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<document id="0D92F19EB9A9CB9D58D43214E9F2551B" ID-DOI="10.11646/zootaxa.3955.3.9" ID-GBIF-Dataset="a09f0c7f-f65f-47af-bebc-9371363b9653" ID-ISSN="1175-5326" ID-Zenodo-Dep="233117" ID-ZooBank="D018AFA7-2EA9-44BF-AB81-ECBAF132E7AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461051944299" checkinUser="plazi" docAuthor="Häckel, Martin &amp; Anichtchenko, Alexandr" docDate="2015" docId="B85E89401001990CFF5C10A6FD62A9BB" docLanguage="en" docName="zt03955p434.pdf" docOrigin="Zootaxa 3955 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspedophorus luzonensis Häckel &amp; Anichtchenko, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="428" masterDocId="4467F1381000990DFFCB117FFFEEAE78" masterDocTitle="Two new Craspedophorus species (Coleoptera: Carabidae: Panagaeini) from the Philippines" masterLastPageNumber="434" masterPageNumber="427" pageNumber="428" updateTime="1698579237837" updateUser="plazi">
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<mods:title id="5340850BFDD3DD373858722E7DA60C86">Two new Craspedophorus species (Coleoptera: Carabidae: Panagaeini) from the Philippines</mods:title>
</mods:titleInfo>
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<mods:namePart id="2DDD2B4931A587511F1164E9B1F85FAD">Häckel, Martin</mods:namePart>
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<mods:namePart id="8B6BA098974DB95FD643239897162817">Anichtchenko, Alexandr</mods:namePart>
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<mods:title id="CF52FF45B442BCF7A634680D19C78C44">Zootaxa</mods:title>
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<mods:date id="FCA4AD14E941050B097A51301AC7FB8C">2015</mods:date>
<mods:detail id="6F89C222CB0EE0EE6C151CFBC06A4704" type="volume">
<mods:number id="6B9FFAAC03CBBA4B0E6CE478FE72B6E6">3955</mods:number>
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<mods:identifier id="BA53E7B5084181D2DF878F8E80DC13AB" type="DOI">10.11646/zootaxa.3955.3.9</mods:identifier>
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<treatment id="B85E89401001990CFF5C10A6FD62A9BB" ID-DOI="http://doi.org/10.5281/zenodo.6111076" ID-GBIF-Taxon="119612163" ID-Zenodo-Dep="6111076" LSID="urn:lsid:plazi:treatment:B85E89401001990CFF5C10A6FD62A9BB" httpUri="http://treatment.plazi.org/id/B85E89401001990CFF5C10A6FD62A9BB" lastPageNumber="428" pageId="1" pageNumber="428">
<subSubSection id="78ED6BDD1001990CFF5C10A6FF05AC6C" pageId="1" pageNumber="428" type="nomenclature">
<paragraph id="304838561001990CFF5C10A6FDA6AF8B" blockId="1.[151,584,473,533]" box="[151,584,473,499]" pageId="1" pageNumber="428">
<heading id="6B008F3A1001990CFF5C10A6FDA6AF8B" bold="true" box="[151,584,473,499]" fontSize="11" level="1" pageId="1" pageNumber="428" reason="1">
<emphasis id="0283E4441001990CFF5C10A6FDA6AF8B" bold="true" box="[151,584,473,499]" pageId="1" pageNumber="428">
<taxonomicName id="F7F743D51001990CFF5C10A6FE0AAF8B" ID-CoL="Z5X4" box="[151,484,473,499]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="0283E4441001990CFF5C10A6FE0AAF8B" bold="true" box="[151,484,473,499]" italics="true" pageId="1" pageNumber="428">Craspedophorus luzonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19B0593F1001990CFE2010A6FDA6AF8B" box="[491,584,473,499]" pageId="1" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="304838561001990CFF5C1083FF05AC6C" blockId="1.[151,584,473,533]" box="[151,235,508,533]" pageId="1" pageNumber="428">
(
<figureCitation id="A8CC24D31001990CFF541083FF0EAC6C" box="[159,224,508,533]" captionStart="FIGURE 1" captionStartId="2.[151,250,1957,1979]" captionTargetBox="[390,1212,213,1932]" captionTargetId="figure@2.[340,1248,195,1933]" captionTargetPageId="2" captionText="FIGURE 1. Craspedophorus luzonensis sp. nov.: A. Holotype, B. Lateral view of aedeagus." httpUri="https://zenodo.org/record/233118/files/figure.png" pageId="1" pageNumber="428">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="78ED6BDD1001990CFF5C133BFF31ACDC" pageId="1" pageNumber="428" type="materials_examined">
<paragraph id="304838561001990CFF5C133BFF31ACDC" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
<emphasis id="0283E4441001990CFF5C133BFEAAAC25" bold="true" box="[151,324,580,605]" pageId="1" pageNumber="428">
<typeStatus id="EF4C86F41001990CFF5C133BFF3CAC25" box="[151,210,580,605]" pageId="1" pageNumber="428">Type</typeStatus>
material.
</emphasis>
<typeStatus id="EF4C86F41001990CFE80133BFE5DAC25" box="[331,435,580,605]" pageId="1" pageNumber="428" type="holotype">Holotype</typeStatus>
♂: “
<collectingCountry id="48E078C61001990CFE23133BFD85AC25" box="[488,619,580,605]" name="Philippines" pageId="1" pageNumber="428">Philippines</collectingCountry>
, N Luzon, Mt. Province, Mt. Polis, 7. 2014, lgt. loc. collector (
<collectionCode id="56E6A0931001990CFAEC133BFA95AC24" box="[1319,1403,580,604]" pageId="1" pageNumber="428">DUBC</collectionCode>
); 1
<typeStatus id="EF4C86F41001990CFF5C1317FF12ACF8" box="[151,252,616,640]" pageId="1" pageNumber="428" type="paratype">Paratype</typeStatus>
, ♀:
<collectingCountry id="48E078C61001990CFEE71318FE44ACF8" box="[300,426,615,640]" name="Philippines" pageId="1" pageNumber="428">Philippines</collectingCountry>
[C Luzon Island], Nueva Vizcaya, Dupax del Sur,
<date id="44491E961001990CFC261318FB48ACF8" box="[1005,1190,615,640]" pageId="1" pageNumber="428" value="2012-11">November 2012</date>
, lgt. Ismael Lumawig (
<collectionCode id="56E6A0931001990CFF5413F3FF3DACDC" box="[159,211,652,676]" httpUri="http://grbio.org/cool/vxkr-andy" name="Centro de Investigaciones Biologicas del Noroeste, S.C. (Mexico)" pageId="1" pageNumber="428">CIB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="78ED6BDD1001990CFF0C13D0FBDAA903" pageId="1" pageNumber="428" type="description">
<paragraph id="304838561001990CFF0C13D0FD39AC94" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
<emphasis id="0283E4441001990CFF0C13D0FE08ACB0" bold="true" box="[199,486,687,712]" pageId="1" pageNumber="428">
Description of
<typeStatus id="EF4C86F41001990CFEB313D0FE0FACB0" box="[376,481,687,712]" pageId="1" pageNumber="428" type="holotype">holotype</typeStatus>
.
</emphasis>
Length
<quantity id="F70F95B31001990CFD8F13CFFD78ACBF" box="[580,662,688,712]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="428" unit="mm" value="17.0">17 mm</quantity>
, width
<quantity id="F70F95B31001990CFD2113D0FCADACBF" box="[746,835,687,712]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="428" unit="mm" value="6.5">6.5 mm</quantity>
. Proportions: Pronotum 1.27x wider than long, 1.67x wider than head, elytra 1.41x wider than pronotum.
</paragraph>
<paragraph id="304838561001990CFF0C1388FC0FADF9" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">Colour: Head, pronotum and elytra black, glossy; mandibles, palps, tarsi, base of labrum, base of antennomere IV lighter, reddish. Legs and antennae ferrugineous, proximal parts of femur darkish. Elytron with two yellow spots; humeral spot reaches from 5th interval (4th stria) to margin and to epipleura, preapical spot reaches from 4th to 8th interval. Dorsal and ventral part of body black, covered by yellow setae.</paragraph>
<paragraph id="304838561001990CFF0C12F2FF11AA22" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
<emphasis id="0283E4441001990CFF0C12F2FEEAADDD" box="[199,260,909,933]" italics="true" pageId="1" pageNumber="428">Head</emphasis>
large, wide, only 1.67x narrower than pronotum; eyes moderately convex; temples short; labrum strongly excavated at anterior side; clypeus smooth, with four long, yellow setae; vertex convex, sparsely punctured and covered by long, reddish setae; anterior part of frons smooth, fairly convex at middle, laterally depressed, rugate, lateral margin with sulcus rimming it from mandibular base to anterior margin of eyes, anterior side arcuate, sinuate before eyes, moderately impressed, smooth, sharply bordered externally, indistinctly bordered medially.
</paragraph>
<paragraph id="304838561001990CFF0C1519FC13AB4A" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
<emphasis id="0283E4441001990CFF0C1519FED8AA05" box="[199,310,1126,1149]" italics="true" pageId="1" pageNumber="428">Pronotum</emphasis>
weakly transverse (length to width ratio 1.27), almost hexagonal, widest at middle; lateral angles widely rounded, weakly arcuate anteriorly; anterior angles weakly rounded and distinctly protruded; anterior margin as wide as neck, flat between angular lobes; lateral margins weakly sinuate toward base; base parallel, oblique laterally near posterior angle; lateral rims punctured, narrowing anteriorly, posteriorly wider and more elevated; disc densely rugate and punctured; sagittal line distinctly impressed, reaching neither anterior margin nor base; basal pits elongate, narrowly and deeply impressed, coarsely punctured.
</paragraph>
<paragraph id="304838561001990CFF0C1442FDB2ABBA" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
<emphasis id="0283E4441001990CFF0C1442FEE3AB2D" box="[199,269,1341,1365]" italics="true" pageId="1" pageNumber="428">Elytra</emphasis>
almost parallel-sided, widest at middle, elongate, slightly convex, weakly tuberose in front of apex; disc flattened; humerus weakly developed; margins rounded, widening posterio-laterally; basal rim indistinct striae deeply impressed and very slightly punctured; intervals moderately convex, sparsely but coarsely punctured, interspaces with distinct microsculpture.
</paragraph>
<paragraph id="304838561001990CFF0C14B1FBF2A871" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
<emphasis id="0283E4441001990CFF0C14B1FEE2AB9D" box="[199,268,1486,1509]" italics="true" pageId="1" pageNumber="428">Venter</emphasis>
black, metepisternum transverse, punctured, weakly elongated and narrowed towards posterior. Ventrites laterally rugate, finely rugate at middle, densely covered by long setae.
</paragraph>
<paragraph id="304838561001990CFF0C176AFAE0A856" blockId="1.[151,1437,580,1987]" box="[199,1294,1557,1582]" pageId="1" pageNumber="428">
Aedeagal median lobe stout, strongly curved laterally, apical lamella short, apex rounded (
<figureCitation id="A8CC24D31001990CFB70176AFB12A856" box="[1211,1276,1557,1582]" captionStart="FIGURE 1" captionStartId="2.[151,250,1957,1979]" captionTargetBox="[390,1212,213,1932]" captionTargetId="figure@2.[340,1248,195,1933]" captionTargetPageId="2" captionText="FIGURE 1. Craspedophorus luzonensis sp. nov.: A. Holotype, B. Lateral view of aedeagus." httpUri="https://zenodo.org/record/233118/files/figure.png" pageId="1" pageNumber="428">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="304838561001990CFF0C1746FBDAA903" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
<emphasis id="0283E4441001990CFF0C1746FE3FA82A" bold="true" box="[199,465,1593,1618]" pageId="1" pageNumber="428">Differential diagnosis.</emphasis>
Based on similar habitus and elytral pattern,
<taxonomicName id="F7F743D51001990CFC201745FB6BA829" box="[1003,1157,1593,1617]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="0283E4441001990CFC201745FB6BA829" box="[1003,1157,1593,1617]" italics="true" pageId="1" pageNumber="428">C. luzonensis</emphasis>
</taxonomicName>
<emphasis id="0283E4441001990CFB441746FB03A82A" bold="true" box="[1167,1261,1593,1618]" pageId="1" pageNumber="428">
<taxonomicNameLabel id="19B0593F1001990CFB441746FB03A82A" box="[1167,1261,1593,1618]" pageId="1" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the
<emphasis id="0283E4441001990CFF5C1722FEE2A80D" box="[151,268,1629,1653]" italics="true" pageId="1" pageNumber="428">“obscurus</emphasis>
species group sensu
<bibRefCitation id="546645A71001990CFDEA1722FC47A80E" author="Hackel" box="[545,937,1629,1654]" pageId="1" pageNumber="428" refString="Hackel, M. &amp; Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel &amp; Kirschenhofer (2014)</bibRefCitation>
. It resembles
<taxonomicName id="F7F743D51001990CFBA91721FE90A8E2" authority="Hackel &amp; Kirschenhofer, 2014" authorityName="Hackel &amp; Kirschenhofer" authorityYear="2014" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="cenwanglao">
<emphasis id="0283E4441001990CFBA91721FAFAA80D" box="[1122,1300,1629,1653]" italics="true" pageId="1" pageNumber="428">C. cenwanglao</emphasis>
<bibRefCitation id="546645A71001990CFAEE1722FE90A8E2" author="Hackel" pageId="1" pageNumber="428" refString="Hackel, M. &amp; Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel &amp; Kirschenhofer, 2014</bibRefCitation>
</taxonomicName>
(
<figureCitation id="A8CC24D31001990CFE4417FEFE3AA8E2" box="[399,468,1665,1690]" captionStart="FIGURE 2. C" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[412,1147,238,1901]" captionTargetId="figure@3.[369,1218,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. C. cenwanglao Häckel &amp; Kirschenhofer, 2014: A. Paratype, B. Lateral view of aedeagus." httpUri="https://zenodo.org/record/233119/files/figure.png" pageId="1" pageNumber="428">Fig. 2</figureCitation>
) from western Guangxi,
<collectingCountry id="48E078C61001990CFD3E17FEFCD5A8E2" box="[757,827,1665,1690]" name="China" pageId="1" pageNumber="428">China</collectingCountry>
, and
<taxonomicName id="F7F743D51001990CFCB217FDFA72A8E2" authority="Hackel &amp; Kirschenhofer, 2014" authorityName="Hackel &amp; Kirschenhofer" authorityYear="2014" box="[889,1436,1665,1690]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="phoupanensis">
<emphasis id="0283E4441001990CFCB217FDFBDBA8E1" box="[889,1077,1665,1689]" italics="true" pageId="1" pageNumber="428">C. phoupanensis</emphasis>
<bibRefCitation id="546645A71001990CFBF517FEFA72A8E2" author="Hackel" box="[1086,1436,1665,1690]" pageId="1" pageNumber="428" refString="Hackel, M. &amp; Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel &amp; Kirschenhofer, 2014</bibRefCitation>
</taxonomicName>
from northern
<collectingCountry id="48E078C61001990CFEF717D9FE9BA8C5" box="[316,373,1702,1725]" name="Laos" pageId="1" pageNumber="428">Laos</collectingCountry>
, particularly in the pronotum. The new species differs from them by somewhat more elongate habitus, less convex elytra, elytra with more grossly and deeply punctured striae and more extended spots; the humeral spot extends from 4th stria (external part of 4th interval) to margin and the preapical spot extends from 3rd stria (4th interval) to 9th interval, contrary to both continental species in which the humeral spot reaches from external part of 5th interval and the preapical spot from 4th stria (5th interval) to 9th interval (
<figureCitation id="A8CC24D31001990CFB721642FAEFA92E" box="[1209,1281,1853,1878]" captionStart="FIGURE 2. C" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[412,1147,238,1901]" captionTargetId="figure@3.[369,1218,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. C. cenwanglao Häckel &amp; Kirschenhofer, 2014: A. Paratype, B. Lateral view of aedeagus." httpUri="https://zenodo.org/record/233119/files/figure.png" pageId="1" pageNumber="428">Fig. 2</figureCitation>
). In addition, intervals are less dense and more coarsely punctured than in
<taxonomicName id="F7F743D51001990CFCF5161DFC3AA902" box="[830,980,1890,1914]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="0283E4441001990CFCF5161DFC3AA902" box="[830,980,1890,1914]" italics="true" pageId="1" pageNumber="428">C. luzonensis</emphasis>
</taxonomicName>
<emphasis id="0283E4441001990CFC10161DFBDAA903" bold="true" box="[987,1076,1890,1915]" pageId="1" pageNumber="428">
<taxonomicNameLabel id="19B0593F1001990CFC10161DFBDAA903" box="[987,1076,1890,1915]" pageId="1" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="78ED6BDD1001990CFF0C16FAFB61A9E6" box="[199,1167,1925,1950]" pageId="1" pageNumber="428" type="etymology">
<paragraph id="304838561001990CFF0C16FAFB61A9E6" blockId="1.[151,1437,580,1987]" box="[199,1167,1925,1950]" pageId="1" pageNumber="428">
<emphasis id="0283E4441001990CFF0C16FAFEA0A9E6" bold="true" box="[199,334,1925,1950]" pageId="1" pageNumber="428">Etymology.</emphasis>
Named after the place of its occurrence - Luzon, island in the
<collectingCountry id="48E078C61001990CFBC216FAFB65A9E6" box="[1033,1163,1925,1950]" name="Philippines" pageId="1" pageNumber="428">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="78ED6BDD1001990CFF0C16D5FD62A9BB" box="[199,652,1962,1987]" pageId="1" pageNumber="428" type="distribution">
<paragraph id="304838561001990CFF0C16D5FD62A9BB" blockId="1.[151,1437,580,1987]" box="[199,652,1962,1987]" pageId="1" pageNumber="428">
<emphasis id="0283E4441001990CFF0C16D5FE8EA9BB" bold="true" box="[199,352,1962,1987]" pageId="1" pageNumber="428">Distribution.</emphasis>
Luzon island,
<collectingCountry id="48E078C61001990CFDCD16D5FD66A9BB" box="[518,648,1962,1987]" name="Philippines" pageId="1" pageNumber="428">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="4E20ADB46CF491FEE838BEC40E765E12">Two new Craspedophorus species (Coleoptera: Carabidae: Panagaeini) from the Philippines</mods:title>
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<mods:namePart id="110E7342BF4B97998B78C4A47C419F88">Häckel, Martin</mods:namePart>
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<treatment id="B85E894010049909FF5C11E8FB68A8D8" ID-DOI="http://doi.org/10.5281/zenodo.6111078" ID-GBIF-Taxon="119612162" ID-Zenodo-Dep="6111078" LSID="urn:lsid:plazi:treatment:B85E894010049909FF5C11E8FB68A8D8" httpUri="http://treatment.plazi.org/id/B85E894010049909FF5C11E8FB68A8D8" lastPageNumber="431" pageId="4" pageNumber="431">
<subSubSection id="78ED6BDD10049909FF5C11E8FF05AEAB" pageId="4" pageNumber="431" type="nomenclature">
<paragraph id="3048385610049909FF5C11E8FD95AECA" blockId="4.[151,635,151,211]" box="[151,635,151,178]" pageId="4" pageNumber="431">
<heading id="6B008F3A10049909FF5C11E8FD95AECA" bold="true" box="[151,635,151,178]" fontSize="11" level="1" pageId="4" pageNumber="431" reason="1">
<emphasis id="0283E44410049909FF5C11E8FD95AECA" bold="true" box="[151,635,151,178]" pageId="4" pageNumber="431">
<taxonomicName id="F7F743D510049909FF5C11E8FDF8AEC9" ID-CoL="Z5WD" box="[151,534,151,177]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="kirschenhoferi" status="sp. nov.">
<emphasis id="0283E44410049909FF5C11E8FDF8AEC9" bold="true" box="[151,534,151,177]" italics="true" pageId="4" pageNumber="431">Craspedophorus kirschenhoferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19B0593F10049909FDD611E7FD95AECA" box="[541,635,152,178]" pageId="4" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3048385610049909FF5C11C5FF05AEAB" blockId="4.[151,635,151,211]" box="[151,235,186,211]" pageId="4" pageNumber="431">
(
<figureCitation id="A8CC24D310049909FF5411C5FF0CAEAB" box="[159,226,186,211]" captionStart="FIGURE 3. C" captionStartId="5.[151,250,1954,1976]" captionText="FIGURE 3. C. kirschenhoferi sp. nov.: A. Holotype, B. Lateral view of aedeagus." pageId="4" pageNumber="431">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="78ED6BDD10049909FF5C107DFAB8AF47" pageId="4" pageNumber="431" type="materials_examined">
<paragraph id="3048385610049909FF5C107DFAB8AF47" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
<emphasis id="0283E44410049909FF5C107DFEAEAF63" bold="true" box="[151,320,258,283]" pageId="4" pageNumber="431">
<typeStatus id="EF4C86F410049909FF5C107DFF3CAF63" box="[151,210,258,283]" pageId="4" pageNumber="431">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="EF4C86F410049909FE9B107DFE56AF63" box="[336,440,258,283]" pageId="4" pageNumber="431" type="holotype">Holotype</typeStatus>
♂: “
<collectingCountry id="48E078C610049909FE39107DFD9BAF63" box="[498,629,258,283]" name="Philippines" pageId="4" pageNumber="431">Philippines</collectingCountry>
, Mindanao, Surigao del Norte.
<date id="44491E9610049909FC29107CFBA7AF63" box="[994,1097,259,283]" pageId="4" pageNumber="431" value="2014-03">III. 2014</date>
, local collector leg. (
<collectionCode id="56E6A09310049909FAF7107CFA7EAF63" box="[1340,1424,259,283]" pageId="4" pageNumber="431">DUBC</collectionCode>
).
<typeStatus id="EF4C86F410049909FF5C1058FF12AF47" box="[151,252,295,319]" pageId="4" pageNumber="431" type="paratype">Paratype</typeStatus>
, 1 ♀: “
<collectingCountry id="48E078C610049909FE821059FE22AF47" box="[329,460,294,319]" name="Philippines" pageId="4" pageNumber="431">Philippines</collectingCountry>
, Mindanao, Bukidnon. Mt. Kalatungan.
<date id="44491E9610049909FC591058FBE5AF47" box="[914,1035,295,319]" pageId="4" pageNumber="431" value="2014-08">VIII. 2014</date>
, local collector leg. (
<collectionCode id="56E6A09310049909FB3E1058FAA5AF47" box="[1269,1355,295,319]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="431">NMPC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="78ED6BDD10049909FF0C1035FE8CA84C" pageId="4" pageNumber="431" type="description">
<paragraph id="3048385610049909FF0C1035FD7EAFFE" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
<emphasis id="0283E44410049909FF0C1035FEB7AF1B" bold="true" box="[199,345,330,355]" pageId="4" pageNumber="431">Description.</emphasis>
<typeStatus id="EF4C86F410049909FEAA1035FE27AF1B" box="[353,457,330,355]" pageId="4" pageNumber="431" type="holotype">Holotype</typeStatus>
length 11.5-12.0 mm, width
<quantity id="F70F95B310049909FCD21035FC4EAF1A" box="[793,928,330,355]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="4" pageNumber="431" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5-4.8 mm</quantity>
. Pronotum 1.37 wider than long, 1.62 wider than head, elytra 1.22x wider than pronotum.
</paragraph>
<paragraph id="3048385610049909FF0C10EDFBBCAC38" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">Colour: Head, pronotum and elytra black. Elytra near margins densely covered by yellow setae, each elytron with two yellow spots. Humeral spot rounded, extending from 4th interval to margin and epipleura. Preapical spot smaller, rounded, reaching from 4th stria to 8th interval, separated from margin. Antennae, epistoma and legs black proximally; antennomeres IV–XI, tarsi and terminal parts of mandibles distally ferrugineous. Venter black, covered sparsely by long, yellow setae. Epipleura light yellowish, laterally near humeral spot.</paragraph>
<paragraph id="3048385610049909FF0C1333FB4CACA9" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
<emphasis id="0283E44410049909FF0C1333FEEAAC1C" box="[199,260,588,612]" italics="true" pageId="4" pageNumber="431">Head</emphasis>
large and wide, only 1.62x narrower than pronotum; frons medially smooth and weakly convex, laterally coarsely punctured, frontal margins rimmed by narrowly sulcate sharp furrows continuing posteriorly and medially between eyes; vertex coarsely punctured; clypeus smooth, shiny, medially upturned; anterior side of labrum concave; eyes strongly convex; temples barely indicated; neck convex, glossy, finely rugate.
</paragraph>
<paragraph id="3048385610049909FF0C13A2FEEBADB5" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
<emphasis id="0283E44410049909FF0C13A2FED8AC8C" box="[199,310,733,756]" italics="true" pageId="4" pageNumber="431">Pronotum</emphasis>
transverse, 1.37x wider than long, almost hexagonal, widest immediately before middle, lateral angles round; anterior margin flat, parallel with base, as long as base; lateral margins narrowing weakly forward; anterior angles indistinct, widely rounded and not extending forward; posterior margins narrow obliquely, in front of posterior angles become weakly sinuate without any indentation; middle of base weakly sinuate, margin weakly oblique; anterior side of lateral rims indistinct, near base wider, more excavated, coarsely punctured; disc weakly convex and punctured more coarsely than rims; sagittal line indistinct, interrupted; basal pits very shallow, weakly indicated.
</paragraph>
<paragraph id="3048385610049909FF0C12A7FB71AA27" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
<emphasis id="0283E44410049909FF0C12A7FEE3AD88" box="[199,269,984,1008]" italics="true" pageId="4" pageNumber="431">Elytra</emphasis>
moderately convex, almost parallel-sided, 1.22x wider than pronotum, weakly widening toward maximum width at middle, distinctly tuberose before apex; basal rim discernible only medial of 4th interval; humeri weakly developed, moderately oblique, widely rounded; striae deeply impressed, coarsely punctured; intervals convex, irregularly punctured; dorsum with black setae, on elytral spots setae yellowish red.
</paragraph>
<paragraph id="3048385610049909FF0C1513FAC7AAFB" blockId="4.[151,1437,258,1696]" box="[199,1321,1131,1156]" pageId="4" pageNumber="431">
<emphasis id="0283E44410049909FF0C1513FEE2AAFB" box="[199,268,1132,1155]" italics="true" pageId="4" pageNumber="431">Venter</emphasis>
black, glossy, metepisterna almost square, narrowed posteriorly, punctured, covered by setae.
</paragraph>
<paragraph id="3048385610049909FF0C15F1FBBBAADF" blockId="4.[151,1437,258,1696]" box="[199,1109,1166,1191]" pageId="4" pageNumber="431">
Aedeagus (damaged in mounting), median lobe strongly curved laterally (
<figureCitation id="A8CC24D310049909FBC815F1FBA9AADF" box="[1027,1095,1166,1191]" captionStart="FIGURE 3. C" captionStartId="5.[151,250,1954,1976]" captionText="FIGURE 3. C. kirschenhoferi sp. nov.: A. Holotype, B. Lateral view of aedeagus." pageId="4" pageNumber="431">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="3048385610049909FF0C15CCFE8CA84C" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
<emphasis id="0283E44410049909FF0C15CCFE3AAAB4" bold="true" box="[199,468,1203,1228]" pageId="4" pageNumber="431">Differential diagnosis.</emphasis>
<taxonomicName id="F7F743D510049909FE2A15CCFCA5AAB3" box="[481,843,1203,1227]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="kirschenhoferi" status="sp. nov.">
<emphasis id="0283E44410049909FE2A15CCFCA5AAB3" box="[481,843,1203,1227]" italics="true" pageId="4" pageNumber="431">Craspedophorus kirschenhoferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19B0593F10049909FC9315CCFC56AAB4" box="[856,952,1203,1228]" pageId="4" pageNumber="431" rank="species">
<emphasis id="0283E44410049909FC9315CCFC56AAB4" bold="true" box="[856,952,1203,1228]" pageId="4" pageNumber="431">sp. nov.</emphasis>
</taxonomicNameLabel>
resembles species of the
<emphasis id="0283E44410049909FB3F15CCFA72AAB3" box="[1268,1436,1203,1227]" italics="true" pageId="4" pageNumber="431">
“
<taxonomicName id="F7F743D510049909FB3715CCFA72AAB3" box="[1276,1436,1203,1227]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="microspilotus">microspilotus</taxonomicName>
</emphasis>
species group sensu
<bibRefCitation id="546645A710049909FEB515A9FD9FAA97" author="Kirschenhofer" box="[382,625,1238,1263]" pageId="4" pageNumber="431" refString="Kirschenhofer, E. (2000) Neue und wenig bekannte Panagaeini der ostlichen Palaarktis sowie der Orientalis. (Coleoptera, Carabidae). Entomofauna, 21 (29), 321 - 372." type="journal article" year="2000">Kirschenhofer (2000)</bibRefCitation>
by its body size and shape, but differs clearly from most species of that group by markedly less transverse, hexagonal pronotum. The two species of that group with most similar pronotum,
<taxonomicName id="F7F743D510049909FED8145FFCA4AB4F" authority="Hackel &amp; Kirschenhofer, 2014" authorityName="Hackel &amp; Kirschenhofer" authorityYear="2014" box="[275,842,1311,1336]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="chiangdaoensis">
<emphasis id="0283E44410049909FED8145FFE0DAB4F" box="[275,483,1311,1335]" italics="true" pageId="4" pageNumber="431">C. chiangdaoensis</emphasis>
<bibRefCitation id="546645A710049909FE271460FCA4AB4F" author="Hackel" box="[492,842,1311,1336]" pageId="4" pageNumber="431" refString="Hackel, M. &amp; Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel &amp; Kirschenhofer, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="F7F743D510049909FC4F145FFA76AB4F" authority="Hackel &amp; Kirschenhofer, 2014" authorityName="Hackel &amp; Kirschenhofer" authorityYear="2014" box="[900,1432,1311,1336]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="kiwlomensis">
<emphasis id="0283E44410049909FC4F145FFBC1AB4F" box="[900,1071,1311,1335]" italics="true" pageId="4" pageNumber="431">C. kiwlomensis</emphasis>
<bibRefCitation id="546645A710049909FBF31460FA76AB4F" author="Hackel" box="[1080,1432,1311,1336]" pageId="4" pageNumber="431" refString="Hackel, M. &amp; Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel &amp; Kirschenhofer, 2014</bibRefCitation>
</taxonomicName>
, differ inter alia by elytral pattern with distinctly larger spots.
<taxonomicName id="F7F743D510049909FCA8143CFBC7AB23" box="[867,1065,1347,1371]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="kirschenhoferi" status="sp. nov.">
<emphasis id="0283E44410049909FCA8143CFBC7AB23" box="[867,1065,1347,1371]" italics="true" pageId="4" pageNumber="431">C. kirschenhoferi</emphasis>
</taxonomicName>
<emphasis id="0283E44410049909FBF8143CFB61AB24" bold="true" box="[1075,1167,1347,1372]" pageId="4" pageNumber="431">
<taxonomicNameLabel id="19B0593F10049909FBF8143CFB61AB24" box="[1075,1167,1347,1372]" pageId="4" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also weakly resembles species of
<taxonomicName id="F7F743D510049909FED81417FE27AB07" box="[275,457,1383,1407]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="basifasciatus">
<emphasis id="0283E44410049909FED81417FE27AB07" box="[275,457,1383,1407]" italics="true" pageId="4" pageNumber="431">C. basifasciatus</emphasis>
</taxonomicName>
group sensu
<bibRefCitation id="546645A710049909FDAD1418FC3AAB07" author="Hackel" box="[614,980,1383,1408]" pageId="4" pageNumber="431" refString="Hackel, M. &amp; Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel &amp; Kirschenhofer (2014)</bibRefCitation>
by similar pronotum, but differs in the elytral pattern with spots without serrate margins. Another very similar but larger (
<quantity id="F70F95B310049909FB5D14F4FAEBABDB" box="[1174,1285,1419,1444]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="431" unit="mm" value="13.5">13.5 mm</quantity>
) species,
<taxonomicName id="F7F743D510049909FA4914F4FDCCABBF" authority="Kirschenhofer, 2011" authorityName="Kirschenhofer" authorityYear="2011" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="soppongensis">
<emphasis id="0283E44410049909FA4914F4FEC0ABBF" italics="true" pageId="4" pageNumber="431">C. soppongensis</emphasis>
Kirschenhofer, 2011
</taxonomicName>
(
<emphasis id="0283E44410049909FDFC14D0FD7CABBF" box="[567,658,1455,1479]" italics="true" pageId="4" pageNumber="431">“kubani</emphasis>
species group) from northern
<collectingCountry id="48E078C610049909FC3614D0FB8FABB0" box="[1021,1121,1455,1480]" name="Thailand" pageId="4" pageNumber="431">Thailand</collectingCountry>
(
<figureCitation id="A8CC24D310049909FBBF14D0FB52ABBF" box="[1140,1212,1455,1480]" captionStart="FIGURE 4. C" captionStartId="6.[151,250,1952,1974]" captionTargetBox="[315,1272,215,1916]" captionTargetId="figure@6.[303,1283,193,1930]" captionTargetPageId="6" captionText="FIGURE 4. C. soppongenis Kirschenhofer, 2011 male, holotype. Scale bar: 10 mm." httpUri="https://zenodo.org/record/233120/files/figure.png" pageId="4" pageNumber="431">Fig. 4</figureCitation>
) has the pronotum similarly shaped but differently sculptured; the elytra shows a similar pattern but is differently sculptured; the sagittal line of pronotum is narrow and deeply impressed, and the pronotal basal pits are narrow, elongated and deeply impressed.
</paragraph>
</subSubSection>
<subSubSection id="78ED6BDD10049909FF0C1740FB41A803" pageId="4" pageNumber="431" type="etymology">
<paragraph id="3048385610049909FF0C1740FB41A803" blockId="4.[151,1437,258,1696]" pageId="4" pageNumber="431">
<emphasis id="0283E44410049909FF0C1740FEA0A820" bold="true" box="[199,334,1599,1624]" pageId="4" pageNumber="431">Etymology.</emphasis>
Named after our colleague and friend Erich Kirschenhofer (Perchtoldsdorf,
<collectingCountry id="48E078C610049909FB7F1740FAE5A820" box="[1204,1291,1599,1624]" name="Austria" pageId="4" pageNumber="431">Austria</collectingCountry>
), a specialist in the Panagaeini and
<taxonomicName id="F7F743D510049909FE45171CFD98A804" box="[398,630,1635,1660]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="tribe" tribe="Licinini">Licinini (Carabidae)</taxonomicName>
. The specific epithet is a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="78ED6BDD10049909FF0C17F8FB68A8D8" box="[199,1158,1671,1696]" pageId="4" pageNumber="431" type="distribution">
<paragraph id="3048385610049909FF0C17F8FB68A8D8" blockId="4.[151,1437,258,1696]" box="[199,1158,1671,1696]" pageId="4" pageNumber="431">
<emphasis id="0283E44410049909FF0C17F8FE8EA8D8" bold="true" box="[199,352,1671,1696]" pageId="4" pageNumber="431">Distribution.</emphasis>
So far known only from two localities in Mindanao Island,
<collectingCountry id="48E078C610049909FBCB17F8FB6CA8D8" box="[1024,1154,1671,1696]" name="Philippines" pageId="4" pageNumber="431">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>