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<mods:title id="9613F3F7C872E59B42E809427B1E72B0">Hidden goby diversity of the Western Indian Ocean region: Glossogobius laticeps (De Vis, 1884) (Teleostei: Gobiidae)</mods:title>
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<mods:namePart id="591916267203A416C6F73C7B87C7F0AD">Zarei, Fatah</mods:namePart>
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<mods:namePart id="F62A10CD8BB9A7AB120C2F8EB4814D4D">Esmaeili, Hamid Reza</mods:namePart>
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<mods:affiliation id="21013FFA7E048B10F831B21F055D2351">Ichthyology and Molecular Systematics Research Laboratory, Department of Biology, School of Science, Shiraz University, Shiraz, Iran</mods:affiliation>
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<mods:affiliation id="47B6D36FF5E14C575AABCC7AE6980B23">Department of Biology, Faculty of Sciences, Lorestan University, Khorramabad, Iran</mods:affiliation>
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<mods:nameIdentifier id="39BC63914497675782D2139FEF1E77B4" type="email">g.sayyadzadeh2021@gmail.com</mods:nameIdentifier>
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<mods:nameIdentifier id="656BD68DAC4E4CF6EEFCA1B820A41A95" type="email">amirhassanmasoomi@gmail.com</mods:nameIdentifier>
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<mods:date id="CB9E6C27F765BD0221338868B1E4F719">2025</mods:date>
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<taxonomicName id="39E10B8BA727667BFF14FF29F068FF6C" authority="(De Vis, 1884)" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[151,592,223,250]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="585" phylum="Chordata" rank="species" species="laticeps">
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<emphasis id="CC95AC1AA727667BFF14FF29F3A3FF6F" bold="true" box="[151,411,223,250]" italics="true" pageId="4" pageNumber="585">Glossogobius laticeps</emphasis>
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(
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<bibRefCitation id="9A700DF9A727667BFE28FF29F070FF6C" author="De Vis, C. W." box="[427,584,223,249]" pageId="4" pageNumber="585" pagination="685 - 698" refId="ref6113" refString="De Vis, C. W. (1884) New fishes in the Queensland Museum. No. 4. Proceedings of the Linnean Society of New South Wales, Series 1, 9 (3), 685-698." type="journal article" year="1884">De Vis, 1884</bibRefCitation>
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)
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<taxonomicName id="39E10B8BA727667BFF14FEBEF3C3FECA" authority="De Vis, 1884: 692" authorityName="De Vis" authorityPageNumber="692" authorityYear="1884" box="[151,507,327,352]" class="Actinopterygii" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="585" phylum="Chordata" rank="species" species="laticeps">
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<emphasis id="CC95AC1AA727667BFF14FEBEF305FEF5" box="[151,317,328,352]" italics="true" pageId="4" pageNumber="585">Eleotris laticeps</emphasis>
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<bibRefCitation id="9A700DF9A727667BFEC0FEBEF3C3FECA" author="De Vis, C. W." box="[323,507,327,352]" pageId="4" pageNumber="585" pagination="685 - 698" refId="ref6113" refString="De Vis, C. W. (1884) New fishes in the Queensland Museum. No. 4. Proceedings of the Linnean Society of New South Wales, Series 1, 9 (3), 685-698." type="journal article" year="1884">De Vis, 1884: 692</bibRefCitation>
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(
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<collectingRegion id="3C25BEEAA727667BFD8BFEBEF0B9FEF5" box="[520,641,328,352]" country="Australia" name="Queensland" pageId="4" pageNumber="585">Queensland</collectingRegion>
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coast).
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</paragraph>
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<emphasis id="CC95AC1AA727667BFF14FE90F359FEEB" box="[151,353,358,382]" italics="true" pageId="4" pageNumber="585">Glossogobius giuris</emphasis>
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:
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<bibRefCitation id="9A700DF9A727667BFEE8FE90F065FEEB" author="Akihito, P. & Meguro, K." box="[363,605,358,383]" pageId="4" pageNumber="585" pagination="127 - 142" refId="ref5784" refString="Akihito, P. & Meguro, K. (1975) Description of a new gobiid fish, Glossogobius aureus, with notes on related species of the genus. Japanese Journal of Ichthyology, 22, 127-142. https://doi.org/10.11369/JJI1950.22.127" type="journal article" year="1975">Akihito & Meguro 1975</bibRefCitation>
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(in part);
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Allen
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<emphasis id="CC95AC1AA727667BFD7BFE91F11BFEEB" box="[760,803,358,382]" italics="true" pageId="4" pageNumber="585">et al</emphasis>
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. 2002
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</bibRefCitation>
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(in part).
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Pusey
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<emphasis id="CC95AC1AA727667BFB82FE91F614FEEB" box="[1025,1068,358,382]" italics="true" pageId="4" pageNumber="585">et al</emphasis>
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. 2004: 440
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</bibRefCitation>
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</treatmentCitation>
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(north-eastern
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<collectingCountry id="86F63098A727667BFAB3FE90F7AAFEEB" box="[1328,1426,358,382]" name="Australia" pageId="4" pageNumber="585">Australia</collectingCountry>
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);
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<bibRefCitation id="9A700DF9A727667BFF44FE73F3A4FE08" author="Pusey, B. J. & Burrows, D. W. & Kennard, M. J. & Perna, C. N. & Unmack, P. J. & Allsop, Q. & Hammer, M. P." box="[199,412,389,413]" pageId="4" pageNumber="585" pagination="1 - 104" refId="ref7024" refString="Pusey, B. J., Burrows, D. W., Kennard, M. J., Perna, C. N., Unmack, P. J., Allsop, Q. & Hammer, M. P. (2017) Freshwater fishes of northern Australia. Zootaxa, 4253 (1), 1-104. https://doi.org/10.11646/zootaxa.4253.1.1" type="journal article" year="2017">
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Pusey
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<emphasis id="CC95AC1AA727667BFE89FE70F30FFE08" box="[266,311,389,413]" italics="true" pageId="4" pageNumber="585">et al</emphasis>
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. 2017: 73
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</bibRefCitation>
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</treatmentCitation>
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(in part).
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<emphasis id="CC95AC1AA727667BFF14FC24F32DFC7F" bold="true" box="[151,277,978,1002]" pageId="4" pageNumber="585">FIGURE 3.</emphasis>
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Live specimen of
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<taxonomicName id="39E10B8BA727667BFE5AFC25F083FC7E" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[473,699,979,1003]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="585" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA727667BFE5AFC25F083FC7E" box="[473,699,979,1003]" italics="true" pageId="4" pageNumber="585">Glossogobius laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
(ZM-CBSU GL1, 73.6 mm SL) from the Gabrik River, Makran basin, southern Iran.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="B6FB2383A727667BFF14FB97F0BFFB7D" pageId="4" pageNumber="585" type="materials_examined">
|
||||
<paragraph id="FE5E7008A727667BFF14FB97F0BFFB7D" blockId="4.[151,1437,1121,2011]" pageId="4" pageNumber="585">
|
||||
<emphasis id="CC95AC1AA727667BFF14FB97F3BEFBEE" bold="true" box="[151,390,1121,1147]" pageId="4" pageNumber="585">Material examined.</emphasis>
|
||||
<materialsCitation id="4E897A55A727667BFE11FB97F0BBFB7D" collectingDate="2023-11-29" collectorName="H. R. Esmaeili & S. H. Hashemi & A. H. Masoumi & S. Echreshavi & Karami & Abbasi" country="Iran" latitude="25.772018" location="Gabrik River" longLatPrecision="1" longitude="58.46697" municipality="Makran" pageId="4" pageNumber="585" specimenCode="ZM-CBSU GL1" specimenCount="10" stateProvince="Hormuzgan Province">
|
||||
<specimenCode id="AE47D873A727667BFE11FB97F075FBEE" box="[402,589,1121,1147]" pageId="4" pageNumber="585">ZM-CBSU GL1</specimenCode>
|
||||
,
|
||||
<specimenCount id="E8E7BB81A727667BFDE3FB97F138FBEE" box="[608,768,1121,1147]" count="10" pageId="4" pageNumber="585" type="generic">10 specimens</specimenCount>
|
||||
, 36.0–
|
||||
<quantity id="3919DDEDA727667BFCCCFB97F180FBEE" box="[847,952,1121,1147]" metricMagnitude="-2" metricUnit="m" metricValue="7.359999999999999" pageId="4" pageNumber="585" unit="mm" value="73.6">73.6 mm</quantity>
|
||||
SL,
|
||||
<location id="FB3E26D3A727667BFC7BFB97F6A9FBEE" LSID="urn:lsid:plazi:treatment:7648C11EA7276677FF14FF29F606FC78:FB3E26D3A727667BFC7BFB97F6A9FBEE" box="[1016,1169,1121,1147]" country="Iran" latitude="25.772018" longLatPrecision="1" longitude="58.46697" municipality="Makran" name="Gabrik River" pageId="4" pageNumber="585" stateProvince="Hormuzgan Province">Gabrik River</location>
|
||||
,
|
||||
<collectingRegion id="3C25BEEAA727667BFB22FB97F7A0FBEE" box="[1185,1432,1121,1147]" country="Iran" name="Hormozgan" pageId="4" pageNumber="585">Hormuzgan Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="1E3AEA72A727667BFF14FB73F2D6FB0A" box="[151,238,1157,1183]" pageId="4" pageNumber="585">Makran</collectingMunicipality>
|
||||
basin,
|
||||
<collectingCountry id="86F63098A727667BFEC2FB73F349FB0A" box="[321,369,1157,1183]" name="Iran" pageId="4" pageNumber="585">Iran</collectingCountry>
|
||||
,
|
||||
