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<mods:title id="774D3B172D086183E8812085F1AF98F5">Rhinonycteridae</mods:title>
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<mods:namePart id="0D3F70787AE24A420FF6E90CCE95FFEC">Don E. Wilson</mods:namePart>
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<mods:namePart id="F23A9D42AC1C28AE7790C0C87A83EF2D">Don E. Wilson</mods:namePart>
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<mods:namePart id="8B851CDD64F6A8427859C9F4BC08C531">Russell A. Mittermeier</mods:namePart>
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<treatment id="0383245F2225977B8BD5F631FD78FE56" ID-DOI="http://doi.org/10.5281/zenodo.6611851" ID-GBIF-Taxon="195730784" ID-Zenodo-Dep="6611851" LSID="urn:lsid:plazi:treatment:0383245F2225977B8BD5F631FD78FE56" httpUri="http://treatment.plazi.org/id/0383245F2225977B8BD5F631FD78FE56" lastPageId="7" lastPageNumber="209" pageId="6" pageNumber="208">
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<subSubSection id="C330C6C22225977A8BD5F631FA76FC8E" box="[1446,1479,824,862]" pageId="6" pageNumber="208" type="multiple">
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<paragraph id="8B9595492225977A8BD5F631FA76FC8E" blockId="6.[1444,2554,823,940]" box="[1446,1479,824,862]" pageId="6" pageNumber="208">
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<heading id="D0DD22252225977A8BD5F631FA76FC8E" box="[1446,1479,824,862]" pageId="6" pageNumber="208">
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@ -53,14 +53,14 @@
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<subSubSection id="C330C6C22225977A8BA4F630F868FC8E" box="[1495,2009,825,862]" pageId="6" pageNumber="208" type="vernacular_names">
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<paragraph id="8B9595492225977A8BA4F630F868FC8E" blockId="6.[1444,2554,823,940]" box="[1495,2009,825,862]" pageId="6" pageNumber="208">
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<heading id="D0DD22252225977A8BA4F630F868FC8E" box="[1495,2009,825,862]" pageId="6" pageNumber="208">
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||||
<vernacularName id="0529E5672225977A8BA4F630F868FC8E" box="[1495,2009,825,862]" pageId="6" pageNumber="208">Grandidiefs Trident Bat</vernacularName>
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<vernacularName id="0529E5672225977A8BA4F630F868FC8E" baseAuthorityName="G. Grandidier" baseAuthorityYear="1912" box="[1495,2009,825,862]" class="Magnoliopsida" family="Fabaceae" genus="Paratriaenops" kingdom="Plantae" language="eng" order="Fabales" pageId="6" pageNumber="208" phylum="Tracheophyta" rank="species" species="auritus">Grandidiefs Trident Bat</vernacularName>
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</heading>
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</paragraph>
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<subSubSection id="C330C6C22225977A867AF633F616FC8D" box="[2057,2471,826,874]" pageId="6" pageNumber="208" type="nomenclature">
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<paragraph id="8B9595492225977A867AF633F616FC8D" blockId="6.[1444,2554,823,940]" box="[2057,2471,826,874]" pageId="6" pageNumber="208">
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||||
<heading id="D0DD22252225977A867AF633F616FC8D" box="[2057,2471,826,874]" pageId="6" pageNumber="208">
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||||
<taxonomicName id="4C2AEECA2225977A867AF633F616FC8D" baseAuthorityName="G. Grandidier" baseAuthorityYear="1912" box="[2057,2471,826,874]" class="Magnoliopsida" family="Fabaceae" genus="Paratriaenops" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="208" phylum="Tracheophyta" rank="species" species="auritus">
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||||
<taxonomicName id="4C2AEECA2225977A867AF633F616FC8D" ID-CoL="4DGHH" baseAuthorityName="G. Grandidier" baseAuthorityYear="1912" box="[2057,2471,826,874]" class="Mammalia" family="Rhinonycteridae" genus="Paratriaenops" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="208" phylum="Chordata" rank="species" species="auritus">
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<emphasis id="B95E495B2225977A867AF633F616FC8D" box="[2057,2471,826,874]" italics="true" pageId="6" pageNumber="208">Paratriaenops auritus</emphasis>
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</taxonomicName>
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</heading>
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@ -70,17 +70,26 @@
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<paragraph id="8B9595492225977A8BD7F678F648FC54" blockId="6.[1444,2554,823,940]" box="[1444,2553,881,904]" pageId="6" pageNumber="208">
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<heading id="D0DD22252225977A8BD7F678F648FC54" box="[1444,2553,881,904]" pageId="6" pageNumber="208">
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<emphasis id="B95E495B2225977A8BD7F678FA5AFC58" bold="true" box="[1444,1515,881,904]" pageId="6" pageNumber="208">French:</emphasis>
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<vernacularName id="0529E5672225977A8B85F67BF955FC54" box="[1526,1764,882,904]" pageId="6" pageNumber="208">Tr aen ps de Grand d er</vernacularName>
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<emphasis id="B95E495B2225977A8884F678F8FCFC58" bold="true" box="[1783,1869,881,904]" pageId="6" pageNumber="208">German:</emphasis>
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<vernacularName id="0529E5672225977A8928F67BF7CBFC55" box="[1883,2170,882,901]" pageId="6" pageNumber="208">Grand drer-Dre zahnblannase</vernacularName>
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<emphasis id="B95E495B2225977A86FDF678F754FC58" bold="true" box="[2190,2277,881,904]" pageId="6" pageNumber="208">Spanish:</emphasis>
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<vernacularName id="0529E5672225977A8687F67BF648FC54" box="[2292,2553,881,901]" pageId="6" pageNumber="208">R non terio de Grındıdıer</vernacularName>
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<vernacularName id="0529E5672225977A8B85F67BF8F6FC58" baseAuthorityName="G. Grandidier" baseAuthorityYear="1912" box="[1526,1863,881,904]" class="Magnoliopsida" family="Fabaceae" genus="Paratriaenops" kingdom="Plantae" language="fra" order="Fabales" pageId="6" pageNumber="208" phylum="Tracheophyta" rank="species" species="auritus">
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Tr aen ps de Grand d er
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<emphasis id="B95E495B2225977A8884F678F8F6FC58" bold="true" box="[1783,1863,881,904]" pageId="6" pageNumber="208">German</emphasis>
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</vernacularName>
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:
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<vernacularName id="0529E5672225977A8928F67BF76EFC58" baseAuthorityName="G. Grandidier" baseAuthorityYear="1912" box="[1883,2271,881,904]" class="Magnoliopsida" family="Fabaceae" genus="Paratriaenops" kingdom="Plantae" language="deu" order="Fabales" pageId="6" pageNumber="208" phylum="Tracheophyta" rank="species" species="auritus">
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Grand drer-Dre zahnblannase
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<emphasis id="B95E495B2225977A86FDF678F76EFC58" bold="true" box="[2190,2271,881,904]" pageId="6" pageNumber="208">Spanish</emphasis>
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</vernacularName>
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:
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<vernacularName id="0529E5672225977A8687F67BF648FC54" baseAuthorityName="G. Grandidier" baseAuthorityYear="1912" box="[2292,2553,881,901]" class="Magnoliopsida" family="Fabaceae" genus="Paratriaenops" kingdom="Plantae" language="eng" order="Fabales" pageId="6" pageNumber="208" phylum="Tracheophyta" rank="species" species="auritus">
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R
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<vernacularName id="0529E5672225977A877BF67EF648FC54" baseAuthorityName="G. Grandidier" baseAuthorityYear="1912" box="[2312,2553,881,901]" class="Magnoliopsida" family="Fabaceae" genus="Paratriaenops" kingdom="Plantae" language="eng" order="Fabales" pageId="6" pageNumber="208" phylum="Tracheophyta" rank="species" species="auritus">non terio de Grındıdıer</vernacularName>
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</vernacularName>
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</heading>
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<paragraph id="8B9595492225977A8BD7F690F8EDFC7C" blockId="6.[1444,2554,823,940]" box="[1444,1884,921,940]" pageId="6" pageNumber="208">
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<heading id="D0DD22252225977A8BD7F690F8EDFC7C" box="[1444,1884,921,940]" pageId="6" pageNumber="208">
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||||
<emphasis id="B95E495B2225977A8BD7F690F926FC7C" bold="true" box="[1444,1687,921,940]" pageId="6" pageNumber="208">Other common names:</emphasis>
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<vernacularName id="0529E5672225977A88D6F690F8EDFC7C" box="[1701,1884,921,940]" pageId="6" pageNumber="208">Golden Trident Bat</vernacularName>
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<vernacularName id="0529E5672225977A88D6F690F8EDFC7C" baseAuthorityName="G. Grandidier" baseAuthorityYear="1912" box="[1701,1884,921,940]" class="Magnoliopsida" family="Fabaceae" genus="Paratriaenops" kingdom="Plantae" language="eng" order="Fabales" pageId="6" pageNumber="208" phylum="Tracheophyta" rank="species" species="auritus">Golden Trident Bat</vernacularName>
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</heading>
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@ -1,47 +1,47 @@
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<mods:title id="09BFA5E34BF85DAAD3F658E22B7AE9B0">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
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<mods:namePart id="D9339C0A239807226096B91EDAD07774">C, Lees David</mods:namePart>
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<treatment id="038747324C23C6001EB72C94FE4726AB" ID-DOI="http://doi.org/10.5281/zenodo.6086460" ID-GBIF-Taxon="120817148" ID-Zenodo-Dep="6086460" LSID="urn:lsid:plazi:treatment:038747324C23C6001EB72C94FE4726AB" httpUri="http://treatment.plazi.org/id/038747324C23C6001EB72C94FE4726AB" lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86">
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<treatment id="038747324C23C6001EB72C94FE4726AB" ID-DOI="http://doi.org/10.5281/zenodo.6086460" ID-GBIF-Taxon="120817148" ID-Zenodo-Dep="6086460" LSID="urn:lsid:plazi:treatment:038747324C23C6001EB72C94FE4726AB" httpUri="http://treatment.plazi.org/id/038747324C23C6001EB72C94FE4726AB" lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
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<subSubSection id="C334A5AF4C23C6021EB72C94FBC12081" pageId="85" pageNumber="86" type="nomenclature">
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<paragraph id="8B91F6244C23C6021EB72C94FDC220A0" blockId="85.[151,1042,441,501]" box="[151,529,441,468]" pageId="85" pageNumber="86">
|
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<heading id="D0D941484C23C6021EB72C94FDC220A0" bold="true" box="[151,529,441,468]" fontSize="11" level="1" pageId="85" pageNumber="86" reason="1">
|
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<emphasis id="B95A2A364C23C6021EB72C94FDC220A0" bold="true" box="[151,529,441,468]" pageId="85" pageNumber="86">
|
||||
<taxonomicName id="4C2E8DA74C23C6021EB72C94FE7520A0" authority="Lees" authorityName="Lees" box="[151,422,441,468]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="borgo" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C23C6021EB72C94FE7520A0" authority="Lees" authorityName="Lees" box="[151,422,441,468]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="borgo" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C23C6021EB72C94FEB620A7" bold="true" box="[151,357,441,467]" italics="true" pageId="85" pageNumber="86">Heteropsis borgo</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -77,7 +77,7 @@ material., Deposition
|
|||
:
|
||||
</emphasis>
|
||||
♂ (
|
||||
<figureCitation id="1315EAA14C23C6021D622F6FFC4F232F" box="[834,924,578,603]" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="85" pageNumber="86">Fig. 27</figureCitation>
|
||||
<figureCitation id="1315EAA14C23C6021D622F6FFC4F232F" box="[834,924,578,603]" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264624" httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="85" pageNumber="86">Fig. 27</figureCitation>
|
||||
A), Anjanaharibe Sud, summit Anjavidibe,
|
||||
<geoCoordinate id="EE1A90E34C23C6021EB72F48FEDC230A" box="[151,271,613,638]" direction="south" orientation="latitude" pageId="85" pageNumber="86" precision="5" value="-14.7396">14.7396°S</geoCoordinate>
|
||||
,
|
||||
|
|
@ -230,11 +230,11 @@ Anjanaharibe Sud, summit Anjavidibe,
|
|||
<subSubSection id="C334A5AF4C23C6021EE729C1FCF22438" pageId="85" pageNumber="86" type="diagnosis">
|
||||
<paragraph id="8B91F6244C23C6021EE729C1FCF22438" blockId="85.[151,1437,542,2030]" pageId="85" pageNumber="86">
|
||||
<emphasis id="B95A2A364C23C6021EE729C1FE922471" bold="true" box="[199,321,1260,1285]" pageId="85" pageNumber="86">Diagnosis.</emphasis>
|
||||
<taxonomicName id="4C2E8DA74C23C6021F6B29C3FDD12471" box="[331,514,1260,1285]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<taxonomicName id="4C2E8DA74C23C6021F6B29C3FDD12471" box="[331,514,1260,1285]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<emphasis id="B95A2A364C23C6021F6B29C3FDD12471" box="[331,514,1260,1285]" italics="true" pageId="85" pageNumber="86">Ht. angulifascia</emphasis>
|
||||
</taxonomicName>
|
||||
is particularly similar, but there are no other closely comparable species. HWV Mb, however, much less indented that in
|
||||
<taxonomicName id="4C2E8DA74C23C6021C4F283CFCF6245D" box="[623,805,1296,1321]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<taxonomicName id="4C2E8DA74C23C6021C4F283CFCF6245D" box="[623,805,1296,1321]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<emphasis id="B95A2A364C23C6021C4F283CFCF6245D" box="[623,805,1296,1321]" italics="true" pageId="85" pageNumber="86">Ht. angulifascia</emphasis>
|
||||
</taxonomicName>
|
||||
, scribing a lobe outward between M2 and M3, running for only a short rather than a long distance just below M2.
|
||||
|
|
@ -247,22 +247,22 @@ is particularly similar, but there are no other closely comparable species. HWV
|
|||
<emphasis id="B95A2A364C20C6011D352DBAFC5921C4" bold="true" box="[789,906,151,176]" pageId="86" pageNumber="87">Variation.</emphasis>
|
||||
♂♂ very similar to each other, without marked variation in eyespot size between the two collection periods (third week of November, late dry season, and second week of February, wet season). The HWV Mb varies in the angularity with which it is kinked out around space- M2–M3, but this ‘kinking’ is not as extreme as in
|
||||
<emphasis id="B95A2A364C20C6011CE62C2EFC51206F" box="[710,898,258,283]" italics="true" pageId="86" pageNumber="87">
|
||||
<taxonomicName id="4C2E8DA74C20C6011CE62C2EFCAD206F" box="[710,894,258,283]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="angulifascia">Ht. angulifascia</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C20C6011CE62C2EFCAD206F" box="[710,894,258,283]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="angulifascia">Ht. angulifascia</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
♀ unknown, but likely to be similar to the ♂, as for the closely related
|
||||
<taxonomicName id="4C2E8DA74C20C6011FB12C0AFD95204A" box="[401,582,293,318]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<taxonomicName id="4C2E8DA74C20C6011FB12C0AFD95204A" box="[401,582,293,318]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<emphasis id="B95A2A364C20C6011FB12C0AFD95204A" box="[401,582,293,318]" italics="true" pageId="86" pageNumber="87">Ht. angulifascia</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C20C6011EB72A19FCCA26D2" httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="86" pageNumber="87" targetBox="[300,1293,370,1814]" targetPageId="86">
|
||||
<caption id="DF51A6AC4C20C6011EB72A19FCCA26D2" ID-DOI="http://doi.org/10.5281/zenodo.264624" httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="86" pageNumber="87" targetBox="[300,1293,370,1814]" targetPageId="86">
|
||||
<paragraph id="8B91F6244C20C6011EB72A19FCCA26D2" blockId="86.[151,1436,1844,1958]" pageId="86" pageNumber="87">
|
||||
<emphasis id="B95A2A364C20C6011EB72A19FEFA263D" bold="true" box="[151,297,1844,1866]" pageId="86" pageNumber="87">FIGURE 27.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
<emphasis id="B95A2A364C20C6011C952A19FD14263D" bold="true" box="[693,711,1844,1865]" pageId="86" pageNumber="87">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C20C6011D762A18FB83263E" authority="Lees" authorityName="Lees" box="[854,1104,1844,1866]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="borgo" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C20C6011D762A18FB83263E" authority="Lees" authorityName="Lees" box="[854,1104,1844,1866]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="borgo" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C20C6011D762A18FBDC263E" box="[854,1039,1844,1866]" italics="true" pageId="86" pageNumber="87">Heteropsis borgo</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -289,7 +289,7 @@ Butler, 1879
|
|||
(‘Antananarivo’, BMNH(E) 674769). Scale bar: 30 mm.
|
||||
<emphasis id="B95A2A364C20C6011AA62A5CFB4B26F2" bold="true" box="[1158,1176,1905,1926]" pageId="86" pageNumber="87">D</emphasis>
|
||||
: ♂ genitalia of
|
||||
<taxonomicName id="4C2E8DA74C20C6011B1A2A5FFA4F26F3" box="[1338,1436,1905,1927]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="borgo">
|
||||
<taxonomicName id="4C2E8DA74C20C6011B1A2A5FFA4F26F3" box="[1338,1436,1905,1927]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="borgo">
|
||||
<emphasis id="B95A2A364C20C6011B1A2A5FFA4F26F3" box="[1338,1436,1905,1927]" italics="true" pageId="86" pageNumber="87">Ht. borgo</emphasis>
|
||||
</taxonomicName>
|
||||
(Anjanaharibe Sud; CK712, 284DL), DV, SV. Scale bar: 3 mm.
|
||||
|
|
@ -316,7 +316,7 @@ range
|
|||
<paragraph id="8B91F6244C21C6001EE72CB9FD6F23DC" blockId="87.[151,1437,151,2015]" pageId="87" pageNumber="88">
|
||||
<emphasis id="B95A2A364C21C6001EE72CB9FE9F20D9" bold="true" box="[199,332,404,429]" pageId="87" pageNumber="88">♂ genitalia</emphasis>
|
||||
: 284DL, PT (
|
||||
<figureCitation id="1315EAA14C21C6001FD12CB9FD9420D8" box="[497,583,404,429]" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="87" pageNumber="88">Fig. 27</figureCitation>
|
||||
<figureCitation id="1315EAA14C21C6001FD12CB9FD9420D8" box="[497,583,404,429]" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264624" httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="87" pageNumber="88">Fig. 27</figureCitation>
|
||||
D): from LV, uncus slightly upbent before middle and recurved down to a sharply hooked point, without a distinct ‘head’ before tip (slightly inflated before tip from DV); tegumen shorter than uncus and constriction at fusion line with vinculum not as pronounced as in some other species. Gnathos flattened distinctly into an ear-like shape (as for, e.g.,
|
||||
<taxonomicName id="4C2E8DA74C21C6001CDA2F2CFC7C236C" box="[762,943,511,536]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<emphasis id="B95A2A364C21C6001CDA2F2CFC7C236C" box="[762,943,511,536]" italics="true" pageId="87" pageNumber="88">Ht. angulifascia</emphasis>
|
||||
|
|
@ -328,7 +328,7 @@ D): from LV, uncus slightly upbent before middle and recurved down to a sharply
|
|||
<paragraph id="8B91F6244C21C6001EE72F99FD4F2261" blockId="87.[151,1437,151,2015]" pageId="87" pageNumber="88">
|
||||
<emphasis id="B95A2A364C21C6001EE72F99FE9D23B9" bold="true" box="[199,334,692,717]" pageId="87" pageNumber="88">Etymology.</emphasis>
|
||||
A very dark and still rather mysterious species of mountain passes, in this case the one on the way to the summit of Anjanaharibe Sud, as in the
|
||||
<taxonomicName id="4C2E8DA74C21C6001CBE2FF5FD302384" box="[670,739,728,752]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="94" phylum="Arthropoda" rank="species" species="borgo">Borgó</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C21C6001CBE2FF5FD302384" box="[670,739,728,752]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="87" pageNumber="94" phylum="Arthropoda" rank="species" species="borgo">Borgó</taxonomicName>
|
||||
pass made famous in F.W. Murnau’s film classic ‘Nosferatu’ and its compelling remake by Werner Herzog.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -347,7 +347,7 @@ locality, only) in November, 2014 by myself and A. Allaoui. The HT ♂ in BMNH o
|
|||
<bibRefCitation id="EFBF8BD54C21C6001FFA2E82FDB622BC" author="Butler" box="[474,613,943,968]" pageId="87" pageNumber="88" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(‘Antananarivo’, Kingdon, BMNH(E) 674769) was examined (
|
||||
<figureCitation id="1315EAA14C21C6001B102E82FA5222BC" box="[1328,1409,943,968]" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="87" pageNumber="88">Fig. 27</figureCitation>
|
||||
<figureCitation id="1315EAA14C21C6001B102E82FA5222BC" box="[1328,1409,943,968]" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264624" httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="87" pageNumber="88">Fig. 27</figureCitation>
|
||||
C) as well as the ♂ and ♀ STs of its synonym
|
||||
<taxonomicName id="4C2E8DA74C21C6001C5A2EF9FC332298" authority="Mabille, 1880" authorityName="Mabille" authorityYear="1880" box="[634,992,980,1005]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="butleri">
|
||||
<emphasis id="B95A2A364C21C6001C5A2EF9FCEF2299" box="[634,828,980,1005]" italics="true" pageId="87" pageNumber="88">Mycalesis butleri</emphasis>
|
||||
|
|
@ -355,7 +355,7 @@ Mabille,
|
|||
<quantity id="4CD65BC14C21C6001D882EF9FC332298" box="[936,992,980,1004]" metricMagnitude="1" metricUnit="m" metricValue="4.7752" pageId="87" pageNumber="88" unit="in" value="1880.0">1880</quantity>
|
||||
</taxonomicName>
|
||||
in BMNH (LT ♂, here designated,
|
||||
<figureCitation id="1315EAA14C21C6001B4F2EF9FF662564" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="87" pageNumber="88">Fig. 27</figureCitation>
|
||||
<figureCitation id="1315EAA14C21C6001B4F2EF9FF662564" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264624" httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="87" pageNumber="88">Fig. 27</figureCitation>
|
||||
B, bearing data: “
|
||||
<typeStatus id="549548864C21C6001F542EDAFE362564" box="[372,485,1015,1040]" pageId="87" pageNumber="88" type="lectotype">Lectotype</typeStatus>
|
||||
/
|
||||
|
|
@ -383,7 +383,7 @@ Torres
|
|||
2001
|
||||
</bibRefCitation>
|
||||
), there was an about 2.15% divergence (8bp different in 415) for a single sample of
|
||||
<taxonomicName id="4C2E8DA74C21C6001D0429FCFC40259C" box="[804,915,1231,1256]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="borgo">
|
||||
<taxonomicName id="4C2E8DA74C21C6001D0429FCFC40259C" box="[804,915,1231,1256]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="borgo">
|
||||
<emphasis id="B95A2A364C21C6001D0429FCFC40259C" box="[804,915,1231,1256]" italics="true" pageId="87" pageNumber="88">Ht. borgo</emphasis>
|
||||
</taxonomicName>
|
||||
(as “
|
||||
|
|
@ -446,7 +446,7 @@ locality.
|
|||
<paragraph id="8B91F6244C21C6001EE72B52FA9827EC" blockId="87.[151,1437,151,2015]" box="[199,1355,1663,1688]" pageId="87" pageNumber="88">
|
||||
<emphasis id="B95A2A364C21C6001EE72B52FE8A27EC" bold="true" box="[199,345,1663,1688]" pageId="87" pageNumber="88">Distribution</emphasis>
|
||||
: endemic to rainforest at Anjanaharibe Sud, as far as is known (
|
||||
<figureCitation id="1315EAA14C21C6001A072B52FBA927EC" box="[1063,1146,1663,1688]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="87" pageNumber="88">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C21C6001A072B52FBA927EC" box="[1063,1146,1663,1688]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="87" pageNumber="88">Fig. 30</figureCitation>
|
||||
C, dark green dot).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C21C6001EE72B89FB772653" blockId="87.[151,1437,151,2015]" pageId="87" pageNumber="88">
|
||||
|
|
@ -473,7 +473,7 @@ downslope, and it is not known if the species occurs on the adjacent real summit
|
|||
<paragraph id="8B91F6244C21C6001EE72A7AFE4726AB" blockId="87.[151,1437,151,2015]" pageId="87" pageNumber="88">
|
||||
<emphasis id="B95A2A364C21C6001EE72A7AFE682604" bold="true" box="[199,443,1879,1904]" pageId="87" pageNumber="88">Referred specimens.</emphasis>
|
||||
♂ (
|
||||
<figureCitation id="1315EAA14C21C6001FCA2A7AFDEE2604" box="[490,573,1879,1904]" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="87" pageNumber="88">Fig. 27</figureCitation>
|
||||
<figureCitation id="1315EAA14C21C6001FCA2A7AFDEE2604" box="[490,573,1879,1904]" captionStart="FIGURE 27" captionStartId="86.[151,250,1844,1866]" captionTargetBox="[300,1293,370,1814]" captionTargetId="figure@86.[279,1306,354,1830]" captionTargetPageId="86" captionText="FIGURE 27. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis borgo Lees, sp. nov. (Anjanaharibe Sud; NHMUK 010289167). B: LT ♂ of Mycalesis butleri Mabille, 1880 (‘ Madgscr’, BMNH (E) 674771). C: HT ♂ of Pseudonympha angulifascia Butler, 1879 (‘ Antananarivo’, BMNH (E) 674769). Scale bar: 30 mm. D: ♂ genitalia of Ht. borgo (Anjanaharibe Sud; CK 712, 284 DL), DV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264624" httpUri="https://zenodo.org/record/264624/files/figure.png" pageId="87" pageNumber="88">Fig. 27</figureCitation>
|
||||
D, male genitalia, rest of voucher lost),
|
||||
<collectingCountry id="F339B6B44C21C6001A252A7AFB5F2604" box="[1029,1164,1879,1904]" name="Madagascar" pageId="87" pageNumber="88">Madagascar</collectingCountry>
|
||||
NE, Anjanaharibe Sud,
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<treatment id="038747324C24C6021EB72BF9FBA92005" ID-DOI="http://doi.org/10.5281/zenodo.6086458" ID-GBIF-Taxon="120817150" ID-Zenodo-Dep="6086458" LSID="urn:lsid:plazi:treatment:038747324C24C6021EB72BF9FBA92005" httpUri="http://treatment.plazi.org/id/038747324C24C6021EB72BF9FBA92005" lastPageId="85" lastPageNumber="85" pageId="82" pageNumber="83">
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<treatment id="038747324C24C6021EB72BF9FBA92005" ID-DOI="http://doi.org/10.5281/zenodo.6086458" ID-GBIF-Taxon="120817150" ID-Zenodo-Dep="6086458" LSID="urn:lsid:plazi:treatment:038747324C24C6021EB72BF9FBA92005" httpUri="http://treatment.plazi.org/id/038747324C24C6021EB72BF9FBA92005" lastPageId="85" lastPageNumber="85" pageId="82" pageNumber="83" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C24C6051EB72BF9FBF52664" pageId="82" pageNumber="83" type="nomenclature">
|
||||
<paragraph id="8B91F6244C24C6051EB72BF9FD99279A" blockId="82.[151,1062,1748,1808]" box="[151,586,1748,1774]" pageId="82" pageNumber="83">
|
||||
<heading id="D0D941484C24C6051EB72BF9FD99279A" bold="true" box="[151,586,1748,1774]" fontSize="11" level="1" pageId="82" pageNumber="83" reason="1">
|
||||
<emphasis id="B95A2A364C24C6051EB72BF9FD99279A" bold="true" box="[151,586,1748,1774]" pageId="82" pageNumber="83">
|
||||
<taxonomicName id="4C2E8DA74C24C6051EB72BF9FE33279A" authority="Lees" authorityName="Lees" box="[151,480,1748,1774]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="oberthueri" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C24C6051EB72BF9FE33279A" authority="Lees" authorityName="Lees" box="[151,480,1748,1774]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="oberthueri" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C24C6051EB72BF9FE4D279A" bold="true" box="[151,414,1748,1774]" italics="true" pageId="82" pageNumber="83">Heteropsis oberthueri</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -62,7 +62,7 @@ Lees
|
|||
] in Aduse-Poku
|
||||
<emphasis id="B95A2A364C24C6051B792A6DFA4F262C" box="[1369,1436,1855,1880]" italics="true" pageId="82" pageNumber="83">et al.,</emphasis>
|
||||
(2015,
|
||||
<figureCitation id="1315EAA14C24C6051EC62A4EFEF4260F" box="[230,295,1891,1916]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="82" pageNumber="83">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C24C6051EC62A4EFEF4260F" box="[230,295,1891,1916]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="82" pageNumber="83">Fig. 1</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -77,7 +77,7 @@ material. Deposition
|
|||
:
|
||||
</emphasis>
|
||||
♂ (
|
||||
<figureCitation id="1315EAA14C24C6051D072AA4FCAD26D6" box="[807,894,1929,1954]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="82" pageNumber="83">Fig. 24</figureCitation>
|
||||
<figureCitation id="1315EAA14C24C6051D072AA4FCAD26D6" box="[807,894,1929,1954]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264621" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="82" pageNumber="83">Fig. 24</figureCitation>
|
||||
D),
|
||||
<collectingCountry id="F339B6B44C24C6051D8C2AA4FBE026D6" box="[940,1075,1929,1954]" name="Madagascar" pageId="82" pageNumber="83">Madagascar</collectingCountry>
|
||||
E, Zahamena NW,
|
||||
|
|
@ -129,7 +129,7 @@ E, Ambavala camp, Zahamena,
|
|||
010289160 [
|
||||
<collectingCountry id="F339B6B44C25C6041A722D92FB5E21A3" box="[1106,1165,191,215]" name="Qatar" pageId="83" pageNumber="84">QTR</collectingCountry>
|
||||
barcode]; ♀ (
|
||||
<figureCitation id="1315EAA14C25C6041B0A2D93FAAF21A3" box="[1322,1404,190,215]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="83" pageNumber="84">Fig. 24</figureCitation>
|
||||
<figureCitation id="1315EAA14C25C6041B0A2D93FAAF21A3" box="[1322,1404,190,215]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264621" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="83" pageNumber="84">Fig. 24</figureCitation>
|
||||
E), E, Zahamena NW,
|
||||
<geoCoordinate id="EE1A90E34C25C6041F4F2DC9FE302189" box="[367,483,225,253]" direction="south" orientation="latitude" pageId="83" pageNumber="84" precision="5" value="-17.5376">17.5376o S</geoCoordinate>
|
||||
,
|
||||
|
|
@ -307,30 +307,30 @@ NW Zahamena,
|
|||
<paragraph id="8B91F6244C25C6041EE72EACFEEF24A2" blockId="83.[151,1437,151,2030]" pageId="83" pageNumber="84">
|
||||
<emphasis id="B95A2A364C25C6041EE72EACFE9222EE" bold="true" box="[199,321,897,922]" pageId="83" pageNumber="84">Diagnosis.</emphasis>
|
||||
The species is particularly similar to
|
||||
<taxonomicName id="4C2E8DA74C25C6041D002EAFFC4B22ED" box="[800,920,898,921]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C25C6041D002EAFFC4B22ED" box="[800,920,898,921]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C25C6041D002EAFFC4B22ED" box="[800,920,898,921]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
described above and in fact few obvious distinguishing features are apparent that are likely to distinguish all examples, but that species is however allopatric, only known from the Angavo massif, and on average, the HWV Mb of
|
||||
<taxonomicName id="4C2E8DA74C25C6041A452EE7FAD92295" box="[1125,1290,968,993]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C25C6041A452EE7FAD92295" box="[1125,1290,968,993]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C25C6041A452EE7FAD92295" box="[1125,1290,968,993]" italics="true" pageId="83" pageNumber="84">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
is relatively straight and there is usually no space-M3 ocellus expressed on the HWD (
|
||||
<figureCitation id="1315EAA14C25C6041DFD2EC1FBFD2571" box="[989,1070,1004,1029]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="83" pageNumber="84">Fig. 24</figureCitation>
|
||||
<figureCitation id="1315EAA14C25C6041DFD2EC1FBFD2571" box="[989,1070,1004,1029]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264621" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="83" pageNumber="84">Fig. 24</figureCitation>
|
||||
D–E). The wings have a slightly greyer cast.
|
||||
<taxonomicName id="4C2E8DA74C25C6041F06293CFE19255D" box="[294,458,1040,1065]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C25C6041F06293CFE19255D" box="[294,458,1040,1065]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C25C6041F06293CFE19255D" box="[294,458,1040,1065]" italics="true" pageId="83" pageNumber="84">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
also strongly resembles
|
||||
<taxonomicName id="4C2E8DA74C25C6041CCD293CFC53255D" box="[749,896,1040,1065]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C25C6041CCD293CFC53255D" box="[749,896,1040,1065]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C25C6041CCD293CFC53255D" box="[749,896,1040,1065]" italics="true" pageId="83" pageNumber="84">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
. It can be distinguished from that last species (which has slightly more conspicuous white spots on the ventral wing surfaces) by the less straight, usually more evenly outangled HWV Mb at space M
|
||||
<quantity id="4CD65BC14C25C6041C6F2975FDA92504" box="[591,634,1111,1136]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="83" pageNumber="84" unit="in" value="2.0">2 in</quantity>
|
||||
<taxonomicName id="4C2E8DA74C25C6041CA02974FCDC2504" box="[640,783,1111,1136]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C25C6041CA02974FCDC2504" box="[640,783,1111,1136]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C25C6041CA02974FCDC2504" box="[640,783,1111,1136]" italics="true" pageId="83" pageNumber="84">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C25C6041D3E297AFCA22504" box="[798,881,1111,1136]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="83" pageNumber="84">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C25C6041D3E297AFCA22504" box="[798,881,1111,1136]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="83" pageNumber="84">Fig. 22</figureCitation>
|
||||
C), and from more yellowish lowland
|
||||
<taxonomicName id="4C2E8DA74C25C6041B3B2974FA4F2504" box="[1307,1436,1111,1136]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C25C6041B3B2974FA4F2504" box="[1307,1436,1111,1136]" italics="true" pageId="83" pageNumber="84">Ht. strigula</emphasis>
|
||||
|
|
@ -340,21 +340,21 @@ which have a conspicuous orange crescent on the HWD, and from
|
|||
<emphasis id="B95A2A364C25C6041DBF2950FBCF25E0" box="[927,1052,1147,1172]" italics="true" pageId="83" pageNumber="84">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
which is smaller and has ocellus space-M3 as well as in space-CuA1 expressed on the HWD (
|
||||
<figureCitation id="1315EAA14C25C6041D7129B2FC7725C3" box="[849,932,1183,1208]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="83" pageNumber="84">Fig. 19</figureCitation>
|
||||
<figureCitation id="1315EAA14C25C6041D7129B2FC7725C3" box="[849,932,1183,1208]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="83" pageNumber="84">Fig. 19</figureCitation>
|
||||
C). The ventrally very similar
|
||||
<taxonomicName id="4C2E8DA74C25C6041ADE298DFA4F25C3" box="[1278,1436,1184,1207]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C25C6041ADE298DFA4F25C3" box="[1278,1436,1184,1207]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C25C6041ADE298DFA4F25C3" box="[1278,1436,1184,1207]" italics="true" pageId="83" pageNumber="84">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
has a strong tendency to expression of a small ocellus M3 on the HWD (
|
||||
<figureCitation id="1315EAA14C25C6041D9C29EEFBDF25AF" box="[956,1036,1219,1244]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="83" pageNumber="84">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C25C6041D9C29EEFBDF25AF" box="[956,1036,1219,1244]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="83" pageNumber="84">Fig. 22</figureCitation>
|
||||
A–B). The potentially sympatric (in ‘Antsianaka’ region)
|
||||
<taxonomicName id="4C2E8DA74C25C6041FAA29C5FDD0258B" box="[394,515,1256,1279]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="anceps">
|
||||
<emphasis id="B95A2A364C25C6041FAA29C5FDD0258B" box="[394,515,1256,1279]" italics="true" pageId="83" pageNumber="84">Ht. anceps</emphasis>
|
||||
</taxonomicName>
|
||||
has a relatively straight HWV Mb and a yellowish ventral cast with sparse light brown irroration and a distinct black ventral androconial patch on the ♂ sternal abdomen (
|
||||
<figureCitation id="1315EAA14C25C6041AEB2827FAF02457" box="[1227,1315,1290,1315]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="83" pageNumber="84">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C25C6041AEB2827FAF02457" box="[1227,1315,1290,1315]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="83" pageNumber="84">Fig. 22</figureCitation>
|
||||
D). The ♂ genitalia are similar to those of
|
||||
<taxonomicName id="4C2E8DA74C25C6041FDB2802FDB82432" box="[507,619,1327,1350]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C25C6041FDB2802FDB82432" box="[507,619,1327,1350]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C25C6041FDB2802FDB82432" box="[507,619,1327,1350]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
, differing by the more strongly inpointed subterminal spine on the valve and the terminal head covered in spinoid setae being enlarged. They also differ from the potentially sympatric
|
||||
|
|
@ -367,11 +367,11 @@ D). The ♂ genitalia are similar to those of
|
|||
, being smaller and with a narrower distal valve tip (
|
||||
<bibRefCitation id="EFBF8BD54C25C6041A28285BFB1624FA" author="Lees" box="[1032,1221,1398,1422]" pageId="83" pageNumber="84" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 107</bibRefCitation>
|
||||
). DNA differences separating
|
||||
<taxonomicName id="4C2E8DA74C25C6041F3128B7FE6324C6" box="[273,432,1433,1458]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C25C6041F3128B7FE6324C6" box="[273,432,1433,1458]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C25C6041F3128B7FE6324C6" box="[273,432,1433,1458]" italics="true" pageId="83" pageNumber="84">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
from
|
||||
<taxonomicName id="4C2E8DA74C25C6041FD428B7FDB724C5" box="[500,612,1434,1457]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C25C6041FD428B7FDB724C5" box="[500,612,1434,1457]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C25C6041FD428B7FDB724C5" box="[500,612,1434,1457]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
are indicated by
|
||||
|
|
@ -387,15 +387,15 @@ Aduse-Poku
|
|||
<paragraph id="8B91F6244C25C6031EE728CDFB4E21A0" blockId="83.[151,1437,151,2030]" lastBlockId="84.[151,1437,151,2013]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
|
||||
<emphasis id="B95A2A364C25C6041EE728CDFE7C248D" bold="true" box="[199,431,1504,1529]" pageId="83" pageNumber="84">Description. Wings</emphasis>
|
||||
: description is similar to that of
|
||||
<taxonomicName id="4C2E8DA74C25C6041D1A28CFFC7D248D" box="[826,942,1506,1529]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C25C6041D1A28CFFC7D248D" box="[826,942,1506,1529]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C25C6041D1A28CFFC7D248D" box="[826,942,1506,1529]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
. Upperside uniform mid brown, with FW space-CuA1 ocellus featuring an orange ocellus ‘ring’ which is sometimes rather ‘squashed’ along the diagonal from mid-costa to tergal angle. FWD space-M1 ocellus small with narrow orange ring. HWD space-CuA1 ocellus is the only one expressed there and is somewhat elliptic, with a narrow orange ring. Darkish brown, diffuse, slightly wavy and uprecurved hair-brush emanates mainly from below vein 1A+2A, as far as mid-vein. Underside greyish brown, similar to
|
||||
<taxonomicName id="4C2E8DA74C25C6041FE52BB8FDEA27D8" box="[453,569,1685,1708]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C25C6041FE52BB8FDEA27D8" box="[453,569,1685,1708]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C25C6041FE52BB8FDEA27D8" box="[453,569,1685,1708]" italics="true" pageId="83" pageNumber="84">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
, rather more uniform and less strongly irrorated than in many
|
||||
<taxonomicName id="4C2E8DA74C25C6041B3C2BB8FA4727D8" box="[1308,1428,1685,1708]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C25C6041B3C2BB8FA4727D8" box="[1308,1428,1685,1708]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C25C6041B3C2BB8FA4727D8" box="[1308,1428,1685,1708]" italics="true" pageId="83" pageNumber="84">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
, rather uniformly irrorated brown and ochreous brown. Space-CuA1 ocellus with orange ring yellowing proximad, following darkish brown concave curve of Mb before it bends back to mid-costa. Space-M1 ocellus FWV a white pupil in HT. On HWV, space-CuA1 ocellus small and slightly elliptic with narrow black iris, also without orange ring trace. HWV space-Rs-M2 ocelli as white pupil-points and space-CuA2 ocellus hardly expressed in HT. HWV Mb darkish brown and fairly straight but minutely irregular, only gently curving, with very small yellow Mf distad of it in space-M2. FWV cell area with four darker brown rather equally spaced transverse Cbs. Basal areas not much darker than areas distad of Mb, and darker brown PMb weakly represented.
|
||||
|
|
@ -425,9 +425,9 @@ series was taken in a few days in early November. Space- M1 ocellus FWV sometime
|
|||
<paragraph id="8B91F6244C22C6031EE72F45FECB2292" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||||
<emphasis id="B95A2A364C22C6031EE72F45FE9823F5" bold="true" box="[199,331,616,641]" pageId="84" pageNumber="85">♂ genitalia</emphasis>
|
||||
: 371 DL (
|
||||
<figureCitation id="1315EAA14C22C6031FE42F45FDCB23F5" box="[452,536,616,641]" captionStart="FIGURE 25" captionStartId="80.[151,250,1045,1067]" captionTargetBox="[254,1341,201,1005]" captionTargetId="figure@80.[231,1356,193,1023]" captionTargetPageId="80" captionText="FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264622/files/figure.png" pageId="84" pageNumber="85">Fig. 25</figureCitation>
|
||||
<figureCitation id="1315EAA14C22C6031FE42F45FDCB23F5" box="[452,536,616,641]" captionStart="FIGURE 25" captionStartId="80.[151,250,1045,1067]" captionTargetBox="[254,1341,201,1005]" captionTargetId="figure@80.[231,1356,193,1023]" captionTargetPageId="80" captionText="FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264622" httpUri="https://zenodo.org/record/264622/files/figure.png" pageId="84" pageNumber="85">Fig. 25</figureCitation>
|
||||
B, PT): description more or less as for
|
||||
<taxonomicName id="4C2E8DA74C22C6031DF42F44FB9523F4" box="[980,1094,617,640]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C22C6031DF42F44FB9523F4" box="[980,1094,617,640]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C22C6031DF42F44FB9523F4" box="[980,1094,617,640]" italics="true" pageId="84" pageNumber="85">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
: from LV, uncus very slightly proud of dorsal curve of tegumen (which has a small proximad notch from DV) and slightly longer than it, fairly straight and parallel-sided to hook without a distinct ‘head’ towards tip (inflated from DV before tip), fairly narrow at hinge with vinculum. Valve without prominent dorsal ‘shoulder’. Valve arm with fairly short ‘neck’ and slight club at tip covered in spinoid setae (its crest particularly raised from DV), with distinct ‘beak’ oriented towards uncus and slightly mesad, with none of valve tip proud of uncus. Gnathos from rather small base with deepened (although not ear-like structure) near base, recurved downwards at tapered tip (appearing sinuate from DV and slightly inrecurved at pointed tip). Saccus not very long and parallel sided, aedeagus slightly shorter than valve and quite recurved, both towards and away from ostium, also proximally uprecurved. ♂ genitalia different from those of potentially sympatric (
|
||||
|
|
@ -469,11 +469,11 @@ and of
|
|||
<bibRefCitation id="EFBF8BD54C22C6031AB929ADFA9425ED" author="Oberthur" box="[1177,1351,1152,1177]" pageId="84" pageNumber="85" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(LT ♂ designated above under
|
||||
<taxonomicName id="4C2E8DA74C22C6031F862988FD9125C8" box="[422,578,1189,1212]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C22C6031F862988FD9125C8" box="[422,578,1189,1212]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C22C6031F862988FD9125C8" box="[422,578,1189,1212]" italics="true" pageId="84" pageNumber="85">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
) were examined but the species strongly resembles
|
||||
<taxonomicName id="4C2E8DA74C22C6031AA92988FB2A25C8" box="[1161,1273,1189,1212]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C22C6031AA92988FB2A25C8" box="[1161,1273,1189,1212]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C22C6031AA92988FB2A25C8" box="[1161,1273,1189,1212]" italics="true" pageId="84" pageNumber="85">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
(see above).
|
||||
|
|
@ -481,7 +481,7 @@ and of
|
|||
<paragraph id="8B91F6244C22C6031EE729EAFEFE245C" blockId="84.[151,1437,151,2013]" pageId="84" pageNumber="85">
|
||||
<emphasis id="B95A2A364C22C6031EE729EAFD202594" bold="true" box="[199,755,1223,1248]" pageId="84" pageNumber="85">Additional information. DNA divergences:</emphasis>
|
||||
COI-5P cluster number BOLD:ACW4996 (exemplar BMAD242-15, DL06-11), about 2.58% divergent to
|
||||
<taxonomicName id="4C2E8DA74C22C6031D3229C0FC592470" box="[786,906,1261,1284]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C22C6031D3229C0FC592470" box="[786,906,1261,1284]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C22C6031D3229C0FC592470" box="[786,906,1261,1284]" italics="true" pageId="84" pageNumber="85">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
(BOLD:AAE4112, exemplar BMAD200-15, DL14Z-051).
|
||||
|
|
@ -493,7 +493,7 @@ COI-5P cluster number BOLD:ACW4996 (exemplar BMAD242-15, DL06-11), about 2.58% d
|
|||
: this species was not known at the time of
|
||||
<bibRefCitation id="EFBF8BD54C22C6031DE92819FB80243F" author="Lees" box="[969,1107,1331,1355]" pageId="84" pageNumber="85" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997)</bibRefCitation>
|
||||
. Closely related to
|
||||
<taxonomicName id="4C2E8DA74C22C6031B0C2819FA4F243F" box="[1324,1436,1332,1355]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C22C6031B0C2819FA4F243F" box="[1324,1436,1332,1355]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C22C6031B0C2819FA4F243F" box="[1324,1436,1332,1355]" italics="true" pageId="84" pageNumber="85">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
(sister in
|
||||
|
|
@ -528,7 +528,7 @@ Aduse-Poku
|
|||
<paragraph id="8B91F6244C22C6031EE72B7CFBA8271E" blockId="84.[151,1437,151,2013]" box="[199,1147,1617,1642]" pageId="84" pageNumber="85">
|
||||
<emphasis id="B95A2A364C22C6031EE72B7CFE8A271E" bold="true" box="[199,345,1617,1642]" pageId="84" pageNumber="85">Distribution</emphasis>
|
||||
: as far as is known, endemic to RNI Zahamena (
|
||||
<figureCitation id="1315EAA14C22C6031D5C2B7CFC1C271D" box="[892,975,1617,1642]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="84" pageNumber="85">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C22C6031D5C2B7CFC1C271D" box="[892,975,1617,1642]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="84" pageNumber="85">Fig. 30</figureCitation>
|
||||
C, brown dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C22C6031EE72B59FC2427F9" blockId="84.[151,1437,151,2013]" box="[199,1015,1652,1677]" pageId="84" pageNumber="85">
|
||||
|
|
@ -612,7 +612,7 @@ _0825 [photo]; ♀, data as above but:
|
|||
06-119, IA432 [isotope voucher], KAP518 [=KA-P518; extract number, sequenced specimen in Aduse-Poku
|
||||
<emphasis id="B95A2A364C22C6031B272AB2FA9426C2" box="[1287,1351,1949,1974]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
|
||||
, 2015, mislabelled as
|
||||
<taxonomicName id="4C2E8DA74C22C6031F1D2AE8FDD826A8" box="[317,523,1989,2012]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="anceps">
|
||||
<taxonomicName id="4C2E8DA74C22C6031F1D2AE8FDD826A8" box="[317,523,1989,2012]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="anceps">
|
||||
<emphasis id="B95A2A364C22C6031F1D2AE8FDD826A8" box="[317,523,1989,2012]" italics="true" pageId="84" pageNumber="85">Heteropsis anceps</emphasis>
|
||||
</taxonomicName>
|
||||
); ♀, data as above but:
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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|
||||
<taxonomicName id="4C2E8DA74C2EC60F1EB72DBAFD5421C6" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[151,647,151,178]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1EB72DBAFD5421C6" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[151,647,151,178]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C2EC60F1EB72DBAFEA121C5" bold="true" box="[151,370,151,177]" italics="true" pageId="88" pageNumber="89">Heteropsis vertigo</emphasis>
|
||||
Lees & Raharitsimba
|
||||
</taxonomicName>
|
||||
|
|
@ -71,7 +71,7 @@ material., Deposition
|
|||
:
|
||||
</emphasis>
|
||||
♂ (
|
||||
<figureCitation id="1315EAA14C2EC60F1D702C08FC7F204A" box="[848,940,293,318]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="88" pageNumber="89">Fig. 28</figureCitation>
|
||||
<figureCitation id="1315EAA14C2EC60F1D702C08FC7F204A" box="[848,940,293,318]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="88" pageNumber="89">Fig. 28</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C2EC60F1DC12C08FBBB204A" box="[993,1128,293,318]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||||
NW, Bekolosy Mt., RS Manongarivo, c.
|
||||
|
|
@ -116,7 +116,7 @@ barcoded specimen],
|
|||
010289174 [
|
||||
<collectingCountry id="F339B6B44C2EC60F1A062C9BFBB220BA" box="[1062,1121,438,462]" name="Qatar" pageId="88" pageNumber="89">QTR</collectingCountry>
|
||||
barcode]; ♀ (
|
||||
<figureCitation id="1315EAA14C2EC60F1B3E2C98FAAF20BA" box="[1310,1404,437,462]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="88" pageNumber="89">Fig. 28</figureCitation>
|
||||
<figureCitation id="1315EAA14C2EC60F1B3E2C98FAAF20BA" box="[1310,1404,437,462]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="88" pageNumber="89">Fig. 28</figureCitation>
|
||||
B),
|
||||
<collectingCountry id="F339B6B44C2EC60F1EB72CF6FECD2080" box="[151,286,475,500]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||||
E, Bekolosy Mt., RS Manongarivo,
|
||||
|
|
@ -140,7 +140,7 @@ NW,
|
|||
010289175 [
|
||||
<collectingCountry id="F339B6B44C2EC60F1FAC2F0FFE14234E" box="[396,455,546,570]" name="Qatar" pageId="88" pageNumber="89">QTR</collectingCountry>
|
||||
barcode]; ♂ (
|
||||
<figureCitation id="1315EAA14C2EC60F1C412F0CFD61234E" box="[609,690,545,570]" captionStart="FIGURE 29" captionStartId="90.[151,250,1420,1442]" captionTargetBox="[179,1434,202,1356]" captionTargetId="figure@90.[161,1455,193,1385]" captionTargetPageId="90" captionText="FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="88" pageNumber="89">Fig. 29</figureCitation>
|
||||
<figureCitation id="1315EAA14C2EC60F1C412F0CFD61234E" box="[609,690,545,570]" captionStart="FIGURE 29" captionStartId="90.[151,250,1420,1442]" captionTargetBox="[179,1434,202,1356]" captionTargetId="figure@90.[161,1455,193,1385]" captionTargetPageId="90" captionText="FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264626" httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="88" pageNumber="89">Fig. 29</figureCitation>
|
||||
A, genitalia),
|
||||
<collectingCountry id="F339B6B44C2EC60F1D6A2F0CFC02234E" box="[842,977,545,570]" name="Madagascar" pageId="88" pageNumber="89">Madagascar</collectingCountry>
|
||||
NW, Bekolosy, RS Manongarivo, ~
|
||||
|
|
@ -268,7 +268,7 @@ Bekolosy Mt., RS Manongarivo, c.
|
|||
<paragraph id="8B91F6244C2EC60F1EE72931FE01247C" blockId="88.[151,1437,258,2028]" pageId="88" pageNumber="89">
|
||||
<emphasis id="B95A2A364C2EC60F1EE72931FE922541" bold="true" box="[199,321,1052,1077]" pageId="88" pageNumber="89">Diagnosis.</emphasis>
|
||||
In its translucence, the extended FWD space-CuA1 Orng of
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1DC92933FBB12540" box="[1001,1122,1053,1077]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1DC92933FBB12540" box="[1001,1122,1053,1077]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<emphasis id="B95A2A364C2EC60F1DC92933FBB12540" box="[1001,1122,1053,1077]" italics="true" pageId="88" pageNumber="89">Ht. vertigo</emphasis>
|
||||
</taxonomicName>
|
||||
is intermediate between
|
||||
|
|
@ -276,11 +276,11 @@ is intermediate between
|
|||
<emphasis id="B95A2A364C2EC60F1B592933FF3E252C" italics="true" pageId="88" pageNumber="89">Ht. turbans</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1F03296CFE5E252C" box="[291,397,1087,1112]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1F03296CFE5E252C" box="[291,397,1087,1112]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<emphasis id="B95A2A364C2EC60F1F03296CFE5E252C" box="[291,397,1087,1112]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1FBB296CFDC7252C" box="[411,532,1089,1112]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1FBB296CFDC7252C" box="[411,532,1089,1112]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<emphasis id="B95A2A364C2EC60F1FBB296CFDC7252C" box="[411,532,1089,1112]" italics="true" pageId="88" pageNumber="89">Ht. vertigo</emphasis>
|
||||
</taxonomicName>
|
||||
in wing pattern resembles
|
||||
|
|
@ -288,21 +288,21 @@ in wing pattern resembles
|
|||
<emphasis id="B95A2A364C2EC60F1D63296CFC11252C" box="[835,962,1087,1112]" italics="true" pageId="88" pageNumber="89">Ht. turbans</emphasis>
|
||||
</taxonomicName>
|
||||
more than it does
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1AB2296CFB2F252C" box="[1170,1276,1087,1112]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1AB2296CFB2F252C" box="[1170,1276,1087,1112]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<emphasis id="B95A2A364C2EC60F1AB2296CFB2F252C" box="[1170,1276,1087,1112]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||||
</taxonomicName>
|
||||
, but the FWD space-CuA1 Orng is largely suffused with orange and only pale whitish-translucent in the anteriad half, especially proximad, rather than throughout it, as in
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1CA529AAFD2025EB" box="[645,755,1158,1183]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1CA529AAFD2025EB" box="[645,755,1158,1183]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<emphasis id="B95A2A364C2EC60F1CA529AAFD2025EB" box="[645,755,1158,1183]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||||
</taxonomicName>
|
||||
, or largely orange as in
|
||||
<emphasis id="B95A2A364C2EC60F1A3829AAFAF925EA" box="[1048,1322,1158,1183]" italics="true" pageId="88" pageNumber="89">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1A3829AAFB4C25EB" box="[1048,1183,1158,1183]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">Ht. turbans</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1A8D29AAFAF925EA" box="[1197,1322,1159,1182]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo">Ht. vertigo</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1A8D29AAFAF925EA" box="[1197,1322,1159,1182]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="vertigo">Ht. vertigo</taxonomicName>
|
||||
</emphasis>
|
||||
lacks the extreme ear-like cupping of the gnathos seen in
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1CF32987FBC825B6" authority="Lees, 1997: 107" authorityName="Lees" authorityPageNumber="107" authorityYear="1997" box="[723,1051,1193,1218]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1CF32987FBC825B6" authority="Lees, 1997: 107" authorityName="Lees" authorityPageNumber="107" authorityYear="1997" box="[723,1051,1193,1218]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<emphasis id="B95A2A364C2EC60F1CF32987FC9125B6" box="[723,834,1193,1218]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C2EC60F1D752987FBC025B6" author="Lees" box="[853,1043,1194,1218]" pageId="88" pageNumber="89" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 107</bibRefCitation>
|
||||
|
|
@ -313,12 +313,12 @@ and
|
|||
<emphasis id="B95A2A364C2EC60F1A7B2987FB0C25B6" box="[1115,1247,1193,1218]" italics="true" pageId="88" pageNumber="89">Ht. turbans</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C2EC60F1AD32984FA9825B6" box="[1267,1355,1193,1218]" captionStart="FIGURE 29" captionStartId="90.[151,250,1420,1442]" captionTargetBox="[179,1434,202,1356]" captionTargetId="figure@90.[161,1455,193,1385]" captionTargetPageId="90" captionText="FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="88" pageNumber="89">Fig. 29</figureCitation>
|
||||
<figureCitation id="1315EAA14C2EC60F1AD32984FA9825B6" box="[1267,1355,1193,1218]" captionStart="FIGURE 29" captionStartId="90.[151,250,1420,1442]" captionTargetBox="[179,1434,202,1356]" captionTargetId="figure@90.[161,1455,193,1385]" captionTargetPageId="90" captionText="FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264626" httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="88" pageNumber="89">Fig. 29</figureCitation>
|
||||
B; NW Zahamena) as well as some related species (e.g.
|
||||
<emphasis id="B95A2A364C2EC60F1C9129E3FC0E2591" box="[689,989,1228,1253]" italics="true" pageId="88" pageNumber="89">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1C9129E3FCBB2591" box="[689,872,1228,1253]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="angulifascia">Ht. angulifascia</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1D5229E3FC0E2591" box="[882,989,1228,1253]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="borgo">Ht. borgo</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1D5229E3FC0E2591" box="[882,989,1228,1253]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="borgo">Ht. borgo</taxonomicName>
|
||||
</emphasis>
|
||||
), and the HWV Mb is much less kinked than in any of these species.
|
||||
</paragraph>
|
||||
|
|
@ -327,7 +327,7 @@ B; NW Zahamena) as well as some related species (e.g.
|
|||
<paragraph id="8B91F6244C2EC60F1EE7283EFB6E26D1" blockId="88.[151,1437,258,2028]" pageId="88" pageNumber="89">
|
||||
<emphasis id="B95A2A364C2EC60F1EE7283EFE7E2458" bold="true" box="[199,429,1299,1324]" pageId="88" pageNumber="89">Description. Wings</emphasis>
|
||||
: Similar to
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1C162839FCDF2458" box="[566,780,1299,1324]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1C162839FCDF2458" box="[566,780,1299,1324]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">
|
||||
<emphasis id="B95A2A364C2EC60F1C162839FCDF2458" box="[566,780,1299,1324]" italics="true" pageId="88" pageNumber="89">Heteropsis turbans</emphasis>
|
||||
</taxonomicName>
|
||||
. Upperside uniform mid brown, except with conspicuous orange Mf/crescent (as in a few other of the
|
||||
|
|
@ -339,11 +339,11 @@ B; NW Zahamena) as well as some related species (e.g.
|
|||
<emphasis id="B95A2A364C2EC60F1DF4281AFB84243B" box="[980,1111,1334,1359]" italics="true" pageId="88" pageNumber="89">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
) at base of HWD space-M2 following curve at base of vein M3. FW space-CuA1 ocellus with orange to pale yellow/translucent ‘ring,’ more like a hexagon that has been compressed along a diagonal running from mid-costa to tergal angle. FWD space-M1 ocellus quite small and elliptic with narrow orange ring. HWD space-CuA1 ocellus the only one expressed there and slightly elliptic, with narrow orange ring. Darkish brown, diffuse, slightly wavy, uprecurved hair brush emanates mainly from space-1A as far as mid-vein. Underside greyish brown, similar to
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1A5628CAFB26248B" box="[1142,1269,1510,1535]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1A5628CAFB26248B" box="[1142,1269,1510,1535]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="turbans">
|
||||
<emphasis id="B95A2A364C2EC60F1A5628CAFB26248B" box="[1142,1269,1510,1535]" italics="true" pageId="88" pageNumber="89">Ht. turbans</emphasis>
|
||||
</taxonomicName>
|
||||
but with a less kinked HWV Mb, and limited irroration, especially in basal area. Space-CuA1 ocellus with orange ring yellowing proximad following reddish brown concave curve of Mb before it bends back to mid-costa. Space-M1 ocellus fairly elliptic, with white pupil and wide narrow black iris, but narrow orange ring. HWV space-CuA1 ocellus large and round to elliptic with narrow pale orange ring. HWV space-Rs-M2 ocelli generally not expressed and space- CuA2 ocellus small and elliptic with faint orange ring. HWV Mb reddish brown and fairly straight but outflexed around M2, generally with a yellow patch (Mf) distad of it in space-M2, and curved back towards the anal angle in space-CuA2. Slight, wavy brown Pml tracks the wing margin and Sml more closely follows the fairly crenate (notably much less so than in the related
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1C592A2CFD36266C" box="[633,741,1791,1816]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<taxonomicName id="4C2E8DA74C2EC60F1C592A2CFD36266C" box="[633,741,1791,1816]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<emphasis id="B95A2A364C2EC60F1C592A2CFD36266C" box="[633,741,1791,1816]" italics="true" pageId="88" pageNumber="89">Ht. sabas</emphasis>
|
||||
</taxonomicName>
|
||||
) HW margin. FWV cell area with four darker brown rather equally spaced transverse Cbs, the outermost of which flank area that is paler than for the inner pair. Basal areas not much darker than areas distad of the Mb, and darker brown PMb weakly represented.
|
||||
|
|
@ -360,13 +360,13 @@ series was caught near the start of the rainy season in mid December. ♀♀ sim
|
|||
<quantity id="4CD65BC14C2EC60F1B7D2A82FF28269F" metricMagnitude="-2" metricUnit="m" metricValue="1.9449999999999998" metricValueMax="1.95" metricValueMin="1.94" pageId="88" pageNumber="89" unit="mm" value="19.45" valueMax="19.5" valueMin="19.4">19.4– 19.5 mm</quantity>
|
||||
(n=2 ♀♀).
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C2FC60E1EB72A64FB0126CF" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="89" pageNumber="90" targetBox="[278,1298,207,1823]" targetPageId="89">
|
||||
<caption id="DF51A6AC4C2FC60E1EB72A64FB0126CF" ID-DOI="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="89" pageNumber="90" targetBox="[278,1298,207,1823]" targetPageId="89">
|
||||
<paragraph id="8B91F6244C2FC60E1EB72A64FB0126CF" blockId="89.[151,1436,1865,1979]" pageId="89" pageNumber="90">
|
||||
<emphasis id="B95A2A364C2FC60E1EB72A64FEF2262A" bold="true" box="[151,289,1865,1887]" pageId="89" pageNumber="90">FIGURE 28.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
<emphasis id="B95A2A364C2FC60E1C582A64FD59262A" bold="true" box="[632,650,1865,1886]" pageId="89" pageNumber="90">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C2FC60E1CDA2A67FB4F262B" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[762,1180,1865,1887]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="vertigo" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C2FC60E1CDA2A67FB4F262B" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[762,1180,1865,1887]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="vertigo" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C2FC60E1CDA2A67FC64262B" box="[762,951,1866,1887]" italics="true" pageId="89" pageNumber="90">Heteropsis vertigo</emphasis>
|
||||
Lees & Raharitsimba
|
||||
</taxonomicName>
|
||||
|
|
@ -377,7 +377,7 @@ Lees & Raharitsimba
|
|||
(Mt. Bekolosy; NHMUK010289173).
|
||||
<emphasis id="B95A2A364C2FC60E1F5D2A45FE5D2609" bold="true" box="[381,398,1896,1917]" pageId="89" pageNumber="90">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C2FC60E1FD22A44FD8C260A" box="[498,607,1897,1918]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<taxonomicName id="4C2E8DA74C2FC60E1FD22A44FD8C260A" box="[498,607,1897,1918]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<emphasis id="B95A2A364C2FC60E1FD22A44FD8C260A" box="[498,607,1897,1918]" italics="true" pageId="89" pageNumber="90">Ht. vertigo</emphasis>
|
||||
</taxonomicName>
|
||||
(Mt. Bekolosy; NHMUK010289175).
|
||||
|
|
@ -407,13 +407,13 @@ Oberthür, 1923
|
|||
(‘Tamatave’; BMNH(E) #674789). Scale bar: 30 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF51A6AC4C2CC60D1EB728A1FBA124B4" httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="90" pageNumber="91" targetBox="[179,1434,202,1356]" targetPageId="90">
|
||||
<caption id="DF51A6AC4C2CC60D1EB728A1FBA124B4" ID-DOI="http://doi.org/10.5281/zenodo.264626" httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="90" pageNumber="91" targetBox="[179,1434,202,1356]" targetPageId="90">
|
||||
<paragraph id="8B91F6244C2CC60D1EB728A1FBA124B4" blockId="90.[151,1436,1420,1472]" pageId="90" pageNumber="91">
|
||||
<emphasis id="B95A2A364C2CC60D1EB728A1FEF724D5" bold="true" box="[151,292,1420,1442]" pageId="90" pageNumber="91">FIGURE 29.</emphasis>
|
||||
♂ genitalia in LV (left), DV (right).
|
||||
<emphasis id="B95A2A364C2CC60D1C9D28A1FD1C24D5" bold="true" box="[701,719,1420,1441]" pageId="90" pageNumber="91">A</emphasis>
|
||||
:
|
||||
<taxonomicName id="4C2E8DA74C2CC60D1CC128A0FB5C24D6" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[737,1167,1420,1442]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="vertigo" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C2CC60D1CC128A0FB5C24D6" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[737,1167,1420,1442]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="vertigo" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C2CC60D1CC128A0FC9D24D6" box="[737,846,1421,1442]" italics="true" pageId="90" pageNumber="91">Heteropsis</emphasis>
|
||||
<emphasis id="B95A2A364C2CC60D1D7928A0FC7224D6" box="[857,929,1421,1442]" italics="true" pageId="90" pageNumber="91">vertigo</emphasis>
|
||||
Lees & Raharitsimba
|
||||
|
|
@ -452,7 +452,7 @@ and
|
|||
<paragraph id="8B91F6244C2CC60C1EE72A2AFC88218C" blockId="90.[151,1437,1511,2005]" lastBlockId="91.[151,1437,151,1910]" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
|
||||
<emphasis id="B95A2A364C2CC60D1EE72A2AFE9D2654" bold="true" box="[199,334,1799,1824]" pageId="90" pageNumber="91">♂ genitalia</emphasis>
|
||||
: 366DL, PT (
|
||||
<figureCitation id="1315EAA14C2CC60D1FD82A2AFD832654" box="[504,592,1799,1824]" captionStart="FIGURE 29" captionStartId="90.[151,250,1420,1442]" captionTargetBox="[179,1434,202,1356]" captionTargetId="figure@90.[161,1455,193,1385]" captionTargetPageId="90" captionText="FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="90" pageNumber="91">Fig. 29</figureCitation>
|
||||
<figureCitation id="1315EAA14C2CC60D1FD82A2AFD832654" box="[504,592,1799,1824]" captionStart="FIGURE 29" captionStartId="90.[151,250,1420,1442]" captionTargetBox="[179,1434,202,1356]" captionTargetId="figure@90.[161,1455,193,1385]" captionTargetPageId="90" captionText="FIGURE 29. ♂ genitalia in LV (left), DV (right). A: Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 159, 366 DL). B: Ht. turbans (Oberthür, 1916) (Zahamena; 370 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264626" httpUri="https://zenodo.org/record/264626/files/figure.png" pageId="90" pageNumber="91">Fig. 29</figureCitation>
|
||||
A): from LV, uncus fairly straight in line with tegumen in non-deflexed position and recurved down to a sharply hooked point, with a distinct ‘head’ before tip; tegumen about same length as uncus and constriction at fusion line with vinculum only moderate, such that tegumen from LV tapers only slightly from a rectangular frame; tegumen features wide and shallow proximad notch from DV. Gnathos on angled base, flattened distinctly before tip into broadened structure (not nearly as earlike as for example in
|
||||
<taxonomicName id="4C2E8DA74C2CC60D1ACE2AB4FABF26C4" box="[1262,1388,1943,1968]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="turbans">
|
||||
<emphasis id="B95A2A364C2CC60D1ACE2AB4FABF26C4" box="[1262,1388,1943,1968]" italics="true" pageId="90" pageNumber="91">Ht. turbans</emphasis>
|
||||
|
|
@ -480,7 +480,7 @@ A direct reference to the vertiginous waterfall directly below the camp at Bekol
|
|||
<paragraph id="8B91F6244C2DC60C1EE72CD2FBB52541" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||||
<emphasis id="B95A2A364C2DC60C1EE72CD2FE9F236C" bold="true" box="[199,332,511,536]" pageId="91" pageNumber="92">Discussion.</emphasis>
|
||||
No historical museum collections of
|
||||
<taxonomicName id="4C2E8DA74C2DC60C1D2C2F2CFC58236C" box="[780,907,513,536]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<taxonomicName id="4C2E8DA74C2DC60C1D2C2F2CFC58236C" box="[780,907,513,536]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<emphasis id="B95A2A364C2DC60C1D2C2F2CFC58236C" box="[780,907,513,536]" italics="true" pageId="91" pageNumber="92">Ht. vertigo</emphasis>
|
||||
</taxonomicName>
|
||||
are known. When first found in the field in
|
||||
|
|
@ -492,11 +492,11 @@ are known. When first found in the field in
|
|||
(indeed
|
||||
<bibRefCitation id="EFBF8BD54C2DC60C1FA02F65FE282314" author="Lees" box="[384,507,584,608]" pageId="91" pageNumber="92" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
|
||||
: 225
|
||||
<figureCitation id="1315EAA14C2DC60C1C1C2F6AFDAD2314" box="[572,638,583,608]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="91" pageNumber="92">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C2DC60C1C1C2F6AFDAD2314" box="[572,638,583,608]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="91" pageNumber="92">Fig. 3</figureCitation>
|
||||
K erroneously shows the distribution of that species extending to the northwest of
|
||||
<collectingCountry id="F339B6B44C2DC60C1F0C2F41FE6A23F1" box="[300,441,620,645]" name="Madagascar" pageId="91" pageNumber="92">Madagascar</collectingCountry>
|
||||
).
|
||||
<taxonomicName id="4C2E8DA74C2DC60C1FEB2F40FD9723F0" box="[459,580,621,644]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<taxonomicName id="4C2E8DA74C2DC60C1FEB2F40FD9723F0" box="[459,580,621,644]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="vertigo">
|
||||
<emphasis id="B95A2A364C2DC60C1FEB2F40FD9723F0" box="[459,580,621,644]" italics="true" pageId="91" pageNumber="92">Ht. vertigo</emphasis>
|
||||
</taxonomicName>
|
||||
has since been found on the adjacent mountain of Antsatrotro in
|
||||
|
|
@ -507,7 +507,7 @@ has since been found on the adjacent mountain of Antsatrotro in
|
|||
<bibRefCitation id="EFBF8BD54C2DC60C1EB72F99FEEF23B9" author="Oberthur" box="[151,316,692,717]" pageId="91" pageNumber="92" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(LT ♂, here designated,
|
||||
<figureCitation id="1315EAA14C2DC60C1C7B2F99FD7D23B8" box="[603,686,692,717]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="91" pageNumber="92">Fig. 28</figureCitation>
|
||||
<figureCitation id="1315EAA14C2DC60C1C7B2F99FD7D23B8" box="[603,686,692,717]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="91" pageNumber="92">Fig. 28</figureCitation>
|
||||
C, specimen bearing labels “
|
||||
<collectingCountry id="F339B6B44C2DC60C1DD02F99FBA923B9" box="[1008,1146,692,717]" name="Madagascar" pageId="91" pageNumber="92">Madagascar</collectingCountry>
|
||||
Antsianaka Perrot Freres 2e Semestre 1893|
|
||||
|
|
@ -533,7 +533,7 @@ Culapa turbans
|
|||
<bibRefCitation id="EFBF8BD54C2DC60C1C672E32FD25224C" author="Oberthur" box="[583,758,799,824]" pageId="91" pageNumber="92" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(LT, here designated,
|
||||
<figureCitation id="1315EAA14C2DC60C1A222E32FB84224C" box="[1026,1111,799,824]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="91" pageNumber="92">Fig. 28</figureCitation>
|
||||
<figureCitation id="1315EAA14C2DC60C1A222E32FB84224C" box="[1026,1111,799,824]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="91" pageNumber="92">Fig. 28</figureCitation>
|
||||
D, specimen bearing labels: “Antsianaka et lac Alaotra 2e Trimestre 1889 Perrot Freres|[copy of
|
||||
<bibRefCitation id="EFBF8BD54C2DC60C1D8C2E69FB862229" author="Oberthur" box="[940,1109,836,861]" pageId="91" pageNumber="92" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür 1916</bibRefCitation>
|
||||
, Pl. 356, f. 3057]|BMNH(E) #674784; automatically then, PLT ♂: BMNH(E) #674786; PLT ♀♀: BMNH(E) #674785) and BMNH(E) #674787, with same locality data).
|
||||
|
|
@ -547,7 +547,7 @@ was considered a synonym of
|
|||
by Lees
|
||||
<emphasis id="B95A2A364C2DC60C1ACC2EA0FAF922D1" box="[1260,1322,908,933]" italics="true" pageId="91" pageNumber="92">et al.,</emphasis>
|
||||
2003 (see also below). Also examined (
|
||||
<figureCitation id="1315EAA14C2DC60C1FCC2E82FD9322BC" box="[492,576,943,968]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="91" pageNumber="92">Fig. 28</figureCitation>
|
||||
<figureCitation id="1315EAA14C2DC60C1FCC2E82FD9322BC" box="[492,576,943,968]" captionStart="FIGURE 28" captionStartId="89.[151,250,1865,1887]" captionTargetBox="[278,1298,207,1823]" captionTargetId="figure@89.[278,1309,193,1844]" captionTargetPageId="89" captionText="FIGURE 28. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis vertigo Lees & Raharitsimba, sp. nov. (Mt. Bekolosy; NHMUK 010289173). B: PT ♀ of Ht. vertigo (Mt. Bekolosy; NHMUK 010289175). C: LT ♂ of Culapa turbans Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674780). D: LT ♂ of Culapa curvatula Oberthür, 1916 (‘ Antsianaka et Lac Alaotra’; BMNH (E) # 674784). E: HT ♂ of Culapa sabas Oberthür, 1923 (‘ Tamatave’; BMNH (E) # 674789). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264625" httpUri="https://zenodo.org/record/264625/files/figure.png" pageId="91" pageNumber="92">Fig. 28</figureCitation>
|
||||
E) was the HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C2DC60C1D182E82FB5F22BC" authority="Oberthur, 1923" authorityName="Oberthur" authorityYear="1923" box="[824,1164,943,968]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<emphasis id="B95A2A364C2DC60C1D182E82FC0722BC" box="[824,980,943,968]" italics="true" pageId="91" pageNumber="92">Culapa sabas</emphasis>
|
||||
|
|
@ -637,7 +637,7 @@ and
|
|||
<paragraph id="8B91F6244C2DC60C1EE72B1AFE5C2701" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||||
<emphasis id="B95A2A364C2DC60C1EE72B1AFE8A2724" bold="true" box="[199,345,1591,1616]" pageId="91" pageNumber="92">Distribution</emphasis>
|
||||
: as far as is known endemic to Réserve Speciale de Manongarivo, Mts. Bekolosy and Antsatrotro (
|
||||
<figureCitation id="1315EAA14C2DC60C1EBF2B71FF212700" box="[159,242,1628,1653]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="91" pageNumber="92">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C2DC60C1EBF2B71FF212700" box="[159,242,1628,1653]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="91" pageNumber="92">Fig. 30</figureCitation>
|
||||
B, fawn dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C2DC60C1EE72B52FEDA27C9" blockId="91.[151,1437,151,1910]" pageId="91" pageNumber="92">
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<taxonomicName id="4C2E8DA74C31C6101EB72DBAFDC621C6" authority="Lees" authorityName="Lees" box="[151,533,151,178]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides" status="sp. nov.">
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<taxonomicName id="4C2E8DA74C31C6101EB72DBAFDC621C6" authority="Lees" authorityName="Lees" box="[151,533,151,178]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides" status="sp. nov.">
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<emphasis id="B95A2A364C31C6101EB72DBAFE0021C5" bold="true" box="[151,467,151,177]" italics="true" pageId="71" pageNumber="72">Heteropsis menamenoides</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -76,7 +76,7 @@ material.,
|
|||
<collectionCode id="ED3F6EE14C31C6101CA52C0BFD0F204B" box="[645,732,294,319]" pageId="71" pageNumber="72">BMNH</collectionCode>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C31C6101D322C0BFCB6204B" box="[786,869,294,319]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="71" pageNumber="72">Fig. 21</figureCitation>
|
||||
<figureCitation id="1315EAA14C31C6101D322C0BFCB6204B" box="[786,869,294,319]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264618" httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="71" pageNumber="72">Fig. 21</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C31C6101DB12C0BFBCB204B" box="[913,1048,294,319]" name="Madagascar" pageId="71" pageNumber="72">Madagascar</collectingCountry>
|
||||
NW, Bekolosy, RS Manongarivo, 950 [880] m, [
|
||||
|
|
@ -136,7 +136,7 @@ extract number],
|
|||
barcode voucher], KA571, KA4084 [=KA-P571, KA-P4084;
|
||||
<collectionCode id="ED3F6EE14C31C6101D782F28FC462369" box="[856,917,517,541]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="71" pageNumber="72">DNA</collectionCode>
|
||||
extract numbers]; ♂ (
|
||||
<figureCitation id="1315EAA14C31C6101AB42F29FB342369" box="[1172,1255,516,541]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="71" pageNumber="72">Fig. 18</figureCitation>
|
||||
<figureCitation id="1315EAA14C31C6101AB42F29FB342369" box="[1172,1255,516,541]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="71" pageNumber="72">Fig. 18</figureCitation>
|
||||
D),
|
||||
<collectingCountry id="F339B6B44C31C6101B352F29FA4F2369" box="[1301,1436,516,541]" name="Madagascar" pageId="71" pageNumber="72">Madagascar</collectingCountry>
|
||||
NW, Bekolosy, RS Manongarivo,
|
||||
|
|
@ -207,7 +207,7 @@ NW, Bekolosy, RS Manongarivo,
|
|||
<subSubSection id="C334A5AF4C31C6101EE72E01FCFF2760" pageId="71" pageNumber="72" type="diagnosis">
|
||||
<paragraph id="8B91F6244C31C6101EE72E01FCFF2760" blockId="71.[151,1437,258,2025]" pageId="71" pageNumber="72">
|
||||
<emphasis id="B95A2A364C31C6101EE72E01FE922231" bold="true" box="[199,321,812,837]" pageId="71" pageNumber="72">Diagnosis.</emphasis>
|
||||
<taxonomicName id="4C2E8DA74C31C6101F692E00FDC02231" box="[329,531,813,837]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="maeva">
|
||||
<taxonomicName id="4C2E8DA74C31C6101F692E00FDC02231" box="[329,531,813,837]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="maeva">
|
||||
<emphasis id="B95A2A364C31C6101F692E00FDC02231" box="[329,531,813,837]" italics="true" pageId="71" pageNumber="72">Heteropsis maeva</emphasis>
|
||||
</taxonomicName>
|
||||
is similar on the dorsal wing surface. In Asia, a group that includes
|
||||
|
|
@ -243,15 +243,15 @@ Mabille 1877
|
|||
: 186 for a photograph) is a synonym according to Lees
|
||||
<emphasis id="B95A2A364C31C6101AED2EB4FAC222C5" box="[1229,1297,920,945]" italics="true" pageId="71" pageNumber="72">et al.,</emphasis>
|
||||
(2003),
|
||||
<taxonomicName id="4C2E8DA74C31C6101B592EB4FEEE22A1" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<taxonomicName id="4C2E8DA74C31C6101B592EB4FEEE22A1" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<emphasis id="B95A2A364C31C6101B592EB4FEEE22A1" italics="true" pageId="71" pageNumber="72">Ht. menamenoides</emphasis>
|
||||
</taxonomicName>
|
||||
has a dark orange Mb and Smb ventrally, with contiguous pale yellow shading distad to the Mb, and dark orange somewhat triangular blotches distal to that. However, in
|
||||
<taxonomicName id="4C2E8DA74C31C6101DE22ECCFB45228D" box="[962,1174,992,1017]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<taxonomicName id="4C2E8DA74C31C6101DE22ECCFB45228D" box="[962,1174,992,1017]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<emphasis id="B95A2A364C31C6101DE22ECCFB45228D" box="[962,1174,992,1017]" italics="true" pageId="71" pageNumber="72">Ht. menamenoides</emphasis>
|
||||
</taxonomicName>
|
||||
, only the space-CuA1 ocellus is expressed as typical of the
|
||||
<taxonomicName id="4C2E8DA74C31C6101C6B292BFD1C2569" box="[587,719,1028,1053]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C31C6101C6B292BFD1C2569" box="[587,719,1028,1053]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C31C6101C6B292BFD1C2569" box="[587,719,1028,1053]" italics="true" pageId="71" pageNumber="72">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group, and the HW margin is less crenulated, whereas in the
|
||||
|
|
@ -261,21 +261,21 @@ group, and the HW margin is less crenulated, whereas in the
|
|||
group, a full complement of ocelli are expressed, especially on the HWV. Within the ‘true’
|
||||
<collectingCountry id="F339B6B44C31C6101EB72961FED72511" box="[151,260,1100,1125]" name="Madagascar" pageId="71" pageNumber="72">Malagasy</collectingCountry>
|
||||
mycalesine fauna, the upperside of
|
||||
<taxonomicName id="4C2E8DA74C31C6101C902963FC572511" box="[688,900,1100,1125]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<taxonomicName id="4C2E8DA74C31C6101C902963FC572511" box="[688,900,1100,1125]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<emphasis id="B95A2A364C31C6101C902963FC572511" box="[688,900,1100,1125]" italics="true" pageId="71" pageNumber="72">Ht. menamenoides</emphasis>
|
||||
</taxonomicName>
|
||||
is similar to that of
|
||||
<taxonomicName id="4C2E8DA74C31C6101AA02963FB242511" box="[1152,1271,1102,1125]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="maeva">
|
||||
<taxonomicName id="4C2E8DA74C31C6101AA02963FB242511" box="[1152,1271,1102,1125]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="maeva">
|
||||
<emphasis id="B95A2A364C31C6101AA02963FB242511" box="[1152,1271,1102,1125]" italics="true" pageId="71" pageNumber="72">Ht. maeva</emphasis>
|
||||
</taxonomicName>
|
||||
, to which the species is apparently closely related, and shares a smooth HW margin, but differs by a much lighter and more uneven orange colouration on both wing surfaces, whereas the HW-crenulate
|
||||
<taxonomicName id="4C2E8DA74C31C6101A2629BBFB4A25D9" box="[1030,1177,1172,1197]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C31C6101A2629BBFB4A25D9" box="[1030,1177,1172,1197]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C31C6101A2629BBFB4A25D9" box="[1030,1177,1172,1197]" italics="true" pageId="71" pageNumber="72">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
is not known from the northwest of
|
||||
<collectingCountry id="F339B6B44C31C6101F112995FE6925A5" box="[305,442,1208,1233]" name="Madagascar" pageId="71" pageNumber="72">Madagascar</collectingCountry>
|
||||
, and has a almost entirely mid brown dorsal wing colouration. Among the only other distinctly ‘orange’
|
||||
<taxonomicName id="4C2E8DA74C31C6101F5529F0FE3E2580" box="[373,493,1245,1268]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C31C6101F5529F0FE3E2580" box="[373,493,1245,1268]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C31C6101F5529F0FE3E2580" box="[373,493,1245,1268]" italics="true" pageId="71" pageNumber="72">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
(in both sexes) known from
|
||||
|
|
@ -283,7 +283,7 @@ is not known from the northwest of
|
|||
, the
|
||||
<collectingCountry id="F339B6B44C31C6101A3429F1FB522581" box="[1044,1153,1244,1269]" name="Madagascar" pageId="71" pageNumber="72">Malagasy</collectingCountry>
|
||||
Region complex of
|
||||
<taxonomicName id="4C2E8DA74C31C6101B5929F3FE18246D" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="narcissus">
|
||||
<taxonomicName id="4C2E8DA74C31C6101B5929F3FE18246D" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="narcissus">
|
||||
<emphasis id="B95A2A364C31C6101B5929F3FED3246C" italics="true" pageId="71" pageNumber="72">Ht. narcissus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C31C6101F30282DFE10246D" author="Fabricius" box="[272,451,1280,1305]" pageId="71" pageNumber="72" refString="ICZN (2015) Opinion 2359 (Case 3573): Papilio narcissus Fabricius, 1798 (currently Heteropsis narcissus; Lepidoptera, NYMPHALIDAE): specific name conserved. Bulletin of Zoological Nomenclature, 72 (2), 171 - 172." type="journal article" year="1798">Fabricius, 1798</bibRefCitation>
|
||||
|
|
@ -298,7 +298,7 @@ and
|
|||
<emphasis id="B95A2A364C31C6101B59282CFF1C2449" italics="true" pageId="71" pageNumber="72">Ht. laeta</emphasis>
|
||||
</taxonomicName>
|
||||
which are similar in dorsal wing pattern, have them, but are paler yellow on underside. Orange ♀ forms of
|
||||
<taxonomicName id="4C2E8DA74C31C6101B59280BFF3F2415" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="erebina">
|
||||
<taxonomicName id="4C2E8DA74C31C6101B59280BFF3F2415" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="erebina">
|
||||
<emphasis id="B95A2A364C31C6101B59280BFF3F2415" italics="true" pageId="71" pageNumber="72">Ht. erebina</emphasis>
|
||||
</taxonomicName>
|
||||
(‘
|
||||
|
|
@ -320,7 +320,7 @@ Mabille, 1884
|
|||
: 56 and
|
||||
<bibRefCitation id="EFBF8BD54C31C6101F1F2841FE2324F1" author="d'Abrera" box="[319,496,1388,1413]" pageId="71" pageNumber="72" refString="d'Abrera, B. L. (1997) Butterflies of the Afrotropical Region. Part 1. Papilionidae, Pieridae, Acraeidae, Danaidae, Satyridae. Hill House Publishers, Melbourne & London, 263 pp." type="book" year="1997">d’Abrera, 1997</bibRefCitation>
|
||||
) are also not confusable because the Mb shows through to the upperside more strongly and delineates an area just proximad of the space-CuA1 ocellus which is highlighted more contrastingly in paler orange.
|
||||
<taxonomicName id="4C2E8DA74C31C6101F14289BFE6624B9" box="[308,437,1460,1485]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="erebina">
|
||||
<taxonomicName id="4C2E8DA74C31C6101F14289BFE6624B9" box="[308,437,1460,1485]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="erebina">
|
||||
<emphasis id="B95A2A364C31C6101F14289BFE6624B9" box="[308,437,1460,1485]" italics="true" pageId="71" pageNumber="72">Ht. erebina</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -328,7 +328,7 @@ and
|
|||
<emphasis id="B95A2A364C31C6101FCE289BFDAD24B9" box="[494,638,1460,1485]" italics="true" pageId="71" pageNumber="72">Ht. antahala</emphasis>
|
||||
</taxonomicName>
|
||||
belong to different clades with different wing shapes and particularly ventral wing patterning. In particular, none of these species have such a distinctly zigzagged orange Sml, that is especially characteristic of the FWV of
|
||||
<taxonomicName id="4C2E8DA74C31C6101C7528D3FCF62761" box="[597,805,1532,1557]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<taxonomicName id="4C2E8DA74C31C6101C7528D3FCF62761" box="[597,805,1532,1557]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<emphasis id="B95A2A364C31C6101C7528D3FCF62761" box="[597,805,1532,1557]" italics="true" pageId="71" pageNumber="72">Ht. menamenoides</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
|
|
@ -339,7 +339,7 @@ belong to different clades with different wing shapes and particularly ventral w
|
|||
<emphasis id="B95A2A364C31C6101EE72B0DFE7F274D" bold="true" box="[199,428,1568,1593]" pageId="71" pageNumber="72">Description. Wings</emphasis>
|
||||
: Upperside orange with wide darkish brown border along margin that widens from costa towards apex in FWD and that is more diffuse on HWD, narrowest at margin and broader at costa and tergal edge. Lighter orange is evident around space-M3 of FWD and HWD. Space-CuA1 ocellus expressed more strongly in FWD than in HWD; Orng not evident and black iris considerably smaller diameter than spacing of veins CuA1 and CuA2. Underside generally of lighter orange cast especially distad of Mb of central symmetry system, and there is a diffuse darker orange band running through arc of ocelli and a very wavy darker orange line before another darker orange Sml that closely tracks the margin on both wings; the margin is fairly smoothly cut and only very gently crenate. The dark orange Mb is relatively straight in the HWV and curves towards the tergal angle of the FWV, tergad of the space-CuA1 ocellus, which is very small in the HT. In FWV, Cbs delineated by two sets of approximately parallel darker orange transverse lines in the FWV cell area, not as evident as in the (anyway brown) species
|
||||
<emphasis id="B95A2A364C31C6101ED22AA7FE4A26D5" box="[242,409,1930,1953]" italics="true" pageId="71" pageNumber="72">
|
||||
<taxonomicName id="4C2E8DA74C31C6101ED22AA7FE4626D5" box="[242,405,1930,1953]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="roussettae">Ht. roussettae</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C31C6101ED22AA7FE4626D5" box="[242,405,1930,1953]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="roussettae">Ht. roussettae</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
Space-M2 ocellus as small white point surrounded by narrow black ring in FWV while space-CuA1 ocellus of HWV only slightly larger; no other ocelli expressed. Basal area strongly strigulated with darker orange lines in both wings, and PMb is more evident in the HWV than FWV.
|
||||
|
|
@ -356,13 +356,13 @@ series were collected over a few adjacent days in mid December. ♀ unknown.
|
|||
<quantity id="4CD65BC14C3EC61F1FEC2C29FDE22068" box="[460,561,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="72" pageNumber="73" unit="mm" value="19.7">19.7 mm</quantity>
|
||||
. ♀ unknown.
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C3EC61F1EB72BC1FE6D2629" httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="72" pageNumber="73" targetBox="[290,1282,340,1732]" targetPageId="72">
|
||||
<caption id="DF51A6AC4C3EC61F1EB72BC1FE6D2629" ID-DOI="http://doi.org/10.5281/zenodo.264618" httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="72" pageNumber="73" targetBox="[290,1282,340,1732]" targetPageId="72">
|
||||
<paragraph id="8B91F6244C3EC61F1EB72BC1FE6D2629" blockId="72.[151,1436,1772,1885]" pageId="72" pageNumber="73">
|
||||
<emphasis id="B95A2A364C3EC61F1EB72BC1FEF52675" bold="true" box="[151,294,1772,1794]" pageId="72" pageNumber="73">FIGURE 21.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
<emphasis id="B95A2A364C3EC61F1C812BC1FD602675" bold="true" box="[673,691,1772,1793]" pageId="72" pageNumber="73">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C3EC61F1D172BC0FB552676" authority="Lees" authorityName="Lees" box="[823,1158,1772,1794]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="menamenoides" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C3EC61F1D172BC0FB552676" authority="Lees" authorityName="Lees" box="[823,1158,1772,1794]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="menamenoides" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C3EC61F1D172BC0FB942676" box="[823,1095,1772,1794]" italics="true" pageId="72" pageNumber="73">Heteropsis menamenoides</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -399,13 +399,13 @@ Mabille, 1877
|
|||
: black ventral abdominal patch between A3–A6 divided centrally by narrow yellow overlying scales, in potential contact with diffusely separated and slightly wavy androconial brush largely emanating from base of space-CuA2 before its fork with vein CuA1 and overlying veins 1A+2A and 3A; the latter symmetrically inflated before midlength in compact ‘tear’-shape (
|
||||
<bibRefCitation id="EFBF8BD54C3FC61E1CF42DB5FC0921C4" author="Lees" box="[724,986,151,176]" pageId="73" pageNumber="74" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="b">
|
||||
Lees, 1997: 97,
|
||||
<figureCitation id="1315EAA14C3FC61E1DA92DBAFC1921C4" box="[905,970,151,176]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="73" pageNumber="74">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C3FC61E1DA92DBAFC1921C4" box="[905,970,151,176]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="73" pageNumber="74">Fig. 3</figureCitation>
|
||||
b
|
||||
</bibRefCitation>
|
||||
), whereas 1A+2A shows very restricted swelling towards the base of 1A. Fairly short and compact Sdb brownish at base and strongly light yellow towards tip overlying small Sdp with small underlying swelling in HW vein R towards end of cell (
|
||||
<bibRefCitation id="EFBF8BD54C3FC61E1AA82DF2FA5D2183" author="Lees" box="[1160,1422,222,247]" pageId="73" pageNumber="74" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="b">
|
||||
Lees, 1997: 97,
|
||||
<figureCitation id="1315EAA14C3FC61E1B1C2DF3FAAE2183" box="[1340,1405,222,247]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="73" pageNumber="74">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C3FC61E1B1C2DF3FAAE2183" box="[1340,1405,222,247]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="73" pageNumber="74">Fig. 3</figureCitation>
|
||||
b
|
||||
</bibRefCitation>
|
||||
); not described in detail here due to scarcity of material.
|
||||
|
|
@ -417,9 +417,9 @@ b
|
|||
<paragraph id="8B91F6244C3FC61E1EE72C46FD702390" blockId="73.[151,1437,151,1690]" pageId="73" pageNumber="74">
|
||||
<emphasis id="B95A2A364C3FC61E1EE72C46FE9D20F0" bold="true" box="[199,334,363,388]" pageId="73" pageNumber="74">♂ genitalia</emphasis>
|
||||
: 231DL (DLBEK94_180;
|
||||
<figureCitation id="1315EAA14C3FC61E1CA82C46FD0820F7" box="[648,731,363,388]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="73" pageNumber="74">Fig. 18</figureCitation>
|
||||
<figureCitation id="1315EAA14C3FC61E1CA82C46FD0820F7" box="[648,731,363,388]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="73" pageNumber="74">Fig. 18</figureCitation>
|
||||
D, PT): from LV, tegumen broader dorsally and very angled proximad (scarcely any notch from DV), very narrow at hinge with vinculum; dorsal edge of tegumen forming a fairly straight line to uncus featuring fairly narrow and evenly deep ‘neck’ distad to hook-tip with distinct dorsal ‘head’ as fairly typical for
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1FC82CFBFDA22099" box="[488,625,468,493]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1FC82CFBFDA22099" box="[488,625,468,493]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C3FC61E1FC82CFBFDA22099" box="[488,625,468,493]" italics="true" pageId="73" pageNumber="74">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group; uncus quite strongly inflated before tip from DV. Gnathos particularly straight and tapering (but slightly sinuate from DV and inrecurved at tip), emanating from rounded base and in the specimen examined, transversing uncus at about 40 degrees. Valve with a long straight dorsal ‘shoulder’. Valve base leads without strong constriction to quite broad and flat valve arm covered in spinoid setae along entirety of fairly truncate tip, featuring a slight distal lobe (incurved from DV), with uncus hook-tip protruding to middle of extension of this truncate tip (from LV). Saccus not very long and aedeagus about same length as valve, strongly uprecurved from midlength and featuring a long proximad ostium, slightly bulbous at tip. Juxta somewhat ‘plate-lipped’ at proximad tip.
|
||||
|
|
@ -464,11 +464,11 @@ at Bekolosy mountain in the RS Manongarivo (
|
|||
of the ‘orange’ species in
|
||||
<collectingCountry id="F339B6B44C3FC61E1AD82EB3FAAC22C3" box="[1272,1407,926,951]" name="Madagascar" pageId="73" pageNumber="74">Madagascar</collectingCountry>
|
||||
in the
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1EE12EEFFE9022AE" box="[193,323,961,986]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1EE12EEFFE9022AE" box="[193,323,961,986]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C3FC61E1EE12EEFFE9022AE" box="[193,323,961,986]" italics="true" pageId="73" pageNumber="74">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group were examined (see under
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1CE42EEFFC8422AE" box="[708,855,961,986]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1CE42EEFFC8422AE" box="[708,855,961,986]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C3FC61E1CE42EEFFC8422AE" box="[708,855,961,986]" italics="true" pageId="73" pageNumber="74">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
), as well as the ♀ HT of
|
||||
|
|
@ -487,7 +487,7 @@ E. & B. Perrot|
|
|||
<paragraph id="8B91F6244C3FC61E1EE72906FEA625D9" blockId="73.[151,1437,151,1690]" pageId="73" pageNumber="74">
|
||||
<emphasis id="B95A2A364C3FC61E1EE72906FD252530" bold="true" box="[199,758,1067,1092]" pageId="73" pageNumber="74">Additional information. DNA divergences:</emphasis>
|
||||
COI-5P cluster number BOLD:AAE4113 (exemplar DLBEK94_179, BMAD075-09, HM404251). The DNA barcode is about 5.5% divergent to an undescribed species, sp. 28 (cluster number BOLD:AAE5458), and about 5.8% pairwise divergent to that of
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1B05295FFA4F25FD" box="[1317,1436,1138,1161]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1B05295FFA4F25FD" box="[1317,1436,1138,1161]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C3FC61E1B05295FFA4F25FD" box="[1317,1436,1138,1161]" italics="true" pageId="73" pageNumber="74">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
(BOLD:AAE4112).
|
||||
|
|
@ -528,7 +528,7 @@ A. Camus, not necessarily the hostplant.
|
|||
<paragraph id="8B91F6244C3FC61E1EE72883FEDD249E" blockId="73.[151,1437,151,1690]" pageId="73" pageNumber="74">
|
||||
<emphasis id="B95A2A364C3FC61E1EE72883FE8A24B3" bold="true" box="[199,345,1454,1479]" pageId="73" pageNumber="74">Distribution</emphasis>
|
||||
: endemic as far as is known to Mt. Bekolosy, Reserve Speciale de Manongarivo (
|
||||
<figureCitation id="1315EAA14C3FC61E1ACE2883FA9324B3" box="[1262,1344,1454,1479]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="73" pageNumber="74">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C3FC61E1ACE2883FA9324B3" box="[1262,1344,1454,1479]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="73" pageNumber="74">Fig. 30</figureCitation>
|
||||
C, slateblue dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C3FC61E1EE728D9FC852779" blockId="73.[151,1437,151,1690]" box="[199,854,1524,1549]" pageId="73" pageNumber="74">
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<treatment id="038747324C34C6111EB72C8FFDC9260E" ID-DOI="http://doi.org/10.5281/zenodo.6086448" ID-GBIF-Taxon="120817147" ID-Zenodo-Dep="6086448" LSID="urn:lsid:plazi:treatment:038747324C34C6111EB72C8FFDC9260E" httpUri="http://treatment.plazi.org/id/038747324C34C6111EB72C8FFDC9260E" lastPageId="70" lastPageNumber="70" pageId="66" pageNumber="67">
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||||
<treatment id="038747324C34C6111EB72C8FFDC9260E" ID-DOI="http://doi.org/10.5281/zenodo.6086448" ID-GBIF-Taxon="120817147" ID-Zenodo-Dep="6086448" LSID="urn:lsid:plazi:treatment:038747324C34C6111EB72C8FFDC9260E" httpUri="http://treatment.plazi.org/id/038747324C34C6111EB72C8FFDC9260E" lastPageId="70" lastPageNumber="70" pageId="66" pageNumber="67" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C34C6151EB72C8FFBC120AA" pageId="66" pageNumber="67" type="nomenclature">
|
||||
<paragraph id="8B91F6244C34C6151EB72C8FFD1620C9" blockId="66.[151,1042,418,478]" box="[151,709,418,445]" pageId="66" pageNumber="67">
|
||||
<heading id="D0D941484C34C6151EB72C8FFD1620C9" bold="true" box="[151,709,418,445]" fontSize="11" level="1" pageId="66" pageNumber="67" reason="1">
|
||||
<emphasis id="B95A2A364C34C6151EB72C8FFD1620C9" bold="true" box="[151,709,418,445]" pageId="66" pageNumber="67">
|
||||
<taxonomicName id="4C2E8DA74C34C6151EB72C8FFD8820C9" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,603,418,445]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C34C6151EB72C8FFD8820C9" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,603,418,445]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C34C6151EB72C8FFE5F20C8" bold="true" box="[151,396,418,444]" italics="true" pageId="66" pageNumber="67">Heteropsis barbarae</emphasis>
|
||||
Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -76,7 +76,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C34C6151CA62F01FD202331" box="[646,755,556,581]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C34C6151D102F01FC552331" box="[816,902,556,581]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
<figureCitation id="1315EAA14C34C6151D102F01FC552331" box="[816,902,556,581]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." figureDoi="http://doi.org/10.5281/zenodo.264617" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C34C6151D962F01FBEE2331" box="[950,1085,556,581]" name="Madagascar" pageId="66" pageNumber="67">Madagascar</collectingCountry>
|
||||
NE, Maitsoarongana, Sikiory River, near camp,
|
||||
|
|
@ -163,7 +163,7 @@ barcode]; ♂, NE, Maitsoarongana, ridge just S. of camp,
|
|||
(E) 1717099 [
|
||||
<collectingCountry id="F339B6B44C34C6151B412E0AFA4F224B" box="[1377,1436,807,831]" name="Qatar" pageId="66" pageNumber="67">QTR</collectingCountry>
|
||||
barcode]; ♀ (
|
||||
<figureCitation id="1315EAA14C34C6151F0C2E61FE532210" box="[300,384,844,869]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
<figureCitation id="1315EAA14C34C6151F0C2E61FE532210" box="[300,384,844,869]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." figureDoi="http://doi.org/10.5281/zenodo.264617" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
B),
|
||||
<collectingCountry id="F339B6B44C34C6151F872E61FDFD2211" box="[423,558,844,869]" name="Madagascar" pageId="66" pageNumber="67">Madagascar</collectingCountry>
|
||||
NE, Sikiory R., riparian,
|
||||
|
|
@ -241,86 +241,86 @@ NE, Maitsoarongana, Sikiory River,
|
|||
<paragraph id="8B91F6244C34C6151EE7296DFC632743" blockId="66.[151,1437,522,2035]" pageId="66" pageNumber="67">
|
||||
<emphasis id="B95A2A364C34C6151EE7296DFE92252D" bold="true" box="[199,321,1088,1113]" pageId="66" pageNumber="67">Diagnosis.</emphasis>
|
||||
The ventral wings of
|
||||
<taxonomicName id="4C2E8DA74C34C6151C17296CFD1A252D" box="[567,713,1088,1113]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C34C6151C17296CFD1A252D" box="[567,713,1088,1113]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C34C6151C17296CFD1A252D" box="[567,713,1088,1113]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
are similar to some other members of the
|
||||
<taxonomicName id="4C2E8DA74C34C6151A82296CFAF0252D" box="[1186,1315,1088,1113]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C34C6151A82296CFAF0252D" box="[1186,1315,1088,1113]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C34C6151A82296CFAF0252D" box="[1186,1315,1088,1113]" italics="true" pageId="66" pageNumber="67">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group that exhibit a crenulate HW margin, notably
|
||||
<taxonomicName id="4C2E8DA74C34C6151C582949FC9E250F" box="[632,845,1122,1147]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="menamenoides" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C34C6151C582949FC9E250F" box="[632,845,1122,1147]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="menamenoides" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C34C6151C582949FC9E250F" box="[632,845,1122,1147]" italics="true" pageId="66" pageNumber="67">Ht. menamenoides</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C34C6151D79294FFC64250F" box="[857,951,1122,1147]" pageId="66" pageNumber="67" rank="species">
|
||||
<emphasis id="B95A2A364C34C6151D79294FFC64250F" bold="true" box="[857,951,1122,1147]" pageId="66" pageNumber="67">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C34C6151DD72949FBAE250F" box="[1015,1149,1122,1147]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C34C6151DD72949FBAE250F" box="[1015,1149,1122,1147]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C34C6151DD72949FBAE250F" box="[1015,1149,1122,1147]" italics="true" pageId="66" pageNumber="67">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
. Upperside dark brown, lacking light orange wash and smooth HW margin of
|
||||
<taxonomicName id="4C2E8DA74C34C6151CD429ABFC1625E9" box="[756,965,1156,1181]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<taxonomicName id="4C2E8DA74C34C6151CD429ABFC1625E9" box="[756,965,1156,1181]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<emphasis id="B95A2A364C34C6151CD429ABFC1625E9" box="[756,965,1156,1181]" italics="true" pageId="66" pageNumber="67">Ht. menamenoides</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C34C6151DF529A9FBF625E9" box="[981,1061,1156,1181]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="66" pageNumber="67">Fig. 21</figureCitation>
|
||||
<figureCitation id="1315EAA14C34C6151DF529A9FBF625E9" box="[981,1061,1156,1181]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264618" httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="66" pageNumber="67">Fig. 21</figureCitation>
|
||||
A) and
|
||||
<taxonomicName id="4C2E8DA74C34C6151A5B29ABFB3D25E9" box="[1147,1262,1158,1181]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="maeva">
|
||||
<emphasis id="B95A2A364C34C6151A5B29ABFB3D25E9" box="[1147,1262,1158,1181]" italics="true" pageId="66" pageNumber="67">Ht. maeva</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C34C6151ADE29A9FA9C25E9" box="[1278,1359,1156,1181]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="66" pageNumber="67">Fig. 21</figureCitation>
|
||||
<figureCitation id="1315EAA14C34C6151ADE29A9FA9C25E9" box="[1278,1359,1156,1181]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264618" httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="66" pageNumber="67">Fig. 21</figureCitation>
|
||||
D) and the HWD orange crescent (Mf) typical of
|
||||
<taxonomicName id="4C2E8DA74C34C6151CA22985FCD625CB" box="[642,773,1190,1215]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C34C6151CA22985FCD625CB" box="[642,773,1190,1215]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C34C6151CA22985FCD625CB" box="[642,773,1190,1215]" italics="true" pageId="66" pageNumber="67">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
. Fresh specimens of
|
||||
<taxonomicName id="4C2E8DA74C34C6151DDF2985FB4725CB" box="[1023,1172,1190,1215]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C34C6151DDF2985FB4725CB" box="[1023,1172,1190,1215]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C34C6151DDF2985FB4725CB" box="[1023,1172,1190,1215]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
are distinctive in their strong slightly dingy orange ventral cast (
|
||||
<figureCitation id="1315EAA14C34C6151C4629E4FD6A2595" box="[614,697,1225,1250]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
<figureCitation id="1315EAA14C34C6151C4629E4FD6A2595" box="[614,697,1225,1250]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." figureDoi="http://doi.org/10.5281/zenodo.264617" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
A,E), and the ventral scales are easily displaced/rubbed. Worn ♀♀ can easily be confused with those of often sympatric
|
||||
<taxonomicName id="4C2E8DA74C34C6151CCB29C1FCB72477" box="[747,868,1260,1283]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="pauper">
|
||||
<emphasis id="B95A2A364C34C6151CCB29C1FCB72477" box="[747,868,1260,1283]" italics="true" pageId="66" pageNumber="67">Ht. pauper</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C34C6151DBB29C1FBFF2470" box="[923,1068,1259,1284]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<taxonomicName id="4C2E8DA74C34C6151DBB29C1FBFF2470" box="[923,1068,1259,1284]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<emphasis id="B95A2A364C34C6151DBB29C1FBFF2470" box="[923,1068,1259,1284]" italics="true" pageId="66" pageNumber="67">Ht. undulans</emphasis>
|
||||
</taxonomicName>
|
||||
. Indeed the ♀ of
|
||||
<taxonomicName id="4C2E8DA74C34C6151AD129C1FA522470" box="[1265,1409,1259,1284]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<taxonomicName id="4C2E8DA74C34C6151AD129C1FA522470" box="[1265,1409,1259,1284]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<emphasis id="B95A2A364C34C6151AD129C1FA522470" box="[1265,1409,1259,1284]" italics="true" pageId="66" pageNumber="67">Ht. undulans</emphasis>
|
||||
</taxonomicName>
|
||||
is sometimes almost indistinguishable (
|
||||
<figureCitation id="1315EAA14C34C6151C172820FD5A2452" box="[567,649,1293,1318]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
<figureCitation id="1315EAA14C34C6151C172820FD5A2452" box="[567,649,1293,1318]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." figureDoi="http://doi.org/10.5281/zenodo.264617" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
D) to that of
|
||||
<taxonomicName id="4C2E8DA74C34C6151D392823FC7F2452" box="[793,940,1293,1318]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C34C6151D392823FC7F2452" box="[793,940,1293,1318]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C34C6151D392823FC7F2452" box="[793,940,1293,1318]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C34C6151D9B2820FBDC2452" box="[955,1039,1293,1318]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
<figureCitation id="1315EAA14C34C6151D9B2820FBDC2452" box="[955,1039,1293,1318]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." figureDoi="http://doi.org/10.5281/zenodo.264617" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
B), but
|
||||
<taxonomicName id="4C2E8DA74C34C6151A412823FB202452" box="[1121,1267,1293,1318]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<taxonomicName id="4C2E8DA74C34C6151A412823FB202452" box="[1121,1267,1293,1318]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<emphasis id="B95A2A364C34C6151A412823FB202452" box="[1121,1267,1293,1318]" italics="true" pageId="66" pageNumber="67">Ht. undulans</emphasis>
|
||||
</taxonomicName>
|
||||
♀♀ often tend to be more flushed with reddish scales on the upperside, while
|
||||
<taxonomicName id="4C2E8DA74C34C6151D87281DFB93243C" box="[935,1088,1327,1352]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C34C6151D87281DFB93243C" box="[935,1088,1327,1352]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C34C6151D87281DFB93243C" box="[935,1088,1327,1352]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
can be distinguished by the combination of rich and dense slightly dingy orange scales on the ventral surface which lack a yellow cast (such as in
|
||||
<taxonomicName id="4C2E8DA74C34C6151E942858FEE724F9" box="[180,308,1396,1421]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C34C6151E942858FEE724F9" box="[180,308,1396,1421]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C34C6151E942858FEE724F9" box="[180,308,1396,1421]" italics="true" pageId="66" pageNumber="67">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
), combined with a less contrasting, pale orange band between Pml and Sml (
|
||||
<figureCitation id="1315EAA14C34C6151AB22859FB3624F8" box="[1170,1253,1396,1421]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
<figureCitation id="1315EAA14C34C6151AB22859FB3624F8" box="[1170,1253,1396,1421]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." figureDoi="http://doi.org/10.5281/zenodo.264617" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
|
||||
C,D,F), which is pale ochreous in both sexes of some populations of
|
||||
<taxonomicName id="4C2E8DA74C34C6151CF528BAFCB524DA" box="[725,870,1429,1454]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<taxonomicName id="4C2E8DA74C34C6151CF528BAFCB524DA" box="[725,870,1429,1454]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<emphasis id="B95A2A364C34C6151CF528BAFCB524DA" box="[725,870,1429,1454]" italics="true" pageId="66" pageNumber="67">Ht. undulans</emphasis>
|
||||
</taxonomicName>
|
||||
(however that is a very variable species), but there will always be a few especially more worn specimens of
|
||||
<taxonomicName id="4C2E8DA74C34C6151D302894FC7124A5" box="[784,930,1464,1489]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C34C6151D302894FC7124A5" box="[784,930,1464,1489]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C34C6151D302894FC7124A5" box="[784,930,1464,1489]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
that are hard to place without dissection. The similarly lowland
|
||||
|
|
@ -328,9 +328,9 @@ that are hard to place without dissection. The similarly lowland
|
|||
<emphasis id="B95A2A364C34C6151F4228F6FE062486" box="[354,469,1499,1522]" italics="true" pageId="66" pageNumber="67">Ht. maeva</emphasis>
|
||||
</taxonomicName>
|
||||
has a brighter orange underside (
|
||||
<figureCitation id="1315EAA14C34C6151D6928F7FC492487" box="[841,922,1498,1523]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="66" pageNumber="67">Fig. 21</figureCitation>
|
||||
<figureCitation id="1315EAA14C34C6151D6928F7FC492487" box="[841,922,1498,1523]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264618" httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="66" pageNumber="67">Fig. 21</figureCitation>
|
||||
D), denser black ventral brush in the ♂, and is distinctive in that its upperside is orange too, except for HWs of western specimens which are usually suffused with brown (but
|
||||
<taxonomicName id="4C2E8DA74C34C6151F392B32FE7F2743" box="[281,428,1566,1591]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C34C6151F392B32FE7F2743" box="[281,428,1566,1591]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C34C6151F392B32FE7F2743" box="[281,428,1566,1591]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
is anyway absent in the west of
|
||||
|
|
@ -342,22 +342,22 @@ is anyway absent in the west of
|
|||
<paragraph id="8B91F6244C34C6131EE72B6DFEC020A4" blockId="66.[151,1437,522,2035]" lastBlockId="68.[151,1437,151,2013]" lastPageId="68" lastPageNumber="69" pageId="66" pageNumber="67">
|
||||
<emphasis id="B95A2A364C34C6151EE72B6DFE7C272D" bold="true" box="[199,431,1600,1625]" pageId="66" pageNumber="67">Description. Wings</emphasis>
|
||||
: Upperside mid-brown with a slight russet brown cast. Space-CuA1 ocellus expressed strongly in FWD; orange ring sub-hexagonal., M1 ocellus very small black white pupilled spot surrounded by a narrow orange ring. HWD ocellus CuA1 fairly small and sub-elliptic, surrounded by a narrow orange ring. Underside has a lighter dingy orange cast, only slightly lighter distad of the Mb. Mb darker orange-brown, gently convex and slightly irregular (relatively straight in the HT HWV) with paler orange highlights distal to it in space- M2 to CuA1. FWV space-CuA1 ocellus similar size to FWD (in the HT), as is FWV M1 ocellus, which is reduced to a very small white point surrounded by a narrow black ring and orange ring that slightly contrasts with background. FWV space-CuA1 ocellus follows concave shape of the FWV darker orange-brown Mb before it bends back toward mid costa, with a ring that is orange near the black iris (which is significantly smaller than on FWD) and is yellowy-orange towards the greatest inflection of the bend in the Mb, forming a ‘spiral arm’ (Ore). HWV space-CuA1 ocellus similar size and shape to that on dorsal surface, surrounded by pale orange ring not much lighter than background; no other ocelli conspicuously expressed on the HWV nor HWD. The HWV basal area is sparsely irrorated with brown strigulae, the area between PMb and Mb more evenly and finely irrorated. A wavy-zigzag darker diffuse brownish line inset from margin follows a double narrow Sml divided by the ground colour, not very ochreous as usual in
|
||||
<taxonomicName id="4C2E8DA74C32C6131C1D2D90FD1D21A1" box="[573,718,188,213]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<taxonomicName id="4C2E8DA74C32C6131C1D2D90FD1D21A1" box="[573,718,188,213]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<emphasis id="B95A2A364C32C6131C1D2D90FD1D21A1" box="[573,718,188,213]" italics="true" pageId="68" pageNumber="69">Ht. undulans</emphasis>
|
||||
</taxonomicName>
|
||||
) that closely tracks the margin of both wings, the margin being relatively smooth in the FW and distinctly crenulate in the HW. The reddish/dark orange-brown HWV PMb is slightly more convex than the Mb and both lines terminate abruptly at 1A. In FWV in the cell area are four dark orange-brown Cbs meeting almost perpendicular at the costa, the outermost pair delineating relatively paler areas, the outermost of which is more convex than the others, with its borders diverging towards the costa (in the HT).
|
||||
<emphasis id="B95A2A364C32C6131EB72C42FED820FC" bold="true" box="[151,267,367,392]" pageId="68" pageNumber="69">Variation.</emphasis>
|
||||
♀♀ similar but larger and slightly paler (
|
||||
<figureCitation id="1315EAA14C32C6131CFB2C42FCFC20FC" box="[731,815,367,392]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="68" pageNumber="69">Fig. 20</figureCitation>
|
||||
<figureCitation id="1315EAA14C32C6131CFB2C42FCFC20FC" box="[731,815,367,392]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." figureDoi="http://doi.org/10.5281/zenodo.264617" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="68" pageNumber="69">Fig. 20</figureCitation>
|
||||
B). Space-CuA1 ocellus FWD is a little wider proximad in some specimens, and FWD M1 ocellus sometimes not visible. HW space-CuA1 ocellus is more circular in some specimens.
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C35C6141EB72A69FE7E26A1" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="67" pageNumber="68" targetBox="[278,1309,211,1838]" targetPageId="67">
|
||||
<caption id="DF51A6AC4C35C6141EB72A69FE7E26A1" ID-DOI="http://doi.org/10.5281/zenodo.264617" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="67" pageNumber="68" targetBox="[278,1309,211,1838]" targetPageId="67">
|
||||
<paragraph id="8B91F6244C35C6141EB72A69FE7E26A1" blockId="67.[151,1436,1860,2005]" pageId="67" pageNumber="68">
|
||||
<emphasis id="B95A2A364C35C6141EB72A69FEF1262D" bold="true" box="[151,290,1860,1882]" pageId="67" pageNumber="68">FIGURE 20.</emphasis>
|
||||
Adults in DV (left), VV (right) for mounted specimens and adults in natural posture.
|
||||
<emphasis id="B95A2A364C35C6141A892A69FB68262D" bold="true" box="[1193,1211,1860,1881]" pageId="67" pageNumber="68">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C35C6141B0F2A68FE7B260D" authority="Lees & Kremen" authorityName="Lees & Kremen" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C35C6141B0F2A68FE7B260D" authority="Lees & Kremen" authorityName="Lees & Kremen" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C35C6141B0F2A68FA4F262E" box="[1327,1436,1861,1882]" italics="true" pageId="67" pageNumber="68">Heteropsis</emphasis>
|
||||
<emphasis id="B95A2A364C35C6141EB72A4EFF26260D" box="[151,245,1891,1913]" italics="true" pageId="67" pageNumber="68">barbarae</emphasis>
|
||||
Lees & Kremen
|
||||
|
|
@ -369,7 +369,7 @@ Lees & Kremen
|
|||
(Maitsoarongana, Makira; BMNH(E) 1717096).
|
||||
<emphasis id="B95A2A364C35C6141A2E2A4EFBCC260C" bold="true" box="[1038,1055,1891,1912]" pageId="67" pageNumber="68">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C35C6141AB12A49FACB260D" box="[1169,1304,1891,1913]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C35C6141AB12A49FACB260D" box="[1169,1304,1891,1913]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C35C6141AB12A49FACB260D" box="[1169,1304,1891,1913]" italics="true" pageId="67" pageNumber="68">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
(Sikiory R., Makira; BMNH(E) 1717100).
|
||||
|
|
@ -388,7 +388,7 @@ Oberthür, 1916
|
|||
(Antanandava, PN Andohahela).
|
||||
<emphasis id="B95A2A364C35C6141FC62A8DFE2426C1" bold="true" box="[486,503,1952,1973]" pageId="67" pageNumber="68">E</emphasis>
|
||||
: ♀ of
|
||||
<taxonomicName id="4C2E8DA74C35C6141C182A8CFD6826C2" box="[568,699,1952,1974]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C35C6141C182A8CFD6826C2" box="[568,699,1952,1974]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C35C6141C182A8CFD6826C2" box="[568,699,1952,1974]" italics="true" pageId="67" pageNumber="68">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
(Makira; 27/01/2003).
|
||||
|
|
@ -441,11 +441,11 @@ by Claire Kremen in the survey of what is now Masoala National Park (
|
|||
, so obviously completely overlooked in historic collecting in the region. All available
|
||||
<typeStatus id="549548864C32C6131ACF2930FAF92541" box="[1263,1322,1053,1077]" pageId="68" pageNumber="69">types</typeStatus>
|
||||
in the
|
||||
<taxonomicName id="4C2E8DA74C32C6131B592930FED4252C" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C32C6131B592930FED4252C" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C32C6131B592930FED4252C" italics="true" pageId="68" pageNumber="69">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C32C6131F65296CFE19252C" box="[325,458,1087,1112]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C32C6131F65296CFE19252C" box="[325,458,1087,1112]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C32C6131F65296CFE19252C" box="[325,458,1087,1112]" italics="true" pageId="68" pageNumber="69">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
groups (all in BMNH) were examined. In particular, among the ‘orange underside’ species, the following specimens are typified. These include two of up to three (although in his original description, Mabille does not state the number of specimens; but see also Mabille [1887]: 77) possible ♂♂ STs in BMNH of
|
||||
|
|
@ -454,7 +454,7 @@ groups (all in BMNH) were examined. In particular, among the ‘orange underside
|
|||
<bibRefCitation id="EFBF8BD54C32C6131FB62981FD8125B0" author="Mabille" box="[406,594,1196,1221]" pageId="68" pageNumber="69" refString="Mabille, P. (1878) Lepidoptera Africana. Bulletin de la Societe Zoologique de France, 3, 81 - 95." type="journal article" year="1878">Mabille, 1878</bibRefCitation>
|
||||
</taxonomicName>
|
||||
; LT ♂, here designated (
|
||||
<figureCitation id="1315EAA14C32C6131DD12981FB8C25B0" box="[1009,1119,1196,1221]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="68" pageNumber="69">Fig. 21</figureCitation>
|
||||
<figureCitation id="1315EAA14C32C6131DD12981FB8C25B0" box="[1009,1119,1196,1221]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264618" httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="68" pageNumber="69">Fig. 21</figureCitation>
|
||||
D): BMNH(E) 673788;
|
||||
<typeStatus id="549548864C32C6131EB729E2FED9259C" box="[151,266,1231,1256]" pageId="68" pageNumber="69" type="lectotype">Lectotype</typeStatus>
|
||||
|MadagascarNo10 [?sic; or ‘MadagascarNW’?]|Ex Coll. CHRIS WARD|Ex Oberthür Coll. Brit. Mus. 1927-3.|
|
||||
|
|
@ -471,7 +471,7 @@ D): BMNH(E) 673788;
|
|||
<bibRefCitation id="EFBF8BD54C32C6131D952811FBB62421" author="Oberthur" box="[949,1125,1340,1365]" pageId="68" pageNumber="69" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, a LT ♂ is here designated (
|
||||
<figureCitation id="1315EAA14C32C6131EBF2872FF20240C" box="[159,243,1375,1400]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="68" pageNumber="69">Fig. 21</figureCitation>
|
||||
<figureCitation id="1315EAA14C32C6131EBF2872FF20240C" box="[159,243,1375,1400]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264618" httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="68" pageNumber="69">Fig. 21</figureCitation>
|
||||
B): the specimen bearing labels “BMNH(E) #674741;
|
||||
<typeStatus id="549548864C32C6131D7E2872FC1C240C" box="[862,975,1375,1400]" pageId="68" pageNumber="69" type="lectotype">Lectotype</typeStatus>
|
||||
|Nord-Madagascar (Antakares) Isokitra a Diego Suarez
|
||||
|
|
@ -494,7 +494,7 @@ are numbered BMNH(E) #674743-674746. Among STs of
|
|||
<bibRefCitation id="EFBF8BD54C32C6131FA12B39FDFC2759" author="Oberthur" box="[385,559,1556,1581]" pageId="68" pageNumber="69" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, a LT ♂ in BMNH is here designated (
|
||||
<figureCitation id="1315EAA14C32C6131DC72B39FBE92758" box="[999,1082,1556,1581]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="68" pageNumber="69">Fig. 21</figureCitation>
|
||||
<figureCitation id="1315EAA14C32C6131DC72B39FBE92758" box="[999,1082,1556,1581]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, (‘ Antakares’; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 (‘ Antakares’; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 (‘ MadagascarNW’; BMNH (E) # 673788). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264618" httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="68" pageNumber="69">Fig. 21</figureCitation>
|
||||
C), the specimen bearing labels “LT|Nord-Madagascar (Antakares) Isokitra à Diego-Suarez
|
||||
<date id="FF90D0E44C32C6131D1E2B1AFBF02724" box="[830,1059,1591,1616]" pageId="68" pageNumber="69" value="1891-05" valueMax="1891-10-31" valueMin="1891-05-00">Mai à Octobre 1891</date>
|
||||
E.&B. Perrot|
|
||||
|
|
@ -537,7 +537,7 @@ were also examined; the LT ♂ is here considered to have designated by
|
|||
: 184 by the text “♂ (
|
||||
<typeStatus id="549548864C32C6131CF72BC1FCEF2671" box="[727,828,1772,1797]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
|
||||
) and ♀ as illustrated”, as illustrated on p. 185; the LT (
|
||||
<figureCitation id="1315EAA14C32C6131EBF2A22FF22265C" box="[159,241,1807,1832]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="68" pageNumber="69">Fig. 20</figureCitation>
|
||||
<figureCitation id="1315EAA14C32C6131EBF2A22FF22265C" box="[159,241,1807,1832]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees & Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." figureDoi="http://doi.org/10.5281/zenodo.264617" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="68" pageNumber="69">Fig. 20</figureCitation>
|
||||
C) bears the labels “
|
||||
<collectingCountry id="F339B6B44C32C6131FF32A22FD8E265C" box="[467,605,1807,1832]" name="Madagascar" pageId="68" pageNumber="69">Madagascar</collectingCountry>
|
||||
Fito
|
||||
|
|
@ -581,7 +581,7 @@ but is misidentified and in fact represents a ♀ of
|
|||
<emphasis id="B95A2A364C33C6121B232DB4FA4021C4" box="[1283,1427,151,176]" italics="true" pageId="69" pageNumber="70">Ht. undulans</emphasis>
|
||||
</taxonomicName>
|
||||
) also would belong. The last form (represented by a ♀ without abdomen;
|
||||
<figureCitation id="1315EAA14C33C6121DE52D91FBC721A0" box="[965,1044,188,213]" captionStart="FIGURE 14" captionStartId="48.[151,250,1802,1824]" captionTargetBox="[299,1270,199,1771]" captionTargetId="figure@48.[287,1299,193,1781]" captionTargetPageId="48" captionText="FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="69" pageNumber="70">Fig. 14</figureCitation>
|
||||
<figureCitation id="1315EAA14C33C6121DE52D91FBC721A0" box="[965,1044,188,213]" captionStart="FIGURE 14" captionStartId="48.[151,250,1802,1824]" captionTargetBox="[299,1270,199,1771]" captionTargetId="figure@48.[287,1299,193,1781]" captionTargetPageId="48" captionText="FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264611" httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="69" pageNumber="70">Fig. 14</figureCitation>
|
||||
E) is here designated LT of
|
||||
<taxonomicName id="4C2E8DA74C33C6121B692D91FE9C218C" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pseudonarcissus">
|
||||
<emphasis id="B95A2A364C33C6121B692D91FE9C218C" italics="true" pageId="69" pageNumber="70">Culapa pseudonarcissus</emphasis>
|
||||
|
|
@ -609,13 +609,13 @@ also captioned it “
|
|||
<paragraph id="8B91F6244C33C6121EE72C42FB472384" blockId="69.[151,1436,151,2013]" pageId="69" pageNumber="70">
|
||||
<emphasis id="B95A2A364C33C6121EE72C42FDA220FC" bold="true" box="[199,625,367,392]" pageId="69" pageNumber="70">Additional information. ♂ genitalia</emphasis>
|
||||
: 141DL (
|
||||
<figureCitation id="1315EAA14C33C6121CC12C42FCE120FC" box="[737,818,367,392]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="69" pageNumber="70">Fig. 18</figureCitation>
|
||||
<figureCitation id="1315EAA14C33C6121CC12C42FCE120FC" box="[737,818,367,392]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="69" pageNumber="70">Fig. 18</figureCitation>
|
||||
C, referred specimen): the basic configuration is fairly typical for
|
||||
<taxonomicName id="4C2E8DA74C33C6121F382CB8FE4F20D9" box="[280,412,404,429]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C33C6121F382CB8FE4F20D9" box="[280,412,404,429]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C33C6121F382CB8FE4F20D9" box="[280,412,404,429]" italics="true" pageId="69" pageNumber="70">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group. From LV, the approximately parallel-sided uncus is very slightly longer than the tegumen, the dorsal margin approximately in line with that of the tegumen and slightly upraised after its brow, with a smoothly round ‘head’ before the ventral hook (inflated from DV before tip), which projects forward rather than down. The gnathos originates on an elliptic base and is gently tapered and straight to an approx. 35-degree angle to uncus in the specimen examined; from DV it features a typical ‘bull’s horn’ sinuate appearance with pointed tips inrecurved. Only a slight constriction is found at hinge with vinculum. The valve has a fairly parallel-sided, flattened arm which is bent over at tip (inwards from DV) exposing a dense coverage of spinoid setae and the uncus tip when not downflexed is about the same extension as, not proud of the valve tips. The saccus is of normal length for the
|
||||
<taxonomicName id="4C2E8DA74C33C6121EC72F98FEB423B9" box="[231,359,692,717]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C33C6121EC72F98FEB423B9" box="[231,359,692,717]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C33C6121EC72F98FEB423B9" box="[231,359,692,717]" italics="true" pageId="69" pageNumber="70">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group and the aedeagus is about the same length as the valve and strongly recurved distad of the (also proximally uprecurved) ostium. The juxta is narrow and not sharply lipped proximad.
|
||||
|
|
@ -625,7 +625,7 @@ group and the aedeagus is about the same length as the valve and strongly recurv
|
|||
<paragraph id="8B91F6244C33C6121EE72FD1FBC822D1" blockId="69.[151,1436,151,2013]" pageId="69" pageNumber="70">
|
||||
<emphasis id="B95A2A364C33C6121EE72FD1FE7A2261" bold="true" box="[199,425,764,789]" pageId="69" pageNumber="70">DNA divergences:</emphasis>
|
||||
COI-5P cluster number BOLD:ACW4995 (exemplar BMAD248-15, DL14M-0033, Marojejy) is 4.86% divergent to
|
||||
<taxonomicName id="4C2E8DA74C33C6121C6C2E0CFD18224C" box="[588,715,801,824]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C33C6121C6C2E0CFD18224C" box="[588,715,801,824]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C33C6121C6C2E0CFD18224C" box="[588,715,801,824]" italics="true" pageId="69" pageNumber="70">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
(BOLD:AAE4112), and 5.63% divergent to
|
||||
|
|
@ -641,7 +641,7 @@ Torres
|
|||
(2001)
|
||||
</bibRefCitation>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C33C6121ECF2E44FE5222F4" box="[239,385,871,896]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<taxonomicName id="4C2E8DA74C33C6121ECF2E44FE5222F4" box="[239,385,871,896]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="barbarae">
|
||||
<emphasis id="B95A2A364C33C6121ECF2E44FE5222F4" box="[239,385,871,896]" italics="true" pageId="69" pageNumber="70">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
(their “
|
||||
|
|
@ -665,7 +665,7 @@ based on 3 ♂♂, and 1 ♀ from Masoala; 390 bp comparable).
|
|||
<paragraph id="8B91F6244C33C6121EE72E82FCBF2564" blockId="69.[151,1436,151,2013]" pageId="69" pageNumber="70">
|
||||
<emphasis id="B95A2A364C33C6121EE72E82FE3422BC" bold="true" box="[199,487,943,968]" pageId="69" pageNumber="70">Phylogeny/sister species</emphasis>
|
||||
: possibly
|
||||
<taxonomicName id="4C2E8DA74C33C6121C7E2E9CFCE322BC" box="[606,816,943,968]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<taxonomicName id="4C2E8DA74C33C6121C7E2E9CFCE322BC" box="[606,816,943,968]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<emphasis id="B95A2A364C33C6121C7E2E9CFCE322BC" box="[606,816,943,968]" italics="true" pageId="69" pageNumber="70">Ht. menamenoides</emphasis>
|
||||
</taxonomicName>
|
||||
(combined tree in
|
||||
|
|
@ -718,7 +718,7 @@ Torres
|
|||
: endemic to the northeastern rainforests of
|
||||
<collectingCountry id="F339B6B44C33C6121D7929E2FC31259C" box="[857,994,1231,1256]" name="Madagascar" pageId="69" pageNumber="70">Madagascar</collectingCountry>
|
||||
, from Marojejy and Masoala south to Ambatovaky (
|
||||
<figureCitation id="1315EAA14C33C6121F1729D9FE5A2478" box="[311,393,1268,1293]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="69" pageNumber="70">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C33C6121F1729D9FE5A2478" box="[311,393,1268,1293]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="69" pageNumber="70">Fig. 30</figureCitation>
|
||||
C, light green dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C33C6121EE7283AFB4A2444" blockId="69.[151,1436,151,2013]" box="[199,1177,1303,1328]" pageId="69" pageNumber="70">
|
||||
|
|
@ -1216,7 +1216,7 @@ NE, Masoala, Be Dinta, R. Anaovandrano,
|
|||
, H. Raharitsimba, GEN 117
|
||||
<collectionCode id="ED3F6EE14C30C6111D0E28B6FC8624C7" box="[814,853,1435,1459]" pageId="70" pageNumber="71">DL</collectionCode>
|
||||
[genitalia]; ♂ (
|
||||
<figureCitation id="1315EAA14C30C6111A2C28B7FBB324C7" box="[1036,1120,1434,1459]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="70" pageNumber="71">Fig. 18</figureCitation>
|
||||
<figureCitation id="1315EAA14C30C6111A2C28B7FBB324C7" box="[1036,1120,1434,1459]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="70" pageNumber="71">Fig. 18</figureCitation>
|
||||
C), NE, Antsamanarana R., Masoala E., [
|
||||
<quantity id="4CD65BC14C30C6111F1128ECFE7F24AD" box="[305,428,1473,1498]" metricMagnitude="2" metricUnit="m" metricValue="2.85" metricValueMax="5.2" metricValueMin="0.5" pageId="70" pageNumber="71" unit="m" value="285.0" valueMax="520.0" valueMin="50.0">50–520 m</quantity>
|
||||
],
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
<document id="96601D57ACD7E80626906081025A1438" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C34C6151EB72DBAFC3D202F" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteropsis strigula" docType="treatment" docVersion="9" lastPageNumber="67" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="67" updateTime="1698662715872" updateUser="ExternalLinkService">
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<mods:title id="779B4647070CC7D3CC00FFD3E797D4D0">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
|
||||
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|
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|
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sub-group
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This sub-group of the
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||||
<paragraph id="8B91F6244C38C6191EB72DBAFD7021C6" blockId="78.[151,1028,151,211]" box="[151,675,151,178]" pageId="78" pageNumber="79">
|
||||
<heading id="D0D941484C38C6191EB72DBAFD7021C6" bold="true" box="[151,675,151,178]" fontSize="11" level="1" pageId="78" pageNumber="79" reason="1">
|
||||
<emphasis id="B95A2A364C38C6191EB72DBAFD7021C6" bold="true" box="[151,675,151,178]" pageId="78" pageNumber="79">
|
||||
<taxonomicName id="4C2E8DA74C38C6191EB72DBAFDEA21C6" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,569,151,178]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tianae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C38C6191EB72DBAFDEA21C6" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,569,151,178]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tianae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C38C6191EB72DBAFEB921C5" bold="true" box="[151,362,151,177]" italics="true" pageId="78" pageNumber="79">Heteropsis tianae</emphasis>
|
||||
Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -76,7 +76,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C38C6191C542C0EFD322048" box="[628,737,291,316]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C38C6191D362C0EFCB92048" box="[790,874,291,316]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191D362C0EFCB92048" box="[790,874,291,316]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264621" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C38C6191DB42C0EFBC82048" box="[916,1051,291,316]" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
|
||||
C, Amboasary-an-ala, Tsaracamp Boogie Pilgrim, Anjozorobe,
|
||||
|
|
@ -106,7 +106,7 @@ Deposition
|
|||
</emphasis>
|
||||
: ♂
|
||||
<emphasis id="B95A2A364C38C6191C7D2CBCFDB520DE" bold="true" box="[605,614,401,426]" pageId="78" pageNumber="79">(</emphasis>
|
||||
<figureCitation id="1315EAA14C38C6191C462CBCFD6B20DE" box="[614,696,401,426]" captionStart="FIGURE 25" captionStartId="80.[151,250,1045,1067]" captionTargetBox="[254,1341,201,1005]" captionTargetId="figure@80.[231,1356,193,1023]" captionTargetPageId="80" captionText="FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264622/files/figure.png" pageId="78" pageNumber="79">Fig. 25</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191C462CBCFD6B20DE" box="[614,696,401,426]" captionStart="FIGURE 25" captionStartId="80.[151,250,1045,1067]" captionTargetBox="[254,1341,201,1005]" captionTargetId="figure@80.[231,1356,193,1023]" captionTargetPageId="80" captionText="FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264622" httpUri="https://zenodo.org/record/264622/files/figure.png" pageId="78" pageNumber="79">Fig. 25</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C38C6191CC52CBCFCBF20DE" box="[741,876,401,426]" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
|
||||
C, Anjozorobe
|
||||
|
|
@ -120,7 +120,7 @@ C, Anjozorobe
|
|||
010289191 [
|
||||
<collectingCountry id="F339B6B44C38C6191CA12C9BFD6F20BA" box="[641,700,438,462]" name="Qatar" pageId="78" pageNumber="79">QTR</collectingCountry>
|
||||
barcode]; ♀ (
|
||||
<figureCitation id="1315EAA14C38C6191D562C98FC0020BA" box="[886,979,437,462]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191D562C98FC0020BA" box="[886,979,437,462]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264621" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
B),
|
||||
<collectingCountry id="F339B6B44C38C6191A242C98FB5820BA" box="[1028,1163,437,462]" name="Madagascar" pageId="78" pageNumber="79">Madagascar</collectingCountry>
|
||||
C, Amboasary-an-ala, Vanjamanitra-Vazimba forest, Anjozorobe; towards start of Vanjamanitra trail,
|
||||
|
|
@ -169,7 +169,7 @@ extract number],
|
|||
” [=Andrangoloaka, E. Lac Mantasoa],
|
||||
<collectionCode id="ED3F6EE14C38C6191CD32F9BFC9923BA" box="[755,842,694,718]" pageId="78" pageNumber="79">BMNH</collectionCode>
|
||||
(E) #674766 (Godman-Salvin Coll.;
|
||||
<figureCitation id="1315EAA14C38C6191AD72F98FA9F23BA" box="[1271,1356,693,718]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191AD72F98FA9F23BA" box="[1271,1356,693,718]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264621" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
C) and
|
||||
<collectionCode id="ED3F6EE14C38C6191EB72FF4FF3D2385" box="[151,238,729,753]" pageId="78" pageNumber="79">BMNH</collectionCode>
|
||||
(E) #674767;
|
||||
|
|
@ -233,11 +233,11 @@ C, Amboasary-an-ala, vic. Anjozorobe,
|
|||
<subSubSection id="C334A5AF4C38C6191EE72ED1FEA22422" pageId="78" pageNumber="79" type="diagnosis">
|
||||
<paragraph id="8B91F6244C38C6191EE72ED1FEA22422" blockId="78.[151,1437,255,1902]" pageId="78" pageNumber="79">
|
||||
<emphasis id="B95A2A364C38C6191EE72ED1FE922561" bold="true" box="[199,321,1020,1045]" pageId="78" pageNumber="79">Diagnosis.</emphasis>
|
||||
<taxonomicName id="4C2E8DA74C38C6191F672ED0FDEF2561" box="[327,572,1020,1045]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C38C6191F672ED0FDEF2561" box="[327,572,1020,1045]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C38C6191F672ED0FDEF2561" box="[327,572,1020,1045]" italics="true" pageId="78" pageNumber="79">Heteropsis oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
, described below, is hard to distinguish from
|
||||
<taxonomicName id="4C2E8DA74C38C6191A182ED0FB742560" box="[1080,1191,1021,1044]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C38C6191A182ED0FB742560" box="[1080,1191,1021,1044]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C38C6191A182ED0FB742560" box="[1080,1191,1021,1044]" italics="true" pageId="78" pageNumber="79">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
except by ♂ genitalia (the gnathos is more flattened at the base than in that species) and by mitochondrial DNA sequence (DNA barcode highly distinct from ‘sp. 28’ to be treated in a subsequent paper, and from
|
||||
|
|
@ -248,33 +248,33 @@ except by ♂ genitalia (the gnathos is more flattened at the base than in that
|
|||
)
|
||||
</taxonomicName>
|
||||
; see under DNA divergences). On average, the HWV Mb of
|
||||
<taxonomicName id="4C2E8DA74C38C6191CEB2945FCEF250B" box="[715,828,1128,1151]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C38C6191CEB2945FCEF250B" box="[715,828,1128,1151]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C38C6191CEB2945FCEF250B" box="[715,828,1128,1151]" italics="true" pageId="78" pageNumber="79">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
is slightly more outdented near space-M2 (
|
||||
<figureCitation id="1315EAA14C38C6191B08294AFAA925F4" box="[1320,1402,1127,1152]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191B08294AFAA925F4" box="[1320,1402,1127,1152]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264621" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
A– C), and the HWV tends to have the space-CuA2 ocellus expressed as small black-ringed ocellus. However,
|
||||
<taxonomicName id="4C2E8DA74C38C6191B5929A1FEDE25B3" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C38C6191B5929A1FEDE25B3" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C38C6191B5929A1FEDE25B3" italics="true" pageId="78" pageNumber="79">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C38C6191F3F2983FEA225B3" box="[287,369,1198,1223]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191F3F2983FEA225B3" box="[287,369,1198,1223]" captionStart="FIGURE 24" captionStartId="79.[151,250,1896,1918]" captionTargetBox="[278,1305,215,1858]" captionTargetId="figure@79.[267,1320,193,1875]" captionTargetPageId="79" captionText="FIGURE 24. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis tianae, Lees & Kremen, sp. nov. (Anjozorobe; NHMUK 010289158). B: PT ♀ of Ht. tianae (Anjozorobe; BMNH (E) # 676763; DNA barcoded specimen). C: Ht. tianae ♀ (‘ Andrangoloaka’; BMNH (E) # 674766, Godman-Salvin Coll.). D: HT ♂ of Heteropsis oberthueri Lees, sp. nov. (Zahamena; NHMUK 010289159). E: PT ♀ of Ht. oberthueri (Zahamena; NHMUK 010289160). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264621" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="78" pageNumber="79">Fig. 24</figureCitation>
|
||||
D–E,
|
||||
<figureCitation id="1315EAA14C38C6191F952983FDDB25B3" box="[437,520,1198,1223]" captionStart="FIGURE 26" captionStartId="82.[151,250,1631,1653]" captionTargetBox="[292,1295,190,1603]" captionTargetId="figure@82.[284,1303,179,1614]" captionTargetPageId="82" captionText="FIGURE 26. Live ♀ adults in natural posture. A: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 5 / 11 / 2006). B: Ht. andasibe (Lees, 2003) (Andasibe; 24 / 10 / 2014). Photos: D. C. Lees." httpUri="https://zenodo.org/record/264623/files/figure.png" pageId="78" pageNumber="79">Fig. 26</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191F952983FDDB25B3" box="[437,520,1198,1223]" captionStart="FIGURE 26" captionStartId="82.[151,250,1631,1653]" captionTargetBox="[292,1295,190,1603]" captionTargetId="figure@82.[284,1303,179,1614]" captionTargetPageId="82" captionText="FIGURE 26. Live ♀ adults in natural posture. A: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 5 / 11 / 2006). B: Ht. andasibe (Lees, 2003) (Andasibe; 24 / 10 / 2014). Photos: D. C. Lees." figureDoi="http://doi.org/10.5281/zenodo.264623" httpUri="https://zenodo.org/record/264623/files/figure.png" pageId="78" pageNumber="79">Fig. 26</figureCitation>
|
||||
A) can be distinguished from the sympatric
|
||||
<taxonomicName id="4C2E8DA74C38C6191A24299DFB4525B3" box="[1028,1174,1198,1223]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C38C6191A24299DFB4525B3" box="[1028,1174,1198,1223]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C38C6191A24299DFB4525B3" box="[1028,1174,1198,1223]" italics="true" pageId="78" pageNumber="79">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
by the distinctly more outangled HWV Mb of that species (
|
||||
<figureCitation id="1315EAA14C38C6191C1029FFFD51259F" box="[560,642,1234,1259]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="78" pageNumber="79">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191C1029FFFD51259F" box="[560,642,1234,1259]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="78" pageNumber="79">Fig. 22</figureCitation>
|
||||
C;
|
||||
<figureCitation id="1315EAA14C38C6191C8029FFFD21259F" box="[672,754,1234,1259]" captionStart="FIGURE 26" captionStartId="82.[151,250,1631,1653]" captionTargetBox="[292,1295,190,1603]" captionTargetId="figure@82.[284,1303,179,1614]" captionTargetPageId="82" captionText="FIGURE 26. Live ♀ adults in natural posture. A: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 5 / 11 / 2006). B: Ht. andasibe (Lees, 2003) (Andasibe; 24 / 10 / 2014). Photos: D. C. Lees." httpUri="https://zenodo.org/record/264623/files/figure.png" pageId="78" pageNumber="79">Fig. 26</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191C8029FFFD21259F" box="[672,754,1234,1259]" captionStart="FIGURE 26" captionStartId="82.[151,250,1631,1653]" captionTargetBox="[292,1295,190,1603]" captionTargetId="figure@82.[284,1303,179,1614]" captionTargetPageId="82" captionText="FIGURE 26. Live ♀ adults in natural posture. A: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 5 / 11 / 2006). B: Ht. andasibe (Lees, 2003) (Andasibe; 24 / 10 / 2014). Photos: D. C. Lees." figureDoi="http://doi.org/10.5281/zenodo.264623" httpUri="https://zenodo.org/record/264623/files/figure.png" pageId="78" pageNumber="79">Fig. 26</figureCitation>
|
||||
B). In the Angavo massif, I observed the last species to prefer habitat only a few metres from the forest margin. The ventrally very similar
|
||||
<taxonomicName id="4C2E8DA74C38C6191DD629DAFB41247A" box="[1014,1170,1271,1294]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C38C6191DD629DAFB41247A" box="[1014,1170,1271,1294]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C38C6191DD629DAFB41247A" box="[1014,1170,1271,1294]" italics="true" pageId="78" pageNumber="79">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
usually has a smoother outcurve to the HWV Mb at space-M2 and strong tendency to expression of a small ocellus in space-M3 on the HWD (
|
||||
<figureCitation id="1315EAA14C38C6191EC92810FEE82422" box="[233,315,1341,1366]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="78" pageNumber="79">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C38C6191EC92810FEE82422" box="[233,315,1341,1366]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="78" pageNumber="79">Fig. 22</figureCitation>
|
||||
AB).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -282,20 +282,20 @@ AB).
|
|||
<paragraph id="8B91F6244C38C6191EE7284CFD66261A" blockId="78.[151,1437,255,1902]" pageId="78" pageNumber="79">
|
||||
<emphasis id="B95A2A364C38C6191EE7284CFE6B240E" bold="true" box="[199,440,1377,1402]" pageId="78" pageNumber="79">Description. Wings:</emphasis>
|
||||
upperside uniform mid brown, FW space-CuA1 ocellus with an orange ocellus ‘ring’ which is sometimes rather hexagonal in shape and eccentric, being wider at proximad edge. FW space-M1 ocellus small with narrow orange ring. HW space-CuA1 ocellus the only one expressed there and somewhat elliptic. Darkish brown diffuse, slightly wavy hair-brush emanating mainly from below vein 1A+2A as far as mid-vein. Underside greyish brown, rather more uniform and less strongly irrorated than in many
|
||||
<taxonomicName id="4C2E8DA74C38C6191AED28DFFA96277D" box="[1229,1349,1522,1545]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C38C6191AED28DFFA96277D" box="[1229,1349,1522,1545]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C38C6191AED28DFFA96277D" box="[1229,1349,1522,1545]" italics="true" pageId="78" pageNumber="79">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
, rather uniformly irrorated brown and ochreous brown. Space-CuA1 ocellus FWV with orange ring yellowing proximad, this Ore following the darkish brown concave curve of the Mb before it bends back to mid-costa. Space-M1 ocellus FWV reduced to white pupil with narrow black iris without trace of orange ring. On HWV, space-CuA1 ocellus slightly elliptic with narrow black iris and small also, without orange ring trace. HWV space-M1-M2 ocelli as white ‘pupil’ points. HWV Mb darkish brown and fairly straight, only gently curving. FWV cell area with four darker brown rather equally spaced transverse Cbs. Basal areas are not much darker than areas distad of the Mb, and a darker brown PMb is weakly represented.
|
||||
<emphasis id="B95A2A364C38C6191C922BC7FCF92677" bold="true" box="[690,810,1770,1795]" pageId="78" pageNumber="79">Variation.</emphasis>
|
||||
♂♂ similar to each other, but seasonal variation has not been quantified. HWV Mb sometimes with a yellow Mf distad of it in space-M2. HWV space-CuA2 ocellus may be slightly expressed as small white pupil with elliptic black iris. ♀♀ similar in pattern, but larger and somewhat lighter dorsally with a more rounded HW shape.
