006-04-18">
<collectingDate id="9ED2A674BDE1146C6DED5EE530E490C3" value="2006-04-18">18 Apr. 2006</collectingDate>
</date>
; coll.
<collectorName id="C5BCD8E60DE96071B7C00B39213B7736">M. Hedin</collectorName>
; MCH 06 _ 066;
<collectionCode id="0720834AC79133168BB216FC56B35488" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="359D68BA0B16B798032320E61CBD97EB" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">G 1087</specimenCode>
</materialsCitation>
; •
<materialsCitation id="90E10794FBD3C57F7B28B281E04E39DA" collectingDate="2006-04-18" collectionCode="SDSU" collectorName="M. Hedin" country="United States of America" latitude="41.9238" location="Beaver Creek" longLatPrecision="6" longitude="-122.8321" municipality="Beaver Creek" specimenCode="G 1088" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="E7C422281F67795AB91FF2309729ED4F" count="1" type="female"></specimenCount>
;
<collectingMunicipality id="CA590767E40860048BEC9D42AEC5EDC9">Beaver Creek</collectingMunicipality>
Rd, near
<location id="DF62CB83427EE3BF67B9EA488BBC7864" LSID="urn:lsid:plazi:treatment:F2A4040E5FF95633B2C103F53A888CFE:DF62CB83427EE3BF67B9EA488BBC7864" country="United States of America" latitude="41.9238" longLatPrecision="6" longitude="-122.8321" municipality="Beaver Creek" name="Beaver Creek" stateProvince="California">Beaver Creek</location>
campground;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EO1BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-122.832100,41.923800]}">
<geoCoordinate id="946446BAC407A63A9633542B36EF2EAB" degrees="41.9238" direction="north" orientation="latitude" precision="5" value="41.9238">41.9238</geoCoordinate>
,
<geoCoordinate id="DDDA943700918FB0E3A55E506264A399" degrees="122.8321" direction="west" orientation="longitude" precision="5" value="-122.8321">- 122.8321</geoCoordinate>
</named-content>
;
<date id="62E50F7155C5F1806B8244616FE92462" value="2006-04-18">
<collectingDate id="2441CFBFB18355271D6B8063EE657557" value="2006-04-18">18 Apr. 2006</collectingDate>
</date>
; coll.
<collectorName id="72A9F0DF087E7FD2DD13B489FAD3CF6E">M. Hedin</collectorName>
; MCH 06 _ 065;
<collectionCode id="18FFEE2F32C11107B65581A449CB8E84" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="63CE5235F51ABDB1DC19CE38E7EE6AE3" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">G 1088</specimenCode>
</materialsCitation>
; •
<materialsCitation id="CFB135EC9B804FE9FEE7855B52EA95F9" collectingDate="2024-07-22" collectorName="M. Hedin &amp; O. Hedin" country="United States of America" latitude="41.9426" location="Fishtrap Creek" longLatPrecision="6" longitude="-122.8024" municipality="Beaver Creek" specimenCount="1" stateProvince="California">
several immatures; FR 11, along
<collectingMunicipality id="CBB177FDEC88D743BC2299288A1A0EA4">Beaver Creek</collectingMunicipality>
, below confluence with
<location id="705822C85C9EFD19E212DE8004FBC979" LSID="urn:lsid:plazi:treatment:F2A4040E5FF95633B2C103F53A888CFE:705822C85C9EFD19E212DE8004FBC979" country="United States of America" latitude="41.9426" longLatPrecision="6" longitude="-122.8024" municipality="Beaver Creek" name="Fishtrap Creek" stateProvince="California">Fishtrap Creek</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EW1BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-122.802400,41.942600]}">
<geoCoordinate id="21D3D2CB2DBD03F623B09AB8448234A6" degrees="41.9426" direction="north" orientation="latitude" precision="5" value="41.9426">41.9426</geoCoordinate>
,
<geoCoordinate id="C4729949E7553AD2F35460DAF03ADCAC" degrees="122.8024" direction="west" orientation="longitude" precision="5" value="-122.8024">- 122.8024</geoCoordinate>
</named-content>
;
<date id="583D13FE2C14C8B0E18E803AC2F724AF" value="2024-07-22">
<collectingDate id="ABCA78D61586EB20AFE8DE561A35DDDB" value="2024-07-22">22 July 2024</collectingDate>
</date>
; coll.