<geoCoordinate id="9BD516CFA727667BFEFEFB73F013FB35" box="[381,555,1157,1184]" degrees="25" direction="north" minutes="46" orientation="latitude" pageId="4" pageNumber="585" precision="1" seconds="19.27" value="25.772018">25°46’19.27”N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="9BD516CFA727667BFDBAFB73F0EEFB35" box="[569,726,1157,1184]" degrees="58" direction="east" minutes="28" orientation="longitude" pageId="4" pageNumber="585" precision="1" seconds="1.09" value="58.46697">58°28’1.09”E</geoCoordinate>
|
||||
, collected by
|
||||
<collectorName id="531415DEA727667BFCF7FB73F62EFB0A" box="[884,1046,1157,1183]" pageId="4" pageNumber="585">H.R. Esmaeili</collectorName>
|
||||
,
|
||||
<collectorName id="531415DEA727667BFBA0FB73F6FBFB0A" box="[1059,1219,1157,1183]" pageId="4" pageNumber="585">S.H. Hashemi</collectorName>
|
||||
,
|
||||
<collectorName id="531415DEA727667BFB4DFB73F742FB0A" box="[1230,1402,1157,1183]" pageId="4" pageNumber="585">A.H. Masoumi</collectorName>
|
||||
,
|
||||
<collectorName id="531415DEA727667BFA05FB73F32EFB56" pageId="4" pageNumber="585">S. Echreshavi</collectorName>
|
||||
,
|
||||
<collectorName id="531415DEA727667BFEA0FB5FF34FFB56" box="[291,375,1193,1219]" pageId="4" pageNumber="585">Karami</collectorName>
|
||||
and
|
||||
<collectorName id="531415DEA727667BFE2DFB5FF038FB56" box="[430,512,1193,1219]" pageId="4" pageNumber="585">Abbasi</collectorName>
|
||||
,
|
||||
<date id="8A5F56C8A727667BFD8EFB5FF0D0FB56" box="[525,744,1193,1219]" pageId="4" pageNumber="585" value="2023-11-29">
|
||||
<collectingDate id="9A1BAF20A727667BFD8EFB5FF0D0FB56" box="[525,744,1193,1219]" pageId="4" pageNumber="585" value="2023-11-29">29 November 2023</collectingDate>
|
||||
</date>
|
||||
(COI sequence IDs: M3735B, M3792B, M3893B; GenBank accession numbers:
|
||||
<accessionNumber id="E1B2EDEBA727667BFEFAFB3BF3CBFB7D" box="[377,499,1229,1256]" pageId="4" pageNumber="585">PQ778063</accessionNumber>
|
||||
–
|
||||
<accessionNumber id="E1B2EDEBA727667BFD81FB3BF043FB7D" box="[514,635,1229,1256]" pageId="4" pageNumber="585">PQ778065</accessionNumber>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="B6FB2383A727667BFF44FB07F32EFA2A" pageId="4" pageNumber="585" type="diagnosis">
|
||||
<paragraph id="FE5E7008A727667BFF44FB07F32EFA2A" blockId="4.[151,1437,1121,2011]" pageId="4" pageNumber="585">
|
||||
<emphasis id="CC95AC1AA727667BFF44FB07F079FA9E" box="[199,577,1265,1291]" italics="true" pageId="4" pageNumber="585">
|
||||
<emphasis id="CC95AC1AA727667BFF44FB07F379FA9E" bold="true" box="[199,321,1265,1291]" italics="true" pageId="4" pageNumber="585">Diagnosis.</emphasis>
|
||||
<taxonomicName id="39E10B8BA727667BFECAFB07F079FA9E" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[329,577,1265,1291]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="585" phylum="Chordata" rank="species" species="laticeps">Glossogobius laticeps</taxonomicName>
|
||||
</emphasis>
|
||||
can be distinguished from its Indian Ocean congeners (see Introduction) by the following combination of characters: (i) head and body pigmented and eyes present; (ii) predorsal area fully scaled to behind eyes, with a distinct naked patch usually present; (iii) suborbital papillae lines 9–10, papillae Line 13 on underside of head, and papillae Line 1a on snout all doubled or tripled; (iv) papillae Line 5 below eye and suborbital papillae Line 7 doubled; and (v) blotch on caudal fin base triangular, without a distinct unpigmented gap posteriorly.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="B6FB2383A7276679FF44FA3EF0D5FEF0" lastPageId="6" lastPageNumber="587" pageId="4" pageNumber="585" type="description">
|
||||
<paragraph id="FE5E7008A727667BFF44FA3EF7ADF8B2" blockId="4.[151,1437,1121,2011]" pageId="4" pageNumber="585">
|
||||
<emphasis id="CC95AC1AA727667BFF44FA3EF382FA76" bold="true" box="[199,442,1480,1507]" italics="true" pageId="4" pageNumber="585">General morphology.</emphasis>
|
||||
Body proportions detailed in
|
||||
<tableCitation id="B36345B3A727667BFC83FA3FF16BFA76" box="[768,851,1481,1507]" captionStart="TABLE 1" captionStartId="2.[151,239,1016,1040]" captionTargetPageId="2" captionText="TABLE 1. Morphometric data of Glossogobius laticeps from the Gabrik River, Makran basin, southern Iran (ZM-CBSU GL1, n=10)." pageId="4" pageNumber="585">Table 1</tableCitation>
|
||||
: body moderately elongate, depth at pelvic-fin origin
|
||||
<quantity id="3919DDEDA727667BFF14FA1BF32FF992" box="[151,279,1517,1544]" metricMagnitude="-1" metricUnit="m" metricValue="1.48844" metricValueMax="1.55448" metricValueMin="1.4223999999999999" pageId="4" pageNumber="585" unit="in" value="5.859999999999999" valueMax="6.12" valueMin="5.6">5.6–6.12 in</quantity>
|
||||
SL, at anal-fin origin
|
||||
<quantity id="3919DDEDA727667BFD90FA1BF099F992" box="[531,673,1517,1544]" metricMagnitude="-1" metricUnit="m" metricValue="1.5722599999999998" metricValueMax="1.6382999999999999" metricValueMin="1.50622" pageId="4" pageNumber="585" unit="in" value="6.1899999999999995" valueMax="6.45" valueMin="5.93">5.93–6.45 in</quantity>
|
||||
SL, laterally compressed posteriorly, with shallow caudal peduncle (depth 0.34–0.42 of length). Head large, length
|
||||
<quantity id="3919DDEDA727667BFD37F9E7F17AF9BE" box="[692,834,1553,1579]" metricMagnitude="-2" metricUnit="m" metricValue="7.2009" metricValueMax="7.5184" metricValueMin="6.8834" pageId="4" pageNumber="585" unit="in" value="2.835" valueMax="2.96" valueMin="2.71">2.71–2.96 in</quantity>
|
||||
SL, width
|
||||
<quantity id="3919DDEDA727667BFC42F9E7F60BF9BE" box="[961,1075,1553,1580]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.4731999999999998" metricValueMin="1.3208" pageId="4" pageNumber="585" unit="in" value="5.5" valueMax="5.8" valueMin="5.2">5.2–5.8 in</quantity>
|
||||
SL, and depth at eye
|
||||
<quantity id="3919DDEDA727667BFAAFF9E7F295F9DA" metricMagnitude="-1" metricUnit="m" metricValue="1.8948399999999999" metricValueMax="2.00914" metricValueMin="1.7805399999999998" pageId="4" pageNumber="585" unit="in" value="7.46" valueMax="7.91" valueMin="7.01">7.01–7.91 in</quantity>
|
||||
SL and 0.69–0.78 width. Postorbital profile sub-horizontal; snout long, length 1.65–1.98 of eye diameter and
|
||||
<quantity id="3919DDEDA727667BFF14F9AFF31BF9E6" box="[151,291,1625,1651]" metricMagnitude="-2" metricUnit="m" metricValue="7.0358" metricValueMax="7.4422" metricValueMin="6.6293999999999995" pageId="4" pageNumber="585" unit="in" value="2.77" valueMax="2.93" valueMin="2.61">2.61–2.93 in</quantity>
|
||||
head length. Anterior nostril at end of short tube, 2–3 nostril diameters from upper lip; posterior nostril as pore without raised rim, located before eye, closer to upper lip than to eye, and about 2 posterior nostril diameters from anterior nostril. Eyes dorsolateral and large, diameter
|
||||
<quantity id="3919DDEDA727667BFCC7F957F1EBF92E" box="[836,979,1697,1724]" metricMagnitude="-1" metricUnit="m" metricValue="1.30175" metricValueMax="1.37668" metricValueMin="1.22682" pageId="4" pageNumber="585" unit="in" value="5.125" valueMax="5.42" valueMin="4.83">4.83–5.42 in</quantity>
|
||||
head length, orbit elevated. Interorbital width
|
||||
<quantity id="3919DDEDA727667BFF5DF933F353F94A" box="[222,363,1733,1760]" metricMagnitude="-2" metricUnit="m" metricValue="3.2258" metricValueMax="3.5305999999999997" metricValueMin="2.921" pageId="4" pageNumber="585" unit="in" value="1.27" valueMax="1.39" valueMin="1.15">1.15–1.39 in</quantity>
|
||||
eye diameter. Mouth angled obliquely upwards; lower jaw significantly prognathous; jaw angle in front of pupil. Teeth in the upper and lower jaws conical, enlarged in the outermost and innermost rows; irregular rows of smaller teeth in between. Gill opening broad, extending just anterior to posterior margin of preoperculum.
|
||||
</paragraph>
|
||||
<paragraph id="FE5E7008A727667AFF44F8C7F395FE88" blockId="4.[151,1437,1121,2011]" lastBlockId="5.[151,1437,151,1043]" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585">
|
||||
<emphasis id="CC95AC1AA727667BFF44F8C7F33CF8DE" bold="true" box="[199,260,1841,1867]" italics="true" pageId="4" pageNumber="585">Fins.</emphasis>
|
||||
D1 VI; D2 I/8–I/9 (I/8:1, I/9:9); A I/8–I/9 (I/8:9, I/9:1); P 18–20 (18:1, 19:3, 20:6). First dorsal fin with rounded to triangular margin; second spine longest, with spines decreasing in size to sixth spine; fin origin well behind pelvic insertion. Second dorsal fin depth equal or sub-equal to first dorsal fin; posterior rays not elongated, reaching about halfway to caudal fin when appressed; fin origin slightly in front of vertical to anal fin origin. Anal fin slightly lower than second dorsal fin; posterior rays elongated like those of second dorsal fin; last ray origin below last ray origin of second dorsal fin. Pectoral fin with rounded margin, not reaching vertical of second dorsal fin origin; all rays branched, uppermost rays not free from membrane. Pelvic disc complete, much longer than wide, ending before or reaching anus; anterior membrane present, without lateral lobes. Caudal fin with rounded margin, shorter than head length.
|
||||
</paragraph>
|
||||
<paragraph id="FE5E7008A726667AFF44FED1F1F7FE60" blockId="5.[151,1437,151,1043]" pageId="5" pageNumber="586">
|
||||
<emphasis id="CC95AC1AA726667AFF44FED1F362FED4" bold="true" box="[199,346,295,321]" italics="true" pageId="5" pageNumber="586">Squamation.</emphasis>
|
||||
Cheek naked; predorsal area fully scaled with cycloid scales but a naked patch behind postorbital pore E present. Operculum with small patch of cycloid scales; occasionally reaches mid-operculum in 1–3 vertical rows. Scale ctenii weakly developed or absent above operculum. Prepelvic area covered with cycloid scales, ending behind posterior preopercular margin; isthmus often naked. Pectoral fin base partly scaled. Abdomen covered in cycloid scales; body covered in ctenoid scales. Caudal peduncle with slightly enlarged scales. LL 30–32 (30:2, 31:4, 32:4); PD 19–25 (19:1, 21:3, 22:2, 24:3, 25:1); TR 9–11 (9:1, 10:5, 11:4).
|
||||
</paragraph>
|
||||
<paragraph id="FE5E7008A726667AFF44FE08F640FD8C" blockId="5.[151,1437,151,1043]" box="[199,1144,510,537]" pageId="5" pageNumber="586">
|
||||
<emphasis id="CC95AC1AA726667AFF44FE08F3A9FD8C" bold="true" box="[199,401,510,537]" pageId="5" pageNumber="586">
|
||||
<emphasis id="CC95AC1AA726667AFF44FE08F37BFD8C" bold="true" box="[199,323,510,537]" italics="true" pageId="5" pageNumber="586">Gill rakers</emphasis>
|
||||
(n=5).
|
||||
</emphasis>
|
||||
Gill rakers on outer face of first arch: 1–2+1+6–8, usually 1+1+8.
|
||||
</paragraph>
|
||||
<paragraph id="FE5E7008A726667AFF44FDD4F286FB86" blockId="5.[151,1437,151,1043]" pageId="5" pageNumber="586">
|
||||
<emphasis id="CC95AC1AA726667AFF44FDD4F3DFFDA8" bold="true" box="[199,487,546,573]" italics="true" pageId="5" pageNumber="586">Head lateral line system.</emphasis>
|
||||
Refer to
|
||||
<figureCitation id="66DA6C8DA726667AFDD6FDD5F08CFDA8" box="[597,692,547,573]" captionStart="FIGURE 4" captionStartId="5.[151,250,1687,1711]" captionTargetBox="[249,1338,1070,1653]" captionTargetId="figure-450@5.[244,1343,1062,1661]" captionTargetPageId="5" captionText="FIGURE 4. Cephalic sensory pores system and papillae of Glossogobius laticeps (ZM-CBSU GL1, 73.6 mm SL) from the Gabrik River, Makran basin, southern Iran. Terminology is described in the text." figureDoi="http://doi.org/10.5281/zenodo.14853706" httpUri="https://zenodo.org/record/14853706/files/figure.png" pageId="5" pageNumber="586">Figure 4</figureCitation>
|
||||
for head sensory pores and papillae pattern. Head pores: anterior oculoscapular canal pores: B′, C, D, E, F, G, and H′; posterior oculoscapular canal pores: K′ and L′; preopercular canal pores: M′, N, and O′. All are paired except for C and D. Sensory papillae: Line 1a: double row starting from pore B′; Line 1b: 1 papilla above anterior nostril; Line 2: single row, continuous across snout; Line 5: below eye, doubled anteriorly, single posteriorly; Line 6: long, anteriorly joins Line 5 behind mid-eye; five suborbital longitudinal lines: 7–11. Line 7: doubled anteriorly, single row posteriorly; axis along papilla line, larger anteriorly. Line 8: larger oval papillae, axis vertical to papilla line. Lines 9 and 10: 3 rows of papillae, axis along papilla line. Line 11: single row, oval papillae, axis vertical to papilla line. Line 12: underside of head, single row, continuous. Line 13: underside of head, doubled and continuous. Line 20: transversal single row of papillae on operculum. Line 21: longitudinal and curved, single row. Line 22: short longitudinal, single row.
|
||||
<collectingRegion id="3C25BEEAA726667AFBD0FC91F6B1FC14" box="[1107,1161,871,897]" country="Myanmar" name="Chin" pageId="5" pageNumber="586">Chin</collectingRegion>
|
||||
partly covered with two clusters of papillae; mental frenum free of papillae. Line 14: long single row behind eye. Line 15: single row from eye to pore K′. Line 17: single row from pore H′ to pore K′. Lines 16 and 18: short single rows, parallel, behind pore L′. Line 19: single transversal row, starting dorsally between pores G and H′, extending downward to just above pore M′.
|
||||
</paragraph>
|
||||
<caption id="AA9E2080A726667AFF14F961F1FEF947" ID-DOI="http://doi.org/10.5281/zenodo.14853706" ID-Zenodo-Dep="14853706" httpUri="https://zenodo.org/record/14853706/files/figure.png" pageId="5" pageNumber="586" startId="5.[151,250,1687,1711]" targetBox="[249,1338,1070,1653]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="FE5E7008A726667AFF14F961F1FEF947" blockId="5.[151,1436,1687,1747]" pageId="5" pageNumber="586">
|
||||
<emphasis id="CC95AC1AA726667AFF14F961F32DF93A" bold="true" box="[151,277,1687,1711]" pageId="5" pageNumber="586">FIGURE 4.</emphasis>
|
||||
Cephalic sensory pores system and papillae of
|
||||
<taxonomicName id="39E10B8BA726667AFC8BF961F1D1F93A" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[776,1001,1687,1711]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="586" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA726667AFC8BF961F1D1F93A" box="[776,1001,1687,1711]" italics="true" pageId="5" pageNumber="586">Glossogobius laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
(ZM-CBSU GL1, 73.6 mm SL) from the Gabrik River, Makran basin, southern Iran. Terminology is described in the text.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="FE5E7008A7266679FF44F8D3F1D8FE88" blockId="5.[151,1437,1829,2036]" lastBlockId="6.[151,1436,151,645]" lastPageId="6" lastPageNumber="587" pageId="5" pageNumber="586">
|
||||
<emphasis id="CC95AC1AA726667AFF44F8D3F39EF8D5" bold="true" box="[199,422,1829,1856]" italics="true" pageId="5" pageNumber="586">Coloration. In life.</emphasis>
|
||||
The typical color pattern of adult specimens as described by
|
||||
<bibRefCitation id="9A700DF9A726667AFBF5F8D0F7A4F8AA" author="Hoese, D. F. & Hammer, M. P." box="[1142,1436,1829,1856]" pageId="5" pageNumber="586" pagination="79 - 115" refId="ref6759" refString="Hoese, D. F. & Hammer, M. P. (2021) A review of the Glossogobius giuris complex in Australia, with wider discussion on nomenclature and possible synonymies. Zootaxa, 4974 (1), 79-115. https://doi.org/10.11646/zootaxa.4974.1.3" type="journal article" year="2021">Hoese & Hammer (2021)</bibRefCitation>
|
||||
is not present in the collected specimens from the Gabrik River. Our specimens, which range in standard length from 36.0 to
|
||||
<quantity id="3919DDEDA726667AFEADF89BF3ADF81D" box="[302,405,1901,1928]" metricMagnitude="-2" metricUnit="m" metricValue="7.359999999999999" pageId="5" pageNumber="586" unit="mm" value="73.6">73.6 mm</quantity>
|
||||
, represent juvenile, young and sub-adult stages: their coloration cannot reflect the patterns exhibited by mature individuals (
|
||||
<figureCitation id="66DA6C8DA726667AFD8AF867F076F83E" box="[521,590,1937,1963]" captionStart="FIGURE 3" captionStartId="4.[151,250,978,1002]" captionTargetBox="[151,1435,435,953]" captionTargetId="figure-77@4.[151,1435,435,953]" captionTargetPageId="4" captionText="FIGURE 3. Live specimen of Glossogobius laticeps (ZM-CBSU GL1, 73.6 mm SL) from the Gabrik River, Makran basin, southern Iran." figureDoi="http://doi.org/10.5281/zenodo.14853704" httpUri="https://zenodo.org/record/14853704/files/figure.png" pageId="5" pageNumber="586">Fig. 3</figureCitation>
|
||||
). The head and body exhibit a white and unpigmented coloration. A weakly developed dark brown bar extends from the anteroventral margin of the eye to the middle of the lips. The body lacks dorsal mottling and brown horizontal stripes above and below the midline. Along the midside, there are five weakly developed horizontally elongated spots, with the rearmost spot on the caudal fin base being the darkest and nearly square in shape. The dorsal, pectoral, pelvic, and anal fin spines, rays, and membranes are all unpigmented, while the rays of the caudal fin display minute dark spots.
|
||||
<emphasis id="CC95AC1AA7256679FD75FF29F192FF6C" bold="true" box="[758,938,223,249]" italics="true" pageId="6" pageNumber="587">In preservative.</emphasis>
|
||||
The head and body are pale white, lacking pigmentation, with a distinct dark spot located at the base of the caudal fin.
|
||||
</paragraph>
|
||||
<paragraph id="FE5E7008A7256679FF44FED0F0D5FEF0" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="587">
|
||||
<emphasis id="CC95AC1AA7256679FF44FED0F38AFED4" bold="true" box="[199,434,294,321]" italics="true" pageId="6" pageNumber="587">Sexual dimorphism.</emphasis>
|
||||
Males have a conical urogenital papilla with a pointed posterior edge; females have a wider, trapezoid papilla with a villous posterior edge.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="B6FB2383A7256679FF44FE98F672FD10" pageId="6" pageNumber="587" type="distribution">
|
||||
<paragraph id="FE5E7008A7256679FF44FE98F672FD10" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="587">
|
||||
<emphasis id="CC95AC1AA7256679FF44FE98F3DAFE1C" bold="true" box="[199,482,366,393]" italics="true" pageId="6" pageNumber="587">Distribution and habitat.</emphasis>
|
||||
Prior to this study,
|
||||
<taxonomicName id="39E10B8BA7256679FD3BFE99F195FE1C" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[696,941,367,393]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA7256679FD3BFE99F195FE1C" box="[696,941,367,393]" italics="true" pageId="6" pageNumber="587">Glossogobius laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
was documented solely in
|
||||
<collectingRegion id="3C25BEEAA7256679FB54FE99F726FE1C" box="[1239,1310,367,393]" country="Indonesia" name="Papua Barat" pageId="6" pageNumber="587">Papua</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FAABFE99F7A0FE1C" box="[1320,1432,367,393]" name="Indonesia" pageId="6" pageNumber="587">Indonesia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FF14FE65F346FE38" box="[151,382,403,429]" name="Papua New Guinea" pageId="6" pageNumber="587">Papua New Guinea</collectingCountry>
|
||||
, the
|
||||
<collectingRegion id="3C25BEEAA7256679FE3DFE65F0AAFE38" box="[446,658,403,429]" country="Australia" name="Northern Territory" pageId="6" pageNumber="587">Northern Territory</collectingRegion>
|
||||
in
|
||||
<collectingCountry id="86F63098A7256679FD43FE65F112FE38" box="[704,810,403,429]" name="Australia" pageId="6" pageNumber="587">Australia</collectingCountry>
|
||||
, and along the
|
||||
<collectingRegion id="3C25BEEAA7256679FC69FE65F656FE38" box="[1002,1134,403,429]" country="Australia" name="Queensland" pageId="6" pageNumber="587">Queensland</collectingRegion>
|
||||
coast from Cooktown to Bundaberg, as well as in
|
||||
<collectingCountry id="86F63098A7256679FE30FE41F3C1FE44" box="[435,505,439,465]" name="China" pageId="6" pageNumber="587">China</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FD87FE41F05EFE44" box="[516,614,439,465]" name="Vietnam" pageId="6" pageNumber="587">Vietnam</collectingCountry>
|
||||
, and
|
||||
<collectingCountry id="86F63098A7256679FD22FE41F11CFE44" box="[673,804,439,465]" name="Bangladesh" pageId="6" pageNumber="587">Bangladesh</collectingCountry>
|
||||
(
|
||||
<figureCitation id="66DA6C8DA7256679FCB0FE41F14DFE44" box="[819,885,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1813,1837]" captionTargetBox="[151,1436,1026,1786]" captionTargetId="figure-377@1.[151,1436,1026,1786]" captionTargetPageId="1" captionText="FIGURE 1. Updated geographical distributions of the four species/clades within the Glossogobius giuris species complex: G. giuris ‘clade B’, G. giuris ‘clade C’, G. laticeps, and G. tenuiformis. These distributions are based on the COI dataset from Hammer et al. (2021), Zarei et al. (2023b), and the present study. The new locality record of G. laticeps from the Gabrik River in the Makran basin, southern Iran, is indicated with a yellow square." figureDoi="http://doi.org/10.5281/zenodo.14853700" httpUri="https://zenodo.org/record/14853700/files/figure.png" pageId="6" pageNumber="587">Fig. 1</figureCitation>
|
||||
). Adults are typically associated with freshwater habitats, while juveniles are occasionally found in estuarine environments (
|
||||
<bibRefCitation id="9A700DF9A7256679FC61FE2DF6C8FE60" author="Hoese, D. F. & Hammer, M. P." box="[994,1264,475,501]" pageId="6" pageNumber="587" pagination="79 - 115" refId="ref6759" refString="Hoese, D. F. & Hammer, M. P. (2021) A review of the Glossogobius giuris complex in Australia, with wider discussion on nomenclature and possible synonymies. Zootaxa, 4974 (1), 79-115. https://doi.org/10.11646/zootaxa.4974.1.3" type="journal article" year="2021">Hoese & Hammer 2021</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="9A700DF9A7256679FB7FFE2DF2EBFD8C" author="Hammer, M. P. & Taillebois, L. & King, A. J. & Crook, D. A. & Wedd, D. & Adams, M. & Unmack, P. J. & Hoese, D. F. & Bertozzi, T." pageId="6" pageNumber="587" pagination="87 - 100" refId="ref6541" refString="Hammer, M. P., Taillebois, L., King, A. J., Crook, D. A., Wedd, D., Adams, M., Unmack, P. J., Hoese, D. F. & Bertozzi, T. (2021) Unravelling the taxonomy and identification of a problematic group of benthic fishes from tropical rivers (Gobiidae: Glossogobius). Journal of Fish Biology, 99 (1), 87-100. https://doi.org/10.1111/jfb.14701" type="journal article" year="2021">