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C39C6181EB72A45FB7926AE" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="79" pageNumber="80" targetBox="[278,1305,215,1858]" targetPageId="79">
|
||||
<caption id="DF51A6AC4C39C6181EB72A45FB7926AE" ID-DOI="http://doi.org/10.5281/zenodo.264621" httpUri="https://zenodo.org/record/264621/files/figure.png" pageId="79" pageNumber="80" targetBox="[278,1305,215,1858]" targetPageId="79">
|
||||
<paragraph id="8B91F6244C39C6181EB72A45FB7926AE" blockId="79.[151,1436,1896,2010]" pageId="79" pageNumber="80">
|
||||
<emphasis id="B95A2A364C39C6181EB72A45FEF62609" bold="true" box="[151,293,1896,1918]" pageId="79" pageNumber="80">FIGURE 24.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
<emphasis id="B95A2A364C39C6181CB82A45FD792609" bold="true" box="[664,682,1896,1917]" pageId="79" pageNumber="80">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C39C6181D0A2A44FB70260A" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[810,1187,1896,1918]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="tianae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C39C6181D0A2A44FB70260A" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[810,1187,1896,1918]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="tianae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C39C6181D0A2A44FC31260A" box="[810,994,1897,1918]" italics="true" pageId="79" pageNumber="80">Heteropsis tianae</emphasis>
|
||||
, Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -306,19 +306,19 @@ Adults in DV (left), VV (right).
|
|||
(Anjozorobe; NHMUK010289158).
|
||||
<emphasis id="B95A2A364C39C6181F5F2AAAFE4326E8" bold="true" box="[383,400,1927,1948]" pageId="79" pageNumber="80">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C39C6181FDF2AA5FDB526E9" box="[511,614,1928,1949]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C39C6181FDF2AA5FDB526E9" box="[511,614,1928,1949]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C39C6181FDF2AA5FDB526E9" box="[511,614,1928,1949]" italics="true" pageId="79" pageNumber="80">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
(Anjozorobe; BMNH(E) #676763; DNA barcoded specimen).
|
||||
<emphasis id="B95A2A364C39C6181AD72AABFADA26E8" bold="true" box="[1271,1289,1926,1948]" pageId="79" pageNumber="80">C</emphasis>
|
||||
:
|
||||
<taxonomicName id="4C2E8DA74C39C6181B392AA5FA5326E9" box="[1305,1408,1928,1949]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C39C6181B392AA5FA5326E9" box="[1305,1408,1928,1949]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C39C6181B392AA5FA5326E9" box="[1305,1408,1928,1949]" italics="true" pageId="79" pageNumber="80">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
♀ (‘Andrangoloaka’; BMNH(E) #674766, Godman-Salvin Coll.).
|
||||
<emphasis id="B95A2A364C39C6181D042A8BFCE526CF" bold="true" box="[804,822,1958,1979]" pageId="79" pageNumber="80">D</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C39C6181D842A8BFB6D26CF" authority="Lees" authorityName="Lees" box="[932,1214,1957,1979]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="oberthueri" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C39C6181D842A8BFB6D26CF" authority="Lees" authorityName="Lees" box="[932,1214,1957,1979]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="oberthueri" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C39C6181D842A8BFB5726CF" box="[932,1156,1957,1979]" italics="true" pageId="79" pageNumber="80">Heteropsis oberthueri</emphasis>
|
||||
<emphasis id="B95A2A364C39C6181AAB2A8BFB6D26CF" bold="true" box="[1163,1214,1958,1979]" pageId="79" pageNumber="80">Lees</emphasis>
|
||||
</taxonomicName>
|
||||
|
|
@ -329,17 +329,17 @@ Adults in DV (left), VV (right).
|
|||
(Zahamena; NHMUK010289159).
|
||||
<emphasis id="B95A2A364C39C6181F5D2AE9FE5D26AD" bold="true" box="[381,398,1988,2009]" pageId="79" pageNumber="80">E</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C39C6181FD42AE8FD5626AE" box="[500,645,1988,2010]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C39C6181FD42AE8FD5626AE" box="[500,645,1988,2010]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C39C6181FD42AE8FD5626AE" box="[500,645,1988,2010]" italics="true" pageId="79" pageNumber="80">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
(Zahamena; NHMUK010289160). Scale bar: 30 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF51A6AC4C26C6071EB72938FC3D253D" httpUri="https://zenodo.org/record/264622/files/figure.png" pageId="80" pageNumber="81" targetBox="[254,1341,201,1005]" targetPageId="80">
|
||||
<caption id="DF51A6AC4C26C6071EB72938FC3D253D" ID-DOI="http://doi.org/10.5281/zenodo.264622" httpUri="https://zenodo.org/record/264622/files/figure.png" pageId="80" pageNumber="81" targetBox="[254,1341,201,1005]" targetPageId="80">
|
||||
<paragraph id="8B91F6244C26C6071EB72938FC3D253D" blockId="80.[151,1436,1044,1097]" pageId="80" pageNumber="81">
|
||||
<emphasis id="B95A2A364C26C6071EB72938FEF1255E" bold="true" box="[151,290,1045,1067]" pageId="80" pageNumber="81">FIGURE 25.</emphasis>
|
||||
♂ genitalia in views stated. A:
|
||||
<taxonomicName id="4C2E8DA74C26C6071C51293BFC05255E" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[625,982,1044,1067]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="tianae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C26C6071C51293BFC05255E" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[625,982,1044,1067]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="tianae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C26C6071C51293BFD0D255F" box="[625,734,1046,1067]" italics="true" pageId="80" pageNumber="81">Heteropsis</emphasis>
|
||||
<emphasis id="B95A2A364C26C6071CC6293BFCF6255F" box="[742,805,1046,1067]" italics="true" pageId="80" pageNumber="81">tianae</emphasis>
|
||||
Lees & Kremen
|
||||
|
|
@ -349,7 +349,7 @@ Lees & Kremen
|
|||
<emphasis id="B95A2A364C26C6071DC32938FBE5255E" bold="true" box="[995,1078,1045,1066]" pageId="80" pageNumber="81">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
(Anjozorobe; 228DL), LV, SV. B:
|
||||
<taxonomicName id="4C2E8DA74C26C6071EB72919FE7D253D" authority="Lees" authorityName="Lees" box="[151,430,1075,1097]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="oberthueri" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C26C6071EB72919FE7D253D" authority="Lees" authorityName="Lees" box="[151,430,1075,1097]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="oberthueri" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C26C6071EB72919FED7253D" box="[151,260,1076,1097]" italics="true" pageId="80" pageNumber="81">Heteropsis</emphasis>
|
||||
<emphasis id="B95A2A364C26C6071F2A291EFEA6253D" box="[266,373,1075,1097]" italics="true" pageId="80" pageNumber="81">oberthueri</emphasis>
|
||||
Lees
|
||||
|
|
@ -376,13 +376,13 @@ Lees
|
|||
: both 1A+2A and 3A have a tapered ‘balloon-like’ inflation, more swollen distad, from base to mid-vein, covered in thin grey-brown scales (
|
||||
<bibRefCitation id="EFBF8BD54C26C6071CB3282BFC3F246A" author="Lees" box="[659,1004,1285,1310]" pageId="80" pageNumber="81" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
|
||||
Lees 1997: 96,
|
||||
<figureCitation id="1315EAA14C26C6071D612828FC51246A" box="[833,898,1285,1310]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="80" pageNumber="81">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C26C6071D612828FC51246A" box="[833,898,1285,1310]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="80" pageNumber="81">Fig. 3</figureCitation>
|
||||
A: sp. 49
|
||||
</bibRefCitation>
|
||||
). HW veins M2, M3, CuA1 and CuA2 are also narrowly inflated throughout their lengths and have specialized scales on the dorsal surface (
|
||||
<bibRefCitation id="EFBF8BD54C26C6071AD62807FF212412" author="Lees" pageId="80" pageNumber="81" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
|
||||
Lees 1997: 96,
|
||||
<figureCitation id="1315EAA14C26C6071EB72860FF082412" box="[151,219,1357,1382]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="80" pageNumber="81">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C26C6071EB72860FF082412" box="[151,219,1357,1382]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="80" pageNumber="81">Fig. 3</figureCitation>
|
||||
A
|
||||
</bibRefCitation>
|
||||
). Abdominal black androconia indistinctly visible ventro-laterally around A2. HWD discocellular brush fawn to yellowish/blonde at tip. Discocellular patch hwd (orange, small, lenticular, composed of narrow yellow scales) (observations on MAD 239–242, MAD244; Anjozorobe, n=5).
|
||||
|
|
@ -394,9 +394,9 @@ A
|
|||
<paragraph id="8B91F6244C26C6071EE72B2FFACD264F" blockId="80.[151,1437,1141,2033]" pageId="80" pageNumber="81">
|
||||
<emphasis id="B95A2A364C26C6071EE72B2FFE9A276F" bold="true" box="[199,329,1538,1563]" pageId="80" pageNumber="81">♂ genitalia</emphasis>
|
||||
: 228DL, PT (
|
||||
<figureCitation id="1315EAA14C26C6071FC62B2FFDEB276F" box="[486,568,1538,1563]" captionStart="FIGURE 25" captionStartId="80.[151,250,1045,1067]" captionTargetBox="[254,1341,201,1005]" captionTargetId="figure@80.[231,1356,193,1023]" captionTargetPageId="80" captionText="FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264622/files/figure.png" pageId="80" pageNumber="81">Fig. 25</figureCitation>
|
||||
<figureCitation id="1315EAA14C26C6071FC62B2FFDEB276F" box="[486,568,1538,1563]" captionStart="FIGURE 25" captionStartId="80.[151,250,1045,1067]" captionTargetBox="[254,1341,201,1005]" captionTargetId="figure@80.[231,1356,193,1023]" captionTargetPageId="80" captionText="FIGURE 25. ♂ genitalia in views stated. A: Heteropsis tianae Lees & Kremen, sp. nov. (Anjozorobe; 228 DL), LV, SV. B: Heteropsis oberthueri Lees, sp. nov. (Zahamena; 371 DL), LV, DV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264622" httpUri="https://zenodo.org/record/264622/files/figure.png" pageId="80" pageNumber="81">Fig. 25</figureCitation>
|
||||
A): from LV, uncus slightly proud of dorsal curve of tegumen and longer than it, fairly straight and parallel-sided to hook without a distinct ‘head’ although slightly broadened ventrally towards tip (evident from the SV), and tegumen fairly narrow at hinge with vinculum; tegumen viewed laterally less tapered ventrad than many species. Valve arm with fairly short neck (distinctly recurved and bowed inwards from the SV) and slight club at tip covered in spinoid setae, with distinct ‘beak’ oriented towards uncus and slightly mesad, uncus tip distinctly proud of valves. Gnathos from rather small base with distinct flattening, more so than in
|
||||
<taxonomicName id="4C2E8DA74C26C6071B592B9AFEDE2787" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C26C6071B592B9AFEDE2787" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C26C6071B592B9AFEDE2787" italics="true" pageId="80" pageNumber="81">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
(although not with ear-like structure) near base, recurved downwards at tapered tip (and inwards from the SV) with slight serration on dorsal surface. Saccus moderate length and parallel sided, aedeagus slightly shorter than valve and strongly recurved twice, towards and away from the also proximally uprecurved ostium.
|
||||
|
|
@ -428,13 +428,13 @@ and of
|
|||
<bibRefCitation id="EFBF8BD54C27C6061D0F2D91FC0C21A1" author="Oberthur" box="[815,991,188,213]" pageId="81" pageNumber="82" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(LT ♂♂ designated above under
|
||||
<taxonomicName id="4C2E8DA74C27C6061B592D90FEDB218C" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C27C6061B592D90FEDB218C" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C27C6061B592D90FEDB218C" italics="true" pageId="81" pageNumber="82">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
; see
|
||||
<figureCitation id="1315EAA14C27C6061F622DF2FE41218C" box="[322,402,223,248]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="81" pageNumber="82">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C27C6061F622DF2FE41218C" box="[322,402,223,248]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="81" pageNumber="82">Fig. 22</figureCitation>
|
||||
C–D) were examined and are clearly different, along with
|
||||
<taxonomicName id="4C2E8DA74C27C6061A002DCCFB13218C" box="[1056,1216,223,248]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C27C6061A002DCCFB13218C" box="[1056,1216,223,248]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C27C6061A002DCCFB13218C" box="[1056,1216,223,248]" italics="true" pageId="81" pageNumber="82">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
, described below.
|
||||
|
|
@ -442,7 +442,7 @@ C–D) were examined and are clearly different, along with
|
|||
<paragraph id="8B91F6244C27C6061EE72C29FDA82081" blockId="81.[151,1437,151,2007]" pageId="81" pageNumber="82">
|
||||
<emphasis id="B95A2A364C27C6061EE72C29FE372069" bold="true" box="[199,484,260,285]" pageId="81" pageNumber="82">Additional information.</emphasis>
|
||||
DNA divergences: COI-5P cluster number BOLD:AAE4112 (exemplar BMAD200- 15, DL14Z-051) is about 2.58% divergent to
|
||||
<taxonomicName id="4C2E8DA74C27C6061CEE2C04FCA42034" box="[718,887,295,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C27C6061CEE2C04FCA42034" box="[718,887,295,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C27C6061CEE2C04FCA42034" box="[718,887,295,320]" italics="true" pageId="81" pageNumber="82">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
(cluster number BOLD:ACW4996, exemplar BMAD242-15, DL06-11, RNI Zahamena) and about 4.17% divergent from ‘sp. 28’ (BOLD:AAE5458), which is not likely to be the sister species. In their dataset for COII (
|
||||
|
|
@ -452,7 +452,7 @@ Torres
|
|||
, 2001
|
||||
</bibRefCitation>
|
||||
),
|
||||
<taxonomicName id="4C2E8DA74C27C6061A3F2C5CFB4320FC" box="[1055,1168,369,392]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C27C6061A3F2C5CFB4320FC" box="[1055,1168,369,392]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C27C6061A3F2C5CFB4320FC" box="[1055,1168,369,392]" italics="true" pageId="81" pageNumber="82">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
(as their “
|
||||
|
|
@ -480,7 +480,7 @@ AY
|
|||
<paragraph id="8B91F6244C27C6061EE72CD2FE8D23F1" blockId="81.[151,1437,151,2007]" pageId="81" pageNumber="82">
|
||||
<emphasis id="B95A2A364C27C6061EE72CD2FE3B236C" bold="true" box="[199,488,511,536]" pageId="81" pageNumber="82">Phylogeny/sister species</emphasis>
|
||||
: probably
|
||||
<taxonomicName id="4C2E8DA74C27C6061C4B2F2CFCDC236C" box="[619,783,511,536]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C27C6061C4B2F2CFCDC236C" box="[619,783,511,536]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<emphasis id="B95A2A364C27C6061C4B2F2CFCDC236C" box="[619,783,511,536]" italics="true" pageId="81" pageNumber="82">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
based on the close relationship of the COI-5P barcode (confirmed by
|
||||
|
|
@ -496,7 +496,7 @@ Torres
|
|||
(2001
|
||||
</bibRefCitation>
|
||||
: 467) suggested a topological relationship of
|
||||
<taxonomicName id="4C2E8DA74C27C6061CAB2F64FD282314" box="[651,763,585,608]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C27C6061CAB2F64FD282314" box="[651,763,585,608]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C27C6061CAB2F64FD282314" box="[651,763,585,608]" italics="true" pageId="81" pageNumber="82">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
(sp. 49) with
|
||||
|
|
@ -535,7 +535,7 @@ Torres
|
|||
<paragraph id="8B91F6244C27C6061EE72E69FC2E22D0" blockId="81.[151,1437,151,2007]" pageId="81" pageNumber="82">
|
||||
<emphasis id="B95A2A364C27C6061EE72E69FE8A2229" bold="true" box="[199,345,836,861]" pageId="81" pageNumber="82">Distribution</emphasis>
|
||||
: endemic to the Angavo massif, including Anjozorobe and Andrangoloaka, as far as is known (
|
||||
<figureCitation id="1315EAA14C27C6061EBF2E4AFF2122F4" box="[159,242,871,896]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="81" pageNumber="82">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C27C6061EBF2E4AFF2122F4" box="[159,242,871,896]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="81" pageNumber="82">Fig. 30</figureCitation>
|
||||
C, dark pink dots). Due to a formatting error, the locality was incorrectly given in Table 1 of
|
||||
<bibRefCitation id="EFBF8BD54C27C6061B2D2E45FF1C22D0" author="Torres" pageId="81" pageNumber="82" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. & Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
|
||||
Torres
|
||||
|
|
@ -568,7 +568,7 @@ Centre, forêt a l’Est du lac de Mantasoa, Andrangoloaka, 27-II/
|
|||
], data as above but: DCL-DB-4469, DCL-DB- 4470; ♂ [
|
||||
<collectionCode id="ED3F6EE14C27C6061F272930FEB22540" box="[263,353,1053,1076]" pageId="81" pageNumber="82">BMNH</collectionCode>
|
||||
; probably referable to
|
||||
<taxonomicName id="4C2E8DA74C27C6061C4C2930FD332540" box="[620,736,1053,1076]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C27C6061C4C2930FD332540" box="[620,736,1053,1076]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C27C6061C4C2930FD332540" box="[620,736,1053,1076]" italics="true" pageId="81" pageNumber="82">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
],
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<treatment id="038747324C3BC61A1EB72BC9FD9D2680" ID-DOI="http://doi.org/10.5281/zenodo.6086454" ID-GBIF-Taxon="120817134" ID-Zenodo-Dep="6086454" LSID="urn:lsid:plazi:treatment:038747324C3BC61A1EB72BC9FD9D2680" httpUri="http://treatment.plazi.org/id/038747324C3BC61A1EB72BC9FD9D2680" lastPageNumber="78" pageId="77" pageNumber="78">
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<treatment id="038747324C3BC61A1EB72BC9FD9D2680" ID-DOI="http://doi.org/10.5281/zenodo.6086454" ID-GBIF-Taxon="120817134" ID-Zenodo-Dep="6086454" LSID="urn:lsid:plazi:treatment:038747324C3BC61A1EB72BC9FD9D2680" httpUri="http://treatment.plazi.org/id/038747324C3BC61A1EB72BC9FD9D2680" lastPageNumber="78" pageId="77" pageNumber="78" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
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<paragraph id="8B91F6244C3BC61A1EB72BC9FE2E278A" blockId="77.[151,509,1764,1790]" box="[151,509,1764,1790]" pageId="77" pageNumber="78">
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<heading id="D0D941484C3BC61A1EB72BC9FE2E278A" bold="true" box="[151,509,1764,1790]" fontSize="11" level="1" pageId="77" pageNumber="78" reason="1">
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<taxonomicName id="4C2E8DA74C3BC61A1EB72BC9FE66278A" box="[151,437,1764,1790]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="angulifascia">
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<taxonomicName id="4C2E8DA74C3BC61A1EB72BC9FE66278A" box="[151,437,1764,1790]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="angulifascia">
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<emphasis id="B95A2A364C3BC61A1EB72BC9FE66278A" bold="true" box="[151,437,1764,1790]" italics="true" pageId="77" pageNumber="78">Heteropsis angulifascia</emphasis>
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</taxonomicName>
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clade
|
||||
|
|
@ -52,9 +52,9 @@ clade
|
|||
In the sampling of Aduse-Poku
|
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<emphasis id="B95A2A364C3BC61A1C3F2A04FDB62635" box="[543,613,1832,1857]" italics="true" pageId="77" pageNumber="78">et al.,</emphasis>
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(2015,
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<figureCitation id="1315EAA14C3BC61A1CE92A05FCC32635" box="[713,784,1832,1857]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="77" pageNumber="78">Fig. 1</figureCitation>
|
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<figureCitation id="1315EAA14C3BC61A1CE92A05FCC32635" box="[713,784,1832,1857]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="77" pageNumber="78">Fig. 1</figureCitation>
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),
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1D0F2A04FB732635" authority="Butler, 1879" authorityName="Butler" authorityYear="1879" box="[815,1184,1832,1857]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="angulifascia">
|
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<taxonomicName id="4C2E8DA74C3BC61A1D0F2A04FB732635" authority="Butler, 1879" authorityName="Butler" authorityYear="1879" box="[815,1184,1832,1857]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="angulifascia">
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<emphasis id="B95A2A364C3BC61A1D0F2A04FC392635" box="[815,1002,1832,1857]" italics="true" pageId="77" pageNumber="78">Ht. angulifascia</emphasis>
|
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(
|
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<bibRefCitation id="EFBF8BD54C3BC61A1A212A05FB4B2635" author="Butler" box="[1025,1176,1832,1857]" pageId="77" pageNumber="78" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
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|
|
@ -67,7 +67,7 @@ In the sampling of Aduse-Poku
|
|||
<emphasis id="B95A2A364C3BC61A1ACD2A04FAE82634" box="[1261,1339,1833,1856]" italics="true" pageId="77" pageNumber="78">anceps</emphasis>
|
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</taxonomicName>
|
||||
’ (=
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1B592A04FEDE2611" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="oberthueri">
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1B592A04FEDE2611" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="oberthueri">
|
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<emphasis id="B95A2A364C3BC61A1B592A04FEDE2611" italics="true" pageId="77" pageNumber="78">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
),
|
||||
|
|
@ -75,7 +75,7 @@ In the sampling of Aduse-Poku
|
|||
<emphasis id="B95A2A364C3BC61A1F052A60FE762611" box="[293,421,1868,1893]" italics="true" pageId="77" pageNumber="78">Ht. turbans</emphasis>
|
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</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1FFD2A60FD9B2611" box="[477,584,1868,1893]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="sabas">
|
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<taxonomicName id="4C2E8DA74C3BC61A1FFD2A60FD9B2611" box="[477,584,1868,1893]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="sabas">
|
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<emphasis id="B95A2A364C3BC61A1FFD2A60FD9B2611" box="[477,584,1868,1893]" italics="true" pageId="77" pageNumber="78">Ht. sabas</emphasis>
|
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</taxonomicName>
|
||||
formed a well-supported molecular clade. This clade, nested within the
|
||||
|
|
@ -83,7 +83,7 @@ formed a well-supported molecular clade. This clade, nested within the
|
|||
<emphasis id="B95A2A364C3BC61A1B592A60FF3D26FD" italics="true" pageId="77" pageNumber="78">Ht. strigula</emphasis>
|
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</taxonomicName>
|
||||
sub-group, and named after
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1C142A5CFD3926FD" box="[564,746,1904,1929]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="angulifascia">
|
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<taxonomicName id="4C2E8DA74C3BC61A1C142A5CFD3926FD" box="[564,746,1904,1929]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="angulifascia">
|
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<emphasis id="B95A2A364C3BC61A1C142A5CFD3926FD" box="[564,746,1904,1929]" italics="true" pageId="77" pageNumber="78">Ht. angulifascia</emphasis>
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</taxonomicName>
|
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, also includes the described species
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<mods:title id="47596728E3071CF66A71F1BC8D6EB4A8">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
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<mods:namePart id="75BB30B92CC99B764F15F3338BED9F2E">C, Lees David</mods:namePart>
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<treatment id="038747324C3FC61A1EB72BCFFAD227D5" ID-DOI="http://doi.org/10.5281/zenodo.6086452" ID-GBIF-Taxon="120817141" ID-Zenodo-Dep="6086452" LSID="urn:lsid:plazi:treatment:038747324C3FC61A1EB72BCFFAD227D5" httpUri="http://treatment.plazi.org/id/038747324C3FC61A1EB72BCFFAD227D5" lastPageId="77" lastPageNumber="78" pageId="73" pageNumber="74">
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<treatment id="038747324C3FC61A1EB72BCFFAD227D5" ID-DOI="http://doi.org/10.5281/zenodo.6086452" ID-GBIF-Taxon="120817141" ID-Zenodo-Dep="6086452" LSID="urn:lsid:plazi:treatment:038747324C3FC61A1EB72BCFFAD227D5" httpUri="http://treatment.plazi.org/id/038747324C3FC61A1EB72BCFFAD227D5" lastPageId="77" lastPageNumber="78" pageId="73" pageNumber="74" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
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<subSubSection id="C334A5AF4C3FC61E1EB72BCFFC2C2669" pageId="73" pageNumber="74" type="nomenclature">
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<paragraph id="8B91F6244C3FC61E1EB72BCFFD012788" blockId="73.[151,1023,1762,1821]" box="[151,722,1762,1788]" pageId="73" pageNumber="74">
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<heading id="D0D941484C3FC61E1EB72BCFFD012788" bold="true" box="[151,722,1762,1788]" fontSize="11" level="1" pageId="73" pageNumber="74" reason="1">
|
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<emphasis id="B95A2A364C3FC61E1EB72BCFFD012788" bold="true" box="[151,722,1762,1788]" pageId="73" pageNumber="74">
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1EB72BCFFDBB2788" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,616,1762,1788]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="roussettae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C3FC61E1EB72BCFFDBB2788" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,616,1762,1788]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="roussettae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C3FC61E1EB72BCFFE4A2788" bold="true" box="[151,409,1762,1788]" italics="true" pageId="73" pageNumber="74">Heteropsis roussettae</emphasis>
|
||||
Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -71,7 +71,7 @@ material., Deposition
|
|||
:
|
||||
</emphasis>
|
||||
♂ (
|
||||
<figureCitation id="1315EAA14C3FC61E1D722A47FC6326F7" box="[850,944,1898,1923]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="73" pageNumber="74">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C3FC61E1D722A47FC6326F7" box="[850,944,1898,1923]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="73" pageNumber="74">Fig. 22</figureCitation>
|
||||
A): ♂,
|
||||
<collectingCountry id="F339B6B44C3FC61E1A322A47FB4826F7" box="[1042,1179,1898,1923]" name="Madagascar" pageId="73" pageNumber="74">Madagascar</collectingCountry>
|
||||
, Montagne d'Ambre,
|
||||
|
|
@ -139,7 +139,7 @@ extract number], IA316 [isotope voucher],
|
|||
010289156 [
|
||||
<collectingCountry id="F339B6B44C3CC61D1A692C28FB572069" box="[1097,1156,261,285]" name="Qatar" pageId="74" pageNumber="75">QTR</collectingCountry>
|
||||
barcode]; ♀ (
|
||||
<figureCitation id="1315EAA14C3CC61D1B062C29FAAF2069" box="[1318,1404,260,285]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="74" pageNumber="75">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C3CC61D1B062C29FAAF2069" box="[1318,1404,260,285]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="74" pageNumber="75">Fig. 22</figureCitation>
|
||||
B),
|
||||
<collectingCountry id="F339B6B44C3CC61D1EB72C07FECD2037" box="[151,286,298,323]" name="Madagascar" pageId="74" pageNumber="75">Madagascar</collectingCountry>
|
||||
N, Montagne d'Ambre, tourist campsite, c.
|
||||
|
|
@ -177,7 +177,7 @@ barcode]; ♂, N, Montagne d'Ambre [
|
|||
010289188 [
|
||||
<collectingCountry id="F339B6B44C3CC61D1D5A2CBAFC6620DB" box="[890,949,407,431]" name="Qatar" pageId="74" pageNumber="75">QTR</collectingCountry>
|
||||
barcode]; ♂ (
|
||||
<figureCitation id="1315EAA14C3CC61D1A442CBBFB6F20DB" box="[1124,1212,406,431]" captionStart="FIGURE 23" captionStartId="76.[151,250,1428,1450]" captionTargetBox="[249,1339,207,1397]" captionTargetId="figure@76.[234,1353,193,1406]" captionTargetPageId="76" captionText="FIGURE 23. Adults in views stated. A: ♂ PT genitalia of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; 33 DL), LV, SV. Scale bar: 3 mm. B: ♂ genitalia and sternal androconial scaling in Ht. roussettae (“ sp. 26 ” modified from Lees, 1997; not to scale), LV. C: ♂ genitalia and sternal androconial scaling in Ht. andasibe (modified from Lees, 1997: where labelled “ antsianakana ”; not to scale), LV." httpUri="https://zenodo.org/record/264620/files/figure.png" pageId="74" pageNumber="75">Fig. 23</figureCitation>
|
||||
<figureCitation id="1315EAA14C3CC61D1A442CBBFB6F20DB" box="[1124,1212,406,431]" captionStart="FIGURE 23" captionStartId="76.[151,250,1428,1450]" captionTargetBox="[249,1339,207,1397]" captionTargetId="figure@76.[234,1353,193,1406]" captionTargetPageId="76" captionText="FIGURE 23. Adults in views stated. A: ♂ PT genitalia of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; 33 DL), LV, SV. Scale bar: 3 mm. B: ♂ genitalia and sternal androconial scaling in Ht. roussettae (“ sp. 26 ” modified from Lees, 1997; not to scale), LV. C: ♂ genitalia and sternal androconial scaling in Ht. andasibe (modified from Lees, 1997: where labelled “ antsianakana ”; not to scale), LV." figureDoi="http://doi.org/10.5281/zenodo.264620" httpUri="https://zenodo.org/record/264620/files/figure.png" pageId="74" pageNumber="75">Fig. 23</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C3CC61D1ACC2CBBFAA020DB" box="[1260,1395,406,431]" name="Madagascar" pageId="74" pageNumber="75">Madagascar</collectingCountry>
|
||||
N, Montagne d'Ambre, W of divide,
|
||||
|
|
@ -261,15 +261,15 @@ Montagne d'Ambre, ‘Les Roussettes’,
|
|||
With a white point emphasised in space-M2 and often M3 of the FW and in M1-M
|
||||
<quantity id="4CD65BC14C3CC61D1AD92FF1FAF72381" box="[1273,1316,732,757]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="74" pageNumber="75" unit="in" value="3.0">3 in</quantity>
|
||||
the HWV,
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1EB72E2DFEE12263" box="[151,306,768,791]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1EB72E2DFEE12263" box="[151,306,768,791]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C3CC61D1EB72E2DFEE12263" box="[151,306,768,791]" italics="true" pageId="74" pageNumber="75">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
looks very like
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1FC52E2DFDA0226C" box="[485,627,767,792]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1FC52E2DFDA0226C" box="[485,627,767,792]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3CC61D1FC52E2DFDA0226C" box="[485,627,767,792]" italics="true" pageId="74" pageNumber="75">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
. It is however allopatric to
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1D852E2DFBE0226C" box="[933,1075,767,792]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1D852E2DFBE0226C" box="[933,1075,767,792]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3CC61D1D852E2DFBE0226C" box="[933,1075,767,792]" italics="true" pageId="74" pageNumber="75">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
, as far as known confined to the northern rainforest of Montagne d’Ambre, and likewise it is allopatric to
|
||||
|
|
@ -280,30 +280,30 @@ looks very like
|
|||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1B0A2E0EFA4F224E" box="[1322,1436,803,826]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1B0A2E0EFA4F224E" box="[1322,1436,803,826]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="tianae">
|
||||
<emphasis id="B95A2A364C3CC61D1B0A2E0EFA4F224E" box="[1322,1436,803,826]" italics="true" pageId="74" pageNumber="75">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1EEC2E6BFEA1222A" box="[204,370,837,862]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="oberthueri" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1EEC2E6BFEA1222A" box="[204,370,837,862]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="oberthueri" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C3CC61D1EEC2E6BFEA1222A" box="[204,370,837,862]" italics="true" pageId="74" pageNumber="75">Ht. oberthueri</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C3CC61D1F5F2E6BFE0D2229" box="[383,478,838,861]" pageId="74" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, which like
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1C572E6BFCDF222A" box="[631,780,837,862]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1C572E6BFCDF222A" box="[631,780,837,862]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3CC61D1C572E6BFCDF222A" box="[631,780,837,862]" italics="true" pageId="74" pageNumber="75">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
are distributed at more central latitudes of the eastern rainforests.
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1F3F2E47FE1322F5" box="[287,448,874,897]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1F3F2E47FE1322F5" box="[287,448,874,897]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C3CC61D1F3F2E47FE1322F5" box="[287,448,874,897]" italics="true" pageId="74" pageNumber="75">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
has a smoother outcurve to the HWV Mb than those species at M3, and in known material there is a strong tendency to expression of an ocellus, albeit small, in space-M3 on the HWD (
|
||||
<figureCitation id="1315EAA14C3CC61D1B0D2EA1FAAC22D0" box="[1325,1407,908,933]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="74" pageNumber="75">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C3CC61D1B0D2EA1FAAC22D0" box="[1325,1407,908,933]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="74" pageNumber="75">Fig. 22</figureCitation>
|
||||
A- B); these features distinguish it from the other four generally similar species. In the ♂ genitalia from LV, compared to
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1E932EFEFE91229F" box="[179,322,978,1003]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1E932EFEFE91229F" box="[179,322,978,1003]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3CC61D1E932EFEFE91229F" box="[179,322,978,1003]" italics="true" pageId="74" pageNumber="75">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
, the tegumen has a relatively straight dorsal edge (and wide and shallow proximad notch from DV), appearing quite extended proximad and the uncus tip extends towards or slightly beyond maximum extension (distad lobe) of valve tips; the reliability of these distinguishing features is unknown. The HWV has a relatively straight Mb (compared to
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1FE42910FCD02520" authority="Mabille, 1877" authorityName="Mabille" authorityYear="1877" box="[452,771,1084,1109]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1FE42910FCD02520" authority="Mabille, 1877" authorityName="Mabille" authorityYear="1877" box="[452,771,1084,1109]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C3CC61D1FE42910FD942521" box="[452,583,1084,1109]" italics="true" pageId="74" pageNumber="75">Ht. strigula</emphasis>
|
||||
(Mabille, 1877)
|
||||
</taxonomicName>
|
||||
|
|
@ -316,15 +316,15 @@ and to
|
|||
<emphasis id="B95A2A364C3CC61D1FE9294DFD972503" box="[457,580,1120,1143]" italics="true" pageId="74" pageNumber="75">Ht. anceps</emphasis>
|
||||
</taxonomicName>
|
||||
and the light orange
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1D1E294DFBC2250C" box="[830,1041,1119,1144]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1D1E294DFBC2250C" box="[830,1041,1119,1144]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<emphasis id="B95A2A364C3CC61D1D1E294DFBC2250C" box="[830,1041,1119,1144]" italics="true" pageId="74" pageNumber="75">Ht. menamenoides</emphasis>
|
||||
</taxonomicName>
|
||||
described above, distinguishing it strongly from members of the
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1FCC29AEFD7225EF" box="[492,673,1154,1179]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1FCC29AEFD7225EF" box="[492,673,1154,1179]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<emphasis id="B95A2A364C3CC61D1FCC29AEFD7225EF" box="[492,673,1154,1179]" italics="true" pageId="74" pageNumber="75">Ht. angulifascia</emphasis>
|
||||
</taxonomicName>
|
||||
clade (
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1CD329AEFCA725EF" box="[755,884,1154,1179]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1CD329AEFCA725EF" box="[755,884,1154,1179]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C3CC61D1CD329AEFCA725EF" box="[755,884,1154,1179]" italics="true" pageId="74" pageNumber="75">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group).
|
||||
|
|
@ -334,11 +334,11 @@ group).
|
|||
<paragraph id="8B91F6244C3CC61D1EE72988FC7627DF" blockId="74.[151,1437,151,1989]" pageId="74" pageNumber="75">
|
||||
<emphasis id="B95A2A364C3CC61D1EE72988FE6625CA" bold="true" box="[199,437,1189,1214]" pageId="74" pageNumber="75">Description. Wings:</emphasis>
|
||||
Upperside uniform dark brown. FW space-CuA1 ocellus with black iris spanning veins CuA1 and CuA2; Orng not perfectly circular, tending to be angled towards base of vein CuA1. FW space-M1 ocellus small, with faint orange ring; M2 ocellus white pupil only evident. In HW, space-CuA1 ocellus elliptic, with narrow dark orange ring, spanning about 1/3 of vein CuA1-CuA2 distance; space-M3 ocellus expressed (unusually for a member of the
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1C24281EFD55243F" box="[516,646,1330,1355]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1C24281EFD55243F" box="[516,646,1330,1355]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C3CC61D1C24281EFD55243F" box="[516,646,1330,1355]" italics="true" pageId="74" pageNumber="75">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group), but smaller, less elliptic. White pupils as points only, in HW spaces-M1-M3 particularly. Underside has ochreous-olive brown cast. FW space-CuA1 ocellus with orange ring that is yellower towards costa tending to be blocked by vein M3, and forming Ore that follows concave curve of dark brown FWV Mb, which as usual curves back towards mid costa. Background colour is slightly yellower olivebrown distad of Mb. Mb in HW relatively straight, as in
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1D3E28EDFC3C24AC" box="[798,1007,1471,1496]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<taxonomicName id="4C2E8DA74C3CC61D1D3E28EDFC3C24AC" box="[798,1007,1471,1496]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="menamenoides">
|
||||
<emphasis id="B95A2A364C3CC61D1D3E28EDFC3C24AC" box="[798,1007,1471,1496]" italics="true" pageId="74" pageNumber="75">Ht. menamenoides</emphasis>
|
||||
</taxonomicName>
|
||||
, and there is a distinct dark brownish convex PMb. Fine irroration of small darker brown line fragments throughout wing and concentrated in basal area, with Cbs on FW comprising two sets of approximately parallel transverse lines in the FW cell area. Very wavy darker diffuse brown line runs submarginally in both wings especially FW (somewhat toothed outward at each vein) and narrow darker brown line tracks wing margin, which is fairly smooth, only very slightly crenate.
|
||||
|
|
@ -362,23 +362,23 @@ group), but smaller, less elliptic. White pupils as points only, in HW spaces-M1
|
|||
Aside from Sdb, dark dispersed Cub consists of brownish hairs emanating from above 1A+2A and 3A that may contact a blackish patch of specialized scales located on abdominal sternite A3–A5 (
|
||||
<bibRefCitation id="EFBF8BD54C3CC61D1B372A32FED8262F" author="Lees" pageId="74" pageNumber="75" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
|
||||
Lees, 1997: 99,
|
||||
<figureCitation id="1315EAA14C3CC61D1EE42A6FFED8262F" box="[196,267,1858,1883]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="74" pageNumber="75">Fig. 5</figureCitation>
|
||||
<figureCitation id="1315EAA14C3CC61D1EE42A6FFED8262F" box="[196,267,1858,1883]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264602" httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="74" pageNumber="75">Fig. 5</figureCitation>
|
||||
</bibRefCitation>
|
||||
). Note that an androconial abdominal patch in this position is not very evident, and does not appear distinctly from superficial examination. The cubital/anal brush(es) partially overlie two ‘balloon’ inflations (swellings) terminating around mid-vein in 1A+2A and 3A, tapering towards base, covered in very small thin grey scales, that in 3A are rather larger than that in 1A+2A (
|
||||
<bibRefCitation id="EFBF8BD54C3CC61D1D202A81FBD426B0" author="Lees" box="[768,1031,1964,1989]" pageId="74" pageNumber="75" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
|
||||
Lees 1997: 96,
|
||||
<figureCitation id="1315EAA14C3CC61D1D8E2A81FC3C26B0" box="[942,1007,1964,1989]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="74" pageNumber="75">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C3CC61D1D8E2A81FC3C26B0" box="[942,1007,1964,1989]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="74" pageNumber="75">Fig. 3</figureCitation>
|
||||
A
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C3DC61C1EB72A11FF29269F" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="75" pageNumber="76" targetBox="[240,1346,208,1812]" targetPageId="75">
|
||||
<caption id="DF51A6AC4C3DC61C1EB72A11FF29269F" ID-DOI="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="75" pageNumber="76" targetBox="[240,1346,208,1812]" targetPageId="75">
|
||||
<paragraph id="8B91F6244C3DC61C1EB72A11FF29269F" blockId="75.[151,1436,1852,2027]" pageId="75" pageNumber="76">
|
||||
<emphasis id="B95A2A364C3DC61C1EB72A11FEF12625" bold="true" box="[151,290,1852,1874]" pageId="75" pageNumber="76">FIGURE 22.</emphasis>
|
||||
Adults in DV (left), VV (right) for mounted specimens and adults in natural posture.
|
||||
<emphasis id="B95A2A364C3DC61C1A892A11FB682625" bold="true" box="[1193,1211,1852,1873]" pageId="75" pageNumber="76">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1B0F2A10FE6A2605" authority="Lees & Kremen" authorityName="Lees & Kremen" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="roussettae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1B0F2A10FE6A2605" authority="Lees & Kremen" authorityName="Lees & Kremen" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="roussettae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C3DC61C1B0F2A10FF2D2605" italics="true" pageId="75" pageNumber="76">Heteropsis roussettae</emphasis>
|
||||
Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -389,7 +389,7 @@ Lees & Kremen
|
|||
(Montagne d’Ambre; NHMUK010289154).
|
||||
<emphasis id="B95A2A364C3DC61C1A212A76FBC12604" bold="true" box="[1025,1042,1883,1904]" pageId="75" pageNumber="76">B</emphasis>
|
||||
: ♀ PT of
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1AAE2A71FAF22605" box="[1166,1313,1884,1905]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1AAE2A71FAF22605" box="[1166,1313,1884,1905]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C3DC61C1AAE2A71FAF22605" box="[1166,1313,1884,1905]" italics="true" pageId="75" pageNumber="76">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
(Montagne d’Ambre; NHMUK010289157).
|
||||
|
|
@ -400,7 +400,7 @@ Lees & Kremen
|
|||
Oberthür, 1916
|
||||
</taxonomicName>
|
||||
(‘Antsianaka’, 1890; BMNH(E) #674757; now
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1EEA2AB7FE0F26DA" authority="Lees (2003)" authorityName="Lees" authorityYear="2003" box="[202,476,1944,1967]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1EEA2AB7FE0F26DA" authority="Lees (2003)" authorityName="Lees" authorityYear="2003" box="[202,476,1944,1967]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3DC61C1EEA2AB7FE9D26DA" box="[202,334,1944,1967]" italics="true" pageId="75" pageNumber="76">Ht. andasibe</emphasis>
|
||||
Lees (2003))
|
||||
</taxonomicName>
|
||||
|
|
@ -414,27 +414,27 @@ Oberthür, 1916
|
|||
(‘Antsianaka’, 1893; BMNH(E) #674762). Scale bar: 30 mm.
|
||||
<emphasis id="B95A2A364C3DC61C1F372A9AFEFB26B8" bold="true" box="[279,296,1975,1996]" pageId="75" pageNumber="76">E</emphasis>
|
||||
: ♀ of
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1F482A95FE2626B9" box="[360,501,1976,1997]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1F482A95FE2626B9" box="[360,501,1976,1997]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C3DC61C1F482A95FE2626B9" box="[360,501,1976,1997]" italics="true" pageId="75" pageNumber="76">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
, in disturbed habitat (Montagne d’Ambre; 09/01/1995);
|
||||
<emphasis id="B95A2A364C3DC61C1A102A9AFBEC26B8" bold="true" box="[1072,1087,1975,1996]" pageId="75" pageNumber="76">F</emphasis>
|
||||
: ♀ of
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1AA02A95FAD326B9" box="[1152,1280,1975,1997]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1AA02A95FAD326B9" box="[1152,1280,1975,1997]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3DC61C1AA02A95FAD326B9" box="[1152,1280,1975,1997]" italics="true" pageId="75" pageNumber="76">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1B2D2A9AFAA226B9" box="[1293,1393,1975,1997]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="84" phylum="Arthropoda" rank="species" species="andasibe">Andasibe</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C3DC61C1B2D2A9AFAA226B9" box="[1293,1393,1975,1997]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="75" pageNumber="84" phylum="Arthropoda" rank="species" species="andasibe">Andasibe</taxonomicName>
|
||||
; 24/ 10/2014).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF51A6AC4C3AC61B1EB728B9FD952772" httpUri="https://zenodo.org/record/264620/files/figure.png" pageId="76" pageNumber="77" targetBox="[249,1339,207,1397]" targetPageId="76">
|
||||
<caption id="DF51A6AC4C3AC61B1EB728B9FD952772" ID-DOI="http://doi.org/10.5281/zenodo.264620" httpUri="https://zenodo.org/record/264620/files/figure.png" pageId="76" pageNumber="77" targetBox="[249,1339,207,1397]" targetPageId="76">
|
||||
<paragraph id="8B91F6244C3AC61B1EB728B9FD952772" blockId="76.[151,1436,1428,1542]" pageId="76" pageNumber="77">
|
||||
<emphasis id="B95A2A364C3AC61B1EB728B9FECC24DD" bold="true" box="[151,287,1428,1450]" pageId="76" pageNumber="77">FIGURE 23.</emphasis>
|
||||
Adults in views stated.
|
||||
<emphasis id="B95A2A364C3AC61B1C2F28B9FDF224DD" bold="true" box="[527,545,1428,1449]" pageId="76" pageNumber="77">A</emphasis>
|
||||
: ♂ PT genitalia of
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1CC228B8FBB624DE" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[738,1125,1428,1450]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="roussettae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1CC228B8FBB624DE" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[738,1125,1428,1450]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="roussettae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C3AC61B1CC228B8FC9C24DE" box="[738,847,1429,1450]" italics="true" pageId="76" pageNumber="77">Heteropsis</emphasis>
|
||||
<emphasis id="B95A2A364C3AC61B1D7528B8FC6F24DE" box="[853,956,1429,1450]" italics="true" pageId="76" pageNumber="77">roussettae</emphasis>
|
||||
Lees & Kremen
|
||||
|
|
@ -446,13 +446,13 @@ Lees & Kremen
|
|||
(Montagne d’Ambre; 33DL), LV, SV. Scale bar: 3 mm.
|
||||
<emphasis id="B95A2A364C3AC61B1FCA289EFE2824BC" bold="true" box="[490,507,1459,1480]" pageId="76" pageNumber="77">B</emphasis>
|
||||
: ♂ genitalia and sternal androconial scaling in
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1DF82899FBB624BD" box="[984,1125,1460,1481]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1DF82899FBB624BD" box="[984,1125,1460,1481]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C3AC61B1DF82899FBB624BD" box="[984,1125,1460,1481]" italics="true" pageId="76" pageNumber="77">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
(“sp. 26” modified from Lees, 1997; not to scale), LV.
|
||||
<emphasis id="B95A2A364C3AC61B1FB928FCFE782493" bold="true" box="[409,427,1489,1511]" pageId="76" pageNumber="77">C</emphasis>
|
||||
: ♂ genitalia and sternal androconial scaling in
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1D8428FFFBFA2493" box="[932,1065,1489,1511]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1D8428FFFBFA2493" box="[932,1065,1489,1511]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3AC61B1D8428FFFBFA2493" box="[932,1065,1489,1511]" italics="true" pageId="76" pageNumber="77">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
(modified from Lees, 1997: where labelled “
|
||||
|
|
@ -469,9 +469,9 @@ Lees & Kremen
|
|||
<paragraph id="8B91F6244C3AC61B1EE72B58FAFC2682" blockId="76.[151,1437,1581,2038]" pageId="76" pageNumber="77">
|
||||
<emphasis id="B95A2A364C3AC61B1EE72B58FE9827FA" bold="true" box="[199,331,1653,1678]" pageId="76" pageNumber="77">♂ genitalia</emphasis>
|
||||
: 33DL (
|
||||
<figureCitation id="1315EAA14C3AC61B1F902B58FDD027FA" box="[432,515,1653,1678]" captionStart="FIGURE 23" captionStartId="76.[151,250,1428,1450]" captionTargetBox="[249,1339,207,1397]" captionTargetId="figure@76.[234,1353,193,1406]" captionTargetPageId="76" captionText="FIGURE 23. Adults in views stated. A: ♂ PT genitalia of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; 33 DL), LV, SV. Scale bar: 3 mm. B: ♂ genitalia and sternal androconial scaling in Ht. roussettae (“ sp. 26 ” modified from Lees, 1997; not to scale), LV. C: ♂ genitalia and sternal androconial scaling in Ht. andasibe (modified from Lees, 1997: where labelled “ antsianakana ”; not to scale), LV." httpUri="https://zenodo.org/record/264620/files/figure.png" pageId="76" pageNumber="77">Fig. 23</figureCitation>
|
||||
<figureCitation id="1315EAA14C3AC61B1F902B58FDD027FA" box="[432,515,1653,1678]" captionStart="FIGURE 23" captionStartId="76.[151,250,1428,1450]" captionTargetBox="[249,1339,207,1397]" captionTargetId="figure@76.[234,1353,193,1406]" captionTargetPageId="76" captionText="FIGURE 23. Adults in views stated. A: ♂ PT genitalia of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; 33 DL), LV, SV. Scale bar: 3 mm. B: ♂ genitalia and sternal androconial scaling in Ht. roussettae (“ sp. 26 ” modified from Lees, 1997; not to scale), LV. C: ♂ genitalia and sternal androconial scaling in Ht. andasibe (modified from Lees, 1997: where labelled “ antsianakana ”; not to scale), LV." figureDoi="http://doi.org/10.5281/zenodo.264620" httpUri="https://zenodo.org/record/264620/files/figure.png" pageId="76" pageNumber="77">Fig. 23</figureCitation>
|
||||
A, PT): from LV, very similar to
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1DA22B5BFBC027FA" box="[898,1043,1653,1678]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1DA22B5BFBC027FA" box="[898,1043,1653,1678]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3AC61B1DA22B5BFBC027FA" box="[898,1043,1653,1678]" italics="true" pageId="76" pageNumber="77">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
(as “
|
||||
|
|
@ -481,12 +481,12 @@ A, PT): from LV, very similar to
|
|||
” in
|
||||
<bibRefCitation id="EFBF8BD54C3AC61B1B3E2B5BFA4B27FA" author="Lees" box="[1310,1432,1654,1678]" pageId="76" pageNumber="77" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
|
||||
: 107,
|
||||
<figureCitation id="1315EAA14C3AC61B1EF12BB4FEE127C6" box="[209,306,1689,1714]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="76" pageNumber="77">
|
||||
<figureCitation id="1315EAA14C3AC61B1EF12BB4FEE127C6" box="[209,306,1689,1714]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="76" pageNumber="77">
|
||||
Figure
|
||||
<quantity id="4CD65BC14C3AC61B1F032BB7FEE127C6" box="[291,306,1690,1714]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="76" pageNumber="77" unit="g" value="7.0">7</quantity>
|
||||
</figureCitation>
|
||||
g), but tegumen, which has fairly straight dorsal edge (and wide and shallow proximad notch from DV), appears quite extended proximad from LV and uncus tip extends towards or slightly beyond maximum extension (the distad lobe) of valve tip. Uncus slightly longer than tegumen and parallel sided from LV with distinct dorsal ‘head’ at tip; slightly inflated before tip from DV. Valves stout and asymmetric at base which also has a small and gently curved ‘shoulder’; dorsal valve margin is the much more recurved one, with a recurved spine facing proximad/mesad. As in
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1C632A63FD062612" box="[579,725,1869,1894]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3AC61B1C632A63FD062612" box="[579,725,1869,1894]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3AC61B1C632A63FD062612" box="[579,725,1869,1894]" italics="true" pageId="76" pageNumber="77">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
, the shallowly bifid ends of valves impart a somewhat ‘crab pincer’-like appearance, heavily armed with spinoid setae along wide, uncus-facing margin. Saccus not long and aedeagus similar length to valve, quite deep and strongly uprecurved distad. The gnathos emanates from a distinctly rectangular base, recurved downward as crossing the fairly narrow and parallel-necked uncus, at an angle of around 40 degrees in the specimen examined. The juxta is narrow proximad and slightly down-lipped.
|
||||
|
|
@ -502,7 +502,7 @@ Named after Les Roussettes (a name likely derived from
|
|||
, a genus of fruit bat; note that the derivation used here though is feminine singular from the French word ‘roussette’, which might approximate the root of both names). Les Roussettes is a rather famous touristic site at PN Montagne d’Ambre and when visited this former forestry station was provided with a ‘
|
||||
<emphasis id="B95A2A364C3BC61A1CEB2C28FD262068" box="[715,757,261,284]" italics="true" pageId="77" pageNumber="78">gite</emphasis>
|
||||
’ around which this low-flying species could be found. The name also reflects that of the closely related species
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1CC42C04FCA02034" box="[740,883,295,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1CC42C04FCA02034" box="[740,883,295,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3BC61A1CC42C04FCA02034" box="[740,883,295,320]" italics="true" pageId="77" pageNumber="78">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
, also named after a familiar tourist destination in
|
||||
|
|
@ -518,12 +518,12 @@ This species was first recognized in the field as “sp. 26” by Claire Kremen
|
|||
(two ♂♂). There is a single historical specimen of this species in PBZT included in the
|
||||
<typeStatus id="549548864C3BC61A1AA22CB8FB6120D9" box="[1154,1202,405,429]" pageId="77" pageNumber="78">type</typeStatus>
|
||||
series. All relevant
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1EB72C94FEDC20A4" box="[151,271,441,464]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1EB72C94FEDC20A4" box="[151,271,441,464]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C3BC61A1EB72C94FEDC20A4" box="[151,271,441,464]" italics="true" pageId="77" pageNumber="78">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
<typeStatus id="549548864C3BC61A1F372C95FE8120A4" box="[279,338,440,464]" pageId="77" pageNumber="78">types</typeStatus>
|
||||
in the
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1F812C94FDF120A4" box="[417,546,439,464]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1F812C94FDF120A4" box="[417,546,439,464]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C3BC61A1F812C94FDF120A4" box="[417,546,439,464]" italics="true" pageId="77" pageNumber="78">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group were examined to ensure recognition of this species as new. The former
|
||||
|
|
@ -538,7 +538,7 @@ were examined: LT ♂, according to the Code, was first designated by
|
|||
: 184) who illustrated a specimen of the first species with a red label next to it and a text which stated “♂ (
|
||||
<typeStatus id="549548864C3BC61A1F2F2F09FEA72349" box="[271,372,548,573]" pageId="77" pageNumber="78" type="holotype">holotype</typeStatus>
|
||||
) as illustrated” (
|
||||
<figureCitation id="1315EAA14C3BC61A1C0C2F09FDAC2348" box="[556,639,548,573]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="77" pageNumber="78">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C3BC61A1C0C2F09FDAC2348" box="[556,639,548,573]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="77" pageNumber="78">Fig. 22</figureCitation>
|
||||
C); bearing labels “BMNH(E) #674757;
|
||||
<collectingCountry id="F339B6B44C3BC61A1A652F09FB1F2349" box="[1093,1228,548,573]" name="Madagascar" pageId="77" pageNumber="78">Madagascar</collectingCountry>
|
||||
Antsianaka Perrot Freres 2e Semestre 1890”. Also examined were the STs of
|
||||
|
|
@ -547,7 +547,7 @@ Antsianaka Perrot Freres 2e Semestre 1890”. Also examined were the STs of
|
|||
<bibRefCitation id="EFBF8BD54C3BC61A1DC82F67FB422317" author="Oberthur" box="[1000,1169,586,611]" pageId="77" pageNumber="78" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
; LT ♂, here designated (
|
||||
<figureCitation id="1315EAA14C3BC61A1EBF2F5CFF2423FE" box="[159,247,625,650]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="77" pageNumber="78">Fig. 22</figureCitation>
|
||||
<figureCitation id="1315EAA14C3BC61A1EBF2F5CFF2423FE" box="[159,247,625,650]" captionStart="FIGURE 22" captionStartId="75.[151,250,1852,1874]" captionTargetBox="[240,1346,208,1812]" captionTargetId="figure@75.[223,1363,195,1828]" captionTargetPageId="75" captionText="FIGURE 22. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis roussettae Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289154). B: ♀ PT of Ht. roussettae (Montagne d’Ambre; NHMUK 010289157). C: LT ♂ of Culapa antsianakana Oberthür, 1916 (‘ Antsianaka’, 1890; BMNH (E) # 674757; now Ht. andasibe Lees (2003 )). D: LT ♂ of Culapa anceps Oberthür, 1916 (‘ Antsianaka’, 1893; BMNH (E) # 674762). Scale bar: 30 mm. E: ♀ of Ht. roussettae, in disturbed habitat (Montagne d’Ambre; 09 / 01 / 1995); F: ♀ of Ht. andasibe (Andasibe; 24 / 10 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264619" httpUri="https://zenodo.org/record/264619/files/figure.png" pageId="77" pageNumber="78">Fig. 22</figureCitation>
|
||||
D), specimen bearing labels “BMNH(E) #674762;
|
||||
<collectingCountry id="F339B6B44C3BC61A1D722F5CFC0A23FE" box="[850,985,625,650]" name="Madagascar" pageId="77" pageNumber="78">Madagascar</collectingCountry>
|
||||
Antsianaka Perrot Freres 2e Semestre 1893”. Regarding
|
||||
|
|
@ -593,7 +593,7 @@ Ward, 1870
|
|||
(HT ♂: “BMNH(E) 673767;
|
||||
<collectingCountry id="F339B6B44C3BC61A1CF72E40FC8D22F2" box="[727,862,877,902]" name="Madagascar" pageId="77" pageNumber="78">Madagascar</collectingCountry>
|
||||
Ex COLL. CHRIS WARD”). See also under
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1B592E43FF2322DE" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1B592E43FF2322DE" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C3BC61A1B592E43FF2322DE" italics="true" pageId="77" pageNumber="78">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
regarding
|
||||
|
|
@ -620,7 +620,7 @@ and
|
|||
no COI-5P barcode yet available. The cytochrome b sequence, 357 bp (BMNH(E) #671644,
|
||||
<accessionNumber id="947D6BC74C3BC61A1FD82ED3FDBB2562" box="[504,616,1022,1046]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819174" pageId="77" pageNumber="78">FJ819174</accessionNumber>
|
||||
) is about 4.75% divergent from
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1DD22ED3FB552562" box="[1010,1158,1021,1046]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1DD22ED3FB552562" box="[1010,1158,1021,1046]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3BC61A1DD22ED3FB552562" box="[1010,1158,1021,1046]" italics="true" pageId="77" pageNumber="78">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
(comparison sequences BMNH(E) #676684
|
||||
|
|
@ -652,11 +652,11 @@ found morphological support for a clade consisting of this species (“TWNSX”)
|
|||
(“
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1FEE29A3FD9225D2" box="[462,577,1166,1190]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="85" phylum="Arthropoda" rank="species" species="anceps">ANCEPS</taxonomicName>
|
||||
”) and
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1CB029A3FCF125D2" box="[656,802,1165,1190]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1CB029A3FCF125D2" box="[656,802,1165,1190]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3BC61A1CB029A3FCF125D2" box="[656,802,1165,1190]" italics="true" pageId="77" pageNumber="78">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
(“ANTSI”), with a topology retained in analysis of ♂ genitalic characters. Although it resembles more closely
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1D18299FFC1F25BE" box="[824,972,1201,1226]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<taxonomicName id="4C2E8DA74C3BC61A1D18299FFC1F25BE" box="[824,972,1201,1226]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="andasibe">
|
||||
<emphasis id="B95A2A364C3BC61A1D18299FFC1F25BE" box="[824,972,1201,1226]" italics="true" pageId="77" pageNumber="78">Ht. andasibe</emphasis>
|
||||
</taxonomicName>
|
||||
, either species might potentially be its allospecific replacement in Northern
|
||||
|
|
@ -697,7 +697,7 @@ Aduse-Poku
|
|||
<paragraph id="8B91F6244C3BC61A1EE728FCFA59249E" blockId="77.[151,1437,151,1697]" box="[199,1418,1489,1514]" pageId="77" pageNumber="78">
|
||||
<emphasis id="B95A2A364C3BC61A1EE728FCFE8A249E" bold="true" box="[199,345,1489,1514]" pageId="77" pageNumber="78">Distribution</emphasis>
|
||||
: only known from Montagne d’Ambre, in the vicinity of Les Roussettes (
|
||||
<figureCitation id="1315EAA14C3BC61A1AB428FCFB34249E" box="[1172,1255,1489,1514]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="77" pageNumber="78">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C3BC61A1AB428FCFB34249E" box="[1172,1255,1489,1514]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="77" pageNumber="78">Fig. 30</figureCitation>
|
||||
C. ochre dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C3BC61A1EE728D8FB75277A" blockId="77.[151,1437,151,1697]" box="[199,1190,1525,1550]" pageId="77" pageNumber="78">
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<mods:identifier id="AFE22B949A76AB5B7203A2DFAA8F7602" type="ZooBank">CFA586DA-10EE-468B-80EE-35351E3845FD</mods:identifier>
|
||||
<mods:classification id="90EE2D0F811AD93BD6432461F43DD3BE">journal article</mods:classification>
|
||||
<mods:identifier id="1D80886579A9758BE0D3937CFBD58312" type="CLB-Dataset">38930</mods:identifier>
|
||||
<mods:identifier id="12566EC861EB4589A0FBE237B3FAB095" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
|
||||
<mods:identifier id="947008AF9141DED1A85FF9704EF2A21F" type="GBIF-Dataset">a4e408c4-2575-4b5c-878b-e1b9010ec619</mods:identifier>
|
||||
<mods:identifier id="1BF70B3D6F83D607F57C891094427C4C" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="9E5E2A10EA27A7D95BE45AF1DA9D659A" type="Zenodo-Dep">264597</mods:identifier>
|
||||
<mods:identifier id="66A4D48E56A84EBEE82E3E01E99831AB" type="ZooBank">CFA586DA-10EE-468B-80EE-35351E3845FD</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="038747324C40C66E1EB72985FAE523DC" ID-DOI="http://doi.org/10.5281/zenodo.6086436" ID-GBIF-Taxon="120817128" ID-Zenodo-Dep="6086436" LSID="urn:lsid:plazi:treatment:038747324C40C66E1EB72985FAE523DC" httpUri="http://treatment.plazi.org/id/038747324C40C66E1EB72985FAE523DC" lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55">
|
||||
<treatment id="038747324C40C66E1EB72985FAE523DC" ID-DOI="http://doi.org/10.5281/zenodo.6086436" ID-GBIF-Taxon="120817128" ID-Zenodo-Dep="6086436" LSID="urn:lsid:plazi:treatment:038747324C40C66E1EB72985FAE523DC" httpUri="http://treatment.plazi.org/id/038747324C40C66E1EB72985FAE523DC" lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C40C6611EB72985FBD62590" pageId="54" pageNumber="55" type="nomenclature">
|
||||
<paragraph id="8B91F6244C40C6611EB72985FDD825B6" blockId="54.[151,1029,1192,1252]" box="[151,523,1192,1218]" pageId="54" pageNumber="55">
|
||||
<heading id="D0D941484C40C6611EB72985FDD825B6" bold="true" box="[151,523,1192,1218]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
|
||||
<emphasis id="B95A2A364C40C6611EB72985FDD825B6" bold="true" box="[151,523,1192,1218]" pageId="54" pageNumber="55">
|
||||
<taxonomicName id="4C2E8DA74C40C6611EB72985FE7325B6" authority="Lees" authorityName="Lees" box="[151,416,1192,1218]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="sogai" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C40C6611EB72985FE7325B6" authority="Lees" authorityName="Lees" box="[151,416,1192,1218]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="sogai" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C40C6611EB72985FE8D25B6" bold="true" box="[151,350,1192,1218]" italics="true" pageId="54" pageNumber="55">Heteropsis sogai</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -76,7 +76,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C40C6611CA52818FD21243A" box="[645,754,1333,1358]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C40C6611D0E2818FC57243A" box="[814,900,1333,1358]" captionStart="FIGURE 16" captionStartId="56.[151,250,1770,1792]" captionTargetBox="[242,1343,191,1739]" captionTargetId="figure@56.[234,1354,177,1752]" captionTargetPageId="56" captionText="FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014)." httpUri="https://zenodo.org/record/264613/files/figure.png" pageId="54" pageNumber="55">Fig. 16</figureCitation>
|
||||
<figureCitation id="1315EAA14C40C6611D0E2818FC57243A" box="[814,900,1333,1358]" captionStart="FIGURE 16" captionStartId="56.[151,250,1770,1792]" captionTargetBox="[242,1343,191,1739]" captionTargetId="figure@56.[234,1354,177,1752]" captionTargetPageId="56" captionText="FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264613" httpUri="https://zenodo.org/record/264613/files/figure.png" pageId="54" pageNumber="55">Fig. 16</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C40C6611D942818FBE8243A" box="[948,1083,1333,1358]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
|
||||
Nord, massif du Tsaratanana, Matsabory, en dessous de l’Andohanisambirano [
|
||||
|
|
@ -105,12 +105,12 @@ Deposition
|
|||
<collectionCode id="ED3F6EE14C40C6611FF42889FDFE24C9" box="[468,557,1444,1469]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
|
||||
</emphasis>
|
||||
: ♀ (
|
||||
<figureCitation id="1315EAA14C40C6611C402889FD6124C9" box="[608,690,1444,1469]" captionStart="FIGURE 16" captionStartId="56.[151,250,1770,1792]" captionTargetBox="[242,1343,191,1739]" captionTargetId="figure@56.[234,1354,177,1752]" captionTargetPageId="56" captionText="FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014)." httpUri="https://zenodo.org/record/264613/files/figure.png" pageId="54" pageNumber="55">Fig. 16</figureCitation>
|
||||
<figureCitation id="1315EAA14C40C6611C402889FD6124C9" box="[608,690,1444,1469]" captionStart="FIGURE 16" captionStartId="56.[151,250,1770,1792]" captionTargetBox="[242,1343,191,1739]" captionTargetId="figure@56.[234,1354,177,1752]" captionTargetPageId="56" captionText="FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264613" httpUri="https://zenodo.org/record/264613/files/figure.png" pageId="54" pageNumber="55">Fig. 16</figureCitation>
|
||||
B), data as HT but DCL-DB-3387;
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C40C6611EE728E5FAAB24AB" blockId="54.[151,1436,1299,2031]" box="[199,1400,1479,1504]" pageId="54" pageNumber="55">
|
||||
♂ (
|
||||
<figureCitation id="1315EAA14C40C6611ECB28EAFEE82494" box="[235,315,1479,1504]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="54" pageNumber="55">Fig. 15</figureCitation>
|
||||
<figureCitation id="1315EAA14C40C6611ECB28EAFEE82494" box="[235,315,1479,1504]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264612" httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="54" pageNumber="55">Fig. 15</figureCitation>
|
||||
C, genitalia), data as HT but DCL-DB-3388| 96
|
||||
<collectionCode id="ED3F6EE14C40C6611D6928EAFCBC2494" box="[841,879,1479,1504]" pageId="54" pageNumber="55">DL</collectionCode>
|
||||
[genitalia]; ♀, data as HT but DCL-DB-3389.
|
||||
|
|
@ -145,15 +145,15 @@ Nord, massif du Tsaratanana, Matsabory, en dessous de l’Andohanisambirano [est
|
|||
<subSubSection id="C334A5AF4C40C6611EE72B78FD1D2634" pageId="54" pageNumber="55" type="diagnosis">
|
||||
<paragraph id="8B91F6244C40C6611EE72B78FD1D2634" blockId="54.[151,1436,1299,2031]" pageId="54" pageNumber="55">
|
||||
<emphasis id="B95A2A364C40C6611EE72B78FE92271A" bold="true" box="[199,321,1621,1646]" pageId="54" pageNumber="55">Diagnosis.</emphasis>
|
||||
<taxonomicName id="4C2E8DA74C40C6611F692B7BFDE9271A" box="[329,570,1621,1646]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C40C6611F692B7BFDE9271A" box="[329,570,1621,1646]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C40C6611F692B7BFDE9271A" box="[329,570,1621,1646]" italics="true" pageId="54" pageNumber="55">Heteropsis subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C40C6611C542B7BFCD22719" box="[628,769,1622,1645]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C40C6611C542B7BFCD22719" box="[628,769,1622,1645]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C40C6611C542B7BFCD22719" box="[628,769,1622,1645]" italics="true" pageId="54" pageNumber="55">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
are very similar, ventrally variegated species, whereas
|
||||
<taxonomicName id="4C2E8DA74C40C6611B592B7BFED027E5" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C40C6611B592B7BFED027E5" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C40C6611B592B7BFED027E5" italics="true" pageId="54" pageNumber="55">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -161,23 +161,23 @@ and
|
|||
<emphasis id="B95A2A364C40C6611F182B57FE6327E5" box="[312,432,1658,1681]" italics="true" pageId="54" pageNumber="55">Ht. pauper</emphasis>
|
||||
</taxonomicName>
|
||||
tend to have the base of FWV costa orange. However, the HWV Mb is much more jagged in
|
||||
<taxonomicName id="4C2E8DA74C40C6611E962BB0FECC27C0" box="[182,287,1693,1716]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C40C6611E962BB0FECC27C0" box="[182,287,1693,1716]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C40C6611E962BB0FECC27C0" box="[182,287,1693,1716]" italics="true" pageId="54" pageNumber="55">Ht. sogai</emphasis>
|
||||
</taxonomicName>
|
||||
than any other species of the
|
||||
<taxonomicName id="4C2E8DA74C40C6611CA02BB0FCCF27C0" box="[640,796,1691,1716]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C40C6611CA02BB0FCCF27C0" box="[640,796,1691,1716]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C40C6611CA02BB0FCCF27C0" box="[640,796,1691,1716]" italics="true" pageId="54" pageNumber="55">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group, whereas
|
||||
<taxonomicName id="4C2E8DA74C40C6611DFD2BB0FBBF27C0" box="[989,1132,1693,1716]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C40C6611DFD2BB0FBBF27C0" box="[989,1132,1693,1716]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C40C6611DFD2BB0FBBF27C0" box="[989,1132,1693,1716]" italics="true" pageId="54" pageNumber="55">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
tends to have a yellowish ventral cast. HWV darker band of the central symmetry system (area between Mb and PMb) narrows more distinctly costad than in any other species of the
|
||||
<taxonomicName id="4C2E8DA74C40C6611CED2BCFFCB9278E" box="[717,874,1761,1786]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C40C6611CED2BCFFCB9278E" box="[717,874,1761,1786]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C40C6611CED2BCFFCB9278E" box="[717,874,1761,1786]" italics="true" pageId="54" pageNumber="55">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group. Two of FWV Cbs taper and meet tergad, demarcating pale ochreous area that is this relatively triangular rather than relatively oblong as in
|
||||
<taxonomicName id="4C2E8DA74C40C6611AC12A28FABE2668" box="[1249,1389,1797,1820]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C40C6611AC12A28FABE2668" box="[1249,1389,1797,1820]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C40C6611AC12A28FABE2668" box="[1249,1389,1797,1820]" italics="true" pageId="54" pageNumber="55">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
, for example. See below regarding androconial organs.
|
||||
|
|
@ -191,17 +191,17 @@ Upperside with rather uniform mid brown background colour. FWD space-CuA1 ocellu
|
|||
<paragraph id="8B91F6244C41C6601EE72E02FEEB22C5" blockId="55.[151,1437,151,2028]" pageId="55" pageNumber="56">
|
||||
<emphasis id="B95A2A364C41C6601EE72E02FE87223C" bold="true" box="[199,340,815,840]" pageId="55" pageNumber="56">Androconia</emphasis>
|
||||
: Sdb HWD dark brown but underlying patch has not been examined in detail. There is a not particularly bulbous anteriad inflation (but with 3–4 septa visible) in R near the anterior portion of the HWD cell, unlike for
|
||||
<taxonomicName id="4C2E8DA74C41C6601F2A2E5BFE7722FA" box="[266,420,885,910]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C41C6601F2A2E5BFE7722FA" box="[266,420,885,910]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C41C6601F2A2E5BFE7722FA" box="[266,420,885,910]" italics="true" pageId="55" pageNumber="56">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C41C6601FF92E5BFDB722F9" box="[473,612,886,909]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C41C6601FF92E5BFDB722F9" box="[473,612,886,909]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C41C6601FF92E5BFDB722F9" box="[473,612,886,909]" italics="true" pageId="55" pageNumber="56">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
(sp. “25”). M2-2A veins in ♂ HW are all linearly inflated (
|
||||
<bibRefCitation id="EFBF8BD54C41C6601AD62E5BFEF822C5" author="Lees" pageId="55" pageNumber="56" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="a">
|
||||
Lees 1997: 96,
|
||||
<figureCitation id="1315EAA14C41C6601EB72EB5FF0B22C5" box="[151,216,920,945]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="55" pageNumber="56">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C41C6601EB72EB5FF0B22C5" box="[151,216,920,945]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="55" pageNumber="56">Fig. 3</figureCitation>
|
||||
a, “39”
|
||||
</bibRefCitation>
|
||||
).
|
||||
|
|
@ -219,11 +219,11 @@ a, “39”
|
|||
<paragraph id="8B91F6244C41C6601EE7292CFCDD2403" blockId="55.[151,1437,151,2028]" pageId="55" pageNumber="56">
|
||||
<emphasis id="B95A2A364C41C6601EE7292CFE9F256E" bold="true" box="[199,332,1025,1050]" pageId="55" pageNumber="56">♂ genitalia</emphasis>
|
||||
: 96DL, PT, MNHN (Tsaratanana;
|
||||
<figureCitation id="1315EAA14C41C6601CFD292CFCE3256E" box="[733,816,1025,1050]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="55" pageNumber="56">Fig. 15</figureCitation>
|
||||
<figureCitation id="1315EAA14C41C6601CFD292CFCE3256E" box="[733,816,1025,1050]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264612" httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="55" pageNumber="56">Fig. 15</figureCitation>
|
||||
C). Small, about
|
||||
<quantity id="4CD65BC14C41C6601DD4292CFB9E256D" box="[1012,1101,1025,1050]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="55" pageNumber="56" unit="mm" value="1.6">1.6 mm</quantity>
|
||||
long, and with configuration typical of the
|
||||
<taxonomicName id="4C2E8DA74C41C6601F172908FE002549" box="[311,467,1060,1085]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C41C6601F172908FE002549" box="[311,467,1060,1085]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C41C6601F172908FE002549" box="[311,467,1060,1085]" italics="true" pageId="55" pageNumber="56">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group; no obvious difference in shape of the genitalic components from other group members (
|
||||
|
|
@ -260,21 +260,21 @@ in MNHN from specimens collected at Tsaratanana in 1966 by P. Griveaud
|
|||
). As a possible endemic of Tsaratanana, it is understandable that there is such little material of this species, all from this single collection in MNHN, except for one specimen in BMNH dating from before 1924, which is not included in the
|
||||
<typeStatus id="549548864C41C6601A1C2B5AFBBF27FB" box="[1084,1132,1655,1679]" pageId="55" pageNumber="56">type</typeStatus>
|
||||
series as it has a dubious locality label (see below). Due to its similarity to the relatively widespread
|
||||
<taxonomicName id="4C2E8DA74C41C6601DCF2BB7FB5927C6" box="[1007,1162,1689,1714]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C41C6601DCF2BB7FB5927C6" box="[1007,1162,1689,1714]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C41C6601DCF2BB7FB5927C6" box="[1007,1162,1689,1714]" italics="true" pageId="55" pageNumber="56">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
, it is also not surprising the species has not been described until now. All other available
|
||||
<typeStatus id="549548864C41C6601DBA2B90FC0627A1" box="[922,981,1725,1749]" pageId="55" pageNumber="56">types</typeStatus>
|
||||
of the
|
||||
<taxonomicName id="4C2E8DA74C41C6601A112B90FB1C27A1" box="[1073,1231,1724,1749]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C41C6601A112B90FB1C27A1" box="[1073,1231,1724,1749]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C41C6601A112B90FB1C27A1" box="[1073,1231,1724,1749]" italics="true" pageId="55" pageNumber="56">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group have been examined (see under
|
||||
<taxonomicName id="4C2E8DA74C41C6601FA82BCDFDCD2783" box="[392,542,1758,1783]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C41C6601FA82BCDFDCD2783" box="[392,542,1758,1783]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C41C6601FA82BCDFDCD2783" box="[392,542,1758,1783]" italics="true" pageId="55" pageNumber="56">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
). As well as the two ♂♂ and two ♀♀ of
|
||||
<taxonomicName id="4C2E8DA74C41C6601DD42BCDFB892783" box="[1012,1114,1760,1783]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C41C6601DD42BCDFB892783" box="[1012,1114,1760,1783]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C41C6601DD42BCDFB892783" box="[1012,1114,1760,1783]" italics="true" pageId="55" pageNumber="56">Ht. sogai</emphasis>
|
||||
</taxonomicName>
|
||||
from Tsaratanana, there is a single ♂ specimen labelled Beforo SE
|
||||
|
|
@ -282,7 +282,7 @@ from Tsaratanana, there is a single ♂ specimen labelled Beforo SE
|
|||
in BMNH. When I revisited the
|
||||
<typeStatus id="549548864C41C6601A192A2FFBBA266E" box="[1081,1129,1794,1818]" pageId="55" pageNumber="56">type</typeStatus>
|
||||
locality (now, of
|
||||
<taxonomicName id="4C2E8DA74C41C6601B0C2A2EFA40266E" box="[1324,1427,1795,1818]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C41C6601B0C2A2EFA40266E" box="[1324,1427,1795,1818]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C41C6601B0C2A2EFA40266E" box="[1324,1427,1795,1818]" italics="true" pageId="55" pageNumber="56">Ht. sogai</emphasis>
|
||||
</taxonomicName>
|
||||
) in
|
||||
|
|
@ -301,13 +301,13 @@ and no place name has been traced exactly as such (in any case, the similar soun
|
|||
</taxonomicName>
|
||||
bears the same locality and accession label.
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C4EC66F1EB72BC7FECA260E" httpUri="https://zenodo.org/record/264613/files/figure.png" pageId="56" pageNumber="57" targetBox="[242,1343,191,1739]" targetPageId="56">
|
||||
<caption id="DF51A6AC4C4EC66F1EB72BC7FECA260E" ID-DOI="http://doi.org/10.5281/zenodo.264613" httpUri="https://zenodo.org/record/264613/files/figure.png" pageId="56" pageNumber="57" targetBox="[242,1343,191,1739]" targetPageId="56">
|
||||
<paragraph id="8B91F6244C4EC66F1EB72BC7FECA260E" blockId="56.[151,1436,1770,1914]" pageId="56" pageNumber="57">
|
||||
<emphasis id="B95A2A364C4EC66F1EB72BC7FEF3278B" bold="true" box="[151,288,1770,1792]" pageId="56" pageNumber="57">FIGURE 16.</emphasis>
|
||||
Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D.C. Lees).
|
||||
<emphasis id="B95A2A364C4EC66F1B7A2BC7FABF278B" bold="true" box="[1370,1388,1770,1791]" pageId="56" pageNumber="57">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1EED2A24FEAB266A" box="[205,376,1801,1822]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="sogai" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1EED2A24FEAB266A" box="[205,376,1801,1822]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="sogai" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C4EC66F1EED2A24FEAB266A" box="[205,376,1801,1822]" italics="true" pageId="56" pageNumber="57">Heteropsis sogai</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
|
|
@ -317,13 +317,13 @@ Adults in DV (left), VV (right) for mounted specimens and adults in natural post
|
|||
(Tsaratanana; DCL-DB-3386).
|
||||
<emphasis id="B95A2A364C4EC66F1D3B2A25FCFF2669" bold="true" box="[795,812,1800,1821]" pageId="56" pageNumber="57">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1DB42A24FC21266A" box="[916,1010,1801,1822]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1DB42A24FC21266A" box="[916,1010,1801,1822]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C4EC66F1DB42A24FC21266A" box="[916,1010,1801,1822]" italics="true" pageId="56" pageNumber="57">Ht. sogai</emphasis>
|
||||
</taxonomicName>
|
||||
(Tsaratanana; DCL-DB-3387). Scale bar: 30 mm.
|
||||
<emphasis id="B95A2A364C4EC66F1ECF2A0AFED22649" bold="true" box="[239,257,1831,1853]" pageId="56" pageNumber="57">C</emphasis>
|
||||
: ♀ of
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1F6A2A05FDDD2649" authority="Lees" authorityName="Lees" box="[330,526,1831,1853]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="kremenae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1F6A2A05FDDD2649" authority="Lees" authorityName="Lees" box="[330,526,1831,1853]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="kremenae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C4EC66F1F6A2A05FEB92649" box="[330,362,1832,1853]" italics="true" pageId="56" pageNumber="57">Ht.</emphasis>
|
||||
<emphasis id="B95A2A364C4EC66F1F522A0AFE072649" box="[370,468,1831,1853]" italics="true" pageId="56" pageNumber="57">kremenae</emphasis>
|
||||
Lees
|
||||
|
|
@ -341,13 +341,13 @@ Lees
|
|||
(Vohitaly, Makira; 30/12/ 2002).
|
||||
<emphasis id="B95A2A364C4EC66F1EFF2A6BFF23262F" bold="true" box="[223,240,1862,1883]" pageId="56" pageNumber="57">E</emphasis>
|
||||
: ♀ of
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1F142A6AFE13262F" box="[308,448,1861,1884]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1F142A6AFE13262F" box="[308,448,1861,1884]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C4EC66F1F142A6AFE13262F" box="[308,448,1861,1884]" italics="true" pageId="56" pageNumber="57">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
(Ambohitantely; 6/12/2006);
|
||||
<emphasis id="B95A2A364C4EC66F1CD32A6BFCD1262F" bold="true" box="[755,770,1862,1883]" pageId="56" pageNumber="57">F</emphasis>
|
||||
: ♀ of
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1D662A6AFBA1262F" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[838,1138,1861,1884]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C4EC66F1D662A6AFBA1262F" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[838,1138,1861,1884]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C4EC66F1D662A6AFCB42628" box="[838,871,1863,1884]" italics="true" pageId="56" pageNumber="57">Ht.</emphasis>
|
||||
<emphasis id="B95A2A364C4EC66F1D4D2A6AFC152628" box="[877,966,1863,1884]" italics="true" pageId="56" pageNumber="57">avaratra</emphasis>
|
||||
Lees & Kremen
|
||||
|
|
@ -372,7 +372,7 @@ locality; and that in late 2013 of R. Ranaivosolo, pers. comm, to an adjacent ar
|
|||
: in the morphological analysis of
|
||||
<bibRefCitation id="EFBF8BD54C4FC66E1D4F2C28FC2B2068" author="Lees" box="[879,1016,260,284]" pageId="57" pageNumber="58" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997)</bibRefCitation>
|
||||
this taxon (“THTNN”) connected to
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1EB72C04FEF62034" box="[151,293,297,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1EB72C04FEF62034" box="[151,293,297,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C4FC66E1EB72C04FEF62034" box="[151,293,297,320]" italics="true" pageId="57" pageNumber="58">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
(“TWNFV”), which might possibly be its sister (as in combined tree in
|
||||
|
|
@ -409,7 +409,7 @@ for a marshy area or small lake within forest.
|
|||
<paragraph id="8B91F6244C4FC66E1EE72F09FE972314" blockId="57.[151,1436,151,680]" pageId="57" pageNumber="58">
|
||||
<emphasis id="B95A2A364C4FC66E1EE72F09FE8A2349" bold="true" box="[199,345,548,573]" pageId="57" pageNumber="58">Distribution</emphasis>
|
||||
: RNI Tsaratanana, where putatively endemic (
|
||||
<figureCitation id="1315EAA14C4FC66E1D542F09FC1A2348" box="[884,969,548,573]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="57" pageNumber="58">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C4FC66E1D542F09FC1A2348" box="[884,969,548,573]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="57" pageNumber="58">Fig. 30</figureCitation>
|
||||
D, mid green dots). See above regarding other localities.
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C4FC66E1EE72F41FD4323F0" blockId="57.[151,1436,151,680]" box="[199,656,620,645]" pageId="57" pageNumber="58">
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<treatment id="038747324C45C6611EB72DBAFED12521" ID-DOI="http://doi.org/10.5281/zenodo.6086434" ID-GBIF-Taxon="120817127" ID-Zenodo-Dep="6086434" LSID="urn:lsid:plazi:treatment:038747324C45C6611EB72DBAFED12521" httpUri="http://treatment.plazi.org/id/038747324C45C6611EB72DBAFED12521" lastPageId="54" lastPageNumber="54" pageId="51" pageNumber="52">
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<subSubSection id="C334A5AF4C45C6641EB72DBAFBC021A7" pageId="51" pageNumber="52" type="nomenclature">
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<emphasis id="B95A2A364C45C6641EB72DBAFD6C21C6" bold="true" box="[151,703,151,178]" pageId="51" pageNumber="52">
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<taxonomicName id="4C2E8DA74C45C6641EB72DBAFD8721C6" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,596,151,178]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C45C6641EB72DBAFD8721C6" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,596,151,178]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C45C6641EB72DBAFE5621C5" bold="true" box="[151,389,151,177]" italics="true" pageId="51" pageNumber="52">Heteropsis avaratra</emphasis>
|
||||
Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -83,7 +83,7 @@ material., Deposition
|
|||
:
|
||||
</emphasis>
|
||||
♂ (
|
||||
<figureCitation id="1315EAA14C45C6641D312C6FFCB3202E" box="[785,864,322,347]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="51" pageNumber="52">Fig. 13</figureCitation>
|
||||
<figureCitation id="1315EAA14C45C6641D312C6FFCB3202E" box="[785,864,322,347]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.264610" httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="51" pageNumber="52">Fig. 13</figureCitation>
|
||||
C),
|
||||
<collectingCountry id="F339B6B44C45C6641DAB2C6FFBC1202F" box="[907,1042,322,347]" name="Madagascar" pageId="51" pageNumber="52">Madagascar</collectingCountry>
|
||||
N, Montagne d'Ambre,
|
||||
|
|
@ -220,7 +220,7 @@ NORD MONTAGNE D’AMBRE LES ROUSSETTES
|
|||
<quantity id="4CD65BC14C45C6641F712E6AFE78222B" box="[337,427,839,863]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="52" unit="m" value="1000.0">1000 m</quantity>
|
||||
<date id="FF90D0E44C45C6641F972E6AFDE32214" box="[439,560,839,864]" pageId="51" pageNumber="52" value="1970-05-25">25-V-1970</date>
|
||||
P. GRIVEAUD|DCL-DB-3381; ♀ [MNHN] (
|
||||
<figureCitation id="1315EAA14C45C6641A6E2E6AFB77222B" box="[1102,1188,839,864]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="51" pageNumber="52">Fig. 13</figureCitation>
|
||||
<figureCitation id="1315EAA14C45C6641A6E2E6AFB77222B" box="[1102,1188,839,864]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.264610" httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="51" pageNumber="52">Fig. 13</figureCitation>
|
||||
D),
|
||||
<collectingCountry id="F339B6B44C45C6641AF52E6AFA4F222B" box="[1237,1436,839,863]" name="Madagascar" pageId="51" pageNumber="52">MADAGASCAR</collectingCountry>
|
||||
NORD Montagne d’Ambre Les Roussettes alt.
|
||||
|
|
@ -275,16 +275,16 @@ N, Montagne d'Ambre,
|
|||
<emphasis id="B95A2A364C45C6641EE7290EFE922548" bold="true" box="[199,321,1059,1084]" pageId="51" pageNumber="52">Diagnosis.</emphasis>
|
||||
compared with
|
||||
<emphasis id="B95A2A364C45C6641FDD2909FCEB2548" box="[509,824,1059,1084]" italics="true" pageId="51" pageNumber="52">
|
||||
<taxonomicName id="4C2E8DA74C45C6641FDD2909FD5C254F" box="[509,655,1060,1083]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">Ht. avaratra</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C45C6641FDD2909FD5C254F" box="[509,655,1060,1083]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">Ht. avaratra</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C45C6641CBC2909FCEB2548" box="[668,824,1059,1084]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="subsimilis">Ht. subsimilis</taxonomicName>
|
||||
</emphasis>
|
||||
has the Rs ocellus on HWV usually quite prominent and
|
||||
<taxonomicName id="4C2E8DA74C45C6641EE82965FEE2252B" box="[200,305,1096,1119]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C45C6641EE82965FEE2252B" box="[200,305,1096,1119]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C45C6641EE82965FEE2252B" box="[200,305,1096,1119]" italics="true" pageId="51" pageNumber="52">Ht. sogai</emphasis>
|
||||
</taxonomicName>
|
||||
has the Mb and PMb strongly dentate between M1 and CuA2. Populations of
|
||||
<taxonomicName id="4C2E8DA74C45C6641AE32965FA82252B" box="[1219,1361,1096,1119]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C45C6641AE32965FA82252B" box="[1219,1361,1096,1119]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C45C6641AE32965FA82252B" box="[1219,1361,1096,1119]" italics="true" pageId="51" pageNumber="52">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
in the northern reaches of the main rainforest belt can be hard to distinguish by wing pattern from the sympatric
|
||||
|
|
@ -292,19 +292,19 @@ in the northern reaches of the main rainforest belt can be hard to distinguish b
|
|||
<emphasis id="B95A2A364C45C6641B592941FED425D3" italics="true" pageId="51" pageNumber="52">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
though easily discriminated by DNA barcoding, but
|
||||
<taxonomicName id="4C2E8DA74C45C6641D4E29A2FC2F25D2" box="[878,1020,1167,1190]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C45C6641D4E29A2FC2F25D2" box="[878,1020,1167,1190]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C45C6641D4E29A2FC2F25D2" box="[878,1020,1167,1190]" italics="true" pageId="51" pageNumber="52">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
lacks an inflated M1-M2 vein (
|
||||
<figureCitation id="1315EAA14C45C6641B4F29A3FF6725BE" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="51" pageNumber="52">Fig. 12</figureCitation>
|
||||
<figureCitation id="1315EAA14C45C6641B4F29A3FF6725BE" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." figureDoi="http://doi.org/10.5281/zenodo.264609" httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="51" pageNumber="52">Fig. 12</figureCitation>
|
||||
E), and the inflated vein below the Sdb basad of the fork of Rs+M1 is more symmetric (more rounded on the anteriad edge) in
|
||||
<taxonomicName id="4C2E8DA74C45C6641F4029FAFE2F259A" box="[352,508,1237,1262]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C45C6641F4029FAFE2F259A" box="[352,508,1237,1262]" italics="true" pageId="51" pageNumber="52">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C45C6641C2E29F8FDB2259A" box="[526,609,1237,1262]" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="51" pageNumber="52">Fig. 12</figureCitation>
|
||||
<figureCitation id="1315EAA14C45C6641C2E29F8FDB2259A" box="[526,609,1237,1262]" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." figureDoi="http://doi.org/10.5281/zenodo.264609" httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="51" pageNumber="52">Fig. 12</figureCitation>
|
||||
D).
|
||||
<taxonomicName id="4C2E8DA74C45C6641CAD29FAFCC8259A" box="[653,795,1239,1262]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C45C6641CAD29FAFCC8259A" box="[653,795,1239,1262]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C45C6641CAD29FAFCC8259A" box="[653,795,1239,1262]" italics="true" pageId="51" pageNumber="52">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
, especially from the topotypical population, has a more prominently ventral yellow cast than in other species including
|
||||
|
|
@ -312,7 +312,7 @@ D).
|
|||
<emphasis id="B95A2A364C45C6641D4029D6FC292466" box="[864,1018,1273,1298]" italics="true" pageId="51" pageNumber="52">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
, and is not variegated with orange as in
|
||||
<taxonomicName id="4C2E8DA74C45C6641E942833FE9A2442" box="[180,329,1309,1334]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C45C6641E942833FE9A2442" box="[180,329,1309,1334]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C45C6641E942833FE9A2442" box="[180,329,1309,1334]" italics="true" pageId="51" pageNumber="52">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
|
|
@ -330,7 +330,7 @@ group. FWD space-CuA1 ocellus with black iris at least spanning CuA1-CuA2, with
|
|||
<emphasis id="B95A2A364C45C6641B592B35FED42720" italics="true" pageId="51" pageNumber="52">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group, and the area between Mb and PMb is finely irrorated in brown to a greater extent than other parts of the wing. FWV space-CuA1 ocellus with black iris same size as upperside and ring fairly subhexagonal and mainly yellow with gradation to orange towards tergal edge. The ring is slightly more pointed towards proximad edge where it hugs a strongly concave rufous brown Mb before it bends back to mid costa and Ore as a yellow hook like the small tail of a comma, representing the end of a spiral arm. FWV space-M1 ocellus as on upperside, slightly more conspicuous with a yellowish ring of background colour. On HWV, space-CuA1 ocellus is the largest as usual and subelliptic and including the concentric yellow ring spanning about 4/5 of veins CuA1-CuA2 (generally smaller than in
|
||||
<taxonomicName id="4C2E8DA74C45C6641F9F2A1AFD85263A" box="[447,598,1845,1870]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C45C6641F9F2A1AFD85263A" box="[447,598,1845,1870]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C45C6641F9F2A1AFD85263A" box="[447,598,1845,1870]" italics="true" pageId="51" pageNumber="52">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
). HWV space-CuA2 ocellus not evident in the HT ♂. HWV space Rs-M3 ocellus reduced to white point often with narrow black iris and Orng blending with background. The FWV cell area has two unequal brown edged more yellow transverse Cbs, the distad one wider. There is no distinctly orange shoulder to the basal FWV costa, which is strigulated evenly with black. The HWV is highlighted with coronal-like yellow-ochreous flares distad of the Mb, particularly in M2, where the Mb is toothed outwards slightly at vein intersections M2, M3 and CuA1, inflexed proximad of ocellus space-CuA1 and bends back towards 1A.
|
||||
|
|
@ -340,11 +340,11 @@ group, and the area between Mb and PMb is finely irrorated in brown to a greater
|
|||
<emphasis id="B95A2A364C42C6631FB72DCDFDE7218C" box="[407,564,223,248]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
, but
|
||||
<taxonomicName id="4C2E8DA74C42C6631C532DCDFCD22183" box="[627,769,224,247]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C42C6631C532DCDFCD22183" box="[627,769,224,247]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C42C6631C532DCDFCD22183" box="[627,769,224,247]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
tends to be a larger species (see also
|
||||
<figureCitation id="1315EAA14C42C6631A9F2DF2FAC1218C" box="[1215,1298,223,248]" captionStart="FIGURE 16" captionStartId="56.[151,250,1770,1792]" captionTargetBox="[242,1343,191,1739]" captionTargetId="figure@56.[234,1354,177,1752]" captionTargetPageId="56" captionText="FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014)." httpUri="https://zenodo.org/record/264613/files/figure.png" pageId="52" pageNumber="53">Fig. 16</figureCitation>
|
||||
<figureCitation id="1315EAA14C42C6631A9F2DF2FAC1218C" box="[1215,1298,223,248]" captionStart="FIGURE 16" captionStartId="56.[151,250,1770,1792]" captionTargetBox="[242,1343,191,1739]" captionTargetId="figure@56.[234,1354,177,1752]" captionTargetPageId="56" captionText="FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014)." figureDoi="http://doi.org/10.5281/zenodo.264613" httpUri="https://zenodo.org/record/264613/files/figure.png" pageId="52" pageNumber="53">Fig. 16</figureCitation>
|
||||
F). Mb well marked and sometimes more jagged-slightly inflexed than in HT, with yellow Mf in space-M2 distad of the Mb sometimes rectangular, although this character is not distinct in the HT specimen. HWV space-CuA2 ocellus pointlike in some specimens, or subelliptic with a pale yellow ring. HWV space-Rs-M3 ocelli usually but not always point-like with that of HWV space-Rs sometimes slightly more evident, as may be that of space-CuA2. FWV Cbs sometimes delineate indistinctly a third yellow band.
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C42C6631EE72C98FC052362" blockId="52.[151,1437,151,2035]" pageId="52" pageNumber="53">
|
||||
|
|
@ -378,9 +378,9 @@ F). Mb well marked and sometimes more jagged-slightly inflexed than in HT, with
|
|||
: 315DL (referred specimen, Joffreville);
|
||||
<bibRefCitation id="EFBF8BD54C42C6631A602E31FED52223" author="Lees" pageId="52" pageNumber="53" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="f">
|
||||
Lees (1997: 106,
|
||||
<figureCitation id="1315EAA14C42C6631B252E36FA942240" box="[1285,1351,795,820]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="52" pageNumber="53">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C42C6631B252E36FA942240" box="[1285,1351,795,820]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="52" pageNumber="53">Fig. 7</figureCitation>
|
||||
f; “25”;
|
||||
<figureCitation id="1315EAA14C42C6631EB72E13FF3B2223" box="[151,232,830,855]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="52" pageNumber="53">Fig. 15</figureCitation>
|
||||
<figureCitation id="1315EAA14C42C6631EB72E13FF3B2223" box="[151,232,830,855]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264612" httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="52" pageNumber="53">Fig. 15</figureCitation>
|
||||
B)
|
||||
</bibRefCitation>
|
||||
. Miniaturised, about
|
||||
|
|
@ -426,7 +426,7 @@ Torres
|
|||
(2001)
|
||||
</bibRefCitation>
|
||||
shows 5.27% pairwise divergence (398 bp compared) of
|
||||
<taxonomicName id="4C2E8DA74C42C6631C582875FCD7241B" box="[632,772,1368,1391]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C42C6631C582875FCD7241B" box="[632,772,1368,1391]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C42C6631C582875FCD7241B" box="[632,772,1368,1391]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
(‘sp. 25’) to
|
||||
|
|
@ -440,16 +440,16 @@ shows 5.27% pairwise divergence (398 bp compared) of
|
|||
: in the morphological analysis of
|
||||
<bibRefCitation id="EFBF8BD54C42C6631D532856FC3F24E7" author="Lees" box="[883,1004,1403,1427]" pageId="52" pageNumber="53" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
|
||||
, this taxon (“TWNFV”) connected to
|
||||
<taxonomicName id="4C2E8DA74C42C6631EB728B2FE8424C2" box="[151,343,1439,1462]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C42C6631EB728B2FE8424C2" box="[151,343,1439,1462]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C42C6631EB728B2FE8424C2" box="[151,343,1439,1462]" italics="true" pageId="52" pageNumber="53">Heteropsis sogai</emphasis>
|
||||
</taxonomicName>
|
||||
(“THRNN”) but in only one (jackknife) tree with support (
|
||||
<bibRefCitation id="EFBF8BD54C42C6631A3B28B2FAFE24C3" author="Lees" box="[1051,1325,1438,1463]" pageId="52" pageNumber="53" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
|
||||
Lees 1997: 157,
|
||||
<figureCitation id="1315EAA14C42C6631AC528B3FAFE24C3" box="[1253,1325,1438,1463]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="52" pageNumber="53">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C42C6631AC528B3FAFE24C3" box="[1253,1325,1438,1463]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="52" pageNumber="53">Fig. 2</figureCitation>
|
||||
</bibRefCitation>
|
||||
). In their analysis of COII sequences (where
|
||||
<taxonomicName id="4C2E8DA74C42C6631C0828EEFD5C24AE" box="[552,655,1475,1498]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C42C6631C0828EEFD5C24AE" box="[552,655,1475,1498]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C42C6631C0828EEFD5C24AE" box="[552,655,1475,1498]" italics="true" pageId="52" pageNumber="53">Ht. sogai</emphasis>
|
||||
</taxonomicName>
|
||||
was the only member of the
|
||||
|
|
@ -465,7 +465,7 @@ Torres
|
|||
(2001
|
||||
</bibRefCitation>
|
||||
: 366) had a meager 64% bootstrap support for a sister relation of
|
||||
<taxonomicName id="4C2E8DA74C42C6631AEE28CAFA89248A" box="[1230,1370,1511,1534]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C42C6631AEE28CAFA89248A" box="[1230,1370,1511,1534]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C42C6631AEE28CAFA89248A" box="[1230,1370,1511,1534]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
, their “
|
||||
|
|
@ -487,7 +487,7 @@ AY
|
|||
<date id="FF90D0E44C42C6631AFE2B24FAE12756" box="[1246,1330,1545,1570]" pageId="52" pageNumber="53" value="45-04-01">040145</date>
|
||||
</accessionNumber>
|
||||
based on one individual from Ranomafana National Park).
|
||||
<taxonomicName id="4C2E8DA74C42C6631CF32B03FCB12731" box="[723,866,1582,1605]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C42C6631CF32B03FCB12731" box="[723,866,1582,1605]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C42C6631CF32B03FCB12731" box="[723,866,1582,1605]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
(as “sp. 25”) was however strongly supported as sister to
|
||||
|
|
@ -501,9 +501,9 @@ Linares
|
|||
(2009
|
||||
</bibRefCitation>
|
||||
, p. 488, their
|
||||
<figureCitation id="1315EAA14C42C6631A872B7CFB38271D" box="[1191,1259,1617,1642]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="52" pageNumber="53">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C42C6631A872B7CFB38271D" box="[1191,1259,1617,1642]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="52" pageNumber="53">Fig. 2</figureCitation>
|
||||
) including COI or also EF-1a and wingless (p. 490, their
|
||||
<figureCitation id="1315EAA14C42C6631C4D2B59FD6127F9" box="[621,690,1652,1677]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="52" pageNumber="53">Fig. 4</figureCitation>
|
||||
<figureCitation id="1315EAA14C42C6631C4D2B59FD6127F9" box="[621,690,1652,1677]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." figureDoi="http://doi.org/10.5281/zenodo.264601" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="52" pageNumber="53">Fig. 4</figureCitation>
|
||||
). However, neither the latter study nor that of
|
||||
<bibRefCitation id="EFBF8BD54C42C6631AE02B59FEDC27C5" pageId="52" pageNumber="53" refString="Kodandaramaiah, U., Lees, D. C., Muller, C. J., Torres, E., Karanth, C. P. & Wahlberg, N. (2010) Phylogenetics and biogeography of a spectacular Old World radiation of butterflies: the subtribe Mycalesina (Lepidoptera: Nymphalidae: Satyrini). BMC Evolutionary Biology, 10, 172. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 10 - 172" type="journal article">
|
||||
Kodandaramaiah
|
||||
|
|
@ -511,11 +511,11 @@ Kodandaramaiah
|
|||
(2010)
|
||||
</bibRefCitation>
|
||||
, which found the same relationship, included
|
||||
<taxonomicName id="4C2E8DA74C42C6631D302BB4FC7527C5" box="[784,934,1688,1713]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C42C6631D302BB4FC7527C5" box="[784,934,1688,1713]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C42C6631D302BB4FC7527C5" box="[784,934,1688,1713]" italics="true" pageId="52" pageNumber="53">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
nor
|
||||
<taxonomicName id="4C2E8DA74C42C6631DF92BB4FB9327C4" box="[985,1088,1689,1712]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C42C6631DF92BB4FB9327C4" box="[985,1088,1689,1712]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C42C6631DF92BB4FB9327C4" box="[985,1088,1689,1712]" italics="true" pageId="52" pageNumber="53">Ht. sogai</emphasis>
|
||||
</taxonomicName>
|
||||
(but that of
|
||||
|
|
@ -525,7 +525,7 @@ Torres
|
|||
2001
|
||||
</bibRefCitation>
|
||||
, who found the same with COII, did include
|
||||
<taxonomicName id="4C2E8DA74C42C6631CA02B90FCC627A1" box="[640,789,1724,1749]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C42C6631CA02B90FCC627A1" box="[640,789,1724,1749]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C42C6631CA02B90FCC627A1" box="[640,789,1724,1749]" italics="true" pageId="52" pageNumber="53">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
as their ‘
|
||||
|
|
@ -539,15 +539,15 @@ was included among the
|
|||
<emphasis id="B95A2A364C42C6631F902BCCFD99278C" box="[432,586,1759,1784]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group.
|
||||
<taxonomicName id="4C2E8DA74C42C6631C822BCCFCEB278C" box="[674,824,1759,1784]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C42C6631C822BCCFCEB278C" box="[674,824,1759,1784]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C42C6631C822BCCFCEB278C" box="[674,824,1759,1784]" italics="true" pageId="52" pageNumber="53">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
is more distant to
|
||||
<taxonomicName id="4C2E8DA74C42C6631A2B2BCCFB44278C" box="[1035,1175,1761,1784]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C42C6631A2B2BCCFB44278C" box="[1035,1175,1761,1784]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C42C6631A2B2BCCFB44278C" box="[1035,1175,1761,1784]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
by COII data (see also description above).
|
||||
<taxonomicName id="4C2E8DA74C42C6631F542A28FE092668" box="[372,474,1797,1820]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C42C6631F542A28FE092668" box="[372,474,1797,1820]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C42C6631F542A28FE092668" box="[372,474,1797,1820]" italics="true" pageId="52" pageNumber="53">Ht. sogai</emphasis>
|
||||
</taxonomicName>
|
||||
might be its sister (as in combined tree in
|
||||
|
|
@ -579,7 +579,7 @@ group have been examined and are clearly distinct with their
|
|||
localities further south in
|
||||
<collectingCountry id="F339B6B44C42C6631C412A9BFD3B26BB" box="[609,744,1974,1999]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
|
||||
(see under
|
||||
<taxonomicName id="4C2E8DA74C42C6631D4A2A9AFBD326BB" box="[874,1024,1974,1999]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C42C6631D4A2A9AFBD326BB" box="[874,1024,1974,1999]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C42C6631D4A2A9AFBD326BB" box="[874,1024,1974,1999]" italics="true" pageId="52" pageNumber="53">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
|
|
@ -596,7 +596,7 @@ localities further south in
|
|||
<paragraph id="8B91F6244C43C6621EE72D96FA562068" blockId="53.[151,1437,151,2031]" pageId="53" pageNumber="54">
|
||||
<emphasis id="B95A2A364C43C6621EE72D96FE9021A0" bold="true" box="[199,323,187,212]" pageId="53" pageNumber="54">Behaviour</emphasis>
|
||||
: unusually for
|
||||
<taxonomicName id="4C2E8DA74C43C6621FD82D90FDA321A0" box="[504,624,189,212]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C43C6621FD82D90FDA321A0" box="[504,624,189,212]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C43C6621FD82D90FDA321A0" box="[504,624,189,212]" italics="true" pageId="53" pageNumber="54">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
, adults have been seen visiting
|
||||
|
|
@ -619,14 +619,14 @@ flowers in Montagne d’Ambre (pers. obs.). Adults fly low and would usually fee
|
|||
<paragraph id="8B91F6244C43C6621EE72C43FD8723BC" blockId="53.[151,1437,151,2031]" pageId="53" pageNumber="54">
|
||||
<emphasis id="B95A2A364C43C6621EE72C43FE8A20F3" bold="true" box="[199,345,366,391]" pageId="53" pageNumber="54">Distribution</emphasis>
|
||||
: Montagne d’Ambre is an important locality but the species occurs in rainforest also in the main rainforest block from Tsaratanana to Makira (
|
||||
<figureCitation id="1315EAA14C43C6621CBC2CBFFD2320DE" box="[668,752,402,427]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="53" pageNumber="54">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C43C6621CBC2CBFFD2320DE" box="[668,752,402,427]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="53" pageNumber="54">Fig. 30</figureCitation>
|
||||
B, red dots). If
|
||||
<emphasis id="B95A2A364C43C6621DBD2CBEFBD820DE" box="[925,1035,403,426]" italics="true" pageId="53" pageNumber="54">
|
||||
<taxonomicName id="4C2E8DA74C43C6621DBD2CBEFBD420DE" box="[925,1031,403,426]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sogai">Ht. sogai</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C43C6621DBD2CBEFBD420DE" box="[925,1031,403,426]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sogai">Ht. sogai</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
not sequenced, is not more closely related,
|
||||
<taxonomicName id="4C2E8DA74C43C6621ED52C9AFE5720BA" box="[245,388,439,462]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C43C6621ED52C9AFE5720BA" box="[245,388,439,462]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C43C6621ED52C9AFE5720BA" box="[245,388,439,462]" italics="true" pageId="53" pageNumber="54">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
might represent
|
||||
|
|
@ -634,7 +634,7 @@ might represent
|
|||
<emphasis id="B95A2A364C43C6621C6C2C9AFD3920BA" box="[588,746,437,462]" italics="true" pageId="53" pageNumber="54">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
’s replacement in Montagne d’Ambre, and in that case,
|
||||
<taxonomicName id="4C2E8DA74C43C6621B592C9AFF2A2085" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C43C6621B592C9AFF2A2085" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C43C6621B592C9AFF2A2085" italics="true" pageId="53" pageNumber="54">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
has achieved sympatry and secondary contact with
|
||||
|
|
@ -656,14 +656,14 @@ assigned a series of specimens in BMNH from Mahassabe [= Mahasoabe], forêt SE F
|
|||
<emphasis id="B95A2A364C43C6621D632F44FC3123F5" box="[835,994,616,641]" italics="true" pageId="53" pageNumber="54">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
or another species (plus this location includes a range of material that seems more characteristic of northeastern localities, for example
|
||||
<taxonomicName id="4C2E8DA74C43C6621B392FA0FE8523BC" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="erebina">
|
||||
<taxonomicName id="4C2E8DA74C43C6621B392FA0FE8523BC" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="erebina">
|
||||
<emphasis id="B95A2A364C43C6621B392FA0FA4F23D1" box="[1305,1436,652,677]" italics="true" pageId="53" pageNumber="54">Ht. erebina</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C43C6621EBE2F82FE9D23BC" author="Oberthur" box="[158,334,687,712]" pageId="53" pageNumber="54" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
; see also
|
||||
<taxonomicName id="4C2E8DA74C43C6621FE12F9CFD9723BC" box="[449,580,687,712]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C43C6621FE12F9CFD9723BC" box="[449,580,687,712]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C43C6621FE12F9CFD9723BC" box="[449,580,687,712]" italics="true" pageId="53" pageNumber="54">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
|
|
@ -1034,7 +1034,7 @@ extract number],
|
|||
; D.C. Lees: DL-05-183, IA205 [isotope voucher]; ♂ [
|
||||
<collectionCode id="ED3F6EE14C40C6611FC52DC5FDED218B" box="[485,574,232,255]" pageId="54" pageNumber="55">BMNH</collectionCode>
|
||||
] (
|
||||
<figureCitation id="1315EAA14C40C6611C762DCAFD792074" box="[598,682,231,256]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="54" pageNumber="55">Fig. 15</figureCitation>
|
||||
<figureCitation id="1315EAA14C40C6611C762DCAFD792074" box="[598,682,231,256]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264612" httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="54" pageNumber="55">Fig. 15</figureCitation>
|
||||
B), N, Joffreville [ca.
|
||||
<geoCoordinate id="EE1A90E34C40C6611D882DC5FBDE2074" box="[936,1037,228,256]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="55" value="-12.483">12.483o S</geoCoordinate>
|
||||
,
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<treatment id="038747324C48C6161EB728DCFCB92691" ID-DOI="http://doi.org/10.5281/zenodo.6086444" ID-GBIF-Taxon="120817126" ID-Zenodo-Dep="6086444" LSID="urn:lsid:plazi:treatment:038747324C48C6161EB728DCFCB92691" httpUri="http://treatment.plazi.org/id/038747324C48C6161EB728DCFCB92691" lastPageId="65" lastPageNumber="66" pageId="62" pageNumber="63">
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<treatment id="038747324C48C6161EB728DCFCB92691" ID-DOI="http://doi.org/10.5281/zenodo.6086444" ID-GBIF-Taxon="120817126" ID-Zenodo-Dep="6086444" LSID="urn:lsid:plazi:treatment:038747324C48C6161EB728DCFCB92691" httpUri="http://treatment.plazi.org/id/038747324C48C6161EB728DCFCB92691" lastPageId="65" lastPageNumber="66" pageId="62" pageNumber="63" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
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<subSubSection id="C334A5AF4C48C6691EB728DCFBC62759" pageId="62" pageNumber="63" type="nomenclature">
|
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<paragraph id="8B91F6244C48C6691EB728DCFDE7277F" blockId="62.[151,1045,1521,1581]" box="[151,564,1521,1547]" pageId="62" pageNumber="63">
|
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<heading id="D0D941484C48C6691EB728DCFDE7277F" bold="true" box="[151,564,1521,1547]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
|
||||
<emphasis id="B95A2A364C48C6691EB728DCFDE7277F" bold="true" box="[151,564,1521,1547]" pageId="62" pageNumber="63">
|
||||
<taxonomicName id="4C2E8DA74C48C6691EB728DCFE1A277F" authority="Lees" authorityName="Lees" box="[151,457,1521,1547]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="lanyvary" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C48C6691EB728DCFE1A277F" authority="Lees" authorityName="Lees" box="[151,457,1521,1547]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="lanyvary" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C48C6691EB728DCFE54277F" bold="true" box="[151,391,1521,1547]" italics="true" pageId="62" pageNumber="63">Heteropsis lanyvary</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -58,7 +58,7 @@ Lees
|
|||
: sp. 67 (
|
||||
<bibRefCitation id="EFBF8BD54C48C6691F982B71FDE22700" author="Lees" box="[440,561,1628,1652]" pageId="62" pageNumber="63" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
|
||||
: 66, where considered the same species as
|
||||
<taxonomicName id="4C2E8DA74C48C6691A322B70FB5E2701" box="[1042,1165,1628,1653]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C48C6691A322B70FB5E2701" box="[1042,1165,1628,1653]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C48C6691A322B70FB5E2701" box="[1042,1165,1628,1653]" italics="true" pageId="62" pageNumber="63">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
|
|
@ -79,7 +79,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C48C6691CBF2B89FCDF27C9" box="[671,780,1700,1725]" pageId="62" pageNumber="63" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C48C6691D762B89FC6127C8" box="[854,946,1700,1725]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="62" pageNumber="63">Fig. 19</figureCitation>
|
||||
<figureCitation id="1315EAA14C48C6691D762B89FC6127C8" box="[854,946,1700,1725]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="62" pageNumber="63">Fig. 19</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C48C6691DC92B89FBA327C9" box="[1001,1136,1700,1725]" name="Madagascar" pageId="62" pageNumber="63">Madagascar</collectingCountry>
|
||||
NW, Tsaratanana, piste Mangindrano-Antetykalambazaha,
|
||||
|
|
@ -111,7 +111,7 @@ Deposition
|
|||
</emphasis>
|
||||
: ♀
|
||||
<emphasis id="B95A2A364C48C6691CAE2A3EFD442658" bold="true" box="[654,663,1811,1836]" pageId="62" pageNumber="63">(</emphasis>
|
||||
<figureCitation id="1315EAA14C48C6691CB72A3EFD26265F" box="[663,757,1811,1836]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="62" pageNumber="63">Fig. 19</figureCitation>
|
||||
<figureCitation id="1315EAA14C48C6691CB72A3EFD26265F" box="[663,757,1811,1836]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="62" pageNumber="63">Fig. 19</figureCitation>
|
||||
B),
|
||||
<collectingCountry id="F339B6B44C48C6691D0C2A3EFC602658" box="[812,947,1811,1836]" name="Madagascar" pageId="62" pageNumber="63">Madagascar</collectingCountry>
|
||||
NW, Tsaratanana, piste Mangindrano- Antetykalambazaha, flat grass-bamboo area,
|
||||
|
|
@ -151,7 +151,7 @@ barcode], IA294 [isotope voucher],
|
|||
010289147 [
|
||||
<collectingCountry id="F339B6B44C48C6691A312AE7FB9F2696" box="[1041,1100,1994,2018]" name="Qatar" pageId="62" pageNumber="63">QTR</collectingCountry>
|
||||
barcode]; ♂ (
|
||||
<figureCitation id="1315EAA14C48C6691AC72AE4FAEB2696" box="[1255,1336,1993,2018]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="62" pageNumber="63">Fig. 18</figureCitation>
|
||||
<figureCitation id="1315EAA14C48C6691AC72AE4FAEB2696" box="[1255,1336,1993,2018]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="62" pageNumber="63">Fig. 18</figureCitation>
|
||||
B, right), same data as HT but DL-05-845, IA192 [isotope voucher], [thoracic fragment for KAP],
|
||||
<collectionCode id="ED3F6EE14C49C6681A942DB5FACD21C4" box="[1204,1310,152,176]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="63" pageNumber="64">NHMUK</collectionCode>
|
||||
010289148 [
|
||||
|
|
@ -193,7 +193,7 @@ voucher; cytochrome b],
|
|||
223-15 [
|
||||
<collectionCode id="ED3F6EE14C49C6681AD62C7FFAE5201E" box="[1270,1334,338,362]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="63" pageNumber="64">DNA</collectionCode>
|
||||
barcode voucher]; B43 [‘andromap’]. ♂ (
|
||||
<figureCitation id="1315EAA14C49C6681C2E2C55FDB120E5" box="[526,610,376,401]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="63" pageNumber="64">Fig. 18</figureCitation>
|
||||
<figureCitation id="1315EAA14C49C6681C2E2C55FDB120E5" box="[526,610,376,401]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="63" pageNumber="64">Fig. 18</figureCitation>
|
||||
B, left), NW, Tsaratanana, [Ampidiranala],
|
||||
<geoCoordinate id="EE1A90E34C49C6681A702C55FB1720E5" box="[1104,1220,373,401]" direction="south" orientation="latitude" pageId="63" pageNumber="64" precision="5" value="-14.1958">14.1958o S</geoCoordinate>
|
||||
,
|
||||
|
|
@ -387,11 +387,11 @@ NW, Tsaratanana, near forest margin,
|
|||
<paragraph id="8B91F6244C49C6681EE72838FED824EE" blockId="63.[151,1436,151,1938]" pageId="63" pageNumber="64">
|
||||
<emphasis id="B95A2A364C49C6681EE72838FE92245A" bold="true" box="[199,321,1301,1326]" pageId="63" pageNumber="64">Diagnosis.</emphasis>
|
||||
The species resembles most closely
|
||||
<taxonomicName id="4C2E8DA74C49C6681CDC283BFCAF245A" box="[764,892,1301,1326]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C49C6681CDC283BFCAF245A" box="[764,892,1301,1326]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C49C6681CDC283BFCAF245A" box="[764,892,1301,1326]" italics="true" pageId="63" pageNumber="64">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
from which there are no obvious ♂ genitalic differences.
|
||||
<taxonomicName id="4C2E8DA74C49C6681F062817FE7B2426" box="[294,424,1337,1362]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C49C6681F062817FE7B2426" box="[294,424,1337,1362]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C49C6681F062817FE7B2426" box="[294,424,1337,1362]" italics="true" pageId="63" pageNumber="64">Ht. turbata</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -399,7 +399,7 @@ and
|
|||
<emphasis id="B95A2A364C49C6681FC92817FDBB2426" box="[489,616,1337,1362]" italics="true" pageId="63" pageNumber="64">Ht. pallida</emphasis>
|
||||
</taxonomicName>
|
||||
can be told apart by a more irregular HWV Mb. The most obvious difference from
|
||||
<taxonomicName id="4C2E8DA74C49C6681F752873FE062402" box="[341,469,1373,1398]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C49C6681F752873FE062402" box="[341,469,1373,1398]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C49C6681F752873FE062402" box="[341,469,1373,1398]" italics="true" pageId="63" pageNumber="64">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
is the darker, much more sombre diffuse dark grey colouration, especially on the underside.
|
||||
|
|
@ -409,34 +409,34 @@ is the darker, much more sombre diffuse dark grey colouration, especially on the
|
|||
<paragraph id="8B91F6244C49C6161EE72888FF382069" blockId="63.[151,1436,151,1938]" lastBlockId="65.[151,1437,151,2021]" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64">
|
||||
<emphasis id="B95A2A364C49C6681EE72888FE7D24CA" bold="true" box="[199,430,1445,1470]" pageId="63" pageNumber="64">Description. Wings</emphasis>
|
||||
: upperside uniform dark brown, with FW space-CuA1 ocellus with a wide orange ring which is generally quite concentric but apparently a little wider and more angled proximad. FWD space-M1 ocellus small and slightly elliptic with very narrow orange ring. HWD space-CuA1 ocellus is the main one expressed there, as well as to lesser extents those of space-M3 and space-M2, as in
|
||||
<taxonomicName id="4C2E8DA74C49C6681DAB2B3FFBD4275E" box="[907,1031,1553,1578]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C49C6681DAB2B3FFBD4275E" box="[907,1031,1553,1578]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C49C6681DAB2B3FFBD4275E" box="[907,1031,1553,1578]" italics="true" pageId="63" pageNumber="64">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
. Hairbrush around space-1A region lacking. Underside dark greyish brown, darker than typical of
|
||||
<taxonomicName id="4C2E8DA74C49C6681D6E2B1BFC1A273A" box="[846,969,1589,1614]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C49C6681D6E2B1BFC1A273A" box="[846,969,1589,1614]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C49C6681D6E2B1BFC1A273A" box="[846,969,1589,1614]" italics="true" pageId="63" pageNumber="64">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
, with fine brown and ochreous irroration, especially in basal parts of wings. FWV space-CuA1 ocellus with large orange ring yellowing slightly proximad following the reddish brown concave curve of the Mb before it bends back indistinctly to mid-costa, terminating as if pinched at the base of vein M3 (a feature also displayed by
|
||||
<taxonomicName id="4C2E8DA74C49C6681D662B8FFC1227CE" box="[838,961,1697,1722]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C49C6681D662B8FFC1227CE" box="[838,961,1697,1722]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C49C6681D662B8FFC1227CE" box="[838,961,1697,1722]" italics="true" pageId="63" pageNumber="64">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
). FWV space-M1 ocellus rather small and slightly elliptic with a white pupil with narrow black iris and with a narrow pale orange ring. On HWV, space- CuA1 ocellus quite small and slightly elliptic with a narrow black iris also and pale orange concentric ring. In HT, no other ocelli expressed on HWV. HWV Mb reddish brown, slightly broader than in
|
||||
<taxonomicName id="4C2E8DA74C49C6681A6E2A22FB1A2652" box="[1102,1225,1805,1830]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C49C6681A6E2A22FB1A2652" box="[1102,1225,1805,1830]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C49C6681A6E2A22FB1A2652" box="[1102,1225,1805,1830]" italics="true" pageId="63" pageNumber="64">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
and fairly straight, but finely irregular, with a yellow Mf distad of it in lower basal part of space-M2. A reddish PMb mirrors shape of Mb at least as far as vein 1A. HWV area distad of Mb lighter due to a higher concentration of ochreous scales than the basal area, but as in
|
||||
<taxonomicName id="4C2E8DA74C49C6681F832A57FDCD26E6" box="[419,542,1913,1938]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C49C6681F832A57FDCD26E6" box="[419,542,1913,1938]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C49C6681F832A57FDCD26E6" box="[419,542,1913,1938]" italics="true" pageId="63" pageNumber="64">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
, more fuscous brown scales predominate in the marginal area from veins Rs to M3 particularly, giving the wing margin a slightly ‘singed’ appearance. FWV cell area with four brown rather equally spaced transverse Cbs delineating spaces highlighted by ochreous-yellow hair-scales. Variation. ♂♂ vary quite a bit in size and wing patterning. HWV ocelli in space M1 and M2 often visible as white points. Sexes similar.
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C36C6171EB72A1AFAC126DD" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="64" pageNumber="65" targetBox="[291,1294,196,1817]" targetPageId="64">
|
||||
<caption id="DF51A6AC4C36C6171EB72A1AFAC126DD" ID-DOI="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="64" pageNumber="65" targetBox="[291,1294,196,1817]" targetPageId="64">
|
||||
<paragraph id="8B91F6244C36C6171EB72A1AFAC126DD" blockId="64.[151,1436,1847,1961]" pageId="64" pageNumber="65">
|
||||
<emphasis id="B95A2A364C36C6171EB72A1AFEF82638" bold="true" box="[151,299,1847,1869]" pageId="64" pageNumber="65">FIGURE 19.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
<emphasis id="B95A2A364C36C6171CE92A1AFD082638" bold="true" box="[713,731,1847,1868]" pageId="64" pageNumber="65">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C36C6171D542A15FB432639" authority="Lees" authorityName="Lees" box="[884,1168,1847,1869]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="lanyvary" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C36C6171D542A15FB432639" authority="Lees" authorityName="Lees" box="[884,1168,1847,1869]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="lanyvary" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C36C6171D542A15FC322639" box="[884,993,1848,1869]" italics="true" pageId="64" pageNumber="65">Heteropsis</emphasis>
|
||||
<emphasis id="B95A2A364C36C6171DD32A1AFB982639" box="[1011,1099,1847,1869]" italics="true" pageId="64" pageNumber="65">lanyvary</emphasis>
|
||||
Lees
|
||||
|
|
@ -448,7 +448,7 @@ Lees
|
|||
(Tsaratanana; NHMUK010289144).
|
||||
<emphasis id="B95A2A364C36C6171F5D2A7BFE5D261F" bold="true" box="[381,398,1878,1899]" pageId="64" pageNumber="65">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C36C6171FD42A7AFDA7261F" box="[500,628,1877,1900]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<taxonomicName id="4C2E8DA74C36C6171FD42A7AFDA7261F" box="[500,628,1877,1900]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<emphasis id="B95A2A364C36C6171FD42A7AFDA7261F" box="[500,628,1877,1900]" italics="true" pageId="64" pageNumber="65">Ht. lanyvary</emphasis>
|
||||
</taxonomicName>
|
||||
(Tsaratanana; NHMUK010289145).
|
||||
|
|
@ -473,7 +473,7 @@ Oberthür, 1916
|
|||
Oberthür, 1916
|
||||
</taxonomicName>
|
||||
(‘Madagascar’; BMNH(E) 673678;
|
||||
<taxonomicName id="4C2E8DA74C36C6171D1F2AB9FB8326DD" authority="Butler, 1880" authorityName="Butler" authorityYear="1880" box="[831,1104,1939,1961]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C36C6171D1F2AB9FB8326DD" authority="Butler, 1880" authorityName="Butler" authorityYear="1880" box="[831,1104,1939,1961]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C36C6171D1F2AB9FCB326DD" box="[831,864,1940,1961]" italics="true" pageId="64" pageNumber="65">Ht.</emphasis>
|
||||
<emphasis id="B95A2A364C36C6171D452ABEFC6326DD" box="[869,944,1939,1961]" italics="true" pageId="64" pageNumber="65">turbata</emphasis>
|
||||
(Butler, 1880))
|
||||
|
|
@ -508,9 +508,9 @@ Oberthür, 1916
|
|||
<paragraph id="8B91F6244C37C6161EE72F09FDAF22D1" blockId="65.[151,1437,151,2021]" pageId="65" pageNumber="66">
|
||||
<emphasis id="B95A2A364C37C6161EE72F09FE9F2349" bold="true" box="[199,332,548,573]" pageId="65" pageNumber="66">♂ genitalia</emphasis>
|
||||
: DL-05-761, PT (
|
||||
<figureCitation id="1315EAA14C37C6161C3D2F09FDA22348" box="[541,625,548,573]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="65" pageNumber="66">Fig. 18</figureCitation>
|
||||
<figureCitation id="1315EAA14C37C6161C3D2F09FDA22348" box="[541,625,548,573]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="65" pageNumber="66">Fig. 18</figureCitation>
|
||||
B, left); DL05-845, PT (
|
||||
<figureCitation id="1315EAA14C37C6161DAA2F09FC0D2348" box="[906,990,548,573]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="65" pageNumber="66">Fig. 18</figureCitation>
|
||||
<figureCitation id="1315EAA14C37C6161DAA2F09FC0D2348" box="[906,990,548,573]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="65" pageNumber="66">Fig. 18</figureCitation>
|
||||
B, right). Tegumen with fairly rounded proximad profile (LV) and rather flat dorsal profile (LV) and very slight brow leading to uncus which is about 1/3 longer than the dorsal edge of the tegumen whose DV features a large proximad rather ‘v’-shaped notch, and an uncus whose depth (LV) increases slightly towards midpoint and features a distinct ‘head’ and a small toothed down-hook at tip, while it is slightly inflated medially (DV). The gnathos emerges from a right-angled base and is widest at its attachment and smoothly downrecurved until not quite in line with tegumen-uncus (LV) and is sinuate and slightly outsplayed (DV), with pointed tips slightly incurved. The valve has a small angled dorsal ‘shoulder’ (LV) and is roughly broad-triangular featuring a stubby and uprecurved arm leading to a small club covered with spinoid setae and with inpointing spine from DV. Saccus bulbous, widening proximad (LV), rather long; aedeagus quite deep and relatively straight with about 3–5 small lateral teeth post-medially (also found in
|
||||
<taxonomicName id="4C2E8DA74C37C6161AF22E44FA9E22F4" box="[1234,1357,871,896]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C37C6161AF22E44FA9E22F4" box="[1234,1357,871,896]" italics="true" pageId="65" pageNumber="66">Ht. ankova</emphasis>
|
||||
|
|
@ -528,7 +528,7 @@ Refers to the practical consideration in finding this species that a trip to Tsa
|
|||
<paragraph id="8B91F6244C37C6161EE72931FDFF2444" blockId="65.[151,1437,151,2021]" pageId="65" pageNumber="66">
|
||||
<emphasis id="B95A2A364C37C6161EE72931FE9F2541" bold="true" box="[199,332,1052,1077]" pageId="65" pageNumber="66">Discussion.</emphasis>
|
||||
Sparse historical material of
|
||||
<taxonomicName id="4C2E8DA74C37C6161C812930FCFC2541" box="[673,815,1052,1077]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<taxonomicName id="4C2E8DA74C37C6161C812930FCFC2541" box="[673,815,1052,1077]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<emphasis id="B95A2A364C37C6161C812930FCFC2541" box="[673,815,1052,1077]" italics="true" pageId="65" pageNumber="66">Ht. lanyvary</emphasis>
|
||||
</taxonomicName>
|
||||
collected by R. Paulian at the
|
||||
|
|
@ -542,11 +542,11 @@ series, but the species was discovered again recently in the field (in
|
|||
locality. All available
|
||||
<typeStatus id="549548864C37C6161F2429A5FEEC25D4" box="[260,319,1160,1184]" pageId="65" pageNumber="66">types</typeStatus>
|
||||
in the
|
||||
<taxonomicName id="4C2E8DA74C37C6161FAD29A4FDDD25D4" box="[397,526,1159,1184]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C37C6161FAD29A4FDDD25D4" box="[397,526,1159,1184]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C37C6161FAD29A4FDDD25D4" box="[397,526,1159,1184]" italics="true" pageId="65" pageNumber="66">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group were examined (see also under
|
||||
<taxonomicName id="4C2E8DA74C37C6161DE229A4FB8D25D4" box="[962,1118,1161,1184]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C37C6161DE229A4FB8D25D4" box="[962,1118,1161,1184]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C37C6161DE229A4FB8D25D4" box="[962,1118,1161,1184]" italics="true" pageId="65" pageNumber="66">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
, and elsewhere below). The most relevant is the HT ♂ of
|
||||
|
|
@ -555,15 +555,15 @@ group were examined (see also under
|
|||
Ward, 1870
|
||||
</taxonomicName>
|
||||
(BMNH(E) 673767) (
|
||||
<figureCitation id="1315EAA14C37C6161A102981FB5125B0" box="[1072,1154,1196,1221]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="65" pageNumber="66">Fig. 19</figureCitation>
|
||||
<figureCitation id="1315EAA14C37C6161A102981FB5125B0" box="[1072,1154,1196,1221]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="65" pageNumber="66">Fig. 19</figureCitation>
|
||||
C), lacking locality, from the Chris Ward collection, which differs in ventral wing colouration from all known samples of
|
||||
<taxonomicName id="4C2E8DA74C37C6161AF929FCFAB7259C" box="[1241,1380,1231,1256]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<taxonomicName id="4C2E8DA74C37C6161AF929FCFAB7259C" box="[1241,1380,1231,1256]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<emphasis id="B95A2A364C37C6161AF929FCFAB7259C" box="[1241,1380,1231,1256]" italics="true" pageId="65" pageNumber="66">Ht. lanyvary</emphasis>
|
||||
</taxonomicName>
|
||||
. See also
|
||||
<figureCitation id="1315EAA14C37C6161EED29D9FECC2478" box="[205,287,1268,1293]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="65" pageNumber="66">Fig. 19</figureCitation>
|
||||
<figureCitation id="1315EAA14C37C6161EED29D9FECC2478" box="[205,287,1268,1293]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="65" pageNumber="66">Fig. 19</figureCitation>
|
||||
and under
|
||||
<taxonomicName id="4C2E8DA74C37C6161F8429D8FDFA2479" box="[420,553,1268,1293]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C37C6161F8429D8FDFA2479" box="[420,553,1268,1293]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C37C6161F8429D8FDFA2479" box="[420,553,1268,1293]" italics="true" pageId="65" pageNumber="66">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
regarding
|
||||
|
|
@ -577,13 +577,13 @@ and
|
|||
<emphasis id="B95A2A364C37C6161A9129D9FA8F2479" box="[1201,1372,1268,1293]" italics="true" pageId="65" pageNumber="66">Culapa pallida</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C37C6161B4E29D9FF602444" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="65" pageNumber="66">Fig. 19</figureCitation>
|
||||
<figureCitation id="1315EAA14C37C6161B4E29D9FF602444" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="65" pageNumber="66">Fig. 19</figureCitation>
|
||||
D) and
|
||||
<taxonomicName id="4C2E8DA74C37C6161F27283AFE782444" box="[263,427,1303,1328]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="ornata">
|
||||
<emphasis id="B95A2A364C37C6161F27283AFE782444" box="[263,427,1303,1328]" italics="true" pageId="65" pageNumber="66">Culapa ornata</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C37C6161F9A283AFDD82444" box="[442,523,1303,1328]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="65" pageNumber="66">Fig. 19</figureCitation>
|
||||
<figureCitation id="1315EAA14C37C6161F9A283AFDD82444" box="[442,523,1303,1328]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="65" pageNumber="66">Fig. 19</figureCitation>
|
||||
E).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C37C6161EE72811FDC424E9" blockId="65.[151,1437,151,2021]" pageId="65" pageNumber="66">
|
||||
|
|
@ -605,7 +605,7 @@ E).
|
|||
<emphasis id="B95A2A364C37C6161FD628E0FDA22491" box="[502,625,1484,1509]" italics="true" pageId="65" pageNumber="66">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
; nor was it treated in any prior molecular works but
|
||||
<taxonomicName id="4C2E8DA74C37C6161A9E28E0FA992491" box="[1214,1354,1484,1509]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<taxonomicName id="4C2E8DA74C37C6161A9E28E0FA992491" box="[1214,1354,1484,1509]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<emphasis id="B95A2A364C37C6161A9E28E0FA992491" box="[1214,1354,1484,1509]" italics="true" pageId="65" pageNumber="66">Ht. lanyvary</emphasis>
|
||||
</taxonomicName>
|
||||
is very likely to be the (at Tsaratanana, sympatric) sister species of
|
||||
|
|
@ -644,7 +644,7 @@ Aduse-Poku
|
|||
<paragraph id="8B91F6244C37C6161EE72BC1FB6C2671" blockId="65.[151,1437,151,2021]" box="[199,1215,1772,1797]" pageId="65" pageNumber="66">
|
||||
<emphasis id="B95A2A364C37C6161EE72BC1FE8A2671" bold="true" box="[199,345,1772,1797]" pageId="65" pageNumber="66">Distribution</emphasis>
|
||||
: endemic as far as is known, to the Tsaratanana massif (
|
||||
<figureCitation id="1315EAA14C37C6161DEF2BC1FBF22670" box="[975,1057,1772,1797]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="65" pageNumber="66">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C37C6161DEF2BC1FBF22670" box="[975,1057,1772,1797]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="65" pageNumber="66">Fig. 30</figureCitation>
|
||||
C, olive dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C37C6161EE72A22FB66265C" blockId="65.[151,1437,151,2021]" box="[199,1205,1807,1832]" pageId="65" pageNumber="66">
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<treatment id="038747324C48C6691EB729A7FABD24EA" ID-DOI="http://doi.org/10.5281/zenodo.6086442" ID-GBIF-Taxon="120817129" ID-Zenodo-Dep="6086442" LSID="urn:lsid:plazi:treatment:038747324C48C6691EB729A7FABD24EA" httpUri="http://treatment.plazi.org/id/038747324C48C6691EB729A7FABD24EA" lastPageNumber="63" pageId="62" pageNumber="63">
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<treatment id="038747324C48C6691EB729A7FABD24EA" ID-DOI="http://doi.org/10.5281/zenodo.6086442" ID-GBIF-Taxon="120817129" ID-Zenodo-Dep="6086442" LSID="urn:lsid:plazi:treatment:038747324C48C6691EB729A7FABD24EA" httpUri="http://treatment.plazi.org/id/038747324C48C6691EB729A7FABD24EA" lastPageNumber="63" pageId="62" pageNumber="63" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
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|
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<heading id="D0D941484C48C6691EB729A7FE7725D1" bold="true" box="[151,420,1162,1189]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
|
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<emphasis id="B95A2A364C48C6691EB729A7FE7725D1" bold="true" box="[151,420,1162,1189]" pageId="62" pageNumber="63">
|
||||
<taxonomicName id="4C2E8DA74C48C6691EB729A7FECF25D0" box="[151,284,1162,1188]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C48C6691EB729A7FECF25D0" box="[151,284,1162,1188]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C48C6691EB729A7FECF25D0" bold="true" box="[151,284,1162,1188]" italics="true" pageId="62" pageNumber="63">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
sub-group
|
||||
|
|
@ -52,13 +52,13 @@ sub-group
|
|||
Aduse-Poku
|
||||
<emphasis id="B95A2A364C48C6691F0929FFFEBA259E" box="[297,361,1233,1258]" italics="true" pageId="62" pageNumber="63">et al.,</emphasis>
|
||||
(2015,
|
||||
<figureCitation id="1315EAA14C48C6691FE129FCFDD2259E" box="[449,513,1233,1258]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="62" pageNumber="63">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C48C6691FE129FCFDD2259E" box="[449,513,1233,1258]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="62" pageNumber="63">Fig. 1</figureCitation>
|
||||
) recovered
|
||||
<taxonomicName id="4C2E8DA74C48C6691CAA29FFFCD6259E" box="[650,773,1233,1258]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C48C6691CAA29FFFCD6259E" box="[650,773,1233,1258]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C48C6691CAA29FFFCD6259E" box="[650,773,1233,1258]" italics="true" pageId="62" pageNumber="63">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C48C6691D1D29FFFC68259E" box="[829,955,1233,1258]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C48C6691D1D29FFFC68259E" box="[829,955,1233,1258]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C48C6691D1D29FFFC68259E" box="[829,955,1233,1258]" italics="true" pageId="62" pageNumber="63">Ht. turbata</emphasis>
|
||||
</taxonomicName>
|
||||
+
|
||||
|
|
@ -66,25 +66,25 @@ and
|
|||
<emphasis id="B95A2A364C48C6691DFA29FFFB86259E" box="[986,1109,1233,1258]" italics="true" pageId="62" pageNumber="63">Ht. pallida</emphasis>
|
||||
</taxonomicName>
|
||||
within a grade at the base of the
|
||||
<taxonomicName id="4C2E8DA74C48C6691E9F29DBFE93247A" box="[191,320,1269,1294]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C48C6691E9F29DBFE93247A" box="[191,320,1269,1294]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C48C6691E9F29DBFE93247A" box="[191,320,1269,1294]" italics="true" pageId="62" pageNumber="63">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group, so it is not certain that this sub-group is monophyletic, unlike the
|
||||
<taxonomicName id="4C2E8DA74C48C6691A4F29DBFB23247A" box="[1135,1264,1269,1294]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C48C6691A4F29DBFB23247A" box="[1135,1264,1269,1294]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C48C6691A4F29DBFB23247A" box="[1135,1264,1269,1294]" italics="true" pageId="62" pageNumber="63">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
sub-group (see below). In
|
||||
<bibRefCitation id="EFBF8BD54C48C6691F332837FE042446" author="Lees" box="[275,471,1305,1330]" pageId="62" pageNumber="63" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997: 156)</bibRefCitation>
|
||||
, these ‘
|
||||
<taxonomicName id="4C2E8DA74C48C6691C0D2837FD7A2446" box="[557,681,1305,1330]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C48C6691C0D2837FD7A2446" box="[557,681,1305,1330]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C48C6691C0D2837FD7A2446" box="[557,681,1305,1330]" italics="true" pageId="62" pageNumber="63">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
subgroup’ species formed a morphological clade with
|
||||
<taxonomicName id="4C2E8DA74C48C6691B382837FA4F2446" box="[1304,1436,1305,1330]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C48C6691B382837FA4F2446" box="[1304,1436,1305,1330]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C48C6691B382837FA4F2446" box="[1304,1436,1305,1330]" italics="true" pageId="62" pageNumber="63">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
(‘FFTFR’). All these species feature expression of an ocellus in space-M3 and sometimes space-M2 of the HWD. The species occur in marshy areas near rainforest margins (
|
||||
<taxonomicName id="4C2E8DA74C48C6691D12284FFC7C240E" box="[818,943,1377,1402]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C48C6691D12284FFC7C240E" box="[818,943,1377,1402]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C48C6691D12284FFC7C240E" box="[818,943,1377,1402]" italics="true" pageId="62" pageNumber="63">Ht. turbata</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<treatment id="038747324C4FC6691EB72A7CFAA72543" ID-DOI="http://doi.org/10.5281/zenodo.6086440" ID-GBIF-Taxon="120817125" ID-Zenodo-Dep="6086440" LSID="urn:lsid:plazi:treatment:038747324C4FC6691EB72A7CFAA72543" httpUri="http://treatment.plazi.org/id/038747324C4FC6691EB72A7CFAA72543" lastPageId="62" lastPageNumber="63" pageId="57" pageNumber="58">
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<treatment id="038747324C4FC6691EB72A7CFAA72543" ID-DOI="http://doi.org/10.5281/zenodo.6086440" ID-GBIF-Taxon="120817125" ID-Zenodo-Dep="6086440" LSID="urn:lsid:plazi:treatment:038747324C4FC6691EB72A7CFAA72543" httpUri="http://treatment.plazi.org/id/038747324C4FC6691EB72A7CFAA72543" lastPageId="62" lastPageNumber="63" pageId="57" pageNumber="58" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
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<subSubSection id="C334A5AF4C4FC66E1EB72A7CFBF026F9" pageId="57" pageNumber="58" type="nomenclature">
|
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<paragraph id="8B91F6244C4FC66E1EB72A7CFC87261F" blockId="57.[151,1059,1873,1933]" box="[151,852,1873,1899]" pageId="57" pageNumber="58">
|
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<heading id="D0D941484C4FC66E1EB72A7CFC87261F" bold="true" box="[151,852,1873,1899]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
|
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<emphasis id="B95A2A364C4FC66E1EB72A7CFC87261F" bold="true" box="[151,852,1873,1899]" pageId="57" pageNumber="58">
|
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<taxonomicName id="4C2E8DA74C4FC66E1EB72A7CFD3A261F" authority="Lees, Allaoui & Aduse-Poku" authorityName="Lees, Allaoui & Aduse-Poku" box="[151,745,1873,1899]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1EB72A7CFD3A261F" authority="Lees, Allaoui & Aduse-Poku" authorityName="Lees, Allaoui & Aduse-Poku" box="[151,745,1873,1899]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C4FC66E1EB72A7CFEAE261F" bold="true" box="[151,381,1873,1899]" italics="true" pageId="57" pageNumber="58">Heteropsis tornado</emphasis>
|
||||
Lees, Allaoui & Aduse-Poku
|
||||
</taxonomicName>
|
||||
|
|
@ -64,7 +64,7 @@ Torres
|
|||
2001
|
||||
</bibRefCitation>
|
||||
: 462—where considered to belong to an unsampled clade); “Heteopsis_sp.nov_
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1F572AF0FE032682" box="[375,464,2013,2038]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado">tornado</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1F572AF0FE032682" box="[375,464,2013,2038]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado">tornado</taxonomicName>
|
||||
” [
|
||||
<emphasis id="B95A2A364C4FC66E1FCA2AF3FDDA2681" box="[490,521,2014,2037]" italics="true" pageId="57" pageNumber="58">sic</emphasis>
|
||||
,
|
||||
|
|
@ -74,13 +74,13 @@ Torres
|
|||
, 2015.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF51A6AC4C4CC66D1EB72A4CFA9D26C0" httpUri="https://zenodo.org/record/264614/files/figure.png" pageId="58" pageNumber="59" targetBox="[237,1351,204,1846]" targetPageId="58">
|
||||
<caption id="DF51A6AC4C4CC66D1EB72A4CFA9D26C0" ID-DOI="http://doi.org/10.5281/zenodo.264614" httpUri="https://zenodo.org/record/264614/files/figure.png" pageId="58" pageNumber="59" targetBox="[237,1351,204,1846]" targetPageId="58">
|
||||
<paragraph id="8B91F6244C4CC66D1EB72A4CFA9D26C0" blockId="58.[151,1436,1889,1972]" pageId="58" pageNumber="59">
|
||||
<emphasis id="B95A2A364C4CC66D1EB72A4CFEF62602" bold="true" box="[151,293,1889,1911]" pageId="58" pageNumber="59">FIGURE 17.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
<emphasis id="B95A2A364C4CC66D1CB72A4CFD7A2602" bold="true" box="[663,681,1889,1910]" pageId="58" pageNumber="59">A</emphasis>
|
||||
: HT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C4CC66D1D092A4FFAE52603" authority="Lees, Allaoui & Aduse-Poku" authorityName="Lees, Allaoui & Aduse-Poku" box="[809,1334,1889,1911]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tornado" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C4CC66D1D092A4FFAE52603" authority="Lees, Allaoui & Aduse-Poku" authorityName="Lees, Allaoui & Aduse-Poku" box="[809,1334,1889,1911]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tornado" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C4CC66D1D092A4FFC452603" box="[809,918,1890,1911]" italics="true" pageId="58" pageNumber="59">Heteropsis</emphasis>
|
||||
<emphasis id="B95A2A364C4CC66D1D822A4CFC212603" box="[930,1010,1889,1911]" italics="true" pageId="58" pageNumber="59">tornado</emphasis>
|
||||
Lees, Allaoui & Aduse-Poku
|
||||
|
|
@ -92,31 +92,31 @@ Lees, Allaoui & Aduse-Poku
|
|||
(NHMUK010289143; Anjanaharibe Sud).
|
||||
<emphasis id="B95A2A364C4CC66D1C7D2AADFDBD26E1" bold="true" box="[605,622,1920,1941]" pageId="58" pageNumber="59">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C4CC66D1CCF2AACFCB826E1" box="[751,875,1919,1942]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C4CC66D1CCF2AACFCB826E1" box="[751,875,1919,1942]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C4CC66D1CCF2AACFCB826E1" box="[751,875,1919,1942]" italics="true" pageId="58" pageNumber="59">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
(Marojejy; KAP-MAD1403).
|
||||
<emphasis id="B95A2A364C4CC66D1A962A52FB1B26E1" bold="true" box="[1206,1224,1919,1941]" pageId="58" pageNumber="59">C</emphasis>
|
||||
: ♂ of
|
||||
<taxonomicName id="4C2E8DA74C4CC66D1B3F2AACFA4F26E1" box="[1311,1436,1919,1942]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C4CC66D1B3F2AACFA4F26E1" box="[1311,1436,1919,1942]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C4CC66D1B3F2AACFA4F26E1" box="[1311,1436,1919,1942]" italics="true" pageId="58" pageNumber="59">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
(‘Mahassabe’) [from colour transparency].
|
||||
<emphasis id="B95A2A364C4CC66D1C6C2AB3FD8D26C7" bold="true" box="[588,606,1950,1971]" pageId="58" pageNumber="59">D</emphasis>
|
||||
: ♀ of
|
||||
<taxonomicName id="4C2E8DA74C4CC66D1C812AB2FCC426C0" box="[673,791,1950,1972]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C4CC66D1C812AB2FCC426C0" box="[673,791,1950,1972]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C4CC66D1C812AB2FCC426C0" box="[673,791,1950,1972]" italics="true" pageId="58" pageNumber="59">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
(‘Mahassabe’; BMNH(E) #674881). Scale bar: 30 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF51A6AC4C4DC66C1EB72A78FF0A2691" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="59" pageNumber="60" targetBox="[234,1357,184,1854]" targetPageId="59">
|
||||
<caption id="DF51A6AC4C4DC66C1EB72A78FF0A2691" ID-DOI="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="59" pageNumber="60" targetBox="[234,1357,184,1854]" targetPageId="59">
|
||||
<paragraph id="8B91F6244C4DC66C1EB72A78FF0A2691" blockId="59.[151,1437,1877,2021]" pageId="59" pageNumber="60">
|
||||
<emphasis id="B95A2A364C4DC66C1EB72A78FEF2261E" bold="true" box="[151,289,1877,1899]" pageId="59" pageNumber="60">FIGURE 18.</emphasis>
|
||||
♂ genitalia in LV (left), DV (right).
|
||||
<emphasis id="B95A2A364C4DC66C1C822A78FD67261E" bold="true" box="[674,692,1877,1898]" pageId="59" pageNumber="60">A</emphasis>
|
||||
:
|
||||
<taxonomicName id="4C2E8DA74C4DC66C1CE32A7BFB6D261F" authority="Lees, Allaoui & Aduse-Poku" authorityName="Lees, Allaoui & Aduse-Poku" box="[707,1214,1877,1899]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="tornado" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C4DC66C1CE32A7BFB6D261F" authority="Lees, Allaoui & Aduse-Poku" authorityName="Lees, Allaoui & Aduse-Poku" box="[707,1214,1877,1899]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="tornado" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C4DC66C1CE32A7BFCE3261F" box="[707,816,1878,1899]" italics="true" pageId="59" pageNumber="60">Heteropsis</emphasis>
|
||||
<emphasis id="B95A2A364C4DC66C1D182A78FC5B261F" box="[824,904,1877,1899]" italics="true" pageId="59" pageNumber="60">tornado</emphasis>
|
||||
Lees, Allaoui & Aduse-Poku
|
||||
|
|
@ -128,20 +128,20 @@ Lees, Allaoui & Aduse-Poku
|
|||
, ♂ genitalia (‘Mahassabe’; 362DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale.
|
||||
<emphasis id="B95A2A364C4DC66C1A9E2A59FB1C26FD" bold="true" box="[1214,1231,1908,1929]" pageId="59" pageNumber="60">B</emphasis>
|
||||
: ♂ genitalia of
|
||||
<taxonomicName id="4C2E8DA74C4DC66C1B5C2A58FF3D26DC" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<taxonomicName id="4C2E8DA74C4DC66C1B5C2A58FF3D26DC" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="lanyvary">
|
||||
<emphasis id="B95A2A364C4DC66C1B5C2A58FF3D26DC" italics="true" pageId="59" pageNumber="60">Ht. lanyvary</emphasis>
|
||||
</taxonomicName>
|
||||
. (left: DL-05-761; right: DL-05-845).
|
||||
<emphasis id="B95A2A364C4DC66C1C5F2ABFFD4226DC" bold="true" box="[639,657,1938,1960]" pageId="59" pageNumber="60">C</emphasis>
|
||||
: ♂ genitalia of
|
||||
<taxonomicName id="4C2E8DA74C4DC66C1D182ABEFBD826DC" box="[824,1035,1938,1960]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C4DC66C1D182ABEFBD826DC" box="[824,1035,1938,1960]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C4DC66C1D182ABEFBD826DC" box="[824,1035,1938,1960]" italics="true" pageId="59" pageNumber="60">Heteropsis barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C4DC66C1A332ABEFBB026DC" box="[1043,1123,1939,1960]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(Antsamanarana R., Masoala; CK30, 141DL).
|
||||
<emphasis id="B95A2A364C4DC66C1F1C2A9CFE9D26B2" bold="true" box="[316,334,1969,1990]" pageId="59" pageNumber="60">D</emphasis>
|
||||
: ♂ genitalia of
|
||||
<taxonomicName id="4C2E8DA74C4DC66C1FCD2A9FFCFD26B3" authority="Lees" authorityName="Lees" box="[493,814,1969,1991]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="menamenoides" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C4DC66C1FCD2A9FFCFD26B3" authority="Lees" authorityName="Lees" box="[493,814,1969,1991]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="menamenoides" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C4DC66C1FCD2A9FFD8926B3" box="[493,602,1970,1991]" italics="true" pageId="59" pageNumber="60">Heteropsis</emphasis>
|
||||
<emphasis id="B95A2A364C4DC66C1C402A9CFD2526B3" box="[608,758,1969,1991]" italics="true" pageId="59" pageNumber="60">menamenoides</emphasis>
|
||||
Lees
|
||||
|
|
@ -163,7 +163,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C4AC66B1C522DBAFD0C21C4" box="[626,735,151,176]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♀ (
|
||||
<figureCitation id="1315EAA14C4AC66B1D322DBAFCB021C4" box="[786,867,151,176]" captionStart="FIGURE 17" captionStartId="58.[151,250,1889,1911]" captionTargetBox="[237,1351,204,1846]" captionTargetId="figure@58.[221,1366,193,1867]" captionTargetPageId="58" captionText="FIGURE 17. Adults in DV (left), VV (right). A: HT ♀ of Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov. (NHMUK 010289143; Anjanaharibe Sud). B: PT ♀ of Ht. tornado (Marojejy; KAP-MAD 1403). C: ♂ of Ht. tornado (‘ Mahassabe’) [from colour transparency]. D: ♀ of Ht. tornado (‘ Mahassabe’; BMNH (E) # 674881). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264614/files/figure.png" pageId="60" pageNumber="61">Fig. 17</figureCitation>
|
||||
<figureCitation id="1315EAA14C4AC66B1D322DBAFCB021C4" box="[786,867,151,176]" captionStart="FIGURE 17" captionStartId="58.[151,250,1889,1911]" captionTargetBox="[237,1351,204,1846]" captionTargetId="figure@58.[221,1366,193,1867]" captionTargetPageId="58" captionText="FIGURE 17. Adults in DV (left), VV (right). A: HT ♀ of Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov. (NHMUK 010289143; Anjanaharibe Sud). B: PT ♀ of Ht. tornado (Marojejy; KAP-MAD 1403). C: ♂ of Ht. tornado (‘ Mahassabe’) [from colour transparency]. D: ♀ of Ht. tornado (‘ Mahassabe’; BMNH (E) # 674881). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264614" httpUri="https://zenodo.org/record/264614/files/figure.png" pageId="60" pageNumber="61">Fig. 17</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C4AC66B1DAF2DBAFBC521C4" box="[911,1046,151,176]" name="Madagascar" pageId="60" pageNumber="61">Madagascar</collectingCountry>
|
||||
NE, R.S. Anjanaharibe Sud, slope above R. Marolakana, road between Befingotra and Analamazava,
|
||||
|
|
@ -195,7 +195,7 @@ Deposition
|
|||
</emphasis>
|
||||
: ♀
|
||||
<emphasis id="B95A2A364C4AC66B1C622C00FD982032" bold="true" box="[578,587,301,326]" pageId="60" pageNumber="61">(</emphasis>
|
||||
<figureCitation id="1315EAA14C4AC66B1C6C2C00FD4D2032" box="[588,670,301,326]" captionStart="FIGURE 17" captionStartId="58.[151,250,1889,1911]" captionTargetBox="[237,1351,204,1846]" captionTargetId="figure@58.[221,1366,193,1867]" captionTargetPageId="58" captionText="FIGURE 17. Adults in DV (left), VV (right). A: HT ♀ of Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov. (NHMUK 010289143; Anjanaharibe Sud). B: PT ♀ of Ht. tornado (Marojejy; KAP-MAD 1403). C: ♂ of Ht. tornado (‘ Mahassabe’) [from colour transparency]. D: ♀ of Ht. tornado (‘ Mahassabe’; BMNH (E) # 674881). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264614/files/figure.png" pageId="60" pageNumber="61">Fig. 17</figureCitation>
|
||||
<figureCitation id="1315EAA14C4AC66B1C6C2C00FD4D2032" box="[588,670,301,326]" captionStart="FIGURE 17" captionStartId="58.[151,250,1889,1911]" captionTargetBox="[237,1351,204,1846]" captionTargetId="figure@58.[221,1366,193,1867]" captionTargetPageId="58" captionText="FIGURE 17. Adults in DV (left), VV (right). A: HT ♀ of Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov. (NHMUK 010289143; Anjanaharibe Sud). B: PT ♀ of Ht. tornado (Marojejy; KAP-MAD 1403). C: ♂ of Ht. tornado (‘ Mahassabe’) [from colour transparency]. D: ♀ of Ht. tornado (‘ Mahassabe’; BMNH (E) # 674881). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264614" httpUri="https://zenodo.org/record/264614/files/figure.png" pageId="60" pageNumber="61">Fig. 17</figureCitation>
|
||||
B),
|
||||
<collectingCountry id="F339B6B44C4AC66B1CE92C00FC832032" box="[713,848,301,326]" name="Madagascar" pageId="60" pageNumber="61">Madagascar</collectingCountry>
|
||||
NE, P.N. Marojejy,
|
||||
|
|
@ -243,26 +243,26 @@ Anjanaharibe Sud, slope above R. Marolakana,
|
|||
<paragraph id="8B91F6244C4AC66B1EE72F01FA54238D" blockId="60.[151,1437,151,2021]" pageId="60" pageNumber="61">
|
||||
<emphasis id="B95A2A364C4AC66B1EE72F01FE922331" bold="true" box="[199,321,556,581]" pageId="60" pageNumber="61">Diagnosis.</emphasis>
|
||||
The species is very distinctive especially on the underside (see
|
||||
<figureCitation id="1315EAA14C4AC66B1A002F01FBA22331" box="[1056,1137,556,581]" captionStart="FIGURE 17" captionStartId="58.[151,250,1889,1911]" captionTargetBox="[237,1351,204,1846]" captionTargetId="figure@58.[221,1366,193,1867]" captionTargetPageId="58" captionText="FIGURE 17. Adults in DV (left), VV (right). A: HT ♀ of Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov. (NHMUK 010289143; Anjanaharibe Sud). B: PT ♀ of Ht. tornado (Marojejy; KAP-MAD 1403). C: ♂ of Ht. tornado (‘ Mahassabe’) [from colour transparency]. D: ♀ of Ht. tornado (‘ Mahassabe’; BMNH (E) # 674881). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264614/files/figure.png" pageId="60" pageNumber="61">Fig. 17</figureCitation>
|
||||
<figureCitation id="1315EAA14C4AC66B1A002F01FBA22331" box="[1056,1137,556,581]" captionStart="FIGURE 17" captionStartId="58.[151,250,1889,1911]" captionTargetBox="[237,1351,204,1846]" captionTargetId="figure@58.[221,1366,193,1867]" captionTargetPageId="58" captionText="FIGURE 17. Adults in DV (left), VV (right). A: HT ♀ of Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov. (NHMUK 010289143; Anjanaharibe Sud). B: PT ♀ of Ht. tornado (Marojejy; KAP-MAD 1403). C: ♂ of Ht. tornado (‘ Mahassabe’) [from colour transparency]. D: ♀ of Ht. tornado (‘ Mahassabe’; BMNH (E) # 674881). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264614" httpUri="https://zenodo.org/record/264614/files/figure.png" pageId="60" pageNumber="61">Fig. 17</figureCitation>
|
||||
) and not readily confused with any other species. It has a superficial resemblance though on the dorsal surface to
|
||||
<emphasis id="B95A2A364C4AC66B1A5B2F7CFA52231D" box="[1147,1409,592,617]" italics="true" pageId="60" pageNumber="61">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1A5B2F7CFB22231D" box="[1147,1265,592,617]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="iboina">Ht. iboina</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1ADD2F7CFA52231D" box="[1277,1409,592,617]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tornado">Ht. tornado</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1ADD2F7CFA52231D" box="[1277,1409,592,617]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tornado">Ht. tornado</taxonomicName>
|
||||
</emphasis>
|
||||
is significantly larger than the typical small size of the four known members of (see
|
||||
<bibRefCitation id="EFBF8BD54C4AC66B1A612F58FB6C23F9" author="Lees" box="[1089,1215,629,653]" pageId="60" pageNumber="61" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
) of the
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1B392F58FA4F23F9" box="[1305,1436,628,653]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1B392F58FA4F23F9" box="[1305,1436,628,653]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C4AC66B1B392F58FA4F23F9" box="[1305,1436,628,653]" italics="true" pageId="60" pageNumber="61">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group (
|
||||
<emphasis id="B95A2A364C4AC66B1EC82FB4FD5523C5" box="[232,646,664,689]" italics="true" pageId="60" pageNumber="61">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1EC82FB4FEB523C5" box="[232,358,664,689]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ankova">Ht. ankova</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1EC82FB4FEB523C5" box="[232,358,664,689]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ankova">Ht. ankova</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1F512FB4FE2D23C5" box="[369,510,664,689]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="lanyvary">Ht. lanyvary</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1F512FB4FE2D23C5" box="[369,510,664,689]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="lanyvary">Ht. lanyvary</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1C292FB4FD5523C5" box="[521,646,664,689]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="turbata">Ht. turbata</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1C292FB4FD5523C5" box="[521,646,664,689]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="turbata">Ht. turbata</taxonomicName>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1C9C2FB4FCE623C5" box="[700,821,664,689]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="pallida">
|
||||
|
|
@ -271,7 +271,7 @@ and
|
|||
) (three members of which formed a paraphylum in the study of Aduse-Poku
|
||||
<emphasis id="B95A2A364C4AC66B1FBC2F90FE0A23A1" box="[412,473,700,725]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
|
||||
, 2015), which like
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1CEA2F90FC8023A1" box="[714,851,700,725]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1CEA2F90FC8023A1" box="[714,851,700,725]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C4AC66B1CEA2F90FC8023A1" box="[714,851,700,725]" italics="true" pageId="60" pageNumber="61">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
routinely show expression of HWD ocellus M3. Including referred specimens, the ♂ and ♀ are similar, but the three known ♀♀ are larger than the one known ♂.