<collectorName id="7BD91E3E16B8C31D3DE1D13EBA66AD24">M. Hedin</collectorName>
,
<collectorName id="43145A499F57C0C7B198178BE128F7F6">O. Hedin</collectorName>
; MCH 24 _ 057
</materialsCitation>
; •
<materialsCitation id="6EBBB601634B0495DEB65C344A3A5C31" collectingDate="2005-07-10" collectingDateMax="2005-07-12" collectingDateMin="2005-07-10" collectionCode="SDSU" collectorName="M. Hedin" country="United States of America" latitude="41.1863" location="Salmon River" longLatPrecision="6" longitude="-123.2148" municipality="Matthews Creek" specimenCode="G 0570" specimenCount="1" specimenCount-female="1" stateProvince="California">
<specimenCount id="DBC39D60F541ABF5CB68CEDFC5151460" count="1" type="female"></specimenCount>
;
<collectingMunicipality id="1601069526CCCD6A1A51E51840D77DCA">Matthews Creek</collectingMunicipality>
campground, on
<location id="60D7D28A6261927C35232AF0BD574162" LSID="urn:lsid:plazi:treatment:F2A4040E5FF95633B2C103F53A888CFE:60D7D28A6261927C35232AF0BD574162" country="United States of America" latitude="41.1863" longLatPrecision="6" longitude="-123.2148" municipality="Matthews Creek" name="Salmon River" stateProvince="California">Salmon River</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E51BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-123.214800,41.186300]}">
<geoCoordinate id="942C02841B36C50515310BFAF66E3300" degrees="41.1863" direction="north" orientation="latitude" precision="5" value="41.1863">41.1863</geoCoordinate>
,
<geoCoordinate id="E7F9E449E32530ADF83198B1D86532B6" degrees="123.2148" direction="west" orientation="longitude" precision="5" value="-123.2148">- 123.2148</geoCoordinate>
</named-content>
;
<date id="BB166738332D4FCE39D1732DA38C4E0C" value="2005-07-10" valueMax="2005-07-12" valueMin="2005-07-10">
<collectingDate id="C7BCB7CC7CBCB4C4669E6814CFC7FB4C" value="2005-07-10" valueMax="2005-07-12" valueMin="2005-07-10">1012 July 2005</collectingDate>
</date>
; coll.
<collectorName id="11114FD2B13639DB7BD66636A6A07A42">M. Hedin</collectorName>
; MCH 05 _ 029;
<collectionCode id="283E2045A5B7C77EC272E04A6B854250" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_
<specimenCode id="0E03B154C4F98E3B5E6C9F57C79707E2" collectionCode="SDSU" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">G 0570</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ED2BG" type="etymology">
<paragraph id="9F397241B06F12C8182EDD53B3AC4659">
<heading id="621F62D07F27BA4C412A45568AAB7505" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="21F67347C6DAD4B70D3840251475B686">
<emphasis id="2479391A2E48F9FC0811874E18030FDF" italics="true">armilla</emphasis>
(L., a ring, bracelet), from the conspicuous ring-like patellar fracture lines (Fig.
<figureCitation id="A48078888DFCFC9E57828D530B1CED19" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Images of live cybaeid spiders. A Neocryphoeca beattyi ♀, Pinal County, AZ, RWM 22 _ 011 B Blabomma sp ♀, San Diego County, CA, MCH 22 _ 020 C Siskiyu armilla sp. nov. immature ♀, Siskiyou County CA, MCH 24 _ 057; patellar fracture line labeled D Siskiyu armilla sp. nov. immature ♂, Siskiyou County CA, MCH 24 _ 057; with silk webbing presumed to belong to the spider E Cybaeota shastae ♀ Chamberlin &amp; Ivie, 1937, East Rosebud Creek, MT F Cybaeus somesbar ♀ Bennett, 2009, Shasta County, CA, MCH 24 _ 012. All images taken by M Hedin, except for Neocryphoeca (RW Mendez)." figureDoi="10.3897/zookeys.1226.140204.figure1" httpUri="https://binary.pensoft.net/fig/1243090">1 C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EX2BG" type="description">
<paragraph id="257227B0DC04B986F3BF357E8B2A95C9">
<heading id="26F67378607605669D5BAA261E962457" reason="title">
Description of
<specimenCount id="83558AF09860F640BB27B4F0CC006A29" count="1" type="male"></specimenCount>
<typeStatus id="6A023A27D5D5505687B5FFDD9479F1E9" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="F0C2FFEC5FAA7C5329C9ED37D7138EF4">
(
<collectionCode id="0C5126CC158AF8B85FC5ED41D636E807" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_ G 1090; Fig.
<figureCitation id="C459783C67781C4A8774F4BBBB83EF0D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siskiyu armilla sp. nov. holotype ♂ (SDSU _ G 1090) palp A ventral view B lateral view. Elongate cymbium, tegular process, embolus, conductor processes, patellar apophysis, and RTA basal and distal processes labeled." figureDoi="10.3897/zookeys.1226.140204.figure6" httpUri="https://binary.pensoft.net/fig/1243095">6 A, B</figureCitation>
) Color in alcohol pale cream to white, fangs slightly darker. Carapace essentially bare, unmarked; abdomen dusky dorsally and ventrally, slightly darker dorsal longitudinal bars posteriorly. Total length 2.3, carapace length 1.15, carapace width 0.85, cephalic region width 0.5, posterior eye row width 0.325. Eye diameters AME: ALE: PME: PLE = 0.025: 0.075: 0.05: 0.075. Leg I article lengths (1.65, 0.5, 1.9, 1.6, 0.9 = 6.4), leg
<collectionCode id="BA017F958B33631D64E92824AEE04791">IV</collectionCode>
article lengths (1.45, 0.4, 1.5, 1.5, 0.8 = 5.6). Leg macrosetae sparse, one or two on tibia
<collectionCode id="AF5742739C7D7E97D7E8079C44E6EEDB">III</collectionCode>
and
<collectionCode id="99131B75F7850FBE19C0D4EE2FC798E1" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
(prolateral and ventral), and metatarsus
<collectionCode id="698E050E5AA18D6F2C14B858A6C1D321">III</collectionCode>
and
<collectionCode id="BCC1180C457D0167F2D7451B3CC1EB9B">IV</collectionCode>
(prolateral and ventral, distal cluster).
</paragraph>
<paragraph id="CA00CFE75D96677D995309F42BD62938">
Male palp with shelf-like distal patellar apophysis. Basal retrolateral tibial apophysis (
<abbrev id="ABBRID0ED3BG" xlink_title="retrolateral tibial apophysis">
<collectionCode id="614B74EBEE8376CB1391266BB42EFE68">RTA</collectionCode>
</abbrev>
) a minute spike, with sclerotized parallel weak ridges bordering a slight excavation, extending to a short and blade-like distal
<abbrev id="ABBRID0EH3BG" xlink_title="retrolateral tibial apophysis">RTA</abbrev>
. Bulb with a short dorsal tegular process at base of whip-like embolus which extends to ~ 3 oclock then projects ventrally, protected by conductor. Triangular conductor with spatulate distal, and sharply pointed basal lobes. Beak-like ventral lobe of conductor enclosing embolus. Median apophysis absent. Cymbium elongate, distal projection approximately as long as bulb.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EL3BG" type="description">
<paragraph id="5F8AD252DA4F6B62B2E22C8F4F62E36F">
<heading id="6BCA21192ACCCED8C4C6EE228855CE1D" reason="title">
Description of
<specimenCount id="206AD1FB7D53E7E273762EE1593AADC8" count="1" type="female"></specimenCount>
<typeStatus id="596CFAF46535377876DC1CEE0E474751" type="paratype">paratype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="8896B6982F3D8A8FFA6B0E25425D0734">
(
<collectionCode id="CF6365B97CCD9937E2DC34C7EE546C38" country="USA" httpUri="http://grbio.org/cool/j1yc-pyd8" name="Severin-McDaniel Insect Collection" type="">SDSU</collectionCode>
_ G 1086 A; Fig.
<figureCitation id="DC12E36FF436FFE8CC71470FA032473A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siskiyu armilla sp. nov. paratype ♀ (SDSU _ G 1086 A) A dorsal view B ventral view C leg IV patellar fracture line D ventral epigynum E dorsal epigynum. Patellar fracture lines and epigynal dorsal ridge, copulatory openings (co), copulatory ducts (cd), bilobed spermathecae and fertilization ducts (fd) labeled." figureDoi="10.3897/zookeys.1226.140204.figure7" httpUri="https://binary.pensoft.net/fig/1243096">7 A E</figureCitation>
) Color in alcohol pale cream to orange, fangs slightly darker. Carapace essentially bare, unmarked; abdomen without markings. Total length 2.9, carapace length 1.3, carapace width 0.925, cephalic region width 0.55, posterior eye row width 0.325. Eye diameters AME: ALE: PME: PLE = 0.025: 1: 0.05: 1. Leg I article lengths (1.57, 0.5, 1.6, 1.4, 0.9 = 5.9), leg
<collectionCode id="9F76985EF393EDF7F1DA0243423B792C">IV</collectionCode>
article lengths (1.55, 0.4, 1.5, 1.5, 0.7 = 5.7). Leg macrosetae sparse, 1 on dorsal patella
<collectionCode id="4DAE0CC5106D50591B5765D7AA9A09A5">III</collectionCode>
and
<collectionCode id="918DB7003E08AC4D3DA4D4ECCA53B811">IV</collectionCode>
, 12 on tibia
<collectionCode id="083F4DDD486C4111B4AAFAEE352C48C0">III</collectionCode>
and
<collectionCode id="A550BB1D82DDFDC687F6290AF36BFD82" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
(prolateral and ventral), and metatarsus
<collectionCode id="81AD84B9FE23B88881D3BFDA7BA452F2">III</collectionCode>
and
<collectionCode id="95092DDECEE83F6200EF8B74F4D371D2">IV</collectionCode>
(prolateral and ventral, distal cluster).
</paragraph>
<paragraph id="208BDE8E461D49F07B5404F7641D51FD">
Epigynal plate with a semicircular ridge along the anterior edge (Fig.
<figureCitation id="0682794C3A57A4695D93F8A939024D95" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siskiyu armilla sp. nov. paratype ♀ (SDSU _ G 1086 A) A dorsal view B ventral view C leg IV patellar fracture line D ventral epigynum E dorsal epigynum. Patellar fracture lines and epigynal dorsal ridge, copulatory openings (co), copulatory ducts (cd), bilobed spermathecae and fertilization ducts (fd) labeled." figureDoi="10.3897/zookeys.1226.140204.figure7" httpUri="https://binary.pensoft.net/fig/1243096">7 D, E</figureCitation>
). Paired lateral ear-shaped copulatory openings, sclerotized laterally and ventrally, lie ventral to this ridge. Thin copulatory ducts arise from copulatory openings and loop dorsally to meet in the midline, then extend ventrally in parallel, diverge at the bottom, then extend dorsally to connect with spermathecae. Paired spermathecae bilobed, medial lobe slightly smaller and more oval-shaped than larger and more spherical lateral lobes, connected by a short, looped duct. Fertilization ducts short and indistinct, extending ventrally from medial lobes.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E23BG" type="variation">
<paragraph id="7BE20565ADAB496D1769050868A50CB7">
<heading id="55B04380083A8E535D8642D894FCAD3F" reason="title">Variation.</heading>
</paragraph>
<paragraph id="A473435E57B159FA9C61CC9DDD80917C">
Female epigyna from the three geographically distant known locations (Cave Junction, Beaver Creek, Salmon River, Fig.
<figureCitation id="5818BA05ACAB37E7E9BFC45288EDB0E6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Species distributions of Siskiyu armilla sp. nov. and Cybaeozyga, including C. furtiva sp. nov. and C. heterops. Locations approximate, see text and Suppl. material 1 for precise location data." figureDoi="10.3897/zookeys.1226.140204.figure10" httpUri="https://binary.pensoft.net/fig/1243099">10</figureCitation>
) are very similar in detail.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EF4BG" type="distribution">
<paragraph id="FAAF98DC148A40A8704BD2DC656A87DB">
<heading id="605CD76F16B7D06493E7D1057A8D9339" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="0168A48EEF3FB15012565354E11C22EB">
Known from five separate locations (three in close vicinity near Beaver Creek), spanning from northern records near Cave Junction,
<collectingRegion id="5F1ED93EB2A6BEB8ADCC92FA6FC28C19" country="United States of America" name="Oregon">Oregon</collectingRegion>
to southern records along the Salmon River in Siskiyou County,
<collectingRegion id="ADCC875ABE6D35A933778D093E013444" country="United States of America" name="California">California</collectingRegion>
(Fig.
<figureCitation id="BBAE288DA13F3C325047F7AD9530F709" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Species distributions of Siskiyu armilla sp. nov. and Cybaeozyga, including C. furtiva sp. nov. and C. heterops. Locations approximate, see text and Suppl. material 1 for precise location data." figureDoi="10.3897/zookeys.1226.140204.figure10" httpUri="https://binary.pensoft.net/fig/1243099">10</figureCitation>
). All collecting events are from mixed coniferous forests (including
<taxonomicName id="8F1E4462729A1C0B13D3B06622CFE7C7" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" kingdom="Plantae" order="Pinales" phylum="Tracheophyta" rank="species" species="menziesii">
<emphasis id="6AD79AF9A7AB33D43F686B65E80C594E" italics="true">Pseudotsuga menziesii</emphasis>
</taxonomicName>
,
<taxonomicName id="877B33C5417C6499292A559117031A40" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="DB672B9266AAE3B49494A8C00123CC4E" italics="true">Pinus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="3CA5E22BB23372A830B7AC1BE7858AAE" class="Magnoliopsida" family="Ericaceae" genus="Arbutus" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="species" species="menziesii">
<emphasis id="F06B8F0E255F94A68D86719EF38C8FD6" italics="true">Arbutus menziesii</emphasis>
</taxonomicName>
) at mid-elevations (
<quantity id="DE2920C918B193956C50792279680BE3" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="8.0" metricValueMin="5.5" unit="m" value="675.0" valueMax="800.0" valueMin="550.0">550800 m</quantity>
) in the Klamath-Siskiyou Mountains. We anticipate additional populations in intervening and neighboring locations.
</paragraph>
<paragraph id="4FBD8CB0D5A5E2960651D6558A74DD99">
Spiders found abundantly beneath rocks in shaded rock piles, often under rocks lying directly on the surface, versus deeper in rock piles. Spiders were generally found without associated webs, although some subadult specimens found along Beaver Creek at Fishtrap Creek (
<collectionCode id="D1859BFA2A87E0BC4F15BDF0EC93CB3A">MCH</collectionCode>
24 _ 057) were found near sparse webbing (Fig.
<figureCitation id="A7E789A97F3E42E1FE4903A68E5E1C58" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Images of live cybaeid spiders. A Neocryphoeca beattyi ♀, Pinal County, AZ, RWM 22 _ 011 B Blabomma sp ♀, San Diego County, CA, MCH 22 _ 020 C Siskiyu armilla sp. nov. immature ♀, Siskiyou County CA, MCH 24 _ 057; patellar fracture line labeled D Siskiyu armilla sp. nov. immature ♂, Siskiyou County CA, MCH 24 _ 057; with silk webbing presumed to belong to the spider E Cybaeota shastae ♀ Chamberlin &amp; Ivie, 1937, East Rosebud Creek, MT F Cybaeus somesbar ♀ Bennett, 2009, Shasta County, CA, MCH 24 _ 012. All images taken by M Hedin, except for Neocryphoeca (RW Mendez)." figureDoi="10.3897/zookeys.1226.140204.figure1" httpUri="https://binary.pensoft.net/fig/1243090">1 D</figureCitation>
). Adults have been collected in April, July, and August; only subadults were found during the July collecting event at Fishtrap Creek (
<collectionCode id="7E00D1922AB3D0EF42F13F8E95F05A71">MCH</collectionCode>
24 _ 057).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ES5BG" type="comments">
<paragraph id="38312683376B56DB52A3BA92DB8F7F58">
<heading id="3079A02DEFB347FC5666C41B17572A1F" reason="title">Comments.</heading>
</paragraph>
<paragraph id="DF9BEAA062F604A949E4D015726A799E">
Discovery of this distinctive taxon adds to our knowledge of the highly diverse flora and fauna of the Klamath-Siskiyou Mountains (
<bibRefCitation id="5241A070B60E919C1A18E2E38F2262F0" author="Kauffmann and Garwood" firstAuthor="Kauffmann" refId="B31" refString="Kauffmann M, Garwood J (2022) The Klamath Mountains: A Natural History. Backcountry Press, Humboldt County, CA." year="2022">Kauffmann and Garwood 2022</bibRefCitation>
). Other globally rare and endemic spider taxa, reflecting also the high phylogenetic diversity of the region, include
<taxonomicName id="0C1D5246E5B1B823807282673F6A18F6" authority="Platnick, 1987" authorityName="Platnick" authorityYear="1987" class="Arachnida" family="Hypochilidae" genus="Hypochilus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="kastoni">
<emphasis id="75456C2183C0F09D5FFB2EECAC15D3B2" italics="true">Hypochilus kastoni</emphasis>
Platnick, 1987
</taxonomicName>
,
<taxonomicName id="077B0793FE514AE2193FB116899F0592" authority="Griswold, Audisio &amp; Ledford, 2012" authorityName="Griswold, Audisio &amp; Ledford" authorityYear="2012" class="Arachnida" family="Trogloraptoridae" genus="Trogloraptor" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="marchingtoni">
<emphasis id="9D42387069E0456E36AD8D7A2A55DF0F" italics="true">Trogloraptor marchingtoni</emphasis>
Griswold, Audisio &amp; Ledford, 2012
</taxonomicName>
, and
<taxonomicName id="480FA9C78E2D8668B5709BB012A1385D" authority="(Chamberlin &amp; Ivie, 1942)" baseAuthorityName="Chamberlin &amp; Ivie" baseAuthorityYear="1942" class="Arachnida" family="Leptonetidae" genus="Calileptoneta" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="sylva">
<emphasis id="DD0A1FC9B4B62F8D592DDE25B56E8BA1" italics="true">Calileptoneta sylva</emphasis>
(Chamberlin &amp; Ivie, 1942)
</taxonomicName>
. Additional efforts are needed to fully understand the arthropod biodiversity of this unique region.
</paragraph>
</subSubSection>
</treatment>
</document>