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<document id="73B8AC00D222C03675AFFCA964B85D8C" ID-DOI="10.1127/njgpa/2024/1195" ID-Zenodo-Dep="15237573" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1744912398416" checkinUser="guilherme" docAuthor="Burckhardt, Daniel, Drohojowska, Jowita, Štarhová Serbina, Liliya &amp; Malenovský, Igor" docDate="2024" docId="03E387941573FF91FCF4FA10FE622873" docLanguage="en" docName="NJbGeolPalaontAbh.311.2.215-227.pdf" docOrigin="Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen (Basel, Switzerland) 311 (2)" docSource="https://doi.org/10.1127/njgpa/2024/1195" docTitle="Diclidophlebia venosa Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="219" masterDocId="FFDAFFEC1572FF94FF89FFE1FF8A2C25" masterDocTitle="First record of jumping plant lice of the family Liviidae (Hemiptera, Sternorrhyncha, Psylloidea) from Dominican amber" masterLastPageNumber="227" masterPageNumber="215" pageNumber="216" updateTime="1744913668726" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="FECDA2F4C83897FC60D407FE459FDB35">First record of jumping plant lice of the family Liviidae (Hemiptera, Sternorrhyncha, Psylloidea) from Dominican amber</mods:title>
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<mods:namePart id="6C0147CF3183255FF59AF083C70FAF35">Burckhardt, Daniel</mods:namePart>
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<mods:namePart id="B57A5493C8A67823EC4785F4695EC2E1">Drohojowska, Jowita</mods:namePart>
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<mods:namePart id="BF8C1A1444C19CE15BB122E5476E8072">Štarhová Serbina, Liliya</mods:namePart>
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<mods:namePart id="66229202430CFF270F591797E3AA743F">Malenovský, Igor</mods:namePart>
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<mods:title id="6EEE6A349153EECD2AC4DFE558A48DAD">Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen</mods:title>
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<mods:date id="33E2244A0E44449D2602177F856637B6">2024</mods:date>
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<mods:title id="7C9F7CF1E17DDA57DDC6963082BC6530">Basel, Switzerland</mods:title>
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<subSubSection id="C35065091573FF95FCF4FA10FB6B2A08" pageId="1" pageNumber="216" type="nomenclature">
<paragraph id="8BF536821573FF95FCF4FA10FB6B2A08" blockId="1.[893,1342,1521,1614]" pageId="1" pageNumber="216">
<heading id="D0BD81EE1573FF95FCF4FA10FB6B2A08" centered="true" fontSize="11" level="2" pageId="1" pageNumber="216" reason="4">
<taxonomicName id="4C4A4D011573FF95FCF4FA10FB0E2A08" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="venosa" status="sp. nov.">
<emphasis id="B93EEA901573FF95FCF4FA10FBF12A2E" box="[893,1147,1521,1547]" italics="true" pageId="1" pageNumber="216">Diclidophlebia venosa</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB1573FF95FB02F9F2FB6B2A08" box="[1163,1249,1555,1581]" pageId="1" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35065091573FF95FB9FF9D5FBF92A97" pageId="1" pageNumber="216" type="description">
<paragraph id="8BF536821573FF95FB9FF9D5FB2C2A6B" blockId="1.[893,1342,1521,1614]" box="[1046,1190,1588,1614]" pageId="1" pageNumber="216">
<figureCitation id="13712A071573FF95FB9FF9D5FB002A6B" box="[1046,1162,1588,1614]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="1" pageNumber="216">Fig. 1A, B</figureCitation>
,
<figureCitation id="13712A071573FF95FB11F9D5FB2C2A6B" box="[1176,1190,1588,1614]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="1" pageNumber="216">2</figureCitation>
</paragraph>
<paragraph id="8BF536821573FF95FCA7F997FBF92A97" blockId="1.[814,1421,1654,1714]" pageId="1" pageNumber="216">
<emphasis id="B93EEA901573FF95FCA7F997FCF32AB5" bold="true" box="[814,889,1654,1680]" pageId="1" pageNumber="216">LSID:</emphasis>
urn:lsid:zoobank.org:act:5BB9A3FC-3E02- 4746-A423-9272D3A3DEA6
</paragraph>
</subSubSection>
<subSubSection id="C35065091573FF95FCA7F93AFA072B33" pageId="1" pageNumber="216" type="etymology">
<paragraph id="8BF536821573FF95FCA7F93AFA072B33" blockId="1.[814,1421,1754,1820]" pageId="1" pageNumber="216">
<emphasis id="B93EEA901573FF95FCA7F93AFC3F2AD0" bold="true" box="[814,949,1755,1781]" pageId="1" pageNumber="216">Etymology:</emphasis>
From the Latin adjective venosus (1
<superScript id="7C3F9BCA1573FF95FAD4F93AFAE22ACC" attach="none" box="[1373,1384,1755,1769]" fontSize="6" pageId="1" pageNumber="216">st</superScript>
/2
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declension) = full of veins, referring to the forked vein R
<subScript id="17CE34C71573FF95FAF6F8EFFA0C2B39" attach="right" box="[1407,1414,1806,1820]" fontSize="6" pageId="1" pageNumber="216">1</subScript>
.
</paragraph>
</subSubSection>
<subSubSection id="C35065091573FF90FCA7F8DEFECF2D72" lastPageId="4" lastPageNumber="219" pageId="1" pageNumber="216" type="materials_examined">
<paragraph id="8BF536821573FF95FCA7F8DEFAC72B7C" blockId="1.[814,1357,1854,1881]" box="[814,1357,1854,1881]" pageId="1" pageNumber="216">
<materialsCitation id="3B223CDF1573FF95FCA7F8DEFAC22B7C" box="[814,1352,1854,1881]" collectionCode="GPAC" country="Dominican Republic" pageId="1" pageNumber="216" specimenCount="1" typeStatus="holotype">
<emphasis id="B93EEA901573FF95FCA7F8DEFC2F2B7C" bold="true" box="[814,933,1855,1881]" pageId="1" pageNumber="216">
<typeStatus id="54F188201573FF95FCA7F8DEFC2A2B7C" box="[814,928,1855,1881]" pageId="1" pageNumber="216">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D4CFD0B1573FF95FC25F8DFFBD12B7D" box="[940,1115,1854,1880]" pageId="1" pageNumber="216" type="male">Male specimen</specimenCount>
, sample #24 (
<collectionCode id="ED5BAE471573FF95FB7CF8DEFACB2B7C" box="[1269,1345,1855,1881]" pageId="1" pageNumber="216">GPAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<paragraph id="8BF536821570FF96FF2FF8EDFB942BB0" blockId="2.[166,1415,1804,1941]" pageId="2" pageNumber="217">
<emphasis id="B93EEA901570FF96FF2FF8EDFF6D2B00" bold="true" box="[166,231,1804,1829]" pageId="2" pageNumber="217">Fig. 1.</emphasis>
Species of the tribe
<taxonomicName id="4C4A4D011570FF96FE30F8ECFDDB2B00" authorityName="VONDRACEK" authorityYear="1963" box="[441,593,1805,1829]" class="Insecta" family="Liviidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="tribe" tribe="Paurocephalini">Paurocephalini</taxonomicName>
in Dominican amber.
<emphasis id="B93EEA901570FF96FCB9F8EDFCC82B01" bold="true" box="[816,834,1804,1828]" pageId="2" pageNumber="217">A</emphasis>
<taxonomicName id="4C4A4D011570FF96FCD0F8ECFA0C2B00" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[857,1414,1805,1829]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="venosa" status="sp. nov.">
<emphasis id="B93EEA901570FF96FCD0F8ECFBB72B01" box="[857,1085,1805,1829]" italics="true" pageId="2" pageNumber="217">Diclidophlebia venosa</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB1570FF96FF2FF8C8FF762B65" box="[166,252,1833,1856]" pageId="2" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
, habitus of male holotype, lateral view.
<emphasis id="B93EEA901570FF96FD3BF8C8FD492B64" bold="true" box="[690,707,1833,1857]" pageId="2" pageNumber="217">B</emphasis>
Same but mid and hind legs of female paratype, ventral view.
<emphasis id="B93EEA901570FF96FF2FF8A5FF3D2B79" bold="true" box="[166,183,1860,1884]" pageId="2" pageNumber="217">C</emphasis>
<taxonomicName id="4C4A4D011570FF96FF42F8A4FD222B78" authority="BURCKHARDT&amp;DROHOJOWSKA" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[203,680,1861,1885]" class="Insecta" family="Liviidae" genus="Klyveria" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="sucina" status="sp. nov.">
<emphasis id="B93EEA901570FF96FF42F8A4FEED2B79" box="[203,359,1861,1885]" italics="true" pageId="2" pageNumber="217">Klyveria sucina</emphasis>
BURCKHARDT&amp;DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB1570FF96FD23F8A4FD722B79" box="[682,760,1861,1884]" pageId="2" pageNumber="217" rank="species">sp.nov.</taxonomicNameLabel>
, habitus of female holotype, dorsal view.
<emphasis id="B93EEA901570FF96FB0DF8A4FB1F2B78" bold="true" box="[1156,1173,1861,1885]" pageId="2" pageNumber="217">D</emphasis>
Same but ventral view.
<emphasis id="B93EEA901570FF96FF2FF880FF3D2B5C" bold="true" box="[166,183,1889,1913]" pageId="2" pageNumber="217">E</emphasis>
<taxonomicName id="4C4A4D011570FF96FF58F880FD542B5C" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[209,734,1889,1913]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="casca" status="sp. nov.">
<emphasis id="B93EEA901570FF96FF58F880FE182B5D" box="[209,402,1889,1913]" italics="true" pageId="2" pageNumber="217">Melanastera casca</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB1570FF96FD6DF880FCBF2B5D" box="[740,821,1889,1912]" pageId="2" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
, habitus of female holotype, dorsal view.
<emphasis id="B93EEA901570FF96FB55F880FB612B5C" bold="true" box="[1244,1259,1889,1913]" pageId="2" pageNumber="217">F</emphasis>
<taxonomicName id="4C4A4D011570FF96FA8FF880FDAD2BB0" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="vetus" status="sp. nov.">
<emphasis id="B93EEA901570FF96FA8FF880FF502BB1" italics="true" pageId="2" pageNumber="217">Melanastera vetus</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB1570FF96FDA5F89CFDF72BB1" box="[556,637,1917,1940]" pageId="2" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
, habitus of female holotype, dorsal view.
</paragraph>
</caption>
<caption id="DF35660A1571FF97FF27F8F2FE102BBF" ID-DOI="http://doi.org/10.5281/zenodo.15237577" ID-Zenodo-Dep="15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="3" pageNumber="218" startId="3.[174,211,1811,1835]" targetBox="[174,1421,151,1790]" targetPageId="3" targetType="figure">
<paragraph id="8BF536821571FF97FF27F8F2FE102BBF" blockId="3.[174,1421,1811,1947]" pageId="3" pageNumber="218">
<emphasis id="B93EEA901571FF97FF27F8F2FF782B0E" bold="true" box="[174,242,1811,1835]" pageId="3" pageNumber="218">Fig. 2.</emphasis>
<taxonomicName id="4C4A4D011571FF97FF76F8F2FCB92B0E" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[255,819,1811,1835]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="venosa" status="sp. nov.">
<emphasis id="B93EEA901571FF97FF76F8F2FE6E2B0E" box="[255,484,1811,1835]" italics="true" pageId="3" pageNumber="218">Diclidophlebia venosa</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB1571FF97FCB3F8F2FC032B0F" box="[826,905,1811,1834]" pageId="3" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93EEA901571FF97FC19F8F2FC282B0E" bold="true" box="[912,930,1811,1835]" pageId="3" pageNumber="218">A</emphasis>
Head, dorsal view.
<emphasis id="B93EEA901571FF97FB0AF8F2FB1E2B0E" bold="true" box="[1155,1172,1811,1835]" pageId="3" pageNumber="218">B</emphasis>
Hind leg, lateral view (entire leg, outer face; apical portion, inner face).
<emphasis id="B93EEA901571FF97FD39F8CEFD482B62" bold="true" box="[688,706,1839,1863]" pageId="3" pageNumber="218">C</emphasis>
Forewing of male holotype.
<emphasis id="B93EEA901571FF97FB8AF8CEFB9F2B62" bold="true" box="[1027,1045,1839,1863]" pageId="3" pageNumber="218">D</emphasis>
Left forewing of female paratype, with two extra veins branching from R
<subScript id="17CE34C71571FF97FDB8F8BAFDB22B4C" attach="left" box="[561,568,1883,1897]" fontSize="6" pageId="3" pageNumber="218">1</subScript>
.
<emphasis id="B93EEA901571FF97FDCDF8AAFDDF2B46" bold="true" box="[580,597,1867,1891]" pageId="3" pageNumber="218">E</emphasis>
Right forewing of the same female paratype, with only one extra vein branching from R
<subScript id="17CE34C71571FF97FF7CF896FF762BA0" attach="left" box="[245,252,1911,1925]" fontSize="6" pageId="3" pageNumber="218">1</subScript>
.
<emphasis id="B93EEA901571FF97FE80F886FE922B5A" bold="true" box="[265,280,1895,1919]" pageId="3" pageNumber="218">F</emphasis>
Male terminalia, lateral view.
<emphasis id="B93EEA901571FF97FDD5F886FDFA2B5A" bold="true" box="[604,624,1895,1919]" pageId="3" pageNumber="218">G</emphasis>
Female terminalia, lateral view.
<emphasis id="B93EEA901571FF97FC43F886FC542B5A" bold="true" box="[970,990,1895,1919]" pageId="3" pageNumber="218">H</emphasis>
Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY.
</paragraph>
</caption>
<paragraph id="8BF536821576FF90FF2FFF77FEBD2CD6" blockId="4.[166,773,150,243]" pageId="4" pageNumber="219">
<materialsCitation id="3B223CDF1576FF90FF2FFF77FF672CF7" collectionCode="GPAC" country="Dominican Republic" location="Cordillera Septentrional" pageId="4" pageNumber="219" specimenCount="1" typeStatus="paratype">
<emphasis id="B93EEA901576FF90FF2FFF77FEA22C95" bold="true" box="[166,296,150,176]" pageId="4" pageNumber="219">
<typeStatus id="54F188201576FF90FF2FFF77FEA92C95" box="[166,291,150,176]" pageId="4" pageNumber="219">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D4CFD0B1576FF90FEBCFF76FDC92C95" box="[309,579,150,177]" pageId="4" pageNumber="219">Two female specimens</specimenCount>
, samples #17(2), #17(3)
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;
<materialsCitation id="3B223CDF1576FF90FF73FF59FEB82CD6" collectionCode="GPAC" country="Dominican Republic" location="Cordillera Septentrional" pageId="4" pageNumber="219" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D4CFD0B1576FF90FF73FF59FE5D2CF4" box="[250,471,183,210]" pageId="4" pageNumber="219">one adult specimen</specimenCount>
without abdomen, sample #22 (
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)
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.
</paragraph>
<paragraph id="8BF536821576FF90FF2FFEFDFECF2D72" blockId="4.[166,773,283,343]" pageId="4" pageNumber="219">
<materialsCitation id="3B223CDF1576FF90FF2FFEFDFECB2D72" country="Dominican Republic" location="Cordillera Septentrional" pageId="4" pageNumber="219">
<emphasis id="B93EEA901576FF90FF2FFEFDFED72D10" bold="true" box="[166,349,283,310]" pageId="4" pageNumber="219">Type locality:</emphasis>
<collectingCountry id="F35D76121576FF90FEFFFEFAFDFF2D10" box="[374,629,283,309]" name="Dominican Republic" pageId="4" pageNumber="219" value="Dominican Republic">Dominican Republic</collectingCountry>
,
<location id="8E9560591576FF90FD1BFEFAFECB2D72" LSID="urn:lsid:plazi:treatment:03E387941573FF91FCF4FA10FE622873:8E9560591576FF90FD1BFEFAFECB2D72" country="Dominican Republic" name="Cordillera Septentrional" pageId="4" pageNumber="219">Cordillera Septentrional</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35065091576FF90FF2FFE9EFBE1289F" pageId="4" pageNumber="219" type="description">
<paragraph id="8BF536821576FF90FF2FFE9EFD3A2EC2" blockId="4.[166,774,383,1816]" pageId="4" pageNumber="219">
<emphasis id="B93EEA901576FF90FF2FFE9EFE1C2DBC" bold="true" box="[166,406,383,409]" pageId="4" pageNumber="219">Description: Adult:</emphasis>
Colouration. Head and thorax light brown. Antenna brown, segments 69 each with dark brown apex, segment 10 entirely brown. Mesoscutum with a median and each a submedian and sublateral longitudinal stripe on either side dark brown. Legs brown, apical metatibial spurs almost black. Forewing with light brown membrane and brown veins; apices of veins along wing margin each with conspicuous black spot; pterostigma greyish brown basally, light brown apically. Abdominal sclerites and terminalia brown, membrane greyish.
</paragraph>
<paragraph id="8BF536821576FF90FF4CFD0FFC3D2831" blockId="4.[166,774,383,1816]" lastBlockId="4.[807,1414,151,1044]" pageId="4" pageNumber="219">
Structure. Head, in lateral view, weakly inclined (&lt;45°) from longitudinal body axis (
<figureCitation id="13712A071576FF90FDE9FCEEFD332F0C" box="[608,697,783,810]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="4" pageNumber="219">
Fig. 1
<collectionCode id="ED5BAE471576FF90FD2BFCEEFD332F0C" box="[674,697,783,809]" pageId="4" pageNumber="219">A</collectionCode>
</figureCitation>
). Vertex trapezoidal, 1.82.5 times as wide as long along coronal suture; covered in granular microsculpture; curved down to genae without separation (
<figureCitation id="13712A071576FF90FD15FC92FD7F2FA8" box="[668,757,883,910]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">
Fig. 2
<collectionCode id="ED5BAE471576FF90FD68FC92FD7F2FA8" box="[737,757,883,909]" pageId="4" pageNumber="219">A</collectionCode>
</figureCitation>
). Genae evenly rounded, lacking processes; in the male the head is strongly compressed dorso-ventrally so that the genae stick out anteriorly looking like short genal processes (
<figureCitation id="13712A071576FF90FEEEFC19FE342836" box="[359,446,1016,1043]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="4" pageNumber="219">
Fig. 1
<collectionCode id="ED5BAE471576FF90FE2FFC18FE342836" box="[422,446,1017,1043]" pageId="4" pageNumber="219">A</collectionCode>
</figureCitation>
,
<figureCitation id="13712A071576FF90FE45FC18FE672836" box="[460,493,1017,1043]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">
2
<collectionCode id="ED5BAE471576FF90FE50FC18FE672836" box="[473,493,1017,1043]" pageId="4" pageNumber="219">A</collectionCode>
</figureCitation>
). Compound eyes hemispherical. Antenna 10-segmented, 1.5 times as long as head width; relative length of flagellar segments as 1.0: 0.3: 0.3: 0.3: 0.3: 0.3: 0.2: 0.2; relative lengths of antennal segment 10 and terminal setae as 1.0: 1.2: 1.1. Pronotum transversely ribbon-shaped, 0.8 times as long as vertex along medial longitudinal body axis. Metapostnotum with low, laterally compressed tooth. Pro- and mesotibiae slender, not widening to apex (
<figureCitation id="13712A071576FF90FF67FAC5FEE4291A" box="[238,366,1316,1343]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="4" pageNumber="219">
Fig. 1
<collectionCode id="ED5BAE471576FF90FEB9FAC4FEC2291A" box="[304,328,1317,1343]" pageId="4" pageNumber="219">A</collectionCode>
,
<collectionCode id="ED5BAE471576FF90FED2FAC4FEE4291A" box="[347,366,1317,1343]" pageId="4" pageNumber="219">B</collectionCode>
</figureCitation>
,
<figureCitation id="13712A071576FF90FEF6FAC4FE15291A" box="[383,415,1317,1343]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">
2
<collectionCode id="ED5BAE471576FF90FE04FAC4FE15291A" box="[397,415,1317,1343]" pageId="4" pageNumber="219">B</collectionCode>
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). Metacoxa with long tubular, apically blunt meracanthus (
<figureCitation id="13712A071576FF90FE7AFAA7FDC12945" box="[499,587,1350,1376]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="4" pageNumber="219">
Fig. 1
<collectionCode id="ED5BAE471576FF90FDBFFAA7FDC12945" box="[566,587,1350,1376]" pageId="4" pageNumber="219">B</collectionCode>
</figureCitation>
,
<figureCitation id="13712A071576FF90FDD7FAA7FDF42945" box="[606,638,1350,1376]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">
2
<collectionCode id="ED5BAE471576FF90FDE5FAA7FDF42945" box="[620,638,1350,1376]" pageId="4" pageNumber="219">B</collectionCode>
</figureCitation>
). Metatibia longer than metafemur, slightly widened apically; bearing (23)+(45) indistinctly grouped apical sclerotised spurs that are not on raised processes, without weakly sclerotised bristles (
<figureCitation id="13712A071576FF90FE6CFA2AFDB129C0" box="[485,571,1483,1510]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="4" pageNumber="219">
Fig. 1
<collectionCode id="ED5BAE471576FF90FDAFFA2AFDB129C0" box="[550,571,1483,1509]" pageId="4" pageNumber="219">B</collectionCode>
</figureCitation>
,
<figureCitation id="13712A071576FF90FDC5FA2AFDE629C0" box="[588,620,1483,1509]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">
2
<collectionCode id="ED5BAE471576FF90FDD3FA2AFDE629C0" box="[602,620,1483,1509]" pageId="4" pageNumber="219">B</collectionCode>
</figureCitation>
). Both metatarsal segments relatively short, subequal in length. Forewing (
<figureCitation id="13712A071576FF90FEABF9EFFEF32A0D" box="[290,377,1550,1576]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="4" pageNumber="219">
Fig. 1
<collectionCode id="ED5BAE471576FF90FEEBF9EFFEF32A0D" box="[354,377,1550,1576]" pageId="4" pageNumber="219">A</collectionCode>
</figureCitation>
,
<figureCitation id="13712A071576FF90FE00F9EFFE432A0D" box="[393,457,1550,1576]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">
2
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<collectionCode id="ED5BAE471576FF90FE31F9EFFE432A0D" box="[440,457,1550,1576]" pageId="4" pageNumber="219">E</collectionCode>
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) oblong-oval; widest in the middle, 2.32.4 times as long as wide, membranous; apex narrowly rounded; vein C+Sc weakly curved, appearing to be slender and well delimited to cell but not clearly visible; costal break developed, close to apex of vein R
<subScript id="17CE34C71576FF90FEC1F926FEDA2AF0" attach="left" box="[328,336,1735,1749]" fontSize="6" pageId="4" pageNumber="219">1</subScript>
; pterostigma wide, coriaceous basally, membranous apically; nodal line not developed; vein R slightly shorter than M+Cu; vein R
<subScript id="17CE34C71576FF90FD19F8EBFD122B3D" attach="left" box="[656,664,1802,1816]" fontSize="6" pageId="4" pageNumber="219">1</subScript>
with one subapical branch, in
<specimenCount id="9D4CFD0B1576FF90FB9DFF76FB1B2C94" box="[1044,1169,151,177]" count="1" pageId="4" pageNumber="219" type="female">one female</specimenCount>
the left forewing has two branches and in a specimen without abdomen both forewings have two branches but arranged asymmetrically; vein Rs relatively straight in basal four fifths, weakly curved towards costal margin in apical fifth; vein M distinctly longer than M
<subScript id="17CE34C71576FF90FB11FEB1FB3A2D7B" attach="left" box="[1176,1200,336,350]" fontSize="6" pageId="4" pageNumber="219">1+2</subScript>
; vein M
<subScript id="17CE34C71576FF90FA9BFEB1FAA12D7B" attach="left" box="[1298,1323,336,350]" fontSize="6" pageId="4" pageNumber="219">1+2</subScript>
weakly curved; vein M
<subScript id="17CE34C71576FF90FC52FE90FC7F2D5A" attach="left" box="[987,1013,369,383]" fontSize="6" pageId="4" pageNumber="219">3+4</subScript>
almost straight; vein Cu
<subScript id="17CE34C71576FF90FA9EFE90FAAC2D5A" attach="left" box="[1303,1318,369,383]" fontSize="6" pageId="4" pageNumber="219">1a</subScript>
straight in basal quarter, strongly curved in apical third; vein Cu
<subScript id="17CE34C71576FF90FCCEFE55FCDD2DE7" attach="left" box="[839,855,436,450]" fontSize="6" pageId="4" pageNumber="219">1b</subScript>
short, straight; cell cu
<subScript id="17CE34C71576FF90FBD1FE55FBEA2DE7" attach="left" box="[1112,1120,436,450]" fontSize="6" pageId="4" pageNumber="219">2</subScript>
long; anal break adjacent to apex of vein Cu
<subScript id="17CE34C71576FF90FB89FE34FB9A2DC6" attach="left" box="[1024,1040,469,483]" fontSize="6" pageId="4" pageNumber="219">1b</subScript>
; surface spinules fine, arranged in transverse rows, visible in cell cu
<subScript id="17CE34C71576FF90FB50FE17FB6B2E21" attach="left" box="[1241,1249,502,516]" fontSize="6" pageId="4" pageNumber="219">2</subScript>
and in apical quarter of wing. Hindwing three quarter of forewing length; with 68 relatively evenly spaced costal setae distal to costal break. Male terminalia as in
<figureCitation id="13712A071576FF90FAA0FDA9FAF52E47" box="[1321,1407,584,610]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="4" pageNumber="219">Fig. 1A</figureCitation>
,
<figureCitation id="13712A071576FF90FCAEFD88FCC92EA6" box="[807,835,617,643]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">2F</figureCitation>
; proctiger tubular, weakly convex posteriorly; subgenital plate elongate, beset with a few moderately long setae in apical two thirds, ventrally; paramere slightly shorter than proctiger, lamellar, strongly narrowed in apical quarter into digitiform process. Female terminalia, in lateral view, cuneate (
<figureCitation id="13712A071576FF90FA97FCF1FAFF2F0F" box="[1310,1397,784,810]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">
Fig. 2
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</figureCitation>
); proctiger almost straight dorsally except for a shallow indentation in the middle, pointed apically, with several moderately long hairs in the middle (
<figureCitation id="13712A071576FF90FAA9FC95FAFF2FAB" box="[1312,1397,884,910]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">
Fig. 2
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</figureCitation>
); subgenital plate slightly shorter than proctiger bearing moderately long setae in the middle; ventral margin hardly curved; irregularly narrowing to pointed apex (
<figureCitation id="13712A071576FF90FCA6FC18FC2C2831" box="[815,934,1017,1044]" captionStart="Fig" captionStartId="3.[174,211,1811,1835]" captionTargetBox="[174,1421,151,1790]" captionTargetId="figure-8@3.[174,1421,151,1790]" captionTargetPageId="3" captionText="Fig. 2. Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view (entire leg, outer face; apical portion, inner face). C Forewing of male holotype. D Left forewing of female paratype, with two extra veins branching from R 1. E Right forewing of the same female paratype, with only one extra vein branching from R 1. F Male terminalia, lateral view. G Female terminalia, lateral view. H Same but ventral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237577" httpUri="https://zenodo.org/record/15237577/files/figure.png" pageId="4" pageNumber="219">
Fig. 2
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,
<collectionCode id="ED5BAE471576FF90FC1BFC1BFC2C2831" box="[914,934,1018,1044]" pageId="4" pageNumber="219">H</collectionCode>
</figureCitation>
).
</paragraph>
<paragraph id="8BF536821576FF90FCAEFBDDFBE1289F" blockId="4.[807,1414,1083,1210]" pageId="4" pageNumber="219">
<emphasis id="B93EEA901576FF90FCAEFBDDFBC42870" bold="true" box="[807,1102,1083,1110]" pageId="4" pageNumber="219">Measurements (in mm):</emphasis>
Head width 0.66; antenna length 0.8; forewing length 1.71.9; metatibia length 0.5; male proctiger length 0.29; paramere length 0.18; female proctiger length 0.43.
</paragraph>
</subSubSection>
<subSubSection id="C35065091576FF91FCAEFB03FE622873" lastPageId="5" lastPageNumber="220" pageId="4" pageNumber="219" type="discussion">
<paragraph id="8BF536821576FF91FCAEFB03FE622873" blockId="4.[807,1414,1250,1816]" lastBlockId="5.[174,781,151,1110]" lastPageId="5" lastPageNumber="220" pageId="4" pageNumber="219">
<emphasis id="B93EEA901576FF90FCAEFB03FC3828D9" bold="true" box="[807,946,1250,1276]" pageId="4" pageNumber="219">Comments:</emphasis>
<taxonomicName id="4C4A4D011576FF90FC34FB03FB4E28D9" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[957,1220,1250,1276]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="venosa">
<emphasis id="B93EEA901576FF90FC34FB03FB4E28D9" box="[957,1220,1250,1276]" italics="true" pageId="4" pageNumber="219">Diclidophlebia venosa</emphasis>
</taxonomicName>
fits the concept of
<taxonomicName id="4C4A4D011576FF90FCC0FAE5FC7F293B" authorityName="CRAWFORD" authorityYear="1920" box="[841,1013,1284,1310]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEA901576FF90FCC0FAE5FC7F293B" box="[841,1013,1284,1310]" italics="true" pageId="4" pageNumber="219">Diclidophlebia</emphasis>
</taxonomicName>
redefined by
<bibRefCitation id="EFDB4B731576FF90FB28FAE5FCE5291A" author="BURCKHARDT, D. &amp; SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEI- ROZ, D. L. &amp; ALENE, D. C. &amp; CHO, G. &amp; PERCY, D. M." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="4" pageNumber="219" pagination="387 - 421" part="201" publicationUrl="10.1093/zoolinnean/zlad128" refId="ref6525" refString="BURCKHARDT, D., SERBINA, L. S., MALENOVSKY, I., QUEI- ROZ, D. L., ALENE, D. C., CHO, G. &amp; PERCY, D. M. (2023): Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data. - Zoological Journal of the Linnean Society, 2023, XX, 1 - 35; advance access publication 9 October 2023. Printed version: (2024), 201: 387-421. DOI: 10.1093/zoolinnean/zlad128" title="Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data" type="journal article" year="2023">BURCKHARDT et al. (2023)</bibRefCitation>
by the pterostigma of the forewing, which is leathery in the proximal half and membranous in the distal half (a synapomorphy of the genus), and the presence of well-sclerotised apical spurs on the metatibia arranged in two groups. Another diagnostic character of
<taxonomicName id="4C4A4D011576FF90FC6CFA2DFC3E2A22" authority="sensu BURCKHARDT et al. (2023)" authorityName="BURCKHARDT" authorityYear="2023" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEA901576FF90FC6CFA2DFB1B29C3" box="[997,1169,1484,1510]" italics="true" pageId="4" pageNumber="219">Diclidophlebia</emphasis>
sensu
<bibRefCitation id="EFDB4B731576FF90FB6EFA2DFC3E2A22" author="BURCKHARDT, D. &amp; SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEI- ROZ, D. L. &amp; ALENE, D. C. &amp; CHO, G. &amp; PERCY, D. M." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="4" pageNumber="219" pagination="387 - 421" part="201" publicationUrl="10.1093/zoolinnean/zlad128" refId="ref6525" refString="BURCKHARDT, D., SERBINA, L. S., MALENOVSKY, I., QUEI- ROZ, D. L., ALENE, D. C., CHO, G. &amp; PERCY, D. M. (2023): Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data. - Zoological Journal of the Linnean Society, 2023, XX, 1 - 35; advance access publication 9 October 2023. Printed version: (2024), 201: 387-421. DOI: 10.1093/zoolinnean/zlad128" title="Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data" type="journal article" year="2023">BURCKHARDT et al. (2023)</bibRefCitation>
</taxonomicName>
, the broadened C+Sc vein of the forewing that is not clearly demarcated from the neighbouring cell, is not clearly visible in the specimens examined.
<taxonomicName id="4C4A4D011576FF90FC27F9B0FB322A4E" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[942,1208,1617,1643]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="venosa">
<emphasis id="B93EEA901576FF90FC27F9B0FB322A4E" box="[942,1208,1617,1643]" italics="true" pageId="4" pageNumber="219">Diclidophlebia venosa</emphasis>
</taxonomicName>
differs from all seven extant
<taxonomicName id="4C4A4D011576FF90FC4EF993FB592AA9" box="[967,1235,1650,1676]" pageId="4" pageNumber="219">
<emphasis id="B93EEA901576FF90FC4EF993FBF92AA9" box="[967,1139,1650,1676]" italics="true" pageId="4" pageNumber="219">Diclidophlebia</emphasis>
species
</taxonomicName>
recognised by
<bibRefCitation id="EFDB4B731576FF90FCAEF975FBDB2A8B" author="BURCKHARDT, D. &amp; SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEI- ROZ, D. L. &amp; ALENE, D. C. &amp; CHO, G. &amp; PERCY, D. M." box="[807,1105,1683,1710]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="4" pageNumber="219" pagination="387 - 421" part="201" publicationUrl="10.1093/zoolinnean/zlad128" refId="ref6525" refString="BURCKHARDT, D., SERBINA, L. S., MALENOVSKY, I., QUEI- ROZ, D. L., ALENE, D. C., CHO, G. &amp; PERCY, D. M. (2023): Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data. - Zoological Journal of the Linnean Society, 2023, XX, 1 - 35; advance access publication 9 October 2023. Printed version: (2024), 201: 387-421. DOI: 10.1093/zoolinnean/zlad128" title="Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data" type="journal article" year="2023">BURCKHARDT et al. (2023)</bibRefCitation>
in the presence of the subapically forked vein R
<subScript id="17CE34C71576FF90FBA1F929FBBA2AF3" attach="left" box="[1064,1072,1736,1750]" fontSize="6" pageId="4" pageNumber="219">1</subScript>
, which is unusual in psyllids and, as far as known, unique in
<taxonomicName id="4C4A4D011576FF90FB2EF936FA812AD4" authorityName="LOW" authorityYear="1879" box="[1191,1291,1751,1777]" class="Insecta" family="Liviidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="225" phylum="Arthropoda" rank="subFamily" subFamily="Liviinae">Liviinae</taxonomicName>
. In two of the
<specimenCount id="9D4CFD0B1576FF90FCDFF919FB832B37" box="[854,1033,1784,1810]" count="4" pageId="4" pageNumber="219" type="generic">four specimens</specimenCount>
at hand a second fork on R
<subScript id="17CE34C71576FF90FAD6F8EBFAED2B3D" attach="left" box="[1375,1383,1802,1816]" fontSize="6" pageId="4" pageNumber="219">1</subScript>
is developed on one or on both wings, but then asymmetically. The asymmetrical nature of this extra-vein suggests that this constitutes an abnormal feature. Specimens with abnormal venation are sometimes also found in extant psyllids.
<taxonomicName id="4C4A4D011577FF91FD81FEFDFC872D13" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[520,781,284,310]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="venosa">
<emphasis id="B93EEA901577FF91FD81FEFDFC872D13" box="[520,781,284,310]" italics="true" pageId="5" pageNumber="220">Diclidophlebia venosa</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C4A4D011577FF91FEBBFEDFFD242D7D" authority="(KLYVER, 1930)" baseAuthorityName="KLYVER" baseAuthorityYear="1930" box="[306,686,317,344]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="fremontiae">
<emphasis id="B93EEA901577FF91FEBBFEDFFE562D72" box="[306,476,317,343]" italics="true" pageId="5" pageNumber="220">D. fremontiae</emphasis>
(
<bibRefCitation id="EFDB4B731577FF91FE70FEDFFD2F2D72" author="KLYVER, F. D." box="[505,677,317,344]" journalOrPublisher="Canadian Entomologist" pageId="5" pageNumber="220" pagination="167 - 175" part="62" publicationUrl="10.4039/Ent62167-8" refId="ref7145" refString="KLYVER, F. D. (1930): Notes on the Chermidae Part 1 (Hemiptera: Homoptera). - Canadian Entomologist, 62: 167-175. DOI: 10.4039/Ent62167-8." title="Notes on the Chermidae Part 1 (Hemiptera: Homoptera)" type="journal article" year="1930">KLYVER, 1930</bibRefCitation>
)
</taxonomicName>
in the unpigmented, elongate oval forewings with the vein M longer than M
<subScript id="17CE34C71577FF91FEF3FE73FE192D85" attach="right" box="[378,403,402,416]" fontSize="6" pageId="5" pageNumber="220">1+2</subScript>
, and with the long cell cu
<subScript id="17CE34C71577FF91FD59FE73FD522D85" attach="right" box="[720,728,402,416]" fontSize="6" pageId="5" pageNumber="220">1</subScript>
, the apically barely expanded metatibia with apical spurs not located on processes, and its occurrence in the New World. These characters indicate a close phylogenetic relationship between the two species.
<taxonomicName id="4C4A4D011577FF91FD53FDE7FE002E65" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="venosa" status="sp. nov.">
<emphasis id="B93EEA901577FF91FD53FDE7FE002E65" italics="true" pageId="5" pageNumber="220">Diclidophlebia venosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57EB1577FF91FE1AFDC6FE662E64" box="[403,492,551,577]" pageId="5" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C4A4D011577FF91FD05FDC6FF532E47" baseAuthorityName="KLYVER" baseAuthorityYear="1930" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="fremontiae">
<emphasis id="B93EEA901577FF91FD05FDC6FF532E47" italics="true" pageId="5" pageNumber="220">D. fremontiae</emphasis>
</taxonomicName>
in the presence of black spots at the apices of the veins along the forewing margin, the greater number of metatibial spurs (1+(23) in the latter), the longer male subgenital plate, the apically stronger and more abruptly narrowed paramere and the longer female terminalia without the upturned apex of the proctiger. The six Old World species have either pigmented forewings (
<taxonomicName id="4C4A4D011577FF91FEA6FCD3FDF42F6E" authority="VONDRACEK, 1964" authorityName="VONDRACEK" authorityYear="1964" box="[303,638,817,846]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="eastopi">
<emphasis id="B93EEA901577FF91FEA6FCD3FE2E2F6E" box="[303,420,818,843]" italics="true" pageId="5" pageNumber="220">D. eastopi</emphasis>
<bibRefCitation id="EFDB4B731577FF91FE22FCD2FDF42F6E" author="VONDRACEK, K." box="[427,638,817,846]" journalOrPublisher="Annals and Magazine of Natural History" pageId="5" pageNumber="220" pagination="289 - 295" part="13" refId="ref7910" refString="VONDRACEK, K. (1964): Diclidophlebia eastopi sp. n., a remarkable psyllid from West Africa. - Annals and Magazine of Natural History, 13 (6): 289-295." title="Diclidophlebia eastopi sp. n., a remarkable psyllid from West Africa" type="journal article" year="1964">VONDRÁČEK, 1964</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4A4D011577FF91FD02FCD3FE622F48" authority="(LI &amp; YANG, 1991)" baseAuthorityName="LI &amp; YANG" baseAuthorityYear="1991" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="excetrodendri">
<emphasis id="B93EEA901577FF91FD02FCD3FF702F49" italics="true" pageId="5" pageNumber="220">D. excetrodendri</emphasis>
(
<bibRefCitation id="EFDB4B731577FF91FE86FCB2FE572F48" author="Li, F. &amp; YANG, C. K." box="[271,477,851,877]" journalOrPublisher="Entomotaxonomia" pageId="5" pageNumber="220" pagination="11 - 19" part="13" refId="ref7227" refString="Li, F. &amp; YANG, C. K. (1991): One new genus, three new species and a known species of psyllids (Homoptera: Psylloidea) from Guanxi, China. - Entomotaxonomia, 13: 11-19." title="One new genus, three new species and a known species of psyllids (Homoptera: Psylloidea) from Guanxi, China" type="journal article" year="1991">
<collectionCode id="ED5BAE471577FF91FE86FCB2FEA22F48" box="[271,296,851,877]" pageId="5" pageNumber="220">LI</collectionCode>
&amp;
<collectionCode id="ED5BAE471577FF91FEDDFCB2FE102F48" box="[340,410,851,877]" pageId="5" pageNumber="220">YANG</collectionCode>
, 1991
</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C4A4D011577FF91FE70FCB2FF6F2FAB" authority="OSISANYA, 1969" authorityName="OSISANYA" authorityYear="1969" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="harrisoni">
<emphasis id="B93EEA901577FF91FE70FCB2FD012F49" box="[505,651,850,876]" italics="true" pageId="5" pageNumber="220">D. harrisoni</emphasis>
<bibRefCitation id="EFDB4B731577FF91FD1EFCB2FF6F2FAB" author="OSISANYA, E. O." journalOrPublisher="Journal of Natural History" pageId="5" pageNumber="220" pagination="71 - 77" part="3" publicationUrl="10.1080/00222936900770081" refId="ref7421" refString="OSISANYA, E. O. (1969): A new species of Diclidophlebia (Homoptera: Psyllidae) from Nigeria. - Journal of Natural History, 3: 71-77. DOI: 10.1080/00222936900770081." title="A new species of Diclidophlebia (Homoptera: Psyllidae) from Nigeria" type="journal article" year="1969">OSISANYA, 1969</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C4A4D011577FF91FEAAFC94FD142FAB" authority="(CRAWFORD, 1919))" authorityName="CRAWFORD" authorityYear="1919" baseAuthorityName="CRAWFORD" baseAuthorityYear="1919" box="[291,670,884,910]" class="Insecta" family="Liviidae" genus="Heteroneura" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="oceanica">
<emphasis id="B93EEA901577FF91FEAAFC94FE3B2FAB" box="[291,433,885,910]" italics="true" pageId="5" pageNumber="220">D. oceanica</emphasis>
(
<bibRefCitation id="EFDB4B731577FF91FE4DFC95FD062FAB" author="CRAWFORD, D. L." box="[452,652,884,910]" journalOrPublisher="Philippine Journal of Science" pageId="5" pageNumber="220" pagination="139 - 207" part="15" publicationUrl="10.5962/bhl.part.11765" refId="ref6674" refString="CRAWFORD, D. L. (1919): The jumping plant lice of the Palaeotropics and the South Pacific Islands - Family Psyllidae, or Chermidae, Homoptera. - Philippine Journal of Science, 15: 139-207. DOI: 10.5962/bhl.part.11765." title="The jumping plant lice of the Palaeotropics and the South Pacific Islands - Family Psyllidae, or Chermidae, Homoptera" type="journal article" year="1919">CRAWFORD, 1919</bibRefCitation>
))
</taxonomicName>
, the vein M
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distinctly longer than M (
<emphasis id="B93EEA901577FF91FD98FC77FF732FF4" italics="true" pageId="5" pageNumber="220">
<taxonomicName id="4C4A4D011577FF91FD98FC77FD002F8A" authorityName="VONDRACEK" authorityYear="1964" box="[529,650,918,943]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="eastopi">D. eastopi</taxonomicName>
,
<taxonomicName id="4C4A4D011577FF91FD1EFC77FDAF2FF5" authority="BURCKHARDT et al., 2006" authorityName="BURCKHARDT" authorityYear="2006" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="leptonychiae">
D. leptonychiae
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and
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<emphasis id="B93EEA901577FF91FDD5FC56FD6E2FF5" box="[604,740,951,976]" italics="true" pageId="5" pageNumber="220">D. oceanica</emphasis>
</taxonomicName>
) or have an apically clearly widened metatibia with apical spurs on processes (
<emphasis id="B93EEA901577FF91FE4FFC1BFE502811" italics="true" pageId="5" pageNumber="220">
<taxonomicName id="4C4A4D011577FF91FE4FFC1BFDCB2836" authorityName="VONDRACEK" authorityYear="1964" box="[454,577,1018,1043]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="eastopi">D. eastopi</taxonomicName>
,
<taxonomicName id="4C4A4D011577FF91FDC6FC1BFD692836" authorityName="OSISANYA" authorityYear="1969" box="[591,739,1017,1043]" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="harrisoni">D. harrisoni</taxonomicName>
,
<taxonomicName id="4C4A4D011577FF91FD78FC1BFECE2811" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="leptonychiae">D. leptonychiae</taxonomicName>
,
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</emphasis>
, and
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in
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et al. 1998
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).
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Genus
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species:
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Genus
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SERBINA, MALENOVSKÝ,QU EIROZ &amp; BURCKHARDT
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in
<bibRefCitation id="EFDB4B731574FF92FBB0FB41FADE289F" author="BURCKHARDT, D. &amp; SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEI- ROZ, D. L. &amp; ALENE, D. C. &amp; CHO, G. &amp; PERCY, D. M." box="[1081,1364,1184,1210]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="6" pageNumber="221" pagination="387 - 421" part="201" publicationUrl="10.1093/zoolinnean/zlad128" refId="ref6525" refString="BURCKHARDT, D., SERBINA, L. S., MALENOVSKY, I., QUEI- ROZ, D. L., ALENE, D. C., CHO, G. &amp; PERCY, D. M. (2023): Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data. - Zoological Journal of the Linnean Society, 2023, XX, 1 - 35; advance access publication 9 October 2023. Printed version: (2024), 201: 387-421. DOI: 10.1093/zoolinnean/zlad128" title="Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data" type="journal article" year="2023">BURCKHARDT et al. 2023</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35065091574FF92FCAEFB02FC23293B" pageId="6" pageNumber="221" type="type_taxon">
<paragraph id="8BF536821574FF92FCAEFB02FC23293B" blockId="6.[807,1414,1250,1311]" pageId="6" pageNumber="221">
<emphasis id="B93EEA901574FF92FCAEFB02FC4628D9" bold="true" box="[807,972,1250,1277]" pageId="6" pageNumber="221">
<typeStatus id="54F188201574FF92FCAEFB02FCE828D8" box="[807,866,1251,1277]" pageId="6" pageNumber="221">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C4A4D011574FF92FC53FB03FC28293A" authority="BURCKHARDT et al., 2005" authorityName="BURCKHARDT" authorityYear="2005" class="Insecta" family="Liviidae" genus="Diclidophlebia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="lucens">
<emphasis id="B93EEA901574FF92FC53FB03FB5328D9" box="[986,1241,1250,1276]" italics="true" pageId="6" pageNumber="221">Diclidophlebia lucens</emphasis>
<bibRefCitation id="EFDB4B731574FF92FB6EFB02FC28293A" author="BURCKHARDT, D. &amp; HANSON, P. &amp; MADRIGAL, L." journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="6" pageNumber="221" pagination="741 - 749" part="107" refId="ref6241" refString="BURCKHARDT, D., HANSON, P. &amp; MADRIGAL, L. (2005): Diclidophlebia lucens, n. sp. (Hemiptera: Psyllidae) from Costa Rica, a potential control agent of Miconia calvescens (Melastomataceae) in Hawaii. - Proceedings of the Entomological Society of Washington, 107 (4): 741-749." title="Diclidophlebia lucens, n. sp. (Hemiptera: Psyllidae) from Costa Rica, a potential control agent of Miconia calvescens (Melastomataceae) in Hawaii" type="journal article" year="2005">BURCKHARDT et al., 2005</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,282 @@
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<mods:title id="E5811E482D821954DBBD3ACF5D062AF5">First record of jumping plant lice of the family Liviidae (Hemiptera, Sternorrhyncha, Psylloidea) from Dominican amber</mods:title>
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<paragraph id="8BF536821574FF92FCBBFAA7FB0B29A7" blockId="6.[818,1402,1350,1444]" pageId="6" pageNumber="221">
<taxonomicName id="4C4A4D011574FF92FCBBFAA7FAF02945" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[818,1402,1350,1376]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="casca" status="sp. nov.">
<emphasis id="B93EEA901574FF92FCBBFAA7FB822945" box="[818,1032,1350,1376]" italics="true" pageId="6" pageNumber="221">Melanastera casca</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB1574FF92FBA5FA89FB0B29A7" box="[1068,1153,1384,1410]" pageId="6" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BF536821574FF92FBA9FA68FB072986" blockId="6.[818,1402,1350,1444]" box="[1056,1165,1417,1444]" pageId="6" pageNumber="221">
<figureCitation id="13712A071574FF92FBA9FA68FBFA2986" box="[1056,1136,1417,1444]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="6" pageNumber="221">Fig. 1E</figureCitation>
,
<figureCitation id="13712A071574FF92FBF6FA68FB072986" box="[1151,1165,1417,1443]" captionStart="Fig" captionStartId="7.[174,211,1011,1035]" captionTargetBox="[174,1421,151,985]" captionTargetId="figure-355@7.[174,1421,151,985]" captionTargetPageId="7" captionText="Fig. 4. Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind legs, lateral view. C Forewing. D Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237581" httpUri="https://zenodo.org/record/15237581/files/figure.png" pageId="6" pageNumber="221">4</figureCitation>
</paragraph>
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<subSubSection id="C35065091574FF92FCAEFA2AFBE42A2D" pageId="6" pageNumber="221" type="description">
<paragraph id="8BF536821574FF92FCAEFA2AFBE42A2D" blockId="6.[807,1414,1483,1544]" pageId="6" pageNumber="221">
<emphasis id="B93EEA901574FF92FCAEFA2AFCF829C0" bold="true" box="[807,882,1483,1509]" pageId="6" pageNumber="221">LSID:</emphasis>
urn:lsid:zoobank.org:act:329F8C6-4FAE-43BE-9547-A6B86A765CAC
</paragraph>
</subSubSection>
<subSubSection id="C35065091574FF92FCAEF9D1FC372AA9" pageId="6" pageNumber="221" type="etymology">
<paragraph id="8BF536821574FF92FCAEF9D1FC372AA9" blockId="6.[807,1414,1584,1676]" pageId="6" pageNumber="221">
<emphasis id="B93EEA901574FF92FCAEF9D1FC392A6F" bold="true" box="[807,947,1584,1610]" pageId="6" pageNumber="221">Etymology:</emphasis>
From the Latin adjective cascus (1
<superScript id="7C3F9BCA1574FF92FADAF9D1FAD42A1B" attach="none" box="[1363,1374,1584,1598]" fontSize="6" pageId="6" pageNumber="221">st</superScript>
/2
<superScript id="7C3F9BCA1574FF92FAFCF9D1FA0C2A1B" attach="none" box="[1397,1414,1584,1598]" fontSize="6" pageId="6" pageNumber="221">nd</superScript>
declension) = ancient, old, archaic, referring to the age of the fossils.
</paragraph>
</subSubSection>
<subSubSection id="C35065091574FF92FCAEF954FC4C2B16" pageId="6" pageNumber="221" type="materials_examined">
<paragraph id="8BF536821574FF92FCAEF954FAD62AEA" blockId="6.[807,1372,1717,1743]" box="[807,1372,1717,1743]" pageId="6" pageNumber="221">
<materialsCitation id="3B223CDF1574FF92FCAEF954FADD2AEA" box="[807,1367,1717,1743]" collectionCode="GPAC" country="Dominican Republic" location="Cordillera Septentrional" pageId="6" pageNumber="221" typeStatus="holotype">
<emphasis id="B93EEA901574FF92FCAEF954FC142AEA" bold="true" box="[807,926,1717,1743]" pageId="6" pageNumber="221">
<typeStatus id="54F188201574FF92FCAEF954FC132AEA" box="[807,921,1717,1743]" pageId="6" pageNumber="221">Holotype</typeStatus>
:
</emphasis>
Female specimen, sample #37 (
<collectionCode id="ED5BAE471574FF92FA8DF954FADA2AEA" box="[1284,1360,1717,1743]" pageId="6" pageNumber="221">GPAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF536821574FF92FCAEF919FC4C2B16" blockId="6.[807,1414,1784,1843]" pageId="6" pageNumber="221">
<materialsCitation id="3B223CDF1574FF92FCAEF919FC482B16" country="Dominican Republic" pageId="6" pageNumber="221" typeStatus="Type">
<emphasis id="B93EEA901574FF92FCAEF919FC572B37" bold="true" box="[807,989,1784,1810]" pageId="6" pageNumber="221">
<typeStatus id="54F188201574FF92FCAEF919FCE82B37" box="[807,866,1784,1810]" pageId="6" pageNumber="221">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F35D76121574FF92FC7EF919FB7C2B37" box="[1015,1270,1784,1810]" name="Dominican Republic" pageId="6" pageNumber="221" value="Dominican Republic">Dominican Republic</collectingCountry>
, Cordillera Septentrional
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF35660A1575FF93FF27FC12FBD12803" ID-DOI="http://doi.org/10.5281/zenodo.15237581" ID-Zenodo-Dep="15237581" httpUri="https://zenodo.org/record/15237581/files/figure.png" pageId="7" pageNumber="222" startId="7.[174,211,1011,1035]" targetBox="[174,1421,151,985]" targetPageId="7" targetType="figure">
<paragraph id="8BF536821575FF93FF27FC12FBD12803" blockId="7.[174,1422,1011,1063]" pageId="7" pageNumber="222">
<emphasis id="B93EEA901575FF93FF27FC12FF79282E" bold="true" box="[174,243,1011,1035]" pageId="7" pageNumber="222">Fig. 4.</emphasis>
<taxonomicName id="4C4A4D011575FF93FF76FC12FC9E282E" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[255,788,1011,1035]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="222" phylum="Arthropoda" rank="species" species="casca" status="sp. nov.">
<emphasis id="B93EEA901575FF93FF76FC12FE48282E" box="[255,450,1011,1035]" italics="true" pageId="7" pageNumber="222">Melanastera casca</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB1575FF93FC95FC12FCE7282F" box="[796,877,1011,1034]" pageId="7" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93EEA901575FF93FCFCFC12FC0D282E" bold="true" box="[885,903,1011,1035]" pageId="7" pageNumber="222">A</emphasis>
Head, dorsal view.
<emphasis id="B93EEA901575FF93FBE7FC12FBF5282E" bold="true" box="[1134,1151,1011,1035]" pageId="7" pageNumber="222">B</emphasis>
Hind legs, lateral view.
<emphasis id="B93EEA901575FF93FF27FBEEFF4A2802" bold="true" box="[174,192,1039,1063]" pageId="7" pageNumber="222">C</emphasis>
Forewing.
<emphasis id="B93EEA901575FF93FECEFBEEFED32802" bold="true" box="[327,345,1039,1063]" pageId="7" pageNumber="222">D</emphasis>
Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY.
</paragraph>
</caption>
<subSubSection id="C35065091575FF93FF27FB9BFB292B0A" pageId="7" pageNumber="222" type="description">
<paragraph id="8BF536821575FF93FF27FB9BFEA628DD" blockId="7.[174,781,1146,1839]" pageId="7" pageNumber="222">
<emphasis id="B93EEA901575FF93FF27FB9BFE1128B1" bold="true" box="[174,411,1146,1172]" pageId="7" pageNumber="222">Description. Adult:</emphasis>
Colouration. Brown. Antennal segments 47 dark apically, segments 810 entirely dark. Forewing with dark veins and colourless membrane.
</paragraph>
<paragraph id="8BF536821575FF93FF45FAE1FAA92A8F" blockId="7.[174,781,1146,1839]" lastBlockId="7.[814,1422,1146,1706]" pageId="7" pageNumber="222">
Structure. Head, in dorsal view, wider than pronotum, slightly narrower than thorax, moderately transverse (
<figureCitation id="13712A071575FF93FEBDFAA3FE012978" box="[308,395,1346,1373]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="7" pageNumber="222">
Fig. 1
<collectionCode id="ED5BAE471575FF93FEFEFAA2FE012978" box="[375,395,1347,1373]" pageId="7" pageNumber="222">E</collectionCode>
</figureCitation>
,
<figureCitation id="13712A071575FF93FE16FAA2FE352978" box="[415,447,1347,1373]" captionStart="Fig" captionStartId="7.[174,211,1011,1035]" captionTargetBox="[174,1421,151,985]" captionTargetId="figure-355@7.[174,1421,151,985]" captionTargetPageId="7" captionText="Fig. 4. Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind legs, lateral view. C Forewing. D Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237581" httpUri="https://zenodo.org/record/15237581/files/figure.png" pageId="7" pageNumber="222">
4
<collectionCode id="ED5BAE471575FF93FE25FAA2FE352978" box="[428,447,1347,1373]" pageId="7" pageNumber="222">A</collectionCode>
</figureCitation>
). Vertex subrectangular, 2.1 times as wide as long; passing smoothly into genae anteriorly; coronal suture not visible basally; median ocellus visible in perpendicular view to vertex; compound eyes, in dorsal view, hemispherical, appressed to head. Antenna (
<figureCitation id="13712A071575FF93FE04FA08FE6E2A26" box="[397,484,1513,1540]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="7" pageNumber="222">
Fig. 1
<collectionCode id="ED5BAE471575FF93FE59FA08FE6E2A26" box="[464,484,1513,1539]" pageId="7" pageNumber="222">E</collectionCode>
</figureCitation>
) probably 10-segmented, but subdivisions not clear; slightly longer than head width. Metacoxa (
<figureCitation id="13712A071575FF93FEF1F9CDFE412A63" box="[376,459,1580,1606]" captionStart="Fig" captionStartId="7.[174,211,1011,1035]" captionTargetBox="[174,1421,151,985]" captionTargetId="figure-355@7.[174,1421,151,985]" captionTargetPageId="7" captionText="Fig. 4. Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind legs, lateral view. C Forewing. D Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237581" httpUri="https://zenodo.org/record/15237581/files/figure.png" pageId="7" pageNumber="222">
Fig. 4
<collectionCode id="ED5BAE471575FF93FE33F9CDFE412A63" box="[442,459,1580,1606]" pageId="7" pageNumber="222">B</collectionCode>
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) with long tubular meracanthus. Metatibia longer than metafemur, weakly widening towards apex; apical metatibial spurs sclerotised, difficult to see, probably between 6 and
<quantity id="4CB29B671575FF93FD1CF971FD4F2A8F" box="[661,709,1680,1706]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="7" pageNumber="222" unit="in" value="8.0">8 in</quantity>
numbers (
<figureCitation id="13712A071575FF93FF67F950FEC82AEE" box="[238,322,1713,1740]" captionStart="Fig" captionStartId="7.[174,211,1011,1035]" captionTargetBox="[174,1421,151,985]" captionTargetId="figure-355@7.[174,1421,151,985]" captionTargetPageId="7" captionText="Fig. 4. Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind legs, lateral view. C Forewing. D Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237581" httpUri="https://zenodo.org/record/15237581/files/figure.png" pageId="7" pageNumber="222">
Fig. 4
<collectionCode id="ED5BAE471575FF93FEBBF950FEC82AEE" box="[306,322,1713,1739]" pageId="7" pageNumber="222">B</collectionCode>
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). Metatarsal segments moderately long, basal segment slightly longer. Forewing (
<figureCitation id="13712A071575FF93FD09F933FD592AC9" box="[640,723,1746,1773]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="7" pageNumber="222">
Fig. 1
<collectionCode id="ED5BAE471575FF93FD36F933FD592AC9" box="[703,723,1746,1772]" pageId="7" pageNumber="222">E</collectionCode>
</figureCitation>
,
<figureCitation id="13712A071575FF93FD6BF933FC8E2AC9" box="[738,772,1746,1772]" captionStart="Fig" captionStartId="7.[174,211,1011,1035]" captionTargetBox="[174,1421,151,985]" captionTargetId="figure-355@7.[174,1421,151,985]" captionTargetPageId="7" captionText="Fig. 4. Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind legs, lateral view. C Forewing. D Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237581" httpUri="https://zenodo.org/record/15237581/files/figure.png" pageId="7" pageNumber="222">
4
<collectionCode id="ED5BAE471575FF93FD78F933FC8E2AC9" box="[753,772,1746,1772]" pageId="7" pageNumber="222">C</collectionCode>
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) oval, widest in the middle; 2.3 times as long as wide, membranous; vein C+Sc moderately curved, slender, distinctly delimited to cell; cell c+sc wide; costal break developed, close to apex of vein R
<subScript id="17CE34C71575FF93FB26FB4FFB3D2899" attach="left" box="[1199,1207,1198,1212]" fontSize="6" pageId="7" pageNumber="222">1</subScript>
; pterostigma moderately wide, with subparallel margin in the median third, not expanded in apical third; entirely membranous; nodal line developed; vein R about as long as M+Cu; vein Rs mostly straight, weakly curved to costal margin basally and subapically; vein M much longer than M
<subScript id="17CE34C71575FF93FC2CFA97FC3729A1" attach="left" box="[933,957,1398,1412]" fontSize="6" pageId="7" pageNumber="222">1+2</subScript>
; vein Cu
<subScript id="17CE34C71575FF93FB97FA97FBA429A1" attach="left" box="[1054,1070,1398,1412]" fontSize="6" pageId="7" pageNumber="222">1a</subScript>
straight at base, curved towards anal margin in apical third; veins M
<subScript id="17CE34C71575FF93FB58FA79FB632983" attach="left" box="[1233,1257,1432,1446]" fontSize="6" pageId="7" pageNumber="222">1+2</subScript>
and M
<subScript id="17CE34C71575FF93FAB7FA79FAD22983" attach="left" box="[1342,1368,1432,1446]" fontSize="6" pageId="7" pageNumber="222">3+4</subScript>
perpendicular to wing margin apically; cell cu
<subScript id="17CE34C71575FF93FA99FA58FA9229E2" attach="left" box="[1296,1304,1465,1479]" fontSize="6" pageId="7" pageNumber="222">2</subScript>
long; anal break adjacent to apex of vein Cu
<subScript id="17CE34C71575FF93FB3CFA3BFB4E29CD" attach="left" box="[1205,1220,1498,1512]" fontSize="6" pageId="7" pageNumber="222">1b</subScript>
; surface spinules covering all cells, forming irregular transverse rows. Male unknown. Female terminalia (
<figureCitation id="13712A071575FF93FB41F9EBFA942A00" box="[1224,1310,1546,1573]" captionStart="Fig" captionStartId="7.[174,211,1011,1035]" captionTargetBox="[174,1421,151,985]" captionTargetId="figure-355@7.[174,1421,151,985]" captionTargetPageId="7" captionText="Fig. 4. Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind legs, lateral view. C Forewing. D Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237581" httpUri="https://zenodo.org/record/15237581/files/figure.png" pageId="7" pageNumber="222">
Fig. 4
<collectionCode id="ED5BAE471575FF93FA85F9EAFA942A00" box="[1292,1310,1547,1573]" pageId="7" pageNumber="222">D</collectionCode>
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) cuneate; proctiger mostly straight dorsally, weakly concave subapically; apex subacute; beset with moderately long hairs across the middle; subgenital plate mostly straight ventrally, slightly upcurved apically.
</paragraph>
<paragraph id="8BF536821575FF93FCA7F932FB292B0A" blockId="7.[814,1421,1746,1839]" pageId="7" pageNumber="222">
<emphasis id="B93EEA901575FF93FCA7F932FBDC2AC9" bold="true" box="[814,1110,1746,1773]" pageId="7" pageNumber="222">Measurements (in mm):</emphasis>
Head width 0.45; antenna length 0.65; forewing length 1.3; metatibia length 0.34; female proctiger length 0.3.
</paragraph>
</subSubSection>
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<paragraph id="8BF53682157AFF9CFF2FFB17FB332963" blockId="8.[166,1414,1270,1350]" pageId="8" pageNumber="223">
<emphasis id="B93EEA90157AFF9CFF2FFB17FF62292B" bold="true" box="[166,232,1270,1294]" pageId="8" pageNumber="223">Fig. 5.</emphasis>
<taxonomicName id="4C4A4D01157AFF9CFF7CFB17FD70292B" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[245,762,1270,1294]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="vetus" status="sp. nov.">
<emphasis id="B93EEA90157AFF9CFF7CFB17FE25292B" box="[245,431,1270,1294]" italics="true" pageId="8" pageNumber="223">Melanastera vetus</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB157AFF9CFC89FB16FCC4292B" box="[768,846,1271,1294]" pageId="8" pageNumber="223" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93EEA90157AFF9CFCDAFB17FCEF292B" bold="true" box="[851,869,1270,1294]" pageId="8" pageNumber="223">A</emphasis>
Head, frontodorsal view.
<emphasis id="B93EEA90157AFF9CFBF4FB17FB04292B" bold="true" box="[1149,1166,1270,1294]" pageId="8" pageNumber="223">B</emphasis>
Hind leg, lateral view.
<emphasis id="B93EEA90157AFF9CFF2FFAF3FF32290F" bold="true" box="[166,184,1298,1322]" pageId="8" pageNumber="223">C</emphasis>
Fragment of right forewing of female holotype, with M-Cu crossvein.
<emphasis id="B93EEA90157AFF9CFC10FAF3FC21290F" bold="true" box="[921,939,1298,1322]" pageId="8" pageNumber="223">D</emphasis>
Incomplete left forewing of female holotype lacking M-Cu crossvein.
<emphasis id="B93EEA90157AFF9CFE2FFACFFE3D2963" bold="true" box="[422,439,1326,1350]" pageId="8" pageNumber="223">E</emphasis>
Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY.
</paragraph>
</caption>
<subSubSection id="C3506509157AFF9CFF2FFA8BFB812B12" pageId="8" pageNumber="223" type="discussion">
<paragraph id="8BF53682157AFF9CFF2FFA8BFB812B12" blockId="8.[166,773,1386,1846]" lastBlockId="8.[807,1414,1386,1847]" pageId="8" pageNumber="223">
<emphasis id="B93EEA90157AFF9CFF2FFA8BFEBA29A1" bold="true" box="[166,304,1386,1412]" pageId="8" pageNumber="223">Comments:</emphasis>
In general habitus, in the venation of the forewings (with the vein
<collectionCode id="ED5BAE47157AFF9CFD9FFA6DFDA32983" box="[534,553,1420,1446]" pageId="8" pageNumber="223">C</collectionCode>
+Sc, which is not widened and clearly delimited from the neighbouring cell, and a uniform membranous pterostigma), in the structure of the hind legs (with several indistinctly grouped, sclerotised apical spurs on metatibiae) and in the female terminalia, the new species is similar to extant Neotropical species of
<taxonomicName id="4C4A4D01157AFF9CFDFFF9B5FE6E2AB5" authority="(BURCKHARDT et al. 2023)" baseAuthorityName="BURCKHARDT" baseAuthorityYear="2023" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEA90157AFF9CFDFFF9B5FC8F2A4B" box="[630,773,1620,1646]" italics="true" pageId="8" pageNumber="223">Melanastera</emphasis>
(
<bibRefCitation id="EFDB4B73157AFF9CFF39F994FE532AAA" author="BURCKHARDT, D. &amp; SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEI- ROZ, D. L. &amp; ALENE, D. C. &amp; CHO, G. &amp; PERCY, D. M." box="[176,473,1653,1680]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="8" pageNumber="223" pagination="387 - 421" part="201" publicationUrl="10.1093/zoolinnean/zlad128" refId="ref6525" refString="BURCKHARDT, D., SERBINA, L. S., MALENOVSKY, I., QUEI- ROZ, D. L., ALENE, D. C., CHO, G. &amp; PERCY, D. M. (2023): Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data. - Zoological Journal of the Linnean Society, 2023, XX, 1 - 35; advance access publication 9 October 2023. Printed version: (2024), 201: 387-421. DOI: 10.1093/zoolinnean/zlad128" title="Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data" type="journal article" year="2023">BURCKHARDT et al. 2023</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C4A4D01157AFF9CFE7CF994FD5B2AAA" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[501,721,1653,1679]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="casca">
<emphasis id="B93EEA90157AFF9CFE7CF994FD5B2AAA" box="[501,721,1653,1679]" italics="true" pageId="8" pageNumber="223">Melanastera casca</emphasis>
</taxonomicName>
differs from the described
<taxonomicName id="4C4A4D01157AFF9CFE41F977FD3E2A95" box="[456,692,1686,1712]" pageId="8" pageNumber="223">
<emphasis id="B93EEA90157AFF9CFE41F977FDDD2A95" box="[456,599,1686,1712]" italics="true" pageId="8" pageNumber="223">Melanastera</emphasis>
species
</taxonomicName>
in the pterostigma of the forewing with mostly subparallel margins rather than widened in the apical third. The characteristic pterostigma shape in
<taxonomicName id="4C4A4D01157AFF9CFDF7F91AFD622B31" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[638,744,1787,1812]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="casca">
<emphasis id="B93EEA90157AFF9CFDF7F91AFD622B31" box="[638,744,1787,1812]" italics="true" pageId="8" pageNumber="223">M. casca</emphasis>
</taxonomicName>
is shared by
<taxonomicName id="4C4A4D01157AFF9CFEABF8FCFC8F2B13" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[290,773,1820,1846]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="vetus" status="sp. nov.">
<emphasis id="B93EEA90157AFF9CFEABF8FCFE0C2B13" box="[290,390,1821,1846]" italics="true" pageId="8" pageNumber="223">M. vetus</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
</taxonomicName>
<taxonomicNameLabel id="A20D57EB157AFF9CFCAEFA8AFCF529A0" box="[807,895,1387,1413]" pageId="8" pageNumber="223" rank="species">sp. nov.</taxonomicNameLabel>
and four undescribed species from southern and southeastern Brazil associated with
<taxonomicName id="4C4A4D01157AFF9CFA81FA6DFA0F2983" box="[1288,1413,1420,1446]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="223" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
(hosts confirmed for three species by the presence of immatures and likely for one species). The four extant species will be described in a monograph on the
<taxonomicName id="4C4A4D01157AFF9CFADBFA11FC222A0E" authorityName="VONDRACEK" authorityYear="1963" class="Insecta" family="Liviidae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="226" phylum="Arthropoda" rank="tribe" tribe="Paurocephalini">Paurocephalini</taxonomicName>
from Brazil (
<bibRefCitation id="EFDB4B73157AFF9CFBCBF9F0FA972A0E" author="SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEIROZ, D. L. &amp; BURCKHARDT, D." box="[1090,1309,1553,1580]" journalOrPublisher="Zootaxa" pageId="8" pageNumber="223" refId="ref7781" refString="SERBINA, L. S., MALENOVSKY, I., QUEIROZ, D. L. &amp; BURCKHARDT, D. (2024): Jumping plant-lice of the tribe Paurocephalini (Hemiptera: Psylloidea: Liviidae) in Brazil. - Zootaxa, submitted." title="Jumping plant-lice of the tribe Paurocephalini (Hemiptera: Psylloidea: Liviidae) in Brazil" type="book" year="2024">SERBINA et al. 2024</bibRefCitation>
).
<taxonomicName id="4C4A4D01157AFF9CFABDF9F0FC3F2A69" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="casca">
<emphasis id="B93EEA90157AFF9CFABDF9F0FC3F2A69" italics="true" pageId="8" pageNumber="223">Melanastera casca</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4A4D01157AFF9CFBC3F9D2FB232A69" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[1098,1193,1587,1612]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="vetus">
<emphasis id="B93EEA90157AFF9CFBC3F9D2FB232A69" box="[1098,1193,1587,1612]" italics="true" pageId="8" pageNumber="223">M. vetus</emphasis>
</taxonomicName>
by the much longer cell cu
<subScript id="17CE34C7157AFF9CFCFAF987FCF12A51" attach="left" box="[883,891,1638,1652]" fontSize="6" pageId="8" pageNumber="223">1</subScript>
of the forewing and the shorter, more robust female terminalia. From the extant species associated with
<taxonomicName id="4C4A4D01157AFF9CFC11F976FB922A94" box="[920,1048,1687,1713]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="223" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
,
<taxonomicName id="4C4A4D01157AFF9CFBACF976FB062A95" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[1061,1164,1687,1712]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="casca">
<emphasis id="B93EEA90157AFF9CFBACF976FB062A95" box="[1061,1164,1687,1712]" italics="true" pageId="8" pageNumber="223">M. casca</emphasis>
</taxonomicName>
differs in the broader forewings (2.2 versus&gt; 2.4) and the relatively short female proctiger with an almost straight dorsal outline (versus relatively long with a distinctly sinuate or concave dorsal outline).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,87 @@
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Genus
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<emphasis id="B93EEA901577FF91FEB5FB9FFE1428BD" box="[316,414,1150,1176]" italics="true" pageId="5" pageNumber="220">Klyveria</emphasis>
BURCKHARDT, SERBINA &amp; MALENOVSKÝ
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in
<bibRefCitation id="EFDB4B731577FF91FE26FB41FD40289F" author="BURCKHARDT, D. &amp; SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEI- ROZ, D. L. &amp; ALENE, D. C. &amp; CHO, G. &amp; PERCY, D. M." box="[431,714,1184,1210]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="5" pageNumber="220" pagination="387 - 421" part="201" publicationUrl="10.1093/zoolinnean/zlad128" refId="ref6525" refString="BURCKHARDT, D., SERBINA, L. S., MALENOVSKY, I., QUEI- ROZ, D. L., ALENE, D. C., CHO, G. &amp; PERCY, D. M. (2023): Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data. - Zoological Journal of the Linnean Society, 2023, XX, 1 - 35; advance access publication 9 October 2023. Printed version: (2024), 201: 387-421. DOI: 10.1093/zoolinnean/zlad128" title="Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data" type="journal article" year="2023">BURCKHARDT et al. 2023</bibRefCitation>
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<emphasis id="B93EEA901577FF91FF27FB02FEC728D9" bold="true" box="[174,333,1250,1277]" pageId="5" pageNumber="220">
<typeStatus id="54F188201577FF91FF27FB02FF6328D8" box="[174,233,1251,1277]" pageId="5" pageNumber="220">Type</typeStatus>
species:
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<taxonomicName id="4C4A4D011577FF91FEDCFB03FD2728D9" baseAuthorityName="BURCKHARDT" baseAuthorityYear="1996" box="[341,685,1250,1276]" class="Insecta" family="Liviidae" genus="Paurocephala" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="crassiflagellata">
<emphasis id="B93EEA901577FF91FEDCFB03FD2728D9" box="[341,685,1250,1276]" italics="true" pageId="5" pageNumber="220">Paurocephala crassiflagellata</emphasis>
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BURCKHARDT, 1996.
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BURCKHARDT &amp; DROHOJOWSKA
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<figureCitation id="13712A071577FF91FE1FFA68FD822981" box="[406,520,1417,1444]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="5" pageNumber="220">Fig. 1C, D</figureCitation>
,
<figureCitation id="13712A071577FF91FD9FFA68FDAE2986" box="[534,548,1417,1443]" captionStart="Fig" captionStartId="6.[166,203,899,923]" captionTargetBox="[166,1413,151,872]" captionTargetId="figure-13@6.[166,1413,151,872]" captionTargetPageId="6" captionText="Fig. 3. Klyveria sucina BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view. C Forewing. D Female terminalia, oblique lateral view; sternite 6 is broken in the middle. Scale = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237579" httpUri="https://zenodo.org/record/15237579/files/figure.png" pageId="5" pageNumber="220">3</figureCitation>
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<emphasis id="B93EEA901577FF91FF27FA2AFF7329C0" bold="true" box="[174,249,1483,1509]" pageId="5" pageNumber="220">LSID:</emphasis>
urn:lsid:zoobank.org:act:7FE418DF-D9EB-4081-A0BC-779B2530276E
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<emphasis id="B93EEA901577FF91FF27F9D1FEB02A6F" bold="true" box="[174,314,1584,1610]" pageId="5" pageNumber="220">Etymology:</emphasis>
From the Latin adjective sucinus (1
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/2
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declension) = of amber, referring to the origin of the species from amber.
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Female specimen, sample #D-14-1146 (
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.
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<materialsCitation id="3B223CDF1577FF91FF27F8F8FD3D2B16" box="[174,695,1817,1843]" collectionCode="GPAC" country="Dominican Republic" location="Cordillera Septentrional" pageId="5" pageNumber="220" typeStatus="paratype">
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<typeStatus id="54F188201577FF91FF27F8F8FEAA2B16" box="[174,288,1817,1843]" pageId="5" pageNumber="220">Paratype</typeStatus>
:
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, sample #31(1) (
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.
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<materialsCitation id="3B223CDF1577FF91FCA7FF76FC432CF7" country="Dominican Republic" pageId="5" pageNumber="220" typeStatus="Type">
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<typeStatus id="54F188201577FF91FCA7FF76FCE32C94" box="[814,873,151,177]" pageId="5" pageNumber="220">Type</typeStatus>
locality:
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<collectingCountry id="F35D76121577FF91FC77FF76FB742C94" box="[1022,1278,151,177]" name="Dominican Republic" pageId="5" pageNumber="220" value="Dominican Republic">Dominican Republic</collectingCountry>
, Cordillera Septentrional
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.
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<emphasis id="B93EEA901577FF91FCA7FF1BFBA02D31" bold="true" box="[814,1066,250,276]" pageId="5" pageNumber="220">Description: Adult:</emphasis>
Colouration. Light brown. Antenna brown, segments 48 with apical third dark brown and segments 9 and 10 entirely dark brown. Metascutum with two brown sublateral dots on either side. Forewing colourless, transparent with dark apices of veins along apical wing margin.
</paragraph>
<paragraph id="8BF536821577FF91FCC4FE22FBA62A0C" blockId="5.[814,1422,250,1577]" pageId="5" pageNumber="220">
Structure. Vertex (
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Fig. 1
<collectionCode id="ED5BAE471577FF91FBEDFE22FBF22DF8" box="[1124,1144,451,477]" pageId="5" pageNumber="220">C</collectionCode>
</figureCitation>
,
<figureCitation id="13712A071577FF91FB01FE22FB2D2DF8" box="[1160,1191,451,477]" captionStart="Fig" captionStartId="6.[166,203,899,923]" captionTargetBox="[166,1413,151,872]" captionTargetId="figure-13@6.[166,1413,151,872]" captionTargetPageId="6" captionText="Fig. 3. Klyveria sucina BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view. C Forewing. D Female terminalia, oblique lateral view; sternite 6 is broken in the middle. Scale = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237579" httpUri="https://zenodo.org/record/15237579/files/figure.png" pageId="5" pageNumber="220">
3
<collectionCode id="ED5BAE471577FF91FB1AFE22FB2D2DF8" box="[1171,1191,451,477]" pageId="5" pageNumber="220">A</collectionCode>
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) rhomboidal, transverse, 2.5 times as wide as long; passing smoothly into genae anteriorly; coronal suture not visible basally; genae not enlarged into processes; frons small, trapezoidal (
<figureCitation id="13712A071577FF91FC08FDA9FC592E47" box="[897,979,584,610]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="5" pageNumber="220">
Fig. 1
<collectionCode id="ED5BAE471577FF91FC35FDA9FC592E47" box="[956,979,584,610]" pageId="5" pageNumber="220">D</collectionCode>
</figureCitation>
); median ocellus visible in perpendicular view to vertex; compound eyes, in dorsal view, subglobular, slightly collard. Clypeus pear-shaped, large, flattened ventrally (
<figureCitation id="13712A071577FF91FB98FD4DFBED2EE3" box="[1041,1127,684,710]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="5" pageNumber="220">
Fig. 1
<collectionCode id="ED5BAE471577FF91FBD8FD4DFBED2EE3" box="[1105,1127,684,710]" pageId="5" pageNumber="220">D</collectionCode>
</figureCitation>
). Antenna 10-segmented, longer than head width; relative length of flagellar segments as 1.0: 0.4: 0.4: 0.4: 0.3: 0.3: 0.4: 0.2; terminal setae not visible. Pronotum, in dorsal view, hardly curved posteriad laterally, 0.6 times as long as vertex along median longitudinal body axis. Metacoxa (
<figureCitation id="13712A071577FF91FCBFFC95FC0D2FAB" box="[822,903,884,910]" captionStart="Fig" captionStartId="6.[166,203,899,923]" captionTargetBox="[166,1413,151,872]" captionTargetId="figure-13@6.[166,1413,151,872]" captionTargetPageId="6" captionText="Fig. 3. Klyveria sucina BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view. C Forewing. D Female terminalia, oblique lateral view; sternite 6 is broken in the middle. Scale = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237579" httpUri="https://zenodo.org/record/15237579/files/figure.png" pageId="5" pageNumber="220">
Fig. 3
<collectionCode id="ED5BAE471577FF91FCFCFC95FC0D2FAB" box="[885,903,884,910]" pageId="5" pageNumber="220">B</collectionCode>
</figureCitation>
) with slender subacute horn-shaped meracanthus. Metatibia (
<figureCitation id="13712A071577FF91FC6DFC74FBBC2F8A" box="[996,1078,917,944]" captionStart="Fig" captionStartId="6.[166,203,899,923]" captionTargetBox="[166,1413,151,872]" captionTargetId="figure-13@6.[166,1413,151,872]" captionTargetPageId="6" captionText="Fig. 3. Klyveria sucina BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view. C Forewing. D Female terminalia, oblique lateral view; sternite 6 is broken in the middle. Scale = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237579" httpUri="https://zenodo.org/record/15237579/files/figure.png" pageId="5" pageNumber="220">
Fig. 3
<collectionCode id="ED5BAE471577FF91FBAAFC74FBBC2F8A" box="[1059,1078,917,943]" pageId="5" pageNumber="220">B</collectionCode>
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) much longer than metafemur, hardly widened apically; lacking sclerotised spurs along its length; bearing a posteriorly open crown of densely spaced apical, weakly sclerotised spurs. Both metatarsal segments moderately long, subequal in length. Forewing (
<figureCitation id="13712A071577FF91FC74FBDAFBD82873" box="[1021,1106,1083,1110]" captionStart="Fig" captionStartId="6.[166,203,899,923]" captionTargetBox="[166,1413,151,872]" captionTargetId="figure-13@6.[166,1413,151,872]" captionTargetPageId="6" captionText="Fig. 3. Klyveria sucina BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view. C Forewing. D Female terminalia, oblique lateral view; sternite 6 is broken in the middle. Scale = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237579" httpUri="https://zenodo.org/record/15237579/files/figure.png" pageId="5" pageNumber="220">
Fig. 3
<collectionCode id="ED5BAE471577FF91FBB6FBDDFBD82873" box="[1087,1106,1084,1110]" pageId="5" pageNumber="220">C</collectionCode>
</figureCitation>
) oval, widest in the middle; 2.3 times as long as wide, membranous; vein C+Sc evenly curved, delimitation to cell not clearly visible; cell c+sc wide; costal break developed, close to apex of vein R
<subScript id="17CE34C71577FF91FC10FB35FC2B28C7" attach="left" box="[921,929,1236,1250]" fontSize="6" pageId="5" pageNumber="220">1</subScript>
; pterostigma indistinct, not clearly visible; nodal line developed; veins R and M+Cu subequal; vein Rs almost straight; vein M shorter than M
<subScript id="17CE34C71577FF91FAE4FAF7FA0F2901" attach="left" box="[1389,1413,1302,1316]" fontSize="6" pageId="5" pageNumber="220">1+2</subScript>
; vein Cu 1a strongly curved towards anal margin; veins M
<subScript id="17CE34C71577FF91FCCEFAB8FCD52942" attach="left" box="[839,863,1369,1383]" fontSize="6" pageId="5" pageNumber="220">1+2</subScript>
and M
<subScript id="17CE34C71577FF91FC3FFAB8FC5A2942" attach="left" box="[950,976,1369,1383]" fontSize="6" pageId="5" pageNumber="220">3+4</subScript>
almost perpendicular to wing margin apically; vein Cu shorter than Cu
<subScript id="17CE34C71577FF91FB3BFA9BFB4829AD" attach="left" box="[1202,1218,1402,1416]" fontSize="6" pageId="5" pageNumber="220">1b</subScript>
; anal break adjacent to apex of vein Cu
<subScript id="17CE34C71577FF91FBBCFA7DFBCE298F" attach="right" box="[1077,1092,1436,1450]" fontSize="6" pageId="5" pageNumber="220">1b</subScript>
; surface spinules not visible. Hindwing almost as long as forewing. Male unknown. Female terminalia (
<figureCitation id="13712A071577FF91FB83FA2DFBD429C3" box="[1034,1118,1484,1510]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="5" pageNumber="220">
Fig. 1
<collectionCode id="ED5BAE471577FF91FBCEFA2DFBD429C3" box="[1095,1118,1484,1510]" pageId="5" pageNumber="220">D</collectionCode>
</figureCitation>
,
<figureCitation id="13712A071577FF91FBE5FA2DFB0629C3" box="[1132,1164,1484,1510]" captionStart="Fig" captionStartId="6.[166,203,899,923]" captionTargetBox="[166,1413,151,872]" captionTargetId="figure-13@6.[166,1413,151,872]" captionTargetPageId="6" captionText="Fig. 3. Klyveria sucina BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, dorsal view. B Hind leg, lateral view. C Forewing. D Female terminalia, oblique lateral view; sternite 6 is broken in the middle. Scale = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237579" httpUri="https://zenodo.org/record/15237579/files/figure.png" pageId="5" pageNumber="220">
3
<collectionCode id="ED5BAE471577FF91FBF1FA2DFB0629C3" box="[1144,1164,1484,1510]" pageId="5" pageNumber="220">D</collectionCode>
</figureCitation>
) partly damaged; both proctiger and subgenital plate with massive base and slender apical process.
</paragraph>
<paragraph id="8BF536821577FF91FCA7F9B0FBFE2A8B" blockId="5.[814,1421,1617,1710]" pageId="5" pageNumber="220">
<emphasis id="B93EEA901577FF91FCA7F9B0FBD02A4E" bold="true" box="[814,1114,1617,1643]" pageId="5" pageNumber="220">Measurements (in mm):</emphasis>
Head width 0.6; antenna length 0.9; forewing length 1.5; metatibia length 0.5; female proctiger length 0.28.
</paragraph>
</subSubSection>
<subSubSection id="C35065091577FF92FCA7F937FC1C2873" lastPageId="6" lastPageNumber="221" pageId="5" pageNumber="220" type="discussion">
<paragraph id="8BF536821577FF92FCA7F937FC1C2873" blockId="5.[814,1422,1750,1844]" lastBlockId="6.[807,1414,1050,1110]" lastPageId="6" lastPageNumber="221" pageId="5" pageNumber="220">
<emphasis id="B93EEA901577FF91FCA7F937FC302AD5" bold="true" box="[814,954,1750,1776]" pageId="5" pageNumber="220">Comments:</emphasis>
The new species has the following characters which diagnose
<taxonomicName id="4C4A4D011577FF91FBB4F919FB2A2B37" box="[1085,1184,1784,1810]" class="Insecta" family="Liviidae" genus="Klyveria" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEA901577FF91FBB4F919FB2A2B37" box="[1085,1184,1784,1810]" italics="true" pageId="5" pageNumber="220">Klyveria</emphasis>
</taxonomicName>
within the
<taxonomicName id="4C4A4D011577FF91FAA4F919FA072B37" authorityName="LOW" authorityYear="1879" box="[1325,1421,1784,1810]" class="Insecta" family="Liviidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="225" phylum="Arthropoda" rank="subFamily" subFamily="Liviinae">Liviinae</taxonomicName>
(
<bibRefCitation id="EFDB4B731577FF91FCB0F8F8FBD02B16" author="BURCKHARDT, D. &amp; SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEI- ROZ, D. L. &amp; ALENE, D. C. &amp; CHO, G. &amp; PERCY, D. M." box="[825,1114,1817,1844]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="5" pageNumber="220" pagination="387 - 421" part="201" publicationUrl="10.1093/zoolinnean/zlad128" refId="ref6525" refString="BURCKHARDT, D., SERBINA, L. S., MALENOVSKY, I., QUEI- ROZ, D. L., ALENE, D. C., CHO, G. &amp; PERCY, D. M. (2023): Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data. - Zoological Journal of the Linnean Society, 2023, XX, 1 - 35; advance access publication 9 October 2023. Printed version: (2024), 201: 387-421. DOI: 10.1093/zoolinnean/zlad128" title="Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data" type="journal article" year="2023">BURCKHARDT et al. 2023</bibRefCitation>
): head with moderately to strongly transverse vertex and subspherical, collared eyes; metatibia with an apical crown of weakly sclerotised spurs, without strong spurs laterally; female terminalia slender and relatively long, subgenital plate with a narrow apical process.
<taxonomicName id="4C4A4D011574FF92FE74FB41FD38289F" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[509,690,1184,1210]" class="Insecta" family="Liviidae" genus="Klyveria" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="sucina">
<emphasis id="B93EEA901574FF92FE74FB41FD38289F" box="[509,690,1184,1210]" italics="true" pageId="6" pageNumber="221">Klyveria sucina</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4A4D011574FF92FF6DFB23FD3F28F9" authority="(BURCKHARDT, 1996)" baseAuthorityName="BURCKHARDT" baseAuthorityYear="1996" box="[228,693,1217,1244]" class="Insecta" family="Liviidae" genus="Paurocephala" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="crassiflagellata">
<emphasis id="B93EEA901574FF92FF6DFB23FE3228FE" box="[228,440,1217,1243]" italics="true" pageId="6" pageNumber="221">K. crassiflagellata</emphasis>
(
<bibRefCitation id="EFDB4B731574FF92FE41FB20FD2128FE" author="BURCKHARDT, D." box="[456,683,1217,1243]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="6" pageNumber="221" pagination="77 - 82" part="43" publicationUrl="10.1002/mmnd.19960430106" refId="ref6136" refString="BURCKHARDT, D. (1996): Two new Paurocephala species from Paraguay (Hemiptera, Psylloidea). - Deutsche Entomologische Zeitschrift, 43: 77-82. DOI: 10.1002/mmnd.19960430106." title="Two new Paurocephala species from Paraguay (Hemiptera, Psylloidea)" type="journal article" year="1996">BURCKHARDT, 1996</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C4A4D011574FF92FD62FB23FD9628D8" authority="(BURCKHARDT, 1996)" baseAuthorityName="BURCKHARDT" baseAuthorityYear="1996" class="Insecta" family="Liviidae" genus="Klyveria" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="setinervis">
<emphasis id="B93EEA901574FF92FD62FB23FE9C28D9" italics="true" pageId="6" pageNumber="221">K. setinervis</emphasis>
(
<bibRefCitation id="EFDB4B731574FF92FEA3FB02FD9828D9" author="BURCKHARDT, D." box="[298,530,1250,1277]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="6" pageNumber="221" pagination="77 - 82" part="43" publicationUrl="10.1002/mmnd.19960430106" refId="ref6136" refString="BURCKHARDT, D. (1996): Two new Paurocephala species from Paraguay (Hemiptera, Psylloidea). - Deutsche Entomologische Zeitschrift, 43: 77-82. DOI: 10.1002/mmnd.19960430106." title="Two new Paurocephala species from Paraguay (Hemiptera, Psylloidea)" type="journal article" year="1996">BURCKHARDT, 1996</bibRefCitation>
)
</taxonomicName>
, the only described species of the genus (
<bibRefCitation id="EFDB4B731574FF92FE17FAE5FDF7293B" author="BURCKHARDT, D." box="[414,637,1284,1310]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="6" pageNumber="221" pagination="77 - 82" part="43" publicationUrl="10.1002/mmnd.19960430106" refId="ref6136" refString="BURCKHARDT, D. (1996): Two new Paurocephala species from Paraguay (Hemiptera, Psylloidea). - Deutsche Entomologische Zeitschrift, 43: 77-82. DOI: 10.1002/mmnd.19960430106." title="Two new Paurocephala species from Paraguay (Hemiptera, Psylloidea)" type="journal article" year="1996">BURCKHARDT 1996</bibRefCitation>
), in the relatively longer head, slender antennal segment 3, and in following details of the forewing: cell c+sc broad (rather than narrow), vein M shorter than M
<subScript id="17CE34C71574FF92FD13FA9BFD3929AD" attach="left" box="[666,691,1402,1416]" fontSize="6" pageId="6" pageNumber="221">1+2</subScript>
(rather than longer) and vein Cu shorter than Cu
<subScript id="17CE34C71574FF92FD14FA7DFD27298F" attach="left" box="[669,685,1436,1450]" fontSize="6" pageId="6" pageNumber="221">1b</subScript>
(rather than longer). The meracanthus of
<taxonomicName id="4C4A4D011574FF92FDAEFA4AFD1929E1" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[551,659,1451,1476]" class="Insecta" family="Liviidae" genus="Klyveria" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="sucina">
<emphasis id="B93EEA901574FF92FDAEFA4AFD1929E1" box="[551,659,1451,1476]" italics="true" pageId="6" pageNumber="221">K. sucina</emphasis>
</taxonomicName>
is slender and tubular rather than horn-shaped as in the other
<taxonomicName id="4C4A4D011574FF92FF2FFA0CFEFB2A22" box="[166,369,1517,1543]" class="Insecta" family="Liviidae" genus="Klyveria" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93EEA901574FF92FF2FFA0CFE832A22" box="[166,265,1517,1543]" italics="true" pageId="6" pageNumber="221">Klyveria</emphasis>
species.
</taxonomicName>
This is probably an artefact due to compression of the specimen. The delimitation of vein C+Sc towards the cell is difficult to see in the
<specimenCount id="9D4CFD0B1574FF92FF5AF9B3FEF52A4E" box="[211,383,1617,1644]" count="2" pageId="6" pageNumber="221" type="generic">two specimens</specimenCount>
at hand of the new species. This makes it difficult to judge whether the species differs in this character from other congeners. According to
<bibRefCitation id="EFDB4B731574FF92FF2FF954FE5A2AEA" author="BURCKHARDT, D. &amp; SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEI- ROZ, D. L. &amp; ALENE, D. C. &amp; CHO, G. &amp; PERCY, D. M." box="[166,464,1717,1744]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="6" pageNumber="221" pagination="387 - 421" part="201" publicationUrl="10.1093/zoolinnean/zlad128" refId="ref6525" refString="BURCKHARDT, D., SERBINA, L. S., MALENOVSKY, I., QUEI- ROZ, D. L., ALENE, D. C., CHO, G. &amp; PERCY, D. M. (2023): Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data. - Zoological Journal of the Linnean Society, 2023, XX, 1 - 35; advance access publication 9 October 2023. Printed version: (2024), 201: 387-421. DOI: 10.1093/zoolinnean/zlad128" title="Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data" type="journal article" year="2023">BURCKHARDT et al. (2023)</bibRefCitation>
the metatarsal segments in
<taxonomicName id="4C4A4D011574FF92FF2FF937FE832AD5" box="[166,265,1750,1776]" class="Insecta" family="Liviidae" genus="Klyveria" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEA901574FF92FF2FF937FE832AD5" box="[166,265,1750,1776]" italics="true" pageId="6" pageNumber="221">Klyveria</emphasis>
</taxonomicName>
are relatively short, but in the fossil species they are moderately long. This character is apparently more variable than suggested by
<bibRefCitation id="EFDB4B731574FF92FDEFF8F8FC3F2810" author="BURCKHARDT, D. &amp; SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEI- ROZ, D. L. &amp; ALENE, D. C. &amp; CHO, G. &amp; PERCY, D. M." box="[614,949,1050,1843]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="6" pageNumber="221" pagination="387 - 421" part="201" publicationUrl="10.1093/zoolinnean/zlad128" refId="ref6525" refString="BURCKHARDT, D., SERBINA, L. S., MALENOVSKY, I., QUEI- ROZ, D. L., ALENE, D. C., CHO, G. &amp; PERCY, D. M. (2023): Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data. - Zoological Journal of the Linnean Society, 2023, XX, 1 - 35; advance access publication 9 October 2023. Printed version: (2024), 201: 387-421. DOI: 10.1093/zoolinnean/zlad128" title="Phylogeny and classification of jumping plant lice of the subfamily Liviinae (Hemiptera: Psylloidea: Liviidae) based on molecular and morphological data" type="journal article" year="2023">BURCKHARDT et al. (2023)</bibRefCitation>
and the generic diagnosis needs to be amended.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="83ACB3CAEFFD370D511F4C22A0D35FE4">First record of jumping plant lice of the family Liviidae (Hemiptera, Sternorrhyncha, Psylloidea) from Dominican amber</mods:title>
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<mods:namePart id="DD6E2A346156727FB7C30409FB2560C8">Burckhardt, Daniel</mods:namePart>
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<mods:namePart id="96F3F369847C6342ADFE9935856AB8C6">Drohojowska, Jowita</mods:namePart>
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<mods:namePart id="D3B9D6EF997358E773BA29AA292C1995">Štarhová Serbina, Liliya</mods:namePart>
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<mods:namePart id="C53A9913169BA08AA110C041569D1790">Malenovský, Igor</mods:namePart>
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<paragraph id="8BF53682157BFF9DFF35FF76FD822CF6" blockId="9.[188,766,151,244]" pageId="9" pageNumber="224">
<heading id="D0BD81EE157BFF9DFF35FF76FD822CF6" centered="true" fontSize="11" level="2" pageId="9" pageNumber="224" reason="4">
<taxonomicName id="4C4A4D01157BFF9DFF35FF76FD742C94" authority="BURCKHARDT &amp; DROHOJOWSKA" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[188,766,151,177]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="224" phylum="Arthropoda" rank="species" species="vetus" status="sp. nov.">
<emphasis id="B93EEA90157BFF9DFF35FF76FE062C95" box="[188,396,151,177]" italics="true" pageId="9" pageNumber="224">Melanastera vetus</emphasis>
BURCKHARDT &amp; DROHOJOWSKA
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<subSubSection id="C3506509157BFF9DFE20FF38FD8B2D72" pageId="9" pageNumber="224" type="description">
<paragraph id="8BF53682157BFF9DFE20FF38FD9B2CD1" blockId="9.[188,766,151,244]" box="[425,529,217,244]" pageId="9" pageNumber="224">
<figureCitation id="13712A07157BFF9DFE20FF38FE7F2CD6" box="[425,501,217,244]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="9" pageNumber="224">Fig. 1F</figureCitation>
,
<figureCitation id="13712A07157BFF9DFD8AFF38FD9B2CD1" box="[515,529,217,244]" captionStart="Fig" captionStartId="8.[166,203,1270,1294]" captionTargetBox="[166,1413,151,1244]" captionTargetId="figure-258@8.[166,1413,151,1244]" captionTargetPageId="8" captionText="Fig. 5. Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, frontodorsal view. B Hind leg, lateral view. C Fragment of right forewing of female holotype, with M-Cu crossvein. D Incomplete left forewing of female holotype lacking M-Cu crossvein. E Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237583" httpUri="https://zenodo.org/record/15237583/files/figure.png" pageId="9" pageNumber="224">5</figureCitation>
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<emphasis id="B93EEA90157BFF9DFF27FEFDFF732D13" bold="true" box="[174,249,284,310]" pageId="9" pageNumber="224">LSID:</emphasis>
urn:lsid:zoobank.org:act:257D080B-093B-4E2D-B6AD-90467B4AB421
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<subSubSection id="C3506509157BFF9DFF27FE61FECE2DF8" pageId="9" pageNumber="224" type="etymology">
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<emphasis id="B93EEA90157BFF9DFF27FE61FEB02DBF" bold="true" box="[174,314,384,410]" pageId="9" pageNumber="224">Etymology:</emphasis>
From the Latin adjective vetus (3
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declension) = old, aged, ancient, referring to the age of the fossils.
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<subSubSection id="C3506509157BFF9DFF27FDE4FEC72EA6" pageId="9" pageNumber="224" type="materials_examined">
<paragraph id="8BF53682157BFF9DFF27FDE4FD692E3A" blockId="9.[174,739,517,544]" box="[174,739,517,544]" pageId="9" pageNumber="224">
<materialsCitation id="3B223CDF157BFF9DFF27FDE4FD542E3A" box="[174,734,517,544]" collectionCode="GPAC" country="Dominican Republic" location="Cordillera Septentrional" pageId="9" pageNumber="224" typeStatus="holotype">
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<typeStatus id="54F18820157BFF9DFF27FDE4FEAA2E3A" box="[174,288,517,543]" pageId="9" pageNumber="224">Holotype</typeStatus>
:
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Female specimen, sample #35 (
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)
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.
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<materialsCitation id="3B223CDF157BFF9DFF27FDA9FEC32EA6" country="Dominican Republic" pageId="9" pageNumber="224" typeStatus="Type">
<emphasis id="B93EEA90157BFF9DFF27FDA9FEEE2E47" bold="true" box="[174,356,584,610]" pageId="9" pageNumber="224">
<typeStatus id="54F18820157BFF9DFF27FDA9FF632E47" box="[174,233,584,610]" pageId="9" pageNumber="224">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F35D7612157BFF9DFEF7FDA9FDF72E47" box="[382,637,584,610]" name="Dominican Republic" pageId="9" pageNumber="224" value="Dominican Republic">Dominican Republic</collectingCountry>
, Cordillera Septentrional
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.
</paragraph>
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<subSubSection id="C3506509157BFF9DFF27FD4DFBF82D5C" pageId="9" pageNumber="224" type="description">
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<emphasis id="B93EEA90157BFF9DFF27FD4DFE202EE3" bold="true" box="[174,426,684,710]" pageId="9" pageNumber="224">Description: Adult:</emphasis>
Colouration. Dark brown. Antenna entirely dark brown. Forewing veins and pterostigma dark brown; membrane colourless, transparent with brown patches at the apex of veins along wing margin.
</paragraph>
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Structure. Head, in dorsal view, wider that pronotum, slightly narrower than thorax, moderately transverse (
<figureCitation id="13712A07157BFF9DFF77FC74FED82F8A" box="[254,338,917,944]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="9" pageNumber="224">
Fig. 1
<collectionCode id="ED5BAE47157BFF9DFEC8FC74FED82F8A" box="[321,338,917,943]" pageId="9" pageNumber="224">F</collectionCode>
</figureCitation>
,
<figureCitation id="13712A07157BFF9DFEEFFC74FE0C2F95" box="[358,390,917,944]" captionStart="Fig" captionStartId="8.[166,203,1270,1294]" captionTargetBox="[166,1413,151,1244]" captionTargetId="figure-258@8.[166,1413,151,1244]" captionTargetPageId="8" captionText="Fig. 5. Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, frontodorsal view. B Hind leg, lateral view. C Fragment of right forewing of female holotype, with M-Cu crossvein. D Incomplete left forewing of female holotype lacking M-Cu crossvein. E Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237583" httpUri="https://zenodo.org/record/15237583/files/figure.png" pageId="9" pageNumber="224">
5
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). Vertex subrhomboidal; passing smoothly into genae anteriorly; coronal suture fully developed; median ocellus not visible; compound eyes, in dorsal view, hemispherical, appressed to head. Antenna 10-segmented, slightly longer than head width; subdivisions only partly visible; relative lengths of antennal segment 10 and terminal setae as 1.0: 0.6: 1.0. Metacoxa (
<figureCitation id="13712A07157BFF9DFE58FB9FFDAF28BC" box="[465,549,1150,1177]" captionStart="Fig" captionStartId="8.[166,203,1270,1294]" captionTargetBox="[166,1413,151,1244]" captionTargetId="figure-258@8.[166,1413,151,1244]" captionTargetPageId="8" captionText="Fig. 5. Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, frontodorsal view. B Hind leg, lateral view. C Fragment of right forewing of female holotype, with M-Cu crossvein. D Incomplete left forewing of female holotype lacking M-Cu crossvein. E Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237583" httpUri="https://zenodo.org/record/15237583/files/figure.png" pageId="9" pageNumber="224">
Fig. 5
<collectionCode id="ED5BAE47157BFF9DFD9DFB9EFDAF28BC" box="[532,549,1151,1177]" pageId="9" pageNumber="224">B</collectionCode>
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) with blunt, tubular meracanthus. Metatibia longer than metafemur, widened apically; bearing sclerotised apical spurs, their number and grouping is not visible (
<figureCitation id="13712A07157BFF9DFDD0FB03FD2528D8" box="[601,687,1250,1277]" captionStart="Fig" captionStartId="8.[166,203,1270,1294]" captionTargetBox="[166,1413,151,1244]" captionTargetId="figure-258@8.[166,1413,151,1244]" captionTargetPageId="8" captionText="Fig. 5. Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, frontodorsal view. B Hind leg, lateral view. C Fragment of right forewing of female holotype, with M-Cu crossvein. D Incomplete left forewing of female holotype lacking M-Cu crossvein. E Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237583" httpUri="https://zenodo.org/record/15237583/files/figure.png" pageId="9" pageNumber="224">
Fig. 5
<collectionCode id="ED5BAE47157BFF9DFD17FB02FD2528D8" box="[670,687,1251,1277]" pageId="9" pageNumber="224">B</collectionCode>
</figureCitation>
). Metatarsal segments moderately long, subequal in length. Forewing (
<figureCitation id="13712A07157BFF9DFEA0FAC4FEF3291A" box="[297,377,1317,1344]" captionStart="Fig" captionStartId="2.[166,203,1804,1828]" captionTargetBox="[166,1413,151,1779]" captionTargetId="figure-13@2.[166,1413,151,1782]" captionTargetPageId="2" captionText="Fig. 1. Species of the tribe Paurocephalini in Dominican amber. A Diclidophlebia venosa BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of male holotype, lateral view. B Same but mid and hind legs of female paratype, ventral view. C KlyveriasucinaBURCKHARDT&amp;DROHOJOWSKAsp.nov.,habitusoffemaleholotype,dorsalview.DSamebutventralview. E Melanastera casca BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view. F Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov., habitus of female holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15237575" httpUri="https://zenodo.org/record/15237575/files/figure.png" pageId="9" pageNumber="224">
Fig. 1
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</figureCitation>
,
<figureCitation id="13712A07157BFF9DFE01FAC4FE412965" box="[392,459,1317,1344]" captionStart="Fig" captionStartId="8.[166,203,1270,1294]" captionTargetBox="[166,1413,151,1244]" captionTargetId="figure-258@8.[166,1413,151,1244]" captionTargetPageId="8" captionText="Fig. 5. Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, frontodorsal view. B Hind leg, lateral view. C Fragment of right forewing of female holotype, with M-Cu crossvein. D Incomplete left forewing of female holotype lacking M-Cu crossvein. E Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237583" httpUri="https://zenodo.org/record/15237583/files/figure.png" pageId="9" pageNumber="224">
5
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,
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) oval, widest in the middle; membranous; vein C+Sc moderately, evenly curved, slender, distinctly delimited to cell; cell c+sc relatively wide; costal break developed, close to apex of vein R
<subScript id="17CE34C7157BFF9DFF71FA5CFE8A29EE" attach="left" box="[248,256,1469,1483]" fontSize="6" pageId="9" pageNumber="224">1</subScript>
; pterostigma moderately wide, with subparallel margin in the median third, not expanded in apical third; entirely membranous; nodal line developed; vein R slightly shorter than M+Cu; vein Rs mostly straight, weakly curved to costal margin subapically; vein M much longer than M
<subScript id="17CE34C7157BFF9DFE73F985FD982A57" attach="left" box="[506,530,1636,1650]" fontSize="6" pageId="9" pageNumber="224">1+2</subScript>
; vein Cu
<subScript id="17CE34C7157BFF9DFDF6F985FD042A57" attach="left" box="[639,654,1636,1650]" fontSize="6" pageId="9" pageNumber="224">1a</subScript>
straight at base, curved towards anal margin subapically; in one wing a crossvein m-cu
<subScript id="17CE34C7157BFF9DFE26F947FE342A91" attach="left" box="[431,446,1702,1716]" fontSize="6" pageId="9" pageNumber="224">1a</subScript>
is developed (
<figureCitation id="13712A07157BFF9DFDEAF975FD332A8A" box="[611,697,1684,1711]" captionStart="Fig" captionStartId="8.[166,203,1270,1294]" captionTargetBox="[166,1413,151,1244]" captionTargetId="figure-258@8.[166,1413,151,1244]" captionTargetPageId="8" captionText="Fig. 5. Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, frontodorsal view. B Hind leg, lateral view. C Fragment of right forewing of female holotype, with M-Cu crossvein. D Incomplete left forewing of female holotype lacking M-Cu crossvein. E Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237583" httpUri="https://zenodo.org/record/15237583/files/figure.png" pageId="9" pageNumber="224">
Fig. 5
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</figureCitation>
); veins M
<subScript id="17CE34C7157BFF9DFF4FF929FF552AF3" attach="left" box="[198,223,1736,1750]" fontSize="6" pageId="9" pageNumber="224">1+2</subScript>
and M
<subScript id="17CE34C7157BFF9DFEBFF929FEDA2AF3" attach="left" box="[310,336,1736,1750]" fontSize="6" pageId="9" pageNumber="224">3+4</subScript>
oblique to wing margin apically; cell cu
<subScript id="17CE34C7157BFF9DFF40F908FF5B2AD2" attach="left" box="[201,209,1769,1783]" fontSize="6" pageId="9" pageNumber="224">2</subScript>
moderately long; anal break adjacent to apex of vein Cu
<subScript id="17CE34C7157BFF9DFE8DF8EBFE9E2B3D" attach="left" box="[260,276,1802,1816]" fontSize="6" pageId="9" pageNumber="224">1b</subScript>
; surface spinules not visible. Male unknown. Female terminalia (
<figureCitation id="13712A07157BFF9DFE00F8F8FE502B11" box="[393,474,1817,1844]" captionStart="Fig" captionStartId="8.[166,203,1270,1294]" captionTargetBox="[166,1413,151,1244]" captionTargetId="figure-258@8.[166,1413,151,1244]" captionTargetPageId="8" captionText="Fig. 5. Melanastera vetus BURCKHARDT &amp; DROHOJOWSKA sp. nov. A Head, frontodorsal view. B Hind leg, lateral view. C Fragment of right forewing of female holotype, with M-Cu crossvein. D Incomplete left forewing of female holotype lacking M-Cu crossvein. E Female terminalia, lateral view. Scales = 0.5 mm. Drawings by A. CORAY." figureDoi="http://doi.org/10.5281/zenodo.15237583" httpUri="https://zenodo.org/record/15237583/files/figure.png" pageId="9" pageNumber="224">
Fig. 5
<collectionCode id="ED5BAE47157BFF9DFE41F8F8FE502B11" box="[456,474,1817,1844]" pageId="9" pageNumber="224">E</collectionCode>
</figureCitation>
) cuneate; proctiger weakly concave dorsally, apex subacute, slightly upturned; beset with long setae across the middle and shorter setae apically; subgenital plate almost straight ventrally, acute apically.
</paragraph>
<paragraph id="8BF53682157BFF9DFCA6FEFDFBF82D5C" blockId="9.[815,1421,284,377]" pageId="9" pageNumber="224">
<emphasis id="B93EEA90157BFF9DFCA6FEFDFBDC2D13" bold="true" box="[815,1110,284,310]" pageId="9" pageNumber="224">Measurements (in mm):</emphasis>
Head width 0.73; antenna length 0.8; forewing length 1.6; metatibia length 0.5; female proctiger length 0.43.
</paragraph>
</subSubSection>
<subSubSection id="C3506509157BFF9DFCA6FE40FACC2F49" pageId="9" pageNumber="224" type="discussion">
<paragraph id="8BF53682157BFF9DFCA6FE40FACC2F49" blockId="9.[814,1422,417,877]" pageId="9" pageNumber="224">
<emphasis id="B93EEA90157BFF9DFCA6FE40FC3F2D9E" bold="true" box="[815,949,417,443]" pageId="9" pageNumber="224">Comments:</emphasis>
<taxonomicName id="4C4A4D01157BFF9DFC49FE40FB042D9E" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[960,1166,417,443]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="224" phylum="Arthropoda" rank="species" species="vetus">
<emphasis id="B93EEA90157BFF9DFC49FE40FB042D9E" box="[960,1166,417,443]" italics="true" pageId="9" pageNumber="224">Melanastera vetus</emphasis>
</taxonomicName>
shares with
<taxonomicName id="4C4A4D01157BFF9DFAACFE43FA072D9E" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[1317,1421,418,443]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="224" phylum="Arthropoda" rank="species" species="casca">
<emphasis id="B93EEA90157BFF9DFAACFE43FA072D9E" box="[1317,1421,418,443]" italics="true" pageId="9" pageNumber="224">M. casca</emphasis>
</taxonomicName>
and four undescribed Brazilian species associated with
<taxonomicName id="4C4A4D01157BFF9DFCA7FE05FB1F2DDB" authority="(SERBINA et al. 2024)" baseAuthorityName="SERBINA" baseAuthorityYear="2024" box="[814,1173,484,510]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="224" phylum="Tracheophyta" rank="family">
Asteraceae (
<bibRefCitation id="EFDB4B73157BFF9DFC3EFE05FB062DDB" author="SERBINA, L. S. &amp; MALENOVSKY, I. &amp; QUEIROZ, D. L. &amp; BURCKHARDT, D." box="[951,1164,484,510]" journalOrPublisher="Zootaxa" pageId="9" pageNumber="224" refId="ref7781" refString="SERBINA, L. S., MALENOVSKY, I., QUEIROZ, D. L. &amp; BURCKHARDT, D. (2024): Jumping plant-lice of the tribe Paurocephalini (Hemiptera: Psylloidea: Liviidae) in Brazil. - Zootaxa, submitted." title="Jumping plant-lice of the tribe Paurocephalini (Hemiptera: Psylloidea: Liviidae) in Brazil" type="book" year="2024">SERBINA et al. 2024</bibRefCitation>
)
</taxonomicName>
the pterostigma of the forewing with subparallel margins (see also comments under
<taxonomicName id="4C4A4D01157BFF9DFCFBFDC6FC582E65" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[882,978,551,576]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="224" phylum="Arthropoda" rank="species" species="casca">
<emphasis id="B93EEA90157BFF9DFCFBFDC6FC582E65" box="[882,978,551,576]" italics="true" pageId="9" pageNumber="224">M. casca</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="4C4A4D01157BFF9DFB0AFDC6FB6E2E65" authorityName="BURCKHARDT &amp; DROHOJOWSKA" authorityYear="2024" box="[1155,1252,551,576]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="224" phylum="Arthropoda" rank="species" species="casca">
<emphasis id="B93EEA90157BFF9DFB0AFDC6FB6E2E65" box="[1155,1252,551,576]" italics="true" pageId="9" pageNumber="224">M. casca</emphasis>
</taxonomicName>
in the forewing with dark dots at the apices of the veins along the margin and a shorter cell cu
<subScript id="17CE34C7157BFF9DFBB2FD9DFBC92EAF" attach="left" box="[1083,1091,636,650]" fontSize="6" pageId="9" pageNumber="224">1</subScript>
, as well as in the longer, dorsally concave female terminalia. From the extant species,
<taxonomicName id="4C4A4D01157BFF9DFCE1FD4CFC4D2EE3" authorityName="Burckhardt &amp; Drohojowska &amp; Štarhová Serbina &amp; Malenovský" authorityYear="2024" box="[872,967,685,710]" class="Insecta" family="Liviidae" genus="Melanastera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="224" phylum="Arthropoda" rank="species" species="vetus" status="sp. nov.">
<emphasis id="B93EEA90157BFF9DFCE1FD4CFC4D2EE3" box="[872,967,685,710]" italics="true" pageId="9" pageNumber="224">M. vetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57EB157BFF9DFC46FD4DFBAE2EE3" box="[975,1060,684,710]" pageId="9" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
differs in the broader forewings (2.4 versus ≥ 2.5) with dark dots, which are restricted to the apices of the veins along the margin but are otherwise absent from the cells. In the extant species associated with
<taxonomicName id="4C4A4D01157BFF9DFC10FCD0FB9F2F6E" box="[921,1045,817,843]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="224" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
, the dark dots on the forewings are either completely absent or present on most cells.
</paragraph>
</subSubSection>
</treatment>
</document>