|
||||
Hammer
|
||||
<emphasis id="CC95AC1AA7256679FAE7FE2DF7ADFE60" box="[1380,1429,475,501]" italics="true" pageId="6" pageNumber="587">et al</emphasis>
|
||||
. 2021
|
||||
</bibRefCitation>
|
||||
). The current record from the Gabrik River in the Makran basin of southern
|
||||
<collectingCountry id="86F63098A7256679FBA0FE09F668FD8C" box="[1059,1104,511,537]" name="Iran" pageId="6" pageNumber="587">Iran</collectingCountry>
|
||||
represents a significant range expansion into the Western Indian Ocean. Most streams in the Makran basin, including the Gabrik River (
|
||||
<figureCitation id="66DA6C8DA7256679FACBFDD5F7B5FDA8" box="[1352,1421,546,573]" captionStart="FIGURE 5" captionStartId="6.[151,250,1525,1549]" captionTargetBox="[238,1349,666,1499]" captionTargetId="figure-264@6.[238,1349,666,1499]" captionTargetPageId="6" captionText="FIGURE 5. Gabrik River, habitat of Glossogobius laticeps in Makran basin, southern Iran." figureDoi="http://doi.org/10.5281/zenodo.14853708" httpUri="https://zenodo.org/record/14853708/files/figure.png" pageId="6" pageNumber="587">Fig. 5</figureCitation>
|
||||
), are characterized by minimal flow, often reducing to isolated pools separated by distances of one kilometer or more, and they frequently experience desiccation along substantial portions of their length.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="AA9E2080A7256679FF14FA03F60DF998" ID-DOI="http://doi.org/10.5281/zenodo.14853708" ID-Zenodo-Dep="14853708" box="[151,1077,1525,1550]" httpUri="https://zenodo.org/record/14853708/files/figure.png" pageId="6" pageNumber="587" startId="6.[151,250,1525,1549]" targetBox="[238,1349,666,1499]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="FE5E7008A7256679FF14FA03F60DF998" blockId="6.[151,1077,1525,1550]" box="[151,1077,1525,1550]" pageId="6" pageNumber="587">
|
||||
<emphasis id="CC95AC1AA7256679FF14FA03F32BF99B" bold="true" box="[151,275,1525,1550]" pageId="6" pageNumber="587">FIGURE 5.</emphasis>
|
||||
Gabrik River, habitat of
|
||||
<taxonomicName id="39E10B8BA7256679FD92FA00F0C8F99B" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[529,752,1526,1550]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA7256679FD92FA00F0C8F99B" box="[529,752,1526,1550]" italics="true" pageId="6" pageNumber="587">Glossogobius laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
in Makran basin, southern Iran.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="B6FB2383A7256677FF44F996F606FC78" lastPageId="8" lastPageNumber="589" pageId="6" pageNumber="587" type="discussion">
|
||||
<paragraph id="FE5E7008A7256678FF44F996F1C8FEF0" blockId="6.[151,1437,1632,2018]" lastBlockId="7.[151,1437,151,357]" lastPageId="7" lastPageNumber="588" pageId="6" pageNumber="587">
|
||||
<emphasis id="CC95AC1AA7256679FF44F996F37DF9EF" bold="true" box="[199,325,1632,1658]" pageId="6" pageNumber="587">Discussion</emphasis>
|
||||
Phylogenetic analysis of mitochondrial COI by
|
||||
<bibRefCitation id="9A700DF9A7256679FCE6F996F660F9EF" author="Hammer, M. P. & Taillebois, L. & King, A. J. & Crook, D. A. & Wedd, D. & Adams, M. & Unmack, P. J. & Hoese, D. F. & Bertozzi, T." box="[869,1112,1632,1658]" pageId="6" pageNumber="587" pagination="87 - 100" refId="ref6541" refString="Hammer, M. P., Taillebois, L., King, A. J., Crook, D. A., Wedd, D., Adams, M., Unmack, P. J., Hoese, D. F. & Bertozzi, T. (2021) Unravelling the taxonomy and identification of a problematic group of benthic fishes from tropical rivers (Gobiidae: Glossogobius). Journal of Fish Biology, 99 (1), 87-100. https://doi.org/10.1111/jfb.14701" type="journal article" year="2021">
|
||||
Hammer
|
||||
<emphasis id="CC95AC1AA7256679FC4DF997F1C7F9EF" box="[974,1023,1632,1658]" italics="true" pageId="6" pageNumber="587">et al</emphasis>
|
||||
. (2021)
|
||||
</bibRefCitation>
|
||||
identified four clades within
|
||||
<taxonomicName id="39E10B8BA7256679FF14F972F341F90B" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[151,377,1668,1694]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA7256679FF14F972F341F90B" box="[151,377,1668,1694]" italics="true" pageId="6" pageNumber="587">Glossogobius giuris</emphasis>
|
||||
</taxonomicName>
|
||||
, indicating cryptic species: (i) clade A was reclassified as
|
||||
<taxonomicName id="39E10B8BA7256679FB96F972F6A8F90B" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[1045,1168,1668,1694]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA7256679FB96F972F6A8F90B" box="[1045,1168,1668,1694]" italics="true" pageId="6" pageNumber="587">G. laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
, recorded in
|
||||
<collectingCountry id="86F63098A7256679FAABF972F7A0F90B" box="[1320,1432,1668,1694]" name="Indonesia" pageId="6" pageNumber="587">Indonesia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FF14F95EF339F957" box="[151,257,1704,1730]" name="Australia" pageId="6" pageNumber="587">Australia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FE8CF95EF36DF957" box="[271,341,1704,1730]" name="China" pageId="6" pageNumber="587">China</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FEE0F95EF3FDF957" box="[355,453,1704,1730]" name="Vietnam" pageId="6" pageNumber="587">Vietnam</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FE50F95EF061F957" box="[467,601,1704,1730]" name="Bangladesh" pageId="6" pageNumber="587">Bangladesh</collectingCountry>
|
||||
, and now the Gabrik River, Makran Basin, southern
|
||||
<collectingCountry id="86F63098A7256679FB46F95EF6CDF957" box="[1221,1269,1704,1730]" name="Iran" pageId="6" pageNumber="587">Iran</collectingCountry>
|
||||
, adding to the country’s freshwater ichthyofauna; (ii) clade B, found in
|
||||
<collectingCountry id="86F63098A7256679FC94F93AF1B9F973" box="[791,897,1740,1766]" name="Australia" pageId="6" pageNumber="587">Australia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FC0FF93AF1F0F973" box="[908,968,1740,1766]" name="India" pageId="6" pageNumber="587">India</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FC57F93AF65BF973" box="[980,1123,1740,1766]" name="South Africa" pageId="6" pageNumber="587">South Africa</collectingCountry>
|
||||
(
|
||||
<figureCitation id="66DA6C8DA7256679FBF1F93AF6F4F973" box="[1138,1228,1740,1766]" captionStart="FIGURE 6" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[151,1436,378,1908]" captionTargetId="figure-122@7.[151,1436,378,1908]" captionTargetPageId="7" captionText="FIGURE 6. Uncatalogued specimens of Glossogobius giuris ‘clade B’ from (A) Mlalazi River, South Africa, and (B) Sarbaz River, Makran basin, southeastern Iran, and G. tenuiformis from (C) South Africa and (D) Wadi Hasik, Oman. Photos by J. Coetzee (A), A. Jouladeh Roudbar (B), O. Gon (C), and H.R. Esmaeili (D)." figureDoi="http://doi.org/10.5281/zenodo.14853710" httpUri="https://zenodo.org/record/14853710/files/figure.png" pageId="6" pageNumber="587">Fig. 6A</figureCitation>
|
||||
), and southeastern
|
||||
<collectingCountry id="86F63098A7256679FF14F906F2FCF89F" box="[151,196,1776,1802]" name="Iran" pageId="6" pageNumber="587">Iran</collectingCountry>
|
||||
(Sarbaz River;
|
||||
<figureCitation id="66DA6C8DA7256679FE03F906F3E0F89F" box="[384,472,1776,1802]" captionStart="FIGURE 6" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[151,1436,378,1908]" captionTargetId="figure-122@7.[151,1436,378,1908]" captionTargetPageId="7" captionText="FIGURE 6. Uncatalogued specimens of Glossogobius giuris ‘clade B’ from (A) Mlalazi River, South Africa, and (B) Sarbaz River, Makran basin, southeastern Iran, and G. tenuiformis from (C) South Africa and (D) Wadi Hasik, Oman. Photos by J. Coetzee (A), A. Jouladeh Roudbar (B), O. Gon (C), and H.R. Esmaeili (D)." figureDoi="http://doi.org/10.5281/zenodo.14853710" httpUri="https://zenodo.org/record/14853710/files/figure.png" pageId="6" pageNumber="587">Fig. 6B</figureCitation>
|
||||
) (
|
||||
<bibRefCitation id="9A700DF9A7256679FE76F906F0DBF89F" author="Hammer, M. P. & Taillebois, L. & King, A. J. & Crook, D. A. & Wedd, D. & Adams, M. & Unmack, P. J. & Hoese, D. F. & Bertozzi, T." box="[501,739,1776,1802]" pageId="6" pageNumber="587" pagination="87 - 100" refId="ref6541" refString="Hammer, M. P., Taillebois, L., King, A. J., Crook, D. A., Wedd, D., Adams, M., Unmack, P. J., Hoese, D. F. & Bertozzi, T. (2021) Unravelling the taxonomy and identification of a problematic group of benthic fishes from tropical rivers (Gobiidae: Glossogobius). Journal of Fish Biology, 99 (1), 87-100. https://doi.org/10.1111/jfb.14701" type="journal article" year="2021">
|
||||
Hammer
|
||||
<emphasis id="CC95AC1AA7256679FDE2F907F0AEF89F" box="[609,662,1776,1802]" italics="true" pageId="6" pageNumber="587">et al</emphasis>
|
||||
. 2021
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="9A700DF9A7256679FD71F906F633F89F" author="Hoese, D. F. & Hammer, M. P." box="[754,1035,1776,1802]" pageId="6" pageNumber="587" pagination="79 - 115" refId="ref6759" refString="Hoese, D. F. & Hammer, M. P. (2021) A review of the Glossogobius giuris complex in Australia, with wider discussion on nomenclature and possible synonymies. Zootaxa, 4974 (1), 79-115. https://doi.org/10.11646/zootaxa.4974.1.3" type="journal article" year="2021">Hoese & Hammer 2021</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="9A700DF9A7256679FB99F906F6C9F89F" author="Zarei, F. & Kovacic, M. & Esmaeili, H. R. & Ghanbarifardi, M." box="[1050,1265,1776,1802]" pageId="6" pageNumber="587" pagination="28 - 50" refId="ref7758" refString="Zarei, F., Kovacic, M., Esmaeili, H. R. & Ghanbarifardi, M. (2023 b) Morphological and molecular characterization of the Makran Glossogobius cf. giuris (Teleostei: Gobiidae) and the Glossogobius giuris species complex taxonomy. Zootaxa, 5346 (1), 28-50. https://doi.org/10.11646/zootaxa.5346.1.2" type="journal article" year="2023">
|
||||
Zarei
|
||||
<emphasis id="CC95AC1AA7256679FBE3F907F6ADF89F" box="[1120,1173,1776,1802]" italics="true" pageId="6" pageNumber="587">et al</emphasis>
|
||||
. 2023b
|
||||
</bibRefCitation>
|
||||
); (iii) clade C, recorded in
|
||||
<collectingCountry id="86F63098A7256679FEA3F8E2F364F8BB" box="[288,348,1812,1838]" name="India" pageId="6" pageNumber="587">India</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FEEAF8E2F3E0F8BB" box="[361,472,1812,1838]" name="Myanmar" pageId="6" pageNumber="587">Myanmar</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FE67F8E2F012F8BB" box="[484,554,1812,1838]" name="Nepal" pageId="6" pageNumber="587">Nepal</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="86F63098A7256679FDB5F8E2F084F8BB" box="[566,700,1812,1838]" name="Bangladesh" pageId="6" pageNumber="587">Bangladesh</collectingCountry>
|
||||
, and
|
||||
<collectingCountry id="86F63098A7256679FD7AF8E2F160F8BB" box="[761,856,1812,1838]" name="Vietnam" pageId="6" pageNumber="587">Vietnam</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="9A700DF9A7256679FCEAF8E2F669F8BB" author="Hammer, M. P. & Taillebois, L. & King, A. J. & Crook, D. A. & Wedd, D. & Adams, M. & Unmack, P. J. & Hoese, D. F. & Bertozzi, T." box="[873,1105,1812,1838]" pageId="6" pageNumber="587" pagination="87 - 100" refId="ref6541" refString="Hammer, M. P., Taillebois, L., King, A. J., Crook, D. A., Wedd, D., Adams, M., Unmack, P. J., Hoese, D. F. & Bertozzi, T. (2021) Unravelling the taxonomy and identification of a problematic group of benthic fishes from tropical rivers (Gobiidae: Glossogobius). Journal of Fish Biology, 99 (1), 87-100. https://doi.org/10.1111/jfb.14701" type="journal article" year="2021">
|
||||
Hammer
|
||||
<emphasis id="CC95AC1AA7256679FC57F8E3F63EF8BB" box="[980,1030,1812,1838]" italics="true" pageId="6" pageNumber="587">et al</emphasis>
|
||||
. 2021
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="9A700DF9A7256679FBDDF8E2F716F8BB" author="Zarei, F. & Kovacic, M. & Esmaeili, H. R. & Ghanbarifardi, M." box="[1118,1326,1812,1838]" pageId="6" pageNumber="587" pagination="28 - 50" refId="ref7758" refString="Zarei, F., Kovacic, M., Esmaeili, H. R. & Ghanbarifardi, M. (2023 b) Morphological and molecular characterization of the Makran Glossogobius cf. giuris (Teleostei: Gobiidae) and the Glossogobius giuris species complex taxonomy. Zootaxa, 5346 (1), 28-50. https://doi.org/10.11646/zootaxa.5346.1.2" type="journal article" year="2023">
|
||||
Zarei
|
||||
<emphasis id="CC95AC1AA7256679FB22F8E3F6ECF8BB" box="[1185,1236,1812,1838]" italics="true" pageId="6" pageNumber="587">et al</emphasis>
|
||||
. 2023b
|
||||
</bibRefCitation>
|
||||
); and (iv) clade D, found in southern Africa (
|
||||
<figureCitation id="66DA6C8DA7256679FDA5F8CEF046F8C7" box="[550,638,1848,1874]" captionStart="FIGURE 6" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[151,1436,378,1908]" captionTargetId="figure-122@7.[151,1436,378,1908]" captionTargetPageId="7" captionText="FIGURE 6. Uncatalogued specimens of Glossogobius giuris ‘clade B’ from (A) Mlalazi River, South Africa, and (B) Sarbaz River, Makran basin, southeastern Iran, and G. tenuiformis from (C) South Africa and (D) Wadi Hasik, Oman. Photos by J. Coetzee (A), A. Jouladeh Roudbar (B), O. Gon (C), and H.R. Esmaeili (D)." figureDoi="http://doi.org/10.5281/zenodo.14853710" httpUri="https://zenodo.org/record/14853710/files/figure.png" pageId="6" pageNumber="587">Fig. 6C</figureCitation>
|
||||
) and
|
||||
<collectingCountry id="86F63098A7256679FD3CF8CEF13BF8C7" box="[703,771,1848,1874]" name="Oman" pageId="6" pageNumber="587">Oman</collectingCountry>
|
||||
(
|
||||
<figureCitation id="66DA6C8DA7256679FC97F8CEF157F8C7" box="[788,879,1848,1874]" captionStart="FIGURE 6" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[151,1436,378,1908]" captionTargetId="figure-122@7.[151,1436,378,1908]" captionTargetPageId="7" captionText="FIGURE 6. Uncatalogued specimens of Glossogobius giuris ‘clade B’ from (A) Mlalazi River, South Africa, and (B) Sarbaz River, Makran basin, southeastern Iran, and G. tenuiformis from (C) South Africa and (D) Wadi Hasik, Oman. Photos by J. Coetzee (A), A. Jouladeh Roudbar (B), O. Gon (C), and H.R. Esmaeili (D)." figureDoi="http://doi.org/10.5281/zenodo.14853710" httpUri="https://zenodo.org/record/14853710/files/figure.png" pageId="6" pageNumber="587">Fig. 6D</figureCitation>
|
||||
), reassigned to
|
||||
<taxonomicName id="39E10B8BA7256679FBA1F8CEF6FDF8C7" authorityName="Fowler" authorityYear="1934" box="[1058,1221,1848,1874]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="tenuiformis">
|
||||
<emphasis id="CC95AC1AA7256679FBA1F8CEF6FDF8C7" box="[1058,1221,1848,1874]" italics="true" pageId="6" pageNumber="587">G. tenuiformis</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="9A700DF9A7256679FB73F8CEF2DDF8E3" author="Al Jufaili, S. M. & Esmaeili, H. R. & Sayyadzadeh, G. & Masoumi, A. H. & Larson, H. K." pageId="6" pageNumber="587" pagination="543 - 554" refId="ref5837" refString="Al Jufaili, S. M., Esmaeili, H. R., Sayyadzadeh, G., Masoumi, A. H. & Larson, H. K. (2022) Redescription of the goby Glossogobius tenuiformis Fowler, 1934 (Teleostei: Gobiidae) and assignment of Oman Glossogobius populations: a morpho-molecular approach. Zootaxa, 5133 (4), 543-554. https://doi.org/10.11646/zootaxa.5133.4.5" type="journal article" year="2022">
|
||||
Al Jufaili
|
||||
<emphasis id="CC95AC1AA7256679FAE0F8CFF7ADF8C7" box="[1379,1429,1848,1874]" italics="true" pageId="6" pageNumber="587">et al</emphasis>
|
||||
. (2022)
|
||||
</bibRefCitation>
|
||||
. According to these studies, clades B and C remain taxonomically unresolved, despite the redescription of
|
||||
<taxonomicName id="39E10B8BA7256679FF3BF876F325F80F" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[184,285,1920,1946]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA7256679FF3BF876F325F80F" box="[184,285,1920,1946]" italics="true" pageId="6" pageNumber="587">G. giuris</emphasis>
|
||||
</taxonomicName>
|
||||
based on clade B (
|
||||
<bibRefCitation id="9A700DF9A7256679FE7CF876F12DF80F" author="Hoese, D. F. & Hammer, M. P." box="[511,789,1920,1946]" pageId="6" pageNumber="587" pagination="79 - 115" refId="ref6759" refString="Hoese, D. F. & Hammer, M. P. (2021) A review of the Glossogobius giuris complex in Australia, with wider discussion on nomenclature and possible synonymies. Zootaxa, 4974 (1), 79-115. https://doi.org/10.11646/zootaxa.4974.1.3" type="journal article" year="2021">Hoese & Hammer 2021</bibRefCitation>
|
||||
). However,
|
||||
<bibRefCitation id="9A700DF9A7256679FC23F876F649F80F" author="Hamilton, F." box="[928,1137,1920,1946]" pageId="6" pageNumber="587" refId="ref6499" refString="Hamilton, F. (1822) An account of the fishes found in the river Ganges and its branches. Printed for A. Constable and company, Edinburgh and London, vii + 405 pp., 39 pls." type="book" year="1822">Hamilton’s (1822)</bibRefCitation>
|
||||
original description of
|
||||
<taxonomicName id="39E10B8BA7256679FA02F876F2E0F82B" baseAuthorityName="Hamilton" baseAuthorityYear="1822" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA7256679FA02F876F2E0F82B" italics="true" pageId="6" pageNumber="587">G. giuris</emphasis>
|
||||
</taxonomicName>
|
||||
from the Ganges River lacks sufficient detail, allowing alignment with all recent redescriptions, including
|
||||
<taxonomicName id="39E10B8BA7256679FF14F83EF303F877" authorityName="Fowler" authorityYear="1934" box="[151,315,1992,2018]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="tenuiformis">
|
||||
<emphasis id="CC95AC1AA7256679FF14F83EF303F877" box="[151,315,1992,2018]" italics="true" pageId="6" pageNumber="587">G. tenuiformis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="39E10B8BA7256679FEC8F83EF3FEF877" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[331,454,1992,2018]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA7256679FEC8F83EF3FEF877" box="[331,454,1992,2018]" italics="true" pageId="6" pageNumber="587">G. laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="39E10B8BA7256679FD84F83EF054F877" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[519,620,1992,2018]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA7256679FD84F83EF054F877" box="[519,620,1992,2018]" italics="true" pageId="6" pageNumber="587">G. giuris</emphasis>
|
||||
</taxonomicName>
|
||||
based on clade B material. There is no type material for
|
||||
<taxonomicName id="39E10B8BA7256679FB7CF83EF75CF877" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[1279,1380,1992,2018]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA7256679FB7CF83EF75CF877" box="[1279,1380,1992,2018]" italics="true" pageId="6" pageNumber="587">G. giuris</emphasis>
|
||||
</taxonomicName>
|
||||
, and the
|
||||
<typeStatus id="215ACEAAA7246678FF40FF61F312FF24" box="[195,298,151,177]" pageId="7" pageNumber="588" type="lectotype">lectotype</typeStatus>
|
||||
designated by
|
||||
<bibRefCitation id="9A700DF9A7246678FE5FFF61F040FF24" author="Fricke, R." box="[476,632,151,177]" pageId="7" pageNumber="588" pagination="1 - 759" refId="ref6228" refString="Fricke, R. (1999) Fishes of the Mascarene Islands (Reunion, Mauritius, Rodriguez). An annotated checklist with descriptions of new species. Theses Zoologicae, 31, 1-759" type="journal article" year="1999">Fricke (1999)</bibRefCitation>
|
||||
relies on Hamilton’s illustration. Clade C also lacks a morphological description (
|
||||
<bibRefCitation id="9A700DF9A7246678FEA6FF4DF035FF40" author="Hammer, M. P. & Taillebois, L. & King, A. J. & Crook, D. A. & Wedd, D. & Adams, M. & Unmack, P. J. & Hoese, D. F. & Bertozzi, T." box="[293,525,187,213]" pageId="7" pageNumber="588" pagination="87 - 100" refId="ref6541" refString="Hammer, M. P., Taillebois, L., King, A. J., Crook, D. A., Wedd, D., Adams, M., Unmack, P. J., Hoese, D. F. & Bertozzi, T. (2021) Unravelling the taxonomy and identification of a problematic group of benthic fishes from tropical rivers (Gobiidae: Glossogobius). Journal of Fish Biology, 99 (1), 87-100. https://doi.org/10.1111/jfb.14701" type="journal article" year="2021">
|
||||
Hammer
|
||||
<emphasis id="CC95AC1AA7246678FE0CFF4DF3FAFF40" box="[399,450,187,213]" italics="true" pageId="7" pageNumber="588">et al</emphasis>
|
||||
. 2021
|
||||
</bibRefCitation>
|
||||
). While
|
||||
<taxonomicName id="39E10B8BA7246678FDEDFF4DF129FF40" authorityName="Fowler" authorityYear="1934" box="[622,785,187,213]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="588" phylum="Chordata" rank="species" species="tenuiformis">
|
||||
<emphasis id="CC95AC1AA7246678FDEDFF4DF129FF40" box="[622,785,187,213]" italics="true" pageId="7" pageNumber="588">G. tenuiformis</emphasis>
|
||||
</taxonomicName>
|
||||
has not been recorded in the Ganges, clades B and C are well documented. Additionally, the recent discovery of
|
||||
<taxonomicName id="39E10B8BA7246678FC8DFF29F1B1FF6C" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[782,905,223,249]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="588" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA7246678FC8DFF29F1B1FF6C" box="[782,905,223,249]" italics="true" pageId="7" pageNumber="588">G. laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
in southern
|
||||
<collectingCountry id="86F63098A7246678FB9AFF29F671FF6C" box="[1049,1097,223,249]" name="Iran" pageId="7" pageNumber="588">Iran</collectingCountry>
|
||||
, i.e., approximately
|
||||
<quantity id="3919DDEDA7246678FAB2FF28F7A4FF6C" box="[1329,1436,222,249]" metricMagnitude="6" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="588" unit="km" value="5500.0">5,500 km</quantity>
|
||||
west of the nearest locality record in
|
||||
<collectingCountry id="86F63098A7246678FDB6FEF5F083FE88" box="[565,699,259,285]" name="Bangladesh" pageId="7" pageNumber="588">Bangladesh</collectingCountry>
|
||||
, suggests a broader distribution across the northern Indian Ocean, possibly including the Ganges. Therefore, a thorough phylogenetic and morphological analysis of fresh specimens from the Ganges is needed to clarify the taxonomy of the
|
||||
<taxonomicName id="39E10B8BA7246678FC9EFEBDF1B9FEF1" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[797,897,331,357]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="588" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA7246678FC9EFEBDF1B9FEF1" box="[797,897,331,357]" italics="true" pageId="7" pageNumber="588">G. giuris</emphasis>
|
||||
</taxonomicName>
|
||||
complex.
|
||||
</paragraph>
|
||||
<caption id="AA9E2080A7246678FF14F878F1AAF87B" ID-DOI="http://doi.org/10.5281/zenodo.14853710" ID-Zenodo-Dep="14853710" httpUri="https://zenodo.org/record/14853710/files/figure.png" pageId="7" pageNumber="588" startId="7.[151,250,1934,1958]" targetBox="[151,1436,378,1908]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="FE5E7008A7246678FF14F878F1AAF87B" blockId="7.[151,1436,1934,2030]" pageId="7" pageNumber="588">
|
||||
<emphasis id="CC95AC1AA7246678FF14F878F32CF833" bold="true" box="[151,276,1934,1958]" pageId="7" pageNumber="588">FIGURE 6.</emphasis>
|
||||
Uncatalogued specimens of
|
||||
<taxonomicName id="39E10B8BA7246678FDBEF878F132F833" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[573,778,1934,1958]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="588" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA7246678FDBEF878F132F833" box="[573,778,1934,1958]" italics="true" pageId="7" pageNumber="588">Glossogobius giuris</emphasis>
|
||||
</taxonomicName>
|
||||
‘clade B’ from (A) Mlalazi River, South Africa, and (B) Sarbaz River, Makran basin, southeastern Iran, and
|
||||
<taxonomicName id="39E10B8BA7246678FDE7F845F0C1F85F" authorityName="Fowler" authorityYear="1934" box="[612,761,1970,1994]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="588" phylum="Chordata" rank="species" species="tenuiformis">
|
||||
<emphasis id="CC95AC1AA7246678FDE7F845F0C1F85F" box="[612,761,1970,1994]" italics="true" pageId="7" pageNumber="588">G. tenuiformis</emphasis>
|
||||
</taxonomicName>
|
||||
from (C) South Africa and (D) Wadi Hasik, Oman. Photos by J. Coetzee (A), A. Jouladeh Roudbar (B), O. Gon (C), and H.R. Esmaeili (D).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="FE5E7008A72B6677FF44FF61F3C4FD64" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="589">
|
||||
The gobies of the
|
||||
<collectingCountry id="86F63098A72B6677FE16FF61F3E1FF24" box="[405,473,151,177]" name="Oman" pageId="8" pageNumber="589">Oman</collectingCountry>
|
||||
Sea and Persian Gulf have not been extensively studied, with data available for only a limited number of species. This is primarily due to factors such as their low economic value, small size, cryptic appearance and behavior, incomplete geographic sampling, their presence in challenging habitats that are difficult to sample, and lack of systematic studies and less attention from ichthyologists. Recent taxonomic studies in these regions however, have led to the description of several new species, including
|
||||
<taxonomicName id="39E10B8BA72B6677FB85FED1F2E1FEF0" authority="(Kovacic et al. 2020)" baseAuthorityName="Kovacic" baseAuthorityYear="2020" family="Gobiidae" genus="Silhouettea" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="ghazalae">
|
||||
<emphasis id="CC95AC1AA72B6677FB85FED1F6C8FED4" box="[1030,1264,295,321]" italics="true" pageId="8" pageNumber="589">Silhouettea ghazalae</emphasis>
|
||||
(
|
||||
<bibRefCitation id="9A700DF9A72B6677FB7DFED1F2E9FEF0" author="Kovacic, M. & Sadeghi, R. & Esmaeili, H. R." pageId="8" pageNumber="589" pagination="49 - 66" refId="ref6879" refString="Kovacic, M., Sadeghi, R. & Esmaeili, H. R. (2020) New species of Silhouettea (Teleostei: Gobiidae) from Qeshm Island, Iran and the DNA barcoding of the Persian Gulf and Oman Sea gobies. Zootaxa, 4750 (1), 49-66. https://doi.org/10.11646/zootaxa.4750.1.3" type="journal article" year="2020">
|
||||
Kovačić
|
||||
<emphasis id="CC95AC1AA72B6677FAE7FED1F7ADFED4" box="[1380,1429,295,321]" italics="true" pageId="8" pageNumber="589">et al</emphasis>
|
||||
. 2020
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="39E10B8BA72B6677FE8FFEBDF0D3FEF0" authority="(Zarei et al. 2022)" baseAuthorityName="Zarei" baseAuthorityYear="2022" box="[268,747,331,357]" family="Gobiidae" genus="Oxyurichthys" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="omanensis">
|
||||
<emphasis id="CC95AC1AA72B6677FE8FFEBDF024FEF1" box="[268,540,331,357]" italics="true" pageId="8" pageNumber="589">Oxyurichthys omanensis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="9A700DF9A72B6677FDAAFEBDF0DBFEF0" author="Zarei, F. & Al Jufaili, S. M. & Esmaeili, H. R." box="[553,739,331,357]" pageId="8" pageNumber="589" pagination="361 - 376" refId="ref7633" refString="Zarei, F., Al Jufaili, S. M. & Esmaeili, H. R. (2022) Oxyurichthys omanensis sp. nov., a new Eyebrow Goby (Teleostei: Gobiidae) from Oman. Zootaxa, 5182 (4), 361-376. https://doi.org/10.11646/zootaxa.5182.4.3" type="journal article" year="2022">
|
||||
Zarei
|
||||
<emphasis id="CC95AC1AA72B6677FDE8FEBDF0A3FEF0" box="[619,667,331,357]" italics="true" pageId="8" pageNumber="589">et al</emphasis>
|
||||
. 2022
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, as well as new records such as
|
||||
<taxonomicName id="39E10B8BA72B6677FBC6FEBDF2E1FE1C" authority="(Bleeker, 1854)" baseAuthorityName="Bleeker" baseAuthorityYear="1854" family="Gobiidae" genus="Favonigobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="reichei">
|
||||
<emphasis id="CC95AC1AA72B6677FBC6FEBDF709FEF0" box="[1093,1329,331,357]" italics="true" pageId="8" pageNumber="589">Favonigobius reichei</emphasis>
|
||||
(Bleeker, 1854)
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="9A700DF9A72B6677FE87FE99F3CDFE1C" author="Sadeghi, R. & Esmaeili, H. R. & Fricke, R. & Larson, H." box="[260,501,367,393]" pageId="8" pageNumber="589" pagination="641 - 645" refId="ref7258" refString="Sadeghi, R., Esmaeili, H. R., Fricke, R. & Larson, H. (2017) New geographical record and morphological features of the Indo-Pacific tropical sand goby, Favonigobius reichei (Bleeker, 1854) from Iranian coast of the Makran Sea (Teleostei, Gobiidae). Check List, 13, 641-645. https://doi.org/10.15560/13.5.641" type="journal article" year="2017">
|
||||
Sadeghi
|
||||
<emphasis id="CC95AC1AA72B6677FEE4FE99F3A1FE1C" box="[359,409,367,393]" italics="true" pageId="8" pageNumber="589">et al</emphasis>
|
||||
. (2017)
|
||||
</bibRefCitation>
|
||||
,
|
||||
<taxonomicName id="39E10B8BA72B6677FD82FE99F1FFFE1C" authority="(Randall, 1994)" baseAuthorityName="Randall" baseAuthorityYear="1994" box="[513,967,367,393]" family="Gobiidae" genus="Coryogalops" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="tessellatus">
|
||||
<emphasis id="CC95AC1AA72B6677FD82FE99F137FE1C" box="[513,783,367,393]" italics="true" pageId="8" pageNumber="589">Coryogalops tessellatus</emphasis>
|
||||
(Randall, 1994)
|
||||
</taxonomicName>
|
||||
by Sadeghi
|
||||
<emphasis id="CC95AC1AA72B6677FBD6FE99F6BFFE1C" box="[1109,1159,367,393]" italics="true" pageId="8" pageNumber="589">et al</emphasis>
|
||||
. (2019a),
|
||||
<taxonomicName id="39E10B8BA72B6677FB7FFE99F3E6FE38" authority="(Bleeker, 1853)" baseAuthorityName="Bleeker" baseAuthorityYear="1853" family="Gobiidae" genus="Cryptocentrus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="cyanotaenia">
|
||||
<emphasis id="CC95AC1AA72B6677FB7FFE99F318FE39" italics="true" pageId="8" pageNumber="589">Cryptocentrus cyanotaenia</emphasis>
|
||||
(Bleeker, 1853)
|
||||
</taxonomicName>
|
||||
by Sadeghi
|
||||
<emphasis id="CC95AC1AA72B6677FDF5FE65F093FE38" box="[630,683,403,429]" italics="true" pageId="8" pageNumber="589">et al</emphasis>
|
||||
. (2019b),
|
||||
<taxonomicName id="39E10B8BA72B6677FCABFE65F74DFE38" authority="(Hoese & Randall, 1982)" baseAuthorityName="Hoese & Randall" baseAuthorityYear="1982" box="[808,1397,403,429]" family="Gobiidae" genus="Stonogobiops" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="nematodes">
|
||||
<emphasis id="CC95AC1AA72B6677FCABFE65F67CFE38" box="[808,1092,403,429]" italics="true" pageId="8" pageNumber="589">Stonogobiops nematodes</emphasis>
|
||||
(Hoese & Randall, 1982)
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="9A700DF9A72B6677FF14FE41F02EFE44" author="Ghanbarifardi, M. & Lagzian, M." box="[151,534,439,465]" pageId="8" pageNumber="589" pagination="385 - 392" refId="ref6334" refString="Ghanbarifardi, M. & Lagzian, M. (2019) Molecular phylogeny of gobies (Teleostei, Gobiidae) from Iranian water bodies with a new record. Russian Journal of Marine Biology, 45, 385-392. https://doi.org/10.1134/S1063074019050055" type="journal article" year="2019">Ghanbarifardi & Lagzian (2019)</bibRefCitation>
|
||||
,
|
||||
<taxonomicName id="39E10B8BA72B6677FDA4FE41F673FE44" authority="(de Beaufort, 1912)" baseAuthorityName="de Beaufort" baseAuthorityYear="1912" box="[551,1099,439,465]" family="Gobiidae" genus="Palutrus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="scapulopunctatus">
|
||||
<emphasis id="CC95AC1AA72B6677FDA4FE41F160FE44" box="[551,856,439,465]" italics="true" pageId="8" pageNumber="589">Palutrus scapulopunctatus</emphasis>
|
||||
(de Beaufort, 1912)
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="39E10B8BA72B6677FB0FFE41F370FE60" authority="(Bleeker, 1854)" baseAuthorityName="Bleeker" baseAuthorityYear="1854" family="Gobiidae" genus="Bathygobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="cocosensis">
|
||||
<emphasis id="CC95AC1AA72B6677FB0FFE41F7A4FE45" box="[1164,1436,439,465]" italics="true" pageId="8" pageNumber="589">Bathygobius cocosensis</emphasis>
|
||||
(Bleeker, 1854)
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="9A700DF9A72B6677FEF4FE2DF08EFE60" author="Sadeghi, R. & Esmaeili, H. R." box="[375,694,475,501]" pageId="8" pageNumber="589" pagination="143 - 149" refId="ref7151" refString="Sadeghi, R. & Esmaeili, H. R. (2019 a) First documentation of an uncommon goby genus and species, Palutrus scapulopunctatus (de Beaufort, 1912) from the Persian Gulf (Teleostei: Gobiidae). Iranian Journal of Ichthyology, 6, 143-149." type="journal article" year="2019">Sadeghi & Esmaeili (2019a</bibRefCitation>
|
||||
, b),
|
||||
<taxonomicName id="39E10B8BA72B6677FD68FE2DF6F8FE60" authority="(Fowler, 1934)" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[747,1216,475,501]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="tenuiformis">
|
||||
<emphasis id="CC95AC1AA72B6677FD68FE2DF635FE60" box="[747,1037,475,501]" italics="true" pageId="8" pageNumber="589">Glossogobius tenuiformis</emphasis>
|
||||
(Fowler, 1934)
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="9A700DF9A72B6677FB6EFE2DF2DCFD8C" author="Al Jufaili, S. M. & Esmaeili, H. R. & Sayyadzadeh, G. & Masoumi, A. H. & Larson, H. K." pageId="8" pageNumber="589" pagination="543 - 554" refId="ref5837" refString="Al Jufaili, S. M., Esmaeili, H. R., Sayyadzadeh, G., Masoumi, A. H. & Larson, H. K. (2022) Redescription of the goby Glossogobius tenuiformis Fowler, 1934 (Teleostei: Gobiidae) and assignment of Oman Glossogobius populations: a morpho-molecular approach. Zootaxa, 5133 (4), 543-554. https://doi.org/10.11646/zootaxa.5133.4.5" type="journal article" year="2022">
|
||||
Al Jufaili
|
||||
<emphasis id="CC95AC1AA72B6677FAE1FE2DF7ADFE60" box="[1378,1429,475,501]" italics="true" pageId="8" pageNumber="589">et al</emphasis>
|
||||
. (2022)
|
||||
</bibRefCitation>
|
||||
,
|
||||
<taxonomicName id="39E10B8BA72B6677FF73FE09F090FD8C" authority="(Bleeker, 1853)" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[240,680,510,537]" class="Actinopterygii" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="acanthopomus">
|
||||
<emphasis id="CC95AC1AA72B6677FF73FE09F3CAFD8C" box="[240,498,511,537]" italics="true" pageId="8" pageNumber="589">Eleotris acanthopomus</emphasis>
|
||||
(Bleeker, 1853)
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="9A700DF9A72B6677FD51FE09F1FEFD8C" author="Esmaeili, H. R. & Jufaili, S. A. & Masoumi, A. H. & Zarei, F." box="[722,966,511,537]" pageId="8" pageNumber="589" pagination="451 - 503" refId="ref6156" refString="Esmaeili, H. R., Jufaili, S. A., Masoumi, A. H. & Zarei, F. (2022) Ichthyodiversity in southeastern Arabian Peninsula: annotated checklist, taxonomy, short description and distribution of Inland fishes of Oman. Zootaxa, 5134 (4), 451-503. https://doi.org/10.11646/zootaxa.5134.4.1" type="journal article" year="2022">
|
||||
Esmaeili
|
||||
<emphasis id="CC95AC1AA72B6677FCB9FE09F153FD8C" box="[826,875,511,537]" italics="true" pageId="8" pageNumber="589">et al</emphasis>
|
||||
. (2022)
|
||||
</bibRefCitation>
|
||||
, and
|
||||
<taxonomicName id="39E10B8BA72B6677FB82FE09F2E1FDA8" authority="(Bleeker, 1860)" baseAuthorityName="Bleeker" baseAuthorityYear="1860" family="Gobiidae" genus="Favonigobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="gymnauchen">
|
||||
<emphasis id="CC95AC1AA72B6677FB82FE09F708FD8C" box="[1025,1328,511,537]" italics="true" pageId="8" pageNumber="589">Favonigobius gymnauchen</emphasis>
|
||||
(Bleeker, 1860)
|
||||
</taxonomicName>
|
||||
and clade B of the
|
||||
<taxonomicName id="39E10B8BA72B6677FE43FDD5F09BFDA9" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[448,675,547,573]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA72B6677FE43FDD5F09BFDA9" box="[448,675,547,573]" italics="true" pageId="8" pageNumber="589">Glossogobius giuris</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="9A700DF9A72B6677FD51FDD5F193FDA8" author="Zarei, F. & Esmaeili, H. R. & Hashemi, S. H." box="[722,939,547,573]" pageId="8" pageNumber="589" pagination="119 - 123" refId="ref7697" refString="Zarei, F., Esmaeili, H. R. & Hashemi, S. H. (2023 a) First record of Favonigobius gymnauchen (Bleeker, 1860) (Teleostei: Gobiidae) in the north-western Indian Ocean: evidence for potential taxonomic diversity. International Journal of Aquatic Biology, 11, 119-123." type="journal article" year="2023">
|
||||
Zarei
|
||||
<emphasis id="CC95AC1AA72B6677FC94FDD5F172FDA8" box="[791,842,547,573]" italics="true" pageId="8" pageNumber="589">et al</emphasis>
|
||||
. (2023a
|
||||
</bibRefCitation>
|
||||
, b). In the Red Sea basin,
|
||||
<bibRefCitation id="9A700DF9A72B6677FB63FDD5F2DAFDF4" author="Golani, D. & Fricke, R." pageId="8" pageNumber="589" pagination="1 - 215" refId="ref6387" refString="Golani, D. & Fricke, R. (2018) Checklist of the Red Sea fishes with delineation of the Gulf of Suez, Gulf of Aqaba, endemism and Lessepsian migrants. Zootaxa, 4509 (1), 1-215. https://doi.org/10.11646/zootaxa.4509.1.1" type="journal article" year="2018">Golani & Fricke (2018)</bibRefCitation>
|
||||
documented 153 goby species. However, a recent update by
|
||||
<bibRefCitation id="9A700DF9A72B6677FC12FDB1F6EFFDF4" author="Bogorodsky, S. V. & Goren, M." box="[913,1239,583,609]" pageId="8" pageNumber="589" pagination="123 - 163" refId="ref5988" refString="Bogorodsky, S. V. & Goren, M. (2023) An updated checklist of the Red Sea gobioid species (Teleostei: Gobiiformes), with four new records. Zoology in the Middle East, 69, 123-163. https://doi.org/10.1080/09397140.2023.2203978" type="journal article" year="2023">Bogorodsky & Goren (2023)</bibRefCitation>
|
||||
added 20 species to the list and corrected the identification of seven species. These findings suggest a significant increase in goby species richness in the northwestern Indian Ocean over the past decade, largely driven by increased sampling, the integration of molecular analyses into taxonomic research, and the re-examination of previously collected specimens stored in zoological collections.
|
||||
</paragraph>
|
||||
<paragraph id="FE5E7008A72B6677FF44FD0DF606FC78" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="589">
|
||||
Although the biology of species within
|
||||
<taxonomicName id="39E10B8BA72B6677FD33FD0DF170FC80" authorityName="Gill" authorityYear="1859" box="[688,840,763,789]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="genus">
|
||||
<emphasis id="CC95AC1AA72B6677FD33FD0DF170FC80" box="[688,840,763,789]" italics="true" pageId="8" pageNumber="589">Glossogobius</emphasis>
|
||||
</taxonomicName>
|
||||
remains insufficiently understood, it is generally assumed that widely distributed species such as
|
||||
<taxonomicName id="39E10B8BA72B6677FD3BFCE9F10AFCAC" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[696,818,799,825]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA72B6677FD3BFCE9F10AFCAC" box="[696,818,799,825]" italics="true" pageId="8" pageNumber="589">G. laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="39E10B8BA72B6677FCC2FCE9F19DFCAD" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[833,933,799,825]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA72B6677FCC2FCE9F19DFCAD" box="[833,933,799,825]" italics="true" pageId="8" pageNumber="589">G. giuris</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="CC95AC1AA72B6677FC2EFCE8F18EFCAD" bold="true" box="[941,950,798,824]" pageId="8" pageNumber="589">‘</emphasis>
|
||||
clade B
|
||||
<emphasis id="CC95AC1AA72B6677FB8FFCE8F62DFCAD" bold="true" box="[1036,1045,798,824]" pageId="8" pageNumber="589">’</emphasis>
|
||||
and
|
||||
<taxonomicName id="39E10B8BA72B6677FBCEFCE9F2F7FCC8" authority="Akihito & Meguro, 1975" authorityName="Akihito & Meguro" authorityYear="1975" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="aureus">
|
||||
<emphasis id="CC95AC1AA72B6677FBCEFCE9F683FCAD" box="[1101,1211,799,825]" italics="true" pageId="8" pageNumber="589">G. aureus</emphasis>
|
||||
<bibRefCitation id="9A700DF9A72B6677FB41FCE9F2F7FCC8" author="Akihito, P. & Meguro, K." pageId="8" pageNumber="589" pagination="127 - 142" refId="ref5784" refString="Akihito, P. & Meguro, K. (1975) Description of a new gobiid fish, Glossogobius aureus, with notes on related species of the genus. Japanese Journal of Ichthyology, 22, 127-142. https://doi.org/10.11369/JJI1950.22.127" type="journal article" year="1975">Akihito & Meguro, 1975</bibRefCitation>
|
||||
</taxonomicName>
|
||||
exhibit amphidromous behavior. These species are typically found in freshwater as adults, with juveniles occasionally inhabiting estuaries. They breed in freshwater, have marine and estuarine larval and post-larval stages, and eventually return to rivers (e.g.,
|
||||
<bibRefCitation id="9A700DF9A72B6677FDB2FC7DF124FC30" author="Hoese, D. F. & Allen, G. R." box="[561,796,907,933]" pageId="8" pageNumber="589" pagination="269 - 285" refId="ref6645" refString="Hoese, D. F. & Allen, G. R. (2012) A review of the amphidromous species of the Glossogobius celebius complex, with description of three new species. Cybium, 35, 269-285. https://doi.org/10.26028/cybium/2011-354-002" type="journal article" year="2012">Hoese & Allen 2012</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="9A700DF9A72B6677FCA4FC7DF60EFC30" author="Hoese, D. F. & Hammer, M. P." box="[807,1078,907,933]" pageId="8" pageNumber="589" pagination="79 - 115" refId="ref6759" refString="Hoese, D. F. & Hammer, M. P. (2021) A review of the Glossogobius giuris complex in Australia, with wider discussion on nomenclature and possible synonymies. Zootaxa, 4974 (1), 79-115. https://doi.org/10.11646/zootaxa.4974.1.3" type="journal article" year="2021">Hoese & Hammer 2021</bibRefCitation>
|
||||
). Consequently, the dispersal of
|
||||
<taxonomicName id="39E10B8BA72B6677FF14FC59F328FC5C" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[151,272,943,969]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA72B6677FF14FC59F328FC5C" box="[151,272,943,969]" italics="true" pageId="8" pageNumber="589">G. laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
to the northwestern Indian Ocean likely occurred via larval transport facilitated by the North Equatorial Current, which flows from east to west, originating south of the Indonesian islands.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
144
data/76/48/C1/7648C11EA72B6677FF14FBC8F7A4FADC.xml
Normal file
144
data/76/48/C1/7648C11EA72B6677FF14FBC8F7A4FADC.xml
Normal file
|
|
@ -0,0 +1,144 @@
|
|||
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<mods:namePart id="600E8251E0A4B8E1562A7429FC78474A">Zarei, Fatah</mods:namePart>
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<mods:nameIdentifier id="9464CA4CB727EE1B22C3F5508A19B0C7" type="email">fataahzarei@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0EABC2D3A78524FB39909A34EC06EBBD">Masoumi, Amir Hassan</mods:namePart>
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<mods:number id="1F0A4545A18144889B443AAE40789A81">5584</mods:number>
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<mods:detail id="30D9DD42A8D33DEE7C66E252D597A943" type="issue">
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<mods:number id="0B81B5E1EC9C6824974B1FDB901CBD18">4</mods:number>
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<mods:start id="068CC12002880F88ABBF63FEF243761D">581</mods:start>
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<mods:location id="029ABF90BBA2681F951296490299A4AA">
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<mods:url id="6EEF915F15E1009FA35F15267F8D26D3">https://doi.org/10.11646/zootaxa.5584.4.9</mods:url>
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||||
</mods:location>
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<mods:classification id="DA17FA0940CCE34239531944383C8296">journal article</mods:classification>
|
||||
<mods:identifier id="1D0DD1322DE2B92C6793100FFD736312" type="DOI">10.11646/zootaxa.5584.4.9</mods:identifier>
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||||
<mods:identifier id="C515D3CBD62A9EC54EF677AA4A9F2021" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F7165F6628174B6D5F3B39120CD662FA" type="Zenodo-Dep">14853698</mods:identifier>
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<mods:identifier id="C76AF5FC1579478A8F5BECECEB5175F2" type="ZooBank">8F8E403B-9D79-4E17-A93E-C76030BB5EAC</mods:identifier>
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</mods:mods>
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<treatment id="7648C11EA72B6677FF14FBC8F7A4FADC" LSID="urn:lsid:plazi:treatment:7648C11EA72B6677FF14FBC8F7A4FADC" httpUri="http://treatment.plazi.org/id/7648C11EA72B6677FF14FBC8F7A4FADC" lastPageNumber="589" pageId="8" pageNumber="589">
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<subSubSection id="B6FB2383A72B6677FF14FBC8F615FBCD" box="[151,1069,1086,1113]" pageId="8" pageNumber="589" type="nomenclature">
|
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<paragraph id="FE5E7008A72B6677FF14FBC8F615FBCD" blockId="8.[151,1069,1086,1113]" box="[151,1069,1086,1113]" pageId="8" pageNumber="589">
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||||
<heading id="A516C764A72B6677FF14FBC8F615FBCD" bold="true" box="[151,1069,1086,1113]" fontSize="11" level="1" pageId="8" pageNumber="589" reason="1">
|
||||
<emphasis id="CC95AC1AA72B6677FF14FBC8F615FBCD" bold="true" box="[151,1069,1086,1113]" pageId="8" pageNumber="589">
|
||||
A key to the species and clades of
|
||||
<taxonomicName id="39E10B8BA72B6677FDBFFBC8F0E5FBCC" ID-CoL="4P99" authority="Gill, 1859" authorityName="Gill" authorityYear="1859" box="[572,733,1086,1113]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="genus">
|
||||
<emphasis id="CC95AC1AA72B6677FDBFFBC8F0E5FBCC" bold="true" box="[572,733,1086,1113]" italics="true" pageId="8" pageNumber="589">Glossogobius</emphasis>
|
||||
</taxonomicName>
|
||||
from the
|
||||
<collectingCountry id="86F63098A72B6677FCD4FBC8F19DFBCD" box="[855,933,1086,1112]" name="Oman" pageId="8" pageNumber="589">Oman</collectingCountry>
|
||||
Sea basin:
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="B6FB2383A72B6677FF14FB75F7A4FADC" pageId="8" pageNumber="589" type="key">
|
||||
<key id="8A702739A72B6677FF14FB75F7A4FADC" pageId="8" pageNumber="589">
|
||||
<keyStep id="451568ADA72B6677FF14FB75F7A4FA9B" pageId="8" pageNumber="589">
|
||||
<paragraph id="FE5E7008A72B6677FF14FB75F7A5FB22" blockId="8.[151,1437,1155,1354]" pageId="8" pageNumber="589">
|
||||
<keyLead id="4510D33DA72B6677FF14FB75F7A5FB22" pageId="8" pageNumber="589">
|
||||
<emphasis id="CC95AC1AA72B6677FF14FB75F28DFB0D" bold="true" box="[151,181,1155,1176]" pageId="8" pageNumber="589">1a.</emphasis>
|
||||
Predorsal area fully scaled to above preopercle margin; all papillae lines on snout, suborbital, and underside of the head single; blotch on caudal fin base triangular, with a distinct unpigmented gap posteriorly........................
|
||||
<taxonomicName id="39E10B8BA72B6677FA8FFB57F7A5FB22" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[1292,1437,1185,1207]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="tenuiformis">
|
||||
<emphasis id="CC95AC1AA72B6677FA8FFB57F7A5FB22" bold="true" box="[1292,1437,1185,1207]" italics="true" pageId="8" pageNumber="589">G. tenuiformis</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="FE5E7008A72B6677FF14FB48F7A4FA9B" blockId="8.[151,1437,1155,1354]" pageId="8" pageNumber="589">
|
||||
<keyLead id="4510D33DA72B6677FF14FB48F7A4FA9B" pageId="8" pageNumber="589">
|
||||
<emphasis id="CC95AC1AA72B6677FF14FB48F28EFB46" bold="true" box="[151,182,1214,1235]" pageId="8" pageNumber="589">1b.</emphasis>
|
||||
Predorsal area fully scaled to behind eyes, with a distinct naked patch usually present; suborbital papillae lines 9–10, papillae Line 13 on underside of head, and papillae Line 1a on snout all doubled or tripled; blotch on caudal fin base triangular, without a distinct unpigmented gap posteriorly.....................................................................
|
||||
<emphasis id="CC95AC1AA72B6677FA13FB0FF7A4FA9B" bold="true" box="[1424,1436,1273,1294]" pageId="8" pageNumber="589">2</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="451568ADA72B6677FF14FAE0F7A4FADC" pageId="8" pageNumber="589">
|
||||
<paragraph id="FE5E7008A72B6677FF14FAE0F7A5FAB9" blockId="8.[151,1437,1155,1354]" box="[151,1437,1302,1324]" pageId="8" pageNumber="589">
|
||||
<keyLead id="4510D33DA72B6677FF14FAE0F7A5FAB9" box="[151,1437,1302,1324]" pageId="8" pageNumber="589">
|
||||
<emphasis id="CC95AC1AA72B6677FF14FAE0F28EFABE" bold="true" box="[151,182,1302,1323]" pageId="8" pageNumber="589">2a.</emphasis>
|
||||
Papillae Line 5 below eye and suborbital papillae Line 7 doubled.......................................
|
||||
<taxonomicName id="39E10B8BA72B6677FAB7FAE0F7A5FAB9" baseAuthorityName="De Vis" baseAuthorityYear="1884" box="[1332,1437,1302,1324]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="laticeps">
|
||||
<emphasis id="CC95AC1AA72B6677FAB7FAE0F7A5FAB9" bold="true" box="[1332,1437,1302,1324]" italics="true" pageId="8" pageNumber="589">G. laticeps</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="FE5E7008A72B6677FF14FAC5F7A4FADC" blockId="8.[151,1437,1155,1354]" box="[151,1436,1331,1354]" pageId="8" pageNumber="589">
|
||||
<keyLead id="4510D33DA72B6677FF14FAC5F7A4FADC" box="[151,1436,1331,1354]" pageId="8" pageNumber="589">
|
||||
<emphasis id="CC95AC1AA72B6677FF14FAC5F28FFADD" bold="true" box="[151,183,1331,1352]" pageId="8" pageNumber="589">2b.</emphasis>
|
||||
Papillae Line 5 below eye and suborbital papillae Line 7 single...................................
|
||||
<emphasis id="CC95AC1AA72B6677FB67FAC5F7A4FADC" bold="true" box="[1252,1436,1331,1353]" pageId="8" pageNumber="589">
|
||||
<taxonomicName id="39E10B8BA72B6677FB67FAC5F703FADD" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[1252,1339,1331,1352]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="giuris">
|
||||
<emphasis id="CC95AC1AA72B6677FB67FAC5F703FADD" bold="true" box="[1252,1339,1331,1352]" italics="true" pageId="8" pageNumber="589">G. giuris</emphasis>
|
||||
</taxonomicName>
|
||||
‘clade B’
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</key>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue