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<document id="F6EC8C8A1BAB5227F864BEC006D1A052" ID-DOI="10.11646/zootaxa.5569.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14703918" ID-ZooBank="332E63B4-97CA-4892-B8C4-DCA205BF91F8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737371648729" checkinUser="plazi" docAuthor="Oh, Jaeseok, Joharchi, Omid, Lee, Seunghwan &amp; Jung, Chuleui" docDate="2025" docId="F66F8788FFCDFFDA9DB1F9D0FA0E40D1" docLanguage="en" docName="zootaxa.5569.1.2.pdf" docOrigin="Zootaxa 5569 (1)" docSource="https://doi.org/10.11646/zootaxa.5569.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmolaelaps Berlese 1903" docType="key" docVersion="3" lastPageNumber="89" masterDocId="0A56FFF0FFECFFF89D26FF8BFF934121" masterDocTitle="New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from the Republic of Korea, with a review of the Korean species of the genus" masterLastPageNumber="92" masterPageNumber="55" pageNumber="88" updateTime="1737478317859" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="83F699C80451AC6DB6DA2100943832E0">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from the Republic of Korea, with a review of the Korean species of the genus</mods:title>
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<mods:namePart id="3480F4283E0B1E4A79D97B42CCC7CBD9">Oh, Jaeseok</mods:namePart>
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<mods:namePart id="39D6E03E0AED655A4412DA73BEEB02D9">Joharchi, Omid</mods:namePart>
<mods:affiliation id="92769870CB7B605DDC4C87140785C92E">Anatis Bioprotection Inc., Saint-Jacques-de-Mineur, Québec, J 0 J 1 Z 0, Canada &amp; Agriculture Science and Technology Institute, Andong National University, Andong, Republic of Korea</mods:affiliation>
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<mods:namePart id="C53404D11C4AD609B6A7E81300662254">Lee, Seunghwan</mods:namePart>
<mods:affiliation id="1AF454874FB891B728508C6B828A18CA">Research Institute of Agriculture and Life Sciences, Seoul National University, Seoul, Republic of Korea</mods:affiliation>
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<mods:namePart id="E49DC24716B4E2529F92561176C88E53">Jung, Chuleui</mods:namePart>
<mods:affiliation id="A827167139CAAF185CFB762CB225191F">Department of Plant medicals, Andong National University, Andong, Republic of Korea &amp; Agriculture Science and Technology Institute, Andong National University, Andong, Republic of Korea</mods:affiliation>
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<heading id="253181F2FFCDFFD99DB1F9D0FC374754" bold="true" box="[151,932,1626,1653]" fontSize="11" level="1" pageId="33" pageNumber="88" reason="1">
<emphasis id="4CB2EA8CFFCDFFD99DB1F9D0FC374754" bold="true" box="[151,932,1626,1653]" pageId="33" pageNumber="88">
Key to species of
<taxonomicName id="B9C64D1DFFCDFFD99C48F9D1FD854754" authorityName="Berlese" authorityYear="1903" box="[366,534,1626,1653]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="4CB2EA8CFFCDFFD99C48F9D1FD854754" bold="true" box="[366,534,1626,1653]" italics="true" pageId="33" pageNumber="88">Cosmolaelaps</emphasis>
</taxonomicName>
occurring in
<collectingCountry id="06D1760EFFCDFFD99F99F9D1FC374754" box="[703,932,1626,1653]" name="South Korea" pageId="33" pageNumber="88">Republic of Korea</collectingCountry>
</emphasis>
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</paragraph>
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<subSubSection id="36DC6515FFCDFFDA9DB1F92BFA0E40D1" lastPageId="34" lastPageNumber="89" pageId="33" pageNumber="88" type="key">
<keyStep id="C5322E3BFFCDFFD99DB1F92BFA0F47F2" pageId="33" pageNumber="88">
<paragraph id="7E79369EFFCDFFD99DB1F92BFA0F4794" blockId="33.[151,1436,1696,2040]" box="[151,1436,1696,1718]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F92BFA0F4794" box="[151,1436,1696,1718]" pageId="33" pageNumber="88">1. Dorsal setae spatulate with or without a small basal protuberance............................................... 2</keyLead>
</paragraph>
<paragraph id="7E79369EFFCDFFD99DB1F936FA0F47F2" blockId="33.[151,1436,1696,2040]" box="[151,1436,1725,1747]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F936FA0F47F2" box="[151,1436,1725,1747]" pageId="33" pageNumber="88">- Dorsal setae awl-shaped, knife or scimitar-like.............................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="C5322E3BFFCDFFD99DB1F951FA0F462C" pageId="33" pageNumber="88">
<paragraph id="7E79369EFFCDFFD99DB1F951FA0F47CE" blockId="33.[151,1436,1696,2040]" box="[151,1436,1754,1776]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F951FA0F47CE" box="[151,1436,1754,1776]" pageId="33" pageNumber="88">2. Dorsal setae without a small basal protuberance............................................................. 3</keyLead>
</paragraph>
<paragraph id="7E79369EFFCDFFD99DB1F973FA0F462C" blockId="33.[151,1436,1696,2040]" box="[151,1436,1784,1806]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F973FA0F462C" box="[151,1436,1784,1806]" pageId="33" pageNumber="88">- Dorsal setae with a small basal protuberance................................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="C5322E3BFFCDFFD99DB1F89EFA0F4644" pageId="33" pageNumber="88">
<paragraph id="7E79369EFFCDFFD99DB1F89EFA0F460B" blockId="33.[151,1436,1696,2040]" box="[151,1436,1813,1835]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F89EFA0F460B" box="[151,1436,1813,1835]" pageId="33" pageNumber="88">
3. Swollen humps on tibia I and basitarsus IIIV, surface of genital shield covered with fine granules.......
<taxonomicName id="B9C64D1DFFCDFFD999C5F89EFAD9460B" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[1251,1354,1813,1834]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="88" phylum="Arthropoda" rank="species" species="oedipus" status="sp. nov.">
<emphasis id="4CB2EA8CFFCDFFD999C5F89EFAD9460B" box="[1251,1354,1813,1834]" italics="true" pageId="33" pageNumber="88">C. oedipus</emphasis>
</taxonomicName>
<emphasis id="4CB2EA8CFFCDFFD99876F89EFA0F460B" bold="true" box="[1360,1436,1813,1834]" pageId="33" pageNumber="88">
<taxonomicNameLabel id="578157F7FFCDFFD99876F89EFA0F460B" box="[1360,1436,1813,1834]" pageId="33" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="7E79369EFFCDFFD99DB1F8B8FA0F4644" blockId="33.[151,1436,1696,2040]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F8B8FA0F4644" pageId="33" pageNumber="88">
- Without any hump-like structures on legs, surface of genital shield covered with lineate reticulation...................................................................................................
<taxonomicName id="B9C64D1DFFCDFFD99989F8DBFAD94644" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[1199,1354,1872,1893]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="88" phylum="Arthropoda" rank="species" species="sansungensis" status="sp. nov.">
<emphasis id="4CB2EA8CFFCDFFD99989F8DBFAD94644" box="[1199,1354,1872,1893]" italics="true" pageId="33" pageNumber="88">C. sansungensis</emphasis>
</taxonomicName>
<emphasis id="4CB2EA8CFFCDFFD99876F8DBFA0F4644" bold="true" box="[1360,1436,1872,1893]" pageId="33" pageNumber="88">
<taxonomicNameLabel id="578157F7FFCDFFD99876F8DBFA0F4644" box="[1360,1436,1872,1893]" pageId="33" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<keyLead id="C53795ABFFCDFFD99DB1F8E6FA0F46A2" box="[151,1436,1901,1923]" pageId="33" pageNumber="88">
4. 41 pairs of dorsal setae spatulate sickle-shaped........................................
<taxonomicName id="B9C64D1DFFCDFFD999ACF8E6FA0F46A2" authority="Ma &amp; Lin" authorityName="Ma &amp; Lin" authorityYear="2009" box="[1162,1436,1901,1923]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="88" phylum="Arthropoda" rank="species" species="robustochaetes">
<emphasis id="4CB2EA8CFFCDFFD999ACF8E6FAA546A3" box="[1162,1334,1901,1922]" italics="true" pageId="33" pageNumber="88">C. robustochaetes</emphasis>
Ma &amp; Lin
</taxonomicName>
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</paragraph>
<paragraph id="7E79369EFFCDFFD99DB1F800FA0F4681" blockId="33.[151,1436,1696,2040]" box="[151,1436,1930,1952]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F800FA0F4681" box="[151,1436,1930,1952]" pageId="33" pageNumber="88">- 40 pairs of dorsal setae spatulate with round, serrated margin................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="C5322E3BFFCDFFD99DB1F82CFA0F46FB" pageId="33" pageNumber="88">
<paragraph id="7E79369EFFCDFFD99DB1F82CFA0F469C" blockId="33.[151,1436,1696,2040]" box="[151,1436,1959,1982]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F82CFA0F469C" box="[151,1436,1959,1982]" pageId="33" pageNumber="88">
5. Dorsal shield with six accessory setae, sternal setae
<emphasis id="4CB2EA8CFFCDFFD99FF3F823FD62469C" box="[725,753,1960,1981]" italics="true" pageId="33" pageNumber="88">st1</emphasis>
on the shield..........................
<taxonomicName id="B9C64D1DFFCDFFD9998DF823FAD9469C" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[1195,1354,1960,1981]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="88" phylum="Arthropoda" rank="species" species="hallasanensis" status="sp. nov.">
<emphasis id="4CB2EA8CFFCDFFD9998DF823FAD9469C" box="[1195,1354,1960,1981]" italics="true" pageId="33" pageNumber="88">C. hallasanensis</emphasis>
</taxonomicName>
<emphasis id="4CB2EA8CFFCDFFD99876F823FA0F469C" bold="true" box="[1360,1436,1960,1981]" pageId="33" pageNumber="88">
<taxonomicNameLabel id="578157F7FFCDFFD99876F823FA0F469C" box="[1360,1436,1960,1981]" pageId="33" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="7E79369EFFCDFFD99DB1F84EFA0F46FB" blockId="33.[151,1436,1696,2040]" box="[151,1436,1989,2011]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F84EFA0F46FB" box="[151,1436,1989,2011]" pageId="33" pageNumber="88">
- Dorsal shield with 10 accessory setae, sternal setae
<emphasis id="4CB2EA8CFFCDFFD99FF7F84EFD7E46FB" box="[721,749,1989,2010]" italics="true" pageId="33" pageNumber="88">st1</emphasis>
off the sternal shield....................
<taxonomicName id="B9C64D1DFFCDFFD99988F84EFAD946FB" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[1198,1354,1989,2010]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="88" phylum="Arthropoda" rank="species" species="cavosternalis" status="sp. nov.">
<emphasis id="4CB2EA8CFFCDFFD99988F84EFAD946FB" box="[1198,1354,1989,2010]" italics="true" pageId="33" pageNumber="88">C. cavosternalis</emphasis>
</taxonomicName>
<emphasis id="4CB2EA8CFFCDFFD99876F84EFA0F46FB" bold="true" box="[1360,1436,1989,2010]" pageId="33" pageNumber="88">
<taxonomicNameLabel id="578157F7FFCDFFD99876F84EFA0F46FB" box="[1360,1436,1989,2010]" pageId="33" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C5322E3BFFCDFFDA9DB1F869FA0F418C" lastPageId="34" lastPageNumber="89" pageId="33" pageNumber="88">
<paragraph id="7E79369EFFCDFFD99DB1F869FA0F46D9" blockId="33.[151,1436,1696,2040]" box="[151,1436,2018,2040]" pageId="33" pageNumber="88">
<keyLead id="C53795ABFFCDFFD99DB1F869FA0F46D9" box="[151,1436,2018,2040]" pageId="33" pageNumber="88">
6. Dorsal setae short and slender, awl-shaped..................................................
<taxonomicName id="B9C64D1DFFCDFFD999FDF868FA0F46D9" authority="(Samsinak)" baseAuthorityName="Samsinak" box="[1243,1436,2018,2040]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="88" phylum="Arthropoda" rank="species" species="hrdyi">
<emphasis id="4CB2EA8CFFCDFFD999FDF868FABA46D6" box="[1243,1321,2018,2040]" italics="true" pageId="33" pageNumber="88">C. hrdyi</emphasis>
(Samšiňák)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7E79369EFFCEFFDA9DB1FF13FA0F418C" blockId="34.[151,1437,151,496]" box="[151,1436,151,173]" pageId="34" pageNumber="89">
<keyLead id="C53795ABFFCEFFDA9DB1FF13FA0F418C" box="[151,1436,151,173]" pageId="34" pageNumber="89">- Dorsal setae relatively expanded, knife or scimitar-like, with conspicuous basal protuberance......................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="C5322E3BFFCEFFDA9DB1FF3EFA0F4003" pageId="34" pageNumber="89">
<paragraph id="7E79369EFFCEFFDA9DB1FF3EFA0F41C9" blockId="34.[151,1437,151,496]" pageId="34" pageNumber="89">
<keyLead id="C53795ABFFCEFFDA9DB1FF3EFA0F41C9" pageId="34" pageNumber="89">
7. Dorsal setae knife-like, not elongate, at most reaching base of following setae, femora II and IV with spatulate setae.............................................................................................
<taxonomicName id="B9C64D1DFFCEFFDA99C4FF59FA0F41C9" authority="(Michael)" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[1250,1436,210,232]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="34" pageNumber="89" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="4CB2EA8CFFCEFFDA99C4FF59FAA441C9" box="[1250,1335,210,232]" italics="true" pageId="34" pageNumber="89">C. vacua</emphasis>
(Michael)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7E79369EFFCEFFDA9DB1FF7BFA0F4003" blockId="34.[151,1437,151,496]" pageId="34" pageNumber="89">
<keyLead id="C53795ABFFCEFFDA9DB1FF7BFA0F4003" pageId="34" pageNumber="89">- Dorsal setae scimitar-like, long, almost all setae reaching well past base of following setae, femora II and IV without spatulate setae............................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="C5322E3BFFCEFFDA9DB1FEA1FA0F407C" pageId="34" pageNumber="89">
<paragraph id="7E79369EFFCEFFDA9DB1FEA1FA0F401E" blockId="34.[151,1437,151,496]" box="[151,1436,298,320]" pageId="34" pageNumber="89">
<keyLead id="C53795ABFFCEFFDA9DB1FEA1FA0F401E" box="[151,1436,298,320]" pageId="34" pageNumber="89">
8. Dorsal setae
<emphasis id="4CB2EA8CFFCEFFDA9C78FEA1FEE4401E" box="[350,375,298,319]" italics="true" pageId="34" pageNumber="89">Z5</emphasis>
shorter than
<emphasis id="4CB2EA8CFFCEFFDA9CD3FEA1FD9F401E" box="[501,524,298,319]" italics="true" pageId="34" pageNumber="89">J5</emphasis>
(ratio of
<emphasis id="4CB2EA8CFFCEFFDA9F40FEA1FDEF401E" box="[614,636,298,319]" italics="true" pageId="34" pageNumber="89">J5</emphasis>
/
<emphasis id="4CB2EA8CFFCEFFDA9FA5FEA1FD0F401E" box="[643,668,298,319]" italics="true" pageId="34" pageNumber="89">Z5</emphasis>
≈ 1.5)...........................................
<taxonomicName id="B9C64D1DFFCEFFDA99CEFEA1FA0F401E" authority="(Gu)" baseAuthorityName="Gu" box="[1256,1436,298,319]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="34" pageNumber="89" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="4CB2EA8CFFCEFFDA99CEFEA1FAFA401E" box="[1256,1385,298,319]" italics="true" pageId="34" pageNumber="89">C. chianensis</emphasis>
(Gu)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7E79369EFFCEFFDA9DB1FEC3FA0F407C" blockId="34.[151,1437,151,496]" box="[151,1436,327,349]" pageId="34" pageNumber="89">
<keyLead id="C53795ABFFCEFFDA9DB1FEC3FA0F407C" box="[151,1436,327,349]" pageId="34" pageNumber="89">
- Dorsal setae
<emphasis id="4CB2EA8CFFCEFFDA9C78FEC3FEE4407C" box="[350,375,328,349]" italics="true" pageId="34" pageNumber="89">Z5</emphasis>
longer than
<emphasis id="4CB2EA8CFFCEFFDA9CD6FEC3FD95407C" box="[496,518,328,349]" italics="true" pageId="34" pageNumber="89">J5</emphasis>
........................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C5322E3BFFCEFFDA9DB1FEEEFA0F4094" pageId="34" pageNumber="89">
<paragraph id="7E79369EFFCEFFDA9DB1FEEEFA0F40B9" blockId="34.[151,1437,151,496]" pageId="34" pageNumber="89">
<keyLead id="C53795ABFFCEFFDA9DB1FEEEFA0F40B9" pageId="34" pageNumber="89">
9. Post-anal seta thickened, sparsely barbed and apically hooked, inserted on a small tubercle,
<emphasis id="4CB2EA8CFFCEFFDA997FFEEEFBE8405B" box="[1113,1147,357,378]" italics="true" pageId="34" pageNumber="89">Jv5</emphasis>
on opisthogastric cuticle well thickened and apically hooked.................................................
<taxonomicName id="B9C64D1DFFCEFFDA9977FE09FA0F40B9" authority="Keum, Jung &amp; Joharchi" authorityName="Keum, Jung &amp; Joharchi" box="[1105,1436,386,408]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="34" pageNumber="89" phylum="Arthropoda" rank="species" species="sejongi">
<emphasis id="4CB2EA8CFFCEFFDA9977FE09FB2140B9" box="[1105,1202,386,408]" italics="true" pageId="34" pageNumber="89">C. sejongi</emphasis>
Keum, Jung &amp; Joharchi
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7E79369EFFCEFFDA9DB1FE2BFA0F4094" blockId="34.[151,1437,151,496]" box="[151,1436,415,437]" pageId="34" pageNumber="89">
<keyLead id="C53795ABFFCEFFDA9DB1FE2BFA0F4094" box="[151,1436,415,437]" pageId="34" pageNumber="89">
- Post-anal seta and
<emphasis id="4CB2EA8CFFCEFFDA9CB3FE2BFE244094" box="[405,439,416,437]" italics="true" pageId="34" pageNumber="89">Jv5</emphasis>
on opisthogastric cuticle simple, slightly thickened....................................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C5322E3BFFCEFFDA9DB1FE36FA0E40D1" pageId="34" pageNumber="89">
<paragraph id="7E79369EFFCEFFDA9DB1FE36FA0E40F3" blockId="34.[151,1437,151,496]" box="[151,1437,445,467]" pageId="34" pageNumber="89">
<keyLead id="C53795ABFFCEFFDA9DB1FE36FA0E40F3" box="[151,1437,445,467]" pageId="34" pageNumber="89">
10. Genital shield notably expanded behind the
<emphasis id="4CB2EA8CFFCEFFDA9FAEFE36FD3740F3" box="[648,676,445,466]" italics="true" pageId="34" pageNumber="89">st5</emphasis>
, epistome with apical projection medially..........
<taxonomicName id="B9C64D1DFFCEFFDA999EFE36FA0E40F3" authority="(Karg)" authorityName="Karg" authorityYear="1985" baseAuthorityName="Karg" baseAuthorityYear="1985" box="[1208,1437,445,466]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="34" pageNumber="89" phylum="Arthropoda" rank="species" species="brevipedestra">
<emphasis id="4CB2EA8CFFCEFFDA999EFE36FAC440F3" box="[1208,1367,445,466]" italics="true" pageId="34" pageNumber="89">C. brevipedestra</emphasis>
(Karg)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7E79369EFFCEFFDA9DB1FE50FA0E40D1" blockId="34.[151,1437,151,496]" box="[151,1437,474,496]" pageId="34" pageNumber="89">
<keyLead id="C53795ABFFCEFFDA9DB1FE50FA0E40D1" box="[151,1437,474,496]" pageId="34" pageNumber="89">
- Genital shield not notably expanded, epistome with round serrated margin......................
<taxonomicName id="B9C64D1DFFCEFFDA999AFE51FA0E40D1" authority="(Ishikawa)" baseAuthorityName="Ishikawa" box="[1212,1437,474,496]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="34" pageNumber="89" phylum="Arthropoda" rank="species" species="hortensis">
<emphasis id="4CB2EA8CFFCEFFDA999AFE51FAA340CE" box="[1212,1328,474,495]" italics="true" pageId="34" pageNumber="89">C. hortensis</emphasis>
(Ishikawa)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,786 @@
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<mods:title id="24984E181532C13026B5336FD257421D">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from the Republic of Korea, with a review of the Korean species of the genus</mods:title>
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<mods:namePart id="C861FE643F5DD6959E039B60DE5DE77F">Oh, Jaeseok</mods:namePart>
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<mods:namePart id="62A4064A2406877BF1A04199293519A6">Joharchi, Omid</mods:namePart>
<mods:affiliation id="71155291FBEB8E9923E528E249292953">Anatis Bioprotection Inc., Saint-Jacques-de-Mineur, Québec, J 0 J 1 Z 0, Canada &amp; Agriculture Science and Technology Institute, Andong National University, Andong, Republic of Korea</mods:affiliation>
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<mods:namePart id="EA4DF3EE1A5DC5247F93F4B953DF4ED2">Lee, Seunghwan</mods:namePart>
<mods:affiliation id="DBAEA8D5A3A1FE41BB0574DED981F47D">Research Institute of Agriculture and Life Sciences, Seoul National University, Seoul, Republic of Korea</mods:affiliation>
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<mods:namePart id="DFCB7F7C9BC3B92D35CD3A2E882781D1">Jung, Chuleui</mods:namePart>
<mods:affiliation id="61E9C94106EC54D94BDB207E94EE5DDE">Department of Plant medicals, Andong National University, Andong, Republic of Korea &amp; Agriculture Science and Technology Institute, Andong National University, Andong, Republic of Korea</mods:affiliation>
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<treatment id="F66F8788FFE6FFEA9DB1FF1CFC2242E8" LSID="urn:lsid:plazi:treatment:F66F8788FFE6FFEA9DB1FF1CFC2242E8" httpUri="http://treatment.plazi.org/id/F66F8788FFE6FFEA9DB1FF1CFC2242E8" lastPageId="18" lastPageNumber="73" pageId="10" pageNumber="65">
<subSubSection id="36DC6515FFE6FFF29DB1FF1CFDC34190" box="[151,592,151,178]" pageId="10" pageNumber="65" type="nomenclature">
<paragraph id="7E79369EFFE6FFF29DB1FF1CFDC34190" blockId="10.[151,592,151,214]" box="[151,592,151,178]" pageId="10" pageNumber="65">
<heading id="253181F2FFE6FFF29DB1FF1CFDC34190" bold="true" box="[151,592,151,178]" fontSize="11" level="1" pageId="10" pageNumber="65" reason="1">
<emphasis id="4CB2EA8CFFE6FFF29DB1FF1CFDC34190" bold="true" box="[151,592,151,178]" pageId="10" pageNumber="65">
<taxonomicName id="B9C64D1DFFE6FFF29DB1FF1CFE784193" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[151,491,151,178]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="hallasanensis" status="sp. nov.">
<emphasis id="4CB2EA8CFFE6FFF29DB1FF1CFE784193" bold="true" box="[151,491,151,178]" italics="true" pageId="10" pageNumber="65">Cosmolaelaps hallasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="578157F7FFE6FFF29CD4FF1CFDC34190" box="[498,592,151,177]" pageId="10" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="36DC6515FFE6FFF29DB1FF30FD9A408C" pageId="10" pageNumber="65" type="description">
<paragraph id="7E79369EFFE6FFF29DB1FF30FED841F7" blockId="10.[151,592,151,214]" box="[151,331,187,214]" pageId="10" pageNumber="65">
(
<figureCitation id="E6FD2A1BFFE6FFF29DB9FF30FED041F7" box="[159,323,187,214]" captionStart-0="FIGURES 2527" captionStart-1="FIGURES 2830" captionStart-2="FIGURES 3138" captionStart-3="FIGURES 3942" captionStart-4="FIGURES 4346" captionStart-5="FIGURES 4752" captionStartId-0="11.[152,270,1943,1967]" captionStartId-1="12.[152,270,1447,1471]" captionStartId-2="13.[152,270,1938,1962]" captionStartId-3="14.[152,270,1493,1517]" captionStartId-4="16.[152,270,1943,1967]" captionStartId-5="17.[152,270,1943,1967]" captionTargetBox-0="[191,1396,196,1904]" captionTargetBox-1="[168,1418,198,1406]" captionTargetBox-2="[184,1402,185,1909]" captionTargetBox-3="[167,1420,197,1453]" captionTargetBox-4="[183,1408,188,1913]" captionTargetBox-5="[197,1396,197,1903]" captionTargetId-0="figure-17@11.[176,1411,181,1919]" captionTargetId-1="figure-26@12.[151,1436,181,1423]" captionTargetId-2="figure-17@13.[181,1406,181,1915]" captionTargetId-3="figure-36@14.[151,1436,181,1469]" captionTargetId-4="figure-26@16.[177,1411,181,1920]" captionTargetId-5="figure-17@17.[182,1405,181,1919]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="16" captionTargetPageId-5="17" captionText-0="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." captionText-1="FIGURES 2830. Cosmolaelaps hallasanensis sp. nov., adult female: 28—Ventral gnathosoma; 29—Epistome; 30— Chelicera." captionText-2="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." captionText-3="FIGURES 3942. Cosmolaelaps hallasanensis sp. nov., adult female: 39—Leg I; 40—Leg II; 41—Leg III; 42—Leg IV." captionText-4="FIGURES 4346. Cosmolaelaps hallasanensis sp. nov., adult male: 43—Line drawing plate of ventral idiosoma; 44—Line drawing plate of chelicera; 45—DIC micrographs of ventral idiosoma; 46—DIC micrographs of femur II." captionText-5="FIGURES 4752. Cosmolaelaps hallasanensis sp. nov., deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.14703930" figureDoi-1="http://doi.org/10.5281/zenodo.14703932" figureDoi-2="http://doi.org/10.5281/zenodo.14703934" figureDoi-3="http://doi.org/10.5281/zenodo.14703938" figureDoi-4="http://doi.org/10.5281/zenodo.14703941" figureDoi-5="http://doi.org/10.5281/zenodo.14703943" httpUri-0="https://zenodo.org/record/14703930/files/figure.png" httpUri-1="https://zenodo.org/record/14703932/files/figure.png" httpUri-2="https://zenodo.org/record/14703934/files/figure.png" httpUri-3="https://zenodo.org/record/14703938/files/figure.png" httpUri-4="https://zenodo.org/record/14703941/files/figure.png" httpUri-5="https://zenodo.org/record/14703943/files/figure.png" pageId="10" pageNumber="65">Figures 2552</figureCitation>
)
</paragraph>
<paragraph id="7E79369EFFE6FFF29DB1FE88FD9A408C" blockId="10.[151,1437,259,789]" pageId="10" pageNumber="65">
<emphasis id="4CB2EA8CFFE6FFF29DB1FE88FE0C403C" bold="true" box="[151,415,259,285]" pageId="10" pageNumber="65">Specimens examined.</emphasis>
<typeStatus id="A17D883CFFE6FFF29C88FE88FD85403C" box="[430,534,259,285]" pageId="10" pageNumber="65" type="holotype">Holotype</typeStatus>
. female, Dosun-dong, Seogwipo-si, Hallasan,
<collectingRegion id="BC02F87CFFE6FFF2996AFE88FB32403C" box="[1100,1185,259,285]" country="South Korea" name="Jeju" pageId="10" pageNumber="65">Jeju-do</collectingRegion>
,
<collectingCountry id="06D1760EFFE6FFF29993FE88FA0B403F" box="[1205,1432,259,286]" name="South Korea" pageId="10" pageNumber="65">Republic of Korea</collectingCountry>
,
<geoCoordinate id="1BF25059FFE6FFF29DB1FEACFEAD4063" box="[151,318,295,322]" degrees="33" direction="north" minutes="20" orientation="latitude" pageId="10" pageNumber="65" precision="1" seconds="58.51" value="33.349586">33°20'58.51&quot;N</geoCoordinate>
<geoCoordinate id="1BF25059FFE6FFF29C6DFEACFE6D4060" box="[331,510,295,321]" degrees="126" direction="east" minutes="30" orientation="longitude" pageId="10" pageNumber="65" precision="1" seconds="01.11" value="126.500305">126°30'01.11&quot;E</geoCoordinate>
, alt.
<quantity id="B93E9B7BFFE6FFF29F19FEACFD084063" box="[575,667,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="10" pageNumber="65" unit="m" value="1350.0">1350 m</quantity>
,
<date id="0A78105EFFE6FFF29F8BFEACFCC54060" box="[685,854,295,321]" pageId="10" pageNumber="65" value="2019-04-13">13 April 2019</date>
, C. Jung and J. Oh coll., soil in the tree cavity, NIBRIV0000862920.
<typeStatus id="A17D883CFFE6FFF29CB7FEC7FE6D4047" box="[401,510,332,358]" pageId="10" pageNumber="65" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="68C0FD17FFE6FFF29F2DFEC0FD144044" box="[523,647,331,357]" count="26" pageId="10" pageNumber="65" type="female">26 females</specimenCount>
,
<specimenCount id="68C0FD17FFE6FFF29FB7FEC0FD664044" box="[657,757,331,357]" count="10" pageId="10" pageNumber="65" type="male">10 males</specimenCount>
and four deutonymph, San 98, Gyorae-ri, Jocheon-eup, Jejusi,
<collectingCountry id="06D1760EFFE6FFF29D9FFEE4FE1F40AB" box="[185,396,367,394]" name="South Korea" pageId="10" pageNumber="65">Republic of Korea</collectingCountry>
,
<date id="0A78105EFFE6FFF29CBEFEE4FDBE40A8" box="[408,557,367,393]" pageId="10" pageNumber="65" value="2021-07-19">19 July 2021</date>
, J. Oh coll., in the nest of
<taxonomicName id="B9C64D1DFFE6FFF29E7FFEE4FBB940AB" box="[857,1066,367,394]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4CB2EA8CFFE6FFF29E7FFEE4FB9040A8" box="[857,1027,367,393]" italics="true" pageId="10" pageNumber="65">Aphaenogaster</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="B9C64D1DFFE6FFF2991FFEE4FB4B40A8" box="[1081,1240,367,393]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="B9C64D1DFFE6FFF299C5FEE4FAF640A8" box="[1251,1381,367,393]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
) and
<specimenCount id="68C0FD17FFE6FFF29DB1FE1FFE93408C" box="[151,256,403,430]" count="1" pageId="10" pageNumber="65" type="male">one male</specimenCount>
, same data as
<typeStatus id="A17D883CFFE6FFF29C86FE18FD95408C" box="[416,518,403,429]" pageId="10" pageNumber="65" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="36DC6515FFE6FFF29DE1FE3CFEC74234" pageId="10" pageNumber="65" type="diagnosis">
<paragraph id="7E79369EFFE6FFF29DE1FE3CFEC74234" blockId="10.[151,1437,259,789]" pageId="10" pageNumber="65">
<emphasis id="4CB2EA8CFFE6FFF29DE1FE3CFE0440F0" bold="true" box="[199,407,439,465]" pageId="10" pageNumber="65">Diagnosis (adult)</emphasis>
Dorsal shield reticulate, with 40 pairs of spatulate-mucronate setae (except
<emphasis id="4CB2EA8CFFE6FFF2983BFE3CFAA040F0" box="[1309,1331,439,465]" italics="true" pageId="10" pageNumber="65">j1</emphasis>
and
<emphasis id="4CB2EA8CFFE6FFF29855FE3CFA1F40F0" box="[1395,1420,439,465]" italics="true" pageId="10" pageNumber="65">z1</emphasis>
), including three pairs of
<emphasis id="4CB2EA8CFFE6FFF29C8DFE57FE5440D4" box="[427,455,476,501]" italics="true" pageId="10" pageNumber="65">Zx</emphasis>
setae and six unpaired
<emphasis id="4CB2EA8CFFE6FFF29FF1FE57FD6340D4" box="[727,752,476,501]" italics="true" pageId="10" pageNumber="65">Jx</emphasis>
setae; presternal area poorly sclerotised, sternal shield with lineate-reticulate ornamentation on lateral surface, distinctly punctate centrally, ratio of shield length/width (at broadest level) ≃ 0.48; genital shield flask-shaped, widened posteriorly behind setae
<emphasis id="4CB2EA8CFFE6FFF29944FDAFFB10431F" box="[1122,1155,548,574]" italics="true" pageId="10" pageNumber="65">st5</emphasis>
, surface reticulated with irregular, mostly longitudinal lines in anterior region, posteriorly with irregular polygonal pattern, basally composed of large cells, shield ratio of length/width (at broadest level) ≃ 1.6, anal shield slightly longer than wide, with ratio of length/width (at broadest level) ≃ 1.18, post-anal seta thickened; opisthogastric setae spatulate, except
<emphasis id="4CB2EA8CFFE6FFF2981CFD04FA064388" box="[1338,1429,655,682]" italics="true" pageId="10" pageNumber="65">Jv1Jv3</emphasis>
,
<emphasis id="4CB2EA8CFFE6FFF29DB1FD38FF5243EC" box="[151,193,691,717]" italics="true" pageId="10" pageNumber="65">Zv1</emphasis>
, and
<emphasis id="4CB2EA8CFFE6FFF29C23FD38FEBC43EC" box="[261,303,691,717]" italics="true" pageId="10" pageNumber="65">Zv2</emphasis>
; fixed digit of chelicera with 1012 teeth of various sizes. Most setae on legs spatulate. In male, holoventral shield bearing 10 pairs of setae. Femur II in both female and male with strong spur-like seta on the ventral side (
<emphasis id="4CB2EA8CFFE6FFF29C0FFD77FED04234" box="[297,323,764,789]" italics="true" pageId="10" pageNumber="65">av</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="36DC6515FFE6FFEA9DB1FCECFAA441F4" lastPageId="18" lastPageNumber="73" pageId="10" pageNumber="65" type="description">
<paragraph id="7E79369EFFE6FFF29DB1FCECFE4C42A0" blockId="10.[151,479,871,934]" box="[151,479,871,897]" pageId="10" pageNumber="65">
<emphasis id="4CB2EA8CFFE6FFF29DB1FCECFE4C42A0" bold="true" box="[151,479,871,897]" pageId="10" pageNumber="65">Description (adult female)</emphasis>
</paragraph>
<paragraph id="7E79369EFFE6FFF29DB1FC00FED84287" blockId="10.[151,479,871,934]" box="[151,331,907,934]" pageId="10" pageNumber="65">
(
<figureCitation id="E6FD2A1BFFE6FFF29DB9FC00FED04287" box="[159,323,907,934]" captionStart-0="FIGURES 2527" captionStart-1="FIGURES 2830" captionStart-2="FIGURES 3138" captionStart-3="FIGURES 3942" captionStartId-0="11.[152,270,1943,1967]" captionStartId-1="12.[152,270,1447,1471]" captionStartId-2="13.[152,270,1938,1962]" captionStartId-3="14.[152,270,1493,1517]" captionTargetBox-0="[191,1396,196,1904]" captionTargetBox-1="[168,1418,198,1406]" captionTargetBox-2="[184,1402,185,1909]" captionTargetBox-3="[167,1420,197,1453]" captionTargetId-0="figure-17@11.[176,1411,181,1919]" captionTargetId-1="figure-26@12.[151,1436,181,1423]" captionTargetId-2="figure-17@13.[181,1406,181,1915]" captionTargetId-3="figure-36@14.[151,1436,181,1469]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." captionText-1="FIGURES 2830. Cosmolaelaps hallasanensis sp. nov., adult female: 28—Ventral gnathosoma; 29—Epistome; 30— Chelicera." captionText-2="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." captionText-3="FIGURES 3942. Cosmolaelaps hallasanensis sp. nov., adult female: 39—Leg I; 40—Leg II; 41—Leg III; 42—Leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.14703930" figureDoi-1="http://doi.org/10.5281/zenodo.14703932" figureDoi-2="http://doi.org/10.5281/zenodo.14703934" figureDoi-3="http://doi.org/10.5281/zenodo.14703938" httpUri-0="https://zenodo.org/record/14703930/files/figure.png" httpUri-1="https://zenodo.org/record/14703932/files/figure.png" httpUri-2="https://zenodo.org/record/14703934/files/figure.png" httpUri-3="https://zenodo.org/record/14703938/files/figure.png" pageId="10" pageNumber="65">Figures 2542</figureCitation>
)
</paragraph>
<paragraph id="7E79369EFFE6FFF29DB1FC5BFE5C42C9" blockId="10.[151,463,976,1000]" box="[151,463,976,1000]" pageId="10" pageNumber="65">
<specimenCount id="68C0FD17FFE6FFF29DB1FC5BFEBA42C9" box="[151,297,976,1000]" count="6" pageId="10" pageNumber="65" type="generic">Six specimens</specimenCount>
were measured.
</paragraph>
<paragraph id="7E79369EFFE6FFF29DB1FB84FAFA4420" blockId="10.[151,1437,1039,2038]" pageId="10" pageNumber="65">
<emphasis id="4CB2EA8CFFE6FFF29DB1FB84FECA4508" bold="true" box="[151,345,1039,1065]" pageId="10" pageNumber="65">Dorsal idiosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFE6FFF29C4FFB84FD9E4508" box="[361,525,1039,1066]" captionStart="FIGURES 2527" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[191,1396,196,1904]" captionTargetId="figure-17@11.[176,1411,181,1919]" captionTargetPageId="11" captionText="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="10" pageNumber="65">Figures 25, 26</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE6FFF29F14FB84FDC24508" box="[562,593,1039,1065]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="10" pageNumber="65">31</figureCitation>
). Dorsal shield oval-shaped, 737790 long, 523538 wide, covering entire idiosoma dorsally; shield with distinct reticulation, bearing 40 pairs of almost uniform spatulate-mucronate setae (with a discernable basal asymmetric protuberance): 22 pairs of podonotal setae and 18 pairs of opisthonotal setae, including three pairs of
<emphasis id="4CB2EA8CFFE6FFF29C85FBF7FE2C45B4" box="[419,447,1148,1173]" italics="true" pageId="10" pageNumber="65">Zx</emphasis>
setae between
<emphasis id="4CB2EA8CFFE6FFF29F4CFBF7FDE545B4" box="[618,630,1148,1173]" italics="true" pageId="10" pageNumber="65">J</emphasis>
and
<emphasis id="4CB2EA8CFFE6FFF29F8BFBF7FD2E45B4" box="[685,701,1148,1173]" italics="true" pageId="10" pageNumber="65">Z</emphasis>
setae and six unpaired supernumerary seta
<emphasis id="4CB2EA8CFFE6FFF29983FBF7FB2D45B4" box="[1189,1214,1148,1173]" italics="true" pageId="10" pageNumber="65">Jx</emphasis>
between
<emphasis id="4CB2EA8CFFE6FFF2980CFBF0FAD745B4" box="[1322,1348,1147,1173]" italics="true" pageId="10" pageNumber="65">J1</emphasis>
and
<emphasis id="4CB2EA8CFFE6FFF2985DFBF7FA0645B7" box="[1403,1429,1148,1174]" italics="true" pageId="10" pageNumber="65">J5</emphasis>
, dorsal setae mostly homogeneous in length and thickness, barely reaching base of next posterior setae (5869), except
<emphasis id="4CB2EA8CFFE6FFF29DCDFB48FE9245FC" box="[235,257,1219,1245]" italics="true" pageId="10" pageNumber="65">j1</emphasis>
(4851) cunate-tricarinate and
<emphasis id="4CB2EA8CFFE6FFF29F54FB48FD1845FC" box="[626,651,1219,1245]" italics="true" pageId="10" pageNumber="65">z1</emphasis>
acuminate (3639) (
<figureCitation id="E6FD2A1BFFE6FFF29E59FB48FBB945FC" box="[895,1066,1219,1246]" captionStart="FIGURES 2527" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[191,1396,196,1904]" captionTargetId="figure-17@11.[176,1411,181,1919]" captionTargetPageId="11" captionText="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="10" pageNumber="65">Figures 25, 26</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE6FFF29970FB48FBE645FC" box="[1110,1141,1219,1245]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="10" pageNumber="65">31</figureCitation>
). Shield with 23 pairs of discernible pore-like structures, including 16 poroids and seven gland openings (
<emphasis id="4CB2EA8CFFE6FFF29906FB6CFBDA4420" box="[1056,1097,1255,1281]" italics="true" pageId="10" pageNumber="65">gd1</emphasis>
,
<emphasis id="4CB2EA8CFFE6FFF2997EFB6CFB114420" box="[1112,1154,1255,1281]" italics="true" pageId="10" pageNumber="65">gd2</emphasis>
,
<emphasis id="4CB2EA8CFFE6FFF299B6FB6CFB454420" box="[1168,1238,1255,1281]" italics="true" pageId="10" pageNumber="65">gd46</emphasis>
, and
<emphasis id="4CB2EA8CFFE6FFF29835FB6CFACB4420" box="[1299,1368,1255,1281]" italics="true" pageId="10" pageNumber="65">gd89</emphasis>
).
</paragraph>
<paragraph id="7E79369EFFE6FFF49DE1FA80FEF547DC" blockId="10.[151,1437,1039,2038]" lastBlockId="12.[151,1437,1583,2041]" lastPageId="12" lastPageNumber="67" pageId="10" pageNumber="65">
<emphasis id="4CB2EA8CFFE6FFF29DE1FA80FE1C4404" bold="true" box="[199,399,1291,1317]" pageId="10" pageNumber="65">Ventral idiosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFE6FFF29CBBFA80FD874404" box="[413,532,1291,1317]" captionStart="FIGURES 2527" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[191,1396,196,1904]" captionTargetId="figure-17@11.[176,1411,181,1919]" captionTargetPageId="11" captionText="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="10" pageNumber="65">Figures 27</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE6FFF29F10FA80FDEC4407" box="[566,639,1291,1318]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="10" pageNumber="65">3235</figureCitation>
). Tritosternum with paired pilose laciniae (119135), columnar base 48 60 long × 1622 wide; presternal area poorly sclerotised, sternal shield length 93112, narrowest between coxae II (123135), widest between coxae II and III (218230), with reticulate ornamentation throughout, except central and posterior parts distinctly punctate (
<figureCitation id="E6FD2A1BFFE6FFF29F33FAFCFD1F44B0" box="[533,652,1399,1425]" captionStart="FIGURES 2527" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[191,1396,196,1904]" captionTargetId="figure-17@11.[176,1411,181,1919]" captionTargetPageId="11" captionText="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="10" pageNumber="65">Figures 27</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE6FFF29F88FAFCFD5F44B0" box="[686,716,1399,1425]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="10" pageNumber="65">33</figureCitation>
), anterior margin of shield conspicuous, posterior margin slightly concave, anterolateral corners narrowly extending between coxae III, distally bearing gland pores
<emphasis id="4CB2EA8CFFE6FFF299D7FA10FA8A4494" box="[1265,1305,1435,1461]" italics="true" pageId="10" pageNumber="65">gvb</emphasis>
; endopodal elements between coxae II and III fused with shield, endopodal elements III/IV elongate, narrow and curved, shield bearing three pairs of smooth long setae (
<emphasis id="4CB2EA8CFFE6FFF29F79FA68FD1344DC" box="[607,640,1507,1533]" italics="true" pageId="10" pageNumber="65">st1</emphasis>
7080,
<emphasis id="4CB2EA8CFFE6FFF29FFFFA68FD6A44DC" box="[729,761,1507,1533]" italics="true" pageId="10" pageNumber="65">st2</emphasis>
6875,
<emphasis id="4CB2EA8CFFE6FFF29E75FA68FCE144DC" box="[851,882,1507,1533]" italics="true" pageId="10" pageNumber="65">st3</emphasis>
6676), extending beyond of next posterior setae, and two pairs of sub-oval poroids (
<emphasis id="4CB2EA8CFFE6FFF29F3FF98CFDA84700" box="[537,571,1543,1569]" italics="true" pageId="10" pageNumber="65">iv1</emphasis>
and
<emphasis id="4CB2EA8CFFE6FFF29F56F98CFD014700" box="[624,658,1543,1569]" italics="true" pageId="10" pageNumber="65">iv2</emphasis>
, mesal to setae
<emphasis id="4CB2EA8CFFE6FFF29E67F98CFCF24700" box="[833,865,1543,1569]" italics="true" pageId="10" pageNumber="65">st1</emphasis>
and between
<emphasis id="4CB2EA8CFFE6FFF29EDFF98CFB8A4700" box="[1017,1049,1543,1569]" italics="true" pageId="10" pageNumber="65">st2</emphasis>
and
<emphasis id="4CB2EA8CFFE6FFF29969F98CFBFC4700" box="[1103,1135,1543,1569]" italics="true" pageId="10" pageNumber="65">st3</emphasis>
, respectively). Metasternal setae
<emphasis id="4CB2EA8CFFE6FFF29DF3F9A0FF654764" box="[213,246,1579,1605]" italics="true" pageId="10" pageNumber="65">st4</emphasis>
(5561) and metasternal poroids located on soft integument. Genital shield flask-shaped, slightly expanded laterally behind setae
<emphasis id="4CB2EA8CFFE6FFF29CABF9DBFE3E474B" box="[397,429,1616,1642]" italics="true" pageId="10" pageNumber="65">st5</emphasis>
(
<figureCitation id="E6FD2A1BFFE6FFF29C9AF9C4FDA64748" box="[444,565,1615,1641]" captionStart="FIGURES 2527" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[191,1396,196,1904]" captionTargetId="figure-17@11.[176,1411,181,1919]" captionTargetPageId="11" captionText="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="10" pageNumber="65">Figures 27</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE6FFF29F7CF9C4FDEA4748" box="[602,633,1615,1641]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="10" pageNumber="65">34</figureCitation>
), length 259280, maximum width 169173, posterior margin rounded, anterior margin of shield convex, overlapping posterior area of sternal shield, reaching base of setae
<emphasis id="4CB2EA8CFFE6FFF2983DF9F8FAA847AC" box="[1307,1339,1651,1677]" italics="true" pageId="10" pageNumber="65">st3</emphasis>
, surface reticulated with irregular, mostly longitudinal lines in anterior region, posteriorly with irregular polygonal pattern, basally composed of large cells, bearing a pair of simple setae
<emphasis id="4CB2EA8CFFE6FFF29E6AF937FCFF47F7" box="[844,876,1724,1750]" italics="true" pageId="10" pageNumber="65">st5</emphasis>
(5867) inserted on lateral margins of shield, near level of posterior edge of coxae IV; paragenital poroids
<emphasis id="4CB2EA8CFFE6FFF29FD9F96BFCB247DB" box="[767,801,1760,1786]" italics="true" pageId="10" pageNumber="65">iv5</emphasis>
located on soft cuticle lateral to shield near seta
<emphasis id="4CB2EA8CFFE6FFF2981CF96BFAC947DB" box="[1338,1370,1760,1786]" italics="true" pageId="10" pageNumber="65">st5</emphasis>
. Anal shield pear-shaped, rounded anteriorly, length 124142, width 110120, anterior half lineate-reticulate, para-anal setae and post-anal seta subequal in length (3136), post-anal seta thickened, cribrum consisting of a terminal tuft with some irregular rows of spicules and a pair of anterior arms extending slightly beyond post-anal seta; anal gland pores (
<emphasis id="4CB2EA8CFFE6FFF29DC2F8E4FE9F46A8" box="[228,268,1903,1929]" italics="true" pageId="10" pageNumber="65">gv3</emphasis>
) on anterolateral margin of anal shield (
<figureCitation id="E6FD2A1BFFE6FFF29FEDF8E4FCD046A8" box="[715,835,1903,1929]" captionStart="FIGURES 2527" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[191,1396,196,1904]" captionTargetId="figure-17@11.[176,1411,181,1919]" captionTargetPageId="11" captionText="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="10" pageNumber="65">Figures 27</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE6FFF29E41F8E4FC1546AB" box="[871,902,1903,1930]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="10" pageNumber="65">35</figureCitation>
). Soft opisthogastric cuticle surrounding genital and anal shields with one pair of elongated metapodal plates (2333 long × 35 wide), six pairs of poroids (five
<emphasis id="4CB2EA8CFFE6FFF29DB1F833FF2A46F0" box="[151,185,1976,2001]" italics="true" pageId="10" pageNumber="65">ivo</emphasis>
;
<emphasis id="4CB2EA8CFFE6FFF29DEFF833FF7846F0" box="[201,235,1976,2001]" italics="true" pageId="10" pageNumber="65">ivp</emphasis>
) and 17 pairs of spatulate setae (3238) (
<emphasis id="4CB2EA8CFFE6FFF29FE3F83CFC6946F0" box="[709,1018,1975,2002]" italics="true" pageId="10" pageNumber="65">Jv1Jv5, Zv1 Zv5, R1R5,</emphasis>
and
<emphasis id="4CB2EA8CFFE6FFF29912F83CFB3446F0" box="[1076,1191,1975,2001]" italics="true" pageId="10" pageNumber="65">UR2UR3</emphasis>
), except
<emphasis id="4CB2EA8CFFE6FFF29837F83CFAF846F0" box="[1297,1387,1975,2001]" italics="true" pageId="10" pageNumber="65">Jv1Jv3</emphasis>
and
<emphasis id="4CB2EA8CFFE6FFF29DB1F850FF6846D4" box="[151,251,2011,2037]" italics="true" pageId="10" pageNumber="65">Zv1Zv2</emphasis>
setiform and slightly longer (4655) (
<figureCitation id="E6FD2A1BFFE6FFF29F89F850FCB846D4" box="[687,811,2011,2037]" captionStart="FIGURES 2527" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[191,1396,196,1904]" captionTargetId="figure-17@11.[176,1411,181,1919]" captionTargetPageId="11" captionText="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="10" pageNumber="65">Figures 27</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE6FFF29E69F850FCFF46D4" box="[847,876,2011,2037]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="10" pageNumber="65">32</figureCitation>
). Two exopodal platelets present between coxae IIIII and IIIIV, exopodal platelets between coxae IIIII small, subtriangular, those behind coxae IIIIV and parapodal platelets fused, surrounding coxa IV and bearing gland pore
<emphasis id="4CB2EA8CFFE0FFF49E00F9D8FCDD474C" box="[806,846,1619,1645]" italics="true" pageId="12" pageNumber="67">gv2</emphasis>
(
<figureCitation id="E6FD2A1BFFE0FFF49E7DF9D8FC42474C" box="[859,977,1619,1645]" captionStart="FIGURES 2527" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[191,1396,196,1904]" captionTargetId="figure-17@11.[176,1411,181,1919]" captionTargetPageId="11" captionText="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="12" pageNumber="67">Figures 27</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE0FFF49ED6F9D8FB9C474C" box="[1008,1039,1619,1645]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="12" pageNumber="67">32</figureCitation>
). Peritrematal shield well developed laterad peritremes, bearing one gland pore and one poroid at level near coxae IIIII, post-stigmatic extension of shield relatively wide, with two pairs of poroids and one pair of gland pores; anterior region of peritrematal shield fused to anterior margin of dorsal shield behind setae
<emphasis id="4CB2EA8CFFE0FFF49FCAF934FC9747F8" box="[748,772,1727,1753]" italics="true" pageId="12" pageNumber="67">z1</emphasis>
. Peritremes long, extending to mid-level of coxa I (near
<emphasis id="4CB2EA8CFFE0FFF4985CF934FA0047F8" box="[1402,1427,1727,1753]" italics="true" pageId="12" pageNumber="67">s1</emphasis>
) (
<figureCitation id="E6FD2A1BFFE0FFF49DB9F968FE8447DC" box="[159,279,1763,1789]" captionStart="FIGURES 2527" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[191,1396,196,1904]" captionTargetId="figure-17@11.[176,1411,181,1919]" captionTargetPageId="11" captionText="FIGURES 2527. Cosmolaelaps hallasanensis sp. nov., adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="12" pageNumber="67">Figures 27</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE0FFF49C1CF968FEC547DC" box="[314,342,1763,1789]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="12" pageNumber="67">32</figureCitation>
).
</paragraph>
<caption id="2AB96616FFE7FFF39DBEF81CFEED46F2" ID-DOI="http://doi.org/10.5281/zenodo.14703930" ID-Zenodo-Dep="14703930" httpUri="https://zenodo.org/record/14703930/files/figure.png" pageId="11" pageNumber="66" startId="11.[152,270,1943,1967]" targetBox="[191,1396,196,1904]" targetPageId="11" targetType="figure">
<paragraph id="7E79369EFFE7FFF39DBEF81CFEED46F2" blockId="11.[151,1436,1943,2003]" pageId="11" pageNumber="66">
<emphasis id="4CB2EA8CFFE7FFF39DBEF81CFD71468E" bold="true" box="[152,738,1943,1967]" pageId="11" pageNumber="66">
FIGURES 2527.
<taxonomicName id="B9C64D1DFFE7FFF39C46F81CFD16468E" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[352,645,1943,1967]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="hallasanensis" status="sp. nov.">
<emphasis id="4CB2EA8CFFE7FFF39C46F81CFD16468E" bold="true" box="[352,645,1943,1967]" italics="true" pageId="11" pageNumber="66">Cosmolaelaps hallasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="578157F7FFE7FFF39FADF81CFD4E468E" box="[651,733,1943,1967]" pageId="11" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
adult female: A—Dorsal idiosoma; 26—Some dorsal setae enlarged; 27—Ventral idiosoma.
</paragraph>
</caption>
<caption id="2AB96616FFE0FFF49DBEFA2CFE9344C2" ID-DOI="http://doi.org/10.5281/zenodo.14703932" ID-Zenodo-Dep="14703932" httpUri="https://zenodo.org/record/14703932/files/figure.png" pageId="12" pageNumber="67" startId="12.[152,270,1447,1471]" targetBox="[168,1418,198,1406]" targetPageId="12" targetType="figure">
<paragraph id="7E79369EFFE0FFF49DBEFA2CFE9344C2" blockId="12.[151,1436,1447,1507]" pageId="12" pageNumber="67">
<emphasis id="4CB2EA8CFFE0FFF49DBEFA2CFC96449E" bold="true" box="[152,773,1447,1471]" pageId="12" pageNumber="67">
FIGURES 2830.
<taxonomicName id="B9C64D1DFFE0FFF49C48FA2CFD09449E" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[366,666,1447,1471]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="67" phylum="Arthropoda" rank="species" species="hallasanensis" status="sp. nov.">
<emphasis id="4CB2EA8CFFE0FFF49C48FA2CFD09449E" bold="true" box="[366,666,1447,1471]" italics="true" pageId="12" pageNumber="67">Cosmolaelaps hallasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="578157F7FFE0FFF49F81FA2CFC92449E" box="[679,769,1447,1471]" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
adult female: 28—Ventral gnathosoma; 29—Epistome; 30— Chelicera.
</paragraph>
</caption>
<paragraph id="7E79369EFFE0FFF69DE1F88DFC1447CB" blockId="12.[151,1437,1583,2041]" lastBlockId="14.[151,1437,1599,2022]" lastPageId="14" lastPageNumber="69" pageId="12" pageNumber="67">
<emphasis id="4CB2EA8CFFE0FFF49DE1F88DFECC4601" bold="true" box="[199,351,1798,1824]" pageId="12" pageNumber="67">Gnathosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFE0FFF49C56F88CFD864600" box="[368,533,1799,1825]" captionStart="FIGURES 2830" captionStartId="12.[152,270,1447,1471]" captionTargetBox="[168,1418,198,1406]" captionTargetId="figure-26@12.[151,1436,181,1423]" captionTargetPageId="12" captionText="FIGURES 2830. Cosmolaelaps hallasanensis sp. nov., adult female: 28—Ventral gnathosoma; 29—Epistome; 30— Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703932" httpUri="https://zenodo.org/record/14703932/files/figure.png" pageId="12" pageNumber="67">Figures 2830</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE0FFF49F1AF88DFD154601" box="[572,646,1798,1824]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="12" pageNumber="67">3638</figureCitation>
). Anterior margin of epistome convex, irregularly denticulate (
<figureCitation id="E6FD2A1BFFE0FFF49877F88CFF244664" captionStart="FIGURES 2830" captionStartId="12.[152,270,1447,1471]" captionTargetBox="[168,1418,198,1406]" captionTargetId="figure-26@12.[151,1436,181,1423]" captionTargetPageId="12" captionText="FIGURES 2830. Cosmolaelaps hallasanensis sp. nov., adult female: 28—Ventral gnathosoma; 29—Epistome; 30— Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703932" httpUri="https://zenodo.org/record/14703932/files/figure.png" pageId="12" pageNumber="67">Figure 29</figureCitation>
). Hypostomal groove greatly widened, with six transverse rows of denticles, each row with 1035 tiny denticles, with smooth anterior and posterior transverse lines, groove wider anteriorly, progressively narrowing from anterior to posterior (
<figureCitation id="E6FD2A1BFFE0FFF49C00F8F8FE0046AC" box="[294,403,1907,1933]" captionStart="FIGURES 2830" captionStartId="12.[152,270,1447,1471]" captionTargetBox="[168,1418,198,1406]" captionTargetId="figure-26@12.[151,1436,181,1423]" captionTargetPageId="12" captionText="FIGURES 2830. Cosmolaelaps hallasanensis sp. nov., adult female: 28—Ventral gnathosoma; 29—Epistome; 30— Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703932" httpUri="https://zenodo.org/record/14703932/files/figure.png" pageId="12" pageNumber="67">Figure 28</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE0FFF49C91F8F9FE4446AD" box="[439,471,1906,1932]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="12" pageNumber="67">36</figureCitation>
). Hypostome with four pairs of smooth setae,
<emphasis id="4CB2EA8CFFE0FFF49EFCF8F8FC6546AC" box="[986,1014,1907,1933]" italics="true" pageId="12" pageNumber="67">h3</emphasis>
(8593)&gt;
<emphasis id="4CB2EA8CFFE0FFF49955F8F8FB1C46AC" box="[1139,1167,1907,1933]" italics="true" pageId="12" pageNumber="67">h1</emphasis>
(6773)&gt;
<emphasis id="4CB2EA8CFFE0FFF4982AF8F8FAB546AD" box="[1292,1318,1907,1932]" italics="true" pageId="12" pageNumber="67">pc</emphasis>
(6570)&gt;
<emphasis id="4CB2EA8CFFE0FFF49DB1F81CFF204690" box="[151,179,1943,1969]" italics="true" pageId="12" pageNumber="67">h2</emphasis>
(3035) (
<figureCitation id="E6FD2A1BFFE0FFF49C06F81CFE1E4690" box="[288,397,1943,1969]" captionStart="FIGURES 2830" captionStartId="12.[152,270,1447,1471]" captionTargetBox="[168,1418,198,1406]" captionTargetId="figure-26@12.[151,1436,181,1423]" captionTargetPageId="12" captionText="FIGURES 2830. Cosmolaelaps hallasanensis sp. nov., adult female: 28—Ventral gnathosoma; 29—Epistome; 30— Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703932" httpUri="https://zenodo.org/record/14703932/files/figure.png" pageId="12" pageNumber="67">Figure 28</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE0FFF49C89F81DFE5C4691" box="[431,463,1942,1968]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="12" pageNumber="67">36</figureCitation>
). Corniculi robust and horn-like, internal malae longer than corniculi, comprising a pair of pilose median projections, and 1012 lateral, longer, tentacle-like, smooth projections; labrum with pilose surface (
<figureCitation id="E6FD2A1BFFE0FFF49DB9F854FE9D46D8" box="[159,270,2015,2041]" captionStart="FIGURES 2830" captionStartId="12.[152,270,1447,1471]" captionTargetBox="[168,1418,198,1406]" captionTargetId="figure-26@12.[151,1436,181,1423]" captionTargetPageId="12" captionText="FIGURES 2830. Cosmolaelaps hallasanensis sp. nov., adult female: 28—Ventral gnathosoma; 29—Epistome; 30— Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703932" httpUri="https://zenodo.org/record/14703932/files/figure.png" pageId="12" pageNumber="67">Figure 28</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE0FFF49C12F855FEC646D9" box="[308,341,2014,2040]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="12" pageNumber="67">36</figureCitation>
); supralabral process not distinct. Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth; palpfemur with seta
<emphasis id="4CB2EA8CFFE2FFF69F8CF9B4FD534778" box="[682,704,1599,1625]" italics="true" pageId="14" pageNumber="69">al</emphasis>
paddle-like; palpgenu with
<emphasis id="4CB2EA8CFFE2FFF69ED3F9B4FB8A4778" box="[1013,1049,1599,1625]" italics="true" pageId="14" pageNumber="69">al1</emphasis>
and
<emphasis id="4CB2EA8CFFE2FFF6996BF9B4FBE24778" box="[1101,1137,1599,1625]" italics="true" pageId="14" pageNumber="69">al2</emphasis>
stout, blunt, thickened and spatulate; palptarsal apotele two-tined (
<figureCitation id="E6FD2A1BFFE2FFF69F74F9E8FD56475F" box="[594,709,1635,1662]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="14" pageNumber="69">Figure 38</figureCitation>
). Fixed digit of chelicera with an offset distal tooth (gabelzhan), followed by 1012 various sized teeth (mostly small), a setaceous pilus dentilis, dorsal cheliceral seta small and prostrate, arthrodial membrane with a rounded flap and normal filaments; cheliceral dorsal and lateral lyrifissures distinct; movable digit with two mid-sized teeth (
<figureCitation id="E6FD2A1BFFE2FFF69F98F944FCA447CB" box="[702,823,1743,1770]" captionStart="FIGURES 2830" captionStartId="12.[152,270,1447,1471]" captionTargetBox="[168,1418,198,1406]" captionTargetId="figure-26@12.[151,1436,181,1423]" captionTargetPageId="12" captionText="FIGURES 2830. Cosmolaelaps hallasanensis sp. nov., adult female: 28—Ventral gnathosoma; 29—Epistome; 30— Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703932" httpUri="https://zenodo.org/record/14703932/files/figure.png" pageId="14" pageNumber="69">Figures 30</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFE2FFF69E7DF95BFCE847CB" box="[859,891,1744,1770]" captionStart="FIGURES 3138" captionStartId="13.[152,270,1938,1962]" captionTargetBox="[184,1402,185,1909]" captionTargetId="figure-17@13.[181,1406,181,1915]" captionTargetPageId="13" captionText="FIGURES 3138. DIC micrographs of Cosmolaelaps hallasanensis sp. nov., adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale." figureDoi="http://doi.org/10.5281/zenodo.14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="14" pageNumber="69">37</figureCitation>
).
</paragraph>
<caption id="2AB96616FFE1FFF59DBEF819FC5046D2" ID-DOI="http://doi.org/10.5281/zenodo.14703934" ID-Zenodo-Dep="14703934" httpUri="https://zenodo.org/record/14703934/files/figure.png" pageId="13" pageNumber="68" startId="13.[152,270,1938,1962]" targetBox="[184,1402,185,1909]" targetPageId="13" targetType="figure">
<paragraph id="7E79369EFFE1FFF59DBEF819FC5046D2" blockId="13.[151,1437,1938,2035]" pageId="13" pageNumber="68">
<emphasis id="4CB2EA8CFFE1FFF59DBEF819FC73468A" bold="true" box="[152,992,1938,1963]" pageId="13" pageNumber="68">
FIGURES 3138. DIC micrographs of
<taxonomicName id="B9C64D1DFFE1FFF59F74F819FCE8468B" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[594,891,1938,1962]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="68" phylum="Arthropoda" rank="species" species="hallasanensis" status="sp. nov.">
<emphasis id="4CB2EA8CFFE1FFF59F74F819FCE8468B" bold="true" box="[594,891,1938,1962]" italics="true" pageId="13" pageNumber="68">Cosmolaelaps hallasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="578157F7FFE1FFF59EA3F818FC48468A" box="[901,987,1939,1963]" pageId="13" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
adult female: 31—Dorsal idiosoma; 32— Ventral idiosoma; 33—Sternal shield; 34—Genital shield; 35—Anal shield; 36—Ventral gnathosoma; 37—Chelicera; 38— Distal portion of palp, with focused on tarsal claw, enlarged section not to scale.
</paragraph>
</caption>
<caption id="2AB96616FFE2FFF69DBEFA5EFAE344CC" ID-DOI="http://doi.org/10.5281/zenodo.14703938" ID-Zenodo-Dep="14703938" box="[152,1392,1493,1517]" httpUri="https://zenodo.org/record/14703938/files/figure.png" pageId="14" pageNumber="69" startId="14.[152,270,1493,1517]" targetBox="[167,1420,197,1453]" targetPageId="14" targetType="figure">
<paragraph id="7E79369EFFE2FFF69DBEFA5EFAE344CC" blockId="14.[152,1392,1493,1517]" box="[152,1392,1493,1517]" pageId="14" pageNumber="69">
<emphasis id="4CB2EA8CFFE2FFF69DBEFA5EFD7044CC" bold="true" box="[152,739,1493,1517]" pageId="14" pageNumber="69">
FIGURES 3942.
<taxonomicName id="B9C64D1DFFE2FFF69C46FA5EFD1544CC" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[352,646,1493,1517]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="69" phylum="Arthropoda" rank="species" species="hallasanensis" status="sp. nov.">
<emphasis id="4CB2EA8CFFE2FFF69C46FA5EFD1544CC" bold="true" box="[352,646,1493,1517]" italics="true" pageId="14" pageNumber="69">Cosmolaelaps hallasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="578157F7FFE2FFF69FAAFA5EFD4C44CC" box="[652,735,1493,1517]" pageId="14" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
adult female: 39—Leg I; 40—Leg II; 41—Leg III; 42—Leg IV.
</paragraph>
</caption>
<paragraph id="7E79369EFFE2FFF69DE1F978FCEF462C" blockId="14.[151,1437,1599,2022]" box="[199,892,1779,1806]" pageId="14" pageNumber="69">
<emphasis id="4CB2EA8CFFE2FFF69DE1F978FE78462F" bold="true" box="[199,491,1779,1806]" pageId="14" pageNumber="69">Insemination structures.</emphasis>
Not seen, apparently unsclerotised.
</paragraph>
<paragraph id="7E79369EFFE2FFF79DE1F893FBE940A8" blockId="14.[151,1437,1599,2022]" lastBlockId="15.[151,1436,151,393]" lastPageId="15" lastPageNumber="70" pageId="14" pageNumber="69">
<emphasis id="4CB2EA8CFFE2FFF69DE1F893FF6C4613" bold="true" box="[199,255,1816,1842]" pageId="14" pageNumber="69">Legs</emphasis>
(
<figureCitation id="E6FD2A1BFFE2FFF69C32F89CFE524613" box="[276,449,1815,1842]" captionStart="FIGURES 3942" captionStartId="14.[152,270,1493,1517]" captionTargetBox="[167,1420,197,1453]" captionTargetId="figure-36@14.[151,1436,181,1469]" captionTargetPageId="14" captionText="FIGURES 3942. Cosmolaelaps hallasanensis sp. nov., adult female: 39—Leg I; 40—Leg II; 41—Leg III; 42—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703938" httpUri="https://zenodo.org/record/14703938/files/figure.png" pageId="14" pageNumber="69">Figures 3942</figureCitation>
). Legs II and III short (551619, 539584), I and IV longer (675787, 706828). Chaetotaxy normal for free-living
<taxonomicName id="B9C64D1DFFE2FFF69F0EF8B0FD344674" box="[552,679,1851,1877]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="69" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
: Leg I (
<figureCitation id="E6FD2A1BFFE2FFF69E21F8B0FCEE4677" box="[775,893,1851,1878]" captionStart="FIGURES 3942" captionStartId="14.[152,270,1493,1517]" captionTargetBox="[167,1420,197,1453]" captionTargetId="figure-36@14.[151,1436,181,1469]" captionTargetPageId="14" captionText="FIGURES 3942. Cosmolaelaps hallasanensis sp. nov., adult female: 39—Leg I; 40—Leg II; 41—Leg III; 42—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703938" httpUri="https://zenodo.org/record/14703938/files/figure.png" pageId="14" pageNumber="69">Figure 39</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/2, 0/1-1 (
<emphasis id="4CB2EA8CFFE2FFF698A0F8B0FA0F4674" box="[1414,1436,1851,1877]" italics="true" pageId="14" pageNumber="69">pl</emphasis>
spatulate), femur 2-2/1, 3/3-2 (posterior dorsal and ventral setae spatulate, except
<emphasis id="4CB2EA8CFFE2FFF69908F8EBFBC4465B" box="[1070,1111,1888,1914]" italics="true" pageId="14" pageNumber="69">pv3</emphasis>
), genu 2-3/2, 3/1-2 (all setae spatulate, except ventral and
<emphasis id="4CB2EA8CFFE2FFF69CC2F808FD9B46BC" box="[484,520,1923,1949]" italics="true" pageId="14" pageNumber="69">al2</emphasis>
), tibia 2-3/2, 3/1-2 (all setae spatulate, except ventral and anterior laterals). Leg II (
<figureCitation id="E6FD2A1BFFE2FFF69D9DF82CFEA346E3" box="[187,304,1959,1986]" captionStart="FIGURES 3942" captionStartId="14.[152,270,1493,1517]" captionTargetBox="[167,1420,197,1453]" captionTargetId="figure-36@14.[151,1436,181,1469]" captionTargetPageId="14" captionText="FIGURES 3942. Cosmolaelaps hallasanensis sp. nov., adult female: 39—Leg I; 40—Leg II; 41—Leg III; 42—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703938" httpUri="https://zenodo.org/record/14703938/files/figure.png" pageId="14" pageNumber="69">Figure 40</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/1-1, femur 2-3/1, 2/2-1 (
<emphasis id="4CB2EA8CFFE2FFF69932F82CFB4A46E0" box="[1044,1241,1959,1985]" italics="true" pageId="14" pageNumber="69">ad1, pd1, pd2, pl</emphasis>
spatulate and
<emphasis id="4CB2EA8CFFE2FFF698A4F823FA0F46E0" box="[1410,1436,1960,1985]" italics="true" pageId="14" pageNumber="69">av</emphasis>
spur-like), genu 2-3/1, 2/1-2 (all setae spatulate, except ventral and anterior laterals, ventral setae thickened), tibia 2-2/1, 2/1-2 (all setae spatulate, except ventral and anterior laterals, ventral setae thickened). Leg III (
<figureCitation id="E6FD2A1BFFE3FFF7983AFF1CFA1C4190" box="[1308,1423,151,177]" captionStart="FIGURES 3942" captionStartId="14.[152,270,1493,1517]" captionTargetBox="[167,1420,197,1453]" captionTargetId="figure-36@14.[151,1436,181,1469]" captionTargetPageId="14" captionText="FIGURES 3942. Cosmolaelaps hallasanensis sp. nov., adult female: 39—Leg I; 40—Leg II; 41—Leg III; 42—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703938" httpUri="https://zenodo.org/record/14703938/files/figure.png" pageId="15" pageNumber="70">Figure 41</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/1-1 (
<emphasis id="4CB2EA8CFFE3FFF79F5CFF30FD0541F4" box="[634,662,187,213]" italics="true" pageId="15" pageNumber="70">ad</emphasis>
spatulate), femur 1-2/1, 1/0-1 (
<emphasis id="4CB2EA8CFFE3FFF79EDAFF30FBB541F4" box="[1020,1062,187,213]" italics="true" pageId="15" pageNumber="70">ad1</emphasis>
spatulate), genu 2-2/1, 2/1-1 (all setae spatulate, except ventrals), tibia: 2-1/1, 2/1-1 (all setae spatulate, except ventrals). Leg IV (
<figureCitation id="E6FD2A1BFFE3FFF799F9FF54FAC041D8" box="[1247,1363,223,249]" captionStart="FIGURES 3942" captionStartId="14.[152,270,1493,1517]" captionTargetBox="[167,1420,197,1453]" captionTargetId="figure-36@14.[151,1436,181,1469]" captionTargetPageId="14" captionText="FIGURES 3942. Cosmolaelaps hallasanensis sp. nov., adult female: 39—Leg I; 40—Leg II; 41—Leg III; 42—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703938" httpUri="https://zenodo.org/record/14703938/files/figure.png" pageId="15" pageNumber="70">Figure 42</figureCitation>
): coxa 0-0/1, 0/0-0, trochanter 1-1/1, 0/1-1 (
<emphasis id="4CB2EA8CFFE3FFF79F1CFE88FDC5403C" box="[570,598,259,285]" italics="true" pageId="15" pageNumber="70">ad</emphasis>
spatulate), femur 1-2/1, 1/0-1 (
<emphasis id="4CB2EA8CFFE3FFF79E9FFE88FBD3403C" box="[953,1088,259,285]" italics="true" pageId="15" pageNumber="70">ad1, ad2, al</emphasis>
, and
<emphasis id="4CB2EA8CFFE3FFF79959FE88FB0A403D" box="[1151,1177,259,284]" italics="true" pageId="15" pageNumber="70">av</emphasis>
spatulate), genu 2-2/1, 3/0-1 (all setae spatulate, except av and pl), tibia 2-1/1, 3/1-2 (all setae spatulate, except ventrals). Tarsi IIIV with 18 setae (3-3/2, 3/2-3 +
<emphasis id="4CB2EA8CFFE3FFF79C82FEC0FE604044" box="[420,499,331,357]" italics="true" pageId="15" pageNumber="70">mv, md</emphasis>
); with some ventral and lateral setae thickened and spatulate, see
<figureCitation id="E6FD2A1BFFE3FFF799F0FEC0FAFC4044" box="[1238,1391,331,357]" captionStart="FIGURES 3942" captionStartId="14.[152,270,1493,1517]" captionTargetBox="[167,1420,197,1453]" captionTargetId="figure-36@14.[151,1436,181,1469]" captionTargetPageId="14" captionText="FIGURES 3942. Cosmolaelaps hallasanensis sp. nov., adult female: 39—Leg I; 40—Leg II; 41—Leg III; 42—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703938" httpUri="https://zenodo.org/record/14703938/files/figure.png" pageId="15" pageNumber="70">Figure 4042</figureCitation>
. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="7E79369EFFE3FFF79DB1FE51FE5B40D5" blockId="15.[151,456,474,537]" box="[151,456,474,500]" pageId="15" pageNumber="70">
<emphasis id="4CB2EA8CFFE3FFF79DB1FE51FE5B40D5" bold="true" box="[151,456,474,500]" pageId="15" pageNumber="70">Description (adult male)</emphasis>
</paragraph>
<paragraph id="7E79369EFFE3FFF79DB1FE74FED84338" blockId="15.[151,456,474,537]" box="[151,331,511,537]" pageId="15" pageNumber="70">
(
<figureCitation id="E6FD2A1BFFE3FFF79DB9FE74FED04338" box="[159,323,511,537]" captionStart="FIGURES 4346" captionStartId="16.[152,270,1943,1967]" captionTargetBox="[183,1408,188,1913]" captionTargetId="figure-26@16.[177,1411,181,1920]" captionTargetPageId="16" captionText="FIGURES 4346. Cosmolaelaps hallasanensis sp. nov., adult male: 43—Line drawing plate of ventral idiosoma; 44—Line drawing plate of chelicera; 45—DIC micrographs of ventral idiosoma; 46—DIC micrographs of femur II." figureDoi="http://doi.org/10.5281/zenodo.14703941" httpUri="https://zenodo.org/record/14703941/files/figure.png" pageId="15" pageNumber="70">Figures 4346</figureCitation>
)
</paragraph>
<paragraph id="7E79369EFFE3FFF79DB1FDC8FE58437A" blockId="15.[151,459,579,603]" box="[151,459,579,603]" pageId="15" pageNumber="70">
<specimenCount id="68C0FD17FFE3FFF79DB1FDC8FEB6437A" box="[151,293,579,603]" count="6" pageId="15" pageNumber="70" type="generic">six specimens</specimenCount>
were measured.
</paragraph>
<paragraph id="7E79369EFFE3FFF79DB1FD02FAD24382" blockId="15.[151,1437,649,1180]" box="[151,1345,649,676]" pageId="15" pageNumber="70">
<emphasis id="4CB2EA8CFFE3FFF79DB1FD02FEF34382" bold="true" box="[151,352,649,675]" pageId="15" pageNumber="70">Dorsal idiosoma.</emphasis>
Dorsal shield 567622 long, 420478 wide; ornamentation and chaetotaxy as in female.
</paragraph>
<paragraph id="7E79369EFFE3FFF79DE1FD26FD5642C6" blockId="15.[151,1437,649,1180]" pageId="15" pageNumber="70">
<emphasis id="4CB2EA8CFFE3FFF79DE1FD26FE0243E6" bold="true" box="[199,401,685,711]" pageId="15" pageNumber="70">Ventral idiosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFE3FFF79CB9FD26FDC443E9" box="[415,599,685,712]" captionStart="FIGURES 4346" captionStartId="16.[152,270,1943,1967]" captionTargetBox="[183,1408,188,1913]" captionTargetId="figure-26@16.[177,1411,181,1920]" captionTargetPageId="16" captionText="FIGURES 4346. Cosmolaelaps hallasanensis sp. nov., adult male: 43—Line drawing plate of ventral idiosoma; 44—Line drawing plate of chelicera; 45—DIC micrographs of ventral idiosoma; 46—DIC micrographs of femur II." figureDoi="http://doi.org/10.5281/zenodo.14703941" httpUri="https://zenodo.org/record/14703941/files/figure.png" pageId="15" pageNumber="70">Figures 43 &amp; 45</figureCitation>
). Sternal, genital, endopodal, ventral and anal shields fused into holoventral shield, 475491 long from anterior to posterior margins of shield, 190195 wide at level of
<emphasis id="4CB2EA8CFFE3FFF799A8FD5AFB3C43CA" box="[1166,1199,721,747]" italics="true" pageId="15" pageNumber="70">st2</emphasis>
, 173179 at
<emphasis id="4CB2EA8CFFE3FFF79819FD5AFAF343CA" box="[1343,1376,721,747]" italics="true" pageId="15" pageNumber="70">st3</emphasis>
level and 261273 at broadest point, posteriorly to coxae IV; shield reticulate throughout (ornamentation only partly drawn on
<figureCitation id="E6FD2A1BFFE3FFF79C2EFC92FE504215" box="[264,451,793,820]" captionStart="FIGURES 4346" captionStartId="16.[152,270,1943,1967]" captionTargetBox="[183,1408,188,1913]" captionTargetId="figure-26@16.[177,1411,181,1920]" captionTargetPageId="16" captionText="FIGURES 4346. Cosmolaelaps hallasanensis sp. nov., adult male: 43—Line drawing plate of ventral idiosoma; 44—Line drawing plate of chelicera; 45—DIC micrographs of ventral idiosoma; 46—DIC micrographs of femur II." figureDoi="http://doi.org/10.5281/zenodo.14703941" httpUri="https://zenodo.org/record/14703941/files/figure.png" pageId="15" pageNumber="70">Figures 43 &amp; 45</figureCitation>
), with five pairs of simple sternal setae (
<emphasis id="4CB2EA8CFFE3FFF79EA8FC92FC4E4212" box="[910,989,793,819]" italics="true" pageId="15" pageNumber="70">st1st5</emphasis>
), and five pairs of smooth ventral setae (
<emphasis id="4CB2EA8CFFE3FFF79D87FCB6FF6F4276" box="[161,252,829,856]" italics="true" pageId="15" pageNumber="70">Jv1Jv3</emphasis>
,
<emphasis id="4CB2EA8CFFE3FFF79C2CFCB6FEA74276" box="[266,308,829,855]" italics="true" pageId="15" pageNumber="70">Zv1</emphasis>
, and
<emphasis id="4CB2EA8CFFE3FFF79C56FCB6FE094276" box="[368,410,829,855]" italics="true" pageId="15" pageNumber="70">Zv2</emphasis>
), plus three smooth circum-anal setae, para-anal setae and post-anal seta subequal in length, post-anal seta thickened, with five pairs of poroids; gland pore
<emphasis id="4CB2EA8CFFE3FFF79E70FCEAFCED425A" box="[854,894,865,891]" italics="true" pageId="15" pageNumber="70">gv2</emphasis>
behind coxa IV not discerned; cribrum with 34 irregular rows of spicules; metapodal platelets narrow and partly fused to holoventral shield (
<figureCitation id="E6FD2A1BFFE3FFF79986FC0EFAC84281" box="[1184,1371,901,928]" captionStart="FIGURES 4346" captionStartId="16.[152,270,1943,1967]" captionTargetBox="[183,1408,188,1913]" captionTargetId="figure-26@16.[177,1411,181,1920]" captionTargetPageId="16" captionText="FIGURES 4346. Cosmolaelaps hallasanensis sp. nov., adult male: 43—Line drawing plate of ventral idiosoma; 44—Line drawing plate of chelicera; 45—DIC micrographs of ventral idiosoma; 46—DIC micrographs of femur II." figureDoi="http://doi.org/10.5281/zenodo.14703941" httpUri="https://zenodo.org/record/14703941/files/figure.png" pageId="15" pageNumber="70">Figures 43 &amp; 45</figureCitation>
). Soft opisthogastric and lateral cuticle with 13 pairs of spatulate setae, except
<emphasis id="4CB2EA8CFFE3FFF79EF7FC22FC6B42E2" box="[977,1016,937,963]" italics="true" pageId="15" pageNumber="70">Jv4</emphasis>
. Peritremes, peritrematal shields and other ventral structures similar to those in female.
</paragraph>
<paragraph id="7E79369EFFE3FFF79DE1FC7AFC3E4556" blockId="15.[151,1437,649,1180]" pageId="15" pageNumber="70">
<emphasis id="4CB2EA8CFFE3FFF79DE1FC7AFECC452A" bold="true" box="[199,351,1009,1035]" pageId="15" pageNumber="70">Gnathosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFE3FFF79C48FC7AFE73452A" box="[366,480,1009,1035]" captionStart="FIGURES 4346" captionStartId="16.[152,270,1943,1967]" captionTargetBox="[183,1408,188,1913]" captionTargetId="figure-26@16.[177,1411,181,1920]" captionTargetPageId="16" captionText="FIGURES 4346. Cosmolaelaps hallasanensis sp. nov., adult male: 43—Line drawing plate of ventral idiosoma; 44—Line drawing plate of chelicera; 45—DIC micrographs of ventral idiosoma; 46—DIC micrographs of femur II." figureDoi="http://doi.org/10.5281/zenodo.14703941" httpUri="https://zenodo.org/record/14703941/files/figure.png" pageId="15" pageNumber="70">Figure 44</figureCitation>
). Fixed digit of chelicera with 1113 teeth of various sizes in addition to apical hook, pilus dentilis setaceous, slightly thicker than in female. Movable digit of chelicera with a relatively large tooth, spermatodactyl digitiform, gently curved, with rounded tip, free portion of spermatodactyl shorter than movable digit (5360). Other gnathosomal structures similar to those in female.
</paragraph>
<paragraph id="7E79369EFFE3FFF79DE1FB0AFA0745BD" blockId="15.[151,1437,649,1180]" box="[199,1428,1153,1180]" pageId="15" pageNumber="70">
<emphasis id="4CB2EA8CFFE3FFF79DE1FB0AFF6C45BA" bold="true" box="[199,255,1153,1179]" pageId="15" pageNumber="70">Legs</emphasis>
(
<figureCitation id="E6FD2A1BFFE3FFF79C28FB0AFE1345BA" box="[270,384,1153,1179]" captionStart="FIGURES 4346" captionStartId="16.[152,270,1943,1967]" captionTargetBox="[183,1408,188,1913]" captionTargetId="figure-26@16.[177,1411,181,1920]" captionTargetPageId="16" captionText="FIGURES 4346. Cosmolaelaps hallasanensis sp. nov., adult male: 43—Line drawing plate of ventral idiosoma; 44—Line drawing plate of chelicera; 45—DIC micrographs of ventral idiosoma; 46—DIC micrographs of femur II." figureDoi="http://doi.org/10.5281/zenodo.14703941" httpUri="https://zenodo.org/record/14703941/files/figure.png" pageId="15" pageNumber="70">Figure 46</figureCitation>
). Chaetotaxy as in female, including strong spur-like seta (
<emphasis id="4CB2EA8CFFE3FFF79934FB09FBBF45BA" box="[1042,1068,1154,1179]" italics="true" pageId="15" pageNumber="70">av</emphasis>
) on the ventral side of femur II.
</paragraph>
<paragraph id="7E79369EFFE3FFF79DB1FB66FE724426" blockId="15.[151,481,1261,1324]" box="[151,481,1261,1287]" pageId="15" pageNumber="70">
<emphasis id="4CB2EA8CFFE3FFF79DB1FB66FE724426" bold="true" box="[151,481,1261,1287]" pageId="15" pageNumber="70">Description (deutonymph)</emphasis>
</paragraph>
<paragraph id="7E79369EFFE3FFF79DB1FA9AFED8440D" blockId="15.[151,481,1261,1324]" box="[151,331,1297,1324]" pageId="15" pageNumber="70">
(
<figureCitation id="E6FD2A1BFFE3FFF79DB9FA9AFED0440D" box="[159,323,1297,1324]" captionStart="FIGURES 4752" captionStartId="17.[152,270,1943,1967]" captionTargetBox="[197,1396,197,1903]" captionTargetId="figure-17@17.[182,1405,181,1919]" captionTargetPageId="17" captionText="FIGURES 4752. Cosmolaelaps hallasanensis sp. nov., deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703943" httpUri="https://zenodo.org/record/14703943/files/figure.png" pageId="15" pageNumber="70">Figures 4752</figureCitation>
)
</paragraph>
<paragraph id="7E79369EFFE3FFF79DB1FADDFE4E444F" blockId="15.[151,477,1366,1390]" box="[151,477,1366,1390]" pageId="15" pageNumber="70">
<specimenCount id="68C0FD17FFE3FFF79DB1FADDFEA4444F" box="[151,311,1366,1390]" count="4" pageId="15" pageNumber="70" type="generic">Four specimens</specimenCount>
were measured.
</paragraph>
<paragraph id="7E79369EFFE3FFF79DB1FA17FAE1474B" blockId="15.[151,1437,1436,2003]" pageId="15" pageNumber="70">
<emphasis id="4CB2EA8CFFE3FFF79DB1FA17FEC44497" bold="true" box="[151,343,1436,1462]" pageId="15" pageNumber="70">Dorsal idiosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFE3FFF79C42FA17FE474497" box="[356,468,1436,1462]" captionStart="FIGURES 4752" captionStartId="17.[152,270,1943,1967]" captionTargetBox="[197,1396,197,1903]" captionTargetId="figure-17@17.[182,1405,181,1919]" captionTargetPageId="17" captionText="FIGURES 4752. Cosmolaelaps hallasanensis sp. nov., deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703943" httpUri="https://zenodo.org/record/14703943/files/figure.png" pageId="15" pageNumber="70">Figure 47</figureCitation>
). Dorsal shield entire 737790 long, 523538 wide, not covering entire dorsal idiosoma, with a narrow incision in each lateral margin, just anteriorly to
<emphasis id="4CB2EA8CFFE3FFF79E5CFA4BFC0544FB" box="[890,918,1472,1498]" italics="true" pageId="15" pageNumber="70">S1</emphasis>
and extending medially about up to level of
<emphasis id="4CB2EA8CFFE3FFF79DB1FA6FFF2344DF" box="[151,176,1508,1534]" italics="true" pageId="15" pageNumber="70">z6</emphasis>
, bearing 40 pairs of almost uniform spatulate-mucronate setae: 22 pairs of podonotal setae and 18 pairs of opisthonotal setae, including three pairs of
<emphasis id="4CB2EA8CFFE3FFF79F5BF982FD0A4703" box="[637,665,1545,1570]" italics="true" pageId="15" pageNumber="70">Zx</emphasis>
setae between
<emphasis id="4CB2EA8CFFE3FFF79E63F982FCC24703" box="[837,849,1545,1570]" italics="true" pageId="15" pageNumber="70">J</emphasis>
and
<emphasis id="4CB2EA8CFFE3FFF79EAFF982FC0A4703" box="[905,921,1545,1570]" italics="true" pageId="15" pageNumber="70">Z</emphasis>
setae and six unpaired supernumerary seta
<emphasis id="4CB2EA8CFFE3FFF798A5F982FA0F4703" box="[1411,1436,1545,1570]" italics="true" pageId="15" pageNumber="70">Jx</emphasis>
between
<emphasis id="4CB2EA8CFFE3FFF79DDDF9A7FE864767" box="[251,277,1580,1606]" italics="true" pageId="15" pageNumber="70">J1</emphasis>
and
<emphasis id="4CB2EA8CFFE3FFF79C6CF9A7FEF74767" box="[330,356,1580,1606]" italics="true" pageId="15" pageNumber="70">J5</emphasis>
, dorsal setae mostly homogeneous in length and thickness, barely reaching base of next posterior setae (3540), except
<emphasis id="4CB2EA8CFFE3FFF79CABF9DBFE31474B" box="[397,418,1616,1642]" italics="true" pageId="15" pageNumber="70">j1</emphasis>
cunate-tricarinate and
<emphasis id="4CB2EA8CFFE3FFF79F82F9DBFD2E474B" box="[676,701,1616,1642]" italics="true" pageId="15" pageNumber="70">z1</emphasis>
acuminate. Pore-like structures essentially as in adult female.
</paragraph>
<paragraph id="7E79369EFFE3FFF79DE1F9FFFE2D468F" blockId="15.[151,1437,1436,2003]" pageId="15" pageNumber="70">
<emphasis id="4CB2EA8CFFE3FFF79DE1F9FFFE0647AF" bold="true" box="[199,405,1652,1678]" pageId="15" pageNumber="70">Ventral idiosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFE3FFF79C81F9FFFD8847AF" box="[423,539,1652,1678]" captionStart="FIGURES 4752" captionStartId="17.[152,270,1943,1967]" captionTargetBox="[197,1396,197,1903]" captionTargetId="figure-17@17.[182,1405,181,1919]" captionTargetPageId="17" captionText="FIGURES 4752. Cosmolaelaps hallasanensis sp. nov., deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703943" httpUri="https://zenodo.org/record/14703943/files/figure.png" pageId="15" pageNumber="70">Figure 48</figureCitation>
). Tritosternum with paired pilose laciniae (84105), trapezoidal base 4349 × 1115 wide. Sternal shield length 255265 and widest 111120, finely reticulate throughout, with four pairs of relatively long setae (
<emphasis id="4CB2EA8CFFE3FFF79CAFF937FE4B47F7" box="[393,472,1724,1751]" italics="true" pageId="15" pageNumber="70">st1st4</emphasis>
) (2765) and three pairs of poroids. Genital setae
<emphasis id="4CB2EA8CFFE3FFF79928F937FBBD47F7" box="[1038,1070,1724,1750]" italics="true" pageId="15" pageNumber="70">st5</emphasis>
(3036) and paragenital poroids on soft cuticle at level of posterior margin of shield; endopodal shields between coxae IIIII and coxae IIIIV absent. Anal shield subtriangular (103110 long, 8494 wide), anterior half lineate-reticulate, para-anal setae and post-anal seta subequal in length (2129), cribrum small, gland pores
<emphasis id="4CB2EA8CFFE3FFF79E6DF8A3FCE04663" box="[843,883,1832,1858]" italics="true" pageId="15" pageNumber="70">gv3</emphasis>
on lateral margins of shield. Soft opisthogastric cuticle as in adult female. Peritrematal shields anteriorly free, not fused with dorsal shield, bearing one gland pore and one poroid at level near coxae IIIII; post-stigmatic extension of shield not developed, two pairs of related poroids on small platelets.
</paragraph>
<paragraph id="7E79369EFFE3FFF79DE1F833FDB946F3" blockId="15.[151,1437,1436,2003]" box="[199,554,1976,2003]" pageId="15" pageNumber="70">
<emphasis id="4CB2EA8CFFE3FFF79DE1F833FEF446F3" bold="true" box="[199,359,1976,2002]" pageId="15" pageNumber="70">Gnathosoma.</emphasis>
similar to adults.
</paragraph>
<caption id="2AB96616FFFCFFE89DBEF81CFB5B46F2" ID-DOI="http://doi.org/10.5281/zenodo.14703941" ID-Zenodo-Dep="14703941" httpUri="https://zenodo.org/record/14703941/files/figure.png" pageId="16" pageNumber="71" startId="16.[152,270,1943,1967]" targetBox="[183,1408,188,1913]" targetPageId="16" targetType="figure">
<paragraph id="7E79369EFFFCFFE89DBEF81CFB5B46F2" blockId="16.[151,1436,1943,2003]" pageId="16" pageNumber="71">
<emphasis id="4CB2EA8CFFFCFFE89DBEF81CFD7A4691" bold="true" box="[152,745,1943,1968]" pageId="16" pageNumber="71">
FIGURES 4346.
<taxonomicName id="B9C64D1DFFFCFFE89C45F81CFD1A468E" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[355,649,1943,1967]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="hallasanensis" status="sp. nov.">
<emphasis id="4CB2EA8CFFFCFFE89C45F81CFD1A468E" bold="true" box="[355,649,1943,1967]" italics="true" pageId="16" pageNumber="71">Cosmolaelaps hallasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="578157F7FFFCFFE89FB6F813FD774691" box="[656,740,1944,1968]" pageId="16" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
adult male: 43—Line drawing plate of ventral idiosoma; 44—Line drawing plate of chelicera; 45—DIC micrographs of ventral idiosoma; 46—DIC micrographs of femur II.
</paragraph>
</caption>
<caption id="2AB96616FFFDFFE99DBEF81CFDA146F2" ID-DOI="http://doi.org/10.5281/zenodo.14703943" ID-Zenodo-Dep="14703943" httpUri="https://zenodo.org/record/14703943/files/figure.png" pageId="17" pageNumber="72" startId="17.[152,270,1943,1967]" targetBox="[197,1396,197,1903]" targetPageId="17" targetType="figure">
<paragraph id="7E79369EFFFDFFE99DBEF81CFDA146F2" blockId="17.[151,1436,1943,2003]" pageId="17" pageNumber="72">
<emphasis id="4CB2EA8CFFFDFFE99DBEF81CFD734691" bold="true" box="[152,736,1943,1968]" pageId="17" pageNumber="72">
FIGURES 4752.
<taxonomicName id="B9C64D1DFFFDFFE99C79F81CFD17468E" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[351,644,1943,1967]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="72" phylum="Arthropoda" rank="species" species="hallasanensis" status="sp. nov.">
<emphasis id="4CB2EA8CFFFDFFE99C79F81CFD17468E" bold="true" box="[351,644,1943,1967]" italics="true" pageId="17" pageNumber="72">Cosmolaelaps hallasanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="578157F7FFFDFFE99FACF813FD4F4691" box="[650,732,1944,1968]" pageId="17" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV.
</paragraph>
</caption>
<paragraph id="7E79369EFFFEFFEA9DE1FF1CFAA441F4" blockId="18.[151,1437,150,969]" pageId="18" pageNumber="73">
<emphasis id="4CB2EA8CFFFEFFEA9DE1FF1CFF6C4190" bold="true" box="[199,255,151,177]" pageId="18" pageNumber="73">Legs</emphasis>
(
<figureCitation id="E6FD2A1BFFFEFFEA9C28FF1CFE254190" box="[270,438,150,177]" captionStart="FIGURES 4752" captionStartId="17.[152,270,1943,1967]" captionTargetBox="[197,1396,197,1903]" captionTargetId="figure-17@17.[182,1405,181,1919]" captionTargetPageId="17" captionText="FIGURES 4752. Cosmolaelaps hallasanensis sp. nov., deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703943" httpUri="https://zenodo.org/record/14703943/files/figure.png" pageId="18" pageNumber="73">Figures 4952</figureCitation>
). Legs II (
<figureCitation id="E6FD2A1BFFFEFFEA9F0EFF1CFD044190" box="[552,663,150,177]" captionStart="FIGURES 4752" captionStartId="17.[152,270,1943,1967]" captionTargetBox="[197,1396,197,1903]" captionTargetId="figure-17@17.[182,1405,181,1919]" captionTargetPageId="17" captionText="FIGURES 4752. Cosmolaelaps hallasanensis sp. nov., deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703943" httpUri="https://zenodo.org/record/14703943/files/figure.png" pageId="18" pageNumber="73">Figure 50</figureCitation>
) (443500) and III (
<figureCitation id="E6FD2A1BFFFEFFEA9E5AFF1CFC7E4190" box="[892,1005,150,177]" captionStart="FIGURES 4752" captionStartId="17.[152,270,1943,1967]" captionTargetBox="[197,1396,197,1903]" captionTargetId="figure-17@17.[182,1405,181,1919]" captionTargetPageId="17" captionText="FIGURES 4752. Cosmolaelaps hallasanensis sp. nov., deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703943" httpUri="https://zenodo.org/record/14703943/files/figure.png" pageId="18" pageNumber="73">Figure 51</figureCitation>
), (433461) short, I (
<figureCitation id="E6FD2A1BFFFEFFEA99FAFF1CFAD84190" box="[1244,1355,151,177]" captionStart="FIGURES 4752" captionStartId="17.[152,270,1943,1967]" captionTargetBox="[197,1396,197,1903]" captionTargetId="figure-17@17.[182,1405,181,1919]" captionTargetPageId="17" captionText="FIGURES 4752. Cosmolaelaps hallasanensis sp. nov., deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703943" httpUri="https://zenodo.org/record/14703943/files/figure.png" pageId="18" pageNumber="73">Figure 49</figureCitation>
) (591 650) and IV (
<figureCitation id="E6FD2A1BFFFEFFEA9C0AFF30FE0841F4" box="[300,411,186,213]" captionStart="FIGURES 4752" captionStartId="17.[152,270,1943,1967]" captionTargetBox="[197,1396,197,1903]" captionTargetId="figure-17@17.[182,1405,181,1919]" captionTargetPageId="17" captionText="FIGURES 4752. Cosmolaelaps hallasanensis sp. nov., deutonymph: 47—Dorsal idiosoma; 48—Ventral idiosoma; 49—Leg I; 50—Leg II; 51—Leg III; 52—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703943" httpUri="https://zenodo.org/record/14703943/files/figure.png" pageId="18" pageNumber="73">Figure 52</figureCitation>
) (590620) longer. Pretarsi, claws, and chaetotaxy of legs IIV as in adult female.
</paragraph>
</subSubSection>
<subSubSection id="36DC6515FFFEFFEA9DE1FF54FE7D403D" pageId="18" pageNumber="73" type="etymology">
<paragraph id="7E79369EFFFEFFEA9DE1FF54FE7D403D" blockId="18.[151,1437,150,969]" pageId="18" pageNumber="73">
<emphasis id="4CB2EA8CFFFEFFEA9DE1FF54FEDD41D8" bold="true" box="[199,334,223,249]" pageId="18" pageNumber="73">Etymology.</emphasis>
The specific name is derived from the name of the region (Hallasan Mountain,
<collectingRegion id="BC02F87CFFFEFFEA99D7FF54FAB241D8" box="[1265,1313,223,249]" country="South Korea" name="Jeju" pageId="18" pageNumber="73">Jeju</collectingRegion>
,
<collectingCountry id="06D1760EFFFEFFEA9809FF54FAE641D8" box="[1327,1397,223,249]" name="South Korea" pageId="18" pageNumber="73">Korea</collectingCountry>
) in which the
<typeStatus id="A17D883CFFFEFFEA9C2AFE88FEAF403C" box="[268,316,259,285]" pageId="18" pageNumber="73">type</typeStatus>
locality occurs.
</paragraph>
</subSubSection>
<subSubSection id="36DC6515FFFEFFEA9DE1FEADFC2242E8" pageId="18" pageNumber="73" type="diagnosis">
<paragraph id="7E79369EFFFEFFEA9DE1FEADFC2242E8" blockId="18.[151,1437,150,969]" pageId="18" pageNumber="73">
<emphasis id="4CB2EA8CFFFEFFEA9DE1FEADFE5D4061" bold="true" box="[199,462,294,320]" pageId="18" pageNumber="73">Differential diagnosis.</emphasis>
Dorsal shield usually bearing 39 pairs of setae (22 pairs on podonotal and 17 pairs on opisthonotal regions including two pairs of
<emphasis id="4CB2EA8CFFFEFFEA9FAEFEC0FD374045" box="[648,676,331,356]" italics="true" pageId="18" pageNumber="73">Zx</emphasis>
setae between
<emphasis id="4CB2EA8CFFFEFFEA9E74FEC0FCCD4045" box="[850,862,331,356]" italics="true" pageId="18" pageNumber="73">J</emphasis>
and
<emphasis id="4CB2EA8CFFFEFFEA9EBEFEC0FC3B4045" box="[920,936,331,356]" italics="true" pageId="18" pageNumber="73">Z</emphasis>
series of each side, as well as 15 unpaired setae (
<emphasis id="4CB2EA8CFFFEFFEA9DC5FEE4FF6F40A9" box="[227,252,367,392]" italics="true" pageId="18" pageNumber="73">Jx</emphasis>
) between
<emphasis id="4CB2EA8CFFFEFFEA9C5FFEE4FE1640A9" box="[377,389,367,392]" italics="true" pageId="18" pageNumber="73">J</emphasis>
series as in most
<taxonomicName id="B9C64D1DFFFEFFEA9F46FEE4FCCE40A8" box="[608,861,367,393]" pageId="18" pageNumber="73">
<emphasis id="4CB2EA8CFFFEFFEA9F46FEE4FD6C40A8" box="[608,767,367,393]" italics="true" pageId="18" pageNumber="73">Cosmolaelaps</emphasis>
species
</taxonomicName>
; but opisthonotal region of dorsal shield with 1 3 extra pairs of setae in some species:
<taxonomicName id="B9C64D1DFFFEFFEA9F42FE18FC1D408C" authority="(Karg, 1981)" baseAuthorityName="Karg" baseAuthorityYear="1981" box="[612,910,403,429]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="anserina">
<emphasis id="4CB2EA8CFFFEFFEA9F42FE18FD78408D" box="[612,747,403,429]" italics="true" pageId="18" pageNumber="73">C. anserina</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9FD8FE18FC16408C" author="Karg, W." box="[766,901,403,429]" pageId="18" pageNumber="73" pagination="213 - 232" refId="ref18894" refString="Karg, W. (1981) Die Untergattung Cosmolaelaps Berlese, 1903 der Raubmilbengattung Hypoaspis Canestrini, 1884 (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 57, 213 - 232. https: // doi. org / 10.1002 / mmnz. 19810570203" type="journal article" year="1981">Karg, 1981</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9EBBFE18FB6F408C" authority="(Karg, 1985)" authorityName="Karg" authorityYear="1985" baseAuthorityName="Karg" baseAuthorityYear="1985" box="[925,1276,402,429]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="brevipedestra">
<emphasis id="4CB2EA8CFFFEFFEA9EBBFE18FBC9408C" box="[925,1114,403,429]" italics="true" pageId="18" pageNumber="73">C. brevipedestra</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA994BFE18FB67408C" author="Karg, W." box="[1133,1268,402,429]" pageId="18" pageNumber="73" pagination="233 - 237" refId="ref18945" refString="Karg, W. (1985) Zwei neue Raubmilbenarten der Untergattung Cosmolaelaps Berlese, 1903 (Gattung Hypoaspis Can, 1884) aus Termitennestern (Acarina, Parasitiformes). Deutsche Entomologische Zeitschrift, 32, 233 - 237. https: // doi. org / 10.1002 / mmnd. 19850320402" type="journal article" year="1985">Karg, 1985</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA982AFE18FE4340F0" authority="(Aswegen and Loots, 1970)" baseAuthorityName="Aswegen and Loots" baseAuthorityYear="1970" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="carvalhoi">
<emphasis id="4CB2EA8CFFFEFFEA982AFE18FA0F408C" box="[1292,1436,403,429]" italics="true" pageId="18" pageNumber="73">C. carvalhoi</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9DB9FE3CFE5B40F0" author="Aswegen, P. I. M. &amp; Loots, G. C." box="[159,456,439,465]" pageId="18" pageNumber="73" pagination="169 - 213" refId="ref17198" refString="Aswegen, P. I. M. &amp; Loots, G. C. (1970) A taxonomic study of the genus Hypoaspis Canestrini sens. lat. (Acari: Laelapidae) in the Ethiopian region. Publicacoes Culturais da Companhia de Diamantes de Angola, 82, 169 - 213." type="journal article" year="1970">Aswegen and Loots, 1970</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9CFAFE3CFCB140F0" authority="Tragardh, 1931" authorityName="Tragardh" authorityYear="1931" box="[476,802,438,465]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="communis">
<emphasis id="4CB2EA8CFFFEFFEA9CFAFE3CFDFD40F1" box="[476,622,439,465]" italics="true" pageId="18" pageNumber="73">C. communis</emphasis>
<bibRefCitation id="1A574B6FFFFEFFEA9F52FE3DFCB140F0" author="Tragardh, I." box="[628,802,438,465]" pageId="18" pageNumber="73" pagination="553 - 628" refId="ref20474" refString="Tragardh, I. (1931) Pacific mites: Acarina from the Juan Fernandez Islands. Natural History of Juan Fernandez and Eastern Islands, Uppsala, 3, 553 - 628." type="journal article" year="1931">Trägårdh, 1931</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9E0BFE3CFBE740F0" authority="(Karg, 2006)" baseAuthorityName="Karg" baseAuthorityYear="2006" box="[813,1140,439,465]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="crossocauda">
<emphasis id="4CB2EA8CFFFEFFEA9E0BFE3CFC4840F0" box="[813,987,439,465]" italics="true" pageId="18" pageNumber="73">C. crossocauda</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9ECFFE3CFBFF40F0" author="Karg, W." box="[1001,1132,439,465]" pageId="18" pageNumber="73" pagination="140 - 169" refId="ref19212" refString="Karg, W. (2006) The systematics of Parasitiformes, especially of Gamasina Leach (Acarina), with new species from Ecuador. Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe, 82, 140 - 169. https: // doi. org / 10.1002 / mmnz. 200600002" type="journal article" year="2006">Karg, 2006</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9959FE3CFA0440F0" authority="(Karg, 1988)" baseAuthorityName="Karg" baseAuthorityYear="1988" box="[1151,1431,439,465]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="cursoria">
<emphasis id="4CB2EA8CFFFEFFEA9959FE3CFB6D40F1" box="[1151,1278,439,465]" italics="true" pageId="18" pageNumber="73">C. cursoria</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA982BFE3CFA1C40F0" author="Karg, W." box="[1293,1423,439,465]" pageId="18" pageNumber="73" pagination="509 - 526" refId="ref19039" refString="Karg, W. (1988) Die Arten der Raubmilbenuntergattung Cosmolaelaps Berlese, 1903 (Gattung Hypoaspis Canestrini, 1884, Acarina, Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik, Geographie und Biologie der Tiere, 115, 509 - 526." type="journal article" year="1988">Karg, 1988</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9DB1FE50FE2E40D4" authority="(Karg, 1996)" baseAuthorityName="Karg" baseAuthorityYear="1996" box="[151,445,475,501]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="euarmata">
<emphasis id="4CB2EA8CFFFEFFEA9DB1FE50FEB040D5" box="[151,291,475,501]" italics="true" pageId="18" pageNumber="73">C. euarmata</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9C14FE50FE2640D4" author="Karg, W." box="[306,437,475,501]" pageId="18" pageNumber="73" pagination="149 - 195" refId="ref19162" refString="Karg, W. (1996) Neue Arten aus Raubmilbengattungen der Gamasina Leach (Acarina, Prasitiformes) mit Indikationen zum Entwicklungsalter. Mitteilungen aus dem Zoologischen Museum in Berlin, 72, 149 - 195. https: // doi. org / 10.1002 / mmnz. 19960720111" type="journal article" year="1996">Karg, 1996</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9CEFFE50FC8140D4" authority="(Karg, 1995)" baseAuthorityName="Karg" baseAuthorityYear="1995" box="[457,786,474,501]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="euventricosa">
<emphasis id="4CB2EA8CFFFEFFEA9CEFFE50FDEB40D5" box="[457,632,475,501]" italics="true" pageId="18" pageNumber="73">C. euventricosa</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9FA1FE50FC9940D4" author="Karg, W." box="[647,778,474,501]" pageId="18" pageNumber="73" pagination="21 - 25" refId="ref19130" refString="Karg, W. (1995) Zur Kenntnis der Raubmilbengattung Cosmolaelaps Berlese, 1903. Abhandlungen und Berichte des Naturkundemuseums Gorlitz, 68 (8), 21 - 25." type="journal article" year="1995">Karg, 1995</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9E38FE50FBC140D4" authority="(Hafez et al., 1982)" baseAuthorityName="Hafez" baseAuthorityYear="1982" box="[798,1106,475,501]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="keni">
<emphasis id="4CB2EA8CFFFEFFEA9E38FE50FCFD40D4" box="[798,878,475,501]" italics="true" pageId="18" pageNumber="73">C. keni</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9E5AFE50FBDA40D4" author="Hafez, S. M. &amp; Elbadry, E. A. &amp; Nasr, A. K." box="[892,1097,475,501]" pageId="18" pageNumber="73" pagination="1 - 15" refId="ref18227" refString="Hafez, S. M., Elbadry, E. A. &amp; Nasr, A. K. (1982) Soil mites of the family Laelapidae from Egypt (Acari: Mesostigmata). Research Bulletin, Faculty of Agriculture, Ain Shams University, 1759, 1 - 15." type="journal article" year="1982">
Hafez
<emphasis id="4CB2EA8CFFFEFFEA9EE1FE50FB9340D4" box="[967,1024,475,501]" italics="true" pageId="18" pageNumber="73">et al.</emphasis>
, 1982
</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA997BFE50FA0440D4" authority="(Karg, 1994)" baseAuthorityName="Karg" baseAuthorityYear="1994" box="[1117,1431,475,501]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="leptolingua">
<emphasis id="4CB2EA8CFFFEFFEA997BFE50FB6D40D4" box="[1117,1278,475,501]" italics="true" pageId="18" pageNumber="73">C. leptolingua</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA982AFE50FA1C40D4" author="Karg, W." box="[1292,1423,475,501]" pageId="18" pageNumber="73" pagination="179 - 216" refId="ref19084" refString="Karg, W. (1994) Raubmilben der cohors Gamasina Leach (Acarina, Parasitiformes) vom Galapagos-Archipel. Mitteilungen aus dem Zoologischen Museum in Berlin, 70, 179 - 216. https: // doi. org / 10.1002 / mmnz. 19940700202" type="journal article" year="1994">Karg, 1994</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9DB1FE74FE0B4338" authority="(Karg, 1987)" baseAuthorityName="Karg" baseAuthorityYear="1987" box="[151,408,511,537]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="lingua">
<emphasis id="4CB2EA8CFFFEFFEA9DB1FE74FE934338" box="[151,256,511,537]" italics="true" pageId="18" pageNumber="73">C. lingua</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9C28FE74FE034338" author="Karg, W." box="[270,400,511,537]" pageId="18" pageNumber="73" pagination="289 - 302" refId="ref18998" refString="Karg, W. (1987) New predatory mites of the genus Hypoaspis Canestrini, 1884 (Acarina, Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik, Geographie und Biologie der Tiere, 114, 289 - 302." type="journal article" year="1987">Karg, 1987</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9C82FE74FD4A4338" authority="(Karg, 1979)" baseAuthorityName="Karg" baseAuthorityYear="1979" box="[420,729,511,537]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="longodigiti">
<emphasis id="4CB2EA8CFFFEFFEA9C82FE74FDAC4338" box="[420,575,511,537]" italics="true" pageId="18" pageNumber="73">C. longodigiti</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9F68FE74FD424338" author="Karg, W." box="[590,721,511,537]" pageId="18" pageNumber="73" pagination="65 - 104" refId="ref18856" refString="Karg, W. (1979) Die Gattung Hypoaspis Canestrini, 1884 (Acarina, Parasitiformes). Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 106, 65 - 104." type="journal article" year="1979">Karg, 1979</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9FC2FE74FB9F4338" authority="(Karg, 1988)" baseAuthorityName="Karg" baseAuthorityYear="1988" box="[740,1036,511,537]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="mahuncai">
<emphasis id="4CB2EA8CFFFEFFEA9FC2FE74FCE04338" box="[740,883,511,537]" italics="true" pageId="18" pageNumber="73">C. mahuncai</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9EA4FE74FB974338" author="Karg, W." box="[898,1028,511,537]" pageId="18" pageNumber="73" pagination="509 - 526" refId="ref19039" refString="Karg, W. (1988) Die Arten der Raubmilbenuntergattung Cosmolaelaps Berlese, 1903 (Gattung Hypoaspis Canestrini, 1884, Acarina, Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik, Geographie und Biologie der Tiere, 115, 509 - 526." type="journal article" year="1988">Karg, 1988</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA993EFE74FA044338" authority="(Gu &amp; Bai, 1992)" baseAuthorityName="Gu &amp; Bai" baseAuthorityYear="1992" box="[1048,1431,511,537]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="paracuneifer">
<emphasis id="4CB2EA8CFFFEFFEA993EFE74FB5A4338" box="[1048,1225,511,537]" italics="true" pageId="18" pageNumber="73">C. paracuneifer</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA99FEFE74FA1C4338" author="Gu, Y. - M. &amp; Bai, X. - L." box="[1240,1423,511,537]" pageId="18" pageNumber="73" pagination="189 - 195" refId="ref18188" refString="Gu, Y. - M. &amp; Bai, X. - L. (1992) Two new species of Hypoaspis from Formicary. Acta Zootaxonomica Sinica, 17 (2), 189 - 195." type="journal article" year="1992">Gu &amp; Bai, 1992</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9DB1FDA8FE54431C" authority="(Karg, 1979)" baseAuthorityName="Karg" baseAuthorityYear="1979" box="[151,455,547,573]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="4CB2EA8CFFFEFFEA9DB1FDA8FEBC431C" box="[151,303,547,573]" italics="true" pageId="18" pageNumber="73">C. penicillata</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9C1BFDA8FE2C431C" author="Karg, W." box="[317,447,547,573]" pageId="18" pageNumber="73" pagination="65 - 104" refId="ref18856" refString="Karg, W. (1979) Die Gattung Hypoaspis Canestrini, 1884 (Acarina, Parasitiformes). Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 106, 65 - 104." type="journal article" year="1979">Karg, 1979</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9CF4FDA8FCC3431C" authority="(Xu &amp; Liang, 1996)" baseAuthorityName="Xu &amp; Liang" baseAuthorityYear="1996" box="[466,848,547,573]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="4CB2EA8CFFFEFFEA9CF4FDA8FDFA431C" box="[466,617,547,573]" italics="true" pageId="18" pageNumber="73">C. reticulatus</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9F51FDA8FCD4431C" author="Xu, X. &amp; Liang, L." box="[631,839,547,573]" pageId="18" pageNumber="73" pagination="189 - 197" refId="ref20543" refString="Xu, X. &amp; Liang, L. (1996) Four new species of the Hypoaspidinae (Acari: Laelapidae) from moss in China. Systematic &amp; Applied Acarology, 1, 189 - 197. https: // doi. org / 10.11158 / saa. 1.1.23" type="journal article" year="1996">Xu &amp; Liang, 1996</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B9C64D1DFFFEFFEA9E7DFDA8FB61431C" authority="(Ma &amp; Lin, 2009)" baseAuthorityName="Ma &amp; Lin" baseAuthorityYear="2009" box="[859,1266,547,573]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="robustochaetes">
<emphasis id="4CB2EA8CFFFEFFEA9E7DFDA8FBB7431C" box="[859,1060,547,573]" italics="true" pageId="18" pageNumber="73">C. robustochaetes</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9917FDA8FB79431C" author="Ma, L. - M. &amp; Lin, J. - Z." box="[1073,1258,547,573]" pageId="18" pageNumber="73" pagination="28 - 39" refId="ref19686" refString="Ma, L. - M. &amp; Lin, J. - Z. (2009) On five new species of the genera Hypoaspis and Cosmolaelaps, with supplementary descriptions of Cosmolaelaps xiajiangensis and Haemolaelaps cordatus (Acari: Mesostigmata: Laelapidae). Acta Arachnologica Sinica, 18, 28 - 39. [in Chinese]" type="journal article" year="2009">Ma &amp; Lin, 2009</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="B9C64D1DFFFEFFEA980DFDA8FE104340" authority="(Pearse et al., 1936)" baseAuthorityName="Pearse" baseAuthorityYear="1936" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="trifidus">
<emphasis id="4CB2EA8CFFFEFFEA980DFDA8FA0F431C" box="[1323,1436,547,573]" italics="true" pageId="18" pageNumber="73">C. trifidus</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9DB9FDCCFEE84340" author="Pearse, A. S. &amp; Patterson, M. T. &amp; Rankin, J. S. &amp; Wharton, G. W." box="[159,379,583,609]" pageId="18" pageNumber="73" pagination="455 - 490" refId="ref20174" refString="Pearse, A. S., Patterson, M. T., Rankin, J. S. &amp; Wharton, G. W. (1936) The ecology of Passalus cornutus Fabricius, a beetle which lives in rotting logs. Ecological Monographs, 6, 455 - 490. https: // doi. org / 10.2307 / 1943239" type="journal article" year="1936">
Pearse
<emphasis id="4CB2EA8CFFFEFFEA9DD5FDCCFEBD4340" box="[243,302,583,609]" italics="true" pageId="18" pageNumber="73">et al.</emphasis>
, 1936
</bibRefCitation>
)
</taxonomicName>
. However,
<taxonomicName id="B9C64D1DFFFEFFEA9F21FDCCFD514340" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[519,706,583,609]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="hallasanensis">
<emphasis id="4CB2EA8CFFFEFFEA9F21FDCCFD514340" box="[519,706,583,609]" italics="true" pageId="18" pageNumber="73">C. hallasanensis</emphasis>
</taxonomicName>
post-anal can be easily distinguished from all those species by the following combination of characters: (1) dorsal shield bearing 40 pairs of spatulate-mucronate setae, including three pairs of
<emphasis id="4CB2EA8CFFFEFFEA9C17FD04FEDE4389" box="[305,333,655,680]" italics="true" pageId="18" pageNumber="73">Zx</emphasis>
setae and six unpaired supernumerary seta (
<emphasis id="4CB2EA8CFFFEFFEA9E1BFD04FCC54389" box="[829,854,655,680]" italics="true" pageId="18" pageNumber="73">Jx</emphasis>
), (2) sternal shield bearing three pairs of long setae, extending beyond of next posterior setae, (3) genital shield flask-shaped, widened posteriorly behind setae
<emphasis id="4CB2EA8CFFFEFFEA986AFD38FAFE43EC" box="[1356,1389,691,717]" italics="true" pageId="18" pageNumber="73">st5</emphasis>
, (4) hypostomal groove greatly widened, with six transverse rows of denticles, each row with 1035 tiny denticles, (5) fixed digit of chelicera with 1012 teeth of various sizes, (6) femur II in both female and male with strong spur-like seta on the ventral side (
<emphasis id="4CB2EA8CFFFEFFEA9C8FFC94FE504219" box="[425,451,799,824]" italics="true" pageId="18" pageNumber="73">av</emphasis>
), (7) Most of setae on legs spatulate.
<taxonomicName id="B9C64D1DFFFEFFEA9E4FFC94FAFE4218" authority="(Gu &amp; Bai, 1992)" baseAuthorityName="Gu &amp; Bai" baseAuthorityYear="1992" box="[873,1389,799,825]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="paracuneifer">
<emphasis id="4CB2EA8CFFFEFFEA9E4FFC94FB334218" box="[873,1184,799,825]" italics="true" pageId="18" pageNumber="73">Cosmolaelaps paracuneifer</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9989FC94FAF74218" author="Gu, Y. - M. &amp; Bai, X. - L." box="[1199,1380,799,825]" pageId="18" pageNumber="73" pagination="189 - 195" refId="ref18188" refString="Gu, Y. - M. &amp; Bai, X. - L. (1992) Two new species of Hypoaspis from Formicary. Acta Zootaxonomica Sinica, 17 (2), 189 - 195." type="journal article" year="1992">Gu &amp; Bai, 1992</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="B9C64D1DFFFEFFEA9DB1FCC8FE57427C" authority="(Berlese, 1883)" baseAuthorityName="Berlese" baseAuthorityYear="1883" box="[151,452,835,861]" class="Arachnida" family="Laelapidae" genus="Laelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="claviger">
<emphasis id="4CB2EA8CFFFEFFEA9DB1FCC8FE80427C" box="[151,275,835,861]" italics="true" pageId="18" pageNumber="73">C. claviger</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9C07FCC8FE2F427C" author="Berlese, A." box="[289,444,835,861]" pageId="18" pageNumber="73" refId="ref17531" refString="Berlese, A. (1883) s. n. Acari, Myriapoda et Scorpiones Hucusque in Italia Reperta, 4, 2. https: // doi. org / 10.5962 / bhl. title. 69269" type="book" year="1883">Berlese, 1883</bibRefCitation>
)
</taxonomicName>
share a general resemblance to
<emphasis id="4CB2EA8CFFFEFFEA9E02FCC8FC77427C" box="[804,996,835,861]" italics="true" pageId="18" pageNumber="73">
<taxonomicName id="B9C64D1DFFFEFFEA9E02FCC8FC73427C" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[804,992,835,861]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="hallasanensis">C. hallasanensis</taxonomicName>
.
</emphasis>
However, there are notable distinctions between the three species.
<taxonomicName id="B9C64D1DFFFEFFEA9CE4FCECFD6242A0" authority="(Berlese, 1883)" baseAuthorityName="Berlese" baseAuthorityYear="1883" box="[450,753,871,897]" class="Arachnida" family="Laelapidae" genus="Laelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="claviger">
<emphasis id="4CB2EA8CFFFEFFEA9CE4FCECFDAD42A0" box="[450,574,871,897]" italics="true" pageId="18" pageNumber="73">C. claviger</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9F6BFCECFD7A42A0" author="Berlese, A." box="[589,745,871,897]" pageId="18" pageNumber="73" refId="ref17531" refString="Berlese, A. (1883) s. n. Acari, Myriapoda et Scorpiones Hucusque in Italia Reperta, 4, 2. https: // doi. org / 10.5962 / bhl. title. 69269" type="book" year="1883">Berlese, 1883</bibRefCitation>
)
</taxonomicName>
possesses 39 pairs of spatulate dorsal setae inserted on small tubercles, whereas
<taxonomicName id="B9C64D1DFFFEFFEA9C4DFC00FDB04284" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[363,547,907,933]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="hallasanensis">
<emphasis id="4CB2EA8CFFFEFFEA9C4DFC00FDB04284" box="[363,547,907,933]" italics="true" pageId="18" pageNumber="73">C. hallasanensis</emphasis>
</taxonomicName>
has 40 pairs of spatulate-mucronate setae. Additionally,
<taxonomicName id="B9C64D1DFFFEFFEA99B3FC00FADE4284" authorityName="Oh &amp; Joharchi &amp; Lee &amp; Jung" authorityYear="2025" box="[1173,1357,907,933]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="hallasanensis">
<emphasis id="4CB2EA8CFFFEFFEA99B3FC00FADE4284" box="[1173,1357,907,933]" italics="true" pageId="18" pageNumber="73">C. hallasanensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="B9C64D1DFFFEFFEA9DF3FC24FDC042E8" authority="(Gu &amp; Bai, 1992)" baseAuthorityName="Gu &amp; Bai" baseAuthorityYear="1992" box="[213,595,943,969]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="paracuneifer">
<emphasis id="4CB2EA8CFFFEFFEA9DF3FC24FE1542E8" box="[213,390,943,969]" italics="true" pageId="18" pageNumber="73">C. paracuneifer</emphasis>
(
<bibRefCitation id="1A574B6FFFFEFFEA9CB3FC24FDD942E8" author="Gu, Y. - M. &amp; Bai, X. - L." box="[405,586,943,969]" pageId="18" pageNumber="73" pagination="189 - 195" refId="ref18188" refString="Gu, Y. - M. &amp; Bai, X. - L. (1992) Two new species of Hypoaspis from Formicary. Acta Zootaxonomica Sinica, 17 (2), 189 - 195." type="journal article" year="1992">Gu &amp; Bai, 1992</bibRefCitation>
)
</taxonomicName>
in the shape of its dorsal setae.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,468 @@
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<mods:title id="613922CD1E8B87F6883D8CDA7A2D2F9C">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from the Republic of Korea, with a review of the Korean species of the genus</mods:title>
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<subSubSection id="36DC6515FFFBFFEF9DB1FF1CFD594190" box="[151,714,151,178]" pageId="23" pageNumber="78" type="nomenclature">
<paragraph id="7E79369EFFFBFFEF9DB1FF1CFD594190" blockId="23.[151,714,151,214]" box="[151,714,151,178]" pageId="23" pageNumber="78">
<heading id="253181F2FFFBFFEF9DB1FF1CFD594190" bold="true" box="[151,714,151,178]" fontSize="11" level="1" pageId="23" pageNumber="78" reason="1">
<taxonomicName id="B9C64D1DFFFBFFEF9DB1FF1CFD594190" authority="Ma &amp; Lin, 2009" authorityName="Ma &amp; Lin" authorityYear="2009" box="[151,714,151,178]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="78" phylum="Arthropoda" rank="species" species="robustochaetes">
<emphasis id="4CB2EA8CFFFBFFEF9DB1FF1CFD594190" bold="true" box="[151,714,151,178]" pageId="23" pageNumber="78">
<emphasis id="4CB2EA8CFFFBFFEF9DB1FF1CFE684193" bold="true" box="[151,507,151,178]" italics="true" pageId="23" pageNumber="78">Cosmolaelaps robustochaetes</emphasis>
<bibRefCitation id="1A574B6FFFFBFFEF9F25FF1CFD594190" author="Ma, L. - M. &amp; Lin, J. - Z." box="[515,714,151,177]" pageId="23" pageNumber="78" pagination="28 - 39" refId="ref19686" refString="Ma, L. - M. &amp; Lin, J. - Z. (2009) On five new species of the genera Hypoaspis and Cosmolaelaps, with supplementary descriptions of Cosmolaelaps xiajiangensis and Haemolaelaps cordatus (Acari: Mesostigmata: Laelapidae). Acta Arachnologica Sinica, 18, 28 - 39. [in Chinese]" type="journal article" year="2009">Ma &amp; Lin, 2009</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="36DC6515FFFBFFEF9DB1FF30FED841F7" box="[151,331,187,214]" pageId="23" pageNumber="78" type="description">
<paragraph id="7E79369EFFFBFFEF9DB1FF30FED841F7" blockId="23.[151,714,151,214]" box="[151,331,187,214]" pageId="23" pageNumber="78">
(
<figureCitation id="E6FD2A1BFFFBFFEF9DB9FF30FED141F7" box="[159,322,187,214]" captionStart-0="FIGURES 6872" captionStart-1="FIGURES 7379" captionStart-2="FIGURES 8083" captionStart-3="FIGURES 8485" captionStartId-0="24.[152,270,1943,1967]" captionStartId-1="25.[152,270,1938,1962]" captionStartId-2="26.[152,270,1844,1868]" captionStartId-3="28.[152,270,1456,1480]" captionTargetBox-0="[187,1403,197,1908]" captionTargetBox-1="[187,1405,189,1910]" captionTargetBox-2="[164,1430,197,1805]" captionTargetBox-3="[168,1419,198,1415]" captionTargetId-0="figure-26@24.[172,1416,181,1919]" captionTargetId-1="figure-17@25.[179,1408,181,1915]" captionTargetId-2="figure-26@26.[151,1435,181,1821]" captionTargetId-3="figure-26@28.[151,1436,181,1432]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="28" captionText-0="FIGURES 6872. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 68—Dorsal idiosoma; 69—Ventral idiosoma; 70— Ventral gnathosoma; 71—Epistome; 72—Chelicera." captionText-1="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." captionText-2="FIGURES 8083. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 80—Leg I; 81—Leg II; 82—Leg III; 83—Leg IV." captionText-3="FIGURES 8485. Cosmolaelaps robustochaetes Ma &amp; Lin, adult male: 84—Ventral idiosoma; 85—Chelicera." figureDoi-0="http://doi.org/10.5281/zenodo.14703951" figureDoi-1="http://doi.org/10.5281/zenodo.14703953" figureDoi-2="http://doi.org/10.5281/zenodo.14703955" figureDoi-3="http://doi.org/10.5281/zenodo.14703957" httpUri-0="https://zenodo.org/record/14703951/files/figure.png" httpUri-1="https://zenodo.org/record/14703953/files/figure.png" httpUri-2="https://zenodo.org/record/14703955/files/figure.png" httpUri-3="https://zenodo.org/record/14703957/files/figure.png" pageId="23" pageNumber="78">Figures 6885</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="36DC6515FFFBFFEF9DB1FE8BFB2E40E0" pageId="23" pageNumber="78" type="reference_group">
<paragraph id="7E79369EFFFBFFEF9DB1FE8BFD0D4039" blockId="23.[151,1033,256,341]" box="[151,670,256,280]" pageId="23" pageNumber="78">
<treatmentCitationGroup id="5ED611B0FFFBFFEF9DB1FE8BFD0D4039" box="[151,670,256,280]" pageId="23" pageNumber="78">
<treatmentCitation id="FF67108FFFFBFFEF9DB1FE8BFD094039" author="Ma, L. - M. &amp; Lin, J. - Z." box="[151,666,256,280]" page="29" pageId="23" pageNumber="78" year="2009">
<taxonomicName id="B9C64D1DFFFBFFEF9DB1FE8BFD094039" authority="Ma &amp; Lin, 2009: 29" authorityName="Ma &amp; Lin" authorityPageNumber="29" authorityYear="2009" box="[151,666,256,280]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="78" phylum="Arthropoda" rank="species" species="robustochaetes">
<emphasis id="4CB2EA8CFFFBFFEF9DB1FE8BFE564039" box="[151,453,256,280]" italics="true" pageId="23" pageNumber="78">Cosmolaelaps robustochaetes</emphasis>
<bibRefCitation id="1A574B6FFFFBFFEF9CEAFE8BFD094039" author="Ma, L. - M. &amp; Lin, J. - Z." box="[460,666,256,280]" pageId="23" pageNumber="78" pagination="28 - 39" refId="ref19686" refString="Ma, L. - M. &amp; Lin, J. - Z. (2009) On five new species of the genera Hypoaspis and Cosmolaelaps, with supplementary descriptions of Cosmolaelaps xiajiangensis and Haemolaelaps cordatus (Acari: Mesostigmata: Laelapidae). Acta Arachnologica Sinica, 18, 28 - 39. [in Chinese]" type="journal article" year="2009">Ma &amp; Lin, 2009: 29</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7E79369EFFFBFFEF9DB1FE94FB9A4017" blockId="23.[151,1033,256,341]" box="[151,1033,286,310]" pageId="23" pageNumber="78">
<treatmentCitationGroup id="5ED611B0FFFBFFEF9DB1FE94FB9A4017" box="[151,1033,286,310]" pageId="23" pageNumber="78">
<taxonomicName id="B9C64D1DFFFBFFEF9DB1FE94FDD44017" box="[151,583,286,310]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="78" phylum="Arthropoda" rank="species" species="robustochaetes" subGenus="Cosmolaelaps">
<emphasis id="4CB2EA8CFFFBFFEF9DB1FE94FE914017" box="[151,258,287,310]" italics="true" pageId="23" pageNumber="78">Hypoaspis</emphasis>
(
<emphasis id="4CB2EA8CFFFBFFEF9C37FE95FE324017" box="[273,417,286,310]" italics="true" pageId="23" pageNumber="78">Cosmolaelaps</emphasis>
)
<emphasis id="4CB2EA8CFFFBFFEF9C89FE95FDD44017" box="[431,583,286,310]" italics="true" pageId="23" pageNumber="78">robustochaetes</emphasis>
</taxonomicName>
.—
<treatmentCitation id="FF67108FFFFBFFEF9F40FE95FCD14017" author="Bai, X. - L. &amp; Ma, L. - M." box="[614,834,286,310]" page="557" pageId="23" pageNumber="78" year="2012">
<bibRefCitation id="1A574B6FFFFBFFEF9F40FE95FCD14017" author="Bai, X. - L. &amp; Ma, L. - M." box="[614,834,286,310]" pageId="23" pageNumber="78" pagination="555 - 558" refId="ref17319" refString="Bai, X. - L. &amp; Ma, L. - M. (2012) A new species of the genus Hypoaspis from Ningxia, China with supplementary description on Hypoaspis subpictus Gu et Bai (Acari: Laelapidae). Acta Zootaxonomica Sinica, 37, 555 - 558. [in Chinese]" type="journal article" year="2012">Bai &amp; Ma, 2012: 557</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="FF67108FFFFBFFEF9E6BFE95FB964017" author="Bai, X. - L. &amp; Yan, Y. &amp; Zhang, T." box="[845,1029,286,310]" page="9" pageId="23" pageNumber="78" year="2016">
<bibRefCitation id="1A574B6FFFFBFFEF9E6BFE95FB964017" author="Bai, X. - L. &amp; Yan, Y. &amp; Zhang, T." box="[845,1029,286,310]" pageId="23" pageNumber="78" pagination="6 - 13" refId="ref17377" refString="Bai, X. - L., Yan, Y. &amp; Zhang, T. (2016) Two new species of Hypoaspis from Helan Mountain Nature Reserve of Ningxia, China (Acari: Laelapidae). Acta Arachnologica Sinica, 25, 6 - 13. [in Chinese]" type="journal article" year="2016">
Bai
<emphasis id="4CB2EA8CFFFBFFEF9E50FE94FC394017" box="[886,938,286,310]" italics="true" pageId="23" pageNumber="78">et al.</emphasis>
, 2016: 9
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7E79369EFFFBFFEF9DB1FEB6FD794074" blockId="23.[151,1033,256,341]" box="[151,746,317,341]" pageId="23" pageNumber="78">
<treatmentCitationGroup id="5ED611B0FFFBFFEF9DB1FEB6FD794074" box="[151,746,317,341]" pageId="23" pageNumber="78">
<taxonomicName id="B9C64D1DFFFBFFEF9DB1FEB6FE564074" authorityName="Ma &amp; Lin" authorityYear="2009" box="[151,453,317,341]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="78" phylum="Arthropoda" rank="species" species="robustochaetes">
<emphasis id="4CB2EA8CFFFBFFEF9DB1FEB6FE564074" box="[151,453,317,341]" italics="true" pageId="23" pageNumber="78">Cosmolaelaps robustochaetes</emphasis>
</taxonomicName>
.—
<treatmentCitation id="FF67108FFFFBFFEF9CC5FEB6FD764074" author="Moreira, G. F. &amp; Klompen, H. &amp; de Moraes, G. J." box="[483,741,317,341]" page="320" pageId="23" pageNumber="78" year="2014">
<bibRefCitation id="1A574B6FFFFBFFEF9CC5FEB6FD764074" author="Moreira, G. F. &amp; Klompen, H. &amp; de Moraes, G. J." box="[483,741,317,341]" pageId="23" pageNumber="78" pagination="317 - 346" refId="ref19966" refString="Moreira, G. F., Klompen, H. &amp; de Moraes, G. J. (2014) Redefinition of Cosmolaelaps Berlese (Acari: Laelapidae) and description of five new species from Brazil. Zootaxa, 3764 (3), 317 - 346. https: // doi. org / 10.11646 / zootaxa. 3764.3.4" type="journal article" year="2014">
Moreira
<emphasis id="4CB2EA8CFFFBFFEF9F1BFEB5FDE34074" box="[573,624,317,341]" italics="true" pageId="23" pageNumber="78">et al.</emphasis>
, 2014: 320
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7E79369EFFFBFFEF9DB1FE08FB2E40E0" blockId="23.[151,1437,387,774]" pageId="23" pageNumber="78">
<emphasis id="4CB2EA8CFFFBFFEF9DB1FE08FE0B40BC" bold="true" box="[151,408,387,413]" pageId="23" pageNumber="78">Specimens examined.</emphasis>
<materialsCitation id="CEAE3CC3FFFBFFEF9C86FE08FB2A40E0" collectingDate="2021-09-29" collectorName="Ulleung-do, GB &amp; J. Oh" country="South Korea" county="Ulleung-gun" latitude="37.496666" location="San" longLatPrecision="19" longitude="130.89166" municipality="Ulleung-eup" pageId="23" pageNumber="78" specimenCount="19" specimenCount-female="11" specimenCount-male="8">
<specimenCount id="68C0FD17FFFBFFEF9C86FE08FDB740BC" box="[416,548,387,413]" count="11" pageId="23" pageNumber="78" type="female">11 Females</specimenCount>
,
<specimenCount id="68C0FD17FFFBFFEF9F16FE08FD2640BC" box="[560,693,387,413]" count="8" pageId="23" pageNumber="78" type="male">eight males</specimenCount>
,
<location id="7B196045FFFBFFEF9FE4FE08FD7F40BC" LSID="urn:lsid:plazi:treatment:F66F8788FFFBFFE39DB1FF1CFDB047DE:7B196045FFFBFFEF9FE4FE08FD7F40BC" box="[706,748,387,413]" country="South Korea" county="Ulleung-gun" latitude="37.496666" longLatPrecision="19" longitude="130.89166" municipality="Ulleung-eup" name="San" pageId="23" pageNumber="78">San</location>
39,
<location id="7B196045FFFBFFEF9E06FE08FC0840BC" LSID="urn:lsid:plazi:treatment:F66F8788FFFBFFE39DB1FF1CFDB047DE:7B196045FFFBFFEF9E06FE08FC0840BC" box="[800,923,387,413]" country="South Korea" county="Ulleung-gun" latitude="37.496666" longLatPrecision="19" longitude="130.89166" municipality="Ulleung-eup" name="Jeodong-ri" pageId="23" pageNumber="78">Jeodong-ri</location>
,
<collectingMunicipality id="9E1DACE4FFFBFFEF9E81FE08FBA540BC" box="[935,1078,387,413]" pageId="23" pageNumber="78">Ulleung-eup</collectingMunicipality>
,
<collectingCounty id="97184E12FFFBFFEF9964FE08FB4140BC" box="[1090,1234,387,413]" pageId="23" pageNumber="78">Ulleung-gun</collectingCounty>
,
<collectorName id="D3335348FFFBFFEF99F9FE08FA0440BC" box="[1247,1431,387,413]" pageId="23" pageNumber="78">Ulleung-do, GB</collectorName>
,
<collectingCountry id="06D1760EFFFBFFEF9DB1FE2CFEFA40E3" box="[151,361,423,450]" name="South Korea" pageId="23" pageNumber="78">Republic of Korea</collectingCountry>
,
<geoCoordinate id="1BF25059FFFBFFEF9C52FE2CFE6440E0" box="[372,503,423,449]" degrees="37" direction="north" minutes="29" orientation="latitude" pageId="23" pageNumber="78" precision="15" seconds="48" value="37.496666">37°29'48&quot;N</geoCoordinate>
<geoCoordinate id="1BF25059FFFBFFEF9CD9FE2CFD0340E3" box="[511,656,423,450]" degrees="130" direction="east" minutes="53" orientation="longitude" pageId="23" pageNumber="78" precision="15" seconds="30" value="130.89166">130°53'30&quot;E</geoCoordinate>
,
<date id="0A78105EFFFBFFEF9FBAFE2CFCEB40E0" box="[668,888,423,449]" pageId="23" pageNumber="78" value="2021-09-29">
<collectingDate id="1A3CE9B6FFFBFFEF9FBAFE2CFCEB40E0" box="[668,888,423,449]" pageId="23" pageNumber="78" value="2021-09-29">29 September 2021</collectingDate>
</date>
,
<collectorName id="D3335348FFFBFFEF9EA5FE23FC2D40E0" box="[899,958,423,450]" pageId="23" pageNumber="78">J. Oh</collectorName>
coll., soil &amp; leaf litter
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="36DC6515FFFBFFEF9DE1FE40FE694227" pageId="23" pageNumber="78" type="diagnosis">
<paragraph id="7E79369EFFFBFFEF9DE1FE40FE694227" blockId="23.[151,1437,387,774]" pageId="23" pageNumber="78">
<emphasis id="4CB2EA8CFFFBFFEF9DE1FE40FE0540C4" bold="true" box="[199,406,459,485]" pageId="23" pageNumber="78">Diagnosis (adult).</emphasis>
Dorsal shield covered with scale-like reticulation posteriorly,with 41 pairs of spatulate, curved scimitar shaped setae (except
<emphasis id="4CB2EA8CFFFBFFEF9CC1FE64FE6E4328" box="[487,509,495,521]" italics="true" pageId="23" pageNumber="78">j1</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9F2DFE64FDB74328" box="[523,548,495,521]" italics="true" pageId="23" pageNumber="78">z1</emphasis>
, and
<emphasis id="4CB2EA8CFFFBFFEF9F44FE7BFD13432B" box="[610,640,496,522]" italics="true" pageId="23" pageNumber="78">Z5</emphasis>
), including 10 unpaired
<emphasis id="4CB2EA8CFFFBFFEF9EB0FE7BFC3C4328" box="[918,943,496,521]" italics="true" pageId="23" pageNumber="78">Jx</emphasis>
setae; presternal area palely sclerotised and eyebrow-shaped, sternal shield with lineate-reticulate ornamentation on lateral surface, distinctly punctate centrally, ratio of shield length/width (at broadest level) ≃ 0.68; genital shield flask-shaped strongly expanded behind coxa IV, surface of shield with irregular polygonal reticulation, shield ratio of length/width (at broadest level) ≃ 1.48, anal shield slightly longer than wide, with ratio of length/width (at broadest level) ≃ 1.24; opisthogastric setae spatulate, except
<emphasis id="4CB2EA8CFFFBFFEF9DCEFD28FE9D439C" box="[232,270,675,701]" italics="true" pageId="23" pageNumber="78">Jv1</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9C3BFD28FED7439C" box="[285,324,675,701]" italics="true" pageId="23" pageNumber="78">Jv2</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9C75FD28FEEE439C" box="[339,381,675,701]" italics="true" pageId="23" pageNumber="78">Zv1</emphasis>
, and
<emphasis id="4CB2EA8CFFFBFFEF9C9DFD28FE75439C" box="[443,486,675,701]" italics="true" pageId="23" pageNumber="78">Zv2</emphasis>
. Fixed digit of chelicera with 910 teeth of various sizes. Most of legs setae simple, except some spatulate setae on trochanter I, femur IIV, and spine like av seta on femur II. In male, holoventral shield bearing 10 pairs of setae.
</paragraph>
</subSubSection>
<subSubSection id="36DC6515FFFBFFE39DB1FCDCFDB047DE" lastPageId="27" lastPageNumber="82" pageId="23" pageNumber="78" type="description">
<paragraph id="7E79369EFFFBFFEF9DB1FCDCFE4C4250" blockId="23.[151,479,855,917]" box="[151,479,855,881]" pageId="23" pageNumber="78">
<emphasis id="4CB2EA8CFFFBFFEF9DB1FCDCFE4C4250" bold="true" box="[151,479,855,881]" pageId="23" pageNumber="78">Description (adult female)</emphasis>
</paragraph>
<paragraph id="7E79369EFFFBFFEF9DB1FCF0FED842B4" blockId="23.[151,479,855,917]" box="[151,331,891,917]" pageId="23" pageNumber="78">
(
<figureCitation id="E6FD2A1BFFFBFFEF9DB9FCF0FED042B4" box="[159,323,891,917]" captionStart-0="FIGURES 6872" captionStart-1="FIGURES 7379" captionStart-2="FIGURES 8083" captionStartId-0="24.[152,270,1943,1967]" captionStartId-1="25.[152,270,1938,1962]" captionStartId-2="26.[152,270,1844,1868]" captionTargetBox-0="[187,1403,197,1908]" captionTargetBox-1="[187,1405,189,1910]" captionTargetBox-2="[164,1430,197,1805]" captionTargetId-0="figure-26@24.[172,1416,181,1919]" captionTargetId-1="figure-17@25.[179,1408,181,1915]" captionTargetId-2="figure-26@26.[151,1435,181,1821]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="FIGURES 6872. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 68—Dorsal idiosoma; 69—Ventral idiosoma; 70— Ventral gnathosoma; 71—Epistome; 72—Chelicera." captionText-1="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." captionText-2="FIGURES 8083. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 80—Leg I; 81—Leg II; 82—Leg III; 83—Leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.14703951" figureDoi-1="http://doi.org/10.5281/zenodo.14703953" figureDoi-2="http://doi.org/10.5281/zenodo.14703955" httpUri-0="https://zenodo.org/record/14703951/files/figure.png" httpUri-1="https://zenodo.org/record/14703953/files/figure.png" httpUri-2="https://zenodo.org/record/14703955/files/figure.png" pageId="23" pageNumber="78">Figures 6883</figureCitation>
)
</paragraph>
<paragraph id="7E79369EFFFBFFEF9DB1FC4BFE4942F9" blockId="23.[151,474,960,984]" box="[151,474,960,984]" pageId="23" pageNumber="78">
<specimenCount id="68C0FD17FFFBFFEF9DB1FC4BFEA642F9" box="[151,309,960,984]" count="5" pageId="23" pageNumber="78" type="generic">Five specimens</specimenCount>
were measured.
</paragraph>
<paragraph id="7E79369EFFFBFFEF9DB1FB8DFC66443D" blockId="23.[151,1437,1029,2028]" pageId="23" pageNumber="78">
<emphasis id="4CB2EA8CFFFBFFEF9DB1FB8DFEC9453E" bold="true" box="[151,346,1029,1056]" pageId="23" pageNumber="78">Dorsal idiosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFFBFFEF9C4CFB8DFE4F4501" box="[362,476,1030,1056]" captionStart="FIGURES 6872" captionStartId="24.[152,270,1943,1967]" captionTargetBox="[187,1403,197,1908]" captionTargetId="figure-26@24.[172,1416,181,1919]" captionTargetPageId="24" captionText="FIGURES 6872. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 68—Dorsal idiosoma; 69—Ventral idiosoma; 70— Ventral gnathosoma; 71—Epistome; 72—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703951" httpUri="https://zenodo.org/record/14703951/files/figure.png" pageId="23" pageNumber="78">Figure 68</figureCitation>
,
<figureCitation id="E6FD2A1BFFFBFFEF9CCFFB8DFDDF4500" box="[489,588,1030,1057]" captionStart="FIGURES 7379" captionStartId="25.[152,270,1938,1962]" captionTargetBox="[187,1405,189,1910]" captionTargetId="figure-17@25.[179,1408,181,1915]" captionTargetPageId="25" captionText="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703953" httpUri="https://zenodo.org/record/14703953/files/figure.png" pageId="23" pageNumber="78">73 &amp; 75</figureCitation>
). Dorsal shield oval-shaped, 684753 long, 456506 wide, covering entire idiosoma dorsally; posterior part of shield with distinct scale-like reticulation, bearing 41 pairs of strongly curved, spatulate, and scimitar shaped setae (with a visible basal asymmetric protuberance) (
<figureCitation id="E6FD2A1BFFFBFFEF9942FBC5FB444548" box="[1124,1239,1102,1129]" captionStart="FIGURES 7379" captionStartId="25.[152,270,1938,1962]" captionTargetBox="[187,1405,189,1910]" captionTargetId="figure-17@25.[179,1408,181,1915]" captionTargetPageId="25" captionText="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703953" httpUri="https://zenodo.org/record/14703953/files/figure.png" pageId="23" pageNumber="78">Figure 75</figureCitation>
), except
<emphasis id="4CB2EA8CFFFBFFEF9866FBC5FAC54549" box="[1344,1366,1102,1128]" italics="true" pageId="23" pageNumber="78">j1</emphasis>
horse ear-like (3143) and
<emphasis id="4CB2EA8CFFFBFFEF9CA3FBF9FE0D45AD" box="[389,414,1138,1164]" italics="true" pageId="23" pageNumber="78">z1</emphasis>
spatulate acicular, without asymmetric protuberance (2836): 22 pairs of podonotal setae, 19 pairs of ophisthonotal setae, and 10 unpaired supernumerary setae
<emphasis id="4CB2EA8CFFFBFFEF9E9CFB1CFC404591" box="[954,979,1175,1200]" italics="true" pageId="23" pageNumber="78">Jx</emphasis>
between
<emphasis id="4CB2EA8CFFFBFFEF9964FB1DFBCF4591" box="[1090,1116,1174,1200]" italics="true" pageId="23" pageNumber="78">J1</emphasis>
and
<emphasis id="4CB2EA8CFFFBFFEF99B1FB1DFB224591" box="[1175,1201,1174,1200]" italics="true" pageId="23" pageNumber="78">J4</emphasis>
, dorsal setae mostly homogeneous in length, barely reaching base of next posterior setae (4287), podonotal setae more curved and thickened than ophisthonotal setae. Shield with nine pairs of discernible pore-like structures, including five poroids (
<emphasis id="4CB2EA8CFFFBFFEF9D87FA89FF57443D" box="[161,196,1282,1308]" italics="true" pageId="23" pageNumber="78">id4</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9DF4FA89FF65443D" box="[210,246,1282,1308]" italics="true" pageId="23" pageNumber="78">id6</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9C22FA89FEAF443D" box="[260,316,1282,1308]" italics="true" pageId="23" pageNumber="78">idm2</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9C6CFA89FE11443D" box="[330,386,1282,1308]" italics="true" pageId="23" pageNumber="78">idm4</emphasis>
, and
<emphasis id="4CB2EA8CFFFBFFEF9C99FA89FE72443D" box="[447,481,1282,1308]" italics="true" pageId="23" pageNumber="78">idx</emphasis>
) and three gland openings (
<emphasis id="4CB2EA8CFFFBFFEF9E3DFA89FCD6443D" box="[795,837,1282,1308]" italics="true" pageId="23" pageNumber="78">gd2</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9E75FA89FCEE443D" box="[851,893,1282,1308]" italics="true" pageId="23" pageNumber="78">gd5</emphasis>
, and
<emphasis id="4CB2EA8CFFFBFFEF9E9CFA89FC76443D" box="[954,997,1282,1308]" italics="true" pageId="23" pageNumber="78">gd6</emphasis>
).
</paragraph>
<paragraph id="7E79369EFFFBFFE39DE1FAADFADE403C" blockId="23.[151,1437,1029,2028]" lastBlockId="27.[151,1437,151,1041]" lastPageId="27" lastPageNumber="82" pageId="23" pageNumber="78">
<emphasis id="4CB2EA8CFFFBFFEF9DE1FAADFE03441E" bold="true" box="[199,400,1317,1344]" pageId="23" pageNumber="78">Ventral idiosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFFBFFEF9CB9FAADFD844461" box="[415,535,1318,1344]" captionStart="FIGURES 6872" captionStartId="24.[152,270,1943,1967]" captionTargetBox="[187,1403,197,1908]" captionTargetId="figure-26@24.[172,1416,181,1919]" captionTargetPageId="24" captionText="FIGURES 6872. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 68—Dorsal idiosoma; 69—Ventral idiosoma; 70— Ventral gnathosoma; 71—Epistome; 72—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703951" httpUri="https://zenodo.org/record/14703951/files/figure.png" pageId="23" pageNumber="78">Figures 69</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFFBFFEF9F1CFAADFDC94461" box="[570,602,1318,1344]" captionStart="FIGURES 7379" captionStartId="25.[152,270,1938,1962]" captionTargetBox="[187,1405,189,1910]" captionTargetId="figure-17@25.[179,1408,181,1915]" captionTargetPageId="25" captionText="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703953" httpUri="https://zenodo.org/record/14703953/files/figure.png" pageId="23" pageNumber="78">74</figureCitation>
). Tritosternum with paired pilose laciniae (101122), columnar base 4054 long × 811 wide; presternal area partly sclerotised, eyebrow-shaped, sternal shield 98124 long, narrowest between coxae II (101111), widest between coxae II and III (170181), covered with reticulate ornamentation throughout, except central and posterior parts distinctly punctate, anterior margin of shield conspicuous, anterolateral corners narrowly extending between coxae III, posterior margin slightly concave. Shield bearing three pairs of long smooth setae (
<emphasis id="4CB2EA8CFFFBFFEF9DF9FA51FF6C44D5" box="[223,255,1498,1524]" italics="true" pageId="23" pageNumber="78">st1</emphasis>
5765,
<emphasis id="4CB2EA8CFFFBFFEF9C7FFA51FEEA44D5" box="[345,377,1498,1524]" italics="true" pageId="23" pageNumber="78">st2</emphasis>
4149,
<emphasis id="4CB2EA8CFFFBFFEF9CF5FA51FE6044D5" box="[467,499,1498,1524]" italics="true" pageId="23" pageNumber="78">st3</emphasis>
3540), not extending beyond next posterior setae, and two pairs of thin rectangular poroids (
<emphasis id="4CB2EA8CFFFBFFEF9DDBFA75FE8C4739" box="[253,287,1534,1560]" italics="true" pageId="23" pageNumber="78">iv1</emphasis>
and
<emphasis id="4CB2EA8CFFFBFFEF9C7FFA75FEEF4739" box="[345,380,1534,1560]" italics="true" pageId="23" pageNumber="78">iv2</emphasis>
, mesal to setae
<emphasis id="4CB2EA8CFFFBFFEF9F13FA75FDC64739" box="[565,597,1534,1560]" italics="true" pageId="23" pageNumber="78">st1</emphasis>
and between
<emphasis id="4CB2EA8CFFFBFFEF9FD3FA75FC864739" box="[757,789,1534,1560]" italics="true" pageId="23" pageNumber="78">st2</emphasis>
and
<emphasis id="4CB2EA8CFFFBFFEF9E76FA75FCE34739" box="[848,880,1534,1560]" italics="true" pageId="23" pageNumber="78">st3</emphasis>
, respectively), metasternal setae
<emphasis id="4CB2EA8CFFFBFFEF99CCFA75FA994739" box="[1258,1290,1534,1560]" italics="true" pageId="23" pageNumber="78">st4</emphasis>
(3242) and metasternal poroids (
<emphasis id="4CB2EA8CFFFBFFEF9CA6F9A9FE31471D" box="[384,418,1570,1596]" italics="true" pageId="23" pageNumber="78">iv3</emphasis>
) located on soft integument; endopodal elements between coxae IIIII fused with shield and endopodal elements IIIIV elongate, narrow, and curved. Genital shield flask-shaped, distinctly expanded laterally past level of setae
<emphasis id="4CB2EA8CFFFBFFEF9C78F9E1FEEC47A5" box="[350,383,1642,1668]" italics="true" pageId="23" pageNumber="78">st5</emphasis>
, length 218248, maximum width 151164, posterior margin rounded, anterior margin of shield slightly convex, abutting posterior margin of sternal shield, surface reticulated with irregular, mostly longitudinal lines in anterior region, posterior part with irregular polygonal pattern, bearing a pair of simple setae
<emphasis id="4CB2EA8CFFFBFFEF9998F939FB4C47ED" box="[1214,1247,1714,1740]" italics="true" pageId="23" pageNumber="78">st5</emphasis>
(4147) inserted on lateral margins of shield, near level of posterior edge of coxae IV. Paragenital poroids
<emphasis id="4CB2EA8CFFFBFFEF995BF95DFB0C47D1" box="[1149,1183,1750,1776]" italics="true" pageId="23" pageNumber="78">iv5</emphasis>
located on soft cuticle lateral to shield near setae
<emphasis id="4CB2EA8CFFFBFFEF9CE6F971FE734635" box="[448,480,1786,1812]" italics="true" pageId="23" pageNumber="78">st5</emphasis>
; anal shield pear-shaped, rounded anteriorly, 107118 long, 8798 wide, anterior half covered with lineate reticulation, para-anal setae (2330) and post-anal seta (1724) simple, cribrum consisting of a terminal tuft with some irregular rows of spicules and a pair of anterior arms extending to level between anal orifice and post-anal seta. Anal gland pores (
<emphasis id="4CB2EA8CFFFBFFEF9F1AF8EDFDF746A1" box="[572,612,1894,1920]" italics="true" pageId="23" pageNumber="78">gv3</emphasis>
) on anterolateral margin of shield; soft opisthogastric cuticle surrounding genital, anal shields, and a pair of elongated metapodal plates (3039 long × 58 wide), with two pairs of poroids (
<emphasis id="4CB2EA8CFFFBFFEF9D86F825FF5146E9" box="[160,194,1966,1992]" italics="true" pageId="23" pageNumber="78">iv5</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9DF7F824FF6046E9" box="[209,243,1967,1992]" italics="true" pageId="23" pageNumber="78">ivo</emphasis>
) and 13 pairs of spatulate setae (1929) (
<emphasis id="4CB2EA8CFFFBFFEF9FE1F825FCB246E9" box="[711,801,1966,1992]" italics="true" pageId="23" pageNumber="78">Jv3Jv5</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9E16F825FC0246E9" box="[816,913,1966,1992]" italics="true" pageId="23" pageNumber="78">Zv3Zv5</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9E86F825FC4046E9" box="[928,979,1966,1992]" italics="true" pageId="23" pageNumber="78">UR2</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF9EC4F825FB8646E9" box="[994,1045,1966,1992]" italics="true" pageId="23" pageNumber="78">UR3</emphasis>
, and
<emphasis id="4CB2EA8CFFFBFFEF9975F825FB0C46E9" box="[1107,1183,1966,1992]" italics="true" pageId="23" pageNumber="78">R1R5</emphasis>
), except
<emphasis id="4CB2EA8CFFFBFFEF9820F825FAB846E9" box="[1286,1323,1966,1992]" italics="true" pageId="23" pageNumber="78">Jv1</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF981CF825FACD46E9" box="[1338,1374,1966,1992]" italics="true" pageId="23" pageNumber="78">Jv2</emphasis>
,
<emphasis id="4CB2EA8CFFFBFFEF984BF825FA0546E9" box="[1389,1430,1966,1992]" italics="true" pageId="23" pageNumber="78">Zv1</emphasis>
, and
<emphasis id="4CB2EA8CFFFBFFEF9DE3F859FF7E46CD" box="[197,237,2002,2028]" italics="true" pageId="23" pageNumber="78">Zv2</emphasis>
setiform (1931). Two exopodal platelets between coxae IIIII small, subtriangular, those behind coxae IIIIV and parapodal platelets fused, surrounding coxa IV and bearing gland pore (
<emphasis id="4CB2EA8CFFF7FFE39EF9FF1CFB944190" box="[991,1031,151,177]" italics="true" pageId="27" pageNumber="82">gv2</emphasis>
); peritrematal shield well developed laterad peritremes, bearing one gland pore and one poroid at level near coxae IIIII, post-stigmatic extension of shield relatively wide, with two pairs of poroids and one pair of gland pores. Anterior region of peritrematal shield fused to anterior margin of dorsal shield behind setae
<emphasis id="4CB2EA8CFFF7FFE39FB7FE88FD39403C" box="[657,682,259,285]" italics="true" pageId="27" pageNumber="82">z1</emphasis>
, peritremes long, extending to mid-level of coxa I (near
<emphasis id="4CB2EA8CFFF7FFE39802FE88FAAE403C" box="[1316,1341,259,285]" italics="true" pageId="27" pageNumber="82">s1</emphasis>
).
</paragraph>
<caption id="2AB96616FFF4FFE09DBEF81CFD3A46F2" ID-DOI="http://doi.org/10.5281/zenodo.14703951" ID-Zenodo-Dep="14703951" httpUri="https://zenodo.org/record/14703951/files/figure.png" pageId="24" pageNumber="79" startId="24.[152,270,1943,1967]" targetBox="[187,1403,197,1908]" targetPageId="24" targetType="figure">
<paragraph id="7E79369EFFF4FFE09DBEF81CFD3A46F2" blockId="24.[151,1436,1943,2003]" pageId="24" pageNumber="79">
<emphasis id="4CB2EA8CFFF4FFE09DBEF81CFC9A468E" bold="true" box="[152,777,1943,1968]" pageId="24" pageNumber="79">
FIGURES 6872.
<taxonomicName id="B9C64D1DFFF4FFE09C79F81CFC97468E" authority="Ma &amp; Lin" authorityName="Ma &amp; Lin" authorityYear="2009" box="[351,772,1943,1968]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="79" phylum="Arthropoda" rank="species" species="robustochaetes">
<emphasis id="4CB2EA8CFFF4FFE09C79F81CFD01468E" bold="true" box="[351,658,1943,1967]" italics="true" pageId="24" pageNumber="79">Cosmolaelaps robustochaetes</emphasis>
Ma &amp; Lin
</taxonomicName>
,
</emphasis>
adult female: 68—Dorsal idiosoma; 69—Ventral idiosoma; 70— Ventral gnathosoma; 71—Epistome; 72—Chelicera.
</paragraph>
</caption>
<caption id="2AB96616FFF5FFE19DBEF819FA1946EE" ID-DOI="http://doi.org/10.5281/zenodo.14703953" ID-Zenodo-Dep="14703953" httpUri="https://zenodo.org/record/14703953/files/figure.png" pageId="25" pageNumber="80" startId="25.[152,270,1938,1962]" targetBox="[187,1405,189,1910]" targetPageId="25" targetType="figure">
<paragraph id="7E79369EFFF5FFE19DBEF819FA1946EE" blockId="25.[151,1436,1938,1999]" pageId="25" pageNumber="80">
<emphasis id="4CB2EA8CFFF5FFE19DBEF819FC63468A" bold="true" box="[152,1008,1938,1963]" pageId="25" pageNumber="80">
FIGURES 7379. DIC micrographs of
<taxonomicName id="B9C64D1DFFF5FFE19F64F819FC79468A" authority="Ma &amp; Lin" authorityName="Ma &amp; Lin" authorityYear="2009" box="[578,1002,1938,1963]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="80" phylum="Arthropoda" rank="species" species="robustochaetes">
<emphasis id="4CB2EA8CFFF5FFE19F64F819FCE6468B" bold="true" box="[578,885,1938,1962]" italics="true" pageId="25" pageNumber="80">Cosmolaelaps robustochaetes</emphasis>
Ma &amp; Lin
</taxonomicName>
,
</emphasis>
adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera.
</paragraph>
</caption>
<caption id="2AB96616FFF6FFE29DBEF8BFFA09466D" ID-DOI="http://doi.org/10.5281/zenodo.14703955" ID-Zenodo-Dep="14703955" box="[152,1434,1844,1869]" httpUri="https://zenodo.org/record/14703955/files/figure.png" pageId="26" pageNumber="81" startId="26.[152,270,1844,1868]" targetBox="[164,1430,197,1805]" targetPageId="26" targetType="figure">
<paragraph id="7E79369EFFF6FFE29DBEF8BFFA09466D" blockId="26.[152,1434,1844,1869]" box="[152,1434,1844,1869]" pageId="26" pageNumber="81">
<emphasis id="4CB2EA8CFFF6FFE29DBEF8BFFC9E466D" bold="true" box="[152,781,1844,1869]" pageId="26" pageNumber="81">
FIGURES 8083.
<taxonomicName id="B9C64D1DFFF6FFE29C46F8BFFC9B466D" authority="Ma &amp; Lin" authorityName="Ma &amp; Lin" authorityYear="2009" box="[352,776,1844,1869]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="81" phylum="Arthropoda" rank="species" species="robustochaetes">
<emphasis id="4CB2EA8CFFF6FFE29C46F8BFFD07466D" bold="true" box="[352,660,1844,1868]" italics="true" pageId="26" pageNumber="81">Cosmolaelaps robustochaetes</emphasis>
Ma &amp; Lin
</taxonomicName>
,
</emphasis>
adult female: 80—Leg I; 81—Leg II; 82—Leg III; 83—Leg IV.
</paragraph>
</caption>
<paragraph id="7E79369EFFF7FFE39DE1FEADFA874388" blockId="27.[151,1437,151,1041]" pageId="27" pageNumber="82">
<emphasis id="4CB2EA8CFFF7FFE39DE1FEADFECC4061" bold="true" box="[199,351,294,320]" pageId="27" pageNumber="82">Gnathosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFF7FFE39C49FEACFD804060" box="[367,531,295,321]" captionStart="FIGURES 6872" captionStartId="24.[152,270,1943,1967]" captionTargetBox="[187,1403,197,1908]" captionTargetId="figure-26@24.[172,1416,181,1919]" captionTargetPageId="24" captionText="FIGURES 6872. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 68—Dorsal idiosoma; 69—Ventral idiosoma; 70— Ventral gnathosoma; 71—Epistome; 72—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703951" httpUri="https://zenodo.org/record/14703951/files/figure.png" pageId="27" pageNumber="82">Figures 7072</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFF7FFE39F1EFEACFD104060" box="[568,643,295,321]" captionStart="FIGURES 7379" captionStartId="25.[152,270,1938,1962]" captionTargetBox="[187,1405,189,1910]" captionTargetId="figure-17@25.[179,1408,181,1915]" captionTargetPageId="25" captionText="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703953" httpUri="https://zenodo.org/record/14703953/files/figure.png" pageId="27" pageNumber="82">7679</figureCitation>
). Anterior margin of epistome convex, irregularly denticulate (
<figureCitation id="E6FD2A1BFFF7FFE39860FEACFF204044" captionStart="FIGURES 6872" captionStartId="24.[152,270,1943,1967]" captionTargetBox="[187,1403,197,1908]" captionTargetId="figure-26@24.[172,1416,181,1919]" captionTargetPageId="24" captionText="FIGURES 6872. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 68—Dorsal idiosoma; 69—Ventral idiosoma; 70— Ventral gnathosoma; 71—Epistome; 72—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703951" httpUri="https://zenodo.org/record/14703951/files/figure.png" pageId="27" pageNumber="82">Figures 71</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFF7FFE39DFDFEC0FF684044" box="[219,251,331,357]" captionStart="FIGURES 7379" captionStartId="25.[152,270,1938,1962]" captionTargetBox="[187,1405,189,1910]" captionTargetId="figure-17@25.[179,1408,181,1915]" captionTargetPageId="25" captionText="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703953" httpUri="https://zenodo.org/record/14703953/files/figure.png" pageId="27" pageNumber="82">78</figureCitation>
); hypostomal groove considerably widened, with six transverse rows of denticles, each row with 1733 tiny denticles, with smooth anterior and posterior transverse lines, groove wider anteriorly, progressively narrowing from anterior to posterior (
<figureCitation id="E6FD2A1BFFF7FFE39C98FE18FDA5408C" box="[446,566,403,429]" captionStart="FIGURES 6872" captionStartId="24.[152,270,1943,1967]" captionTargetBox="[187,1403,197,1908]" captionTargetId="figure-26@24.[172,1416,181,1919]" captionTargetPageId="24" captionText="FIGURES 6872. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 68—Dorsal idiosoma; 69—Ventral idiosoma; 70— Ventral gnathosoma; 71—Epistome; 72—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703951" httpUri="https://zenodo.org/record/14703951/files/figure.png" pageId="27" pageNumber="82">Figures 70</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFF7FFE39F71FE18FDE4408C" box="[599,631,403,429]" captionStart="FIGURES 7379" captionStartId="25.[152,270,1938,1962]" captionTargetBox="[187,1405,189,1910]" captionTargetId="figure-17@25.[179,1408,181,1915]" captionTargetPageId="25" captionText="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703953" httpUri="https://zenodo.org/record/14703953/files/figure.png" pageId="27" pageNumber="82">76</figureCitation>
). Hypostome with four pairs of smooth setae,
<emphasis id="4CB2EA8CFFF7FFE39953FE18FB02408C" box="[1141,1169,403,429]" italics="true" pageId="27" pageNumber="82">h3</emphasis>
(5370)&gt;
<emphasis id="4CB2EA8CFFF7FFE3982AFE18FAB4408C" box="[1292,1319,403,429]" italics="true" pageId="27" pageNumber="82">h1</emphasis>
(4753)&gt;
<emphasis id="4CB2EA8CFFF7FFE39DB1FE3CFF2240F1" box="[151,177,439,464]" italics="true" pageId="27" pageNumber="82">pc</emphasis>
(4145)&gt;
<emphasis id="4CB2EA8CFFF7FFE39C09FE3CFEDF40F0" box="[303,332,439,465]" italics="true" pageId="27" pageNumber="82">h2</emphasis>
(2127). Corniculi robust and horn-like, internal malae longer than corniculi, comprising a pair of pilose median projections, and 1115 lateral, longer, tentacle-like, smooth projections; labrum with pilose median surface, supralabral process indistinct; chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth, palpfemur with seta
<emphasis id="4CB2EA8CFFF7FFE39CFBFDA8FE60431C" box="[477,499,547,573]" italics="true" pageId="27" pageNumber="82">al</emphasis>
thickened; palpgenu with
<emphasis id="4CB2EA8CFFF7FFE39E3DFDA8FCAC431C" box="[795,831,547,573]" italics="true" pageId="27" pageNumber="82">al1</emphasis>
and
<emphasis id="4CB2EA8CFFF7FFE39E53FDA8FC0A431C" box="[885,921,547,573]" italics="true" pageId="27" pageNumber="82">al2</emphasis>
stout, blunt, and thickened. Palptarsal apotele two-tined (
<figureCitation id="E6FD2A1BFFF7FFE39C36FDCCFE134340" box="[272,384,583,609]" captionStart="FIGURES 7379" captionStartId="25.[152,270,1938,1962]" captionTargetBox="[187,1405,189,1910]" captionTargetId="figure-17@25.[179,1408,181,1915]" captionTargetPageId="25" captionText="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703953" httpUri="https://zenodo.org/record/14703953/files/figure.png" pageId="27" pageNumber="82">Figure 77</figureCitation>
); fixed digit of chelicera with an offset distal tooth (gabelzhan), followed by 910 various sized teeth, a setaceous pilus dentilis, dorsal cheliceral seta small and prostrate, arthrodial membrane with a rounded flap and normal filaments, dorsal and lateral lyrifissures distinct, movable digit bidentate (
<figureCitation id="E6FD2A1BFFF7FFE39970FD04FB574388" box="[1110,1220,655,681]" captionStart="FIGURES 6872" captionStartId="24.[152,270,1943,1967]" captionTargetBox="[187,1403,197,1908]" captionTargetId="figure-26@24.[172,1416,181,1919]" captionTargetPageId="24" captionText="FIGURES 6872. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 68—Dorsal idiosoma; 69—Ventral idiosoma; 70— Ventral gnathosoma; 71—Epistome; 72—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703951" httpUri="https://zenodo.org/record/14703951/files/figure.png" pageId="27" pageNumber="82">Figure 72</figureCitation>
&amp;
<figureCitation id="E6FD2A1BFFF7FFE399CEFD04FA9B4388" box="[1256,1288,655,681]" captionStart="FIGURES 7379" captionStartId="25.[152,270,1938,1962]" captionTargetBox="[187,1405,189,1910]" captionTargetId="figure-17@25.[179,1408,181,1915]" captionTargetPageId="25" captionText="FIGURES 7379. DIC micrographs of Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 73—Dorsal idiosoma; 74— Ventral idiosoma; 75—Some dorsal setae enlarged; 76—Ventral gnathosoma; 77—palp digit; 78—Epistome; 79—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703953" httpUri="https://zenodo.org/record/14703953/files/figure.png" pageId="27" pageNumber="82">79</figureCitation>
).
</paragraph>
<paragraph id="7E79369EFFF7FFE39DE1FD39FCEF43EC" blockId="27.[151,1437,151,1041]" box="[199,892,690,717]" pageId="27" pageNumber="82">
<emphasis id="4CB2EA8CFFF7FFE39DE1FD39FE7843EC" bold="true" box="[199,491,690,717]" pageId="27" pageNumber="82">Insemination structures.</emphasis>
Not seen, apparently unsclerotised.
</paragraph>
<paragraph id="7E79369EFFF7FFE39DE1FD5CFC3F4530" blockId="27.[151,1437,151,1041]" pageId="27" pageNumber="82">
<emphasis id="4CB2EA8CFFF7FFE39DE1FD5CFF6C43D0" bold="true" box="[199,255,727,753]" pageId="27" pageNumber="82">Legs</emphasis>
(
<figureCitation id="E6FD2A1BFFF7FFE39C35FD5CFE2C43D0" box="[275,447,727,753]" captionStart="FIGURES 8083" captionStartId="26.[152,270,1844,1868]" captionTargetBox="[164,1430,197,1805]" captionTargetId="figure-26@26.[151,1435,181,1821]" captionTargetPageId="26" captionText="FIGURES 8083. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 80—Leg I; 81—Leg II; 82—Leg III; 83—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703955" httpUri="https://zenodo.org/record/14703955/files/figure.png" pageId="27" pageNumber="82">Figures 8083</figureCitation>
). Leg II (471533) and III (427501) short, I (595667) and IV (635727) longer. Chaetotaxy normal for free-living
<taxonomicName id="B9C64D1DFFF7FFE39F0EFD70FD344234" box="[552,679,763,789]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="27" pageNumber="82" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
: Leg I (
<figureCitation id="E6FD2A1BFFF7FFE39E21FD70FCEE4234" box="[775,893,763,789]" captionStart="FIGURES 8083" captionStartId="26.[152,270,1844,1868]" captionTargetBox="[164,1430,197,1805]" captionTargetId="figure-26@26.[151,1435,181,1821]" captionTargetPageId="26" captionText="FIGURES 8083. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 80—Leg I; 81—Leg II; 82—Leg III; 83—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703955" httpUri="https://zenodo.org/record/14703955/files/figure.png" pageId="27" pageNumber="82">Figure 80</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/2, 0/1-1 (
<emphasis id="4CB2EA8CFFF7FFE398A0FD70FA0F4234" box="[1414,1436,763,789]" italics="true" pageId="27" pageNumber="82">al</emphasis>
spatulate), femur 2-2/1, 3/3-2 (
<emphasis id="4CB2EA8CFFF7FFE39CDAFC94FDB54218" box="[508,550,799,825]" italics="true" pageId="27" pageNumber="82">pd2</emphasis>
and
<emphasis id="4CB2EA8CFFF7FFE39F45FC94FD1F4218" box="[611,652,799,825]" italics="true" pageId="27" pageNumber="82">pd3</emphasis>
spatulate), genu 2-3/2, 3/1-2, tibia 2-3/2, 3/1-2. Leg II (
<figureCitation id="E6FD2A1BFFF7FFE3983CFC94FA1D4218" box="[1306,1422,799,825]" captionStart="FIGURES 8083" captionStartId="26.[152,270,1844,1868]" captionTargetBox="[164,1430,197,1805]" captionTargetId="figure-26@26.[151,1435,181,1821]" captionTargetPageId="26" captionText="FIGURES 8083. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 80—Leg I; 81—Leg II; 82—Leg III; 83—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703955" httpUri="https://zenodo.org/record/14703955/files/figure.png" pageId="27" pageNumber="82">Figure 81</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/1-1, femur 2-3/1, 2/2-1 (
<emphasis id="4CB2EA8CFFF7FFE39E4EFCC8FC47427C" box="[872,980,835,861]" italics="true" pageId="27" pageNumber="82">ad1, pd1,</emphasis>
and
<emphasis id="4CB2EA8CFFF7FFE39937FCC8FBA9427C" box="[1041,1082,835,861]" italics="true" pageId="27" pageNumber="82">pd2</emphasis>
spatulate,
<emphasis id="4CB2EA8CFFF7FFE3999FFCC8FB40427D" box="[1209,1235,835,860]" italics="true" pageId="27" pageNumber="82">av</emphasis>
spine-like), genu 2-3/1, 2/1-2, tibia 2-2/1, 2/1-2. Leg III (
<figureCitation id="E6FD2A1BFFF7FFE39F78FCECFD4242A0" box="[606,721,871,897]" captionStart="FIGURES 8083" captionStartId="26.[152,270,1844,1868]" captionTargetBox="[164,1430,197,1805]" captionTargetId="figure-26@26.[151,1435,181,1821]" captionTargetPageId="26" captionText="FIGURES 8083. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 80—Leg I; 81—Leg II; 82—Leg III; 83—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703955" httpUri="https://zenodo.org/record/14703955/files/figure.png" pageId="27" pageNumber="82">Figure 82</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/1-1, femur 1-2/1, 1/0-1 (
<emphasis id="4CB2EA8CFFF7FFE39D87FC00FF594284" box="[161,202,907,933]" italics="true" pageId="27" pageNumber="82">ad1</emphasis>
spatulate), genu 2-2/1, 2/1-1, tibia: 2-1/1, 2/1-1. Leg IV (
<figureCitation id="E6FD2A1BFFF7FFE39E6EFC00FC2A4284" box="[840,953,907,933]" captionStart="FIGURES 8083" captionStartId="26.[152,270,1844,1868]" captionTargetBox="[164,1430,197,1805]" captionTargetId="figure-26@26.[151,1435,181,1821]" captionTargetPageId="26" captionText="FIGURES 8083. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 80—Leg I; 81—Leg II; 82—Leg III; 83—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703955" httpUri="https://zenodo.org/record/14703955/files/figure.png" pageId="27" pageNumber="82">Figure 83</figureCitation>
): coxa 0-0/1, 0/0-0, trochanter 1-1/1, 0/1-1, femur 1-2/1, 1/0-1 (
<emphasis id="4CB2EA8CFFF7FFE39C53FC24FE4442E8" box="[373,471,943,969]" italics="true" pageId="27" pageNumber="82">ad1, ad2</emphasis>
spatulate), genu 2-2/1, 3/0-1, tibia 2-1/1, 3/1-2. Tarsi IIIV with 18 setae (3-3/2, 3/2-3 +
<emphasis id="4CB2EA8CFFF7FFE39D89FC58FF6C42CC" box="[175,255,979,1005]" italics="true" pageId="27" pageNumber="82">mv, md</emphasis>
); with some ventral and lateral setae thickened and spatulate, see
<figureCitation id="E6FD2A1BFFF7FFE39ED6FC58FB0542CC" box="[1008,1174,979,1005]" captionStart="FIGURES 8083" captionStartId="26.[152,270,1844,1868]" captionTargetBox="[164,1430,197,1805]" captionTargetId="figure-26@26.[151,1435,181,1821]" captionTargetPageId="26" captionText="FIGURES 8083. Cosmolaelaps robustochaetes Ma &amp; Lin, adult female: 80—Leg I; 81—Leg II; 82—Leg III; 83—Leg IV." figureDoi="http://doi.org/10.5281/zenodo.14703955" httpUri="https://zenodo.org/record/14703955/files/figure.png" pageId="27" pageNumber="82">Figures 8183</figureCitation>
. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="7E79369EFFF7FFE39DB1FBE9FE5B455D" blockId="27.[151,456,1122,1185]" box="[151,456,1122,1148]" pageId="27" pageNumber="82">
<emphasis id="4CB2EA8CFFF7FFE39DB1FBE9FE5B455D" bold="true" box="[151,456,1122,1148]" pageId="27" pageNumber="82">Description (adult male)</emphasis>
</paragraph>
<paragraph id="7E79369EFFF7FFE39DB1FB0CFEF34580" blockId="27.[151,456,1122,1185]" box="[151,352,1158,1185]" pageId="27" pageNumber="82">
(
<figureCitation id="E6FD2A1BFFF7FFE39DB9FB0CFECB4580" box="[159,344,1158,1185]" captionStart="FIGURES 8485" captionStartId="28.[152,270,1456,1480]" captionTargetBox="[168,1419,198,1415]" captionTargetId="figure-26@28.[151,1436,181,1432]" captionTargetPageId="28" captionText="FIGURES 8485. Cosmolaelaps robustochaetes Ma &amp; Lin, adult male: 84—Ventral idiosoma; 85—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703957" httpUri="https://zenodo.org/record/14703957/files/figure.png" pageId="27" pageNumber="82">Figures 84 &amp; 85</figureCitation>
)
</paragraph>
<paragraph id="7E79369EFFF7FFE39DB1FB40FE4945C2" blockId="27.[151,474,1227,1251]" box="[151,474,1227,1251]" pageId="27" pageNumber="82">
<specimenCount id="68C0FD17FFF7FFE39DB1FB40FEA645C2" box="[151,309,1227,1251]" count="5" pageId="27" pageNumber="82" type="generic">Five specimens</specimenCount>
were measured.
</paragraph>
<paragraph id="7E79369EFFF7FFE39DB1FA9AFAD2440A" blockId="27.[151,1437,1297,1792]" box="[151,1345,1297,1324]" pageId="27" pageNumber="82">
<emphasis id="4CB2EA8CFFF7FFE39DB1FA9AFEF3440A" bold="true" box="[151,352,1297,1323]" pageId="27" pageNumber="82">Dorsal idiosoma.</emphasis>
Dorsal shield 508535 long, 347372 wide; ornamentation and chaetotaxy as in female.
</paragraph>
<paragraph id="7E79369EFFF7FFE39DE1FABEFD17476A" blockId="27.[151,1437,1297,1792]" pageId="27" pageNumber="82">
<emphasis id="4CB2EA8CFFF7FFE39DE1FABEFE02446E" bold="true" box="[199,401,1333,1359]" pageId="27" pageNumber="82">Ventral idiosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFF7FFE39CB8FABEFD9D446E" box="[414,526,1333,1359]" captionStart="FIGURES 8485" captionStartId="28.[152,270,1456,1480]" captionTargetBox="[168,1419,198,1415]" captionTargetId="figure-26@28.[151,1436,181,1432]" captionTargetPageId="28" captionText="FIGURES 8485. Cosmolaelaps robustochaetes Ma &amp; Lin, adult male: 84—Ventral idiosoma; 85—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703957" httpUri="https://zenodo.org/record/14703957/files/figure.png" pageId="27" pageNumber="82">Figure 84</figureCitation>
). Sternal, genital, endopodal, ventral and anal shields fused into holoventral shield, 391409 long from anterior to posterior margins of shield, 95107 wide at level of
<emphasis id="4CB2EA8CFFF7FFE39975FAD2FBE04452" box="[1107,1139,1369,1395]" italics="true" pageId="27" pageNumber="82">st2</emphasis>
, 102106 at
<emphasis id="4CB2EA8CFFF7FFE3982DFAD2FABF4452" box="[1291,1324,1369,1395]" italics="true" pageId="27" pageNumber="82">st3</emphasis>
level and 249268 at broadest point, posteriorly to coxae IV; shield reticulate throughout (ornamentation only partly drawn on
<figureCitation id="E6FD2A1BFFF7FFE39D9CFA2AFEBB449A" box="[186,296,1441,1467]" captionStart="FIGURES 8485" captionStartId="28.[152,270,1456,1480]" captionTargetBox="[168,1419,198,1415]" captionTargetId="figure-26@28.[151,1436,181,1432]" captionTargetPageId="28" captionText="FIGURES 8485. Cosmolaelaps robustochaetes Ma &amp; Lin, adult male: 84—Ventral idiosoma; 85—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703957" httpUri="https://zenodo.org/record/14703957/files/figure.png" pageId="27" pageNumber="82">Figure 84</figureCitation>
), with five pairs of simple sternal setae (
<emphasis id="4CB2EA8CFFF7FFE39FCDFA2AFCA9449D" box="[747,826,1441,1468]" italics="true" pageId="27" pageNumber="82">st1st5</emphasis>
), and five pairs of smooth ventral setae (
<emphasis id="4CB2EA8CFFF7FFE39824FA2AFACD449A" box="[1282,1374,1441,1468]" italics="true" pageId="27" pageNumber="82">Jv1Jv3</emphasis>
,
<emphasis id="4CB2EA8CFFF7FFE3984DFA2AFA06449A" box="[1387,1429,1441,1467]" italics="true" pageId="27" pageNumber="82">Zv1</emphasis>
, and
<emphasis id="4CB2EA8CFFF7FFE39DE1FA4EFF6244FE" box="[199,241,1477,1503]" italics="true" pageId="27" pageNumber="82">Zv2</emphasis>
), plus three smooth circum-anal setae, para-anal setae and post-anal seta subequal in length, with six pairs of poroids; cribrum with 34 irregular rows of spicules; metapodal platelets completely fused to holoventral shield. Soft ophisthogastric and lateral cuticle with 12 pairs of spatulate setae. Peritremes, peritrematal shields, and other ventral structures similar to those in female.
</paragraph>
<paragraph id="7E79369EFFF7FFE39DE1F9DEFF7847FA" blockId="27.[151,1437,1297,1792]" pageId="27" pageNumber="82">
<emphasis id="4CB2EA8CFFF7FFE39DE1F9DEFECC474E" bold="true" box="[199,351,1621,1647]" pageId="27" pageNumber="82">Gnathosoma</emphasis>
(
<figureCitation id="E6FD2A1BFFF7FFE39C57F9DEFE774751" box="[369,484,1621,1648]" captionStart="FIGURES 8485" captionStartId="28.[152,270,1456,1480]" captionTargetBox="[168,1419,198,1415]" captionTargetId="figure-26@28.[151,1436,181,1432]" captionTargetPageId="28" captionText="FIGURES 8485. Cosmolaelaps robustochaetes Ma &amp; Lin, adult male: 84—Ventral idiosoma; 85—Chelicera." figureDoi="http://doi.org/10.5281/zenodo.14703957" httpUri="https://zenodo.org/record/14703957/files/figure.png" pageId="27" pageNumber="82">Figure 85</figureCitation>
). Fixed digit of chelicera with 11 teeth of various sizes in addition to apical hook. Movable digit of chelicera monodentate, spermatodactyl digitform, gently curved, with rounded tip, free portion of spermatodactyl shorter than movable digit (3438). Other gnathosomal and cheliceral structures similar to those in female.
</paragraph>
<paragraph id="7E79369EFFF7FFE39DE1F96EFDB047DE" blockId="27.[151,1437,1297,1792]" box="[199,547,1765,1792]" pageId="27" pageNumber="82">
<emphasis id="4CB2EA8CFFF7FFE39DE1F96EFE9547DE" bold="true" box="[199,262,1765,1791]" pageId="27" pageNumber="82">Legs.</emphasis>
Chaetotaxy as in female.
</paragraph>
</subSubSection>
</treatment>
</document>