|
||||
|
|
@ -282,11 +282,11 @@ routinely show expression of HWD ocellus M3. Including referred specimens, the
|
|||
<emphasis id="B95A2A364C4AC66B1EE72E29FE7C2269" bold="true" box="[199,431,772,797]" pageId="60" pageNumber="61">Description. Wings</emphasis>
|
||||
: upperside uniform mid brown, with a reddish-orange flush in the basal areas of both wings. FWD space-CuA1 ocellus with black iris more than spanning veins CuA1-CuA2. Orng very large, subhexagonal, orange, spanning from upper part of space-M3 down to upper half of 1A. Space-M1 ocellus is slightly elliptic (especially on the HT right FW) and quite large in comparison with
|
||||
<collectingCountry id="F339B6B44C4AC66B1A782E5DFB1622FD" box="[1112,1221,880,905]" name="Madagascar" pageId="60" pageNumber="61">Malagasy</collectingCountry>
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1AED2E5CFA9622FC" box="[1229,1349,881,904]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1AED2E5CFA9622FC" box="[1229,1349,881,904]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C4AC66B1AED2E5CFA9622FC" box="[1229,1349,881,904]" italics="true" pageId="60" pageNumber="61">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
, with a quite wide black iris and narrow orange ring, which extends slightly into space-M2. In HWD three subelliptic ocelli are expressed, the smallest in space-M2, but whose ring is almost contiguous with the ring of the largest of these ocelli in space-M3 whose ring nearly spans veins M3-CuA1, the second largest and most elliptic below it in space-CuA1 spanning about ¾ of inter-vein distance CuA1-CuA2. This is an eyespot expression configuration seen in the
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1EC42908FEB72549" box="[228,356,1060,1085]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1EC42908FEB72549" box="[228,356,1060,1085]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C4AC66B1EC42908FEB72549" box="[228,356,1060,1085]" italics="true" pageId="60" pageNumber="61">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
clade, which is reminiscent also of ‘
|
||||
|
|
@ -298,30 +298,30 @@ clade, which is reminiscent also of ‘
|
|||
(
|
||||
<collectingCountry id="F339B6B44C4AC66B1DFD2909FB962549" box="[989,1093,1060,1085]" name="Marshall Islands" pageId="60" pageNumber="61">Marshall</collectingCountry>
|
||||
, 1880) in the Western Ghats. Proximad of the smallest HWD ocellus, whose ring is about half the diameter of the largest, an orange Mf (median fleck) or crescent mark hugs vein M3 as far as its intersection with Mb (faintly visible on HWD). This Mf is a characteristic of quite a few species of the
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1CAD29BCFCC225DD" box="[653,785,1168,1193]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1CAD29BCFCC225DD" box="[653,785,1168,1193]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C4AC66B1CAD29BCFCC225DD" box="[653,785,1168,1193]" italics="true" pageId="60" pageNumber="61">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group, most markedly so in
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1A4529BCFB3925DD" box="[1125,1258,1168,1193]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1A4529BCFB3925DD" box="[1125,1258,1168,1193]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C4AC66B1A4529BCFB3925DD" box="[1125,1258,1168,1193]" italics="true" pageId="60" pageNumber="61">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
. HW distinctly crenate (as strongly so as in
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1FF92998FDBC25B9" box="[473,623,1204,1229]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="bicristata">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1FF92998FDBC25B9" box="[473,623,1204,1229]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="bicristata">
|
||||
<emphasis id="B95A2A364C4AC66B1FF92998FDBC25B9" box="[473,623,1204,1229]" italics="true" pageId="60" pageNumber="61">Ht. bicristata</emphasis>
|
||||
</taxonomicName>
|
||||
), outwardly reflexed to each darker part of the fringe (forming in effect very convex tails) at the end of veins Rs-CuA1. Underside has an ochreous yellow cast, irrorated with rufous brown especially proximad of the Mb. FWV space-CuA1 Orng varying from light orange to darker orange towards the tergum, where occupying the width of space-1A, its Ore (ocellus ring extension) forming a ‘tornado’ pattern comprising the pale light orange-yellow outer arm of, in effect, a spiral., The paler yellowish colour of this outer arm of the spiral contrasts with the eccentric, elliptic to sub-hexagonal shape of the FWV Orng; such a pattern is evident, with the proximad yellow arm not usually as well separated costad, in a few other
|
||||
<collectingCountry id="F339B6B44C4AC66B1A8E28A1FAC824D1" box="[1198,1307,1420,1445]" name="Madagascar" pageId="60" pageNumber="61">Malagasy</collectingCountry>
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1B0428A0FA4F24D0" box="[1316,1436,1421,1444]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1B0428A0FA4F24D0" box="[1316,1436,1421,1444]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C4AC66B1B0428A0FA4F24D0" box="[1316,1436,1421,1444]" italics="true" pageId="60" pageNumber="61">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
which would be placed in different clades, such as
|
||||
<emphasis id="B95A2A364C4AC66B1CC0289CFC2824BD" box="[736,1019,1456,1481]" italics="true" pageId="60" pageNumber="61">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1CC0289CFC8524BD" box="[736,854,1456,1481]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="iboina">Ht. iboina</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1D43289CFC2824BD" box="[867,1019,1456,1481]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="kremenae">Ht. kremenae</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1D43289CFC2824BD" box="[867,1019,1456,1481]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="kremenae">Ht. kremenae</taxonomicName>
|
||||
</emphasis>
|
||||
, and
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1A1C289CFA4B24BD" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[1084,1432,1456,1481]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<taxonomicName id="4C2E8DA74C4AC66B1A1C289CFA4B24BD" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[1084,1432,1456,1481]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="undulans">
|
||||
<emphasis id="B95A2A364C4AC66B1A1C289CFB1E24BD" box="[1084,1229,1456,1481]" italics="true" pageId="60" pageNumber="61">Ht. undulans</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C4AC66B1AFE289DFA4324BD" author="Oberthur" box="[1246,1424,1456,1481]" pageId="60" pageNumber="61" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
|
|
@ -370,11 +370,11 @@ Refers in particular to the tornado-like markings, the sporadic rarity of this s
|
|||
of a relictual member of a greatly diversified lineage (Aduse- Poku
|
||||
<emphasis id="B95A2A364C4BC66A1EF72FF4FEC32384" box="[215,272,727,752]" italics="true" pageId="61" pageNumber="62">et al.</emphasis>
|
||||
, 2015,
|
||||
<figureCitation id="1315EAA14C4BC66A1F442FFAFE772384" box="[356,420,727,752]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="61" pageNumber="62">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C4BC66A1F442FFAFE772384" box="[356,420,727,752]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="61" pageNumber="62">Fig. 1</figureCitation>
|
||||
). In different endemic radiating mycalesine clades in
|
||||
<collectingCountry id="F339B6B44C4BC66A1A222FFAFB582384" box="[1026,1163,727,752]" name="Madagascar" pageId="61" pageNumber="62">Madagascar</collectingCountry>
|
||||
, there is a trend towards reduction of the number of ocelli expressed in the upperside HW, so its configuration might also be correlated with a typically more open habitat, such as the marshy areas typical for
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1DA22E0CFC2C224C" box="[898,1023,799,824]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1DA22E0CFC2C224C" box="[898,1023,799,824]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C4BC66A1DA22E0CFC2C224C" box="[898,1023,799,824]" italics="true" pageId="61" pageNumber="62">Ht. turbata</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
|
|
@ -388,13 +388,13 @@ This species was first discovered by myself in an uncurated drawer of the Oberth
|
|||
), with a unique ♂ and ♀ labelled as from ‘Mahassabe’ (see Referred specimens), collection dating pre-1923 (
|
||||
<bibRefCitation id="EFBF8BD54C4BC66A1C792EA1FCD022D1" author="Lees" box="[601,771,908,933]" pageId="61" pageNumber="62" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 65</bibRefCitation>
|
||||
). Because of the distance of the locality of these specimens from recent material and the uncertainty of the labels (see also above under
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1A082E9CFB4022BC" box="[1064,1171,945,968]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1A082E9CFB4022BC" box="[1064,1171,945,968]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="sogai">
|
||||
<emphasis id="B95A2A364C4BC66A1A082E9CFB4022BC" box="[1064,1171,945,968]" italics="true" pageId="61" pageNumber="62">Ht. sogai</emphasis>
|
||||
</taxonomicName>
|
||||
) they are not actually included in the
|
||||
<typeStatus id="549548864C4BC66A1F662EF8FEA52299" box="[326,374,981,1005]" pageId="61" pageNumber="62">type</typeStatus>
|
||||
series. The exceptional rarity of this distinctive species, despite intensive prior sampling from the 1890s to the present in the Fianarantsoa (?) and Marojejy-Anjanaharibe Sud areas, is noteworthy. Perhaps this species has naturally small populations, or normally occupies a particular niche that has not been properly sampled and difficult of access, such as open cliff slopes, like the spectacular habitat facing ‘Camp II Marojejya’. Mahasoabe (‘Mahassabe’;
|
||||
<figureCitation id="1315EAA14C4BC66A1FE62949FDCA2508" box="[454,537,1124,1149]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="61" pageNumber="62">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C4BC66A1FE62949FDCA2508" box="[454,537,1124,1149]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="61" pageNumber="62">Fig. 30</figureCitation>
|
||||
D, lower brown dot) is a village located at around
|
||||
<quantity id="4CD65BC14C4BC66A1A742949FB792508" box="[1108,1194,1124,1148]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="61" pageNumber="62" unit="m" value="1100.0">1100 m</quantity>
|
||||
, and its surroundings are largely deforested today, but the nearest primary forest is over
|
||||
|
|
@ -402,17 +402,17 @@ D, lower brown dot) is a village located at around
|
|||
to the east, and assumed plantations or secondary forest can be viewed on Google Earth from
|
||||
<quantity id="4CD65BC14C4BC66A1CD92981FCE125B1" box="[761,818,1196,1221]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="61" pageNumber="62" unit="km" value="4.0">4 km</quantity>
|
||||
to the west. The corresponding printed label is perhaps anyway as mentioned before unreliable, considering that that the same label is placed on series of characteristically northern and lowland species like
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1C1229D8FD662479" box="[562,693,1268,1293]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="erebina">
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1C1229D8FD662479" box="[562,693,1268,1293]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="erebina">
|
||||
<emphasis id="B95A2A364C4BC66A1C1229D8FD662479" box="[562,693,1268,1293]" italics="true" pageId="61" pageNumber="62">Ht. erebina</emphasis>
|
||||
</taxonomicName>
|
||||
. The species is distinctive enough to be unmistakable, but all available
|
||||
<typeStatus id="549548864C4BC66A1F262835FE922444" box="[262,321,1304,1328]" pageId="61" pageNumber="62">types</typeStatus>
|
||||
in the
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1FB72834FDE72444" box="[407,564,1303,1328]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1FB72834FDE72444" box="[407,564,1303,1328]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C4BC66A1FB72834FDE72444" box="[407,564,1303,1328]" italics="true" pageId="61" pageNumber="62">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1C512834FD262444" box="[625,757,1303,1328]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1C512834FD262444" box="[625,757,1303,1328]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C4BC66A1C512834FD262444" box="[625,757,1303,1328]" italics="true" pageId="61" pageNumber="62">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
groups were examined. In particular I checked the STs of
|
||||
|
|
@ -423,7 +423,7 @@ groups were examined. In particular I checked the STs of
|
|||
(which was synonymised by
|
||||
<bibRefCitation id="EFBF8BD54C4BC66A1D652811FC262420" author="d'Abrera" box="[837,1013,1340,1365]" pageId="61" pageNumber="62" refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp." type="book" year="1980">d’Abrera, 1980</bibRefCitation>
|
||||
: 182 with
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1A4E2810FA442420" authority="Butler, 1880" authorityName="Butler" authorityYear="1880" box="[1134,1431,1340,1365]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1A4E2810FA442420" authority="Butler, 1880" authorityName="Butler" authorityYear="1880" box="[1134,1431,1340,1365]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C4BC66A1A4E2810FB3E2421" box="[1134,1261,1340,1365]" italics="true" pageId="61" pageNumber="62">Ht. turbata</emphasis>
|
||||
(Butler, 1880)
|
||||
</taxonomicName>
|
||||
|
|
@ -432,9 +432,9 @@ groups were examined. In particular I checked the STs of
|
|||
<emphasis id="B95A2A364C4BC66A1D872872FB82240C" box="[935,1105,1375,1400]" italics="true" pageId="61" pageNumber="62">Culapa ornata</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C4BC66A1A462872FB6F240C" box="[1126,1212,1375,1400]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="61" pageNumber="62">Fig. 19</figureCitation>
|
||||
<figureCitation id="1315EAA14C4BC66A1A462872FB6F240C" box="[1126,1212,1375,1400]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="61" pageNumber="62">Fig. 19</figureCitation>
|
||||
E, representing
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1B59284CFF3924E9" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C4BC66A1B59284CFF3924E9" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C4BC66A1B59284CFF3924E9" italics="true" pageId="61" pageNumber="62">Ht. turbata</emphasis>
|
||||
</taxonomicName>
|
||||
), here designated: BMNH(E) 673678,
|
||||
|
|
@ -447,7 +447,7 @@ of
|
|||
<bibRefCitation id="EFBF8BD54C4BC66A1FFB288AFD5B24B4" author="Oberthur" box="[475,648,1447,1472]" pageId="61" pageNumber="62" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(LT ♂, missing abdomen,
|
||||
<figureCitation id="1315EAA14C4BC66A1DE5288AFBC424B4" box="[965,1047,1447,1472]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="61" pageNumber="62">Fig. 19</figureCitation>
|
||||
<figureCitation id="1315EAA14C4BC66A1DE5288AFBC424B4" box="[965,1047,1447,1472]" captionStart="FIGURE 19" captionStartId="64.[151,250,1847,1869]" captionTargetBox="[291,1294,196,1817]" captionTargetId="figure@64.[283,1304,193,1826]" captionTargetPageId="64" captionText="FIGURE 19. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis lanyvary Lees, sp. nov. (Tsaratanana; NHMUK 010289144). B: PT ♀ of Ht. lanyvary (Tsaratanana; NHMUK 010289145). C: HT ♂ of Mycalesis ankova Ward, 1870 (‘ Madagascar’; BMNH (E) 673767). D: LT ♂ of Culapa pallida Oberthür, 1916 (‘ Antsianaka’, BMNH (E) 673775). E: LT ♂ of Culapa ornata Oberthür, 1916 (‘ Madagascar’; BMNH (E) 673678; Ht. turbata (Butler, 1880 )). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264616" httpUri="https://zenodo.org/record/264616/files/figure.png" pageId="61" pageNumber="62">Fig. 19</figureCitation>
|
||||
D, here designated, bearing label: “
|
||||
<collectingCountry id="F339B6B44C4BC66A1E8128E3FEFE2493" box="[161,301,1486,1511]" name="Madagascar" pageId="61" pageNumber="62">Madagascar</collectingCountry>
|
||||
, Antsianaka, Perrot Freres, 2e Semestre 1890”|BMNH(E) 673775”; PLT ♂, BMNH(E) 673779 and ♀♀ BMNH(E) 673776 (illustrated in
|
||||
|
|
@ -485,27 +485,27 @@ Mabille, 1880
|
|||
<paragraph id="8B91F6244C4BC6691EE72A38FE522183" blockId="61.[151,1437,151,2018]" lastBlockId="62.[151,1437,151,1079]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
|
||||
<emphasis id="B95A2A364C4BC66A1EE72A38FE3A265A" bold="true" box="[199,489,1813,1838]" pageId="61" pageNumber="62">Additional information.</emphasis>
|
||||
♂ genitalia: 362DL (referred specimen,
|
||||
<figureCitation id="1315EAA14C4BC66A1DEE2A38FBF0265A" box="[974,1059,1813,1838]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="61" pageNumber="62">Fig. 18</figureCitation>
|
||||
<figureCitation id="1315EAA14C4BC66A1DEE2A38FBF0265A" box="[974,1059,1813,1838]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui & Aduse-Poku, sp. nov., ♂ genitalia (‘ Mahassabe’; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264615" httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="61" pageNumber="62">Fig. 18</figureCitation>
|
||||
A): about
|
||||
<quantity id="4CD65BC14C4BC66A1ABD2A3BFB302659" box="[1181,1251,1814,1838]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="61" pageNumber="62" unit="mm" value="2.0">2 mm</quantity>
|
||||
long; from LV, tegumen with bent down to fairly straight uncus leading to downward hook at tip without a distinct ‘head’. Gnathos from square base narrow and tapered, slightly uprecurved to tip. Hinge with vinculum wider than half maximum dimension of tegumen. Saccus rather short, stubby and bulbous. Aedeagus distinctly longer than valve, fairly stout, uprecurved distad of ostium. Valves fairly short, arms uprecurved and leading to club, which has spinoid setae at the end and a distinct ‘beak’ pointing dorsad/mesad. The overall impression is a similarity to genitalia of some of the
|
||||
<taxonomicName id="4C2E8DA74C48C6691EE22DB4FE9621C4" box="[194,325,151,176]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C48C6691EE22DB4FE9621C4" box="[194,325,151,176]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C48C6691EE22DB4FE9621C4" box="[194,325,151,176]" italics="true" pageId="62" pageNumber="63">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group (valves not symmetrically ‘skittle’-shaped’ from LV as in
|
||||
<taxonomicName id="4C2E8DA74C48C6691A182DB4FB0621C4" box="[1080,1237,151,176]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C48C6691A182DB4FB0621C4" box="[1080,1237,151,176]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C48C6691A182DB4FB0621C4" box="[1080,1237,151,176]" italics="true" pageId="62" pageNumber="63">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group and rather uncus is more straight from LV;
|
||||
<bibRefCitation id="EFBF8BD54C48C6691C262D90FDAC21A0" author="Lees" box="[518,639,188,212]" pageId="62" pageNumber="63" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
|
||||
: 106,
|
||||
<figureCitation id="1315EAA14C48C6691CE32D91FCD521A1" box="[707,774,188,213]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="62" pageNumber="63">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C48C6691CE32D91FCD521A1" box="[707,774,188,213]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="62" pageNumber="63">Fig. 7</figureCitation>
|
||||
f). The genitalia of
|
||||
<taxonomicName id="4C2E8DA74C48C6691DC12D90FB8C21A1" box="[993,1119,188,213]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C48C6691DC12D90FB8C21A1" box="[993,1119,188,213]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C48C6691DC12D90FB8C21A1" box="[993,1119,188,213]" italics="true" pageId="62" pageNumber="63">Ht. turbata</emphasis>
|
||||
</taxonomicName>
|
||||
are actually quite similar to those of
|
||||
<taxonomicName id="4C2E8DA74C48C6691ED72DCCFEA9218C" box="[247,378,223,248]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C48C6691ED72DCCFEA9218C" box="[247,378,223,248]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C48C6691ED72DCCFEA9218C" box="[247,378,223,248]" italics="true" pageId="62" pageNumber="63">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
|
|
@ -515,20 +515,20 @@ are actually quite similar to those of
|
|||
<paragraph id="8B91F6244C48C6691EE72C29FDD32010" blockId="62.[151,1437,151,1079]" pageId="62" pageNumber="63">
|
||||
<emphasis id="B95A2A364C48C6691EE72C29FE4C2069" bold="true" box="[199,415,260,285]" pageId="62" pageNumber="63">DNA divergences</emphasis>
|
||||
: COI-5P cluster number BOLD:ACW4939 (exemplar BMAD192-15, DL14A-0245), closest to members of
|
||||
<taxonomicName id="4C2E8DA74C48C6691FB42C04FDC52034" box="[404,534,295,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C48C6691FB42C04FDC52034" box="[404,534,295,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C48C6691FB42C04FDC52034" box="[404,534,295,320]" italics="true" pageId="62" pageNumber="63">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group, e.g. 6.7% divergent to
|
||||
<taxonomicName id="4C2E8DA74C48C6691D4D2C04FBD32034" box="[877,1024,295,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C48C6691D4D2C04FBD32034" box="[877,1024,295,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C48C6691D4D2C04FBD32034" box="[877,1024,295,320]" italics="true" pageId="62" pageNumber="63">Ht. barbarae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C48C6691A272C05FB8A204B" box="[1031,1113,296,319]" pageId="62" pageNumber="63" rank="species">sp. nov</taxonomicNameLabel>
|
||||
and 8.3% to
|
||||
<taxonomicName id="4C2E8DA74C48C6691ACF2C04FABF2034" box="[1263,1388,295,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C48C6691ACF2C04FABF2034" box="[1263,1388,295,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C48C6691ACF2C04FABF2034" box="[1263,1388,295,320]" italics="true" pageId="62" pageNumber="63">Ht. turbata</emphasis>
|
||||
</taxonomicName>
|
||||
and a similar distance to
|
||||
<taxonomicName id="4C2E8DA74C48C6691F5E2C60FE292011" box="[382,506,332,357]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C48C6691F5E2C60FE292011" box="[382,506,332,357]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C48C6691F5E2C60FE292011" box="[382,506,332,357]" italics="true" pageId="62" pageNumber="63">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
|
|
@ -538,28 +538,28 @@ and a similar distance to
|
|||
:
|
||||
<bibRefCitation id="EFBF8BD54C48C6691FD22C5DFD6220FC" author="Lees" box="[498,689,367,392]" pageId="62" pageNumber="63" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997: 156)</bibRefCitation>
|
||||
had over 96% jackknife support for a sister relation of
|
||||
<taxonomicName id="4C2E8DA74C48C6691B3A2C5CFA4F20FC" box="[1306,1436,367,392]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C48C6691B3A2C5CFA4F20FC" box="[1306,1436,367,392]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C48C6691B3A2C5CFA4F20FC" box="[1306,1436,367,392]" italics="true" pageId="62" pageNumber="63">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
(“FFTFR”) with the
|
||||
<taxonomicName id="4C2E8DA74C48C6691FAC2CB8FDDE20D9" box="[396,525,404,429]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C48C6691FAC2CB8FDDE20D9" box="[396,525,404,429]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C48C6691FAC2CB8FDDE20D9" box="[396,525,404,429]" italics="true" pageId="62" pageNumber="63">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
‘clade’ of (then three) species, but this resolution appeared only in the total evidence parsimony analysis. The species would therefore be tentatively placed in the
|
||||
<taxonomicName id="4C2E8DA74C48C6691A462C94FB3B20A4" box="[1126,1256,439,464]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C48C6691A462C94FB3B20A4" box="[1126,1256,439,464]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C48C6691A462C94FB3B20A4" box="[1126,1256,439,464]" italics="true" pageId="62" pageNumber="63">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group based on morphological features and support. Molecular sequencing of multiple genes provides evidence
|
||||
<taxonomicName id="4C2E8DA74C48C6691AD82CF0FAAC2081" box="[1272,1407,476,501]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C48C6691AD82CF0FAAC2081" box="[1272,1407,476,501]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C48C6691AD82CF0FAAC2081" box="[1272,1407,476,501]" italics="true" pageId="62" pageNumber="63">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
is sister to all others of the
|
||||
<taxonomicName id="4C2E8DA74C48C6691F8C2F29FDFE236F" box="[428,557,514,539]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C48C6691F8C2F29FDFE236F" box="[428,557,514,539]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C48C6691F8C2F29FDFE236F" box="[428,557,514,539]" italics="true" pageId="62" pageNumber="63">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group, including the ‘
|
||||
<emphasis id="B95A2A364C48C6691D042CD2FCFA2379" box="[804,809,511,525]" italics="true" pageId="62" pageNumber="63">‘</emphasis>
|
||||
<taxonomicName id="4C2E8DA74C48C6691D0A2F29FC76236F" box="[810,933,514,539]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C48C6691D0A2F29FC76236F" box="[810,933,514,539]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C48C6691D0A2F29FC76236F" box="[810,933,514,539]" italics="true" pageId="62" pageNumber="63">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
’ sub-group (Aduse-Poku
|
||||
|
|
@ -588,7 +588,7 @@ group, including the ‘
|
|||
<paragraph id="8B91F6244C48C6691EE72FD3FA572263" blockId="62.[151,1437,151,1079]" box="[199,1412,766,791]" pageId="62" pageNumber="63">
|
||||
<emphasis id="B95A2A364C48C6691EE72FD3FE8A2263" bold="true" box="[199,345,766,791]" pageId="62" pageNumber="63">Distribution</emphasis>
|
||||
: Marojejy, Anjanaharibe Sud; (?) Mahasoabe, SE of Fianarantsoa (
|
||||
<figureCitation id="1315EAA14C48C6691A6B2FD3FB4E2263" box="[1099,1181,766,791]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="62" pageNumber="63">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C48C6691A6B2FD3FB4E2263" box="[1099,1181,766,791]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="62" pageNumber="63">Fig. 30</figureCitation>
|
||||
D, dark brown dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C48C6691EE72E0FFB1D224F" blockId="62.[151,1437,151,1079]" box="[199,1230,802,827]" pageId="62" pageNumber="63">
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<taxonomicName id="4C2E8DA74C4FC66E1EB72FD1FEAF2262" box="[151,380,764,790]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
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<emphasis id="B95A2A364C4FC66E1EB72FD1FEAF2262" bold="true" box="[151,380,764,790]" italics="true" pageId="57" pageNumber="58">Heteropsis strigula</emphasis>
|
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</taxonomicName>
|
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group
|
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|
|
@ -52,11 +52,11 @@ group
|
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The
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1EEB2E6BFE9E222A" box="[203,333,837,862]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
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<taxonomicName id="4C2E8DA74C4FC66E1EEB2E6BFE9E222A" box="[203,333,837,862]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
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<emphasis id="B95A2A364C4FC66E1EEB2E6BFE9E222A" box="[203,333,837,862]" italics="true" pageId="57" pageNumber="58">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
species group (the ‘
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C152E68FD5F222A" box="[565,652,837,862]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C152E68FD5F222A" box="[565,652,837,862]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C4FC66E1C152E68FD5F222A" box="[565,652,837,862]" italics="true" pageId="57" pageNumber="58">strigula</emphasis>
|
||||
</taxonomicName>
|
||||
15 ’ group of
|
||||
|
|
@ -66,7 +66,7 @@ Torres
|
|||
, 2001
|
||||
</bibRefCitation>
|
||||
), now including
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1A842E6BFAFA222A" box="[1188,1321,837,862]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1A842E6BFAFA222A" box="[1188,1321,837,862]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C4FC66E1A842E6BFAFA222A" box="[1188,1321,837,862]" italics="true" pageId="57" pageNumber="58">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
described here (Aduse-Poku
|
||||
|
|
@ -82,7 +82,7 @@ Torres
|
|||
2001
|
||||
</bibRefCitation>
|
||||
) supported by mitochondrial DNA (COII and cytochrome b). Eight species sampled within it that include
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C8D2ED0FC172561" authority="Ward, 1870" authorityName="Ward" authorityYear="1870" box="[685,964,1020,1045]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C8D2ED0FC172561" authority="Ward, 1870" authorityName="Ward" authorityYear="1870" box="[685,964,1020,1045]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C4FC66E1C8D2ED0FCFB2561" box="[685,808,1020,1045]" italics="true" pageId="57" pageNumber="58">Ht. ankova</emphasis>
|
||||
(Ward, 1870)
|
||||
</taxonomicName>
|
||||
|
|
@ -92,15 +92,15 @@ Lees,
|
|||
<quantity id="4CD65BC14C4FC66E1AA12ED0FB6A2561" box="[1153,1209,1021,1045]" metricMagnitude="1" metricUnit="m" metricValue="5.07238" pageId="57" pageNumber="58" unit="in" value="1997.0">1997</quantity>
|
||||
</bibRefCitation>
|
||||
in a separate group, (the
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1EEB290CFE94254D" box="[203,327,1056,1081]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1EEB290CFE94254D" box="[203,327,1056,1081]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C4FC66E1EEB290CFE94254D" box="[203,327,1056,1081]" italics="true" pageId="57" pageNumber="58">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
clade of the
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1FFD290CFD8C254D" box="[477,607,1056,1081]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1FFD290CFD8C254D" box="[477,607,1056,1081]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C4FC66E1FFD290CFD8C254D" box="[477,607,1056,1081]" italics="true" pageId="57" pageNumber="58">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group that sometimes included
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1DEC290CFB82254D" box="[972,1105,1056,1081]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1DEC290CFB82254D" box="[972,1105,1056,1081]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C4FC66E1DEC290CFB82254D" box="[972,1105,1056,1081]" italics="true" pageId="57" pageNumber="58">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
) are united by a single C->T transversion in the highly conserved 28SDDFF region that is not observed elsewhere in the
|
||||
|
|
@ -114,17 +114,17 @@ Monaghan
|
|||
). Now with the ten gene study of Aduse-Poku
|
||||
<emphasis id="B95A2A364C4FC66E1A3D2944FB8F25F5" box="[1053,1116,1128,1153]" italics="true" pageId="57" pageNumber="58">et al.,</emphasis>
|
||||
(2015, see
|
||||
<figureCitation id="1315EAA14C4FC66E1AFC2945FACE25F4" box="[1244,1309,1128,1153]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="57" pageNumber="58">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C4FC66E1AFC2945FACE25F4" box="[1244,1309,1128,1153]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="57" pageNumber="58">Fig. 1</figureCitation>
|
||||
and Suppl. File S4) that included 13 sampled members, the slightly expanded
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1D8529A0FBFA25D1" box="[933,1065,1164,1189]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1D8529A0FBFA25D1" box="[933,1065,1164,1189]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C4FC66E1D8529A0FBFA25D1" box="[933,1065,1164,1189]" italics="true" pageId="57" pageNumber="58">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group is shown to be the wellsupported sister of the
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1FB7299CFDE225BD" box="[407,561,1200,1225]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1FB7299CFDE225BD" box="[407,561,1200,1225]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C4FC66E1FB7299CFDE225BD" box="[407,561,1200,1225]" italics="true" pageId="57" pageNumber="58">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
clade, which might possibly be (with submarginal bootstrap support) sister to a clade consisting of
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1FA129F8FDC02598" box="[385,531,1237,1260]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="narcissus">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1FA129F8FDC02598" box="[385,531,1237,1260]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="narcissus">
|
||||
<emphasis id="B95A2A364C4FC66E1FA129F8FDC02598" box="[385,531,1237,1260]" italics="true" pageId="57" pageNumber="58">Ht. narcissus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
|
|
@ -133,11 +133,11 @@ clade, which might possibly be (with submarginal bootstrap support) sister to a
|
|||
<bibRefCitation id="EFBF8BD54C4FC66E1C4729F9FC8C2599" author="Balletto" box="[615,863,1236,1261]" pageId="57" pageNumber="58" refString="Balletto, E. & Lees, D. C. (2012) Case 3573 - Papilio narcissus Fabricius, 1798 (currently Heteropsis narcissus; Lepidoptera, NYMPHALIDAE): proposed conservation of the specific name. Bulletin of Zoological Nomenclature, 69, 260 - 265." type="journal article" year="2012">Balletto & Lees, 2012</bibRefCitation>
|
||||
</taxonomicName>
|
||||
) +
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1DA329F8FBCF2599" box="[899,1052,1236,1261]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankaratra">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1DA329F8FBCF2599" box="[899,1052,1236,1261]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankaratra">
|
||||
<emphasis id="B95A2A364C4FC66E1DA329F8FBCF2599" box="[899,1052,1236,1261]" italics="true" pageId="57" pageNumber="58">Ht. ankaratra</emphasis>
|
||||
</taxonomicName>
|
||||
. Five sampled members of the
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1B5929F8FF3D2465" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1B5929F8FF3D2465" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C4FC66E1B5929F8FF3D2465" italics="true" pageId="57" pageNumber="58">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group were submarginally supported (pp=0.86) in the three-gene study of
|
||||
|
|
@ -147,19 +147,19 @@ Kodandaramaiah
|
|||
(2010)
|
||||
</bibRefCitation>
|
||||
. As for the
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1F342830FE632441" box="[276,432,1308,1333]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1F342830FE632441" box="[276,432,1308,1333]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C4FC66E1F342830FE632441" box="[276,432,1308,1333]" italics="true" pageId="57" pageNumber="58">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group, ♂♂ and ♀♀ are similar, apart mainly from size and androconial and/or inflated HW vein differences. ♀♀ of some species are as yet unknown. Both sexes are attracted strongly to fruit as adults, do not fly high, and seem to specialise on forest (or sometimes marshland) grasses. Most species (apart from
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1B592848FF2324D5" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1B592848FF2324D5" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C4FC66E1B592848FF2324D5" italics="true" pageId="57" pageNumber="58">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
, and from the members
|
||||
<emphasis id="B95A2A364C4FC66E1C2628A4FD5B24D5" box="[518,648,1416,1441]" italics="true" pageId="57" pageNumber="58">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C2628A4FD5624D5" box="[518,645,1416,1441]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankova">Ht. ankova</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C2628A4FD5624D5" box="[518,645,1416,1441]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankova">Ht. ankova</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1CB028A4FC6724D4" authority="Butler, 1880" authorityName="Butler" authorityYear="1880" box="[656,948,1416,1441]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1CB028A4FC6724D4" authority="Butler, 1880" authorityName="Butler" authorityYear="1880" box="[656,948,1416,1441]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C4FC66E1CB028A4FCDD24D5" box="[656,782,1416,1441]" italics="true" pageId="57" pageNumber="58">Ht. turbata</emphasis>
|
||||
(Butler, 1880)
|
||||
</taxonomicName>
|
||||
|
|
@ -175,15 +175,15 @@ that were recovered as sister to the rest of the group in
|
|||
: 156, although equivocally, see
|
||||
<bibRefCitation id="EFBF8BD54C4FC66E1A5C2880FB2824B1" author="Lees" box="[1148,1275,1453,1477]" pageId="57" pageNumber="58" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
: 168, and also apart from
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1F3328FCFE7C249C" box="[275,431,1489,1512]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1F3328FCFE7C249C" box="[275,431,1489,1512]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="roussettae">
|
||||
<emphasis id="B95A2A364C4FC66E1F3328FCFE7C249C" box="[275,431,1489,1512]" italics="true" pageId="57" pageNumber="58">Ht. roussettae</emphasis>
|
||||
</taxonomicName>
|
||||
described below) show reduction in ocellus expression in the HW, so that only the space- CuA1 ocellus is expressed, as apparently parallelled in the
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1D7728D8FC252779" box="[855,1014,1524,1549]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1D7728D8FC252779" box="[855,1014,1524,1549]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C4FC66E1D7728D8FC252779" box="[855,1014,1524,1549]" italics="true" pageId="57" pageNumber="58">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group, as described above. Var i ou s species show from LV a combination of relatively straight or gently sinuate ‘neck’ to the uncus before the apical hook, a relatively asymmetric valve base and the extensive development of spinoid setae on the valve tips, and a few members (belonging to
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1FFE2B4CFD45270D" box="[478,662,1632,1657]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1FFE2B4CFD45270D" box="[478,662,1632,1657]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="angulifascia">
|
||||
<emphasis id="B95A2A364C4FC66E1FFE2B4CFD45270D" box="[478,662,1632,1657]" italics="true" pageId="57" pageNumber="58">Ht. angulifascia</emphasis>
|
||||
</taxonomicName>
|
||||
clade) show a striking planification of the gnathos into an ear-like structure, and wavy androconial tufts from HW space-CuA2 sometimes correlate with ventral androconial scale patches. The main radiation occurs throughout the eastern rainforest biome, but one member,
|
||||
|
|
@ -196,7 +196,7 @@ clade) show a striking planification of the gnathos into an ear-like structure,
|
|||
, also occurs in Western
|
||||
<collectingCountry id="F339B6B44C4FC66E1FCA2BE1FDA72791" box="[490,628,1740,1765]" name="Madagascar" pageId="57" pageNumber="58">Madagascar</collectingCountry>
|
||||
.
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C5D2BE0FD282791" box="[637,763,1740,1765]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C5D2BE0FD282791" box="[637,763,1740,1765]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="turbata">
|
||||
<emphasis id="B95A2A364C4FC66E1C5D2BE0FD282791" box="[637,763,1740,1765]" italics="true" pageId="57" pageNumber="58">Ht. turbata</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -204,11 +204,11 @@ and
|
|||
<emphasis id="B95A2A364C4FC66E1D112BE0FC792791" box="[817,938,1740,1765]" italics="true" pageId="57" pageNumber="58">Ht. pallida</emphasis>
|
||||
</taxonomicName>
|
||||
are unusual in that they occur outside forest margins in marshy areas, whereas
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C382BDCFD40267D" box="[536,659,1776,1801]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1C382BDCFD40267D" box="[536,659,1776,1801]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C4FC66E1C382BDCFD40267D" box="[536,659,1776,1801]" italics="true" pageId="57" pageNumber="58">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
as well as
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1D2D2BDCFC49267D" box="[781,922,1776,1801]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="lanyvary" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C4FC66E1D2D2BDCFC49267D" box="[781,922,1776,1801]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="lanyvary" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C4FC66E1D2D2BDCFC49267D" box="[781,922,1776,1801]" italics="true" pageId="57" pageNumber="58">Ht. lanyvary</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C4FC66E1D812BDDFC29267D" box="[929,1018,1776,1801]" pageId="57" pageNumber="58" rank="species">
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<treatment id="038747324C50C67F1EB72C0FFB4522CC" ID-DOI="http://doi.org/10.5281/zenodo.6086426" ID-GBIF-Taxon="120817123" ID-Zenodo-Dep="6086426" LSID="urn:lsid:plazi:treatment:038747324C50C67F1EB72C0FFB4522CC" httpUri="http://treatment.plazi.org/id/038747324C50C67F1EB72C0FFB4522CC" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
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<treatment id="038747324C50C67F1EB72C0FFB4522CC" ID-DOI="http://doi.org/10.5281/zenodo.6086426" ID-GBIF-Taxon="120817123" ID-Zenodo-Dep="6086426" LSID="urn:lsid:plazi:treatment:038747324C50C67F1EB72C0FFB4522CC" httpUri="http://treatment.plazi.org/id/038747324C50C67F1EB72C0FFB4522CC" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
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<subSubSection id="C334A5AF4C50C6711EB72C0FFBC4202A" pageId="38" pageNumber="39" type="nomenclature">
|
||||
<paragraph id="8B91F6244C50C6711EB72C0FFDE32049" blockId="38.[151,1047,290,350]" box="[151,560,290,317]" pageId="38" pageNumber="39">
|
||||
<heading id="D0D941484C50C6711EB72C0FFDE32049" bold="true" box="[151,560,290,317]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
|
||||
<emphasis id="B95A2A364C50C6711EB72C0FFDE32049" bold="true" box="[151,560,290,317]" pageId="38" pageNumber="39">
|
||||
<taxonomicName id="4C2E8DA74C50C6711EB72C0FFE152049" authority="Lees" authorityName="Lees" box="[151,454,290,317]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C50C6711EB72C0FFE152049" authority="Lees" authorityName="Lees" box="[151,454,290,317]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C50C6711EB72C0FFE572048" bold="true" box="[151,388,290,316]" italics="true" pageId="38" pageNumber="39">Heteropsis pauliani</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -76,7 +76,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C50C6711C5D2C83FD3920B3" box="[637,746,430,455]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C50C6711D022C83FCA520B3" box="[802,886,430,455]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="38" pageNumber="39">Fig. 10</figureCitation>
|
||||
<figureCitation id="1315EAA14C50C6711D022C83FCA520B3" box="[802,886,430,455]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="38" pageNumber="39">Fig. 10</figureCitation>
|
||||
C,
|
||||
<collectionCode id="ED3F6EE14C50C6711DB82C82FC2120B3" box="[920,1010,431,455]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="38" pageNumber="39">MNHN</collectionCode>
|
||||
),
|
||||
|
|
@ -152,7 +152,7 @@ extract, cytochrome b];
|
|||
<paragraph id="8B91F6244C50C6711EE72E2DFF3722A2" blockId="38.[151,1437,394,2031]" pageId="38" pageNumber="39">
|
||||
<emphasis id="B95A2A364C50C6711EE72E2DFE79226D" bold="true" box="[199,426,768,793]" pageId="38" pageNumber="39">Deposition MNHN</emphasis>
|
||||
: ♀ (
|
||||
<figureCitation id="1315EAA14C50C6711FC52E2DFDE9226D" box="[485,570,768,793]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="38" pageNumber="39">Fig. 10</figureCitation>
|
||||
<figureCitation id="1315EAA14C50C6711FC52E2DFDE9226D" box="[485,570,768,793]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="38" pageNumber="39">Fig. 10</figureCitation>
|
||||
D), [
|
||||
<collectingCountry id="F339B6B44C50C6711C4F2E2DFD2A226D" box="[623,761,768,793]" name="Madagascar" pageId="38" pageNumber="39">Madagascar</collectingCountry>
|
||||
NW], Mt. Tsaratanana,
|
||||
|
|
@ -224,7 +224,7 @@ Nord, massif du Tsaratanana, en dessous de l’Andohanisambirano [position estim
|
|||
<paragraph id="8B91F6244C50C6711EE72993FB20246B" blockId="38.[151,1437,394,2031]" pageId="38" pageNumber="39">
|
||||
<emphasis id="B95A2A364C50C6711EE72993FE9225A3" bold="true" box="[199,321,1214,1239]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
|
||||
No really similar species even in the
|
||||
<taxonomicName id="4C2E8DA74C50C6711D3929EDFC7625A3" box="[793,933,1214,1239]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C50C6711D3929EDFC7625A3" box="[793,933,1214,1239]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C50C6711D3929EDFC7625A3" box="[793,933,1214,1239]" italics="true" pageId="38" pageNumber="39">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group. This species is very distinctive in possessing tails on the HW margin, giving it a superficial resemblance at least dorsally to a species of the
|
||||
|
|
@ -232,7 +232,7 @@ group. This species is very distinctive in possessing tails on the HW margin, gi
|
|||
<emphasis id="B95A2A364C50C6711B5929C9FF28246B" italics="true" pageId="38" pageNumber="39">Ht. antahala</emphasis>
|
||||
</taxonomicName>
|
||||
group. The long tails distinguish it also from the perhaps quite closely related
|
||||
<taxonomicName id="4C2E8DA74C50C6711A4B2825FB3F246B" box="[1131,1260,1288,1311]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C50C6711A4B2825FB3F246B" box="[1131,1260,1288,1311]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C50C6711A4B2825FB3F246B" box="[1131,1260,1288,1311]" italics="true" pageId="38" pageNumber="39">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
|
|
@ -264,7 +264,7 @@ wingspan.
|
|||
<paragraph id="8B91F6244C51C6701EE72CD2FA4F2384" blockId="39.[151,1437,151,1977]" pageId="39" pageNumber="40">
|
||||
<emphasis id="B95A2A364C51C6701EE72CD2FE9A236C" bold="true" box="[199,329,511,536]" pageId="39" pageNumber="40">♂ genitalia</emphasis>
|
||||
: 97DL (PT,
|
||||
<figureCitation id="1315EAA14C51C6701FF42CD2FDF0236C" box="[468,547,511,536]" captionStart="FIGURE 11" captionStartId="37.[151,250,1177,1199]" captionTargetBox="[264,1363,233,1142]" captionTargetId="figure@37.[212,1379,190,1160]" captionTargetPageId="37" captionText="FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL)." httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="39" pageNumber="40">Fig. 11</figureCitation>
|
||||
<figureCitation id="1315EAA14C51C6701FF42CD2FDF0236C" box="[468,547,511,536]" captionStart="FIGURE 11" captionStartId="37.[151,250,1177,1199]" captionTargetBox="[264,1363,233,1142]" captionTargetId="figure@37.[212,1379,190,1160]" captionTargetPageId="37" captionText="FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL)." figureDoi="http://doi.org/10.5281/zenodo.264608" httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="39" pageNumber="40">Fig. 11</figureCitation>
|
||||
B): from LV, tegumen with distinct brow towards scythe-hooked uncus which is relatively broad at base and narrow in middle. Uncus slightly longer than dorsal profile of tegumen. Constriction at hinge with vinculus is relatively broad, about 2/3 length of tegumen. Gnathos fairly straight and tapered and slightly downcurved pointing towards dorsal edge of uncus. Valves quite long with a slight ventral elbow at base of arm and flattened blades strongly covered in spinoid setae on mesad face of distal club, those at the edge appearing like saw-tooth serrae. Valves protrude almost the full length of club beyond uncus tip. Saccus not especially long and up-bent. Aedeagus nearly full length of valve, quite long proximad of ostium and slightly recurved distad of it.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -274,7 +274,7 @@ B): from LV, tegumen with distinct brow towards scythe-hooked uncus which is rel
|
|||
After the pioneering French entomologist the late Rector Renaud Paulian (
|
||||
<date id="FF90D0E44C51C6701AF02FD1FA802260" box="[1232,1363,764,788]" pageId="39" pageNumber="40" value="1913" valueMax="2003">1913–2003</date>
|
||||
), who described two
|
||||
<taxonomicName id="4C2E8DA74C51C6701F1D2E0CFE66224C" box="[317,437,801,824]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C51C6701F1D2E0CFE66224C" box="[317,437,801,824]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C51C6701F1D2E0CFE66224C" box="[317,437,801,824]" italics="true" pageId="39" pageNumber="40">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
taxa and discovered the first known specimens of this species on Madagascar’s highest mountain a lifetime ago.
|
||||
|
|
@ -294,11 +294,11 @@ in
|
|||
. All available
|
||||
<typeStatus id="549548864C51C6701F5A2EF8FE662299" box="[378,437,981,1005]" pageId="39" pageNumber="40">types</typeStatus>
|
||||
within the
|
||||
<taxonomicName id="4C2E8DA74C51C6701C192EF8FD122299" box="[569,705,980,1005]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C51C6701C192EF8FD122299" box="[569,705,980,1005]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C51C6701C192EF8FD122299" box="[569,705,980,1005]" italics="true" pageId="39" pageNumber="40">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group have been examined and the species is distinct from any
|
||||
<taxonomicName id="4C2E8DA74C51C6701EB72ED4FEDC2564" box="[151,271,1017,1040]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C51C6701EB72ED4FEDC2564" box="[151,271,1017,1040]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C51C6701EB72ED4FEDC2564" box="[151,271,1017,1040]" italics="true" pageId="39" pageNumber="40">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
hitherto described.
|
||||
|
|
@ -306,7 +306,7 @@ hitherto described.
|
|||
<paragraph id="8B91F6244C51C6701EE72931FC742420" blockId="39.[151,1437,151,1977]" pageId="39" pageNumber="40">
|
||||
<emphasis id="B95A2A364C51C6701EE72931FE342541" bold="true" box="[199,487,1052,1077]" pageId="39" pageNumber="40">Additional information.</emphasis>
|
||||
DNA divergences: COI-5P cluster number BOLD:ACW4997 (exemplar MTSBR, BMAD234-15), 6.08% divergent to
|
||||
<taxonomicName id="4C2E8DA74C51C6701C10296CFD4C252C" box="[560,671,1089,1112]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C51C6701C10296CFD4C252C" box="[560,671,1089,1112]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C51C6701C10296CFD4C252C" box="[560,671,1089,1112]" italics="true" pageId="39" pageNumber="40">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
(Anjanaharibe Sud; exemplar BMAD137-15, DL14A-0196, cluster number BOLD:AAK5839) and with a similar divergence to a few other related species. In their COII dataset (
|
||||
|
|
@ -316,7 +316,7 @@ Torres
|
|||
2001
|
||||
</bibRefCitation>
|
||||
), with 422 bp comparable,
|
||||
<taxonomicName id="4C2E8DA74C51C6701C8429A4FCF825D4" box="[676,811,1159,1184]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<taxonomicName id="4C2E8DA74C51C6701C8429A4FCF825D4" box="[676,811,1159,1184]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<emphasis id="B95A2A364C51C6701C8429A4FCF825D4" box="[676,811,1159,1184]" italics="true" pageId="39" pageNumber="40">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
(accession
|
||||
|
|
@ -327,7 +327,7 @@ AY
|
|||
), based on a single individual, is closest (5% pairwise divergent) to “
|
||||
<emphasis id="B95A2A364C51C6701C092980FD9725B0" box="[553,580,1197,1220]" italics="true" pageId="39" pageNumber="40">H.</emphasis>
|
||||
sp. 32” (?also =
|
||||
<taxonomicName id="4C2E8DA74C51C6701CDB2980FCB225B0" box="[763,865,1197,1220]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C51C6701CDB2980FCB225B0" box="[763,865,1197,1220]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C51C6701CDB2980FCB225B0" box="[763,865,1197,1220]" italics="true" pageId="39" pageNumber="40">H. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
|
|
@ -336,7 +336,7 @@ AY
|
|||
<date id="FF90D0E44C51C6701DBA2981FC2325B0" box="[922,1008,1196,1220]" pageId="39" pageNumber="40" value="43-04-01">040143</date>
|
||||
</accessionNumber>
|
||||
), based on one ♂ from Andohahela. It is considerably more pairwise divergent, about 7.67%, in the same COII dataset (417 bp compared) to
|
||||
<taxonomicName id="4C2E8DA74C51C6701B3B29FCFA4F259C" box="[1307,1436,1233,1256]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C51C6701B3B29FCFA4F259C" box="[1307,1436,1233,1256]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C51C6701B3B29FCFA4F259C" box="[1307,1436,1233,1256]" italics="true" pageId="39" pageNumber="40">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
(their ‘
|
||||
|
|
@ -347,9 +347,9 @@ AY
|
|||
<date id="FF90D0E44C51C6701F8429D9FE2A2479" box="[420,505,1268,1293]" pageId="39" pageNumber="40" value="50-04-01">040150</date>
|
||||
</accessionNumber>
|
||||
, based on two ♂♂ from Ankazomivady, an exemplar of which is illustrated in
|
||||
<figureCitation id="1315EAA14C51C6701B5029D9FF752444" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="39" pageNumber="40">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C51C6701B5029D9FF752444" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="39" pageNumber="40">Fig. 7</figureCitation>
|
||||
E). This last taxon is also only marginally different from, and undoubtedly conspecific with (see
|
||||
<taxonomicName id="4C2E8DA74C51C6701B372834FA4F2444" box="[1303,1436,1305,1328]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C51C6701B372834FA4F2444" box="[1303,1436,1305,1328]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C51C6701B372834FA4F2444" box="[1303,1436,1305,1328]" italics="true" pageId="39" pageNumber="40">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
description below) ‘
|
||||
|
|
@ -374,17 +374,17 @@ Torres
|
|||
: sister group uncertain, without agreement so far between molecules and morphology (but see Aduse-Poku
|
||||
<emphasis id="B95A2A364C51C6701C3828A8FD8324E9" box="[536,592,1412,1437]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
|
||||
, 2015; 2016, in press, where sister to
|
||||
<taxonomicName id="4C2E8DA74C51C6701DDB28A8FBAF24E8" box="[1019,1148,1413,1436]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C51C6701DDB28A8FBAF24E8" box="[1019,1148,1413,1436]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C51C6701DDB28A8FBAF24E8" box="[1019,1148,1413,1436]" italics="true" pageId="39" pageNumber="40">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
in combined tree). Based on parsimony analysis of morphological characters,
|
||||
<bibRefCitation id="EFBF8BD54C51C6701CC22885FC2124B4" author="Lees" box="[738,1010,1447,1472]" pageId="39" pageNumber="40" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
|
||||
Lees (1997: 156,
|
||||
<figureCitation id="1315EAA14C51C6701D87288AFC3424B4" box="[935,999,1447,1472]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="39" pageNumber="40">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C51C6701D87288AFC3424B4" box="[935,999,1447,1472]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="39" pageNumber="40">Fig. 1</figureCitation>
|
||||
)
|
||||
</bibRefCitation>
|
||||
had over 96% jackknife support for a sister relationship between
|
||||
<taxonomicName id="4C2E8DA74C51C6701FEC28E0FD872491" box="[460,596,1484,1509]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C51C6701FEC28E0FD872491" box="[460,596,1484,1509]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C51C6701FEC28E0FD872491" box="[460,596,1484,1509]" italics="true" pageId="39" pageNumber="40">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C51C6701C7D28E1FD642491" box="[605,695,1484,1509]" pageId="39" pageNumber="40" rank="species">
|
||||
|
|
@ -393,18 +393,18 @@ had over 96% jackknife support for a sister relationship between
|
|||
(“THRSV”) and
|
||||
<emphasis id="B95A2A364C51C6701D5F28E0FC702490" box="[895,931,1485,1508]" italics="true" pageId="39" pageNumber="40">Ht.</emphasis>
|
||||
76 (“SVTSX”) +
|
||||
<taxonomicName id="4C2E8DA74C51C6701A5328E0FB262490" box="[1139,1269,1485,1508]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C51C6701A5328E0FB262490" box="[1139,1269,1485,1508]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C51C6701A5328E0FB262490" box="[1139,1269,1485,1508]" italics="true" pageId="39" pageNumber="40">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
(“TWNTW”), while
|
||||
<taxonomicName id="4C2E8DA74C51C6701EFE28DCFE52277C" box="[222,385,1519,1544]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C51C6701EFE28DCFE52277C" box="[222,385,1519,1544]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C51C6701EFE28DCFE52277C" box="[222,385,1519,1544]" italics="true" pageId="39" pageNumber="40">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C51C6701FAB28C2FE35277C" box="[395,486,1519,1544]" pageId="39" pageNumber="40" rank="species">
|
||||
<emphasis id="B95A2A364C51C6701FAB28C2FE35277C" bold="true" box="[395,486,1519,1544]" pageId="39" pageNumber="40">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
(THRTB) and
|
||||
<taxonomicName id="4C2E8DA74C51C6701CB828DCFCDA277C" box="[664,777,1521,1544]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C51C6701CB828DCFCDA277C" box="[664,777,1521,1544]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C51C6701CB828DCFCDA277C" box="[664,777,1521,1544]" italics="true" pageId="39" pageNumber="40">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
(THRTN) fell in the next branch down. In their cladistic parsimony analysis of COII sequences,
|
||||
|
|
@ -414,7 +414,7 @@ Torres
|
|||
(2001
|
||||
</bibRefCitation>
|
||||
: 467) found a topological relationship, although with meagre 51% bootstrap support, for a clade consisting of
|
||||
<taxonomicName id="4C2E8DA74C51C6701D3F2B14FC7B2724" box="[799,936,1591,1616]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<taxonomicName id="4C2E8DA74C51C6701D3F2B14FC7B2724" box="[799,936,1591,1616]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<emphasis id="B95A2A364C51C6701D3F2B14FC7B2724" box="[799,936,1591,1616]" italics="true" pageId="39" pageNumber="40">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
(their
|
||||
|
|
@ -424,13 +424,13 @@ Torres
|
|||
sp. 74’ + ‘
|
||||
<emphasis id="B95A2A364C51C6701F472B70FEA82700" box="[359,379,1629,1652]" italics="true" pageId="39" pageNumber="40">H</emphasis>
|
||||
. sp. 32’ (taxa from Andohahela and Anjanaharibe Sud, respectively that were fully supported as sisters; but they did not include
|
||||
<taxonomicName id="4C2E8DA74C51C6701C092BACFD4A27EC" box="[553,665,1665,1688]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C51C6701C092BACFD4A27EC" box="[553,665,1665,1688]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C51C6701C092BACFD4A27EC" box="[553,665,1665,1688]" italics="true" pageId="39" pageNumber="40">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<emphasis id="B95A2A364C51C6701C8C2BACFD1327EC" box="[684,704,1665,1688]" italics="true" pageId="39" pageNumber="40">H</emphasis>
|
||||
. sp. 13) nor
|
||||
<taxonomicName id="4C2E8DA74C51C6701D762BACFC2B27EC" box="[854,1016,1663,1688]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<taxonomicName id="4C2E8DA74C51C6701D762BACFC2B27EC" box="[854,1016,1663,1688]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<emphasis id="B95A2A364C51C6701D762BACFC2B27EC" box="[854,1016,1663,1688]" italics="true" pageId="39" pageNumber="40">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
|
|
@ -455,18 +455,18 @@ AY
|
|||
<date id="FF90D0E44C51C6701B692BC1FA4F2671" box="[1353,1436,1772,1797]" pageId="39" pageNumber="40" value="69-04-01">040169</date>
|
||||
</accessionNumber>
|
||||
(
|
||||
<taxonomicName id="4C2E8DA74C51C6701E802A3CFD8D265C" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[160,606,1807,1832]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="laetifica">
|
||||
<taxonomicName id="4C2E8DA74C51C6701E802A3CFD8D265C" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[160,606,1807,1832]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="laetifica">
|
||||
<emphasis id="B95A2A364C51C6701E802A3CFE56265C" box="[160,389,1807,1832]" italics="true" pageId="39" pageNumber="40">Heteropsis laetifica</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C51C6701FBD2A22FD84265C" author="Oberthur" box="[413,599,1807,1832]" pageId="39" pageNumber="40" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, Analalava, a species in the
|
||||
<taxonomicName id="4C2E8DA74C51C6701DF52A3CFB8C265C" box="[981,1119,1807,1832]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C51C6701DF52A3CFB8C265C" box="[981,1119,1807,1832]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C51C6701DF52A3CFB8C265C" box="[981,1119,1807,1832]" italics="true" pageId="39" pageNumber="40">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group), whereas the real morphospecies 74 should be either closely related, if not conspecific, with
|
||||
<taxonomicName id="4C2E8DA74C51C6701DC82A18FB322638" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[1000,1249,1844,1868]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C51C6701DC82A18FB322638" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[1000,1249,1844,1868]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C51C6701DC82A18FB8A2638" box="[1000,1113,1845,1868]" italics="true" pageId="39" pageNumber="40">Ht. viettei</emphasis>
|
||||
Lees, 2003
|
||||
</taxonomicName>
|
||||
|
|
@ -496,7 +496,7 @@ Lees, 2003
|
|||
<paragraph id="8B91F6244C5EC67F1EE72C0AFBA020FC" blockId="40.[151,1437,151,952]" pageId="40" pageNumber="41">
|
||||
<emphasis id="B95A2A364C5EC67F1EE72C0AFE8A2034" bold="true" box="[199,345,295,320]" pageId="40" pageNumber="41">Distribution</emphasis>
|
||||
: endemic to the mountain ranges between Tsaratanana and Anjanaharibe Sud, as far as is known (
|
||||
<figureCitation id="1315EAA14C5EC67F1EBF2C61FF222010" box="[159,241,332,357]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="40" pageNumber="41">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C5EC67F1EBF2C61FF222010" box="[159,241,332,357]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="40" pageNumber="41">Fig. 30</figureCitation>
|
||||
A, dark green dots). Due to a formatting error, the locality was incorrectly given in Table 1 of
|
||||
<bibRefCitation id="EFBF8BD54C5EC67F1B2F2C60FF1C20FC" author="Torres" pageId="40" pageNumber="41" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. & Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
|
||||
Torres
|
||||
|
|
@ -516,13 +516,13 @@ as Andohahela, instead of its true provenance in the line below, Anjanaharibe Su
|
|||
<paragraph id="8B91F6244C5EC67F1EE72C9AFBE12349" blockId="40.[151,1437,151,952]" pageId="40" pageNumber="41">
|
||||
<emphasis id="B95A2A364C5EC67F1EE72C9AFEB420A4" bold="true" box="[199,359,439,464]" pageId="40" pageNumber="41">Conservation</emphasis>
|
||||
: probably the highest elevation
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1CFE2C94FC8520A4" box="[734,854,441,464]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1CFE2C94FC8520A4" box="[734,854,441,464]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C5EC67F1CFE2C94FC8520A4" box="[734,854,441,464]" italics="true" pageId="40" pageNumber="41">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
in
|
||||
<collectingCountry id="F339B6B44C5EC67F1DA02C9AFBD920A4" box="[896,1034,439,464]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
|
||||
. Not likely in imminent danger by habitat destruction.
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1F542CF0FE292081" box="[372,506,476,501]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1F542CF0FE292081" box="[372,506,476,501]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<emphasis id="B95A2A364C5EC67F1F542CF0FE292081" box="[372,506,476,501]" italics="true" pageId="40" pageNumber="41">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
likely occurs on a number of high and rather inaccessible peaks or ridges across its range. It might however be endangered by global warming. No individuals were found in a lower elevational transect up to
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<mods:classification id="9CA2AB236E95AD62A14E513318DDA246">journal article</mods:classification>
|
||||
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||||
<mods:identifier id="44795459B52DA66B2FD93BB176FF86C8" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
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||||
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<treatment id="038747324C54C6711EB72966FF2221A3" ID-DOI="http://doi.org/10.5281/zenodo.6086422" ID-GBIF-Taxon="120817122" ID-Zenodo-Dep="6086422" LSID="urn:lsid:plazi:treatment:038747324C54C6711EB72966FF2221A3" httpUri="http://treatment.plazi.org/id/038747324C54C6711EB72966FF2221A3" lastPageId="38" lastPageNumber="38" pageId="34" pageNumber="35">
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||||
<treatment id="038747324C54C6711EB72966FF2221A3" ID-DOI="http://doi.org/10.5281/zenodo.6086422" ID-GBIF-Taxon="120817122" ID-Zenodo-Dep="6086422" LSID="urn:lsid:plazi:treatment:038747324C54C6711EB72966FF2221A3" httpUri="http://treatment.plazi.org/id/038747324C54C6711EB72966FF2221A3" lastPageId="38" lastPageNumber="38" pageId="34" pageNumber="35" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C54C6751EB72966FBC425F2" pageId="34" pageNumber="35" type="nomenclature">
|
||||
<paragraph id="8B91F6244C54C6751EB72966FD942511" blockId="34.[151,1047,1099,1158]" box="[151,583,1099,1125]" pageId="34" pageNumber="35">
|
||||
<heading id="D0D941484C54C6751EB72966FD942511" bold="true" box="[151,583,1099,1125]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
|
||||
<emphasis id="B95A2A364C54C6751EB72966FD942511" bold="true" box="[151,583,1099,1125]" pageId="34" pageNumber="35">
|
||||
<taxonomicName id="4C2E8DA74C54C6751EB72966FE0E2511" authority="Lees" authorityName="Lees" box="[151,477,1099,1125]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C54C6751EB72966FE0E2511" authority="Lees" authorityName="Lees" box="[151,477,1099,1125]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C54C6751EB72966FE482511" bold="true" box="[151,411,1099,1125]" italics="true" pageId="34" pageNumber="35">Heteropsis westwoodi</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -64,7 +64,7 @@ Lees
|
|||
] (Aduse-Poku
|
||||
<emphasis id="B95A2A364C54C6751AC4299AFAF125BA" box="[1252,1314,1205,1230]" italics="true" pageId="34" pageNumber="35">et al.,</emphasis>
|
||||
2015:
|
||||
<figureCitation id="1315EAA14C54C6751B502998FF772586" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="34" pageNumber="35">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C54C6751B502998FF772586" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="34" pageNumber="35">Fig. 1</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -78,7 +78,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C54C6751C5229D0FD0C2462" box="[626,735,1277,1302]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C54C6751D3229D0FCB02462" box="[786,867,1277,1302]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="34" pageNumber="35">Fig. 10</figureCitation>
|
||||
<figureCitation id="1315EAA14C54C6751D3229D0FCB02462" box="[786,867,1277,1302]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="34" pageNumber="35">Fig. 10</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C54C6751DAE29D0FBC62462" box="[910,1045,1277,1302]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
|
||||
SE, Vohitrasiva, near ridge, "
|
||||
|
|
@ -171,17 +171,17 @@ E, Vohitrasiva, Andringitra,
|
|||
<paragraph id="8B91F6244C54C6751EE72B82FE2D2640" blockId="34.[151,1437,1205,2022]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B95A2A364C54C6751EE72B82FE9227BC" bold="true" box="[199,321,1711,1736]" pageId="34" pageNumber="35">Diagnosis.</emphasis>
|
||||
There is only one particularly similar species to
|
||||
<taxonomicName id="4C2E8DA74C54C6751D452B9CFBD527BC" box="[869,1030,1711,1736]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<taxonomicName id="4C2E8DA74C54C6751D452B9CFBD527BC" box="[869,1030,1711,1736]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<emphasis id="B95A2A364C54C6751D452B9CFBD527BC" box="[869,1030,1711,1736]" italics="true" pageId="34" pageNumber="35">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<taxonomicName id="4C2E8DA74C54C6751A352B9CFB5727BC" box="[1045,1156,1713,1736]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C54C6751A352B9CFB5727BC" box="[1045,1156,1713,1736]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C54C6751A352B9CFB5727BC" box="[1045,1156,1713,1736]" italics="true" pageId="34" pageNumber="35">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
has a similar white band distad of the HWV Mb, but is generally smaller and much more widely distributed (sympatric in Fianarantsoa province) in
|
||||
<collectingCountry id="F339B6B44C54C6751F0C2BDAFE652664" box="[300,438,1783,1808]" name="Madagascar" pageId="34" pageNumber="35">Madagascar</collectingCountry>
|
||||
. Among related species,
|
||||
<taxonomicName id="4C2E8DA74C54C6751CFA2BD5FC882664" box="[730,859,1783,1808]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C54C6751CFA2BD5FC882664" box="[730,859,1783,1808]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C54C6751CFA2BD5FC882664" box="[730,859,1783,1808]" italics="true" pageId="34" pageNumber="35">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
♂♂ have yellow rather than whitish highlighting distad of the Mb in some forms.
|
||||
|
|
@ -193,9 +193,9 @@ has a similar white band distad of the HWV Mb, but is generally smaller and much
|
|||
: dorsal surface fairly uniform dark brown, darker in area of cell and above towards costa. FWD space-CuA1 ocellus expressed, roughly circular, its black iris spanning inter-vein distance CuA1-CuA2, with a narrow concentric dark orange ring. HWD space-cuA1 ocellus elliptic, with a black iris spanning more than half intervein distance CuA1-CuA2, and a dark orange ring relatively slightly broader than in the FWD and nearly spanning the intervein distance. No other ocelli expressed on FWD. A dark Sml quite closely hugs the margin in both wings and both surfaces. HW margin moderately and evenly crenulate, with rather blunt dark blackish brown tails on the otherwise violet-grey-brown fringe. Ventral surface with a light grey-violet cast, especially on the HW. FWV ocellus space-CuA1 is circular and same size as on the FWD with a concentric quite narrow orange ring. M1 ocellus FWV fairly small and circular with black iris and faint narrow orange ring. A small ocellus with very small black iris is expressed just below the M1 ocellus FWV. In the HWV the space-CuA1 ocellus is as on the HWD and space-R5-M3 ocelli are expressed as very small white points. Mb irregular, strongly concave from the margin to M2 where it forms a cusp, then strongly and irregularly indented and maximally concave in space-CuA1, convex again in space-CuA2 and concave again to vein 1A+2A where it terminates. The Mb HWV delineates a relatively darker brown irrorated area proximad to the PMb that only slightly repeats the shape of the Mb. Distad of the Mb from veins M2-1A is a clear white band, also irregular, with prominence in space M2 and maximum width proximad of space-CuA1 ocellus, tapering to 1A+2A with a slight expansion in Space-CuA2. Distad of this white band is a slightly paler brown area irrorated less densely with darker brown than the band in the more basal area. Patterning on the FWV is fairly uniform, darker proximad of the Mb that is concave around the space-CuA1 ocellus and recurves back towards mid-costa, with indications of three or four paler stripes above the cell. There is slightly paler brown highlighting in premarginal area just proximad of the dark Sml.
|
||||
<emphasis id="B95A2A364C55C6741A442FA1FB0823D1" bold="true" box="[1124,1243,652,677]" pageId="35" pageNumber="36">Variation.</emphasis>
|
||||
A PT ♂ (DL95- 0001) has a much stronger violet-grey cast, especially to the HWV. Sexually dimorphic (
|
||||
<figureCitation id="1315EAA14C55C6741A542F82FB1623BC" box="[1140,1221,687,712]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="35" pageNumber="36">Fig. 10</figureCitation>
|
||||
<figureCitation id="1315EAA14C55C6741A542F82FB1623BC" box="[1140,1221,687,712]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="35" pageNumber="36">Fig. 10</figureCitation>
|
||||
A–B), ♀ larger and lighter in referred specimen BMNH(E) #674842 (
|
||||
<figureCitation id="1315EAA14C55C6741CE32FFEFCC52398" box="[707,790,723,748]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="35" pageNumber="36">Fig. 10</figureCitation>
|
||||
<figureCitation id="1315EAA14C55C6741CE32FFEFCC52398" box="[707,790,723,748]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="35" pageNumber="36">Fig. 10</figureCitation>
|
||||
B), but similar in ventral patterning, although with a more contrasting darker band between the HWV PMb and Mb.
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C55C6741EE72E36FC0F2223" blockId="35.[151,1437,151,1999]" pageId="35" pageNumber="36">
|
||||
|
|
@ -223,12 +223,12 @@ B), but similar in ventral patterning, although with a more contrasting darker b
|
|||
: 236DL, PT ♂ from Vohitraseva (
|
||||
<bibRefCitation id="EFBF8BD54C55C6741CEC2EDFFB02257E" author="Lees" box="[716,1233,1009,1034]" pageId="35" pageNumber="36" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="i">
|
||||
Lees, 1997: 109,
|
||||
<figureCitation id="1315EAA14C55C6741DB02EDCFC00257E" box="[912,979,1009,1034]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="35" pageNumber="36">
|
||||
<figureCitation id="1315EAA14C55C6741DB02EDCFC00257E" box="[912,979,1009,1034]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="35" pageNumber="36">
|
||||
Fig.
|
||||
<date id="FF90D0E44C55C6741DE42EDCFC00257E" box="[964,979,1009,1034]" pageId="35" pageNumber="36">7</date>
|
||||
</figureCitation>
|
||||
i, “13B”; see
|
||||
<figureCitation id="1315EAA14C55C6741A4A2EDCFB6A257E" box="[1130,1209,1009,1034]" captionStart="FIGURE 11" captionStartId="37.[151,250,1177,1199]" captionTargetBox="[264,1363,233,1142]" captionTargetId="figure@37.[212,1379,190,1160]" captionTargetPageId="37" captionText="FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL)." httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="35" pageNumber="36">Fig. 11</figureCitation>
|
||||
<figureCitation id="1315EAA14C55C6741A4A2EDCFB6A257E" box="[1130,1209,1009,1034]" captionStart="FIGURE 11" captionStartId="37.[151,250,1177,1199]" captionTargetBox="[264,1363,233,1142]" captionTargetId="figure@37.[212,1379,190,1160]" captionTargetPageId="37" captionText="FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL)." figureDoi="http://doi.org/10.5281/zenodo.264608" httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="35" pageNumber="36">Fig. 11</figureCitation>
|
||||
A
|
||||
</bibRefCitation>
|
||||
). From LV, a very shallowly domed tegumen with a rather straight dorsal edge profile and slight brow leads to a ‘hand-scythe’-shaped uncus, which is fairly evenly deep except towards the tip of the hook, where it features more of a ‘head’ dorsad; very thin from DV. Tegumen with wide ‘U’-shaped proximad notch, from LV proximad profile bending round sharply to a constriction at junction with vinculum. This ‘waist’ is over half the maximum dimension of the tegumen (and thus the tegumen does not have a quadrate shape from LV). Valve with a very prominent convex dorsal ‘shoulder’ making up part of an oblong shape which leads to the narrow quite tapering and fairly straight valve arms, which are ventrally slightly elbowed and with a very slightly expanded club extending beyond the uncus tip, with a dorsal tooth and with a slightly incurved, rather blunt and serrate distal region (pointed and somewhat incurved from DV); this club covered in an array of spinoid setae on its mesad face. Gnathos from rightangled base pointing in line with the uncus tip in its porrect position, relatively straight and tapered to pointed tip and with negligible serration; from DV slightly sinuate and inrecurved at tip. Saccus moderately extended and markedly inflated proximad, less than a third of total valve length. Aedeagus very thin, dorsoventrally flattened and uprecurved in distad half, with a long ostium proximad, proximad tip rather semicircular from DV. Juxta quite prominent proximad and keel-like from LV, bilobed from DV.
|
||||
|
|
@ -238,11 +238,11 @@ A
|
|||
<paragraph id="8B91F6244C55C6741EE72B24FD45271E" blockId="35.[151,1437,151,1999]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B95A2A364C55C6741EE72B24FE9D2756" bold="true" box="[199,334,1545,1570]" pageId="35" pageNumber="36">Etymology.</emphasis>
|
||||
After John Obadiah Westwood, founder of the genus
|
||||
<taxonomicName id="4C2E8DA74C55C6741D942B27FBFF2755" box="[948,1068,1546,1569]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C55C6741D942B27FBFF2755" box="[948,1068,1546,1569]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C55C6741D942B27FBFF2755" box="[948,1068,1546,1569]" italics="true" pageId="35" pageNumber="36">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
and describer of the remarkable
|
||||
<taxonomicName id="4C2E8DA74C55C6741EB72B03FEA02732" box="[151,371,1581,1606]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C55C6741EB72B03FEA02732" box="[151,371,1581,1606]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C55C6741EB72B03FEA02732" box="[151,371,1581,1606]" italics="true" pageId="35" pageNumber="36">Heteropsis drepana</emphasis>
|
||||
</taxonomicName>
|
||||
. The Hope Department of Entomology at Oxford, that Westwood founded, contains a number of other interesting species from
|
||||
|
|
@ -270,22 +270,22 @@ was examined and belongs to a not closely related species, as discussed above. T
|
|||
<bibRefCitation id="EFBF8BD54C55C6741DB92BF2FBFB278C" author="Butler" box="[921,1064,1759,1784]" pageId="35" pageNumber="36" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||||
</taxonomicName>
|
||||
was also examined (BMNH(E) #674838) together with its genitalic slide dissection (P. Viette no. 4847; see also above) and this species (now known as
|
||||
<taxonomicName id="4C2E8DA74C55C6741F362A05FDA02634" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[278,627,1832,1856]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C55C6741F362A05FDA02634" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[278,627,1832,1856]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C55C6741F362A05FE32264B" box="[278,481,1832,1855]" italics="true" pageId="35" pageNumber="36">Heteropsis viettei</emphasis>
|
||||
Lees, 2003
|
||||
</taxonomicName>
|
||||
) is clearly not conspecific, despite the superficial similarity to
|
||||
<taxonomicName id="4C2E8DA74C55C6741B592A05FEDE2610" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<taxonomicName id="4C2E8DA74C55C6741B592A05FEDE2610" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<emphasis id="B95A2A364C55C6741B592A05FEDE2610" italics="true" pageId="35" pageNumber="36">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
of Pl. 5,
|
||||
<figureCitation id="1315EAA14C55C6741F542A66FE122610" box="[372,449,1867,1892]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="35" pageNumber="36">Figs. 6</figureCitation>
|
||||
<figureCitation id="1315EAA14C55C6741F542A66FE122610" box="[372,449,1867,1892]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="35" pageNumber="36">Figs. 6</figureCitation>
|
||||
,
|
||||
<figureCitation id="1315EAA14C55C6741FE62A66FE062610" box="[454,469,1867,1892]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="35" pageNumber="36">7</figureCitation>
|
||||
<figureCitation id="1315EAA14C55C6741FE62A66FE062610" box="[454,469,1867,1892]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="35" pageNumber="36">7</figureCitation>
|
||||
of
|
||||
<bibRefCitation id="EFBF8BD54C55C6741FDA2A66FD7C2610" box="[506,687,1867,1892]" pageId="35" pageNumber="36" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter">Mabille ([1885]</bibRefCitation>
|
||||
). The specimen represents
|
||||
<taxonomicName id="4C2E8DA74C55C6741DFF2A61FB9E2617" box="[991,1101,1868,1891]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C55C6741DFF2A61FB9E2617" box="[991,1101,1868,1891]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C55C6741DFF2A61FB9E2617" box="[991,1101,1868,1891]" italics="true" pageId="35" pageNumber="36">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
, as discussed above.
|
||||
|
|
@ -297,13 +297,13 @@ of
|
|||
2015) from Ambondrombe).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF51A6AC4C52C6731EB72BE4FE61264E" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="36" pageNumber="37" targetBox="[272,1316,212,1704]" targetPageId="36">
|
||||
<caption id="DF51A6AC4C52C6731EB72BE4FE61264E" ID-DOI="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="36" pageNumber="37" targetBox="[272,1316,212,1704]" targetPageId="36">
|
||||
<paragraph id="8B91F6244C52C6731EB72BE4FE61264E" blockId="36.[151,1436,1737,1850]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B95A2A364C52C6731EB72BE4FEF127AA" bold="true" box="[151,290,1737,1759]" pageId="36" pageNumber="37">FIGURE 10.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
<emphasis id="B95A2A364C52C6731CA42BE4FD4527AA" bold="true" box="[644,662,1737,1758]" pageId="36" pageNumber="37">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C52C6731D2C2BE7FBFA27AB" authority="Lees" authorityName="Lees" box="[780,1065,1737,1759]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C52C6731D2C2BE7FBFA27AB" authority="Lees" authorityName="Lees" box="[780,1065,1737,1759]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C52C6731D2C2BE7FC3D27AB" box="[780,1006,1737,1759]" italics="true" pageId="36" pageNumber="37">Heteropsis westwoodi</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -314,13 +314,13 @@ Lees
|
|||
(Vohitrasiva, Andringitra; NHMUK010289130).
|
||||
<emphasis id="B95A2A364C52C6731F5D2BC5FE5D2789" bold="true" box="[381,398,1768,1789]" pageId="36" pageNumber="37">B</emphasis>
|
||||
: referred ♀ of
|
||||
<taxonomicName id="4C2E8DA74C52C6731C052BC4FD652789" box="[549,694,1767,1790]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<taxonomicName id="4C2E8DA74C52C6731C052BC4FD652789" box="[549,694,1767,1790]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<emphasis id="B95A2A364C52C6731C052BC4FD652789" box="[549,694,1767,1790]" italics="true" pageId="36" pageNumber="37">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
(Andohahela, Antananandava, Andohahela; BMNH(E) #674842).
|
||||
<emphasis id="B95A2A364C52C6731B7B2BCAFABE2789" bold="true" box="[1371,1389,1767,1789]" pageId="36" pageNumber="37">C</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C52C6731EEB2A2AFEAD2668" authority="Lees" authorityName="Lees" box="[203,382,1798,1820]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C52C6731EEB2A2AFEAD2668" authority="Lees" authorityName="Lees" box="[203,382,1798,1820]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C52C6731EEB2A2AFE952668" box="[203,326,1798,1820]" italics="true" pageId="36" pageNumber="37">Ht. pauliani</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -331,7 +331,7 @@ Lees
|
|||
(Tsaratanana; DCL-DB-2966; MNHN).
|
||||
<emphasis id="B95A2A364C52C6731D552A2AFC542668" bold="true" box="[885,903,1799,1820]" pageId="36" pageNumber="37">D</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C52C6731DCD2A2AFBBB2668" box="[1005,1128,1798,1820]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<taxonomicName id="4C2E8DA74C52C6731DCD2A2AFBBB2668" box="[1005,1128,1798,1820]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<emphasis id="B95A2A364C52C6731DCD2A2AFBBB2668" box="[1005,1128,1798,1820]" italics="true" pageId="36" pageNumber="37">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
(Tsaratanana; DCL-DB-2975; MNHN). Scale bar: 30 mm.
|
||||
|
|
@ -341,21 +341,21 @@ Lees
|
|||
<paragraph id="8B91F6244C52C6731EE72A4AFB5E269F" blockId="36.[151,1436,1895,2028]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B95A2A364C52C6731EE72A4AFE3426F4" bold="true" box="[199,487,1895,1920]" pageId="36" pageNumber="37">Phylogeny/sister species</emphasis>
|
||||
: apparently most closely related to
|
||||
<taxonomicName id="4C2E8DA74C52C6731DA72A45FBDB26F4" box="[903,1032,1895,1920]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C52C6731DA72A45FBDB26F4" box="[903,1032,1895,1920]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C52C6731DA72A45FBDB26F4" box="[903,1032,1895,1920]" italics="true" pageId="36" pageNumber="37">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
(cluster number BOLD:ACW5694 and to
|
||||
<taxonomicName id="4C2E8DA74C52C6731EC82AA0FE8A26D0" box="[232,345,1933,1956]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C52C6731EC82AA0FE8A26D0" box="[232,345,1933,1956]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C52C6731EC82AA0FE8A26D0" box="[232,345,1933,1956]" italics="true" pageId="36" pageNumber="37">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
(BOLDAAK5839).
|
||||
<bibRefCitation id="EFBF8BD54C52C6731C662AA1FD1C26D0" author="Lees" box="[582,719,1932,1956]" pageId="36" pageNumber="37" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997)</bibRefCitation>
|
||||
recovered
|
||||
<taxonomicName id="4C2E8DA74C52C6731D712AA0FC2026D0" box="[849,1011,1931,1956]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<taxonomicName id="4C2E8DA74C52C6731D712AA0FC2026D0" box="[849,1011,1931,1956]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<emphasis id="B95A2A364C52C6731D712AA0FC2026D0" box="[849,1011,1931,1956]" italics="true" pageId="36" pageNumber="37">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
(as ‘THRTB’) as sister to
|
||||
<taxonomicName id="4C2E8DA74C52C6731B0B2AA0FA4F26D0" box="[1323,1436,1933,1956]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C52C6731B0B2AA0FA4F26D0" box="[1323,1436,1933,1956]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C52C6731B0B2AA0FA4F26D0" box="[1323,1436,1933,1956]" italics="true" pageId="36" pageNumber="37">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
(‘THRTN’), based on all morphological evidence, genitalic characters, and non-genitalic characters, but without any jackknife support.
|
||||
|
|
@ -365,16 +365,16 @@ Aduse-Poku
|
|||
(2016
|
||||
</bibRefCitation>
|
||||
, in press) confirm it as sister to
|
||||
<taxonomicName id="4C2E8DA74C52C6731A382AF8FB552698" box="[1048,1158,2005,2028]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C52C6731A382AF8FB552698" box="[1048,1158,2005,2028]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C52C6731A382AF8FB552698" box="[1048,1158,2005,2028]" italics="true" pageId="36" pageNumber="37">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C53C6721EB729B4FC6025BA" httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="37" pageNumber="38" targetBox="[264,1363,233,1142]" targetPageId="37">
|
||||
<caption id="DF51A6AC4C53C6721EB729B4FC6025BA" ID-DOI="http://doi.org/10.5281/zenodo.264608" httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="37" pageNumber="38" targetBox="[264,1363,233,1142]" targetPageId="37">
|
||||
<paragraph id="8B91F6244C53C6721EB729B4FC6025BA" blockId="37.[151,1436,1177,1230]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B95A2A364C53C6721EB729B4FEF225DA" bold="true" box="[151,289,1177,1199]" pageId="37" pageNumber="38">FIGURE 11.</emphasis>
|
||||
♂ genitalia in LV (left), DV (right). A: ♂ genitalia of
|
||||
<taxonomicName id="4C2E8DA74C53C6721D5B29B7FB4A25DB" authority="Lees" authorityName="Lees" box="[891,1177,1177,1199]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C53C6721D5B29B7FB4A25DB" authority="Lees" authorityName="Lees" box="[891,1177,1177,1199]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C53C6721D5B29B7FB8E25DB" box="[891,1117,1177,1199]" italics="true" pageId="37" pageNumber="38">Heteropsis westwoodi</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -383,7 +383,7 @@ Lees
|
|||
<emphasis id="B95A2A364C53C6721A8829B7FB2E25DB" bold="true" box="[1192,1277,1178,1199]" pageId="37" pageNumber="38">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
(Antanandava, Andohahela; 236DL). B:
|
||||
<taxonomicName id="4C2E8DA74C53C6721FB92994FDC725BA" box="[409,532,1208,1230]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<taxonomicName id="4C2E8DA74C53C6721FB92994FDC725BA" box="[409,532,1208,1230]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<emphasis id="B95A2A364C53C6721FB92994FDC725BA" box="[409,532,1208,1230]" italics="true" pageId="37" pageNumber="38">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
, ♂ genitalia, PT ♂ (Tsaratanana; 97DL).
|
||||
|
|
@ -415,7 +415,7 @@ Lees
|
|||
: endemic to the southeastern mountains of
|
||||
<collectingCountry id="F339B6B44C53C6721D6F2884FC0524B6" box="[847,982,1449,1474]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
|
||||
from Andohahela north to Andringitra, Mt. Ambondrombe and Mt Maharira, Ranomafana National Park (
|
||||
<figureCitation id="1315EAA14C53C6721DA128E0FC012492" box="[897,978,1485,1510]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="37" pageNumber="38">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C53C6721DA128E0FC012492" box="[897,978,1485,1510]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="37" pageNumber="38">Fig. 30</figureCitation>
|
||||
A, pink dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C53C6721EE728DCFB35277E" blockId="37.[151,1437,1269,2035]" box="[199,1254,1521,1546]" pageId="37" pageNumber="38">
|
||||
|
|
@ -496,7 +496,7 @@ extract number],
|
|||
243-15 [
|
||||
<collectionCode id="ED3F6EE14C53C6721FAC2A31FE1F2640" box="[396,460,1820,1844]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="37" pageNumber="38">DNA</collectionCode>
|
||||
barcode number]; ♀ (
|
||||
<figureCitation id="1315EAA14C53C6721CE32A36FCC62640" box="[707,789,1819,1844]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="37" pageNumber="38">Fig. 10</figureCitation>
|
||||
<figureCitation id="1315EAA14C53C6721CE32A36FCC62640" box="[707,789,1819,1844]" captionStart="FIGURE 10" captionStartId="36.[151,250,1737,1759]" captionTargetBox="[272,1316,212,1704]" captionTargetId="figure@36.[258,1329,193,1715]" captionTargetPageId="36" captionText="FIGURE 10. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis westwoodi Lees, sp. nov. (Vohitrasiva, Andringitra; NHMUK 010289130). B: referred ♀ of Ht. westwoodi (Andohahela, Antananandava, Andohahela; BMNH (E) # 674842). C: HT ♂ of Ht. pauliani Lees, sp. nov. (Tsaratanana; DCL-DB- 2966; MNHN). D: PT ♀ of Ht. pauliani (Tsaratanana; DCL-DB- 2975; MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264607" httpUri="https://zenodo.org/record/264607/files/figure.png" pageId="37" pageNumber="38">Fig. 10</figureCitation>
|
||||
B), SE, PN Andohahela, Antananandava,
|
||||
<quantity id="4CD65BC14C53C6721AC62A31FAEF2647" box="[1254,1340,1820,1844]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="38" unit="m" value="1200.0">1200 m</quantity>
|
||||
,
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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|
||||
<emphasis id="B95A2A364C5AC67B1EB729EDFE3A25AF" bold="true" box="[151,489,1216,1243]" pageId="44" pageNumber="45">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1EB729EDFE4425AE" box="[151,407,1216,1242]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1EB729EDFE4425AE" box="[151,407,1216,1242]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C5AC67B1EB729EDFE4425AE" bold="true" box="[151,407,1216,1242]" italics="true" pageId="44" pageNumber="45">Heteropsis subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group
|
||||
|
|
@ -50,7 +50,7 @@ group
|
|||
</paragraph>
|
||||
<paragraph id="8B91F6244C5AC67A1EB7282AFCB32348" blockId="44.[151,1436,1287,2035]" lastBlockId="45.[151,1437,151,573]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
|
||||
The ‘
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1EF2282AFE912454" box="[210,322,1287,1312]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1EF2282AFE912454" box="[210,322,1287,1312]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C5AC67B1EF2282AFE912454" box="[210,322,1287,1312]" italics="true" pageId="44" pageNumber="45">subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
7’ species group (after
|
||||
|
|
@ -143,7 +143,7 @@ Linares
|
|||
presented a supported phylogeography of relationships of three species. Recently though, Aduse-Poku
|
||||
<emphasis id="B95A2A364C5AC67B1A022B06FBB12737" box="[1058,1122,1578,1603]" italics="true" pageId="44" pageNumber="45">et al.,</emphasis>
|
||||
(2015,
|
||||
<figureCitation id="1315EAA14C5AC67B1A9A2B07FB2A2737" box="[1210,1273,1578,1603]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="44" pageNumber="45">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C5AC67B1A9A2B07FB2A2737" box="[1210,1273,1578,1603]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="44" pageNumber="45">Fig. 1</figureCitation>
|
||||
) found strong support for three sampled species namely
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1C4C2B62FCD62713" box="[620,773,1614,1639]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C5AC67B1C4C2B62FCD62713" box="[620,773,1614,1639]" italics="true" pageId="44" pageNumber="45">Ht. subsimilis</emphasis>
|
||||
|
|
@ -157,40 +157,40 @@ and
|
|||
<emphasis id="B95A2A364C5AC67B1DE12B62FB8E2712" box="[961,1117,1615,1638]" italics="true" pageId="44" pageNumber="45">Ht. comorana</emphasis>
|
||||
</taxonomicName>
|
||||
, which clade they recovered as sister to 13 sampled species of the
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1C692B5EFD1827FF" box="[585,715,1650,1675]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1C692B5EFD1827FF" box="[585,715,1650,1675]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C5AC67B1C692B5EFD1827FF" box="[585,715,1650,1675]" italics="true" pageId="44" pageNumber="45">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group, including
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1DB92B5EFBCE27FF" box="[921,1053,1650,1675]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tornado" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1DB92B5EFBCE27FF" box="[921,1053,1650,1675]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tornado" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C5AC67B1DB92B5EFBCE27FF" box="[921,1053,1650,1675]" italics="true" pageId="44" pageNumber="45">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C5AC67B1A062B5FFB5227FF" box="[1062,1153,1650,1675]" pageId="44" pageNumber="45" rank="species">
|
||||
<emphasis id="B95A2A364C5AC67B1A062B5FFB5227FF" bold="true" box="[1062,1153,1650,1675]" pageId="44" pageNumber="45">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
and the
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1AC62B5EFAB027FF" box="[1254,1379,1650,1675]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1AC62B5EFAB027FF" box="[1254,1379,1650,1675]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ankova">
|
||||
<emphasis id="B95A2A364C5AC67B1AC62B5EFAB027FF" box="[1254,1379,1650,1675]" italics="true" pageId="44" pageNumber="45">Ht. ankova</emphasis>
|
||||
</taxonomicName>
|
||||
subgroup (see below). ♂ genitalia have valve tips with a stout inward projecting spine that fall far short of the uncus extension; from LV, the uncus is inflated before its tip. The ♂ genitalia are in fact remarkably miniaturised compared to other
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1F492BF2FE322782" box="[361,481,1759,1782]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1F492BF2FE322782" box="[361,481,1759,1782]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C5AC67B1F492BF2FE322782" box="[361,481,1759,1782]" italics="true" pageId="44" pageNumber="45">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
, only around
|
||||
<quantity id="4CD65BC14C5AC67B1C5D2BF2FD012782" box="[637,722,1759,1783]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="44" pageNumber="45" unit="mm" value="1.7">1.7 mm</quantity>
|
||||
in length (apart from those of
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1A082BF2FB782783" box="[1064,1195,1758,1783]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1A082BF2FB782783" box="[1064,1195,1758,1783]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tornado">
|
||||
<emphasis id="B95A2A364C5AC67B1A082BF2FB782783" box="[1064,1195,1758,1783]" italics="true" pageId="44" pageNumber="45">Ht. tornado</emphasis>
|
||||
</taxonomicName>
|
||||
which is about
|
||||
<quantity id="4CD65BC14C5AC67B1B7C2BF2FA4F2782" box="[1372,1436,1759,1783]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="45" unit="mm" value="2.0">2 mm</quantity>
|
||||
long, it can be seen in figures here that those of the
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1CFD2A2EFC8D266F" box="[733,862,1794,1819]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1CFD2A2EFC8D266F" box="[733,862,1794,1819]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C5AC67B1CFD2A2EFC8D266F" box="[733,862,1794,1819]" italics="true" pageId="44" pageNumber="45">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group are generally between
|
||||
<quantity id="4CD65BC14C5AC67B1A8C2A2EFAD9266E" box="[1196,1290,1795,1819]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="44" pageNumber="45" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||||
long and the
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1EB72A0AFECE264B" box="[151,285,1830,1855]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1EB72A0AFECE264B" box="[151,285,1830,1855]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C5AC67B1EB72A0AFECE264B" box="[151,285,1830,1855]" italics="true" pageId="44" pageNumber="45">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -202,15 +202,15 @@ groups are more than three mm long and exhibit extension of particular parts). I
|
|||
<emphasis id="B95A2A364C5AC67B1C792A42FD2026F3" box="[601,755,1902,1927]" italics="true" pageId="44" pageNumber="45">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group, yet only in the
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1DD42A42FBA926F3" box="[1012,1146,1902,1927]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1DD42A42FBA926F3" box="[1012,1146,1902,1927]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C5AC67B1DD42A42FBA926F3" box="[1012,1146,1902,1927]" italics="true" pageId="44" pageNumber="45">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group is there a tendency to reduction or even loss (the supra-discocellular patch of
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1D092ABEFC7126DE" box="[809,930,1939,1962]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="narova">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1D092ABEFC7126DE" box="[809,930,1939,1962]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="narova">
|
||||
<emphasis id="B95A2A364C5AC67B1D092ABEFC7126DE" box="[809,930,1939,1962]" italics="true" pageId="44" pageNumber="45">Ht. narova</emphasis>
|
||||
</taxonomicName>
|
||||
) of parts of the androconial system (perhaps regained in
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1F052A9AFE6226BB" box="[293,433,1974,1999]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C5AC67B1F052A9AFE6226BB" box="[293,433,1974,1999]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C5AC67B1F052A9AFE6226BB" box="[293,433,1974,1999]" italics="true" pageId="44" pageNumber="45">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
only). Atrophication of the genitalic system may be compensated for in a different, possibly androconial character system. Phenotypically, adult ♂♂ and ♀♀ of the
|
||||
|
|
@ -218,7 +218,7 @@ only). Atrophication of the genitalic system may be compensated for in a differe
|
|||
<emphasis id="B95A2A364C5AC67B1A0F2AF6FB182687" box="[1071,1227,2010,2035]" italics="true" pageId="44" pageNumber="45">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group are similar, nearly monomorphic except in size, but for simple androconia in the ♂ discocellular region and an inflated HW vein system also found in the
|
||||
<taxonomicName id="4C2E8DA74C5BC67A1FCC2D90FDBD21A1" box="[492,622,188,213]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C5BC67A1FCC2D90FDBD21A1" box="[492,622,188,213]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C5BC67A1FCC2D90FDBD21A1" box="[492,622,188,213]" italics="true" pageId="45" pageNumber="46">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group. In the
|
||||
|
|
@ -226,11 +226,11 @@ group. In the
|
|||
<emphasis id="B95A2A364C5BC67A1D302D90FC7F21A1" box="[784,940,188,213]" italics="true" pageId="45" pageNumber="46">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group, this system features a set of linearly rather than bulbously inflated veins in the ♂ HW (
|
||||
<figureCitation id="1315EAA14C5BC67A1CC32DF2FCEA218C" box="[739,825,223,248]" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="45" pageNumber="46">Fig. 12</figureCitation>
|
||||
<figureCitation id="1315EAA14C5BC67A1CC32DF2FCEA218C" box="[739,825,223,248]" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." figureDoi="http://doi.org/10.5281/zenodo.264609" httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="45" pageNumber="46">Fig. 12</figureCitation>
|
||||
;
|
||||
<bibRefCitation id="EFBF8BD54C5BC67A1D682DCDFC1A218C" author="Lees" box="[840,969,224,248]" pageId="45" pageNumber="46" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
: 96). As predominant in the
|
||||
<taxonomicName id="4C2E8DA74C5BC67A1B382DCCFA4F218C" box="[1304,1436,223,248]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C5BC67A1B382DCCFA4F218C" box="[1304,1436,223,248]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C5BC67A1B382DCCFA4F218C" box="[1304,1436,223,248]" italics="true" pageId="45" pageNumber="46">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group, all species have the reduced ocellus configuration of the HWD, with only that in space CuA1 expressed as well as often that in space Rs; the morphological ‘total evidence’ phylogenetic hypotheses of
|
||||
|
|
@ -240,7 +240,7 @@ that the
|
|||
<emphasis id="B95A2A364C5BC67A1EE12C60FE8F2011" box="[193,348,332,357]" italics="true" pageId="45" pageNumber="46">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group is sister to only part of the
|
||||
<taxonomicName id="4C2E8DA74C5BC67A1CC02C60FCB12011" box="[736,866,332,357]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C5BC67A1CC02C60FCB12011" box="[736,866,332,357]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C5BC67A1CC02C60FCB12011" box="[736,866,332,357]" italics="true" pageId="45" pageNumber="46">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group (see also
|
||||
|
|
@ -256,10 +256,10 @@ Kodandaramaiah
|
|||
(2010
|
||||
</bibRefCitation>
|
||||
,
|
||||
<figureCitation id="1315EAA14C5BC67A1F342CB9FE8420D8" box="[276,343,404,429]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="45" pageNumber="46">Fig. 3</figureCitation>
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<figureCitation id="1315EAA14C5BC67A1F342CB9FE8420D8" box="[276,343,404,429]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="45" pageNumber="46">Fig. 3</figureCitation>
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) would imply that this configuration of HWD reduction is convergent between these two sister species groups (only with a gain of expression in HWD space-M
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<emphasis id="B95A2A364C5BC67A1D842C94FB9220A4" box="[932,1089,441,464]" italics="true" pageId="45" pageNumber="46">Ht. roussettae</emphasis>
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<taxonomicName id="4C2E8DA74C5BC67A1EB72FAAFE0B23D6" authority="Lees" authorityName="Lees" box="[151,472,647,674]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="kremenae" status="sp. nov.">
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<taxonomicName id="4C2E8DA74C5BC67A1EB72FAAFE0B23D6" authority="Lees" authorityName="Lees" box="[151,472,647,674]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="kremenae" status="sp. nov.">
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<emphasis id="B95A2A364C5BC67A1EB72FAAFE4523D5" bold="true" box="[151,406,647,673]" italics="true" pageId="45" pageNumber="46">Heteropsis kremenae</emphasis>
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Lees
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</taxonomicName>
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|
|
@ -76,7 +76,7 @@ material., Deposition
|
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<typeStatus id="549548864C5BC67A1CBD2E39FCD92259" box="[669,778,788,813]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
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</emphasis>
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: ♂ (
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<figureCitation id="1315EAA14C5BC67A1D732E39FC7D2258" box="[851,942,788,813]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="45" pageNumber="46">Fig. 13</figureCitation>
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<figureCitation id="1315EAA14C5BC67A1D732E39FC7D2258" box="[851,942,788,813]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.264610" httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="45" pageNumber="46">Fig. 13</figureCitation>
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A),
|
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<collectingCountry id="F339B6B44C5BC67A1DC52E39FBBF2259" box="[997,1132,788,813]" name="Madagascar" pageId="45" pageNumber="46">Madagascar</collectingCountry>
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NE, Ambavaony R. [= Ambanivony R.], R. Onive (Ankavanana), E Masoala,
|
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|
|
@ -153,7 +153,7 @@ barcode]; ♂, NE, Ambavaony R., E. Masoala,
|
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010289138 [
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<collectingCountry id="F339B6B44C5BC67A1D0C2934FCB42545" box="[812,871,1049,1073]" name="Qatar" pageId="45" pageNumber="46">QTR</collectingCountry>
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barcode]; ♀ (
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<figureCitation id="1315EAA14C5BC67A1A2A2935FB8D2545" box="[1034,1118,1048,1073]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="45" pageNumber="46">Fig. 13</figureCitation>
|
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<figureCitation id="1315EAA14C5BC67A1A2A2935FB8D2545" box="[1034,1118,1048,1073]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." figureDoi="http://doi.org/10.5281/zenodo.264610" httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="45" pageNumber="46">Fig. 13</figureCitation>
|
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B), NE, Masoala Peninsula, Manosona,
|
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<specimenCount id="9D283DAD4C5BC67A1F3C2912FE8C2523" box="[284,351,1087,1112]" pageId="45" pageNumber="46" type="generic">
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|
|
@ -199,7 +199,7 @@ voucher], E4 [egg voucher: “3 greenish eggs expressed 29/11”],
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010289140 [
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<collectingCountry id="F339B6B44C5BC67A1DDC2983FBE425B2" box="[1020,1079,1198,1222]" name="Qatar" pageId="45" pageNumber="46">QTR</collectingCountry>
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barcode]; ♂ (
|
||||
<figureCitation id="1315EAA14C5BC67A1AC72980FAEE25B2" box="[1255,1341,1197,1222]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="45" pageNumber="46">Fig. 15</figureCitation>
|
||||
<figureCitation id="1315EAA14C5BC67A1AC72980FAEE25B2" box="[1255,1341,1197,1222]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264612" httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="45" pageNumber="46">Fig. 15</figureCitation>
|
||||
A), NE, Masoala E, Ambanivony [=Ambavaony],
|
||||
<geoCoordinate id="EE1A90E34C5BC67A1C5029F9FD052599" box="[624,726,1233,1261]" direction="south" orientation="latitude" pageId="45" pageNumber="46" precision="55" value="-15.288">15.288o S</geoCoordinate>
|
||||
,
|
||||
|
|
@ -331,7 +331,7 @@ Apparently most similar to
|
|||
<emphasis id="B95A2A364C5BC67A1F442A28FDD12669" box="[356,514,1796,1821]" italics="true" pageId="45" pageNumber="46">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C5BC67A1C602A28FD1C2668" box="[576,719,1797,1820]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C5BC67A1C602A28FD1C2668" box="[576,719,1797,1820]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C5BC67A1C602A28FD1C2668" box="[576,719,1797,1820]" italics="true" pageId="45" pageNumber="46">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C5BC67A1CFA2A29FCE42669" box="[730,823,1796,1821]" pageId="45" pageNumber="46" rank="species">
|
||||
|
|
@ -385,11 +385,11 @@ showed for one specimen that the veins that were significantly inflated along th
|
|||
<paragraph id="8B91F6244C58C6791EE72981FE80277C" blockId="46.[151,1437,151,2015]" pageId="46" pageNumber="47">
|
||||
<emphasis id="B95A2A364C58C6791EE72981FE9F25B1" bold="true" box="[199,332,1196,1221]" pageId="46" pageNumber="47">♂ genitalia</emphasis>
|
||||
: 19DL, PT (
|
||||
<figureCitation id="1315EAA14C58C6791FC12981FDE025B1" box="[481,563,1196,1221]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="46" pageNumber="47">Fig. 15</figureCitation>
|
||||
<figureCitation id="1315EAA14C58C6791FC12981FDE025B1" box="[481,563,1196,1221]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264612" httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="46" pageNumber="47">Fig. 15</figureCitation>
|
||||
A);
|
||||
<bibRefCitation id="EFBF8BD54C58C6791C422980FC3125B0" author="Lees" box="[610,994,1196,1221]" pageId="46" pageNumber="47" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="f">
|
||||
Lees (1997: 106,
|
||||
<figureCitation id="1315EAA14C58C6791D0D2981FCA225B1" box="[813,881,1196,1221]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="46" pageNumber="47">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C58C6791D0D2981FCA225B1" box="[813,881,1196,1221]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="46" pageNumber="47">Fig. 7</figureCitation>
|
||||
f; “14A”)
|
||||
</bibRefCitation>
|
||||
: from LV: miniaturised, about
|
||||
|
|
@ -450,7 +450,7 @@ were also examined (LT ♂, here designated, that illustrated by
|
|||
<bibRefCitation id="EFBF8BD54C58C6791D0B2A53FC6426E3" author="Butler" box="[811,951,1918,1943]" pageId="46" pageNumber="47" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(BMNH(E) #674882) was examined (
|
||||
<figureCitation id="1315EAA14C58C6791B4F2A53FF6726CF" captionStart="FIGURE 14" captionStartId="48.[151,250,1802,1824]" captionTargetBox="[299,1270,199,1771]" captionTargetId="figure@48.[287,1299,193,1781]" captionTargetPageId="48" captionText="FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="46" pageNumber="47">Fig. 14</figureCitation>
|
||||
<figureCitation id="1315EAA14C58C6791B4F2A53FF6726CF" captionStart="FIGURE 14" captionStartId="48.[151,250,1802,1824]" captionTargetBox="[299,1270,199,1771]" captionTargetId="figure@48.[287,1299,193,1781]" captionTargetPageId="48" captionText="FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264611" httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="46" pageNumber="47">Fig. 14</figureCitation>
|
||||
A), with which species
|
||||
<taxonomicName id="4C2E8DA74C58C6791F9C2A8FFCFF26CF" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[444,812,1954,1979]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="undulata">
|
||||
<emphasis id="B95A2A364C58C6791F9C2A8FFDA926CF" box="[444,634,1954,1979]" italics="true" pageId="46" pageNumber="47">Culapa undulata</emphasis>
|
||||
|
|
@ -461,7 +461,7 @@ A), with which species
|
|||
: 185 was unable to locate) has been synonymised (Lees
|
||||
<emphasis id="B95A2A364C58C6791FFF2AE5FDCD26AB" box="[479,542,1990,2015]" italics="true" pageId="46" pageNumber="47">et al.,</emphasis>
|
||||
2003) (LT ♂, here designated,
|
||||
<figureCitation id="1315EAA14C58C6791D5D2AEBFC1E26AB" box="[893,973,1990,2015]" captionStart="FIGURE 14" captionStartId="48.[151,250,1802,1824]" captionTargetBox="[299,1270,199,1771]" captionTargetId="figure@48.[287,1299,193,1781]" captionTargetPageId="48" captionText="FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="46" pageNumber="47">Fig. 14</figureCitation>
|
||||
<figureCitation id="1315EAA14C58C6791D5D2AEBFC1E26AB" box="[893,973,1990,2015]" captionStart="FIGURE 14" captionStartId="48.[151,250,1802,1824]" captionTargetBox="[299,1270,199,1771]" captionTargetId="figure@48.[287,1299,193,1781]" captionTargetPageId="48" captionText="FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264611" httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="46" pageNumber="47">Fig. 14</figureCitation>
|
||||
C, bearing labels “
|
||||
<typeStatus id="549548864C58C6791ABA2AEBFADE26AB" box="[1178,1293,1990,2015]" pageId="46" pageNumber="47" type="lectotype">Lectotype</typeStatus>
|
||||
|
|
||||
|
|
@ -485,7 +485,7 @@ Culapa undulata
|
|||
) of
|
||||
<taxonomicName id="4C2E8DA74C47C6661DCE28C5FACB2775" authority="Oberthur" authorityName="Oberthur" box="[1006,1304,1512,1537]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="undulata">Culapa undulata Oberthür</taxonomicName>
|
||||
”|BMNH(E) #674883; Oberthür specified as STs 190 other ♂♂ (including BMNH(E) #674885) and 58 ♀♀ (including BMNH(E) #674884,
|
||||
<figureCitation id="1315EAA14C47C6661FAA2B1DFE0F273D" box="[394,476,1584,1609]" captionStart="FIGURE 14" captionStartId="48.[151,250,1802,1824]" captionTargetBox="[299,1270,199,1771]" captionTargetId="figure@48.[287,1299,193,1781]" captionTargetPageId="48" captionText="FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="49" pageNumber="50">Fig. 14</figureCitation>
|
||||
<figureCitation id="1315EAA14C47C6661FAA2B1DFE0F273D" box="[394,476,1584,1609]" captionStart="FIGURE 14" captionStartId="48.[151,250,1802,1824]" captionTargetBox="[299,1270,199,1771]" captionTargetId="figure@48.[287,1299,193,1781]" captionTargetPageId="48" captionText="FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 (‘ Antananarivo’; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 (‘ Antsianaka’; BMNH (E) # 674888). E: LT ♀ of Culapa pauper ‘ sp. or var. ’ pseudonarcissus Oberthür, 1916 (‘ Antakares’; BMNH (E) # 674890). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264611" httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="49" pageNumber="50">Fig. 14</figureCitation>
|
||||
B, illustrated in
|
||||
<bibRefCitation id="EFBF8BD54C47C6661CB62B1DFC93273D" author="Oberthur" box="[662,832,1584,1609]" pageId="49" pageNumber="50" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür 1916</bibRefCitation>
|
||||
: P. 367, f. 3065 and BMNH(E) #674886), which are then automatically PLTs of
|
||||
|
|
@ -494,13 +494,13 @@ B, illustrated in
|
|||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF51A6AC4C59C6781EB72AADFE492686" httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="47" pageNumber="48" targetBox="[243,1342,223,1890]" targetPageId="47">
|
||||
<caption id="DF51A6AC4C59C6781EB72AADFE492686" ID-DOI="http://doi.org/10.5281/zenodo.264610" httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="47" pageNumber="48" targetBox="[243,1342,223,1890]" targetPageId="47">
|
||||
<paragraph id="8B91F6244C59C6781EB72AADFE492686" blockId="47.[151,1436,1920,2034]" pageId="47" pageNumber="48">
|
||||
<emphasis id="B95A2A364C59C6781EB72AADFEF126E1" bold="true" box="[151,290,1920,1942]" pageId="47" pageNumber="48">FIGURE 13.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
<emphasis id="B95A2A364C59C6781CA22AADFD4726E1" bold="true" box="[642,660,1920,1941]" pageId="47" pageNumber="48">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C59C6781D292AACFBF226E2" authority="Lees" authorityName="Lees" box="[777,1057,1920,1942]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kremenae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C59C6781D292AACFBF226E2" authority="Lees" authorityName="Lees" box="[777,1057,1920,1942]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kremenae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C59C6781D292AACFC0C26E2" box="[777,991,1920,1942]" italics="true" pageId="47" pageNumber="48">Heteropsis kremenae</emphasis>
|
||||
, Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -511,13 +511,13 @@ Adults in DV (left), VV (right).
|
|||
(Ambavaony R., Masoala; NHMUK010289135).
|
||||
<emphasis id="B95A2A364C59C6781FA02AB2FE4226C0" bold="true" box="[384,401,1951,1972]" pageId="47" pageNumber="48">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C59C6781C232A8DFD5E26C1" box="[515,653,1951,1973]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C59C6781C232A8DFD5E26C1" box="[515,653,1951,1973]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C59C6781C232A8DFD5E26C1" box="[515,653,1951,1973]" italics="true" pageId="47" pageNumber="48">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
(Masoala; NHMUK010289139).
|
||||
<emphasis id="B95A2A364C59C6781DCF2AB2FBD226C1" bold="true" box="[1007,1025,1951,1973]" pageId="47" pageNumber="48">C</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C59C6781A572A8DFF3826A0" authority="Lees & Kremen" authorityName="Lees & Kremen" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C59C6781A572A8DFF3826A0" authority="Lees & Kremen" authorityName="Lees & Kremen" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C59C6781A572A8DFA9526C1" box="[1143,1350,1952,1973]" italics="true" pageId="47" pageNumber="48">Heteropsis avaratra</emphasis>
|
||||
Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -528,13 +528,13 @@ Lees & Kremen
|
|||
(Montagne d’Ambre; NHMUK010289141).
|
||||
<emphasis id="B95A2A364C59C6781D382A93FCF926A7" bold="true" box="[792,810,1982,2003]" pageId="47" pageNumber="48">D</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C59C6781DB52A92FBC626A0" box="[917,1045,1983,2004]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C59C6781DB52A92FBC626A0" box="[917,1045,1983,2004]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C59C6781DB52A92FBC626A0" box="[917,1045,1983,2004]" italics="true" pageId="47" pageNumber="48">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
(Montagne d’Ambre, Les Roussettes; DCL-DB-3385, MNHN).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF51A6AC4C46C6671EB72A27FEC026EE" httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="48" pageNumber="49" targetBox="[299,1270,199,1771]" targetPageId="48">
|
||||
<caption id="DF51A6AC4C46C6671EB72A27FEC026EE" ID-DOI="http://doi.org/10.5281/zenodo.264611" httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="48" pageNumber="49" targetBox="[299,1270,199,1771]" targetPageId="48">
|
||||
<paragraph id="8B91F6244C46C6671EB72A27FEC026EE" blockId="48.[151,1436,1802,1946]" pageId="48" pageNumber="49">
|
||||
<emphasis id="B95A2A364C46C6671EB72A27FEF3266B" bold="true" box="[151,288,1802,1824]" pageId="48" pageNumber="49">FIGURE 14.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
|
|
@ -579,13 +579,13 @@ Oberthür, 1916
|
|||
(‘Antakares’; BMNH(E) #674890). Scale bar: 30 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF51A6AC4C47C6661EB7286AFC5624EE" httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="49" pageNumber="50" targetBox="[386,1205,210,1306]" targetPageId="49">
|
||||
<caption id="DF51A6AC4C47C6661EB7286AFC5624EE" ID-DOI="http://doi.org/10.5281/zenodo.264612" httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="49" pageNumber="50" targetBox="[386,1205,210,1306]" targetPageId="49">
|
||||
<paragraph id="8B91F6244C47C6661EB7286AFC5624EE" blockId="49.[151,1436,1351,1434]" pageId="49" pageNumber="50">
|
||||
<emphasis id="B95A2A364C47C6661EB7286AFEF22428" bold="true" box="[151,289,1351,1373]" pageId="49" pageNumber="50">FIGURE 15.</emphasis>
|
||||
♂ genitalia in LV (left), DV (right) unless otherwise stated.
|
||||
<emphasis id="B95A2A364C47C6661DBC286AFC7D2428" bold="true" box="[924,942,1351,1372]" pageId="49" pageNumber="50">A</emphasis>
|
||||
:
|
||||
<taxonomicName id="4C2E8DA74C47C6661D9E2865FB1C2429" authority="Lees" authorityName="Lees" box="[958,1231,1351,1373]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="kremenae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C47C6661D9E2865FB1C2429" authority="Lees" authorityName="Lees" box="[958,1231,1351,1373]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="kremenae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C47C6661D9E2865FB462429" box="[958,1173,1351,1373]" italics="true" pageId="49" pageNumber="50">Heteropsis kremenae</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -596,7 +596,7 @@ Lees
|
|||
(Masoala, Ambanivony; 19DL).
|
||||
<emphasis id="B95A2A364C47C6661F5B284BFE5F240F" bold="true" box="[379,396,1382,1403]" pageId="49" pageNumber="50">B</emphasis>
|
||||
:
|
||||
<taxonomicName id="4C2E8DA74C47C6661FBB284AFD1A2408" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[411,713,1382,1404]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C47C6661FBB284AFD1A2408" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[411,713,1382,1404]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C47C6661FBB284AFDC82408" box="[411,539,1383,1404]" italics="true" pageId="49" pageNumber="50">Ht. avaratra</emphasis>
|
||||
Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -607,7 +607,7 @@ Lees & Kremen
|
|||
(315DL, referred specimen, Joffreville).
|
||||
<emphasis id="B95A2A364C47C6661AF0284BFB312408" bold="true" box="[1232,1250,1382,1404]" pageId="49" pageNumber="50">C</emphasis>
|
||||
:
|
||||
<taxonomicName id="4C2E8DA74C47C6661AD1284AFF1A24EE" authority="Lees" authorityName="Lees" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="sogai" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C47C6661AD1284AFF1A24EE" authority="Lees" authorityName="Lees" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="sogai" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C47C6661AD1284AFA4F2408" box="[1265,1436,1383,1404]" italics="true" pageId="49" pageNumber="50">Heteropsis sogai</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -621,7 +621,7 @@ PT (Tsaratanana; 96DL, MNHN), LV, SV. Scale bar: 3 mm.
|
|||
<paragraph id="8B91F6244C47C6651EE72B55FBE02068" blockId="49.[151,1436,1473,2005]" lastBlockId="50.[151,1437,151,1930]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
|
||||
<emphasis id="B95A2A364C47C6661EE72B55FD1827E5" bold="true" box="[199,715,1656,1681]" pageId="49" pageNumber="50">Additional information. DNA divergences</emphasis>
|
||||
: cluster number BOLD:ACW4937 (exemplar BMAD249-15, Marojejy), diverges by 5.47% from that of
|
||||
<taxonomicName id="4C2E8DA74C47C6661CA32BB3FCDA27C1" box="[643,777,1694,1717]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C47C6661CA32BB3FCDA27C1" box="[643,777,1694,1717]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C47C6661CA32BB3FCDA27C1" box="[643,777,1694,1717]" italics="true" pageId="49" pageNumber="50">H. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
(cluster number BOLD:AAD0195, exemplar BMAD016- 09) and by about 5.5% from that of
|
||||
|
|
@ -635,13 +635,13 @@ Torres
|
|||
(2001)
|
||||
</bibRefCitation>
|
||||
dataset, a COII sequence for
|
||||
<taxonomicName id="4C2E8DA74C47C6661FFC2BCBFDA12789" box="[476,626,1764,1789]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C47C6661FFC2BCBFDA12789" box="[476,626,1764,1789]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C47C6661FFC2BCBFDA12789" box="[476,626,1764,1789]" italics="true" pageId="49" pageNumber="50">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
(their “
|
||||
<emphasis id="B95A2A364C47C6661CE92BCBFD242789" box="[713,759,1766,1789]" italics="true" pageId="49" pageNumber="50">Hen</emphasis>
|
||||
. sp. 14A”) from Masoala is 7.03% pairwise divergent to
|
||||
<taxonomicName id="4C2E8DA74C47C6661B592BCBFF2A2655" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C47C6661B592BCBFF2A2655" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C47C6661B592BCBFF2A2655" italics="true" pageId="49" pageNumber="50">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
from Montagne d’Ambre (
|
||||
|
|
@ -673,7 +673,7 @@ AY
|
|||
<date id="FF90D0E44C47C6661D852A7CFC28261D" box="[933,1019,1873,1897]" pageId="49" pageNumber="50" value="33-04-01">040133</date>
|
||||
</accessionNumber>
|
||||
, 414 bp compared). Meanwhile,
|
||||
<taxonomicName id="4C2E8DA74C47C6661B592A7FFF2A26F9" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C47C6661B592A7FFF2A26F9" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C47C6661B592A7FFF2A26F9" italics="true" pageId="49" pageNumber="50">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -735,7 +735,7 @@ Linares
|
|||
<paragraph id="8B91F6244C44C6651EE72C0AFF2D20D8" blockId="50.[151,1437,151,1930]" pageId="50" pageNumber="51">
|
||||
<emphasis id="B95A2A364C44C6651EE72C0AFE362034" bold="true" box="[199,485,295,320]" pageId="50" pageNumber="51">Phylogeny/sister species</emphasis>
|
||||
:
|
||||
<taxonomicName id="4C2E8DA74C44C6651FD42C04FD582034" box="[500,651,295,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C44C6651FD42C04FD582034" box="[500,651,295,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C44C6651FD42C04FD582034" box="[500,651,295,320]" italics="true" pageId="50" pageNumber="51">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
is likely most closely related to
|
||||
|
|
@ -743,13 +743,13 @@ is likely most closely related to
|
|||
<emphasis id="B95A2A364C44C6651DDA2C04FB472034" box="[1018,1172,295,320]" italics="true" pageId="50" pageNumber="51">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C44C6651AEC2C04FA8B2034" box="[1228,1368,297,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<taxonomicName id="4C2E8DA74C44C6651AEC2C04FA8B2034" box="[1228,1368,297,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<emphasis id="B95A2A364C44C6651AEC2C04FA8B2034" box="[1228,1368,297,320]" italics="true" pageId="50" pageNumber="51">Ht. avaratra</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
<bibRefCitation id="EFBF8BD54C44C6651B472C05FF322010" author="Lees" pageId="50" pageNumber="51" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997)</bibRefCitation>
|
||||
did not resolve the position of
|
||||
<taxonomicName id="4C2E8DA74C44C6651C1F2C60FD062011" box="[575,725,332,357]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<taxonomicName id="4C2E8DA74C44C6651C1F2C60FD062011" box="[575,725,332,357]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="kremenae">
|
||||
<emphasis id="B95A2A364C44C6651C1F2C60FD062011" box="[575,725,332,357]" italics="true" pageId="50" pageNumber="51">Ht. kremenae</emphasis>
|
||||
</taxonomicName>
|
||||
‘FRTNA’] within the
|
||||
|
|
@ -794,7 +794,7 @@ Aduse-Poku
|
|||
<paragraph id="8B91F6244C44C6651EE72FA2FCC923B8" blockId="50.[151,1437,151,1930]" pageId="50" pageNumber="51">
|
||||
<emphasis id="B95A2A364C44C6651EE72FA2FE8A23DC" bold="true" box="[199,345,655,680]" pageId="50" pageNumber="51">Distribution</emphasis>
|
||||
: endemic to lowland rainforest in the northeast of the rainforest belt and Masoala Peninsula (
|
||||
<figureCitation id="1315EAA14C44C6651B4E2FA2FF6623B8" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="50" pageNumber="51">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C44C6651B4E2FA2FF6623B8" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="50" pageNumber="51">Fig. 30</figureCitation>
|
||||
B, dark blue dots), with a preference for riparian areas.
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C44C6651EE72FFAFB582384" blockId="50.[151,1437,151,1930]" box="[199,1163,727,752]" pageId="50" pageNumber="51">
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
<document id="427EA61E09F951BC09627A407C5A60B7" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C5EC67C1EB7292BFD3E27E9" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteropsis imerina Lees, sp. nov." docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="41" updateTime="1698662715872" updateUser="ExternalLinkService">
|
||||
<mods:mods id="2ED04E853205CA47BCEA91CFC41F8F3B" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="F60672CFA242BB30308590156336FFBD">
|
||||
<mods:title id="DD4D57D5CCEF941B6C9D5DFFBC4759B0">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
|
||||
<document id="4468AD885170DB362DD62D2EF85BD2EA" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C5EC67C1EB7292BFD3E27E9" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Heteropsis imerina Lees, sp. nov." docType="treatment" docVersion="12" lastPageNumber="43" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="41" updateTime="1735917000942" updateUser="ExternalLinkService">
|
||||
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|
||||
<mods:titleInfo id="B914F0A4B1C7EEEA04E0205B21051B34">
|
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<mods:title id="828D5633C66D766B6F15BE4E8B4C8CA4">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
|
||||
</mods:titleInfo>
|
||||
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<mods:namePart id="D3C626801AFB4B769BAE2E2A78B632AB">C, Lees David</mods:namePart>
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<mods:date id="65C6BF2CA4BDDF36F12BD1FBEB30B08F">2016</mods:date>
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<mods:identifier id="EE3E51197DEAD020A53B211F6B4830B6" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
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<treatment id="038747324C5EC67C1EB7292BFD3E27E9" ID-DOI="http://doi.org/10.5281/zenodo.6086428" ID-GBIF-Taxon="120817089" ID-Zenodo-Dep="6086428" LSID="urn:lsid:plazi:treatment:038747324C5EC67C1EB7292BFD3E27E9" httpUri="http://treatment.plazi.org/id/038747324C5EC67C1EB7292BFD3E27E9" lastPageId="43" lastPageNumber="43" pageId="40" pageNumber="41">
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<treatment id="038747324C5EC67C1EB7292BFD3E27E9" ID-DOI="http://doi.org/10.5281/zenodo.6086428" ID-GBIF-Taxon="120817089" ID-Zenodo-Dep="6086428" LSID="urn:lsid:plazi:treatment:038747324C5EC67C1EB7292BFD3E27E9" httpUri="http://treatment.plazi.org/id/038747324C5EC67C1EB7292BFD3E27E9" lastPageId="43" lastPageNumber="43" pageId="40" pageNumber="41" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C5EC67F1EB7292BFBCA2535" pageId="40" pageNumber="41" type="nomenclature">
|
||||
<paragraph id="8B91F6244C5EC67F1EB7292BFDF92554" blockId="40.[151,1049,1030,1089]" box="[151,554,1030,1056]" pageId="40" pageNumber="41">
|
||||
<heading id="D0D941484C5EC67F1EB7292BFDF92554" bold="true" box="[151,554,1030,1056]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
|
||||
<emphasis id="B95A2A364C5EC67F1EB7292BFDF92554" bold="true" box="[151,554,1030,1056]" pageId="40" pageNumber="41">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1EB7292BFE6C2554" authority="Lees" authorityName="Lees" box="[151,447,1030,1056]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1EB7292BFE6C2554" authority="Lees" authorityName="Lees" box="[151,447,1030,1056]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C5EC67F1EB7292BFEAE2554" bold="true" box="[151,381,1030,1056]" italics="true" pageId="40" pageNumber="41">Heteropsis imerina</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -85,7 +85,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C5EC67F1C5B2995FD3B25A5" box="[635,744,1208,1233]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C5EC67F1D002995FCB525A5" box="[800,870,1208,1233]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="40" pageNumber="41">Fig. 9</figureCitation>
|
||||
<figureCitation id="1315EAA14C5EC67F1D002995FCB525A5" box="[800,870,1208,1233]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="40" pageNumber="41">Fig. 9</figureCitation>
|
||||
C),
|
||||
<collectingCountry id="F339B6B44C5EC67F1DB22995FBCA25A5" box="[914,1049,1208,1233]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
|
||||
E, Mantadia National Park, Jofa, near bridge,
|
||||
|
|
@ -147,7 +147,7 @@ barcode];
|
|||
E. route d’Anosibe
|
||||
<date id="FF90D0E44C5EC67F1D7428A3FC6324D3" box="[852,944,1422,1447]" pageId="40" pageNumber="41" value="1966-06">VI-1966</date>
|
||||
/MUSEUM PARIS COLL. G. DUJARDIN DEL. ET R. VIOSSAT|DCL-DB-2908; ♀ (
|
||||
<figureCitation id="1315EAA14C5EC67F1CA3289FFD1B24BF" box="[643,712,1458,1483]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="40" pageNumber="41">Fig. 9</figureCitation>
|
||||
<figureCitation id="1315EAA14C5EC67F1CA3289FFD1B24BF" box="[643,712,1458,1483]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="40" pageNumber="41">Fig. 9</figureCitation>
|
||||
D),
|
||||
<collectingCountry id="F339B6B44C5EC67F1CD5289FFCAF24BF" box="[757,892,1458,1483]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
|
||||
Est, route de Lakato km 15, Ankasoka
|
||||
|
|
@ -186,41 +186,41 @@ E., Mantadia National Park, Jofa, near bridge,
|
|||
<paragraph id="8B91F6244C5EC67F1EE72B48FB94260C" blockId="40.[151,1436,1136,2019]" pageId="40" pageNumber="41">
|
||||
<emphasis id="B95A2A364C5EC67F1EE72B48FE92270A" bold="true" box="[199,321,1637,1662]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
|
||||
Very distinct by its shorter HW tails from the somewhat similar
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1A6F2B4BFB09270A" box="[1103,1242,1637,1662]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1A6F2B4BFB09270A" box="[1103,1242,1637,1662]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pauliani" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C5EC67F1A6F2B4BFB09270A" box="[1103,1242,1637,1662]" italics="true" pageId="40" pageNumber="41">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C5EC67F1AC62B48FA96270A" box="[1254,1349,1637,1662]" pageId="40" pageNumber="41" rank="species">
|
||||
<emphasis id="B95A2A364C5EC67F1AC62B48FA96270A" bold="true" box="[1254,1349,1637,1662]" pageId="40" pageNumber="41">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
, where potentially (see discussion) sympatric (from RS Anjanaharibe Sud to RNI Tsaratanana).
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1AA22BA7FA8427D5" box="[1154,1367,1672,1697]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1AA22BA7FA8427D5" box="[1154,1367,1672,1697]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C5EC67F1AA22BA7FA8427D5" box="[1154,1367,1672,1697]" italics="true" pageId="40" pageNumber="41">Heteropsis harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
is the species most likely to be confused in the main central-eastern rainforest block, from which
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1A852B83FAF427B0" box="[1189,1319,1709,1733]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1A852B83FAF427B0" box="[1189,1319,1709,1733]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C5EC67F1A852B83FAF427B0" box="[1189,1319,1709,1733]" italics="true" pageId="40" pageNumber="41">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
differs by lack of an androconial patch on the FWV, by the proximad inflexion of the HWV medial band around ocellus space-CuA1 being rather sharply angled, and by the more evenly pronounced HW tails of the former, as for the other small dark species (in the case of ♂♂).
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1C8B2A34FC8D2644" box="[683,862,1815,1840]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="vanewrighti" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1C8B2A34FC8D2644" box="[683,862,1815,1840]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="vanewrighti" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C5EC67F1C8B2A34FC8D2644" box="[683,862,1815,1840]" italics="true" pageId="40" pageNumber="41">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C5EC67F1D472A3AFC172644" box="[871,964,1815,1840]" pageId="40" pageNumber="41" rank="species">
|
||||
<emphasis id="B95A2A364C5EC67F1D472A3AFC172644" bold="true" box="[871,964,1815,1840]" pageId="40" pageNumber="41">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1DF42A34FB962644" box="[980,1093,1817,1840]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1DF42A34FB962644" box="[980,1093,1817,1840]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C5EC67F1DF42A34FB962644" box="[980,1093,1817,1840]" italics="true" pageId="40" pageNumber="41">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1AA22A34FAF72644" box="[1154,1316,1815,1840]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1AA22A34FAF72644" box="[1154,1316,1815,1840]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<emphasis id="B95A2A364C5EC67F1AA22A34FAF72644" box="[1154,1316,1815,1840]" italics="true" pageId="40" pageNumber="41">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
are easily distinguished from
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1F532A10FE252620" box="[371,502,1853,1876]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1F532A10FE252620" box="[371,502,1853,1876]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C5EC67F1F532A10FE252620" box="[371,502,1853,1876]" italics="true" pageId="40" pageNumber="41">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
by their whitish highlighting distad of the HWV Mb, which in
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1AEC2A10FA9D2620" box="[1228,1358,1853,1876]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C5EC67F1AEC2A10FA9D2620" box="[1228,1358,1853,1876]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C5EC67F1AEC2A10FA9D2620" box="[1228,1358,1853,1876]" italics="true" pageId="40" pageNumber="41">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
♂♂ is yellowish, sometimes with a reddish suffusion proximad of the space-CuA1 ocellus.
|
||||
|
|
@ -230,15 +230,15 @@ by their whitish highlighting distad of the HWV Mb, which in
|
|||
<paragraph id="8B91F6244C5EC67E1EE72AAEFD3E2223" blockId="40.[151,1436,1136,2019]" lastBlockId="41.[151,1437,151,2037]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
|
||||
<emphasis id="B95A2A364C5EC67F1EE72AAEFE7E26E8" bold="true" box="[199,429,1923,1948]" pageId="40" pageNumber="41">Description. Wings</emphasis>
|
||||
: upperside uniform mid to dark brown (costad area not much darker) with space-CuA1 ocellus well expressed on FWD, with a round concentric orange-reddish ring, its black iris almost spanning or spanning CuA1-CuA2, and the Orng compromising these veins and moderately narrow and eccentric. FWD space- M1 ocellus not evident. HWD space-CuA1 ocellus also hardly evident. FWV space-CuA1 ocellus similar size to FWD and marginally more eccentric with mid orange ring slightly skewed to tergum. Space-M1 ocellus much smaller and with a small black iris and a faint mid orange ring; that in space-M2 almost reduced to a small white spot. HWV space-CuA1 ocellus relatively small compared with others of
|
||||
<taxonomicName id="4C2E8DA74C5FC67E1DF42C29FB882068" box="[980,1115,259,284]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C5FC67E1DF42C29FB882068" box="[980,1115,259,284]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C5FC67E1DF42C29FB882068" box="[980,1115,259,284]" italics="true" pageId="41" pageNumber="42">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group and subelliptic with a narrow mid orange ring. Other HWV ocelli, particularly those of spaces Rs-M3, also of space-CuA2-1A, expressed as white spots. Ventral wing background colour with a distinct dark reddish brown cast. FWV more uniform than HWV but paler towards tergal angle with a pale ochreous mark near costa towards apex, HWV lighter brown distad of Mb. This dark russet brown Mb is irregularly snaking, concave around its intersection with M1, convex around that with M2, curving back to the deepest concavity in space-CuA1 and convex again near vein CuA2 before terminating at 1A. This line is highlighted with ochreous admixture with reddish brown scales distad of it and sombrely shaded proximad of it, with a more consistent irroration with russet brown contrasting with more ochreous brown scales in the basal areas; the basal HWV areas are thus relatively dark in the ♂, while the PMb is indistinct and basad of it is also ochreous highlighted. Before the margin in both wings is a slightly more ochreous highlighting and a double thin russet-brown Pml (pre-marginal line), the outer of which follows the margin, which is lightly crenate. The slightly more prominent but short dark brown ‘tail’ is at the end of M3, which can be a field character distinguishing from similar members of the
|
||||
<taxonomicName id="4C2E8DA74C5FC67E1CCD2F9CFCA023BC" box="[749,883,687,712]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C5FC67E1CCD2F9CFCA023BC" box="[749,883,687,712]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C5FC67E1CCD2F9CFCA023BC" box="[749,883,687,712]" italics="true" pageId="41" pageNumber="42">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group, particularly in the ♀ (
|
||||
<figureCitation id="1315EAA14C5FC67E1A972F82FB2923BC" box="[1207,1274,687,712]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="41" pageNumber="42">Fig. 9</figureCitation>
|
||||
<figureCitation id="1315EAA14C5FC67E1A972F82FB2923BC" box="[1207,1274,687,712]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="41" pageNumber="42">Fig. 9</figureCitation>
|
||||
D).
|
||||
<emphasis id="B95A2A364C5FC67E1B042F82FA4F23BC" bold="true" box="[1316,1436,687,712]" pageId="41" pageNumber="42">Variation.</emphasis>
|
||||
♂♂ vary quite a lot in their underside colouration, and the HWD space-CuA1 ocellus is sometimes expressed. White spots not always evident on HWV. ♀ is similar but larger, lighter brown on upperside and with more ochreous cast on upperside with PMb and Mb russet brown on HWV, thus more pronounced. In apparent dry seasonal forms, the HWV ocelli can be very reduced.
|
||||
|
|
@ -280,7 +280,7 @@ by Claire Kremen (
|
|||
) and as sp. 76 by the same worker in
|
||||
<date id="FF90D0E44C5FC67E1EB72822FEC6245C" box="[151,277,1295,1320]" pageId="41" pageNumber="42" value="1995" valueMax="1996">1995–1996</date>
|
||||
at Anjanaharibe Sud, but there were earlier collections, mostly represented in MNHN. The morphospecies ‘sp. 76’ (Anjanaharibe Sud) is definitely referable to
|
||||
<taxonomicName id="4C2E8DA74C5FC67E1DBA2819FBC8243F" box="[922,1051,1332,1355]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C5FC67E1DBA2819FBC8243F" box="[922,1051,1332,1355]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C5FC67E1DBA2819FBC8243F" box="[922,1051,1332,1355]" italics="true" pageId="41" pageNumber="42">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
, although its ♂ genitalia had been considered distinct in
|
||||
|
|
@ -288,7 +288,7 @@ at Anjanaharibe Sud, but there were earlier collections, mostly represented in M
|
|||
; spp. ‘22’ and ‘76’ were recovered anyway as sister taxa in
|
||||
<bibRefCitation id="EFBF8BD54C5FC67E1AF72875FA4B241B" author="Lees" box="[1239,1432,1366,1391]" pageId="41" pageNumber="42" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997: 156)</bibRefCitation>
|
||||
, but the recent discovery by myself of additional populations with adult phenotype similar to
|
||||
<taxonomicName id="4C2E8DA74C5FC67E1AF62851FA8824E7" box="[1238,1371,1404,1427]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C5FC67E1AF62851FA8824E7" box="[1238,1371,1404,1427]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C5FC67E1AF62851FA8824E7" box="[1238,1371,1404,1427]" italics="true" pageId="41" pageNumber="42">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
from Manongarivo through Tsaratanana to Marojejy and Anjanaharibe Sud suggest the species is indeed quite widespread. A deeper study of variation in the genitalic variation would be interesting, especially if backed up by molecular sequences; the study of
|
||||
|
|
@ -304,11 +304,11 @@ also showed minimal differences in COII (see below).
|
|||
: 154DL (not in
|
||||
<typeStatus id="549548864C5FC67E1D612B27FCA22756" box="[833,881,1546,1570]" pageId="41" pageNumber="42">type</typeStatus>
|
||||
series,
|
||||
<figureCitation id="1315EAA14C5FC67E1DEF2B24FBC72756" box="[975,1044,1545,1570]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="41" pageNumber="42">Fig. 8</figureCitation>
|
||||
<figureCitation id="1315EAA14C5FC67E1DEF2B24FBC72756" box="[975,1044,1545,1570]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264605" httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="41" pageNumber="42">Fig. 8</figureCitation>
|
||||
C): from LV, tegumen with fairly straight dorsal profile (strong v-shaped notch proximad from DV), leading to narrowly scythe-hooked uncus without a strong ventral ‘dewlap’. Gnathos relatively thin and sinuate, tapering, first downcurved and then upcurved (also sinuate from DV). Constriction in division with vinculum about half maximum dimension of tegumen. Vinculum not strongly bowed and broadening towards base of saccus. Valves relatively long, with distinctly angled shoulder, and stretched out valve base narrowing gradually to valve arm which is slightly longer than valve base with distinct ventral ‘elbow’, leading to spatulate tips with a limited number of spinoid setae on inner face/at tip (valve arms gently incurved from DV without a strong terminal spine but with a small number of teeth on mesad edge towards tip). Valve tips protrude considerably proud of maximum extension of uncus tip (
|
||||
<bibRefCitation id="EFBF8BD54C5FC67E1B462A05FEA12610" author="Lees" pageId="41" pageNumber="42" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="i">
|
||||
Lees 1997: 109,
|
||||
<figureCitation id="1315EAA14C5FC67E1F012A66FEB92610" box="[289,362,1867,1892]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="41" pageNumber="42">
|
||||
<figureCitation id="1315EAA14C5FC67E1F012A66FEB92610" box="[289,362,1867,1892]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="41" pageNumber="42">
|
||||
Fig.
|
||||
<date id="FF90D0E44C5FC67E1F7A2A66FEB92610" box="[346,362,1867,1892]" pageId="41" pageNumber="42">7</date>
|
||||
</figureCitation>
|
||||
|
|
@ -333,7 +333,7 @@ i
|
|||
, BMNH(E) #676685
|
||||
<accessionNumber id="947D6BC74C5CC67D1F212DF2FEA2218C" box="[257,369,223,248]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819267" pageId="42" pageNumber="43">FJ819267</accessionNumber>
|
||||
) is about 4.2% pairwise divergent to those of
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1D5D2DCDFC2F218C" box="[893,1020,223,248]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1D5D2DCDFC2F218C" box="[893,1020,223,248]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C5CC67D1D5D2DCDFC2F218C" box="[893,1020,223,248]" italics="true" pageId="42" pageNumber="43">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
described here (BMNH(E) #671648
|
||||
|
|
@ -347,25 +347,25 @@ Monaghan
|
|||
, 2009
|
||||
</bibRefCitation>
|
||||
; additional members of the
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1A902C29FAE42068" box="[1200,1335,259,284]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1A902C29FAE42068" box="[1200,1335,259,284]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C5CC67D1A902C29FAE42068" box="[1200,1335,259,284]" italics="true" pageId="42" pageNumber="43">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group in that study include
|
||||
<emphasis id="B95A2A364C5CC67D1F582C05FCB1204B" box="[376,866,294,319]" italics="true" pageId="42" pageNumber="43">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1F582C05FDE3204B" box="[376,560,294,319]" class="Insecta" family="Nymphalidae" genus="Erebia" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="passandava">Ht. passandava</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1C622C05FD10204B" box="[578,707,296,319]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="narova">Ht. narova</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1C622C05FD10204B" box="[578,707,296,319]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="narova">Ht. narova</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1CF52C05FCB1204B" box="[725,866,294,319]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="difficilis">Ht. difficilis</taxonomicName>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1D852C05FBC5204B" box="[933,1046,296,319]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="strato">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1D852C05FBC5204B" box="[933,1046,296,319]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="strato">
|
||||
<emphasis id="B95A2A364C5CC67D1D852C05FBC5204B" box="[933,1046,296,319]" italics="true" pageId="42" pageNumber="43">Ht. strato</emphasis>
|
||||
</taxonomicName>
|
||||
). A 706-bp relatively conserved 28SDDFF sequence (BMNH(E) #676640,
|
||||
<accessionNumber id="947D6BC74C5CC67D1CA82C67FD2B2017" box="[648,760,330,355]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ817836" pageId="42" pageNumber="43">FJ817836</accessionNumber>
|
||||
) as aligned is identical to those of
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1A802C61FADC2017" box="[1184,1295,332,355]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="strato">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1A802C61FADC2017" box="[1184,1295,332,355]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="strato">
|
||||
<emphasis id="B95A2A364C5CC67D1A802C61FADC2017" box="[1184,1295,332,355]" italics="true" pageId="42" pageNumber="43">Ht. strato</emphasis>
|
||||
</taxonomicName>
|
||||
(BMNH(E) #671544
|
||||
|
|
@ -401,13 +401,13 @@ Torres
|
|||
2001
|
||||
</bibRefCitation>
|
||||
had ‘sp. 76’ as sister to
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1EDB2CF7FEAC2085" box="[251,383,474,497]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1EDB2CF7FEAC2085" box="[251,383,474,497]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C5CC67D1EDB2CF7FEAC2085" box="[251,383,474,497]" italics="true" pageId="42" pageNumber="43">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
(their ‘
|
||||
<emphasis id="B95A2A364C5CC67D1FF82CF7FDD42085" box="[472,519,474,497]" italics="true" pageId="42" pageNumber="43">Hen</emphasis>
|
||||
. sp. 22’) with 100% bootstrap support. In their dataset, with 417 bp compared, ignoring two differences at the 5’ end,
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1C6F2CD3FD032361" box="[591,720,510,533]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1C6F2CD3FD032361" box="[591,720,510,533]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C5CC67D1C6F2CD3FD032361" box="[591,720,510,533]" italics="true" pageId="42" pageNumber="43">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
|
|
@ -427,19 +427,19 @@ AY
|
|||
, based on 2 ♂♂ from RS Anjanaharibe Sud). According to the ‘total evidence’ morphological analysis of
|
||||
<bibRefCitation id="EFBF8BD54C5CC67D1B472F68FEB923F5" author="Lees" pageId="42" pageNumber="43" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
|
||||
Lees (1997: 156,
|
||||
<figureCitation id="1315EAA14C5CC67D1F3F2F45FEB323F5" box="[287,352,616,641]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="42" pageNumber="43">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C5CC67D1F3F2F45FEB323F5" box="[287,352,616,641]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="42" pageNumber="43">Fig. 1</figureCitation>
|
||||
)
|
||||
</bibRefCitation>
|
||||
, there was 0.963 jackknife support for a sister relationship of these exemplars with
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1B342F44FA4823F5" box="[1300,1435,616,641]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1B342F44FA4823F5" box="[1300,1435,616,641]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<emphasis id="B95A2A364C5CC67D1B342F44FA4823F5" box="[1300,1435,616,641]" italics="true" pageId="42" pageNumber="43">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
(“sp. 37”). There is some support for
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1C152FA0FD6523D0" box="[565,694,653,676]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1C152FA0FD6523D0" box="[565,694,653,676]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C5CC67D1C152FA0FD6523D0" box="[565,694,653,676]" italics="true" pageId="42" pageNumber="43">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
as the sister of
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1D422FA0FC3A23D1" box="[866,1001,652,677]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1D422FA0FC3A23D1" box="[866,1001,652,677]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<emphasis id="B95A2A364C5CC67D1D422FA0FC3A23D1" box="[866,1001,652,677]" italics="true" pageId="42" pageNumber="43">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
in the combined tree of
|
||||
|
|
@ -474,7 +474,7 @@ occurs among native climbing bamboos, which it might feed on.
|
|||
<paragraph id="8B91F6244C5CC67D1EE72EE0FAE125ED" blockId="42.[151,1437,151,2030]" pageId="42" pageNumber="43">
|
||||
<emphasis id="B95A2A364C5CC67D1EE72EE0FE8A2292" bold="true" box="[199,345,973,998]" pageId="42" pageNumber="43">Distribution</emphasis>
|
||||
: widespread in the principal eastern rainforest belt and also in high plateau relics notably Ankazomivady south of Ambositra (
|
||||
<figureCitation id="1315EAA14C5CC67D1C1D2EDCFD42257E" box="[573,657,1009,1034]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="42" pageNumber="43">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C5CC67D1C1D2EDCFD42257E" box="[573,657,1009,1034]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="42" pageNumber="43">Fig. 30</figureCitation>
|
||||
A, turquoise dots). The locality was incorrectly given in Table 1 of
|
||||
<bibRefCitation id="EFBF8BD54C5CC67D1EB7293BFEB4255A" author="Torres" box="[151,359,1045,1070]" pageId="42" pageNumber="43" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. & Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
|
||||
Torres
|
||||
|
|
@ -484,7 +484,7 @@ Torres
|
|||
(which contains a formatting glitch) as ‘Anjanaharibe Sud, 1240 m’ instead of the line below, ‘
|
||||
<quantity id="4CD65BC14C5CC67D1EBC2914FF342526" box="[156,231,1081,1106]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="42" pageNumber="43" unit="km" value="32.0">32 km</quantity>
|
||||
S. of Ambositra’. The species extends to the northern mountain range of the rainforest biome. ‘Sp. 76’ from RS Anjanaharibe Sud is doubtless referable to
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1CF72973FC8C2500" box="[727,863,1117,1141]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1CF72973FC8C2500" box="[727,863,1117,1141]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C5CC67D1CF72973FC8C2500" box="[727,863,1117,1141]" italics="true" pageId="42" pageNumber="43">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
, as for material (‘sp. 65’) from Bekolosy (RS Manongarivo) and RNI Tsaratanana, while this is a species for which phylogeographic study is desirable.
|
||||
|
|
@ -516,7 +516,7 @@ E, Bemoara camp (going out from to Ambavala), Zahamena PN [northwestern sector],
|
|||
, D.C. Lees:
|
||||
<collectionCode id="ED3F6EE14C5CC67D1ADB283FFAF0245F" box="[1275,1315,1298,1323]" pageId="42" pageNumber="43">DL</collectionCode>
|
||||
06-023; ♂,
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1EB72818FF3D243A" box="[151,238,1333,1358]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="93" phylum="Arthropoda" rank="species" species="imerina">Imerina</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1EB72818FF3D243A" box="[151,238,1333,1358]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="93" phylum="Arthropoda" rank="species" species="imerina">Imerina</taxonomicName>
|
||||
<collectingCountry id="F339B6B44C5CC67D1EDA2818FE55243A" box="[250,390,1333,1358]" name="Madagascar" pageId="42" pageNumber="43">Madagascar</collectingCountry>
|
||||
|1910 E. La Moult |
|
||||
<collectionCode id="ED3F6EE14C5CC67D1C4A281BFD69243A" box="[618,698,1334,1358]" pageId="42" pageNumber="43">COLL</collectionCode>
|
||||
|
|
@ -525,7 +525,7 @@ E, Bemoara camp (going out from to Ambavala), Zahamena PN [northwestern sector],
|
|||
|
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1A11281BFB30243A" box="[1073,1251,1333,1358]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andravahana">H. andravahana</taxonomicName>
|
||||
ab. marmorata, Auriv. [sic]|DCL-DB-2897; ♀,
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1FDB2874FD812406" box="[507,594,1369,1394]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="93" phylum="Arthropoda" rank="species" species="imerina">Imerina</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C5CC67D1FDB2874FD812406" box="[507,594,1369,1394]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="93" phylum="Arthropoda" rank="species" species="imerina">Imerina</taxonomicName>
|
||||
<collectingCountry id="F339B6B44C5CC67D1C7A2874FD352406" box="[602,742,1369,1394]" name="Madagascar" pageId="42" pageNumber="43">Madagascar</collectingCountry>
|
||||
|1910 E. La Moult |DCL-DB-2911; ♂ [
|
||||
<collectionCode id="ED3F6EE14C5CC67D1ABC2877FB292405" box="[1180,1274,1370,1393]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="42" pageNumber="43">MNHN</collectionCode>
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
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<mods:identifier id="EFE889BC7B4544E5DF72C01CB84857FF" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
|
||||
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|
||||
<mods:identifier id="EEB6080C488E07B681A5CA5B9C80D445" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="F6FA9DC62E4CB38D046199795EADA2A3" type="Zenodo-Dep">264597</mods:identifier>
|
||||
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|
||||
<mods:classification id="C5CE80407B002F3AD4C1A4615FA50D39">journal article</mods:classification>
|
||||
<mods:identifier id="AFC687081CD9E45CF402158F01DF2765" type="CLB-Dataset">38930</mods:identifier>
|
||||
<mods:identifier id="2AB8199F125F7DA74D5E420020613EA9" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
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||||
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||||
<mods:identifier id="4D48DB612C35989B9DDD5D0F9DF1FF95" type="Zenodo-Dep">264597</mods:identifier>
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|
||||
</mods:mods>
|
||||
<treatment id="038747324C62C6401EB728FCFB6F2681" ID-DOI="http://doi.org/10.5281/zenodo.6086416" ID-GBIF-Taxon="120817091" ID-Zenodo-Dep="6086416" LSID="urn:lsid:plazi:treatment:038747324C62C6401EB728FCFB6F2681" httpUri="http://treatment.plazi.org/id/038747324C62C6401EB728FCFB6F2681" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
|
||||
<treatment id="038747324C62C6401EB728FCFB6F2681" ID-DOI="http://doi.org/10.5281/zenodo.6086416" ID-GBIF-Taxon="120817091" ID-Zenodo-Dep="6086416" LSID="urn:lsid:plazi:treatment:038747324C62C6401EB728FCFB6F2681" httpUri="http://treatment.plazi.org/id/038747324C62C6401EB728FCFB6F2681" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C62C6431EB728FCFC2E249F" box="[151,1021,1489,1515]" pageId="20" pageNumber="21" type="nomenclature">
|
||||
<paragraph id="8B91F6244C62C6431EB728FCFC2E249F" blockId="20.[151,1021,1489,1515]" box="[151,1021,1489,1515]" pageId="20" pageNumber="21">
|
||||
<heading id="D0D941484C62C6431EB728FCFC2E249F" bold="true" box="[151,1021,1489,1515]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
|
||||
<emphasis id="B95A2A364C62C6431EB728FCFC2E249F" bold="true" box="[151,1021,1489,1515]" pageId="20" pageNumber="21">
|
||||
<taxonomicName id="4C2E8DA74C62C6431EB728FCFE51249F" box="[151,386,1489,1515]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C62C6431EB728FCFE51249F" ID-CoL="3LCPQ" box="[151,386,1489,1515]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C62C6431EB728FCFE51249F" bold="true" box="[151,386,1489,1515]" italics="true" pageId="20" pageNumber="21">Heteropsis drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group (including subgenus
|
||||
|
|
@ -57,11 +57,11 @@ group (including subgenus
|
|||
<subSubSection id="C334A5AF4C62C6421EB72B35FBE9241B" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="reference_group">
|
||||
<paragraph id="8B91F6244C62C6431EB72B35FA4F2694" blockId="20.[151,1436,1560,2017]" pageId="20" pageNumber="21">
|
||||
The
|
||||
<taxonomicName id="4C2E8DA74C62C6431EEE2B34FE892745" box="[206,346,1560,1585]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C62C6431EEE2B34FE892745" box="[206,346,1560,1585]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C62C6431EEE2B34FE892745" box="[206,346,1560,1585]" italics="true" pageId="20" pageNumber="21">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group (comprising the ‘
|
||||
<taxonomicName id="4C2E8DA74C62C6431C5E2B34FD1F2744" box="[638,716,1561,1584]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="narova">
|
||||
<taxonomicName id="4C2E8DA74C62C6431C5E2B34FD1F2744" box="[638,716,1561,1584]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="narova">
|
||||
<emphasis id="B95A2A364C62C6431C5E2B34FD1F2744" box="[638,716,1561,1584]" italics="true" pageId="20" pageNumber="21">narova</emphasis>
|
||||
</taxonomicName>
|
||||
-group’ of
|
||||
|
|
@ -82,11 +82,11 @@ Lees
|
|||
<emphasis id="B95A2A364C62C6431D642B10FC652720" box="[836,950,1597,1620]" italics="true" pageId="20" pageNumber="21">Henotesia</emphasis>
|
||||
</taxonomicName>
|
||||
clade’, with the addition of
|
||||
<taxonomicName id="4C2E8DA74C62C6431ACF2B10FABD2721" box="[1263,1390,1596,1621]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C62C6431ACF2B10FABD2721" box="[1263,1390,1596,1621]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C62C6431ACF2B10FABD2721" box="[1263,1390,1596,1621]" italics="true" pageId="20" pageNumber="21">H. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C62C6431EB72B4CFEE1270D" box="[151,306,1632,1657]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fuliginosa">
|
||||
<taxonomicName id="4C2E8DA74C62C6431EB72B4CFEE1270D" box="[151,306,1632,1657]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fuliginosa">
|
||||
<emphasis id="B95A2A364C62C6431EB72B4CFEE1270D" box="[151,306,1632,1657]" italics="true" pageId="20" pageNumber="21">Ht. fuliginosa</emphasis>
|
||||
</taxonomicName>
|
||||
; see Aduse-Poku
|
||||
|
|
@ -129,7 +129,7 @@ with it, is based on
|
|||
(Mabille, 1877)
|
||||
</taxonomicName>
|
||||
. The
|
||||
<taxonomicName id="4C2E8DA74C62C6431FA72A38FDDE2659" box="[391,525,1812,1837]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C62C6431FA72A38FDDE2659" box="[391,525,1812,1837]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C62C6431FA72A38FDDE2659" box="[391,525,1812,1837]" italics="true" pageId="20" pageNumber="21">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group, or at least the
|
||||
|
|
@ -151,7 +151,7 @@ Kodandaramaiah
|
|||
clade is the dorso-ventrally flattened aedeagus (
|
||||
<bibRefCitation id="EFBF8BD54C62C6431D732A70FC002601" author="Lees" box="[851,979,1885,1909]" pageId="20" pageNumber="21" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
). However, the entire
|
||||
<taxonomicName id="4C2E8DA74C62C6431AEC2A70FA802601" box="[1228,1363,1884,1909]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C62C6431AEC2A70FA802601" box="[1228,1363,1884,1909]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C62C6431AEC2A70FA802601" box="[1228,1363,1884,1909]" italics="true" pageId="20" pageNumber="21">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group includes moderately to highly sexually dimorphic species (
|
||||
|
|
@ -163,18 +163,18 @@ group includes moderately to highly sexually dimorphic species (
|
|||
<emphasis id="B95A2A364C62C6431D142A88FC7626C8" box="[820,933,1957,1980]" italics="true" pageId="20" pageNumber="21">Henotesia</emphasis>
|
||||
</taxonomicName>
|
||||
clade, had 92% bootstrap support as sister to
|
||||
<taxonomicName id="4C2E8DA74C62C6431EB72AE4FEF32695" box="[151,288,1992,2017]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C62C6431EB72AE4FEF32695" box="[151,288,1992,2017]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C62C6431EB72AE4FEF32695" box="[151,288,1992,2017]" italics="true" pageId="20" pageNumber="21">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
, and this clade 87% support as sister to the hitherto hard to place
|
||||
<taxonomicName id="4C2E8DA74C62C6431A0E2AE4FB182695" box="[1070,1227,1992,2017]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fuliginosa">
|
||||
<taxonomicName id="4C2E8DA74C62C6431A0E2AE4FB182695" box="[1070,1227,1992,2017]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fuliginosa">
|
||||
<emphasis id="B95A2A364C62C6431A0E2AE4FB182695" box="[1070,1227,1992,2017]" italics="true" pageId="20" pageNumber="21">Ht. fuliginosa</emphasis>
|
||||
</taxonomicName>
|
||||
. The sister of
|
||||
<emphasis id="B95A2A364C62C6431B592AE4FA4F2694" box="[1401,1436,1993,2016]" italics="true" pageId="20" pageNumber="21">Ht.</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C63C6421EB72DBAFEBC2384" blockId="21.[151,1437,151,2030]" pageId="21" pageNumber="22">
|
||||
<taxonomicName id="4C2E8DA74C63C6421EB72DBAFF2721C4" box="[151,244,151,176]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C63C6421EB72DBAFF2721C4" box="[151,244,151,176]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C63C6421EB72DBAFF2721C4" box="[151,244,151,176]" italics="true" pageId="21" pageNumber="22">drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group is recovered with significant bootstrap support (
|
||||
|
|
@ -189,11 +189,11 @@ Aduse-Poku
|
|||
</taxonomicName>
|
||||
group’ of
|
||||
<collectingCountry id="F339B6B44C63C6421F422D91FE1C21A1" box="[354,463,188,213]" name="Madagascar" pageId="21" pageNumber="22">Malagasy</collectingCountry>
|
||||
<taxonomicName id="4C2E8DA74C63C6421FFA2D90FD8121A0" ID-CoL="8HKH8" box="[474,594,189,212]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C63C6421FFA2D90FD8121A0" ID-CoL="8HKH8" box="[474,594,189,212]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C63C6421FFA2D90FD8121A0" box="[474,594,189,212]" italics="true" pageId="21" pageNumber="22">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
that included
|
||||
<taxonomicName id="4C2E8DA74C63C6421CDE2D90FC8F21A1" box="[766,860,188,213]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="vola">
|
||||
<taxonomicName id="4C2E8DA74C63C6421CDE2D90FC8F21A1" box="[766,860,188,213]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="vola">
|
||||
<emphasis id="B95A2A364C63C6421CDE2D90FC8F21A1" box="[766,860,188,213]" italics="true" pageId="21" pageNumber="22">Ht. vola</emphasis>
|
||||
</taxonomicName>
|
||||
, that was previously (and erroneously) placed in (subgenus)
|
||||
|
|
@ -212,18 +212,18 @@ group also comprised
|
|||
(Ward, 1870)
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName id="4C2E8DA74C63C6421B0E2DCCFEEB2068" authority="Butler, 1879" authorityName="Butler" authorityYear="1879" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="parva">
|
||||
<taxonomicName id="4C2E8DA74C63C6421B0E2DCCFEEB2068" authority="Butler, 1879" authorityName="Butler" authorityYear="1879" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="parva">
|
||||
<emphasis id="B95A2A364C63C6421B0E2DCCFA4F218C" box="[1326,1436,225,248]" italics="true" pageId="21" pageNumber="22">Ht. parva</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C63C6421EBE2C29FEE32068" author="Butler" box="[158,304,260,285]" pageId="21" pageNumber="22" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and members of the
|
||||
<taxonomicName id="4C2E8DA74C63C6421C172C28FD6E2069" box="[567,701,260,285]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="avelona">
|
||||
<taxonomicName id="4C2E8DA74C63C6421C172C28FD6E2069" box="[567,701,260,285]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="avelona">
|
||||
<emphasis id="B95A2A364C63C6421C172C28FD6E2069" box="[567,701,260,285]" italics="true" pageId="21" pageNumber="22">Ht. avelona</emphasis>
|
||||
</taxonomicName>
|
||||
group (=
|
||||
<taxonomicName id="4C2E8DA74C63C6421D0D2C29FC4A2069" box="[813,921,260,285]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bicristata">
|
||||
<taxonomicName id="4C2E8DA74C63C6421D0D2C29FC4A2069" box="[813,921,260,285]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bicristata">
|
||||
<emphasis id="B95A2A364C63C6421D0D2C29FC4A2069" box="[813,921,260,285]" italics="true" pageId="21" pageNumber="22">bicristata</emphasis>
|
||||
</taxonomicName>
|
||||
group of
|
||||
|
|
@ -233,24 +233,24 @@ Torres
|
|||
, 2001
|
||||
</bibRefCitation>
|
||||
), including
|
||||
<taxonomicName id="4C2E8DA74C63C6421B592C28FE582034" authority="Ward, 1870" authorityName="Ward" authorityYear="1870" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="avelona">
|
||||
<taxonomicName id="4C2E8DA74C63C6421B592C28FE582034" authority="Ward, 1870" authorityName="Ward" authorityYear="1870" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="avelona">
|
||||
<emphasis id="B95A2A364C63C6421B592C28FF232034" italics="true" pageId="21" pageNumber="22">Ht. avelona</emphasis>
|
||||
(Ward, 1870)
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName id="4C2E8DA74C63C6421F882C04FCD12034" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[424,770,295,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="uniformis">
|
||||
<taxonomicName id="4C2E8DA74C63C6421F882C04FCD12034" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[424,770,295,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="uniformis">
|
||||
<emphasis id="B95A2A364C63C6421F882C04FDEE2034" box="[424,573,296,320]" italics="true" pageId="21" pageNumber="22">Ht. uniformis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C63C6421C6B2C0AFD292034" author="Oberthur" box="[587,762,295,320]" pageId="21" pageNumber="22" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C63C6421D2D2C04FB8D2034" authority="Mabille, 1880" authorityName="Mabille" authorityYear="1880" box="[781,1118,295,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="parvidens">
|
||||
<taxonomicName id="4C2E8DA74C63C6421D2D2C04FB8D2034" authority="Mabille, 1880" authorityName="Mabille" authorityYear="1880" box="[781,1118,295,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="parvidens">
|
||||
<emphasis id="B95A2A364C63C6421D2D2C04FC762034" box="[781,933,295,320]" italics="true" pageId="21" pageNumber="22">Ht. parvidens</emphasis>
|
||||
(Mabille, 1880)
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C2E8DA74C63C6421AB92C04FF082010" authority="Mabille, 1878" authorityName="Mabille" authorityYear="1878" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bicristata">
|
||||
<taxonomicName id="4C2E8DA74C63C6421AB92C04FF082010" authority="Mabille, 1878" authorityName="Mabille" authorityYear="1878" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bicristata">
|
||||
<emphasis id="B95A2A364C63C6421AB92C04FAFD2034" box="[1177,1326,295,320]" italics="true" pageId="21" pageNumber="22">Ht. bicristata</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C63C6421B1C2C0AFF002010" author="Mabille" pageId="21" pageNumber="22" refString="Mabille, P. (1878) Lepidoptera Africana. Bulletin de la Societe Zoologique de France, 3, 81 - 95." type="journal article" year="1878">Mabille, 1878</bibRefCitation>
|
||||
|
|
@ -261,13 +261,13 @@ Torres
|
|||
<emphasis id="B95A2A364C63C6421C962C60FCF42011" box="[694,807,332,357]" italics="true" pageId="21" pageNumber="22">Ht. iboina</emphasis>
|
||||
</taxonomicName>
|
||||
group. In general, the question of the number and limits of subgenera in
|
||||
<taxonomicName id="4C2E8DA74C63C6421F702C5CFE1B20FC" ID-CoL="8HKH8" box="[336,456,369,392]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C63C6421F702C5CFE1B20FC" ID-CoL="8HKH8" box="[336,456,369,392]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C63C6421F702C5CFE1B20FC" box="[336,456,369,392]" italics="true" pageId="21" pageNumber="22">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
is left open here. If the name
|
||||
<emphasis id="B95A2A364C63C6421D052C42FC7320FC" box="[805,928,367,392]" italics="true" pageId="21" pageNumber="22">Houlbertia</emphasis>
|
||||
Oberthür was restricted to a clade of highly sexually dimorphic species that include
|
||||
<taxonomicName id="4C2E8DA74C63C6421C492CB8FD3720D8" box="[617,740,405,428]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="narova">
|
||||
<taxonomicName id="4C2E8DA74C63C6421C492CB8FD3720D8" box="[617,740,405,428]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="narova">
|
||||
<emphasis id="B95A2A364C63C6421C492CB8FD3720D8" box="[617,740,405,428]" italics="true" pageId="21" pageNumber="22">Ht. narova</emphasis>
|
||||
</taxonomicName>
|
||||
, excluding
|
||||
|
|
@ -277,7 +277,7 @@ Oberthür was restricted to a clade of highly sexually dimorphic species that in
|
|||
according to Aduse-Poku
|
||||
<emphasis id="B95A2A364C63C6421B792CB8FA4F20D9" box="[1369,1436,404,429]" italics="true" pageId="21" pageNumber="22">et al.,</emphasis>
|
||||
(2015,
|
||||
<figureCitation id="1315EAA14C63C6421EC72C9AFEFA20A4" box="[231,297,439,464]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C63C6421EC72C9AFEFA20A4" box="[231,297,439,464]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
|
||||
), with
|
||||
<taxonomicName id="4C2E8DA74C63C6421F5D2C94FDEE20A4" ID-CoL="92BDD" authority="Butler" authorityName="Butler" box="[381,573,439,464]" class="Insecta" family="Nymphalidae" genus="Henotesia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C63C6421F5D2C94FE3C20A4" box="[381,495,441,464]" italics="true" pageId="21" pageNumber="22">Henotesia</emphasis>
|
||||
|
|
@ -296,18 +296,18 @@ without a subgeneric name. While more faithful to traditional taxonomy in
|
|||
and
|
||||
<emphasis id="B95A2A364C63C6421DB22CD2FBDE236C" box="[914,1037,511,536]" italics="true" pageId="21" pageNumber="22">Houlbertia</emphasis>
|
||||
with the nominotypical subgenus
|
||||
<taxonomicName id="4C2E8DA74C63C6421EB72F08FE422349" ID-CoL="8HKH8" authority="Westwood" authorityName="Westwood" box="[151,401,548,573]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C63C6421EB72F08FE422349" ID-CoL="8HKH8" authority="Westwood" authorityName="Westwood" box="[151,401,548,573]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C63C6421EB72F08FEDC2348" box="[151,271,549,572]" italics="true" pageId="21" pageNumber="22">Heteropsis</emphasis>
|
||||
Westwood
|
||||
</taxonomicName>
|
||||
, perhaps with the addition of the previously enigmatic
|
||||
<taxonomicName id="4C2E8DA74C63C6421A272F08FB702349" box="[1031,1187,548,573]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fuliginosa">
|
||||
<taxonomicName id="4C2E8DA74C63C6421A272F08FB702349" box="[1031,1187,548,573]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fuliginosa">
|
||||
<emphasis id="B95A2A364C63C6421A272F08FB702349" box="[1031,1187,548,573]" italics="true" pageId="21" pageNumber="22">Ht. fuliginosa</emphasis>
|
||||
</taxonomicName>
|
||||
, forming together the well-supported clade in Aduse-Poku
|
||||
<emphasis id="B95A2A364C63C6421C192F64FDAA2314" box="[569,633,583,608]" italics="true" pageId="21" pageNumber="22">et al.,</emphasis>
|
||||
(2015,
|
||||
<figureCitation id="1315EAA14C63C6421CF12F6AFCC12314" box="[721,786,583,608]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C63C6421CF12F6AFCC12314" box="[721,786,583,608]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
|
||||
). However, such an action, that would abandon names in prior use as historic curiosities, awaits a forthcoming study. It should also be noted that without explicit justification (perhaps merely because it seemed different to other African hairy-eyed mycalesines),
|
||||
<bibRefCitation id="EFBF8BD54C63C6421B172FA2FF0F23B8" author="d'Abrera" pageId="21" pageNumber="22" refString="d'Abrera, B. L. (1997) Butterflies of the Afrotropical Region. Part 1. Papilionidae, Pieridae, Acraeidae, Danaidae, Satyridae. Hill House Publishers, Melbourne & London, 263 pp." type="book" year="1997">d’Abrera (1997</bibRefCitation>
|
||||
: 226) included the African species ‘
|
||||
|
|
@ -324,7 +324,7 @@ Aduse-Poku
|
|||
</paragraph>
|
||||
<paragraph id="8B91F6244C63C6421EE72FD1FBE9241B" blockId="21.[151,1437,151,2030]" pageId="21" pageNumber="22">
|
||||
Species of the
|
||||
<taxonomicName id="4C2E8DA74C63C6421F4C2FD0FE202261" box="[364,499,764,789]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C63C6421F4C2FD0FE202261" box="[364,499,764,789]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C63C6421F4C2FD0FE202261" box="[364,499,764,789]" italics="true" pageId="21" pageNumber="22">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group with the most vivid blue upperside ♂♂ such as
|
||||
|
|
@ -334,7 +334,7 @@ group with the most vivid blue upperside ♂♂ such as
|
|||
occur at lower elevations than those with dark ♂♂, further discussed below. The last group tend to be found in more montane areas, up to over
|
||||
<quantity id="4CD65BC14C63C6421F762E6EFE7E222F" box="[342,429,835,859]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="22" unit="m" value="2300.0">2300 m</quantity>
|
||||
, in the case of
|
||||
<taxonomicName id="4C2E8DA74C63C6421C742E69FD092228" box="[596,730,835,860]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<taxonomicName id="4C2E8DA74C63C6421C742E69FD092228" box="[596,730,835,860]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<emphasis id="B95A2A364C63C6421C742E69FD092228" box="[596,730,835,860]" italics="true" pageId="21" pageNumber="22">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
, which is as high as any mycalesines are found in
|
||||
|
|
@ -358,7 +358,7 @@ Endl.
|
|||
), whereas the opposite sex is regularly attracted to fruit. Adults sometimes feed on
|
||||
<emphasis id="B95A2A364C63C6421CDE2E83FC9622B1" box="[766,837,942,965]" italics="true" pageId="21" pageNumber="22">in situ</emphasis>
|
||||
fruits. Species fly from understory to lower canopy, males following lightgaps, sometimes resting on top of clumps of bamboo, whereas
|
||||
<taxonomicName id="4C2E8DA74C63C6421A6C2EFFFB00229D" box="[1100,1235,976,1001]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C63C6421A6C2EFFFB00229D" box="[1100,1235,976,1001]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C63C6421A6C2EFFFB00229D" box="[1100,1235,976,1001]" italics="true" pageId="21" pageNumber="22">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
males hilltop and rest on top of ridge bushes in sunny weather. Many if not all species seems to be closely associated with different
|
||||
|
|
@ -369,7 +369,7 @@ of forest bamboo, but early stage data is very sparse. Stands of bamboo (such as
|
|||
S.Dransf.
|
||||
</taxonomicName>
|
||||
in the lowlands) can sometimes support large sympatrically flying populations of several closely related species. ♀♀ of the more sexually dimorphic members of the
|
||||
<taxonomicName id="4C2E8DA74C63C6421D6F2972FC052503" box="[847,982,1118,1143]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C63C6421D6F2972FC052503" box="[847,982,1118,1143]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C63C6421D6F2972FC052503" box="[847,982,1118,1143]" italics="true" pageId="21" pageNumber="22">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group tend to be very sedentary resting low in the undergrowth on leaf litter (where they are cryptic with fallen leaves including those of bamboo), and may not be noticed at all when not venturing off paths. The ♀♀ of
|
||||
|
|
@ -414,7 +414,7 @@ along with
|
|||
; see
|
||||
<bibRefCitation id="EFBF8BD54C63C6421EB72819FEC42438" author="Lees" box="[151,279,1332,1356]" pageId="21" pageNumber="22" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
). The more montane members of the
|
||||
<taxonomicName id="4C2E8DA74C63C6421C9F2819FC942438" box="[703,839,1331,1356]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C63C6421C9F2819FC942438" box="[703,839,1331,1356]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C63C6421C9F2819FC942438" box="[703,839,1331,1356]" italics="true" pageId="21" pageNumber="22">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group have ♀♀ that tend to be vertical trunk-resting (as do often the ♂♂, likely explaining their less sexually dimorphic wing patterns).
|
||||
|
|
@ -423,7 +423,7 @@ group have ♀♀ that tend to be vertical trunk-resting (as do often the ♂♂
|
|||
<subSubSection id="C334A5AF4C63C6421EE72857FCFF26BE" pageId="21" pageNumber="22" type="description">
|
||||
<paragraph id="8B91F6244C63C6421EE72857FCFF26BE" blockId="21.[151,1437,151,2030]" pageId="21" pageNumber="22">
|
||||
Within the
|
||||
<taxonomicName id="4C2E8DA74C63C6421F662856FE1D24E7" box="[326,462,1402,1427]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C63C6421F662856FE1D24E7" box="[326,462,1402,1427]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C63C6421F662856FE1D24E7" box="[326,462,1402,1427]" italics="true" pageId="21" pageNumber="22">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group are two or more groups of phenetically similar, moderately sexually dimorphic species, which inhabit montane forest ridges between
|
||||
|
|
@ -431,9 +431,9 @@ group are two or more groups of phenetically similar, moderately sexually dimorp
|
|||
. Two sampled members in the phylogeny of Aduse-Poku
|
||||
<emphasis id="B95A2A364C63C6421F0A28EFFEB924AE" box="[298,362,1473,1498]" italics="true" pageId="21" pageNumber="22">et al.,</emphasis>
|
||||
(2015,
|
||||
<figureCitation id="1315EAA14C63C6421FE328ECFDD724AE" box="[451,516,1473,1498]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C63C6421FE328ECFDD724AE" box="[451,516,1473,1498]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
|
||||
) are
|
||||
<taxonomicName id="4C2E8DA74C63C6421C6228EFFCE524AE" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[578,822,1474,1498]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C63C6421C6228EFFCE524AE" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[578,822,1474,1498]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C63C6421C6228EFFD6124AD" box="[578,690,1474,1497]" italics="true" pageId="21" pageNumber="22">Ht. viettei</emphasis>
|
||||
Lees, 2003
|
||||
</taxonomicName>
|
||||
|
|
@ -444,7 +444,7 @@ and
|
|||
<emphasis id="B95A2A364C63C6421D8428ECFBEA24AE" box="[932,1081,1473,1498]" italics="true" pageId="21" pageNumber="22">andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
’ (=
|
||||
<taxonomicName id="4C2E8DA74C63C6421A4428EFFAD624AE" box="[1124,1285,1473,1498]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C63C6421A4428EFFAD624AE" box="[1124,1285,1473,1498]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="westwoodi" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C63C6421A4428EFFAD624AE" box="[1124,1285,1473,1498]" italics="true" pageId="21" pageNumber="22">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C63C6421B2E28ECFABA24AE" box="[1294,1385,1473,1498]" pageId="21" pageNumber="22" rank="species">
|
||||
|
|
@ -455,7 +455,7 @@ and
|
|||
<emphasis id="B95A2A364C63C6421FC928CBFDA32489" box="[489,624,1508,1533]" italics="true" pageId="21" pageNumber="22">Ht. obscura</emphasis>
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName id="4C2E8DA74C63C6421CB428CBFC0A2489" authority="Mabille, 1880" authorityName="Mabille" authorityYear="1880" box="[660,985,1508,1533]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="difficilis">
|
||||
<taxonomicName id="4C2E8DA74C63C6421CB428CBFC0A2489" authority="Mabille, 1880" authorityName="Mabille" authorityYear="1880" box="[660,985,1508,1533]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="difficilis">
|
||||
<emphasis id="B95A2A364C63C6421CB428CBFCCF2489" box="[660,796,1508,1533]" italics="true" pageId="21" pageNumber="22">Ht. difficilis</emphasis>
|
||||
(Mabille, 1880)
|
||||
</taxonomicName>
|
||||
|
|
@ -475,22 +475,22 @@ sp. 13’
|
|||
: 417; now including sp. 74), and sp. 13B, respectively, of
|
||||
<bibRefCitation id="EFBF8BD54C63C6421AC52B01FA4A2730" author="Lees" box="[1253,1433,1579,1604]" pageId="21" pageNumber="22" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997: 64)</bibRefCitation>
|
||||
. This species pair exhibit varying degrees of whitish or cream highlighting distal of the HWV Mb. The
|
||||
<taxonomicName id="4C2E8DA74C63C6421B3D2B7DFA4F271C" box="[1309,1436,1615,1640]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C63C6421B3D2B7DFA4F271C" box="[1309,1436,1615,1640]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C63C6421B3D2B7DFA4F271C" box="[1309,1436,1615,1640]" italics="true" pageId="21" pageNumber="22">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
sub-group of the
|
||||
<taxonomicName id="4C2E8DA74C63C6421F752B59FE0827FF" box="[341,475,1650,1675]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C63C6421F752B59FE0827FF" box="[341,475,1650,1675]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C63C6421F752B59FE0827FF" box="[341,475,1650,1675]" italics="true" pageId="21" pageNumber="22">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group represents a tightly knit species complex with dark-uppersides and includes
|
||||
<taxonomicName id="4C2E8DA74C63C6421B592B59FF3827DB" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="harveyi" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C63C6421B592B59FF3827DB" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="harveyi" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C63C6421B592B59FF3827DB" italics="true" pageId="21" pageNumber="22">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C63C6421ED22BBBFE9F27DB" box="[242,332,1686,1711]" pageId="21" pageNumber="22" rank="species">
|
||||
<emphasis id="B95A2A364C63C6421ED22BBBFE9F27DB" bold="true" box="[242,332,1686,1711]" pageId="21" pageNumber="22">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C63C6421FA42BBAFDE727DB" box="[388,564,1686,1711]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="vanewrighti" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C63C6421FA42BBAFDE727DB" box="[388,564,1686,1711]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="vanewrighti" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C63C6421FA42BBAFDE727DB" box="[388,564,1686,1711]" italics="true" pageId="21" pageNumber="22">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C63C6421C1B2BBBFD4627DB" box="[571,661,1686,1711]" pageId="21" pageNumber="22" rank="species">
|
||||
|
|
@ -505,27 +505,27 @@ Torres
|
|||
, 2001
|
||||
</bibRefCitation>
|
||||
). The
|
||||
<taxonomicName id="4C2E8DA74C63C6421A672B97FB1627A6" box="[1095,1221,1721,1746]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C63C6421A672B97FB1627A6" box="[1095,1221,1721,1746]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C63C6421A672B97FB1627A6" box="[1095,1221,1721,1746]" italics="true" pageId="21" pageNumber="22">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
sub-group exhibits the presence or absence of a russet or dark patch on the underside of the FW along vein 1A+2A (in
|
||||
<collectingCountry id="F339B6B44C63C6421B2F2BF1FA4B2781" box="[1295,1432,1756,1781]" name="Madagascar" pageId="21" pageNumber="22">Madagascar</collectingCountry>
|
||||
, only the
|
||||
<taxonomicName id="4C2E8DA74C63C6421EDA2A2CFEAE266D" box="[250,381,1792,1817]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="avelona">
|
||||
<taxonomicName id="4C2E8DA74C63C6421EDA2A2CFEAE266D" box="[250,381,1792,1817]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="avelona">
|
||||
<emphasis id="B95A2A364C63C6421EDA2A2CFEAE266D" box="[250,381,1792,1817]" italics="true" pageId="21" pageNumber="22">Ht. avelona</emphasis>
|
||||
</taxonomicName>
|
||||
group of mycalesines also exhibit a FWV patch). Another dark non-reflective upperside pair, more similar to
|
||||
<taxonomicName id="4C2E8DA74C63C6421F762A08FE1A2648" box="[342,457,1829,1852]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C63C6421F762A08FE1A2648" box="[342,457,1829,1852]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C63C6421F762A08FE1A2648" box="[342,457,1829,1852]" italics="true" pageId="21" pageNumber="22">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName id="4C2E8DA74C63C6421FCF2A08FD472649" box="[495,660,1828,1853]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<taxonomicName id="4C2E8DA74C63C6421FCF2A08FD472649" box="[495,660,1828,1853]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<emphasis id="B95A2A364C63C6421FCF2A08FD472649" box="[495,660,1828,1853]" italics="true" pageId="21" pageNumber="22">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
, although not represented in the phylogeny of Aduse-Poku
|
||||
<emphasis id="B95A2A364C63C6421B782A08FA4F2649" box="[1368,1436,1828,1853]" italics="true" pageId="21" pageNumber="22">et al.,</emphasis>
|
||||
(2015), and so with relationship unresolved, is
|
||||
<taxonomicName id="4C2E8DA74C63C6421C852A65FCF5262B" box="[677,806,1864,1887]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="imerina" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C63C6421C852A65FCF5262B" box="[677,806,1864,1887]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="imerina" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C63C6421C852A65FCF5262B" box="[677,806,1864,1887]" italics="true" pageId="21" pageNumber="22">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C63C6421D0D2A6AFC542614" box="[813,903,1863,1888]" pageId="21" pageNumber="22" rank="species">
|
||||
|
|
@ -540,7 +540,7 @@ Torres
|
|||
, 2001
|
||||
</bibRefCitation>
|
||||
) and
|
||||
<taxonomicName id="4C2E8DA74C63C6421F1B2A41FE1126F0" box="[315,450,1899,1924]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<taxonomicName id="4C2E8DA74C63C6421F1B2A41FE1126F0" box="[315,450,1899,1924]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pauliani">
|
||||
<emphasis id="B95A2A364C63C6421F1B2A41FE1126F0" box="[315,450,1899,1924]" italics="true" pageId="21" pageNumber="22">Ht. pauliani</emphasis>
|
||||
</taxonomicName>
|
||||
(sp. 37 of
|
||||
|
|
@ -575,13 +575,13 @@ locality, the first and main part of the short description matches at least the
|
|||
: Pl. 5, Fig. 6.7). However, this first part does not match the ♂ of the taxon represented by the “var.” in the latter work and illustrated in
|
||||
<bibRefCitation id="EFBF8BD54C60C6411CEA2C29FCAB2069" author="Mabille" box="[714,888,260,285]" pageId="22" pageNumber="23" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter" year="1885">Mabille [1885]</bibRefCitation>
|
||||
: Pl. 5,
|
||||
<figureCitation id="1315EAA14C60C6411DEA2C29FBDC2068" box="[970,1039,260,285]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="22" pageNumber="23">Fig. 8</figureCitation>
|
||||
<figureCitation id="1315EAA14C60C6411DEA2C29FBDC2068" box="[970,1039,260,285]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264605" httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="22" pageNumber="23">Fig. 8</figureCitation>
|
||||
). The additional details in
|
||||
<bibRefCitation id="EFBF8BD54C60C6411B652C29FF362034" author="Mabille" pageId="22" pageNumber="23" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter" year="1885">Mabille ([1885</bibRefCitation>
|
||||
-7]) suggests that the original description had been based on at least three phenotypically divergent specimens, two ♂ and one ♀ (that illustrated dorsally by
|
||||
<bibRefCitation id="EFBF8BD54C60C6411D372C61FC192011" box="[791,970,332,357]" pageId="22" pageNumber="23" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter">Mabille, [1885]</bibRefCitation>
|
||||
: Pl. 5,
|
||||
<figureCitation id="1315EAA14C60C6411A372C61FB892010" box="[1047,1114,332,357]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
|
||||
<figureCitation id="1315EAA14C60C6411A372C61FB892010" box="[1047,1114,332,357]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C60C6411EE72C42FC952229" blockId="22.[151,1437,151,2016]" pageId="22" pageNumber="23">
|
||||
|
|
@ -596,7 +596,7 @@ of
|
|||
<emphasis id="B95A2A364C60C6411CD32C9AFBD020A4" box="[755,1027,439,464]" italics="true" pageId="22" pageNumber="23">Mycalesis andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
had been traced in the three main feasible repositories for Mabille material, BMNH and MNHN (DCL, pers. obs.), or ZHMU (W. Mey, pers. comm.). Although the ♂ (
|
||||
<figureCitation id="1315EAA14C60C6411F412CD2FE6D236C" box="[353,446,511,536]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="22" pageNumber="23">Fig. 6A</figureCitation>
|
||||
<figureCitation id="1315EAA14C60C6411F412CD2FE6D236C" box="[353,446,511,536]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="22" pageNumber="23">Fig. 6A</figureCitation>
|
||||
, ‘6, 7’) might have been a good choice for the
|
||||
<typeStatus id="549548864C60C6411DD82CD2FB8C236C" box="[1016,1119,511,536]" pageId="22" pageNumber="23" type="lectotype">lectotype</typeStatus>
|
||||
of
|
||||
|
|
@ -621,7 +621,7 @@ identity and is now numbered as BMNH(E) #674841. Because of its wide and distinc
|
|||
for ‘
|
||||
<emphasis id="B95A2A364C60C6411DAB2FD0FC4C2260" box="[907,927,765,788]" italics="true" pageId="22" pageNumber="23">H</emphasis>
|
||||
. 13B’), to correspond to ‘sp. 13B’ (described below as
|
||||
<taxonomicName id="4C2E8DA74C60C6411F2E2E0CFE66224C" box="[270,437,799,824]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<taxonomicName id="4C2E8DA74C60C6411F2E2E0CFE66224C" box="[270,437,799,824]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<emphasis id="B95A2A364C60C6411F2E2E0CFE66224C" box="[270,437,799,824]" italics="true" pageId="22" pageNumber="23">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
, a species apparently endemic to the southeastern mountains of
|
||||
|
|
@ -639,7 +639,7 @@ Mabille, 1880
|
|||
. This last taxon is illustrated in
|
||||
<bibRefCitation id="EFBF8BD54C60C6411EB72ED7FE802567" box="[151,339,1018,1043]" pageId="22" pageNumber="23" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter">Mabille, ([1885]</bibRefCitation>
|
||||
, Pl. 7B, ‘2’, ‘2a’) and reproduced here (
|
||||
<figureCitation id="1315EAA14C60C6411D332ED7FCBE2567" box="[787,877,1018,1043]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="22" pageNumber="23">Fig. 6A</figureCitation>
|
||||
<figureCitation id="1315EAA14C60C6411D332ED7FCBE2567" box="[787,877,1018,1043]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="22" pageNumber="23">Fig. 6A</figureCitation>
|
||||
, ‘9’).
|
||||
<taxonomicName id="4C2E8DA74C60C6411D932ED1FBE62567" box="[947,1077,1018,1043]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="difficilis">
|
||||
<emphasis id="B95A2A364C60C6411D932ED1FBE62567" box="[947,1077,1018,1043]" italics="true" pageId="22" pageNumber="23">M. difficilis</emphasis>
|
||||
|
|
@ -673,7 +673,7 @@ var.
|
|||
). Although it is has not previously been clear what species is represented by the ♀ whose dorsal surface only is illustrated in
|
||||
<bibRefCitation id="EFBF8BD54C60C6411DD92983FB6125B3" box="[1017,1202,1198,1223]" pageId="22" pageNumber="23" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter">Mabille ([1885]</bibRefCitation>
|
||||
: Pl. 5,
|
||||
<figureCitation id="1315EAA14C60C6411B282983FA9D25B3" box="[1288,1358,1198,1223]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
|
||||
<figureCitation id="1315EAA14C60C6411B282983FA9D25B3" box="[1288,1358,1198,1223]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
|
||||
) as an example of
|
||||
<taxonomicName id="4C2E8DA74C60C6411F3C29FFFDFB259F" box="[284,552,1234,1259]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andravahana">
|
||||
<emphasis id="B95A2A364C60C6411F3C29FFFDFB259F" box="[284,552,1234,1259]" italics="true" pageId="22" pageNumber="23">Mycalesis andravahana</emphasis>
|
||||
|
|
@ -683,7 +683,7 @@ var.
|
|||
<emphasis id="B95A2A364C60C6411D4A29D5FC23247B" box="[874,1008,1270,1295]" italics="true" pageId="22" pageNumber="23">M. difficilis</emphasis>
|
||||
</taxonomicName>
|
||||
. In fact the resemblance to a recent photograph of a living ♀ from Anjanaharibe Sud is striking (
|
||||
<figureCitation id="1315EAA14C60C6411D7D2837FC702447" box="[861,931,1306,1331]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="22" pageNumber="23">Fig. 6</figureCitation>
|
||||
<figureCitation id="1315EAA14C60C6411D7D2837FC702447" box="[861,931,1306,1331]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="22" pageNumber="23">Fig. 6</figureCitation>
|
||||
C). About the original ♀ specimen, Mabille ([1887]: 58) stated “La femelle est d'une couleur plus claire que le mâle, surtout sur le disque. L'ocelle des ailes supérieures est très grand, d'un fauve clair; l'ocelle des inférieures est très petit. En dessous, les premières ailes ont les zébrures blanchâtres jusque dans la cellule; l'espace terminal est d'un gris cendré. Aux inférieures, il y a les mêmes dessins, mais plus délayés; la bandelette blanche est d'un gris sale. Les autres parties de l'insecte sont comme dans le
|
||||
<typeStatus id="549548864C60C6411F7328FDFE50249C" box="[339,387,1488,1512]" pageId="22" pageNumber="23">type</typeStatus>
|
||||
mâle que nous avons décrit plus haut”. Just below this he stated “L'envergure est de 36 millimetres chez le ♂, de 44 chez la ♀.
|
||||
|
|
@ -699,7 +699,7 @@ had the same provenance (Mabille, 1880), but as implied above, would represent a
|
|||
<emphasis id="B95A2A364C60C6411B5D2B11FEFF2703" italics="true" pageId="22" pageNumber="23">M. andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
coming from the Grose-Smith collection (BMNH(E) #674841;
|
||||
<figureCitation id="1315EAA14C60C6411DDD2B73FBEC2703" box="[1021,1087,1630,1655]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="22" pageNumber="23">Fig. 6</figureCitation>
|
||||
<figureCitation id="1315EAA14C60C6411DDD2B73FBEC2703" box="[1021,1087,1630,1655]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="22" pageNumber="23">Fig. 6</figureCitation>
|
||||
D) measures
|
||||
<quantity id="4CD65BC14C60C6411AF32B73FAEB2703" box="[1235,1336,1630,1655]" metricMagnitude="-2" metricUnit="m" metricValue="5.02" pageId="22" pageNumber="23" unit="mm" value="50.2">50.2 mm</quantity>
|
||||
absolute wingspan and between
|
||||
|
|
@ -707,11 +707,11 @@ absolute wingspan and between
|
|||
FWL. However, all four wings are fixed to the body with the FW tergal edges angled in a non-orthogonal orientation, with blobs of red sealing wax. My digital reconstruction of the wings shows that the apex to apex dimension, when reconstructed as
|
||||
<bibRefCitation id="EFBF8BD54C60C6411D2B2BE7FC122797" box="[779,961,1738,1763]" pageId="22" pageNumber="23" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter">Mabille ([1885]</bibRefCitation>
|
||||
: Pl. 5,
|
||||
<figureCitation id="1315EAA14C60C6411A2E2BE7FB822797" box="[1038,1105,1738,1763]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
|
||||
<figureCitation id="1315EAA14C60C6411A2E2BE7FB822797" box="[1038,1105,1738,1763]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
|
||||
) actually would measure
|
||||
<quantity id="4CD65BC14C60C6411B522BE6FF3B2673" metricMagnitude="-2" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="22" pageNumber="23" unit="mm" value="44.5" valueMax="45.0" valueMin="44.0">44– 45 mm</quantity>
|
||||
apex to apex. It should be acknowledged that in general,
|
||||
<taxonomicName id="4C2E8DA74C60C6411DA42BDDFC2F2673" box="[900,1020,1776,1799]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C60C6411DA42BDDFC2F2673" box="[900,1020,1776,1799]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C60C6411DA42BDDFC2F2673" box="[900,1020,1776,1799]" italics="true" pageId="22" pageNumber="23">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
are in general hard to identify from uppersides, more so for a painting of a ♀.
|
||||
|
|
@ -746,7 +746,7 @@ of
|
|||
, Lees
|
||||
<emphasis id="B95A2A364C61C6401C1C2DCCFDAF218C" box="[572,636,223,248]" italics="true" pageId="23" pageNumber="24">et al.,</emphasis>
|
||||
2003) does not disagree with Mabille’s ([1885-7]) redescription in any significant aspect. The dorsal illustration would indeed represent a quite accurate portrayal of the original specimen. There is indication, especially on the non hand-coloured copy of Mabille’s work available on Biodiversity Heritage Library (
|
||||
<figureCitation id="1315EAA14C61C6401FD52C61FDEA2011" box="[501,569,332,357]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6</figureCitation>
|
||||
<figureCitation id="1315EAA14C61C6401FD52C61FDEA2011" box="[501,569,332,357]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6</figureCitation>
|
||||
B), of the pale HWV postmedian area showing through to the dorsal surface, just as it does in BMNH(E) #674841, and as clearly indicated in the illustration of
|
||||
<taxonomicName id="4C2E8DA74C61C6401A6A2C42FF3520DB" authority="in Mabille" authorityName="in Mabille" authorityYear="1885" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="difficilis">
|
||||
<emphasis id="B95A2A364C61C6401A6A2C42FACD20FC" box="[1098,1310,367,392]" italics="true" pageId="23" pageNumber="24">Mycalesis difficilis</emphasis>
|
||||
|
|
@ -754,7 +754,7 @@ in
|
|||
<bibRefCitation id="EFBF8BD54C61C6401B652C42FF3520DB" author="Mabille" pageId="23" pageNumber="24" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter" year="1885">Mabille ([1885</bibRefCitation>
|
||||
</taxonomicName>
|
||||
: Pl. 7B) (
|
||||
<figureCitation id="1315EAA14C61C6401F6D2CBBFE7920DB" box="[333,426,406,431]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6A</figureCitation>
|
||||
<figureCitation id="1315EAA14C61C6401F6D2CBBFE7920DB" box="[333,426,406,431]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6A</figureCitation>
|
||||
). The relative size of dorsal FW and HW ocelli and their rings in these two versions of Mabille’s illustration (Pl. 5, ‘9’) is also convincingly accurate as corresponding.
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C61C6401EE72CF3FE02236F" blockId="23.[151,1436,151,2038]" pageId="23" pageNumber="24">
|
||||
|
|
@ -818,17 +818,17 @@ var.
|
|||
, has dull metallic blue upperside ♂♂, is not known from the Southeast of
|
||||
<collectingCountry id="F339B6B44C61C6401A5F2FD6FADF2260" box="[1151,1292,763,788]" name="Madagascar" pageId="23" pageNumber="24">Madagascar</collectingCountry>
|
||||
). ‘Sp. 12’, as known from Ranomafana, is now described below as
|
||||
<taxonomicName id="4C2E8DA74C61C6401CDA2E0DFCAA2243" box="[762,889,798,823]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C61C6401CDA2E0DFCAA2243" box="[762,889,798,823]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C61C6401CDA2E0DFCAA2243" box="[762,889,798,823]" italics="true" pageId="23" pageNumber="24">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
(a manuscript name of
|
||||
<bibRefCitation id="EFBF8BD54C61C6401AA82E0DFADB2243" author="Lees" box="[1160,1288,799,823]" pageId="23" pageNumber="24" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="4C2E8DA74C61C6401B3D2E0DFA4F2243" box="[1309,1436,798,823]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C61C6401B3D2E0DFA4F2243" box="[1309,1436,798,823]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C61C6401B3D2E0DFA4F2243" box="[1309,1436,798,823]" italics="true" pageId="23" pageNumber="24">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
not only has ♂♂ with very dark brown uppersides, but also lacks the distinctly white postmedial band which as mentioned above is present in
|
||||
<taxonomicName id="4C2E8DA74C61C6401C2B2E4AFD53220A" box="[523,640,871,894]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C61C6401C2B2E4AFD53220A" box="[523,640,871,894]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C61C6401C2B2E4AFD53220A" box="[523,640,871,894]" italics="true" pageId="23" pageNumber="24">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
. It is not certain if the species without HWV whitish postmedian highlighting represented as “var.” of
|
||||
|
|
@ -836,11 +836,11 @@ not only has ♂♂ with very dark brown uppersides, but also lacks the distinct
|
|||
<emphasis id="B95A2A364C61C6401C692EA4FC8A22D6" box="[585,857,905,930]" italics="true" pageId="23" pageNumber="24">Mycalesis andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
in Mabille’s description and redescription (1878; [1885]: Pl. 5,
|
||||
<figureCitation id="1315EAA14C61C6401F1C2E80FE5022B1" box="[316,387,941,966]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="23" pageNumber="24">Fig. 8</figureCitation>
|
||||
<figureCitation id="1315EAA14C61C6401F1C2E80FE5022B1" box="[316,387,941,966]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264605" httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="23" pageNumber="24">Fig. 8</figureCitation>
|
||||
; [1887], text) and reproduced here in
|
||||
<figureCitation id="1315EAA14C61C6401D642E80FC4E22B2" box="[836,925,941,966]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6A</figureCitation>
|
||||
<figureCitation id="1315EAA14C61C6401D642E80FC4E22B2" box="[836,925,941,966]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6A</figureCitation>
|
||||
is referable to
|
||||
<taxonomicName id="4C2E8DA74C61C6401A732E83FB0622B2" box="[1107,1237,941,966]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C61C6401A732E83FB0622B2" box="[1107,1237,941,966]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C61C6401A732E83FB0622B2" box="[1107,1237,941,966]" italics="true" pageId="23" pageNumber="24">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
, but it might be. However,
|
||||
|
|
@ -852,15 +852,15 @@ described a similar ‘var.’ lacking the white postmedian HWV band as ‘
|
|||
ab.
|
||||
<emphasis id="B95A2A364C61C6401F4D2ED8FE382578" box="[365,491,1013,1036]" italics="true" pageId="23" pageNumber="24">marmorata</emphasis>
|
||||
’ (specimen represented in
|
||||
<figureCitation id="1315EAA14C61C6401D1F2EDEFC592578" box="[831,906,1011,1036]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="23" pageNumber="24">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C61C6401D1F2EDEFC592578" box="[831,906,1011,1036]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="23" pageNumber="24">Fig. 7</figureCitation>
|
||||
C). Its HT ♂ (‘TYPUS’, specimen NHRS- TOBI000000050) was examined by DCL in SNHM. This is an unavailable name (clearly infrasubspecific, Art 45.6.2), but the specimen matches well
|
||||
<taxonomicName id="4C2E8DA74C61C6401C762911FD072527" box="[598,724,1082,1107]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C61C6401C762911FD072527" box="[598,724,1082,1107]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C61C6401C762911FD072527" box="[598,724,1082,1107]" italics="true" pageId="23" pageNumber="24">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
, in fact much more clearly than does that illustrated in
|
||||
<bibRefCitation id="EFBF8BD54C61C6401B652917FF3D2503" pageId="23" pageNumber="24" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter">Mabille ([1885]</bibRefCitation>
|
||||
: Pl. 5, 8). As judged by Mabille’s [1885] plate,
|
||||
<taxonomicName id="4C2E8DA74C61C6401D112972FC602503" box="[817,947,1118,1143]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C61C6401D112972FC602503" box="[817,947,1118,1143]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C61C6401D112972FC602503" box="[817,947,1118,1143]" italics="true" pageId="23" pageNumber="24">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
may have been one of the species under Mabille’s eyes when he described
|
||||
|
|
@ -868,19 +868,19 @@ may have been one of the species under Mabille’s eyes when he described
|
|||
<emphasis id="B95A2A364C61C6401C0929AEFD3425EE" box="[553,743,1153,1178]" italics="true" pageId="23" pageNumber="24">M. andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
, even though not the first mentioned, or focal, form of the description. In
|
||||
<taxonomicName id="4C2E8DA74C61C6401F68298BFE1925CA" box="[328,458,1189,1214]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C61C6401F68298BFE1925CA" box="[328,458,1189,1214]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C61C6401F68298BFE1925CA" box="[328,458,1189,1214]" italics="true" pageId="23" pageNumber="24">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
, although the ventral irrorated pattern and general colouration resemble the ventral illustration in
|
||||
<bibRefCitation id="EFBF8BD54C61C6401F1929E5FE222595" box="[313,497,1224,1249]" pageId="23" pageNumber="24" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter">Mabille ([1885]</bibRefCitation>
|
||||
: Pl. 5,
|
||||
<figureCitation id="1315EAA14C61C6401C6529E5FD592595" box="[581,650,1224,1249]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="23" pageNumber="24">Fig. 8</figureCitation>
|
||||
<figureCitation id="1315EAA14C61C6401C6529E5FD592595" box="[581,650,1224,1249]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264605" httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="23" pageNumber="24">Fig. 8</figureCitation>
|
||||
), generally the median and pre-median HWV bands are much more clearly delineated. This is the case in the HT of Aurivillius’ aberration, and in most specimens of
|
||||
<taxonomicName id="4C2E8DA74C61C6401ACA29C0FABA2471" box="[1258,1385,1260,1285]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C61C6401ACA29C0FABA2471" box="[1258,1385,1260,1285]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C61C6401ACA29C0FABA2471" box="[1258,1385,1260,1285]" italics="true" pageId="23" pageNumber="24">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
, but not so for Mabille’s ♂ ‘var.’ (
|
||||
<figureCitation id="1315EAA14C61C6401FC12822FDE8245C" box="[481,571,1295,1320]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6A</figureCitation>
|
||||
<figureCitation id="1315EAA14C61C6401FC12822FDE8245C" box="[481,571,1295,1320]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6A</figureCitation>
|
||||
, ‘8’).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C61C6401EE72819FED5241B" blockId="23.[151,1436,151,2038]" pageId="23" pageNumber="24">
|
||||
|
|
@ -889,7 +889,7 @@ To resolve the
|
|||
<emphasis id="B95A2A364C61C6401F51281EFDAC2438" box="[369,639,1331,1356]" italics="true" pageId="23" pageNumber="24">Mycalesis andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
problem, the following synonymy of
|
||||
<taxonomicName id="4C2E8DA74C61C6401A132819FA9A2438" box="[1075,1353,1331,1356]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andravahana">
|
||||
<taxonomicName id="4C2E8DA74C61C6401A132819FA9A2438" box="[1075,1353,1331,1356]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andravahana">
|
||||
<emphasis id="B95A2A364C61C6401A132819FA9A2438" box="[1075,1353,1331,1356]" italics="true" pageId="23" pageNumber="24">Heteropsis andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
is thus proposed:
|
||||
|
|
@ -897,20 +897,20 @@ is thus proposed:
|
|||
</subSubSection>
|
||||
<subSubSection id="C334A5AF4C61C6401EE72856FB6F2681" pageId="23" pageNumber="24" type="reference_group">
|
||||
<paragraph id="8B91F6244C61C6401EE72856FB442489" blockId="23.[151,1436,151,2038]" pageId="23" pageNumber="24">
|
||||
<taxonomicName id="4C2E8DA74C61C6401EE72856FD4424E6" authority="Mabille, 1878" authorityName="Mabille" authorityYear="1878" box="[199,663,1402,1427]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andravahana">
|
||||
<taxonomicName id="4C2E8DA74C61C6401EE72856FD4424E6" authority="Mabille, 1878" authorityName="Mabille" authorityYear="1878" box="[199,663,1402,1427]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andravahana">
|
||||
<emphasis id="B95A2A364C61C6401EE72856FE0F24E7" box="[199,476,1402,1427]" italics="true" pageId="23" pageNumber="24">Heteropsis andravahana</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C61C6401FCB2857FD5D24E6" author="Mabille" box="[491,654,1402,1427]" pageId="23" pageNumber="24" refString="Mabille, P. (1878) Lepidoptera Africana. Bulletin de la Societe Zoologique de France, 3, 81 - 95." type="journal article" year="1878">Mabille, 1878</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. LT ♀, here designated (
|
||||
<figureCitation id="1315EAA14C61C6401D8C2857FC2324E7" box="[940,1008,1402,1427]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6</figureCitation>
|
||||
<figureCitation id="1315EAA14C61C6401D8C2857FC2324E7" box="[940,1008,1402,1427]" captionStart="FIGURE 6. A" captionStartId="24.[151,250,1718,1740]" captionTargetBox="[210,1375,193,1675]" captionTargetId="figure@24.[210,1377,193,1694]" captionTargetPageId="24" captionText="FIGURE 6. A: pastiche of illustrations from Mabille ([1885]) in Grandidier’s work, Figs 6, 7, 8, 9 from Pl. 5, Figs 2, 2 a from Pl. 7 B. The ♂ “ var. ” in (Pl. 5: Fig. 8) of that work might possibly represent Ht. harveyi Lees & Kremen, sp. nov., while the ♂ in (Pl. 5, Figs 7, 8) probably represents Ht. viettei (Lees, 2003) and the ♀ upperside in (Pl. 5: Fig. 9) probably represents the same species, described as Mycalesis difficilis Mabille, 1880 (probable HT illustrated: Pl. 5, Figs 2, 2 a). B: alternative illustration prior to hand colouring (from copy of Mabille ([1885]) in BHL online library) of Mabille’s Pl. 5: Fig. 9; C: Ht. andravahana (Mabille, 1878), stat. rev. ♀ (Anjanaharibe Sud 25 / 11 / 2014, photo: D. C. Lees), representing a dry seasonal form of this species, for comparison with D: chosen LT ♀ of Mycalesis andravahana, DV, VV (BMNH (E) # 674841; considered to represent the ♀ specimen depicted by Mabille ([1885]: Pl. 5. Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.264603" httpUri="https://zenodo.org/record/264603/files/figure.png" pageId="23" pageNumber="24">Fig. 6</figureCitation>
|
||||
D), lacking abdomen (the only known surviving potential ST, from the Grose-Smith collection): BMNH(E) #674841, bearing labels “LT (round purplebordered label)/
|
||||
<typeStatus id="549548864C61C6401F6828EDFE5224AD" box="[328,385,1472,1497]" pageId="23" pageNumber="24">Type</typeStatus>
|
||||
HT (round red-bordered label)/
|
||||
<typeStatus id="549548864C61C6401CC628EDFCFC24AD" box="[742,815,1472,1497]" pageId="23" pageNumber="24">TYPE</typeStatus>
|
||||
. [white red printed label]/Ex Grose-Smith, 1910/Joicey Bequest Brit. Mus. 1934-120./[Colour photocopy of
|
||||
<figureCitation id="1315EAA14C61C6401CC528C9FCF52489" box="[741,806,1508,1533]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="23" pageNumber="24">Fig. 9</figureCitation>
|
||||
<figureCitation id="1315EAA14C61C6401CC528C9FCF52489" box="[741,806,1508,1533]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="23" pageNumber="24">Fig. 9</figureCitation>
|
||||
from plate 5 of Mabille, 1887]”;
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C61C6401EE72B2AFF332730" blockId="23.[151,1436,151,2038]" pageId="23" pageNumber="24">
|
||||
|
|
@ -958,7 +958,7 @@ LT ♂, here designated (of 4 potential STs), lacking abdomen: BMNH(E) #674827,
|
|||
(round red-bordered label)/
|
||||
<collectingCountry id="F339B6B44C61C6401EB72A28FEF3266A" box="[151,288,1797,1822]" name="Madagascar" pageId="23" pageNumber="24">Madagascar</collectingCountry>
|
||||
, Antsianaka, Perrot Freres, 2e Semestre 1893/Ex Oberthür Coll. Brit. Mus. 1927-3/
|
||||
<taxonomicName id="4C2E8DA74C61C6401AE42A28FEA72636" authority="Obthr." authorityName="Obthr." class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="undulosa" variety="luctuosa">
|
||||
<taxonomicName id="4C2E8DA74C61C6401AE42A28FEA72636" authority="Obthr." authorityName="Obthr." class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="variety" species="undulosa" variety="luctuosa">
|
||||
<emphasis id="B95A2A364C61C6401AE42A28FA4F266A" box="[1220,1436,1797,1822]" italics="true" pageId="23" pageNumber="24">Gallienia undulosa</emphasis>
|
||||
var.
|
||||
<emphasis id="B95A2A364C61C6401EE72A04FEF52636" box="[199,294,1833,1858]" italics="true" pageId="23" pageNumber="24">luctuosa</emphasis>
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<mods:end id="924E7DDD447C36DB23CBE69CAD635D6D">97</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:classification id="A75BECD616631328149946F8099C08D8">journal article</mods:classification>
|
||||
<mods:identifier id="E0C6A2898F83AF39F5860BAB93879432" type="CLB-Dataset">38930</mods:identifier>
|
||||
<mods:identifier id="758F32C21D9888F930B8756DD5B3915F" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
|
||||
<mods:identifier id="D4720CD5EAAEF3EFB93F776E5227C429" type="GBIF-Dataset">a4e408c4-2575-4b5c-878b-e1b9010ec619</mods:identifier>
|
||||
<mods:identifier id="41826FCD1948C553990B21EF966EDE3E" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="CAA636D40FD451FF08AB018E44A8DE38" type="Zenodo-Dep">264597</mods:identifier>
|
||||
<mods:identifier id="5A99037BF16FD25BC32BED2137DF43F5" type="ZooBank">CFA586DA-10EE-468B-80EE-35351E3845FD</mods:identifier>
|
||||
<mods:classification id="877AF16C96421AB6937FDCF7CF7344DD">journal article</mods:classification>
|
||||
<mods:identifier id="591009EEDDB94852102171A05AA9108C" type="CLB-Dataset">38930</mods:identifier>
|
||||
<mods:identifier id="2BEB1131783D2C214443EBD102CCE351" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
|
||||
<mods:identifier id="A122C2D82C85FA96999D099BCD84F4A6" type="GBIF-Dataset">a4e408c4-2575-4b5c-878b-e1b9010ec619</mods:identifier>
|
||||
<mods:identifier id="C257DAE6F20C79E1411F73178F9E8B40" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="F07582AFEF66DE523E1C5D52D3D00B7E" type="Zenodo-Dep">264597</mods:identifier>
|
||||
<mods:identifier id="904BD7AE055588F6351F8723DAF208DC" type="ZooBank">CFA586DA-10EE-468B-80EE-35351E3845FD</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="038747324C66C6431EB72E14FCDB240A" ID-DOI="http://doi.org/10.5281/zenodo.6086414" ID-GBIF-Taxon="120817087" ID-Zenodo-Dep="6086414" LSID="urn:lsid:plazi:treatment:038747324C66C6431EB72E14FCDB240A" httpUri="http://treatment.plazi.org/id/038747324C66C6431EB72E14FCDB240A" lastPageId="20" lastPageNumber="21" pageId="16" pageNumber="17">
|
||||
<treatment id="038747324C66C6431EB72E14FCDB240A" ID-DOI="http://doi.org/10.5281/zenodo.6086414" ID-GBIF-Taxon="120817087" ID-Zenodo-Dep="6086414" LSID="urn:lsid:plazi:treatment:038747324C66C6431EB72E14FCDB240A" httpUri="http://treatment.plazi.org/id/038747324C66C6431EB72E14FCDB240A" lastPageId="20" lastPageNumber="21" pageId="16" pageNumber="17" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C66C6471EB72E14FBC62201" pageId="16" pageNumber="17" type="nomenclature">
|
||||
<paragraph id="8B91F6244C66C6471EB72E14FCD92227" blockId="16.[151,1045,825,885]" box="[151,778,825,851]" pageId="16" pageNumber="17">
|
||||
<heading id="D0D941484C66C6471EB72E14FCD92227" bold="true" box="[151,778,825,851]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
|
||||
<emphasis id="B95A2A364C66C6471EB72E14FCD92227" bold="true" box="[151,778,825,851]" pageId="16" pageNumber="17">
|
||||
<taxonomicName id="4C2E8DA74C66C6471EB72E14FD4C2227" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[151,671,825,851]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C66C6471EB72E14FD4C2227" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[151,671,825,851]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C66C6471EB72E14FE582227" bold="true" box="[151,395,825,851]" italics="true" pageId="16" pageNumber="17">Heteropsis hazovola</emphasis>
|
||||
Lees & Raharitsimba
|
||||
</taxonomicName>
|
||||
|
|
@ -70,7 +70,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C66C6471CA82EEFFD2622AF" box="[648,757,962,987]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C66C6471D142EEFFCAE22AF" box="[820,893,962,987]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="16" pageNumber="17">Fig. 4</figureCitation>
|
||||
<figureCitation id="1315EAA14C66C6471D142EEFFCAE22AF" box="[820,893,962,987]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." figureDoi="http://doi.org/10.5281/zenodo.264601" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="16" pageNumber="17">Fig. 4</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C66C6471D8D2EEFFBE722AF" box="[941,1076,962,987]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
|
||||
NE, Ambery R., Masoala E.,
|
||||
|
|
@ -108,7 +108,7 @@ barcode].
|
|||
010289117 [
|
||||
<collectingCountry id="F339B6B44C66C6471C7D2954FD4B25E5" box="[605,664,1145,1169]" name="Qatar" pageId="16" pageNumber="17">QTR</collectingCountry>
|
||||
barcode]; ♀ (
|
||||
<figureCitation id="1315EAA14C66C6471D142955FCAB25E5" box="[820,888,1144,1169]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="16" pageNumber="17">Fig. 4</figureCitation>
|
||||
<figureCitation id="1315EAA14C66C6471D142955FCAB25E5" box="[820,888,1144,1169]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." figureDoi="http://doi.org/10.5281/zenodo.264601" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="16" pageNumber="17">Fig. 4</figureCitation>
|
||||
B),
|
||||
<collectingCountry id="F339B6B44C66C6471DBF2955FBF525E5" box="[927,1062,1144,1169]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
|
||||
NE, Antsamanarana R., Masoala E., [
|
||||
|
|
@ -167,7 +167,7 @@ voucher; cytochrome b], IA302 [isotope voucher],
|
|||
barcode voucher], KA2053 [=KA-P2053;
|
||||
<collectionCode id="ED3F6EE14C66C6471C34287AFD82241B" box="[532,593,1367,1391]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="16" pageNumber="17">DNA</collectionCode>
|
||||
extract number]; ♂ (
|
||||
<figureCitation id="1315EAA14C66C6471D60287BFC50241B" box="[832,899,1366,1391]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="16" pageNumber="17">Fig. 5</figureCitation>
|
||||
<figureCitation id="1315EAA14C66C6471D60287BFC50241B" box="[832,899,1366,1391]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264602" httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="16" pageNumber="17">Fig. 5</figureCitation>
|
||||
B), data as above but: 17:02, D.C. Lees:
|
||||
<collectionCode id="ED3F6EE14C66C6471B72287AFAA9241B" box="[1362,1402,1367,1391]" pageId="16" pageNumber="17">DL</collectionCode>
|
||||
93- 0017; 26
|
||||
|
|
@ -209,7 +209,7 @@ NE, Masoala E., Ambery R.,
|
|||
<paragraph id="8B91F6244C66C6471EE728CBFD6D2615" blockId="16.[151,1436,926,2032]" pageId="16" pageNumber="17">
|
||||
<emphasis id="B95A2A364C66C6471EE728CBFE92248B" bold="true" box="[199,321,1510,1535]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
|
||||
this species has rather short and less pointed HW tails compared with
|
||||
<taxonomicName id="4C2E8DA74C66C6471A6F28C5FB08248B" box="[1103,1243,1510,1535]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471A6F28C5FB08248B" box="[1103,1243,1510,1535]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C66C6471A6F28C5FB08248B" box="[1103,1243,1510,1535]" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -221,11 +221,11 @@ and
|
|||
<emphasis id="B95A2A364C66C6471F832B26FDF52757" box="[419,550,1546,1571]" italics="true" pageId="16" pageNumber="17">Ht. ankoma</emphasis>
|
||||
</taxonomicName>
|
||||
. ♂♂ of
|
||||
<taxonomicName id="4C2E8DA74C66C6471CA12B26FCDC2757" box="[641,783,1546,1571]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471CA12B26FCDC2757" box="[641,783,1546,1571]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C66C6471CA12B26FCDC2757" box="[641,783,1546,1571]" italics="true" pageId="16" pageNumber="17">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
have a dense black latero-ventral androconial patch on the abdominal sternum, whereas
|
||||
<taxonomicName id="4C2E8DA74C66C6471FC52B03FDA72732" box="[485,628,1581,1606]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471FC52B03FDA72732" box="[485,628,1581,1606]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C66C6471FC52B03FDA72732" box="[485,628,1581,1606]" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -233,11 +233,11 @@ and
|
|||
<emphasis id="B95A2A364C66C6471C8E2B03FCE02732" box="[686,819,1581,1606]" italics="true" pageId="16" pageNumber="17">Ht. ankoma</emphasis>
|
||||
</taxonomicName>
|
||||
(particularly similar superficially) lack obvious black ventral abdominal scales in the ♂. Ventral wing pattern is very similar to other species in the group including
|
||||
<taxonomicName id="4C2E8DA74C66C6471B592B7FFF2A27F9" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471B592B7FFF2A27F9" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C66C6471B592B7FFF2A27F9" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C66C6471F2D2B5BFE7127F9" box="[269,418,1652,1677]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471F2D2B5BFE7127F9" box="[269,418,1652,1677]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C66C6471F2D2B5BFE7127F9" box="[269,418,1652,1677]" italics="true" pageId="16" pageNumber="17">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
, and to a lesser extent,
|
||||
|
|
@ -252,15 +252,15 @@ and
|
|||
<typeStatus id="549548864C66C6471FC22BB4FDC127C5" box="[482,530,1689,1713]" pageId="16" pageNumber="17">type</typeStatus>
|
||||
series of
|
||||
<emphasis id="B95A2A364C66C6471CA12BB4FCCA27C5" box="[641,793,1688,1713]" italics="true" pageId="16" pageNumber="17">
|
||||
<taxonomicName id="4C2E8DA74C66C6471CA12BB4FCC627C5" box="[641,789,1688,1713]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">Ht. hazovola</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C66C6471CA12BB4FCC627C5" box="[641,789,1688,1713]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">Ht. hazovola</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
On the dorsum, in fresh specimens,
|
||||
<taxonomicName id="4C2E8DA74C66C6471A982BB4FA9627C5" box="[1208,1349,1688,1713]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471A982BB4FA9627C5" box="[1208,1349,1688,1713]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C66C6471A982BB4FA9627C5" box="[1208,1349,1688,1713]" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
has the darkest wing colouration of these species, whereas in
|
||||
<taxonomicName id="4C2E8DA74C66C6471CD42B90FC5727A0" box="[756,900,1723,1748]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471CD42B90FC5727A0" box="[756,900,1723,1748]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C66C6471CD42B90FC5727A0" box="[756,900,1723,1748]" italics="true" pageId="16" pageNumber="17">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
, the dorsal wing background colour tends to be reddish brown.
|
||||
|
|
@ -268,11 +268,11 @@ has the darkest wing colouration of these species, whereas in
|
|||
<emphasis id="B95A2A364C66C6471F6D2BCDFE0B278C" box="[333,472,1759,1784]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||||
</taxonomicName>
|
||||
has an even lighter mid-brown dorsal colour than
|
||||
<taxonomicName id="4C2E8DA74C66C6471A102BCDFB10278C" box="[1072,1219,1759,1784]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471A102BCDFB10278C" box="[1072,1219,1759,1784]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C66C6471A102BCDFB10278C" box="[1072,1219,1759,1784]" italics="true" pageId="16" pageNumber="17">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
, but generally has space-M3 ocellus HWD strongly expressed in the ♂.
|
||||
<taxonomicName id="4C2E8DA74C66C6471CCD2A2EFCAF266F" box="[749,892,1794,1819]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471CCD2A2EFCAF266F" box="[749,892,1794,1819]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C66C6471CCD2A2EFCAF266F" box="[749,892,1794,1819]" italics="true" pageId="16" pageNumber="17">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
shares with
|
||||
|
|
@ -297,7 +297,7 @@ has a darker FWD margin). HWD this uniform dull reddish brown colour with a dark
|
|||
<emphasis id="B95A2A364C66C6471B592AF5FA4F269B" box="[1401,1436,2008,2031]" italics="true" pageId="16" pageNumber="17">Ht.</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C67C6461EB72DBAFD922553" blockId="17.[151,1437,151,2035]" pageId="17" pageNumber="18">
|
||||
<taxonomicName id="4C2E8DA74C67C6461EB72DBAFF2A21C4" box="[151,249,151,176]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C67C6461EB72DBAFF2A21C4" box="[151,249,151,176]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C67C6461EB72DBAFF2A21C4" box="[151,249,151,176]" italics="true" pageId="17" pageNumber="18">alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
, more similar to the configuration in
|
||||
|
|
@ -313,11 +313,11 @@ where they are relatively short. FW space-CuA1 ocellus weakly expressed. FWD spa
|
|||
<emphasis id="B95A2A364C67C6461CD22C98FC5020B8" box="[754,899,435,460]" italics="true" pageId="17" pageNumber="18">Ht. antahala</emphasis>
|
||||
</taxonomicName>
|
||||
group. Highly irrorated pattern of dark brown strigulae and yellow flecks on a lilac-washed light brown background, but the lilac wash is only strongly evident in the HT and one of the PT ♂♂, and to a lesser extent in the PT ♀, which has a generally lighter greyer-brown colouration on either surface. A striking pattern of ventral irrorations (dark strigulae) which are most finely divided proximad of the jagged dark brown Mb, but continue distad of it in both wings except around the ocelli. The very thick Mb is sharply inflexed around the CuA1 ocelli in both wings, quite jagged, strongly concave in space-M3- CuA1, and again in space-1A, fairly straight to mid costa from just below vein M3. FWV Mb forms an angle distad near vein M2, and there is another dark FWV band slightly proximad but along the irroration, the PMb is not distinct in either wing. There is a prominent yellow patch (highlight) at the base of space-M2 just distad of the Mb. Such underside patterns are not particularly special for
|
||||
<taxonomicName id="4C2E8DA74C67C6461CDF2FD9FC5D2278" box="[767,910,755,780]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C67C6461CDF2FD9FC5D2278" box="[767,910,755,780]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C67C6461CDF2FD9FC5D2278" box="[767,910,755,780]" italics="true" pageId="17" pageNumber="18">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
: in particular,
|
||||
<taxonomicName id="4C2E8DA74C67C6461A0F2FD9FB682278" box="[1071,1211,755,780]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C67C6461A0F2FD9FB682278" box="[1071,1211,755,780]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C67C6461A0F2FD9FB682278" box="[1071,1211,755,780]" italics="true" pageId="17" pageNumber="18">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -327,7 +327,7 @@ and
|
|||
are very similar and confusable ventrally, but the
|
||||
<typeStatus id="549548864C67C6461CBA2E3AFD19225B" box="[666,714,791,815]" pageId="17" pageNumber="18">type</typeStatus>
|
||||
material of
|
||||
<taxonomicName id="4C2E8DA74C67C6461D732E3AFC31225B" box="[851,994,790,815]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C67C6461D732E3AFC31225B" box="[851,994,790,815]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C67C6461D732E3AFC31225B" box="[851,994,790,815]" italics="true" pageId="17" pageNumber="18">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
seems more deeply washed with violet colour.
|
||||
|
|
@ -353,7 +353,7 @@ the ♀ PT. ♀ PT HWV ocellus expression similar to the ♂ HT but larger, exce
|
|||
<emphasis id="B95A2A364C67C6461F9429B2FD9725C3" box="[436,580,1182,1207]" italics="true" pageId="17" pageNumber="18">Ht. antahala</emphasis>
|
||||
</taxonomicName>
|
||||
group (the black spot in
|
||||
<figureCitation id="1315EAA14C67C6461D4729B3FC7F25C3" box="[871,940,1182,1207]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
|
||||
<figureCitation id="1315EAA14C67C6461D4729B3FC7F25C3" box="[871,940,1182,1207]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." figureDoi="http://doi.org/10.5281/zenodo.264601" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
|
||||
A, right, FWV, is a broken wing fragment). Blackish brush emanating near base of space-CuA1, as usual for
|
||||
<taxonomicName id="4C2E8DA74C67C6461DB629EEFBFC25AF" box="[918,1071,1218,1243]" class="Magnoliopsida" family="Fabaceae" genus="Smithia" kingdom="Plantae" order="Fabales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="paradoxa">
|
||||
<emphasis id="B95A2A364C67C6461DB629EEFBFC25AF" box="[918,1071,1218,1243]" italics="true" pageId="17" pageNumber="18">Ht. paradoxa</emphasis>
|
||||
|
|
@ -367,23 +367,23 @@ group, over reflexed veins 1A+2A + 3A, and potentially contacting a broad and co
|
|||
<emphasis id="B95A2A364C67C6461ADC2826FA462457" box="[1276,1429,1290,1315]" italics="true" pageId="17" pageNumber="18">Ht. paradoxa</emphasis>
|
||||
</taxonomicName>
|
||||
, despite a weakly developed hindwing brush arising in the cubital region), that is present in
|
||||
<taxonomicName id="4C2E8DA74C67C6461ABF2802FAFC2433" box="[1183,1327,1326,1351]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C67C6461ABF2802FAFC2433" box="[1183,1327,1326,1351]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C67C6461ABF2802FAFC2433" box="[1183,1327,1326,1351]" italics="true" pageId="17" pageNumber="18">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C67C6461B1F2803FA502433" box="[1343,1411,1326,1351]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
|
||||
<figureCitation id="1315EAA14C67C6461B1F2803FA502433" box="[1343,1411,1326,1351]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." figureDoi="http://doi.org/10.5281/zenodo.264601" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
|
||||
C, PT ♂) but not evident in
|
||||
<taxonomicName id="4C2E8DA74C67C6461F8D287EFDE3241F" box="[429,560,1362,1387]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ankoma">
|
||||
<emphasis id="B95A2A364C67C6461F8D287EFDE3241F" box="[429,560,1362,1387]" italics="true" pageId="17" pageNumber="18">Ht. ankoma</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C67C6461C1F287FFD51241E" box="[575,642,1362,1387]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
|
||||
<figureCitation id="1315EAA14C67C6461C1F287FFD51241E" box="[575,642,1362,1387]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." figureDoi="http://doi.org/10.5281/zenodo.264601" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
|
||||
D, LT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C67C6461D23287EFC53241F" box="[771,896,1362,1387]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ankoma">
|
||||
<emphasis id="B95A2A364C67C6461D23287EFC53241F" box="[771,896,1362,1387]" italics="true" pageId="17" pageNumber="18">M. ankoma</emphasis>
|
||||
</taxonomicName>
|
||||
) and indeed in
|
||||
<taxonomicName id="4C2E8DA74C67C6461A0D287EFB68241F" box="[1069,1211,1362,1387]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C67C6461A0D287EFB68241F" box="[1069,1211,1362,1387]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C67C6461A0D287EFB68241F" box="[1069,1211,1362,1387]" italics="true" pageId="17" pageNumber="18">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
it is more extensive and obvious than in others of the group that exhibit ventral blackish androconial scales, namely
|
||||
|
|
@ -394,7 +394,7 @@ it is more extensive and obvious than in others of the group that exhibit ventra
|
|||
</emphasis>
|
||||
and
|
||||
<emphasis id="B95A2A364C67C6461F1028B6FE1724C6" box="[304,452,1435,1458]" italics="true" pageId="17" pageNumber="18">
|
||||
<taxonomicName id="4C2E8DA74C67C6461F1028B6FE1224C6" box="[304,449,1435,1458]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="masoura">Ht. masoura</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C67C6461F1028B6FE1224C6" box="[304,449,1435,1458]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="masoura">Ht. masoura</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
|
|
@ -405,7 +405,7 @@ and
|
|||
<emphasis id="B95A2A364C67C6461D7D2B2AFC30276B" box="[861,995,1542,1567]" italics="true" pageId="17" pageNumber="18">Ht. mabillei</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C67C6461A3A2B2AFB75276B" box="[1050,1190,1542,1567]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C67C6461A3A2B2AFB75276B" box="[1050,1190,1542,1567]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C67C6461A3A2B2AFB75276B" box="[1050,1190,1542,1567]" italics="true" pageId="17" pageNumber="18">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
|
|
@ -413,7 +413,7 @@ and
|
|||
<paragraph id="8B91F6244C67C6461EE72B07FED92687" blockId="17.[151,1437,151,2035]" pageId="17" pageNumber="18">
|
||||
<emphasis id="B95A2A364C67C6461EE72B07FE992737" bold="true" box="[199,330,1578,1603]" pageId="17" pageNumber="18">♂ genitalia</emphasis>
|
||||
: 26DL (PT,
|
||||
<figureCitation id="1315EAA14C67C6461FF82B07FDC92737" box="[472,538,1578,1603]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="17" pageNumber="18">Fig. 5</figureCitation>
|
||||
<figureCitation id="1315EAA14C67C6461FF82B07FDC92737" box="[472,538,1578,1603]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264602" httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="17" pageNumber="18">Fig. 5</figureCitation>
|
||||
B, LV, DV): tegumen shallowly domed from LV (wide concave proximad notch from DV), with strong brow, tegumen much longer than sickle-shaped uncus with strong ventrad ‘dewlap’-like cusp and angularly raised ‘head’ (characteristic of
|
||||
<taxonomicName id="4C2E8DA74C67C6461CFB2B5EFCB827FF" box="[731,875,1650,1675]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="antahala">
|
||||
<emphasis id="B95A2A364C67C6461CFB2B5EFCB827FF" box="[731,875,1650,1675]" italics="true" pageId="17" pageNumber="18">Ht. antahala</emphasis>
|
||||
|
|
@ -425,21 +425,21 @@ group), leading to hook with fairly straight edge (uncus is narrow to hook from
|
|||
in particular (
|
||||
<bibRefCitation id="EFBF8BD54C67C6461D3D2A0AFC1D264B" author="Lees" box="[797,974,1830,1855]" pageId="17" pageNumber="18" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997: 105</bibRefCitation>
|
||||
), protruding well beyond uncus tip at tip, slightly inbent towards midline with a short infacing spine, and a few serrae mesad towards tip (unlike
|
||||
<taxonomicName id="4C2E8DA74C67C6461B592A66FF2A26F3" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C67C6461B592A66FF2A26F3" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C67C6461B592A66FF2A26F3" italics="true" pageId="17" pageNumber="18">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
; 146DL, Fianarantsoa, illustrated in
|
||||
<figureCitation id="1315EAA14C67C6461CBC2A43FD0C26F3" box="[668,735,1902,1927]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="17" pageNumber="18">Fig. 5</figureCitation>
|
||||
<figureCitation id="1315EAA14C67C6461CBC2A43FD0C26F3" box="[668,735,1902,1927]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264602" httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="17" pageNumber="18">Fig. 5</figureCitation>
|
||||
C, which has a large number of teeth along the inside edge of the valves). Saccus strongly uprecurved, inflated towards tip. Aedeagus almost the same length as valves, slightly uprecurved towards blunt distal end, and broadening towards a cup-like ostium. Juxta narrow, not prominent proximad.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF51A6AC4C64C6451EB72A42FA442695" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="18" pageNumber="19" targetBox="[266,1320,200,1853]" targetPageId="18">
|
||||
<caption id="DF51A6AC4C64C6451EB72A42FA442695" ID-DOI="http://doi.org/10.5281/zenodo.264601" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="18" pageNumber="19" targetBox="[266,1320,200,1853]" targetPageId="18">
|
||||
<paragraph id="8B91F6244C64C6451EB72A42FA442695" blockId="18.[151,1437,1903,2017]" pageId="18" pageNumber="19">
|
||||
<emphasis id="B95A2A364C64C6451EB72A42FEC126F0" bold="true" box="[151,274,1903,1925]" pageId="18" pageNumber="19">FIGURE 4.</emphasis>
|
||||
Adults in DV (left), VV (right) unless otherwise stated.
|
||||
<emphasis id="B95A2A364C64C6451D6B2A42FC8E26F0" bold="true" box="[843,861,1903,1924]" pageId="18" pageNumber="19">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C64C6451DE62A5DFAA726F1" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[966,1396,1903,1925]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hazovola" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C64C6451DE62A5DFAA726F1" authority="Lees & Raharitsimba" authorityName="Lees & Raharitsimba" box="[966,1396,1903,1925]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hazovola" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C64C6451DE62A5DFB4626F1" box="[966,1173,1903,1925]" italics="true" pageId="18" pageNumber="19">Heteropsis hazovola</emphasis>
|
||||
Lees & Raharitsimba
|
||||
</taxonomicName>
|
||||
|
|
@ -450,13 +450,13 @@ Lees & Raharitsimba
|
|||
(Masoala; NHMUK010289116).
|
||||
<emphasis id="B95A2A364C64C6451C3D2AA3FDFD26D7" bold="true" box="[541,558,1934,1955]" pageId="18" pageNumber="19">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C64C6451CB82AA2FCC826D7" box="[664,795,1933,1956]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C64C6451CB82AA2FCC826D7" box="[664,795,1933,1956]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C64C6451CB82AA2FCC826D7" box="[664,795,1933,1956]" italics="true" pageId="18" pageNumber="19">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
(Masoala; NHMUK010289118).
|
||||
<emphasis id="B95A2A364C64C6451A542AA0FB5526D7" bold="true" box="[1140,1158,1933,1955]" pageId="18" pageNumber="19">C</emphasis>
|
||||
: Abdomen of HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C64C6451B5C2AA2FF2026B6" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C64C6451B5C2AA2FF2026B6" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C64C6451B5C2AA2FF2026B6" italics="true" pageId="18" pageNumber="19">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
(VV, left).
|
||||
|
|
@ -481,25 +481,25 @@ Mabille, 1878
|
|||
(BMNH(E) 672618). Scale bar: 30 mm (C–D with separate scale).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF51A6AC4C65C6441EB72B44FDE427C8" httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="19" pageNumber="20" targetBox="[259,1327,202,1604]" targetPageId="19">
|
||||
<caption id="DF51A6AC4C65C6441EB72B44FDE427C8" ID-DOI="http://doi.org/10.5281/zenodo.264602" httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="19" pageNumber="20" targetBox="[259,1327,202,1604]" targetPageId="19">
|
||||
<paragraph id="8B91F6244C65C6441EB72B44FDE427C8" blockId="19.[151,1436,1641,1724]" pageId="19" pageNumber="20">
|
||||
<emphasis id="B95A2A364C65C6441EB72B44FEC1270A" bold="true" box="[151,274,1641,1663]" pageId="19" pageNumber="20">FIGURE 5.</emphasis>
|
||||
♂ genitalia in LV (left), DV (right).
|
||||
<emphasis id="B95A2A364C65C6441CA22B44FD47270A" bold="true" box="[642,660,1641,1662]" pageId="19" pageNumber="20">A</emphasis>
|
||||
: ♂ genitalia of
|
||||
<taxonomicName id="4C2E8DA74C65C6441D122B47FBD1270B" box="[818,1026,1642,1663]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<taxonomicName id="4C2E8DA74C65C6441D122B47FBD1270B" box="[818,1026,1642,1663]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<emphasis id="B95A2A364C65C6441D122B47FBD1270B" box="[818,1026,1642,1663]" italics="true" pageId="19" pageNumber="20">Heteropsis mimetica</emphasis>
|
||||
</taxonomicName>
|
||||
(Anjanaharibe Sud; 278DL, CK529).
|
||||
<emphasis id="B95A2A364C65C6441BA42B44FA46270A" bold="true" box="[1412,1429,1641,1662]" pageId="19" pageNumber="20">B</emphasis>
|
||||
: ♂ genitalia of PT of
|
||||
<taxonomicName id="4C2E8DA74C65C6441FAC2BA4FDC627E9" box="[396,533,1671,1694]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C65C6441FAC2BA4FDC627E9" box="[396,533,1671,1694]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C65C6441FAC2BA4FDC627E9" box="[396,533,1671,1694]" italics="true" pageId="19" pageNumber="20">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
(Masoala; NHMUK010289179, 26DL).
|
||||
<emphasis id="B95A2A364C65C6441DF12BAAFC3027E9" bold="true" box="[977,995,1671,1693]" pageId="19" pageNumber="20">C</emphasis>
|
||||
: ♂ genitalia of
|
||||
<taxonomicName id="4C2E8DA74C65C6441A802BA4FAF627E9" box="[1184,1317,1671,1694]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alaokola" status="stat. nov.">
|
||||
<taxonomicName id="4C2E8DA74C65C6441A802BA4FAF627E9" box="[1184,1317,1671,1694]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alaokola" status="stat. nov.">
|
||||
<emphasis id="B95A2A364C65C6441A802BA4FAF627E9" box="[1184,1317,1671,1694]" italics="true" pageId="19" pageNumber="20">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
|
|
@ -550,7 +550,7 @@ Mabille,
|
|||
</bibRefCitation>
|
||||
</taxonomicName>
|
||||
in BMNH, plus an additional ST in MNHN ex Chris Ward from the Galichon collection, DCL-DB-2982). A LT has already been designated for
|
||||
<taxonomicName id="4C2E8DA74C62C6431A2E2DF2FB33218C" box="[1038,1248,223,248]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C62C6431A2E2DF2FB33218C" box="[1038,1248,223,248]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C62C6431A2E2DF2FB33218C" box="[1038,1248,223,248]" italics="true" pageId="20" pageNumber="21">Gallienia alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
(see above), and as there is more possibility for confusion in this group, it is necessary to select another LT. The ♂ LT of
|
||||
|
|
@ -558,11 +558,11 @@ in BMNH, plus an additional ST in MNHN ex Chris Ward from the Galichon collectio
|
|||
<emphasis id="B95A2A364C62C6431B0C2C29FF232034" italics="true" pageId="20" pageNumber="21">Mycalesis ankoma</emphasis>
|
||||
</taxonomicName>
|
||||
is here designated as that illustrated here,
|
||||
<figureCitation id="1315EAA14C62C6431CF52C0AFCCB2034" box="[725,792,295,320]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="20" pageNumber="21">Fig. 4</figureCitation>
|
||||
<figureCitation id="1315EAA14C62C6431CF52C0AFCCB2034" box="[725,792,295,320]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." figureDoi="http://doi.org/10.5281/zenodo.264601" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="20" pageNumber="21">Fig. 4</figureCitation>
|
||||
D–E (there are several very similar STs but it is possible this is also the ♂ illustrated in Pl. 6
|
||||
<figureCitation id="1315EAA14C62C6431C162C61FD5F2010" box="[566,652,332,357]" captionStart="FIGURE 11" captionStartId="37.[151,250,1177,1199]" captionTargetBox="[264,1363,233,1142]" captionTargetId="figure@37.[212,1379,190,1160]" captionTargetPageId="37" captionText="FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL)." httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="20" pageNumber="21">Figs 11</figureCitation>
|
||||
<figureCitation id="1315EAA14C62C6431C162C61FD5F2010" box="[566,652,332,357]" captionStart="FIGURE 11" captionStartId="37.[151,250,1177,1199]" captionTargetBox="[264,1363,233,1142]" captionTargetId="figure@37.[212,1379,190,1160]" captionTargetPageId="37" captionText="FIGURE 11. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis westwoodi Lees, sp. nov. (Antanandava, Andohahela; 236 DL). B: Ht. pauliani, ♂ genitalia, PT ♂ (Tsaratanana; 97 DL)." figureDoi="http://doi.org/10.5281/zenodo.264608" httpUri="https://zenodo.org/record/264608/files/figure.png" pageId="20" pageNumber="21">Figs 11</figureCitation>
|
||||
,
|
||||
<figureCitation id="1315EAA14C62C6431CB92C61FD662010" box="[665,693,332,356]" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="20" pageNumber="21">12</figureCitation>
|
||||
<figureCitation id="1315EAA14C62C6431CB92C61FD662010" box="[665,693,332,356]" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." figureDoi="http://doi.org/10.5281/zenodo.264609" httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="20" pageNumber="21">12</figureCitation>
|
||||
of
|
||||
<bibRefCitation id="EFBF8BD54C62C6431CFE2C61FC582011" author="Mabille" box="[734,907,332,357]" pageId="20" pageNumber="21" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter" year="1885">Mabille [1885]</bibRefCitation>
|
||||
), bearing the labels “
|
||||
|
|
@ -576,12 +576,12 @@ of
|
|||
<emphasis id="B95A2A364C62C6431F8F2CF0FDFD2081" box="[431,558,476,501]" italics="true" pageId="20" pageNumber="21">M. ankoma</emphasis>
|
||||
</taxonomicName>
|
||||
from the Grose-Smith collection (BMNH(E) 672618) is illustrated in
|
||||
<figureCitation id="1315EAA14C62C6431B662CF1FA5B2080" box="[1350,1416,476,501]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="20" pageNumber="21">
|
||||
<figureCitation id="1315EAA14C62C6431B662CF1FA5B2080" box="[1350,1416,476,501]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." figureDoi="http://doi.org/10.5281/zenodo.264601" httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="20" pageNumber="21">
|
||||
Fig.
|
||||
<specimenCount id="9D283DAD4C62C6431B5A2CF1FA5B2080" box="[1402,1416,476,500]" pageId="20" pageNumber="21" type="generic">4</specimenCount>
|
||||
</figureCitation>
|
||||
F.
|
||||
<taxonomicName id="4C2E8DA74C62C6431EB72F2CFEF5236C" box="[151,294,511,536]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C62C6431EB72F2CFEF5236C" box="[151,294,511,536]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C62C6431EB72F2CFEF5236C" box="[151,294,511,536]" italics="true" pageId="20" pageNumber="21">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
now constitutes the first described butterfly species apparently endemic to the Masoala National Park (Kremen
|
||||
|
|
@ -591,7 +591,7 @@ now constitutes the first described butterfly species apparently endemic to the
|
|||
<paragraph id="8B91F6244C62C6431EE72F6AFE36222B" blockId="20.[151,1437,151,1406]" pageId="20" pageNumber="21">
|
||||
<emphasis id="B95A2A364C62C6431EE72F6AFD352314" bold="true" box="[199,742,583,608]" pageId="20" pageNumber="21">Additional information. DNA divergences</emphasis>
|
||||
: COI-5P cluster number BOLD:ACW4994 (exemplars BMAD228-15 to BMAD230-15 from Masoala), 5.02% divergent to a still undescribed species (BMAD147-15, DL95-0005, Vohitraseva) which is in turn about 2–2.4% divergent to other members, whether conspecific or not, of cluster BOLD:ACW4938.
|
||||
<taxonomicName id="4C2E8DA74C62C6431FE82F98FD8923B9" box="[456,602,692,717]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C62C6431FE82F98FD8923B9" box="[456,602,692,717]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C62C6431FE82F98FD8923B9" box="[456,602,692,717]" italics="true" pageId="20" pageNumber="21">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
appears more closely related to
|
||||
|
|
@ -599,13 +599,13 @@ appears more closely related to
|
|||
<emphasis id="B95A2A364C62C6431DF22F98FB8923B9" box="[978,1114,692,717]" italics="true" pageId="20" pageNumber="21">Ht. mabillei</emphasis>
|
||||
</taxonomicName>
|
||||
in fact than to
|
||||
<taxonomicName id="4C2E8DA74C62C6431B2D2F98FA4F23B9" box="[1293,1436,692,717]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C62C6431B2D2F98FA4F23B9" box="[1293,1436,692,717]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C62C6431B2D2F98FA4F23B9" box="[1293,1436,692,717]" italics="true" pageId="20" pageNumber="21">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
(see above regarding the shallow divergence of exemplars in Aduse-Poku
|
||||
<emphasis id="B95A2A364C62C6431A2D2FF4FB982384" box="[1037,1099,727,752]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
|
||||
, 2010) but only 140 bp are currently available (BMAD131-15, DL14M0-0003, Anjozorobe, for which there is a G rather than A or T in the 517th nucleotide position) and this stretch is 3.1% divergent from the barcode of the HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C62C6431B272E09FA4F224F" box="[1287,1436,802,827]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C62C6431B272E09FA4F224F" box="[1287,1436,802,827]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C62C6431B272E09FA4F224F" box="[1287,1436,802,827]" italics="true" pageId="20" pageNumber="21">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
(BMAD229-15, DL93-0002).
|
||||
|
|
@ -615,7 +615,7 @@ in fact than to
|
|||
<paragraph id="8B91F6244C62C6431EE72E47FC5422BF" blockId="20.[151,1437,151,1406]" pageId="20" pageNumber="21">
|
||||
<emphasis id="B95A2A364C62C6431EE72E47FE3622F7" bold="true" box="[199,485,874,899]" pageId="20" pageNumber="21">Phylogeny/sister species</emphasis>
|
||||
: the sister taxon of
|
||||
<taxonomicName id="4C2E8DA74C62C6431CE62E41FC8522F7" box="[710,854,874,899]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<taxonomicName id="4C2E8DA74C62C6431CE62E41FC8522F7" box="[710,854,874,899]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hazovola">
|
||||
<emphasis id="B95A2A364C62C6431CE62E41FC8522F7" box="[710,854,874,899]" italics="true" pageId="20" pageNumber="21">Ht. hazovola</emphasis>
|
||||
</taxonomicName>
|
||||
(as “sp. 53”) was unresolved in
|
||||
|
|
@ -625,7 +625,7 @@ in fact than to
|
|||
<emphasis id="B95A2A364C62C6431CE72EBDFC9F22D3" box="[711,844,910,935]" italics="true" pageId="20" pageNumber="21">Ht. ankoma</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C62C6431DA52EBDFBC022D3" box="[901,1043,910,935]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C62C6431DA52EBDFBC022D3" box="[901,1043,910,935]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C62C6431DA52EBDFBC022D3" box="[901,1043,910,935]" italics="true" pageId="20" pageNumber="21">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
(“sp. 56”;
|
||||
|
|
@ -666,7 +666,7 @@ Aduse-Poku
|
|||
<paragraph id="8B91F6244C62C6431EE729A7FEDD25B3" blockId="20.[151,1437,151,1406]" pageId="20" pageNumber="21">
|
||||
<emphasis id="B95A2A364C62C6431EE729A7FE8A25D7" bold="true" box="[199,345,1162,1187]" pageId="20" pageNumber="21">Distribution</emphasis>
|
||||
: endemic as far as is known to the eastern lowlands of the Masoala Peninsula (
|
||||
<figureCitation id="1315EAA14C62C6431AD229A7FA9625D7" box="[1266,1349,1162,1187]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="20" pageNumber="21">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C62C6431AD229A7FA9625D7" box="[1266,1349,1162,1187]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="20" pageNumber="21">Fig. 30</figureCitation>
|
||||
A, light blue dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C62C6431EE729FFFB94259F" blockId="20.[151,1437,151,1406]" box="[199,1095,1234,1259]" pageId="20" pageNumber="21">
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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|
||||
<taxonomicName id="4C2E8DA74C69C6481EB72C61FE3C2012" authority="Lees" authorityName="Lees" box="[151,495,332,358]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanewrighti" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C69C6481EB72C61FE3C2012" authority="Lees" authorityName="Lees" box="[151,495,332,358]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanewrighti" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C69C6481EB72C61FE7D2012" bold="true" box="[151,430,332,358]" italics="true" pageId="31" pageNumber="32">Heteropsis vanewrighti</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -70,7 +70,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C69C6481C592CFAFD352084" box="[633,742,471,496]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C69C6481D3D2CFAFCB22084" box="[797,865,471,496]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
|
||||
<figureCitation id="1315EAA14C69C6481D3D2CFAFCB22084" box="[797,865,471,496]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C69C6481DAE2CFAFBC62084" box="[910,1045,471,496]" name="Madagascar" pageId="31" pageNumber="32">Madagascar</collectingCountry>
|
||||
NW, Bekolosy, RS Manongarivo,
|
||||
|
|
@ -104,7 +104,7 @@ Deposition
|
|||
:
|
||||
</emphasis>
|
||||
♀ (
|
||||
<figureCitation id="1315EAA14C69C6481C482F6AFD7D2314" box="[616,686,583,608]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
|
||||
<figureCitation id="1315EAA14C69C6481C482F6AFD7D2314" box="[616,686,583,608]" captionStart="FIGURE 9" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[262,1325,218,1725]" captionTargetId="figure@32.[252,1334,193,1747]" captionTargetPageId="32" captionText="FIGURE 9. Adults in DV (left), VV (right) A: HT ♂ of Heteropsis vanewrighti Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289124). B: PT ♀ of Ht. vanewrighti (Mt. Bekolosy; NHMUK 010289125). C: HT ♂ of Ht. imerina Lees, sp. nov. (Mantadia; NHMUK 010289132). D: PT ♀ of Ht. imerina (Ankasoka, route Lakato; DCL-DB- 2906, MNHN). Scale bar: 30 mm." figureDoi="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
|
||||
B),
|
||||
<collectingCountry id="F339B6B44C69C6481CF92F6AFCB32314" box="[729,864,583,608]" name="Madagascar" pageId="31" pageNumber="32">Madagascar</collectingCountry>
|
||||
NW, Bekolosy Mt., RS Manongarivo,
|
||||
|
|
@ -351,12 +351,12 @@ NW, Bekolosy, RS Manongarivo,
|
|||
<paragraph id="8B91F6244C69C6481EE72B5BFF0726BA" blockId="31.[151,1437,436,1998]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B95A2A364C69C6481EE72B5BFE9227FB" bold="true" box="[199,321,1654,1679]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
|
||||
Especially in ♂♂,
|
||||
<taxonomicName id="4C2E8DA74C69C6481C042B5AFD0427FB" box="[548,727,1654,1679]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C69C6481C042B5AFD0427FB" box="[548,727,1654,1679]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C69C6481C042B5AFD0427FB" box="[548,727,1654,1679]" italics="true" pageId="31" pageNumber="32">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
is the smallest species of
|
||||
<collectingCountry id="F339B6B44C69C6481A2B2B5BFBAB27FB" box="[1035,1144,1654,1679]" name="Madagascar" pageId="31" pageNumber="32">Malagasy</collectingCountry>
|
||||
<taxonomicName id="4C2E8DA74C69C6481AA22B5AFB2927FA" box="[1154,1274,1655,1678]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C69C6481AA22B5AFB2927FA" box="[1154,1274,1655,1678]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C69C6481AA22B5AFB2927FA" box="[1154,1274,1655,1678]" italics="true" pageId="31" pageNumber="32">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
so far known (only in Africa are found smaller mycalesines such as
|
||||
|
|
@ -367,7 +367,7 @@ so far known (only in Africa are found smaller mycalesines such as
|
|||
:
|
||||
<bibRefCitation id="EFBF8BD54C69C6481AAC2BB4FA9927C6" author="Condamin" box="[1164,1354,1689,1714]" pageId="31" pageNumber="32" refString="Condamin, M. (1973) Monographie du Genre Bicyclus (Lepidoptera: Satyridae). Institut Fondamental d' Afrique Noire, Dakar, 324 pp." type="book" year="1973">Condamin, 1973</bibRefCitation>
|
||||
). It can normally be distinguished from
|
||||
<taxonomicName id="4C2E8DA74C69C6481FDF2B93FDAD27A2" box="[511,638,1725,1750]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C69C6481FDF2B93FDAD27A2" box="[511,638,1725,1750]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C69C6481FDF2B93FDAD27A2" box="[511,638,1725,1750]" italics="true" pageId="31" pageNumber="32">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
by the lack of an androconial patch on the FWV, and by its restriction, as presently known, to the extreme northwestern part of the rainforest biome. In Marojejy, northeastern
|
||||
|
|
@ -381,27 +381,27 @@ Torres
|
|||
2001
|
||||
</bibRefCitation>
|
||||
), which could be confused with
|
||||
<taxonomicName id="4C2E8DA74C69C6481C672A05FD2A2634" box="[583,761,1831,1856]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C69C6481C672A05FD2A2634" box="[583,761,1831,1856]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C69C6481C672A05FD2A2634" box="[583,761,1831,1856]" italics="true" pageId="31" pageNumber="32">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
, which sometimes exhibits extremely reduced FWV patch. However, counting from the start of the standard 358 bp COI-5P barcode, a barcoded individual of
|
||||
<taxonomicName id="4C2E8DA74C69C6481ACC2A61FA4F2610" box="[1260,1436,1867,1892]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C69C6481ACC2A61FA4F2610" box="[1260,1436,1867,1892]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C69C6481ACC2A61FA4F2610" box="[1260,1436,1867,1892]" italics="true" pageId="31" pageNumber="32">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
from Bekolosy (CCDB-02225-F04, BMAD064-09) has the following five potentially autapomorphic nucleotide states relative to other members (in parentheses) of the
|
||||
<taxonomicName id="4C2E8DA74C69C6481D342ABEFC4026DF" box="[788,915,1938,1963]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C69C6481D342ABEFC4026DF" box="[788,915,1938,1963]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C69C6481D342ABEFC4026DF" box="[788,915,1938,1963]" italics="true" pageId="31" pageNumber="32">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
group: 268, 292, 547: T(C); 352: A(G); 451: C(T).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF51A6AC4C56C6771EB72BC5FF15262E" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="32" pageNumber="33" targetBox="[262,1325,218,1725]" targetPageId="32">
|
||||
<caption id="DF51A6AC4C56C6771EB72BC5FF15262E" ID-DOI="http://doi.org/10.5281/zenodo.264606" httpUri="https://zenodo.org/record/264606/files/figure.png" pageId="32" pageNumber="33" targetBox="[262,1325,218,1725]" targetPageId="32">
|
||||
<paragraph id="8B91F6244C56C6771EB72BC5FF15262E" blockId="32.[151,1436,1768,1882]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B95A2A364C56C6771EB72BC5FECF2789" bold="true" box="[151,284,1768,1790]" pageId="32" pageNumber="33">FIGURE 9.</emphasis>
|
||||
Adults in DV (left), VV (right)
|
||||
<emphasis id="B95A2A364C56C6771C832BC5FD662789" bold="true" box="[675,693,1768,1789]" pageId="32" pageNumber="33">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C56C6771D652BC4FBA8278A" authority="Lees" authorityName="Lees" box="[837,1147,1768,1790]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="vanewrighti" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C56C6771D652BC4FBA8278A" authority="Lees" authorityName="Lees" box="[837,1147,1768,1790]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="vanewrighti" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C56C6771D652BC4FBE9278A" box="[837,1082,1768,1790]" italics="true" pageId="32" pageNumber="33">Heteropsis vanewrighti</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -412,13 +412,13 @@ Lees
|
|||
(Mt. Bekolosy; NHMUK010289124).
|
||||
<emphasis id="B95A2A364C56C6771F5D2A2AFE5D2668" bold="true" box="[381,398,1799,1820]" pageId="32" pageNumber="33">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C56C6771FD22A25FD432668" box="[498,656,1798,1821]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C56C6771FD22A25FD432668" box="[498,656,1798,1821]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C56C6771FD22A25FD432668" box="[498,656,1798,1821]" italics="true" pageId="32" pageNumber="33">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
(Mt. Bekolosy; NHMUK010289125).
|
||||
<emphasis id="B95A2A364C56C6771A3B2A2BFBFE2668" bold="true" box="[1051,1069,1798,1820]" pageId="32" pageNumber="33">C</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C56C6771AB52A25FA912668" authority="Lees" authorityName="Lees" box="[1173,1346,1799,1821]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="imerina" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C56C6771AB52A25FA912668" authority="Lees" authorityName="Lees" box="[1173,1346,1799,1821]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="imerina" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C56C6771AB52A25FAD92669" box="[1173,1290,1800,1821]" italics="true" pageId="32" pageNumber="33">Ht. imerina</emphasis>
|
||||
Lees
|
||||
</taxonomicName>
|
||||
|
|
@ -429,7 +429,7 @@ Lees
|
|||
(Mantadia; NHMUK010289132).
|
||||
<emphasis id="B95A2A364C56C6771FD72A0BFDDA264F" bold="true" box="[503,521,1830,1851]" pageId="32" pageNumber="33">D</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C56C6771C582A0BFD3C264F" box="[632,751,1830,1851]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C56C6771C582A0BFD3C264F" box="[632,751,1830,1851]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C56C6771C582A0BFD3C264F" box="[632,751,1830,1851]" italics="true" pageId="32" pageNumber="33">Ht. imerina</emphasis>
|
||||
</taxonomicName>
|
||||
(Ankasoka, route Lakato; DCL-DB-2906, MNHN). Scale bar: 30 mm.
|
||||
|
|
@ -469,7 +469,7 @@ as revised here, with a near right angle inflexion around vein CuA1, curving con
|
|||
<paragraph id="8B91F6244C57C6761EE729AAFEDD277C" blockId="33.[151,1437,151,2013]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B95A2A364C57C6761EE729AAFEA625D4" bold="true" box="[199,373,1159,1184]" pageId="33" pageNumber="34">Male genitalia</emphasis>
|
||||
(
|
||||
<figureCitation id="1315EAA14C57C6761FA829AAFE1D25D4" box="[392,462,1159,1184]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="33" pageNumber="34">Fig. 8</figureCitation>
|
||||
<figureCitation id="1315EAA14C57C6761FA829AAFE1D25D4" box="[392,462,1159,1184]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264605" httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="33" pageNumber="34">Fig. 8</figureCitation>
|
||||
B, 233DL, PT): from LV: tegumen shallowly domed (with small concave proximad notch from DV), with fairly distinct brow down to scythe-hooked uncus. Gnathos fairly straight, thin and tapered, in line with up-hook of uncus in specimen examined (slightly incurved and not sinuate from DV bending round from stout oblong base). Tegumen-vinculum division slightly more than maximum dimension of tegumen. Valve base fits in rather oblong frame with rather angular ‘shoulder’ at base of dorsal edge, leading abruptly to gently elbowed rather parallel-sided valve arm which has a very slender ‘head’ for club with stout small spine at tip, uprecurved and significantly proud of uncus tip (strongly incurved at tip from DV and with a few short distad teeth). Aedeagus very long and thin, dorso-ventrally flattened, more than one and a half times length of valve, gently and smoothly uprecurved distad from ostium, which is itself long and cupped proximad, winged into two lobes proximad from DV. Saccus relatively long, and juxta featuring a prominent down-lip, which also has small side lobes.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -499,7 +499,7 @@ at Bekolosy mountain in the Réserve Spéciale de Manongarivo (
|
|||
<bibRefCitation id="EFBF8BD54C57C6761A502BEAFB282794" author="Butler" box="[1136,1275,1735,1760]" pageId="33" pageNumber="34" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(=
|
||||
<taxonomicName id="4C2E8DA74C57C6761B042BE4FEB22670" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C57C6761B042BE4FEB22670" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C57C6761B042BE4FF0F2670" italics="true" pageId="33" pageNumber="34">Heteropsis viettei</emphasis>
|
||||
Lees, 2003
|
||||
</taxonomicName>
|
||||
|
|
@ -512,11 +512,11 @@ ab.
|
|||
<emphasis id="B95A2A364C57C6761A6F2A3CFB1E265C" box="[1103,1229,1809,1832]" italics="true" pageId="33" pageNumber="34">marmorata</emphasis>
|
||||
<bibRefCitation id="EFBF8BD54C57C6761AF52A22FA4B265C" author="Aurivillius" box="[1237,1432,1807,1832]" pageId="33" pageNumber="34" refString="Aurivillius, P. O. C. (1911) In: Seitz, A. (Ed.), (1908 - 1925) Die Gross-Schmetterlinge der Erde. (2) 13 - Afrikanischen Faunengebietes, A. Kerner, Stuttgart, pp. 1 - 613, pls. 1 - 80." type="journal article" year="1911">Aurivillius, 1911</bibRefCitation>
|
||||
. This and a large series of specimens referred to
|
||||
<taxonomicName id="4C2E8DA74C57C6761C9D2A18FC412639" box="[701,914,1844,1869]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C57C6761C9D2A18FC412639" box="[701,914,1844,1869]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C57C6761C9D2A18FC412639" box="[701,914,1844,1869]" italics="true" pageId="33" pageNumber="34">Heteropsis harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
in the Oberthür collection are clearly distinct from
|
||||
<taxonomicName id="4C2E8DA74C57C6761EF52A74FE542604" box="[213,391,1879,1904]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C57C6761EF52A74FE542604" box="[213,391,1879,1904]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C57C6761EF52A74FE542604" box="[213,391,1879,1904]" italics="true" pageId="33" pageNumber="34">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
in being generally larger in size, possessing a FWV androconial patch, and occurring much further south in the central to southeastern parts of the rainforest biome.
|
||||
|
|
@ -537,7 +537,7 @@ HM
|
|||
<date id="FF90D0E44C54C6751EE52D91FECE21A1" box="[197,285,188,213]" pageId="34" pageNumber="35" value="20-06-24">240620</date>
|
||||
</accessionNumber>
|
||||
); for more details see under
|
||||
<taxonomicName id="4C2E8DA74C54C6751C7A2D90FD0B21A1" box="[602,728,188,213]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C54C6751C7A2D90FD0B21A1" box="[602,728,188,213]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C54C6751C7A2D90FD0B21A1" box="[602,728,188,213]" italics="true" pageId="34" pageNumber="35">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
|
|
@ -547,15 +547,15 @@ HM
|
|||
<paragraph id="8B91F6244C54C6751EE72DF2FD462034" blockId="34.[151,1437,151,1016]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B95A2A364C54C6751EE72DF2FE36218C" bold="true" box="[199,485,223,248]" pageId="34" pageNumber="35">Phylogeny/sister species</emphasis>
|
||||
: without a detailed phylogeographic analysis of the
|
||||
<taxonomicName id="4C2E8DA74C54C6751A1D2DCCFB6E218C" box="[1085,1213,223,248]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C54C6751A1D2DCCFB6E218C" box="[1085,1213,223,248]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C54C6751A1D2DCCFB6E218C" box="[1085,1213,223,248]" italics="true" pageId="34" pageNumber="35">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
-complex, the sister taxon of
|
||||
<taxonomicName id="4C2E8DA74C54C6751ED92C28FE7A2069" box="[249,425,260,285]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C54C6751ED92C28FE7A2069" box="[249,425,260,285]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C54C6751ED92C28FE7A2069" box="[249,425,260,285]" italics="true" pageId="34" pageNumber="35">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
is not yet certain, but the closest currently described relative is
|
||||
<taxonomicName id="4C2E8DA74C54C6751A4C2C28FB3A2069" box="[1132,1257,260,285]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C54C6751A4C2C28FB3A2069" box="[1132,1257,260,285]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C54C6751A4C2C28FB3A2069" box="[1132,1257,260,285]" italics="true" pageId="34" pageNumber="35">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
|
|
@ -593,11 +593,11 @@ sp. 20.
|
|||
<emphasis id="B95A2A364C54C6751EE72CD2FE8A236C" bold="true" box="[199,345,511,536]" pageId="34" pageNumber="35">Distribution</emphasis>
|
||||
: Bekolosy and Antsatrotro mountains (where found in 2011), Réserve Spéciale de Manongarivo, to where, as far as is known,
|
||||
<emphasis id="B95A2A364C54C6751FFC2F08FD482348" box="[476,667,548,573]" italics="true" pageId="34" pageNumber="35">
|
||||
<taxonomicName id="4C2E8DA74C54C6751FFC2F08FD5F2349" box="[476,652,548,573]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="vanewrighti">Ht. vanewrighti</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C54C6751FFC2F08FD5F2349" box="[476,652,548,573]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="vanewrighti">Ht. vanewrighti</taxonomicName>
|
||||
i
|
||||
</emphasis>
|
||||
s endemic (
|
||||
<figureCitation id="1315EAA14C54C6751D3B2F09FCBF2348" box="[795,876,548,573]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="34" pageNumber="35">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C54C6751D3B2F09FCBF2348" box="[795,876,548,573]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="34" pageNumber="35">Fig. 30</figureCitation>
|
||||
A, purple dots).
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C54C6751EE72F6AFB272314" blockId="34.[151,1437,151,1016]" box="[199,1268,583,608]" pageId="34" pageNumber="35">
|
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|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
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<treatment id="038747324C6DC6481EB72DBAFBA8218A" ID-DOI="http://doi.org/10.5281/zenodo.6086418" ID-GBIF-Taxon="120817090" ID-Zenodo-Dep="6086418" LSID="urn:lsid:plazi:treatment:038747324C6DC6481EB72DBAFBA8218A" httpUri="http://treatment.plazi.org/id/038747324C6DC6481EB72DBAFBA8218A" lastPageId="31" lastPageNumber="31" pageId="27" pageNumber="28">
|
||||
<treatment id="038747324C6DC6481EB72DBAFBA8218A" ID-DOI="http://doi.org/10.5281/zenodo.6086418" ID-GBIF-Taxon="120817090" ID-Zenodo-Dep="6086418" LSID="urn:lsid:plazi:treatment:038747324C6DC6481EB72DBAFBA8218A" httpUri="http://treatment.plazi.org/id/038747324C6DC6481EB72DBAFBA8218A" lastPageId="31" lastPageNumber="31" pageId="27" pageNumber="28" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C6DC64C1EB72DBAFBC621A7" pageId="27" pageNumber="28" type="nomenclature">
|
||||
<paragraph id="8B91F6244C6DC64C1EB72DBAFD6121C6" blockId="27.[151,1045,151,211]" box="[151,690,151,178]" pageId="27" pageNumber="28">
|
||||
<heading id="D0D941484C6DC64C1EB72DBAFD6121C6" bold="true" box="[151,690,151,178]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
|
||||
<emphasis id="B95A2A364C6DC64C1EB72DBAFD6121C6" bold="true" box="[151,690,151,178]" pageId="27" pageNumber="28">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1EB72DBAFD9421C6" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,583,151,178]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1EB72DBAFD9421C6" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[151,583,151,178]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C6DC64C1EB72DBAFEAA21C5" bold="true" box="[151,377,151,177]" italics="true" pageId="27" pageNumber="28">Heteropsis harveyi</emphasis>
|
||||
Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -63,7 +63,7 @@ Mabille
|
|||
♂ “var.” (
|
||||
<bibRefCitation id="EFBF8BD54C6DC64C1D642DD2FC21206C" author="Mabille" box="[836,1010,255,280]" pageId="27" pageNumber="28" refString="Mabille, P. (1885 - 1887) Histoire, Physique, Naturelle et Politique de Madagascar. Vol. 18 - 19. Imprimerie Nationale, Paris, 364 pp., 55 pls. [Grandidier, A. [1885 - 1887] (Ed.)] http: // dx. doi. org / 10.5962 / bhl. title. 1599" type="book chapter" year="1885">Mabille [1885]</bibRefCitation>
|
||||
: Pl. 5
|
||||
<figureCitation id="1315EAA14C6DC64C1A182DD2FBA8206C" box="[1080,1147,255,280]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="27" pageNumber="28">Fig. 8</figureCitation>
|
||||
<figureCitation id="1315EAA14C6DC64C1A182DD2FBA8206C" box="[1080,1147,255,280]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264605" httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="27" pageNumber="28">Fig. 8</figureCitation>
|
||||
, [1887: 57–59]);
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -122,14 +122,14 @@ Ackery
|
|||
<emphasis id="B95A2A364C6DC64C1A7D2CFAFB262084" box="[1117,1269,471,496]" italics="true" pageId="27" pageNumber="28">Andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
provenant de Fianarantsoa et d'Antsianaka” (subsequently known as
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1D202CD0FB9B2360" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[768,1096,508,532]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1D202CD0FB9B2360" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[768,1096,508,532]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C6DC64C1D202CD0FC102360" box="[768,963,509,532]" italics="true" pageId="27" pageNumber="28">Heteropsis viettei</emphasis>
|
||||
Lees, 2003
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFBF8BD54C6DC64C1A742CD0FB002360" author="Lees" box="[1108,1235,508,532]" pageId="27" pageNumber="28" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
: 62; 391–392, but here treated as
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1F772F0CFDA0234C" box="[343,627,543,568]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andravahana">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1F772F0CFDA0234C" box="[343,627,543,568]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andravahana">
|
||||
<emphasis id="B95A2A364C6DC64C1F772F0CFDA0234C" box="[343,627,543,568]" italics="true" pageId="27" pageNumber="28">Heteropsis andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
), “et 78 ♂♂ et 45 ♀♀ du supposé
|
||||
|
|
@ -137,7 +137,7 @@ Lees, 2003
|
|||
<emphasis id="B95A2A364C6DC64C1A612F32FB5A234C" box="[1089,1161,543,568]" italics="true" pageId="27" pageNumber="28">Wardii</emphasis>
|
||||
</taxonomicName>
|
||||
, capturés également à Fianarantsoa et dans l’Antsianaka"; referred to here as
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1D322F68FC412329" box="[786,914,580,605]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1D322F68FC412329" box="[786,914,580,605]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6DC64C1D322F68FC412329" box="[786,914,580,605]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
; All 78 ♂♂ and 13 of the ♀♀ of Oberthür’s “
|
||||
|
|
@ -178,7 +178,7 @@ Torres
|
|||
<paragraph id="8B91F6244C6DC64C1EE72FF9FAED2398" blockId="27.[151,1437,255,2020]" box="[199,1342,724,749]" pageId="27" pageNumber="28">
|
||||
<treatmentCitationGroup id="AB3ED10A4C6DC64C1EE72FF9FAED2398" box="[199,1342,724,749]" pageId="27" pageNumber="28">
|
||||
“
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1EF32FF8FDBD2398" authority="Lees" authorityName="Lees" box="[211,622,724,749]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi" subGenus="Henotesia">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1EF32FF8FDBD2398" authority="Lees" authorityName="Lees" box="[211,622,724,749]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi" subGenus="Henotesia">
|
||||
<emphasis id="B95A2A364C6DC64C1EF32FF8FE982398" box="[211,331,725,748]" italics="true" pageId="27" pageNumber="28">Heteropsis</emphasis>
|
||||
(
|
||||
<emphasis id="B95A2A364C6DC64C1F7C2FF8FE1D2398" box="[348,462,725,748]" italics="true" pageId="27" pageNumber="28">Henotesia</emphasis>
|
||||
|
|
@ -205,7 +205,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C6DC64C1CBD2FDAFCD92264" box="[669,778,759,784]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C6DC64C1D732FDAFC722264" box="[851,929,759,784]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="27" pageNumber="28">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C6DC64C1D732FDAFC722264" box="[851,929,759,784]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="27" pageNumber="28">Fig. 7</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C6DC64C1DF72FDAFB8D2264" box="[983,1118,759,784]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
|
||||
E, Sahavondronina VOI, Ranomafana National Park,
|
||||
|
|
@ -245,7 +245,7 @@ Deposition
|
|||
010289120 [
|
||||
<collectingCountry id="F339B6B44C6DC64C1FAC2EA6FE1422D7" box="[396,455,907,931]" name="Qatar" pageId="27" pageNumber="28">QTR</collectingCountry>
|
||||
barcode]; ♀ (
|
||||
<figureCitation id="1315EAA14C6DC64C1C452EA7FD7422D7" box="[613,679,906,931]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="27" pageNumber="28">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C6DC64C1C452EA7FD7422D7" box="[613,679,906,931]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="27" pageNumber="28">Fig. 7</figureCitation>
|
||||
B),
|
||||
<date id="FF90D0E44C6DC64C1CF12EA6FC9522D7" box="[721,838,907,931]" pageId="27" pageNumber="28" value="2014-11-08">8/11/2014</date>
|
||||
: 10:57, D.C. Lees:
|
||||
|
|
@ -351,33 +351,33 @@ E, Sahavondronina VOI, Ranomafana National Park,
|
|||
<subSubSection id="C334A5AF4C6DC64C1EE72881FE4C260C" pageId="27" pageNumber="28" type="diagnosis">
|
||||
<paragraph id="8B91F6244C6DC64C1EE72881FE4C260C" blockId="27.[151,1437,255,2020]" pageId="27" pageNumber="28">
|
||||
<emphasis id="B95A2A364C6DC64C1EE72881FE9224B1" bold="true" box="[199,321,1452,1477]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1F6D2880FDD224B1" box="[333,513,1452,1477]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1F6D2880FDD224B1" box="[333,513,1452,1477]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C6DC64C1F6D2880FDD224B1" box="[333,513,1452,1477]" italics="true" pageId="27" pageNumber="28">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
and an undescribed species (‘
|
||||
<emphasis id="B95A2A364C6DC64C1D4A2880FC5524B0" box="[874,902,1453,1476]" italics="true" pageId="27" pageNumber="28">Ht</emphasis>
|
||||
. sp. 20’) are both generally smaller than
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1B592880FF38249D" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1B592880FF38249D" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6DC64C1B592880FF38249D" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
and the former occurs only in NW
|
||||
<collectingCountry id="F339B6B44C6DC64C1C5728FDFCD3249D" box="[631,768,1488,1513]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
|
||||
, whereas
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1D4E28FCFC3F249D" box="[878,1004,1488,1513]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1D4E28FCFC3F249D" box="[878,1004,1488,1513]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6DC64C1D4E28FCFC3F249D" box="[878,1004,1488,1513]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
is generally similar to
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1ACC28FCFA4F249D" box="[1260,1436,1488,1513]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1ACC28FCFA4F249D" box="[1260,1436,1488,1513]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C6DC64C1ACC28FCFA4F249D" box="[1260,1436,1488,1513]" italics="true" pageId="27" pageNumber="28">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
in wing colouration, but in all known specimens shows a prominent androconial patch on the FWV. As currently recognized,
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1F3F2B31FE4E2740" box="[287,413,1563,1588]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1F3F2B31FE4E2740" box="[287,413,1563,1588]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6DC64C1F3F2B31FE4E2740" box="[287,413,1563,1588]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
occurs in the centre-east of the montane rainforest biome at latitudes south of about
|
||||
<geoCoordinate id="EE1A90E34C6DC64C1B6C2B36FA452747" box="[1356,1430,1559,1588]" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="5555" value="-17.5">17.5o S</geoCoordinate>
|
||||
, whereas
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1EDA2B6DFE78272C" box="[250,427,1599,1624]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1EDA2B6DFE78272C" box="[250,427,1599,1624]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C6DC64C1EDA2B6DFE78272C" box="[250,427,1599,1624]" italics="true" pageId="27" pageNumber="28">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
is only known from the extreme northwestern part of the rainforest biome, but
|
||||
|
|
@ -385,19 +385,19 @@ is only known from the extreme northwestern part of the rainforest biome, but
|
|||
. sp. 20 appears to occur partially sympatrically, distributed from the NE to CE of
|
||||
<collectingCountry id="F339B6B44C6DC64C1A212B4EFB582708" box="[1025,1163,1635,1660]" name="Madagascar" pageId="27" pageNumber="28">Madagascar</collectingCountry>
|
||||
.
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1AB82B49FAE82708" box="[1176,1339,1635,1660]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1AB82B49FAE82708" box="[1176,1339,1635,1660]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="westwoodi">
|
||||
<emphasis id="B95A2A364C6DC64C1AB82B49FAE82708" box="[1176,1339,1635,1660]" italics="true" pageId="27" pageNumber="28">Ht. westwoodi</emphasis>
|
||||
</taxonomicName>
|
||||
is quite distinctive from either of these species, having a more uniform colouration on the FWV towards the costa than any of these species, while
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1F8F2B81FDF127B7" box="[431,546,1708,1731]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1F8F2B81FDF127B7" box="[431,546,1708,1731]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C6DC64C1F8F2B81FDF127B7" box="[431,546,1708,1731]" italics="true" pageId="27" pageNumber="28">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
also has a prominent pale or whitish highlighting distal of the HWV Mb. Counting from the start of the 658 bp COI-5P barcode, the following four putatively fixed nucleotide differences in the DNA barcode appear (n=4) to be diagnostic within the
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1D042BD9FC712678" box="[804,930,1779,1804]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1D042BD9FC712678" box="[804,930,1779,1804]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6DC64C1D042BD9FC712678" box="[804,930,1779,1804]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
complex (cluster number BOLD:ACW5694) throughout its range, where in parentheses is specified the un-derived state: 274 C(T); 298 T(C); 466 A(G); 498 C(A). DNA barcoding is currently the best means for determining material of
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1A0A2A11FB782620" box="[1066,1195,1851,1876]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6DC64C1A0A2A11FB782620" box="[1066,1195,1851,1876]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6DC64C1A0A2A11FB782620" box="[1066,1195,1851,1876]" italics="true" pageId="27" pageNumber="28">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
among material that exhibit the FWV patch.
|
||||
|
|
@ -415,17 +415,17 @@ among material that exhibit the FWV patch.
|
|||
’). Distad of the central HWV band and highlighting it there is a more obscure dark violet to grey-brown region, and the FWV pattern is generally more dark-obscure and less marbled but with about four ochreous strigulae (Costal bars, henceforth ‘Cbs’) perpendicular to the costa beyond mid-costa.
|
||||
<emphasis id="B95A2A364C6AC64B1F352FD1FE592261" bold="true" box="[277,394,764,789]" pageId="28" pageNumber="29">Variation.</emphasis>
|
||||
♂♂ similar, but vary in the intensity of the darker brown irroration and Mb on the HWV. Sexually dimorphic, but marginally so. The ♂ and ♀ are one of the most similar of the
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1A5C2E0CFAD0224C" box="[1148,1283,799,824]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1A5C2E0CFAD0224C" box="[1148,1283,799,824]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C6AC64B1A5C2E0CFAD0224C" box="[1148,1283,799,824]" italics="true" pageId="28" pageNumber="29">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group and in fact the only pair in the
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1F862E68FDFD2229" box="[422,558,836,861]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1F862E68FDFD2229" box="[422,558,836,861]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C6AC64B1F862E68FDFD2229" box="[422,558,836,861]" italics="true" pageId="28" pageNumber="29">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
group to have been correctly matched by
|
||||
<bibRefCitation id="EFBF8BD54C6AC64B1A292E69FB102229" author="Oberthur" box="[1033,1219,836,861]" pageId="28" pageNumber="29" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür (1916)</bibRefCitation>
|
||||
, but the ♀ is larger and paler. The ♂ and ♀ have been photographed in copula (
|
||||
<figureCitation id="1315EAA14C6AC64B1D0D2E4AFCA322F4" box="[813,880,871,896]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="28" pageNumber="29">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C6AC64B1D0D2E4AFCA322F4" box="[813,880,871,896]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="28" pageNumber="29">Fig. 7</figureCitation>
|
||||
D). The ♀ is similar to the ♂, but rather larger and lighter brown on either surface, with a similar ocellus configuration, although the space-CuA1 ocelli tend to be larger in either wing.
|
||||
</paragraph>
|
||||
<paragraph id="8B91F6244C6AC64B1EE72EF9FE3A252C" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||||
|
|
@ -459,18 +459,18 @@ After Don Harvey, who first pointed out (to Claire Kremen) the FWV androconial p
|
|||
<paragraph id="8B91F6244C6AC64B1EE7283AFE3E26E1" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B95A2A364C6AC64B1EE7283AFE9F2444" bold="true" box="[199,332,1303,1328]" pageId="28" pageNumber="29">Discussion.</emphasis>
|
||||
Apart from
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1FFF2834FD812444" box="[479,594,1305,1328]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tianae" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1FFF2834FD812444" box="[479,594,1305,1328]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tianae" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C6AC64B1FFF2834FD812444" box="[479,594,1305,1328]" italics="true" pageId="28" pageNumber="29">Ht. tianae</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A269974D4C6AC64B1C7C283AFD6B2444" box="[604,696,1303,1328]" pageId="28" pageNumber="29" rank="species">
|
||||
<emphasis id="B95A2A364C6AC64B1C7C283AFD6B2444" bold="true" box="[604,696,1303,1328]" pageId="28" pageNumber="29">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1CE92834FC992444" box="[713,842,1303,1328]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1CE92834FC992444" box="[713,842,1303,1328]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6AC64B1CE92834FC992444" box="[713,842,1303,1328]" italics="true" pageId="28" pageNumber="29">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
is the
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1DBC2834FBC72444" box="[924,1044,1305,1328]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1DBC2834FBC72444" box="[924,1044,1305,1328]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C6AC64B1DBC2834FBC72444" box="[924,1044,1305,1328]" italics="true" pageId="28" pageNumber="29">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
species described here which has longest been known in historical collections (in BMNH, MNHN, and SNHM, at least). The first modern field specimens of this species were recognized by C. Kremen in
|
||||
|
|
@ -481,19 +481,19 @@ in
|
|||
what is now Ranomafana National Park (
|
||||
<bibRefCitation id="EFBF8BD54C6AC64B1EBF28A9FE9924E9" author="Lees" box="[159,330,1412,1437]" pageId="28" pageNumber="29" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 64</bibRefCitation>
|
||||
). Until now, recognition of this species has been taxonomically very confusing (as detailed above). Compared with closely related species, it can readily be distinguished androconially and allopatrically from
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1B592884FECE2491" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1B592884FECE2491" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C6AC64B1B592884FECE2491" italics="true" pageId="28" pageNumber="29">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
. However, as mentioned above, morphological and genetic variation across the potential full range of
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1EB728DCFECD277C" box="[151,286,1519,1544]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1EB728DCFECD277C" box="[151,286,1519,1544]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6AC64B1EB728DCFECD277C" box="[151,286,1519,1544]" italics="true" pageId="28" pageNumber="29">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
needs further examination and this is amenable to a phylogeographic analysis, to see if its circumscription might eventually comprise one or more potentially allopatric or sympatric morphospecies that also have a ventral androconial patch, ‘sp. 20’ (same as ‘sp. 77’?) known from the northeastern montane rainforests of the island (see also
|
||||
<bibRefCitation id="EFBF8BD54C6AC64B1FAB2B70FDC32700" author="Lees" box="[395,528,1628,1652]" pageId="28" pageNumber="29" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
). As mentioned above, the HT ♂ (NHRS-TOBI000000050;
|
||||
<figureCitation id="1315EAA14C6AC64B1AC42B71FAFF2700" box="[1252,1324,1628,1653]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="28" pageNumber="29">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C6AC64B1AC42B71FAFF2700" box="[1252,1324,1628,1653]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="28" pageNumber="29">Fig. 7</figureCitation>
|
||||
C) of
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1B592B70FEFD27EC" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="andrahahana">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1B592B70FEFD27EC" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="andrahahana">
|
||||
<emphasis id="B95A2A364C6AC64B1B592B70FEFD27EC" italics="true" pageId="28" pageNumber="29">Ht. andrahahana</emphasis>
|
||||
</taxonomicName>
|
||||
ab.
|
||||
|
|
@ -504,12 +504,12 @@ Aurivillius was examined in SNHM and is considered to be conspecific, but the na
|
|||
<bibRefCitation id="EFBF8BD54C6AC64B1DBF2B89FBFA27C8" author="Butler" box="[927,1065,1700,1725]" pageId="28" pageNumber="29" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(BMNH(E) 697802); now called
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1EB72BE4FE352794" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[151,486,1736,1760]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1EB72BE4FE352794" authority="Lees, 2003" authorityName="Lees" authorityYear="2003" box="[151,486,1736,1760]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C6AC64B1EB72BE4FE8E2794" box="[151,349,1737,1760]" italics="true" pageId="28" pageNumber="29">Heteropsis viettei</emphasis>
|
||||
Lees, 2003
|
||||
</taxonomicName>
|
||||
, a species potentially confused with
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1DAE2BE4FBDC2794" box="[910,1039,1735,1760]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1DAE2BE4FBDC2794" box="[910,1039,1735,1760]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6AC64B1DAE2BE4FBDC2794" box="[910,1039,1735,1760]" italics="true" pageId="28" pageNumber="29">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
in Mabille’s (1878) description of
|
||||
|
|
@ -517,7 +517,7 @@ in Mabille’s (1878) description of
|
|||
<emphasis id="B95A2A364C6AC64B1EB72BC1FE742671" box="[151,423,1772,1797]" italics="true" pageId="28" pageNumber="29">Mycalesis andravahana</emphasis>
|
||||
</taxonomicName>
|
||||
, is not conspecific and the genitalia are different (P. Viette dissection slide no. 4847). Differences in the lateral shape of the tegumen and the extent of the valve relative to the uncus tip in this slide to other preparations of
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1FBD2A18FDC02638" box="[413,531,1845,1868]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1FBD2A18FDC02638" box="[413,531,1845,1868]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="viettei">
|
||||
<emphasis id="B95A2A364C6AC64B1FBD2A18FDC02638" box="[413,531,1845,1868]" italics="true" pageId="28" pageNumber="29">Ht. viettei</emphasis>
|
||||
</taxonomicName>
|
||||
is considered to represent distortion towards two dimensions due to slide preparation, with distortion occurring most noticeably in the structures most susceptible to pressing, and not in the aedeagus or its relative length.
|
||||
|
|
@ -527,14 +527,14 @@ is considered to represent distortion towards two dimensions due to slide prepar
|
|||
: (
|
||||
<bibRefCitation id="EFBF8BD54C6AC64B1CAC2A8DFDB326A9" author="Lees" pageId="28" pageNumber="29" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="i">
|
||||
Lees, 1997: 109,
|
||||
<figureCitation id="1315EAA14C6AC64B1D752A8DFC4926CD" box="[853,922,1952,1977]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="28" pageNumber="29">
|
||||
<figureCitation id="1315EAA14C6AC64B1D752A8DFC4926CD" box="[853,922,1952,1977]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="28" pageNumber="29">
|
||||
Fig.
|
||||
<date id="FF90D0E44C6AC64B1DAB2A8DFC4926CD" box="[907,922,1952,1977]" pageId="28" pageNumber="29">7</date>
|
||||
</figureCitation>
|
||||
i, “12”, voucher 98DL,
|
||||
<figureCitation id="1315EAA14C6AC64B1A8A2A8DFB3E26CC" box="[1194,1261,1952,1977]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="28" pageNumber="29">Fig. 8</figureCitation>
|
||||
<figureCitation id="1315EAA14C6AC64B1A8A2A8DFB3E26CC" box="[1194,1261,1952,1977]" captionStart="FIGURE 8" captionStartId="26.[151,250,1929,1951]" captionTargetBox="[192,1379,232,1885]" captionTargetId="figure@26.[161,1426,193,1908]" captionTargetPageId="26" captionText="FIGURE 8. ♂ genitalia in LV (left), DV (right). A: Heteropsis cf. harveyi (see text) (Anjozorobe; 98 DL). B: Ht. vanewrighti Lees, sp. nov. (Mt. Bekolosy; 233 DL). C: Ht. imerina Lees, sp. nov. (Ankazomivady; 154 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264605" httpUri="https://zenodo.org/record/264605/files/figure.png" pageId="28" pageNumber="29">Fig. 8</figureCitation>
|
||||
A, specimen of
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1EB72AE8FE4326A9" box="[151,400,1988,2013]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6AC64B1EB72AE8FE4326A9" box="[151,400,1988,2013]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6AC64B1EB72AE8FEDC26A8" box="[151,271,1989,2012]" italics="true" pageId="28" pageNumber="29">Heteropsis</emphasis>
|
||||
cf.
|
||||
<emphasis id="B95A2A364C6AC64B1F1C2AE9FE4326A9" box="[316,400,1988,2013]" italics="true" pageId="28" pageNumber="29">harveyi</emphasis>
|
||||
|
|
@ -542,17 +542,17 @@ cf.
|
|||
from Anjozorobe
|
||||
</bibRefCitation>
|
||||
). From LV, a very shallowly domed tegumen with a rather straight dorsal edge profile and a distinctly angled brow leads to a hand-scythe shaped uncus whose ventral edge at the base is downbent as if like a ‘dewlap’ and with a rather narrow and pointed hook. Tegumen dorsal profile features a quite narrow v-shaped proximad notch and lateral profile bends round to constriction at division with vinculum, which is over half the maximum dimension of the tegumen (and thus the tegumen does not have a quadrate shape from LV). Valve with a very prominent convex dorsal shoulder and valve base very broad leading abruptly to the narrow quite tapering and sub-sinuate valve arms, which are ventrally elbowed and with only a slight indication of a ventral ‘head’ at the unexpanded club, covered with fine setae and sporting a stout spine, inrecurved from DV, where somewhat serrate on mesad edge, pointing towards the uncus tip, which falls well short of the maximum uncus extension, in fact points to the middle of the ‘hook’. Gnathos from right-angled base pointing in line with the uncus tip from LV where relatively straight blade-like, tapered to tip, recurved inward from DV. Saccus quite extended and inflated proximad, less than a third of total valve length (it is longer in the
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1A292F2CFB6B236C" box="[1033,1208,511,536]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1A292F2CFB6B236C" box="[1033,1208,511,536]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C6BC64A1A292F2CFB6B236C" box="[1033,1208,511,536]" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
specimen described below). Aedeagus particularly thin and dorsoventrally flattened, very gently uprecurved distad of the ostium, also quite extended proximad of it, distad tip quite pointed, about 1.75 x total length of valve. Juxta quite prominent proximad and rounded like a small plate from DV. A specimen referred to
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1DD42F40FBA723F1" box="[1012,1140,620,645]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1DD42F40FBA723F1" box="[1012,1140,620,645]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6BC64A1DD42F40FBA723F1" box="[1012,1140,620,645]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
from Miaranony,
|
||||
<quantity id="4CD65BC14C6BC64A1B652F41FA4F23F0" box="[1349,1436,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="29" pageNumber="30" unit="m" value="1080.0">1080 m</quantity>
|
||||
(156DL) is very similar in proportions to 98DL, with valve terminating in an inwardly pointing spine that falls short of uncus tip, but has a saccus about 1/4 longer and the hook of uncus about 1/3 deeper (from LV). A future more satisfactory resolution of genitalic variation within the
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1D672FF4FC152384" box="[839,966,727,752]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1D672FF4FC152384" box="[839,966,727,752]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6BC64A1D672FF4FC152384" box="[839,966,727,752]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
complex (currently containing four COI- 5P BIN clusters) should be based on DNA barcoded material.
|
||||
|
|
@ -568,16 +568,16 @@ locality and BMAD270-15, DL-4-963 from Ambondrombe) which is 1.52% divergent to
|
|||
. sp. 20 (exemplar BMAD213- 15 DL14A-0387, Anjanaharibe Sud, cluster number BOLD:ACW5777), 1.67% divergent to
|
||||
<emphasis id="B95A2A364C6BC64A1AE92E44FB3622F4" box="[1225,1253,873,896]" italics="true" pageId="29" pageNumber="30">Ht</emphasis>
|
||||
. sp.
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1B012E4AFA4F22F4" box="[1313,1436,871,896]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1B012E4AFA4F22F4" box="[1313,1436,871,896]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
cf.
|
||||
<emphasis id="B95A2A364C6BC64A1B682E4AFA4F22F4" box="[1352,1436,871,896]" italics="true" pageId="29" pageNumber="30">harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
(BMAD268-15, DL14AM-0025 from Anjozorobe, cluster number BOLD:ACW5778) and 1.98% divergent to
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1B592EA0FECE22BC" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1B592EA0FECE22BC" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C6BC64A1B592EA0FECE22BC" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
(BMAD064-04, Bekolosy, cluster number BOLD:AAE4111). The 357 bp cytochrome b sequence of
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1EB72EF8FECA2299" box="[151,281,980,1005]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1EB72EF8FECA2299" box="[151,281,980,1005]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6BC64A1EB72EF8FECA2299" box="[151,281,980,1005]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
(exemplars BMNH(E) 671648
|
||||
|
|
@ -587,7 +587,7 @@ cf.
|
|||
. sp. 20 (as
|
||||
<emphasis id="B95A2A364C6BC64A1A312ED4FBFE2564" box="[1041,1069,1017,1040]" italics="true" pageId="29" pageNumber="30">Ht</emphasis>
|
||||
. sp. 77; exemplar CK678 from Anjanaharibe Sud) and to
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1FE62930FD942541" box="[454,583,1052,1077]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1FE62930FD942541" box="[454,583,1052,1077]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6BC64A1FE62930FE312540" box="[454,482,1053,1076]" italics="true" pageId="29" pageNumber="30">Ht</emphasis>
|
||||
.
|
||||
<emphasis id="B95A2A364C6BC64A1FD32931FD942541" box="[499,583,1052,1077]" italics="true" pageId="29" pageNumber="30">harveyi</emphasis>
|
||||
|
|
@ -595,11 +595,11 @@ cf.
|
|||
(exemplar BMNH(E) #676467
|
||||
<accessionNumber id="947D6BC74C6BC64A1D982931FBFA2541" box="[952,1065,1052,1077]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819454" pageId="29" pageNumber="30">FJ819454</accessionNumber>
|
||||
, DL-05-385 from Ranomafana), and is about 1.56% divergent to
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1C29296CFD68252C" box="[521,699,1087,1112]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1C29296CFD68252C" box="[521,699,1087,1112]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C6BC64A1C29296CFD68252C" box="[521,699,1087,1112]" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
(exemplar DLBEK94-130, BMNH(E) #697875 from Bekolosy), whereas it is about 4.2% pairwise divergent to sequences of
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1D6F2948FBFB2508" box="[847,1064,1125,1148]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1D6F2948FBFB2508" box="[847,1064,1125,1148]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="imerina">
|
||||
<emphasis id="B95A2A364C6BC64A1D6F2948FBFB2508" box="[847,1064,1125,1148]" italics="true" pageId="29" pageNumber="30">Heteropsis imerina</emphasis>
|
||||
</taxonomicName>
|
||||
(BMNH(E) #676683
|
||||
|
|
@ -620,7 +620,7 @@ group in that study included
|
|||
<emphasis id="B95A2A364C6BC64A1F662980FD5425B0" box="[326,647,1196,1221]" italics="true" pageId="29" pageNumber="30">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1F662980FE2A25B1" box="[326,505,1196,1221]" class="Insecta" family="Nymphalidae" genus="Erebia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="passandava">Ht. passandava</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1C262980FD5125B0" box="[518,642,1197,1220]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="narova">Ht. narova</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1C262980FD5125B0" box="[518,642,1197,1220]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="narova">Ht. narova</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
‘
|
||||
|
|
@ -635,7 +635,7 @@ group in that study included
|
|||
<emphasis id="B95A2A364C6BC64A1A262981FBBE25B1" bold="true" box="[1030,1133,1196,1221]" pageId="29" pageNumber="30">stat. rev.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
] and
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1A912980FACD25B0" box="[1201,1310,1197,1220]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="strato">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1A912980FACD25B0" box="[1201,1310,1197,1220]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="strato">
|
||||
<emphasis id="B95A2A364C6BC64A1A912980FACD25B0" box="[1201,1310,1197,1220]" italics="true" pageId="29" pageNumber="30">Ht. strato</emphasis>
|
||||
</taxonomicName>
|
||||
). No COII sequence is available on GenBank (see below).
|
||||
|
|
@ -643,11 +643,11 @@ group in that study included
|
|||
<paragraph id="8B91F6244C6BC64A1EE729D9FB79277C" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="30">
|
||||
<emphasis id="B95A2A364C6BC64A1EE729D9FE342479" bold="true" box="[199,487,1268,1293]" pageId="29" pageNumber="30">Phylogeny/sister species</emphasis>
|
||||
: closely related to
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1CE829D8FCA82479" box="[712,891,1268,1293]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1CE829D8FCA82479" box="[712,891,1268,1293]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C6BC64A1CE829D8FCA82479" box="[712,891,1268,1293]" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||||
</taxonomicName>
|
||||
, and to two (?) still undescribed species in the same cryptic complex. In their cladistic analysis of COII sequences, not including
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1A692834FECE2421" authority="Torres et al., (2001: 465)" authorityName="Torres et al." authorityPageNumber="465" authorityYear="2001" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1A692834FECE2421" authority="Torres et al., (2001: 465)" authorityName="Torres et al." authorityPageNumber="465" authorityYear="2001" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vanewrighti">
|
||||
<emphasis id="B95A2A364C6BC64A1A692834FB292444" box="[1097,1274,1303,1328]" italics="true" pageId="29" pageNumber="30">Ht. vanewrighti</emphasis>
|
||||
,
|
||||
<bibRefCitation id="EFBF8BD54C6BC64A1B2B2835FF0F2420" author="Torres" pageId="29" pageNumber="30" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. & Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
|
||||
|
|
@ -658,7 +658,7 @@ Torres
|
|||
: 465)
|
||||
</taxonomicName>
|
||||
had full support for a sister relation of
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1CCC2810FCBE2421" box="[748,877,1340,1365]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1CCC2810FCBE2421" box="[748,877,1340,1365]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6BC64A1CCC2810FCBE2421" box="[748,877,1340,1365]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
(their “
|
||||
|
|
@ -674,7 +674,7 @@ Torres
|
|||
2001
|
||||
</bibRefCitation>
|
||||
[here considered to be the same as “sp. 20” from Masoala] appears from putatively fixed differences in its DNA barcodes (n=10 currently on BOLD; cluster number BOLD:ACW5777) not to be conspecific with
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1CBD28DCFCC8277C" box="[669,795,1519,1544]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1CBD28DCFCC8277C" box="[669,795,1519,1544]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6BC64A1CBD28DCFCC8277C" box="[669,795,1519,1544]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
(as also treated by
|
||||
|
|
@ -707,19 +707,19 @@ Torres
|
|||
: from Fianarantsoa/Ranomafana (including
|
||||
<typeStatus id="549548864C6BC64A1D482BC0FC4B2671" box="[872,920,1773,1797]" pageId="29" pageNumber="30">type</typeStatus>
|
||||
locality) probably at least as far north as Andasibe-Mantadia in the eastern rainforest biome, south to Ambondrombe (according to DNA barcode) and possibly as far South as Chaines Anosyennes (referred specimen) (
|
||||
<figureCitation id="1315EAA14C6BC64A1DB02A19FC372638" box="[912,996,1844,1869]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="29" pageNumber="30">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C6BC64A1DB02A19FC372638" box="[912,996,1844,1869]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="29" pageNumber="30">Fig. 30</figureCitation>
|
||||
D, lilac dots). Light brown dots in
|
||||
<figureCitation id="1315EAA14C6BC64A1B502A19FF602604" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="29" pageNumber="30">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C6BC64A1B502A19FF602604" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="29" pageNumber="30">Fig. 30</figureCitation>
|
||||
D represent localities for specimens morphologically similar to
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1DAA2A74FBD92604" box="[906,1034,1879,1904]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C6BC64A1DAA2A74FBD92604" box="[906,1034,1879,1904]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C6BC64A1DAA2A74FBD92604" box="[906,1034,1879,1904]" italics="true" pageId="29" pageNumber="30">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
falling outside the known range of this species, and where the presence of cluster BOLD:ACW5694 has not been confirmed. Populations in the more central Angavo Massif (exemplar BMAD268-15, cluster number BOLD:ACW5778, Anjozorobe) might represent a different species. However, data is currently insufficient to indicate whether the illustrated genitalia from Anjozorobe represent
|
||||
<taxonomicName id="4C2E8DA74C68C6491FB02DB4FDDD21C4" box="[400,526,151,176]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C68C6491FB02DB4FDDD21C4" box="[400,526,151,176]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C68C6491FB02DB4FDDD21C4" box="[400,526,151,176]" italics="true" pageId="30" pageNumber="31">Ht. harveyi</emphasis>
|
||||
</taxonomicName>
|
||||
or not (hence the labelling ‘
|
||||
<taxonomicName id="4C2E8DA74C68C6491D6D2DB4FC2321C4" box="[845,1008,151,176]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<taxonomicName id="4C2E8DA74C68C6491D6D2DB4FC2321C4" box="[845,1008,151,176]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="harveyi">
|
||||
<emphasis id="B95A2A364C68C6491D6D2DB4FCBA21C4" box="[845,873,153,176]" italics="true" pageId="30" pageNumber="31">Ht</emphasis>
|
||||
. cf.
|
||||
<emphasis id="B95A2A364C68C6491DBC2DBAFC2321C4" box="[924,1008,151,176]" italics="true" pageId="30" pageNumber="31">harveyi</emphasis>
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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|
||||
<treatment id="038747324C78C6471EB729E2FC722384" ID-DOI="http://doi.org/10.5281/zenodo.6086412" ID-GBIF-Taxon="120817077" ID-Zenodo-Dep="6086412" LSID="urn:lsid:plazi:treatment:038747324C78C6471EB729E2FC722384" httpUri="http://treatment.plazi.org/id/038747324C78C6471EB729E2FC722384" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="15">
|
||||
<treatment id="038747324C78C6471EB729E2FC722384" ID-DOI="http://doi.org/10.5281/zenodo.6086412" ID-GBIF-Taxon="120817077" ID-Zenodo-Dep="6086412" LSID="urn:lsid:plazi:treatment:038747324C78C6471EB729E2FC722384" httpUri="http://treatment.plazi.org/id/038747324C78C6471EB729E2FC722384" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="15" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C78C6471EB729E2FC722384" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="nomenclature">
|
||||
<paragraph id="8B91F6244C78C6591EB729E2FE34259E" blockId="14.[151,487,1231,1258]" box="[151,487,1231,1258]" pageId="14" pageNumber="15">
|
||||
<heading id="D0D941484C78C6591EB729E2FE34259E" bold="true" box="[151,487,1231,1258]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
|
||||
<emphasis id="B95A2A364C78C6591EB729E2FE34259E" bold="true" box="[151,487,1231,1258]" pageId="14" pageNumber="15">
|
||||
<taxonomicName id="4C2E8DA74C78C6591EB729E2FEA5259D" box="[151,374,1231,1257]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C78C6591EB729E2FEA5259D" ID-CoL="953DN" box="[151,374,1231,1257]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C78C6591EB729E2FEA5259D" bold="true" box="[151,374,1231,1257]" italics="true" pageId="14" pageNumber="15">Gallienia alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
problem
|
||||
|
|
@ -50,9 +50,9 @@ problem
|
|||
</paragraph>
|
||||
<paragraph id="8B91F6244C78C6471EB7283BFC722384" blockId="14.[151,1436,1302,2014]" lastBlockId="16.[151,1437,151,752]" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
|
||||
The left image (#3043) in
|
||||
<figureCitation id="1315EAA14C78C6591F9E283BFE2D245B" box="[446,510,1302,1327]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C78C6591F9E283BFE2D245B" box="[446,510,1302,1327]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
A represents the specimen (BMNH(E) 672621) that P. Viette intended to represent the LT of
|
||||
<taxonomicName id="4C2E8DA74C78C6591F282817FD402427" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[264,659,1338,1363]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C78C6591F282817FD402427" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[264,659,1338,1363]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C78C6591F282817FE0E2427" box="[264,477,1338,1363]" italics="true" pageId="14" pageNumber="15">Gallienia alaokola</emphasis>
|
||||
<bibRefCitation id="EFBF8BD54C78C6591FC42817FD402427" author="Oberthur" box="[484,659,1338,1363]" pageId="14" pageNumber="15" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
</taxonomicName>
|
||||
|
|
@ -63,11 +63,11 @@ A represents the specimen (BMNH(E) 672621) that P. Viette intended to represent
|
|||
) illustrated as [a ♀ of] ‘
|
||||
<emphasis id="B95A2A364C78C6591ECA28A9FEDA24EF" box="[234,265,1412,1435]" italics="true" pageId="14" pageNumber="15">M.</emphasis>
|
||||
?
|
||||
<taxonomicName id="4C2E8DA74C78C6591F0828AFFE5924EF" box="[296,394,1410,1435]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C78C6591F0828AFFE5924EF" box="[296,394,1410,1435]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C78C6591F0828AFFE5924EF" box="[296,394,1410,1435]" italics="true" pageId="14" pageNumber="15">alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
’. I contend, considering dorsal colouration, ocellus expression in space-M3 and relatively more pointed HW tails, that both specimens in fact represent
|
||||
<taxonomicName id="4C2E8DA74C78C6591D6F288AFC0524CB" box="[847,982,1446,1471]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mabillei">
|
||||
<taxonomicName id="4C2E8DA74C78C6591D6F288AFC0524CB" box="[847,982,1446,1471]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mabillei">
|
||||
<emphasis id="B95A2A364C78C6591D6F288AFC0524CB" box="[847,982,1446,1471]" italics="true" pageId="14" pageNumber="15">Ht. mabillei</emphasis>
|
||||
</taxonomicName>
|
||||
(both surfaces of the HT ♂ of
|
||||
|
|
@ -76,11 +76,11 @@ A represents the specimen (BMNH(E) 672621) that P. Viette intended to represent
|
|||
<bibRefCitation id="EFBF8BD54C78C6591EDC28E7FE582497" author="Butler" box="[252,395,1482,1507]" pageId="14" pageNumber="15" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, BMNH(E) 672624, are figured in
|
||||
<figureCitation id="1315EAA14C78C6591D0028E7FCB72497" box="[800,868,1482,1507]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C78C6591D0028E7FCB72497" box="[800,868,1482,1507]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
C). Moreover, although
|
||||
<bibRefCitation id="EFBF8BD54C78C6591A5828E7FAF82497" author="d'Abrera" box="[1144,1323,1482,1507]" pageId="14" pageNumber="15" refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp." type="book" year="1980">d’Abrera (1980</bibRefCitation>
|
||||
: 190) had placed a red label next to photograph of “
|
||||
<taxonomicName id="4C2E8DA74C78C6591C4C28C2FD2F2773" box="[620,764,1518,1543]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C78C6591C4C28C2FD2F2773" box="[620,764,1518,1543]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C78C6591C4C28C2FD2F2773" box="[620,764,1518,1543]" italics="true" pageId="14" pageNumber="15">M. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
”, the accompanying text says “♀ (?
|
||||
|
|
@ -94,7 +94,7 @@ would be extremely unsatisfactory in this difficult-to-distinguish species group
|
|||
, considering in particular his qualification of use of the word 'type' by a question mark (Art. 74.5). A proportion of Oberthür’s
|
||||
<typeStatus id="549548864C78C6591DA92B8EFC3327CF" box="[905,992,1699,1723]" pageId="14" pageNumber="15" type="syntype">syntype</typeStatus>
|
||||
specimens of
|
||||
<taxonomicName id="4C2E8DA74C78C6591AA72B8FFA8F27CF" box="[1159,1372,1698,1723]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C78C6591AA72B8FFA8F27CF" box="[1159,1372,1698,1723]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C78C6591AA72B8FFA8F27CF" box="[1159,1372,1698,1723]" italics="true" pageId="14" pageNumber="15">Gallienia alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
from Fianarantsoa and some also from Antsianaka (the current region around RNI Zahamena) have apparently generally shorter HW tails than in the HT of
|
||||
|
|
@ -103,21 +103,21 @@ from Fianarantsoa and some also from Antsianaka (the current region around RNI Z
|
|||
Butler
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C78C6591D682BC7FC592677" box="[840,906,1770,1795]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C78C6591D682BC7FC592677" box="[840,906,1770,1795]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
C) and a much darker upperside. They lack the ocellus expression in both space-M3 and CuA1 of the ♂ HW upperside, with sometimes neither ocellus expressed (this variation in his mixed series is alluded to in Oberthür’s 1916: 205–206 description). Among Oberthür’s 75
|
||||
<typeStatus id="549548864C78C6591EB72A7AFF2F261B" box="[151,252,1879,1903]" pageId="14" pageNumber="15" type="syntype">syntypes</typeStatus>
|
||||
, a LT ♂ specimen of
|
||||
<taxonomicName id="4C2E8DA74C78C6591FC92A7BFD68261B" box="[489,699,1878,1903]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C78C6591FC92A7BFD68261B" box="[489,699,1878,1903]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C78C6591FC92A7BFD68261B" box="[489,699,1878,1903]" italics="true" pageId="14" pageNumber="15">Gallienia alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
is here therefore chosen to resolve the problem. This LT ♂ is here designated as the specimen (
|
||||
<figureCitation id="1315EAA14C78C6591FC22A50FDF526E2" box="[482,550,1917,1942]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C78C6591FC22A50FDF526E2" box="[482,550,1917,1942]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||||
B) bearing data labels “
|
||||
<collectingCountry id="F339B6B44C78C6591D182A50FC1726E2" box="[824,964,1917,1942]" name="Madagascar" pageId="14" pageNumber="15">Madagascar</collectingCountry>
|
||||
, Fianarantsoa, Perrot Frères, 2e Semestre 1892|Ex Oberthür Coll. Brit. Mus. 1927-3|Photographed by B. d'Abrera 77/78|BMNH(E) 1717102 [
|
||||
<collectingCountry id="F339B6B44C78C6591B252A8FFA9326CE" box="[1285,1344,1954,1978]" name="Qatar" pageId="14" pageNumber="15">QTR</collectingCountry>
|
||||
label]”. This is also the “
|
||||
<taxonomicName id="4C2E8DA74C78C6591F482AEBFE1E26A9" box="[360,461,1990,2013]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="18" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<taxonomicName id="4C2E8DA74C78C6591F482AEBFE1E26A9" box="[360,461,1990,2013]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="18" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<emphasis id="B95A2A364C78C6591F482AEBFE1E26A9" box="[360,461,1990,2013]" italics="true" pageId="14" pageNumber="15">Masoura</emphasis>
|
||||
</taxonomicName>
|
||||
? sp.” illustrated by
|
||||
|
|
@ -125,7 +125,7 @@ label]”. This is also the “
|
|||
: 229) in DV/VV, following preference for a previously illustrated specimen (Recommendation 74B of the code), although in this case, not the one that the original author intended. This action is also in the spirit of the Code as it promotes stability and obviates the need to describe a new species within Oberthür’s syntypic series. The majority of the series of
|
||||
<bibRefCitation id="EFBF8BD54C79C6581A8A2ABDFAB926DD" author="Oberthur" box="[1194,1386,1936,1961]" pageId="15" pageNumber="16" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür (1916)</bibRefCitation>
|
||||
’s
|
||||
<taxonomicName id="4C2E8DA74C79C6581BA42ABCFF2A26BA" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C79C6581BA42ABCFF2A26BA" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C79C6581BA42ABCFF2A26BA" italics="true" pageId="15" pageNumber="16">G. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
STs from “Fianarantsoa” belong to
|
||||
|
|
@ -143,13 +143,13 @@ STs from “Fianarantsoa” belong to
|
|||
(see
|
||||
<bibRefCitation id="EFBF8BD54C66C6471DE32C28FB902068" author="Lees" box="[963,1091,260,284]" pageId="16" pageNumber="17" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
: 99) and also, more markedly,
|
||||
<taxonomicName id="4C2E8DA74C66C6471EB72C04FEF42034" box="[151,295,297,320]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<taxonomicName id="4C2E8DA74C66C6471EB72C04FEF42034" box="[151,295,297,320]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<emphasis id="B95A2A364C66C6471EB72C04FEF42034" box="[151,295,297,320]" italics="true" pageId="16" pageNumber="17">Ht. masoura</emphasis>
|
||||
</taxonomicName>
|
||||
. It (“sp. 56”;
|
||||
<bibRefCitation id="EFBF8BD54C66C6471FF22C05FD862034" author="Lees" box="[466,597,296,320]" pageId="16" pageNumber="17" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
: 105) also has multiply toothed long valves, which do not have such a projected sharply inbent tip, as does the genitalia of Oberthür’s ♂ ‘iconotype’ of
|
||||
<taxonomicName id="4C2E8DA74C66C6471A0C2C60FB7D2011" box="[1068,1198,332,357]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471A0C2C60FB7D2011" box="[1068,1198,332,357]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C66C6471A0C2C60FB7D2011" box="[1068,1198,332,357]" italics="true" pageId="16" pageNumber="17">G. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
(=
|
||||
|
|
@ -157,7 +157,7 @@ STs from “Fianarantsoa” belong to
|
|||
<emphasis id="B95A2A364C66C6471AEE2C60FA862011" box="[1230,1365,332,357]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||||
</taxonomicName>
|
||||
) (
|
||||
<figureCitation id="1315EAA14C66C6471B4E2C61FF7620FC" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C66C6471B4E2C61FF7620FC" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||||
D). This feature is not in fact evident in the HT specimen of
|
||||
<taxonomicName id="4C2E8DA74C66C6471D7F2C42FBAB20FC" authority="Butler" authorityName="Butler" box="[863,1144,367,392]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||||
<emphasis id="B95A2A364C66C6471D7F2C42FBFB20FC" box="[863,1064,367,392]" italics="true" pageId="16" pageNumber="17">Strabena mabillei</emphasis>
|
||||
|
|
@ -168,7 +168,7 @@ Butler
|
|||
<emphasis id="B95A2A364C66C6471ABD2C94FAF920A4" box="[1181,1322,439,464]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||||
</taxonomicName>
|
||||
(in fresh specimens,
|
||||
<taxonomicName id="4C2E8DA74C66C6471F3E2CF0FE7C2081" box="[286,431,476,501]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471F3E2CF0FE7C2081" box="[286,431,476,501]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C66C6471F3E2CF0FE7C2081" box="[286,431,476,501]" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
, as here fixed, also has a darker dorsal colouration). Note the androconial scales on abdominal sternum third segment and the inwards recurved valve tips characteristic of
|
||||
|
|
@ -176,25 +176,25 @@ Butler
|
|||
<emphasis id="B95A2A364C66C6471A532F2CFB29236C" box="[1139,1274,511,536]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C66C6471B2A2CD2FA9E236C" box="[1290,1357,511,536]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C66C6471B2A2CD2FA9E236C" box="[1290,1357,511,536]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||||
D) and also (
|
||||
<figureCitation id="1315EAA14C66C6471EF42F09FEC62348" box="[212,277,548,573]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C66C6471EF42F09FEC62348" box="[212,277,548,573]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||||
A,C) that
|
||||
<taxonomicName id="4C2E8DA74C66C6471FA62F08FDDF2349" box="[390,524,548,573]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||||
<emphasis id="B95A2A364C66C6471FA62F08FDDF2349" box="[390,524,548,573]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||||
</taxonomicName>
|
||||
has relatively long tails especially at M3, a lighter brown dorsal colouration, and three or four white stripes on the FWV tending to be more strongly represented as marks on the FWD costa, than the species/specimen here selected to represent
|
||||
<taxonomicName id="4C2E8DA74C66C6471C952F40FC9023F1" box="[693,835,620,645]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471C952F40FC9023F1" box="[693,835,620,645]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C66C6471C952F40FC9023F1" box="[693,835,620,645]" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C66C6471D742F41FC4423F0" box="[852,919,620,645]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||||
<figureCitation id="1315EAA14C66C6471D742F41FC4423F0" box="[852,919,620,645]" captionStart="FIGURE 3" captionStartId="15.[151,250,1650,1672]" captionTargetBox="[303,1292,195,1608]" captionTargetId="figure@15.[275,1307,177,1626]" captionTargetPageId="15" captionText="FIGURE 3. Adults in DV (left), VV (right) unless otherwise stated. A: depictions (‘ iconotypes’) of Gallienia alaokola Oberthür (1916) ♂, left (VV) and ♀, right (DV), representing Oberthür’s preference for the type specimens of this nominal species, both referred here to Ht. mabillei (Butler, 1879). B: designated LT ♂ of G. alaokola Oberthür, 1916 (‘ Fianarantsoa’; BMNH (E) 1717102). C: HT ♂ of Strabena mabillei Butler, 1879 (BMNH (E) 672624). Scale bar: 30 mm. D: ♂ abdomen of Oberthür collection specimen of Ht. mabillei (left, VV). E: PLT ♂ abdomen of G. alaokola (Fianarantsoa; BMNH (E) 672621) (right, VV)." figureDoi="http://doi.org/10.5281/zenodo.264600" httpUri="https://zenodo.org/record/264600/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||||
B). It may be noted that samples respectively representing
|
||||
<taxonomicName id="4C2E8DA74C66C6471F092FBCFE6323DC" box="[297,432,655,680]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mabillei">
|
||||
<emphasis id="B95A2A364C66C6471F092FBCFE6323DC" box="[297,432,655,680]" italics="true" pageId="16" pageNumber="17">Ht. mabillei</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C66C6471FC72FBCFDA023DC" box="[487,627,655,680]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C66C6471FC72FBCFDA023DC" box="[487,627,655,680]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C66C6471FC72FBCFDA023DC" box="[487,627,655,680]" italics="true" pageId="16" pageNumber="17">Ht. alaokola</emphasis>
|
||||
</taxonomicName>
|
||||
(KAP524 [KA-P524] = DL-4-233, Anjozorobe; KAP537 [KA-P537] = DL-02-20, Makira, in the study of Aduse-Poku
|
||||
|
|
|
|||
|
|
@ -1,51 +1,51 @@
|
|||
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<mods:title id="59BBF3558881CAAF9B667E42AEBA128E">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
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<mods:title id="341B2EA9CC223BC5612A6D93D846422B">Zootaxa</mods:title>
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<treatment id="038747324C7BC6591EB728FCFDFA2509" ID-DOI="http://doi.org/10.5281/zenodo.6086410" ID-GBIF-Taxon="120817095" ID-Zenodo-Dep="6086410" LSID="urn:lsid:plazi:treatment:038747324C7BC6591EB728FCFDFA2509" httpUri="http://treatment.plazi.org/id/038747324C7BC6591EB728FCFDFA2509" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<treatment id="038747324C7BC6591EB728FCFDFA2509" ID-DOI="http://doi.org/10.5281/zenodo.6086410" ID-GBIF-Taxon="120817095" ID-Zenodo-Dep="6086410" LSID="urn:lsid:plazi:treatment:038747324C7BC6591EB728FCFDFA2509" httpUri="http://treatment.plazi.org/id/038747324C7BC6591EB728FCFDFA2509" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
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<subSubSection id="C334A5AF4C7BC6591EB728FCFB8B2349" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="nomenclature">
|
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<paragraph id="8B91F6244C7BC65A1EB728FCFBBC249F" blockId="13.[151,1135,1489,1515]" box="[151,1135,1489,1515]" pageId="13" pageNumber="14">
|
||||
<heading id="D0D941484C7BC65A1EB728FCFBBC249F" bold="true" box="[151,1135,1489,1515]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
|
||||
<emphasis id="B95A2A364C7BC65A1EB728FCFBBC249F" bold="true" box="[151,1135,1489,1515]" pageId="13" pageNumber="14">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1EB728FCFE59249F" box="[151,394,1489,1515]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="antahala">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1EB728FCFE59249F" box="[151,394,1489,1515]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="antahala">
|
||||
<emphasis id="B95A2A364C7BC65A1EB728FCFE59249F" bold="true" box="[151,394,1489,1515]" italics="true" pageId="13" pageNumber="14">Heteropsis antahala</emphasis>
|
||||
</taxonomicName>
|
||||
group (subgenera
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C5428FCFD32249F" box="[628,737,1489,1515]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="18" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C5428FCFD32249F" box="[628,737,1489,1515]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="18" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<emphasis id="B95A2A364C7BC65A1C5428FCFD32249F" bold="true" box="[628,737,1489,1515]" italics="true" pageId="13" pageNumber="14">Masoura</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
|
|
@ -59,7 +59,7 @@ and
|
|||
</paragraph>
|
||||
<paragraph id="8B91F6244C7BC6591EB72B34FB8B2349" blockId="13.[151,1437,1560,2017]" lastBlockId="14.[151,1437,151,1149]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||||
A new species is described here in the
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C472B34FD2A2745" box="[615,761,1560,1585]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="antahala">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C472B34FD2A2745" box="[615,761,1560,1585]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="antahala">
|
||||
<emphasis id="B95A2A364C7BC65A1C472B34FD2A2745" box="[615,761,1560,1585]" italics="true" pageId="13" pageNumber="14">Ht. antahala</emphasis>
|
||||
</taxonomicName>
|
||||
group, and so it is convenient to characterise its nominal species. The LT ♂ of
|
||||
|
|
@ -68,7 +68,7 @@ group, and so it is convenient to characterise its nominal species. The LT ♂ o
|
|||
Ward, 1872
|
||||
</taxonomicName>
|
||||
is here designated as the specimen (
|
||||
<figureCitation id="1315EAA14C7BC65A1A892B11FB3E2721" box="[1193,1261,1596,1621]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="13" pageNumber="14">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7BC65A1A892B11FB3E2721" box="[1193,1261,1596,1621]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="13" pageNumber="14">Fig. 2</figureCitation>
|
||||
F) bearing data “
|
||||
<typeStatus id="549548864C7BC65A1E812B4DFEC6270D" box="[161,277,1632,1657]" pageId="13" pageNumber="14" type="lectotype">Lectotype</typeStatus>
|
||||
|
|
||||
|
|
@ -99,7 +99,7 @@ as well as the more strongly HWtailed mycalesines traditionally placed in the su
|
|||
<bibRefCitation id="EFBF8BD54C7BC65A1D5D2A39FBFD2659" author="Hemming" box="[893,1070,1812,1837]" pageId="13" pageNumber="14" refString="Hemming, F. (1964) Annotationes Lepidopterologicae. Part 4. Hepburn & Sons, London, 180 pp." type="book" year="1964">Hemming 1964</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(see below). Here, the status of
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1EB72A14FE362625" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[151,485,1848,1873]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1EB72A14FE362625" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[151,485,1848,1873]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="alaokola">
|
||||
<emphasis id="B95A2A364C7BC65A1EB72A14FEF12625" box="[151,290,1848,1873]" italics="true" pageId="13" pageNumber="14">Ht. alaokola</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C7BC65A1F102A15FE0E2625" author="Oberthur" box="[304,477,1848,1873]" pageId="13" pageNumber="14" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
|
||||
|
|
@ -118,10 +118,10 @@ group (
|
|||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B95A2A364C7BC65A1C822A70FCE52601" box="[674,822,1884,1909]" italics="true" pageId="13" pageNumber="14">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C822A70FCE12601" box="[674,818,1884,1909]" class="Insecta" family="Tettigoniidae" genus="Gallienia" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="alaokola">Ht. alaokola</taxonomicName>
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C822A70FCE12601" box="[674,818,1884,1909]" class="Insecta" family="Nymphalidae" genus="Gallienia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="alaokola">Ht. alaokola</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1D1F2A70FBBC2601" authority="Butler, 1879" authorityName="Butler" authorityYear="1879" box="[831,1135,1884,1909]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mabillei">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1D1F2A70FBBC2601" authority="Butler, 1879" authorityName="Butler" authorityYear="1879" box="[831,1135,1884,1909]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mabillei">
|
||||
<emphasis id="B95A2A364C7BC65A1D1F2A70FC152601" box="[831,966,1884,1909]" italics="true" pageId="13" pageNumber="14">Ht. mabillei</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C7BC65A1DF62A71FBBB2601" author="Butler" box="[982,1128,1884,1909]" pageId="13" pageNumber="14" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4, 227 - 246. http: // dx. doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
|
||||
|
|
@ -140,7 +140,7 @@ in
|
|||
<emphasis id="B95A2A364C78C6591CF92DB4FCBB21C4" box="[729,872,151,176]" italics="true" pageId="14" pageNumber="15">Ht. antahala</emphasis>
|
||||
</taxonomicName>
|
||||
group; see also
|
||||
<taxonomicName id="4C2E8DA74C78C6591A3F2DB4FB6921C4" box="[1055,1210,151,176]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<taxonomicName id="4C2E8DA74C78C6591A3F2DB4FB6921C4" box="[1055,1210,151,176]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="subsimilis">
|
||||
<emphasis id="B95A2A364C78C6591A3F2DB4FB6921C4" box="[1055,1210,151,176]" italics="true" pageId="14" pageNumber="15">Ht. subsimilis</emphasis>
|
||||
</taxonomicName>
|
||||
group, in respect of possible breakdown in pre-mating isolating system. It may be necessary to inspect androconial features of ♂♂ closely to assist in identification. This group are canopy specialists attracted to ripe fruit as adults (sometimes feeding on fruit
|
||||
|
|
@ -152,7 +152,7 @@ group, in respect of possible breakdown in pre-mating isolating system. It may b
|
|||
) observed to roost gregariously as adults (
|
||||
<bibRefCitation id="EFBF8BD54C78C6591CA12C61FCD52010" author="Lees" box="[641,774,332,356]" pageId="14" pageNumber="15" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
). The ♂♂ indulge in vigorous hilltopping behaviour on ridgetops, sitting atop bushes just like
|
||||
<taxonomicName id="4C2E8DA74C78C6591C6E2C5CFD0520FC" box="[590,726,367,392]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C78C6591C6E2C5CFD0520FC" box="[590,726,367,392]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C78C6591C6E2C5CFD0520FC" box="[590,726,367,392]" italics="true" pageId="14" pageNumber="15">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
, and making canopy chases. Unfortunately there are as yet no biological observations on the large and obvious monomorphic
|
||||
|
|
@ -160,7 +160,7 @@ group, in respect of possible breakdown in pre-mating isolating system. It may b
|
|||
<emphasis id="B95A2A364C78C6591D412CB9FC1D20D9" box="[865,974,404,429]" italics="true" pageId="14" pageNumber="15">Mylothris</emphasis>
|
||||
</taxonomicName>
|
||||
mimic,
|
||||
<taxonomicName id="4C2E8DA74C78C6591A0B2CB8FB6420D8" box="[1067,1207,405,428]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<taxonomicName id="4C2E8DA74C78C6591A0B2CB8FB6420D8" box="[1067,1207,405,428]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<emphasis id="B95A2A364C78C6591A0B2CB8FB6420D8" box="[1067,1207,405,428]" italics="true" pageId="14" pageNumber="15">Ht. masoura</emphasis>
|
||||
</taxonomicName>
|
||||
; its female is one of the largest mycalesines globally (around
|
||||
|
|
@ -195,7 +195,7 @@ and four sampled members of the
|
|||
group: Aduse-Poku
|
||||
<emphasis id="B95A2A364C78C6591C372F40FD8123F1" box="[535,594,620,645]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
|
||||
, 2015,
|
||||
<figureCitation id="1315EAA14C78C6591C8A2F41FD3C23F0" box="[682,751,620,645]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C78C6591C8A2F41FD3C23F0" box="[682,751,620,645]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
|
||||
; which was submarginally supported (pp=0.91) for the two exemplars in
|
||||
<bibRefCitation id="EFBF8BD54C78C6591F122FA2FD5223DC" author="Kodandaramaiah" box="[306,641,655,680]" pageId="14" pageNumber="15" refString="Kodandaramaiah, U., Lees, D. C., Muller, C. J., Torres, E., Karanth, C. P. & Wahlberg, N. (2010) Phylogenetics and biogeography of a spectacular Old World radiation of butterflies: the subtribe Mycalesina (Lepidoptera: Nymphalidae: Satyrini). BMC Evolutionary Biology, 10, 172. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 10 - 172" type="journal article" year="2010">
|
||||
Kodandaramaiah
|
||||
|
|
@ -207,17 +207,17 @@ Kodandaramaiah
|
|||
the ♂ HW (
|
||||
<bibRefCitation id="EFBF8BD54C78C6591A092F99FB0023B9" author="Lees" box="[1065,1235,692,717]" pageId="14" pageNumber="15" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997: 76</bibRefCitation>
|
||||
) and overlying at least, vein 1A, a character which is only sporadically seen elsewhere, e.g. in the African ‘
|
||||
<taxonomicName id="4C2E8DA74C78C6591A5D2FF4FB262384" box="[1149,1269,729,752]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C78C6591A5D2FF4FB262384" box="[1149,1269,729,752]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C78C6591A5D2FF4FB262384" box="[1149,1269,729,752]" italics="true" pageId="14" pageNumber="15">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
’
|
||||
<emphasis id="B95A2A364C78C6591B252FFAFA982384" box="[1285,1355,727,752]" italics="true" pageId="14" pageNumber="15">peitho</emphasis>
|
||||
(Plötz, 1880) group and in
|
||||
<taxonomicName id="4C2E8DA74C78C6591F512FD0FE242261" box="[369,503,764,789]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<taxonomicName id="4C2E8DA74C78C6591F512FD0FE242261" box="[369,503,764,789]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="drepana">
|
||||
<emphasis id="B95A2A364C78C6591F512FD0FE242261" box="[369,503,764,789]" italics="true" pageId="14" pageNumber="15">Ht. drepana</emphasis>
|
||||
</taxonomicName>
|
||||
and (2) the ocelli in hwv space M2, CuA1 and particularly CuA2, where the ventral pattern is pronounced (i.e. not in
|
||||
<taxonomicName id="4C2E8DA74C78C6591C2B2E0CFD44224C" box="[523,663,801,824]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<taxonomicName id="4C2E8DA74C78C6591C2B2E0CFD44224C" box="[523,663,801,824]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<emphasis id="B95A2A364C78C6591C2B2E0CFD44224C" box="[523,663,801,824]" italics="true" pageId="14" pageNumber="15">Ht. masoura</emphasis>
|
||||
</taxonomicName>
|
||||
, but in
|
||||
|
|
@ -229,11 +229,11 @@ and (2) the ocelli in hwv space M2, CuA1 and particularly CuA2, where the ventra
|
|||
<emphasis id="B95A2A364C78C6591B592E44FED122D1" italics="true" pageId="14" pageNumber="15">Ht. paradoxa</emphasis>
|
||||
</taxonomicName>
|
||||
, except in
|
||||
<taxonomicName id="4C2E8DA74C78C6591FA22EA0FDC322D0" box="[386,528,909,932]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<taxonomicName id="4C2E8DA74C78C6591FA22EA0FDC322D0" box="[386,528,909,932]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="masoura">
|
||||
<emphasis id="B95A2A364C78C6591FA22EA0FDC322D0" box="[386,528,909,932]" italics="true" pageId="14" pageNumber="15">Ht. masoura</emphasis>
|
||||
</taxonomicName>
|
||||
, which has the upper surface suffused with white. This FWD trend can also be seen in ♀♀ of some
|
||||
<taxonomicName id="4C2E8DA74C78C6591FA02E9CFE2B22BC" box="[384,504,945,968]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C78C6591FA02E9CFE2B22BC" box="[384,504,945,968]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C78C6591FA02E9CFE2B22BC" box="[384,504,945,968]" italics="true" pageId="14" pageNumber="15">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
(e.g.
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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<treatment id="038747324C7CC65A1EB72934FCB3240A" ID-DOI="http://doi.org/10.5281/zenodo.6086408" ID-GBIF-Taxon="120817094" ID-Zenodo-Dep="6086408" LSID="urn:lsid:plazi:treatment:038747324C7CC65A1EB72934FCB3240A" httpUri="http://treatment.plazi.org/id/038747324C7CC65A1EB72934FCB3240A" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
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<taxonomicName id="4C2E8DA74C7CC65D1EB72934FDA72547" authority="Lees and Kremen" authorityName="Lees and Kremen" box="[151,628,1049,1075]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimetica" status="sp. nov.">
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<emphasis id="B95A2A364C7CC65D1EB72934FE5D2547" bold="true" box="[151,398,1049,1075]" italics="true" pageId="10" pageNumber="11">Heteropsis mimetica</emphasis>
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Lees and Kremen
|
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</taxonomicName>
|
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|
|
@ -58,7 +58,7 @@ Lees and Kremen
|
|||
: “sp. 72” (
|
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<bibRefCitation id="EFBF8BD54C7CC65D1FF129A9FCD625E8" author="Lees" box="[465,773,1156,1181]" pageId="10" pageNumber="11" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
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||||
Lees 1997: 56,
|
||||
<figureCitation id="1315EAA14C7CC65D1C5F29A9FD6C25E8" box="[639,703,1156,1181]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
|
||||
<figureCitation id="1315EAA14C7CC65D1C5F29A9FD6C25E8" box="[639,703,1156,1181]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." figureDoi="http://doi.org/10.5281/zenodo.264604" httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
|
||||
G, 7H
|
||||
</bibRefCitation>
|
||||
).
|
||||
|
|
@ -74,7 +74,7 @@ material., Deposition
|
|||
<typeStatus id="549548864C7CC65D1C542987FD3225B7" box="[628,737,1194,1219]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: ♂ (
|
||||
<figureCitation id="1315EAA14C7CC65D1D362987FC8925B7" box="[790,858,1194,1219]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7CC65D1D362987FC8925B7" box="[790,858,1194,1219]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
|
||||
A),
|
||||
<collectingCountry id="F339B6B44C7CC65D1DA52987FBDF25B7" box="[901,1036,1194,1219]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
|
||||
NE, Anjanaharibe Sud,
|
||||
|
|
@ -170,7 +170,7 @@ voucher];
|
|||
(E) 1717112 [
|
||||
<collectingCountry id="F339B6B44C7CC65D1B4128DFFA4F277E" box="[1377,1436,1522,1546]" name="Qatar" pageId="10" pageNumber="11">QTR</collectingCountry>
|
||||
barcode]; ♀ (
|
||||
<figureCitation id="1315EAA14C7CC65D1F0C2B38FEBC275A" box="[300,367,1557,1582]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7CC65D1F0C2B38FEBC275A" box="[300,367,1557,1582]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
|
||||
B), data as HT but: CK517,
|
||||
<collectionCode id="ED3F6EE14C7CC65D1C892B3BFD35275A" box="[681,742,1558,1582]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="10" pageNumber="11">DNA</collectionCode>
|
||||
[
|
||||
|
|
@ -233,7 +233,7 @@ NE, Anjanaharibe Sud,
|
|||
<paragraph id="8B91F6244C7CC65D1EE72A39FDBF26C8" blockId="10.[151,1437,1156,1981]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B95A2A364C7CC65D1EE72A39FE922659" bold="true" box="[199,321,1812,1837]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
|
||||
The combination in
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1C0C2A38FD6E2658" box="[556,701,1813,1836]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1C0C2A38FD6E2658" box="[556,701,1813,1836]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<emphasis id="B95A2A364C7CC65D1C0C2A38FD6E2658" box="[556,701,1813,1836]" italics="true" pageId="10" pageNumber="11">Ht. mimetica</emphasis>
|
||||
</taxonomicName>
|
||||
of the space-M3 as well as space-CuA1 ocellus expressed on the HWD and a white-translucent ocellus ring (hereafter ‘Orng’) extended close to the base of space M3 and CuA
|
||||
|
|
@ -243,23 +243,23 @@ the FWD, forming a rectangle, as in some forms of
|
|||
<emphasis id="B95A2A364C7CC65D1CDE2A70FC522601" box="[766,897,1884,1909]" italics="true" pageId="10" pageNumber="11">Ht. turbans</emphasis>
|
||||
</taxonomicName>
|
||||
, is diagnostic for this species for both known populations. In the closely related allospecies
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1C882AACFCF626EC" box="[680,805,1921,1944]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1C882AACFCF626EC" box="[680,805,1921,1944]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<emphasis id="B95A2A364C7CC65D1C882AACFCF626EC" box="[680,805,1921,1944]" italics="true" pageId="10" pageNumber="11">Ht. cowani</emphasis>
|
||||
</taxonomicName>
|
||||
, the Orng is nearly circular for all known populations. This trait is consistent in all
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1FF42A88FDB626C8" box="[468,613,1957,1980]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1FF42A88FDB626C8" box="[468,613,1957,1980]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<emphasis id="B95A2A364C7CC65D1FF42A88FDB626C8" box="[468,613,1957,1980]" italics="true" pageId="10" pageNumber="11">Ht. mimetica</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF51A6AC4C7DC65C1EB72A61FC8126A9" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="11" pageNumber="12" targetBox="[266,1318,198,1839]" targetPageId="11">
|
||||
<caption id="DF51A6AC4C7DC65C1EB72A61FC8126A9" ID-DOI="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="11" pageNumber="12" targetBox="[266,1318,198,1839]" targetPageId="11">
|
||||
<paragraph id="8B91F6244C7DC65C1EB72A61FC8126A9" blockId="11.[151,1436,1868,2013]" pageId="11" pageNumber="12">
|
||||
<emphasis id="B95A2A364C7DC65C1EB72A61FEC72615" bold="true" box="[151,276,1868,1890]" pageId="11" pageNumber="12">FIGURE 2.</emphasis>
|
||||
Adults in DV (left), VV (right).
|
||||
<emphasis id="B95A2A364C7DC65C1C4C2A61FDAD2615" bold="true" box="[620,638,1868,1889]" pageId="11" pageNumber="12">A</emphasis>
|
||||
: HT ♂ of
|
||||
<taxonomicName id="4C2E8DA74C7DC65C1CCE2A60FBBC2616" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[750,1135,1868,1890]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mimetica" status="sp. nov.">
|
||||
<taxonomicName id="4C2E8DA74C7DC65C1CCE2A60FBBC2616" authority="Lees & Kremen" authorityName="Lees & Kremen" box="[750,1135,1868,1890]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mimetica" status="sp. nov.">
|
||||
<emphasis id="B95A2A364C7DC65C1CCE2A60FC132616" box="[750,960,1869,1890]" italics="true" pageId="11" pageNumber="12">Heteropsis mimetica</emphasis>
|
||||
Lees & Kremen
|
||||
</taxonomicName>
|
||||
|
|
@ -270,7 +270,7 @@ Lees & Kremen
|
|||
(Anjanaharibe Sud; BMNH(E) 1717105),
|
||||
<emphasis id="B95A2A364C7DC65C1F592A46FE5926F4" bold="true" box="[377,394,1899,1920]" pageId="11" pageNumber="12">B</emphasis>
|
||||
: PT ♀ of
|
||||
<taxonomicName id="4C2E8DA74C7DC65C1FD42A41FDAB26F5" box="[500,632,1900,1921]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<taxonomicName id="4C2E8DA74C7DC65C1FD42A41FDAB26F5" box="[500,632,1900,1921]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<emphasis id="B95A2A364C7DC65C1FD42A41FDAB26F5" box="[500,632,1900,1921]" italics="true" pageId="11" pageNumber="12">Ht. mimetica</emphasis>
|
||||
</taxonomicName>
|
||||
(Anjanaharibe Sud; BMNH(E) 1717113).
|
||||
|
|
@ -293,7 +293,7 @@ Oberthür, 1923
|
|||
Ward, 1872
|
||||
</taxonomicName>
|
||||
([Madagascar], BMNH(E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different
|
||||
<taxonomicName id="4C2E8DA74C7DC65C1CB92AE5FCD526A9" box="[665,774,1992,2013]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C7DC65C1CB92AE5FCD526A9" box="[665,774,1992,2013]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C7DC65C1CB92AE5FCD526A9" box="[665,774,1992,2013]" italics="true" pageId="11" pageNumber="12">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
clades.
|
||||
|
|
@ -303,20 +303,20 @@ clades.
|
|||
<paragraph id="8B91F6244C7AC65B1EE72DBAFE152260" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B95A2A364C7AC65B1EE72DBAFE6121C4" bold="true" box="[199,434,151,176]" pageId="12" pageNumber="13">Description. Wings.</emphasis>
|
||||
Upperside uniform mid brown, except light brown in the area around the expressed ocelli of HWD where the ventral white area shows through. On FW, space-MI ocellus expressed as conspicuous white spot. Space-CuA1 ocellus has narrow whitish-orange ring set in a somewhat rectangular white-translucent area whose lower proximad corner extends close to the base of space-CuA1, as in
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1DDB2C28FAFB2068" authority="Oberthur, 1923" authorityName="Oberthur" authorityYear="1923" box="[1019,1320,260,285]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1DDB2C28FAFB2068" authority="Oberthur, 1923" authorityName="Oberthur" authorityYear="1923" box="[1019,1320,260,285]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<emphasis id="B95A2A364C7AC65B1DDB2C28FBB62069" box="[1019,1125,260,285]" italics="true" pageId="12" pageNumber="13">Ht. sabas</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFBF8BD54C7AC65B1A532C29FAF32068" author="Oberthur" box="[1139,1312,260,285]" pageId="12" pageNumber="13" refString="Oberthur, C. (1923) Lepidopteres de Madagascar et d'Afrique tropicale. Etudes de Lepidopterologie Comparee, 21, 119 - 155." type="journal article" year="1923">Oberthür, 1923</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C7AC65B1B182C29FAA82068" box="[1336,1403,260,285]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7AC65B1B182C29FAA82068" box="[1336,1403,260,285]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
E), rather than
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1F392C04FE452034" box="[281,406,297,320]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<emphasis id="B95A2A364C7AC65B1F392C04FE452034" box="[281,406,297,320]" italics="true" pageId="12" pageNumber="13">Ht. cowani</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C7AC65B1F872C0AFE3A2034" box="[423,489,295,320]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7AC65B1F872C0AFE3A2034" box="[423,489,295,320]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
C-D Fianarantsoa material). Only two ocelli are conspicuously expressed in HWD, that of space-M3 and space-CuA1, both subelliptic and surrounded by a similarly shaped light orange ring. HW margin has two conspicuous mid brown Smls (submarginal lines) following the somewhat crenate margin, slightly more crenate than typical in
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1FC02CB8FD8E20D8" box="[480,605,405,428]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<emphasis id="B95A2A364C7AC65B1FC02CB8FD8E20D8" box="[480,605,405,428]" italics="true" pageId="12" pageNumber="13">Ht. cowani</emphasis>
|
||||
|
|
@ -332,7 +332,7 @@ is almost identical in patterning on either surface except for the proximad exte
|
|||
.
|
||||
<emphasis id="B95A2A364C7AC65B1C9F2F99FCE623B9" bold="true" box="[703,821,692,717]" pageId="12" pageNumber="13">Variation.</emphasis>
|
||||
Not much variation is evident between individuals of
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1EB72FF4FEFB2384" box="[151,296,729,752]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1EB72FF4FEFB2384" box="[151,296,729,752]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<emphasis id="B95A2A364C7AC65B1EB72FF4FEFB2384" box="[151,296,729,752]" italics="true" pageId="12" pageNumber="13">Ht. mimetica</emphasis>
|
||||
</taxonomicName>
|
||||
and there only one ♂ was dissected. Sexes similar, but ♀ larger, and may have more extensive white area near FW ventral apex.
|
||||
|
|
@ -364,13 +364,13 @@ and there only one ♂ was dissected. Sexes similar, but ♀ larger, and may hav
|
|||
<paragraph id="8B91F6244C7AC65B1EE72912FA8D24E9" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B95A2A364C7AC65B1EE72912FE98252C" bold="true" box="[199,331,1087,1112]" pageId="12" pageNumber="13">♂ genitalia</emphasis>
|
||||
: 278DL, PT (
|
||||
<figureCitation id="1315EAA14C7AC65B1FCD2912FDE1252C" box="[493,562,1087,1112]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
|
||||
<figureCitation id="1315EAA14C7AC65B1FCD2912FDE1252C" box="[493,562,1087,1112]" captionStart="FIGURE 5" captionStartId="19.[151,250,1641,1663]" captionTargetBox="[259,1327,202,1604]" captionTargetId="figure@19.[234,1353,193,1619]" captionTargetPageId="19" captionText="FIGURE 5. ♂ genitalia in LV (left), DV (right). A: ♂ genitalia of Heteropsis mimetica (Anjanaharibe Sud; 278 DL, CK 529). B: ♂ genitalia of PT of Ht. hazovola (Masoala; NHMUK 010289179, 26 DL). C: ♂ genitalia of Ht. alaokola, stat. rev. (Fianarantsoa; 146 DL). Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.264602" httpUri="https://zenodo.org/record/264602/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
|
||||
A, LV, DV). Tegumen with fairly straight dorsal profile from LV, where it is relatively quadrate (slight proximad notch from DV). Gnathos narrow with small rounded projection distad at base arising almost perpendicular to base of uncus in its porrect position and curving strongly towards its forwardprojecting and pointed tips (from DV gnathos bends in and out strongly from its stout base). Vinculum strongly arched proximad with moderate constriction with tegumen. Valves with strongly rounded ‘shoulder’ at base, broad throughout with valve arm not much less broad than valve base, fairly parallel sided and spatulate, tips extending further than maximum extension of uncus, with an area of spinoid setae on inner face of tip (not as extensive as in
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1EB72810FEC12420" box="[151,274,1341,1364]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<emphasis id="B95A2A364C7AC65B1EB72810FEC12420" box="[151,274,1341,1364]" italics="true" pageId="12" pageNumber="13">Ht. cowani</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1F692810FE342421" box="[329,487,1340,1365]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1F692810FE342421" box="[329,487,1340,1365]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7AC65B1F692810FE342421" box="[329,487,1340,1365]" italics="true" pageId="12" pageNumber="13">Ht. exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
|
|
@ -380,7 +380,7 @@ and
|
|||
<emphasis id="B95A2A364C7AC65B1EB7284CFEC6240C" box="[151,277,1377,1400]" italics="true" pageId="12" pageNumber="13">Ht. cowani</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1F72284CFE21240C" box="[338,498,1375,1400]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1F72284CFE21240C" box="[338,498,1375,1400]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7AC65B1F72284CFE21240C" box="[338,498,1375,1400]" italics="true" pageId="12" pageNumber="13">Ht. exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
) and inflated towards tip, while juxta is quite broad and down-lipped proximad. Aedeagus narrowly cupped beyond ostium and quite recurved away from body, fairly thin and parallel-sided.
|
||||
|
|
@ -390,20 +390,20 @@ and
|
|||
<paragraph id="8B91F6244C7AC65B1EE7288AFC902701" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B95A2A364C7AC65B1EE7288AFE9D24B4" bold="true" box="[199,334,1447,1472]" pageId="12" pageNumber="13">Etymology.</emphasis>
|
||||
Refers to the seemingly dual mimetic colour pattern (
|
||||
<figureCitation id="1315EAA14C7AC65B1D95288AFC2824B4" box="[949,1019,1447,1472]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7AC65B1D95288AFC2824B4" box="[949,1019,1447,1472]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
), on the upperside to
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1AD32884FA8D24B4" box="[1267,1374,1447,1472]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1AD32884FA8D24B4" box="[1267,1374,1447,1472]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<emphasis id="B95A2A364C7AC65B1AD32884FA8D24B4" box="[1267,1374,1447,1472]" italics="true" pageId="12" pageNumber="13">Ht. sabas</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C7AC65B1B4E288AFF752490" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7AC65B1B4E288AFF752490" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
E), and on the underside to some forms of
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1CA428E0FC7D2490" authority="Ward, 1872" authorityName="Ward" authorityYear="1872" box="[644,942,1484,1509]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="antahala">
|
||||
<emphasis id="B95A2A364C7AC65B1CA428E0FCC12491" box="[644,786,1484,1509]" italics="true" pageId="12" pageNumber="13">Ht. antahala</emphasis>
|
||||
(Ward, 1872)
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1315EAA14C7AC65B1D9D28E1FBD22490" box="[957,1025,1484,1509]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7AC65B1D9D28E1FBD22490" box="[957,1025,1484,1509]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
F); some
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1A4A28E1FB1C2491" box="[1130,1231,1484,1509]" class="Insecta" family="Nymphalidae" genus="Strabena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C7AC65B1A4A28E1FB1C2491" box="[1130,1231,1484,1509]" italics="true" pageId="12" pageNumber="13">Strabena</emphasis>
|
||||
|
|
@ -421,15 +421,15 @@ and
|
|||
exhibit similar HWV patterns (see e.g.
|
||||
<bibRefCitation id="EFBF8BD54C7AC65B1AE228C2FAA2277C" author="d'Abrera" box="[1218,1393,1519,1544]" pageId="12" pageNumber="13" refString="d'Abrera, B. L. (1997) Butterflies of the Afrotropical Region. Part 1. Papilionidae, Pieridae, Acraeidae, Danaidae, Satyridae. Hill House Publishers, Melbourne & London, 263 pp." type="book" year="1997">d’Abrera, 1997</bibRefCitation>
|
||||
). In these butterflies, there is a similar configuration on the underside of ocelli set in a mainly white area. Unlike in most of the
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1F3C2B14FE4E2724" box="[284,413,1591,1616]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1F3C2B14FE4E2724" box="[284,413,1591,1616]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="strigula">
|
||||
<emphasis id="B95A2A364C7AC65B1F3C2B14FE4E2724" box="[284,413,1591,1616]" italics="true" pageId="12" pageNumber="13">Ht. strigula</emphasis>
|
||||
</taxonomicName>
|
||||
group, but as in others of the
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1CCE2B14FC582724" box="[750,907,1591,1616]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1CCE2B14FC582724" box="[750,907,1591,1616]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7AC65B1CCE2B14FC582724" box="[750,907,1591,1616]" italics="true" pageId="12" pageNumber="13">Ht. exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
group, and in some other more early diverging clades of
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1F222B70FEA92700" box="[258,378,1629,1652]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1F222B70FEA92700" box="[258,378,1629,1652]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C7AC65B1F222B70FEA92700" box="[258,378,1629,1652]" italics="true" pageId="12" pageNumber="13">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
, the HW space-M3 ocellus is expressed.
|
||||
|
|
@ -447,7 +447,7 @@ This species was first recognized in the field by Claire Kremen at Anjanaharibe
|
|||
<emphasis id="B95A2A364C7AC65B1DC42BE4FB8D2794" box="[996,1118,1737,1760]" italics="true" pageId="12" pageNumber="13">Ht. cowani</emphasis>
|
||||
</taxonomicName>
|
||||
in BMNH and MNHN from Fianarantsoa (
|
||||
<figureCitation id="1315EAA14C7AC65B1F152BC1FEA42670" box="[309,375,1772,1797]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7AC65B1F152BC1FEA42670" box="[309,375,1772,1797]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
C-D), Ranomafana, Andrambovato or other collected localities of author or others (Vohiparara/ Sahavondronina, Ambondrombe, Anjozorobe). In
|
||||
<taxonomicName id="4C2E8DA74C7AC65B1CEE2A3CFC99265C" box="[718,842,1809,1832]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<emphasis id="B95A2A364C7AC65B1CEE2A3CFC99265C" box="[718,842,1809,1832]" italics="true" pageId="12" pageNumber="13">Ht. cowani</emphasis>
|
||||
|
|
@ -460,7 +460,7 @@ were examined. The LT ♂ of
|
|||
Butler, 1880
|
||||
</taxonomicName>
|
||||
is here designated as the specimen (
|
||||
<figureCitation id="1315EAA14C7AC65B1C962A7AFD242604" box="[694,759,1879,1904]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7AC65B1C962A7AFD242604" box="[694,759,1879,1904]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||||
C) bearing labels “
|
||||
<typeStatus id="549548864C7AC65B1DE72A75FBE82604" box="[967,1083,1880,1904]" pageId="12" pageNumber="13" type="lectotype">Lectotype</typeStatus>
|
||||
|
|
||||
|
|
@ -489,7 +489,7 @@ of
|
|||
, pl. 569, figs 4909, 4910 from ‘Fianarantsoa’, were also examined; the ♂ specimen, that is here designated the
|
||||
<typeStatus id="549548864C7BC65A1C652D91FD7F21A1" box="[581,684,188,213]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
|
||||
(
|
||||
<figureCitation id="1315EAA14C7BC65A1C9E2D91FCD021A0" box="[702,771,188,213]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="13" pageNumber="14">Fig. 2</figureCitation>
|
||||
<figureCitation id="1315EAA14C7BC65A1C9E2D91FCD021A0" box="[702,771,188,213]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." figureDoi="http://doi.org/10.5281/zenodo.264599" httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="13" pageNumber="14">Fig. 2</figureCitation>
|
||||
D), bears the data “
|
||||
<typeStatus id="549548864C7BC65A1DC12D91FB8721A1" box="[993,1108,188,213]" pageId="13" pageNumber="14" type="lectotype">Lectotype</typeStatus>
|
||||
|
|
||||
|
|
@ -517,7 +517,7 @@ is clearly synonymous with
|
|||
<emphasis id="B95A2A364C7BC65A1D082CB8FC7420D8" box="[808,935,405,428]" italics="true" pageId="13" pageNumber="14">Ht. cowani</emphasis>
|
||||
</taxonomicName>
|
||||
(BMAD122-15, DL14Z-050, Anjozorobe), whereas about 10% divergent to
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C2B2C94FD7A20A4" box="[523,681,439,464]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C2B2C94FD7A20A4" box="[523,681,439,464]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7BC65A1C2B2C94FD7A20A4" box="[523,681,439,464]" italics="true" pageId="13" pageNumber="14">Ht. exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
|
|
@ -527,11 +527,11 @@ is clearly synonymous with
|
|||
<emphasis id="B95A2A364C7BC65A1AA32CCDFB2D2083" box="[1155,1278,480,503]" italics="true" pageId="13" pageNumber="14">Ht. cowani</emphasis>
|
||||
</taxonomicName>
|
||||
has a G and a C (as in
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1ED82F29FE45236F" box="[248,406,514,539]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1ED82F29FE45236F" box="[248,406,514,539]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7BC65A1ED82F29FE45236F" box="[248,406,514,539]" italics="true" pageId="13" pageNumber="14">Ht. exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
),
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1F8E2F29FD93236F" box="[430,576,516,539]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1F8E2F29FD93236F" box="[430,576,516,539]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<emphasis id="B95A2A364C7BC65A1F8E2F29FD93236F" box="[430,576,516,539]" italics="true" pageId="13" pageNumber="14">Ht. mimetica</emphasis>
|
||||
</taxonomicName>
|
||||
has an A and a C (n=
|
||||
|
|
@ -565,11 +565,11 @@ growing on quartzite substrate, next to a stream (pers. obs.). Similar habitat o
|
|||
<paragraph id="8B91F6244C7BC65A1EE72E0FFB1422BF" blockId="13.[151,1437,151,1406]" pageId="13" pageNumber="14">
|
||||
<emphasis id="B95A2A364C7BC65A1EE72E0FFE90224F" bold="true" box="[199,323,802,827]" pageId="13" pageNumber="14">Behaviour</emphasis>
|
||||
: flies low, apparently mimetic of
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C9C2E09FCF5224F" box="[700,806,802,827]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1C9C2E09FCF5224F" box="[700,806,802,827]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sabas">
|
||||
<emphasis id="B95A2A364C7BC65A1C9C2E09FCF5224F" box="[700,806,802,827]" italics="true" pageId="13" pageNumber="14">Ht. sabas</emphasis>
|
||||
</taxonomicName>
|
||||
in flight, as first noted by Claire Kremen (pers. comm.). All individuals were caught low down along or near the path;
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1D482E65FC28222B" box="[872,1019,840,863]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<taxonomicName id="4C2E8DA74C7BC65A1D482E65FC28222B" box="[872,1019,840,863]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mimetica">
|
||||
<emphasis id="B95A2A364C7BC65A1D482E65FC28222B" box="[872,1019,840,863]" italics="true" pageId="13" pageNumber="14">Ht. mimetica</emphasis>
|
||||
</taxonomicName>
|
||||
, like
|
||||
|
|
@ -602,7 +602,7 @@ seems reluctant to come to fruit bait. The key functional aspect of this whiteni
|
|||
endemic to the rainforest of northeast
|
||||
<collectingCountry id="F339B6B44C7BC65A1AFF296FFAB5252F" box="[1247,1382,1090,1115]" name="Madagascar" pageId="13" pageNumber="14">Madagascar</collectingCountry>
|
||||
and apparently extremely localized, known from only two very localised sites (
|
||||
<figureCitation id="1315EAA14C7BC65A1DFB294BFBFE250B" box="[987,1069,1126,1151]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="13" pageNumber="14">Fig. 30</figureCitation>
|
||||
<figureCitation id="1315EAA14C7BC65A1DFB294BFBFE250B" box="[987,1069,1126,1151]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." figureDoi="http://doi.org/10.5281/zenodo.264627" httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="13" pageNumber="14">Fig. 30</figureCitation>
|
||||
A, yellow dots). The species was searched for in its
|
||||
<typeStatus id="549548864C7BC65A1F4529A6FE4625D7" box="[357,405,1163,1187]" pageId="13" pageNumber="14">type</typeStatus>
|
||||
locality in
|
||||
|
|
|
|||
|
|
@ -1,47 +1,47 @@
|
|||
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|
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|
||||
<mods:identifier id="46E1474D27908C9107C9A7B590D14C6B" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
|
||||
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|
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<mods:identifier id="C6A8BA125DD185D1E9E617CF6CA39184" type="Zenodo-Dep">264597</mods:identifier>
|
||||
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|
||||
</mods:mods>
|
||||
<treatment id="038747324C7CC65D1EB72CBFFA5F22B2" ID-DOI="http://doi.org/10.5281/zenodo.6086406" ID-GBIF-Taxon="120817106" ID-Zenodo-Dep="6086406" LSID="urn:lsid:plazi:treatment:038747324C7CC65D1EB72CBFFA5F22B2" httpUri="http://treatment.plazi.org/id/038747324C7CC65D1EB72CBFFA5F22B2" lastPageNumber="11" pageId="10" pageNumber="11">
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<treatment id="038747324C7CC65D1EB72CBFFA5F22B2" ID-DOI="http://doi.org/10.5281/zenodo.6086406" ID-GBIF-Taxon="120817106" ID-Zenodo-Dep="6086406" LSID="urn:lsid:plazi:treatment:038747324C7CC65D1EB72CBFFA5F22B2" httpUri="http://treatment.plazi.org/id/038747324C7CC65D1EB72CBFFA5F22B2" lastPageNumber="11" pageId="10" pageNumber="11" scope_family="Nymphalidae" scope_subFamily="Satyrinae" scope_tribe="Satyrini">
|
||||
<subSubSection id="C334A5AF4C7CC65D1EB72CBFFA5F22B2" pageId="10" pageNumber="11" type="nomenclature">
|
||||
<paragraph id="8B91F6244C7CC65D1EB72CBFFE3920D8" blockId="10.[151,490,402,428]" box="[151,490,402,428]" pageId="10" pageNumber="11">
|
||||
<heading id="D0D941484C7CC65D1EB72CBFFE3920D8" bold="true" box="[151,490,402,428]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
|
||||
<emphasis id="B95A2A364C7CC65D1EB72CBFFE3920D8" bold="true" box="[151,490,402,428]" pageId="10" pageNumber="11">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1EB72CBFFE4A20D8" box="[151,409,402,428]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1EB72CBFFE4A20D8" ID-CoL="94373" box="[151,409,402,428]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7CC65D1EB72CBFFE4A20D8" bold="true" box="[151,409,402,428]" italics="true" pageId="10" pageNumber="11">Heteropsis exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
group
|
||||
|
|
@ -52,37 +52,37 @@ group
|
|||
A small clade supported by multiple genes (Aduse-Poku
|
||||
<emphasis id="B95A2A364C7CC65D1D312CF7FC912086" box="[785,834,473,498]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
|
||||
., 2015,
|
||||
<figureCitation id="1315EAA14C7CC65D1DBC2CF4FC052085" box="[924,982,473,498]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="10" pageNumber="11">Fig 1</figureCitation>
|
||||
<figureCitation id="1315EAA14C7CC65D1DBC2CF4FC052085" box="[924,982,473,498]" captionStart="FIGURE 1" captionStartId="7.[151,250,1711,1733]" captionTargetBox="[166,1422,492,1675]" captionTargetId="figure@7.[150,1437,481,1690]" captionTargetPageId="7" captionText="FIGURE 1. Venation and wing patterning diagram of a fairly typical species of Heteropsis from Madagascar (Heteropis antahala group; ♂), DV. Sdbp = Supra-discocellular brush and patch. Cub = Cubital brush. Ventral banding: Cb = Costal bars. Mf = Median fleck (pale highlight). Mb = Median band. PMb = Premedian band. Ore = Ocellus ring extension. Pml = Premarginal line. Sml = Submarginal line." figureDoi="http://doi.org/10.5281/zenodo.264598" httpUri="https://zenodo.org/record/264598/files/figure.png" pageId="10" pageNumber="11">Fig 1</figureCitation>
|
||||
; Suppl. S4) for two sampled species,
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1B592CF7FED92362" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1B592CF7FED92362" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7CC65D1B592CF7FED92362" italics="true" pageId="10" pageNumber="11">Ht. exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1F652CD3FDBD2362" authority="Butler, 1880" authorityName="Butler" authorityYear="1880" box="[325,622,509,534]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1F652CD3FDBD2362" authority="Butler, 1880" authorityName="Butler" authorityYear="1880" box="[325,622,509,534]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<emphasis id="B95A2A364C7CC65D1F652CD3FE102361" box="[325,451,510,533]" italics="true" pageId="10" pageNumber="11">Ht. cowani</emphasis>
|
||||
(Butler, 1880)
|
||||
</taxonomicName>
|
||||
, with an intriguing published topology (albeit with moderate bootstrap support) as sister to all other
|
||||
<collectingCountry id="F339B6B44C7CC65D1FFE2F0CFD98234E" box="[478,587,545,570]" name="Madagascar" pageId="10" pageNumber="11">Malagasy</collectingCountry>
|
||||
Region
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1C8B2F0FFCF0234D" box="[683,803,546,569]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1C8B2F0FFCF0234D" box="[683,803,546,569]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C7CC65D1C8B2F0FFCF0234D" box="[683,803,546,569]" italics="true" pageId="10" pageNumber="11">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
. This clade is also morphologically characterised (
|
||||
<bibRefCitation id="EFBF8BD54C7CC65D1B7F2F0CFF03232A" author="Lees" pageId="10" pageNumber="11" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees, 1997</bibRefCitation>
|
||||
) by the following combination of features: HW ocellus strongly expressed in dorsal space-M3 as well as space-CuA1; spatulate valves, toothed mesad near the slightly folded tip which projects beyond the uncus; gnathos with basal curved projection which is unique among
|
||||
<collectingCountry id="F339B6B44C7CC65D1CC72FA0FC8723D2" box="[743,852,653,678]" name="Madagascar" pageId="10" pageNumber="11">Malagasy</collectingCountry>
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1D7B2FA3FC0023D1" box="[859,979,654,677]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1D7B2FA3FC0023D1" box="[859,979,654,677]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B95A2A364C7CC65D1D7B2FA3FC0023D1" box="[859,979,654,677]" italics="true" pageId="10" pageNumber="11">Heteropsis</emphasis>
|
||||
</taxonomicName>
|
||||
and developed (at least in
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1ADF2FA3FA4F23D2" box="[1279,1436,653,678]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1ADF2FA3FA4F23D2" box="[1279,1436,653,678]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7CC65D1ADF2FA3FAC823D1" box="[1279,1307,654,677]" italics="true" pageId="10" pageNumber="11">Ht</emphasis>
|
||||
.
|
||||
<emphasis id="B95A2A364C7CC65D1B092FA0FA4F23D2" box="[1321,1436,653,678]" italics="true" pageId="10" pageNumber="11">exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1EE72F9FFE9023BD" box="[199,323,690,713]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1EE72F9FFE9023BD" box="[199,323,690,713]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cowani">
|
||||
<emphasis id="B95A2A364C7CC65D1EE72F9FFE9023BD" box="[199,323,690,713]" italics="true" pageId="10" pageNumber="11">Ht. cowani</emphasis>
|
||||
</taxonomicName>
|
||||
) with its main arm downward arching over the uncus (
|
||||
|
|
@ -90,7 +90,7 @@ and
|
|||
). The vinculum is strongly bowed proximad from LV. See
|
||||
<bibRefCitation id="EFBF8BD54C7CC65D1FD32FF8FD4B239A" author="Paulian" box="[499,664,725,750]" pageId="10" pageNumber="11" refString="Paulian, R. (1951) Etudes sur les lepidopteres malgaches. II. Nouveaux satyrides (1). Memoires de l'Institut Scientifique de Madagascar, (A), 6, 387 - 394." type="journal article" year="1951">Paulian (1951)</bibRefCitation>
|
||||
for variation in the ♂ genitalia of
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1A362FFBFB61239A" box="[1046,1202,725,750]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1A362FFBFB61239A" box="[1046,1202,725,750]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7CC65D1A362FFBFB61239A" box="[1046,1202,725,750]" italics="true" pageId="10" pageNumber="11">Ht. exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
(under its synonyms
|
||||
|
|
@ -110,13 +110,13 @@ Linares
|
|||
: 488–489), who include representatives from the
|
||||
<typeStatus id="549548864C7CC65D1D7E2E33FC5D2242" box="[862,910,798,822]" pageId="10" pageNumber="11">type</typeStatus>
|
||||
localities of both nominal taxa and find no evidence for reciprocal monophyly among the respective populations. The biology of the
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1ABE2E6FFAE6222E" box="[1182,1333,833,858]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1ABE2E6FFAE6222E" box="[1182,1333,833,858]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7CC65D1ABE2E6FFAE6222E" box="[1182,1333,833,858]" italics="true" pageId="10" pageNumber="11">H. exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
group is unknown, but these are low flying forest interior butterflies likely to be associated with low grasses, which seem rarely attracted to fruit. They tend to occur in localised and bright forest pockets with a very humid aspect such as near streams, up to around
|
||||
<quantity id="4CD65BC14C7CC65D1FE62E83FDC822B1" box="[454,539,942,966]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="11" unit="m" value="1800.0">1800 m</quantity>
|
||||
elevation.
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1CB62E83FCE022B2" box="[662,819,941,966]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<taxonomicName id="4C2E8DA74C7CC65D1CB62E83FCE022B2" box="[662,819,941,966]" class="Insecta" family="Nymphalidae" genus="Heteropsis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="exocellata">
|
||||
<emphasis id="B95A2A364C7CC65D1CB62E83FCE022B2" box="[662,819,941,966]" italics="true" pageId="10" pageNumber="11">Ht. exocellata</emphasis>
|
||||
</taxonomicName>
|
||||
has been observed hilltopping on a ridge (pers. obs.).
|
||||
|
|
|
|||
|
|
@ -1,400 +0,0 @@
|
|||
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<mods:title id="CAFDE6510999EB617926C3BA3134F511">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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|
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<mods:namePart id="909C54E3DB4EC3C49F9A5CC4A3162CAD">Reyes-Hernández, José L.</mods:namePart>
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<mods:affiliation id="F36B29595082475E7C284EC59686A67D">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="DE9E636B2AB15DE425A7A4238AF40684" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
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<mods:namePart id="57E29C7BAE2BE2A9186B973E6225C737">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="911BEBAA962759A02746A44437403755">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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<mods:namePart id="A61467CE4590F174702920D0826A58E4">Shaw, Josh Jenkins</mods:namePart>
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<mods:affiliation id="B24EDC7BFD820B91E2B906E1D2C8DFA7">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:namePart id="C48F6FD7A0F809CD0A0051C42414F10F">Solodovnikov, Alexey</mods:namePart>
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<mods:title id="0DEDFFAEFFB26113E035166C10D952B9">Zool. J. Linn. Soc.</mods:title>
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||||
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|
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|
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<mods:number id="0EDC4D50DB25C54C8A0A13FEA59B7A1E">2023-10-28</mods:number>
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<mods:url id="B206804E1D23E050C91E24A121D5825A">https://doi.org/10.1093/zoolinnean/zlad143</mods:url>
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<mods:classification id="47F92DBAE0638EDB0BFE8F519DCC52C5">journal article</mods:classification>
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<mods:identifier id="C821D83533E175AFFF9FEB0B7DC942A1" type="DOI">10.1093/zoolinnean/zlad143</mods:identifier>
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<treatment id="03D187F3FF802E06FF09124AFE08F7CF" LSID="urn:lsid:plazi:treatment:03D187F3FF802E06FF09124AFE08F7CF" httpUri="http://treatment.plazi.org/id/03D187F3FF802E06FF09124AFE08F7CF" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
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||||
<subSubSection id="C362656EFF802E05FF09124AFD36F2FB" box="[157,719,878,905]" pageId="33" pageNumber="34" type="nomenclature">
|
||||
<paragraph id="8BC736E5FF802E05FF09124AFD36F2FB" blockId="33.[157,719,878,905]" box="[157,719,878,905]" pageId="33" pageNumber="34">
|
||||
<heading id="D08F8189FF802E05FF09124AFD36F2FB" bold="true" box="[157,719,878,905]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="34" reason="2">
|
||||
<taxonomicName id="4C784D66FF802E05FF09124AFD36F2FB" ID-CoL="72MP2" authority="(Fairmaire and Germain, 1862)" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[157,719,878,905]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF802E05FF09124AFD36F2FB" bold="true" box="[157,719,878,905]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FF09124AFE84F2FA" bold="true" box="[157,381,878,905]" italics="true" pageId="33" pageNumber="34">Loncovilius semiflavus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF802E05FE1F124AFD3DF2FB" author="Fairmaire L & Germain P" box="[395,708,878,904]" pageId="33" pageNumber="34" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF802E05FF5D1284FD5AF2C9" box="[201,675,928,954]" pageId="33" pageNumber="34" type="description">
|
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<paragraph id="8BC736E5FF802E05FF5D1284FD5AF2C9" blockId="33.[201,675,928,954]" box="[201,675,928,954]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FF5D1284FD5AF2C9" bold="true" box="[201,675,928,954]" pageId="33" pageNumber="34">
|
||||
(
|
||||
<figureCitation id="13432A60FF802E05FF471284FEDAF2C9" box="[211,291,928,954]" captionStart="Figure 6" captionStartId="14.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@14.[168,1434,144,1840]" captionTargetPageId="14" captionText="Figure 6. Habitus of Loncovilius species. A, male of L. edwardsianus. B, male of L. cantharoides. C, female of L. semiflavus. D, male of L. barclayi." pageId="33" pageNumber="34">Figs 6C</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FF802E05FEB91284FEB6F2C9" box="[301,335,928,954]" captionStart="Figure 7" captionStartId="15.[113,178,1868,1892]" captionTargetBox="[153,1419,163,1840]" captionTargetId="figure-6@15.[153,1419,144,1840]" captionTargetPageId="15" captionText="Figure 7. Female habitus of Loncovilius species.A, L. lividipennis. B, L. carlsbergi. Head. C, L. hammondi. D, L. semiflavus. E, L. carlsbergi. F, Male mesotarsi of L. cantharoides." pageId="33" pageNumber="34">7D</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FF802E05FECC1284FE6EF2C9" box="[344,407,928,954]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="33" pageNumber="34">9D, G</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FF802E05FE341284FE17F2C9" box="[416,494,928,954]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="33" pageNumber="34">11A–C</figureCitation>
|
||||
, 12G, 13G, 14G)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF802E05FFE512F5FDF5F597" pageId="33" pageNumber="34" type="reference_group">
|
||||
<paragraph id="8BC736E5FF802E05FFE512F5FDF5F597" blockId="33.[112,763,977,1252]" pageId="33" pageNumber="34">
|
||||
<bibRefCitation id="EFE94B14FF802E05FFE512F5FE16F29A" author="Fairmaire L & Germain P" box="[113,495,977,1001]" pageId="33" pageNumber="34" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain 1862: 428</bibRefCitation>
|
||||
(original description as
|
||||
<taxonomicName id="4C784D66FF802E05FFE512D5FEC8F57A" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[113,305,1009,1033]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF802E05FFE512D5FEC8F57A" box="[113,305,1009,1033]" italics="true" pageId="33" pageNumber="34">Quedius semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
);
|
||||
<bibRefCitation id="EFE94B14FF802E05FEC512D5FD36F57B" author="Gemminger M & Harold E" box="[337,719,1008,1033]" pageId="33" pageNumber="34" pagination="425 - 752" refId="ref36690" refString="Gemminger M, Harold E. Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus. Vol. 2. Dytiscidae, Gyrinidae, Hydrophilidae, Staphylinidae, Pselaphidae, Gnostidae, Paussidae, Scydmaenidae, Silphidae, Trichopterygidae, Scaphidiidae. Monachii [Munich]: E. H. Gummi, 1868, 425 - 752." type="book chapter" year="1868">Gemminger and Harold 1868: 570</bibRefCitation>
|
||||
(as synonym of
|
||||
<taxonomicName id="4C784D66FF802E05FF7F1534FE54F55B" authorityName="Fairmaire and Germain" authorityYear="1862" box="[235,429,1040,1064]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF802E05FF7F1534FE54F55B" box="[235,429,1040,1064]" italics="true" pageId="33" pageNumber="34">Quedius lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
);
|
||||
<bibRefCitation id="EFE94B14FF802E05FE511534FF61F534" author="Bernhauer M & Schubert K" pageId="33" pageNumber="34" pagination="409 - 98" refId="ref36154" refString="Bernhauer M, Schubert K. In: Staphylinidae V, Schenkling S (eds), Coleopterorum Catalogus 5 (67). Berlin: Junk, 1916, 409 - 98." type="book chapter" year="1916">Bernhauer and Schubert 1916: 434</bibRefCitation>
|
||||
(as
|
||||
<taxonomicName id="4C784D66FF802E05FF5C150BFE78F534" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[200,385,1071,1095]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF802E05FF5C150BFE78F534" box="[200,385,1071,1095]" italics="true" pageId="33" pageNumber="34">Quedius semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
, catalogue);
|
||||
<bibRefCitation id="EFE94B14FF802E05FD9E150BFD2DF534" author="Germain P" box="[522,724,1071,1095]" pageId="33" pageNumber="34" pagination="391 - 445" refId="ref36747" refString="Germain P. Apuntes Entomolojicos. Rectificaciones, anotaciones complementarias, i descripciones de siete especies nuevas de coleopteros. Anales de la Universidad de Chile 1903; 113: 391 - 445." type="journal article" year="1903">Germain 1903: 443</bibRefCitation>
|
||||
(as
|
||||
<taxonomicName id="4C784D66FF802E05FFE0156BFEB2F514" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[116,331,1103,1127]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF802E05FFE0156BFEB2F514" box="[116,331,1103,1127]" italics="true" pageId="33" pageNumber="34">Loncovilius semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
, characters);
|
||||
<bibRefCitation id="EFE94B14FF802E05FE4E156BFD03F514" author="Coiffait H & Saiz F" box="[474,762,1103,1127]" pageId="33" pageNumber="34" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz 1966: 411</bibRefCitation>
|
||||
(as
|
||||
<emphasis id="B90CEAF7FF802E05FF00154AFE7EF5F5" box="[148,391,1134,1158]" italics="true" pageId="33" pageNumber="34">
|
||||
<taxonomicName id="4C784D66FF802E05FF00154AFEFBF5F5" box="[148,258,1134,1158]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus" sensu="stricto">Loncovilius</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF802E05FE92154BFED8F5F5" box="[262,289,1135,1158]" pageId="33" pageNumber="34" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
<taxonomicName id="4C784D66FF802E05FEB0154AFE7EF5F5" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[292,391,1134,1158]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">semiflavus</taxonomicName>
|
||||
</emphasis>
|
||||
; characters); Coiffait and Sáiz, 1968: 365 (as
|
||||
<emphasis id="B90CEAF7FF802E05FF5215A9FE47F5D6" box="[198,446,1165,1189]" italics="true" pageId="33" pageNumber="34">
|
||||
<taxonomicName id="4C784D66FF802E05FF5215A9FECDF5D6" box="[198,308,1165,1189]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus" sensu="stricto">Loncovilius</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF802E05FEAE15AAFEAFF5D6" box="[314,342,1166,1189]" pageId="33" pageNumber="34" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
<taxonomicName id="4C784D66FF802E05FEC815A9FE47F5D6" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[348,446,1165,1189]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">semiflavus</taxonomicName>
|
||||
</emphasis>
|
||||
, checklist);
|
||||
<bibRefCitation id="EFE94B14FF802E05FDA915A9FD2CF5D6" author="Saiz F" box="[573,725,1165,1189]" pageId="33" pageNumber="34" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz 1971: 385</bibRefCitation>
|
||||
(as
|
||||
<emphasis id="B90CEAF7FF802E05FFE71589FE95F5B6" box="[115,364,1197,1221]" italics="true" pageId="33" pageNumber="34">
|
||||
<taxonomicName id="4C784D66FF802E05FFE71589FF18F5B6" box="[115,225,1197,1221]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus" sensu="stricto">Loncovilius</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF802E05FF7C1589FEFAF5B7" box="[232,259,1197,1220]" pageId="33" pageNumber="34" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
<taxonomicName id="4C784D66FF802E05FE9E1589FE95F5B6" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[266,364,1197,1221]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">semiflavus</taxonomicName>
|
||||
</emphasis>
|
||||
; additional material);
|
||||
<bibRefCitation id="EFE94B14FF802E05FDC31589FF5CF597" author="Herman LH" pageId="33" pageNumber="34" pagination="3021 - 840" refId="ref36981" refString="Herman LH. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine Group (Part 3). Staphylininae: Staphylinini: (Quediina + Xanthopygina), Xantholinini. Staphylinidae Incertae Sedis. Fossils, Protactinae †. Bulletin of the American Museum of Natural History 2001 b; 265: 3021 - 840." type="journal article" year="2001">Herman 2001b: 3083</bibRefCitation>
|
||||
(as
|
||||
<taxonomicName id="4C784D66FF802E05FF5A15E8FE5DF597" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[206,420,1228,1252]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF802E05FF5A15E8FE5DF597" box="[206,420,1228,1252]" italics="true" pageId="33" pageNumber="34">Loncovilius semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
; catalog).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF802E05FFE51422FD3AF7DC" pageId="33" pageNumber="34" type="materials_examined">
|
||||
<paragraph id="8BC736E5FF802E05FFE51422FE00F705" blockId="33.[113,763,1285,1654]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FFE51422FE94F46E" box="[113,365,1285,1309]" italics="true" pageId="33" pageNumber="34">
|
||||
Type material.
|
||||
<materialsCitation id="3B103CB8FF802E05FE9D1422FEEAF4C9" collectionCode="RBINS" collectorName="A. Fauvel" determinerName="Coiff. & Saiz / J. L. Reyes-Hernandez" location="Quedius semiflavus Fairm and Germ" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
|
||||
<typeStatus id="54C38847FF802E05FE9D1422FE91F46E" box="[265,360,1286,1309]" pageId="33" pageNumber="34" type="lectotype">Lectotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D7EFD6CFF802E05FEE21421FE50F46E" box="[374,425,1285,1309]" pageId="33" pageNumber="34" type="male">male</specimenCount>
|
||||
, pinned, with genitalia in a separate microvial, with labels as follows: Coll. et det
|
||||
<collectorName id="268D5333FF802E05FD021401FD02F44E" box="[662,763,1317,1341]" pageId="33" pageNumber="34">A. Fauvel</collectorName>
|
||||
<taxonomicName id="4C784D66FF802E05FFE51460FDAEF42F" authority="Fairm and Germ. R. I. S." authorityName="Fairm and Germ. R. I. S." box="[113,599,1348,1372]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<location id="8EA7603EFF802E05FFE51460FDF5F42F" LSID="urn:lsid:plazi:treatment:03D187F3FF802E06FF09124AFE08F7CF:8EA7603EFF802E05FFE51460FDF5F42F" box="[113,524,1348,1372]" name="Quedius semiflavus Fairm and Germ" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FFE51460FECFF42F" box="[113,310,1348,1372]" italics="true" pageId="33" pageNumber="34">Quedius semiflavus</emphasis>
|
||||
Fairm and Germ
|
||||
</location>
|
||||
. R.I.S.
|
||||
</taxonomicName>
|
||||
c.N.B. 17.479/ Concepcion/Lectotype [Red label]/
|
||||
<taxonomicName id="4C784D66FF802E05FDA51447FD66F408" box="[561,671,1379,1403]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF802E05FDA51447FD66F408" box="[561,671,1379,1403]" italics="true" pageId="33" pageNumber="34">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<emphasis id="B90CEAF7FF802E05FD471440FD22F408" box="[723,731,1380,1403]" italics="true" pageId="33" pageNumber="34">s</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF802E05FD751440FD10F408" box="[737,745,1380,1403]" italics="true" pageId="33" pageNumber="34">s</emphasis>
|
||||
.)
|
||||
<taxonomicName id="4C784D66FF802E05FFE514A7FF2DF4E8" box="[113,212,1411,1435]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF802E05FFE514A7FF2DF4E8" box="[113,212,1411,1435]" italics="true" pageId="33" pageNumber="34">semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
1964 det.
|
||||
<determinerName id="7E1BDA21FF802E05FEDF14A7FE7CF4E8" box="[331,389,1411,1435]" pageId="33" pageNumber="34">Coiff.</determinerName>
|
||||
et
|
||||
<determinerName id="7E1BDA21FF802E05FE3E14A7FD33F4E8" box="[426,714,1411,1435]" pageId="33" pageNumber="34">Sáiz/J. L. Reyes-Hernández</determinerName>
|
||||
det. 2022 [
|
||||
<collectionCode id="ED69AE20FF802E05FF201486FEF1F4C9" box="[180,264,1442,1466]" collectionName="RBINS" pageId="33" pageNumber="34">RBINS</collectionCode>
|
||||
].
|
||||
</materialsCitation>
|
||||
</emphasis>
|
||||
<materialsCitation id="3B103CB8FF802E05FE8E1486FE00F705" collectionCode="RBINS" collectorName="A. Fauvel" determinerName="Saiz / J. L. Reyes-Hernandez" location="Quedius semiflavus Fairm and Germ" pageId="33" pageNumber="34" specimenCount="1" typeStatus="paralectotype">
|
||||
<emphasis id="B90CEAF7FF802E05FE8E1486FE55F4C9" box="[282,428,1442,1466]" italics="true" pageId="33" pageNumber="34">
|
||||
<typeStatus id="54C38847FF802E05FE8E1486FE5EF4C9" box="[282,423,1442,1466]" pageId="33" pageNumber="34" type="paralectotype">Paralectotypes</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
Coll. et det.
|
||||
<collectorName id="268D5333FF802E05FDAE1486FD67F4C9" box="[570,670,1442,1466]" pageId="33" pageNumber="34">A. Fauvel</collectorName>
|
||||
,
|
||||
<taxonomicName id="4C784D66FF802E05FD3E1486FDEAF4AA" authority="Fairm and Germ., R. I. Sc. N. B." authorityName="Fairm and Germ., R. I. Sc. N. B." class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<location id="8EA7603EFF802E05FD3E1486FE6BF4AA" LSID="urn:lsid:plazi:treatment:03D187F3FF802E06FF09124AFE08F7CF:8EA7603EFF802E05FD3E1486FE6BF4AA" name="Quedius semiflavus Fairm and Germ" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FD3E1486FF2DF4AA" italics="true" pageId="33" pageNumber="34">Quedius semiflavus</emphasis>
|
||||
Fairm and Germ
|
||||
</location>
|
||||
., R.I.Sc.N.B.
|
||||
</taxonomicName>
|
||||
17.479/paralectotype [red paper]/
|
||||
<taxonomicName id="4C784D66FF802E05FF6314C5FE2AF48A" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[247,467,1505,1529]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF802E05FF6314C5FE2AF48A" box="[247,467,1505,1529]" italics="true" pageId="33" pageNumber="34">Loncovilius semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
, 1964 det. Coiff. et
|
||||
<determinerName id="7E1BDA21FF802E05FD2314C5FEBEF76B" pageId="33" pageNumber="34">Sáiz/J. L. Reyes-Hernández</determinerName>
|
||||
det. 2022; concepcion/
|
||||
<taxonomicName id="4C784D66FF802E05FDDF1724FF40F744" authority="Fairm. Germ." authorityName="Fairm. Germ." class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF802E05FDDF1724FD57F76B" box="[587,686,1536,1560]" italics="true" pageId="33" pageNumber="34">semiflavus</emphasis>
|
||||
Fairm. Germ.
|
||||
</taxonomicName>
|
||||
/R.I.Sc.N.B. 17.479, Coll. et det. A. Fauvel/paralectotype [red paper]/
|
||||
<taxonomicName id="4C784D66FF802E05FF67171BFE32F724" box="[243,459,1599,1623]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF802E05FF67171BFE32F724" box="[243,459,1599,1623]" italics="true" pageId="33" pageNumber="34">Loncovilius semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
, 1964 det. Coiff. et Saiz/J. L. Reyes-Hernández det. 2022 [
|
||||
<collectionCode id="ED69AE20FF802E05FE0E177AFE17F705" box="[410,494,1630,1654]" collectionName="RBINS" pageId="33" pageNumber="34">RBINS</collectionCode>
|
||||
].
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF802E05FFE517B3FD3AF7DC" blockId="33.[113,707,1687,1711]" box="[113,707,1687,1711]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FFE517B3FE94F7DC" box="[113,365,1687,1711]" italics="true" pageId="33" pageNumber="34">Other material examined:</emphasis>
|
||||
Supporting Information, File S7.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF802E05FFE517F5FE5DF635" pageId="33" pageNumber="34" type="diagnosis">
|
||||
<paragraph id="8BC736E5FF802E05FFE517F5FE5DF635" blockId="33.[113,762,1745,1863]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FFE517F5FF2EF79B" box="[113,215,1745,1768]" italics="true" pageId="33" pageNumber="34">Diagnosis:</emphasis>
|
||||
Elytra translucent yellow or yellowish-brown. Male tergite X with broad U-shape emargination medially (
|
||||
<figureCitation id="13432A60FF802E05FD0417D4FD11F67B" box="[656,744,1776,1800]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="33" pageNumber="34">Fig. 12G</figureCitation>
|
||||
); female tergite X slightly emarginate medio-apically (
|
||||
<figureCitation id="13432A60FF802E05FD04162BFD10F654" box="[656,745,1807,1831]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="33" pageNumber="34">Fig. 13G</figureCitation>
|
||||
). Aedeagus as in
|
||||
<figureCitation id="13432A60FF802E05FE98160BFE67F634" box="[268,414,1839,1863]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="33" pageNumber="34">Figure 11A–C</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF802E06FFE5164DFE85F34C" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
|
||||
<paragraph id="8BC736E5FF802E05FFE5164DFB52F075" blockId="33.[113,764,1896,1951]" lastBlockId="33.[809,1461,144,1954]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FFE5164DFF11F6F3" box="[113,232,1897,1920]" italics="true" pageId="33" pageNumber="34">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFF802E05FE07164FFE54F60C" box="[403,429,1899,1919]" count="1" pageId="33" pageNumber="34" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FF802E05FD0A164DFD48F6F3" box="[670,689,1897,1920]" italics="true" pageId="33" pageNumber="34">N</emphasis>
|
||||
= 10]: FBL = 3.82–4.19 (4.02); TL = 6.25–6.88 (6.54); HW = 1–1.05 (1.03); HL = 0.98–1.09 (1.02); HW/HL = 0.96–1.04 (1.01); PW = 1.28–1.44 (1.35); PL = 1.28–1.43 (1.35); PW/ PL = 0.99–1.01 (1); EW = 1.66–1.8 (1.74); EL = 1.5–1.8 (1.65); PW/HW = 1.27–1.37 (1.32).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF802E05FCD1102AFBD1F0B1" blockId="33.[809,1461,144,1954]" pageId="33" pageNumber="34">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFF802E05FC751034FC02F057" box="[993,1019,272,292]" count="1" pageId="33" pageNumber="34" type="female">♀</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FF802E05FB4F102AFB17F056" box="[1243,1262,270,293]" italics="true" pageId="33" pageNumber="34">N</emphasis>
|
||||
= 10]: FBL = 4.1– 4.45 (4.29); TL = 7.2–8.6 (7.77); HW = 1.08–1.15 (1.1); HL = 1.04–1.13 (1.06); HW/HL = 1–1.06 (1.04); PW = 1.4– 1.5 (1.47); PL = 1.38–1.5 (1.46); PW/PL = 0.98–1.05 (1.01); EW = 1.85–1.9 (1.88); EL = 1.67–1.82 (1.77); PW/ HW = 1.27–1.39 (1.34).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF802E05FCD110EDFB04F32C" blockId="33.[809,1461,144,1954]" pageId="33" pageNumber="34">Head and pronotum yellow-orange or reddish-brown, usually females darker than males; elytra translucent yellow; antennae, mouthparts, and legs yellow or yellowish-brown; mesosternum, metasternum, metacoxa and abdomen dark reddish-brown; abdominal segments with pale posterior margin.</paragraph>
|
||||
<paragraph id="8BC736E5FF802E05FCD11342FC67F43C" blockId="33.[809,1461,144,1954]" pageId="33" pageNumber="34">
|
||||
Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small size. Eyes medium size (EYL/HL = x = 0.5), from 1.2 to 1.4 times as long as temples (in lateral view); distance between eyes about 1.2 times as long as length of eye in males and 1.39 times in females. Antennomeres 2 and 3 subequal in length; antennomeres 4 and 5 subequal in length; antennomeres 5 to 10 gradually becoming shorter apicad; a11 is from 1.88 to 2 times as long as a
|
||||
<quantity id="4C809B00FF802E05FAE912BBFA4AF2C4" box="[1405,1459,927,951]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="33" pageNumber="34" unit="in" value="10.0">10 in</quantity>
|
||||
males and from 1.6 to 1.8 times in females. Basal puncture single; parocular puncture usually single too, distinctly separated from eye margin; posterior frontal puncture located anterior to temporal puncture; ventral basal ridge rather straight, almost united with gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge distinct only laterally, nuchal and infraorbital ridges fused; gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF802E05FCD11472FC7EF759" blockId="33.[809,1461,144,1954]" pageId="33" pageNumber="34">Pronotum about as wide as long, strongly convex, evenly curved at sides, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to three times as long as, diameter of punctures. Mesosternum with three or four macrosetae arranged in a row medially.</paragraph>
|
||||
<paragraph id="8BC736E5FF802E05FCD11715FC31F656" blockId="33.[809,1461,144,1954]" pageId="33" pageNumber="34">Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate.</paragraph>
|
||||
<paragraph id="8BC736E5FF802E06FCD11608FD91F0F0" blockId="33.[809,1461,144,1954]" lastBlockId="34.[129,780,144,575]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||||
Abdominal tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area; tergite VIII distinctly emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sterniteVIIIwithstrong (
|
||||
<figureCitation id="13432A60FF832E06FEEF118BFE36F1B4" box="[379,463,175,199]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="34" pageNumber="35">Fig.14G</figureCitation>
|
||||
),V-shapedemarginationmedially; female sternite VIII with weak apical medial emargination; male sternite IX with arcuate emargination medially, its basal portion distinctly longer than distal portion; male tergite X with broad U-shape emargination medially (
|
||||
<figureCitation id="13432A60FF832E06FD841009FD9EF036" box="[528,615,301,325]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="34" pageNumber="35">Fig. 12G</figureCitation>
|
||||
); female tergite X slightly emarginate medio-apically (
|
||||
<figureCitation id="13432A60FF832E06FD9E1068FD9DF017" box="[522,612,332,356]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="34" pageNumber="35">Fig. 13G</figureCitation>
|
||||
); ovipositor, its second gonocoxite with one macroseta medially.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF832E06FF0810AFFE85F34C" blockId="34.[129,780,144,575]" pageId="34" pageNumber="35">
|
||||
Aedeagus as in
|
||||
<figureCitation id="13432A60FF832E06FED410AFFE2CF0D0" box="[320,469,395,419]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="34" pageNumber="35">Figure 11A–C</figureCitation>
|
||||
, its length 1.21; median lobe rod-like with moderately emarginate apex; paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging in a trapezoidal slightly emarginate apex, with small translucent ‘peg setae’, in lateral view almost straight and slightly concave apically.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF832E06FF151345FDF6F246" pageId="34" pageNumber="35" type="distribution">
|
||||
<paragraph id="8BC736E5FF832E06FF151345FDF6F246" blockId="34.[128,778,609,821]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B90CEAF7FF832E06FF151345FD9FF30A" box="[129,614,609,633]" italics="true" pageId="34" pageNumber="35">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FF832E06FE1E1345FD9FF30A" box="[394,614,609,633]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="semiflavus">Loncovilius semiflavus</taxonomicName>
|
||||
</emphasis>
|
||||
is known only from
|
||||
<collectingCountry id="F36F7675FF832E06FF5513A4FF02F3EB" box="[193,251,640,664]" name="Chile" pageId="34" pageNumber="35">Chile</collectingCountry>
|
||||
, from the
|
||||
<collectingRegion id="49BCF807FF832E06FEE913A4FE3AF3EB" box="[381,451,640,664]" country="Chile" name="Biobio" pageId="34" pageNumber="35">Biobío</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FF832E06FE4113A4FDEEF3EB" box="[469,535,640,664]" country="Chile" name="Maule" pageId="34" pageNumber="35">Maule</collectingRegion>
|
||||
, Ñuble, and
|
||||
<collectingRegion id="49BCF807FF832E06FD2513A4FCF0F3EB" box="[689,777,640,664]" country="Chile" name="Region Metropolitana" pageId="34" pageNumber="35">Santiago</collectingRegion>
|
||||
Metropolitan regions where it occurs from
|
||||
<quantity id="4C809B00FF832E06FDA713BBFD3FF3C4" box="[563,710,671,695]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="12.0" metricValueMin="2.0" pageId="34" pageNumber="35" unit="m" value="700.0" valueMax="1200.0" valueMin="200.0">200 to 1200 m</quantity>
|
||||
of elevation. Its distribution is confined to the northern portions of the Valdivian temperate forest ecoregion. It has been collected by sifting leaf litter and beating the forest vegetation and flowers, e.g. from
|
||||
<taxonomicName id="4C784D66FF832E06FF4B1239FE89F247" box="[223,368,797,820]" class="Magnoliopsida" family="Winteraceae" genus="Drimys" kingdom="Plantae" order="Canellales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="winteri">
|
||||
<emphasis id="B90CEAF7FF832E06FF4B1239FE89F247" box="[223,368,797,820]" italics="true" pageId="34" pageNumber="35">Drimys winteri</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C784D66FF832E06FEE81239FDFDF246" box="[380,516,797,821]" class="Magnoliopsida" family="Winteraceae" kingdom="Plantae" order="Canellales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="family">Winteraceae</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF832E06FF151272FE08F7CF" pageId="34" pageNumber="35" type="discussion">
|
||||
<paragraph id="8BC736E5FF832E06FF151272FE08F7CF" blockId="34.[127,781,854,1724]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B90CEAF7FF832E06FF151272FE2BF21D" box="[129,466,854,878]" italics="true" pageId="34" pageNumber="35">
|
||||
Remarks:
|
||||
<taxonomicName id="4C784D66FF832E06FF7B1272FE2BF21D" baseAuthorityName="Fairmaire & Germain" baseAuthorityYear="1862" box="[239,466,854,878]" class="Magnoliopsida" family="Winteraceae" genus="Lonvcovilius" kingdom="Plantae" order="Canellales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="semiflavus">Lonvcovilius semiflavus</taxonomicName>
|
||||
</emphasis>
|
||||
was described as a species of
|
||||
<taxonomicName id="4C784D66FF832E06FF151251FF28F2FE" box="[129,209,885,909]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF832E06FF151251FF28F2FE" box="[129,209,885,909]" italics="true" pageId="34" pageNumber="35">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
based on an unspecified number of specimens from ‘Concepcion’ (Concepción city in Central
|
||||
<collectingCountry id="F36F7675FF832E06FDDB12B1FD73F2DE" box="[591,650,917,941]" name="Chile" pageId="34" pageNumber="35">Chile</collectingCountry>
|
||||
) (
|
||||
<bibRefCitation id="EFE94B14FF832E06FD3312B1FEBBF2BF" author="Fairmaire L & Germain P" pageId="34" pageNumber="35" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain 1862</bibRefCitation>
|
||||
).
|
||||
<bibRefCitation id="EFE94B14FF832E06FECC1290FD67F2BF" author="Gemminger M & Harold E" box="[344,670,948,972]" pageId="34" pageNumber="35" pagination="425 - 752" refId="ref36690" refString="Gemminger M, Harold E. Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus. Vol. 2. Dytiscidae, Gyrinidae, Hydrophilidae, Staphylinidae, Pselaphidae, Gnostidae, Paussidae, Scydmaenidae, Silphidae, Trichopterygidae, Scaphidiidae. Monachii [Munich]: E. H. Gummi, 1868, 425 - 752." type="book chapter" year="1868">Gemminger and Harold (1868)</bibRefCitation>
|
||||
listed it as a synonym of
|
||||
<taxonomicName id="4C784D66FF832E06FE9A12F7FCFDF298" authority="Fairmaire and Germain, 1862" authorityName="Fairmaire and Germain" authorityYear="1862" box="[270,772,979,1003]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF832E06FE9A12F7FE2BF298" box="[270,466,979,1003]" italics="true" pageId="34" pageNumber="35">Quedius lividipennis</emphasis>
|
||||
<bibRefCitation id="EFE94B14FF832E06FE4312F7FCFDF298" author="Fairmaire L & Germain P" box="[471,772,979,1003]" pageId="34" pageNumber="35" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, but
|
||||
<bibRefCitation id="EFE94B14FF832E06FF3312D7FEB6F578" author="Germain P" box="[167,335,1011,1035]" pageId="34" pageNumber="35" pagination="391 - 445" refId="ref36747" refString="Germain P. Apuntes Entomolojicos. Rectificaciones, anotaciones complementarias, i descripciones de siete especies nuevas de coleopteros. Anales de la Universidad de Chile 1903; 113: 391 - 445." type="journal article" year="1903">Germain (1903)</bibRefCitation>
|
||||
considered both species valid when erecting a new genus
|
||||
<taxonomicName id="4C784D66FF832E06FF6A1536FE95F559" box="[254,364,1042,1066]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF832E06FF6A1536FE95F559" box="[254,364,1042,1066]" italics="true" pageId="34" pageNumber="35">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
for them.
|
||||
<bibRefCitation id="EFE94B14FF832E06FE451536FD3FF559" author="Coiffait H & Saiz F" box="[465,710,1042,1066]" pageId="34" pageNumber="35" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz (1966)</bibRefCitation>
|
||||
examined
|
||||
<specimenCount id="9D7EFD6CFF832E06FF261515FEAFF53A" box="[178,342,1073,1097]" count="3" pageId="34" pageNumber="35" type="generic">three specimens</specimenCount>
|
||||
from the collection of A. Fauvel in Brussels that they considered
|
||||
<typeStatus id="54C38847FF832E06FEF41575FE43F51A" box="[352,442,1105,1129]" pageId="34" pageNumber="35" type="syntype">syntypes</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C784D66FF832E06FE761575FD9AF51A" baseAuthorityName="Fairmaire & Germain" baseAuthorityYear="1862" box="[482,611,1105,1129]" class="Magnoliopsida" family="Winteraceae" genus="Lonvcovilius" kingdom="Plantae" order="Canellales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF832E06FE761575FE08F51B" box="[482,497,1105,1128]" italics="true" pageId="34" pageNumber="35">L</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF832E06FD941575FD9AF51A" box="[512,611,1105,1129]" italics="true" pageId="34" pageNumber="35">semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
and designated
|
||||
<specimenCount id="9D7EFD6CFF832E06FF151554FF19F5FB" box="[129,224,1136,1160]" count="1" pageId="34" pageNumber="35" type="male">one male</specimenCount>
|
||||
as a
|
||||
<typeStatus id="54C38847FF832E06FE821554FE81F5FB" box="[278,376,1136,1160]" pageId="34" pageNumber="35" type="lectotype">lectotype</typeStatus>
|
||||
. Morphology and labels of these
|
||||
<specimenCount id="9D7EFD6CFF832E06FD401554FF15F5D4" count="3" pageId="34" pageNumber="35" type="generic">three specimens</specimenCount>
|
||||
match the original description and the type locality, respectively. According to Camousseight (1980), the type material of the species described by Philibert Germain is kept at the National Museum of
|
||||
<collectingCountry id="F36F7675FF832E06FECB15C9FE6EF476" box="[351,407,1261,1285]" name="Chile" pageId="34" pageNumber="35">Chile</collectingCountry>
|
||||
in
|
||||
<collectingRegion id="49BCF807FF832E06FE2F15C9FDEDF476" box="[443,532,1261,1285]" country="Chile" name="Region Metropolitana" pageId="34" pageNumber="35">Santiago</collectingRegion>
|
||||
. However, even though Camousseight (1980) listed type material for
|
||||
<taxonomicName id="4C784D66FF832E06FDEF1429FCF0F456" authorityName="Fairmaire and Germain" authorityYear="1862" box="[635,777,1293,1317]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF832E06FDEF1429FCF0F456" box="[635,777,1293,1317]" italics="true" pageId="34" pageNumber="35">L. lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
(see below that species) and some other
|
||||
<taxonomicName id="4C784D66FF832E06FDB61408FD49F437" authorityName="Seevers" authorityYear="1944" box="[546,688,1324,1348]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="tribe" tribe="Amblyopinini">Amblyopinini</taxonomicName>
|
||||
as being deposited there, he did not mention any types of
|
||||
<emphasis id="B90CEAF7FF832E06FD1C1468FCF4F410" box="[648,781,1355,1379]" italics="true" pageId="34" pageNumber="35">
|
||||
<taxonomicName id="4C784D66FF832E06FD1C1468FCF1F410" baseAuthorityName="Fairmaire & Germain" baseAuthorityYear="1862" box="[648,776,1355,1379]" class="Magnoliopsida" family="Winteraceae" genus="Lonvcovilius" kingdom="Plantae" order="Canellales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="semiflavus">L. semiflavus</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
Presumably, they could have disappeared, as some other material did during a period when this collection was neglected (Camousseight, 1980), while the material from RBINS studied by
|
||||
<bibRefCitation id="EFE94B14FF832E06FF3414EDFE63F492" author="Coiffait H & Saiz F" box="[160,410,1481,1505]" pageId="34" pageNumber="35" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz (1966)</bibRefCitation>
|
||||
are
|
||||
<typeStatus id="54C38847FF832E06FE5214EDFDD9F492" box="[454,544,1481,1505]" pageId="34" pageNumber="35" type="syntype">syntypes</typeStatus>
|
||||
that P. Germain could have shared with A. Fauvel, a prominent colleague of that time. Based on that material,
|
||||
<bibRefCitation id="EFE94B14FF832E06FE1F1723FD6EF76C" author="Coiffait H & Saiz F" box="[395,663,1543,1567]" pageId="34" pageNumber="35" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz (1966)</bibRefCitation>
|
||||
illustrated the aedeagus of
|
||||
<taxonomicName id="4C784D66FF832E06FEBE1703FE53F74C" baseAuthorityName="Fairmaire & Germain" baseAuthorityYear="1862" box="[298,426,1575,1599]" class="Magnoliopsida" family="Winteraceae" genus="Lonvcovilius" kingdom="Plantae" order="Canellales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF832E06FEBE1703FEC0F74D" box="[298,313,1575,1598]" italics="true" pageId="34" pageNumber="35">L</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF832E06FED31703FE53F74C" box="[327,426,1575,1599]" italics="true" pageId="34" pageNumber="35">semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
and, together with
|
||||
<taxonomicName id="4C784D66FF832E06FDED1703FCFDF74C" authorityName="Fairmaire and Germain" authorityYear="1862" box="[633,772,1575,1599]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF832E06FDED1703FD71F74D" box="[633,648,1575,1598]" italics="true" pageId="34" pageNumber="35">L</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF832E06FD021703FCFDF74C" box="[662,772,1575,1599]" italics="true" pageId="34" pageNumber="35">lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, treated it as a member of the subgenus
|
||||
<emphasis id="B90CEAF7FF832E06FD951762FD77F72D" box="[513,654,1606,1630]" italics="true" pageId="34" pageNumber="35">
|
||||
<taxonomicName id="4C784D66FF832E06FD951762FD96F72D" box="[513,623,1606,1630]" pageId="34" pageNumber="35" rank="subGenus" subGenus="Loncovilius">Loncovilius</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF832E06FDE71763FD77F72D" box="[627,654,1607,1630]" pageId="34" pageNumber="35" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
. Apart from the types, a specimen from La Jaula in Curicó Province of
|
||||
<collectingCountry id="F36F7675FF832E06FD441741FCF1F70E" box="[720,776,1637,1661]" name="Chile" pageId="34" pageNumber="35">Chile</collectingCountry>
|
||||
listed in
|
||||
<bibRefCitation id="EFE94B14FF832E06FF4F17A1FEAAF7EE" author="Saiz F" box="[219,339,1669,1693]" pageId="34" pageNumber="35" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz (1971)</bibRefCitation>
|
||||
, was the only additional material reported for this species prior to our revision.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,112 +0,0 @@
|
|||
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<mods:title id="B05D381E4695A5E308DCADECCBA22778">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<paragraph id="8BC736E5FF812E04FEFB1086FDE2F0CE" blockId="32.[367,539,418,445]" box="[367,539,418,445]" pageId="32" pageNumber="33">
|
||||
<heading id="D08F8189FF812E04FEFB1086FDE2F0CE" bold="true" box="[367,539,418,445]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="33" reason="2">
|
||||
<emphasis id="B90CEAF7FF812E04FEFB1086FDE2F0CE" bold="true" box="[367,539,418,445]" pageId="32" pageNumber="33">
|
||||
<taxonomicName id="4C784D66FF812E04FEFB1086FE2EF0CE" ID-CoL="72MP2" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[367,471,418,445]" class="Insecta" family="Staphylinidae" genus="Loncovilius" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF812E04FEFB1086FE2EF0CE" bold="true" box="[367,471,418,445]" italics="true" pageId="32" pageNumber="33">semiflavus</emphasis>
|
||||
</taxonomicName>
|
||||
group
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF812E04FF1510C7FE31F368" pageId="32" pageNumber="33" type="description">
|
||||
<paragraph id="8BC736E5FF812E04FF1510C7FE31F368" blockId="32.[129,777,483,539]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B90CEAF7FF812E04FF1510C7FE0DF088" box="[129,500,483,507]" italics="true" pageId="32" pageNumber="33">
|
||||
Included species:
|
||||
<taxonomicName id="4C784D66FF812E04FEBB10C7FE0DF088" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[303,500,483,507]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">Loncovilius barclayi</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A23F578CFF812E04FE6A10C0FDB7F088" box="[510,590,484,507]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C784D66FF812E04FD1C10C0FE3DF368" authority="(Fairmaire and Germain, 1862)" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="semiflavus">
|
||||
<emphasis id="B90CEAF7FF812E04FD1C10C0FD6EF088" box="[648,663,484,507]" italics="true" pageId="32" pageNumber="33">L</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF812E04FD3210C7FCF0F088" box="[678,777,483,507]" italics="true" pageId="32" pageNumber="33">semiflavus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF812E04FF1F1327FE41F368" author="Fairmaire L & Germain P" box="[139,440,515,539]" pageId="32" pageNumber="33" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF812E04FF151319FD7EF2BF" pageId="32" pageNumber="33" type="diagnosis">
|
||||
<paragraph id="8BC736E5FF812E04FF151319FD7EF2BF" blockId="32.[128,778,572,972]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B90CEAF7FF812E04FF151319FF1EF327" box="[129,231,573,596]" italics="true" pageId="32" pageNumber="33">Diagnosis:</emphasis>
|
||||
Head about as wide as long, without coarse non-setiferous punctures; nuchal ridge absent dorsally, nuchal and infraorbital ridges joined; gula without isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin; PMP (usually reduced); females with antennomere 11 distinctly longer than antennomere 10 (a11:a10 ratio>1.5). Pronotum outline evenly curved, with one SLSP. Protibiae not sexually dimorphic, in both sexes without laterodorsal row of thick spines. Tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area near apical margin (
|
||||
<figureCitation id="13432A60FF812E04FE6F1251FDBEF2FE" box="[507,583,885,909]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="32" pageNumber="33">Fig. 9D</figureCitation>
|
||||
); tergite VIII with medial apical emargination in both sexes; ovipositor, its second gonocoxite with one macroseta medially (
|
||||
<figureCitation id="13432A60FF812E04FDBF1290FD8FF2BF" box="[555,630,948,972]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="32" pageNumber="33">Fig. 9G</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,343 +0,0 @@
|
|||
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<mods:title id="596529CA5D2086D7BA59DEA1CF053C8D">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="98DBA76E441147F2B0F42766764BE6AF">Reyes-Hernández, José L.</mods:namePart>
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<mods:affiliation id="86F82622885BEACA5CE91B896FFD1A73">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="5A027CD8D720741B816FC5239A1346CE" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
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</mods:name>
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</mods:role>
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<mods:namePart id="91CB7BA5C7675D3AD503703E26C8A559">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="60E0DA68202183D29EF82CCC5DF03094">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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</mods:role>
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<mods:namePart id="A5112FB422FDA9F898C39FACCA4D4CBD">Shaw, Josh Jenkins</mods:namePart>
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<mods:affiliation id="51CEEAA7404AEEBB1D44C1D9499887CB">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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</mods:role>
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<mods:namePart id="2A3DE6D17DAAC4809E37F3625D8C628E">Solodovnikov, Alexey</mods:namePart>
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<mods:affiliation id="2530A2B704D1C1A09A7B53AAF403D617">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:title id="73B5D2FCDD7EE672314084AD1DD8F54D">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="5893F5EC8139780B1B3FBF0051F1CCA6">2024</mods:date>
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<mods:title id="A640D5BDD34E4CDFA6E69295BC90D188">Zool. J. Linn. Soc.</mods:title>
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<mods:identifier id="6002E70B827AEC3474DBCBD4E037AB10" type="ZooBank">CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding</mods:identifier>
|
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<treatment id="03D187F3FF812E05FEAC12D2FE55F23C" LSID="urn:lsid:plazi:treatment:03D187F3FF812E05FEAC12D2FE55F23C" httpUri="http://treatment.plazi.org/id/03D187F3FF812E05FEAC12D2FE55F23C" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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<subSubSection id="C362656EFF812E04FEAC12D2FDA8F562" box="[312,593,1014,1041]" pageId="32" pageNumber="33" type="nomenclature">
|
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<heading id="D08F8189FF812E04FEAC12D2FDA8F562" bold="true" box="[312,593,1014,1041]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="33" reason="2">
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<emphasis id="B90CEAF7FF812E04FEAC12D2FDA8F562" bold="true" box="[312,593,1014,1041]" pageId="32" pageNumber="33">
|
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<taxonomicName id="4C784D66FF812E04FEAC12D2FDF8F562" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[312,513,1014,1041]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
|
||||
<emphasis id="B90CEAF7FF812E04FEAC12D2FDF8F562" bold="true" box="[312,513,1014,1041]" italics="true" pageId="32" pageNumber="33">Loncovilius barclayi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF812E04FD9112D3FDA8F562" box="[517,593,1015,1041]" pageId="32" pageNumber="33" rank="species">sp.nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF812E05FE851511FE55F23C" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
|
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<paragraph id="8BC736E5FF812E04FE851511FD83F53C" blockId="32.[273,634,1077,1104]" box="[273,634,1077,1104]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B90CEAF7FF812E04FE851511FD83F53C" bold="true" box="[273,634,1077,1104]" pageId="32" pageNumber="33">
|
||||
(
|
||||
<figureCitation id="13432A60FF812E04FE8F1511FE95F53C" box="[283,364,1077,1103]" captionStart="Figure 6" captionStartId="14.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@14.[168,1434,144,1840]" captionTargetPageId="14" captionText="Figure 6. Habitus of Loncovilius species. A, male of L. edwardsianus. B, male of L. cantharoides. C, female of L. semiflavus. D, male of L. barclayi." pageId="32" pageNumber="33">Figs 6D</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FF812E04FEE11511FE3BF523" box="[373,450,1077,1104]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="32" pageNumber="33">11D–F</figureCitation>
|
||||
, 12H, 13H, 14D)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF812E04FF151552FE09F5DE" blockId="32.[129,777,1142,1197]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B90CEAF7FF812E04FF151552FE97F5FD" box="[129,366,1142,1166]" italics="true" pageId="32" pageNumber="33">Zoobank registration:</emphasis>
|
||||
<uri id="FFE93AE7FF812E04FE041552FE15F5DE" pageId="32" pageNumber="33">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFDE0C30FF812E04FD1B1552FE15F5DE" pageId="32" pageNumber="33">B426E064- 0842-4911-A48A-C7A73A73B5E0</uuid>
|
||||
</uri>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF812E04FF1515EBFE5BF6C4" blockId="32.[128,779,1231,1975]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B90CEAF7FF812E04FF1515EBFEE8F594" box="[129,273,1231,1255]" italics="true" pageId="32" pageNumber="33">Type material.</emphasis>
|
||||
<materialsCitation id="3B103CB8FF812E04FE8D15EBFD88F437" collectingDate="1986-01-10" collectorName="L. E. Pena" country="Chile" location="Valdivia" municipality="Purulon" pageId="32" pageNumber="33" specimenCode="FMNH-INS 0000 023 665" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<emphasis id="B90CEAF7FF812E04FE8D15EBFE80F594" box="[281,377,1231,1255]" italics="true" pageId="32" pageNumber="33">
|
||||
<typeStatus id="54C38847FF812E04FE8D15EBFE8DF594" box="[281,372,1231,1255]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
male, pinned, with genitalia in a separate microvial, with labels as follows:
|
||||
<collectingCountry id="F36F7675FF812E04FDDE15CAFD64F475" box="[586,669,1262,1286]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
|
||||
:
|
||||
<location id="8EA7603EFF812E04FD2415CAFCFFF475" LSID="urn:lsid:plazi:treatment:03D187F3FF812E05FEAC12D2FE55F23C:8EA7603EFF812E04FD2415CAFCFFF475" box="[688,774,1262,1286]" country="Chile" municipality="Purulon" name="Valdivia" pageId="32" pageNumber="33">Valdivia</location>
|
||||
:
|
||||
<collectingMunicipality id="6BA3AC9FFF812E04FF151429FF2CF456" box="[129,213,1293,1317]" pageId="32" pageNumber="33">Purulón</collectingMunicipality>
|
||||
[-39.47, -72.69],
|
||||
<date id="FFC61025FF812E04FE121429FE11F456" box="[390,488,1293,1317]" pageId="32" pageNumber="33" value="1986-01-10">
|
||||
<collectingDate id="EF82E9CDFF812E04FE121429FE11F456" box="[390,488,1293,1317]" pageId="32" pageNumber="33" value="1986-01-10">10.I.1986</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FF812E04FE78142AFDB6F456" box="[492,591,1294,1317]" pageId="32" pageNumber="33">L.E. Peña</collectorName>
|
||||
, leg. FIELD MUS. NAT. HIST./
|
||||
<specimenCount id="9D7EFD6CFF812E04FE801414FED7F437" box="[276,302,1328,1348]" count="1" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFF812E04FEA31409FD88F437" box="[311,625,1324,1349]" pageId="32" pageNumber="33">FMNH-INS 0000 023 665</specimenCode>
|
||||
</materialsCitation>
|
||||
/
|
||||
<typeStatus id="54C38847FF812E04FDED1408FCF0F437" box="[633,777,1324,1348]" pageId="32" pageNumber="33" type="holotype">HOLOTYPE</typeStatus>
|
||||
<taxonomicName id="4C784D66FF812E04FF151468FEB3F417" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[129,330,1356,1380]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi">
|
||||
<emphasis id="B90CEAF7FF812E04FF151468FEB3F417" box="[129,330,1356,1380]" italics="true" pageId="32" pageNumber="33">Loncovilius barclayi</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
|
||||
<materialsCitation id="3B103CB8FF812E04FD361448FF51F492" collectingDate="1986-01-10" collectorName="L. E. Pena" country="Chile" location="Purulon" municipality="Valdivia" pageId="32" pageNumber="33" specimenCode="FMNH-INS 0000 023 667" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||||
<emphasis id="B90CEAF7FF812E04FD361448FCF0F4F0" box="[674,777,1388,1411]" italics="true" pageId="32" pageNumber="33">
|
||||
<typeStatus id="54C38847FF812E04FD361448FCFDF4F0" box="[674,772,1388,1411]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
<collectingCountry id="F36F7675FF812E04FF1514AFFF2DF4D0" box="[129,212,1419,1443]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingMunicipality id="6BA3AC9FFF812E04FF7714AFFEC0F4D0" box="[227,313,1419,1443]" pageId="32" pageNumber="33">Valdivia</collectingMunicipality>
|
||||
:
|
||||
<location id="8EA7603EFF812E04FEDC14AFFE65F4D0" LSID="urn:lsid:plazi:treatment:03D187F3FF812E05FEAC12D2FE55F23C:8EA7603EFF812E04FEDC14AFFE65F4D0" box="[328,412,1419,1443]" country="Chile" municipality="Valdivia" name="Purulon" pageId="32" pageNumber="33">Purulón</location>
|
||||
[-39.47, -72.69],
|
||||
<date id="FFC61025FF812E04FDF114AFFD3EF4D0" box="[613,711,1419,1443]" pageId="32" pageNumber="33" value="1986-01-10">
|
||||
<collectingDate id="EF82E9CDFF812E04FDF114AFFD3EF4D0" box="[613,711,1419,1443]" pageId="32" pageNumber="33" value="1986-01-10">10.I.1986</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FF812E04FD4714AFFF4CF4B2" pageId="32" pageNumber="33">L. E. Peña</collectorName>
|
||||
, leg. FIELD MUS. NAT. HIST./
|
||||
<specimenCount id="9D7EFD6CFF812E04FE691489FDEEF4B2" box="[509,535,1453,1473]" count="1" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFF812E04FDB4148EFF51F492" pageId="32" pageNumber="33">FMNH-INS 0000 023 667</specimenCode>
|
||||
</materialsCitation>
|
||||
/
|
||||
<taxonomicName id="4C784D66FF812E04FF3B14EDFD95F492" authority="J. L. Reyes-Hernandez" authorityName="J. L. Reyes-Hernandez" box="[175,620,1481,1505]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi">Loncovilius barclayi J. L. Reyes-Hernández</taxonomicName>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFF812E04FDE814EDFD5AF493" box="[636,675,1481,1505]" count="1" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
|
||||
FMNH];
|
||||
<materialsCitation id="3B103CB8FF812E04FF1514CDFEFFF72D" collectingDate="1986-01-10" collectingDateMax="1993-01-20" collectingDateMin="1986-01-10" collectionCode="NHM, FMNH" collectorName="L. E. Pena & Coll. T Curkovic" country="Chile" county="Valdivia" determinerName="Newton" location="Purulon" municipality="Nahuelbuta" pageId="32" pageNumber="33" specimenCode="FMNH-INS 0000 023 666" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF812E04FF1514CDFF2DF772" box="[129,212,1513,1537]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingCounty id="62A64E69FF812E04FF7714CDFEC0F772" box="[227,313,1513,1537]" pageId="32" pageNumber="33">Valdivia</collectingCounty>
|
||||
:
|
||||
<location id="8EA7603EFF812E04FEDD14CDFE64F772" LSID="urn:lsid:plazi:treatment:03D187F3FF812E05FEAC12D2FE55F23C:8EA7603EFF812E04FEDD14CDFE64F772" box="[329,413,1513,1537]" country="Chile" county="Valdivia" municipality="Nahuelbuta" name="Purulon" pageId="32" pageNumber="33">Purulón</location>
|
||||
[-39.47, -72.69],
|
||||
<date id="FFC61025FF812E04FDF114CDFD3EF772" box="[613,711,1513,1537]" pageId="32" pageNumber="33" value="1986-01-10">
|
||||
<collectingDate id="EF82E9CDFF812E04FDF114CDFD3EF772" box="[613,711,1513,1537]" pageId="32" pageNumber="33" value="1986-01-10">10.I.1986</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FF812E04FD4014CDFF4CF76C" pageId="32" pageNumber="33">L. E. Peña</collectorName>
|
||||
, leg. FIELD MUS. NAT. HIST./
|
||||
<specimenCount id="9D7EFD6CFF812E04FE69172FFDEEF76C" box="[509,535,1547,1567]" count="1" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFF812E04FDB4172CFF51F74C" pageId="32" pageNumber="33">FMNH-INS 0000 023 666</specimenCode>
|
||||
/
|
||||
<taxonomicName id="4C784D66FF812E04FF241703FD78F74C" authority="J. L. Reyes-Hernandez" authorityName="J. L. Reyes-Hernandez" box="[176,641,1575,1599]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi">Loncovilius barclayi J. L. Reyes-Hernández</taxonomicName>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFF812E04FD021703FD47F74D" box="[662,702,1575,1599]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FF812E04FD5D170CFCF0F74C" box="[713,777,1576,1599]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="32" pageNumber="33">NHM</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FF812E04FF321763FF05F72D" box="[166,252,1607,1630]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="32" pageNumber="33">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF812E04FE9A1763FF71F7EE" collectingDate="1986-01-10" collectingDateMax="1993-01-20" collectingDateMin="1986-01-10" collectionCode="NHM, FMNH" collectorName="L. E. Pena & Coll. T Curkovic" country="Chile" county="Valdivia" determinerName="Newton" location="Purulon" municipality="Nahuelbuta" pageId="32" pageNumber="33" specimenCode="FMNH-INS 0000 023 666" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
|
||||
<collectingMunicipality id="6BA3AC9FFF812E04FE9A1763FE7FF72C" box="[270,390,1607,1631]" pageId="32" pageNumber="33">Nahuelbuta</collectingMunicipality>
|
||||
IX
|
||||
<location id="8EA7603EFF812E04FE251763FDC4F72C" LSID="urn:lsid:plazi:treatment:03D187F3FF812E05FEAC12D2FE55F23C:8EA7603EFF812E04FE251763FDC4F72C" box="[433,573,1607,1631]" country="Chile" county="Valdivia" municipality="Nahuelbuta" name="Region-Chile" pageId="32" pageNumber="33">Region-Chile</location>
|
||||
,
|
||||
<date id="FFC61025FF812E04FDDE1763FD48F72C" box="[586,689,1607,1631]" pageId="32" pageNumber="33" value="1993-01-20">
|
||||
<collectingDate id="EF82E9CDFF812E04FDDE1763FD48F72C" box="[586,689,1607,1631]" pageId="32" pageNumber="33" value="1993-01-20">20-I-1993</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="268D5333FF812E04FD291763FF1BF70D" pageId="32" pageNumber="33">Coll. T Curkovic</collectorName>
|
||||
/
|
||||
<taxonomicName id="4C784D66FF812E04FF7E1742FE1CF70D" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[234,485,1638,1662]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="semiflavus">Loncovilius? semiflavus</taxonomicName>
|
||||
det.
|
||||
<determinerName id="7E1BDA21FF812E04FDB11742FD80F70E" box="[549,633,1638,1661]" pageId="32" pageNumber="33">Newton</determinerName>
|
||||
2000 [
|
||||
<specimenCount id="9D7EFD6CFF812E04FD441742FD0EF70E" box="[720,759,1638,1662]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF812E04FF1317A1FEE3F7A8" collectingDate="1981-01-22" collectionCode="MNHN, MNNC" collectorName="Valdivia & Purulon & Pena" country="Chile" determinerName="Newton" location="Nahuelbuta Parque" municipality="Nahuelbuta" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Region Metropolitana" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF812E04FF1317A1FF19F7EE" box="[135,224,1669,1693]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
|
||||
:] COLL. CERDA
|
||||
<collectionCode id="ED69AE20FF812E04FE5917A2FDD8F7EE" box="[461,545,1670,1693]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="32" pageNumber="33">MNHN</collectionCode>
|
||||
CHILE/
|
||||
<location id="8EA7603EFF812E04FD1917A1FF30F7CF" LSID="urn:lsid:plazi:treatment:03D187F3FF812E05FEAC12D2FE55F23C:8EA7603EFF812E04FD1917A1FF30F7CF" country="Chile" municipality="Nahuelbuta" name="Nahuelbuta Parque" pageId="32" pageNumber="33" stateProvince="Region Metropolitana">Nahuelbuta Parque</location>
|
||||
[-37.80, -73.01]
|
||||
<date id="FFC61025FF812E04FEE71781FE25F7CE" box="[371,476,1701,1725]" pageId="32" pageNumber="33" value="1981-01-22">
|
||||
<collectingDate id="EF82E9CDFF812E04FEE71781FE25F7CE" box="[371,476,1701,1725]" pageId="32" pageNumber="33" value="1981-01-22">22.1.1981</collectingDate>
|
||||
</date>
|
||||
. P. [illegible] B/Santiago Mus [
|
||||
<collectionCode id="ED69AE20FF812E04FF1217E0FF25F7AF" box="[134,220,1732,1756]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="32" pageNumber="33">MNNC</collectionCode>
|
||||
<specimenCount id="9D7EFD6CFF812E04FF7317E0FEF7F7A8" box="[231,270,1732,1756]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
|
||||
]
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B103CB8FF812E04FEBF17E0FF08F649" collectingDate="2005-01-25" collectingDateMax="2005-01-31" collectingDateMin="2005-01-25" collectionCode="NHMD, NMPC" collectorName="Sv Bily" country="Chile" elevation="1200" location="Nahuelbuta National Park" pageId="32" pageNumber="33" specimenCode="916914" specimenCount="1" specimenCount-male="1" stateProvince="P. N. Nahuelbula" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF812E04FEBF17E0FE82F7AF" box="[299,379,1732,1756]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
|
||||
, IX r.,
|
||||
<date id="FFC61025FF812E04FE5F17E0FDB7F7AF" box="[459,590,1732,1756]" pageId="32" pageNumber="33" value="2005-01-25" valueMax="2005-01-31" valueMin="2005-01-25">
|
||||
<collectingDate id="EF82E9CDFF812E04FE5F17E0FDB7F7AF" box="[459,590,1732,1756]" pageId="32" pageNumber="33" value="2005-01-25" valueMax="2005-01-31" valueMin="2005-01-25">25-31.i.2005</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectingRegion id="49BCF807FF812E04FDC917E0FCF0F7AF" box="[605,777,1732,1756]" pageId="32" pageNumber="33">P.N. Nahuelbula</collectingRegion>
|
||||
[
|
||||
<collectingRegion id="49BCF807FF812E04FF1217C7FF0AF788" box="[134,243,1763,1787]" country="Chile" name="Araucania" pageId="32" pageNumber="33">Araucanía</collectingRegion>
|
||||
,
|
||||
<location id="8EA7603EFF812E04FE9617C7FDE5F788" LSID="urn:lsid:plazi:treatment:03D187F3FF812E05FEAC12D2FE55F23C:8EA7603EFF812E04FE9617C7FDE5F788" box="[258,540,1763,1787]" country="Chile" name="Nahuelbuta National Park" pageId="32" pageNumber="33" stateProvince="P. N. Nahuelbula">Nahuelbuta National Park</location>
|
||||
]
|
||||
<geoCoordinate id="EE4C5022FF812E04FDB917C7FD6CF788" box="[557,661,1762,1787]" pageId="32" pageNumber="33">
|
||||
37°48
|
||||
<emphasis id="B90CEAF7FF812E04FDFD17C6FD96F788" box="[617,623,1762,1787]" italics="true" pageId="32" pageNumber="33">ʹ</emphasis>
|
||||
530
|
||||
</geoCoordinate>
|
||||
<geoCoordinate id="EE4C5022FF812E04FD3417C7FCF0F788" box="[672,777,1762,1787]" pageId="32" pageNumber="33">
|
||||
73°00
|
||||
<emphasis id="B90CEAF7FF812E04FD4917C6FD1AF788" box="[733,739,1762,1787]" italics="true" pageId="32" pageNumber="33">ʹ</emphasis>
|
||||
954
|
||||
</geoCoordinate>
|
||||
[-37.80, -73.01],
|
||||
<quantity id="4C809B00FF812E04FEAD1627FE77F669" box="[313,398,1795,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="33" unit="m" value="1200.0">
|
||||
<elevation id="0055D1D6FF812E04FEAD1627FE77F669" box="[313,398,1795,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="33" unit="m" value="1200.0">1200 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<collectorName id="268D5333FF812E04FE091627FE11F668" box="[413,488,1795,1819]" pageId="32" pageNumber="33">Sv Bíly</collectorName>
|
||||
lgt./
|
||||
<collectionCode id="ED69AE20FF812E04FD891627FD8DF668" box="[541,628,1795,1819]" collectionName="NHMD" pageId="32" pageNumber="33">NHMD</collectionCode>
|
||||
<specimenCode id="DBDE9E9EFF812E04FDEA1627FD35F668" box="[638,716,1795,1819]" pageId="32" pageNumber="33">916914</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFF812E04FD751627FCF0F669" box="[737,777,1795,1819]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FF812E04FF151606FF2CF649" box="[129,213,1826,1850]" collectionName="Czech Republic, Prague, National Museum (Natural History)" pageId="32" pageNumber="33">NMPC</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF812E04FF651606FE9EF60B" collectingDate="2005-01-25" collectingDateMax="2005-01-31" collectingDateMin="2005-01-25" collectionCode="MNNC, FMNH" collectorName="Fairmaire" country="Chile" elevation="1200" location="CHILE" pageId="32" pageNumber="33" specimenCode="FMNH-INS 0000 024 190" specimenCount="1" specimenCount-male="1" stateProvince="Araucania" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF812E04FF651606FEBEF649" box="[241,327,1826,1850]" name="Chile" pageId="32" pageNumber="33">CHILE</collectingCountry>
|
||||
: no exact location] col.
|
||||
<collectorName id="268D5333FF812E04FDCE1606FD3FF649" box="[602,710,1826,1850]" pageId="32" pageNumber="33">Fairmaire</collectorName>
|
||||
/1251/
|
||||
<taxonomicName id="4C784D66FF812E04FF151665FEA8F62A" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[129,337,1857,1881]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="semiflavus" status="sp. nov.">Quedius semiflavus</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF812E04FEF41665FE75F62A" box="[352,396,1857,1881]" pageId="32" pageNumber="33" rank="species">n.sp.</taxonomicNameLabel>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFF812E04FE071665FD30F62A" box="[403,713,1857,1881]" pageId="32" pageNumber="33">FMNH-INS 0000 024 190</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFF812E04FD751665FCF0F62B" box="[737,777,1857,1881]" count="1" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FF812E04FF151645FF2AF60A" box="[129,211,1889,1913]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="32" pageNumber="33">MNNC</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FF812E04FE9C1645FEA4F60B" box="[264,349,1889,1912]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="32" pageNumber="33">FMNH</collectionCode>
|
||||
].
|
||||
</materialsCitation>
|
||||
All
|
||||
<typeStatus id="54C38847FF812E04FE091645FDF9F60A" box="[413,512,1889,1913]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
|
||||
with label: PARATYPE
|
||||
<taxonomicName id="4C784D66FF812E04FF1516A4FEB3F6EB" box="[129,330,1920,1944]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="barclayi">
|
||||
<emphasis id="B90CEAF7FF812E04FF1516A4FEB3F6EB" box="[129,330,1920,1944]" italics="true" pageId="32" pageNumber="33">Loncovilius barclayi</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF812E04FCAD11B5FB1BF076" blockId="32.[825,1474,144,262]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B90CEAF7FF812E04FCAD11B5FC66F1DB" box="[825,927,145,168]" italics="true" pageId="32" pageNumber="33">Diagnosis:</emphasis>
|
||||
Elytra yellow or yellowish-brown with black marks around mesoscutellum and on suture. Male tergite X trilobed (
|
||||
<figureCitation id="13432A60FF812E04FCD011EBFC66F194" box="[836,927,207,231]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="32" pageNumber="33">Fig. 12H</figureCitation>
|
||||
); female tergite X deeply emarginate medio-apically (
|
||||
<figureCitation id="13432A60FF812E04FCD011CAFC64F075" box="[836,925,238,262]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="32" pageNumber="33">Fig. 13H</figureCitation>
|
||||
). Aedeagus as in
|
||||
<figureCitation id="13432A60FF812E04FBDA11CAFB25F075" box="[1102,1244,238,262]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="32" pageNumber="33">Figure 11D–F</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF812E04FCAD101FFB15F09C" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">
|
||||
<emphasis id="B90CEAF7FF812E04FCAD101FFC49F021" box="[825,944,315,338]" italics="true" pageId="32" pageNumber="33">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFF812E04FBF5101AFB82F021" box="[1121,1147,318,338]" count="1" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FF812E04FAE7101FFA7FF021" box="[1395,1414,315,338]" italics="true" pageId="32" pageNumber="33">N</emphasis>
|
||||
= 6]: FBL = 2.42–3.86 (3.56); TL = 7.13–7.5 (7.32); HW = 0.9– 0.98 (0.95); HL = 0.9–0.98 (0.95); HW/HL = 0.99–1.03 (1); PW = 1.19–1.31 (1.25); PL = 1.25–1.3 (1.28); PW/ PL = 0.93–1.02 (0.98); EW = 1.54–1.75 (1.64); EL = 1.45– 1.65 (1.56); PW/HW = 1.28–1.34 (1.31).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF812E04FCC110D3FAAEF33E" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFF812E04FB9F10DEFBDCF37D" box="[1035,1061,506,526]" count="1" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||||
(
|
||||
<emphasis id="B90CEAF7FF812E04FBC510D3FB9DF37D" box="[1105,1124,503,526]" italics="true" pageId="32" pageNumber="33">N</emphasis>
|
||||
= 1): FBL = 4.03; TL = 7.5; HW = 1.05; HL = 1; HW/HL = 1.05; PW = 1.38; PL = 1.38; PW/PL = 1; EW = 1.7; EL = 1.65; PW/HW = 1.31.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF812E04FCC11371FC3FF27A" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">Head and pronotum yellow-orange; antennae, mouthparts, and legs yellow or yellowish-brown; elytra yellow or yellowish-brown with dark mark around mesoscutellum and on suture; meso- and metasternum, all coxae, femora and abdomen dark reddish-brown; abdominal tergites and sternites with pale posterior margin.</paragraph>
|
||||
<paragraph id="8BC736E5FF812E04FCC11235FC14F4C8" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">
|
||||
Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small-size. Eyes medium size (EYL/HL = x = 0.52), from 1.4 to 1.55 times longer than length of temples (in lateral view); distance between eyes about equal to length of eye. Antennomeres 2 and 3 subequal in length; antennomeres 4 to 6 subequal in length; antennomeres 6 to 10 gradually reduced in length; a11 1.75 to 1.88 times as long as a
|
||||
<quantity id="4C809B00FF812E04FC0C156EFC37F511" box="[920,974,1098,1122]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="32" pageNumber="33" unit="in" value="10.0">10 in</quantity>
|
||||
males and
|
||||
<quantity id="4C809B00FF812E04FBDC156EFB68F511" box="[1096,1169,1098,1122]" metricMagnitude="-2" metricUnit="m" metricValue="4.2926" pageId="32" pageNumber="33" unit="in" value="1.69">1.69 in</quantity>
|
||||
females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; ventral basal ridge more or less straight in checkmark shape, almost united with gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present only laterally, fused with infraorbital ridge; gular sutures moderately separated. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF812E04FCC114E6FA42F704" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">Pronotum about as wide as long, strongly convex, evenly curved, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (or process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to two or four times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
|
||||
<paragraph id="8BC736E5FF812E04FCC1175AFABCF621" blockId="32.[823,1476,315,1968]" pageId="32" pageNumber="33">Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate.</paragraph>
|
||||
<paragraph id="8BC736E5FF812E05FCC1167DFE02F0D0" blockId="32.[823,1476,315,1968]" lastBlockId="33.[113,763,144,607]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||||
Abdominal tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area; tergite VIII distinctly emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with strong, V-shaped emargination medially (
|
||||
<figureCitation id="13432A60FF802E05FF3D11CAFEFFF075" box="[169,262,238,262]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="33" pageNumber="34">Fig. 14H</figureCitation>
|
||||
); female sternite VIII with weak apical medial emargination; male sternite IX not emarginate medio-apically, its basal portion distinctly longer than distal portion; male tergite X trilobed (
|
||||
<figureCitation id="13432A60FF802E05FE8F1068FE8FF017" box="[283,374,332,356]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="33" pageNumber="34">Fig. 12H</figureCitation>
|
||||
); female tergite X deeply emarginate medio-apically (
|
||||
<figureCitation id="13432A60FF802E05FE8A104FFE84F0F0" box="[286,381,363,387]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="33" pageNumber="34">Fig. 13H</figureCitation>
|
||||
); ovipositor, its second gonocoxite with one macroseta medially (
|
||||
<figureCitation id="13432A60FF802E05FE3110AFFE13F0D0" box="[421,490,395,419]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="33" pageNumber="34">Fig 9G</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF802E05FF19108EFD02F32C" blockId="33.[113,763,144,607]" pageId="33" pageNumber="34">
|
||||
Aedeagus as in
|
||||
<figureCitation id="13432A60FF802E05FEB8108EFE43F0B1" box="[300,442,426,450]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="33" pageNumber="34">Figure 11D–F</figureCitation>
|
||||
, its length ~ 1.46; median lobe rod-like, in parameral view apex deeply emarginate; internal sac with pair of long I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging into a deeply emarginate apex, with dark peg setae, in lateral view almost straight and slightly concave apically.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF802E05FFE513A4FD03F3C4" blockId="33.[113,763,640,695]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FFE513A4FF27F3EB" box="[113,222,640,664]" italics="true" pageId="33" pageNumber="34">Etymology:</emphasis>
|
||||
The species epithet is a patronym in recognition of Maxwell V.L.Barclay, in particular for his earlier mentoring of J.J.S..
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF802E05FFE513FDFE55F23C" blockId="33.[113,763,729,847]" pageId="33" pageNumber="34">
|
||||
<emphasis id="B90CEAF7FF802E05FFE513FDFDD7F382" box="[113,558,729,753]" italics="true" pageId="33" pageNumber="34">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FF802E05FEF913FDFDD7F382" box="[365,558,729,753]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="barclayi">Loncovilius barclayi</taxonomicName>
|
||||
</emphasis>
|
||||
is known only from
|
||||
<collectingCountry id="F36F7675FF802E05FFE613DCFF55F263" box="[114,172,760,784]" name="Chile" pageId="33" pageNumber="34">Chile</collectingCountry>
|
||||
, from the
|
||||
<collectingRegion id="49BCF807FF802E05FE8E13DCFE78F263" box="[282,385,760,784]" country="Chile" name="Araucania" pageId="33" pageNumber="34">Araucanía</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="49BCF807FF802E05FE2313DCFDEDF263" box="[439,532,760,784]" country="Chile" name="Los Rios" pageId="33" pageNumber="34">Los Ríos</collectingRegion>
|
||||
regions, in the north-central portion of the Valdivian temperate forest ecoregion. It is unknown how it was collected.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,349 +0,0 @@
|
|||
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<mods:title id="025E1F54FD6E4A6B7180FE697FDA8FE4">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:affiliation id="DCD0FD4E8D9F3BEF7C67772EB92F6D45">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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|
||||
<mods:identifier id="A45AFE61A29AD533D1F7A320419856D5" type="Zenodo-Dep">14541979</mods:identifier>
|
||||
<mods:identifier id="72782EBEA2840F77CF60FE3205970D07" type="ZooBank">CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03D187F3FF822E00FCC41098FA3AF32C" LSID="urn:lsid:plazi:treatment:03D187F3FF822E00FCC41098FA3AF32C" httpUri="http://treatment.plazi.org/id/03D187F3FF822E00FCC41098FA3AF32C" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
|
||||
<subSubSection id="C362656EFF822E07FCC41098FA74F0A5" box="[848,1421,444,471]" pageId="35" pageNumber="36" type="nomenclature">
|
||||
<paragraph id="8BC736E5FF822E07FCC41098FA74F0A5" blockId="35.[848,1421,444,471]" box="[848,1421,444,471]" pageId="35" pageNumber="36">
|
||||
<heading id="D08F8189FF822E07FCC41098FA74F0A5" bold="true" box="[848,1421,444,471]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="36" reason="2">
|
||||
<taxonomicName id="4C784D66FF822E07FCC41098FA74F0A5" ID-CoL="3VYHS" authority="(Fairmaire and Germain, 1862)" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[848,1421,444,471]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF822E07FCC41098FA74F0A5" bold="true" box="[848,1421,444,471]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B90CEAF7FF822E07FCC41098FBC2F0A4" bold="true" box="[848,1083,444,471]" italics="true" pageId="35" pageNumber="36">Loncovilius lividipennis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF822E07FBDD1099FA7BF0A4" author="Fairmaire L & Germain P" box="[1097,1410,445,471]" pageId="35" pageNumber="36" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF822E07FC2310C8FADEF375" box="[951,1319,492,518]" pageId="35" pageNumber="36" type="description">
|
||||
<paragraph id="8BC736E5FF822E07FC2310C8FADEF375" blockId="35.[951,1319,492,518]" box="[951,1319,492,518]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B90CEAF7FF822E07FC2310C8FADEF375" bold="true" box="[951,1319,492,518]" pageId="35" pageNumber="36">
|
||||
(
|
||||
<figureCitation id="13432A60FF822E07FC5510C8FBE8F375" box="[961,1041,492,518]" captionStart="Figure 7" captionStartId="15.[113,178,1868,1892]" captionTargetBox="[153,1419,163,1840]" captionTargetId="figure-6@15.[153,1419,144,1840]" captionTargetPageId="15" captionText="Figure 7. Female habitus of Loncovilius species.A, L. lividipennis. B, L. carlsbergi. Head. C, L. hammondi. D, L. semiflavus. E, L. carlsbergi. F, Male mesotarsi of L. cantharoides." pageId="35" pageNumber="36">Figs 7A</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FF822E07FB8E10C8FBC1F375" box="[1050,1080,492,518]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="35" pageNumber="36">9E</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FF822E07FBD610C8FB69F375" box="[1090,1168,492,518]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="35" pageNumber="36">11G, H</figureCitation>
|
||||
, 12I, 13I, 14I)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF822E07FCBE1307FB33F2E0" pageId="35" pageNumber="36" type="reference_group">
|
||||
<paragraph id="8BC736E5FF822E07FCBE1307FB33F2E0" blockId="35.[809,1460,547,915]" pageId="35" pageNumber="36">
|
||||
<bibRefCitation id="EFE94B14FF822E07FCBE1307FB60F348" author="Fairmaire L & Germain P" box="[810,1177,547,571]" pageId="35" pageNumber="36" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain 1862: 430</bibRefCitation>
|
||||
(as
|
||||
<taxonomicName id="4C784D66FF822E07FB581307FA54F348" authorityName="Fairmaire & Germain" authorityYear="1862" box="[1228,1453,547,571]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF822E07FB581307FA54F348" box="[1228,1453,547,571]" italics="true" pageId="35" pageNumber="36">Philonthus lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, original description); Fauvel, 1866: 338 (as
|
||||
<taxonomicName id="4C784D66FF822E07FB7E1366FA57F329" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[1258,1454,578,602]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF822E07FB7E1366FA57F329" box="[1258,1454,578,602]" italics="true" pageId="35" pageNumber="36">Quedius lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, additional characters);
|
||||
<bibRefCitation id="EFE94B14FF822E07FBBC1345FA4AF30A" author="Bernhauer M & Schubert K" box="[1064,1459,609,633]" pageId="35" pageNumber="36" pagination="409 - 98" refId="ref36154" refString="Bernhauer M, Schubert K. In: Staphylinidae V, Schenkling S (eds), Coleopterorum Catalogus 5 (67). Berlin: Junk, 1916, 409 - 98." type="book chapter" year="1916">Bernhauer and Schubert 1916: 427</bibRefCitation>
|
||||
(as
|
||||
<taxonomicName id="4C784D66FF822E07FCC613A5FBE3F3EA" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[850,1050,641,665]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF822E07FCC613A5FBE3F3EA" box="[850,1050,641,665]" italics="true" pageId="35" pageNumber="36">Quedius lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, catalogue); Scheerpeltz, 1972: 23 [as
|
||||
<emphasis id="B90CEAF7FF822E07FCBE1384FB84F3CB" box="[810,1149,672,696]" italics="true" pageId="35" pageNumber="36">
|
||||
<taxonomicName id="4C784D66FF822E07FCBE1384FB8EF3CB" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[810,1143,672,696]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis" subGenus="Loncovilius">Quedius (Loncovilius) lividipennis</taxonomicName>
|
||||
;
|
||||
</emphasis>
|
||||
additional material];
|
||||
<bibRefCitation id="EFE94B14FF822E07FAC31384FC6CF3A4" author="Germain P" pageId="35" pageNumber="36" pagination="391 - 445" refId="ref36747" refString="Germain P. Apuntes Entomolojicos. Rectificaciones, anotaciones complementarias, i descripciones de siete especies nuevas de coleopteros. Anales de la Universidad de Chile 1903; 113: 391 - 445." type="journal article" year="1903">Germain 1903: 443</bibRefCitation>
|
||||
(as
|
||||
<taxonomicName id="4C784D66FF822E07FC50139BFB51F3A4" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[964,1192,703,727]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF822E07FC50139BFB51F3A4" box="[964,1192,703,727]" italics="true" pageId="35" pageNumber="36">Loncovilius lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, characters);
|
||||
<bibRefCitation id="EFE94B14FF822E07FAA3139BFC3EF384" author="Coiffait H & Saiz F" pageId="35" pageNumber="36" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz 1966: 409</bibRefCitation>
|
||||
(as
|
||||
<emphasis id="B90CEAF7FF822E07FC6213FBFB07F384" box="[1014,1278,735,759]" italics="true" pageId="35" pageNumber="36">
|
||||
<taxonomicName id="4C784D66FF822E07FC6213FBFB9DF384" box="[1014,1124,735,759]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" sensu="stricto">Loncovilius</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF822E07FBF913FBFB71F385" box="[1133,1160,735,758]" pageId="35" pageNumber="36" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
<taxonomicName id="4C784D66FF822E07FB0413FBFB07F384" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[1168,1278,735,759]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis">lividipennis</taxonomicName>
|
||||
</emphasis>
|
||||
; characters; additional material); Coiffait and Sáiz, 1968: 365 (as
|
||||
<emphasis id="B90CEAF7FF822E07FAD113DAFC47F246" italics="true" pageId="35" pageNumber="36">
|
||||
<taxonomicName id="4C784D66FF822E07FAD113DAFA4AF265" box="[1349,1459,766,790]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" sensu="stricto">Loncovilius</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF822E07FCBE123AFCBFF246" box="[810,838,798,821]" pageId="35" pageNumber="36" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
<taxonomicName id="4C784D66FF822E07FCC41239FC47F246" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[848,958,797,821]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis">lividipennis</taxonomicName>
|
||||
</emphasis>
|
||||
, checklist);
|
||||
<bibRefCitation id="EFE94B14FF822E07FBD31239FB10F246" author="Saiz F" box="[1095,1257,797,821]" pageId="35" pageNumber="36" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz 1971: 385</bibRefCitation>
|
||||
(as
|
||||
<emphasis id="B90CEAF7FF822E07FA8B1239FC61F226" italics="true" pageId="35" pageNumber="36">
|
||||
<taxonomicName id="4C784D66FF822E07FA8B1239FA74F246" box="[1311,1421,797,821]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" sensu="stricto">Loncovilius</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF822E07FA0C123AFA4DF246" box="[1432,1460,798,821]" pageId="35" pageNumber="36" sensu="stricto">s.s.</taxonomicNameLabel>
|
||||
<taxonomicName id="4C784D66FF822E07FCBE1219FC61F226" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[810,920,829,853]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis">lividipennis</taxonomicName>
|
||||
</emphasis>
|
||||
; additional material);
|
||||
<bibRefCitation id="EFE94B14FF822E07FB031219FA7FF226" author="Herman LH" box="[1175,1414,829,853]" pageId="35" pageNumber="36" pagination="3021 - 840" refId="ref36981" refString="Herman LH. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine Group (Part 3). Staphylininae: Staphylinini: (Quediina + Xanthopygina), Xantholinini. Staphylinidae Incertae Sedis. Fossils, Protactinae †. Bulletin of the American Museum of Natural History 2001 b; 265: 3021 - 840." type="journal article" year="2001">Herman 2001b: 3083</bibRefCitation>
|
||||
(as
|
||||
<taxonomicName id="4C784D66FF822E07FCBE1278FBF3F207" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[810,1034,860,884]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF822E07FCBE1278FBF3F207" box="[810,1034,860,884]" italics="true" pageId="35" pageNumber="36">Loncovilius lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, catalogue);
|
||||
<bibRefCitation id="EFE94B14FF822E07FB1F1278FA4AF207" author="Jenkins Shaw J & Zyla D & Solodovnikov A" box="[1163,1459,860,884]" pageId="35" pageNumber="36" pagination="430 - 46" refId="ref37147" refString="Jenkins Shaw J, Zyla D, Solodovnikov A. Molecular phylogeny illuminates Amblyopinini (Coleoptera: Staphylinidae) rove beetles as a target for systematic and evolutionary research. Systematic Entomology 2020 a; 45: 430 - 46." type="journal article" year="2020">
|
||||
Jenkins Shaw
|
||||
<emphasis id="B90CEAF7FF822E07FA801279FAC5F207" box="[1300,1340,860,884]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
|
||||
., 2020: 337
|
||||
</bibRefCitation>
|
||||
(as
|
||||
<emphasis id="B90CEAF7FF822E07FCDA125FFBBBF2E0" box="[846,1090,891,915]" italics="true" pageId="35" pageNumber="36">
|
||||
<taxonomicName id="4C784D66FF822E07FCDA125FFC45F2E0" box="[846,956,891,915]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Loncovilius</taxonomicName>
|
||||
nr.
|
||||
<taxonomicName id="4C784D66FF822E07FC4B125FFBBBF2E0" baseAuthorityName="Fairmaire & Germain" baseAuthorityYear="1862" box="[991,1090,891,915]" class="Magnoliopsida" family="Winteraceae" genus="Lonvcovilius" kingdom="Plantae" order="Canellales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="semiflavus">semiflavus</taxonomicName>
|
||||
</emphasis>
|
||||
; phylogeny).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF822E07FCBE1291FAC2F2BE" box="[810,1339,949,973]" pageId="35" pageNumber="36" type="materials_examined">
|
||||
<paragraph id="8BC736E5FF822E07FCBE1291FAC2F2BE" blockId="35.[810,1339,949,973]" box="[810,1339,949,973]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B90CEAF7FF822E07FCBE1291FC10F2BE" box="[810,1001,949,973]" italics="true" pageId="35" pageNumber="36">Material examined:</emphasis>
|
||||
Supporting Information, File S7.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF822E07FCBE12CEFAB2F54C" pageId="35" pageNumber="36" type="diagnosis">
|
||||
<paragraph id="8BC736E5FF822E07FCBE12CEFAB2F54C" blockId="35.[809,1459,1001,1088]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B90CEAF7FF822E07FCBE12CEFC69F572" box="[810,912,1002,1025]" italics="true" pageId="35" pageNumber="36">Diagnosis:</emphasis>
|
||||
Pronotum very rarely with APP. Elytra yellow or yellowish-brown. Sternite III with basal transverse carina sharply pointed medioapically. Aedeagus as in
|
||||
<figureCitation id="13432A60FF822E07FB26150CFABDF54C" box="[1202,1348,1064,1088]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="35" pageNumber="36">Figure 11G, H</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF822E00FCBE1579FD22F4FE" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
|
||||
<paragraph id="8BC736E5FF822E07FCBE1579FB29F462" blockId="35.[810,1461,1117,1987]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B90CEAF7FF822E07FCBE1579FC58F507" box="[810,929,1117,1140]" italics="true" pageId="35" pageNumber="36">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFF822E07FBD81544FB9FF507" box="[1100,1126,1120,1140]" count="1" pageId="35" pageNumber="36" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FF822E07FAC31579FA93F507" box="[1367,1386,1117,1140]" italics="true" pageId="35" pageNumber="36">N</emphasis>
|
||||
= 10]: FBL = 3.09–3.36 (3.22); TL = 5.15–6.86 (6.20); HW = 0.79– 0.84 (0.81); HL = 0.78–0.84 (0.81); HW/HL = 0.98–1.02 (1); PW = 1.05–1.17 (1.11); PL = 0.97–1.08 (1.02); PW/ PL = 1.08–1.11 (1.09); EW = 1.38–1.84 (1.56); EL = 1.33– 1.44 (1.4); PW/HW = 1.31–1.42 (1.37).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FCD1143DFB3AF4BE" blockId="35.[810,1461,1117,1987]" pageId="35" pageNumber="36">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFF822E07FB941438FBE3F443" box="[1024,1050,1308,1328]" count="1" pageId="35" pageNumber="36" type="female">♀</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FF822E07FAC3143DFA93F443" box="[1367,1386,1305,1328]" italics="true" pageId="35" pageNumber="36">N</emphasis>
|
||||
= 10]: FBL = 3.13–3.68 (3.42); TL = 5.87–6.68 (6.29); HW = 0.78– 0.89 (0.85); HL = 0.8–0.9 (0.85); HW/HL = 0.98–1.02 (1); PW = 0.95–1.24 (1.14); PL = 0.89–1.16 (1.06); PW/ PL = 1.04–1.11 (1.08); EW = 1.54–1.82 (1.64); EL = 1.33– 1.63 (1.5); PW/HW = 1.1–1.41 (1.35).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FCD114F1FB3CF719" blockId="35.[810,1461,1117,1987]" pageId="35" pageNumber="36">Most of antennae, mouthparts, pronotum, elytra, legs yellow or yellowish-brown; last three or five antennomeres and pronotum sometimes darker; head, mandibles, pro-, meso-, and metasternum, metacoxal, and abdomen reddish-brown; tergites and sternites with pale posterior margin.</paragraph>
|
||||
<paragraph id="8BC736E5FF822E00FCD11755FE8FF34C" blockId="35.[810,1461,1117,1987]" lastBlockId="36.[127,779,144,1421]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
|
||||
Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with several setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.47), from 1.07 to 1.44 times as long as temples (in lateral view) in males and
|
||||
<quantity id="4C809B00FF822E07FBA91609FB2CF636" box="[1085,1237,1837,1861]" metricMagnitude="-2" metricUnit="m" metricValue="2.9337" metricValueMax="3.175" metricValueMin="2.6924" pageId="35" pageNumber="36" unit="in" value="1.155" valueMax="1.25" valueMin="1.06">1.06 to 1.25 in</quantity>
|
||||
females; distance between eyes about 1.16 times as long as length of eye in males and 1.35 times in females. Antennomeres 2 and 3 subequal in length; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a11 from 1.88 to 2.2 times long as a
|
||||
<quantity id="4C809B00FF822E00FA0D168FFF6CF1DB" lastPageId="36" lastPageNumber="37" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="10.0">10 in</quantity>
|
||||
males and from
|
||||
<quantity id="4C809B00FF852E00FED111B4FE24F1DB" box="[325,477,144,168]" metricMagnitude="-2" metricUnit="m" metricValue="4.6482" metricValueMax="4.9276" metricValueMin="4.3688" pageId="36" pageNumber="37" unit="in" value="1.83" valueMax="1.94" valueMin="1.72">1.72 to 1.94 in</quantity>
|
||||
females. Basal and parocular punctures usually single; posterior frontal puncture located posterior to temporal puncture; without small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight, almost united with gular sutures; postgenal ridge present; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than previous and more or less subconical.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF852E00FF081363FD5DF388" blockId="36.[127,779,144,1421]" pageId="36" pageNumber="37">Pronotum slightly transverse, convex, evenly curved, with two PPDS, usually without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to one or three times as long as, diameter of punctures. Mesosternum with seven macrosetae arranged in two rows medially.</paragraph>
|
||||
<paragraph id="8BC736E5FF852E00FF081227FD45F285" blockId="36.[127,779,144,1421]" pageId="36" pageNumber="37">Protibiae usually sexually dimorphic, with laterodorsal row of thick spines only in females; rarely in males too. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 moderately longer than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate.</paragraph>
|
||||
<paragraph id="8BC736E5FF852E00FF0812D9FCFFF41D" blockId="36.[127,779,144,1421]" pageId="36" pageNumber="37">
|
||||
Abdominal tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with deep sharp V-shaped emargination medially (
|
||||
<figureCitation id="13432A60FF852E00FF1F159DFF20F5A2" box="[139,217,1209,1233]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="36" pageNumber="37">Fig. 14I</figureCitation>
|
||||
); female sternite VIII with arcuate emargination medially; male sternite IX with V-shaped emargination medially, its basal portion moderately longer than distal portion; male tergite X with deep U-shaped emargination medially (
|
||||
<figureCitation id="13432A60FF852E00FD1E1433FD2EF45C" box="[650,727,1303,1327]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="36" pageNumber="37">Fig. 12I</figureCitation>
|
||||
); female tergite X with V-shaped emargination medially (
|
||||
<figureCitation id="13432A60FF852E00FD3C1413FD0FF43C" box="[680,758,1335,1359]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="36" pageNumber="37">Fig. 13I</figureCitation>
|
||||
); ovipositor, its second gonocoxite with one macroseta medially.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF852E00FF081451FD22F4FE" blockId="36.[127,779,144,1421]" box="[156,731,1397,1421]" pageId="36" pageNumber="37">
|
||||
Aedeagus as in
|
||||
<figureCitation id="13432A60FF852E00FEA31451FE30F4FE" box="[311,457,1397,1421]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="36" pageNumber="37">Figure 11G, H</figureCitation>
|
||||
; its total length 1.05–1.15.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF852E00FF151483FE2FF664" pageId="36" pageNumber="37" type="distribution">
|
||||
<paragraph id="8BC736E5FF852E00FF151483FE2FF664" blockId="36.[128,778,1447,1816]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B90CEAF7FF852E00FF151483FD91F4CC" box="[129,616,1447,1471]" italics="true" pageId="36" pageNumber="37">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FF852E00FE171483FD91F4CC" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[387,616,1447,1471]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lividipennis">Loncovilius lividipennis</taxonomicName>
|
||||
</emphasis>
|
||||
is known from
|
||||
<collectingCountry id="F36F7675FF852E00FF1514E3FF40F4AC" box="[129,185,1479,1503]" name="Chile" pageId="36" pageNumber="37">Chile</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="F36F7675FF852E00FF7C14E3FEA9F4AC" box="[232,336,1479,1503]" name="Argentina" pageId="36" pageNumber="37">Argentina</collectingCountry>
|
||||
. In
|
||||
<collectingCountry id="F36F7675FF852E00FEE714E3FE54F4AC" box="[371,429,1479,1503]" name="Chile" pageId="36" pageNumber="37">Chile</collectingCountry>
|
||||
, it occurs in the
|
||||
<collectingRegion id="49BCF807FF852E00FDDA14E3FD4EF4AC" box="[590,695,1479,1503]" country="Chile" name="Araucania" pageId="36" pageNumber="37">Araucanía</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FF852E00FD5414E3FCFFF4AC" box="[704,774,1479,1503]" country="Chile" name="Biobio" pageId="36" pageNumber="37">Biobío</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FF852E00FF1414C2FF13F48D" box="[128,234,1510,1534]" country="Chile" name="Los Lagos" pageId="36" pageNumber="37">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FF852E00FF6014C2FEA9F48D" box="[244,336,1510,1534]" country="Chile" name="Los Rios" pageId="36" pageNumber="37">Los Ríos</collectingRegion>
|
||||
, and Ñuble regions. In
|
||||
<collectingCountry id="F36F7675FF852E00FDAB14C2FD5EF48D" box="[575,679,1510,1534]" name="Argentina" pageId="36" pageNumber="37">Argentina</collectingCountry>
|
||||
, it is confined to the westernmost portions of the
|
||||
<collectingRegion id="49BCF807FF852E00FD8A1721FD89F76E" box="[542,624,1541,1565]" country="Argentina" name="Chubut" pageId="36" pageNumber="37">Chubut</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FF852E00FDED1721FD23F76E" box="[633,730,1541,1565]" country="Argentina" name="Neuquen" pageId="36" pageNumber="37">Neuquén</collectingRegion>
|
||||
, and
|
||||
<collectingRegion id="49BCF807FF852E00FF141701FF16F74E" box="[128,239,1573,1597]" country="Argentina" name="Rio Negro" pageId="36" pageNumber="37">Río Negro</collectingRegion>
|
||||
provinces. The species occurs from
|
||||
<quantity id="4C809B00FF852E00FDF81700FCF3F74F" box="[620,778,1572,1597]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.6" metricValueMin="0.55" pageId="36" pageNumber="37" unit="m" value="1075.0" valueMax="1600.0" valueMin="550.0">550 to 1600 m</quantity>
|
||||
elevation; it is reported in the
|
||||
<emphasis id="B90CEAF7FF852E00FE241760FD73F72F" box="[432,650,1604,1628]" italics="true" pageId="36" pageNumber="37">
|
||||
<taxonomicName id="4C784D66FF852E00FE241760FDEBF728" box="[432,530,1604,1627]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="genus">Araucaria</taxonomicName>
|
||||
-
|
||||
<taxonomicName id="4C784D66FF852E00FD8E1760FD73F72F" box="[538,650,1604,1628]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
|
||||
</emphasis>
|
||||
forests from central and north-central portions of the Valdivian temperate forest ecoregion. It has been collected using Malaise and window traps, beating the forest vegetation, attracted to ultraviolet lights, sifted from moss and leaf litter, and directly from male cones of
|
||||
<taxonomicName id="4C784D66FF852E00FF1617C5FEB0F78B" box="[130,329,1761,1784]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="araucana">
|
||||
<emphasis id="B90CEAF7FF852E00FF1617C5FEB0F78B" box="[130,329,1761,1784]" italics="true" pageId="36" pageNumber="37">Araucaria araucana</emphasis>
|
||||
</taxonomicName>
|
||||
and flowers, e.g. from
|
||||
<taxonomicName id="4C784D66FF852E00FDD217C5FCF0F78A" box="[582,777,1761,1785]" class="Magnoliopsida" family="Loranthaceae" genus="Desmaria" kingdom="Plantae" order="Santalales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="mutabilis">
|
||||
<emphasis id="B90CEAF7FF852E00FDD217C5FCF0F78A" box="[582,777,1761,1785]" italics="true" pageId="36" pageNumber="37">Desmaria mutabilis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C784D66FF852E00FF1C1624FEE0F66B" box="[136,281,1792,1816]" class="Magnoliopsida" family="Loranthaceae" kingdom="Plantae" order="Santalales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="family">Loranthaceae</taxonomicName>
|
||||
) on
|
||||
<taxonomicName id="4C784D66FF852E00FED11624FE2FF664" box="[325,470,1792,1816]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="undetermined">
|
||||
<emphasis id="B90CEAF7FF852E00FED11624FE4CF66B" box="[325,437,1792,1816]" italics="true" pageId="36" pageNumber="37">Nothofagus</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF852E00FF15161DFA3AF32C" pageId="36" pageNumber="37" type="discussion">
|
||||
<paragraph id="8BC736E5FF852E00FF15161DFA3AF32C" blockId="36.[129,778,1849,1967]" lastBlockId="36.[825,1476,144,607]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B90CEAF7FF852E00FF15161DFF22F622" box="[129,219,1849,1873]" italics="true" pageId="36" pageNumber="37">Remarks:</emphasis>
|
||||
This species was described in the genus
|
||||
<taxonomicName id="4C784D66FF852E00FD31161DFE06F602" authority="(Fairmaire and Germain 1862: 430)" baseAuthorityName="Fairmaire and Germain" baseAuthorityPageNumber="430" baseAuthorityYear="1862" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FD31161DFCF3F622" box="[677,778,1849,1873]" italics="true" pageId="36" pageNumber="37">Philonthus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF852E00FF1F167DFE01F602" author="Fairmaire L & Germain P" box="[139,504,1881,1905]" pageId="36" pageNumber="37" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain 1862: 430</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
based on an unspecified number of
|
||||
<typeStatus id="54C38847FF852E00FF6F165CFEAAF6E3" box="[251,339,1912,1936]" pageId="36" pageNumber="37" type="syntype">syntypes</typeStatus>
|
||||
from the Andean forests near Chillan, a city in central
|
||||
<collectingCountry id="F36F7675FF852E00FE8816B3FEADF6DC" box="[284,340,1943,1967]" name="Chile" pageId="36" pageNumber="37">Chile</collectingCountry>
|
||||
. Fauvel (1866) moved the species to the genus
|
||||
<taxonomicName id="4C784D66FF852E00FCE811B4FC33F1DB" box="[892,970,144,168]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FCE811B4FC33F1DB" box="[892,970,144,168]" italics="true" pageId="36" pageNumber="37">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
, while
|
||||
<bibRefCitation id="EFE94B14FF852E00FB8C11B4FB3AF1DB" author="Germain P" box="[1048,1219,144,168]" pageId="36" pageNumber="37" pagination="391 - 445" refId="ref36747" refString="Germain P. Apuntes Entomolojicos. Rectificaciones, anotaciones complementarias, i descripciones de siete especies nuevas de coleopteros. Anales de la Universidad de Chile 1903; 113: 391 - 445." type="journal article" year="1903">Germain (1903)</bibRefCitation>
|
||||
to his genus
|
||||
<taxonomicName id="4C784D66FF852E00FAC711B4FA45F1DB" box="[1363,1468,144,168]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FAC711B4FA45F1DB" box="[1363,1468,144,168]" italics="true" pageId="36" pageNumber="37">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
. Scheerpeltz (1972), who treated
|
||||
<taxonomicName id="4C784D66FF852E00FB1C118BFB0BF1B4" box="[1160,1266,175,199]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FB1C118BFB0BF1B4" box="[1160,1266,175,199]" italics="true" pageId="36" pageNumber="37">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
at the rank of a subgenus of
|
||||
<taxonomicName id="4C784D66FF852E00FC0311EBFC1CF194" box="[919,997,207,231]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FC0311EBFC1CF194" box="[919,997,207,231]" italics="true" pageId="36" pageNumber="37">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
, listed significant new material for this species.
|
||||
<bibRefCitation id="EFE94B14FF852E00FCAD11CAFBCBF075" author="Coiffait H & Saiz F" box="[825,1074,238,262]" pageId="36" pageNumber="37" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz (1966)</bibRefCitation>
|
||||
provided the species redescription and first illustration of the aedeagus based on the non-type material that comes from localities other than ‘Chillan’.
|
||||
<bibRefCitation id="EFE94B14FF852E00FA9B1009FA7FF036" author="Saiz F" box="[1295,1414,301,325]" pageId="36" pageNumber="37" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz (1971)</bibRefCitation>
|
||||
listed a few more specimens. Camousseight (1980) lists a ‘
|
||||
<typeStatus id="54C38847FF852E00FADB1068FA5EF017" box="[1359,1447,332,356]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
|
||||
N 2267–2272’ of
|
||||
<taxonomicName id="4C784D66FF852E00FC45104FFB5CF0F0" authorityName="Fairmaire & Germain" authorityYear="1862" box="[977,1189,363,387]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF852E00FC45104FFB5CF0F0" box="[977,1189,363,387]" italics="true" pageId="36" pageNumber="37">Philonthus lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
from Germain’s collection at the National Museum of
|
||||
<collectingCountry id="F36F7675FF852E00FBA310AFFB97F0D0" box="[1079,1134,395,419]" name="Chile" pageId="36" pageNumber="37">Chile</collectingCountry>
|
||||
in
|
||||
<collectingRegion id="49BCF807FF852E00FB1B10AFFB1CF0D0" box="[1167,1253,395,419]" country="Chile" name="Region Metropolitana" pageId="36" pageNumber="37">Santiago</collectingRegion>
|
||||
, which is apparently a
|
||||
<typeStatus id="54C38847FF852E00FCAF108EFC72F0B1" box="[827,907,426,450]" pageId="36" pageNumber="37" type="syntype">syntype</typeStatus>
|
||||
.
|
||||
<bibRefCitation id="EFE94B14FF852E00FC00108EFB73F0B1" author="Coiffait H & Saiz F" box="[916,1162,426,450]" pageId="36" pageNumber="37" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz (1966)</bibRefCitation>
|
||||
denoted one of the specimens they examined as a
|
||||
<typeStatus id="54C38847FF852E00FC6010EEFBBFF091" box="[1012,1094,458,482]" pageId="36" pageNumber="37" type="neotype">neotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C784D66FF852E00FBF410EEFB18F092" authorityName="Fairmaire & Germain" authorityYear="1862" box="[1120,1249,457,481]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF852E00FBF410EEFB18F092" box="[1120,1249,457,481]" italics="true" pageId="36" pageNumber="37">L.lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
. Although we were unable to examine any
|
||||
<typeStatus id="54C38847FF852E00FB9610CDFBA3F372" box="[1026,1114,489,513]" pageId="36" pageNumber="37" type="syntype">syntypes</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C784D66FF852E00FBED10CDFB02F372" authorityName="Fairmaire & Germain" authorityYear="1862" box="[1145,1275,489,513]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF852E00FBED10CDFB02F372" box="[1145,1275,489,513]" italics="true" pageId="36" pageNumber="37">L. lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
, we assume that the currently accepted concept of this species is correct, because the only other externally similar species,
|
||||
<taxonomicName id="4C784D66FF852E00FB35130CFAEBF34C" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[1185,1298,551,575]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="carlsbergi" status="sp. nov.">
|
||||
<emphasis id="B90CEAF7FF852E00FB35130CFAEBF34C" box="[1185,1298,551,575]" italics="true" pageId="36" pageNumber="37">L. carlsbergi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF852E00FA82130CFA98F34C" box="[1302,1377,552,575]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, is known only further south, from the Magellanic subpolar forests ecoregion.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,110 +0,0 @@
|
|||
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<emphasis id="B90CEAF7FF832E06FEFD17FDFE25F787" bold="true" box="[361,476,1753,1780]" italics="true" pageId="34" pageNumber="35">lividipennis</emphasis>
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|
||||
group
|
||||
</emphasis>
|
||||
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|
||||
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|
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|
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Included species:
|
||||
<taxonomicName id="4C784D66FF832E06FEBE162FFE02F650" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[298,507,1803,1827]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="carlsbergi" status="sp. nov.">Loncovilius carlsbergi</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A23F578CFF832E06FD94162FFDB4F651" box="[512,589,1803,1826]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C784D66FF832E06FD16162FFE3DF631" authority="(Fairmaire and Germain, 1862)" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF832E06FD16162FFD76F651" box="[642,655,1803,1826]" italics="true" pageId="34" pageNumber="35">L</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF832E06FD0F162FFCF0F650" box="[667,777,1803,1827]" italics="true" pageId="34" pageNumber="35">lividipennis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF832E06FF1F160EFE41F631" author="Fairmaire L & Germain P" box="[139,440,1834,1858]" pageId="34" pageNumber="35" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
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)
|
||||
</taxonomicName>
|
||||
.
|
||||
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|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF832E06FF15167EFABFF0F0" pageId="34" pageNumber="35" type="diagnosis">
|
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|
||||
<emphasis id="B90CEAF7FF832E06FF15167EFF1EF602" box="[129,231,1882,1905]" italics="true" pageId="34" pageNumber="35">Diagnosis:</emphasis>
|
||||
Head about as wide as long, without coarse non-setiferous punctures; nuchal ridge complete dorsally, connected with infraorbital ridge; gula without isodiametric microsculpture in the middle; posterior frontal puncture located posterior to temporal puncture; postmandibular ridge located close to eye margin; PMP (usually reduced); females with antennomere 11 distinctly longer than antennomere 10 (a11:a10 ratio>1.5). Pronotum outline evenly curved. Tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area near to apical margin (
|
||||
<figureCitation id="13432A60FF832E06FAB51009FA91F036" box="[1313,1384,301,325]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="34" pageNumber="35">Fig. 9E</figureCitation>
|
||||
); tergite VIII with medial apical emargination in both sexes; ovipositor, its second gonocoxite with one macroseta medially.
|
||||
</paragraph>
|
||||
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|
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|
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|
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|
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@ -1,406 +0,0 @@
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<mods:roleTerm id="8FFAE0F042052FDE4D51E5E73A2D1C75">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="54B7A294CD44F15A4EC35915CC8DAC4F">Solodovnikov, Alexey</mods:namePart>
|
||||
<mods:affiliation id="8096D8938E122C69E9107798DD0A08B4">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="9E641086B5CA4BDFDC8D40A3625CA38F">text</mods:typeOfResource>
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||||
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||||
<mods:titleInfo id="2ECE20BE3AD0B995643348763DC95A34">
|
||||
<mods:title id="3AB9562788344B26A1BE0024926AAAC0">Zoological Journal of the Linnean Society</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="4D4257D8B7061C25B66354FFE3F8944C">
|
||||
<mods:date id="59B6294B6E515C391186B06BC4E9B1F3">2024</mods:date>
|
||||
<mods:detail id="C07090571D9C56BF87939E0726F41B54" type="series">
|
||||
<mods:title id="7F132A2D823E1B8F6071FC1F3310AFE6">Zool. J. Linn. Soc.</mods:title>
|
||||
</mods:detail>
|
||||
<mods:detail id="13DEED170F180C786D3E77B975FB51AE" type="pubDate">
|
||||
<mods:number id="2E9CC57A837F4A1B34F9215E7826D98D">2023-10-28</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="65DD83E91455F3FAE7E7A3171229DCB4" type="volume">
|
||||
<mods:number id="8B68814C288B10CF31BB8335111E0FDE">202</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="E711356FD0A3BD9032E2EAE5088B325C" type="issue">
|
||||
<mods:number id="937D7B79CED8F2B0E17D9AA481B6F026">1</mods:number>
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<mods:extent id="746867213C0AC87DD2DB12091290176E" unit="page">
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<mods:start id="C2735C9A9395A4B7F68173964884D36A">1</mods:start>
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<mods:end id="F4A6949336B024AD1C25585FBDE85B53">42</mods:end>
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||||
</mods:extent>
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||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="E41754C34601E88E360F9AD322DBA9D7">
|
||||
<mods:url id="626CECDE066B9DB629D01AFC7CBC5E6F">https://doi.org/10.1093/zoolinnean/zlad143</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="201DF6EAE46CC71BA94DA1DE619EDF55">journal article</mods:classification>
|
||||
<mods:identifier id="ABC5714D8F6BCC30FE7B54F07F52301C" type="DOI">10.1093/zoolinnean/zlad143</mods:identifier>
|
||||
<mods:identifier id="87CB081194D9C7DDEC589DC94FF3B61B" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="D03B4A3759106A5EC7F305717811512B" type="Zenodo-Dep">14541979</mods:identifier>
|
||||
<mods:identifier id="AEAD97124A410229B49E4602ECD9C46A" type="ZooBank">CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03D187F3FF832E07FC7D1086FB1DF0EE" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE" httpUri="http://treatment.plazi.org/id/03D187F3FF832E07FC7D1086FB1DF0EE" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
|
||||
<subSubSection id="C362656EFF832E06FC7D1086FAEBF0CE" box="[1001,1298,418,445]" pageId="34" pageNumber="35" type="nomenclature">
|
||||
<paragraph id="8BC736E5FF832E06FC7D1086FAEBF0CE" blockId="34.[1001,1298,418,445]" box="[1001,1298,418,445]" pageId="34" pageNumber="35">
|
||||
<heading id="D08F8189FF832E06FC7D1086FAEBF0CE" bold="true" box="[1001,1298,418,445]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="35" reason="2">
|
||||
<emphasis id="B90CEAF7FF832E06FC7D1086FAEBF0CE" bold="true" box="[1001,1298,418,445]" pageId="34" pageNumber="35">
|
||||
<taxonomicName id="4C784D66FF832E06FC7D1086FB38F0CE" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[1001,1217,418,445]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="carlsbergi" status="sp. nov.">
|
||||
<emphasis id="B90CEAF7FF832E06FC7D1086FB38F0CE" bold="true" box="[1001,1217,418,445]" italics="true" pageId="34" pageNumber="35">Loncovilius carlsbergi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF832E06FB521087FAEBF0CE" box="[1222,1298,419,445]" pageId="34" pageNumber="35" rank="species">sp.nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF832E07FC5210F6FB1DF0EE" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
|
||||
<paragraph id="8BC736E5FF832E06FC5210F6FACFF09F" blockId="34.[966,1334,466,492]" box="[966,1334,466,492]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B90CEAF7FF832E06FC5210F6FACFF09F" bold="true" box="[966,1334,466,492]" pageId="34" pageNumber="35">
|
||||
(
|
||||
<figureCitation id="13432A60FF832E06FC4410F6FBC1F09F" box="[976,1080,466,492]" captionStart="Figure 7" captionStartId="15.[113,178,1868,1892]" captionTargetBox="[153,1419,163,1840]" captionTargetId="figure-6@15.[153,1419,144,1840]" captionTargetPageId="15" captionText="Figure 7. Female habitus of Loncovilius species.A, L. lividipennis. B, L. carlsbergi. Head. C, L. hammondi. D, L. semiflavus. E, L. carlsbergi. F, Male mesotarsi of L. cantharoides." pageId="34" pageNumber="35">Figs 7B, E</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FF832E06FBD510F6FB98F09F" box="[1089,1121,466,492]" captionStart="Figure 8" captionStartId="16.[129,194,1829,1853]" captionTargetBox="[183,1419,150,1799]" captionTargetId="figure-8@16.[183,1419,144,1799]" captionTargetPageId="16" captionText="Figure 8. A, pronotum of L. carlsbergi. AMRSP = anterior marginal row of setiferous punctures, APP = additional paired punctures adjacent to AMRSP, LLSP = large lateral setiferous puncture, LPSP = large posterior setiferous puncture, PPDS = paired punctures on dorsal serial, SLSP = sublateral setiferous puncture.B, L. cantharoides ventral view of mesotarsomere 3 with pale adhesive setae with terminal plate.C, L. cantharoides ventral view of metatarsomere 4, apical margin emarginated (bilobed), with pale adhesive setae without terminal plate. D, L. cantharoides tergites VI and VII.Black arrow up = anterior transverse basal carina (ATBC) laterally straight continuing to paratergites, white arrow down = ATBC descending laterally without reaching tergite margin.Head in ventral view of Loncovilius species. E, L. hammondi. F, L. lividipennis. PMP = postmandibular puncture, PMR = postmandibular ridge, EM = eye margin." pageId="34" pageNumber="35">8A</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FF832E06FBFF10F6FB5AF09F" box="[1131,1187,466,492]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="34" pageNumber="35">11I, J</figureCitation>
|
||||
, 12J, 13J, 14J)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF832E06FCAD1327FB48F348" blockId="34.[825,1474,515,571]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B90CEAF7FF832E06FCAD1327FBE7F368" box="[825,1054,515,539]" italics="true" pageId="34" pageNumber="35">Zoobank registration:</emphasis>
|
||||
<uri id="FFE93AE7FF832E06FBA31327FB54F348" pageId="34" pageNumber="35">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFDE0C30FF832E06FAA61327FB54F348" pageId="34" pageNumber="35">C2A7BAAE-3D40-4ED6-9C8D-09602F86B9B3</uuid>
|
||||
</uri>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF832E06FCAD1379FBFEF772" blockId="34.[824,1475,604,1537]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B90CEAF7FF832E06FCAD1379FC32F307" box="[825,971,604,628]" italics="true" pageId="34" pageNumber="35">Type material.</emphasis>
|
||||
<materialsCitation id="3B103CB8FF832E06FC401378FAD2F3A1" collectingDate="1958-12-04" collectionCode="NHM, NHMD" collectorName="Fr Kuschel" country="Chile" location="Chilean Antarctica, Puerto Eden" pageId="34" pageNumber="35" specimenCode="916967" specimenCount="1" stateProvince="Magallanes" typeStatus="holotype">
|
||||
<emphasis id="B90CEAF7FF832E06FC401378FBCDF307" box="[980,1076,604,628]" italics="true" pageId="34" pageNumber="35">
|
||||
<typeStatus id="54C38847FF832E06FC401378FBD6F307" box="[980,1071,604,628]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
male, pinned, with labels as follows: [
|
||||
<collectingCountry id="F36F7675FF832E06FCD4135FFC6FF3E0" box="[832,918,635,659]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FF832E06FC31135FFBE1F3E0" box="[933,1048,635,659]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="34" pageNumber="35">Magallanes</collectingRegion>
|
||||
and
|
||||
<location id="8EA7603EFF832E06FBC3135FFA44F3E0" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FBC3135FFA44F3E0" box="[1111,1469,635,659]" country="Chile" name="Chilean Antarctica, Puerto Eden" pageId="34" pageNumber="35" stateProvince="Magallanes">
|
||||
Chilean
|
||||
<taxonomicName id="4C784D66FF832E06FB21135FFA44F3E0" authority=", Puerto Eden" authorityName="Puerto Eden" box="[1205,1469,635,659]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="antarctica">Antarctica, Puerto Edén</taxonomicName>
|
||||
</location>
|
||||
]
|
||||
<location id="8EA7603EFF832E06FCAD13BFFBB9F3C0" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FCAD13BFFBB9F3C0" box="[825,1088,667,691]" country="Chile" name="Wellington Is. Pto. Eden" pageId="34" pageNumber="35" stateProvince="Magallanes">Wellington Is. Pto. Eden</location>
|
||||
[-49.11, -74.40]
|
||||
<date id="FFC61025FF832E06FB6A13BEFA91F3C1" box="[1278,1384,666,690]" pageId="34" pageNumber="35" value="1958-12-04">
|
||||
<collectingDate id="EF82E9CDFF832E06FB6A13BEFA91F3C1" box="[1278,1384,666,690]" pageId="34" pageNumber="35" value="1958-12-04">4.XII1958</collectingDate>
|
||||
</date>
|
||||
/CHILE
|
||||
<collectorName id="268D5333FF832E06FCAD139EFC56F3A1" box="[825,943,698,722]" pageId="34" pageNumber="35">Fr Kuschel</collectorName>
|
||||
/
|
||||
<collectionCode id="ED69AE20FF832E06FC23139EFC0FF3A1" box="[951,1014,698,722]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
[blue label]/
|
||||
<collectionCode id="ED69AE20FF832E06FBEB139EFB2CF3A1" box="[1151,1237,698,722]" collectionName="NHMD" pageId="34" pageNumber="35">NHMD</collectionCode>
|
||||
<specimenCode id="DBDE9E9EFF832E06FB48139EFAD2F3A1" box="[1244,1323,698,722]" pageId="34" pageNumber="35">916967</specimenCode>
|
||||
</materialsCitation>
|
||||
/
|
||||
<typeStatus id="54C38847FF832E06FAA7139EFA3BF3A1" box="[1331,1474,698,722]" pageId="34" pageNumber="35" type="holotype">HOLOTYPE</typeStatus>
|
||||
<taxonomicName id="4C784D66FF832E06FCAF13FDFBE8F382" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[827,1041,729,753]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="carlsbergi">
|
||||
<emphasis id="B90CEAF7FF832E06FCAF13FDFBE8F382" box="[827,1041,729,753]" italics="true" pageId="34" pageNumber="35">Loncovilius carlsbergi</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at NHM.
|
||||
<materialsCitation id="3B103CB8FF832E06FACF13DDFB57F23C" collectingDate="1958-12-04" collectingDateMax="1958-12-27" collectingDateMin="1958-11-20" collectorName="Wellington Is. Pto. Eden & I. Puerto Eden & Pto. Eden Wellington Is. & Pena. Munoz, HM & Fr. Kuschel & P. Voillians" country="Chile" determinerName="A. Newton" elevation="12" location="Chilean Antarctica, Puerto Eden" municipality="Fr Kuschel" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="18" specimenCount-male="21" stateProvince="Magallanes" typeStatus="paratype">
|
||||
<emphasis id="B90CEAF7FF832E06FACF13DDFA3BF263" box="[1371,1474,761,784]" italics="true" pageId="34" pageNumber="35">
|
||||
<typeStatus id="54C38847FF832E06FACF13DDFA44F263" box="[1371,1469,761,784]" pageId="34" pageNumber="35" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
[
|
||||
<collectingCountry id="F36F7675FF832E06FCD4123CFC6FF243" box="[832,918,792,816]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FF832E06FC31123CFBE1F243" box="[933,1048,792,816]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="34" pageNumber="35">Magallanes</collectingRegion>
|
||||
and
|
||||
<location id="8EA7603EFF832E06FBC3123CFA44F243" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FBC3123CFA44F243" box="[1111,1469,792,816]" country="Chile" municipality="Fr Kuschel" name="Chilean Antarctica, Puerto Eden" pageId="34" pageNumber="35" stateProvince="Magallanes">
|
||||
Chilean
|
||||
<taxonomicName id="4C784D66FF832E06FB21123CFA44F243" authority=", Puerto Eden" authorityName="Puerto Eden" box="[1205,1469,792,816]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="antarctica">Antarctica, Puerto Edén</taxonomicName>
|
||||
</location>
|
||||
] Wellington Is. Pto. Eden
|
||||
<date id="FFC61025FF832E06FBA31213FB5FF23C" box="[1079,1190,823,847]" pageId="34" pageNumber="35" value="1958-12-04">
|
||||
<collectingDate id="EF82E9CDFF832E06FBA31213FB5FF23C" box="[1079,1190,823,847]" pageId="34" pageNumber="35" value="1958-12-04">4.XII.1958</collectingDate>
|
||||
</date>
|
||||
/
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF832E06FB3A1213FA90F2DE" collectingDate="1958-11-20" collectingDateMax="1958-12-27" collectingDateMin="1958-11-20" collectionCode="NHM, NHMD, CZUG, FMNH" collectorName="Wellington Is. Pto. Eden & I. Puerto Eden & Pto. Eden Wellington Is. & Pena. Munoz, HM & Fr. Kuschel & P. Voillians" country="Chile" determinerName="A. Newton" elevation="12" location="CHILE" municipality="Fr Kuschel" pageId="34" pageNumber="35" specimenCount="18" specimenCount-female="10" specimenCount-male="8" stateProvince="Magallanes y de la Antartica Chilena" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF832E06FB3A1213FAF8F23C" box="[1198,1281,823,847]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
|
||||
Fr Kuschel/
|
||||
<collectionCode id="ED69AE20FF832E06FA171213FA3BF23C" box="[1411,1474,823,847]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
[blue label] [
|
||||
<specimenCount id="9D7EFD6CFF832E06FC2F1273FC1DF21D" box="[955,996,855,879]" count="2" pageId="34" pageNumber="35" type="male">2♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFF832E06FC7C1273FBF6F21D" box="[1000,1039,855,879]" count="6" pageId="34" pageNumber="35" type="female">6♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FB801273FBAEF21D" box="[1044,1111,855,878]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFF832E06FBCB1273FB7FF21D" box="[1119,1158,855,879]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FB1E1273FB24F21D" box="[1162,1245,855,878]" collectionName="NHMD" pageId="34" pageNumber="35">NHMD</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FF832E06FA961273FABCF21D" box="[1282,1349,855,878]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFF832E06FADA1273FA8CF21D" box="[1358,1397,855,879]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FAED1273FA3BF21C" box="[1401,1474,855,879]" collectionName="Mexico, Jalisco, Guadalajara, Universidad de Guadalajara, Centro de Estudios en Zoologia, Entomologia" pageId="34" pageNumber="35">CZUG</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FF832E06FCCE1252FC64F2FE" box="[858,925,886,909]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFF832E06FC321252FC36F2FE" box="[934,975,886,910]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FC401252FBDBF2FE" box="[980,1058,886,909]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="34" pageNumber="35">FMNH</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FF832E06FBDD1252FB69F2FE" box="[1097,1168,886,909]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
]; [same data as above except:]
|
||||
<date id="FFC61025FF832E06FCAD12B1FC54F2DE" box="[825,941,917,941]" pageId="34" pageNumber="35" value="1958-11-20">
|
||||
<collectingDate id="EF82E9CDFF832E06FCAD12B1FC54F2DE" box="[825,941,917,941]" pageId="34" pageNumber="35" value="1958-11-20">20.XI.1958</collectingDate>
|
||||
</date>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFF832E06FC2F12B1FC1CF2DF" box="[955,997,917,941]" count="5" pageId="34" pageNumber="35" type="male">5♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFF832E06FC7F12B1FBEBF2DF" box="[1003,1042,917,941]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FB8D12B2FBA5F2DE" box="[1049,1116,918,941]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFF832E06FBF212B1FB74F2DF" box="[1126,1165,917,941]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FB0012B2FB1BF2DE" box="[1172,1250,918,941]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="34" pageNumber="35">FMNH</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FF832E06FA9F12B2FAABF2DE" box="[1291,1362,918,941]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF832E06FAFD12B1FC77F559" collectingDate="1958-11-29" collectingDateMax="1958-12-27" collectingDateMin="1958-11-20" collectionCode="NHM" collectorName="Wellington Is. Pto. Eden & I. Puerto Eden & Pto. Eden Wellington Is. & Pena. Munoz, HM & Fr. Kuschel & P. Voillians" country="Chile" determinerName="A. Newton" elevation="12" location="Nothofagus antarctica" municipality="Fr Kuschel" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="1" specimenCount-male="21" stateProvince="Magallanes" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF832E06FAFD12B1FA46F2DE" box="[1385,1471,917,941]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FF832E06FCAE1291FC54F2BE" box="[826,941,949,973]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="34" pageNumber="35">Magallanes</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="49BCF807FF832E06FC7A1291FB43F2BE" box="[1006,1210,949,973]" pageId="34" pageNumber="35">Chilean Antarctica</collectingRegion>
|
||||
]
|
||||
<taxonomicName id="4C784D66FF832E06FB591291FA42F2BE" box="[1229,1467,949,973]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="antarctica">
|
||||
<location id="8EA7603EFF832E06FB591291FA42F2BE" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FB591291FA42F2BE" box="[1229,1467,949,973]" country="Chile" municipality="Fr Kuschel" name="Nothofagus antarctica" pageId="34" pageNumber="35" stateProvince="Magallanes">Nothofagus antarctica</location>
|
||||
</taxonomicName>
|
||||
/ CHILE HE
|
||||
<location id="8EA7603EFF832E06FC5412F0FBACF29F" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FC5412F0FBACF29F" box="[960,1109,980,1004]" country="Chile" municipality="Fr Kuschel" name="I. Puerto Eden" pageId="34" pageNumber="35" stateProvince="Magallanes">I.Puerto Eden</location>
|
||||
,
|
||||
<location id="8EA7603EFF832E06FBF012F0FAFAF29F" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FBF012F0FAFAF29F" box="[1124,1283,980,1004]" country="Chile" municipality="Fr Kuschel" name="Isla Wellington" pageId="34" pageNumber="35" stateProvince="Magallanes">Isla Wellington</location>
|
||||
[-49.126, -74.40] 490S
|
||||
<quantity id="4C809B00FF832E06FCE712D7FC51F578" box="[883,936,1011,1035]" metricMagnitude="1" metricUnit="m" metricValue="1.2192" pageId="34" pageNumber="35" unit="ft" value="40.0">
|
||||
<elevation id="0055D1D6FF832E06FCE712D7FC51F578" box="[883,936,1011,1035]" metricMagnitude="1" metricUnit="m" metricValue="1.2192" pageId="34" pageNumber="35" unit="ft" value="40.0">40 ft.</elevation>
|
||||
</quantity>
|
||||
<date id="FFC61025FF832E06FC3812D7FBE0F578" box="[940,1049,1011,1035]" pageId="34" pageNumber="35" value="1958-11-29">
|
||||
<collectingDate id="EF82E9CDFF832E06FC3812D7FBE0F578" box="[940,1049,1011,1035]" pageId="34" pageNumber="35" value="1958-11-29">29.xi.1958</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<collectionCode id="ED69AE20FF832E06FBB512D7FB9DF578" box="[1057,1124,1011,1035]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFF832E06FBE012D7FB62F579" box="[1140,1179,1011,1035]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FB3512D0FB11F578" box="[1185,1256,1012,1035]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
]; [same data as above except:]
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF832E06FC011537FB38F558" box="[917,1217,1043,1067]" collectingDate="1958-11-20" collectingDateMax="1958-12-27" collectingDateMin="1958-11-20" collectionCode="NHM" collectorName="Wellington Is. Pto. Eden & I. Puerto Eden & Pto. Eden Wellington Is. & Pena. Munoz, HM & Fr. Kuschel & P. Voillians" country="Chile" determinerName="A. Newton" elevation="12" location="CHILE" municipality="Fr Kuschel" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="1" specimenCount-male="21" stateProvince="Magallanes y de la Antartica Chilena" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF832E06FC011537FC1AF558" box="[917,995,1043,1067]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
|
||||
HE6. [
|
||||
<specimenCount id="9D7EFD6CFF832E06FBA71537FBA3F559" box="[1075,1114,1043,1067]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FBF61537FB50F559" box="[1122,1193,1043,1066]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF832E06FB551537FB96F5DB" collectingDate="2015-12-07" collectingDateMax="1958-12-27" collectingDateMin="1958-11-20" collectionCode="MCZ, F, G, FMNH" collectorName="Wellington Is. Pto. Eden & I. Puerto Eden & Pto. Eden Wellington Is. & Pena. Munoz, HM & Fr. Kuschel & P. Voillians" country="Chile" determinerName="A. Newton" elevation="12" location="Pto. Eden Wellington Is" municipality="Fr Kuschel" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="18" specimenCount-male="1" stateProvince="Magallanes and Chilean Antarctica" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF832E06FB551537FAEEF558" box="[1217,1303,1043,1067]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FF832E06FAB61537FC05F539" pageId="34" pageNumber="35">Magallanes and Chilean Antarctica</collectingRegion>
|
||||
] S. CHILE
|
||||
<location id="8EA7603EFF832E06FBF91516FAA5F53A" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FBF91516FAA5F53A" box="[1133,1372,1074,1098]" country="Chile" municipality="Fr Kuschel" name="Pto. Eden Wellington Is" pageId="34" pageNumber="35" stateProvince="Magallanes and Chilean Antarctica">Pto. Edén Wellington Is</location>
|
||||
. [-49.126, -74.40]
|
||||
<date id="FFC61025FF832E06FC181575FC10F51A" box="[908,1001,1105,1129]" pageId="34" pageNumber="35" value="2015-12-07">
|
||||
<collectingDate id="EF82E9CDFF832E06FC181575FC10F51A" box="[908,1001,1105,1129]" pageId="34" pageNumber="35" value="2015-12-07">Dec.7-15</collectingDate>
|
||||
</date>
|
||||
,’62 PJDarlinton/FIELD MUSEUM ex
|
||||
<collectionCode id="ED69AE20FF832E06FA131575FA38F51A" box="[1415,1473,1105,1129]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" pageId="34" pageNumber="35">MCZ</collectionCode>
|
||||
(retained duplic.)/
|
||||
<taxonomicName id="4C784D66FF832E06FB931555FB0EF5FA" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[1031,1271,1137,1161]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lividipennis">
|
||||
<emphasis id="B90CEAF7FF832E06FB931555FB0EF5FA" box="[1031,1271,1137,1161]" italics="true" pageId="34" pageNumber="35">Loncovilius lividipennis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<collectionCode id="ED69AE20FF832E06FA861555FADAF5FA" box="[1298,1315,1137,1161]" pageId="34" pageNumber="35">F</collectionCode>
|
||||
+
|
||||
<collectionCode id="ED69AE20FF832E06FAD41555FAAFF5FA" box="[1344,1366,1137,1161]" pageId="34" pageNumber="35">G</collectionCode>
|
||||
) det.
|
||||
<determinerName id="7E1BDA21FF832E06FA3F1555FC74F5D4" pageId="34" pageNumber="35">A. Newton</determinerName>
|
||||
1993 [
|
||||
<specimenCount id="9D7EFD6CFF832E06FC4215B4FC04F5D4" box="[982,1021,1168,1192]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FB9715B4FBA0F5D4" box="[1027,1113,1168,1191]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="34" pageNumber="35">FMNH</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF832E06FBFB15B4FA90F475" collectingDate="1958-12-27" collectingDateMax="1958-12-27" collectingDateMin="1958-11-20" collectionCode="NHM, MNNC" collectorName="Pena. Munoz" country="Chile" determinerName="A. Newton" elevation="9" location="Pena. Munoz" municipality="Nothofagus antarctica" pageId="34" pageNumber="35" specimenCount="8" specimenCount-female="3" specimenCount-male="4" stateProvince="Magallanes and Chilean Antarctica" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF832E06FBFB15B4FB3CF5DB" box="[1135,1221,1168,1192]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FF832E06FB5915B4FC51F5B4" pageId="34" pageNumber="35">Magallanes and Chilean Antarctica</collectingRegion>
|
||||
]
|
||||
<taxonomicName id="4C784D66FF832E06FC5E158BFB31F5B4" box="[970,1224,1199,1223]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="antarctica">
|
||||
<collectingMunicipality id="6BA3AC9FFF832E06FC5E158BFB31F5B4" box="[970,1224,1199,1223]" country="Chile" municipality="Fr Kuschel" name="Nothofagus antarctica" pageId="34" pageNumber="35" stateProvince="Magallanes and Chilean Antarctica">Nothofagus antarctica</collectingMunicipality>
|
||||
</taxonomicName>
|
||||
/CHILE
|
||||
<location id="8EA7603EFF832E06FAA81594FA47F5B4" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FAA81594FA47F5B4" box="[1340,1470,1200,1223]" country="Chile" municipality="Nothofagus antarctica" name="Pena. Munoz" pageId="34" pageNumber="35" stateProvince="Magallanes and Chilean Antarctica">Pena.Munoz</location>
|
||||
, HM3.
|
||||
<location id="8EA7603EFF832E06FCE715EBFC35F594" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FCE715EBFC35F594" box="[883,972,1231,1255]" country="Chile" municipality="Nothofagus antarctica" name="Gamero" pageId="34" pageNumber="35" stateProvince="Magallanes and Chilean Antarctica">Gamero</location>
|
||||
.
|
||||
<quantity id="4C809B00FF832E06FC4815EBFBB9F595" box="[988,1088,1231,1255]" metricMagnitude="0" metricUnit="m" metricValue="9.144" metricValueMax="12.192" metricValueMin="6.096" pageId="34" pageNumber="35" unit="ft" value="30.0" valueMax="40.0" valueMin="20.0">
|
||||
<elevation id="0055D1D6FF832E06FC4815EBFBB9F595" box="[988,1088,1231,1255]" metricMagnitude="0" metricUnit="m" metricValue="9.144" metricValueMax="12.192" metricValueMin="6.096" pageId="34" pageNumber="35" unit="ft" value="30.0" valueMax="40.0" valueMin="20.0">20–40 ft.</elevation>
|
||||
</quantity>
|
||||
[-52.33, -73.53,
|
||||
<quantity id="4C809B00FF832E06FB6A15EBFAD6F594" box="[1278,1327,1231,1255]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="34" pageNumber="35" unit="m" value="12.0">
|
||||
<elevation id="0055D1D6FF832E06FB6A15EBFAD6F594" box="[1278,1327,1231,1255]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="34" pageNumber="35" unit="m" value="12.0">12m</elevation>
|
||||
</quantity>
|
||||
]
|
||||
<date id="FFC61025FF832E06FAD715EAFA43F595" box="[1347,1466,1230,1254]" pageId="34" pageNumber="35" value="1958-12-27">
|
||||
<collectingDate id="EF82E9CDFF832E06FAD715EAFA43F595" box="[1347,1466,1230,1254]" pageId="34" pageNumber="35" value="1958-12-27">27.xii.1958</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<collectionCode id="ED69AE20FF832E06FCAD15CAFC80F476" box="[825,889,1262,1285]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFF832E06FC1C15CAFC48F476" box="[904,945,1262,1286]" count="3" pageId="34" pageNumber="35" type="male">3♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFF832E06FC2C15CAFC26F476" box="[952,991,1262,1286]" count="2" pageId="34" pageNumber="35" type="female">2♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FC7215CAFBD0F476" box="[998,1065,1262,1285]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFF832E06FBA015CAFBA2F476" box="[1076,1115,1262,1286]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFF832E06FBF515CAFB70F476" box="[1121,1161,1262,1286]" count="1" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FB1B15CAFB18F475" box="[1167,1249,1262,1286]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="34" pageNumber="35">MNNC</collectionCode>
|
||||
<specimenCount id="9D7EFD6CFF832E06FB7C15CAFAFCF476" box="[1256,1285,1262,1285]" pageId="34" pageNumber="35" type="generic">ex.</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FA9F15CAFAABF476" box="[1291,1362,1262,1285]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF832E06FAFD15CAFC4FF410" collectingDate="1958-12-27" collectingDateMax="1958-12-27" collectingDateMin="1958-11-20" collectionCode="NHM" collectorName="Fr. Kuschel" country="Chile" determinerName="A. Newton" elevation="12" location="Munoz Gamero" municipality="Fr Kuschel" pageId="34" pageNumber="35" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Magallanes and Chilean Antarctica" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FF832E06FAFD15CAFA46F475" box="[1385,1471,1262,1286]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FF832E06FCAF1429FB48F456" box="[827,1201,1293,1317]" pageId="34" pageNumber="35">Magallanes and Chilean Antarctica</collectingRegion>
|
||||
]
|
||||
<location id="8EA7603EFF832E06FB54142AFA91F456" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FB54142AFA91F456" box="[1216,1384,1293,1317]" country="Chile" municipality="Fr Kuschel" name="Munoz Gamero" pageId="34" pageNumber="35" stateProvince="Magallanes and Chilean Antarctica">Munoz Gamero</location>
|
||||
[-52.33, -73.53]
|
||||
<date id="FFC61025FF832E06FC061408FBF7F437" box="[914,1038,1324,1348]" pageId="34" pageNumber="35" value="1958-12-27">
|
||||
<collectingDate id="EF82E9CDFF832E06FC061408FBF7F437" box="[914,1038,1324,1348]" pageId="34" pageNumber="35" value="1958-12-27">27.XII.1958</collectingDate>
|
||||
</date>
|
||||
/CHILE
|
||||
<collectorName id="268D5333FF832E06FBE11409FB0CF436" box="[1141,1269,1325,1349]" pageId="34" pageNumber="35">Fr. Kuschel</collectorName>
|
||||
/Shore/
|
||||
<collectionCode id="ED69AE20FF832E06FAD01409FA7DF436" box="[1348,1412,1325,1349]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFF832E06FA0C1409FA3BF437" box="[1432,1474,1325,1349]" count="6" pageId="34" pageNumber="35" type="male">6♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFF832E06FCAD1468FC99F410" box="[825,864,1356,1380]" count="3" pageId="34" pageNumber="35" type="female">3♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FCFA1468FC48F410" box="[878,945,1356,1379]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FF832E06FC571468FC4AF4B2" collectingDate="1958-11-20" collectingDateMax="1958-12-27" collectingDateMin="1958-11-20" collectionCode="NHMD, NHM" collectorName="Fr Kuschel" country="Chile" determinerName="A. Newton" elevation="12" location="P. Voillians" municipality="Fr Kuschel" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="18" specimenCount-male="1" stateProvince="Magallanes and Chilean Antarctica" typeStatus="paratype">
|
||||
<specimenCount id="9D7EFD6CFF832E06FC571468FC13F410" box="[963,1002,1356,1380]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FC6C1468FBB2F410" box="[1016,1099,1356,1379]" collectionName="NHMD" pageId="34" pageNumber="35">NHMD</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FF832E06FB171468FB33F410" box="[1155,1226,1356,1379]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
]; [
|
||||
<collectingCountry id="F36F7675FF832E06FB7D1468FAC6F417" box="[1257,1343,1356,1380]" name="Chile" pageId="34" pageNumber="35">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FF832E06FADB1468FBCAF4F0" pageId="34" pageNumber="35">Magallanes and Chilean Antarctica</collectingRegion>
|
||||
]
|
||||
<location id="8EA7603EFF832E06FBD7144FFB52F4F0" LSID="urn:lsid:plazi:treatment:03D187F3FF832E07FC7D1086FB1DF0EE:8EA7603EFF832E06FBD7144FFB52F4F0" box="[1091,1195,1387,1411]" country="Chile" municipality="Fr Kuschel" name="P. Voillians" pageId="34" pageNumber="35" stateProvince="Magallanes and Chilean Antarctica">P.Voillians</location>
|
||||
[Puerto Williams, -54.93, -67.619] 10.1-1959/CHILE
|
||||
<collectorName id="268D5333FF832E06FBF714AFFB22F4D0" box="[1123,1243,1419,1443]" pageId="34" pageNumber="35">Fr Kuschel</collectorName>
|
||||
/4/
|
||||
<collectionCode id="ED69AE20FF832E06FB6D14AFFAC3F4D0" box="[1273,1338,1419,1443]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFF832E06FADE14AFFA8BF4D1" box="[1354,1394,1419,1443]" count="1" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FF832E06FAED14AFFA3BF4D0" box="[1401,1474,1419,1443]" collectionName="Mexico, Jalisco, Guadalajara, Universidad de Guadalajara, Centro de Estudios en Zoologia, Entomologia" pageId="34" pageNumber="35">CZUG</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FF832E06FCF7148EFC50F4B2" box="[867,937,1450,1473]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="35">NHM</collectionCode>
|
||||
].
|
||||
</materialsCitation>
|
||||
All paratypes with label: PARATYPE
|
||||
<taxonomicName id="4C784D66FF832E06FAC0148EFC6CF492" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="carlsbergi">
|
||||
<emphasis id="B90CEAF7FF832E06FAC0148EFC6CF492" italics="true" pageId="34" pageNumber="35">Loncovilius carlsbergi</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF832E06FCAD1707FB6EF70B" blockId="34.[824,1474,1570,1657]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B90CEAF7FF832E06FCAD1707FC66F749" box="[825,927,1571,1594]" italics="true" pageId="34" pageNumber="35">Diagnosis:</emphasis>
|
||||
Pronotum with APP. Elytra yellowish-brown. Sternite III with basal transverse carina evenly converging at an obtuse angle. Aedeagus as in
|
||||
<figureCitation id="13432A60FF832E06FB811745FB68F70B" box="[1045,1169,1633,1657]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="34" pageNumber="35">Figure 11I, J</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF832E06FCAD17BFFB43F63C" blockId="34.[825,1475,1690,1996]" pageId="34" pageNumber="35">
|
||||
<emphasis id="B90CEAF7FF832E06FCAD17BFFC49F7C1" box="[825,944,1691,1714]" italics="true" pageId="34" pageNumber="35">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFF832E06FBF517B9FB82F7C2" box="[1121,1147,1693,1713]" count="1" pageId="34" pageNumber="35" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FF832E06FAE717BFFA7FF7C1" box="[1395,1414,1691,1714]" italics="true" pageId="34" pageNumber="35">N</emphasis>
|
||||
= 6]: FBL = 3.5–3.95 (3.69); TL = 6.47–7.17 (6.71); HW = 0.87– 0.95 (0.9); HL = 0.89–0.95 (0.91); HW/HL = 0.95–1.03 (0.99); PW = 1.19–1.3 (1.25); PL = 1.11–1.2 (1.16); PW/ PL = 1.03–1.13 (1.08); EW = 1.75–1.86 (1.81); EL = 1.46–1.8 (1.61); PW/HW = 1.36–1.44 (1.38).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF832E07FCC11672FE96F1B4" blockId="34.[825,1475,1690,1996]" lastBlockId="35.[112,764,144,1985]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFF832E06FC64167DFBF3F61E" box="[1008,1034,1881,1901]" count="1" pageId="34" pageNumber="35" type="female">♀</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FF832E06FB7E1673FB04F61D" box="[1258,1277,1879,1902]" italics="true" pageId="34" pageNumber="35">N</emphasis>
|
||||
= 6]: FBL = 3.73– 3.94 (3.85); TL = 6.82–8.54 (7.92); HW = 0.91–0.96 (0.94); HL = 0.93–0.97 (0.95); HW/HL = 0.98–1 (0.99); PW = 1.2– 1.36 (1.28); PL = 1.2–1.26 (1.23); PW/PL = 1–1.09 (1.04); EW = 1.71–1.96 (1.84); EL = 1.59–1.74 (1.67); PW/ HW = 1.32–1.42 (1.36).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FF1911EBFEABF036" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">Head, mandibles, legs, last antennomeres, and last palpomeres dark-brown; first palpomeres and antennomeres, and elytra yellow; pronotum yellowish-brown; tergites and sternites with pale posterior margin.</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FF191068FE68F500" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">
|
||||
Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with several setiferous punctures of medium and small-size. Eyes small size (EYL/HL = x = 0.45), from 1.27 to 1.35 times as long as temples (in lateral view) in males and from 1 to 1.16 times in females; distance between eyes about 1.24 times as long as length of eye in males and 1.41 times in females. Antennomeres 2 and 3 subequal in length; antennomeres 4 to 7 subequal in length; antennomeres 8 antennomeres 10 subequal in length; a11 from 1.62 to 2 times as long as a
|
||||
<quantity id="4C809B00FF822E07FE001381FE30F3CE" box="[404,457,677,701]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="10.0">10 in</quantity>
|
||||
males and from
|
||||
<quantity id="4C809B00FF822E07FDE81380FF7CF3AF" metricMagnitude="-2" metricUnit="m" metricValue="4.1275" metricValueMax="4.2926" metricValueMin="3.9624" pageId="35" pageNumber="36" unit="in" value="1.625" valueMax="1.69" valueMin="1.56">1.56 to 1.69 in</quantity>
|
||||
females. Basal and parocular punctures usually single; posterior frontal puncture located posterior to temporal puncture; without small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight, ends distinctly far from gular sutures; postgenal ridge present; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated in males and widely in females. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than previous and more or less subconical.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FF19155FFD87F45C" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">Pronotum slightly transverse, convex, evenly curved, with two PPDS, with an APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to three or five times as long as, diameter of punctures. Mesosternum with five macrosetae arranged in two rows medially.</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FF191413FF3CF759" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">Protibiae sexually dimorphic, with laterodorsal row of thick spines only in females. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 moderately longer than 5, metatarsomere 4 ventrally with apical margin bilobed.</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FF191715FD37F6D1" blockId="35.[112,764,144,1985]" pageId="35" pageNumber="36">
|
||||
Abdominal tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina evenly converging at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with rectangular V-shaped emargination medially (
|
||||
<figureCitation id="13432A60FF822E07FFE817C9FF33F676" box="[124,202,1773,1797]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="35" pageNumber="36">Fig. 14J</figureCitation>
|
||||
); female sternite VIII with arcuate emargination medially; male sternite IX with obtuse emargination medially, its basal portion moderately longer than distal portion; male tergite X with U-shaped emargination medially (
|
||||
<figureCitation id="13432A60FF822E07FDDA166FFD64F610" box="[590,669,1867,1891]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="35" pageNumber="36">Fig. 12J</figureCitation>
|
||||
); female tergite X with U-shaped emargination medially (
|
||||
<figureCitation id="13432A60FF822E07FDFE164FFD43F6F0" box="[618,698,1899,1923]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="35" pageNumber="36">Fig. 13J</figureCitation>
|
||||
); ovipositor, its second gonocoxite with one macroseta medially.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FF19168DFD4FF6B2" blockId="35.[112,764,144,1985]" box="[141,694,1961,1985]" pageId="35" pageNumber="36">
|
||||
Aedeagus as in
|
||||
<figureCitation id="13432A60FF822E07FEBC168DFE5CF6B2" box="[296,421,1961,1985]" captionStart="Figure 11" captionStartId="19.[113,178,1868,1892]" captionTargetBox="[161,1413,148,1837]" captionTargetId="figure-6@19.[156,1417,144,1840]" captionTargetPageId="19" captionText="Figure 11. Aedeagus of Loncovilius species. A–C, L. semiflavus. D–F, L. barclayi. G, H, L. lividipennis. I, J, L. carlsbergi. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally." pageId="35" pageNumber="36">Figure 11I, J</figureCitation>
|
||||
; its total length 1.35–1.45.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FCBE11B4FAADF076" blockId="35.[810,1459,144,262]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B90CEAF7FF822E07FCBE11B4FC60F1DB" box="[810,921,144,168]" italics="true" pageId="35" pageNumber="36">Etymology:</emphasis>
|
||||
The species epithet is in honour of the Denmarkbased Carlsberg beer brewery, whose generous grants and beverages have facilitated much research in the Solodovnikov Lab. In addition, the male aedeagus resembles a bottle opener.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FF822E07FCBE1003FB1DF0EE" blockId="35.[809,1459,295,413]" pageId="35" pageNumber="36">
|
||||
<emphasis id="B90CEAF7FF822E07FCBE1003FB15F04C" box="[810,1260,295,319]" italics="true" pageId="35" pageNumber="36">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FF822E07FB8B1003FB15F04C" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[1055,1260,295,319]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="carlsbergi">Loncovilius carlsbergi</taxonomicName>
|
||||
</emphasis>
|
||||
is known only from the Magallanes and Chilean
|
||||
<collectingCountry id="F36F7675FF822E07FBD31063FB48F02C" box="[1095,1201,327,351]" name="Antarctica" pageId="35" pageNumber="36">Antarctica</collectingCountry>
|
||||
Region of
|
||||
<collectingCountry id="F36F7675FF822E07FA891063FAAEF02C" box="[1309,1367,327,351]" name="Chile" pageId="35" pageNumber="36">Chile</collectingCountry>
|
||||
, where it occurs in lowlands confined to Magellanic subpolar forest ecoregions. It is unknown how it was collected.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,205 +0,0 @@
|
|||
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<mods:mods id="8EA266BCE2495FD6F2FA51747538E7A4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="ED3876EDC359174EC9CE22CB94385751">
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<mods:title id="1588669968D4989751D93C42A62F07D3">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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</mods:role>
|
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<mods:namePart id="AE0832755A54A7A90FF57F541D7CA7F7">Reyes-Hernández, José L.</mods:namePart>
|
||||
<mods:affiliation id="6D9F8D65D6FED07FC9BCE5AFE406C15F">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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||||
<mods:nameIdentifier id="077AC09D8F776D27DB77447B4106E068" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
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</mods:role>
|
||||
<mods:namePart id="5B2899D386AB28E1EDEF1757F0D01137">Hansen, Aslak Kappel</mods:namePart>
|
||||
<mods:affiliation id="4AD109DD58283E8BD83FD156A5C5BC80">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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<mods:namePart id="C3429EE72F25528BC259B865946A30D0">Shaw, Josh Jenkins</mods:namePart>
|
||||
<mods:affiliation id="FDAFC247C59433AD78E865EB0EB77108">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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|
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<mods:namePart id="0580CC9BAFFE8AF1B05463AD34A1018F">Solodovnikov, Alexey</mods:namePart>
|
||||
<mods:affiliation id="01B4A5FDCA4A109B0305829704B0C4A4">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:date id="E828714A51E4937C8149B204DE9E0D26">2024</mods:date>
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<mods:title id="6EB6D09B6706BB3708E10EF48400DB4B">Zool. J. Linn. Soc.</mods:title>
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|
||||
<mods:url id="B00B9D4EC380FE3F1A839AB0C6C54A4A">https://doi.org/10.1093/zoolinnean/zlad143</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="19B1EF8495A06C27442885E9F6B05B19">journal article</mods:classification>
|
||||
<mods:identifier id="A7BA7FA712617641A89F11B6A33787E8" type="DOI">10.1093/zoolinnean/zlad143</mods:identifier>
|
||||
<mods:identifier id="1367335F564CCA85F4D6C626D71FECEA" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="1578CE6B6345CECEEE2C8A024255CE9E" type="Zenodo-Dep">14541979</mods:identifier>
|
||||
<mods:identifier id="ED45D74D20CBB81130EA1FC5EA45A122" type="ZooBank">CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03D187F3FF842E01FF3913F8FB72F0F0" ID-DOI="http://doi.org/10.5281/zenodo.14542017" ID-Zenodo-Dep="14542017" LSID="urn:lsid:plazi:treatment:03D187F3FF842E01FF3913F8FB72F0F0" httpUri="http://treatment.plazi.org/id/03D187F3FF842E01FF3913F8FB72F0F0" lastPageNumber="38" pageId="37" pageNumber="38">
|
||||
<subSubSection id="C362656EFF842E01FF3913F8FD47F385" box="[173,702,731,758]" pageId="37" pageNumber="38" type="nomenclature">
|
||||
<paragraph id="8BC736E5FF842E01FF3913F8FD47F385" blockId="37.[173,702,731,758]" box="[173,702,731,758]" pageId="37" pageNumber="38">
|
||||
<heading id="D08F8189FF842E01FF3913F8FD47F385" bold="true" box="[173,702,731,758]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="3">
|
||||
<emphasis id="B90CEAF7FF842E01FF3913F8FD47F385" bold="true" box="[173,702,731,758]" pageId="37" pageNumber="38">
|
||||
Genus
|
||||
<taxonomicName id="4C784D66FF842E01FF6313F8FDA6F385" authority="Coiffait and Saiz, 1966" authorityName="Coiffait and Saiz" authorityYear="1966" box="[247,607,731,758]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus" status="stat. nov.">
|
||||
<emphasis id="B90CEAF7FF842E01FF6313F8FE95F385" bold="true" box="[247,364,732,758]" italics="true" pageId="37" pageNumber="38">Lienturius</emphasis>
|
||||
<bibRefCitation id="EFE94B14FF842E01FEE513F8FDA6F385" author="Coiffait H & Saiz F" box="[369,607,731,758]" pageId="37" pageNumber="38" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz, 1966</bibRefCitation>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF842E01FDF713F8FD47F385" box="[611,702,732,758]" pageId="37" pageNumber="38">stat. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FE14122FFE15F256" box="[384,492,779,805]" pageId="37" pageNumber="38" type="description">
|
||||
<paragraph id="8BC736E5FF842E01FE14122FFE15F256" blockId="37.[384,492,779,805]" box="[384,492,779,805]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FE14122FFE15F256" bold="true" box="[384,492,779,805]" pageId="37" pageNumber="38">
|
||||
(
|
||||
<figureCitation id="13432A60FF842E01FE1E122FFE18F256" box="[394,481,779,805]" captionStart="Figure 15" captionStartId="38.[129,194,1066,1090]" captionTargetBox="[129,1472,144,1039]" captionTargetId="figure-496@38.[129,1473,144,1039]" captionTargetPageId="38" captionText="Figure 15. Habitus of Amblyopinini genera.A, Apoquedius aeneipennis (Fairmaire and Germain, 1862). B, Lienturius leiocephalus (Solier, 1849). C, Sphingoquedius discoideus (Fairmaire and Germain, 1862)." pageId="37" pageNumber="38">Fig. 15B</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FFE51266FEC1F284" pageId="37" pageNumber="38" type="reference_group">
|
||||
<paragraph id="8BC736E5FF842E01FFE51266FEC1F284" blockId="37.[113,763,834,1015]" pageId="37" pageNumber="38">
|
||||
<bibRefCitation id="EFE94B14FF842E01FFE51266FE6BF229" author="Coiffait H & Saiz F" box="[113,402,834,858]" pageId="37" pageNumber="38" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz 1966: 403</bibRefCitation>
|
||||
[original description; as subgenus of
|
||||
<taxonomicName id="4C784D66FF842E01FF1F1245FF00F20A" box="[139,249,865,889]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FF1F1245FF00F20A" box="[139,249,865,889]" italics="true" pageId="37" pageNumber="38">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
; species included:
|
||||
<taxonomicName id="4C784D66FF842E01FE231246FDDEF20A" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[439,551,866,889]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="aeneipennis">
|
||||
<emphasis id="B90CEAF7FF842E01FE231246FDDEF20A" box="[439,551,866,889]" italics="true" pageId="37" pageNumber="38">aeneipennis</emphasis>
|
||||
</taxonomicName>
|
||||
(as
|
||||
<taxonomicName id="4C784D66FF842E01FDDA1245FD84F20A" authorityName="Blackwelder" authorityYear="1944" box="[590,637,865,889]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="heeri">
|
||||
<emphasis id="B90CEAF7FF842E01FDDA1245FD84F20A" box="[590,637,865,889]" italics="true" pageId="37" pageNumber="38">heeri</emphasis>
|
||||
</taxonomicName>
|
||||
),
|
||||
<taxonomicName id="4C784D66FF842E01FD071245FD0DF20A" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[659,756,865,889]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="discoideus">
|
||||
<emphasis id="B90CEAF7FF842E01FD071245FD0DF20A" box="[659,756,865,889]" italics="true" pageId="37" pageNumber="38">discoideus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C784D66FF842E01FFE512A5FF3EF2EB" box="[113,199,897,920]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="germaini">
|
||||
<emphasis id="B90CEAF7FF842E01FFE512A5FF3EF2EB" box="[113,199,897,920]" italics="true" pageId="37" pageNumber="38">germaini</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C784D66FF842E01FF6812A5FE89F2EA" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Solier" baseAuthorityYear="1849" box="[252,368,897,921]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="leiocephalus">
|
||||
<emphasis id="B90CEAF7FF842E01FF6812A5FE89F2EA" box="[252,368,897,921]" italics="true" pageId="37" pageNumber="38">leiocephalus</emphasis>
|
||||
</taxonomicName>
|
||||
];
|
||||
<bibRefCitation id="EFE94B14FF842E01FE1312A5FDFAF2EA" author="Saiz F" box="[391,515,897,921]" pageId="37" pageNumber="38" pagination="8 - 11" refId="ref38216" refString="Saiz F. Actualizacion de las especies de estafilinidos descritos en la obra de Gay. Noticiario Mensual del Museo Nacional de Historia Natural 1969; 13: 8 - 11." type="journal article" year="1969">Sáiz 1969: 9</bibRefCitation>
|
||||
(subgenus of
|
||||
<taxonomicName id="4C784D66FF842E01FD1B12A5FD0AF2EB" authorityName="Coiffait and Saiz" authorityYear="1966" box="[655,755,897,920]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FD1B12A5FD0AF2EB" box="[655,755,897,920]" italics="true" pageId="37" pageNumber="38">Lienturius</emphasis>
|
||||
</taxonomicName>
|
||||
; notes);
|
||||
<bibRefCitation id="EFE94B14FF842E01FF511284FE91F2CB" author="Saiz F" box="[197,360,928,952]" pageId="37" pageNumber="38" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz 1971: 382</bibRefCitation>
|
||||
, 383 (subgenus of
|
||||
<taxonomicName id="4C784D66FF842E01FDAF1285FD66F2CB" authorityName="Coiffait and Saiz" authorityYear="1966" box="[571,671,929,952]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FDAF1285FD66F2CB" box="[571,671,929,952]" italics="true" pageId="37" pageNumber="38">Lienturius</emphasis>
|
||||
</taxonomicName>
|
||||
; characters;
|
||||
<collectingCountry id="F36F7675FF842E01FF31129BFF1AF2A4" box="[165,227,959,983]" name="Chile" pageId="37" pageNumber="38">Chile</collectingCountry>
|
||||
);
|
||||
<bibRefCitation id="EFE94B14FF842E01FF6112E4FE70F2A4" author="Newton A" box="[245,393,959,983]" pageId="37" pageNumber="38" refId="ref37551" refString="Newton A. StaphBase: Staphyliniformia world catalog database (version Aug 2022). In: Roskov Y, Ower G, Orrell T et al. (eds), Species 2000 and ITIS Catalogue of Life Checklist, 25 th August 2022. Leiden: Naturalis, 2022. (12 Jun 2023, date last accessed)." type="book" year="2022">Newton, 2022</bibRefCitation>
|
||||
[synonym of
|
||||
<taxonomicName id="4C784D66FF842E01FD88129BFED7F284" authority="Scheerpeltz, 1972" authorityName="Scheerpeltz" authorityYear="1972" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="subGenus" subGenus="Apoquedius">
|
||||
<emphasis id="B90CEAF7FF842E01FD88129BFD95F2A4" box="[540,620,959,983]" italics="true" pageId="37" pageNumber="38">Quedius</emphasis>
|
||||
(
|
||||
<emphasis id="B90CEAF7FF842E01FDE9129BFD16F2A4" box="[637,751,959,983]" italics="true" pageId="37" pageNumber="38">Apoquedius</emphasis>
|
||||
) Scheerpeltz, 1972
|
||||
</taxonomicName>
|
||||
].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FFE5153DFD03F543" box="[113,762,1048,1072]" pageId="37" pageNumber="38" type="type_taxon">
|
||||
<paragraph id="8BC736E5FF842E01FFE5153DFD03F543" blockId="37.[113,762,1048,1072]" box="[113,762,1048,1072]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FFE5153DFF14F543" box="[113,237,1049,1072]" italics="true" pageId="37" pageNumber="38">
|
||||
<typeStatus id="54C38847FF842E01FFE5153DFF58F543" box="[113,161,1049,1072]" pageId="37" pageNumber="38">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C784D66FF842E01FF63153CFD87F543" authority="(Solier, 1849)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Solier" baseAuthorityYear="1849" box="[247,638,1048,1072]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="leiocephalus" status="comb. nov.">
|
||||
<emphasis id="B90CEAF7FF842E01FF63153CFE13F543" bold="true" box="[247,490,1048,1072]" italics="true" pageId="37" pageNumber="38">Lienturius leiocephalus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF842E01FE6E153CFD8AF543" author="Solier AJJ" box="[506,627,1048,1072]" pageId="37" pageNumber="38" pagination="105 - 380" refId="ref38501" refString="Solier AJJ. Orden III. Coleopteros. In: Gay C (ed.), Historia Fisica y Politica de Chile. Zoologia. Vol. 4 C. Paris: Gay, 1849, 105 - 380, 414 - 511, pls 1 - 12." type="book chapter" year="1849">Solier, 1849</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
<emphasis id="B90CEAF7FF842E01FD17153CFD03F543" bold="true" box="[643,762,1048,1072]" pageId="37" pageNumber="38">
|
||||
<taxonomicNameLabel id="A23F578CFF842E01FD17153CFD03F543" box="[643,762,1048,1072]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FFE51576FE35F7EE" pageId="37" pageNumber="38" type="diagnosis">
|
||||
<paragraph id="8BC736E5FF842E01FFE51576FE35F7EE" blockId="37.[112,764,1105,1693]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FFE51576FF2EF51A" box="[113,215,1106,1129]" italics="true" pageId="37" pageNumber="38">Diagnosis:</emphasis>
|
||||
The phylogenetic relationship of
|
||||
<taxonomicName id="4C784D66FF842E01FDB91576FD68F51A" authorityName="Coiffait and Saiz" authorityYear="1966" box="[557,657,1106,1129]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FDB91576FD68F51A" box="[557,657,1106,1129]" italics="true" pageId="37" pageNumber="38">Lienturius</emphasis>
|
||||
</taxonomicName>
|
||||
to the rest of the
|
||||
<taxonomicName id="4C784D66FF842E01FF251555FEC6F5FA" authorityName="Seevers" authorityYear="1944" box="[177,319,1137,1161]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="tribe" tribe="Amblyopinini">Amblyopinini</taxonomicName>
|
||||
genera remains unknown, so it is likely that the diagnosis of this genus will change as we wait for the results of our pending phylogenomic research. For the moment the only species of this genus is distinguished based on the combination of the following characters: head with three or more paraocular punctures on each side (not in a row), of which at least one is close to the eye margin and one is far from the eye margin and the remaining paraocular punctures, posterior frontal puncture anterior to temporal puncture; flexible postcoxal hypomeral extension (process) rounded and not interrupted by inferior line; elytra without humeral spines or spine-like setae; two short empodial setae on each tarsomere 5; tergites III–VI with PTBC, sternite III with basal transverse carina converging abruptly at an acute angle with rounded tip.
|
||||
<taxonomicName id="4C784D66FF842E01FE38172DFDE9F753" authorityName="Coiffait and Saiz" authorityYear="1966" box="[428,528,1545,1568]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FE38172DFDE9F753" box="[428,528,1545,1568]" italics="true" pageId="37" pageNumber="38">Lienturius</emphasis>
|
||||
</taxonomicName>
|
||||
is easily distinguished from
|
||||
<taxonomicName id="4C784D66FF842E01FF3E1703FEE1F74C" box="[170,280,1575,1599]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FF3E1703FEE1F74C" box="[170,280,1575,1599]" italics="true" pageId="37" pageNumber="38">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
by the absence of a complete frontoclypeal suture; ventral side of meso- and metatarsi in males and females without pale adhesive setae; presence of protergal glands; presence of PTBC in tergites III to VI.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FFE5179BFD29F7A4" box="[113,720,1727,1751]" pageId="37" pageNumber="38" type="distribution">
|
||||
<paragraph id="8BC736E5FF842E01FFE5179BFD29F7A4" blockId="37.[113,720,1727,1751]" box="[113,720,1727,1751]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FFE5179BFF16F7A4" box="[113,239,1727,1751]" italics="true" pageId="37" pageNumber="38">Distribution:</emphasis>
|
||||
Only known from the Southern zone of
|
||||
<collectingCountry id="F36F7675FF842E01FD05179BFD32F7A4" box="[657,715,1727,1751]" name="Chile" pageId="37" pageNumber="38">Chile</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FFE517CAFB72F0F0" pageId="37" pageNumber="38" type="discussion">
|
||||
<paragraph id="8BC736E5FF842E01FFE517CAFB72F0F0" blockId="37.[113,763,1774,1986]" lastBlockId="37.[810,1459,144,387]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FFE517CAFF36F675" box="[113,207,1774,1798]" italics="true" pageId="37" pageNumber="38">Remarks:</emphasis>
|
||||
The new combination implemented here [
|
||||
<taxonomicName id="4C784D66FF842E01FD0317CBFE81F656" authority="(Solier, 1849)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Solier" baseAuthorityYear="1849" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="leiocephalus">
|
||||
<emphasis id="B90CEAF7FF842E01FD0317CBFF1CF656" italics="true" pageId="37" pageNumber="38">Lienturius leiocephalus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF842E01FF601629FE95F656" author="Solier AJJ" box="[244,364,1805,1829]" pageId="37" pageNumber="38" pagination="105 - 380" refId="ref38501" refString="Solier AJJ. Orden III. Coleopteros. In: Gay C (ed.), Historia Fisica y Politica de Chile. Zoologia. Vol. 4 C. Paris: Gay, 1849, 105 - 380, 414 - 511, pls 1 - 12." type="book chapter" year="1849">Solier, 1849</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
] is supported by the phylogenetic analysis based on eight homoplasious synapomorphies: right mandible with proximal tooth and distal tooth, distal tooth not bifid, space between proximal and distal teeth narrow and smooth (character 24, state 2); labrum, shape of anterior margin (only sclerotized part, excluding apical membrane where present) entire, without emargination (character 28, state 1); three or more paraocular punctures (character 41, state 2); posterior frontal puncture anterior to temporal puncture (character 42, state 0); pronotum with SLSP (character 55, state 1); protarsi with tarsomeres 1–3 as long as wide (character 63, state 0); tergite VI with PTBC (character 89, state 1); sternite III with basal transverse carina converging abruptly at an acute angle with rounded tip (character 99, state 3).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,349 +0,0 @@
|
|||
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<mods:title id="4A567A292A79F5BBBC096FCAF2C2F2EF">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:affiliation id="4B6D679AE22738A265D8D3ACD05F813F">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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<mods:affiliation id="4919B8E091834AEC836FA36F29DF7B02">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:affiliation id="497B35A39180F863D347CBC2632E6395">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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Genus
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<taxonomicName id="4C784D66FF842E01FC3B108AFAE0F0BB" ID-CoL="8XPYL" authority="Bernhauer, 1941" authorityName="Bernhauer" authorityYear="1941" box="[943,1305,430,457]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus" status="stat. nov.">
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Bernhauer, 1941
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</emphasis>
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<paragraph id="8BC736E5FF842E01FBA310C6FB5FF08F" blockId="37.[1079,1190,482,508]" box="[1079,1190,482,508]" pageId="37" pageNumber="38">
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<emphasis id="B90CEAF7FF842E01FBA310C6FB5FF08F" bold="true" box="[1079,1190,482,508]" pageId="37" pageNumber="38">
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(
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<figureCitation id="13432A60FF842E01FBD610C6FB62F08F" box="[1090,1179,482,508]" captionStart="Figure 15" captionStartId="38.[129,194,1066,1090]" captionTargetBox="[129,1472,144,1039]" captionTargetId="figure-496@38.[129,1473,144,1039]" captionTargetPageId="38" captionText="Figure 15. Habitus of Amblyopinini genera.A, Apoquedius aeneipennis (Fairmaire and Germain, 1862). B, Lienturius leiocephalus (Solier, 1849). C, Sphingoquedius discoideus (Fairmaire and Germain, 1862)." pageId="37" pageNumber="38">Fig. 15C</figureCitation>
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)
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</emphasis>
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</paragraph>
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<taxonomicName id="4C784D66FF842E01FCBE133DFB43F342" authority="Bernhauer 1941: 27" authorityName="Bernhauer" authorityPageNumber="27" authorityYear="1941" box="[810,1210,537,561]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="B90CEAF7FF842E01FCBE133DFC3BF342" box="[810,962,537,561]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
|
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Bernhauer 1941: 27
|
||||
</taxonomicName>
|
||||
(original description; species included:
|
||||
<taxonomicName id="4C784D66FF842E01FC7A131CFBCBF323" authorityName="Bernhauer" authorityYear="1941" box="[1006,1074,568,592]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="strandi">
|
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<emphasis id="B90CEAF7FF842E01FC7A131CFBCBF323" box="[1006,1074,568,592]" italics="true" pageId="37" pageNumber="38">strandi</emphasis>
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||||
</taxonomicName>
|
||||
); Klimaszewski
|
||||
<emphasis id="B90CEAF7FF842E01FB7E131DFAE3F323" box="[1258,1306,568,592]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
|
||||
. (1996): 151 (
|
||||
<taxonomicName id="4C784D66FF842E01FCA11373FC34F31C" authorityName="Bernhauer" authorityYear="1941" box="[821,973,599,623]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FCA11373FC34F31C" box="[821,973,599,623]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
; endemic to
|
||||
<collectingCountry id="F36F7675FF842E01FBC2137CFB1FF31C" box="[1110,1254,599,623]" name="New Zealand" pageId="37" pageNumber="38">New Zealand</collectingCountry>
|
||||
);
|
||||
<bibRefCitation id="EFE94B14FF842E01FB6D1373FC0FF3FC" author="Solodovnikov A & Schomann A" pageId="37" pageNumber="38" pagination="443 - 66" refId="ref38647" refString="Solodovnikov A, Schomann A. Revised systematics and biogeography of ' Quediina' of Subsaharan Africa: new phylogenetic insights into the rove beetle tribe Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology 2009; 34: 443 - 66." type="journal article" year="2009">Solodovnikov and Schomann 2009: 34</bibRefCitation>
|
||||
(included in phylogenetic analysis);
|
||||
<bibRefCitation id="EFE94B14FF842E01FAFC1353FB7DF3DD" author="Jenkins Shaw J & Solodovnikov A" pageId="37" pageNumber="38" pagination="1 - 25" refId="ref37050" refString="Jenkins Shaw J, Solodovnikov A. Systematic and biogeographic review of the Staphylinini rove beetles of Lord Howe Island with description of new species and taxonomic changes (Coleoptera, Staphylinidae). ZooKeys 2016; 638: 1 - 25. https: // doi. org / 10.3897 / zookeys. 638.10883" type="journal article" year="2016">Jenkins Shaw and Solodovnikov (2016)</bibRefCitation>
|
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: 22 (absent on Lord Howe Island); Solodovnikov and Brunke (2016): 43 (new combination); Solodovnikov and
|
||||
<bibRefCitation id="EFE94B14FF842E01FBAD13F1FAEFF39E" author="Jenkins Shaw J & Zyla D & Solodovnikov A" box="[1081,1302,725,749]" pageId="37" pageNumber="38" pagination="713 - 22" refId="ref37105" refString="Jenkins Shaw J, Zyla D, Solodovnikov A. A spectacular new genus of Staphylinini rove beetle from the tropical Andes and its phylogenetic assessment (Coleoptera: Staphylinidae). Invertebrate Systematics 2017; 31: 713 - 22." type="journal article" year="2017">Jenkins Shaw (2017)</bibRefCitation>
|
||||
: 318 (notes on relationship);
|
||||
<bibRefCitation id="EFE94B14FF842E01FC2F13D0FB1FF27F" author="Jenkins Shaw J & Zyla D & Solodovnikov A" box="[955,1254,756,780]" pageId="37" pageNumber="38" pagination="713 - 22" refId="ref37105" refString="Jenkins Shaw J, Zyla D, Solodovnikov A. A spectacular new genus of Staphylinini rove beetle from the tropical Andes and its phylogenetic assessment (Coleoptera: Staphylinidae). Invertebrate Systematics 2017; 31: 713 - 22." type="journal article" year="2017">
|
||||
Jenkins Shaw
|
||||
<emphasis id="B90CEAF7FF842E01FBDD13D1FB88F27F" box="[1097,1137,756,780]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
|
||||
. 2017: 714
|
||||
</bibRefCitation>
|
||||
(included in phylogenetic analysis);
|
||||
<bibRefCitation id="EFE94B14FF842E01FC721237FAEAF258" author="Jenkins Shaw J & Zyla D & Solodovnikov A" box="[998,1299,787,811]" pageId="37" pageNumber="38" pagination="430 - 46" refId="ref37147" refString="Jenkins Shaw J, Zyla D, Solodovnikov A. Molecular phylogeny illuminates Amblyopinini (Coleoptera: Staphylinidae) rove beetles as a target for systematic and evolutionary research. Systematic Entomology 2020 a; 45: 430 - 46." type="journal article" year="2020">
|
||||
Jenkins Shaw
|
||||
<emphasis id="B90CEAF7FF842E01FBE21230FB67F258" box="[1142,1182,787,811]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
|
||||
. 2020a: 45
|
||||
</bibRefCitation>
|
||||
(notes on relationship);
|
||||
<bibRefCitation id="EFE94B14FF842E01FC011217FB54F238" author="Jenkins Shaw J & Solodovnikov A & Bai M" box="[917,1197,819,843]" pageId="37" pageNumber="38" pagination="1082 - 8" refId="ref37188" refString="Jenkins Shaw J, Solodovnikov A, Bai M et al. An amblyopinine rove beetle (Coleoptera, Staphylinidae, Staphylininae, Amblyopinini) from the earliest Miocene Foulden Maar fossil-Lagerstatte, New Zealand. Journal of Paleontology 2020 b; 94: 1082 - 8. https: // doi. org / 10.1017 / jpa. 2020.35" type="journal article" year="2020">
|
||||
Jenkins Shaw
|
||||
<emphasis id="B90CEAF7FF842E01FB8B1217FBB1F238" box="[1055,1096,819,843]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
|
||||
. 2020b: 2
|
||||
</bibRefCitation>
|
||||
(description of new fossil species).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FCBE12B1FAB8F2DF" box="[810,1345,916,940]" pageId="37" pageNumber="38" type="type_taxon">
|
||||
<paragraph id="8BC736E5FF842E01FCBE12B1FAB8F2DF" blockId="37.[810,1345,916,940]" box="[810,1345,916,940]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FCBE12B1FC5FF2DF" box="[810,934,917,940]" italics="true" pageId="37" pageNumber="38">
|
||||
<typeStatus id="54C38847FF842E01FCBE12B1FCA3F2DF" box="[810,858,917,940]" pageId="37" pageNumber="38">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C784D66FF842E01FC3F12B0FAC5F2DF" authority="Bernhauer, 1941" authorityName="Bernhauer" authorityYear="1941" box="[939,1340,916,940]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="strandi">
|
||||
<emphasis id="B90CEAF7FF842E01FC3F12B0FB74F2DF" box="[939,1165,916,940]" italics="true" pageId="37" pageNumber="38">Sphingoquedius strandi</emphasis>
|
||||
Bernhauer, 1941
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FCBE12E9FA74F530" pageId="37" pageNumber="38" type="taxon_list">
|
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<paragraph id="8BC736E5FF842E01FCBE12E9FA74F530" blockId="37.[810,1459,973,1091]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FCBE12E9FC3EF296" box="[810,967,973,997]" italics="true" pageId="37" pageNumber="38">Included species:</emphasis>
|
||||
<taxonomicName id="4C784D66FF842E01FC4512E9FACDF296" authority="(Saiz 1971)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Saiz" baseAuthorityYear="1971" box="[977,1332,973,997]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="brevis" status="comb. nov.">
|
||||
<emphasis id="B90CEAF7FF842E01FC4512E9FB41F296" bold="true" box="[977,1208,973,997]" italics="true" pageId="37" pageNumber="38">Sphingoquedius brevis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF842E01FB5C12E9FAD0F296" author="Saiz F" box="[1224,1321,973,997]" pageId="37" pageNumber="38" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz 1971</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
<emphasis id="B90CEAF7FF842E01FAAD12E9FA57F296" bold="true" box="[1337,1454,973,997]" pageId="37" pageNumber="38">
|
||||
<taxonomicNameLabel id="A23F578CFF842E01FAAD12E9FA57F296" box="[1337,1454,973,997]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C784D66FF842E01FCBE12C9FB15F576" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[810,1260,1004,1029]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="discoideus" status="comb. nov.">
|
||||
<emphasis id="B90CEAF7FF842E01FCBE12C9FC54F577" bold="true" box="[810,941,1004,1029]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FCBE12C9FCCEF576" bold="true" box="[810,823,1005,1029]" italics="true" pageId="37" pageNumber="38">S</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF842E01FCD612C8FC54F577" bold="true" box="[834,941,1004,1028]" italics="true" pageId="37" pageNumber="38">discoideus</emphasis>
|
||||
</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF842E01FC2F12C9FB19F576" author="Fairmaire L & Germain P" box="[955,1248,1005,1029]" pageId="37" pageNumber="38" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
<emphasis id="B90CEAF7FF842E01FB6412C9FA93F576" bold="true" box="[1264,1386,1005,1029]" pageId="37" pageNumber="38">
|
||||
<taxonomicNameLabel id="A23F578CFF842E01FB6412C9FA9CF576" box="[1264,1381,1005,1029]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
,
|
||||
</emphasis>
|
||||
<taxonomicName id="4C784D66FF842E01FAFA12C9FBCDF557" authority="Jenkins Shaw et al., 2020 b" authorityName="Jenkins Shaw" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="meto">
|
||||
<emphasis id="B90CEAF7FF842E01FAFA12C9FA82F577" box="[1390,1403,1005,1028]" italics="true" pageId="37" pageNumber="38">S</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF842E01FA1012C9FA4AF577" box="[1412,1459,1005,1028]" italics="true" pageId="37" pageNumber="38">meto</emphasis>
|
||||
<bibRefCitation id="EFE94B14FF842E01FCBE1528FBCDF557" author="Jenkins Shaw J & Solodovnikov A & Bai M" box="[810,1076,1036,1060]" pageId="37" pageNumber="38" pagination="1082 - 8" refId="ref37188" refString="Jenkins Shaw J, Solodovnikov A, Bai M et al. An amblyopinine rove beetle (Coleoptera, Staphylinidae, Staphylininae, Amblyopinini) from the earliest Miocene Foulden Maar fossil-Lagerstatte, New Zealand. Journal of Paleontology 2020 b; 94: 1082 - 8. https: // doi. org / 10.1017 / jpa. 2020.35" type="journal article" year="2020">Jenkins Shaw et al., 2020b</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C784D66FF842E01FBAA1528FAE0F557" authority="(Saiz, 1971)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Saiz" baseAuthorityYear="1971" box="[1086,1305,1036,1060]" class="Insecta" family="Staphylinidae" genus="Cheilocolpus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nanus" status="comb. nov.">
|
||||
<emphasis id="B90CEAF7FF842E01FBAA1528FB6FF550" bold="true" box="[1086,1174,1036,1060]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FBAA1528FBB3F557" bold="true" box="[1086,1098,1036,1060]" italics="true" pageId="37" pageNumber="38">S</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF842E01FBC31528FB6FF550" bold="true" box="[1111,1174,1036,1059]" italics="true" pageId="37" pageNumber="38">nanus</emphasis>
|
||||
</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF842E01FB321528FAF7F557" author="Saiz F" box="[1190,1294,1036,1060]" pageId="37" pageNumber="38" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz, 1971</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
<emphasis id="B90CEAF7FF842E01FA8B1528FA6FF557" bold="true" box="[1311,1430,1036,1060]" pageId="37" pageNumber="38">
|
||||
<taxonomicNameLabel id="A23F578CFF842E01FA8B1528FA6FF557" box="[1311,1430,1036,1060]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C784D66FF842E01FA351528FB8EF530" authority="(Duvivier, 1883)" baseAuthorityName="Duvivier" baseAuthorityYear="1883" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="novaezeelandiae">
|
||||
<emphasis id="B90CEAF7FF842E01FA351528FA57F550" box="[1441,1454,1036,1059]" italics="true" pageId="37" pageNumber="38">S</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF842E01FCBE150FFC3CF530" box="[810,965,1067,1091]" italics="true" pageId="37" pageNumber="38">novaezeelandiae</emphasis>
|
||||
(Duvivier, 1883)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C784D66FF842E01FB141508FA70F530" authority="Bernhauer, 1941" authorityName="Bernhauer" authorityYear="1941" box="[1152,1417,1067,1091]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="strandi">
|
||||
<emphasis id="B90CEAF7FF842E01FB141508FB74F530" box="[1152,1165,1068,1091]" italics="true" pageId="37" pageNumber="38">S</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF842E01FB0C150FFB22F530" box="[1176,1243,1067,1091]" italics="true" pageId="37" pageNumber="38">strandi</emphasis>
|
||||
Bernhauer, 1941
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FCBE1542FAEDF4C4" pageId="37" pageNumber="38" type="diagnosis">
|
||||
<paragraph id="8BC736E5FF842E01FCBE1542FAEDF4C4" blockId="37.[810,1460,1125,1463]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FCBE1542FC69F50E" box="[810,912,1126,1149]" italics="true" pageId="37" pageNumber="38">Diagnosis:</emphasis>
|
||||
The diagnosis of
|
||||
<taxonomicName id="4C784D66FF842E01FBC11541FB14F50E" authorityName="Bernhauer" authorityYear="1941" box="[1109,1261,1125,1149]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FBC11541FB14F50E" box="[1109,1261,1125,1149]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
is likely to change pending phylogenomic results; however, these species can be tentatively placed in
|
||||
<taxonomicName id="4C784D66FF842E01FC6A1580FB6FF5CF" authorityName="Bernhauer" authorityYear="1941" box="[1022,1174,1188,1212]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FC6A1580FB6FF5CF" box="[1022,1174,1188,1212]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
based on the large eyes and presence of patches of pale radiating setae on abdominal tergites III and IV. American
|
||||
<taxonomicName id="4C784D66FF842E01FB8C15C7FB49F588" authorityName="Bernhauer" authorityYear="1941" box="[1048,1200,1251,1275]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FB8C15C7FB49F588" box="[1048,1200,1251,1275]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
are easily distinguished from
|
||||
<taxonomicName id="4C784D66FF842E01FCF01426FC2BF469" box="[868,978,1282,1306]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FCF01426FC2BF469" box="[868,978,1282,1306]" italics="true" pageId="37" pageNumber="38">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
by empodial setae as long as the tarsal claws; ventral side of meso- and metatarsi in males and females without pale adhesive setae; presence of protergal glands; presence of patches of pale radiating setae on abdominal tergites; sternite III with basal transverse carina converges abruptly at an acute angle with rounded tip (projection sides invaginated).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E01FCBE14FFFC82F702" pageId="37" pageNumber="38" type="distribution">
|
||||
<paragraph id="8BC736E5FF842E01FCBE14FFFC82F702" blockId="37.[810,1459,1499,1649]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FCBE14FFFC51F480" box="[810,936,1499,1523]" italics="true" pageId="37" pageNumber="38">Distribution:</emphasis>
|
||||
After the taxonomic changes implemented here,
|
||||
<taxonomicName id="4C784D66FF842E01FCBE14DFFC3BF760" authorityName="Bernhauer" authorityYear="1941" box="[810,962,1531,1555]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FCBE14DFFC3BF760" box="[810,962,1531,1555]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
occurs in
|
||||
<collectingCountry id="F36F7675FF842E01FBAB14DFFB37F760" box="[1087,1230,1531,1555]" name="New Zealand" pageId="37" pageNumber="38">New Zealand</collectingCountry>
|
||||
and southern South America. A single fossil species (
|
||||
<taxonomicName id="4C784D66FF842E01FBEC173EFABCF741" box="[1144,1349,1562,1586]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="meto">
|
||||
<emphasis id="B90CEAF7FF842E01FBEC173EFABCF741" box="[1144,1349,1562,1586]" italics="true" pageId="37" pageNumber="38">Sphingoquedius meto</emphasis>
|
||||
</taxonomicName>
|
||||
) is known from the earliest Miocene of
|
||||
<collectingCountry id="F36F7675FF842E01FBC3171EFB19F722" box="[1111,1248,1593,1617]" name="New Zealand" pageId="37" pageNumber="38">New Zealand</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF842E01FB61171DFC93F702" author="Jenkins Shaw J & Solodovnikov A & Bai M" pageId="37" pageNumber="38" pagination="1082 - 8" refId="ref37188" refString="Jenkins Shaw J, Solodovnikov A, Bai M et al. An amblyopinine rove beetle (Coleoptera, Staphylinidae, Staphylininae, Amblyopinini) from the earliest Miocene Foulden Maar fossil-Lagerstatte, New Zealand. Journal of Paleontology 2020 b; 94: 1082 - 8. https: // doi. org / 10.1017 / jpa. 2020.35" type="journal article" year="2020">
|
||||
Jenkins Shaw
|
||||
<emphasis id="B90CEAF7FF842E01FA11171EFA54F722" box="[1413,1453,1593,1617]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
|
||||
. 2020b
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF842E02FCBE17B4FE04F4D2" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="discussion">
|
||||
<paragraph id="8BC736E5FF842E02FCBE17B4FF2AF5B5" blockId="37.[809,1460,1680,1986]" lastBlockId="38.[129,778,1166,1441]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
|
||||
<emphasis id="B90CEAF7FF842E01FCBE17B4FC71F7DB" box="[810,904,1680,1704]" italics="true" pageId="37" pageNumber="38">Remarks:</emphasis>
|
||||
The new combinations implemented here [
|
||||
<taxonomicName id="4C784D66FF842E01FCA1178BFB6BF7B4" authority="(Saiz, 1971)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Saiz" baseAuthorityYear="1971" box="[821,1170,1711,1735]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nanus">
|
||||
<emphasis id="B90CEAF7FF842E01FCA1178BFBF4F7B4" box="[821,1037,1711,1735]" italics="true" pageId="37" pageNumber="38">Sphingoquedius nanus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF842E01FB89178BFB7FF7B4" author="Saiz F" box="[1053,1158,1711,1735]" pageId="37" pageNumber="38" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz, 1971</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C784D66FF842E01FB0A178BFC93F794" authority="(Saiz, 1971)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Saiz" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="brevis">
|
||||
<emphasis id="B90CEAF7FF842E01FB0A178BFA8DF7B4" box="[1182,1396,1711,1735]" italics="true" pageId="37" pageNumber="38">Sphingoquedius brevis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF842E01FA10178BFCA7F794" author="Saiz F" pageId="37" pageNumber="38" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz, 1971</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C784D66FF842E01FC1617EBFC93F675" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="discoideus">
|
||||
<emphasis id="B90CEAF7FF842E01FC1617EBFB75F794" box="[898,1164,1743,1767]" italics="true" pageId="37" pageNumber="38">Sphingoquedius discoideus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF842E01FB3C17EBFCA7F675" author="Fairmaire L & Germain P" pageId="37" pageNumber="38" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
] are supported by the phylogenetic analysis where species of
|
||||
<taxonomicName id="4C784D66FF842E01FCE11629FBF4F656" authorityName="Bernhauer" authorityYear="1941" box="[885,1037,1805,1829]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FCE11629FBF4F656" box="[885,1037,1805,1829]" italics="true" pageId="37" pageNumber="38">Sphingoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
are united based on two unique synapomorphies: empodial setae longer or at least as long as tarsal claws (character 62, state 1; empodial setae missing in
|
||||
<taxonomicName id="4C784D66FF842E01FAD41668FA58F617" authorityName="Bernhauer" authorityYear="1941" box="[1344,1441,1868,1892]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="strandi">
|
||||
<emphasis id="B90CEAF7FF842E01FAD41668FAB4F610" box="[1344,1357,1868,1891]" italics="true" pageId="37" pageNumber="38">S</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FF842E01FAC91668FA58F617" box="[1373,1441,1868,1892]" italics="true" pageId="37" pageNumber="38">strandi</emphasis>
|
||||
</taxonomicName>
|
||||
); abdomen with tergites III and IV bearing patches of pale radiating setae (character 93, state 2; also see:
|
||||
<bibRefCitation id="EFE94B14FF842E01FB7B16AFFC95F6B1" author="Jenkins Shaw J & Solodovnikov A & Bai M" pageId="37" pageNumber="38" pagination="1082 - 8" refId="ref37188" refString="Jenkins Shaw J, Solodovnikov A, Bai M et al. An amblyopinine rove beetle (Coleoptera, Staphylinidae, Staphylininae, Amblyopinini) from the earliest Miocene Foulden Maar fossil-Lagerstatte, New Zealand. Journal of Paleontology 2020 b; 94: 1082 - 8. https: // doi. org / 10.1017 / jpa. 2020.35" type="journal article" year="2020">
|
||||
Jenkins Shaw
|
||||
<emphasis id="B90CEAF7FF842E01FA1516AFFA54F6D0" box="[1409,1453,1931,1955]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
|
||||
. 2020b
|
||||
</bibRefCitation>
|
||||
: fig. 2.3); and a single homoplasious synapomorphy: postmandibular ridge located close to eye margin (character 16, state 1).
|
||||
</paragraph>
|
||||
<caption id="DF07666DFF872E02FF15150EFD44F52C" pageId="38" pageNumber="39" startId="38.[129,194,1066,1090]" targetBox="[129,1472,144,1039]" targetPageId="38" targetType="figure">
|
||||
<paragraph id="8BC736E5FF872E02FF15150EFD44F52C" blockId="38.[129,1473,1066,1119]" pageId="38" pageNumber="39">
|
||||
<emphasis id="B90CEAF7FF872E02FF15150EFF1CF530" bold="true" box="[129,229,1066,1091]" pageId="38" pageNumber="39">Figure 15.</emphasis>
|
||||
Habitus of
|
||||
<taxonomicName id="4C784D66FF872E02FEC6150FFE2FF530" authorityName="Seevers" authorityYear="1944" box="[338,470,1067,1091]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="tribe" tribe="Amblyopinini">Amblyopinini</taxonomicName>
|
||||
genera. A,
|
||||
<taxonomicName id="4C784D66FF872E02FDAA150FFBBDF530" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[574,1092,1067,1091]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="aeneipennis">
|
||||
<emphasis id="B90CEAF7FF872E02FDAA150FFCEDF531" box="[574,788,1067,1091]" italics="true" pageId="38" pageNumber="39">Apoquedius aeneipennis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF872E02FCB7150FFBC0F530" author="Fairmaire L & Germain P" box="[803,1081,1067,1091]" pageId="38" pageNumber="39" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. B,
|
||||
<taxonomicName id="4C784D66FF872E02FBF1150FFA45F530" authority="(Solier, 1849)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Solier" baseAuthorityYear="1849" box="[1125,1468,1067,1091]" class="Insecta" family="Staphylinidae" genus="Lienturius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="leiocephalus">
|
||||
<emphasis id="B90CEAF7FF872E02FBF1150FFACAF530" box="[1125,1331,1067,1091]" italics="true" pageId="38" pageNumber="39">Lienturius leiocephalus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF872E02FAD5150FFA48F530" author="Solier AJJ" box="[1345,1457,1067,1091]" pageId="38" pageNumber="39" pagination="105 - 380" refId="ref38501" refString="Solier AJJ. Orden III. Coleopteros. In: Gay C (ed.), Historia Fisica y Politica de Chile. Zoologia. Vol. 4 C. Paris: Gay, 1849, 105 - 380, 414 - 511, pls 1 - 12." type="book chapter" year="1849">Solier, 1849</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. C,
|
||||
<taxonomicName id="4C784D66FF872E02FF0F1563FD41F52C" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[155,696,1095,1119]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="discoideus">
|
||||
<emphasis id="B90CEAF7FF872E02FF0F1563FE7EF52C" box="[155,391,1095,1119]" italics="true" pageId="38" pageNumber="39">Sphingoquedius discoideus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF872E02FE021563FD54F52C" author="Fairmaire L & Germain P" box="[406,685,1095,1119]" pageId="38" pageNumber="39" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BC736E5FF872E02FF0815EAFE04F4D2" blockId="38.[129,778,1166,1441]" pageId="38" pageNumber="39">
|
||||
Numerous described and undescribed species from
|
||||
<collectingCountry id="F36F7675FF872E02FD4D15EAFF2DF476" name="New Zealand" pageId="38" pageNumber="39">New Zealand</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="F36F7675FF872E02FE9815C9FE9EF476" box="[268,359,1261,1285]" name="Australia" pageId="38" pageNumber="39">Australia</collectingCountry>
|
||||
may also belong to
|
||||
<taxonomicName id="4C784D66FF872E02FDD615C9FD23F476" authorityName="Bernhauer" authorityYear="1941" box="[578,730,1261,1285]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF872E02FDD615C9FD23F476" box="[578,730,1261,1285]" italics="true" pageId="38" pageNumber="39">Sphingoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
and will be dealt with separately pending further study. Even though the generic placement of the new combinations implemented here may change further, their placement in
|
||||
<taxonomicName id="4C784D66FF872E02FDCC146FFD09F410" authorityName="Bernhauer" authorityYear="1941" box="[600,752,1355,1379]" class="Insecta" family="Staphylinidae" genus="Sphingoquedius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF872E02FDCC146FFD09F410" box="[600,752,1355,1379]" italics="true" pageId="38" pageNumber="39">Sphingoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
is supported by the phylogenetic analysis and can at least be regarded as a step in the right direction.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,216 +0,0 @@
|
|||
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<mods:title id="26DCFE20C66715DB517642F647B09361">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="4767438DA03D97BA8627C1FC0D1A85C5">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="74EE4D8973366A53F1F3C33ACB065FD8">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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Genus
|
||||
<taxonomicName id="4C784D66FF852E00FC5B135AFAEEF3EB" authority="Scheerpeltz, 1972" authorityName="Scheerpeltz" authorityYear="1972" box="[975,1303,638,665]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus" status="stat. nov.">
|
||||
<emphasis id="B90CEAF7FF852E00FC5B135AFBACF3EA" bold="true" box="[975,1109,638,665]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
|
||||
Scheerpeltz, 1972
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFF852E00FA88135AFA8FF3EB" box="[1308,1398,638,664]" pageId="36" pageNumber="37">stat. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
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|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF852E00FBD31389FB4DF3B4" box="[1095,1204,685,711]" pageId="36" pageNumber="37" type="description">
|
||||
<paragraph id="8BC736E5FF852E00FBD31389FB4DF3B4" blockId="36.[1095,1204,685,711]" box="[1095,1204,685,711]" pageId="36" pageNumber="37">
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||||
<emphasis id="B90CEAF7FF852E00FBD31389FB4DF3B4" bold="true" box="[1095,1204,685,711]" pageId="36" pageNumber="37">
|
||||
(
|
||||
<figureCitation id="13432A60FF852E00FBC61389FB50F3B4" box="[1106,1193,685,711]" captionStart="Figure 15" captionStartId="38.[129,194,1066,1090]" captionTargetBox="[129,1472,144,1039]" captionTargetId="figure-496@38.[129,1473,144,1039]" captionTargetPageId="38" captionText="Figure 15. Habitus of Amblyopinini genera.A, Apoquedius aeneipennis (Fairmaire and Germain, 1862). B, Lienturius leiocephalus (Solier, 1849). C, Sphingoquedius discoideus (Fairmaire and Germain, 1862)." pageId="36" pageNumber="37">Fig. 15A</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF852E00FCAD13C0FAB2F229" pageId="36" pageNumber="37" type="reference_group">
|
||||
<paragraph id="8BC736E5FF852E00FCAD13C0FAB2F229" blockId="36.[825,1474,740,858]" pageId="36" pageNumber="37">
|
||||
<taxonomicName id="4C784D66FF852E00FCAD13C0FB51F38F" authority="Scheerpeltz, 1972: 24" authorityName="Scheerpeltz" authorityPageNumber="24" authorityYear="1972" box="[825,1192,740,764]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FCAD13C0FC52F38F" box="[825,939,740,764]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
|
||||
Scheerpeltz, 1972: 24
|
||||
</taxonomicName>
|
||||
, 25 (original description; as subgenus of
|
||||
<taxonomicName id="4C784D66FF852E00FC7B1227FBC6F268" box="[1007,1087,771,795]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FC7B1227FBC6F268" box="[1007,1087,771,795]" italics="true" pageId="36" pageNumber="37">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
; included species:
|
||||
<taxonomicName id="4C784D66FF852E00FA891220FA74F268" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" box="[1309,1421,772,795]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aeneipennis">
|
||||
<emphasis id="B90CEAF7FF852E00FA891220FA74F268" box="[1309,1421,772,795]" italics="true" pageId="36" pageNumber="37">aeneipennis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C784D66FF852E00FCAD1207FC61F248" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1912" box="[825,920,803,827]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="piciformis">
|
||||
<emphasis id="B90CEAF7FF852E00FCAD1207FC61F248" box="[825,920,803,827]" italics="true" pageId="36" pageNumber="37">piciformis</emphasis>
|
||||
</taxonomicName>
|
||||
);
|
||||
<bibRefCitation id="EFE94B14FF852E00FC281207FB6BF248" author="Herman LH" box="[956,1170,803,827]" pageId="36" pageNumber="37" pagination="1 - 82" refId="ref36950" refString="Herman LH. Nomenclatural changes in the Staphylinidae (Insecta: Coleoptera). Bulletin of the American Museum of Natural History 2001 a; 264: 1 - 82." type="journal article" year="2001">Herman 2001a: 24</bibRefCitation>
|
||||
(synonym of
|
||||
<taxonomicName id="4C784D66FF852E00FAD61207FA49F248" box="[1346,1456,803,827]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FAD61207FA49F248" box="[1346,1456,803,827]" italics="true" pageId="36" pageNumber="37">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
);
|
||||
<bibRefCitation id="EFE94B14FF852E00FCAD1266FC35F229" author="Newton A" box="[825,972,834,858]" pageId="36" pageNumber="37" refId="ref37551" refString="Newton A. StaphBase: Staphyliniformia world catalog database (version Aug 2022). In: Roskov Y, Ower G, Orrell T et al. (eds), Species 2000 and ITIS Catalogue of Life Checklist, 25 th August 2022. Leiden: Naturalis, 2022. (12 Jun 2023, date last accessed)." type="book" year="2022">Newton, 2022</bibRefCitation>
|
||||
(synonym of
|
||||
<taxonomicName id="4C784D66FF852E00FBCD1266FABEF229" authorityName="Coiffait & Saiz" authorityYear="1966" box="[1113,1351,834,858]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="subGenus" subGenus="Lienturius">
|
||||
<emphasis id="B90CEAF7FF852E00FBCD1266FB3EF229" box="[1113,1223,834,858]" italics="true" pageId="36" pageNumber="37">Loncovilius</emphasis>
|
||||
(
|
||||
<emphasis id="B90CEAF7FF852E00FB431267FAC2F229" box="[1239,1339,835,858]" italics="true" pageId="36" pageNumber="37">Lienturius</emphasis>
|
||||
)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF852E00FCAD1258FC0DF2C0" pageId="36" pageNumber="37" type="type_taxon">
|
||||
<paragraph id="8BC736E5FF852E00FCAD1258FC0DF2C0" blockId="36.[825,1474,891,947]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B90CEAF7FF852E00FCAD1258FC57F2E0" box="[825,942,892,915]" italics="true" pageId="36" pageNumber="37">
|
||||
<typeStatus id="54C38847FF852E00FCAD1258FC90F2E0" box="[825,873,892,915]" pageId="36" pageNumber="37">Type</typeStatus>
|
||||
genus:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C784D66FF852E00FC22125FFC81F2C0" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aeneipennis" status="comb. nov.">
|
||||
<emphasis id="B90CEAF7FF852E00FC22125FFB4FF2E0" bold="true" box="[950,1206,891,915]" italics="true" pageId="36" pageNumber="37">Apoquedius aeneipennis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF852E00FB5D125FFC94F2C0" author="Fairmaire L & Germain P" pageId="36" pageNumber="37" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
<emphasis id="B90CEAF7FF852E00FCEA12BFFC0DF2C0" bold="true" box="[894,1012,923,947]" pageId="36" pageNumber="37">
|
||||
<taxonomicNameLabel id="A23F578CFF852E00FCEA12BFFC0DF2C0" box="[894,1012,923,947]" pageId="36" pageNumber="37" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF852E00FCAD12F0FC49F558" pageId="36" pageNumber="37" type="taxon_list">
|
||||
<paragraph id="8BC736E5FF852E00FCAD12F0FC49F558" blockId="36.[825,1474,980,1067]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B90CEAF7FF852E00FCAD12F0FC1DF29F" box="[825,996,980,1004]" italics="true" pageId="36" pageNumber="37">Included species:</emphasis>
|
||||
<taxonomicName id="4C784D66FF852E00FB9212F0FC18F578" authority="(Fairmaire and Germain, 1862)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Fairmaire and Germain" baseAuthorityYear="1862" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aeneipennis" status="comb. nov.">
|
||||
<emphasis id="B90CEAF7FF852E00FB9212F0FAF7F29F" bold="true" box="[1030,1294,980,1004]" italics="true" pageId="36" pageNumber="37">Apoquedius aeneipennis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FF852E00FABD12F0FC2FF578" author="Fairmaire L & Germain P" pageId="36" pageNumber="37" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
<emphasis id="B90CEAF7FF852E00FC7D12D7FB9BF578" bold="true" box="[1001,1122,1011,1035]" pageId="36" pageNumber="37">
|
||||
<taxonomicNameLabel id="A23F578CFF852E00FC7D12D7FB9BF578" box="[1001,1122,1011,1035]" pageId="36" pageNumber="37" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
,
|
||||
<taxonomicName id="4C784D66FF852E00FBFB12D7FA3BF578" authority="(Bernhauer, 1912)" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" baseAuthorityName="Bernhauer" baseAuthorityYear="1912" box="[1135,1474,1011,1035]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="piciformis" status="comb. nov.">
|
||||
<emphasis id="B90CEAF7FF852E00FBFB12D7FB01F578" bold="true" box="[1135,1272,1011,1035]" italics="true" pageId="36" pageNumber="37">A. piciformis</emphasis>
|
||||
(Bernhauer, 1912)
|
||||
</taxonomicName>
|
||||
<emphasis id="B90CEAF7FF852E00FCAD1537FC49F558" bold="true" box="[825,944,1043,1067]" pageId="36" pageNumber="37">
|
||||
<taxonomicNameLabel id="A23F578CFF852E00FCAD1537FC49F558" box="[825,944,1043,1067]" pageId="36" pageNumber="37" rank="species">comb. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF852E00FCAD1569FB8EF70B" pageId="36" pageNumber="37" type="diagnosis">
|
||||
<paragraph id="8BC736E5FF852E00FCAD1569FB8EF70B" blockId="36.[825,1476,1100,1657]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B90CEAF7FF852E00FCAD1569FBD1F517" box="[825,1064,1100,1124]" italics="true" pageId="36" pageNumber="37">
|
||||
Diagnosis:
|
||||
<taxonomicName id="4C784D66FF852E00FC221568FBD1F517" authorityName="Scheerpeltz" authorityYear="1972" box="[950,1064,1100,1124]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Apoquedius</taxonomicName>
|
||||
</emphasis>
|
||||
can be diagnosed based on the head with frontoclypeal suture distinct only laterally; wide neck; paraocular punctures present; apical segment of maxillary palpi fusiform; apical segment of labial palpi fusiform, widest at the middle portion; apical labial palpomere distinctly longer than penultimate labial palpomere; flexible postcoxal hypomeral extension or process elongated and thin; two short empodial setae on each tarsal 5 segment; metacoxae with more than four spines on the posterior surface; abdomen with apical row of setae at the anterior margin of tergites and sternites III–VI; anterior margin of tergites III–VI distinctly crenulate.
|
||||
<taxonomicName id="4C784D66FF852E00FB5314A1FAC0F4EE" authorityName="Scheerpeltz" authorityYear="1972" box="[1223,1337,1413,1437]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FB5314A1FAC0F4EE" box="[1223,1337,1413,1437]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
is easily distinguished from
|
||||
<taxonomicName id="4C784D66FF852E00FC711481FBAAF4CE" box="[997,1107,1445,1469]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FC711481FBAAF4CE" box="[997,1107,1445,1469]" italics="true" pageId="36" pageNumber="37">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
by the shape of the maxillary palps, which are fusiform; the shape of the postcoxal process which is thin and elongated; ventral side of meso- and metatarsi in males and females without pale adhesive setae; presence of protergal glands; presence of PTBC in tergites III to V; the most apical row of setae at the apical margin of tergites and sternites III to VI giving a crenulated appearance.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF852E00FCAD17BEFC5CF7A2" pageId="36" pageNumber="37" type="distribution">
|
||||
<paragraph id="8BC736E5FF852E00FCAD17BEFC5CF7A2" blockId="36.[825,1474,1690,1745]" pageId="36" pageNumber="37">
|
||||
<emphasis id="B90CEAF7FF852E00FCAD17BEFC4EF7C1" box="[825,951,1690,1714]" italics="true" pageId="36" pageNumber="37">Distribution:</emphasis>
|
||||
As of now,
|
||||
<taxonomicName id="4C784D66FF852E00FBA917BEFB56F7C1" authorityName="Scheerpeltz" authorityYear="1972" box="[1085,1199,1690,1714]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FBA917BEFB56F7C1" box="[1085,1199,1690,1714]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
is restricted to
|
||||
<collectingCountry id="F36F7675FF852E00FACE17BEFA6BF7C1" box="[1370,1426,1690,1714]" name="Chile" pageId="36" pageNumber="37">Chile</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="F36F7675FF852E00FCAD179DFC58F7A2" box="[825,929,1721,1745]" name="Argentina" pageId="36" pageNumber="37">Argentina</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFF852E01FCAD17D7FE64F3CE" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="discussion">
|
||||
<paragraph id="8BC736E5FF852E01FCAD17D7FE64F3CE" blockId="36.[825,1474,1779,1959]" lastBlockId="37.[112,764,144,701]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
|
||||
<emphasis id="B90CEAF7FF852E00FCAD17D7FC6EF678" box="[825,919,1779,1803]" italics="true" pageId="36" pageNumber="37">Remarks:</emphasis>
|
||||
The monophyly of
|
||||
<taxonomicName id="4C784D66FF852E00FB1017D7FB0FF678" authorityName="Scheerpeltz" authorityYear="1972" box="[1156,1270,1779,1803]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF852E00FB1017D7FB0FF678" box="[1156,1270,1779,1803]" italics="true" pageId="36" pageNumber="37">Apoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
is supported by a unique synapomorphy: flexible postcoxal hypomeral extension or process elongated and thin (character 58, state 1); and 11 homoplastic synapomorphies: head with frontoclypeal suture distinct only laterally (character 1, state 1); infraorbital ridge (IOR) not merged with postgenal ridge (PGR) (character 17, state 1); apical segment of maxillary palpi more or less fusiform, widest at its middle portion (character 29, state 0); apical segment of labial palpi more or less fusiform, widest at the middle portion (character 32, state 0); apical labial palpomere distinctly longer than penultimate labial palpomere (character 33, state 1); two parocular punctures on each side (character 41, state 1); basisternum from about as long as to moderately longer than furcasternum (character 46, state 0); protibiae, apical tibial spur distinctly outside tibial margin (character 66, state 1); more than four spines on the posterior surface of metacoxae (character 73, state 1); mesothorax with sternopleural (anapleural) suture transverse, or nearly transverse (very slightly oblique) (character 79, state 0); abdomen with apical row of setae at the anterior margin of tergites and sternites III–VI and anterior margin of tergites III–VI distinctly crenulate (character 90, state 1). Several undescribed species from southern South America and one described species from
|
||||
<collectingRegion id="49BCF807FF842E01FEF913A1FE36F3EE" box="[365,463,645,669]" country="Australia" name="Tasmania" pageId="37" pageNumber="38">Tasmania</collectingRegion>
|
||||
likely belong to
|
||||
<taxonomicName id="4C784D66FF842E01FD1C13A1FD03F3EE" authorityName="Scheerpeltz" authorityYear="1972" box="[648,762,645,669]" class="Insecta" family="Staphylinidae" genus="Apoquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF842E01FD1C13A1FD03F3EE" box="[648,762,645,669]" italics="true" pageId="37" pageNumber="38">Apoquedius</emphasis>
|
||||
</taxonomicName>
|
||||
pending further examination.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
File diff suppressed because one or more lines are too long
|
|
@ -1,201 +0,0 @@
|
|||
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<mods:title id="6CBCBF541E311736016D746BF91EE899">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:affiliation id="98DAA8CED925D81F7FEC1103C8CDE344">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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|
||||
<paragraph id="8BC736E5FFB62E33FDFC1373FC45F303" blockId="23.[140,1434,598,1909]" box="[616,956,598,625]" pageId="23" pageNumber="24">
|
||||
Key to the species of
|
||||
<taxonomicName id="4C784D66FFB62E33FCDD1372FC45F303" ID-CoL="8X7JJ" authority="Germain, 1903" authorityName="Germain" authorityYear="1903" box="[841,956,598,624]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FFB62E33FCDD1372FC45F303" box="[841,956,598,624]" italics="true" pageId="23" pageNumber="24">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFB62E3CFF1813B3FA5EF08E" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="key">
|
||||
<key id="FFE961D4FFB62E3CFF1813B3FA5EF08E" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
|
||||
<keyStep id="308C2E40FFB62E33FF1813B3FA61F2D9" pageId="23" pageNumber="24">
|
||||
<paragraph id="8BC736E5FFB62E33FF1813B3FA61F27E" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF1813B3FA61F27E" pageId="23" pageNumber="24">1. Head transverse, round in outline; disc with coarse non-setiferous punctures (Fig. 7C); postmandibular ridge distinctly separated from eye margin; PMP conspicuous (Fig. 8E); mandibles with a shallow dorso-lateral groove; right mandible with distal tooth bifid; left mandible only with proximal, non-bifid tooth. Abdominal tergites III–V with two patches of setiferous punctures separated by an impunctate area (Fig. 9A); second gonocoxite without macrosetae (Fig. 9F)..................................2</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB62E33FF181231FA61F2D9" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF181231FA61F2D9" pageId="23" pageNumber="24">- Head about as wide as long or elongate, its outline mostly longitudinally elliptical; disc without coarse non-setiferous punctures (Fig. 7D, E); postmandibular ridge situated very close to eye margin; PMP reduced and hard to see (Fig. 8F); mandibles without dorso-lateral groove; right mandible with distal tooth not bifid; left mandible with proximal and distal teeth, the latter not bifid, short and truncated, space between proximal and distal teeth rough. Abdominal tergites III–V not as described above; second gonocoxite with at least one macroseta (Fig. 9G, H) ................................................................................4</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="308C2E40FFB62E33FF181295FA61F554" pageId="23" pageNumber="24">
|
||||
<paragraph id="8BC736E5FFB62E33FF181295FA61F29A" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF181295FA61F29A" pageId="23" pageNumber="24">
|
||||
2. Disc of head without microsculpture; antennal insertions situated closer to frontoclypeus than to eye. Aedeagus as in Figure 10A–C........................................................................................................................................
|
||||
<emphasis id="B90CEAF7FFB62E33FC6B12F5FB31F29B" box="[1023,1224,977,1001]" italics="true" pageId="23" pageNumber="24">Loncovilius germaini</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FFB62E33FB4312F5FA74F29A" author="Scheerpeltz O" box="[1239,1421,977,1001]" pageId="23" pageNumber="24" pagination="989 - 1500" refId="ref38278" refString="Scheerpeltz O. Staphylinidae VII. In: Schenkling S (ed.), Coleopterorum Catalogus, Vol. 6. Berlin: Junk, 1933, 989 - 1500." type="book chapter" year="1933">Scheerpeltz, 1933</bibRefCitation>
|
||||
)
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB62E33FF1812D4FA61F554" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF1812D4FA61F554" pageId="23" pageNumber="24">- Disc of head with microsculpture; antennal insertions situated at equal distance to frontoclypeus and to the eye, or closer to the eye. Aedeagus not as above ....................................................................................................................................................................3</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="308C2E40FFB62E33FF18150BFA60F5D6" pageId="23" pageNumber="24">
|
||||
<paragraph id="8BC736E5FFB62E33FF18150BFA60F515" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF18150BFA60F515" pageId="23" pageNumber="24">
|
||||
3. Microsculpture on dorsal surface of head meshed to isodiametric. Pronotum and elytral disc with few and scattered coarse non-setiferous punctures. Aedeagus as in Figure 10G, H .................................................................
|
||||
<emphasis id="B90CEAF7FFB62E33FBFE156AFABBF515" box="[1130,1346,1102,1126]" italics="true" pageId="23" pageNumber="24">Loncovilius impunctus</emphasis>
|
||||
<emphasis id="B90CEAF7FFB62E33FAD3156AFA60F515" bold="true" box="[1351,1433,1102,1126]" pageId="23" pageNumber="24">sp. nov.</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB62E33FF18154AFA60F5D6" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF18154AFA60F5D6" pageId="23" pageNumber="24">
|
||||
- Microsculpture on dorsal surface of head as transverse waves. Pronotum and elytral disc with conspicuous coarse non-setiferous punctures. Aedeagus as in Figure 10D–F...........................................................................
|
||||
<emphasis id="B90CEAF7FFB62E33FBF115A9FABBF5D6" box="[1125,1346,1165,1189]" italics="true" pageId="23" pageNumber="24">Loncovilius hammondi</emphasis>
|
||||
<emphasis id="B90CEAF7FFB62E33FAD315A9FA60F5D6" bold="true" box="[1351,1433,1165,1189]" pageId="23" pageNumber="24">sp. nov.</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="308C2E40FFB62E33FF181588FA61F452" pageId="23" pageNumber="24">
|
||||
<paragraph id="8BC736E5FFB62E33FF181588FA61F590" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF181588FA61F590" pageId="23" pageNumber="24">4. Head: nuchal ridge present dorsally and laterally. Abdomen: tergites III and IV with more than three rows of punctures, anterior half punctate (Fig. 9B, E) ...................................................................................................................................................................5</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB62E33FF1815CFFA61F452" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF1815CFFA61F452" pageId="23" pageNumber="24">- Head: nuchal ridge missing dorsally, present laterally. Abdomen: tergites III and IV with less than three rows of punctures, anterior half impunctate (Fig. 9C, D).........................................................................................................................................................7</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="308C2E40FFB62E33FF18140DFA61F48E" pageId="23" pageNumber="24">
|
||||
<paragraph id="8BC736E5FFB62E33FF18140DFA60F4EC" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF18140DFA60F4EC" pageId="23" pageNumber="24">
|
||||
5. Protibia in both sexes without a row of thick laterodorsal spines; first metatarsomere shorter than fifth or at most both tarsomeres subequal. Abdominal tergites III and IV with several rows of sparse and fine punctures of moderate density (Fig. 9B); sternite III with basal transverse carina strongly arcuate; aedeagus as in Figure 10I–K ........................................................... ..........................................................................................................................................................................
|
||||
<emphasis id="B90CEAF7FFB62E33FBE114A3FAB8F4EC" box="[1141,1345,1415,1439]" italics="true" pageId="23" pageNumber="24">Loncovilius variabilis</emphasis>
|
||||
<emphasis id="B90CEAF7FFB62E33FAD314A3FA60F4EC" bold="true" box="[1351,1433,1415,1439]" pageId="23" pageNumber="24">sp. nov.</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB62E33FF181483FA61F48E" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF181483FA61F48E" pageId="23" pageNumber="24">- Protibia with a row of thick laterodorsal spines present at least in females; first metatarsomere moderately longer than fifth. Abdominal tergites III and IV with several rows of punctures of moderate size and density (Fig. 9E); sternite III with basal transverse carina projecting medially in a sharp or obtuse angle; aedeagus not as above ...............................................................6</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="308C2E40FFB62E33FF181721FA60F7CA" pageId="23" pageNumber="24">
|
||||
<paragraph id="8BC736E5FFB62E33FF181721FA61F728" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF181721FA61F728" pageId="23" pageNumber="24">
|
||||
6. Pronotum rarely with additional paired punctures adjacent to anterior marginal row of setiferous punctures (Fig. 8A). Abdominal sternite III with basal transverse carina sharply pointed medioapically; aedeagus as in Figure 11G, H ........................ ...........................................................................................................................
|
||||
<emphasis id="B90CEAF7FFB62E33FCE41767FBA8F728" box="[880,1105,1603,1627]" italics="true" pageId="23" pageNumber="24">Loncovilius lividipennis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FFB62E33FBF41767FA74F728" author="Fairmaire L & Germain P" box="[1120,1421,1603,1627]" pageId="23" pageNumber="24" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB62E33FF181747FA60F7CA" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF181747FA60F7CA" pageId="23" pageNumber="24">
|
||||
- Pronotum always with additional paired punctures adjacent to anterior marginal row of setiferous punctures (Fig. 8A). Abdominal sternite III with basal transverse carina evenly converging at an obtuse angle; aedeagus as in Figure 11I, J ..........................................................................................................................................................................
|
||||
<emphasis id="B90CEAF7FFB62E33FBE71785FABBF7CA" box="[1139,1346,1697,1721]" italics="true" pageId="23" pageNumber="24">Loncovilius carlsbergi</emphasis>
|
||||
<emphasis id="B90CEAF7FFB62E33FAD31785FA60F7CA" bold="true" box="[1351,1433,1697,1721]" pageId="23" pageNumber="24">sp. nov.</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="308C2E40FFB62E33FF1817E5FA61F606" pageId="23" pageNumber="24">
|
||||
<paragraph id="8BC736E5FFB62E33FF1817E5FA61F664" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF1817E5FA61F664" pageId="23" pageNumber="24">7. Head and pronotum microsculpture consisting of isodiametric meshes. Head elongate. Pronotum without sublateral setiferous punctures. Males: aedeagus, paramere strongly produced over apex of median lobe. Females: second gonocoxite with three or more macrosetae (Fig. 9G) ...........................................................................................................................................................8</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB62E33FF18163BFA61F606" blockId="23.[140,1434,598,1909]" pageId="23" pageNumber="24">
|
||||
<keyLead id="308995D0FFB62E33FF18163BFA61F606" pageId="23" pageNumber="24">- Head and pronotum with very fine microsculpture of long transverse and diagonal waves. Head about as wide as long. Pronotum with sublateral setiferous punctures. Males: aedeagus, paramere slightly shorter or produced over apex of median lobe. Females: second gonocoxite with one or two macrosetae (Fig. 9H).........................................................................................9</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="308C2E40FFB92E3CFF0F11A8FA51F032" pageId="24" pageNumber="25">
|
||||
<paragraph id="8BC736E5FFB92E3CFF0F11A8FA51F190" blockId="24.[155,1448,140,509]" pageId="24" pageNumber="25">
|
||||
<keyLead id="308995D0FFB92E3CFF0F11A8FA51F190" pageId="24" pageNumber="25">
|
||||
8. Head with small setiferous punctures between frontoclypeal and anterior frontal punctures. Pronotum outline evenly curved. Protibia with a row of thick laterodorsal spines only in females. Elytra rarely with weak bright metallic iridescence ....................................................................................................................................................................
|
||||
<emphasis id="B90CEAF7FFB92E3CFBF711EFFAA8F190" box="[1123,1361,203,227]" italics="true" pageId="24" pageNumber="25">Loncovilius cantharoides</emphasis>
|
||||
<emphasis id="B90CEAF7FFB92E3CFAC211EFFA51F190" bold="true" box="[1366,1448,203,227]" pageId="24" pageNumber="25">sp. nov.</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFF0F11CFFA51F032" blockId="24.[155,1448,140,509]" pageId="24" pageNumber="25">
|
||||
<keyLead id="308995D0FFB92E3CFF0F11CFFA51F032" pageId="24" pageNumber="25">
|
||||
- Head without small setiferous punctures between frontoclypeal and anterior frontal punctures. Pronotum outline hexagonal. Protibia in both sexes without a row of thick laterodorsal spines. Elytra with bright greenish, bluish, or purplish metallic iridescence (Figs 1A, 6A).........................................................................................
|
||||
<emphasis id="B90CEAF7FFB92E3CFB8D100DFAF6F032" box="[1049,1295,297,321]" italics="true" pageId="24" pageNumber="25">Loncovilius edwardsianus</emphasis>
|
||||
(Korge, 1963)
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="308C2E40FFB92E3CFF0F106CFA5EF08E" pageId="24" pageNumber="25">
|
||||
<paragraph id="8BC736E5FFB92E3CFF0F106CFA51F0EC" blockId="24.[155,1448,140,509]" pageId="24" pageNumber="25">
|
||||
<keyLead id="308995D0FFB92E3CFF0F106CFA51F0EC" pageId="24" pageNumber="25">
|
||||
9 Elytra with brown-black marking around mesoscutellum and on suture (Fig. 6D). All femora reddish-brown. Male sternite IX without median emargination at apex; male tergite X trilobed (Fig.12H); aedeagus as in Figure 11D–F .............................. ............................................................................................................................................................................
|
||||
<emphasis id="B90CEAF7FFB92E3CFB0510A3FAA8F0EC" box="[1169,1361,391,415]" italics="true" pageId="24" pageNumber="25">Loncovilius barclayi</emphasis>
|
||||
<emphasis id="B90CEAF7FFB92E3CFAC210A3FA51F0EC" bold="true" box="[1366,1448,391,415]" pageId="24" pageNumber="25">sp. nov.</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFF0F1083FA5EF08E" blockId="24.[155,1448,140,509]" pageId="24" pageNumber="25">
|
||||
<keyLead id="308995D0FFB92E3CFF0F1083FA5EF08E" pageId="24" pageNumber="25">
|
||||
- Elytra without dark mark around mesoscutellum and on suture. All femora yellow or yellowish-brown; male sternite IX with median emargination at apex; male tergite X emarginate medio-apically (Fig. 12G); aedeagus as in Figure 11A–C .............................................................................................................................
|
||||
<emphasis id="B90CEAF7FFB92E3CFC1E10C1FB99F08E" box="[906,1120,485,509]" italics="true" pageId="24" pageNumber="25">Loncovilius semiflavus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B14FFB92E3CFBE410C1FA65F08E" author="Fairmaire L & Germain P" box="[1136,1436,485,509]" pageId="24" pageNumber="25" pagination="405 - 56" refId="ref36627" refString="Fairmaire L, Germain P. Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France 1862; 4: 405 - 56." type="journal article" year="1862">Fairmaire and Germain, 1862</bibRefCitation>
|
||||
)
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</key>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,285 +0,0 @@
|
|||
<document id="C11C07884DFBD03BA35AD78BC6EEAC72" ID-DOI="10.1093/zoolinnean/zlad143" ID-ISSN="0024-4082" ID-Zenodo-Dep="14541979" ID-ZooBank="CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732839935034" checkinUser="plazi" docAuthor="Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins & Solodovnikov, Alexey" docDate="2024" docId="03D187F3FFB82E3EFE8E175FFEF9F74C" docLanguage="en" docName="zlad143.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlad143" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Loncovilius hammondi Reyes-Hernández, Hansen, Shaw & Solodovnikov, 2024, sp.nov." docType="treatment" docVersion="2" lastPageNumber="27" masterDocId="FFE8FF8BFFA12E24FF941124FFF9F173" masterDocTitle="Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="26" updateTime="1734824012822" updateUser="ExternalLinkService">
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<mods:titleInfo id="5CFCFAA33BC7359DC3FDAE9C8576B011">
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<mods:title id="02685188EED87427E93CF6B3C5A8D6C6">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="1E86C738B58DE7F204FC7EF2B0DD2A56">Reyes-Hernández, José L.</mods:namePart>
|
||||
<mods:affiliation id="99840F2148EC000F32457D644C5451C7">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="0AF8A97BA45066BD430F30712A34165C" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
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</mods:role>
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||||
<mods:namePart id="E9703B50801EF488660D9F1ABA5ABF9A">Hansen, Aslak Kappel</mods:namePart>
|
||||
<mods:affiliation id="3BD17445BAA8EAF5B8EA1DA781007C3F">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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</mods:role>
|
||||
<mods:namePart id="267F4F0D426ECF02BC41D64DB579984A">Shaw, Josh Jenkins</mods:namePart>
|
||||
<mods:affiliation id="D7A0809073490EA1541D5E0951BECB0D">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
|
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</mods:name>
|
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|
||||
<mods:roleTerm id="1E4DF1492938876464FE6E29CF5C3B78">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="77A0FAC02E03C2648430ACE6DD184AFF">Solodovnikov, Alexey</mods:namePart>
|
||||
<mods:affiliation id="5FA1B9ED6D42A3D715BD717D8C136820">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:typeOfResource id="F340B80FD873D5FC8C6F42B3C5069622">text</mods:typeOfResource>
|
||||
<mods:relatedItem id="301CFE45102A0A77BE83298CA20BA162" type="host">
|
||||
<mods:titleInfo id="90855BCA57D6FF2896EA4ACD58E03321">
|
||||
<mods:title id="F6CFEC3DDEAD5D27884400AAD4AF2D62">Zoological Journal of the Linnean Society</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="E2DB2304BF0A272139A4C050F2C8FBA9">
|
||||
<mods:date id="9907780F3BC73546157022289FE1A278">2024</mods:date>
|
||||
<mods:detail id="D6085189713AEE8D5AF7970765BFDCD9" type="series">
|
||||
<mods:title id="E748ED44E5FA0805137DAFB55378846A">Zool. J. Linn. Soc.</mods:title>
|
||||
</mods:detail>
|
||||
<mods:detail id="83CFDBE025939464D3E45F688CA2D59F" type="pubDate">
|
||||
<mods:number id="8DD916FED5B4171E84344D323614DF3C">2023-10-28</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="F998DB0A84A19FAB7E9E1B7BE91E9136" type="volume">
|
||||
<mods:number id="71C83463C5DA2ACD37C487F7505A6C89">202</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="F7189EE8104CA8FA4A6269C31376CAA6" type="issue">
|
||||
<mods:number id="A37D2E42377A122F309CDE91C199107F">1</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="EF06745D4779E0E62D916B0EFC41E66B" unit="page">
|
||||
<mods:start id="4FC5C2F4019F26E8E9FF6DEEBECCBB5E">1</mods:start>
|
||||
<mods:end id="D2CC7425365230300F07084240F66F21">42</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="586F9D65BE495123BA6989059D04FC9A">
|
||||
<mods:url id="723DEF59E42E0ECDFFC37D15146F624A">https://doi.org/10.1093/zoolinnean/zlad143</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="7D5F8C58A58548E6CAAEBFE98837D488">journal article</mods:classification>
|
||||
<mods:identifier id="78534EAF50226EC95DC44CC67C457A2C" type="DOI">10.1093/zoolinnean/zlad143</mods:identifier>
|
||||
<mods:identifier id="3F314EDEA7B511B65AE11B54C76E91A5" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="4DEE7968ED2E57F0B93705BAE411E1BB" type="Zenodo-Dep">14541979</mods:identifier>
|
||||
<mods:identifier id="E455AAFDD7EEED974E0091ACAF5AC669" type="ZooBank">CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03D187F3FFB82E3EFE8E175FFEF9F74C" ID-DOI="http://doi.org/10.5281/zenodo.14541985" ID-Zenodo-Dep="14541985" LSID="urn:lsid:plazi:treatment:03D187F3FFB82E3EFE8E175FFEF9F74C" httpUri="http://treatment.plazi.org/id/03D187F3FFB82E3EFE8E175FFEF9F74C" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
|
||||
<subSubSection id="C362656EFFB82E3DFE8E175FFDA8F7E5" box="[282,593,1659,1686]" pageId="25" pageNumber="26" type="nomenclature">
|
||||
<paragraph id="8BC736E5FFB82E3DFE8E175FFDA8F7E5" blockId="25.[282,593,1659,1686]" box="[282,593,1659,1686]" pageId="25" pageNumber="26">
|
||||
<heading id="D08F8189FFB82E3DFE8E175FFDA8F7E5" bold="true" box="[282,593,1659,1686]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
|
||||
<emphasis id="B90CEAF7FFB82E3DFE8E175FFDA8F7E5" bold="true" box="[282,593,1659,1686]" pageId="25" pageNumber="26">
|
||||
<taxonomicName id="4C784D66FFB82E3DFE8E175FFDF8F7E5" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[282,513,1659,1686]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hammondi" status="sp. nov.">
|
||||
<emphasis id="B90CEAF7FFB82E3DFE8E175FFDF8F7E5" bold="true" box="[282,513,1659,1686]" italics="true" pageId="25" pageNumber="26">Loncovilius hammondi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFFB82E3DFD911758FDA8F7E5" box="[517,593,1660,1686]" pageId="25" pageNumber="26" rank="species">sp.nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFB82E3EFF601789FEF9F74C" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
|
||||
<paragraph id="8BC736E5FFB82E3DFF601789FD81F7B4" blockId="25.[244,632,1709,1736]" box="[244,632,1709,1736]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B90CEAF7FFB82E3DFF601789FD81F7B4" bold="true" box="[244,632,1709,1736]" pageId="25" pageNumber="26">
|
||||
(
|
||||
<figureCitation id="13432A60FFB82E3DFF6B1789FEB5F7BB" box="[255,332,1709,1736]" captionStart="Figure 5" captionStartId="13.[113,178,1868,1892]" captionTargetBox="[154,1419,145,1840]" captionTargetId="figure-6@13.[153,1420,144,1840]" captionTargetPageId="13" captionText="Figure 5. Female habitus of Loncovilius species.A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis." pageId="25" pageNumber="26">Figs 5B</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB82E3DFEC21789FE8EF7B4" box="[342,375,1709,1735]" captionStart="Figure 7" captionStartId="15.[113,178,1868,1892]" captionTargetBox="[153,1419,163,1840]" captionTargetId="figure-6@15.[153,1419,144,1840]" captionTargetPageId="15" captionText="Figure 7. Female habitus of Loncovilius species.A, L. lividipennis. B, L. carlsbergi. Head. C, L. hammondi. D, L. semiflavus. E, L. carlsbergi. F, Male mesotarsi of L. cantharoides." pageId="25" pageNumber="26">7C</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB82E3DFE151789FE34F7BB" box="[385,461,1709,1736]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="25" pageNumber="26">10D–F</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB82E3DFE421789FDFBF7B4" box="[470,514,1709,1735]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="25" pageNumber="26">12B</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB82E3DFD981789FDC1F7B4" box="[524,568,1709,1735]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="25" pageNumber="26">13B</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB82E3DFDD61789FD95F7B4" box="[578,620,1709,1735]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="25" pageNumber="26">14B</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFFE517C5FE26F66B" blockId="25.[113,762,1761,1816]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B90CEAF7FFB82E3DFFE517C5FEAFF78A" box="[113,342,1761,1785]" italics="true" pageId="25" pageNumber="26">Zoobank registration:</emphasis>
|
||||
<uri id="FFE93AE7FFB82E3DFEE417C5FE22F66B" pageId="25" pageNumber="26">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFDE0C30FFB82E3DFDFA17C5FE22F66B" pageId="25" pageNumber="26">4DCCE49B-5875-4A73-9DA6-203BC4455E20</uuid>
|
||||
</uri>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFFE5161FFA8AF353" blockId="25.[113,763,1850,1999]" lastBlockId="25.[809,1460,144,544]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B90CEAF7FFB82E3DFFE5161FFEF0F621" box="[113,265,1850,1874]" italics="true" pageId="25" pageNumber="26">Type material.</emphasis>
|
||||
<materialsCitation id="3B103CB8FFB82E3DFE8C161EFD0BF6E2" collectingDate="1958-12-04" collectorName="Wellington Is. Pto. Eden" country="Chile" location="Fr Kuschel" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="B90CEAF7FFB82E3DFE8C161EFE81F621" box="[280,376,1850,1874]" italics="true" pageId="25" pageNumber="26">
|
||||
<typeStatus id="54C38847FFB82E3DFE8C161EFE8AF621" box="[280,371,1850,1874]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
male, mounted, with genitalia in a separate microvial, with labels as follows:
|
||||
<collectorName id="268D5333FFB82E3DFDF0167DFF27F6E2" pageId="25" pageNumber="26">Wellington Is. Pto. Eden</collectorName>
|
||||
[-49.11, -74.40]
|
||||
<date id="FFC61025FFB82E3DFE31165CFDF6F6E3" box="[421,527,1912,1936]" pageId="25" pageNumber="26" value="1958-12-04">
|
||||
<collectingDate id="EF82E9CDFFB82E3DFE31165CFDF6F6E3" box="[421,527,1912,1936]" pageId="25" pageNumber="26" value="1958-12-04">4.XII1958</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<collectingCountry id="F36F7675FFB82E3DFD83165CFD90F6E3" box="[535,617,1912,1936]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
|
||||
<location id="8EA7603EFFB82E3DFDE3165DFD0BF6E2" LSID="urn:lsid:plazi:treatment:03D187F3FFB82E3EFE8E175FFEF9F74C:8EA7603EFFB82E3DFDE3165DFD0BF6E2" box="[631,754,1913,1937]" country="Chile" name="Fr Kuschel" pageId="25" pageNumber="26">Fr Kuschel</location>
|
||||
</materialsCitation>
|
||||
/
|
||||
<typeStatus id="54C38847FFB82E3DFFE516BCFF05F6C3" box="[113,252,1944,1968]" pageId="25" pageNumber="26" type="holotype">HOLOTYPE</typeStatus>
|
||||
<taxonomicName id="4C784D66FFB82E3DFE9916BCFE0FF6C3" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[269,502,1944,1968]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hammondi">
|
||||
<emphasis id="B90CEAF7FFB82E3DFE9916BCFE0FF6C3" box="[269,502,1944,1968]" italics="true" pageId="25" pageNumber="26">Loncovilius hammondi</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at NHM.
|
||||
<materialsCitation id="3B103CB8FFB82E3DFCE811B5FA96F353" collectingDate="1958-11-10" collectingDateMax="1982-12-29" collectingDateMin="1958-11-10" collectorName="Wellington Is. Pto. Eden & Chiloe S. Pedro & Fr Kuschel & Qued. & Loncovilius' & Cautin Pr. Volcan Villarrica" country="Chile" county="Flowers of Perneuya" elevation="1120" latitude="-49.0" location="Puerto Eden" municipality="Wellington" pageId="25" pageNumber="26" specimenCount="18" specimenCount-female="10" specimenCount-male="8" typeStatus="paratype">
|
||||
<emphasis id="B90CEAF7FFB82E3DFCE811B5FC1AF1DB" box="[892,995,145,168]" italics="true" pageId="25" pageNumber="26">
|
||||
<typeStatus id="54C38847FFB82E3DFCE811B5FC27F1DB" box="[892,990,145,168]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
<collectorName id="268D5333FFB82E3DFC6411B4FB04F1DB" box="[1008,1277,144,168]" pageId="25" pageNumber="26">Wellington Is. Pto. Eden</collectorName>
|
||||
[-49.11, -74.40]
|
||||
<date id="FFC61025FFB82E3DFCBE118BFC60F1B4" box="[810,921,175,199]" pageId="25" pageNumber="26" value="1958-12-04">
|
||||
<collectingDate id="EF82E9CDFFB82E3DFCBE118BFC60F1B4" box="[810,921,175,199]" pageId="25" pageNumber="26" value="1958-12-04">4.XII.1958</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<collectingCountry id="F36F7675FFB82E3DFC35118BFC0DF1B4" box="[929,1012,175,199]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
|
||||
Fr Kuschel [
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFBEA118BFB51F1B5" box="[1150,1192,175,199]" count="4" pageId="25" pageNumber="26" type="male">4♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFB24118BFB2EF1B5" box="[1200,1239,175,199]" count="8" pageId="25" pageNumber="26" type="female">8♀</specimenCount>
|
||||
NHM;
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFAA4118BFAAEF1B5" box="[1328,1367,175,199]" count="1" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
|
||||
NHMD ex. NHM;
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFC0B11EBFC3FF195" box="[927,966,207,231]" count="1" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
|
||||
CZUG ex. NHM];
|
||||
<taxonomicName id="4C784D66FFB82E3DFB0A11EBFAEEF194" box="[1182,1303,207,231]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
|
||||
forest/
|
||||
<collectingCountry id="F36F7675FFB82E3DFAF111EBFA4AF194" box="[1381,1459,207,231]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
|
||||
HE6.
|
||||
<location id="8EA7603EFFB82E3DFCCA11CAFC1CF075" LSID="urn:lsid:plazi:treatment:03D187F3FFB82E3EFE8E175FFEF9F74C:8EA7603EFFB82E3DFCCA11CAFC1CF075" box="[862,997,238,262]" country="Chile" county="Flowers of Perneuya" latitude="-49.0" municipality="Wellington" name="Puerto Eden" pageId="25" pageNumber="26">Puerto Eden</location>
|
||||
Isla
|
||||
<collectingMunicipality id="6BA3AC9FFFB82E3DFB8D11CAFB72F075" box="[1049,1163,238,262]" pageId="25" pageNumber="26">Wellington</collectingMunicipality>
|
||||
[-49.11, -74.40]
|
||||
<geoCoordinate id="EE4C5022FFB82E3DFAD611CAFA83F075" box="[1346,1402,238,262]" degrees="49" direction="south" orientation="latitude" pageId="25" pageNumber="26" precision="55555" value="-49.0">49° S</geoCoordinate>
|
||||
I200 ss.
|
||||
<date id="FFC61025FFB82E3DFCD11029FC53F056" box="[837,938,269,293]" pageId="25" pageNumber="26" value="1958-12-02">
|
||||
<collectingDate id="EF82E9CDFFB82E3DFCD11029FC53F056" box="[837,938,269,293]" pageId="25" pageNumber="26" value="1958-12-02">2.xii.1958</collectingDate>
|
||||
</date>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFC2F1029FC1BF057" box="[955,994,269,293]" count="1" pageId="25" pageNumber="26" type="female">1♀</specimenCount>
|
||||
MNNC ex. NHM];
|
||||
<collectorName id="268D5333FFB82E3DFB2E1029FAA5F056" box="[1210,1372,269,293]" pageId="25" pageNumber="26">Chiloé S. Pedro</collectorName>
|
||||
[-43.32, -73.72]
|
||||
<date id="FFC61025FFB82E3DFC151008FC0AF037" box="[897,1011,300,324]" pageId="25" pageNumber="26" value="1958-11-10">
|
||||
<collectingDate id="EF82E9CDFFB82E3DFC151008FC0AF037" box="[897,1011,300,324]" pageId="25" pageNumber="26" value="1958-11-10">10.XI.1958</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<collectingCountry id="F36F7675FFB82E3DFC6F1008FBB6F037" box="[1019,1103,300,324]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
|
||||
<collectorName id="268D5333FFB82E3DFBCD1009FB36F036" box="[1113,1231,301,325]" pageId="25" pageNumber="26">Fr Kuschel</collectorName>
|
||||
/
|
||||
<collectingCounty id="62A64E69FFB82E3DFB431009FA4DF036" box="[1239,1460,301,325]" pageId="25" pageNumber="26">Flowers of Perneưya</collectingCounty>
|
||||
mucronata/
|
||||
<collectorName id="268D5333FFB82E3DFC331068FC1FF017" box="[935,998,332,356]" pageId="25" pageNumber="26">Qued.</collectorName>
|
||||
sp. 8/‘
|
||||
<collectorName id="268D5333FFB82E3DFBB71068FB53F017" box="[1059,1194,332,356]" pageId="25" pageNumber="26">Loncovilius’</collectorName>
|
||||
<taxonomicName id="4C784D66FFB82E3DFB3B1068FC65F0F0" authority="Scheerp. P. M. Hammond" class="Insecta" family="Staphylinidae" genus="Loncovilius" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">cf germaini Scheerp. P.M. Hammond</taxonomicName>
|
||||
det. 1980 [
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFB82104FFBC7F0F1" box="[1046,1086,363,387]" count="1" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
|
||||
NHM];
|
||||
<collectingCountry id="F36F7675FFB82E3DFB0A104FFB08F0F0" box="[1182,1265,363,387]" name="Chile" pageId="25" pageNumber="26">CHILE</collectingCountry>
|
||||
:
|
||||
<collectorName id="268D5333FFB82E3DFB68104FFC71F0D0" pageId="25" pageNumber="26">Cautín Pr. Volcán Villarrica</collectorName>
|
||||
[-39.36, -71.94],
|
||||
<quantity id="4C809B00FFB82E3DFBAC10AFFB7BF0D0" box="[1080,1154,395,419]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="25" pageNumber="26" unit="m" value="1120.0">
|
||||
<elevation id="0055D1D6FFB82E3DFBAC10AFFB7BF0D0" box="[1080,1154,395,419]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="25" pageNumber="26" unit="m" value="1120.0">1120m</elevation>
|
||||
</quantity>
|
||||
, site 654
|
||||
<date id="FFC61025FFB82E3DFB4A10AEFA8BF0D1" box="[1246,1394,394,418]" pageId="25" pageNumber="26" value="1982-12-15" valueMax="1982-12-29" valueMin="1982-12-15">
|
||||
<collectingDate id="EF82E9CDFFB82E3DFB4A10AEFA8BF0D1" box="[1246,1394,394,418]" pageId="25" pageNumber="26" value="1982-12-15" valueMax="1982-12-29" valueMin="1982-12-15">15-29.xii.1982</collectingDate>
|
||||
</date>
|
||||
Noth. Dombeyi-Saxe-gothea with
|
||||
<taxonomicName id="4C784D66FFB82E3DFBDA108EFA50F0B1" authority="A. Newton and M. Nayer" authorityName="A. Newton and M. Nayer" box="[1102,1449,426,450]" class="Magnoliopsida" family="Winteraceae" genus="Drimys" kingdom="Plantae" order="Canellales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">Drimys A.Newton and M.Ŋayer</taxonomicName>
|
||||
/
|
||||
<collectingMethod id="52394EF2FFB82E3DFCBE10EDFC4BF092" box="[810,946,457,481]" pageId="25" pageNumber="26">window trap</collectingMethod>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFC5C10EDFC16F093" box="[968,1007,457,481]" count="1" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFC6F10EDFBDBF093" box="[1019,1058,457,481]" count="1" pageId="25" pageNumber="26" type="female">1♀</specimenCount>
|
||||
FMNH]. All
|
||||
<typeStatus id="54C38847FFB82E3DFB5610EEFADCF091" box="[1218,1317,458,482]" pageId="25" pageNumber="26" type="paratype">paratypes</typeStatus>
|
||||
are supplied with the label: PAÞTYPE
|
||||
<taxonomicName id="4C784D66FFB82E3DFBC710CDFACDF372" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[1107,1332,489,513]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hammondi">
|
||||
<emphasis id="B90CEAF7FFB82E3DFBC710CDFACDF372" box="[1107,1332,489,513]" italics="true" pageId="25" pageNumber="26">Loncovilius hammondi</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFCBE1366FA8CF247" blockId="25.[810,1460,577,821]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B90CEAF7FFB82E3DFCBE1366FC69F32A" box="[810,912,578,601]" italics="true" pageId="25" pageNumber="26">Diagnosis:</emphasis>
|
||||
Head dorsally with microsculpture of transverse waves; gula with isodiametric microsculpture in the middle; females with a11 distinctly longer than a10 (a11:a10 ratio>1.5). Elytra reddish-brown, disc with coarse non-setiferous punctures and only a few rows of setiferous punctures, epipleura evenly setose. Protibiae not sexually dimorphic, in both sexes with laterodorsal row of thick spines. Tergite VIII with medial apical emargination in both sexes. Aedeagus as in
|
||||
<figureCitation id="13432A60FFB82E3DFB741239FA96F246" box="[1248,1391,797,821]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="25" pageNumber="26">Figure 10D–F</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFCBE1273FB67F578" blockId="25.[809,1461,854,2006]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B90CEAF7FFB82E3DFCBE1273FC58F21D" box="[810,929,855,878]" italics="true" pageId="25" pageNumber="26">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFBC5127DFB92F21E" box="[1105,1131,857,877]" count="1" pageId="25" pageNumber="26" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFB82E3DFAF01273FA8EF21D" box="[1380,1399,855,878]" italics="true" pageId="25" pageNumber="26">N</emphasis>
|
||||
= 6]: FBL = 3.5–3.91 (3.67); TL = 6.7–7.31 (7.01); HW = 1.06– 1.16 (1.11); HL = 0.94–1.02 (0.97); HW/HL = 1.12–1.16 (1.14); PW = 1.36–1.51 (1.43); PL = 1.16–1.26 (1.21); PW/ PL = 1.17–1.2 (1.19); EW = 1.65–1.82 (1.76); EL = 1.38–1.63 (1.49); PW/HW = 1.29–1.3 (1.29).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFCD11536FB67F5B4" blockId="25.[809,1461,854,2006]" pageId="25" pageNumber="26">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFB82E3DFC751531FC02F55A" box="[993,1019,1045,1065]" count="1" pageId="25" pageNumber="26" type="female">♀</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFB82E3DFB4F1537FB17F559" box="[1243,1262,1043,1066]" italics="true" pageId="25" pageNumber="26">N</emphasis>
|
||||
= 6]: FBL = 3.88– 4.22 (4.04); TL = 7.06–8.88 (7.81); HW = 1.13–1.21 (1.18); HL = 0.98–1.07 (1.03); HW/HL = 1.13–1.19 (1.15); PW = 1.5–1.67 (1.58); PL = 1.23–1.37 (1.31); PW/ PL = 1.18–1.22 (1.20); EW = 1.9–2.3 (2.05); EL = 1.67–1.78 (1.7); PW/HW = 1.31–1.38 (1.33).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFCD115EAFC2FF437" blockId="25.[809,1461,854,2006]" pageId="25" pageNumber="26">Head black to brown-black; most of antennae, pronotum, mouthparts, elytra, legs, and abdomen reddish-brown; pronotum, apical antennomeres, tibiae, and disc of abdominal segments darker.</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3EFCD1146FFF0DF056" blockId="25.[809,1461,854,2006]" lastBlockId="26.[127,779,144,1171]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
|
||||
Head transverse; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse waves, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes small (EYL/HL = x = 0.47), from 1.3 to 1.8 times longer than temples (in lateral view); distance between eyes about 1.7 times as long as length of eye in males and 1.68 times in females. Antennal insertions situated at equal distance from frontoclypeus and from eye; antennomeres 2 and 3 usually subequal in length; antennomeres 4 to 6 gradually reduced in length; antennomeres 7 to 10 subequal in length; antennomere 11 is from 1.53 to 1.69 times as long as antennomere
|
||||
<quantity id="4C809B00FFB82E3DFBC91780FB69F7CF" box="[1117,1168,1700,1724]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="10.0">10 in</quantity>
|
||||
males and from 1.63 to 1.71 times in females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; without small setiferous punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and emarginate apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than previous and more or less subconical.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFF081009FCF9F372" blockId="26.[127,779,144,1171]" pageId="26" pageNumber="27">Pronotum transverse, convex, evenly curved, with one PPDS and with an APP, without SLSP, with several coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra with epipleura evenly setiferous punctate; disc with only a few rows of setiferous punctures and with several coarse non-setiferous punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFF08132CFF36F388" blockId="26.[127,779,144,1171]" pageId="26" pageNumber="27">Protibiae not sexually dimorphic, with laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin slightly sinuate.</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFF081227FEA2F500" blockId="26.[127,779,144,1171]" pageId="26" pageNumber="27">
|
||||
Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line; tergite VIII emarginate medio-apically. Sternum III with basal transverse carina strongly arcuate; lateral tergal sclerites IX laterally flaưened only in males. Male sternite VIII with a broad V-shaped emargination medially (
|
||||
<figureCitation id="13432A60FFBB2E3EFDD212BBFD5AF2C4" box="[582,675,927,951]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="26" pageNumber="27">Fig. 14B</figureCitation>
|
||||
); female sternite VIII with a U-shaped emargination medially; male sternite IX with an obtuse emargination medially, its basal portion shorter than distal portion; male tergite X truncate medially (
|
||||
<figureCitation id="13432A60FFBB2E3EFF2E1539FEEAF546" box="[186,275,1053,1077]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="26" pageNumber="27">Fig. 12B</figureCitation>
|
||||
); female tergite X truncate to slightly arcuate medially (
|
||||
<figureCitation id="13432A60FFBB2E3EFF7F1518FEB8F527" box="[235,321,1084,1108]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="26" pageNumber="27">Fig. 13B</figureCitation>
|
||||
); ovipositor, its second gonocoxite without macrosetae medially.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFF08155FFD41F5E0" blockId="26.[127,779,144,1171]" box="[156,696,1147,1171]" pageId="26" pageNumber="27">
|
||||
Aedeagus. As in
|
||||
<figureCitation id="13432A60FFBB2E3EFED6155FFE2AF5E0" box="[322,467,1147,1171]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="26" pageNumber="27">Figure 10D–F</figureCitation>
|
||||
; its total length ~1.21.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFF151590FDAFF459" blockId="26.[128,778,1204,1322]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFF151590FF09F5BF" box="[129,240,1204,1228]" italics="true" pageId="26" pageNumber="27">Etymology:</emphasis>
|
||||
Ŋe species epithet isa patronym in recognition of the late Peter. M. Hammond, a prominent coleopterist who also conducted some work on this genus, which was unpublished but which we were able to trace through his labels.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFF15146FFEF9F74C" blockId="26.[129,777,1355,1599]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFF15146FFD69F410" box="[129,656,1355,1379]" italics="true" pageId="26" pageNumber="27">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FFBB2E3EFE37146FFD69F410" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[419,656,1355,1379]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hammondi">Loncovilius hammondi</taxonomicName>
|
||||
</emphasis>
|
||||
is known only from
|
||||
<collectingCountry id="F36F7675FFBB2E3EFF62144FFEC8F4F0" box="[246,305,1387,1411]" name="Chile" pageId="26" pageNumber="27">Chile</collectingCountry>
|
||||
, from the
|
||||
<collectingRegion id="49BCF807FFBB2E3EFE3C144FFDEDF4F0" box="[424,532,1387,1411]" country="Chile" name="Araucania" pageId="26" pageNumber="27">Araucanía</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBB2E3EFDB6144FFD6AF4F0" box="[546,659,1387,1411]" country="Chile" name="Los Lagos" pageId="26" pageNumber="27">Los Lagos</collectingRegion>
|
||||
, as well as from
|
||||
<collectingRegion id="49BCF807FFBB2E3EFF5714AEFEC1F4D1" box="[195,312,1418,1442]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="26" pageNumber="27">Magallanes</collectingRegion>
|
||||
and Chilean
|
||||
<collectingCountry id="F36F7675FFBB2E3EFE4B14AEFDB2F4D1" box="[479,587,1418,1442]" name="Antarctica" pageId="26" pageNumber="27">Antarctica</collectingCountry>
|
||||
regions. Known distribution consists of three widely separated areas, two of them confined to the Valdivian temperate forest and one to the Magellanic subpolar forest. It has been collected using window traps, and directly from flowers of
|
||||
<taxonomicName id="4C784D66FFBB2E3EFDAF172CFCF0F76C" box="[571,777,1544,1567]" class="Magnoliopsida" family="Campanulaceae" genus="Pernetya" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Asterales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="mucronata">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFDAF172CFCF0F76C" box="[571,777,1544,1567]" italics="true" pageId="26" pageNumber="27">Pernetya mucronata</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C784D66FFBB2E3EFF1C1703FF0CF74C" box="[136,245,1575,1599]" class="Magnoliopsida" family="Ericaceae" kingdom="Plantae" order="Ericales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="family">Ericaceae</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,126 +0,0 @@
|
|||
<document id="AACFB4F729CF0E3A2A4685756DF00BDF" ID-DOI="10.1093/zoolinnean/zlad143" ID-ISSN="0024-4082" ID-Zenodo-Dep="14541979" ID-ZooBank="CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1732839935034" checkinUser="plazi" docAuthor="Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins & Solodovnikov, Alexey" docDate="2024" docId="03D187F3FFB92E3CFEE21311FD2DF2AF" docLanguage="en" docName="zlad143.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlad143" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Loncovilius germaini" docType="treatment" docVersion="3" lastPageNumber="25" masterDocId="FFE8FF8BFFA12E24FF941124FFF9F173" masterDocTitle="Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="25" updateTime="1734984611405" updateUser="julia">
|
||||
<mods:mods id="78FFFFD5C95A1605FED483F9792754ED" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="DFBBC46CD310A871DE40EC387C7088CA">
|
||||
<mods:title id="C94B8D544F6CA1F1D3E66F7A5DC540FC">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="2D0F57C63D1D8A6DDDF740458EEF2FD7" type="personal">
|
||||
<mods:role id="6AA8EBFF0E3AA3A57C3C39E8FF8598B0">
|
||||
<mods:roleTerm id="509B7019BC1081A022B202F3E8AF2758">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="D33A21205292C6F2EDD1C91121ED4461">Reyes-Hernández, José L.</mods:namePart>
|
||||
<mods:affiliation id="9CAE0D7132AA40131E8CD7DDB03278E9">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
|
||||
<mods:nameIdentifier id="0B5E42AF30E821F7D45A17B44AFE51DC" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name id="E82920CD245AA1A933EA7495A54F77EF" type="personal">
|
||||
<mods:role id="1A1B6B0C6F8973A9A93A488E629D5ECC">
|
||||
<mods:roleTerm id="E1943ED88BDF03313C3B2850C2280326">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="B1184FF2A5F45AB7FF45DBDF3559ACB5">Hansen, Aslak Kappel</mods:namePart>
|
||||
<mods:affiliation id="8E2FA850E792C84E76F5E08148018E4B">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="0114B195225741B67B418C5E7B9D6B85" type="personal">
|
||||
<mods:role id="FE157D202C586D3879DCCF6CF90820D1">
|
||||
<mods:roleTerm id="DFE716FA662DFDAF322E98A7463A07B7">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
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group
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<taxonomicName id="4C784D66FFB92E3CFEA71351FD1AF3FD" authority="(Scheerpeltz, 1933)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[307,739,629,654]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini">
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<emphasis id="B90CEAF7FFB92E3CFEA71351FDFDF3FE" box="[307,516,629,653]" italics="true" pageId="24" pageNumber="25">Loncovilius germaini</emphasis>
|
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(
|
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<bibRefCitation id="EFE94B14FFB92E3CFD8E1351FD21F3FD" author="Scheerpeltz O" box="[538,728,629,654]" pageId="24" pageNumber="25" pagination="989 - 1500" refId="ref38278" refString="Scheerpeltz O. Staphylinidae VII. In: Schenkling S (ed.), Coleopterorum Catalogus, Vol. 6. Berlin: Junk, 1933, 989 - 1500." type="book chapter" year="1933">Scheerpeltz, 1933</bibRefCitation>
|
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)
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</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C784D66FFB92E3CFD611352FF12F3DE" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="hammondi" status="sp. nov.">
|
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<emphasis id="B90CEAF7FFB92E3CFD611352FCFDF3FE" box="[757,772,630,653]" italics="true" pageId="24" pageNumber="25">L</emphasis>
|
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.
|
||||
<emphasis id="B90CEAF7FFB92E3CFF1513B1FF12F3DE" box="[129,235,661,685]" italics="true" pageId="24" pageNumber="25">hammondi</emphasis>
|
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|
||||
<taxonomicNameLabel id="A23F578CFFB92E3CFF7B13B1FEC4F3DF" box="[239,317,661,684]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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, and
|
||||
<taxonomicName id="4C784D66FFB92E3CFEE413B2FE16F3DE" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[368,495,661,685]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="impunctus" status="sp. nov.">
|
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<emphasis id="B90CEAF7FFB92E3CFEE413B2FE86F3DE" box="[368,383,662,685]" italics="true" pageId="24" pageNumber="25">L</emphasis>
|
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.
|
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<emphasis id="B90CEAF7FFB92E3CFE1E13B2FE16F3DE" box="[394,495,662,685]" italics="true" pageId="24" pageNumber="25">impunctus</emphasis>
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<taxonomicNameLabel id="A23F578CFFB92E3CFE6013B1FDB9F3DF" box="[500,576,661,684]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="B90CEAF7FFB92E3CFF1513EEFF1DF392" box="[129,228,714,737]" italics="true" pageId="24" pageNumber="25">Diagnosis:</emphasis>
|
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Head transverse, with coarse non-setiferous punctures; nuchal ridge complete dorsally, nuchal and infraorbital ridges joined; postmandibular ridge distinctly separated from eye margin and well-developed PMP (
|
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<figureCitation id="13432A60FFB92E3CFE1C1203FE34F233" box="[392,461,807,832]" captionStart="Figure 8" captionStartId="16.[129,194,1829,1853]" captionTargetBox="[183,1419,150,1799]" captionTargetId="figure-8@16.[183,1419,144,1799]" captionTargetPageId="16" captionText="Figure 8. A, pronotum of L. carlsbergi. AMRSP = anterior marginal row of setiferous punctures, APP = additional paired punctures adjacent to AMRSP, LLSP = large lateral setiferous puncture, LPSP = large posterior setiferous puncture, PPDS = paired punctures on dorsal serial, SLSP = sublateral setiferous puncture.B, L. cantharoides ventral view of mesotarsomere 3 with pale adhesive setae with terminal plate.C, L. cantharoides ventral view of metatarsomere 4, apical margin emarginated (bilobed), with pale adhesive setae without terminal plate. D, L. cantharoides tergites VI and VII.Black arrow up = anterior transverse basal carina (ATBC) laterally straight continuing to paratergites, white arrow down = ATBC descending laterally without reaching tergite margin.Head in ventral view of Loncovilius species. E, L. hammondi. F, L. lividipennis. PMP = postmandibular puncture, PMR = postmandibular ridge, EM = eye margin." pageId="24" pageNumber="25">Fig. 8E</figureCitation>
|
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;
|
||||
<bibRefCitation id="EFE94B14FFB92E3CFE4D1203FD70F233" author="Brunke AJ & Zyla D & Yamamoto S" box="[473,649,807,832]" pageId="24" pageNumber="25" pagination="166 - 97" refId="ref36256" refString="Brunke AJ, Zyla D, Yamamoto S et al. Baltic amber Staphylinini (Coleoptera: Staphylinidae: Staphylininae): a rove beetle fauna on the eve of our modern climate. Zoological Journal of the Linnean Society 2019; 187: 166 - 97." type="journal article" year="2019">
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Brunke
|
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<emphasis id="B90CEAF7FFB92E3CFDB2120CFDB4F24C" box="[550,589,807,831]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
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. 2019
|
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</bibRefCitation>
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: fig. 1); right mandible with distal tooth bifid; left mandible only with proximal, non-bifid tooth. Pronotum transverse, outline evenly curved. Tergites III and IV with two patches of punctures of moderate size and density, with a middle impunctate line (
|
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<figureCitation id="13432A60FFB92E3CFDAA1281FD7EF2CE" box="[574,647,933,957]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="24" pageNumber="25">Fig. 9A</figureCitation>
|
||||
); ovipositor, its second gonocoxite without macroseta medially (
|
||||
<figureCitation id="13432A60FFB92E3CFDEB12E0FD3DF2AF" box="[639,708,964,988]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="24" pageNumber="25">Fig. 9F</figureCitation>
|
||||
).
|
||||
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|
||||
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|
||||
<mods:classification id="20357BA40C1881FBB5F072C087D5B616">journal article</mods:classification>
|
||||
<mods:identifier id="7FDC526E6FCBCE92199F4AD14C4252B9" type="DOI">10.1093/zoolinnean/zlad143</mods:identifier>
|
||||
<mods:identifier id="0D28D596D0AD887DA364CD66EA18B0ED" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="6FF410CF36FB3521B8BEB0596B45731B" type="Zenodo-Dep">14541979</mods:identifier>
|
||||
<mods:identifier id="AEE2C2C9C5457368812A15CBF1955210" type="ZooBank">CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03D187F3FFB92E3DFF6412DFFF2FF72D" LSID="urn:lsid:plazi:treatment:03D187F3FFB92E3DFF6412DFFF2FF72D" httpUri="http://treatment.plazi.org/id/03D187F3FFB92E3DFF6412DFFF2FF72D" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||||
<subSubSection id="C362656EFFB92E3CFF6412DFFD63F566" box="[240,666,1019,1046]" pageId="24" pageNumber="25" type="nomenclature">
|
||||
<paragraph id="8BC736E5FFB92E3CFF6412DFFD63F566" blockId="24.[240,666,1019,1046]" box="[240,666,1019,1046]" pageId="24" pageNumber="25">
|
||||
<heading id="D08F8189FFB92E3CFF6412DFFD63F566" bold="true" box="[240,666,1019,1046]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
|
||||
<taxonomicName id="4C784D66FFB92E3CFF6412DFFD63F566" ID-CoL="3VYHQ" authority="(Scheerpeltz, 1933)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[240,666,1019,1046]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini">
|
||||
<emphasis id="B90CEAF7FFB92E3CFF6412DFFD63F566" bold="true" box="[240,666,1019,1046]" pageId="24" pageNumber="25">
|
||||
<emphasis id="B90CEAF7FFB92E3CFF6412DFFE3BF565" bold="true" box="[240,450,1019,1046]" italics="true" pageId="24" pageNumber="25">Loncovilius germaini</emphasis>
|
||||
(Scheerpeltz, 1933)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFB92E3DFE94150FFF2FF72D" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
|
||||
<paragraph id="8BC736E5FFB92E3CFE94150FFD73F536" blockId="24.[256,650,1067,1093]" box="[256,650,1067,1093]" pageId="24" pageNumber="25">
|
||||
<emphasis id="B90CEAF7FFB92E3CFE94150FFD73F536" bold="true" box="[256,650,1067,1093]" pageId="24" pageNumber="25">
|
||||
(
|
||||
<figureCitation id="13432A60FFB92E3CFE9E150FFEA3F536" box="[266,346,1067,1093]" captionStart="Figure 5" captionStartId="13.[113,178,1868,1892]" captionTargetBox="[154,1419,145,1840]" captionTargetId="figure-6@13.[153,1420,144,1840]" captionTargetPageId="13" captionText="Figure 5. Female habitus of Loncovilius species.A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis." pageId="24" pageNumber="25">Figs 5A</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB92E3CFEF0150FFE7AF536" box="[356,387,1067,1093]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="24" pageNumber="25">9A</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB92E3CFE19150FFE22F536" box="[397,475,1067,1093]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="24" pageNumber="25">10A–C</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB92E3CFE70150FFDEBF536" box="[484,530,1067,1093]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="24" pageNumber="25">12A</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB92E3CFD88150FFDB3F536" box="[540,586,1067,1093]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="24" pageNumber="25">13A</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFB92E3CFDC7150FFD86F536" box="[595,639,1067,1093]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="24" pageNumber="25">14A</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFF151545FD36F4E0" blockId="24.[129,779,1121,1428]" pageId="24" pageNumber="25">
|
||||
Germain 1903: 412 (as
|
||||
<taxonomicName id="4C784D66FFB92E3CFEE31545FDAFF50A" box="[375,598,1121,1145]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cribripennis">
|
||||
<emphasis id="B90CEAF7FFB92E3CFEE31545FDAFF50A" box="[375,598,1121,1145]" italics="true" pageId="24" pageNumber="25">Philonthus cribripennis</emphasis>
|
||||
</taxonomicName>
|
||||
, original description); Bernhauer and Schubert 1914: 332 (as
|
||||
<taxonomicName id="4C784D66FFB92E3CFDD315A5FCFAF5EA" authorityName="Bernhauer & K. Schubert" authorityYear="1914" box="[583,771,1153,1177]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chilensis">
|
||||
<emphasis id="B90CEAF7FFB92E3CFDD315A5FCFAF5EA" box="[583,771,1153,1177]" italics="true" pageId="24" pageNumber="25">Philonthus chilensis</emphasis>
|
||||
</taxonomicName>
|
||||
, replacement name); Scheerpeltz 1933: 1344 (as
|
||||
<taxonomicName id="4C784D66FFB92E3CFD351584FF2EF5A4" authorityName="Scheerpeltz" authorityYear="1933" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini">
|
||||
<emphasis id="B90CEAF7FFB92E3CFD351584FF2EF5A4" italics="true" pageId="24" pageNumber="25">Philonthus germaini</emphasis>
|
||||
</taxonomicName>
|
||||
, replacement name); Coiffait and Sáiz 1966: 406 (as
|
||||
<taxonomicName id="4C784D66FFB92E3CFF1515FBFE37F584" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[129,462,1247,1271]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini" subGenus="Lienturius">
|
||||
<emphasis id="B90CEAF7FFB92E3CFF1515FBFF16F584" box="[129,239,1247,1271]" italics="true" pageId="24" pageNumber="25">Loncovilius</emphasis>
|
||||
(
|
||||
<emphasis id="B90CEAF7FFB92E3CFE9515C4FE9CF584" box="[257,357,1248,1271]" italics="true" pageId="24" pageNumber="25">Lienturius</emphasis>
|
||||
)
|
||||
<emphasis id="B90CEAF7FFB92E3CFEEC15C4FE37F584" box="[376,462,1248,1271]" italics="true" pageId="24" pageNumber="25">germaini</emphasis>
|
||||
</taxonomicName>
|
||||
, characters); Coiffait and Sáiz 1968: 365 [as
|
||||
<taxonomicName id="4C784D66FFB92E3CFE8315DAFD98F465" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[279,609,1278,1302]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini" subGenus="Lienturius">
|
||||
<emphasis id="B90CEAF7FFB92E3CFE8315DAFE7CF465" box="[279,389,1278,1302]" italics="true" pageId="24" pageNumber="25">Loncovilius</emphasis>
|
||||
(
|
||||
<emphasis id="B90CEAF7FFB92E3CFE0115DBFE00F465" box="[405,505,1279,1302]" italics="true" pageId="24" pageNumber="25">Lienturius</emphasis>
|
||||
)
|
||||
<emphasis id="B90CEAF7FFB92E3CFD9E15DBFD98F465" box="[522,609,1279,1302]" italics="true" pageId="24" pageNumber="25">germaini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90CEAF7FFB92E3CFDF615DBFD91F465" box="[610,616,1279,1302]" italics="true" pageId="24" pageNumber="25">;</emphasis>
|
||||
checklist]; Sáiz 1971: 385 [as
|
||||
<taxonomicName id="4C784D66FFB92E3CFE8F1439FD90F446" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[283,617,1309,1334]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini" subGenus="Lienturius">
|
||||
<emphasis id="B90CEAF7FFB92E3CFE8F1439FE70F446" box="[283,393,1309,1333]" italics="true" pageId="24" pageNumber="25">Loncovilius</emphasis>
|
||||
(
|
||||
<emphasis id="B90CEAF7FFB92E3CFE08143AFDF9F446" box="[412,512,1310,1333]" italics="true" pageId="24" pageNumber="25">Lienturius</emphasis>
|
||||
)
|
||||
<emphasis id="B90CEAF7FFB92E3CFD87143AFD90F446" box="[531,617,1310,1333]" italics="true" pageId="24" pageNumber="25">germaini</emphasis>
|
||||
</taxonomicName>
|
||||
; listed without new data]; Herman 2001a: 26 (as
|
||||
<taxonomicName id="4C784D66FFB92E3CFE451419FD6BF426" authorityName="Bernhauer and Schubert" authorityYear="1914" box="[465,658,1341,1365]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chilensis">
|
||||
<emphasis id="B90CEAF7FFB92E3CFE451419FD6BF426" box="[465,658,1341,1365]" italics="true" pageId="24" pageNumber="25">Loncovilius chilensis</emphasis>
|
||||
</taxonomicName>
|
||||
, nomenclature); Herman 2001b: 3083 (as
|
||||
<taxonomicName id="4C784D66FFB92E3CFE5D1478FD72F407" authorityName="Bernhauer and Schubert" authorityYear="1914" box="[457,651,1372,1396]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chilensis">
|
||||
<emphasis id="B90CEAF7FFB92E3CFE5D1478FD72F407" box="[457,651,1372,1396]" italics="true" pageId="24" pageNumber="25">Loncovilius chilensis</emphasis>
|
||||
</taxonomicName>
|
||||
, catalogue); Newton 2017: 22 (as
|
||||
<taxonomicName id="4C784D66FFB92E3CFEC9145FFDDFF4E0" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[349,550,1403,1427]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="germaini">
|
||||
<emphasis id="B90CEAF7FFB92E3CFEC9145FFDDFF4E0" box="[349,550,1403,1427]" italics="true" pageId="24" pageNumber="25">Loncovilius germaini</emphasis>
|
||||
</taxonomicName>
|
||||
, nomenclature).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFF151494FD6BF4BB" blockId="24.[129,658,1456,1480]" box="[129,658,1456,1480]" pageId="24" pageNumber="25">
|
||||
<emphasis id="B90CEAF7FFB92E3CFF151494FEB9F4BB" box="[129,320,1456,1480]" italics="true" pageId="24" pageNumber="25">Material eoamined:</emphasis>
|
||||
Supporting Information, File S7.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFF1514FAFCFDF7C3" blockId="24.[128,778,1501,1713]" pageId="24" pageNumber="25">
|
||||
<emphasis id="B90CEAF7FFB92E3CFF1514FAFF1CF486" box="[129,229,1502,1525]" italics="true" pageId="24" pageNumber="25">Diagnosis:</emphasis>
|
||||
Head dorsally without microsculpture; gula without isodiametric microsculpture in the middle; females with a11 distinctly longer than a10 (a11:a10 ratio <1.5). Elytra, including epipleura, evenly setose and with several coarse non-setiferous punctures. Protibiae sexually dimorphic, with laterodorsal row of thick spines only present in females. Tergite VIII with medial apical emargination only in females. Aedeagus as in
|
||||
<figureCitation id="13432A60FFB92E3CFDF917BDFD07F7C2" box="[621,766,1689,1713]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="24" pageNumber="25">Figure 10A–C</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFF1517EDFE0CF60E" blockId="24.[129,779,1736,1980]" pageId="24" pageNumber="25">
|
||||
<emphasis id="B90CEAF7FFB92E3CFF1517EDFF0EF793" box="[129,247,1737,1760]" italics="true" pageId="24" pageNumber="25">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFB92E3CFE3C17EFFE3BF7AC" box="[424,450,1739,1759]" count="1" pageId="24" pageNumber="25" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFB92E3CFD2E17EDFD34F793" box="[698,717,1737,1760]" italics="true" pageId="24" pageNumber="25">N</emphasis>
|
||||
= 6]: FBL = 2.9–3.44 (3.19); TL = 5.6–6.25 (6.01); HW = 0.87– 1.01 (0.94); HL = 0.76–0.86 (0.82); HW/HL = 1.12–1.2 (1.14); PW = 1.11–1.29 (1.21); PL = 0.97–1.18 (1.08); PW/ PL = 1.1–1.14 (1.12); EW = 1.4–1.64 (1.52); EL = 1.16–1.4 (1.3); PW/HW = 1.26–1.33 (1.29).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFF0816A1FBA7F3DE" blockId="24.[129,779,1736,1980]" lastBlockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFB92E3CFEA316A3FEA8F6E8" box="[311,337,1927,1947]" count="1" pageId="24" pageNumber="25" type="female">♀</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFB92E3CFDA516A1FDBDF6EF" box="[561,580,1925,1948]" italics="true" pageId="24" pageNumber="25">N</emphasis>
|
||||
= 6]: FBL = 3.36– 3.87 (3.6); TL = 7.31–7.77 (7.54); HW = 0.96–1.12 (1.03); HL = 0.85–0.95 (0.91); HW/HL = 1.11–1.18 (1.13); PW = 1.25–1.48 (1.34); PL = 1.13–1.28 (1.2); PW/PL = 1.1– 1.16 (1.12); EW = 1.5–1.88 (1.68); EL = 1.38–1.64 (1.49); PW/HW = 1.26–1.33 (1.3).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFCC11390FB6FF259" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">Head black to brown-black; mouthparts, antennae, pronotum, legs, disc of elytra, and abdominal segments dark reddish-brown; epipleura, abdominal tergites, and sternites with posterior margin light reddish-brown.</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFCC11215FA4CF769" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">
|
||||
Head transverse; dorsally and ventrally glossy with few micropunctures, dorsally without microsculpture, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.46), from 1.4 to 1.86 times longer than the length of temples (in lateral view); distance between eyes about 1.68 times as long as length of the eye in males and 1.8 times in females. Antennal insertions situated closer to frontoclypeus than to eye; antennomeres 2 and 3 subequal in length; antennomeres 5 to 10 subequal in length; antennomere 11 from 1.53 to 1.69 times longer than antennomere
|
||||
<quantity id="4C809B00FFB92E3CFB15154FFB4FF5F0" box="[1153,1206,1131,1155]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="24" pageNumber="25" unit="in" value="10.0">10 in</quantity>
|
||||
males and from
|
||||
<quantity id="4C809B00FFB92E3CFAF5154FFCB7F5D1" metricMagnitude="-2" metricUnit="m" metricValue="3.937" metricValueMax="4.064" metricValueMin="3.81" pageId="24" pageNumber="25" unit="in" value="1.55" valueMax="1.6" valueMin="1.5">1.5 to 1.6 in</quantity>
|
||||
females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; no small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and entire apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than penultimate and more or less subconical.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFCC11706FA3AF7A5" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">Pronotum transverse, convex, evenly curved, usually with one PPDS with additional paired punctures adjacent to AMRSP (APP), without SLSP, with few coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra evenly setose and with several coarse non-setiferous punctures.Mesosternumwithfourmacrosetaearrangedinarowmedially.</paragraph>
|
||||
<paragraph id="8BC736E5FFB92E3CFCC117FAFA3AF6C2" blockId="24.[824,1476,567,1970]" pageId="24" pageNumber="25">In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin slightly sinuate. Protibiae sexually dimorphic, with laterodorsal row of thick spines only in females. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4withpaleadhesivesetae,withoutterminalplate.</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFF1911B4FF2AF353" blockId="25.[113,763,144,576]" pageId="25" pageNumber="26">
|
||||
Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line (
|
||||
<figureCitation id="13432A60FFB82E3DFFE811EBFF33F194" box="[124,202,207,231]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="25" pageNumber="26">Fig. 9A</figureCitation>
|
||||
); in females tergite VIII emarginate medio-apically. Sternite III with basal transverse carina converging evenly at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with deep V-shaped emargination medially (
|
||||
<figureCitation id="13432A60FFB82E3DFEE71068FE36F017" box="[371,463,332,356]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="25" pageNumber="26">Fig. 14A</figureCitation>
|
||||
); female sternite VIII with U-shaped emargination medially; male sternite IX with obtuse emargination medially, its basal portion shorter than distal portion; male tergite X subtruncate medially (
|
||||
<figureCitation id="13432A60FFB82E3DFDAB108EFD63F0B1" box="[575,666,426,450]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="25" pageNumber="26">Fig. 12A</figureCitation>
|
||||
); female tergite X with U-shaped emargination medially (
|
||||
<figureCitation id="13432A60FFB82E3DFD1F10EDFD11F092" box="[651,744,457,481]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="25" pageNumber="26">Fig. 13A</figureCitation>
|
||||
); ovipositor, its second gonocoxite without macrosetae medially (
|
||||
<figureCitation id="13432A60FFB82E3DFFE8132CFF3BF353" box="[124,194,520,544]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="25" pageNumber="26">Fig. 9F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFF191303FD49F34C" blockId="25.[113,763,144,576]" box="[141,688,551,576]" pageId="25" pageNumber="26">
|
||||
Aedeagus. As in
|
||||
<figureCitation id="13432A60FFB82E3DFEA71303FE3CF34C" box="[307,453,551,575]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="25" pageNumber="26">Figure 10A–C</figureCitation>
|
||||
; its total length ~ 1.21.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFFE51345FF15F247" blockId="25.[113,764,609,821]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B90CEAF7FFB82E3DFFE51345FDAFF30B" box="[113,598,609,633]" italics="true" pageId="25" pageNumber="26">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FFB82E3DFE131345FDAFF30B" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[391,598,609,633]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">Loncovilius germaini</taxonomicName>
|
||||
</emphasis>
|
||||
is known only from
|
||||
<collectingCountry id="F36F7675FFB82E3DFF3E13A4FF1DF3EB" box="[170,228,640,664]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
|
||||
, from the
|
||||
<collectingRegion id="49BCF807FFB82E3DFEC413A4FE43F3EB" box="[336,442,640,664]" country="Chile" name="Araucania" pageId="25" pageNumber="26">Araucanía</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFB82E3DFE5113A4FDFFF3EB" box="[453,518,640,664]" country="Chile" name="Aysen" pageId="25" pageNumber="26">Aysén</collectingRegion>
|
||||
, and
|
||||
<collectingRegion id="49BCF807FFB82E3DFDA913A4FD5EF3EB" box="[573,679,640,664]" country="Chile" name="Los Lagos" pageId="25" pageNumber="26">Los Lagos</collectingRegion>
|
||||
regions where it occurs in the eastern portion of the Valdivian temperate forest ecoregion, from
|
||||
<quantity id="4C809B00FFB82E3DFE25139AFDACF3A5" box="[433,597,702,726]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.45" metricValueMin="0.75" pageId="25" pageNumber="26" unit="m" value="1100.0" valueMax="1450.0" valueMin="750.0">750 to 1450 m</quantity>
|
||||
of elevation. It has been collected by sissing leaf liưer and moss, also by using Malaise and window traps, as well as carrion and dung baited pitfall traps.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFB82E3DFFE51272FF2FF72D" blockId="25.[112,763,854,1630]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B90CEAF7FFB82E3DFFE51272FE44F21D" box="[113,445,854,878]" italics="true" pageId="25" pageNumber="26">
|
||||
Remarks:
|
||||
<taxonomicName id="4C784D66FFB82E3DFF7E1272FE44F21D" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[234,445,854,878]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">Loncovilius germaini</taxonomicName>
|
||||
</emphasis>
|
||||
was described as
|
||||
<taxonomicName id="4C784D66FFB82E3DFD051272FF1CF2FE" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cribripennis">
|
||||
<emphasis id="B90CEAF7FFB82E3DFD051272FF1CF2FE" italics="true" pageId="25" pageNumber="26">Philonthus cribripennis</emphasis>
|
||||
</taxonomicName>
|
||||
based on an unspecified number of
|
||||
<typeStatus id="54C38847FFB82E3DFDD81252FD5EF2FD" box="[588,679,886,910]" pageId="25" pageNumber="26" type="syntype">syntypes</typeStatus>
|
||||
from an uncertain type locality in
|
||||
<collectingCountry id="F36F7675FFB82E3DFEE212B1FE48F2DE" box="[374,433,917,941]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
|
||||
. Ŋis name was found to be preoccupied and replaced by another preoccupied name
|
||||
<taxonomicName id="4C784D66FFB82E3DFD041291FE54F298" authority="Bernhauer and Schubert 1914" authorityName="Bernhauer and Schubert" authorityYear="1914" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chilensis">
|
||||
<emphasis id="B90CEAF7FFB82E3DFD041291FD67F2BF" box="[656,670,949,972]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FFB82E3DFD3E1290FD03F2BF" box="[682,762,948,972]" italics="true" pageId="25" pageNumber="26">chilensis</emphasis>
|
||||
Bernhauer and Schubert 1914
|
||||
</taxonomicName>
|
||||
; the laưer was replaced again by
|
||||
<taxonomicName id="4C784D66FFB82E3DFFE612D7FE51F578" authority="Scheerpeltz, 1933" authorityName="Scheerpeltz" authorityYear="1933" box="[114,424,1011,1035]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">
|
||||
<emphasis id="B90CEAF7FFB82E3DFFE612D7FF79F579" box="[114,128,1011,1034]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FFB82E3DFF1A12D7FF1DF579" box="[142,228,1011,1034]" italics="true" pageId="25" pageNumber="26">germaini</emphasis>
|
||||
Scheerpeltz, 1933
|
||||
</taxonomicName>
|
||||
. Coiffiat and Sáiz (1966, 1968) and Sáiz (1971) treated
|
||||
<taxonomicName id="4C784D66FFB82E3DFEEC1537FE12F559" authorityName="Scheerpeltz" authorityYear="1933" box="[376,491,1042,1066]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">
|
||||
<emphasis id="B90CEAF7FFB82E3DFEEC1537FE7EF559" box="[376,391,1043,1066]" italics="true" pageId="25" pageNumber="26">L</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FFB82E3DFE011537FE12F559" box="[405,491,1043,1066]" italics="true" pageId="25" pageNumber="26">germaini</emphasis>
|
||||
</taxonomicName>
|
||||
as a member of the subgenus
|
||||
<taxonomicName id="4C784D66FFB82E3DFF201516FEE0F53A" box="[180,281,1074,1097]" pageId="25" pageNumber="26" rank="subGenus" subGenus="Lienturius">
|
||||
<emphasis id="B90CEAF7FFB82E3DFF201516FEE0F53A" box="[180,281,1074,1097]" italics="true" pageId="25" pageNumber="26">Lienturius</emphasis>
|
||||
</taxonomicName>
|
||||
of the genus
|
||||
<taxonomicName id="4C784D66FFB82E3DFE311515FDEDF53A" authorityName="Germain" authorityYear="1903" box="[421,532,1073,1097]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FFB82E3DFE311515FDEDF53A" box="[421,532,1073,1097]" italics="true" pageId="25" pageNumber="26">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
without adding much data about morphology or distribution of this species, for which even the aedeagus hitherto remained not illustrated. Coiffait and Sáiz (1966) mentioned that the species was known from a female
|
||||
<typeStatus id="54C38847FFB82E3DFF59158BFED3F5B4" box="[205,298,1199,1223]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||||
kept in the collection of the Natural History Museum of
|
||||
<collectingCountry id="F36F7675FFB82E3DFF7A15EAFED0F595" box="[238,297,1230,1254]" name="Chile" pageId="25" pageNumber="26">Chile</collectingCountry>
|
||||
. At the same time, Camousseight (1980) did not list this species among the type material in Germain’s collection in this museum. We did not find any material that could be interpreted as types of
|
||||
<emphasis id="B90CEAF7FFB82E3DFE1A1409FDF1F437" box="[398,520,1325,1348]" italics="true" pageId="25" pageNumber="26">
|
||||
<taxonomicName id="4C784D66FFB82E3DFE1A1409FDFDF437" authorityName="Scheerpeltz" authorityYear="1933" box="[398,516,1325,1348]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="germaini">L. germaini</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
Given the uncertainty about type material in the original description and the catalogue by Camousseight (1980), the type status of the female listed by Coiffait and Sáiz (1966) as ‘holotype’ must be verified in the future. For now, we have matched the identity of this species based on the description by Coiffait and Sáiz (1966). As explained by Newton (2017), Herman’s (2001a) interpretation of both names as replacements for only
|
||||
<taxonomicName id="4C784D66FFB82E3DFD8E172CFD5FF76C" box="[538,678,1543,1567]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cribripennis">
|
||||
<emphasis id="B90CEAF7FFB82E3DFD8E172CFD5FF76C" box="[538,678,1543,1567]" italics="true" pageId="25" pageNumber="26">P. cribripennis</emphasis>
|
||||
</taxonomicName>
|
||||
and his resurrection of
|
||||
<taxonomicName id="4C784D66FFB82E3DFE841703FE8EF74C" authorityName="Bernhauer and Schubert" authorityYear="1914" box="[272,375,1575,1599]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="chilensis">
|
||||
<emphasis id="B90CEAF7FFB82E3DFE841703FE8EF74C" box="[272,375,1575,1599]" italics="true" pageId="25" pageNumber="26">P. chilensis</emphasis>
|
||||
</taxonomicName>
|
||||
as the valid name for this species was incorrect.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,935 +0,0 @@
|
|||
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<mods:title id="204F546B3A657320430A0D74A7C9B24C">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="46C092C27582873BA9563778DF32F645">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="3CCE2BC6654AE27F09D5410E5D268FF6">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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<mods:number id="54DF03712F603085AAA9A719CDD671AD">202</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="9670CBFB9D1295433867F5204CFAA5C5" type="issue">
|
||||
<mods:number id="525B570C4A0DEA6A127EBD50974E3C6C">1</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="30C34324D8C9AE41EDED7659EE15D80B" unit="page">
|
||||
<mods:start id="48C765DD09B4C7CA14E493DE968D314D">1</mods:start>
|
||||
<mods:end id="9ADEE73246A45B2324C95AADC7D08FA7">42</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="AE5D13302A53C564BFE1B1907D214D48">
|
||||
<mods:url id="E9199325612FC1390693664BDC001F44">https://doi.org/10.1093/zoolinnean/zlad143</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="6453168EE6FBFE7F5CB8E9BDA7A677EF">journal article</mods:classification>
|
||||
<mods:identifier id="76607354D304A87E776D394B0DD35E93" type="DOI">10.1093/zoolinnean/zlad143</mods:identifier>
|
||||
<mods:identifier id="E4C4EC239AF36FA6F0508700F30E850D" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="B6B19AD8F268ACDBA32A3507C6AF390A" type="Zenodo-Dep">14541979</mods:identifier>
|
||||
<mods:identifier id="EC13F46434041DB4EB63E470D1640BCB" type="ZooBank">CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03D187F3FFBA2E39FC481020FEE0F51A" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A" httpUri="http://treatment.plazi.org/id/03D187F3FFBA2E39FC481020FEE0F51A" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
|
||||
<subSubSection id="C362656EFFBA2E3FFC481020FAF8F06C" box="[988,1281,260,287]" pageId="27" pageNumber="28" type="nomenclature">
|
||||
<paragraph id="8BC736E5FFBA2E3FFC481020FAF8F06C" blockId="27.[988,1281,260,287]" box="[988,1281,260,287]" pageId="27" pageNumber="28">
|
||||
<heading id="D08F8189FFBA2E3FFC481020FAF8F06C" bold="true" box="[988,1281,260,287]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="28" reason="2">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFC481020FAF8F06C" bold="true" box="[988,1281,260,287]" pageId="27" pageNumber="28">
|
||||
<taxonomicName id="4C784D66FFBA2E3FFC481020FB48F06C" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[988,1201,260,287]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFC481020FB48F06C" bold="true" box="[988,1201,260,287]" italics="true" pageId="27" pageNumber="28">Loncovilius variabilis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFFBA2E3FFB211021FAF8F06C" box="[1205,1281,261,287]" pageId="27" pageNumber="28" rank="species">sp.nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBA2E3FFC011010FB52F0EF" pageId="27" pageNumber="28" type="description">
|
||||
<paragraph id="8BC736E5FFBA2E3FFC011010FAB1F03D" blockId="27.[917,1352,308,334]" box="[917,1352,308,334]" pageId="27" pageNumber="28">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFC011010FAB1F03D" bold="true" box="[917,1352,308,334]" pageId="27" pageNumber="28">
|
||||
(
|
||||
<figureCitation id="13432A60FFBA2E3FFC341010FC16F03D" box="[928,1007,308,334]" captionStart="Figure 1" captionStartId="1.[113,178,1870,1894]" captionTargetBox="[152,1420,144,1842]" captionTargetId="figure-6@1.[152,1420,144,1842]" captionTargetPageId="1" captionText="Figure 1. Habitats of Loncovilius. A, L. edwardsianus (Korge, 1963) live specimen in the Valdivian Temperate Rainforest in Chiloé, Los Lagos, Chile (Photograph: Matías Gargiulo, through iNaturalist). B, PN Puyehue, Chile, L. lividipennis was collected in mosses on the stones at river banks and under stones in accumulations along banks (photograph: Martin Fikáček). C, L. variabilis sp. nov. found in a flower of the Chiloé archipelago (photograph:Matías Gargiulo, through iNaturalist)." pageId="27" pageNumber="28">Figs 1C</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBA2E3FFC6E1010FBE2F03D" box="[1018,1051,308,334]" captionStart="Figure 5" captionStartId="13.[113,178,1868,1892]" captionTargetBox="[154,1419,145,1840]" captionTargetId="figure-6@13.[153,1420,144,1840]" captionTargetPageId="13" captionText="Figure 5. Female habitus of Loncovilius species.A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis." pageId="27" pageNumber="28">5D</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBA2E3FFBB11010FBBBF03D" box="[1061,1090,308,334]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="27" pageNumber="28">9B</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBA2E3FFBD91010FB6DF03D" box="[1101,1172,308,334]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="27" pageNumber="28">10I–K</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBA2E3FFB091010FB35F03D" box="[1181,1228,308,334]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="27" pageNumber="28">12D</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBA2E3FFB421010FAFDF03D" box="[1238,1284,308,334]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="27" pageNumber="28">13D</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBA2E3FFA9A1010FAC4F03D" box="[1294,1341,308,334]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="27" pageNumber="28">14D</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBA2E3FFCBE1041FB52F0EF" blockId="27.[810,1459,357,412]" pageId="27" pageNumber="28">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFCBE1041FBECF00E" box="[810,1045,357,381]" italics="true" pageId="27" pageNumber="28">Zoobank registration:</emphasis>
|
||||
<uri id="FFE93AE7FFBA2E3FFBA01041FB51F0EF" pageId="27" pageNumber="28">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFDE0C30FFBA2E3FFAA41041FB51F0EF" pageId="27" pageNumber="28">6008BBBE-2EF2-4459-8ACD-5AA4E242AA4A</uuid>
|
||||
</uri>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBA2E38FCBE109BFDBEF656" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="materials_examined">
|
||||
<paragraph id="8BC736E5FFBA2E38FCBE109BFDBEF656" blockId="27.[809,1460,446,1974]" lastBlockId="28.[127,779,144,1829]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFCBE109BFBE1F0A5" box="[810,1048,446,470]" italics="true" pageId="27" pageNumber="28">
|
||||
Type material.
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFC2D109AFB1FF3A3" collectingDate="1997-01-23" collectingDateMax="1997-01-25" collectingDateMin="1997-01-23" collectorName="A. Newton & M. Thayer" country="Chile" elevation="230" latitude="-41.852585" location="Palena Pr." longLatPrecision="1" longitude="-72.70292" municipality="Hualaihue" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Los Lagos" typeStatus="holotype">
|
||||
<typeStatus id="54C38847FFBA2E3FFC2D109AFBEAF0A5" box="[953,1043,446,470]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB89109AFBB6F0A5" box="[1053,1103,446,470]" pageId="27" pageNumber="28" type="male">male</specimenCount>
|
||||
, pinned, with genitalia in a separate microvial, with labels as follows:
|
||||
<collectingCountry id="F36F7675FFBA2E3FFBFA10F9FB39F086" box="[1134,1216,477,501]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
: [
|
||||
<collectingRegion id="49BCF807FFBA2E3FFB5910FAFAC0F085" box="[1229,1337,478,502]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFAD410F9FA54F086" box="[1344,1453,477,501]" pageId="27" pageNumber="28">Hualaihué</collectingMunicipality>
|
||||
]
|
||||
<location id="8EA7603EFFBA2E3FFCBD10D9FC69F367" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFCBD10D9FC69F367" box="[809,912,509,533]" country="Chile" latitude="-41.852585" longLatPrecision="1" longitude="-72.70292" municipality="Hualaihue" name="Palena Pr." pageId="27" pageNumber="28" stateProvince="Los Lagos">Palena Pr.</location>
|
||||
:
|
||||
<location id="8EA7603EFFBA2E3FFC0C10D9FBB8F366" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFC0C10D9FBB8F366" box="[920,1089,509,533]" country="Chile" latitude="-41.852585" longLatPrecision="1" longitude="-72.70292" municipality="Hualaihue" name="Austral Highway" pageId="27" pageNumber="28" stateProvince="Los Lagos">Austral Highway</location>
|
||||
, km 62.9 (
|
||||
<quantity id="4C809B00FFBA2E3FFB3C10D9FB09F366" box="[1192,1264,509,533]" metricMagnitude="3" metricUnit="m" metricValue="6.7" pageId="27" pageNumber="28" unit="km" value="6.7">6.7 km</quantity>
|
||||
S Contao turnoff),
|
||||
<quantity id="4C809B00FFBA2E3FFCBE1338FC9EF347" box="[810,871,540,564]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="28" unit="m" value="230.0">
|
||||
<elevation id="0055D1D6FFBA2E3FFCBE1338FC9EF347" box="[810,871,540,564]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="28" unit="m" value="230.0">230m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFCFB1338FC17F347" box="[879,1006,538,564]" degrees="41" direction="south" minutes="51.155" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="-41.852585">
|
||||
41°51.155
|
||||
<emphasis id="B90CEAF7FFBA2E3FFC42133EFC25F340" box="[982,988,538,563]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFC631338FB79F340" box="[1015,1152,538,564]" degrees="72" direction="west" minutes="42.175" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="-72.70292">
|
||||
72°42.175
|
||||
<emphasis id="B90CEAF7FFBA2E3FFBCA133EFB9DF340" box="[1118,1124,538,563]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
[-41.85, -72.70],
|
||||
<date id="FFC61025FFBA2E3FFAB91338FA56F347" box="[1325,1455,540,564]" pageId="27" pageNumber="28" value="1997-01-23" valueMax="1997-01-25" valueMin="1997-01-23">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFAB91338FA56F347" box="[1325,1455,540,564]" pageId="27" pageNumber="28" value="1997-01-23" valueMax="1997-01-25" valueMin="1997-01-23">23-25.i.1997</collectingDate>
|
||||
</date>
|
||||
, young secondary Valdivian rainforest;/beating flowers
|
||||
<taxonomicName id="4C784D66FFBA2E3FFAFF131FFC7BF300" class="Magnoliopsida" family="Ericaceae" genus="Tepulia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="stipularis">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFAFF131FFC7BF300" italics="true" pageId="27" pageNumber="28">Tepulia stipularis</emphasis>
|
||||
</taxonomicName>
|
||||
(myrtaceae) [
|
||||
<taxonomicName id="4C784D66FFBA2E3FFB83137FFB70F300" box="[1047,1161,603,627]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
|
||||
]
|
||||
<collectorName id="268D5333FFBA2E3FFB0D137FFAF4F301" box="[1177,1293,603,627]" pageId="27" pageNumber="28">A. Newton</collectorName>
|
||||
and
|
||||
<collectorName id="268D5333FFBA2E3FFAD3137FFA4AF300" box="[1351,1459,603,627]" pageId="27" pageNumber="28">M. Thayer</collectorName>
|
||||
FIELD MUSEUM NAT. HIST./HOLOTYPE
|
||||
<taxonomicName id="4C784D66FFBA2E3FFAD2135EFC7BF3C2" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="variabilis">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFAD2135EFC7BF3C2" italics="true" pageId="27" pageNumber="28">Loncovilius variabilis</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
|
||||
</materialsCitation>
|
||||
</emphasis>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFB7B139DFC0AF25D" collectingDate="1978-01-02" collectionCode="FMNH" collectorName="L. E. Pena" country="Chile" location="Cautin Temuco" municipality="Vilcun" pageId="27" pageNumber="28" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Araucania" typeStatus="paratype">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFB7B139DFAADF3A3" box="[1263,1364,697,720]" italics="true" pageId="27" pageNumber="28">
|
||||
<typeStatus id="54C38847FFBA2E3FFB7B139DFAB6F3A3" box="[1263,1359,697,720]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
<collectingCountry id="F36F7675FFBA2E3FFACA139DFA49F3A2" box="[1374,1456,697,721]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
: [
|
||||
<collectingRegion id="49BCF807FFBA2E3FFCA413FCFC65F383" box="[816,924,728,752]" country="Chile" name="Araucania" pageId="27" pageNumber="28">Araucanía</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFC3813FCFC0BF383" box="[940,1010,728,752]" pageId="27" pageNumber="28">Vilcún</collectingMunicipality>
|
||||
]
|
||||
<location id="8EA7603EFFBA2E3FFB9113FCFB52F39C" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFB9113FCFB52F39C" box="[1029,1195,728,752]" country="Chile" municipality="Vilcun" name="Cautin Temuco" pageId="27" pageNumber="28" stateProvince="Araucania">Cautín Temuco</location>
|
||||
,
|
||||
<location id="8EA7603EFFBA2E3FFB2813FCFAC3F383" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFB2813FCFAC3F383" box="[1212,1338,728,752]" country="Chile" municipality="Vilcun" name="Cherquenco" pageId="27" pageNumber="28" stateProvince="Araucania">Cherquenco</location>
|
||||
[~ -38.69, -72.00]
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFC1513D3FC22F27C" box="[897,987,759,783]" pageId="27" pageNumber="28">I-II.1978</collectingDate>
|
||||
<collectorName id="268D5333FFBA2E3FFC7C13DCFBA3F27C" box="[1000,1114,760,783]" pageId="27" pageNumber="28">L. E. Peña</collectorName>
|
||||
leg. FIELD MUS. NAT. HIST. [
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFCA61233FCA2F25D" box="[818,859,791,815]" count="3" pageId="27" pageNumber="28" type="male">3♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFCF01233FC72F25D" box="[868,907,791,815]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFC001233FC10F25D" box="[916,1001,791,814]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="27" pageNumber="28">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFC681233FBCDF2FF" collectingDate="2003-12-01" collectingDateMax="2003-12-10" collectingDateMin="2003-12-01" collectionCode="NHM, NHMD, CZUG" collectorName="P. M. Hammond / Below" country="Chile" location="Queulat N. P." pageId="27" pageNumber="28" specimenCount="14" specimenCount-female="6" specimenCount-male="6" stateProvince="Aysen">
|
||||
<collectingCountry id="F36F7675FFBA2E3FFC681233FBB7F25C" box="[1020,1102,791,815]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
: Reg. XI [
|
||||
<collectingRegion id="49BCF807FFBA2E3FFB2C1233FB04F25C" box="[1208,1277,791,815]" country="Chile" name="Aysen" pageId="27" pageNumber="28">Aysén</collectingRegion>
|
||||
]
|
||||
<location id="8EA7603EFFBA2E3FFA981233FA6BF25D" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFA981233FA6BF25D" box="[1292,1426,791,815]" country="Chile" name="Queulat N. P." pageId="27" pageNumber="28" stateProvince="Aysen">Queulat N.P.</location>
|
||||
[~ -44.38, -72.25]
|
||||
<collectorName id="268D5333FFBA2E3FFC5B1212FB3BF23D" box="[975,1218,822,846]" pageId="27" pageNumber="28">P.M. Hammond/Below</collectorName>
|
||||
base
|
||||
<date id="FFC61025FFBA2E3FFA961212FA96F23D" box="[1282,1391,822,846]" pageId="27" pageNumber="28" value="2003-12-01" valueMax="2003-12-10" valueMin="2003-12-01">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFA961212FA96F23D" box="[1282,1391,822,846]" pageId="27" pageNumber="28" value="2003-12-01" valueMax="2003-12-10" valueMin="2003-12-01">1-10.xii.03</collectingDate>
|
||||
</date>
|
||||
shrub flowers/
|
||||
<collectionCode id="ED69AE20FFBA2E3FFC121271FC31F21E" box="[902,968,853,877]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="27" pageNumber="28">NHM</collectionCode>
|
||||
/[
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFC431271FC05F21F" box="[983,1020,853,877]" count="5" pageId="27" pageNumber="28" type="male">5♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB921271FBD4F21F" box="[1030,1069,853,877]" count="4" pageId="27" pageNumber="28" type="female">4♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFBA31272FB83F21E" box="[1079,1146,854,877]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="27" pageNumber="28">NHM</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB1C1271FB57F21F" box="[1160,1198,853,877]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB2C1271FB26F21F" box="[1208,1247,853,877]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFB7D1272FAC5F21E" box="[1257,1340,854,877]" collectionName="NHMD" pageId="27" pageNumber="28">NHMD</collectionCode>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFAD21272FA9AF21E" box="[1350,1379,854,877]" pageId="27" pageNumber="28" type="generic">ex.</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFAF81272FA56F21E" box="[1388,1455,854,877]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="27" pageNumber="28">NHM</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFCBE1251FCA8F2FF" box="[810,849,885,909]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFCCF1251FC5AF2FE" box="[859,931,885,909]" collectionName="Mexico, Jalisco, Guadalajara, Universidad de Guadalajara, Centro de Estudios en Zoologia, Entomologia" pageId="27" pageNumber="28">CZUG</collectionCode>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFC391251FC33F2FF" box="[941,970,885,908]" pageId="27" pageNumber="28" type="generic">ex.</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFC471251FBE0F2FF" box="[979,1049,885,908]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="27" pageNumber="28">NHM</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFBA01251FAD1F2D8" collectingDate="2003-11-21" collectingDateMax="2003-12-12" collectingDateMin="2003-11-21" collectingMethod="Malaise trap" collectionCode="NHM" location="Glacier Path" municipality="Glacier Path" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1">
|
||||
same data as above except:]
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFAFE1251FCA1F2DF" pageId="27" pageNumber="28">Glacier Path</collectingMunicipality>
|
||||
<date id="FFC61025FFBA2E3FFCF612B0FC09F2DF" box="[866,1008,916,940]" pageId="27" pageNumber="28" value="2003-11-21" valueMax="2003-12-12" valueMin="2003-11-21">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFCF612B0FC09F2DF" box="[866,1008,916,940]" pageId="27" pageNumber="28" value="2003-11-21" valueMax="2003-12-12" valueMin="2003-11-21">21.xi-12.xii.03</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<collectingMethod id="52394EF2FFBA2E3FFC6312B0FB78F2D8" box="[1015,1153,916,940]" pageId="27" pageNumber="28">Malaise trap</collectingMethod>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB0112B0FB45F2D8" box="[1173,1212,916,940]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFB5312B0FAF4F2D8" box="[1223,1293,916,939]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="27" pageNumber="28">NHM</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFABC12B0FC6CF57A" collectingDate="2003-11-21" collectingDateMax="2003-12-12" collectingDateMin="2003-11-21" collectionCode="NHM" country="Chile" location="Base area Beating [or] Base Camp Area" municipality="Hualaihue" pageId="27" pageNumber="28" specimenCount="15" specimenCount-female="7" specimenCount-male="8" stateProvince="Los Lagos">
|
||||
same data as above except:]
|
||||
<location id="8EA7603EFFBA2E3FFC581290FA89F2BF" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFC581290FA89F2BF" box="[972,1392,947,972]" country="Chile" municipality="Hualaihue" name="Base area Beating [or] Base Camp Area" pageId="27" pageNumber="28" stateProvince="Los Lagos">Base area Beating [or] Base Camp Area</location>
|
||||
<date id="FFC61025FFBA2E3FFAED1297FC7BF298" pageId="27" pageNumber="28" value="2003-11-21" valueMax="2003-12-12" valueMin="2003-11-21">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFAED1297FC7BF298" pageId="27" pageNumber="28" value="2003-11-21" valueMax="2003-12-12" valueMin="2003-11-21">21.xi-12.xii.03</collectingDate>
|
||||
</date>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFC0112F7FC47F299" box="[917,958,979,1003]" count="8" pageId="27" pageNumber="28" type="male">8♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFC5E12F7FC08F299" box="[970,1009,979,1002]" count="7" pageId="27" pageNumber="28" type="female">7♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFC6812F7FBBBF299" box="[1020,1090,979,1002]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="27" pageNumber="28">NHM</collectionCode>
|
||||
];
|
||||
<collectingCountry id="F36F7675FFBA2E3FFBCC12F7FB53F298" box="[1112,1194,979,1003]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
: [
|
||||
<collectingRegion id="49BCF807FFBA2E3FFB2B12F7FAC8F298" box="[1215,1329,979,1003]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFAD412F7FA54F298" box="[1344,1453,979,1003]" pageId="27" pageNumber="28">Hualaihué</collectingMunicipality>
|
||||
]
|
||||
<location id="8EA7603EFFBA2E3FFCBE12D6FC6BF57A" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFCBE12D6FC6BF57A" box="[810,914,1010,1034]" country="Chile" municipality="Hualaihue" name="Palena Pr." pageId="27" pageNumber="28" stateProvince="Los Lagos">Palena Pr.</location>
|
||||
:
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFC0E12D6FAD7F5F4" collectingDate="1997-01-23" collectingDateMax="1997-01-25" collectingDateMin="1997-01-23" collectionCode="FMNH" collectorName="A. Newton & M. Thayer" elevation="230" latitude="-41.8525" location="Contao turnoff" longLatPrecision="11" longitude="-72.702835" municipality="Austral Highway" pageId="27" pageNumber="28" specimenCount="5" specimenCount-male="5">
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFC0E12D6FBBDF579" box="[922,1092,1010,1034]" pageId="27" pageNumber="28">Austral Highway</collectingMunicipality>
|
||||
, km 62.9 (
|
||||
<locationDeviation id="9A752FC7FFBA2E3FFB3812D6FAFBF579" bearing="180.0" box="[1196,1282,1010,1034]" distance="6700.0" pageId="27" pageNumber="28">
|
||||
<quantity id="4C809B00FFBA2E3FFB3812D6FB16F579" box="[1196,1263,1010,1034]" metricMagnitude="3" metricUnit="m" metricValue="6.7" pageId="27" pageNumber="28" unit="km" value="6.7">6.7km</quantity>
|
||||
S
|
||||
</locationDeviation>
|
||||
<location id="8EA7603EFFBA2E3FFA9C12D6FA50F579" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFA9C12D6FA50F579" box="[1288,1449,1010,1034]" latitude="-41.8525" longLatPrecision="11" longitude="-72.702835" municipality="Austral Highway" name="Contao turnoff" pageId="27" pageNumber="28">Contao turnoff</location>
|
||||
),
|
||||
<quantity id="4C809B00FFBA2E3FFCBF1535FC91F55A" box="[811,872,1041,1065]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="28" unit="m" value="230.0">
|
||||
<elevation id="0055D1D6FFBA2E3FFCBF1535FC91F55A" box="[811,872,1041,1065]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="28" unit="m" value="230.0">230m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFCE21535FC1DF55A" box="[886,996,1041,1065]" degrees="41" direction="south" minutes="51.15" orientation="latitude" pageId="27" pageNumber="28" precision="9" value="-41.8525">41°51.15’S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFC671535FB95F55A" box="[1011,1132,1041,1065]" degrees="72" direction="west" minutes="42.17" orientation="longitude" pageId="27" pageNumber="28" precision="9" value="-72.702835">72°42.17’W</geoCoordinate>
|
||||
[-41.85, -72.70],
|
||||
<date id="FFC61025FFBA2E3FFABA1535FA49F55A" box="[1326,1456,1041,1065]" pageId="27" pageNumber="28" value="1997-01-23" valueMax="1997-01-25" valueMin="1997-01-23">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFABA1535FA49F55A" box="[1326,1456,1041,1065]" pageId="27" pageNumber="28" value="1997-01-23" valueMax="1997-01-25" valueMin="1997-01-23">23-25.i.1997</collectingDate>
|
||||
</date>
|
||||
, young secondary Valdivian rainforest;/beating flowers
|
||||
<taxonomicName id="4C784D66FFBA2E3FFAFF1515FC7BF51B" class="Magnoliopsida" family="Ericaceae" genus="Tepulia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="stipularis">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFAFF1515FC7BF51B" italics="true" pageId="27" pageNumber="28">Tepulia stipularis</emphasis>
|
||||
</taxonomicName>
|
||||
(myrtaceae) [
|
||||
<taxonomicName id="4C784D66FFBA2E3FFB831574FB70F51B" box="[1047,1161,1104,1128]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
|
||||
]
|
||||
<collectorName id="268D5333FFBA2E3FFB0D1574FAF4F514" box="[1177,1293,1104,1128]" pageId="27" pageNumber="28">A. Newton</collectorName>
|
||||
and
|
||||
<collectorName id="268D5333FFBA2E3FFAD31574FA4AF51B" box="[1351,1459,1104,1128]" pageId="27" pageNumber="28">M. Thayer</collectorName>
|
||||
FIELD MUSEUM NAT. HIST [
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB03154BFB39F5F5" box="[1175,1216,1135,1159]" count="5" pageId="27" pageNumber="28" type="male">5♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFB5B1554FADDF5F4" box="[1231,1316,1136,1159]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="27" pageNumber="28">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFAA9154BFB80F5B6" collectingDate="1985-02" collectionCode="FMNH" collectorName="J. Zuniga" country="Chile" location="Llanquihue Lago Chapo" pageId="27" pageNumber="28" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Los Lagos">
|
||||
<collectingCountry id="F36F7675FFBA2E3FFAA9154BFA8CF5F4" box="[1341,1397,1135,1159]" name="Chile" pageId="27" pageNumber="28">Chile</collectingCountry>
|
||||
[
|
||||
<collectingRegion id="49BCF807FFBA2E3FFA1D1554FC93F5D4" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
|
||||
]
|
||||
<location id="8EA7603EFFBA2E3FFCED15ABFB8EF5D4" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFCED15ABFB8EF5D4" box="[889,1143,1167,1191]" country="Chile" name="Llanquihue Lago Chapo" pageId="27" pageNumber="28" stateProvince="Los Lagos">Llanquihue Lago Chapo</location>
|
||||
[-41.40, -72.52]
|
||||
<date id="FFC61025FFBA2E3FFABA15ABFA4AF5D5" box="[1326,1459,1166,1191]" pageId="27" pageNumber="28" value="1985-02">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFABA15ABFA4AF5D5" box="[1326,1459,1166,1191]" pageId="27" pageNumber="28" value="1985-02">febrero 1985</collectingDate>
|
||||
</date>
|
||||
leg.
|
||||
<collectorName id="268D5333FFBA2E3FFCC7158AFC57F5B5" box="[851,942,1198,1222]" pageId="27" pageNumber="28">J. Zuñiga</collectorName>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFC2A158AFC1FF5B6" box="[958,998,1198,1222]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFC78158AFBEAF5B6" box="[1004,1043,1198,1222]" count="3" pageId="27" pageNumber="28" type="female">3♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFB8E158AFB96F5B6" box="[1050,1135,1198,1221]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="27" pageNumber="28">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFBEB158AFAA3F476" collectingDate="1972-12" collectionCode="FMNH" collectorName="L. E. Pena" location="HORNOHUINCO SW. Lago Chapo Lianquihue" municipality="Llanquihue" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Los Lagos">
|
||||
<location id="8EA7603EFFBA2E3FFBEB158AFC0AF596" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFBEB158AFC0AF596" municipality="Llanquihue" name="HORNOHUINCO SW. Lago Chapo Lianquihue" pageId="27" pageNumber="28" stateProvince="Los Lagos">HORNOHUINCO SW. Lago Chapo Lianquihue</location>
|
||||
[
|
||||
<collectingRegion id="49BCF807FFBA2E3FFB9315EAFB85F595" box="[1031,1148,1230,1254]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFB1915E9FAFAF596" box="[1165,1283,1229,1253]" pageId="27" pageNumber="28">Llanquihue</collectingMunicipality>
|
||||
, -41.40, -72.52] 21,24-
|
||||
<date id="FFC61025FFBA2E3FFCF815C9FC31F476" box="[876,968,1261,1285]" pageId="27" pageNumber="28" value="1972-12">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFCF815C9FC31F476" box="[876,968,1261,1285]" pageId="27" pageNumber="28" value="1972-12">Dic.1972</collectingDate>
|
||||
</date>
|
||||
Coll:
|
||||
<collectorName id="268D5333FFBA2E3FFB9315C9FB9DF476" box="[1031,1124,1261,1285]" pageId="27" pageNumber="28">L.E.Pena</collectorName>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFBE815C9FB5AF477" box="[1148,1187,1261,1285]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB2615C9FB21F477" box="[1202,1240,1261,1285]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFB7315C9FAC5F477" box="[1255,1340,1261,1284]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="27" pageNumber="28">FMNH</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFACF15C9FB57F410" collectingDate="1957-02-08" collectionCode="CNCI, CNC" country="Chile" location="Pena" municipality="Rio Gol-Gol Osorno Prov" pageId="27" pageNumber="28" specimenCode="L2004-28, FMNH-INS 0000 024 263-265" specimenCount="4" specimenCount-male="4" stateProvince="Los Lagos">
|
||||
<collectingCountry id="F36F7675FFBA2E3FFACF15C9FA49F476" box="[1371,1456,1261,1285]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBA2E3FFCBE1428FC61F457" box="[810,920,1292,1316]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
|
||||
]
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFC311428FB4DF450" box="[933,1204,1292,1316]" pageId="27" pageNumber="28">Rio Gol-Gol Osorno Prov</collectingMunicipality>
|
||||
. [~ -40.66, -72.18]/
|
||||
<date id="FFC61025FFBA2E3FFA111428FCBDF430" pageId="27" pageNumber="28" value="1957-02-08">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFA111428FCBDF430" pageId="27" pageNumber="28" value="1957-02-08">II-8- 57</collectingDate>
|
||||
</date>
|
||||
<location id="8EA7603EFFBA2E3FFCC4140FFC7DF430" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFCC4140FFC7DF430" box="[848,900,1323,1347]" country="Chile" municipality="Rio Gol-Gol Osorno Prov" name="Pena" pageId="27" pageNumber="28" stateProvince="Los Lagos">Peña</location>
|
||||
/
|
||||
<collectingCountry id="F36F7675FFBA2E3FFC18140FFC22F430" box="[908,987,1323,1347]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
[green label]/
|
||||
<collectionCode id="ED69AE20FFBA2E3FFBED140FFB45F430" box="[1145,1212,1323,1347]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="27" pageNumber="28">CNCI</collectionCode>
|
||||
<specimenCode id="DBDE9E9EFFBA2E3FFB53140FFAD6F430" box="[1223,1327,1323,1347]" pageId="27" pageNumber="28">L2004-28</specimenCode>
|
||||
/
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFAA3140AFAA8F431" box="[1335,1361,1326,1346]" count="1" pageId="27" pageNumber="28" type="male">♂</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFFBA2E3FFACE140FFBD9F411" pageId="27" pageNumber="28">FMNH-INS 0000 024 263-265</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFBA6146FFBA2F411" box="[1074,1115,1355,1379]" count="3" pageId="27" pageNumber="28" type="male">3♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFBF1146FFB5AF410" box="[1125,1187,1355,1379]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="27" pageNumber="28">CNC</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFB2C146FFAEAF4B3" collectingDate="1998-01-18" collectionCode="FMNH, MNNC" collectorName="T. Cekalovic" country="Chile" location="Chiloe" municipality="Chiloe Pr." pageId="27" pageNumber="28" specimenCode="FMNH-INS 0000 024 165-167" specimenCount="4" specimenCount-female="1" specimenCount-male="2">
|
||||
<collectingCountry id="F36F7675FFBA2E3FFB2C146FFAF3F410" box="[1208,1290,1355,1379]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFA83146FFA7AF411" box="[1303,1411,1355,1379]" pageId="27" pageNumber="28">Chiloé Pr.</collectingMunicipality>
|
||||
,
|
||||
<collectingCountry id="F36F7675FFBA2E3FFA04146FFA4AF410" box="[1424,1459,1355,1379]" name="Iceland" pageId="27" pageNumber="28">Isla</collectingCountry>
|
||||
<location id="8EA7603EFFBA2E3FFCBE144EFC89F4F1" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFCBE144EFC89F4F1" box="[810,880,1386,1410]" country="Chile" municipality="Chiloe Pr." name="Chiloe" pageId="27" pageNumber="28">Chiloé</location>
|
||||
,
|
||||
<location id="8EA7603EFFBA2E3FFCEA144EFBFBF4F1" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFCEA144EFBFBF4F1" box="[894,1026,1386,1410]" country="Chile" municipality="Chiloe Pr." name="Compu Alto" pageId="27" pageNumber="28">Compu Alto</location>
|
||||
[-42.87, -73.70],
|
||||
<date id="FFC61025FFBA2E3FFB55144EFAD8F4F1" box="[1217,1313,1386,1410]" pageId="27" pageNumber="28" value="1998-01-18">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFB55144EFAD8F4F1" box="[1217,1313,1386,1410]" pageId="27" pageNumber="28" value="1998-01-18">18.i.1998</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="268D5333FFBA2E3FFAA4144EFA4AF4F1" box="[1328,1459,1386,1410]" pageId="27" pageNumber="28">T. Cekalovic</collectorName>
|
||||
leg. FIELD MUS. NAT. HIST./
|
||||
<specimenCode id="DBDE9E9EFFBA2E3FFBF814ADFA4AF4D2" box="[1132,1459,1417,1441]" pageId="27" pageNumber="28">FMNH-INS 0000 024 165-167</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFCA1148DFCA2F4B3" box="[821,859,1449,1473]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFCFC148DFC40F4B3" box="[872,953,1449,1472]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="27" pageNumber="28">FMNH</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFC5C148DFC16F4B3" box="[968,1007,1449,1473]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFC6F148DFBDBF4B3" box="[1019,1058,1449,1473]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFBBA148DFB86F4B2" box="[1070,1151,1449,1473]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="27" pageNumber="28">MNNC</collectionCode>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB18148DFB50F4B3" box="[1164,1193,1449,1472]" pageId="27" pageNumber="28" type="generic">ex.</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFB20148DFAF0F4B3" box="[1204,1289,1449,1472]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="27" pageNumber="28">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFA8B148DFBFFF76C" collectingDate="1998-01-18" collectionCode="SEMC" collectorName="T. Cekalovic" country="Chile" county="Isla Chiloe" location="Compu Alto" municipality="Prov. Chiloe" pageId="27" pageNumber="28" specimenCode="SM0722732 KUNHM-ENT" specimenCount="1" specimenCount-male="1">
|
||||
<collectingCountry id="F36F7675FFBA2E3FFA8B148DFA88F4B2" box="[1311,1393,1449,1473]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFA14148DFC8AF493" pageId="27" pageNumber="28">Prov. Chiloé</collectingMunicipality>
|
||||
:
|
||||
<collectingCounty id="62A64E69FFBA2E3FFCEB14ECFC09F493" box="[895,1008,1480,1504]" pageId="27" pageNumber="28">Isla Chiloé</collectingCounty>
|
||||
,
|
||||
<location id="8EA7603EFFBA2E3FFC6814ECFB87F493" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFC6814ECFB87F493" box="[1020,1150,1480,1504]" country="Chile" county="Isla Chiloe" municipality="Prov. Chiloe" name="Compu Alto" pageId="27" pageNumber="28">Compu Alto</location>
|
||||
[-42.87, -73.70]
|
||||
<date id="FFC61025FFBA2E3FFAA614ECFA6EF493" box="[1330,1431,1480,1504]" pageId="27" pageNumber="28" value="1998-01-18">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFAA614ECFA6EF493" box="[1330,1431,1480,1504]" pageId="27" pageNumber="28" value="1998-01-18">18-I-1998</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FFBA2E3FFA0B14ECFC76F48C" pageId="27" pageNumber="28">T. Cekalovic</collectorName>
|
||||
<accessionNumber id="942BAB06FFBA2E3FFC0E14C3FBBBF48C" box="[922,1090,1511,1535]" pageId="27" pageNumber="28">CHIL1C98 001</accessionNumber>
|
||||
/[Barcode]
|
||||
<specimenCode id="DBDE9E9EFFBA2E3FFB2B14C3FCA6F76D" pageId="27" pageNumber="28">SM0722732 KUNHM-ENT</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFCE51723FC61F76D" box="[881,920,1543,1567]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFC351723FC17F76C" box="[929,1006,1543,1567]" collectionName="USA, Kansas, Lawrence, University of Kansas, Snow Entomological Museum" pageId="27" pageNumber="28">SEMC</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFB931723FBDAF72E" collectingDate="1969-01-22" collectionCode="FMNH" collectorName="L. E. Pena" country="Chile" county="Puerto Montt" location="Prov. Llanquihue Correntoso" municipality="Prov. Llanquihue Correntoso" pageId="27" pageNumber="28" specimenCount="2" specimenCount-male="2" stateProvince="Los Lagos">
|
||||
<collectingCountry id="F36F7675FFBA2E3FFB931723FBA5F76C" box="[1031,1116,1543,1567]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBA2E3FFBF31723FB2DF76C" box="[1127,1236,1543,1567]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="62A64E69FFBA2E3FFB4B1723FA88F76D" box="[1247,1393,1543,1566]" pageId="27" pageNumber="28">Puerto Montt</collectingCounty>
|
||||
]
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFA141723FBE0F74D" pageId="27" pageNumber="28">Prov. Llanquihue Correntoso</collectingMunicipality>
|
||||
[~ -41.45, -72.66]
|
||||
<date id="FFC61025FFBA2E3FFB4B1702FA80F74D" box="[1247,1401,1574,1598]" pageId="27" pageNumber="28" value="1969-01-22">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFB4B1702FA80F74D" box="[1247,1401,1574,1598]" pageId="27" pageNumber="28" value="1969-01-22">22-Enero-1969</collectingDate>
|
||||
</date>
|
||||
Coll:
|
||||
<collectorName id="268D5333FFBA2E3FFCB81762FC7EF72E" box="[812,903,1606,1629]" pageId="27" pageNumber="28">L.E.Pena</collectorName>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFC001761FC44F72F" box="[916,957,1605,1629]" count="2" pageId="27" pageNumber="28" type="male">2♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFC501762FBE0F72E" box="[964,1049,1606,1629]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="27" pageNumber="28">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFBBE1761FAA3F789" collectingDate="1997-01-17" collectionCode="FMNH" collectorName="A. Newton & M. Thayer" country="Chile" county="Chonchi" elevation="130" latitude="-42.7788" location="Miraflores" longLatPrecision="6" longitude="-73.7953" municipality="Chiloe Pr." pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Los Lagos">
|
||||
<collectingCountry id="F36F7675FFBA2E3FFBBE1761FB85F72E" box="[1066,1148,1605,1629]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
: [
|
||||
<collectingRegion id="49BCF807FFBA2E3FFB181762FB03F72D" box="[1164,1274,1606,1630]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="62A64E69FFBA2E3FFA901761FA98F72E" box="[1284,1377,1605,1629]" pageId="27" pageNumber="28">Chonchi</collectingCounty>
|
||||
]
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFAFA1761FCB5F70F" pageId="27" pageNumber="28">Chiloé Pr.</collectingMunicipality>
|
||||
:
|
||||
<location id="8EA7603EFFBA2E3FFCCD1741FC3AF70E" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFCCD1741FC3AF70E" box="[857,963,1637,1661]" country="Chile" county="Chonchi" latitude="-42.7788" longLatPrecision="6" longitude="-73.7953" municipality="Chiloe Pr." name="Miraflores" pageId="27" pageNumber="28" stateProvince="Los Lagos">Miraflores</location>
|
||||
, road to (
|
||||
<quantity id="4C809B00FFBA2E3FFBA61741FB8FF70E" box="[1074,1142,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="27" pageNumber="28" unit="km" value="0.6">
|
||||
<locationDeviation id="9A752FC7FFBA2E3FFBA61741FB8FF70E" box="[1074,1142,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="27" pageNumber="28" unit="km" value="0.6">0.6km</locationDeviation>
|
||||
</quantity>
|
||||
W Hwy 5),
|
||||
<quantity id="4C809B00FFBA2E3FFA9C1741FABCF70E" box="[1288,1349,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="28" unit="m" value="130.0">
|
||||
<elevation id="0055D1D6FFBA2E3FFA9C1741FABCF70E" box="[1288,1349,1637,1661]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="27" pageNumber="28" unit="m" value="130.0">130m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFAC01741FCCEF7EF" degrees="42" direction="south" minutes="46.73" orientation="latitude" pageId="27" pageNumber="28" precision="9" value="-42.77883">
|
||||
42°46.73
|
||||
<emphasis id="B90CEAF7FFBA2E3FFA391747FA4AF70F" box="[1453,1459,1635,1660]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFCDD17A0FC2BF7E8" box="[841,978,1666,1692]" degrees="73" direction="west" minutes="47.71" orientation="longitude" pageId="27" pageNumber="28" precision="9" value="-73.795166">
|
||||
73°47.71
|
||||
<emphasis id="B90CEAF7FFBA2E3FFC3717A6FC50F7E8" box="[931,937,1666,1691]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
[
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFC7E17A0FBB2F7EF" box="[1002,1099,1668,1692]" degrees="42.7788" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="-42.7788">-42.7788</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFBCB17A0FB47F7EF" box="[1119,1214,1668,1692]" degrees="73.7953" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="-73.7953">-73.7953</geoCoordinate>
|
||||
]
|
||||
<date id="FFC61025FFBA2E3FFB4217A0FACCF7EF" box="[1238,1333,1668,1692]" pageId="27" pageNumber="28" value="1997-01-17">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFB4217A0FACCF7EF" box="[1238,1333,1668,1692]" pageId="27" pageNumber="28" value="1997-01-17">17.i.1997</collectingDate>
|
||||
</date>
|
||||
, secondary Valdivian rainforest:/beating flowering Cald-cluvia paniculata [
|
||||
<taxonomicName id="4C784D66FFBA2E3FFCA117E7FBFEF7A8" box="[821,1031,1731,1755]" class="Magnoliopsida" family="Cunoniaceae" genus="Caldcluvia" kingdom="Plantae" order="Oxalidales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="paniculata">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFCA117E7FBFEF7A8" box="[821,1031,1731,1755]" italics="true" pageId="27" pageNumber="28">Caldcluvia paniculata</emphasis>
|
||||
</taxonomicName>
|
||||
] (
|
||||
<taxonomicName id="4C784D66FFBA2E3FFB8B17E7FB50F7A8" box="[1055,1193,1731,1755]" class="Magnoliopsida" family="Cunoniaceae" kingdom="Plantae" order="Oxalidales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Cunoniaceae</taxonomicName>
|
||||
)
|
||||
<collectorName id="268D5333FFBA2E3FFB2317E7FAD9F7A8" box="[1207,1312,1731,1755]" pageId="27" pageNumber="28">A.Newton</collectorName>
|
||||
and
|
||||
<collectorName id="268D5333FFBA2E3FFAC717E7FA4AF7A8" box="[1363,1459,1731,1755]" pageId="27" pageNumber="28">M.Thayer</collectorName>
|
||||
994 FIELD MUS. NAT. HIST. [
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB1C17C6FB56F78A" box="[1160,1199,1762,1786]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFB2D17C6FB19F78A" box="[1209,1248,1762,1786]" count="1" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFB7F17C6FAB9F78A" box="[1259,1344,1762,1785]" pageId="27" pageNumber="28">FMNH</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E3FFACF17C6FBE9F6E5" collectingDate="2014-12-13" collectingDateMax="2015-05" collectingDateMin="2014-12-13" collectionCode="NHMW" collectorName="R. Vila" country="Chile" determinerName="V. Assing" elevation="40" latitude="-42.458557" location="peninsula Huequi under logs termas, Porcelana" longLatPrecision="1" longitude="-72.45416" municipality="Hualaihue" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Los Lagos">
|
||||
<collectingCountry id="F36F7675FFBA2E3FFACF17C6FA49F789" box="[1371,1456,1762,1786]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBA2E3FFCBE1626FC60F669" box="[810,921,1794,1818]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFC3C1625FBECF66A" box="[936,1045,1793,1817]" pageId="27" pageNumber="28">Hualaihué</collectingMunicipality>
|
||||
] 11 Chile X,
|
||||
<location id="8EA7603EFFBA2E3FFB261625FBF7F64A" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFB261625FBF7F64A" country="Chile" latitude="-42.458557" longLatPrecision="1" longitude="-72.45416" municipality="Hualaihue" name="peninsula Huequi under logs termas, Porcelana" pageId="27" pageNumber="28" stateProvince="Los Lagos">peninsula Huequi under logs termas, Porcelana</location>
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFB811605FB5BF64A" box="[1045,1186,1823,1849]" degrees="42" direction="south" minutes="27" orientation="latitude" pageId="27" pageNumber="28" precision="1" seconds="30.8" value="-42.458557">
|
||||
42°27
|
||||
<emphasis id="B90CEAF7FFBA2E3FFBC5163BFBAEF64B" box="[1105,1111,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
|
||||
30.8
|
||||
<emphasis id="B90CEAF7FFBA2E3FFB16163BFB74F64B" box="[1154,1165,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹʹ</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
<geoCoordinate id="EE4C5022FFBA2E3FFB3E1605FAB8F64B" box="[1194,1345,1823,1849]" degrees="72" direction="west" minutes="27" orientation="longitude" pageId="27" pageNumber="28" precision="1" seconds="15.0" value="-72.45416">
|
||||
72°27
|
||||
<emphasis id="B90CEAF7FFBA2E3FFB71163BFB12F64B" box="[1253,1259,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹ</emphasis>
|
||||
15.0
|
||||
<emphasis id="B90CEAF7FFBA2E3FFA83163BFADBF64B" box="[1303,1314,1823,1848]" italics="true" pageId="27" pageNumber="28">ʹʹ</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
[-42.4585, -72.4541]
|
||||
<quantity id="4C809B00FFBA2E3FFC0C1664FC36F624" box="[920,975,1856,1880]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="m" value="40.0">
|
||||
<elevation id="0055D1D6FFBA2E3FFC0C1664FC36F624" box="[920,975,1856,1880]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="m" value="40.0">40 m</elevation>
|
||||
</quantity>
|
||||
<collectorName id="268D5333FFBA2E3FFC431664FBE6F62B" box="[983,1055,1856,1880]" pageId="27" pageNumber="28">R. Vila</collectorName>
|
||||
leg.
|
||||
<date id="FFC61025FFBA2E3FFBC71664FB3BF62B" box="[1107,1218,1856,1880]" pageId="27" pageNumber="28" value="2014-12-13">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFBC71664FB3BF62B" box="[1107,1218,1856,1880]" pageId="27" pageNumber="28" value="2014-12-13">13.xii.2014</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<taxonomicName id="4C784D66FFBA2E3FFB5E1664FA91F624" authority="sp" authorityName="Sp" box="[1226,1384,1856,1880]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="30" phylum="Arthropoda" rank="genus">Loncovilius sp</taxonomicName>
|
||||
det.
|
||||
<determinerName id="7E1BDA21FFBA2E3FFA0B1664FC94F604" pageId="27" pageNumber="28">V. Assing</determinerName>
|
||||
<date id="FFC61025FFBA2E3FFCE1167BFC33F604" box="[885,970,1887,1911]" bridgedPair="/" pageId="27" pageNumber="28" value="2015-05">
|
||||
<collectingDate id="EF82E9CDFFBA2E3FFCE1167BFC33F604" box="[885,970,1887,1911]" bridgedPair="/" pageId="27" pageNumber="28" value="2015-05">2015/V.</collectingDate>
|
||||
</date>
|
||||
Assing Coll. J Jenkins Shaw det. 2016 [yellow label] [
|
||||
<specimenCount id="9D7EFD6CFFBA2E3FFCE2165BFC67F6E5" box="[886,926,1919,1943]" count="1" pageId="27" pageNumber="28" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBA2E3FFC33165BFBFFF6E5" box="[935,1030,1919,1942]" pageId="27" pageNumber="28">NHMW</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBA2E38FB8D165BFF19F076" collectingDate="1982-12-19" collectingDateMax="1982-12-26" collectingDateMin="1982-12-19" collectionCode="FMNH" collectorName="A. Newton & M. Thayer" country="Chile" determinerName="A. Newton" elevation="430" lastPageId="28" lastPageNumber="29" location="Osorno Prov. Parque Nac. Puyehue" municipality="Puyehue National Park" pageId="27" pageNumber="28" specimenCode="FMNH-INS 0000 024 163" specimenCount="3" specimenCount-female="2" stateProvince="Los Lagos">
|
||||
<collectingCountry id="F36F7675FFBA2E3FFB8D165BFB92F6E4" box="[1049,1131,1919,1943]" name="Chile" pageId="27" pageNumber="28">CHILE</collectingCountry>
|
||||
: [
|
||||
<collectingRegion id="49BCF807FFBA2E3FFBE9165BFB14F6E4" box="[1149,1261,1919,1943]" country="Chile" name="Los Lagos" pageId="27" pageNumber="28">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BA3AC9FFFBA2E3FFB6E165BFCA5F6C5" pageId="27" pageNumber="28">Puyehue National Park</collectingMunicipality>
|
||||
]
|
||||
<location id="8EA7603EFFBA2E3FFCF816BAFB11F6C5" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFCF816BAFB11F6C5" box="[876,1256,1950,1974]" country="Chile" municipality="Puyehue National Park" name="Osorno Prov. Parque Nac. Puyehue" pageId="27" pageNumber="28" stateProvince="Los Lagos">Osorno Prov. Parque Nac. Puyehue</location>
|
||||
,
|
||||
<locationDeviation id="9A752FC7FFBA2E3FFB6216BAFAA9F6C6" bearing="90.0" box="[1270,1360,1950,1974]" distance="4100.0" pageId="27" pageNumber="28">
|
||||
<quantity id="4C809B00FFBA2E3FFB6216BAFACFF6C5" box="[1270,1334,1950,1974]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="27" pageNumber="28" unit="km" value="4.1">4.1km</quantity>
|
||||
E
|
||||
</locationDeviation>
|
||||
<location id="8EA7603EFFBA2E3FFACF16BAFA4DF6C5" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBA2E3FFACF16BAFA4DF6C5" box="[1371,1460,1950,1974]" country="Chile" municipality="Puyehue National Park" name="Anticura" pageId="27" pageNumber="28" stateProvince="Los Lagos">Anticura</location>
|
||||
[-40.66, -72.12],
|
||||
<quantity id="4C809B00FFBD2E38FEA111B4FE8BF1DB" box="[309,370,144,168]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="28" pageNumber="29" unit="m" value="430.0">
|
||||
<elevation id="0055D1D6FFBD2E38FEA111B4FE8BF1DB" box="[309,370,144,168]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="28" pageNumber="29" unit="m" value="430.0">430m</elevation>
|
||||
</quantity>
|
||||
, trap site 662
|
||||
<date id="FFC61025FFBD2E38FD9A11B4FD58F1DB" box="[526,673,144,168]" pageId="28" pageNumber="29" value="1982-12-19" valueMax="1982-12-26" valueMin="1982-12-19">
|
||||
<collectingDate id="EF82E9CDFFBD2E38FD9A11B4FD58F1DB" box="[526,673,144,168]" pageId="28" pageNumber="29" value="1982-12-19" valueMax="1982-12-26" valueMin="1982-12-19">19-26.xii.1982</collectingDate>
|
||||
</date>
|
||||
Valdivian rainfor.
|
||||
<collectorName id="268D5333FFBD2E38FF5B1194FEC0F1BB" box="[207,313,176,200]" pageId="28" pageNumber="29">A.Newton</collectorName>
|
||||
and
|
||||
<collectorName id="268D5333FFBD2E38FEFE118BFE3EF1B4" box="[362,455,175,199]" pageId="28" pageNumber="29">M.Thayer</collectorName>
|
||||
/Screen- sweeping/
|
||||
<taxonomicName id="4C784D66FFBD2E38FD1B118BFF64F195" authority="sp." authorityName="Sp." class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||||
<location id="8EA7603EFFBD2E38FD1B118BFCF0F1B4" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBD2E38FD1B118BFCF0F1B4" box="[655,777,175,199]" country="Chile" municipality="Puyehue National Park" name="Loncovilius" pageId="28" pageNumber="29" stateProvince="Los Lagos">Loncovilius</location>
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FF1511EBFF64F195" box="[129,157,207,230]" pageId="28" pageNumber="29" type="generic">sp.</specimenCount>
|
||||
</taxonomicName>
|
||||
7 det.
|
||||
<determinerName id="7E1BDA21FFBD2E38FF7111EBFEB7F194" box="[229,334,207,231]" pageId="28" pageNumber="29">A.Newton</determinerName>
|
||||
1984/
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FE0111F6FE56F195" box="[405,431,210,230]" count="1" pageId="28" pageNumber="29" type="female">♀</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFFBD2E38FE2311EBFD28F194" box="[439,721,207,231]" pageId="28" pageNumber="29">FMNH-INS 0000 024 163</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FD7611EBFCF0F195" box="[738,777,207,231]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FF1511CAFF2FF076" box="[129,214,238,261]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="28" pageNumber="29">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FF7211CAFDCFF0F0" collectingDate="1984-12-20" collectingDateMax="1985-02-08" collectingDateMin="1984-12-20" collectionCode="FMNH" country="Chile" elevation="500" location="Osorno Prov" municipality="Puyehue National Park" pageId="28" pageNumber="29" specimenCode="FMNH-INS 0000 024 169" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Los Lagos">
|
||||
<collectingCountry id="F36F7675FFBD2E38FF7211CAFEC1F075" box="[230,312,238,262]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
: [
|
||||
<collectingRegion id="49BCF807FFBD2E38FEDC11CAFE4FF075" box="[328,438,238,262]" country="Chile" name="Los Lagos" pageId="28" pageNumber="29">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6BA3AC9FFFBD2E38FE5411CAFD56F075" box="[448,687,238,262]" pageId="28" pageNumber="29">Puyehue National Park</collectingMunicipality>
|
||||
]
|
||||
<location id="8EA7603EFFBD2E38FD2F11CAFF4DF056" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBD2E38FD2F11CAFF4DF056" country="Chile" municipality="Puyehue National Park" name="Osorno Prov" pageId="28" pageNumber="29" stateProvince="Los Lagos">Osorno Prov</location>
|
||||
.,
|
||||
<location id="8EA7603EFFBD2E38FF541029FDE1F056" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBD2E38FF541029FDE1F056" box="[192,536,269,294]" country="Chile" municipality="Puyehue National Park" name="Puyehue Nat. Pk. Aguas Calientes" pageId="28" pageNumber="29" stateProvince="Los Lagos">Puyehue Nat. Pk. Aguas Calientes</location>
|
||||
[-40.73, -72.30],
|
||||
<quantity id="4C809B00FFBD2E38FD5E1029FCFEF056" box="[714,775,269,293]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="28" pageNumber="29" unit="m" value="500.0">
|
||||
<elevation id="0055D1D6FFBD2E38FD5E1029FCFEF056" box="[714,775,269,293]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="28" pageNumber="29" unit="m" value="500.0">500m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFC61025FFBD2E38FF151008FF07F037" box="[129,254,300,324]" pageId="28" pageNumber="29" value="1984-12-20">
|
||||
<collectingDate id="EF82E9CDFFBD2E38FF151008FF07F037" box="[129,254,300,324]" pageId="28" pageNumber="29" value="1984-12-20">20.XII.1984</collectingDate>
|
||||
</date>
|
||||
-
|
||||
<date id="FFC61025FFBD2E38FE911008FEA4F037" box="[261,349,300,324]" pageId="28" pageNumber="29" value="1985-02-08">
|
||||
<collectingDate id="EF82E9CDFFBD2E38FE911008FEA4F037" box="[261,349,300,324]" pageId="28" pageNumber="29" value="1985-02-08">8.II.1985</collectingDate>
|
||||
</date>
|
||||
/FMHD#85-930
|
||||
<location id="8EA7603EFFBD2E38FDB61009FCFFF036" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBD2E38FDB61009FCFFF036" box="[546,774,301,325]" country="Chile" municipality="Puyehue National Park" name="Derumbes for. Trail" pageId="28" pageNumber="29" stateProvince="Los Lagos">Derumbes for. Trail</location>
|
||||
, S.andJ. Peck, P#85-45, FIT FIELD MUSEUM NAT HIST/
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FD77106BFD04F010" box="[739,765,335,355]" count="1" pageId="28" pageNumber="29" type="female">♀</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFFBD2E38FF15104FFE6EF0F0" box="[129,407,363,387]" pageId="28" pageNumber="29">FMNH-INS 0000 024 169</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FE3C104FFE36F0F1" box="[424,463,363,387]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FE431048FDD5F0F0" box="[471,556,364,387]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="28" pageNumber="29">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FDAA104FFD52F373" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FDAA104FFD69F0F0" box="[574,656,363,387]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
: [
|
||||
<collectingRegion id="49BCF807FFBD2E38FD351048FCFDF0F0" box="[673,772,363,387]" country="Chile" name="Los Rios" pageId="28" pageNumber="29">Los Ríos</collectingRegion>
|
||||
]
|
||||
<collectingMunicipality id="6BA3AC9FFFBD2E38FF1510AFFEF6F0D1" box="[129,271,395,419]" pageId="28" pageNumber="29">Valdivia Prov</collectingMunicipality>
|
||||
.,
|
||||
<locationDeviation id="9A752FC7FFBD2E38FEB410AFFE74F0D0" bearing="135.0" box="[288,397,395,419]" distance="26000.0" pageId="28" pageNumber="29">
|
||||
<quantity id="4C809B00FFBD2E38FEB410AFFE9FF0D0" box="[288,358,395,419]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="28" pageNumber="29" unit="km" value="26.0">
|
||||
<locationDeviation id="9A752FC7FFBD2E38FEB410AFFE9FF0D0" box="[288,358,395,419]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="28" pageNumber="29" unit="km" value="26.0">26 km</locationDeviation>
|
||||
</quantity>
|
||||
SE
|
||||
</locationDeviation>
|
||||
<location id="8EA7603EFFBD2E38FE0C10AFFDF2F0D0" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBD2E38FE0C10AFFDF2F0D0" box="[408,523,395,419]" pageId="28" pageNumber="29">Panguipulli</location>
|
||||
[-39.75, -72.15]
|
||||
<quantity id="4C809B00FFBD2E38FD5D10AFFCFCF0D0" box="[713,773,395,419]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="m" value="300.0">
|
||||
<elevation id="0055D1D6FFBD2E38FD5D10AFFCFCF0D0" box="[713,773,395,419]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="m" value="300.0">
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FD5D10AFFCFCF0D0" box="[713,773,395,419]" pageId="28" pageNumber="29" type="male">300m</specimenCount>
|
||||
</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFC61025FFBD2E38FF15108EFF04F0B1" box="[129,253,426,450]" pageId="28" pageNumber="29" value="1984-12-16">
|
||||
<collectingDate id="EF82E9CDFFBD2E38FF15108EFF04F0B1" box="[129,253,426,450]" pageId="28" pageNumber="29" value="1984-12-16">16.XII.1984</collectingDate>
|
||||
</date>
|
||||
-
|
||||
<date id="FFC61025FFBD2E38FE91108EFE93F0B1" box="[261,362,426,450]" pageId="28" pageNumber="29" value="1985-02-11">
|
||||
<collectingDate id="EF82E9CDFFBD2E38FE91108EFE93F0B1" box="[261,362,426,450]" pageId="28" pageNumber="29" value="1985-02-11">11.II.1985</collectingDate>
|
||||
</date>
|
||||
/FMHD#85-921,
|
||||
<location id="8EA7603EFFBD2E38FDBA108EFF68F092" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBD2E38FDBA108EFF68F092" pageId="28" pageNumber="29">
|
||||
<taxonomicName id="4C784D66FFBD2E38FDBA108EFD5CF0B1" box="[558,677,426,450]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
|
||||
remnant S.
|
||||
</location>
|
||||
andJ.
|
||||
<location id="8EA7603EFFBD2E38FF4510EDFEFDF092" LSID="urn:lsid:plazi:treatment:03D187F3FFBA2E39FC481020FEE0F51A:8EA7603EFFBD2E38FF4510EDFEFDF092" box="[209,260,457,481]" pageId="28" pageNumber="29">Peck</location>
|
||||
, P#85-34, carrion trap FIELD MUSEUM NAT HIST/
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FF5C10C8FF1BF373" box="[200,226,492,512]" count="1" pageId="28" pageNumber="29" type="female">♀</specimenCount>
|
||||
/FMNH-INS 0000 024 168 [
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FD8F10CDFDBAF373" box="[539,579,489,513]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FDD810CDFD58F373" box="[588,673,489,512]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="28" pageNumber="29">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FD2010CDFF7EF30D" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FD2010CDFCFFF372" box="[692,774,489,513]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
: [
|
||||
<collectingRegion id="49BCF807FFBD2E38FF13132CFF1EF353" box="[135,231,520,544]" country="Chile" name="Los Rios" pageId="28" pageNumber="29">Los Ríos</collectingRegion>
|
||||
] Prov. Valdivia: Parque Oncol (Cruce Sendero Tepual) Rio Cruces [-39.72, -73.27]
|
||||
<date id="FFC61025FFBD2E38FE321303FE07F34C" box="[422,510,551,575]" pageId="28" pageNumber="29" value="2001-01-08">8-I-2001</date>
|
||||
T Cekalovic CHIL1CO1 005/[Barcode] SM0723095 KUNHM-ENT [
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FDED1363FD59F32D" box="[633,672,583,607]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FD261363FD06F32C" box="[690,767,583,607]" collectionName="USA, Kansas, Lawrence, University of Kansas, Snow Entomological Museum" pageId="28" pageNumber="29">SEMC</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FF131342FD48F3EE" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FF131342FF25F30D" box="[135,220,614,638]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBD2E38FF7C1342FEB1F30D" box="[232,328,614,638]" country="Chile" name="Los Rios" pageId="28" pageNumber="29">Los Ríos</collectingRegion>
|
||||
] 85/Süd-Chile
|
||||
<date id="FFC61025FFBD2E38FE791342FDC1F30D" box="[493,568,614,638]" pageId="28" pageNumber="29" value="90-01-31">31.1.90</date>
|
||||
Puerto Puyehue [~ -40.65, -72.33] leg.G.H. Schwabe [
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FE6413A1FDE3F3EF" box="[496,538,645,669]" count="2" pageId="28" pageNumber="29" type="male">2♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FDB113A1FDB5F3EF" box="[549,588,645,669]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
|
||||
SDEI]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FD2513A1FF7EF268" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FD2513A1FCFFF3EE" box="[689,774,645,669]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBD2E38FF151381FF0AF3CE" box="[129,243,677,701]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
|
||||
and Chilean
|
||||
<collectingCountry id="F36F7675FFBD2E38FEE71381FE19F3CE" box="[371,480,677,701]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
|
||||
] Welling Is. Pto. Eden [-49.11, -74.40]
|
||||
<date id="FFC61025FFBD2E38FF4213E0FEB6F3AF" box="[214,335,708,732]" pageId="28" pageNumber="29" value="1958-12-13">13.XII.1958</date>
|
||||
/
|
||||
<collectingCountry id="F36F7675FFBD2E38FECC13E0FE52F3AF" box="[344,427,708,732]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
Fr Kuschel/
|
||||
<taxonomicName id="4C784D66FFBD2E38FDA213E0FD74F3AF" box="[566,653,708,732]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
|
||||
nigroflavus Tottenham
|
||||
<typeStatus id="54C38847FFBD2E38FE9413C0FEB9F388" box="[256,320,740,763]" pageId="28" pageNumber="29">TYPE</typeStatus>
|
||||
[manuscript name]/
|
||||
<collectionCode id="ED69AE20FFBD2E38FDB213C7FD91F388" box="[550,616,739,763]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FD1513C7FD50F389" box="[641,681,739,763]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FD2C13C0FD07F388" box="[696,766,740,763]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FF131227FEC5F20A" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FF131227FF25F268" box="[135,220,771,795]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBD2E38FF7D1227FEA2F268" box="[233,347,771,795]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
|
||||
and Chilean
|
||||
<collectingCountry id="F36F7675FFBD2E38FE791227FDA3F268" box="[493,602,771,795]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
|
||||
] Welling Is. Pto. Eden [-49.11, -74.40]
|
||||
<date id="FFC61025FFBD2E38FEF41206FE20F249" box="[352,473,802,826]" pageId="28" pageNumber="29" value="1958-12-13">13.XII.1958</date>
|
||||
/
|
||||
<collectingCountry id="F36F7675FFBD2E38FE751206FDCDF249" box="[481,564,802,826]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
Fr Kuschel/
|
||||
<taxonomicName id="4C784D66FFBD2E38FD261206FCF0F249" box="[690,777,802,826]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
|
||||
nigroflavus Tottenham PARATYPE [manuscript name]/
|
||||
<collectionCode id="ED69AE20FFBD2E38FD531265FCF0F22A" box="[711,777,833,857]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FF1D1245FF4BF20B" box="[137,178,865,889]" count="3" pageId="28" pageNumber="29" type="male">3♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FF221245FF24F20B" box="[182,221,865,889]" count="2" pageId="28" pageNumber="29" type="female">2♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FF751245FEDEF20B" box="[225,295,865,888]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FEA81245FF7FF546" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FEA81245FE68F20A" box="[316,401,865,889]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBD2E38FE0C1245FDF3F20A" box="[408,522,865,889]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
|
||||
and Chilean
|
||||
<collectingCountry id="F36F7675FFBD2E38FD1F1245FD05F20A" box="[651,764,865,889]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
|
||||
]/
|
||||
<typeStatus id="54C38847FFBD2E38FF1512A4FF22F2EB" box="[129,219,896,920]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
|
||||
/Welling Is. Pto. Eden [-49.11, -74.40]
|
||||
<date id="FFC61025FFBD2E38FDE712A4FD12F2EB" box="[627,747,896,920]" pageId="28" pageNumber="29" value="1958-11-30">30.XI.1958</date>
|
||||
/C. E. Tottenham collection. B.M. 1974-587/
|
||||
<collectingCountry id="F36F7675FFBD2E38FDA512BBFD7BF2C4" box="[561,642,927,951]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
Fr Kuschel/ nigroflavus Tot [manuscript name]/
|
||||
<collectionCode id="ED69AE20FFBD2E38FDAB129BFD7BF2A4" box="[575,642,959,983]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
/PARATYPE
|
||||
<taxonomicName id="4C784D66FFBD2E38FF1512FAFEAAF285" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[129,339,990,1014]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
|
||||
<emphasis id="B90CEAF7FFBD2E38FF1512FAFEAAF285" box="[129,339,990,1014]" italics="true" pageId="28" pageNumber="29">Loncovilius variabilis</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2023/
|
||||
<collectionCode id="ED69AE20FFBD2E38FE3412D9FDFCF566" box="[416,517,1021,1045]" collectionName="NHMUK" pageId="28" pageNumber="29">NHMUK</collectionCode>
|
||||
015025170 [
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FD1812D9FD4AF567" box="[652,691,1021,1045]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FD2C12DAFD07F566" box="[696,766,1022,1045]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FF131539FDB0F5E1" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FF131539FF25F546" box="[135,220,1053,1077]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBD2E38FF7D1539FEA2F546" box="[233,347,1053,1077]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="28" pageNumber="29">Magallanes</collectingRegion>
|
||||
and Chilean
|
||||
<collectingCountry id="F36F7675FFBD2E38FE791539FDA3F546" box="[493,602,1053,1077]" name="Antarctica" pageId="28" pageNumber="29">Antarctica</collectingCountry>
|
||||
] Welling Is. Pto. Eden [-49.11, -74.40]
|
||||
<date id="FFC61025FFBD2E38FEFA1518FE1EF527" box="[366,487,1084,1108]" pageId="28" pageNumber="29" value="1958-12-13">13.XII.1958</date>
|
||||
/
|
||||
<collectingCountry id="F36F7675FFBD2E38FE641518FDBAF527" box="[496,579,1084,1108]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
Fr Kuschel/Qued. Sp. 5/
|
||||
<taxonomicName id="4C784D66FFBD2E38FF29157FFE98F500" authority="sp." authorityName="Sp." box="[189,353,1115,1139]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||||
Loncovilius
|
||||
<taxonomicNameLabel id="A23F578CFFBD2E38FED11578FE98F500" box="[325,353,1116,1139]" pageId="28" pageNumber="29" rank="species">sp.</taxonomicNameLabel>
|
||||
</taxonomicName>
|
||||
Nov. P.M. Hammond det. 1980/
|
||||
<collectionCode id="ED69AE20FFBD2E38FD5C157FFCF3F500" box="[712,778,1115,1139]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
[Blue label]/
|
||||
<collectionCode id="ED69AE20FFBD2E38FE92155FFEA2F5E0" box="[262,347,1147,1171]" collectionName="NHMD" pageId="28" pageNumber="29">NHMD</collectionCode>
|
||||
917069 [
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FE53155FFE17F5E1" box="[455,494,1147,1171]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FE6C155FFDC7F5E1" box="[504,574,1147,1170]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FDC7155FFE7BF477" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FDC7155FFD5CF5E0" box="[595,677,1147,1171]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBD2E38FD26155FFCF0F5E0" box="[690,777,1147,1171]" country="Chile" name="Region Metropolitana" pageId="28" pageNumber="29">Santiago</collectingRegion>
|
||||
[
|
||||
<collectingRegion id="49BCF807FFBD2E38FF1215BEFF18F5C1" box="[134,225,1178,1202]" country="Chile" name="Region Metropolitana" pageId="28" pageNumber="29">Santiago</collectingRegion>
|
||||
Metropolitan]: El Alfalfal [~ -33.51, -70.20]
|
||||
<date id="FFC61025FFBD2E38FD5C15BEFCF3F5C1" box="[712,778,1178,1202]" pageId="28" pageNumber="29" value="1977-01">I.1977</date>
|
||||
L.E.Peña, leg. FIELD MUS. NAT. HIST./FMNH-INS 0000 024 157–162 [
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FE8815FDFEBAF583" box="[284,323,1241,1265]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FEDF15FDFE8BF583" box="[331,370,1241,1265]" count="2" pageId="28" pageNumber="29" type="female">2♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FEEE15FDFE32F583" box="[378,459,1241,1264]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="28" pageNumber="29">FMNH</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FE4315FDFE07F583" box="[471,510,1241,1265]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FD9215FDFDD4F583" box="[518,557,1241,1265]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FDA115FDFD71F583" box="[565,648,1241,1264]" collectionName="NHMD" pageId="28" pageNumber="29">NHMD</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FFBD2E38FD2015FDFCFCF583" box="[692,773,1241,1264]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="28" pageNumber="29">FMNH</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FF1515DCFF5EF47C" box="[129,167,1272,1296]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FF3A15DCFF0FF463" box="[174,246,1272,1296]" collectionName="Mexico, Jalisco, Guadalajara, Universidad de Guadalajara, Centro de Estudios en Zoologia, Entomologia" pageId="28" pageNumber="29">CZUG</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FFBD2E38FEB415DCFE88F47C" box="[288,369,1272,1295]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="28" pageNumber="29">FMNH</collectionCode>
|
||||
;
|
||||
<superScript id="7C0D9BADFFBD2E38FEE915D2FE7BF477" attach="right" box="[381,386,1270,1284]" fontSize="6" pageId="28" pageNumber="29">*</superScript>
|
||||
</materialsCitation>
|
||||
this is potentially a labelling error. The
|
||||
<taxonomicName id="4C784D66FFBD2E38FF16143CFF0DF45C" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[130,244,1303,1327]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
|
||||
<emphasis id="B90CEAF7FFBD2E38FF16143CFF76F45C" box="[130,143,1304,1327]" italics="true" pageId="28" pageNumber="29">L</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FFBD2E38FF081433FF0DF45C" box="[156,244,1303,1327]" italics="true" pageId="28" pageNumber="29">variabilis</emphasis>
|
||||
</taxonomicName>
|
||||
model shown in
|
||||
<figureCitation id="13432A60FFBD2E38FE3F1433FDF8F45C" box="[427,513,1303,1327]" captionStart="Figure 3" captionStartId="11.[113,178,1776,1800]" captionTargetBox="[114,1458,144,1740]" captionTargetId="figure-6@11.[114,1458,144,1749]" captionTargetPageId="11" captionText="Figure 3. Ecological niche modelling of Loncovilius species. A–D, potential suitable areas predicted with free extrapolation in South America. Binarized projection with 5% threshold for suitability values and mobility-oriented parity analysis (MOP) showing areas for which current model predictions are strictly based on extrapolation. E–H, predictions under shared socio-economic pathways (SSPs) 126, and agreement between different global climate models (GCMs), and MOP agreement among GCMs with strict extrapolative conditions under the low emissions.I–L, predictions under SSPs 585, agreement between different GCMs, and MOP agreement among GCMs with strict extrapolative conditions under the high emissions." pageId="28" pageNumber="29">Figure 3</figureCitation>
|
||||
does not include this occurrence. The sets of variables and the calibration results that do include this occurrence can be consulted in the Supporting Information,
|
||||
<tableCitation id="C6FA035EFFBD2E38FE9D1451FE9BF4FE" box="[265,354,1397,1421]" captionStart="Table 1" captionStartId="3.[115,170,893,917]" captionTargetPageId="3" captionText="Table 1.Taxon sample for the phylogenetic analysis (left column) and resulting taxonomic changes (right column)." pageId="28" pageNumber="29">Table S1</tableCitation>
|
||||
, and in the Supporting Information, File S6]; [
|
||||
<materialsCitation id="3B103CB8FFBD2E38FF2F14B1FF28F779" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FF2F14B1FEE9F4DE" box="[187,272,1429,1453]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
: unknown locality and date]
|
||||
<collectingCountry id="F36F7675FFBD2E38FDD514B1FD8EF4DE" box="[577,631,1429,1453]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
|
||||
./
|
||||
<collectingCountry id="F36F7675FFBD2E38FD1614B1FD41F4DE" box="[642,696,1429,1453]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
|
||||
[Green round label]/Sharp Coll 1905-313./
|
||||
<taxonomicName id="4C784D66FFBD2E38FDB11490FD78F4BF" box="[549,641,1460,1484]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
|
||||
nigroflavus Tottenham PARATYPE [manuscript name]/
|
||||
<collectionCode id="ED69AE20FFBD2E38FDE914F7FD46F498" box="[637,703,1491,1515]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FD7614F7FCF0F499" box="[738,777,1491,1515]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FF1514D7FF3EF779" box="[129,199,1523,1546]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="28" pageNumber="29">NHM</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FF4214D7FF7FF7D4" pageId="28" pageNumber="29">
|
||||
[
|
||||
<collectingCountry id="F36F7675FFBD2E38FF4914D7FECBF778" box="[221,306,1523,1547]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
: unknown locality and date]
|
||||
<collectingCountry id="F36F7675FFBD2E38FDC714D7FD7FF778" box="[595,646,1523,1547]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
|
||||
Sharp Coll./
|
||||
<taxonomicName id="4C784D66FFBD2E38FF151736FF21F759" box="[129,216,1554,1578]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
|
||||
nigroflavus Brnh (Typ.) [manuscript name]/BRIT. MUSEUM DON. ARROW/B. Typ. ohne Flg.d streifen/Chicago NHMus M.Bernhauer Collection/Bernhauer MS name? Current genus:
|
||||
<taxonomicName id="4C784D66FFBD2E38FE891754FE6CF7FB" box="[285,405,1648,1672]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Loncovilius</taxonomicName>
|
||||
teste A.Newton 2000 [
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FDEA1754FD5CF7F4" box="[638,677,1648,1672]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FD3E1754FD06F7F4" box="[682,767,1648,1671]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="28" pageNumber="29">FMNH</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBD2E38FF1217ABFDBEF656" pageId="28" pageNumber="29">
|
||||
<collectingCountry id="F36F7675FFBD2E38FF1217ABFF22F7D4" box="[134,219,1679,1703]" name="Chile" pageId="28" pageNumber="29">CHILE</collectingCountry>
|
||||
: unknown locality and date] 53793/Reed/
|
||||
<collectingCountry id="F36F7675FFBD2E38FD3117ABFD22F7D4" box="[677,731,1679,1703]" name="Chile" pageId="28" pageNumber="29">Chili</collectingCountry>
|
||||
/Fry Coll. 1905.100/Chicago NHMus M.Bernhauer Collection/
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FD771796FD04F7B5" box="[739,765,1714,1734]" count="1" pageId="28" pageNumber="29" type="male">♂</specimenCount>
|
||||
/ FMNH-INS 0000 024 171 [
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FE2517EAFE21F796" box="[433,472,1742,1766]" count="1" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBD2E38FE7617EAFDCFF796" box="[482,566,1742,1765]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="28" pageNumber="29">FMNH</collectionCode>
|
||||
]. All
|
||||
<typeStatus id="54C38847FFBD2E38FDE417EAFD28F795" box="[624,721,1742,1766]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
|
||||
with label: PARATYPE
|
||||
<taxonomicName id="4C784D66FFBD2E38FED317C9FDECF676" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[327,533,1773,1797]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
|
||||
<emphasis id="B90CEAF7FFBD2E38FED317C9FDECF676" box="[327,533,1773,1797]" italics="true" pageId="28" pageNumber="29">Loncovilius variabilis</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBD2E38FF151663FDB0F62D" box="[129,585,1862,1886]" pageId="28" pageNumber="29" type="diagnosis">
|
||||
<paragraph id="8BC736E5FFBD2E38FF151663FDB0F62D" blockId="28.[129,585,1862,1886]" box="[129,585,1862,1886]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B90CEAF7FFBD2E38FF151663FF1EF62D" box="[129,231,1863,1886]" italics="true" pageId="28" pageNumber="29">Diagnosis:</emphasis>
|
||||
Same as
|
||||
<taxonomicName id="4C784D66FFBD2E38FED71662FE65F62D" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[323,412,1862,1886]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variabilis">
|
||||
<emphasis id="B90CEAF7FFBD2E38FED71662FE65F62D" box="[323,412,1862,1886]" italics="true" pageId="28" pageNumber="29">variabilis</emphasis>
|
||||
</taxonomicName>
|
||||
group diagnosis.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBD2E39FF1516A4FD88F353" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
|
||||
<paragraph id="8BC736E5FFBD2E38FF1516A4FA9AF075" blockId="28.[129,778,1919,1975]" lastBlockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
|
||||
<emphasis id="B90CEAF7FFBD2E38FF1516A4FF0EF6E4" box="[129,247,1920,1943]" italics="true" pageId="28" pageNumber="29">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FE4216A6FE09F6E5" box="[470,496,1922,1942]" count="1" pageId="28" pageNumber="29" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFBD2E38FF1516BBFF6DF6C5" box="[129,148,1951,1974]" italics="true" pageId="28" pageNumber="29">N</emphasis>
|
||||
= 10]: FBL = 2.79–3.19 (2.98); TL = 5.12–6.25 (5.61); HW = 0.72–0.8 (0.77); HL = 0.74–0.83 (0.79); HW/ HL = 0.95–1 (0.97); PW = 0.95–1.05 (1); PL = 0.88–0.98 (0.93); PW/PL = 1.02–1.09 (1.07); EW = 1.31–1.54 (1.38); EL = 1.17–1.43 (1.25); PW/HW = 1.23–1.33 (1.30).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBD2E38FCC1102AFB92F0B1" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFBD2E38FB9B1034FBD0F057" box="[1039,1065,272,292]" count="1" pageId="28" pageNumber="29" type="female">♀</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFBD2E38FAF2102AFA80F056" box="[1382,1401,270,293]" italics="true" pageId="28" pageNumber="29">N</emphasis>
|
||||
= 10]: FBL = 3.05–3.53 (3.33); TL = 5.5–7.44 (6.4); HW = 0.79– 0.86 (0.83); HL = 0.82–0.9 (0.85); HW/HL = 0.94–1 (0.97); PW = 1.05–1.17 (1.10); PL = 1–1.08 (1.04); PW/PL = 1.03– 1.09 (1.06); EW = 1.36–1.74 (1.56); EL = 1.22–1.55 (1.43); PW/HW = 1.29–1.39 (1.34).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBD2E38FCC110EDFA72F34C" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">Head, pronotum, and elytra black to brown-black, head darker; antennae, mouthparts, and legs yellowish-brown; abdominal tergites and sternites variable in coloration, from dark reddish-brown to brown-black with pale posterior margin.</paragraph>
|
||||
<paragraph id="8BC736E5FFBD2E38FCC11363FAAEF41D" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
|
||||
Head from about as wide as long to, usually, slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.47), from 1.03 to 1.29 times as long as temples (in lateral view); distance between eyes about 1.25 times as long as length of eye in males and 1.34 times in females. Antennomere 3 slightly longer than antennomere 2; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a11 1.53-1.72 times as long as a
|
||||
<quantity id="4C809B00FFBD2E38FBD812A4FB86F2EB" box="[1100,1151,896,920]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="28" pageNumber="29" unit="in" value="10.0">10 in</quantity>
|
||||
males and
|
||||
<quantity id="4C809B00FFBD2E38FB7B12A4FA90F2EB" box="[1263,1385,896,920]" metricMagnitude="-2" metricUnit="m" metricValue="4.0259" metricValueMax="4.318" metricValueMin="3.7338" pageId="28" pageNumber="29" unit="in" value="1.585" valueMax="1.7" valueMin="1.47">1.47-1.70 in</quantity>
|
||||
females. Basal puncture usually double; parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; no small setiferous punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge more or less straight in checkmark shape, almost united with gular sutures; postgenal ridge reduced to small trace near gular sutures; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with isodiametric microsculpture medially, gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with sub-rectangular sclerotized region and entire to slightly notched apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical to fusiform apical palpomere.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBD2E38FCC11452FAD0F759" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">Pronotum slightly wider than long, convex, evenly curved, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to 5 or even 10 times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
|
||||
<paragraph id="8BC736E5FFBD2E38FCC11715FB73F676" blockId="28.[824,1476,144,1985]" pageId="28" pageNumber="29">
|
||||
Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. Mesotarsomeres 1–4 not sexually dimorphic, in both sexes with pale adhesive setae and mesotarsomeres
|
||||
<quantity id="4C809B00FFBD2E38FB8A17ABFB9FF7D4" box="[1054,1126,1679,1703]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="28" pageNumber="29" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1–3 in</quantity>
|
||||
both sexes with terminal plate. Metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBD2E39FCC11629FD37F036" blockId="28.[824,1476,144,1985]" lastBlockId="29.[113,765,144,544]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||||
Abdominal tergites III and IV with several rows of sparse and fine punctures of moderate density, without sizeable impunctate area; tergite VIII only slightly emarginate medio-apically in females. Sternite III with basal transverse carina strongly arcuate; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with broad and shallow U-shaped emargination medially (
|
||||
<figureCitation id="13432A60FFBC2E39FEFD11B4FE3BF1DB" box="[361,450,144,168]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="29" pageNumber="30">Fig. 14D</figureCitation>
|
||||
); female sternite VIII without emargination; male sternite IX without emargination medially (apex subtruncate), its basal portion moderately longer than distal portion; male tergite X with anterior margin subtruncate (
|
||||
<figureCitation id="13432A60FFBC2E39FFE81029FF2FF056" box="[124,214,269,293]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="29" pageNumber="30">Fig. 12D</figureCitation>
|
||||
); female tergite X apically truncate (
|
||||
<figureCitation id="13432A60FFBC2E39FDCA1029FD41F056" box="[606,696,269,293]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="29" pageNumber="30">Fig. 13D</figureCitation>
|
||||
); ovipositor, its second gonocoxite with one macroseta medially.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBC2E39FF191068FD88F353" blockId="29.[113,765,144,544]" pageId="29" pageNumber="30">
|
||||
Aedeagus as in
|
||||
<figureCitation id="13432A60FFBC2E39FEB81068FE4CF017" box="[300,437,332,356]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="29" pageNumber="30">Figure 10I–K</figureCitation>
|
||||
; its total length ~ 0.95. Median lobe rod-like, apex not emarginate; internal sac with median vertical sclerite and pair of short I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe; wider than median lobe; converging into a subrounded apex, slightly projected in the middle; with dark peg setae; in lateral view paramere almost straight, slightly concave apically.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBC2E39FFE51365FE41F266" pageId="29" pageNumber="30" type="etymology">
|
||||
<paragraph id="8BC736E5FFBC2E39FFE51365FE41F266" blockId="29.[113,763,577,789]" pageId="29" pageNumber="30">
|
||||
<emphasis id="B90CEAF7FFBC2E39FFE51365FF18F32A" box="[113,225,577,601]" italics="true" pageId="29" pageNumber="30">Etymology:</emphasis>
|
||||
The species epithet is the Latin adjective
|
||||
<taxonomicName id="4C784D66FFBC2E39FD011365FD0DF32A" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[661,756,577,601]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="variabilis">
|
||||
‘
|
||||
<emphasis id="B90CEAF7FFBC2E39FD0F1365FD0DF32A" box="[667,756,577,601]" italics="true" pageId="29" pageNumber="30">variabilis</emphasis>
|
||||
</taxonomicName>
|
||||
’ (variable) and refers to a notable intraspecific morphological variation of this taxon. This variation can occur between individuals from the same locality, e.g. it can be completely blackishbrown forms and others with a reddish-brown abdomen. The head punctation also varies, and loss or duplication of paraocular and basal punctures is common.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBC2E39FFE51213FEE0F51A" pageId="29" pageNumber="30" type="distribution">
|
||||
<paragraph id="8BC736E5FFBC2E39FFE51213FEE0F51A" blockId="29.[113,764,823,1129]" pageId="29" pageNumber="30">
|
||||
<emphasis id="B90CEAF7FFBC2E39FFE51213FDA0F23C" box="[113,601,823,847]" italics="true" pageId="29" pageNumber="30">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FFBC2E39FEEB1213FDA0F23C" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[383,601,823,847]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="variabilis">Loncovilius variabilis</taxonomicName>
|
||||
</emphasis>
|
||||
is known only from
|
||||
<collectingCountry id="F36F7675FFBC2E39FF241272FF15F21D" box="[176,236,854,878]" name="Chile" pageId="29" pageNumber="30">Chile</collectingCountry>
|
||||
, from the following regions:
|
||||
<collectingRegion id="49BCF807FFBC2E39FDA31272FD5CF21D" box="[567,677,854,878]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBC2E39FD201272FD0FF21D" box="[692,758,854,878]" country="Chile" name="Aysen" pageId="29" pageNumber="30">Aysén</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBC2E39FFE51252FF1CF2FD" box="[113,229,886,910]" country="Chile" name="Los Lagos" pageId="29" pageNumber="30">Los Lagos</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBC2E39FF611252FEA0F2FE" box="[245,345,885,909]" country="Chile" name="Los Rios" pageId="29" pageNumber="30">Los Ríos</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBC2E39FEFD1251FE1AF2FE" box="[361,483,885,909]" country="Chile" name="Magallanes y de la Antartica Chilena" pageId="29" pageNumber="30">Magallanes</collectingRegion>
|
||||
, and Chilean
|
||||
<collectingCountry id="F36F7675FFBC2E39FD111251FD0FF2FE" box="[645,758,885,909]" name="Antarctica" pageId="29" pageNumber="30">Antarctica</collectingCountry>
|
||||
, and from Santiago Metropolitan. The species occurs from lowlands to
|
||||
<quantity id="4C809B00FFBC2E39FF611290FEB1F2B8" box="[245,328,948,972]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="m" value="1300.0">1300 m</quantity>
|
||||
of elevation in the mountains; confined to various
|
||||
<typeStatus id="54C38847FFBC2E39FF7812F0FEDCF29F" box="[236,293,980,1004]" pageId="29" pageNumber="30">types</typeStatus>
|
||||
of forests in the Magellanic subpolar and Valdivian temperate forest ecoregions. It has been collected using Malaise traps, and directly from flowers, e.g. from
|
||||
<taxonomicName id="4C784D66FFBC2E39FFE51515FEC1F53A" box="[113,312,1073,1097]" class="Magnoliopsida" family="Myrtaceae" genus="Tepualia" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="stipularis">
|
||||
<emphasis id="B90CEAF7FFBC2E39FFE51515FEC1F53A" box="[113,312,1073,1097]" italics="true" pageId="29" pageNumber="30">Tepualia stipularis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C784D66FFBC2E39FEDA1515FE3CF53A" box="[334,453,1073,1097]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
|
||||
) and
|
||||
<taxonomicName id="4C784D66FFBC2E39FD861515FD03F53A" box="[530,762,1073,1097]" class="Magnoliopsida" family="Cunoniaceae" genus="Caldcluvia" kingdom="Plantae" order="Oxalidales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="paniculata">
|
||||
<emphasis id="B90CEAF7FFBC2E39FD861515FD03F53A" box="[530,762,1073,1097]" italics="true" pageId="29" pageNumber="30">Caldcluvia paniculata</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C784D66FFBC2E39FFED1575FEF4F51A" box="[121,269,1105,1129]" class="Magnoliopsida" family="Cunoniaceae" kingdom="Plantae" order="Oxalidales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Cunoniaceae</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,107 +0,0 @@
|
|||
<document id="9871233E3957C86DDC16F4A87B3F7F26" ID-DOI="10.1093/zoolinnean/zlad143" ID-ISSN="0024-4082" ID-Zenodo-Dep="14541979" ID-ZooBank="CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1732839935034" checkinUser="plazi" docAuthor="Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins & Solodovnikov, Alexey" docDate="2024" docId="03D187F3FFBA2E3FFEF11447FC16F1B4" docLanguage="en" docName="zlad143.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlad143" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Loncovilius variabilis Reyes-Hernández & Hansen & Shaw & Solodovnikov 2024" docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="FFE8FF8BFFA12E24FF941124FFF9F173" masterDocTitle="Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="28" updateTime="1734984611405" updateUser="julia">
|
||||
<mods:mods id="5CA83A87927B11D5A674565DBDF3BE1A" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="4377A146473801298CBF0157441B341D">
|
||||
<mods:title id="05D1A001EC0E1C3459E01231A75213D0">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="36C4042EE7F81A22232C8633AA2DCEA5" type="personal">
|
||||
<mods:role id="E0FB49BCB799347FCA280E86AC2FF57E">
|
||||
<mods:roleTerm id="FA13AFFB56A7B6429F9173ED9F5D3240">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="E8BC98E942DB27EE01C090A4A4C35102">Reyes-Hernández, José L.</mods:namePart>
|
||||
<mods:affiliation id="46D9A93CF1DD813040C03149FF9BACDC">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
|
||||
<mods:nameIdentifier id="3D08DE9C81F5B98F4A9DA103A0F21989" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name id="515238EAE38CEAFDA2C5404D27E5905E" type="personal">
|
||||
<mods:role id="46E5A29EB7A9FA7EEF92490ED4E87C44">
|
||||
<mods:roleTerm id="B9197B70A5FE641EF93FCDE798AC7DBE">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="702487F9FB4858EE4DF115E5198EE529">Hansen, Aslak Kappel</mods:namePart>
|
||||
<mods:affiliation id="0A400FF1F6EDD6C6DE89B10044A8EEB5">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="781EA4334D3FC303D071050CC1F000D1" type="personal">
|
||||
<mods:role id="F7F05CBC09C15A85709B9E0C6D02CB02">
|
||||
<mods:roleTerm id="AE6E5E3C2B4A7A88AC544011C6792863">Author</mods:roleTerm>
|
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</mods:role>
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<emphasis id="B90CEAF7FFBA2E3FFEF11447FE3AF40D" bold="true" box="[357,451,1379,1406]" italics="true" pageId="27" pageNumber="28">variabilis</emphasis>
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group
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Included species:
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<taxonomicName id="4C784D66FFBA2E3FFE821480FE1BF4CF" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[278,482,1444,1468]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">Loncovilius variabilis</taxonomicName>
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<taxonomicNameLabel id="A23F578CFFBA2E3FFE731480FDD7F4C8" box="[487,558,1444,1467]" pageId="27" pageNumber="28" rank="species">sp. nov</taxonomicNameLabel>
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..
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<emphasis id="B90CEAF7FFBA2E3FFFE514FAFF20F486" box="[113,217,1502,1525]" italics="true" pageId="27" pageNumber="28">Diagnosis:</emphasis>
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Head usually elongate, without coarse non-setiferous punctures; nuchal ridge complete dorsally, joined with infraorbital ridge; gula with isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin (
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<figureCitation id="13432A60FFBA2E3FFFE8177FFF3CF700" box="[124,197,1627,1651]" captionStart="Figure 8" captionStartId="16.[129,194,1829,1853]" captionTargetBox="[183,1419,150,1799]" captionTargetId="figure-8@16.[183,1419,144,1799]" captionTargetPageId="16" captionText="Figure 8. A, pronotum of L. carlsbergi. AMRSP = anterior marginal row of setiferous punctures, APP = additional paired punctures adjacent to AMRSP, LLSP = large lateral setiferous puncture, LPSP = large posterior setiferous puncture, PPDS = paired punctures on dorsal serial, SLSP = sublateral setiferous puncture.B, L. cantharoides ventral view of mesotarsomere 3 with pale adhesive setae with terminal plate.C, L. cantharoides ventral view of metatarsomere 4, apical margin emarginated (bilobed), with pale adhesive setae without terminal plate. D, L. cantharoides tergites VI and VII.Black arrow up = anterior transverse basal carina (ATBC) laterally straight continuing to paratergites, white arrow down = ATBC descending laterally without reaching tergite margin.Head in ventral view of Loncovilius species. E, L. hammondi. F, L. lividipennis. PMP = postmandibular puncture, PMR = postmandibular ridge, EM = eye margin." pageId="27" pageNumber="28">Fig. 8F</figureCitation>
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); PMP (usually reduced); females with a11 distinctly longer than a10 (a11:a10 ratio>1.5). Pronotum slightly wider than long, its outline evenly curved. Elytra with even but sparse setiferous punctation on disc and epipleura. Protibiae not sexually dimorphic, in both sexes without laterodorsal row of thick spines. Mesotarsomeres 1–4 not sexually dimorphic, in both sexes with pale adhesive setae and mesotarsomeres
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<quantity id="4C809B00FFBA2E3FFD261633FD03F65C" box="[690,762,1815,1839]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="27" pageNumber="28" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1–3 in</quantity>
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both sexes with terminal plate. Tergites III and IV with several rows of fine punctures of moderate density, without sizeable impunctate area near to posterior margin (
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<figureCitation id="13432A60FFBA2E3FFDB31651FD97F6FE" box="[551,622,1909,1933]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="27" pageNumber="28">Fig. 9B</figureCitation>
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); tergite VIII only slightly emarginate medio-apically in females; ovipositor, its second gonocoxite with one macroseta medially. Aedeagus as in
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<figureCitation id="13432A60FFBA2E3FFCCA118BFC11F1B4" box="[862,1000,175,199]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="27" pageNumber="28">Figure 10I–K</figureCitation>
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.
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||||
<taxonomicName id="4C784D66FFBB2E3EFEB8177AFDF4F70B" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[300,525,1630,1657]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="impunctus" status="sp. nov.">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFEB8177AFDF4F70B" bold="true" box="[300,525,1630,1657]" italics="true" pageId="26" pageNumber="27">Loncovilius impunctus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFFBB2E3EFD85177AFDA4F70B" box="[529,605,1630,1656]" pageId="26" pageNumber="27" rank="species">sp.nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBB2E3EFE8617B8FE15F67B" pageId="26" pageNumber="27" type="description">
|
||||
<paragraph id="8BC736E5FFBB2E3EFE8617B8FD81F7C5" blockId="26.[274,632,1692,1719]" box="[274,632,1692,1719]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFE8617B8FD81F7C5" bold="true" box="[274,632,1692,1719]" pageId="26" pageNumber="27">
|
||||
(
|
||||
<figureCitation id="13432A60FFBB2E3EFE8917B9FE94F7C4" box="[285,365,1693,1719]" captionStart="Figure 5" captionStartId="13.[113,178,1868,1892]" captionTargetBox="[154,1419,145,1840]" captionTargetId="figure-6@13.[153,1420,144,1840]" captionTargetPageId="13" captionText="Figure 5. Female habitus of Loncovilius species.A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis." pageId="26" pageNumber="27">Figs 5C</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBB2E3EFEE317B9FE3CF7C4" box="[375,453,1693,1719]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="26" pageNumber="27">10G, H</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBB2E3EFE5B17B9FE04F7C4" box="[463,509,1693,1719]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="26" pageNumber="27">12C</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBB2E3EFD9317B9FDCCF7C4" box="[519,565,1693,1719]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="26" pageNumber="27">13C</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBB2E3EFDAB17B9FD94F7C4" box="[575,621,1693,1719]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="26" pageNumber="27">14C</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFF1517F5FE15F67B" blockId="26.[129,777,1744,1800]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFF1517F5FE93F79B" box="[129,362,1745,1769]" italics="true" pageId="26" pageNumber="27">Zoobank registration:</emphasis>
|
||||
<uri id="FFE93AE7FFBB2E3EFE1D17F4FE1EF67B" pageId="26" pageNumber="27">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FFDE0C30FFBB2E3EFD1E17F4FE1EF67B" pageId="26" pageNumber="27">45463DC0- 4ECA-4B24-8220-24008B5BE531</uuid>
|
||||
</uri>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBB2E3EFF15160EFBACF0B1" pageId="26" pageNumber="27" type="materials_examined">
|
||||
<paragraph id="8BC736E5FFBB2E3EFF15160EFAA4F010" blockId="26.[127,778,1833,1983]" lastBlockId="26.[824,1474,144,450]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFF15160EFEF6F632" box="[129,271,1833,1857]" italics="true" pageId="26" pageNumber="27">Type material.</emphasis>
|
||||
<materialsCitation id="3B103CB8FFBB2E3EFE80160DFF5FF6EC" collectingDate="1954-07" collectorName="L. Pena" country="Chile" location="Cherquenco" pageId="26" pageNumber="27" specimenCode="FMNH-INS 0000 024 367" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFE80160DFE8DF632" box="[276,372,1833,1857]" italics="true" pageId="26" pageNumber="27">
|
||||
<typeStatus id="54C38847FFBB2E3EFE80160DFE96F632" box="[276,367,1833,1857]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
<specimenCount id="9D7EFD6CFFBB2E3EFEE8160DFE56F632" box="[380,431,1833,1857]" pageId="26" pageNumber="27" type="male">male</specimenCount>
|
||||
, mounted, with genitalia in a separate microvial, with labels as follows:
|
||||
<collectingCountry id="F36F7675FFBB2E3EFD99166DFD99F612" box="[525,608,1865,1889]" name="Chile" pageId="26" pageNumber="27">CHILE</collectingCountry>
|
||||
:
|
||||
<location id="8EA7603EFFBB2E3EFDFD166DFD10F612" LSID="urn:lsid:plazi:treatment:03D187F3FFBB2E3FFEB8177AFD30F437:8EA7603EFFBB2E3EFDFD166DFD10F612" box="[617,745,1865,1889]" country="Chile" name="Cherquenco" pageId="26" pageNumber="27">Cherquenco</location>
|
||||
[~ -38.70, -72.01]
|
||||
<date id="FFC61025FFBB2E3EFE88164CFE83F6F3" box="[284,378,1896,1920]" pageId="26" pageNumber="27" value="1954-07">
|
||||
<collectingDate id="EF82E9CDFFBB2E3EFE88164CFE83F6F3" box="[284,378,1896,1920]" pageId="26" pageNumber="27" value="1954-07">VII-1954</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FFBB2E3EFEEB164CFE32F60C" box="[383,459,1896,1919]" pageId="26" pageNumber="27">L. Pena</collectorName>
|
||||
, leg./♂/
|
||||
<specimenCode id="DBDE9E9EFFBB2E3EFDB7164CFF5FF6EC" pageId="26" pageNumber="27">FMNH-INS 0000 024 367</specimenCode>
|
||||
</materialsCitation>
|
||||
/
|
||||
<typeStatus id="54C38847FFBB2E3EFF3A16A3FEC7F6EC" box="[174,318,1927,1951]" pageId="26" pageNumber="27" type="holotype">HOLOTYPE</typeStatus>
|
||||
<taxonomicName id="4C784D66FFBB2E3EFED016A3FDE5F6EC" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[324,540,1927,1951]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="impunctus">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFED016A3FDE5F6EC" box="[324,540,1927,1951]" italics="true" pageId="26" pageNumber="27">Loncovilius impunctus</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH.
|
||||
<materialsCitation id="3B103CB8FFBB2E3EFC0E11B5FC6CF195" collectingDate="1954-07" collectorName="L. Pena" country="Chile" location="Cherquenco" pageId="26" pageNumber="27" specimenCode="FMNH-INS 0000 024 368-369" specimenCount="1" specimenCount-female="3" typeStatus="paratype">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFC0E11B5FBF9F1DB" box="[922,1024,145,168]" italics="true" pageId="26" pageNumber="27">
|
||||
<typeStatus id="54C38847FFBB2E3EFC0E11B5FC05F1DB" box="[922,1020,145,168]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
<collectingCountry id="F36F7675FFBB2E3EFB9911B4FB99F1DB" box="[1037,1120,144,168]" name="Chile" pageId="26" pageNumber="27">CHILE</collectingCountry>
|
||||
:
|
||||
<location id="8EA7603EFFBB2E3EFBE411B4FB09F1DB" LSID="urn:lsid:plazi:treatment:03D187F3FFBB2E3FFEB8177AFD30F437:8EA7603EFFBB2E3EFBE411B4FB09F1DB" box="[1136,1264,144,168]" country="Chile" name="Cherquenco" pageId="26" pageNumber="27">Cherquenco</location>
|
||||
[~ -38.70, -72.01]
|
||||
<date id="FFC61025FFBB2E3EFCAD118BFC6EF1B4" box="[825,919,175,199]" pageId="26" pageNumber="27" value="1954-07">
|
||||
<collectingDate id="EF82E9CDFFBB2E3EFCAD118BFC6EF1B4" box="[825,919,175,199]" pageId="26" pageNumber="27" value="1954-07">VII-1954</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FFBB2E3EFC091194FC12F1B4" box="[925,1003,176,199]" pageId="26" pageNumber="27">L. Pena</collectorName>
|
||||
, leg./♀/
|
||||
<specimenCode id="DBDE9E9EFFBB2E3EFBD2118BFA72F1B4" box="[1094,1419,175,199]" pageId="26" pageNumber="27">FMNH-INS 0000 024 368-369</specimenCode>
|
||||
[2♀ FMNH]
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B103CB8FFBB2E3EFC3011EBFAECF056" collectingDate="1976-12-19" collectingDateMax="1976-12-21" collectingDateMin="1976-12-19" collectionCode="CNCI" collectorName="H. F. Howden" country="Chile" elevation="1100" location="Malleco Prov." pageId="26" pageNumber="27" specimenCode="FMNH-INS 0000 024 465" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FFBB2E3EFC3011EBFC0EF194" box="[932,1015,207,231]" name="Chile" pageId="26" pageNumber="27">CHILE</collectingCountry>
|
||||
:
|
||||
<location id="8EA7603EFFBB2E3EFB9011EBFB6FF195" LSID="urn:lsid:plazi:treatment:03D187F3FFBB2E3FFEB8177AFD30F437:8EA7603EFFBB2E3EFB9011EBFB6FF195" box="[1028,1174,207,231]" country="Chile" name="Malleco Prov." pageId="26" pageNumber="27">Malleco Prov.</location>
|
||||
<locationDeviation id="9A752FC7FFBB2E3EFB3411EBFA9CF195" box="[1184,1381,207,231]" location="Manzanar" pageId="26" pageNumber="27">
|
||||
<quantity id="4C809B00FFBB2E3EFB3411EBFB22F194" box="[1184,1243,207,231]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="km" value="20.0">20km</quantity>
|
||||
E Manzanar
|
||||
</locationDeviation>
|
||||
[-38.44, -71.51]
|
||||
<quantity id="4C809B00FFBB2E3EFC0411CAFC20F075" box="[912,985,238,262]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="27" unit="m" value="1100.0">
|
||||
<elevation id="0055D1D6FFBB2E3EFC0411CAFC20F075" box="[912,985,238,262]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="27" unit="m" value="1100.0">1100m</elevation>
|
||||
</quantity>
|
||||
<date id="FFC61025FFBB2E3EFC7011CAFB80F075" box="[996,1145,238,262]" pageId="26" pageNumber="27" value="1976-12-19" valueMax="1976-12-21" valueMin="1976-12-19">
|
||||
<collectingDate id="EF82E9CDFFBB2E3EFC7011CAFB80F075" box="[996,1145,238,262]" pageId="26" pageNumber="27" value="1976-12-19" valueMax="1976-12-21" valueMin="1976-12-19">19-21.xii.1976</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FFBB2E3EFB1011CAFADBF075" box="[1156,1314,238,262]" pageId="26" pageNumber="27">H. F. Howden</collectorName>
|
||||
/
|
||||
<collectionCode id="ED69AE20FFBB2E3EFABE11CAFA95F075" box="[1322,1388,238,262]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="26" pageNumber="27">CNCI</collectionCode>
|
||||
L2004- 28/
|
||||
<specimenCount id="9D7EFD6CFFBB2E3EFCF41034FC83F057" box="[864,890,272,292]" count="1" pageId="26" pageNumber="27" type="female">♀</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFFBB2E3EFC161029FB6CF056" box="[898,1173,269,293]" pageId="26" pageNumber="27">FMNH-INS 0000 024 465</specimenCode>
|
||||
[1♀ CNC]
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3B103CB8FFBB2E3EFA8A1029FAA1F010" collectingDate="1962-01-11" collectionCode="FMNH, MNNC" collectorName="Coll. L. Pena" country="Chile" location="Tolhuaco" pageId="26" pageNumber="27" specimenCode="FMNH-INS 0000 024 364-366" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
|
||||
<collectingCountry id="F36F7675FFBB2E3EFA8A1029FAA2F056" box="[1310,1371,269,293]" name="Chile" pageId="26" pageNumber="27">Chile</collectingCountry>
|
||||
:
|
||||
<location id="8EA7603EFFBB2E3EFAF61029FA3BF056" LSID="urn:lsid:plazi:treatment:03D187F3FFBB2E3FFEB8177AFD30F437:8EA7603EFFBB2E3EFAF61029FA3BF056" box="[1378,1474,269,293]" country="Chile" name="Tolhuaco" pageId="26" pageNumber="27">Tolhuaco</location>
|
||||
[~ -38.23, -71.71]
|
||||
<date id="FFC61025FFBB2E3EFC6C1009FBC4F036" box="[1016,1085,301,325]" pageId="26" pageNumber="27" value="1962-01-11">
|
||||
<collectingDate id="EF82E9CDFFBB2E3EFC6C1009FBC4F036" box="[1016,1085,301,325]" pageId="26" pageNumber="27" value="1962-01-11">11.I.62</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<collectorName id="268D5333FFBB2E3EFBD21009FB2EF036" box="[1094,1239,301,325]" pageId="26" pageNumber="27">Coll. L. Peña</collectorName>
|
||||
/
|
||||
<specimenCount id="9D7EFD6CFFBB2E3EFB741014FB03F037" box="[1248,1274,304,324]" count="1" pageId="26" pageNumber="27" type="male">♂</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFFBB2E3EFA961009FC38F017" pageId="26" pageNumber="27">FMNH-INS 0000 024 364–366</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBB2E3EFC5A1068FC01F010" box="[974,1016,332,356]" count="2" pageId="26" pageNumber="27" type="male">2♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBB2E3EFC691068FBB6F010" box="[1021,1103,332,355]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="26" pageNumber="27">FMNH</collectionCode>
|
||||
;
|
||||
<specimenCount id="9D7EFD6CFFBB2E3EFBCC1068FB86F010" box="[1112,1151,332,356]" count="1" pageId="26" pageNumber="27" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBB2E3EFB111068FB2EF017" box="[1157,1239,332,356]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="26" pageNumber="27">MNNC</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FFBB2E3EFB6A1068FAAAF010" box="[1278,1363,332,355]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="26" pageNumber="27">FMNH</collectionCode>
|
||||
]
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFCC1104FFBACF0B1" blockId="26.[824,1474,144,450]" pageId="26" pageNumber="27">
|
||||
All
|
||||
<typeStatus id="54C38847FFBB2E3EFC171048FC1FF0F7" box="[899,998,364,388]" pageId="26" pageNumber="27" type="paratype">paratypes</typeStatus>
|
||||
are supplied with the label: PARATYPE
|
||||
<taxonomicName id="4C784D66FFBB2E3EFCAD10AFFBEFF0D1" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[825,1046,395,419]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="impunctus">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFCAD10AFFBEFF0D1" box="[825,1046,395,419]" italics="true" pageId="26" pageNumber="27">Loncovilius impunctus</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBB2E3EFCAD10C0FAF9F3A5" pageId="26" pageNumber="27" type="diagnosis">
|
||||
<paragraph id="8BC736E5FFBB2E3EFCAD10C0FAF9F3A5" blockId="26.[824,1475,483,727]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFCAD10C0FC66F088" box="[825,927,484,507]" italics="true" pageId="26" pageNumber="27">Diagnosis:</emphasis>
|
||||
Head dorsally with meshed microsculpture; gula with isodiametric microsculpture in the middle; females with antennomere 11 (a11) distinctly longer than antennomere 10 (a10; a11:a10 ratio>1.5). Elytra reddish-brown, disc glossy with only a few rows of setiferous punctures, epipleura evenly setose. Protibiae not sexually dimorphic, in both sexes with laterodorsal row of thick spines. Tergite VIII in both sexes with medial apical emargination. Aedeagus as in
|
||||
<figureCitation id="13432A60FFBB2E3EFBF3139BFB03F3A5" box="[1127,1274,703,727]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="26" pageNumber="27">Figure 10G, H</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBB2E3FFCAD13DDFD68F527" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
|
||||
<paragraph id="8BC736E5FFBB2E3EFCAD13DDFBCEF2DE" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">
|
||||
<emphasis id="B90CEAF7FFBB2E3EFCAD13DDFC49F263" box="[825,944,761,784]" italics="true" pageId="26" pageNumber="27">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFBB2E3EFBF513DFFB82F27C" box="[1121,1147,763,783]" count="1" pageId="26" pageNumber="27" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFBB2E3EFAE713DDFA7FF263" box="[1395,1414,761,784]" italics="true" pageId="26" pageNumber="27">N</emphasis>
|
||||
= 3]: FBL = 3.35–3.59 (3.48); HW = 1.01–1.1 (1.07); HL = 0.85– 0.93 (0.89); HW/HL = 1.18–1.21 (1.19); PW = 1.4– 1.44 (1.42); PL = 1.2–1.25 (1.23); PW/PL = 1.14–1.17 (1.15); EW = 1.67–1.7 (1.68); EL = 1.3–1.44 (1.36); PW/ HW = 1.28–1.39 (1.33).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFCC11290FB43F51A" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFBB2E3EFC641293FBF3F2B8" box="[1008,1034,951,971]" count="1" pageId="26" pageNumber="27" type="female">♀</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFBB2E3EFB7E1291FB04F2BF" box="[1258,1277,949,972]" italics="true" pageId="26" pageNumber="27">N</emphasis>
|
||||
= 4]: FBL = 3.71– 3.93 (3.81); TL = 7.5–7.53 (7.52); HW = 1.17–1.18 (1.18); HL = 0.93–0.97 (0.95); HW/HL = 1.22–1.26 (1.24); PW = 1.54–1.59 (1.56); PL = 1.29–1.37 (1.32); PW/ PL = 1.16–1.19 (1.18); EW = 1.82–1.9 (1.87); EL = 1.46–1.59 (1.54); PW/HW = 1.32–1.35 (1.33).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFCC11554FC78F5B4" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">Head, mouthparts, and legs black to brown-black; pronotum, antennae, elytra, and abdomen dark reddish-brown, elytra lighter.</paragraph>
|
||||
<paragraph id="8BC736E5FFBB2E3EFCC115EAFBA0F6C4" blockId="26.[824,1476,760,1975]" pageId="26" pageNumber="27">
|
||||
Head transverse; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse meshes, with coarse non-setiferous punctures; posterior angles indistinct with few setiferous punctures of medium and small size. Eyes of medium size (EYL/HL = x = 0.52), from 1.51 to 1.64 times longer than temples (in lateral view); distance between eyes around 1.8 times the length of eye. Antennal insertions situated at equal distance from frontoclypeus and from eye; antennomeres 2 and 3 subequal in length; antennomeres 4 to 10 gradually reduced in length; a11 from 1.54 to 1.75 times longer than a
|
||||
<quantity id="4C809B00FFBB2E3EFAD914CCFA78F773" box="[1357,1409,1512,1536]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="26" pageNumber="27" unit="in" value="10.0">10 in</quantity>
|
||||
males and from
|
||||
<quantity id="4C809B00FFBB2E3EFC331723FBBEF76C" box="[935,1095,1543,1567]" metricMagnitude="-2" metricUnit="m" metricValue="4.0893999999999995" metricValueMax="4.2672" metricValueMin="3.9116" pageId="26" pageNumber="27" unit="in" value="1.6099999999999999" valueMax="1.68" valueMin="1.54">1.54 to 1.68 in</quantity>
|
||||
females. Basal and parocular punctures usually single; posterior frontal puncture located anterior to temporal puncture; without small setose punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge rather straight in L-shape, almost united with gular sutures; postgenal ridge absent; postmandibular ridge distinctly separated from eye margin; well-developed PMP; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula with microsculpture medially, gular sutures moderately separated. Mandibles with dorsolateral groove; labrum with subrectangular sclerotized region and emarginate apical margin; mentum with seta beta only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than penultimate and more or less subconical.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBA2E3FFF1911B4FDD2F036" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">Pronotum transverse, convex, evenly curved, with one PPDS and an APP, without SLSP, with few coarse non-setiferous punctures; flexible postcoxal hypomeral extension (process) interrupted by inferior line. Elytra with epipleura evenly setose; disc with only a few rows of setiferous punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
|
||||
<paragraph id="8BC736E5FFBA2E3FFF191068FF39F34C" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">Protibiae not sexually dimorphic, with laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrallywith apical margin slightly sinuate.</paragraph>
|
||||
<paragraph id="8BC736E5FFBA2E3FFF191363FD9CF2EB" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">
|
||||
Abdominal tergites III and IV with two patches of punctures of moderate size and density, with middle impunctate line; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina strongly arcuate; lateral tergal sclerites IX laterally flattened only in males. Male sternite VIII with broad V-shaped emargination medially (
|
||||
<figureCitation id="13432A60FFBA2E3FFEE713C7FE29F388" box="[371,464,739,763]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="27" pageNumber="28">Fig. 14C</figureCitation>
|
||||
); female sternite VIII with U-shaped emargination medially; male sternite IX with obtuse emargination medially, its basal portion shorter than distal portion; male tergite X truncate medially (
|
||||
<figureCitation id="13432A60FFBA2E3FFD941265FDAEF22A" box="[512,599,833,857]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="27" pageNumber="28">Fig. 12C</figureCitation>
|
||||
); female tergite X truncate to slightly arcuate medially (
|
||||
<figureCitation id="13432A60FFBA2E3FFD961245FDA0F20A" box="[514,601,865,889]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="27" pageNumber="28">Fig. 13C</figureCitation>
|
||||
); ovipositor, its second gonocoxite without macrosetae medially.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBA2E3FFF1912BBFD68F527" blockId="27.[113,764,144,1108]" pageId="27" pageNumber="28">
|
||||
Aedeagus as in
|
||||
<figureCitation id="13432A60FFBA2E3FFEA612BBFE35F2C4" box="[306,460,927,951]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="27" pageNumber="28">Figure 10G, H</figureCitation>
|
||||
; its total length 1.2. Median lobe bulky, in parameral view with truncated apex; internal sac with pair of long I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging into M-shape apex, with dark peg setae, in lateral view almost straight and slightly concave apically.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBA2E3FFFE51551FDD3F5BF" pageId="27" pageNumber="28" type="etymology">
|
||||
<paragraph id="8BC736E5FFBA2E3FFFE51551FDD3F5BF" blockId="27.[113,762,1141,1228]" pageId="27" pageNumber="28">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFFE51551FF18F5FE" box="[113,225,1141,1165]" italics="true" pageId="27" pageNumber="28">Etymology:</emphasis>
|
||||
The species epithet is derived from the Latin prefix ‘
|
||||
<emphasis id="B90CEAF7FFBA2E3FFFE315B1FF68F5DF" box="[119,145,1173,1196]" italics="true" pageId="27" pageNumber="28">im</emphasis>
|
||||
’ (not) and adjective ‘
|
||||
<emphasis id="B90CEAF7FFBA2E3FFEFA15B1FE38F5DF" box="[366,449,1173,1196]" italics="true" pageId="27" pageNumber="28">punctate</emphasis>
|
||||
’ (dotted), referring to the disc of pronotum and elytra mostly impunctate.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBA2E3FFFE515C9FD30F437" pageId="27" pageNumber="28" type="distribution">
|
||||
<paragraph id="8BC736E5FFBA2E3FFFE515C9FD30F437" blockId="27.[112,762,1261,1348]" pageId="27" pageNumber="28">
|
||||
<emphasis id="B90CEAF7FFBA2E3FFFE515C9FDC7F476" box="[113,574,1261,1285]" italics="true" pageId="27" pageNumber="28">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FFBA2E3FFEF315C9FDC7F476" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[359,574,1261,1285]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="impunctus">Loncovilius impunctus</taxonomicName>
|
||||
</emphasis>
|
||||
is known from the
|
||||
<collectingRegion id="49BCF807FFBA2E3FFFE41429FEDFF456" box="[112,294,1293,1317]" country="Chile" name="Araucania" pageId="27" pageNumber="28">Araucanía Region</collectingRegion>
|
||||
in
|
||||
<collectingCountry id="F36F7675FFBA2E3FFED01429FE85F456" box="[324,380,1293,1317]" name="Chile" pageId="27" pageNumber="28">Chile</collectingCountry>
|
||||
where it is confined to Valdivian temperate forest ecoregion. It is unknown how it was collected.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,110 +0,0 @@
|
|||
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<mods:title id="6086130BC013FBA690A06CC0B1898CDD">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="C41A1C90C6EC55BE26F0A9AD40C0473A">Reyes-Hernández, José L.</mods:namePart>
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<mods:namePart id="E1D4EDB605D12A82CDD9D7F3AB9411C7">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="B92B9A4DC29AA01661480532587DAD07">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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<mods:namePart id="23357EE258B2B7A83D7E0BAC50751D9A">Solodovnikov, Alexey</mods:namePart>
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<mods:affiliation id="575C26E879853D27DD20F790C4D2745B">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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|
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<taxonomicName id="4C784D66FFBC2E39FEDB15AAFE21F5DA" ID-CoL="3VYHP" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[335,472,1166,1193]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBC2E39FEDB15AAFE21F5DA" bold="true" box="[335,472,1166,1193]" italics="true" pageId="29" pageNumber="30">edwardsianus</emphasis>
|
||||
</taxonomicName>
|
||||
group
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBC2E39FFE515EBFE6AF475" pageId="29" pageNumber="30" type="taxon_list">
|
||||
<paragraph id="8BC736E5FFBC2E39FFE515EBFE6AF475" blockId="29.[113,763,1231,1286]" pageId="29" pageNumber="30">
|
||||
<emphasis id="B90CEAF7FFBC2E39FFE515EBFDD5F594" box="[113,556,1231,1255]" italics="true" pageId="29" pageNumber="30">
|
||||
Included species:
|
||||
<taxonomicName id="4C784D66FFBC2E39FEA415EBFDD5F594" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[304,556,1231,1255]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cantharoides" status="sp. nov.">Loncovilius cantharoides</taxonomicName>
|
||||
</emphasis>
|
||||
<taxonomicNameLabel id="A23F578CFFBC2E39FDA915EBFD61F595" box="[573,664,1231,1254]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, and
|
||||
<taxonomicName id="4C784D66FFBC2E39FD7215EBFE76F475" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBC2E39FD7215EBFD0CF595" box="[742,757,1231,1254]" italics="true" pageId="29" pageNumber="30">L</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FFBC2E39FFE515CAFF0DF475" box="[113,244,1262,1286]" italics="true" pageId="29" pageNumber="30">edwardsianus</emphasis>
|
||||
(Korge, 1963)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBC2E39FFE5140FFE13F72E" pageId="29" pageNumber="30" type="diagnosis">
|
||||
<paragraph id="8BC736E5FFBC2E39FFE5140FFE13F72E" blockId="29.[113,764,1323,1629]" pageId="29" pageNumber="30">
|
||||
<emphasis id="B90CEAF7FFBC2E39FFE5140FFF2EF431" box="[113,215,1323,1346]" italics="true" pageId="29" pageNumber="30">Diagnosis:</emphasis>
|
||||
Head elongate, without coarse non-setiferous punctures; nuchal and infraorbital ridges not joined; gula with isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin; PMP strongly reduced or rarely absent; females with a11 slightly longer than a10 (a11:a10 ratio <1.5). Pronotum without SLSP. Tergites III and IV with only a few rows of coarse, sparse punctures, with large impunctate area near apical margin (
|
||||
<figureCitation id="13432A60FFBC2E39FEAB1722FE73F76D" box="[319,394,1542,1566]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="29" pageNumber="30">Fig. 9C</figureCitation>
|
||||
); tergite VIII without medial apical emargination; ovipositor, its second gonocoxite with three or more macrosetae medially (
|
||||
<figureCitation id="13432A60FFBC2E39FE191761FE20F72E" box="[397,473,1605,1629]" captionStart="Figure 9" captionStartId="17.[113,178,1867,1891]" captionTargetBox="[153,1420,143,1718]" captionTargetId="figure-7@17.[153,1420,143,1839]" captionTargetPageId="17" captionText="Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus." pageId="29" pageNumber="30">Fig. 9H</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,812 +0,0 @@
|
|||
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<mods:mods id="0C9FADEB22DA2B1A6B787FE3F007B06A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="ABF8475C551F70EEDD87C519A3172249">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="3ACC5FAB55D95E67352C4D92575EA5E7">Reyes-Hernández, José L.</mods:namePart>
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<mods:affiliation id="790328C71C383A6F06100CFA2D6C9B20">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="A9A2018B60652501896724A5C2D2DD44" type="email">jl.reyeshdez@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2EFBFF2A563D08D0B0BF70B5250E0128">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="48355B99F8C01DB229BC72B91AFBC7B0">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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<mods:namePart id="59B9CF1EA3B774D21052E9A01681BFE7">Shaw, Josh Jenkins</mods:namePart>
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<mods:affiliation id="C9A77CFF6167243CFAF4B3A2CBCA982A">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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</mods:role>
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<mods:namePart id="D62148151ABF1EEBA71E512B7CBF39EB">Solodovnikov, Alexey</mods:namePart>
|
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<mods:affiliation id="08041533C64C222C11E7E8E2EE5D779A">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark</mods:affiliation>
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<mods:date id="5D3E6090A8BF3D37A1E1C2035D2156D9">2024</mods:date>
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<mods:title id="A7EB71BD399E0B642023C7366E728E29">Zool. J. Linn. Soc.</mods:title>
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||||
<treatment id="03D187F3FFBC2E3AFE8617A3FAFCF4B2" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2" httpUri="http://treatment.plazi.org/id/03D187F3FFBC2E3AFE8617A3FAFCF4B2" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
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|
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<paragraph id="8BC736E5FFBC2E39FE8617A3FDA3F7D1" blockId="29.[274,602,1671,1698]" box="[274,602,1671,1698]" pageId="29" pageNumber="30">
|
||||
<heading id="D08F8189FFBC2E39FE8617A3FDA3F7D1" bold="true" box="[274,602,1671,1698]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
|
||||
<emphasis id="B90CEAF7FFBC2E39FE8617A3FDA3F7D1" bold="true" box="[274,602,1671,1698]" pageId="29" pageNumber="30">
|
||||
<taxonomicName id="4C784D66FFBC2E39FE8617A3FDF0F7D1" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[274,521,1671,1698]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cantharoides" status="sp. nov.">
|
||||
<emphasis id="B90CEAF7FFBC2E39FE8617A3FDF0F7D1" bold="true" box="[274,521,1671,1698]" italics="true" pageId="29" pageNumber="30">Loncovilius cantharoides</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFFBC2E39FD9A17ACFDA3F7D1" box="[526,602,1672,1698]" pageId="29" pageNumber="30" rank="species">sp.nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBC2E39FF6317E2FE04F64D" pageId="29" pageNumber="30" type="description">
|
||||
<paragraph id="8BC736E5FFBC2E39FF6317E2FD8CF793" blockId="29.[247,629,1734,1760]" box="[247,629,1734,1760]" pageId="29" pageNumber="30">
|
||||
<emphasis id="B90CEAF7FFBC2E39FF6317E2FD8CF793" bold="true" box="[247,629,1734,1760]" pageId="29" pageNumber="30">
|
||||
(
|
||||
<figureCitation id="13432A60FFBC2E39FE9617E2FEB6F793" box="[258,335,1734,1760]" captionStart="Figure 6" captionStartId="14.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@14.[168,1434,144,1840]" captionTargetPageId="14" captionText="Figure 6. Habitus of Loncovilius species. A, male of L. edwardsianus. B, male of L. cantharoides. C, female of L. semiflavus. D, male of L. barclayi." pageId="29" pageNumber="30">Figs 6B</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBC2E39FECD17E2FE8DF793" box="[345,372,1734,1760]" captionStart="Figure 7" captionStartId="15.[113,178,1868,1892]" captionTargetBox="[153,1419,163,1840]" captionTargetId="figure-6@15.[153,1419,144,1840]" captionTargetPageId="15" captionText="Figure 7. Female habitus of Loncovilius species.A, L. lividipennis. B, L. carlsbergi. Head. C, L. hammondi. D, L. semiflavus. E, L. carlsbergi. F, Male mesotarsi of L. cantharoides." pageId="29" pageNumber="30">7F</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBC2E39FEEA17E2FE2BF793" box="[382,466,1734,1760]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="29" pageNumber="30">10O–Q</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBC2E39FE4817E2FDFCF793" box="[476,517,1734,1760]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="29" pageNumber="30">12F</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBC2E39FD9A17E2FDCEF793" box="[526,567,1734,1760]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="29" pageNumber="30">13F</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBC2E39FDD417E2FD93F793" box="[576,618,1734,1760]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="29" pageNumber="30">14F</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBC2E39FFE51623FE04F64D" blockId="29.[113,762,1798,1854]" pageId="29" pageNumber="30">
|
||||
<emphasis id="B90CEAF7FFBC2E39FFE51623FE99F66D" box="[113,352,1799,1823]" italics="true" pageId="29" pageNumber="30">Zoobank registration:</emphasis>
|
||||
<uri id="FFE93AE7FFBC2E39FE171622FE00F64D" pageId="29" pageNumber="30">
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||||
urn:lsid:zoobank.org:act:
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||||
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|
||||
</uri>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBC2E3AFFE51644FF42F003" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="materials_examined">
|
||||
<paragraph id="8BC736E5FFBC2E3AFFE51644FD2CF062" blockId="29.[113,762,1887,1974]" lastBlockId="30.[128,780,155,368]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||||
<emphasis id="B90CEAF7FFBC2E39FFE51644FEFCF604" box="[113,261,1887,1911]" italics="true" pageId="29" pageNumber="30">Type material.</emphasis>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FE9B167BFBA8F056" collectingDate="1982-12-13" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="NHMD" collectorName="Km E Malalcahuello & A. Newton & M. Thayer" country="Chile" determinerName="J. Jenkins Shaw" elevation="1080" location="Malleco Prov." municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="916904" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="B90CEAF7FFBC2E39FE9B167BFE96F604" box="[271,367,1887,1911]" italics="true" pageId="29" pageNumber="30">
|
||||
<typeStatus id="54C38847FFBC2E39FE9B167BFE93F604" box="[271,362,1887,1911]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
male, mounted, with labels as follows:
|
||||
<collectingCountry id="F36F7675FFBC2E39FF24165BFEFAF6E4" box="[176,259,1919,1943]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<location id="8EA7603EFFBC2E39FE9A165BFE67F6E5" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FE9A165BFE67F6E5" box="[270,414,1919,1943]" country="Chile" municipality="Malalcahuello" name="Malleco Prov." pageId="29" pageNumber="30">Malleco Prov.</location>
|
||||
6.5
|
||||
<collectorName id="268D5333FFBC2E39FE59165BFD5AF6E4" box="[461,675,1919,1943]" pageId="29" pageNumber="30">Km E Malalcahuello</collectorName>
|
||||
<quantity id="4C809B00FFBC2E39FD38165BFD0FF6E4" box="[684,758,1919,1943]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="29" pageNumber="30" unit="m" value="1080.0">
|
||||
<elevation id="0055D1D6FFBC2E39FD38165BFD0FF6E4" box="[684,758,1919,1943]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="29" pageNumber="30" unit="m" value="1080.0">1080m</elevation>
|
||||
</quantity>
|
||||
, trap site 651
|
||||
<date id="FFC61025FFBC2E39FE9616BAFE6EF6C5" box="[258,407,1950,1974]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FE9616BAFE6EF6C5" box="[258,407,1950,1974]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">13-31.xii.1982</collectingDate>
|
||||
</date>
|
||||
North. Dombeyi with Chusquea
|
||||
<collectorName id="268D5333FFBC2E39FCBE11B4FC5AF1D4" box="[810,931,144,168]" pageId="29" pageNumber="30">A. Newton</collectorName>
|
||||
and
|
||||
<collectorName id="268D5333FFBC2E39FC7711B4FBA8F1DB" box="[995,1105,144,168]" pageId="29" pageNumber="30">M. Thayer</collectorName>
|
||||
/window trap/
|
||||
<taxonomicName id="4C784D66FFBC2E39FB6411B4FA63F1DB" authority="Sp." authorityName="Sp." box="[1264,1434,144,168]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Loncovilius Sp.</taxonomicName>
|
||||
5 Solodovnikov det. 2002/
|
||||
<taxonomicName id="4C784D66FFBC2E39FBA21194FA78F1B4" authority="J. Jenkins Shaw" authorityName="J. Jenkins Shaw" box="[1078,1409,175,199]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cantharoides">
|
||||
<emphasis id="B90CEAF7FFBC2E39FBA21194FBBCF1B4" box="[1078,1093,176,199]" italics="true" pageId="29" pageNumber="30">L</emphasis>
|
||||
.
|
||||
<emphasis id="B90CEAF7FFBC2E39FBC2118BFB28F1B4" box="[1110,1233,175,199]" italics="true" pageId="29" pageNumber="30">cantharoides</emphasis>
|
||||
<determinerName id="7E1BDA21FFBC2E39FB481194FA78F1B4" box="[1244,1409,175,199]" pageId="29" pageNumber="30">J. Jenkins Shaw</determinerName>
|
||||
</taxonomicName>
|
||||
det. 2016/
|
||||
<collectionCode id="ED69AE20FFBC2E39FCFD11EBFC47F194" box="[873,958,207,231]" collectionName="NHMD" pageId="29" pageNumber="30">NHMD</collectionCode>
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FC5F11EBFBE2F194" box="[971,1051,207,231]" pageId="29" pageNumber="30">916904</specimenCode>
|
||||
/HOLOTYPE
|
||||
<taxonomicName id="4C784D66FFBC2E39FB2A11EBFA4AF194" box="[1214,1459,207,231]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cantharoides">
|
||||
<emphasis id="B90CEAF7FFBC2E39FB2A11EBFA4AF194" box="[1214,1459,207,231]" italics="true" pageId="29" pageNumber="30">Loncovilius cantharoides</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNM.
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FBCC102AFB14F0B2" collectingDate="1982-12-13" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="FMNH" collectorName="A. Newton & M. Thayer" country="Chile" county="Malleco Prov" elevation="1080" location="Malleco Prov" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="FMNH-INS 0000 023 458-463, 465-466, 467-471, 472-474, 476-477, 478, 481, 484-485" specimenCount="22" specimenCount-female="9" specimenCount-male="13" typeStatus="paratype">
|
||||
<emphasis id="B90CEAF7FFBC2E39FBCC102AFB47F056" box="[1112,1214,270,293]" italics="true" pageId="29" pageNumber="30">
|
||||
<typeStatus id="54C38847FFBC2E39FBCC102AFB43F056" box="[1112,1210,270,293]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
<collectingCountry id="F36F7675FFBC2E39FB531029FAE3F056" box="[1223,1306,269,293]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingCounty id="62A64E69FFBC2E39FAB01029FA49F056" box="[1316,1456,269,293]" pageId="29" pageNumber="30">Malleco Prov</collectingCounty>
|
||||
.
|
||||
<quantity id="4C809B00FFBC2E39FCBE1008FC8BF036" box="[810,882,300,325]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="29" pageNumber="30" unit="km" value="6.5">
|
||||
<locationDeviation id="9A752FC7FFBC2E39FCBE1008FC8BF036" box="[810,882,300,325]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="29" pageNumber="30" unit="km" value="6.5">6.5 km</locationDeviation>
|
||||
</quantity>
|
||||
E Malalcahuello [-38.46, -71.51],
|
||||
<quantity id="4C809B00FFBC2E39FB431009FAD8F036" box="[1239,1313,301,325]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="29" pageNumber="30" unit="m" value="1080.0">
|
||||
<elevation id="0055D1D6FFBC2E39FB431009FAD8F036" box="[1239,1313,301,325]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="29" pageNumber="30" unit="m" value="1080.0">1080m</elevation>
|
||||
</quantity>
|
||||
, trap site 651,
|
||||
<date id="FFC61025FFBC2E39FCB81068FC39F017" box="[812,960,332,356]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FCB81068FC39F017" box="[812,960,332,356]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">13-31.xii.1982</collectingDate>
|
||||
</date>
|
||||
<taxonomicName id="4C784D66FFBC2E39FC461068FB83F017" box="[978,1146,332,356]" pageId="29" pageNumber="30" rank="species" species="dombeyi">Noth. dombeyi</taxonomicName>
|
||||
[
|
||||
<taxonomicName id="4C784D66FFBC2E39FB011068FA90F017" box="[1173,1385,332,356]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="dombeyi">
|
||||
<emphasis id="B90CEAF7FFBC2E39FB011068FA90F017" box="[1173,1385,332,356]" italics="true" pageId="29" pageNumber="30">Nothofagus dombeyi</emphasis>
|
||||
</taxonomicName>
|
||||
] with Chusquea
|
||||
<collectorName id="268D5333FFBC2E39FC0C1048FBFAF0F7" box="[920,1027,364,388]" pageId="29" pageNumber="30">A.Newton</collectorName>
|
||||
and
|
||||
<collectorName id="268D5333FFBC2E39FBA3104FFB6EF0F0" box="[1079,1175,363,387]" pageId="29" pageNumber="30">M.Thayer</collectorName>
|
||||
/window trap/
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FAA4104FFC04F0D1" pageId="29" pageNumber="30">FMNH-INS 0000 023 458–463</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FB9810AEFB90F0D1" box="[1036,1129,394,418]" pageId="29" pageNumber="30">465–466</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FBED10AFFB2EF0D0" box="[1145,1239,395,419]" pageId="29" pageNumber="30">467–471</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FB7210AFFABDF0D0" box="[1254,1348,395,419]" pageId="29" pageNumber="30">472–474</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FAC710AFFA48F0D0" box="[1363,1457,395,419]" pageId="29" pageNumber="30">476–477</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FCBE108EFCAAF0B1" box="[810,851,426,450]" pageId="29" pageNumber="30">478</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FCF1108EFC77F0B1" box="[869,910,426,450]" pageId="29" pageNumber="30">481</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FC34108EFC05F0B1" box="[928,1020,426,450]" pageId="29" pageNumber="30">484–485</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FB86108EFBB0F0B2" box="[1042,1097,426,450]" count="13" pageId="29" pageNumber="30" type="male">13♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FBCC108EFB86F0B2" box="[1112,1151,426,450]" count="9" pageId="29" pageNumber="30" type="female">9♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FB1A108EFB1DF0B2" box="[1166,1252,426,449]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="29" pageNumber="30">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FB68108EFC73F32D" collectingDate="1984-12-12" collectingDateMax="1985-02-16" collectingDateMin="1984-12-12" collectionCode="FMNH" collectorName="Km E Malalcahuello" country="Chile" county="Malleco Prov" elevation="1500" location="Curacautin" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="FMNH-INS 0000 023 490-532, 602-613" specimenCount="59" specimenCount-female="24" specimenCount-male="35" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FB68108EFAB6F0B1" box="[1276,1359,426,450]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingCounty id="62A64E69FFBC2E39FAF4108EFCA5F092" pageId="29" pageNumber="30">Malleco Prov</collectingCounty>
|
||||
.,
|
||||
<quantity id="4C809B00FFBC2E39FCF810EDFC56F092" box="[876,943,457,481]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="km" value="40.0">
|
||||
<locationDeviation id="9A752FC7FFBC2E39FCF810EDFC56F092" box="[876,943,457,481]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="km" value="40.0">40 km</locationDeviation>
|
||||
</quantity>
|
||||
W [
|
||||
<quantity id="4C809B00FFBC2E39FC4B10EDFBDFF092" box="[991,1062,457,481]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="km" value="40.0">
|
||||
<locationDeviation id="9A752FC7FFBC2E39FC4B10EDFBDFF092" box="[991,1062,457,481]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="km" value="40.0">40 km</locationDeviation>
|
||||
</quantity>
|
||||
E]
|
||||
<location id="8EA7603EFFBC2E39FBC410EDFB3BF092" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FBC410EDFB3BF092" box="[1104,1218,457,481]" country="Chile" county="Malleco Prov" municipality="Malalcahuello" name="Curacautin" pageId="29" pageNumber="30">Curacautin</location>
|
||||
[-38.44, -71.50],
|
||||
<quantity id="4C809B00FFBC2E39FAEB10EDFCB9F373" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">
|
||||
<elevation id="0055D1D6FFBC2E39FAEB10EDFCB9F373" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">1500 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFC61025FFBC2E39FCCD10CDFC2CF372" box="[857,981,489,513]" pageId="29" pageNumber="30" value="1984-12-12">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FCCD10CDFC2CF372" box="[857,981,489,513]" pageId="29" pageNumber="30" value="1984-12-12">12.XII.1984</collectingDate>
|
||||
</date>
|
||||
-
|
||||
<date id="FFC61025FFBC2E39FB9310CCFB97F373" box="[1031,1134,488,512]" pageId="29" pageNumber="30" value="1985-02-16">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FB9310CCFB97F373" box="[1031,1134,488,512]" pageId="29" pageNumber="30" value="1985-02-16">16.II.1985</collectingDate>
|
||||
</date>
|
||||
/FMHD#85-905,
|
||||
<emphasis id="B90CEAF7FFBC2E39FAAF10CDFA4AF373" box="[1339,1459,489,513]" italics="true" pageId="29" pageNumber="30">
|
||||
<taxonomicName id="4C784D66FFBC2E39FAAF10CDFA52F372" box="[1339,1451,489,513]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
|
||||
-
|
||||
</emphasis>
|
||||
<taxonomicName id="4C784D66FFBC2E39FCBE132CFBFCF353" authority=", S. Peck, P" authorityName="S. Peck, P" box="[810,1029,520,544]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="B90CEAF7FFBC2E39FCBE132CFC75F36C" box="[810,908,520,543]" italics="true" pageId="29" pageNumber="30">Araucaria</emphasis>
|
||||
, S. Peck, P
|
||||
</taxonomicName>
|
||||
#85-19, Malaise FIELD MUSEUM NAT HIST/
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FCFA1303FB3FF34C" box="[878,1222,551,575]" pageId="29" pageNumber="30">FMNH-INS 0000 023 490–532</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FB461303FAD4F34C" box="[1234,1325,551,575]" pageId="29" pageNumber="30">602–613</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FAAA1303FA8CF34D" box="[1342,1397,551,575]" count="35" pageId="29" pageNumber="30" type="male">35♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FAEB1303FA4AF34D" box="[1407,1459,551,575]" count="24" pageId="29" pageNumber="30" type="female">24♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FCBE1363FC79F32D" box="[810,896,583,606]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="29" pageNumber="30">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FC1A1363FB00F3A8" collectingDate="1982-12-13" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="FMNH" collectorName="A. Newton & M. Thayer" country="Chile" county="Malleco Prov" elevation="1570" location="E Malalcahuello" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="FMNH-INS 0000 023 475, 478, 480, 482, 486-489" specimenCount="8" specimenCount-female="5" specimenCount-male="3" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FC1A1363FC18F32C" box="[910,993,583,607]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingCounty id="62A64E69FFBC2E39FC7C1363FB89F32D" box="[1000,1136,583,607]" pageId="29" pageNumber="30">Malleco Prov</collectingCounty>
|
||||
.
|
||||
<quantity id="4C809B00FFBC2E39FBE31363FB4FF32C" box="[1143,1206,583,607]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="km" value="14.0">
|
||||
<locationDeviation id="9A752FC7FFBC2E39FBE31363FB4FF32C" box="[1143,1206,583,607]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="km" value="14.0">14 km</locationDeviation>
|
||||
</quantity>
|
||||
<location id="8EA7603EFFBC2E39FB2E1363FAA4F32C" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FB2E1363FAA4F32C" box="[1210,1373,583,607]" country="Chile" county="Malleco Prov" municipality="Malalcahuello" name="E Malalcahuello" pageId="29" pageNumber="30">E Malalcahuello</location>
|
||||
[-38.43, -71.48]
|
||||
<quantity id="4C809B00FFBC2E39FC141342FC33F30D" box="[896,970,614,638]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="29" pageNumber="30" unit="m" value="1570.0">
|
||||
<elevation id="0055D1D6FFBC2E39FC141342FC33F30D" box="[896,970,614,638]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="29" pageNumber="30" unit="m" value="1570.0">1570m</elevation>
|
||||
</quantity>
|
||||
, trap site 649,
|
||||
<date id="FFC61025FFBC2E39FBE71342FAFEF30D" box="[1139,1287,614,638]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FBE71342FAFEF30D" box="[1139,1287,614,638]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">13-31.xii.1982</collectingDate>
|
||||
</date>
|
||||
Noth.
|
||||
<taxonomicName id="4C784D66FFBC2E39FCBE13A1FC75F3EE" box="[810,908,645,669]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">pumilio-Araucaria</taxonomicName>
|
||||
f. [
|
||||
<emphasis id="B90CEAF7FFBC2E39FC2D13A1FB10F3EE" box="[953,1257,645,669]" italics="true" pageId="29" pageNumber="30">
|
||||
<taxonomicName id="4C784D66FFBC2E39FC2D13A1FB87F3EE" box="[953,1150,645,669]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="pumilio">Nothofagus pumilio</taxonomicName>
|
||||
-
|
||||
<taxonomicName id="4C784D66FFBC2E39FB1313A2FB10F3EE" box="[1159,1257,646,669]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">Araucaria</taxonomicName>
|
||||
</emphasis>
|
||||
forest],
|
||||
<collectorName id="268D5333FFBC2E39FADC13A2FA4AF3ED" box="[1352,1459,646,670]" pageId="29" pageNumber="30">A.Newton</collectorName>
|
||||
and
|
||||
<collectorName id="268D5333FFBC2E39FCCF1381FC43F3CE" box="[859,954,677,701]" pageId="29" pageNumber="30">M.Thayer</collectorName>
|
||||
/window trap 649/
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FB1C1381FA56F3CF" box="[1160,1455,676,701]" pageId="29" pageNumber="30">FMNH-INS 0000 023 475</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FCBE13E0FCAAF3AF" box="[810,851,708,732]" pageId="29" pageNumber="30">478</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FCCB13E0FC71F3AF" box="[863,904,708,732]" pageId="29" pageNumber="30">480</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FC0113E0FC47F3AF" box="[917,958,708,732]" pageId="29" pageNumber="30">482</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FC5E13E0FBDFF3AF" box="[970,1062,708,732]" pageId="29" pageNumber="30">486–489</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FBA313E0FB99F3A8" box="[1079,1120,708,732]" count="3" pageId="29" pageNumber="30" type="male">3♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FBFD13E0FB69F3A8" box="[1129,1168,708,732]" count="5" pageId="29" pageNumber="30" type="female">5♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FB0D13E0FB16F3A8" box="[1177,1263,708,731]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="29" pageNumber="30">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FA9613E0FC0EF22A" collectingDate="1982-12-13" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="FMNH" collectorName="A. Newton & M. Thayer" country="Chile" county="Malleco Prov" elevation="1350" location="E Malacahuello" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="FMNH-INS 0000 023 595" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FA9613E0FAACF3AF" box="[1282,1365,708,732]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingCounty id="62A64E69FFBC2E39FAF413E0FCA2F388" pageId="29" pageNumber="30">Malleco Prov</collectingCounty>
|
||||
.
|
||||
<quantity id="4C809B00FFBC2E39FCF313C7FC5BF388" box="[871,930,739,763]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="km" value="12.0">
|
||||
<locationDeviation id="9A752FC7FFBC2E39FCF313C7FC5BF388" box="[871,930,739,763]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="km" value="12.0">12km</locationDeviation>
|
||||
</quantity>
|
||||
<location id="8EA7603EFFBC2E39FC3F13C0FBB4F388" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FC3F13C0FBB4F388" box="[939,1101,739,763]" country="Chile" county="Malleco Prov" municipality="Malalcahuello" name="E Malacahuello" pageId="29" pageNumber="30">E Malacahuello</location>
|
||||
[-38.43, -71.49]
|
||||
<quantity id="4C809B00FFBC2E39FA9213C7FAA9F388" box="[1286,1360,739,763]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="29" pageNumber="30" unit="m" value="1350.0">
|
||||
<elevation id="0055D1D6FFBC2E39FA9213C7FAA9F388" box="[1286,1360,739,763]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="29" pageNumber="30" unit="m" value="1350.0">1350m</elevation>
|
||||
</quantity>
|
||||
, trap site 650
|
||||
<date id="FFC61025FFBC2E39FCCD1227FC14F268" box="[857,1005,771,795]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FCCD1227FC14F268" box="[857,1005,771,795]" pageId="29" pageNumber="30" value="1982-12-13" valueMax="1982-12-31" valueMin="1982-12-13">13-31.xii.1982</collectingDate>
|
||||
</date>
|
||||
Noth. dombeyi-Araucaria f.
|
||||
<collectorName id="268D5333FFBC2E39FA8E1227FA7CF268" box="[1306,1413,771,795]" pageId="29" pageNumber="30">A.Newton</collectorName>
|
||||
and
|
||||
<collectorName id="268D5333FFBC2E39FCBE1206FC73F249" box="[810,906,802,826]" pageId="29" pageNumber="30">M.Thayer</collectorName>
|
||||
/carrion trap (squid)/
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FB1F1201FB5CF24A" box="[1163,1189,805,825]" count="1" pageId="29" pageNumber="30" type="male">♂</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FB391206FCA8F22A" pageId="29" pageNumber="30">FMNH-INS 0000 023 595</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FCF11265FC74F22B" box="[869,909,833,857]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FC031266FC14F22A" box="[919,1005,834,857]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="29" pageNumber="30">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FB951265FC8DF2C4" collectingDate="1976-12-19" collectingDateMax="1976-12-21" collectingDateMin="1976-12-19" collectionCode="CNCI, CNC" collectorName="H. F. Howden" country="Chile" county="MallecoProv" elevation="1100" location="MallecoProv" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="FMNH-INS 0000 023 593-594" specimenCount="3" specimenCount-male="3" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FB951265FBA8F22A" box="[1025,1105,833,857]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
,
|
||||
<collectingCounty id="62A64E69FFBC2E39FBCA1265FB18F22A" box="[1118,1249,833,857]" pageId="29" pageNumber="30">MallecoProv</collectingCounty>
|
||||
.
|
||||
<quantity id="4C809B00FFBC2E39FB7A1265FAD3F22A" box="[1262,1322,833,857]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="km" value="20.0">
|
||||
<locationDeviation id="9A752FC7FFBC2E39FB7A1265FAD3F22A" box="[1262,1322,833,857]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="km" value="20.0">20km</locationDeviation>
|
||||
</quantity>
|
||||
E Manzanar [-38.44, -71.51]
|
||||
<quantity id="4C809B00FFBC2E39FC4D1245FBD8F20A" box="[985,1057,865,889]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="m" value="1100.0">
|
||||
<elevation id="0055D1D6FFBC2E39FC4D1245FBD8F20A" box="[985,1057,865,889]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="m" value="1100.0">1100m</elevation>
|
||||
</quantity>
|
||||
<date id="FFC61025FFBC2E39FBBE1245FB46F20A" box="[1066,1215,865,889]" pageId="29" pageNumber="30" value="1976-12-19" valueMax="1976-12-21" valueMin="1976-12-19">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FBBE1245FB46F20A" box="[1066,1215,865,889]" pageId="29" pageNumber="30" value="1976-12-19" valueMax="1976-12-21" valueMin="1976-12-19">19-21.xii.1976</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FFBC2E39FB531245FAB4F20A" box="[1223,1357,865,889]" pageId="29" pageNumber="30">H.F.Howden</collectorName>
|
||||
/Beating/
|
||||
<collectionCode id="ED69AE20FFBC2E39FCBE12A4FC92F2EB" box="[810,875,896,920]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="29" pageNumber="30">CNCI</collectionCode>
|
||||
L2004-28/
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FC7812A7FBFFF2E4" box="[1004,1030,899,919]" count="1" pageId="29" pageNumber="30" type="male">♂</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FB9B12A4FA88F2EB" box="[1039,1393,896,920]" pageId="29" pageNumber="30">FMNH-INS 0000 023 593-594</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FA1D12A4FA4AF2E4" box="[1417,1459,896,920]" count="2" pageId="29" pageNumber="30" type="male">2♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FCBE12BBFC90F2C4" box="[810,873,927,951]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="29" pageNumber="30">CNC</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FCE812BBFB1AF285" collectingDate="1976-12-21" collectingDateMax="1976-12-22" collectingDateMin="1976-12-21" collectionCode="W, FMNH" collectorName="L. E. Pena" country="Chile" county="Malleco" elevation="1080" location="Las Raices" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="FMNH-INS 0000 023 596" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FCE812BBFC36F2C4" box="[892,975,927,951]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingCounty id="62A64E69FFBC2E39FC4E12BBFBC8F2C4" box="[986,1073,927,951]" pageId="29" pageNumber="30">Malleco</collectingCounty>
|
||||
:
|
||||
<location id="8EA7603EFFBC2E39FBA81284FB52F2C4" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FBA81284FB52F2C4" box="[1084,1195,927,951]" country="Chile" county="Malleco" municipality="Malalcahuello" name="Las Raices" pageId="29" pageNumber="30">Las Raices</location>
|
||||
(
|
||||
<collectionCode id="ED69AE20FFBC2E39FB2E12BBFB2CF2C4" box="[1210,1237,927,951]" pageId="29" pageNumber="30">W</collectionCode>
|
||||
of) [-38.54, -71.45],
|
||||
<date id="FFC61025FFBC2E39FCBE129BFC30F2A4" box="[810,969,959,983]" pageId="29" pageNumber="30" value="1976-12-21" valueMax="1976-12-22" valueMin="1976-12-21">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FCBE129BFC30F2A4" box="[810,969,959,983]" pageId="29" pageNumber="30" value="1976-12-21" valueMax="1976-12-22" valueMin="1976-12-21">21-22.XII.1976</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FFBC2E39FC43129BFBCBF2A5" box="[983,1074,959,982]" pageId="29" pageNumber="30">L.E.Peña</collectorName>
|
||||
leg. FIELD MUSE. NAT. HIST./
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FCBE12FAFBC5F285" box="[810,1084,990,1014]" pageId="29" pageNumber="30">FMNH-INS 0000 023 596</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FBDD12FAFB8BF286" box="[1097,1138,990,1014]" count="2" pageId="29" pageNumber="30" type="male">2♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FBEC12FAFB37F286" box="[1144,1230,990,1013]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="29" pageNumber="30">FMNH</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FB7012FAFB26F546" collectingDate="1959-01-27" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="CNC" collectorName="Km E Malalcahuello" country="Chile" county="Rio Blanco Malleco Prov" elevation="1080" location="Rio Blanco Malleco Prov" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="928331-928333" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Araucania" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FB7012FAFAC3F285" box="[1252,1338,990,1014]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBC2E39FAD612FAFA54F285" box="[1346,1453,990,1014]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
|
||||
]
|
||||
<collectingCounty id="62A64E69FFBC2E39FCBE12D9FBB2F566" box="[810,1099,1021,1045]" pageId="29" pageNumber="30">Rio Blanco Malleco Prov</collectingCounty>
|
||||
. [-39.10, -71.61]/
|
||||
<date id="FFC61025FFBC2E39FA8112D9FA9DF566" box="[1301,1380,1021,1045]" pageId="29" pageNumber="30" value="1959-01-27">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FA8112D9FA9DF566" box="[1301,1380,1021,1045]" pageId="29" pageNumber="30" value="1959-01-27">I-27-59</collectingDate>
|
||||
</date>
|
||||
Pena/
|
||||
<collectionCode id="ED69AE20FFBC2E39FCBE1539FC9AF546" box="[810,867,1053,1077]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="29" pageNumber="30">CNC</collectionCode>
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FCF81539FBE9F546" box="[876,1040,1053,1077]" pageId="29" pageNumber="30">928331-928333</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FBB01539FBB2F547" box="[1060,1099,1053,1077]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FBC11539FB85F547" box="[1109,1148,1053,1077]" count="2" pageId="29" pageNumber="30" type="female">2♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FB121539FB3CF546" box="[1158,1221,1053,1077]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="29" pageNumber="30">CNC</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FB741539FB34F5E1" collectingDate="1977-01-18" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="FMNH" collectorName="Km E Malalcahuello" country="Chile" determinerName="Newton" elevation="1080" location="Cabrerias 1100 metro Cord. Nahuelbuta" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="FMNH-INS 0000 023 577" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Araucania" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FB741539FACFF546" box="[1248,1334,1053,1077]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBC2E39FAD61539FA54F546" box="[1346,1453,1053,1077]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
|
||||
]
|
||||
<location id="8EA7603EFFBC2E39FCBE1518FB0DF527" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FCBE1518FB0DF527" box="[810,1268,1084,1108]" country="Chile" municipality="Malalcahuello" name="Cabrerias 1100 metro Cord. Nahuelbuta" pageId="29" pageNumber="30" stateProvince="Araucania">Cabrerias 1100 metro Cord. Nahuelbuta</location>
|
||||
[-37.71, -73.03]
|
||||
<date id="FFC61025FFBC2E39FCBE157FFC76F500" box="[810,911,1115,1139]" pageId="29" pageNumber="30" value="1977-01-18">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FCBE157FFC76F500" box="[810,911,1115,1139]" pageId="29" pageNumber="30" value="1977-01-18">18.I.1977</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<taxonomicName id="4C784D66FFBC2E39FC0D157FFBC0F500" authority="sp." authorityName="Sp." box="[921,1081,1115,1139]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FFBC2E39FC0D157FFBFEF500" box="[921,1031,1115,1139]" italics="true" pageId="29" pageNumber="30">Loncovilius</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
det.
|
||||
<determinerName id="7E1BDA21FFBC2E39FB181578FB19F500" box="[1164,1248,1116,1139]" pageId="29" pageNumber="30">Newton</determinerName>
|
||||
2000/
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FAA2157AFAA9F501" box="[1334,1360,1118,1138]" count="1" pageId="29" pageNumber="30" type="male">♂</specimenCount>
|
||||
/
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FACD157FFC0CF5E1" pageId="29" pageNumber="30">FMNH-INS 0000 023 577</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FB9E155FFBC8F5E1" box="[1034,1073,1147,1171]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FBA8155FFB9AF5E1" box="[1084,1123,1147,1171]" count="2" pageId="29" pageNumber="30" type="female">2♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FBFA155FFB3DF5E1" box="[1134,1220,1147,1170]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="29" pageNumber="30">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FB4C155FFA96F583" collectingDate="1976-12-14" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="FMNH" collectorName="Km E Malalcahuello" country="Chile" county="Malleco" elevation="1100" location="Cord. Nahuelbuta, Cabreria" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="FMNH-INS 0000 023 585-590, 592, 600" specimenCount="8" specimenCount-female="2" specimenCount-male="6" stateProvince="Araucania" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FB4C155FFAD2F5E0" box="[1240,1323,1147,1171]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
: [
|
||||
<collectingRegion id="49BCF807FFBC2E39FAAA155FFA54F5E0" box="[1342,1453,1147,1171]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
|
||||
]
|
||||
<collectingCounty id="62A64E69FFBC2E39FCBE15BEFC78F5C1" box="[810,897,1178,1202]" pageId="29" pageNumber="30">Malleco</collectingCounty>
|
||||
:
|
||||
<location id="8EA7603EFFBC2E39FC1815BEFB56F5C1" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FC1815BEFB56F5C1" box="[908,1199,1178,1202]" country="Chile" county="Malleco" municipality="Malalcahuello" name="Cord. Nahuelbuta, Cabreria" pageId="29" pageNumber="30" stateProvince="Araucania">Cord. Nahuelbuta, Cabreria</location>
|
||||
[-37.71, -73.03]
|
||||
<quantity id="4C809B00FFBC2E39FAF115BEFA56F5C1" box="[1381,1455,1178,1202]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="m" value="1100.0">
|
||||
<elevation id="0055D1D6FFBC2E39FAF115BEFA56F5C1" box="[1381,1455,1178,1202]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="29" pageNumber="30" unit="m" value="1100.0">1100m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFC61025FFBC2E39FCBE159DFC5EF5A2" box="[810,935,1209,1233]" pageId="29" pageNumber="30" value="1976-12-14">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FCBE159DFC5EF5A2" box="[810,935,1209,1233]" pageId="29" pageNumber="30" value="1976-12-14">14.XII.1976</collectingDate>
|
||||
</date>
|
||||
L.E.Peña leg. FIELD MUS. NAT. HIST./
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FACE159DFBD1F583" pageId="29" pageNumber="30">FMNH-INS 0000 023 585-590</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FBA315FCFB99F583" box="[1079,1120,1240,1264]" pageId="29" pageNumber="30">592</specimenCode>
|
||||
,
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FBFA15FDFB6CF582" box="[1134,1173,1241,1265]" pageId="29" pageNumber="30">600</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FB3C15FDFB2BF583" box="[1192,1234,1241,1265]" count="6" pageId="29" pageNumber="30" type="male">6♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FB4915FDFAFDF583" box="[1245,1284,1241,1265]" count="2" pageId="29" pageNumber="30" type="female">2♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FA8415FDFA9FF583" box="[1296,1382,1241,1264]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="29" pageNumber="30">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FAEF15FDFC5EF43C" collectingDate="2013-12-23" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectorName="P. N. Lonquimay" country="Chile" county="Curacautin" elevation="1080" latitude="-38.44" location="Curacautin" longLatPrecision="6" longitude="-71.51" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Araucania" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FAEF15FDFA4AF582" box="[1403,1459,1241,1265]" name="Chile" pageId="29" pageNumber="30">Chile</collectingCountry>
|
||||
[
|
||||
<collectingRegion id="49BCF807FFBC2E39FCA415DCFC64F463" box="[816,925,1272,1296]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="62A64E69FFBC2E39FC5715DCFBC5F463" box="[963,1084,1272,1296]" pageId="29" pageNumber="30">Curacautín</collectingCounty>
|
||||
]:
|
||||
<collectorName id="268D5333FFBC2E39FBF215DCFAFEF463" box="[1126,1287,1272,1296]" pageId="29" pageNumber="30">P.N.Lonquimay</collectorName>
|
||||
,
|
||||
<date id="FFC61025FFBC2E39FAB815DCFA57F463" box="[1324,1454,1272,1296]" pageId="29" pageNumber="30" value="2013-12-23">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FAB815DCFA57F463" box="[1324,1454,1272,1296]" pageId="29" pageNumber="30" value="2013-12-23">23/12/2013</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FCBE1433FC80F45C" box="[810,889,1303,1327]" degrees="38.44" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="555" value="-38.44">38.44°S</geoCoordinate>
|
||||
–
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FC131433FC18F45C" box="[903,993,1303,1327]" degrees="71.51" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="-71.51">71.51°W</geoCoordinate>
|
||||
[
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FC781433FBB7F45C" box="[1004,1102,1303,1327]" degrees="38.4400" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="5" value="-38.44">-38.4400</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FBC31433FB40F45C" box="[1111,1209,1303,1327]" degrees="71.5100" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="5" value="-71.51">-71.5100</geoCoordinate>
|
||||
], litter layer small stream [
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FCA01413FCA5F43D" box="[820,860,1335,1359]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
|
||||
TSC];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FC251413FBD6F4FE" collectingDate="2013-12-05" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectorName="Km E Malalcahuello" country="Chile" county="Pucon" elevation="1080" latitude="-39.43" location="Palguin" longLatPrecision="6" longitude="-71.79" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Araucania" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FC251413FC10F43C" box="[945,1001,1335,1359]" name="Chile" pageId="29" pageNumber="30">Chile</collectingCountry>
|
||||
[
|
||||
<collectingRegion id="49BCF807FFBC2E39FC6D1413FB9FF43C" box="[1017,1126,1335,1359]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="62A64E69FFBC2E39FBE01413FB44F43C" box="[1140,1213,1335,1359]" pageId="29" pageNumber="30">Pucón</collectingCounty>
|
||||
]:
|
||||
<location id="8EA7603EFFBC2E39FB441413FAE6F43C" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FB441413FAE6F43C" box="[1232,1311,1335,1359]" country="Chile" county="Pucon" latitude="-39.43" longLatPrecision="6" longitude="-71.79" municipality="Malalcahuello" name="Palguin" pageId="29" pageNumber="30" stateProvince="Araucania">Palguin</location>
|
||||
,
|
||||
<date id="FFC61025FFBC2E39FAB81412FA56F43D" box="[1324,1455,1334,1358]" pageId="29" pageNumber="30" value="2013-12-05">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FAB81412FA56F43D" box="[1324,1455,1334,1358]" pageId="29" pageNumber="30" value="2013-12-05">05/12/2013</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FCBE1472FC8EF41D" box="[810,887,1366,1390]" degrees="39.43" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="555" value="-39.43">39.43°S</geoCoordinate>
|
||||
–
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FC111472FC18F41D" box="[901,993,1366,1390]" degrees="71.79" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="-71.79">71.79°W</geoCoordinate>
|
||||
[
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FC7B1472FBA8F41D" box="[1007,1105,1366,1390]" degrees="39.4300" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="5" value="-39.43">-39.4300</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FBC81472FB47F41D" box="[1116,1214,1366,1390]" degrees="71.7900" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="5" value="-71.79">-71.7900</geoCoordinate>
|
||||
], litter layer/
|
||||
<taxonomicName id="4C784D66FFBC2E39FAD11472FA4AF41D" authorityName="Reyes-Hernández & Hansen & Shaw & Solodovnikov" authorityYear="2024" box="[1349,1459,1366,1390]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus" status="n. sp.">
|
||||
<emphasis id="B90CEAF7FFBC2E39FAD11472FA4AF41D" box="[1349,1459,1366,1390]" italics="true" pageId="29" pageNumber="30">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFFBC2E39FCBE1452FC9DF4FE" box="[810,868,1398,1421]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FCEC1451FC59F4FF" box="[888,928,1397,1421]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FC3F1451FC2BF4FF" box="[939,978,1397,1421]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
|
||||
TSC]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FBBB1451FB06F759" collectingDate="1999-12-09" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="NHMO" collectorName="V. I. Gusarov" country="Chile" county="Melipeuco" determinerName="A. Solodovnikov" elevation="1200" latitude="-38.6291" location="E Temuco" longLatPrecision="6" longitude="-71.6203" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCount="2" specimenCount-male="2" stateProvince="Araucania" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FBBB1451FB7CF4FE" box="[1071,1157,1397,1421]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="49BCF807FFBC2E39FB071451FB05F4FE" box="[1171,1276,1397,1421]" country="Chile" name="Araucania" pageId="29" pageNumber="30">Araucanía</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="62A64E69FFBC2E39FA9F1451FA85F4FE" box="[1291,1404,1397,1421]" pageId="29" pageNumber="30">Melipeuco</collectingCounty>
|
||||
]
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FA18145CFA5FF4FF" box="[1420,1446,1400,1420]" count="1" pageId="29" pageNumber="30" type="male">♂</specimenCount>
|
||||
/ Chile, La Araucania (IX Región),
|
||||
<quantity id="4C809B00FFBC2E39FB0A14B0FB1DF4DE" box="[1182,1252,1428,1453]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="29" pageNumber="30" unit="km" value="85.0">
|
||||
<locationDeviation id="9A752FC7FFBC2E39FB0A14B0FB1DF4DE" box="[1182,1252,1428,1453]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="29" pageNumber="30" unit="km" value="85.0">85 km</locationDeviation>
|
||||
</quantity>
|
||||
<location id="8EA7603EFFBC2E39FB7B14B1FAA7F4DF" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FB7B14B1FAA7F4DF" box="[1263,1374,1429,1452]" country="Chile" county="Melipeuco" latitude="-38.6291" longLatPrecision="6" longitude="-71.6203" municipality="Malalcahuello" name="E Temuco" pageId="29" pageNumber="30" stateProvince="Araucania">E Temuco</location>
|
||||
,
|
||||
<quantity id="4C809B00FFBC2E39FAF914B1FA4AF4DE" box="[1389,1459,1429,1453]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="km" value="26.0">
|
||||
<locationDeviation id="9A752FC7FFBC2E39FAF914B1FA4AF4DE" box="[1389,1459,1429,1453]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="km" value="26.0">26 km</locationDeviation>
|
||||
</quantity>
|
||||
<location id="8EA7603EFFBC2E39FCBE1490FC24F4BF" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FCBE1490FC24F4BF" box="[810,989,1460,1484]" country="Chile" county="Melipeuco" latitude="-38.6291" longLatPrecision="6" longitude="-71.6203" municipality="Malalcahuello" name="NNE Melipeuco" pageId="29" pageNumber="30" stateProvince="Araucania">NNE Melipeuco</location>
|
||||
,
|
||||
<location id="8EA7603EFFBC2E39FC7B1490FAE0F4BF" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FC7B1490FAE0F4BF" box="[1007,1305,1460,1484]" country="Chile" county="Melipeuco" latitude="-38.6291" longLatPrecision="6" longitude="-71.6203" municipality="Malalcahuello" name="Parque Nacional Corguillio" pageId="29" pageNumber="30" stateProvince="Araucania">Parque Nacional Corguillío</location>
|
||||
, NE coast of Laguna Conguillío,
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FB9914F7FB85F498" box="[1037,1148,1490,1515]" degrees="38" direction="south" minutes="38.24" orientation="latitude" pageId="29" pageNumber="30" precision="9" value="-38.637333">
|
||||
38°38.24
|
||||
<emphasis id="B90CEAF7FFBC2E39FBFC14F6FB97F498" box="[1128,1134,1490,1515]" italics="true" pageId="29" pageNumber="30">ʹ</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
, W
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FB2814F7FACFF498" box="[1212,1334,1490,1515]" degrees="71" direction="west" minutes="36.74" orientation="longitude" pageId="29" pageNumber="30" precision="9" value="-71.612335">
|
||||
71°36.74
|
||||
<emphasis id="B90CEAF7FFBC2E39FA8C14F6FAE7F498" box="[1304,1310,1490,1515]" italics="true" pageId="29" pageNumber="30">ʹ</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
[
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FAD914F7FA56F498" box="[1357,1455,1491,1515]" degrees="38.6291" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="5" value="-38.6291">-38.6291</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FCBE14D7FC75F778" box="[810,908,1523,1547]" degrees="71.6203" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="5" value="-71.6203">-71.6203</geoCoordinate>
|
||||
],
|
||||
<quantity id="4C809B00FFBC2E39FC3714D7FC03F779" box="[931,1018,1523,1547]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="m" value="1200.0">
|
||||
<elevation id="0055D1D6FFBC2E39FC3714D7FC03F779" box="[931,1018,1523,1547]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="m" value="1200.0">1200 m</elevation>
|
||||
</quantity>
|
||||
<collectorName id="268D5333FFBC2E39FB9314D7FB87F778" box="[1031,1150,1523,1547]" pageId="29" pageNumber="30">V.I.Gusarov</collectorName>
|
||||
<date id="FFC61025FFBC2E39FB1814D7FB17F778" box="[1164,1262,1523,1547]" pageId="29" pageNumber="30" value="1999-12-09">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FB1814D7FB17F778" box="[1164,1262,1523,1547]" pageId="29" pageNumber="30" value="1999-12-09">9.xii.1999</collectingDate>
|
||||
</date>
|
||||
/
|
||||
<taxonomicName id="4C784D66FFBC2E39FB6214D7FA5BF779" authority="sp." authorityName="Sp." box="[1270,1442,1523,1547]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||||
Loncovilius
|
||||
<taxonomicNameLabel id="A23F578CFFBC2E39FA1114D7FA5BF779" box="[1413,1442,1523,1546]" pageId="29" pageNumber="30" rank="species">sp.</taxonomicNameLabel>
|
||||
</taxonomicName>
|
||||
n.
|
||||
<determinerName id="7E1BDA21FFBC2E39FCBE1736FC36F759" box="[810,975,1554,1578]" pageId="29" pageNumber="30">A.Solodovnikov</determinerName>
|
||||
det. 2004 [
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FBF61736FB73F75A" box="[1122,1162,1554,1578]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FB0D1736FB0DF759" box="[1177,1268,1554,1578]" pageId="29" pageNumber="30">NHMO</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FA9A1736FA4AF7D4" collectingDate="1984-12-06" collectingDateMax="1985-02-19" collectingDateMin="1984-12-06" collectionCode="FMNH" collectorName="Km E Malalcahuello" country="Chile" county="Nuble" elevation="1700" location="SE Chillan, Trancas nr Termas" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCode="FMNH-INS 0000 023 578-584" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FA9A1736FA98F759" box="[1294,1377,1554,1578]" name="Chile" pageId="29" pageNumber="30">CHILE</collectingCountry>
|
||||
:
|
||||
<collectingCounty id="62A64E69FFBC2E39FAE71736FA4AF759" box="[1395,1459,1554,1578]" pageId="29" pageNumber="30">Nuble</collectingCounty>
|
||||
[Ñuble] Prov.,
|
||||
<quantity id="4C809B00FFBC2E39FC5D1715FBF5F73A" box="[969,1036,1585,1609]" metricMagnitude="4" metricUnit="m" metricValue="7.2" pageId="29" pageNumber="30" unit="km" value="72.0">
|
||||
<locationDeviation id="9A752FC7FFBC2E39FC5D1715FBF5F73A" box="[969,1036,1585,1609]" metricMagnitude="4" metricUnit="m" metricValue="7.2" pageId="29" pageNumber="30" unit="km" value="72.0">72 km</locationDeviation>
|
||||
</quantity>
|
||||
<location id="8EA7603EFFBC2E39FB811715FAA0F73A" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FB811715FAA0F73A" box="[1045,1369,1585,1609]" country="Chile" county="Nuble" municipality="Malalcahuello" name="SE Chillan, Trancas nr Termas" pageId="29" pageNumber="30">SE Chillan, Trancas nr Termas</location>
|
||||
[-36.90, -71.44],
|
||||
<quantity id="4C809B00FFBC2E39FC0E1775FC1DF71A" box="[922,996,1617,1641]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="29" pageNumber="30" unit="m" value="1700.0">
|
||||
<elevation id="0055D1D6FFBC2E39FC0E1775FC1DF71A" box="[922,996,1617,1641]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="29" pageNumber="30" unit="m" value="1700.0">1700m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFC61025FFBC2E39FB931774FB81F71B" box="[1031,1144,1616,1640]" pageId="29" pageNumber="30" value="1984-12-06">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FB931774FB81F71B" box="[1031,1144,1616,1640]" pageId="29" pageNumber="30" value="1984-12-06">6.XII.1984</collectingDate>
|
||||
</date>
|
||||
-
|
||||
<date id="FFC61025FFBC2E39FB141774FB1FF71B" box="[1152,1254,1616,1640]" pageId="29" pageNumber="30" value="1985-02-19">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FB141774FB1FF71B" box="[1152,1254,1616,1640]" pageId="29" pageNumber="30" value="1985-02-19">19.II.1985</collectingDate>
|
||||
</date>
|
||||
/FMNHD#85-893,
|
||||
<taxonomicName id="4C784D66FFBC2E39FCBE1754FC5AF7FB" box="[810,931,1648,1672]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
|
||||
forest, S.Peck, P#85-8 FIT FIELD MUSEUM NAT HIST/
|
||||
<specimenCode id="DBDE9E9EFFBC2E39FCFA17ABFB28F7D4" box="[878,1233,1679,1703]" pageId="29" pageNumber="30">FMNH-INS 0000 023 578–584</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FB7317ABFAE9F7D5" box="[1255,1296,1679,1703]" count="3" pageId="29" pageNumber="30" type="male">3♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FA8A17ABFABCF7D5" box="[1310,1349,1679,1703]" count="4" pageId="29" pageNumber="30" type="female">4♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FAC717B4FA50F7D4" box="[1363,1449,1680,1703]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="29" pageNumber="30">FMNH</collectionCode>
|
||||
];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FCBE178BFB38F676" collectingDate="1978-12-04" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="NHMD" collectorName="Km E Malalcahuello" country="Argentina" elevation="770" location="Lago Nahuel Huapi Puerto Blest" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Rio Negro" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FCBE178BFC3DF7B4" box="[810,964,1711,1735]" name="Argentina" pageId="29" pageNumber="30">ARGENTINA</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBC2E39FC5B178BFBC4F7B4" box="[975,1085,1711,1735]" country="Argentina" name="Rio Negro" pageId="29" pageNumber="30">Rio Negro</collectingRegion>
|
||||
8:
|
||||
<location id="8EA7603EFFBC2E39FBCB178BFA4AF7B4" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FBCB178BFA4AF7B4" box="[1119,1459,1711,1735]" country="Argentina" municipality="Malalcahuello" name="Lago Nahuel Huapi Puerto Blest" pageId="29" pageNumber="30" stateProvince="Rio Negro">Lago Nahuel Huapi Puerto Blest</location>
|
||||
[-41.02, -71.81],
|
||||
<quantity id="4C809B00FFBC2E39FC7117EAFBD5F796" box="[997,1068,1742,1766]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="29" pageNumber="30" unit="m" value="770.0">
|
||||
<elevation id="0055D1D6FFBC2E39FC7117EAFBD5F796" box="[997,1068,1742,1766]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="29" pageNumber="30" unit="m" value="770.0">770 m</elevation>
|
||||
</quantity>
|
||||
<date id="FFC61025FFBC2E39FBA317EAFB65F795" box="[1079,1180,1742,1766]" pageId="29" pageNumber="30" value="1978-12-04">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FBA317EAFB65F795" box="[1079,1180,1742,1766]" pageId="29" pageNumber="30" value="1978-12-04">4.xii.1978</collectingDate>
|
||||
</date>
|
||||
<location id="8EA7603EFFBC2E39FB3317EAFA4CF796" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FB3317EAFA4CF796" box="[1191,1461,1742,1766]" country="Argentina" municipality="Malalcahuello" name="Mision Cientifica Danesa" pageId="29" pageNumber="30" stateProvince="Rio Negro">Mision Cientifica Danesa</location>
|
||||
[pale orange label
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FC6917C9FBDDF677" box="[1021,1060,1773,1797]" count="2" pageId="29" pageNumber="30" type="male">2♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FBA517C9FBA1F677" box="[1073,1112,1773,1797]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FBF217CAFB44F676" box="[1126,1213,1774,1797]" collectionName="NHMD" pageId="29" pageNumber="30">NHMD</collectionCode>
|
||||
;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FB5A17C9FC7CF657" collectingDate="1982-12-13" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="MNNC, NHMD" collectorName="Km E Malalcahuello" country="Chile" elevation="1080" location="Malleco Prov." municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FB5A17C9FB0CF677" box="[1230,1269,1773,1797]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FA9717C9FAD3F677" box="[1283,1322,1773,1797]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FAA317C9FA70F676" box="[1335,1417,1773,1797]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="29" pageNumber="30">MNNC</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FFBC2E39FCBE1629FC78F657" box="[810,897,1805,1828]" collectionName="NHMD" pageId="29" pageNumber="30">NHMD</collectionCode>
|
||||
;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FC191629FB2AF656" box="[909,1235,1805,1829]" collectingDate="1982-12-13" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="CZUG, NHMD" collectorName="Km E Malalcahuello" country="Chile" elevation="1080" location="Malleco Prov." municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FC191629FC4DF657" box="[909,948,1805,1829]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FC281629FC1AF657" box="[956,995,1805,1829]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FC7F1629FBCDF656" box="[1003,1076,1805,1829]" collectionName="Mexico, Jalisco, Guadalajara, Universidad de Guadalajara, Centro de Estudios en Zoologia, Entomologia" pageId="29" pageNumber="30">CZUG</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FFBC2E39FBF41629FB43F657" box="[1120,1210,1805,1828]" collectionName="NHMD" pageId="29" pageNumber="30">NHMD</collectionCode>
|
||||
]; [
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E39FB471629FA4AF630" collectingDate="1978-12-16" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="NHMD" collectorName="Km E Malalcahuello" country="Argentina" elevation="1080" location="ARGENTINA" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBC2E39FB471629FA8DF656" box="[1235,1396,1805,1829]" name="Argentina" pageId="29" pageNumber="30">ARGENTINA</collectingCountry>
|
||||
: same locality, only the date different (
|
||||
<date id="FFC61025FFBC2E39FBE01608FB14F637" box="[1140,1261,1836,1860]" pageId="29" pageNumber="30" value="1978-12-16">
|
||||
<collectingDate id="EF82E9CDFFBC2E39FBE01608FB14F637" box="[1140,1261,1836,1860]" pageId="29" pageNumber="30" value="1978-12-16">16.xii.1978</collectingDate>
|
||||
</date>
|
||||
)
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FB681608FADAF630" box="[1276,1315,1836,1860]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FABE1608FAA8F630" box="[1322,1361,1836,1860]" count="3" pageId="29" pageNumber="30" type="female">3♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FACC1608FA56F630" box="[1368,1455,1836,1859]" collectionName="NHMD" pageId="29" pageNumber="30">NHMD</collectionCode>
|
||||
;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBC2E3AFCB8166FFEE6F1A0" collectingDate="1982-12-13" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="NHM, NHMD" collectorName="Paula Sackmann" country="Argentina" determinerName="V. I. Gusarov" elevation="150" lastPageId="30" lastPageNumber="31" latitude="-41.3525" location="Bariloche forest, Nothofagus antartica, unburnt" longLatPrecision="1" longitude="-71.62625" municipality="Malalcahuello" pageId="29" pageNumber="30" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Rio Negro Prov." typeStatus="holotype">
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FCB8166FFCA8F611" box="[812,849,1867,1891]" count="1" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBC2E39FCCB166FFC7FF611" box="[863,902,1867,1891]" count="1" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBC2E39FC001668FC2DF610" box="[916,980,1868,1891]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="29" pageNumber="30">NHM</collectionCode>
|
||||
ex.
|
||||
<collectionCode id="ED69AE20FFBC2E39FB991668FB9EF610" box="[1037,1127,1868,1891]" collectionName="NHMD" pageId="29" pageNumber="30">NHMD</collectionCode>
|
||||
];
|
||||
<collectingCountry id="F36F7675FFBC2E39FBEB166FFB1EF610" box="[1151,1255,1867,1891]" name="Argentina" pageId="29" pageNumber="30">Argentina</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBC2E39FB6C166FFA49F610" box="[1272,1456,1867,1892]" country="Argentina" name="Rio Negro" pageId="29" pageNumber="30">Rio Negro Prov.</collectingRegion>
|
||||
,
|
||||
<location id="8EA7603EFFBC2E39FCBE164FFAC1F6F0" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBC2E39FCBE164FFAC1F6F0" box="[810,1336,1899,1923]" country="Argentina" latitude="-41.3525" longLatPrecision="1" longitude="-71.62625" municipality="Malalcahuello" name="Bariloche forest, Nothofagus antartica, unburnt" pageId="29" pageNumber="30" stateProvince="Rio Negro Prov.">
|
||||
Bariloche forest,
|
||||
<taxonomicName id="4C784D66FFBC2E39FC7F164FFB9DF6F0" box="[1003,1124,1899,1923]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
|
||||
antartica, unburnt
|
||||
</location>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FAD8164FFA4AF6F0" box="[1356,1459,1897,1923]" degrees="41" direction="south" minutes="21" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="9" value="-41.352497">
|
||||
41°21
|
||||
<emphasis id="B90CEAF7FFBC2E39FA1C164DFA77F6F1" box="[1416,1422,1897,1922]" italics="true" pageId="29" pageNumber="30">ʹ</emphasis>
|
||||
9
|
||||
<emphasis id="B90CEAF7FFBC2E39FA0E164DFA5CF6F1" box="[1434,1445,1897,1922]" italics="true" pageId="29" pageNumber="30">ʹʹ</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FCBE16AEFC28F6D2" box="[810,977,1928,1954]" degrees="71" direction="west" minutes="37" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="34.5" value="-71.62625">
|
||||
71°37
|
||||
<emphasis id="B90CEAF7FFBC2E39FCF216ACFC95F6D2" box="[870,876,1928,1953]" italics="true" pageId="29" pageNumber="30">ʹ</emphasis>
|
||||
34.5
|
||||
<emphasis id="B90CEAF7FFBC2E39FC0D16ACFC5DF6D2" box="[921,932,1928,1953]" italics="true" pageId="29" pageNumber="30">ʹʹ</emphasis>
|
||||
W
|
||||
</geoCoordinate>
|
||||
, [
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FC7B16AEFB92F6D1" box="[1007,1131,1930,1954]" degrees="41.352500" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="1" value="-41.3525">-41.352500</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE4C5022FFBC2E39FB1016AEFB06F6D1" box="[1156,1279,1930,1954]" degrees="71.626250" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="1" value="-71.62625">-71.626250</geoCoordinate>
|
||||
] 823 ±
|
||||
<quantity id="4C809B00FFBC2E39FAC916AEFA56F6D2" box="[1373,1455,1930,1954]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="150.0">
|
||||
<elevation id="0055D1D6FFBC2E39FAC916AEFA56F6D2" box="[1373,1455,1930,1954]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="150.0">150 m</elevation>
|
||||
</quantity>
|
||||
, ABCONF17 P.Sackmann/Argentina, Patagonia ABCONF17
|
||||
<collectorName id="268D5333FFBF2E3AFF1511BFFED0F1C0" box="[129,297,155,179]" pageId="30" pageNumber="31">Paula Sackmann</collectorName>
|
||||
leg./
|
||||
<taxonomicName id="4C784D66FFBF2E3AFECE11BFFE02F1C0" authority="sp." authorityName="Sp." box="[346,507,155,179]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="30" phylum="Arthropoda" rank="genus">Loncovilius sp.</taxonomicName>
|
||||
<specimenCount id="9D7EFD6CFFBF2E3AFD9711BBFDE4F1C0" box="[515,541,159,179]" count="1" pageId="30" pageNumber="31" type="female">♀</specimenCount>
|
||||
<determinerName id="7E1BDA21FFBF2E3AFDB611B8FD58F1C7" box="[546,673,156,180]" pageId="30" pageNumber="31">V.I. Gusarov</determinerName>
|
||||
det. 2002 [
|
||||
<specimenCount id="9D7EFD6CFFBF2E3AFF1F119FFF4BF1A1" box="[139,178,187,211]" count="1" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
|
||||
NHMO];
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBF2E3AFEB3119FFD2CF062" collectingDate="1972-01-22" collectingDateMax="1982-12-31" collectingDateMin="1982-12-13" collectionCode="AMNH, FMNH" collectorName="L. Herman" country="Argentina" elevation="1080" location="Cerro Malo" municipality="Malalcahuello" pageId="30" pageNumber="31" specimenCode="FMNH-INS 0000 023 597-599" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Neuquen" typeStatus="holotype">
|
||||
<collectingCountry id="F36F7675FFBF2E3AFEB3119FFE38F1A0" box="[295,449,187,211]" name="Argentina" pageId="30" pageNumber="31">ARGENTINA</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBF2E3AFE58119FFDC9F1A0" box="[460,560,187,211]" country="Argentina" name="Neuquen" pageId="30" pageNumber="31">Neuquén</collectingRegion>
|
||||
:
|
||||
<location id="8EA7603EFFBF2E3AFDAE119FFD4CF1A0" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBF2E3AFDAE119FFD4CF1A0" box="[570,693,187,211]" country="Argentina" municipality="Malalcahuello" name="Cerro Malo" pageId="30" pageNumber="31" stateProvince="Neuquen">Cerro Malo</location>
|
||||
,
|
||||
<quantity id="4C809B00FFBF2E3AFD54119FFD17F1A0" box="[704,750,187,211]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="31" unit="km" value="6.0">
|
||||
<locationDeviation id="9A752FC7FFBF2E3AFD54119FFD17F1A0" box="[704,750,187,211]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="31" unit="km" value="6.0">6km</locationDeviation>
|
||||
</quantity>
|
||||
<location id="8EA7603EFFBF2E3AFD61119FFF32F181" LSID="urn:lsid:plazi:treatment:03D187F3FFBC2E3AFE8617A3FAFCF4B2:8EA7603EFFBF2E3AFD61119FFF32F181" country="Argentina" municipality="Malalcahuello" name="N Pucara" pageId="30" pageNumber="31" stateProvince="Neuquen">N Pucara</location>
|
||||
,
|
||||
<date id="FFC61025FFBF2E3AFF4711FFFEAFF181" box="[211,342,218,242]" pageId="30" pageNumber="31" value="1972-01-22">
|
||||
<collectingDate id="EF82E9CDFFBF2E3AFF4711FFFEAFF181" box="[211,342,218,242]" pageId="30" pageNumber="31" value="1972-01-22">Jan. 22, 1972</collectingDate>
|
||||
</date>
|
||||
<collectorName id="268D5333FFBF2E3AFECF11FFFE3CF181" box="[347,453,219,242]" pageId="30" pageNumber="31">L.Herman</collectorName>
|
||||
874/
|
||||
<collectionCode id="ED69AE20FFBF2E3AFE6E11FEFDB7F181" box="[506,590,218,242]" collectionName="USA, New York, New York, American Museum of Natural History" pageId="30" pageNumber="31">AMNH</collectionCode>
|
||||
1489 [pale orange label]/
|
||||
<specimenCode id="DBDE9E9EFFBF2E3AFF5611DDFDF5F062" box="[194,524,249,274]" pageId="30" pageNumber="31">FMNH-INS 0000 023 597–599</specimenCode>
|
||||
[
|
||||
<specimenCount id="9D7EFD6CFFBF2E3AFD8811DEFDBDF062" box="[540,580,250,274]" count="1" pageId="30" pageNumber="31" type="male">1♂</specimenCount>
|
||||
<specimenCount id="9D7EFD6CFFBF2E3AFDDD11DEFD89F062" box="[585,624,250,274]" count="2" pageId="30" pageNumber="31" type="female">2♀</specimenCount>
|
||||
<collectionCode id="ED69AE20FFBF2E3AFDE211DEFD32F062" box="[630,715,250,273]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="30" pageNumber="31">FMNH</collectionCode>
|
||||
].
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBF2E3AFF08103DFF42F003" blockId="30.[128,780,155,368]" pageId="30" pageNumber="31">
|
||||
All
|
||||
<typeStatus id="54C38847FFBF2E3AFF2B103DFEDBF042" box="[191,290,281,305]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
|
||||
with label: PARATYPE
|
||||
<taxonomicName id="4C784D66FFBF2E3AFD88103DFCF3F042" box="[540,778,281,305]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="cantharoides">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFD88103DFCF3F042" box="[540,778,281,305]" italics="true" pageId="30" pageNumber="31">Loncovilius cantharoides</emphasis>
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBF2E3AFF1510B6FD71F355" pageId="30" pageNumber="31" type="diagnosis">
|
||||
<paragraph id="8BC736E5FFBF2E3AFF1510B6FD71F355" blockId="30.[128,778,401,550]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFF1510B6FF1EF0DA" box="[129,231,402,425]" italics="true" pageId="30" pageNumber="31">Diagnosis:</emphasis>
|
||||
Nuchal ridge with small portion absent dorsally. Pronotum outline evenly curved. Elytra yellow or yellowish-brown with brown-black mark around mesoscutellum and suture. Protibiae sexually dimorphic, with laterodorsal row of thick spines only females. Aedeagus as in
|
||||
<figureCitation id="13432A60FFBF2E3AFE7F132AFD7BF355" box="[491,642,526,550]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="30" pageNumber="31">Figure 10O–Q</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBF2E3AFF15136CFA3AF566" pageId="30" pageNumber="31" type="description">
|
||||
<paragraph id="8BC736E5FFBF2E3AFF15136CFE0CF38F" blockId="30.[128,780,583,1986]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFF15136CFF0EF32C" box="[129,247,584,607]" italics="true" pageId="30" pageNumber="31">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFBF2E3AFE3C136FFE3BF32C" box="[424,450,587,607]" count="1" pageId="30" pageNumber="31" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFBF2E3AFD2E136CFD34F32C" box="[698,717,584,607]" italics="true" pageId="30" pageNumber="31">N</emphasis>
|
||||
= 5]: FBL = 3.98–4.34 (4.22); TL = 7.18–8.4 (7.59); HW = 0.93– 1.08 (1); HL = 1.03–1.23 (1.16); HW/HL = 0.88–0.91 (0.89); PW = 1.28–1.36 (1.32); PL = 1.25–1.36 (1.31); PW/ PL = 1–1.04 (1.02); EW = 1.74–1.92 (1.84); EL = 1.59–1.81 (1.7); PW/HW = 1.26–1.28 (1.27).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBF2E3AFF081220FE86F2CB" blockId="30.[128,780,583,1986]" pageId="30" pageNumber="31">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFBF2E3AFEA31223FEA8F268" box="[311,337,775,795]" count="1" pageId="30" pageNumber="31" type="female">♀</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFBF2E3AFDA51220FDBDF268" box="[561,580,772,795]" italics="true" pageId="30" pageNumber="31">N</emphasis>
|
||||
= 5]: FBL = 4.43– 4.67 (4.55); TL = 8.13–8.47 (8.3); HW = 1.18–1.22 (1.20); HL = 1.25–1.33 (1.29); HW/HL = 0.92–0.94 (0.93); PW = 1.44–1.49 (1.47);PL = 1.4–1.44 (1.42);PW/PL = 1.03– 1.03 (1.03); EW = 2–2.1 (2.05); EL = 1.78–1.9 (1.84); PW/ HW = 4.43–4.67 (4.55).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBF2E3AFF08129BFD13F526" blockId="30.[128,780,583,1986]" pageId="30" pageNumber="31">Head and pronotum brown-black with slight greenish, bluish, or golden overtones, with yellow margins; antennae, mouthparts, and legs yellow or yellowish-brown; elytra yellow with brown-black mark around mesoscutellum and suture; tergites and sternites dark reddish-brown with pale posterior margin.</paragraph>
|
||||
<paragraph id="8BC736E5FFBF2E3AFF081578FD9EF617" blockId="30.[128,780,583,1986]" pageId="30" pageNumber="31">
|
||||
Head elongate; dorsally and ventrally dull with metallic iridescence; with micropunctures and microsculpture of transverse isodiametric meshes, without coarse non-setiferous punctures; posterior angles indistinct with several setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.44), from 1.05 to 1.11 times as long as temples (in lateral view); distance between eyes about 1.24 times as long as length of eye in both sexes. Antennomere 3 moderately longer than antennomere 2; antennomeres 4 and 5 subequal in length; antennomeres 8 to 10 subequal in length; a11 from 1.52 to 1.7 times long as a
|
||||
<quantity id="4C809B00FFBF2E3AFE1E14B2FE45F4DE" box="[394,444,1429,1454]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="30" pageNumber="31" unit="in" value="10.0">10 in</quantity>
|
||||
males and from
|
||||
<quantity id="4C809B00FFBF2E3AFDF114B2FD13F4DE" box="[613,746,1429,1454]" metricMagnitude="-2" metricUnit="m" metricValue="3.6576" metricValueMax="3.7592" metricValueMin="3.5559999999999996" pageId="30" pageNumber="31" unit="in" value="1.44" valueMax="1.48" valueMin="1.4">1.4 to 1.48 in</quantity>
|
||||
females. Basal and parocular punctures usually double; posterior frontal puncture located anterior to temporal puncture; with small setiferous punctures between frontoclypeal and anterior frontal punctures: ventral basal ridge in checkmark shape, almost united with gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and a little dorsally, not fused with infraorbital ridge; gula with conspicuous isodiametric microsculpture medially, gular sutures widely and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBF2E3AFF081648FADFF194" blockId="30.[128,780,583,1986]" lastBlockId="30.[824,1475,144,1046]" pageId="30" pageNumber="31">Pronotum about as wide as long, convex, evenly curved, with two PPDS, without APP, without SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to four or five times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially.</paragraph>
|
||||
<paragraph id="8BC736E5FFBF2E3AFCC111CAFC6AF092" blockId="30.[824,1475,144,1046]" pageId="30" pageNumber="31">Protibiae sexually dimorphic, with laterodorsal row of thick spines only females. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 1–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed.</paragraph>
|
||||
<paragraph id="8BC736E5FFBF2E3AFCC110CDFBF7F22A" blockId="30.[824,1475,144,1046]" pageId="30" pageNumber="31">
|
||||
Abdominal tergites III and IV with only a few rows of coarse, sparse punctures, with large, impunctate area; tergite VIII not emarginate medio-apically. Sternite III with basal transverse carina evenly converging at obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with U-shaped emargination medially (
|
||||
<figureCitation id="13432A60FFBF2E3AFB0013A1FB1FF3EE" box="[1172,1254,645,669]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="30" pageNumber="31">Fig. 14F</figureCitation>
|
||||
); female sternite VIII without emargination; male sternite IX without emargination medially, with rounded apex, its basal portion shorter than distal portion; male tergite X with anterior margin subtruncate (
|
||||
<figureCitation id="13432A60FFBF2E3AFA0813C7FC98F268" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="30" pageNumber="31">Fig. 12F</figureCitation>
|
||||
); female tergite X medio-apically distinctly subacute to acute (
|
||||
<figureCitation id="13432A60FFBF2E3AFC171206FC23F249" box="[899,986,802,826]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="30" pageNumber="31">Fig. 13F</figureCitation>
|
||||
); ovipositor, its second gonocoxite with three macrosetae medially.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBF2E3AFCC11245FA3AF566" blockId="30.[824,1475,144,1046]" pageId="30" pageNumber="31">
|
||||
Aedeagus as in
|
||||
<figureCitation id="13432A60FFBF2E3AFC6B1245FB64F20A" box="[1023,1181,865,889]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="30" pageNumber="31">Figure 10O–Q</figureCitation>
|
||||
; its total length 1.25–1.31. Median lobe rod-like; internal sac with pair of long inverted L-shaped copulatory sclerites. Paramere distinctly produced over apex of median lobe; wider than median lobe; converging to a subrectangular apex; with small translucent peg setae; in lateral view almost straight, curved away from median lobe apically.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBF2E3AFCAD1513FC47F5DE" pageId="30" pageNumber="31" type="etymology">
|
||||
<paragraph id="8BC736E5FFBF2E3AFCAD1513FC47F5DE" blockId="30.[825,1474,1079,1197]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFCAD1513FC51F53C" box="[825,936,1079,1103]" italics="true" pageId="30" pageNumber="31">Etymology:</emphasis>
|
||||
The species epithet is derived from the Greek κανθαΡίς (
|
||||
<emphasis id="B90CEAF7FFBF2E3AFC3C1572FBFDF51D" box="[936,1028,1110,1134]" italics="true" pageId="30" pageNumber="31">kantharís</emphasis>
|
||||
; soldier beetles) and suffix -ειδής (
|
||||
<emphasis id="B90CEAF7FFBF2E3AFA101572FA43F51D" box="[1412,1466,1106,1134]" italics="true" pageId="30" pageNumber="31">
|
||||
-eid
|
||||
<emphasis id="B90CEAF7FFBF2E3AFA3C1576FA4AF51D" bold="true" box="[1448,1459,1106,1134]" italics="true" pageId="30" pageNumber="31">ḗ</emphasis>
|
||||
s
|
||||
</emphasis>
|
||||
; like), as the habitus resembles some species of the beetle family
|
||||
<taxonomicName id="4C784D66FFBF2E3AFCAE15B1FC43F5DE" authorityName="Imhoff" authorityYear="1856" box="[826,954,1173,1197]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBF2E3AFCAD15EAFAFCF4B2" pageId="30" pageNumber="31" type="distribution">
|
||||
<paragraph id="8BC736E5FFBF2E3AFCAD15EAFAFCF4B2" blockId="30.[825,1477,1230,1473]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFCAD15EAFAD1F595" box="[825,1320,1230,1254]" italics="true" pageId="30" pageNumber="31">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FFBF2E3AFBAC15EAFAD1F595" box="[1080,1320,1230,1254]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="cantharoides">Loncovilius cantharoides</taxonomicName>
|
||||
</emphasis>
|
||||
is known from
|
||||
<collectingCountry id="F36F7675FFBF2E3AFCAF15C9FC8CF476" box="[827,885,1261,1285]" name="Chile" pageId="30" pageNumber="31">Chile</collectingCountry>
|
||||
, from the
|
||||
<collectingRegion id="49BCF807FFBF2E3AFC4C15C9FBC6F476" box="[984,1087,1261,1285]" country="Chile" name="Araucania" pageId="30" pageNumber="31">Araucanía</collectingRegion>
|
||||
and Ñuble regions, and from the westernmost portions of the
|
||||
<collectingRegion id="49BCF807FFBF2E3AFBAE1429FB63F456" box="[1082,1178,1293,1317]" country="Argentina" name="Neuquen" pageId="30" pageNumber="31">Neuquén</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="49BCF807FFBF2E3AFB5A1429FAC5F456" box="[1230,1340,1293,1317]" country="Argentina" name="Rio Negro" pageId="30" pageNumber="31">Río Negro</collectingRegion>
|
||||
provinces of
|
||||
<collectingCountry id="F36F7675FFBF2E3AFCA81408FC5DF437" box="[828,932,1324,1348]" name="Argentina" pageId="30" pageNumber="31">Argentina</collectingCountry>
|
||||
. The species occurs from
|
||||
<quantity id="4C809B00FFBF2E3AFB3D1408FAB9F430" box="[1193,1344,1324,1348]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="30" pageNumber="31" unit="m" value="1200.0" valueMax="1600.0" valueMin="800.0">800 to 1600 m</quantity>
|
||||
of elevation; reported in the
|
||||
<emphasis id="B90CEAF7FFBF2E3AFC711468FB46F410" box="[997,1215,1355,1379]" italics="true" pageId="30" pageNumber="31">
|
||||
<taxonomicName id="4C784D66FFBF2E3AFC711468FBBEF410" box="[997,1095,1356,1379]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">Araucaria</taxonomicName>
|
||||
-
|
||||
<taxonomicName id="4C784D66FFBF2E3AFBDB146FFB46F410" box="[1103,1215,1355,1379]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">Nothofagus</taxonomicName>
|
||||
</emphasis>
|
||||
forests from the central and north-central portions of the Valdivian temperate forest ecoregion. It has been collected using Malaise, window, carrion and dung-baited pitfall traps, and sifting leaf litter.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,373 +0,0 @@
|
|||
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<mods:title id="716890BD3ED2EF073BC8D7F86157DF06">Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="9F1AB15876C8A1C666DEAB256C7182A5">Reyes-Hernández, José L.</mods:namePart>
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<mods:affiliation id="224E1EF61C7B345D31EBF46E839CF13A">Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany</mods:affiliation>
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<mods:url id="48BF6E922371E7BC1241F3FD5432EAA2">https://doi.org/10.1093/zoolinnean/zlad143</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="EECD9D77B9376F4392D90D31D50E7C7F">journal article</mods:classification>
|
||||
<mods:identifier id="B990CEB9A95788813766534628D0957F" type="DOI">10.1093/zoolinnean/zlad143</mods:identifier>
|
||||
<mods:identifier id="0E6AD2BA660D1E4D8DAD4263E15BAC55" type="ISSN">0024-4082</mods:identifier>
|
||||
<mods:identifier id="ECDCE035DF20A5F56F430290E317065E" type="Zenodo-Dep">14541979</mods:identifier>
|
||||
<mods:identifier id="8163F90BE136B8749B2B0D2E25F609B5" type="ZooBank">CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="03D187F3FFBF2E04FC2414C0FE3AF0F0" LSID="urn:lsid:plazi:treatment:03D187F3FFBF2E04FC2414C0FE3AF0F0" httpUri="http://treatment.plazi.org/id/03D187F3FFBF2E04FC2414C0FE3AF0F0" lastPageId="32" lastPageNumber="32" pageId="30" pageNumber="31">
|
||||
<subSubSection id="C362656EFFBF2E3AFC2414C0FAB5F48D" box="[944,1356,1508,1535]" pageId="30" pageNumber="31" type="nomenclature">
|
||||
<paragraph id="8BC736E5FFBF2E3AFC2414C0FAB5F48D" blockId="30.[944,1356,1508,1535]" box="[944,1356,1508,1535]" pageId="30" pageNumber="31">
|
||||
<heading id="D08F8189FFBF2E3AFC2414C0FAB5F48D" bold="true" box="[944,1356,1508,1535]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
|
||||
<taxonomicName id="4C784D66FFBF2E3AFC2414C0FAB5F48D" ID-CoL="3VYHP" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[944,1356,1508,1535]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFC2414C0FAB5F48D" bold="true" box="[944,1356,1508,1535]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFC2414C0FB56F48C" bold="true" box="[944,1199,1508,1535]" italics="true" pageId="30" pageNumber="31">Loncovilius edwardsianus</emphasis>
|
||||
(Korge, 1963)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBF2E3AFC29173AFAC6F74B" box="[957,1343,1566,1592]" pageId="30" pageNumber="31" type="description">
|
||||
<paragraph id="8BC736E5FFBF2E3AFC29173AFAC6F74B" blockId="30.[957,1343,1566,1592]" box="[957,1343,1566,1592]" pageId="30" pageNumber="31">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFC29173AFAC6F74B" bold="true" box="[957,1343,1566,1592]" pageId="30" pageNumber="31">
|
||||
(
|
||||
<figureCitation id="13432A60FFBF2E3AFC53173AFBEFF74B" box="[967,1046,1566,1592]" captionStart="Figure 1" captionStartId="1.[113,178,1870,1894]" captionTargetBox="[152,1420,144,1842]" captionTargetId="figure-6@1.[152,1420,144,1842]" captionTargetPageId="1" captionText="Figure 1. Habitats of Loncovilius. A, L. edwardsianus (Korge, 1963) live specimen in the Valdivian Temperate Rainforest in Chiloé, Los Lagos, Chile (Photograph: Matías Gargiulo, through iNaturalist). B, PN Puyehue, Chile, L. lividipennis was collected in mosses on the stones at river banks and under stones in accumulations along banks (photograph: Martin Fikáček). C, L. variabilis sp. nov. found in a flower of the Chiloé archipelago (photograph:Matías Gargiulo, through iNaturalist)." pageId="30" pageNumber="31">Figs 1A</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBF2E3AFBB4173AFBB9F74B" box="[1056,1088,1566,1592]" captionStart="Figure 6" captionStartId="14.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@14.[168,1434,144,1840]" captionTargetPageId="14" captionText="Figure 6. Habitus of Loncovilius species. A, male of L. edwardsianus. B, male of L. cantharoides. C, female of L. semiflavus. D, male of L. barclayi." pageId="30" pageNumber="31">6A</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBF2E3AFBDE173AFB6EF74B" box="[1098,1175,1566,1592]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="30" pageNumber="31">10L–N</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBF2E3AFB35173AFB35F74B" box="[1185,1228,1566,1592]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="30" pageNumber="31">12E</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBF2E3AFB42173AFAF8F74B" box="[1238,1281,1566,1592]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="30" pageNumber="31">13E</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A60FFBF2E3AFA9E173AFACDF74B" box="[1290,1332,1566,1592]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="30" pageNumber="31">14E</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBF2E3AFCAD177CFBD1F619" pageId="30" pageNumber="31" type="reference_group">
|
||||
<paragraph id="8BC736E5FFBF2E3AFCAD177CFC40F7FC" blockId="30.[825,1475,1623,1898]" pageId="30" pageNumber="31">
|
||||
=
|
||||
<taxonomicName id="4C784D66FFBF2E3AFCDC1773FB26F71C" authority="(Bernhauer, 1935)" baseAuthorityName="Bernhauer" baseAuthorityYear="1935" box="[840,1247,1623,1647]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsi">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFCDC1773FBEDF71C" box="[840,1044,1623,1647]" italics="true" pageId="30" pageNumber="31">Loncovilius edwardsi</emphasis>
|
||||
(Bernhauer, 1935)
|
||||
</taxonomicName>
|
||||
nec
|
||||
<taxonomicName id="4C784D66FFBF2E3AFA871773FC4CF7FC" authority="Sharp, 1886" authorityName="Bernh. Nec Sharp" authorityYear="1886" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsi">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFA871773FA3BF71C" box="[1299,1474,1623,1647]" italics="true" pageId="30" pageNumber="31">Quedius edwardsi</emphasis>
|
||||
Sharp, 1886
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBF2E3AFCC117B2FBD1F619" blockId="30.[825,1475,1623,1898]" pageId="30" pageNumber="31">
|
||||
Bernhauer, 1935: 92 (as
|
||||
<taxonomicName id="4C784D66FFBF2E3AFBCF17B2FAF3F7DD" authorityName="Bernh. Nec Sharp" authorityYear="1886" box="[1115,1290,1686,1710]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsi">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFBCF17B2FAF3F7DD" box="[1115,1290,1686,1710]" italics="true" pageId="30" pageNumber="31">Quedius edwardsi</emphasis>
|
||||
</taxonomicName>
|
||||
, original description); Korge, 1963 (as
|
||||
<taxonomicName id="4C784D66FFBF2E3AFBDF1791FAD7F7BE" authorityName="Korge" authorityYear="1963" box="[1099,1326,1717,1741]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFBDF1791FAD7F7BE" box="[1099,1326,1717,1741]" italics="true" pageId="30" pageNumber="31">Quedius edwardsianus</emphasis>
|
||||
</taxonomicName>
|
||||
, replacement name);
|
||||
<bibRefCitation id="EFE94B14FFBF2E3AFC0417F1FB3DF79E" author="Coiffait H & Saiz F" box="[912,1220,1749,1773]" pageId="30" pageNumber="31" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz 1966: 413</bibRefCitation>
|
||||
(
|
||||
<taxonomicName id="4C784D66FFBF2E3AFB4817F1FA45F79E" authorityName="Korge" authorityYear="1963" box="[1244,1468,1749,1773]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFB4817F1FA45F79E" box="[1244,1468,1749,1773]" italics="true" pageId="30" pageNumber="31">Quedius edwardsianus</emphasis>
|
||||
</taxonomicName>
|
||||
, redescription without new material or data); Coiffait and Sáiz, 1968: 365 (listed as
|
||||
<taxonomicName id="4C784D66FFBF2E3AFB851637FB14F658" authorityName="Korge" authorityYear="1963" box="[1041,1261,1811,1835]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFB851637FB14F658" box="[1041,1261,1811,1835]" italics="true" pageId="30" pageNumber="31">Quedius edwardsianus</emphasis>
|
||||
</taxonomicName>
|
||||
, without new data);
|
||||
<bibRefCitation id="EFE94B14FFBF2E3AFCAD1617FC03F638" author="Newton AF" box="[825,1018,1843,1867]" pageId="30" pageNumber="31" pagination="1 - 52" refId="ref37528" refString="Newton AF. Nomenclatural and taxonomic changes in Staphyliniformia (Coleoptera). Insecta Mundi 2017; 0595: 1 - 52." type="book chapter" year="2017">Newton 2017: 28</bibRefCitation>
|
||||
(as
|
||||
<taxonomicName id="4C784D66FFBF2E3AFBA41617FAD4F638" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[1072,1325,1843,1867]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFBA41617FAD4F638" box="[1072,1325,1843,1867]" italics="true" pageId="30" pageNumber="31">Loncovilius edwardsianus</emphasis>
|
||||
</taxonomicName>
|
||||
, moved from
|
||||
<taxonomicName id="4C784D66FFBF2E3AFCAD1676FC70F619" box="[825,905,1874,1898]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFCAD1676FC70F619" box="[825,905,1874,1898]" italics="true" pageId="30" pageNumber="31">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
to
|
||||
<taxonomicName id="4C784D66FFBF2E3AFC3D1676FBEEF619" box="[937,1047,1874,1898]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFC3D1676FBEEF619" box="[937,1047,1874,1898]" italics="true" pageId="30" pageNumber="31">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBF2E3BFCAD16A8FD47F3C4" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="materials_examined">
|
||||
<paragraph id="8BC736E5FFBF2E3BFCAD16A8FE87F30E" blockId="30.[825,1474,1931,1987]" lastBlockId="31.[113,763,144,638]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
|
||||
<emphasis id="B90CEAF7FFBF2E3AFCAD16A8FBB9F6D0" box="[825,1088,1931,1955]" italics="true" pageId="30" pageNumber="31">
|
||||
Type material.
|
||||
<materialsCitation id="3B103CB8FFBF2E3BFC4816A8FEB4F010" collectingDate="1926-12-04" collectingDateMax="1926-12-10" collectingDateMin="1926-12-04" collectionCode="FMNH" collectorName="M. Edwards" country="Chile" determinerName="Bernhauer" lastPageId="31" lastPageNumber="32" location="Casa Pangue" municipality="Llanquihue prov." pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
|
||||
<typeStatus id="54C38847FFBF2E3AFC4816A8FBC2F6D0" box="[988,1083,1932,1955]" pageId="30" pageNumber="31" type="lectotype">Lectotype</typeStatus>
|
||||
: (here designated):
|
||||
<specimenCount id="9D7EFD6CFFBF2E3AFAA416AFFA9AF6D0" box="[1328,1379,1931,1955]" pageId="30" pageNumber="31" type="male">male</specimenCount>
|
||||
, pinned, with genitalia in a separate microvial, with labels as follows: S.
|
||||
<collectingCountry id="F36F7675FFBE2E3BFFE511B4FF57F1DB" box="[113,174,144,168]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
|
||||
:
|
||||
<collectingMunicipality id="6BA3AC9FFFBE2E3BFF2811B4FE96F1D4" box="[188,367,144,168]" pageId="31" pageNumber="32">Llanquihue prov.</collectingMunicipality>
|
||||
F. and
|
||||
<collectorName id="268D5333FFBE2E3BFE5211B4FDB1F1DB" box="[454,584,144,168]" pageId="31" pageNumber="32">M. Edwards</collectorName>
|
||||
. B.M. 1927–63/
|
||||
<location id="8EA7603EFFBE2E3BFFE5118BFF02F1BB" LSID="urn:lsid:plazi:treatment:03D187F3FFBF2E04FC2414C0FE3AF0F0:8EA7603EFFBE2E3BFFE5118BFF02F1BB" box="[113,251,175,200]" country="Chile" municipality="Llanquihue prov." name="Casa Pangue" pageId="31" pageNumber="32">Casa Pangue</location>
|
||||
[-41.04, -71.87]
|
||||
<date id="FFC61025FFBE2E3BFE55118BFDBFF1B4" box="[449,582,175,199]" pageId="31" pageNumber="32" value="1926-12-04" valueMax="1926-12-10" valueMin="1926-12-04">
|
||||
<collectingDate id="EF82E9CDFFBE2E3BFE55118BFDBFF1B4" box="[449,582,175,199]" pageId="31" pageNumber="32" value="1926-12-04" valueMax="1926-12-10" valueMin="1926-12-04">4-10.xii.1926</collectingDate>
|
||||
</date>
|
||||
./
|
||||
<determinerName id="7E1BDA21FFBE2E3BFDC5118BFD3FF1B4" box="[593,710,175,199]" pageId="31" pageNumber="32">Bernhauer</determinerName>
|
||||
det. [unreadable]/Chicago NHM M. Bernhauer collection/
|
||||
<taxonomicName id="4C784D66FFBE2E3BFFE511CAFE9EF075" authority="Brh." authorityName="Bernhauer" authorityYear="2024" box="[113,359,238,262]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi" status="sp. nov.">Quedius edwardsi Brh.</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFFBE2E3BFEE411CAFE51F075" box="[368,424,238,262]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
|
||||
/
|
||||
<taxonomicName id="4C784D66FFBE2E3BFE2411CAFD72F075" authority="Bernhauer" authorityName="Bernhauer" authorityYear="2024" box="[432,651,238,262]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi">Edwardsi Bernhauer</taxonomicName>
|
||||
<taxonomicName id="4C784D66FFBE2E3BFD0211CAFD0BF075" box="[662,754,238,262]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
|
||||
/ LECTOTYPE
|
||||
<taxonomicName id="4C784D66FFBE2E3BFE811029FD3AF056" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[277,707,269,293]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFE811029FDEFF056" box="[277,534,269,293]" italics="true" pageId="31" pageNumber="32">Loncovilius edwardsianus</emphasis>
|
||||
(Korge, 1963)
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at
|
||||
<collectionCode id="ED69AE20FFBE2E3BFF6C1068FEB0F010" box="[248,329,332,355]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="31" pageNumber="32">FMNH</collectionCode>
|
||||
.
|
||||
</materialsCitation>
|
||||
</emphasis>
|
||||
<materialsCitation id="3B103CB8FFBE2E3BFEC61068FD02F353" collectingDate="1926-12-04" collectingDateMax="1926-12-10" collectingDateMin="1926-12-04" collectionCode="NHMUK" collectorName="M. Edwards" country="Chile" determinerName="M. K. Thayer" location="Casa Pangue" municipality="Llanquihue prov." pageId="31" pageNumber="32" specimenCode="NHMUK015024785" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFEC61068FE25F017" box="[338,476,332,356]" italics="true" pageId="31" pageNumber="32">
|
||||
<typeStatus id="54C38847FFBE2E3BFEC61068FE2EF017" box="[338,471,332,356]" pageId="31" pageNumber="32" type="paralectotype">Paralectotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
<specimenCount id="9D7EFD6CFFBE2E3BFE751068FDEDF017" box="[481,532,332,356]" pageId="31" pageNumber="32" type="male">male</specimenCount>
|
||||
, pinned, with genitalia attached to the card, with labels as follows: S.
|
||||
<collectingCountry id="F36F7675FFBE2E3BFDAB104FFD85F0F0" box="[575,636,363,387]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
|
||||
:
|
||||
<collectingMunicipality id="6BA3AC9FFFBE2E3BFD10104FFF5CF0D1" pageId="31" pageNumber="32">Llanquihue prov.</collectingMunicipality>
|
||||
F. and
|
||||
<collectorName id="268D5333FFBE2E3BFF6410AFFE97F0D0" box="[240,366,395,419]" pageId="31" pageNumber="32">M. Edwards</collectorName>
|
||||
. B. M. 1927–63/
|
||||
<location id="8EA7603EFFBE2E3BFD8F10AFFD5BF0D0" LSID="urn:lsid:plazi:treatment:03D187F3FFBF2E04FC2414C0FE3AF0F0:8EA7603EFFBE2E3BFD8F10AFFD5BF0D0" box="[539,674,395,419]" country="Chile" municipality="Llanquihue prov." name="Casa Pangue" pageId="31" pageNumber="32">Casa Pangue</location>
|
||||
[-41.04, -71.87]
|
||||
<date id="FFC61025FFBE2E3BFF43108EFEA6F0B1" box="[215,351,426,450]" pageId="31" pageNumber="32" value="1926-12-04" valueMax="1926-12-10" valueMin="1926-12-04">
|
||||
<collectingDate id="EF82E9CDFFBE2E3BFF43108EFEA6F0B1" box="[215,351,426,450]" pageId="31" pageNumber="32" value="1926-12-04" valueMax="1926-12-10" valueMin="1926-12-04">4-10.xii.1926</collectingDate>
|
||||
</date>
|
||||
./
|
||||
<taxonomicName id="4C784D66FFBE2E3BFEFF108EFE30F0B1" box="[363,457,426,450]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
|
||||
<taxonomicName id="4C784D66FFBE2E3BFE74108EFD77F0B1" authority="Bernh" authorityName="Bernhauer" authorityYear="2024" box="[480,654,426,450]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi" status="sp. nov.">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFE74108EFDCFF0B1" box="[480,566,426,450]" italics="true" pageId="31" pageNumber="32">edwardsi</emphasis>
|
||||
Bernh
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F578CFFBE2E3BFD33108EFD08F0B1" box="[679,753,426,450]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
|
||||
/ LECTOTYPE C.E.Tottenham of
|
||||
<taxonomicName id="4C784D66FFBE2E3BFE5110EDFD0BF092" authority="Bernh. Nec Sharp" authorityName="Bernh. Nec Sharp" authorityYear="1886" box="[453,754,457,481]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi">Q.edwardsi Bernh.Nec Sharp</taxonomicName>
|
||||
/
|
||||
<taxonomicName id="4C784D66FFBE2E3BFFE510CDFF30F372" box="[113,201,489,513]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
|
||||
ultor n. n. Tottenham/edwardsi repl. By
|
||||
<taxonomicName id="4C784D66FFBE2E3BFDE410CDFE6DF353" authority="Korge 1963 M. K. Nayer" authorityName="Korge" authorityYear="1963" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
edwardsianus Korge 1963
|
||||
<determinerName id="7E1BDA21FFBE2E3BFF6B132CFE6DF353" box="[255,404,520,544]" pageId="31" pageNumber="32">M. K. Thayer</determinerName>
|
||||
</taxonomicName>
|
||||
det. 2005/
|
||||
<specimenCode id="DBDE9E9EFFBE2E3BFD86132CFD0AF353" box="[530,755,520,544]" pageId="31" pageNumber="32">NHMUK015024785</specimenCode>
|
||||
/
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B103CB8FFBE2E3BFFE51303FE80F30E" collectionCode="NHM" pageId="31" pageNumber="32" specimenCount="1" typeStatus="paralectotype">
|
||||
<typeStatus id="54C38847FFBE2E3BFFE51303FEBCF34C" box="[113,325,551,575]" pageId="31" pageNumber="32">PARALECTOTYPE</typeStatus>
|
||||
<taxonomicName id="4C784D66FFBE2E3BFEC71303FD03F34C" authority="(Korge, 1963)" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[339,762,551,575]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFEC71303FDA8F34C" box="[339,593,551,575]" italics="true" pageId="31" pageNumber="32">Loncovilius edwardsianus</emphasis>
|
||||
(Korge, 1963)
|
||||
</taxonomicName>
|
||||
des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2023. Deposited at
|
||||
<collectionCode id="ED69AE20FFBE2E3BFEA31342FE80F30E" box="[311,377,614,637]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="31" pageNumber="32">NHM</collectionCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBE2E3BFFE513BBFD47F3C4" blockId="31.[113,702,671,695]" box="[113,702,671,695]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFFE513BBFE95F3C4" box="[113,364,671,695]" italics="true" pageId="31" pageNumber="32">Other material examined:</emphasis>
|
||||
Supporting Information, File S7.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBE2E3BFFE513FDFDDEF21E" pageId="31" pageNumber="32" type="diagnosis">
|
||||
<paragraph id="8BC736E5FFBE2E3BFFE513FDFDDEF21E" blockId="31.[113,763,729,878]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFFE513FDFF2EF383" box="[113,215,729,752]" italics="true" pageId="31" pageNumber="32">Diagnosis:</emphasis>
|
||||
Nuchal ridge absent dorsally. Pronotum outline hexagonal. Elytra yellowish-brown to brown-black with greenish, bluish, or purplish iridescence. Protibiae without sexual dimorphism, both males and females without laterodorsal row of thick spines. Aedeagus as in
|
||||
<figureCitation id="13432A60FFBE2E3BFE1B1272FDD8F21D" box="[399,545,854,878]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="31" pageNumber="32">Figure 10L–N</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBE2E3BFFE512B7FAA4F498" pageId="31" pageNumber="32" type="description">
|
||||
<paragraph id="8BC736E5FFBE2E3BFFE512B7FE0BF534" blockId="31.[113,765,915,1973]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFFE512B7FF11F2D9" box="[113,232,915,938]" italics="true" pageId="31" pageNumber="32">Description:</emphasis>
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFBE2E3BFE0D12B2FE4AF2D9" box="[409,435,918,938]" count="1" pageId="31" pageNumber="32" type="male">♂</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFBE2E3BFD3F12B7FD47F2D9" box="[683,702,915,938]" italics="true" pageId="31" pageNumber="32">N</emphasis>
|
||||
= 6]: FBL = 3.95–4.21 (4.1); TL = 7.92–8 (7.95); HW = 1–1.11 (1.05); HL = 1.13–1.25 (1.19); HW/HL = 0.83–0.92 (0.88); PW = 1.15–1.24 (1.21); PL = 1.15–1.25 (1.21); PW/ PL = 0.95–1.03 (1); EW = 1.76–1.9 (1.81); EL = 1.67–1.71 (1.69); PW/HW = 1.11–1.21 (1.15).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBE2E3BFF19156BFE5AF470" blockId="31.[113,765,915,1973]" pageId="31" pageNumber="32">
|
||||
Measurements
|
||||
<specimenCount id="9D7EFD6CFFBE2E3BFEBC1576FEBBF515" box="[296,322,1106,1126]" count="1" pageId="31" pageNumber="32" type="female">♀</specimenCount>
|
||||
[min–max (average);
|
||||
<emphasis id="B90CEAF7FFBE2E3BFDB6156BFDCCF515" box="[546,565,1103,1126]" italics="true" pageId="31" pageNumber="32">N</emphasis>
|
||||
= 3]: FBL = 4.47– 4.76 (4.59); TL = 7.32–8.56 (7.76); HW = 1.16–1.21 (1.18); HL = 1.3–1.36 (1.33); HW/HL = 0.87–0.9 (0.89); PW = 1.33–1.5 (1.4); PL = 1.31–1.44 (1.37); PW/PL = 1.01– 1.04 (1.02); EW = 1.98–2.13 (2.06); EL = 1.83–1.96 (1.9); PW/HW = 1.15–1.24 (1.18).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBE2E3BFF19142FFF39F4AC" blockId="31.[113,765,915,1973]" pageId="31" pageNumber="32">Head and pronotum dark brown to black with metallic blue overtones; antennae, mouthparts, pronotum at margins and legs yellowish-brown; elytra yellowish-brown to brown-black with greenish, bluish, or purplish iridescence, sometimes darker around mesocutellum and at suture; abdominal tergites and sternites brown-black or dark reddish-brown with pale posterior margin.</paragraph>
|
||||
<paragraph id="8BC736E5FFBE2E3BFF1914C2FC67F092" blockId="31.[113,765,915,1973]" lastBlockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">
|
||||
Head elongate; dorsally and ventrally with iridescence and with micropunctures and microsculpture of transverse isodiametric meshes, without coarse non-setiferous punctures; posterior angles indistinct with moderate setiferous punctures of medium and small-size. Eyes small (EYL/HL = x = 0.44), from 1.18 to 1.46 times as long as temples (in lateral view) in males and
|
||||
<quantity id="4C809B00FFBE2E3BFF351786FEC2F7C9" box="[161,315,1698,1722]" metricMagnitude="-2" metricUnit="m" metricValue="2.3876" metricValueMax="2.6416" metricValueMin="2.1336" pageId="31" pageNumber="32" unit="in" value="0.94" valueMax="1.04" valueMin="0.84">0.84 to 1.04 in</quantity>
|
||||
females; distance between eyes about 1.13 times as long as length of eye in males and 1.44 times in females. Antennomere 3 moderately longer than antennomere 2; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a11 from 1.52 to 1.75 times as long as a
|
||||
<quantity id="4C809B00FFBE2E3BFD5D163BFD02F644" box="[713,763,1823,1847]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="31" pageNumber="32" unit="in" value="10.0">10 in</quantity>
|
||||
males and from
|
||||
<quantity id="4C809B00FFBE2E3BFE87161BFE6EF624" box="[275,407,1855,1879]" metricMagnitude="-2" metricUnit="m" metricValue="3.3401" metricValueMax="3.5559999999999996" metricValueMin="3.1242" pageId="31" pageNumber="32" unit="in" value="1.315" valueMax="1.4" valueMin="1.23">1.23 to 1.4 in</quantity>
|
||||
females. Basal and parocular punctures usually double; posterior frontal puncture located anterior to temporal puncture; without small setose punctures between frontoclypeal and anterior frontal punctures; ventral basal ridge more or less straight in checkmark shape, ending distinctly far from gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present only laterally, not fused with infraorbital ridge; gula with conspicuous isodiametric microsculpture medially, gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBE2E3BFCD110CDFB48F3CF" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">Pronotum about as wide as long, convex, hexagonal, with two PPDS, without APP, without SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to two or even up to six times as long as, diameter of punctures. Mesosternum with six macrosetae arranged in a row medially. Metasternum with several macrosetae.</paragraph>
|
||||
<paragraph id="8BC736E5FFBE2E3BFCD113E0FA63F2EB" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 1–4 with pale adhesive setae, without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin bilobed.</paragraph>
|
||||
<paragraph id="8BC736E5FFBE2E3BFCD112BBFBC0F463" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">
|
||||
Abdominal tergites III and IV with only a few rows of coarse, sparse punctures, with large impunctate area; tergite VIII not emarginate medio-apically. Sternite III with basal transverse carina usually evenly converging at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with U-shaped emargination medially (
|
||||
<figureCitation id="13432A60FFBE2E3BFB691518FAADF527" box="[1277,1364,1084,1108]" captionStart="Figure 14" captionStartId="22.[129,194,1868,1892]" captionTargetBox="[168,1434,144,1840]" captionTargetId="figure-7@22.[168,1434,144,1840]" captionTargetPageId="22" captionText="Figure 14. Male sternite VIII of Loncovilius species. A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="31" pageNumber="32">Fig. 14E</figureCitation>
|
||||
); female sternite VIII without emargination; male sternite IX without emargination medially, with rounded apex, its basal portion shorter than distal portion; male tergite X apically subtruncate (
|
||||
<figureCitation id="13432A60FFBE2E3BFCA1159DFC72F5A2" box="[821,907,1209,1233]" captionStart="Figure 12" captionStartId="20.[129,194,1869,1893]" captionTargetBox="[168,1434,145,1841]" captionTargetId="figure-7@20.[168,1434,145,1841]" captionTargetPageId="20" captionText="Figure 12. Male tergite X of Loncovilius species. A, L. germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F,L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="31" pageNumber="32">Fig. 12E</figureCitation>
|
||||
); female tergite X with apical margin distinctly subacute to acute (
|
||||
<figureCitation id="13432A60FFBE2E3BFC5215FDFBE3F582" box="[966,1050,1241,1265]" captionStart="Figure 13" captionStartId="21.[113,178,1868,1892]" captionTargetBox="[186,1368,154,1839]" captionTargetId="figure-6@21.[153,1419,144,1840]" captionTargetPageId="21" captionText="Figure 13. Female tergite X of Loncovilius species.A, Loncovilius germaini. B, L. hammondi. C, L. impunctus. D, L. variabilis. E, L. edwardsianus. F, L. cantharoides. G, L. semiflavus. H, L. barclayi. I, L. lividipennis. J, L. carlsbergi." pageId="31" pageNumber="32">Fig. 13E</figureCitation>
|
||||
); ovipositor, its second gonocoxite with three macrosetae medially.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E5FFBE2E3BFCD11433FAA4F498" blockId="31.[809,1461,144,1516]" pageId="31" pageNumber="32">
|
||||
Aedeagus as in
|
||||
<figureCitation id="13432A60FFBE2E3BFC661433FB72F45C" box="[1010,1163,1303,1327]" captionStart="Figure 10" captionStartId="18.[128,193,1867,1891]" captionTargetBox="[195,1408,147,1835]" captionTargetId="figure-8@18.[190,1412,143,1839]" captionTargetPageId="18" captionText="Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally." pageId="31" pageNumber="32">Figure 10L–N</figureCitation>
|
||||
; its total length 1.25–1.31. Median lobe rod-like, its apex in parameral view slightly V-shaped, emarginate, internal sac with pair of long inverted L-shaped copulatory sclerites. Paramere distinctly produced over apex of median lobe; wider than median lobe; converging to a round apex; with small translucent peg setae; in lateral view almost straight, curved away from median lobe apically.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBE2E3BFCBE1729FC31F78C" pageId="31" pageNumber="32" type="distribution">
|
||||
<paragraph id="8BC736E5FFBE2E3BFCBE1729FC31F78C" blockId="31.[810,1460,1549,1792]" pageId="31" pageNumber="32">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFCBE1729FAE3F756" box="[810,1306,1549,1573]" italics="true" pageId="31" pageNumber="32">
|
||||
Distribution and habitat:
|
||||
<taxonomicName id="4C784D66FFBE2E3BFBB01729FAE3F756" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[1060,1306,1549,1573]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">Loncovilius edwardsianus</taxonomicName>
|
||||
</emphasis>
|
||||
is known from
|
||||
<collectingCountry id="F36F7675FFBE2E3BFCBE1708FC9DF737" box="[810,868,1580,1604]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
|
||||
, from the
|
||||
<collectingRegion id="49BCF807FFBE2E3BFC471708FBC5F737" box="[979,1084,1580,1604]" country="Chile" name="Araucania" pageId="31" pageNumber="32">Araucanía</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBE2E3BFBDC1708FB71F737" box="[1096,1160,1580,1604]" country="Chile" name="Aysen" pageId="31" pageNumber="32">Aysén</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBE2E3BFB001708FB23F737" box="[1172,1242,1580,1604]" country="Chile" name="Biobio" pageId="31" pageNumber="32">Biobío</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="49BCF807FFBE2E3BFB721708FAABF737" box="[1254,1362,1580,1604]" country="Chile" name="Los Lagos" pageId="31" pageNumber="32">Los Lagos</collectingRegion>
|
||||
, and
|
||||
<collectingRegion id="49BCF807FFBE2E3BFA191708FCA0F710" country="Chile" name="Los Rios" pageId="31" pageNumber="32">Los Ríos</collectingRegion>
|
||||
regions. In addition, it occurs in the westernmost portions of the
|
||||
<collectingRegion id="49BCF807FFBE2E3BFCFA174FFC47F7F0" box="[878,958,1643,1667]" country="Argentina" name="Chubut" pageId="31" pageNumber="32">Chubut</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="49BCF807FFBE2E3BFC66174FFBA7F7F0" box="[1010,1118,1643,1667]" country="Argentina" name="Rio Negro" pageId="31" pageNumber="32">Río Negro</collectingRegion>
|
||||
provinces of
|
||||
<collectingCountry id="F36F7675FFBE2E3BFB7E174FFAABF7F0" box="[1258,1362,1643,1667]" name="Argentina" pageId="31" pageNumber="32">Argentina</collectingCountry>
|
||||
. The species occurs from
|
||||
<quantity id="4C809B00FFBE2E3BFC4B17AEFB84F7D2" box="[991,1149,1674,1698]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="12.0" metricValueMin="2.0" pageId="31" pageNumber="32" unit="m" value="700.0" valueMax="1200.0" valueMin="200.0">200 to 1200 m</quantity>
|
||||
elevation; it is reported from wet sclerophyll and open
|
||||
<taxonomicName id="4C784D66FFBE2E3BFBA4178DFB59F7B2" box="[1072,1184,1705,1729]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="genus">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFBA4178DFB59F7B2" box="[1072,1184,1705,1729]" italics="true" pageId="31" pageNumber="32">Nothofagus</emphasis>
|
||||
</taxonomicName>
|
||||
forests along the Valdivian temperate forest ecoregion. It has been collected by sifting leaf litter and moss.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656EFFBE2E04FCBE1605FE3AF0F0" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="discussion">
|
||||
<paragraph id="8BC736E5FFBE2E04FCBE1605FE3AF0F0" blockId="31.[809,1459,1825,1975]" lastBlockId="32.[127,780,144,387]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFCBE1605FB78F64A" box="[810,1153,1825,1849]" italics="true" pageId="31" pageNumber="32">
|
||||
Remarks:
|
||||
<taxonomicName id="4C784D66FFBE2E3BFC1A1605FB78F64A" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[910,1153,1825,1849]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">Loncovilius edwardsianus</taxonomicName>
|
||||
</emphasis>
|
||||
was described under the preoccupied name
|
||||
<taxonomicName id="4C784D66FFBE2E3BFC521665FB94F62A" authorityName="Bernh. Nec Sharp" authorityYear="1886" box="[966,1133,1857,1881]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsi">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFC521665FB94F62A" box="[966,1133,1857,1881]" italics="true" pageId="31" pageNumber="32">Quedius edwardsi</emphasis>
|
||||
</taxonomicName>
|
||||
based on an unspecified number of
|
||||
<typeStatus id="54C38847FFBE2E3BFCD51644FC63F60B" box="[833,922,1888,1912]" pageId="31" pageNumber="32" type="syntype">syntypes</typeStatus>
|
||||
from Casa Pangue in Llanquihue Province in Southern
|
||||
<collectingCountry id="F36F7675FFBE2E3BFCBD165BFC9BF6E4" box="[809,866,1919,1943]" name="Chile" pageId="31" pageNumber="32">Chile</collectingCountry>
|
||||
. Korge (1963) replaced the name with
|
||||
<taxonomicName id="4C784D66FFBE2E3BFB7316A4FA7AF6E4" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[1255,1411,1919,1943]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="edwardsianus">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFB7316A4FA7AF6E4" box="[1255,1411,1919,1943]" italics="true" pageId="31" pageNumber="32">Q. edwardsianus</emphasis>
|
||||
</taxonomicName>
|
||||
and, interestingly, this species was treated in the genus
|
||||
<taxonomicName id="4C784D66FFBE2E3BFAD516BBFA69F6C4" box="[1345,1424,1951,1975]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FFBE2E3BFAD516BBFA69F6C4" box="[1345,1424,1951,1975]" italics="true" pageId="31" pageNumber="32">Quedius</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="EFE94B14FF812E04FF1511B4FE93F1DB" author="Coiffait H & Saiz F" box="[129,362,144,168]" pageId="32" pageNumber="33" pagination="385 - 414" refId="ref36501" refString="Coiffait H, Saiz F. Les Quediini du Chili (Col. Staphylinidae). Annales de la Societe Entomologique de France (n. ser.) 1966; 2: 385 - 414." type="journal article" year="1966">Coiffait and Sáiz (1966</bibRefCitation>
|
||||
, 1968) who neither studied the type material nor reported any new material. Presumably, all mentioned authors knew this species only from literature because even
|
||||
<bibRefCitation id="EFE94B14FF812E04FD7711EBFF30F075" author="Saiz F" pageId="32" pageNumber="33" pagination="364 - 92" refId="ref38248" refString="Saiz F. Sur les Quediini du Chili (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 1971; 106: 364 - 92." type="journal article" year="1971">Sáiz (1971)</bibRefCitation>
|
||||
did not mention it in the fauna of
|
||||
<collectingCountry id="F36F7675FF812E04FDB311CAFD99F075" box="[551,608,238,262]" name="Chile" pageId="32" pageNumber="33">Chile</collectingCountry>
|
||||
. The species was transferred to the genus
|
||||
<taxonomicName id="4C784D66FF812E04FEFF1029FE2FF056" box="[363,470,269,293]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B90CEAF7FF812E04FEFF1029FE2FF056" box="[363,470,269,293]" italics="true" pageId="32" pageNumber="33">Loncovilius</emphasis>
|
||||
</taxonomicName>
|
||||
much later by
|
||||
<bibRefCitation id="EFE94B14FF812E04FDF0102AFCFAF056" author="Newton AF" box="[612,771,269,293]" pageId="32" pageNumber="33" pagination="1 - 52" refId="ref37528" refString="Newton AF. Nomenclatural and taxonomic changes in Staphyliniformia (Coleoptera). Insecta Mundi 2017; 0595: 1 - 52." type="book chapter" year="2017">Newton (2017)</bibRefCitation>
|
||||
. To fix the identity of the species, a male from the Bernhauer’s collection at the FMNH, which is clearly a
|
||||
<typeStatus id="54C38847FF812E04FD971068FDADF017" box="[515,596,332,356]" pageId="32" pageNumber="33" type="syntype">syntype</typeStatus>
|
||||
, is here designated as a
|
||||
<typeStatus id="54C38847FF812E04FF3C104FFEFCF0F0" box="[168,261,363,387]" pageId="32" pageNumber="33" type="lectotype">lectotype</typeStatus>
|
||||
of
|
||||
<emphasis id="B90CEAF7FF812E04FEB01048FE3AF0F0" box="[292,451,363,387]" italics="true" pageId="32" pageNumber="33">
|
||||
<taxonomicName id="4C784D66FF812E04FEB01048FE47F0F0" baseAuthorityName="Korge" baseAuthorityYear="1963" box="[292,446,363,387]" class="Insecta" family="Staphylinidae" genus="Loncovilius" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="edwardsianus">L. edwardsianus</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
184
data/22/E1/F9/22E1F903849F59E9B57DA8AA89046C6D.xml
Normal file
184
data/22/E1/F9/22E1F903849F59E9B57DA8AA89046C6D.xml
Normal file
|
|
@ -0,0 +1,184 @@
|
|||
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<mods:titleInfo id="EB7E22A099DE85B1A013BA6107E3731B">
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<mods:title id="AC03D5A28E46B34A749C3EC35E19075E">New wasps of † Falsiformicidae from mid-Cretaceous Kachin amber</mods:title>
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|
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<mods:affiliation id="06FC4F9AAB800F29763D6D0A3B80DBAE">A. A. Borissiak Palaeontological Institute, Russian Academy of Sciences, Moscow, 117647, Russia</mods:affiliation>
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<mods:namePart id="3BDB2C50DC73D12FB17575E39BDA0AF5">Vilhelmsen, Lars</mods:namePart>
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<mods:affiliation id="69530E142556AF70EE41F881875CA242">College of Life Sciences, Capital Normal University, Beijing, 100048, China</mods:affiliation>
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<mods:number id="A1187AA6C0D8A03B1E2F5CD745B05815">2025-01-03</mods:number>
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<treatment id="22E1F903849F59E9B57DA8AA89046C6D" ID-arpha="22E1F903-849F-59E9-B57D-A8AA89046C6D" LSID="urn:lsid:plazi:treatment:22E1F903849F59E9B57DA8AA89046C6D" httpUri="http://treatment.plazi.org/id/22E1F903849F59E9B57DA8AA89046C6D">
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||||
<subSubSection id="077082BACB38840BDE870E58E0FEEEDD" type="nomenclature">
|
||||
<paragraph id="200D1944FA25E24DD72FD05AB039D9AA">
|
||||
<heading id="7FF25799D2B1FB60B556D27E29E7A50D" reason="label">Genus</heading>
|
||||
<taxonomicName id="CE4A2B956E5CCBE8D9FCEFE754DF432F" authority="Cockx & McKellar, 2016" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="16CF1063C563E152470AC9FD54582073" italics="true">Siccibythus</emphasis>
|
||||
<bibRefCitation id="455C7D37A7D0CB27C5573564C029D12A" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD & McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx & McKellar, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ELLAE" type="type species">
|
||||
<paragraph id="F1EFF210974AA5521F10D5DAE57A9ED1">
|
||||
<heading id="600FED4707A0002563B99720265889B2" reason="title">
|
||||
<typeStatus id="95E9CDDE0B33E1E98AAA8B462B9BA28A">Type</typeStatus>
|
||||
species.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="CB3623490BF04C4CB7696222AEE01D2C">
|
||||
<taxonomicName id="9B0F5C689D0BFD74E929822B43ADB8BE" authority="Cockx & McKellar, 2016" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="musculosus">
|
||||
<emphasis id="F6F22818D3A6D5ED66F30B663CB73B69" italics="true">Siccibythus musculosus</emphasis>
|
||||
<bibRefCitation id="C75CF54C484AF4D2D61097BEE206D97D" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD & McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx & McKellar, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E3LAE" type="Species included">
|
||||
<paragraph id="914B5145DF3D3FD06D2CFBBD7DF6F83F">
|
||||
<heading id="39DDBEA0B3CA2BA3D1CB745820AEF5F9" reason="title">Species included.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7428A7A30362209D2B7EBFED50BF3661">
|
||||
<taxonomicName id="6D1B2709CFC7C16AD31EC283BDD3C524" authority="Cockx & McKellar, 2016" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="musculosus">
|
||||
<emphasis id="C599A4735E8542A614FE32CA97DDEBD6" italics="true">Siccibythus musculosus</emphasis>
|
||||
<bibRefCitation id="D9F34B8E83B60F8E2F819EBFA1238A7B" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD & McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx & McKellar, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="03C2A99D1CEF74BA0255D9D315FE6867" authority="Rasnitsyn, Zhang, Muller & Zhang, 2020" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="martynovae">
|
||||
<emphasis id="16053B694982E448ACB936A621B4FC4B" italics="true">S. martynovae</emphasis>
|
||||
<bibRefCitation id="DE20A915209D4BA09210798350E4CC5A" DOI="10.11646/palaeoentomology.3.6.10" author="Rasnitsyn AP & Zhang Q & Müller P & Zhang HC" journalOrPublisher="Palaeoentomology" pagination="582-596" refId="B20" refString="Rasnitsyn AP, Zhang Q, Muller P, Zhang HC (2020) On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.). Palaeoentomology 3: 582 - 596. https: // doi. org / 10.11646 / palaeoentomology. 3.6. 10" title="On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.)." volume="3" year="2020">Rasnitsyn, Zhang, Müller & Zhang, 2020</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="150BE709757E9FFBCC0D7B1D6BABFF1F" authority="Rasnitsyn, Zhang, Muller & Zhang, 2020" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="oculatus">
|
||||
<emphasis id="06BAD4BE6546023F1C3BAA9A40AD035C" italics="true">S. oculatus</emphasis>
|
||||
<bibRefCitation id="D5E1C31F544D3D65E72EBD10196ABD07" DOI="10.11646/palaeoentomology.3.6.10" author="Rasnitsyn AP & Zhang Q & Müller P & Zhang HC" journalOrPublisher="Palaeoentomology" pagination="582-596" refId="B20" refString="Rasnitsyn AP, Zhang Q, Muller P, Zhang HC (2020) On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.). Palaeoentomology 3: 582 - 596. https: // doi. org / 10.11646 / palaeoentomology. 3.6. 10" title="On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.)." volume="3" year="2020">Rasnitsyn, Zhang, Müller & Zhang, 2020</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="0B8038E397278BE98B391CBBAB00C5FF" authority="Rasnitsyn, Zhang, Muller & Zhang, 2020" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ohmkuhnlei">
|
||||
<emphasis id="3BB5166AB91140DCD1E68234792A13EB" italics="true">S. ohmkuhnlei</emphasis>
|
||||
<bibRefCitation id="1FD4BF5908DBAA23CECE311B1DCD328C" DOI="10.11646/palaeoentomology.3.6.10" author="Rasnitsyn AP & Zhang Q & Müller P & Zhang HC" journalOrPublisher="Palaeoentomology" pagination="582-596" refId="B20" refString="Rasnitsyn AP, Zhang Q, Muller P, Zhang HC (2020) On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.). Palaeoentomology 3: 582 - 596. https: // doi. org / 10.11646 / palaeoentomology. 3.6. 10" title="On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.)." volume="3" year="2020">Rasnitsyn, Zhang, Müller & Zhang, 2020</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="3E7F3B2F668E7126D2D31B04FDF298BE" authority="Rasnitsyn, Zhang, Muller & Zhang, 2020" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pallidus">
|
||||
<emphasis id="D9BEBFB33DF76197CCC3EE7089D677BF" italics="true">S. pallidus</emphasis>
|
||||
<bibRefCitation id="D6F04D00E21582FD22AE8394B173FEDA" DOI="10.11646/palaeoentomology.3.6.10" author="Rasnitsyn AP & Zhang Q & Müller P & Zhang HC" journalOrPublisher="Palaeoentomology" pagination="582-596" refId="B20" refString="Rasnitsyn AP, Zhang Q, Muller P, Zhang HC (2020) On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.). Palaeoentomology 3: 582 - 596. https: // doi. org / 10.11646 / palaeoentomology. 3.6. 10" title="On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.)." volume="3" year="2020">Rasnitsyn, Zhang, Müller & Zhang, 2020</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="21FE28BEED13652F739A2679C89CEDFE" authority="Rasnitsyn, Zhang, Muller & Zhang, 2020" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="paulus">
|
||||
<emphasis id="3F307E414B58B602FF45AD08B31C5F45" italics="true">S. paulus</emphasis>
|
||||
<bibRefCitation id="9326249568E5320D4E897F9616F5544C" DOI="10.11646/palaeoentomology.3.6.10" author="Rasnitsyn AP & Zhang Q & Müller P & Zhang HC" journalOrPublisher="Palaeoentomology" pagination="582-596" refId="B20" refString="Rasnitsyn AP, Zhang Q, Muller P, Zhang HC (2020) On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.). Palaeoentomology 3: 582 - 596. https: // doi. org / 10.11646 / palaeoentomology. 3.6. 10" title="On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.)." volume="3" year="2020">Rasnitsyn, Zhang, Müller & Zhang, 2020</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="13DDC84942B801AFC9E841EE4CD174D9" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
|
||||
<emphasis id="C1B858EC4B353F9A11FB1FBF5F920413" italics="true">S. aristovi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="ED4EFF9D495B71D3CA875A70992570E6" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="B46286E8FC496E91A7C1245B05A70542" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
|
||||
<emphasis id="A5F125C8F9087CAB79C06D5DA6DEDF5F" italics="true">S. robustus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="F25BA940CF630B3CCB8F48C2C32E91E7" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E1OAE" type="Revised diagnosis">
|
||||
<paragraph id="E2087BA92199D27BAE978B2359D19366">
|
||||
<heading id="1D799EDCED70E9528A50AD69A67EC76F" reason="title">Revised diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="A42282A38E4CB458E51E193FFAA26DB0">
|
||||
Propodeum weakly sculptured with longitudinal carinae; antenna with 13 segments in male,
|
||||
<quantity id="C1B0153F35A5944C660C080DC39C84DC" metricMagnitude="-1" metricUnit="m" metricValue="3.048" unit="in" value="12.0">12 in</quantity>
|
||||
female, scape elongate and flattened, pedicel weakly pyriform; three or four labial palpomeres; fore wing with no cells closed with tubular veins in its middle and distal part, vein 1 m-cu at most only partially tubular; vein
|
||||
<collectionCode id="097927B7E8E15649F37E6B20549F0D81" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
|
||||
1 absent apical to pterostigma, and apical parts of M, M + Cu and A (distally of Cu-a) spectral. Fore tarsus and mid tarsus with large, bilobed arolium, smaller arolium present on hind tarsus, claws simple.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E6OAE" type="remarks">
|
||||
<paragraph id="A3B2113E96560815C2BBF75B3B47DB3D">
|
||||
<heading id="544A9689B5B4774B42DDA13A540A8FDA" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="14D40D6339CD2AD5F14C76697DBB501F">
|
||||
†
|
||||
<taxonomicName id="23EB8E79A75BB2B30516C339A0F9F250" authority="Cockx & McKellar, 2016" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="69F0B7B0DF28849444A732AD7D51585B" italics="true">Siccibythus</emphasis>
|
||||
<bibRefCitation id="3B77115D08F1F3FD4BB9F829ADA3A3B7" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD & McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx & McKellar, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
differs †
|
||||
<taxonomicName id="05F173914A3F8EB47C15C8274A181CC0" authority="Rasnitsyn, 1975" authorityName="Rasnitsyn" authorityYear="1975" class="Insecta" family="Falsiformicidae" genus="Falsiformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="67E2F21373D580887056FBF42F0B0ADC" italics="true">Falsiformica</emphasis>
|
||||
<bibRefCitation id="A92EEE848CA438BE1F483A9E1B01B37B" author="Rasnitsyn AP" journalOrPublisher="Transaction of the Paleontological Institute, Academy of Sciences of the USSR" pagination="1-134" refId="B19" refString="Rasnitsyn AP (1975) Hymenoptera Apocrita of Mesozoic. Transaction of the Paleontological Institute, Academy of Sciences of the USSR 147: 1 - 134." title="Hymenoptera Apocrita of Mesozoic." volume="147" year="1975">Rasnitsyn, 1975</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and †
|
||||
<emphasis id="14E4748F7C16BE4BE546828F3ACC6E51" italics="true">Neophenax</emphasis>
|
||||
<bibRefCitation id="32E23E632B001438AAFFFD85123D386E" author="Engel MS" journalOrPublisher="Entomologist’s Monthly Magazine" pagination="157-168" refId="B5" refString="Engel MS (2022) The first New World Falsiformicidae (Hymenoptera): a new genus in Upper Cretaceous amber from New Jersey. Entomologist's Monthly Magazine 158: 157 - 168. https: // doi: 10.31184 / M 00138908.1583. 4100" title="The first New World Falsiformicidae (Hymenoptera): a new genus in Upper Cretaceous amber from New Jersey." url="https://doi: 10.31184/M00138908.1583.4100" volume="158" year="2022">Engel, 2022</bibRefCitation>
|
||||
by having fore wing with vein 1 m-cu at most partially tubular and pronotum with dorsal surface longer than wide or, rarely, subquadrate. †
|
||||
<taxonomicName id="82FF9A82ABA797846D269752C5D1A595" authority="Cockx & McKellar, 2016" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="662D3D20BD6ED36EB9F7197CE76E3171" italics="true">Siccibythus</emphasis>
|
||||
<bibRefCitation id="E6ED324532A9AAE2062276CF5ABEB6E9" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD & McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx & McKellar, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
differs from †
|
||||
<emphasis id="47B9F2BD702BA06AD17E19F72B4CFFF3" italics="true">Falsiformix</emphasis>
|
||||
<bibRefCitation id="2E82BC1013E720A7063FEEEA003CC959" DOI="10.1016/j.cretres.2024.106057" author="Zhang Q & Jia N & Li YZ & Wang JX & Rasnitsyn AP" refId="B32" refString="Zhang Q, Jia N, Li YZ, Wang JX, Rasnitsyn AP (2024) New genus of falsiformicid wasps (Hymenoptera, Vespoidea: Falsiformicidae) with brachypterous male from middle Cretaceous Kachin amber in Myanmar. Cretaceous Research. https: // doi. org / 10.1016 / j. cretres. 2024.106057" year="2024">
|
||||
Zhang & Rasnitsyn,
|
||||
<quantity id="D92841FC04F7E2021127A7A74A92D3D8" metricMagnitude="1" metricUnit="m" metricValue="5.14096" unit="in" value="2024.0">2024 in</quantity>
|
||||
</bibRefCitation>
|
||||
having a fore wing longer than mesosoma.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
325
data/54/BF/FD/54BFFD0035125A8E8C223856B09A20C9.xml
Normal file
325
data/54/BF/FD/54BFFD0035125A8E8C223856B09A20C9.xml
Normal file
|
|
@ -0,0 +1,325 @@
|
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<mods:namePart id="D532FDCB9B6925B7FB85FB83D199E110">Wang, Zhen</mods:namePart>
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<mods:affiliation id="AF533B7EEC2ABE0317DDF19EECA7B409">A. A. Borissiak Palaeontological Institute, Russian Academy of Sciences, Moscow, 117647, Russia</mods:affiliation>
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|
||||
<taxonomicName id="98E9670D0E28D688FD65BFB3D130C522" authority="Wang, Rasnitsyn, Perkovsky & Vilhelmsen" authorityName="Wang, Rasnitsyn, Perkovsky & Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
|
||||
<emphasis id="A93B2FEEBB724234BF0F665FB771FC30" italics="true">Siccibythus robustus</emphasis>
|
||||
Wang, Rasnitsyn, Perkovsky & Vilhelmsen
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="ADB335495D60C33EB041C8DB77EE196D" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ERRAE" type="diagnosis">
|
||||
<paragraph id="270A5371052612084E86955A94028F69">
|
||||
<heading id="CF3866132522EE16DED9F6200EFE663B" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="62365BF98B298FA68D53B04A8681B50E">
|
||||
The length of the scape is less than 0.3 times the length of the flagellum. Fore wing with 2 Rs distal of r-rs several times as long as r-rs; Cu nebulous. Fore tibial spur long, thin and acute apically. Metabasitarsus longer than tarsomeres 2–4 combined. Propodeum outline angular. Anterior outline of 1
|
||||
<superScript id="7D273D01FD09119CECCFE3500725BAE9">st</superScript>
|
||||
metasomal tergum somewhat angular.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EZRAE" type="etymology">
|
||||
<paragraph id="1371E3E9B64602FDDF81B5D881025A6D">
|
||||
<heading id="25DE3CF6C1E1E6B356E123FA8E47FFB6" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="CD4B1763213D9097C3600F5FB13B6346">
|
||||
The species is named after the Latin adjective
|
||||
<emphasis id="406EC3B919B06489CF788D93B73E0A8C" italics="true">robustus</emphasis>
|
||||
, meaning “ thick ” and referring to the thick fore femur.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EJSAE" type="type material">
|
||||
<paragraph id="1AB59F69F647F37D93FA52DBD3EC99AB">
|
||||
<heading id="92318F059220CEDD2BFEC6D42D3B81D2" reason="title">Type material.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="152F21776301F101E33D3838F8F14730">
|
||||
<emphasis id="6F81167E535749A0996FB758A6FA3702" bold="true" italics="true">
|
||||
<typeStatus id="A226A76C6D0DEB41C0EE82E5CC8250A9" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
• female. No. CNU-HYM-MA 2016212. A well-preserved and complete specimen (Figs
|
||||
<figureCitation id="F0AE785FF8CCA057DC6D4F6C6168F3A0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A habitus in lateral view B head in lateral view C fore wing venation." figureDoi="10.3897/jhr.98.136200.figure1" httpUri="https://binary.pensoft.net/fig/1213364">1</figureCitation>
|
||||
,
|
||||
<figureCitation id="7C07252D79C0AC43468C73734DEA1BA4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2</figureCitation>
|
||||
).
|
||||
<emphasis id="3503CFC9F8C882580586022A6014DF54" bold="true" italics="true">
|
||||
<typeStatus id="9B95C509049CDFEE7206A0CE3F80490E" type="paratype">Paratype</typeStatus>
|
||||
</emphasis>
|
||||
• male. No. CNU-HYM-MA 2016213. A well-preserved and complete specimen (Figs
|
||||
<figureCitation id="08BF8AB8B5EB3BD09E0BC257DDC4A8A3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photographs of Siccibythus robustus sp. nov. (Male paratype; CNU-HYM-MA- 2016213) A habitus in lateral view B head in lateral view." figureDoi="10.3897/jhr.98.136200.figure3" httpUri="https://binary.pensoft.net/fig/1213366">3</figureCitation>
|
||||
,
|
||||
<figureCitation id="559707BA08738F3563BA48B368E6872E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photographs of Siccibythus robustus sp. nov. (Male paratype; CNU-HYM-MA- 2016213) A mesosoma in dorsal view B fore wing venation C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure4" httpUri="https://binary.pensoft.net/fig/1213367">4</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="EC9CFF30FB197EF612DFD47327D9A658" ID-DOI="10.3897/jhr.98.136200.figure1" ID-arpha="A42DFDDD-6CFF-5B6F-9EFB-E456B1085D1F" httpUri="https://binary.pensoft.net/fig/1213364" startId="F1">
|
||||
<paragraph id="DB3C0097E06EBD5B44D90134492C3BF2">
|
||||
<label id="920DA0E53F8B552A7DF5395BFA19421C">Figure 1.</label>
|
||||
</paragraph>
|
||||
<paragraph id="106342625C7D3CA0C9E1BBA31644CC4C">
|
||||
Photographs of
|
||||
<taxonomicName id="9C77CFF93CB280E015B756F3E945C9B6" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
|
||||
<emphasis id="420C3A988589D2F56145DC71F9302EE7" italics="true">Siccibythus robustus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="4123FBBFE381A12501D9AFEA9F970823" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(Female holotype; CNU-HYM-MA- 2016212)
|
||||
<emphasis id="C4AFB60CC9C0A6DE552E0A9FB0B96A5B" bold="true">A</emphasis>
|
||||
habitus in lateral view
|
||||
<emphasis id="0871FAE2DA1D69A035EAE9252A283574" bold="true">B</emphasis>
|
||||
head in lateral view
|
||||
<emphasis id="548FE8D4AD72623E639C0882EF0C1D93" bold="true">C</emphasis>
|
||||
fore wing venation.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="E3C9E3AFA754ED81DA42B3A4637B606F" ID-DOI="10.3897/jhr.98.136200.figure2" ID-arpha="FFE4DB87-AAE0-5845-8D6E-4EB8D33E1480" httpUri="https://binary.pensoft.net/fig/1213365" startId="F2">
|
||||
<paragraph id="FC584D3F013CDC035299237C7BC1916C">
|
||||
<label id="91CBDDF218914FF5BACEDD78431FCFB0">Figure 2.</label>
|
||||
</paragraph>
|
||||
<paragraph id="E2AEC1027376027C8C12FA408CB9048A">
|
||||
Photographs of
|
||||
<taxonomicName id="F05CE8D80811E21ACA83F0AB44832331" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
|
||||
<emphasis id="A503E1BBD672ED0FC4E7E5E453314E85" italics="true">Siccibythus robustus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="70107FF25A110E5BFE0CEB7BBA84E5CE" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(Female holotype; CNU-HYM-MA- 2016212)
|
||||
<emphasis id="A8D3C9C81F9240040ABA4364953D85D3" bold="true">A</emphasis>
|
||||
mesosoma in dorsal view
|
||||
<emphasis id="76DFD7EC0F82A43A29D1A8EDB94A0379" bold="true">B</emphasis>
|
||||
mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae
|
||||
<emphasis id="E443B9723F46074361DD0BB078374B78" bold="true">C</emphasis>
|
||||
hind leg in lateral view
|
||||
<emphasis id="71CC2DDF41D4969F168CB2D0E3BD16B6" bold="true">D</emphasis>
|
||||
metasoma in lateral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="4C041BB5504A7F966761441C0AD37B0F" ID-DOI="10.3897/jhr.98.136200.figure3" ID-arpha="E6E96A44-67D9-5C8C-8AC1-725919698C9E" httpUri="https://binary.pensoft.net/fig/1213366" startId="F3">
|
||||
<paragraph id="DDC36C3A5C5FE02127F913C635104ECA">
|
||||
<label id="BD1C3E014A66C632D75D27D8CBB2E79A">Figure 3.</label>
|
||||
</paragraph>
|
||||
<paragraph id="9CA13883CDB30320179AFF61F5033F2D">
|
||||
Photographs of
|
||||
<taxonomicName id="15CB95FF3F725A73AA266AF593DFAB5A" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
|
||||
<emphasis id="5E14813E805812CB554E8842A261BE0B" italics="true">Siccibythus robustus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="EB77FF5F954EEAE645A87C38F074FC44" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(Male paratype; CNU-HYM-MA- 2016213)
|
||||
<emphasis id="D4C44BA6AC5B7102965D4636117CD1FB" bold="true">A</emphasis>
|
||||
habitus in lateral view
|
||||
<emphasis id="D0522B955611230B0A57E7500D013E7D" bold="true">B</emphasis>
|
||||
head in lateral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="112A143F703AAA2F521ECC430DEA8218" ID-DOI="10.3897/jhr.98.136200.figure4" ID-arpha="CB887498-BF95-53AC-9F83-9B4FB0C07766" httpUri="https://binary.pensoft.net/fig/1213367" startId="F4">
|
||||
<paragraph id="758DA2637FFEA198A558E80A4AF73038">
|
||||
<label id="B1948BF22B9DA31C456EFC1ECC806500">Figure 4.</label>
|
||||
</paragraph>
|
||||
<paragraph id="77E61C5C564E9080008AB26A1E21FED3">
|
||||
Photographs of
|
||||
<taxonomicName id="555C3492C6AA0D7FA2C965D139AED29D" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
|
||||
<emphasis id="C270CDB3F177F3DEE87A001653547A3D" italics="true">Siccibythus robustus</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="1CD08DF755BAAC10C1A42BF82C72DD66" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(Male paratype; CNU-HYM-MA- 2016213)
|
||||
<emphasis id="8142C8CB273495AB206F37096425949E" bold="true">A</emphasis>
|
||||
mesosoma in dorsal view
|
||||
<emphasis id="AE26FDDF69E9F45F0D0AFC3489F6F5E6" bold="true">B</emphasis>
|
||||
fore wing venation
|
||||
<emphasis id="C649A50D860B8C6972188A1392BB56DA" bold="true">C</emphasis>
|
||||
hind leg in lateral view
|
||||
<emphasis id="102EF47FA8868F0233B4E672B3111591" bold="true">D</emphasis>
|
||||
metasoma in lateral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EBXAE" type="type locality">
|
||||
<paragraph id="87637630DA09704E8407130B16C2360B">
|
||||
<heading id="7250C516F78A5B627BD90BC70107DD15" reason="title">Type locality and horizon.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="E85ADEF8FE91882A4E270D5B37D31C3E">
|
||||
The amber specimens were collected from
|
||||
<collectingRegion id="9273C124D947FEB65FF68DAD75511F18" country="Myanmar" name="Kachin">Kachin</collectingRegion>
|
||||
, northern
|
||||
<collectingCountry id="761A619A613B55AD812456233696EF4B" name="Myanmar">Myanmar</collectingCountry>
|
||||
, lowermost Cenomanian, mid-Cretaceous, which was dated at 98.79 ± 0.62 Ma (
|
||||
<bibRefCitation id="7AB70E8ED6B23D562A687550CCF820CC" DOI="10.1016/S1367-9120(02)00044-5" author="Cruickshank and Ko" firstAuthor="Cruickshank" journalOrPublisher="Journal of Asian Earth Sciences" pagination="441-455" refId="B3" refString="Cruickshank RD, Ko K (2003) Geology of an amber locality in the Hukawng Valley, Northern Myanmar. Journal of Asian Earth Sciences 21: 441–455. https://doi.org/10.1016/S1367-9120 (02) 00044-5" title="Geology of an amber locality in the Hukawng Valley, Northern Myanmar." volume="21" year="2003">Cruickshank and Ko 2003</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="AB5387F18AD060B8D1DB155832A8E821" DOI="10.1016/j.cretres.2012.03.014" author="Shi" etAl="et al." firstAuthor="Shi" journalOrPublisher="Cretaceous Research" pagination="155-163" refId="B25" refString="Shi G, Grimaldi DA, Harlow GE, Wang J, Wang J, Yang M, Lei W, Li Q, Li X (2012) Age constraint on Burmese amber based on U – Pb dating of zircons. Cretaceous Research 37: 155–163. https://doi.org/10.1016/j.cretres.2012.03.014" title="Age constraint on Burmese amber based on U - Pb dating of zircons." volume="37" year="2012">Shi et al. 2012</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EPXAE" type="description">
|
||||
<paragraph id="94D6F86FA75D3A644FEA36DFCC5ECE85">
|
||||
<heading id="FA8765CA507A0ED611BF7849919B01A4" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="FA8251C47E484315660742FAABDB8C01">
|
||||
<emphasis id="40ADF2920AD2D12BF606559A45E41602" bold="true">Female.</emphasis>
|
||||
Head globose, with eyes bulging, occupying most of lateral parts of head (except for wide, convex temples), almost reaching mandibular bases, occipital carina complete, reaching hypostomal carina well behind mandibular base, vertex and frons narrow, longer than wide, ocelli in small acute triangle well distant from eyes, interantennal space flat, clypeus short, simple. Antenna slender, with 12 antennomeres in female (
|
||||
<quantity id="2C41D53EAEF5D2C76FB7FEE026567228" metricMagnitude="-1" metricUnit="m" metricValue="3.302" unit="in" value="13.0">13 in</quantity>
|
||||
male), each antennomere is approximately 1–2 times longer than wide, scape slightly longer than pedicel and first flagellomere combined, pedicel longer than first flagellomere, flagellomeres slightly shortening gradually toward apex except for shorter first flagellomere and longer apical segment (Fig.
|
||||
<figureCitation id="F699C6D80D7DE1866B99A965E3FF1D70" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A habitus in lateral view B head in lateral view C fore wing venation." figureDoi="10.3897/jhr.98.136200.figure1" httpUri="https://binary.pensoft.net/fig/1213364">1 B</figureCitation>
|
||||
). Labrum not visible externally, mandibles small, overlapping, maxillary and labial palps not visible (
|
||||
<typeStatus id="CC6E94F21DCA027726E016FF8F97042A" type="paratype">paratype</typeStatus>
|
||||
can be observed mandibles with three teeth, maxillary palp with four palpomeres visible, three apical long, narrow).
|
||||
</paragraph>
|
||||
<paragraph id="85AD161937E8B8CD793093CD4EB1DC17">
|
||||
Pronotum posterior margin extended, with pronotum accounting for 0.4 times the length of mesosoma, pronotum short declivitous part delimited by transverse groove posteriorly. Propleura long, slender, abutting throughout ventrally. Mesoscutum convex between notauli; notauli complete, parapsides and lateral edges incomplete; notauli widely spaced anteriorly, converging to almost meet posteriorly; transscutal articulation distinct; mesoscutellum triangular, narrow, the length is 0.4 times its maximum width, weakly convex, carinate laterally. Meso- and metapleuron smooth, metapleuron delimited from propodeum with incomplete row of pits (Fig.
|
||||
<figureCitation id="0C3A0FF577D004AA276D4B0F51FA8E49" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2 A, B</figureCitation>
|
||||
). Metanotum short, with indistinct metascutellum in female (metanotum short, with distinct metascutellum in male). Propodeum with clear dorsal and posterior surfaces meeting in angle, dorsal with longitudinal and one transverse carinae, posterior surface with lateral longitudinal carinae, sublateral longitudinal carinae and supracoxal transverse carinae (Fig.
|
||||
<figureCitation id="96DF58117974BD85ED122880D02D6115" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2 B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="4D74DB5987C0A293884A2AC0C80596D8">
|
||||
Fore wing distal of basal veins (1 Rs and 1 M), r-rs and 1 cu-a without tubular veins, except for pterostigma and half section of 2 Rs distal of r-rs, all other distal veins nebulous in female (fore wing distal of basal vein (1 Rs and 1 M) and cu-a with no tubular veins, except for pterostigma, r-rs and most of Rs distal of r-rs, all other veins nebulous in male). Pterostigma prominent, longer than wide; vein 1 Rs straight, shorter than 1 M, arise from Sc +
|
||||
<collectionCode id="9BA04AB4F566B4382800CC66E952BE73" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
|
||||
just proximal to pterostigma; 1 cu-a shorter than 1 M, placed distinctly before it, parallel to 2 cu-a; Cu parallel to A; vein A extending distal to 1 cu-a, and distal part tubular, extending distal to 2 cu-a as nebulous vein, both 1 cu-a and 2 cu-a reaching A; r-rs slightly curved, arising beyond midlength of pterostigma; r-m not present (Fig.
|
||||
<figureCitation id="BF173D951FE3C1C7839D1920E5B60A7C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A habitus in lateral view B head in lateral view C fore wing venation." figureDoi="10.3897/jhr.98.136200.figure1" httpUri="https://binary.pensoft.net/fig/1213364">1 C</figureCitation>
|
||||
). Hind wing with thin, tubular veins Rs arising from Sc +
|
||||
<collectionCode id="D53173D05D11AB263FF2D013F3E30ED3" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
|
||||
; A present; other veins not visible. Seven hamuli present.
|
||||
</paragraph>
|
||||
<paragraph id="355B566CBAD0CE561DED1F0FFC8EDEFC">
|
||||
Fore coxa swollen, triangular, nearly as long as wide; fore trochanter short, fore trochantellus not visible in female (fore trochantellus trapezoidal, the ratio compared to the length of the fore femur is approximately 1:
|
||||
<quantity id="0CECB28AB6386D2B07078DD99A5DB1DE" metricMagnitude="-1" metricUnit="m" metricValue="4.318" unit="in" value="17.0">17 in</quantity>
|
||||
male); fore femur swollen in middle, twice as long as wide (calcar long, thin, gently curved and acute apically in male); fore tibia nearly as long as femur; fore tarsus shorter than fore tibia, basitarsomere shorter than combined length of remaining tarsomeres in female (basitarsomere longer than combined length of remaining tarsomeres in male). Mid coxa slender, smaller than both fore and hind coxa, twice as long as wide; mid trochanter short, mid trochantellus very short and ring-like, approximately 0.1 times as long as femur; mid femur swollen in middle, twice as long as wide; mid tibia slenderer than femur, three times as long as wide, tibia shorter than femur; mid tarsus nearly as long as mid tibia, basitarsomere shorter than combined length of remaining tarsomeres. Hind coxae elongate, swollen, three times as long as wide, with lamella dorsally, basally in female (no lamella in male); hind trochanter short, bulging dorsally, hind trochantellus very short and ring-like (ca. 0.02 as long as femur); hind femur swollen in middle, nearly three times as long as wide in female (nearly six times as long as wide in male); hind tibia very slender, slightly longer than femur; hind tarsus nearly as long as hind tibia, basitarsomere slightly shorter than combined length of remaining tarsomeres (Fig.
|
||||
<figureCitation id="71158E2214C902A03F2178B40782DDD7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2 C</figureCitation>
|
||||
). Claw simple, arolium small. Tibial spur formula 1 / 2 / 2.
|
||||
</paragraph>
|
||||
<paragraph id="56B0BBCE11F19F376D53E8523A694C09">
|
||||
First metasomal segment short, tergum node-like, female with anterior outline, low triangular, second metasomal segment longest and highest, following segments gradually smaller, apical metasomal sternum high, with dorsal sides overlapping (Fig.
|
||||
<figureCitation id="FB3C2D48856086166F6CBBB1F47BF0D6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2 D</figureCitation>
|
||||
) (male first metasomal tergum with fore surface convex, dorsal surface short, rather flat, elevated above hind surface; seventh metasomal sternum with two long upcurved teeth).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EXYAE" type="description">
|
||||
<paragraph id="8EE4CBF29EBF7AFA3D20105DFB7F399A">
|
||||
<heading id="22C51808E09F262FE0D6F038695042B0" reason="title">Measurements (mm).</heading>
|
||||
</paragraph>
|
||||
<paragraph id="6E51014A65FF4706A344F062FE044C61">
|
||||
<typeStatus id="E0B2232FA29A58944CC703C39E3EE201" type="holotype">Holotype</typeStatus>
|
||||
female. Total body length (excluding sting and antenna) 3.93; head length 0.47, width 0.61, height 0.53; scape length 0.25, pedicel length 0.13, the first flagellum length 0.08, total flagellum length 0.87; fore wing length 2.22; hind wing length 1.56; fore coxa length 0.30, fore trochanter length 0.57, fore femur length 0.58, fore tibia length 0.55, fore tarsal length 0.32; mid coxa length 0.31, mid trochanter length 0.09, mid trochantellus 0.05, mid femur length 0.50, mid tibia length 0.38, mid tarsal length 0.38; hind coxa length 0.44, hind trochanter length 0.14, hind trochantellus 0.02, hind femur length 0.67, hind tibia length 0.78, hind tarsal length 1.02; the first metasomal segment length 0.28, metasoma length 2.18.
|
||||
</paragraph>
|
||||
<paragraph id="174A11C3E7381316AB511D9DB0E4F8C8">
|
||||
<typeStatus id="CC6EA19B5BB394F69313BF79D8D4DF11" type="paratype">Paratype</typeStatus>
|
||||
male. Total body length (excluding antenna) 3.54; head length 0.50, width 0.42, height 0.50; scape length 0.25, pedicel length 0.16, the first flagellum length 0.09; total flagellum length 0.99; fore wing length 1.98; hind wing length 1.47; fore coxa length 0.21, fore trochanter length 0.08, fore trochantellus 0.03, fore femur length 0.50, fore tibia length 0.45, fore tarsal length 0.32; mid coxa length 0.12, mid trochanter length 0.10, fore trochantellus 0.02, mid femur length 0.40, mid tibia length 0.30, mid tarsal length 0.34; hind coxa length 0.28, hind trochanter length 0.12, hind femur length 0.55, hind tibia length 0.60, hind tarsal length 0.57; the first metasomal segment length 0.25, metasoma length 1.62.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E4YAE" type="remarks">
|
||||
<paragraph id="FB1D82DF19A11C69D6E8DA02344D0233">
|
||||
<heading id="D63F8971A268B49E5C26BA4D31D8AA11" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="578CBC58B01488B7356B8689B8AC3322">
|
||||
The new species is classified in †
|
||||
<taxonomicName id="599B0064B45DACD5A3A150E87FC7F15E" authority="Cockx & McKellar, 2016" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="8F6EE06B86B82173886BC8DBC82FA071" italics="true">Siccibythus</emphasis>
|
||||
<bibRefCitation id="DE416655AA41DB0EBA1F33A52EBD27BE" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD & McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx & McKellar, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
based on the fore wing with no cells closed with tubular veins in its middle and distal part (
|
||||
<bibRefCitation id="78E05863837D24B5350572C242D1F2B3" DOI="10.11646/palaeoentomology.3.6.10" author="Rasnitsyn" etAl="et al." firstAuthor="Rasnitsyn" journalOrPublisher="Palaeoentomology" pagination="582-596" refId="B20" refString="Rasnitsyn AP, Zhang Q, Müller P, Zhang HC (2020) On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.). Palaeoentomology 3: 582–596. https://doi.org/10.11646/palaeoentomology.3.6.10" title="On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.)." volume="3" year="2020">Rasnitsyn et al. 2020</bibRefCitation>
|
||||
). However, the new species is differentiated from
|
||||
<taxonomicName id="58BDB0F64D7243F492774E3A309686DB" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="martynovae">
|
||||
<emphasis id="99BD5F86901C6FDEF1AF17F1D38DF0B8" italics="true">Siccibythus martynovae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="0C211A049947632DF234F307CE7D43E8" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pallidus">
|
||||
<emphasis id="F4968C9B505511E04429EA0A4A0237AD" italics="true">S. pallidus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="917667F797B549320B2D7288192A08CB" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="oculatus">
|
||||
<emphasis id="93C746DA90AF6D27A2761044CD63056E" italics="true">S. oculatus</emphasis>
|
||||
</taxonomicName>
|
||||
by fore wing with 2 Rs distal of r-rs several times as long as r-rs, basitarsus longer than tarsomeres 2–4 combined (vs. 2 Rs distal of r-rs at most about as long as r-rs, basitarsus not longer than tarsomeres 2–4 combined in
|
||||
<taxonomicName id="09A6C6D38A30EC858B2F37772D232175" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="martynovae">
|
||||
<emphasis id="1EFDED134CCEC00E6ED3622DC5D74D10" italics="true">S. martynovae</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="7BC8EEABC15FF4462D8D3A3C886CB067" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pallidus">
|
||||
<emphasis id="15810D245CC3E615B9402B4900433F25" italics="true">S. pallidus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4CAE7DD5CB3685815926698B0C74BC42" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="oculatus">
|
||||
<emphasis id="768DF06C37D712CA66A445653A52DE8A" italics="true">S. oculatus</emphasis>
|
||||
</taxonomicName>
|
||||
). It is distinguished from
|
||||
<taxonomicName id="FF4B8F78C2C47BE809658F28B1330821" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="musculosus">
|
||||
<emphasis id="AD001712E618813C823F90B3DFDE584C" italics="true">S. musculosus</emphasis>
|
||||
</taxonomicName>
|
||||
by the following characters: 1) Cu is nebulous (vs. Cu is tubular); 2) calcar long, thin and acute apical (vs. calcar short, bifid apically). It is differentiated from
|
||||
<taxonomicName id="C1F14698F4E18B5199879FBE5C41E758" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="paulus">
|
||||
<emphasis id="F46570DC92EB65744A6F5D4A3AC765BE" italics="true">S. paulus</emphasis>
|
||||
</taxonomicName>
|
||||
by the length of the scape being less than 0.3 times the length of the flagellum (vs. the scape is longer than or equal to 0.3 times of the flagellum). It is distinguished from
|
||||
<taxonomicName id="71F0186EAEFBF2C32BFCF0DA9DCE266F" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ohmkuhnlei">
|
||||
<emphasis id="01E175FA52540E6714C62DDB1FB830B6" italics="true">S. ohmkuhnlei</emphasis>
|
||||
</taxonomicName>
|
||||
by the following characters: 1) anterior outline of 1
|
||||
<superScript id="A83D8D7CF8F465D48004A31FEBFDEAD1">st</superScript>
|
||||
metasomal tergum somewhat angular (vs. straight); 2) propodeum outline angular (vs. more smoothly curved).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
275
data/7B/67/07/7B670705B95353228DE96C96DD49E7A3.xml
Normal file
275
data/7B/67/07/7B670705B95353228DE96C96DD49E7A3.xml
Normal file
|
|
@ -0,0 +1,275 @@
|
|||
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<mods:mods id="B245CD56578B8445C65DA466E27FA6B0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BEA99FDC1CDA9E916F05035051372DBC">
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<mods:title id="929660F95F0F561053B7585EA22C32E1">New wasps of † Falsiformicidae from mid-Cretaceous Kachin amber</mods:title>
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<mods:namePart id="3BCB2FD336519CA391B0184007258833">Wang, Zhen</mods:namePart>
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<mods:affiliation id="A5F58B64F351A7F8A1BDC9165BDEA443">College of Life Sciences, Capital Normal University, Beijing, 100048, China</mods:affiliation>
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<mods:namePart id="24A9356571DC211B5AC93F62B67CC76C">Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation id="BDC825CF9A94F1FFB1D5B542418430C5">A. A. Borissiak Palaeontological Institute, Russian Academy of Sciences, Moscow, 117647, Russia</mods:affiliation>
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<mods:namePart id="4497B136D97A322DF9EC82D30C555349">Vilhelmsen, Lars</mods:namePart>
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<mods:nameIdentifier id="7D4F34E97180BF4B8391DF29238E6D95" type="ORCID">0000-0002-5593-5722</mods:nameIdentifier>
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<mods:namePart id="1F0685A6B4FC7AC5A49309A9A77D9824">Gao, Taiping</mods:namePart>
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<mods:part id="D29B141072BDAB34C779B4E19B359157">
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<mods:number id="08C9A1BC94F460D886F2610AC4629CDD">2025-01-03</mods:number>
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<mods:identifier id="6BCD62E6746A807591F855E1DE4C7613" type="ZooBank">77736D1C-30F9-4D6C-8310-3A04121CACE2</mods:identifier>
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</mods:mods>
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<treatment id="7B670705B95353228DE96C96DD49E7A3" ID-arpha="7B670705-B953-5322-8DE9-6C96DD49E7A3" ID-zoobank="https://zoobank.org/C0027405-25CC-48BB-AD21-8FCA112E21F0" LSID="urn:lsid:plazi:treatment:7B670705B95353228DE96C96DD49E7A3" LSID-ZBK="urn:lsid:zoobank.org:act:C0027405-25CC-48BB-AD21-8FCA112E21F0" httpUri="http://treatment.plazi.org/id/7B670705B95353228DE96C96DD49E7A3">
|
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<subSubSection id="CDBE9E3391390AFAEB15843DDDA73A92" type="nomenclature">
|
||||
<paragraph id="FA1A7CFAAB8F0F7F3BEF763076FB84E8">
|
||||
<taxonomicName id="1050EDADCE5205A07A0D36E3E1020B30" authority="Wang, Rasnitsyn, Perkovsky & Vilhelmsen" authorityName="Wang, Rasnitsyn, Perkovsky & Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
|
||||
<emphasis id="FA1AEE3227921FA2492CAE606B70C362" italics="true">Siccibythus aristovi</emphasis>
|
||||
Wang, Rasnitsyn, Perkovsky & Vilhelmsen
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="58F2B1114EA1684BCE7C89F5004C0DD9" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E34AE" type="diagnosis">
|
||||
<paragraph id="C0CE1C096649597B1A870FB43EA8252B">
|
||||
<heading id="A8CD3B84ABA4553E1BBFEEB9F0CC4FB3" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="07FAA2C035100AA8E51F3ED300979AEF">Fore wing with 2 Rs distal of r-rs at most about as long as r-rs; Rs + M and Cu both nebulous; A behind 1 cu-a not pigmented (except for very base). Fore tibial spur short, bifid apically. Basitarsus not longer than tarsomeres 2–4 combined. Pronotum short, propodeal horns long.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EB5AE" type="etymology">
|
||||
<paragraph id="3A2C5B908FB64911ADD8E1A90900257F">
|
||||
<heading id="B6F5FCD956CF3CCA74B523D4ADA1C60C" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="24B111D3FFC247AF0FA0F18D37312802">
|
||||
The specific name “
|
||||
<emphasis id="CD8525821D6801B228DE757893C3EA9B" italics="true">aristovi</emphasis>
|
||||
” is dedicated to the memory of the late Dr. Daniil Sergeevich Aristov, for his outstanding contribution to palaeoentomology.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ER5AE" type="type material">
|
||||
<paragraph id="6A67FDA9ABDC420344A33B47FB9800B7">
|
||||
<heading id="6EAFD42892662108502502D88C7BD766" reason="title">Type material.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="F7F3799DEA90C310DA25549ECC513411">
|
||||
<emphasis id="23CBBD796B71D8420A68F1AA80BD494F" bold="true" italics="true">
|
||||
<typeStatus id="ADD3EE3BA589341585273690D66BC93E" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
• female. No. CNU-HYM-MA 2016214. A well-preserved and complete specimen (Figs
|
||||
<figureCitation id="B1B2CF1FAB8FB3913B45C15FD2CDF6EC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A habitus in dorsal view B habitus in lateral view." figureDoi="10.3897/jhr.98.136200.figure5" httpUri="https://binary.pensoft.net/fig/1213368">5</figureCitation>
|
||||
,
|
||||
<figureCitation id="015120464B11CE1A2BE5A1D0FEAC3D51" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="FA85A54E913004BCE28E6258F95D15AE" ID-DOI="10.3897/jhr.98.136200.figure5" ID-arpha="23B8DC68-F7F1-5D46-94A4-B92BC5D5D76B" httpUri="https://binary.pensoft.net/fig/1213368" startId="F5">
|
||||
<paragraph id="2094A10D15D8ABA20594E7C63D89A5D9">
|
||||
<label id="2F9FD6E1D1F2039D3A1FBFF37EF08A15">Figure 5.</label>
|
||||
</paragraph>
|
||||
<paragraph id="72D34D4CCAF54A770BCA315645201E10">
|
||||
Photographs of
|
||||
<taxonomicName id="2D940FFD9D88738018770A13BFB808CD" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
|
||||
<emphasis id="256075F34C8B158D70D2F57AD857D652" italics="true">Siccibythus aristovi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="8B9CF498DBF7B61E1AE11B0A380D2920" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(Female holotype; CNU-HYM-MA- 2016214)
|
||||
<emphasis id="84C90237F123EA4100EBC9CD28A6EB2C" bold="true">A</emphasis>
|
||||
habitus in dorsal view
|
||||
<emphasis id="E1F4BB78E1D7ECFC0484AB3B746EF4D1" bold="true">B</emphasis>
|
||||
habitus in lateral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="7438A2D4CF0C2481BC257CE76249A2CF" ID-DOI="10.3897/jhr.98.136200.figure6" ID-arpha="558F7053-A129-5650-99D8-8479B810E09A" httpUri="https://binary.pensoft.net/fig/1213369" startId="F6">
|
||||
<paragraph id="858FBD0B7BFFA6769A4E7AF06EE056B7">
|
||||
<label id="F8F22A0BC4DBB9E0D2FAC7DBDD2B6E9D">Figure 6.</label>
|
||||
</paragraph>
|
||||
<paragraph id="157B5187014E0A7695EAFB15800C85B6">
|
||||
Photographs of
|
||||
<taxonomicName id="AD2791CC4F375E78481B7CDB93F715B4" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
|
||||
<emphasis id="35413E489406873A430DCFDEF64EB1D6" italics="true">Siccibythus aristovi</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="E1EAC4F372FB775E203426194203FAA7" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(Female holotype; CNU-HYM-MA- 2016214)
|
||||
<emphasis id="D0C9DCB20006CC21AE41A34283AEE8E1" bold="true">A</emphasis>
|
||||
head in lateral view
|
||||
<emphasis id="2683DD1F1AC5B93FD9BD5FC7BD4EF748" bold="true">B</emphasis>
|
||||
mouthparts
|
||||
<emphasis id="2B3621D333F587F0E2F190A6866B2016" bold="true">C</emphasis>
|
||||
mesosoma in lateral view, black arrow indicates the carinae
|
||||
<emphasis id="C5171604F303287BA580EE75F5429487" bold="true">D</emphasis>
|
||||
mesosoma in dorsal view
|
||||
<emphasis id="7B449FD0ECCDC48167DC4A64B87BBE6F" bold="true">E</emphasis>
|
||||
fore wing venation
|
||||
<emphasis id="0554A827528FCBACEB09CAB27B80E1E5" bold="true">F</emphasis>
|
||||
metasoma in lateral view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EFBAG" type="type locality">
|
||||
<paragraph id="5CF1C2D447366ACFDD2B0261A75E2A07">
|
||||
<heading id="667E5F991882758B4B64B9EAA091B76B" reason="title">Type locality and horizon.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="29B3A6E8E933494C0DB95B8640C87608">
|
||||
The amber specimen was collected from
|
||||
<collectingRegion id="00F0564B3DB959A9AACD7326A20DC0E2" country="Myanmar" name="Kachin">Kachin</collectingRegion>
|
||||
, northern
|
||||
<collectingCountry id="9F463E19AC482388677765E681184B11" name="Myanmar">Myanmar</collectingCountry>
|
||||
, lowermost Cenomanian, mid-Cretaceous, which was dated at 98.79 ± 0.62 Ma (
|
||||
<bibRefCitation id="33BE68BBF3224DF39C4D04C2306B41F4" DOI="10.1016/S1367-9120(02)00044-5" author="Cruickshank and Ko" firstAuthor="Cruickshank" journalOrPublisher="Journal of Asian Earth Sciences" pagination="441-455" refId="B3" refString="Cruickshank RD, Ko K (2003) Geology of an amber locality in the Hukawng Valley, Northern Myanmar. Journal of Asian Earth Sciences 21: 441–455. https://doi.org/10.1016/S1367-9120 (02) 00044-5" title="Geology of an amber locality in the Hukawng Valley, Northern Myanmar." volume="21" year="2003">Cruickshank and Ko 2003</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="6C482AF1AC389557F1209BE41BE8DAA4" DOI="10.1016/j.cretres.2012.03.014" author="Shi" etAl="et al." firstAuthor="Shi" journalOrPublisher="Cretaceous Research" pagination="155-163" refId="B25" refString="Shi G, Grimaldi DA, Harlow GE, Wang J, Wang J, Yang M, Lei W, Li Q, Li X (2012) Age constraint on Burmese amber based on U – Pb dating of zircons. Cretaceous Research 37: 155–163. https://doi.org/10.1016/j.cretres.2012.03.014" title="Age constraint on Burmese amber based on U - Pb dating of zircons." volume="37" year="2012">Shi et al. 2012</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ETBAG" type="description">
|
||||
<paragraph id="19A98DEC328B2D4D63C61D80D237FDD6">
|
||||
<heading id="1138E481BFF6647D54AC112F54D03B65" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="02724523D63E8060355BE48887187CEA">
|
||||
<emphasis id="BCDD5E2927D50971546CCC56744F35AA" bold="true">Female.</emphasis>
|
||||
Body brown, metasoma dark brown. Head globose, with eyes bulging, occupying three-quarters of head in lateral view, almost reaching mandibular bases, occipital carina complete, reaching hypostomal carina well behind mandibular base, ocelli in small acute triangle well distant from eyes, interantennal space flat. Antenna narrow, with 12 antennomeres, each antennomere is approximately 1–2 times longer than wide, scape shorter than pedicel, pedicel longer than first flagellomere, flagellomeres slightly shortening gradually toward apex except for longer apical segment (Fig.
|
||||
<figureCitation id="7902E25D3EB7E86E972CCA483AC4E087" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 A</figureCitation>
|
||||
). Mandibles with three teeth, not overlapping; labrum not visible; maxillary palp short with six elongate palpomeres, labial palp short, with four slightly elongate to subquadrate palpomeres (Fig.
|
||||
<figureCitation id="E6A459043219E2FE3E993BD1F8BC2374" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="6D3793C7486B5C103E4440B072142D8E">
|
||||
Pronotum with long dorsal part shorter than wide before lateral lobes, constricted near midlength, with fore declivitous part short, delimited by transverse groove posteriorly. Propleura long, slender, in lateral view, the length is 4 times its width, and abutting throughout. Mesoscutum with deep notauli widely spaced anteriorly, converging to almost meet posteriorly; transscutal articulation distinct; mesoscutellum rather short, wide apically, weakly convex. Meso- and metapleurae smooth. Metanotum short, with metascutellum very short and wide. Propodeum with dorsal and posterior surfaces meeting in angle, dorsal with longitudinal carina, and with transverse one midway between sublateral carinae, posterior surface with lateral longitudinal carinae, sublateral longitudinal carinae and supracoxal transverse carinae (Fig.
|
||||
<figureCitation id="FE54A8E8338C4695A4E97926966F5579" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 C, D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="165E37707DCD62287E130078425F844D">
|
||||
Fore wing basal veins (C, Sc +
|
||||
<collectionCode id="7AE4C02EEB7DAB14F1EEB53817E740C7" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
|
||||
, M + Cu, A, 1 Rs and 1 M), r-rs, short pterostigma, part of 2 Rs distal of r-rs and cu-a tubular, other veins nebulous. Pterostigma prominent, longer than wide; vein 1 Rs straight, shorter than 1 M, arise from Sc +
|
||||
<collectionCode id="D18C1B5552760985368B8DFDB14F42C5" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
|
||||
just proximal to pterostigma; 1 cu-a shorter than 1 M, parallel to 2 cu-a, placed slightly basally of 1 M; 1 Cu parallel to A; vein A extends distal up to 1 cu-a only; r-rs straight, arising beyond midlength of pterostigma; r-m not present (Fig.
|
||||
<figureCitation id="53770044FADC31CCA21C326E9B60C2EC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 E</figureCitation>
|
||||
). Hind wing with Sc +
|
||||
<collectionCode id="81B9D5B208BEF5512D04C282C1613A3E" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
|
||||
; other veins not visible.
|
||||
</paragraph>
|
||||
<paragraph id="320039E7456960E2D4F61DC461BFE47A">Fore trochanter present, fore trochantellus ring-like; fore femur slightly swollen in middle, nearly three times as long as wide; fore tibia shorter than femur, calcar acute (not bifid); fore tarsus shorter than fore tibia, basitarsomere shorter than combined length of remaining tarsomeres. Mid trochanter present, mid trochantellus ring-like; mid femur slightly swollen in middle, four times as long as wide; mid tibia nearly as long as femur, six times as long as wide; mid tarsus shorter than mid tibia, basitarsomere shorter than combined length remaining tarsomeres. Hind trochanter short, bulging dorsally, hind trochantellus ring-like; hind femur slightly swollen in middle, nearly four times as long as wide; hind tibia slender than femur, longer than femur; hind tarsus nearly as long as hind tibia, remaining tarsomeres combined shorter than basitarsomere. Claw simple, arolium small. Tibial spur formula 1 / 2 / 2.</paragraph>
|
||||
<paragraph id="06A1C006A8F41699B95069D266C10698">
|
||||
First metasomal segment short, tergum node-like, triangular in lateral view, with anterior surface concave, second segment slightly longer and higher than following, following gradually smaller, apical (sixth) sternum high, apparently folded, with upper sides and apical orifice apparently not specialized, tergum 7 as visible (in lateral view) small but more or less sclerotized, with spiracle not enlarged (Fig.
|
||||
<figureCitation id="51A5402712162F471B21C9208AA9D599" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 F</figureCitation>
|
||||
). Sting straight, apparently short, without teeth.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EWCAG" type="description">
|
||||
<paragraph id="0196B03107F33418C9477AEB5EB3B9D9">
|
||||
<heading id="142FD024AC6C4A251910F2AA8964D328" reason="title">Measurements (mm).</heading>
|
||||
</paragraph>
|
||||
<paragraph id="AF2F1D01F00804FDB31C7EFAFDD1B2A1">
|
||||
<typeStatus id="60534AEFF89EB0922C76B9B52C35ABCF" type="holotype">Holotype</typeStatus>
|
||||
female. Total body length (excluding sting and antenna) 4.14; head length 0.94, width 0.62, height 0.48; scape length 0.20, pedicel length 0.19, the first flagellum length 0.14, total flagellum length 1.11; fore wing length 1.95; hind wing length 1.20; fore coxa length 0.27, fore trochantellus 0.06, fore femur length 0.64, fore tibia length 0.49, fore tarsal length 0.64; mid coxa length 0.25, mid trochanter length 0.10, mid trochantellus 0.04, mid femur length 0.60, mid tibia length 0.64, mid tarsal length 0.39; hind coxa length 0.23, hind trochanter length 0.10, hind trochantellus 0.06, hind femur length 0.69, hind tibia length 0.58, hind tarsal length 0.79; the first metasomal segment length 0.29, metasoma length 1.66.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E2CAG" type="remarks">
|
||||
<paragraph id="1E9A11545C8FB4751359AE9A4EC2CACC">
|
||||
<heading id="6D8784AFA07B709724D4369DF475B554" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="92EFCD66B272E4B2BBE5DF2DC33034E2">
|
||||
The new species is classified in †
|
||||
<taxonomicName id="16512B109E6E596E1A35960B6ACC879B" authority="Cockx & McKellar, 2016" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="48716356E4F83DF8B0D83D6AE9996A0D" italics="true">Siccibythus</emphasis>
|
||||
<bibRefCitation id="AA5EF34503A8AF3912AD19A31CC9BBC3" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD & McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx & McKellar, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
based on the fore wing with no cells closed with tubular veins in its middle and distal part (
|
||||
<bibRefCitation id="EE75EF1F9B60707724DA9EF51B127BC3" DOI="10.11646/palaeoentomology.3.6.10" author="Rasnitsyn" etAl="et al." firstAuthor="Rasnitsyn" journalOrPublisher="Palaeoentomology" pagination="582-596" refId="B20" refString="Rasnitsyn AP, Zhang Q, Müller P, Zhang HC (2020) On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.). Palaeoentomology 3: 582–596. https://doi.org/10.11646/palaeoentomology.3.6.10" title="On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.)." volume="3" year="2020">Rasnitsyn et al. 2020</bibRefCitation>
|
||||
). It is differentiated from
|
||||
<taxonomicName id="4798421EFC510891D4BD29B9C78AB2D4" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="musculosus">
|
||||
<emphasis id="7A2554C75ABE2D5BB364513CED4F6709" italics="true">Siccibythus musculosus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="5AAB368797027804C4E381A761DD908D" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="paulus">
|
||||
<emphasis id="6AEBB685155A2105F4974B9D84E43374" italics="true">S. paulus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="8364D6CC3400DA567C018B610706C77A" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ohmkuhnlei">
|
||||
<emphasis id="2DB0BDC3C322A1C64FBA888AF917BFD9" italics="true">S. ohmkuhnlei</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="2674496A124A6E51EAEF368ABC9BB4EF" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus">
|
||||
<emphasis id="EF21A7EF736265A1ED9694D1757217CF" italics="true">S. robustus</emphasis>
|
||||
</taxonomicName>
|
||||
by fore wing with 2 Rs distal of r-rs shorter than r-rs, basitarsus not longer than tarsomeres 2–4 combined (vs. 2 Rs distal of r-rs several times as long as r-rs, basitarsus longer than tarsomeres 2–4 combined in
|
||||
<taxonomicName id="65C1CC51DAED0057F54CF782C2E810EA" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="musculosus">
|
||||
<emphasis id="7837D722073DB4384EAB954D1FC84962" italics="true">S. musculosus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="59C5BAFE0F1C29A968A0CE3440BA5E4D" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="paulus">
|
||||
<emphasis id="7435FCD0AF5512462A0E3ECC85CB2396" italics="true">S. paulus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="589A379A0616AC48DF96FA8817481D35" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ohmkuhnlei">
|
||||
<emphasis id="7C83623B986214A6E899608E9F7721C0" italics="true">S. ohmkuhnlei</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="B93DEA4D1374D2C73BC09AF0440DC1BB" authorityName="Wang & Rasnitsyn & Perkovsky & Vilhelmsen & Gao" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus">
|
||||
<emphasis id="D5EC16E9A650603FE6A9E58218D2001A" italics="true">S. robustus</emphasis>
|
||||
</taxonomicName>
|
||||
). It is distinguished from
|
||||
<taxonomicName id="676BF01993866DCD134526AA0906A9AB" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="martynovae">
|
||||
<emphasis id="1628283EE7DE6D9B6415594625812700" italics="true">S. martynovae</emphasis>
|
||||
</taxonomicName>
|
||||
by fore wing veins Rs + M and Cu both being nebulous (vs. Rs + M and Cu are both tubular in
|
||||
<taxonomicName id="63EBCCE5986DFDFB3C426BE37D504C7C" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="martynovae">
|
||||
<emphasis id="C38EF60D9B5F86D7203068C5C5391B10" italics="true">S. martynovae</emphasis>
|
||||
</taxonomicName>
|
||||
). It is differentiated from
|
||||
<taxonomicName id="2BA0E64431454C6FD5CD930B126BDC5D" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pallidus">
|
||||
<emphasis id="F0937FD25FCB9F255C0C0A9A53CC5F57" italics="true">S. pallidus</emphasis>
|
||||
</taxonomicName>
|
||||
by the following characters: 1) fore wing with 2 A not pigmented (except for very base) (vs. fore wing with 2 A pigmented); 2) pronotum constricted in the middle (vs. not constricted). It is distinguished from
|
||||
<taxonomicName id="F883EF78E486B0F14ED8BCB03925DBA5" authorityName="Rasnitsyn, Zhang, Muller & Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="oculatus">
|
||||
<emphasis id="F552A4A1C3F59475FC5CA718388AF49C" italics="true">S. oculatus</emphasis>
|
||||
</taxonomicName>
|
||||
by pronotum short, propodeal horns long (vs. pronotum long, propodeal horns short).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
119
data/EC/53/11/EC531111868A5194B7C353CA91169FF3.xml
Normal file
119
data/EC/53/11/EC531111868A5194B7C353CA91169FF3.xml
Normal file
|
|
@ -0,0 +1,119 @@
|
|||
<document id="EC81AF8A0A8C4C6C4016BCA08D61E0EE" ID-DOI="10.3897/jhr.98.136200" ID-ZooBank="77736D1C-30F9-4D6C-8310-3A04121CACE2" ID-publisher-id="136200" URI-arpha="CFF2E897-2633-5F4E-8369-35D589D56708" URI-zoobank="http://zoobank.org/77736D1C-30F9-4D6C-8310-3A04121CACE2" article-type="research-article" checkinTime="1735917671054" checkinUser="pensoft" docAuthor="Wang, Zhen, Rasnitsyn, Alexandr P., Perkovsky, Evgeny E., Vilhelmsen, Lars & Gao, Taiping" docDate="2025" docId="EC531111868A5194B7C353CA91169FF3" docLanguage="en" docName="Journal of Hymenoptera Research 98: 1-18" docOrigin="Journal of Hymenoptera Research 98" docSource="https://jhr.pensoft.net/article/136200/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Falsiformicidae Rasnitsyn 1975" docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="18" masterDocId="CFF2E89726335F4E836935D589D56708" masterDocTitle="New wasps of † Falsiformicidae from mid-Cretaceous Kachin amber" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1735917735267" updateUser="pensoft">
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||||
<mods:mods id="BDC2A8C703B25E5B2C437F8C7657FE3E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="94E13EB9BD8B577CB5A14FB454B2A68C">
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<mods:title id="2FEF2419BC4BD96898E7B0A683789991">New wasps of † Falsiformicidae from mid-Cretaceous Kachin amber</mods:title>
|
||||
</mods:titleInfo>
|
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<mods:name id="6A66089BFFACF104DBA57EBA1C831FBE" type="personal">
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<mods:role id="BB62959521A374F076167EA27E8FE60A">
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<mods:roleTerm id="2C77EFE8ED6F73BC8E646647EBC7D532">Author</mods:roleTerm>
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||||
</mods:role>
|
||||
<mods:namePart id="0A4438126204C49D55F61C3EC43D03E2">Wang, Zhen</mods:namePart>
|
||||
<mods:nameIdentifier id="85F6DC4EA40A164C3F1F9A2C698D5956" type="ORCID">https://orcid.org/0009-0002-1981-8323</mods:nameIdentifier>
|
||||
<mods:affiliation id="7ADDA2644AC9BF63ED31BD86B72A312A">College of Life Sciences, Capital Normal University, Beijing, 100048, China</mods:affiliation>
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||||
</mods:name>
|
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<mods:name id="85E21B4CDAF9CA253B72BDF9053EEC12" type="personal">
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<mods:role id="9133C619ED22E7D4290135C747F0CFAB">
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<mods:roleTerm id="6E1CFF0F6A88FE79D66C52815C1738C3">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="6A265CCF38672FB2D0392BCAA2218304">Rasnitsyn, Alexandr P.</mods:namePart>
|
||||
<mods:affiliation id="B81E5682F981C57421DB668151835AD1">A. A. Borissiak Palaeontological Institute, Russian Academy of Sciences, Moscow, 117647, Russia</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="B0241D83634F67438D5C69920E47FFDB" type="personal">
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<mods:role id="DCAD6F41E98BB0DFE4DB32803AE4282C">
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<mods:roleTerm id="5B85AA7882AA9BC051F56363F403A117">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="6E8FBA16D338392C83B9C0E2D32687AD">Perkovsky, Evgeny E.</mods:namePart>
|
||||
<mods:nameIdentifier id="42B779BCA77A23DD8E805558FC7DC414" type="ORCID">0000-0002-7959-4379</mods:nameIdentifier>
|
||||
<mods:affiliation id="DEA66E989E113FC39D512ABD31BF2739">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Copenhagen, 2100, Denmark</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="4ADF33A83760F4BD85C30B649CC74B12" type="personal">
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<mods:role id="034F0E5A34E99A0180B9C0FBAB6ECC2F">
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<mods:roleTerm id="72FBC0B64F2541F81FFDF4569611C8C9">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="A234402ECFD8CD2D0A1EB424F85FFA69">Vilhelmsen, Lars</mods:namePart>
|
||||
<mods:nameIdentifier id="2EF2C3E92A6DA3B37BDD9C5C188E5D27" type="ORCID">0000-0002-5593-5722</mods:nameIdentifier>
|
||||
<mods:affiliation id="647DE7F39A767365968F30B7C7A6C79C">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Copenhagen, 2100, Denmark</mods:affiliation>
|
||||
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|
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<mods:name id="C42640268D3B5A9D86B6888BF3B94B22" type="personal">
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<mods:roleTerm id="4C689C63CF71EF05D2E8C0631B0B4DB1">Author</mods:roleTerm>
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||||
</mods:role>
|
||||
<mods:namePart id="8A4702DC1D4E7D53198B2B8178CD1B0D">Gao, Taiping</mods:namePart>
|
||||
<mods:nameIdentifier id="B5110263B4328B30A7AF8F06CE1D0CD0" type="ORCID">0000-0002-8118-8708</mods:nameIdentifier>
|
||||
<mods:affiliation id="1C1DC726247CEB8150EA6C0DABDA86CB">College of Life Sciences, Capital Normal University, Beijing, 100048, China</mods:affiliation>
|
||||
</mods:name>
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<mods:typeOfResource id="B387E247365FA1DC0AE65CFA2FDB0B63">text</mods:typeOfResource>
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<mods:titleInfo id="70564C3FFC9183BD50D11763F1F95ED9">
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<mods:title id="C471A4BE1E6A319BBC2800661CFE52C7">Journal of Hymenoptera Research</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part id="D123C9DC123E2BBC878044440A3C13CC">
|
||||
<mods:date id="A8D9A5086D33A84B4A78B1E49AF80754">2025</mods:date>
|
||||
<mods:detail id="153C4B84975852EA784549175F0B561E" type="pubDate">
|
||||
<mods:number id="A4344E0347E9CAE0E3F89520112F3BCD">2025-01-03</mods:number>
|
||||
</mods:detail>
|
||||
<mods:detail id="310AF30D6158E985AA04210D26D89AF2" type="volume">
|
||||
<mods:number id="5E798D86C4A7473473410EC4ACCBE30D">98</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="2B0D4F0E54C50D52430C6692F16E4570" unit="page">
|
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<mods:start id="116C305B4D82BF08C61A8670D21A6FBE">1</mods:start>
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</mods:extent>
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</mods:part>
|
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|
||||
<mods:classification id="0EB67CC62E617C145A0F6331E84C979A">journal article</mods:classification>
|
||||
<mods:identifier id="7DE394C46D1FE83AD9C2800E09A7BCD2" type="DOI">10.3897/jhr.98.136200</mods:identifier>
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<mods:identifier id="8F27540BC7BD6C700BDF222A5D980C95" type="ZooBank">77736D1C-30F9-4D6C-8310-3A04121CACE2</mods:identifier>
|
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</mods:mods>
|
||||
<treatment id="EC531111868A5194B7C353CA91169FF3" ID-arpha="EC531111-868A-5194-B7C3-53CA91169FF3" LSID="urn:lsid:plazi:treatment:EC531111868A5194B7C353CA91169FF3" httpUri="http://treatment.plazi.org/id/EC531111868A5194B7C353CA91169FF3">
|
||||
<subSubSection id="0A4C08DDBB6A507F51CB862A6321E71F" type="nomenclature">
|
||||
<paragraph id="DD2AFE71D0F8DAC3018755A9B7A58850">
|
||||
<heading id="83C532EE592177F90D53F6CA38CC1BD2" reason="label">Family †</heading>
|
||||
<taxonomicName id="423EE7C618D69811ED7905643AC5693A" authority="Rasnitsyn, 1975" authorityName="Rasnitsyn" authorityYear="1975" class="Insecta" family="Falsiformicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="family">
|
||||
Falsiformicidae
|
||||
<bibRefCitation id="ED085237EED0BB07CC33A5841A47EC60" author="Rasnitsyn AP" journalOrPublisher="Transaction of the Paleontological Institute, Academy of Sciences of the USSR" pagination="1-134" refId="B19" refString="Rasnitsyn AP (1975) Hymenoptera Apocrita of Mesozoic. Transaction of the Paleontological Institute, Academy of Sciences of the USSR 147: 1 - 134." title="Hymenoptera Apocrita of Mesozoic." volume="147" year="1975">Rasnitsyn, 1975</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E3IAE" type="type genus">
|
||||
<paragraph id="A5D41DC4DFC07D128B2EC33A6A3C7016">
|
||||
<heading id="0B656EF093141622840ECDF61640F970" reason="title">
|
||||
<typeStatus id="6DA9B20EB097243E729D39DAB8FD560F">Type</typeStatus>
|
||||
genus.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B42F968C96ECF6F54F1A81C111D18C95">
|
||||
†
|
||||
<taxonomicName id="C1C3211AA63E00EC09AA1AC532C20405" authority="Rasnitsyn, 1975" authorityName="Rasnitsyn" authorityYear="1975" class="Insecta" family="Falsiformicidae" genus="Falsiformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="2DDCB9032717FBA78536835616801F96" italics="true">Falsiformica</emphasis>
|
||||
<bibRefCitation id="1B3F4A0B1AC982A10F64C2994A812681" author="Rasnitsyn AP" journalOrPublisher="Transaction of the Paleontological Institute, Academy of Sciences of the USSR" pagination="1-134" refId="B19" refString="Rasnitsyn AP (1975) Hymenoptera Apocrita of Mesozoic. Transaction of the Paleontological Institute, Academy of Sciences of the USSR 147: 1 - 134." title="Hymenoptera Apocrita of Mesozoic." volume="147" year="1975">Rasnitsyn, 1975</bibRefCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EJJAE" type="Genera included">
|
||||
<paragraph id="629B171F1311064448083BEB4EA04AB9">
|
||||
<heading id="A909AC47E3EA05BBABB87787CAE72594" reason="title">Genera included.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="F1EA1E5E5DE425C27C73B61DE6A2FC31">
|
||||
†
|
||||
<taxonomicName id="DC637B786FDC506D268BD90E9766D5B5" authority="Rasnitsyn, 1975" authorityName="Rasnitsyn" authorityYear="1975" class="Insecta" family="Falsiformicidae" genus="Falsiformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3A4BDF319063F541A3B3ECF68CCB2A75" italics="true">Falsiformica</emphasis>
|
||||
<bibRefCitation id="B1CE1273211DC1C984FEDE7184415F63" author="Rasnitsyn AP" journalOrPublisher="Transaction of the Paleontological Institute, Academy of Sciences of the USSR" pagination="1-134" refId="B19" refString="Rasnitsyn AP (1975) Hymenoptera Apocrita of Mesozoic. Transaction of the Paleontological Institute, Academy of Sciences of the USSR 147: 1 - 134." title="Hymenoptera Apocrita of Mesozoic." volume="147" year="1975">Rasnitsyn, 1975</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, †
|
||||
<taxonomicName id="63863578582AF668D0B64DFBA0D2A909" authority="Cockx & McKellar, 2016" authorityName="Cockx & McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="69BEA02DDF59D0CBD9A3A62FE1667A32" italics="true">Siccibythus</emphasis>
|
||||
<bibRefCitation id="7A8BC4840510FCECA4CFCB28645F6B57" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD & McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx & McKellar, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, †
|
||||
<emphasis id="7C3700686DE696682DF735EA63B25E8E" italics="true">Neophenax</emphasis>
|
||||
<bibRefCitation id="19679402356DE3A49DF83DADF79FA416" author="Engel MS" journalOrPublisher="Entomologist’s Monthly Magazine" pagination="157-168" refId="B5" refString="Engel MS (2022) The first New World Falsiformicidae (Hymenoptera): a new genus in Upper Cretaceous amber from New Jersey. Entomologist's Monthly Magazine 158: 157 - 168. https: // doi: 10.31184 / M 00138908.1583. 4100" title="The first New World Falsiformicidae (Hymenoptera): a new genus in Upper Cretaceous amber from New Jersey." url="https://doi: 10.31184/M00138908.1583.4100" volume="158" year="2022">Engel, 2022</bibRefCitation>
|
||||
,
|
||||
<emphasis id="4B7C80485E82A5550A5CE3A8F828B5AC" italics="true">Falsiformix</emphasis>
|
||||
<bibRefCitation id="9B10E0A747FC847CB0D7111DEA68AF74" DOI="10.1016/j.cretres.2024.106057" author="Zhang Q & Jia N & Li YZ & Wang JX & Rasnitsyn AP" refId="B32" refString="Zhang Q, Jia N, Li YZ, Wang JX, Rasnitsyn AP (2024) New genus of falsiformicid wasps (Hymenoptera, Vespoidea: Falsiformicidae) with brachypterous male from middle Cretaceous Kachin amber in Myanmar. Cretaceous Research. https: // doi. org / 10.1016 / j. cretres. 2024.106057" year="2024">Zhang & Rasnitsyn, 2024</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue