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<mods:title id="24B3E3FA392C0E3E8AD82F1CC2F577B1">New and little-known Lepturinae from succinite (Coleoptera Cerambycidae)</mods:title>
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<taxonomicName id="4C294D716861B61CFDA5FE8BC5922D6F" ID-CoL="4CRC8" authority="Vitali, 2005" authorityName="Vitali" authorityYear="2005" box="[624,1007,331,355]" class="Insecta" family="Cerambycidae" genus="Paracorymbia" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="antiqua">
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<emphasis id="B95DEAE06861B61CFDA5FE8BC5102D6F" bold="true" box="[624,877,331,355]" italics="true" pageId="1">Paracorymbia antiqua</emphasis>
<bibRefCitation id="EFB84B036861B61CFCB8FE8BC5922D6F" author="Vitali F." box="[877,1007,331,355]" pageId="1" pagination="530 - 538" refId="ref5280" refString="Vitali F. 2005. Notes about the European fossil Lepturinae and the description of a new species (Coleoptera, Cerambycidae, Lepturinae). Lambillionea, CV (4): 530 - 538." type="journal article" year="2005">Vitali, 2005</bibRefCitation>
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(
<figureCitation id="13122A776861B61CFDADFEAAC4CB2D8E" box="[632,694,362,386]" captionStart="Fig" captionStartId="1.[114,153,1138,1162]" captionTargetBox="[114,1096,447,1101]" captionTargetId="figure-56@1.[114,1096,447,1101]" captionTargetPageId="1" captionText="Fig. 1.Paracorymbia (Vadoniocorymbia) antiquaVitali, 2005, Holotype, detail of the low- er margin of the eye." pageId="1">Fig. 1</figureCitation>
)
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Fig. 1.
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<emphasis id="B95DEAE06861B61CFF6AFBB2C4ED2886" box="[191,656,1138,1162]" italics="true" pageId="1">Paracorymbia (Vadoniocorymbia) antiqua</emphasis>
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, Holotype, detail of the low- er margin of the eye.
</paragraph>
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<emphasis id="B95DEAE06861B61CFFA7FB0EC71728EA" bold="true" box="[114,362,1230,1254]" pageId="1">Examined material.</emphasis>
<typeStatus id="549288506861B61CFEAFFB0EC79D28EA" box="[378,480,1230,1254]" pageId="1" type="holotype">Holotype</typeStatus>
, female, Baltic amber, ex coll. C. Gröhn, authors coll. FS3BS3 (
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); female, Baltic amber, ex coll. M. Veta; authors coll. FS73BS42.
</paragraph>
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<emphasis id="B95DEAE06861B61CFFA7FA46C7182992" bold="true" box="[114,357,1414,1438]" pageId="1">Differential diagnosis</emphasis>
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<paragraph id="8B9636F26861B61CFFA7FA65C42D2A79" blockId="1.[114,593,1414,1653]" pageId="1">
As remarked in the original description (Vitali 2015),
<taxonomicName id="4C294D716861B61CFF2EFA04C78B29D7" authorityName="Vitali" authorityYear="2005" box="[251,502,1476,1500]" class="Insecta" family="Cerambycidae" genus="Paracorymbia" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="B95DEAE06861B61CFF2EFA04C78B29D7" box="[251,502,1476,1500]" italics="true" pageId="1">Paracorymbia antiqua</emphasis>
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shows pronotum with apical furrow and truncate elytral apex like
<taxonomicName id="4C294D716861B61CFEF1F9C1C7C32A15" authorityName="Miroshnikov" authorityYear="1998" box="[292,446,1537,1561]" class="Insecta" family="Cerambycidae" genus="Paracorymbia" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE06861B61CFEF1F9C1C7C32A15" box="[292,446,1537,1561]" italics="true" pageId="1">Paracorymbia</emphasis>
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. However, it differs from all congeners of the Recent in the convex under margin of the eyes (concave in the extant species). This archaic
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190 character is almost present in the extant species of the genus
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<emphasis id="B95DEAE06861B61CFCA8FB2DC5AA2909" box="[893,983,1261,1285]" italics="true" pageId="1">Vadonia</emphasis>
Mulsant, 1863
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, which however show rounded elytral apex and pronotum without apical collar.
</paragraph>
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Female genitalia (exposed in the
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) are analogue to the ones of the extant
<taxonomicName id="4C294D716861B61CFBF7FA47C4D029B1" authorityName="Mulsant" authorityYear="1863" class="Insecta" family="Cerambycidae" genus="Vadonia" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE06861B61CFBF7FA47C4D029B1" italics="true" pageId="1">Vadonia</emphasis>
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and
<taxonomicName id="4C294D716861B61CFD34FA65C50629B1" authorityName="Miroshnikov" authorityYear="1998" box="[737,891,1445,1469]" class="Insecta" family="Cerambycidae" genus="Paracorymbia" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE06861B61CFD34FA65C50629B1" box="[737,891,1445,1469]" italics="true" pageId="1">Paracorymbia</emphasis>
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.
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Due to its mixed characters,
<taxonomicName id="4C294D716861B61CFC72FA23C26B29F6" authorityName="Vitali" authorityYear="2005" box="[935,1046,1507,1530]" class="Insecta" family="Cerambycidae" genus="Paracorymbia" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="B95DEAE06861B61CFC72FA23C26B29F6" box="[935,1046,1507,1530]" italics="true" pageId="1">P. antiqua</emphasis>
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was interpreted as a quite archaic member of
<taxonomicName id="4C294D716861B61CFDA5F9E0C5772A34" authorityName="Miroshnikov" authorityYear="1998" box="[624,778,1568,1592]" class="Insecta" family="Cerambycidae" genus="Paracorymbia" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE06861B61CFDA5F9E0C5772A34" box="[624,778,1568,1592]" italics="true" pageId="1">Paracorymbia</emphasis>
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, close to the
<emphasis id="B95DEAE06861B61CFC61F9E0C2772A34" box="[948,1034,1568,1592]" italics="true" pageId="1">pallens-</emphasis>
group and as a probable link with
<emphasis id="B95DEAE06861B61CFC42F9FEC5842A5A" box="[919,1017,1598,1622]" italics="true" pageId="1">
<taxonomicName id="4C294D716861B61CFC42F9FEC5882A5A" authorityName="Mulsant" authorityYear="1863" box="[919,1013,1598,1622]" class="Insecta" family="Cerambycidae" genus="Vadonia" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">Vadonia</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B9636F26862B61FFF68FF72C7CF2D48" blockId="2.[189,668,178,324]" pageId="2">
However, considering that this species can be inserted in neither extant subgenera (
<bibRefCitation id="EFB84B036862B61FFF11FF2FC7E22D0B" author="Miroshnikov A. I." box="[196,415,239,263]" pageId="2" pagination="384 - 420" refId="ref5000" refString="Miroshnikov A. I. 1998 a. A new classification of longicorn beetles of the Anoplodera complex, Tribe Lepturini (Coleoptera, Cerambycidae) of Holoarctic fauna, I. Entomologicescoe Obozrenie, LXXVII (2): 384 - 420." type="journal article" year="1998">Miroshnikov 1998a</bibRefCitation>
, b),
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<emphasis id="B95DEAE06862B61FFE02FF2FC4EB2D0B" box="[471,662,239,263]" italics="true" pageId="2">Vadoniocorymbia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6862B61FFF68FECEC7532D29" box="[189,302,269,293]" pageId="2" rank="subGenus">n. subgen.</taxonomicNameLabel>
is here introduced to accommodate this fossil species.
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</subSubSection>
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<paragraph id="8B9636F26863B61EFDA5FD9FC49D2E7B" blockId="3.[624,1087,576,631]" box="[624,736,607,631]" pageId="3">
(
<figureCitation id="13122A776863B61EFDADFD9FC4A52E7B" box="[632,728,607,631]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[624,663,1408,1432]" captionStartId-1="4.[189,228,828,852]" captionStartId-2="4.[699,738,889,913]" captionTargetBox-0="[624,1098,661,1371]" captionTargetBox-1="[189,1042,172,791]" captionTargetBox-2="[204,523,910,1590]" captionTargetId-0="figure-180@3.[624,1098,661,1371]" captionTargetId-1="figure-11@4.[189,1042,171,791]" captionTargetId-2="figure-21@4.[189,538,882,1601]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="4" captionText-0="Fig. 4. Pedostrangalia ostensackeni n. sp., Holotype, lateral view." captionText-1="Fig. 5. Pedostrangalia ostensackenin. sp., Holotype, dorsal view." captionText-2="Fig. 6. Pedostrangalia ostensackeni n. sp., Holotype, reconstruction." pageId="3">Figs. 4-6</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF56667A6864B619FF68FCFCC5002F58" box="[189,893,828,852]" pageId="4" startId="4.[189,228,828,852]" targetBox="[189,1042,172,791]" targetPageId="4" targetType="figure">
<paragraph id="8B9636F26864B619FF68FCFCC5002F58" blockId="4.[189,893,828,852]" box="[189,893,828,852]" pageId="4">
Fig. 5.
<taxonomicName id="4C294D716864B619FEDCFCFCC43E2F58" authorityName="Vitali" authorityYear="2023" box="[265,579,828,852]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="ostensackeni" status="sp. nov.">
<emphasis id="B95DEAE06864B619FEDCFCFCC43E2F58" box="[265,579,828,852]" italics="true" pageId="4">Pedostrangalia ostensackeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6864B619FD96FCFCC4032F5F" box="[579,638,828,851]" pageId="4" rank="species">n. sp.</taxonomicNameLabel>
, Holotype, dorsal view.
</paragraph>
</caption>
<caption id="DF56667A6864B619FD6EFCB9C5B12FBC" pageId="4" startId="4.[699,738,889,913]" targetBox="[204,523,910,1590]" targetPageId="4" targetType="figure">
<paragraph id="8B9636F26864B619FD6EFCB9C5B12FBC" blockId="4.[699,1178,889,944]" pageId="4">
Fig. 6.
<taxonomicName id="4C294D716864B619FCDAFCB9C2312F9D" authorityName="Vitali" authorityYear="2023" box="[783,1100,889,913]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="ostensackeni" status="sp. nov.">
<emphasis id="B95DEAE06864B619FCDAFCB9C2312F9D" box="[783,1100,889,913]" italics="true" pageId="4">Pedostrangalia ostensackeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6864B619FB85FCBAC2E82F9D" box="[1104,1173,890,913]" pageId="4" rank="species">n. sp.</taxonomicNameLabel>
, Holotype, reconstruction.
</paragraph>
</caption>
<subSubSection id="C33365796864B619FD6EFC15C5772970" pageId="4" type="materials_examined">
<paragraph id="8B9636F26864B619FD6EFC15C4922826" blockId="4.[699,1178,981,1066]" pageId="4">
<emphasis id="B95DEAE06864B619FD6EFC15C5572FE1" bold="true" box="[699,810,981,1005]" pageId="4">
<typeStatus id="549288506864B619FD6EFC15C5582FE1" box="[699,805,981,1005]" pageId="4" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male, Baltic amber, ex coll. M. Veta 210415, authors coll. FS83BS51 (
<figureCitation id="13122A776864B619FBBFFC34C49F2826" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[624,663,1408,1432]" captionStartId-1="4.[189,228,828,852]" captionTargetBox-0="[624,1098,661,1371]" captionTargetBox-1="[189,1042,172,791]" captionTargetId-0="figure-180@3.[624,1098,661,1371]" captionTargetId-1="figure-11@4.[189,1042,171,791]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 4. Pedostrangalia ostensackeni n. sp., Holotype, lateral view." captionText-1="Fig. 5. Pedostrangalia ostensackenin. sp., Holotype, dorsal view." pageId="4">Fig. 4-5</figureCitation>
).
</paragraph>
<paragraph id="8B9636F26864B619FD6EFB90C5772970" blockId="4.[699,1178,1104,1404]" pageId="4">
The beetle is preserved inside a circular piece of amber measuring ~
<quantity id="4CD19B176864B619FC33FBAFC24E288A" box="[998,1075,1135,1159]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" unit="mm" value="20.0">20 mm</quantity>
in diameter. It lacks antennomeres
<collectionCode id="ED38AE376864B619FC3DFB4EC24628A9" box="[1000,1083,1166,1189]" pageId="4">VIII-XI</collectionCode>
and the body looks to have been subject to a strong deformation of the amber. The humeri look laminiform, the coxae look flattened and pulled backwards, while the abdomen is almost dislocated backwards. The amber piece contains one stellate fagacean trichome.
</paragraph>
</subSubSection>
<subSubSection id="C33365796864B618FD6EFA61C7F328D0" lastPageId="5" pageId="4" type="diagnosis">
<paragraph id="8B9636F26864B619FD6EFA61C5D229B5" blockId="4.[699,1179,1441,1649]" box="[699,943,1441,1465]" pageId="4">
<emphasis id="B95DEAE06864B619FD6EFA61C5D229B5" bold="true" box="[699,943,1441,1465]" pageId="4">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="8B9636F26864B619FD6EFA00C2E72A7D" blockId="4.[699,1179,1441,1649]" pageId="4">
The fossil has been identified as a member of the genus
<taxonomicName id="4C294D716864B619FC9FFA1EC59329FA" authorityName="Sokolov" authorityYear="1897" box="[842,1006,1502,1526]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE06864B619FC9FFA1EC59329FA" box="[842,1006,1502,1526]" italics="true" pageId="4">Pedostrangalia</emphasis>
</taxonomicName>
(s. str.) for the following combination of characters: prosternum convex, not forming an angle with the intercoxal process (
<taxonomicName id="4C294D716864B619FC1DF9FAC24F2A5E" authorityName="Latreille" authorityYear="1802" box="[968,1074,1594,1618]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="tribe" tribe="Lepturini">Lepturini</taxonomicName>
); cheeks developed: head with distinct angulated
</paragraph>
<paragraph id="8B9636F26864B618FBB9F966C6962D6F" blockId="4.[1132,1172,1702,1726]" lastBlockId="5.[114,593,178,355]" lastPageId="5" pageId="4">193 temples; pronotum without apical collar and with acute basal angles; elytra not shortened, posteriorly convergent and not round- ed at apex; antennae long and not strongly dentate. In addition, the pronotum is laterally winkled.</paragraph>
<paragraph id="8B9636F26865B618FFA7FE48C6E82DF0" blockId="5.[114,592,392,508]" pageId="5">The peculiar character of the furrowed tarsi in not visible, but other genera show either strongly reduced temples or a pronotal collar.</paragraph>
<paragraph id="8B9636F26865B618FFA7FDE2C6972F1C" blockId="5.[114,593,546,784]" pageId="5">
With respect to
<taxonomicName id="4C294D716865B618FEF4FDE2C7EE2E35" authorityName="Vitali" authorityYear="2014" box="[289,403,546,569]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="pristina">
<emphasis id="B95DEAE06865B618FEF4FDE2C7EE2E35" box="[289,403,546,569]" italics="true" pageId="5">P. pristina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C294D716865B618FE76FDE2C42C2E36" authorityName="Vitali" authorityYear="2023" box="[419,593,546,570]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="ostensackeni" status="n. sp.">
<emphasis id="B95DEAE06865B618FE76FDE2C42C2E36" box="[419,593,546,570]" italics="true" pageId="5">P. ostensackeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6865B618FFA7FD81C6D12E54" box="[114,172,577,600]" pageId="5" rank="species">n. sp.</taxonomicNameLabel>
shows coloration entirely black, habitus more elongated, pronotum more cylindrical and with lateral angles, elytra 3.3 as long as wide at shoulders (
<quantity id="4CD19B176865B618FE48FD5CC7982EB8" box="[413,485,668,692]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" pageId="5" unit="in" value="2.6">2.6 in</quantity>
<taxonomicName id="4C294D716865B618FE3CFD5DC6F02EDF" authorityName="Vitali" authorityYear="2014" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="pristina">
<emphasis id="B95DEAE06865B618FE3CFD5DC6F02EDF" italics="true" pageId="5">P. pristina</emphasis>
</taxonomicName>
) and more slender antennae, whose last antennomere is pedunculate or sub-articulated.
</paragraph>
<paragraph id="8B9636F26865B618FFA7FCF5C7F328D0" blockId="5.[114,593,821,1244]" pageId="5">
With respect to the congeners of the Recent,
<taxonomicName id="4C294D716865B618FF7AFC95C7222F60" authorityName="Vitali" authorityYear="2023" box="[175,351,852,876]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="ostensackeni" status="sp. n.">
<emphasis id="B95DEAE06865B618FF7AFC95C7222F60" box="[175,351,852,876]" italics="true" pageId="5">P. ostensackeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6865B618FEB7FC95C7E22F60" box="[354,415,853,876]" pageId="5" rank="species">sp. n.</taxonomicNameLabel>
seems to show most affinity with the Caucasian
<taxonomicName id="4C294D716865B618FE00FCB4C7632FA6" authority="Sama, 1996" authorityName="Sama" authorityYear="1996" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="tokatensis">
<emphasis id="B95DEAE06865B618FE00FCB4C6F32FA5" italics="true" pageId="5">P. tokatensis</emphasis>
<bibRefCitation id="EFB84B036865B618FF4CFC52C7632FA6" author="Sama G." box="[153,286,914,938]" pageId="5" pagination="101 - 116" refId="ref5137" refString="Sama G. 1996. Contribution a la connaissance des Longicornes de Grece et d'Asie Mineure (Coleoptera, Cerambycidae). Biocosme Mesogeen, 12 (4): 101 - 116." type="journal article" year="1996">Sama, 1996</bibRefCitation>
</taxonomicName>
, which is characterised by analogous habitus and an atypical furrow behind the posterior margin of the eyes (unnoticed in the original description). The fossil differs in the pronotum laterally toothed, the longer elytra spines and (possibly) the black body colour.
<taxonomicName id="4C294D716865B618FE76FB8AC42C286D" authorityName="Vitali" authorityYear="2023" box="[419,593,1097,1121]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="ostensackeni" status="sp. n.">
<emphasis id="B95DEAE06865B618FE76FB8AC42C286D" box="[419,593,1097,1121]" italics="true" pageId="5">P. ostensackeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6865B618FFA7FBA9C6D2288C" box="[114,175,1129,1152]" pageId="5" rank="species">sp. n.</taxonomicNameLabel>
reminds also of some
<taxonomicName id="4C294D716865B618FE64FBA8C439288C" authorityName="Lobl" authorityYear="2010" box="[433,580,1128,1152]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="subGenus" subGenus="Neosphenalia">
<emphasis id="B95DEAE06865B618FE64FBA8C439288C" box="[433,580,1128,1152]" italics="true" pageId="5">Neosphenalia</emphasis>
</taxonomicName>
, e.g., the Manchurian
<taxonomicName id="4C294D716865B618FE7EFB48C72B28B1" authority="(Motschulsky, 1860)" baseAuthorityName="Motschulsky" baseAuthorityYear="1860" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="B95DEAE06865B618FE7EFB48C4372893" box="[427,586,1159,1183]" italics="true" pageId="5">P. femoralis</emphasis>
(Motschulsky, 1860)
</taxonomicName>
, which especially differs in the shorter temples.
</paragraph>
</subSubSection>
<subSubSection id="C33365796865B61BFFA7FAC8C4912CE4" lastPageId="6" pageId="5" type="description">
<paragraph id="8B9636F26865B618FFA7FAC8C68B292C" blockId="5.[114,246,1288,1312]" box="[114,246,1288,1312]" pageId="5">
<emphasis id="B95DEAE06865B618FFA7FAC8C68B292C" bold="true" box="[114,246,1288,1312]" pageId="5">Description</emphasis>
</paragraph>
<paragraph id="8B9636F26865B618FFA7FA85C7D829B5" blockId="5.[114,593,1349,1465]" pageId="5">
Undetermined sex, probably male; length
<quantity id="4CD19B176865B618FFA7FAA4C6B62970" box="[114,203,1380,1404]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="5" unit="mm" value="7.5">7.5 mm</quantity>
. General habitus small, elongated; body seemingly black, ventral side covered with dense light pubescence.
</paragraph>
<paragraph id="8B9636F26865B618FFA7FA1FC42C2A7D" blockId="5.[114,593,1503,1649]" pageId="5">Head relatively short; cheeks developed but relatively short; clypeus and forehead transverse; antennal tubercles widely separated, fairly elevated; eyes relatively close to the base of the mandibles, emarginate at</paragraph>
<paragraph id="8B9636F26865B618FFA7F966C5C82F87" blockId="5.[114,154,1702,1726]" lastBlockId="5.[624,1103,178,907]" pageId="5">194 the upper side, uniformly convex at the un- der one, finely faceted; temples relatively long, inferiorly as long as cheeks, parallel-sided, abruptly converging backward, transversely furrowed behind the posterior margin of the eyes; neck as long as temples. Last (IV) maxillar palpomere widened and transversely truncate at apex; III palpomere widened at apex, three-fourth as long as IV; II palpomere elongated, as long as IV. Antennae 11-segmented, last antennomere seemingly peduculate or sub-articulated, inserted between the eyes, hardly reaching the elytral apex, glabrous, extremely finely and densely punctured; scape feebly bowed; pedicle twice longer than broad, nearly one-fourth as long as scape; antennomere III more than one-fourth longer than scape; antennomere IV hardly longer than scape; antennomere V as long as III; following antennomeres decreasing in length, except for the last, about as long as VII; proportions according to the formula: 3.8: 1.0: 5.0: 4.0: 5.0: 3.8: 3.2: 2.8: 2.8: 2.7: 3.1.</paragraph>
<paragraph id="8B9636F26865B618FDA5FC70C4B428F7" blockId="5.[624,1103,944,1275]" pageId="5">Prothorax transverse, bell-shaped; sides with a stout angle before the middle; hind angles acute, embracing the elytral base; apex concave, deeply furrowed and bordered; base posteriorly lobed in the middle, finely grooved; disc feebly convex above, without longitudinal furrow, entirely covered with an extremely fine dense puncturing and indistinct pubescence. Scutellum small, elongate, forming an isosceles triangle.</paragraph>
<paragraph id="8B9636F26865B618FDA5FAE0C4C92A40" blockId="5.[624,1103,1312,1612]" pageId="5">Elytra long, 3.7 times as long as pronotum, 3.3 times as long as wide at the shoulders, flat above, feebly constricted at the sides after humeri, then nearly parallel-sided, apically tapered shortly before apex and obliquely emarginated at the apex; marginal apex acute; surface covered with a coarsely dense puncturing and an extremely fine, nearly invisible, short recumbent pubescence.</paragraph>
<paragraph id="8B9636F26866B61BFF68FF72C6852D29" blockId="6.[189,669,178,293]" pageId="6">Ventral side convex, apparently impunctate and covered with a dense recumbent pubescence; prosternum nearly straight in lateral view.</paragraph>
<paragraph id="8B9636F26866B61BFF68FE8BC4912CE4" blockId="6.[189,668,331,754]" lastBlockId="6.[699,1178,177,232]" pageId="6">Legs long; femora slightly club-shaped; tibiae linear, rectilinearly truncated at the apex, finely punctured and pubescent; apex of mesotibiae armed with two equal spines; apex of metatibiae with two unequal spines, the longest one being one-third as long as metatarsomere I. Metatarsi narrow, long, about one-fifth shorter than the metatibiae; metatasomere I about as long as the following articles together; metatarsomere II one-half longer than III; metatarsomere III one-fourth as long as I, incised at the apex more than one-half of its length; metatarsomere IV minute; onychium as long as II (proportions according to the formula: 4: 1.5: 1:?: 1.5).</paragraph>
</subSubSection>
<subSubSection id="C33365796866B61BFD6EFECDC2F22DD1" pageId="6" type="etymology">
<paragraph id="8B9636F26866B61BFD6EFECDC54B2D29" blockId="6.[699,822,269,293]" box="[699,822,269,293]" pageId="6">
<emphasis id="B95DEAE06866B61BFD6EFECDC54B2D29" bold="true" box="[699,822,269,293]" pageId="6">Etymology</emphasis>
</paragraph>
<paragraph id="8B9636F26866B61BFD6EFE8BC2F22DD1" blockId="6.[699,1178,331,478]" pageId="6">
This species is dedicated to Carl-Robert Osten-Sacken (1828-1906), eminent dipterologist of Baltic German origin and Russian consul in the
<collectingCountry id="F33E76626866B61BFC8EFE67C5932DB3" box="[859,1006,423,447]" name="United States of America" pageId="6">United States</collectingCountry>
for reminding the strength links among European peoples.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="038087E46866B617FD6EFDE8C58B2A21" LSID="urn:lsid:plazi:treatment:038087E46866B617FD6EFDE8C58B2A21" httpUri="http://treatment.plazi.org/id/038087E46866B617FD6EFDE8C58B2A21" lastPageId="10" pageId="6">
<subSubSection id="C33365796866B61BFD6EFDE8C2102E4C" box="[699,1133,552,576]" pageId="6" type="nomenclature">
<paragraph id="8B9636F26866B61BFD6EFDE8C2102E4C" blockId="6.[699,1133,552,607]" box="[699,1133,552,576]" pageId="6">
<heading id="D0DE819E6866B61BFD6EFDE8C2102E4C" bold="true" box="[699,1133,552,576]" fontSize="10" level="2" pageId="6" reason="4">
<emphasis id="B95DEAE06866B61BFD6EFDE8C2102E4C" bold="true" box="[699,1133,552,576]" pageId="6">
<taxonomicName id="4C294D716866B61BFD6EFDE8C2592E4C" authorityName="Vitali" authorityYear="2023" box="[699,1060,552,576]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. nov.">
<emphasis id="B95DEAE06866B61BFD6EFDE8C2592E4C" bold="true" box="[699,1060,552,576]" italics="true" pageId="6">Pedostrangalia (s. str.) rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6866B61BFBFEFDE8C2102E4C" box="[1067,1133,552,576]" pageId="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33365796866B61BFD6EFD86C54F2E53" box="[699,818,582,607]" pageId="6" type="description">
<paragraph id="8B9636F26866B61BFD6EFD86C54F2E53" blockId="6.[699,1133,552,607]" box="[699,818,582,607]" pageId="6">
(
<figureCitation id="13122A776866B61BFD16FD86C5572E53" box="[707,810,582,607]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="6.[189,228,1404,1428]" captionStartId-1="7.[114,153,822,846]" captionStartId-2="7.[114,153,1528,1552]" captionStartId-3="8.[189,228,822,846]" captionStartId-4="8.[189,228,1528,1552]" captionStartId-5="9.[114,153,1024,1048]" captionTargetBox-0="[189,1171,754,1368]" captionTargetBox-1="[114,1096,171,785]" captionTargetBox-2="[114,1096,876,1490]" captionTargetBox-3="[189,1171,171,785]" captionTargetBox-4="[189,1171,876,1490]" captionTargetBox-5="[129,572,188,969]" captionTargetId-0="figure-190@6.[189,1171,754,1368]" captionTargetId-1="figure-4@7.[114,1096,171,785]" captionTargetId-2="figure-15@7.[114,1096,876,1490]" captionTargetId-3="figure-11@8.[189,1171,171,785]" captionTargetId-4="figure-22@8.[189,1171,876,1490]" captionTargetId-5="figure-4@9.[114,579,171,986]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="Fig. 7.Pedostrangalia rovnensisn. sp., male, Holotype, dorsal view." captionText-1="Fig. 8.Pedostrangalia rovnensisn. sp., male, Holotype, lateral view." captionText-2="Fig. 9. Pedostrangalia rovnensisn. sp., male, Holotype, ventral view." captionText-3="Fig. 10.Pedostrangalia rovnensisn. sp., female, Paratype, anterior view." captionText-4="Fig. 11.Pedostrangalia rovnensisn. sp., female, Paratype, posterior view." captionText-5="Fig. 12.Pedostrangalia rovnensisn. sp. reconstruction." pageId="6">Figs 7-12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33365796866B614FD6EFD43C6E72AB2" lastPageId="9" pageId="6" type="materials_examined">
<paragraph id="8B9636F26866B61BFD6EFD43C5DA2ED5" blockId="6.[699,1178,643,729]" pageId="6">
<materialsCitation id="3B413CAF6866B61BFD6EFD43C5DA2ED5" collectionCode="R" collectorName="J. Damzen" location="Rovno" pageId="6" specimenCount="1" stateProvince="Rivne" typeStatus="holotype">
<emphasis id="B95DEAE06866B61BFD6EFD43C5572E97" bold="true" box="[699,810,643,667]" pageId="6">
<typeStatus id="549288506866B61BFD6EFD43C5582E97" box="[699,805,643,667]" pageId="6" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male,
<collectingRegion id="49EDF8106866B61BFCABFD44C5B82E97" box="[894,965,644,667]" country="Ukraine" name="Rivne" pageId="6">Rovno</collectingRegion>
amber, ex coll.
<collectorName id="26DC53246866B61BFB56FD44C5692EB6" pageId="6">J. Damzen</collectorName>
JDC-10269
<collectionCode id="ED38AE376866B61BFC66FD62C5B82EB6" box="[947,965,674,698]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6">R</collectionCode>
, authors coll. FS86BS54 (
<figureCitation id="13122A776866B61BFCE8FD01C5E42ED5" box="[829,921,705,729]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[189,228,1404,1428]" captionStartId-1="7.[114,153,822,846]" captionStartId-2="7.[114,153,1528,1552]" captionTargetBox-0="[189,1171,754,1368]" captionTargetBox-1="[114,1096,171,785]" captionTargetBox-2="[114,1096,876,1490]" captionTargetId-0="figure-190@6.[189,1171,754,1368]" captionTargetId-1="figure-4@7.[114,1096,171,785]" captionTargetId-2="figure-15@7.[114,1096,876,1490]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionText-0="Fig. 7.Pedostrangalia rovnensisn. sp., male, Holotype, dorsal view." captionText-1="Fig. 8.Pedostrangalia rovnensisn. sp., male, Holotype, lateral view." captionText-2="Fig. 9. Pedostrangalia rovnensisn. sp., male, Holotype, ventral view." pageId="6">Figs 7-9</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="DF56667A6866B61BFF68FABCC5E12998" box="[189,924,1404,1429]" pageId="6" startId="6.[189,228,1404,1428]" targetBox="[189,1171,754,1368]" targetPageId="6" targetType="figure">
<paragraph id="8B9636F26866B61BFF68FABCC5E12998" blockId="6.[189,924,1404,1429]" box="[189,924,1404,1429]" pageId="6">
Fig. 7.
<taxonomicName id="4C294D716866B61BFEDCFABDC45D2998" authorityName="Vitali" authorityYear="2023" box="[265,544,1405,1429]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. nov.">
<emphasis id="B95DEAE06866B61BFEDCFABDC45D2998" box="[265,544,1405,1429]" italics="true" pageId="6">Pedostrangalia rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6866B61BFDF5FABDC4212998" box="[544,604,1405,1428]" pageId="6" rank="species">n. sp.</taxonomicNameLabel>
, male, Holotype, dorsal view.
</paragraph>
</caption>
<paragraph id="8B9636F26866B61BFBB9F966C2E92AB2" blockId="6.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="6">195</paragraph>
<caption id="DF56667A6867B61AFFA7FCF6C52F2F42" box="[114,850,822,846]" pageId="7" startId="7.[114,153,822,846]" targetBox="[114,1096,171,785]" targetPageId="7" targetType="figure">
<paragraph id="8B9636F26867B61AFFA7FCF6C52F2F42" blockId="7.[114,850,822,846]" box="[114,850,822,846]" pageId="7">
Fig. 8.
<taxonomicName id="4C294D716867B61AFF68FCF6C7A82F42" authorityName="Vitali" authorityYear="2023" box="[189,469,822,846]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. nov.">
<emphasis id="B95DEAE06867B61AFF68FCF6C7A82F42" box="[189,469,822,846]" italics="true" pageId="7">Pedostrangalia rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6867B61AFE00FCF7C46D2F42" box="[469,528,823,846]" pageId="7" rank="species">n. sp.</taxonomicNameLabel>
, male, Holotype, lateral view.
</paragraph>
</caption>
<caption id="DF56667A6867B61AFFA7FA38C5242A1C" box="[114,857,1528,1552]" pageId="7" startId="7.[114,153,1528,1552]" targetBox="[114,1096,876,1490]" targetPageId="7" targetType="figure">
<paragraph id="8B9636F26867B61AFFA7FA38C5242A1C" blockId="7.[114,857,1528,1552]" box="[114,857,1528,1552]" pageId="7">
Fig. 9.
<taxonomicName id="4C294D716867B61AFF68FA38C7A82A03" authorityName="Vitali" authorityYear="2023" box="[189,469,1528,1552]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. nov.">
<emphasis id="B95DEAE06867B61AFF68FA38C7A82A03" box="[189,469,1528,1552]" italics="true" pageId="7">Pedostrangalia rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6867B61AFE00FA38C46D2A03" box="[469,528,1528,1551]" pageId="7" rank="species">n. sp.</taxonomicNameLabel>
, male, Holotype, ventral view.
</paragraph>
</caption>
<paragraph id="8B9636F26867B61AFFA7F966C6E72AB2" blockId="7.[114,154,1702,1726]" box="[114,154,1702,1726]" pageId="7">196</paragraph>
<caption id="DF56667A6868B615FF68FCF6C5B72F42" box="[189,970,822,847]" pageId="8" startId="8.[189,228,822,846]" targetBox="[189,1171,171,785]" targetPageId="8" targetType="figure">
<paragraph id="8B9636F26868B615FF68FCF6C5B72F42" blockId="8.[189,970,822,847]" box="[189,970,822,847]" pageId="8">
Fig. 10.
<taxonomicName id="4C294D716868B615FEC3FCF6C4532F42" authorityName="Vitali" authorityYear="2023" box="[278,558,822,846]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. nov.">
<emphasis id="B95DEAE06868B615FEC3FCF6C4532F42" box="[278,558,822,846]" italics="true" pageId="8">Pedostrangalia rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6868B615FDFBFCF7C4142F42" box="[558,617,823,846]" pageId="8" rank="species">n. sp.</taxonomicNameLabel>
, female, Paratype, anterior view.
</paragraph>
</caption>
<caption id="DF56667A6868B615FF68FA38C5AB2A1C" box="[189,982,1528,1552]" pageId="8" startId="8.[189,228,1528,1552]" targetBox="[189,1171,876,1490]" targetPageId="8" targetType="figure">
<paragraph id="8B9636F26868B615FF68FA38C5AB2A1C" blockId="8.[189,982,1528,1552]" box="[189,982,1528,1552]" pageId="8">
Fig. 11.
<taxonomicName id="4C294D716868B615FEC3FA38C4532A03" authorityName="Vitali" authorityYear="2023" box="[278,558,1528,1552]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. nov.">
<emphasis id="B95DEAE06868B615FEC3FA38C4532A03" box="[278,558,1528,1552]" italics="true" pageId="8">Pedostrangalia rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6868B615FDFBFA38C4142A03" box="[558,617,1528,1551]" pageId="8" rank="species">n. sp.</taxonomicNameLabel>
, female, Paratype, posterior view.
</paragraph>
</caption>
<paragraph id="8B9636F26868B614FBB9F966C5BF2D8E" blockId="8.[1132,1172,1702,1726]" lastBlockId="9.[624,1103,178,447]" lastPageId="9" pageId="8">
197 32x28x
<quantity id="4CD19B176869B614FD14FF72C5692CC5" box="[705,788,178,202]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" unit="mm" value="10.0">10 mm</quantity>
. The specimen, which lays folded on itself with an open elytron, is perfectly preserved but mostly covered with milky turbidity, except for the right eye, the first four articles of the right antenna and some parts of the legs. The extended ovipositor implies that it is female.
</paragraph>
<paragraph id="8B9636F26869B614FDA5FE48C5CC2DB3" blockId="9.[624,1103,178,447]" pageId="9">
The amber piece also contains a minute unidentified beetle,
<quantity id="4CD19B176869B614FCFFFE67C5062DB2" box="[810,891,423,447]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" unit="mm" value="1.2">1.2 mm</quantity>
long
</paragraph>
<caption id="DF56667A6869B614FFA7FBC0C680283A" pageId="9" startId="9.[114,153,1024,1048]" targetBox="[129,572,188,969]" targetPageId="9" targetType="figure">
<paragraph id="8B9636F26869B614FFA7FBC0C680283A" blockId="9.[114,586,1024,1078]" pageId="9">
Fig. 12.
<taxonomicName id="4C294D716869B614FF1BFBC0C79A281B" authorityName="Vitali" authorityYear="2023" box="[206,487,1024,1048]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. nov.">
<emphasis id="B95DEAE06869B614FF1BFBC0C79A281B" box="[206,487,1024,1048]" italics="true" pageId="9">Pedostrangalia rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6869B614FE3CFBC0C459281B" box="[489,548,1024,1047]" pageId="9" rank="species">n. sp.</taxonomicNameLabel>
reconstruction.
</paragraph>
</caption>
<paragraph id="8B9636F26869B614FFA7FB9CC6A52920" blockId="9.[114,593,1116,1324]" pageId="9">
The beetle is preserved inside a subtriangular piece of amber measuring about 48x28x
<quantity id="4CD19B176869B614FF16FB59C76C28BD" box="[195,273,1177,1201]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="9" unit="mm" value="12.0">12 mm</quantity>
. It is perfectly preserved, but the head and the right ventral side is covered with milky turbidity. The narrowly conical pygidium implies that this specimen is a male.
</paragraph>
<paragraph id="8B9636F26869B614FFA7FA91C69829AA" blockId="9.[114,593,1361,1446]" pageId="9">The amber piece contains numerous wood debris but seemingly, not further zoological inclusions.</paragraph>
<paragraph id="8B9636F26869B614FFA7FA0BC7052A2D" blockId="9.[114,592,1483,1569]" pageId="9">
<materialsCitation id="3B413CAF6869B614FFA7FA0BC7092A2D" collectionCode="R" collectorName="J. Damzen" location="Rovno" pageId="9" specimenCount="1" stateProvince="Rivne" typeStatus="paratype">
<emphasis id="B95DEAE06869B614FFA7FA0BC69C29EF" bold="true" box="[114,225,1483,1507]" pageId="9">
<typeStatus id="549288506869B614FFA7FA0BC6A129EF" box="[114,220,1483,1507]" pageId="9" type="paratype">Paratype</typeStatus>
.
</emphasis>
Female,
<collectingRegion id="49EDF8106869B614FE91FA0CC7F629EF" box="[324,395,1484,1507]" country="Ukraine" name="Rivne" pageId="9">Rovno</collectingRegion>
amber, ex coll.
<collectorName id="26DC53246869B614FDECFA0CC6B62A0E" pageId="9">J. Damzen</collectorName>
JDC-11826
<collectionCode id="ED38AE376869B614FEBFFA2AC7012A0E" box="[362,380,1514,1538]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9">R</collectionCode>
, authors coll. FS93BS61 (
<figureCitation id="13122A776869B614FF26F9C9C7162A2D" box="[243,363,1545,1569]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,228,822,846]" captionStartId-1="8.[189,228,1528,1552]" captionTargetBox-0="[189,1171,171,785]" captionTargetBox-1="[189,1171,876,1490]" captionTargetId-0="figure-11@8.[189,1171,171,785]" captionTargetId-1="figure-22@8.[189,1171,876,1490]" captionTargetPageId-0="8" captionTargetPageId-1="8" captionText-0="Fig. 10.Pedostrangalia rovnensisn. sp., female, Paratype, anterior view." captionText-1="Fig. 11.Pedostrangalia rovnensisn. sp., female, Paratype, posterior view." pageId="9">Figs 10-11</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9636F26869B614FFA7F986C42C2A71" blockId="9.[114,593,1606,1661]" pageId="9">The beetle is preserved inside a subtriangular piece of amber measuring about</paragraph>
<paragraph id="8B9636F26869B614FFA7F966C6E72AB2" blockId="9.[114,154,1702,1726]" box="[114,154,1702,1726]" pageId="9">198</paragraph>
</subSubSection>
<subSubSection id="C33365796869B617FDA5FE24C70D2EFE" lastPageId="10" pageId="9" type="diagnosis">
<paragraph id="8B9636F26869B614FDA5FE24C51E2DF0" blockId="9.[624,867,484,508]" box="[624,867,484,508]" pageId="9">
<emphasis id="B95DEAE06869B614FDA5FE24C51E2DF0" bold="true" box="[624,867,484,508]" pageId="9">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="8B9636F26869B614FDA5FDE2C55F2F60" blockId="9.[624,1103,546,1152]" pageId="9">
Both fossils have been identified as members of the genus
<emphasis id="B95DEAE06869B614FC91FD80C2322E54" box="[836,1103,576,600]" italics="true" pageId="9">
<taxonomicName id="4C294D716869B614FC91FD80C5952E54" authorityName="Sokolov" authorityYear="1897" box="[836,1000,576,600]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">Pedostrangalia</taxonomicName>
(s. str.)
</emphasis>
for the following combination of characters: prosternum convex, not forming an angle with the intercoxal process (
<taxonomicName id="4C294D716869B614FC01FD5CC23E2EB8" authorityName="Latreille" authorityYear="1802" box="[980,1091,668,692]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="tribe" tribe="Lepturini">Lepturini</taxonomicName>
); cheeks developed: head with distinct angulated temples; pronotum without apical collar and with acute basal angles; elytra not shortened, posteriorly convergent and not rounded at apex; antennae long and not strongly dentate.
</paragraph>
<paragraph id="8B9636F26869B614FDA5FCB3C5E0288C" blockId="9.[624,1103,546,1152]" pageId="9">
As for the previous species, the peculiar character of the furrowed tarsi in not visible. In absence of this character, the species might be attributed to the Nearctic genus
<taxonomicName id="4C294D716869B614FDA5FC2EC2672809" authority="Linsley &amp; Chemsak, 1976" authorityName="Linsley &amp; Chemsak" authorityYear="1976" box="[624,1050,1005,1030]" class="Insecta" family="Cerambycidae" genus="Pygoleptura" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE06869B614FDA5FC2EC489280A" box="[624,756,1006,1030]" italics="true" pageId="9">Pygoleptura</emphasis>
Linsley &amp; Chemsak, 1976
</taxonomicName>
, but male antennae are not dentate and female antennae are not short as in this genus. Oth- er genera show either strongly reduced temples or a pronotal collar.
</paragraph>
<paragraph id="8B9636F26869B614FDA5FB66C5F72934" blockId="9.[624,1103,1189,1336]" pageId="9">
<taxonomicName id="4C294D716869B614FDA5FB66C5F128B1" authorityName="Vitali" authorityYear="2023" box="[624,908,1190,1214]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. nov.">
<emphasis id="B95DEAE06869B614FDA5FB66C5F128B1" box="[624,908,1190,1214]" italics="true" pageId="9">Pedostrangalia rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6869B614FC47FB66C5AC28B1" box="[914,977,1190,1213]" pageId="9" rank="species">n. sp.</taxonomicNameLabel>
is characterised by elytral apex obliquely truncate, without marginal spine, and pronotum very slightly elongated, strongly sinuous at sides and without lateral angles.
</paragraph>
<paragraph id="8B9636F26869B614FDA5FA9DC4AE2A67" blockId="9.[624,1103,1373,1643]" pageId="9">
These characters are sufficient to separate it from the fossil
<taxonomicName id="4C294D716869B614FCC1FABDC5F82998" authorityName="Vitali" authorityYear="2014" box="[788,901,1405,1428]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="pristina">
<emphasis id="B95DEAE06869B614FCC1FABDC5F82998" box="[788,901,1405,1428]" italics="true" pageId="9">P. pristina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C294D716869B614FC6FFABDC4EA29BF" authorityName="Vitali" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="ostensackeni" status="n. sp.">
<emphasis id="B95DEAE06869B614FC6FFABDC4EA29BF" italics="true" pageId="9">P. ostensackeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6869B614FD48FA5BC4A329BE" box="[669,734,1435,1458]" pageId="9" rank="species">n. sp.</taxonomicNameLabel>
, since both show spined elytral apex. Moreover, the pronotum is trapezoidal (
<taxonomicName id="4C294D716869B614FD74FA19C56E29FC" authorityName="Vitali" authorityYear="2014" box="[673,787,1497,1520]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="pristina">
<emphasis id="B95DEAE06869B614FD74FA19C56E29FC" box="[673,787,1497,1520]" italics="true" pageId="9">P. pristina</emphasis>
</taxonomicName>
) or laterally winkled (
<taxonomicName id="4C294D716869B614FBDCFA19C49C2A03" authorityName="Vitali" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="ostensackeni">
<emphasis id="B95DEAE06869B614FBDCFA19C49C2A03" italics="true" pageId="9">P. ostensackeni</emphasis>
</taxonomicName>
). The coloration of
<taxonomicName id="4C294D716869B614FC17FA38C2352A02" authorityName="Vitali" authorityYear="2023" box="[962,1096,1527,1551]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. n.">
<emphasis id="B95DEAE06869B614FC17FA38C2352A02" box="[962,1096,1527,1551]" italics="true" pageId="9">P. rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B6869B614FDA5F9D6C4D32A21" box="[624,686,1558,1581]" pageId="9" rank="species">sp. n.</taxonomicNameLabel>
is entirely black (like
<taxonomicName id="4C294D716869B614FC60F9D6C4EC2A40" authorityName="Vitali" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="ostensackeni">
<emphasis id="B95DEAE06869B614FC60F9D6C4EC2A40" italics="true" pageId="9">P. ostensackeni</emphasis>
</taxonomicName>
), but body and limbs are robust (like
<taxonomicName id="4C294D716869B614FBE4F9F5C4BE2A67" authorityName="Vitali" authorityYear="2014" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="pristina">
<emphasis id="B95DEAE06869B614FBE4F9F5C4BE2A67" italics="true" pageId="9">P. pristina</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B9636F2686AB617FF68FF10C70D2EFE" blockId="10.[189,668,208,754]" pageId="10">
With respect to the congeners of the Recent,
<taxonomicName id="4C294D71686AB617FF2DFF30C7012D0A" authorityName="Vitali" authorityYear="2023" box="[248,380,239,263]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. n.">
<emphasis id="B95DEAE0686AB617FF2DFF30C7012D0A" box="[248,380,239,263]" italics="true" pageId="10">P. rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B686AB617FE51FF2FC7C32D0A" box="[388,446,239,262]" pageId="10" rank="species">sp. n.</taxonomicNameLabel>
should have looked somehow similar to the Anatolian
<taxonomicName id="4C294D71686AB617FDE9FECEC7F02D48" authority="(Pic, 1892)" baseAuthorityName="Pic" baseAuthorityYear="1892" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="verticenigra">
<emphasis id="B95DEAE0686AB617FDE9FECEC7732D48" italics="true" pageId="10">P. verticenigra</emphasis>
(Pic, 1892)
</taxonomicName>
and the Balkan
<taxonomicName id="4C294D71686AB617FDEBFEEDC7D92D6F" authority="(German, 1822)" baseAuthorityName="German" baseAuthorityYear="1822" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B95DEAE0686AB617FDEBFEEDC6922D6F" italics="true" pageId="10">P. verticalis</emphasis>
(German, 1822)
</taxonomicName>
but with much longer temples. Both species belong in fact to the subgenus
<taxonomicName id="4C294D71686AB617FEFFFE48C43C2DAC" authority="Lobl, 2010" authorityName="Lobl" authorityYear="2010" box="[298,577,392,416]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="subGenus" subGenus="Neosphenalia">
<emphasis id="B95DEAE0686AB617FEFFFE48C7C12DAC" box="[298,444,392,416]" italics="true" pageId="10">Neosphenalia</emphasis>
Löbl, 2010
</taxonomicName>
. Elytral apex and pronotal shape remind as well of the European
<taxonomicName id="4C294D71686AB617FEB6FE06C7992DD1" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[355,484,454,477]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="revestita">
<emphasis id="B95DEAE0686AB617FEB6FE06C7992DD1" box="[355,484,454,477]" italics="true" pageId="10">P. revestita</emphasis>
</taxonomicName>
, especially the melanistic form
<emphasis id="B95DEAE0686AB617FEA4FE24C7982DF0" box="[369,485,484,508]" italics="true" pageId="10">fulvilabris</emphasis>
Mulsant, but elytra are not carinate and the pronotum is without lateral angles as in this species and other ones of this genus. Possibly,
<taxonomicName id="4C294D71686AB617FD92FD81C77D2E7A" authorityName="Vitali" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="rovnensis" status="sp. n.">
<emphasis id="B95DEAE0686AB617FD92FD81C77D2E7A" italics="true" pageId="10">P. rovnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B686AB617FEDEFD9FC7352E7A" box="[267,328,607,630]" pageId="10" rank="species">sp. n.</taxonomicNameLabel>
shows more affinity with the Chinese
<taxonomicName id="4C294D71686AB617FECBFDBEC4182E9A" authority="Holzschuh, 1999" authorityName="Holzschuh" authorityYear="1999" box="[286,613,637,662]" class="Insecta" family="Cerambycidae" genus="Pedostrangalia" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="signifera">
<emphasis id="B95DEAE0686AB617FECBFDBEC7E22E9A" box="[286,415,638,662]" italics="true" pageId="10">P. signifera</emphasis>
Holzschuh, 1999
</taxonomicName>
, but the pronotum is not so enlarged at base as in this species and the coloration is completely different.
</paragraph>
</subSubSection>
<subSubSection id="C3336579686AB617FF68FCD7C5B22998" pageId="10" type="description">
<paragraph id="8B9636F2686AB617FF68FCD7C73C2F23" blockId="10.[189,321,791,815]" box="[189,321,791,815]" pageId="10">
<emphasis id="B95DEAE0686AB617FF68FCD7C73C2F23" bold="true" box="[189,321,791,815]" pageId="10">Description</emphasis>
</paragraph>
<paragraph id="8B9636F2686AB617FF68FC94C46E2FC4" blockId="10.[189,669,852,968]" pageId="10">
Male: body length
<quantity id="4CD19B17686AB617FE41FC94C7902F60" box="[404,493,852,876]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="10" unit="mm" value="8.3">8.3 mm</quantity>
. Female: Body length ~
<quantity id="4CD19B17686AB617FECCFCB3C7142F86" box="[281,361,883,907]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" unit="mm" value="10.0">10 mm</quantity>
(difficult guessable due to the position). General habitus small, elongated, drop-shaped; body black.
</paragraph>
<paragraph id="8B9636F2686AB617FF68FC2DC4992D0B" blockId="10.[189,669,1005,1643]" lastBlockId="10.[699,1178,178,263]" pageId="10">Head relatively short; cheeks developed; clypeus and forehead transverse; antennal tubercles widely separated, fairly elevated; eyes far from the base of the mandibles, finely faceted; however, their shape not guessable; temples relatively long, a bit shorter than cheeks, parallel-sided, abruptly converging backward; neck almost long. Palps not observable. Antennae 11- segmented, inserted between the eyes, hardly reaching the elytral apex, finely pubescent, extremely finely and densely punctured; scape sub-linear; pedicle as long as broad or hardly elongate, less than one-fourth as long as scape; antennomere III slightly longer than scape; antennomere IV slightly shorter than scape; antennomere V the longest, nearly one-third longer than scape; antennomeres VI and VII sub-equal; following antennomeres decreasing in length, except for the last, about as long as III; proportions according to the formula: 4.2: 1.0: 4.5: 4.0: 5.4: 4.8: 4.8: 4.2: 3.9: 3.5: 4.5.</paragraph>
<paragraph id="8B9636F2686AB617FD6EFEECC5D62E7B" blockId="10.[699,1179,300,631]" pageId="10">Prothorax slightly elongate, bell-shaped, regularly enlarged posteriorly, hind angles short and acute, shortly embracing the elytral base; apex straight, finely grooved; sides without winkles; base posteriorly lobed in the middle, finely grooved; disc convex above, without longitudinal furrow, entirely covered with an extremely fine dense puncturing and a fine short recumbent pubescence. Scutellum small, forming an equilateral triangle.</paragraph>
<paragraph id="8B9636F2686AB617FD6EFD5CC5C82F87" blockId="10.[699,1179,668,907]" pageId="10">Elytra relatively short, 2.15 times as long as wide at the shoulders, slightly more than 3 times as long as pronotum, flat above, nearly parallel-sided, feebly tapered posteriorly; apex almost transversely truncate, without marginal spine; surface covered with a coarsely fine puncturing and a fine short recumbent pubescence.</paragraph>
<paragraph id="8B9636F2686AB617FD6EFC70C5572809" blockId="10.[699,1179,944,1029]" pageId="10">Ventral side convex; prosternum regularly convex in lateral view; other characters not guessable.</paragraph>
<paragraph id="8B9636F2686AB617FD6EFBEBC5B22998" blockId="10.[699,1179,1067,1428]" pageId="10">Legs relatively robust; femora slightly club-shaped; tibiae linear, rectilinearly truncated at the apex, finely punctured and densely pubescent; apex of metatibiae with two subequal spines, 0.4 as long as metatarsomere I. Metatarsi long, a bit shorter than metatibiae; metatasomere I as long as the following articles together; metatarsomere II nearly one-half as long as I; metatarsomere III one-third shorter than II; onychium as long as III (proportions according to the formula: 7: 3: 2.5:?: 2.5).</paragraph>
</subSubSection>
<subSubSection id="C3336579686AB617FD6EFA79C58B2A21" pageId="10" type="etymology">
<paragraph id="8B9636F2686AB617FD6EFA79C54B29DD" blockId="10.[699,822,1465,1489]" box="[699,822,1465,1489]" pageId="10">
<emphasis id="B95DEAE0686AB617FD6EFA79C54B29DD" bold="true" box="[699,822,1465,1489]" pageId="10">Etymology</emphasis>
</paragraph>
<paragraph id="8B9636F2686AB617FD6EFA37C58B2A21" blockId="10.[699,1178,1527,1581]" pageId="10">
The specific epithet refers to the origin of this amber:
<collectingRegion id="49EDF810686AB617FCECF9D6C5FD2A21" box="[825,896,1558,1581]" country="Ukraine" name="Rivne" pageId="10">Rovno</collectingRegion>
(
<collectingCountry id="F33E7662686AB617FC5BF9D5C5972A21" box="[910,1002,1557,1581]" name="Ukraine" pageId="10">Ukraine</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,193 @@
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<treatment id="038087E4686BB611FFA7FF72C5632FEB" LSID="urn:lsid:plazi:treatment:038087E4686BB611FFA7FF72C5632FEB" httpUri="http://treatment.plazi.org/id/038087E4686BB611FFA7FF72C5632FEB" lastPageId="12" pageId="11">
<subSubSection id="C3336579686BB616FFA7FF72C7C82CC5" box="[114,437,177,202]" pageId="11" type="nomenclature">
<paragraph id="8B9636F2686BB616FFA7FF72C7C82CC5" blockId="11.[114,437,177,202]" box="[114,437,177,202]" pageId="11">
<heading id="D0DE819E686BB616FFA7FF72C7C82CC5" bold="true" box="[114,437,177,202]" centered="true" fontSize="10" level="2" pageId="11" reason="4">
<emphasis id="B95DEAE0686BB616FFA7FF72C7C82CC5" bold="true" box="[114,437,177,202]" pageId="11">
<taxonomicName id="4C294D71686BB616FFA7FF72C7042CC6" authorityName="Vitali" authorityYear="2023" box="[114,377,178,202]" class="Insecta" family="Cerambycidae" genus="Anoplodera" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="volyniensis" status="sp. nov.">
<emphasis id="B95DEAE0686BB616FFA7FF72C7042CC6" bold="true" box="[114,377,178,202]" italics="true" pageId="11">Anoplodera volyniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B686BB616FEACFF71C7C82CC5" box="[377,437,177,201]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3336579686BB616FDA5FF72C6DD2961" pageId="11" type="description">
<paragraph id="8B9636F2686BB616FDA5FF72C6912FF1" blockId="11.[624,718,178,202]" lastBlockId="11.[114,593,936,1389]" pageId="11">
(
<materialsCitation id="3B413CAF686BB616FDADFF72C6912FF1" collectionCode="R" collectorName="J. Damzen" location="Rovno" pageId="11" specimenCount="1" stateProvince="Rivne" typeStatus="holotype">
<figureCitation id="13122A77686BB616FDADFF72C4B82CC6" box="[632,709,178,202]" captionStart="Fig" captionStartId="11.[114,153,874,898]" captionTargetBox="[114,1081,232,837]" captionTargetId="figure-10@11.[114,1081,232,837]" captionTargetPageId="11" captionText="Fig. 13.Anoplodera volyniensis n. sp. Holotype" pageId="11">Fig. 13</figureCitation>
)
<emphasis id="B95DEAE0686BB616FFA7FC68C69C2FCC" bold="true" box="[114,225,936,960]" pageId="11">
<typeStatus id="54928850686BB616FFA7FC68C6A12FCC" box="[114,220,936,960]" pageId="11" type="holotype">Holotype</typeStatus>
.
</emphasis>
male,
<collectingRegion id="49EDF810686BB616FEE7FC68C7042FB3" box="[306,377,936,959]" country="Ukraine" name="Rivne" pageId="11">Rovno</collectingRegion>
amber, ex coll.
<collectorName id="26DC5324686BB616FDECFC68C6B62FD2" pageId="11">J. Damzen</collectorName>
JDC-9448
<collectionCode id="ED38AE37686BB616FEB4FC07C70E2FD3" box="[353,371,967,991]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11">R</collectionCode>
, authors coll. FS89BS57.
</materialsCitation>
</paragraph>
<caption id="DF56667A686BB616FFA7FCAAC4122F8E" box="[114,623,874,898]" pageId="11" startId="11.[114,153,874,898]" targetBox="[114,1081,232,837]" targetPageId="11" targetType="figure">
<paragraph id="8B9636F2686BB616FFA7FCAAC4122F8E" blockId="11.[114,623,874,898]" box="[114,623,874,898]" pageId="11">
Fig. 13.
<taxonomicName id="4C294D71686BB616FF1EFCAAC7B82F8E" authorityName="Vitali" authorityYear="2023" box="[203,453,874,898]" class="Insecta" family="Cerambycidae" genus="Anoplodera" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="volyniensis" status="sp. nov.">
<emphasis id="B95DEAE0686BB616FF1EFCAAC7B82F8E" box="[203,453,874,898]" italics="true" pageId="11">Anoplodera volyniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E579B686BB616FE19FCABC47B2F8E" box="[460,518,875,898]" pageId="11" rank="species">n. sp.</taxonomicNameLabel>
Holotype
</paragraph>
</caption>
<paragraph id="8B9636F2686BB616FFA7FBC4C767289A" blockId="11.[114,593,936,1389]" pageId="11">
The insect is preserved inside a prismatic piece of amber measuring about 44x35x
<quantity id="4CD19B17686BB616FDE4FBE2C6E22855" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="11" unit="mm" value="18.0">18 mm</quantity>
. It is perfectly preserved but the posterior part of the corps is covered with a fine milky turbidity.
</paragraph>
<paragraph id="8B9636F2686BB616FFA7FB5DC6DD2961" blockId="11.[114,593,936,1389]" pageId="11">
The amber piece also contains minute zoological inclusions: a specimen of
<taxonomicName id="4C294D71686BB616FFA7FB1AC76928FE" authorityName="Latreille" authorityYear="1802" box="[114,276,1242,1266]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="family">Chrysomelidae</taxonomicName>
Alticini, another identified beetle (only elytra visible), one
<taxonomicName id="4C294D71686BB616FE0DFB3AC6E72923" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="11" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
<taxonomicName id="4C294D71686BB616FF74FAD8C768293C" baseAuthorityName="Forster" baseAuthorityYear="1856" box="[161,277,1304,1328]" class="Insecta" family="Encyrtidae" kingdom="Animalia" order="Hymenoptera" pageId="11" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
and several dipterans of different families (
<taxonomicName id="4C294D71686BB616FEF9FAF6C7B62942" authorityName="Newman" authorityYear="1835" box="[300,459,1334,1358]" class="Insecta" family="Cecidomyidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" phylum="Arthropoda" rank="family">Cecidomyidae</taxonomicName>
,
<taxonomicName id="4C294D71686BB616FE08FAF6C4312942" box="[477,588,1334,1358]" class="Insecta" family="Tipulidae" kingdom="Animalia" order="Diptera" pageId="11" phylum="Arthropoda" rank="family">Tipulidae</taxonomicName>
, etc.)
</paragraph>
</subSubSection>
<subSubSection id="C3336579686BB611FFA7FA52C7CF2DAC" lastPageId="12" pageId="11" type="diagnosis">
<paragraph id="8B9636F2686BB616FFA7FA52C71829A6" blockId="11.[114,357,1426,1450]" box="[114,357,1426,1450]" pageId="11">
<emphasis id="B95DEAE0686BB616FFA7FA52C71829A6" bold="true" box="[114,357,1426,1450]" pageId="11">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="8B9636F2686BB616FFA7FA10C42D2A6E" blockId="11.[114,593,1488,1634]" pageId="11">The identification at the level of genus was rather complicated due to the position of the beetle, which did not allow observing exactly the pronotal shape, especially, the basal angles. The pronotum is convex, almost</paragraph>
<paragraph id="8B9636F2686BB616FFA7F966C57D2855" blockId="11.[114,154,1702,1726]" lastBlockId="11.[624,1103,936,1113]" pageId="11">
200 cylindrical, evidently narrower than elytral base, while elytra are parallel-sided and rounded at apex. This character set reminds some basal genera like
<taxonomicName id="4C294D71686BB616FCA6FBC4C2772810" box="[883,1034,1028,1052]" class="Insecta" family="Cerambycidae" genus="Grammoptera" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686BB616FCA6FBC4C2772810" box="[883,1034,1028,1052]" italics="true" pageId="11">Grammoptera</emphasis>
</taxonomicName>
,
<emphasis id="B95DEAE0686BB616FBCFFBC4C2772836" italics="true" pageId="11">
<taxonomicName id="4C294D71686BB616FBCFFBC4C4C82836" authorityName="Mulsant" authorityYear="1863" class="Insecta" family="Cerambycidae" genus="Alosterna" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">Alosterna</taxonomicName>
,
<taxonomicName id="4C294D71686BB616FD0DFBE2C5242836" authorityName="Mulsant" authorityYear="1839" box="[728,857,1058,1082]" class="Insecta" family="Cerambycidae" genus="Anoplodera" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">Anoplodera</taxonomicName>
,
<taxonomicName id="4C294D71686BB616FCAEFBE2C2772836" authorityName="Casey" authorityYear="1913" box="[891,1034,1058,1082]" class="Insecta" family="Cerambycidae" genus="Xestoleptura" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">Xestoleptura</taxonomicName>
</emphasis>
and
<taxonomicName id="4C294D71686BB616FDA5FB81C4872855" authorityName="Casey" authorityYear="1913" box="[624,762,1089,1113]" class="Insecta" family="Cerambycidae" genus="Strangalepta" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686BB616FDA5FB81C4872855" box="[624,762,1089,1113]" italics="true" pageId="11">Strangalepta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9636F2686BB616FDA5FBBEC48229C5" blockId="11.[624,1103,1150,1481]" pageId="11">
Finally, the fossil has been identified as a member of the genus
<taxonomicName id="4C294D71686BB616FCA1FB5DC58F28B9" authorityName="Mulsant" authorityYear="1839" box="[884,1010,1181,1205]" class="Insecta" family="Cerambycidae" genus="Anoplodera" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686BB616FCA1FB5DC58F28B9" box="[884,1010,1181,1205]" italics="true" pageId="11">Anoplodera</emphasis>
</taxonomicName>
on the basis of the following characters: 1) pronotal base without transversal furrow and 2) with a small indentation at each side; 3) antennae angulated externally from the fifth article; 4) elytral base straight. The first character excludes
<taxonomicName id="4C294D71686BB616FC8CFA95C59F2961" authorityName="Casey" authorityYear="1913" box="[857,994,1365,1389]" class="Insecta" family="Cerambycidae" genus="Strangalepta" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686BB616FC8CFA95C59F2961" box="[857,994,1365,1389]" italics="true" pageId="11">Strangalepta</emphasis>
</taxonomicName>
, second and third ones exclude
<taxonomicName id="4C294D71686BB616FCABFAB4C2682980" box="[894,1045,1396,1420]" class="Insecta" family="Cerambycidae" genus="Grammoptera" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686BB616FCABFAB4C2682980" box="[894,1045,1396,1420]" italics="true" pageId="11">Grammoptera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C294D71686BB616FDA5FA52C4A529A6" authorityName="Mulsant" authorityYear="1863" box="[624,728,1426,1450]" class="Insecta" family="Cerambycidae" genus="Alosterna" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686BB616FDA5FA52C4A529A6" box="[624,728,1426,1450]" italics="true" pageId="11">Alosterna</emphasis>
</taxonomicName>
, while the fourth one excludes
<taxonomicName id="4C294D71686BB616FDA5FA71C48529C5" authorityName="Casey" authorityYear="1913" box="[624,760,1457,1481]" class="Insecta" family="Cerambycidae" genus="Xestoleptura" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686BB616FDA5FA71C48529C5" box="[624,760,1457,1481]" italics="true" pageId="11">Xestoleptura</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9636F2686BB611FDA5FA2EC7CF2DAC" blockId="11.[624,1103,1518,1634]" lastBlockId="12.[189,668,178,417]" lastPageId="12" pageId="11">
The antennal proportions (article III longer than IV and a bit shorter than V) correspond to the extant
<taxonomicName id="4C294D71686BB616FC80F9ECC5AE2A48" authorityName="Mulsant" authorityYear="1839" box="[853,979,1580,1604]" class="Insecta" family="Cerambycidae" genus="Anoplodera" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686BB616FC80F9ECC5AE2A48" box="[853,979,1580,1604]" italics="true" pageId="11">Anoplodera</emphasis>
</taxonomicName>
; however, this fossil does not perfectly fit with this genus. In fact it shows long temples (as
<taxonomicName id="4C294D71686CB611FF68FF10C7382CE4" authorityName="Casey" authorityYear="1913" box="[189,325,208,232]" class="Insecta" family="Cerambycidae" genus="Xestoleptura" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686CB611FF68FF10C7382CE4" box="[189,325,208,232]" italics="true" pageId="12">Xestoleptura</emphasis>
</taxonomicName>
) and a pubescence of semierect setae, such as several cerambycids in Baltic amber but no extant
<taxonomicName id="4C294D71686CB611FDC4FECEC4F22D2A" authorityName="Mulsant" authorityYear="1839" box="[529,655,270,294]" class="Insecta" family="Cerambycidae" genus="Anoplodera" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEAE0686CB611FDC4FECEC4F22D2A" box="[529,655,270,294]" italics="true" pageId="12">Anoplodera</emphasis>
</taxonomicName>
. Possibly, it belongs to a new undescribed (sub)genus, but I prefer waiting for further more observable specimens before proposing a new genus name.
</paragraph>
</subSubSection>
<subSubSection id="C3336579686CB611FF68FE05C5CC2F1C" pageId="12" type="description">
<paragraph id="8B9636F2686CB611FF68FE05C73C2DD1" blockId="12.[189,321,453,477]" box="[189,321,453,477]" pageId="12">
<emphasis id="B95DEAE0686CB611FF68FE05C73C2DD1" bold="true" box="[189,321,453,477]" pageId="12">Description</emphasis>
</paragraph>
<paragraph id="8B9636F2686CB611FF68FDC3C4FC2E36" blockId="12.[189,668,515,570]" pageId="12">
Undetermined sex, probably male; length
<quantity id="4CD19B17686CB611FF68FDE2C76F2E35" box="[189,274,546,570]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="12" unit="mm" value="8.3">8.3 mm</quantity>
. General habitus small, elongated.
</paragraph>
<paragraph id="8B9636F2686CB611FF68FD9FC44728F7" blockId="12.[189,669,607,1275]" pageId="12">Head relatively short; cheeks almost developed but; clypeus and forehead complanar; antennal tubercles widely separated, fairly elevated; eyes relatively close to the base of the mandibles, emarginate at the upper side, uniformly convex at the under one, finely faceted; temples long, as long as the hind eye-lobes, feebly converging backward; neck one-third as long as temples. Palpi not visible. Antennae 11-segmented, almost robust, inserted between the eyes, hardly reaching the elytral apex, covered with dense recumbent pubescence; scape sub-linear; pedicle scarcely longer than broad, about one-third as long as scape; antennomere III one-fourth longer than scape; antennomere IV one-tenth shorter than III; antennomere V the longest, twice as long as scape; following antennomeres decreasing in length, except for the last one (proportions according to the formula: 3.2: 1.0: 4.0: 3.8: 6.4: 5.8: 5.8: 5.4: 5.2: 4.2: 4.4).</paragraph>
<paragraph id="8B9636F2686CB611FF68FAE0C76B2A40" blockId="12.[189,669,1312,1612]" pageId="12">Prothorax elongate, cylindrical, (hind angles rounded), evidently narrower than the elytral base; apex straight and finely grooved; base with a small indentation at each side, finely grooved; disc convex above, without longitudinal furrow, everywhere covered with a fine dense puncturing and some oblique strong black setae. Scutellum small, elongate, forming an isosceles triangle.</paragraph>
<paragraph id="8B9636F2686CB611FD6EFF72C5FF2D6E" blockId="12.[699,1179,177,355]" pageId="12">Elytra long, 6.5 times as long as pronotum, flat above, parallel-sided, widely rounded at apex; surface covered with a coarsely dense puncturing, a fine short recumbent pubescence and a sparse pubescence of oblique strong black setae.</paragraph>
<paragraph id="8B9636F2686CB611FD6EFE48C5C92DF0" blockId="12.[699,1178,392,508]" pageId="12">Ventral side convex, covered with a dense recumbent short pubescence; prosternum feebly convex in lateral view; procoxal cavities posteriorly closed.</paragraph>
<paragraph id="8B9636F2686CB611FD6EFDE2C5CC2F1C" blockId="12.[699,1179,546,784]" pageId="12">Legs relatively short; femora slightly club-shaped; tibiae linear, rectilinearly truncated at the apex, densely pubescent; apex of mesotibiae armed with two sub-equal spines; apex of metatibiae with two unequal spines, the longest one one-third as long as metatarsomere I; metatarsi long, their proportions not guessable.</paragraph>
</subSubSection>
<subSubSection id="C3336579686CB611FD6EFCF5C5632FEB" pageId="12" type="etymology">
<paragraph id="8B9636F2686CB611FD6EFCF5C54B2F41" blockId="12.[699,822,821,845]" box="[699,822,821,845]" pageId="12">
<emphasis id="B95DEAE0686CB611FD6EFCF5C54B2F41" bold="true" box="[699,822,821,845]" pageId="12">Etymology</emphasis>
</paragraph>
<paragraph id="8B9636F2686CB611FD6EFCB3C5632FEB" blockId="12.[699,1179,883,999]" pageId="12">
The specific epithet refers to Volhynia, the historic region, today shared by
<collectingCountry id="F33E7662686CB611FB96FC52C2EB2FA6" box="[1091,1174,914,938]" name="Poland" pageId="12">Poland</collectingCountry>
,
<collectingCountry id="F33E7662686CB611FD6EFC70C5712FC4" box="[699,780,944,968]" name="Belarus" pageId="12">Belarus</collectingCountry>
and
<collectingCountry id="F33E7662686CB611FC9FFC70C5DE2FC4" box="[842,931,944,968]" name="Ukraine" pageId="12">Ukraine</collectingCountry>
, where
<collectingRegion id="49EDF810686CB611FC2BFC71C2382FC4" box="[1022,1093,945,968]" country="Ukraine" name="Rivne" pageId="12">Rovno</collectingRegion>
amber is mined.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,335 @@
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<mods:title id="5ABCAA35CB8F74BA1E415290D4DCA179">Taxonomic notes on the genus Hexagonia Kirby, 1825 with description of three new species from the Philippines (Coleoptera: Carabidae: Hexagoniini)</mods:title>
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<emphasis id="B95AEAC2FFA5B11DFCE13752FAC5FDDF" bold="true" box="[850,1360,544,569]" pageId="8" pageNumber="63">
<taxonomicName id="4C2E4D53FFA5B11DFCE13752FAB8FDDF" authority="Anichtchenko" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[850,1325,544,569]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="nigrocyanea">
<emphasis id="B95AEAC2FFA5B11DFCE13752FBE2FDDE" bold="true" box="[850,1143,545,569]" italics="true" pageId="8" pageNumber="63">Hexagonia nigrocyanea</emphasis>
Anichtchenko
</taxonomicName>
&amp;
</emphasis>
</heading>
</paragraph>
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Sciaky
<taxonomicNameLabel id="A26957B9FFA5B11DFC163733FC74FDBF" box="[933,993,576,600]" pageId="8" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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(
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,
<figureCitation id="13152A55FFA5B11DFC013712FC59FD9F" box="[946,972,609,632]" captionStart="Figs" captionStartId="10.[302,352,1399,1422]" captionTargetBox="[307,1358,191,1362]" captionTargetId="figure-177@10.[302,1360,189,1366]" captionTargetPageId="10" captionText="Figs. 711. Aedeagus of Hexagonia, A dorsal view, B lateral view. 7 H. andrewesi JedliËka, 1935, Holotype; 8 H. castanea JedliËka, 1936 syn. n. Cotype; 9 H. sauteri Dupuis, 1912, Holotype. Scale bar 1 mm. 10 H. amicitia sp. n., Holotype; 11 H. nigrocyanea sp. n. Holotype. Scale bar 1 mm." pageId="8" pageNumber="63">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA5B11DFCE137D3FB4CFC3F" pageId="8" pageNumber="63" type="materials_examined">
<paragraph id="8B9136D0FFA5B11DFCE137D3FB4CFC3F" blockId="8.[850,1361,672,984]" pageId="8" pageNumber="63">
<emphasis id="B95AEAC2FFA5B11DFCE137D3FC69FD5F" bold="true" box="[850,1020,672,696]" pageId="8" pageNumber="63">Type material:</emphasis>
<materialsCitation id="3B463C8DFFA5B11DFBB037D2FA99FD3F" collectingDate="2019-02" country="Philippines" location="Mindanao" municipality="Dominorog" pageId="8" pageNumber="63" specimenCount="1" typeStatus="holotype">
<typeStatus id="54958872FFA5B11DFBB037D2FBFFFD5F" box="[1027,1130,673,696]" pageId="8" pageNumber="63" type="holotype">Holotype</typeStatus>
, male
<collectingCountry id="F3397640FFA5B11DFB7D37D2FAD8FD5F" box="[1230,1357,673,696]" name="Philippines" pageId="8" pageNumber="63">Philippines</collectingCountry>
,
<location id="8EF1600BFFA5B11DFCE137B2FC2AFD3F" LSID="urn:lsid:plazi:treatment:038787C6FFA5B11EFCE13752FEBFFABF:8EF1600BFFA5B11DFCE137B2FC2AFD3F" box="[850,959,705,728]" country="Philippines" municipality="Dominorog" name="Mindanao" pageId="8" pageNumber="63">Mindanao</location>
|
<collectingMunicipality id="6BF5ACAAFFA5B11DFC6637B2FBC4FD3F" box="[981,1105,705,728]" pageId="8" pageNumber="63">Dominorog</collectingMunicipality>
|
<date id="FF901010FFA5B11DFBD437B2FB22FD3F" box="[1127,1207,705,728]" pageId="8" pageNumber="63" value="2019-02">
<collectingDate id="EFD4E9F8FFA5B11DFBD437B2FB22FD3F" box="[1127,1207,705,728]" pageId="8" pageNumber="63" value="2019-02">II.2019</collectingDate>
</date>
” (cRS)
</materialsCitation>
.
<materialsCitation id="3B463C8DFFA5B11DFAAB37B2FB09FCFF" collectingDate="2019-05" country="Philippines" location="Dominorog" municipality="Mindanao" pageId="8" pageNumber="63" specimenCount="1" specimenCount-female="1" stateProvince="Bukidnon" typeStatus="paratype">
<typeStatus id="54958872FFA5B11DFAAB37B2FC18FD1F" pageId="8" pageNumber="63" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D28FD59FFA5B11DFC263792FC78FD1F" box="[917,1005,737,760]" count="1" pageId="8" pageNumber="63" type="female">1 female</specimenCount>
<collectingCountry id="F3397640FFA5B11DFBBE3792FB12FD1F" box="[1037,1159,737,760]" name="Philippines" pageId="8" pageNumber="63">Philippines</collectingCountry>
,
<collectingMunicipality id="6BF5ACAAFFA5B11DFB3C3792FB6CFD1F" box="[1167,1273,737,760]" pageId="8" pageNumber="63">Mindanao</collectingMunicipality>
|
<collectingRegion id="49EAF832FFA5B11DFABB3792FCECFCFF" country="Philippines" name="Bukidnon" pageId="8" pageNumber="63">Bukidnon</collectingRegion>
|
<location id="8EF1600BFFA5B11DFC343672FC6AFCFF" LSID="urn:lsid:plazi:treatment:038787C6FFA5B11EFCE13752FEBFFABF:8EF1600BFFA5B11DFC343672FC6AFCFF" box="[903,1023,769,792]" country="Philippines" municipality="Mindanao" name="Dominorog" pageId="8" pageNumber="63" stateProvince="Bukidnon">Dominorog</location>
<date id="FF901010FFA5B11DFBB03672FBDBFCFF" box="[1027,1102,769,792]" pageId="8" pageNumber="63" value="2019-05">
<collectingDate id="EFD4E9F8FFA5B11DFBB03672FBDBFCFF" box="[1027,1102,769,792]" pageId="8" pageNumber="63" value="2019-05">V.2019</collectingDate>
</date>
” (cRS)
</materialsCitation>
;
<materialsCitation id="3B463C8DFFA5B11DFB173672FC07FCBF" collectingDate="2017-10" country="Philippines" location="Maitum" municipality="Mindanao" pageId="8" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="Sarangani" typeStatus="paratype">
<specimenCount id="9D28FD59FFA5B11DFB173672FB7CFCFF" box="[1188,1257,769,792]" count="1" pageId="8" pageNumber="63" type="male">1 male</specimenCount>
<collectingCountry id="F3397640FFA5B11DFABA3672FC1BFCDF" name="Philippines" pageId="8" pageNumber="63">Philippines</collectingCountry>
,
<collectingMunicipality id="6BF5ACAAFFA5B11DFC2A3652FB93FCDF" box="[921,1030,801,824]" pageId="8" pageNumber="63">Mindanao</collectingMunicipality>
|
<collectingRegion id="49EAF832FFA5B11DFBAA3652FB10FCDF" box="[1049,1157,801,824]" country="Philippines" name="Sarangani" pageId="8" pageNumber="63">Sarangani</collectingRegion>
|
<location id="8EF1600BFFA5B11DFB2B3652FB78FCDF" LSID="urn:lsid:plazi:treatment:038787C6FFA5B11EFCE13752FEBFFABF:8EF1600BFFA5B11DFB2B3652FB78FCDF" box="[1176,1261,801,824]" country="Philippines" municipality="Mindanao" name="Maitum" pageId="8" pageNumber="63" stateProvince="Sarangani">Maitum</location>
<date id="FF901010FFA5B11DFB473652FAD2FCDF" box="[1268,1351,801,824]" pageId="8" pageNumber="63" value="2017-10">
<collectingDate id="EFD4E9F8FFA5B11DFB473652FAD2FCDF" box="[1268,1351,801,824]" pageId="8" pageNumber="63" value="2017-10">X.2017</collectingDate>
</date>
” (cRS)
</materialsCitation>
;
<materialsCitation id="3B463C8DFFA5B11DFC283632FB04FC9F" collectingDate="2019-11" country="Philippines" location="Wao" municipality="Mindanao" pageId="8" pageNumber="63" specimenCount="1" specimenCount-female="1" stateProvince="Lanao del Sur" typeStatus="paratype">
<specimenCount id="9D28FD59FFA5B11DFC283632FC61FCBF" box="[923,1012,833,856]" count="1" pageId="8" pageNumber="63" type="female">1 female</specimenCount>
<collectingCountry id="F3397640FFA5B11DFBA03632FB18FCBF" box="[1043,1165,833,856]" name="Philippines" pageId="8" pageNumber="63">Philippines</collectingCountry>
,
<collectingMunicipality id="6BF5ACAAFFA5B11DFB253632FA95FCBF" box="[1174,1280,833,856]" pageId="8" pageNumber="63">Mindanao</collectingMunicipality>
|
<collectingRegion id="49EAF832FFA5B11DFABC3632FC0BFC9F" country="Philippines" name="Lanao del Sur" pageId="8" pageNumber="63">Lanao del Sur</collectingRegion>
|
<location id="8EF1600BFFA5B11DFC1D3612FC4BFC9F" LSID="urn:lsid:plazi:treatment:038787C6FFA5B11EFCE13752FEBFFABF:8EF1600BFFA5B11DFC1D3612FC4BFC9F" box="[942,990,865,888]" country="Philippines" municipality="Mindanao" name="Wao" pageId="8" pageNumber="63" stateProvince="Lanao del Sur">Wao</location>
<date id="FF901010FFA5B11DFC573612FBAAFC9F" box="[996,1087,865,888]" pageId="8" pageNumber="63" value="2019-11">
<collectingDate id="EFD4E9F8FFA5B11DFC573612FBAAFC9F" box="[996,1087,865,888]" pageId="8" pageNumber="63" value="2019-11">XI.2019</collectingDate>
</date>
” (cRS)
</materialsCitation>
;
<materialsCitation id="3B463C8DFFA5B11DFB283612FC01FC5F" collectingDate="2021-11" country="Philippines" location="Wao" municipality="Mindanao" pageId="8" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="Lanao del Sur" typeStatus="paratype">
<specimenCount id="9D28FD59FFA5B11DFB283612FB77FC9F" box="[1179,1250,865,888]" count="1" pageId="8" pageNumber="63" type="male">1 male</specimenCount>
<collectingCountry id="F3397640FFA5B11DFAB53612FC1BFC7F" name="Philippines" pageId="8" pageNumber="63">Philippines</collectingCountry>
,
<collectingMunicipality id="6BF5ACAAFFA5B11DFC2436F2FB92FC7F" box="[919,1031,897,920]" pageId="8" pageNumber="63">Mindanao</collectingMunicipality>
,
<collectingRegion id="49EAF832FFA5B11DFBA336F2FB31FC7F" box="[1040,1188,897,920]" country="Philippines" name="Lanao del Sur" pageId="8" pageNumber="63">Lanao del Sur</collectingRegion>
,
<location id="8EF1600BFFA5B11DFB1C36F2FB77FC7F" LSID="urn:lsid:plazi:treatment:038787C6FFA5B11EFCE13752FEBFFABF:8EF1600BFFA5B11DFB1C36F2FB77FC7F" box="[1199,1250,897,920]" country="Philippines" municipality="Mindanao" name="Wao" pageId="8" pageNumber="63" stateProvince="Lanao del Sur">Wao</location>
,
<date id="FF901010FFA5B11DFB5836F2FAD3FC7F" box="[1259,1350,897,920]" pageId="8" pageNumber="63" value="2021-11">
<collectingDate id="EFD4E9F8FFA5B11DFB5836F2FAD3FC7F" box="[1259,1350,897,920]" pageId="8" pageNumber="63" value="2021-11">XI.2021</collectingDate>
</date>
” (cRS)
</materialsCitation>
;
<materialsCitation id="3B463C8DFFA5B11DFC1336D2FAA3FC5F" box="[928,1334,929,952]" collectingDate="2021-10" country="Philippines" location="Wao" municipality="Mindanao" pageId="8" pageNumber="63" specimenCount="1" specimenCount-female="1" stateProvince="Lanao del Sur" typeStatus="paratype">
<specimenCount id="9D28FD59FFA5B11DFC1336D2FC6AFC5F" box="[928,1023,929,952]" count="1" pageId="8" pageNumber="63" type="female">1 female</specimenCount>
idem but “
<date id="FF901010FFA5B11DFB3C36D2FB77FC5F" box="[1167,1250,929,952]" pageId="8" pageNumber="63" value="2021-10">
<collectingDate id="EFD4E9F8FFA5B11DFB3C36D2FB77FC5F" box="[1167,1250,929,952]" pageId="8" pageNumber="63" value="2021-10">X.2021</collectingDate>
</date>
” (cRS)
</materialsCitation>
;
<materialsCitation id="3B463C8DFFA5B11DFAF036D2FB40FC3F" collectingDate="2021-09" country="Philippines" location="Wao" municipality="Mindanao" pageId="8" pageNumber="63" specimenCount="1" specimenCount-female="1" stateProvince="Lanao del Sur" typeStatus="paratype">
<specimenCount id="9D28FD59FFA5B11DFAF036D2FC0EFC3F" count="1" pageId="8" pageNumber="63" type="female">1 female</specimenCount>
idem but “
<date id="FF901010FFA5B11DFB9536B2FB17FC3F" box="[1062,1154,961,984]" pageId="8" pageNumber="63" value="2021-09">
<collectingDate id="EFD4E9F8FFA5B11DFB9536B2FB17FC3F" box="[1062,1154,961,984]" pageId="8" pageNumber="63" value="2021-09">IX.2021</collectingDate>
</date>
” (cRS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA5B11DFCE13173FC36FB5F" pageId="8" pageNumber="63" type="diagnosis">
<paragraph id="8B9136D0FFA5B11DFCE13173FC36FB5F" blockId="8.[850,1361,1024,1208]" pageId="8" pageNumber="63">
<emphasis id="B95AEAC2FFA5B11DFCE13173FC51FBFF" bold="true" box="[850,964,1024,1048]" pageId="8" pageNumber="63">Diagnosis.</emphasis>
Easily distinguishable from all known species in the Oriental region for its combination of dark bluish elytra and rather trapezoid head but without a groove connecting the fovea behind the eye with the postorbital pore, as in
<taxonomicName id="4C2E4D53FFA5B11DFB4231F2FC09FB5F" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="panumanod">
<emphasis id="B95AEAC2FFA5B11DFB4231F2FC09FB5F" italics="true" pageId="8" pageNumber="63">H. panumanod</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA5B11FFCE13193FD41F93B" lastPageId="10" lastPageNumber="65" pageId="8" pageNumber="63" type="description">
<paragraph id="8B9136D0FFA5B11DFCE13193FC49FA9F" blockId="8.[850,1361,1248,1400]" pageId="8" pageNumber="63">
<emphasis id="B95AEAC2FFA5B11DFCE13193FC4CFB1F" bold="true" box="[850,985,1248,1272]" pageId="8" pageNumber="63">Description.</emphasis>
Body length: 12,114,2 mm. Width/ length of head: 0.96; width/length of pronotum: 1.07; length/ width of elytra: 1.63­ 1.65. Body uniformly black, pronotum and elytra with slight bluish luster.
</paragraph>
<paragraph id="8B9136D0FFA5B11DFCE130D2FAC5F93F" blockId="8.[850,1361,1441,1752]" pageId="8" pageNumber="63">Head strongly trapezoid and depressed, slightly narrower than the pronotum. Eyes moderately large and markedly protruded, 0.5 times as long as temples; temples protruding; posterior border almost perpendicula. The shape of the head is slightly different in the male specimens compared to the female ones: the temples are more protruding in the males than in the females, indicating a probable sexual dimorphism in this character. Constriction of neck weak and shallow, neck</paragraph>
<paragraph id="8B9136D0FFA4B11CFD8F35E2FC38FF43" blockId="9.[572,941,145,164]" box="[572,941,145,164]" pageId="9" pageNumber="64">
<emphasis id="B95AEAC2FFA4B11CFD8F35E2FC38FF43" box="[572,941,145,164]" italics="true" pageId="9" pageNumber="64">
Anichtchenko A., Sciaky R.,
<taxonomicName id="4C2E4D53FFA4B11CFC8435E2FCECFF43" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[823,889,145,164]" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="genus">Medina</taxonomicName>
M.N.
</emphasis>
</paragraph>
<paragraph id="8B9136D0FFA4B11CFF5035B2FD18FD9F" blockId="9.[227,738,193,632]" pageId="9" pageNumber="64">
narrow. Mandibles elongate, almost straight in basal two thirds, incurved towards apex. Palpi elongate, narrowed towards apex, smooth.
<collectingRegion id="49EAF832FFA4B11CFD143472FE99FEDF" country="Myanmar" name="Chin" pageId="9" pageNumber="64">Mentum</collectingRegion>
with wide triangular tooth, bisetose.Antenna short, attaining the lateral pore of pronotum; three basal antennomeres and basal half of antennomere 4 glabrous. Clypeal suture shallow, laterally indistinct; clypeus slightly emarginate at apex. Labrum with deep wide triangular incision, asymmetric, right lobe longer. Frons with two irregularly shaped, shallow impressions. Supraorbital pore connected by a shallow groove with the postorbital pore. Surface with extremely fine transverse microreticulation, iridescent.
</paragraph>
<paragraph id="8B9136D0FFA4B11CFCB535B2FC77FEFF" blockId="9.[774,1285,193,280]" pageId="9" pageNumber="64">of 1213 pores. Intervals impunctate and with weak microreticulation, consisting of polygonal meshes, very glossy.</paragraph>
<paragraph id="8B9136D0FFA4B11CFCB53432FE3CFB3F" blockId="9.[774,1286,321,440]" lastBlockId="9.[227,738,673,1240]" pageId="9" pageNumber="64">Legs of average size, tarsomere 3 of all tarsi two times shorter than basal one; tarsomere 4 wide and very deeply excised, with dense brush­like pilosity on ventral surface. Tarsal claws edentate. Pronotum wider than the head, obovate, widest at apical quarter; dorsal surface depressed. Anterior margin slightly convex, anterior angles completely rounded. Lateral margins evenly curved, immediately in front of base slightly excised; base straight, basal angles obtuse and slightly produced. Median line deep, impunctate, almost attaining anterior margin and base; both transverse sulci obsolete. Basal grooves starts right from the posterior angles in form of short, smooth, diagonal impressions. Anterior margin and base not margined, lateral borders narrowly margined, marginal gutter slightly wider near lateral pore. Anterior lateral seta situated at widest point, posterior lateral seta absent. Dorsal surface smooth, with some very shallow irregular transverse sulci, with extremely fine transverse microreticulation, slightly iridescent.</paragraph>
<paragraph id="8B9136D0FFA4B11FFF503072FDDCF9DB" blockId="9.[227,737,1281,1752]" lastBlockId="10.[302,812,1541,1596]" lastPageId="10" lastPageNumber="65" pageId="9" pageNumber="64">Elytra elongate, markedly depressed, parallel sided. Base oblique, humeri rounded; lateral margins straight. Basal margin straight and forming an angle at the junction with lateral margin, at position between striae 4 and 5. Scutellar stria long and connected with stria 1. All striae deeply impressed and moderately punctate. Intervals flat and smooth. Scutellar seta situated at origin of stria 1. Interval 3 with three setiferous pores: anterior one situated in the stria 3 at the level of the end of scutellar stria; median one situated in apical third of elytra near stria 2; posterior one situated close to apex, near stria 2. Interval 5 of elytra with one pore located opposite the middle between discal and apical pores. Marginal series consisting Three basal tarsomeres of all legs triangular, very wide and dorsally smooth.</paragraph>
<caption id="DF516658FFA4B11CFCB533EEFBFDF933" pageId="9" pageNumber="64" startId="9.[774,813,1693,1716]" targetBox="[774,1281,481,1657]" targetPageId="9" targetType="figure">
<paragraph id="8B9136D0FFA4B11CFCB533EEFBFDF933" blockId="9.[774,1285,1693,1748]" pageId="9" pageNumber="64">
Fig. 6. Habitus of
<taxonomicName id="4C2E4D53FFA4B11CFC7533EDFB52F952" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[966,1223,1694,1717]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="nigrocyanea" status="sp. nov.">
<emphasis id="B95AEAC2FFA4B11CFC7533EDFB52F952" box="[966,1223,1694,1717]" italics="true" pageId="9" pageNumber="64">Hexagonia nigrocyanea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26957B9FFA4B11CFB7E33EEFA90F953" box="[1229,1285,1693,1716]" pageId="9" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
Holotype, male. Scale bar 5 mm.
</paragraph>
</caption>
<caption id="DF516658FFA7B11FFE9D3004FAC4FA29" pageId="10" pageNumber="65" startId="10.[302,352,1399,1422]" targetBox="[307,1358,191,1362]" targetPageId="10" targetType="figure">
<paragraph id="8B9136D0FFA7B11FFE9D3004FAC4FA29" blockId="10.[302,1361,1398,1487]" pageId="10" pageNumber="65">
Figs. 711. Aedeagus of
<taxonomicName id="4C2E4D53FFA7B11FFD8F300BFD27FA68" authorityName="Kirby" authorityYear="1825" box="[572,690,1400,1423]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B95AEAC2FFA7B11FFD8F300BFD27FA68" box="[572,690,1400,1423]" italics="true" pageId="10" pageNumber="65">Hexagonia</emphasis>
</taxonomicName>
, A dorsal view, B lateral view. 7
<taxonomicName id="4C2E4D53FFA7B11FFBE9300BFEF0FA49" authority="JedliEka, 1935" authorityName="JedliEka" authorityYear="1935" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B95AEAC2FFA7B11FFBE9300BFB7FFA68" box="[1114,1258,1400,1423]" italics="true" pageId="10" pageNumber="65">H. andrewesi</emphasis>
<bibRefCitation id="EFBF4B21FFA7B11FFB423004FEF0FA49" author="JedliEka A." pageId="10" pageNumber="65" refId="ref6710" refString="JedliEka A., 1935. Novy Druh rodu Hexagonia (Col. Carab.). &quot; asopis &quot; eskoslovenske spoleEnosti entomologicke 32: 89." type="journal volume" year="1935">JedliËka, 1935</bibRefCitation>
</taxonomicName>
, Holotype; 8
<taxonomicName id="4C2E4D53FFA7B11FFDB230EBFC8CFA49" authority="JedliEka, 1936" authorityName="JedliEka" authorityYear="1936" box="[513,793,1430,1455]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="castanea" status="syn. n.">
<emphasis id="B95AEAC2FFA7B11FFDB230EBFDEBFA48" box="[513,638,1432,1455]" italics="true" pageId="10" pageNumber="65">H. castanea</emphasis>
<bibRefCitation id="EFBF4B21FFA7B11FFD3130E4FC8CFA49" author="JedliEka A." box="[642,793,1430,1455]" pageId="10" pageNumber="65" pagination="51 - 55" refId="ref6737" refString="JedliEka A., 1936. Neue Carabiden aus Ostasien. (XI Teil). Entomologisches Nachrichtenblatt 10 (1): 51 - 55." type="journal article" year="1936">JedliËka, 1936</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A26957B9FFA7B11FFCAD30E4FCF7FA49" box="[798,866,1431,1454]" pageId="10" pageNumber="65">syn. n.</taxonomicNameLabel>
Cotype; 9
<taxonomicName id="4C2E4D53FFA7B11FFC5030EBFB4BFA49" authority="Dupuis, 1912" authorityName="Dupuis" authorityYear="1912" box="[995,1246,1431,1455]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B95AEAC2FFA7B11FFC5030EBFBDEFA48" box="[995,1099,1432,1455]" italics="true" pageId="10" pageNumber="65">H. sauteri</emphasis>
Dupuis, 1912
</taxonomicName>
, Holotype. Scale bar 1 mm. 10
<taxonomicName id="4C2E4D53FFA7B11FFDBA30CBFDE8FA28" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[521,637,1464,1487]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="amicitia" status="sp. n.">
<emphasis id="B95AEAC2FFA7B11FFDBA30CBFDE8FA28" box="[521,637,1464,1487]" italics="true" pageId="10" pageNumber="65">H. amicitia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26957B9FFA7B11FFD3230C4FD2CFA29" box="[641,697,1463,1486]" pageId="10" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
, Holotype; 11
<taxonomicName id="4C2E4D53FFA7B11FFCD330CBFC6BFA28" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[864,1022,1464,1487]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="nigrocyanea" status="sp. n.">
<emphasis id="B95AEAC2FFA7B11FFCD330CBFC6BFA28" box="[864,1022,1464,1487]" italics="true" pageId="10" pageNumber="65">H. nigrocyanea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26957B9FFA7B11FFBB030C4FBAFFA29" box="[1027,1082,1463,1486]" pageId="10" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
Holotype. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="8B9136D0FFA7B11FFE9D3316FD41F93B" blockId="10.[302,813,1637,1756]" pageId="10" pageNumber="65">Aedeagus (Fig. ??) elongate and very thin in lateral view. Apex of median lobe moderately long and slightly bent to the lower side. Endophallus with large semicircular sclerite.</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA7B11EFCE13376FC38FF43" lastPageId="11" lastPageNumber="66" pageId="10" pageNumber="65" type="distribution">
<paragraph id="8B9136D0FFA7B11FFCE13376FBCFF925" blockId="10.[850,1361,1541,1730]" pageId="10" pageNumber="65">
<emphasis id="B95AEAC2FFA7B11FFCE13376FC72F9FA" bold="true" box="[850,999,1541,1565]" pageId="10" pageNumber="65">Distribution.</emphasis>
Only known from the island of Mindanao, in three different localities in the provinces of
<collectingRegion id="49EAF832FFA7B11FFC6A3334FBD0F9B9" box="[985,1093,1607,1630]" country="Philippines" name="Bukidnon" pageId="10" pageNumber="65">Bukidnon</collectingRegion>
,
<collectingRegion id="49EAF832FFA7B11FFBFD3334FB2DF9B9" box="[1102,1208,1607,1630]" country="Philippines" name="Sarangani" pageId="10" pageNumber="65">Sarangani</collectingRegion>
and
<collectingRegion id="49EAF832FFA7B11FFB5B3334FCE3F998" country="Philippines" name="Lanao del Sur" pageId="10" pageNumber="65">Lanao del Sur</collectingRegion>
, therefore occupying almost all the island of Mindanao. We can expect to find it in different areas of the same island.
</paragraph>
<paragraph id="8B9136D0FFA6B11EFD8F35E2FC38FF43" blockId="11.[572,941,145,164]" box="[572,941,145,164]" pageId="11" pageNumber="66">
<emphasis id="B95AEAC2FFA6B11EFD8F35E2FC38FF43" box="[572,941,145,164]" italics="true" pageId="11" pageNumber="66">
Anichtchenko A., Sciaky R.,
<taxonomicName id="4C2E4D53FFA6B11EFC8435E2FCECFF43" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[823,889,145,164]" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="66" phylum="Arthropoda" rank="genus">Medina</taxonomicName>
M.N.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA6B11EFF5035B3FEBFFABF" pageId="11" pageNumber="66" type="etymology">
<paragraph id="8B9136D0FFA6B11EFF5035B3FD5FFEFF" blockId="11.[227,737,192,280]" pageId="11" pageNumber="66">
<emphasis id="B95AEAC2FFA6B11EFF5035B3FEF1FF3F" bold="true" box="[227,356,192,216]" pageId="11" pageNumber="66">Etymology.</emphasis>
This specific epithet alludes to the peculiar color of this species: the Latin term
<taxonomicName id="4C2E4D53FFA6B11EFF503472FEFCFEFF" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[227,361,257,280]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="nigrocyanea">
<emphasis id="B95AEAC2FFA6B11EFF503472FEFCFEFF" box="[227,361,257,280]" italics="true" pageId="11" pageNumber="66">nigrocyanea</emphasis>
</taxonomicName>
means, in fact, black and bluish.
</paragraph>
<paragraph id="8B9136D0FFA6B11EFF503433FDA3FE1F" blockId="11.[227,737,320,504]" pageId="11" pageNumber="66">
<emphasis id="B95AEAC2FFA6B11EFF503433FEC7FEBF" bold="true" box="[227,338,320,344]" pageId="11" pageNumber="66">Affinities.</emphasis>
This species is the largest of the genus in the
<collectingCountry id="F3397640FFA6B11EFE9A3412FE37FE9F" box="[297,418,353,376]" name="Philippines" pageId="11" pageNumber="66">Philippines</collectingCountry>
and one of the largest in the whole world. It seems related to the sympatric
<taxonomicName id="4C2E4D53FFA6B11EFF5034D2FE17FE5F" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[227,386,417,440]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="panumanod">
<emphasis id="B95AEAC2FFA6B11EFF5034D2FE17FE5F" box="[227,386,417,440]" italics="true" pageId="11" pageNumber="66">H. panumanod</emphasis>
</taxonomicName>
, although well distinct. As far as we know these are the only two metallic species of
<taxonomicName id="4C2E4D53FFA6B11EFF4C3492FEE0FE1F" authorityName="Kirby" authorityYear="1825" box="[255,373,481,504]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B95AEAC2FFA6B11EFF4C3492FEE0FE1F" box="[255,373,481,504]" italics="true" pageId="11" pageNumber="66">Hexagonia</emphasis>
</taxonomicName>
known until now.
</paragraph>
<paragraph id="8B9136D0FFA6B11EFF503753FD77FD1F" blockId="11.[227,738,544,760]" pageId="11" pageNumber="66">
<emphasis id="B95AEAC2FFA6B11EFF503753FE38FDDF" bold="true" box="[227,429,544,568]" pageId="11" pageNumber="66">General remarks.</emphasis>
The genus
<taxonomicName id="4C2E4D53FFA6B11EFD9B3752FD0BFDDF" authorityName="Kirby" authorityYear="1825" box="[552,670,545,568]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B95AEAC2FFA6B11EFD9B3752FD0BFDDF" box="[552,670,545,568]" italics="true" pageId="11" pageNumber="66">Hexagonia</emphasis>
</taxonomicName>
in the
<collectingCountry id="F3397640FFA6B11EFF503732FE09FDBF" box="[227,412,577,600]" name="Philippines" pageId="11" pageNumber="66">Philippine islands</collectingCountry>
seems mainly distributed in the island of Mindanao and its vicinity, in fact, four out of the five species until now known from this country have been found here, while only
<taxonomicName id="4C2E4D53FFA6B11EFD2637D2FE9DFD3F" authorityName="Dupuis" authorityYear="1912" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B95AEAC2FFA6B11EFD2637D2FE9DFD3F" italics="true" pageId="11" pageNumber="66">H. sauteri</emphasis>
</taxonomicName>
is known from the island of Luzon, beyond
<collectingCountry id="F3397640FFA6B11EFF503792FEA4FD1F" box="[227,305,737,760]" name="Taiwan" pageId="11" pageNumber="66">Taiwan</collectingCountry>
, from where it had been initially described.
</paragraph>
<paragraph id="8B9136D0FFA6B11EFF503652FE0FFB9F" blockId="11.[227,737,801,1144]" pageId="11" pageNumber="66">
<bibRefCitation id="EFBF4B21FFA6B11EFF503652FE75FCDF" author="Darlington P. J. Jr." box="[227,480,801,824]" pageId="11" pageNumber="66" pagination="1 - 253" refId="ref6640" refString="Darlington P. J. Jr. 1968. The carabid beetles of New Guinea. Part III. Harpalinae (continued): Perigonini to Pseudomorphini. Bulletin of The Museum of Comparative Zoology 137: 1 - 253." type="journal article" year="1968">Darlington (1968: 203)</bibRefCitation>
wrote when describing
<taxonomicName id="4C2E4D53FFA6B11EFF503632FE3FFCBF" authorityName="Darlington" authorityYear="1968" box="[227,426,833,856]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="papua">
<emphasis id="B95AEAC2FFA6B11EFF503632FE3FFCBF" box="[227,426,833,856]" italics="true" pageId="11" pageNumber="66">Hexagonia papua</emphasis>
</taxonomicName>
: “One or more similar but apparently undescribed species occur in the
<collectingCountry id="F3397640FFA6B11EFD073612FED4FC7F" name="Philippines" pageId="11" pageNumber="66">Philippines</collectingCountry>
.”. This sentence seems to prove that he had seen some specimens from this area belonging to the group of reddish brown species with a black spot at the elytral apex. These species are very numerous in South­East Asia, and only seeing abundant material could allow to decide whether they are new species or species known from other areas.
</paragraph>
<paragraph id="8B9136D0FFA6B11EFF5031D2FEBFFABF" blockId="11.[227,737,1185,1368]" pageId="11" pageNumber="66">
In any case, we think that the knowledge on this genus in the
<collectingCountry id="F3397640FFA6B11EFEDB31B2FE77FB3F" box="[360,482,1217,1240]" name="Philippines" pageId="11" pageNumber="66">Philippines</collectingCountry>
are still incomplete and that certainly some new species and new data on the species already known will be discovered as the faunistic exploration of this archipelago continues.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,177 @@
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<mods:title id="63D468CBEB4A2350099C313DC22B8DEA">Taxonomic notes on the genus Hexagonia Kirby, 1825 with description of three new species from the Philippines (Coleoptera: Carabidae: Hexagoniini)</mods:title>
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<treatment id="038787C6FFA8B112FCB537F3FDA4FEDF" LSID="urn:lsid:plazi:treatment:038787C6FFA8B112FCB537F3FDA4FEDF" httpUri="http://treatment.plazi.org/id/038787C6FFA8B112FCB537F3FDA4FEDF" lastPageId="7" lastPageNumber="62" pageId="5" pageNumber="60">
<subSubSection id="C334655BFFA8B110FCB537F3FC3DFD7F" box="[774,936,640,664]" pageId="5" pageNumber="60" type="nomenclature">
<paragraph id="8B9136D0FFA8B110FCB537F3FC3DFD7F" blockId="5.[774,1285,608,696]" box="[774,936,640,664]" pageId="5" pageNumber="60">
<heading id="D0D981BCFFA8B110FCB537F3FC3DFD7F" bold="true" box="[774,936,640,664]" fontSize="9" level="2" pageId="5" pageNumber="60" reason="6">
<emphasis id="B95AEAC2FFA8B110FCB537F3FC3DFD7F" bold="true" box="[774,936,640,664]" pageId="5" pageNumber="60">
<taxonomicName id="4C2E4D53FFA8B110FCB537F3FCF5FD7F" ID-CoL="5MH6" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[774,864,640,664]" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus" status="sp. n.">Medina</taxonomicName>
,
<taxonomicNameLabel id="A26957B9FFA8B110FCDF37F3FC3DFD7F" box="[876,936,640,664]" pageId="5" pageNumber="60" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA8B110FCB537D2FC1FFD5F" box="[774,906,673,696]" pageId="5" pageNumber="60" type="description">
<paragraph id="8B9136D0FFA8B110FCB537D2FC1FFD5F" blockId="5.[774,1285,608,696]" box="[774,906,673,696]" pageId="5" pageNumber="60">
(
<figureCitation id="13152A55FFA8B110FCBD37D2FCCFFD5F" box="[782,858,673,696]" captionStart="Fig" captionStartId="6.[302,338,1219,1242]" captionTargetBox="[306,811,333,1183]" captionTargetId="figure-620@6.[302,812,333,1183]" captionTargetPageId="6" captionText="Fig. 5. Habitus of Hexagonia amicitia sp. n. Holotype, male. Scale bar 5 mm." pageId="5" pageNumber="60">Figs. 5</figureCitation>
,
<figureCitation id="13152A55FFA8B110FCD537D2FC17FD5F" box="[870,898,673,696]" captionStart="Figs" captionStartId="10.[302,352,1399,1422]" captionTargetBox="[307,1358,191,1362]" captionTargetId="figure-177@10.[302,1360,189,1366]" captionTargetPageId="10" captionText="Figs. 711. Aedeagus of Hexagonia, A dorsal view, B lateral view. 7 H. andrewesi JedliËka, 1935, Holotype; 8 H. castanea JedliËka, 1936 syn. n. Cotype; 9 H. sauteri Dupuis, 1912, Holotype. Scale bar 1 mm. 10 H. amicitia sp. n., Holotype; 11 H. nigrocyanea sp. n. Holotype. Scale bar 1 mm." pageId="5" pageNumber="60">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA8B110FCB53793FC6CFCBF" pageId="5" pageNumber="60" type="materials_examined">
<paragraph id="8B9136D0FFA8B110FCB53793FC6CFCBF" blockId="5.[774,1285,736,856]" pageId="5" pageNumber="60">
<emphasis id="B95AEAC2FFA8B110FCB53793FCE1FD1F" bold="true" box="[774,884,736,760]" pageId="5" pageNumber="60">Material:</emphasis>
<materialsCitation id="3B463C8DFFA8B110FCCE3792FB68FCFF" collectingDate="2018-10" collectionCode="DUBC" location="Mindanao" municipality="Maguing" pageId="5" pageNumber="60" specimenCount="1" stateProvince="Lanao del Sur" typeStatus="holotype">
<typeStatus id="54958872FFA8B110FCCE3792FC71FD1F" box="[893,996,737,760]" pageId="5" pageNumber="60" type="holotype">Holotype</typeStatus>
, male
<collectingMunicipality id="6BF5ACAAFFA8B110FBFD3792FB21FD1F" box="[1102,1204,737,760]" pageId="5" pageNumber="60">Maguing</collectingMunicipality>
,
<collectingRegion id="49EAF832FFA8B110FB713792FCC8FCFF" country="Philippines" name="Lanao del Sur" pageId="5" pageNumber="60">Lanao Del Sur</collectingRegion>
,
<location id="8EF1600BFFA8B110FCDD3672FC49FCFF" LSID="urn:lsid:plazi:treatment:038787C6FFA8B112FCB537F3FDA4FEDF:8EF1600BFFA8B110FCDD3672FC49FCFF" box="[878,988,769,792]" municipality="Maguing" name="Mindanao" pageId="5" pageNumber="60" stateProvince="Lanao del Sur">Mindanao</location>
|
<date id="FF901010FFA8B110FC443672FB1CFCFF" box="[1015,1161,769,792]" pageId="5" pageNumber="60" value="2018-10">
<collectingDate id="EFD4E9F8FFA8B110FC443672FB1CFCFF" box="[1015,1161,769,792]" pageId="5" pageNumber="60" value="2018-10">october 2018</collectingDate>
</date>
” (
<collectionCode id="ED3FAE15FFA8B110FB1A3672FB61FCFF" box="[1193,1268,769,792]" pageId="5" pageNumber="60">DUBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B463C8DFFA8B110FCB53652FC60FCBF" collectingDate="2013-11" country="Philippines" location="Mindanao" municipality="Mt. Apo" pageId="5" pageNumber="60" specimenCount="1" typeStatus="paratype">
<typeStatus id="54958872FFA8B110FCB53652FCF2FCDF" box="[774,871,801,824]" pageId="5" pageNumber="60" type="paratype">Paratype</typeStatus>
, female
<collectingCountry id="F3397640FFA8B110FC683652FBCCFCDF" box="[987,1113,801,824]" name="Philippines" pageId="5" pageNumber="60">Philippines</collectingCountry>
,
<location id="8EF1600BFFA8B110FBD13652FB5AFCDF" LSID="urn:lsid:plazi:treatment:038787C6FFA8B112FCB537F3FDA4FEDF:8EF1600BFFA8B110FBD13652FB5AFCDF" box="[1122,1231,801,824]" country="Philippines" municipality="Mt. Apo" name="Mindanao" pageId="5" pageNumber="60">Mindanao</location>
|
<collectingMunicipality id="6BF5ACAAFFA8B110FB6C3652FCA1FCBF" pageId="5" pageNumber="60">Mt. Apo</collectingMunicipality>
|
<date id="FF901010FFA8B110FCFB3632FC37FCBF" box="[840,930,833,856]" pageId="5" pageNumber="60" value="2013-11">
<collectingDate id="EFD4E9F8FFA8B110FCFB3632FC37FCBF" box="[840,930,833,856]" pageId="5" pageNumber="60" value="2013-11">XI.2013</collectingDate>
</date>
” (cRS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA8B110FCB536F3FC5AFC3F" pageId="5" pageNumber="60" type="diagnosis">
<paragraph id="8B9136D0FFA8B110FCB536F3FC5AFC3F" blockId="5.[774,1285,896,984]" pageId="5" pageNumber="60">
<emphasis id="B95AEAC2FFA8B110FCB536F3FCEBFC7F" bold="true" box="[774,894,896,920]" pageId="5" pageNumber="60">Diagnosis.</emphasis>
This species is very similar to
<taxonomicName id="4C2E4D53FFA8B110FB5836F2FCE1FC5F" authorityName="JedliEka" authorityYear="1935" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B95AEAC2FFA8B110FB5836F2FCE1FC5F" italics="true" pageId="5" pageNumber="60">H. andrewesi</emphasis>
</taxonomicName>
; the diagnostic differences are listed under that species.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA8B113FCB53173FAC4FAFF" lastPageId="6" lastPageNumber="61" pageId="5" pageNumber="60" type="description">
<paragraph id="8B9136D0FFA8B110FCB53173FCC7FB7F" blockId="5.[774,1285,1024,1176]" pageId="5" pageNumber="60">
<emphasis id="B95AEAC2FFA8B110FCB53173FC07FBFF" bold="true" box="[774,914,1024,1048]" pageId="5" pageNumber="60">Description.</emphasis>
Body length: 8.910.0 mm. Width/ length of head: 1; Width/length of pronotum: 1.06; length/ width of elytra: 1.7. Body, legs and palpi dark brown or uniformly black; antennae brown.
</paragraph>
<paragraph id="8B9136D0FFA8B113FCB531B2FBCFFEBF" blockId="5.[774,1286,1217,1752]" lastBlockId="6.[850,1361,193,344]" lastPageId="6" lastPageNumber="61" pageId="5" pageNumber="60">
Head as long as wide, depressed, slightly narrower than the prothorax. Eye moderately large and markedly protruded, 0.75 times as long as temples; temples not protruding; gradually rounded to the constriction of the neck. Mandibles elongate, slightly incurved in basal two thirds, then strongly incurved towards apex. Palpi elongate, narrowed towards apex, smooth.
<collectingRegion id="49EAF832FFA8B110FC4830D2FBC3FA5F" box="[1019,1110,1441,1464]" country="Myanmar" name="Chin" pageId="5" pageNumber="60">Mentum</collectingRegion>
with wide triangular tooth, bisetose. Antenna relatively long, last antennomere go beyond the middle lenght of pronotum; three basal antennomeres and basal half of antennomere 4 glabrous. Clypeus with two pores near anterior angles; with wide, transverse and rugate depression, clypeal suture very deep and complete; apical margin of clypeus straight. Labrum transverse and symmetric, slightly bisinuate apically. Frons with two large irregularly shaped, deep impressions. Posterior half of impressions rugate. Postorbital pore isolated. Surface with indistinct microreticulation, shiny. pore and completely obsolete in basal fifth.Anterior lateral seta situated at widest point, posterior lateral seta absent. Dorsal surface smooth, with very shallow transverse sulci. Microreticulation transverse and very fine.
</paragraph>
<paragraph id="8B9136D0FFABB113FCE134F2FAA4FC7F" blockId="6.[850,1361,385,920]" pageId="6" pageNumber="61">Elytra elongate, markedly depressed, parallel sided. Base straight, humeri widely rounded; lateral margins straight. Basal margin concave, connected at an obtuse angle with lateral margin at position between striae 4 and 5. Scutellar stria long, not connected with first stria.All striae deeply impressed and almost smooth, impunctate. Intervals flat and smooth. Scutellar seta situated at base of stria 1. Interval 3 with three setiferous pores, the anterior pore situated in the 3rd stria at the level of apice of scutellar stria; the discal pore situated immediately behind the middle of elytra near second stria; posterior one situated in apical fifth, near the stria 2. Interval 5 of elytra with one pore. Marginal series consisting of 1415 pores. Intervals impunctate and with well visible microreticulation, consisting of polygonal meshes, glossy.</paragraph>
<paragraph id="8B9136D0FFABB113FCE136B2FBAAFB9F" blockId="6.[850,1361,961,1144]" pageId="6" pageNumber="61">Legs of average size. Three basal tarsomeres of all legs triangular and wide; dorsally with longitudinal rugosity, especially in mid­ and hind tarsa; tarsomere 4 wide and very deeply excised, with dense brush­like pilosity on ventral surface. Tarsal claws edentate.</paragraph>
<caption id="DF516658FFABB113FE9D31B0FD05FB1D" pageId="6" pageNumber="61" startId="6.[302,338,1219,1242]" targetBox="[306,811,333,1183]" targetPageId="6" targetType="figure">
<paragraph id="8B9136D0FFABB113FE9D31B0FD05FB1D" blockId="6.[302,813,1219,1274]" pageId="6" pageNumber="61">
Fig. 5. Habitus of
<taxonomicName id="4C2E4D53FFABB113FDB631B0FD76FB3D" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[517,739,1219,1242]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="amicitia" status="sp. nov.">
<emphasis id="B95AEAC2FFABB113FDB631B0FD76FB3D" box="[517,739,1219,1242]" italics="true" pageId="6" pageNumber="61">Hexagonia amicitia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26957B9FFABB113FD5D31B0FCB8FB3D" box="[750,813,1219,1242]" pageId="6" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
Holotype, male. Scale bar 5 mm.
</paragraph>
</caption>
<paragraph id="8B9136D0FFABB113FE9D3042FAC4FAFF" blockId="6.[302,813,1329,1743]" lastBlockId="6.[850,1361,1185,1304]" pageId="6" pageNumber="61">
Pronotum wider than head, widest at apical third, almost as long as wide; dorsal surface depressed. Anterior margin straight, anterior angles completely rounded. Lateral margins evenly curved, in basal fifth of the lenght parallel sided, with long sinuation before hind angles; basal angles right, not produced; base straight. Median line deep, impunctate, not reaching anterior margin of pronotum; anterior transverse sulcus obsoleted, posterior sulcus wide and shallow, connecting apices of very shallow basal grooves. Apex and base not margined, lateral borders narrowly margined, marginal sulcus slightly wider near lateral Aedeagus (
<figureCitation id="13152A55FFABB113FC6031D2FBB1FB5F" box="[979,1060,1185,1208]" captionStart="Figs" captionStartId="10.[302,352,1399,1422]" captionTargetBox="[307,1358,191,1362]" captionTargetId="figure-177@10.[302,1360,189,1366]" captionTargetPageId="10" captionText="Figs. 711. Aedeagus of Hexagonia, A dorsal view, B lateral view. 7 H. andrewesi JedliËka, 1935, Holotype; 8 H. castanea JedliËka, 1936 syn. n. Cotype; 9 H. sauteri Dupuis, 1912, Holotype. Scale bar 1 mm. 10 H. amicitia sp. n., Holotype; 11 H. nigrocyanea sp. n. Holotype. Scale bar 1 mm." pageId="6" pageNumber="61">Fig. 10</figureCitation>
) elongate, ventral surface almost straight to apex. Apex of median lobe moderately long and slightly bent to the lower side. Endophallus with big semicircular sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFABB113FCE13033FA81FA3F" pageId="6" pageNumber="61" type="distribution">
<paragraph id="8B9136D0FFABB113FCE13033FA81FA3F" blockId="6.[850,1360,1344,1496]" pageId="6" pageNumber="61">
<emphasis id="B95AEAC2FFABB113FCE13033FC73FABF" bold="true" box="[850,998,1344,1368]" pageId="6" pageNumber="61">Distribution.</emphasis>
Known from two different localities of the island of Mindanao, one situated in the province of
<collectingRegion id="49EAF832FFABB113FBB230F2FB0FFA7F" box="[1025,1178,1409,1432]" country="Philippines" name="Lanao del Sur" pageId="6" pageNumber="61">Lanao del Sur</collectingRegion>
, the other in the province of
<collectingRegion id="49EAF832FFABB113FC6130D2FBF8FA5F" box="[978,1133,1441,1464]" country="Philippines" name="Davao del Sur" pageId="6" pageNumber="61">Davao del Sur</collectingRegion>
; probably it is widely diffused in the whole island of Mindanao.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFABB112FCE13373FDA4FEDF" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61" type="etymology">
<paragraph id="8B9136D0FFABB113FCE13373FB89F99F" blockId="6.[850,1361,1536,1656]" pageId="6" pageNumber="61">
<emphasis id="B95AEAC2FFABB113FCE13373FC47F9FF" bold="true" box="[850,978,1536,1560]" pageId="6" pageNumber="61">Etymology.</emphasis>
The specific epithet derives from the Latin
<taxonomicName id="4C2E4D53FFABB113FC213352FC62F9DF" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[914,1015,1569,1592]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="amicitia">“amicitia</taxonomicName>
” (fem.) friendship, in allusion to significance of friendship and collaboration in science and life.
</paragraph>
<paragraph id="8B9136D0FFABB113FCE133D3FAC5F93F" blockId="6.[850,1360,1696,1752]" pageId="6" pageNumber="61">
<emphasis id="B95AEAC2FFABB113FCE133D3FC51F95F" bold="true" box="[850,964,1696,1720]" pageId="6" pageNumber="61">Affinities.</emphasis>
This species is similar to
<taxonomicName id="4C2E4D53FFABB113FB5433D2FC14F93F" authorityName="JedliEka" authorityYear="1935" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B95AEAC2FFABB113FB5433D2FC14F93F" italics="true" pageId="6" pageNumber="61">H. andrewesi</emphasis>
</taxonomicName>
, although smaller. The overall aspect is very
</paragraph>
<paragraph id="8B9136D0FFAAB112FD8F35E2FC38FF43" blockId="7.[572,941,145,164]" box="[572,941,145,164]" pageId="7" pageNumber="62">
<emphasis id="B95AEAC2FFAAB112FD8F35E2FC38FF43" box="[572,941,145,164]" italics="true" pageId="7" pageNumber="62">
Anichtchenko A., Sciaky R.,
<taxonomicName id="4C2E4D53FFAAB112FC8435E2FCECFF43" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[823,889,145,164]" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">Medina</taxonomicName>
M.N.
</emphasis>
</paragraph>
<paragraph id="8B9136D0FFAAB112FF5035B2FDA4FEDF" blockId="7.[227,737,193,312]" pageId="7" pageNumber="62">similar, but some details ot the elytral punctuation, the different length/width ratio of the elytra, and shape of the head and of the pronotum allow to distinguish between the two.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,159 @@
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<mods:title id="108FAFA2600D8F0CE12BEA2F2D86E755">Taxonomic notes on the genus Hexagonia Kirby, 1825 with description of three new species from the Philippines (Coleoptera: Carabidae: Hexagoniini)</mods:title>
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<subSubSection id="C334655BFFAAB112FF5034D3FE40FE5F" box="[227,469,416,440]" pageId="7" pageNumber="62" type="nomenclature">
<paragraph id="8B9136D0FFAAB112FF5034D3FE40FE5F" blockId="7.[227,737,384,440]" box="[227,469,416,440]" pageId="7" pageNumber="62">
<emphasis id="B95AEAC2FFAAB112FF5034D3FEEBFE5F" bold="true" box="[227,382,416,440]" pageId="7" pageNumber="62">
<taxonomicName id="4C2E4D53FFAAB112FF5034D3FEAEFE5F" ID-CoL="5MH6" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[227,315,416,440]" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="genus" status="sp. n.">Medina</taxonomicName>
<taxonomicNameLabel id="A26957B9FFAAB112FEF234D3FEEBFE5F" box="[321,382,416,440]" pageId="7" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13152A55FFAAB112FE3E34D2FE58FE5F" box="[397,461,417,440]" captionStart="Fig" captionStartId="7.[774,813,1164,1187]" captionTargetBox="[774,1282,193,1129]" captionTargetId="figure-505@7.[774,1284,189,1131]" captionTargetPageId="7" captionText="Fig. 4. Habitus of Hexagonia panumanod sp. n. Holotype, female. Scale bar 5 mm." pageId="7" pageNumber="62">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAAB112FF503493FE23FDBF" pageId="7" pageNumber="62" type="materials_examined">
<paragraph id="8B9136D0FFAAB112FF503493FE23FDBF" blockId="7.[227,737,480,600]" pageId="7" pageNumber="62">
<emphasis id="B95AEAC2FFAAB112FF503493FE12FE1F" bold="true" box="[227,391,480,504]" pageId="7" pageNumber="62">Type material:</emphasis>
<materialsCitation id="3B463C8DFFAAB112FE3F3492FE23FDBF" collectingDate="2022-05-04" collectionCode="DUBC" country="Philippines" county="Eureka Mt." elevation="730" latitude="8.691617" location="Mindanao" longLatPrecision="1" longitude="125.23785" municipality="Gingoog" pageId="7" pageNumber="62" specimenCount="1" typeStatus="holotype">
<typeStatus id="54958872FFAAB112FE3F3492FE64FE1F" box="[396,497,481,504]" pageId="7" pageNumber="62" type="holotype">Holotype</typeStatus>
, female
<collectingCountry id="F3397640FFAAB112FDD23492FD48FE1F" box="[609,733,481,504]" name="Philippines" pageId="7" pageNumber="62">Philippines</collectingCountry>
,
<location id="8EF1600BFFAAB112FF503772FECEFDFF" LSID="urn:lsid:plazi:treatment:038787C6FFAAB11DFF5034D3FBBEFE3F:8EF1600BFFAAB112FF503772FECEFDFF" box="[227,347,513,536]" country="Philippines" county="Eureka Mt." latitude="8.691617" longLatPrecision="1" longitude="125.23785" municipality="Gingoog" name="Mindanao" pageId="7" pageNumber="62">Mindanao</location>
,
<collectingMunicipality id="6BF5ACAAFFAAB112FEC33772FE42FDFF" box="[368,471,513,536]" pageId="7" pageNumber="62">Gingoog</collectingMunicipality>
|
<collectingCounty id="62F04E5CFFAAB112FE443772FD17FDFF" box="[503,642,513,536]" pageId="7" pageNumber="62">Eureka Mt.</collectingCounty>
,
<quantity id="4CD69B35FFAAB112FD243772FD4CFDFF" box="[663,729,513,536]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="7" pageNumber="62" unit="m" value="730.0">
<elevation id="0003D1E3FFAAB112FD243772FD4CFDFF" box="[663,729,513,536]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="7" pageNumber="62" unit="m" value="730.0">730m</elevation>
</quantity>
,
<geoCoordinate id="EE1A5017FFAAB112FF503752FEF3FDDF" box="[227,358,545,568]" degrees="8" direction="north" minutes="41.4970" orientation="latitude" pageId="7" pageNumber="62" precision="1" value="8.691617">8°41.4970N</geoCoordinate>
|
<geoCoordinate id="EE1A5017FFAAB112FECB3752FD80FDDF" box="[376,533,545,568]" degrees="125" direction="east" minutes="14.2710" orientation="longitude" pageId="7" pageNumber="62" precision="1" value="125.23785">125°14.2710E</geoCoordinate>
,
<date id="FF901010FFAAB112FD933752FDE9FDDF" box="[544,636,545,568]" pageId="7" pageNumber="62" value="2022-05-04">
<collectingDate id="EFD4E9F8FFAAB112FD933752FDE9FDDF" box="[544,636,545,568]" pageId="7" pageNumber="62" value="2022-05-04">4.v.2022</collectingDate>
</date>
| M.
<taxonomicName id="4C2E4D53FFAAB112FD073752FE84FDBF" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="genus">Medina</taxonomicName>
leg.” (
<collectionCode id="ED3FAE15FFAAB112FEEA3732FE3FFDBF" box="[345,426,577,600]" pageId="7" pageNumber="62">DUBC</collectionCode>
).
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</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAAB112FF5037F3FD25FCFF" pageId="7" pageNumber="62" type="diagnosis">
<paragraph id="8B9136D0FFAAB112FF5037F3FD25FCFF" blockId="7.[227,737,640,792]" pageId="7" pageNumber="62">
<emphasis id="B95AEAC2FFAAB112FF5037F3FEC1FD7F" bold="true" box="[227,340,640,664]" pageId="7" pageNumber="62">Diagnosis.</emphasis>
Easily distinguishable from all known species in the Oriental region in the combination of metallic blue elytra and strongly trapezoid head with the fovea behind the eye connected by a shallow groove with the postorbital pore.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAAB11DFF503633FD36FAFF" lastPageId="8" lastPageNumber="63" pageId="7" pageNumber="62" type="description">
<paragraph id="8B9136D0FFAAB112FF503633FD48FC1F" blockId="7.[227,737,832,1016]" pageId="7" pageNumber="62">
<emphasis id="B95AEAC2FFAAB112FF503633FEFBFCBF" bold="true" box="[227,366,832,856]" pageId="7" pageNumber="62">Description.</emphasis>
Body length:
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. Width/length of head:1.13; width/length of pronotum:1.1; length/ width of elytra: 1.7. Head and pronotum black with weak bluish sheen, elytra black with bright blue luster. Mandibles dark brown, palpi brown, antenna and legs uniformly black. Lower surface black.
</paragraph>
<paragraph id="8B9136D0FFAAB112FF503152FD28F99F" blockId="7.[227,738,1057,1656]" pageId="7" pageNumber="62">
Head strongly trapezoid and depressed, slightly narrower than the pronotum. Eye moderately large and markedly protruded, 0.5 times as long as temples; temples protruding; posterior border almost perpendicular; constriction of neck weak and shallow, neck narrow. Mandibles elongate, almost straight in basal two thirds, incurved towards apex. Palpi elongate, narrowed towards apex, smooth.
<collectingRegion id="49EAF832FFAAB112FE883052FE01FADF" box="[315,404,1313,1336]" country="Myanmar" name="Chin" pageId="7" pageNumber="62">Mentum</collectingRegion>
with wide triangular tooth, bisetose. Antenna short, attaining the lateral pore of pronotum; three basal antennomeres and basal half of antennomere 4 glabrous. Clypeal suture shallow, laterally indistinct; clypeus slightly emarginate at apex. Labrum with deep wide triangular incision, asymmetric, right lobe longer. Frons with two irregularly shaped, shallow impressions. Supraorbital pore connected by a shallow groove with the postorbital pore. Surface with extremely fine transverse microreticulation, iridescent.
</paragraph>
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<paragraph id="8B9136D0FFAAB112FCB531FFFBE8FB23" blockId="7.[774,1285,1164,1220]" pageId="7" pageNumber="62">
Fig. 4. Habitus of
<taxonomicName id="4C2E4D53FFAAB112FC7931FFFB50FB44" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[970,1221,1164,1187]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="panumanod" status="sp. nov.">
<emphasis id="B95AEAC2FFAAB112FC7931FFFB50FB44" box="[970,1221,1164,1187]" italics="true" pageId="7" pageNumber="62">Hexagonia panumanod</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26957B9FFAAB112FB7831FFFA90FB44" box="[1227,1285,1164,1187]" pageId="7" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
Holotype, female. Scale bar 5 mm.
</paragraph>
</caption>
<paragraph id="8B9136D0FFAAB11DFF5033D2FD9CFD1F" blockId="7.[227,737,1697,1752]" lastBlockId="8.[302,813,193,760]" lastPageId="8" lastPageNumber="63" pageId="7" pageNumber="62">Pronotum wider than the head, obovate, widest at apical quarter; dorsal surface depressed.Anterior margin slightly convex, anterior angles completely rounded. Lateral margins evenly curved, immediately in front of base slightly excised; base straight, basal angles obtuse and slightly produced. Median line deep, impunctate, almost attaining anterior margin and base; both transverse sulci obsoleted. Basal grooves starts right from the posterior angles in form of short, smooth, diagonal impressions. Anterior margin and base not margined, lateral borders narrowly margined, marginal sulcus slightly wider near lateral pore. Anterior lateral seta situated at widest point, posterior lateral seta absent. Dorsal surface smooth, with some very shallow irregular transverse sulci, with extremely fine transverse microreticulation, slightly iridescent. Elytra elongate, markedly depressed, parallel sided. Base oblique, humeri rounded; lateral margins straight. Basal margin straight and forming a curve at the junction with lateral margin, between striae 4 and 5. Scutellar stria long and connected with stria 1. All striae deeply impressed and moderately punctate. Intervals flat and smooth. Scutellar seta situated at origin of stria 1. Interval 3 with three setiferous pores, the anterior pore situated in the stria 3 at the level of the end of scutellar stria; the discal pore situated in apical third of elytra near stria 2; posterior one situated close to apex, near stria 2. Interval 5 of elytra with one pore located opposite the middle between discal and apical pores. Marginal series consisting of 1213 pores. Intervals impunctate and with weak microreticulation, consisting of polygonal meshes, very glossy.</paragraph>
<paragraph id="8B9136D0FFA5B11DFE9D3652FDDCFC3F" blockId="8.[302,813,801,984]" pageId="8" pageNumber="63">Legs rather short, tarsomere 3 of all tarsi two times shorter than basal one; tarsomere 4 wide and very deeply excised, with dense brush­like pilosity on ventral surface. Tarsal claws edentate. Three basal tarsomeres of all legs triangular, very wide and dorsally smooth.</paragraph>
<paragraph id="8B9136D0FFA5B11DFE9D3172FE43FBFF" blockId="8.[302,470,1025,1048]" box="[302,470,1025,1048]" pageId="8" pageNumber="63">Male unknown.</paragraph>
<paragraph id="8B9136D0FFA5B11DFE9D3132FD36FAFF" blockId="8.[302,813,1089,1304]" pageId="8" pageNumber="63">Female genitalia. Gonocoxite 1 apically with five elongate setae along ventro­apical rim. Gonocoxite 2 rather narrow, triangular, slightly curved, with obtusely rounded apex; with three short and stout ensiform setae on the external­apical edge and one large suboval setiferous pit with seta near apex on the medio­ventral surface.</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA5B11DFE9D3033FD27FA7F" pageId="8" pageNumber="63" type="distribution">
<paragraph id="8B9136D0FFA5B11DFE9D3033FD27FA7F" blockId="8.[302,813,1344,1432]" pageId="8" pageNumber="63">
<emphasis id="B95AEAC2FFA5B11DFE9D3033FE2BFABF" bold="true" box="[302,446,1344,1368]" pageId="8" pageNumber="63">Distribution.</emphasis>
Known only from the
<typeStatus id="54958872FFA5B11DFD143032FD41FABF" box="[679,724,1345,1368]" pageId="8" pageNumber="63">type</typeStatus>
locality, situated in the province of
<collectingRegion id="49EAF832FFA5B11DFDFD3012FC9BFA9F" box="[590,782,1377,1400]" country="Philippines" name="Misamis Oriental" pageId="8" pageNumber="63">Misamis Oriental</collectingRegion>
. in the north of the island of Mindanao.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA5B11DFE9D30B3FBBEFE3F" pageId="8" pageNumber="63" type="etymology">
<paragraph id="8B9136D0FFA5B11DFE9D30B3FD43F9FF" blockId="8.[302,813,1472,1560]" pageId="8" pageNumber="63">
<emphasis id="B95AEAC2FFA5B11DFE9D30B3FE27FA3F" bold="true" box="[302,434,1472,1496]" pageId="8" pageNumber="63">Etymology.</emphasis>
This new species is named after
<emphasis id="B95AEAC2FFA5B11DFE843092FE2DFA1F" box="[311,440,1505,1528]" italics="true" pageId="8" pageNumber="63">Panumanod</emphasis>
, the guardian of the animals of the Higaunon Tribe in Northern Mindanao.
</paragraph>
<paragraph id="8B9136D0FFA5B11DFE9D3333FBBEFE3F" blockId="8.[302,813,1600,1752]" lastBlockId="8.[850,1361,193,472]" pageId="8" pageNumber="63">
<emphasis id="B95AEAC2FFA5B11DFE9D3333FE35F9BF" bold="true" box="[302,416,1600,1624]" pageId="8" pageNumber="63">Affinities.</emphasis>
This species is very peculiar within the genus in the metallic color and the strongly trapezoid head, with the supraorbital pore connected by a groove with the postorbital pore. In these two characters it is strongly reminiscent of some species of the Madagascan genus
<taxonomicName id="4C2E4D53FFA5B11DFB4235B2FBAAFF1F" authority="Fairmaire, 1896" authorityName="Fairmaire" authorityYear="1896" class="Insecta" family="Carabidae" genus="Omphreoides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B95AEAC2FFA5B11DFB4235B2FC1EFF1F" italics="true" pageId="8" pageNumber="63">Omphreoides</emphasis>
Fairmaire, 1896
</taxonomicName>
(
<bibRefCitation id="EFBF4B21FFA5B11DFBFE3592FB72FF1F" author="Jeannel R." box="[1101,1255,225,248]" pageId="8" pageNumber="63" pagination="373 - 766" refId="ref6681" refString="Jeannel R., 1948. Coleopteres carabiques de la region malgache (deuxieme partie). - Faune de l'Empire Francaise, 10: 373 - 766." type="journal article" year="1948">Jeannel, 1948</bibRefCitation>
). The following species (
<taxonomicName id="4C2E4D53FFA5B11DFC4C3472FB36FEFF" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[1023,1187,257,280]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="nigrocyanea" status="n. sp.">
<emphasis id="B95AEAC2FFA5B11DFC4C3472FB36FEFF" box="[1023,1187,257,280]" italics="true" pageId="8" pageNumber="63">H. nigrocyanea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26957B9FFA5B11DFB1A3472FB77FEFF" box="[1193,1250,257,280]" pageId="8" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
) is clearly similar in size, shape and general characters, but some of the features are less evident (the trapezoid head, the metallic color, ecc.). These species seem to form a small natural group within the genus, apparently without relationships with any other species group.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,318 @@
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<mods:title id="58DFFF8B6BEF30C66AFE2DC1E5BFE615">Taxonomic notes on the genus Hexagonia Kirby, 1825 with description of three new species from the Philippines (Coleoptera: Carabidae: Hexagoniini)</mods:title>
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<treatment id="038787C6FFAEB110FCB534D2FA96FDFF" LSID="urn:lsid:plazi:treatment:038787C6FFAEB110FCB534D2FA96FDFF" httpUri="http://treatment.plazi.org/id/038787C6FFAEB110FCB534D2FA96FDFF" lastPageId="5" lastPageNumber="60" pageId="3" pageNumber="58">
<subSubSection id="C334655BFFAEB116FCB534D2FB3CFE5F" box="[774,1193,416,441]" pageId="3" pageNumber="58" type="nomenclature">
<paragraph id="8B9136D0FFAEB116FCB534D2FB3CFE5F" blockId="3.[774,1281,416,504]" box="[774,1193,416,441]" pageId="3" pageNumber="58">
<heading id="D0D981BCFFAEB116FCB534D2FB3CFE5F" bold="true" box="[774,1193,416,441]" fontSize="10" level="2" pageId="3" pageNumber="58" reason="4">
<taxonomicName id="4C2E4D53FFAEB116FCB534D2FB3CFE5F" ID-CoL="6LTFX" authority="Dupuis, 1912: 308" authorityName="Dupuis" authorityPageNumber="308" authorityYear="1912" box="[774,1193,416,441]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B95AEAC2FFAEB116FCB534D2FB3CFE5F" bold="true" box="[774,1193,416,441]" pageId="3" pageNumber="58">
<emphasis id="B95AEAC2FFAEB116FCB534D2FC41FE5E" bold="true" box="[774,980,417,441]" italics="true" pageId="3" pageNumber="58">Hexagonia sauteri</emphasis>
Dupuis, 1912: 308
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAEB116FCB534B2FC26FE3F" box="[774,947,449,472]" pageId="3" pageNumber="58" type="description">
<paragraph id="8B9136D0FFAEB116FCB534B2FC26FE3F" blockId="3.[774,1281,416,504]" box="[774,947,449,472]" pageId="3" pageNumber="58">
(
<figureCitation id="13152A55FFAEB116FCBD34B2FCE3FE3F" box="[782,886,449,472]" captionStart="Figs" captionStartId="4.[302,352,1206,1229]" captionTargetBox="[302,1360,189,1172]" captionTargetId="figure-25@4.[302,1360,189,1172]" captionTargetPageId="4" captionText="Figs. 23. Habitus of Hexagonia. 2 H. sauteri Dupuis, 1912 Holotype, male; 3 H. castanea JedliËka, 1936 syn. n., Cotype, male. Scale bar 5 mm." pageId="3" pageNumber="58">Figs. 23</figureCitation>
,
<figureCitation id="13152A55FFAEB116FC3134B2FC3EFE3F" box="[898,939,449,472]" captionStart="Figs" captionStartId="10.[302,352,1399,1422]" captionTargetBox="[307,1358,191,1362]" captionTargetId="figure-177@10.[302,1360,189,1366]" captionTargetPageId="10" captionText="Figs. 711. Aedeagus of Hexagonia, A dorsal view, B lateral view. 7 H. andrewesi JedliËka, 1935, Holotype; 8 H. castanea JedliËka, 1936 syn. n. Cotype; 9 H. sauteri Dupuis, 1912, Holotype. Scale bar 1 mm. 10 H. amicitia sp. n., Holotype; 11 H. nigrocyanea sp. n. Holotype. Scale bar 1 mm." pageId="3" pageNumber="58">89</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAEB116FCB53492FA94FE1F" box="[774,1281,479,504]" pageId="3" pageNumber="58" type="reference_group">
<paragraph id="8B9136D0FFAEB116FCB53492FA94FE1F" blockId="3.[774,1281,416,504]" box="[774,1281,479,504]" pageId="3" pageNumber="58">
<treatmentCitationGroup id="AB3E11FEFFAEB116FCB53492FA94FE1F" box="[774,1281,479,504]" pageId="3" pageNumber="58">
<treatmentCitation id="0A8F10C1FFAEB116FCB53492FB24FE1F" author="JedliEka A." box="[774,1201,479,504]" page="55" pageId="3" pageNumber="58" year="1936">
<taxonomicName id="4C2E4D53FFAEB116FCB53492FB24FE1F" ID-CoL="6LTF9" authority="JedliEka, 1936: 55" authorityName="JedliEka" authorityPageNumber="55" authorityYear="1936" box="[774,1201,479,504]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="castanea" status="syn. nov.">
<emphasis id="B95AEAC2FFAEB116FCB53492FC71FE1F" box="[774,996,481,504]" italics="true" pageId="3" pageNumber="58">Hexagonia castanea</emphasis>
<bibRefCitation id="EFBF4B21FFAEB116FC583492FB24FE1F" author="JedliEka A." box="[1003,1201,479,504]" pageId="3" pageNumber="58" pagination="51 - 55" refId="ref6737" refString="JedliEka A., 1936. Neue Carabiden aus Ostasien. (XI Teil). Entomologisches Nachrichtenblatt 10 (1): 51 - 55." type="journal article" year="1936">JedliËka, 1936: 55</bibRefCitation>
</taxonomicName>
</treatmentCitation>
<emphasis id="B95AEAC2FFAEB116FB043493FA94FE1F" bold="true" box="[1207,1281,480,504]" pageId="3" pageNumber="58">
<taxonomicNameLabel id="A26957B9FFAEB116FB043493FA94FE1F" box="[1207,1281,480,504]" pageId="3" pageNumber="58">syn. n.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAEB116FCB53753FA90FA3F" pageId="3" pageNumber="58" type="materials_examined">
<paragraph id="8B9136D0FFAEB116FCB53753FB9AFADF" blockId="3.[774,1285,544,1336]" pageId="3" pageNumber="58">
<emphasis id="B95AEAC2FFAEB116FCB53753FC28FDDF" bold="true" box="[774,957,544,568]" pageId="3" pageNumber="58">Type material:</emphasis>
<materialsCitation id="3B463C8DFFAEB116FC7A3752FB13FD3F" collectionCode="BMNH" collectorName="Ex. Deutsch &amp; H. E. Andrewes Coll." country="Taiwan" determinerName="Dupuis" location="Formosa" pageId="3" pageNumber="58" specimenCount="1" typeStatus="holotype">
<typeStatus id="54958872FFAEB116FC7A3752FBA7FDDF" box="[969,1074,545,568]" pageId="3" pageNumber="58" type="holotype">Holotype</typeStatus>
, male “Kankau [= Hengchun] |
<collectingCountry id="F3397640FFAEB116FC0B3732FB83FDBF" box="[952,1046,577,600]" name="Taiwan" pageId="3" pageNumber="58">Formosa</collectingCountry>
| Sauter IV.09”, “
<collectorName id="26DB5306FFAEB116FB6C3732FCC9FD9F" pageId="3" pageNumber="58">Ex. Deutsch</collectorName>
| Ent. Mus”, “
<collectorName id="26DB5306FFAEB116FC593712FB54FD9F" box="[1002,1217,609,632]" pageId="3" pageNumber="58">H.E. Andrewes Coll.</collectorName>
| B.M. 1945­ 97”, “
<taxonomicName id="4C2E4D53FFAEB116FC3037F2FC6FFD7F" authorityName="Kirby" authorityYear="1825" box="[899,1018,641,664]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">Hexagonia</taxonomicName>
| sauteri | Dupuis [handwritten]”, “
<determinerName id="7E4DDA14FFAEB116FC3B37D2FC42FD5F" box="[904,983,673,696]" pageId="3" pageNumber="58">Dupuis</determinerName>
det”, “Typus [red label]”, “NHMUK 014404802” (
<collectionCode id="ED3FAE15FFAEB116FB9A37B2FBE9FD3F" box="[1065,1148,705,728]" pageId="3" pageNumber="58">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B463C8DFFAEB116FB2A37B2FBE2FC9F" collectionCode="BMNH" collectorName="Ex. Coll. Berl. &amp; Coll. Berl. &amp; H. E. Andrewes Coll." country="Taiwan" determinerName="P. Dupuis" location="Formosa" pageId="3" pageNumber="58" specimenCount="1" typeStatus="paratype">
<typeStatus id="54958872FFAEB116FB2A37B2FB68FD3F" box="[1177,1277,705,728]" pageId="3" pageNumber="58" type="paratype">Paratype</typeStatus>
, 1 ex
<collectingCountry id="F3397640FFAEB116FCEA3792FC2FFD1F" box="[857,954,737,760]" name="Taiwan" pageId="3" pageNumber="58">Formosa</collectingCountry>
| Kankau IV.09 | Sauter”, “
<collectorName id="26DB5306FFAEB116FB6C3792FCEDFCFF" pageId="3" pageNumber="58">Ex. Coll. Berl.</collectorName>
| Zool. Mus.”, “Cotype [green round label]”, “
<taxonomicName id="4C2E4D53FFAEB116FCC03652FBD1FCDF" authorityName="Dupuis" authorityYear="1912" box="[883,1092,801,824]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="sauteri">Hexagonia sauteri</taxonomicName>
| det.
<determinerName id="7E4DDA14FFAEB116FB263652FA90FCDF" box="[1173,1285,801,824]" pageId="3" pageNumber="58">P. Dupuis</determinerName>
1913”, “
<collectorName id="26DB5306FFAEB116FCD13632FBD0FCBF" box="[866,1093,833,856]" pageId="3" pageNumber="58">H.E. Andrewes Coll.</collectorName>
| B.M. 1945­ 97”, “NHMUK 014404769” (
<collectionCode id="ED3FAE15FFAEB116FBAE3612FBFBFC9F" box="[1053,1134,865,888]" pageId="3" pageNumber="58">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B463C8DFFAEB116FB3B3612FBEAFBFF" collectingDate="1914-02-06" collectionCode="BMNH" collectorName="Coll. Bottcher" country="Philippines" determinerName="Ing. JedliEka" location="Type" pageId="3" pageNumber="58" specimenCount="1" stateProvince="Manila" typeStatus="holotype">
<typeStatus id="54958872FFAEB116FB3B3612FB78FC9F" box="[1160,1261,865,888]" pageId="3" pageNumber="58" type="holotype">Holotype</typeStatus>
<location id="8EF1600BFFAEB116FCA336F2FCDDFC7F" LSID="urn:lsid:plazi:treatment:038787C6FFAEB110FCB534D2FA96FDFF:8EF1600BFFAEB116FCA336F2FCDDFC7F" box="[784,840,897,920]" country="Philippines" name="Type" pageId="3" pageNumber="58" stateProvince="Manila">Type</location>
[red round label]”, “
<collectingCountry id="F3397640FFAEB116FB9C36F2FB63FC7F" box="[1071,1270,897,920]" name="Philippines" pageId="3" pageNumber="58">Philippine Islands</collectingCountry>
|
<collectingRegion id="49EAF832FFAEB116FCB536D2FCC7FC5F" box="[774,850,929,952]" country="Philippines" name="Manila" pageId="3" pageNumber="58">Manila</collectingRegion>
|
<date id="FF901010FFAEB116FCD736D2FC5DFC5F" box="[868,968,929,952]" pageId="3" pageNumber="58" value="1914-02-06">
<collectingDate id="EFD4E9F8FFAEB116FCD736D2FC5DFC5F" box="[868,968,929,952]" pageId="3" pageNumber="58" value="1914-02-06">6.II.1914</collectingDate>
</date>
|
<collectorName id="26DB5306FFAEB116FC6A36D2FBE3FC5F" box="[985,1142,929,952]" pageId="3" pageNumber="58">Coll. Bottcher</collectorName>
”, “Philippine Is.|
<collectorName id="26DB5306FFAEB116FC9A36B2FC2EFC3F" box="[809,955,961,984]" pageId="3" pageNumber="58">Coll. Bottcher</collectorName>
| B.M. 1929­ 201”, “
<taxonomicName id="4C2E4D53FFAEB116FB3536B2FB6CFC3F" authorityName="Kirby" authorityYear="1825" box="[1158,1273,961,984]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">Hexagonia</taxonomicName>
| castanea
<taxonomicNameLabel id="A26957B9FFAEB116FCD83692FC0AFC1F" box="[875,927,993,1016]" pageId="3" pageNumber="58" rank="species">sp.n.</taxonomicNameLabel>
| det.
<determinerName id="7E4DDA14FFAEB116FC5A3692FBE0FC1F" box="[1001,1141,991,1016]" pageId="3" pageNumber="58">Ing. JedliËka</determinerName>
[red label]”, “NHMUK 014404930” (
<collectionCode id="ED3FAE15FFAEB116FB903172FBE0FBFF" box="[1059,1141,1025,1048]" pageId="3" pageNumber="58">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B463C8DFFAEB116FB223172FB9EFADF" collectingDate="1914-02-06" collectionCode="BMNH, SDEI" collectorName="Coll. Bottcher" country="Philippines" determinerName="Ing. JedliEka &amp; Dupuis" location="Cotype" pageId="3" pageNumber="58" specimenCount="2" stateProvince="Manila" typeStatus="paratype">
<typeStatus id="54958872FFAEB116FB223172FB6BFBFF" box="[1169,1278,1025,1048]" pageId="3" pageNumber="58" type="paratype">Paratypes</typeStatus>
, 5 ex
<location id="8EF1600BFFAEB116FCE03152FC34FBDF" LSID="urn:lsid:plazi:treatment:038787C6FFAEB110FCB534D2FA96FDFF:8EF1600BFFAEB116FCE03152FC34FBDF" box="[851,929,1057,1080]" country="Philippines" name="Cotype" pageId="3" pageNumber="58" stateProvince="Manila">Cotype</location>
[yellow round label]”, “
<collectingCountry id="F3397640FFAEB116FB243152FCC6FBBF" name="Philippines" pageId="3" pageNumber="58">Philippine Islands</collectingCountry>
|
<collectingRegion id="49EAF832FFAEB116FCDD3132FC2EFBBF" box="[878,955,1089,1112]" country="Philippines" name="Manila" pageId="3" pageNumber="58">Manila</collectingRegion>
|
<date id="FF901010FFAEB116FC663132FBACFBBF" box="[981,1081,1089,1112]" pageId="3" pageNumber="58" value="1914-02-06">
<collectingDate id="EFD4E9F8FFAEB116FC663132FBACFBBF" box="[981,1081,1089,1112]" pageId="3" pageNumber="58" value="1914-02-06">6.II.1914</collectingDate>
</date>
|
<collectorName id="26DB5306FFAEB116FBE73132FB67FBBF" box="[1108,1266,1089,1112]" pageId="3" pageNumber="58">Coll. Bottcher</collectorName>
”, “Philippine Is.|
<collectorName id="26DB5306FFAEB116FC193112FBD5FB9F" box="[938,1088,1121,1144]" pageId="3" pageNumber="58">Coll. Bottcher</collectorName>
| B.M. 1929­ 201”, “
<taxonomicName id="4C2E4D53FFAEB116FCA331F2FC12FB7F" authorityName="Kirby" authorityYear="1825" box="[784,903,1153,1176]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">Hexagonia</taxonomicName>
| castanea
<taxonomicNameLabel id="A26957B9FFAEB116FC4831F2FBBAFB7F" box="[1019,1071,1153,1176]" pageId="3" pageNumber="58" rank="species">sp.n.</taxonomicNameLabel>
| det.
<determinerName id="7E4DDA14FFAEB116FBDC31F2FB6EFB7F" box="[1135,1275,1151,1176]" pageId="3" pageNumber="58">Ing. JedliËka</determinerName>
” (
<collectionCode id="ED3FAE15FFAEB116FCBD31D2FCF4FB5F" box="[782,865,1185,1208]" pageId="3" pageNumber="58">BMNH</collectionCode>
). 2 ex “Kankau |
<collectingCountry id="F3397640FFAEB116FBA031D2FBF9FB5F" box="[1043,1132,1185,1208]" name="Taiwan" pageId="3" pageNumber="58">Formosa</collectingCountry>
| Sauter IV.09”, “
<determinerName id="7E4DDA14FFAEB116FCA331B2FCF5FB3F" box="[784,864,1217,1240]" pageId="3" pageNumber="58">Dupuis</determinerName>
det”, “Typus [red label]”, “
<taxonomicName id="4C2E4D53FFAEB116FB3D31B2FCDBFB1F" authorityName="Dupuis" authorityYear="1912" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="sauteri">Hexagonia sauteri</taxonomicName>
| Dupuis”, “
<typeStatus id="54958872FFAEB116FC673192FBAEFB1F" box="[980,1083,1249,1272]" pageId="3" pageNumber="58" type="syntype">Syntypus</typeStatus>
[red label]”, “DEI
<taxonomicName id="4C2E4D53FFAEB116FCB53072FCE9FAFF" authorityName="Linnaeus" authorityYear="1758" box="[774,892,1281,1304]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="3" pageNumber="58" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
| # 200406”;
<emphasis id="B95AEAC2FFAEB116FBBF3072FBFCFAFF" box="[1036,1129,1281,1304]" italics="true" pageId="3" pageNumber="58">idem but</emphasis>
“DEI
<taxonomicName id="4C2E4D53FFAEB116FB1D3072FCBBFADF" authorityName="Linnaeus" authorityYear="1758" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="3" pageNumber="58" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
| # 200407” (
<collectionCode id="ED3FAE15FFAEB116FC713052FB91FADF" box="[962,1028,1313,1336]" pageId="3" pageNumber="58">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9136D0FFAEB116FCB53013FA90FA3F" blockId="3.[774,1285,1376,1496]" pageId="3" pageNumber="58">
<emphasis id="B95AEAC2FFAEB116FCB53013FC20FA9F" bold="true" box="[774,949,1376,1400]" pageId="3" pageNumber="58">Other material:</emphasis>
<materialsCitation id="3B463C8DFFAEB116FC0A3012FA90FA3F" collectingDate="2016-03" collectionCode="BMNH" collectorName="Coll. Bottcher, B. M." country="Philippines" location="Dingalan" municipality="Sierra Madre" pageId="3" pageNumber="58" specimenCount="1" stateProvince="Aurora">
1 ex “Philippine Is.,
<collectorName id="26DB5306FFAEB116FB2B3012FCEEFA7F" pageId="3" pageNumber="58">Coll. Bottcher, B.M.</collectorName>
1929­ 201”, “NHMUK 014404905” (
<collectionCode id="ED3FAE15FFAEB116FCBD30D2FCF0FA5F" box="[782,869,1441,1464]" pageId="3" pageNumber="58">BMNH</collectionCode>
); 1 ex female
<collectingCountry id="F3397640FFAEB116FBA530D2FB00FA5F" box="[1046,1173,1441,1464]" name="Philippines" pageId="3" pageNumber="58">Philippines</collectingCountry>
, E Luzon |
<collectingMunicipality id="6BF5ACAAFFAEB116FCB530B2FC00FA3F" box="[774,917,1473,1496]" pageId="3" pageNumber="58">Sierra Madre</collectingMunicipality>
,
<collectingRegion id="49EAF832FFAEB116FC2F30B2FC7DFA3F" box="[924,1000,1473,1496]" country="Philippines" name="Aurora" pageId="3" pageNumber="58">Aurora</collectingRegion>
|
<location id="8EF1600BFFAEB116FC4430B2FBCCFA3F" LSID="urn:lsid:plazi:treatment:038787C6FFAEB110FCB534D2FA96FDFF:8EF1600BFFAEB116FC4430B2FBCCFA3F" box="[1015,1113,1473,1496]" country="Philippines" municipality="Sierra Madre" name="Dingalan" pageId="3" pageNumber="58" stateProvince="Aurora">Dingalan</location>
<date id="FF901010FFAEB116FBED30B2FB2DFA3F" box="[1118,1208,1473,1496]" pageId="3" pageNumber="58" value="2016-03">
<collectingDate id="EFD4E9F8FFAEB116FBED30B2FB2DFA3F" box="[1118,1208,1473,1496]" pageId="3" pageNumber="58" value="2016-03">III.2016</collectingDate>
</date>
” (cRS)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAEB116FCB53373FA90F99F" pageId="3" pageNumber="58" type="diagnosis">
<paragraph id="8B9136D0FFAEB116FCB53373FA90F99F" blockId="3.[774,1285,1536,1656]" pageId="3" pageNumber="58">
<emphasis id="B95AEAC2FFAEB116FCB53373FC15F9FF" bold="true" box="[774,896,1536,1560]" pageId="3" pageNumber="58">Diagnosis.</emphasis>
Instantly recognizable among all other oriental species of the genus by the combination of bifid mentum, short and wide mandibles, and several pores in elytral intervals 3 and 5.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAEB110FCB533D3FC2FFF1F" lastPageId="5" lastPageNumber="60" pageId="3" pageNumber="58" type="description">
<paragraph id="8B9136D0FFAEB116FCB533D3FA93F93F" blockId="3.[774,1286,1696,1752]" pageId="3" pageNumber="58">
<emphasis id="B95AEAC2FFAEB116FCB533D3FC3FF95F" bold="true" box="[774,938,1696,1720]" pageId="3" pageNumber="58">Redescription.</emphasis>
Body length:
<quantity id="4CD69B35FFAEB116FB8933D2FB23F95F" box="[1082,1206,1697,1720]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" metricValueMax="9.5" metricValueMin="8.3" pageId="3" pageNumber="58" unit="mm" value="8.9" valueMax="9.5" valueMin="8.3">8.3­ 9.5 mm</quantity>
. Width/ length of head: 1.27; width/length of pronotum:
</paragraph>
<paragraph id="8B9136D0FFA9B111FEF035E2FAA9FF43" blockId="4.[323,1340,144,164]" box="[323,1340,144,164]" pageId="4" pageNumber="59">
<emphasis id="B95AEAC2FFA9B111FEF035E2FDDBFF43" box="[323,590,145,164]" italics="true" pageId="4" pageNumber="59">Taxonomic notes on the genus</emphasis>
<taxonomicName id="4C2E4D53FFA9B111FDE635E3FCB4FF43" authority="Kirby, 1825" authorityName="Kirby" authorityYear="1825" box="[597,801,144,164]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="genus">
Hexagonia
<bibRefCitation id="EFBF4B21FFA9B111FD0935E2FCB4FF43" author="Kirby, W. A." box="[698,801,145,164]" pageId="4" pageNumber="59" pagination="563 - 572" refId="ref6764" refString="Kirby, W. A. 1825. description of such genera and species of insects, alluded to in the &quot; Introduction to Entomology &quot; of Messrs. KIRBY and SPENCE, as appear not to have been before sufficiently noticed or described. The Transactions of the Linnean Society of London, 14: 563 - 572." type="journal article" year="1825">
<emphasis id="B95AEAC2FFA9B111FD0935E2FCB4FF43" box="[698,801,145,164]" italics="true" pageId="4" pageNumber="59">Kirby, 1825</emphasis>
</bibRefCitation>
</taxonomicName>
<emphasis id="B95AEAC2FFA9B111FC9B35E2FAA9FF43" box="[808,1340,145,164]" italics="true" pageId="4" pageNumber="59">
with description of three new species from the
<collectingCountry id="F3397640FFA9B111FB7B35E2FAB8FF43" box="[1224,1325,145,164]" name="Philippines" pageId="4" pageNumber="59">Philippines</collectingCountry>
...
</emphasis>
</paragraph>
<caption id="DF516658FFA9B111FE9D31C5FCE5FB0A" pageId="4" pageNumber="59" startId="4.[302,352,1206,1229]" targetBox="[302,1360,189,1172]" targetPageId="4" targetType="figure">
<paragraph id="8B9136D0FFA9B111FE9D31C5FCE5FB0A" blockId="4.[302,1360,1206,1262]" pageId="4" pageNumber="59">
Figs. 23. Habitus of
<taxonomicName id="4C2E4D53FFA9B111FD9231C4FD02FB29" authorityName="Kirby" authorityYear="1825" box="[545,663,1207,1230]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B95AEAC2FFA9B111FD9231C4FD02FB29" box="[545,663,1207,1230]" italics="true" pageId="4" pageNumber="59">Hexagonia</emphasis>
</taxonomicName>
. 2
<taxonomicName id="4C2E4D53FFA9B111FD6731C4FC4AFB2A" authority="Dupuis, 1912" authorityName="Dupuis" authorityYear="1912" box="[724,991,1206,1230]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B95AEAC2FFA9B111FD6731C4FCD7FB29" box="[724,834,1207,1230]" italics="true" pageId="4" pageNumber="59">H. sauteri</emphasis>
Dupuis, 1912
</taxonomicName>
Holotype, male; 3
<taxonomicName id="4C2E4D53FFA9B111FB7E31C4FE5FFB0A" authority="JedliEka, 1936" authorityName="JedliEka" authorityYear="1936" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="castanea" status="syn. n.">
<emphasis id="B95AEAC2FFA9B111FB7E31C4FAC5FB29" box="[1229,1360,1207,1230]" italics="true" pageId="4" pageNumber="59">H. castanea</emphasis>
<bibRefCitation id="EFBF4B21FFA9B111FE9D31A5FE5FFB0A" author="JedliEka A." box="[302,458,1237,1262]" pageId="4" pageNumber="59" pagination="51 - 55" refId="ref6737" refString="JedliEka A., 1936. Neue Carabiden aus Ostasien. (XI Teil). Entomologisches Nachrichtenblatt 10 (1): 51 - 55." type="journal article" year="1936">JedliËka, 1936</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A26957B9FFA9B111FE6331A5FD8CFB0A" box="[464,537,1238,1261]" pageId="4" pageNumber="59">syn. n.</taxonomicNameLabel>
, Cotype, male. Scale bar 5 mm.
</paragraph>
</caption>
<paragraph id="8B9136D0FFA9B111FE9D3053FD20FA90" blockId="4.[302,813,1312,1399]" pageId="4" pageNumber="59">1.17­ 1.2; length/ width of elytra: 1.45­ 1.56. Body uniformly brown, sometimes tibiae, sides of elytra and mandibles slightly darker.</paragraph>
<paragraph id="8B9136D0FFA9B111FE9D30D3FAC5F9B0" blockId="4.[302,813,1440,1751]" lastBlockId="4.[850,1361,1312,1623]" pageId="4" pageNumber="59">
Head trapezoid, dorsally depressed, narrower than the prothorax. Eyes large and markedly protruded, almost as long as temples; temples protruding; gradually rounded to the constriction of the neck. Neck constriction well marked, narrow and deep. Mandibles relatively short, wide and gradually incurved towards apex. Palpi elongate, narrowed towards apex, smooth.
<collectingRegion id="49EAF832FFA9B111FD9933F3FD10F970" box="[554,645,1664,1687]" country="Myanmar" name="Chin" pageId="4" pageNumber="59">Mentum</collectingRegion>
bifid, bisetose. Antenna relatively short, just reaching lateral pore of pronotum; three basal antennomeres and basal half of antennomere 4 glabrous. Clypeus with two pores and an irregular transverse depression, apical margin of clypeus slightly concave; clypeal suture very strong and deep, laterally effaced. Labrum transverse and slightly asymmetric, margin with wide rounded incision, right angle slightly longer than left. Frons with two deep and wide impressions, connected by a third weaker impression between them. Postorbital pore isolated. Surface with indistinct microreticulation, shiny.
</paragraph>
<paragraph id="8B9136D0FFA9B111FCE133F3FAC4F930" blockId="4.[850,1361,1664,1751]" pageId="4" pageNumber="59">Pronotum much wider than head, markedly transverse, widest near apical third; dorsal surface depressed.Anterior margin slightly concave, ante­</paragraph>
<paragraph id="8B9136D0FFA8B110FD8F35E2FC38FF43" blockId="5.[572,941,145,164]" box="[572,941,145,164]" pageId="5" pageNumber="60">
<emphasis id="B95AEAC2FFA8B110FD8F35E2FC38FF43" box="[572,941,145,164]" italics="true" pageId="5" pageNumber="60">
Anichtchenko A., Sciaky R.,
<taxonomicName id="4C2E4D53FFA8B110FC8435E2FCECFF43" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[823,889,145,164]" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="genus">Medina</taxonomicName>
M.N.
</emphasis>
</paragraph>
<paragraph id="8B9136D0FFA8B110FF5035B2FED4FD7F" blockId="5.[227,737,193,664]" pageId="5" pageNumber="60">rior angles completely rounded. Lateral margins form a slight and obtuse angle opposite the lateral setiferous pore; with evident excision before hind angles; basal angles right and moderately produced; base of pronotum straight. Median line narrow and deep, impunctate, almost reaching anterior and posterior margins of pronotum; anterior and posterior transverse sulcus obsolete. Anterior margin and base not margined; lateral borders margined, marginal sulcus gradually widened from angles of pronotum to setiferous pore. Lateral setiferous pore situated in widest point of pronotum, posterior pore absent. Disc transversally rugose, with hardly visible transverse microreticulation.</paragraph>
<paragraph id="8B9136D0FFA8B110FF5037B2FE67FB3F" blockId="5.[227,738,705,1240]" pageId="5" pageNumber="60">Elytra elongate, markedly depressed, very slightly widened in apical half. Humeri rounded. Lateral margins straight. Basal margin oblique, connected at a very obtuse angle with lateral margin at position between striae 4 and 5. Scutellar stria long, not connected with stria 1. All striae deeply impressed and crenulate. Intervals flat and smooth. Scutellar seta situated at base of 1st stria. Interval 3 with 47 setiferous pores (usually 67), two anterior pores closely spaced and situated near stria 3 at the level of end of scutellar stria; the discal and apical pores irregularly placed in apical half. Interval 5 of elytra with 2 pores, asymmetrically and irregularly placed in apical half. Marginal series consisting of 1214 pores. Intervals impunctate and with well visible squamiform microreticulation, glossy.</paragraph>
<paragraph id="8B9136D0FFA8B110FF503072FDBFFA7F" blockId="5.[227,737,1281,1432]" pageId="5" pageNumber="60">Legs of average size. Three basal tarsomeres of all legs triangular, very wide and dorsally smooth; tarsomere 4 of all tarsi wide and very deeply excised, with dense brush­like pilosity on ventral surface. Tarsal claws edentate.</paragraph>
<paragraph id="8B9136D0FFA8B110FF5030B2FD3AF9FF" blockId="5.[227,738,1473,1560]" pageId="5" pageNumber="60">
Median lobe of aedeagus (
<figureCitation id="13152A55FFA8B110FE4E30B2FDF6FA3F" box="[509,611,1473,1496]" captionStart="Figs" captionStartId="10.[302,352,1399,1422]" captionTargetBox="[307,1358,191,1362]" captionTargetId="figure-177@10.[302,1360,189,1366]" captionTargetPageId="10" captionText="Figs. 711. Aedeagus of Hexagonia, A dorsal view, B lateral view. 7 H. andrewesi JedliËka, 1935, Holotype; 8 H. castanea JedliËka, 1936 syn. n. Cotype; 9 H. sauteri Dupuis, 1912, Holotype. Scale bar 1 mm. 10 H. amicitia sp. n., Holotype; 11 H. nigrocyanea sp. n. Holotype. Scale bar 1 mm." pageId="5" pageNumber="60">Figs. 89</figureCitation>
) elongated, apical third of median lobe slightly downturned. Endophallus with big semicircular sclerite.
</paragraph>
<paragraph id="8B9136D0FFA8B110FF503332FD9FF9BF" blockId="5.[227,522,1601,1624]" box="[227,522,1601,1624]" pageId="5" pageNumber="60">Female genitalia unstudied.</paragraph>
<paragraph id="8B9136D0FFA8B110FF5033F3FC2FFF1F" blockId="5.[227,737,1664,1752]" lastBlockId="5.[774,1285,193,248]" pageId="5" pageNumber="60">
<emphasis id="B95AEAC2FFA8B110FF5033F3FE38F97F" bold="true" box="[227,429,1664,1688]" pageId="5" pageNumber="60">Taxonomic notes.</emphasis>
After careful study of
<typeStatus id="54958872FFA8B110FD0133F2FD75F97F" box="[690,736,1665,1688]" pageId="5" pageNumber="60">type</typeStatus>
material of
<taxonomicName id="4C2E4D53FFA8B110FEED33D2FDF4F95F" authority="Dupuis, 1912" authorityName="Dupuis" authorityYear="1912" box="[350,609,1697,1720]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B95AEAC2FFA8B110FEED33D2FE5CF95F" box="[350,457,1697,1720]" italics="true" pageId="5" pageNumber="60">H. sauteri</emphasis>
Dupuis, 1912
</taxonomicName>
and
<taxonomicName id="4C2E4D53FFA8B110FD2733D2FE54F93F" authority="JedliEka, 1936" authorityName="JedliEka" authorityYear="1936" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B95AEAC2FFA8B110FD2733D2FE8AF93F" italics="true" pageId="5" pageNumber="60">H. castanea</emphasis>
<bibRefCitation id="EFBF4B21FFA8B110FE9633B2FE54F93F" author="JedliEka A." box="[293,449,1727,1752]" pageId="5" pageNumber="60" pagination="51 - 55" refId="ref6737" refString="JedliEka A., 1936. Neue Carabiden aus Ostasien. (XI Teil). Entomologisches Nachrichtenblatt 10 (1): 51 - 55." type="journal article" year="1936">JedliËka, 1936</bibRefCitation>
</taxonomicName>
no differences were found between the two species. The new synonymy is established here.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFA8B110FCB53453FA96FDFF" pageId="5" pageNumber="60" type="distribution">
<paragraph id="8B9136D0FFA8B110FCB53453FBEBFEBF" blockId="5.[774,1285,288,344]" pageId="5" pageNumber="60">
<emphasis id="B95AEAC2FFA8B110FCB53453FC09FEDF" bold="true" box="[774,924,288,312]" pageId="5" pageNumber="60">Distribution.</emphasis>
<collectingCountry id="F3397640FFA8B110FC153452FC7CFEDF" box="[934,1001,289,312]" name="China" pageId="5" pageNumber="60">China</collectingCountry>
,
<collectingCountry id="F3397640FFA8B110FC4B3452FBA3FEDF" box="[1016,1078,289,312]" name="Japan" pageId="5" pageNumber="60">Japan</collectingCountry>
(Ryukyu Islands),
<collectingCountry id="F3397640FFA8B110FCB53432FCCDFEBF" box="[774,856,321,344]" name="Taiwan" pageId="5" pageNumber="60">Taiwan</collectingCountry>
,
<collectingCountry id="F3397640FFA8B110FCD13432FBBDFEBF" box="[866,1064,321,344]" name="Philippines" pageId="5" pageNumber="60">Philippine Islands</collectingCountry>
: Luzon.
</paragraph>
<paragraph id="8B9136D0FFA8B110FCB534F3FA96FDFF" blockId="5.[774,1285,384,536]" pageId="5" pageNumber="60">
<emphasis id="B95AEAC2FFA8B110FCB534F3FCEFFE7F" bold="true" box="[774,890,384,408]" pageId="5" pageNumber="60">Affinities.</emphasis>
This species seems rather isolated within the genus in the rather short and wide elytra, crenulate striae and several points on intervals 3 and 5. Beyond this, it is the only species known to day from an island different from Mindanao.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,196 @@
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<mods:title id="5774CF9994E48358E5FAA473FD387585">Taxonomic notes on the genus Hexagonia Kirby, 1825 with description of three new species from the Philippines (Coleoptera: Carabidae: Hexagoniini)</mods:title>
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</mods:role>
<mods:namePart id="0E43C26E366EE64E0AD6F5B9581AAFC5">Medina, Milton Norman</mods:namePart>
</mods:name>
<mods:typeOfResource id="195E137499B1D31832BC0B752E738962">text</mods:typeOfResource>
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<mods:titleInfo id="961192F82556F2EEC9A16266122B6F4B">
<mods:title id="F413C4ABDD17BA53A318F5664985B40B">Baltic Journal of Coleopterology</mods:title>
</mods:titleInfo>
<mods:part id="C589D965E57101BD2D2398E48555140C">
<mods:date id="BE5FC753F8FADD95CEFA95E70C084069">2022</mods:date>
<mods:detail id="3829E59E137E3DFA7E4D91FAD3A5F281" type="volume">
<mods:number id="E5405A42CA5147A0DA81334F9E2D934A">22</mods:number>
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<mods:number id="18074D308672627A40E17C2E0C261707">1</mods:number>
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<mods:start id="1E50474D3434B2514F406065A05AE7AD">55</mods:start>
<mods:end id="B274C3DDDADA69CCCB66C1AD7C5A2CD1">66</mods:end>
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<mods:identifier id="068797DCBED0C0EB6B97F4118F4F667E" type="ISSN">1407-8619</mods:identifier>
</mods:mods>
<treatment id="038787C6FFAFB116FE9D35B2FA95FEBF" LSID="urn:lsid:plazi:treatment:038787C6FFAFB116FE9D35B2FA95FEBF" httpUri="http://treatment.plazi.org/id/038787C6FFAFB116FE9D35B2FA95FEBF" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57">
<subSubSection id="C334655BFFAFB117FE9D35B2FD63FF3F" box="[302,758,191,217]" pageId="2" pageNumber="57" type="nomenclature">
<paragraph id="8B9136D0FFAFB117FE9D35B2FD63FF3F" blockId="2.[302,758,191,248]" box="[302,758,191,217]" pageId="2" pageNumber="57">
<heading id="D0D981BCFFAFB117FE9D35B2FD63FF3F" bold="true" box="[302,758,191,217]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="57" reason="4">
<treatmentCitationGroup id="AB3E11FEFFAFB117FE9D35B2FD63FF3F" box="[302,758,191,217]" pageId="2" pageNumber="57">
<treatmentCitation id="0A8F10C1FFAFB117FE9D35B2FD63FF3F" author="JedliEka A." box="[302,758,191,217]" page="89" pageId="2" pageNumber="57" year="1935">
<taxonomicName id="4C2E4D53FFAFB117FE9D35B2FD63FF3F" ID-CoL="6LTDK" authority="JedliEka, 1935: 89" authorityName="JedliEka" authorityPageNumber="89" authorityYear="1935" box="[302,758,191,217]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B95AEAC2FFAFB117FE9D35B2FD63FF3F" bold="true" box="[302,758,191,217]" pageId="2" pageNumber="57">
<emphasis id="B95AEAC2FFAFB117FE9D35B2FDB5FF3E" bold="true" box="[302,544,193,217]" italics="true" pageId="2" pageNumber="57">Hexagonia andrewesi</emphasis>
<bibRefCitation id="EFBF4B21FFAFB117FD9435B3FD63FF3F" author="JedliEka A." box="[551,758,191,216]" pageId="2" pageNumber="57" refId="ref6710" refString="JedliEka A., 1935. Novy Druh rodu Hexagonia (Col. Carab.). &quot; asopis &quot; eskoslovenske spoleEnosti entomologicke 32: 89." type="journal volume" year="1935">JedliËka, 1935: 89</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAFB117FE9D3592FC29FF1F" pageId="2" pageNumber="57" type="description">
<paragraph id="8B9136D0FFAFB117FE9D3592FE31FF1F" blockId="2.[302,758,191,248]" box="[302,420,225,248]" pageId="2" pageNumber="57">
(
<figureCitation id="13152A55FFAFB117FE853592FE14FF1F" box="[310,385,225,248]" captionStart="Fig" captionStartId="2.[302,340,1245,1268]" captionTargetBox="[302,811,297,1211]" captionTargetId="figure-679@2.[302,812,288,1213]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Hexagonia andrewesi JedliËka, 1935, Holotype, male. Scale bar 5 mm." pageId="2" pageNumber="57">Figs. 1</figureCitation>
,
<figureCitation id="13152A55FFAFB117FE3D3592FE09FF1F" box="[398,412,225,248]" captionStart="Figs" captionStartId="10.[302,352,1399,1422]" captionTargetBox="[307,1358,191,1362]" captionTargetId="figure-177@10.[302,1360,189,1366]" captionTargetPageId="10" captionText="Figs. 711. Aedeagus of Hexagonia, A dorsal view, B lateral view. 7 H. andrewesi JedliËka, 1935, Holotype; 8 H. castanea JedliËka, 1936 syn. n. Cotype; 9 H. sauteri Dupuis, 1912, Holotype. Scale bar 1 mm. 10 H. amicitia sp. n., Holotype; 11 H. nigrocyanea sp. n. Holotype. Scale bar 1 mm." pageId="2" pageNumber="57">7</figureCitation>
)
</paragraph>
<paragraph id="8B9136D0FFAFB117FCE135B2FC29FF1F" blockId="2.[850,1360,193,248]" pageId="2" pageNumber="57">
014404850” &amp;
<emphasis id="B95AEAC2FFAFB117FC4535B2FBBFFF3F" box="[1014,1066,193,216]" italics="true" pageId="2" pageNumber="57">idem</emphasis>
but “NHMUK 014404853” (BMNH).
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAFB117FCE13453FB78FDFF" pageId="2" pageNumber="57" type="diagnosis">
<paragraph id="8B9136D0FFAFB117FCE13453FB78FDFF" blockId="2.[850,1360,288,536]" pageId="2" pageNumber="57">
<emphasis id="B95AEAC2FFAFB117FCE13453FC52FEDF" bold="true" box="[850,967,288,312]" pageId="2" pageNumber="57">Diagnosis.</emphasis>
Very similar to
<taxonomicName id="4C2E4D53FFAFB117FBDD3452FB70FEDF" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[1134,1253,289,312]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="amicitia" status="sp. n.">
<emphasis id="B95AEAC2FFAFB117FBDD3452FB70FEDF" box="[1134,1253,289,312]" italics="true" pageId="2" pageNumber="57">H. amicitia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26957B9FFAFB117FB593452FAB1FEDF" box="[1258,1316,289,312]" pageId="2" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
, but different in the longer and more prominent temples; asymetric labrum; pronotum without lateral setiferous pores, slightly produced hind angles of pronotum and different length/width ratio of elytra. In addition it can be diagnosed by the presence of obtuse humeral angle and absence of setiferous pores in interval 5 of elytra.
</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAFB117FCE13733FBDEFD3F" pageId="2" pageNumber="57" type="description">
<paragraph id="8B9136D0FFAFB117FCE13733FBDEFD3F" blockId="2.[850,1361,576,728]" pageId="2" pageNumber="57">
<emphasis id="B95AEAC2FFAFB117FCE13733FC6CFDBF" bold="true" box="[850,1017,576,600]" pageId="2" pageNumber="57">Redescription.</emphasis>
Body length: 11.5 12.0 mm. Width/length of head: 0.92; width/length of pronotum: 0.9; length/width of elytra: 1.7. Body, antennae and legs uniformly black or slightly brownish, palpi brown.
</paragraph>
</subSubSection>
<caption id="DF516658FFAFB117FE9D31AEFD41FAF3" pageId="2" pageNumber="57" startId="2.[302,340,1245,1268]" targetBox="[302,811,297,1211]" targetPageId="2" targetType="figure">
<paragraph id="8B9136D0FFAFB117FE9D31AEFD41FAF3" blockId="2.[302,812,1243,1300]" pageId="2" pageNumber="57">
Fig. 1. Habitus of
<taxonomicName id="4C2E4D53FFAFB117FE5431AEFEF3FAF3" authority="JedliEka, 1935" authorityName="JedliEka" authorityYear="1935" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B95AEAC2FFAFB117FE5431AEFD5FFB13" box="[487,714,1245,1268]" italics="true" pageId="2" pageNumber="57">Hexagonia andrewesi</emphasis>
<bibRefCitation id="EFBF4B21FFAFB117FD7C31AEFEF3FAF3" author="JedliEka A." pageId="2" pageNumber="57" refId="ref6710" refString="JedliEka A., 1935. Novy Druh rodu Hexagonia (Col. Carab.). &quot; asopis &quot; eskoslovenske spoleEnosti entomologicke 32: 89." type="journal volume" year="1935">JedliËka, 1935</bibRefCitation>
</taxonomicName>
, Holotype, male. Scale bar 5 mm.
</paragraph>
</caption>
<subSubSection id="C334655BFFAFB116FE9D3031FD9FFABF" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57" type="materials_examined">
<paragraph id="8B9136D0FFAFB117FE9D3031FC5DFA7F" blockId="2.[302,813,1346,1754]" lastBlockId="2.[850,1361,769,1432]" pageId="2" pageNumber="57">
<emphasis id="B95AEAC2FFAFB117FE9D3031FE4EFABD" bold="true" box="[302,475,1346,1370]" pageId="2" pageNumber="57">Type material:</emphasis>
<materialsCitation id="3B463C8DFFAFB117FE563030FE01F9FD" collectingDate="1915-12" collectionCode="BMNH" collectorName="Coll. Bottcher" country="Philippines" determinerName="Ing. JedliEka" location="Dinagat" pageId="2" pageNumber="57" specimenCount="1" stateProvince="Dinagat Islands" typeStatus="holotype">
<typeStatus id="54958872FFAFB117FE563030FDD9FABD" box="[485,588,1347,1370]" pageId="2" pageNumber="57" type="holotype">Holotype</typeStatus>
, male
<collectingCountry id="F3397640FFAFB117FD093030FEEFFA9D" name="Philippines" pageId="2" pageNumber="57">Philippine Islands</collectingCountry>
| Dingat Is. [
<collectingRegion id="49EAF832FFAFB117FDBA3010FDF1FA9D" box="[521,612,1379,1402]" country="Philippines" name="Dinagat Islands" pageId="2" pageNumber="57">Dinagat</collectingRegion>
] | 163 |
<date id="FF901010FFAFB117FD0E3010FCB5FA9D" box="[701,800,1379,1402]" pageId="2" pageNumber="57" value="1915-12">
<collectingDate id="EFD4E9F8FFAFB117FD0E3010FCB5FA9D" box="[701,800,1379,1402]" pageId="2" pageNumber="57" value="1915-12">XII.1915</collectingDate>
</date>
|
<collectorName id="26DB5306FFAFB117FE9D30F0FE5EFA7D" box="[302,459,1411,1434]" pageId="2" pageNumber="57">Coll. Bottcher</collectorName>
”, “Philippine Is. |
<collectorName id="26DB5306FFAFB117FD3930F0FCB7FA7D" box="[650,802,1411,1434]" pageId="2" pageNumber="57">Coll. Bottcher</collectorName>
| B.M. 1929­ 201”, “
<taxonomicName id="4C2E4D53FFAFB117FE4530D0FDF8FA5D" authorityName="Kirby" authorityYear="1825" box="[502,621,1443,1466]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">Hexagonia</taxonomicName>
| andrewesi | type sp. n. [handwritten]| det.
<determinerName id="7E4DDA14FFAFB117FDF730B0FD46FA3D" box="[580,723,1473,1498]" pageId="2" pageNumber="57">Ing. JedliËka</determinerName>
”, “Type [round red label]”, “NHMUK 014404858” (
<collectionCode id="ED3FAE15FFAFB117FE853370FE19F9FD" box="[310,396,1539,1562]" pageId="2" pageNumber="57">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B463C8DFFAFB117FE283370FE00F95D" collectingDate="1915-12" collectionCode="BMNH" collectorName="Coll. Bottcher" country="Philippines" determinerName="Ing. JedliEka" location="Dinagat" pageId="2" pageNumber="57" specimenCount="1" stateProvince="Dinagat Islands" typeStatus="paratype">
<typeStatus id="54958872FFAFB117FE283370FD90F9FD" box="[411,517,1539,1562]" pageId="2" pageNumber="57" type="paratype">Paratypes</typeStatus>
, 2 ex “Philippine Is. |
<collectorName id="26DB5306FFAFB117FD4B3370FE1EF9DD" pageId="2" pageNumber="57">Coll. Bottcher</collectorName>
| B.M. 1929­ 201”, “
<taxonomicName id="4C2E4D53FFAFB117FDD33350FD42F9DD" authorityName="Kirby" authorityYear="1825" box="[608,727,1571,1594]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">Hexagonia</taxonomicName>
| andrewesi | sp. n. [handwritten]| det.
<determinerName id="7E4DDA14FFAFB117FD3D3330FC88F9BD" box="[654,797,1601,1626]" pageId="2" pageNumber="57">Ing. JedliËka</determinerName>
”, “Cotype [round yellow label]”, “NHMUK 014404929” &amp; idem but “NHMUK 014404872” (
<collectionCode id="ED3FAE15FFAFB117FE8533D0FE18F95D" box="[310,397,1699,1722]" pageId="2" pageNumber="57">BMNH</collectionCode>
)
</materialsCitation>
.
<emphasis id="B95AEAC2FFAFB117FE1333D1FDCCF95D" bold="true" box="[416,601,1698,1722]" pageId="2" pageNumber="57">Other material:</emphasis>
2 ex “Philippine Is. | Coll. Bottcher | B.M. 1929­ 201”, “NHMUK Head with elongate hexagonal shape; dorsally depressed, slightly narrower than the prothorax. Eye moderately large and markedly protruded, 0.6 times as long as temples; temples not protruding; gradually rounded to the constriction of the neck. Mandibles elongate, slightly incurved in basal two thirds, then strongly incurved towards apex. Palpi elongate, narrowed towards apex, smooth. Mentum with wide triangular tooth, bisetose. Antenna relatively long, two apical antennomeres go beyond the middle lenght of pronotum; three basal antennomeres and basal half of antennomere 4 glabrous. Clypeus with two pores and two shallow pits, apical margin of clypeus slightly concave; clypeal suture very shallow and laterally effaced. Labrum transverse and strongly asymmetric, margin with wide triangular incision, right angle much longer than left. Frons with two deep, elongate, parallel sided pits. Postorbital pore isolated. Surface with indistinct microreticulation, shiny.
</paragraph>
<paragraph id="8B9136D0FFAFB117FCE130B2FAC5F93F" blockId="2.[850,1361,1473,1752]" pageId="2" pageNumber="57">Pronotum wider than the head, widest at the middle, almost as long as wide; dorsal surface depressed.Anterior margin straight, anterior angles completely rounded. Lateral margins evenly curved, with short sinuation before hind angles; basal angles right, slightly produced; base of pronotum concave. Median line very narrow and moderately deep, impunctate, almost reaching anterior and posterior margins of pronotum;</paragraph>
<paragraph id="8B9136D0FFAEB116FD8F35E2FC38FF43" blockId="3.[572,941,145,164]" box="[572,941,145,164]" pageId="3" pageNumber="58">
<emphasis id="B95AEAC2FFAEB116FD8F35E2FC38FF43" box="[572,941,145,164]" italics="true" pageId="3" pageNumber="58">
Anichtchenko A., Sciaky R.,
<taxonomicName id="4C2E4D53FFAEB116FC8435E2FCECFF43" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[823,889,145,164]" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">Medina</taxonomicName>
M.N.
</emphasis>
</paragraph>
<paragraph id="8B9136D0FFAEB116FF5035B2FED5FE7F" blockId="3.[227,738,193,408]" pageId="3" pageNumber="58">anterior and posterior transverse sulcus obsoleted. Apex and base not margined; lateral borders narrowly margined, marginal gutter relatively wide in the middle part of pronotum. Lateral and posterior setiferous setae absent. Dorsal surface smooth, with hardly visible transverse microreticulation.</paragraph>
<paragraph id="8B9136D0FFAEB116FF5034B2FD54FC5F" blockId="3.[227,737,449,952]" pageId="3" pageNumber="58">Elytra elongate, markedly depressed, parallel sided. Base straight, humeri with obtuse humeral angle; lateral margins straight. Basal margin concave, connected at an obtuse angle with lateral margin at position between striae 4 and 5. Scutellar stria long, but very effaced. All striae deeply impressed and smooth; sutural stria effaced near scutellum. Intervals flat and smooth. Scutellar seta situated at base of stria 1. Interval 3 with two setiferous pores, the anterior pore situated in stria 3 at the level of the end of scutellar stria; discal pore absent; posterior one situated in apical fourth, near stria 2. Marginal series widely interrupted in the middle, consisting of 910 pores. Intervals impunctate and with well visible microreticulation, consisting of polygonal meshes, glossy.</paragraph>
<paragraph id="8B9136D0FFAEB116FF503692FE12FB9F" blockId="3.[227,737,993,1144]" pageId="3" pageNumber="58">Legs of average size. Three basal tarsomeres of all legs triangular, very wide and dorsally smooth; tarsomere 4 wide and very deeply excised, with dense brush­like pilosity on ventral surface. Tarsal claws edentate.</paragraph>
<paragraph id="8B9136D0FFAEB116FF5031D2FE79FAFF" blockId="3.[227,737,1185,1304]" pageId="3" pageNumber="58">
Aedeagus (
<figureCitation id="13152A55FFAEB116FEE631D2FE07FB5F" box="[341,402,1185,1208]" captionStart="Figs" captionStartId="10.[302,352,1399,1422]" captionTargetBox="[307,1358,191,1362]" captionTargetId="figure-177@10.[302,1360,189,1366]" captionTargetPageId="10" captionText="Figs. 711. Aedeagus of Hexagonia, A dorsal view, B lateral view. 7 H. andrewesi JedliËka, 1935, Holotype; 8 H. castanea JedliËka, 1936 syn. n. Cotype; 9 H. sauteri Dupuis, 1912, Holotype. Scale bar 1 mm. 10 H. amicitia sp. n., Holotype; 11 H. nigrocyanea sp. n. Holotype. Scale bar 1 mm." pageId="3" pageNumber="58">Fig. 7</figureCitation>
) elongate, ventral surface almost straight to apex.Apex of median lobe moderately long and slightly downturned. Endophallus with big semicircular sclerite.
</paragraph>
<paragraph id="8B9136D0FFAEB116FF503032FD9FFABF" blockId="3.[227,522,1345,1368]" box="[227,522,1345,1368]" pageId="3" pageNumber="58">Female genitalia unstudied.</paragraph>
</subSubSection>
<subSubSection id="C334655BFFAEB116FF5030F3FA95FEBF" pageId="3" pageNumber="58" type="distribution">
<paragraph id="8B9136D0FFAEB116FF5030F3FE56F9DF" blockId="3.[227,737,1408,1592]" pageId="3" pageNumber="58">
<emphasis id="B95AEAC2FFAEB116FF5030F3FEE3FA7F" bold="true" box="[227,374,1408,1432]" pageId="3" pageNumber="58">Distribution.</emphasis>
This species known with certainty only from
<collectingRegion id="49EAF832FFAEB116FEE230D2FE33FA5F" box="[337,422,1441,1464]" country="Philippines" name="Dinagat Islands" pageId="3" pageNumber="58">Dinagat</collectingRegion>
island (
<collectingRegion id="49EAF832FFAEB116FE4430D2FD4FFA5F" box="[503,730,1441,1464]" country="Philippines" name="Dinagat Islands" pageId="3" pageNumber="58">Dinagat Islands prov.</collectingRegion>
, Caraga region). It is an island situated to the north­east of Mindanao. All the other known specimens bear generic labels mentioning only “
<collectingCountry id="F3397640FFAEB116FF5F3352FE20F9DF" box="[236,437,1569,1592]" name="Philippines" pageId="3" pageNumber="58">Philippine Islands</collectingCountry>
”.
</paragraph>
<paragraph id="8B9136D0FFAEB116FF503313FA95FEBF" blockId="3.[227,737,1632,1752]" lastBlockId="3.[774,1285,193,344]" pageId="3" pageNumber="58">
<emphasis id="B95AEAC2FFAEB116FF503313FEDBF99F" bold="true" box="[227,334,1632,1656]" pageId="3" pageNumber="58">Affinities</emphasis>
. This species is similar to
<taxonomicName id="4C2E4D53FFAEB116FDD03312FD4FF99F" authorityName="Anichtchenko &amp; Sciaky &amp; Medina" authorityYear="2022" box="[611,730,1633,1656]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="amicitia">
<emphasis id="B95AEAC2FFAEB116FDD03312FD4FF99F" box="[611,730,1633,1656]" italics="true" pageId="3" pageNumber="58">H. amicitia</emphasis>
</taxonomicName>
, although larger. The two share a similar general structure, but differ in the elytral punctuation, the length/width ratio of the elytra, and the shape of the head and of the pronotum. The only species, among those known to us, to which there two can be compared is
<taxonomicName id="4C2E4D53FFAEB116FC673472FA94FEFF" authority="Van de Pool, 1889" authorityName="Van de Pool" authorityYear="1889" box="[980,1281,257,280]" class="Insecta" family="Carabidae" genus="Hexagonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B95AEAC2FFAEB116FC673472FBA9FEFF" box="[980,1084,257,280]" italics="true" pageId="3" pageNumber="58">H. nigrita</emphasis>
Van de Pool, 1889
</taxonomicName>
, widely diffused in south­east Asia from
<collectingCountry id="F3397640FFAEB116FB003452FB7FFEDF" box="[1203,1258,289,312]" name="India" pageId="3" pageNumber="58">India</collectingCountry>
to
<collectingCountry id="F3397640FFAEB116FCB53432FCE7FEBF" box="[774,882,321,344]" name="Indonesia" pageId="3" pageNumber="58">Indonesia</collectingCountry>
, but not known from the
<collectingCountry id="F3397640FFAEB116FBCC3432FB69FEBF" box="[1151,1276,321,344]" name="Philippines" pageId="3" pageNumber="58">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="7701650CADEB8EF910E03C8EE37555CC">A new species of Callimetopus Blanchard, 1853 (Coleoptera: Cerambycidae) from Panay Island, Philippines</mods:title>
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<mods:namePart id="65A19A5A94CF26D43007CCE1C2BA3435">Cabras, Analyn Anzano</mods:namePart>
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<emphasis id="B954EADB81582C729565C1AFFB8F6698" bold="true" box="[774,1051,427,451]" italics="true" pageId="1" pageNumber="134">Callimetopus panayensis</emphasis>
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(
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)
</paragraph>
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<emphasis id="B954EADB81582C729565C20FFC326578" bold="true" box="[774,934,523,547]" pageId="1" pageNumber="134">Type material</emphasis>
</paragraph>
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<typeStatus id="549B886B81582C729565C22FFCE46518" box="[774,880,555,579]" pageId="1" pageNumber="134" type="holotype">Holotype</typeStatus>
:
</emphasis>
male. White handwritten label:
<collectingCountry id="F337765981582C7292DEC228FCD66538" name="Philippines" pageId="1" pageNumber="134">Philippines</collectingCountry>
, Panay /
<collectingRegion id="49E4F82B81582C7295CFC248FC656538" box="[940,1009,588,611]" country="Philippines" name="Aklan" pageId="1" pageNumber="134">Aklan</collectingRegion>
, 09.2018, local collector leg. // Red handwritten label:
<typeStatus id="549B886B81582C72922DC268FB6665D8" box="[1102,1266,620,643]" pageId="1" pageNumber="134" type="holotype">HOLOTYPUS</typeStatus>
: /
<emphasis id="B954EADB81582C729565C288FBB165F8" box="[774,1061,652,675]" italics="true" pageId="1" pageNumber="134">
<taxonomicName id="4C204D4A81582C729565C288FC0265F8" authorityName="Blanchard" authorityYear="1853" box="[774,918,652,675]" class="Insecta" family="Cerambycidae" genus="Callimetopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="genus">Callimetopus</taxonomicName>
/
<taxonomicName id="4C204D4A81582C7295CEC288FBB165F8" authorityName="Barševskis &amp; Cabras &amp; Medina" authorityYear="2022" box="[941,1061,652,675]" class="Insecta" family="Cerambycidae" genus="Callimetopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="panayensis" status="sp. nov.">panayensis</taxonomicName>
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<emphasis id="B954EADB81582C72924EC288FBEE65F8" box="[1069,1146,652,675]" italics="true" pageId="1" pageNumber="134">sp. nov</emphasis>
.
</taxonomicNameLabel>
/ A.Barśevskis, M.N.Medina / &amp; A.Cabras det. 2022. [handwrited]. Deposited in
<collectionCode id="ED31AE0C81582C729592C2C8FBAB65B8" box="[1009,1087,716,739]" pageId="1" pageNumber="134">DUBC</collectionCode>
.
</paragraph>
<paragraph id="8B9F36C981582C729680C3C5FD7663C3" blockId="1.[227,738,961,1176]" pageId="1" pageNumber="134">
This paper presents the fifth species of
<taxonomicName id="4C204D4A81582C7294E0C3C5FE8B64A3" authorityName="Blanchard" authorityYear="1853" class="Insecta" family="Cerambycidae" genus="Callimetopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B954EADB81582C7294E0C3C5FE8B64A3" italics="true" pageId="1" pageNumber="134">Callimetopus</emphasis>
</taxonomicName>
from Panay Island. Four species were previously known from this island:
<taxonomicName id="4C204D4A81582C729409C405FEF66363" authority="Heller, 1924" authorityName="Heller" authorityYear="1924" class="Insecta" family="Cerambycidae" genus="Callimetopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="cordifer">
<emphasis id="B954EADB81582C729409C405FD756343" box="[618,737,1025,1048]" italics="true" pageId="1" pageNumber="134">C. cordifer</emphasis>
Heller, 1924
</taxonomicName>
,
<taxonomicName id="4C204D4A81582C72970AC425FDF86363" authority="Breuning 1960" authorityName="Breuning" authorityYear="1960" box="[361,620,1057,1080]" class="Insecta" family="Cerambycidae" genus="Callimetopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="B954EADB81582C72970AC425FE5B6363" box="[361,463,1057,1080]" italics="true" pageId="1" pageNumber="134">C. griseus</emphasis>
Breuning 1960
</taxonomicName>
;
<taxonomicName id="4C204D4A81582C729417C425FE066303" authority="Barsevskis 2019" authorityName="Barsevskis" authorityYear="2019" class="Insecta" family="Cerambycidae" genus="Callimetopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="kalninsi">
<emphasis id="B954EADB81582C729417C425FD746363" box="[628,736,1057,1080]" italics="true" pageId="1" pageNumber="134">C. kalninsi</emphasis>
<bibRefCitation id="EFB14B3881582C729680C445FE066303" author="Barsevskis A." box="[227,402,1087,1112]" pageId="1" pageNumber="134" pagination="7 - 10" refId="ref2053" refString="Barsevskis A. 2019. A new species of the genus Callimetopus Blanchard, 1853 (Coleoptera: Cerambycidae) from Luzon Island. Studies and Reports. Taxonomical Series, 15 (1): 7 - 10." type="journal article" year="2019">Barśevskis 2019</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C204D4A81582C7297F8C445FD4E6303" authority="(Schultze, 1920)" baseAuthorityName="Schultze" baseAuthorityYear="1920" box="[411,730,1089,1112]" class="Insecta" family="Cerambycidae" genus="Callimetopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="panayanus">
<emphasis id="B954EADB81582C7297F8C445FDBD6303" box="[411,553,1089,1112]" italics="true" pageId="1" pageNumber="134">C. panayanus</emphasis>
(Schultze, 1920)
</taxonomicName>
. Detailed description and photographic reference of other species in the
<collectingCountry id="F337765981582C7297A1C485FDA263C3" box="[450,566,1153,1176]" name="Philippines" pageId="1" pageNumber="134">Philippines</collectingCountry>
is also provided.
</paragraph>
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</treatment>
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(
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)
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<materialsCitation id="3B4D26550461FFDEFFD9FD5781ED70FE" country="Russia" county="Baltic" location="Baltic" municipality="Fanti" pageId="1" specimenCount="1" typeStatus="holotype">
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<typeStatus id="549E92AA0461FFDEFFD9FD5781C6709A" box="[114,226,647,671]" pageId="1" type="holotype">Holotype</typeStatus>
.
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<location id="8EFA7AD30461FFDEFF44FD57800A709A" LSID="urn:lsid:plazi:treatment:038C9D1E0461FFDDFFD9FDFA820073D8:8EFA7AD30461FFDEFF44FD57800A709A" box="[239,302,647,671]" country="Russia" county="Baltic" municipality="Fanti" name="Baltic" pageId="1">Baltic</location>
amber, collection
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, access code BaA08RU (BaA =
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amber; 08 = sequential number; RU =
<collectingCountry id="F3326C980461FFDEFFD9FD33819A70FE" box="[114,190,739,763]" name="Russia" pageId="1">Russia</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9A2C080461FFDEFFD9FCF080367152" blockId="1.[114,593,800,855]" pageId="1">
<emphasis id="B951F01A0461FFDEFFD9FCF08030713D" bold="true" box="[114,276,800,824]" pageId="1">Type locality.</emphasis>
Yantarny mine,
<collectingRegion id="49E1E2EA0461FFDEFE60FCF0819D7152" country="Russia" name="Kaliningrad" pageId="1">Kaliningrad region</collectingRegion>
,
<collectingCountry id="F3326C980461FFDEFF6FFCEF802A7152" box="[196,270,831,855]" name="Russia" pageId="1">Russia</collectingCountry>
.
</paragraph>
<paragraph id="8B9A2C080461FFDEFFD9FCAC800971D4" blockId="1.[114,592,892,977]" pageId="1">
<emphasis id="B951F01A0461FFDEFFD9FCAC803D7191" bold="true" box="[114,281,892,916]" pageId="1">Type horizon.</emphasis>
Middle Eocene: Lutetian (47.841.2 Mya) to late Eocene: Priabonian (37.833.9 Mya).
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830461FFDDFFD9FC2783A077D4" lastPageId="2" pageId="1" type="diagnosis">
<paragraph id="8B9A2C080461FFDEFFD9FC2783757450" blockId="1.[114,593,1015,1621]" pageId="1">
<emphasis id="B951F01A0461FFDEFFD9FC278051760A" bold="true" box="[114,373,1015,1039]" pageId="1">Differential diagnosis.</emphasis>
Compared to the new species described here,
<taxonomicName id="4C25578B0461FFDEFFD9FBE4818C766E" authority="Schaufuss, 1892" authorityName="Schaufuss" authorityYear="1892" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="cerambyx">
<emphasis id="B951F01A0461FFDEFFD9FBE480E57649" box="[114,449,1076,1100]" italics="true" pageId="1">Cacomorphocerus cerambyx</emphasis>
<bibRefCitation id="EFB451F90461FFDEFE7CFBE4818C766E" author="Schaufuss C. F. C." pageId="1" refId="ref3297" refString="Schaufuss C. F. C. 1892. Preussens Bernstein-Kafer. Neue Formen aus der Helm'schen Sammlung im Danziger Provinzialmuseum. Berliner Entomologische Zeitschrift 36 (I): 53 - 64 [1891]." type="book" year="1892">Schaufuss, 1892</bibRefCitation>
</taxonomicName>
shows a different pronotum, which is square, with narrower apical margin and a longitudinal impression on the disc (
<bibRefCitation id="EFB451F90461FFDEFFD2FB7F800976C2" author="Schaufuss C. F. C." box="[121,301,1199,1223]" pageId="1" refId="ref3297" refString="Schaufuss C. F. C. 1892. Preussens Bernstein-Kafer. Neue Formen aus der Helm'schen Sammlung im Danziger Provinzialmuseum. Berliner Entomologische Zeitschrift 36 (I): 53 - 64 [1891]." type="book" year="1892">Schaufuss 1892</bibRefCitation>
;
<bibRefCitation id="EFB451F90461FFDEFE94FB7F819676E0" author="Fanti F. &amp; Kupryjanowicz J." pageId="1" pagination="149 - 153" refId="ref2527" refString="Fanti F., Kupryjanowicz J. 2018. Discovery of a new fossil soldier beetle in Eocene Baltic amber, with the establishment of the new tribe Cacomorphocerini. Annales de Paleontologie 104: 149 - 153." type="journal article" year="2018">Fanti &amp; Kupryjanowicz 2018</bibRefCitation>
).
<taxonomicName id="4C25578B0461FFDEFF70FB1E80987701" authority="Fanti &amp; Kupryjanowicz, 2018" authorityName="Fanti &amp; Kupryjanowicz" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="wiszniewskii">
<emphasis id="B951F01A0461FFDEFF70FB1E837476E3" box="[219,592,1230,1254]" italics="true" pageId="1">Cacomorphocerus wiszniewskii</emphasis>
<bibRefCitation id="EFB451F90461FFDEFFD9FB3C80987701" author="Fanti F. &amp; Kupryjanowicz J." box="[114,444,1260,1284]" pageId="1" pagination="149 - 153" refId="ref2527" refString="Fanti F., Kupryjanowicz J. 2018. Discovery of a new fossil soldier beetle in Eocene Baltic amber, with the establishment of the new tribe Cacomorphocerini. Annales de Paleontologie 104: 149 - 153." type="journal article" year="2018">Fanti &amp; Kupryjanowicz, 2018</bibRefCitation>
</taxonomicName>
differs from the new species in the subrectangular pronotum, with sides slightly concave from base to the apical third, where a very small denticle is present, and in the less saucer-shaped antennomeres IV-VI (
<bibRefCitation id="EFB451F90461FFDEFE48FA55804F77B9" author="Fanti F. &amp; Kupryjanowicz J." pageId="1" pagination="149 - 153" refId="ref2527" refString="Fanti F., Kupryjanowicz J. 2018. Discovery of a new fossil soldier beetle in Eocene Baltic amber, with the establishment of the new tribe Cacomorphocerini. Annales de Paleontologie 104: 149 - 153." type="journal article" year="2018">Fanti &amp; Kupryjanowicz 2018</bibRefCitation>
).
<taxonomicName id="4C25578B0461FFDEFE2EFA7480DE77DE" authority="(Kuska &amp; Kania, 2010)" baseAuthorityName="Kuska &amp; Kania" baseAuthorityYear="2010" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="jantaricus">
<emphasis id="B951F01A0461FFDEFE2EFA7481C177DF" italics="true" pageId="1">Cacomorphocerus jantaricus</emphasis>
(
<bibRefCitation id="EFB451F90461FFDEFF58FA1380D777DE" author="Kuska A. &amp; Kania I." box="[243,499,1475,1499]" pageId="1" pagination="49 - 56" refId="ref2935" refString="Kuska A., Kania I. 2010. New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa 2400: 49 - 56." type="journal article" year="2010">Kuśka &amp; Kania, 2010</bibRefCitation>
)
</taxonomicName>
shows antennomeres differently shaped and pronotum more rectangular, with sides slightly protruding laterally near the apical margin (
<bibRefCitation id="EFB451F90461FFDEFF7BF9ED80EC7450" author="Kuska A. &amp; Kania I." box="[208,456,1597,1621]" pageId="1" pagination="49 - 56" refId="ref2935" refString="Kuska A., Kania I. 2010. New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa 2400: 49 - 56." type="journal article" year="2010">Kuśka &amp; Kania 2010</bibRefCitation>
). The other
</paragraph>
<caption id="DF5A7C800461FFDEFDDBFB8882A9770F" pageId="1" startId="1.[624,663,1112,1136]" targetBox="[624,1070,263,1074]" targetPageId="1" targetType="figure">
<paragraph id="8B9A2C080461FFDEFDDBFB8882A9770F" blockId="1.[624,1103,1112,1290]" pageId="1">
Fig. 1.
<taxonomicName id="4C25578B0461FFDEFD6AFB8882D57675" authorityName="Sarker &amp; Talukder" authorityYear="2023" box="[705,1009,1112,1136]" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="incurvus" status="sp. nov.">
<emphasis id="B951F01A0461FFDEFD6AFB8882D57675" box="[705,1009,1112,1136]" italics="true" pageId="1">Cacomorphocerus incurvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624D610461FFDEFC5FFB88856B7675" box="[1012,1103,1112,1136]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
in Baltic amber (code BaA08RU). A: Holotype, dorsal view, scale bar = 1.0 mm; B: Holotype, ventral view, scale bar = 1.0 mm; C: Holotype, detail of head (ventral view), scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B9A2C080461FFDEFFD9F976829477A6" blockId="1.[114,154,1702,1726]" lastBlockId="1.[624,1103,1327,1627]" pageId="1">
234
<emphasis id="B951F01A0461FFDEFDDBFAFF82257742" bold="true" box="[624,769,1327,1351]" pageId="1">Description.</emphasis>
Sex undefined, adult, robust, winged. Body length: about
<quantity id="4CDD81ED0461FFDEFC6FFA9E856B7763" box="[964,1103,1358,1382]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="7.5" metricValueMin="7.3" pageId="1" unit="mm" value="7.4" valueMax="7.5" valueMin="7.3">7.3-7.5 mm</quantity>
(the specimen is folded), elytra
<quantity id="4CDD81ED0461FFDEFC7DFABC856F7781" box="[982,1099,1388,1412]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.1" metricValueMin="5.0" pageId="1" unit="mm" value="5.05" valueMax="5.1" valueMin="5.0">5-5.1 mm</quantity>
. Entirely blackish-dark brown.
</paragraph>
<paragraph id="8B9A2C080461FFDDFDDBFA7A801372ED" blockId="1.[624,1103,1327,1627]" lastBlockId="2.[189,669,178,1489]" lastPageId="2" pageId="1">Head elongated, rounded behind the eyes, covered with long setae and a shallow punctation. Eyes roundish, convex and prominent, inserted in the upper-lateral part of the head. Mandibles elongated, falciform. Maxillary palps 4-segmented; last palpomere securiform. Labial palps 3- segmented.</paragraph>
<paragraph id="8B9A2C080462FFDDFF16FF3F80CA7090" blockId="2.[189,669,178,1489]" pageId="2">Antennae 12-segmented, short, reaching about the elytral half, strongly pubescent; scape short, stout, club-shaped; antennomere II very short, globular, about 2.7 times shorter than scape; antennomeres IIIVII saucer-shaped and asymmetrically inflated; antennomere VIII strongly saucer-shaped and asymmetrically dilated with apical lobes, sturdier than previous ones; antennomere IX squared, thinner than previous one; antennomeres XXII elongated, filiform, rather robust, X-XI slightly enlarged apically, last antennomere thinner and rounded at apex.</paragraph>
<paragraph id="8B9A2C080462FFDDFF16FD4C804B7169" blockId="2.[189,669,178,1489]" pageId="2">Pronotum transverse, about as wide as head; apical and basal margins straight; sides straight, without denticles; surface without impressions, covered with shallow granules and several setae. Scutellum triangular, with pointed apex which is rounded at apex.</paragraph>
<paragraph id="8B9A2C080462FFDDFF16FCA3837F7600" blockId="2.[189,669,178,1489]" pageId="2">Elytra long, covering and surpassing the last abdominal segment, parallel-sided and with rounded apex, wider than pronotum; surface smooth and pubescent. Hind wings transparent, slightly longer than elytra.</paragraph>
<paragraph id="8B9A2C080462FFDDFF16FBDC831E771C" blockId="2.[189,669,178,1489]" pageId="2">Metasternum wide, short, robust, rounded apically and pubescent. Sternites transverse and pubescent. Legs slender and short; coxae very large; trochanters elongated, rounded apically; femora cylindrical, straight, more robust than tibiae; tibiae cylindrical, pro- and mesotibiae very slightly longer than pro- and mesofemora, metatibiae longer than metafemora.</paragraph>
<caption id="DF5A7C800462FFDDFD10FBEA823076AB" pageId="2" startId="2.[699,738,1082,1106]" targetBox="[699,1184,754,1045]" targetPageId="2" targetType="figure">
<paragraph id="8B9A2C080462FFDDFD10FBEA823076AB" blockId="2.[699,1179,1082,1198]" pageId="2">
Fig. 2.
<taxonomicName id="4C25578B0462FFDDFCA7FBEA85577654" authorityName="Sarker &amp; Talukder" authorityYear="2023" box="[780,1139,1082,1106]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="amicitiae" status="sp. nov." subGenus="Malthinus">
<emphasis id="B951F01A0462FFDDFCA7FBEA825B7657" box="[780,895,1082,1106]" italics="true" pageId="2">Malthinus</emphasis>
(
<emphasis id="B951F01A0462FFDDFC20FBEA82D37657" box="[907,1015,1082,1106]" italics="true" pageId="2">Malthinus</emphasis>
)
<emphasis id="B951F01A0462FFDDFBA2FBEA85577654" box="[1033,1139,1082,1105]" italics="true" pageId="2">amicitiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624D610462FFDDFBDEFBEA83CD7675" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
in Baltic amber (code BaA09RU). Holotype, dorso-lateral view, scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B9A2C080462FFDDFF16FAF083A077D4" blockId="2.[189,669,178,1489]" pageId="2">Tarsi 5-segmented, short; tarsomere I elongated, tarsomeres II and III triangular and shorter than I, tarsomere IV bilobed, tarsomere V elongated but not extremely thin and enlarged apically; claws simple with a very small and obtuse tooth at base.</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDDFF16FA27850D72ED" pageId="2" type="etymology">
<paragraph id="8B9A2C080462FFDDFF16FA27800F746E" blockId="2.[189,668,1527,1643]" pageId="2">
<emphasis id="B951F01A0462FFDDFF16FA278060740A" bold="true" box="[189,324,1527,1551]" pageId="2">Etymology.</emphasis>
The specific epithet derives from the Latin adjective
<taxonomicName id="4C25578B0462FFDDFE55F9C583427428" authorityName="Sarker &amp; Talukder" authorityYear="2023" box="[510,614,1557,1581]" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="incurvus">
<emphasis id="B951F01A0462FFDDFDA1F9C683427428" box="[522,614,1558,1581]" italics="true" pageId="2">incurvus</emphasis>
</taxonomicName>
” = curved, in reference to the pronotum and head bent.
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FF61850D72ED" blockId="2.[699,1178,177,232]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FF61824072CC" bold="true" box="[699,868,177,201]" pageId="2">Syninclusions.</emphasis>
Enhydro and air bubbles, botanical fragments, stellate hairs.
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDDFD10FEDD820073D8" pageId="2" type="discussion">
<paragraph id="8B9A2C080462FFDDFD10FEDD820073D8" blockId="2.[699,1179,269,477]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FEDD82147320" bold="true" box="[699,816,269,293]" pageId="2">Remarks.</emphasis>
The oblong reddish yellow amber piece measures approximately 41x20x
<quantity id="4CDD81ED0462FFDDFCA6FE9B827B7366" box="[781,863,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" unit="mm" value="10.0">10 mm</quantity>
. The inclusion is complete, with some folded parts and an open elytron showing the hind wing. Some parts of the body are slightly covered with white emulsion.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,185 @@
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<subSubSection id="C33F7F830462FFDDFD10FDD383D3703C" pageId="2" type="nomenclature">
<paragraph id="8B9A2C080462FFDDFD10FDD385BE701E" blockId="2.[699,1178,515,631]" box="[699,1178,515,539]" pageId="2">
<heading id="D0D29B640462FFDDFD10FDD385BE701E" bold="true" box="[699,1178,515,539]" centered="true" fontSize="10" level="2" pageId="2" reason="3">
<emphasis id="B951F01A0462FFDDFD10FDD385BE701E" bold="true" box="[699,1178,515,539]" pageId="2">
Subfamily
<taxonomicName id="4C25578B0462FFDDFCE4FDD385B1701E" authority="Kiesenwetter" authorityName="Kiesenwetter" authorityYear="1852" box="[847,1173,515,539]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="subFamily" subFamily="Malthininae">Malthininae Kiesenwetter</taxonomicName>
,
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FDF183D3703C" blockId="2.[699,1178,515,631]" box="[699,759,545,569]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FDF183D3703C" bold="true" box="[699,759,545,569]" pageId="2">1852</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDDFD10FD90822F70D6" pageId="2" type="description">
<paragraph id="8B9A2C080462FFDDFD10FD908572705D" blockId="2.[699,1178,515,631]" box="[699,1110,576,600]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FD908572705D" bold="true" box="[699,1110,576,600]" pageId="2">
Tribe
<taxonomicName id="4C25578B0462FFDDFD54FD908572705D" authority="Kiesenwetter, 1852" authorityName="Kiesenwetter" authorityYear="1852" box="[767,1110,576,600]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="tribe" tribe="Malthinini">Malthinini Kiesenwetter, 1852</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FD8F850C7072" blockId="2.[699,1178,515,631]" box="[699,1064,607,631]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FD8F850C7072" bold="true" box="[699,1064,607,631]" pageId="2">
Genus
<taxonomicName id="4C25578B0462FFDDFCA1FD8F850C7072" authority="Latreille, 1806" authorityName="Latreille" authorityYear="1806" box="[778,1064,607,631]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B951F01A0462FFDDFCA1FD8F82A77072" bold="true" box="[778,899,607,631]" italics="true" pageId="2">Malthinus</emphasis>
Latreille, 1806
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FD4C85B770B1" blockId="2.[699,1171,668,723]" box="[699,1171,668,692]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FD4C83EC70B1" bold="true" box="[699,712,668,692]" pageId="2"></emphasis>
<taxonomicName id="4C25578B0462FFDDFD62FD4C851570B1" authorityName="Sarker &amp; Talukder" authorityYear="2023" box="[713,1073,668,692]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="amicitiae" status="sp. nov." subGenus="Malthinus">
<emphasis id="B951F01A0462FFDDFD62FD4C829870B1" bold="true" box="[713,956,668,692]" pageId="2">
<emphasis id="B951F01A0462FFDDFD62FD4C826570B1" bold="true" box="[713,833,668,692]" italics="true" pageId="2">Malthinus</emphasis>
(
<emphasis id="B951F01A0462FFDDFCE1FD4C829870B1" bold="true" box="[842,956,668,692]" italics="true" pageId="2">Malthinus</emphasis>
</emphasis>
)amicitiae
</taxonomicName>
<emphasis id="B951F01A0462FFDDFB9CFD4C85B770B1" bold="true" box="[1079,1171,668,692]" pageId="2">
<taxonomicNameLabel id="A2624D610462FFDDFB9CFD4C85B770B1" box="[1079,1171,668,692]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FD6B822F70D6" blockId="2.[699,1171,668,723]" box="[699,779,699,723]" pageId="2">
(
<figureCitation id="131E308D0462FFDDFD68FD6B822770D6" box="[707,771,699,723]" captionStart="Fig" captionStartId="2.[699,738,1082,1106]" captionTargetBox="[699,1184,754,1045]" captionTargetId="figure-329@2.[699,1184,754,1045]" captionTargetPageId="2" captionText="Fig. 2.Malthinus(Malthinus)amicitiaesp. nov. in Baltic amber (code BaA09RU). Holotype, dorso-lateral view, scale bar = 1.0 mm." pageId="2">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDDFD10FB038253741B" pageId="2" type="materials_examined">
<paragraph id="8B9A2C080462FFDDFD10FB0382367742" blockId="2.[699,1178,1235,1351]" pageId="2">
<materialsCitation id="3B4D26550462FFDDFD10FB0382367742" country="Russia" county="Baltic" location="Baltic" municipality="Fanti" pageId="2" specimenCount="1" typeStatus="holotype">
<emphasis id="B951F01A0462FFDDFD10FB03821476EE" bold="true" box="[699,816,1235,1259]" pageId="2">
<typeStatus id="549E92AA0462FFDDFD10FB03820F76EE" box="[699,811,1235,1259]" pageId="2" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EFA7AD30462FFDDFC92FB03825C76EE" LSID="urn:lsid:plazi:treatment:038C9D1E0462FFDCFD10FDD382BD7770:8EFA7AD30462FFDDFC92FB03825C76EE" box="[825,888,1235,1259]" country="Russia" county="Baltic" municipality="Fanti" name="Baltic" pageId="2">Baltic</location>
amber, collection
<collectingMunicipality id="6BFEB6720462FFDDFBFDFB0385B176EE" box="[1110,1173,1235,1259]" pageId="2">Fanti</collectingMunicipality>
, access code BaA09RU (BaA =
<collectingCounty id="62FB54840462FFDDFBFFFB2285BE770F" box="[1108,1178,1266,1290]" pageId="2">Baltic</collectingCounty>
amber; 09 = sequential number; RU =
<collectingCountry id="F3326C980462FFDDFD10FAFF82237742" box="[699,775,1327,1351]" name="Russia" pageId="2">Russia</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FABC827F77A6" blockId="2.[699,1178,1388,1443]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FABC82797781" bold="true" box="[699,861,1388,1412]" pageId="2">Type locality.</emphasis>
Yantarny mine,
<collectingRegion id="49E1E2EA0462FFDDFBBFFABC822677A6" country="Russia" name="Kaliningrad" pageId="2">Kaliningrad region</collectingRegion>
,
<collectingCountry id="F3326C980462FFDDFCA6FA5B827377A6" box="[781,855,1419,1443]" name="Russia" pageId="2">Russia</collectingCountry>
.
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FA188253741B" blockId="2.[699,1178,1480,1566]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FA18824777E5" bold="true" box="[699,867,1480,1504]" pageId="2">Type horizon.</emphasis>
Middle Eocene: Lutetian (47.841.2 Mya) to late Eocene: Priabonian (37.833.9 Mya).
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDCFD10F99381D173D8" lastPageId="3" pageId="2" type="diagnosis">
<paragraph id="8B9A2C080462FFDDFD10F99385BE747F" blockId="2.[699,1178,1603,1658]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10F993829C745E" bold="true" box="[699,952,1603,1627]" pageId="2">Differential diagnosis.</emphasis>
The new species is easily recognizable by its neck gradually
</paragraph>
<paragraph id="8B9A2C080462FFDCFBC7F97681D173D8" blockId="2.[1132,1172,1702,1726]" lastBlockId="3.[114,593,178,477]" lastPageId="3" pageId="2">
235 constricted posteriorly. It is similar only to the extinct
<taxonomicName id="4C25578B0463FFDCFF55FF01818E7302" authority="Kuska &amp; Kania, 2010" authorityName="Kuska &amp; Kania" authorityYear="2010" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="danieli">
<emphasis id="B951F01A0463FFDCFF55FF01805272ED" box="[254,374,208,232]" italics="true" pageId="3">M. danieli</emphasis>
<bibRefCitation id="EFB451F90463FFDCFE23FF00818E7302" author="Kuska A. &amp; Kania I." pageId="3" pagination="49 - 56" refId="ref2935" refString="Kuska A., Kania I. 2010. New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa 2400: 49 - 56." type="journal article" year="2010">Kuśka &amp; Kania, 2010</bibRefCitation>
</taxonomicName>
, from which it differs in the pronotum not narrower anteriorly, the elytra without impressed punctuation and the different colour (
<bibRefCitation id="EFB451F90463FFDCFF61FE9B80E27366" author="Kuska A. &amp; Kania I." box="[202,454,331,355]" pageId="3" pagination="49 - 56" refId="ref2935" refString="Kuska A., Kania I. 2010. New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa 2400: 49 - 56." type="journal article" year="2010">Kuśka &amp; Kania 2010</bibRefCitation>
). The other fossil
<taxonomicName id="4C25578B0463FFDCFF19FEBA80567387" box="[178,370,362,386]" pageId="3">
<emphasis id="B951F01A0463FFDCFF19FEBA803A7387" box="[178,286,362,386]" italics="true" pageId="3">Malthinus</emphasis>
species
</taxonomicName>
show head strongly restricted behind eyes and dotted elytra (
<bibRefCitation id="EFB451F90463FFDCFFD2FE7780B973BA" author="Fanti F. &amp; Damgaard A. L." box="[121,413,423,447]" pageId="3" pagination="1 - 32" refId="ref2486" refString="Fanti F., Damgaard A. L. 2018. Fossil soldier beetles from Baltic amber of the Anders Damgaard amber collection (Coleoptera Cantharidae). Baltic Journal of Coleopterology 18 (1): 1 - 32." type="journal article" year="2018">Fanti &amp; Damgaard 2018</bibRefCitation>
;
<bibRefCitation id="EFB451F90463FFDCFE1AFE7781CD73D8" author="Pankowski M. G. &amp; Fanti F." pageId="3" pagination="548 - 556" refId="ref3008" refString="Pankowski M. G., Fanti F. 2022. Two new fossil species of soldier beetles (Coleoptera, Cantharidae, Malthi-ninae) from Baltic amber. Zootaxa 5165 (4): 548 - 556." type="journal article" year="2022">Pankowski &amp; Fanti 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830463FFDCFFD9FDD382CA71CD" pageId="3" type="description">
<paragraph id="8B9A2C080463FFDCFFD9FDD381F6705D" blockId="3.[114,593,515,1643]" pageId="3">
<emphasis id="B951F01A0463FFDCFFD9FDD38027701E" bold="true" box="[114,259,515,539]" pageId="3">Description.</emphasis>
Female, adult, robust, winged. Body length: about 6.0-
<quantity id="4CDD81ED0463FFDCFE2EFDF280C7703F" box="[389,483,546,570]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="3" unit="mm" value="6.2">6.2 mm</quantity>
. Entirely blackish.
</paragraph>
<paragraph id="8B9A2C080463FFDCFFD9FD8F8000718E" blockId="3.[114,593,515,1643]" pageId="3">Head almost completely exposed, neck gradually constricted posteriorly, equipped with several long setae and impressed small punctation. Eyes large, convex, rounded, inserted in the lateral-upper part of head. Mandibles falciform. Maxillary palps 4- segmented; last palpomere globularelongated and apically pointed. Labial palps 3-segmented; last palpomere globular and apically pointed.</paragraph>
<paragraph id="8B9A2C080463FFDCFFD9FC42800076FE" blockId="3.[114,593,515,1643]" pageId="3">Antennae 11-segmented, filiform, relatively short, surpassing the elytral half, covered with several setae; scape elongated, club-shaped; antennomere II short; antennomeres III-IV sub-equal, elongated, rather robust; antennomeres V-VII sub-equal, slightly longer than previous ones; antennomere VIII longer and thinner than previous ones; antennomeres IX-X sub-equal, shorter and very slightly sturdier than previous one; antennomere XI elongated, rounded at apex.</paragraph>
<paragraph id="8B9A2C080463FFDCFFD9FAD18064746E" blockId="3.[114,593,515,1643]" pageId="3">Pronotum rectangular, slightly longer than wide, slightly narrower than head; apical margin very slightly rounded and not bordered; anterior corners rounded; sides straight and strongly bordered; basal margin straight and strongly bordered; posterior corners almost acute; surface irregular, gibbous on the basal part and concave near the apical margin, equipped with some long setae, without punctation. Scutellum triangular, wide at base, pointed apically, with rounded apex.</paragraph>
<paragraph id="8B9A2C080463FFDCFFD9F97683F57387" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1103,178,968]" pageId="3">236 Elytra not particularly short, revealing the last abdominal segment uncovered, slightly wider than pronotum, parallel-sided, not dehiscent, rounded at apex; surface rather wrinkled, covered with several long setae and without punctation. Hind wings as long as elytra.</paragraph>
<paragraph id="8B9A2C080463FFDCFDDBFE5882857115" blockId="3.[624,1103,178,968]" pageId="3">Metasternum with slightly rounded posterior margin, covered with dispersed short setae and very shallow punctation. Sternites transverse, wide, pubescent. Last tergite and last sternite wide and rounded. Legs robust, pubescent; coxae elongated, stout, enlarged at base; trochanters elongated, rounded at apex; femora enlarged, curved in the middle; tibiae cylindrical and thin, equipped with an apical spur, protibiae shorter than profemora, meso- and metatibiae longer than meso- and metafemora.</paragraph>
<paragraph id="8B9A2C080463FFDCFDDBFCC782CA71CD" blockId="3.[624,1103,178,968]" pageId="3">Tarsi 5-segmented, pubescent; tarsomere I elongated; tarsomere II about 1.4 times shorter than tarsomere I; tarsomere III shorter than previous one; tarsomere IV bilobed; tarsomere V short, thin, curved; claws simple, robust, without tooth.</paragraph>
</subSubSection>
<subSubSection id="C33F7F830463FFDCFDDBFC3D827676B8" pageId="3" type="etymology">
<paragraph id="8B9A2C080463FFDCFDDBFC3D82897664" blockId="3.[624,1103,1005,1121]" pageId="3">
<emphasis id="B951F01A0463FFDCFDDBFC3D83D47600" bold="true" box="[624,752,1005,1029]" pageId="3">Etymology.</emphasis>
The specific epithet derives from the Latin noun “
<emphasis id="B951F01A0463FFDCFC1CFBDC852A7621" box="[951,1038,1036,1060]" italics="true" pageId="3">ămīcĭtĭa</emphasis>
” = friendship, in reference to the friendship that binds the two co-authors.
</paragraph>
<paragraph id="8B9A2C080463FFDCFDDBFB57827676B8" blockId="3.[624,1102,1159,1213]" pageId="3">
<emphasis id="B951F01A0463FFDCFDDBFB578236769A" bold="true" box="[624,786,1159,1183]" pageId="3">Syninclusions.</emphasis>
Botanical fragments, air bubbles, stellate hair.
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830463FFDCFDDBFB3382BD7770" pageId="3" type="discussion">
<paragraph id="8B9A2C080463FFDCFDDBFB3382BD7770" blockId="3.[624,1103,1251,1397]" pageId="3">
<emphasis id="B951F01A0463FFDCFDDBFB3383FB76FE" bold="true" box="[624,735,1251,1275]" pageId="3">Remarks.</emphasis>
The oblong yellow amber piece measures approximately 30x20x
<quantity id="4CDD81ED0463FFDCFC7BFAD18537771C" box="[976,1043,1281,1305]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="3" unit="mm" value="9.0">9 mm</quantity>
. The inclusion is complete and clearly visible on one side, while it has shows some oxidations on the other one.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,248 @@
<document id="E14C8E1B4A292CCC06EC496041DAB09D" ID-DOI="10.59893/bjc.23(2).004" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657628258" checkinUser="felipe" docAuthor="Minkina, Łukasz" docDate="2023" docId="038D87AAEE00FFA2A1601491FF30F96E" docLanguage="en" docName="BalticJColeopterol.23.2.205-208.pdf" docOrigin="Baltic Journal of Coleopterology (Basel, Switzerland) 23 (2)" docSource="http://dx.doi.org/10.3389/fpsyt.2024.1436142" docStyle="DocumentStyle:514B3A4252435B0AFD496C900C6D7572.1:BalticJColeop.2023-.journal_article" docStyleId="514B3A4252435B0AFD496C900C6D7572" docStyleName="BalticJColeop.2023-.journal_article" docStyleVersion="1" docTitle="Airapus madangensis Minkina, 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="208" masterDocId="FFB4FFD2EE01FFA1A1121637FFAAFFD0" masterDocTitle="A new species of the genusAirapus Stebnicka &amp; Howden, 1996 (Coleoptera: Scarabaeidae: Aphodiinae) from Papua New Guinea" masterLastPageNumber="208" masterPageNumber="205" pageNumber="206" updateTime="1722690880365" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="211FD9629890EF9FC39C095667E818EF">A new species of the genusAirapus Stebnicka &amp; Howden, 1996 (Coleoptera: Scarabaeidae: Aphodiinae) from Papua New Guinea</mods:title>
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<mods:date id="22FA4A2845FE4CC94943DD965AE6B914">2023</mods:date>
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<treatment id="038D87AAEE00FFA2A1601491FF30F96E" LSID="urn:lsid:plazi:treatment:038D87AAEE00FFA2A1601491FF30F96E" httpUri="http://treatment.plazi.org/id/038D87AAEE00FFA2A1601491FF30F96E" lastPageId="3" lastPageNumber="208" pageId="1" pageNumber="206">
<subSubSection id="C33E6537EE00FFA0A1601491FE11FD6D" box="[114,443,678,702]" pageId="1" pageNumber="206" type="nomenclature">
<paragraph id="8B9B36BCEE00FFA0A1601491FE11FD6D" blockId="1.[114,443,678,702]" box="[114,443,678,702]" pageId="1" pageNumber="206">
<heading id="D0D381D0EE00FFA0A1601491FE11FD6D" box="[114,443,678,702]" fontSize="10" level="2" pageId="1" pageNumber="206" reason="7">
<taxonomicName id="4C244D3FEE00FFA0A1601491FEF5FD6E" authorityName="Minkina" authorityYear="2023" box="[114,351,678,702]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="206" phylum="Arthropoda" rank="species" species="madangensis" status="sp. nov.">
<emphasis id="B950EAAEEE00FFA0A1601491FEF5FD6E" bold="true" box="[114,351,678,702]" italics="true" pageId="1" pageNumber="206">Airapus madangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357D5EE00FFA0A0741491FE11FD6D" box="[358,443,678,701]" pageId="1" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E6537EE00FFA0A16014D4FF76FD2A" box="[114,220,738,763]" pageId="1" pageNumber="206" type="description">
<paragraph id="8B9B36BCEE00FFA0A16014D4FF76FD2A" blockId="1.[114,220,738,763]" box="[114,220,738,763]" pageId="1" pageNumber="206">
(
<figureCitation id="131F2A39EE00FFA0A16B14D4FF78FD2A" box="[121,210,738,763]" captionStart-0="Figs 1-3" captionStart-1="Figs 4-5" captionStartId-0="1.[624,670,1480,1504]" captionStartId-1="2.[189,235,1258,1282]" captionTargetBox-0="[628,1108,848,1443]" captionTargetBox-1="[198,672,611,1204]" captionTargetId-0="figure-392@1.[624,1112,846,1443]" captionTargetId-1="figure-391@2.[189,677,600,1218]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs 1-3.Airapus madangensis sp. nov.,♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." captionText-1="Figs 4-5. Airapus madangensis sp. nov.,♀, holotype: 4- head; 5- epipharynx. Fig. 4-5: scale lines: Fig. 4: 1.0 mm; Fig. 5: 0.2 mm." pageId="1" pageNumber="206">Figs 1-5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E6537EE00FFA0A1601517FED9FB78" pageId="1" pageNumber="206" type="materials_examined">
<paragraph id="8B9B36BCEE00FFA0A1601517FE6CFC87" blockId="1.[114,593,800,855]" pageId="1" pageNumber="206">
<emphasis id="B950EAAEEE00FFA0A1601517FEA4FCE8" bold="true" box="[114,270,800,824]" pageId="1" pageNumber="206">Type locality.</emphasis>
<collectingCountry id="F333762CEE00FFA0A0071516FE42FCE8" box="[277,488,800,824]" name="Papua New Guinea" pageId="1" pageNumber="206">Papua New Guinea</collectingCountry>
,
<collectingRegion id="49E0F85EEE00FFA0A0E11517FF06FC86" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="206">Madang prov.</collectingRegion>
, Hansa Bay, Laing Island.
</paragraph>
<paragraph id="8B9B36BCEE00FFA0A3621685FC97FEB2" blockId="1.[624,1103,178,355]" pageId="1" pageNumber="206">coarse, with variable in size, rounded punctures; in the middle very dense, coarse, with somewhat variable in size, longitudinal punctures; anteriorly moderately dense, not so coarse, irregular in size, with rounded punctures.</paragraph>
<paragraph id="8B9B36BCEE00FFA0A36217BFFCF7FCFF" blockId="1.[624,1103,392,815]" pageId="1" pageNumber="206">
Epipharynx (
<figureCitation id="131F2A39EE00FFA0A3EF17BFFCEAFE70" box="[765,832,392,416]" captionStart="Figs 4-5" captionStartId="2.[189,235,1258,1282]" captionTargetBox="[198,672,611,1204]" captionTargetId="figure-391@2.[189,677,600,1218]" captionTargetPageId="2" captionText="Figs 4-5. Airapus madangensis sp. nov.,♀, holotype: 4- head; 5- epipharynx. Fig. 4-5: scale lines: Fig. 4: 1.0 mm; Fig. 5: 0.2 mm." pageId="1" pageNumber="206">Fig. 5</figureCitation>
) transverse, with lateral sides broadly rounded, anterior margin of pedia concavely arcuate, corypha below of margin of pedia, indistinct, with two very short, thick celtes. Mesoepitorma shortened basally, somewhat pear-shaped. Acanthopariae with dense, very short and thin chaetae; acropariae with dense, short and thiner chaetae than on acanthopariae; chaetopariae with dense belt of thick, long chaetae; adelochaetae similar to chaetopariae, but less dense; prophobae very short and thin chaetae; chaetopediae without chaetae. Tormae long.
</paragraph>
<paragraph id="8B9B36BCEE00FFA0A160154BFEDEFBDF" blockId="1.[114,593,892,1192]" pageId="1" pageNumber="206">
<emphasis id="B950EAAEEE00FFA0A160154BFE85FC44" bold="true" box="[114,303,892,916]" pageId="1" pageNumber="206">Type material.</emphasis>
<materialsCitation id="3B4C3CE1EE00FFA0A058154BFEDEFBDF" collectingDate="1981-06-08" collectionCode="R, IRSN" collectorName="Coll. I. &amp; Sc. N. B. &amp; W. Laing Island &amp; J. Van Goethem" country="Papua New Guinea" county="Laing Island" location="Bay" municipality="Hansa" pageId="1" pageNumber="206" specimenCount="1" specimenCount-female="1" stateProvince="Madang" typeStatus="holotype">
<emphasis id="B950EAAEEE00FFA0A058154BFE12FC44" bold="true" box="[330,440,892,916]" pageId="1" pageNumber="206">
<typeStatus id="549F881EEE00FFA0A058154BFE12FC44" box="[330,440,892,916]" pageId="1" pageNumber="206" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="9D22FD35EE00FFA0A0C4154BFE40FC43" box="[470,490,892,915]" count="1" pageId="1" pageNumber="206" type="female"></specimenCount>
):
<collectorName id="26D1536AEE00FFA0A307154BFF2AFC63" pageId="1" pageNumber="206">Coll. I.</collectorName>
<collectionCode id="ED35AE79EE00FFA0A16D15ACFF3FFC63" box="[127,149,923,947]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="206">R</collectionCode>
.
<collectorName id="26D1536AEE00FFA0A18B15ACFF47FC63" box="[153,237,923,947]" pageId="1" pageNumber="206">Sc.N.B.</collectorName>
|
<collectingCountry id="F333762CEE00FFA0A01315ACFE7BFC63" box="[257,465,923,947]" name="Papua New Guinea" pageId="1" pageNumber="206">Papua New Guinea</collectingCountry>
|
<collectorName id="26D1536AEE00FFA0A0F615ACFF1CFC01" pageId="1" pageNumber="206">
W.
<collectingCounty id="62FA4E30EE00FFA0A31E15ACFF1CFC01" pageId="1" pageNumber="206">Laing Island</collectingCounty>
</collectorName>
,
<collectingMunicipality id="6BFFACC6EE00FFA0A1D4158DFEA3FC01" box="[198,265,954,977]" pageId="1" pageNumber="206">Hansa</collectingMunicipality>
|
<location id="8EFB6067EE00FFA0A035158DFEFEFC02" LSID="urn:lsid:plazi:treatment:038D87AAEE00FFA2A1601491FF30F96E:8EFB6067EE00FFA0A035158DFEFEFC02" box="[295,340,954,978]" country="Papua New Guinea" county="Laing Island" municipality="Hansa" name="Bay" pageId="1" pageNumber="206" stateProvince="Madang">Bay</location>
,
<collectingRegion id="49E0F85EEE00FFA0A077158EFDAAFC01" box="[357,512,953,977]" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="206">Madang prov.</collectingRegion>
|
<location id="8EFB6067EE00FFA0A30C158DFDFBFC01" LSID="urn:lsid:plazi:treatment:038D87AAEE00FFA2A1601491FF30F96E:8EFB6067EE00FFA0A30C158DFDFBFC01" box="[542,593,954,977]" country="Papua New Guinea" county="Laing Island" municipality="Hansa" name="Wet" pageId="1" pageNumber="206" stateProvince="Madang">Wet</location>
leaf litter | I.G.: 26373 |
<date id="FF9A107CEE00FFA0A09B15EFFE44FC20" box="[393,494,984,1008]" pageId="1" pageNumber="206" value="1981-06-08">
<collectingDate id="EFDEE994EE00FFA0A09B15EFFE44FC20" box="[393,494,984,1008]" pageId="1" pageNumber="206" value="1981-06-08">8.vi.1981</collectingDate>
</date>
| leg.
<collectorName id="26D1536AEE00FFA0A32B15EEFEADFBDF" pageId="1" pageNumber="206">J. Van Goethem</collectorName>
|| (
<collectionCode id="ED35AE79EE00FFA0A03415C0FEC2FBDF" box="[294,360,1015,1039]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="206">IRSN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9B36BCEE00FFA0A1601222FED9FB78" blockId="1.[114,593,892,1192]" pageId="1" pageNumber="206">
<materialsCitation id="3B4C3CE1EE00FFA0A1601222FE27FBFE" box="[114,397,1045,1070]" collectionCode="R" collectorName="Coll. I." pageId="1" pageNumber="206" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B950EAAEEE00FFA0A1601222FF70FBFD" bold="true" box="[114,218,1045,1069]" pageId="1" pageNumber="206">
<typeStatus id="549F881EEE00FFA0A1601222FF70FBFD" box="[114,218,1045,1069]" pageId="1" pageNumber="206" type="paratype">Paratype</typeStatus>
</emphasis>
(
<specimenCount id="9D22FD35EE00FFA0A1E01221FEACFBFD" box="[242,262,1046,1069]" count="1" pageId="1" pageNumber="206" type="female"></specimenCount>
):
<collectorName id="26D1536AEE00FFA0A0371222FEDDFBFE" box="[293,375,1045,1070]" pageId="1" pageNumber="206">Coll. I.</collectorName>
<collectionCode id="ED35AE79EE00FFA0A0651221FE27FBFE" box="[375,397,1046,1070]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="206">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B4C3CE1EE00FFA0A0821221FEC5FB78" collectingDate="1978-05-09" collectionCode="ISEA" collectorName="Sc. N. B. &amp; J. Van Goethem" country="Papua New Guinea" location="Hills" pageId="1" pageNumber="206" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectorName id="26D1536AEE00FFA0A0821221FE4FFBFE" box="[400,485,1046,1070]" pageId="1" pageNumber="206">Sc.N.B.</collectorName>
|
<collectingCountry id="F333762CEE00FFA0A31B1221FEABFB9C" name="Papua New Guinea" pageId="1" pageNumber="206">Papua New Guinea</collectingCountry>
|
<location id="8EFB6067EE00FFA0A0341203FEF0FB9C" LSID="urn:lsid:plazi:treatment:038D87AAEE00FFA2A1601491FF30F96E:8EFB6067EE00FFA0A0341203FEF0FB9C" box="[294,346,1076,1100]" country="Papua New Guinea" name="Hills" pageId="1" pageNumber="206" stateProvince="Madang">Hills</location>
<locationDeviation id="9A292F9EEE00FFA0A0781203FD9FFB9C" box="[362,565,1076,1100]" location="Bogia" pageId="1" pageNumber="206">
<quantity id="4CDC9B59EE00FFA0A0781203FE03FB9C" box="[362,425,1076,1100]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="206" unit="km" value="4.0">4 km</quantity>
NW Bogia
</locationDeviation>
|
<collectingRegion id="49E0F85EEE00FFA0A1601264FEA1FBBA" box="[114,267,1107,1131]" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="206">Madang prov.</collectingRegion>
, forest | gallery, wet leaf litter | near tree, I.G.: 25848 |
<date id="FF9A107CEE00FFA0A09C1246FE41FB59" box="[398,491,1137,1161]" pageId="1" pageNumber="206" value="1978-05-09">
<collectingDate id="EFDEE994EE00FFA0A09C1246FE41FB59" box="[398,491,1137,1161]" pageId="1" pageNumber="206" value="1978-05-09">9.v.1978</collectingDate>
</date>
| leg.
<collectorName id="26D1536AEE00FFA0A32B1245FEADFB78" pageId="1" pageNumber="206">J. Van Goethem</collectorName>
|| (
<collectionCode id="ED35AE79EE00FFA0A03412A7FECDFB78" box="[294,359,1168,1192]" pageId="1" pageNumber="206">ISEA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E6537EE00FFA3A16012FAFB8FFA01" lastPageId="2" lastPageNumber="207" pageId="1" pageNumber="206" type="description">
<paragraph id="8B9B36BCEE00FFA0A16012FAFD99FAF3" blockId="1.[114,593,1229,1315]" pageId="1" pageNumber="206">
<emphasis id="B950EAAEEE00FFA0A16012FAFEA9FB35" bold="true" box="[114,259,1229,1253]" pageId="1" pageNumber="206">Description.</emphasis>
Dorsum (
<figureCitation id="131F2A39EE00FFA0A07E12FAFE01FB36" box="[364,427,1229,1254]" captionStart="Figs 1-3" captionStartId="1.[624,670,1480,1504]" captionTargetBox="[628,1108,848,1443]" captionTargetId="figure-392@1.[624,1112,846,1443]" captionTargetPageId="1" captionText="Figs 1-3.Airapus madangensis sp. nov.,♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." pageId="1" pageNumber="206">Fig. 1</figureCitation>
). Body length
<quantity id="4CDC9B59EE00FFA0A16012DBFF6CFAD3" box="[114,198,1260,1284]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="206" unit="mm" value="5.7">5.7 mm</quantity>
, elongate, shiny, blackish, sides and apex of elytra with distinct macrosetation.
</paragraph>
<paragraph id="8B9B36BCEE00FFA0A160137FFDFBF9A4" blockId="1.[114,593,1352,1652]" pageId="1" pageNumber="206">
Head (
<figureCitation id="131F2A39EE00FFA0A1AF137FFEA8FAB0" box="[189,258,1352,1376]" captionStart="Figs 4-5" captionStartId="2.[189,235,1258,1282]" captionTargetBox="[198,672,611,1204]" captionTargetId="figure-391@2.[189,677,600,1218]" captionTargetPageId="2" captionText="Figs 4-5. Airapus madangensis sp. nov.,♀, holotype: 4- head; 5- epipharynx. Fig. 4-5: scale lines: Fig. 4: 1.0 mm; Fig. 5: 0.2 mm." pageId="1" pageNumber="206">Fig. 4</figureCitation>
) relatively large, trapezoidal, convex, shiny, without microreticulation. Clypeus gently bordered, widely sinuate anteriorly, denticulate on sides, widely rounded laterally, not notched before genae, clypeal border without macrosetae. Genae obtuse, very distinctly exceeding eyes, with few very short, thin macrosetae. Frontal suture not noticeable, without gibbosities. Punctation simple: basally very dense, quite
</paragraph>
<caption id="DF5B6634EE00FFA0A36213FFFB9CF9CD" pageId="1" pageNumber="206" startId="1.[624,670,1480,1504]" targetBox="[628,1108,848,1443]" targetPageId="1" targetType="figure">
<paragraph id="8B9B36BCEE00FFA0A36213FFFB9CF9CD" blockId="1.[624,1103,1480,1565]" pageId="1" pageNumber="206">
Figs 1-3.
<taxonomicName id="4C244D3FEE00FFA0A3C413FFFC17FA30" authorityName="Minkina" authorityYear="2023" box="[726,957,1480,1504]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="206" phylum="Arthropoda" rank="species" species="madangensis" status="sp. nov.">
<emphasis id="B950EAAEEE00FFA0A3C413FFFC17FA30" box="[726,957,1480,1504]" italics="true" pageId="1" pageNumber="206">Airapus madangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357D5EE00FFA0A2DE13FEFB82FA30" box="[972,1064,1481,1504]" pageId="1" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
,♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B9B36BCEE00FFA3A1601091FD2CFE6E" blockId="1.[114,154,1702,1726]" lastBlockId="2.[189,669,178,447]" lastPageId="2" lastPageNumber="207" pageId="1" pageNumber="206">206 Pronotum transverse, approximately as wide as base of elytra, widest in the middle, convex, shiny, without microreticulation, with simple, quite regularly distributed punctation, punctures are coarser on disc, quite irregular in shape, mainly rounded. Base not distinctly bordered by a groove, anteriorly and on sides not bordered. Sides and base moderately long, quite thick macrosetae. Anterior angles widely rounded. Base before hind angles truncate.</paragraph>
<paragraph id="8B9B36BCEE03FFA3A1AF17D3FDB6FDEA" blockId="2.[189,668,484,570]" pageId="2" pageNumber="207">Scutellum small, triangular, with ogival sides, with few small punctures, moderately shiny, without microreticulation.</paragraph>
<caption id="DF5B6634EE03FFA3A1AF12DDFD20FAEE" pageId="2" pageNumber="207" startId="2.[189,235,1258,1282]" targetBox="[198,672,611,1204]" targetPageId="2" targetType="figure">
<paragraph id="8B9B36BCEE03FFA3A1AF12DDFD20FAEE" blockId="2.[189,668,1257,1343]" pageId="2" pageNumber="207">
Figs 4-5.
<taxonomicName id="4C244D3FEE03FFA3A03612DDFDA1FAD2" authorityName="Minkina" authorityYear="2023" box="[292,523,1258,1282]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="madangensis" status="sp. nov.">
<emphasis id="B950EAAEEE03FFA3A03612DDFDA1FAD2" box="[292,523,1258,1282]" italics="true" pageId="2" pageNumber="207">Airapus madangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357D5EE03FFA3A30B12DDFDDCFAD1" box="[537,630,1258,1281]" pageId="2" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
,♀, holotype: 4- head; 5- epipharynx. Fig. 4-5: scale lines: Fig. 4: 1.0 mm; Fig. 5: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B9B36BCEE03FFA3A1AF1353FBFDFE70" blockId="2.[189,668,1380,1650]" lastBlockId="2.[699,1179,178,416]" pageId="2" pageNumber="207">Elytra elongate, convex, parallel, shiny, with weak microreticulation (which is more distinct on sides and before apex), with short and thin macrosetae on sides and before apex; with small indistinct humeral denticles; with ten striae and ten intervals. Intervals shiny, convex, with quite regularly distributed, simple, coarse punctation; part of punctures nearby sides connected with punctures of striae and because of it striae seems indenting intervals margins here very distinctly; on sides because of that structure intervals seems to be somewhat tuberculate. Striae distinctly, quite densely punctate, with medium sized punctures; punctures distinctly indenting margins of intervals. All striae joined together before apex.</paragraph>
<paragraph id="8B9B36BCEE03FFA3A3A917F1FC1BFC18" blockId="2.[699,1179,453,968]" pageId="2" pageNumber="207">Legs. Femora shiny, without microreticulation, moderately coarsely and sparsely punctate, with punctures bearing very short macrosetae. Protibiae distinctly tridentate laterally, proximally not serrulate; dorsal side smooth, shiny, with few very fine punctures; apical spur long, moderately broad, gently downwardly and inwardly bent, with apex rounded. Meso- and metatibiae fimbriate apically with row of short spinules of equal length. Apex of metatibiae without accessory spine. Metatibiae superior apical spur very slightly shorter than basimetatarsomere, latter approximately as long as three of next metatarsomeres combined. Claws short, thin, distinctly arcuate.</paragraph>
<paragraph id="8B9B36BCEE03FFA3A3A915D9FCFDFBD5" blockId="2.[699,855,1006,1029]" box="[699,855,1006,1029]" pageId="2" pageNumber="207">Macropterous.</paragraph>
<paragraph id="8B9B36BCEE03FFA3A3A9121CFB8FFA01" blockId="2.[699,1178,1067,1489]" pageId="2" pageNumber="207">
Venter (
<figureCitation id="131F2A39EE03FFA3A20C121CFCC6FB93" box="[798,876,1067,1091]" captionStart="Figs 1-3" captionStartId="1.[624,670,1480,1504]" captionTargetBox="[628,1108,848,1443]" captionTargetId="figure-392@1.[624,1112,846,1443]" captionTargetPageId="1" captionText="Figs 1-3.Airapus madangensis sp. nov.,♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." pageId="2" pageNumber="207">Fig. 2</figureCitation>
). Meso-metaventral plate shiny, weakly convex, with distinct, shallow, quite narrow longitudinal furrow in the middle; furrow with deeper part nearby base and nearby apex, additionaly with reversed-heart shaped concavity before apex; surface with moderately coarse, sparse punctures. Abdominal ventrites moderately shiny, anteriorly weakly fluted, with weak microreticulation, with sparse, moderately coarsely, irregularly spaced punctatures; all punctures bearing very short and quite thin macrosetae. Pygidium with similar structure to ventrites.
</paragraph>
</subSubSection>
<subSubSection id="C33E6537EE03FFA2A3A913C0FF30F96E" lastPageId="3" lastPageNumber="208" pageId="2" pageNumber="207" type="etymology">
<paragraph id="8B9B36BCEE03FFA3A3A913C0FBE3F99C" blockId="2.[699,1179,1527,1612]" pageId="2" pageNumber="207">
<emphasis id="B950EAAEEE03FFA3A3A913C0FCEEF9DF" bold="true" box="[699,836,1527,1551]" pageId="2" pageNumber="207">Etymology.</emphasis>
Toponymic; an adjective derived from the name of
<collectingRegion id="49E0F85EEE03FFA3A2C01022FB3CF9FD" box="[978,1174,1557,1581]" country="Papua New Guinea" name="Madang" pageId="2" pageNumber="207">Madang province</collectingRegion>
, where the new species was collected.
</paragraph>
<paragraph id="8B9B36BCEE03FFA3A57E1091FB3EF96E" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="207">207</paragraph>
<paragraph id="8B9B36BCEE02FFA2A1601686FDA7FF38" blockId="3.[114,593,177,232]" pageId="3" pageNumber="208">
<emphasis id="B950EAAEEE02FFA2A1601686FF5FFF19" bold="true" box="[114,245,177,201]" pageId="3" pageNumber="208">Variability.</emphasis>
Body length:
<quantity id="4CDC9B59EE02FFA2A0B81686FDE7FF19" box="[426,589,177,201]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="5.7" metricValueMin="5.6" pageId="3" pageNumber="208" unit="mm" value="5.65" valueMax="5.7" valueMin="5.6">5.6 5.7mm</quantity>
. Punctation of body is weakly variable.
</paragraph>
<paragraph id="8B9B36BCEE02FFA2A160173AFE67FEF5" blockId="3.[114,461,269,293]" box="[114,461,269,293]" pageId="3" pageNumber="208">
<emphasis id="B950EAAEEE02FFA2A160173AFEFCFEF5" bold="true" box="[114,342,269,293]" pageId="3" pageNumber="208">Sexual dimorphism.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B9B36BCEE02FFA2A160177CFEC7F9BB" blockId="3.[114,593,331,1643]" pageId="3" pageNumber="208">
<emphasis id="B950EAAEEE02FFA2A160177CFF72FEB3" bold="true" box="[114,216,331,355]" pageId="3" pageNumber="208">Affinity.</emphasis>
Newly described species at first glance, due to relatively regularly convex body, not very distinctly deplanate anterior angles of the pronotum and lack of typical deplanations on sides of the pronotum, as well as the relatively normally developed elytral intervals with macrosetae only on the sides and before apex, seems to be more similar to the genus
<taxonomicName id="4C244D3FEE02FFA2A0421477FDE7FD88" authority="Harold, 1867" authorityName="Harold" authorityYear="1867" box="[336,589,576,600]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAAEEE02FFA2A0421477FE07FD88" box="[336,429,576,600]" italics="true" pageId="3" pageNumber="208">Ataenius</emphasis>
Harold, 1867
</taxonomicName>
. Besides, large distance between mesocoxae, typical shape of meso-metaventral plate, denticulate clypeus and typical punctation of head definitely place newly described species in the genus
<taxonomicName id="4C244D3FEE02FFA2A07214EDFE8BFCC0" authority="Stebnicka &amp; Howden, 1996" authorityName="Stebnicka &amp; Howden" authorityYear="1996" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAAEEE02FFA2A07214EDFE1FFD22" box="[352,437,730,754]" italics="true" pageId="3" pageNumber="208">Airapus</emphasis>
Stebnicka &amp; Howden, 1996
</taxonomicName>
.
<taxonomicName id="4C244D3FEE02FFA2A03F14CFFDB6FCC0" authorityName="Minkina" authorityYear="2023" box="[301,540,760,784]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="madangensis" status="sp. nov.">
<emphasis id="B950EAAEEE02FFA2A03F14CFFDB6FCC0" box="[301,540,760,784]" italics="true" pageId="3" pageNumber="208">Airapus madangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357D5EE02FFA2A33E14CEFF0BFCFE" pageId="3" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from all other known species due to unique structure of elytral interals (quite smooth on the disc, slightly granulate on the sides; with characteristic punctation); additionaly it cannot be including to any group of species created by
<bibRefCitation id="EFB54B4DEE02FFA2A1FB15F8FE0BFC37" author="Stebnicka Z." box="[233,417,975,999]" pageId="3" pageNumber="208" pagination="833 - 895" refId="ref2021" refString="Stebnicka Z. 1998. The Aphodiinae of New Guinea, the Bismarck Archipelago and the Solomon Islands Invertebrate Taxonomy 12 (5): 833 - 895." type="journal article" year="1998">Stebnicka (1998)</bibRefCitation>
and due of it is easily distinguishable from other species from
<collectingCountry id="F333762CEE02FFA2A1A3123AFE20FBF4" box="[177,394,1036,1060]" name="Papua New Guinea" pageId="3" pageNumber="208">Papua New Guinea</collectingCountry>
. Additional body composites that allow it to be distinguished from other species are shape of meso-metaventral plate and shape of scutellum both of which are distinctive and unique among different species of the genus. In any case, the two features mentioned need to be studied in the future in different manuscript, as they are not easy to determine (unique shapes of concavities before apex of meso-metaventral plates, unique clustering of scutellum punctation, etc.), and also with this features it will probably be easier in the future to determine the relationship of affinities between species and possible new genera (as
<taxonomicName id="4C244D3FEE02FFA2A1BC13C0FEA9F9DF" authorityName="Stebnicka &amp; Howden" authorityYear="1996" box="[174,259,1527,1551]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAAEEE02FFA2A1BC13C0FEA9F9DF" box="[174,259,1527,1551]" italics="true" pageId="3" pageNumber="208">Airapus</emphasis>
</taxonomicName>
or even
<taxonomicName id="4C244D3FEE02FFA2A0B613C0FDABF9DF" authorityName="Harold" authorityYear="1867" box="[420,513,1527,1551]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAAEEE02FFA2A0B613C0FDABF9DF" box="[420,513,1527,1551]" italics="true" pageId="3" pageNumber="208">Ataenius</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFB54B4DEE02FFA2A1601022FE84F9FE" author="Stebnicka Z." box="[114,302,1557,1582]" pageId="3" pageNumber="208" pagination="1 - 155" refId="ref2050" refString="Stebnicka Z. 2007. The genus Ataenius Harold, 1867 of New World. Iconography. Krakow: Institute of Systematics and Evolution of Animals Polish Academy of Sciences: 1 - 155." type="book chapter" year="2007">Stebnicka, 2007</bibRefCitation>
) probably paraphyletic, due to the presence of unique features between some species).
</paragraph>
<paragraph id="8B9B36BCEE02FFA2A1601091FF30F96E" blockId="3.[114,154,1702,1726]" box="[114,154,1702,1726]" pageId="3" pageNumber="208">208</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,183 @@
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<mods:title id="574153ABF8732DDED90789FBB5A01E03">Description of two new Therates Latreille, 1816 species from Philippines (Coleoptera: Cicindelidae)</mods:title>
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<paragraph id="8B9822006609FFEDD646FEA2947DFEAA" blockId="1.[774,1066,260,315]" box="[774,1066,260,284]" pageId="1" pageNumber="76">
<heading id="D0D0956C6609FFEDD646FEA2947DFEAA" bold="true" box="[774,1066,260,284]" fontSize="10" level="2" pageId="1" pageNumber="76" reason="4">
<emphasis id="B953FE126609FFEDD646FEA2947DFEAA" bold="true" box="[774,1066,260,284]" pageId="1" pageNumber="76">
<taxonomicName id="4C2759836609FFEDD646FEA293BFFEAA" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[774,1000,260,284]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="barsevskisi" status="sp. nov.">
<emphasis id="B953FE126609FFEDD646FEA293BFFEAA" bold="true" box="[774,1000,260,284]" italics="true" pageId="1" pageNumber="76">Therates barsevskisi</emphasis>
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<taxonomicNameLabel id="A26043696609FFEDD6AEFEA2947DFEAA" box="[1006,1066,260,284]" pageId="1" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="8B9822006609FFEDD646FE82932AFE8D" blockId="1.[774,1066,260,315]" box="[774,893,292,315]" pageId="1" pageNumber="76">
(
<figureCitation id="131C3E856609FFEDD64EFE829323FE8D" box="[782,884,292,315]" captionStart="Figs" captionStartId="1.[227,277,1702,1725]" captionTargetBox="[332,1180,851,1665]" captionTargetId="figure-257@1.[329,1181,847,1666]" captionTargetPageId="1" captionText="Figs. 12. Habitus of Therates barsevskisi sp. n.: 1 holotype, male; 2 paratype, female. Scale line = 1 sm." pageId="1" pageNumber="76">Figs. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33D718B6609FFEDD646FEC294B2FDAD" pageId="1" pageNumber="76" type="materials_examined">
<paragraph id="8B9822006609FFEDD646FEC294B2FDAD" blockId="1.[774,1285,356,539]" pageId="1" pageNumber="76">
<emphasis id="B953FE126609FFEDD646FEC293F8FECA" bold="true" box="[774,943,356,380]" pageId="1" pageNumber="76">Type material:</emphasis>
<materialsCitation id="3B4F285D6609FFEDD6F5FEC2938CFE0D" collectionCode="DUBC" country="Philippines" location="Antique" pageId="1" pageNumber="76" specimenCount="1" stateProvince="Antique" typeStatus="holotype">
<typeStatus id="549C9CA26609FFEDD6F5FEC2944BFECD" box="[949,1052,356,379]" pageId="1" pageNumber="76" type="holotype">Holotype</typeStatus>
, male
<collectingCountry id="F33062906609FFEDD13DFEC294AEFECD" box="[1149,1273,356,379]" name="Philippines" pageId="1" pageNumber="76">Philippines</collectingCountry>
| W Visayas,
<collectingRegion id="49E3ECE26609FFEDD6C1FE229380FE2D" box="[897,983,388,411]" country="Philippines" name="Antique" pageId="1" pageNumber="76">Antique</collectingRegion>
| Panay 2018. VII | local collector leg” (
<collectionCode id="ED36BAC56609FFEDD6C3FE029383FE0D" box="[899,980,420,443]" pageId="1" pageNumber="76">DUBC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F285D6609FFEDD6A3FE029556FE6D" collectingDate="2017-07" country="Philippines" location="Panay isl." pageId="1" pageNumber="76" specimenCount="1" specimenCount-female="1" stateProvince="Antique" typeStatus="paratype">
<typeStatus id="549C9CA26609FFEDD6A3FE029407FE0D" box="[995,1104,420,443]" pageId="1" pageNumber="76" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D21E9896609FFEDD119FE0294E3FE0D" box="[1113,1204,420,443]" count="1" pageId="1" pageNumber="76" type="female">1 female</specimenCount>
<collectingCountry id="F33062906609FFEDD198FE02930BFE6D" name="Philippines" pageId="1" pageNumber="76">Philippines</collectingCountry>
|
<location id="8EF874DB6609FFEDD62DFE629381FE6D" LSID="urn:lsid:plazi:treatment:038E93166609FFEED646FEA29465FDCE:8EF874DB6609FFEDD62DFE629381FE6D" box="[877,982,452,475]" country="Philippines" name="Panay isl." pageId="1" pageNumber="76" stateProvince="Antique">Panay isl.</location>
|
<collectingRegion id="49E3ECE26609FFEDD6A6FE62946AFE6D" box="[998,1085,452,475]" country="Philippines" name="Antique" pageId="1" pageNumber="76">Antique</collectingRegion>
<date id="FF9904C06609FFEDD102FE6294FBFE6D" box="[1090,1196,452,475]" pageId="1" pageNumber="76" value="2017-07">
<collectingDate id="EFDDFD286609FFEDD102FE6294FBFE6D" box="[1090,1196,452,475]" pageId="1" pageNumber="76" value="2017-07">VII. 2017</collectingDate>
</date>
” (cJW)
</materialsCitation>
;
<materialsCitation id="3B4F285D6609FFEDD646FE4294B6FDAD" collectingDate="2018-06" collectorName="W. Visayas &amp; M. Madia-as" country="Philippines" location="Panay island" pageId="1" pageNumber="76" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D21E9896609FFEDD646FE429303FE4D" box="[774,852,484,507]" count="1" pageId="1" pageNumber="76" type="male">1 male</specimenCount>
,
<specimenCount id="9D21E9896609FFEDD626FE42939DFE4D" box="[870,970,484,507]" count="1" pageId="1" pageNumber="76" type="female">1 female</specimenCount>
<collectingCountry id="F33062906609FFEDD6BAFE429420FE4D" box="[1018,1143,484,507]" name="Philippines" pageId="1" pageNumber="76">Philippines</collectingCountry>
,
<collectorName id="26D247D66609FFEDD1C9FE4294AAFE4D" box="[1161,1277,484,507]" pageId="1" pageNumber="76">W.Visayas</collectorName>
,
<location id="8EF874DB6609FFEDD646FDA293C5FDAD" LSID="urn:lsid:plazi:treatment:038E93166609FFEED646FEA29465FDCE:8EF874DB6609FFEDD646FDA293C5FDAD" box="[774,914,516,539]" country="Philippines" name="Panay island" pageId="1" pageNumber="76">Panay island</location>
,
<collectorName id="26D247D66609FFEDD6DDFDA2947EFDAD" box="[925,1065,516,539]" pageId="1" pageNumber="76">M. Madia­as</collectorName>
,
<date id="FF9904C06609FFEDD174FDA294D8FDAD" box="[1076,1167,516,539]" pageId="1" pageNumber="76" value="2018-06">
<collectingDate id="EFDDFD286609FFEDD174FDA294D8FDAD" box="[1076,1167,516,539]" pageId="1" pageNumber="76" value="2018-06">VI.2018</collectingDate>
</date>
” (cRS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D718B6609FFEDD646FDE294B1FCAD" pageId="1" pageNumber="76" type="diagnosis">
<paragraph id="8B9822006609FFEDD646FDE294B1FCAD" blockId="1.[774,1285,580,795]" pageId="1" pageNumber="76">
<emphasis id="B953FE126609FFEDD646FDE2932BFDEA" bold="true" box="[774,892,580,604]" pageId="1" pageNumber="76">Diagnosis.</emphasis>
This new species belongs to
<taxonomicName id="4C2759836609FFEDD1FCFDE29364FDCD" baseAuthorityName="Fabricius" baseAuthorityYear="1801" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B953FE126609FFEDD189FDE29364FDCD" italics="true" pageId="1" pageNumber="76">fasciatus</emphasis>
</taxonomicName>
group, characterized by clypeus without setae, and by elytra without tumescence at mid length. By its elytral pattern it is similar to
<taxonomicName id="4C2759836609FFEDD1B1FD0293F1FD6D" authorityName="W. Horn" authorityYear="1928" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="pseudosemperi">
<emphasis id="B953FE126609FFEDD1B1FD0293F1FD6D" italics="true" pageId="1" pageNumber="76">T. pseudosemperi</emphasis>
</taxonomicName>
(Mindoro isl.) and
<taxonomicName id="4C2759836609FFEDD130FD629552FD6D" authorityName="Zettel &amp; Pangantihon" authorityYear="2017" box="[1136,1285,708,731]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="negrosicola">
<emphasis id="B953FE126609FFEDD130FD629552FD6D" box="[1136,1285,708,731]" italics="true" pageId="1" pageNumber="76">T. negrosicola</emphasis>
</taxonomicName>
(Negros isl.), but this new species can be easily distinguished by its entirely black pronotum.
</paragraph>
</subSubSection>
<caption id="DF5872886609FFEDD5A3F9009135F96B" pageId="1" pageNumber="76" startId="1.[227,277,1702,1725]" targetBox="[332,1180,851,1665]" targetPageId="1" targetType="figure">
<paragraph id="8B9822006609FFEDD5A3F9009135F96B" blockId="1.[227,1285,1702,1757]" pageId="1" pageNumber="76">
Figs. 12. Habitus of
<taxonomicName id="4C2759836609FFEDD491F90192F8F908" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[465,687,1703,1726]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="barsevskisi" status="sp. nov.">
<emphasis id="B953FE126609FFEDD491F90192F8F908" box="[465,687,1703,1726]" italics="true" pageId="1" pageNumber="76">Therates barsevskisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26043696609FFEDD7F7F90092A3F90B" box="[695,756,1702,1725]" pageId="1" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
: 1 holotype, male; 2 paratype, female. Scale line = 1 sm.
</paragraph>
</caption>
<subSubSection id="C33D718B660AFFEED46EFF66944AFECE" pageId="2" pageNumber="77" type="description">
<paragraph id="8B982200660AFFEED46EFF669133FF4E" blockId="2.[302,812,192,248]" pageId="2" pageNumber="77">
<emphasis id="B953FE12660AFFEED46EFF6691EFFF6E" bold="true" box="[302,440,192,216]" pageId="2" pageNumber="77">Description.</emphasis>
Body length 11.0­
<quantity id="4CDF8FE5660AFFEED733FF679282FF6E" box="[627,725,193,216]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="77" unit="mm" value="11.5">11.5 mm</quantity>
, mean = 11.1.
</paragraph>
<paragraph id="8B982200660AFFEED46EFE879345FECE" blockId="2.[302,812,289,376]" pageId="2" pageNumber="77">Head and pronotum black with light gold luster; elytra black with gold luster and orange transverse bands. Labrum, mouth parts and legs orange.</paragraph>
<paragraph id="8B982200660AFFEED46EFE07937BFD8E" blockId="2.[302,813,417,920]" pageId="2" pageNumber="77">Head without microsculpture, temples not protruding, slightly convergent towards pronotum; clypeus without setae; frons wide, frontal grooves shallow, gradually curved; eyes very protruding; each supra­orbital plate with one seta. Labrum</paragraph>
<paragraph id="8B982200660AFFEED46EFDE7937AFC2E" blockId="2.[302,813,417,920]" pageId="2" pageNumber="77">
(
<figureCitation id="131C3E85660AFFEED476FDE791C1FDEE" box="[310,406,577,600]" captionStart="Figs" captionStartId="3.[884,934,1327,1350]" captionTargetBox="[229,848,779,1447]" captionTargetId="figure-302@3.[226,851,779,1448]" captionTargetPageId="3" captionText="Figs. 47. Labrum of Therates spp. 4 T. barsevskisi sp. n., male; 5 idem, female; 67 T. insolitus sp. n., female. Scale line = 1 mm." pageId="2" pageNumber="77">Figs. 67</figureCitation>
) relatively long; with extremely shallow microsculpture, almost isodiametric on the disc and slightly transverse on sides; with nine sub marginal setae and ten teeth: with six short frontal central teeths, with rounded apices in four central and slightly pointed in lateral ones, as well as a pair of slightly longer and narrower latero­apical teeth and longer and pointed latero­basal teeth on both sides. Mandibles orange with dark brown tooths. Antennae relatively long; blackish brown, scape and pedicel yellow, except blackish dorsal side.
</paragraph>
<paragraph id="8B982200660AFFEED46EFC67944AFECE" blockId="2.[302,813,961,1016]" lastBlockId="2.[850,1361,193,376]" pageId="2" pageNumber="77">
Pronotum as long as wide, smooth; midline barely visible; anterior and posterior grooves glabrous. Aedeagus (
<figureCitation id="131C3E85660AFFEED68BFF67945CFF6E" box="[971,1035,193,216]" captionStart="Fig" captionStartId="3.[884,923,1665,1688]" captionTargetBox="[228,849,1478,1752]" captionTargetId="figure-393@3.[226,851,1475,1754]" captionTargetPageId="3" captionText="Fig. 8. Aedeagus of Therates barsevskisi sp. n., holotype. Scale line = 1 mm." pageId="2" pageNumber="77">Fig. 8</figureCitation>
) relatively large; with a wide, gradually curved basal bulb; ventral side of median lobe straight; apical third moderately downturned, apex relatively wide and rounded; internal sack with big and thick flagellum, hookshaped at the base.
</paragraph>
</subSubSection>
<subSubSection id="C33D718B660AFFEED612FE069555FE0E" box="[850,1282,416,440]" pageId="2" pageNumber="77" type="distribution">
<paragraph id="8B982200660AFFEED612FE069555FE0E" blockId="2.[850,1282,416,440]" box="[850,1282,416,440]" pageId="2" pageNumber="77">
<emphasis id="B953FE12660AFFEED612FE069388FE0E" bold="true" box="[850,991,416,440]" pageId="2" pageNumber="77">Distribution</emphasis>
.
<collectingCountry id="F3306290660AFFEED6ACFE07943FFE0E" box="[1004,1128,417,440]" name="Philippines" pageId="2" pageNumber="77">Philippines</collectingCountry>
, Panay island.
</paragraph>
</subSubSection>
<subSubSection id="C33D718B660AFFEED612FE469465FDCE" pageId="2" pageNumber="77" type="etymology">
<paragraph id="8B982200660AFFEED612FE469465FDCE" blockId="2.[850,1360,480,632]" pageId="2" pageNumber="77">
<emphasis id="B953FE12660AFFEED612FE46939CFE4E" bold="true" box="[850,971,480,504]" pageId="2" pageNumber="77">Etymology</emphasis>
. Species is named after our friend and colleague Dr. Arvids Bar
<emphasis id="B953FE12660AFFEED122FE59943BFDAE" bold="true" box="[1122,1132,511,536]" pageId="2" pageNumber="77">š</emphasis>
evskis, a coleopterist in
<collectingRegion id="49E3ECE2660AFFEED62BFD8793B4FD8E" box="[875,995,545,568]" country="Latvia" name="Daugavpils" pageId="2" pageNumber="77">Daugavpils</collectingRegion>
University (
<collectingCountry id="F3306290660AFFEED123FD8794FEFD8E" box="[1123,1193,545,568]" name="Latvia" pageId="2" pageNumber="77">Latvia</collectingCountry>
) in appreciation of his contribution to the study of coleopterofauna of
<collectingCountry id="F3306290660AFFEED6F1FDC79479FDCE" box="[945,1070,609,632]" name="Philippines" pageId="2" pageNumber="77">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,131 @@
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<mods:title id="F0604B5BEDEC13D12D732CD07EE04496">Baltic Journal of Coleopterology</mods:title>
</mods:titleInfo>
<mods:part id="6314112EA7355D07C622D24A4B89CFD4">
<mods:date id="B9864CD1FD0FE7FE8475236C823582CB">2008</mods:date>
<mods:detail id="55E77315E6FA4BBC80AB839C0240BC8B" type="volume">
<mods:number id="FEBD3C22F5B57CC6543E033CE002F473">8</mods:number>
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</mods:mods>
<treatment id="038F8784FF93FF9BFCCC48AE438DFD38" ID-DOI="http://doi.org/10.5281/zenodo.13199519" ID-Zenodo-Dep="13199519" LSID="urn:lsid:plazi:treatment:038F8784FF93FF9BFCCC48AE438DFD38" httpUri="http://treatment.plazi.org/id/038F8784FF93FF9BFCCC48AE438DFD38" lastPageId="3" pageId="1">
<subSubSection id="C33C6519FF93FF99FCCC48AE468DFBD8" pageId="1" type="nomenclature">
<paragraph id="8B993692FF93FF99FCCC48AE46ACFC79" blockId="1.[810,1154,957,981]" box="[810,1154,957,981]" pageId="1">
<heading id="D0D181FEFF93FF99FCCC48AE46ACFC79" bold="true" box="[810,1154,957,981]" fontSize="10" level="2" pageId="1" reason="4">
<taxonomicName id="4C264D11FF93FF99FCCC48AE46ACFC79" ID-CoL="3M55S" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" box="[810,1154,957,981]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="cruentus">
<emphasis id="B952EA80FF93FF99FCCC48AE46ACFC79" bold="true" box="[810,1154,957,981]" pageId="1">
<emphasis id="B952EA80FF93FF99FCCC48AE41FBFC79" bold="true" box="[810,981,957,981]" italics="true" pageId="1">Hister cruentus</emphasis>
Erichson, 1834
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B993692FF93FF99FCCC48EF468DFBD8" blockId="1.[810,1320,1020,1140]" pageId="1">
There are
<specimenCount id="9D20FD1BFF93FF99FC7448EF463EFBB8" box="[914,1040,1020,1044]" count="4" pageId="1" type="generic">4 specimens</specimenCount>
from the type-series. Thus, a designation of the
<typeStatus id="549D8830FF93FF99FBE54F0F466EFB98" box="[1027,1088,1052,1076]" pageId="1" type="lectotype">lecto-</typeStatus>
and
<typeStatus id="549D8830FF93FF99FB924F0F4721FB98" box="[1140,1295,1052,1076]" pageId="1" type="paralectotype">paralectotypes</typeStatus>
is needed.Additionally, the figures of the male genital structure are given (Figs. 11-15).
</paragraph>
</subSubSection>
<subSubSection id="C33C6519FF93FF9BFCCC4F8E438DFD38" lastPageId="3" pageId="1" type="description">
<paragraph id="8B993692FF93FF99FCCC4F8E4688FA18" blockId="1.[810,1321,1180,1460]" pageId="1">
<typeStatus id="549D8830FF93FF99FCCC4F8E41BBFB19" box="[810,917,1181,1205]" pageId="1" type="lectotype">Lectotype</typeStatus>
: a female, 1) [white, printed] 48809, 2) [green, handwritten] [
<collectingCountry id="F3317602FF93FF99FBEE4FAF4611FB78" box="[1032,1087,1212,1236]" name="South Africa" pageId="1">RSA</collectingCountry>
] Caffr.[aria], Krebs, 3) [green, handwritten]
<taxonomicName id="4C264D11FF93FF99FBE74FCE4650FB58" authority="Er." authorityName="Erichson" authorityYear="1834" box="[1025,1150,1245,1268]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="cruentus">cruentus Er.</taxonomicName>
, 4) [red, printed] Type, 5) [blue, printed] Hist.-Coll. (
<taxonomicName id="4C264D11FF93FF99FB404FEF470EFAB8" authorityName="Linnaeus" authorityYear="1758" box="[1190,1312,1276,1300]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) Nr 48809,
<taxonomicName id="4C264D11FF93FF99FC7D4E0F464FFA98" authority="Er." authorityName="Erichson" authorityYear="1834" box="[923,1121,1308,1332]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="cruentus">Hister cruentus Er.</taxonomicName>
<superScript id="7C539BDAFF93FF99FB874E084647FA85" attach="none" box="[1121,1129,1307,1321]" fontSize="6" pageId="1">*</superScript>
, Caffraria, Krebs, Zool. Mus. Berlin, 6) [red, printed]
<typeStatus id="549D8830FF93FF99FB5B4E2F470DFAF8" box="[1213,1315,1340,1364]" pageId="1" type="syntype">Syntypus</typeStatus>
,
<taxonomicName id="4C264D11FF93FF99FCCC4E4F468EFAD8" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" box="[810,1184,1372,1396]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="cruentus">Hister cruentus Erichson, 1834</taxonomicName>
, labeled by MNHUB, 2007, 7) [white, printed]
<typeStatus id="549D8830FF93FF99FB704E6E4724FA39" box="[1174,1290,1405,1429]" pageId="1" type="lectotype">Lectotypus</typeStatus>
, 8) [white, printed] Des. S.
<bibRefCitation id="EFB74B63FF93FF99FBF94E8E468CFA18" author="Mazur S." box="[1055,1186,1436,1460]" pageId="1" pagination="129 - 140" refId="ref3101" refString="Mazur S. 2007. Notes on some African Histerini (Coleoptera: Histeridae) with description of a new genus Quassarus gen. n. Studies and Reports of District Museum Prague-East Taxonomical Series 3 (1 - 2): 129 - 140." type="book chapter" year="2007">Mazur 2007</bibRefCitation>
.
</paragraph>
<paragraph id="8B993692FF93FF9BFCCC4ECF401AFF78" blockId="1.[810,1320,1500,1652]" lastBlockId="3.[262,766,124,660]" lastPageId="3" pageId="1">
<typeStatus id="549D8830FF93FF99FCCC4ECF41E8FA58" box="[810,966,1500,1524]" pageId="1" type="paralectotype">Paralectotypes</typeStatus>
: I, a male, 1) [green, handwritten] [
<collectingCountry id="F3317602FF93FF99FCD64EEF4149F9B8" box="[816,871,1532,1556]" name="South Africa" pageId="1">RSA</collectingCountry>
, Capland] Pr. b. sp., Tollin, 2) [red, printed] Type, 3) [blue, printed] Hist.-Coll. (
<taxonomicName id="4C264D11FF93FF99FB404D0F470EF998" authorityName="Linnaeus" authorityYear="1758" box="[1190,1312,1564,1588]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) Nr 48809,
<taxonomicName id="4C264D11FF93FF99FC794D2F4642F9F8" authority="Er." authorityName="Erichson" authorityYear="1834" box="[927,1132,1596,1620]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="cruentus">Hister cruentus Er.</taxonomicName>
<superScript id="7C539BDAFF93FF99FB8A4D28465AF9E5" attach="none" box="[1132,1140,1595,1609]" fontSize="6" pageId="1">*</superScript>
, Pr. b. sp., Zool. Mus. Berlin, 4) [red, printed]
<typeStatus id="549D8830FF93FF99FB954D4F46F7F9D8" box="[1139,1241,1628,1652]" pageId="1" type="syntype">Syntypus</typeStatus>
,
<taxonomicName id="4C264D11FF93FF9BFB034D4F403CFF38" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" lastPageId="3" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="cruentus">Hister cruentus Erichson, 1834</taxonomicName>
, labeled by MNHUB, 2007, 5) [white, printed]
<typeStatus id="549D8830FF91FF9BFE184B8F40BAFF18" box="[510,660,156,180]" pageId="3" type="paralectotype">Paralectotypus</typeStatus>
, 6) [white, printed] Des. S.
<bibRefCitation id="EFB74B63FF91FF9BFE4B4BAE401EFF78" author="Mazur S." box="[429,560,188,212]" pageId="3" pagination="129 - 140" refId="ref3101" refString="Mazur S. 2007. Notes on some African Histerini (Coleoptera: Histeridae) with description of a new genus Quassarus gen. n. Studies and Reports of District Museum Prague-East Taxonomical Series 3 (1 - 2): 129 - 140." type="book chapter" year="2007">Mazur 2007</bibRefCitation>
.
</paragraph>
<paragraph id="8B993692FF91FF9BFEE04BCE401AFE18" blockId="3.[262,766,124,660]" pageId="3">
II, sex undetermined, 1) [red, printed] Type, 2) [blue, printed] Hist.-Coll. (
<taxonomicName id="4C264D11FF91FF9BFDF34BEF40A7FEB8" authorityName="Linnaeus" authorityYear="1758" box="[533,649,252,276]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) Nr 48809,
<taxonomicName id="4C264D11FF91FF9BFEE04A0F43E6FE98" authority="Er." authorityName="Erichson" authorityYear="1834" box="[262,456,284,308]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="cruentus">Hister cruentus Er.</taxonomicName>
<superScript id="7C539BDAFF91FF9BFE214A0843E0FE85" attach="none" box="[455,462,283,297]" fontSize="6" pageId="3">*</superScript>
, Caffraria, Krebs, Zool. Mus. Berlin, 3) [red, printed]
<typeStatus id="549D8830FF91FF9BFE1E4A2F4077FEF8" box="[504,601,316,340]" pageId="3" type="syntype">Syntypus</typeStatus>
,
<taxonomicName id="4C264D11FF91FF9BFD844A2F439EFED8" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="cruentus">Hister cruentus Erichson, 1834</taxonomicName>
, labeled by MNHUB 2007, 4) [white, printed]
<typeStatus id="549D8830FF91FF9BFE2D4A6F4056FE38" box="[459,632,380,404]" pageId="3" type="paralectotype">Paralectotypus</typeStatus>
, 5) [white, printed] Des. S.
<bibRefCitation id="EFB74B63FF91FF9BFE4B4A8E401EFE18" author="Mazur S." box="[429,560,412,436]" pageId="3" pagination="129 - 140" refId="ref3101" refString="Mazur S. 2007. Notes on some African Histerini (Coleoptera: Histeridae) with description of a new genus Quassarus gen. n. Studies and Reports of District Museum Prague-East Taxonomical Series 3 (1 - 2): 129 - 140." type="book chapter" year="2007">Mazur 2007</bibRefCitation>
.
</paragraph>
<paragraph id="8B993692FF91FF9BFEE04AAE438DFD38" blockId="3.[262,766,124,660]" pageId="3">
III, sex undetermined, 1) [red, printed] Type, 2) [blue, printed] Hist.-Coll. (
<taxonomicName id="4C264D11FF91FF9BFDF34ACF40A7FE58" authorityName="Linnaeus" authorityYear="1758" box="[533,649,476,500]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) Nr 48809,
<taxonomicName id="4C264D11FF91FF9BFEE04AEF43E6FDB8" authority="Er." authorityName="Erichson" authorityYear="1834" box="[262,456,508,532]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="cruentus">Hister cruentus Er.</taxonomicName>
<superScript id="7C539BDAFF91FF9BFE2E4AE843E1FDA5" attach="none" box="[456,463,507,521]" fontSize="6" pageId="3">*</superScript>
, Pr. b. sp., Zool. Mus. Berlin, 3) [red, printed]
<typeStatus id="549D8830FF91FF9BFE37490F4012FD98" box="[465,572,540,564]" pageId="3" type="syntype">Syntypus</typeStatus>
,
<taxonomicName id="4C264D11FF91FF9BFDAD490F4385FDF8" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="cruentus">Hister cruentus Erichson, 1834</taxonomicName>
, labeled by MNHUB, 4) [white, printed]
<typeStatus id="549D8830FF91FF9BFE86494F43D0FDD8" box="[352,510,604,628]" pageId="3" type="paralectotype">Paralectotypus</typeStatus>
, 5) [white, printed] Des. S.
<bibRefCitation id="EFB74B63FF91FF9BFEF9496E43B1FD38" author="Mazur S." box="[287,415,636,660]" pageId="3" pagination="129 - 140" refId="ref3101" refString="Mazur S. 2007. Notes on some African Histerini (Coleoptera: Histeridae) with description of a new genus Quassarus gen. n. Studies and Reports of District Museum Prague-East Taxonomical Series 3 (1 - 2): 129 - 140." type="book chapter" year="2007">Mazur 2007</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,169 @@
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<mods:title id="C1D9339CEE712BEFFBF83C22B27E34DD">Two new species of the genusAirapus Stebnicka &amp; Howden, 1996 (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
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<heading id="D0CC81D1FFB5FFB6FF8E64C1D4BFFE1A" box="[114,387,466,490]" fontSize="10" level="2" pageId="1" pageNumber="36" reason="7">
<taxonomicName id="4C3B4D3EFFB5FFB6FF8E64C1D413FE19" ID-CoL="C399Z" authorityName="Bellmann &amp; Minkina" authorityYear="2023" box="[114,303,466,490]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="rudolphi" status="sp. nov.">
<emphasis id="B94FEAAFFFB5FFB6FF8E64C1D413FE19" bold="true" box="[114,303,466,490]" italics="true" pageId="1" pageNumber="36">Airapus rudolphi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57D4FFB5FFB6FED264C1D4BFFE1A" box="[302,387,466,489]" pageId="1" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="C3216536FFB5FFB6FF8E671CD42CFDD4" box="[114,272,527,551]" pageId="1" pageNumber="36" type="description">
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<emphasis id="B94FEAAFFFB5FFB6FF8E671CD42CFDD4" bold="true" box="[114,272,527,551]" pageId="1" pageNumber="36">
(
<figureCitation id="13002A38FFB5FFB6FF85671CD5EBFDD4" box="[121,215,527,551]" captionStart="Figs 1-3" captionStartId="1.[114,160,1170,1194]" captionTargetBox="[119,592,586,1124]" captionTargetId="figure-371@1.[114,601,582,1133]" captionTargetPageId="1" captionText="Figs 1-3. Airapus rudolphisp. nov., ♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." pageId="1" pageNumber="36">Figs 1-3</figureCitation>
;
<figureCitation id="13002A38FFB5FFB6FF18671CD434FDD4" box="[228,264,527,551]" captionStart="Figs 7-10" captionStartId="4.[189,235,1137,1161]" captionTargetBox="[189,675,640,1099]" captionTargetId="figure-405@4.[189,675,631,1099]" captionTargetPageId="4" captionText="Figs 7-10. Heads and epipharyngi: 7- A. rudolphisp. nov., ♀, holotype, head; 8- A. yunnanensis sp. nov.,♀, holotype, head; 9- A. rudolphisp. nov., ♀, holotype, epipharynx; 10- A. yunnanensis sp. nov.,♀, holotype, epipharynx. Figs 7, 9: scale lines: 1.0 mm. Figs 8, 10: scale lines: 0.2 mm." pageId="1" pageNumber="36">7-8</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF446635FFB5FFB6FF8E6181D704FB14" pageId="1" pageNumber="36" startId="1.[114,160,1170,1194]" targetBox="[119,592,586,1124]" targetPageId="1" targetType="figure">
<paragraph id="8B8436BDFFB5FFB6FF8E6181D704FB14" blockId="1.[114,593,1170,1256]" pageId="1" pageNumber="36">
Figs 1-3.
<taxonomicName id="4C3B4D3EFFB5FFB6FF0D6180D489FB58" authorityName="Bellmann &amp; Minkina" authorityYear="2023" box="[241,437,1171,1195]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="rudolphi" status="sp. nov.">
<emphasis id="B94FEAAFFFB5FFB6FF0D6180D489FB58" box="[241,437,1171,1195]" italics="true" pageId="1" pageNumber="36">Airapus rudolphi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57D4FFB5FFB6FE496180D722FB59" box="[437,542,1171,1194]" pageId="1" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C3216536FFB5FFB6FF8E601ED496FA2E" pageId="1" pageNumber="36" type="materials_examined">
<paragraph id="8B8436BDFFB5FFB6FF8E601ED482FAB7" blockId="1.[114,593,1293,1348]" pageId="1" pageNumber="36">
<materialsCitation id="3B533CE0FFB5FFB6FF8E601ED486FAB7" country="Indonesia" location="Ketambe" municipality="Kutacane" pageId="1" pageNumber="36" specimenCount="1" stateProvince="Aceh" typeStatus="holotype">
<emphasis id="B94FEAAFFFB5FFB6FF8E601ED432FAD6" bold="true" box="[114,270,1293,1317]" pageId="1" pageNumber="36">
<typeStatus id="5480881FFFB5FFB6FF8E601ED590FAD6" box="[114,172,1293,1317]" pageId="1" pageNumber="36">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32C762DFFB5FFB6FEEB601ED4BEFAD6" box="[279,386,1293,1317]" name="Indonesia" pageId="1" pageNumber="36">Indonesia</collectingCountry>
, N-Sumatra,
<collectingRegion id="49FFF85FFFB5FFB6FDEE601ED5EEFAB7" country="Indonesia" name="Aceh" pageId="1" pageNumber="36">Aceh province</collectingRegion>
,
<collectingMunicipality id="6BE0ACC7FFB5FFB6FF20603FD47DFAB0" box="[220,321,1324,1347]" pageId="1" pageNumber="36">Kutacane</collectingMunicipality>
/
<location id="8EE46066FFB5FFB6FEAA603FD486FAB7" LSID="urn:lsid:plazi:treatment:039287ABFFB5FFB5FF8E64C1D6CAFC5A:8EE46066FFB5FFB6FEAA603FD486FAB7" box="[342,442,1324,1348]" country="Indonesia" municipality="Kutacane" name="Ketambe" pageId="1" pageNumber="36" stateProvince="Aceh">Ketambe</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8436BDFFB5FFB6FF8E607AD496FA2E" blockId="1.[114,593,1385,1501]" pageId="1" pageNumber="36">
<emphasis id="B94FEAAFFFB5FFB6FF8E607AD421FA72" bold="true" box="[114,285,1385,1409]" pageId="1" pageNumber="36">Type material.</emphasis>
<materialsCitation id="3B533CE0FFB5FFB6FEDA607AD496FA2E" collectingDate="2013-12-02" collectingDateMax="2013-12-07" collectingDateMin="2013-12-02" collectorName="Horst Rudolph" country="Indonesia" location="Ketambe" municipality="Kutacane" pageId="1" pageNumber="36" specimenCount="1" specimenCount-female="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<typeStatus id="5480881FFFB5FFB6FEDA607AD4B6FA72" box="[294,394,1385,1409]" pageId="1" pageNumber="36" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D3DFD34FFB5FFB6FE606079D48CFA72" box="[412,432,1386,1409]" count="1" pageId="1" pageNumber="36" type="female"></specimenCount>
): “
<collectingCountry id="F32C762DFFB5FFB6FE2F607AD76DFA72" box="[467,593,1385,1409]" name="Indonesia" pageId="1" pageNumber="36">Indonesien</collectingCountry>
N.
<collectingRegion id="49FFF85FFFB5FFB6FF74609BD5D8FA53" box="[136,228,1416,1440]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="36">Sumatra</collectingRegion>
| Aceh-Prov. Umg. von |
<collectingMunicipality id="6BE0ACC7FFB5FFB6FF8E60B4D5EBFA4D" box="[114,215,1447,1470]" pageId="1" pageNumber="36">Kutacane</collectingMunicipality>
|
<location id="8EE46066FFB5FFB6FF0060B5D460FA4D" LSID="urn:lsid:plazi:treatment:039287ABFFB5FFB5FF8E64C1D6CAFC5A:8EE46066FFB5FFB6FF0060B5D460FA4D" box="[252,348,1446,1470]" country="Indonesia" municipality="Kutacane" name="Ketambe" pageId="1" pageNumber="36" stateProvince="Sumatera Utara">Ketambe</location>
|
<date id="FF85107DFFB5FFB6FE7D60B4D594FA2E" pageId="1" pageNumber="36" value="2013-12-02" valueMax="2013-12-07" valueMin="2013-12-02">
<collectingDate id="EFC1E995FFB5FFB6FE7D60B4D594FA2E" pageId="1" pageNumber="36" value="2013-12-02" valueMax="2013-12-07" valueMin="2013-12-02">02.-07. Dezember 2013</collectingDate>
</date>
| leg.
<collectorName id="26CE536BFFB5FFB6FF1460D6D4BAFA2E" box="[232,390,1477,1501]" pageId="1" pageNumber="36">Horst Rudolph</collectorName>
||”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3216536FFB5FFB5FF8E6311D1BFFDAB" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36" type="description">
<paragraph id="8B8436BDFFB5FFB6FF8E6311D433F985" blockId="1.[114,593,1538,1654]" pageId="1" pageNumber="36">
<emphasis id="B94FEAAFFFB5FFB6FF8E6311D43FF9E9" bold="true" box="[114,259,1538,1562]" pageId="1" pageNumber="36">Description.</emphasis>
Dorsum (
<figureCitation id="13002A38FFB5FFB6FE906311D491F9E9" box="[364,429,1538,1562]" captionStart="Figs 1-3" captionStartId="1.[114,160,1170,1194]" captionTargetBox="[119,592,586,1124]" captionTargetId="figure-371@1.[114,601,582,1133]" captionTargetPageId="1" captionText="Figs 1-3. Airapus rudolphisp. nov., ♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." pageId="1" pageNumber="36">Fig. 1</figureCitation>
). Body length
<quantity id="4CC39B58FFB5FFB6FF8E6332D5F2F9CA" box="[114,206,1569,1593]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="36" unit="mm" value="3.6">3.6 mm</quantity>
, elongate, shiny, light brownish, sides and apex of elytra with indistinct macrosetation.
</paragraph>
<paragraph id="8B8436BDFFB5FFB6FF8E63B5D686FCDC" blockId="1.[114,141,1702,1726]" lastBlockId="1.[624,1104,178,815]" pageId="1" pageNumber="36">
36 Head (
<figureCitation id="13002A38FFB5FFB6FD4765A1D63CFF39" box="[699,768,178,202]" captionStart="Figs 7-10" captionStartId="4.[189,235,1137,1161]" captionTargetBox="[189,675,640,1099]" captionTargetId="figure-405@4.[189,675,631,1099]" captionTargetPageId="4" captionText="Figs 7-10. Heads and epipharyngi: 7- A. rudolphisp. nov., ♀, holotype, head; 8- A. yunnanensis sp. nov.,♀, holotype, head; 9- A. rudolphisp. nov., ♀, holotype, epipharynx; 10- A. yunnanensis sp. nov.,♀, holotype, epipharynx. Figs 7, 9: scale lines: 1.0 mm. Figs 8, 10: scale lines: 0.2 mm." pageId="1" pageNumber="36">Fig. 7</figureCitation>
) relatively large, trapezoidal, convex, shiny, without microreticulation. Clypeus gently bordered, widely triangularly sinuate anteriorly, denticulate on sides, widely rounded laterally, not notched before genae, clypeal border without macrosetae. Genae obtuse, very distinctly exceeding eyes, with few very short, thin macrosetae. Frontal suture noticeable on sides, without gibbosities. Punctation simple: basally very dense, quite coarse, with variable in size, punctures rounded; in the middle very dense, coarse, with somewhat variable in size, longitudinal punctures; anteriorly with moderately dense, fine, regular in size, somewhat elevated punctures; anterior border of clypeus (with fine punctures) divided from the middle part by quite convex transverse tubercle, which is somewhat lower in the middle.
</paragraph>
<paragraph id="8B8436BDFFB5FFB6FD8C6647D63CFB4E" blockId="1.[624,1103,852,1213]" pageId="1" pageNumber="36">
Epipharynx (
<figureCitation id="13002A38FFB5FFB6FCFC6647D679FC9F" box="[768,837,852,876]" captionStart="Figs 7-10" captionStartId="4.[189,235,1137,1161]" captionTargetBox="[189,675,640,1099]" captionTargetId="figure-405@4.[189,675,631,1099]" captionTargetPageId="4" captionText="Figs 7-10. Heads and epipharyngi: 7- A. rudolphisp. nov., ♀, holotype, head; 8- A. yunnanensis sp. nov.,♀, holotype, head; 9- A. rudolphisp. nov., ♀, holotype, epipharynx; 10- A. yunnanensis sp. nov.,♀, holotype, epipharynx. Figs 7, 9: scale lines: 1.0 mm. Figs 8, 10: scale lines: 0.2 mm." pageId="1" pageNumber="36">Fig. 8</figureCitation>
) transverse, with sides broadly rounded, anterior margin of pedia concavely arcuate, corypha above of margin of pedia, distinct, with two very short, thick celtes. Mesoepitorma shortened basally, somewhat pear-shaped. Chaetopariae with dense belt of thick, long chaetae; adelochaetae similar to chaetopariae, but thicker and shorter; chaetopediae with only one very short and thin chaeta on each side; angustofenestrae dense, short and thick. Tormae long.
</paragraph>
<paragraph id="8B8436BDFFB5FFB5FD8C61F0D420FF1B" blockId="1.[624,1103,1251,1643]" lastBlockId="2.[189,668,178,232]" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36">Pronotum transverse, as wide as base of elytra, widest in the middle, convex, shiny, without microreticulation, with double punctation: larger punctures moderately coarse, dense, regularly distributes, somewhat irregular in size, smaller punctures not fine, not so dense, irregularly distributed. Base quite distinctly bordered by a thin and shallow groove, sides bordered quite widely and somewhat more deeply than base; anteriorly not bordered. Sides and base with moderately long, quite thick macrosetae. Anterior angles distinctly rounded. Base before hind angles widely rounded.</paragraph>
<paragraph id="8B8436BDFFB6FFB5FF41641ED43DFE71" blockId="2.[189,668,269,386]" pageId="2" pageNumber="37">Scutellum small, triangular, with widely rounded sides, with a few small punctures, moderately shiny, without microreticulation.</paragraph>
<paragraph id="8B8436BDFFB6FFB5FF4164B4D49CFC79" blockId="2.[189,668,423,907]" pageId="2" pageNumber="37">Elytra elongate, convex, parallel, shiny, without microreticulation, with short and thin, barely noticeable macrosetae on sides and before apex; with small but distinct humeral denticles; with ten striae and ten intervals. Intervals shiny, convex, with row of not fine punctures on external half and with row of moderately coarse punctures on internal half, which are connected with punctures of rows; due to this the striae seem to indent the margins of the intervals margins very distinctly. Striae distinctly, quite densely punctate, with medium sized punctures; punctures distinctly indenting margins of intervals. All striae joined together before apex.</paragraph>
<paragraph id="8B8436BDFFB6FFB5FF4166A3D7A9FC3B" blockId="2.[189,661,944,968]" box="[189,661,944,968]" pageId="2" pageNumber="37">Pygidium with similar structure to ventrites.</paragraph>
<paragraph id="8B8436BDFFB6FFB5FF4166FDD42DF9BF" blockId="2.[189,669,1005,1612]" pageId="2" pageNumber="37">Legs. Femora shiny, without microreticulation, not coarsely and sparsely punctate; profemora basally and apically bordered, meso- and metafemora bordered only apically. Mesofemora Widely separated from each other. Protibiae distinctly tridentate laterally, proximally not serrulate; somewhat inwardly sinuate; dorsal side smooth, shiny, with few very fine punctures; apical spur long, moderately broad, gently downwardly and outwardly bent, with apex acute. Meso- and metatibiae fimbriate apically with row of short spinules of equal length. Apex of metatibiae without accessory spine. Metatibiae superior apical spur equal length as basimetatarsomere, latter approximately as long as following three metatarsomeres combined. Claws short, very thin, distinctly arcuate.</paragraph>
<paragraph id="8B8436BDFFB6FFB5FD4765A1D66BFF3A" blockId="2.[699,855,178,201]" box="[699,855,178,201]" pageId="2" pageNumber="37">Macropterous.</paragraph>
<paragraph id="8B8436BDFFB6FFB5FD4765FCD1BFFDAB" blockId="2.[699,1179,239,600]" pageId="2" pageNumber="37">
Venter (
<figureCitation id="13002A38FFB6FFB5FCE265FCD650FEF4" box="[798,876,239,263]" captionStart="Figs 1-3" captionStartId="1.[114,160,1170,1194]" captionTargetBox="[119,592,586,1124]" captionTargetId="figure-371@1.[114,601,582,1133]" captionTargetPageId="1" captionText="Figs 1-3. Airapus rudolphisp. nov., ♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." pageId="2" pageNumber="37">Fig. 2</figureCitation>
). Meso-metaventral plate shiny, weakly convex, with distinct, quite deep, quite narrow longitudinal furrow in the middle; surface with moderately coarse, somewhat variable in size, sparse punctures. Process of meso-metaventral plate without any groove or recess, below of base there is no additional modifications. Abdominal ventrites shiny, anteriorly weakly fluted, without microreticulation, with moderately dense, moderately coarsely, quite regularly spaced punctures.
</paragraph>
</subSubSection>
<subSubSection id="C3216536FFB6FFB5FD47678FD6CAFC5A" pageId="2" pageNumber="37" type="etymology">
<paragraph id="8B8436BDFFB6FFB5FD47678FD6F0FD01" blockId="2.[699,1179,668,754]" pageId="2" pageNumber="37">
<emphasis id="B94FEAAFFFB6FFB5FD47678FD678FD47" bold="true" box="[699,836,668,692]" pageId="2" pageNumber="37">Etymology.</emphasis>
Patronymic; an adjective derived from the name of
<typeStatus id="5480881FFFB6FFB5FBCA67A8D1A6FD20" box="[1078,1178,699,723]" pageId="2" pageNumber="37" type="holotype">holotype</typeStatus>
collector: Horst Rudolph.
</paragraph>
<paragraph id="8B8436BDFFB6FFB5FD476604D68AFCDC" blockId="2.[699,950,791,815]" box="[699,950,791,815]" pageId="2" pageNumber="37">
<emphasis id="B94FEAAFFFB6FFB5FD476604D603FCDC" bold="true" box="[699,831,791,815]" pageId="2" pageNumber="37">Variability.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B8436BDFFB6FFB5FD476647D12BFC9F" blockId="2.[699,1047,852,876]" box="[699,1047,852,876]" pageId="2" pageNumber="37">
<emphasis id="B94FEAAFFFB6FFB5FD476647D69CFC9F" bold="true" box="[699,928,852,876]" pageId="2" pageNumber="37">Sexual dimorphism.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B8436BDFFB6FFB5FD476682D6CAFC5A" blockId="2.[699,1014,913,938]" box="[699,1014,913,938]" pageId="2" pageNumber="37">
<emphasis id="B94FEAAFFFB6FFB5FD476682D61EFC5A" bold="true" box="[699,802,913,937]" pageId="2" pageNumber="37">Affinity.</emphasis>
See discussion part.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,206 @@
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<mods:title id="3704AAA7678D801EC48B20B6046E4648">Two new species of the genusAirapus Stebnicka &amp; Howden, 1996 (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
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<subSubSection id="C3216536FFB6FFB5FD4766FDD13CFBF6" box="[699,1024,1006,1030]" pageId="2" pageNumber="37" type="nomenclature">
<paragraph id="8B8436BDFFB6FFB5FD4766FDD13CFBF6" blockId="2.[699,1024,1006,1030]" box="[699,1024,1006,1030]" pageId="2" pageNumber="37">
<heading id="D0CC81D1FFB6FFB5FD4766FDD13CFBF6" box="[699,1024,1006,1030]" fontSize="10" level="2" pageId="2" pageNumber="37" reason="7">
<taxonomicName id="4C3B4D3EFFB6FFB5FD4766FDD699FBF5" ID-CoL="C399N" authorityName="Bellmann &amp; Minkina" authorityYear="2023" box="[699,933,1006,1030]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="yunnanensis" status="sp. nov.">
<emphasis id="B94FEAAFFFB6FFB5FD4766FDD699FBF5" bold="true" box="[699,933,1006,1030]" italics="true" pageId="2" pageNumber="37">Airapus yunnanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57D4FFB6FFB5FC5766FDD13CFBF6" box="[939,1024,1006,1029]" pageId="2" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3216536FFB6FFB5FD476138D65BFBB0" box="[699,871,1067,1091]" pageId="2" pageNumber="37" type="description">
<paragraph id="8B8436BDFFB6FFB5FD476138D65BFBB0" blockId="2.[699,871,1067,1091]" box="[699,871,1067,1091]" pageId="2" pageNumber="37">
<emphasis id="B94FEAAFFFB6FFB5FD476138D65BFBB0" bold="true" box="[699,871,1067,1091]" pageId="2" pageNumber="37">
(
<figureCitation id="13002A38FFB6FFB5FD3E6138D61DFBB0" box="[706,801,1067,1091]" captionStart="Figs 4-6" captionStartId="3.[114,160,1218,1242]" captionTargetBox="[119,595,693,1173]" captionTargetId="figure-410@3.[114,599,692,1181]" captionTargetPageId="3" captionText="Figs 4-6. Airapus yunnanensissp. nov., ♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." pageId="2" pageNumber="37">Figs 4-6</figureCitation>
;
<figureCitation id="13002A38FFB6FFB5FCD16138D663FBB0" box="[813,863,1067,1091]" captionStart="Figs 7-10" captionStartId="4.[189,235,1137,1161]" captionTargetBox="[189,675,640,1099]" captionTargetId="figure-405@4.[189,675,631,1099]" captionTargetPageId="4" captionText="Figs 7-10. Heads and epipharyngi: 7- A. rudolphisp. nov., ♀, holotype, head; 8- A. yunnanensis sp. nov.,♀, holotype, head; 9- A. rudolphisp. nov., ♀, holotype, epipharynx; 10- A. yunnanensis sp. nov.,♀, holotype, epipharynx. Figs 7, 9: scale lines: 1.0 mm. Figs 8, 10: scale lines: 0.2 mm." pageId="2" pageNumber="37">9-10</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3216536FFB6FFB5FD47617BD110FACB" pageId="2" pageNumber="37" type="materials_examined">
<paragraph id="8B8436BDFFB6FFB5FD47617BD61DFB6C" blockId="2.[699,1178,1128,1183]" pageId="2" pageNumber="37">
<materialsCitation id="3B533CE0FFB6FFB5FD47617BD621FB6C" country="China" location="Gulingin" municipality="Honghe" pageId="2" pageNumber="37" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B94FEAAFFFB6FFB5FD47617BD65DFB73" bold="true" box="[699,865,1128,1152]" pageId="2" pageNumber="37">
<typeStatus id="5480881FFFB6FFB5FD47617BD7C9FB73" box="[699,757,1128,1152]" pageId="2" pageNumber="37">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32C762DFFB6FFB5FC8F617BD68BFB73" box="[883,951,1128,1152]" name="China" pageId="2" pageNumber="37">China</collectingCountry>
:
<collectingRegion id="49FFF85FFFB6FFB5FC30617AD118FB73" box="[972,1060,1129,1152]" country="China" name="Yunnan" pageId="2" pageNumber="37">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BE0ACC7FFB6FFB5FBC5617BD1AAFB73" box="[1081,1174,1128,1152]" pageId="2" pageNumber="37">Honghe</collectingMunicipality>
,
<location id="8EE46066FFB6FFB5FD476194D621FB6C" LSID="urn:lsid:plazi:treatment:039287ABFFB6FFB3FD4766FDD6C9FD47:8EE46066FFB6FFB5FD476194D621FB6C" box="[699,797,1159,1183]" country="China" municipality="Honghe" name="Gulingin" pageId="2" pageNumber="37" stateProvince="Yunnan">Gulingin</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8436BDFFB6FFB5FD4761D7D110FACB" blockId="2.[699,1178,1220,1336]" pageId="2" pageNumber="37">
<emphasis id="B94FEAAFFFB6FFB5FD4761D7D651FB2F" bold="true" box="[699,877,1220,1244]" pageId="2" pageNumber="37">Type material.</emphasis>
<materialsCitation id="3B533CE0FFB6FFB5FC8161D7D110FACB" collectingDate="2019-06-11" collectorName="A. Weigel" country="China" elevation="520" latitude="22.71861" location="Gulingin" longLatPrecision="21" longitude="103.998886" municipality="Honghe" pageId="2" pageNumber="37" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="5480881FFFB6FFB5FC8161D7D6DCFB2F" box="[893,992,1220,1244]" pageId="2" pageNumber="37" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D3DFD34FFB6FFB5FC0561D6D131FB2F" box="[1017,1037,1221,1244]" count="1" pageId="2" pageNumber="37" type="female"></specimenCount>
): “
<collectingCountry id="F32C762DFFB6FFB5FBCA61D7D1B3FB2F" box="[1078,1167,1220,1244]" name="China" pageId="2" pageNumber="37">CHINA</collectingCountry>
,
<collectingRegion id="49FFF85FFFB6FFB5FD4761F0D628FB09" box="[699,788,1251,1274]" country="China" name="Yunnan" pageId="2" pageNumber="37">Yunnan</collectingRegion>
:
<collectingMunicipality id="6BE0ACC7FFB6FFB5FCDE61F0D64AFB08" box="[802,886,1251,1275]" pageId="2" pageNumber="37">Honghe</collectingMunicipality>
|
<location id="8EE46066FFB6FFB5FC6D61F0D6CFFB08" LSID="urn:lsid:plazi:treatment:039287ABFFB6FFB3FD4766FDD6C9FD47:8EE46066FFB6FFB5FC6D61F0D6CFFB08" box="[913,1011,1251,1275]" country="China" latitude="22.71861" longLatPrecision="21" longitude="103.998886" municipality="Honghe" name="Gulingin" pageId="2" pageNumber="37" stateProvince="Yunnan">Gulingin</location>
,
<quantity id="4CC39B58FFB6FFB5FBFD61F0D17DFB08" box="[1025,1089,1251,1275]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="37" unit="m" value="520.0">
<elevation id="0016D18EFFB6FFB5FBFD61F0D17DFB08" box="[1025,1089,1251,1275]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="37" unit="m" value="520.0">520m</elevation>
</quantity>
, LFF, |
<date id="FF85107DFFB6FFB5FD476012D60BFAEA" box="[699,823,1281,1305]" pageId="2" pageNumber="37" value="2019-06-11">
<collectingDate id="EFC1E995FFB6FFB5FD476012D60BFAEA" box="[699,823,1281,1305]" pageId="2" pageNumber="37" value="2019-06-11">11.VI.2019</collectingDate>
</date>
,
<geoCoordinate id="EE0F507AFFB6FFB5FCAC6012D6F0FAEA" box="[848,972,1281,1305]" degrees="22" direction="north" minutes="43" orientation="latitude" pageId="2" pageNumber="37" precision="15" seconds="7" value="22.71861">22°437N</geoCoordinate>
|
<geoCoordinate id="EE0F507AFFB6FFB5FC066012D1A9FAEA" box="[1018,1173,1281,1305]" degrees="103" direction="east" minutes="59" orientation="longitude" pageId="2" pageNumber="37" precision="15" seconds="56" value="103.998886">103°5956E</geoCoordinate>
, primary | forest, leg.
<collectorName id="26CE536BFFB6FFB5FC676033D13BFACB" box="[923,1031,1312,1336]" pageId="2" pageNumber="37">A. Weigel</collectorName>
||”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3216536FFB6FFB3FD47604ED11FFE91" lastPageId="4" lastPageNumber="39" pageId="2" pageNumber="37" type="description">
<paragraph id="8B8436BDFFB6FFB5FD47604ED1AEFA40" blockId="2.[699,1178,1373,1459]" pageId="2" pageNumber="37">
<emphasis id="B94FEAAFFFB6FFB5FD47604ED670FA86" bold="true" box="[699,844,1373,1397]" pageId="2" pageNumber="37">Description.</emphasis>
Dorsum (
<figureCitation id="13002A38FFB6FFB5FC46604ED6C2FA86" box="[954,1022,1373,1397]" captionStart="Figs 4-6" captionStartId="3.[114,160,1218,1242]" captionTargetBox="[119,595,693,1173]" captionTargetId="figure-410@3.[114,599,692,1181]" captionTargetPageId="3" captionText="Figs 4-6. Airapus yunnanensissp. nov., ♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." pageId="2" pageNumber="37">Fig. 4</figureCitation>
).Body length
<quantity id="4CC39B58FFB6FFB5FD47606FD62CFA67" box="[699,784,1404,1428]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="37" unit="mm" value="3.9">3.9 mm</quantity>
, elongate, shiny, blackish, sides and apex of elytra with indistinct macrosetation.
</paragraph>
<paragraph id="8B8436BDFFB6FFB5FD4760CBD1A7F999" blockId="2.[699,1179,1496,1643]" pageId="2" pageNumber="37">
Head (
<figureCitation id="13002A38FFB6FFB5FCFB60CBD670FA03" box="[775,844,1496,1520]" captionStart="Figs 7-10" captionStartId="4.[189,235,1137,1161]" captionTargetBox="[189,675,640,1099]" captionTargetId="figure-405@4.[189,675,631,1099]" captionTargetPageId="4" captionText="Figs 7-10. Heads and epipharyngi: 7- A. rudolphisp. nov., ♀, holotype, head; 8- A. yunnanensis sp. nov.,♀, holotype, head; 9- A. rudolphisp. nov., ♀, holotype, epipharynx; 10- A. yunnanensis sp. nov.,♀, holotype, epipharynx. Figs 7, 9: scale lines: 1.0 mm. Figs 8, 10: scale lines: 0.2 mm." pageId="2" pageNumber="37">Fig. 9</figureCitation>
) relatively large, trapezoidal, convex, shiny, without microreticulation. Clypeus gently bordered, widely, triangularly sinuate anteriorly, denticulate on sides, widely rounded laterally, not
</paragraph>
<paragraph id="8B8436BDFFB6FFB4FB8563B5D480FD66" blockId="2.[1145,1172,1702,1726]" lastBlockId="3.[114,593,178,661]" lastPageId="3" lastPageNumber="38" pageId="2" pageNumber="37">37 notched before genae, clypeal border without macrosetae. Genae obtuse, very distinctly exceeding eyes, with few very short, thin macrosetae. Frontal suture noticeable on sides, without gibbosities. Punctation simple: basally very dense, quite coarse, with variable in size, rounded punctures; in the middle very dense, coarse, with somewhat variable in size, longitudinal punctures; anteriorly moderately dense, fine, regular in size, with somewhat elevated punctures; anterior border of clypeus (with fine punctures) divided from the middle part by distinctly convex transverse tubercle, which is somewhat lower in the middle.</paragraph>
<paragraph id="8B8436BDFFB7FFB4FD8C65A1D63CFF1B" blockId="3.[624,1103,178,692]" pageId="3" pageNumber="38">angustofenestrae dense, short and thick. Tormae long.</paragraph>
<paragraph id="8B8436BDFFB7FFB4FD8C65FCD7F3FD47" blockId="3.[624,1103,178,692]" pageId="3" pageNumber="38">Pronotum transverse, somewhat wider than base of elytra, widest in the middle, convex, shiny, without microreticulation, with double punctation: larger punctures moderately coarse, dense, regularly distributes, somewhat irregular in size, smaller punctures fine, dense, regularly distributed. Base indistinctly bordered by a thin and shallow groove, sides bordered quite widely and somewhat more deeply than base; anteriorly not bordered. Sides and base with moderately long, quite thick macrosetae. Anterior angles quite distinctly rounded. Base before hind angles widely rounded.</paragraph>
<paragraph id="8B8436BDFFB7FFB4FD8C67C9D6F2FCDC" blockId="3.[624,1103,730,815]" pageId="3" pageNumber="38">Scutellum small, triangular, with ogival sides, with few small punctures, moderately shiny, without microreticulation.</paragraph>
<caption id="DF446635FFB7FFB4FF8E61D1D704FAE4" pageId="3" pageNumber="38" startId="3.[114,160,1218,1242]" targetBox="[119,595,693,1173]" targetPageId="3" targetType="figure">
<paragraph id="8B8436BDFFB7FFB4FF8E61D1D704FAE4" blockId="3.[114,593,1218,1304]" pageId="3" pageNumber="38">
Figs 4-6.
<taxonomicName id="4C3B4D3EFFB7FFB4FF1D61D0D4F9FB28" authorityName="Bellmann &amp; Minkina" authorityYear="2023" box="[225,453,1219,1243]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="yunnanensis" status="sp. nov.">
<emphasis id="B94FEAAFFFB7FFB4FF1D61D0D4F9FB28" box="[225,453,1219,1243]" italics="true" pageId="3" pageNumber="38">Airapus yunnanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57D4FFB7FFB4FE3861D0D71AFB29" box="[452,550,1219,1242]" pageId="3" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B8436BDFFB7FFB4FF8E602ED76DF99A" blockId="3.[114,593,1341,1641]" pageId="3" pageNumber="38">
Epipharynx (
<figureCitation id="13002A38FFB7FFB4FF03602ED473FAA6" box="[255,335,1341,1365]" captionStart="Figs 7-10" captionStartId="4.[189,235,1137,1161]" captionTargetBox="[189,675,640,1099]" captionTargetId="figure-405@4.[189,675,631,1099]" captionTargetPageId="4" captionText="Figs 7-10. Heads and epipharyngi: 7- A. rudolphisp. nov., ♀, holotype, head; 8- A. yunnanensis sp. nov.,♀, holotype, head; 9- A. rudolphisp. nov., ♀, holotype, epipharynx; 10- A. yunnanensis sp. nov.,♀, holotype, epipharynx. Figs 7, 9: scale lines: 1.0 mm. Figs 8, 10: scale lines: 0.2 mm." pageId="3" pageNumber="38">Fig. 10</figureCitation>
) transverse, with sides broadly rounded, anterior margin of pedia concavely arcuate, corypha above margin of pedia, distinct, with two very short, thick celtes. Mesoepitorma shortened basally, somewhat pear-shaped. Chaetopariae with dense belt of thick, long chaetae; adelochaetae similar to chaetopariae, but thicker and shorter; chaetopediae with only one very short and thin chaeta on each side;
</paragraph>
<paragraph id="8B8436BDFFB7FFB4FF8E63B5D796FACB" blockId="3.[114,141,1702,1726]" lastBlockId="3.[624,1103,852,1336]" pageId="3" pageNumber="38">38 Elytra elongate, convex, parallel, shiny, without microreticulation, with short and thin, barely noticeable macrosetae on sides and before apex; with small but distinct humeral denticles; with ten striae and ten intervals. Intervals shiny, convex, with row of not fine punctures on external half and with row of moderately coarse punctures on internal half, which are connected with punctures of rows; due to this thestriae seem to indent intervals margins here very distinctly. Striae distinctly, quite densely punctate, with medium sized punctures; punctures distinctly indenting margins of intervals. All striae joined together before apex.</paragraph>
<paragraph id="8B8436BDFFB7FFB4FD8C604ED174FA86" blockId="3.[624,1096,1373,1397]" box="[624,1096,1373,1397]" pageId="3" pageNumber="38">Pygidium with similar structure to ventrites.</paragraph>
<paragraph id="8B8436BDFFB7FFB3FD8C6088D4D5FDAB" blockId="3.[624,1103,1435,1643]" lastBlockId="4.[189,669,178,601]" lastPageId="4" lastPageNumber="39" pageId="3" pageNumber="38">Legs. Femora shiny, without microreticulation, moderately coarsely and quite densely punctate; profemora basally and apically bordered, meso- and metafemora bordered only apically. Mesofemora are in moderately large distance between each other. Protibiae distinctly tridentate laterally, proximally not serrulate; somewhat inwardly sinuate; dorsal side smooth, shiny, with few very fine punctures; apical spur long, moderately broad, gently downwardly and outwardly bent, with apex acute. Meso- and metatibiae fimbriate apically with row of short spinules of equal length. Apex of metatibiae without accessory spine. Metatibiae superior apical spur slightly shorter than basimetatarsomere, latter approximately as long as following three and half metatarsomeres combined. Claws short, very thin, distinctly arcuate.</paragraph>
<caption id="DF446635FFB0FFB3FF416162D755FAB3" pageId="4" pageNumber="39" startId="4.[189,235,1137,1161]" targetBox="[189,675,640,1099]" targetPageId="4" targetType="figure">
<paragraph id="8B8436BDFFB0FFB3FF416162D755FAB3" blockId="4.[189,668,1137,1345]" pageId="4" pageNumber="39">
Figs 7-10. Heads and epipharyngi: 7-
<taxonomicName id="4C3B4D3EFFB0FFB3FD836162D425FB5B" authorityName="Bellmann &amp; Minkina" authorityYear="2023" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="rudolphi" status="sp. nov.">
<emphasis id="B94FEAAFFFB0FFB3FD836162D425FB5B" italics="true" pageId="4" pageNumber="39">A. rudolphi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57D4FFB0FFB3FEE56183D446FB54" box="[281,378,1168,1191]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, holotype, head; 8-
<taxonomicName id="4C3B4D3EFFB0FFB3FD836183D47DFB34" authorityName="Bellmann &amp; Minkina" authorityYear="2023" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="yunnanensis" status="sp. nov.">
<emphasis id="B94FEAAFFFB0FFB3FD836183D47DFB34" italics="true" pageId="4" pageNumber="39">A. yunnanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57D4FFB0FFB3FEB561BCD49BFB35" box="[329,423,1199,1222]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
,♀, holotype, head; 9-
<taxonomicName id="4C3B4D3EFFB0FFB3FF4161DED469FB16" authorityName="Bellmann &amp; Minkina" authorityYear="2023" box="[189,341,1229,1253]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="rudolphi" status="sp. nov.">
<emphasis id="B94FEAAFFFB0FFB3FF4161DED469FB16" box="[189,341,1229,1253]" italics="true" pageId="4" pageNumber="39">A. rudolphi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57D4FFB0FFB3FEA961DED4F3FB17" box="[341,463,1229,1252]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, holotype, epipharynx; 10-
<taxonomicName id="4C3B4D3EFFB0FFB3FE9361FFD72DFAF7" authorityName="Bellmann &amp; Minkina" authorityYear="2023" box="[367,529,1260,1284]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="yunnanensis" status="sp. nov.">
<emphasis id="B94FEAAFFFB0FFB3FE9361FFD72DFAF7" box="[367,529,1260,1284]" italics="true" pageId="4" pageNumber="39">A. yunnanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57D4FFB0FFB3FDE461FFD74AFAF0" box="[536,630,1260,1283]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
,♀, holotype, epipharynx. Figs 7, 9: scale lines: 1.0 mm. Figs 8, 10: scale lines: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8436BDFFB0FFB3FF416074D465FA8D" blockId="4.[189,345,1383,1406]" box="[189,345,1383,1406]" pageId="4" pageNumber="39">Macropterous.</paragraph>
<paragraph id="8B8436BDFFB0FFB3FF4160B7D11FFE91" blockId="4.[189,669,1443,1652]" lastBlockId="4.[699,1178,178,355]" pageId="4" pageNumber="39">
Venter (
<figureCitation id="13002A38FFB0FFB3FEDC60B7D451FA48" box="[288,365,1443,1468]" captionStart="Figs 4-6" captionStartId="3.[114,160,1218,1242]" captionTargetBox="[119,595,693,1173]" captionTargetId="figure-410@3.[114,599,692,1181]" captionTargetPageId="3" captionText="Figs 4-6. Airapus yunnanensissp. nov., ♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." pageId="4" pageNumber="39">Fig. 5</figureCitation>
). Meso-metaventral plate shiny, weakly convex, with distinct, quite deep, quite narrow longitudinal furrowin the middle; surface with moderately coarse, somewhat variable in size, moderately dense punctures. Process of meso-metaventral plate without any groove or recess, below of base there is no additional modifications. Abdominal ventrites shiny, anteriorly weakly fluted, with out micro reticulation, with moderately dense, moderately coarsely, quite regularly spaced punctures.
</paragraph>
</subSubSection>
<subSubSection id="C3216536FFB0FFB3FD47649BD6C9FD47" pageId="4" pageNumber="39" type="etymology">
<paragraph id="8B8436BDFFB0FFB3FD47649BD61FFE0F" blockId="4.[699,1179,392,508]" pageId="4" pageNumber="39">
<emphasis id="B94FEAAFFFB0FFB3FD47649BD678FE53" bold="true" box="[699,836,392,416]" pageId="4" pageNumber="39">Etymology.</emphasis>
Toponymic; an adjective derived from the name of
<collectingRegion id="49FFF85FFFB0FFB3FC2864B4D1AAFE4C" box="[980,1174,423,447]" country="China" name="Yunnan" pageId="4" pageNumber="39">Yunnan province</collectingRegion>
, where
<typeStatus id="5480881FFFB0FFB3FCF064D6D655FE2E" box="[780,873,453,477]" pageId="4" pageNumber="39" type="holotype">holotype</typeStatus>
of the new species was collected.
</paragraph>
<paragraph id="8B8436BDFFB0FFB3FD476732D68AFDC9" blockId="4.[699,950,545,570]" box="[699,950,545,570]" pageId="4" pageNumber="39">
<emphasis id="B94FEAAFFFB0FFB3FD476732D603FDCA" bold="true" box="[699,831,545,569]" pageId="4" pageNumber="39">Variability.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B8436BDFFB0FFB3FD47674CD12BFD84" blockId="4.[699,1047,607,631]" box="[699,1047,607,631]" pageId="4" pageNumber="39">
<emphasis id="B94FEAAFFFB0FFB3FD47674CD69CFD84" bold="true" box="[699,928,607,631]" pageId="4" pageNumber="39">Sexual dimorphism.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B8436BDFFB0FFB3FD47678FD6C9FD47" blockId="4.[699,1013,668,692]" box="[699,1013,668,692]" pageId="4" pageNumber="39">
<emphasis id="B94FEAAFFFB0FFB3FD47678FD61EFD47" bold="true" box="[699,802,668,692]" pageId="4" pageNumber="39">Affinity.</emphasis>
See discussion part.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,235 @@
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<mods:title id="64772D5974CB51E795DE96F520C09E7F">Two new fossil Silinae (Coleoptera, Cantharidae) from Baltic amber</mods:title>
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<mods:namePart id="A80D8FC22CE99A4558CC69BBB98BC519">Bukejs, Andris</mods:namePart>
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<mods:namePart id="14C9B9D38EE4AB72F4453C9B387F75B2">Fanti, Fabrizio</mods:namePart>
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<paragraph id="8B8036B9B169FF8EFDE8FF3A1FC4FF05" blockId="1.[624,907,177,232]" box="[624,907,177,202]" pageId="1" pageNumber="332">
<heading id="D0C881D5B169FF8EFDE8FF3A1FC4FF05" bold="true" box="[624,907,177,202]" fontSize="10" level="2" pageId="1" pageNumber="332" reason="4">
<emphasis id="B94BEAABB169FF8EFDE8FF3A1FC4FF05" bold="true" box="[624,907,177,202]" pageId="1" pageNumber="332">
<taxonomicName id="4C3F4D3AB169FF8EFDE8FF3A1F7FFF06" authorityName="Bukejs &amp; Fanti" authorityYear="2023" box="[624,816,178,202]" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
<emphasis id="B94BEAABB169FF8EFDE8FF3A1F7FFF06" bold="true" box="[624,816,178,202]" italics="true" pageId="1" pageNumber="332">Podosilis groehni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857D0B169FF8EFCAEFF391FC4FF05" box="[822,907,177,201]" pageId="1" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3256532B169FF8EFDE8FF581EF0FF24" box="[624,703,208,232]" pageId="1" pageNumber="332" type="description">
<paragraph id="8B8036B9B169FF8EFDE8FF581EF0FF24" blockId="1.[624,907,177,232]" box="[624,703,208,232]" pageId="1" pageNumber="332">
(
<figureCitation id="13042A3CB169FF8EFDE0FF581EF9FF24" box="[632,694,208,232]" captionStart="Fig" captionStartId="2.[189,228,913,937]" captionTargetBox="[399,1006,192,876]" captionTargetId="figure-256@2.[399,1006,192,876]" captionTargetPageId="2" captionText="Fig. 1.Podosilis groehni sp. nov., holotype, No 5202 [GPIH]: A habitus, dorsal view; B habitus, ventral view." pageId="1" pageNumber="332">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3256532B169FF8EFDE8FE851FEAFAD5" pageId="1" pageNumber="332" type="materials_examined">
<paragraph id="8B8036B9B169FF8EFDE8FE851F84FDBB" blockId="1.[624,1103,269,631]" pageId="1" pageNumber="332">
<emphasis id="B94BEAABB169FF8EFDE8FE851F55FEE9" bold="true" box="[624,794,269,293]" pageId="1" pageNumber="332">Type material.</emphasis>
<typeStatus id="5484881BB169FF8EFCBCFE851FC4FEE9" box="[804,907,269,293]" pageId="1" pageNumber="332" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED2EAE7CB169FF8EFC01FE861F98FEEA" box="[921,983,270,294]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="1" pageNumber="332">GPIH</collectionCode>
no. 5202,
<collectionCode id="ED2EAE7CB169FF8EFDE8FEA41EF5FE88" box="[624,698,300,324]" pageId="1" pageNumber="332">CCGG</collectionCode>
no. 8701 (ex coll. Jonas Damzen JDC-12438); adult, male. A complete beetle with partially exposed metathoracic wings is included in a transparent, yellow amber piece with dimensions of 36×
<quantity id="4CC79B5CB169FF8EFC2FFE2F184BFE72" box="[951,1028,423,447]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="332" unit="mm" value="19.0">19 mm</quantity>
and a maximum thickness of
<quantity id="4CC79B5CB169FF8EFC1CFE4D1F82FE11" box="[900,973,453,477]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="332" unit="mm" value="6.0">6 mm</quantity>
; preserved without supplementary fixation. Syninclusions: one Brachycera (Diptera) specimen, two Nematocera (Diptera) specimen, seven?
<taxonomicName id="4C3F4D3AB169FF8EFD5FFDC81FE5FD94" box="[711,938,576,600]" class="Insecta" family="Melolonthidae" genus="Acarina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="subGenus" subGenus="Arachnida">Acarina (Arachnida)</taxonomicName>
nymphs, and few stellate
<taxonomicName id="4C3F4D3AB169FF8EFD68FDD71F1CFDBB" authorityName="Dumortier" authorityYear="1829" box="[752,851,607,631]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="332" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
trichomes.
</paragraph>
<paragraph id="8B8036B9B169FF8EFDE8FD141FD0FD3E" blockId="1.[624,1103,668,754]" pageId="1" pageNumber="332">
<materialsCitation id="3B573CE4B169FF8EFDE8FD141FD5FD3E" country="Russia" county="Sambian Peninsula" location="Amber" municipality="Yantarny" pageId="1" pageNumber="332" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B94BEAABB169FF8EFDE8FD141F57FD78" bold="true" box="[624,792,668,692]" pageId="1" pageNumber="332">
<typeStatus id="5484881BB169FF8EFDE8FD141EE7FD78" box="[624,680,668,692]" pageId="1" pageNumber="332">Type</typeStatus>
locality.
</emphasis>
<location id="8EE06062B169FF8EFCBAFD141F23FD78" LSID="urn:lsid:plazi:treatment:039687AFB169FF8CFDE8FF3A1FCAFBE8:8EE06062B169FF8EFCBAFD141F23FD78" box="[802,876,668,692]" country="Russia" county="Sambian Peninsula" municipality="Yantarny" name="Amber" pageId="1" pageNumber="332" stateProvince="Kaliningrad">Amber</location>
mine in
<collectingMunicipality id="6BE4ACC3B169FF8EFC70FD151800FD79" box="[1000,1103,669,693]" pageId="1" pageNumber="332">Yantarny</collectingMunicipality>
settlement, the
<collectingCounty id="62E14E35B169FF8EFCA8FD331841FD1F" box="[816,1038,699,723]" pageId="1" pageNumber="332">Sambian Peninsula</collectingCounty>
, the
<collectingRegion id="49FBF85BB169FF8EFDE8FD521F09FD3E" box="[624,838,730,754]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="332">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="F3287629B169FF8EFCC8FD521FD5FD3E" box="[848,922,730,754]" name="Russia" pageId="1" pageNumber="332">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8036B9B169FF8EFDE8FC9F1F1BFC47" blockId="1.[624,1103,791,907]" pageId="1" pageNumber="332">
<emphasis id="B94BEAABB169FF8EFDE8FC9F1F6FFCE3" bold="true" box="[624,800,791,815]" pageId="1" pageNumber="332">
<typeStatus id="5484881BB169FF8EFDE8FC9F1EE7FCE3" box="[624,680,791,815]" pageId="1" pageNumber="332">Type</typeStatus>
stratum.
</emphasis>
Baltic amber; Middle Upper Eocene (
<bibRefCitation id="EFAE4B48B169FF8EFC81FCBD1FB0FC82" author="Sadowski E. - M. &amp; Schmidt A. R. &amp; Seyfullah L. J. &amp; Kunzmann L." box="[793,1023,821,846]" pageId="1" pageNumber="332" pagination="1 - 73" refId="ref4468" refString="Sadowski E. - M., Schmidt A. R., Seyfullah L. J., Kunzmann L. 2017. Conifers of the ' Baltic amber forest' and their palaeoecological significance. Stapfia 106: 1 - 73." type="journal article" year="2017">Sadowski et al. 2017</bibRefCitation>
,
<bibRefCitation id="EFAE4B48B169FF8EFB94FCBE1804FC82" author="Sadowski E. - M. &amp; Schmidt A. R. &amp; Denk T." box="[1036,1099,822,846]" pageId="1" pageNumber="332" refId="ref4511" refString="Sadowski E. - M., Schmidt A. R., Denk T. 2020. Staminate inflorescences with in situ pollen from Eocene Baltic amber reveal high diversity in Fagaceae (oak family). Willdenowia: 50: 405 - 517 + Supplement." type="book" year="2020">2020</bibRefCitation>
;
<bibRefCitation id="EFAE4B48B169FF8EFDE8FCDC1F2FFCA0" author="Seyfullah L. J. &amp; Beimforde C. &amp; Dal Corso J. &amp; Perrichot V. &amp; Rikkinen J. &amp; Schmidt A. R." box="[624,864,852,876]" pageId="1" pageNumber="332" pagination="1684 - 1714" refId="ref4559" refString="Seyfullah L. J., Beimforde C., Dal Corso J., Perrichot V., Rikkinen J., Schmidt A. R. 2018. Production and preservation of resins - past and present. Biological Reviews 93: 1684 - 1714." type="journal article" year="2018">Seyfullah et al. 2018</bibRefCitation>
;
<bibRefCitation id="EFAE4B48B169FF8EFCE8FCDC1804FCA0" author="Bukejs A. &amp; Alekseev V. I. &amp; Pollock D. A." box="[880,1099,852,876]" pageId="1" pageNumber="332" pagination="261 - 273" refId="ref3689" refString="Bukejs A., Alekseev V. I., Pollock D. A. 2019. Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa 4664 (2): 261 - 273." type="journal article" year="2019">Bukejs et al. 2019</bibRefCitation>
;
<bibRefCitation id="EFAE4B48B169FF8EFDE8FCFB1F07FC47" author="Kasinski J. R. &amp; Kramarska R. &amp; Slodkowska B. &amp; Sivkov V. &amp; Piwocki M." box="[624,840,883,907]" pageId="1" pageNumber="332" pagination="29 - 53" refId="ref4100" refString="Kasinski J. R., Kramarska R., Slodkowska B., Sivkov V., Piwocki M. 2020. Paleocene and Eocene deposits on the eastern margin of the Gulf of Gdansk (Yantarny P- 1 borehole, Kaliningrad region, Russia). Geological Quarterly 64 (1): 29 - 53." type="journal article" year="2020">Kasiński et al. 2020</bibRefCitation>
).
</paragraph>
<paragraph id="8B8036B9B169FF8EFDE8FC381FEAFAD5" blockId="1.[624,1103,944,1305]" pageId="1" pageNumber="332">
<emphasis id="B94BEAABB169FF8EFDE8FC381FC6FC04" bold="true" box="[624,905,944,968]" pageId="1" pageNumber="332">Systematic placement.</emphasis>
The specimen considered here was assigned to the genus
<taxonomicName id="4C3F4D3AB169FF8EFDE8FC661E9CFBCA" authorityName="Wittmer" authorityYear="1978" box="[624,723,1006,1030]" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB169FF8EFDE8FC661E9CFBCA" box="[624,723,1006,1030]" italics="true" pageId="1" pageNumber="332">Podosilis</emphasis>
</taxonomicName>
based on a combination of the following morphological characters: (1) internal claws of all legs apically bifurcate (simple in
<taxonomicName id="4C3F4D3AB169FF8EFD74FBC11F7BFBAD" authorityName="Reitter" authorityYear="1887" box="[748,820,1097,1121]" class="Insecta" family="Cantharidae" genus="Eusilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB169FF8EFD74FBC11F7BFBAD" box="[748,820,1097,1121]" italics="true" pageId="1" pageNumber="332">Eusilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3F4D3AB169FF8EFCDEFBC11F3BFBAD" authorityName="Charpentier" authorityYear="1825" box="[838,884,1097,1121]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB169FF8EFCDEFBC11F3BFBAD" box="[838,884,1097,1121]" italics="true" pageId="1" pageNumber="332">Silis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D3AB169FF8EFC2AFBC11840FBAD" authorityName="Kazantsev" authorityYear="2011" box="[946,1039,1097,1121]" class="Insecta" family="Cantharidae" genus="Autosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB169FF8EFC2AFBC11840FBAD" box="[946,1039,1097,1121]" italics="true" pageId="1" pageNumber="332">Autosilis</emphasis>
</taxonomicName>
, and only the internal claw of forelegs apically bifurcate in
<taxonomicName id="4C3F4D3AB169FF8EFC9BFB0F1F12FB53" authorityName="Wittmer" authorityYear="1977" box="[771,861,1159,1183]" class="Insecta" family="Cantharidae" genus="Asiosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB169FF8EFC9BFB0F1F12FB53" box="[771,861,1159,1183]" italics="true" pageId="1" pageNumber="332">Asiosilis</emphasis>
</taxonomicName>
), (2) claws without denticle basally (all claws with large denticle in
<taxonomicName id="4C3F4D3AB169FF8EFD71FB4C1F0CFB10" authorityName="Wittmer" authorityYear="1977" box="[745,835,1220,1244]" class="Insecta" family="Cantharidae" genus="Asiosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB169FF8EFD71FB4C1F0CFB10" box="[745,835,1220,1244]" italics="true" pageId="1" pageNumber="332">Asiosilis</emphasis>
</taxonomicName>
), and (3) tarsomere 4 is widest than others tarsomeres (narrow as others tarsomeres in
<taxonomicName id="4C3F4D3AB169FF8EFCD5FA891FDAFAD5" authorityName="Reitter" authorityYear="1887" box="[845,917,1281,1305]" class="Insecta" family="Cantharidae" genus="Eusilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB169FF8EFCD5FA891FDAFAD5" box="[845,917,1281,1305]" italics="true" pageId="1" pageNumber="332">Eusilis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3256532B169FF8DFDE8FAB71F81FFBA" lastPageId="2" lastPageNumber="333" pageId="1" pageNumber="332" type="diagnosis">
<paragraph id="8B8036B9B169FF8EFDE8FAB71E93F9A7" blockId="1.[624,1109,1343,1643]" pageId="1" pageNumber="332">
<emphasis id="B94BEAABB169FF8EFDE8FAB71F35FA9B" bold="true" box="[624,890,1343,1367]" pageId="1" pageNumber="332">Differential diagnosis.</emphasis>
<taxonomicName id="4C3F4D3AB169FF8EFC1BFAB71807FA9B" authorityName="Bukejs &amp; Fanti" authorityYear="2023" box="[899,1096,1343,1367]" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
<emphasis id="B94BEAABB169FF8EFC1BFAB71807FA9B" box="[899,1096,1343,1367]" italics="true" pageId="1" pageNumber="332">Podosilis groehni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857D0B169FF8EFDE8FAD61E99FAB9" box="[624,726,1374,1397]" pageId="1" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C3F4D3AB169FF8EFC05FAD61F6FFA58" authority="Kazantsev, 2020" authorityName="Kazantsev" authorityYear="2020" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="gedaniensis">
<emphasis id="B94BEAABB169FF8EFC05FAD61807FABA" box="[925,1096,1374,1398]" italics="true" pageId="1" pageNumber="332">P. gedaniensis</emphasis>
<bibRefCitation id="EFAE4B48B169FF8EFDE8FAF51F6FFA58" author="Kazantsev S. V." box="[624,800,1404,1428]" pageId="1" pageNumber="332" pagination="260 - 268" refId="ref4231" refString="Kazantsev S. V. 2020. New Baltic amber soldier beetles (Insecta: Coleoptera: Cantharidae) with some taxonomic notes. Palaeoentomology 3 (3): 260 - 268." type="journal article" year="2020">Kazantsev, 2020</bibRefCitation>
</taxonomicName>
(the single fossil species of the genus, the
<typeStatus id="5484881BB169FF8EFC83FA131F37FA7F" box="[795,888,1435,1459]" pageId="1" pageNumber="332" type="holotype">holotype</typeStatus>
is female) by the obsolete elytral humeral costae (distinct in
<taxonomicName id="4C3F4D3AB169FF8EFBA3FA321EA1FA3C" authorityName="Kazantsev" authorityYear="2020" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="gedaniensis">
<emphasis id="B94BEAABB169FF8EFBA3FA321EA1FA3C" italics="true" pageId="1" pageNumber="332">P. gedaniensis</emphasis>
</taxonomicName>
); this character cannot be considered as sexual dimorphism. Also, the new species differs from the extant species of
<taxonomicName id="4C3F4D3AB169FF8EFD08F9BC1EB5F980" authorityName="Wittmer" authorityYear="1978" box="[656,762,1588,1612]" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB169FF8EFD08F9BC1EB5F980" box="[656,762,1588,1612]" italics="true" pageId="1" pageNumber="332">Podosilis</emphasis>
</taxonomicName>
in having a extremely smaller body size.
</paragraph>
<paragraph id="8B8036B9B16AFF8DFE1BFFEB1F81FFBA" blockId="2.[387,974,99,118]" box="[387,974,99,118]" pageId="2" pageNumber="333">
<emphasis id="B94BEAABB16AFF8DFE1BFFEB1F81FFBA" box="[387,974,99,118]" italics="true" pageId="2" pageNumber="333">
Two new fossil
<taxonomicName id="4C3F4D3AB16AFF8DFD9FFFEB1E0CFFBA" authorityName="Mulsant" authorityYear="1862" box="[519,579,99,118]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="subFamily" subFamily="Silinae">Silinae</taxonomicName>
(
<taxonomicName id="4C3F4D3AB16AFF8DFDD6FFEB1EFFFFBA" authorityName="Linnaeus" authorityYear="1758" box="[590,688,99,118]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="2" pageNumber="333" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
,
<taxonomicName id="4C3F4D3AB16AFF8DFD22FFEB1F68FFBA" authorityName="Imhoff" authorityYear="1856" box="[698,807,99,118]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="333" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
) from Baltic amber
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF406631B16AFF8DFF25FC191DEDFC04" pageId="2" pageNumber="333" startId="2.[189,228,913,937]" targetBox="[399,1006,192,876]" targetPageId="2" targetType="figure">
<paragraph id="8B8036B9B16AFF8DFF25FC191DEDFC04" blockId="2.[189,1172,913,968]" pageId="2" pageNumber="333">
Fig. 1.
<taxonomicName id="4C3F4D3AB16AFF8DFE92FC191D89FC65" authorityName="Bukejs &amp; Fanti" authorityYear="2023" box="[266,454,913,937]" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="333" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
<emphasis id="B94BEAABB16AFF8DFE92FC191D89FC65" box="[266,454,913,937]" italics="true" pageId="2" pageNumber="333">Podosilis groehni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857D0B16AFF8DFE55FC1A1E6CFC65" box="[461,547,914,937]" pageId="2" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, No 5202 [GPIH]: A habitus, dorsal view; B habitus, ventral view.
</paragraph>
</caption>
<subSubSection id="C3256532B16AFF8CFF25FC651FA0FD1E" lastPageId="3" lastPageNumber="334" pageId="2" pageNumber="333" type="description">
<paragraph id="8B8036B9B16AFF8DFF25FC651EC0FAD5" blockId="2.[189,669,1005,1305]" pageId="2" pageNumber="333">
<emphasis id="B94BEAABB16AFF8DFF25FC651D01FBC9" bold="true" box="[189,334,1005,1029]" pageId="2" pageNumber="333">Description.</emphasis>
Body elongate-oval, slightly flatened dorsally; sparsely covered with homogenous, short, semierect setae; integument unicolorous brown (as preserved). Measurements: total body length 3.0 mm (from anterior margin of head to elytral apex); pronotum length
<quantity id="4CC79B5CB16AFF8DFDECFB2D1CA6FB17" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="333" unit="mm" value="0.5">0.5 mm</quantity>
, pronotum maximum width
<quantity id="4CC79B5CB16AFF8DFDAEFB4C1ED7FB17" box="[566,664,1220,1244]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="333" unit="mm" value="0.8">0.8 mm</quantity>
; elytra length
<quantity id="4CC79B5CB16AFF8DFEFBFB6B1D8CFB36" box="[355,451,1251,1275]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="333" unit="mm" value="2.4">2.4 mm</quantity>
, elytra maximum width
<quantity id="4CC79B5CB16AFF8DFE99FA891D1AFAD5" box="[257,341,1281,1305]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="333" unit="mm" value="1.2">1.2 mm</quantity>
, elytra width at base 1.0 mm.
</paragraph>
<paragraph id="8B8036B9B16AFF8DFF25FAB618EDF9A6" blockId="2.[189,675,1342,1642]" lastBlockId="2.[699,1186,1005,1642]" pageId="2" pageNumber="333">Head short, transverse, pubescent, with fine punctation, partially covered by pronotum; frons convex. Compound eyes large, hemispherical and strongly prominent, inserted laterally. Mandibles large, simple, and falciform. Maxillary palpi 4-segmented, palpomeres unequal in length, terminal palpomere weakly securiform. Labial palps 3-segmented, terminal palpomere securiform. Antennae inserted far from inner margin of eyes, rather long, extending about to posterior one-third of elytral length; 11-segmented, filiform, all antennomeres covered with short, semierect setae; scape subcylindrical, dilated apically, elongate, 2.5× longer than wide, robust, curved; antennomere 2 shortest, subtrapezoidal, dilated apically, slightly elongate, 1.25× longer than wide, about 0.5× as long as scape; antennomere 3 subcylindrical, slightly dilated apically, elongate, 2.25× longer than wide, slightly shorter than antennomere 4; antennomeres 48 equal in size and shape, robust, subcylindrical, dilated apically, elongate, 2.5× longer than wide, about 2.0× as long as antennomere 2; antennomeres 910 equal equal in size and shape, robust, subcylindrical, dilated apically, slightly shorter than antennomere 8; antennomere 11 elongate-oval, 3.1× longer than wide, narrowly rounded at apex;</paragraph>
<paragraph id="8B8036B9B16AFF8DFBF4F92E18DBF972" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="333">333</paragraph>
<paragraph id="8B8036B9B16BFF8CFFEAFF3A1E42FF24" blockId="3.[114,593,178,723]" pageId="3" pageNumber="334">relative length ratios of antennomeres 11 equal to 10:5:9:10:10:10:10:10:9:9:11.</paragraph>
<paragraph id="8B8036B9B16BFF8CFFEAFF671E4CFD1F" blockId="3.[114,593,178,723]" pageId="3" pageNumber="334">Pronotum subtrapezoidal, transverse, 1.6× wider than long, widest in posterior one-third of its length, slightly wider than head; punctation fine and sparse on disc, and distinctly denser laterally; disc convex, with subtriangular impressions basally and suboval impressions anterolaterally. Pronotal apical margin slightly rounded; basal margin almost straight; lateral margins rounded, incised near base and near anterior angles, protruding posteromedially with two small and obtuse tooth. Anterior angles obtuse, slightly projecting; posterior angles acute, distinctly projecting posteriad forming denticle. Prohypomera densely covered with fine punctation.</paragraph>
<paragraph id="8B8036B9B16BFF8CFFEAFD701E04FB71" blockId="3.[114,599,760,1336]" pageId="3" pageNumber="334">Scutellar shield subtriangular with widely rounded apex, with few fine punctutes. Elytra wider than pronotum, elongate-oval, 2.0× longer than wide, subparallel-sided in anterior half and slightly widened in posterior half, rounded apically, completely covering abdomen; humeral costae obsolete, slightly distinct in anterior two-thirds of elytral lenght; with large oval impression post-medially; elytral punctation small and rather dense, distance between punctures about 0.71.3× diameter of one puncture, punctures distinctly denser laterally. Metanepisternum subtriangular, narrowed posteriad, with fine punctation.</paragraph>
<paragraph id="8B8036B9B16BFF8CFFEAFB4C1E04FAD5" blockId="3.[114,599,760,1336]" pageId="3" pageNumber="334">Metaventrite wide, with rounded posterior margin, covered with fine punctation; disc convex, apparently with discrimen.</paragraph>
<paragraph id="8B8036B9B16BFF8CFFEAFAA81DB5FAF4" blockId="3.[114,599,760,1336]" box="[114,506,1312,1336]" pageId="3" pageNumber="334">Metathoracic wings fully developed.</paragraph>
<paragraph id="8B8036B9B16BFF8CFFEAFAD61E1EF980" blockId="3.[114,593,1373,1612]" pageId="3" pageNumber="334">Legs rather long, slender, pubescent. Procoxae conical, large; mesocoxae narrowly oval, convex; metacoxae narrow, strongly transverse. Trochanters elongate, convex. Femora narrow, slightly flattened, straight. Tibiae subcylindrical, apparently with two spines apically, pro- and mesotibiae nearly as long as pro- and</paragraph>
<paragraph id="8B8036B9B16BFF8CFFEAF92E1859FE30" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1109,178,508]" pageId="3" pageNumber="334">334 mesofemora respectively, metatibiae distinctly longer than metafemora. Tarsal formula 5-5-5; tarsomere 1 longest, elongate, slightly dilated apically; tarsomere 2 shorter than tarsomere 1; tarsomere 3 subtriangular, dilated apically, shorter than tarsomere 2; tarsomere 4 widest, deeply bilobed; tarsomere 5 subcylindrical, elongate, curved, enlarged apically. Pretarsal claws without basal tooth, inner claw of all legs bifurcate apically.</paragraph>
<paragraph id="8B8036B9B16BFF8CFDE8FDAA1FA0FD1E" blockId="3.[624,1103,546,723]" pageId="3" pageNumber="334">Abdomen with seven visible sternites (ventrites), covered fine, inconspicuous punctation; sternite 7 with elongate lobe with rounded apex; ultimate tergite longer and wider than ultimate sternite, with prominent conical lateral processes.</paragraph>
</subSubSection>
<subSubSection id="C3256532B16BFF8CFDE8FD701EB0FC47" pageId="3" pageNumber="334" type="etymology">
<paragraph id="8B8036B9B16BFF8CFDE8FD701EB0FC47" blockId="3.[624,1103,760,907]" pageId="3" pageNumber="334">
<emphasis id="B94BEAABB16BFF8CFDE8FD701EB8FCDC" bold="true" box="[624,759,760,784]" pageId="3" pageNumber="334">Etymology.</emphasis>
The specific epithet is a patronym; the new species is named in honor of Mr. Carsten Gröhn (Glinde,
<collectingCountry id="F3287629B16BFF8CFDE8FCDC1E97FCA0" box="[624,728,852,876]" name="Germany" pageId="3" pageNumber="334">Germany</collectingCountry>
), an enthusiast and specialist in Baltic amber.
</paragraph>
</subSubSection>
<subSubSection id="C3256532B16BFF8CFDE8FC381FCAFBE8" pageId="3" pageNumber="334" type="discussion">
<paragraph id="8B8036B9B16BFF8CFDE8FC381FCAFBE8" blockId="3.[624,1103,944,1060]" pageId="3" pageNumber="334">
<emphasis id="B94BEAABB16BFF8CFDE8FC381E95FC04" bold="true" box="[624,730,944,968]" pageId="3" pageNumber="334">Remark.</emphasis>
The lateral margins of pronotum incised and denticulated, and the ultimate abdominal sternite long and narrow indicate that the specimen is male.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,285 @@
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<subSubSection id="C3256532B16BFF8CFDE8FB2D1FEEFB71" box="[624,929,1189,1213]" pageId="3" pageNumber="334" type="nomenclature">
<paragraph id="8B8036B9B16BFF8CFDE8FB2D1FEEFB71" blockId="3.[624,929,1189,1244]" box="[624,929,1189,1213]" pageId="3" pageNumber="334">
<heading id="D0C881D5B16BFF8CFDE8FB2D1FEEFB71" bold="true" box="[624,929,1189,1213]" fontSize="10" level="2" pageId="3" pageNumber="334" reason="4">
<emphasis id="B94BEAABB16BFF8CFDE8FB2D1FEEFB71" bold="true" box="[624,929,1189,1213]" pageId="3" pageNumber="334">
<taxonomicName id="4C3F4D3AB16BFF8CFDE8FB2D1F04FB71" authorityName="Bukejs &amp; Fanti" authorityYear="2023" box="[624,843,1189,1213]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="carsteni" status="sp. nov." subGenus="Silis">
<emphasis id="B94BEAABB16BFF8CFDE8FB2D1EE9FB71" bold="true" box="[624,678,1189,1213]" italics="true" pageId="3" pageNumber="334">Silis</emphasis>
(
<emphasis id="B94BEAABB16BFF8CFD36FB2D1E92FB71" bold="true" box="[686,733,1189,1213]" italics="true" pageId="3" pageNumber="334">Silis</emphasis>
)
<emphasis id="B94BEAABB16BFF8CFD75FB2D1F04FB71" bold="true" box="[749,843,1189,1213]" italics="true" pageId="3" pageNumber="334">carsteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857D0B16BFF8CFCD4FB2D1FEEFB71" box="[844,929,1189,1213]" pageId="3" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3256532B16BFF8CFDE8FB4C1E8FFB10" box="[624,704,1220,1244]" pageId="3" pageNumber="334" type="description">
<paragraph id="8B8036B9B16BFF8CFDE8FB4C1E8FFB10" blockId="3.[624,929,1189,1244]" box="[624,704,1220,1244]" pageId="3" pageNumber="334">
(
<figureCitation id="13042A3CB16BFF8CFDE0FB4C1EF7FB10" box="[632,696,1220,1244]" captionStart="Fig" captionStartId="4.[189,228,792,816]" captionTargetBox="[332,1118,179,753]" captionTargetId="figure-338@4.[332,1118,179,753]" captionTargetPageId="4" captionText="Fig. 2.Silis(Silis) carsteni sp. nov., holotype, No 5203 [GPIH]: A habitus, dorsal view; B habitus, ventral view." pageId="3" pageNumber="334">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3256532B16BFF8BFDE8FA891F87FC67" lastPageId="4" lastPageNumber="335" pageId="3" pageNumber="334" type="materials_examined">
<paragraph id="8B8036B9B16BFF8CFDE8FA891EAFF9E1" blockId="3.[624,1103,1281,1581]" pageId="3" pageNumber="334">
<emphasis id="B94BEAABB16BFF8CFDE8FA891F55FAD5" bold="true" box="[624,794,1281,1305]" pageId="3" pageNumber="334">Type material.</emphasis>
<materialsCitation id="3B573CE4B16BFF8CFCBCFA891EAFF9E1" collectionCode="GPIH, CCGG" collectorName="Jonas Damzen &amp; Syninclusions" pageId="3" pageNumber="334" specimenCount="1" typeStatus="holotype">
<typeStatus id="5484881BB16BFF8CFCBCFA891FC4FAD5" box="[804,907,1281,1305]" pageId="3" pageNumber="334" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED2EAE7CB16BFF8CFC01FA891F98FAD5" box="[921,983,1281,1305]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="3" pageNumber="334">GPIH</collectionCode>
no. 5203,
<collectionCode id="ED2EAE7CB16BFF8CFDE8FAA81EF5FAF4" box="[624,698,1312,1336]" pageId="3" pageNumber="334">CCGG</collectionCode>
no. 8702, (ex coll.
<collectorName id="26CA536FB16BFF8CFC30FAA91807FAF4" box="[936,1096,1313,1336]" pageId="3" pageNumber="334">Jonas Damzen</collectorName>
JDC-12439); adult, female. A complete beetle with partially exposed metathoracic wings is included in a transparent, yellow amber piece with dimensions of 31×
<quantity id="4CC79B5CB16BFF8CFC64FA131800FA7E" box="[1020,1103,1435,1459]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="334" unit="mm" value="16.0">16 mm</quantity>
and a maximum thickness of
<quantity id="4CC79B5CB16BFF8CFC6CFA311805FA1D" box="[1012,1098,1465,1489]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="334" unit="mm" value="5.0">5 mm</quantity>
; preserved without supplementary fixation.
<collectorName id="26CA536FB16BFF8CFDE8FA7F1F45F9C3" box="[624,778,1527,1551]" pageId="3" pageNumber="334">Syninclusions</collectorName>
: few stellate
<taxonomicName id="4C3F4D3AB16BFF8CFC7DFA7F1800F9C3" authorityName="Dumortier" authorityYear="1829" box="[997,1103,1527,1551]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="334" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
trichomes.
</materialsCitation>
</paragraph>
<paragraph id="8B8036B9B16CFF8BFE1BFFEB1F81FFBA" blockId="4.[387,974,99,118]" box="[387,974,99,118]" pageId="4" pageNumber="335">
<emphasis id="B94BEAABB16CFF8BFE1BFFEB1F81FFBA" box="[387,974,99,118]" italics="true" pageId="4" pageNumber="335">
Two new fossil
<taxonomicName id="4C3F4D3AB16CFF8BFD9FFFEB1E0CFFBA" authorityName="Mulsant" authorityYear="1862" box="[519,579,99,118]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="subFamily" subFamily="Silinae">Silinae</taxonomicName>
(
<taxonomicName id="4C3F4D3AB16CFF8BFDD6FFEB1EFFFFBA" authorityName="Linnaeus" authorityYear="1758" box="[590,688,99,118]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="335" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
,
<taxonomicName id="4C3F4D3AB16CFF8BFD22FFEB1F68FFBA" authorityName="Imhoff" authorityYear="1856" box="[698,807,99,118]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
) from Baltic amber
</emphasis>
</paragraph>
<caption id="DF406631B16CFF8BFF25FC901D81FC83" pageId="4" pageNumber="335" startId="4.[189,228,792,816]" targetBox="[332,1118,179,753]" targetPageId="4" targetType="figure">
<paragraph id="8B8036B9B16CFF8BFF25FC901D81FC83" blockId="4.[189,1179,792,847]" pageId="4" pageNumber="335">
Fig. 2.
<taxonomicName id="4C3F4D3AB16CFF8BFE95FC901DADFCFC" authorityName="Bukejs &amp; Fanti" authorityYear="2023" box="[269,482,792,817]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="carsteni" status="sp. nov." subGenus="Silis">
<emphasis id="B94BEAABB16CFF8BFE95FC901D0EFCFC" box="[269,321,792,816]" italics="true" pageId="4" pageNumber="335">Silis</emphasis>
(
<emphasis id="B94BEAABB16CFF8BFED4FC901D35FCFC" box="[332,378,792,816]" italics="true" pageId="4" pageNumber="335">Silis</emphasis>
)
<emphasis id="B94BEAABB16CFF8BFE14FC911DADFCFC" box="[396,482,793,816]" italics="true" pageId="4" pageNumber="335">carsteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857D0B16CFF8BFE72FC911E0DFCFC" box="[490,578,793,816]" pageId="4" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, No 5203 [GPIH]: A habitus, dorsal view; B habitus, ventral view.
</paragraph>
</caption>
<paragraph id="8B8036B9B16CFF8BFF25FCFC1DA3FC06" blockId="4.[189,668,884,970]" pageId="4" pageNumber="335">
<materialsCitation id="3B573CE4B16CFF8BFF25FCFC1DA7FC06" country="Russia" county="Sambian Peninsula" location="Amber" municipality="Yantarny" pageId="4" pageNumber="335" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B94BEAABB16CFF8BFF25FCFC1D2AFC40" bold="true" box="[189,357,884,908]" pageId="4" pageNumber="335">
<typeStatus id="5484881BB16CFF8BFF25FCFC1CBAFC40" box="[189,245,884,908]" pageId="4" pageNumber="335">Type</typeStatus>
locality.
</emphasis>
<location id="8EE06062B16CFF8BFEE8FCFC1DF5FC40" LSID="urn:lsid:plazi:treatment:039687AFB16BFF89FDE8FB2D1ED2FB8F:8EE06062B16CFF8BFEE8FCFC1DF5FC40" box="[368,442,884,908]" country="Russia" county="Sambian Peninsula" municipality="Yantarny" name="Amber" pageId="4" pageNumber="335" stateProvince="Kaliningrad">Amber</location>
mine in
<collectingMunicipality id="6BE4ACC3B16CFF8BFDADFCFD1ED3FC41" box="[565,668,885,909]" pageId="4" pageNumber="335">Yantarny</collectingMunicipality>
settlement, the
<collectingCounty id="62E14E35B16CFF8BFEE5FC1B1E13FC67" box="[381,604,915,939]" pageId="4" pageNumber="335">Sambian Peninsula</collectingCounty>
, the
<collectingRegion id="49FBF85BB16CFF8BFF25FC3A1DDBFC06" box="[189,404,946,970]" country="Russia" name="Kaliningrad" pageId="4" pageNumber="335">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="F3287629B16CFF8BFE06FC3A1DA7FC06" box="[414,488,946,970]" name="Russia" pageId="4" pageNumber="335">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8036B9B16CFF8BFF25FC671DEEFBAF" blockId="4.[189,669,1007,1123]" pageId="4" pageNumber="335">
<emphasis id="B94BEAABB16CFF8BFF25FC671D21FBCB" bold="true" box="[189,366,1007,1031]" pageId="4" pageNumber="335">
<typeStatus id="5484881BB16CFF8BFF25FC671CBAFBCB" box="[189,245,1007,1031]" pageId="4" pageNumber="335">Type</typeStatus>
stratum.
</emphasis>
Baltic amber; Middle Upper Eocene (
<bibRefCitation id="EFAE4B48B16CFF8BFEFEFB851E02FBEA" author="Sadowski E. - M. &amp; Schmidt A. R. &amp; Seyfullah L. J. &amp; Kunzmann L." box="[358,589,1037,1062]" pageId="4" pageNumber="335" pagination="1 - 73" refId="ref4468" refString="Sadowski E. - M., Schmidt A. R., Seyfullah L. J., Kunzmann L. 2017. Conifers of the ' Baltic amber forest' and their palaeoecological significance. Stapfia 106: 1 - 73." type="journal article" year="2017">Sadowski et al. 2017</bibRefCitation>
,
<bibRefCitation id="EFAE4B48B16CFF8BFDC1FB861ED7FBEA" author="Sadowski E. - M. &amp; Schmidt A. R. &amp; Denk T." box="[601,664,1038,1062]" pageId="4" pageNumber="335" refId="ref4511" refString="Sadowski E. - M., Schmidt A. R., Denk T. 2020. Staminate inflorescences with in situ pollen from Eocene Baltic amber reveal high diversity in Fagaceae (oak family). Willdenowia: 50: 405 - 517 + Supplement." type="book" year="2020">2020</bibRefCitation>
;
<bibRefCitation id="EFAE4B48B16CFF8BFF25FBA41DE1FB88" author="Seyfullah L. J. &amp; Beimforde C. &amp; Dal Corso J. &amp; Perrichot V. &amp; Rikkinen J. &amp; Schmidt A. R." box="[189,430,1068,1092]" pageId="4" pageNumber="335" pagination="1684 - 1714" refId="ref4559" refString="Seyfullah L. J., Beimforde C., Dal Corso J., Perrichot V., Rikkinen J., Schmidt A. R. 2018. Production and preservation of resins - past and present. Biological Reviews 93: 1684 - 1714." type="journal article" year="2018">Seyfullah et al. 2018</bibRefCitation>
;
<bibRefCitation id="EFAE4B48B16CFF8BFE26FBA41ED7FB88" author="Bukejs A. &amp; Alekseev V. I. &amp; Pollock D. A." box="[446,664,1068,1092]" pageId="4" pageNumber="335" pagination="261 - 273" refId="ref3689" refString="Bukejs A., Alekseev V. I., Pollock D. A. 2019. Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa 4664 (2): 261 - 273." type="journal article" year="2019">Bukejs et al. 2019</bibRefCitation>
;
<bibRefCitation id="EFAE4B48B16CFF8BFF25FBC31DD9FBAF" author="Kasinski J. R. &amp; Kramarska R. &amp; Slodkowska B. &amp; Sivkov V. &amp; Piwocki M." box="[189,406,1099,1123]" pageId="4" pageNumber="335" pagination="29 - 53" refId="ref4100" refString="Kasinski J. R., Kramarska R., Slodkowska B., Sivkov V., Piwocki M. 2020. Paleocene and Eocene deposits on the eastern margin of the Gulf of Gdansk (Yantarny P- 1 borehole, Kaliningrad region, Russia). Geological Quarterly 64 (1): 29 - 53." type="journal article" year="2020">Kasiński et al. 2020</bibRefCitation>
).
</paragraph>
<paragraph id="8B8036B9B16CFF8BFF25FB001F87FC67" blockId="4.[189,668,1160,1614]" lastBlockId="4.[699,1178,884,939]" pageId="4" pageNumber="335">
<emphasis id="B94BEAABB16CFF8BFF25FB001D9EFB6C" bold="true" box="[189,465,1160,1184]" pageId="4" pageNumber="335">Systematic placement.</emphasis>
The specimen considered here was assigned to the genus
<taxonomicName id="4C3F4D3AB16CFF8BFF25FB4E1CA4FB12" authorityName="Charpentier" authorityYear="1825" box="[189,235,1222,1246]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB16CFF8BFF25FB4E1CA4FB12" box="[189,235,1222,1246]" italics="true" pageId="4" pageNumber="335">Silis</emphasis>
</taxonomicName>
based on a combination of the following morphological characters: (1) the pronotum non-incised at lateral sides (incised in females of
<taxonomicName id="4C3F4D3AB16CFF8BFE78FAAA1E0CFAF6" authorityName="Wittmer" authorityYear="1978" box="[480,579,1314,1338]" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB16CFF8BFE78FAAA1E0CFAF6" box="[480,579,1314,1338]" italics="true" pageId="4" pageNumber="335">Podosilis</emphasis>
</taxonomicName>
), with impressions (without impressions in
<taxonomicName id="4C3F4D3AB16CFF8BFF25FAD71D55FABB" authorityName="Kazantsev" authorityYear="2011" box="[189,282,1375,1399]" class="Insecta" family="Cantharidae" genus="Autosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB16CFF8BFF25FAD71D55FABB" box="[189,282,1375,1399]" italics="true" pageId="4" pageNumber="335">Autosilis</emphasis>
</taxonomicName>
), (2) abdomen with six visible sternites (seven sternites in females of
<taxonomicName id="4C3F4D3AB16CFF8BFF25FA141D6FFA78" authorityName="Wittmer" authorityYear="1978" box="[189,288,1436,1460]" class="Insecta" family="Cantharidae" genus="Podosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB16CFF8BFF25FA141D6FFA78" box="[189,288,1436,1460]" italics="true" pageId="4" pageNumber="335">Podosilis</emphasis>
</taxonomicName>
), (3) tarsomere 4 wider than others tarsomeres (tarsomere 4 as wide as tarsomeres
<quantity id="4CC79B5CB16CFF8BFEA0FA521DCCFA3E" box="[312,387,1498,1522]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="4" pageNumber="335" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">13 in</quantity>
<taxonomicName id="4C3F4D3AB16CFF8BFE1CFA521D83FA3E" authorityName="Reitter" authorityYear="1887" box="[388,460,1498,1522]" class="Insecta" family="Cantharidae" genus="Eusilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB16CFF8BFE1CFA521D83FA3E" box="[388,460,1498,1522]" italics="true" pageId="4" pageNumber="335">Eusilis</emphasis>
</taxonomicName>
), (4) claws simple, internal claw with obtuse and small tooth basally (all claws with large denticle basally in
<taxonomicName id="4C3F4D3AB16CFF8BFF47F9BE1D76F982" authorityName="Wittmer" authorityYear="1977" box="[223,313,1590,1614]" class="Insecta" family="Cantharidae" genus="Asiosilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAABB16CFF8BFF47F9BE1D76F982" box="[223,313,1590,1614]" italics="true" pageId="4" pageNumber="335">Asiosilis</emphasis>
</taxonomicName>
), and (5) the distal margin of ultimate tergite without conspicuous processes at lateral sides.
</paragraph>
</subSubSection>
<subSubSection id="C3256532B16CFF8AFD23FC581D9EFF24" lastPageId="5" lastPageNumber="336" pageId="4" pageNumber="335" type="diagnosis">
<paragraph id="8B8036B9B16CFF8BFD23FC5818D5F9A0" blockId="4.[699,1179,976,1644]" pageId="4" pageNumber="335">
<emphasis id="B94BEAABB16CFF8BFD23FC581F89FC24" bold="true" box="[699,966,976,1000]" pageId="4" pageNumber="335">Differential diagnosis.</emphasis>
A new species resembles
<taxonomicName id="4C3F4D3AB16CFF8BFCB7FC671FD3FBEA" authority="Fanti &amp; M. G. Pankowski, 2022" authorityName="Fanti &amp; M. G. Pankowski" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="boninoi">
<emphasis id="B94BEAABB16CFF8BFCB7FC671F12FBCB" box="[815,861,1007,1031]" italics="true" pageId="4" pageNumber="335">Silis</emphasis>
(
<emphasis id="B94BEAABB16CFF8BFCE8FC781FE4FBCB" box="[880,939,1008,1031]" italics="true" pageId="4" pageNumber="335">s. str.</emphasis>
)
<emphasis id="B94BEAABB16CFF8BFC24FC671842FBCB" box="[956,1037,1007,1031]" italics="true" pageId="4" pageNumber="335">boninoi</emphasis>
Fanti &amp; M. G. Pankowski, 2022
</taxonomicName>
(Baltic amber; male), but differs in flat and widely explanate pronotal lateral sides, pronotum widest basally (in contrast to pronotum widest in the middle in
<taxonomicName id="4C3F4D3AB16CFF8BFCC3FB001F84FB6C" authorityName="Fanti &amp; M. G. Pankowski" authorityYear="2022" box="[859,971,1160,1184]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="boninoi">
<emphasis id="B94BEAABB16CFF8BFCC3FB001F84FB6C" box="[859,971,1160,1184]" italics="true" pageId="4" pageNumber="335">S. boninoi</emphasis>
</taxonomicName>
), obtuse posterior pronotal angles (in contrast to acute triangular porterior pronotal angles in
<taxonomicName id="4C3F4D3AB16CFF8BFB18FB4E1F42FB30" authorityName="Fanti &amp; M. G. Pankowski" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="boninoi">
<emphasis id="B94BEAABB16CFF8BFB18FB4E1F42FB30" italics="true" pageId="4" pageNumber="335">S. boninoi</emphasis>
</taxonomicName>
), larger pronotum, different shape of antennomere 1, and elytra wider at humeri.
<taxonomicName id="4C3F4D3AB16CFF8BFC8CFAAA1FBAFAF5" authorityName="Bukejs &amp; Fanti" authorityYear="2023" box="[788,1013,1314,1338]" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="carsteni" status="sp. nov.">
<emphasis id="B94BEAABB16CFF8BFC8CFAAA1F0DFAF6" box="[788,834,1314,1338]" italics="true" pageId="4" pageNumber="335">Silis</emphasis>
(
<emphasis id="B94BEAABB16CFF8BFCCAFAAA1FC5FAF5" box="[850,906,1314,1337]" italics="true" pageId="4" pageNumber="335">s. str.</emphasis>
)
<emphasis id="B94BEAABB16CFF8BFC01FAAA1FBAFAF5" box="[921,1013,1314,1337]" italics="true" pageId="4" pageNumber="335">carsteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27857D0B16CFF8BFC6DFAAA1806FAF5" box="[1013,1097,1314,1337]" pageId="4" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C3F4D3AB16CFF8BFD60FAC81EBEFABB" authority="Parisi &amp; Fanti, 2019" authorityName="Parisi &amp; Fanti" authorityYear="2019" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="lombardii">
<emphasis id="B94BEAABB16CFF8BFD60FAC81F63FA94" box="[760,812,1344,1368]" italics="true" pageId="4" pageNumber="335">Silis</emphasis>
(
<emphasis id="B94BEAABB16CFF8BFCAFFAC91F3EFA94" box="[823,881,1345,1368]" italics="true" pageId="4" pageNumber="335">s. str.</emphasis>
)
<emphasis id="B94BEAABB16CFF8BFC1AFAC81FA5FA94" box="[898,1002,1344,1368]" italics="true" pageId="4" pageNumber="335">lombardii</emphasis>
<bibRefCitation id="EFAE4B48B16CFF8BFC6BFAC81EBEFABB" author="Parisi F. &amp; Fanti F." pageId="4" pageNumber="335" pagination="119 - 123" refId="ref4358" refString="Parisi F., Fanti F. 2019. On the first Sili s Charpentier, 1825 from Baltic amber (Coleoptera, Cantharidae). Fossil Record 22 (2): 119 - 123." type="journal article" year="2019">Parisi &amp; Fanti, 2019</bibRefCitation>
</taxonomicName>
(Baltic amber; male) by the considerably smaller body size. A new species differs from Dominican amber species
<taxonomicName id="4C3F4D3AB16CFF8BFC8AFA331F36FA3E" authority="Fanti &amp; M. G. Pankowski, 2021" authorityName="Fanti &amp; M. G. Pankowski" authorityYear="2021" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="hegnai">
<emphasis id="B94BEAABB16CFF8BFC8AFA331F0FFA1F" box="[786,832,1467,1491]" italics="true" pageId="4" pageNumber="335">Silis</emphasis>
(
<emphasis id="B94BEAABB16CFF8BFCCBFA341FC1FA1F" box="[851,910,1468,1491]" italics="true" pageId="4" pageNumber="335">s. str.</emphasis>
)
<emphasis id="B94BEAABB16CFF8BFC38FA331FBFFA1F" box="[928,1008,1467,1491]" italics="true" pageId="4" pageNumber="335">hegnai</emphasis>
Fanti &amp; M. G. Pankowski, 2021
</taxonomicName>
and
<taxonomicName id="4C3F4D3AB16CFF8BFC2FFA52182CF9DC" authority="Fanti &amp; M. G. Pankowski, 2021" authorityName="Fanti &amp; M. G. Pankowski" authorityYear="2021" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="curleri">
<emphasis id="B94BEAABB16CFF8BFC2FFA521FAAFA3E" box="[951,997,1498,1522]" italics="true" pageId="4" pageNumber="335">Silis</emphasis>
(
<emphasis id="B94BEAABB16CFF8BFC61FA521878FA3D" box="[1017,1079,1498,1521]" italics="true" pageId="4" pageNumber="335">s. str.</emphasis>
)
<emphasis id="B94BEAABB16CFF8BFBD3FA5218DCFA3E" box="[1099,1171,1498,1522]" italics="true" pageId="4" pageNumber="335">curleri</emphasis>
Fanti &amp; M. G. Pankowski, 2021
</taxonomicName>
by distinctly smaller body size; and from
<taxonomicName id="4C3F4D3AB16CFF8BFBFEF99F18D5F981" authority="Wittmer, 1963" authorityName="Wittmer" authorityYear="1963" class="Insecta" family="Cantharidae" genus="Silis" higherTaxonomySource="GBIF" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="chiapasensis">
<emphasis id="B94BEAABB16CFF8BFBFEF99F18D5F9E3" box="[1126,1178,1559,1583]" italics="true" pageId="4" pageNumber="335">Silis</emphasis>
(
<emphasis id="B94BEAABB16CFF8BFD5CF9BE1F5BF981" box="[708,788,1590,1613]" italics="true" pageId="4" pageNumber="335">s. str.</emphasis>
)
<emphasis id="B94BEAABB16CFF8BFCA3F9BE1F8CF982" box="[827,963,1590,1614]" italics="true" pageId="4" pageNumber="335">chiapasensis</emphasis>
<bibRefCitation id="EFAE4B48B16CFF8BFC7AF9BD18D5F981" author="Wittmer W." box="[994,1178,1589,1613]" pageId="4" pageNumber="335" refId="ref4639" refString="Wittmer W. 1963. A new cantharid from the Chiapas amber of Mexico. University of California Publications in Entomology 31: 53, pl. 3 (p. 59)." type="journal volume" year="1963">Wittmer, 1963</bibRefCitation>
</taxonomicName>
(described from Mexican amber) by the
</paragraph>
<paragraph id="8B8036B9B16CFF8BFBF4F92E18DBF972" blockId="4.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="4" pageNumber="335">335</paragraph>
<paragraph id="8B8036B9B16DFF8AFFEAFF3A1D9EFF24" blockId="5.[114,593,178,232]" pageId="5" pageNumber="336">smaller body size and the longer elytra comparated to abdominal length.</paragraph>
</subSubSection>
<subSubSection id="C3256532B16DFF89FFEAFE851E42FD59" lastPageId="6" lastPageNumber="337" pageId="5" pageNumber="336" type="description">
<paragraph id="8B8036B9B16DFF8AFFEAFE851D11FDF5" blockId="5.[114,593,269,570]" pageId="5" pageNumber="336">
<emphasis id="B94BEAABB16DFF8AFFEAFE851CB3FEE9" bold="true" box="[114,252,269,293]" pageId="5" pageNumber="336">Description.</emphasis>
Body elongate-oval, flatened; sparsely covered with homogenous, short, semierect to erect setae; integument unicolorous dark brown (as preserved). Measurements: total body length
<quantity id="4CC79B5CB16DFF8AFE6AFE001E1EFE6C" box="[498,593,392,416]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="336" unit="mm" value="2.8">2.8 mm</quantity>
(from anterior margin of head to elytral apex); pronotum length
<quantity id="4CC79B5CB16DFF8AFEE5FE4D1D9BFE11" box="[381,468,453,477]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="336" unit="mm" value="0.5">0.5 mm</quantity>
, pronotum maximum width 1.0 mm; elytra length
<quantity id="4CC79B5CB16DFF8AFDB1FE6C1CD1FDD6" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="336" unit="mm" value="2.3">2.3 mm</quantity>
, elytra maximum width
<quantity id="4CC79B5CB16DFF8AFE33FD8B1E4EFDD6" box="[427,513,515,539]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="336" unit="mm" value="1.3">1.3 mm</quantity>
, elytra width at base 1.0 mm.
</paragraph>
<paragraph id="8B8036B9B16DFF8AFFEAFDD71D1AF9E1" blockId="5.[114,600,607,1581]" pageId="5" pageNumber="336">Head partially covered by pronotum, small, short, transverse, covered with fine punctation, distinctly narrower than anterior margin of pronotum. Compound eyes rather small, nearly hemispherical, strongly prominent, inserted in upper and lateral parts of head. Mandibles falciform, simple. Maxillary palpi 4-segmented, unequal in length, terminal palpomere securiform. Labial palps 3-segmented, terminal palpomere securiform. Antennae inserted far from from inner margin of eyes, relatively short, extending about to middle of elytra, 11-segmented, filiform, all antennomeres covered with short, semierect setae; scape longest, strongly elongate, about 3.0× longer than wide, not particularly robust, curved, dorso-apically enlarged and sligthly protruding forward; antennomere 2 shortest, cylindrical, about 0.6× as long as scape; antennomeres 35 equal in size and shape, subcylindrical, dilated apically, 2.4× as long as wide, robust, about 1.2× as long as antennomere 2; antennomeres 610 sub-equal in size and shape, subcylindrical, dilated apically, 2.5× as long as wide, slightly shorter than antennomere 5; antennomere XI elongate oval with acute and long apex, about 3.5× as long as wide; relative length ratios of antennomeres 11 equal to 9:5:6:6:6:5:5:5:5:5:7.</paragraph>
<paragraph id="8B8036B9B16DFF8AFFEAF92E1F58FD1F" blockId="5.[114,154,1702,1726]" lastBlockId="5.[624,1103,178,723]" pageId="5" pageNumber="336">336 Pronotum trapezoidal, strongly transverse, about 2.0× wider than long, widest in posterior one-fourth of its length, distinctly wider than head; punctation fine and sparse on disc and denser laterally; disc convex, with subtriangular impressions basally and suboval impressions anterolaterally; pronotal sides widely explanate. Pronotal anterior margin rounded medially; posterior margin slightly concave laterally and distinctly protruding medially forming wide lobe; lateral margins weakly rounded, non-incised, with two obtuse and small tooth (one at anterior angles and one near posterior margin). Anterior angles obtuse, rounded, slightly projecting; posterior angles obtuse. Prohypomera covered with fine punctation,</paragraph>
<paragraph id="8B8036B9B16DFF8AFDE8FD701E9AFB71" blockId="5.[624,1103,760,1213]" pageId="5" pageNumber="336">Scutellar shield subtrapezoidal, rounded apically, with fine punctation. Elytra at humeri wide and slightly narrower than pronotum, while in posterior one-third of elytral length wider than pronotum, elongate-oval, 1.8× longer than wide, subparallel-sided in anterior two-thirds and strongly widened in posterior one-third, rounded apically, completely covering abdomen; humeri distinct; humeral costae sharp, distinct in anterior two-thirds of elytral lenght elytral punctation small and rather dense, distance between punctures apparently lesser than diameter of one puncture.</paragraph>
<paragraph id="8B8036B9B16DFF8AFDE8FB6B1867FA9B" blockId="5.[624,1103,1251,1367]" pageId="5" pageNumber="336">Metaventrite wide, with rounded posterior margin, covered with fine punctation; disc convex. Metathoracic wings fully developed, completely covered by elytra.</paragraph>
<paragraph id="8B8036B9B16DFF89FDE8FAF51D36FE73" blockId="5.[624,1103,1404,1643]" lastBlockId="6.[189,668,178,447]" lastPageId="6" lastPageNumber="337" pageId="5" pageNumber="336">Legs rather long, slender, pubescent. Pro- and mesocoxae subconical, robust, metacoxae narrow, strongly transverse. Trochanters elongate with rounded apex. Femora slightly enlarged, flattened, slightly curved. Tibiae subcylindrical, with apical spur, nearly as long as femora. Tarsal formula 5-5-5; tarsomere 1 longest, elongate; tarsomere 2 slightly shorter than tarsomere 1; tarsomere 3 subtrapezoidal, slightly dilated apically, wider than previous ones, shorter than tarsomere 2; tarsomere 4 widest, deeply bilobed; tarsomere 5 subcylindrical, strongly elongate, slender, curved. Pretarsal claws simple, internal claws with obtuse and small basal tooth.</paragraph>
<paragraph id="8B8036B9B16EFF89FF25FE6C1E42FD59" blockId="6.[189,668,484,661]" pageId="6" pageNumber="337">Abdomen with six visible sternites (ventrites), covered with fine punctation; ultimate sternite wide, transverse, with emarginated posterior margin; ultimate tergite concave apically, longer and narrower than ultimate sternite.</paragraph>
</subSubSection>
<subSubSection id="C3256532B16EFF89FF25FD331D03FC81" pageId="6" pageNumber="337" type="etymology">
<paragraph id="8B8036B9B16EFF89FF25FD331D03FC81" blockId="6.[189,668,699,845]" pageId="6" pageNumber="337">
<emphasis id="B94BEAABB16EFF89FF25FD331D0AFD1F" bold="true" box="[189,325,699,723]" pageId="6" pageNumber="337">Etymology.</emphasis>
The specific epithet is a patronym; the new species is named in honor of Mr. Carsten Gröhn (Glinde,
<collectingCountry id="F3287629B16EFF89FF25FC9F1D6AFCE3" box="[189,293,791,815]" name="Germany" pageId="6" pageNumber="337">Germany</collectingCountry>
), an enthusiast and specialist in Baltic amber.
</paragraph>
</subSubSection>
<subSubSection id="C3256532B16EFF89FF25FCFB1ED2FB8F" pageId="6" pageNumber="337" type="discussion">
<paragraph id="8B8036B9B16EFF89FF25FCFB1ED2FB8F" blockId="6.[189,669,883,1091]" pageId="6" pageNumber="337">
<emphasis id="B94BEAABB16EFF89FF25FCFB1D67FC47" bold="true" box="[189,296,883,907]" pageId="6" pageNumber="337">Remark.</emphasis>
The lateral margins of pronotum without long lobes or teeth, the proportions of elytra (lenght/widht ratio), widely explanate elytral margins, relatively short anttenomeres, as well as ultimate abdominal sternite wide and emarginated apically indicate that the specimen is female.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,128 @@
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<mods:title id="E804DCC9E2FBF642EADE407DE7B52A34">NewNidella Gressit &amp; Rondon 1970 (Coleoptera: Cerambycidae) species from Palawan</mods:title>
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<paragraph id="8B8136C67A7AFFC8FDEE86C8FC2AFD09" blockId="1.[624,909,649,704]" box="[624,909,649,674]" pageId="1" pageNumber="260">
<heading id="D0C981AA7A7AFFC8FDEE86C8FC2AFD09" bold="true" box="[624,909,649,674]" fontSize="10" level="2" pageId="1" pageNumber="260" reason="4">
<emphasis id="B94AEAD47A7AFFC8FDEE86C8FC2AFD09" bold="true" box="[624,909,649,674]" pageId="1" pageNumber="260">
<taxonomicName id="4C3E4D457A7AFFC8FDEE86C8FC96FD0A" authorityName="Barševskis &amp; Zeltiņa &amp; Pepito" authorityYear="2023" box="[624,817,650,674]" class="Insecta" family="Cerambycidae" genus="Nidella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="260" phylum="Arthropoda" rank="species" species="palawana" status="sp. nov.">
<emphasis id="B94AEAD47A7AFFC8FDEE86C8FC96FD0A" bold="true" box="[624,817,650,674]" italics="true" pageId="1" pageNumber="260">Nidella palawana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27957AF7A7AFFC8FCA686CBFC2AFD09" box="[824,909,649,673]" pageId="1" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C324654D7A7AFFCBFDEE86EAFE03F996" lastPageId="2" lastPageNumber="261" pageId="1" pageNumber="260" type="description">
<paragraph id="8B8136C67A7AFFC8FDEE86EAFD18FD68" blockId="1.[624,909,649,704]" box="[624,703,680,704]" pageId="1" pageNumber="260">
(
<figureCitation id="13052A437A7AFFC8FDE686EAFD11FD68" box="[632,694,680,704]" captionStart="Fig" captionStartId="2.[189,228,930,954]" captionTargetBox="[196,643,171,924]" captionTargetId="figure-334@2.[189,659,171,924]" captionTargetPageId="2" captionText="Fig. 1.Nidella palawana sp. nov. Type specimen: Holotype: Female. /Philippines: Palawan Isl., / Roxas, 06.2023, / local collector leg./ [handwritten]; // HOLOTYPUS:/ Nidella / palawana sp. nov. / A.Barševskis, Z.Zeltiņa, / M.J.T. Pepito det. 2023 [red label, handwritten] (DUBC)." pageId="1" pageNumber="260">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="8B8136C67A7AFFC8FDEE86A7FD58FCB4" blockId="1.[624,1095,741,796]" pageId="1" pageNumber="260">
<emphasis id="B94AEAD47A7AFFC8FDEE86A7FCDDFD55" bold="true" box="[624,890,741,765]" pageId="1" pageNumber="260">General distribution:</emphasis>
<collectingCountry id="F32976567A7AFFC8FC1286A4FBAEFD56" box="[908,1033,742,766]" name="Philippines" pageId="1" pageNumber="260">Philippines</collectingCountry>
:
<collectingRegion id="49FAF8247A7AFFC8FBBA86A4FD17FCB4" country="Philippines" name="Palawan" pageId="1" pageNumber="260">Palawan</collectingRegion>
Island.
</paragraph>
<paragraph id="8B8136C67A7AFFC8FDEE8703FCB3FC7C" blockId="1.[624,1103,833,980]" pageId="1" pageNumber="260">
<emphasis id="B94AEAD47A7AFFC8FDEE8703FCA6FCF1" bold="true" box="[624,769,833,857]" pageId="1" pageNumber="260">Description.</emphasis>
Body elongated, narrow, subparallel, black. Elytra dorsally with six white spots, but pronotum laterally with white spot. Body lenght 13.0 mm, maximal width:
<quantity id="4CC69B237A7AFFC8FD2587FEFCB7FC7C" box="[699,784,956,980]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="260" unit="mm" value="1.2">1.2 mm</quantity>
.
</paragraph>
<paragraph id="8B8136C67A7AFFC8FDEE87B8FC6EF9B3" blockId="1.[624,1103,1018,1563]" pageId="1" pageNumber="260">Head flattened, with rectangular apical portion and convex, bilobate yellow-brown eyes. Dorsal surface of head with coarse punctures and sparse pubescence, especially along the raised margins, with short, longitudinal line between thick and extended antennal bases. Frontal portion of head vith elongated middle line. Cheeks covered with dense silvery-grey pubescence. Labrum brown, narrow, pubescent. Clypeus brown, transverse, shiny. Mandibles brown, with sharp apex, massive, wide, shiny, lateral sides with pubescence. Antennae slender, relatively long. Female antennae shorter than body, covered with fine punctures and fine, sparse pubescence; antennomeres 15 thickened apically, antennomeres 12 black, remaining antennomeres brown.</paragraph>
<paragraph id="8B8136C67A79FFCBFD2584F0FCCDFE17" blockId="2.[699,1178,178,447]" pageId="2" pageNumber="261">Elytra almost parallel-sided, in the middle narrowed, flattened dorsally, with distinct, slightly raised shoulders hump. Elytra black, matt, with weak reflection and microsculpture. Each elytron with three yellow spots. Punctation of elytra very dense, not arranged in longitudinal rows. Apical margins of elytra near suture with small, sharp extension.</paragraph>
<paragraph id="8B8136C67A79FFCBFD2585A6FBD5FD1C" blockId="2.[699,1179,484,692]" pageId="2" pageNumber="261">Ventro-lateral surface of body black, shinny, particularly covered with pubescence. Legs dichromatic, with yellow-brown or brown tibia and tarsus, and forelegs and middle legs with dark femora, while hind legs femora are dark with a yellow-brown base, covered with very fine pubescence.</paragraph>
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<paragraph id="8B8136C67A79FFCBFF2387E0FD87FC12" blockId="2.[189,668,930,1169]" box="[189,544,930,954]" pageId="2" pageNumber="261">
Fig. 1.
<taxonomicName id="4C3E4D457A79FFCBFE9787E0FE60FC12" authorityName="Barševskis &amp; Zeltiņa &amp; Pepito" authorityYear="2023" box="[265,455,930,954]" class="Insecta" family="Cerambycidae" genus="Nidella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="palawana" status="sp. nov.">
<emphasis id="B94AEAD47A79FFCBFE9787E0FE60FC12" box="[265,455,930,954]" italics="true" pageId="2" pageNumber="261">Nidella palawana</emphasis>
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<paragraph id="8B8136C67A79FFCBFF238782FE87FB39" blockId="2.[189,668,930,1169]" pageId="2" pageNumber="261">
<emphasis id="B94AEAD47A79FFCBFF238782FD83FC70" bold="true" box="[189,548,960,984]" pageId="2" pageNumber="261">Type specimen: Holotype:</emphasis>
Female.
<emphasis id="B94AEAD47A79FFCBFF23879DFF63FC5F" bold="true" box="[189,196,991,1015]" pageId="2" pageNumber="261">/</emphasis>
Philippines: Palawan Isl., / Roxas, 06.2023, / local collector leg./ [handwritten]; // HOLOTYPUS:/
<taxonomicName id="4C3E4D457A79FFCBFE59805FFDB2FB9D" authorityName="Gressit &amp; Rondon" authorityYear="1970" box="[455,533,1053,1077]" class="Insecta" family="Cerambycidae" genus="Nidella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="262" phylum="Arthropoda" rank="genus">Nidella</taxonomicName>
/ palawana
<taxonomicNameLabel id="A27957AF7A79FFCBFF238079FEB5FBFB" box="[189,274,1083,1107]" pageId="2" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
/ A.Barševskis, Z.Zeltiņa, / M.J.T. Pepito det. 2023 [red label, handwritten] (DUBC).
</paragraph>
</caption>
<paragraph id="8B8136C67A79FFCBFF2380F4FE2FFA2E" blockId="2.[189,668,1206,1414]" pageId="2" pageNumber="261">Pronotum black, cylindrical, elongated, neck-shaped narrowed in apical and basal portions, in the middle portion laterally with large triangular spot of silver-grey, dense pubescence. Pronotal disc without pubescence, with dense and coarse punctures and wrinkles.</paragraph>
<paragraph id="8B8136C67A79FFCBFF2381E9FE03F996" blockId="2.[189,669,1451,1598]" pageId="2" pageNumber="261">Scutellum small, with dense pubescence. Pars stridens partially visible because it is under the posterior edge of the pronotum, forming smooth spot, with very fine transverse microsculpture.</paragraph>
</subSubSection>
<subSubSection id="C324654D7A79FFCBFD25869BFC34FC02" pageId="2" pageNumber="261" type="diagnosis">
<paragraph id="8B8136C67A79FFCBFD25869BFC34FC02" blockId="2.[699,1179,729,938]" pageId="2" pageNumber="261">
<emphasis id="B94AEAD47A79FFCBFD25869BFC6CFD59" bold="true" box="[699,971,729,753]" pageId="2" pageNumber="261">Differential diagnosis.</emphasis>
Regarding the shape of the body, the new species differs from all other species of this genus by six yellow spots on elytra and triangular lateral spot in the middle of pronotum. No other species of this genus known so far have such characteristics.
</paragraph>
</subSubSection>
<subSubSection id="C324654D7A79FFCBFD25878CFC94FB8C" pageId="2" pageNumber="261" type="etymology">
<paragraph id="8B8136C67A79FFCBFD25878CFC94FB8C" blockId="2.[699,1178,974,1060]" pageId="2" pageNumber="261">
<emphasis id="B94AEAD47A79FFCBFD25878CFC9BFC4E" bold="true" box="[699,828,974,998]" pageId="2" pageNumber="261">Etymology.</emphasis>
This species is named after the
<collectingRegion id="49FAF8247A79FFCBFD2587AFFCB2FBAD" box="[699,789,1005,1029]" country="Philippines" name="Palawan" pageId="2" pageNumber="261">Palawan</collectingRegion>
island, where it occurs [
<collectingRegion id="49FAF8247A79FFCBFBBA87AFFB26FBAD" box="[1060,1153,1005,1029]" country="Philippines" name="Palawan" pageId="2" pageNumber="261">Palawan</collectingRegion>
-
<taxonomicName id="4C3E4D457A79FFCBFD25804EFC83FB8C" authorityName="Barševskis &amp; Zeltiņa &amp; Pepito" authorityYear="2023" box="[699,804,1036,1060]" class="Insecta" family="Cerambycidae" genus="Nidella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="palawana">
<emphasis id="B94AEAD47A79FFCBFD25804EFC83FB8C" box="[699,804,1036,1060]" italics="true" pageId="2" pageNumber="261">palawana</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:role id="B97AD595997F00227932B33A64D0C9DE">
<mods:roleTerm id="FBB70E4909C5E0EA469D77475183D23E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C088FF8EE8C4A70AFF38A9DDCC8676B2">Wentzel, Danuta</mods:namePart>
</mods:name>
<mods:name id="0D61DC00E7C96FF6354C947A9AE31869" type="personal">
<mods:role id="D94993826A2129E127E4A6A196F1207E">
<mods:roleTerm id="CF851CAA0F0A4EEAC8594C60EE22525A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EBFA75E9F2EE823234B8FD314DD088FD">Bonino, Enrico</mods:namePart>
</mods:name>
<mods:name id="FAB39D7948DE9B3477CD4AF58C04B50D" type="personal">
<mods:role id="E47FBA87675E38AD03B2281C378512BE">
<mods:roleTerm id="5AA2D54CE5EDE9FE4817B33FE09E97F6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3F7C6E7F2E65ADAF9C195B5DA768C41">Fanti, Fabrizio</mods:namePart>
</mods:name>
<mods:typeOfResource id="860C7D419BA4B28F1CDE69F1AAE6156E">text</mods:typeOfResource>
<mods:relatedItem id="45174A495AFE8564A835ADAD5D1744F2" type="host">
<mods:titleInfo id="7DC7B940E48C4BB9A3557B357170DB6F">
<mods:title id="AC4C5005CD4B4FF3F638A453338C88ED">Baltic Journal of Coleopterology</mods:title>
</mods:titleInfo>
<mods:part id="8C28B57564866961BD16D0ADD545E9A9">
<mods:date id="9C97F166CA2BA79BD748F097CC02E099">2022</mods:date>
<mods:detail id="FDA5DDC711E747179480ACAC93E839E8" type="volume">
<mods:number id="4A3B6BF10CAAE4E2D7B979B62A95B120">22</mods:number>
</mods:detail>
<mods:detail id="8BC214362480ECC7C3999FA249768FCC" type="issue">
<mods:number id="A422A12990D2AA540837D1A2A5F11AAD">2</mods:number>
</mods:detail>
<mods:extent id="93388B3D0B803A783671ECB443F1ECDF" unit="page">
<mods:start id="CD0E44BB1967D8B4114D111835853189">313</mods:start>
<mods:end id="6E6B2917EB43A822B20D9293F8A965EF">318</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7D8B3E2FD60425B977F22A6C0264EF04">journal article</mods:classification>
<mods:identifier id="8D1899066400E69B28D56D85ECD156CD" type="ISSN">1407-8619</mods:identifier>
</mods:mods>
<treatment id="039887EDFF876F63FFB2FB5A5D56FCC2" LSID="urn:lsid:plazi:treatment:039887EDFF876F63FFB2FB5A5D56FCC2" httpUri="http://treatment.plazi.org/id/039887EDFF876F63FFB2FB5A5D56FCC2" lastPageId="3" lastPageNumber="316" pageId="1" pageNumber="314">
<subSubSection id="C32B6570FF876F61FFB2FB5A5C52FB2E" box="[113,590,1225,1249]" pageId="1" pageNumber="314" type="nomenclature">
<paragraph id="8B8E36FBFF876F61FFB2FB5A5C52FB2E" blockId="1.[113,590,1225,1280]" box="[113,590,1225,1249]" pageId="1" pageNumber="314">
<heading id="D0C68197FF876F61FFB2FB5A5C52FB2E" bold="true" box="[113,590,1225,1249]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="314" reason="4">
<emphasis id="B945EAE9FF876F61FFB2FB5A5C52FB2E" bold="true" box="[113,590,1225,1249]" pageId="1" pageNumber="314">
<taxonomicName id="4C314D78FF876F61FFB2FB5A5FEEFB2E" authorityName="Wentzel &amp; Bonino &amp; Fanti" authorityYear="2022" box="[113,498,1225,1249]" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="314" phylum="Arthropoda" rank="species" species="thomasiwentzeli" status="sp. nov.">
<emphasis id="B945EAE9FF876F61FFB2FB5A5FEEFB2E" bold="true" box="[113,498,1225,1249]" italics="true" pageId="1" pageNumber="314">Cacomorphocerus thomasiwentzeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765792FF876F61FE3BFB5A5C52FB2E" box="[504,590,1225,1249]" pageId="1" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B6570FF876F61FFB2FB7A5EFAFACF" box="[113,230,1257,1280]" pageId="1" pageNumber="314" type="description">
<paragraph id="8B8E36FBFF876F61FFB2FB7A5EFAFACF" blockId="1.[113,590,1225,1280]" box="[113,230,1257,1280]" pageId="1" pageNumber="314">
(
<figureCitation id="130A2A7EFF876F61FFBAFB7A5EC2FACF" box="[121,222,1257,1280]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[151,190,1546,1569]" captionStartId-1="3.[113,152,1665,1688]" captionTargetBox-0="[151,1172,151,1512]" captionTargetBox-1="[113,1134,1211,1631]" captionTargetId-0="figure-14@2.[151,1172,151,1512]" captionTargetId-1="figure-590@3.[113,1134,1211,1631]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Cacomorphocerus thomasiwentzeli sp. nov. in Baltic amber. A: Holotype, lateral view, bar = 1.0 mm; B: Holotype, detail of last abdominal segments, bar = 0.5 mm." captionText-1="Fig. 2. Cacomorphocerus thomasiwentzeli sp. nov. in Baltic amber. Reconstruction of antenna." pageId="1" pageNumber="314">Figs. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32B6570FF876F61FFB2FA8C5EF4F9EB" pageId="1" pageNumber="314" type="materials_examined">
<paragraph id="8B8E36FBFF876F61FFB2FA8C5C2DFAB8" blockId="1.[113,604,1311,1399]" pageId="1" pageNumber="314">
<materialsCitation id="3B593CA6FF876F61FFB2FA8C5C2DFAB8" country="Germany" county="Ennepetal" location="Male" municipality="Danuta Wentzel" pageId="1" pageNumber="314" specimenCount="1" typeStatus="holotype">
<emphasis id="B945EAE9FF876F61FFB2FA8C5EC2FAF8" bold="true" box="[113,222,1311,1335]" pageId="1" pageNumber="314">
<typeStatus id="548A8859FF876F61FFB2FA8C5EC4FAF8" box="[113,216,1311,1335]" pageId="1" pageNumber="314" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EEE6020FF876F61FF22FAB35F05FAF8" LSID="urn:lsid:plazi:treatment:039887EDFF876F63FFB2FB5A5D56FCC2:8EEE6020FF876F61FF22FAB35F05FAF8" box="[225,281,1312,1335]" country="Germany" county="Ennepetal" municipality="Danuta Wentzel" name="Male" pageId="1" pageNumber="314">Male</location>
, adult specimen in
<location id="8EEE6020FF876F61FE1AFAB35C0BFAF8" LSID="urn:lsid:plazi:treatment:039887EDFF876F63FFB2FB5A5D56FCC2:8EEE6020FF876F61FE1AFAB35C0BFAF8" box="[473,535,1312,1335]" country="Germany" county="Ennepetal" municipality="Danuta Wentzel" name="Baltic" pageId="1" pageNumber="314">Baltic</location>
amber piece, with the
<location id="8EEE6020FF876F61FEC5FAD35F4AFA98" LSID="urn:lsid:plazi:treatment:039887EDFF876F63FFB2FB5A5D56FCC2:8EEE6020FF876F61FEC5FAD35F4AFA98" box="[262,342,1344,1367]" country="Germany" county="Ennepetal" municipality="Danuta Wentzel" name="Catalog" pageId="1" pageNumber="314">Catalog</location>
number
<quantity id="4CC99B1EFF876F61FE6FFAD35FF7FA98" box="[428,491,1344,1367]" metricMagnitude="1" metricUnit="m" metricValue="1.1480799999999998" pageId="1" pageNumber="314" unit="in" value="452.0">452 in</quantity>
the
<collectingMunicipality id="6BEAAC81FF876F61FDD1FAD35ED4FAB8" pageId="1" pageNumber="314">Danuta Wentzel</collectingMunicipality>
collection (
<collectingCounty id="62EF4E77FF876F61FE86FAF35FA8FAB8" box="[325,436,1376,1399]" pageId="1" pageNumber="314">Ennepetal</collectingCounty>
,
<collectingCountry id="F326766BFF876F61FE7DFAF35C3AFAB8" box="[446,550,1376,1399]" name="Germany" pageId="1" pageNumber="314">Germany</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8E36FBFF876F61FFB2FA055C2FFA62" blockId="1.[113,563,1430,1454]" box="[113,563,1430,1454]" pageId="1" pageNumber="314">
<emphasis id="B945EAE9FF876F61FFB2FA055F14FA61" bold="true" box="[113,264,1430,1454]" pageId="1" pageNumber="314">Type locality.</emphasis>
<collectingRegion id="49F5F819FF876F61FECDFA055FC6FA62" box="[270,474,1430,1453]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="314">Kaliningrad region</collectingRegion>
,
<collectingCountry id="F326766BFF876F61FE27FA055C33FA62" box="[484,559,1430,1453]" name="Russia" pageId="1" pageNumber="314">Russia</collectingCountry>
.
</paragraph>
<paragraph id="8B8E36FBFF876F61FFB2FA5E5EF4F9EB" blockId="1.[113,605,1485,1572]" pageId="1" pageNumber="314">
<emphasis id="B945EAE9FF876F61FFB2FA5E5F17FA2A" bold="true" box="[113,267,1485,1509]" pageId="1" pageNumber="314">Type horizon.</emphasis>
Middle Eocene: Lutetian (47.8 41.2 Mya) to late Eocene: Priabonian (37.8 33.9 Mya).
</paragraph>
</subSubSection>
<subSubSection id="C32B6570FF876F61FFB2F9D75DB3FEDB" pageId="1" pageNumber="314" type="diagnosis">
<paragraph id="8B8E36FBFF876F61FFB2F9D75C41F954" blockId="1.[113,605,1604,1691]" pageId="1" pageNumber="314">
<emphasis id="B945EAE9FF876F61FFB2F9D75F7AF993" bold="true" box="[113,358,1604,1628]" pageId="1" pageNumber="314">Differential diagnosis.</emphasis>
The most similar
<taxonomicName id="4C314D78FF876F61FDDEF9D75FABF9B4" authority="Schaufuss, 1892" authorityName="Schaufuss" authorityYear="1892" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAE9FF876F61FDDEF9D75EE2F9B4" italics="true" pageId="1" pageNumber="314">Cacomorphocerus</emphasis>
<bibRefCitation id="EFA04B0AFF876F61FEC4F9F75FABF9B4" author="Schaufuss C. F. C." box="[263,439,1636,1659]" pageId="1" pageNumber="314" pagination="53 - 64" refId="ref2626" refString="Schaufuss C. F. C. 1892. Preussens Bernstein- Kafer. Neue Formen aus der Helm'schen Sammlung im Danziger Provinzialmuseum. Berliner Entomologische Zeitschrift 36 (I): 53 - 64. [1891]" type="book chapter" year="1892">Schaufuss, 1892</bibRefCitation>
</taxonomicName>
with 12 antennomeres is
<taxonomicName id="4C314D78FF876F61FF3BF9175C41F954" box="[248,605,1668,1691]" pageId="1" pageNumber="314">
<emphasis id="B945EAE9FF876F61FF3BF9175C41F954" box="[248,605,1668,1691]" italics="true" pageId="1" pageNumber="314">Cacomorphocerus wiszniewskii</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8B8E36FBFF876F61FD40FF085DB3FEDB" blockId="1.[643,1134,155,276]" pageId="1" pageNumber="314">
<bibRefCitation id="EFA04B0AFF876F61FD40FF085DDFFF7D" author="Fanti F. &amp; Kupryjanowicz J." box="[643,963,155,178]" pageId="1" pageNumber="314" pagination="149 - 153" refId="ref2118" refString="Fanti F., Kupryjanowicz J. 2018. Discovery of a new fossil soldier beetle in Eocene Baltic amber, with the establishment of the new tribe Cacomorphocerini. Annales de Paleontologie 104: 149 - 153." type="journal article" year="2018">Fanti &amp; Kupryjanowicz, 2018</bibRefCitation>
from which the new species differs for the antennomere VIII longer and more curved and antennomere IX more transverse and thinner.
</paragraph>
</subSubSection>
<subSubSection id="C32B6570FF876F63FD40FEA75C32FC9D" lastPageId="3" lastPageNumber="316" pageId="1" pageNumber="314" type="description">
<paragraph id="8B8E36FBFF876F61FD40FEA75A73FE21" blockId="1.[643,1135,308,494]" pageId="1" pageNumber="314">
<emphasis id="B945EAE9FF876F61FD40FEA75D12FE83" bold="true" box="[643,782,308,332]" pageId="1" pageNumber="314">Description.</emphasis>
Adult, alate. Male, based on last sternite extremely small and transverse, and based on the presence of long appendage which is probably part of the aedeagus. Body length:
<quantity id="4CC99B1EFF876F61FD40FE255CC8FE02" box="[643,724,438,461]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="314" unit="mm" value="6.2">6.2 mm</quantity>
(the body is curled up and in life probably reached about
<quantity id="4CC99B1EFF876F61FC92FE445D93FE21" box="[849,911,471,494]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="314" unit="mm" value="7.0">7 mm</quantity>
). Entirely dark brown.
</paragraph>
<paragraph id="8B8E36FBFF876F63FD40FD9D5C32FC9D" blockId="1.[643,1135,526,1687]" lastBlockId="3.[113,605,155,850]" lastPageId="3" lastPageNumber="316" pageId="1" pageNumber="314">Head rounded at sides, short and slightly transverse, as large as pronotum, equipped with long setae and shallow punctuation. Eyes rounded, convex and very prominent, inserted laterally to the head in the upper part. Mandibles elongated, falciform, apparently without tooth. Maxillary palps 4­ segmented with the last palpomere strongly securiform. Labial palps 3­ segmented. Antennae pubescent equipped with long setae, surpassing the half of elytra and almost reaching the apex, 12­ segmented, inserted almost close to the eyes; scape elongated, stout, not clubshaped; antennomere II short, narrower near the base, about 2.6 times shorter than scape; antennomeres IIIIV longer than previous one, stout and slightly enlarged at apex; antennomeres V VI slightly sturdier and shorter than previous ones, triangular shaped; antennomere VII slightly saucer­shaped and slightly asymmetrical dilated, more transverse than previous ones; antennomere VIII asymmetric, rather elongated, strongly curved and deformed, apex concave; antennomere IX asymmetric, strongly saucer­shaped, thin, strongly transverse; antennomeres XXII very elongated and filiform, antennomere XI shorter than X and last antennomere which is the longest and has a rounded apex. Pronotum sub­rectangular that does not cover the head, longer than wide, sides straight and finely bordered, anterior margin straight and finely bordered, posterior margin very slightly undulate, surface bulged on the disc equipped with few and very long setae and without punctuation. Scutellum triangular­shaped. Elytra long which covers and surpasses the last abdominal segments, parallel sided and with rounded apex, slightly wider than pronotum, surface with small punctuation and with long setae. Posterior wings semi­transparent, longer than elytra. Sternum elongated and pubescent, ventrites transverse and pubescent, last ventrite apparently very short and transverse, last tergite very broad and emarginate at apex, probably part of edeagus visible with a long and strongly pubescent appendix which is very thin at base and enlarged and rounded at apex. Legs slender, long and pubescent; coxae wide, subsquare; trochanters elongated; femurs cylindrical, straight, sturdier than tibiae; tibiae slender, cylindrical, with an apical spur which is curved and robust, pro­ and mesotibiae shorter than pro­ and mesofemora, metatibiae slightly longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere elongated, slightly shorter than the second and third together; second elongated; third tarsomere almost triangular, not bilobed at sides; fourth tarsomere strongly bilobed; fifth tarsomere thin and very elongated; claws simple with an obtuse tooth at base. Female unknown.</paragraph>
</subSubSection>
<caption id="DF4E6673FF846F62FF54F9995DD7F98E" pageId="2" pageNumber="315" startId="2.[151,190,1546,1569]" targetBox="[151,1172,151,1512]" targetPageId="2" targetType="figure">
<paragraph id="8B8E36FBFF846F62FF54F9995DD7F98E" blockId="2.[151,1172,1546,1601]" pageId="2" pageNumber="315">
Fig. 1.
<taxonomicName id="4C314D78FF846F62FF24F9995C7CF9EE" authorityName="Wentzel &amp; Bonino &amp; Fanti" authorityYear="2022" box="[231,608,1546,1569]" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="thomasiwentzeli" status="sp. nov.">
<emphasis id="B945EAE9FF846F62FF24F9995C7CF9EE" box="[231,608,1546,1569]" italics="true" pageId="2" pageNumber="315">Cacomorphocerus thomasiwentzeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765792FF846F62FDAAF9995CA1F9EE" box="[617,701,1546,1569]" pageId="2" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
in Baltic amber. A: Holotype, lateral view, bar = 1.0 mm; B: Holotype, detail of last abdominal segments, bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C32B6570FF856F63FFB2FCE25CEBFDFA" pageId="3" pageNumber="316" type="etymology">
<paragraph id="8B8E36FBFF856F63FFB2FCE25C4EFC67" blockId="3.[113,605,881,936]" pageId="3" pageNumber="316">
<emphasis id="B945EAE9FF856F63FFB2FCE25EF1FC46" bold="true" box="[113,237,881,905]" pageId="3" pageNumber="316">Etymology.</emphasis>
Named in honor of Thomas Wentzel, husband of the first author (Danuta Wentzel).
</paragraph>
<paragraph id="8B8E36FBFF856F63FFB2FC5B5F00FC30" blockId="3.[113,605,968,1023]" pageId="3" pageNumber="316">
<emphasis id="B945EAE9FF856F63FFB2FC5B5F0CFC2F" bold="true" box="[113,272,968,992]" pageId="3" pageNumber="316">Syninclusions.</emphasis>
Stellate hairs, botanical remains and air bubbles.
</paragraph>
<paragraph id="8B8E36FBFF856F63FFB2FB8D5CEBFDFA" blockId="3.[113,605,1054,1174]" lastBlockId="3.[643,1135,155,565]" pageId="3" pageNumber="316">
<emphasis id="B945EAE9FF856F63FFB2FB8D5F70FBF9" bold="true" box="[113,364,1054,1078]" pageId="3" pageNumber="316">Systematic placement.</emphasis>
The last maxillary palpomere securiform, tarsal formula 5­ 5­ 5, elongated elytra, pronotum without lobes and teeth at sides, and the antennae 12­ segmented with some antennomeres saucer­shaped, unequivocally make the new species belong to the subfamily
<taxonomicName id="4C314D78FF856F63FD08FF4F5DCFFF3C" authority="Imhoff, 1856" authorityName="Imhoff" authorityYear="1856" box="[715,979,220,243]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="subFamily" subFamily="Cantharinae">Cantharinae Imhoff, 1856</taxonomicName>
and to the tribe
<taxonomicName id="4C314D78FF856F63FD40FF6F5CABFEFC" authority="Fanti &amp; Kupryjanowicz, 2018" authorityName="Fanti &amp; Kupryjanowicz" authorityYear="2018" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="tribe" tribe="Cacomorphocerini">
Cacomorphocerini
<bibRefCitation id="EFA04B0AFF856F63FC99FF6F5CABFEFC" author="Fanti F. &amp; Kupryjanowicz J." pageId="3" pageNumber="316" pagination="149 - 153" refId="ref2118" refString="Fanti F., Kupryjanowicz J. 2018. Discovery of a new fossil soldier beetle in Eocene Baltic amber, with the establishment of the new tribe Cacomorphocerini. Annales de Paleontologie 104: 149 - 153." type="journal article" year="2018">Fanti &amp; Kupryjanowicz, 2018</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFA04B0AFF856F63FD01FE8F5D75FEFC" author="Brancucci M." box="[706,873,284,307]" pageId="3" pageNumber="316" pagination="215 - 388" refId="ref1892" refString="Brancucci M. 1980. Morphologie comparee, evolution et systematique des Cantharidae (Insecta: Coleoptera). Entomologica Basiliensia 5: 215 - 388." type="journal article" year="1980">Brancucci 1980</bibRefCitation>
; Ramsdale 2002;
<bibRefCitation id="EFA04B0AFF856F63FBDEFE8F5D7CFE9C" author="Fanti F. &amp; Kupryjanowicz J." pageId="3" pageNumber="316" pagination="149 - 153" refId="ref2118" refString="Fanti F., Kupryjanowicz J. 2018. Discovery of a new fossil soldier beetle in Eocene Baltic amber, with the establishment of the new tribe Cacomorphocerini. Annales de Paleontologie 104: 149 - 153." type="journal article" year="2018">Fanti &amp; Kupryjanowicz 2018</bibRefCitation>
;
<bibRefCitation id="EFA04B0AFF856F63FCABFEAF5A76FE9C" author="Fanti F. &amp; Pankowski M. K." box="[872,1130,316,339]" pageId="3" pageNumber="316" pagination="589 - 595" refId="ref2198" refString="Fanti F., Pankowski M. K. 2019. A new soldier beetle of the extinct tribe Cacomorphocerini Fanti &amp; Kupryjanowicz, 2018. Zootaxa 4651 (3): 589 - 595." type="journal article" year="2019">Fanti &amp; Pankowski 2019</bibRefCitation>
;
<bibRefCitation id="EFA04B0AFF856F63FD40FECE5D7DFEBB" author="Poinar G. O. Jr. &amp; Fanti F." box="[643,865,349,372]" pageId="3" pageNumber="316" pagination="99 - 105" refId="ref2509" refString="Poinar G. O. Jr., Fanti F. 2019. A new fossil soldier beetle (Coleoptera: Cantharidae) of the genus Cacomorphocerus Schaufuss, 1892 from Baltic amber. Palaeodiversity 12 (1): 99 - 105. Ramsdale A. S. 2002. Family 64. Cantharidae Imhoff 1856. In: Arnett R. H. Jr., Thomas M. C., Skelley P. E., Frank J. H. (Eds.), American Beetles. Vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press, Boca Raton, Florida, pp. 202 - 218." type="book chapter" year="2019">Poinar &amp; Fanti 2019</bibRefCitation>
;
<bibRefCitation id="EFA04B0AFF856F63FCAFFECE5CA1FE5B" author="Kazantsev S. V. &amp; Perkovsky E. E." pageId="3" pageNumber="316" pagination="395 - 400" refId="ref2350" refString="Kazantsev S. V., Perkovsky E. E. 2020. The first Cacomorphocerus species (Coleoptera, Cantharidae) from Rovno amber: a second species of the genus with 11 - segmented antennae. Zootaxa 4751 (2): 395 - 400." type="journal article" year="2020">Kazantsev &amp; Perkovsky 2020</bibRefCitation>
). The new species belongs to the genus
<taxonomicName id="4C314D78FF856F63FD40FE0E5A1FFE7B" authority="Schaufuss, 1892" authorityName="Schaufuss" authorityYear="1892" box="[643,1027,413,436]" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAE9FF856F63FD40FE0E5D54FE7B" box="[643,840,413,436]" italics="true" pageId="3" pageNumber="316">Cacomorphocerus</emphasis>
<bibRefCitation id="EFA04B0AFF856F63FC92FE0E5A1FFE7B" author="Schaufuss C. F. C." box="[849,1027,413,436]" pageId="3" pageNumber="316" pagination="53 - 64" refId="ref2626" refString="Schaufuss C. F. C. 1892. Preussens Bernstein- Kafer. Neue Formen aus der Helm'schen Sammlung im Danziger Provinzialmuseum. Berliner Entomologische Zeitschrift 36 (I): 53 - 64. [1891]" type="book chapter" year="1892">Schaufuss, 1892</bibRefCitation>
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based on its 12 antennomeres with some of these saucer­shaped and antennomeres 1012 filiform, tarsomere 3 straight and not bilobed (
<bibRefCitation id="EFA04B0AFF856F63FBCAFE6D5CF7FDFA" author="Bukejs A. &amp; Fanti F. &amp; McKellar R. C." pageId="3" pageNumber="316" pagination="1 - 7" refId="ref1978" refString="Bukejs A., Fanti F., McKellar R. C. 2019 b. A new species of Cacomorphocerus Schaufuss, 1892 (Coleoptera: Cantharidae) from Baltic amber with a key to known species. Palaeontologia Electronica 22.2.28 A: 1 - 7." type="book chapter" year="2019">Bukejs et al. 2019b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B6570FF856F63FD40FDC65D56FCC2" pageId="3" pageNumber="316" type="discussion">
<paragraph id="8B8E36FBFF856F63FD40FDC65D56FCC2" blockId="3.[643,1134,597,781]" pageId="3" pageNumber="316">
<emphasis id="B945EAE9FF856F63FD40FDC65CEDFDA2" bold="true" box="[643,753,597,621]" pageId="3" pageNumber="316">Remarks.</emphasis>
Piece of amber that measures 1.88x
<quantity id="4CC99B1EFF856F63FD40FDE65CCCFD43" box="[643,720,629,652]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="316" unit="cm" value="1.2">1.2 cm</quantity>
. The inclusion is complete, except for the right metathoracic leg preserved up to part of the third tarsomere. The left side is poorly visible, and therefore also the last antennomeres of the left antenna.
</paragraph>
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</treatment>
</document>

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@ -0,0 +1,73 @@
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Tribe
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<emphasis id="B944EAD6391EFFF0FFD6FB63FDF2C40B" bold="true" box="[114,518,1259,1283]" pageId="1">Subtrribe Auletorhinina Voss, 1935</emphasis>
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Genus
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Voss, 1922
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Subgenus
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View file

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<emphasis id="B940792E3817FFD3FDA73B5AFD605000" bold="true" box="[624,759,576,600]" italics="true" pageId="1" pageNumber="228">Globicornis</emphasis>
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<emphasis id="B940792E3817FFD3FDA73B45FD50502F" bold="true" box="[624,711,607,631]" pageId="1" pageNumber="228">sp. nov.</emphasis>
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(
<figureCitation id="130FB9B93817FFD3FDAF3B67FD4A50CE" box="[632,733,637,662]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[114,153,1028,1052]" captionStartId-1="3.[114,153,1546,1570]" captionTargetBox-0="[177,1064,338,1022]" captionTargetBox-1="[215,987,1114,1539]" captionTargetId-0="figure-96@3.[177,1064,338,1022]" captionTargetId-1="figure-99@3.[215,987,1114,1539]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionText-0="Fig. 1. Globicornis samlandensis sp. nov., holotype, No RSKM_PAL_A66 [RSM]: A habitus, dorsal view; B habitus, ventral view." captionText-1="Fig. 2.Globicornis samlandensis sp. nov., holotype, No RSKM_PAL_A66 [RSM], details of forebody, ventral view. Abbreviations: a7a10 antennomeres 710 respectively; ey compound eye; pc prosternal “collar“." pageId="1" pageNumber="228">Figs. 12</figureCitation>
)
</paragraph>
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<emphasis id="B940792E3817FFD3FDA73BA1FCEF508B" bold="true" box="[624,888,698,723]" pageId="1" pageNumber="228">Type material.</emphasis>
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:
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_PAL_A66 [
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], (ex coll.
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JDC-10352); adult, sex unknown.
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almost complete beetle (with left metatarsus missing) is included in a transparent, yellow amber piece with dimensions of 21×
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and a maximum thickness of
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; preserved without supplementary fixation.
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: one
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(
<collectorName id="26C1C0EA3817FFD3FDA03AD5FD5C51BF" box="[631,715,975,999]" pageId="1" pageNumber="228">Diptera</collectorName>
) specimen, and few small stellate
<taxonomicName id="4C34DEBF3817FFD3FDA73AF4FD44565E" authorityName="Dumortier" authorityYear="1829" box="[624,723,1006,1030]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="228" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
trichomes.
</materialsCitation>
</paragraph>
<paragraph id="8B8BA53C3817FFD3FDA73D31FCC356C7" blockId="1.[624,1103,1067,1183]" pageId="1" pageNumber="228">
<emphasis id="B940792E3817FFD3FDA73D31FCB6561B" bold="true" box="[624,801,1067,1091]" pageId="1" pageNumber="228">
<typeStatus id="548F1B9E3817FFD3FDA73D31FD3D561B" box="[624,682,1067,1091]" pageId="1" pageNumber="228">Type</typeStatus>
stratum.
</emphasis>
Baltic amber; MiddleUpper Eocene (
<bibRefCitation id="EFA5D8CD3817FFD3FCCE3D53FC685639" author="Sadowski E. - M. &amp; Seyfullah L. J. &amp; Schmidt A. R. &amp; Kunzmann L." box="[793,1023,1097,1121]" pageId="1" pageNumber="228" pagination="1 - 73" refId="ref2778" refString="Sadowski E. - M., Seyfullah L. J., Schmidt A. R., Kunzmann L. 2017. Conifers of the ' Baltic amber forest' and their palaeoecological significance. Stapfia 106: 1 - 73." type="journal article" year="2017">Sadowski et al. 2017</bibRefCitation>
,
<bibRefCitation id="EFA5D8CD3817FFD3FBDB3D53FBDC5639" author="Sadowski E. - M. &amp; Schmidt A. R. &amp; Denk T." box="[1036,1099,1097,1121]" pageId="1" pageNumber="228" pagination="405 - 517" refId="ref2821" refString="Sadowski E. - M., Schmidt A. R., Denk T. 2020. Staminate inflorescences with in situ pollen from Eocene Baltic amber reveal high diversity in Fagaceae (oak family). Willdenowia: 50: 405 - 517." type="journal article" year="2020">2020</bibRefCitation>
;
<bibRefCitation id="EFA5D8CD3817FFD3FDA73D72FCF756D8" author="Seyfullah L. J. &amp; Beimforde C. &amp; Dal Corso J. &amp; Perrichot V. &amp; Rikkinen J. &amp; Schmidt A. R." box="[624,864,1128,1152]" pageId="1" pageNumber="228" pagination="1684 - 1714" refId="ref2867" refString="Seyfullah L. J., Beimforde C., Dal Corso J., Perrichot V., Rikkinen J., Schmidt A. R. 2018. Production and preservation of resins - past and present. Biological Reviews 93: 1684 - 1714." type="journal article" year="2018">Seyfullah et al. 2018</bibRefCitation>
;
<bibRefCitation id="EFA5D8CD3817FFD3FCA73D72FBDC56D8" author="Bukejs A &amp; Alekseev VI &amp; Pollock DA" box="[880,1099,1128,1152]" pageId="1" pageNumber="228" pagination="261 - 273" refId="ref2303" refString="Bukejs A, Alekseev VI, Pollock DA. 2019. Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa 4664 (2): 261 - 273." type="journal article" year="2019">Bukejs et al. 2019</bibRefCitation>
;
<bibRefCitation id="EFA5D8CD3817FFD3FDA73D9DFCDF56C7" author="Kasinski J. R. &amp; Kramarska R. &amp; Slodkowska B. &amp; Sivkov V. &amp; Piwocki M." box="[624,840,1159,1183]" pageId="1" pageNumber="228" pagination="29 - 53" refId="ref2694" refString="Kasinski J. R., Kramarska R., Slodkowska B., Sivkov V., Piwocki M. 2020. Paleocene and Eocene deposits on the eastern margin of the Gulf of Gdansk (Yantarny P- 1 bore hole, Kaliningrad region, Russia). Geological Quarterly 64: 29 - 53." type="journal article" year="2020">Kasiński et al. 2020</bibRefCitation>
).
</paragraph>
<paragraph id="8B8BA53C3817FFD3FDA73DDEFC8F5760" blockId="1.[624,1103,1219,1336]" pageId="1" pageNumber="228">
<materialsCitation id="3B5CAF613817FFD3FDA73DDEFC835760" country="Russia" location="Yantarny" municipality="Sambian" pageId="1" pageNumber="228" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B940792E3817FFD3FDA73DDEFCBD5684" bold="true" box="[624,810,1219,1244]" pageId="1" pageNumber="228">
<typeStatus id="548F1B9E3817FFD3FDA73DDEFD3D5684" box="[624,682,1220,1244]" pageId="1" pageNumber="228">Type</typeStatus>
locality.
</emphasis>
<location id="8EEBF3E73817FFD3FC863DDFFC235685" LSID="urn:lsid:plazi:treatment:039D142A3817FFD6FDA73B5AFBEF537D:8EEBF3E73817FFD3FC863DDFFC235685" box="[849,948,1221,1245]" country="Russia" municipality="Sambian" name="Yantarny" pageId="1" pageNumber="228" stateProvince="Kaliningrad">Yantarny</location>
settlement (formerly Palmnicken), the
<collectingMunicipality id="6BEF3F463817FFD3FC763DF9FC6956A3" box="[929,1022,1251,1275]" pageId="1" pageNumber="228">Sambian</collectingMunicipality>
(Samland) peninsula, the
<collectingRegion id="49F06BDE3817FFD3FCB23C1BFBDC5741" box="[869,1099,1281,1305]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="228">Kaliningrad Region</collectingRegion>
, western
<collectingCountry id="F323E5AC3817FFD3FD1E3C3AFC835760" box="[713,788,1312,1336]" name="Russia" pageId="1" pageNumber="228">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32EF6B73817FFD0FDA73C47FCC857CC" lastPageId="2" lastPageNumber="229" pageId="1" pageNumber="228" type="description">
<paragraph id="8B8BA53C3817FFD0FDA73C47FDD9533A" blockId="1.[624,1103,1373,1643]" lastBlockId="2.[189,669,178,355]" lastPageId="2" lastPageNumber="229" pageId="1" pageNumber="228">
<emphasis id="B940792E3817FFD3FDA73C47FC96572E" bold="true" box="[624,769,1373,1398]" pageId="1" pageNumber="228">Description.</emphasis>
Body elongate-subovoid, about 1.8× longer than wide, moderately convex; integument unicolorous dark brown (as preserved). Pubescence includes: pronotum and elytra densely covered with rather long, semierect, unicolorous dark setae (some setae seem whitish because of minute gas layer surrounding them, or separation from the surrounding amber); ventrum with finer, recumbent, dark setation. Measurements: total body length
<quantity id="4CCC08D93814FFD0FF6A39F5FE8E535E" box="[189,281,239,263]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="229" unit="mm" value="3.2">3.2 mm</quantity>
, maximum body width
<quantity id="4CCC08D93814FFD0FDE239F5FD00535E" box="[565,663,239,263]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="229" unit="mm" value="1.8">1.8 mm</quantity>
; pronotum length
<quantity id="4CCC08D93814FFD0FE7D3814FD8F537D" box="[426,536,270,294]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="229" unit="mm" value="0.8">0.8 mm</quantity>
, pronotal maximum width
<quantity id="4CCC08D93814FFD0FEAF3836FE58531C" box="[376,463,300,324]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="229" unit="mm" value="1.6">1.6 mm</quantity>
; elytral length
<quantity id="4CCC08D93814FFD0FDA33836FF7E533A" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="229" unit="mm" value="2.4">2.4 mm</quantity>
, elytral maximum width
<quantity id="4CCC08D93814FFD0FE223851FDDD533A" box="[501,586,331,355]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="229" unit="mm" value="1.8">1.8 mm</quantity>
.
</paragraph>
<paragraph id="8B8BA53C3814FFD0FF6A3892FEF55639" blockId="2.[189,669,392,1121]" pageId="2" pageNumber="229">Head densely covered with fine punctation. Frontal median ocellus present. Compound eyes oval, rather large, convex, and entire. Antennae with ten antennomeres, and clavate; antennae short, extending about to posterior one-third of pronotal length; three apical antennomeres forming compact club, slightly longer than combined length of antennomeres 27; scape subcylindrical, elongate, large; antennomere 2 hemispherical, shorter and narrower than scape, and distinctly wider than antennomere 3; antennomeres 37 small, subquadrate to transverse; antennomere 8 trapezoidal, distinctly dilated apically, transverse, 1.25× wider than long, and markedly larger than antennomere 7; antennomere 9 trapezoidal, weakly dilated apically, transverse, 1.5× wider than long, as long as antennomere 8; antennomere 10 subtriangular with rounded apex, about as long as wide, slightly narrower than antennomere 9, short, about 1.25× as long as antennomere 9.</paragraph>
<paragraph id="8B8BA53C3814FFD0FF6A3D9DFDC657A8" blockId="2.[189,668,1159,1520]" pageId="2" pageNumber="229">Pronotum strongly transverse, 2.0× wider than long, bell-shaped, widest posteriorly; lateral margins of pronotum widely rounded, posterior margin bisinuate, anterior margin rounded; pronotal punctation fine and sparse, with distance between punctures distinctly larger than diameter of one puncture. Hypomeron with distinct antennal cavity; finely and densely punctate. Prosternum with dense, small punctation; forming “collar” under which mouthparts fit when head is retracted.</paragraph>
<paragraph id="8B8BA53C3814FFD0FF6A3F0FFD0B5432" blockId="2.[189,668,1557,1643]" pageId="2" pageNumber="229">Scutellar shield small, triangular with pointed apex, transverse, about 1.5× wider than long, with few fine puctures,</paragraph>
<paragraph id="8B8BA53C3814FFD0FD6C39A8FC015292" blockId="2.[699,918,178,202]" box="[699,918,178,202]" pageId="2" pageNumber="229">apparently glabrous.</paragraph>
<paragraph id="8B8BA53C3814FFD0FD6C39F5FC595115" blockId="2.[699,1179,239,845]" pageId="2" pageNumber="229">Elytra 1.1× longer than wide, widest within anterior one-quarter of length, narrowed posteriad, with anterior margin deeply concave, and humeral callus weak; elytral surface densely covered with small, irregular punctation (punctures larger than pronotal punctures), distance between punctures 1.02.0× diameter of one puncture; interspaces with micropunctation. Epipleura narrow, slightly impressed, wider at humeri and gradually narrowing posteriad, incomplete, not reaching elytral apex. Metaventrite with disc convex; densely covered with small punctation, distance between punctures apparently smaller than diameter of one puncture. Metepisternum wide, subrectangular, about 4.0× longer than wide, with dense and small punctures, distance between punctures smaller than diameter of one puncture.</paragraph>
<paragraph id="8B8BA53C3814FFD0FD6C3A69FBB756E5" blockId="2.[699,1179,883,1213]" pageId="2" pageNumber="229">Legs slender, rather long, covered with fine punctures and short, recumbent setae. Femora flattened, weakly spindle-shaped, with longitudinal groove ventrally. Tibia straight, subcylindrical, about 0.5× as wide as femoral width; protibia without spines, slightly shorter than meso- and metatibia. Tarsi thin, moderately long, about 0.5× as long as tibiae; protarsus slightly shorter than meso- and metatarsus. Pretarsal claws thin and rather long, free, simple.</paragraph>
<paragraph id="8B8BA53C3814FFD0FD6C3DF9FCC857CC" blockId="2.[699,1178,1251,1428]" pageId="2" pageNumber="229">Abdomen with five visible ventrites; ventrite 5 with widely rounded apical margin; cuticle densely covered with small punctation; relative length ratios of ventrites 15 equal to 11:11:9:8:10 (measured medially).</paragraph>
</subSubSection>
<subSubSection id="C32EF6B73814FFD6FD6C3CA3FD8F53DA" lastPageId="4" lastPageNumber="231" pageId="2" pageNumber="229" type="diagnosis">
<paragraph id="8B8BA53C3814FFD0FD6C3CA3FB0D5433" blockId="2.[699,1179,1465,1643]" pageId="2" pageNumber="229">
<emphasis id="B940792E3814FFD0FD6C3CA3FC75578A" bold="true" box="[699,994,1465,1490]" pageId="2" pageNumber="229">Differential diagnosis.</emphasis>
<taxonomicName id="4C34DEBF3814FFD0FBC33CA3FCDC57A8" authorityName="Bukejs &amp; Háva &amp; McKellar &amp; Alekseev" authorityYear="2023" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="229" phylum="Arthropoda" rank="species" species="samlandensis" status="sp. nov.">
<emphasis id="B940792E3814FFD0FBC33CA3FCDC57A8" italics="true" pageId="2" pageNumber="229">Globicornis samlandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273C4553814FFD0FC8D3CC3FC2057A8" box="[858,951,1497,1520]" pageId="2" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
differs from other extinct species of the genus (
<taxonomicName id="4C34DEBF3814FFD0FC2C3CEDFB1A5457" authorityName="Hava, Prokop et Herrmann" authorityYear="2006" box="[1019,1165,1527,1551]" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="229" phylum="Arthropoda" rank="species" species="ambericus">
<emphasis id="B940792E3814FFD0FC2C3CEDFB995456" box="[1019,1038,1527,1550]" italics="true" pageId="2" pageNumber="229">G</emphasis>
.
<emphasis id="B940792E3814FFD0FBC93CEDFB1A5457" box="[1054,1165,1527,1551]" italics="true" pageId="2" pageNumber="229">ambericus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C34DEBF3814FFD0FD6C3F0CFC1B5475" authorityName="Hava et Damgaard" authorityYear="2015" box="[699,908,1557,1581]" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="229" phylum="Arthropoda" rank="species" species="ingelehmannae">
<emphasis id="B940792E3814FFD0FD6C3F0CFC1B5475" box="[699,908,1557,1581]" italics="true" pageId="2" pageNumber="229">G. ingelehmannae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C34DEBF3814FFD0FC713F0CFBB15475" authorityName="Bukejs et Hava" authorityYear="2018" box="[934,1062,1557,1581]" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="229" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="B940792E3814FFD0FC713F0CFC2E5475" box="[934,953,1558,1581]" italics="true" pageId="2" pageNumber="229">G</emphasis>
.
<emphasis id="B940792E3814FFD0FC043F0FFBB15475" box="[979,1062,1557,1581]" italics="true" pageId="2" pageNumber="229">groehni</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C34DEBF3814FFD0FBAD3F0CFC855414" authorityName="Hava" authorityYear="2008" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="229" phylum="Arthropoda" rank="species" species="rakovici">
<emphasis id="B940792E3814FFD0FBAD3F0CFB1A5475" box="[1146,1165,1558,1581]" italics="true" pageId="2" pageNumber="229">G</emphasis>
.
<emphasis id="B940792E3814FFD0FD6C3F2EFC855414" box="[699,786,1588,1612]" italics="true" pageId="2" pageNumber="229">rakovici</emphasis>
</taxonomicName>
) in possessing a combination of the following characters: (1) antennomere 10
</paragraph>
<paragraph id="8B8BA53C3814FFD1FBBB3FBCFBDF52B0" blockId="2.[1132,1172,1702,1726]" lastBlockId="3.[624,1096,178,232]" lastPageId="3" lastPageNumber="230" pageId="2" pageNumber="229">229 subtriangular with rounded apex, about as long as wide, short, not longer than antennomeres 8 and 9 combined; (2) pronotum and elytra unicolorous dark brown, and covered with unicolorous dark setae (without patches). See also key below.</paragraph>
<caption id="DF4BF5B43815FFD1FFA53D1EFDFE5663" pageId="3" pageNumber="230" startId="3.[114,153,1028,1052]" targetBox="[177,1064,338,1022]" targetPageId="3" targetType="figure">
<paragraph id="8B8BA53C3815FFD1FFA53D1EFDFE5663" blockId="3.[114,1096,1028,1083]" pageId="3" pageNumber="230">
Fig. 1.
<taxonomicName id="4C34DEBF3815FFD1FF113D1EFE775644" authorityName="Bukejs &amp; Háva &amp; McKellar &amp; Alekseev" authorityYear="2023" box="[198,480,1028,1052]" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="230" phylum="Arthropoda" rank="species" species="samlandensis" status="sp. nov.">
<emphasis id="B940792E3815FFD1FF113D1EFE775644" box="[198,480,1028,1052]" italics="true" pageId="3" pageNumber="230">Globicornis samlandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273C4553815FFD1FE3C3D1FFDD15644" box="[491,582,1029,1052]" pageId="3" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, No RSKM_PAL_A66 [RSM]: A habitus, dorsal view; B habitus, ventral view.
</paragraph>
</caption>
<caption id="DF4BF5B43815FFD1FFA53F10FD8A5407" pageId="3" pageNumber="230" startId="3.[114,153,1546,1570]" targetBox="[215,987,1114,1539]" targetPageId="3" targetType="figure">
<paragraph id="8B8BA53C3815FFD1FFA53F10FD8A5407" blockId="3.[114,1097,1546,1632]" pageId="3" pageNumber="230">
Fig. 2.
<taxonomicName id="4C34DEBF3815FFD1FF163F10FE4F547A" authorityName="Bukejs &amp; Háva &amp; McKellar &amp; Alekseev" authorityYear="2023" box="[193,472,1546,1570]" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="230" phylum="Arthropoda" rank="species" species="samlandensis" status="sp. nov.">
<emphasis id="B940792E3815FFD1FF163F10FE4F547A" box="[193,472,1546,1570]" italics="true" pageId="3" pageNumber="230">Globicornis samlandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273C4553815FFD1FE373F10FDAE5479" box="[480,569,1546,1569]" pageId="3" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, No RSKM_PAL_A66 [RSM], details of forebody, ventral view. Abbreviations: a7a10 antennomeres 710 respectively; ey compound eye; pc prosternal “collar“.
</paragraph>
</caption>
<paragraph id="8B8BA53C3815FFD6FFA53FBCFD8F53DA" blockId="3.[114,154,1702,1726]" lastBlockId="4.[189,668,178,386]" lastPageId="4" lastPageNumber="231" pageId="3" pageNumber="230">
230
<emphasis id="B940792E3812FFD6FF6A39A8FEA85292" bold="true" box="[189,319,178,202]" pageId="4" pageNumber="231">Etymology.</emphasis>
The species name
<taxonomicName id="4C34DEBF3812FFD6FDD839A8FF4D52B0" authorityName="Bukejs &amp; Háva &amp; McKellar &amp; Alekseev" authorityYear="2023" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="samlandensis">“samlandensis</taxonomicName>
” is toponymic and derived from “das Samland”, the German name of the Sambi- an peninsula, where the main deposits of Baltic amber are located, and where the amber piece with the
<typeStatus id="548F1B9E3812FFD6FE7F3851FD92533B" box="[424,517,331,355]" pageId="4" pageNumber="231" type="holotype">holotype</typeStatus>
of the carpet beetle described here originated.
</paragraph>
</subSubSection>
<subSubSection id="C32EF6B73812FFD6FF6A38DCFBEF537D" pageId="4" pageNumber="231" type="key">
<paragraph id="8B8BA53C3812FFD6FF6A38DCFECE5062" blockId="4.[189,668,453,570]" pageId="4" pageNumber="231">
<emphasis id="B940792E3812FFD6FF6A38DCFDC053A4" bold="true" pageId="4" pageNumber="231">
KEY TO
<taxonomicName id="4C34DEBF3812FFD6FE8338DFFD0B5386" box="[340,668,453,478]" class="Insecta" family="Dermestidae" genus="Globicornis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B940792E3812FFD6FE8338DFFD9C5385" bold="true" box="[340,523,453,477]" italics="true" pageId="4" pageNumber="231">GLOBICORNIS</emphasis>
SPECIES
</taxonomicName>
KNOWN FROM BALTIC AMBER
</emphasis>
(according to
<bibRefCitation id="EFA5D8CD3812FFD6FE8E3B19FDCF5043" author="Bukejs A. &amp; Hava J." box="[345,600,515,539]" pageId="4" pageNumber="231" pagination="395 - 400" refId="ref2263" refString="Bukejs A., Hava J. 2018. A new species of Globicornis Latreille (Coleopetra: Dermestidae) from Baltic amber, with key to fossil species. Zootaxa 4483 (2): 395 - 400." type="journal article" year="2018">Bukejs &amp; Háva (2018)</bibRefCitation>
, with modifications)
</paragraph>
<paragraph id="8B8BA53C3812FFD6FF6A3B45FD025177" blockId="4.[189,668,607,815]" pageId="4" pageNumber="231">1. Antennomere 10 circular to subtriangular with rounded apex, short, not longer than combined length of antennomeres 8 and 9 ................................................................... 2 Antennomere 10 triangular, long, longer than combined length of antennomeres 8 and 9 .......................................................... 4</paragraph>
<paragraph id="8B8BA53C3812FFD6FF6A3A4EFDA55190" blockId="4.[189,668,852,1121]" pageId="4" pageNumber="231">
2. Body bicolorous, pronotum and elytra black with orange-red spots; antennomere 10 circular; body length
<quantity id="4CCC08D93812FFD6FE1E3A8BFD8A51F1" box="[457,541,913,937]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="231" unit="mm" value="2.5">2.5 mm</quantity>
.................
<taxonomicName id="4C34DEBF3812FFD6FF6A3AAAFDA55190" authorityName="Hava" authorityYear="2008" box="[189,562,944,968]" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="rakovici" subGenus="Globicornis">
<emphasis id="B940792E3812FFD6FF6A3AAAFEAB5190" box="[189,316,944,968]" italics="true" pageId="4" pageNumber="231">Globicornis</emphasis>
(
<emphasis id="B940792E3812FFD6FE9B3AAAFE5C5190" box="[332,459,944,968]" italics="true" pageId="4" pageNumber="231">Globicornis</emphasis>
)
<emphasis id="B940792E3812FFD6FE0C3AAAFDA55190" box="[475,562,944,968]" italics="true" pageId="4" pageNumber="231">rakovici</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8B8BA53C3812FFD6FF6A3AD5FD1C5639" blockId="4.[189,668,852,1121]" pageId="4" pageNumber="231">
Body unicolorous, dark brown or black, pronotum and elytra without paler cuticular spots; antennomere 10 subtriangular with rounded apex; body length not less than
<quantity id="4CCC08D93812FFD6FDA33D31FF705639" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="231" unit="mm" value="2.7">2.7 mm</quantity>
........................................................... 3
</paragraph>
<paragraph id="8B8BA53C3812FFD6FF6A3D9DFDB95760" blockId="4.[189,669,1159,1489]" pageId="4" pageNumber="231">
3. Pronotum and elytra with bicolourous setae: black, and yellowish to white, with paler setae forming patches; antennomere 10 transverse, about 1.5× longer than wide; body length
<quantity id="4CCC08D93812FFD6FE533C1BFE615741" box="[388,502,1281,1305]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="231" unit="mm" value="3.6">3.6 mm</quantity>
..................
<taxonomicName id="4C34DEBF3812FFD6FF6A3C3AFDB95760" authorityName="Bukejs et Hava" authorityYear="2018" box="[189,558,1312,1336]" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="groehni" subGenus="Globicornis">
<emphasis id="B940792E3812FFD6FF6A3C3AFEAB5760" box="[189,316,1312,1336]" italics="true" pageId="4" pageNumber="231">Globicornis</emphasis>
(
<emphasis id="B940792E3812FFD6FE9B3C3AFE5C5760" box="[332,459,1312,1336]" italics="true" pageId="4" pageNumber="231">Globicornis</emphasis>
)
<emphasis id="B940792E3812FFD6FE0C3C3AFDB95760" box="[475,558,1312,1336]" italics="true" pageId="4" pageNumber="231">groehni</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8B8BA53C3812FFD6FF6A3C25FDD75789" blockId="4.[189,669,1159,1489]" pageId="4" pageNumber="231">
Pronotum and elytra with unicolorous dark setae, without patches; antennomere 10 about as long as wide; body length
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...….................……..….
<taxonomicName id="4C34DEBF3812FFD6FDC13C81FE725789" authorityName="Bukejs &amp; Háva &amp; McKellar &amp; Alekseev" authorityYear="2023" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="samlandensis" status="sp. nov." subGenus="Globicornis">
<emphasis id="B940792E3812FFD6FDC13C81FD0257EB" box="[534,661,1435,1459]" italics="true" pageId="4" pageNumber="231">Globicornis</emphasis>
(
<emphasis id="B940792E3812FFD6FF113CA3FED25789" box="[198,325,1465,1489]" italics="true" pageId="4" pageNumber="231">Globicornis</emphasis>
)
<emphasis id="B940792E3812FFD6FE823CA3FE725789" box="[341,485,1465,1489]" italics="true" pageId="4" pageNumber="231">samlandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273C4553812FFD6FE3B3CA0FDD75789" box="[492,576,1466,1489]" pageId="4" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B8BA53C3812FFD6FF6A3CEDFDD15433" blockId="4.[189,668,1527,1643]" pageId="4" pageNumber="231">
4. Body covered with white setation; pronotum less transverse, 1.8× wider than long; body length 3.0 mm ...........................
<taxonomicName id="4C34DEBF3812FFD6FF6A3F49FDD15433" authorityName="Hava, Prokop et Herrmann" authorityYear="2006" box="[189,582,1619,1643]" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="ambericus" subGenus="Hadrotoma">
<emphasis id="B940792E3812FFD6FF6A3F49FED45433" box="[189,323,1619,1643]" italics="true" pageId="4" pageNumber="231">Globicornis</emphasis>
(
<emphasis id="B940792E3812FFD6FE9B3F49FE505433" box="[332,455,1619,1643]" italics="true" pageId="4" pageNumber="231">Hadrotoma</emphasis>
)
<emphasis id="B940792E3812FFD6FE003F49FDD15433" box="[471,582,1619,1643]" italics="true" pageId="4" pageNumber="231">ambericus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8B8BA53C3812FFD6FD6C39A8FBEF537D" blockId="4.[699,1178,178,294]" pageId="4" pageNumber="231">
Body covered with black setation; pronotum more transverse, 2.3× wider than long; body length
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..........................
<taxonomicName id="4C34DEBF3812FFD6FD6C3817FBEF537D" authorityName="Hava et Damgaard" authorityYear="2015" box="[699,1144,269,294]" class="Insecta" family="Dermestidae" genus="Globicornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="ingelehmannae" subGenus="Hadrotoma">
<emphasis id="B940792E3812FFD6FD6C3817FCD6537D" box="[699,833,269,293]" italics="true" pageId="4" pageNumber="231">Globicornis</emphasis>
(
<emphasis id="B940792E3812FFD6FC9D3817FC52537D" box="[842,965,269,293]" italics="true" pageId="4" pageNumber="231">Hadrotoma</emphasis>
)
<emphasis id="B940792E3812FFD6FC023817FBEF537D" box="[981,1144,269,293]" italics="true" pageId="4" pageNumber="231">ingelehmannae</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,173 @@
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<emphasis id="B975EAC9FFEA825AFFC1F9E4FD83F9DE" bold="true" box="[114,550,1571,1595]" pageId="1" pageNumber="264">
<taxonomicName id="4C014D58FFEA825AFFC1F9E4FE6FF9DE" authorityName="Barševskis &amp; Torrejos" authorityYear="2023" box="[114,458,1571,1595]" class="Insecta" family="Cerambycidae" genus="Borneogrammus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="hamiguitanus" status="sp. nov.">
<emphasis id="B975EAC9FFEA825AFFC1F9E4FE6FF9DE" bold="true" box="[114,458,1571,1595]" italics="true" pageId="1" pageNumber="264">Borneogrammus hamiguitanus</emphasis>
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<taxonomicNameLabel id="A24657B2FFEA825AFE62F9E4FD83F9DE" box="[465,550,1571,1595]" pageId="1" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="B975EAC9FFEA825AFFC1F983FF66F9B9" bold="true" box="[114,195,1604,1628]" pageId="1" pageNumber="264">
(
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)
</emphasis>
</paragraph>
<paragraph id="8BBE36DBFFEA825AFFC1F961FCC7FAC1" blockId="1.[114,154,1702,1726]" lastBlockId="1.[624,1103,1077,1316]" pageId="1" pageNumber="264">
<materialsCitation id="3B693C86FFEA825AFFC1F961FCFBFAC1" collectingDate="2023-05-27" collectionCode="DUBC" collectorName="A. Barsevskis" country="Philippines" determinerName="A. Barsevskis &amp; Ch &amp; Torrejos" location="Mt. Hamiguitan" municipality="Mindanao Isl." pageId="1" pageNumber="264" specimenCount="264" stateProvince="Davao Oriental" typeStatus="holotype">
<specimenCount id="9D07FD52FFEA825AFFC1F961FD0DFBA8" box="[114,680,1077,1726]" count="264" pageId="1" pageNumber="264" type="generic">
264
<emphasis id="B975EAC9FFEA825AFDC3FBF2FD0DFBA8" bold="true" box="[624,680,1077,1101]" pageId="1" pageNumber="264">Type</emphasis>
</specimenCount>
specimens:
<typeStatus id="54BA8879FFEA825AFCC0FBF2FC40FBA8" box="[883,997,1077,1101]" pageId="1" pageNumber="264" type="holotype">Holotype</typeStatus>
: Male.
<emphasis id="B975EAC9FFEA825AFDC3FB94FDD2FB8E" bold="true" box="[624,631,1107,1131]" pageId="1" pageNumber="264">/</emphasis>
<collectingCountry id="F316764BFFEA825AFDC4FB93FD56FB89" box="[631,755,1108,1132]" name="Philippines" pageId="1" pageNumber="264">Philippines</collectingCountry>
: /
<collectingMunicipality id="6BDAACA1FFEA825AFC9BFB93FC68FB89" box="[808,973,1108,1132]" pageId="1" pageNumber="264">Mindanao Isl.</collectingMunicipality>
, /
<collectingRegion id="49C5F839FFEA825AFBB0FB93FD6FFB6E" country="Philippines" name="Davao Oriental" pageId="1" pageNumber="264">Davao Oriental</collectingRegion>
, / Pujada Penins., /
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env., /
<date id="FFBF101BFFEA825AFD7AFB56FCE1FB4C" box="[713,836,1169,1193]" pageId="1" pageNumber="264" value="2023-05-27">
<collectingDate id="EFFBE9F3FFEA825AFD7AFB56FCE1FB4C" box="[713,836,1169,1193]" pageId="1" pageNumber="264" value="2023-05-27">27.05.2023</collectingDate>
</date>
, /
<collectorName id="26F4530DFFEA825AFCDEFB56FC5FFB4C" box="[877,1018,1169,1193]" pageId="1" pageNumber="264">A.Barševskis</collectorName>
leg. / [handwritten]; //
<typeStatus id="54BA8879FFEA825AFC9FFB77FC6BFB2D" box="[812,974,1200,1224]" pageId="1" pageNumber="264" type="holotype">HOLOTYPUS</typeStatus>
:/
<taxonomicName id="4C014D58FFEA825AFC50FB77FD63FB03" authorityName="Vives &amp; Heffern" authorityYear="2016" class="Insecta" family="Cerambycidae" genus="Borneogrammus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="genus">Borneogrammus</taxonomicName>
/ hamiguitanus
<taxonomicNameLabel id="A24657B2FFEA825AFC74FB09FBBBFB03" box="[967,1054,1230,1254]" pageId="1" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
/
<determinerName id="7E62DA1FFFEA825AFDC3FB2AFCA5FAE0" box="[624,768,1261,1285]" pageId="1" pageNumber="264">A.Barševskis</determinerName>
,
<determinerName id="7E62DA1FFFEA825AFCBEFB2AFC8AFAE0" box="[781,815,1261,1285]" pageId="1" pageNumber="264">Ch</determinerName>
,
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det. [red label, handwritten] (
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)
</materialsCitation>
.
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<emphasis id="B975EAC9FFEA825AFDC3FA8EFCD0FA84" bold="true" box="[624,885,1353,1377]" pageId="1" pageNumber="264">General distribution:</emphasis>
<collectingCountry id="F316764BFFEA825AFC32FA8EFC5BFA84" box="[897,1022,1353,1377]" name="Philippines" pageId="1" pageNumber="264">Philippines</collectingCountry>
: Mindanao island.
</paragraph>
<paragraph id="8BBE36DBFFEA825AFDC3FA62FCD2F9DD" blockId="1.[624,1103,1445,1592]" pageId="1" pageNumber="264">
<emphasis id="B975EAC9FFEA825AFDC3FA62FCA4FA58" bold="true" box="[624,769,1445,1469]" pageId="1" pageNumber="264">Description.</emphasis>
Dark - brown, elongated, with parallel sides, elytra with coarse punctures and two interrupted narrow bands of yellow pubescence. Body lenght
<quantity id="4CF99B3EFFEA825AFC21F9C6FC5DF9FC" box="[914,1016,1537,1561]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="1" pageNumber="264" unit="mm" value="11.3">11.3 mm</quantity>
. Maximal body width
<quantity id="4CF99B3EFFEA825AFCADF9E7FCD6F9DD" box="[798,883,1568,1592]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="264" unit="mm" value="2.7">2.7 mm</quantity>
.
</paragraph>
<paragraph id="8BBE36DBFFE98259FD08FF75FBABFEE2" blockId="2.[699,1178,178,263]" pageId="2" pageNumber="265">transverse, very narrow, yellow, shiny, with nummerous long setae. Mandibles sharp, relatively massive, dark-brown.</paragraph>
<paragraph id="8BBE36DBFFE98259FD08FEEBFC46FE19" blockId="2.[699,1178,300,508]" pageId="2" pageNumber="265">Pronotum slightly elongate, cylindric, with small sharp denticle at lateraal margin. Sides of pronotum very slightly constricted before apical margin, densely and coarsely punctate. Dorsal disc of pronotum wis elongate narrow middle line and two lateral lines of yellow pubescence.</paragraph>
<paragraph id="8BBE36DBFFE98259FD08FDE5FBD7FD70" blockId="2.[699,1178,546,661]" pageId="2" pageNumber="265">
Scutellum small, slightly elongated, with dense yellow pubescence.
<emphasis id="B975EAC9FFE98259FC49FD86FB3FFDBD" box="[1018,1178,576,600]" italics="true" pageId="2" pageNumber="265">Pars stridens</emphasis>
slightly visible in the form of a smooth area with very fine transverse microsculpture.
</paragraph>
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<paragraph id="8BBE36DBFFE98259FF0EFCBCFEA1FC34" blockId="2.[189,668,891,977]" pageId="2" pageNumber="265">
Fig. 2.
<taxonomicName id="4C014D58FFE98259FE9DFCBBFE1EFC57" authority="Vives &amp; Heffern 2016" authorityName="Vives &amp; Heffern" authorityYear="2016" class="Insecta" family="Cerambycidae" genus="Borneogrammus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="species" species="albomaculatus">
<emphasis id="B975EAC9FFE98259FE9DFCBBFD33FC71" box="[302,662,892,916]" italics="true" pageId="2" pageNumber="265">Borneogrammus albomaculatus</emphasis>
<bibRefCitation id="EF904B2AFFE98259FF0EFC5DFE1EFC57" author="Vives E &amp; Heffern D." box="[189,443,922,946]" pageId="2" pageNumber="265" pagination="107 - 128" refId="ref1455" refString="Vives E, Heffern D. 2016. Descriptions and notes on Oriental longhorned beetles, mostly from Borneo, Part III (Coleoptera, Cerambycidae, Lamiinae). Les Cahiers Magellanes, N 0 23: 107 - 128." type="journal article" year="2016">Vives &amp; Heffern 2016</bibRefCitation>
</taxonomicName>
(holotype (
<bibRefCitation id="EF904B2AFFE98259FDF0FC5DFF56FC34" author="Berzak L." pageId="2" pageNumber="265" refId="ref1298" refString="Berzak L. 2023. Old World Cerambycidae Catalog. http: // bezbycids. com / byc iddb / wdefault. asp? w = o [accessed: 01.11. 2023]." type="url" year="2023">Berzak, 2023</bibRefCitation>
))
</paragraph>
</caption>
<paragraph id="8BBE36DBFFE98259FF0EFC31FD5CFC2D" blockId="2.[189,669,1014,1651]" lastBlockId="2.[699,1179,699,968]" pageId="2" pageNumber="265">Head hypognathous, quadrangular, with sparse, brown pubescence. The surface of the head is flat, even, slightly convex towards the front. Frons even, coarsely punctate. Between relatively big, slightly convex eyes weakly defined short midline, but between antennal tubercles with deep wide impression. Cheeks smooth, shiny, slightly extended, with coarse punctures and covered with fine, pubescence. The portions of the face around the cheeks are covered with relatively longer yellow-brown pubescence. Antennae relatively longer than apex of elytra, with dark first antennomere and brown remaining antennomeres, covered with very fine pubescence. Inside of antennomeres with a band of longer mixed dark and pale bristles, which extend more at the tips of the antennae, where the antennomeres resemble a brush. Labrum transverse, narrow, covered with long setae. Clypeus Elytra parallel, slightly narrowed in the middle, each elytron is rounded at the end; coarsely and irregularly punctate. Elytra with a raised suture and a raised longitudinal line covered with yellow pubescence, interrupted in some places, but at apical portion with a two additional parallel interrupted rudimentary lines of small yellow spots.</paragraph>
<paragraph id="8BBE36DBFFE98259FD08FC2AFBDDFB84" blockId="2.[699,1178,1005,1121]" pageId="2" pageNumber="265">Underside of body with yellow pubescence of different density. Legs short, massive, dark-brown, with very fine, pubescence and some erect hairs. Tarsomeres dark brown.</paragraph>
</subSubSection>
<subSubSection id="C31B6550FFE98259FD08FB41FB31F95B" pageId="2" pageNumber="265" type="diagnosis">
<paragraph id="8BBE36DBFFE98259FD08FB41FC17F9EA" blockId="2.[699,1178,1158,1551]" pageId="2" pageNumber="265">
<emphasis id="B975EAC9FFE98259FD08FB41FC1CFB7B" bold="true" box="[699,953,1158,1182]" pageId="2" pageNumber="265">Differential diagnosis.</emphasis>
Comparing the external shape of the surface of the new species with second species of this genus
<taxonomicName id="4C014D58FFE98259FBD0FB02FC8FFAFC" authority="Vives &amp; Heffern 2016" authorityName="Vives &amp; Heffern" authorityYear="2016" class="Insecta" family="Cerambycidae" genus="Borneogrammus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="species" species="albomaculatus" status="sp. nov.">
<emphasis id="B975EAC9FFE98259FBD0FB02FC4BFB1E" italics="true" pageId="2" pageNumber="265">Borneogrammus albomaculatus</emphasis>
<bibRefCitation id="EF904B2AFFE98259FC49FB24FC8FFAFC" author="Vives E &amp; Heffern D." pageId="2" pageNumber="265" pagination="107 - 128" refId="ref1455" refString="Vives E, Heffern D. 2016. Descriptions and notes on Oriental longhorned beetles, mostly from Borneo, Part III (Coleoptera, Cerambycidae, Lamiinae). Les Cahiers Magellanes, N 0 23: 107 - 128." type="journal article" year="2016">Vives &amp; Heffern 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A24657B2FFE98259FC8AFAC6FC18FAFC" box="[825,957,1281,1305]" pageId="2" pageNumber="265" rank="species">new species</taxonomicNameLabel>
differs in different pattern of elytra. The new species has elytra with with a raised suture and a raised longitudinal line of yellow pubescence, interrupted in some places, but at apical portion - a two additional parallel interrupted rudimentary lines of small yellow spots. Elytra of
<taxonomicName id="4C014D58FFE98259FC94FA1EFC4EFA15" authorityName="Vives &amp; Heffern" authorityYear="2016" box="[807,1003,1496,1520]" class="Insecta" family="Cerambycidae" genus="Borneogrammus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="species" species="albomaculatus">
<emphasis id="B975EAC9FFE98259FC94FA1EFC4EFA15" box="[807,1003,1496,1520]" italics="true" pageId="2" pageNumber="265">B. albomaculatus</emphasis>
</taxonomicName>
evenly covered with small white spots.
</paragraph>
<paragraph id="8BBE36DBFFE98259FBDFF961FB31F95B" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="265">265</paragraph>
</subSubSection>
<subSubSection id="C31B6550FFE88258FFC1FF76FEECFEE2" pageId="3" pageNumber="266" type="etymology">
<paragraph id="8BBE36DBFFE88258FFC1FF76FEECFEE2" blockId="3.[114,592,177,263]" pageId="3" pageNumber="266">
<emphasis id="B975EAC9FFE88258FFC1FF76FF51FF2C" bold="true" box="[114,244,177,201]" pageId="3" pageNumber="266">Etymology.</emphasis>
This species isnamed after its place of origin - Mt. Hamiguitan [Hamiguitan -
<taxonomicName id="4C014D58FFE88258FF1AFF28FE9FFEE2" authorityName="Barševskis &amp; Torrejos" authorityYear="2023" box="[169,314,239,263]" class="Insecta" family="Cerambycidae" genus="Borneogrammus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="266" phylum="Arthropoda" rank="species" species="hamiguitanus">
<emphasis id="B975EAC9FFE88258FF1AFF28FE9FFEE2" box="[169,314,239,263]" italics="true" pageId="3" pageNumber="266">hamiguitanus</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,221 @@
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<mods:title id="E089BED11E1B15D758BC3FF00AF264EB">New species of Broscosoma Rosenhauer, 1846 (Coleoptera: Carabidae) from China</mods:title>
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<treatment id="03AA87BAFFD91455FCEEA8D2FCB30DEA" LSID="urn:lsid:plazi:treatment:03AA87BAFFD91455FCEEA8D2FCB30DEA" httpUri="http://treatment.plazi.org/id/03AA87BAFFD91455FCEEA8D2FCB30DEA" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<subSubSection id="C3196527FFD91456FCEEA8D2FBA50BBB" box="[810,1113,192,216]" pageId="1" pageNumber="20" type="nomenclature">
<paragraph id="8BBC36ACFFD91456FCEEA8D2FBA50BBB" blockId="1.[810,1113,192,216]" box="[810,1113,192,216]" pageId="1" pageNumber="20">
<heading id="D0F481C0FFD91456FCEEA8D2FBA50BBB" bold="true" box="[810,1113,192,216]" fontSize="10" level="2" pageId="1" pageNumber="20" reason="4">
<emphasis id="B977EABEFFD91456FCEEA8D2FBA50BBB" bold="true" box="[810,1113,192,216]" italics="true" pageId="1" pageNumber="20">
<taxonomicName id="4C034D2FFFD91456FCEEA8D2FBE40BBB" ID-CoL="NDXW" authorityName="Barševskis" authorityYear="2010" box="[810,1048,192,216]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="valainisi" status="sp. nov.">Broscosoma valainisi</taxonomicName>
<taxonomicNameLabel id="A24457C5FFD91456FBDBA8D2FBA50BBB" box="[1055,1113,192,216]" pageId="1" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3196527FFD91456FCEEA912FAD40AB4" pageId="1" pageNumber="20" type="materials_examined">
<paragraph id="8BBC36ACFFD91456FCEEA912FB970A34" blockId="1.[810,1320,255,343]" pageId="1" pageNumber="20">
<materialsCitation id="3B6B3CF1FFD91456FCEEA912FB970A34" collectingDate="2005-10-08" collectionCode="V" collectorName="Patrikeev" country="China" elevation="2000" latitude="33.883335" location="Mts. Taibeichan Nat. Res." longLatPrecision="1202" longitude="107.816666" municipality="Tschingling Mts." pageId="1" pageNumber="20" specimenCount="1" stateProvince="Shanxi" typeStatus="holotype">
<emphasis id="B977EABEFFD91456FCEEA912FC660A7B" bold="true" box="[810,922,256,280]" pageId="1" pageNumber="20">
<typeStatus id="54B8880EFFD91456FCEEA912FC690A7B" box="[810,917,256,280]" pageId="1" pageNumber="20" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female.
<collectingCountry id="F314763CFFD91456FC38A8EDFBBF0A74" box="[1020,1091,255,279]" name="China" pageId="1" pageNumber="20">China</collectingCountry>
:
<collectingRegion id="49C7F84EFFD91456FB88A8EDFB600A74" box="[1100,1180,255,279]" country="China" name="Shanxi" pageId="1" pageNumber="20">Shanxi</collectingRegion>
,
<collectingMunicipality id="6BD8ACD6FFD91456FB62A8EDFCA40A54" pageId="1" pageNumber="20">
Tschingling
<location id="8EDC6077FFD91456FCEEA932FCA40A54" LSID="urn:lsid:plazi:treatment:03AA87BAFFD91455FCEEA8D2FCB30DEA:8EDC6077FFD91456FCEEA932FCA40A54" box="[810,856,288,311]" country="China" latitude="33.883335" longLatPrecision="1202" longitude="107.816666" municipality="Tschingling Mts." name="Mts. Taibeichan Nat. Res." pageId="1" pageNumber="20" stateProvince="Shanxi">Mts.</location>
</collectingMunicipality>
Taibeichan Nat. Res.,
<geoCoordinate id="EE37506BFFD91456FBF7A90DFB7B0A54" box="[1075,1159,286,311]" degrees="33" direction="north" minutes="53" orientation="latitude" pageId="1" pageNumber="20" precision="925" value="33.883335">
33
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53N
</geoCoordinate>
<geoCoordinate id="EE37506BFFD91456FB48A90DFB100A54" box="[1164,1260,286,311]" degrees="107" direction="east" minutes="49" orientation="longitude" pageId="1" pageNumber="20" precision="925" value="107.816666">
107
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49E
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,
<quantity id="4CFB9B49FFD91456FB31A90DFCBD0A34" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="20" unit="m" value="2000.0">
<elevation id="002ED19FFFD91456FB31A90DFCBD0A34" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="20" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="FFBD106CFFD91456FC8DA92DFC3D0A34" box="[841,961,319,343]" pageId="1" pageNumber="20" value="2005-10-08">
<collectingDate id="EFF9E984FFD91456FC8DA92DFC3D0A34" box="[841,961,319,343]" pageId="1" pageNumber="20" value="2005-10-08">10.08.2005</collectingDate>
</date>
.,
<collectionCode id="ED12AE69FFD91456FC08A92DFC210A34" box="[972,989,319,343]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="20" type="Museum">V</collectionCode>
.
<collectorName id="26F6537AFFD91456FC24A92DFBBC0A34" box="[992,1088,319,343]" pageId="1" pageNumber="20">Patrikeev</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BBC36ACFFD91456FCEEA992FAD40AB4" blockId="1.[810,1320,383,471]" pageId="1" pageNumber="20">
<emphasis id="B977EABEFFD91456FCEEA992FC3A0AFB" bold="true" box="[810,966,384,408]" pageId="1" pageNumber="20">Type locality.</emphasis>
<collectingCountry id="F314763CFFD91456FC0AA96DFBE80AF4" box="[974,1044,383,407]" name="China" pageId="1" pageNumber="20">China</collectingCountry>
,
<collectingRegion id="49C7F84EFFD91456FBE5A96DFB8D0AF4" box="[1057,1137,383,407]" country="China" name="Shanxi" pageId="1" pageNumber="20">Shanxi</collectingRegion>
. This is the first species of the genus
<taxonomicName id="4C034D2FFFD91456FBD3A9B3FADB0AD4" authority="Rosenhauer" authorityName="Rosenhauer" authorityYear="1846" box="[1047,1319,415,440]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B977EABEFFD91456FBD3A9B3FB610ADB" box="[1047,1181,417,440]" italics="true" pageId="1" pageNumber="20">Broscosoma</emphasis>
Rosenhauer
</taxonomicName>
that has been found in
<collectingRegion id="49C7F84EFFD91456FBD7A9ADFB430AB4" box="[1043,1215,447,471]" country="China" name="Shanxi" pageId="1" pageNumber="20">Shanxi Province</collectingRegion>
in
<collectingCountry id="F314763CFFD91456FB24A9ADFAD80AB4" box="[1248,1316,447,471]" name="China" pageId="1" pageNumber="20">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3196527FFD91455FCEEAA12FBC609D7" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" type="description">
<paragraph id="8BBC36ACFFD91456FCEEAA12FB3F09D4" blockId="1.[810,1320,511,695]" pageId="1" pageNumber="20">
<emphasis id="B977EABEFFD91456FCEEAA12FC53097B" bold="true" box="[810,943,512,536]" pageId="1" pageNumber="20">Description.</emphasis>
Female. The body is slender, convex, dark brown. Its surface is dark, plain but with bronze metallic lustre. Its length is
<quantity id="4CFB9B49FFD91456FB51AA2DFB090934" box="[1173,1269,575,599]" metricMagnitude="-3" metricUnit="m" metricValue="8.11" pageId="1" pageNumber="20" unit="mm" value="8.11">8.11 mm</quantity>
. The length of elytra is
<quantity id="4CFB9B49FFD91456FC2FAA4DFBB30914" box="[1003,1103,607,631]" metricMagnitude="-3" metricUnit="m" metricValue="4.02" pageId="1" pageNumber="20" unit="mm" value="4.02">4.02 mm</quantity>
, but their maximum width is
<quantity id="4CFB9B49FFD91456FC46AA6DFC1E09F4" box="[898,994,639,663]" metricMagnitude="-3" metricUnit="m" metricValue="2.69" pageId="1" pageNumber="20" unit="mm" value="2.69">2.69 mm</quantity>
. The length of its thorax is
<quantity id="4CFB9B49FFD91456FB3DAA6DFCA909D4" metricMagnitude="-3" metricUnit="m" metricValue="2.19" pageId="1" pageNumber="20" unit="mm" value="2.19">2.19 mm</quantity>
, but its maximum width is
<quantity id="4CFB9B49FFD91456FBA5AA8DFB4209D4" box="[1121,1214,671,695]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="1" pageNumber="20" unit="mm" value="1.64">1.64 mm</quantity>
.
</paragraph>
<paragraph id="8BBC36ACFFD91456FCEEAACDFBDA089B" blockId="1.[810,1321,735,1016]" pageId="1" pageNumber="20">The beetles head is pointed, with visibly convex surface, dark, lustrous, its front side is russet-brown. Its mouth organs and antennae are russet-brown, the antennae are a little lighter in colour. Its eyes, similar to most of the species of this genus, are small and not projecting outward. The frontal grooves irregular and deep, in the back side they are placed distantly from each other towards the eyes.</paragraph>
<paragraph id="8BBC36ACFFD91456FCEEAC0DFCA60F94" blockId="1.[810,1320,1055,1271]" pageId="1" pageNumber="20">The thorax is extended, the correlation between its length and width is 1.3. It is noticeably contracted in the main part and forms a neck-like joint before the elytra. Its upper part is curved, smooth and lustrous. On the thorax disk there is a centre line that is a little impressed in its front part.</paragraph>
<paragraph id="8BBC36ACFFD91456FCEEAD0DFBC20EF4" blockId="1.[810,1320,1311,1431]" pageId="1" pageNumber="20">The legs are dichromatic and lustrous. The femora are dark-brown or black, but in the bottom and top parts they are lighter, i.e., they are brown with russet-brown tarsus.</paragraph>
<paragraph id="8BBC36ACFFD91455FCEEADADFBC609D7" blockId="1.[810,1321,1471,1623]" lastBlockId="2.[886,1397,124,692]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
The elytra are noticeably egg-shaped with widely rounded shoulders. They are black or dark-brown, a little lighter in the top part. The surface is smooth, without micro-sculpture, it has bronze sheen. The beetle has well defined punctate. The with bronze lustre. It differs from
<taxonomicName id="4C034D2FFFDA1455FB0EA86FFC0C0BD7" authority="Sciaky &amp; Facchini" authorityName="Sciaky &amp; Facchini" authorityYear="2005" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="stefani">
<emphasis id="B977EABEFFDA1455FB0EA86FFAD70BF7" box="[1226,1323,124,148]" italics="true" pageId="2" pageNumber="21">B. stefani</emphasis>
Sciaky &amp; Facchini
</taxonomicName>
in its colouring and more extended and slender body. The body colouring of
<taxonomicName id="4C034D2FFFDA1455FA98A8AFFB780B97" authority="Sciaky &amp; Facchini" authorityName="Sciaky &amp; Facchini" authorityYear="2005" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="stefani">
<emphasis id="B977EABEFFDA1455FA98A8AFFC410B97" italics="true" pageId="2" pageNumber="21">B. stefani</emphasis>
Sciaky &amp; Facchini
</taxonomicName>
is black, with greenish or bluish hue, head with collar constriction deep and punctate, but the new species have dark brown body with bronze lustre. The elytra are more extended than those of
<taxonomicName id="4C034D2FFFDA1455FB6CA94FFC240AF7" authority="Sciaky &amp; Facchini" authorityName="Sciaky &amp; Facchini" authorityYear="2005" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="stefani">
<emphasis id="B977EABEFFDA1455FB6CA94FFAF10A17" box="[1192,1293,348,372]" italics="true" pageId="2" pageNumber="21">B. stefani</emphasis>
Sciaky &amp; Facchini
</taxonomicName>
, too. The species have different areas of distribution as well:
<taxonomicName id="4C034D2FFFDA1455FBAFA98FFABB0AD7" authority="Deuve" authorityName="Deuve" authorityYear="1990" box="[1131,1351,412,436]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="sichuanum">
<emphasis id="B977EABEFFDA1455FBAFA98FFB010AD7" box="[1131,1277,412,436]" italics="true" pageId="2" pageNumber="21">B. sichuanum</emphasis>
Deuve
</taxonomicName>
and
<taxonomicName id="4C034D2FFFDA1455FCB2A9AFFB520AB7" authority="Sciaky &amp; Facchini" authorityName="Sciaky &amp; Facchini" authorityYear="2005" box="[886,1198,444,468]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="stefani">
<emphasis id="B977EABEFFDA1455FCB2A9AFFC220AB7" box="[886,990,444,468]" italics="true" pageId="2" pageNumber="21">B. stefani</emphasis>
Sciaky &amp; Facchini
</taxonomicName>
are to be found in
<collectingRegion id="49C7F84EFFDA1455FCB2A9CEFBCF0A97" box="[886,1075,476,500]" country="China" name="Sichuan" pageId="2" pageNumber="21">Sichuan Province</collectingRegion>
in
<collectingCountry id="F314763CFFDA1455FB92A9CEFB670A97" box="[1110,1179,476,500]" name="China" pageId="2" pageNumber="21">China</collectingCountry>
, but the new species so far is known only from
<collectingRegion id="49C7F84EFFDA1455FB41A9EEFACA0977" box="[1157,1334,508,532]" country="China" name="Shanxi" pageId="2" pageNumber="21">Shanxi province</collectingRegion>
. Most of the species found in Eastern Asia are distributed in small areas, the only exception being
<taxonomicName id="4C034D2FFFDA1455FC7DAA4FFB600917" authority="Deuve" authorityName="Deuve" authorityYear="1990" box="[953,1180,604,628]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="sichuanum">
<emphasis id="B977EABEFFDA1455FC7DAA4FFBB20917" box="[953,1102,604,628]" italics="true" pageId="2" pageNumber="21">B. sichuanum</emphasis>
Deuve
</taxonomicName>
, which has recently been discovered in a larger territory in
<collectingRegion id="49C7F84EFFDA1455FADFAA6EFC2F09D7" country="China" name="Sichuan" pageId="2" pageNumber="21">Sichuan Province</collectingRegion>
in
<collectingCountry id="F314763CFFDA1455FC36AA8EFBCA09D7" box="[1010,1078,668,692]" name="China" pageId="2" pageNumber="21">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3196527FFDA1455FCB2AACFFCB30DEA" pageId="2" pageNumber="21" type="etymology">
<paragraph id="8BBC36ACFFDA1455FCB2AACFFC220897" blockId="2.[886,1397,732,1012]" pageId="2" pageNumber="21">
<emphasis id="B977EABEFFDA1455FCB2AACFFC060996" bold="true" box="[886,1018,733,757]" pageId="2" pageNumber="21">Etymology.</emphasis>
This species is named after my colleague and disciple Uldis Valainis (Institute of Systematic Biology,
<collectingRegion id="49C7F84EFFDA1455FBBCAB0EFB0F0857" box="[1144,1267,796,820]" country="Latvia" name="Daugavpils" pageId="2" pageNumber="21">Daugavpils</collectingRegion>
University,
<collectingRegion id="49C7F84EFFDA1455FCB2AB2EFC0A0837" box="[886,1014,828,852]" country="Latvia" name="Daugavpils" pageId="2" pageNumber="21">Daugavpils</collectingRegion>
,
<collectingCountry id="F314763CFFDA1455FBC2AB2EFBB20837" box="[1030,1102,828,852]" name="Latvia" pageId="2" pageNumber="21">Latvia</collectingCountry>
) who, having studied the ground beetles of the genus
<taxonomicName id="4C034D2FFFDA1455FB5EAB4EFABB0817" authority="Latr." authorityName="Latr." box="[1178,1351,860,884]" class="Insecta" family="Carabidae" genus="Omophron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B977EABEFFDA1455FB5EAB4EFAF00817" box="[1178,1292,860,884]" italics="true" pageId="2" pageNumber="21">Omophron</emphasis>
Latr.
</taxonomicName>
, has greatly contributed to the development of carabidology. The name of the species has been coined on the basis of his surname “Valainis”
<emphasis id="B977EABEFFDA1455FCB2ABCEFC220897" box="[886,990,988,1012]" italics="true" pageId="2" pageNumber="21">
<taxonomicName id="4C034D2FFFDA1455FCB2ABCEFC260897" authorityName="Barševskis" authorityYear="2010" box="[886,986,988,1012]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="valainisi">valainisi</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="DF7C6624FFDA1455FE96AC48FD390F11" box="[338,709,1114,1138]" pageId="2" pageNumber="21" startId="2.[338,378,1114,1138]" targetType="figure">
<paragraph id="8BBC36ACFFDA1455FE96AC48FD390F11" blockId="2.[338,709,1114,1138]" box="[338,709,1114,1138]" pageId="2" pageNumber="21">
Fig. 1.
<taxonomicName id="4C034D2FFFDA1455FE59AC49FD7B0F11" authorityName="Barševskis" authorityYear="2010" box="[413,647,1114,1138]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="valainisi" status="sp. nov.">Broscosoma valainisi</taxonomicName>
<taxonomicNameLabel id="A24457C5FFDA1455FD49AC49FD390F11" box="[653,709,1115,1138]" pageId="2" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BBC36ACFFDA1455FE96ACA3FE220E2A" blockId="2.[338,849,1201,1385]" pageId="2" pageNumber="21">first punctate line is impressed in and markedly punctate and it reaches the apex of the elytra. The rest of the punctate lines are smoothed down. The correlation index of the elytra length and width is 1.49.</paragraph>
<paragraph id="8BBC36ACFFDA1455FE96AD43FDB70E0A" blockId="2.[338,849,1201,1385]" box="[338,587,1361,1385]" pageId="2" pageNumber="21">The male is not known.</paragraph>
<paragraph id="8BBC36ACFFDA1455FE96AD80FCB30DEA" blockId="2.[338,849,1425,1673]" pageId="2" pageNumber="21">
<emphasis id="B977EABEFFDA1455FE96AD80FE1A0EC9" bold="true" box="[338,486,1426,1450]" pageId="2" pageNumber="21">Comparison.</emphasis>
This species is very similar to
<taxonomicName id="4C034D2FFFDA1455FCFCAD80FDAF0EAA" authority="Deuve, 1990" authorityName="Deuve" authorityYear="1990" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="sichuanum">
<emphasis id="B977EABEFFDA1455FCFCAD80FE3B0EAA" italics="true" pageId="2" pageNumber="21">B. sichuanum</emphasis>
<bibRefCitation id="EF924B5DFFDA1455FE08ADA0FDAF0EAA" author="Deuve T." box="[460,595,1457,1481]" pageId="2" pageNumber="21" refId="ref1745" refString="Deuve T. 1990. Nouveaux Carabidae et Broscidae des montagnes tibeto-himalayennes" type="book" year="1990">Deuve, 1990</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C034D2FFFDA1455FD41ADA0FE120E8A" authority="Sciaky &amp; Facchini, 2005" authorityName="Sciaky &amp; Facchini" authorityYear="2005" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="stefani">
<emphasis id="B977EABEFFDA1455FD41ADA0FD150EAA" box="[645,745,1457,1481]" italics="true" pageId="2" pageNumber="21">B. stefani</emphasis>
<bibRefCitation id="EF924B5DFFDA1455FD2BADA3FE120E8A" author="Sciaky R. &amp; Facchini S." pageId="2" pageNumber="21" pagination="1 - 12" refId="ref1942" refString="Sciaky R. &amp; Facchini S. 2005. Revision of the Chinese Broscosoma Rosenhauer, 1846, with descriptions of two new species (Coleoptera: Carabidae, Broscinae). Koleopterologische Rundschau, 75: 1 - 12." type="journal article" year="2005">Sciaky &amp; Facchini, 2005</bibRefCitation>
</taxonomicName>
. It differs from the former one with the different colouring and more slender and extended habitus. The colour of
<taxonomicName id="4C034D2FFFDA1455FD7EAE00FE6B0D2A" authority="Deuve" authorityName="Deuve" authorityYear="1990" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="sichuanum">
<emphasis id="B977EABEFFDA1455FD7EAE00FCAC0D4A" box="[698,848,1553,1577]" italics="true" pageId="2" pageNumber="21">B. sichuanum</emphasis>
Deuve
</taxonomicName>
is green or bluish green, head with collar constriction impunctate or sparsely punctate, but the colouring of
<emphasis id="B977EABEFFDA1455FE3AAE60FD530DEA" box="[510,687,1649,1673]" italics="true" pageId="2" pageNumber="21">
<taxonomicName id="4C034D2FFFDA1455FE3AAE60FD840DEA" authorityName="Barševskis" authorityYear="2010" box="[510,632,1649,1673]" class="Insecta" family="Carabidae" genus="Broscosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="valainisi" status="sp.n.">B. valainisi</taxonomicName>
<taxonomicNameLabel id="A24457C5FFDA1455FDB9AE60FD530DEA" box="[637,687,1650,1673]" pageId="2" pageNumber="21" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is black-brown
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,239 @@
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<mods:title id="5C0A7B6AEB50D4F91DE0CE31DF817A02">A new species of the genus Trichomacrocyrtus Yoshitake, 2018 (Curculionidae: Entiminae: Pachyrhynchini) from Luzon Island, Philippines</mods:title>
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<treatment id="03B587ECFFB0FFD4277E9794A214E825" LSID="urn:lsid:plazi:treatment:03B587ECFFB0FFD4277E9794A214E825" httpUri="http://treatment.plazi.org/id/03B587ECFFB0FFD4277E9794A214E825" lastPageId="3" lastPageNumber="440" pageId="1" pageNumber="438">
<subSubSection id="C3066571FFB0FFD6277E9794A0DBEFA9" box="[113,486,395,419]" pageId="1" pageNumber="438" type="nomenclature">
<paragraph id="8BA336FAFFB0FFD6277E9794A0DBEFA9" blockId="1.[113,486,395,450]" box="[113,486,395,419]" pageId="1" pageNumber="438">
<heading id="D0EB8196FFB0FFD6277E9794A0DBEFA9" bold="true" box="[113,486,395,419]" fontSize="10" level="2" pageId="1" pageNumber="438" reason="4">
<emphasis id="B968EAE8FFB0FFD6277E9794A0DBEFA9" bold="true" box="[113,486,395,419]" pageId="1" pageNumber="438">
<taxonomicName id="4C1C4D79FFB0FFD6277E9794A0B7EFA9" ID-CoL="B7QV3" authorityName="Rukmane-Brbale" authorityYear="2022" box="[113,394,395,419]" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="nubes" status="sp. nov.">
<emphasis id="B968EAE8FFB0FFD6277E9794A0B7EFA9" bold="true" box="[113,394,395,419]" italics="true" pageId="1" pageNumber="438">Trichomacrocyrtus nubes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5793FFB0FFD6269E9794A0DBEFA9" box="[401,486,395,419]" pageId="1" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3066571FFB0FFD6277E97B4A1E1EFC8" box="[113,220,427,450]" pageId="1" pageNumber="438" type="description">
<paragraph id="8BA336FAFFB0FFD6277E97B4A1E1EFC8" blockId="1.[113,486,395,450]" box="[113,220,427,450]" pageId="1" pageNumber="438">
(
<figureCitation id="13272A7FFFB0FFD6277697B4A185EFC8" box="[121,184,427,450]" captionStart="Figure 1" captionStartId="2.[151,221,1633,1656]" captionTargetBox="[156,1169,517,1600]" captionTargetId="figure-21@2.[151,1172,513,1602]" captionTargetPageId="2" captionText="Figure 1. Trichomacrocyrtus nubes sp. nov. A Holotype male, dorsal view. B paratype female, dorsal view." pageId="1" pageNumber="438">Fig. 1</figureCitation>
,
<figureCitation id="13272A7FFFB0FFD627C997B4A1E9EFC8" box="[198,212,427,450]" captionStart="Figure 2" captionStartId="3.[113,183,908,931]" captionTargetBox="[113,1132,153,874]" captionTargetId="figure-234@3.[113,1134,151,876]" captionTargetPageId="3" captionText="Figure 2. AC. Male genitalia of Trichomacrocyrtus nubes sp. nov. A aedeagus in lateral view, B aedeagus in dorsal view, C sternite IX in dorsal view; DE. Female genitalia of Trichomacrocyrtus nubes sp. nov., D sternite VIII in ventral view, E ovipositor in dorsal view, F spermatheca." pageId="1" pageNumber="438">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066571FFB0FFD6277E97FDA0E7EDDB" pageId="1" pageNumber="438" type="materials_examined">
<paragraph id="8BA336FAFFB0FFD6277E97FDA1E9ECB3" blockId="1.[113,605,482,697]" pageId="1" pageNumber="438">
<emphasis id="B968EAE8FFB0FFD6277E97FDA01EEFF0" bold="true" box="[113,291,482,506]" pageId="1" pageNumber="438">Type material.</emphasis>
<materialsCitation id="3B743CA7FFB0FFD6262197FDA1E9ECB3" collectingDate="2020-05" collectionCode="DUBC" country="Philippines" county="Bontoc" location="Luzon" municipality="Barlig" pageId="1" pageNumber="438" specimenCount="2" typeStatus="holotype">
<typeStatus id="54A78858FFB0FFD6262197FDA0ABEFF3" box="[302,406,482,505]" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
, male: “
<collectingCountry id="F30B766AFFB0FFD626F297FDA186EC13" name="Philippines" pageId="1" pageNumber="438">PHILIPPINES</collectingCountry>
/
<location id="8EC36021FFB0FFD627C3941DA02EEC13" LSID="urn:lsid:plazi:treatment:03B587ECFFB0FFD4277E9794A214E825:8EC36021FFB0FFD627C3941DA02EEC13" box="[204,275,514,537]" country="Philippines" county="Bontoc" municipality="Barlig" name="Luzon" pageId="1" pageNumber="438">Luzon</location>
,
<collectingMunicipality id="6BC7AC80FFB0FFD62613941DA05CEC13" box="[284,353,514,537]" pageId="1" pageNumber="438">Barlig</collectingMunicipality>
,
<collectingCounty id="62C24E76FFB0FFD62665941DA08AEC13" box="[362,439,514,537]" pageId="1" pageNumber="438">Bontoc</collectingCounty>
/
<date id="FFA2103AFFB0FFD626C8941DA325EC13" box="[455,536,514,537]" pageId="1" pageNumber="438" value="2020-05">
<collectingDate id="EFE6E9D2FFB0FFD626C8941DA325EC13" box="[455,536,514,537]" pageId="1" pageNumber="438" value="2020-05">V. 2020</collectingDate>
</date>
/ local collector leg.” (white rectangular label, printed); “
<typeStatus id="54A78858FFB0FFD627BF945DA002EC53" box="[176,319,578,601]" pageId="1" pageNumber="438" type="holotype">HOLOTYPE</typeStatus>
/ male /
<taxonomicName id="4C1C4D79FFB0FFD6269B945DA193EC73" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="nubes">
<emphasis id="B968EAE8FFB0FFD6269B945DA193EC73" italics="true" pageId="1" pageNumber="438">Trichomacrocyrtus nubes</emphasis>
</taxonomicName>
/ Rukmane­Brbale 2022 / det. RukmaneBrbale 2022” (red rectangular label, printed) (
<collectionCode id="ED0DAE3FFFB0FFD6277694BDA1F4ECB3" box="[121,201,674,697]" pageId="1" pageNumber="438">DUBC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA336FAFFB0FFD6277E94C6A0E7EDDB" blockId="1.[113,605,729,977]" pageId="1" pageNumber="438">
<materialsCitation id="3B743CA7FFB0FFD6277E94C6A0E8EDDB" pageId="1" pageNumber="438" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54A78858FFB0FFD6277E94C6A1E8ECFA" box="[113,213,729,752]" pageId="1" pageNumber="438" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D1AFD73FFB0FFD627EB94C6A015ECFA" box="[228,296,729,752]" count="1" pageId="1" pageNumber="438" type="male">1 male</specimenCount>
,
<specimenCount id="9D1AFD73FFB0FFD6263F94C6A0B1ECFA" box="[304,396,729,752]" count="1" pageId="1" pageNumber="438" type="female">1 female</specimenCount>
): Both with the same white rectangular printed locality label as
<typeStatus id="54A78858FFB0FFD6252A94E6A19CED3A" pageId="1" pageNumber="438" type="holotype">holotype</typeStatus>
; with the following red printed labels: “
<typeStatus id="54A78858FFB0FFD6253B9506A1EBED5A" pageId="1" pageNumber="438" type="paratype">PARATYPE</typeStatus>
/ male /
<taxonomicName id="4C1C4D79FFB0FFD626329526A372ED5A" authorityName="Rukmane-Brbale" authorityYear="2022" box="[317,591,825,848]" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="nubes">
<emphasis id="B968EAE8FFB0FFD626329526A372ED5A" box="[317,591,825,848]" italics="true" pageId="1" pageNumber="438">Trichomacrocyrtus nubes</emphasis>
</taxonomicName>
/ Rukmane­Brbale 2022 / det. Rukmane­Brbale 2022” (); “
<typeStatus id="54A78858FFB0FFD6262F9566A09BED9A" box="[288,422,889,912]" pageId="1" pageNumber="438" type="paratype">PARATYPE</typeStatus>
/ female /
<taxonomicName id="4C1C4D79FFB0FFD625009566A006EDBB" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="nubes">
<emphasis id="B968EAE8FFB0FFD625009566A006EDBB" italics="true" pageId="1" pageNumber="438">Trichomacrocyrtus nubes</emphasis>
</taxonomicName>
/ Rukmane­Brbale 2022 / det. Rukmane­Brbale 2022” ()
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066571FFB0FFD4277E95EDA21EEB74" lastPageId="3" lastPageNumber="440" pageId="1" pageNumber="438" type="description">
<paragraph id="8BA336FAFFB0FFD6277E95EDA012EB00" blockId="1.[113,608,1010,1290]" pageId="1" pageNumber="438">
<emphasis id="B968EAE8FFB0FFD6277E95EDA1C1EA00" bold="true" box="[113,252,1010,1034]" pageId="1" pageNumber="438">Description.</emphasis>
Male. Measurements: LB: 13.1­ 15.8 (
<typeStatus id="54A78858FFB0FFD627A2920CA02FEA20" box="[173,274,1043,1066]" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
15.8; mean 14.45); LP: 3.7­ 4.6 (
<typeStatus id="54A78858FFB0FFD62777922CA1E1EA40" box="[120,220,1075,1098]" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
4.6; mean 4.3); LE: 10.0­ 11.7 (
<typeStatus id="54A78858FFB0FFD62510922CA1A2EA60" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
11.7; mean 10.85); WE: 6.4­ 8.3 (
<typeStatus id="54A78858FFB0FFD626F7924CA361EA60" box="[504,604,1107,1130]" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
8.3; mean 7.35); WP: 3.9­ 4.4 (
<typeStatus id="54A78858FFB0FFD626CA926CA317EA80" box="[453,554,1139,1162]" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
4.4; mean 4.15); LR: 2.3­ 3.1 (
<typeStatus id="54A78858FFB0FFD62689928CA0D6EAA0" box="[390,491,1171,1194]" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
3.1; mean 2.7); WR: 2.1­ 2.3 (
<typeStatus id="54A78858FFB0FFD6264992ACA096EAC0" box="[326,427,1203,1226]" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
2.3; mean 2.2). N=2 for all measurements. Dorsal habitus as shown in
<figureCitation id="13272A7FFFB0FFD627D892ECA016EB00" box="[215,299,1267,1290]" captionStart="Figure 1" captionStartId="2.[151,221,1633,1656]" captionTargetBox="[156,1169,517,1600]" captionTargetId="figure-21@2.[151,1172,513,1602]" captionTargetPageId="2" captionText="Figure 1. Trichomacrocyrtus nubes sp. nov. A Holotype male, dorsal view. B paratype female, dorsal view." pageId="1" pageNumber="438">Fig. 1A</figureCitation>
.
</paragraph>
<paragraph id="8BA336FAFFB0FFD6277E9336A191E84A" blockId="1.[113,605,1321,1600]" pageId="1" pageNumber="438">Integument black, very shiny, underside with weaker lustre; markings of green round to recumbent scales on elytra, pronotum, rostrum, femur and underside; metasternum densely covered with green round scales, mingled with long, light hairs, strongly pubescent; elongated patch of scales on each side of ventrite I; rest of the ventrites without scales, mingled with long light hairs.</paragraph>
<paragraph id="8BA336FAFFB0FFD6277E907FA3D6ED94" blockId="1.[113,605,1632,1687]" lastBlockId="1.[643,1134,155,926]" pageId="1" pageNumber="438">Head sub ovate, nearly smooty except area between eyes with stronger puncture and rugosity; genae without scales, mingled with short light hairs; forehead strongly wrinkled between eyes, nearly 3 times as wide as eye width; patch of green round scales from the middle of forehead to medial part of rostrum; rostrum slightly pubescent at basal halh, then strongly pubescent to apex; short light hairs from antennal scape to labrum; in dorsal contour narrowed at subbasal part, slightly widened to medial part and straigth to apical part, subapical part slightly narrowed; dorsally with longitudinal groove from middle of rostrum to medial part of forehead, transverse groove at subbasal part and deep sub ovate depression from apical half to base of the forehead, apical part with moderate bulge; eyes not convex, relatively small, not prominent from the outline of the head. Antennae slender; segment I nearly 1.5 times as long as segment II, 2 times as long as wide, segment II 1.5 times longer than segment III, 1.5 times longer than wide, segments IIIVII sub­equal in size, gradually increasing to club, wider than long, club lanceolate, nearly 2 times longer than wide; longer than wide, LR/ WR 1.35.</paragraph>
<paragraph id="8BA336FAFFB0FFD6258C95A1A3E9EB36" blockId="1.[643,1134,958,1340]" pageId="1" pageNumber="438">Prothorax sub cylindrical, slightly longer than wide, LP/WP 1.05, with the following scaly markings: 1) transverse line of scales slightly before the middle circumsribing prothorax; 2) longitudinal line on each dorso­lateral parts from medial part to dorsal edge of prothorax; 3) lagre scally patch on vantral part; dorsal part finely punctured, slightly pubescent; widest at the middle, in dorsal contour narrowest along apical margin, increased to widest midline, then rounded and decreased to basal 1/5, straight to basal margin.</paragraph>
<paragraph id="8BA336FAFFB0FFD5258C9343A1F3EFD8" blockId="1.[643,1134,1372,1687]" lastBlockId="2.[151,643,155,466]" lastPageId="2" lastPageNumber="439" pageId="1" pageNumber="438">
Elytra sub­lanceolate, intervals smooth, very slightly punctured, without pubescence except apex with short light hairs; widest in the middle; in dorsal contour increased from basal margin to widest middle, rounded and decreased to apical 1/2 and more strongly decreased to extended apex; each elytron with the following scaly markings: 1) longitudinal line of scales from interval III to IV, slightly wider at base, decreasing in apical direction, from basal margin to basal 1/2; 2) longitudinal line of scales along lateral edge in all length; 3) irregular, cloud­like shape patch on medial part of elytra, from suture to lateral edge in width and from basal 3/4 to apical 1/
<quantity id="4CE49B1FFFB3FFD5257B96E4A191EF38" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="439" unit="in" value="2.0">2 in</quantity>
length, patch bare in the middle; 4) longitudinal line from middle on interval II to interval III from apical 1/2 to apex; LE/WE: 1.41; nearly twice as wide as prothorax, WE/WP: 1.89; more than two times as long as prothorax, LE/LP: 2.54.
</paragraph>
<paragraph id="8BA336FAFFB3FFD525A79684A562EF92" blockId="2.[680,1172,155,408]" pageId="2" pageNumber="439">Front coxa with fine patch of scales, slightly pubescent; each femora with fine patch of scales on apical part along internal margin, without pubescence, with short light hairs along outer margin; front tibia with short light hairs in all length and longer hairs along internal margin and apical part; mid and hind tibia with long, dense golden hairs along internal margin.</paragraph>
<paragraph id="8BA336FAFFB3FFD425A797A7A1D9EB3F" blockId="2.[680,1086,440,463]" lastBlockId="3.[113,605,1085,1333]" lastPageId="3" lastPageNumber="440" pageId="2" pageNumber="439">
Aedegal body as shown in
<figureCitation id="13272A7FFFB3FFD524C997A7A504EFC5" box="[966,1081,440,463]" captionStart="Figure 2" captionStartId="3.[113,183,908,931]" captionTargetBox="[113,1132,153,874]" captionTargetId="figure-234@3.[113,1134,151,876]" captionTargetPageId="3" captionText="Figure 2. AC. Male genitalia of Trichomacrocyrtus nubes sp. nov. A aedeagus in lateral view, B aedeagus in dorsal view, C sternite IX in dorsal view; DE. Female genitalia of Trichomacrocyrtus nubes sp. nov., D sternite VIII in ventral view, E ovipositor in dorsal view, F spermatheca." pageId="2" pageNumber="439">Fig. 2AC</figureCitation>
. Female. Measurements: LB: 13.9; LP: 3.6; LE: 10.2; WE: 7.9; WP: 4.0; LR: 2.4; WR: 1.9. N=1 for all measurements. Elytra signifficantly wider than male, apex more strongly extended, with two peaks; LE/WE 1.29; scale colour brighter, with ogange and yellow scales. Dorsal habitus as shown in
<figureCitation id="13272A7FFFB2FFD4260D92E1A067EB1F" box="[258,346,1278,1301]" captionStart="Figure 1" captionStartId="2.[151,221,1633,1656]" captionTargetBox="[156,1169,517,1600]" captionTargetId="figure-21@2.[151,1172,513,1602]" captionTargetPageId="2" captionText="Figure 1. Trichomacrocyrtus nubes sp. nov. A Holotype male, dorsal view. B paratype female, dorsal view." pageId="3" pageNumber="440">Fig. 1B</figureCitation>
. Genitalia as shown in
<figureCitation id="13272A7FFFB2FFD4277E9301A1E0EB3F" box="[113,221,1310,1333]" captionStart="Figure 2" captionStartId="3.[113,183,908,931]" captionTargetBox="[113,1132,153,874]" captionTargetId="figure-234@3.[113,1134,151,876]" captionTargetPageId="3" captionText="Figure 2. AC. Male genitalia of Trichomacrocyrtus nubes sp. nov. A aedeagus in lateral view, B aedeagus in dorsal view, C sternite IX in dorsal view; DE. Female genitalia of Trichomacrocyrtus nubes sp. nov., D sternite VIII in ventral view, E ovipositor in dorsal view, F spermatheca." pageId="3" pageNumber="440">Fig. 2DF</figureCitation>
.
</paragraph>
<caption id="DF636672FFB3FFD52798907EA02AE892" pageId="2" pageNumber="439" startId="2.[151,221,1633,1656]" targetBox="[156,1169,517,1600]" targetPageId="2" targetType="figure">
<paragraph id="8BA336FAFFB3FFD52798907EA02AE892" blockId="2.[151,1172,1632,1688]" pageId="2" pageNumber="439">
Figure 1.
<taxonomicName id="4C1C4D79FFB3FFD527F4907EA334E872" authorityName="Rukmane-Brbale" authorityYear="2022" box="[251,521,1633,1656]" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="439" phylum="Arthropoda" rank="species" species="nubes" status="sp. nov.">
<emphasis id="B968EAE8FFB3FFD527F4907EA334E872" box="[251,521,1633,1656]" italics="true" pageId="2" pageNumber="439">Trichomacrocyrtus nubes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5793FFB3FFD52501907EA362E872" box="[526,607,1633,1656]" pageId="2" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B968EAE8FFB3FFD5256B907FA34AE872" bold="true" box="[612,631,1632,1656]" pageId="2" pageNumber="439">A</emphasis>
Holotype male, dorsal view.
<emphasis id="B968EAE8FFB3FFD524B4907FA2F0E872" bold="true" box="[955,973,1632,1656]" pageId="2" pageNumber="439">B</emphasis>
paratype female, dorsal view.
</paragraph>
</caption>
<caption id="DF636672FFB2FFD4277E9593A1C3EA09" pageId="3" pageNumber="440" startId="3.[113,183,908,931]" targetBox="[113,1132,153,874]" targetPageId="3" targetType="figure">
<paragraph id="8BA336FAFFB2FFD4277E9593A1C3EA09" blockId="3.[113,1134,907,1027]" pageId="3" pageNumber="440">
Figure 2.
<emphasis id="B968EAE8FFB2FFD427D89594A036EDA9" bold="true" box="[215,267,907,931]" pageId="3" pageNumber="440">AC</emphasis>
. Male genitalia of
<taxonomicName id="4C1C4D79FFB2FFD426DD9593A3DDEDA9" authorityName="Rukmane-Brbale" authorityYear="2022" box="[466,736,908,931]" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="nubes" status="sp. nov.">
<emphasis id="B968EAE8FFB2FFD426DD9593A3DDEDA9" box="[466,736,908,931]" italics="true" pageId="3" pageNumber="440">Trichomacrocyrtus nubes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5793FFB2FFD425E89593A205EDA9" box="[743,824,908,931]" pageId="3" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B968EAE8FFB2FFD424319594A26CEDA9" bold="true" box="[830,849,907,931]" pageId="3" pageNumber="440">A</emphasis>
aedeagus in lateral view,
<emphasis id="B968EAE8FFB2FFD4277E95B4A1BEEDC9" bold="true" box="[113,131,939,963]" pageId="3" pageNumber="440">B</emphasis>
aedeagus in dorsal view,
<emphasis id="B968EAE8FFB2FFD426BE95B4A0F9EDC9" bold="true" box="[433,452,939,963]" pageId="3" pageNumber="440">C</emphasis>
sternite IX in dorsal view;
<emphasis id="B968EAE8FFB2FFD4240995B4A205EDC9" bold="true" box="[774,824,939,963]" pageId="3" pageNumber="440">DE</emphasis>
. Female genitalia of
<taxonomicName id="4C1C4D79FFB2FFD4231095B3A006EDE9" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="nubes" status="sp. nov.">
<emphasis id="B968EAE8FFB2FFD4231095B3A006EDE9" italics="true" pageId="3" pageNumber="440">Trichomacrocyrtus nubes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5793FFB2FFD4264D95D3A0AEEDE9" box="[322,403,972,995]" pageId="3" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="B968EAE8FFB2FFD426AD95D4A088EDE9" bold="true" box="[418,437,971,995]" pageId="3" pageNumber="440">D</emphasis>
sternite VIII in ventral view,
<emphasis id="B968EAE8FFB2FFD4240995D4A225EDE9" bold="true" box="[774,792,971,995]" pageId="3" pageNumber="440">E</emphasis>
ovipositor in dorsal view,
<emphasis id="B968EAE8FFB2FFD4234695D4A564EDE9" bold="true" box="[1097,1113,971,995]" pageId="3" pageNumber="440">F</emphasis>
spermatheca.
</paragraph>
</caption>
<paragraph id="8BA336FAFFB2FFD4277E934AA21EEB74" blockId="3.[113,606,1365,1582]" lastBlockId="3.[643,1135,1085,1406]" pageId="3" pageNumber="440">
Differential analyses.
<taxonomicName id="4C1C4D79FFB2FFD426579349A093EB67" authorityName="Rukmane-Brbale" authorityYear="2022" box="[344,430,1366,1389]" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="nubes" status="sp. nov.">
<emphasis id="B968EAE8FFB2FFD426579349A093EB67" box="[344,430,1366,1389]" italics="true" pageId="3" pageNumber="440">T. nubes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B5793FFB2FFD426BC934AA33EEB66" box="[435,515,1365,1388]" pageId="3" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
in general appearance is similar to
<taxonomicName id="4C1C4D79FFB2FFD426AC9369A034EBA7" authority="Schultze, 1917" authorityName="Schultze" authorityYear="1917" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="hieroglyphicus">
<emphasis id="B968EAE8FFB2FFD426AC9369A360EB87" box="[419,605,1398,1421]" italics="true" pageId="3" pageNumber="440">T. hieroglyphicus</emphasis>
<bibRefCitation id="EF8D4B0BFFB2FFD4277E9389A034EBA7" author="Schultze, W." box="[113,265,1430,1453]" pageId="3" pageNumber="440" pagination="249 - 259" refId="ref2070" refString="Schultze, W. 1917. Fourth contribution to the Coleoptera fauna of the Philippines. Philippine Journal of Science, Manilla, 12 (4), 249 - 259." type="journal article" year="1917">Schultze, 1917</bibRefCitation>
</taxonomicName>
from the same
<collectingRegion id="49D8F818FFB2FFD426AE9389A367EBA7" box="[417,602,1430,1453]" country="Philippines" name="Benguet" pageId="3" pageNumber="440">Benguet province</collectingRegion>
. The new species are easyly distinguishable by the following features: 1) different pronotal markings: dorsal transverse line is wider and not interrupted in medial part in new species, but narrower and medially interrupted in
<emphasis id="B968EAE8FFB2FFD4231C9222A3D6EA7F" italics="true" pageId="3" pageNumber="440">
<taxonomicName id="4C1C4D79FFB2FFD4231C9222A3D8EA7F" authorityName="Schultze" authorityYear="1917" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="hierogliphicus">T. hierogliphicus</taxonomicName>
;
</emphasis>
2) different elytral markings: longitudinal line along interval II in new species is interrupted by large cloud­like spot, the new species also lack longitudinal line on dorso­lateral part of each elytron, while markings on elytra of
<taxonomicName id="4C1C4D79FFB2FFD4235492FCA221EB11" authorityName="Schultze" authorityYear="1917" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="hierogliphicus">
<emphasis id="B968EAE8FFB2FFD4235492FCA221EB11" italics="true" pageId="3" pageNumber="440">T. hierogliphicus</emphasis>
</taxonomicName>
are solid in all length; 3) bigger eyes in
<emphasis id="B968EAE8FFB2FFD425DA933AA210EB36" box="[725,813,1317,1340]" italics="true" pageId="3" pageNumber="440">
P.
<taxonomicName id="4C1C4D79FFB2FFD425E0933AA210EB36" authorityName="Rukmane-Brbale" authorityYear="2022" box="[751,813,1317,1340]" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="nubes">nubes</taxonomicName>
</emphasis>
; 4) different shape of aedegal body, with more curved apical opening in
<taxonomicName id="4C1C4D79FFB2FFD423549359A220EB74" authorityName="Schultze" authorityYear="1917" class="Insecta" family="Curculionidae" genus="Trichomacrocyrtus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="hierogliphicus">
<emphasis id="B968EAE8FFB2FFD423549359A220EB74" italics="true" pageId="3" pageNumber="440">T. hierogliphicus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066571FFB2FFD4258C9381A56AEBBF" box="[643,1111,1438,1462]" pageId="3" pageNumber="440" type="distribution">
<paragraph id="8BA336FAFFB2FFD4258C9381A56AEBBF" blockId="3.[643,1111,1438,1462]" box="[643,1111,1438,1462]" pageId="3" pageNumber="440">
<emphasis id="B968EAE8FFB2FFD4258C9381A228EBBC" bold="true" box="[643,789,1438,1462]" pageId="3" pageNumber="440">Distribution.</emphasis>
Luzon Island, Barlig, Bontoc.
</paragraph>
</subSubSection>
<subSubSection id="C3066571FFB2FFD4258C93C9A214E825" pageId="3" pageNumber="440" type="etymology">
<paragraph id="8BA336FAFFB2FFD4258C93C9A214E825" blockId="3.[643,1134,1494,1583]" pageId="3" pageNumber="440">
<emphasis id="B968EAE8FFB2FFD4258C93C9A23EEBE4" bold="true" box="[643,771,1494,1518]" pageId="3" pageNumber="440">Etymology.</emphasis>
The new species is named after its cloud­like scally markings on the elytra. Cloud on Latin­nubes.
</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="D0E5A64EFFDA5D72FDCCE38AFC2018B2" bold="true" box="[624,985,404,429]" fontSize="10" level="2" pageId="1" pageNumber="120" reason="4">
<taxonomicName id="4C126AA1FFDA5D72FDCCE38AFC2018B2" ID-CoL="PK32" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" box="[624,985,404,429]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="granosa">
<emphasis id="B966CD30FFDA5D72FDCCE38AFC2018B2" bold="true" box="[624,985,404,429]" pageId="1" pageNumber="120">
<emphasis id="B966CD30FFDA5D72FDCCE38AFCB018B3" bold="true" box="[624,841,405,429]" italics="true" pageId="1" pageNumber="120">Calidiopsis granosa</emphasis>
<bibRefCitation id="EF836CD3FFDA5D72FCF3E38BFC2018B2" author="Heller K. M." box="[847,985,404,428]" pageId="1" pageNumber="120" pagination="3 - 9" refId="ref1360" refString="Heller K. M. 1923. Neue philippinische- Kafer. Stettiner Entomologische Zeitung, 84 (1): 3 - 9." type="journal article" year="1923">Heller, 1923</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAD1122FFDA5D72FDCCE3ACFD1118D5" blockId="1.[624,985,404,459]" box="[624,744,435,459]" pageId="1" pageNumber="120">(Fig. 1,2,3)</paragraph>
</subSubSection>
<caption id="DF6D41AAFFDA5D72FFCEE45EFF631FA0" pageId="1" pageNumber="120" startId="1.[114,184,1601,1625]" targetBox="[129,1015,558,1557]" targetPageId="1" targetType="figure">
<paragraph id="8BAD1122FFDA5D72FFCEE45EFF631FA0" blockId="1.[114,1095,1600,1726]" pageId="1" pageNumber="120">
Figure 1.
<taxonomicName id="4C126AA1FFDA5D72FF65E45EFDB91F47" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" box="[217,576,1601,1625]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="granosa">
<emphasis id="B966CD30FFDA5D72FF65E45EFE4E1F47" box="[217,439,1601,1625]" italics="true" pageId="1" pageNumber="120">Calidiopsis granosa</emphasis>
<bibRefCitation id="EF836CD3FFDA5D72FE0BE45EFDB91F47" author="Heller K. M." box="[439,576,1601,1625]" pageId="1" pageNumber="120" pagination="3 - 9" refId="ref1360" refString="Heller K. M. 1923. Neue philippinische- Kafer. Stettiner Entomologische Zeitung, 84 (1): 3 - 9." type="journal article" year="1923">Heller, 1923</bibRefCitation>
</taxonomicName>
.
<emphasis id="B966CD30FFDA5D72FDF6E45FFDA41F46" bold="true" box="[586,605,1600,1624]" pageId="1" pageNumber="120">A</emphasis>
male, dorsal view.
<emphasis id="B966CD30FFDA5D72FCFEE45FFCAD1F46" bold="true" box="[834,852,1600,1624]" pageId="1" pageNumber="120">B</emphasis>
female, dorsal view. 120
</paragraph>
</caption>
<caption id="DF6D41AAFFD95D71FF01E65EFD791D69" pageId="2" pageNumber="121" startId="2.[189,259,1089,1113]" targetBox="[220,652,188,1067]" targetPageId="2" targetType="figure">
<paragraph id="8BAD1122FFD95D71FF01E65EFD791D69" blockId="2.[189,1179,1089,1143]" pageId="2" pageNumber="121">
Figure 2. Male genitalia of
<taxonomicName id="4C126AA1FFD95D71FE4EE65EFC9A1D47" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" box="[498,867,1089,1113]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="granosa">
<emphasis id="B966CD30FFD95D71FE4EE65EFD341D47" box="[498,717,1089,1113]" italics="true" pageId="2" pageNumber="121">Calidiopsis granosa</emphasis>
<bibRefCitation id="EF836CD3FFD95D71FD6BE65EFC9A1D47" author="Heller K. M." box="[727,867,1089,1113]" pageId="2" pageNumber="121" pagination="3 - 9" refId="ref1360" refString="Heller K. M. 1923. Neue philippinische- Kafer. Stettiner Entomologische Zeitung, 84 (1): 3 - 9." type="journal article" year="1923">Heller, 1923</bibRefCitation>
</taxonomicName>
. Aedeagus in lateral view, sternite IX in dorsal view. Scale bar 1mm.
</paragraph>
</caption>
<subSubSection id="C30842A9FFD95D77FF01E6A4FE15185A" lastPageId="4" lastPageNumber="123" pageId="2" pageNumber="121" type="description">
<paragraph id="8BAD1122FFD95D71FF01E6A4FE341F3A" blockId="2.[189,668,1211,1572]" pageId="2" pageNumber="121">
<emphasis id="B966CD30FFD95D71FF01E6A4FE211DCD" bold="true" box="[189,472,1211,1235]" pageId="2" pageNumber="121">Original description:</emphasis>
<taxonomicName id="4C126AA1FFD95D71FDA0E6A4FEE21DEC" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="granosa" status="sp. nov.">
<emphasis id="B966CD30FFD95D71FDA0E6A4FEE21DEC" italics="true" pageId="2" pageNumber="121">Calidiopsis granosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255704BFFD95D71FEA1E6C5FEAE1DEC" box="[285,343,1242,1266]" pageId="2" pageNumber="121" rank="species">sp. n.</taxonomicNameLabel>
<collectingCountry id="F30551B2FFD95D71FEE3E6C5FE661DEC" box="[351,415,1242,1266]" name="Niger" pageId="2" pageNumber="121">Niger</collectingCountry>
, parcenigrosetosa, rostro dorso dense albido, prothorace elytrisque ubique, illosqamulis minutissimis parcisque, his squamulis paulomaioribuc in loculis acervatis, canis; prothorace granulis magnitudine diversa dense obsito; elytrisseriato-granulatis, granulisnitidis, spatium sigulum explentibus, pedibus subtiliter glauco-squamosis. Long. 9-10,5, lat. 3,5-4,5 mm. Insula: Basilan (N. 7314, 15 110, 13 178). (
<bibRefCitation id="EF836CD3FFD95D71FE8AE413FE391F3A" author="Heller K. M." box="[310,448,1548,1572]" pageId="2" pageNumber="121" pagination="3 - 9" refId="ref1360" refString="Heller K. M. 1923. Neue philippinische- Kafer. Stettiner Entomologische Zeitung, 84 (1): 3 - 9." type="journal article" year="1923">Heller, 1923</bibRefCitation>
).
</paragraph>
<paragraph id="8BAD1122FFD95D71FD07E6A4FBA71F18" blockId="2.[699,1179,1211,1542]" pageId="2" pageNumber="121">
<emphasis id="B966CD30FFD95D71FD07E6A4FC841DCD" bold="true" box="[699,893,1211,1235]" pageId="2" pageNumber="121">Notes on types:</emphasis>
<materialsCitation id="3B7A1B7FFFD95D71FC39E6A4FBA71F18" collectionCode="MTD" country="Germany" county="Typus'" location="Dresden" municipality="Baker'" pageId="2" pageNumber="121" specimenCount="2" typeStatus="holotype">
The original description was based on
<specimenCount id="9D14DAABFFD95D71FCEEE6C5FC081DEC" box="[850,1009,1242,1266]" count="2" pageId="2" pageNumber="121" type="generic">two specimens</specimenCount>
now deposited in
<collectionCode id="ED0389E7FFD95D71FD5CE6E6FCE21C0F" box="[736,795,1273,1297]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="2" pageNumber="121">MTD</collectionCode>
(
<location id="8ECD47F9FFD95D71FC8FE6E6FC691C0F" LSID="urn:lsid:plazi:treatment:03BBA034FFDA5D77FDCCE38AFE5C1B24:8ECD47F9FFD95D71FC8FE6E6FC691C0F" box="[819,912,1273,1297]" country="Germany" county="Typus'" municipality="Baker'" name="Dresden" pageId="2" pageNumber="121">Dresden</location>
,
<collectingCountry id="F30551B2FFD95D71FC18E6E6FBF01C0F" box="[932,1033,1273,1297]" name="Germany" pageId="2" pageNumber="121">Germany</collectingCountry>
) both from
<location id="8ECD47F9FFD95D71FD07E708FC4D1C31" LSID="urn:lsid:plazi:treatment:03BBA034FFDA5D77FDCCE38AFE5C1B24:8ECD47F9FFD95D71FD07E708FC4D1C31" box="[699,948,1303,1327]" country="Germany" county="Typus'" municipality="Baker'" name="Island of Basilan. The" pageId="2" pageNumber="121">Island of Basilan. The</location>
label data of
<typeStatus id="54A9AF80FFD95D71FBEAE708FD101C50" pageId="2" pageNumber="121" type="holotype">Holotype</typeStatus>
include five labels: 1922 / 6 (yellow rectangular label); Island of /
<location id="8ECD47F9FFD95D71FBF8E74BFB631C72" LSID="urn:lsid:plazi:treatment:03BBA034FFDA5D77FDCCE38AFE5C1B24:8ECD47F9FFD95D71FBF8E74BFB631C72" box="[1092,1178,1364,1388]" country="Germany" county="Typus'" municipality="Baker'" name="Basilan" pageId="2" pageNumber="121">Basilan</location>
/
<collectingMunicipality id="6BC98B58FFD95D71FD5CE76CFCD41C95" box="[736,813,1395,1419]" pageId="2" pageNumber="121">Baker</collectingMunicipality>
(brown rectangular label); 7314 (brown rectangular label);
<emphasis id="B966CD30FFD95D71FBCFE78DFCEA1CD7" italics="true" pageId="2" pageNumber="121">
<taxonomicName id="4C126AA1FFD95D71FBC5E78DFCEA1CD7" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="granosa">C. granosa</taxonomicName>
</emphasis>
/
<collectingCounty id="62CC69AEFFD95D71FC8FE7AEFC7A1CD7" box="[819,899,1457,1481]" pageId="2" pageNumber="121">Typus</collectingCounty>
(red rectangular label); Staatl. Museum fur / Tierkunde. Dresden (white rectangular label) (
<figureCitation id="13290DA7FFD95D71FBB3E7F1FBA91F18" box="[1039,1104,1518,1542]" captionStart="Figure 3" captionStartId="3.[114,184,880,904]" captionTargetBox="[114,1097,171,873]" captionTargetId="figure-305@3.[114,1097,171,873]" captionTargetPageId="3" captionText="Figure 3.Calidiopsis granosa Heller, 1923: Paratype (MTD)." pageId="2" pageNumber="121">Fig. 3</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAD1122FFD95D71FBD0E4B9FB6D1FA0" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="121">121</paragraph>
<paragraph id="8BAD1122FFD85D70FDB8E27CFD411968" blockId="3.[516,696,99,118]" box="[516,696,99,118]" pageId="3" pageNumber="122">
<emphasis id="B966CD30FFD85D70FDB8E27CFD411968" box="[516,696,99,118]" italics="true" pageId="3" pageNumber="122">Rukmane-Bārbale A.</emphasis>
</paragraph>
<caption id="DF6D41AAFFD85D70FFCEE16FFCFA1A96" box="[114,771,880,904]" pageId="3" pageNumber="122" startId="3.[114,184,880,904]" targetBox="[114,1097,171,873]" targetPageId="3" targetType="figure">
<paragraph id="8BAD1122FFD85D70FFCEE16FFCFA1A96" blockId="3.[114,771,880,904]" box="[114,771,880,904]" pageId="3" pageNumber="122">
Figure 3.
<taxonomicName id="4C126AA1FFD85D70FF65E16FFDB81A96" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" box="[217,577,880,904]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="granosa">
<emphasis id="B966CD30FFD85D70FF65E16FFE481A96" box="[217,433,880,904]" italics="true" pageId="3" pageNumber="122">Calidiopsis granosa</emphasis>
<bibRefCitation id="EF836CD3FFD85D70FE0BE16FFDB81A96" author="Heller K. M." box="[439,577,880,904]" pageId="3" pageNumber="122" pagination="3 - 9" refId="ref1360" refString="Heller K. M. 1923. Neue philippinische- Kafer. Stettiner Entomologische Zeitung, 84 (1): 3 - 9." type="journal article" year="1923">Heller, 1923</bibRefCitation>
</taxonomicName>
: Paratype (MTD).
</paragraph>
</caption>
<paragraph id="8BAD1122FFD85D70FFCEE1D4FDB21F76" blockId="3.[114,593,971,1640]" pageId="3" pageNumber="122">
<emphasis id="B966CD30FFD85D70FFCEE1D4FDFA1AFD" bold="true" box="[114,515,971,995]" pageId="3" pageNumber="122">External morphological features:</emphasis>
Male:
<emphasis id="B966CD30FFD85D70FFCEE1F5FF541D1C" bold="true" box="[114,173,1002,1026]" pageId="3" pageNumber="122">Body</emphasis>
black, with markings of single light scales on rostrum dorsally and elytra; femur, tibia and tarsus bare black or marked with single, light scales of round shape; antennae bare or marked with single, light scales of round shape;
<emphasis id="B966CD30FFD85D70FE3EE69CFE1F1D85" bold="true" box="[386,486,1155,1179]" pageId="3" pageNumber="122">Rostrum</emphasis>
smooth, widened to apex, with light pubescence to longer, light hairs along impressions; deep medial longitudinal groove dorsally; two deep impressions at base near the eyes; two large impressions along antennal scape.
<emphasis id="B966CD30FFD85D70FFCEE724FF5E1C4D" bold="true" box="[114,167,1339,1363]" pageId="3" pageNumber="122">Eyes</emphasis>
large, strongly bulging from the outline of the head, two times as wide as forehead each.
<emphasis id="B966CD30FFD85D70FF44E767FE951C8E" bold="true" box="[248,364,1400,1424]" pageId="3" pageNumber="122">Antennae</emphasis>
with scape slightly longer than funicle and club, scape with long setae on all length, widened to apex;
<emphasis id="B966CD30FFD85D70FFCEE7CAFF1E1CF3" bold="true" box="[114,231,1493,1517]" pageId="3" pageNumber="122">Prothorax</emphasis>
strongly rugose, in dorsal contour widest just before the middle, in lateral contour curved along apical 1/2; without markings of light scales;
<emphasis id="B966CD30FFD85D70FE38E42FFE2A1F56" bold="true" box="[388,467,1584,1608]" pageId="3" pageNumber="122">Elytra</emphasis>
ellipsoidal, strongly granulated, with expressed longi-
</paragraph>
<paragraph id="8BAD1122FFD85D70FFCEE4B9FC181DA4" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1103,972,1210]" pageId="3" pageNumber="122">
122 tudinal intervals; in dorsal contour widest just at the middle, in lateral contour gradually rounded, peak just at the middle; apical half of elytra with single setae; rare single light scales of round shape mingled on all length, density of scales vary along specimens. Dorsal habitus as showed in
<figureCitation id="13290DA7FFD85D70FC53E69CFBB31D85" box="[1007,1098,1155,1179]" captionStart="Figure 1" captionStartId="1.[114,184,1601,1625]" captionTargetBox="[129,1015,558,1557]" captionTargetId="figure-100@1.[113,1037,538,1594]" captionTargetPageId="1" captionText="Figure 1.Calidiopsis granosaHeller, 1923. A male, dorsal view.B female, dorsal view. 120" pageId="3" pageNumber="122">Fig. 1A</figureCitation>
. Aedegal body as showed in
<figureCitation id="13290DA7FFD85D70FC20E6BDFC241DA4" box="[924,989,1186,1210]" captionStart="Figure 2" captionStartId="2.[189,259,1089,1113]" captionTargetBox="[220,652,188,1067]" captionTargetId="figure-168@2.[188,672,170,1082]" captionTargetPageId="2" captionText="Figure 2. Male genitalia of Calidiopsis granosa Heller, 1923. Aedeagus in lateral view, sternite IX in dorsal view. Scale bar 1mm." pageId="3" pageNumber="122">Fig. 2</figureCitation>
.
</paragraph>
<paragraph id="8BAD1122FFD85D70FDCCE6FFFCD71C6C" blockId="3.[624,1103,1248,1394]" pageId="3" pageNumber="122">
Female: Bigger, prothorax and elytra more strongly rounded, hairs on legs (femur, tibia, tarsus), antenae and elytra longer, dense. Otherwise as in males. Dorsal habitus as shown in
<figureCitation id="13290DA7FFD85D70FD64E745FCD31C6C" box="[728,810,1370,1394]" captionStart="Figure 1" captionStartId="1.[114,184,1601,1625]" captionTargetBox="[129,1015,558,1557]" captionTargetId="figure-100@1.[113,1037,538,1594]" captionTargetPageId="1" captionText="Figure 1.Calidiopsis granosaHeller, 1923. A male, dorsal view.B female, dorsal view. 120" pageId="3" pageNumber="122">Fig. 1B</figureCitation>
.
</paragraph>
<paragraph id="8BAD1122FFD85D77FDCCE7A9FE15185A" blockId="3.[624,1103,1462,1640]" lastBlockId="4.[189,668,178,324]" lastPageId="4" lastPageNumber="123" pageId="3" pageNumber="122">
<emphasis id="B966CD30FFD85D70FDCCE7A9FCDF1CD0" bold="true" box="[624,806,1462,1486]" pageId="3" pageNumber="122">Measurements:</emphasis>
Male: LB: 6.8 - 8.6 (mean 7.7), LE: 4.5 - 5.9 (mean 5.2), LP: 2.1 - 2.6 (mean 2.4), LR: 1.0 - 1.3 (mean 1.2), WE: 3.7 - 4.1 (mean 3.9), WP: 1.9 - 2.0 (mean 1.95), WR: 0.8 - 1.0 (mean 0.9) (N=4 for all measurements); Female: LB: 8.5 9.2 (mean 8.8), LE: 6.0 6.7 (mean 6.4), LP: 2.8 2.9 (mean 2.85), LR: 1.2 - 1.4 (mean 1.3), WE: 4.6 4.8 (mean 4.7), WP: 2.3 2.6 (mean 2.45), WR: 1.1 1.3 (mean 1.2) (N=3 for all measurements).
</paragraph>
</subSubSection>
<subSubSection id="C30842A9FFDF5D77FF01E376FD9A18C0" pageId="4" pageNumber="123" type="materials_examined">
<paragraph id="8BAD1122FFDF5D77FF01E376FD9A18C0" blockId="4.[189,668,361,478]" pageId="4" pageNumber="123">
<emphasis id="B966CD30FFDF5D77FF01E376FE50189F" bold="true" box="[189,425,361,385]" pageId="4" pageNumber="123">Material examined:</emphasis>
<materialsCitation id="3B7A1B7FFFDF5D77FE10E375FDA618C0" collectingDate="2022-08" country="Philippines" location="Siocon" municipality="Mindanao" pageId="4" pageNumber="123" specimenCount="14" specimenCount-female="3" specimenCount-male="4" stateProvince="Zamboanga">
<specimenCount id="9D14DAABFFDF5D77FE10E375FE1C189C" box="[428,485,362,386]" count="7" pageId="4" pageNumber="123" type="generic">7 ex.</specimenCount>
(
<specimenCount id="9D14DAABFFDF5D77FE4BE375FDA8189C" box="[503,593,362,386]" count="4" pageId="4" pageNumber="123" type="male">4 males</specimenCount>
,
<specimenCount id="9D14DAABFFDF5D77FDDCE375FF0418BE" count="3" pageId="4" pageNumber="123" type="female">3 females</specimenCount>
)
<collectingCountry id="F30551B2FFDF5D77FEA4E397FE4618BE" box="[280,447,392,416]" name="Philippines" pageId="4" pageNumber="123">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="6BC98B58FFDF5D77FE62E397FDB418BE" box="[478,589,392,416]" pageId="4" pageNumber="123">Mindanao</collectingMunicipality>
,
<collectingRegion id="49D6DFC0FFDF5D77FDE0E397FEF418A1" country="Philippines" name="Zamboanga" pageId="4" pageNumber="123">Zamboanga</collectingRegion>
,
<location id="8ECD47F9FFDF5D77FEA4E3B8FE9B18A1" LSID="urn:lsid:plazi:treatment:03BBA034FFDA5D77FDCCE38AFE5C1B24:8ECD47F9FFDF5D77FEA4E3B8FE9B18A1" box="[280,354,423,447]" country="Philippines" municipality="Mindanao" name="Siocon" pageId="4" pageNumber="123" stateProvince="Zamboanga">Siocon</location>
/
<date id="FFAC37E2FFDF5D77FEC4E3B8FE1118A1" box="[376,488,423,447]" pageId="4" pageNumber="123" value="2022-08">
<collectingDate id="EFE8CE0AFFDF5D77FEC4E3B8FE1118A1" box="[376,488,423,447]" pageId="4" pageNumber="123" value="2022-08">VIII. 2022</collectingDate>
</date>
/ local collector leg. (white rectangular label, printed)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30842A9FFDF5D77FF01E01CFE5C1B24" pageId="4" pageNumber="123" type="distribution">
<paragraph id="8BAD1122FFDF5D77FF01E01CFE5C1B24" blockId="4.[189,668,515,570]" pageId="4" pageNumber="123">
<emphasis id="B966CD30FFDF5D77FF01E01CFEA11B05" bold="true" box="[189,344,515,539]" pageId="4" pageNumber="123">Distribution:</emphasis>
<collectingRegion id="49D6DFC0FFDF5D77FEE0E01CFE551B05" box="[348,428,515,539]" country="Philippines" name="Basilan" pageId="4" pageNumber="123">Basilan</collectingRegion>
Island and Southern
<collectingRegion id="49D6DFC0FFDF5D77FF01E03DFEC21B24" box="[189,315,546,570]" country="Philippines" name="Zamboanga" pageId="4" pageNumber="123">Zamboanga</collectingRegion>
(Siocon).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,215 @@
<document id="51257DE4BC6C60DEF71EF5A6818112F8" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722658458551" checkinUser="felipe" docAuthor="Minkina, Ÿukasz, Anichtchenko, Alexander, Vasiljeva, Anastasija &amp; Skelley, Paul E." docDate="2022" docId="03BD44335E50FFEDFC8C27F700CDFC08" docLanguage="en" docName="BalticJColeopterol.22.1.81-101.pdf" docOrigin="Baltic Journal of Coleopterology 22 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.4:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="4" docTitle="Rhyparus bacanensis Minkina, Anichtchenko, Vasiljeva &amp; Skelley, 2022, sp. nov." docType="treatment" docVersion="1" lastPageNumber="84" masterDocId="FF843C4B5E51FFEEFF8A22690417FFB0" masterDocTitle="Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females" masterLastPageNumber="101" masterPageNumber="81" pageNumber="82" updateTime="1722691352965" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="DF817891782C4F21F71F6ED5987A8007">Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females</mods:title>
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<mods:namePart id="F7F47C2499AE526C8BFD1D0833F0FFA2">Minkina, Ÿukasz</mods:namePart>
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<mods:namePart id="3A7EE5E1C97FB509DD47082F3650A91C">Anichtchenko, Alexander</mods:namePart>
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<mods:title id="A3E439AD49C61B44D43759B207BABBD0">Baltic Journal of Coleopterology</mods:title>
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<taxonomicName id="4C148EA65E50FFEFFC8C27F707E3FA06" ID-CoL="9WHZQ" authorityName="Minkina &amp; Anichtchenko &amp; Vasiljeva &amp; Skelley" authorityYear="2022" box="[774,1012,1438,1462]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="82" phylum="Arthropoda" rank="species" species="bacanensis" status="sp. nov.">
<emphasis id="B96029375E50FFEFFC8C27F707E3FA06" bold="true" box="[774,1012,1438,1462]" italics="true" pageId="1" pageNumber="82">Rhyparus bacanensis</emphasis>
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(
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,
<figureCitation id="132FE9A05E50FFEFFC0827D70788FA65" box="[898,927,1470,1493]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 2025. Heads: 20 R. bacanensis sp. nov.,, holotype; 21 R. buruensis sp. nov.,, holotype; 22 R. fijiensis sp. nov.,, holotype; 23 R. stebnickae sp. nov.,, holotype; 24 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 25 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." pageId="1" pageNumber="82">20</figureCitation>
,
<figureCitation id="132FE9A05E50FFEFFC2127D707D0FA65" box="[939,967,1470,1493]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="1" pageNumber="82">26</figureCitation>
)
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locality.
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, Moluccas
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,
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,
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,
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SE of Makian
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vill
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.
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<emphasis id="B96029375E50FFEFFC8C243707ADF9C6" bold="true" box="[774,954,1630,1654]" pageId="1" pageNumber="82">Type material.</emphasis>
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<typeStatus id="54AF4B875E50FFEFFC4F24360039F9C6" box="[965,1070,1631,1654]" pageId="1" pageNumber="82" type="holotype">Holotype</typeStatus>
():
<collectingCountry id="F303B5B55E50FFEFFBFB243600C8F9C6" box="[1137,1247,1631,1654]" name="Indonesia" pageId="1" pageNumber="82">Indonesia</collectingCountry>
, N Moluccas |
<quantity id="4CEC58C05E50FFEFFC0A24E907E3F927" box="[896,1012,1664,1687]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="1" pageNumber="82" unit="m" value="625.0" valueMax="750.0" valueMin="500.0">
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.,
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| SE slopes of
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SE of Makian
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vill. |
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, leg.
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|| (
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).
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<paragraph id="8BABF5255E53FFECFEB522F80157FF14" blockId="2.[319,1344,145,164]" box="[319,1344,145,164]" pageId="2" pageNumber="83">
<emphasis id="B96029375E53FFECFEB522F80157FF14" box="[319,1344,145,164]" italics="true" pageId="2" pageNumber="83">
Four new species of the genus
<taxonomicName id="4C148EA65E53FFECFDDF22F80728FF14" authority="Westwood, 1845" authorityName="Westwood" authorityYear="1845" box="[597,831,145,164]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">Rhyparus Westwood, 1845</taxonomicName>
(
<taxonomicName id="4C148EA65E53FFECFCC722F807A5FF14" authorityName="Linnaeus" authorityYear="1758" box="[845,946,145,164]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="2" pageNumber="83" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C148EA65E53FFECFC3422F8002FFF14" authorityName="Latreille" authorityYear="1802" box="[958,1080,145,164]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
:
<taxonomicName id="4C148EA65E53FFECFBCE22F800BEFF14" authorityName="Leach" authorityYear="1815" box="[1092,1193,145,164]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="subFamily" subFamily="Aphodiinae">Aphodiinae</taxonomicName>
) with modified...
</emphasis>
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Figs. 13.
<taxonomicName id="4C148EA65E53FFECFE2A26020692FB32" authorityName="Minkina &amp; Anichtchenko &amp; Vasiljeva &amp; Skelley" authorityYear="2022" box="[416,645,1131,1154]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="bacanensis" status="sp. nov.">
<emphasis id="B96029375E53FFECFE2A26020692FB32" box="[416,645,1131,1154]" italics="true" pageId="2" pageNumber="83">Rhyparus bacanensis</emphasis>
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<taxonomicNameLabel id="A253944C5E53FFECFD06260306C9FB31" box="[652,734,1130,1153]" pageId="2" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
,
</paragraph>
<paragraph id="8BABF5255E53FFECFEA426E3065FFB11" blockId="2.[302,741,1130,1185]" box="[302,584,1162,1185]" pageId="2" pageNumber="83">view. Scale lines: 1.0 mm.</paragraph>
</caption>
<subSubSection id="C30EA6AE5E53FFEDFEA426B30038FD48" lastPageId="3" lastPageNumber="84" pageId="2" pageNumber="83" type="description">
<paragraph id="8BABF5255E53FFECFEA426B306CBFA41" blockId="2.[302,813,1242,1521]" pageId="2" pageNumber="83">
<emphasis id="B96029375E53FFECFEA426B30679FB42" bold="true" box="[302,622,1242,1266]" pageId="2" pageNumber="83">
Description of the
<typeStatus id="54AF4B875E53FFECFD8E26B3067EFB42" box="[516,617,1242,1266]" pageId="2" pageNumber="83" type="holotype">holotype</typeStatus>
.
</emphasis>
Dorsum (
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). Length:
<quantity id="4CEC58C05E53FFECFE09269305C4FAA1" box="[387,467,1274,1297]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="2" pageNumber="83" unit="mm" value="7.2">7.2 mm</quantity>
; maximum width:
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. Body large­sized for members of this genus, elongate, not so distinctly convex, flattened in central part; weakly shiny; apparently almost glabrous, though partly clothed with very small yellowish macrosetae on head and all longitudinal costae on pronotum and elytra. Brownish to dark brown; antennae, tarsomeres and mouth parts pale brown.
</paragraph>
<paragraph id="8BABF5255E53FFECFEA424730153FA61" blockId="2.[302,813,1562,1745]" lastBlockId="2.[850,1361,1242,1489]" pageId="2" pageNumber="83">
Head (
<figureCitation id="132FE9A05E53FFECFEFC247305D3F981" box="[374,452,1562,1585]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 2025. Heads: 20 R. bacanensis sp. nov.,, holotype; 21 R. buruensis sp. nov.,, holotype; 22 R. fijiensis sp. nov.,, holotype; 23 R. stebnickae sp. nov.,, holotype; 24 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 25 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." pageId="2" pageNumber="83">Fig. 20</figureCitation>
) moderately shiny, tops of costae distinctly shiny; transversely sub­hexagonal; clypeus trapezoidal in outline, anteriorly truncate, on sides weakly upturned as obtuse, weak tooth, and later sinuous on either side; genae distinctly more excavate than eyes; clypeal disc distinctly convex, ringed by a deep groove; convexity with a pair of quite distinct, very short, convergent ridges, nearly on whole surface with distinct, fine punctures bearing small macrosetae. Frons with four distinct, longitudinal ridges with similar structure as ridges on clypeal convexity. Head covered by quite regularly spaced, quite dense, moderately large punctures bearing small macrosetae.
</paragraph>
<paragraph id="8BABF5255E53FFECFCD827930146F961" blockId="2.[850,1361,1530,1745]" pageId="2" pageNumber="83">Pronotum very weakly shiny, tops of costae distinctly shiny; with eight distinct costae and seven intercostae, with two lateral, rounded lobes on each side. Anterior lobes somewhat higher and distinctly narrower than posterior, on the top are the widest part of pronotum. Costae of middle, third and fourth pair not interrupted in basal part</paragraph>
<paragraph id="8BABF5255E52FFEDFD8A22F807FFFF14" blockId="3.[512,1000,145,164]" box="[512,1000,145,164]" pageId="3" pageNumber="84">
<emphasis id="B96029375E52FFEDFD8A22F807FFFF14" box="[512,1000,145,164]" italics="true" pageId="3" pageNumber="84">Minkina Ł., Anichtchenko A., Vasiljeva A., Skelley P.E.</emphasis>
</paragraph>
<paragraph id="8BABF5255E52FFEDFF6922A80675FE48" blockId="3.[227,737,193,504]" pageId="3" pageNumber="84">of apical half, very gently convergent, distinctly convergent in the middle of apical third; second pair of costae distinctly interrupted in basal part of apical half; costae on each side with very small punctures bearing very small macrosetae. All intercostae in anterior part without additional short costae. Median intercostae with dense punctures, quite distinctly concentrated around median part into longitudinal line, all intercostae in basal part with distinct, dense punctation.</paragraph>
<paragraph id="8BABF5255E52FFEDFF6920480628FD88" blockId="3.[227,575,545,568]" box="[227,575,545,568]" pageId="3" pageNumber="84">Scutellum almost imperceptible.</paragraph>
<paragraph id="8BABF5255E52FFEDFF69200805AFFB68" blockId="3.[227,737,609,1240]" pageId="3" pageNumber="84">Elytra moderately shiny, tops of costae and preapical glandular area more distinctly shiny. Each elytron with six elevated costae, and five flat intercostae. Costae on sides with very small punctures bearing very small macrosetae. Preapical glandular area relatively small. Intercostae first to fourth with two distinct rows of punctures; on third intercostae there is very short additional costae with very short additional row of punctures here, however there are simple, sparsely located, much smaller punctures punctures almost until apex. Fifth intercosta with one row of punctures; in basal part of fourth intercosta there is region with extremely short additional costa, with some additional punctures.External caudal bulb distinctly reduced, area between external caudal bulb and sides of elytra not divided; external and medio­internal caudal bulbs not divided; medio­internal caudal bulb shortened, transversally rounded, internally stretched.</paragraph>
<paragraph id="8BABF5255E52FFEDFF692768052BFA28" blockId="3.[227,737,1281,1432]" pageId="3" pageNumber="84">
Pygidium (
<figureCitation id="132FE9A05E52FFEDFEDF276805B7FAA8" box="[341,416,1281,1304]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="3" pageNumber="84">Fig. 26</figureCitation>
) with dense, irregularly spaced punctation, with weak longitudinal rib in the middle and deep excisionon its sides; with distinctly longitudinal apex in the shape of distinct triangle.
</paragraph>
<paragraph id="8BABF5255E52FFEDFF6927A800EEFEE8" blockId="3.[227,737,1473,1752]" lastBlockId="3.[774,1285,193,344]" pageId="3" pageNumber="84">
Venter (
<figureCitation id="132FE9A05E52FFEDFEB127A80568FA68" box="[315,383,1473,1496]" captionStart="Figs" captionStartId="2.[302,352,1130,1153]" captionTargetBox="[302,1361,189,1185]" captionTargetId="figure-284@2.[412,1253,189,1096]" captionTargetPageId="2" captionText="Figs. 13. Rhyparus bacanensis sp. nov., view. Scale lines: 1.0 mm." pageId="3" pageNumber="84">Fig. 2</figureCitation>
) moderately shiny. Meso­metaventral plate flattened in the middle, with distinct, narrow, quite deep longitudinal furrow; punctation of meso­metaventral plate dense, quite regularly spaced, irregular in size; all punctures bearing short macrosetae.Abdominal ventrites moderately shiny, on sides with rows of punctures; with an additional punctured furrow in basal part. Last abdominal ventrite with dense punctures, which are about one and half time larger than on last but one ventrite; in basal half in the middle part with dense, very deep, longitudinal rows; in apical part in the middle with very deep groove, clearly located much below apex of pygidium.
</paragraph>
<paragraph id="8BABF5255E52FFEDFC8C23E80111FE48" blockId="3.[774,1286,385,504]" pageId="3" pageNumber="84">Meso­ and metafemora with two very indistinct tubercles on lower border; all femora shiny, with regular, very distinct, rather small, very dense punctation; all punctures bearing small macrosetae.</paragraph>
<paragraph id="8BABF5255E52FFEDFC8C204907E6FD88" blockId="3.[774,1009,544,568]" box="[774,1009,544,568]" pageId="3" pageNumber="84">
<emphasis id="B96029375E52FFEDFC8C2049076DFD88" bold="true" box="[774,890,544,568]" pageId="3" pageNumber="84">Variation.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BABF5255E52FFEDFC8C20090038FD48" blockId="3.[774,1285,608,760]" pageId="3" pageNumber="84">
<emphasis id="B96029375E52FFEDFC8C200907FAFDC8" bold="true" box="[774,1005,608,632]" pageId="3" pageNumber="84">Sexual dimorphism.</emphasis>
Unknown. When we look at shape of meso­metaventral plate and apex of mesotibiae we can suppose that sexual dimorphism it is like typical for a whole genus (except pygidium and last ventrite).
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E52FFEDFC8C214900CDFC08" pageId="3" pageNumber="84" type="etymology">
<paragraph id="8BABF5255E52FFEDFC8C21490788FCC8" blockId="3.[774,1285,800,888]" pageId="3" pageNumber="84">
<emphasis id="B96029375E52FFEDFC8C21490794FC88" bold="true" box="[774,899,800,824]" pageId="3" pageNumber="84">Etymology.</emphasis>
Toponymic; an adjective derived from the name of Bacan Island where the
<typeStatus id="54AF4B875E52FFEDFB2C21280113FCE8" box="[1190,1284,833,856]" pageId="3" pageNumber="84" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
<paragraph id="8BABF5255E52FFEDFC8C21C900CDFC08" blockId="3.[774,1242,928,952]" box="[774,1242,928,952]" pageId="3" pageNumber="84">
<emphasis id="B96029375E52FFEDFC8C21C90771FC08" bold="true" box="[774,870,928,952]" pageId="3" pageNumber="84">Affinity.</emphasis>
See discussion and tables 1 and 2.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,218 @@
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<mods:title id="7E639EE9DB5BDA37DC094BCA51FD006B">Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females</mods:title>
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<mods:namePart id="B82396F0C8C84C37CF9D35EF88B523F2">Minkina, Ÿukasz</mods:namePart>
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<mods:namePart id="4911B4C9955C92EAD1A5291EE1DA3881">Vasiljeva, Anastasija</mods:namePart>
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<mods:namePart id="D9BA324FE08FBBB9AAE14CEF69F93568">Skelley, Paul E.</mods:namePart>
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<paragraph id="8BABF5255E52FFEDFC8C26680028FBA8" blockId="3.[774,1087,1025,1080]" box="[774,1087,1025,1049]" pageId="3" pageNumber="84">
<heading id="D0E342495E52FFEDFC8C26680028FBA8" box="[774,1087,1025,1049]" fontSize="10" level="2" pageId="3" pageNumber="84" reason="7">
<taxonomicName id="4C148EA65E52FFEDFC8C266807F2FBA9" ID-CoL="9WHZK" authorityName="Minkina &amp; Anichtchenko &amp; Vasiljeva &amp; Skelley" authorityYear="2022" box="[774,997,1025,1049]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="buruensis" status="sp. nov.">
<emphasis id="B96029375E52FFEDFC8C266807F2FBA9" bold="true" box="[774,997,1025,1049]" italics="true" pageId="3" pageNumber="84">Rhyparus buruensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253944C5E52FFEDFC6626680028FBA8" box="[1004,1087,1025,1048]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E52FFEDFC8C264807D8FB88" box="[774,975,1057,1080]" pageId="3" pageNumber="84" type="description">
<paragraph id="8BABF5255E52FFEDFC8C264807D8FB88" blockId="3.[774,1087,1025,1080]" box="[774,975,1057,1080]" pageId="3" pageNumber="84">
(
<figureCitation id="132FE9A05E52FFEDFC8426480761FB88" box="[782,886,1057,1080]" captionStart="Figs" captionStartId="4.[302,352,1693,1716]" captionTargetBox="[483,1187,751,1656]" captionTargetId="figure-275@4.[478,1187,748,1659]" captionTargetPageId="4" captionText="Figs. 46. Rhyparus buruensis sp. nov.,, holotype: 4 dorsal view; 5 ventral view; 6 lateral view. Scale lines: 1.0 mm." pageId="3" pageNumber="84">Figs. 46</figureCitation>
,
<figureCitation id="132FE9A05E52FFEDFC0826480788FB88" box="[898,927,1057,1080]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 2025. Heads: 20 R. bacanensis sp. nov.,, holotype; 21 R. buruensis sp. nov.,, holotype; 22 R. fijiensis sp. nov.,, holotype; 23 R. stebnickae sp. nov.,, holotype; 24 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 25 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." pageId="3" pageNumber="84">21</figureCitation>
,
<figureCitation id="132FE9A05E52FFEDFC21264807D0FB88" box="[939,967,1057,1080]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="3" pageNumber="84">27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E52FFEDFC8C26090043FACA" pageId="3" pageNumber="84" type="materials_examined">
<paragraph id="8BABF5255E52FFEDFC8C2609005DFB28" blockId="3.[774,1285,1120,1176]" pageId="3" pageNumber="84">
<materialsCitation id="3B7CFF785E52FFEDFC8C26090051FB28" country="Indonesia" county="Ramaja Mount" location="Buru Island" municipality="Ilat" pageId="3" pageNumber="84" specimenCount="1" typeStatus="holotype">
<emphasis id="B96029375E52FFEDFC8C2609078CFBC8" bold="true" box="[774,923,1120,1144]" pageId="3" pageNumber="84">
<typeStatus id="54AF4B875E52FFEDFC8C26090729FBC8" box="[774,830,1120,1144]" pageId="3" pageNumber="84">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F303B5B55E52FFEDFC152608001EFBC8" box="[927,1033,1121,1144]" name="Indonesia" pageId="3" pageNumber="84">Indonesia</collectingCountry>
, Moluccas,
<location id="8ECBA3FE5E52FFEDFB0E26080116FBC8" LSID="urn:lsid:plazi:treatment:03BD44335E52FFEBFC8C266800CDFE08:8ECBA3FE5E52FFEDFB0E26080116FBC8" box="[1156,1281,1121,1144]" country="Indonesia" county="Ramaja Mount" municipality="Ilat" name="Buru Island" pageId="3" pageNumber="84">Buru Island</location>
,
<collectingMunicipality id="6BCF6F5F5E52FFEDFC8C26E8073DFB28" box="[774,810,1153,1176]" pageId="3" pageNumber="84">Ilat</collectingMunicipality>
vill., env.,
<collectingCounty id="62CA8DA95E52FFEDFC2826E80051FB28" box="[930,1094,1153,1176]" pageId="3" pageNumber="84">Ramaja Mount</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="8BABF5255E52FFEDFC8C26A80043FACA" blockId="3.[774,1286,1217,1403]" pageId="3" pageNumber="84">
<emphasis id="B96029375E52FFEDFC8C26A807BBFB69" bold="true" box="[774,940,1217,1241]" pageId="3" pageNumber="84">Type material.</emphasis>
<materialsCitation id="3B7CFF785E52FFEDFC3A26A80116FAAA" collectingDate="2012-12" collectionCode="Y, MCN" collectorName="S. Jakl" country="Indonesia" county="Ramaja Mt." elevation="275" location="Buru Island" municipality="Ilat" pageId="3" pageNumber="84" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AF4B875E52FFEDFC3A26A80003FB68" box="[944,1044,1217,1240]" pageId="3" pageNumber="84" type="holotype">Holotype</typeStatus>
():
<collectingCountry id="F303B5B55E52FFEDFBC226A800A7FB68" box="[1096,1200,1217,1240]" name="Indonesia" pageId="3" pageNumber="84">Indonesia</collectingCountry>
| C Moluccas: see
<location id="8ECBA3FE5E52FFEDFC0F268A0012FB4A" LSID="urn:lsid:plazi:treatment:03BD44335E52FFEBFC8C266800CDFE08:8ECBA3FE5E52FFEDFC0F268A0012FB4A" box="[901,1029,1251,1274]" country="Indonesia" county="Ramaja Mt." municipality="Ilat" name="Buru Island" pageId="3" pageNumber="84">Buru Island</location>
|
<quantity id="4CEC58C05E52FFEDFB91268A009AFB4A" box="[1051,1165,1251,1274]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="3" pageNumber="84" unit="m" value="275.0" valueMax="350.0" valueMin="200.0">
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</quantity>
. |
<collectingMunicipality id="6BCF6F5F5E52FFEDFB2D268A00DCFB4A" box="[1191,1227,1251,1274]" pageId="3" pageNumber="84">Ilat</collectingMunicipality>
vill., env.,
<collectingCounty id="62CA8DA95E52FFEDFCB1276A07A3FAAA" box="[827,948,1283,1306]" pageId="3" pageNumber="84">Ramaja Mt.</collectingCounty>
|
<date id="FFAAD3E55E52FFEDFC4B276A0001FAAA" box="[961,1046,1283,1306]" pageId="3" pageNumber="84" value="2012-12">
<collectingDate id="EFEE2A0D5E52FFEDFC4B276A0001FAAA" box="[961,1046,1283,1306]" pageId="3" pageNumber="84" value="2012-12">xii.2012</collectingDate>
</date>
| leg.
<collectorName id="26E190F35E52FFEDFBC7276A0087FAAA" box="[1101,1168,1283,1306]" pageId="3" pageNumber="84">S. Jakl</collectorName>
|| (
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<collectionCode id="ED056DE05E52FFEDFB21276900ACFAAA" box="[1195,1211,1280,1306]" pageId="3" pageNumber="84">Ÿ</collectionCode>
</emphasis>
<collectionCode id="ED056DE05E52FFEDFB31276A00ECFAAA" box="[1211,1275,1283,1306]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="3" pageNumber="84" type="Herbarium">MCN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7CFF785E52FFEDFC8C274A0047FACA" collectingDate="2013-06" collectionCode="Y, MCN" country="Indonesia" county="Ramaja Mt." elevation="200" location="Buru Island" municipality="Ilat" pageId="3" pageNumber="84" specimenCount="1" typeStatus="paratype">
<typeStatus id="54AF4B875E52FFEDFC8C274A0773FA8A" box="[774,868,1315,1338]" pageId="3" pageNumber="84" type="paratype">Paratype</typeStatus>
():
<collectingCountry id="F303B5B55E52FFEDFC16274A0012FA8A" box="[924,1029,1315,1338]" name="Indonesia" pageId="3" pageNumber="84">Indonesia</collectingCountry>
| C Moluccas: see
<location id="8ECBA3FE5E52FFEDFB45274A075FFAEA" LSID="urn:lsid:plazi:treatment:03BD44335E52FFEBFC8C266800CDFE08:8ECBA3FE5E52FFEDFB45274A075FFAEA" country="Indonesia" county="Ramaja Mt." municipality="Ilat" name="Buru Island" pageId="3" pageNumber="84">Buru Island</location>
|
<quantity id="4CEC58C05E52FFEDFCD7272A07D6FAEA" box="[861,961,1347,1370]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.5" metricValueMin="0.5" pageId="3" pageNumber="84" unit="m" value="200.0" valueMax="350.0" valueMin="50.0">
<elevation id="003912165E52FFEDFCD7272A07D6FAEA" box="[861,961,1347,1370]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.5" metricValueMin="0.5" pageId="3" pageNumber="84" unit="m" value="200.0" valueMax="350.0" valueMin="50.0">50­ 350m</elevation>
</quantity>
. |
<collectingMunicipality id="6BCF6F5F5E52FFEDFC53272A07EAFAEA" box="[985,1021,1347,1370]" pageId="3" pageNumber="84">Ilat</collectingMunicipality>
vill., env.,
<collectingCounty id="62CA8DA95E52FFEDFBF2272A00EEFAEA" box="[1144,1273,1347,1370]" pageId="3" pageNumber="84">Ramaja Mt.</collectingCounty>
|
<date id="FFAAD3E55E52FFEDFC8C270A074FFACA" box="[774,856,1379,1402]" pageId="3" pageNumber="84" value="2013-06">
<collectingDate id="EFEE2A0D5E52FFEDFC8C270A074FFACA" box="[774,856,1379,1402]" pageId="3" pageNumber="84" value="2013-06">vi.2013</collectingDate>
</date>
| local coll. || (
<emphasis id="B96029375E52FFEDFC7C27080011FACB" bold="true" box="[1014,1030,1377,1403]" pageId="3" pageNumber="84">
<collectionCode id="ED056DE05E52FFEDFC7C27080011FACB" box="[1014,1030,1377,1403]" pageId="3" pageNumber="84">Ÿ</collectionCode>
</emphasis>
<collectionCode id="ED056DE05E52FFEDFB8C270A005FFACA" box="[1030,1096,1379,1402]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="3" pageNumber="84" type="Herbarium">MCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E52FFEBFC8C27CA0038FF48" lastPageId="5" lastPageNumber="86" pageId="3" pageNumber="84" type="description">
<paragraph id="8BABF5255E52FFEDFC8C27CA0097F90A" blockId="3.[774,1285,1443,1722]" pageId="3" pageNumber="84">
<emphasis id="B96029375E52FFEDFC8C27CA0050FA0B" bold="true" box="[774,1095,1443,1467]" pageId="3" pageNumber="84">
Description of the
<typeStatus id="54AF4B875E52FFEDFC5727CA0055FA0B" box="[989,1090,1443,1467]" pageId="3" pageNumber="84" type="holotype">holotype</typeStatus>
.
</emphasis>
Dorsum (
<figureCitation id="132FE9A05E52FFEDFB3E27CA00EFFA0A" box="[1204,1272,1443,1466]" captionStart="Figs" captionStartId="4.[302,352,1693,1716]" captionTargetBox="[483,1187,751,1656]" captionTargetId="figure-275@4.[478,1187,748,1659]" captionTargetPageId="4" captionText="Figs. 46. Rhyparus buruensis sp. nov.,, holotype: 4 dorsal view; 5 ventral view; 6 lateral view. Scale lines: 1.0 mm." pageId="3" pageNumber="84">Fig. 4</figureCitation>
). Length:
<quantity id="4CEC58C05E52FFEDFCD127AA07BCFA6A" box="[859,939,1475,1498]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="84" unit="mm" value="7.3">7.3 mm</quantity>
; maximum width:
<quantity id="4CEC58C05E52FFEDFBED27AA00D2FA6A" box="[1127,1221,1475,1498]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="3" pageNumber="84" unit="mm" value="2.45">2.45 mm</quantity>
. Body large­sized for members of this genus, elongate, not so distinctly convex, flattened in central part; shiny; apparently almost glabrous, though partly clothed with very small yellowish macrosetae on head and all longitudinal costae on pronotum and elytra. Brownish to dark brown; antennae, tarsomeres and mouth parts pale brown.
</paragraph>
<paragraph id="8BABF5255E55FFEAFEA422A805F8FDC8" blockId="4.[302,813,193,632]" pageId="4" pageNumber="85">
Head (
<figureCitation id="132FE9A05E55FFEAFEFC22A805D1FF68" box="[374,454,193,216]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 2025. Heads: 20 R. bacanensis sp. nov.,, holotype; 21 R. buruensis sp. nov.,, holotype; 22 R. fijiensis sp. nov.,, holotype; 23 R. stebnickae sp. nov.,, holotype; 24 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 25 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." pageId="4" pageNumber="85">Fig. 21</figureCitation>
) weakly shiny; transversely sub­hexagonal; clypeus trapezoidal in outline, anteriorly weakly sinuate, on sides weakly upturned as obtuse, quite distinct tooth, and later sinuous on either side; genae distinctly more excavate than eyes; clypeal disc distinctly convex, ringed by a deep groove; convexity with a pair of quite distinct, very short, convergent ridges, on whole surface with distinct, fine punctures bearing small macrosetae. Frons with four distinct, longitudinal ridges with similar structure as ridges on clypeal convexity. Head covered by quite regularly spaced, quite dense, moderately large punctures bearing small macrosetae.
</paragraph>
<paragraph id="8BABF5255E55FFEAFCD822A80146FD08" blockId="4.[850,1361,193,696]" pageId="4" pageNumber="85">Pronotum quite distinctly shiny; with eight distinct costae and seven intercostae, with two lateral, rounded lobes on each side.Anterior lobes somewhat higherand distinctly narrower than posterior, on the top are the widest part of pronotum. Costae of third and fourth pair not interrupted in basal part of apical half, very gently convergent, distinctly convergent in the middle of apical third; median and second pair of costae distinctly interrupted in basal part of apical half; costae on each side with very small punctures bearing very small macrosetae. All intercostae in anterior part without additional short costae. Median intercostae with dense punctures, weakly concentrated around median part into longitudinal line, all intercostae in basal part with distinct, dense punctation.</paragraph>
<caption id="DF6BA5AD5E55FFEAFEA424F4065FF964" pageId="4" pageNumber="85" startId="4.[302,352,1693,1716]" targetBox="[483,1187,751,1656]" targetPageId="4" targetType="figure">
<paragraph id="8BABF5255E55FFEAFEA424F4065FF964" blockId="4.[302,1361,1693,1748]" pageId="4" pageNumber="85">
Figs. 46.
<taxonomicName id="4C148EA65E55FFEAFE2B24F7066EF905" authorityName="Minkina &amp; Anichtchenko &amp; Vasiljeva &amp; Skelley" authorityYear="2022" box="[417,633,1694,1717]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="buruensis" status="sp. nov.">
<emphasis id="B96029375E55FFEAFE2B24F7066EF905" box="[417,633,1694,1717]" italics="true" pageId="4" pageNumber="85">Rhyparus buruensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253944C5E55FFEAFD0A24F406C4F904" box="[640,723,1693,1716]" pageId="4" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
,, holotype: 4 dorsal view; 5 ventral view; 6 lateral view. Scale lines: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BABF5255E54FFEBFD8A22F807FFFF14" blockId="5.[512,1000,145,164]" box="[512,1000,145,164]" pageId="5" pageNumber="86">
<emphasis id="B96029375E54FFEBFD8A22F807FFFF14" box="[512,1000,145,164]" italics="true" pageId="5" pageNumber="86">Minkina Ł., Anichtchenko A., Vasiljeva A., Skelley P.E.</emphasis>
</paragraph>
<paragraph id="8BABF5255E54FFEBFF6922A80628FF68" blockId="5.[227,575,193,216]" box="[227,575,193,216]" pageId="5" pageNumber="86">Scutellum almost imperceptible.</paragraph>
<paragraph id="8BABF5255E54FFEBFF69236806B9FD48" blockId="5.[227,737,257,760]" pageId="5" pageNumber="86">Elytra shiny. Each elytron with six elevated costae, and five flat intercostae. Costae on sides with very small punctures bearing very small macrosetae. Preapical glandular area relatively small. Intercostae first to fourth with two distinct rows of punctures; on third intercostae there is short additional costae with additional row of punctures untill half of length of elytra. Fifth intercosta with one row of punctures; in basal part of fourth intercosta there is region with extremely short additional costa, with some additional punctures here. External caudal bulb distinctly reduced, area between external caudal bulb and sides of elytra not divided; external and medio­internal caudal bulbs not divided; medio­internal caudal bulb shortened, transversally rounded, internally stretched.</paragraph>
<paragraph id="8BABF5255E54FFEBFF6921480582FC08" blockId="5.[227,737,801,952]" pageId="5" pageNumber="86">
Pygidium (
<figureCitation id="132FE9A05E54FFEBFEDF21480588FC88" box="[341,415,801,824]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="5" pageNumber="86">Fig. 27</figureCitation>
) with dense, irregularly spaced punctation, with weak longitudinal rib in the middle, and deep excision on its sides; with distinctly longitudinal apex in the shape of volcano with sinuate top.
</paragraph>
<paragraph id="8BABF5255E54FFEBFF69218806C2FAC8" blockId="5.[227,738,993,1400]" pageId="5" pageNumber="86">
Venter (
<figureCitation id="132FE9A05E54FFEBFEB121880597FC48" box="[315,384,993,1016]" captionStart="Figs" captionStartId="4.[302,352,1693,1716]" captionTargetBox="[483,1187,751,1656]" captionTargetId="figure-275@4.[478,1187,748,1659]" captionTargetPageId="4" captionText="Figs. 46. Rhyparus buruensis sp. nov.,, holotype: 4 dorsal view; 5 ventral view; 6 lateral view. Scale lines: 1.0 mm." pageId="5" pageNumber="86">Fig. 5</figureCitation>
) shiny. Meso­metaventral plate flattened in the middle, with distinct, narrow, quite deep longitudinal furrow; punctation of meso­metaventral plate dense, regularly spaced, irregular in size; all punctures bearing short macrosetae. Abdominal ventrites moderately shiny, on sides with rows of punctures; with an additional punctured furrow in basal part. Last abdominal ventrite with dense punctures, which are similar size as on last but one ventrite; in basal half in the middle part with dense, very deep, longitudinal rows; in apical part in the middle with very deep groove, clearly located much below apex of pygidium.
</paragraph>
<paragraph id="8BABF5255E54FFEBFF6927C8069CF988" blockId="5.[227,737,1441,1624]" pageId="5" pageNumber="86">Mesofemora with two very indistinct tubercles on lower border; metafemora without any tubercles on lower border; all femora shiny, with regular, very distinct, rather small, very dense punctation; all punctures bearing small macrosetae.</paragraph>
<paragraph id="8BABF5255E54FFEBFF69242905DBF9E8" blockId="5.[227,737,1441,1624]" box="[227,460,1600,1624]" pageId="5" pageNumber="86">
<emphasis id="B96029375E54FFEBFF6924290541F9E8" bold="true" box="[227,342,1600,1624]" pageId="5" pageNumber="86">Variation.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BABF5255E54FFEBFF6924E90038FF48" blockId="5.[227,737,1664,1752]" lastBlockId="5.[774,1285,193,248]" pageId="5" pageNumber="86">
<emphasis id="B96029375E54FFEBFF6924E905DEF928" bold="true" box="[227,457,1664,1688]" pageId="5" pageNumber="86">Sexual dimorphism.</emphasis>
Unknown. When we look at shape of meso­metaventral plate and apex of mesotibiae we can suppose that sexual dimorphism it is like typical for a whole genus (except pygidium and last ventrite).
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E54FFEBFC8C234900CDFE08" pageId="5" pageNumber="86" type="etymology">
<paragraph id="8BABF5255E54FFEBFC8C23490788FEC8" blockId="5.[774,1285,288,376]" pageId="5" pageNumber="86">
<emphasis id="B96029375E54FFEBFC8C23490794FE88" bold="true" box="[774,899,288,312]" pageId="5" pageNumber="86">Etymology.</emphasis>
Toponymic; an adjective derived from the name of Buru Island where the
<typeStatus id="54AF4B875E54FFEBFB2C23280113FEE8" box="[1190,1284,321,344]" pageId="5" pageNumber="86" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
<paragraph id="8BABF5255E54FFEBFC8C23C900CDFE08" blockId="5.[774,1242,416,440]" box="[774,1242,416,440]" pageId="5" pageNumber="86">
<emphasis id="B96029375E54FFEBFC8C23C90771FE08" bold="true" box="[774,870,416,440]" pageId="5" pageNumber="86">Affinity.</emphasis>
See discussion and tables 1 and 2.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,347 @@
<document id="C15F74CD0DDC99AF511175A6B54FFE09" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722658458551" checkinUser="felipe" docAuthor="Minkina, Ÿukasz, Anichtchenko, Alexander, Vasiljeva, Anastasija &amp; Skelley, Paul E." docDate="2022" docId="03BD44335E54FFE9FC8C206800CDFD48" docLanguage="en" docName="BalticJColeopterol.22.1.81-101.pdf" docOrigin="Baltic Journal of Coleopterology 22 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.4:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="4" docTitle="Rhyparus fijiensis Minkina, Anichtchenko, Vasiljeva &amp; Skelley, 2022, sp. nov." docType="treatment" docVersion="1" lastPageNumber="88" masterDocId="FF843C4B5E51FFEEFF8A22690417FFB0" masterDocTitle="Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females" masterLastPageNumber="101" masterPageNumber="81" pageNumber="86" updateTime="1722691352965" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="99FD355019D85701CDE6C855C36565E3">Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females</mods:title>
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<mods:namePart id="BCE81067C5E2E4210AEAE381656963D7">Skelley, Paul E.</mods:namePart>
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<subSubSection id="C30EA6AE5E54FFEBFC8C2068003EFDA8" box="[774,1065,513,537]" pageId="5" pageNumber="86" type="nomenclature">
<paragraph id="8BABF5255E54FFEBFC8C2068003EFDA8" blockId="5.[774,1065,513,568]" box="[774,1065,513,537]" pageId="5" pageNumber="86">
<heading id="D0E342495E54FFEBFC8C2068003EFDA8" box="[774,1065,513,537]" fontSize="10" level="2" pageId="5" pageNumber="86" reason="7">
<taxonomicName id="4C148EA65E54FFEBFC8C206807C7FDA9" ID-CoL="9WHZW" authorityName="Minkina &amp; Anichtchenko &amp; Vasiljeva &amp; Skelley" authorityYear="2022" box="[774,976,513,537]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="species" species="fijiensis" status="sp. nov.">
<emphasis id="B96029375E54FFEBFC8C206807C7FDA9" bold="true" box="[774,976,513,537]" italics="true" pageId="5" pageNumber="86">Rhyparus fijiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253944C5E54FFEBFC5C2068003EFDA8" box="[982,1065,513,536]" pageId="5" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E54FFEBFC8C204807D8FD88" box="[774,975,545,568]" pageId="5" pageNumber="86" type="description">
<paragraph id="8BABF5255E54FFEBFC8C204807D8FD88" blockId="5.[774,1065,513,568]" box="[774,975,545,568]" pageId="5" pageNumber="86">
(
<figureCitation id="132FE9A05E54FFEBFC8420480760FD88" box="[782,887,545,568]" captionStart="Figs" captionStartId="6.[302,352,1693,1716]" captionTargetBox="[491,1182,755,1657]" captionTargetId="figure-272@6.[487,1184,749,1658]" captionTargetPageId="6" captionText="Figs. 79. Rhyparus fijiensis sp. nov.,, holotype: 7 dorsal view; 8 ventral view; 9 lateral view. Scale lines: 1.0 mm." pageId="5" pageNumber="86">Figs. 79</figureCitation>
,
<figureCitation id="132FE9A05E54FFEBFC0820480788FD88" box="[898,927,545,568]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 2025. Heads: 20 R. bacanensis sp. nov.,, holotype; 21 R. buruensis sp. nov.,, holotype; 22 R. fijiensis sp. nov.,, holotype; 23 R. stebnickae sp. nov.,, holotype; 24 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 25 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." pageId="5" pageNumber="86">22</figureCitation>
,
<figureCitation id="132FE9A05E54FFEBFC21204807D0FD88" box="[939,967,545,568]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="5" pageNumber="86">28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E54FFEBFC8C2009008BFB2A" pageId="5" pageNumber="86" type="materials_examined">
<paragraph id="8BABF5255E54FFEBFC8C200900CBFDC8" blockId="5.[774,1244,608,632]" box="[774,1244,608,632]" pageId="5" pageNumber="86">
<materialsCitation id="3B7CFF785E54FFEBFC8C200900CFFDC8" box="[774,1240,608,632]" country="Fiji" location="Viti Levu Island" municipality="Navai" pageId="5" pageNumber="86" specimenCount="1" typeStatus="holotype">
<emphasis id="B96029375E54FFEBFC8C20090788FDC8" bold="true" box="[774,927,608,632]" pageId="5" pageNumber="86">
<typeStatus id="54AF4B875E54FFEBFC8C20090729FDC8" box="[774,830,608,632]" pageId="5" pageNumber="86">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F303B5B55E54FFEBFC2F200807DAFDC8" box="[933,973,609,632]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
,
<location id="8ECBA3FE5E54FFEBFC522008009DFDC8" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFC522008009DFDC8" box="[984,1162,609,632]" country="Fiji" municipality="Navai" name="Viti Levu Island" pageId="5" pageNumber="86">Viti Levu Island</location>
,
<collectingMunicipality id="6BCF6F5F5E54FFEBFB1E200800CFFDC8" box="[1172,1240,609,632]" pageId="5" pageNumber="86">Navai</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8BABF5255E54FFEBFC8C20C8008BFB2A" blockId="5.[774,1285,673,1178]" pageId="5" pageNumber="86">
<emphasis id="B96029375E54FFEBFC8C20C807A7FD09" bold="true" box="[774,944,673,697]" pageId="5" pageNumber="86">Type material.</emphasis>
<materialsCitation id="3B7CFF785E54FFEBFC3320C8003CFD6A" collectingDate="2016-07" collectionCode="Y, MCN" country="Fiji" location="Levu" municipality="Navai" pageId="5" pageNumber="86" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AF4B875E54FFEBFC3320C8000AFD08" box="[953,1053,673,696]" pageId="5" pageNumber="86" type="holotype">Holotype</typeStatus>
():
<collectingCountry id="F303B5B55E54FFEBFBD320C80068FD08" box="[1113,1151,673,696]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
| Viti
<location id="8ECBA3FE5E54FFEBFB4D20C80116FD08" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFB4D20C80116FD08" box="[1223,1281,673,696]" country="Fiji" municipality="Navai" name="Levu" pageId="5" pageNumber="86">Levu</location>
,
<collectingMunicipality id="6BCF6F5F5E54FFEBFC8C20AA0751FD6A" box="[774,838,707,730]" pageId="5" pageNumber="86">Navai</collectingMunicipality>
|
<date id="FFAAD3E55E54FFEBFCD220AA07A6FD6A" box="[856,945,707,730]" pageId="5" pageNumber="86" value="2016-07">
<collectingDate id="EFEE2A0D5E54FFEBFCD220AA07A6FD6A" box="[856,945,707,730]" pageId="5" pageNumber="86" value="2016-07">vii.2016</collectingDate>
</date>
|| (
<emphasis id="B96029375E54FFEBFC5B20A907F6FD6A" bold="true" box="[977,993,704,730]" pageId="5" pageNumber="86">
<collectionCode id="ED056DE05E54FFEBFC5B20A907F6FD6A" box="[977,993,704,730]" pageId="5" pageNumber="86">Ÿ</collectionCode>
</emphasis>
<collectionCode id="ED056DE05E54FFEBFC6B20AA0034FD6A" box="[993,1059,707,730]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="5" pageNumber="86" type="Herbarium">MCN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7CFF785E54FFEBFBBE20AA0117FD4A" collectingDate="2016-12" collectionCode="Y, MCN" country="Fiji" location="Levu" municipality="Navai" pageId="5" pageNumber="86" specimenCount="1" typeStatus="paratype">
<typeStatus id="54AF4B875E54FFEBFBBE20AA008AFD6A" box="[1076,1181,707,730]" pageId="5" pageNumber="86" type="paratype">Paratypes</typeStatus>
(27exx.): 2exx.:
<collectingCountry id="F303B5B55E54FFEBFCC6208A0767FD4A" box="[844,880,739,762]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
| Viti
<location id="8ECBA3FE5E54FFEBFC23208A07F6FD4A" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFC23208A07F6FD4A" box="[937,993,739,762]" country="Fiji" municipality="Navai" name="Levu" pageId="5" pageNumber="86">Levu</location>
,
<collectingMunicipality id="6BCF6F5F5E54FFEBFC60208A003EFD4A" box="[1002,1065,739,762]" pageId="5" pageNumber="86">Navai</collectingMunicipality>
|
<date id="FFAAD3E55E54FFEBFBBD208A009AFD4A" box="[1079,1165,739,762]" pageId="5" pageNumber="86" value="2016-12">
<collectingDate id="EFEE2A0D5E54FFEBFBBD208A009AFD4A" box="[1079,1165,739,762]" pageId="5" pageNumber="86" value="2016-12">xii.2016</collectingDate>
</date>
|| (
<emphasis id="B96029375E54FFEBFB23208900AEFD4A" bold="true" box="[1193,1209,736,762]" pageId="5" pageNumber="86">
<collectionCode id="ED056DE05E54FFEBFB23208900AEFD4A" box="[1193,1209,736,762]" pageId="5" pageNumber="86">Ÿ</collectionCode>
</emphasis>
<collectionCode id="ED056DE05E54FFEBFB33208A00EEFD4A" box="[1209,1273,739,762]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="5" pageNumber="86" type="Herbarium">MCN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7CFF785E54FFEBFC8C216A004AFC8A" collectingDate="2016-12" collectionCode="CNCW, DUBC, ISEA" country="Fiji" location="Levu" municipality="Navai" pageId="5" pageNumber="86" specimenCount="2" typeStatus="paratype">
6exx.:
<collectingCountry id="F303B5B55E54FFEBFCDA216A0761FCAA" box="[848,886,771,794]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
| Viti
<location id="8ECBA3FE5E54FFEBFC3D216A07E6FCAA" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFC3D216A07E6FCAA" box="[951,1009,771,794]" country="Fiji" municipality="Navai" name="Levu" pageId="5" pageNumber="86">Levu</location>
,
<collectingMunicipality id="6BCF6F5F5E54FFEBFC76216A002BFCAA" box="[1020,1084,771,794]" pageId="5" pageNumber="86">Navai</collectingMunicipality>
|
<date id="FFAAD3E55E54FFEBFBC5216A00BEFCAA" box="[1103,1193,771,794]" pageId="5" pageNumber="86" value="2016-12">
<collectingDate id="EFEE2A0D5E54FFEBFBC5216A00BEFCAA" box="[1103,1193,771,794]" pageId="5" pageNumber="86" value="2016-12">xii.2016</collectingDate>
</date>
|| (4exx.
<collectionCode id="ED056DE05E54FFEBFC8C214A0744FC8A" box="[774,851,803,826]" pageId="5" pageNumber="86">CNCW</collectionCode>
,
<specimenCount id="9D123EAC5E54FFEBFCD5214A079CFC8A" box="[863,907,803,826]" count="1" pageId="5" pageNumber="86" type="generic">1ex.</specimenCount>
<collectionCode id="ED056DE05E54FFEBFC1A214A07CBFC8A" box="[912,988,803,826]" pageId="5" pageNumber="86">DUBC</collectionCode>
,
<specimenCount id="9D123EAC5E54FFEBFC6F214A0005FC8A" box="[997,1042,803,826]" count="1" pageId="5" pageNumber="86" type="generic">1ex.</specimenCount>
<collectionCode id="ED056DE05E54FFEBFB9C214A0042FC8A" box="[1046,1109,803,826]" pageId="5" pageNumber="86">ISEA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7CFF785E54FFEBFBEF214A07F7FCCA" collectingDate="1975-07-31" collectionCode="NZAC" collectorName="P. A. Maddison" country="Fiji" location="Levu" municipality="Savura Creek" pageId="5" pageNumber="86" specimenCount="1" typeStatus="paratype">
2exx.:
<collectingCountry id="F303B5B55E54FFEBFB21214A00C7FC8A" box="[1195,1232,803,826]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
| Viti
<location id="8ECBA3FE5E54FFEBFC8C212A072BFCEA" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFC8C212A072BFCEA" box="[774,828,835,858]" country="Fiji" municipality="Savura Creek" name="Levu" pageId="5" pageNumber="86">Levu</location>
|
<collectingMunicipality id="6BCF6F5F5E54FFEBFCC0212A07CFFCEA" box="[842,984,835,858]" pageId="5" pageNumber="86">Savura Creek</collectingMunicipality>
,
<date id="FFAAD3E55E54FFEBFC6A212A0041FCEA" box="[992,1110,835,858]" pageId="5" pageNumber="86" value="1975-07-31">
<collectingDate id="EFEE2A0D5E54FFEBFC6A212A0041FCEA" box="[992,1110,835,858]" pageId="5" pageNumber="86" value="1975-07-31">31.vii.1975</collectingDate>
</date>
|
<collectorName id="26E190F35E54FFEBFBEE212A0116FCEA" box="[1124,1281,835,858]" pageId="5" pageNumber="86">P.A. Maddison</collectorName>
, M.V. light || (
<collectionCode id="ED056DE05E54FFEBFC01210A07CEFCCA" box="[907,985,867,890]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="86" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7CFF785E54FFEBFC60210A0116FC2A" collectingDate="1985-08-04" collectionCode="NZAC" collectorName="T. Kepa" country="Fiji" location="Levu" municipality="Colo-i-Savu" pageId="5" pageNumber="86" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D123EAC5E54FFEBFC60210A000EFCCA" box="[1002,1049,867,890]" count="1" pageId="5" pageNumber="86" type="generic">1ex.</specimenCount>
:
<collectingCountry id="F303B5B55E54FFEBFBA8210A0050FCCA" box="[1058,1095,867,890]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
| Viti
<location id="8ECBA3FE5E54FFEBFB08210A00AFFCCA" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFB08210A00AFFCCA" box="[1154,1208,867,890]" country="Fiji" municipality="Colo-i-Savu" name="Levu" pageId="5" pageNumber="86">Levu</location>
|
<collectingMunicipality id="6BCF6F5F5E54FFEBFB43210A075EFC2A" pageId="5" pageNumber="86">Colo­i­Savu</collectingMunicipality>
|
<date id="FFAAD3E55E54FFEBFCDD21EA07D0FC2A" box="[855,967,899,922]" pageId="5" pageNumber="86" value="1985-08-04">
<collectingDate id="EFEE2A0D5E54FFEBFCDD21EA07D0FC2A" box="[855,967,899,922]" pageId="5" pageNumber="86" value="1985-08-04">4.viii.1985</collectingDate>
</date>
|
<collectorName id="26E190F35E54FFEBFC5921EA0032FC2A" box="[979,1061,899,922]" pageId="5" pageNumber="86">T. Kepa</collectorName>
, M.V. light || (
<collectionCode id="ED056DE05E54FFEBFB2721EA00EEFC2A" box="[1197,1273,899,922]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="86" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7CFF785E54FFEBFC8C21CA0086FC6A" collectingDate="1975-01-18" collectionCode="NZAC" collectorName="P. A. Maddison" country="Fiji" location="Levu" municipality="Colo-i-Savu" pageId="5" pageNumber="86" specimenCount="1" typeStatus="paratype">
2exx.:
<collectingCountry id="F303B5B55E54FFEBFCC521CA0762FC0A" box="[847,885,931,954]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
| Viti
<location id="8ECBA3FE5E54FFEBFC3F21CA07FBFC0A" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFC3F21CA07FBFC0A" box="[949,1004,931,954]" country="Fiji" municipality="Colo-i-Savu" name="Levu" pageId="5" pageNumber="86">Levu</location>
|
<collectingMunicipality id="6BCF6F5F5E54FFEBFC7421CA0095FC0A" box="[1022,1154,931,954]" pageId="5" pageNumber="86">Colo­i­Savu</collectingMunicipality>
|
<date id="FFAAD3E55E54FFEBFB1921CA00EDFC0A" box="[1171,1274,931,954]" pageId="5" pageNumber="86" value="1975-01-18">
<collectingDate id="EFEE2A0D5E54FFEBFB1921CA00EDFC0A" box="[1171,1274,931,954]" pageId="5" pageNumber="86" value="1975-01-18">18.i.1975</collectingDate>
</date>
|
<collectorName id="26E190F35E54FFEBFC8C21AA07B0FC6A" box="[774,935,963,986]" pageId="5" pageNumber="86">P.A. Maddison</collectorName>
, M.V. light || (
<collectionCode id="ED056DE05E54FFEBFBB021AA009EFC6A" box="[1082,1161,963,986]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="86" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7CFF785E54FFEBFB1021AA0767FBAA" collectingDate="1977-10-19" collectionCode="NZAC" collectorName="G. Kuschel" country="Fiji" elevation="750" location="Levu" municipality="Nandrau" pageId="5" pageNumber="86" specimenCount="1" typeStatus="paratype">
4exx.:
<collectingCountry id="F303B5B55E54FFEBFB6A21AA0112FC6A" box="[1248,1285,963,986]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
| Viti
<location id="8ECBA3FE5E54FFEBFCB3218A0779FC4A" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFCB3218A0779FC4A" box="[825,878,995,1018]" country="Fiji" municipality="Nandrau" name="Levu" pageId="5" pageNumber="86">Levu</location>
|
<collectingMunicipality id="6BCF6F5F5E54FFEBFCF1218A07C4FC4A" box="[891,979,995,1018]" pageId="5" pageNumber="86">Nandrau</collectingMunicipality>
|
<quantity id="4CEC58C05E54FFEBFC6B218A0009FC4A" box="[993,1054,995,1018]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="86" unit="m" value="750.0">
<elevation id="003912165E54FFEBFC6B218A0009FC4A" box="[993,1054,995,1018]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="86" unit="m" value="750.0">750m</elevation>
</quantity>
,
<date id="FFAAD3E55E54FFEBFBAC218A009BFC4A" box="[1062,1164,995,1018]" pageId="5" pageNumber="86" value="1977-10-19">
<collectingDate id="EFEE2A0D5E54FFEBFBAC218A009BFC4A" box="[1062,1164,995,1018]" pageId="5" pageNumber="86" value="1977-10-19">19.x.1977</collectingDate>
</date>
|
<collectorName id="26E190F35E54FFEBFB11218A0112FC4A" box="[1179,1285,995,1018]" pageId="5" pageNumber="86">G.Kuschel</collectorName>
|| (
<collectionCode id="ED056DE05E54FFEBFC96266A077EFBAA" box="[796,873,1027,1050]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="86" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7CFF785E54FFEBFCF2266A002DFBEA" collectingDate="1977-10-10" collectionCode="FSCA, NZAC" collectorName="G. Kuschel" country="Fiji" location="Levu" municipality="Savu" pageId="5" pageNumber="86" specimenCount="1" typeStatus="paratype">
9exx.:
<collectingCountry id="F303B5B55E54FFEBFC36266A07F7FBAA" box="[956,992,1027,1050]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
| Viti
<location id="8ECBA3FE5E54FFEBFB92266A0059FBAA" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFB92266A0059FBAA" box="[1048,1102,1027,1050]" country="Fiji" municipality="Savu" name="Levu" pageId="5" pageNumber="86">Levu</location>
|
<collectingMunicipality id="6BCF6F5F5E54FFEBFBD6266A0087FBAA" box="[1116,1168,1027,1050]" pageId="5" pageNumber="86">Savu</collectingMunicipality>
|
<date id="FFAAD3E55E54FFEBFB17266A0112FBAA" box="[1181,1285,1027,1050]" pageId="5" pageNumber="86" value="1977-10-10">
<collectingDate id="EFEE2A0D5E54FFEBFB17266A0112FBAA" box="[1181,1285,1027,1050]" pageId="5" pageNumber="86" value="1977-10-10">10.x.1977</collectingDate>
</date>
|
<collectorName id="26E190F35E54FFEBFC99264A079DFB8A" box="[787,906,1059,1082]" pageId="5" pageNumber="86">G. Kuschel</collectorName>
[duplicate specimens in alcohol] || (6exx.
<collectionCode id="ED056DE05E54FFEBFCDB262A078FFBEA" box="[849,920,1091,1114]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="86" type="Museum">FSCA</collectionCode>
; 3exx.
<collectionCode id="ED056DE05E54FFEBFC6F262A0024FBEA" box="[997,1075,1091,1114]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="86" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7CFF785E54FFEBFBCC262A008EFB2A" collectingDate="1975-10-03" collectingDateMax="1975-10-06" collectingDateMin="1975-10-03" collectionCode="NZAC" collectorName="P. S. Maddison" country="Fiji" location="Vanjua Levu" pageId="5" pageNumber="86" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D123EAC5E54FFEBFBCC262A0061FBEA" box="[1094,1142,1091,1114]" count="1" pageId="5" pageNumber="86" type="generic">1ex.</specimenCount>
:
<collectingCountry id="F303B5B55E54FFEBFB08262A00B0FBEA" box="[1154,1191,1091,1114]" name="Fiji" pageId="5" pageNumber="86">Fiji</collectingCountry>
|
<location id="8ECBA3FE5E54FFEBFB30262A072AFBCA" LSID="urn:lsid:plazi:treatment:03BD44335E54FFE9FC8C206800CDFD48:8ECBA3FE5E54FFEBFB30262A072AFBCA" country="Fiji" name="Vanjua Levu" pageId="5" pageNumber="86">Vanjua Levu</location>
| “forest nr. Niurudi Navonu” |
<date id="FFAAD3E55E54FFEBFB03260A0117FBCA" box="[1161,1280,1123,1146]" pageId="5" pageNumber="86" value="1975-10-03" valueMax="1975-10-06" valueMin="1975-10-03">
<collectingDate id="EFEE2A0D5E54FFEBFB03260A0117FBCA" box="[1161,1280,1123,1146]" pageId="5" pageNumber="86" value="1975-10-03" valueMax="1975-10-06" valueMin="1975-10-03">3­ 6.x.1975</collectingDate>
</date>
,
<collectorName id="26E190F35E54FFEBFC8C26EA07BCFB2A" box="[774,939,1155,1178]" pageId="5" pageNumber="86">P. S. Maddison</collectorName>
, M.V. light || (
<collectionCode id="ED056DE05E54FFEBFBCB26EA0087FB2A" box="[1089,1168,1155,1178]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="5" pageNumber="86" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E54FFE9FC8C26AA0048FD88" lastPageId="7" lastPageNumber="88" pageId="5" pageNumber="86" type="description">
<paragraph id="8BABF5255E54FFEBFC8C26AA0097FA6A" blockId="5.[774,1285,1219,1498]" pageId="5" pageNumber="86">
<emphasis id="B96029375E54FFEBFC8C26AA0050FB6B" bold="true" box="[774,1095,1219,1243]" pageId="5" pageNumber="86">
Description of the
<typeStatus id="54AF4B875E54FFEBFC5726AA0055FB6B" box="[989,1090,1219,1243]" pageId="5" pageNumber="86" type="holotype">holotype</typeStatus>
.
</emphasis>
Dorsum (
<figureCitation id="132FE9A05E54FFEBFB3E26AA00EFFB6A" box="[1204,1272,1219,1242]" captionStart="Figs" captionStartId="6.[302,352,1693,1716]" captionTargetBox="[491,1182,755,1657]" captionTargetId="figure-272@6.[487,1184,749,1658]" captionTargetPageId="6" captionText="Figs. 79. Rhyparus fijiensis sp. nov.,, holotype: 7 dorsal view; 8 ventral view; 9 lateral view. Scale lines: 1.0 mm." pageId="5" pageNumber="86">Fig. 7</figureCitation>
). Length: 8.0 mm; maximum width:
<quantity id="4CEC58C05E54FFEBFBFB268A00D3FB4A" box="[1137,1220,1251,1274]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="86" unit="mm" value="2.3">2.3 mm</quantity>
. Body large­sized for members of this genus, elongate, not so distinctly convex, flattened in central part; shiny; apparently almost glabrous, though partly clothed with very small yellowish macrosetae on head and all longitudinal costae on pronotum and elytra. Brownish to dark brown; antennae, tarsomeres and mouth parts pale brown.
</paragraph>
<paragraph id="8BABF5255E54FFE8FC8C246A0651FE28" blockId="5.[774,1285,1539,1754]" lastBlockId="6.[302,813,193,408]" lastPageId="6" lastPageNumber="87" pageId="5" pageNumber="86">
Head (
<figureCitation id="132FE9A05E54FFEBFCC5246A0789F9AA" box="[847,926,1539,1562]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 2025. Heads: 20 R. bacanensis sp. nov.,, holotype; 21 R. buruensis sp. nov.,, holotype; 22 R. fijiensis sp. nov.,, holotype; 23 R. stebnickae sp. nov.,, holotype; 24 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 25 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." pageId="5" pageNumber="86">Fig. 22</figureCitation>
) weakly shiny; transversely sub­hexagonal; clypeus trapezoidal in outline, anteriorly weakly sinuate, on sides weakly upturned as obtuse, quite distinct tooth, and later sinuous on either side; genae distinctly more excavate than eyes; clypeal disc distinctly convex, ringed by a deep groove; convexity with a pair of quite distinct, very short, convergent ridges, nearly on whole surface with distinct, fine punctures bearing small macrosetae. Frons with four distinct, longitudinal ridges with similar structure as ridges on clypeal convexity. Head covered by quite regularly spaced, quite dense, moderately large punctures bearing small macrosetae.
</paragraph>
<paragraph id="8BABF5255E57FFE8FEA423A800C5FE08" blockId="6.[302,813,449,696]" lastBlockId="6.[850,1362,193,440]" pageId="6" pageNumber="87">Pronotum shiny; with eight distinct costae and seven intercostae, with two lateral, rounded lobes on each side. Anterior lobes very weakly higher and distinctly narrower than posterior, on the top are the widest part of pronotum. Costae of third and fourth pair not interrupted in basal part of apical half, very gently convergent, distinctly convergent in the middle of apical third; median and second pair of costae distinctly interrupted in basal part of apical half; costae on each side with very small punctures bearing very small macrosetae. All intercostae in anterior part without additional short costae. Median intercostae with dense punctures, distinctly concentrated around median part into longitudinal line, all intercostae in basal part with distinct, dense punctation.</paragraph>
<paragraph id="8BABF5255E57FFE8FCD8238800B9FE48" blockId="6.[850,1198,481,504]" box="[850,1198,481,504]" pageId="6" pageNumber="87">Scutellum almost imperceptible.</paragraph>
<paragraph id="8BABF5255E57FFE8FCD820480147FD08" blockId="6.[850,1361,545,696]" pageId="6" pageNumber="87">Elytra shiny. Each elytron with six elevated costae, and five flat intercostae. Costae on sides with very small punctures bearing very small macrosetae. Preapical glandular area relatively small. Intercostae first to fourth with two distinct rows of</paragraph>
<caption id="DF6BA5AD5E57FFE8FEA424F4061EF964" pageId="6" pageNumber="87" startId="6.[302,352,1693,1716]" targetBox="[491,1182,755,1657]" targetPageId="6" targetType="figure">
<paragraph id="8BABF5255E57FFE8FEA424F4061EF964" blockId="6.[302,1361,1693,1748]" pageId="6" pageNumber="87">
Figs. 79.
<taxonomicName id="4C148EA65E57FFE8FE1724F7064AF905" authorityName="Minkina &amp; Anichtchenko &amp; Vasiljeva &amp; Skelley" authorityYear="2022" box="[413,605,1694,1717]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="fijiensis" status="sp. nov.">
<emphasis id="B96029375E57FFE8FE1724F7064AF905" box="[413,605,1694,1717]" italics="true" pageId="6" pageNumber="87">Rhyparus fijiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253944C5E57FFE8FDE824F406A4F904" box="[610,691,1693,1716]" pageId="6" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
,, holotype: 7 dorsal view; 8 ventral view; 9 lateral view. Scale lines: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BABF5255E56FFE9FD8A22F807FFFF14" blockId="7.[512,1000,145,164]" box="[512,1000,145,164]" pageId="7" pageNumber="88">
<emphasis id="B96029375E56FFE9FD8A22F807FFFF14" box="[512,1000,145,164]" italics="true" pageId="7" pageNumber="88">Minkina Ł., Anichtchenko A., Vasiljeva A., Skelley P.E.</emphasis>
</paragraph>
<paragraph id="8BABF5255E56FFE9FF6922A806F5FDA8" blockId="7.[227,738,193,536]" pageId="7" pageNumber="88">punctures; on third intercostae there is short additional costae with additional row of punctures untill half of length of elytra. Fifth intercosta with one row of punctures; in basal part of fourth intercosta there is region with extremely short additional costa, with some additional punctures here. External caudal bulb distinctly reduced, area between external caudal bulb and sides of elytra not divided; external and medio­internal caudal bulbs not divided; medio­internal caudal bulb shortened, transversally rounded, internally stretched.</paragraph>
<paragraph id="8BABF5255E56FFE9FF69202805E0FD68" blockId="7.[227,737,577,728]" pageId="7" pageNumber="88">
Pygidium (
<figureCitation id="132FE9A05E56FFE9FEDF20280588FDE8" box="[341,415,577,600]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="7" pageNumber="88">Fig. 28</figureCitation>
) with dense, irregularly spaced punctation, with distinct longitudinal rib in the middle and deep concavitieson its sides; with distinctly longitudinal apex in the shape of volcano with narrow, truncate top.
</paragraph>
<paragraph id="8BABF5255E56FFE9FF69216805B0FB08" blockId="7.[227,737,769,1208]" pageId="7" pageNumber="88">
Venter (
<figureCitation id="132FE9A05E56FFE9FEB121680597FCA8" box="[315,384,769,792]" captionStart="Figs" captionStartId="6.[302,352,1693,1716]" captionTargetBox="[491,1182,755,1657]" captionTargetId="figure-272@6.[487,1184,749,1658]" captionTargetPageId="6" captionText="Figs. 79. Rhyparus fijiensis sp. nov.,, holotype: 7 dorsal view; 8 ventral view; 9 lateral view. Scale lines: 1.0 mm." pageId="7" pageNumber="88">Fig. 8</figureCitation>
) shiny. Meso­metaventral plate flattened in the middle, with distinct, narrow, quite deep longitudinal furrow; punctation of meso­metaventral plate dense, regularly spaced, irregular in size; all punctures bearing short macrosetae. Abdominal ventrites moderately shiny, on sides with rows of punctures; with an additional punctured furrow in basal part. Last abdominal ventrite with dense punctures, which are slightly larger than on last but one ventrite; in basal half in the middle part with dense, very deep, longitudinal rows; in apical part in the middle with very deep groove, clearly located much below apex of pygidium.
</paragraph>
<paragraph id="8BABF5255E56FFE9FF692688067DFAC8" blockId="7.[227,737,1249,1400]" pageId="7" pageNumber="88">Mesofemora with two distinct tubercles on lower border; metafemora without any tubercles on lower border; all femora shiny, with regular, very distinct, rather small, very dense punctation; all punctures bearing small macrosetae.</paragraph>
<paragraph id="8BABF5255E56FFE9FF6927C900C9FEA8" blockId="7.[227,737,1440,1752]" lastBlockId="7.[774,1285,193,280]" pageId="7" pageNumber="88">
<emphasis id="B96029375E56FFE9FF6927C90541FA08" bold="true" box="[227,342,1440,1464]" pageId="7" pageNumber="88">Variation.</emphasis>
Length of body:
<quantity id="4CEC58C05E56FFE9FD9F27C8068DFA08" box="[533,666,1441,1464]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" metricValueMax="8.2" metricValueMin="7.4" pageId="7" pageNumber="88" unit="mm" value="7.8" valueMax="8.2" valueMin="7.4">7.48.2 mm</quantity>
. Intercostae of pronotum and elytra usually are shiny, but sometimes less shiny untill subopaque. Lateral lobes of pronotum may be more or less distinctly developed. Density of punctation of pronotal and elytral intercostae is quite distinctly variable. Degree of concentration of punctures in median intercostae of pronotum is quite variable. Usually elytral costae are convex and narrow, rarely may be flattened and quite wide. Additional rows of fine punctures observed usually on third elytral intercostae between main rows, but there are specimens with some punctures on all rows.
</paragraph>
<paragraph id="8BABF5255E56FFE9FC8C23290048FD88" blockId="7.[774,1285,320,568]" pageId="7" pageNumber="88">
<emphasis id="B96029375E56FFE9FC8C232907F1FEE8" bold="true" box="[774,998,320,344]" pageId="7" pageNumber="88">Sexual dimorphism.</emphasis>
Shape of meso­metaventral plate (in both sexes), apex of mesotibiae (in both sexes) and shape of last abdominal ventrite and pygidium of males are typical for sexual dimorphism in genus
<taxonomicName id="4C148EA65E56FFE9FC2E23A8001FFE68" authorityName="Westwood" authorityYear="1845" box="[932,1032,449,472]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B96029375E56FFE9FC2E23A8001FFE68" box="[932,1032,449,472]" italics="true" pageId="7" pageNumber="88">Rhyparus</emphasis>
</taxonomicName>
. Last abdominal ventrite and pygidium of females is distinctly elongate at apex (
<figureCitation id="132FE9A05E56FFE9FCCF20680784FDA8" box="[837,915,513,536]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="7" pageNumber="88">Fig. 28</figureCitation>
). Last abdominal ventrite and pygidium of males are not modified.
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E56FFE9FC8C200900CDFD48" pageId="7" pageNumber="88" type="etymology">
<paragraph id="8BABF5255E56FFE9FC8C200907F4FD08" blockId="7.[774,1285,608,696]" pageId="7" pageNumber="88">
<emphasis id="B96029375E56FFE9FC8C2009079DFDC8" bold="true" box="[774,906,608,632]" pageId="7" pageNumber="88">Etymology.</emphasis>
Toponymic; an adjective derived from the name of
<collectingCountry id="F303B5B55E56FFE9FC4620E80051FD28" box="[972,1094,641,664]" name="Fiji" pageId="7" pageNumber="88">Fiji Islands</collectingCountry>
where the
<typeStatus id="54AF4B875E56FFE9FB5C20E80113FD28" box="[1238,1284,641,664]" pageId="7" pageNumber="88">type</typeStatus>
series was collected.
</paragraph>
<paragraph id="8BABF5255E56FFE9FC8C208900CDFD48" blockId="7.[774,1242,736,760]" box="[774,1242,736,760]" pageId="7" pageNumber="88">
<emphasis id="B96029375E56FFE9FC8C20890771FD48" bold="true" box="[774,870,736,760]" pageId="7" pageNumber="88">Affinity.</emphasis>
See discussion and tables 1 and 2.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,302 @@
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<mods:title id="9D143417A25EFDF6C430E6147A55AA9E">Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females</mods:title>
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<mods:namePart id="27F3936D711855B2DBF49F7AC44E1A54">Minkina, Ÿukasz</mods:namePart>
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<mods:namePart id="8F4C45D3A49BD225704E65E3A37CAF08">Skelley, Paul E.</mods:namePart>
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<taxonomicName id="4C148EA65E56FFE9FC8C212807F9FCE9" ID-CoL="9WHZG" authorityName="Minkina &amp; Anichtchenko &amp; Vasiljeva &amp; Skelley" authorityYear="2022" box="[774,1006,833,857]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="stebnickae" status="sp. nov.">
<emphasis id="B96029375E56FFE9FC8C212807F9FCE9" bold="true" box="[774,1006,833,857]" italics="true" pageId="7" pageNumber="88">Rhyparus stebnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253944C5E56FFE9FC7E21280050FCE8" box="[1012,1095,833,856]" pageId="7" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E56FFE9FC8C210807FDFCC8" box="[774,1002,865,888]" pageId="7" pageNumber="88" type="description">
<paragraph id="8BABF5255E56FFE9FC8C210807FDFCC8" blockId="7.[774,1095,833,888]" box="[774,1002,865,888]" pageId="7" pageNumber="88">
(
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,
<figureCitation id="132FE9A05E56FFE9FC17210807ADFCC8" box="[925,954,865,888]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 2025. Heads: 20 R. bacanensis sp. nov.,, holotype; 21 R. buruensis sp. nov.,, holotype; 22 R. fijiensis sp. nov.,, holotype; 23 R. stebnickae sp. nov.,, holotype; 24 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 25 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." pageId="7" pageNumber="88">23</figureCitation>
,
<figureCitation id="132FE9A05E56FFE9FC4C210807F5FCC8" box="[966,994,865,888]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="7" pageNumber="88">29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E56FFE9FC8C21C900EBF96A" pageId="7" pageNumber="88" type="materials_examined">
<paragraph id="8BABF5255E56FFE9FC8C21C907EEFC48" blockId="7.[774,1285,928,1016]" pageId="7" pageNumber="88">
<materialsCitation id="3B7CFF785E56FFE9FC8C21C907E2FC48" country="Indonesia" county="Arfak Mts" elevation="1190" location="Dueibei" municipality="Manokwari distr." pageId="7" pageNumber="88" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<emphasis id="B96029375E56FFE9FC8C21C9078EFC08" bold="true" box="[774,921,928,952]" pageId="7" pageNumber="88">
<typeStatus id="54AF4B875E56FFE9FC8C21C9072AFC08" box="[774,829,928,952]" pageId="7" pageNumber="88">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F303B5B55E56FFE9FC1721C80012FC08" box="[925,1029,929,952]" name="Indonesia" pageId="7" pageNumber="88">Indonesia</collectingCountry>
,
<collectingRegion id="49D03BC75E56FFE9FB8721C800A9FC08" box="[1037,1214,929,952]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="88">West Papua prov.</collectingRegion>
,
<collectingCounty id="62CA8DA95E56FFE9FB4221C80725FC68" pageId="7" pageNumber="88">Arfak Mts</collectingCounty>
,
<collectingMunicipality id="6BCF6F5F5E56FFE9FCB721A807E2FC68" box="[829,1013,961,984]" pageId="7" pageNumber="88">Manokwari distr.</collectingMunicipality>
,
<location id="8ECBA3FE5E56FFE9FB8921A8004FFC68" LSID="urn:lsid:plazi:treatment:03BD44335E56FFE4FC8C212806F3FE68:8ECBA3FE5E56FFE9FB8921A8004FFC68" box="[1027,1112,961,984]" country="Indonesia" county="Arfak Mts" municipality="Manokwari distr." name="Dueibei" pageId="7" pageNumber="88" stateProvince="Papua Barat">Dueibei</location>
env., cca
<locationDeviation id="9A19EC075E56FFE9FB4B21A80789FC48" location="Warmare" pageId="7" pageNumber="88">
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S of Warmare
</locationDeviation>
,
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</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BABF5255E56FFE9FC8C264800EBF96A" blockId="7.[774,1286,1057,1754]" pageId="7" pageNumber="88">
<emphasis id="B96029375E56FFE9FC8C264807A7FB89" bold="true" box="[774,944,1057,1081]" pageId="7" pageNumber="88">Type material.</emphasis>
<typeStatus id="54AF4B875E56FFE9FC322648000BFB88" box="[952,1052,1057,1080]" pageId="7" pageNumber="88" type="holotype">Holotype</typeStatus>
():
<collectingCountry id="F303B5B55E56FFE9FBD2264800D3FB88" box="[1112,1220,1057,1080]" name="Indonesia" pageId="7" pageNumber="88">Indonesia</collectingCountry>
,
<collectingRegion id="49D03BC75E56FFE9FB5A2648079EFBEA" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="88">West Papua prov.</collectingRegion>
| Arfak Mts,
<quantity id="4CEC58C05E56FFE9FBA8262A007CFBEA" box="[1058,1131,1091,1114]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="7" pageNumber="88" unit="m" value="1190.0">1190m</quantity>
alt. | Dueibei env.,
<date id="FFAAD3E55E56FFE9FCB5260A07F1FBCA" box="[831,998,1123,1146]" pageId="7" pageNumber="88" value="2008-02-10" valueMax="2008-02-28" valueMin="2008-02-10">10.ii­28.ii.2008</date>
| cca
<quantity id="4CEC58C05E56FFE9FBAA260A0074FBCA" box="[1056,1123,1123,1146]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="88" unit="km" value="20.0">20 km</quantity>
S of Warmare | Manokwari distr, St. Jakl lgt. || (
<emphasis id="B96029375E56FFE9FBED26E90060FB2A" bold="true" box="[1127,1143,1152,1178]" pageId="7" pageNumber="88">
<collectionCode id="ED056DE05E56FFE9FBED26E90060FB2A" box="[1127,1143,1152,1178]" pageId="7" pageNumber="88">Ÿ</collectionCode>
</emphasis>
<collectionCode id="ED056DE05E56FFE9FBFD26EA00AEFB2A" box="[1143,1209,1155,1178]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="7" pageNumber="88" type="Herbarium">MCN</collectionCode>
).
<typeStatus id="54AF4B875E56FFE9FB4626EA072BFB0A" pageId="7" pageNumber="88" type="paratype">Paratypes</typeStatus>
(10exx.):
<specimenCount id="9D123EAC5E56FFE9FC2E26CA07C5FB0A" box="[932,978,1187,1210]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
: Coll. I.R.Sc.N.B. |
<collectingCountry id="F303B5B55E56FFE9FB1826CA0759FB6A" name="Papua New Guinea" pageId="7" pageNumber="88">Papua New Guinea</collectingCountry>
| Wau,
<collectingRegion id="49D03BC75E56FFE9FC1926AA07F5FB6A" box="[915,994,1219,1242]" country="Papua New Guinea" name="Morobe" pageId="7" pageNumber="88">Morobe</collectingRegion>
prob., | Wau Ecology Institute, |
<quantity id="4CEC58C05E56FFE9FC98268A0772FB4A" box="[786,869,1251,1274]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="88" unit="m" value="1200.0">1200 m</quantity>
, at light | I.G.: 26373 |
<date id="FFAAD3E55E56FFE9FBD2268A00DCFB4A" box="[1112,1227,1251,1274]" pageId="7" pageNumber="88" value="1981-06-18">18.vi.1981</date>
| leg. J. Van Goethem || (
<collectionCode id="ED056DE05E56FFE9FC4D276A0000FAAA" box="[967,1047,1283,1306]" pageId="7" pageNumber="88">DUBC</collectionCode>
);
<specimenCount id="9D123EAC5E56FFE9FBAD276A0040FAAA" box="[1063,1111,1283,1306]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
: Coll.R.Isc.N.B. | Canopy mission P.N.G. |
<collectingRegion id="49D03BC75E56FFE9FB9A274A00DEFA8A" box="[1040,1225,1315,1338]" country="Papua New Guinea" name="Madang" pageId="7" pageNumber="88">Madang province</collectingRegion>
| Batteta
<date id="FFAAD3E55E56FFE9FCBE272A07BAFAEA" box="[820,941,1347,1370]" pageId="7" pageNumber="88" value="1993-06-19">19­ vi­1993</date>
| light M1 | leg. Olivier Missa || (
<collectionCode id="ED056DE05E56FFE9FC84270A074CFACA" box="[782,859,1379,1402]" pageId="7" pageNumber="88">DUBC</collectionCode>
);
<specimenCount id="9D123EAC5E56FFE9FCE0270A078EFACA" box="[874,921,1379,1402]" count="2" pageId="7" pageNumber="88" type="generic">2ex.</specimenCount>
:
<collectingCountry id="F303B5B55E56FFE9FC15270A0072FACA" box="[927,1125,1379,1402]" name="Papua New Guinea" pageId="7" pageNumber="88">Papua New Guinea</collectingCountry>
: |
<collectingRegion id="49D03BC75E56FFE9FBFE270A00E9FACA" box="[1140,1278,1379,1402]" country="Papua New Guinea" name="Morobe" pageId="7" pageNumber="88">Morobe Prov.</collectingRegion>
: Wau; |
<date id="FFAAD3E55E56FFE9FCD927EA07D6FA2A" box="[851,961,1411,1434]" pageId="7" pageNumber="88" value="1982-02-21">21.ii.1982</date>
|| light trap | R.T.Bell, Coll. || (
<specimenCount id="9D123EAC5E56FFE9FC8527CA072CFA0A" box="[783,827,1443,1466]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
<collectionCode id="ED056DE05E56FFE9FCB527CA079AFA0A" box="[831,909,1443,1466]" pageId="7" pageNumber="88">CNCW</collectionCode>
;
<specimenCount id="9D123EAC5E56FFE9FC1227CA07D2FA0A" box="[920,965,1443,1466]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
NHNG);
<specimenCount id="9D123EAC5E56FFE9FBA327CA004FFA0A" box="[1065,1112,1443,1466]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
: Papua Nlle
<collectingCountry id="F303B5B55E56FFE9FB5F27CA073CFA6A" name="Guinea-Bissau" pageId="7" pageNumber="88">Guinée</collectingCountry>
| Morobe i.(19)80 | env de Gurakor | W.G.Ullrich || (
<collectionCode id="ED056DE05E56FFE9FCC3278A079FFA4A" box="[841,904,1507,1530]" pageId="7" pageNumber="88">ISEA</collectionCode>
);
<specimenCount id="9D123EAC5E56FFE9FC12278A07D0FA4A" box="[920,967,1507,1530]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
: New
<collectingCountry id="F303B5B55E56FFE9FB8F278A0043FA4A" box="[1029,1108,1507,1530]" name="Guinea-Bissau" pageId="7" pageNumber="88">Guinea</collectingCountry>
: (NE) | Wau,
<collectingRegion id="49D03BC75E56FFE9FB52278A0778F9AA" country="Papua New Guinea" name="Morobe" pageId="7" pageNumber="88">Morobe distr.</collectingRegion>
|
<quantity id="4CEC58C05E56FFE9FCF7246A07DEF9AA" box="[893,969,1539,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="88" unit="m" value="1200.0">1200m</quantity>
, 2­ 10.xi.(19)61 || J. &amp; M. Sedlacek | M.V. light trap || H.F. Howden, 1992 || (
<emphasis id="B96029375E56FFE9FC8524290708F9EA" bold="true" box="[783,799,1600,1626]" pageId="7" pageNumber="88">
<collectionCode id="ED056DE05E56FFE9FC8524290708F9EA" box="[783,799,1600,1626]" pageId="7" pageNumber="88">Ÿ</collectionCode>
</emphasis>
<collectionCode id="ED056DE05E56FFE9FC94242A074AF9EA" box="[798,861,1603,1626]" country="USA" lsid="urn:lsid:biocol.org:col:13564" name="McNeese State University" pageId="7" pageNumber="88" type="Herbarium">MCN</collectionCode>
); 3exx.:
<collectingCountry id="F303B5B55E56FFE9FC25242A0078F9EA" box="[943,1135,1603,1626]" name="Papua New Guinea" pageId="7" pageNumber="88">Papua New Guinea</collectingCountry>
| Wau,
<quantity id="4CEC58C05E56FFE9FB3C242A0111F9EA" box="[1206,1286,1603,1626]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="7" pageNumber="88" unit="ft" value="4000.0">4000 ft.</quantity>
|
<date id="FFAAD3E55E56FFE9FC9A240A07B2F9CA" box="[784,933,1635,1658]" pageId="7" pageNumber="88" value="1974-06-24" valueMax="1974-06-30" valueMin="1974-06-24">24­ 30.VI.1974</date>
| H.F. Howden ||
<taxonomicName id="4C148EA65E56FFE9FBC1240A00BAF9CA" authorityName="Westwood" authorityYear="1845" box="[1099,1197,1635,1658]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="genus">Rhyparus</taxonomicName>
|
<taxonomicName id="4C148EA65E56FFE9FB36240A073EF92A" authority="Ar." authorityName="Arrow" authorityYear="1935" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="striatus">striatus Ar.</taxonomicName>
| dt. Stebnicka (2exx.
<collectionCode id="ED056DE05E56FFE9FB9424EA0066F92A" box="[1054,1137,1667,1690]" pageId="7" pageNumber="88">CMNC</collectionCode>
;
<specimenCount id="9D123EAC5E56FFE9FBF624EA00BEF92A" box="[1148,1193,1667,1690]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
<collectionCode id="ED056DE05E56FFE9FB3B24EA00EEF92A" box="[1201,1273,1667,1690]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="7" pageNumber="88" type="Museum">FSCA</collectionCode>
);
<specimenCount id="9D123EAC5E56FFE9FC8C24CA0720F90A" box="[774,823,1699,1722]" count="1" pageId="7" pageNumber="88" type="generic">1ex.</specimenCount>
: Papua N.
<collectingCountry id="F303B5B55E56FFE9FC4024CA000FF90A" box="[970,1048,1699,1722]" name="Guinea-Bissau" pageId="7" pageNumber="88">Guinea</collectingCountry>
| Mainyanda
<date id="FFAAD3E55E56FFE9FB3824CA00E1F90A" box="[1202,1270,1699,1722]" pageId="7" pageNumber="88" value="1980-01">i.1980</date>
|
<quantity id="4CEC58C05E56FFE9FC8C24AA0753F96A" box="[774,836,1731,1754]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="88" unit="km" value="25.0">25km</quantity>
W Bulolo | W.G. Ullrich
<quantity id="4CEC58C05E56FFE9FBD824AA0098F96A" box="[1106,1167,1731,1754]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="88" unit="m" value="600.0">600m</quantity>
|| (
<collectionCode id="ED056DE05E56FFE9FB2524AA00E6F96A" box="[1199,1265,1731,1754]" pageId="7" pageNumber="88">ISEA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E59FFE7FEA422A90048FB59" lastPageId="9" lastPageNumber="90" pageId="8" pageNumber="89" type="description">
<paragraph id="8BABF5255E59FFE6FEA422A9073AFE68" blockId="8.[302,813,192,472]" pageId="8" pageNumber="89">
<emphasis id="B96029375E59FFE6FEA422A9067FFF68" bold="true" box="[302,616,192,216]" pageId="8" pageNumber="89">
Description of the
<typeStatus id="54AF4B875E59FFE6FE7422A90674FF68" box="[510,611,192,216]" pageId="8" pageNumber="89" type="holotype">holotype</typeStatus>
.
</emphasis>
Dorsum (
<figureCitation id="132FE9A05E59FFE6FD5B22A80737FF68" box="[721,800,193,216]" captionStart="Figs" captionStartId="8.[302,352,1693,1716]" captionTargetBox="[450,1200,751,1654]" captionTargetId="figure-239@8.[448,1205,750,1660]" captionTargetPageId="8" captionText="Figs. 1012. Rhyparus stebnickae sp. nov.,, holotype: 10 dorsal view; 11 ventral view; 12 lateral view. Scale lines: 1.0 mm." pageId="8" pageNumber="89">Fig. 10</figureCitation>
). Length:
<quantity id="4CEC58C05E59FFE6FE0F228805CFFF48" box="[389,472,225,248]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="89" unit="mm" value="6.5">6.5 mm</quantity>
; maximum width:
<quantity id="4CEC58C05E59FFE6FD13228806FCFF48" box="[665,747,225,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="89" unit="mm" value="1.9">1.9 mm</quantity>
. Body moderate­ to large­sized for members of this genus, elongate, not so distinctly convex, flattened in central part; shiny; apparently almost glabrous, though partly clothed with very small yellowish macrosetae on head and all longitudinal costae on pronotum and elytra. Brownish to dark brown; antennae, tarsomeres and mouth parts pale brown.
</paragraph>
<paragraph id="8BABF5255E59FFE6FEA4206807C5FE08" blockId="8.[302,812,513,696]" lastBlockId="8.[850,1361,193,440]" pageId="8" pageNumber="89">
Head (
<figureCitation id="132FE9A05E59FFE6FEF8206805ACFDA8" box="[370,443,513,536]" captionStart="Figs" captionStartId="13.[227,277,1463,1486]" captionTargetBox="[349,1168,359,1425]" captionTargetId="figure-16@13.[342,1170,355,1428]" captionTargetPageId="13" captionText="Figs. 2025. Heads: 20 R. bacanensis sp. nov.,, holotype; 21 R. buruensis sp. nov.,, holotype; 22 R. fijiensis sp. nov.,, holotype; 23 R. stebnickae sp. nov.,, holotype; 24 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 25 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 1.0 mm." pageId="8" pageNumber="89">Fig. 23</figureCitation>
) shiny; transversely sub­hexagonal; clypeus trapezoidal in outline, anteriorly weakly sinuate, on sides weakly upturned as obtuse, quite distinct tooth, and later sinuous on either side; genae distinctly more excavate than eyes; clypeal disc distinctly convex, ringed by a deep groove; convexity with a pair of very indistinct, very short, convergent ridges, nearly on whole surface with distinct, fine punctures bearing small macrosetae. Frons with four distinct, longitudinal ridges with similar structure as ridges on clypeal convexity. Head covered by quite regularly spaced, quite dense, moderately large punctures bearing small macrosetae.
</paragraph>
<paragraph id="8BABF5255E59FFE6FCD823880147FD08" blockId="8.[850,1361,481,696]" pageId="8" pageNumber="89">Pronotum shiny; with eight low but distinct costae and seven intercostae, with two lateral, rounded lobes on each side.Anterior lobes somewhat lower and distinctly narrower than posterior; posterior lobes on the top are the widest part of pronotum. Costae of third and fourth pair not interrupted in basal part of apical half, very gently convergent,</paragraph>
<caption id="DF6BA5AD5E59FFE6FEA424F40685F964" pageId="8" pageNumber="89" startId="8.[302,352,1693,1716]" targetBox="[450,1200,751,1654]" targetPageId="8" targetType="figure">
<paragraph id="8BABF5255E59FFE6FEA424F40685F964" blockId="8.[302,1360,1693,1748]" pageId="8" pageNumber="89">
Figs. 1012.
<taxonomicName id="4C148EA65E59FFE6FE3624F7068AF905" authorityName="Minkina &amp; Anichtchenko &amp; Vasiljeva &amp; Skelley" authorityYear="2022" box="[444,669,1694,1717]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="stebnickae" status="sp. nov.">
<emphasis id="B96029375E59FFE6FE3624F7068AF905" box="[444,669,1694,1717]" italics="true" pageId="8" pageNumber="89">Rhyparus stebnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253944C5E59FFE6FD2E24F406E0F904" box="[676,759,1693,1716]" pageId="8" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
,, holotype: 10 dorsal view; 11 ventral view; 12 lateral view. Scale lines: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BABF5255E58FFE7FD8A22F807FFFF14" blockId="9.[512,1000,145,164]" box="[512,1000,145,164]" pageId="9" pageNumber="90">
<emphasis id="B96029375E58FFE7FD8A22F807FFFF14" box="[512,1000,145,164]" italics="true" pageId="9" pageNumber="90">Minkina Ł., Anichtchenko A., Vasiljeva A., Skelley P.E.</emphasis>
</paragraph>
<paragraph id="8BABF5255E58FFE7FF6922A806F5FE4D" blockId="9.[227,738,193,509]" pageId="9" pageNumber="90">distinctly convergent in the middle of apical third; median pair of costae in apical half become very low but without any cutpoint of interruption; second pair of costae distinctly interrupted in basal part of apical half; costae on each side with small punctures bearing very small macrosetae.All intercostae in anterior part without additional short costae. Median intercostae with dense punctures, very distinctly concentrated around median part, all intercostae in basal part with distinct, dense punctation.</paragraph>
<paragraph id="8BABF5255E58FFE7FF69204E0628FD8E" blockId="9.[227,575,551,574]" box="[227,575,551,574]" pageId="9" pageNumber="90">Scutellum almost imperceptible.</paragraph>
<paragraph id="8BABF5255E58FFE7FF6920010624FB38" blockId="9.[227,737,616,1160]" pageId="9" pageNumber="90">Elytra shiny. Each elytron with six elevated but flattened on the top costae, and five flat intercostae. Costae on sides with very small punctures bearing very small macrosetae. Preapical glandular area relatively small. Intercostae first to fourth with two distinct rows of punctures; on third intercostae there is short additional costae with additional row of punctures untill half of length of elytra. Fifth intercosta with one row of punctures; in basal part of fourth intercosta there is region with extremely short additional costa, with some additional punctures here. External caudal bulb distinctly reduced, area between external caudal bulb and sides of elytra not divided; external and medio­internal caudal bulbs not divided; medio­internal caudal bulb shortened, transverse, internally stretched.</paragraph>
<paragraph id="8BABF5255E58FFE7FF6926DB05D9FAFB" blockId="9.[227,737,1202,1355]" pageId="9" pageNumber="90">
Pygidium (
<figureCitation id="132FE9A05E58FFE7FEDF26DB05B7FB79" box="[341,416,1202,1225]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="9" pageNumber="90">Fig. 29</figureCitation>
) with dense, irregularly spaced punctation, with very distinct longitudinal rib in the middle and deep excision on its sides; with distinctly longitudinal apex in the shape of volcano with sinuate top.
</paragraph>
<paragraph id="8BABF5255E58FFE7FF69271C0763FEA9" blockId="9.[227,737,1397,1745]" lastBlockId="9.[774,1285,193,281]" pageId="9" pageNumber="90">
Venter (
<figureCitation id="132FE9A05E58FFE7FEB3271C0590FA3C" box="[313,391,1397,1420]" captionStart="Figs" captionStartId="8.[302,352,1693,1716]" captionTargetBox="[450,1200,751,1654]" captionTargetId="figure-239@8.[448,1205,750,1660]" captionTargetPageId="8" captionText="Figs. 1012. Rhyparus stebnickae sp. nov.,, holotype: 10 dorsal view; 11 ventral view; 12 lateral view. Scale lines: 1.0 mm." pageId="9" pageNumber="90">Fig. 11</figureCitation>
) shiny. Meso­metaventral plate flattened in the middle, with distinct, narrow, quite deep longitudinal furrow; punctation of meso­metaventral plate dense, regularly spaced, irregular in size; all punctures bearing short macrosetae. Abdominal ventrites moderately shiny, on sides with rows of punctures; with an additional punctured furrow in basal part. Last abdominal ventrite with dense punctures, which are slightly larger than on last but one ventrite; in basal half in the middle part with dense, very deep, longitudinal rows; in apical part in the middle with very deep groove, clearly located much below apex of pygidium.
</paragraph>
<paragraph id="8BABF5255E58FFE7FC8C232A009AFE6C" blockId="9.[774,1285,323,476]" pageId="9" pageNumber="90">Mesofemora with two distinct tubercles on lower border; metafemora with very weak tubercles on lower border; all femora shiny, with regular, very distinct, rather small, very dense punctation; all punctures bearing small macrosetae.</paragraph>
<paragraph id="8BABF5255E58FFE7FC8C206F0055FC74" blockId="9.[774,1286,518,964]" pageId="9" pageNumber="90">
<emphasis id="B96029375E58FFE7FC8C206F076DFDAE" bold="true" box="[774,890,518,542]" pageId="9" pageNumber="90">Variation.</emphasis>
Length of body: 6.2­ 7.0 mm. Intercostae of pronotum and elytra usually are shiny, but sometimes may be somewhat less shiny. Lateral lobes of pronotum may be more or less weakly developed. Density of punctation of pronotal and elytral intercostae is quite distinctly variable. Degree of concentration of punctures in median intercostae of pronotum is very weakly variable. Usually elytral costae are flattened and wide, rarely may be somewhat more distinctly convex and quite narrow. Additional rows of fine punctures observed usually on third elytral intercostae between main rows, but there are specimens with simple punctures on all rows.
</paragraph>
<paragraph id="8BABF5255E58FFE7FC8C21870048FB59" blockId="9.[774,1285,1006,1257]" pageId="9" pageNumber="90">
<emphasis id="B96029375E58FFE7FC8C218707F1FBB6" bold="true" box="[774,998,1006,1030]" pageId="9" pageNumber="90">Sexual dimorphism.</emphasis>
Shape of meso­metaventral plate (in both sexes), apex of mesotibiae (in both sexes) and shape of last abdominal ventrite and pygidium of males are typical for sexual dimorphism in genus
<taxonomicName id="4C148EA65E58FFE7FC2E2618001FFB38" authorityName="Westwood" authorityYear="1845" box="[932,1032,1137,1160]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B96029375E58FFE7FC2E2618001FFB38" box="[932,1032,1137,1160]" italics="true" pageId="9" pageNumber="90">Rhyparus</emphasis>
</taxonomicName>
. Last abdominal ventrite and pygidium of females is distinctly elongate at apex (
<figureCitation id="132FE9A05E58FFE7FCCF26DB0784FB79" box="[837,915,1202,1225]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="9" pageNumber="90">Fig. 29</figureCitation>
). Last abdominal ventrite and pygidium of males are not modified.
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E58FFE4FC8C277A06F3FE68" lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90" type="etymology">
<paragraph id="8BABF5255E58FFE7FC8C277A00FAFADB" blockId="9.[774,1285,1298,1387]" pageId="9" pageNumber="90">
<emphasis id="B96029375E58FFE7FC8C277A0791FA9B" bold="true" box="[774,902,1299,1323]" pageId="9" pageNumber="90">Etymology.</emphasis>
Patronymic. Dedicated to
<emphasis id="B96029375E58FFE7FB12277A0112FA9A" bold="true" box="[1176,1285,1298,1322]" pageId="9" pageNumber="90">Zdzis˘awa</emphasis>
Stebnicka who noticed unusual sexual dimorphism in the genus
<taxonomicName id="4C148EA65E58FFE7FC5C273D002AFADB" authorityName="Westwood" authorityYear="1845" box="[982,1085,1364,1387]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="genus">Rhyparus</taxonomicName>
at the first time.
</paragraph>
<paragraph id="8BABF5255E58FFE4FC8C27FC06F3FE68" blockId="9.[774,1285,1429,1745]" lastBlockId="10.[302,813,193,472]" lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90">
<emphasis id="B96029375E58FFE7FC8C27FC0773FA1D" bold="true" box="[774,868,1429,1453]" pageId="9" pageNumber="90">Affinity.</emphasis>
See discussion and tables 1 and 2. Additional problem is that some specimens of
<taxonomicName id="4C148EA65E58FFE7FB6727DF07F3FA5D" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96029375E58FFE7FB6727DF074EFA5E" italics="true" pageId="9" pageNumber="90">R. gracilis</emphasis>
Arrow, 1905
</taxonomicName>
(which them
<taxonomicName id="4C148EA65E58FFE7FBFC27BE0112FA5E" authorityName="Minkina &amp; Anichtchenko &amp; Vasiljeva &amp; Skelley" authorityYear="2022" box="[1142,1285,1495,1518]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="stebnickae" status="sp. nov.">
<emphasis id="B96029375E58FFE7FBFC27BE0112FA5E" box="[1142,1285,1495,1518]" italics="true" pageId="9" pageNumber="90">R. stebnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253944C5E58FFE7FC8C279E074DF9BE" box="[774,858,1527,1550]" pageId="9" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
was collected on the same localities on
<collectingCountry id="F303B5B55E58FFE7FC8C247E07CFF99E" box="[774,984,1559,1582]" name="Papua New Guinea" pageId="9" pageNumber="90">Papua New Guinea</collectingCountry>
) may have similar structure of elytral costae, identique structure of caudal bulbs and somewhat similar shape of lateral lobes of pronotum (posterior lateral lobes can be widely, regularly rounded, weakly developed, with very small sinuation between them and anterior lateral lobes). Anyway females of
<taxonomicName id="4C148EA65E5BFFE4FDD822A806D5FF68" authorityName="Arrow" authorityYear="1905" box="[594,706,193,216]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96029375E5BFFE4FDD822A806D5FF68" box="[594,706,193,216]" italics="true" pageId="10" pageNumber="91">R. gracilis</emphasis>
</taxonomicName>
have normally developed pygidium and last abdominal ventrite. Males of
<taxonomicName id="4C148EA65E5BFFE4FE6323680642FEA8" authorityName="Arrow" authorityYear="1905" box="[489,597,257,280]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96029375E5BFFE4FE6323680642FEA8" box="[489,597,257,280]" italics="true" pageId="10" pageNumber="91">R. gracilis</emphasis>
</taxonomicName>
can be distinguished by: more distinctly convex body, posterior lateral lobes of pronotum never more distinctly developed than anterior, punctures in median intercostae of pronotum never so distinctly concentrated nearby the middle (never create so distinct longitudinal line) and elytral intercostae usually matt.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,268 @@
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<mods:title id="E440B12D0B04F48E4F644777094CC0DF">Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females</mods:title>
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<paragraph id="8BABF5255E5BFFE4FEA4200506F2FD33" blockId="10.[302,741,620,676]" box="[302,741,620,644]" pageId="10" pageNumber="91">
<heading id="D0E342495E5BFFE4FEA4200506F2FD33" box="[302,741,620,644]" fontSize="10" level="2" pageId="10" pageNumber="91" reason="7">
<taxonomicName id="4C148EA65E5BFFE4FEA4200506F2FD33" ID-CoL="4SV3W" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[302,741,620,644]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="approximans">
<emphasis id="B96029375E5BFFE4FEA420050625FD34" bold="true" box="[302,562,620,644]" italics="true" pageId="10" pageNumber="91">Rhyparus approximans</emphasis>
Fairmaire, 1893
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30EA6AE5E5BFFE4FEA420E40610FD14" box="[302,519,653,676]" pageId="10" pageNumber="91" type="description">
<paragraph id="8BABF5255E5BFFE4FEA420E40610FD14" blockId="10.[302,741,620,676]" box="[302,519,653,676]" pageId="10" pageNumber="91">
(
<figureCitation id="132FE9A05E5BFFE4FEBC20E40591FD14" box="[310,390,653,676]" captionStart="Figs" captionStartId="14.[302,352,1464,1487]" captionTargetBox="[463,1196,355,1425]" captionTargetId="figure-122@14.[462,1196,355,1428]" captionTargetPageId="14" captionText="Figs. 2633.Abdominal apexes with black­white miniatures of pygidium on left­down side: 26 R. bacanensis sp. nov.,, holotype; 27 R. buruensis sp. nov.,, holotype; 28 R. fijiensis sp. nov.,, holotype; 29 R. stebnickae sp. nov.,, holotype; 30 R. approximans Fairmaire, 1893,; 31 R. loebli Paulian, 1983,; 32 R. paraphilippinensis Ochi &amp; Kakizoe, 2021,; 33 R. sepikensis Stebnicka, 1998,, paratype. Scale lines: 0.5 mm." pageId="10" pageNumber="91">Fig. 30</figureCitation>
,
<figureCitation id="132FE9A05E5BFFE4FE1820E405C0FD14" box="[402,471,653,676]" captionStart="Figs" captionStartId="15.[227,277,896,919]" captionTargetBox="[227,1285,189,951]" captionTargetId="figure-16@15.[479,1035,189,860]" captionTargetPageId="15" captionText="Figs. 3436. Rhyparus approximans Fairmaire, 1893, lateral view. Scale lines: 1.0 mm." pageId="10" pageNumber="91">3436</figureCitation>
, 40)
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E5BFFE4FEA420A7073AFC18" pageId="10" pageNumber="91" type="reference_group">
<paragraph id="8BABF5255E5BFFE4FEA420A706DCFCB5" blockId="10.[302,813,717,936]" pageId="10" pageNumber="91">
<taxonomicName id="4C148EA65E5BFFE4FEA420A70731FD54" ID-CoL="4SV3W" authority="Fairmaire, 1893: 145" authorityName="Fairmaire" authorityPageNumber="145" authorityYear="1893" box="[302,806,717,741]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="approximans">
<emphasis id="B96029375E5BFFE4FEA420A70639FD55" box="[302,558,718,741]" italics="true" pageId="10" pageNumber="91">Rhyparus approximans</emphasis>
Fairmaire, 1893: 145
</taxonomicName>
.
<typeStatus id="54AF4B875E5BFFE4FEA420870574FCB5" box="[302,355,750,773]" pageId="10" pageNumber="91">Type</typeStatus>
locality: “Bornéo occ., Sambas”.
</paragraph>
<paragraph id="8BABF5255E5BFFE4FEA421660709FC96" blockId="10.[302,813,717,936]" box="[302,798,783,806]" pageId="10" pageNumber="91">
<treatmentCitationGroup id="AB04D20B5E5BFFE4FEA421660709FC96" box="[302,798,783,806]" pageId="10" pageNumber="91">
<taxonomicName id="4C148EA65E5BFFE4FEA42166063FFC96" ID-CoL="4SV3W" authorityName="Fairmaire" authorityYear="1893" box="[302,552,783,806]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="approximans">
<emphasis id="B96029375E5BFFE4FEA42166063FFC96" box="[302,552,783,806]" italics="true" pageId="10" pageNumber="91">Rhyparus approximans</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AB5D3345E5BFFE4FDBD2166070DFC96" author="Balthasar V." box="[567,794,783,806]" page="610" pageId="10" pageNumber="91" year="1964">
<bibRefCitation id="EF8588D45E5BFFE4FDBD2166070DFC96" author="Balthasar V." box="[567,794,783,806]" pageId="10" pageNumber="91" pagination="1 - 652" refId="ref8914" refString="Balthasar V. 1964: Monographie der Scarabaeidae und Aphodidae der palaearktischen und Orientalischen Region Aphodiidae. Tchechoslowakische Akademie der Wissenschaften Prag. 3: 1 - 652." type="book chapter" year="1964">Balthasar, 1964: 610</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BABF5255E5BFFE4FEA4214605B8FCD7" blockId="10.[302,813,717,936]" pageId="10" pageNumber="91">
<taxonomicName id="4C148EA65E5BFFE4FEA421460623FCF6" ID-CoL="4SV3W" authorityName="Fairmaire" authorityYear="1893" box="[302,564,815,838]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="approximans">
<emphasis id="B96029375E5BFFE4FEA421460623FCF6" box="[302,564,815,838]" italics="true" pageId="10" pageNumber="91">Rhyparus approximans</emphasis>
</taxonomicName>
: Dellacasa, 1988: 89 (catalogue).
</paragraph>
<paragraph id="8BABF5255E5BFFE4FEA4211806E2FC37" blockId="10.[302,813,717,936]" box="[302,757,880,904]" pageId="10" pageNumber="91">
<treatmentCitationGroup id="AB04D20B5E5BFFE4FEA4211806E2FC37" box="[302,757,880,904]" pageId="10" pageNumber="91">
<taxonomicName id="4C148EA65E5BFFE4FEA42118063FFC38" ID-CoL="4SV3W" authorityName="Fairmaire" authorityYear="1893" box="[302,552,881,904]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="approximans">
<emphasis id="B96029375E5BFFE4FEA42118063FFC38" box="[302,552,881,904]" italics="true" pageId="10" pageNumber="91">Rhyparus approximans</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AB5D3345E5BFFE4FDBD211906E6FC37" author="Bordat P." box="[567,753,880,903]" page="92" pageId="10" pageNumber="91" year="1996">
<bibRefCitation id="EF8588D45E5BFFE4FDBD211906E6FC37" author="Bordat P." box="[567,753,880,903]" pageId="10" pageNumber="91" pagination="83 - 93" refId="ref8943" refString="Bordat P. 1996: Quatre nouvelles esp E ces de Rhyparus de Borneo. Nouvelle Revue d'Entomologie, 13 (1): 83 - 93." type="journal article" year="1996">Bordat, 1996: 92</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BABF5255E5BFFE4FEA421F8073AFC18" blockId="10.[302,813,717,936]" box="[302,813,913,936]" pageId="10" pageNumber="91">
<treatmentCitationGroup id="AB04D20B5E5BFFE4FEA421F8073AFC18" box="[302,813,913,936]" pageId="10" pageNumber="91">
<taxonomicName id="4C148EA65E5BFFE4FEA421F80632FC18" ID-CoL="4SV3W" authorityName="Fairmaire" authorityYear="1893" box="[302,549,913,936]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="approximans">
<emphasis id="B96029375E5BFFE4FEA421F80632FC18" box="[302,549,913,936]" italics="true" pageId="10" pageNumber="91">Rhyparus approximans</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AB5D3345E5BFFE4FDB821F8073EFC18" author="Mencl L. &amp; Rakovic M. &amp; Kral D." box="[562,809,913,936]" page="493" pageId="10" pageNumber="91" year="2013">
<bibRefCitation id="EF8588D45E5BFFE4FDB821F8073EFC18" author="Mencl L. &amp; Rakovic M. &amp; Kral D." box="[562,809,913,936]" pageId="10" pageNumber="91" pagination="487 - 498" refId="ref9083" refString="Mencl L., Rakovic M. &amp; Kral D. 2013: A new species of the genus Rhyparus from the Oriental Region having an accessory costa on each elytron. Studies and Reports. Taxonomical Series 9 (2): 487 - 498." type="journal article" year="2013">
Mencl
<emphasis id="B96029375E5BFFE4FDF621F806A7FC18" box="[636,688,913,936]" italics="true" pageId="10" pageNumber="91">et al.</emphasis>
, 2013: 493
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E5BFFE4FEA421BB06D3FADF" pageId="10" pageNumber="91" type="materials_examined">
<paragraph id="8BABF5255E5BFFE4FEA421BB06D3FADF" blockId="10.[302,813,978,1391]" pageId="10" pageNumber="91">
<emphasis id="B96029375E5BFFE4FEA421BB069CFC5A" bold="true" box="[302,651,978,1002]" pageId="10" pageNumber="91">Additional material examined.</emphasis>
(6exx.): 2exx.:
<collectingCountry id="F303B5B55E5BFFE4FEA4219B0586FBB9" box="[302,401,1010,1033]" name="Malaysia" pageId="10" pageNumber="91">Malaysia</collectingCountry>
, Borneo,Trus Madi Mts,
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, local coll. (
<collectionCode id="ED056DE05E5BFFE4FE2D267A05E9FB9A" box="[423,510,1043,1066]" pageId="10" pageNumber="91">CNCW</collectionCode>
);
<materialsCitation id="3B7CFF785E5BFFE4FD9A267A0599FBDB" collectingDate="2013-04-04" collectionCode="DUBC" collectorName="N Borneo" country="Malaysia" location="Mt. Trus Madi" municipality="Keningau distr." pageId="10" pageNumber="91" specimenCount="1" stateProvince="Sabah">
<specimenCount id="9D123EAC5E5BFFE4FD9A267A0656FB9A" box="[528,577,1043,1066]" count="1" pageId="10" pageNumber="91" type="generic">1ex.</specimenCount>
:
<collectingCountry id="F303B5B55E5BFFE4FDC6267A06A5FB9A" box="[588,690,1043,1066]" name="Malaysia" pageId="10" pageNumber="91">Malaysia</collectingCountry>
,
<collectorName id="26E190F35E5BFFE4FD36267A073EFB9A" box="[700,809,1043,1066]" pageId="10" pageNumber="91">N Borneo</collectorName>
,
<collectingRegion id="49D03BC75E5BFFE4FEA4265D0565FBFB" box="[302,370,1076,1099]" country="Malaysia" name="Sabah" pageId="10" pageNumber="91">Sabah</collectingRegion>
,
<collectingMunicipality id="6BCF6F5F5E5BFFE4FEF1265D060CFBFB" box="[379,539,1076,1099]" pageId="10" pageNumber="91">Keningau distr.</collectingMunicipality>
,
<location id="8ECBA3FE5E5BFFE4FDAD265D06D6FBFB" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE4FEA4200506E0FA00:8ECBA3FE5E5BFFE4FDAD265D06D6FBFB" box="[551,705,1076,1099]" country="Malaysia" municipality="Keningau distr." name="Mt. Trus Madi" pageId="10" pageNumber="91" stateProvince="Sabah">Mt. Trus Madi</location>
,
<date id="FFAAD3E55E5BFFE4FD40265D073AFBFB" box="[714,813,1076,1099]" pageId="10" pageNumber="91" value="2013-04-04">
<collectingDate id="EFEE2A0D5E5BFFE4FD40265D073AFBFB" box="[714,813,1076,1099]" pageId="10" pageNumber="91" value="2013-04-04">4.iv.2013</collectingDate>
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<emphasis id="B96029375E5BFFE4FEA427F005D5FA01" bold="true" box="[302,450,1433,1457]" pageId="10" pageNumber="91">Distribution.</emphasis>
Known from Borneo Island.
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View file

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<taxonomicName id="4C148EA65E5BFFE4FCD8275D07A3FADB" ID-CoL="8TJR4" authority="Ochi &amp; Kakizoe, 2021: 31" authorityName="Ochi &amp; Kakizoe" authorityPageNumber="31" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="paraphilippinensis">
<emphasis id="B96029375E5BFFE4FCD8275D009AFAFB" box="[850,1165,1332,1355]" italics="true" pageId="10" pageNumber="91">Rhyparus paraphilippinensis</emphasis>
<materialsCitation id="3B7CFF785E5BFFE4FB1F275D07A3FADB" country="Philippines" county="Mindanao Island" location="Sibagat" municipality="Caraga Region" pageId="10" pageNumber="91" specimenCount="1" stateProvince="Agusan del Sur">
<bibRefCitation id="EF8588D45E5BFFE4FB1F275D07A3FADB" author="Ochi, T. &amp; Kakizoe, S. &amp; Minkina, Y. &amp; Kawahara, M. &amp; Cabras, A. A." pageId="10" pageNumber="91" pagination="17 - 42" refId="ref9157" refString="Ochi, T., Kakizoe, S., Minkina, Y., Kawahara, M. &amp; Cabras, A. A. 2021: A revisional study of the genus Rhyparus from the Philippines. Kogane, 24: 17 - 42." type="journal article" year="2021">Ochi &amp; Kakizoe, 2021: 31</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, figs: 5, 18, 30, 38, 45.
<typeStatus id="54AF4B875E5BFFE4FB30273D00E7FADB" box="[1210,1264,1364,1387]" pageId="10" pageNumber="91">Type</typeStatus>
locality: “
<collectingCountry id="F303B5B55E5BFFE4FCD6271C07CCFA3C" box="[860,987,1397,1420]" name="Philippines" pageId="10" pageNumber="91">Philippines</collectingCountry>
,
<collectingCounty id="62CA8DA95E5BFFE4FC6F271C0089FA3C" box="[997,1182,1397,1420]" pageId="10" pageNumber="91">Mindanao Island</collectingCounty>
,
<collectingMunicipality id="6BCF6F5F5E5BFFE4FB23271C015AFA3C" box="[1193,1357,1397,1420]" pageId="10" pageNumber="91">Caraga Region</collectingMunicipality>
,
<collectingRegion id="49D03BC75E5BFFE4FCD827FC07E6FA1C" box="[850,1009,1429,1452]" country="Philippines" name="Agusan del Sur" pageId="10" pageNumber="91">Agusan del sur</collectingRegion>
,
<location id="8ECBA3FE5E5BFFE4FC7527FC0042FA1C" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5BFFE4FC7527FC0042FA1C" box="[1023,1109,1429,1452]" country="Philippines" county="Mindanao Island" municipality="Caraga Region" name="Sibagat" pageId="10" pageNumber="91" stateProvince="Agusan del Sur">Sibagat</location>
”.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BABF5255E5BFFE4FCD827DF0123FA5D" blockId="10.[850,1361,1332,1518]" pageId="10" pageNumber="91">
<treatmentCitationGroup id="AB04D20B5E5BFFE4FCD827DF0123FA5D" pageId="10" pageNumber="91">
<taxonomicName id="4C148EA65E5BFFE4FCD827DF00B9FA7D" ID-CoL="4SV4J" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[850,1198,1462,1485]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96029375E5BFFE4FCD827DF0001FA7D" box="[850,1046,1462,1485]" italics="true" pageId="10" pageNumber="91">Rhyparus gracilis</emphasis>
Arrow, 1905
</taxonomicName>
:
<treatmentCitation id="0AB5D3345E5BFFE4FB3327DF0012FA5D" author="Anichtchenko, A. V. &amp; Minkina, L. &amp; Vasiljeva, A. &amp; Medina, M. N. D." page="102" pageId="10" pageNumber="91" year="2021">
<bibRefCitation id="EF8588D45E5BFFE4FB3327DF0012FA5D" author="Anichtchenko, A. V. &amp; Minkina, L. &amp; Vasiljeva, A. &amp; Medina, M. N. D." pageId="10" pageNumber="91" pagination="99 - 111" refId="ref8853" refString="Anichtchenko, A. V., Minkina, L., Vasiljeva, A. &amp; Medina, M. N. D. 2021: A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao. A cta entomologica Musei nationalis Pragae 61 (1): 99 - 111." type="journal article" year="2021">
Anichtchenko
<emphasis id="B96029375E5BFFE4FCD827BE079EFA5E" box="[850,905,1495,1518]" italics="true" pageId="10" pageNumber="91">et al.</emphasis>
, 2021: 102
</bibRefCitation>
</treatmentCitation>
, figs: 3a­c, 9, 15, 21, 27a­b.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E5BFFE5FCD8247E0697FDA8" lastPageId="11" lastPageNumber="92" pageId="10" pageNumber="91" type="materials_examined">
<paragraph id="8BABF5255E5BFFE4FCD8247E0147F961" blockId="10.[850,1361,1559,1745]" pageId="10" pageNumber="91">
<emphasis id="B96029375E5BFFE4FCD8247E00B1F99F" bold="true" box="[850,1190,1559,1583]" pageId="10" pageNumber="91">Additional material examined.</emphasis>
(22exx.):
<materialsCitation id="3B7CFF785E5BFFE4FA85247E0009F961" collectingDate="2022-03-31" collectingDateMax="2022-04-03" collectingDateMin="2022-03-31" collectionCode="NMPC, DUBC" collectorName="Bocak &amp; A. Vasiljeva" country="Philippines" county="Eureka Mt." elevation="730" latitude="8.691617" location="Mindanao" longLatPrecision="1" longitude="125.23785" municipality="Gingoog" pageId="10" pageNumber="91" specimenCount="1">
8exx.:
<collectingCountry id="F303B5B55E5BFFE4FCD8245107DBF9FF" box="[850,972,1592,1615]" name="Philippines" pageId="10" pageNumber="91">Philippines</collectingCountry>
,
<location id="8ECBA3FE5E5BFFE4FC5F24510054F9FF" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5BFFE4FC5F24510054F9FF" box="[981,1091,1592,1615]" country="Philippines" county="Eureka Mt." latitude="8.691617" longLatPrecision="1" longitude="125.23785" municipality="Gingoog" name="Mindanao" pageId="10" pageNumber="91">Mindanao</location>
,
<location id="8ECBA3FE5E5BFFE4FBC6245100F7F9FF" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5BFFE4FBC6245100F7F9FF" box="[1100,1248,1592,1615]" country="Philippines" county="Eureka Mt." latitude="8.691617" longLatPrecision="1" longitude="125.23785" municipality="Gingoog" name="Bagongsilang" pageId="10" pageNumber="91">Bagongsilang</location>
, 1996, leg.
<collectorName id="26E190F35E5BFFE4FCD824300780F9C0" box="[850,919,1625,1648]" pageId="10" pageNumber="91">Bocak</collectorName>
(
<collectionCode id="ED056DE05E5BFFE4FC2F243007E0F9C0" box="[933,1015,1625,1648]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="91">NMPC</collectionCode>
); 3exx.:
<collectingCountry id="F303B5B55E5BFFE4FBDF243000C6F9C0" box="[1109,1233,1625,1648]" name="Philippines" pageId="10" pageNumber="91">Philippines</collectingCountry>
,
<location id="8ECBA3FE5E5BFFE4FB562430015BF9C0" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5BFFE4FB562430015BF9C0" box="[1244,1356,1625,1648]" country="Philippines" county="Eureka Mt." latitude="8.691617" longLatPrecision="1" longitude="125.23785" municipality="Gingoog" name="Mindanao" pageId="10" pageNumber="91">Mindanao</location>
,
<collectingMunicipality id="6BCF6F5F5E5BFFE4FCD8241007A0F920" box="[850,951,1657,1680]" pageId="10" pageNumber="91">Gingoog</collectingMunicipality>
,
<collectingCounty id="62CA8DA95E5BFFE4FC4124100045F920" box="[971,1106,1657,1680]" pageId="10" pageNumber="91">Eureka Mt.</collectingCounty>
,
<quantity id="4CEC58C05E5BFFE4FBEC241000B0F920" box="[1126,1191,1657,1680]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="91" unit="m" value="730.0">
<elevation id="003912165E5BFFE4FBEC241000B0F920" box="[1126,1191,1657,1680]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="91" unit="m" value="730.0">730m</elevation>
</quantity>
,
<geoCoordinate id="EE2093E25E5BFFE4FB312410015FF920" box="[1211,1352,1657,1680]" degrees="8" direction="north" minutes="41. 4970" orientation="latitude" pageId="10" pageNumber="91" precision="1" value="8.691617">8°41.4970N</geoCoordinate>
,
<geoCoordinate id="EE2093E25E5BFFE4FCD824F307F8F901" box="[850,1007,1690,1713]" degrees="125" direction="east" minutes="14.2710" orientation="longitude" pageId="10" pageNumber="91" precision="1" value="125.23785">125°14.2710E</geoCoordinate>
,
<date id="FFAAD3E55E5BFFE4FC7724F300ACF901" box="[1021,1211,1690,1713]" pageId="10" pageNumber="91" value="2022-03-31" valueMax="2022-04-03" valueMin="2022-03-31">
<collectingDate id="EFEE2A0D5E5BFFE4FC7724F300ACF901" box="[1021,1211,1690,1713]" pageId="10" pageNumber="91" value="2022-03-31" valueMax="2022-04-03" valueMin="2022-03-31">31.iii 3.iv.2022</collectingDate>
</date>
, leg.
<collectorName id="26E190F35E5BFFE4FB7D24F30768F961" pageId="10" pageNumber="91">A.Vasiljeva</collectorName>
(
<collectionCode id="ED056DE05E5BFFE4FC0624D307CBF961" box="[908,988,1722,1745]" pageId="10" pageNumber="91">DUBC</collectionCode>
);
<specimenCount id="9D123EAC5E5BFFE4FC6724D30009F961" box="[1005,1054,1722,1745]" count="1" pageId="10" pageNumber="91" type="generic">1ex.</specimenCount>
</materialsCitation>
: the same but: leg. A.Anicht­
</paragraph>
<paragraph id="8BABF5255E5AFFE5FD8A22F807FFFF14" blockId="11.[512,1000,145,164]" box="[512,1000,145,164]" pageId="11" pageNumber="92">
<emphasis id="B96029375E5AFFE5FD8A22F807FFFF14" box="[512,1000,145,164]" italics="true" pageId="11" pageNumber="92">Minkina Ł., Anichtchenko A., Vasiljeva A., Skelley P.E.</emphasis>
</paragraph>
<paragraph id="8BABF5255E5AFFE5FF6922A80697FDA8" blockId="11.[227,740,193,536]" pageId="11" pageNumber="92">
<materialsCitation id="3B7CFF785E5AFFE5FF6922A805D5FE48" collectingDate="2022-03-21" collectingDateMax="2022-03-28" collectingDateMin="2022-03-21" collectionCode="DUBC" collectorName="A. Vasiljeva &amp; Apo Mt. Rng" country="Philippines" county="Mindanao" elevation="1200" latitude="7.12135" location="Claveria rd." longLatPrecision="1" longitude="125.31263" municipality="Misamis" pageId="11" pageNumber="92" specimenCount="3" stateProvince="Davao del Sur">
chenko (
<collectionCode id="ED056DE05E5AFFE5FECB22A80585FF68" box="[321,402,193,216]" pageId="11" pageNumber="92">DUBC</collectionCode>
);
<specimenCount id="9D123EAC5E5AFFE5FE2C22A805C0FF68" box="[422,471,193,216]" count="1" pageId="11" pageNumber="92" type="generic">1ex.</specimenCount>
:
<collectingCountry id="F303B5B55E5AFFE5FE6E22A80677FF68" box="[484,608,193,216]" name="Philippines" pageId="11" pageNumber="92">Philippines</collectingCountry>
,
<collectingCounty id="62CA8DA95E5AFFE5FDE722A806CAFF68" box="[621,733,193,216]" pageId="11" pageNumber="92">Mindanao</collectingCounty>
,
<collectingMunicipality id="6BCF6F5F5E5AFFE5FF6922880555FF48" box="[227,322,225,248]" pageId="11" pageNumber="92">Misamis</collectingMunicipality>
or.,
<location id="8ECBA3FE5E5AFFE5FEF122880615FF48" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FEF122880615FF48" box="[379,514,225,248]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Claveria rd." pageId="11" pageNumber="92" stateProvince="Davao del Sur">Claveria rd.</location>
,
<quantity id="4CEC58C05E5AFFE5FD9E22880677FF48" box="[532,608,225,248]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="92" unit="m" value="1200.0">
<elevation id="003912165E5AFFE5FD9E22880677FF48" box="[532,608,225,248]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="92" unit="m" value="1200.0">1200m</elevation>
</quantity>
, 8.701522, 125.014624,
<date id="FFAAD3E55E5AFFE5FEF02368060EFEA8" box="[378,537,257,280]" pageId="11" pageNumber="92" value="2022-03-26" valueMax="2022-03-28" valueMin="2022-03-26">
<collectingDate id="EFEE2A0D5E5AFFE5FEF02368060EFEA8" box="[378,537,257,280]" pageId="11" pageNumber="92" value="2022-03-26" valueMax="2022-03-28" valueMin="2022-03-26">26­28.iii.2022</collectingDate>
</date>
, leg.
<collectorName id="26E190F35E5AFFE5FDD5236806F6FEA8" box="[607,737,257,280]" pageId="11" pageNumber="92">A.Vasiljeva</collectorName>
(
<collectionCode id="ED056DE05E5AFFE5FF602348052EFE88" box="[234,313,289,312]" pageId="11" pageNumber="92">DUBC</collectionCode>
);
<specimenCount id="9D123EAC5E5AFFE5FEC32348056FFE88" box="[329,376,289,312]" count="1" pageId="11" pageNumber="92" type="generic">1ex.</specimenCount>
:
<collectingCountry id="F303B5B55E5AFFE5FE0B234805EEFE88" box="[385,505,289,312]" name="Philippines" pageId="11" pageNumber="92">Philippines</collectingCountry>
,
<location id="8ECBA3FE5E5AFFE5FD8823480678FE88" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FD8823480678FE88" box="[514,623,289,312]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mindanao" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mindanao</location>
,
<collectingRegion id="49D03BC75E5AFFE5FDF223480510FEE8" country="Philippines" name="Davao del Sur" pageId="11" pageNumber="92">Davao del Sur</collectingRegion>
,
<collectorName id="26E190F35E5AFFE5FE98232805B7FEE8" box="[274,416,321,344]" pageId="11" pageNumber="92">
<location id="8ECBA3FE5E5AFFE5FE982328057BFEE8" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FE982328057BFEE8" box="[274,364,321,344]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Apo Mt." pageId="11" pageNumber="92" stateProvince="Davao del Sur">
Apo
<location id="8ECBA3FE5E5AFFE5FECF2328057BFEE8" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FECF2328057BFEE8" box="[325,364,321,344]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mt. Rng" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mt.</location>
</location>
Rng
</collectorName>
,
<location id="8ECBA3FE5E5AFFE5FE2023280602FEE8" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FE2023280602FEE8" box="[426,533,321,344]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mabanlas" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mabanlas</location>
,
<quantity id="4CEC58C05E5AFFE5FD942328067CFEE8" box="[542,619,321,344]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="92" unit="m" value="1200.0">
<elevation id="003912165E5AFFE5FD942328067CFEE8" box="[542,619,321,344]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="92" unit="m" value="1200.0">1200m</elevation>
</quantity>
, 7.120251, 125.317203,
<date id="FFAAD3E55E5AFFE5FEFD23080603FEC8" box="[375,532,353,376]" pageId="11" pageNumber="92" value="2022-03-23" valueMax="2022-03-24" valueMin="2022-03-23">
<collectingDate id="EFEE2A0D5E5AFFE5FEFD23080603FEC8" box="[375,532,353,376]" pageId="11" pageNumber="92" value="2022-03-23" valueMax="2022-03-24" valueMin="2022-03-23">23­24.iii.2022</collectingDate>
</date>
, leg.
<collectorName id="26E190F35E5AFFE5FDDD230806F6FEC8" box="[599,737,353,376]" pageId="11" pageNumber="92">A. Vasiljeva</collectorName>
(
<collectionCode id="ED056DE05E5AFFE5FF6023E8052CFE28" box="[234,315,385,408]" pageId="11" pageNumber="92">DUBC</collectionCode>
);
<specimenCount id="9D123EAC5E5AFFE5FEC423E80568FE28" box="[334,383,385,408]" count="1" pageId="11" pageNumber="92" type="generic">1ex.</specimenCount>
: the same but:
<collectingCountry id="F303B5B55E5AFFE5FDA923E80688FE28" box="[547,671,385,408]" name="Philippines" pageId="11" pageNumber="92">Philippines</collectingCountry>
,
<location id="8ECBA3FE5E5AFFE5FD2123E80531FE08" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FD2123E80531FE08" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mindanao" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mindanao</location>
,
<collectingRegion id="49D03BC75E5AFFE5FEBA23C805DEFE08" box="[304,457,417,440]" country="Philippines" name="Davao del Sur" pageId="11" pageNumber="92">Davao del Sur</collectingRegion>
,
<collectorName id="26E190F35E5AFFE5FE5F23C80671FE08" box="[469,614,417,440]" pageId="11" pageNumber="92">
<location id="8ECBA3FE5E5AFFE5FE5F23C80626FE08" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FE5F23C80626FE08" box="[469,561,417,440]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Apo Mt." pageId="11" pageNumber="92" stateProvince="Davao del Sur">
Apo
<location id="8ECBA3FE5E5AFFE5FD8023C80626FE08" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FD8023C80626FE08" box="[522,561,417,440]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mt. Rng" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mt.</location>
</location>
Rng
</collectorName>
,
<location id="8ECBA3FE5E5AFFE5FDFB23C806CBFE08" LSID="urn:lsid:plazi:treatment:03BD44335E5BFFE5FCD826D800F2FF68:8ECBA3FE5E5AFFE5FDFB23C806CBFE08" box="[625,732,417,440]" country="Philippines" county="Mindanao" latitude="7.12135" longLatPrecision="1" longitude="125.31263" municipality="Misamis" name="Mabanlas" pageId="11" pageNumber="92" stateProvince="Davao del Sur">Mabanlas</location>
,
<quantity id="4CEC58C05E5AFFE5FF6923A80523FE68" box="[227,308,449,472]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="92" unit="m" value="1500.0">
<elevation id="003912165E5AFFE5FF6923A80523FE68" box="[227,308,449,472]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="92" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<geoCoordinate id="EE2093E25E5AFFE5FEDA23A805C4FE68" box="[336,467,449,472]" degrees="7" direction="north" minutes="7. 2810" orientation="latitude" pageId="11" pageNumber="92" precision="1" value="7.12135">7 °7.2810N</geoCoordinate>
,
<geoCoordinate id="EE2093E25E5AFFE5FE7A23A8068AFE68" box="[496,669,449,472]" degrees="125" direction="east" minutes="18. 7580" orientation="longitude" pageId="11" pageNumber="92" precision="1" value="125.31263">125°18.7580E</geoCoordinate>
,
<date id="FFAAD3E55E5AFFE5FD3023A8054FFE48" pageId="11" pageNumber="92" value="2022-03-21" valueMax="2022-03-23" valueMin="2022-03-21">
<collectingDate id="EFEE2A0D5E5AFFE5FD3023A8054FFE48" pageId="11" pageNumber="92" value="2022-03-21" valueMax="2022-03-23" valueMin="2022-03-21">21­ 23.iii.2022</collectingDate>
</date>
(
<collectionCode id="ED056DE05E5AFFE5FEE3238805ADFE48" box="[361,442,481,504]" pageId="11" pageNumber="92">DUBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7CFF785E5AFFE5FE5A2388066AFDA8" collectingDate="2022-03-21" collectingDateMax="2022-03-24" collectingDateMin="2022-03-21" collectionCode="DUBC" collectorName="A. Vasiljeva" country="Philippines" pageId="11" pageNumber="92" specimenCount="1" stateProvince="Davao del Sur">
7exx.: the same but:
<date id="FFAAD3E55E5AFFE5FD372388054CFDA8" pageId="11" pageNumber="92" value="2022-03-21" valueMax="2022-03-24" valueMin="2022-03-21">
<collectingDate id="EFEE2A0D5E5AFFE5FD372388054CFDA8" pageId="11" pageNumber="92" value="2022-03-21" valueMax="2022-03-24" valueMin="2022-03-21">21­ 24.iii.2022</collectingDate>
</date>
, leg.
<collectorName id="26E190F35E5AFFE5FE1820680601FDA8" box="[402,534,513,536]" pageId="11" pageNumber="92">A. Vasiljeva</collectorName>
(
<collectionCode id="ED056DE05E5AFFE5FDAE20680662FDA8" box="[548,629,513,536]" pageId="11" pageNumber="92">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E5AFFE5FF69202905FFFD08" pageId="11" pageNumber="92" type="discussion">
<paragraph id="8BABF5255E5AFFE5FF69202905FFFD08" blockId="11.[227,737,576,696]" pageId="11" pageNumber="92">
<emphasis id="B96029375E5AFFE5FF692029054BFDE8" bold="true" box="[227,348,576,600]" pageId="11" pageNumber="92">Comment.</emphasis>
Specimens from paper of
<bibRefCitation id="EF8588D45E5AFFE5FD07202805A1FDC8" author="Anichtchenko, A. V. &amp; Minkina, L. &amp; Vasiljeva, A. &amp; Medina, M. N. D." pageId="11" pageNumber="92" pagination="99 - 111" refId="ref8853" refString="Anichtchenko, A. V., Minkina, L., Vasiljeva, A. &amp; Medina, M. N. D. 2021: A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao. A cta entomologica Musei nationalis Pragae 61 (1): 99 - 111." type="journal article" year="2021">
Anichtchenko
<emphasis id="B96029375E5AFFE5FEB320080566FDC8" box="[313,369,609,632]" italics="true" pageId="11" pageNumber="92">et al.</emphasis>
, 2021
</bibRefCitation>
identified by authors as
<taxonomicName id="4C148EA65E5AFFE5FD4020080521FD28" authorityName="Arrow" authorityYear="1905" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="92" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96029375E5AFFE5FD4020080521FD28" italics="true" pageId="11" pageNumber="92">R. gracilis</emphasis>
</taxonomicName>
indeed are
<taxonomicName id="4C148EA65E5AFFE5FE2520E80680FD28" authorityName="Ochi &amp; Kakizoe" authorityYear="2021" box="[431,663,641,664]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="92" phylum="Arthropoda" rank="species" species="paraphilippinensis">
<emphasis id="B96029375E5AFFE5FE2520E80680FD28" box="[431,663,641,664]" italics="true" pageId="11" pageNumber="92">R. paraphilippinensis</emphasis>
</taxonomicName>
. Additionally, see: discussion.
</paragraph>
</subSubSection>
<subSubSection id="C30EA6AE5E5AFFE5FC8C22A900F2FF68" box="[774,1253,192,216]" pageId="11" pageNumber="92" type="distribution">
<paragraph id="8BABF5255E5AFFE5FC8C22A900F2FF68" blockId="11.[774,1253,192,216]" box="[774,1253,192,216]" pageId="11" pageNumber="92">
<emphasis id="B96029375E5AFFE5FC8C22A9078CFF68" bold="true" box="[774,923,192,216]" pageId="11" pageNumber="92">Distribution.</emphasis>
<collectingCountry id="F303B5B55E5AFFE5FC2B22A80008FF68" box="[929,1055,193,216]" name="Philippines" pageId="11" pageNumber="92">Philippines</collectingCountry>
: Mindanao Island.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,267 @@
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<mods:title id="93FD9CDBE830E8AC98168221317FEB49">Description of two new Neocollyris W. Horn, 1901 species from Vietnam (Coleoptera: Cicindelidae)</mods:title>
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<mods:namePart id="8C89188B1864C91828E9872E365D0C65">Wiesner, Jürgen</mods:namePart>
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<mods:date id="806FE3C4F20597C7AB892C7F00AC9000">2022</mods:date>
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<treatment id="03C02E688964FFAB12270CA0B0CD986E" LSID="urn:lsid:plazi:treatment:03C02E688964FFAB12270CA0B0CD986E" httpUri="http://treatment.plazi.org/id/03C02E688964FFAB12270CA0B0CD986E" lastPageId="5" lastPageNumber="72" pageId="3" pageNumber="70">
<subSubSection id="C373CCF58964FFAD12270CA0B5249D90" box="[774,1133,448,473]" pageId="3" pageNumber="70" type="nomenclature">
<paragraph id="8BD69F7E8964FFAD12270CA0B5249D90" blockId="3.[774,1133,448,473]" box="[774,1133,448,473]" pageId="3" pageNumber="70">
<heading id="D09E28128964FFAD12270CA0B5249D90" box="[774,1133,448,473]" fontSize="10" level="2" pageId="3" pageNumber="70" reason="7">
<taxonomicName id="4C69E4FD8964FFAD12270CA0B5629D91" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[774,1067,449,473]" class="Insecta" family="Carabidae" genus="Neocollyris" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. nov.">
<emphasis id="B91D436C8964FFAD12270CA0B2CD9D91" bold="true" box="[774,900,449,473]" italics="true" pageId="3" pageNumber="70">Neocollyris</emphasis>
(s. str.) hirsuta
</taxonomicName>
<emphasis id="B91D436C8964FFAD15100CA1B5249D90" bold="true" box="[1073,1133,448,472]" pageId="3" pageNumber="70">
<taxonomicNameLabel id="A22EFE178964FFAD15100CA1B5249D90" box="[1073,1133,448,472]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58964FFAD12270F61B2239E10" pageId="3" pageNumber="70" type="materials_examined">
<paragraph id="8BD69F7E8964FFAD12270F61B2239E10" blockId="3.[774,1285,512,600]" pageId="3" pageNumber="70">
<emphasis id="B91D436C8964FFAD12270F61B2F19E50" bold="true" box="[774,952,512,536]" pageId="3" pageNumber="70">Type material:</emphasis>
<materialsCitation id="3B0195238964FFAD12E20F60B2239E10" collectingDate="2018-04" collectionCode="DUBC" country="Vietnam" location="So'n Tra Mt." municipality="Nang Pro" pageId="3" pageNumber="70" specimenCount="1" stateProvince="Da Nang" typeStatus="holotype">
<typeStatus id="54D221DC8964FFAD12E20F60B5639E50" box="[963,1066,513,536]" pageId="3" pageNumber="70" type="holotype">Holotype</typeStatus>
, male
<collectingCountry id="F37EDFEE8964FFAD15810F60B5B49E50" box="[1184,1277,513,536]" name="Vietnam" pageId="3" pageNumber="70">Vietnam</collectingCountry>
,
<collectingMunicipality id="6BB205048964FFAD12270F40B23A9E70" box="[774,883,545,568]" pageId="3" pageNumber="70">Nang Pro</collectingMunicipality>
,
<collectingRegion id="49AD519C8964FFAD12A00F40B2AE9E70" box="[897,999,545,568]" country="Vietnam" name="Da Nang" pageId="3" pageNumber="70">Da Nang</collectingRegion>
, |
<location id="8EB6C9A58964FFAD15250F40B5DC9E70" LSID="urn:lsid:plazi:treatment:03C02E688964FFAB12270CA0B0CD986E:8EB6C9A58964FFAD15250F40B5DC9E70" box="[1028,1173,545,568]" country="Vietnam" municipality="Nang Pro" name="So'n Tra Mt." pageId="3" pageNumber="70" stateProvince="Da Nang">Son Trà Mt.</location>
,
<date id="FFD7B9BE8964FFAD15820F40B5B29E70" box="[1187,1275,545,568]" pageId="3" pageNumber="70" value="2018-04">
<collectingDate id="EF9340568964FFAD15820F40B5B29E70" box="[1187,1275,545,568]" pageId="3" pageNumber="70" value="2018-04">IV.2018</collectingDate>
</date>
” (
<collectionCode id="ED7807BB8964FFAD122F0F20B2169E10" box="[782,863,577,600]" pageId="3" pageNumber="70">DUBC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58964FFAD12270FE1B57F9FB0" pageId="3" pageNumber="70" type="diagnosis">
<paragraph id="8BD69F7E8964FFAD12270FE1B57F9FB0" blockId="3.[774,1285,640,1016]" pageId="3" pageNumber="70">
<emphasis id="B91D436C8964FFAD12270FE1B2359ED0" bold="true" box="[774,892,640,664]" pageId="3" pageNumber="70">Diagnosis.</emphasis>
This new species is similar in body size, color, and dense pubescence to the newly described
<taxonomicName id="4C69E4FD8964FFAD12550FA0B5619E90" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[884,1064,705,728]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="trichocephala" status="sp. n.">
<emphasis id="B91D436C8964FFAD12550FA0B5619E90" box="[884,1064,705,728]" italics="true" pageId="3" pageNumber="70">N. trichocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD150C0FA0B52E9E90" box="[1069,1127,705,728]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
, but
<taxonomicName id="4C69E4FD8964FFAD15BB0FA0B44D9E90" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[1178,1284,705,728]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. n.">
<emphasis id="B91D436C8964FFAD15BB0FA0B44D9E90" box="[1178,1284,705,728]" italics="true" pageId="3" pageNumber="70">N. hirsuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD12270F80B2089EB0" box="[774,833,737,760]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
can be differentiated by its smooth head. Elytra with dark brown, barely visible but evident sub­ humeral and transversal elytral pattern, while in
<taxonomicName id="4C69E4FD8964FFAD12450E20B5509F10" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[868,1049,833,856]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="trichocephala" status="sp. n.">
<emphasis id="B91D436C8964FFAD12450E20B5509F10" box="[868,1049,833,856]" italics="true" pageId="3" pageNumber="70">N. trichocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD15010E20B5139F10" box="[1056,1114,833,856]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
, elytra lack this pattern. Labrum of
<taxonomicName id="4C69E4FD8964FFAD12F90E00B5C79F30" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[984,1166,865,888]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="trichocephala" status="sp. n.">
<emphasis id="B91D436C8964FFAD12F90E00B5C79F30" box="[984,1166,865,888]" italics="true" pageId="3" pageNumber="70">N. trichocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD15B50E00B5879F30" box="[1172,1230,865,888]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
with thin microsculpture, while in
<taxonomicName id="4C69E4FD8964FFAD15650EE0B5E69FD0" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[1092,1199,897,920]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. n.">
<emphasis id="B91D436C8964FFAD15650EE0B5E69FD0" box="[1092,1199,897,920]" italics="true" pageId="3" pageNumber="70">N. hirsuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD15970EE0B5B99FD0" box="[1206,1264,897,920]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
it is smooth. Aedeagus of
<taxonomicName id="4C69E4FD8964FFAD12DE0EC0B5219FF0" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[1023,1128,929,952]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. n.">
<emphasis id="B91D436C8964FFAD12DE0EC0B5219FF0" box="[1023,1128,929,952]" italics="true" pageId="3" pageNumber="70">N. hirsuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD154C0EC0B5EF9FF0" box="[1133,1190,929,952]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
, is larger than in
<taxonomicName id="4C69E4FD8964FFAD127F0EA0B55F9F90" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[862,1046,961,984]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="trichocephala" status="sp. n.">
<emphasis id="B91D436C8964FFAD127F0EA0B55F9F90" box="[862,1046,961,984]" italics="true" pageId="3" pageNumber="70">N. trichocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD15010EA0B5149F90" box="[1056,1117,961,984]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
and with apex longer and less downturned.
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58964FFAB12270941B3929F0A" lastPageId="5" lastPageNumber="72" pageId="3" pageNumber="70" type="description">
<paragraph id="8BD69F7E8964FFAD12270941B44D9870" blockId="3.[774,1284,1056,1112]" box="[774,1284,1056,1080]" pageId="3" pageNumber="70">
<emphasis id="B91D436C8964FFAD12270941B2DA9870" bold="true" box="[774,915,1056,1080]" pageId="3" pageNumber="70">Description.</emphasis>
Body length
<quantity id="4C91329B8964FFAD15050940B5CD9870" box="[1060,1156,1057,1080]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="3" pageNumber="70" unit="mm" value="12.8">12.8 mm</quantity>
in
<typeStatus id="54D221DC8964FFAD15870940B44D9870" box="[1190,1284,1057,1080]" pageId="3" pageNumber="70" type="holotype">holotype</typeStatus>
</paragraph>
<paragraph id="8BD69F7E8964FFAD12270920B2179810" blockId="3.[774,1284,1056,1112]" box="[774,862,1089,1112]" pageId="3" pageNumber="70">
(
<figureCitation id="135283FB8964FFAD122F0920B2199810" box="[782,848,1089,1112]" captionStart="Figs" captionStartId="1.[226,276,1698,1721]" captionTargetBox="[228,1284,986,1663]" captionTargetId="figure-337@1.[226,840,984,1663]" captionTargetPageId="1" captionText="Figs. 13. Habitus of Neocollyris (s. str.) spp.: 1 N. trichocephalus sp. n., holotype; 2 idem, paratype. 3 N. hirsutus sp. n., Holotype. Scale line = 1 sm." pageId="3" pageNumber="70">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DF16CFF68964FFAD11C30BE5B39D9A93" pageId="3" pageNumber="70" startId="3.[226,276,1668,1691]" targetBox="[352,1157,1144,1642]" targetPageId="3" targetType="figure">
<paragraph id="8BD69F7E8964FFAD11C30BE5B39D9A93" blockId="3.[226,1285,1668,1755]" pageId="3" pageNumber="70">
Figs. 912. Palpi of
<taxonomicName id="4C69E4FD8964FFAD109C0BE5B3709AD3" authorityName="W. Horn" authorityYear="1901" box="[445,569,1668,1691]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B91D436C8964FFAD109C0BE5B3709AD3" box="[445,569,1668,1691]" italics="true" pageId="3" pageNumber="70">Neocollyris</emphasis>
</taxonomicName>
(s. str.) spp. 9 maxillary palpus of
<taxonomicName id="4C69E4FD8964FFAD12E60BE5B5CF9AD3" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[967,1158,1668,1691]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="trichocephalus" status="sp. n.">
<emphasis id="B91D436C8964FFAD12E60BE5B5CF9AD3" box="[967,1158,1668,1691]" italics="true" pageId="3" pageNumber="70">N. trichocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD15AC0BE5B5809AD3" box="[1165,1225,1668,1691]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
; 10 labial palpus of
<taxonomicName id="4C69E4FD8964FFAD10B40BC5B31E9AF3" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[405,599,1700,1723]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="trichocephalus" status="sp. n.">
<emphasis id="B91D436C8964FFAD10B40BC5B31E9AF3" box="[405,599,1700,1723]" italics="true" pageId="3" pageNumber="70">N. trichocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD13410BC5B3D49AF3" box="[608,669,1700,1723]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
; 11 maxillary palpus of
<taxonomicName id="4C69E4FD8964FFAD12E30BC5B5739AF3" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[962,1082,1700,1723]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. n.">
<emphasis id="B91D436C8964FFAD12E30BC5B5739AF3" box="[962,1082,1700,1723]" italics="true" pageId="3" pageNumber="70">N. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD15620BC5B5C89AF3" box="[1091,1153,1700,1723]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
; 12 labial palpus of
<taxonomicName id="4C69E4FD8964FFAD106C0BA5B08A9A93" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[333,451,1732,1755]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. n.">
<emphasis id="B91D436C8964FFAD106C0BA5B08A9A93" box="[333,451,1732,1755]" italics="true" pageId="3" pageNumber="70">N. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178964FFAD10E80BA5B34A9A93" box="[457,515,1732,1755]" pageId="3" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
Scale line = 1 mm.
</paragraph>
</caption>
<caption id="DF16CFF68963FFAA100F0F7AB2609E3A" pageId="4" pageNumber="71" startId="4.[302,348,539,562]" targetBox="[304,810,192,509]" targetPageId="4" targetType="figure">
<paragraph id="8BD69F7E8963FFAA100F0F7AB2609E3A" blockId="4.[302,812,539,626]" pageId="4" pageNumber="71">
Figs 1314. Head in frontal view and labrum of
<taxonomicName id="4C69E4FD8963FFAA100F0F5AB0E39E1A" authorityName="W. Horn" authorityYear="1901" box="[302,426,571,594]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91D436C8963FFAA100F0F5AB0E39E1A" box="[302,426,571,594]" italics="true" pageId="4" pageNumber="71">Neocollyris</emphasis>
</taxonomicName>
(s.str.) spp. 13
<taxonomicName id="4C69E4FD8963FFAA13550F5AB2659E1A" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[628,812,571,594]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="trichocephala" status="sp. n.">
<emphasis id="B91D436C8963FFAA13550F5AB2659E1A" box="[628,812,571,594]" italics="true" pageId="4" pageNumber="71">N. trichocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178963FFAA100F0F3AB0239E3A" box="[302,362,603,626]" pageId="4" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
; 14
<taxonomicName id="4C69E4FD8963FFAA108D0F3AB35E9E3A" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[428,535,603,626]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. n.">
<emphasis id="B91D436C8963FFAA108D0F3AB35E9E3A" box="[428,535,603,626]" italics="true" pageId="4" pageNumber="71">N. hirsuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178963FFAA133F0F3AB3119E3A" box="[542,600,603,626]" pageId="4" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
Scale line = 1 mm.
</paragraph>
</caption>
<caption id="DF16CFF68963FFAA100F087AB0B199DA" pageId="4" pageNumber="71" startId="4.[302,352,1307,1330]" targetBox="[305,810,669,1273]" targetPageId="4" targetType="figure">
<paragraph id="8BD69F7E8963FFAA100F087AB0B199DA" blockId="4.[302,813,1307,1426]" pageId="4" pageNumber="71">
Figs. 1517. Aedeagus of
<taxonomicName id="4C69E4FD8963FFAA1377087AB39A997A" authorityName="W. Horn" authorityYear="1901" box="[598,723,1307,1330]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91D436C8963FFAA1377087AB39A997A" box="[598,723,1307,1330]" italics="true" pageId="4" pageNumber="71">Neocollyris</emphasis>
</taxonomicName>
(s. str.) spp.: 1516
<taxonomicName id="4C69E4FD8963FFAA10E7085AB332991A" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[454,635,1339,1362]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="trichocephala" status="sp. n.">
<emphasis id="B91D436C8963FFAA10E7085AB332991A" box="[454,635,1339,1362]" italics="true" pageId="4" pageNumber="71">N. trichocephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178963FFAA13A0085AB3F5991A" box="[641,700,1339,1362]" pageId="4" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
, Holotype and paratype; 17
<taxonomicName id="4C69E4FD8963FFAA1323083AB326993A" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[514,623,1371,1394]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. n.">
<emphasis id="B91D436C8963FFAA1323083AB326993A" box="[514,623,1371,1394]" italics="true" pageId="4" pageNumber="71">N. hirsuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178963FFAA1356083AB3FD993A" box="[631,692,1371,1394]" pageId="4" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
, Holotype. Scale line = 1 mm.
</paragraph>
</caption>
<paragraph id="8BD69F7E8963FFAA100F08A0B02699B0" blockId="4.[302,813,1473,1528]" pageId="4" pageNumber="71">Body color uniformly violet with strong purple luster.</paragraph>
<paragraph id="8BD69F7E8963FFAA100F0B40B4189CB1" blockId="4.[302,813,1569,1752]" lastBlockId="4.[850,1361,193,932]" pageId="4" pageNumber="71">
Head (
<figureCitation id="135283FB8963FFAA10560B40B0979A70" box="[375,478,1569,1592]" captionStart="Figs" captionStartId="2.[302,352,1130,1153]" captionTargetBox="[444,1228,194,1096]" captionTargetId="figure-342@2.[440,1231,189,1098]" captionTargetPageId="2" captionText="Figs. 48. Neocollyris (s. str.) spp.: 45 head, pronotum, and basal part of elytra, right lateral view; 68 head, dorsal view. 4, 6, 7 N. trichocephalus sp. n.; 5, 8 N. hirsutus sp. n., Holotype. Scale line = 1 mm." pageId="4" pageNumber="71">Figs. 5, 8</figureCitation>
) oblong, HL/HW = 0.82, with thin microsculpture consisting of polygonal meshes, temples protruding and evenly rounded towards base; clypeus with two setae; frons narrow, frontal grooves shallow, almost straight, subparallel in anterior third, slightly divergent in posterior third; eyes moderately protruding; each supra­orbital plate with 2 long setae. Labrum
</paragraph>
<paragraph id="8BD69F7E8963FFAA12730C63B4189E16" blockId="4.[850,1361,193,932]" pageId="4" pageNumber="71">
(
<figureCitation id="135283FB8963FFAA127B0C63B2E09D51" box="[858,937,258,281]" captionStart="Figs 1314" captionStartId="4.[302,348,539,562]" captionTargetBox="[304,810,192,509]" captionTargetId="figure-590@4.[302,812,189,509]" captionTargetPageId="4" captionText="Figs 1314. Head in frontal view and labrum of Neocollyris (s.str.) spp. 13 N. trichocephala sp. n.; 14 N. hirsuta sp. n. Scale line = 1 mm." pageId="4" pageNumber="71">Fig. 14</figureCitation>
) relatively short and transverse; unicolored, violet; with microsculpture, consisting of polygonal meshes on the center, and transverse meshes on the sides; with eight long sub marginal setae and seven apical teeth: with three short and wide frontal central teeth with rounded apices, as well as a pair of slightly longer and narrower latero­apical teeth and longer and pointed latero­basal teeth on both sides. Mandibles black with violet luster on the base. Palpi entirely black, maxillary with the shape of isosceles triangle
</paragraph>
<paragraph id="8BD69F7E8963FFAA12730F09B4189E37" blockId="4.[850,1361,193,932]" box="[850,1361,616,639]" pageId="4" pageNumber="71">
(
<figureCitation id="135283FB8963FFAA127B0F09B2EB9E37" box="[858,930,616,639]" captionStart="Figs" captionStartId="3.[226,276,1668,1691]" captionTargetBox="[352,1157,1144,1642]" captionTargetId="figure-434@3.[348,1157,1142,1645]" captionTargetPageId="3" captionText="Figs. 912. Palpi of Neocollyris (s. str.) spp. 9 maxillary palpus of N. trichocephalus sp. n.; 10 labial palpus of N. trichocephalus sp. n.; 11 maxillary palpus of N. hirsutus sp. n.; 12 labial palpus of N. hirsutus sp. n. Scale line = 1 mm." pageId="4" pageNumber="71">Fig. 11</figureCitation>
) and labial with the shape of right triangle
</paragraph>
<paragraph id="8BD69F7E8963FFAA12730FE8B5F39FEC" blockId="4.[850,1361,193,932]" pageId="4" pageNumber="71">
(
<figureCitation id="135283FB8963FFAA127B0FE8B2EE9EE8" box="[858,935,649,672]" captionStart="Figs" captionStartId="3.[226,276,1668,1691]" captionTargetBox="[352,1157,1144,1642]" captionTargetId="figure-434@3.[348,1157,1142,1645]" captionTargetPageId="3" captionText="Figs. 912. Palpi of Neocollyris (s. str.) spp. 9 maxillary palpus of N. trichocephalus sp. n.; 10 labial palpus of N. trichocephalus sp. n.; 11 maxillary palpus of N. hirsutus sp. n.; 12 labial palpus of N. hirsutus sp. n. Scale line = 1 mm." pageId="4" pageNumber="71">Fig. 12</figureCitation>
). Antennae relatively long; antennomere 3 the longest; scape and pedicel dark violet, antennomeres 34 violet, with yellow apices; antennomere 5 yellow, with violet basal third; antennomeres 611 brown. Scape with a single apical seta, antennomeres 34 glabrous except for 6 very short setae on ventral ridge, apical half of antennomere 5 and antennomeres 611 densely pubescent with very short yellow setae.
</paragraph>
<paragraph id="8BD69F7E8963FFAA12730EAFB5FD9903" blockId="4.[850,1361,974,1355]" pageId="4" pageNumber="71">
Pronotum (
<figureCitation id="135283FB8963FFAA12E20EAFB54A9FAD" box="[963,1027,974,997]" captionStart="Figs" captionStartId="1.[226,276,1698,1721]" captionTargetBox="[228,1284,986,1663]" captionTargetId="figure-337@1.[226,840,984,1663]" captionTargetPageId="1" captionText="Figs. 13. Habitus of Neocollyris (s. str.) spp.: 1 N. trichocephalus sp. n., holotype; 2 idem, paratype. 3 N. hirsutus sp. n., Holotype. Scale line = 1 sm." pageId="4" pageNumber="71">Figs 3</figureCitation>
,
<figureCitation id="135283FB8963FFAA152C0EAFB5539FAD" box="[1037,1050,974,997]" captionStart="Figs" captionStartId="2.[302,352,1130,1153]" captionTargetBox="[444,1228,194,1096]" captionTargetId="figure-342@2.[440,1231,189,1098]" captionTargetPageId="2" captionText="Figs. 48. Neocollyris (s. str.) spp.: 45 head, pronotum, and basal part of elytra, right lateral view; 68 head, dorsal view. 4, 6, 7 N. trichocephalus sp. n.; 5, 8 N. hirsutus sp. n., Holotype. Scale line = 1 mm." pageId="4" pageNumber="71">5</figureCitation>
) with a shallow collar, without anterior hump, moderately long, LP/WP = 1.83, distinctly expanded in basal third; disc almost flat in lateral view, with shallow, transverse wrinkles in anterior half, relatively densely and unevenly punctured by setiferous pores, bearing very long white hairs; punctures on base of pronotum twice as large and denser as on disc; anterior sulcus relatively wide and shallow, while posterior one more narrow and deeper. Prosternum, pro­episternum, mesothorax and mesosternum densely pubescent with long, white hairs.
</paragraph>
<paragraph id="8BD69F7E8963FFAA12730814B4199A99" blockId="4.[850,1361,1397,1745]" pageId="4" pageNumber="71">
Elytra (
<figureCitation id="135283FB8963FFAA12830814B2A899C4" box="[930,993,1397,1420]" captionStart="Figs" captionStartId="1.[226,276,1698,1721]" captionTargetBox="[228,1284,986,1663]" captionTargetId="figure-337@1.[226,840,984,1663]" captionTargetPageId="1" captionText="Figs. 13. Habitus of Neocollyris (s. str.) spp.: 1 N. trichocephalus sp. n., holotype; 2 idem, paratype. 3 N. hirsutus sp. n., Holotype. Scale line = 1 sm." pageId="4" pageNumber="71">Fig. 3</figureCitation>
) moderately long, EL/EW = 3.33, parallel sided, metallic violet with light purple luster; elytral maculation dark brown, barely visible, consisting of sub­ humeral elongate spot, and transversal middle spot; covered with isodiametric microsculpture; disc with numerous regularly distributed, rounded, blue pits, slightly elongate and smaller in apical fifth; almost all pits with very small setiferous pore on anterior margin, bearing extremely long white hairs; suture moderately protruding. Sutural tooth on apices
</paragraph>
<paragraph id="8BD69F7E8962FFAB13560DF0B2389CEC" blockId="5.[631,881,145,164]" box="[631,881,145,164]" pageId="5" pageNumber="72">
<emphasis id="B91D436C8962FFAB13560DF0B2389CEC" box="[631,881,145,164]" italics="true" pageId="5" pageNumber="72">Anichtchenko A., Wiesner J.</emphasis>
</paragraph>
<paragraph id="8BD69F7E8962FFAB11C30DA0B3A89CB1" blockId="5.[226,737,193,249]" pageId="5" pageNumber="72">of elytra obtuse, not prominent, exterior angle of elytra obtuse and angulate. Scutellum dark blue.</paragraph>
<paragraph id="8BD69F7E8962FFAB11C30C42B3AB9D33" blockId="5.[226,738,291,379]" pageId="5" pageNumber="72">Abdominal sternites black with violet luster, with few thin and long pale setae on each side. All coxae, as well as fore­ and mid­trochanters black.</paragraph>
<paragraph id="8BD69F7E8962FFAB11C30CC4B3099E16" blockId="5.[226,737,421,606]" pageId="5" pageNumber="72">
Profemora dark violet, ventrally dark brown with violet reflections. Meso­ and metafemora dark brown with violet reflections, dorsally and apically violet. All tibiae dark violet. Pro­ and mesotarsi uniformly violet; metatarsi light yellow, with two apical tarsomeres violet (
<figureCitation id="135283FB8962FFAB10D00F26B37B9E16" box="[497,562,583,606]" captionStart="Figs" captionStartId="1.[226,276,1698,1721]" captionTargetBox="[228,1284,986,1663]" captionTargetId="figure-337@1.[226,840,984,1663]" captionTargetPageId="1" captionText="Figs. 13. Habitus of Neocollyris (s. str.) spp.: 1 N. trichocephalus sp. n., holotype; 2 idem, paratype. 3 N. hirsutus sp. n., Holotype. Scale line = 1 sm." pageId="5" pageNumber="72">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BD69F7E8962FFAB11C30FE8B3929F0A" blockId="5.[226,737,649,834]" pageId="5" pageNumber="72">
Aedeagus (
<figureCitation id="135283FB8962FFAB10760FE8B0E89EE8" box="[343,417,649,672]" captionStart="Figs" captionStartId="4.[302,352,1307,1330]" captionTargetBox="[305,810,669,1273]" captionTargetId="figure-592@4.[302,812,666,1274]" captionTargetPageId="4" captionText="Figs. 1517. Aedeagus of Neocollyris (s. str.) spp.: 1516 N. trichocephala sp. n., Holotype and paratype; 17 N. hirsuta sp. n., Holotype. Scale line = 1 mm." pageId="5" pageNumber="72">Fig. 17</figureCitation>
) relatively long, 2.0
<quantity id="4C91329B8962FFAB135F0FE8B3949EE8" box="[638,733,649,672]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="5" pageNumber="72" unit="mm" value="2.15">2.15 mm</quantity>
; with a wide, gradually curved basal bulb; ventral side of median lobe straight; apical part relatively long, strongly downturned, apex narrow and acute; internal sack with short and thick flagellum, and with thin undulate sclerite at base of flagellum.
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58962FFAB11C30E0DB3F99FCB" box="[226,688,876,900]" pageId="5" pageNumber="72" type="distribution">
<paragraph id="8BD69F7E8962FFAB11C30E0DB3F99FCB" blockId="5.[226,688,876,900]" box="[226,688,876,900]" pageId="5" pageNumber="72">
<emphasis id="B91D436C8962FFAB11C30E0DB03E9FCC" bold="true" box="[226,375,876,900]" pageId="5" pageNumber="72">Distribution.</emphasis>
<collectingCountry id="F37EDFEE8962FFAB105D0E0DB0929FCB" box="[380,475,876,899]" name="Vietnam" pageId="5" pageNumber="72">Vietnam</collectingCountry>
: province
<collectingRegion id="49AD519C8962FFAB13680E0DB3E59FCB" box="[585,684,876,899]" country="Vietnam" name="Da Nang" pageId="5" pageNumber="72">Da Nang</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58962FFAB11C30ECCB0CD986E" pageId="5" pageNumber="72" type="etymology">
<paragraph id="8BD69F7E8962FFAB11C30ECCB0CD986E" blockId="5.[226,737,941,1062]" pageId="5" pageNumber="72">
<emphasis id="B91D436C8962FFAB11C30ECCB02E9F8D" bold="true" box="[226,359,941,965]" pageId="5" pageNumber="72">Etymology.</emphasis>
The name of the new species is derived from the Latin
<taxonomicName id="4C69E4FD8962FFAB10F50EAFB3629FAD" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[468,555,974,997]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="hirsuta">“hirsuta</taxonomicName>
roughly hairy, because of unusually long pubescence of pronotum and elytra.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="8BD69F7E8966FFAF11C30FEAB3DD9EEB" blockId="1.[226,660,651,675]" box="[226,660,651,675]" pageId="1" pageNumber="68">
<heading id="D09E28128966FFAF11C30FEAB3DD9EEB" box="[226,660,651,675]" fontSize="10" level="2" pageId="1" pageNumber="68" reason="7">
<taxonomicName id="4C69E4FD8966FFAF11C30FEAB3189EEB" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[226,593,651,675]" class="Insecta" family="Carabidae" genus="Neocollyris" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="trichocephala" status="sp. nov.">
<emphasis id="B91D436C8966FFAF11C30FEAB0299EEB" bold="true" box="[226,352,651,675]" italics="true" pageId="1" pageNumber="68">Neocollyris</emphasis>
(s. str.) trichocephala
</taxonomicName>
<emphasis id="B91D436C8966FFAF13790FEAB3DD9EEB" bold="true" box="[600,660,651,675]" pageId="1" pageNumber="68">
<taxonomicNameLabel id="A22EFE178966FFAF13790FEAB3DD9EEB" box="[600,660,651,675]" pageId="1" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58966FFAF11C30FAAB0299F0B" pageId="1" pageNumber="68" type="materials_examined">
<paragraph id="8BD69F7E8966FFAF11C30FAAB0299F0B" blockId="1.[226,737,715,835]" pageId="1" pageNumber="68">
<emphasis id="B91D436C8966FFAF11C30FAAB0DD9EAB" bold="true" box="[226,404,715,739]" pageId="1" pageNumber="68">Type material:</emphasis>
<materialsCitation id="3B0195238966FFAF10BE0FADB00A9F6B" collectingDate="2018-04" collectionCode="DUBC" country="Vietnam" location="So'n Tra Mt." municipality="Nang Pro" pageId="1" pageNumber="68" specimenCount="1" stateProvince="Da Nang" typeStatus="holotype">
<typeStatus id="54D221DC8966FFAF10BE0FADB34F9EAB" box="[415,518,716,739]" pageId="1" pageNumber="68" type="holotype">Holotype</typeStatus>
, male
<collectingCountry id="F37EDFEE8966FFAF135D0FADB3909EAB" box="[636,729,716,739]" name="Vietnam" pageId="1" pageNumber="68">Vietnam</collectingCountry>
,
<collectingMunicipality id="6BB205048966FFAF11C30F8DB0069F4B" box="[226,335,748,771]" pageId="1" pageNumber="68">Nang Pro</collectingMunicipality>
,
<collectingRegion id="49AD519C8966FFAF107C0F8DB08A9F4B" box="[349,451,748,771]" country="Vietnam" name="Da Nang" pageId="1" pageNumber="68">Da Nang</collectingRegion>
, |
<location id="8EB6C9A58966FFAF10C00F8DB3389F4B" LSID="urn:lsid:plazi:treatment:03C02E688966FFAD11C30FEAB5479D30:8EB6C9A58966FFAF10C00F8DB3389F4B" box="[481,625,748,771]" country="Vietnam" municipality="Nang Pro" name="So'n Tra Mt." pageId="1" pageNumber="68" stateProvince="Da Nang">Son Trà Mt.</location>
,
<date id="FFD7B9BE8966FFAF135E0F8DB39F9F4B" box="[639,726,748,771]" pageId="1" pageNumber="68" value="2018-04">
<collectingDate id="EF9340568966FFAF135E0F8DB39F9F4B" box="[639,726,748,771]" pageId="1" pageNumber="68" value="2018-04">IV.2018</collectingDate>
</date>
” (
<collectionCode id="ED7807BB8966FFAF11CB0E6DB0729F6B" box="[234,315,780,803]" pageId="1" pageNumber="68">DUBC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54D221DC8966FFAF106C0E6DB0E79F6B" box="[333,430,780,803]" pageId="1" pageNumber="68" type="paratype">Paratype</typeStatus>
,
<specimenCount id="9D6F54F78966FFAF10980E6DB34D9F6B" box="[441,516,780,803]" count="1" pageId="1" pageNumber="68" type="male">1 male</specimenCount>
, with the same label data (cJW).
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58966FFAF11C30E0AB2949CB0" pageId="1" pageNumber="68" type="diagnosis">
<paragraph id="8BD69F7E8966FFAF11C30E0AB2949CB0" blockId="1.[226,737,875,931]" lastBlockId="1.[774,1284,193,248]" pageId="1" pageNumber="68">
<emphasis id="B91D436C8966FFAF11C30E0AB0119FCB" bold="true" box="[226,344,875,899]" pageId="1" pageNumber="68">Diagnosis.</emphasis>
This new species can be easily differentiated from all known species of the genus by its head, with 2525 large setiferous pores bearing long hairs (
<figureCitation id="135283FB8966FFAF12A10D80B2869CB0" box="[896,975,225,248]" captionStart="Figs 1314" captionStartId="4.[302,348,539,562]" captionTargetBox="[304,810,192,509]" captionTargetId="figure-590@4.[302,812,189,509]" captionTargetPageId="4" captionText="Figs 1314. Head in frontal view and labrum of Neocollyris (s.str.) spp. 13 N. trichocephala sp. n.; 14 N. hirsuta sp. n. Scale line = 1 mm." pageId="1" pageNumber="68">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58966FFAD12270C40B0199810" lastPageId="3" lastPageNumber="70" pageId="1" pageNumber="68" type="description">
<paragraph id="8BD69F7E8966FFAF12270C40B5059D11" blockId="1.[774,1285,289,345]" pageId="1" pageNumber="68">
<emphasis id="B91D436C8966FFAF12270C40B2DA9D71" bold="true" box="[774,915,289,313]" pageId="1" pageNumber="68">Description.</emphasis>
Body length
<quantity id="4C91329B8966FFAF15000C43B5C99D71" box="[1057,1152,290,313]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="1" pageNumber="68" unit="mm" value="10.7">10.7 mm</quantity>
in
<typeStatus id="54D221DC8966FFAF15810C43B4489D71" box="[1184,1281,290,313]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
,
<quantity id="4C91329B8966FFAF12270C23B2189D11" box="[774,849,322,345]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="68" unit="mm" value="11.0">11 mm</quantity>
in
<typeStatus id="54D221DC8966FFAF12520C23B2999D11" box="[883,976,322,345]" pageId="1" pageNumber="68" type="paratype">paratype</typeStatus>
(
<figureCitation id="135283FB8966FFAF12FF0C23B5769D11" box="[990,1087,322,345]" captionStart="Figs" captionStartId="1.[226,276,1698,1721]" captionTargetBox="[228,1284,986,1663]" captionTargetId="figure-337@1.[226,840,984,1663]" captionTargetPageId="1" captionText="Figs. 13. Habitus of Neocollyris (s. str.) spp.: 1 N. trichocephalus sp. n., holotype; 2 idem, paratype. 3 N. hirsutus sp. n., Holotype. Scale line = 1 sm." pageId="1" pageNumber="68">Figs 1, 2</figureCitation>
).
</paragraph>
<paragraph id="8BD69F7E8966FFAF12270CE2B5209DF2" blockId="1.[774,1285,387,442]" pageId="1" pageNumber="68">Body color uniformly violet with weak purple luster on the head and pronotum.</paragraph>
<paragraph id="8BD69F7E8966FFAC12270C82B264993C" blockId="1.[774,1285,483,926]" lastBlockId="2.[302,813,1277,1748]" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68">
Head (
<figureCitation id="135283FB8966FFAF126A0C82B28D9DB2" box="[843,964,483,506]" captionStart="Figs" captionStartId="2.[302,352,1130,1153]" captionTargetBox="[444,1228,194,1096]" captionTargetId="figure-342@2.[440,1231,189,1098]" captionTargetPageId="2" captionText="Figs. 48. Neocollyris (s. str.) spp.: 45 head, pronotum, and basal part of elytra, right lateral view; 68 head, dorsal view. 4, 6, 7 N. trichocephalus sp. n.; 5, 8 N. hirsutus sp. n., Holotype. Scale line = 1 mm." pageId="1" pageNumber="68">Figs. 4, 6, 7</figureCitation>
) oblong, HL/HW = 0.870.89, with thin microsculpture consisting of polygonal meshes, temples protruding and evenly rounded; clypeus with two setae; frons narrow, frontal grooves shallow, slightly divergent from clypeus towards vertex; eyes moderately protruding; each supra­orbital plate with 510 long setae; vertex and occiput with 2526 large setiferous pores, bearing long setae. Labrum (
<figureCitation id="135283FB8966FFAF15600F84B5C69EB4" box="[1089,1167,741,764]" captionStart="Figs 1314" captionStartId="4.[302,348,539,562]" captionTargetBox="[304,810,192,509]" captionTargetId="figure-590@4.[302,812,189,509]" captionTargetPageId="4" captionText="Figs 1314. Head in frontal view and labrum of Neocollyris (s.str.) spp. 13 N. trichocephala sp. n.; 14 N. hirsuta sp. n. Scale line = 1 mm." pageId="1" pageNumber="68">Fig. 13</figureCitation>
) short and transverse; unicolored, violet; smooth, without micro reticulation, with eight long sub marginal setae and seven apical teeth: with three short and wide central teeth, with obtuse apices, as well as a pair of slightly longer and narrow latero­apical teeth and relatively smaller latero­basal teeth with pointed apex on both sides. Mandibles black with violet luster on the base. Palpi entirely black, maxillary with the shape of isosceles triangle
</paragraph>
<caption id="DF16CFF68966FFAF11C30BC3B2259A91" pageId="1" pageNumber="68" startId="1.[226,276,1698,1721]" targetBox="[228,1284,986,1663]" targetPageId="1" targetType="figure">
<paragraph id="8BD69F7E8966FFAF11C30BC3B2259A91" blockId="1.[226,1285,1698,1754]" pageId="1" pageNumber="68">
Figs. 13. Habitus of
<taxonomicName id="4C69E4FD8966FFAF10F50BC2B3199AF2" authorityName="W. Horn" authorityYear="1901" box="[468,592,1699,1722]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B91D436C8966FFAF10F50BC2B3199AF2" box="[468,592,1699,1722]" italics="true" pageId="1" pageNumber="68">Neocollyris</emphasis>
</taxonomicName>
(s. str.) spp.: 1
<taxonomicName id="4C69E4FD8966FFAF12360BC2B2939AF2" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[791,986,1699,1722]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="trichocephalus" status="sp. n.">
<emphasis id="B91D436C8966FFAF12360BC2B2939AF2" box="[791,986,1699,1722]" italics="true" pageId="1" pageNumber="68">N. trichocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178966FFAF12C30BC3B5569AF1" box="[994,1055,1698,1721]" pageId="1" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
, holotype; 2 idem, paratype. 3
<taxonomicName id="4C69E4FD8966FFAF10540BA2B0A29A92" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[373,491,1731,1754]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. n.">
<emphasis id="B91D436C8966FFAF10540BA2B0A29A92" box="[373,491,1731,1754]" italics="true" pageId="1" pageNumber="68">N. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178966FFAF10D30BA3B3649A91" box="[498,557,1730,1753]" pageId="1" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
, Holotype. Scale line = 1 sm.
</paragraph>
</caption>
<caption id="DF16CFF68965FFAC100F090BB0F19889" pageId="2" pageNumber="69" startId="2.[302,352,1130,1153]" targetBox="[444,1228,194,1096]" targetPageId="2" targetType="figure">
<paragraph id="8BD69F7E8965FFAC100F090BB0F19889" blockId="2.[302,1360,1130,1217]" pageId="2" pageNumber="69">
Figs. 48.
<taxonomicName id="4C69E4FD8965FFAC10BC090BB35098C9" authorityName="W. Horn" authorityYear="1901" box="[413,537,1130,1153]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B91D436C8965FFAC10BC090BB35098C9" box="[413,537,1130,1153]" italics="true" pageId="2" pageNumber="69">Neocollyris</emphasis>
</taxonomicName>
(s. str.) spp.: 45 head, pronotum, and basal part of elytra, right lateral view; 68 head, dorsal view. 4, 6, 7
<taxonomicName id="4C69E4FD8965FFAC13B509EBB21A98E9" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[660,851,1162,1185]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="trichocephalus" status="sp. n.">
<emphasis id="B91D436C8965FFAC13B509EBB21A98E9" box="[660,851,1162,1185]" italics="true" pageId="2" pageNumber="69">N. trichocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178965FFAC127B09EBB2DC98E9" box="[858,917,1162,1185]" pageId="2" pageNumber="69" rank="species">sp. n.</taxonomicNameLabel>
; 5, 8
<taxonomicName id="4C69E4FD8965FFAC12C309EBB51198E9" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2022" box="[994,1112,1162,1185]" class="Insecta" family="Carabidae" genus="Neocollyris" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. n.">
<emphasis id="B91D436C8965FFAC12C309EBB51198E9" box="[994,1112,1162,1185]" italics="true" pageId="2" pageNumber="69">N. hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFE178965FFAC157F09EBB5D098E9" box="[1118,1177,1162,1185]" pageId="2" pageNumber="69" rank="species">sp. n.</taxonomicNameLabel>
, Holotype. Scale line = 1 mm.
</paragraph>
</caption>
<paragraph id="8BD69F7E8965FFAC100F081CB26599DC" blockId="2.[302,813,1277,1748]" box="[302,812,1405,1428]" pageId="2" pageNumber="69">
(
<figureCitation id="135283FB8965FFAC1017081CB03D99DC" box="[310,372,1405,1428]" captionStart="Figs" captionStartId="3.[226,276,1668,1691]" captionTargetBox="[352,1157,1144,1642]" captionTargetId="figure-434@3.[348,1157,1142,1645]" captionTargetPageId="3" captionText="Figs. 912. Palpi of Neocollyris (s. str.) spp. 9 maxillary palpus of N. trichocephalus sp. n.; 10 labial palpus of N. trichocephalus sp. n.; 11 maxillary palpus of N. hirsutus sp. n.; 12 labial palpus of N. hirsutus sp. n. Scale line = 1 mm." pageId="2" pageNumber="69">Fig. 9</figureCitation>
) and labial with the shape of right triangle
</paragraph>
<paragraph id="8BD69F7E8965FFAC100F08FCB24F9A9C" blockId="2.[302,813,1277,1748]" pageId="2" pageNumber="69">
(
<figureCitation id="135283FB8965FFAC101708FCB0CD99FC" box="[310,388,1437,1460]" captionStart="Figs" captionStartId="3.[226,276,1668,1691]" captionTargetBox="[352,1157,1144,1642]" captionTargetId="figure-434@3.[348,1157,1142,1645]" captionTargetPageId="3" captionText="Figs. 912. Palpi of Neocollyris (s. str.) spp. 9 maxillary palpus of N. trichocephalus sp. n.; 10 labial palpus of N. trichocephalus sp. n.; 11 maxillary palpus of N. hirsutus sp. n.; 12 labial palpus of N. hirsutus sp. n. Scale line = 1 mm." pageId="2" pageNumber="69">Fig. 10</figureCitation>
). Antennae relatively long; antennomere 3 the longest; scape and pedicel dark violet, antennomeres 34 violet, with brown apices; antennomere 5 violet, with brown ring in central part; antennomeres 611 dark brown, almost black, with violet reflections. Scape with a single apical seta, antennomeres 34 glabrous except for 56 thin and short setae on ventral ridge, apical half of antennomere 5 and antennomeres 611 densely pubescent with very short yellow setae.
</paragraph>
<paragraph id="8BD69F7E8965FFAC1273099CB40B9A3C" blockId="2.[850,1361,1277,1652]" pageId="2" pageNumber="69">
Pronotum (
<figureCitation id="135283FB8965FFAC12EC099CB57A995C" box="[973,1075,1277,1300]" captionStart="Figs" captionStartId="1.[226,276,1698,1721]" captionTargetBox="[228,1284,986,1663]" captionTargetId="figure-337@1.[226,840,984,1663]" captionTargetPageId="1" captionText="Figs. 13. Habitus of Neocollyris (s. str.) spp.: 1 N. trichocephalus sp. n., holotype; 2 idem, paratype. 3 N. hirsutus sp. n., Holotype. Scale line = 1 sm." pageId="2" pageNumber="69">Figs 1, 2</figureCitation>
,
<figureCitation id="135283FB8965FFAC1563099CB519995C" box="[1090,1104,1277,1300]" captionStart="Figs" captionStartId="2.[302,352,1130,1153]" captionTargetBox="[444,1228,194,1096]" captionTargetId="figure-342@2.[440,1231,189,1098]" captionTargetPageId="2" captionText="Figs. 48. Neocollyris (s. str.) spp.: 45 head, pronotum, and basal part of elytra, right lateral view; 68 head, dorsal view. 4, 6, 7 N. trichocephalus sp. n.; 5, 8 N. hirsutus sp. n., Holotype. Scale line = 1 mm." pageId="2" pageNumber="69">4</figureCitation>
) with a shallow collar, without anterior hump, moderately long, PL/ PW = 1.69­1.72, distinctly expanded in basal third; disc slightly convex in lateral view, with indistinct, sparse, shallow, transverse wrinkles in anterior half, unevenly punctured by setiferous pores, bearing very long white hairs; punctures on base of pronotum twice as large as on disc; anterior sulcus relatively wide and shallow, while posterior one more narrow and deeper. Prosternum, pro­episternum, mesothorax and mesosternum densely pubescent with long, white hairs.
</paragraph>
<paragraph id="8BD69F7E8965FFAC12730BFCB4199A9C" blockId="2.[850,1360,1693,1748]" pageId="2" pageNumber="69">
Elytra (
<figureCitation id="135283FB8965FFAC12870BFCB5449AFC" box="[934,1037,1693,1716]" captionStart="Figs" captionStartId="1.[226,276,1698,1721]" captionTargetBox="[228,1284,986,1663]" captionTargetId="figure-337@1.[226,840,984,1663]" captionTargetPageId="1" captionText="Figs. 13. Habitus of Neocollyris (s. str.) spp.: 1 N. trichocephalus sp. n., holotype; 2 idem, paratype. 3 N. hirsutus sp. n., Holotype. Scale line = 1 sm." pageId="2" pageNumber="69">Figs 1, 2</figureCitation>
) moderately long, EL/EW = 2.74­2.83, parallel sided, metallic violet with
</paragraph>
<paragraph id="8BD69F7E8964FFAD13560DF0B2389CEC" blockId="3.[631,881,145,164]" box="[631,881,145,164]" pageId="3" pageNumber="70">
<emphasis id="B91D436C8964FFAD13560DF0B2389CEC" box="[631,881,145,164]" italics="true" pageId="3" pageNumber="70">Anichtchenko A., Wiesner J.</emphasis>
</paragraph>
<paragraph id="8BD69F7E8964FFAD11C30DA0B0DE9DB0" blockId="3.[226,738,193,504]" pageId="3" pageNumber="70">light purple luster and without elytral maculation; covered with isodiametric microsculpture; disc with numerous regularly distributed, rounded, blue pits, slightly elongate and smaller in apical fifth; almost all pits with very small setiferous pore on anterior margin, bearing extremely long white hairs; suture moderately protruding. Sutural tooth on apice of elytra right, prominent, well visible, exterior angle of elytra obtuse and rounded. Scutellum black.</paragraph>
<paragraph id="8BD69F7E8964FFAD11C30F40B3749E30" blockId="3.[226,736,545,632]" pageId="3" pageNumber="70">Abdominal sternites black with violet luster, sternites with thin pale setae. All coxae, as well as fore­ and mid­trochanters black.</paragraph>
<paragraph id="8BD69F7E8964FFAD11C30FC0B3169F10" blockId="3.[226,737,673,856]" pageId="3" pageNumber="70">
Profemora dark violet, ventrally dark brown with violet reflections. Meso­ and metafemora dark brown with violet reflections, dorsally and apically violet. All tibiae dark violet. Pro­ and mesotarsi uniformly violet; metatarsi light yellow, with two apical tarsomeres violet (
<figureCitation id="135283FB8964FFAD10D00E20B31B9F10" box="[497,594,833,856]" captionStart="Figs" captionStartId="1.[226,276,1698,1721]" captionTargetBox="[228,1284,986,1663]" captionTargetId="figure-337@1.[226,840,984,1663]" captionTargetPageId="1" captionText="Figs. 13. Habitus of Neocollyris (s. str.) spp.: 1 N. trichocephalus sp. n., holotype; 2 idem, paratype. 3 N. hirsutus sp. n., Holotype. Scale line = 1 sm." pageId="3" pageNumber="70">Figs 1, 2</figureCitation>
).
</paragraph>
<paragraph id="8BD69F7E8964FFAD11C30EE0B0199810" blockId="3.[226,737,897,1112]" pageId="3" pageNumber="70">
Aedeagus (
<figureCitation id="135283FB8964FFAD10780EE0B0869FD0" box="[345,463,897,920]" captionStart="Figs" captionStartId="4.[302,352,1307,1330]" captionTargetBox="[305,810,669,1273]" captionTargetId="figure-592@4.[302,812,666,1274]" captionTargetPageId="4" captionText="Figs. 1517. Aedeagus of Neocollyris (s. str.) spp.: 1516 N. trichocephala sp. n., Holotype and paratype; 17 N. hirsuta sp. n., Holotype. Scale line = 1 mm." pageId="3" pageNumber="70">Figs 15, 16</figureCitation>
) relatively long, 2.0
<quantity id="4C91329B8964FFAD13930EE0B0469FF0" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="3" pageNumber="70" unit="mm" value="2.15">2.15 mm</quantity>
; with a wide, gradually curved basal bulb; ventral side of median lobe straight; apical part relatively long, strongly downturned, apex narrow and acute; internal sack with short and thick flagellum, and with thin undulate sclerite at base of flagellum.
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58964FFAD12270DA1B59D9C90" box="[774,1236,192,216]" pageId="3" pageNumber="70" type="distribution">
<paragraph id="8BD69F7E8964FFAD12270DA1B59D9C90" blockId="3.[774,1236,192,216]" box="[774,1236,192,216]" pageId="3" pageNumber="70">
<emphasis id="B91D436C8964FFAD12270DA1B2D29C90" bold="true" box="[774,923,192,216]" pageId="3" pageNumber="70">Distribution.</emphasis>
<collectingCountry id="F37EDFEE8964FFAD12810DA0B5499C90" box="[928,1024,193,216]" name="Vietnam" pageId="3" pageNumber="70">Vietnam</collectingCountry>
: province
<collectingRegion id="49AD519C8964FFAD154C0DA0B5989C90" box="[1133,1233,193,216]" country="Vietnam" name="Da Nang" pageId="3" pageNumber="70">Da Nang</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C373CCF58964FFAD12270C61B5479D30" pageId="3" pageNumber="70" type="etymology">
<paragraph id="8BD69F7E8964FFAD12270C61B5479D30" blockId="3.[774,1285,256,376]" pageId="3" pageNumber="70">
<emphasis id="B91D436C8964FFAD12270C61B2CF9D50" bold="true" box="[774,902,256,280]" pageId="3" pageNumber="70">Etymology.</emphasis>
The name of the new species is Latinized form from the Greek “
<emphasis id="B91D436C8964FFAD151C0C41B5E99D70" bold="true" box="[1085,1184,288,312]" pageId="3" pageNumber="70">tricwto´ j</emphasis>
hairy, and “
<emphasis id="B91D436C8964FFAD121E0C21B2D69D10" bold="true" box="[831,927,320,344]" pageId="3" pageNumber="70">ke´ faloj</emphasis>
head, because of its unusually punctate and hairy head.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,180 @@
<document id="F55F8848D27927026288B7E57EDB3542" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722658991509" checkinUser="felipe" docAuthor="Legalov, Andrei A." docDate="2022" docId="03C08785783A8D29FF421B55FAA4A139" docLanguage="en" docName="BalticJColeopterol.22.1.123-126.pdf" docOrigin="Baltic Journal of Coleopterology 22 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.4:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="4" docTitle="Dryophthorus bacanensis Legalov 2022, sp. nov." docType="treatment" docVersion="1" lastPageNumber="125" masterDocId="FFF9FFFD783B8D2BFFA11834FFB4A441" masterDocTitle="A new species of the genus Dryophthorus Germar, 1823 (Coleoptera, Curculionidae) from Maluku Islands" masterLastPageNumber="126" masterPageNumber="123" pageNumber="124" updateTime="1722691601247" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="1AD6E16F5FD29811BAAAA08162193025">A new species of the genus Dryophthorus Germar, 1823 (Coleoptera, Curculionidae) from Maluku Islands</mods:title>
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<mods:namePart id="F45D7C7315F3B63B5076024491CB8C73">Legalov, Andrei A.</mods:namePart>
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<paragraph id="8BD63693783A8D2AFF421B55FD70A738" blockId="1.[227,708,865,921]" box="[227,708,865,889]" pageId="1" pageNumber="124">
<heading id="D09E81FF783A8D2AFF421B55FD70A738" bold="true" box="[227,708,865,889]" fontSize="10" level="2" pageId="1" pageNumber="124" reason="4">
<emphasis id="B91DEA81783A8D2AFF421B55FD70A738" bold="true" box="[227,708,865,889]" pageId="1" pageNumber="124">
<taxonomicName id="4C694D10783A8D2AFF421B55FDD5A738" authority="Legalov" authorityName="Legalov" authorityYear="2022" box="[227,609,865,889]" class="Insecta" family="Dryophthoridae" genus="Dryophthorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="bacanensis" status="sp. nov.">
<emphasis id="B91DEA81783A8D2AFF421B55FDB4A738" bold="true" box="[227,512,865,889]" italics="true" pageId="1" pageNumber="124">Dryophthorus bacanensis</emphasis>
Legalov
</taxonomicName>
,
<taxonomicNameLabel id="A22E57FA783A8D2AFDCF1B55FD70A738" box="[622,708,865,889]" pageId="1" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3736518783A8D2AFF421BB6FB67A719" pageId="1" pageNumber="124" type="description">
<paragraph id="8BD63693783A8D2AFF421BB6FE80A7D8" blockId="1.[227,708,865,921]" box="[227,308,898,921]" pageId="1" pageNumber="124">
(
<figureCitation id="13522A16783A8D2AFF4A1BB6FE9DA7D8" box="[235,297,898,921]" captionStart="Fig" captionStartId="1.[774,813,1634,1657]" captionTargetBox="[783,1277,884,1601]" captionTargetId="figure-581@1.[781,1278,883,1603]" captionTargetPageId="1" captionText="Fig. 1. Dryophthorus bacanensis sp. nov. male: a habitus, holotype, dorsally, b aedeagus, paratype, dorsally. Scale bar = 1.0 mm for a, d; 0.2 mm for b." pageId="1" pageNumber="124">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="8BD63693783A8D2AFCA718F5FAB2A679" blockId="1.[774,1286,193,568]" pageId="1" pageNumber="124">as long as and about 0.7 times as narrow as scape. Antennomeres 3­ 5 subconical. Antennomere 3 about 0.9 times as long as wide in apex, 0.8 times as long as and 0.9 times as narrow as antennomere 2. Antennomere 4 0.5 times as long as wide in apex, about 0.8 times as long as and about 1.3 times as wide as antennomere 3. Antennomere 5 about 0.4 times as long as wide, about 0.8 times as long as and about 1.2 times as wide as antennomere 4.Antennal club compact, not truncate, about 1.6 times as long as wide, slightly shorter than antennomeres 2­ 5 combined, with tomentose apex.</paragraph>
<paragraph id="8BD63693783A8D2AFCA71A55FB67A719" blockId="1.[774,1285,609,856]" pageId="1" pageNumber="124">Pronotum campanulate, 1.3­ 1.4 times as long as wide at apex, slightly longer than wide at midlength and at pronotal base. Pronotal disk weakly convex dorsally, coarsely punctate. Intervals between points smaller than their diameter. Maximum width before middle. Base of pronotum distinctly narrower than base of elytra. Scutellum small, triangular, about 1.4 times as long as wide.</paragraph>
</subSubSection>
<subSubSection id="C3736518783A8D2AFF421BF5FE5CA018" pageId="1" pageNumber="124" type="materials_examined">
<paragraph id="8BD63693783A8D2AFF421BF5FE5CA018" blockId="1.[227,737,961,1113]" pageId="1" pageNumber="124">
<emphasis id="B91DEA81783A8D2AFF421BF5FE39A798" bold="true" box="[227,397,961,985]" pageId="1" pageNumber="124">Type material:</emphasis>
<materialsCitation id="3B013CCE783A8D2AFE341BF5FE0BA078" collectingDate="2008-05-02" collectingDateMax="2008-05-12" collectingDateMin="2008-05-02" collectionCode="ISEA" collectorName="Bacan Isl., SE" country="Indonesia" county="St. Jakl" elevation="535" location="Male" municipality="Mt. Sibela" pageId="1" pageNumber="124" specimenCount="1" typeStatus="holotype">
<emphasis id="B91DEA81783A8D2AFE341BF5FDB0A798" bold="true" box="[405,516,961,985]" pageId="1" pageNumber="124">
<typeStatus id="54D28831783A8D2AFE341BF5FDB4A798" box="[405,512,961,985]" pageId="1" pageNumber="124" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EB66048783A8D2AFDAD1BF6FDF7A798" LSID="urn:lsid:plazi:treatment:03C08785783A8D29FF421B55FAA4A139:8EB66048783A8D2AFDAD1BF6FDF7A798" box="[524,579,962,985]" country="Indonesia" county="St. Jakl" municipality="Mt. Sibela" name="Male" pageId="1" pageNumber="124">Male</location>
(
<collectionCode id="ED78AE56783A8D2AFDF31BF6FD27A798" box="[594,659,962,985]" pageId="1" pageNumber="124">ISEA</collectionCode>
),
<collectingCountry id="F37E7603783A8D2AFD061BF6FEA8A7B8" name="Indonesia" pageId="1" pageNumber="124">Indonesia</collectingCountry>
, N Moluccas,
<collectorName id="269C5345783A8D2AFE6B1BD6FDDBA7B8" box="[458,623,994,1017]" pageId="1" pageNumber="124">
<location id="8EB66048783A8D2AFE6B1BD6FD89A7B8" LSID="urn:lsid:plazi:treatment:03C08785783A8D29FF421B55FAA4A139:8EB66048783A8D2AFE6B1BD6FD89A7B8" box="[458,573,994,1017]" country="Indonesia" county="St. Jakl" municipality="Mt. Sibela" name="Bacan Isl." pageId="1" pageNumber="124">Bacan Isl.</location>
, SE
</collectorName>
slopes of
<collectingMunicipality id="6BB2ACE9783A8D2AFF421C36FEFBA058" box="[227,335,1026,1049]" pageId="1" pageNumber="124">Mt.Sibela</collectingMunicipality>
,
<locationDeviation id="9A642FB1783A8D2AFEF81C36FD9CA058" box="[345,552,1026,1049]" location="Makian" pageId="1" pageNumber="124">
<quantity id="4C919B76783A8D2AFEF81C36FE3AA058" box="[345,398,1026,1049]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="124" unit="km" value="5.0">5 km</quantity>
SE of Makian
</locationDeviation>
vill.,
<quantity id="4C919B76783A8D2AFDC71C36FD68A058" box="[614,732,1026,1049]" metricMagnitude="2" metricUnit="m" metricValue="5.35" metricValueMax="5.7" metricValueMin="5.0" pageId="1" pageNumber="124" unit="m" value="535.0" valueMax="570.0" valueMin="500.0">
<elevation id="0044D1A0783A8D2AFDC71C36FD68A058" box="[614,732,1026,1049]" metricMagnitude="2" metricUnit="m" metricValue="5.35" metricValueMax="5.7" metricValueMin="5.0" pageId="1" pageNumber="124" unit="m" value="535.0" valueMax="570.0" valueMin="500.0">500­ 570 m</elevation>
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,
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<collectingDate id="EF93E9BB783A8D2AFF421C16FED3A078" box="[227,359,1058,1081]" pageId="1" pageNumber="124" value="2008-05-02" valueMax="2008-05-12" valueMin="2008-05-02">2­ 12.V.2008</collectingDate>
</date>
,
<collectingCounty id="62B74E1F783A8D2AFED11C16FE0BA078" box="[368,447,1058,1081]" pageId="1" pageNumber="124">St. Jakl</collectingCounty>
</materialsCitation>
.
<materialsCitation id="3B013CCE783A8D2AFE661C16FD69A078" box="[455,733,1058,1081]" collectionCode="ISEA" country="Indonesia" pageId="1" pageNumber="124" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54D28831783A8D2AFE661C16FD85A078" box="[455,561,1058,1081]" pageId="1" pageNumber="124" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D6FFD1A783A8D2AFD981C16FD3DA078" box="[569,649,1058,1081]" count="2" pageId="1" pageNumber="124" type="male">2 males</specimenCount>
(
<collectionCode id="ED78AE56783A8D2AFD341C16FD61A078" box="[661,725,1058,1081]" pageId="1" pageNumber="124">ISEA</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B013CCE783A8D2AFF421C76FE50A018" box="[227,484,1090,1113]" collectionCode="ISEA" country="Indonesia" pageId="1" pageNumber="124" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D6FFD1A783A8D2AFF421C76FEFEA018" box="[227,330,1090,1113]" count="2" pageId="1" pageNumber="124" type="female">2 females</specimenCount>
(
<collectionCode id="ED78AE56783A8D2AFEF91C76FE2DA018" box="[344,409,1090,1113]" pageId="1" pageNumber="124">ISEA</collectionCode>
), idem
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3736518783A8D29FF421CB5FA82A059" lastPageId="2" lastPageNumber="125" pageId="1" pageNumber="124" type="description">
<paragraph id="8BD63693783A8D2AFF421CB5FD7EA0F8" blockId="1.[227,738,1153,1209]" pageId="1" pageNumber="124">
<emphasis id="B91DEA81783A8D2AFF421CB5FED3A0D8" bold="true" box="[227,359,1153,1177]" pageId="1" pageNumber="124">Description.</emphasis>
Body black, with matted pubescence. Antennae, apex of tibiae and tarsi red­brown.
</paragraph>
<paragraph id="8BD63693783A8D29FF421CD5FD33A59D" blockId="1.[227,737,1249,1753]" lastBlockId="2.[302,813,193,476]" lastPageId="2" lastPageNumber="125" pageId="1" pageNumber="124">
<emphasis id="B91DEA81783A8D2AFF421CD5FE92A0B8" bold="true" box="[227,294,1249,1273]" pageId="1" pageNumber="124">Male:</emphasis>
Head subconical. Mandibles small. Rostrum quite short, 0.7 times as long as pronotum, 2.2 times as long as wide at apex, 1.8­ 2.2 times as long as wide at midlength, 1.4­ 2.3 times as long as wide at base, weakly curved, sparsely punctate. Apex of rostrum finely punctate. Eyes large, linear, 2.4 times as wide as length, not protruding from contour of head, linear, separated ventrally, coarsely faceted. Forehead flat, 0.8 times as narrow as rostrum base width. Temples 1.4­ 1.8 times as long as eye length. Antennal scrobes directed base of rostrum. Antennae inserted before middle of rostrum. Scape long, about 4.6 times as long as wide in apex, extends beyond back of eye. Funicle 4­ segmented.Antennomere 2 suboval, 1.4 times as long as wide in apex, about 0.2 times Elytra almost subparallel, at base 1.5­ 1.6 times as long as wide, at midlength 1.6 times as long as wide, at apical fourth 2.4­ 2.5 times as long as wide, 2.0 times as long as pronotum. Humeri weakly flattened. Elytral striae distinct, wide. Stria 9 short, fused with stria 10 at level of metacoxae. Interstriae convex, narrow, 0.5­ 0.6 times as narrow as width of striae. Interstria 4 fused with interstriae 1 and 2 apically.
</paragraph>
<caption id="DF16661B783A8D2AFCA71E56FC2DA298" pageId="1" pageNumber="124" startId="1.[774,813,1634,1657]" targetBox="[783,1277,884,1601]" targetPageId="1" targetType="figure">
<paragraph id="8BD63693783A8D2AFCA71E56FC2DA298" blockId="1.[774,1285,1634,1753]" pageId="1" pageNumber="124">
Fig. 1.
<taxonomicName id="4C694D10783A8D2AFCF21E56FBDDA238" authorityName="Legalov" authorityYear="2022" box="[851,1129,1634,1657]" class="Insecta" family="Dryophthoridae" genus="Dryophthorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="bacanensis" status="sp. nov.">
<emphasis id="B91DEA81783A8D2AFCF21E56FBDDA238" box="[851,1129,1634,1657]" italics="true" pageId="1" pageNumber="124">Dryophthorus bacanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FA783A8D2AFBD11E56FB77A238" box="[1136,1219,1634,1657]" pageId="1" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
male: a habitus, holotype, dorsally, b aedeagus, paratype, dorsally. Scale bar = 1.0 mm for a, d; 0.2 mm for b.
</paragraph>
</caption>
<paragraph id="8BD6369378398D29FE8F1A32FE25A740" blockId="2.[302,813,518,769]" pageId="2" pageNumber="125">Prosternum punctate, without postocular lobes. Precoxal portion of prosternum equal to procoxal cavity length. Postcoxal portion of prosternum short, about 0.5 times as long as procoxal cavity. Procoxal cavities widely separated. Metepistrnum concealed by elytra. Metaventrite 2.5.1 times as long as length of metacoxa, flattened, coarsely punctate.</paragraph>
<paragraph id="8BD6369378398D29FE8F1B1FFDA0A067" blockId="2.[302,813,811,1062]" pageId="2" pageNumber="125">Abdomen weakly convex ventrally, punctate. Ventrites 1 and 2 weekly impressed. Ventrite 1 slightly shorter than length of metacoxa. Ventrite 2 1.4 times as long as ventrite 1. Ventrite 3 0.3 times shorter than ventrite 2. Ventrite 4 slightly shorter than ventrite 3. Ventrite 5 3.6 times as long as ventrite 4, densely punctate. Pygidium impressed in meddle.</paragraph>
<paragraph id="8BD6369378398D29FE8F1C64FB4CA5F9" blockId="2.[302,814,1104,1680]" lastBlockId="2.[850,1361,193,440]" pageId="2" pageNumber="125">Procoxae spherical. Metacoxae almost transverse. Femora slightly thickened, without tooth. Metafemora not extending beyond apex of abdomen. Tibiae weakly curved, with large uncus.Tarsi long. Tarsomeres 1­ 3 subconical. Tarsomere 4 square. Tarsomere 5 elongate.Tarsal claws free, divergent. Protarsi: tarsomeres 1­ 3 subequal in width; tarsomere 1 about 1.4 times as long as wide; tarsomere 2 about 1.1 times as long as wide, 0.8 times as long as tarsomere 1; tarsomere 3 about 1.4 times as long as wide, about 1.3 times as long as tarsomere 2; tarsomeres 4 and 5 subequal in width; tarsomere 4 equal in length and width, 0.4 times as long as and about 0.6 times as narrow as tarsomere 3; tarsomere 5 about 3.8 times as long as wide, about 3.8 times as long as tarsomere 4. Metatarsi: tarsomeres 1­3 subequal in width; tarsomere 1 2.0 times as long as wide; tarsomere 2 about 1.3 times as long as wide, about 0.7 times as long as tarsomere 1; tarsomere 3 about 1.7 times as long as wide, about 1.3 times as long as tarsomere 2; tarsomere 4 equal in length and width, 0.5 times as long as and about 0.7 times as narrow as tarsomere 3; tarsomere 5 3.2 times as long as wide, about 4.0 times as long as and about 1.3 times as wide as tarsomere 4.</paragraph>
<paragraph id="8BD6369378398D29FCF319D5FBC0A659" blockId="2.[850,1361,481,536]" pageId="2" pageNumber="125">
Total body length (without rostrum)
<quantity id="4C919B7678398D29FB7119D5FAF9A5B9" box="[1232,1357,481,504]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="2" pageNumber="125" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.6­ 2.7 mm</quantity>
. Length of rostrum
<quantity id="4C919B7678398D29FBBA1A35FBC4A659" box="[1051,1136,513,536]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="125" unit="mm" value="0.6">0.6 mm</quantity>
.
</paragraph>
<paragraph id="8BD6369378398D29FCF31A74FA82A059" blockId="2.[850,1364,576,1048]" pageId="2" pageNumber="125">
<emphasis id="B91DEA8178398D29FCF31A74FC1AA619" bold="true" box="[850,942,576,600]" pageId="2" pageNumber="125">Female:</emphasis>
Rostrum 0.7­ 0.8 times as long as pronotum, 2.3­ 2.6 times as long as wide at apex, 2.8­ 2.9 times as long as wide at midlength, 2.4­ 2.6 times as long as wide at base. Pronotum 1.4­ 1.5 times as long as wide at apex, 1.1 times as long as wide at midlength and 1.1­ 1.2 times as long as wide at pronotal base. Elytra at base 1.4­ 1.8 times as long as wide at midlength and 2.7­ 3.0 times as long as wide at apical fourth, 2.0 times as long as pronotum. Precoxal portion of prosternum about 1.1 times as long as procoxal cavity length. Postcoxal portion of prosternum about 0.4 times as long as procoxal cavity. Ventrites 1 and 2 weekly convex. Total body length (without rostrum)
<quantity id="4C919B7678398D29FC141C35FBBEA059" box="[949,1034,1025,1048]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="125" unit="mm" value="2.7">2.7 mm</quantity>
. Length of rostrum
<quantity id="4C919B7678398D29FB7C1C35FA85A059" box="[1245,1329,1025,1048]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="125" unit="mm" value="0.6">0.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C373651878398D29FCF31C74FBDEA0B9" pageId="2" pageNumber="125" type="diagnosis">
<paragraph id="8BD6369378398D29FCF31C74FBDEA0B9" blockId="2.[850,1361,1088,1272]" pageId="2" pageNumber="125">
<emphasis id="B91DEA8178398D29FCF31C74FC7CA019" bold="true" box="[850,968,1088,1112]" pageId="2" pageNumber="125">Diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="4C694D1078398D29FABC1C75FB50A039" authority="Voss, 1940" authorityName="Voss" authorityYear="1940" class="Insecta" family="Dryophthoridae" genus="Dryophthorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="subtruncatus">
<emphasis id="B91DEA8178398D29FABC1C75FBE9A039" italics="true" pageId="2" pageNumber="125">Dryophthorus subtruncatus</emphasis>
<bibRefCitation id="EFF84B6278398D29FBC61C55FB50A039" author="Voss E." box="[1127,1252,1121,1144]" pageId="2" pageNumber="125" pagination="17 - 93" refId="ref2244" refString="Voss E. 1940. Uber Russelkafer der Indomalayischen Subregion, vorwiegend von Java (Col.; Curc.). Teil I. (82. Beitrag zur Kenntnis der Curculioniden). Tidschrift voor Entomologie, 83, 17 - 93." type="journal article" year="1940">Voss, 1940</bibRefCitation>
</taxonomicName>
from the
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but differs in the smaller body sizes, glabrous apex of the rostrum, punctate, not rugose rostrum in basal half, and the apex of the aedeagus impressed in middle.
</paragraph>
</subSubSection>
<subSubSection id="C373651878398D29FCF31D14FB1AA179" box="[850,1198,1312,1336]" pageId="2" pageNumber="125" type="etymology">
<paragraph id="8BD6369378398D29FCF31D14FB1AA179" blockId="2.[850,1198,1312,1336]" box="[850,1198,1312,1336]" pageId="2" pageNumber="125">
<emphasis id="B91DEA8178398D29FCF31D14FC60A179" bold="true" box="[850,980,1312,1336]" pageId="2" pageNumber="125">Etymology.</emphasis>
From Bacan Island.
</paragraph>
</subSubSection>
<subSubSection id="C373651878398D29FCF31D54FAA4A139" box="[850,1296,1376,1400]" pageId="2" pageNumber="125" type="distribution">
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<emphasis id="B91DEA8178398D29FCF31D54FC52A139" bold="true" box="[850,998,1376,1400]" pageId="2" pageNumber="125">Distribution.</emphasis>
<collectingCountry id="F37E760378398D29FC4D1D55FBEDA139" box="[1004,1113,1377,1400]" name="Indonesia" pageId="2" pageNumber="125">Indonesia</collectingCountry>
:
<collectingCountry id="F37E760378398D29FBC31D55FAB8A139" box="[1122,1292,1377,1400]" name="Indonesia" pageId="2" pageNumber="125">Maluku Islands</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,236 @@
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<mods:title id="3895AE4BEBB5DEF4130A7E4FDD4F5532">A new species and new data of Cosmodiscus Sloan, 1907 (Coleoptera, Carabidae, Pterostichinae) from the Philippines</mods:title>
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<taxonomicName id="4C6ED9E7F1606B244320FAFCFDF0FAED" authority="Anichtchenko" authorityName="Anichtchenko" authorityYear="2023" box="[114,592,1294,1318]" class="Insecta" family="Carabidae" genus="Cosmodiscus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="shavrini" status="sp.">
<emphasis id="B91A7E76F1606B244320FAFCFDF0FAED" bold="true" box="[114,592,1294,1318]" pageId="1" pageNumber="126">
<emphasis id="B91A7E76F1606B244320FAFCFEC9FAED" bold="true" box="[114,361,1294,1318]" italics="true" pageId="1" pageNumber="126">Cosmodiscus shavrini</emphasis>
Anichtchenko, sp.
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<emphasis id="B91A7E76F1606B244320FADEFF02FA8F" bold="true" box="[114,162,1324,1348]" pageId="1" pageNumber="126">nov.</emphasis>
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<emphasis id="B91A7E76F1606B244320FA9BFF7CFA4A" bold="true" box="[114,220,1385,1409]" pageId="1" pageNumber="126">Material.</emphasis>
<materialsCitation id="3B06A839F1606B2443BDFA9BFE12FA37" collectingDate="2019-05-04" collectionCode="V, DUBC" country="Philippines" county="Mindanao" elevation="900" latitude="7.2764726" location="Pantukan" longLatPrecision="1" longitude="128.17023" municipality="Araibo" pageId="1" pageNumber="126" specimenCount="1" stateProvince="Campostela Valley" typeStatus="holotype">
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<typeStatus id="54D51CC6F1606B2443BDFA9BFEF7FA4A" box="[239,343,1385,1409]" pageId="1" pageNumber="126" type="holotype">Holotype</typeStatus>
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, male: “
<collectingCountry id="F379E2F4F1606B244298FA98FDEDFA49" box="[458,589,1386,1410]" name="Philippines" pageId="1" pageNumber="126">Philippines</collectingCountry>
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,
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, /
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,
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, /
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,
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, Shavrin A.
<collectionCode id="ED7F3AA1F1606B2443BEFA16FEA2FA37" box="[236,258,1508,1532]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="126" type="Museum">V</collectionCode>
. leg.” (
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)
</materialsCitation>
.
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.
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: “
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,
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,
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/
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<collectingDate id="EF947D4CF1606B244320F9D0FF48F9F1" box="[114,232,1570,1594]" pageId="1" pageNumber="126" value="2014-07">July 2014</collectingDate>
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” (
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)
</materialsCitation>
;
<materialsCitation id="3B06A839F1606B24423AF9D0FEB9F9BC" collectingDate="2015-01" collectionCode="DUBC" country="Philippines" location="Mindanao" municipality="Masara" pageId="1" pageNumber="126" specimenCount="1" specimenCount-female="1" stateProvince="Compostela" typeStatus="paratype">
<specimenCount id="9D6869EDF1606B24423AF9D0FE74F9F1" box="[360,468,1570,1594]" count="1" pageId="1" pageNumber="126" type="female">1 female</specimenCount>
: “
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,
<collectingRegion id="49AA6C86F1606B244320F9B2FEECF993" box="[114,332,1600,1624]" country="Philippines" name="Compostela" pageId="1" pageNumber="126">Compostela Valley</collectingRegion>
,
<location id="8EB1F4BFF1606B244233F9B2FE6DF993" LSID="urn:lsid:plazi:treatment:03C71372F1606B264320FAFCFDE4FC67:8EB1F4BFF1606B244233F9B2FE6DF993" box="[353,461,1600,1624]" country="Philippines" municipality="Masara" name="Mindanao" pageId="1" pageNumber="126" stateProvince="Compostela">Mindanao</location>
/
<date id="FFD084A4F1606B2442A5F9B2FF0AF9BC" pageId="1" pageNumber="126" value="2015-01">
<collectingDate id="EF947D4CF1606B2442A5F9B2FF0AF9BC" pageId="1" pageNumber="126" value="2015-01">January 2015</collectingDate>
</date>
” (
<collectionCode id="ED7F3AA1F1606B244393F9ADFEB1F9BC" box="[193,273,1631,1655]" pageId="1" pageNumber="126">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD1A264F1606B244320F954FB9BFD18" blockId="1.[114,154,1702,1726]" lastBlockId="1.[624,1103,177,723]" pageId="1" pageNumber="126">
126
<emphasis id="B91A7E76F1606B244122FF43FD44FF02" bold="true" box="[624,740,177,201]" pageId="1" pageNumber="126">Diagnosis.</emphasis>
New species is similar to
<taxonomicName id="4C6ED9E7F1606B244762FF40FC23FF23" authority="(Bates, 1873)" baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Cosmodiscus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="platynotus">
<emphasis id="B91A7E76F1606B244762FF40FD7FFF22" italics="true" pageId="1" pageNumber="126">C. platynotus</emphasis>
(Bates, 1873)
</taxonomicName>
by its uniformly black body color, but easily distinguishable by strongly transverse pronotum, with maximum width at the middle and not dentate humeri, i.e. in
<taxonomicName id="4C6ED9E7F1606B24405DFEB9FC3EFEA8" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[783,926,331,355]" class="Insecta" family="Carabidae" genus="Cosmodiscus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="platynotus">
<emphasis id="B91A7E76F1606B24405DFEB9FC3EFEA8" box="[783,926,331,355]" italics="true" pageId="1" pageNumber="126">C. platynotus</emphasis>
</taxonomicName>
pronotum trapezoidal, and broadest distinctly behind the middle and humeri slightly dentate. By the transverse shape of pronotum, new species is similar with
<taxonomicName id="4C6ED9E7F1606B24407DFE34FD08FE37" authority="Darlington, 1962" authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Cosmodiscus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B91A7E76F1606B24407DFE34FC1CFE15" box="[815,956,454,478]" italics="true" pageId="1" pageNumber="126">C. brunneus</emphasis>
<bibRefCitation id="EFFFDF95F1606B24409CFE37FD08FE37" author="Darlington P. J." pageId="1" pageNumber="126" pagination="321 - 564" refId="ref1814" refString="Darlington P. J. 1962. The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini. Bulletin of the Museum of Comparative Zoology at Harvard College 126 (3): 321 - 564." type="journal article" year="1962">Darlington, 1962</bibRefCitation>
</taxonomicName>
, known from New
<collectingCountry id="F379E2F4F1606B2440D4FE16FC77FE37" box="[902,983,484,508]" name="Guinea-Bissau" pageId="1" pageNumber="126">Guinea</collectingCountry>
. It can be differentiated by concave anterior margin of pronotum, with anteriorly prominent angles, and by completely rounded posterior angles of pronotum, i.e. in
<taxonomicName id="4C6ED9E7F1606B2440EEFDADFBE8FDBC" authorityName="Darlington" authorityYear="1962" box="[956,1096,607,631]" class="Insecta" family="Carabidae" genus="Cosmodiscus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B91A7E76F1606B2440EEFDADFBE8FDBC" box="[956,1096,607,631]" italics="true" pageId="1" pageNumber="126">C. brunneus</emphasis>
</taxonomicName>
anterior margin straight, anterior angles of pronotum not prominent and posterior angles obtuse but distinct (
<bibRefCitation id="EFFFDF95F1606B244021FD49FB8FFD18" author="Darlington P. J." box="[883,1071,699,723]" pageId="1" pageNumber="126" pagination="321 - 564" refId="ref1814" refString="Darlington P. J. 1962. The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini. Bulletin of the Museum of Comparative Zoology at Harvard College 126 (3): 321 - 564." type="journal article" year="1962">Darlington, 1962</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C374F1EFF1606B274122FD0AFBB8FA9C" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126" type="description">
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<emphasis id="B91A7E76F1606B244122FD0AFD54FCDB" bold="true" box="[624,756,760,784]" pageId="1" pageNumber="126">Description</emphasis>
. BL =
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. Body
</paragraph>
<paragraph id="8BD1A264F1606B244122FCE5FC1EFB88" blockId="1.[624,1103,760,1091]" pageId="1" pageNumber="126">
(
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) shiny black, explanate lateral margins of pronotum, reflexed lateral margin of elytra, legs, antennae, mouthparts and labrum yellow. Ventral side black, except for redmiddle of pro-, meso- and meta-sternum. Dorsal microsculpture meshed, very superficial, almost obsolete on head, strongly transverse on pronotum and elytra, granulate in marginal groove of the latter, not well traceable on pronotal disc.
</paragraph>
<paragraph id="8BD1A264F1606B274122FB9AFDD3FE74" blockId="1.[624,1103,1128,1643]" lastBlockId="2.[189,669,178,447]" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126">Head with abruptly prominent eyes and a nearly imperceptible neck constriction on sides. HW/PW = 0.56 1.57. Frontal sulci short, deep, very slightly diverging based, disappearing on a level with anterior supraocular setae. Mandibles short, laterally rounded. Labrum truncate, weakly concave at middle. Antennae reaching pronotal base. Pronotum strongly transverse, PW/PL = 1.6 1.61. Sides evenly rounded, lateral margins slightly reflexed, borderline between explanate margins and disc obliterate basally. Margins finely beaded throughout except basally, apical bead very distinct, medially flat and much wider than laterally. Base truncate and gently trisinuate; basal angles obtuse and rounded. Basal sulci moderately deep, indistinctly converging basally, much shallow basally. Apical margin rather deeply concaved between obtuse and rounded apical angles. Median line fine, nearly entire, obliterate just basally. Disc convex and smooth; anterior and lateral margins, and interval between basal sulci irregularly and very shallowly rugate; basal angles slightly convex and smooth.</paragraph>
<paragraph id="8BD1A264F1636B2741E9FF40FC36FE6B" blockId="2.[699,1178,178,416]" pageId="2" pageNumber="127">vatedin apical third and toward lateral margin. Scutellar setigerous pores present, on base of stria II, slightly removed back from basal margin with distance of diameter of pore or so. Discal pore indistinct. Stria VII with two apical setae. Umbilicate series consist of 15 setigerous pores (6 humeral, 1 lateral and 8 apical).</paragraph>
<paragraph id="8BD1A264F1636B2741E9FE37FC4AFD7F" blockId="2.[699,1179,453,692]" pageId="2" pageNumber="127">Prosternum, proepisterna and abdominal sterna smooth. Anterior margin of mesoepisterna, metepisterna and metasternum densely shallowly punctate. Anterior margin of abdominal sternites IVVI with pair of setae; sternite III asetose. Prosternum without median groove, prosternal process convex and apically beaded.</paragraph>
<paragraph id="8BD1A264F1636B2741E9FD28FB96FBCE" blockId="2.[699,1179,730,1029]" pageId="2" pageNumber="127">Legs. Protarsomeres I-IV of males symmetrically dilated; in females protarsomeres I and II modified, with internal angle strongly extended forward. Protibia with three latero-apical spines. Metafemur bisetose. Mesotibia with four spiniform anterolateral setae, inner setal brush consisting of 7+1 setae; metatibia with two spiniform seta at middle of external ridge. First meso- and metatarsomere 1 externally sulcate.</paragraph>
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<paragraph id="8BD1A264F1636B2743EFFB01FE84FA82" blockId="2.[189,668,1267,1353]" pageId="2" pageNumber="127">
Fig. 1. Habitus of
<taxonomicName id="4C6ED9E7F1636B2742F0FB06FE29FAE1" authority="Anichtchenko" authorityName="Anichtchenko" authorityYear="2023" class="Insecta" family="Carabidae" genus="Cosmodiscus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="shavrini" status="sp.">
<emphasis id="B91A7E76F1636B2742F0FB06FD35FAC7" box="[418,661,1268,1292]" italics="true" pageId="2" pageNumber="127">Cosmodiscus shavrini</emphasis>
Anichtchenko,
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<emphasis id="B91A7E76F1636B27423BFAE0FE29FAE1" bold="true" box="[361,393,1298,1322]" pageId="2" pageNumber="127">sp.</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="B91A7E76F1636B2742CAFAE0FE68FAE1" bold="true" box="[408,456,1298,1322]" pageId="2" pageNumber="127">nov.</emphasis>
(holotype). Scale bar 1mm.
</paragraph>
</caption>
<paragraph id="8BD1A264F1636B2743EFFA9CFBB8FA9C" blockId="2.[189,669,1390,1659]" lastBlockId="2.[699,1178,1067,1367]" pageId="2" pageNumber="127">
Elytra wide, EL/EW =1.36 1.38, slightly wider than pronotum, EW/PW =1.09 1.16, almost parallel-sided in basal 3/4, widest behind middle; slightly sinuated before apices; apices rounded combined. Humeri rounded, without humeral tooth. Striae entire, deep, minutely punctate, much deeper apically and toward lateral margin. Intervals moderately convex, more ele- Male genitalia (
<figureCitation id="1355BEE1F1636B27403CFBD9FC15FB88" box="[878,949,1067,1091]" captionStart="Fig" captionStartId="3.[114,153,732,756]" captionTargetBox="[117,595,191,688]" captionTargetId="figure-285@3.[114,598,171,695]" captionTargetPageId="3" captionText="Fig. 2. Male genitalia ofCosmodiscus shavrini Anichtchenko,sp. nov. Scale bar 1mm. a genital ring; b median lobe of aedeagus; c left paramere; d right paramere. Scale bar 1mm." pageId="2" pageNumber="127">Fig. 2</figureCitation>
). Genital ring (
<figureCitation id="1355BEE1F1636B274738FBD9FD76FBAA" captionStart="Fig" captionStartId="3.[114,153,732,756]" captionTargetBox="[117,595,191,688]" captionTargetId="figure-285@3.[114,598,171,695]" captionTargetPageId="3" captionText="Fig. 2. Male genitalia ofCosmodiscus shavrini Anichtchenko,sp. nov. Scale bar 1mm. a genital ring; b median lobe of aedeagus; c left paramere; d right paramere. Scale bar 1mm." pageId="2" pageNumber="127">Fig. 2a</figureCitation>
) with short triangular process. Median lobe of aedeagus (Fig. 6b) in overall appearance similar with other species of the genus, i.e. the basal part is curved at a right angle slightly before the middle of its length, apex slightly curved down in lateral view, internal sac without sclerotized pieces. Right paramere elongate and narrow, left paramere large and rounded.
</paragraph>
</subSubSection>
<subSubSection id="C374F1EFF1636B2641E9FA8EFC17FE74" lastPageId="3" lastPageNumber="128" pageId="2" pageNumber="127" type="etymology">
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<emphasis id="B91A7E76F1636B2741E9FA8EFCE4FA5F" bold="true" box="[699,836,1404,1428]" pageId="2" pageNumber="127">Etymology.</emphasis>
Named after Dr. Alexey Shavrin, best friend and colleague, a specialist in
<taxonomicName id="4C6ED9E7F1636B2741E9FA4BFCF1FA1A" box="[699,849,1465,1489]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
, who caught this species during one of our joint expeditions in the
<collectingCountry id="F379E2F4F1636B2741E9FA05FC97F9C4" box="[699,823,1527,1551]" name="Philippines" pageId="2" pageNumber="127">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8BD1A264F1636B26473EF954FC17FE74" blockId="2.[1132,1172,1702,1726]" lastBlockId="3.[624,1103,178,447]" lastPageId="3" lastPageNumber="128" pageId="2" pageNumber="127">
127 president Roy G. Ponce; to the Research, Innovation, and Extension (RIE) Division of DOrSU; to Jose (SirToto) Letchoncito, Jr. for the assistance in acquiring our GP at DENR XI. To Nestor Badilla, ourlocal guide in Maragusan,
<collectingRegion id="49AA6C86F1626B264004FEB9FC3BFEA8" box="[854,923,331,355]" country="Philippines" name="Davao" pageId="3" pageNumber="128">Davao</collectingRegion>
de Oro. To our good friends who accompany us in the field, Dexter Patalita, and Alexey Shavrin to whom the species is named.
</paragraph>
</subSubSection>
<caption id="DF11F2ECF1626B264320FD2EFEFBFCA4" pageId="3" pageNumber="128" startId="3.[114,153,732,756]" targetBox="[117,595,191,688]" targetPageId="3" targetType="figure">
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Fig. 2. Male genitalia of
<taxonomicName id="4C6ED9E7F1626B26422EFD2EFEC3FCD9" pageId="3" pageNumber="128">
<emphasis id="B91A7E76F1626B26422EFD2EFF38FCD8" italics="true" pageId="3" pageNumber="128">Cosmodiscus shavrini</emphasis>
Anichtchenko,
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<emphasis id="B91A7E76F1626B264216FD08FEC3FCD9" bold="true" box="[324,355,762,786]" pageId="3" pageNumber="128">sp.</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="B91A7E76F1626B264239FD08FE3BFCD9" bold="true" box="[363,411,762,786]" pageId="3" pageNumber="128">nov.</emphasis>
Scale bar 1mm. a genital ring; b median lobe of aedeagus; c left paramere; d right paramere. Scale bar 1mm.
</paragraph>
</caption>
<subSubSection id="C374F1EFF1626B264320FC66FDE4FC67" box="[114,580,916,940]" pageId="3" pageNumber="128" type="distribution">
<paragraph id="8BD1A264F1626B264320FC66FDE4FC67" blockId="3.[114,580,916,940]" box="[114,580,916,940]" pageId="3" pageNumber="128">
<emphasis id="B91A7E76F1626B264320FC66FF5DFC67" bold="true" box="[114,253,916,940]" pageId="3" pageNumber="128">Distribution</emphasis>
. Known from Mindanao only.
</paragraph>
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</document>

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<emphasis id="B91A7E76F1626B264320FC23FE36FC22" bold="true" box="[114,406,977,1001]" italics="true" pageId="3" pageNumber="128">Cosmodiscus rufolimbatus</emphasis>
<bibRefCitation id="EFFFDF95F1626B2642CFFC23FD9EFC22" author="Jedlicka A." box="[413,574,977,1001]" pageId="3" pageNumber="128" refId="ref1857" refString="Jedlicka A. 1936. Cosmodiscusrufolimbatus sp. n. Casopis Ceske Spolecnosti Entomologicke 33: 103." type="journal volume" year="1936">Jedlička, 1936</bibRefCitation>
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<emphasis id="B91A7E76F1626B264320FBFDFF47FBEC" box="[114,231,1039,1063]" italics="true" pageId="3" pageNumber="128">louwerensi</emphasis>
Straneo, 1940: 215 (Saleier Is., S Celebes);
<bibRefCitation id="EFFFDF95F1626B2643AAFBDFFE4DFB8E" author="Darlington P. J." box="[248,493,1069,1093]" pageId="3" pageNumber="128" pagination="321 - 564" refId="ref1814" refString="Darlington P. J. 1962. The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini. Bulletin of the Museum of Comparative Zoology at Harvard College 126 (3): 321 - 564." type="journal article" year="1962">Darlington, 1962: 515</bibRefCitation>
, syn. by Fedorenko, 2021: 426.
</paragraph>
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<emphasis id="B91A7E76F1626B264320FB7BFF7EFB6A" bold="true" box="[114,222,1161,1185]" pageId="3" pageNumber="128">Material:</emphasis>
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1f
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,
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,
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, Banaue,
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<collectingDate id="EF947D4CF1626B264251FB5AFEFBFB0B" box="[259,347,1192,1216]" pageId="3" pageNumber="128" value="2016-06">VI.2016</collectingDate>
</date>
(
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)
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.
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<emphasis id="B91A7E76F1626B264320FB17FF4DFB36" bold="true" box="[114,237,1253,1277]" pageId="3" pageNumber="128">Comments</emphasis>
. Species described from Mindanao (
<bibRefCitation id="EFFFDF95F1626B26439AFAF6FE0DFAD7" author="Jedlicka A." box="[200,429,1284,1308]" pageId="3" pageNumber="128" refId="ref1857" refString="Jedlicka A. 1936. Cosmodiscusrufolimbatus sp. n. Casopis Ceske Spolecnosti Entomologicke 33: 103." type="journal volume" year="1936">Jedlička, 1936: 103</bibRefCitation>
), also known from Sulawesi. First record from Luzon.
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<mods:title id="F6030292CA9D9BE9842819BB8D138263">Baltic Journal of Coleopterology</mods:title>
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<mods:part id="31A0EFBAAADED8AC5B654F287AA8890F">
<mods:date id="2AF7D7FBA62A24491A3084586AE90EA3">2022</mods:date>
<mods:detail id="EFED60181E7B560E7C32334D511B3901" type="volume">
<mods:number id="C86A54886FBCAF9F11CA6E69306C3C47">22</mods:number>
</mods:detail>
<mods:detail id="B165F07DB3616AED4B594CF34E385121" type="issue">
<mods:number id="FCC86652C85200424B5979792521B1DB">2</mods:number>
</mods:detail>
<mods:extent id="D92FE410559DD815217EDD4A93639398" unit="page">
<mods:start id="14E639DF288F19A1654EC38ECD3EA239">421</mods:start>
<mods:end id="C7A65B9C010AAC7DC2DFD5D7326EFEDF">428</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AEEC3200994A50ACF0AC032343F0221B">journal article</mods:classification>
<mods:identifier id="F10192F77871E5DEB1BBB9A69D9C4C1F" type="ISSN">1407-8619</mods:identifier>
</mods:mods>
<treatment id="03C9879E8B00D04BFF277D3EFCB2A243" LSID="urn:lsid:plazi:treatment:03C9879E8B00D04BFF277D3EFCB2A243" httpUri="http://treatment.plazi.org/id/03C9879E8B00D04BFF277D3EFCB2A243" lastPageId="6" lastPageNumber="427" pageId="4" pageNumber="425">
<subSubSection id="C37A65038B00D049FF277D3EFE1CA179" box="[151,498,1548,1572]" pageId="4" pageNumber="425" type="nomenclature">
<paragraph id="8BDF36888B00D049FF277D3EFE1CA179" blockId="4.[151,498,1548,1604]" box="[151,498,1548,1572]" pageId="4" pageNumber="425">
<heading id="D09781E48B00D049FF277D3EFE1CA179" bold="true" box="[151,498,1548,1572]" fontSize="10" level="2" pageId="4" pageNumber="425" reason="4">
<emphasis id="B914EA9A8B00D049FF277D3EFE1CA179" bold="true" box="[151,498,1548,1572]" pageId="4" pageNumber="425">
<taxonomicName id="4C604D0B8B00D049FF277D3EFE78A179" ID-CoL="B7QSB" authorityName="Rukmane-Brbale" authorityYear="2022" box="[151,406,1548,1572]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FF277D3EFE78A179" bold="true" box="[151,406,1548,1572]" italics="true" pageId="4" pageNumber="425">Excalidiopsis bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FE2C7D3EFE1CA179" box="[412,498,1548,1572]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B00D049FF277D1FFEDCA119" box="[151,306,1581,1604]" pageId="4" pageNumber="425" type="description">
<paragraph id="8BDF36888B00D049FF277D1FFEDCA119" blockId="4.[151,498,1548,1604]" box="[151,306,1581,1604]" pageId="4" pageNumber="425">
(
<figureCitation id="135B2A0D8B00D049FF2F7D1FFF0EA119" box="[159,224,1581,1604]" captionStart="Figure 2" captionStartId="5.[113,183,1633,1656]" captionTargetBox="[117,1132,651,1599]" captionTargetId="figure-272@5.[113,1134,648,1600]" captionTargetPageId="5" captionText="Figure 2. Excalidiopsis bramanti sp. nov. A Holotype male, dorsal view. B paratype female, dorsal view." pageId="4" pageNumber="425">Fig. 2</figureCitation>
,
<figureCitation id="135B2A0D8B00D049FF5C7D1FFEC7A119" box="[236,297,1581,1604]" captionStart="Figure 3" captionStartId="6.[151,222,663,686]" captionTargetBox="[152,1171,152,628]" captionTargetId="figure-52@6.[151,1172,151,629]" captionTargetPageId="6" captionText="Figure 3. AC. Dorsaw view of head and antennae of selected species. A Excalidiopsis palawanensis sp nov. B Excalidiopsis bramanti sp. nov. C Calidiopsis bilineatus Rukmane, 2020; DE. Lateral view of selected species. D Excalidiopsis palawanensis sp. nov. E Excalidiopsis bramanti sp. nov." pageId="4" pageNumber="425">3B, E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B00D049FF277D51FBCDA553" pageId="4" pageNumber="425" type="materials_examined">
<paragraph id="8BDF36888B00D049FF277D51FD6CA1C6" blockId="4.[151,643,1635,1691]" pageId="4" pageNumber="425">
<emphasis id="B914EA9A8B00D049FF277D51FEABA126" bold="true" box="[151,325,1635,1659]" pageId="4" pageNumber="425">Type material.</emphasis>
<materialsCitation id="3B083CD58B00D049FEE07D51FD6CA1C6" collectingDate="2018-10" country="Philippines" location="Brokes Point" pageId="4" pageNumber="425" specimenCount="1" stateProvince="Palawan" typeStatus="holotype">
<emphasis id="B914EA9A8B00D049FEE07D51FDE0A126" bold="true" box="[336,526,1635,1659]" pageId="4" pageNumber="425">
<typeStatus id="54DB882A8B00D049FEE07D51FE54A126" box="[336,442,1635,1659]" pageId="4" pageNumber="425" type="holotype">Holotype</typeStatus>
. Male.
</emphasis>
<collectingCountry id="F37776188B00D049FD957D56FF0FA1C6" name="Philippines" pageId="4" pageNumber="425">PHILIPPINES</collectingCountry>
/
<collectingRegion id="49A4F86A8B00D049FF4C7DB6FEB8A1C6" box="[252,342,1668,1691]" country="Philippines" name="Palawan" pageId="4" pageNumber="425">Palawan</collectingRegion>
/
<location id="8EBF60538B00D049FEC17DB6FE10A1C6" LSID="urn:lsid:plazi:treatment:03C9879E8B00D04BFF277D3EFCB2A243:8EBF60538B00D049FEC17DB6FE10A1C6" box="[369,510,1668,1691]" country="Philippines" name="Brokes Point" pageId="4" pageNumber="425" stateProvince="Palawan">Brokes Point</location>
/
<date id="FFDE10488B00D049FDA97DB6FD9FA1C6" box="[537,625,1668,1691]" pageId="4" pageNumber="425" value="2018-10">
<collectingDate id="EF9AE9A08B00D049FDA97DB6FD9FA1C6" box="[537,625,1668,1691]" pageId="4" pageNumber="425" value="2018-10">X. 2018</collectingDate>
</date>
/
</materialsCitation>
</paragraph>
<paragraph id="8BDF36888B00D049FD187BA9FBEBA649" blockId="4.[680,1172,155,276]" pageId="4" pageNumber="425">
<materialsCitation id="3B083CD58B00D049FD187BA9FBEFA649" collectionCode="DUBC" pageId="4" pageNumber="425" specimenCount="1" typeStatus="holotype">
local collector leg.” (white rectangular label, printed) // “
<typeStatus id="54DB882A8B00D049FCAE7B8EFC43A78E" box="[798,941,188,211]" pageId="4" pageNumber="425" type="holotype">HOLOTYPE</typeStatus>
/ Male /
<taxonomicName id="4C604D0B8B00D049FBB57B8EFCE4A7AE" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="bramanti">
<emphasis id="B914EA9A8B00D049FBB57B8EFCE4A7AE" italics="true" pageId="4" pageNumber="425">Excalidiopsis bramanti</emphasis>
</taxonomicName>
Rukmane­Brbale, 2022” (red rectangular label, printed) (
<collectionCode id="ED71AE4D8B00D049FC197BCFFC17A649" box="[937,1017,253,276]" pageId="4" pageNumber="425">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF36888B00D049FD187A06FBCDA553" blockId="4.[680,1172,308,527]" pageId="4" pageNumber="425">
<materialsCitation id="3B083CD58B00D049FD187A06FBCEA553" collectingDate="2018-09" collectingDateMax="2018-10" collectingDateMin="2018-09" collectionCode="DUBC" country="Philippines" location="With" pageId="4" pageNumber="425" specimenCount="2" stateProvince="Palawan" typeStatus="paratype">
<emphasis id="B914EA9A8B00D049FD187A06FCF8A611" bold="true" box="[680,790,308,332]" pageId="4" pageNumber="425">
<typeStatus id="54DB882A8B00D049FD187A06FCF8A611" box="[680,790,308,332]" pageId="4" pageNumber="425" type="paratype">Paratypes</typeStatus>
</emphasis>
(3, 2): “
<collectingCountry id="F37776188B00D049FC367A06FBCEA616" box="[902,1056,308,331]" name="Philippines" pageId="4" pageNumber="425">PHILIPPINES</collectingCountry>
/
<collectingRegion id="49A4F86A8B00D049FB9F7A06FB68A616" box="[1071,1158,308,331]" country="Philippines" name="Palawan" pageId="4" pageNumber="425">Palawan</collectingRegion>
/ Brokes Point /
<date id="FFDE10488B00D049FCFA7A67FC44A631" box="[842,938,341,364]" pageId="4" pageNumber="425" value="2018-09">
<collectingDate id="EF9AE9A08B00D049FCFA7A67FC44A631" box="[842,938,341,364]" pageId="4" pageNumber="425" value="2018-09">IX. 2018</collectingDate>
</date>
/ local collector leg.” (2, 2) // “
<collectingCountry id="F37776188B00D049FC987A47FC26A6D1" box="[808,968,373,396]" name="Philippines" pageId="4" pageNumber="425">PHILIPPINES</collectingCountry>
/
<collectingRegion id="49A4F86A8B00D049FC6B7A47FBDBA6D1" box="[987,1077,373,396]" country="Philippines" name="Palawan" pageId="4" pageNumber="425">Palawan</collectingRegion>
/ Brokes Point /
<date id="FFDE10488B00D049FD5F7AA7FCAFA6F1" box="[751,833,405,428]" pageId="4" pageNumber="425" value="2018-10">
<collectingDate id="EF9AE9A08B00D049FD5F7AA7FCAFA6F1" box="[751,833,405,428]" pageId="4" pageNumber="425" value="2018-10">X. 2018</collectingDate>
</date>
/ local collector leg.” (1) (white rectangular labels, printed).
<location id="8EBF60538B00D049FC7B7A84FC11A690" LSID="urn:lsid:plazi:treatment:03C9879E8B00D04BFF277D3EFCB2A243:8EBF60538B00D049FC7B7A84FC11A690" box="[971,1023,438,461]" country="Philippines" name="With" pageId="4" pageNumber="425" stateProvince="Palawan">With</location>
additional red label: “
<typeStatus id="54DB882A8B00D049FD467AE5FC93A6B3" box="[758,893,471,494]" pageId="4" pageNumber="425" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C604D0B8B00D049FC287AE5FB7AA6B3" authorityName="Rukmane-Bārbale" authorityYear="2022" box="[920,1172,471,494]" class="Insecta" family="Curculionidae" genus="Exlacidiopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="bramanti">
<emphasis id="B914EA9A8B00D049FC287AE5FB7AA6B3" box="[920,1172,471,494]" italics="true" pageId="4" pageNumber="425">Exlacidiopsis bramanti</emphasis>
</taxonomicName>
Rukmane­Brbale, 2022” (
<collectionCode id="ED71AE4D8B00D049FC787AC5FBF6A553" box="[968,1048,503,526]" pageId="4" pageNumber="425">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B00D04BFD18791CFCAAA4E7" lastPageId="6" lastPageNumber="427" pageId="4" pageNumber="425" type="description">
<paragraph id="8BDF36888B00D049FD18791CFCEFA417" blockId="4.[680,1173,558,842]" pageId="4" pageNumber="425">
<emphasis id="B914EA9A8B00D049FD18791CFCDFA51B" bold="true" box="[680,817,558,582]" pageId="4" pageNumber="425">Description.</emphasis>
Male. Measurements: LB: 7.4­ 8.3 (
<typeStatus id="54DB882A8B00D049FD00797DFCE1A53B" box="[688,783,591,614]" pageId="4" pageNumber="425" type="holotype">holotype</typeStatus>
7.8, mean 7.87); LP: 2.0­ 2.7 (
<typeStatus id="54DB882A8B00D049FBEB797DFD38A5DB" pageId="4" pageNumber="425" type="holotype">holotype</typeStatus>
2.2, mean 2.3); LE: 5.4­ 5.7 (
<typeStatus id="54DB882A8B00D049FBB7795DFB88A5DB" box="[1031,1126,623,646]" pageId="4" pageNumber="425" type="holotype">holotype</typeStatus>
5.6, mean 5.57); LR: 1.4­ 1.5 (
<typeStatus id="54DB882A8B00D049FC7179A2FBCEA5FA" box="[961,1056,656,679]" pageId="4" pageNumber="425" type="holotype">holotype</typeStatus>
1.4, mean 1.42); WP: 2.1­ 2.2 (
<typeStatus id="54DB882A8B00D049FC357982FC0AA59A" box="[901,996,688,711]" pageId="4" pageNumber="425" type="holotype">holotype</typeStatus>
2.1, mean 2.1); WE: 4.1­ 4.4 (
<typeStatus id="54DB882A8B00D049FC8E79E3FC73A5B5" box="[830,925,721,744]" pageId="4" pageNumber="425" type="holotype">holotype</typeStatus>
4.4, mean 4.22); WR: 1.1­1.2 (
<typeStatus id="54DB882A8B00D049FCBA79C0FC84A454" box="[778,874,754,777]" pageId="4" pageNumber="425" type="holotype">holotype</typeStatus>
1.2, mean 1.17). N=4 for all measurements. Dorsal habitus as shown in
<figureCitation id="135B2A0D8B00D049FD187801FD13A417" box="[680,765,819,842]" captionStart="Figure 2" captionStartId="5.[113,183,1633,1656]" captionTargetBox="[117,1132,651,1599]" captionTargetId="figure-272@5.[113,1134,648,1600]" captionTargetPageId="5" captionText="Figure 2. Excalidiopsis bramanti sp. nov. A Holotype male, dorsal view. B paratype female, dorsal view." pageId="4" pageNumber="425">Fig. 2A</figureCitation>
.
</paragraph>
<paragraph id="8BDF36888B00D049FD187858FD1CA35E" blockId="4.[680,1172,874,1027]" pageId="4" pageNumber="425">Body rather big, with markings of shiny green scales of round to eliptic shape; prothorax and elytra strongly rugose, head punctured at apical part, rostrum and legs strongly punctured in all length.</paragraph>
<paragraph id="8BDF36888B00D049FD187F11FBC9A17F" blockId="4.[680,1172,1059,1570]" pageId="4" pageNumber="425">Forehead dorsally with longitudinal medial groove, separated from rostrum by sulcus at medial portion between eyes; eyes small, unprominent from outline of the head dorsally, widest just in the middle; rostrum longer than wide, LR/WR: 1.17; marked with round scales dorsally at basal half; eliptic shape scales at lateral parts along apical half; mingled with short light hairs in all length; antennal scrobes with dorsal furrow less sharp, wide V­shaped, peak rounded, just before the middle. Antennal scape without green scales in all length; segments of funicle I to VII subequal in length except segment II, segment II 1.5 times longer than rest, marked with round green scales; club slightly longer than wide, divided into three equal parts.</paragraph>
<paragraph id="8BDF36888B00D048FD187D70FDC7A66F" blockId="4.[680,1172,1602,1690]" lastBlockId="5.[113,605,155,306]" lastPageId="5" lastPageNumber="426" pageId="4" pageNumber="425">Prothorax nearly same length and width, LP/ WP: 1.05; with the following markings of shiny green scales: 1) longidutinal medial line medially at disc in all length; 2) narrow longitudinal line at each dorso­lateral part in all length; 3) ventral part sparsely covered with round scales; dorsal contour widest just before the middle, in lateral contour slightly bulged.</paragraph>
<paragraph id="8BDF36888B01D048FFC17A63FD03A6CF" blockId="5.[113,606,337,584]" lastBlockId="5.[643,1134,155,402]" pageId="5" pageNumber="426">Elytra ovate, bulged, longer than wide: LE/WE: 1.27; apex more or less rounded; finer granulation, intervals less pronounced; each elytron with the following markings of shiny green scales: 1) three elongated patches at basal part, two from subbasal to basal 1/3, one along interval III, second redirected laterally and one just before the middle along intervals IV to V; 2) one medial patch along interval VII; 3) four patches at apical part, two at apicl 1/2, one along interval III and one along interval V, two more patches along interval III, one at subapical part, one before apex; 4) two sutural patches, one just before the middle, one at apical 3/4; 5) three small patches along lateral margin scattered in all length.</paragraph>
<paragraph id="8BDF36888B01D048FD337A83FBA7A515" blockId="5.[643,1135,433,584]" pageId="5" pageNumber="426">
Female. Measurements: LB: 8.3­ 8.4 (mean 8.35); LP: 2.5; LW: 5.8­ 5.9 (mean 5.85); LR: 1.5; WP: 2.3­ 2.4 (mean 2.35); WE: 4.9­ 5.0 (mean 4.95); WR: 1.3­ 1.4 (mean 1.35). N=2 for all measurements. Dorsal habitus as shown in
<figureCitation id="135B2A0D8B01D048FC437903FBABA515" box="[1011,1093,561,584]" captionStart="Figure 2" captionStartId="5.[113,183,1633,1656]" captionTargetBox="[117,1132,651,1599]" captionTargetId="figure-272@5.[113,1134,648,1600]" captionTargetPageId="5" captionText="Figure 2. Excalidiopsis bramanti sp. nov. A Holotype male, dorsal view. B paratype female, dorsal view." pageId="5" pageNumber="426">Fig. 2B</figureCitation>
.
</paragraph>
<caption id="DF1F66008B01D048FFC17D53FF1CA1C5" pageId="5" pageNumber="426" startId="5.[113,183,1633,1656]" targetBox="[117,1132,651,1599]" targetPageId="5" targetType="figure">
<paragraph id="8BDF36888B01D048FFC17D53FF1CA1C5" blockId="5.[113,1134,1632,1688]" pageId="5" pageNumber="426">
Figure 2.
<taxonomicName id="4C604D0B8B01D048FF687D53FE3FA125" authorityName="Rukmane-Brbale" authorityYear="2022" box="[216,465,1633,1656]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B01D048FF687D53FE3FA125" box="[216,465,1633,1656]" italics="true" pageId="5" pageNumber="426">Excalidiopsis bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B01D048FE677D53FDC4A125" box="[471,554,1633,1656]" pageId="5" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B914EA9A8B01D048FD807D52FDADA125" bold="true" box="[560,579,1632,1656]" pageId="5" pageNumber="426">A</emphasis>
Holotype male, dorsal view.
<emphasis id="B914EA9A8B01D048FC207D52FC4CA125" bold="true" box="[912,930,1632,1656]" pageId="5" pageNumber="426">B</emphasis>
paratype female, dorsal view.
</paragraph>
</caption>
<caption id="DF1F66008B02D04BFF2779A5FE06A453" pageId="6" pageNumber="427" startId="6.[151,222,663,686]" targetBox="[152,1171,152,628]" targetPageId="6" targetType="figure">
<paragraph id="8BDF36888B02D04BFF2779A5FE06A453" blockId="6.[151,1172,662,782]" pageId="6" pageNumber="427">
Figure 3.
<emphasis id="B914EA9A8B02D04BFEB879A4FED2A5F3" bold="true" box="[264,316,662,686]" pageId="6" pageNumber="427">AC</emphasis>
. Dorsaw view of head and antennae of selected species.
<emphasis id="B914EA9A8B02D04BFC7B79A4FC30A5F3" bold="true" box="[971,990,662,686]" pageId="6" pageNumber="427">A</emphasis>
<taxonomicName id="4C604D0B8B02D04BFBB079A5FEC6A593" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="B914EA9A8B02D04BFBB079A5FEC6A593" italics="true" pageId="6" pageNumber="427">Excalidiopsis palawanensis</emphasis>
</taxonomicName>
sp nov.
<emphasis id="B914EA9A8B02D04BFE327984FE7AA593" bold="true" box="[386,404,694,718]" pageId="6" pageNumber="427">B</emphasis>
<taxonomicName id="4C604D0B8B02D04BFE1F7985FD46A593" authorityName="Rukmane-Brbale" authorityYear="2022" box="[431,680,695,718]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B02D04BFE1F7985FD46A593" box="[431,680,695,718]" italics="true" pageId="6" pageNumber="427">Excalidiopsis bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B02D04BFD1F7985FCEFA593" box="[687,769,695,718]" pageId="6" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B914EA9A8B02D04BFCB87984FCF5A593" bold="true" box="[776,795,694,718]" pageId="6" pageNumber="427">C</emphasis>
<taxonomicName id="4C604D0B8B02D04BFC867985FF3EA5B3" authority="Rukmane, 2020" authorityName="Rukmane" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="bilineatus">
<emphasis id="B914EA9A8B02D04BFC867985FBCEA593" box="[822,1056,695,718]" italics="true" pageId="6" pageNumber="427">Calidiopsis bilineatus</emphasis>
Rukmane, 2020
</taxonomicName>
;
<emphasis id="B914EA9A8B02D04BFF6F79E4FEFCA5B3" bold="true" box="[223,274,726,750]" pageId="6" pageNumber="427">DE</emphasis>
. Lateral view of selected species.
<emphasis id="B914EA9A8B02D04BFD2A79E4FD43A5B3" bold="true" box="[666,685,726,750]" pageId="6" pageNumber="427">D</emphasis>
<taxonomicName id="4C604D0B8B02D04BFD7F79E5FC11A5B3" authorityName="Rukmane-Brbale" authorityYear="2022" box="[719,1023,727,750]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B02D04BFD7F79E5FC11A5B3" box="[719,1023,727,750]" italics="true" pageId="6" pageNumber="427">Excalidiopsis palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B02D04BFBB979E5FB8EA5B3" box="[1033,1120,727,750]" pageId="6" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B914EA9A8B02D04BFBDA79E4FB92A5B3" bold="true" box="[1130,1148,726,750]" pageId="6" pageNumber="427">E</emphasis>
<taxonomicName id="4C604D0B8B02D04BFF2779C5FE7EA453" authorityName="Rukmane-Brbale" authorityYear="2022" box="[151,400,759,782]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B02D04BFF2779C5FE7EA453" box="[151,400,759,782]" italics="true" pageId="6" pageNumber="427">Excalidiopsis bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B02D04BFE2679C5FE06A453" box="[406,488,759,782]" pageId="6" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BDF36888B02D04BFD187871FCAAA4E7" blockId="6.[680,1172,835,954]" pageId="6" pageNumber="427">
<emphasis id="B914EA9A8B02D04BFD187871FC65A406" bold="true" box="[680,907,835,859]" pageId="6" pageNumber="427">Differential analyses.</emphasis>
The new species is similar to
<taxonomicName id="4C604D0B8B02D04BFD717851FC87A427" authorityName="Rukmane-Brbale" authorityYear="2022" box="[705,873,867,890]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B02D04BFD717851FC87A427" box="[705,873,867,890]" italics="true" pageId="6" pageNumber="427">E. palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B02D04BFCDE7851FC55A427" box="[878,955,867,890]" pageId="6" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
, differences between species are lister in description of
<taxonomicName id="4C604D0B8B02D04BFBAE78B1FD05A4E7" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B02D04BFBAE78B1FD05A4E7" italics="true" pageId="6" pageNumber="427">E. palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B02D04BFD427891FCAAA4E7" box="[754,836,931,954]" pageId="6" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B02D04BFD1878EBFC07A4AC" box="[680,1001,985,1009]" pageId="6" pageNumber="427" type="distribution">
<paragraph id="8BDF36888B02D04BFD1878EBFC07A4AC" blockId="6.[680,1001,985,1009]" box="[680,1001,985,1009]" pageId="6" pageNumber="427">
<emphasis id="B914EA9A8B02D04BFD1878EBFCD5A4AC" bold="true" box="[680,827,985,1009]" pageId="6" pageNumber="427">Distribution.</emphasis>
<collectingRegion id="49A4F86A8B02D04BFCF078E8FC74A4AC" box="[832,922,986,1009]" country="Philippines" name="Palawan" pageId="6" pageNumber="427">Palawan</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B02D04BFD187F22FCB2A243" pageId="6" pageNumber="427" type="etymology">
<paragraph id="8BDF36888B02D04BFD187F22FB7BA3FA" blockId="6.[680,1173,1040,1191]" pageId="6" pageNumber="427">
<emphasis id="B914EA9A8B02D04BFD187F22FCC4A375" bold="true" box="[680,810,1040,1064]" pageId="6" pageNumber="427">Etymology.</emphasis>
The new species is named after enthusiastic entomologists, brothers
<taxonomicName id="4C604D0B8B02D04BFB807F02FBA4A33A" authority="Alessandro and Andrea (Lucca, Itally)" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="bramanti">Bramanti Alessandro and Andrea (Lucca, Itally)</taxonomicName>
in appreciation of their contribution to Curculionid taxonomy, especially to fauna of
<collectingRegion id="49A4F86A8B02D04BFC437FA2FBA5A3FA" box="[1011,1099,1168,1191]" country="Philippines" name="Palawan" pageId="6" pageNumber="427">Palawan</collectingRegion>
Island.
</paragraph>
<paragraph id="8BDF36888B02D04BFD187FF5FCB2A243" blockId="6.[680,1172,1223,1310]" pageId="6" pageNumber="427">
<emphasis id="B914EA9A8B02D04BFD187FF5FCFCA382" bold="true" box="[680,786,1223,1247]" pageId="6" pageNumber="427">Mimicry.</emphasis>
The species consist in mimetic relationship with
<taxonomicName id="4C604D0B8B02D04BFCFA7FD5FCB6A243" authority="Yoshitake, 2013" authorityName="Yoshitake" authorityYear="2013" class="Insecta" family="Curculionidae" genus="Expachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B914EA9A8B02D04BFCFA7FD5FB7AA3A3" box="[842,1172,1255,1278]" italics="true" pageId="6" pageNumber="427">Expachyrhynchus granulatus</emphasis>
<bibRefCitation id="EFF14B798B02D04BFD187E35FCB6A243" author="Yoshitake H." box="[680,856,1287,1310]" pageId="6" pageNumber="427" pagination="1 - 8" refId="ref3835" refString="Yoshitake H. 2013. A new genus and two new species of the tribe Pachyrhynchini (Coleoptera: Curculionidae) from Palawan Island, the Philippines. Esakia, 53: 1 - 8." type="journal article" year="2013">Yoshitake, 2013</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,148 @@
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<mods:title id="6EE9021BC69671EDEA1699618F31E752">Anew genus and two new species of the tribe Celeuthetini Lacordaire, 1863 (Coleoptera: Curculionidae) from the Philippines</mods:title>
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<treatment id="03C9879E8B05D04FFD337BE7FD6DA383" ID-DOI="http://doi.org/10.5281/zenodo.13199633" ID-Zenodo-Dep="13199633" LSID="urn:lsid:plazi:treatment:03C9879E8B05D04FFD337BE7FD6DA383" httpUri="http://treatment.plazi.org/id/03C9879E8B05D04FFD337BE7FD6DA383" lastPageId="2" lastPageNumber="423" pageId="1" pageNumber="422">
<subSubSection id="C37A65038B05D04CFD337BE7FC6EA7B1" box="[643,896,212,237]" pageId="1" pageNumber="422" type="nomenclature">
<paragraph id="8BDF36888B05D04CFD337BE7FC6EA7B1" blockId="1.[643,896,212,237]" box="[643,896,212,237]" pageId="1" pageNumber="422">
<heading id="D09781E48B05D04CFD337BE7FC6EA7B1" bold="true" box="[643,896,212,237]" fontSize="10" level="2" pageId="1" pageNumber="422" reason="4">
<emphasis id="B914EA9A8B05D04CFD337BE7FC6EA7B1" bold="true" box="[643,896,212,237]" pageId="1" pageNumber="422">
<taxonomicName id="4C604D0B8B05D04CFD337BE7FCF8A7B0" ID-CoL="B7QMY" authorityName="Rukmane-Brbale" authorityYear="2022" box="[643,790,213,237]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914EA9A8B05D04CFD337BE7FCF8A7B0" bold="true" box="[643,790,213,237]" italics="true" pageId="1" pageNumber="422">Excalidiopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B05D04CFCAC7BE6FC6EA7B1" box="[796,896,212,236]" pageId="1" pageNumber="422" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B05D04CFD337A39FD41A61E" pageId="1" pageNumber="422" type="type_taxon">
<paragraph id="8BDF36888B05D04CFD337A39FD41A61E" blockId="1.[643,1134,267,323]" pageId="1" pageNumber="422">
<emphasis id="B914EA9A8B05D04CFD337A39FCF7A67E" bold="true" box="[643,793,267,291]" pageId="1" pageNumber="422">
<typeStatus id="54DB882A8B05D04CFD337A39FD55A67E" box="[643,699,267,291]" pageId="1" pageNumber="422">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C604D0B8B05D04CFC907A3EFBA6A67E" ID-CoL="B7QL9" authorityName="Rukmane-Brbale" authorityYear="2022" box="[800,1096,268,291]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B05D04CFC907A3EFBA6A67E" box="[800,1096,268,291]" italics="true" pageId="1" pageNumber="422">Excalidiopsis palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B05D04CFBFF7A3EFD41A61E" pageId="1" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B05D04FFD337A51FE38A30D" lastPageId="2" lastPageNumber="423" pageId="1" pageNumber="422" type="description">
<paragraph id="8BDF36888B05D04CFD337A51FB80A561" blockId="1.[643,1134,355,572]" pageId="1" pageNumber="422">
<emphasis id="B914EA9A8B05D04CFD337A51FCE0A626" bold="true" box="[643,782,355,379]" pageId="1" pageNumber="422">Description.</emphasis>
Body medium sized, males and females of nearly same size, last beeing bigger, with wider elytra; prothorax and elytra strongly rugose except smooth underside; head smooth, rostrum and legs granulated; body covered with round to elliptic bright, shiny scales of various colour forming scally spots, lines and patches.
</paragraph>
<paragraph id="8BDF36888B05D04CFD33796EFBB7A34B" blockId="1.[643,1134,604,1046]" pageId="1" pageNumber="422">Forehead straight, separated from rostrum by sulcus; eyes small, slightly prominent to unprominent from the outline of the head; rostrum longer than wide; dorsal surface with longitudinal medial groove from medial part to base; frontal part straight to slightly widened apically, ventral part widened apically; dorsal contour of forehaed and rostrum decreased from widened apex; lateral contour more or less straigth; antennal scrobes dorso­lateral position, open posteriorly, narrow triangular, dorsal furrow more or less sharp, wide V­shaped, with peak before the middle, ventral furrow curved downvards, rised to apex, end just before inner margin of eye.</paragraph>
<paragraph id="8BDF36888B05D04CFD337F04FD0AA272" blockId="1.[643,1134,1078,1327]" pageId="1" pageNumber="422">Antennae with scape reaching medial part of prothorax; scape flattened dorsally, widened apically, nearly same length as funicle, mingled with short light hairs; funicle mingld with longer light hairs, club short, more or less equal in length and width; segments I to VII subequal in size, with II beeing same to 1.5 times longer than rest.</paragraph>
<paragraph id="8BDF36888B05D04CFD337E7CFBDFA2BA" blockId="1.[643,1134,1358,1511]" pageId="1" pageNumber="422">Pronotum with rugose puncture, mingled with short hairs, with longitudinal lines of glossy scales; dorsal contour nearly straigth, slightly widened at medial part, lateral contour straight to slighty convex before the medial part.</paragraph>
<paragraph id="8BDF36888B05D04CFD337D34FC4EA163" blockId="1.[643,1134,1542,1598]" pageId="1" pageNumber="422">Coxa and coxal cavities connate. Hind tibiae with semi­enclosed corbel.</paragraph>
<paragraph id="8BDF36888B05D04FFD337D6CFDF2A7AE" blockId="1.[643,1134,1630,1685]" lastBlockId="2.[151,643,155,243]" lastPageId="2" lastPageNumber="423" pageId="1" pageNumber="422">Elytra lanceolate, convex dorsally, widest along middle, apex extended; with well expressed intervals of strongly rugose puncture rows; each elytron with seven to ten scally patches and two more sutural patches of shiny scales.</paragraph>
<paragraph id="8BDF36888B06D04FFF277A20FE38A30D" blockId="2.[151,643,274,1104]" pageId="2" pageNumber="423">
<emphasis id="B914EA9A8B06D04FFF277A20FE62A677" bold="true" box="[151,396,274,298]" pageId="2" pageNumber="423">Differential analyses.</emphasis>
Among 11 Philippine distributed genera of the tribe
<taxonomicName id="4C604D0B8B06D04FFE707A01FDD1A617" authorityName="Lacordaire" authorityYear="1863" box="[448,575,307,330]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="tribe" tribe="Celeuthetini">Celeuthetini</taxonomicName>
,
<taxonomicName id="4C604D0B8B06D04FFDF77A01FF1CA637" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914EA9A8B06D04FFDF77A01FF1CA637" italics="true" pageId="2" pageNumber="423">Excalidiopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B06D04FFF4C7A61FE8FA637" box="[252,353,339,362]" pageId="2" pageNumber="423" rank="genus">gen. nov.</taxonomicNameLabel>
is most closely related to
<taxonomicName id="4C604D0B8B06D04FFF277A41FE7AA6D7" authority="Heller, 1913" authorityName="Heller" authorityYear="1913" box="[151,404,371,394]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B914EA9A8B06D04FFF277A41FEE0A6D7" box="[151,270,371,394]" italics="true" pageId="2" pageNumber="423">Calidiopsis</emphasis>
Heller, 1913
</taxonomicName>
by the following features: antennal scape long, reaching medial to hind part of prothorax, antennal scape of nearly same length as funicle, prothorax and elytra strongly rugose, prothorax small, slender, nearly straigth dorsally, longer than wide, elytra with strongly extended apex dorsally. The new genus can be distinguished from
<taxonomicName id="4C604D0B8B06D04FFEEF7967FE38A531" authorityName="Heller" authorityYear="1913" box="[351,470,597,620]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B914EA9A8B06D04FFEEF7967FE38A531" box="[351,470,597,620]" italics="true" pageId="2" pageNumber="423">Calidiopsis</emphasis>
</taxonomicName>
by the following characters: 1) large, strongly prominent eyes in
<taxonomicName id="4C604D0B8B06D04FFF2779A4FEE5A5F0" authorityName="Heller" authorityYear="1913" box="[151,267,662,685]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B914EA9A8B06D04FFF2779A4FEE5A5F0" box="[151,267,662,685]" italics="true" pageId="2" pageNumber="423">Calidiopsis</emphasis>
</taxonomicName>
, while small, unprominent or slightly prominent in
<taxonomicName id="4C604D0B8B06D04FFE947984FE5AA590" authorityName="Rukmane-Brbale" authorityYear="2022" box="[292,436,694,717]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914EA9A8B06D04FFE947984FE5AA590" box="[292,436,694,717]" italics="true" pageId="2" pageNumber="423">Excalidiopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B06D04FFE087984FDF8A590" box="[440,534,694,717]" pageId="2" pageNumber="423" rank="genus">gen. nov.</taxonomicNameLabel>
; 2) antennal scape of
<taxonomicName id="4C604D0B8B06D04FFEA579E4FE62A5B0" authorityName="Heller" authorityYear="1913" box="[277,396,726,749]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B914EA9A8B06D04FFEA579E4FE62A5B0" box="[277,396,726,749]" italics="true" pageId="2" pageNumber="423">Calidiopsis</emphasis>
</taxonomicName>
covered with very long, dark hairs while hairs on antennal scape of
<taxonomicName id="4C604D0B8B06D04FFDF679C5FF1CA473" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914EA9A8B06D04FFDF679C5FF1CA473" italics="true" pageId="2" pageNumber="423">Excalidiopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B06D04FFF487825FEB6A473" box="[248,344,791,814]" pageId="2" pageNumber="423" rank="genus">gen. nov.</taxonomicNameLabel>
short; 3) I and II segment of funicle differ in length from segments III to VII in
<taxonomicName id="4C604D0B8B06D04FFF037865FEC3A433" authorityName="Heller" authorityYear="1913" box="[179,301,855,878]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B914EA9A8B06D04FFF037865FEC3A433" box="[179,301,855,878]" italics="true" pageId="2" pageNumber="423">Calidiopsis</emphasis>
</taxonomicName>
, I beeing from 1.5 times longer and III beeing up to three times longer than segments III to VII, segments of
<taxonomicName id="4C604D0B8B06D04FFE7478AAFDBCA4F2" authorityName="Rukmane-Brbale" authorityYear="2022" box="[452,594,920,943]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914EA9A8B06D04FFE7478AAFDBCA4F2" box="[452,594,920,943]" italics="true" pageId="2" pageNumber="423">Excalidiopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B06D04FFDE678AAFF2BA492" pageId="2" pageNumber="423" rank="genus">gen. nov.</taxonomicNameLabel>
subequal in size, with II being same or slightly longer than I to VII; 4) club of
<taxonomicName id="4C604D0B8B06D04FFD9278EAFF20A34D" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914EA9A8B06D04FFD9278EAFF20A34D" italics="true" pageId="2" pageNumber="423">Excalidiopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B06D04FFF6778CAFED4A352" box="[215,314,1016,1039]" pageId="2" pageNumber="423" rank="genus">gen. nov.</taxonomicNameLabel>
short, nearly same length and width while club of
<taxonomicName id="4C604D0B8B06D04FFEC57F2BFE01A36D" authorityName="Heller" authorityYear="1913" box="[373,495,1049,1072]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B914EA9A8B06D04FFEC57F2BFE01A36D" box="[373,495,1049,1072]" italics="true" pageId="2" pageNumber="423">Calidiopsis</emphasis>
</taxonomicName>
long, at least twice as long as wide (
<figureCitation id="135B2A0D8B06D04FFE387F0BFE27A30D" box="[392,457,1081,1104]" captionStart="Figure 3" captionStartId="6.[151,222,663,686]" captionTargetBox="[152,1171,152,628]" captionTargetId="figure-52@6.[151,1172,151,629]" captionTargetPageId="6" captionText="Figure 3. AC. Dorsaw view of head and antennae of selected species. A Excalidiopsis palawanensis sp nov. B Excalidiopsis bramanti sp. nov. C Calidiopsis bilineatus Rukmane, 2020; DE. Lateral view of selected species. D Excalidiopsis palawanensis sp. nov. E Excalidiopsis bramanti sp. nov." pageId="2" pageNumber="423">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B06D04FFF277F42FDB3A3DA" box="[151,605,1136,1160]" pageId="2" pageNumber="423" type="distribution">
<paragraph id="8BDF36888B06D04FFF277F42FDB3A3DA" blockId="2.[151,605,1136,1160]" box="[151,605,1136,1160]" pageId="2" pageNumber="423">
<emphasis id="B914EA9A8B06D04FFF277F42FEC4A3D5" bold="true" box="[151,298,1136,1160]" pageId="2" pageNumber="423">Distribution.</emphasis>
<collectingCountry id="F37776188B06D04FFE9F7F42FE44A3DA" box="[303,426,1136,1159]" name="Philippines" pageId="2" pageNumber="423">Philippines</collectingCountry>
,
<collectingRegion id="49A4F86A8B06D04FFE047F42FDE0A3DA" box="[436,526,1136,1159]" country="Philippines" name="Palawan" pageId="2" pageNumber="423">Palawan</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B06D04FFF277F94FD6DA383" pageId="2" pageNumber="423" type="etymology">
<paragraph id="8BDF36888B06D04FFF277F94FD6DA383" blockId="2.[151,643,1190,1246]" pageId="2" pageNumber="423">
<emphasis id="B914EA9A8B06D04FFF277F94FEF6A3E3" bold="true" box="[151,280,1190,1214]" pageId="2" pageNumber="423">Etymology.</emphasis>
The generic name is derived from similarity with closely related genus
<taxonomicName id="4C604D0B8B06D04FFDB77FF5FD95A383" authorityName="Heller" authorityYear="1913" box="[519,635,1223,1246]" class="Insecta" family="Curculionidae" genus="Calidiopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B914EA9A8B06D04FFDB77FF5FD95A383" box="[519,635,1223,1246]" italics="true" pageId="2" pageNumber="423">Calidiopsis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,288 @@
<document id="D72F19FADE43C22DA70D9676738AA8EA" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722656919139" checkinUser="felipe" docAuthor="Rukmane-B rbale, Anita" docDate="2022" docId="03C9879E8B06D049FF277E27FF3BA288" docLanguage="en" docName="BalticJColeopterol.22.2.421-428.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Excalidiopsis palawanensis rbale, 2022, sp. nov." docType="treatment" docVersion="1" lastPageNumber="425" masterDocId="FFF0FFE68B04D04DFFB07B32FFEEA75D" masterDocTitle="Anew genus and two new species of the tribe Celeuthetini Lacordaire, 1863 (Coleoptera: Curculionidae) from the Philippines" masterLastPageNumber="428" masterPageNumber="421" pageNumber="423" updateTime="1722690729176" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="66D04BF642FC413869BA92ED8C3508EB">Anew genus and two new species of the tribe Celeuthetini Lacordaire, 1863 (Coleoptera: Curculionidae) from the Philippines</mods:title>
</mods:titleInfo>
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<mods:namePart id="99AD49904E08EC9A8AE98DCD10C09729">Rukmane-B rbale, Anita</mods:namePart>
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<mods:part id="980B1CF939AC7678B6F55CC9A86407F4">
<mods:date id="BD59F91F0EF3AAF83DF3C3F197E80449">2022</mods:date>
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</mods:mods>
<treatment id="03C9879E8B06D049FF277E27FF3BA288" LSID="urn:lsid:plazi:treatment:03C9879E8B06D049FF277E27FF3BA288" httpUri="http://treatment.plazi.org/id/03C9879E8B06D049FF277E27FF3BA288" lastPageId="4" lastPageNumber="425" pageId="2" pageNumber="423">
<subSubSection id="C37A65038B06D04FFF277E27FDCFA271" box="[151,545,1300,1325]" pageId="2" pageNumber="423" type="nomenclature">
<paragraph id="8BDF36888B06D04FFF277E27FDCFA271" blockId="2.[151,545,1300,1356]" box="[151,545,1300,1325]" pageId="2" pageNumber="423">
<heading id="D09781E48B06D04FFF277E27FDCFA271" bold="true" box="[151,545,1300,1325]" fontSize="10" level="2" pageId="2" pageNumber="423" reason="4">
<emphasis id="B914EA9A8B06D04FFF277E27FDCFA271" bold="true" box="[151,545,1300,1325]" pageId="2" pageNumber="423">
<taxonomicName id="4C604D0B8B06D04FFF277E27FE2BA270" ID-CoL="B7QL9" authorityName="Rukmane-Brbale" authorityYear="2022" box="[151,453,1301,1325]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B06D04FFF277E27FE2BA270" bold="true" box="[151,453,1301,1325]" italics="true" pageId="2" pageNumber="423">Excalidiopsis palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B06D04FFE7B7E26FDCFA271" box="[459,545,1300,1324]" pageId="2" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B06D04FFF277E07FEBEA211" box="[151,336,1333,1356]" pageId="2" pageNumber="423" type="description">
<paragraph id="8BDF36888B06D04FFF277E07FEBEA211" blockId="2.[151,545,1300,1356]" box="[151,336,1333,1356]" pageId="2" pageNumber="423">
(
<figureCitation id="135B2A0D8B06D04FFF2F7E07FF30A211" box="[159,222,1333,1356]" captionStart="Figure 1" captionStartId="3.[113,183,1636,1659]" captionTargetBox="[118,1130,579,1606]" captionTargetId="figure-206@3.[113,1134,579,1607]" captionTargetPageId="3" captionText="Figure 1. Excalidiopsis palawanensis sp. nov. A Holotype male, dorsal view. B paratype female, dorsal view." pageId="2" pageNumber="423">Fig. 1</figureCitation>
,
<figureCitation id="135B2A0D8B06D04FFF5C7E07FEC0A211" box="[236,302,1333,1356]" captionStart="Figure 3" captionStartId="6.[151,222,663,686]" captionTargetBox="[152,1171,152,628]" captionTargetId="figure-52@6.[151,1172,151,629]" captionTargetPageId="6" captionText="Figure 3. AC. Dorsaw view of head and antennae of selected species. A Excalidiopsis palawanensis sp nov. B Excalidiopsis bramanti sp. nov. C Calidiopsis bilineatus Rukmane, 2020; DE. Lateral view of selected species. D Excalidiopsis palawanensis sp. nov. E Excalidiopsis bramanti sp. nov." pageId="2" pageNumber="423">3A, D</figureCitation>
,
<figureCitation id="135B2A0D8B06D04FFE8A7E07FEA6A211" box="[314,328,1333,1356]" captionStart="Figure 4" captionStartId="6.[151,221,1569,1592]" captionTargetBox="[237,559,834,1537]" captionTargetId="figure-364@6.[234,561,831,1539]" captionTargetPageId="6" captionText="Figure 4. AB. Male genitalia of Excalidiopsis palawanensis sp. nov. A aedeagus in lateral view. B sternite IX in dorsal view. Scale bar 1 mm." pageId="2" pageNumber="423">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B06D04FFF277E5EFCE5A628" pageId="2" pageNumber="423" type="materials_examined">
<paragraph id="8BDF36888B06D04FFF277E5EFDFBA179" blockId="2.[151,643,1388,1572]" pageId="2" pageNumber="423">
<emphasis id="B914EA9A8B06D04FFF277E5EFEABA2D9" bold="true" box="[151,325,1388,1412]" pageId="2" pageNumber="423">Type material.</emphasis>
<materialsCitation id="3B083CD58B06D04FFEE07E5EFDFBA179" collectingDate="2019-10" collectionCode="DUBC" country="Philippines" location="Roxas" municipality="Magara" pageId="2" pageNumber="423" specimenCount="2" stateProvince="Palawan" typeStatus="holotype">
<emphasis id="B914EA9A8B06D04FFEE07E5EFDE0A2D9" bold="true" box="[336,526,1388,1412]" pageId="2" pageNumber="423">
<typeStatus id="54DB882A8B06D04FFEE07E5EFE54A2D9" box="[336,442,1388,1412]" pageId="2" pageNumber="423" type="holotype">Holotype</typeStatus>
. Male.
</emphasis>
<collectingCountry id="F37776188B06D04FFD947E5EFF0FA2FE" name="Philippines" pageId="2" pageNumber="423">PHILIPPINES</collectingCountry>
/
<collectingRegion id="49A4F86A8B06D04FFF477EBEFEBFA2FE" box="[247,337,1420,1443]" country="Philippines" name="Palawan" pageId="2" pageNumber="423">Palawan</collectingRegion>
/
<collectingMunicipality id="6BBBACF28B06D04FFED77EBEFE55A2FE" box="[359,443,1420,1443]" pageId="2" pageNumber="423">Magara</collectingMunicipality>
,
<location id="8EBF60538B06D04FFE767EBEFDE7A2FE" LSID="urn:lsid:plazi:treatment:03C9879E8B06D049FF277E27FF3BA288:8EBF60538B06D04FFE767EBEFDE7A2FE" box="[454,521,1420,1443]" country="Philippines" municipality="Magara" name="Roxas" pageId="2" pageNumber="423" stateProvince="Palawan">Roxas</location>
/
<date id="FFDE10488B06D04FFDAE7EBEFD9AA2FE" box="[542,628,1420,1443]" pageId="2" pageNumber="423" value="2019-10">
<collectingDate id="EF9AE9A08B06D04FFDAE7EBEFD9AA2FE" box="[542,628,1420,1443]" pageId="2" pageNumber="423" value="2019-10">X. 2019</collectingDate>
</date>
/ local collector leg.” (white rectangular label, printed). “
<typeStatus id="54DB882A8B06D04FFEB27EFFFE7FA2B9" box="[258,401,1485,1508]" pageId="2" pageNumber="423" type="holotype">HOLOTYPE</typeStatus>
/ Male /
<taxonomicName id="4C604D0B8B06D04FFE437EFFFEC4A159" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="B914EA9A8B06D04FFE437EFFFEC4A159" italics="true" pageId="2" pageNumber="423">Excalidiopsis palawanensis</emphasis>
</taxonomicName>
Rukmane­Brbale, 2022” (red rectangular label, printed) (
<collectionCode id="ED71AE4D8B06D04FFE097D3FFDE7A179" box="[441,521,1549,1572]" pageId="2" pageNumber="423">DUBC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BDF36888B06D04FFF277D76FD6DA1C1" blockId="2.[151,643,1604,1692]" pageId="2" pageNumber="423">
<materialsCitation id="3B083CD58B06D04FFF277D76FEEBA1C1" collectingDate="2019-10" country="Philippines" location="Roxas" municipality="Magara" pageId="2" pageNumber="423" specimenCount="1" stateProvince="Palawan" typeStatus="paratype">
<emphasis id="B914EA9A8B06D04FFF277D76FEE6A101" bold="true" box="[151,264,1604,1628]" pageId="2" pageNumber="423">
<typeStatus id="54DB882A8B06D04FFF277D76FEE6A101" box="[151,264,1604,1628]" pageId="2" pageNumber="423" type="paratype">Paratypes</typeStatus>
</emphasis>
(4, 8): “
<collectingCountry id="F37776188B06D04FFECC7D76FDF4A106" box="[380,538,1604,1627]" name="Philippines" pageId="2" pageNumber="423">PHILIPPINES</collectingCountry>
/
<collectingRegion id="49A4F86A8B06D04FFD9A7D76FD6DA106" box="[554,643,1604,1627]" country="Philippines" name="Palawan" pageId="2" pageNumber="423">Palawan</collectingRegion>
/
<collectingMunicipality id="6BBBACF28B06D04FFF147D57FF19A121" box="[164,247,1637,1660]" pageId="2" pageNumber="423">Magara</collectingMunicipality>
,
<location id="8EBF60538B06D04FFEB07D57FEACA121" LSID="urn:lsid:plazi:treatment:03C9879E8B06D049FF277E27FF3BA288:8EBF60538B06D04FFEB07D57FEACA121" box="[256,322,1637,1660]" country="Philippines" municipality="Magara" name="Roxas" pageId="2" pageNumber="423" stateProvince="Palawan">Roxas</location>
/
<date id="FFDE10488B06D04FFEE37D57FE48A121" box="[339,422,1637,1660]" pageId="2" pageNumber="423" value="2019-10">
<collectingDate id="EF9AE9A08B06D04FFEE37D57FE48A121" box="[339,422,1637,1660]" pageId="2" pageNumber="423" value="2019-10">X. 2019</collectingDate>
</date>
/ local collector leg.” (3, 6) //
</materialsCitation>
<materialsCitation id="3B083CD58B06D04FFEA37DB7FD91A1C1" box="[275,639,1669,1692]" country="Philippines" location="Palawan" pageId="2" pageNumber="423" specimenCount="1" stateProvince="Palawan" typeStatus="paratype">
<collectingCountry id="F37776188B06D04FFEA37DB7FE5FA1C1" box="[275,433,1669,1692]" name="Philippines" pageId="2" pageNumber="423">PHILIPPINES</collectingCountry>
/
<collectingRegion id="49A4F86A8B06D04FFE717DB7FDF4A1C1" box="[449,538,1669,1692]" country="Philippines" name="Palawan" pageId="2" pageNumber="423">Palawan</collectingRegion>
/ Magara
</materialsCitation>
,
</paragraph>
<paragraph id="8BDF36888B06D04FFD187BA9FCE5A628" blockId="2.[680,1172,155,373]" pageId="2" pageNumber="423">
<materialsCitation id="3B083CD58B06D04FFD187BA9FCE6A628" collectingDate="2020-01" collectingDateMax="2021-10" collectingDateMin="2020-01" collectionCode="DUBC" country="Philippines" location="All" municipality="Roxas" pageId="2" pageNumber="423" specimenCount="1" stateProvince="Palawan" typeStatus="paratype">
<collectingMunicipality id="6BBBACF28B06D04FFD187BA9FD05A7EF" box="[680,747,155,178]" pageId="2" pageNumber="423">Roxas</collectingMunicipality>
/
<date id="FFDE10488B06D04FFCB57BA9FCBCA7EF" box="[773,850,155,178]" pageId="2" pageNumber="423" value="2020-01">
<collectingDate id="EF9AE9A08B06D04FFCB57BA9FCBCA7EF" box="[773,850,155,178]" pageId="2" pageNumber="423" value="2020-01">I. 2020</collectingDate>
</date>
/ local collector leg.” (1) // “
<collectingCountry id="F37776188B06D04FFD027B8EFCBCA78E" box="[690,850,188,211]" name="Philippines" pageId="2" pageNumber="423">PHILIPPINES</collectingCountry>
/
<collectingRegion id="49A4F86A8B06D04FFCDB7B8EFC2BA78E" box="[875,965,188,211]" country="Philippines" name="Palawan" pageId="2" pageNumber="423">Palawan</collectingRegion>
/ Magara, Roxas /
<date id="FFDE10488B06D04FFD187BEEFD10A7AE" box="[680,766,220,243]" pageId="2" pageNumber="423" value="2021-10">
<collectingDate id="EF9AE9A08B06D04FFD187BEEFD10A7AE" box="[680,766,220,243]" pageId="2" pageNumber="423" value="2021-10">X. 2021</collectingDate>
</date>
/ local collector leg.” (1, 1) (all on white rectangular labels, printed).
<location id="8EBF60538B06D04FFB857BCFFBB6A649" LSID="urn:lsid:plazi:treatment:03C9879E8B06D049FF277E27FF3BA288:8EBF60538B06D04FFB857BCFFBB6A649" box="[1077,1112,253,276]" country="Philippines" municipality="Roxas" name="All" pageId="2" pageNumber="423" stateProvince="Palawan">All</location>
with additional red label: “
<typeStatus id="54DB882A8B06D04FFC227A2FFBF4A669" box="[914,1050,285,308]" pageId="2" pageNumber="423" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C604D0B8B06D04FFB827A2FFC96A608" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="B914EA9A8B06D04FFB827A2FFC96A608" italics="true" pageId="2" pageNumber="423">Excalidiopsis palawanensis</emphasis>
</taxonomicName>
Rukmane­Brbale, 2022” (
<collectionCode id="ED71AE4D8B06D04FFD007A6CFCEEA628" box="[688,768,350,373]" pageId="2" pageNumber="423">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B06D049FD187AA7FE09A3D0" lastPageId="4" lastPageNumber="425" pageId="2" pageNumber="423" type="description">
<paragraph id="8BDF36888B06D04FFD187AA7FB81A5CD" blockId="2.[680,1172,405,656]" pageId="2" pageNumber="423">
<emphasis id="B914EA9A8B06D04FFD187AA7FCDFA6F0" bold="true" box="[680,817,405,429]" pageId="2" pageNumber="423">Description.</emphasis>
Male. Measuremants: LB: 6.7­ 7.2 (
<typeStatus id="54DB882A8B06D04FFD007A84FCE1A690" box="[688,783,438,461]" pageId="2" pageNumber="423" type="holotype">holotype</typeStatus>
7.1, mean 6.98); LP: 1.7­ 1.9 (
<typeStatus id="54DB882A8B06D04FFBEB7A84FD3BA6B3" pageId="2" pageNumber="423" type="holotype">holotype</typeStatus>
1.9, mean 1.82); LE: 5.0­ 5.3 (
<typeStatus id="54DB882A8B06D04FFBBA7AE5FB86A6B3" box="[1034,1128,471,494]" pageId="2" pageNumber="423" type="holotype">holotype</typeStatus>
5.2, mean 5.16); LR: 1.2­ 1.3 (
<typeStatus id="54DB882A8B06D04FFC717AC5FBCEA553" box="[961,1056,503,526]" pageId="2" pageNumber="423" type="holotype">holotype</typeStatus>
1.3, mean 1.26); WP: 1.7­ 1.8 (
<typeStatus id="54DB882A8B06D04FFCCF792AFC30A572" box="[895,990,536,559]" pageId="2" pageNumber="423" type="holotype">holotype</typeStatus>
1.8, mean 1.76); WE: 3.6­ 3.8 (
<typeStatus id="54DB882A8B06D04FFC86790AFC7BA512" box="[822,917,568,591]" pageId="2" pageNumber="423" type="holotype">holotype</typeStatus>
3.8, mean 3.7); WR: 1.1 (
<typeStatus id="54DB882A8B06D04FFD00796BFCE1A52D" box="[688,783,601,624]" pageId="2" pageNumber="423" type="holotype">holotype</typeStatus>
1.1, mean 1.1). N=5 for all measurements. Dorsal habitus as shown in
<figureCitation id="135B2A0D8B06D04FFBA8794BFB85A5CD" box="[1048,1131,633,656]" captionStart="Figure 1" captionStartId="3.[113,183,1636,1659]" captionTargetBox="[118,1130,579,1606]" captionTargetId="figure-206@3.[113,1134,579,1607]" captionTargetPageId="3" captionText="Figure 1. Excalidiopsis palawanensis sp. nov. A Holotype male, dorsal view. B paratype female, dorsal view." pageId="2" pageNumber="423">Fig. 1A</figureCitation>
.
</paragraph>
<paragraph id="8BDF36888B06D04FFD187982FBCFA5B5" blockId="2.[680,1172,688,744]" pageId="2" pageNumber="423">Body black, with markings of glossy yellow to goldish, round to recumbent scales.</paragraph>
<paragraph id="8BDF36888B06D04FFD18783AFCA3A3D8" blockId="2.[680,1172,776,1157]" pageId="2" pageNumber="423">Forehead nearly three times as wide as eye width; eyes slightly prominent from the outline of the head, widest just after the middle dorsally; LR/ WR 1.18; rostrum dorsally covered with round, shiny scales except apical 1/2; frontal part of rostrum slightly widened apically; in lateral contour slightly increased just before the middle, sulcus with impression at base of the eye; lateral parts covered with round shiny scales at basal part and with scales and short light hairs at apical part; dorsal furrow of antennal scrobe sharp, wide V­shaped.</paragraph>
<paragraph id="8BDF36888B06D04FFD187F97FC6BA29C" blockId="2.[680,1172,1189,1473]" pageId="2" pageNumber="423">Antennal scape covered with eliptic scales from base to apical 1/2, short light hairs on all length; segments I to VII subegual in length, except segment II of slightly longer length, nearly same lenght and width, covered with eliptic shiny scales; club short, slightly longer whan wide, divided into three parts, first and third part beeing same width and second narrower; club without shiny scales.</paragraph>
<paragraph id="8BDF36888B06D04EFD187ED3FF42A649" blockId="2.[680,1172,1505,1690]" lastBlockId="3.[113,605,155,276]" lastPageId="3" lastPageNumber="424" pageId="2" pageNumber="423">Pronotum nearly as long as wide, LP/WP: 1.05, with the following markings of round to recumbent shiny scales: 1) longitudinal medial line at disc dorsally in all length; 2) wide longitudinal line at each dorso­lateral edge in all length; 3) large patch from one latero­ventral edge to other; dorsal contour slightly widened just before the middle; lateral contour straight; both basal and apical concavities covered with short, light hairs.</paragraph>
<paragraph id="8BDF36888B07D04EFFC17A06FE84A553" blockId="3.[113,605,308,526]" pageId="3" pageNumber="424">Femora with scally markings at basal part, bare before the middle, with markings from middle to apical half and bare to apex, mid femora with scaly markings from base to apical half, mingled with light hairs in all length; tarsus covered with scales, mingled with longer light hairs, hairs longer along inner margin.</paragraph>
<paragraph id="8BDF36888B07D049FD337BA9FE64A668" blockId="3.[643,1134,155,530]" lastBlockId="4.[151,643,155,309]" lastPageId="4" lastPageNumber="425" pageId="3" pageNumber="424">Elytra longer than wide, LE/WE 1.37, widest just in the middle, each elytron with the following scally patches: 1) two longitudinal patches at basal 1/2, one along intervals II to III, second along intervals VI to VII; 2) four patches at medial part, one redirected apically, from middle of interval I to middle of interval III, two patches along interval V, one redirected basally, one apically, one medial patch redirected laterally along interval VII; 3) triangular­shaped patch at apical part before apex; 4) two sutural patches, one before the middle, one from apical 1/2; 5) lateral margin with solid line of scales that in some cases are interupted and divided in smaller patches; in lateral contour slightly impressed just before the middle, bulged along apical 1/2 and decreased to apex.</paragraph>
<caption id="DF1F66008B07D04EFFC17D56FEA8A1C6" pageId="3" pageNumber="424" startId="3.[113,183,1636,1659]" targetBox="[118,1130,579,1606]" targetPageId="3" targetType="figure">
<paragraph id="8BDF36888B07D04EFFC17D56FEA8A1C6" blockId="3.[113,1134,1636,1691]" pageId="3" pageNumber="424">
Figure 1.
<taxonomicName id="4C604D0B8B07D04EFF6E7D56FDE6A126" authorityName="Rukmane-Brbale" authorityYear="2022" box="[222,520,1636,1659]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B07D04EFF6E7D56FDE6A126" box="[222,520,1636,1659]" italics="true" pageId="3" pageNumber="424">Excalidiopsis palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B07D04EFDA27D56FD89A126" box="[530,615,1636,1659]" pageId="3" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B914EA9A8B07D04EFDC07D56FD6DA121" bold="true" box="[624,643,1636,1660]" pageId="3" pageNumber="424">A</emphasis>
Holotype male, dorsal view.
<emphasis id="B914EA9A8B07D04EFC507D56FC1CA121" bold="true" box="[992,1010,1636,1660]" pageId="3" pageNumber="424">B</emphasis>
paratype female, dorsal view.
</paragraph>
</caption>
<paragraph id="8BDF36888B00D049FF277A64FE10A630" blockId="4.[151,510,342,365]" box="[151,510,342,365]" pageId="4" pageNumber="425">
Male genitalia as shown in
<figureCitation id="135B2A0D8B00D049FE0A7A64FE14A630" box="[442,506,342,365]" captionStart="Figure 4" captionStartId="6.[151,221,1569,1592]" captionTargetBox="[237,559,834,1537]" captionTargetId="figure-364@6.[234,561,831,1539]" captionTargetPageId="6" captionText="Figure 4. AB. Male genitalia of Excalidiopsis palawanensis sp. nov. A aedeagus in lateral view. B sternite IX in dorsal view. Scale bar 1 mm." pageId="4" pageNumber="425">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="8BDF36888B00D049FF277ABFFEBAA515" blockId="4.[151,643,397,584]" pageId="4" pageNumber="425">
Female. Measurements: LB: 7.6­8.0 (mean 7.78); LP: 1.9­ 2.1 (mean 2); LE: 5.5­ 5.9 (mean 5.74); LR: 1.3­ 1.4 (mean 1.36); WP: 1.7­ 1.9 (mean 1.8); WE: 4.1­ 4.3 (mean 4.22); WR: 1.1. N=5 for all measuremants. Dorsal habitus as shown in
<figureCitation id="135B2A0D8B00D049FF4E7903FEA1A515" box="[254,335,561,584]" captionStart="Figure 1" captionStartId="3.[113,183,1636,1659]" captionTargetBox="[118,1130,579,1606]" captionTargetId="figure-206@3.[113,1134,579,1607]" captionTargetPageId="3" captionText="Figure 1. Excalidiopsis palawanensis sp. nov. A Holotype male, dorsal view. B paratype female, dorsal view." pageId="4" pageNumber="425">Fig. 1B</figureCitation>
.
</paragraph>
<paragraph id="8BDF36888B00D049FF27795BFE09A3D0" blockId="4.[151,643,617,1165]" pageId="4" pageNumber="425">
<emphasis id="B914EA9A8B00D049FF27795BFE6CA5DC" bold="true" box="[151,386,617,641]" pageId="4" pageNumber="425">Differential analyses.</emphasis>
The new species is similar to second species of the genus
<taxonomicName id="4C604D0B8B00D049FE4479B8FD6DA5FC" authorityName="Rukmane-Brbale" authorityYear="2022" box="[500,643,650,673]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914EA9A8B00D049FE4479B8FD6DA5FC" box="[500,643,650,673]" italics="true" pageId="4" pageNumber="425">Excalidiopsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FF277999FEEFA59F" box="[151,257,683,706]" pageId="4" pageNumber="425" rank="genus">gen. nov.</taxonomicNameLabel>
:
<taxonomicName id="4C604D0B8B00D049FEA47999FDF3A59F" authorityName="Rukmane-Brbale" authorityYear="2022" box="[276,541,683,706]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FEA47999FDF3A59F" box="[276,541,683,706]" italics="true" pageId="4" pageNumber="425">Excalidiopsis bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FD997999FD6DA59F" box="[553,643,683,706]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
Species might be distinguished by the following morphological features: 1) scally markings in
<taxonomicName id="4C604D0B8B00D049FF27783FFEAFA479" authorityName="Rukmane-Brbale" authorityYear="2022" box="[151,321,781,804]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FF27783FFEAFA479" box="[151,321,781,804]" italics="true" pageId="4" pageNumber="425">E. palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FEF6783FFE7AA479" box="[326,404,781,804]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
of goldish to yellowish colour, while greenish in
<taxonomicName id="4C604D0B8B00D049FE15781CFDCAA418" authorityName="Rukmane-Brbale" authorityYear="2022" box="[421,548,814,837]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FE15781CFDCAA418" box="[421,548,814,837]" italics="true" pageId="4" pageNumber="425">E. bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FD9A781CFD95A418" box="[554,635,814,837]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
; 2) segment II nearly same size as rest in
<taxonomicName id="4C604D0B8B00D049FD80787DFF16A4DA" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FD80787DFF16A4DA" italics="true" pageId="4" pageNumber="425">E. palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FF4D785DFEA2A4DB" box="[253,332,879,902]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
while at least 1.5 times longer than rest in
<taxonomicName id="4C604D0B8B00D049FEA078A2FE60A4FA" authorityName="Rukmane-Brbale" authorityYear="2022" box="[272,398,912,935]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FEA078A2FE60A4FA" box="[272,398,912,935]" italics="true" pageId="4" pageNumber="425">E. bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FE2378A2FE0DA4FA" box="[403,483,912,935]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
; 3) sulcus laterally with groove at base of the eye in
<taxonomicName id="4C604D0B8B00D049FD9D7883FF1AA4B4" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FD9D7883FF1AA4B4" italics="true" pageId="4" pageNumber="425">E. palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FF4878E0FEABA4B4" box="[248,325,978,1001]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
and just before the distal margin of eye in
<taxonomicName id="4C604D0B8B00D049FF4778C1FE9BA357" authorityName="Rukmane-Brbale" authorityYear="2022" box="[247,373,1011,1034]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FF4778C1FE9BA357" box="[247,373,1011,1034]" italics="true" pageId="4" pageNumber="425">E. bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FECB78C1FE25A357" box="[379,459,1011,1034]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
; 4) dorsal furrow sharp in
<taxonomicName id="4C604D0B8B00D049FF5F7F21FE75A377" authorityName="Rukmane-Brbale" authorityYear="2022" box="[239,411,1043,1066]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FF5F7F21FE75A377" box="[239,411,1043,1066]" italics="true" pageId="4" pageNumber="425">E. palawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FE2F7F21FE00A377" box="[415,494,1043,1066]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
, but moderate in
<taxonomicName id="4C604D0B8B00D049FF017F06FEC0A316" authorityName="Rukmane-Brbale" authorityYear="2022" box="[177,302,1076,1099]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FF017F06FEC0A316" box="[177,302,1076,1099]" italics="true" pageId="4" pageNumber="425">E. bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FE837F06FE6CA316" box="[307,386,1076,1099]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
; 5) elytra of
<taxonomicName id="4C604D0B8B00D049FDB67F06FD6CA316" authorityName="Rukmane-Brbale" authorityYear="2022" box="[518,642,1076,1099]" class="Insecta" family="Curculionidae" genus="Excalidiopsis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="bramanti" status="sp. nov.">
<emphasis id="B914EA9A8B00D049FDB67F06FD6CA316" box="[518,642,1076,1099]" italics="true" pageId="4" pageNumber="425">E. bramanti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757E18B00D049FF277F67FF08A331" box="[151,230,1109,1132]" pageId="4" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
wider, more strongly rounded, in lateral contour more stronghy bulging.
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B00D049FF277F9EFE36A399" box="[151,472,1196,1220]" pageId="4" pageNumber="425" type="distribution">
<paragraph id="8BDF36888B00D049FF277F9EFE36A399" blockId="4.[151,472,1196,1220]" box="[151,472,1196,1220]" pageId="4" pageNumber="425">
<emphasis id="B914EA9A8B00D049FF277F9EFEC4A399" bold="true" box="[151,298,1196,1220]" pageId="4" pageNumber="425">Distribution.</emphasis>
<collectingRegion id="49A4F86A8B00D049FE9F7F9FFE67A399" box="[303,393,1197,1220]" country="Philippines" name="Palawan" pageId="4" pageNumber="425">Palawan</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C37A65038B00D049FF277FD6FF3BA288" pageId="4" pageNumber="425" type="etymology">
<paragraph id="8BDF36888B00D049FF277FD6FD80A241" blockId="4.[151,643,1252,1308]" pageId="4" pageNumber="425">
<emphasis id="B914EA9A8B00D049FF277FD6FEF6A3A1" bold="true" box="[151,280,1252,1276]" pageId="4" pageNumber="425">Etymology.</emphasis>
The new species name is latinized name of
<collectingCountry id="F37776188B00D049FF447E37FEDBA241" box="[244,309,1285,1308]" name="Iceland" pageId="4" pageNumber="425">Island</collectingCountry>
where species is distrubuted.
</paragraph>
<paragraph id="8BDF36888B00D049FF277E0EFF3BA288" blockId="4.[151,643,1340,1493]" pageId="4" pageNumber="425">
<emphasis id="B914EA9A8B00D049FF277E0EFEEEA209" bold="true" box="[151,256,1340,1364]" pageId="4" pageNumber="425">Mimicry.</emphasis>
The species consist in mimetic relationship with
<taxonomicName id="4C604D0B8B00D049FE827E6FFE64A2C9" authority="Yoshitake, 2013" authorityName="Yoshitake" authorityYear="2013" class="Insecta" family="Curculionidae" genus="Expachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="chloromaculatus">
<emphasis id="B914EA9A8B00D049FE827E6FFF22A2C9" italics="true" pageId="4" pageNumber="425">Expachyrhynchus chloromaculatus</emphasis>
<bibRefCitation id="EFF14B798B00D049FF677E4FFE64A2C9" author="Yoshitake H." box="[215,394,1405,1428]" pageId="4" pageNumber="425" pagination="1 - 8" refId="ref3835" refString="Yoshitake H. 2013. A new genus and two new species of the tribe Pachyrhynchini (Coleoptera: Curculionidae) from Palawan Island, the Philippines. Esakia, 53: 1 - 8." type="journal article" year="2013">Yoshitake, 2013</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C604D0B8B00D049FE767E4FFF3EA288" authority="Rukmane, Bramanti &amp; Bramanti, 2020" authorityName="Rukmane, Bramanti &amp; Bramanti" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Evrostopolycatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="B914EA9A8B00D049FE767E4FFEC8A2E8" italics="true" pageId="4" pageNumber="425">Evrostopolycatus palawanensis</emphasis>
Rukmane,
<bibRefCitation id="EFF14B798B00D049FE2B7EAFFF3EA288" author="Bramanti A. &amp; Bramanti A." pageId="4" pageNumber="425" pagination="87 - 94" refId="ref3785" refString="Rukmane-B ' rbale A., Bramanti A., Bramanti A. 2020. A new genus of the trive Polocatini Marshal, 1956 (Coleoptera: Curculionidae: Entiminae) from the Philippines, Baltic Journal of Coleopterology, 20 (1): 87 - 94." type="journal article" year="2020">Bramanti &amp; Bramanti, 2020</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="03CB87ED015AFFE4C103CA4FFCDE8F88" LSID="urn:lsid:plazi:treatment:03CB87ED015AFFE4C103CA4FFCDE8F88" httpUri="http://treatment.plazi.org/id/03CB87ED015AFFE4C103CA4FFCDE8F88" lastPageId="7" lastPageNumber="182" pageId="6" pageNumber="181">
<subSubSection id="C3786570015AFFE5C103CA4FFB4A8A5B" box="[850,1249,1376,1400]" pageId="6" pageNumber="181" type="nomenclature">
<paragraph id="8BDD36FB015AFFE5C103CA4FFB4A8A5B" blockId="6.[850,1249,1376,1431]" box="[850,1249,1376,1400]" pageId="6" pageNumber="181">
<heading id="D0958197015AFFE5C103CA4FFB4A8A5B" box="[850,1249,1376,1400]" fontSize="10" level="2" pageId="6" pageNumber="181" reason="7">
<taxonomicName id="4C624D78015AFFE5C103CA4FFB4A8A5B" ID-CoL="4SV65" authority="Schmidt, 1916" authorityName="Schmidt" authorityYear="1916" box="[850,1249,1376,1400]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="B916EAE9015AFFE5C103CA4FFB968A54" bold="true" box="[850,1085,1376,1400]" italics="true" pageId="6" pageNumber="181">Rhyparus verrucosus</emphasis>
Schmidt, 1916
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3786570015AFFE5C103CAAFFC778ABB" box="[850,988,1408,1431]" pageId="6" pageNumber="181" type="description">
<paragraph id="8BDD36FB015AFFE5C103CAAFFC778ABB" blockId="6.[850,1249,1376,1431]" box="[850,988,1408,1431]" pageId="6" pageNumber="181">
(
<figureCitation id="13592A7E015AFFE5C10BCAAFFC7F8ABB" box="[858,980,1408,1431]" captionStart="Figs 1316" captionStartId="7.[227,273,1189,1212]" captionTargetBox="[333,1182,192,1154]" captionTargetId="figure-241@7.[330,1184,189,1157]" captionTargetPageId="7" captionText="Figs 1316. Dorsal views and abdominal apexes: 13 dorsal view of Rhyparus denticollis, 14 dorsal view of R. verrucosus (Bali Island), 15 abdominal apex of R. verrucosus (holotype, Sumatra), 16 abdominal apex of R. jakli sp. nov. (scale lines: 1.0 mm)." pageId="6" pageNumber="181">Figs 14, 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3786570015AFFE4C103CAEDFCDE8F88" lastPageId="7" lastPageNumber="182" pageId="6" pageNumber="181" type="materials_examined">
<paragraph id="8BDD36FB015AFFE5C103CAEDFB8E8937" blockId="6.[850,1361,1474,1563]" pageId="6" pageNumber="181">
<emphasis id="B916EAE9015AFFE5C103CAEDFB9E8AF6" bold="true" box="[850,1077,1474,1498]" pageId="6" pageNumber="181">Examined material.</emphasis>
<materialsCitation id="3B0A3CA6015AFFE5C66BCAEDFB8A8937" collectingDate="2004-11" collectionCode="Y" country="Indonesia" elevation="1200" location="Bedugul region" pageId="6" pageNumber="181" specimenCount="1" stateProvince="Bali">
<specimenCount id="9D64FD72015AFFE5C66BCAEDFBDA8AF5" box="[1082,1137,1474,1497]" count="1" pageId="6" pageNumber="181" type="generic">1 ex.</specimenCount>
:
<collectingCountry id="F375766B015AFFE5C62ACAEDFB4C8AF5" box="[1147,1255,1474,1497]" name="Indonesia" pageId="6" pageNumber="181">Indonesia</collectingCountry>
,
<collectingRegion id="49A6F819015AFFE5C6A1CAEDFAB48AF5" box="[1264,1311,1474,1497]" country="Indonesia" name="Bali" pageId="6" pageNumber="181">Bali</collectingRegion>
,
<location id="8EBD6020015AFFE5C77BCAEDFC768AD6" LSID="urn:lsid:plazi:treatment:03CB87ED015AFFE4C103CA4FFCDE8F88:8EBD6020015AFFE5C77BCAEDFC768AD6" country="Indonesia" name="Bedugul region" pageId="6" pageNumber="181" stateProvince="Bali">Bedugul region</location>
,
<quantity id="4C9A9B1E015AFFE5C1B6CACCFB9C8AD6" box="[999,1079,1507,1530]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="181" unit="m" value="1200.0">
<elevation id="004FD1C8015AFFE5C1B6CACCFB9C8AD6" box="[999,1079,1507,1530]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="181" unit="m" value="1200.0">1200m</elevation>
</quantity>
.,
<date id="FFDC103B015AFFE5C614CACCFB3D8AD6" box="[1093,1174,1507,1530]" pageId="6" pageNumber="181" value="2004-11">
<collectingDate id="EF98E9D3015AFFE5C614CACCFB3D8AD6" box="[1093,1174,1507,1530]" pageId="6" pageNumber="181" value="2004-11">xi.2004</collectingDate>
</date>
[
<emphasis id="B916EAE9015AFFE5C6F4CACDFB1E8AD7" bold="true" box="[1189,1205,1506,1531]" pageId="6" pageNumber="181">
<collectionCode id="ED73AE3E015AFFE5C6F4CACDFB1E8AD7" box="[1189,1205,1506,1531]" pageId="6" pageNumber="181">Ÿ</collectionCode>
</emphasis>
MCN] [photographed specimen]
</materialsCitation>
.
</paragraph>
<paragraph id="8BDD36FB015AFFE5C103C96AFB4B8951" blockId="6.[850,1360,1605,1661]" pageId="6" pageNumber="181">
<emphasis id="B916EAE9015AFFE5C103C96AFBAB8971" bold="true" box="[850,1024,1605,1629]" pageId="6" pageNumber="181">Short comment.</emphasis>
Previously was known only from
<collectingRegion id="49A6F819015AFFE5C103C949FC048951" box="[850,943,1638,1661]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="181">Sumatra</collectingRegion>
. First record for
<collectingRegion id="49A6F819015AFFE5C632C949FB3B8951" box="[1123,1168,1638,1661]" country="Indonesia" name="Bali" pageId="6" pageNumber="181">Bali</collectingRegion>
Island.
</paragraph>
<paragraph id="8BDD36FB015BFFE4C025CFBEFCDE8F88" blockId="7.[628,885,145,164]" box="[628,885,145,164]" pageId="7" pageNumber="182">
<emphasis id="B916EAE9015BFFE4C025CFBEFCDE8F88" box="[628,885,145,164]" italics="true" pageId="7" pageNumber="182">Minkina Ł., Anichtchenko A.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,238 @@
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<mods:title id="D401D4E43DC8DC75AC31D8300004FB6F">Studies on genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with description of two new species from Southeast Asia</mods:title>
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</mods:mods>
<treatment id="03CB87ED015AFFE5C103CE2BFBA88A38" LSID="urn:lsid:plazi:treatment:03CB87ED015AFFE5C103CE2BFBA88A38" httpUri="http://treatment.plazi.org/id/03CB87ED015AFFE5C103CE2BFBA88A38" lastPageNumber="181" pageId="6" pageNumber="181">
<subSubSection id="C3786570015AFFE5C103CE2BFB428E30" box="[850,1257,260,284]" pageId="6" pageNumber="181" type="nomenclature">
<paragraph id="8BDD36FB015AFFE5C103CE2BFB428E30" blockId="6.[850,1257,260,316]" box="[850,1257,260,284]" pageId="6" pageNumber="181">
<heading id="D0958197015AFFE5C103CE2BFB428E30" box="[850,1257,260,284]" fontSize="10" level="2" pageId="6" pageNumber="181" reason="7">
<taxonomicName id="4C624D78015AFFE5C103CE2BFB428E30" ID-CoL="4SV4F" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[850,1257,260,284]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="B916EAE9015AFFE5C103CE2BFB9D8E30" bold="true" box="[850,1078,260,284]" italics="true" pageId="6" pageNumber="181">Rhyparus denticollis</emphasis>
Fairmaire, 1893
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3786570015AFFE5C103CE0AFC1A8E10" box="[850,945,293,316]" pageId="6" pageNumber="181" type="description">
<paragraph id="8BDD36FB015AFFE5C103CE0AFC1A8E10" blockId="6.[850,1257,260,316]" box="[850,945,293,316]" pageId="6" pageNumber="181">
(
<figureCitation id="13592A7E015AFFE5C10BCE0AFC0C8E10" box="[858,935,293,316]" captionStart="Figs 1316" captionStartId="7.[227,273,1189,1212]" captionTargetBox="[333,1182,192,1154]" captionTargetId="figure-241@7.[330,1184,189,1157]" captionTargetPageId="7" captionText="Figs 1316. Dorsal views and abdominal apexes: 13 dorsal view of Rhyparus denticollis, 14 dorsal view of R. verrucosus (Bali Island), 15 abdominal apex of R. verrucosus (holotype, Sumatra), 16 abdominal apex of R. jakli sp. nov. (scale lines: 1.0 mm)." pageId="6" pageNumber="181">Fig. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3786570015AFFE5C103CE49FBA88A38" pageId="6" pageNumber="181" type="materials_examined">
<paragraph id="8BDD36FB015AFFE5C103CE49FC128C80" blockId="6.[850,1361,358,940]" pageId="6" pageNumber="181">
<emphasis id="B916EAE9015AFFE5C103CE49FB9E8E52" bold="true" box="[850,1077,358,382]" pageId="6" pageNumber="181">Examined material.</emphasis>
<materialsCitation id="3B0A3CA6015AFFE5C66BCE48FB9B8EB3" collectionCode="NMPC" collectorName="Palembang" country="Indonesia" location="Sumatra" pageId="6" pageNumber="181" specimenCount="1" stateProvince="Sumatera Utara">
<specimenCount id="9D64FD72015AFFE5C66BCE48FBDA8E52" box="[1082,1137,359,382]" count="1" pageId="6" pageNumber="181" type="generic">1 ex.</specimenCount>
:
<collectingCountry id="F375766B015AFFE5C62ACE48FB4C8E52" box="[1147,1255,359,382]" name="Indonesia" pageId="6" pageNumber="181">Indonesia</collectingCountry>
,
<collectingRegion id="49A6F819015AFFE5C6A1CE48FAE68E52" box="[1264,1357,359,382]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="181">Sumatra</collectingRegion>
,
<collectorName id="2697532D015AFFE5C103CEA7FC618EB3" box="[850,970,392,415]" pageId="6" pageNumber="181">Palembang</collectorName>
[
<collectionCode id="ED73AE3E015AFFE5C184CEA7FB818EB3" box="[981,1066,392,415]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="6" pageNumber="181">NMPC</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B0A3CA6015AFFE5C66BCEA7FBA68D0E" collectingDate="1989-11" collectionCode="PBCS" collectorName="D. C. Darling &amp; Malaise Tp. &amp; H. &amp; A. Howden" country="Indonesia" county="Leuser Nat. Pk." elevation="350" latitude="3.6833334" location="Primary Rainforest" longLatPrecision="1307" longitude="97.65" municipality="Ketambe Res. Sta." pageId="6" pageNumber="181" specimenCount="1" stateProvince="Sumatera Utara">
<specimenCount id="9D64FD72015AFFE5C66BCEA7FBDA8EB3" box="[1082,1137,392,415]" count="1" pageId="6" pageNumber="181" type="generic">1 ex.</specimenCount>
:
<collectingCountry id="F375766B015AFFE5C62BCEA7FB4C8EB3" box="[1146,1255,392,415]" name="Indonesia" pageId="6" pageNumber="181">Indonesia</collectingCountry>
:
<collectingRegion id="49A6F819015AFFE5C6A1CEA7FAE68EB3" box="[1264,1357,392,415]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="181">Sumatra</collectingRegion>
,
<collectingRegion id="49A6F819015AFFE5C103CE87FC268E93" box="[850,909,424,447]" country="Indonesia" name="Aceh" pageId="6" pageNumber="181">Aceh</collectingRegion>
, G.
<collectingCounty id="62BC4E77015AFFE5C1E2CE87FBF08E93" box="[947,1115,424,447]" pageId="6" pageNumber="181">Leuser Nat. Pk.</collectingCounty>
,
<collectingMunicipality id="6BB9AC81015AFFE5C635CE87FA808E93" box="[1124,1323,424,447]" pageId="6" pageNumber="181">Ketambe Res. Sta.</collectingMunicipality>
,
<date id="FFDC103B015AFFE5C764CE87FC218ECC" pageId="6" pageNumber="181" value="1989-11">
<collectingDate id="EF98E9D3015AFFE5C764CE87FC218ECC" pageId="6" pageNumber="181" value="1989-11">xi. 1989</collectingDate>
</date>
,
<collectorName id="2697532D015AFFE5C1C2CEE6FB8E8ECC" box="[915,1061,457,480]" pageId="6" pageNumber="181">D. C. Darling</collectorName>
,
<collectorName id="2697532D015AFFE5C67FCEE6FB068ECC" box="[1070,1197,457,480]" pageId="6" pageNumber="181">Malaise Tp.</collectorName>
<quantity id="4C9A9B1E015AFFE5C6E0CEE6FB5A8ECC" box="[1201,1265,457,480]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="181" unit="m" value="350.0">
<elevation id="004FD1C8015AFFE5C6E0CEE6FB5A8ECC" box="[1201,1265,457,480]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="181" unit="m" value="350.0">350m</elevation>
</quantity>
,
<location id="8EBD6020015AFFE5C6ABCEE6FC6E8D2D" LSID="urn:lsid:plazi:treatment:03CB87ED015AFFE5C103CE2BFBA88A38:8EBD6020015AFFE5C6ABCEE6FC6E8D2D" country="Indonesia" county="Leuser Nat. Pk." latitude="3.6833334" longLatPrecision="1307" longitude="97.65" municipality="Ketambe Res. Sta." name="Primary Rainforest" pageId="6" pageNumber="181" stateProvince="Sumatera Utara">Primary Rainforest</location>
,
<geoCoordinate id="EE56503C015AFFE5C180CEC5FB898D2D" box="[977,1058,490,513]" degrees="3" direction="north" minutes="41" orientation="latitude" pageId="6" pageNumber="181" precision="925" value="3.6833334">3°41´N</geoCoordinate>
,
<geoCoordinate id="EE56503C015AFFE5C67ECEC5FB218D2D" box="[1071,1162,490,513]" degrees="97" direction="east" minutes="39" orientation="longitude" pageId="6" pageNumber="181" precision="925" value="97.65">97°39´E</geoCoordinate>
,
<collectorName id="2697532D015AFFE5C6C6CEC5FAFB8D2D" box="[1175,1360,490,513]" pageId="6" pageNumber="181">H. &amp; A. Howden</collectorName>
collection [
<collectionCode id="ED73AE3E015AFFE5C190CD24FBAC8D0E" box="[961,1031,523,546]" pageId="6" pageNumber="181">PBCS</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B0A3CA6015AFFE5C647CD24FBE58DA8" collectingDate="2014-12-19" collectionCode="Y" collectorName="Stanislav Jakl" country="Indonesia" elevation="1325" location="Env." municipality="Mt. Sanggul" pageId="6" pageNumber="181" specimenCount="1" stateProvince="Sumatera Barat">
<specimenCount id="9D64FD72015AFFE5C647CD24FBE28D0E" box="[1046,1097,523,546]" count="1" pageId="6" pageNumber="181" type="generic">1 ex.</specimenCount>
:
<collectingCountry id="F375766B015AFFE5C600CD24FB1C8D0E" box="[1105,1207,523,546]" name="Indonesia" pageId="6" pageNumber="181">Indonesia</collectingCountry>
,
<collectingRegion id="49A6F819015AFFE5C6EFCD24FAE78D0E" box="[1214,1356,523,546]" country="Indonesia" name="Sumatera Barat" pageId="6" pageNumber="181">West Sumatra</collectingRegion>
,
<collectingMunicipality id="6BB9AC81015AFFE5C103CD03FC7E8D6F" box="[850,981,556,579]" pageId="6" pageNumber="181">Mt. Sanggul</collectingMunicipality>
, cca
<locationDeviation id="9A6F2FD9015AFFE5C654CD03FAFB8D6F" box="[1029,1360,556,579]" location="Payakumbuh Landai" pageId="6" pageNumber="181">
<quantity id="4C9A9B1E015AFFE5C654CD03FBEA8D6F" box="[1029,1089,556,579]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="181" unit="km" value="35.0">35km</quantity>
N of Payakumbuh Landai
</locationDeviation>
vill.
<location id="8EBD6020015AFFE5C1D8CD62FC108D48" LSID="urn:lsid:plazi:treatment:03CB87ED015AFFE5C103CE2BFBA88A38:8EBD6020015AFFE5C1D8CD62FC108D48" box="[905,955,589,612]" country="Indonesia" municipality="Mt. Sanggul" name="Env." pageId="6" pageNumber="181" stateProvince="Sumatera Barat">Env.</location>
,
<quantity id="4C9A9B1E015AFFE5C19DCD62FBF28D48" box="[972,1113,589,612]" metricMagnitude="3" metricUnit="m" metricValue="1.325" metricValueMax="1.4" metricValueMin="1.25" pageId="6" pageNumber="181" unit="m" value="1325.0" valueMax="1400.0" valueMin="1250.0">
<elevation id="004FD1C8015AFFE5C19DCD62FBF28D48" box="[972,1113,589,612]" metricMagnitude="3" metricUnit="m" metricValue="1.325" metricValueMax="1.4" metricValueMin="1.25" pageId="6" pageNumber="181" unit="m" value="1325.0" valueMax="1400.0" valueMin="1250.0">1250­1400m</elevation>
</quantity>
alt.,
<date id="FFDC103B015AFFE5C6C9CD62FABD8D48" box="[1176,1302,589,612]" pageId="6" pageNumber="181" value="2014-12-19">
<collectingDate id="EF98E9D3015AFFE5C6C9CD62FABD8D48" box="[1176,1302,589,612]" pageId="6" pageNumber="181" value="2014-12-19">19.xii.2014</collectingDate>
</date>
, leg.
<collectorName id="2697532D015AFFE5C103CD42FC4E8DA8" box="[850,997,621,644]" pageId="6" pageNumber="181">Stanislav Jakl</collectorName>
[
<emphasis id="B916EAE9015AFFE5C1A5CD43FBAF8DA9" bold="true" box="[1012,1028,620,645]" pageId="6" pageNumber="181">
<collectionCode id="ED73AE3E015AFFE5C1A5CD43FBAF8DA9" box="[1012,1028,620,645]" pageId="6" pageNumber="181">Ÿ</collectionCode>
</emphasis>
MCN]
</materialsCitation>
;
<materialsCitation id="3B0A3CA6015AFFE5C609CD42FB718DCB" collectingDate="1996-06-24" collectingDateMax="1996-07-01" collectingDateMin="1996-06-24" collectionCode="ISEA" country="Malaysia" location="Punggul Resort" municipality="Balu" pageId="6" pageNumber="181" specimenCount="1" stateProvince="Sabah">
<specimenCount id="9D64FD72015AFFE5C609CD42FB258DA8" box="[1112,1166,621,644]" count="1" pageId="6" pageNumber="181" type="generic">1 ex.</specimenCount>
:
<collectingCountry id="F375766B015AFFE5C6C9CD42FB548DA8" box="[1176,1279,621,644]" name="Malaysia" pageId="6" pageNumber="181">Malaysia</collectingCountry>
:
<collectingRegion id="49A6F819015AFFE5C759CD42FAE68DA8" box="[1288,1357,621,644]" country="Malaysia" name="Sabah" pageId="6" pageNumber="181">Sabah</collectingRegion>
,
<collectingMunicipality id="6BB9AC81015AFFE5C103CDA1FC2C8D89" box="[850,903,654,677]" pageId="6" pageNumber="181">Balu</collectingMunicipality>
,
<location id="8EBD6020015AFFE5C1C0CDA1FB9D8D89" LSID="urn:lsid:plazi:treatment:03CB87ED015AFFE5C103CE2BFBA88A38:8EBD6020015AFFE5C1C0CDA1FB9D8D89" box="[913,1078,654,677]" country="Malaysia" municipality="Balu" name="Punggul Resort" pageId="6" pageNumber="181" stateProvince="Sabah">Punggul Resort</location>
env.
<date id="FFDC103B015AFFE5C63FCDA1FA8F8D89" box="[1134,1316,654,677]" pageId="6" pageNumber="181" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">
<collectingDate id="EF98E9D3015AFFE5C63FCDA1FA8F8D89" box="[1134,1316,654,677]" pageId="6" pageNumber="181" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">24.vi.­ 1.vii.1996</collectingDate>
</date>
, vegetation debris and forest floor litter accumulated around large treens near river [
<collectionCode id="ED73AE3E015AFFE5C6C0CDFFFB7F8DCB" box="[1169,1236,720,743]" pageId="6" pageNumber="181">ISEA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B0A3CA6015AFFE5C6B2CDFFFB7A8C65" collectingDate="2012-08-18" collectionCode="Y" collectorName="N Borneo &amp; A. Klimenko" country="Malaysia" elevation="1160" location="Trus Modi Mt." municipality="Keningau distr." pageId="6" pageNumber="181" specimenCount="1" stateProvince="Sabah">
<specimenCount id="9D64FD72015AFFE5C6B2CDFFFAB08DCB" box="[1251,1307,720,743]" count="1" pageId="6" pageNumber="181" type="generic">1 ex.</specimenCount>
:
<collectingCountry id="F375766B015AFFE5C775CDFFFC3F8C2B" name="Malaysia" pageId="6" pageNumber="181">Malaysia</collectingCountry>
,
<collectorName id="2697532D015AFFE5C1F1CDDFFBA58C2B" box="[928,1038,752,775]" pageId="6" pageNumber="181">N Borneo</collectorName>
,
<collectingRegion id="49A6F819015AFFE5C64BCDDFFBF48C2B" box="[1050,1119,752,775]" country="Malaysia" name="Sabah" pageId="6" pageNumber="181">Sabah</collectingRegion>
,
<collectingMunicipality id="6BB9AC81015AFFE5C63ACDDFFABA8C2B" box="[1131,1297,752,775]" pageId="6" pageNumber="181">Keningau distr.</collectingMunicipality>
,
<location id="8EBD6020015AFFE5C771CDDFFC1C8C04" LSID="urn:lsid:plazi:treatment:03CB87ED015AFFE5C103CE2BFBA88A38:8EBD6020015AFFE5C771CDDFFC1C8C04" country="Malaysia" municipality="Keningau distr." name="Trus Modi Mt." pageId="6" pageNumber="181" stateProvince="Sabah">Trus Modi Mt.</location>
,
<quantity id="4C9A9B1E015AFFE5C1EECC3EFBA78C04" box="[959,1036,785,808]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="6" pageNumber="181" unit="m" value="1160.0">
<elevation id="004FD1C8015AFFE5C1EECC3EFBA78C04" box="[959,1036,785,808]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="6" pageNumber="181" unit="m" value="1160.0">1160m</elevation>
</quantity>
.,
<date id="FFDC103B015AFFE5C649CC3EFB3C8C04" box="[1048,1175,785,808]" pageId="6" pageNumber="181" value="2012-08-18">
<collectingDate id="EF98E9D3015AFFE5C649CC3EFB3C8C04" box="[1048,1175,785,808]" pageId="6" pageNumber="181" value="2012-08-18">18.viii.2012</collectingDate>
</date>
, leg.
<collectorName id="2697532D015AFFE5C699CC3EFAFB8C04" box="[1224,1360,785,808]" pageId="6" pageNumber="181">A. Klimenko</collectorName>
[
<emphasis id="B916EAE9015AFFE5C10ACC1EFCC08C66" bold="true" box="[859,875,817,842]" pageId="6" pageNumber="181">
<collectionCode id="ED73AE3E015AFFE5C10ACC1EFCC08C66" box="[859,875,817,842]" pageId="6" pageNumber="181">Ÿ</collectionCode>
</emphasis>
MCN] [photographed specimen]
</materialsCitation>
;
<materialsCitation id="3B0A3CA6015AFFE5C68FCC1DFC398CA7" collectorName="Kinabalu N. P. &amp; Poring" country="Malaysia" elevation="500" location="Borneo" pageId="6" pageNumber="181" specimenCount="1" stateProvince="Sabah">
<specimenCount id="9D64FD72015AFFE5C68FCC1DFABC8C65" box="[1246,1303,818,841]" count="1" pageId="6" pageNumber="181" type="generic">1 ex.</specimenCount>
:
<collectingCountry id="F375766B015AFFE5C775CC1DFC3F8C46" name="Malaysia" pageId="6" pageNumber="181">Malaysia</collectingCountry>
,
<location id="8EBD6020015AFFE5C1F3CC7CFC5F8C46" LSID="urn:lsid:plazi:treatment:03CB87ED015AFFE5C103CE2BFBA88A38:8EBD6020015AFFE5C1F3CC7CFC5F8C46" box="[930,1012,851,874]" country="Malaysia" name="Borneo" pageId="6" pageNumber="181" stateProvince="Sabah">Borneo</location>
,
<collectingRegion id="49A6F819015AFFE5C652CC7CFBE38C46" box="[1027,1096,851,874]" country="Malaysia" name="Sabah" pageId="6" pageNumber="181">Sabah</collectingRegion>
,
<collectorName id="2697532D015AFFE5C607CC7CFB5B8C46" box="[1110,1264,851,874]" pageId="6" pageNumber="181">Kinabalu N.P.</collectorName>
,
<collectorName id="2697532D015AFFE5C753CC7CFAE68C46" box="[1282,1357,851,874]" pageId="6" pageNumber="181">Poring</collectorName>
,
<quantity id="4C9A9B1E015AFFE5C103CC5BFC398CA7" box="[850,914,884,907]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="181" unit="m" value="500.0">
<specimenCount id="9D64FD72015AFFE5C103CC5BFC398CA7" box="[850,914,884,907]" count="500" pageId="6" pageNumber="181" type="generic">
<elevation id="004FD1C8015AFFE5C103CC5BFC398CA7" box="[850,914,884,907]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="181" unit="m" value="500.0">500m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation id="3B0A3CA6015AFFE5C1C2CC5BFC1E8C80" collectingDate="1997-12-02" collectionCode="Y" collectorName="D. Grimm" country="Malaysia" pageId="6" pageNumber="181" specimenCount="1" stateProvince="Sabah">
., at light,
<date id="FFDC103B015AFFE5C646CC5BFB258CA7" box="[1047,1166,884,907]" pageId="6" pageNumber="181" value="1997-12-02">
<collectingDate id="EF98E9D3015AFFE5C646CC5BFB258CA7" box="[1047,1166,884,907]" pageId="6" pageNumber="181" value="1997-12-02">2.xii.1997</collectingDate>
</date>
, leg.
<collectorName id="2697532D015AFFE5C689CC5BFAFB8CA7" box="[1240,1360,884,907]" pageId="6" pageNumber="181">D. Grimm</collectorName>
[
<emphasis id="B916EAE9015AFFE5C10ACCBCFCC08C80" bold="true" box="[859,875,915,940]" pageId="6" pageNumber="181">
<collectionCode id="ED73AE3E015AFFE5C10ACCBCFCC08C80" box="[859,875,915,940]" pageId="6" pageNumber="181">Ÿ</collectionCode>
</emphasis>
MCN]
</materialsCitation>
.
</paragraph>
<paragraph id="8BDD36FB015AFFE5C103CCF9FBA88A38" blockId="6.[850,1361,982,1300]" pageId="6" pageNumber="181">
<emphasis id="B916EAE9015AFFE5C103CCF9FBA68CC2" bold="true" box="[850,1037,982,1006]" pageId="6" pageNumber="181">Short comment.</emphasis>
Previously known from
<collectingRegion id="49A6F819015AFFE5C770CCF9FAFA8CC1" box="[1313,1361,982,1005]" country="Indonesia" name="Jawa Timur" pageId="6" pageNumber="181">Java</collectingRegion>
and Borneo. Additional records for Borneo. First records for
<collectingRegion id="49A6F819015AFFE5C196CB37FB898B03" box="[967,1058,1048,1071]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="181">Sumatra</collectingRegion>
. Specimens from Borneo and
<collectingRegion id="49A6F819015AFFE5C103CB17FC028B63" box="[850,937,1080,1103]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="181">Sumatra</collectingRegion>
are almost identique. Only one very weak difference were observed in caudal bulbs structure, i.e. sinuation between median and internal protrusions is slightly less distinct in population from
<collectingRegion id="49A6F819015AFFE5C1DCCB93FC418BFF" box="[909,1002,1212,1235]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="181">Sumatra</collectingRegion>
. In opinion of authors such small feature is not enough to describe this population as separate taxa.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,303 @@
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<mods:title id="FA5EE4C204EC71762F7F277137B402C2">Studies on genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with description of two new species from Southeast Asia</mods:title>
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<mods:date id="5141C3A3B4A3034EBE879F1F161A29C6">2022</mods:date>
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<treatment id="03CB87ED015DFFE0C157CE2BFCDE8B43" LSID="urn:lsid:plazi:treatment:03CB87ED015DFFE0C157CE2BFCDE8B43" httpUri="http://treatment.plazi.org/id/03CB87ED015DFFE0C157CE2BFCDE8B43" lastPageId="3" lastPageNumber="178" pageId="1" pageNumber="176">
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<paragraph id="8BDD36FB015DFFE2C157CE2BFBB08E30" blockId="1.[774,1051,260,316]" box="[774,1051,260,284]" pageId="1" pageNumber="176">
<heading id="D0958197015DFFE2C157CE2BFBB08E30" box="[774,1051,260,284]" fontSize="10" level="2" pageId="1" pageNumber="176" reason="7">
<taxonomicName id="4C624D78015DFFE2C157CE2BFC6A8E30" ID-CoL="9WHZZ" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[774,961,260,284]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="176" phylum="Arthropoda" rank="species" species="merkli" status="sp. nov.">
<emphasis id="B916EAE9015DFFE2C157CE2BFC6A8E30" bold="true" box="[774,961,260,284]" italics="true" pageId="1" pageNumber="176">Rhyparus merkli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2255792015DFFE2C196CE2AFBB08E30" box="[967,1051,261,284]" pageId="1" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3786570015DFFE2C157CE0AFC7E8E10" box="[774,981,293,316]" pageId="1" pageNumber="176" type="description">
<paragraph id="8BDD36FB015DFFE2C157CE0AFC7E8E10" blockId="1.[774,1051,260,316]" box="[774,981,293,316]" pageId="1" pageNumber="176">
(
<figureCitation id="13592A7E015DFFE2C15FCE0AFCC48E10" box="[782,879,293,316]" captionStart="Figs 13" captionStartId="2.[302,348,1693,1716]" captionTargetBox="[441,1217,692,1651]" captionTargetId="figure-267@2.[430,1225,688,1659]" captionTargetPageId="2" captionText="Figs 13. Rhyparus merkli sp. nov.,, holotype: 1 dorsal view, 2 ventral view, 3 lateral view (scale lines: 1.0 mm)." pageId="1" pageNumber="176">Figs 13</figureCitation>
,
<figureCitation id="13592A7E015DFFE2C12ACE0AFC218E10" box="[891,906,293,316]" captionStart="Figs 7, 8" captionStartId="5.[774,820,724,747]" captionTargetBox="[776,1283,190,691]" captionTargetId="figure-490@5.[774,1284,189,691]" captionTargetPageId="5" captionText="Figs 7, 8. Heads: 7 Rhyparus merkli sp. nov., 8 R. jakli sp. nov. (scale lines: Fig. 7: 0.5mm, Fig. 8: 1.0mm)." pageId="1" pageNumber="176">7</figureCitation>
,
<figureCitation id="13592A7E015DFFE2C1C7CE0AFC608E10" box="[918,971,293,316]" captionStart="Figs 912" captionStartId="5.[774,819,1595,1618]" captionTargetBox="[779,1282,888,1559]" captionTargetId="figure-488@5.[774,1284,882,1561]" captionTargetPageId="5" captionText="Figs 912.Aedeagi and epipharyngi:9 aedeagus of Rhyparus merkli sp. nov., 10 aedeagus of R. jakli sp. nov., 11 epipharynx of R. merkli sp. nov., 12 epipharynx of R. jakli sp. nov. (scale lines: Figs 9, 10: 0.5mm, Figs 11, 12: 1.0mm)." pageId="1" pageNumber="176">9, 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3786570015DFFE2C157CE49FB478C65" pageId="1" pageNumber="176" type="materials_examined">
<paragraph id="8BDD36FB015DFFE2C157CE49FC668EB3" blockId="1.[774,1284,358,415]" pageId="1" pageNumber="176">
<materialsCitation id="3B0A3CA6015DFFE2C157CE49FC628EB3" country="Malaysia" location="Tanah Rata" municipality="Cameron Highlands" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<emphasis id="B916EAE9015DFFE2C157CE49FC318E52" bold="true" box="[774,922,358,382]" pageId="1" pageNumber="176">
<typeStatus id="54D98859015DFFE2C157CE49FC968E52" box="[774,829,358,382]" pageId="1" pageNumber="176">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F375766B015DFFE2C1CFCE48FBAA8E52" box="[926,1025,359,382]" name="Malaysia" pageId="1" pageNumber="176">Malaysia</collectingCountry>
,
<collectingRegion id="49A6F819015DFFE2C658CE48FBF28E52" box="[1033,1113,359,382]" country="Malaysia" name="Pahang" pageId="1" pageNumber="176">Pahang</collectingRegion>
,
<collectingMunicipality id="6BB9AC81015DFFE2C633CE48FCE98EB3" pageId="1" pageNumber="176">Cameron Highlands</collectingMunicipality>
,
<location id="8EBD6020015DFFE2C11DCEA7FC628EB3" LSID="urn:lsid:plazi:treatment:03CB87ED015DFFE0C157CE2BFCDE8B43:8EBD6020015DFFE2C11DCEA7FC628EB3" box="[844,969,392,415]" country="Malaysia" municipality="Cameron Highlands" name="Tanah Rata" pageId="1" pageNumber="176" stateProvince="Pahang">Tanah Rata</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDD36FB015DFFE2C157CEE6FB478C65" blockId="1.[774,1285,457,841]" pageId="1" pageNumber="176">
<emphasis id="B916EAE9015DFFE2C157CEE6FC028ECD" bold="true" box="[774,937,457,481]" pageId="1" pageNumber="176">Type material.</emphasis>
<materialsCitation id="3B0A3CA6015DFFE2C1FCCEE6FC178D6F" collectingDate="1995-03-23" collectingDateMax="1995-03-31" collectingDateMin="1995-03-23" collectionCode="ISEA" collectorName="O. Merkl" country="Malaysia" location="Tanah Rata" municipality="Cameron Highlands" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<typeStatus id="54D98859015DFFE2C1FCCEE6FBA48ECC" box="[941,1039,457,480]" pageId="1" pageNumber="176" type="holotype">Holotype</typeStatus>
():
<collectingCountry id="F375766B015DFFE2C614CEE6FB038ECC" box="[1093,1192,457,480]" name="Malaysia" pageId="1" pageNumber="176">Malaysia</collectingCountry>
,
<collectingRegion id="49A6F819015DFFE2C6E0CEE6FAAA8ECC" box="[1201,1281,457,480]" country="Malaysia" name="Pahang" pageId="1" pageNumber="176">Pahang</collectingRegion>
,
<collectingMunicipality id="6BB9AC81015DFFE2C157CEC5FC4E8D2D" box="[774,997,490,513]" pageId="1" pageNumber="176">Cameron Highlands</collectingMunicipality>
,
<location id="8EBD6020015DFFE2C1A0CEC5FBDB8D2D" LSID="urn:lsid:plazi:treatment:03CB87ED015DFFE0C157CE2BFCDE8B43:8EBD6020015DFFE2C1A0CEC5FBDB8D2D" box="[1009,1136,490,513]" country="Malaysia" municipality="Cameron Highlands" name="Tanah Rata" pageId="1" pageNumber="176" stateProvince="Pahang">Tanah Rata</location>
, from illuminated, white­washed walls,
<date id="FFDC103B015DFFE2C661CD24FB798D0E" box="[1072,1234,523,546]" pageId="1" pageNumber="176" value="1995-03-23" valueMax="1995-03-31" valueMin="1995-03-23">
<collectingDate id="EF98E9D3015DFFE2C661CD24FB798D0E" box="[1072,1234,523,546]" pageId="1" pageNumber="176" value="1995-03-23" valueMax="1995-03-31" valueMin="1995-03-23">23.­ 31.iii.1995</collectingDate>
</date>
, leg.
<collectorName id="2697532D015DFFE2C157CD03FCCD8D6F" box="[774,870,556,579]" pageId="1" pageNumber="176">O. Merkl</collectorName>
[
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]
</materialsCitation>
.
<materialsCitation id="3B0A3CA6015DFFE2C192CD03FB828DEA" collectingDate="1995-03-29" collectionCode="ISEA, DUBC" collectorName="O. Merkl &amp; Szikossy" country="Malaysia" location="Cameron Highlands" pageId="1" pageNumber="176" specimenCount="2" stateProvince="Pahang" typeStatus="paratype">
<typeStatus id="54D98859015DFFE2C192CD03FB818D6F" box="[963,1066,556,579]" pageId="1" pageNumber="176" type="paratype">Paratypes</typeStatus>
(6 exx.): 4 exx.:
<collectingCountry id="F375766B015DFFE2C689CD03FCEE8D48" name="Malaysia" pageId="1" pageNumber="176">Malaysia</collectingCountry>
,
<collectingRegion id="49A6F819015DFFE2C11FCD62FC368D48" box="[846,925,589,612]" country="Malaysia" name="Pahang" pageId="1" pageNumber="176">Pahang</collectingRegion>
,
<location id="8EBD6020015DFFE2C1F4CD62FBD38D48" LSID="urn:lsid:plazi:treatment:03CB87ED015DFFE0C157CE2BFCDE8B43:8EBD6020015DFFE2C1F4CD62FBD38D48" box="[933,1144,589,612]" country="Malaysia" name="Cameron Highlands" pageId="1" pageNumber="176" stateProvince="Pahang">Cameron Highlands</location>
,
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<quantity id="4C9A9B1E015DFFE2C62ECD62FB048D48" box="[1151,1199,589,612]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="176" unit="km" value="2.0">2km</quantity>
S Tanah Rata on Tapah Road
</locationDeviation>
, montane rainforest,
<date id="FFDC103B015DFFE2C697CD42FC958D89" pageId="1" pageNumber="176" value="1995-03-29">
<collectingDate id="EF98E9D3015DFFE2C697CD42FC958D89" pageId="1" pageNumber="176" value="1995-03-29">29.iii. 1995</collectingDate>
</date>
, leg.
<collectorName id="2697532D015DFFE2C12ACDA1FC4B8D89" box="[891,992,654,677]" pageId="1" pageNumber="176">O. Merkl</collectorName>
&amp;
<collectorName id="2697532D015DFFE2C654CDA1FBCE8D89" box="[1029,1125,654,677]" pageId="1" pageNumber="176">Szikossy</collectorName>
[2 exx.
<collectionCode id="ED73AE3E015DFFE2C693CDA1FAAB8D89" box="[1218,1280,654,677]" pageId="1" pageNumber="176">ISEA</collectionCode>
,
<specimenCount id="9D64FD72015DFFE2C157CD80FC908DEA" box="[774,827,687,710]" count="1" pageId="1" pageNumber="176" type="generic">1 ex.</specimenCount>
<emphasis id="B916EAE9015DFFE2C111CD82FCFB8DEA" bold="true" box="[832,848,685,710]" pageId="1" pageNumber="176">Ÿ</emphasis>
MCN,
<specimenCount id="9D64FD72015DFFE2C1CBCD80FC668DEA" box="[922,973,687,710]" count="1" pageId="1" pageNumber="176" type="generic">1 ex.</specimenCount>
<collectionCode id="ED73AE3E015DFFE2C183CD80FB888DEA" box="[978,1059,687,710]" pageId="1" pageNumber="176">DUBC</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B0A3CA6015DFFE2C663CD80FC158C04" collectingDate="1995-03-21" collectingDateMax="1995-04-02" collectingDateMin="1995-03-21" collectionCode="ISEA" collectorName="O. Merkl" country="Malaysia" location="Tanah Rata" municipality="Cameron Highlands" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Pahang" typeStatus="paratype">
<specimenCount id="9D64FD72015DFFE2C663CD80FBC28DEA" box="[1074,1129,687,710]" count="1" pageId="1" pageNumber="176" type="generic">1 ex.</specimenCount>
:
<collectingCountry id="F375766B015DFFE2C623CD80FB738DEA" box="[1138,1240,687,710]" name="Malaysia" pageId="1" pageNumber="176">Malaysia</collectingCountry>
,
<collectingRegion id="49A6F819015DFFE2C6B0CD80FC968DCB" country="Malaysia" name="Pahang" pageId="1" pageNumber="176">Pahang</collectingRegion>
,
<collectingMunicipality id="6BB9AC81015DFFE2C119CDFFFB888DCB" box="[840,1059,720,743]" pageId="1" pageNumber="176">Cameron Highlands</collectingMunicipality>
,
<location id="8EBD6020015DFFE2C67FCDFFFB018DCB" LSID="urn:lsid:plazi:treatment:03CB87ED015DFFE0C157CE2BFCDE8B43:8EBD6020015DFFE2C67FCDFFFB018DCB" box="[1070,1194,720,743]" country="Malaysia" municipality="Cameron Highlands" name="Tanah Rata" pageId="1" pageNumber="176" stateProvince="Pahang">Tanah Rata</location>
, edge of degraded rainforest, at light,
<date id="FFDC103B015DFFE2C67CCDDFFB7D8C2B" box="[1069,1238,752,775]" pageId="1" pageNumber="176" value="1995-03-21" valueMax="1995-04-02" valueMin="1995-03-21">
<collectingDate id="EF98E9D3015DFFE2C67CCDDFFB7D8C2B" box="[1069,1238,752,775]" pageId="1" pageNumber="176" value="1995-03-21" valueMax="1995-04-02" valueMin="1995-03-21">21.iii.­ 2.iv.1995</collectingDate>
</date>
, leg.
<collectorName id="2697532D015DFFE2C157CC3EFCC38C04" box="[774,872,785,808]" pageId="1" pageNumber="176">O. Merkl</collectorName>
[
<collectionCode id="ED73AE3E015DFFE2C122CC3EFC1C8C04" box="[883,951,785,808]" pageId="1" pageNumber="176">ISEA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B0A3CA6015DFFE2C196CC3EFB438C65" collectingDate="1985-05" collectionCode="ISEA" collectorName="Wong" country="Malaysia" location="Cameron Highlands" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Perak" typeStatus="paratype">
<specimenCount id="9D64FD72015DFFE2C196CC3EFC558C04" box="[967,1022,785,808]" count="1" pageId="1" pageNumber="176" type="generic">1 ex.</specimenCount>
: W
<collectingCountry id="F375766B015DFFE2C674CC3EFB278C04" box="[1061,1164,785,808]" name="Malaysia" pageId="1" pageNumber="176">Malaysia</collectingCountry>
:
<collectingRegion id="49A6F819015DFFE2C6C5CC3EFB7F8C04" box="[1172,1236,785,808]" country="Malaysia" name="Perak" pageId="1" pageNumber="176">Perak</collectingRegion>
,
<location id="8EBD6020015DFFE2C68FCC3EFC688C65" LSID="urn:lsid:plazi:treatment:03CB87ED015DFFE0C157CE2BFCDE8B43:8EBD6020015DFFE2C68FCC3EFC688C65" country="Malaysia" name="Cameron Highlands" pageId="1" pageNumber="176" stateProvince="Perak">Cameron Highlands</location>
,
<date id="FFDC103B015DFFE2C19FCC1DFBB38C65" box="[974,1048,818,841]" pageId="1" pageNumber="176" value="1985-05">
<collectingDate id="EF98E9D3015DFFE2C19FCC1DFBB38C65" box="[974,1048,818,841]" pageId="1" pageNumber="176" value="1985-05">v.1985</collectingDate>
</date>
, leg.
<collectorName id="2697532D015DFFE2C600CC1DFB3B8C65" box="[1105,1168,818,841]" pageId="1" pageNumber="176">Wong</collectorName>
[
<collectionCode id="ED73AE3E015DFFE2C6CCCC1DFB4A8C65" box="[1181,1249,818,841]" pageId="1" pageNumber="176">ISEA</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3786570015DFFE0C157CC5CFDA78935" lastPageId="3" lastPageNumber="178" pageId="1" pageNumber="176" type="description">
<paragraph id="8BDD36FB015DFFE2C157CC5CFAAD8B03" blockId="1.[774,1286,883,1071]" pageId="1" pageNumber="176">
<emphasis id="B916EAE9015DFFE2C157CC5CFC388CA7" bold="true" box="[774,915,883,907]" pageId="1" pageNumber="176">Description.</emphasis>
Dorsum (
<figureCitation id="13592A7E015DFFE2C655CC5BFBEE8CA7" box="[1028,1093,884,907]" captionStart="Figs 13" captionStartId="2.[302,348,1693,1716]" captionTargetBox="[441,1217,692,1651]" captionTargetId="figure-267@2.[430,1225,688,1659]" captionTargetPageId="2" captionText="Figs 13. Rhyparus merkli sp. nov.,, holotype: 1 dorsal view, 2 ventral view, 3 lateral view (scale lines: 1.0 mm)." pageId="1" pageNumber="176">Fig. 1</figureCitation>
). Body length of
<typeStatus id="54D98859015DFFE2C157CCBAFCCF8C80" box="[774,868,917,940]" pageId="1" pageNumber="176" type="holotype">holotype</typeStatus>
6.0mm. Moderate in size as member of the genus. Elongate­oval, except of preapical parts of elytra, inner part of legs and strongly convex elements as costae mat, brownishblack; except punctures, tops of costae and legs glabrous.
</paragraph>
<paragraph id="8BDD36FB015DFFE2C157CB76FB218AF5" blockId="1.[774,1285,1113,1497]" pageId="1" pageNumber="176">
Head (
<figureCitation id="13592A7E015DFFE2C11BCB76FC2C8B5C" box="[842,903,1113,1136]" captionStart="Figs 7, 8" captionStartId="5.[774,820,724,747]" captionTargetBox="[776,1283,190,691]" captionTargetId="figure-490@5.[774,1284,189,691]" captionTargetPageId="5" captionText="Figs 7, 8. Heads: 7 Rhyparus merkli sp. nov., 8 R. jakli sp. nov. (scale lines: Fig. 7: 0.5mm, Fig. 8: 1.0mm)." pageId="1" pageNumber="176">Fig. 7</figureCitation>
) trapezoidal, clypeus anteriorly truncate, with indistinct teeth on sides, laterally distinctly emarginated and next with widely rounded angles, relatively distinctly separated from distinctly protruding, rounded genae. Clypeocentral disc distinctly convex, ringed by deep grove, with pair of distinct, moderately convex, shiny only on top, somewhat longitudinal tubercles. Clypeocentral disc relatively sparsely punctate. Frons with four short, relatively narrow longitudinal costae. Punctation of head relatively regular, all punctures bearing short macrosetae.
</paragraph>
<paragraph id="8BDD36FB015DFFE2C157C92BFBF98993" blockId="1.[774,1285,1540,1727]" pageId="1" pageNumber="176">
Epipharynx (
<figureCitation id="13592A7E015DFFE2C1DEC92BFC728937" box="[911,985,1540,1563]" captionStart="Figs 912" captionStartId="5.[774,819,1595,1618]" captionTargetBox="[779,1282,888,1559]" captionTargetId="figure-488@5.[774,1284,882,1561]" captionTargetPageId="5" captionText="Figs 912.Aedeagi and epipharyngi:9 aedeagus of Rhyparus merkli sp. nov., 10 aedeagus of R. jakli sp. nov., 11 epipharynx of R. merkli sp. nov., 12 epipharynx of R. jakli sp. nov. (scale lines: Figs 9, 10: 0.5mm, Figs 11, 12: 1.0mm)." pageId="1" pageNumber="176">Fig. 11</figureCitation>
) distinctly transverse, rather distinctly sinuate at middle, corypha with several long celter, with two lateral lobes bearing tuft of smaller and finer celtes. Epitorma broad, somewhat oval­shaped, with dense chaetopedia laterally. Tormae moderately long.
</paragraph>
<paragraph id="8BDD36FB015EFFE1C37FCFEEFE5D8D14" blockId="2.[302,813,193,568]" pageId="2" pageNumber="177">Pronotum transverse, with six distinct, costae and seven longitudinal intercostae; on each side of costae there is row of short and small yellowish macrosetae. Lateral margin with two distinctly protruding lobes; anterior lobes sharply triangular, much more developed than broadly triangular posterior. Paramedian costae continuous, almost parallel. Discolateral costae relatively widely interrupted in apical half. Median intercosta with relatively dense, but concentrated medially, medium sized punctures, lateral intercostae almost without punctures.</paragraph>
<paragraph id="8BDD36FB015EFFE1C37FCD4EFBA68D54" blockId="2.[302,650,609,632]" lastBlockId="2.[850,1361,193,632]" pageId="2" pageNumber="177">Scutellum almost imperceptible. Elytra elongate, relatively wide, widest before the middle. Each elytron with five distinct costae and five intercostae; on each side of costae there is row of short and small yellowish macrosetae. Fourth costae widened basally; covered by relatively dense, moderately large punctures. Intercostae flat, all of them with two rows of medium sized punctures. Postdiscal bulbs relatively small. Caudal bulbs well developed; external protrusion distinctly divided from median and internal which are fused into one and visible as one not so large elongately­spherical bulb. Pygidium with elevated area basally; medially with relatively deep grooves on sides.</paragraph>
<caption id="DF1D6673015EFFE1C37FC9B2FDBD89F8" pageId="2" pageNumber="177" startId="2.[302,348,1693,1716]" targetBox="[441,1217,692,1651]" targetPageId="2" targetType="figure">
<paragraph id="8BDD36FB015EFFE1C37FC9B2FDBD89F8" blockId="2.[302,1360,1693,1748]" pageId="2" pageNumber="177">
Figs 13.
<taxonomicName id="4C624D78015EFFE1C3C8C9B1FDE78999" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[409,588,1694,1717]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="177" phylum="Arthropoda" rank="species" species="merkli" status="sp. nov.">
<emphasis id="B916EAE9015EFFE1C3C8C9B1FDE78999" box="[409,588,1694,1717]" italics="true" pageId="2" pageNumber="177">Rhyparus merkli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2255792015EFFE1C003C9B2FD0F8998" box="[594,676,1693,1716]" pageId="2" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
,, holotype: 1 dorsal view, 2 ventral view, 3 lateral view (scale lines: 1.0 mm).
</paragraph>
</caption>
<paragraph id="8BDD36FB015FFFE0C025CFBEFCDE8F88" blockId="3.[628,885,145,164]" box="[628,885,145,164]" pageId="3" pageNumber="178">
<emphasis id="B916EAE9015FFFE0C025CFBEFCDE8F88" box="[628,885,145,164]" italics="true" pageId="3" pageNumber="178">Minkina Ł., Anichtchenko A.</emphasis>
</paragraph>
<paragraph id="8BDD36FB015FFFE0C2B2CFEEFE2B8FF4" blockId="3.[227,384,193,216]" box="[227,384,193,216]" pageId="3" pageNumber="178">Macropterous.</paragraph>
<paragraph id="8BDD36FB015FFFE0C2B2CE2CFE9D8EF3" blockId="3.[227,737,259,479]" pageId="3" pageNumber="178">
Venter matt, brownishblack (
<figureCitation id="13592A7E015FFFE0C05ECE2CFDE48E36" box="[527,591,259,282]" captionStart="Figs 13" captionStartId="2.[302,348,1693,1716]" captionTargetBox="[441,1217,692,1651]" captionTargetId="figure-267@2.[430,1225,688,1659]" captionTargetPageId="2" captionText="Figs 13. Rhyparus merkli sp. nov.,, holotype: 1 dorsal view, 2 ventral view, 3 lateral view (scale lines: 1.0 mm)." pageId="3" pageNumber="178">Fig. 2</figureCitation>
). Meso­metaventral plate flat, with distinct, relatively shallow, wide median impression in basal 2/3; relatively densely covered by medium sized punctures bearing very thin macrosetae.Abdomal ventrites separated by relatively large, irregular in shape impressed sutures.
</paragraph>
<paragraph id="8BDD36FB015FFFE0C2B2CD26FE498CE7" blockId="3.[227,737,521,971]" pageId="3" pageNumber="178">Legs sturdy, matt, with a lot of small punctured with eracted short setae. Mesofemora with two distinct teeth on basal border; metafemora with two indistinct teeth there. All femora with dense, not so fine, quite regular punctation bearing very thin macrosetation. Protibia tridentate apically, straight, with distinct, relatively large first dent. Mesotibiae narrow at base, widest before middle of length, and next narrowed to apex; at apex with widened apical projection. Metatibiae with similar structure as mesotibiae, but widest nearby middle of length and with more distinctly developed apical projection. Metatibiae as general with distinct sinuations.</paragraph>
<paragraph id="8BDD36FB015FFFE0C2B2CCDAFE208BA3" blockId="3.[227,737,1013,1167]" pageId="3" pageNumber="178">
Aedeagus (
<figureCitation id="13592A7E015FFFE0C30ACCDAFE308B20" box="[347,411,1013,1036]" captionStart="Figs 912" captionStartId="5.[774,819,1595,1618]" captionTargetBox="[779,1282,888,1559]" captionTargetId="figure-488@5.[774,1284,882,1561]" captionTargetPageId="5" captionText="Figs 912.Aedeagi and epipharyngi:9 aedeagus of Rhyparus merkli sp. nov., 10 aedeagus of R. jakli sp. nov., 11 epipharynx of R. merkli sp. nov., 12 epipharynx of R. jakli sp. nov. (scale lines: Figs 9, 10: 0.5mm, Figs 11, 12: 1.0mm)." pageId="3" pageNumber="178">Fig. 9</figureCitation>
) Phallobase much longer than parameres, moderately de­curved toward apex in lateral view. Parameres small, thin, semitransparent, visible as weak protrusion of each side of median lobe.
</paragraph>
<paragraph id="8BDD36FB015FFFE0C2B2CB95FDE18A59" blockId="3.[227,738,1210,1397]" pageId="3" pageNumber="178">
<emphasis id="B916EAE9015FFFE0C2B2CB95FECA8BFE" bold="true" box="[227,353,1210,1234]" pageId="3" pageNumber="178">Variability.</emphasis>
Body length: 5.8­ 6.3mm. Punctation of intercostae of pronotum is slightly variable. Shape of caudal bulbs is slightly variable: sinuation between median and internal protrusion is somewhat variable, sometimes it is well visible, and sometimes sinuation is weak.
</paragraph>
<paragraph id="8BDD36FB015FFFE0C2B2CAB0FDA78935" blockId="3.[227,737,1439,1561]" pageId="3" pageNumber="178">
<emphasis id="B916EAE9015FFFE0C2B2CAB0FE678A9B" bold="true" box="[227,460,1439,1463]" pageId="3" pageNumber="178">Sexual dimorphism.</emphasis>
Characteristic for genus. Visible in shape of meso­ and metatibiae as larger sinuosity and flattening with presence of elongate projection at apex in males.
</paragraph>
</subSubSection>
<subSubSection id="C3786570015FFFE0C2B2C96CFCDE8B43" pageId="3" pageNumber="178" type="etymology">
<paragraph id="8BDD36FB015FFFE0C2B2C96CFEC98991" blockId="3.[227,737,1603,1725]" pageId="3" pageNumber="178">
<emphasis id="B916EAE9015FFFE0C2B2C96CFECC8977" bold="true" box="[227,359,1603,1627]" pageId="3" pageNumber="178">Etymology.</emphasis>
Patronymic. Name of newly described species is dedicated to its collector and our colleque who passed away some time ago: Otto Merkl.
</paragraph>
<paragraph id="8BDD36FB015FFFE0C157CFEEFCDE8B43" blockId="3.[774,1285,193,1135]" pageId="3" pageNumber="178">
<emphasis id="B916EAE9015FFFE0C157CFEEFCCE8FF5" bold="true" box="[774,869,193,217]" pageId="3" pageNumber="178">Affinity.</emphasis>
Because of compilation of features, i.e. relatively medium size of body; well­developed lateral lobes of pronotum, with anterior lobe distinctly larger than posterior; relatively short and wide elytra, each intercostae of elytra with two rows of medium sized punctures and external protrusion of caudal bulbs distinctly, sinuately divided from median this species can be confuzed only with
<taxonomicName id="4C624D78015FFFE0C121CEE7FB078EF3" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[880,1196,456,479]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="B916EAE9015FFFE0C121CEE7FC578EF3" box="[880,1020,456,479]" italics="true" pageId="3" pageNumber="178">R. denticollis</emphasis>
Fairmaire, 1893
</taxonomicName>
(see fig. 13) and
<taxonomicName id="4C624D78015FFFE0C130CEC6FC978D0C" authority="Anichtchenko et al., 2021" authorityName="Anichtchenko" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="mindanaoensis">
<emphasis id="B916EAE9015FFFE0C130CEC6FB898D2C" box="[865,1058,489,512]" italics="true" pageId="3" pageNumber="178">R. mindanaoensis</emphasis>
<bibRefCitation id="EFF34B0A015FFFE0C679CEC7FC978D0C" author="Anichtchenko A. &amp; Minkina Y. &amp; Vasiljeva A. &amp; Medina M. N. D." pageId="3" pageNumber="178" pagination="99 - 111" refId="ref4145" refString="Anichtchenko A., Minkina Y., Vasiljeva A., Medina M. N. D. 2021. A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae, 61 (1): 99 - 111." type="journal article" year="2021">
Anichtchenko
<emphasis id="B916EAE9015FFFE0C696CEC6FB548D2C" box="[1223,1279,489,512]" italics="true" pageId="3" pageNumber="178">et al.</emphasis>
, 2021
</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFF34B0A015FFFE0C127CD26FB3F8D0C" author="Anichtchenko A. &amp; Minkina Y. &amp; Vasiljeva A. &amp; Medina M. N. D." box="[886,1172,521,545]" pageId="3" pageNumber="178" pagination="99 - 111" refId="ref4145" refString="Anichtchenko A., Minkina Y., Vasiljeva A., Medina M. N. D. 2021. A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae, 61 (1): 99 - 111." type="journal article" year="2021">
Anichtchenko
<emphasis id="B916EAE9015FFFE0C644CD25FBE78D0D" box="[1045,1100,522,545]" italics="true" pageId="3" pageNumber="178">et al.</emphasis>
, 2021
</bibRefCitation>
: figs 5a­c, 11, 17, 23, 29a­b). From both species can be easily distinguished by punctation of intercostae of pronotum (
<taxonomicName id="4C624D78015FFFE0C109CD43FC4A8DAF" authorityName="Fairmaire" authorityYear="1893" box="[856,993,620,643]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="B916EAE9015FFFE0C109CD43FC4A8DAF" box="[856,993,620,643]" italics="true" pageId="3" pageNumber="178">R. denticollis</emphasis>
</taxonomicName>
have all intercostae densely punctate;
<taxonomicName id="4C624D78015FFFE0C120CDA2FB998D88" box="[881,1074,653,676]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="mindanaoensis">
<emphasis id="B916EAE9015FFFE0C120CDA2FB998D88" box="[881,1074,653,676]" italics="true" pageId="3" pageNumber="178">R. mindanaoensis</emphasis>
</taxonomicName>
have median intercosta similarly, but much more densely punctate and third intercostae with small area with dense punctation in basal part) and shape of caudal bulbs (
<taxonomicName id="4C624D78015FFFE0C141CC3FFC098C0B" authorityName="Fairmaire" authorityYear="1893" box="[784,930,784,807]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="B916EAE9015FFFE0C141CC3FFC098C0B" box="[784,930,784,807]" italics="true" pageId="3" pageNumber="178">R. denticollis</emphasis>
</taxonomicName>
as well
<taxonomicName id="4C624D78015FFFE0C654CC3FFB638C0B" box="[1029,1224,784,807]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="mindanaoensis">
<emphasis id="B916EAE9015FFFE0C654CC3FFB638C0B" box="[1029,1224,784,807]" italics="true" pageId="3" pageNumber="178">R. mindanaoensis</emphasis>
</taxonomicName>
have median and internal protrusions of caudal bulbs well distinguishable, with more or less distinct sinuation between them) (compare figs: 1, 13 and 5a fromAnichtchenko
<emphasis id="B916EAE9015FFFE0C1B6CCBCFBBC8C86" box="[999,1047,915,938]" italics="true" pageId="3" pageNumber="178">et al.</emphasis>
, 2021).What interesting aedeagus of
<taxonomicName id="4C624D78015FFFE0C1DDCC9BFC458CE7" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[908,1006,948,971]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="merkli" status="sp. nov.">
<emphasis id="B916EAE9015FFFE0C1DDCC9BFC458CE7" box="[908,1006,948,971]" italics="true" pageId="3" pageNumber="178">R. merkli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2255792015FFFE0C1A5CC9BFBEE8CE7" box="[1012,1093,948,971]" pageId="3" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
is more distinctly de­curved than aedeagus of
<taxonomicName id="4C624D78015FFFE0C67ECCFAFB168CC0" authorityName="Fairmaire" authorityYear="1893" box="[1071,1213,981,1004]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="B916EAE9015FFFE0C67ECCFAFB168CC0" box="[1071,1213,981,1004]" italics="true" pageId="3" pageNumber="178">R. denticollis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C624D78015FFFE0C6BFCCFAFC038B20" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="mindanaoensis">
<emphasis id="B916EAE9015FFFE0C6BFCCFAFC038B20" italics="true" pageId="3" pageNumber="178">R. mindanaoensis</emphasis>
</taxonomicName>
(compare figs: 9 and 29a from
<bibRefCitation id="EFF34B0A015FFFE0C157CB39FB888B01" author="Anichtchenko A. &amp; Minkina Y. &amp; Vasiljeva A. &amp; Medina M. N. D." box="[774,1059,1046,1069]" pageId="3" pageNumber="178" pagination="99 - 111" refId="ref4145" refString="Anichtchenko A., Minkina Y., Vasiljeva A., Medina M. N. D. 2021. A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae, 61 (1): 99 - 111." type="journal article" year="2021">
Anichtchenko
<emphasis id="B916EAE9015FFFE0C1F7CB39FC768B01" box="[934,989,1046,1069]" italics="true" pageId="3" pageNumber="178">et al.</emphasis>
, 2021
</bibRefCitation>
) what is relatively rarely observed distinguishal feature among genus
<taxonomicName id="4C624D78015FFFE0C157CB77FCC68B43" authorityName="Westwood" authorityYear="1845" box="[774,877,1112,1135]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B916EAE9015FFFE0C157CB77FCC68B43" box="[774,877,1112,1135]" italics="true" pageId="3" pageNumber="178">Rhyparus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,298 @@
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<mods:title id="8632FF8D696BB07A611A8DD1B7D4E56C">Studies on genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with description of two new species from Southeast Asia</mods:title>
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<mods:date id="CBAF6D1706362C6C26AEE819E276736F">2022</mods:date>
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<subSubSection id="C3786570015FFFE0C157CB95FBAF8BFD" box="[774,1028,1210,1234]" pageId="3" pageNumber="178" type="nomenclature">
<paragraph id="8BDD36FB015FFFE0C157CB95FBAF8BFD" blockId="3.[774,1035,1210,1266]" box="[774,1028,1210,1234]" pageId="3" pageNumber="178">
<heading id="D0958197015FFFE0C157CB95FBAF8BFD" box="[774,1028,1210,1234]" fontSize="10" level="2" pageId="3" pageNumber="178" reason="7">
<taxonomicName id="4C624D78015FFFE0C157CB95FC018BFE" ID-CoL="9WJ2Q" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[774,938,1210,1234]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="jakli" status="sp. nov.">
<emphasis id="B916EAE9015FFFE0C157CB95FC018BFE" bold="true" box="[774,938,1210,1234]" italics="true" pageId="3" pageNumber="178">Rhyparus jakli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2255792015FFFE0C1E0CB95FBAF8BFD" box="[945,1028,1210,1233]" pageId="3" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3786570015FFFE0C157CBF4FBA08BDE" box="[774,1035,1243,1266]" pageId="3" pageNumber="178" type="description">
<paragraph id="8BDD36FB015FFFE0C157CBF4FBA08BDE" blockId="3.[774,1035,1210,1266]" box="[774,1035,1243,1266]" pageId="3" pageNumber="178">
(
<figureCitation id="13592A7E015FFFE0C15FCBF4FCC48BDE" box="[782,879,1243,1266]" captionStart="Figs 46" captionStartId="4.[302,348,1132,1155]" captionTargetBox="[302,1360,189,1187]" captionTargetId="figure-346@4.[451,1213,189,1098]" captionTargetPageId="4" captionText="Figs 46. Rhyparus jakli sp. nov., (scale lines: 1.0 mm)." pageId="3" pageNumber="178">Figs 46</figureCitation>
,
<figureCitation id="13592A7E015FFFE0C12ACBF4FC218BDE" box="[891,906,1243,1266]" captionStart="Figs 7, 8" captionStartId="5.[774,820,724,747]" captionTargetBox="[776,1283,190,691]" captionTargetId="figure-490@5.[774,1284,189,691]" captionTargetPageId="5" captionText="Figs 7, 8. Heads: 7 Rhyparus merkli sp. nov., 8 R. jakli sp. nov. (scale lines: Fig. 7: 0.5mm, Fig. 8: 1.0mm)." pageId="3" pageNumber="178">8</figureCitation>
,
<figureCitation id="13592A7E015FFFE0C1C7CBF4FC778BDE" box="[918,988,1243,1266]" captionStart="Figs 912" captionStartId="5.[774,819,1595,1618]" captionTargetBox="[779,1282,888,1559]" captionTargetId="figure-488@5.[774,1284,882,1561]" captionTargetPageId="5" captionText="Figs 912.Aedeagi and epipharyngi:9 aedeagus of Rhyparus merkli sp. nov., 10 aedeagus of R. jakli sp. nov., 11 epipharynx of R. merkli sp. nov., 12 epipharynx of R. jakli sp. nov. (scale lines: Figs 9, 10: 0.5mm, Figs 11, 12: 1.0mm)." pageId="3" pageNumber="178">10, 12</figureCitation>
,
<figureCitation id="13592A7E015FFFE0C1B9CBF4FBAF8BDE" box="[1000,1028,1243,1266]" captionStart="Figs 1316" captionStartId="7.[227,273,1189,1212]" captionTargetBox="[333,1182,192,1154]" captionTargetId="figure-241@7.[330,1184,189,1157]" captionTargetPageId="7" captionText="Figs 1316. Dorsal views and abdominal apexes: 13 dorsal view of Rhyparus denticollis, 14 dorsal view of R. verrucosus (Bali Island), 15 abdominal apex of R. verrucosus (holotype, Sumatra), 16 abdominal apex of R. jakli sp. nov. (scale lines: 1.0 mm)." pageId="3" pageNumber="178">16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3786570015FFFE0C157CA33FB198AFB" pageId="3" pageNumber="178" type="materials_examined">
<paragraph id="8BDD36FB015FFFE0C157CA33FC158A78" blockId="3.[774,1284,1308,1364]" pageId="3" pageNumber="178">
<materialsCitation id="3B0A3CA6015FFFE0C157CA33FC158A78" collectorName="W Timor" country="Indonesia" location="Buraen." pageId="3" pageNumber="178" specimenCount="1" typeStatus="holotype">
<emphasis id="B916EAE9015FFFE0C157CA33FC0A8A18" bold="true" box="[774,929,1308,1332]" pageId="3" pageNumber="178">
<typeStatus id="54D98859015FFFE0C157CA33FC958A18" box="[774,830,1308,1332]" pageId="3" pageNumber="178">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F375766B015FFFE0C1F8CA32FBBE8A18" box="[937,1045,1309,1332]" name="Indonesia" pageId="3" pageNumber="178">Indonesia</collectingCountry>
,
<collectorName id="2697532D015FFFE0C670CA32FB298A18" box="[1057,1154,1309,1332]" pageId="3" pageNumber="178">W Timor</collectorName>
,
<locationDeviation id="9A6F2FD9015FFFE0C6C3CA32FCF58A78" location="Kupang" pageId="3" pageNumber="178">
<quantity id="4C9A9B1E015FFFE0C6C3CA32FB648A18" box="[1170,1231,1309,1332]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="178" unit="km" value="50.0">50km</quantity>
S of Kupang
</locationDeviation>
,
<location id="8EBD6020015FFFE0C139CA12FC158A78" LSID="urn:lsid:plazi:treatment:03CB87ED015FFFE5C157CB95FE028991:8EBD6020015FFFE0C139CA12FC158A78" box="[872,958,1341,1364]" country="Indonesia" name="Buraen." pageId="3" pageNumber="178">Buraen.</location>
</materialsCitation>
</paragraph>
<paragraph id="8BDD36FB015FFFE0C157CA51FB198AFB" blockId="3.[774,1288,1406,1496]" pageId="3" pageNumber="178">
<emphasis id="B916EAE9015FFFE0C157CA51FC1C8ABA" bold="true" box="[774,951,1406,1430]" pageId="3" pageNumber="178">Type material.</emphasis>
<materialsCitation id="3B0A3CA6015FFFE0C193CA50FB198AFB" collectingDate="2006-01-26" collectingDateMax="2006-02-09" collectingDateMin="2006-01-26" collectionCode="Y" collectorName="W Timor &amp; Stanislav Jakl" country="Indonesia" elevation="350" location="Buraen" pageId="3" pageNumber="178" specimenCount="350" typeStatus="holotype">
<typeStatus id="54D98859015FFFE0C193CA50FB828ABA" box="[962,1065,1407,1430]" pageId="3" pageNumber="178" type="holotype">Holotype</typeStatus>
():
<collectingCountry id="F375766B015FFFE0C63FCA50FB718ABA" box="[1134,1242,1407,1430]" name="Indonesia" pageId="3" pageNumber="178">Indonesia</collectingCountry>
,
<collectorName id="2697532D015FFFE0C6BDCA50FCEC8A9B" pageId="3" pageNumber="178">W Timor</collectorName>
,
<locationDeviation id="9A6F2FD9015FFFE0C102CA8FFBB08A9B" box="[851,1051,1440,1463]" location="Kupang" pageId="3" pageNumber="178">
<quantity id="4C9A9B1E015FFFE0C102CA8FFC3B8A9B" box="[851,912,1440,1463]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="178" unit="km" value="50.0">50km</quantity>
S of Kupang
</locationDeviation>
,
<location id="8EBD6020015FFFE0C675CA8FFBDF8A9B" LSID="urn:lsid:plazi:treatment:03CB87ED015FFFE5C157CB95FE028991:8EBD6020015FFFE0C675CA8FFBDF8A9B" box="[1060,1140,1440,1463]" country="Indonesia" name="Buraen" pageId="3" pageNumber="178">Buraen</location>
,
<specimenCount id="9D64FD72015FFFE0C62CCA8FFB6F8A9B" box="[1149,1220,1440,1463]" count="350" pageId="3" pageNumber="178" type="generic">
<quantity id="4C9A9B1E015FFFE0C62CCA8FFB6B8A9B" box="[1149,1216,1440,1463]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="178" unit="m" value="350.0">
<elevation id="004FD1C8015FFFE0C62CCA8FFB6B8A9B" box="[1149,1216,1440,1463]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="178" unit="m" value="350.0">350m</elevation>
</quantity>
.
</specimenCount>
,
<date id="FFDC103B015FFFE0C69DCA8FFCDD8AFB" pageId="3" pageNumber="178" value="2006-01-26" valueMax="2006-02-09" valueMin="2006-01-26">
<collectingDate id="EF98E9D3015FFFE0C69DCA8FFCDD8AFB" pageId="3" pageNumber="178" value="2006-01-26" valueMax="2006-02-09" valueMin="2006-01-26">26.i.­ 09.ii.2006</collectingDate>
</date>
, leg.
<collectorName id="2697532D015FFFE0C1E1CAEFFBEF8AFB" box="[944,1092,1472,1495]" pageId="3" pageNumber="178">Stanislav Jakl</collectorName>
[
<emphasis id="B916EAE9015FFFE0C605CA90FBCF8AF4" bold="true" box="[1108,1124,1471,1496]" pageId="3" pageNumber="178">
<collectionCode id="ED73AE3E015FFFE0C605CA90FBCF8AF4" box="[1108,1124,1471,1496]" pageId="3" pageNumber="178">Ÿ</collectionCode>
</emphasis>
MCN].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3786570015FFFE5C157C92DFD228EF3" lastPageId="6" lastPageNumber="181" pageId="3" pageNumber="178" type="description">
<paragraph id="8BDD36FB015FFFE0C157C92DFBCE8991" blockId="3.[774,1285,1538,1725]" pageId="3" pageNumber="178">
<emphasis id="B916EAE9015FFFE0C157C92DFC388936" bold="true" box="[774,915,1538,1562]" pageId="3" pageNumber="178">Description.</emphasis>
Dorsum (
<figureCitation id="13592A7E015FFFE0C655C92DFBE38935" box="[1028,1096,1538,1561]" captionStart="Figs 46" captionStartId="4.[302,348,1132,1155]" captionTargetBox="[302,1360,189,1187]" captionTargetId="figure-346@4.[451,1213,189,1098]" captionTargetPageId="4" captionText="Figs 46. Rhyparus jakli sp. nov., (scale lines: 1.0 mm)." pageId="3" pageNumber="178">Fig. 4</figureCitation>
). Body length of
<typeStatus id="54D98859015FFFE0C157C90CFCCF8916" box="[774,868,1571,1594]" pageId="3" pageNumber="178" type="holotype">holotype</typeStatus>
8.4mm. Large in size as member of the genus. Elongate, except of 1/3 preapical parts of elytra and strongly convex elements as costae weakly shiny, brownishblack; except punctures, tops of costae and legs glabrous.
</paragraph>
<paragraph id="8BDD36FB0158FFE7C321CFBEFAA48F88" blockId="4.[368,1295,145,164]" box="[368,1295,145,164]" pageId="4" pageNumber="179">
<emphasis id="B916EAE90158FFE7C321CFBEFAA48F88" box="[368,1295,145,164]" italics="true" pageId="4" pageNumber="179">
Studies on genus
<taxonomicName id="4C624D780158FFE7C05CCFBEFD5C8F88" authority="Westwood, 1845" authorityName="Westwood" authorityYear="1845" box="[525,759,145,164]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="genus">Rhyparus Westwood, 1845</taxonomicName>
(
<taxonomicName id="4C624D780158FFE7C154CFBEFCC18F88" authorityName="Linnaeus" authorityYear="1758" box="[773,874,145,164]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="179" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C624D780158FFE7C126CFBEFC5A8F88" authorityName="Latreille" authorityYear="1802" box="[887,1009,145,164]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
:
<taxonomicName id="4C624D780158FFE7C1ACCFBEFBC98F88" authorityName="Leach" authorityYear="1815" box="[1021,1122,145,164]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="subFamily" subFamily="Aphodiinae">Aphodiinae</taxonomicName>
) with description...
</emphasis>
</paragraph>
<caption id="DF1D66730158FFE7C37FCB43FDBD8B8F" pageId="4" pageNumber="179" startId="4.[302,348,1132,1155]" targetBox="[302,1360,189,1187]" targetPageId="4" targetType="figure">
<paragraph id="8BDD36FB0158FFE7C37FCB43FD368BAF" blockId="4.[302,669,1132,1187]" box="[302,669,1132,1155]" pageId="4" pageNumber="179">
Figs 46.
<taxonomicName id="4C624D780158FFE7C3CACB43FD918BAF" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[411,570,1132,1155]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="jakli" status="sp. nov.">
<emphasis id="B916EAE90158FFE7C3CACB43FD918BAF" box="[411,570,1132,1155]" italics="true" pageId="4" pageNumber="179">Rhyparus jakli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557920158FFE7C013CB43FD3E8BAF" box="[578,661,1132,1155]" pageId="4" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
,
</paragraph>
<paragraph id="8BDD36FB0158FFE7C37FCBA3FDBD8B8F" blockId="4.[302,669,1132,1187]" box="[302,534,1164,1187]" pageId="4" pageNumber="179">(scale lines: 1.0 mm).</paragraph>
</caption>
<paragraph id="8BDD36FB0158FFE7C37FCBCFFD1A897B" blockId="4.[302,813,1248,1623]" pageId="4" pageNumber="179">
Head (
<figureCitation id="13592A7E0158FFE7C32DCBCFFE6B8BDB" box="[380,448,1248,1271]" captionStart="Figs 7, 8" captionStartId="5.[774,820,724,747]" captionTargetBox="[776,1283,190,691]" captionTargetId="figure-490@5.[774,1284,189,691]" captionTargetPageId="5" captionText="Figs 7, 8. Heads: 7 Rhyparus merkli sp. nov., 8 R. jakli sp. nov. (scale lines: Fig. 7: 0.5mm, Fig. 8: 1.0mm)." pageId="4" pageNumber="179">Fig. 8</figureCitation>
) trapezoidal, clypeus anteriorly weakly sinuate, with relatively distinct teeth on sides, laterally distinctly emarginated and next with widely rounded angles, relatively not so relatively distinctly separated from distinctly protruding, rounded genae. Clypeocentral disc relatively large, distinctly convex, ringed by not so deep grove, with pair of indistinct, weakly convex, shiny, longitudinal tubercles. Clypeocentral disc densely punctate. Frons with four short, longitudinal costae. Punctation of head relatively regular, all punctures bearing short macrosetae.
</paragraph>
<paragraph id="8BDD36FB0158FFE7C37FC9AFFC498A15" blockId="4.[302,813,1664,1751]" lastBlockId="4.[850,1361,1249,1337]" pageId="4" pageNumber="179">
Epipharynx (
<figureCitation id="13592A7E0158FFE7C3E6C9AFFDA989BB" box="[439,514,1664,1687]" captionStart="Figs 912" captionStartId="5.[774,819,1595,1618]" captionTargetBox="[779,1282,888,1559]" captionTargetId="figure-488@5.[774,1284,882,1561]" captionTargetPageId="5" captionText="Figs 912.Aedeagi and epipharyngi:9 aedeagus of Rhyparus merkli sp. nov., 10 aedeagus of R. jakli sp. nov., 11 epipharynx of R. merkli sp. nov., 12 epipharynx of R. jakli sp. nov. (scale lines: Figs 9, 10: 0.5mm, Figs 11, 12: 1.0mm)." pageId="4" pageNumber="179">Fig. 12</figureCitation>
) distinctly transverse, rather distinctly sinuate at middle, corypha with several long celter, with two lateral lobes bearing tuft of smaller and finer celtes. Epitorma broad, somewhat rectangular, with dense chaetopedia laterally. Tormae long.
</paragraph>
<paragraph id="8BDD36FB0158FFE7C103CA4BFB0389EF" blockId="4.[850,1361,1380,1731]" pageId="4" pageNumber="179">Pronotum transverse, with six distinct, costae and seven longitudinal intercostae; on each side of costae there is row of short and small yellowish macrosetae. Lateral margin with two distinctly protruding lobes; anterior lobes sharply triangular with relatively rounded top, somewhat less developed than broadly triangular posterior which have somewhat rounded top too. Paramedian costae continuous, parallelogram. Discolateral costae interrupted in apical half. All intercosta with dense, medium sized punctures.</paragraph>
<paragraph id="8BDD36FB0159FFE6C025CFBEFCDE8F88" blockId="5.[628,885,145,164]" box="[628,885,145,164]" pageId="5" pageNumber="180">
<emphasis id="B916EAE90159FFE6C025CFBEFCDE8F88" box="[628,885,145,164]" italics="true" pageId="5" pageNumber="180">Minkina Ł., Anichtchenko A.</emphasis>
</paragraph>
<paragraph id="8BDD36FB0159FFE6C2B2CFEEFD948FF4" blockId="5.[227,575,193,216]" box="[227,575,193,216]" pageId="5" pageNumber="180">Scutellum almost imperceptible.</paragraph>
<paragraph id="8BDD36FB0159FFE6C2B2CE2EFEEA8C50" blockId="5.[227,738,257,892]" pageId="5" pageNumber="180">Elytra elongate, relatively narrow, widest before the middle. Each elytron with five distinct costae and five intercostae; on each side of costae there is row of short and small yellowish macrosetae. Fourth costae widened basally; covered by relatively dense, moderately large punctures. Intercostae flat, all of them with two rows of medium sized punctures; sometimes simple additional punctures are visible between them especially basaly on second intercosta; third intercosta with distinct row of relatively small punctures between two rows of medium sized punctures. Postdiscal bulbs relatively moderate in size. Caudal bulbs moderately strongly developed; external protrusion quite distinctly divided from median and internal which are fused into one and visible as transverse, elongate bulb; only very week sinuation between them. Pygidium with elevated area basally; medially with relatively deep grooves on sides.</paragraph>
<caption id="DF1D66730159FFE6C157CDFBFC058C07" pageId="5" pageNumber="180" startId="5.[774,820,724,747]" targetBox="[776,1283,190,691]" targetPageId="5" targetType="figure">
<paragraph id="8BDD36FB0159FFE6C157CDFBFC058C07" blockId="5.[774,1285,724,811]" pageId="5" pageNumber="180">
Figs 7, 8. Heads: 7
<taxonomicName id="4C624D780159FFE6C1BECDFBFB088DC7" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[1007,1187,724,747]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="merkli" status="sp. nov.">
<emphasis id="B916EAE90159FFE6C1BECDFBFB088DC7" box="[1007,1187,724,747]" italics="true" pageId="5" pageNumber="180">Rhyparus merkli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557920159FFE6C6FACDFBFB568DC7" box="[1195,1277,724,747]" pageId="5" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
, 8
<taxonomicName id="4C624D780159FFE6C17ECDDBFCD68C27" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[815,893,756,779]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="jakli" status="sp. nov.">
<emphasis id="B916EAE90159FFE6C17ECDDBFCD68C27" box="[815,893,756,779]" italics="true" pageId="5" pageNumber="180">R. jakli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557920159FFE6C1D5CDDBFC7C8C27" box="[900,983,756,779]" pageId="5" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
(scale lines: Fig. 7: 0.5mm, Fig. 8: 1.0mm).
</paragraph>
</caption>
<paragraph id="8BDD36FB0159FFE6C2B2CC89FE2B8C91" blockId="5.[227,384,934,957]" box="[227,384,934,957]" pageId="5" pageNumber="180">Macropterous.</paragraph>
<paragraph id="8BDD36FB0159FFE6C2B2CCC8FE258A2F" blockId="5.[227,737,999,1283]" pageId="5" pageNumber="180">
Venter weakly shiny, brownishblack (
<figureCitation id="13592A7E0159FFE6C0DFCCC8FD7B8CD2" box="[654,720,999,1022]" captionStart="Figs 46" captionStartId="4.[302,348,1132,1155]" captionTargetBox="[302,1360,189,1187]" captionTargetId="figure-346@4.[451,1213,189,1098]" captionTargetPageId="4" captionText="Figs 46. Rhyparus jakli sp. nov., (scale lines: 1.0 mm)." pageId="5" pageNumber="180">Fig. 5</figureCitation>
). Meso­metaventral plate concave, with distinct, relatively shallow, relatively thin, somewhat arrow­shaped median impression in basal 4/5; densely covered by medium sized punctures bearing very thin macrosetae; punctures here distinctly variable in size. Abdomal ventrites separated by not so large, but deep, quite regular in shape impressed sutures.
</paragraph>
<paragraph id="8BDD36FB0159FFE6C2B2CA02FDE689E6" blockId="5.[227,738,1325,1738]" pageId="5" pageNumber="180">Legs sturdy, shiny, with a lot of small punctured with eracted short setae. Mesofemora with distinct teeth on basal border, metafemora without teeth there. All femora with very dense, not so fine, quite regular punctation bearing very thin macrosetation. Protibia tridentate apically, straight, with rather distinct, relatively small basal dent. Mesotibiae narrow at base, widest in basal third of length, and next narrowed to apex; at apex with widened apical projection. Metatibiae with similar structure as mesotibiae, but with more distinctly developed apical projection. Metatibiae as general relatively straight.</paragraph>
<caption id="DF1D66730159FFE6C157C914FB5189FE" pageId="5" pageNumber="180" startId="5.[774,819,1595,1618]" targetBox="[779,1282,888,1559]" targetPageId="5" targetType="figure">
<paragraph id="8BDD36FB0159FFE6C157C914FB5189FE" blockId="5.[774,1285,1595,1746]" pageId="5" pageNumber="180">
Figs 912.Aedeagi and epipharyngi: 9 aedeagus of
<taxonomicName id="4C624D780159FFE6C173C974FC7F895E" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[802,980,1627,1650]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="merkli" status="sp. nov.">
<emphasis id="B916EAE90159FFE6C173C974FC7F895E" box="[802,980,1627,1650]" italics="true" pageId="5" pageNumber="180">Rhyparus merkli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557920159FFE6C188C974FB81895E" box="[985,1066,1627,1650]" pageId="5" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
, 10 aedeagus of
<taxonomicName id="4C624D780159FFE6C6BFC974FC9D89BE" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="jakli" status="sp. nov.">
<emphasis id="B916EAE90159FFE6C6BFC974FC9D89BE" italics="true" pageId="5" pageNumber="180">R. jakli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557920159FFE6C16EC954FC3989BE" box="[831,914,1659,1682]" pageId="5" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
, 11 epipharynx of
<taxonomicName id="4C624D780159FFE6C62BC954FB7589BE" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[1146,1246,1659,1682]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="merkli" status="sp. nov.">
<emphasis id="B916EAE90159FFE6C62BC954FB7589BE" box="[1146,1246,1659,1682]" italics="true" pageId="5" pageNumber="180">R. merkli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557920159FFE6C6B7C954FC98899E" pageId="5" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
, 12 epipharynx of
<taxonomicName id="4C624D780159FFE6C647C9B4FBCF899E" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[1046,1124,1691,1714]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="jakli" status="sp. nov.">
<emphasis id="B916EAE90159FFE6C647C9B4FBCF899E" box="[1046,1124,1691,1714]" italics="true" pageId="5" pageNumber="180">R. jakli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557920159FFE6C63DC9B4FB14899E" box="[1132,1215,1691,1714]" pageId="5" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
(scale lines: Figs 9, 10: 0.5mm, Figs 11, 12: 1.0mm).
</paragraph>
</caption>
<paragraph id="8BDD36FB015AFFE5C37FCFEEFE7C8E70" blockId="6.[302,813,193,348]" pageId="6" pageNumber="181">
Aedeagus (
<figureCitation id="13592A7E015AFFE5C3F5CFEEFE448FF4" box="[420,495,193,216]" captionStart="Figs 912" captionStartId="5.[774,819,1595,1618]" captionTargetBox="[779,1282,888,1559]" captionTargetId="figure-488@5.[774,1284,882,1561]" captionTargetPageId="5" captionText="Figs 912.Aedeagi and epipharyngi:9 aedeagus of Rhyparus merkli sp. nov., 10 aedeagus of R. jakli sp. nov., 11 epipharynx of R. merkli sp. nov., 12 epipharynx of R. jakli sp. nov. (scale lines: Figs 9, 10: 0.5mm, Figs 11, 12: 1.0mm)." pageId="6" pageNumber="181">Fig. 10</figureCitation>
) Phallobase much longer than parameres, moderately de­curved toward apex in lateral view. Parameres small, thin, semitransparent, visible as weak protrusion of each side of median lobe.
</paragraph>
<paragraph id="8BDD36FB015AFFE5C37FCEA9FD8D8EB1" blockId="6.[302,550,390,414]" box="[302,550,390,414]" pageId="6" pageNumber="181">
<emphasis id="B916EAE9015AFFE5C37FCEA9FE048EB2" bold="true" box="[302,431,390,414]" pageId="6" pageNumber="181">Variability.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BDD36FB015AFFE5C37FCEE8FD228EF3" blockId="6.[302,649,455,479]" box="[302,649,455,479]" pageId="6" pageNumber="181">
<emphasis id="B916EAE9015AFFE5C37FCEE8FDB88EF3" bold="true" box="[302,531,455,479]" pageId="6" pageNumber="181">Sexual dimorphism.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3786570015AFFE5C37FCD26FE028991" pageId="6" pageNumber="181" type="etymology">
<paragraph id="8BDD36FB015AFFE5C37FCD26FE628D6D" blockId="6.[302,813,521,577]" pageId="6" pageNumber="181">
<emphasis id="B916EAE9015AFFE5C37FCD26FE018D0D" bold="true" box="[302,426,521,545]" pageId="6" pageNumber="181">Etymology.</emphasis>
Patronymic. Dedicated to its collector: Stanislav Jakl.
</paragraph>
<paragraph id="8BDD36FB015AFFE5C37FCD44FE028991" blockId="6.[302,813,619,1725]" pageId="6" pageNumber="181">
<emphasis id="B916EAE9015AFFE5C37FCD44FE258DAF" bold="true" box="[302,398,619,643]" pageId="6" pageNumber="181">Affinity.</emphasis>
Because of combination of features: large size of body, dense punctation of all intercostae of pronotum and external protrusion of caudal bulbs quite distinctly divided from median protrusion
<taxonomicName id="4C624D78015AFFE5C3F2CDC0FE5B8C2A" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[419,496,751,774]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="jakli" status="sp. nov.">
<emphasis id="B916EAE9015AFFE5C3F2CDC0FE5B8C2A" box="[419,496,751,774]" italics="true" pageId="6" pageNumber="181">R. jakli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2255792015AFFE5C3A4CDC0FDEC8C2A" box="[501,583,751,774]" pageId="6" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
can be confused only with two Oceanian species:
<taxonomicName id="4C624D78015AFFE5C037CC3FFD538C0B" authorityName="Fairmaire" authorityYear="1893" box="[614,760,784,807]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="B916EAE9015AFFE5C037CC3FFD538C0B" box="[614,760,784,807]" italics="true" pageId="6" pageNumber="181">R. denticollis</emphasis>
</taxonomicName>
(see fig. 13) and
<taxonomicName id="4C624D78015AFFE5C3E1CC1EFD588C6B" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[432,755,816,840]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="B916EAE9015AFFE5C3E1CC1EFDCD8C64" box="[432,614,817,840]" italics="true" pageId="6" pageNumber="181">R. philippinensis</emphasis>
Arrow, 1905
</taxonomicName>
(see:
<bibRefCitation id="EFF34B0A015AFFE5C37FCC7EFDE38C44" author="Anichtchenko A. &amp; Minkina Y. &amp; Vasiljeva A. &amp; Medina M. N. D." box="[302,584,849,873]" pageId="6" pageNumber="181" pagination="99 - 111" refId="ref4145" refString="Anichtchenko A., Minkina Y., Vasiljeva A., Medina M. N. D. 2021. A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae, 61 (1): 99 - 111." type="journal article" year="2021">
Anichtchenko
<emphasis id="B916EAE9015AFFE5C39DCC7DFDAA8C45" box="[460,513,850,873]" italics="true" pageId="6" pageNumber="181">et al.</emphasis>
, 2021
</bibRefCitation>
: figs 6a­c, 12, 18, 24, 30a­b). However
<taxonomicName id="4C624D78015AFFE5C3B8CC5DFDDC8CA5" authorityName="Fairmaire" authorityYear="1893" box="[489,631,882,905]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="B916EAE9015AFFE5C3B8CC5DFDDC8CA5" box="[489,631,882,905]" italics="true" pageId="6" pageNumber="181">R. denticollis</emphasis>
</taxonomicName>
has: smaller size of body; lateral lobes of pronotum much more distinct, anterior lobe larger than posterior; elytra proportionally shorter and wider, with distinct sinuation between median and inner protrusion of caudal bulb (compare figs: 4 and 13).
<taxonomicName id="4C624D78015AFFE5C09ACB39FE2C8B62" authorityName="Arrow" authorityYear="1905" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="B916EAE9015AFFE5C09ACB39FE2C8B62" italics="true" pageId="6" pageNumber="181">R. philippinensis</emphasis>
</taxonomicName>
has: smaller size of body (moderate in size species); lateral lobes of pronotum widely rounded. Median and internal protrusions of pronotum differently directed less transverse (compare figs: 4 and 6a from
<bibRefCitation id="EFF34B0A015AFFE5C01BCB95FEC38BDE" author="Anichtchenko A. &amp; Minkina Y. &amp; Vasiljeva A. &amp; Medina M. N. D." pageId="6" pageNumber="181" pagination="99 - 111" refId="ref4145" refString="Anichtchenko A., Minkina Y., Vasiljeva A., Medina M. N. D. 2021. A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae). Acta Entomologica Musei Nationalis Pragae, 61 (1): 99 - 111." type="journal article" year="2021">
Anichtchenko
<emphasis id="B916EAE9015AFFE5C0BDCB95FC8D8BFD" box="[748,806,1210,1233]" italics="true" pageId="6" pageNumber="181">et al.</emphasis>
, 2021
</bibRefCitation>
). However when we base on distributional data, size of body, general proportions, shape of ventral side of body (similar meso­metaventral plate and abdominal ventrites) we can suspect that most closely related to
<taxonomicName id="4C624D78015AFFE5C002CA71FD0B8A59" authorityName="Minkina &amp; Anichtchenko" authorityYear="2022" box="[595,672,1374,1397]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="jakli" status="sp. nov.">
<emphasis id="B916EAE9015AFFE5C002CA71FD0B8A59" box="[595,672,1374,1397]" italics="true" pageId="6" pageNumber="181">R. jakli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2255792015AFFE5C0F7CA71FD538A59" box="[678,760,1374,1397]" pageId="6" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
is
<taxonomicName id="4C624D78015AFFE5C144CA71FD908ABA" authority="Schmidt, 1916" authorityName="Schmidt" authorityYear="1916" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="B916EAE9015AFFE5C144CA71FE0B8ABA" italics="true" pageId="6" pageNumber="181">R. verrucosus</emphasis>
Schmidt, 1916
</taxonomicName>
(see
<figureCitation id="13592A7E015AFFE5C03FCA50FD128ABA" box="[622,697,1407,1430]" captionStart="Figs 1316" captionStartId="7.[227,273,1189,1212]" captionTargetBox="[333,1182,192,1154]" captionTargetId="figure-241@7.[330,1184,189,1157]" captionTargetPageId="7" captionText="Figs 1316. Dorsal views and abdominal apexes: 13 dorsal view of Rhyparus denticollis, 14 dorsal view of R. verrucosus (Bali Island), 15 abdominal apex of R. verrucosus (holotype, Sumatra), 16 abdominal apex of R. jakli sp. nov. (scale lines: 1.0 mm)." pageId="6" pageNumber="181">Fig. 14</figureCitation>
). Both species are very easily distingushable by punctation of intercostae of pronotum (
<taxonomicName id="4C624D78015AFFE5C00FCAEEFD598AF4" authorityName="Schmidt" authorityYear="1916" box="[606,754,1473,1496]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="B916EAE9015AFFE5C00FCAEEFD598AF4" box="[606,754,1473,1496]" italics="true" pageId="6" pageNumber="181">R. verrucosus</emphasis>
</taxonomicName>
have densely punctate only median intercosta), shape of lateral lobes of pronotum (
<taxonomicName id="4C624D78015AFFE5C033C92DFD5E8935" authorityName="Schmidt" authorityYear="1916" box="[610,757,1538,1561]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="B916EAE9015AFFE5C033C92DFD5E8935" box="[610,757,1538,1561]" italics="true" pageId="6" pageNumber="181">R. verrucosus</emphasis>
</taxonomicName>
have somewhat rounded lobes), shape of caudal bulb (clearly less developed in
<taxonomicName id="4C624D78015AFFE5C019C96BFD778977" authorityName="Schmidt" authorityYear="1916" box="[584,732,1604,1627]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="B916EAE9015AFFE5C019C96BFD778977" box="[584,732,1604,1627]" italics="true" pageId="6" pageNumber="181">R. verrucosus</emphasis>
</taxonomicName>
) (compare figs: 4 and 14) and different shape of last abdominal ventrite and pygidium (compare figs: 15 and 16).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="BA5706833C824120CC76313A9F26D1F4">A new species of the genus Procleomenes Gressit &amp; Rondon, 1970 (Coleoptera: Cerambycidae) from the Guangxi, China</mods:title>
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<paragraph id="8BDAE40DFFA62B4FFFFFFD5827C3FD37" blockId="1.[113,456,707,763]" box="[113,456,707,731]" pageId="1" pageNumber="396">
<heading id="D0925361FFA62B4FFFFFFD5827C3FD37" bold="true" box="[113,456,707,731]" fontSize="10" level="2" pageId="1" pageNumber="396" reason="4">
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<taxonomicName id="4C659F8EFFA62B4FFFFFFD582760FD37" ID-CoL="BS966" authorityName="Barševskis" authorityYear="2022" box="[113,363,707,731]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="medinai" status="sp. nov.">
<emphasis id="B911381FFFA62B4FFFFFFD582760FD37" bold="true" box="[113,363,707,731]" italics="true" pageId="1" pageNumber="396">Procleomenes medinai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2228564FFA62B4FFEFCFD5827C3FD37" box="[370,456,707,731]" pageId="1" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BDAE40DFFA62B4FFFFFFD7F26C9FD17" blockId="1.[113,456,707,763]" box="[113,194,740,763]" pageId="1" pageNumber="396">
(
<figureCitation id="135EF888FFA62B4FFFF7FD7F26BCFD17" box="[121,183,740,763]" captionStart="Fig" captionStartId="1.[113,152,1589,1612]" captionTargetBox="[116,601,811,1558]" captionTargetId="figure-486@1.[113,604,809,1560]" captionTargetPageId="1" captionText="Fig. 1. Procleomenes medinai sp. nov. (holotypus)" pageId="1" pageNumber="396">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37FB786FFA62B4FFD0DFF002221FE80" pageId="1" pageNumber="396" type="materials_examined">
<paragraph id="8BDAE40DFFA62B4FFD0DFF00226CFED8" blockId="1.[643,1135,155,308]" pageId="1" pageNumber="396">
<emphasis id="B911381FFFA62B4FFD0DFF00252DFF5F" bold="true" box="[643,806,155,179]" pageId="1" pageNumber="396">Type material.</emphasis>
<materialsCitation id="3B0DEE50FFA62B4FFCA4FF0025B7FF1F" country="China" location="Dayao Shan Mt." pageId="1" pageNumber="396" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="54DE5AAFFFA62B4FFCA4FF0025C1FF5E" box="[810,970,155,178]" pageId="1" pageNumber="396" type="holotype">HOLOTYPUS</typeStatus>
, female:
<collectingCountry id="F372A49DFFA62B4FFBA6FF002260FF5E" box="[1064,1131,155,178]" name="China" pageId="1" pageNumber="396">China</collectingCountry>
:
<collectingRegion id="49A12AEFFFA62B4FFD0DFF2724E9FF3F" box="[643,738,188,211]" country="China" name="Guangxi" pageId="1" pageNumber="396">Guangxi</collectingRegion>
, /
<location id="8EBAB2D6FFA62B4FFD73FF2725BAFF3F" LSID="urn:lsid:plazi:treatment:03CC551BFFA62B4CFFFFFD5827FAFA39:8EBAB2D6FFA62B4FFD73FF2725BAFF3F" box="[765,945,188,211]" country="China" name="Dayao Shan Mt." pageId="1" pageNumber="396" stateProvince="Guangxi">Dayao Shan Mt.</location>
, / 05.2020./ local collector leg. [handwritten]; //
</materialsCitation>
<materialsCitation id="3B0DEE50FFA62B4FFC4FFF47226FFED8" collectionCode="DUBC" collectorName="A. Bar" country="China" pageId="1" pageNumber="396" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="54DE5AAFFFA62B4FFC4FFF472268FF1F" box="[961,1123,220,243]" pageId="1" pageNumber="396" type="holotype">HOLOTYPUS</typeStatus>
:/
<taxonomicName id="4C659F8EFFA62B4FFD0DFF66251DFEF8" authorityName="Gressit &amp; Rondon" authorityYear="1970" box="[643,790,253,276]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="397" phylum="Arthropoda" rank="genus">Procleomenes</taxonomicName>
/ medinai
<taxonomicNameLabel id="A2228564FFA62B4FFC0EFF6625C4FEF8" box="[896,975,253,276]" pageId="1" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
/
<collectorName id="269081DBFFA62B4FFC50FF662229FEF8" box="[990,1058,253,276]" pageId="1" pageNumber="396">A. Bar</collectorName>
<emphasis id="B911381FFFA62B4FFBACFF602227FEF8" bold="true" box="[1058,1068,251,276]" pageId="1" pageNumber="396">š</emphasis>
evskis descr. 2022 [red label, handwritten] (
<collectionCode id="ED747CC8FFA62B4FFB82FE862257FED8" box="[1036,1116,285,308]" pageId="1" pageNumber="396">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDAE40DFFA62B4FFD0DFECF2221FE80" blockId="1.[643,1066,340,364]" box="[643,1066,340,364]" pageId="1" pageNumber="396">
<emphasis id="B911381FFFA62B4FFD0DFECF257FFE80" bold="true" box="[643,884,340,364]" pageId="1" pageNumber="396">General distribution:</emphasis>
<collectingCountry id="F372A49DFFA62B4FFCF4FECE25B5FE80" box="[890,958,341,364]" name="China" pageId="1" pageNumber="396">China</collectingCountry>
:
<collectingRegion id="49A12AEFFFA62B4FFC48FECE222EFE80" box="[966,1061,341,364]" country="China" name="Guangxi" pageId="1" pageNumber="396">Guangxi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C37FB786FFA62B4CFD0DFE102735FD1A" lastPageId="2" lastPageNumber="397" pageId="1" pageNumber="396" type="description">
<paragraph id="8BDAE40DFFA62B4FFD0DFE1024C3FDE8" blockId="1.[643,1135,395,516]" pageId="1" pageNumber="396">
<emphasis id="B911381FFFA62B4FFD0DFE102501FE4F" bold="true" box="[643,778,395,419]" pageId="1" pageNumber="396">Description.</emphasis>
Body length:
<quantity id="4C9D49E8FFA62B4FFC17FE1725E1FE4F" box="[921,1002,396,419]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="1" pageNumber="396" unit="mm" value="6.6">6.6 mm</quantity>
, body width:
<quantity id="4C9D49E8FFA62B4FFD0DFE3724DDFE2F" box="[643,726,428,451]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="396" unit="mm" value="1.3">1.3 mm</quantity>
. Dorsal surface dark. Elytra flat, shiny, with three transverse somewhat transparent bands.
</paragraph>
<paragraph id="8BDAE40DFFA62B4FFD0DFDBE25BEFB88" blockId="1.[643,1135,549,1124]" pageId="1" pageNumber="396">Head small, quadrangular, transverse, flattened, with sparse pubescence and fine microsculpture. Eyes large, extended. Cheeks narrow, not extended,with sparse pubescence. Head between antennal bases with wide deep impression. Clypeus narrow, shiny, with transverse deep impression in basal part. Labrum brown, shiny, with yellow setae. Mandibles brown, wide, shiny, with very acute apices. Maxillary palpomeres brown. Antennae dark­brown. Basal antennomere elongate and thickened, with fine microsculpture and relatively short setation; antennomere 2 short, yellow, covered with fine setae; antennomeres 34 yellow in basal portions, dark­brown in apical portions, covered with pubescence; remaining antennomeres unicolor dark­brown. Inner sides of all antennomeres with row of some long setae.</paragraph>
<paragraph id="8BDAE40DFFA62B4FFD0DFB1E223FF9EE" blockId="1.[643,1135,1157,1538]" pageId="1" pageNumber="396">
Pronotum elongate, subcylindrical, with dorsaland lateral tubercles. Length:
<quantity id="4C9D49E8FFA62B4FFC44FB3E2215FB50" box="[970,1054,1189,1212]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="396" unit="mm" value="1.6">1.6 mm</quantity>
, width:
<quantity id="4C9D49E8FFA62B4FFD0DFB5D24DAFB31" box="[643,721,1222,1245]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="396" unit="mm" value="0.7">0.7 mm</quantity>
before tubercles and
<quantity id="4C9D49E8FFA62B4FFC26FB5D25FDFB31" box="[936,1014,1222,1245]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="396" unit="mm" value="1.1">1.1 mm</quantity>
with lateral tubercles; anterior and posterior borders prominent; pronotum distinctly longer thanwide. Pronotal disc with a pair of lateral protruded tubercles and five dorsal tubercles. Pronotum shiny, with very fine microsculptureand very fine, sparse, coarse punctation in middle; lateral portionsswith numerous long setae. Pars stridens partly visible under basal margin ofpronotum, emargined with impressed lines laterally.
</paragraph>
<paragraph id="8BDAE40DFFA62B4CFD0DF9B927F2FD53" blockId="1.[643,1135,1570,1690]" lastBlockId="2.[151,644,155,703]" lastPageId="2" lastPageNumber="397" pageId="1" pageNumber="396">
Dorsal surface of elytra flat, shiny, without pubescence, dark, with three transverse almost transparent bands. Elytra narrowing in about middle and slightly broadened posteriorly; apical margin of elytra not reaching apex of abdomen. Apices of elytra truncate, brown, with rudimental angles, not extended in sharp denticles. Anteromedian portions of each elytron with oval impression at base near suture, with very sparse and fine punctation; each puncture bearing a long yellow seta. Width of elytra at shoulders:
<quantity id="4C9D49E8FFA52B4CFF19FE1A26E6FE74" box="[151,237,385,408]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="397" unit="mm" value="1.2">1.2 mm</quantity>
. Length of elytra:
<quantity id="4C9D49E8FFA52B4CFE3AFE1A2402FE74" box="[436,521,385,408]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="397" unit="mm" value="3.1">3.1 mm</quantity>
. Surface of elytra with fine rows of relatively sparse punctures. Ventral surface of body smooth, with sparse punctation and, with irregular rudimental pubescence. Legs relatively long and thin, with thickened apical portions of femora. Widest part of femora black, otherportions yellow. Tibia dark­brown, flattened atexternal margin, with fringe of long setae. Tarsomeres yellow, covered by sparseand golden pubescence.
</paragraph>
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<paragraph id="8BDAE40DFFA62B4FFFFFF9AE26B8F981" blockId="1.[113,605,1589,1645]" pageId="1" pageNumber="396">
Fig. 1.
<taxonomicName id="4C659F8EFFA62B4FFF4FF9AD27BDF9A1" authorityName="Barševskis" authorityYear="2022" box="[193,438,1590,1613]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="medinai" status="sp. nov.">
<emphasis id="B911381FFFA62B4FFF4FF9AD27BDF9A1" box="[193,438,1590,1613]" italics="true" pageId="1" pageNumber="396">Procleomenes medinai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2228564FFA62B4FFE31F9AE2418F9A0" box="[447,531,1589,1612]" pageId="1" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
(holotypus)
</paragraph>
</caption>
<paragraph id="8BDAE40DFFA52B4CFF19FD442735FD1A" blockId="2.[151,318,735,758]" box="[151,318,735,758]" pageId="2" pageNumber="397">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="C37FB786FFA52B4CFF19FC8C270AFB34" pageId="2" pageNumber="397" type="diagnosis">
<paragraph id="8BDAE40DFFA52B4CFF19FC8C270AFB34" blockId="2.[151,643,791,1241]" pageId="2" pageNumber="397">
<emphasis id="B911381FFFA52B4CFF19FC8C2784FCC3" bold="true" box="[151,399,791,815]" pageId="2" pageNumber="397">Differential diagnosis.</emphasis>
<taxonomicName id="4C659F8EFFA52B4CFE1DFC8C2488FCC2" authorityName="Barševskis" authorityYear="2022" box="[403,643,791,814]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="medinai" status="sp. nov.">
<emphasis id="B911381FFFA52B4CFE1DFC8C2488FCC2" box="[403,643,791,814]" italics="true" pageId="2" pageNumber="397">Procleomenes medinai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2228564FFA52B4CFF19FCA326E3FCA3" box="[151,232,824,847]" pageId="2" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
with a relatively dark surface is similar to
<taxonomicName id="4C659F8EFFA52B4CFF3EFCC227B6FC9C" authority="Holzschuh, 2007" authorityName="Holzschuh" authorityYear="2007" box="[176,445,857,880]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B911381FFFA52B4CFF3EFCC2270CFC9C" box="[176,263,857,880]" italics="true" pageId="2" pageNumber="397">P. morio</emphasis>
<bibRefCitation id="EFF499FCFFA52B4CFE82FCC227B6FC9C" author="Holzschuh C." box="[268,445,857,880]" pageId="2" pageNumber="397" pagination="177 - 286" refId="ref1397" refString="Holzschuh C. 2007. Beschreibung von 80 neuen Bockkafern aus der orientalischen und palaearktischen Region, vorwiegend aus China, Laos und Borneo (Coleoptera, Cerambycidae). Entomologica Basiliensa et Collectionis Frey 29: 177 - 286." type="journal article" year="2007">Holzschuh, 2007</bibRefCitation>
</taxonomicName>
, known from
<collectingCountry id="F372A49DFFA52B4CFDC7FCC22475FC9C" box="[585,638,857,880]" name="Laos" pageId="2" pageNumber="397">Laos</collectingCountry>
, but differs from it by the different surface pattern. The first postbasal transverse transparent band of a new species is broad and triangular, from the base of the elytra at the scutellum almost extending the lateral margin, enclosing the dark shoulder (this band of
<taxonomicName id="4C659F8EFFA52B4CFE08FB8527EBFBD9" authorityName="Holzschuh" authorityYear="2007" box="[390,480,1054,1077]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B911381FFFA52B4CFE08FB8527EBFBD9" box="[390,480,1054,1077]" italics="true" pageId="2" pageNumber="397">P. morio</emphasis>
</taxonomicName>
is smaller, narrower, not reaching the base and lateral margin, does not surround the shoulder). Remaining second and third dorsal tranverse bands of a new species are very narrow (relatively broader in
<taxonomicName id="4C659F8EFFA52B4CFF19FB5926FAFB35" authorityName="Holzschuh" authorityYear="2007" box="[151,241,1218,1241]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="B911381FFFA52B4CFF19FB5926FAFB35" box="[151,241,1218,1241]" italics="true" pageId="2" pageNumber="397">P. morio</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37FB786FFA52B4CFF19FB6227FAFA39" pageId="2" pageNumber="397" type="etymology">
<paragraph id="8BDAE40DFFA52B4CFF19FB6227FAFA39" blockId="2.[151,643,1273,1493]" pageId="2" pageNumber="397">
<emphasis id="B911381FFFA52B4CFF19FB622711FAFD" bold="true" box="[151,282,1273,1297]" pageId="2" pageNumber="397">Etymology.</emphasis>
This species is named after my friend and excellent colleague, Philippine entomologist Dr. Milton Norman Medina (
<collectingRegion id="49A12AEFFFA52B4CFDB5FAA0272DFA9E" country="Philippines" name="Davao Oriental" pageId="2" pageNumber="397">Davao Oriental State</collectingRegion>
University,
<collectingCountry id="F372A49DFFA52B4CFE2DFAC02410FA9E" box="[419,539,1371,1394]" name="Philippines" pageId="2" pageNumber="397">Philippines</collectingCountry>
) in appreciation of friendship, excellent cooperation, and in gratitude for his contribution to the studies of
<taxonomicName id="4C659F8EFFA52B4CFF3AFA252722FA39" authorityName="Linnaeus" authorityYear="1758" box="[180,297,1470,1493]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="2" pageNumber="397" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
in the
<collectingCountry id="F372A49DFFA52B4CFEFCFA2527E5FA39" box="[370,494,1470,1493]" name="Philippines" pageId="2" pageNumber="397">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="F31E2B1522C1BEE80B8AE0DFB1CD7954">A new record of Oxynopterus harmseni Candèze, 1885 (Coleoptera: Elateridae) from Sumatra</mods:title>
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<mods:namePart id="EB610A7840A78AE38A7023E803269724">Vanaga, Anarina</mods:namePart>
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<mods:namePart id="C7D955089862D7AA125D987A1688A6D7">Barševskis, ArvÓds</mods:namePart>
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<mods:title id="B816FD73106B5B371605319B6A9CFEE8">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="BEF9B8C1C53D5BCB050270DBDA4063DD">2022</mods:date>
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<paragraph id="8BC736ADFFF6DF6E4790B94EFBD5FED7" blockId="1.[643,1057,322,378]" box="[643,1057,322,346]" pageId="1" pageNumber="392">
<heading id="D08F81C1FFF6DF6E4790B94EFBD5FED7" bold="true" box="[643,1057,322,346]" fontSize="10" level="2" pageId="1" pageNumber="392" reason="4">
<taxonomicName id="4C784D2EFFF6DF6E4790B94EFBD5FED7" authority="Candeze, 1885" authorityName="Candeze" authorityYear="1885" box="[643,1057,322,346]" class="Insecta" family="Elateridae" genus="Oxynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="harmsen">
<emphasis id="B90CEABFFFF6DF6E4790B94EFBD5FED7" bold="true" box="[643,1057,322,346]" pageId="1" pageNumber="392">
<emphasis id="B90CEABFFFF6DF6E4790B94EFC82FED7" bold="true" box="[643,886,322,346]" italics="true" pageId="1" pageNumber="392">Oxynopterus harmsen</emphasis>
<bibRefCitation id="EFE94B5CFFF6DF6E466EB94EFBD5FED7" author="Candeze E. C." box="[893,1057,322,346]" pageId="1" pageNumber="392" pagination="120 - 121" refId="ref1309" refString="Candeze E. C. 1885. Deux especes nouvelles d'Elaterides. Notes from the Leyden Museum, 7 (2): 120 - 121." type="journal article" year="1885">Candèze, 1885</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C3626526FFF6DF6E4790B96FFD03FEF7" box="[643,759,355,378]" pageId="1" pageNumber="392" type="description">
<paragraph id="8BC736ADFFF6DF6E4790B96FFD03FEF7" blockId="1.[643,1057,322,378]" box="[643,759,355,378]" pageId="1" pageNumber="392">
(
<figureCitation id="13432A28FFF6DF6E4798B96FFD1BFEF7" box="[651,751,355,378]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[151,190,1669,1692]" captionStartId-1="3.[113,152,782,805]" captionTargetBox-0="[164,1149,488,1635]" captionTargetBox-1="[115,864,153,745]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Oxynopterus harmseni (A dorsal view, B lateral view, C ventral view)." captionText-1="Fig. 2. Male genitalia of Oxynopterus harmseni (A aedeagus, B abdominal tergite IXX, C abdominal tergite XI)." pageId="1" pageNumber="392">Figs. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3626526FFF6DF6E4790B996FBC1FE7F" pageId="1" pageNumber="392" type="reference_group">
<paragraph id="8BC736ADFFF6DF6E4790B996FBC1FE7F" blockId="1.[643,1135,410,498]" pageId="1" pageNumber="392">
<emphasis id="B90CEABFFFF6DF6E4790B996FCFFFE3F" bold="true" box="[643,779,410,434]" pageId="1" pageNumber="392">References:</emphasis>
<bibRefCitation id="EFE94B5CFFF6DF6E4605B996FC41FE3C" author="Candeze E. C." box="[790,949,410,433]" pageId="1" pageNumber="392" pagination="120 - 121" refId="ref1309" refString="Candeze E. C. 1885. Deux especes nouvelles d'Elaterides. Notes from the Leyden Museum, 7 (2): 120 - 121." type="journal article" year="1885">Candèze 1885</bibRefCitation>
,
<bibRefCitation id="EFE94B5CFFF6DF6E46D4B996FC09FE3C" author="Candeze E. C." box="[967,1021,410,433]" pageId="1" pageNumber="392" refId="ref1336" refString="Candeze E. C. 1891. Catalogue metodique des Elaterides connus en 1890. xii + 246 pp., Liege." type="book" year="1891">1891</bibRefCitation>
;
<bibRefCitation id="EFE94B5CFFF6DF6E4103B996FD48FE5F" author="Schwarz O." pageId="1" pageNumber="392" pagination="1 - 224" refId="ref1641" refString="Schwarz O. 1906. Coleoptera, Fam. Elateridae. In: Wytsman P. (ed.), Genera Insectorum, 46 AB: 1 - 224." type="book chapter" year="1906">Schwarz 1906</bibRefCitation>
;
<bibRefCitation id="EFE94B5CFFF6DF6E47DAB9B7FC76FE5F" author="Schenkling S." box="[713,898,443,466]" pageId="1" pageNumber="392" pagination="1 - 263" refId="ref1529" refString="Schenkling S. 1925. Elateridae I. In: Junk W., Schenkling S. (eds.), Coleopterorum Catalogus, 80: 1 - 263." type="journal article" year="1925">Schenkling 1925</bibRefCitation>
;
<bibRefCitation id="EFE94B5CFFF6DF6E4682B9B7FBCCFE5F" author="Fleutiaux E." box="[913,1080,443,466]" pageId="1" pageNumber="392" pagination="95 - 112" refId="ref1359" refString="Fleutiaux E. 1926. Remarques et observations sur le Catalogue des Elateridae. 1 re partie de M. S. Schenkling. Annales de la Societe Entomologique de France, 95: 95 - 112." type="journal article" year="1926">Fleutiaux 1926</bibRefCitation>
;
<bibRefCitation id="EFE94B5CFFF6DF6E4156B9B7FCA9FE7F" author="Van Zwaluvenburg R. H." pageId="1" pageNumber="392" pagination="393 - 432" refId="ref1672" refString="Van Zwaluvenburg R. H. 1936. The elaterid beetles of the Philippine Islands. The Philippine Journal of Science, 39: 393 - 432." type="journal article" year="1936">Van Zwaluvenburg 1936</bibRefCitation>
;
<bibRefCitation id="EFE94B5CFFF6DF6E4674B9D7FC1BFE7F" author="Suzuki W." box="[871,1007,475,498]" pageId="1" pageNumber="392" pagination="326 - 334" refId="ref1560" refString="Suzuki W. 2001. Notes on three species of the genus Oxynopterus (Coleoptera, Elateridae) from Southeast Asia, with description of a new species. Elytra, 29 (2): 326 - 334." type="journal article" year="2001">Suzuki 2001</bibRefCitation>
,
<bibRefCitation id="EFE94B5CFFF6DF6E46EAB9D7FBC5FE7F" author="Suzuki W." box="[1017,1073,475,498]" pageId="1" pageNumber="392" pagination="217 - 222" refId="ref1600" refString="Suzuki W. 2011. A new subgenus of the genus Oxynopterus Hope (Coleoptera: Elateridae) from South East Asia. Spec. Publ. Jpn. Soc. Scarabaeoidology, Tokyo, 1: 217 - 222." type="journal article" year="2011">2011</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3626526FFF6DF6E4790BA1EFCFFFD6E" pageId="1" pageNumber="392" type="materials_examined">
<paragraph id="8BC736ADFFF6DF6E4790BA1EFD33FD06" blockId="1.[643,1134,530,651]" pageId="1" pageNumber="392">
<emphasis id="B90CEABFFFF6DF6E4790BA1EFC97FDA7" bold="true" box="[643,867,530,554]" pageId="1" pageNumber="392">Material examined:</emphasis>
<materialsCitation id="3B103CF0FFF6DF6E467ABA1EFB91FDA4" box="[873,1125,530,554]" country="Indonesia" location="Sumatra" pageId="1" pageNumber="392" specimenCount="1" stateProvince="Sumatera Utara">
<emphasis id="B90CEABFFFF6DF6E467ABA1EFC15FDA7" bold="true" box="[873,993,530,554]" pageId="1" pageNumber="392">
<collectingCountry id="F36F763DFFF6DF6E467ABA1EFC28FDA7" box="[873,988,530,554]" name="Indonesia" pageId="1" pageNumber="392">Indonesia</collectingCountry>
:
</emphasis>
<collectingRegion id="49BCF84FFFF6DF6E46F4BA1EFBB4FDA4" box="[999,1088,530,553]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="392">Sumatra</collectingRegion>
Isl
</materialsCitation>
.,
<materialsCitation id="3B103CF0FFF6DF6E4790BA3FFD37FD06" country="Indonesia" location="Bengkulu" pageId="1" pageNumber="392" specimenCount="1" specimenCount-male="1" stateProvince="Bengkulu">
<collectingRegion id="49BCF84FFFF6DF6E4790BA3FFD1EFDC7" box="[643,746,563,586]" country="Indonesia" name="Bengkulu" pageId="1" pageNumber="392">Bengkulu</collectingRegion>
, 05.2019; (
<specimenCount id="9D7EFD24FFF6DF6E4649BA3FFC6BFDC7" box="[858,927,563,586]" count="1" pageId="1" pageNumber="392" type="male">1 male</specimenCount>
, local collector leg.). The habitus as in
<figureCitation id="13432A28FFF6DF6E4629BA58FC83FDE6" box="[826,887,596,619]" captionStart="Fig" captionStartId="2.[151,190,1669,1692]" captionTargetBox="[164,1149,488,1635]" captionTargetPageId="2" captionText="Fig. 1. Oxynopterus harmseni (A dorsal view, B lateral view, C ventral view)." pageId="1" pageNumber="392">Fig. 1</figureCitation>
, and the aedeagus as in
<figureCitation id="13432A28FFF6DF6E4790BA78FD37FD06" box="[643,707,628,651]" captionStart="Fig" captionStartId="3.[113,152,782,805]" captionTargetBox="[115,864,153,745]" captionTargetPageId="3" captionText="Fig. 2. Male genitalia of Oxynopterus harmseni (A aedeagus, B abdominal tergite IXX, C abdominal tergite XI)." pageId="1" pageNumber="392">Fig. 2</figureCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736ADFFF6DF6E4790BAA7FCFFFD6E" blockId="1.[643,1134,683,739]" pageId="1" pageNumber="392">
<emphasis id="B90CEABFFFF6DF6E4790BAA7FC87FD4E" bold="true" box="[643,883,683,707]" pageId="1" pageNumber="392">General distribution:</emphasis>
<collectingCountry id="F36F763DFFF6DF6E4664BAA7FC16FD4F" box="[887,994,683,706]" name="Indonesia" pageId="1" pageNumber="392">Indonesia</collectingCountry>
:
<collectingRegion id="49BCF84FFFF6DF6E46F9BAA7FBB6FD4F" box="[1002,1090,683,706]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="392">Sumatra</collectingRegion>
and Nias islands.
</paragraph>
</subSubSection>
<subSubSection id="C3626526FFF6DF6C4790BB0FFC7CFD65" lastPageId="3" lastPageNumber="394" pageId="1" pageNumber="392" type="discussion">
<paragraph id="8BC736ADFFF6DF6E4790BB0FFBEDFBB2" blockId="1.[643,1134,771,1087]" pageId="1" pageNumber="392">
<emphasis id="B90CEABFFFF6DF6E4790BB0FFD06FC96" bold="true" box="[643,754,771,795]" pageId="1" pageNumber="392">Remarks.</emphasis>
<taxonomicName id="4C784D2EFFF6DF6E47EFBB0FFC00FC97" authorityName="Candeze" authorityYear="1885" box="[764,1012,771,794]" class="Insecta" family="Elateridae" genus="Oxynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="harmseni">
<emphasis id="B90CEABFFFF6DF6E47EFBB0FFC00FC97" box="[764,1012,771,794]" italics="true" pageId="1" pageNumber="392">Oxynopterus harmseni</emphasis>
</taxonomicName>
was originally described in 1885 from
<collectingCountry id="F36F763DFFF6DF6E46B6BB28FBFFFCB6" box="[933,1035,804,827]" name="Indonesia" pageId="1" pageNumber="392">Indonesia</collectingCountry>
,
<collectingRegion id="49BCF84FFFF6DF6E4101BB28FB9EFCB6" box="[1042,1130,804,827]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="392">Sumatra</collectingRegion>
, pay des Battaks (
<bibRefCitation id="EFE94B5CFFF6DF6E4629BB48FC2CFCD6" author="Candeze E. C." box="[826,984,836,859]" pageId="1" pageNumber="392" pagination="120 - 121" refId="ref1309" refString="Candeze E. C. 1885. Deux especes nouvelles d'Elaterides. Notes from the Leyden Museum, 7 (2): 120 - 121." type="journal article" year="1885">Candèze 1885</bibRefCitation>
). After that, it was mentioned only twice in the scientific literature (except catalogues or check­lists). It was firstly recorded 116 years after the original description (
<bibRefCitation id="EFE94B5CFFF6DF6E47E4BBCAFC77FC50" author="Suzuki W." box="[759,899,966,989]" pageId="1" pageNumber="392" pagination="326 - 334" refId="ref1560" refString="Suzuki W. 2001. Notes on three species of the genus Oxynopterus (Coleoptera, Elateridae) from Southeast Asia, with description of a new species. Elytra, 29 (2): 326 - 334." type="journal article" year="2001">Suzuki 2001</bibRefCitation>
,
<bibRefCitation id="EFE94B5CFFF6DF6E4686BBCAFC3EFC50" author="Suzuki W." box="[917,970,966,989]" pageId="1" pageNumber="392" pagination="217 - 222" refId="ref1600" refString="Suzuki W. 2011. A new subgenus of the genus Oxynopterus Hope (Coleoptera: Elateridae) from South East Asia. Spec. Publ. Jpn. Soc. Scarabaeoidology, Tokyo, 1: 217 - 222." type="journal article" year="2011">2011</bibRefCitation>
). The
<tableCitation id="C6FA0316FFF6DF6E4109BBCAFB9AFC50" box="[1050,1134,966,989]" captionStart="Table 1" captionStartId="2.[151,210,155,178]" captionText="Table 1. Published data on the representation of specimens of O. harmseni in the collections" pageId="1" pageNumber="392">Table 1</tableCitation>
shows the published data on the representation of the species in the collections, the number of specimens, sex and published records.
</paragraph>
<paragraph id="8BC736ADFFF6DF6E4790BC53FCF0FA36" blockId="1.[643,1134,1119,1467]" pageId="1" pageNumber="392">
<materialsCitation id="3B103CF0FFF6DF6E4790BC53FCF4FA36" county="It" location="Faunistic" municipality="Perhaps" pageId="1" pageNumber="392" specimenCount="9" specimenCount-female="1" specimenCount-male="4" stateProvince="Sumatera Utara">
Regarding the occurrence of
<taxonomicName id="4C784D2EFFF6DF6E46D4BC53FBBBFBFB" authorityName="Candeze" authorityYear="1885" box="[967,1103,1119,1142]" class="Insecta" family="Elateridae" genus="Oxynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="harmseni">
<emphasis id="B90CEABFFFF6DF6E46D4BC53FBBBFBFB" box="[967,1103,1119,1142]" italics="true" pageId="1" pageNumber="392">O. harmseni</emphasis>
</taxonomicName>
in world collections, currently only
<specimenCount id="9D7EFD24FFF6DF6E46C1BC73FB9AFB1B" box="[978,1134,1151,1174]" count="4" pageId="1" pageNumber="392" type="generic">four specimens</specimenCount>
of this species (
<specimenCount id="9D7EFD24FFF6DF6E4621BCACFC40FB3A" box="[818,948,1184,1207]" count="3" pageId="1" pageNumber="392" type="male">three males</specimenCount>
,
<specimenCount id="9D7EFD24FFF6DF6E46D2BCACFBC8FB3A" box="[961,1084,1184,1207]" count="1" pageId="1" pageNumber="392" type="female">one female</specimenCount>
) are available in the scientific literature, of which data on
<specimenCount id="9D7EFD24FFF6DF6E47CDBCEDFCB6FB75" box="[734,834,1249,1272]" count="1" pageId="1" pageNumber="392" type="male">one male</specimenCount>
is presented in the present study.
<collectingCounty id="62A64E21FFF6DF6E47DBBD0EFD2CFA94" box="[712,728,1282,1305]" pageId="1" pageNumber="392">It</collectingCounty>
is probably a very rare species whose biology and ecology are still unknown.
<collectingMunicipality id="6BA3ACD7FFF6DF6E4109BD2EFB9AFAB4" box="[1050,1134,1314,1337]" pageId="1" pageNumber="392">Perhaps</collectingMunicipality>
this species is in danger and needs real protection.
<location id="8EA76076FFF6DF6E47ABBD6FFCEDFAF7" LSID="urn:lsid:plazi:treatment:03D187BBFFF6DF6C4790B94EFC7CFD65:8EA76076FFF6DF6E47ABBD6FFCEDFAF7" box="[696,793,1379,1402]" county="It" municipality="Perhaps" name="Faunistic" pageId="1" pageNumber="392" stateProvince="Sumatera Utara">Faunistic</location>
studies in
<collectingRegion id="49BCF84FFFF6DF6E4695BD6FFC2BFAF7" box="[902,991,1379,1402]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="392">Sumatra</collectingRegion>
are needed to clarify the distribution and population status of this species
</materialsCitation>
.
</paragraph>
<caption id="DF076625FFF5DF6D4584B897FB9FFF3F" box="[151,1131,155,178]" pageId="2" pageNumber="393" startId="2.[151,210,155,178]" targetType="table">
<paragraph id="8BC736ADFFF5DF6D4584B897FB9FFF3F" blockId="2.[151,1131,155,178]" box="[151,1131,155,178]" pageId="2" pageNumber="393">
Table 1. Published data on the representation of specimens of
<taxonomicName id="4C784D2EFFF5DF6D4639B897FC5AFF3F" authorityName="Candeze" authorityYear="1885" box="[810,942,155,178]" class="Insecta" family="Elateridae" genus="Oxynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="harmseni">
<emphasis id="B90CEABFFFF5DF6D4639B897FC5AFF3F" box="[810,942,155,178]" italics="true" pageId="2" pageNumber="393">O. harmseni</emphasis>
</taxonomicName>
in the collections
</paragraph>
</caption>
<paragraph id="8BC736ADFFF5DF6D443AB8EEFB8EFE22" pageId="2" pageNumber="393">
<table id="F978C40DFFF52090458CB8D8FB7FFE33" box="[159,1163,212,446]" gridcols="4" gridrows="5" pageId="2" pageNumber="393">
<tr id="354834EFFFF52090458CB8D8FB7FFE88" box="[159,1163,212,261]" gridrow="0" pageId="2" pageNumber="393">
<th id="76995D93FFF52090458CB8D8FDE3FE88" box="[159,535,212,261]" gridcol="0" gridrow="0" pageId="2" pageNumber="393">Collections</th>
<th id="76995D93FFF520904720B8D8FD2EFE88" box="[563,730,212,261]" gridcol="1" gridrow="0" pageId="2" pageNumber="393">Number of speci­ mens, gender</th>
<th id="76995D93FFF5209047E7B8D8FC47FE88" box="[756,947,212,261]" gridcol="2" gridrow="0" pageId="2" pageNumber="393">Locality</th>
<th id="76995D93FFF5209046DCB8D8FB7FFE88" box="[975,1163,212,261]" gridcol="3" gridrow="0" pageId="2" pageNumber="393">References</th>
</tr>
<tr id="354834EFFFF52090458CB902FB7FFEAE" box="[159,1163,270,291]" gridrow="1" pageId="2" pageNumber="393">
<th id="76995D93FFF52090458CB902FDE3FEAE" box="[159,535,270,291]" gridcol="0" gridrow="1" pageId="2" pageNumber="393">Coll. H.Arimoto (Japan)</th>
<td id="76995D93FFF520904720B902FD2EFEAE" box="[563,730,270,291]" gridcol="1" gridrow="1" pageId="2" pageNumber="393">1, female</td>
<td id="76995D93FFF5209047E7B902FC47FEAE" box="[756,947,270,291]" gridcol="2" gridrow="1" pageId="2" pageNumber="393">Solok, W.Sumatra</td>
<td id="76995D93FFF5209046DCB902FB7FFEAE" box="[975,1163,270,291]" gridcol="3" gridrow="1" pageId="2" pageNumber="393">
<bibRefCitation id="EFE94B5CFFF5DF6D46DCB902FBBAFEAE" author="Suzuki W." box="[975,1102,270,291]" pageId="2" pageNumber="393" pagination="217 - 222" refId="ref1600" refString="Suzuki W. 2011. A new subgenus of the genus Oxynopterus Hope (Coleoptera: Elateridae) from South East Asia. Spec. Publ. Jpn. Soc. Scarabaeoidology, Tokyo, 1: 217 - 222." type="journal article" year="2011">Suzuki, 2011</bibRefCitation>
</td>
</tr>
<tr id="354834EFFFF52090458CB923FB7FFEC9" box="[159,1163,303,324]" gridrow="2" pageId="2" pageNumber="393">
<th id="76995D93FFF52090458CB923FDE3FEC9" box="[159,535,303,324]" gridcol="0" gridrow="2" pageId="2" pageNumber="393">Coll. W.Suzuki (Japan)</th>
<td id="76995D93FFF520904720B923FD2EFEC9" box="[563,730,303,324]" gridcol="1" gridrow="2" pageId="2" pageNumber="393">1, male</td>
<td id="76995D93FFF5209047E7B923FC47FEC9" box="[756,947,303,324]" gridcol="2" gridrow="2" pageId="2" pageNumber="393">Padang, W Sumatra</td>
<td id="76995D93FFF5209046DCB923FB7FFEC9" box="[975,1163,303,324]" gridcol="3" gridrow="2" pageId="2" pageNumber="393">
<bibRefCitation id="EFE94B5CFFF5DF6D46DCB923FBA5FEC9" author="Suzuki W." box="[975,1105,303,324]" pageId="2" pageNumber="393" pagination="326 - 334" refId="ref1560" refString="Suzuki W. 2001. Notes on three species of the genus Oxynopterus (Coleoptera, Elateridae) from Southeast Asia, with description of a new species. Elytra, 29 (2): 326 - 334." type="journal article" year="2001">Suzuki, 2001</bibRefCitation>
,
<bibRefCitation id="EFE94B5CFFF5DF6D414FB923FB7FFEC9" author="Suzuki W." box="[1116,1163,303,324]" pageId="2" pageNumber="393" pagination="217 - 222" refId="ref1600" refString="Suzuki W. 2011. A new subgenus of the genus Oxynopterus Hope (Coleoptera: Elateridae) from South East Asia. Spec. Publ. Jpn. Soc. Scarabaeoidology, Tokyo, 1: 217 - 222." type="journal article" year="2011">2011</bibRefCitation>
</td>
</tr>
<tr id="354834EFFFF52090458CB95CFB7FFE0C" box="[159,1163,336,385]" gridrow="3" pageId="2" pageNumber="393">
<th id="76995D93FFF52090458CB95CFDE3FE0C" box="[159,535,336,385]" gridcol="0" gridrow="3" pageId="2" pageNumber="393">Coll. Natural History Museum, London, UK</th>
<td id="76995D93FFF520904720B95CFD2EFE0C" box="[563,730,336,385]" gridcol="1" gridrow="3" pageId="2" pageNumber="393">1, male</td>
<td id="76995D93FFF5209047E7B95CFC47FE0C" box="[756,947,336,385]" gridcol="2" gridrow="3" pageId="2" pageNumber="393">Niasisl.</td>
<td id="76995D93FFF5209046DCB95CFB7FFE0C" box="[975,1163,336,385]" gridcol="3" gridrow="3" pageId="2" pageNumber="393">
<bibRefCitation id="EFE94B5CFFF5DF6D46DCB951FBBBFEFF" author="Suzuki W." box="[975,1103,349,370]" pageId="2" pageNumber="393" pagination="326 - 334" refId="ref1560" refString="Suzuki W. 2001. Notes on three species of the genus Oxynopterus (Coleoptera, Elateridae) from Southeast Asia, with description of a new species. Elytra, 29 (2): 326 - 334." type="journal article" year="2001">Suzuki, 2001</bibRefCitation>
</td>
</tr>
<tr id="354834EFFFF52090458CB981FB7FFE33" box="[159,1163,397,446]" gridrow="4" pageId="2" pageNumber="393">
<th id="76995D93FFF52090458CB981FDE3FE33" box="[159,535,397,446]" gridcol="0" gridrow="4" pageId="2" pageNumber="393">Beetles Coll. of Daugavpils University, Ilgas, Latvia</th>
<td id="76995D93FFF520904720B981FD2EFE33" box="[563,730,397,446]" gridcol="1" gridrow="4" pageId="2" pageNumber="393">1, male</td>
<td id="76995D93FFF5209047E7B981FC47FE33" box="[756,947,397,446]" gridcol="2" gridrow="4" pageId="2" pageNumber="393">Bengkulu, Sumatra</td>
<td id="76995D93FFF5209046DCB981FB7FFE33" box="[975,1163,397,446]" gridcol="3" gridrow="4" pageId="2" pageNumber="393">The present study</td>
</tr>
</table>
</paragraph>
<caption id="DF076625FFF5DF6D4584BE89FBFFF911" box="[151,1035,1669,1692]" pageId="2" pageNumber="393" startId="2.[151,190,1669,1692]" targetBox="[164,1149,488,1635]" targetPageId="2" targetType="figure">
<paragraph id="8BC736ADFFF5DF6D4584BE89FBFFF911" blockId="2.[151,1035,1669,1692]" box="[151,1035,1669,1692]" pageId="2" pageNumber="393">
Fig. 1.
<taxonomicName id="4C784D2EFFF5DF6D45F1BE89FE20F911" authorityName="Candeze" authorityYear="1885" box="[226,468,1669,1692]" class="Insecta" family="Elateridae" genus="Oxynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="harmseni">
<emphasis id="B90CEABFFFF5DF6D45F1BE89FE20F911" box="[226,468,1669,1692]" italics="true" pageId="2" pageNumber="393">Oxynopterus harmseni</emphasis>
</taxonomicName>
(A dorsal view, B lateral view, C ventral view).
</paragraph>
</caption>
<paragraph id="8BC736ADFFF4DF6C4666BADCFC7CFD65" blockId="3.[885,904,720,744]" box="[885,904,720,744]" pageId="3" pageNumber="394">C</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,366 @@
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<mods:title id="75D2F0652DB51838802A1A85DB5B8AD6">Two new species of MetapocyrtusHeller, 1912 (Coleoptera, Curculionidae, Entiminae) from Lanao and Panay, Philippines</mods:title>
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<paragraph id="8BC736FF4F223B35C126106BFABF414E" blockId="4.[850,1361,518,606]" box="[850,1361,518,542]" pageId="4" pageNumber="189">
<heading id="D08F81934F223B35C126106BFABF414E" bold="true" box="[850,1361,518,542]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="189" reason="4">
<emphasis id="B90CEAED4F223B35C126106BFABF414E" bold="true" box="[850,1361,518,542]" pageId="4" pageNumber="189">
<taxonomicName id="4C784D7C4F223B35C126106BFACE414E" ID-CoL="B7QKN" authority="Cabras" authorityName="Cabras &amp; Villegas &amp; Dagta &amp; Medina &amp; Barševskis" authorityYear="2022" box="[850,1312,518,542]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="ilgas" subGenus="Metapocyrtus">
<emphasis id="B90CEAED4F223B35C126106BFC09414E" bold="true" box="[850,999,518,542]" italics="true" pageId="4" pageNumber="189">Metapocyrtus</emphasis>
(
<emphasis id="B90CEAED4F223B35C180106BFB66414E" bold="true" box="[1012,1160,518,542]" italics="true" pageId="4" pageNumber="189">Metapocyrtus</emphasis>
)
<emphasis id="B90CEAED4F223B35C6E2106BFB24414E" bold="true" box="[1174,1226,518,542]" italics="true" pageId="4" pageNumber="189">ilgas</emphasis>
Cabras
</taxonomicName>
and
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC736FF4F223B35C126104AFBE0416F" blockId="4.[850,1361,518,606]" box="[850,1038,551,575]" pageId="4" pageNumber="189">
<heading id="D08F81934F223B35C126104AFBE0416F" bold="true" box="[850,1038,551,575]" fontSize="9" level="2" pageId="4" pageNumber="189" reason="6">
<emphasis id="B90CEAED4F223B35C126104AFBE0416F" bold="true" box="[850,1038,551,575]" pageId="4" pageNumber="189">
Medina,
<taxonomicNameLabel id="A23F57964F223B35C1CC104AFBE0416F" box="[952,1038,551,575]" pageId="4" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36265744F223B35C126102AFC35410E" box="[850,987,583,606]" pageId="4" pageNumber="189" type="description">
<paragraph id="8BC736FF4F223B35C126102AFC35410E" blockId="4.[850,1361,518,606]" box="[850,987,583,606]" pageId="4" pageNumber="189">
(
<figureCitation id="13432A7A4F223B35C12E102AFC3A410E" box="[858,980,583,606]" captionStart="Figure 2" captionStartId="5.[226,296,1693,1716]" captionTargetBox="[256,1257,192,1654]" captionTargetId="figure-56@5.[254,1260,189,1659]" captionTargetPageId="5" captionText="Figure 2. Metapocyrtus (Metapocyrtus) ilgas sp. nov. A, C, Holotype male; A. dorsal view, C. lateral view. B, D, Paratype female; B. dorsal view, D. lateral view" pageId="4" pageNumber="189">Fig. 2 AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36265744F223B35C12610E5FC374776" pageId="4" pageNumber="189" type="materials_examined">
<paragraph id="8BC736FF4F223B35C12610E5FBF940D3" blockId="4.[850,1361,648,1062]" pageId="4" pageNumber="189">
<materialsCitation id="3B103CA24F223B35C12610E5FC3041B1" location="December" municipality="Panay" pageId="4" pageNumber="189" specimenCount="1" stateProvince="Antique" typeStatus="holotype">
<emphasis id="B90CEAED4F223B35C12610E5FC5841F0" bold="true" box="[850,950,648,672]" pageId="4" pageNumber="189">
<typeStatus id="54C3885D4F223B35C12610E5FC5841F0" box="[850,950,648,672]" pageId="4" pageNumber="189" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13432A7A4F223B35C1B510E4FBC041F0" box="[961,1070,649,672]" captionStart="Figure 2" captionStartId="5.[226,296,1693,1716]" captionTargetBox="[256,1257,192,1654]" captionTargetId="figure-56@5.[254,1260,189,1659]" captionTargetPageId="5" captionText="Figure 2. Metapocyrtus (Metapocyrtus) ilgas sp. nov. A, C, Holotype male; A. dorsal view, C. lateral view. B, D, Paratype female; B. dorsal view, D. lateral view" pageId="4" pageNumber="189">Figs 2A,C</figureCitation>
), male: Philippines­ /
<collectingMunicipality id="6BA3AC854F223B35C77010E4FAAD41F0" box="[1284,1347,649,672]" pageId="4" pageNumber="189">Panay</collectingMunicipality>
/
<collectingRegion id="49BCF81D4F223B35C12610C4FC474190" box="[850,937,681,704]" country="Philippines" name="Antique" pageId="4" pageNumber="189">Antique</collectingRegion>
/
<location id="8EA760244F223B35C1CB10C4FBC34190" LSID="urn:lsid:plazi:treatment:03D187E94F223B39C126106BFABE43F4:8EA760244F223B35C1CB10C4FBC34190" box="[959,1069,681,704]" municipality="Panay" name="December" pageId="4" pageNumber="189" stateProvince="Antique">December</location>
, 2020 / coll. LC (typed on white card) //
</materialsCitation>
<materialsCitation id="3B103CA24F223B35C19010A7FBFD40D3" collectionCode="UMCRC, PNMNH" country="Philippines" county="National Museum" location="Presently" municipality="National Museum of Natural History" pageId="4" pageNumber="189" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C3885D4F223B35C19010A7FB9C41B1" box="[996,1138,714,737]" pageId="4" pageNumber="189" type="holotype">HOLOTYPE</typeStatus>
male /
<taxonomicName id="4C784D7C4F223B35C6C810A7FBD84052" authorityName="Cabras &amp; Villegas &amp; Dagta &amp; Medina &amp; Barševskis" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="ilgas" subGenus="Metapocyrtus">
<emphasis id="B90CEAED4F223B35C6C810A7FABE41B1" box="[1212,1360,714,737]" italics="true" pageId="4" pageNumber="189">Metapocyrtus</emphasis>
(
<emphasis id="B90CEAED4F223B35C12F1086FC014052" box="[859,1007,747,770]" italics="true" pageId="4" pageNumber="189">Metapocyrtus</emphasis>
)
<emphasis id="B90CEAED4F223B35C6751086FBD84052" box="[1025,1078,747,770]" italics="true" pageId="4" pageNumber="189">ilgas</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFE94B0E4F223B35C63A1087FC664072" author="Cabras A. A. &amp; Medina M. N. &amp; Bollino M." pageId="4" pageNumber="189" pagination="139 - 154" refId="ref4204" refString="Cabras A. A., Medina M. N., Bollino M. 2021 a. Two new species of the genus Metapocyrtus Heller, 1912 (Coleoptera, curculionidae, Entiminae, Pachyrhynchini), subgenus Orthocyrtus Heller, 1912, from Mindanao island, Philippines. ZooKeys, 1029: 139 - 154. https: // doi. org / 10.3897 / zookeys. 1029.63023" type="journal article" year="2021">CABRAS &amp; MEDINA, 2021</bibRefCitation>
(typed on red card).
<location id="8EA760244F223B35C6101166FB264072" LSID="urn:lsid:plazi:treatment:03D187E94F223B39C126106BFABE43F4:8EA760244F223B35C6101166FB264072" box="[1124,1224,779,802]" country="Philippines" county="National Museum" municipality="National Museum of Natural History" name="Presently" pageId="4" pageNumber="189">Presently</location>
in
<collectionCode id="ED69AE3A4F223B35C69D1166FAA34072" box="[1257,1357,779,802]" pageId="4" pageNumber="189">UMCRC</collectionCode>
, will be deposited in
<collectingMunicipality id="6BA3AC854F223B35C6571146FC4D4033" pageId="4" pageNumber="189">National Museum of Natural History</collectingMunicipality>
(
<collectionCode id="ED69AE3A4F223B35C1C51121FBF84033" box="[945,1046,844,867]" pageId="4" pageNumber="189">PNMNH</collectionCode>
) under the
<collectingCounty id="62A64E734F223B35C6FB1121FABE4033" box="[1167,1360,844,867]" pageId="4" pageNumber="189">National Museum</collectingCounty>
of the
<collectingCountry id="F36F766F4F223B35C1E21101FBFD40D3" box="[918,1043,876,899]" name="Philippines" pageId="4" pageNumber="189">Philippines</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736FF4F223B35C12611E1FC374776" blockId="4.[850,1361,648,1062]" pageId="4" pageNumber="189">
<materialsCitation id="3B103CA24F223B35C12611E1FAA34094" collectionCode="DUBC" county="Philippines-Panay" location="Presently" municipality="January" pageId="4" pageNumber="189" specimenCount="1" stateProvince="Antique" typeStatus="paratype">
<emphasis id="B90CEAED4F223B35C12611E1FC2B40F4" bold="true" box="[850,965,908,932]" pageId="4" pageNumber="189">
<typeStatus id="54C3885D4F223B35C12611E1FC2B40F4" box="[850,965,908,932]" pageId="4" pageNumber="189" type="paratype">Paratypes</typeStatus>
</emphasis>
(13, 6):
<collectingCounty id="62A64E734F223B35C63F11E0FAFE40F4" box="[1099,1296,909,932]" pageId="4" pageNumber="189">Philippines­Panay</collectingCounty>
/
<collectingRegion id="49BCF81D4F223B35C75211E0FC6B4094" country="Philippines" name="Antique" pageId="4" pageNumber="189">Antique</collectingRegion>
/
<collectingMunicipality id="6BA3AC854F223B35C1E011C0FC0F4094" box="[916,993,941,964]" pageId="4" pageNumber="189">January</collectingMunicipality>
, 2018 / coll. LC.
<location id="8EA760244F223B35C6FC11C0FB084094" LSID="urn:lsid:plazi:treatment:03D187E94F223B39C126106BFABE43F4:8EA760244F223B35C6FC11C0FB084094" box="[1160,1254,941,964]" county="Philippines-Panay" municipality="January" name="Presently" pageId="4" pageNumber="189" stateProvince="Antique">Presently</location>
in
<collectionCode id="ED69AE3A4F223B35C77611C0FAA34094" box="[1282,1357,941,964]" pageId="4" pageNumber="189">DUBC</collectionCode>
</materialsCitation>
.
<typeStatus id="54C3885D4F223B35C12611A3FC5440B5" box="[850,954,974,997]" pageId="4" pageNumber="189" type="paratype">Paratypes</typeStatus>
has additional red label:
<typeStatus id="54C3885D4F223B35C6CA11A3FAAC40B5" box="[1214,1346,974,997]" pageId="4" pageNumber="189" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C784D7C4F223B35C1261182FB664755" authorityName="Heller" authorityYear="1912" box="[850,1160,1006,1030]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="subGenus" subGenus="Metapocyrtus">
<emphasis id="B90CEAED4F223B35C1261182FC0C4756" box="[850,994,1007,1030]" italics="true" pageId="4" pageNumber="189">Metapocyrtus</emphasis>
(
<emphasis id="B90CEAED4F223B35C1841182FB6E4756" box="[1008,1152,1007,1030]" italics="true" pageId="4" pageNumber="189">Metapocyrtus</emphasis>
)
</taxonomicName>
/
<bibRefCitation id="EFE94B0E4F223B35C6ED1183FC374776" author="Cabras A. &amp; Medina M. N. &amp; Bollino M." pageId="4" pageNumber="189" pagination="115 - 128" refId="ref4348" refString="Cabras A., Medina M. N., Bollino M. 2022. Three new species of the genus Metapocyrtus Heller, 1912, subgenus Orthocyrtus Heller 1912 (Coleoptera, Curculionidae, Entiminae, Pachyrhynchini), from Mindanao Island, Philippines. ZooKeys, 1088: 115 - 128. doi: 10.3897 / zookeys. 1088.79021 https: // zookeys. pensoft. net" type="journal article" year="2022">CABRAS &amp; MEDINA, 2022</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36265744F223B35C126163DFB784799" pageId="4" pageNumber="189" type="diagnosis">
<paragraph id="8BC736FF4F223B35C126163DFB784799" blockId="4.[850,1361,1104,1225]" pageId="4" pageNumber="189">
<emphasis id="B90CEAED4F223B35C126163DFC264738" bold="true" box="[850,968,1104,1128]" pageId="4" pageNumber="189">Diagnosis.</emphasis>
<taxonomicName id="4C784D7C4F223B35C1A4163DFABF4737" authorityName="Cabras &amp; Villegas &amp; Dagta &amp; Medina &amp; Barševskis" authorityYear="2022" box="[976,1361,1104,1127]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="ilgas" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B90CEAED4F223B35C1A4163DFB8B4737" box="[976,1125,1104,1127]" italics="true" pageId="4" pageNumber="189">Metapocyrtus</emphasis>
(
<emphasis id="B90CEAED4F223B35C602163DFAE44737" box="[1142,1290,1104,1127]" italics="true" pageId="4" pageNumber="189">Metapocyrtus</emphasis>
)
<emphasis id="B90CEAED4F223B35C768163DFABF4737" box="[1308,1361,1104,1127]" italics="true" pageId="4" pageNumber="189">ilgas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57964F223B35C126161CFC4D47D8" box="[850,931,1137,1160]" pageId="4" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from any other
<taxonomicName id="4C784D7C4F223B35C681161CFC0D47F8" pageId="4" pageNumber="189">
<emphasis id="B90CEAED4F223B35C681161CFC7C47F8" italics="true" pageId="4" pageNumber="189">Metapocyrtus</emphasis>
species
</taxonomicName>
in Panay Island based on its unique pronotal and elytral markings.
</paragraph>
</subSubSection>
<subSubSection id="C36265744F223B37C126169FFABF46FC" lastPageId="6" lastPageNumber="191" pageId="4" pageNumber="189" type="description">
<paragraph id="8BC736FF4F223B35C126169FFB16461B" blockId="4.[850,1361,1266,1355]" pageId="4" pageNumber="189">
<emphasis id="B90CEAED4F223B35C126169FFBC2465A" bold="true" box="[850,1068,1266,1290]" pageId="4" pageNumber="189">Description. Male.</emphasis>
Dimensions: LB:
<quantity id="4C809B1A4F223B35C68A169EFAA4465A" box="[1278,1354,1267,1290]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="4" pageNumber="189" unit="mm" value="9.3">9.3mm</quantity>
. LR:
<quantity id="4C809B1A4F223B35C1FD177EFC39467A" box="[905,983,1299,1322]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="189" unit="mm" value="1.8">1.8mm</quantity>
. WR:
<quantity id="4C809B1A4F223B35C65D177EFB99467A" box="[1065,1143,1299,1322]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="189" unit="mm" value="1.8">1.8mm</quantity>
. LP:
<quantity id="4C809B1A4F223B35C6CB177EFAE2467A" box="[1215,1292,1299,1322]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="189" unit="mm" value="3.5">3.5mm</quantity>
. WP:
<quantity id="4C809B1A4F223B35C1261759FC4F461B" box="[850,929,1332,1355]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="189" unit="mm" value="3.5">3.5mm</quantity>
. LE:
<quantity id="4C809B1A4F223B35C1AD1759FBC6461B" box="[985,1064,1332,1355]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="189" unit="mm" value="5.8">5.8mm</quantity>
. WE:
<quantity id="4C809B1A4F223B35C61D1759FB59461B" box="[1129,1207,1332,1355]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="189" unit="mm" value="4.3">4.3mm</quantity>
. N=1.
</paragraph>
<paragraph id="8BC736FF4F223B35C1261718FB4A46FC" blockId="4.[850,1360,1397,1452]" pageId="4" pageNumber="189">Integument black. Body surface, rostrum, head, and underside with weak luster.</paragraph>
<paragraph id="8BC736FF4F223B35C12617BBFABE4581" blockId="4.[850,1361,1494,1745]" pageId="4" pageNumber="189">
<emphasis id="B90CEAED4F223B35C12617BBFC6046BE" bold="true" box="[850,910,1494,1518]" pageId="4" pageNumber="189">Head</emphasis>
finely punctured on dorsum with few sparse pubescence, frons with sparse pink, round scales, lateroventral parts below the eye with sparse pink elliptical scales with very few piliform scales, forehead between eyes nearly flattish, midline groove distinct extending from the vertex towards the transverse groove. Eyes small­sized and feebly convex, barely protruding from the outline of
</paragraph>
<paragraph id="8BC736FF4F233B34C3B212FCFBCC43F4" blockId="5.[454,1058,143,164]" box="[454,1058,143,164]" pageId="5" pageNumber="190">
<emphasis id="B90CEAED4F233B34C3B212FCFBCC43F4" box="[454,1058,143,164]" italics="true" pageId="5" pageNumber="190">Cabras A.A., Villegas J., Dagta M.J., Medina M.N.D., Barṥevskis A.</emphasis>
</paragraph>
<caption id="DF0766774F233B34C29614F0FC404584" pageId="5" pageNumber="190" startId="5.[226,296,1693,1716]" targetBox="[256,1257,192,1654]" targetPageId="5" targetType="figure">
<paragraph id="8BC736FF4F233B34C29614F0FC404584" blockId="5.[226,1284,1693,1749]" pageId="5" pageNumber="190">
Figure 2.
<taxonomicName id="4C784D7C4F233B34C33914F3FD2045E5" authorityName="Cabras &amp; Villegas &amp; Dagta &amp; Medina &amp; Barševskis" authorityYear="2022" box="[333,718,1693,1717]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="ilgas" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B90CEAED4F233B34C33914F3FE0F45E5" box="[333,481,1694,1717]" italics="true" pageId="5" pageNumber="190">Metapocyrtus</emphasis>
(
<emphasis id="B90CEAED4F233B34C38714F3FD6945E5" box="[499,647,1694,1717]" italics="true" pageId="5" pageNumber="190">Metapocyrtus</emphasis>
)
<emphasis id="B90CEAED4F233B34C0ED14F3FD2045E5" box="[665,718,1694,1717]" italics="true" pageId="5" pageNumber="190">ilgas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57964F233B34C0A114F0FCC445E4" box="[725,810,1693,1716]" pageId="5" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B90CEAED4F233B34C13C14F0FC6A45E5" bold="true" box="[840,900,1693,1717]" pageId="5" pageNumber="190">A, C,</emphasis>
Holotype male;
<emphasis id="B90CEAED4F233B34C64914F0FBB845E5" bold="true" box="[1085,1110,1693,1717]" pageId="5" pageNumber="190">A.</emphasis>
dorsal view,
<emphasis id="B90CEAED4F233B34C69F14F0FAEA45E5" bold="true" box="[1259,1284,1693,1717]" pageId="5" pageNumber="190">C.</emphasis>
lateral view.
<emphasis id="B90CEAED4F233B34C31914D0FE484585" bold="true" box="[365,422,1725,1749]" pageId="5" pageNumber="190">B, D,</emphasis>
Paratype female;
<emphasis id="B90CEAED4F233B34C01C14D0FD6E4585" bold="true" box="[616,640,1725,1749]" pageId="5" pageNumber="190">B.</emphasis>
dorsal view,
<emphasis id="B90CEAED4F233B34C17B14D0FCC74585" bold="true" box="[783,809,1725,1749]" pageId="5" pageNumber="190">D.</emphasis>
lateral view
</paragraph>
</caption>
<paragraph id="8BC736FF4F203B37C35A12ACFB7440B5" blockId="6.[302,813,193,1745]" lastBlockId="6.[850,1361,193,997]" pageId="6" pageNumber="191">
head.
<emphasis id="B90CEAED4F203B37C31912ACFE3C4389" bold="true" box="[365,466,193,217]" pageId="6" pageNumber="191">Rostrum</emphasis>
weakly rugose with coarse punctures on basal 2/3
<superScript id="7C0D9BB74F203B37C071128DFDFC43BE" attach="left" box="[517,530,224,238]" fontSize="6" pageId="6" pageNumber="191">rd</superScript>
and finely punctured on apical third, as long as wide (LR/WR:
<quantity id="4C809B1A4F203B37C0AD136FFCCA4249" box="[729,804,258,281]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="191" unit="mm" value="1.8">1.8mm</quantity>
/
<quantity id="4C809B1A4F203B37C35A134EFE95426A" box="[302,379,291,314]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="191" unit="mm" value="1.8">1.8mm</quantity>
), dorsum with sparse and adpressed brownish setae, lateral surface before antennal scrobe with sparse peach elliptical scales interspersed with minute adpressed piliform scales and below antennal scrobe are long white adpressed, piliform scales, anterolateral margin with whitish, long suberect setae, ventral surface with long suberect whitish setae; transverse basal groove deep and highly distinct; longitudinal groove along midline distinct extending up to basal half; dorsum with two oblique subparallel ridges which gradually declines towards dorsolateral margins; lateral sides with weakly expanded apicad. Antennal scape slightly longer than funicle, scape reaching a little beyond the hind margin of eye, covered with subadpressed fine light­colored hairs, and funicle with suberect yellowish hairs. Funicular segments I and II almost of the same length, three times longer than wide; segments IIIVII nearly as long as wide; club sub­ellipsoidal, nearly 3 times longer than wide.
<emphasis id="B90CEAED4F203B37C04211C0FD424095" bold="true" box="[566,684,941,965]" pageId="6" pageNumber="191">Prothorax</emphasis>
globose, as long as wide (LP/WP: 3.5/
<quantity id="4C809B1A4F203B37C04611A3FD9540B5" box="[562,635,974,997]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="191" unit="mm" value="3.5">3.5mm</quantity>
), finely punctured especially towards the anterior margin, widest at middle, moderately convex on dorsal surface, dorsal contour highest point at the middle. Prothorax with the following scaly markings of pink round scales: a) stripe at the anterior margin widened laterally, b) two subtriangular scaly markings on each side of disc starting at posterior margin tapered towards the middle, and c) broad patch before the coxa confluent with the anterior stripe.
<emphasis id="B90CEAED4F203B37C30D177EFE2C467B" bold="true" box="[377,450,1299,1323]" pageId="6" pageNumber="191">Elytra</emphasis>
subovate, longer than wide (LE/ WE: 5.8/
<quantity id="4C809B1A4F203B37C3FB1759FE30461B" box="[399,478,1332,1355]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="191" unit="mm" value="4.3">4.3mm</quantity>
), slightly wider and moderately longer than prothorax (WE/WP: 4.3/
<quantity id="4C809B1A4F203B37C0DB1739FD12463B" box="[687,764,1364,1387]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="191" unit="mm" value="3.5">3.5mm</quantity>
, LE/ LP: 5.8/
<quantity id="4C809B1A4F203B37C3F31718FE3B46DC" box="[391,469,1397,1420]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="191" unit="mm" value="3.5">3.5mm</quantity>
), distinctly punctured with very minute pubescence, moderately convex, dorsal contour highest at middle, lateral contour evenly arcuate, widest at middle. Each elytron with the following scaly markings of pink and tangerine round scales: a) thick transverse basal stripe from stria II towards lateral margin, b) one longitudinal thick stripe on interval III extending from base to apex, c) thick transverse medial band from suture towards lateral margin broadened laterally, d) one long marginal stripe extending from base towards apex confluent with the basal, and medial stripe, and the long stripe at interval III, and e) skewed subcircular spot on apical third confluent with the long stripe on interval II.
<emphasis id="B90CEAED4F203B37C76F134FFABF426A" bold="true" box="[1307,1361,290,314]" pageId="6" pageNumber="191">Legs</emphasis>
with moderately clavate femora. Femora black covered with adpressed setae and light orange elliptical scales near the apical margin. Tibiae covered with subadpressed yellowish setae, weakly serrate along inner edge with few protruding teeth. Fore and mid tibiae bear a mucro at apex. Tarsomeres pubescent. Forecoxae covered with colored piliform scales and with light orange elliptical scales; mesocoxae and metacoxae covered with hairs. Mesoventrite covered with light colored hairs and with light orange round scales on distal ends. Metaventrite sparsely covered with light colored piliform scales and with light orange round scales on distal ends. Ventrite I slightly depressed on disc, covered with light colored piliform scales and with light yellow to orange round scales towards lateral margin. Ventrite II to V sparsely covered with whitish hairs and piliform scales which tends to get denser at distal ends. Ventrite V flattened, apical half finely coarsely rugose, with minute hairs.
</paragraph>
<paragraph id="8BC736FF4F203B37C1261662FAF04776" blockId="6.[850,1310,1039,1062]" box="[850,1310,1039,1062]" pageId="6" pageNumber="191">
Male aedeagus as shown in
<figureCitation id="13432A7A4F203B37C60A1662FAF84776" box="[1150,1302,1039,1062]" captionStart="Figures 3" captionStartId="7.[226,306,1661,1684]" captionTargetBox="[336,1172,245,1583]" captionTargetId="figure-75@7.[272,1241,189,1629]" captionTargetPageId="7" captionText="Figures 3 AE. Male genitalia of Metapocyrtus spp. AC Metapocyrtus (Metapocyrtus) ilgas sp. nov. DE Metapocyrtus (Orthocyrtus) quadratus sp. nov. A, D aedeagus in lateral view B, F idem in dorsal view C sternite IX in dorsal view E aedeagus in dorsolateral view" pageId="6" pageNumber="191">Figures 3 DF</figureCitation>
.
</paragraph>
<paragraph id="8BC736FF4F203B37C126163DFBF747F8" blockId="6.[850,1361,1104,1192]" pageId="6" pageNumber="191">
<emphasis id="B90CEAED4F203B37C126163DFC424738" bold="true" box="[850,940,1104,1128]" pageId="6" pageNumber="191">Female.</emphasis>
Dimensions: LB: 10.0mm: LR:
<quantity id="4C809B1A4F203B37C68A163DFAA34737" box="[1278,1357,1104,1127]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="191" unit="mm" value="1.5">1.5mm</quantity>
: WR: 1.0mm. LP: 4.0mm. WP:4.0mm. LE: 6.0mm. WE: 4.0mm. N=1.
</paragraph>
<paragraph id="8BC736FF4F203B37C12616BFFB3847B9" blockId="6.[850,1238,1234,1257]" box="[850,1238,1234,1257]" pageId="6" pageNumber="191">
Habitus as shown in
<figureCitation id="13432A7A4F203B37C64516BFFB3C47B9" box="[1073,1234,1234,1257]" captionStart="Figure 2" captionStartId="5.[226,296,1693,1716]" captionTargetBox="[256,1257,192,1654]" captionTargetId="figure-56@5.[254,1260,189,1659]" captionTargetPageId="5" captionText="Figure 2. Metapocyrtus (Metapocyrtus) ilgas sp. nov. A, C, Holotype male; A. dorsal view, C. lateral view. B, D, Paratype female; B. dorsal view, D. lateral view" pageId="6" pageNumber="191">Figures 2 B, D</figureCitation>
.
</paragraph>
<paragraph id="8BC736FF4F203B37C126177EFABF46FC" blockId="6.[850,1361,1299,1452]" pageId="6" pageNumber="191">Females differ from males in the following: a) the scaly markings in head, rostrum, pronotum, elytra and legs are yellow­green, b) elytra longer and moderately wider, and d) ventrite I slightly convex on disc. Otherwise female similar to the male.</paragraph>
</subSubSection>
<subSubSection id="C36265744F203B37C12617BBFBCA4520" pageId="6" pageNumber="191" type="etymology">
<paragraph id="8BC736FF4F203B37C12617BBFBCA4520" blockId="6.[850,1361,1494,1648]" pageId="6" pageNumber="191">
<emphasis id="B90CEAED4F203B37C12617BBFC3D46BE" bold="true" box="[850,979,1494,1518]" pageId="6" pageNumber="191">Etymology.</emphasis>
<taxonomicName id="4C784D7C4F203B37C1AC17BAFABE46BE" authorityName="Cabras &amp; Villegas &amp; Dagta &amp; Medina &amp; Barševskis" authorityYear="2022" box="[984,1360,1494,1518]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="ilgas" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B90CEAED4F203B37C1AC17BAFB8246BE" box="[984,1132,1495,1518]" italics="true" pageId="6" pageNumber="191">Metapocyrtus</emphasis>
(
<emphasis id="B90CEAED4F203B37C60E17BAFAE046BE" box="[1146,1294,1495,1518]" italics="true" pageId="6" pageNumber="191">Metapocyrtus</emphasis>
)
<emphasis id="B90CEAED4F203B37C76817BAFABE46BE" box="[1308,1360,1495,1518]" italics="true" pageId="6" pageNumber="191">ilgas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57964F203B37C126179AFC4A455E" box="[850,932,1527,1550]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
is named after the beetle research center of
<collectingRegion id="49BCF81D4F203B37C11A147AFC07457E" box="[878,1001,1559,1582]" country="Latvia" name="Daugavpils" pageId="6" pageNumber="191">Daugavpils</collectingRegion>
University in appreciation for the successful cooperation which led to the discovery of this new species.
</paragraph>
</subSubSection>
<subSubSection id="C36265744F203B39C12614F4FABE43F4" lastPageId="8" lastPageNumber="193" pageId="6" pageNumber="191" type="distribution">
<paragraph id="8BC736FF4F203B37C12614F4FAFA4581" blockId="6.[850,1360,1689,1746]" pageId="6" pageNumber="191">
<emphasis id="B90CEAED4F203B37C12614F4FC0045E1" bold="true" box="[850,1006,1689,1713]" pageId="6" pageNumber="191">Distribution.</emphasis>
<taxonomicName id="4C784D7C4F203B37C18D14F7FC684582" authorityName="Cabras &amp; Villegas &amp; Dagta &amp; Medina &amp; Barševskis" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="ilgas" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B90CEAED4F203B37C18D14F7FB7B45E1" box="[1017,1173,1690,1713]" italics="true" pageId="6" pageNumber="191">Metapocyrtus</emphasis>
(
<emphasis id="B90CEAED4F203B37C6DF14F7FAA945E1" box="[1195,1351,1690,1713]" italics="true" pageId="6" pageNumber="191">Metapocyrtus</emphasis>
)
<emphasis id="B90CEAED4F203B37C12614D6FC684582" box="[850,902,1723,1746]" italics="true" pageId="6" pageNumber="191">ilgas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57964F203B37C1F914D7FC0E4581" box="[909,992,1722,1745]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
is known in its
<typeStatus id="54C3885D4F203B37C6FD14D7FB594581" box="[1161,1207,1722,1745]" pageId="6" pageNumber="191">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BC736FF4F213B36C3B212FCFBCC43F4" blockId="7.[454,1058,143,164]" box="[454,1058,143,164]" pageId="7" pageNumber="192">
<emphasis id="B90CEAED4F213B36C3B212FCFBCC43F4" box="[454,1058,143,164]" italics="true" pageId="7" pageNumber="192">Cabras A.A., Villegas J., Dagta M.J., Medina M.N.D., Barṥevskis A.</emphasis>
</paragraph>
<caption id="DF0766774F213B36C2961410FC004584" pageId="7" pageNumber="192" startId="7.[226,306,1661,1684]" targetBox="[336,1172,245,1583]" targetPageId="7" targetType="figure">
<paragraph id="8BC736FF4F213B36C2961410FC004584" blockId="7.[226,1285,1661,1749]" pageId="7" pageNumber="192">
Figures 3 AE. Male genitalia of
<taxonomicName id="4C784D7C4F213B36C04F1413FCFB45C5" box="[571,789,1661,1685]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B90CEAED4F213B36C04F1413FD2345C5" box="[571,717,1662,1685]" italics="true" pageId="7" pageNumber="192">Metapocyrtus</emphasis>
spp.
<emphasis id="B90CEAED4F213B36C1761410FCFB45C5" bold="true" box="[770,789,1661,1685]" pageId="7" pageNumber="192">A</emphasis>
</taxonomicName>
<emphasis id="B90CEAED4F213B36C1621410FCD845C5" bold="true" box="[790,822,1661,1685]" pageId="7" pageNumber="192">C</emphasis>
<taxonomicName id="4C784D7C4F213B36C14F1413FB4145C5" authorityName="Cabras &amp; Villegas &amp; Dagta &amp; Medina &amp; Barševskis" authorityYear="2022" box="[827,1199,1661,1685]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="ilgas" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B90CEAED4F213B36C14F1413FC2345C5" box="[827,973,1662,1685]" italics="true" pageId="7" pageNumber="192">Metapocyrtus</emphasis>
(
<emphasis id="B90CEAED4F213B36C1AF1413FB8345C5" box="[987,1133,1662,1685]" italics="true" pageId="7" pageNumber="192">Metapocyrtus</emphasis>
)
<emphasis id="B90CEAED4F213B36C60F1413FB4145C5" box="[1147,1199,1662,1685]" italics="true" pageId="7" pageNumber="192">ilgas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57964F213B36C6C71410FAEB45C4" box="[1203,1285,1661,1684]" pageId="7" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B90CEAED4F213B36C29614F0FEFB45E5" bold="true" box="[226,277,1693,1717]" pageId="7" pageNumber="192">DE</emphasis>
<taxonomicName id="4C784D7C4F213B36C36914F3FD2A45E5" authorityName="Cabras &amp; Villegas &amp; Dagta &amp; Medina &amp; Barševskis" authorityYear="2022" box="[285,708,1693,1717]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="192" phylum="Arthropoda" rank="species" species="quadratus" status="sp. nov." subGenus="Orthocyrtus">
<emphasis id="B90CEAED4F213B36C36914F3FE5F45E5" box="[285,433,1694,1717]" italics="true" pageId="7" pageNumber="192">Metapocyrtus</emphasis>
(
<emphasis id="B90CEAED4F213B36C3B714F3FDA845E5" box="[451,582,1694,1717]" italics="true" pageId="7" pageNumber="192">Orthocyrtus</emphasis>
)
<emphasis id="B90CEAED4F213B36C02314F3FD2A45E5" box="[599,708,1694,1717]" italics="true" pageId="7" pageNumber="192">quadratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57964F213B36C0B814F0FCCF45E4" box="[716,801,1693,1716]" pageId="7" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B90CEAED4F213B36C15D14F0FCB145E5" bold="true" box="[809,863,1693,1717]" pageId="7" pageNumber="192">A, D</emphasis>
aedeagus in lateral view
<emphasis id="B90CEAED4F213B36C60C14F0FB4745E5" bold="true" box="[1144,1193,1693,1717]" pageId="7" pageNumber="192">B, F</emphasis>
idem in dorsal view
<emphasis id="B90CEAED4F213B36C31114D0FE964585" bold="true" box="[357,376,1725,1749]" pageId="7" pageNumber="192">C</emphasis>
sternite IX in dorsal view
<emphasis id="B90CEAED4F213B36C0E314D0FD474585" bold="true" box="[663,681,1725,1749]" pageId="7" pageNumber="192">E</emphasis>
aedeagus in dorsolateral view
</paragraph>
</caption>
<paragraph id="8BC736FF4F2E3B39C35A12FCFABE43F4" blockId="8.[302,1360,145,164]" box="[302,1360,145,164]" pageId="8" pageNumber="193">
<emphasis id="B90CEAED4F2E3B39C35A12FCFABE43F4" box="[302,1360,145,164]" italics="true" pageId="8" pageNumber="193">
Two new species of
<taxonomicName id="4C784D7C4F2E3B39C3A212FCFD5243F4" authority="Heller, 1912" authorityName="Heller" authorityYear="1912" box="[470,700,145,164]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="193" phylum="Arthropoda" rank="genus">Metapocyrtus Heller, 1912</taxonomicName>
(
<taxonomicName id="4C784D7C4F2E3B39C0B312FCFCC243F4" authorityName="Linnaeus" authorityYear="1758" box="[711,812,145,164]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="8" pageNumber="193" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
,
<taxonomicName id="4C784D7C4F2E3B39C14012FCFC5D43F4" box="[820,947,145,164]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="193" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
,
<taxonomicName id="4C784D7C4F2E3B39C1CF12FCFBFB43F4" box="[955,1045,145,164]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="193" phylum="Arthropoda" rank="subFamily" subFamily="Entiminae">Entiminae</taxonomicName>
) from Lanao and Panay,
<collectingCountry id="F36F766F4F2E3B39C69912FCFABE43F4" box="[1261,1360,145,164]" name="Philippines" pageId="8" pageNumber="193">Philippines</collectingCountry>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,179 @@
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<mods:title id="CA406803357453F083B5C05E9734149B">A new genus and species of Ernobiinae (Coleoptera: Ptinidae) from Eocene Baltic amber of the Sambian Peninsula</mods:title>
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<subSubSection id="C3616515FC401F37FD8B4789FC71FE71" box="[624,940,766,791]" pageId="1" type="nomenclature">
<paragraph id="8BC4369EFC401F37FD8B4789FC71FE71" blockId="1.[624,940,766,791]" box="[624,940,766,791]" pageId="1">
<heading id="D08C81F2FC401F37FD8B4789FC71FE71" bold="true" box="[624,940,766,791]" fontSize="10" level="2" pageId="1" reason="4">
<emphasis id="B90FEA8CFC401F37FD8B4789FC71FE71" bold="true" box="[624,940,766,791]" pageId="1">
Genus
<taxonomicName id="4C7B4D1DFC401F37FD444788FC9DFE71" authorityName="Alekseev &amp; Bukejs &amp; Háva &amp; Zahradník" authorityYear="2023" box="[703,832,767,791]" class="Insecta" family="Ptinidae" genus="Eocenobius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FEA8CFC401F37FD444788FC9DFE71" bold="true" box="[703,832,767,791]" italics="true" pageId="1">Eocenobius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57F7FC401F37FCBD4788FC71FE71" box="[838,940,767,791]" pageId="1" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3616515FC401F37FD8B464BFC7EFE14" pageId="1" type="materials_examined">
<paragraph id="8BC4369EFC401F37FD8B464BFC7EFE14" blockId="1.[624,1102,828,882]" pageId="1">
<typeStatus id="54C0883CFC401F37FD8B464BFD7AFE32" box="[624,679,828,852]" pageId="1">Type</typeStatus>
species:
<taxonomicName id="4C7B4D1DFC401F37FCFE464BFBC2FE35" authorityName="Alekseev &amp; Bukejs &amp; Háva &amp; Zahradník" authorityYear="2023" box="[773,1055,828,852]" class="Insecta" family="Ptinidae" genus="Eocenobius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="praestigitator" status="sp. nov.">
<emphasis id="B90FEA8CFC401F37FCFE464BFBC2FE35" box="[773,1055,828,852]" italics="true" pageId="1">Eocenobius praestigitator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57F7FC401F37FBD1464BFD7CFE14" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
, by present designation.
</paragraph>
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<subSubSection id="C3616515FC401F34FD8B46E0FDF4FF3E" lastPageId="2" pageId="1" type="diagnosis">
<paragraph id="8BC4369EFC401F37FD8B46E0FBFBF984" blockId="1.[624,1103,919,1250]" pageId="1">
<emphasis id="B90FEA8CFC401F37FD8B46E0FCAAFED6" bold="true" box="[624,887,919,944]" pageId="1">Differential diagnosis.</emphasis>
<taxonomicName id="4C7B4D1DFC401F37FC7046EFFBD5FED6" authorityName="Alekseev &amp; Bukejs &amp; Háva &amp; Zahradník" authorityYear="2023" box="[907,1032,920,944]" class="Insecta" family="Ptinidae" genus="Eocenobius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FEA8CFC401F37FC7046EFFBD5FED6" box="[907,1032,920,944]" italics="true" pageId="1">Eocenobius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57F7FC401F37FBE046EFFD42FEA8" pageId="1" rank="genus">gen. nov.</taxonomicNameLabel>
is characterized by the following combination of characters:(1) antennae 11- segmented withfiliform antennomeres 38, and elongate, three-segmented club; (2) lateral margins of pronotum complete; (3) elytra short and forming oval outline; (4) dorsal pubescence thin, sparse, long, and erect; (5) meso- and metacoxae widely separated; (6) elytral punctation relatively sparse; and (7) metatrochanters angulate.
</paragraph>
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The newly described genus can be distinguished from the genus
<taxonomicName id="4C7B4D1DFC401F37FC834051FC07F858" box="[888,986,1318,1342]" class="Insecta" family="Anobiidae" genus="Ernobius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC401F37FC834051FC07F858" box="[888,986,1318,1342]" italics="true" pageId="1">Ernobius</emphasis>
</taxonomicName>
in having shorter elytra (1.5× longer than wide in the new genus and 1.92.9× longer than wide in
<taxonomicName id="4C7B4D1DFC401F37FD6240F5FD26F8FC" box="[665,763,1410,1434]" class="Insecta" family="Anobiidae" genus="Ernobius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC401F37FD6240F5FD26F8FC" box="[665,763,1410,1434]" italics="true" pageId="1">Ernobius</emphasis>
</taxonomicName>
), sparser elytral punctation (distance between punctures usually less than diameter of one puncture in
<taxonomicName id="4C7B4D1DFC401F37FC2C40B7FBE4F8BE" box="[983,1081,1472,1496]" class="Insecta" family="Anobiidae" genus="Ernobius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC401F37FC2C40B7FBE4F8BE" box="[983,1081,1472,1496]" italics="true" pageId="1">Ernobius</emphasis>
</taxonomicName>
), erect dorsal pubescence (recumbent in
<taxonomicName id="4C7B4D1DFC401F37FD8B408AFD0FFB73" box="[624,722,1533,1557]" class="Insecta" family="Anobiidae" genus="Ernobius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC401F37FD8B408AFD0FFB73" box="[624,722,1533,1557]" italics="true" pageId="1">Ernobius</emphasis>
</taxonomicName>
), as well as comparatively wide separated meso- and metacoxae.
<taxonomicName id="4C7B4D1DFC401F37FC20436BFD55FB34" authorityName="Alekseev &amp; Bukejs &amp; Háva &amp; Zahradník" authorityYear="2023" class="Insecta" family="Ptinidae" genus="Eocenobius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FEA8CFC401F37FC20436BFD55FB34" italics="true" pageId="1">Eocenobius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57F7FC401F37FD63434CFCD8FB34" box="[664,773,1595,1619]" pageId="1" rank="genus">gen. nov.</taxonomicNameLabel>
can additionally be distinguished from extinct
<taxonomicName id="4C7B4D1DFC401F34FC50432EFE12FDAC" authority="Zahradnik et Hava, 2014" authorityName="Zahradnik et Hava" authorityYear="2014" class="Insecta" family="Ptinidae" genus="Tuberernobius" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="2" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC401F37FC50432EFB95FB17" box="[939,1096,1625,1649]" italics="true" pageId="1">Tuberernobius</emphasis>
<bibRefCitation id="EFEA4B6FFC431F34FF4645C5FE12FDAC" author="Zahradnik P. &amp; Hava J." box="[189,463,178,202]" pageId="2" pagination="629 - 646" refId="ref4627" refString="Zahradnik P., Hava J. 2014. New Ptinidae (Coleoptera: Bostrichoidea) from Baltic amber with a list of known fossil species. Studies and Reports. Taxonomical Series, 10 (2): 629 - 646." type="journal article" year="2014">
Zahradník et Háva,
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</taxonomicName>
in the absence of knob-like tubercles on the pronotum; from
<taxonomicName id="4C7B4D1DFC431F34FF464598FE60FC61" authority="Fall, 1905" authorityName="Fall" authorityYear="1905" box="[189,445,239,263]" class="Insecta" family="Anobiidae" genus="Paralobium" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC431F34FF464598FEE0FC61" box="[189,317,239,263]" italics="true" pageId="2">Paralobium</emphasis>
Fall,
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</taxonomicName>
in possessing filiform antennomeres 38;and from the genera
<taxonomicName id="4C7B4D1DFC431F34FF1D445AFDC4FC22" authority="Thomson, 1863" authorityName="Thomson" authorityYear="1863" box="[230,537,300,324]" class="Insecta" family="Anobiidae" genus="Episernus" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC431F34FF1D445AFE8CFC22" box="[230,337,301,324]" italics="true" pageId="2">Episernus</emphasis>
Thomson, 1863
</taxonomicName>
and
<taxonomicName id="4C7B4D1DFC431F34FD93445AFE29FC05" authority="Roubal, 1917" authorityName="Roubal" authorityYear="1917" class="Insecta" family="Anobiidae" genus="Episernomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC431F34FD93445AFE8BFC05" italics="true" pageId="2">Episernomorphus</emphasis>
Roubal,
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</taxonomicName>
in having 11- segmented antennae. For comparison with extinct
<taxonomicName id="4C7B4D1DFC431F34FEED44FFFE56FCC6" authorityName="Pic" authorityYear="1912" box="[278,395,392,416]" class="Insecta" family="Ptinidae" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="subFamily" subFamily="Ernobiinae">Ernobiinae</taxonomicName>
taxa of Baltic amber, also see the key presented below. Additionally,
<taxonomicName id="4C7B4D1DFC431F34FF0F44B2FEACFCBB" authorityName="Alekseev &amp; Bukejs &amp; Háva &amp; Zahradník" authorityYear="2023" box="[244,369,453,477]" class="Insecta" family="Ptinidae" genus="Eocenobius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FEA8CFC431F34FF0F44B2FEACFCBB" box="[244,369,453,477]" italics="true" pageId="2">Eocenobius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57F7FC431F34FE8244B1FE00FCBB" box="[377,477,454,478]" pageId="2" rank="genus">gen. nov.</taxonomicNameLabel>
shows similarity to extant
<taxonomicName id="4C7B4D1DFC431F34FED64493FDC3FC9A" authority="Dejean, 1821" authorityName="Dejean" authorityYear="1821" box="[301,542,484,508]" class="Insecta" family="Ptinidae" genus="Ochina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC431F34FED64493FEA1FC9A" box="[301,380,484,508]" italics="true" pageId="2">Ochina</emphasis>
Dejean,
<quantity id="4C839B7BFC431F34FE124493FDC3FC9A" box="[489,542,484,508]" metricMagnitude="1" metricUnit="m" metricValue="4.62534" pageId="2" unit="in" value="1821.0">1821</quantity>
</taxonomicName>
in having comparatively short elytra. However,
<taxonomicName id="4C7B4D1DFC431F34FDA24774FF05FF5F" authorityName="Dejean" authorityYear="1821" class="Insecta" family="Ptinidae" genus="Ochina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEA8CFC431F34FDA24774FF05FF5F" italics="true" pageId="2">Ochina</emphasis>
</taxonomicName>
differs from the new genus in having weakly serrate antennomeres 38.
</paragraph>
</subSubSection>
<subSubSection id="C3616515FC431F34FF46470CFDD5FE4A" pageId="2" type="etymology">
<paragraph id="8BC4369EFC431F34FF46470CFDD5FE4A" blockId="2.[189,668,635,812]" pageId="2">
<emphasis id="B90FEA8CFC431F34FF46470CFE9BFFF5" bold="true" box="[189,326,635,659]" pageId="2">Etymology.</emphasis>
The name of the new genus is a compound word and combines “Eocene” (the name of the
<typeStatus id="54C0883CFC431F34FE7147CFFE6AFFB6" box="[394,439,696,720]" pageId="2">type</typeStatus>
species geological epoch) and “-
<emphasis id="B90FEA8CFC431F34FEA047A1FE5AFF88" box="[347,391,726,750]" italics="true" pageId="2">bius</emphasis>
” (the Latin root of the
<typeStatus id="54C0883CFC431F34FF464781FF36FE68" box="[189,235,758,782]" pageId="2">type</typeStatus>
genus of the tribe to which the genus belongs). Gender is masculine.
</paragraph>
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<subSubSection id="C3616515FC431F34FF464639FDEAFEC5" pageId="2" type="discussion">
<paragraph id="8BC4369EFC431F34FF464639FDEAFEC5" blockId="2.[189,669,846,932]" pageId="2">
<emphasis id="B90FEA8CFC431F34FF464639FEF1FE00" bold="true" box="[189,300,846,870]" pageId="2">Remarks.</emphasis>
The new genus is monotypic. Therefore, the generic diagnosis is currently identical to that of the
<typeStatus id="54C0883CFC431F34FE5546FBFE01FEC2" box="[430,476,908,932]" pageId="2">type</typeStatus>
species.
</paragraph>
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@ -0,0 +1,81 @@
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<mods:title id="B001B726A70E738D17ABD7DEE70F5E88">A new genus and species of Ernobiinae (Coleoptera: Ptinidae) from Eocene Baltic amber of the Sambian Peninsula</mods:title>
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Tribe
<taxonomicName id="4C7B4D1DFC401F37FD4D4445FC4DFC2D" authority="Pic, 1912" authorityName="Pic" authorityYear="1912" box="[694,912,306,331]" class="Insecta" family="Ptinidae" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="tribe" tribe="Ernobiini">Ernobiini Pic, 1912</taxonomicName>
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<emphasis id="B90FEA8CFC401F37FD8B4407FD38FCEE" bold="true" box="[624,741,368,392]" pageId="1">Remarks.</emphasis>
The fossil specimen under consideration is placed in the tribe
<taxonomicName id="4C7B4D1DFC401F37FC1F44F9FB92FCC0" authorityName="Pic" authorityYear="1912" box="[996,1103,398,422]" class="Insecta" family="Ptinidae" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="tribe" tribe="Ernobiini">Ernobiini</taxonomicName>
within the subfamily
<taxonomicName id="4C7B4D1DFC401F37FC9B44DAFC08FCA3" authorityName="Pic" authorityYear="1912" box="[864,981,429,453]" class="Insecta" family="Ptinidae" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="subFamily" subFamily="Ernobiinae">Ernobiinae</taxonomicName>
(
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) based on: having antennae that are 11- segmented, with a loose, 3-segmented club; tarsal formula 5-5-5; antennal insertions widely separated; head free, prognathous, not extended posteriorly; elytra with irregular punctation; legs not received in thoracic cavities; prothorax not excavated beneath for reception of profemora; and pronotum with lateral margins.
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<taxonomicName id="4C7B4D1DFC431F34FF464695FE0AFE9C" authorityName="Alekseev &amp; Bukejs &amp; Háva &amp; Zahradník" authorityYear="2023" box="[189,471,994,1018]" class="Insecta" family="Ptinidae" genus="Eocenobius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="praestigitator" status="sp. nov.">
<emphasis id="B90FEA8CFC431F34FF464695FE0AFE9C" bold="true" box="[189,471,994,1018]" italics="true" pageId="2">Eocenobius praestigitator</emphasis>
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(
<figureCitation id="13402A1BFC431F34FF3E4177FEF7F97E" box="[197,298,1024,1048]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[114,153,1589,1613]" captionStartId-1="4.[189,228,922,946]" captionTargetBox-0="[114,1073,171,1552]" captionTargetBox-1="[189,1181,171,885]" captionTargetId-0="figure-38@3.[114,1073,171,1552]" captionTargetId-1="figure-271@4.[189,1181,171,885]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 1. Eocenobius praestigitator sp. nov., holotype, No AWI-165 [CVIA]: A habitus, dorsal view; B habitus, ventro-lateral view. Scale bar = 1 mm." captionText-1="Fig. 2. Eocenobius praestigitator sp. nov., holotype, No AWI-165 [CVIA]: A habitus, ventral view; B antenna (reconstruction)." pageId="2">Figs. 12</figureCitation>
)
</paragraph>
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<subSubSection id="C3616515FC431F34FF46414AFECEF8A7" pageId="2" type="materials_examined">
<paragraph id="8BC4369EFC431F34FF46414AFE67F80F" blockId="2.[189,669,1084,1385]" pageId="2">
<emphasis id="B90FEA8CFC431F34FF46414AFEABF933" bold="true" box="[189,374,1084,1109]" pageId="2">Type material.</emphasis>
<typeStatus id="54C0883CFC431F34FE7A414AFE35F933" box="[385,488,1085,1109]" pageId="2" type="holotype">Holotype</typeStatus>
: No AWI-165 [
<collectionCode id="ED6AAE5BFC431F34FF3F412BFED0F912" box="[196,269,1116,1140]" pageId="2">CVIA</collectionCode>
]; adult, sex unknown. Almost complete beetle (right antenna absent) included in a transparent, yellow, flat amber piece of irregular form with dimensions of 21×
<quantity id="4C839B7BFC431F34FD8741CFFF3AF988" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" unit="mm" value="13.0">13 mm</quantity>
and a maximum thickness of
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; preserved without supplementary fixation. Syninclusions: several detrital particles, two plant trichomes, two legs of an insect (likely Diptera: Tipuloidea).
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<typeStatus id="54C0883CFC431F34FF4640FBFF2AF8C2" box="[189,247,1420,1444]" pageId="2">Type</typeStatus>
stratum.
</emphasis>
Baltic amber; MiddleLate Eocene.
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<emphasis id="B90FEA8CFC431F34FD4045C6FC91FDAC" bold="true" box="[699,844,177,202]" pageId="2">Description.</emphasis>
Measurements: body length (from anterior margin of pronotum to elytral apex along midline)
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, body maximum width across both elytra
<quantity id="4C839B7BFC431F34FB9E447AFD35FC22" metricMagnitude="-3" metricUnit="m" metricValue="1.36" pageId="2" unit="mm" value="1.36">1.36 mm</quantity>
; head length
<quantity id="4C839B7BFC431F34FC87445BFC3DFC22" box="[892,992,300,324]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="2" unit="mm" value="0.57">0.57 mm</quantity>
, head maximum width across eyes
<quantity id="4C839B7BFC431F34FC7A443CFC38FC04" box="[897,997,331,355]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="2" unit="mm" value="0.69">0.69 mm</quantity>
; pronotal length
<quantity id="4C839B7BFC431F34FD40441DFCF8FCE7" box="[699,805,362,386]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="2" unit="mm" value="0.43">0.43 mm</quantity>
, pronotal maximum width
<quantity id="4C839B7BFC431F34FB9E441DFD35FCC6" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="2" unit="mm" value="1.14">1.14 mm</quantity>
; elytral length
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, elytral maximum width
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. Body moderately convex; integument dark brown (as preserved); pubescence conspicuous, long, thin, erect.
</paragraph>
<paragraph id="8BC4369EFC431F34FD40474AFCA2F8A8" blockId="2.[699,1179,573,1486]" pageId="2">
Head prognathous, mostly hidden in dorsal view, almost as long as wide (including eyes); finely punctate, with distance between punctures 1.01.5× diameter of one puncture. Compound eyes protruding, large, finely facetted, hemispherical, weakly impressed near antennal bases. Ommatidial setae absent. Antennae inserted at inner margin of eyes; antennal insertions widely separated, distance between antennal insertions equal to length of antennomeres 15. Antennae 11-segmented, with loose, 3- segmented club; antennae moderately long, reaching abdominal ventrite
<quantity id="4C839B7BFC431F34FBF346BBFBE7FE82" box="[1032,1082,972,996]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" unit="in" value="1.0">1 in</quantity>
ventral position. Antennomere 1 subcylindrical, elongate, 2.0× longer than wide; antennomere 2 subovate, slightly dilated apically, 1.5× longer than wide; antennomere 3 elongate, 2.5× longer than wide; antennomeres 48subequal in length, elongate, 1.51.8× longer than wide; antennomeres 910 elongate, slightly flattened, about 3.3× longer than antennomere8; antennomere 11 spindle-shaped, with acute apex, 3.0× longer than wide. Relative length ratios of antennomeres 111 equal to 6:4:5:3:3:3:3:3:10: 10:11. Apical maxillary palpomere short, cylindrical, slightly longer than wide, truncated apically. Apical labial palpomere triangular in outline.
</paragraph>
</subSubSection>
<subSubSection id="C3616515FC431F33FF464092FEFEFE2B" lastPageId="5" pageId="2" type="materials_examined">
<paragraph id="8BC4369EFC431F34FF464092FED1FB5C" blockId="2.[189,668,1508,1594]" pageId="2">
<materialsCitation id="3B133CC3FC431F34FF464092FED5FB5C" country="Russia" location="Yantarny village" pageId="2" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B90FEA8CFC431F34FF464092FEB9F89B" bold="true" box="[189,356,1508,1533]" pageId="2">
<typeStatus id="54C0883CFC431F34FF464092FF2AF89B" box="[189,247,1509,1533]" pageId="2">Type</typeStatus>
locality.
</emphasis>
<location id="8EA46045FC431F34FE914092FDFEF89A" LSID="urn:lsid:plazi:treatment:03D28788FC431F33FF464695FF39FE81:8EA46045FC431F34FE914092FDFEF89A" box="[362,547,1508,1533]" country="Russia" name="Yantarny village" pageId="2" stateProvince="Kaliningrad">Yantarny village</location>
(formerly Palmnicken), the
<collectingRegion id="49BFF87CFC431F34FE704374FDB4FB7D" box="[395,617,1539,1563]" country="Russia" name="Kaliningrad" pageId="2">Kaliningrad Region</collectingRegion>
, W
<collectingCountry id="F36C760EFC431F34FF464355FED5FB5C" box="[189,264,1570,1594]" name="Russia" pageId="2">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC4369EFC431F34FB8243D1FB49FBD8" blockId="2.[1145,1172,1702,1726]" box="[1145,1172,1702,1726]" pageId="2">53</paragraph>
<caption id="DF046616FC421F35FF894342FCC0FB0D" pageId="3" startId="3.[114,153,1589,1613]" targetBox="[114,1073,171,1552]" targetPageId="3" targetType="figure">
<paragraph id="8BC4369EFC421F35FF894342FCC0FB0D" blockId="3.[114,1103,1589,1644]" pageId="3">
Fig. 1.
<taxonomicName id="4C7B4D1DFC421F35FF3E4342FE02FB2B" authorityName="Alekseev &amp; Bukejs &amp; Háva &amp; Zahradník" authorityYear="2023" box="[197,479,1589,1613]" class="Insecta" family="Ptinidae" genus="Eocenobius" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="praestigitator" status="sp. nov.">
<emphasis id="B90FEA8CFC421F35FF3E4342FE02FB2B" box="[197,479,1589,1613]" italics="true" pageId="3">Eocenobius praestigitator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57F7FC421F35FE124341FD9EFB2B" box="[489,579,1590,1613]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, No AWI-165 [CVIA]: A habitus, dorsal view; B habitus, ventro-lateral view. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8BC4369EFC421F35FF8943D1FF50FBD8" blockId="3.[114,141,1702,1726]" box="[114,141,1702,1726]" pageId="3">54</paragraph>
<paragraph id="8BC4369EFC451F32FF464514FB4EFD10" blockId="4.[189,1171,99,118]" box="[189,1171,99,118]" pageId="4">
<emphasis id="B90FEA8CFC451F32FF464514FB4EFD10" box="[189,1171,99,118]" italics="true" pageId="4">
A new genus and species of
<taxonomicName id="4C7B4D1DFC451F32FE514514FDD7FD10" authorityName="Pic" authorityYear="1912" box="[426,522,99,118]" class="Insecta" family="Ptinidae" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="subFamily" subFamily="Ernobiinae">Ernobiinae</taxonomicName>
(
<taxonomicName id="4C7B4D1DFC451F32FDED4514FDA4FD10" authorityName="Linnaeus" authorityYear="1758" box="[534,633,99,118]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C7B4D1DFC451F32FD794514FD16FD10" authorityName="Latreille" authorityYear="1802" box="[642,715,99,118]" class="Insecta" family="Ptinidae" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="family">Ptinidae</taxonomicName>
) from Eocene Baltic amber of the Sambian Peninsula
</emphasis>
</paragraph>
<caption id="DF046616FC451F32FF4646EDFD55FEB7" pageId="4" startId="4.[189,228,922,946]" targetBox="[189,1181,171,885]" targetPageId="4" targetType="figure">
<paragraph id="8BC4369EFC451F32FF4646EDFD55FEB7" blockId="4.[189,1178,922,977]" pageId="4">
Fig. 2.
<taxonomicName id="4C7B4D1DFC451F32FEEB46EDFDF7FED4" authorityName="Alekseev &amp; Bukejs &amp; Háva &amp; Zahradník" authorityYear="2023" box="[272,554,922,946]" class="Insecta" family="Ptinidae" genus="Eocenobius" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="praestigitator" status="sp. nov.">
<emphasis id="B90FEA8CFC451F32FEEB46EDFDF7FED4" box="[272,554,922,946]" italics="true" pageId="4">Eocenobius praestigitator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57F7FC451F32FDCF46ECFD53FED4" box="[564,654,923,946]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, No AWI-165 [CVIA]: A habitus, ventral view; B antenna (reconstruction).
</paragraph>
</caption>
<paragraph id="8BC4369EFC451F32FF464680FDCDF8BC" blockId="4.[189,669,1014,1498]" pageId="4">Pronotum slightly narrower than elytra at base, strongly transverse, 2.65× wider than long, with maximum width at base; lateral sides and anterior edge finely margined. Pronotal punctation fine, denser at base, about as large as punctation of head; distance between punctures equal to 1.02.0× diameter of one puncture. Pronotal pubescence fine, erect, comparatively short, with setae inconspicuous on disc and longer laterally. Posterior pronotal angles obtuse, rounded; anterior angles rectangular with rounded apices. Anterior pronotal margin rounded, without excision, and not raised; posterior edge bisinuate; lateral sides almost straight, smooth, reflexed.</paragraph>
<paragraph id="8BC4369EFC451F32FF464377FC44F89F" blockId="4.[189,668,1536,1652]" lastBlockId="4.[699,1179,1014,1529]" pageId="4">Scutellar shield small, semicircular, transverse. Elytra short and oval, moderately convex, with maximum width occurring near mid-length, 1.5× longer than wide, 4.74× as long as pronotal length; humeri rounded, distinct. Elytral punctation irregular, fine, with punctures slightly larger than punctation of pronotum and head; distance between punctures equal to 1.52.0× diameter of one puncture. Elytral pubescence fine, semierect and shorter on disc, erect and longer laterally. Epipleura narrow extending to abdominal ventrite 1. Metaventrite with disc convex; covered with coarse, rather dense punctation; posterior margin between metacoxae straight, with small triangular notch in middle. Mesocoxaeoval, separated by distance equal to 1.5× their transverse diameter. Metacoxae strongly transverse, separated by distance equal to their length.</paragraph>
<paragraph id="8BC4369EFC451F32FD404369FB47FB15" blockId="4.[699,1178,1566,1652]" pageId="4">All legs are relatively consistent in shape, and finely punctate. Mesocoxaeoval, transverse; metacoxae strongly transverse,</paragraph>
<paragraph id="8BC4369EFC451F33FB8243D1FDF5FC9A" blockId="4.[1145,1172,1702,1726]" lastBlockId="5.[114,593,178,508]" lastPageId="5" pageId="4">55 grooved for reception of metatibiae. Metatrochanters large, transverse, angulate. Femora elongate-oval, flattened. Tibiae thin, almost straight, nearly as long as femora. Meso- and metatibiae with one apical spur. Tarsal formula 5-5-5. Tarsi withtarsomeres not lobed; tarsomere 1 the longest; tarsomeres 35 widened, short, equal in length. Relative length ratios of metatarsomeres 15 equal to 10:5:3:3:3. Pretarsal claws strongly curved, thickened at base.</paragraph>
<paragraph id="8BC4369EFC441F33FF894755FE22FFD2" blockId="5.[114,593,546,692]" pageId="5">Abdomen with five visible, freely articulat- ed ventrites, covered with rather densepunctation; ventrite 5 rounded apically. Relative length ratios of ventrites 15 equal to 12:15:12:10:13 (measured medially).</paragraph>
<paragraph id="8BC4369EFC441F33FF8947ADFEFEFE2B" blockId="5.[114,592,730,845]" pageId="5">
<emphasis id="B90FEA8CFC441F33FF8947ADFF69FF94" bold="true" box="[114,180,730,754]" pageId="5">Note.</emphasis>
Procoxal area and prothorax not visible in
<typeStatus id="54C0883CFC441F33FF46478EFF36FE77" box="[189,235,761,785]" pageId="5">type</typeStatus>
specimen due to head position and milky covering of inclusion within surrounding amber.
</paragraph>
</subSubSection>
<subSubSection id="C3616515FC441F33FF894604FF39FE81" pageId="5" type="etymology">
<paragraph id="8BC4369EFC441F33FF894604FF39FE81" blockId="5.[114,593,883,999]" pageId="5">
<emphasis id="B90FEA8CFC441F33FF894604FF29FEED" bold="true" box="[114,244,883,907]" pageId="5">Etymology.</emphasis>
Specific epithet is Latin word
<taxonomicName id="4C7B4D1DFC441F33FF8946E5FED8FECF" authorityName="Alekseev &amp; Bukejs &amp; Háva &amp; Zahradník" authorityYear="2023" box="[114,261,914,937]" class="Insecta" family="Ptinidae" genus="Eocenobius" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="praestigitator">
<emphasis id="B90FEA8CFC441F33FF8946E5FED8FECF" box="[114,261,914,937]" italics="true" pageId="5">praestigitator</emphasis>
</taxonomicName>
, used as noun in apposition and meaning “trickster, juggler, deceiver, impostor”.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,113 @@
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<mods:title id="2412321C70A99AA90A2795B79B073F52">To The Knowledge Onmetapocyrtusheller, 1912 (Coleoptera: Curculionidae: Pachyrhynchini) Species From Sibuyan Island, With Description Of New Species</mods:title>
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<subSubSection id="C360651CFFF8FFAF07785DFBFB88FE6D" box="[624,1081,465,490]" pageId="1" pageNumber="276" type="nomenclature">
<paragraph id="8BC53697FFF8FFAF07785DFBFB88FE6D" blockId="1.[624,1081,465,520]" box="[624,1081,465,490]" pageId="1" pageNumber="276">
<heading id="D08D81FBFFF8FFAF07785DFBFB88FE6D" bold="true" box="[624,1081,465,490]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="276" reason="4">
<taxonomicName id="4C7A4D14FFF8FFAF07785DFBFB88FE6D" ID-CoL="42CL6" authority="Schultze, 1925" authorityName="Schultze" authorityYear="1925" box="[624,1081,465,490]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="vestitus" subGenus="Trachycyrtus">
<emphasis id="B90EEA85FFF8FFAF07785DFBFB88FE6D" bold="true" box="[624,1081,465,490]" pageId="1" pageNumber="276">
<emphasis id="B90EEA85FFF8FFAF07785DFBFD24FE6E" bold="true" box="[624,661,466,490]" italics="true" pageId="1" pageNumber="276">M.</emphasis>
(
<emphasis id="B90EEA85FFF8FFAF07965DFBFC9EFE6E" bold="true" box="[670,815,466,490]" italics="true" pageId="1" pageNumber="276">Trachycyrtus</emphasis>
)
<emphasis id="B90EEA85FFF8FFAF06365DFBFC27FE6E" bold="true" box="[830,918,466,490]" italics="true" pageId="1" pageNumber="276">vestitus</emphasis>
<bibRefCitation id="EFEB4B66FFF8FFAF069E5DF8FB88FE6D" author="Schultze W." box="[918,1081,465,489]" pageId="1" pageNumber="276" pagination="131 - 310" refId="ref2426" refString="Schultze W. 1925. A monograph of the pachyrrhynchid group of the Brachyderinse, Curculionidme: Part III. The genera Apocyrtidius Heller and Metapocyrtus Heller. The Philippine Journal of Science 26 (2): 131 - 310." type="journal article" year="1925">Schultze, 1925</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C360651CFFF8FFAF07785DD9FD0CF9C7" pageId="1" pageNumber="276" type="description">
<paragraph id="8BC53697FFF8FFAF07785DD9FD0EFD8C" blockId="1.[624,1081,465,520]" box="[624,703,496,520]" pageId="1" pageNumber="276">
(
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)
</paragraph>
<caption id="DF05661FFFF8FFAF057A58D1FC39FA94" box="[114,904,1272,1296]" pageId="1" pageNumber="276" startId="1.[114,153,1272,1296]" targetBox="[117,1100,579,1255]" targetPageId="1" targetType="figure">
<paragraph id="8BC53697FFF8FFAF057A58D1FC39FA94" blockId="1.[114,904,1272,1296]" box="[114,904,1272,1296]" pageId="1" pageNumber="276">
Fig. 1.
<taxonomicName id="4C7A4D14FFF8FFAF05B558D1FDCFFA94" authority="Schultze, 1925" authorityName="Schultze" authorityYear="1925" box="[189,638,1272,1296]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="vestitus" subGenus="Trachycyrtus">
<emphasis id="B90EEA85FFF8FFAF05B558D1FE6DFA94" box="[189,476,1272,1296]" italics="true" pageId="1" pageNumber="276">M. (Trachycyrtus) vestitus</emphasis>
<bibRefCitation id="EFEB4B66FFF8FFAF04D458D1FDCFFA94" author="Schultze W." box="[476,638,1272,1296]" pageId="1" pageNumber="276" pagination="131 - 310" refId="ref2426" refString="Schultze W. 1925. A monograph of the pachyrrhynchid group of the Brachyderinse, Curculionidme: Part III. The genera Apocyrtidius Heller and Metapocyrtus Heller. The Philippine Journal of Science 26 (2): 131 - 310." type="journal article" year="1925">Schultze, 1925</bibRefCitation>
</taxonomicName>
, Holotype, male (MTD).
</paragraph>
</caption>
<paragraph id="8BC53697FFF8FFAF057A591CFDE1F9E6" blockId="1.[114,593,1333,1634]" pageId="1" pageNumber="276">
<emphasis id="B90EEA85FFF8FFAF057A591CFF47FAC9" bold="true" box="[114,246,1333,1357]" pageId="1" pageNumber="276">Description</emphasis>
(
<bibRefCitation id="EFEB4B66FFF8FFAF040C591CFE18FAC9" author="Schultze W." box="[260,425,1333,1357]" pageId="1" pageNumber="276" pagination="131 - 310" refId="ref2426" refString="Schultze W. 1925. A monograph of the pachyrrhynchid group of the Brachyderinse, Curculionidme: Part III. The genera Apocyrtidius Heller and Metapocyrtus Heller. The Philippine Journal of Science 26 (2): 131 - 310." type="journal article" year="1925">Schultze, 1925</bibRefCitation>
): Black, with sparsely scattered pale-yellow wish scales. Elytra short, subovate, with indistinct longitudinal rows of minutely fine hairs and sparsely scattered scales. Rostrum densely confluently punctured, beset with fine whitish hair, basal half with a rather strongly pronounced medial depression and two irregular, sub rugose longitudinal ridges. Basal transverse groove confluent with a pit
</paragraph>
<paragraph id="8BC53697FFF8FFAF057A5A8FFD0CF9C7" blockId="1.[114,154,1702,1726]" lastBlockId="1.[624,1103,1333,1603]" pageId="1" pageNumber="276">276 like depression on sides, in front of eye, and the latter united with antennal scrobe. Front rugosely punctured with a sharply defined medial groove and some scattered scales. Prothorax sub globular, densely, and uniformly roundish granulate, sparsely scattered with yellowish scales. Legs dark brown, moderately densely beset with setae.</paragraph>
</subSubSection>
<subSubSection id="C360651CFFFBFFAC05B55C98FEB8FE06" pageId="2" pageNumber="277" type="materials_examined">
<paragraph id="8BC53697FFFBFFAC05B55C98FEB8FE06" blockId="2.[189,668,177,386]" pageId="2" pageNumber="277">
<emphasis id="B90EEA85FFFBFFAC05B55C98FE24FF4D" bold="true" box="[189,405,177,201]" pageId="2" pageNumber="277">Material examined</emphasis>
:
<specimenCount id="9D7CFD1EFFFBFFAC04AC5C9BFE6DFF4E" box="[420,476,178,202]" count="6" pageId="2" pageNumber="277" type="generic">6 ex.</specimenCount>
: Male
<typeStatus id="54C18835FFFBFFAC072D5C9BFDEDFF4E" box="[549,604,178,202]" pageId="2" pageNumber="277">Type</typeStatus>
(
<figureCitation id="13412A12FFFBFFAC07645C9BFF78FF6C" captionStart="Fig" captionStartId="1.[114,153,1272,1296]" captionTargetBox="[117,1100,579,1255]" captionTargetId="figure-214@1.[114,1108,570,1264]" captionTargetPageId="1" captionText="Fig. 1.M. (Trachycyrtus) vestitusSchultze, 1925, Holotype, male (MTD)." pageId="2" pageNumber="277">Fig. 1</figureCitation>
) //
<collectingRegion id="49BEF875FFFBFFAC05FF5CF9FEE8FF6C" box="[247,345,208,232]" country="Philippines" name="Romblon" pageId="2" pageNumber="277">Romblon</collectingRegion>
Isl. /
<date id="FFC41057FFFBFFAC04B45CF9FD86FF6C" box="[444,567,208,232]" pageId="2" pageNumber="277" value="1904-06">June 1904</date>
/
<collectionCode id="ED6BAE52FFFBFFAC076D5CF9FDCCFF6C" box="[613,637,208,232]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="277">R</collectionCode>
.C. McGregor // Coll. W. Schultze / Ankauf 1942 // Staatl. Museum fur / Tierkunde Dresden //
<taxonomicName id="4C7A4D14FFFBFFAC043F5D04FE61FEC1" authorityName="HELLER" authorityYear="1912" box="[311,464,301,325]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEA85FFFBFFAC043F5D04FE61FEC1" box="[311,464,301,325]" italics="true" pageId="2" pageNumber="277">Metapocyrtus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C7A4D14FFFBFFAC04EB5D04FD88FEC1" authorityName="Schultze" authorityYear="1925" box="[483,569,301,325]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="277" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="B90EEA85FFFBFFAC04EB5D04FD88FEC1" box="[483,569,301,325]" italics="true" pageId="2" pageNumber="277">vestitus</emphasis>
</taxonomicName>
<specimenCount id="9D7CFD1EFFFBFFAC07335D04FDFEFEC0" box="[571,591,301,324]" count="1" pageId="2" pageNumber="277" type="male"></specimenCount>
/ Det. W. Schultze
<emphasis id="B90EEA85FFFBFFAC045C5D62FE05FEE7" box="[340,436,331,355]" italics="true" pageId="2" pageNumber="277">Schultze</emphasis>
/
<typeStatus id="54C18835FFFBFFAC04D85D62FDB7FEE7" box="[464,518,331,355]" pageId="2" pageNumber="277">Type</typeStatus>
// (all from
<collectionCode id="ED6BAE52FFFBFFAC05B55D43FF4CFE06" box="[189,253,362,386]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="2" pageNumber="277">MTD</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,222 @@
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<mods:title id="DAE101E849945EA4BB9A15390E194B27">To The Knowledge Onmetapocyrtusheller, 1912 (Coleoptera: Curculionidae: Pachyrhynchini) Species From Sibuyan Island, With Description Of New Species</mods:title>
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<treatment id="03D38781FFFAFFA8057A5949FC38FDF3" LSID="urn:lsid:plazi:treatment:03D38781FFFAFFA8057A5949FC38FDF3" httpUri="http://treatment.plazi.org/id/03D38781FFFAFFA8057A5949FC38FDF3" lastPageId="6" lastPageNumber="281" pageId="3" pageNumber="278">
<subSubSection id="C360651CFFFAFFAD057A5949FDB1FAFC" box="[114,512,1376,1400]" pageId="3" pageNumber="278" type="nomenclature">
<paragraph id="8BC53697FFFAFFAD057A5949FDB1FAFC" blockId="3.[114,593,1376,1584]" box="[114,512,1376,1400]" pageId="3" pageNumber="278">
<heading id="D08D81FBFFFAFFAD057A5949FDB1FAFC" bold="true" box="[114,512,1376,1400]" fontSize="10" level="2" pageId="3" pageNumber="278" reason="4">
<emphasis id="B90EEA85FFFAFFAD057A5949FDB1FAFC" bold="true" box="[114,512,1376,1400]" pageId="3" pageNumber="278">
<taxonomicName id="4C7A4D14FFFAFFAD057A5949FE18FAFC" ID-CoL="C3BVC" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[114,425,1376,1400]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="romblonensis" status="sp. nov.">
<emphasis id="B90EEA85FFFAFFAD057A5949FE18FAFC" bold="true" box="[114,425,1376,1400]" italics="true" pageId="3" pageNumber="278">Metapocyrtus romblonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57FEFFFAFFAD04A15949FDB1FAFC" box="[425,512,1376,1400]" pageId="3" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C360651CFFFAFFAA057A5956FBC8FFF2" lastPageId="4" lastPageNumber="279" pageId="3" pageNumber="278" type="materials_examined">
<paragraph id="8BC53697FFFAFFAD057A5956FE0EF9B4" blockId="3.[114,593,1376,1584]" pageId="3" pageNumber="278">
<emphasis id="B90EEA85FFFAFFAD057A5956FEFBFA13" bold="true" box="[114,330,1407,1431]" pageId="3" pageNumber="278">Type material.</emphasis>
<materialsCitation id="3B123CCAFFFAFFAD047C5956FE28FA77" collectingDate="2022-09" country="Philippines" location="Sibuyan" pageId="3" pageNumber="278" specimenCount="1" stateProvince="Romblon" typeStatus="holotype">
<typeStatus id="54C18835FFFAFFAD047C5956FE6BFA13" box="[372,474,1407,1431]" pageId="3" pageNumber="278" type="holotype">Holotype</typeStatus>
, male:
<collectingCountry id="F36D7607FFFAFFAD057259B4FE90FA31" box="[122,289,1437,1461]" name="Philippines" pageId="3" pageNumber="278">PHILIPPINES</collectingCountry>
/
<location id="8EA5604CFFFAFFAD043559B4FE26FA31" LSID="urn:lsid:plazi:treatment:03D38781FFFAFFA8057A5949FC38FDF3:8EA5604CFFFAFFAD043559B4FE26FA31" box="[317,407,1437,1461]" country="Philippines" name="Sibuyan" pageId="3" pageNumber="278" stateProvince="Romblon">Sibuyan</location>
,
<collectingRegion id="49BEF875FFFAFFAD04AF59B4FDBAFA31" box="[423,523,1437,1461]" country="Philippines" name="Romblon" pageId="3" pageNumber="278">Romblon</collectingRegion>
/
<date id="FFC41057FFFAFFAD072059B4FF16FA50" pageId="3" pageNumber="278" value="2022-09">
<collectingDate id="EF80E9BFFFFAFFAD072059B4FF16FA50" pageId="3" pageNumber="278" value="2022-09">IX. 2022</collectingDate>
</date>
/ local collector leg. (white rectangular label, printed) //
</materialsCitation>
<typeStatus id="54C18835FFFAFFAD04AE59F2FD8DFA77" box="[422,572,1499,1523]" pageId="3" pageNumber="278" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C7A4D14FFFAFFAD057A59D3FE98F996" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[114,297,1530,1554]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="romblonensis">
<emphasis id="B90EEA85FFFAFFAD057A59D3FE98F996" box="[114,297,1530,1554]" italics="true" pageId="3" pageNumber="278">M. romblonensis</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023 (red rectangular label, printed).
</paragraph>
<paragraph id="8BC53697FFFAFFAD057A5A8FFCA1FA77" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1103,1376,1523]" pageId="3" pageNumber="278">
<materialsCitation id="3B123CCAFFFAFFAD057A5A8FFCBAFA77" collectionCode="DUBC" pageId="3" pageNumber="278" specimenCount="284" specimenCount-female="5" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D7CFD1EFFFAFFAD057A5A8FFD6DFAFC" box="[114,732,1376,1726]" count="278" pageId="3" pageNumber="278" type="generic" typeStatus="paratype">278 Paratypes</specimenCount>
: (
<specimenCount id="9D7CFD1EFFFAFFAD07FB5949FCF1FAFC" box="[755,832,1376,1400]" count="1" pageId="3" pageNumber="278" type="male">1 male</specimenCount>
,
<specimenCount id="9D7CFD1EFFFAFFAD06455949FC09FAFC" box="[845,952,1376,1400]" count="5" pageId="3" pageNumber="278" type="female">5 females</specimenCount>
) all with the same location label as
<typeStatus id="54C18835FFFAFFAD06975956FC4EFA13" box="[927,1023,1407,1431]" pageId="3" pageNumber="278" type="holotype">holotype</typeStatus>
, with additional red label:
<typeStatus id="54C18835FFFAFFAD067859B4FBB2FA31" box="[880,1027,1437,1461]" pageId="3" pageNumber="278" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C7A4D14FFFAFFAD012459B7FCB7FA51" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="romblonensis">
<emphasis id="B90EEA85FFFAFFAD012459B7FCB7FA51" italics="true" pageId="3" pageNumber="278">M. romblonensis</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023 (all in
<collectionCode id="ED6BAE52FFFAFFAD07BD59F2FCB5FA77" box="[693,772,1499,1523]" pageId="3" pageNumber="278">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC53697FFFDFFAA05D05C4AFBC8FFF2" blockId="4.[216,1145,99,118]" box="[216,1145,99,118]" pageId="4" pageNumber="279">
<emphasis id="B90EEA85FFFDFFAA05D05C4AFBC8FFF2" box="[216,1145,99,118]" italics="true" pageId="4" pageNumber="279">
To the knowledge on metapocyrtus
<bibRefCitation id="EFEB4B66FFFDFFAA070F5C4AFDDFFFF2" author="Heller K. M." box="[519,622,99,118]" pageId="4" pageNumber="279" pagination="295 - 402" refId="ref2341" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science 7 (5): 295 - 402." type="journal article" year="1912">heller, 1912</bibRefCitation>
(coleoptera: curculionidae:
<taxonomicName id="4C7A4D14FFFDFFAA066D5C4AFC5AFFF2" authorityName="Schoenherr" authorityYear="1826" box="[869,1003,99,118]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Pachyrhynchini">pachyrhynchini</taxonomicName>
) species from …
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF05661FFFFDFFAA05B55F1BFC71FCCE" box="[189,960,818,842]" pageId="4" pageNumber="279" startId="4.[189,228,818,842]" targetBox="[189,1174,171,810]" targetPageId="4" targetType="figure">
<paragraph id="8BC53697FFFDFFAA05B55F1BFC71FCCE" blockId="4.[189,960,818,842]" box="[189,960,818,842]" pageId="4" pageNumber="279">
Fig. 4.
<taxonomicName id="4C7A4D14FFFDFFAA04015F1BFD07FCCE" authority="Heller, 1912" authorityName="Heller" authorityYear="1912" box="[265,694,818,842]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="striatus" subGenus="Metapocyrtus">
<emphasis id="B90EEA85FFFDFFAA04015F1BFD9FFCCE" box="[265,558,818,842]" italics="true" pageId="4" pageNumber="279">M. (Metapocyrtus) striatus</emphasis>
<bibRefCitation id="EFEB4B66FFFDFFAA07265F1BFD07FCCE" author="Heller K. M." box="[558,694,818,842]" pageId="4" pageNumber="279" pagination="295 - 402" refId="ref2341" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science 7 (5): 295 - 402." type="journal article" year="1912">Heller, 1912</bibRefCitation>
</taxonomicName>
, Holotype, male (MTD).
</paragraph>
</caption>
<subSubSection id="C360651CFFFDFFA805B55FA4FB25FE3B" lastPageId="6" lastPageNumber="281" pageId="4" pageNumber="279" type="description">
<paragraph id="8BC53697FFFDFFAA05B55FA4FEECFB86" blockId="4.[189,668,909,1026]" pageId="4" pageNumber="279">
<emphasis id="B90EEA85FFFDFFAA05B55FA4FDAFFC21" bold="true" box="[189,542,909,933]" pageId="4" pageNumber="279">
Description.
<typeStatus id="54C18835FFFDFFAA04505FA4FE73FC21" box="[344,450,909,933]" pageId="4" pageNumber="279" type="holotype">Holotype</typeStatus>
, Male.
</emphasis>
(
<figureCitation id="13412A12FFFDFFAA07385FA4FD22FC21" box="[560,659,909,933]" captionStart="Fig" captionStartId="5.[114,153,1127,1151]" captionTargetBox="[127,1057,271,1119]" captionTargetId="figure-243@5.[114,1084,263,1120]" captionTargetPageId="5" captionText="Fig. 5. Dorsal habitus ofM. romblonensis sp. nov., male, holotype (A); B growths at elytral shoulder; C penis in lateral view; D sternite IX in dorsal view; E sternite VIII in dorsal view; F ovipositor; G spermatheca." pageId="4" pageNumber="279">Fig. 5A</figureCitation>
) Dimensions: LB: 8,6, LR: 1,9, WR: 1,3, LP: 3,2, WP: 3,2, LE: 5,4, WE: 3,5. N = 1 for all measurements.
</paragraph>
<paragraph id="8BC53697FFFDFFAA05B5580EFDABFB1F" blockId="4.[189,668,1063,1179]" pageId="4" pageNumber="279">
<emphasis id="B90EEA85FFFDFFAA05B5580EFEF7FBBB" bold="true" box="[189,326,1063,1087]" pageId="4" pageNumber="279">Coloration.</emphasis>
Integument black. Body surface, rostrum, head, and underside moderately shiny, with markings of round, metallic, green to reddish scales.
</paragraph>
<paragraph id="8BC53697FFFDFFAA05B558E9FE42F98F" blockId="4.[189,668,1216,1639]" pageId="4" pageNumber="279">
<emphasis id="B90EEA85FFFDFFAA05B558E9FEB6FB5C" bold="true" box="[189,263,1216,1240]" pageId="4" pageNumber="279">Head.</emphasis>
Dorsal surface and lateral sides without scally markings; surface between eyes with single white setae; genae covered with white to greenish shiny hair-like scales, increasing intensity to ventral surface; forehead flattish; moderate medial longitudinal furrow from base of the eyes to base of the forehead; transverse groove at the base of the eyes confluent with medial furrow; eyes medium-sized, moderately convex from outline of head.
</paragraph>
<paragraph id="8BC53697FFFDFFAA05B55A38FB8FFB1F" blockId="4.[189,668,1216,1639]" lastBlockId="4.[699,1179,909,1179]" pageId="4" pageNumber="279">
<emphasis id="B90EEA85FFFDFFAA05B55A38FE9FF9AD" bold="true" box="[189,302,1553,1577]" pageId="4" pageNumber="279">Rostrum.</emphasis>
Longer than wide (LR/WR: 1,46), moderately rugose on all length; basal half covered with white setae; finely punctured on all length; without colourful scales; dorsum with deep medial furrow from base to anterior ½ forming moderate medial concavity; in dorsal contour nearly straight; in lateral contour convex dorsally, with moderate dorsal bulge; moderate furrow from base of eye to antennal scape; ventral part covered with setae and longer, white hairs; lateral parts without puncture.
</paragraph>
<paragraph id="8BC53697FFFDFFAA07B358E9FC16FA2B" blockId="4.[699,1179,1216,1455]" pageId="4" pageNumber="279">
<emphasis id="B90EEA85FFFDFFAA07B358E9FC9EFB5C" bold="true" box="[699,815,1216,1240]" pageId="4" pageNumber="279">Antennae</emphasis>
with antennal scape nearly same length as funicle, scape reaching beyond hind margin of eye; sparse setae at anterior half; funicular segment 1,5 times longer than II, segment II 1,5 times longer than III, segments III VI as long as wide, segment VII wider than long, club sub-ellipsoidal, 2,5 times as long as wide.
</paragraph>
<paragraph id="8BC53697FFFDFFAA07B359FDFB2AF9E3" blockId="4.[699,1179,1492,1639]" pageId="4" pageNumber="279">
<emphasis id="B90EEA85FFFDFFAA07B359FDFC86FA68" bold="true" box="[699,823,1492,1516]" pageId="4" pageNumber="279">Prothorax</emphasis>
sub globular, as long as wide (LP/WP: 1), strongly punctured with sparse corresponding setae; widest just before the middle, moderately convex in dorsal view; with moderate medial longitudinal groove at
</paragraph>
<paragraph id="8BC53697FFFDFFAB01645A8FFBAFFF6C" blockId="4.[1132,1172,1702,1726]" lastBlockId="5.[624,1103,178,232]" lastPageId="5" lastPageNumber="280" pageId="4" pageNumber="279">279 disc; with the following markings of round, metallic green to goldish, yellowish shales: two roundish dots medially at disc, each redirected laterally; confluent anterior and posterior transverse groove at all length.</paragraph>
<caption id="DF05661FFFFCFFAB057A584EFDA3FB38" pageId="5" pageNumber="280" startId="5.[114,153,1127,1151]" targetBox="[127,1057,271,1119]" targetPageId="5" targetType="figure">
<paragraph id="8BC53697FFFCFFAB057A584EFDA3FB38" blockId="5.[114,1103,1127,1212]" pageId="5" pageNumber="280">
Fig. 5. Dorsal habitus of
<taxonomicName id="4C7A4D14FFFCFFAB047F584EFD9BFBFB" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[375,554,1127,1151]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="280" phylum="Arthropoda" rank="species" species="romblonensis" status="sp. nov.">
<emphasis id="B90EEA85FFFCFFAB047F584EFD9BFBFB" box="[375,554,1127,1151]" italics="true" pageId="5" pageNumber="280">M. romblonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57FEFFFCFFAB0727584EFD34FBFB" box="[559,645,1127,1151]" pageId="5" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
, male, holotype (A); B growths at elytral shoulder; C penis in lateral view; D sternite IX in dorsal view; E sternite VIII in dorsal view; F ovipositor; G spermatheca.
</paragraph>
</caption>
<paragraph id="8BC53697FFFCFFAB057A58C8FDE1F9ED" blockId="5.[114,593,1249,1641]" pageId="5" pageNumber="280">
<emphasis id="B90EEA85FFFCFFAB057A58C8FF70FB7D" bold="true" box="[114,193,1249,1273]" pageId="5" pageNumber="280">Elytra</emphasis>
sub-ellipsoidal, (LE/WE: 1,54), slightly wider than prothorax and moderately longer than prothorax (WE/WP: 1,09; LE/WP: 1,68); striate punctate near suture, irregular puncture towards dorsolateral sides; in dorsal contour widest just at the middle, slightly convex; each elytron with the following scally markings of round, metallic green to goldish, yellow scales: 1) two roundish spots at medial part, one along intervals II to III, one redirected laterally; 2) two roundish spots at apical part along intervals II to III, one along apical ½,
</paragraph>
<paragraph id="8BC53697FFFCFFAB057A5A8FFD5FFA35" blockId="5.[114,154,1702,1726]" lastBlockId="5.[624,1103,1249,1457]" pageId="5" pageNumber="280">280 one redirected to apex; spots can vary in size from smaller to larger, some exemplars have two additional roundish spots at sub-basal part, one along intervals II to III, one redirected laterally; larger growths at each elytral shoulder that vary in number along individuals.</paragraph>
<paragraph id="8BC53697FFFCFFA8077859FFFDE8FEC0" blockId="5.[624,1103,1494,1641]" lastBlockId="6.[189,669,178,324]" lastPageId="6" lastPageNumber="281" pageId="5" pageNumber="280">
<emphasis id="B90EEA85FFFCFFAB077859FFFD61FA6A" bold="true" box="[624,720,1494,1518]" pageId="5" pageNumber="280">Femora</emphasis>
covered with rare white to golden setae and hair-like scales in all length; femora, tibia and tarsus without scally markings; fore-tibia with longer, brownish setae that are more intense along internal margin and towards anterior part and moderate denticules along internal margin; mid and hind tibiae with less dense setae, without denticles along internal margin. Male genitalia as shown in
<figureCitation id="13412A12FFFFFFA804EA5D05FDE4FEC0" box="[482,597,300,324]" captionStart="Fig" captionStartId="5.[114,153,1127,1151]" captionTargetBox="[127,1057,271,1119]" captionTargetId="figure-243@5.[114,1084,263,1120]" captionTargetPageId="5" captionText="Fig. 5. Dorsal habitus ofM. romblonensis sp. nov., male, holotype (A); B growths at elytral shoulder; C penis in lateral view; D sternite IX in dorsal view; E sternite VIII in dorsal view; F ovipositor; G spermatheca." pageId="6" pageNumber="281">Fig. 5C, D</figureCitation>
.
</paragraph>
<paragraph id="8BC53697FFFFFFA805B55D40FE54FD9F" blockId="6.[189,669,361,539]" pageId="6" pageNumber="281">
<emphasis id="B90EEA85FFFFFFA805B55D40FEAFFE05" bold="true" box="[189,286,361,385]" pageId="6" pageNumber="281">Female.</emphasis>
Elytra more ovate; genitalia as shown in
<figureCitation id="13412A12FFFFFFA804345DA1FE34FE24" box="[316,389,392,416]" captionStart="Fig" captionStartId="5.[114,153,1127,1151]" captionTargetBox="[127,1057,271,1119]" captionTargetId="figure-243@5.[114,1084,263,1120]" captionTargetPageId="5" captionText="Fig. 5. Dorsal habitus ofM. romblonensis sp. nov., male, holotype (A); B growths at elytral shoulder; C penis in lateral view; D sternite IX in dorsal view; E sternite VIII in dorsal view; F ovipositor; G spermatheca." pageId="6" pageNumber="281">Fig. 5</figureCitation>
E-G; spots on disc of prothorax redirected posteriorly, with additional transverse line of scales along anterior margin; disc of prothorax without longitudinal medial groove.
</paragraph>
<paragraph id="8BC53697FFFFFFA805B55E69FB25FE3B" blockId="6.[189,668,576,631]" lastBlockId="6.[699,1179,178,447]" pageId="6" pageNumber="281">
<emphasis id="B90EEA85FFFFFFA805B55E69FE01FDDC" bold="true" box="[189,432,576,600]" pageId="6" pageNumber="281">Differential analyses.</emphasis>
<taxonomicName id="4C7A4D14FFFFFFA804B05E69FDDFFDDC" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[440,622,576,600]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="romblonensis" status="sp. nov.">
<emphasis id="B90EEA85FFFFFFA804B05E69FDDFFDDC" box="[440,622,576,600]" italics="true" pageId="6" pageNumber="281">M. romblonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57FEFFFFFFA8077F5E69FF5DFDF3" pageId="6" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
in general appearance is unique for fauna of Sibuyan but can be related to
<taxonomicName id="4C7A4D14FFFFFFA8017F5C9BFBBEFF6C" authority="Heller, 1912" authorityName="Heller" authorityYear="1912" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Metapocyrtus">
<emphasis id="B90EEA85FFFFFFA8017F5C9BFCCAFF6D" italics="true" pageId="6" pageNumber="281">M. (M.) puncticollis</emphasis>
<bibRefCitation id="EFEB4B66FFFFFFA8068A5CF9FBBEFF6C" author="Heller K. M." box="[898,1039,208,232]" pageId="6" pageNumber="281" pagination="295 - 402" refId="ref2341" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science 7 (5): 295 - 402." type="journal article" year="1912">Heller, 1912</bibRefCitation>
</taxonomicName>
by general appearance and scally markings of metallic green colour. The new species can be distinguished by unique growths at each elytral shoulder, dorsally convex rostrum in lateral view that is widest at base (
<taxonomicName id="4C7A4D14FFFFFFA8017F5D43FCE1FE24" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="romblonensis">
<emphasis id="B90EEA85FFFFFFA8017F5D43FCE1FE24" italics="true" pageId="6" pageNumber="281">M. romblonensis</emphasis>
</taxonomicName>
sp. new.) not at sub-apical part (
<taxonomicName id="4C7A4D14FFFFFFA807CC5D8EFB90FE3B" authority="Heller, 1912" authorityName="Heller" authorityYear="1912" box="[708,1057,423,447]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="puncticollis" subGenus="Metapocyrtus">
<emphasis id="B90EEA85FFFFFFA807CC5D8EFC2AFE3B" box="[708,923,423,447]" italics="true" pageId="6" pageNumber="281">M. (M.) puncticollis</emphasis>
<bibRefCitation id="EFEB4B66FFFFFFA806935D8EFB90FE3B" author="Heller K. M." box="[923,1057,423,447]" pageId="6" pageNumber="281" pagination="295 - 402" refId="ref2341" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science 7 (5): 295 - 402." type="journal article" year="1912">Heller, 1912</bibRefCitation>
</taxonomicName>
) (
<figureCitation id="13412A12FFFFFFA8013D5D8EFB36FE3B" box="[1077,1159,423,447]" captionStart="Fig" captionStartId="6.[189,228,1413,1437]" captionTargetBox="[190,1136,662,1384]" captionTargetId="figure-235@6.[189,1152,662,1407]" captionTargetPageId="6" captionText="Fig. 6. Dorsal (A, B) and lateral (C, D) view ofM. (Metapocyrtus) puncticollisHeller, 1912 (A, C) andM. romblonensis sp. nov. (B, D)." pageId="6" pageNumber="281">Fig. 6C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C360651CFFFFFFA807B35DCDFBDAFE78" box="[699,1131,484,508]" pageId="6" pageNumber="281" type="distribution">
<paragraph id="8BC53697FFFFFFA807B35DCDFBDAFE78" blockId="6.[699,1131,484,508]" box="[699,1131,484,508]" pageId="6" pageNumber="281">
<emphasis id="B90EEA85FFFFFFA807B35DCDFCE5FE78" bold="true" box="[699,852,484,508]" pageId="6" pageNumber="281">Distribution.</emphasis>
Sibuyan Island,
<collectingRegion id="49BEF875FFFFFFA801085DCDFBD6FE78" box="[1024,1127,484,508]" country="Philippines" name="Romblon" pageId="6" pageNumber="281">Romblon</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C360651CFFFFFFA807B35E08FC38FDF3" pageId="6" pageNumber="281" type="etymology">
<paragraph id="8BC53697FFFFFFA807B35E08FC38FDF3" blockId="6.[699,1179,545,631]" pageId="6" pageNumber="281">
<emphasis id="B90EEA85FFFFFFA807B35E08FC8CFDBD" bold="true" box="[699,829,545,569]" pageId="6" pageNumber="281">Etymology.</emphasis>
The specific epithet is a toponym to the
<typeStatus id="54C18835FFFFFFA8067F5E69FC14FDDC" box="[887,933,576,600]" pageId="6" pageNumber="281">type</typeStatus>
locality to
<collectingRegion id="49BEF875FFFFFFA801395E69FB2AFDDC" box="[1073,1179,576,600]" country="Philippines" name="Romblon" pageId="6" pageNumber="281">Romblon</collectingRegion>
Island,
<collectingCountry id="F36D7607FFFFFFA806025E76FC37FDF3" box="[778,902,607,631]" name="Philippines" pageId="6" pageNumber="281">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,122 @@
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<mods:title id="9FEBE4D853E4C7F4E92064F7A38A6186">To The Knowledge Onmetapocyrtusheller, 1912 (Coleoptera: Curculionidae: Pachyrhynchini) Species From Sibuyan Island, With Description Of New Species</mods:title>
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<subSubSection id="C360651CFFFBFFAC07B359B2FBC1FA37" box="[699,1136,1435,1459]" pageId="2" pageNumber="277" type="nomenclature">
<paragraph id="8BC53697FFFBFFAC07B359B2FBC1FA37" blockId="2.[699,1136,1435,1489]" box="[699,1136,1435,1459]" pageId="2" pageNumber="277">
<heading id="D08D81FBFFFBFFAC07B359B2FBC1FA37" bold="true" box="[699,1136,1435,1459]" fontSize="10" level="2" pageId="2" pageNumber="277" reason="4">
<taxonomicName id="4C7A4D14FFFBFFAC07B359B2FBC1FA37" ID-CoL="42CH5" authority="Heller, 1912" authorityName="Heller" authorityYear="1912" box="[699,1136,1435,1459]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="277" phylum="Arthropoda" rank="species" species="striatus" subGenus="Metapocyrtus">
<emphasis id="B90EEA85FFFBFFAC07B359B2FBC1FA37" bold="true" box="[699,1136,1435,1459]" pageId="2" pageNumber="277">
<emphasis id="B90EEA85FFFBFFAC07B359B2FC57FA37" bold="true" box="[699,998,1435,1459]" italics="true" pageId="2" pageNumber="277">M. (Metapocyrtus) striatus</emphasis>
<bibRefCitation id="EFEB4B66FFFBFFAC06EE59B2FBC1FA37" author="Heller K. M." box="[998,1136,1435,1459]" pageId="2" pageNumber="277" pagination="295 - 402" refId="ref2341" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science 7 (5): 295 - 402." type="journal article" year="1912">Heller, 1912</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C360651CFFFBFFAD07B35990FDA8FE3B" lastPageId="3" lastPageNumber="278" pageId="2" pageNumber="277" type="description">
<paragraph id="8BC53697FFFBFFAC07B35990FCBAFA55" blockId="2.[699,1136,1435,1489]" box="[699,779,1465,1489]" pageId="2" pageNumber="277">
(
<figureCitation id="13412A12FFFBFFAC07CB5990FCB2FA55" box="[707,771,1465,1489]" captionStart="Fig" captionStartId="4.[189,228,818,842]" captionTargetBox="[189,1174,171,810]" captionTargetId="figure-301@4.[189,1174,171,810]" captionTargetPageId="4" captionText="Fig. 4.M. (Metapocyrtus) striatusHeller, 1912, Holotype, male (MTD)." pageId="2" pageNumber="277">Fig. 4</figureCitation>
)
</paragraph>
<paragraph id="8BC53697FFFBFFAC07B359DEFB2BF9EF" blockId="2.[699,1178,1527,1643]" pageId="2" pageNumber="277">
<emphasis id="B90EEA85FFFBFFAC07B359DEFCF4F98B" bold="true" box="[699,837,1527,1551]" pageId="2" pageNumber="277">Description</emphasis>
(
<bibRefCitation id="EFEB4B66FFFBFFAC065459DEFC46F98B" author="Heller K. M." box="[860,1015,1527,1551]" pageId="2" pageNumber="277" pagination="295 - 402" refId="ref2341" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science 7 (5): 295 - 402." type="journal article" year="1912">Heller, 1912</bibRefCitation>
): Pronotum roughly dotted; with single round to elongated white scales dorsally on disc; anterior and posterior edge of pronotum with
</paragraph>
<paragraph id="8BC53697FFFBFFAD01645A8FFE5AFE83" blockId="2.[1132,1172,1702,1726]" lastBlockId="3.[114,593,178,263]" lastPageId="3" lastPageNumber="278" pageId="2" pageNumber="277">277 sparse white scales; elytra with regular puncture rows and corresponding longitudinal rows of white scales.</paragraph>
<paragraph id="8BC53697FFFAFFAD057A5D05FDA8FE3B" blockId="3.[114,593,300,447]" pageId="3" pageNumber="278">
<emphasis id="B90EEA85FFFAFFAD057A5D05FF50FEC0" bold="true" box="[114,225,300,324]" pageId="3" pageNumber="278">Relation:</emphasis>
Related to
<taxonomicName id="4C7A4D14FFFAFFAD04695D04FEA5FEE7" authority="Schultze, 1925" authorityName="Schultze" authorityYear="1925" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="sibuyanensis" subGenus="Metapocyrtus">
<emphasis id="B90EEA85FFFAFFAD04695D04FDE0FEC1" box="[353,593,301,325]" italics="true" pageId="3" pageNumber="278">M. (M.) sibuyanensis</emphasis>
<bibRefCitation id="EFEB4B66FFFAFFAD057A5D62FEA5FEE7" author="Schultze W." box="[114,276,331,355]" pageId="3" pageNumber="278" pagination="131 - 310" refId="ref2426" refString="Schultze W. 1925. A monograph of the pachyrrhynchid group of the Brachyderinse, Curculionidme: Part III. The genera Apocyrtidius Heller and Metapocyrtus Heller. The Philippine Journal of Science 26 (2): 131 - 310." type="journal article" year="1925">Schultze, 1925</bibRefCitation>
</taxonomicName>
, but can be distinguished by different sculpture of elytra, presence of white longitudinal scale stripes on elytra and shorter rostrum with stronger sculpture.
</paragraph>
</subSubSection>
<subSubSection id="C360651CFFFAFFAD07785C98FCBCFE24" pageId="3" pageNumber="278" type="materials_examined">
<paragraph id="8BC53697FFFAFFAD07785C98FBF8FE06" blockId="3.[624,1103,177,416]" pageId="3" pageNumber="278">
<emphasis id="B90EEA85FFFAFFAD07785C98FCE1FF4D" bold="true" box="[624,848,177,201]" pageId="3" pageNumber="278">Material examined</emphasis>
:
<materialsCitation id="3B123CCAFFFAFFAD06605C9BFCEBFEA1" collectionCode="R" collectorName="Male &amp; Acc No. &amp; Lot &amp; Govt. Lab. Coll" pageId="3" pageNumber="278" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="holotype">
<specimenCount id="9D7CFD1EFFFAFFAD06605C9BFC77FF4E" box="[872,966,178,202]" count="3" pageId="3" pageNumber="278" type="male">3 males</specimenCount>
,
<specimenCount id="9D7CFD1EFFFAFFAD06D35C9BFBFDFF4E" box="[987,1100,178,202]" count="1" pageId="3" pageNumber="278" type="female">1 female</specimenCount>
:
<collectorName id="268F5341FFFAFFAD07785CF9FD16FF6C" box="[624,679,208,232]" pageId="3" pageNumber="278">Male</collectorName>
<typeStatus id="54C18835FFFAFFAD07A75CF9FD6CFF6C" box="[687,733,208,232]" pageId="3" pageNumber="278">type</typeStatus>
(
<figureCitation id="13412A12FFFAFFAD07E65CF9FC80FF6C" box="[750,817,208,232]" captionStart="Fig" captionStartId="4.[189,228,818,842]" captionTargetBox="[189,1174,171,810]" captionTargetId="figure-301@4.[189,1174,171,810]" captionTargetPageId="4" captionText="Fig. 4.M. (Metapocyrtus) striatusHeller, 1912, Holotype, male (MTD)." pageId="3" pageNumber="278">Fig. 4</figureCitation>
) //
<collectorName id="268F5341FFFAFFAD06525CF9FC04FF6C" box="[858,949,208,232]" pageId="3" pageNumber="278">Acc No.</collectorName>
1990. /
<collectorName id="268F5341FFFAFFAD011B5CF9FB89FF6C" box="[1043,1080,208,232]" pageId="3" pageNumber="278">Lot</collectorName>
/
<collectorName id="268F5341FFFAFFAD07785CC6FCBBFE83" box="[624,778,239,263]" pageId="3" pageNumber="278">Govt.Lab.Coll</collectorName>
// 1911 / 7 //
<typeStatus id="54C18835FFFAFFAD06935CC6FC52FE83" box="[923,995,239,263]" pageId="3" pageNumber="278">Typus</typeStatus>
! / striatus // Collected by /
<collectionCode id="ED6BAE52FFFAFFAD06405D24FCEBFEA1" box="[840,858,269,293]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="278">R</collectionCode>
</materialsCitation>
.McGregor. // Staatl. Museum fur / Tierkunde Dresden //
<materialsCitation id="3B123CCAFFFAFFAD01295D05FBF4FE06" collectionCode="MTD" collectorName="C. M. Weber." location="Romblon" pageId="3" pageNumber="278" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Romblon" typeStatus="holotype">
<specimenCount id="9D7CFD1EFFFAFFAD01295D05FD1FFEE7" count="2" pageId="3" pageNumber="278" type="male">two males</specimenCount>
and
<specimenCount id="9D7CFD1EFFFAFFAD07E95D62FCE7FEE7" box="[737,854,331,355]" count="1" pageId="3" pageNumber="278" type="female">one female</specimenCount>
with occurrence label: //
<collectingRegion id="49BEF875FFFAFFAD078C5D43FD59FE06" box="[644,744,362,386]" country="Philippines" name="Romblon" pageId="3" pageNumber="278">Romblon</collectingRegion>
Isl /
<collectorName id="268F5341FFFAFFAD06135D43FC11FE06" box="[795,928,362,386]" pageId="3" pageNumber="278">C.M.Weber.</collectorName>
// (all in
<collectionCode id="ED6BAE52FFFAFFAD06F55D43FB8CFE06" box="[1021,1085,362,386]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="3" pageNumber="278">MTD</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC53697FFFAFFAD07785DA1FCBCFE24" blockId="3.[624,1103,177,416]" box="[624,781,392,416]" pageId="3" pageNumber="278">
(
<figureCitation id="13412A12FFFAFFAD07705DA1FD08FE24" box="[632,697,392,416]" captionStart="Fig" captionStartId="5.[114,153,1127,1151]" captionTargetBox="[127,1057,271,1119]" captionTargetId="figure-243@5.[114,1084,263,1120]" captionTargetPageId="5" captionText="Fig. 5. Dorsal habitus ofM. romblonensis sp. nov., male, holotype (A); B growths at elytral shoulder; C penis in lateral view; D sternite IX in dorsal view; E sternite VIII in dorsal view; F ovipositor; G spermatheca." pageId="3" pageNumber="278">Fig. 5</figureCitation>
,
<figureCitation id="13412A12FFFAFFAD07CC5DA1FCB4FE24" box="[708,773,392,416]" captionStart="Fig" captionStartId="6.[189,228,1413,1437]" captionTargetBox="[190,1136,662,1384]" captionTargetId="figure-235@6.[189,1152,662,1407]" captionTargetPageId="6" captionText="Fig. 6. Dorsal (A, B) and lateral (C, D) view ofM. (Metapocyrtus) puncticollisHeller, 1912 (A, C) andM. romblonensis sp. nov. (B, D)." pageId="3" pageNumber="278">6B, D</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,156 @@
<document id="1950CDA5E3F5685355F3F7DF9E62A613" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Müller, Patrick" docDate="2022" docId="03DAD52BFF80FFCDFD19BF275E00A466" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus temporiscapsula Fanti &amp; Müller, 2022, sp. nov." docType="treatment" docVersion="1" lastPageNumber="370" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="368" updateTime="1722690849856" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="677CBE23EFB32B9EF1A6EAA214504252">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="92A78B21BA0D13D5612155FB76117F63">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="1072BCAB5A91EFB3E9A669404812E607">Müller, Patrick</mods:namePart>
</mods:name>
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<mods:title id="A506AEED59E4D44EFAC662AD0195BE34">Baltic Journal of Coleopterology</mods:title>
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<mods:part id="BF1F15212165A66EEA1F7F37DC12267A">
<mods:date id="2151DF73050C9D5375981E263F592382">2022</mods:date>
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<mods:number id="FF6DE634C1E608C8B048DE61846A5923">22</mods:number>
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</mods:mods>
<treatment id="03DAD52BFF80FFCDFD19BF275E00A466" LSID="urn:lsid:plazi:treatment:03DAD52BFF80FFCDFD19BF275E00A466" httpUri="http://treatment.plazi.org/id/03DAD52BFF80FFCDFD19BF275E00A466" lastPageId="39" lastPageNumber="370" pageId="37" pageNumber="368">
<subSubSection id="C36937B6FF80FFCFFD19BF275B6DA35D" box="[643,1067,533,557]" pageId="37" pageNumber="368" type="nomenclature">
<paragraph id="8BCC643DFF80FFCFFD19BF275B6DA35D" blockId="37.[643,1067,533,588]" box="[643,1067,533,557]" pageId="37" pageNumber="368">
<heading id="D084D351FF80FFCFFD19BF275B6DA35D" bold="true" box="[643,1067,533,557]" fontSize="10" level="2" pageId="37" pageNumber="368" reason="4">
<emphasis id="B907B82FFF80FFCFFD19BF275B6DA35D" bold="true" box="[643,1067,533,557]" pageId="37" pageNumber="368">
<taxonomicName id="4C731FBEFF80FFCFFD19BF275C89A35D" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[643,975,533,557]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="368" phylum="Arthropoda" rank="species" species="temporiscapsula" status="sp. nov.">
<emphasis id="B907B82FFF80FFCFFD19BF275C89A35D" bold="true" box="[643,975,533,557]" italics="true" pageId="37" pageNumber="368">Sanaungulus temporiscapsula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF80FFCFFC4CBF275B6DA35D" box="[982,1067,533,557]" pageId="37" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF80FFCFFD19BF075DA6A33C" box="[643,736,565,588]" pageId="37" pageNumber="368" type="description">
<paragraph id="8BCC643DFF80FFCFFD19BF075DA6A33C" blockId="37.[643,1067,533,588]" box="[643,736,565,588]" pageId="37" pageNumber="368">
(
<figureCitation id="134878B8FF80FFCFFD11BF075D91A33C" box="[651,727,565,588]" captionStart="Fig" captionStartId="38.[151,190,1635,1658]" captionTargetBox="[151,1172,380,1605]" captionTargetId="figure-38@38.[151,1172,380,1605]" captionTargetPageId="38" captionText="Fig. 21. Sanaungulus temporiscapsula sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view." pageId="37" pageNumber="368">Fig. 21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF80FFCFFD19BF5E5CE4A2C0" pageId="37" pageNumber="368" type="materials_examined">
<paragraph id="8BCC643DFF80FFCFFD19BF5E5B69A273" blockId="37.[643,1134,620,771]" pageId="37" pageNumber="368">
<materialsCitation id="3B1B6E60FF80FFCFFD19BF5E5B69A273" collectingDate="2021-12-16" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="37" pageNumber="368" specimenCode="BUB4314" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFF80FFCFFD19BF5E5DA9A3F4" bold="true" box="[643,751,620,644]" pageId="37" pageNumber="368">
<typeStatus id="54C8DA9FFF80FFCFFD19BF5E5DACA3F4" box="[643,746,620,644]" pageId="37" pageNumber="368" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FF80FFCFFD69BF5E5C04A3F3" box="[755,834,620,643]" pageId="37" pageNumber="368">Female</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FF80FFCFFB8BBF5E5B28A3F3" box="[1041,1134,620,643]" pageId="37" pageNumber="368">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFF80FFCFFD10BFBE5D9CA3D3" box="[650,730,652,675]" country="Myanmar" name="Kachin" pageId="37" pageNumber="368">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FF80FFCFFCE9BFBE5C80A3D3" LSID="urn:lsid:plazi:treatment:03DAD52BFF80FFCDFD19BF275E00A466:8EAC32E6FF80FFCFFCE9BFBE5C80A3D3" box="[883,966,652,675]" county="Female" municipality="Burmese" name="Catalog" pageId="37" pageNumber="368" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFF80FFCFFD54BF9E5C10A3B3" box="[718,854,684,707]" pageId="37" pageNumber="368" value="2021-12-16">
<collectingDate id="EF89BB15FF80FFCFFD54BF9E5C10A3B3" box="[718,854,684,707]" pageId="37" pageNumber="368" value="2021-12-16">
2021 XII
<quantity id="4C8BC9D8FF80FFCFFCA1BF9E5C10A3B3" box="[827,854,684,707]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="37" pageNumber="368" unit="in" value="16.0">16</quantity>
</collectingDate>
</date>
in the
<location id="8EAC32E6FF80FFCFFC3CBF9E5DB4A393" LSID="urn:lsid:plazi:treatment:03DAD52BFF80FFCDFD19BF275E00A466:8EAC32E6FF80FFCFFC3CBF9E5DB4A393" county="Female" municipality="Burmese" name="Bayerische Staatssammlung" pageId="37" pageNumber="368" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FF80FFCFFC87BFFE5CF6A393" LSID="urn:lsid:plazi:treatment:03DAD52BFF80FFCDFD19BF275E00A466:8EAC32E6FF80FFCFFC87BFFE5CF6A393" box="[797,944,716,739]" county="Female" municipality="Burmese" name="Palaontologie" pageId="37" pageNumber="368" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FF80FFCFFC7EBFFE5B00A393" LSID="urn:lsid:plazi:treatment:03DAD52BFF80FFCDFD19BF275E00A466:8EAC32E6FF80FFCFFC7EBFFE5B00A393" box="[996,1094,716,739]" county="Female" municipality="Burmese" name="Geologie" pageId="37" pageNumber="368" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FF80FFCFFD19BFDE5D99A273" LSID="urn:lsid:plazi:treatment:03DAD52BFF80FFCDFD19BF275E00A466:8EAC32E6FF80FFCFFD19BFDE5D99A273" box="[643,735,748,771]" county="Female" municipality="Burmese" name="Muller's" pageId="37" pageNumber="368" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FF80FFCFFC29BFDE5B62A273" box="[947,1060,748,771]" collectionCode="BUB" pageId="37" pageNumber="368">BUB4314</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFF80FFCFFD19BE105C93A22A" blockId="37.[643,1134,802,858]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19BE105C5AA24A" bold="true" box="[643,796,802,826]" pageId="37" pageNumber="368">Type locality.</emphasis>
<collectingCountry id="F36424ADFF80FFCFFCBEBE115CD6A24A" box="[804,912,803,826]" name="Myanmar" pageId="37" pageNumber="368">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFF80FFCFFC01BE115B63A24A" box="[923,1061,803,826]" country="Myanmar" name="Kachin" pageId="37" pageNumber="368">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFF80FFCFFD19BE4B5CE4A2C0" blockId="37.[643,1134,889,944]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19BE4B5C50A2E1" bold="true" box="[643,790,889,913]" pageId="37" pageNumber="368">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF80FFCFFD19BEE25C9AA557" pageId="37" pageNumber="368" type="diagnosis">
<paragraph id="8BCC643DFF80FFCFFD19BEE25C9AA557" blockId="37.[643,1134,976,1063]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19BEE25CC6A298" bold="true" box="[643,896,976,1000]" pageId="37" pageNumber="368">Differential diagnosis.</emphasis>
The species is easily distinguishable by its antennomeres IIIV that are short and subequal in length.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF80FFCDFD19B9745D57A222" lastPageId="39" lastPageNumber="370" pageId="37" pageNumber="368" type="description">
<paragraph id="8BCC643DFF80FFCFFD19B9745D88A5CE" blockId="37.[643,1134,1094,1214]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19B9745C48A52E" bold="true" box="[643,782,1094,1118]" pageId="37" pageNumber="368">Description.</emphasis>
Adult, winged, slender. Female, defined on the shape of the last urites and short antennae. Body length
<quantity id="4C8BC9D8FF80FFCFFCE4B9B55C93A5EE" box="[894,981,1159,1182]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="37" pageNumber="368" unit="mm" value="3.9">3.9 mm</quantity>
. Entirely dark brown.
</paragraph>
<paragraph id="8BCC643DFF80FFCDFD19B9EF5D57A222" blockId="37.[643,1134,1245,1684]" lastBlockId="39.[113,605,155,850]" lastPageId="39" lastPageNumber="370" pageId="37" pageNumber="368">Head transverse, strongly constricted (triangular­shaped) behind the eyes, fitted with rugosity and punctuation. Eyes very prominent, very large, roundish, inserted in the upper and lateral part of the head, inter­ocular dorsal distance approximately 1.8 times greater than the eye diameter. Mandibles falciform. Maxillary palps 4­ segmented with the last palpomere elongated and securiform.Antennae 11­ segmented, filiform, rather robust, short, slightly surpassing the humeral zone of the elytra and not reaching the half of elytra, antennal insertion in the eyes proximity; scape elongated, slender, filiform and not enlarged apically; antennomeres IIIV very short, almost globular, subequal in length with the fourth antennomere slightly serrate apically and slightly longer; antennomere V slightly longer than previous one, antennomeres VIVIII about 1.5 times longer than antennomere IV; antennomeres IXX shorter than previous ones, and slightly stouter; antennomere XI filiform, squared at apex; all the antennomeres with short pubescence. Pronotum transverse, as wide as head, slightly narrow anteriorly, anterior margin rounded and bordered, posterior margin almost straight, sides slightly restricted anteriorly and expanded posteriorly, posterior corners obtuse, pronotum disc convex and slightly wrinkled. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, at humeri wider than pronotum, dehiscent posteriorly, rounded apexes, surface with impressed punctation gathered in striae, pubescent. Metathoracic wings transparent, longer than elytra. Sternum elongated, sternites wide and transverse, last tergite small with two short lobes, last sternite small and rounded. Legs thin, posterior legs long, pubescent; coxae massive; trochanters rounded; femora nearly straight, more robust than tibiae; tibiae slightly flattened, thin, enlarged apically, without spurs near the apex, pro­ and mesotibiae strongly shorter than pro­ and mesofemora, metatibiae very longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 2.0 times longer than second; third tarsomere short and enlarged apically; fourth strongly bilobed; fifth tarsomere very elongated, thin; claws simple, apparently without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FF83FFCCFF0DBB515ED6A7EA" pageId="38" pageNumber="369" startId="38.[151,190,1635,1658]" targetBox="[151,1172,380,1605]" targetPageId="38" targetType="figure">
<paragraph id="8BCC643DFF83FFCCFF0DBB515ED6A7EA" blockId="38.[151,1172,1635,1690]" pageId="38" pageNumber="369">
Fig. 21.
<taxonomicName id="4C731FBEFF83FFCCFF6ABB515D77A70A" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[240,561,1635,1658]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="369" phylum="Arthropoda" rank="species" species="temporiscapsula" status="sp. nov.">
<emphasis id="B907B82FFF83FFCCFF6ABB515D77A70A" box="[240,561,1635,1658]" italics="true" pageId="38" pageNumber="369">Sanaungulus temporiscapsula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF83FFCCFDA2BB515DCCA70A" box="[568,650,1635,1658]" pageId="38" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FF82FFCDFFEBBE435EBFA50F" pageId="39" pageNumber="370" type="etymology">
<paragraph id="8BCC643DFF82FFCDFFEBBE435E1DA538" blockId="39.[113,605,881,1096]" pageId="39" pageNumber="370">
<emphasis id="B907B82FFF82FFCDFFEBBE435FBCA2F9" bold="true" box="[113,250,881,905]" pageId="39" pageNumber="370">Etymology.</emphasis>
Derived from the Latin noun “
<emphasis id="B907B82FFF82FFCDFFE7BEA05F9CA2D9" box="[125,218,914,937]" italics="true" pageId="39" pageNumber="370">temporis</emphasis>
” (
<emphasis id="B907B82FFF82FFCDFF6EBEA05E79A2D9" box="[244,319,914,937]" italics="true" pageId="39" pageNumber="370">tempus</emphasis>
) = time, and the Latin noun “
<emphasis id="B907B82FFF82FFCDFFE7BE805F97A2B9" box="[125,209,946,969]" italics="true" pageId="39" pageNumber="370">capsula</emphasis>
” = capsule. Thus = time capsule. In reference to the fact that the amber piece has embedded the new species making it reach us through time. Specific epithet is to be treated as noun in apposition.
</paragraph>
<paragraph id="8BCC643DFF82FFCDFFEBB95A5EBFA50F" blockId="39.[113,505,1128,1152]" box="[113,505,1128,1152]" pageId="39" pageNumber="370">
<emphasis id="B907B82FFF82FFCDFFEBB95A5E52A5F0" bold="true" box="[113,276,1128,1152]" pageId="39" pageNumber="370">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF82FFCDFFEBB9AC5E00A466" pageId="39" pageNumber="370" type="discussion">
<paragraph id="8BCC643DFF82FFCDFFEBB9AC5E00A466" blockId="39.[113,605,1182,1302]" pageId="39" pageNumber="370">
<emphasis id="B907B82FFF82FFCDFFEBB9AC5F9BA5C6" bold="true" box="[113,221,1182,1206]" pageId="39" pageNumber="370">Remarks.</emphasis>
Piece of clear, golden amber that measures 14x7x
<quantity id="4C8BC9D8FF82FFCDFF75B98D5E02A5A6" box="[239,324,1215,1238]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="39" pageNumber="370" unit="mm" value="2.5">2.5 mm</quantity>
. The inclusion is complete except for the left antenna broken and almost completely missing.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="0E8F786F319F3E077A7A69E59E3E8477" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Müller, Patrick" docDate="2022" docId="03DAD52BFF82FFC3FFEBB87E5F9CA38A" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus ypogaeum Fanti &amp; Müller, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="372" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="370" updateTime="1722691394301" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="798E5520DCA7E02A5E8A8FB4AE2E095B">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="B9A01AE7BA7162CDC698F15AA99C5DB1">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="7C878E7D2F81F5B48448F9603BAE0039">Müller, Patrick</mods:namePart>
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<mods:date id="911376E9BDB6F07880D06B0B8C3CE7E8">2022</mods:date>
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<subSubSection id="C36937B6FF82FFCDFFEBB87E5E91A414" box="[113,471,1356,1380]" pageId="39" pageNumber="370" type="nomenclature">
<paragraph id="8BCC643DFF82FFCDFFEBB87E5E91A414" blockId="39.[113,471,1356,1411]" box="[113,471,1356,1380]" pageId="39" pageNumber="370">
<heading id="D084D351FF82FFCDFFEBB87E5E91A414" bold="true" box="[113,471,1356,1380]" fontSize="10" level="2" pageId="39" pageNumber="370" reason="4">
<emphasis id="B907B82FFF82FFCDFFEBB87E5E91A414" bold="true" box="[113,471,1356,1380]" pageId="39" pageNumber="370">
<taxonomicName id="4C731FBEFF82FFCDFFEBB87E5E3AA414" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[113,380,1356,1380]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="370" phylum="Arthropoda" rank="species" species="ypogaeum" status="sp. nov.">
<emphasis id="B907B82FFF82FFCDFFEBB87E5E3AA414" bold="true" box="[113,380,1356,1380]" italics="true" pageId="39" pageNumber="370">Sanaungulus ypogaeum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF82FFCDFE18B87E5E91A414" box="[386,471,1356,1380]" pageId="39" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF82FFCDFFEBB85E5F89A4F3" box="[113,207,1388,1411]" pageId="39" pageNumber="370" type="description">
<paragraph id="8BCC643DFF82FFCDFFEBB85E5F89A4F3" blockId="39.[113,471,1356,1411]" box="[113,207,1388,1411]" pageId="39" pageNumber="370">
(
<figureCitation id="134878B8FF82FFCDFFE3B85E5F81A4F3" box="[121,199,1388,1411]" captionStart="Fig" captionStartId="40.[151,189,1639,1662]" captionTargetBox="[151,1172,385,1610]" captionTargetId="figure-18@40.[151,1172,385,1610]" captionTargetPageId="40" captionText="Fig. 22. Sanaungulus ypogaeum sp. nov. in Burmese amber.A: Holotype, dorsal view; B: Holotype, ventral view." pageId="39" pageNumber="370">Fig. 22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF82FFCDFFEBB8905CE4A1A2" pageId="39" pageNumber="370" type="materials_examined">
<paragraph id="8BCC643DFF82FFCDFFEBB8905D58A74A" blockId="39.[113,605,1442,1594]" pageId="39" pageNumber="370">
<materialsCitation id="3B1B6E60FF82FFCDFFEBB8905D58A74A" collectingDate="2021-12" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="39" pageNumber="370" specimenCode="BUB4165" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFF82FFCDFFEBB8905F9BA4CA" bold="true" box="[113,221,1442,1466]" pageId="39" pageNumber="370">
<typeStatus id="54C8DA9FFF82FFCDFFEBB8905F9EA4CA" box="[113,216,1442,1466]" pageId="39" pageNumber="370" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FF82FFCDFF7BB8915E76A4CA" box="[225,304,1443,1466]" pageId="39" pageNumber="370">Female</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FF82FFCDFD9AB8915D1BA4CA" box="[512,605,1443,1466]" pageId="39" pageNumber="370">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFF82FFCDFFE3B8F15F8FA4AA" box="[121,201,1475,1498]" country="Myanmar" name="Kachin" pageId="39" pageNumber="370">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FF82FFCDFEF8B8F15EF3A4AA" LSID="urn:lsid:plazi:treatment:03DAD52BFF82FFC3FFEBB87E5F9CA38A:8EAC32E6FF82FFCDFEF8B8F15EF3A4AA" box="[354,437,1475,1498]" county="Female" municipality="Burmese" name="Catalog" pageId="39" pageNumber="370" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFF82FFCDFF27B8D15E67A48A" box="[189,289,1507,1530]" pageId="39" pageNumber="370" value="2021-12">
<collectingDate id="EF89BB15FF82FFCDFF27B8D15E67A48A" box="[189,289,1507,1530]" pageId="39" pageNumber="370" value="2021-12">2021 XII</collectingDate>
</date>
<quantity id="4C8BC9D8FF82FFCDFEB0B8D15E24A48A" box="[298,354,1507,1530]" metricMagnitude="0" metricUnit="m" metricValue="1.7018" pageId="39" pageNumber="370" unit="in" value="67.0">67 in</quantity>
the
<location id="8EAC32E6FF82FFCDFE0FB8D15FA6A76A" LSID="urn:lsid:plazi:treatment:03DAD52BFF82FFC3FFEBB87E5F9CA38A:8EAC32E6FF82FFCDFE0FB8D15FA6A76A" county="Female" municipality="Burmese" name="Bayerische Staatssammlung" pageId="39" pageNumber="370" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FF82FFCDFE96BB315ED9A76A" LSID="urn:lsid:plazi:treatment:03DAD52BFF82FFC3FFEBB87E5F9CA38A:8EAC32E6FF82FFCDFE96BB315ED9A76A" box="[268,415,1539,1562]" county="Female" municipality="Burmese" name="Palaontologie" pageId="39" pageNumber="370" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FF82FFCDFE49BB315D73A76A" LSID="urn:lsid:plazi:treatment:03DAD52BFF82FFC3FFEBB87E5F9CA38A:8EAC32E6FF82FFCDFE49BB315D73A76A" box="[467,565,1539,1562]" county="Female" municipality="Burmese" name="Geologie" pageId="39" pageNumber="370" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FF82FFCDFFEBBB115F8BA74A" LSID="urn:lsid:plazi:treatment:03DAD52BFF82FFC3FFEBB87E5F9CA38A:8EAC32E6FF82FFCDFFEBBB115F8BA74A" box="[113,205,1571,1594]" county="Female" municipality="Burmese" name="Muller's" pageId="39" pageNumber="370" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FF82FFCDFE38BB115D54A74A" box="[418,530,1571,1594]" collectionCode="BUB" pageId="39" pageNumber="370">BUB4165</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFF82FFCDFFEBBB6B5E82A7E0" blockId="39.[113,605,1625,1680]" pageId="39" pageNumber="370">
<emphasis id="B907B82FFF82FFCDFFEBBB6B5E4CA701" bold="true" box="[113,266,1625,1649]" pageId="39" pageNumber="370">Type locality.</emphasis>
<collectingCountry id="F36424ADFF82FFCDFE89BB6B5E39A700" box="[275,383,1625,1648]" name="Myanmar" pageId="39" pageNumber="370">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFF82FFCDFE10BB6B5D52A700" box="[394,532,1625,1648]" country="Myanmar" name="Kachin" pageId="39" pageNumber="370">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFF82FFCDFD19BDA85CE4A1A2" blockId="39.[643,1134,154,210]" pageId="39" pageNumber="370">
<emphasis id="B907B82FFF82FFCDFD19BDA85C50A1C2" bold="true" box="[643,790,154,178]" pageId="39" pageNumber="370">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF82FFCDFD19BDC35CD0A358" pageId="39" pageNumber="370" type="diagnosis">
<paragraph id="8BCC643DFF82FFCDFD19BDC35CD0A358" blockId="39.[643,1135,241,552]" pageId="39" pageNumber="370">
<emphasis id="B907B82FFF82FFCDFD19BDC35C35A079" bold="true" box="[643,883,241,265]" pageId="39" pageNumber="370">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFF82FFCDFCE2BDC05B28A079" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[888,1134,242,265]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="370" phylum="Arthropoda" rank="species" species="ypogaeum" status="sp. nov.">
<emphasis id="B907B82FFF82FFCDFCE2BDC05B28A079" box="[888,1134,242,265]" italics="true" pageId="39" pageNumber="370">Sanaungulus ypogaeum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF82FFCDFD19BC235D90A058" box="[643,726,273,296]" pageId="39" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
differs from other species of
<taxonomicName id="4C731FBEFF82FFCDFB8FBC205DF9A039" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF82FFCDFB8FBC205DF9A039" italics="true" pageId="39" pageNumber="370">Sanaungulus</emphasis>
</taxonomicName>
with filiform antennae:
<taxonomicName id="4C731FBEFF82FFCDFC45BC005C9DA018" authority="(Hsiao &amp; Huang, 2018)" baseAuthorityName="Hsiao &amp; Huang" baseAuthorityYear="2018" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="370" phylum="Arthropoda" rank="species" species="ruicheni">
<emphasis id="B907B82FFF82FFCDFC45BC005D9AA019" italics="true" pageId="39" pageNumber="370">Sanaungulus ruicheni</emphasis>
(
<bibRefCitation id="EFE219CCFF82FFCDFD72BC635C95A018" author="Hsiao Y. &amp; Huang C. - L." box="[744,979,337,360]" pageId="39" pageNumber="370" pagination="257 - 263" refId="ref20597" refString="Hsiao Y., Huang C. - L. 2018. Taxonomic revision on the genus Ornatomalthinus Poinar and Fanti (Coleoptera: Cantharidae), with description of a new species from the Cretaceous Burmese amber. Cretaceous Research 92: 257 - 263." type="journal article" year="2018">Hsiao &amp; Huang, 2018</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C731FBEFF82FFCDFC7EBC605C9BA0F8" authority="Fanti &amp; Damgaard, 2020" authorityName="Fanti &amp; Damgaard" authorityYear="2020" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="370" phylum="Arthropoda" rank="species" species="morellii">
<emphasis id="B907B82FFF82FFCDFC7EBC605D93A0F9" italics="true" pageId="39" pageNumber="370">Sanaungulus morellii</emphasis>
<bibRefCitation id="EFE219CCFF82FFCDFD40BC435C9BA0F8" author="Fanti F. &amp; Damgaard A. L." box="[730,989,369,392]" pageId="39" pageNumber="370" pagination="125 - 139" refId="ref20405" refString="Fanti F., Damgaard A. L. 2020. Fossil soldier beetles of the Anders Damgaard amber collection. Fourth update. Baltic Journal of Coleopterology 20 (2): 125 - 139." type="journal article" year="2020">Fanti &amp; Damgaard, 2020</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C731FBEFF82FFCDFC7CBC405B50A0D8" authority="Fanti &amp; Damgaard, 2020" authorityName="Fanti &amp; Damgaard" authorityYear="2020" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="370" phylum="Arthropoda" rank="species" species="rosenzweigi">
<emphasis id="B907B82FFF82FFCDFC7CBC405C44A0D9" italics="true" pageId="39" pageNumber="370">Sanaungulus rosenzweigi</emphasis>
<bibRefCitation id="EFE219CCFF82FFCDFC92BCA35B50A0D8" author="Fanti F. &amp; Damgaard A. L." box="[776,1046,401,424]" pageId="39" pageNumber="370" pagination="125 - 139" refId="ref20405" refString="Fanti F., Damgaard A. L. 2020. Fossil soldier beetles of the Anders Damgaard amber collection. Fourth update. Baltic Journal of Coleopterology 20 (2): 125 - 139." type="journal article" year="2020">Fanti &amp; Damgaard, 2020</bibRefCitation>
</taxonomicName>
, and the species described here based on its pronotum that is strongly narrow anteriorly and with a rounded anterior margin (
<bibRefCitation id="EFE219CCFF82FFCDFC10BCC35B2CA378" author="Hsiao Y. &amp; Huang C. - L." box="[906,1130,497,520]" pageId="39" pageNumber="370" pagination="257 - 263" refId="ref20597" refString="Hsiao Y., Huang C. - L. 2018. Taxonomic revision on the genus Ornatomalthinus Poinar and Fanti (Coleoptera: Cantharidae), with description of a new species from the Cretaceous Burmese amber. Cretaceous Research 92: 257 - 263." type="journal article" year="2018">Hsiao &amp; Huang 2018</bibRefCitation>
;
<bibRefCitation id="EFE219CCFF82FFCDFD19BF235CCCA358" author="Fanti F. &amp; Damgaard A. L." box="[643,906,529,552]" pageId="39" pageNumber="370" pagination="125 - 139" refId="ref20405" refString="Fanti F., Damgaard A. L. 2020. Fossil soldier beetles of the Anders Damgaard amber collection. Fourth update. Baltic Journal of Coleopterology 20 (2): 125 - 139." type="journal article" year="2020">Fanti &amp; Damgaard 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF82FFC3FD19BF7A5E02A0E4" lastPageId="41" lastPageNumber="372" pageId="39" pageNumber="370" type="description">
<paragraph id="8BCC643DFF82FFCDFD19BF7A5C29A3AF" blockId="39.[643,1134,584,735]" pageId="39" pageNumber="370">
<emphasis id="B907B82FFF82FFCDFD19BF7A5C48A310" bold="true" box="[643,782,584,608]" pageId="39" pageNumber="370">Description.</emphasis>
Adult, winged, slender. Female, defined on the shape of the last urites and on the basis of the short antennae. Body length
<quantity id="4C8BC9D8FF82FFCDFD19BF9A5D91A3CF" box="[643,727,680,703]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="39" pageNumber="370" unit="mm" value="4.9">4.9 mm</quantity>
. Head and pronotum testaceous­brown, elytra and legs brown.
</paragraph>
<paragraph id="8BCC643DFF82FFC3FD19BFCD5E02A0E4" blockId="39.[643,1135,767,1686]" lastBlockId="41.[113,605,156,404]" lastPageId="41" lastPageNumber="372" pageId="39" pageNumber="370">Head elongated, very narrow, strongly constricted (triangular­shaped) behind the eyes, fitted with rugosity and impressed punctuation. Eyes prominent, very large, roundish, inserted in the upper and dorsolateral part of the head, inter­ocular dorsal distance about 0.6 times greater than the eye diameter. Mandibles elongated, robust, falciform, without teeth. Maxillary palps 4­ segmented with the last palpomere strongly securiform. Labial palps 3­ segmented, with the last palpomere slightly securiform.Antennae 11­ segmented, filiform, robust, short and slightly surpassing the humeral zone of the elytra, antennal insertion in the eyes proximity; scape elongated, robust, club­shaped; antennomere II very short, globular; antennomere III short, thinner and longer than second; antennomere IV elongated, the longest; antennomeres VVII about 0.2 times shorter than antennomere IV; antennomeres VIIIX shorter than previous ones; antennomere XI filiform, longer than previous ones, rounded at apex; all the antennomeres with short pubescence. Pronotum transverse, wider than head, narrow anteriorly, anterior margin rounded and very slightly bordered, posterior margin almost straight, sides rounded, posterior corners obtuse, pronotum disc smooth. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, at humeri wider than pronotum and as wide as pronotum at the apex, humeral zone enlarged, posteriorly parallel­sided, rounded apexes, surface sligthly pubescent and with a very small rugosity. Metathoracic wings almost completely covered by elytra. Sternum elongated with posterior margin rounded, sternites narrow and transverse, last tergite deeply incised apically and divided in two lobes, last sternite small and rounded. Legs thin, relatively long, pubescent; coxae massive; trochanters elongated, with rounded apex; femora nearly straight, more robust than tibiae, cylindrical; tibiae flattened, thin, without spurs near the apex, protibiae slightly longer than profemora, metatibiae very longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 2.0 times longer than second; third tarsomere short and bilobed; fourth strongly bilobed; fifth tarsomere elongated, thin, slightly flat and curved; claws simple, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FF8DFFC2FF0DBB555E59A7EE" pageId="40" pageNumber="371" startId="40.[151,189,1639,1662]" targetBox="[151,1172,385,1610]" targetPageId="40" targetType="figure">
<paragraph id="8BCC643DFF8DFFC2FF0DBB555E59A7EE" blockId="40.[151,1172,1639,1694]" pageId="40" pageNumber="371">
Fig. 22.
<taxonomicName id="4C731FBEFF8DFFC2FF71BB5A5EA5A70F" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[235,483,1640,1663]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="371" phylum="Arthropoda" rank="species" species="ypogaeum" status="sp. nov.">
<emphasis id="B907B82FFF8DFFC2FF71BB5A5EA5A70F" box="[235,483,1640,1663]" italics="true" pageId="40" pageNumber="371">Sanaungulus ypogaeum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF8DFFC2FE7DBB555D70A70E" box="[487,566,1639,1662]" pageId="40" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber.A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FF8CFFC3FFEBBC815EBFA3F3" pageId="41" pageNumber="372" type="etymology">
<paragraph id="8BCC643DFF8CFFC3FFEBBC815D0BA33C" blockId="41.[113,605,435,588]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FFEBBC815FBCA0BB" bold="true" box="[113,250,435,459]" pageId="41" pageNumber="372">Etymology.</emphasis>
Derived from the Latin noun
<taxonomicName id="4C731FBEFF8CFFC3FFEBBCE65FAAA09B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[113,236,468,491]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="species" species="ypogaeum">
<emphasis id="B907B82FFF8CFFC3FFE7BCE65FAAA09B" box="[125,236,468,491]" italics="true" pageId="41" pageNumber="372">ypogaeum</emphasis>
</taxonomicName>
(
<emphasis id="B907B82FFF8CFFC3FF67BCE65E79A09B" box="[253,319,468,491]" italics="true" pageId="41" pageNumber="372">hypog</emphasis>
Á
<emphasis id="B907B82FFF8CFFC3FED6BCE65E2AA09B" box="[332,364,468,491]" italics="true" pageId="41" pageNumber="372">um</emphasis>
)” = hypogeum, tomb. In reference to the fact that the amber that has incorporated the specimen is its tomb. Specific epithet is to be treated as noun in apposition.
</paragraph>
<paragraph id="8BCC643DFF8CFFC3FFEBBF595EBFA3F3" blockId="41.[113,505,619,643]" box="[113,505,619,643]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FFEBBF595E52A3F3" bold="true" box="[113,276,619,643]" pageId="41" pageNumber="372">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8CFFC3FFEBBF905F9CA38A" pageId="41" pageNumber="372" type="discussion">
<paragraph id="8BCC643DFF8CFFC3FFEBBF905F9CA38A" blockId="41.[113,605,674,762]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FFEBBF905FA6A3CA" bold="true" box="[113,224,674,698]" pageId="41" pageNumber="372">Remarks.</emphasis>
Piece of clear, golden amber in Cabochon cut 18x14x
<quantity id="4C8BC9D8FF8CFFC3FEDEBFF15ED8A3AA" box="[324,414,707,730]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="41" pageNumber="372" unit="mm" value="2.8">2.8 mm</quantity>
. The inclusion is complete.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,169 @@
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<mods:title id="EAFDE2A1B5AD41E4FA80FE9BA689CCD4">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="8F8A3EEF4B9EE293F96684067730FE1A">Fanti, Fabrizio</mods:namePart>
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<treatment id="03DAD52BFF85FFC8FF0DBE635B00A1F0" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0" httpUri="http://treatment.plazi.org/id/03DAD52BFF85FFC8FF0DBE635B00A1F0" lastPageId="34" lastPageNumber="365" pageId="32" pageNumber="363">
<subSubSection id="C36937B6FF85FFCAFF0DBE635EFDA219" box="[151,443,849,873]" pageId="32" pageNumber="363" type="nomenclature">
<paragraph id="8BCC643DFF85FFCAFF0DBE635EFDA219" blockId="32.[151,443,849,904]" box="[151,443,849,873]" pageId="32" pageNumber="363">
<heading id="D084D351FF85FFCAFF0DBE635EFDA219" bold="true" box="[151,443,849,873]" fontSize="10" level="2" pageId="32" pageNumber="363" reason="4">
<emphasis id="B907B82FFF85FFCAFF0DBE635EFDA219" bold="true" box="[151,443,849,873]" pageId="32" pageNumber="363">
<taxonomicName id="4C731FBEFF85FFCAFF0DBE635E26A219" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[151,352,849,873]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="363" phylum="Arthropoda" rank="species" species="nilsi" status="sp. nov.">
<emphasis id="B907B82FFF85FFCAFF0DBE635E26A219" bold="true" box="[151,352,849,873]" italics="true" pageId="32" pageNumber="363">Sanaungulus nilsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF85FFCAFEFCBE635EFDA219" box="[358,443,849,873]" pageId="32" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF85FFCAFF0DBE435FB2A2F8" box="[151,244,881,904]" pageId="32" pageNumber="363" type="description">
<paragraph id="8BCC643DFF85FFCAFF0DBE435FB2A2F8" blockId="32.[151,443,849,904]" box="[151,244,881,904]" pageId="32" pageNumber="363">
(
<figureCitation id="134878B8FF85FFCAFF05BE435FAAA2F8" box="[159,236,881,904]" captionStart="Fig" captionStartId="33.[113,152,1409,1432]" captionTargetBox="[113,1134,151,1376]" captionTargetId="figure-28@33.[113,1134,151,1376]" captionTargetPageId="33" captionText="Fig. 18. Sanaungulus nilsi sp. nov. in Burmese amber: A: Holotype, lateral view; B: Holotype, ventral view." pageId="32" pageNumber="363">Fig. 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF85FFCAFF0DBE9A5EF0A59F" pageId="32" pageNumber="363" type="materials_examined">
<paragraph id="8BCC643DFF85FFCAFF0DBE9A5D02A530" blockId="32.[151,643,936,1088]" pageId="32" pageNumber="363">
<materialsCitation id="3B1B6E60FF85FFCAFF0DBE9A5D02A530" collectingDate="2021-12" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="32" pageNumber="363" specimenCode="BUB4393" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFF85FFCAFF0DBE9A5E42A2B0" bold="true" box="[151,260,936,960]" pageId="32" pageNumber="363">
<typeStatus id="54C8DA9FFF85FFCAFF0DBE9A5FB8A2B0" box="[151,254,936,960]" pageId="32" pageNumber="363" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FF85FFCAFE9DBE9A5E10A2CF" box="[263,342,936,959]" pageId="32" pageNumber="363">Female</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FF85FFCAFDBCBE9A5DC5A2CF" box="[550,643,936,959]" pageId="32" pageNumber="363">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFF85FFCAFF05BEFA5FA9A2AF" box="[159,239,968,991]" country="Myanmar" name="Kachin" pageId="32" pageNumber="363">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FF85FFCAFE12BEFA5E9DA2AF" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFE12BEFA5E9DA2AF" box="[392,475,968,991]" county="Female" municipality="Burmese" name="Catalog" pageId="32" pageNumber="363" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFF85FFCAFF79BEDB5E01A570" box="[227,327,1001,1024]" pageId="32" pageNumber="363" value="2021-12">
<collectingDate id="EF89BB15FF85FFCAFF79BEDB5E01A570" box="[227,327,1001,1024]" pageId="32" pageNumber="363" value="2021-12">2021 XII</collectingDate>
</date>
<quantity id="4C8BC9D8FF85FFCAFECABEDB5ECEA570" box="[336,392,1001,1024]" metricMagnitude="0" metricUnit="m" metricValue="1.6256" pageId="32" pageNumber="363" unit="in" value="64.0">64 in</quantity>
the
<location id="8EAC32E6FF85FFCAFE21BEDB5E40A550" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFE21BEDB5E40A550" county="Female" municipality="Burmese" name="Bayerische Staatssammlung" pageId="32" pageNumber="363" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FF85FFCAFEA8B93B5E83A550" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFEA8B93B5E83A550" box="[306,453,1033,1056]" county="Female" municipality="Burmese" name="Palaontologie" pageId="32" pageNumber="363" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FF85FFCAFE63B93B5D1DA550" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFE63B93B5D1DA550" box="[505,603,1033,1056]" county="Female" municipality="Burmese" name="Geologie" pageId="32" pageNumber="363" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FF85FFCAFF0DB91B5FB4A530" LSID="urn:lsid:plazi:treatment:03DAD52BFF85FFC8FF0DBE635B00A1F0:8EAC32E6FF85FFCAFF0DB91B5FB4A530" box="[151,242,1065,1088]" county="Female" municipality="Burmese" name="Muller's" pageId="32" pageNumber="363" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FF85FFCAFE5DB91B5D7EA530" box="[455,568,1065,1088]" collectionCode="BUB" pageId="32" pageNumber="363">BUB4393</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFF85FFCAFF0DB9525EACA5E7" blockId="32.[151,643,1120,1175]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DB9525E76A508" bold="true" box="[151,304,1120,1144]" pageId="32" pageNumber="363">Type locality.</emphasis>
<collectingCountry id="F36424ADFF85FFCAFEA2B9525EE2A507" box="[312,420,1120,1143]" name="Myanmar" pageId="32" pageNumber="363">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFF85FFCAFE2AB9525D7CA507" box="[432,570,1120,1143]" country="Myanmar" name="Kachin" pageId="32" pageNumber="363">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFF85FFCAFF0DB9855EF0A59F" blockId="32.[151,643,1207,1263]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DB9855E6CA5BF" bold="true" box="[151,298,1207,1231]" pageId="32" pageNumber="363">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF85FFCAFF0DB83C5FA4A4F6" pageId="32" pageNumber="363" type="diagnosis">
<paragraph id="8BCC643DFF85FFCAFF0DB83C5FA4A4F6" blockId="32.[151,643,1294,1414]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DB83C5ED2A456" bold="true" box="[151,404,1294,1318]" pageId="32" pageNumber="363">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFF85FFCAFE06B83D5D1AA456" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[412,604,1295,1318]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="363" phylum="Arthropoda" rank="species" species="nilsi" status="sp. nov.">
<emphasis id="B907B82FFF85FFCAFE06B83D5D1AA456" box="[412,604,1295,1318]" italics="true" pageId="32" pageNumber="363">Sanaungulus nilsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF85FFCAFDFEB83C5F83A436" pageId="32" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
differs from other species of
<taxonomicName id="4C731FBEFF85FFCAFE63B81D5DC5A436" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[505,643,1327,1350]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="363" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF85FFCAFE63B81D5DC5A436" box="[505,643,1327,1350]" italics="true" pageId="32" pageNumber="363">Sanaungulus</emphasis>
</taxonomicName>
with filiform antennae by its very elongated pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF85FFCBFF0DB8945DF9A1F0" lastPageId="33" lastPageNumber="364" pageId="32" pageNumber="363" type="description">
<paragraph id="8BCC643DFF85FFCAFF0DB8945EEFA76E" blockId="32.[151,643,1446,1566]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DB8945E65A4CE" bold="true" box="[151,291,1446,1470]" pageId="32" pageNumber="363">Description.</emphasis>
Adult, winged, slender. Female, defined on the basis of short antennae and large abdomen. Body length
<quantity id="4C8BC9D8FF85FFCAFE10B8D55E9BA48E" box="[394,477,1511,1534]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="32" pageNumber="363" unit="mm" value="3.3">3.3 mm</quantity>
. Entirely brown with testaceous abdomen.
</paragraph>
<paragraph id="8BCC643DFF85FFCAFF0DBB0C5C40A7E2" blockId="32.[151,643,1598,1685]" lastBlockId="32.[680,1172,155,1682]" pageId="32" pageNumber="363">Head rounded, slightly constricted (triangular­shaped) behind the eyes, equipped with dense pubescence and shallow punctation. Eyes prominent, large, roundish, inserted in the upper and lateral part of the head. Mandibles elongated, falciform, without teeth. Maxillary palps 4­ segmented with the last palpomere very elongated and securiform. Labial palps 3­ segmented, with the last palpomere elongated and securiform. Antennae 11­segmented, filiform, robust, short, reaching half of elytra and the first abdominal segment, antennal insertion in the eyes proximity; scape robust, club­shaped; antennomeres IIIII short, subequal, thinner than first; antennomere IV elongated, the longest; antennomeres VX slightly shorter than antennomere IV, subequal, with antennomeres V VIII sturdier than others; antennomere XI filiform, longer than previous ones, enlarged in the middle, rounded at apex; all the antennomeres with short pubescence. Pronotum very elongated, subrectangular, as wide as head, concave in the middle, anterior margin straight, posterior margin slightly undulated and strongly bordered, sides straight and bordered, corners obtuse, pronotum disc concave and pubescent. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, at humeri slightly wider than pronotum, posteriorly parallel­sided and slighty dehiscent apically, rounded apexes, surface pubescent and with a very small rugosity. Metathoracic wings longer than elytra, slightly surpassing the half of abdomen. Sternum elongated with posterior margin rounded, sternites large and transverse, last tergite elongated with rounded apex and narrower than penultimate tergite which is large and transverse, last sternite wide and enlarged at sides and slightly pointed apically. Legs thin, relatively long, pubescent; coxae massive; trochanters elongated, with rounded apex; femora slightly curved, more robust than tibiae, slightly flat; tibiae flattened, thin, with spurs near the apex, protibiae as long as profemora, mesotibiae slightly longer than mesofemora, metatibiae longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 0.4 times longer than second; third tarsomere short and slightly bilobed; fourth strongly bilobed; fifth tarsomere elongated, thin, slightly flat and curved; claws simple, without denticle at the base.</paragraph>
<paragraph id="8BCC643DFF84FFCBFDBABD5F5DF9A1F0" blockId="33.[544,703,109,128]" box="[544,703,109,128]" pageId="33" pageNumber="364">
<emphasis id="B907B82FFF84FFCBFDBABD5F5DF9A1F0" box="[544,703,109,128]" italics="true" pageId="33" pageNumber="364">Fanti F., Müller P.</emphasis>
</paragraph>
</subSubSection>
<caption id="DF0C34B5FF84FFCBFFEBB8B35FBCA4C8" pageId="33" pageNumber="364" startId="33.[113,152,1409,1432]" targetBox="[113,1134,151,1376]" targetPageId="33" targetType="figure">
<paragraph id="8BCC643DFF84FFCBFFEBB8B35FBCA4C8" blockId="33.[113,1134,1409,1464]" pageId="33" pageNumber="364">
Fig. 18.
<taxonomicName id="4C731FBEFF84FFCBFF56B8B35ECAA4E8" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[204,396,1409,1432]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="364" phylum="Arthropoda" rank="species" species="nilsi" status="sp. nov.">
<emphasis id="B907B82FFF84FFCBFF56B8B35ECAA4E8" box="[204,396,1409,1432]" italics="true" pageId="33" pageNumber="364">Sanaungulus nilsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF84FFCBFE0EB8B35EA1A4E8" box="[404,487,1409,1432]" pageId="33" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber: A: Holotype, lateral view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FF84FFCBFFEBB8DD5ECDA7F1" pageId="33" pageNumber="364" type="etymology">
<paragraph id="8BCC643DFF84FFCBFFEBB8DD5EF3A758" blockId="33.[113,604,1519,1576]" pageId="33" pageNumber="364">
<emphasis id="B907B82FFF84FFCBFFEBB8DD5FA9A777" bold="true" box="[113,239,1519,1543]" pageId="33" pageNumber="364">Etymology.</emphasis>
Named in honor of Nils Müller, son of the second author (Patrick).
</paragraph>
<paragraph id="8BCC643DFF84FFCBFFEBBB7A5ECDA7F1" blockId="33.[113,605,1608,1665]" pageId="33" pageNumber="364">
<emphasis id="B907B82FFF84FFCBFFEBBB7A5E5EA710" bold="true" box="[113,280,1608,1632]" pageId="33" pageNumber="364">Syninclusions.</emphasis>
<taxonomicName id="4C731FBEFF84FFCBFEB8BB7B5E81A710" box="[290,455,1609,1632]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="364" phylum="Arthropoda" rank="family">Cecidomyiidae</taxonomicName>
(leg), cicada (leg), air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF84FFC8FD19B8DD5B00A1F0" lastPageId="34" lastPageNumber="365" pageId="33" pageNumber="364" type="discussion">
<paragraph id="8BCC643DFF84FFCBFD19B8DD5DFBA739" blockId="33.[643,1134,1519,1609]" pageId="33" pageNumber="364">
<emphasis id="B907B82FFF84FFCBFD19B8DD5DB4A777" bold="true" box="[643,754,1519,1543]" pageId="33" pageNumber="364">Remarks.</emphasis>
Piece of clear, golden amber that measures 19x7.5x
<quantity id="4C8BC9D8FF84FFCBFCA1BB235CCAA758" box="[827,908,1553,1576]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="33" pageNumber="364" unit="mm" value="3.3">3.3 mm</quantity>
. The inclusion is complete.
</paragraph>
<paragraph id="8BCC643DFF87FFC8FF7CBD5F5B00A1F0" blockId="34.[230,1094,109,128]" box="[230,1094,109,128]" pageId="34" pageNumber="365">
<emphasis id="B907B82FFF87FFC8FF7CBD5F5B00A1F0" box="[230,1094,109,128]" italics="true" pageId="34" pageNumber="365">
Fossil
<taxonomicName id="4C731FBEFF87FFC8FEBBBD5F5ECBA1F0" authorityName="Imhoff" authorityYear="1856" box="[289,397,109,128]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
from the Cretaceous Burmese (
<collectingRegion id="49B7AADFFF87FFC8FD3ABD5F5D99A1F0" box="[672,735,109,128]" country="Myanmar" name="Kachin" pageId="34" pageNumber="365">Kachin</collectingRegion>
) amber of the Patrick Müller collection..
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,158 @@
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<mods:title id="23AB2BB159F47ADB0EBB67BBD31279FA">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<emphasis id="B907B82FFF86FFC9FD19B9C95CE1A463" bold="true" box="[643,935,1275,1299]" italics="true" pageId="35" pageNumber="366">Sanaungulus peteriruedeli</emphasis>
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<taxonomicNameLabel id="A2340554FF86FFC9FC37B9C95B44A463" box="[941,1026,1275,1299]" pageId="35" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C36937B6FF86FFC9FD19B8295DA6A442" box="[643,736,1307,1330]" pageId="35" pageNumber="366" type="description">
<paragraph id="8BCC643DFF86FFC9FD19B8295DA6A442" blockId="35.[643,1026,1275,1330]" box="[643,736,1307,1330]" pageId="35" pageNumber="366">
(
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)
</paragraph>
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<paragraph id="8BCC643DFF86FFC9FD19B8605C88A49A" blockId="35.[643,1134,1362,1514]" pageId="35" pageNumber="366">
<materialsCitation id="3B1B6E60FF86FFC9FD19B8605C88A49A" collectionCode="BUB" county="Sex" location="Catalog" municipality="Burmese" pageId="35" pageNumber="366" specimenCode="BUB4460" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFF86FFC9FD19B8605DB7A41A" bold="true" box="[643,753,1362,1386]" pageId="35" pageNumber="366">
<typeStatus id="54C8DA9FFF86FFC9FD19B8605DABA41A" box="[643,749,1362,1386]" pageId="35" pageNumber="366" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FF86FFC9FD62B8605C66A419" box="[760,800,1362,1385]" pageId="35" pageNumber="366">Sex</collectingCounty>
undefined, adult specimen in a
<collectingMunicipality id="6BA8FE47FF86FFC9FD19B8405D99A4F9" box="[643,735,1394,1417]" pageId="35" pageNumber="366">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFF86FFC9FD70B8405C7EA4F9" box="[746,824,1394,1417]" country="Myanmar" name="Kachin" pageId="35" pageNumber="366">Kachin</collectingRegion>
) amber piece:
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number SNHM­
<quantity id="4C8BC9D8FF86FFC9FD41B8A15C75A4DA" box="[731,819,1427,1450]" metricMagnitude="2" metricUnit="m" metricValue="1.527048" pageId="35" pageNumber="366" unit="in" value="6012.0">6012 in</quantity>
the
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(ex
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collection number:
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).
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<paragraph id="8BCC643DFF86FFC9FD19BB385C93A731" blockId="35.[643,1134,1546,1601]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FD19BB385C5AA752" bold="true" box="[643,796,1546,1570]" pageId="35" pageNumber="366">Type locality.</emphasis>
<collectingCountry id="F36424ADFF86FFC9FCBEBB385CD6A751" box="[804,912,1546,1569]" name="Myanmar" pageId="35" pageNumber="366">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFF86FFC9FC01BB385B63A751" box="[923,1061,1546,1569]" country="Myanmar" name="Kachin" pageId="35" pageNumber="366">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFF86FFC9FD19BB535CE4A7E9" blockId="35.[643,1134,1633,1689]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FD19BB535C50A709" bold="true" box="[643,790,1633,1657]" pageId="35" pageNumber="366">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
<paragraph id="8BCC643DFF81FFCEFF7CBD5F5B00A1F0" blockId="36.[230,1094,109,128]" box="[230,1094,109,128]" pageId="36" pageNumber="367">
<emphasis id="B907B82FFF81FFCEFF7CBD5F5B00A1F0" box="[230,1094,109,128]" italics="true" pageId="36" pageNumber="367">
Fossil
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from the Cretaceous Burmese (
<collectingRegion id="49B7AADFFF81FFCEFD3ABD5F5D99A1F0" box="[672,735,109,128]" country="Myanmar" name="Kachin" pageId="36" pageNumber="367">Kachin</collectingRegion>
) amber of the Patrick Müller collection..
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF0C34B5FF81FFCEFF0DB84D5ED6A4C6" pageId="36" pageNumber="367" startId="36.[151,190,1407,1430]" targetBox="[151,1172,151,1376]" targetPageId="36" targetType="figure">
<paragraph id="8BCC643DFF81FFCEFF0DB84D5ED6A4C6" blockId="36.[151,1172,1407,1462]" pageId="36" pageNumber="367">
Fig. 20.
<taxonomicName id="4C731FBEFF81FFCEFF6CB84D5D52A4E6" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[246,532,1407,1430]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="367" phylum="Arthropoda" rank="species" species="peteriruedeli" status="sp. nov.">
<emphasis id="B907B82FFF81FFCEFF6CB84D5D52A4E6" box="[246,532,1407,1430]" italics="true" pageId="36" pageNumber="367">Sanaungulus peteriruedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF81FFCEFD84B84D5D35A4E6" box="[542,627,1407,1430]" pageId="36" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FF81FFCEFF0DB8DE5EB5A715" pageId="36" pageNumber="367" type="diagnosis">
<paragraph id="8BCC643DFF81FFCEFF0DB8DE5EB5A715" blockId="36.[151,643,1516,1637]" pageId="36" pageNumber="367">
<emphasis id="B907B82FFF81FFCEFF0DB8DE5ED6A774" bold="true" box="[151,400,1516,1540]" pageId="36" pageNumber="367">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFF81FFCEFE0FB8DE5F8DA754" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="367" phylum="Arthropoda" rank="species" species="peteriruedeli" status="sp. nov.">
<emphasis id="B907B82FFF81FFCEFE0FB8DE5F8DA754" italics="true" pageId="36" pageNumber="367">Sanaungulus peteriruedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF81FFCEFF49BB3F5E61A754" box="[211,295,1549,1572]" pageId="36" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by its antennae with very large and robust antennomeres, in particular antennomeres IIIVII.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF81FFCFFD32B8D95F89A7E7" lastPageId="37" lastPageNumber="368" pageId="36" pageNumber="367" type="description">
<paragraph id="8BCC643DFF81FFCEFD32B8D95BD3A752" blockId="36.[680,1173,1515,1570]" pageId="36" pageNumber="367">
<emphasis id="B907B82FFF81FFCEFD32B8D95C75A773" bold="true" box="[680,819,1515,1539]" pageId="36" pageNumber="367">Description.</emphasis>
Adult, winged, rather robust. Sex undefined. Body length
<quantity id="4C8BC9D8FF81FFCEFC06BB395CAAA752" box="[924,1004,1547,1570]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="36" pageNumber="367" unit="mm" value="4.1">4.1 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFF81FFCFFD32BB705F89A7E7" blockId="36.[680,1172,1602,1689]" lastBlockId="37.[113,606,155,1687]" lastPageId="37" lastPageNumber="368" pageId="36" pageNumber="367">Head transverse, constricted (triangular­shaped) behind the eyes, surface irregular. Eyes prominent, very large, roundish, inserted in the upper and lateral part of the head. Mandibles elongated, robust, falciform. Maxillary palps 4­ segmented with the last palpomere strongly elongated and securiform. Labial palps 3­ segmented, with the last palpomere securiform. Antennae 11­ segmented, slightly surpassing half of elytra and not reaching half of abdomen, antennal insertion far from the eyes; scape club­shaped, elongated, robust; antennomere II very short, small, swollen apically; antennomeres IIIIV extremely wide and robust, flat, stouter than scape; antennomeres VVI robust, less wide and slightly longer than previous ones; antennomeres VIIX robust, narrower than previous ones; antennomere XI robust, with rounded apex; all the antennomeres pubescent. Pronotum very elongated, slightly narrower than head, anterior margin undulate and not bordered, posterior margin slightly curved in the middle and slightly bordered, sides straight except for a small tooth near the apex and not bordered, corners obtuse, pronotum disc irregular with large drafts. Scutellum triangular­shaped, wide basally with rounded apex. Elytra short which reveals some abdominal segments, at humeri strongly wider than pronotum, posteriorly parallel­sided and slightly dehiscent apically, rounded apexes, surface smooth (only few and very small punctuation) with pubescence. Metathoracic wings transparent, approximatively as long as elytra. Sternum very elongated and slender, sternites transverse and pubescent, last sternite small, penultimate abdominal segment with a long lobe at the apical corners. Legs thin, long, slightly pubescent; coxae massive; trochanters elongated, robust, with thin and rounded apex; femora curved, cylindrical, slightly sturdier than tibiae; tibiae, thin, subcylindrical, long, with a small spur near the apex, pro­ and mesotibiae slightly longer than pro­ and mesofemora, metatibiae longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 1.1­ 1.2 times longer than second; third tarsomere short, triangular­shaped; fourth strongly bilobed; fifth tarsomere extremely elongated, thin, cylindrical and curved; claws simple, long, pointed, without denticle at the base.</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF80FFCFFD19BDA85B51A038" pageId="37" pageNumber="368" type="etymology">
<paragraph id="8BCC643DFF80FFCFFD19BDA85C68A182" blockId="37.[643,1134,154,242]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19BDA85C45A1C2" bold="true" box="[643,771,154,178]" pageId="37" pageNumber="368">Etymology.</emphasis>
Named in honor of Peter Rüdel, friend of the second author (Patrick), and a great amber collector.
</paragraph>
<paragraph id="8BCC643DFF80FFCFFD19BC235B51A038" blockId="37.[643,1134,273,328]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19BC235C60A059" bold="true" box="[643,806,273,297]" pageId="37" pageNumber="368">Syninclusions.</emphasis>
Detritus, air bubbles (small), a specimen of
<taxonomicName id="4C731FBEFF80FFCFFC90BC035C39A038" authorityName="Handlirsch" authorityYear="1906" box="[778,895,305,328]" class="Insecta" family="Berothidae" kingdom="Animalia" order="Neuroptera" pageId="37" pageNumber="368" phylum="Arthropoda" rank="family">Berothidae</taxonomicName>
(
<taxonomicName id="4C731FBEFF80FFCFFC17BC035B4AA038" box="[909,1036,305,328]" class="Insecta" kingdom="Animalia" order="Neuroptera" pageId="37" pageNumber="368" phylum="Arthropoda" rank="order">Neuroptera</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF80FFCFFD19BC5A5CE8A0AF" pageId="37" pageNumber="368" type="discussion">
<paragraph id="8BCC643DFF80FFCFFD19BC5A5CE8A0AF" blockId="37.[643,1134,360,479]" pageId="37" pageNumber="368">
<emphasis id="B907B82FFF80FFCFFD19BC5A5DB7A0F0" bold="true" box="[643,753,360,384]" pageId="37" pageNumber="368">Remarks.</emphasis>
Piece of brownish Burmese amber that measures 14x9x
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. The inclusion is complete and only opacified in the last sternites due to the presence of emulsion.
</paragraph>
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</treatment>
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<mods:part id="26421E1D0AF3D9F8B5B9F00B62CA79E0">
<mods:date id="704AD13BFFF97BA96639F67AAD223E0E">2022</mods:date>
<mods:detail id="CA0EF4DFD3269BB7892638307EFDE1DE" type="volume">
<mods:number id="3616F87848A0B9BE1091E9460E1106E7">22</mods:number>
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<treatment id="03DAD52BFF87FFC9FF0DB8DD5CE7A5B4" ID-DOI="http://doi.org/10.5281/zenodo.13199833" ID-Zenodo-Dep="13199833" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4" httpUri="http://treatment.plazi.org/id/03DAD52BFF87FFC9FF0DB8DD5CE7A5B4" lastPageId="35" lastPageNumber="366" pageId="34" pageNumber="365">
<subSubSection id="C36937B6FF87FFC8FF0DB8DD5D47A777" box="[151,513,1519,1543]" pageId="34" pageNumber="365" type="nomenclature">
<paragraph id="8BCC643DFF87FFC8FF0DB8DD5D47A777" blockId="34.[151,513,1519,1574]" box="[151,513,1519,1543]" pageId="34" pageNumber="365">
<heading id="D084D351FF87FFC8FF0DB8DD5D47A777" bold="true" box="[151,513,1519,1543]" fontSize="10" level="2" pageId="34" pageNumber="365" reason="4">
<emphasis id="B907B82FFF87FFC8FF0DB8DD5D47A777" bold="true" box="[151,513,1519,1543]" pageId="34" pageNumber="365">
<taxonomicName id="4C731FBEFF87FFC8FF0DB8DD5EE0A777" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[151,422,1519,1543]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="species" species="perkovskyi" status="sp. nov.">
<emphasis id="B907B82FFF87FFC8FF0DB8DD5EE0A777" bold="true" box="[151,422,1519,1543]" italics="true" pageId="34" pageNumber="365">Sanaungulus perkovskyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF87FFC8FE36B8DD5D47A777" box="[428,513,1519,1543]" pageId="34" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF87FFC8FF0DBB3D5FB2A756" box="[151,244,1551,1574]" pageId="34" pageNumber="365" type="description">
<paragraph id="8BCC643DFF87FFC8FF0DBB3D5FB2A756" blockId="34.[151,513,1519,1574]" box="[151,244,1551,1574]" pageId="34" pageNumber="365">
(
<figureCitation id="134878B8FF87FFC8FF05BB3D5FAAA756" box="[159,236,1551,1574]" captionStart="Fig" captionStartId="34.[151,189,1409,1432]" captionTargetBox="[151,1172,151,1376]" captionTargetId="figure-73@34.[151,1172,151,1376]" captionTargetPageId="34" captionText="Fig.19. Sanaungulus perkovskyi sp. nov. in Burmese amber.A: Holotype, dorsal view; B: Holotype, ventral view." pageId="34" pageNumber="365">Fig. 19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF87FFC9FF0DBB775ED6A1A3" lastPageId="35" lastPageNumber="366" pageId="34" pageNumber="365" type="materials_examined">
<paragraph id="8BCC643DFF87FFC8FF0DBB775B13A756" blockId="34.[151,643,1605,1693]" lastBlockId="34.[680,1171,1519,1574]" pageId="34" pageNumber="365">
<materialsCitation id="3B1B6E60FF87FFC8FF0DBB775B13A756" collectingDate="2021-12" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="34" pageNumber="365" specimenCode="BUB3888" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFF87FFC8FF0DBB775E40A72D" bold="true" box="[151,262,1605,1629]" pageId="34" pageNumber="365">
<typeStatus id="54C8DA9FFF87FFC8FF0DBB775E47A72D" box="[151,257,1605,1629]" pageId="34" pageNumber="365" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FF87FFC8FE96BB745E03A72D" box="[268,325,1606,1629]" pageId="34" pageNumber="365">Male</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FF87FFC8FDBEBB745DC5A72D" box="[548,643,1606,1629]" pageId="34" pageNumber="365">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFF87FFC8FF05BB545FA9A70D" box="[159,239,1638,1661]" country="Myanmar" name="Kachin" pageId="34" pageNumber="365">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FF87FFC8FE12BB545E9DA70D" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FE12BB545E9DA70D" box="[392,475,1638,1661]" county="Male" municipality="Burmese" name="Catalog" pageId="34" pageNumber="365" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFF87FFC8FF79BBB45E01A7ED" box="[227,327,1670,1693]" pageId="34" pageNumber="365" value="2021-12">
<collectingDate id="EF89BB15FF87FFC8FF79BBB45E01A7ED" box="[227,327,1670,1693]" pageId="34" pageNumber="365" value="2021-12">2021 XII</collectingDate>
</date>
<quantity id="4C8BC9D8FF87FFC8FECABBB45ECEA7ED" box="[336,392,1670,1693]" metricMagnitude="0" metricUnit="m" metricValue="1.7526" pageId="34" pageNumber="365" unit="in" value="69.0">69 in</quantity>
the
<location id="8EAC32E6FF87FFC8FE21BBB45C51A776" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FE21BBB45C51A776" box="[443,791,1519,1693]" county="Male" municipality="Burmese" name="Bayerische Staatssammlung" pageId="34" pageNumber="365" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FF87FFC8FCD9B8DD5C90A776" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FCD9B8DD5C90A776" box="[835,982,1519,1542]" county="Male" municipality="Burmese" name="Palaontologie" pageId="34" pageNumber="365" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FF87FFC8FB90B8DD5B2AA776" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FB90B8DD5B2AA776" box="[1034,1132,1519,1542]" county="Male" municipality="Burmese" name="Geologie" pageId="34" pageNumber="365" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FF87FFC8FD32BB3D5C42A756" LSID="urn:lsid:plazi:treatment:03DAD52BFF87FFC9FF0DB8DD5CE7A5B4:8EAC32E6FF87FFC8FD32BB3D5C42A756" box="[680,772,1551,1574]" county="Male" municipality="Burmese" name="Muller's" pageId="34" pageNumber="365" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FF87FFC8FC43BB3D5B0FA756" box="[985,1097,1551,1574]" collectionCode="BUB" pageId="34" pageNumber="365">BUB3888</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFF87FFC8FD32BB775CBDA70D" blockId="34.[680,1172,1605,1661]" pageId="34" pageNumber="365">
<emphasis id="B907B82FFF87FFC8FD32BB775C07A72D" bold="true" box="[680,833,1605,1629]" pageId="34" pageNumber="365">Type locality.</emphasis>
<collectingCountry id="F36424ADFF87FFC8FCD0BB745CF0A72D" box="[842,950,1606,1629]" name="Myanmar" pageId="34" pageNumber="365">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFF87FFC8FC5BBB745B0DA72D" box="[961,1099,1606,1629]" country="Myanmar" name="Kachin" pageId="34" pageNumber="365">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFF86FFC9FFEBBDA95ED6A1A3" blockId="35.[113,605,155,211]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FFEBBDA95E43A1C3" bold="true" box="[113,261,155,179]" pageId="35" pageNumber="366">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF86FFC9FFEBBDC15D15A3FA" pageId="35" pageNumber="366" type="diagnosis">
<paragraph id="8BCC643DFF86FFC9FFEBBDC15D15A3FA" blockId="35.[113,606,243,650]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FFEBBDC15E26A07B" bold="true" box="[113,352,243,267]" pageId="35" pageNumber="366">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFF86FFC9FEFFBDC15D1BA07A" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[357,605,243,266]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="366" phylum="Arthropoda" rank="species" species="perkovskyi" status="sp. nov.">
<emphasis id="B907B82FFF86FFC9FEFFBDC15D1BA07A" box="[357,605,243,266]" italics="true" pageId="35" pageNumber="366">Sanaungulus perkovskyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF86FFC9FFEBBC215F86A05A" box="[113,192,275,298]" pageId="35" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by a strongly transverse pronotum and pectinate antennae (the antennal process in antennomeres IVVII).
<taxonomicName id="4C731FBEFF86FFC9FE4FBC615F93A0DA" authority="Fanti, Damgaard &amp; Ellenberger, 2018" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="366" phylum="Arthropoda" rank="species" species="ghitaenoerbyae">
<emphasis id="B907B82FFF86FFC9FE4FBC615E51A0FA" italics="true" pageId="35" pageNumber="366">Sanaungulus ghitaenoerbyae</emphasis>
<bibRefCitation id="EFE219CCFF86FFC9FEBABC415F93A0DA" author="Fanti F. &amp; Damgaard A. L. &amp; Ellenberger S." pageId="35" pageNumber="366" pagination="170 - 177" refId="ref20442" refString="Fanti F., Damgaard A. L., Ellenberger S. 2018. Two new genera of Cantharidae from Burmese amber of the Hukawng Valley (Insecta, Coleoptera). Cretaceous Research 86: 170 - 177." type="journal article" year="2018">Fanti, Damgaard &amp; Ellenberger, 2018</bibRefCitation>
</taxonomicName>
is the most similar taxon and differs from the new species because of its more transverse head and its pronotum with its sides and posterior margin not straight (Fanti et al. 2018).
<taxonomicName id="4C731FBEFF86FFC9FFEBBF215FA3A33A" authority="Fanti &amp; Damgaard, 2019" authorityName="Fanti &amp; Damgaard" authorityYear="2019" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="366" phylum="Arthropoda" rank="species" species="troelsikloevedali">
<emphasis id="B907B82FFF86FFC9FFEBBF215EF0A35A" box="[113,438,531,554]" italics="true" pageId="35" pageNumber="366">Sanaungulus troelsikloevedali</emphasis>
<bibRefCitation id="EFE219CCFF86FFC9FE5BBF215FA3A33A" author="Fanti F. &amp; Damgaard A. L." pageId="35" pageNumber="366" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti &amp; Damgaard, 2019</bibRefCitation>
</taxonomicName>
differs from the new species described here by its longer antennae with more slender last antennomeres (
<bibRefCitation id="EFE219CCFF86FFC9FEA4BF415D01A3FA" author="Fanti F. &amp; Damgaard A. L." box="[318,583,627,650]" pageId="35" pageNumber="366" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti &amp; Damgaard 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF86FFC9FFEBBF9B5CEAA2EE" pageId="35" pageNumber="366" type="description">
<paragraph id="8BCC643DFF86FFC9FFEBBF9B5EC8A251" blockId="35.[113,605,681,801]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FFEBBF9B5FB0A3B1" bold="true" box="[113,246,681,705]" pageId="35" pageNumber="366">Description.</emphasis>
Adult, winged, rather robust. Male, defined on the basis of the last abdominal segment triangular shaped. Body length
<quantity id="4C8BC9D8FF86FFC9FD99BFD85D1FA271" box="[515,601,746,769]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="35" pageNumber="366" unit="mm" value="3.2">3.2 mm</quantity>
. Entirely testaceous­brown.
</paragraph>
<paragraph id="8BCC643DFF86FFC9FFEBBE725CEAA2EE" blockId="35.[113,606,832,1687]" lastBlockId="35.[643,1135,155,926]" pageId="35" pageNumber="366">Head not particularly transverse, constricted (triangular­shaped) behind the eyes, pubescent with shallow punctation. Eyes prominent, large, roundish, inserted in the upper and lateral part of the head. Mandibles elongated, falciform. Maxillary palps 4­ segmented with the last palpomere securiform. Labial palps 3­ segmented, with the last palpomere securiform. Antennae 11­ segmented, pectinate, approximately slightly surpassing the half of elytra, antennal insertion far from the eyes; scape club­shaped, robust; antennomeres IIIII short with antennomere II sturdier and swollen apically; antennomeres IV VI extremely long, each equipped with an antennal process at apex which is very long and robust and slightly pointed apically; antennomere VII shorter than previous ones, equipped with a long antennal process; antennomere VIII sturdier and shorter than previous one and without the antennal process; antennomeres IXX very robust, flat, enlarged at sides; antennomere XI filiform, less sturdier than previous ones, with rounded apex; all the antennomeres pubescent with setae very short.Pronotum transverse, wider than head, anterior margin undulate and strongly bordered, posterior margin straight and bordered, sides straight and bordered, corners obtuse, pronotum disc pubescent with shallow punctation. Scutellum triangular­shaped, very wide at base, with rounded apex. Elytra short which reveals some abdominal segments, robust, wider than pronotum, posteriorly parallel­sided and not dehiscent apically, rounded apexes, surface with coarse and impressed punctation gathered in striae. Metathoracic wings transparent, longer than elytra. Sternum elongated with posterior margin almost straight, sternites transverse and pubescent, last sternite small and triangular shaped. Legs thin, long, slightly pubescent; coxae massive; trochanters small, rounded; femora almost straight, slightly flat, sturdier than tibiae; tibiae, thin, cylindrical, long, without spurs near the apex, pro­ and mesotibiae very slightly longer than pro­ and mesofemora, metatibiae longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 1.4 times longer than second; third tarsomere short, triangular­shaped; fourth strongly bilobed with lobes elongated and rounded apically; fifth tarsomere elongated, thin, cylindrical and curved; claws simple, long, pointed, without denticle at the base.</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF86FFC9FD19BE8C5B29A55C" pageId="35" pageNumber="366" type="etymology">
<paragraph id="8BCC643DFF86FFC9FD19BE8C5B29A285" blockId="35.[643,1135,958,1013]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FD19BE8C5C44A2A6" bold="true" box="[643,770,958,982]" pageId="35" pageNumber="366">Etymology.</emphasis>
Named in honor of Evgeny E. Perkovsky (
<collectingCountry id="F36424ADFF86FFC9FD42BEEC5C76A285" box="[728,816,990,1013]" name="Ukraine" pageId="35" pageNumber="366">Ukraine</collectingCountry>
), a well­known palaeontologist.
</paragraph>
<paragraph id="8BCC643DFF86FFC9FD19B9275B29A55C" blockId="35.[643,1135,1045,1069]" box="[643,1135,1045,1069]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FD19B9275C58A55D" bold="true" box="[643,798,1045,1069]" pageId="35" pageNumber="366">Syninclusions.</emphasis>
Air bubbles (very large), detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF86FFC9FD19B97E5CE7A5B4" pageId="35" pageNumber="366" type="discussion">
<paragraph id="8BCC643DFF86FFC9FD19B97E5CE7A5B4" blockId="35.[643,1134,1100,1220]" pageId="35" pageNumber="366">
<emphasis id="B907B82FFF86FFC9FD19B97E5DB7A514" bold="true" box="[643,753,1100,1124]" pageId="35" pageNumber="366">Remarks.</emphasis>
Piece of very clear amber that measures 12x8x
<quantity id="4C8BC9D8FF86FFC9FC9EB95F5C00A5F4" box="[772,838,1133,1156]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="366" unit="mm" value="2.0">2 mm</quantity>
. The inclusion is complete, but many big bubbles in the amber cover the majority of its ventral part.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,158 @@
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<mods:title id="7143C1D0A78D63CE6A964818E63025FF">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<treatment id="03DAD52BFF8CFFC1FD19BFDA5FA4A7EA" LSID="urn:lsid:plazi:treatment:03DAD52BFF8CFFC1FD19BFDA5FA4A7EA" httpUri="http://treatment.plazi.org/id/03DAD52BFF8CFFC1FD19BFDA5FA4A7EA" lastPageId="43" lastPageNumber="374" pageId="41" pageNumber="372">
<subSubSection id="C36937B6FF8CFFC3FD19BFDA5CAFA38F" box="[643,1001,743,768]" pageId="41" pageNumber="372" type="nomenclature">
<paragraph id="8BCC643DFF8CFFC3FD19BFDA5CAFA38F" blockId="41.[643,1001,743,799]" box="[643,1001,743,768]" pageId="41" pageNumber="372">
<heading id="D084D351FF8CFFC3FD19BFDA5CAFA38F" bold="true" box="[643,1001,743,768]" fontSize="10" level="2" pageId="41" pageNumber="372" reason="4">
<emphasis id="B907B82FFF8CFFC3FD19BFDA5CAFA38F" bold="true" box="[643,1001,743,768]" pageId="41" pageNumber="372">
<taxonomicName id="4C731FBEFF8CFFC3FD19BFDA5CC8A270" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[643,910,744,768]" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="species" species="burmiticus" status="sp. nov.">
<emphasis id="B907B82FFF8CFFC3FD19BFDA5CC8A270" bold="true" box="[643,910,744,768]" italics="true" pageId="41" pageNumber="372">Vitalfranzius burmiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF8CFFC3FC0EBFD55CAFA38F" box="[916,1001,743,767]" pageId="41" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8CFFC3FD19BE3A5DA6A26F" box="[643,736,776,799]" pageId="41" pageNumber="372" type="description">
<paragraph id="8BCC643DFF8CFFC3FD19BE3A5DA6A26F" blockId="41.[643,1001,743,799]" box="[643,736,776,799]" pageId="41" pageNumber="372">
(
<figureCitation id="134878B8FF8CFFC3FD11BE3A5D9EA26F" box="[651,728,776,799]" captionStart="Fig" captionStartId="42.[151,190,1636,1659]" captionTargetBox="[151,1171,383,1608]" captionTargetId="figure-153@42.[151,1171,383,1608]" captionTargetPageId="42" captionText="Fig. 23. Vitalfranzius burmiticus gen. et sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view." pageId="41" pageNumber="372">Fig. 23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8CFFC3FD19BE0D5CE4A5F8" pageId="41" pageNumber="372" type="materials_examined">
<paragraph id="8BCC643DFF8CFFC3FD19BE0D5B69A2A9" blockId="41.[643,1134,831,985]" pageId="41" pageNumber="372">
<materialsCitation id="3B1B6E60FF8CFFC3FD19BE0D5B69A2A9" collectingDate="2021-12-15" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="41" pageNumber="372" specimenCode="BUB4313" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFF8CFFC3FD19BE0D5DA9A227" bold="true" box="[643,751,831,855]" pageId="41" pageNumber="372">
<typeStatus id="54C8DA9FFF8CFFC3FD19BE0D5DACA227" box="[643,746,831,855]" pageId="41" pageNumber="372" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FF8CFFC3FD69BE0D5C04A226" box="[755,834,831,854]" pageId="41" pageNumber="372">Female</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FF8CFFC3FB8BBE0D5B28A226" box="[1041,1134,831,854]" pageId="41" pageNumber="372">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFF8CFFC3FD10BE525D9CA207" box="[650,730,864,887]" country="Myanmar" name="Kachin" pageId="41" pageNumber="372">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FF8CFFC3FCE9BE525C80A207" LSID="urn:lsid:plazi:treatment:03DAD52BFF8CFFC1FD19BFDA5FA4A7EA:8EAC32E6FF8CFFC3FCE9BE525C80A207" box="[883,966,864,887]" county="Female" municipality="Burmese" name="Catalog" pageId="41" pageNumber="372" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFF8CFFC3FD54BEB25C10A2E7" box="[718,854,896,919]" pageId="41" pageNumber="372" value="2021-12-15">
<collectingDate id="EF89BB15FF8CFFC3FD54BEB25C10A2E7" box="[718,854,896,919]" pageId="41" pageNumber="372" value="2021-12-15">
2021 XII
<quantity id="4C8BC9D8FF8CFFC3FCA1BEB25C10A2E7" box="[827,854,896,919]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="41" pageNumber="372" unit="in" value="15.0">15</quantity>
</collectingDate>
</date>
in the
<location id="8EAC32E6FF8CFFC3FC3CBEB25DB4A2C8" LSID="urn:lsid:plazi:treatment:03DAD52BFF8CFFC1FD19BFDA5FA4A7EA:8EAC32E6FF8CFFC3FC3CBEB25DB4A2C8" county="Female" municipality="Burmese" name="Bayerische Staatssammlung" pageId="41" pageNumber="372" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FF8CFFC3FC87BE935CF6A2C8" LSID="urn:lsid:plazi:treatment:03DAD52BFF8CFFC1FD19BFDA5FA4A7EA:8EAC32E6FF8CFFC3FC87BE935CF6A2C8" box="[797,944,929,952]" county="Female" municipality="Burmese" name="Palaontologie" pageId="41" pageNumber="372" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FF8CFFC3FC7EBE935B00A2C8" LSID="urn:lsid:plazi:treatment:03DAD52BFF8CFFC1FD19BFDA5FA4A7EA:8EAC32E6FF8CFFC3FC7EBE935B00A2C8" box="[996,1094,929,952]" county="Female" municipality="Burmese" name="Geologie" pageId="41" pageNumber="372" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FF8CFFC3FD19BEF05D99A2A9" LSID="urn:lsid:plazi:treatment:03DAD52BFF8CFFC1FD19BFDA5FA4A7EA:8EAC32E6FF8CFFC3FD19BEF05D99A2A9" box="[643,735,962,985]" county="Female" municipality="Burmese" name="Muller's" pageId="41" pageNumber="372" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FF8CFFC3FC29BEF05B65A2A9" box="[947,1059,962,985]" collectionCode="BUB" pageId="41" pageNumber="372">BUB4313</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFF8CFFC3FD19BECA5C93A540" blockId="41.[643,1134,1016,1072]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FD19BECA5C5AA560" bold="true" box="[643,796,1016,1040]" pageId="41" pageNumber="372">Type locality.</emphasis>
<collectingCountry id="F36424ADFF8CFFC3FCBEBECB5CD6A560" box="[804,912,1017,1040]" name="Myanmar" pageId="41" pageNumber="372">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFF8CFFC3FC01BECB5B63A560" box="[923,1061,1017,1040]" country="Myanmar" name="Kachin" pageId="41" pageNumber="372">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFF8CFFC3FD19B9625CE4A5F8" blockId="41.[643,1134,1104,1160]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FD19B9625C50A518" bold="true" box="[643,790,1104,1128]" pageId="41" pageNumber="372">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8CFFC3FD19B99A5C9AA4F2" pageId="41" pageNumber="372" type="diagnosis">
<paragraph id="8BCC643DFF8CFFC3FD19B99A5C9AA4F2" blockId="41.[643,1134,1192,1410]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FD19B99A5C3AA5B0" bold="true" box="[643,892,1192,1216]" pageId="41" pageNumber="372">Differential diagnosis.</emphasis>
Smooth and very short elytra, the rounded head behind the eyes, antennae filiform, and the last abdominal segment slightly forked apically make this new species belong to the genus
<taxonomicName id="4C731FBEFF8CFFC3FCC9B8195C9DA432" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[851,987,1323,1346]" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B907B82FFF8CFFC3FCC9B8195C9DA432" box="[851,987,1323,1346]" italics="true" pageId="41" pageNumber="372">Vitalfranzius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF8CFFC3FC45B8185B07A432" box="[991,1089,1322,1346]" pageId="41" pageNumber="372" rank="genus">gen. nov.</taxonomicNameLabel>
The new species is characterized by filiform antennae and its transverse pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8CFFC1FD19B8905EF7A47D" lastPageId="43" lastPageNumber="374" pageId="41" pageNumber="372" type="description">
<paragraph id="8BCC643DFF8CFFC3FD19B8905B21A76B" blockId="41.[643,1135,1442,1563]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FD19B8905C48A4CA" bold="true" box="[643,782,1442,1466]" pageId="41" pageNumber="372">Description.</emphasis>
Adult, winged, rather robust. Female, defined on the shape of the last abdominal segment and on the basis of the short antennae. Body length 5.0 mm. Entirely reddish­brown.
</paragraph>
<paragraph id="8BCC643DFF8CFFC1FD19BB095EF7A47D" blockId="41.[643,1134,1595,1684]" lastBlockId="43.[113,605,155,1293]" lastPageId="43" lastPageNumber="374" pageId="41" pageNumber="372">Head strongly rounded, slightly transverse, slightly convex, not constricted (triangular­shaped) behind the eyes, surface smooth. Eyes prominent, large, roundish, inserted in the upper and dorsolateral part of the head. Mandibles elongated, falciform. Maxillary palps 4­ segmented with the last palpomere securiform. Labial palps 3­ segmented.Antennae 11­ segmented, filiform, slender, short, almost reaching the half of elytra but not reaching the half of abdomen, antennal insertion in the eyes proximity; scape elongated, robust, club­shaped; antennomere II short, globular; antennomeres IIIIX subequal in length, very elongated, longer than previous one; antennomere X slightly shorter and stouter than previous ones; antennomere XI filiform, longer than antennomere VIII, rounded at apex; antennomeres pubescent. Pronotum transverse, slightly wider than head, anterior and posterior margin almost straight and bordered, sides straight and bordered, corners obtuse, pronotum disc smooth. Scutellum robust, triangular­shaped. Elytra short which reveals various abdominal segments, at humeri wider than pronotum and as wide as pronotum at the center and apex, humeral zone rather enlarged, posteriorly parallel­sided, rounded apexes, suture robust and evident, surface completely smooth without pubescence and punctation. Metathoracic wings transparent, longer than elytra, almost reaching the apex od abdomen. Sternum elongated, subrectangular, with posterior margin slightly rounded, sternites very wide and transverse, last tergite robust with forked apex equipped with incised concavity at the center, last sternite initially robust and globular and apically forked with short and robust lobes slightly incised at the center. Legs robust, very short, pubescent with several short setae; coxae very massive and rounded; trochanters elongated and curved, with rounded apex; femora at anterior margin straight and posterior margin curved, more robust than tibiae, slightly flat; tibiae cylindrical, thin, with a spur near the apex, longer than femora, metatibiae slightly curved. Tarsal formula 5­ 5­ 5; first tarsomere long, about 1.8 times longer than second; third tarsomere short, triangular and slightly bilobed at sides; fourth globular and bilobed; fifth tarsomere elongated, thin, flat and curved; claws apparently simple without tooth.</paragraph>
</subSubSection>
<caption id="DF0C34B5FF8FFFC0FF0DBB565ED6A7EB" pageId="42" pageNumber="373" startId="42.[151,190,1636,1659]" targetBox="[151,1171,383,1608]" targetPageId="42" targetType="figure">
<paragraph id="8BCC643DFF8FFFC0FF0DBB565ED6A7EB" blockId="42.[151,1172,1636,1691]" pageId="42" pageNumber="373">
Fig. 23.
<taxonomicName id="4C731FBEFF8FFFC0FF75BB565EA9A70B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[239,495,1636,1659]" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="373" phylum="Arthropoda" rank="species" species="burmiticus" status="gen. et sp. nov.">
<emphasis id="B907B82FFF8FFFC0FF75BB565EA9A70B" box="[239,495,1636,1659]" italics="true" pageId="42" pageNumber="373">Vitalfranzius burmiticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF8FFFC0FE6EBB565DD4A70B" box="[500,658,1636,1659]" pageId="42" pageNumber="373" rank="species">gen. et sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FF8EFFC1FFEBB81F5EBFA4CE" pageId="43" pageNumber="374" type="etymology">
<paragraph id="8BCC643DFF8EFFC1FFEBB81F5D16A4F6" blockId="43.[113,605,1325,1414]" pageId="43" pageNumber="374">
<emphasis id="B907B82FFF8EFFC1FFEBB81F5FB4A435" bold="true" box="[113,242,1325,1349]" pageId="43" pageNumber="374">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C731FBEFF8EFFC1FE49B81C5D17A435" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[467,593,1326,1349]" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="374" phylum="Arthropoda" rank="species" species="burmiticus">
<emphasis id="B907B82FFF8EFFC1FE45B81C5D17A435" box="[479,593,1326,1349]" italics="true" pageId="43" pageNumber="374">burmiticus</emphasis>
</taxonomicName>
” derives from the occurrence of this fossil in “burmite”, another name for Burmese amber.
</paragraph>
<paragraph id="8BCC643DFF8EFFC1FFEBB8955EBFA4CE" blockId="43.[113,505,1447,1471]" box="[113,505,1447,1471]" pageId="43" pageNumber="374">
<emphasis id="B907B82FFF8EFFC1FFEBB8955E52A4CF" bold="true" box="[113,276,1447,1471]" pageId="43" pageNumber="374">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8EFFC1FFEBB8EC5FA4A7EA" pageId="43" pageNumber="374" type="discussion">
<paragraph id="8BCC643DFF8EFFC1FFEBB8EC5FA4A7EA" blockId="43.[113,605,1502,1690]" pageId="43" pageNumber="374">
<emphasis id="B907B82FFF8EFFC1FFEBB8EC5F9BA486" bold="true" box="[113,221,1502,1526]" pageId="43" pageNumber="374">Remarks.</emphasis>
Piece of clear, golden amber that measures 20x11x
<quantity id="4C8BC9D8FF8EFFC1FF65B8CD5E07A766" box="[255,321,1535,1558]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="43" pageNumber="374" unit="mm" value="4.0">4 mm</quantity>
. Two legs of the inclusion are not present/visible, and its head and pronotum are partially covered and opacized by the amber surface as well as the ventral part (with emulsion).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,149 @@
<document id="F4796C00D1F18C6FA74345D88E31B3F0" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Müller, Patrick" docDate="2022" docId="03DAD52BFF8CFFC3FFEBBE025B57A3C1" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Vitalfranzius Fanti &amp; Müller, 2022, gen. nov." docType="treatment" docVersion="1" lastPageNumber="372" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="372" updateTime="1722690849856" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="499F9E04125EE389BFCE198C71889E9D">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="2A1ADD2BBE61A39FB0E97ECD309C5F12">Müller, Patrick</mods:namePart>
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<mods:date id="BDE2E3E707C23485648B1FD9D5749A3F">2022</mods:date>
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<subSubSection id="C36937B6FF8CFFC3FFEBBE025EFEA238" box="[113,440,816,841]" pageId="41" pageNumber="372" type="nomenclature">
<paragraph id="8BCC643DFF8CFFC3FFEBBE025EFEA238" blockId="41.[113,440,816,841]" box="[113,440,816,841]" pageId="41" pageNumber="372">
<heading id="D084D351FF8CFFC3FFEBBE025EFEA238" bold="true" box="[113,440,816,841]" fontSize="10" level="2" pageId="41" pageNumber="372" reason="4">
<emphasis id="B907B82FFF8CFFC3FFEBBE025EFEA238" bold="true" box="[113,440,816,841]" pageId="41" pageNumber="372">
Genus
<taxonomicName id="4C731FBEFF8CFFC3FF5ABE035E08A239" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[192,334,817,841]" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B907B82FFF8CFFC3FF5ABE035E08A239" bold="true" box="[192,334,817,841]" italics="true" pageId="41" pageNumber="372">Vitalfranzius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF8CFFC3FECFBE025EFEA238" box="[341,440,816,840]" pageId="41" pageNumber="372" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8CFFC3FFEBBE555EBAA2CF" pageId="41" pageNumber="372" type="type_taxon">
<paragraph id="8BCC643DFF8CFFC3FFEBBE555EBAA2CF" blockId="41.[113,605,871,959]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FFEBBE555E45A20F" bold="true" box="[113,259,871,895]" pageId="41" pageNumber="372">
<typeStatus id="54C8DA9FFF8CFFC3FFEBBE555FEFA20F" box="[113,169,871,895]" pageId="41" pageNumber="372">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C731FBEFF8CFFC3FE92BE5A5F95A2EF" authority="Fanti &amp; P. Muller" authorityName="Fanti &amp; P. Muller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="species" species="burmiticus" status="sp. nov.">
<emphasis id="B907B82FFF8CFFC3FE92BE5A5D41A20F" box="[264,519,872,895]" italics="true" pageId="41" pageNumber="372">Vitalfranzius burmiticus</emphasis>
Fanti &amp; P. Müller
</taxonomicName>
<taxonomicNameLabel id="A2340554FF8CFFC3FF43BEBA5E6CA2EF" box="[217,298,904,927]" pageId="41" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
, by present designation. The genus currently includes two species.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8CFFC3FFEBBEED5D18A527" pageId="41" pageNumber="372" type="etymology">
<paragraph id="8BCC643DFF8CFFC3FFEBBEED5D18A527" blockId="41.[113,606,991,1111]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FFEBBEED5FB7A287" bold="true" box="[113,241,991,1015]" pageId="41" pageNumber="372">Etymology.</emphasis>
Named in honor of the great friend of the first author, Francesco Vitali (Museum of
<collectingCountry id="F36424ADFF8CFFC3FF11B9125E55A547" box="[139,275,1056,1079]" name="Luxembourg" pageId="41" pageNumber="372">Luxembourg</collectingCountry>
). Derived from his entomological nickname “vitalfranz”. The gender is masculine.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8CFFC3FFEBB9445B03A329" pageId="41" pageNumber="372" type="diagnosis">
<paragraph id="8BCC643DFF8CFFC3FFEBB9445D1AA7E2" blockId="41.[113,605,1142,1682]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FFEBB9445FA0A5FF" bold="true" box="[113,230,1142,1167]" pageId="41" pageNumber="372">Diagnosis.</emphasis>
The maxillary palps 4­ segmented with the last palpomere securiform, the tarsal formula 5­ 5­ 5, a pronotum without modified lateral margin, tarsomeres IIIIV without ventral lobes, smooth elytra, and the absence of a long neck, permits to assign this new genus belonging to the family
<taxonomicName id="4C731FBEFF8CFFC3FF6CB80A5E35A43F" authorityName="Imhoff" authorityYear="1856" box="[246,371,1336,1359]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
and its subfamily
<taxonomicName id="4C731FBEFF8CFFC3FDB0B80A5F8DA400" authorityName="Imhoff" authorityYear="1856" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="subFamily" subFamily="Cantharinae">Cantharinae</taxonomicName>
. The new genus is characterized by smooth and very short elytra that leave various abdominal segments uncovered, the rounded head behind the eyes, short and robust legs, and the last abdominal segment slightly forked apically. Four other genera from Burmese amber have very short elytra:
<taxonomicName id="4C731FBEFF8CFFC3FEE5BB295ED1A722" authority="Fanti &amp; Ellenberger, 2018" authorityName="Fanti &amp; Ellenberger" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8CFFC3FEE5BB295D1AA742" box="[383,604,1563,1586]" italics="true" pageId="41" pageNumber="372">Hukawngichthyurus</emphasis>
<bibRefCitation id="EFE219CCFF8CFFC3FFEBBB095ED1A722" author="Fanti F. &amp; Ellenberger S." box="[113,407,1595,1618]" pageId="41" pageNumber="372" pagination="201 - 204" refId="ref20520" refString="Fanti F., Ellenberger S. 2018. A new fossil genus of soldier beetles (Coleoptera: Cantharidae) from mid-Cretaceous Burmese amber: A probable case of adaptive convergence. Cretaceous Research 92: 201 - 204." type="journal article" year="2018">Fanti &amp; Ellenberger, 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C731FBEFF8CFFC3FE3FBB095E0AA702" authority="Poinar &amp; Fanti, 2016" authorityName="Poinar &amp; Fanti" authorityYear="2016" class="Insecta" family="Cantharidae" genus="Ornatomalthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8CFFC3FE3FBB095D1BA722" box="[421,605,1595,1618]" italics="true" pageId="41" pageNumber="372">Ornatomalthinus</emphasis>
<bibRefCitation id="EFE219CCFF8CFFC3FFEBBB695E0AA702" author="Poinar G. O. Jr. &amp; Fanti F." box="[113,332,1627,1650]" pageId="41" pageNumber="372" pagination="1 - 7" refId="ref20956" refString="Poinar G. O. Jr., Fanti F. 2016. New fossil soldier beetles (Coleoptera: Cantharidae) in Burmese, Baltic and Dominican amber. Palaeodiversity 9: 1 - 7." type="journal article" year="2016">Poinar &amp; Fanti, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C731FBEFF8CFFC3FECEBB695EE7A7E2" authority="Fanti, Damgaard &amp; Ellenberger, 2018" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8CFFC3FECEBB695E9DA702" box="[340,475,1627,1650]" italics="true" pageId="41" pageNumber="372">Sanaungulus</emphasis>
<bibRefCitation id="EFE219CCFF8CFFC3FE7ABB695EE7A7E2" author="Fanti F. &amp; Damgaard A. L. &amp; Ellenberger S." pageId="41" pageNumber="372" pagination="170 - 177" refId="ref20442" refString="Fanti F., Damgaard A. L., Ellenberger S. 2018. Two new genera of Cantharidae from Burmese amber of the Hukawng Valley (Insecta, Coleoptera). Cretaceous Research 86: 170 - 177." type="journal article" year="2018">Fanti, Damgaard &amp; Ellenberger, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C731FBEFF8CFFC3FE47BB495D1AA7E2" authorityName="Yang, Liu &amp; Zhao" authorityYear="2022" box="[477,604,1659,1682]" class="Insecta" family="Cantharidae" genus="Brevipterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8CFFC3FE47BB495D1AA7E2" box="[477,604,1659,1682]" italics="true" pageId="41" pageNumber="372">Brevipterus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BCC643DFF8CFFC3FD19BDA95B28A1C2" blockId="41.[643,1134,156,601]" box="[643,1134,155,178]" pageId="41" pageNumber="372">
Y. Yang, H. Liu &amp; W. Zhao (
<bibRefCitation id="EFE219CCFF8CFFC3FC5DBDA95B28A1C2" author="Poinar G. O. Jr. &amp; Fanti F." box="[967,1134,155,178]" pageId="41" pageNumber="372" pagination="1 - 7" refId="ref20956" refString="Poinar G. O. Jr., Fanti F. 2016. New fossil soldier beetles (Coleoptera: Cantharidae) in Burmese, Baltic and Dominican amber. Palaeodiversity 9: 1 - 7." type="journal article" year="2016">Poinar &amp; Fanti</bibRefCitation>
</paragraph>
<paragraph id="8BCC643DFF8CFFC3FD19BD8E5B03A329" blockId="41.[643,1134,156,601]" pageId="41" pageNumber="372">
2016; Fanti et al. 2018;
<bibRefCitation id="EFE219CCFF8CFFC3FCF0BD8E5B2CA1A3" author="Fanti F. &amp; Ellenberger S." box="[874,1130,188,211]" pageId="41" pageNumber="372" pagination="201 - 204" refId="ref20520" refString="Fanti F., Ellenberger S. 2018. A new fossil genus of soldier beetles (Coleoptera: Cantharidae) from mid-Cretaceous Burmese amber: A probable case of adaptive convergence. Cretaceous Research 92: 201 - 204." type="journal article" year="2018">Fanti &amp; Ellenberger 2018</bibRefCitation>
;
<bibRefCitation id="EFE219CCFF8CFFC3FD19BDEE5C7AA183" author="Zhao W. &amp; Liu H. &amp; Geiser M. F. &amp; Yang Y." box="[643,828,220,243]" pageId="41" pageNumber="372" pagination="608 - 621" refId="ref21441" refString="Zhao W., Liu H., Geiser M. F., Yang Y. 2022. Morphology and geometric morphometrics unveil a new genus of Cantharidae (Coleoptera, Elateroidea) from mid-Cretaceous Burmese amber, with a preliminary investigation on the phylogenetic position. Invertebrate Systematic 36 (7): 608 - 621 + supplementary material." type="journal article" year="2022">Zhao et al. 2022</bibRefCitation>
). The most similar is
<taxonomicName id="4C731FBEFF8CFFC3FBB7BDEE5C47A064" authorityName="Poinar &amp; Fanti" authorityYear="2016" class="Insecta" family="Cantharidae" genus="Ornatomalthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8CFFC3FBB7BDEE5C47A064" italics="true" pageId="41" pageNumber="372">Ornatomalthinus</emphasis>
</taxonomicName>
which has a similar rounded head, but the new genus differs for the smooth elytra (with striations provided of relief points in
<taxonomicName id="4C731FBEFF8CFFC3FBB5BC0C5C47A006" authorityName="Poinar &amp; Fanti" authorityYear="2016" class="Insecta" family="Cantharidae" genus="Ornatomalthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8CFFC3FBB5BC0C5C47A006" italics="true" pageId="41" pageNumber="372">Ornatomalthinus</emphasis>
</taxonomicName>
), the smooth head and pronotum (wrinkled and rugose in
<taxonomicName id="4C731FBEFF8CFFC3FCE5BC4D5B72A0E6" authorityName="Poinar &amp; Fanti" authorityYear="2016" box="[895,1076,383,406]" class="Insecta" family="Cantharidae" genus="Ornatomalthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8CFFC3FCE5BC4D5B72A0E6" box="[895,1076,383,406]" italics="true" pageId="41" pageNumber="372">Ornatomalthinus</emphasis>
</taxonomicName>
), and particularly by the last abdominal segment forked (rounded in
<taxonomicName id="4C731FBEFF8CFFC3FC95BCF25C8FA0A7" authorityName="Poinar &amp; Fanti" authorityYear="2016" box="[783,969,448,471]" class="Insecta" family="Cantharidae" genus="Ornatomalthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8CFFC3FC95BCF25C8FA0A7" box="[783,969,448,471]" italics="true" pageId="41" pageNumber="372">Ornatomalthinus</emphasis>
</taxonomicName>
).
<taxonomicName id="4C731FBEFF8CFFC3FC79BCF25B28A0A7" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[995,1134,448,471]" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B907B82FFF8CFFC3FC79BCF25B28A0A7" box="[995,1134,448,471]" italics="true" pageId="41" pageNumber="372">Vitalfranzius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF8CFFC3FD19BCD25DA5A087" box="[643,739,480,503]" pageId="41" pageNumber="372" rank="genus">gen. nov.</taxonomicNameLabel>
is also similar to
<taxonomicName id="4C731FBEFF8CFFC3FC04BCD35B5CA088" authorityName="Yang, Liu &amp; Zhao" authorityYear="2022" box="[926,1050,481,504]" class="Insecta" family="Cantharidae" genus="Brevipterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8CFFC3FC04BCD35B5CA088" box="[926,1050,481,504]" italics="true" pageId="41" pageNumber="372">Brevipterus</emphasis>
</taxonomicName>
but differs for the head strongly rounded and shorter, the pronotum not subreniform, elytra completely smooth and for the antennae not pectinate.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8CFFC3FD19BF4B5B57A3C1" pageId="41" pageNumber="372" type="distribution">
<paragraph id="8BCC643DFF8CFFC3FD19BF4B5B57A3C1" blockId="41.[643,1134,633,689]" pageId="41" pageNumber="372">
<emphasis id="B907B82FFF8CFFC3FD19BF4B5C53A3E1" bold="true" box="[643,789,633,657]" pageId="41" pageNumber="372">Distribution.</emphasis>
Currently known only from the Cretaceous Burmese (
<collectingRegion id="49B7AADFFF8CFFC3FCF1BFA85CFDA3C1" box="[875,955,666,689]" country="Myanmar" name="Kachin" pageId="41" pageNumber="372">Kachin</collectingRegion>
) amber.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,167 @@
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<emphasis id="B907B82FFF8EFFC1FD19BDA95C39A1C3" bold="true" box="[643,895,155,179]" italics="true" pageId="43" pageNumber="374">Vitalfranzius cretaceus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF8EFFC1FC1FBDA95C9CA1C3" box="[901,986,155,179]" pageId="43" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C36937B6FF8EFFC1FD19BD8E5DA6A1A3" box="[643,736,188,211]" pageId="43" pageNumber="374" type="description">
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(
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)
</paragraph>
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<subSubSection id="C36937B6FF8EFFC1FD19BDC15CE4A34C" pageId="43" pageNumber="374" type="materials_examined">
<paragraph id="8BCC643DFF8EFFC1FD19BDC15B1EA0FC" blockId="43.[643,1134,243,396]" pageId="43" pageNumber="374">
<materialsCitation id="3B1B6E60FF8EFFC1FD19BDC15B1EA0FC" collectingDate="2021-12" collectionCode="BUB" county="Sex" location="Catalog" municipality="Burmese" pageId="43" pageNumber="374" specimenCode="BUB4236" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFF8EFFC1FD19BDC15DB7A07B" bold="true" box="[643,753,243,267]" pageId="43" pageNumber="374">
<typeStatus id="54C8DA9FFF8EFFC1FD19BDC15DABA07B" box="[643,749,243,267]" pageId="43" pageNumber="374" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FF8EFFC1FD62BDC15C66A07A" box="[760,800,243,266]" pageId="43" pageNumber="374">Sex</collectingCounty>
undefined, adult specimen in a
<collectingMunicipality id="6BA8FE47FF8EFFC1FD19BC265D99A05B" box="[643,735,276,299]" pageId="43" pageNumber="374">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFF8EFFC1FD70BC265C7EA05B" box="[746,824,276,299]" country="Myanmar" name="Kachin" pageId="43" pageNumber="374">Kachin</collectingRegion>
) amber piece:
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number SNSB­BSPG
<date id="FFCD42FDFF8EFFC1FC81BC065C39A03B" box="[795,895,308,331]" pageId="43" pageNumber="374" value="2021-12">
<collectingDate id="EF89BB15FF8EFFC1FC81BC065C39A03B" box="[795,895,308,331]" pageId="43" pageNumber="374" value="2021-12">2021 XII</collectingDate>
</date>
<quantity id="4C8BC9D8FF8EFFC1FC13BC065C85A03B" box="[905,963,308,331]" metricMagnitude="0" metricUnit="m" metricValue="1.6764" pageId="43" pageNumber="374" unit="in" value="66.0">66 in</quantity>
the
<location id="8EAC32E6FF8EFFC1FC6DBC065C6AA01C" LSID="urn:lsid:plazi:treatment:03DAD52BFF8EFFC7FD19BDA95D56A5B6:8EAC32E6FF8EFFC1FC6DBC065C6AA01C" county="Sex" municipality="Burmese" name="Bayerische Staatssammlung" pageId="43" pageNumber="374" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FF8EFFC1FCC8BC675CA6A01C" LSID="urn:lsid:plazi:treatment:03DAD52BFF8EFFC7FD19BDA95D56A5B6:8EAC32E6FF8EFFC1FCC8BC675CA6A01C" box="[850,992,341,364]" county="Sex" municipality="Burmese" name="Palaontologie" pageId="43" pageNumber="374" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FF8EFFC1FB95BC675B28A01C" LSID="urn:lsid:plazi:treatment:03DAD52BFF8EFFC7FD19BDA95D56A5B6:8EAC32E6FF8EFFC1FB95BC675B28A01C" box="[1039,1134,341,364]" county="Sex" municipality="Burmese" name="Geologie" pageId="43" pageNumber="374" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FF8EFFC1FD31BC475C41A0FC" LSID="urn:lsid:plazi:treatment:03DAD52BFF8EFFC7FD19BDA95D56A5B6:8EAC32E6FF8EFFC1FD31BC475C41A0FC" box="[683,775,373,396]" county="Sex" municipality="Burmese" name="Muller's" pageId="43" pageNumber="374" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FF8EFFC1FC41BC475B0BA0FC" box="[987,1101,373,396]" collectionCode="BUB" pageId="43" pageNumber="374">BUB4236</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFF8EFFC1FD19BC9E5C93A094" blockId="43.[643,1134,428,484]" pageId="43" pageNumber="374">
<emphasis id="B907B82FFF8EFFC1FD19BC9E5C5AA0B4" bold="true" box="[643,796,428,452]" pageId="43" pageNumber="374">Type locality.</emphasis>
<collectingCountry id="F36424ADFF8EFFC1FCBEBC9E5CD6A0B3" box="[804,912,428,451]" name="Myanmar" pageId="43" pageNumber="374">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFF8EFFC1FC01BC9E5B63A0B3" box="[923,1061,428,451]" country="Myanmar" name="Kachin" pageId="43" pageNumber="374">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFF8EFFC1FD19BF365CE4A34C" blockId="43.[643,1134,516,572]" pageId="43" pageNumber="374">
<emphasis id="B907B82FFF8EFFC1FD19BF365C50A36C" bold="true" box="[643,790,516,540]" pageId="43" pageNumber="374">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8EFFC1FD19BF695D92A207" pageId="43" pageNumber="374" type="diagnosis">
<paragraph id="8BCC643DFF8EFFC1FD19BF695D92A207" blockId="43.[643,1135,603,887]" pageId="43" pageNumber="374">
<emphasis id="B907B82FFF8EFFC1FD19BF695C3AA303" bold="true" box="[643,892,603,627]" pageId="43" pageNumber="374">Differential diagnosis.</emphasis>
Smooth and very short elytra, the rounded head behind the eyes, antennae serrate, and the last abdominal segment slightly forked apically make this new species belong to the genus
<taxonomicName id="4C731FBEFF8EFFC1FCC9BFEC5C9DA385" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[851,987,734,757]" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="374" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B907B82FFF8EFFC1FCC9BFEC5C9DA385" box="[851,987,734,757]" italics="true" pageId="43" pageNumber="374">Vitalfranzius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF8EFFC1FC7ABFEC5B06A385" box="[992,1088,734,757]" pageId="43" pageNumber="374" rank="genus">gen. nov.</taxonomicNameLabel>
The new species is easily distinguishable from other species of
<taxonomicName id="4C731FBEFF8EFFC1FD69BE2D5C3DA246" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[755,891,799,822]" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="374" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFF8EFFC1FD69BE2D5C3DA246" box="[755,891,799,822]" italics="true" pageId="43" pageNumber="374">Vitalfranzius</emphasis>
</taxonomicName>
by its serrate antennae and a rounded pronotum at the sides and anterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF8EFFC7FD19BEA55E40A3CB" lastPageId="45" lastPageNumber="376" pageId="43" pageNumber="374" type="description">
<paragraph id="8BCC643DFF8EFFC1FD19BEA55B29A29F" blockId="43.[643,1135,919,1007]" pageId="43" pageNumber="374">
<emphasis id="B907B82FFF8EFFC1FD19BEA55C4BA2DF" bold="true" box="[643,781,919,943]" pageId="43" pageNumber="374">Description.</emphasis>
Adult, winged, rather robust. Sex undefined. Body length 4.0 + head
<quantity id="4C8BC9D8FF8EFFC1FC75BE8A5B78A2BF" box="[1007,1086,952,975]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="43" pageNumber="374" unit="mm" value="0.8">0.8 mm</quantity>
(detached from the body). Entirely reddish­brown.
</paragraph>
<paragraph id="8BCC643DFF8EFFC7FD19B93D5E40A3CB" blockId="43.[643,1134,1039,1681]" lastBlockId="45.[113,605,155,699]" lastPageId="45" lastPageNumber="376" pageId="43" pageNumber="374">Head strongly rounded, slightly transverse, slightly convex, not constricted (triangular­shaped) behind the eyes, surface smooth. Eyes prominent, large, convex, roundish­oblong, inserted in the upper and lateral part of the head. Mandibles very elongated and robust, falciform with pointed apex, equipped with a robust tooth inserted about at the two third of the mandibles length. Maxillary palps 4­ segmented with the last palpomere elongated, securiform with apex rounded. Labial palps 3­ segmented. Antennae 11­ segmented, serrate, rather robust, very short, millimetrically reaches the humeral zone of the elytra, antennal insertion far from the eyes; scape elongated, robust, club­shaped; antennomere II extremely short, globular; antennomeres IIIXI subequal in length, short, robust, each equipped with a short and robust antennal process slightly elongated with rounded apex, but antennomeres IIIIV slightly longer than the following, and the antennomeres IXXI stouter than previous ones with the lobe more squared­irregulaar apically and wider (also the antennomere VIII has the antennal process squared­irregular apically). Pronotum strongly rounded, slightly wider than head, anterior margin strongly rounded, posterior margin almost straight, sides strongly rounded, anterior corners completely rounded and not visible, posterior corners small and obtuse, pronotum disc smooth. Scutellum distorted, not well visible. Elytra short, at humeri wider than pronotum and narrower than pronotum at the apex, posteriorly slightly dehiscent, rounded apexes, suture robust and evident, surface smooth with very short setae and without punctation. Metathoracic wings transparent, longer than elytra. Sternum elongated, rectangular, with posterior margin irregular. Legs robust, very short, pubescent with several short setae; coxae massive, very elongated, with rounded apex; trochanters elongated, with rounded apex; femora slightly curved, more robust than tibiae, cylindrical; tibiae slightly flat, thin, with a spur near the apex, shorter than femora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 1.5 times longer than second; third tarsomere short, triangular; fourth strongly bilobed; fifth tarsomere elongated, thin, flat and curved; claws simple without tooth.</paragraph>
</subSubSection>
<caption id="DF0C34B5FF89FFC6FF0DBB565ED6A7EB" pageId="44" pageNumber="375" startId="44.[151,190,1636,1659]" targetBox="[151,1172,380,1605]" targetPageId="44" targetType="figure">
<paragraph id="8BCC643DFF89FFC6FF0DBB565ED6A7EB" blockId="44.[151,1172,1636,1691]" pageId="44" pageNumber="375">
Fig. 24.
<taxonomicName id="4C731FBEFF89FFC6FF6ABB565EA2A70B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[240,484,1636,1659]" class="Insecta" family="Cantharidae" genus="Vitalfranzius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="375" phylum="Arthropoda" rank="species" species="cretaceus" status="gen. et sp. nov.">
<emphasis id="B907B82FFF89FFC6FF6ABB565EA2A70B" box="[240,484,1636,1659]" italics="true" pageId="44" pageNumber="375">Vitalfranzius cretaceus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FF89FFC6FE71BB565DCDA70B" box="[491,651,1636,1659]" pageId="44" pageNumber="375" rank="species">gen. et sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FF88FFC7FFEBBFE95EBFA2DB" pageId="45" pageNumber="376" type="etymology">
<paragraph id="8BCC643DFF88FFC7FFEBBFE95FEDA204" blockId="45.[113,605,731,884]" pageId="45" pageNumber="376">
<emphasis id="B907B82FFF88FFC7FFEBBFE95FB4A383" bold="true" box="[113,242,731,755]" pageId="45" pageNumber="376">Etymology.</emphasis>
From the Latin adjective “
<emphasis id="B907B82FFF88FFC7FD87BFE95D75A382" box="[541,563,731,754]" italics="true" pageId="45" pageNumber="376">cr</emphasis>
Á
<emphasis id="B907B82FFF88FFC7FDA5BFE95D00A382" box="[575,582,731,754]" italics="true" pageId="45" pageNumber="376">t</emphasis>
-
<emphasis id="B907B82FFF88FFC7FFEBBFCE5FE4A263" box="[113,162,764,787]" italics="true" pageId="45" pageNumber="376">ceus</emphasis>
” = cretaceous. In reference to the geological period (Cretaceous, derived from the Latin “
<emphasis id="B907B82FFF88FFC7FFE7BE0F5FD5A224" box="[125,147,829,852]" italics="true" pageId="45" pageNumber="376">cr</emphasis>
Á
<emphasis id="B907B82FFF88FFC7FF05BE0F5FF2A224" box="[159,180,829,852]" italics="true" pageId="45" pageNumber="376">ta</emphasis>
” = chalk) in which the amber was deposited.
</paragraph>
<paragraph id="8BCC643DFF88FFC7FFEBBEA65EBFA2DB" blockId="45.[113,505,916,940]" box="[113,505,916,940]" pageId="45" pageNumber="376">
<emphasis id="B907B82FFF88FFC7FFEBBEA65E52A2DC" bold="true" box="[113,276,916,940]" pageId="45" pageNumber="376">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF88FFC7FFEBBEF95D56A5B6" pageId="45" pageNumber="376" type="discussion">
<paragraph id="8BCC643DFF88FFC7FFEBBEF95D56A5B6" blockId="45.[113,605,971,1222]" pageId="45" pageNumber="376">
<emphasis id="B907B82FFF88FFC7FFEBBEF95FA7A293" bold="true" box="[113,225,971,995]" pageId="45" pageNumber="376">Remarks.</emphasis>
Piece of clear, golden amber that measures 20x12x
<quantity id="4C8BC9D8FF88FFC7FEB3BEDE5E20A573" box="[297,358,1004,1027]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="45" pageNumber="376" unit="mm" value="3.0">3 mm</quantity>
. The inclusion lacks the whole abdomen, while the head is present but completely detached from the body. Furthermore, the left antenna is preserved until the ninth antennomere. Some debris and the surface with reddish­black spots could mistakenly suggest the presence of a fronto­clypeal suture.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,177 @@
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<mods:title id="5E67099E8104AEF5D6A18B77EC80C1C1">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="A8697A2540CED202F8E0A19300E975EE">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="BD0108CE936884A30B58F4D6FB55E818">Müller, Patrick</mods:namePart>
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<mods:date id="F623A8C41E32B01F683740D468214361">2022</mods:date>
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<subSubSection id="C36937B6FFA0FFEFFFEBBB6B5EEDA701" box="[113,427,1625,1649]" pageId="5" pageNumber="336" type="nomenclature">
<paragraph id="8BCC643DFFA0FFEFFFEBBB6B5EEDA701" blockId="5.[113,427,1625,1680]" box="[113,427,1625,1649]" pageId="5" pageNumber="336">
<heading id="D084D351FFA0FFEFFFEBBB6B5EEDA701" bold="true" box="[113,427,1625,1649]" fontSize="10" level="2" pageId="5" pageNumber="336" reason="3">
<emphasis id="B907B82FFFA0FFEFFFEBBB6B5EEDA701" bold="true" box="[113,427,1625,1649]" pageId="5" pageNumber="336">
<taxonomicName id="4C731FBEFFA0FFEFFFEBBB6B5E09A701" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[113,335,1625,1649]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="thiloi" status="sp. nov.">
<emphasis id="B907B82FFFA0FFEFFFEBBB6B5E09A701" bold="true" box="[113,335,1625,1649]" italics="true" pageId="5" pageNumber="336">Elektrokleinia thiloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA0FFEFFECFBB6B5EEDA701" box="[341,427,1625,1649]" pageId="5" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA0FFEFFFEBBB4B5F87A7E0" box="[113,193,1657,1680]" pageId="5" pageNumber="336" type="description">
<paragraph id="8BCC643DFFA0FFEFFFEBBB4B5F87A7E0" blockId="5.[113,427,1625,1680]" box="[113,193,1657,1680]" pageId="5" pageNumber="336">
(
<figureCitation id="134878B8FFA0FFEFFFE3BB4B5FFEA7E0" box="[121,184,1657,1680]" captionStart="Fig" captionStartId="6.[151,190,1640,1663]" captionTargetBox="[151,1172,385,1610]" captionTargetId="figure-98@6.[151,1172,385,1610]" captionTargetPageId="6" captionText="Fig. 3. Elektrokleinia thiloi sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view." pageId="5" pageNumber="336">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA0FFEFFD19BDA95CE4A094" pageId="5" pageNumber="336" type="materials_examined">
<paragraph id="8BCC643DFFA0FFEFFD19BDA95B1EA044" blockId="5.[643,1134,155,308]" pageId="5" pageNumber="336">
<materialsCitation id="3B1B6E60FFA0FFEFFD19BDA95B1EA044" collectingDate="2021-12-20" collectionCode="BUB" county="Probable" location="Catalog" municipality="Burmese" pageId="5" pageNumber="336" specimenCode="BUB4392" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFA0FFEFFD19BDA95DB7A1C3" bold="true" box="[643,753,155,179]" pageId="5" pageNumber="336">
<typeStatus id="54C8DA9FFFA0FFEFFD19BDA95DABA1C3" box="[643,749,155,179]" pageId="5" pageNumber="336" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFA0FFEFFD62BDA95C11A1C2" box="[760,855,155,178]" pageId="5" pageNumber="336">Probable</collectingCounty>
male, adult specimen in a
<collectingMunicipality id="6BA8FE47FFA0FFEFFD19BD8E5D99A1A3" box="[643,735,188,211]" pageId="5" pageNumber="336">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFA0FFEFFD70BD8E5C7EA1A3" box="[746,824,188,211]" country="Myanmar" name="Kachin" pageId="5" pageNumber="336">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFA0FFEFFC50BD8E5B5CA1A3" LSID="urn:lsid:plazi:treatment:03DAD52BFFA0FFEDFFEBBB6B5ED9A541:8EAC32E6FFA0FFEFFC50BD8E5B5CA1A3" box="[970,1050,188,211]" county="Probable" municipality="Burmese" name="Catalog" pageId="5" pageNumber="336" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFA0FFEFFC81BDEE5CE2A183" box="[795,932,220,243]" pageId="5" pageNumber="336" value="2021-12-20">
<collectingDate id="EF89BB15FFA0FFEFFC81BDEE5CE2A183" box="[795,932,220,243]" pageId="5" pageNumber="336" value="2021-12-20">
2021 XII
<quantity id="4C8BC9D8FFA0FFEFFC13BDEE5CE2A183" box="[905,932,220,243]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="336" unit="in" value="20.0">20</quantity>
</collectingDate>
</date>
in the
<location id="8EAC32E6FFA0FFEFFC6DBDEE5C6AA064" LSID="urn:lsid:plazi:treatment:03DAD52BFFA0FFEDFFEBBB6B5ED9A541:8EAC32E6FFA0FFEFFC6DBDEE5C6AA064" county="Probable" municipality="Burmese" name="Bayerische Staatssammlung" pageId="5" pageNumber="336" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFA0FFEFFCC8BDCF5CA6A064" LSID="urn:lsid:plazi:treatment:03DAD52BFFA0FFEDFFEBBB6B5ED9A541:8EAC32E6FFA0FFEFFCC8BDCF5CA6A064" box="[850,992,253,276]" county="Probable" municipality="Burmese" name="Palaontologie" pageId="5" pageNumber="336" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFA0FFEFFB95BDCF5B28A064" LSID="urn:lsid:plazi:treatment:03DAD52BFFA0FFEDFFEBBB6B5ED9A541:8EAC32E6FFA0FFEFFB95BDCF5B28A064" box="[1039,1134,253,276]" county="Probable" municipality="Burmese" name="Geologie" pageId="5" pageNumber="336" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFA0FFEFFD31BC2F5C41A044" LSID="urn:lsid:plazi:treatment:03DAD52BFFA0FFEDFFEBBB6B5ED9A541:8EAC32E6FFA0FFEFFD31BC2F5C41A044" box="[683,775,285,308]" county="Probable" municipality="Burmese" name="Muller's" pageId="5" pageNumber="336" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFA0FFEFFC41BC2F5B0AA044" box="[987,1100,285,308]" collectionCode="BUB" pageId="5" pageNumber="336">BUB4392</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFA0FFEFFD19BC665C93A0FC" blockId="5.[643,1134,340,396]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFD19BC665C5AA01C" bold="true" box="[643,796,340,364]" pageId="5" pageNumber="336">Type locality.</emphasis>
<collectingCountry id="F36424ADFFA0FFEFFCBEBC675CD6A01C" box="[804,912,341,364]" name="Myanmar" pageId="5" pageNumber="336">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFA0FFEFFC01BC675B63A01C" box="[923,1061,341,364]" country="Myanmar" name="Kachin" pageId="5" pageNumber="336">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFA0FFEFFD19BC9E5CE4A094" blockId="5.[643,1134,428,484]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFD19BC9E5C50A0B4" bold="true" box="[643,790,428,452]" pageId="5" pageNumber="336">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA0FFEFFD19BF365CCBA2B2" pageId="5" pageNumber="336" type="diagnosis">
<paragraph id="8BCC643DFFA0FFEFFD19BF365CCBA2B2" blockId="5.[643,1134,516,962]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFD19BF365C3DA36C" bold="true" box="[643,891,516,540]" pageId="5" pageNumber="336">Differential diagnosis.</emphasis>
Head restricted behind the eyes, short elytra that do not reach the last abdominal segments and equipped with punctuation gathered in striae, plus a pronotum that is little rounded at the sides, make this new species belong to the genus
<taxonomicName id="4C731FBEFFA0FFEFFCD2BF955C9EA3CE" authorityName="Ellenberger &amp; Fanti" authorityYear="2019" box="[840,984,679,702]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA0FFEFFCD2BF955C9EA3CE" box="[840,984,679,702]" italics="true" pageId="5" pageNumber="336">Elektrokleinia</emphasis>
</taxonomicName>
(
<taxonomicName id="4C731FBEFFA0FFEFFC78BF955B26A3CE" authorityName="Imhoff" authorityYear="1856" box="[994,1120,679,702]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
).
<taxonomicName id="4C731FBEFFA0FFEFFD19BFF55B2BA3AE" authority="Ellenberger &amp; Fanti, 2019" authorityName="Ellenberger &amp; Fanti" authorityYear="2019" box="[643,1133,711,734]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B907B82FFFA0FFEFFD19BFF55C12A3AE" box="[643,852,711,734]" italics="true" pageId="5" pageNumber="336">Elektrokleinia picta</emphasis>
<bibRefCitation id="EFE219CCFFA0FFEFFCC0BFF55B2BA3AE" author="Ellenberger S. &amp; Fanti F." box="[858,1133,711,734]" pageId="5" pageNumber="336" pagination="594 - 600" refId="ref20177" refString="Ellenberger S., Fanti F. 2019. New Cretaceous soldier beetle (Cantharidae) from Burmese amber with preserved coloration on the elytra. Zootaxa 4609 (3): 594 - 600." type="journal article" year="2019">Ellenberger &amp; Fanti, 2019</bibRefCitation>
</taxonomicName>
is the
<taxonomicName id="4C731FBEFFA0FFEFFD52BFDA5C18A38F" authorityName="Ellenberger &amp; Fanti" authorityYear="2019" box="[712,862,744,767]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA0FFEFFD52BFDA5C18A38F" box="[712,862,744,767]" italics="true" pageId="5" pageNumber="336">Elektrokleinia</emphasis>
</taxonomicName>
most similar to
<taxonomicName id="4C731FBEFFA0FFEFFB8DBFDA5C43A250" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="thiloi" status="sp. nov.">
<emphasis id="B907B82FFFA0FFEFFB8DBFDA5C43A250" italics="true" pageId="5" pageNumber="336">Elektrokleinia thiloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA0FFEFFC93BE3A5C10A26F" box="[777,854,776,799]" pageId="5" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
, but differs by its pronotum that is less rounded anteriorly and with a less deep concavity in the first basal third of the sides (
<bibRefCitation id="EFE219CCFFA0FFEFFD13BE585CCBA2F1" author="Ellenberger S. &amp; Fanti F." box="[649,909,874,897]" pageId="5" pageNumber="336" pagination="594 - 600" refId="ref20177" refString="Ellenberger S., Fanti F. 2019. New Cretaceous soldier beetle (Cantharidae) from Burmese amber with preserved coloration on the elytra. Zootaxa 4609 (3): 594 - 600." type="journal article" year="2019">Ellenberger &amp; Fanti 2019</bibRefCitation>
).
<taxonomicName id="4C731FBEFFA0FFEFFC01BE585C90A2D1" authority="(Hsiao, Y. Li, Ren &amp; Pang, 2021)" authorityName="Fanti &amp; Müller" authorityYear="2022" baseAuthorityName="Hsiao, Y. Li, Ren &amp; Pang" baseAuthorityYear="2021" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="panna">
<emphasis id="B907B82FFFA0FFEFFC01BE585B28A2F1" box="[923,1134,874,897]" italics="true" pageId="5" pageNumber="336">Elektrokleinia panna</emphasis>
(Hsiao, Y. Li, Ren &amp; Pang, 2021)
</taxonomicName>
has the pronotum narrowest anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA0FFEDFD19BED05E8CA27D" lastPageId="7" lastPageNumber="338" pageId="5" pageNumber="336" type="description">
<paragraph id="8BCC643DFFA0FFEFFD19BED05B29A54A" blockId="5.[643,1135,994,1082]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFD19BED05C48A28A" bold="true" box="[643,782,994,1018]" pageId="5" pageNumber="336">Description.</emphasis>
Adult, alate, rather robust. Probable male, for the last sternite deeply incised. Entirely brown­testaceous.Body length:
<quantity id="4C8BC9D8FFA0FFEFFB83B9115B2CA54A" box="[1049,1130,1059,1082]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="336" unit="mm" value="5.1">5.1 mm</quantity>
.
</paragraph>
<paragraph id="8BCC643DFFA0FFEDFD19B9685E8CA27D" blockId="5.[643,1135,1114,1690]" lastBlockId="7.[113,606,155,781]" lastPageId="7" lastPageNumber="338" pageId="5" pageNumber="336">Head transverse, triangular behind the eyes, slightly convex dorsally, partially covered by the pronotum, surface smooth. Eyes very prominent, subelliptical, inserted laterally to the head, inter­ocular dorsal distance about 2.5 times greater than eye diameter. Mandibles elongated, falciform. Maxillary palps 4­ segmented, with the first palpomere thin and elongated, second palpomere robust, third palpomere almost triangular, last palpomere securiform. Labial palps 3­ segmented with the last palpomere securiform. Antennae 11­ segmented, filiform, inserted in the upper part of the head and not particularly close to the eyes, short, not reaching the half of elytra; scape club­shaped; antennomeres IIIII subequal, shorter than first; antennomere IV as long as antennomeres IIIII together; antennomeres VX subequal, shorter than antennomere IV; antennomere XI elongate, with rounded apex; all antennomeres with very short setae. Pronotum transverse, wider than head, anterior margin rounded and bordered, sides irregular and slightly bordered and with a concavity in the first basal third, surface flat without pubescence. Scutellum triangular­shaped with rounded apex. Elytra wider than pronotum, parallel­sided, rounded at apex, not covering the last two abdominal segments, surface slightly pubescent presenting coarse punctation gathered in striae. Metathoracic wings transparent, slightly longer than elytra, not surpassing the last abdominal segment. Sternum subquadrate with a rounded posterior margin. Sternites transverse, last two tergites as a caudal appendage, last tergite narrower than penultimate tergite, last sternite with the posterior margin with a deep incision in the middle. Legs robust and with pubescence; coxae short, robust; trochanters elongated with rounded apex; femora almost straight; tibiae thin, elongate, cylindrical and enlarged at the apex, with a robust spur at apex, protibiae as long as profemora, mesotibiae slightly longer than mesofemora, metatibiae longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere about 2.1 times longer than second tarsomere; third tarsomere very slightly shorter than second tarsomere and with a lobe at sides; fourth tarsomere bilobed with the lobes very elongated; fifth tarsomere elongate, thin, slightly enlarged apically; claws simple without tooth at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFA3FFECFF0DBB5A5E59A7EF" pageId="6" pageNumber="337" startId="6.[151,190,1640,1663]" targetBox="[151,1172,385,1610]" targetPageId="6" targetType="figure">
<paragraph id="8BCC643DFFA3FFECFF0DBB5A5E59A7EF" blockId="6.[151,1172,1640,1695]" pageId="6" pageNumber="337">
Fig. 3.
<taxonomicName id="4C731FBEFFA3FFECFF7EBB5A5EFCA70F" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[228,442,1640,1663]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="thiloi" status="sp. nov.">
<emphasis id="B907B82FFFA3FFECFF7EBB5A5EFCA70F" box="[228,442,1640,1663]" italics="true" pageId="6" pageNumber="337">Elektrokleinia thiloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA3FFECFE58BB5A5D53A70F" box="[450,533,1640,1663]" pageId="6" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFA2FFEDFFEBBE1E5EBFA2EA" pageId="7" pageNumber="338" type="etymology">
<paragraph id="8BCC643DFFA2FFEDFFEBBE1E5EA7A213" blockId="7.[113,605,812,867]" pageId="7" pageNumber="338">
<emphasis id="B907B82FFFA2FFEDFFEBBE1E5FB4A234" bold="true" box="[113,242,812,836]" pageId="7" pageNumber="338">Etymology.</emphasis>
Named in honor of Thilo Müller, son of the second author (Patrick).
</paragraph>
<paragraph id="8BCC643DFFA2FFEDFFEBBEB15EBFA2EA" blockId="7.[113,505,899,923]" box="[113,505,899,923]" pageId="7" pageNumber="338">
<emphasis id="B907B82FFFA2FFEDFFEBBEB15E52A2EB" bold="true" box="[113,276,899,923]" pageId="7" pageNumber="338">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA2FFEDFFEBBE8B5ED9A541" pageId="7" pageNumber="338" type="discussion">
<paragraph id="8BCC643DFFA2FFEDFFEBBE8B5ED9A541" blockId="7.[113,605,953,1073]" pageId="7" pageNumber="338">
<emphasis id="B907B82FFFA2FFEDFFEBBE8B5FA7A2A1" bold="true" box="[113,225,953,977]" pageId="7" pageNumber="338">Remarks.</emphasis>
Piece of clear, golden amber that measures 20x11x
<quantity id="4C8BC9D8FFA2FFEDFEB1BEE85E2DA281" box="[299,363,986,1009]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="338" unit="mm" value="5.0">5 mm</quantity>
. The inclusion is complete, except for the left mesothoracic leg preserved until part of the tibia.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,171 @@
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<mods:title id="174A9AEF9B1A3523086B449A0EF4CF0E">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="C475C88C979DADA34C08F6AC629A9AE1">Fanti, Fabrizio</mods:namePart>
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<subSubSection id="C36937B6FFA2FFEDFFEBB9555EA1A50F" box="[113,487,1127,1151]" pageId="7" pageNumber="338" type="nomenclature">
<paragraph id="8BCC643DFFA2FFEDFFEBB9555EA1A50F" blockId="7.[113,487,1127,1182]" box="[113,487,1127,1151]" pageId="7" pageNumber="338">
<heading id="D084D351FFA2FFEDFFEBB9555EA1A50F" bold="true" box="[113,487,1127,1151]" fontSize="10" level="2" pageId="7" pageNumber="338" reason="3">
<emphasis id="B907B82FFFA2FFEDFFEBB9555EA1A50F" bold="true" box="[113,487,1127,1151]" pageId="7" pageNumber="338">
<taxonomicName id="4C731FBEFFA2FFEDFFEBB9555ECAA50F" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[113,396,1127,1151]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="species" species="zahradniki" status="sp. nov.">
<emphasis id="B907B82FFFA2FFEDFFEBB9555ECAA50F" bold="true" box="[113,396,1127,1151]" italics="true" pageId="7" pageNumber="338">Elektrokleinia zahradniki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA2FFEDFE08B9555EA1A50F" box="[402,487,1127,1151]" pageId="7" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA2FFEDFFEBB9B55F87A5EE" box="[113,193,1159,1182]" pageId="7" pageNumber="338" type="description">
<paragraph id="8BCC643DFFA2FFEDFFEBB9B55F87A5EE" blockId="7.[113,487,1127,1182]" box="[113,193,1159,1182]" pageId="7" pageNumber="338">
(
<figureCitation id="134878B8FFA2FFEDFFE3B9B55FFFA5EE" box="[121,185,1159,1182]" captionStart="Fig" captionStartId="8.[151,190,1636,1659]" captionTargetBox="[151,1172,386,1607]" captionTargetId="figure-92@8.[151,1172,382,1607]" captionTargetPageId="8" captionText="Fig. 4. Elektrokleinia zahradniki sp. nov. in Burmese amber. A: Holotype, dorso­lateral view; B: Holotype, dorsal view." pageId="7" pageNumber="338">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA2FFEDFFEBB98F5ED6A772" pageId="7" pageNumber="338" type="materials_examined">
<paragraph id="8BCC643DFFA2FFEDFFEBB98F5D01A425" blockId="7.[113,605,1213,1365]" pageId="7" pageNumber="338">
<materialsCitation id="3B1B6E60FFA2FFEDFFEBB98F5D01A425" collectingDate="2021-12-18" collectionCode="BUB" county="Probable" location="Catalog" municipality="Burmese" pageId="7" pageNumber="338" specimenCode="BUB4327" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFA2FFEDFFEBB98F5FA6A5A5" bold="true" box="[113,224,1213,1237]" pageId="7" pageNumber="338">
<typeStatus id="54C8DA9FFFA2FFEDFFEBB98F5F9DA5A5" box="[113,219,1213,1237]" pageId="7" pageNumber="338" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFA2FFEDFF7DB98C5E00A5A5" box="[231,326,1214,1237]" pageId="7" pageNumber="338">Probable</collectingCounty>
female, adult specimen in a
<collectingMunicipality id="6BA8FE47FFA2FFEDFF1BB9EC5F9DA585" box="[129,219,1246,1269]" pageId="7" pageNumber="338">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFA2FFEDFF7FB9EC5E77A585" box="[229,305,1246,1269]" country="Myanmar" name="Kachin" pageId="7" pageNumber="338">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFA2FFEDFE27B9EC5D4DA585" LSID="urn:lsid:plazi:treatment:03DAD52BFFA2FFE3FFEBB9555FFDA377:8EAC32E6FFA2FFEDFE27B9EC5D4DA585" box="[445,523,1246,1269]" county="Probable" municipality="Burmese" name="Catalog" pageId="7" pageNumber="338" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFA2FFEDFE90B9CC5ED5A465" box="[266,403,1278,1301]" pageId="7" pageNumber="338" value="2021-12-18">
<collectingDate id="EF89BB15FFA2FFEDFE90B9CC5ED5A465" box="[266,403,1278,1301]" pageId="7" pageNumber="338" value="2021-12-18">
2021 XII
<quantity id="4C8BC9D8FFA2FFEDFEE2B9CC5ED5A465" box="[376,403,1278,1301]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="7" pageNumber="338" unit="in" value="18.0">18</quantity>
</collectingDate>
</date>
in the
<location id="8EAC32E6FFA2FFEDFE7CB9CC5E5CA445" LSID="urn:lsid:plazi:treatment:03DAD52BFFA2FFE3FFEBB9555FFDA377:8EAC32E6FFA2FFEDFE7CB9CC5E5CA445" county="Probable" municipality="Burmese" name="Bayerische Staatssammlung" pageId="7" pageNumber="338" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFA2FFEDFEDBB82C5E89A445" LSID="urn:lsid:plazi:treatment:03DAD52BFFA2FFE3FFEBB9555FFDA377:8EAC32E6FFA2FFEDFEDBB82C5E89A445" box="[321,463,1310,1333]" county="Probable" municipality="Burmese" name="Palaontologie" pageId="7" pageNumber="338" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFA2FFEDFE64B82C5D1BA445" LSID="urn:lsid:plazi:treatment:03DAD52BFFA2FFE3FFEBB9555FFDA377:8EAC32E6FFA2FFEDFE64B82C5D1BA445" box="[510,605,1310,1333]" county="Probable" municipality="Burmese" name="Geologie" pageId="7" pageNumber="338" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFA2FFEDFF00B80C5FB0A425" LSID="urn:lsid:plazi:treatment:03DAD52BFFA2FFE3FFEBB9555FFDA377:8EAC32E6FFA2FFEDFF00B80C5FB0A425" box="[154,246,1342,1365]" county="Probable" municipality="Burmese" name="Muller's" pageId="7" pageNumber="338" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFA2FFEDFE50B80C5D7AA425" box="[458,572,1342,1365]" collectionCode="BUB" pageId="7" pageNumber="338">BUB4327</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFA2FFEDFFEBB8465E82A4DB" blockId="7.[113,605,1396,1451]" pageId="7" pageNumber="338">
<emphasis id="B907B82FFFA2FFEDFFEBB8465E4CA4FC" bold="true" box="[113,266,1396,1420]" pageId="7" pageNumber="338">Type locality.</emphasis>
<collectingCountry id="F36424ADFFA2FFEDFE89B8465E39A4FB" box="[275,383,1396,1419]" name="Myanmar" pageId="7" pageNumber="338">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFA2FFEDFE10B8465D52A4FB" box="[394,532,1396,1419]" country="Myanmar" name="Kachin" pageId="7" pageNumber="338">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFA2FFEDFFEBB8F95ED6A772" blockId="7.[113,605,1483,1538]" pageId="7" pageNumber="338">
<emphasis id="B907B82FFFA2FFEDFFEBB8F95E43A493" bold="true" box="[113,261,1483,1507]" pageId="7" pageNumber="338">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA2FFEDFFEBBB135C1FA0E2" pageId="7" pageNumber="338" type="diagnosis">
<paragraph id="8BCC643DFFA2FFEDFFEBBB135C1FA0E2" blockId="7.[113,605,1569,1689]" lastBlockId="7.[643,1134,155,402]" pageId="7" pageNumber="338">
<emphasis id="B907B82FFFA2FFEDFFEBBB135E2CA749" bold="true" box="[113,362,1569,1593]" pageId="7" pageNumber="338">Differential diagnosis.</emphasis>
Head restricted behind the eyes makes this new species belong to the genus
<taxonomicName id="4C731FBEFFA2FFEDFF29BB505E02A709" authorityName="Ellenberger &amp; Fanti" authorityYear="2019" box="[179,324,1634,1657]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA2FFEDFF29BB505E02A709" box="[179,324,1634,1657]" italics="true" pageId="7" pageNumber="338">Elektrokleinia</emphasis>
</taxonomicName>
and not to the similar genus
<taxonomicName id="4C731FBEFFA2FFEDFFEBBBB05FB4A7E9" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[113,242,1666,1689]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA2FFEDFFEBBBB05FB4A7E9" box="[113,242,1666,1689]" italics="true" pageId="7" pageNumber="338">Burmomiles</emphasis>
</taxonomicName>
, which has a head rounded behind the eyes.
<taxonomicName id="4C731FBEFFA2FFEDFD73BDA95CBFA1C2" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[745,1017,155,178]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="species" species="zahradniki" status="sp. nov.">
<emphasis id="B907B82FFFA2FFEDFD73BDA95CBFA1C2" box="[745,1017,155,178]" italics="true" pageId="7" pageNumber="338">Elektrokleinia zahradniki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA2FFEDFB9BBDA95B13A1C2" box="[1025,1109,155,178]" pageId="7" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
is distinguishable from other
<taxonomicName id="4C731FBEFFA2FFEDFC01BD895B76A1A2" authorityName="Ellenberger &amp; Fanti" authorityYear="2019" box="[923,1072,187,210]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA2FFEDFC01BD895B76A1A2" box="[923,1072,187,210]" italics="true" pageId="7" pageNumber="338">Elektrokleinia</emphasis>
</taxonomicName>
by its last maxillary palpomere very elongated and not particularly securiform (strongly securiform in the already known species). It also differs by the antennomeral lengths and pronotal shape, and the sculpture of the elytra is less prominent and much smoother.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA2FFE2FD19BC835B3EA047" lastPageId="8" lastPageNumber="339" pageId="7" pageNumber="338" type="description">
<paragraph id="8BCC643DFFA2FFEDFD19BC835B29A358" blockId="7.[643,1135,433,552]" pageId="7" pageNumber="338">
<emphasis id="B907B82FFFA2FFEDFD19BC835C48A0B9" bold="true" box="[643,782,433,457]" pageId="7" pageNumber="338">Description.</emphasis>
Adult, alate, rather robust. Probable female, for the short antennae. Head and pronotum testaceous­brown, elytra dark brown, antennae and legs brown. Body length:
<quantity id="4C8BC9D8FFA2FFEDFB83BF235B2DA358" box="[1049,1131,529,552]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="7" pageNumber="338" unit="mm" value="5.2">5.2 mm</quantity>
.
</paragraph>
<paragraph id="8BCC643DFFA2FFE2FD19BF7A5B3EA047" blockId="7.[643,1135,584,1686]" lastBlockId="8.[680,1172,156,311]" lastPageId="8" lastPageNumber="339" pageId="7" pageNumber="338">Head slightly transverse, rounded and narrower (restricted) behind the eyes, slightly convex dorsally, partially covered by the pronotum, surface smooth and pubescent equipped with short and not erected setae. Eyes little prominent, subelliptical, inserted laterally to the head. Mandibles elongated, falciform. Maxillary palps 4­ segmented, last palpomere very elongated and not particularly securiform with the external side convex. Labial palps 3­ segmented, with the last palpomere securiform, very curved and with pointed apex.Antennae 11­ segmented, filiform, inserted in frons and far to the eyes, short, reaching the humeral zone of elytra; scape elongated, slightly club­shaped; antennomere II about 1.3­ 1.4 times shorter than first; antennomeres IIIX subequal, robust, enlarged and slightly longer than second antennomere; antennomere XI thinner than previous ones, with squared­irregular apex; all antennomeres with very short setae. Pronotum very slightly transverse, wider than head, anterior margin rounded and very slightly bordered, posterior margin straight and strongly bordered, sides irregular and slightly bordered and enlarged in the first basal third, surface flat without pubescence and with shallow punctaction. Scutellum triangular­shaped, elongated, with rounded apex. Elytra wider than pronotum, parallel­sided with humeral zone enlarged, rounded at apex, not covering the last abdominal segment, surface very slightly pubescent presenting small coarse punctation gathered in striae little noticeable. Metathoracic wings covered by elytra. Sternum subquadrate. Sternites transverse. Legs rather robust, short; coxae short, robust; trochanters elongated with rounded apex; femora almost straight and flat; tibiae thin, elongate, cylindrical and slightly enlarged at the apex, protibiae shorter than profemora, mesotibiae nearly as long as mesofemora, metatibiae longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere elongated; second tarsomere slightly shorter than first; third tarsomere triangular­shaped, fourth tarsomere strongly bilobed with the lobes very elongated; fifth tarsomere elongate, thin, curved; claws simple apparently without tooth at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFADFFE2FF0DBB565ECEA7EB" pageId="8" pageNumber="339" startId="8.[151,190,1636,1659]" targetBox="[151,1172,386,1607]" targetPageId="8" targetType="figure">
<paragraph id="8BCC643DFFADFFE2FF0DBB565ECEA7EB" blockId="8.[151,1172,1636,1691]" pageId="8" pageNumber="339">
Fig. 4.
<taxonomicName id="4C731FBEFFADFFE2FF78BB565EB7A70B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[226,497,1636,1659]" class="Insecta" family="Cantharidae" genus="Elektrokleinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="zahradniki" status="sp. nov.">
<emphasis id="B907B82FFFADFFE2FF78BB565EB7A70B" box="[226,497,1636,1659]" italics="true" pageId="8" pageNumber="339">Elektrokleinia zahradniki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFADFFE2FE6DBB565D0FA70B" box="[503,585,1636,1659]" pageId="8" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorso­lateral view; B: Holotype, dorsal view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFACFFE3FFEBBDA95FFFA03C" pageId="9" pageNumber="340" type="etymology">
<paragraph id="8BCC643DFFACFFE3FFEBBDA95E04A184" blockId="9.[113,605,155,244]" pageId="9" pageNumber="340">
<emphasis id="B907B82FFFACFFE3FFEBBDA95FB4A1C3" bold="true" box="[113,242,155,179]" pageId="9" pageNumber="340">Etymology.</emphasis>
Named in honor of Petr Zahradník (
<collectingCountry id="F36424ADFFACFFE3FFE3BD8E5E6DA1A3" box="[121,299,188,211]" name="Czech Republic" pageId="9" pageNumber="340">Czech Republic</collectingCountry>
), a well­known specialist of the family
<taxonomicName id="4C731FBEFFACFFE3FF79BDEF5E78A184" authorityName="Latreille" authorityYear="1802" box="[227,318,221,244]" class="Insecta" family="Ptinidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="family">Ptinidae</taxonomicName>
.
</paragraph>
<paragraph id="8BCC643DFFACFFE3FFEBBC265FFFA03C" blockId="9.[113,605,276,332]" pageId="9" pageNumber="340">
<emphasis id="B907B82FFFACFFE3FFEBBC265E52A05C" bold="true" box="[113,276,276,300]" pageId="9" pageNumber="340">Syninclusions.</emphasis>
Air bubbles, detritus, little ant beetle.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFACFFE3FFEBBC5E5FFDA377" pageId="9" pageNumber="340" type="discussion">
<paragraph id="8BCC643DFFACFFE3FFEBBC5E5FFDA377" blockId="9.[113,605,364,519]" pageId="9" pageNumber="340">
<emphasis id="B907B82FFFACFFE3FFEBBC5E5F9BA0F4" bold="true" box="[113,221,364,388]" pageId="9" pageNumber="340">Remarks.</emphasis>
Piece of clear, golden amber that measures 17x10x
<quantity id="4C8BC9D8FFACFFE3FF65BCBF5E78A0D4" box="[255,318,397,420]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="340" unit="mm" value="8.0">8 mm</quantity>
. The inclusion is complete. The ventral part is little visible and part of the abdomen is covered by emulsion like white­cotton.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,162 @@
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<mods:title id="2D909D3C8F2BB3EE9C9554019856D3D0">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="74D32CE393B1F595DFB643B25AF08031">Fanti, Fabrizio</mods:namePart>
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<mods:date id="D62D1B98C1D73E882BCCAC44F043038D">2022</mods:date>
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<paragraph id="8BCC643DFFA4FFEBFD19BFD85CF4A272" blockId="1.[643,946,746,801]" box="[643,946,746,770]" pageId="1" pageNumber="332">
<heading id="D084D351FFA4FFEBFD19BFD85CF4A272" bold="true" box="[643,946,746,770]" fontSize="10" level="2" pageId="1" pageNumber="332" reason="4">
<emphasis id="B907B82FFFA4FFEBFD19BFD85CF4A272" bold="true" box="[643,946,746,770]" pageId="1" pageNumber="332">
<taxonomicName id="4C731FBEFFA4FFEBFD19BFD85C10A272" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[643,854,746,770]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="parisii" status="sp. nov.">
<emphasis id="B907B82FFFA4FFEBFD19BFD85C10A272" bold="true" box="[643,854,746,770]" italics="true" pageId="1" pageNumber="332">Burmomiles parisii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA4FFEBFCC7BFD85CF4A272" box="[861,946,746,770]" pageId="1" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA4FFEBFD19BE385D95A251" box="[643,723,778,801]" pageId="1" pageNumber="332" type="description">
<paragraph id="8BCC643DFFA4FFEBFD19BE385D95A251" blockId="1.[643,946,746,801]" box="[643,723,778,801]" pageId="1" pageNumber="332">
(
<figureCitation id="134878B8FFA4FFEBFD11BE385D8EA251" box="[651,712,778,801]" captionStart="Fig" captionStartId="2.[151,190,1639,1662]" captionTargetBox="[151,1172,384,1609]" captionTargetId="figure-18@2.[151,1172,384,1609]" captionTargetPageId="2" captionText="Fig. 1. Burmomiles parisii sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view." pageId="1" pageNumber="332">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA4FFEBFD19BE725CF1A5F5" pageId="1" pageNumber="332" type="materials_examined">
<paragraph id="8BCC643DFFA4FFEBFD19BE725C46A2A8" blockId="1.[643,1135,832,984]" pageId="1" pageNumber="332">
<materialsCitation id="3B1B6E60FFA4FFEBFD19BE725C46A2A8" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="1" pageNumber="332" specimenCode="BUB4457" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFA4FFEBFD19BE725DA9A228" bold="true" box="[643,751,832,856]" pageId="1" pageNumber="332">
<typeStatus id="54C8DA9FFFA4FFEBFD19BE725DACA228" box="[643,746,832,856]" pageId="1" pageNumber="332" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFA4FFEBFD69BE735C04A228" box="[755,834,833,856]" pageId="1" pageNumber="332">Female</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFA4FFEBFB8BBE735B28A228" box="[1041,1134,833,856]" pageId="1" pageNumber="332">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFA4FFEBFD10BE535D9FA208" box="[650,729,865,888]" country="Myanmar" name="Kachin" pageId="1" pageNumber="332">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFA4FFEBFCF4BE535C86A208" LSID="urn:lsid:plazi:treatment:03DAD52BFFA4FFE9FD19BFD85C59A062:8EAC32E6FFA4FFEBFCF4BE535C86A208" box="[878,960,865,888]" county="Female" municipality="Burmese" name="Catalog" pageId="1" pageNumber="332" stateProvince="Kachin">Catalog</location>
number
<quantity id="4C8BC9D8FFA4FFEBFB83BE535D8BA2E8" metricMagnitude="2" metricUnit="m" metricValue="1.525016" pageId="1" pageNumber="332" unit="in" value="6004.0">SNHM­6004 in</quantity>
the
<location id="8EAC32E6FFA4FFEBFD69BEB35C5DA2C8" LSID="urn:lsid:plazi:treatment:03DAD52BFFA4FFE9FD19BFD85C59A062:8EAC32E6FFA4FFEBFD69BEB35C5DA2C8" county="Female" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="1" pageNumber="332" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FFA4FFEBFCDDBE935CE4A2C8" LSID="urn:lsid:plazi:treatment:03DAD52BFFA4FFE9FD19BFD85C59A062:8EAC32E6FFA4FFEBFCDDBE935CE4A2C8" box="[839,930,929,952]" county="Female" municipality="Burmese" name="Muller's" pageId="1" pageNumber="332" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFA4FFEBFD19BEF35DB2A2A8" box="[643,756,961,984]" collectionCode="BUB" pageId="1" pageNumber="332">BUB4457</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFA4FFEBFD19BEC55C93A55E" blockId="1.[643,1134,1015,1070]" pageId="1" pageNumber="332">
<emphasis id="B907B82FFFA4FFEBFD19BEC55C5AA57F" bold="true" box="[643,796,1015,1039]" pageId="1" pageNumber="332">Type locality.</emphasis>
<collectingCountry id="F36424ADFFA4FFEBFCBEBEC55CD6A57E" box="[804,912,1015,1038]" name="Myanmar" pageId="1" pageNumber="332">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFA4FFEBFC01BEC55B63A57E" box="[923,1061,1015,1038]" country="Myanmar" name="Kachin" pageId="1" pageNumber="332">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFA4FFEBFD19B97F5CF1A5F5" blockId="1.[643,1134,1101,1157]" pageId="1" pageNumber="332">
<emphasis id="B907B82FFFA4FFEBFD19B97F5C5BA515" bold="true" box="[643,797,1101,1125]" pageId="1" pageNumber="332">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA4FFEBFD19B9965C45A7EC" pageId="1" pageNumber="332" type="diagnosis">
<paragraph id="8BCC643DFFA4FFEBFD19B9965C45A7EC" blockId="1.[643,1137,1188,1692]" pageId="1" pageNumber="332">
<emphasis id="B907B82FFFA4FFEBFD19B9965C3FA5CC" bold="true" box="[643,889,1188,1212]" pageId="1" pageNumber="332">Differential diagnosis.</emphasis>
The genus
<taxonomicName id="4C731FBEFFA4FFEBFC74B9975B28A5CC" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[1006,1134,1189,1212]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA4FFEBFC74B9975B28A5CC" box="[1006,1134,1189,1212]" italics="true" pageId="1" pageNumber="332">Burmomiles</emphasis>
</taxonomicName>
is characterized by short elytra that do not reach the last abdominal segments, tarsal formula 5­ 5­ 5, elytra with shallow but dense punctuation arranged in striae, a pronotum enlarged and rounded at sides, the head rounded, and the unequal maxillary palpomeres with the last segment securiform (Fanti et al. 2018), all features present in the new species described here. The new taxon differs from other
<taxonomicName id="4C731FBEFFA4FFEBFC2BB8F75B77A4AC" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[945,1073,1477,1500]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA4FFEBFC2BB8F75B77A4AC" box="[945,1073,1477,1500]" italics="true" pageId="1" pageNumber="332">Burmomiles</emphasis>
</taxonomicName>
by its much more transverse pronotum. Furthermore, the other
<taxonomicName id="4C731FBEFFA4FFEBFD7FBB375C20A76C" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[741,870,1541,1564]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA4FFEBFD7FBB375C20A76C" box="[741,870,1541,1564]" italics="true" pageId="1" pageNumber="332">Burmomiles</emphasis>
</taxonomicName>
have pectinate antennae.
<taxonomicName id="4C731FBEFFA4FFEBFD19BB175B28A74C" authority="Yang, Geiser &amp; Liu, 2021" authorityName="Yang, Geiser &amp; Liu" authorityYear="2021" box="[643,1134,1573,1596]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B907B82FFFA4FFEBFD19BB175C24A74C" box="[643,866,1573,1596]" italics="true" pageId="1" pageNumber="332">Burmomiles laticollis</emphasis>
<bibRefCitation id="EFE219CCFFA4FFEBFCFCBB175B28A74C" author="Yang Y. &amp; Zhao W. &amp; Geiser M. F. &amp; Zhang W. &amp; Bai M. &amp; Liu H." box="[870,1134,1573,1596]" pageId="1" pageNumber="332" refId="ref21251" refString="Yang Y., Zhao W., Geiser M. F., Zhang W., Bai M., Liu H. 2021. Review of the genus Burmomiles Fanti, Damgaard &amp; Ellenberger from mid-Cretaceous Burmese amber (Coleoptera, Cantharidae). Cretaceous Research 128: 104961 [9 pp.]." type="book" year="2021">Yang, Geiser &amp; Liu, 2021</bibRefCitation>
</taxonomicName>
has a similar pronotum, but it has shorter elytra (less punctated) and pectinate antennae (
<bibRefCitation id="EFE219CCFFA4FFEBFBAFBB575DBEA7EC" author="Yang Y. &amp; Zhao W. &amp; Geiser M. F. &amp; Zhang W. &amp; Bai M. &amp; Liu H." pageId="1" pageNumber="332" refId="ref21251" refString="Yang Y., Zhao W., Geiser M. F., Zhang W., Bai M., Liu H. 2021. Review of the genus Burmomiles Fanti, Damgaard &amp; Ellenberger from mid-Cretaceous Burmese amber (Coleoptera, Cantharidae). Cretaceous Research 128: 104961 [9 pp.]." type="book" year="2021">Yang et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA7FFE9FF0DBDA95E55A4B7" lastPageId="3" lastPageNumber="334" pageId="2" pageNumber="333" type="description">
<paragraph id="8BCC643DFFA7FFE8FF0DBDA95D24A184" blockId="2.[151,644,155,244]" pageId="2" pageNumber="333">
<emphasis id="B907B82FFFA7FFE8FF0DBDA95E65A1C3" bold="true" box="[151,291,155,179]" pageId="2" pageNumber="333">Description.</emphasis>
Adult, alate, robust. Female, for the shape of last sternite and short antennae. Entirely dark brown. Body length: 3.0 mm.
</paragraph>
<paragraph id="8BCC643DFFA7FFE9FF0DBC265E55A4B7" blockId="2.[151,643,276,332]" lastBlockId="3.[113,606,155,1479]" lastPageId="3" lastPageNumber="334" pageId="2" pageNumber="333">Head rounded behind the eyes, partially covered by the pronotum, surface with shallow punctation. Eyes prominent, subelliptical, inserted laterally to the head. Mandibles falciform. Maxillary palps 4­ segmented, with the last palpomere securiform. Labial palps 3­ segmented with the last palpomere securiform. Antennae 11­ segmented, filiform, inserted in the upper part of the head and not particularly close to the eyes, short, slightly surpassing the humeral zone of the elytra; scape club­shaped, very robust; antennomere II short, very wide and globular; antennomere III narrower than previous one and globular; antennomeres IVX subequal, robust, swollen, longer than antennomeres IIIII together; antennomere XI elongate, pointed apically; all antennomeres with setae. Pronotum very transverse, wider than head, anterior margin straight and slightly bordered, posterior margin almost straight, sides slightly curved and slightly bordered, surface completely flat with pubescence (just perceptible). Scutellum triangular­shaped with rounded apex. Elytra wider than pronotum, robust and wide, rounded at apex, not covering the last abdominal segment, wider at humeri, surface presenting a coarse and wide punctation gathered in evident striae visible for the entire elytral length. Metathoracic wings covered by elytra, and not visible. Sternum subquadrate with almost straight posterior margin. Sternites transverse and wide, last two tergites as a caudal large appendage, last sternite rounded presenting two slender and divergent lobes (oviduct). Legs very robust and with short and fitted pubescence; coxae short, robust; trochanters elongated with rounded apex; femora slightly curved, sturdier than tibiae; tibiae thin, elongate, cylindrical and slightly flat apically, pro­ and mesotibiae as long as pro­ and mesofemora, metatibiae longer than metafemora, with a robust spur at apex. Tarsal formula 5­ 5­ 5; first tarsomere about 1.3 times longer than second; second tarsomere elongated; third tarsomere triangular, very slightly enlarged at corners; fourth tarsomere flat, wide, very slightly lobated at corners; fifth tarsomere very elongated, thin, slightly curved, slightly enlarged apically; claws simple, very long, apparently without tooth at the base (with the base slightly wider).</paragraph>
</subSubSection>
<caption id="DF0C34B5FFA7FFE8FF0DBB555E59A7EE" pageId="2" pageNumber="333" startId="2.[151,190,1639,1662]" targetBox="[151,1172,384,1609]" targetPageId="2" targetType="figure">
<paragraph id="8BCC643DFFA7FFE8FF0DBB555E59A7EE" blockId="2.[151,1172,1639,1694]" pageId="2" pageNumber="333">
Fig. 1.
<taxonomicName id="4C731FBEFFA7FFE8FF7FBB555EF2A70E" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[229,436,1639,1662]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="333" phylum="Arthropoda" rank="species" species="parisii" status="sp. nov.">
<emphasis id="B907B82FFFA7FFE8FF7FBB555EF2A70E" box="[229,436,1639,1662]" italics="true" pageId="2" pageNumber="333">Burmomiles parisii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA7FFE8FE26BB555D49A70E" box="[444,527,1639,1662]" pageId="2" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFA6FFE9FFEBB8D55E4CA7E7" pageId="3" pageNumber="334" type="etymology">
<paragraph id="8BCC643DFFA6FFE9FFEBB8D55E4DA730" blockId="3.[113,605,1511,1600]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FFEBB8D55FA8A48F" bold="true" box="[113,238,1511,1535]" pageId="3" pageNumber="334">Etymology.</emphasis>
Named in honor of Francesco Parisi (University of Florence,
<collectingCountry id="F36424ADFFA6FFE9FEF4BB3A5EE7A76F" box="[366,417,1544,1567]" name="Italy" pageId="3" pageNumber="334">Italy</collectingCountry>
), friend of the first author (Fanti).
</paragraph>
<paragraph id="8BCC643DFFA6FFE9FFEBBB525E4CA7E7" blockId="3.[113,605,1632,1687]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FFEBBB525E48A708" bold="true" box="[113,270,1632,1656]" pageId="3" pageNumber="334">Syninclusions.</emphasis>
Air bubbles, detritus, Hymenoptera (
<quantity id="4C8BC9D8FFA6FFE9FF3DBBB25FB8A7E7" box="[167,254,1664,1687]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="334" unit="mm" value="0.7">0.7 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA6FFE9FD19BDA85C59A062" pageId="3" pageNumber="334" type="discussion">
<paragraph id="8BCC643DFFA6FFE9FD19BDA85C59A062" blockId="3.[643,1134,154,274]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FD19BDA85DB4A1C2" bold="true" box="[643,754,154,178]" pageId="3" pageNumber="334">Remarks.</emphasis>
Piece of clear, golden amber that measures 11x10x
<quantity id="4C8BC9D8FFA6FFE9FCAFBD895CCEA1A2" box="[821,904,187,210]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="334" unit="mm" value="4.5">4.5 mm</quantity>
. The inclusion is complete, but the pronotum and ventral side are not clearly visible.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,173 @@
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<mods:title id="5B94414E8E0F8680E0DBF8FBF898F627">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="0642BB9B2D07B9275986249E43C5FC38">Fanti, Fabrizio</mods:namePart>
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<paragraph id="8BCC643DFFA6FFE9FD19BC725CE8A028" blockId="3.[643,942,320,375]" box="[643,942,320,344]" pageId="3" pageNumber="334">
<heading id="D084D351FFA6FFE9FD19BC725CE8A028" bold="true" box="[643,942,320,344]" fontSize="10" level="2" pageId="3" pageNumber="334" reason="4">
<emphasis id="B907B82FFFA6FFE9FD19BC725CE8A028" bold="true" box="[643,942,320,344]" pageId="3" pageNumber="334">
<taxonomicName id="4C731FBEFFA6FFE9FD19BC725C17A028" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[643,849,320,344]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="volpei" status="sp. nov.">
<emphasis id="B907B82FFFA6FFE9FD19BC725C17A028" bold="true" box="[643,849,320,344]" italics="true" pageId="3" pageNumber="334">Burmomiles volpei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA6FFE9FCC2BC725CE8A028" box="[856,942,320,344]" pageId="3" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA6FFE9FD19BC525D94A007" box="[643,722,352,375]" pageId="3" pageNumber="334" type="description">
<paragraph id="8BCC643DFFA6FFE9FD19BC525D94A007" blockId="3.[643,942,320,375]" box="[643,722,352,375]" pageId="3" pageNumber="334">
(
<figureCitation id="134878B8FFA6FFE9FD11BC525D8CA007" box="[651,714,352,375]" captionStart="Fig" captionStartId="4.[151,190,1639,1662]" captionTargetBox="[151,1172,397,1610]" captionTargetId="figure-135@4.[151,1172,397,1610]" captionTargetPageId="4" captionText="Fig. 2. Burmomiles volpei sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view." pageId="3" pageNumber="334">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA6FFE9FD19BCA55CE4A3AA" pageId="3" pageNumber="334" type="materials_examined">
<paragraph id="8BCC643DFFA6FFE9FD19BCA55B69A35E" blockId="3.[643,1134,407,558]" pageId="3" pageNumber="334">
<materialsCitation id="3B1B6E60FFA6FFE9FD19BCA55B69A35E" collectingDate="2021-12" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="3" pageNumber="334" specimenCode="BUB4027" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFA6FFE9FD19BCA55DB7A0DF" bold="true" box="[643,753,407,431]" pageId="3" pageNumber="334">
<typeStatus id="54C8DA9FFFA6FFE9FD19BCA55DABA0DF" box="[643,749,407,431]" pageId="3" pageNumber="334" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFA6FFE9FD6DBCA55C76A0DE" box="[759,816,407,430]" pageId="3" pageNumber="334">Male</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFA6FFE9FB95BCA55B28A0DE" box="[1039,1134,407,430]" pageId="3" pageNumber="334">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFA6FFE9FD10BC855D9CA0BE" box="[650,730,439,462]" country="Myanmar" name="Kachin" pageId="3" pageNumber="334">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFA6FFE9FCE9BC855C80A0BE" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FCE9BC855C80A0BE" box="[883,966,439,462]" county="Male" municipality="Burmese" name="Catalog" pageId="3" pageNumber="334" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFA6FFE9FD54BCE55C74A09E" box="[718,818,471,494]" pageId="3" pageNumber="334" value="2021-12">
<collectingDate id="EF89BB15FFA6FFE9FD54BCE55C74A09E" box="[718,818,471,494]" pageId="3" pageNumber="334" value="2021-12">2021 XII</collectingDate>
</date>
<quantity id="4C8BC9D8FFA6FFE9FCA1BCE55C32A09E" box="[827,884,471,494]" metricMagnitude="0" metricUnit="m" metricValue="1.8796" pageId="3" pageNumber="334" unit="in" value="74.0">74 in</quantity>
the
<location id="8EAC32E6FFA6FFE9FC3CBCE55DB4A37E" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FC3CBCE55DB4A37E" county="Male" municipality="Burmese" name="Bayerische Staatssammlung" pageId="3" pageNumber="334" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFA6FFE9FC87BCC55CF6A37E" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FC87BCC55CF6A37E" box="[797,944,503,526]" county="Male" municipality="Burmese" name="Palaontologie" pageId="3" pageNumber="334" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFA6FFE9FC7EBCC55B00A37E" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FC7EBCC55B00A37E" box="[996,1094,503,526]" county="Male" municipality="Burmese" name="Geologie" pageId="3" pageNumber="334" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFA6FFE9FD19BF255D99A35E" LSID="urn:lsid:plazi:treatment:03DAD52BFFA6FFEFFD19BC725FE2A406:8EAC32E6FFA6FFE9FD19BF255D99A35E" box="[643,735,535,558]" county="Male" municipality="Burmese" name="Muller's" pageId="3" pageNumber="334" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFA6FFE9FC29BF255B62A35E" box="[947,1060,535,558]" collectionCode="BUB" pageId="3" pageNumber="334">BUB4027</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFA6FFE9FD19BF7F5C93A3F4" blockId="3.[643,1134,589,644]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FD19BF7F5C5AA315" bold="true" box="[643,796,589,613]" pageId="3" pageNumber="334">Type locality.</emphasis>
<collectingCountry id="F36424ADFFA6FFE9FCBEBF7F5CD6A314" box="[804,912,589,612]" name="Myanmar" pageId="3" pageNumber="334">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFA6FFE9FC01BF7F5B63A314" box="[923,1061,589,612]" country="Myanmar" name="Kachin" pageId="3" pageNumber="334">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFA6FFE9FD19BF915CE4A3AA" blockId="3.[643,1134,675,730]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FD19BF915C50A3CB" bold="true" box="[643,790,675,699]" pageId="3" pageNumber="334">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA6FFE9FD19BFCA5B0DA580" pageId="3" pageNumber="334" type="diagnosis">
<paragraph id="8BCC643DFFA6FFE9FD19BFCA5B0DA580" blockId="3.[643,1134,760,1264]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FD19BFCA5C38A260" bold="true" box="[643,894,760,784]" pageId="3" pageNumber="334">Differential diagnosis.</emphasis>
Head rounded behind the eyes, short elytra that do not reach the last abdominal segments and which are equipped with punctuation gathered in striae, plus a pronotum rounded at the sides, make this new species belong to the genus
<taxonomicName id="4C731FBEFFA6FFE9FC30BEAB5B6DA2C0" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[938,1067,921,944]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA6FFE9FC30BEAB5B6DA2C0" box="[938,1067,921,944]" italics="true" pageId="3" pageNumber="334">Burmomiles</emphasis>
</taxonomicName>
(
<taxonomicName id="4C731FBEFFA6FFE9FBA0BEAB5D98A2A0" authorityName="Imhoff" authorityYear="1856" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
).
<taxonomicName id="4C731FBEFFA6FFE9FD6BBE8B5CFAA2A0" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[753,956,953,976]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="volpei" status="sp. nov.">
<emphasis id="B907B82FFFA6FFE9FD6BBE8B5CFAA2A0" box="[753,956,953,976]" italics="true" pageId="3" pageNumber="334">Burmomiles volpei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA6FFE9FC5CBE8B5B5DA2A0" box="[966,1051,953,976]" pageId="3" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C731FBEFFA6FFE9FD6DBEEB5C9CA560" authority="Fanti, Damgaard &amp; Ellenberger, 2018" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="willerslevorum">
<emphasis id="B907B82FFFA6FFE9FD6DBEEB5B60A280" box="[759,1062,985,1008]" italics="true" pageId="3" pageNumber="334">Burmomiles willerslevorum</emphasis>
<bibRefCitation id="EFE219CCFFA6FFE9FBAABEEB5C9CA560" author="Fanti F. &amp; Damgaard A. L. &amp; Ellenberger S." pageId="3" pageNumber="334" pagination="170 - 177" refId="ref20442" refString="Fanti F., Damgaard A. L., Ellenberger S. 2018. Two new genera of Cantharidae from Burmese amber of the Hukawng Valley (Insecta, Coleoptera). Cretaceous Research 86: 170 - 177." type="journal article" year="2018">Fanti, Damgaard &amp; Ellenberger, 2018</bibRefCitation>
</taxonomicName>
, but it differs based on the slightly longer elytra, and by its pronotum less protruded at the sides and less protruded in the middle of the anterior margin (Fanti et al. 2018). Instead,
<taxonomicName id="4C731FBEFFA6FFE9FC0FB94B5CD6A5C0" authority="Fanti &amp; Damgaard, 2019" authorityName="Fanti &amp; Damgaard" authorityYear="2019" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="blixenae">
<emphasis id="B907B82FFFA6FFE9FC0FB94B5B2BA5E0" box="[917,1133,1145,1168]" italics="true" pageId="3" pageNumber="334">Burmomiles blixenae</emphasis>
<bibRefCitation id="EFE219CCFFA6FFE9FD19B9AB5CD6A5C0" author="Fanti F. &amp; Damgaard A. L." box="[643,912,1177,1200]" pageId="3" pageNumber="334" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti &amp; Damgaard, 2019</bibRefCitation>
</taxonomicName>
has a pronotum that is more bordered and shorter elytra with a different sculpture (
<bibRefCitation id="EFE219CCFFA6FFE9FCADB9EB5B06A580" author="Fanti F. &amp; Damgaard A. L." box="[823,1088,1241,1264]" pageId="3" pageNumber="334" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti &amp; Damgaard 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA6FFEFFD19B83C5E25A542" lastPageId="5" lastPageNumber="336" pageId="3" pageNumber="334" type="description">
<paragraph id="8BCC643DFFA6FFE9FD19B83C5C2CA415" blockId="3.[643,1134,1294,1381]" pageId="3" pageNumber="334">
<emphasis id="B907B82FFFA6FFE9FD19B83C5C48A456" bold="true" box="[643,782,1294,1318]" pageId="3" pageNumber="334">Description.</emphasis>
Adult, alate, rather robust. Male, for the shape of last sternite. Entirely dark brown. Body length:
<quantity id="4C8BC9D8FFA6FFE9FC88B87C5C21A415" box="[786,871,1358,1381]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="334" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
<paragraph id="8BCC643DFFA6FFEFFD19B8B75E25A542" blockId="3.[643,1134,1413,1692]" lastBlockId="5.[113,605,155,1074]" lastPageId="5" lastPageNumber="336" pageId="3" pageNumber="334">Head rounded behind the eyes, partially covered by the pronotum, surface with sparce punctation. Eyes very prominent, rounded, inserted laterally to the head. Mandibles falciform. Maxillary palps 4­ segmented, with the last palpomere securiform. Labial palps 3­ segmented with the last palpomere securiform. Antennae 11­ segmented, pectinate, inserted in the frons, relatively long, surpassing the half of the elytra; scape club­shaped, not particularly robust, slightly enlarged apically; antennomere II very short, globular; antennomere III short, slightly longer than second, triangular­shaped; antennomeres IVIX subequal, robust, swollen, over two times longer than antennomeres IIIII together, each antennomere with a long process inserted near the base; antennomere X leaner than previous ones but subequal in length, with a long process near the base; antennomere XI elongate, sturdier, squadrate apically; all antennomeres with setae.Pronotum transverse, wider than head, anterior margin rounded and slightly bordered, posterior margin almost straight and bordered, sides rather curved, surface almost flat with pubescence. Scutellum triangular­shaped, elongated, with a subtruncate apex. Elytra wider than pronotum, robust, rounded at apex, not covering the last abdominal segment (and very slightly the penultimate), slightly wider at humeri, surface presenting a coarse and very wide punctation gathered in evident striae visible for the entire elytral length. Metathoracic wings slightly infuscate, longer than elytra. Sternum elongate and robust. Sternites transverse, last tergite as a caudal appendage which is relative short and rounded apically, last sternite narrower than last tergite. Legs long and slender with short and fitted pubescence; coxae short, robust; trochanters elongated with rounded apex; femora curved, sturdier than tibiae; tibiae thin, elongated, cylindrical, longer than femora, with a long spur at apex. Tarsal formula 5­ 5­ 5; first tarsomere elongated, about 1.4 times longer than second; second tarsomere elongated (pro­ and mesotarsomere triangular­shaped); third tarsomere triangular, shorter than previous one; fourth tarsomere lobated at corners; fifth tarsomere very elongated, thin, curved, slightly enlarged apically; claws simple, very long, apparently without tooth at the base (with the base not enlarged).</paragraph>
</subSubSection>
<caption id="DF0C34B5FFA1FFEEFF0DBB555E59A7EE" pageId="4" pageNumber="335" startId="4.[151,190,1639,1662]" targetBox="[151,1172,397,1610]" targetPageId="4" targetType="figure">
<paragraph id="8BCC643DFFA1FFEEFF0DBB555E59A7EE" blockId="4.[151,1172,1639,1694]" pageId="4" pageNumber="335">
Fig. 2.
<taxonomicName id="4C731FBEFFA1FFEEFF7CBB5A5EF6A70F" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[230,432,1640,1663]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="volpei" status="sp. nov.">
<emphasis id="B907B82FFFA1FFEEFF7CBB5A5EF6A70F" box="[230,432,1640,1663]" italics="true" pageId="4" pageNumber="335">Burmomiles volpei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA1FFEEFE22BB555D4AA70E" box="[440,524,1639,1662]" pageId="4" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFA0FFEFFFEBB9635EBFA5AF" pageId="5" pageNumber="336" type="etymology">
<paragraph id="8BCC643DFFA0FFEFFFEBB9635FAAA5D8" blockId="5.[113,605,1105,1192]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFFEBB9635FB4A519" bold="true" box="[113,242,1105,1129]" pageId="5" pageNumber="336">Etymology.</emphasis>
Named in honor of Felice Volpe (Montella, Avellino,
<collectingCountry id="F36424ADFFA0FFEFFECEB9435ECFA5F8" box="[340,393,1137,1160]" name="Italy" pageId="5" pageNumber="336">Italy</collectingCountry>
), dear friend of the first author.
</paragraph>
<paragraph id="8BCC643DFFA0FFEFFFEBB9FA5EBFA5AF" blockId="5.[113,505,1224,1248]" box="[113,505,1224,1248]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFFEBB9FA5E52A590" bold="true" box="[113,276,1224,1248]" pageId="5" pageNumber="336">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA0FFEFFFEBB9CC5FE2A406" pageId="5" pageNumber="336" type="discussion">
<paragraph id="8BCC643DFFA0FFEFFFEBB9CC5FE2A406" blockId="5.[113,605,1278,1398]" pageId="5" pageNumber="336">
<emphasis id="B907B82FFFA0FFEFFFEBB9CC5F9BA466" bold="true" box="[113,221,1278,1302]" pageId="5" pageNumber="336">Remarks.</emphasis>
Piece of rounded, clear, golden amber that measures 12x12x
<quantity id="4C8BC9D8FFA0FFEFFEFAB82D5EE7A446" box="[352,417,1311,1334]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="336" unit="mm" value="3.0">3 mm</quantity>
. To the inclusion lacks almost completely the legs of the right side.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,178 @@
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<mods:title id="B03A34EB5CA28F639AEE5FEBCB138C90">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<subSubSection id="C36937B6FFA9FFE6FD32B9E05B30A59A" box="[680,1142,1234,1258]" pageId="12" pageNumber="343" type="nomenclature">
<paragraph id="8BCC643DFFA9FFE6FD32B9E05B30A59A" blockId="12.[680,1142,1234,1290]" box="[680,1142,1234,1258]" pageId="12" pageNumber="343">
<heading id="D084D351FFA9FFE6FD32B9E05B30A59A" bold="true" box="[680,1142,1234,1258]" fontSize="10" level="2" pageId="12" pageNumber="343" reason="4">
<emphasis id="B907B82FFFA9FFE6FD32B9E05B30A59A" bold="true" box="[680,1142,1234,1258]" pageId="12" pageNumber="343">
<taxonomicName id="4C731FBEFFA9FFE6FD32B9E05B5DA59A" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[680,1051,1234,1258]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="343" phylum="Arthropoda" rank="species" species="dominikiweissbachi" status="sp. nov.">
<emphasis id="B907B82FFFA9FFE6FD32B9E05B5DA59A" bold="true" box="[680,1051,1234,1258]" italics="true" pageId="12" pageNumber="343">Sanaungulus dominikiweissbachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA9FFE6FBBAB9E05B30A59A" box="[1056,1142,1234,1258]" pageId="12" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA9FFE6FD32B9C15DBEA47A" box="[680,760,1267,1290]" pageId="12" pageNumber="343" type="description">
<paragraph id="8BCC643DFFA9FFE6FD32B9C15DBEA47A" blockId="12.[680,1142,1234,1290]" box="[680,760,1267,1290]" pageId="12" pageNumber="343">
(
<figureCitation id="134878B8FFA9FFE6FD2AB9C15DB6A47A" box="[688,752,1267,1290]" captionStart="Fig" captionStartId="13.[113,152,793,816]" captionTargetBox="[113,1134,151,763]" captionTargetId="figure-342@13.[113,1134,151,763]" captionTargetPageId="13" captionText="Fig. 7. Sanaungulus dominikiweissbachi sp. nov. in Burmese amber. Holotype and Paratype (a1: dorsal view; b2: ventral view)." pageId="12" pageNumber="343">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA9FFE7FD32B8185DF9A1F0" lastPageId="13" lastPageNumber="344" pageId="12" pageNumber="343" type="materials_examined">
<paragraph id="8BCC643DFFA9FFE6FD32B8185CC2A494" blockId="12.[680,1172,1322,1508]" pageId="12" pageNumber="343">
<materialsCitation id="3B1B6E60FFA9FFE6FD32B8185CC6A494" collectingDate="2021-12" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="12" pageNumber="343" specimenCode="BUB4059" specimenCount="2" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFA9FFE6FD32B8185CEFA432" bold="true" box="[680,937,1322,1346]" pageId="12" pageNumber="343">
<typeStatus id="54C8DA9FFFA9FFE6FD32B8185C4AA432" box="[680,780,1322,1346]" pageId="12" pageNumber="343" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54C8DA9FFFA9FFE6FCA5B8185CE3A432" box="[831,933,1322,1346]" pageId="12" pageNumber="343" type="paratype">Paratype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFA9FFE6FC37B8185CA4A431" box="[941,994,1322,1345]" pageId="12" pageNumber="343">Male</collectingCounty>
and female, adult specimens in a single
<collectingMunicipality id="6BA8FE47FFA9FFE6FC11B8795CAFA412" box="[907,1001,1355,1378]" pageId="12" pageNumber="343">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFA9FFE6FC6FB8795B03A412" box="[1013,1093,1355,1378]" country="Myanmar" name="Kachin" pageId="12" pageNumber="343">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFA9FFE6FD74B8595C07A4F2" LSID="urn:lsid:plazi:treatment:03DAD52BFFA9FFE4FD32B9E05CE1A0E4:8EAC32E6FFA9FFE6FD74B8595C07A4F2" box="[750,833,1387,1410]" county="Male" municipality="Burmese" name="Catalog" pageId="12" pageNumber="343" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFA9FFE6FBA9B8595BD2A4F2" box="[1075,1172,1387,1410]" pageId="12" pageNumber="343" value="2021-12">
<collectingDate id="EF89BB15FFA9FFE6FBA9B8595BD2A4F2" box="[1075,1172,1387,1410]" pageId="12" pageNumber="343" value="2021-12">2021 XII</collectingDate>
</date>
<quantity id="4C8BC9D8FFA9FFE6FD32B8BE5D98A4D3" box="[680,734,1420,1443]" metricMagnitude="0" metricUnit="m" metricValue="1.905" pageId="12" pageNumber="343" unit="in" value="75.0">75 in</quantity>
the
<location id="8EAC32E6FFA9FFE6FC93B8BE5B75A4D3" LSID="urn:lsid:plazi:treatment:03DAD52BFFA9FFE4FD32B9E05CE1A0E4:8EAC32E6FFA9FFE6FC93B8BE5B75A4D3" box="[777,1075,1420,1443]" county="Male" municipality="Burmese" name="Bayerische Staatssammlung" pageId="12" pageNumber="343" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFA9FFE6FBC7B8BE5C4AA4B4" LSID="urn:lsid:plazi:treatment:03DAD52BFFA9FFE4FD32B9E05CE1A0E4:8EAC32E6FFA9FFE6FBC7B8BE5C4AA4B4" county="Male" municipality="Burmese" name="Palaontologie" pageId="12" pageNumber="343" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFA9FFE6FCA4B89F5CD9A4B4" LSID="urn:lsid:plazi:treatment:03DAD52BFFA9FFE4FD32B9E05CE1A0E4:8EAC32E6FFA9FFE6FCA4B89F5CD9A4B4" box="[830,927,1453,1476]" county="Male" municipality="Burmese" name="Geologie" pageId="12" pageNumber="343" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFA9FFE6FC51B89F5B60A4B4" LSID="urn:lsid:plazi:treatment:03DAD52BFFA9FFE4FD32B9E05CE1A0E4:8EAC32E6FFA9FFE6FC51B89F5B60A4B4" box="[971,1062,1453,1476]" county="Male" municipality="Burmese" name="Muller's" pageId="12" pageNumber="343" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFA9FFE6FC92B8FF5C3FA494" box="[776,889,1485,1508]" collectionCode="BUB" pageId="12" pageNumber="343">BUB4059</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCC643DFFA9FFE6FD32BB365CBDA74C" blockId="12.[680,1172,1540,1596]" pageId="12" pageNumber="343">
<emphasis id="B907B82FFFA9FFE6FD32BB365C07A76C" bold="true" box="[680,833,1540,1564]" pageId="12" pageNumber="343">Type locality.</emphasis>
<collectingCountry id="F36424ADFFA9FFE6FCD0BB365CF0A76B" box="[842,950,1540,1563]" name="Myanmar" pageId="12" pageNumber="343">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFA9FFE6FC5BBB365B0DA76B" box="[961,1099,1540,1563]" country="Myanmar" name="Kachin" pageId="12" pageNumber="343">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFA9FFE6FD32BB6E5C81A7E4" blockId="12.[680,1172,1628,1684]" pageId="12" pageNumber="343">
<emphasis id="B907B82FFFA9FFE6FD32BB6E5C7AA704" bold="true" box="[680,828,1628,1652]" pageId="12" pageNumber="343">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
<paragraph id="8BCC643DFFA8FFE7FDBABD5F5DF9A1F0" blockId="13.[544,703,109,128]" box="[544,703,109,128]" pageId="13" pageNumber="344">
<emphasis id="B907B82FFFA8FFE7FDBABD5F5DF9A1F0" box="[544,703,109,128]" italics="true" pageId="13" pageNumber="344">Fanti F., Müller P.</emphasis>
</paragraph>
</subSubSection>
<caption id="DF0C34B5FFA8FFE7FFEBBE2B5EF0A220" pageId="13" pageNumber="344" startId="13.[113,152,793,816]" targetBox="[113,1134,151,763]" targetPageId="13" targetType="figure">
<paragraph id="8BCC643DFFA8FFE7FFEBBE2B5EF0A220" blockId="13.[113,1134,793,848]" pageId="13" pageNumber="344">
Fig. 7.
<taxonomicName id="4C731FBEFFA8FFE7FF26BE285D59A241" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[188,543,794,817]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="dominikiweissbachi" status="sp. nov.">
<emphasis id="B907B82FFFA8FFE7FF26BE285D59A241" box="[188,543,794,817]" italics="true" pageId="13" pageNumber="344">Sanaungulus dominikiweissbachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA8FFE7FDBFBE2B5D31A240" box="[549,631,793,816]" pageId="13" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. Holotype and Paratype (a1: dorsal view; b2: ventral view).
</paragraph>
</caption>
<subSubSection id="C36937B6FFA8FFE7FFEBBEB65E2EA52C" pageId="13" pageNumber="344" type="diagnosis">
<paragraph id="8BCC643DFFA8FFE7FFEBBEB65E2EA52C" blockId="13.[113,605,900,1116]" pageId="13" pageNumber="344">
<emphasis id="B907B82FFFA8FFE7FFEBBEB65E29A2EC" bold="true" box="[113,367,900,924]" pageId="13" pageNumber="344">Differential diagnosis.</emphasis>
The specimen of
<taxonomicName id="4C731FBEFFA8FFE7FDA3BEB75EF4A2CC" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="344" phylum="Arthropoda" rank="species" species="dominikiweissbachi" status="sp. nov.">
<emphasis id="B907B82FFFA8FFE7FDA3BEB75EF4A2CC" italics="true" pageId="13" pageNumber="344">Sanaungulus dominikiweissbachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFA8FFE7FE2DBE975D40A2CC" box="[439,518,933,956]" pageId="13" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
features a male and female, the first Burmese fossil soldier beetles documented preserved during mating. The species is characterized by long (usually only one abdominal segment remains uncovered) and smooth elytra.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA8FFE4FFEBB9485D70A403" lastPageId="14" lastPageNumber="345" pageId="13" pageNumber="344" type="description">
<paragraph id="8BCC643DFFA8FFE7FFEBB9485D61A5E1" blockId="13.[113,551,1146,1170]" box="[113,551,1146,1170]" pageId="13" pageNumber="344">
<emphasis id="B907B82FFFA8FFE7FFEBB9485FBAA5E2" bold="true" box="[113,252,1146,1170]" pageId="13" pageNumber="344">Description.</emphasis>
Male and female in mating.
</paragraph>
<paragraph id="8BCC643DFFA8FFE7FFEBB99D5E29A456" blockId="13.[113,605,1199,1318]" pageId="13" pageNumber="344">
<materialsCitation id="3B1B6E60FFA8FFE7FFEBB99D5E2DA456" collectionCode="MALE" pageId="13" pageNumber="344" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED62FCF8FFA8FFE7FFEBB99D5FFAA5B6" box="[113,188,1199,1222]" pageId="13" pageNumber="344">MALE</collectionCode>
(
<typeStatus id="54C8DA9FFFA8FFE7FF52B99D5E6DA5B6" box="[200,299,1199,1222]" pageId="13" pageNumber="344" type="holotype">Holotype</typeStatus>
A1): adult, winged, slender. Male, defined on the basis of the slender abdomen and visible aedeagus. Body length
<quantity id="4C8BC9D8FFA8FFE7FD9DB9DD5D1CA476" box="[519,602,1263,1286]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="344" unit="mm" value="2.7">2.7 mm</quantity>
. Entirely reddish­brown
</materialsCitation>
.
</paragraph>
<paragraph id="8BCC643DFFA8FFE4FFEBB8765D58A329" blockId="13.[113,605,1348,1691]" lastBlockId="14.[151,643,155,601]" lastPageId="14" lastPageNumber="345" pageId="13" pageNumber="344">Head slightly transverse, constricted (triangular­shaped) behind the eyes, surface with shallow punctation. Eyes prominent, very large, roundish, inserted in the upper and lateral part of the head. Mandibles elongated, falciform. Maxillary palps 4­ segmented with the last palpomere strongly elongated and securiform. Labial palps 3­ segmented, with the last palpomere securiform.Antennae 11­ segmented, pectinate, relatively long, reaching the half of elytra, antennal insertion very far from the eyes; scape strongly elongated, strongly restricted in the middle and globular at apex; antennomeres IIIII short, subequal in length, globular­elongated, antennomere III at apex with a long and robust antennal process; antennomeres IVVII very long, rather robust, each with an antennal process at apex which is very long and emarginated at apex; antennomeres VIIIIX shorter than previous ones, each with the same antennal process at apex; antennomere X as long as the previous ones, without the antennal process; antennomere XI robust, with rounded apex; all the antennomeres slightly pubescent. Pronotum elongated, almost spherical, slightly wider than head, anterior margin rounded, posterior margin almost straight, sides curved with a small and rounded tooth near the middle, corners obtuse, pronotum disc irregular. Scutellum triangular­shaped. Elytra elongated that do not cover the last abdominal segment, wider than pronotum, parallel­sided, rounded apexes, surface completely smooth with pubescence. Metathoracic wings transparent, very slightly longer than elytra. Last tergite elongated. Aedeagus trilobated with the lobes irregular, long and not plate­like. Legs thin, not particularly long, slightly pubescent; coxae massive; trochanters elongated, robust, rounded apex; femora almost straight, cylindrical, very sturdier than tibiae; tibiae, thin, sub­cylindrical, long, with a thin and long spur near the apex, pro­ and mesotibiae longer than pro­ and mesofemora, metatibiae very longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 1.5 times longer than second; third tarsomere short, about 1.5 times shorter than second; fourth strongly bilobed with lobes very long and pointed apically; fifth tarsomere extremely elongated, thin, slightly flat, curved; claws simple, long, pointed, without denticle at the base.</paragraph>
<paragraph id="8BCC643DFFABFFE4FF0DBF4B5DC5A3A1" blockId="14.[151,643,633,721]" pageId="14" pageNumber="345">
FEMALE (
<typeStatus id="54C8DA9FFFABFFE4FE97BF4B5E2DA3E0" box="[269,363,633,656]" pageId="14" pageNumber="345" type="paratype">Paratype</typeStatus>
B2): adult, winged, slen­ der. Body length
<quantity id="4C8BC9D8FFABFFE4FEA4BFA85EC8A3C1" box="[318,398,666,689]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="14" pageNumber="345" unit="mm" value="2.9">2.9 mm</quantity>
.Entirely reddish­brown. Morphologically very similar to the
<typeStatus id="54C8DA9FFFABFFE4FD82BF885D39A3A1" box="[536,639,698,721]" pageId="14" pageNumber="345" type="holotype">Holotype</typeStatus>
.
</paragraph>
<paragraph id="8BCC643DFFABFFE4FF0DBFC05D70A403" blockId="14.[151,644,754,1395]" pageId="14" pageNumber="345">Head transverse, constricted behind the eyes. Eyes prominent and large. Mandibles falciform. Last maxillary palpomere securiform. Antennae 11­ segmented, filiform, rather long, reaching the half of abdomen, antennal insertion very far from the eyes; scape strongly elongated, restricted in the middle and globular at apex; antennomeres IIIII short, subequal in length, globular­elongated; antennomeres IVV long, rather robust; antennomeres VIX slightly shorter and less sturdier than previous ones; antennomere XI filiform, elongated, with rounded apex; all the antennomeres slightly pubescent and without antennal process. Pronotum almost spherical. Elytra elongated that do not cover the last abdominal segment and partially the penultimate, completely smooth. Metathoracic wings transparent, very slightly longer than elytra. Sternum elongated with rounded apex. Sternites transverse and pubescent, last sternite wide.</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFABFFE4FF0DB8A15D59A772" pageId="14" pageNumber="345" type="etymology">
<paragraph id="8BCC643DFFABFFE4FF0DB8A15DC2A4BA" blockId="14.[151,644,1427,1482]" pageId="14" pageNumber="345">
<emphasis id="B907B82FFFABFFE4FF0DB8A15E50A4DB" bold="true" box="[151,278,1427,1451]" pageId="14" pageNumber="345">Etymology.</emphasis>
Named in honor of Dominik Weissbach, dear friend of the second author (Patrick).
</paragraph>
<paragraph id="8BCC643DFFABFFE4FF0DB8D85D59A772" blockId="14.[151,543,1514,1538]" box="[151,543,1514,1538]" pageId="14" pageNumber="345">
<emphasis id="B907B82FFFABFFE4FF0DB8D85E7CA772" bold="true" box="[151,314,1514,1538]" pageId="14" pageNumber="345">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFABFFE4FF0DBB135CE1A0E4" pageId="14" pageNumber="345" type="discussion">
<paragraph id="8BCC643DFFABFFE4FF0DBB135CE1A0E4" blockId="14.[151,643,1569,1690]" lastBlockId="14.[680,1172,155,404]" pageId="14" pageNumber="345">
<emphasis id="B907B82FFFABFFE4FF0DBB135E45A749" bold="true" box="[151,259,1569,1593]" pageId="14" pageNumber="345">Remarks.</emphasis>
Piece of clear, golden amber that measures 18x13x
<quantity id="4C8BC9D8FFABFFE4FEBFBB705E22A729" box="[293,356,1602,1625]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="345" unit="mm" value="3.0">3 mm</quantity>
. The inclusion is complete, but the male (A1) has the left antenna detached approximately from the end of the antennomere VI and the mesothoracic legs detached from the tibia or femur but with all these parts still present and preserved in the amber piece near the body. While, the female (B2) is complete with only the elytra detached and move upwards. It is not possible to take a proper picture of the other side of specimens as there is a resin layer covering most of this part.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,154 @@
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<mods:title id="7C59BD159179249CFC73BB57F20EC48E">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<paragraph id="8BCC643DFFABFFE4FD32BCF85CB4A092" blockId="14.[680,1010,458,514]" box="[680,1010,458,482]" pageId="14" pageNumber="345">
<heading id="D084D351FFABFFE4FD32BCF85CB4A092" bold="true" box="[680,1010,458,482]" fontSize="10" level="2" pageId="14" pageNumber="345" reason="4">
<emphasis id="B907B82FFFABFFE4FD32BCF85CB4A092" bold="true" box="[680,1010,458,482]" pageId="14" pageNumber="345">
<taxonomicName id="4C731FBEFFABFFE4FD32BCF85CD3A092" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[680,917,458,482]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="dunlopi" status="sp. nov.">
<emphasis id="B907B82FFFABFFE4FD32BCF85CD3A092" bold="true" box="[680,917,458,482]" italics="true" pageId="14" pageNumber="345">Sanaungulus dunlopi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFABFFE4FC07BCF85CB4A092" box="[925,1010,458,482]" pageId="14" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFABFFE4FD32BCD95DBEA372" box="[680,760,491,514]" pageId="14" pageNumber="345" type="description">
<paragraph id="8BCC643DFFABFFE4FD32BCD95DBEA372" blockId="14.[680,1010,458,514]" box="[680,760,491,514]" pageId="14" pageNumber="345">
(
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)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFABFFE4FD32BF135C81A218" pageId="14" pageNumber="345" type="materials_examined">
<paragraph id="8BCC643DFFABFFE4FD32BF135B13A3CA" blockId="14.[680,1172,545,698]" pageId="14" pageNumber="345">
<materialsCitation id="3B1B6E60FFABFFE4FD32BF135B13A3CA" collectingDate="2021-12" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="14" pageNumber="345" specimenCode="BUB3886" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFABFFE4FD32BF135C52A349" bold="true" box="[680,788,545,569]" pageId="14" pageNumber="345">
<typeStatus id="54C8DA9FFFABFFE4FD32BF135C49A349" box="[680,783,545,569]" pageId="14" pageNumber="345" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFABFFE4FC82BF105C21A349" box="[792,871,546,569]" pageId="14" pageNumber="345">Female</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFABFFE4FBADBF105BD2A349" box="[1079,1172,546,569]" pageId="14" pageNumber="345">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFABFFE4FD2ABF705C46A329" box="[688,768,578,601]" country="Myanmar" name="Kachin" pageId="14" pageNumber="345">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFABFFE4FC03BF705CAAA329" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FC03BF705CAAA329" box="[921,1004,578,601]" county="Female" municipality="Burmese" name="Catalog" pageId="14" pageNumber="345" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFABFFE4FD6EBF505C1EA309" box="[756,856,610,633]" pageId="14" pageNumber="345" value="2021-12">
<collectingDate id="EF89BB15FFABFFE4FD6EBF505C1EA309" box="[756,856,610,633]" pageId="14" pageNumber="345" value="2021-12">2021 XII</collectingDate>
</date>
<quantity id="4C8BC9D8FFABFFE4FCFBBF505CDFA309" box="[865,921,610,633]" metricMagnitude="0" metricUnit="m" metricValue="1.7271999999999998" pageId="14" pageNumber="345" unit="in" value="68.0">68 in</quantity>
the
<location id="8EAC32E6FFABFFE4FC56BF505C51A3E9" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FC56BF505C51A3E9" county="Female" municipality="Burmese" name="Bayerische Staatssammlung" pageId="14" pageNumber="345" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFABFFE4FCD9BFB05C90A3E9" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FCD9BFB05C90A3E9" box="[835,982,642,665]" county="Female" municipality="Burmese" name="Palaontologie" pageId="14" pageNumber="345" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFABFFE4FB90BFB05B2AA3E9" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FB90BFB05B2AA3E9" box="[1034,1132,642,665]" county="Female" municipality="Burmese" name="Geologie" pageId="14" pageNumber="345" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFABFFE4FD32BF915C42A3CA" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FD32BF915C42A3CA" box="[680,772,675,698]" county="Female" municipality="Burmese" name="Muller's" pageId="14" pageNumber="345" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFABFFE4FC43BF915B0FA3CA" box="[985,1097,675,698]" collectionCode="BUB" pageId="14" pageNumber="345">BUB3886</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFABFFE4FD32BFEB5CBDA261" blockId="14.[680,1172,729,785]" pageId="14" pageNumber="345">
<emphasis id="B907B82FFFABFFE4FD32BFEB5C07A381" bold="true" box="[680,833,729,753]" pageId="14" pageNumber="345">Type locality.</emphasis>
<collectingCountry id="F36424ADFFABFFE4FCD0BFE85CF0A381" box="[842,950,730,753]" name="Myanmar" pageId="14" pageNumber="345">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFABFFE4FC5BBFE85B0DA381" box="[961,1099,730,753]" country="Myanmar" name="Kachin" pageId="14" pageNumber="345">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFABFFE4FD32BE025C81A218" blockId="14.[680,1172,816,872]" pageId="14" pageNumber="345">
<emphasis id="B907B82FFFABFFE4FD32BE025C7AA238" bold="true" box="[680,828,816,840]" pageId="14" pageNumber="345">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFABFFE4FD32BEBA5B77A513" pageId="14" pageNumber="345" type="diagnosis">
<paragraph id="8BCC643DFFABFFE4FD32BEBA5B77A513" blockId="14.[680,1172,904,1123]" pageId="14" pageNumber="345">
<emphasis id="B907B82FFFABFFE4FD32BEBA5CDBA2D0" bold="true" box="[680,925,904,928]" pageId="14" pageNumber="345">Differential diagnosis.</emphasis>
The new species is characterized by an elongated pronotum with a strongly rounded anterior margin, and it is equipped with robust antennomeres. The genus
<taxonomicName id="4C731FBEFFABFFE4FD32B9395C6AA552" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[680,812,1035,1058]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="345" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFABFFE4FD32B9395C6AA552" box="[680,812,1035,1058]" italics="true" pageId="14" pageNumber="345">Sanaungulus</emphasis>
</taxonomicName>
is characterized by very short elytra often with deep punctuation arranged in striae, and by a head restricted behind eyes.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFABFFFAFD32B9B15D52A289" lastPageId="16" lastPageNumber="347" pageId="14" pageNumber="345" type="description">
<paragraph id="8BCC643DFFABFFE4FD32B9B15C65A46C" blockId="14.[680,1172,1155,1308]" pageId="14" pageNumber="345">
<emphasis id="B907B82FFFABFFE4FD32B9B15C72A5EB" bold="true" box="[680,820,1155,1179]" pageId="14" pageNumber="345">Description.</emphasis>
Adult, winged, slender. Female, defined on the shape of the last abdominal segments, and on the basis of short antennae. Body length about
<quantity id="4C8BC9D8FFABFFE4FCABB9D75CC7A58C" box="[817,897,1253,1276]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="345" unit="mm" value="3.2">3.2 mm</quantity>
(the body is curled). Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFABFFFAFD32B80E5D52A289" blockId="14.[680,1172,1340,1689]" lastBlockId="16.[151,644,155,1017]" lastPageId="16" lastPageNumber="347" pageId="14" pageNumber="345">Head short, constricted (triangular­shaped) behind the eyes, smooth, without punctuation, fitted with few long setae. Eyes very prominent, large, roundish, inserted in the lateral part of the head, inter­ocular dorsal distance about 1.3 times greater than eye diameter. Mandibles elongated, falciform. Maxillary palps 4­ segmented, with the last palpomere securiform.Antennae 11­ segmented, filiform, robust, short, slightly surpassing the middle of the elytra; scape very elongated and club­shaped; antennomere II short; antennomeres IIIVIII subequal in length, very robust, without antennal process, enlarged at the external corner which is rounded; antennomere IX approximately as long as previous ones, slightly narrower; antennomere X narrower than previous one, slightly enlarged at the external corner; antennomere XI eleongated, strongly squared apically; antennomeres pubescent. Pronotum very elongated, very slightly narrower than head, anterior margin strongly rounded and bordered, posterior margin rather straight and slightly bordered, corners rather acute, sides straight and slightly bordered, pronotum disc not flat and very slightly wrinkled with shallow punctuation. Scutellum triangular­shaped with pointed apex. Elytra very short which reveals many abdominal segments, wider than pronotum at humery, strongly dehiscent posteriorly, rounded apexes, surface strongly ruguose equipped with several short setae. Metathoracic wings transparent, very short, very slightly surpassing the elytra. Sternum elongated with posterior margin rounded, sternites strongly transverse, robust, equipped with short setae; last tergite short, wide, emarginate at apex; last sternite transverse, robust, short, as long as last tergite, with the posterior margin straight. Legs thin, pro­ and mesolegs short, metalegs elongated and slender, pubescent; coxae massive; trochanters elongated and rounded apically; femora nearly straight, slightly sturdier than tibiae (expecially the metafemora); tibiae cylindrical, thin, without spurs near the apex, shorter than femora. Tarsal formula 5­ 5­ 5; first tarsomere elongated and thin; second tarsomere slightly shorter than first; third tarsomere slightly elongated and triangular­shaped; fourth strongly bilobed with the lobes very long; fifth tarsomere elongated, flat and enlarged apically; claws simple, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFAAFFE5FFEBBB555E79A7EE" pageId="15" pageNumber="346" startId="15.[113,152,1639,1662]" targetBox="[113,1134,385,1610]" targetPageId="15" targetType="figure">
<paragraph id="8BCC643DFFAAFFE5FFEBBB555E79A7EE" blockId="15.[113,1134,1639,1694]" pageId="15" pageNumber="346">
Fig. 8.
<taxonomicName id="4C731FBEFFAAFFE5FF27BB5A5ED9A70F" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[189,415,1640,1663]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="346" phylum="Arthropoda" rank="species" species="dunlopi" status="sp. nov.">
<emphasis id="B907B82FFFAAFFE5FF27BB5A5ED9A70F" box="[189,415,1640,1663]" italics="true" pageId="15" pageNumber="346">Sanaungulus dunlopi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFAAFFE5FE3FBB555EB1A70E" box="[421,503,1639,1662]" pageId="15" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventro­lateral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFB5FFFAFF0DB92A5D59A5D6" pageId="16" pageNumber="347" type="etymology">
<paragraph id="8BCC643DFFB5FFFAFF0DB92A5DC2A500" blockId="16.[151,644,1048,1136]" pageId="16" pageNumber="347">
<emphasis id="B907B82FFFB5FFFAFF0DB92A5E52A540" bold="true" box="[151,276,1048,1072]" pageId="16" pageNumber="347">Etymology.</emphasis>
Named in honor of Jason A. Dunlop (Museum für Naturkunde,
<collectingRegion id="49B7AADFFFB5FFFAFE3FB90B5EAFA520" box="[421,489,1081,1104]" country="Germany" name="Berlin" pageId="16" pageNumber="347">Berlin</collectingRegion>
), a well­known specialist of fossil arachnids and their relatives.
</paragraph>
<paragraph id="8BCC643DFFB5FFFAFF0DB9BD5D59A5D6" blockId="16.[151,543,1167,1191]" box="[151,543,1167,1191]" pageId="16" pageNumber="347">
<emphasis id="B907B82FFFB5FFFAFF0DB9BD5E7CA5D7" bold="true" box="[151,314,1167,1191]" pageId="16" pageNumber="347">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB5FFFAFF0DB9F75D6BA46D" pageId="16" pageNumber="347" type="discussion">
<paragraph id="8BCC643DFFB5FFFAFF0DB9F75D6BA46D" blockId="16.[151,643,1221,1309]" pageId="16" pageNumber="347">
<emphasis id="B907B82FFFB5FFFAFF0DB9F75E41A5AD" bold="true" box="[151,263,1221,1245]" pageId="16" pageNumber="347">Remarks.</emphasis>
Piece of clear, golden amber that measures 10x9x
<quantity id="4C8BC9D8FFB5FFFAFEDCB9D45ED9A58D" box="[326,415,1254,1277]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="347" unit="mm" value="3.5">3.5 mm</quantity>
. In the inclusion the right antenna and a leg are not visible.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,162 @@
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<mods:title id="2ED6B55E25032C18EC861D63DE95163D">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<paragraph id="8BCC643DFFACFFE3FFEBBFA45D68A3DE" blockId="9.[113,558,662,718]" box="[113,558,662,686]" pageId="9" pageNumber="340">
<heading id="D084D351FFACFFE3FFEBBFA45D68A3DE" bold="true" box="[113,558,662,686]" fontSize="10" level="2" pageId="9" pageNumber="340" reason="4">
<emphasis id="B907B82FFFACFFE3FFEBBFA45D68A3DE" bold="true" box="[113,558,662,686]" pageId="9" pageNumber="340">
<taxonomicName id="4C731FBEFFACFFE3FFEBBFA45E95A3DE" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[113,467,662,686]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="barsevskisi" status="sp. nov.">
<emphasis id="B907B82FFFACFFE3FFEBBFA45E95A3DE" bold="true" box="[113,467,662,686]" italics="true" pageId="9" pageNumber="340">Hukawngichthyurus barsevskisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFACFFE3FE43BFA45D68A3DE" box="[473,558,662,686]" pageId="9" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFACFFE3FFEBBF855F87A3BE" box="[113,193,695,718]" pageId="9" pageNumber="340" type="description">
<paragraph id="8BCC643DFFACFFE3FFEBBF855F87A3BE" blockId="9.[113,558,662,718]" box="[113,193,695,718]" pageId="9" pageNumber="340">
(
<figureCitation id="134878B8FFACFFE3FFE3BF855FFFA3BE" box="[121,185,695,718]" captionStart="Fig" captionStartId="10.[151,190,1639,1662]" captionTargetBox="[151,1172,386,1611]" captionTargetId="figure-18@10.[151,1172,386,1611]" captionTargetPageId="10" captionText="Fig. 5. Hukawngichthyurus barsevskisi sp. nov. in Burmese amber. AB: Holotype, dorso­lateral views." pageId="9" pageNumber="340">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFACFFE3FFEBBFDC5ED6A549" pageId="9" pageNumber="340" type="materials_examined">
<paragraph id="8BCC643DFFACFFE3FFEBBFDC5D58A2F9" blockId="9.[113,605,750,905]" pageId="9" pageNumber="340">
<materialsCitation id="3B1B6E60FFACFFE3FFEBBFDC5D58A2F9" collectingDate="2021-12-19" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="9" pageNumber="340" specimenCode="BUB4328" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFACFFE3FFEBBFDC5FA6A276" bold="true" box="[113,224,750,774]" pageId="9" pageNumber="340">
<typeStatus id="54C8DA9FFFACFFE3FFEBBFDC5F9DA276" box="[113,219,750,774]" pageId="9" pageNumber="340" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFACFFE3FF7DBFDC5E66A275" box="[231,288,750,773]" pageId="9" pageNumber="340">Male</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFACFFE3FE64BFDC5D1BA275" box="[510,605,750,773]" pageId="9" pageNumber="340">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFACFFE3FFE3BE3D5F8FA256" box="[121,201,783,806]" country="Myanmar" name="Kachin" pageId="9" pageNumber="340">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFACFFE3FEF8BE3D5EF3A256" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FEF8BE3D5EF3A256" box="[354,437,783,806]" county="Male" municipality="Burmese" name="Catalog" pageId="9" pageNumber="340" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFACFFE3FF27BE025E03A237" box="[189,325,816,839]" pageId="9" pageNumber="340" value="2021-12-19">
<collectingDate id="EF89BB15FFACFFE3FF27BE025E03A237" box="[189,325,816,839]" pageId="9" pageNumber="340" value="2021-12-19">
2021 XII
<quantity id="4C8BC9D8FFACFFE3FEB0BE025E03A237" box="[298,325,816,839]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="9" pageNumber="340" unit="in" value="19.0">19</quantity>
</collectingDate>
</date>
in the
<location id="8EAC32E6FFACFFE3FE0FBE025FA6A218" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FE0FBE025FA6A218" county="Male" municipality="Burmese" name="Bayerische Staatssammlung" pageId="9" pageNumber="340" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFACFFE3FE96BE635ED9A218" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FE96BE635ED9A218" box="[268,415,849,872]" county="Male" municipality="Burmese" name="Palaontologie" pageId="9" pageNumber="340" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFACFFE3FE49BE635D73A218" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FE49BE635D73A218" box="[467,565,849,872]" county="Male" municipality="Burmese" name="Geologie" pageId="9" pageNumber="340" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFACFFE3FFEBBE405F8BA2F9" LSID="urn:lsid:plazi:treatment:03DAD52BFFACFFE0FFEBBFA45BD2A03C:8EAC32E6FFACFFE3FFEBBE405F8BA2F9" box="[113,205,882,905]" county="Male" municipality="Burmese" name="Muller's" pageId="9" pageNumber="340" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFACFFE3FE38BE405D55A2F9" box="[418,531,882,905]" collectionCode="BUB" pageId="9" pageNumber="340">BUB4328</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFACFFE3FFEBBE9B5E82A291" blockId="9.[113,605,937,993]" pageId="9" pageNumber="340">
<emphasis id="B907B82FFFACFFE3FFEBBE9B5E4CA2B1" bold="true" box="[113,266,937,961]" pageId="9" pageNumber="340">Type locality.</emphasis>
<collectingCountry id="F36424ADFFACFFE3FE89BE9B5E39A2B0" box="[275,383,937,960]" name="Myanmar" pageId="9" pageNumber="340">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFACFFE3FE10BE9B5D52A2B0" box="[394,532,937,960]" country="Myanmar" name="Kachin" pageId="9" pageNumber="340">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFACFFE3FFEBB9335ED6A549" blockId="9.[113,605,1025,1081]" pageId="9" pageNumber="340">
<emphasis id="B907B82FFFACFFE3FFEBB9335E43A569" bold="true" box="[113,261,1025,1049]" pageId="9" pageNumber="340">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFACFFE3FFEBB96B5E11A76B" pageId="9" pageNumber="340" type="diagnosis">
<paragraph id="8BCC643DFFACFFE3FFEBB96B5E11A76B" blockId="9.[113,605,1113,1563]" pageId="9" pageNumber="340">
<emphasis id="B907B82FFFACFFE3FFEBB96B5E3EA501" bold="true" box="[113,376,1113,1137]" pageId="9" pageNumber="340">Differential diagnosis.</emphasis>
The new species is distinguishable from other
<taxonomicName id="4C731FBEFFACFFE3FE12B9495D1AA5E2" authorityName="Fanti &amp; Ellenberger" authorityYear="2018" box="[392,604,1147,1170]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFACFFE3FE12B9495D1AA5E2" box="[392,604,1147,1170]" italics="true" pageId="9" pageNumber="340">Hukawngichthyurus</emphasis>
</taxonomicName>
by its pectinate antennae. We attribute this new species to this genus on the basis of the very short and irregular antennomeres IIIII, for the pronotum with depressions and for the last abdominal segments in a kind of “fish tail.” The antennae are often a variable character. For example, in the fossil genus
<emphasis id="B907B82FFFACFFE3FEB3B8525EF2A407" box="[297,436,1376,1399]" italics="true" pageId="9" pageNumber="340">
<taxonomicName id="4C731FBEFFACFFE3FEB3B8525EF6A407" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[297,432,1376,1399]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="genus">Sanaungulus</taxonomicName>
,
</emphasis>
there are species with pectinate or filiform antennae. Unlike what
<bibRefCitation id="EFE219CCFFACFFE3FFEBB8905E6AA4C9" author="Yang Y. &amp; Zhao W. &amp; Geiser M. F. &amp; Zhang W. &amp; Bai M. &amp; Liu H." box="[113,300,1442,1465]" pageId="9" pageNumber="340" refId="ref21251" refString="Yang Y., Zhao W., Geiser M. F., Zhang W., Bai M., Liu H. 2021. Review of the genus Burmomiles Fanti, Damgaard &amp; Ellenberger from mid-Cretaceous Burmese amber (Coleoptera, Cantharidae). Cretaceous Research 128: 104961 [9 pp.]." type="book" year="2021">Yang et al. (2021)</bibRefCitation>
say about
<taxonomicName id="4C731FBEFFACFFE3FE00B8905EE1A4AA" authority="(Hsiao &amp; Huang, 2018)" baseAuthorityName="Hsiao &amp; Huang" baseAuthorityYear="2018" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="ruicheni">
<emphasis id="B907B82FFFACFFE3FE00B8905FD9A4AA" italics="true" pageId="9" pageNumber="340">Sanaungulus ruicheni</emphasis>
(
<bibRefCitation id="EFE219CCFFACFFE3FF34B8F15ED9A4AA" author="Hsiao Y. &amp; Huang C. - L." box="[174,415,1475,1498]" pageId="9" pageNumber="340" pagination="257 - 263" refId="ref20597" refString="Hsiao Y., Huang C. - L. 2018. Taxonomic revision on the genus Ornatomalthinus Poinar and Fanti (Coleoptera: Cantharidae), with description of a new species from the Cretaceous Burmese amber. Cretaceous Research 92: 257 - 263." type="journal article" year="2018">Hsiao &amp; Huang, 2018</bibRefCitation>
)
</taxonomicName>
, this is not a distinguishing character, being extremely variable even in living genera.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFACFFE0FFEBBB095EF6A1A3" lastPageId="10" lastPageNumber="341" pageId="9" pageNumber="340" type="description">
<paragraph id="8BCC643DFFACFFE3FFEBBB095EA6A7E4" blockId="9.[113,605,1595,1684]" pageId="9" pageNumber="340">
<emphasis id="B907B82FFFACFFE3FFEBBB095FBCA723" bold="true" box="[113,250,1595,1619]" pageId="9" pageNumber="340">Description.</emphasis>
Adult, alate, slender. Male, for the shape of the last abdominal segment. Entirely dark brown. Body length:
<quantity id="4C8BC9D8FFACFFE3FE12BB4F5E9AA7E4" box="[392,476,1661,1684]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="340" unit="mm" value="3.4">3.4 mm</quantity>
.
</paragraph>
<paragraph id="8BCC643DFFACFFE0FD19BDA95EF6A1A3" blockId="9.[643,1134,155,1682]" lastBlockId="10.[151,643,155,211]" lastPageId="10" lastPageNumber="341" pageId="9" pageNumber="340">Head rounded, convex, slightly restricted behind the eyes, surface with shallow punctation and short setae. Eyes large and prominent, subelliptical, inserted laterally to the head. Rostrum elongated. Mandibles falciform. Maxillary palps 4­ segmented, last palpomere securiform with rounded apex. Labial palps 3­ segmented, last palpomere stout and slightly securiform.Antennae 11­ segmented, relatively short, pectinate, surpassing the half of elytra and reaching the half of abdomen, insertion of the antennae close to the eyes and in their lower part; scape robust, elongated, club­shaped; antennomere II very short, moniliform­globular, asymmetrical; antennomere III short, sturdier and longer than second, slightly irregular­distorted; antennomeres IVX pectinate, equipped with a lobe which is very long (longer than antennomeres), rounded at apex and inserted in the basal part of antennomeres; antennomere XI filiform, rounded at apex; all antennomeres punctate with not erected and very short setae. Pronotum transverse, slightly wider than head, anterior margin undulate, posterior margin undulate and bordered, sides irregular (almost straight), surface irregular with draft and depressions (particularly near the anterior margin) and equipped with short setae. Scutellum triangular. Elytra wider than pronotum, parallel­sided, rounded and slightly dehiscent at apex, rather short and revealing the last three abdominal segments, surface smooth without punctation, equipped with few little and thin striae. Metathoracic wings transparent, folded, very slightly surpassing the elytra. Metasternum sub­quadrate; sternites transverse. Last urite with an apical lobe in each side, which is extremely long, thin, with rounded apex. Legs slender, rather long, with pubescence; coxae elongated and robust; trochanters globular­shaped, irregular, with apex rounded; femora robust and not curved, almost flat, sturdier and longer than tibiae; tibiae cylindrical, thin, without spurs at apex. Tarsal formula 5­ 5­ 5; first tarsomere elongated, about 1.2 times longer than second; second tarsomere thin at base and slightly enlarged apically; third tarsomere triangular and shorter than second; fourth tarsomere bilobed with the lobes very long and not particularly wide; fifth tarsomere very long; claws simple without denticles or lobes.</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFAFFFE0FF0DBDC15C96A1A3" pageId="10" pageNumber="341" type="etymology">
<paragraph id="8BCC643DFFAFFFE0FF0DBDC15EDAA03C" blockId="10.[151,643,237,332]" pageId="10" pageNumber="341">
<emphasis id="B907B82FFFAFFFE0FF0DBDC15E5FA07B" bold="true" box="[151,281,243,267]" pageId="10" pageNumber="341">Etymology.</emphasis>
Named in honor of ArvÓds Barševskis (
<collectingRegion id="49B7AADFFFAFFFE0FF6FBC265E36A05B" box="[245,368,276,299]" country="Latvia" name="Daugavpils" pageId="10" pageNumber="341">Daugavpils</collectingRegion>
University,
<collectingCountry id="F36424ADFFAFFFE0FE62BC265D06A05B" box="[504,576,276,299]" name="Latvia" pageId="10" pageNumber="341">Latvia</collectingCountry>
), dear friend of the first author.
</paragraph>
<paragraph id="8BCC643DFFAFFFE0FD32BDA95C96A1A3" blockId="10.[680,1172,155,211]" pageId="10" pageNumber="341">
<emphasis id="B907B82FFFAFFFE0FD32BDA95C14A1C3" bold="true" box="[680,850,155,179]" pageId="10" pageNumber="341">Syninclusions.</emphasis>
Remains of another soldier beetle, air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFAFFFE0FD32BDC15BD2A03C" pageId="10" pageNumber="341" type="discussion">
<paragraph id="8BCC643DFFAFFFE0FD32BDC15BD2A03C" blockId="10.[680,1172,243,332]" pageId="10" pageNumber="341">
<emphasis id="B907B82FFFAFFFE0FD32BDC15C52A07B" bold="true" box="[680,788,243,267]" pageId="10" pageNumber="341">Remarks.</emphasis>
Piece of clear, golden amber that measures 11x9x
<quantity id="4C8BC9D8FFAFFFE0FC87BC265C10A05B" box="[797,854,276,299]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="341" unit="mm" value="3.0">3 mm</quantity>
.The inclusion is complete except for the left anterior leg preserved until the tibia.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,161 @@
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<mods:title id="F64EAADABE4A56A98A484F0B12C64C77">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<paragraph id="8BCC643DFFAEFFE1FFEBBDA85D5DA1C2" blockId="11.[113,539,154,210]" box="[113,539,154,178]" pageId="11" pageNumber="342">
<heading id="D084D351FFAEFFE1FFEBBDA85D5DA1C2" bold="true" box="[113,539,154,178]" fontSize="10" level="2" pageId="11" pageNumber="342" reason="4">
<emphasis id="B907B82FFFAEFFE1FFEBBDA85D5DA1C2" bold="true" box="[113,539,154,178]" pageId="11" pageNumber="342">
<taxonomicName id="4C731FBEFFAEFFE1FFEBBDA85EF9A1C2" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[113,447,154,178]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="342" phylum="Arthropoda" rank="species" species="ditaddeoi" status="sp. nov.">
<emphasis id="B907B82FFFAEFFE1FFEBBDA85EF9A1C2" bold="true" box="[113,447,154,178]" italics="true" pageId="11" pageNumber="342">Hukawngichthyurus ditaddeoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFAEFFE1FE5FBDA85D5DA1C2" box="[453,539,154,178]" pageId="11" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFAEFFE1FFEBBD895F87A1A2" box="[113,193,187,210]" pageId="11" pageNumber="342" type="description">
<paragraph id="8BCC643DFFAEFFE1FFEBBD895F87A1A2" blockId="11.[113,539,154,210]" box="[113,193,187,210]" pageId="11" pageNumber="342">
(
<figureCitation id="134878B8FFAEFFE1FFE3BD895FFFA1A2" box="[121,185,187,210]" captionStart="Fig" captionStartId="11.[113,152,1636,1659]" captionTargetBox="[113,1134,382,1607]" captionTargetId="figure-98@11.[113,1134,382,1607]" captionTargetPageId="11" captionText="Fig. 6. Hukawngichthyurus ditaddeoi sp. nov. in Burmese amber. A: Holotype, ventro­lateral view; B: Holotype, dorso­lateral view." pageId="11" pageNumber="342">Fig. 6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFAEFFE6FFEBBDC35EF0A1A3" lastPageId="12" lastPageNumber="343" pageId="11" pageNumber="342" type="materials_examined">
<paragraph id="8BCC643DFFAEFFE1FFEBBDC35D8FA182" blockId="11.[113,605,241,328]" lastBlockId="11.[643,1134,155,242]" pageId="11" pageNumber="342">
<materialsCitation id="3B1B6E60FFAEFFE1FFEBBDC35D8FA182" collectingDate="2021-12-17" collectionCode="BUB" county="Probable" location="Catalog" municipality="Burmese" pageId="11" pageNumber="342" specimenCode="BUB 4326" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFAEFFE1FFEBBDC35FA5A079" bold="true" box="[113,227,241,265]" pageId="11" pageNumber="342">
<typeStatus id="54C8DA9FFFAEFFE1FFEBBDC35F9DA079" box="[113,219,241,265]" pageId="11" pageNumber="342" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFAEFFE1FF77BDC35E08A078" box="[237,334,241,264]" pageId="11" pageNumber="342">Probable</collectingCounty>
female, adult specimen in a
<collectingMunicipality id="6BA8FE47FFAEFFE1FF3FBC235E42A058" box="[165,260,273,296]" pageId="11" pageNumber="342">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFAEFFE1FE8EBC235E22A058" box="[276,356,273,296]" country="Myanmar" name="Kachin" pageId="11" pageNumber="342">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFAEFFE1FD90BC235D1BA058" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FD90BC235D1BA058" box="[522,605,273,296]" county="Probable" municipality="Burmese" name="Catalog" pageId="11" pageNumber="342" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFAEFFE1FEC4BC035EA6A038" box="[350,480,305,328]" pageId="11" pageNumber="342" value="2021-12-17">
<collectingDate id="EF89BB15FFAEFFE1FEC4BC035EA6A038" box="[350,480,305,328]" pageId="11" pageNumber="342" value="2021-12-17">
2021 XII
<quantity id="4C8BC9D8FFAEFFE1FE5FBC035EA6A038" box="[453,480,305,328]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="11" pageNumber="342" unit="in" value="17.0">17</quantity>
</collectingDate>
</date>
in the
<location id="8EAC32E6FFAEFFE1FDB3BC035CC5A1C2" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FDB3BC035CC5A1C2" box="[553,899,155,328]" county="Probable" municipality="Burmese" name="Bayerische Staatssammlung" pageId="11" pageNumber="342" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFAEFFE1FC34BDA95B07A1C2" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FC34BDA95B07A1C2" box="[942,1089,155,178]" county="Probable" municipality="Burmese" name="Palaontologie" pageId="11" pageNumber="342" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFAEFFE1FD19BD895DA2A1A2" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FD19BD895DA2A1A2" box="[643,740,187,210]" county="Probable" municipality="Burmese" name="Geologie" pageId="11" pageNumber="342" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFAEFFE1FC95BD895C2FA1A2" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FC95BD895C2FA1A2" box="[783,873,187,210]" county="Probable" municipality="Burmese" name="Muller's" pageId="11" pageNumber="342" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFAEFFE1FBADBD895DFBA182" collectionCode="BUB" pageId="11" pageNumber="342">BUB 4326</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFAEFFE1FD19BC235C93A038" blockId="11.[643,1134,273,328]" pageId="11" pageNumber="342">
<emphasis id="B907B82FFFAEFFE1FD19BC235C5AA059" bold="true" box="[643,796,273,297]" pageId="11" pageNumber="342">Type locality.</emphasis>
<collectingCountry id="F36424ADFFAEFFE1FCBEBC235CD6A058" box="[804,912,273,296]" name="Myanmar" pageId="11" pageNumber="342">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFAEFFE1FC01BC235B63A058" box="[923,1061,273,296]" country="Myanmar" name="Kachin" pageId="11" pageNumber="342">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<caption id="DF0C34B5FFAEFFE1FFEBBB565D4EA7EB" pageId="11" pageNumber="342" startId="11.[113,152,1636,1659]" targetBox="[113,1134,382,1607]" targetPageId="11" targetType="figure">
<paragraph id="8BCC643DFFAEFFE1FFEBBB565D4EA7EB" blockId="11.[113,1134,1636,1691]" pageId="11" pageNumber="342">
Fig. 6.
<taxonomicName id="4C731FBEFFAEFFE1FF59BB565D4EA70B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[195,520,1636,1659]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="342" phylum="Arthropoda" rank="species" species="ditaddeoi" status="sp. nov.">
<emphasis id="B907B82FFFAEFFE1FF59BB565D4EA70B" box="[195,520,1636,1659]" italics="true" pageId="11" pageNumber="342">Hukawngichthyurus ditaddeoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFAEFFE1FD88BB565D21A70B" box="[530,615,1636,1659]" pageId="11" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, ventro­lateral view; B: Holotype, dorso­lateral view.
</paragraph>
</caption>
<paragraph id="8BCC643DFFA9FFE6FF0DBDA95EF0A1A3" blockId="12.[151,643,155,211]" pageId="12" pageNumber="343">
<emphasis id="B907B82FFFA9FFE6FF0DBDA95E6CA1C3" bold="true" box="[151,298,155,179]" pageId="12" pageNumber="343">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA9FFE6FF0DBDC15E86A0BF" pageId="12" pageNumber="343" type="diagnosis">
<paragraph id="8BCC643DFFA9FFE6FF0DBDC15E86A0BF" blockId="12.[151,643,243,463]" pageId="12" pageNumber="343">
<emphasis id="B907B82FFFA9FFE6FF0DBDC15EDFA07B" bold="true" box="[151,409,243,267]" pageId="12" pageNumber="343">Differential diagnosis.</emphasis>
The relatively small body size, the pronotum irregular with a convex draft, and the last urites in a kind of “fish tail” make this new species belong to the genus
<emphasis id="B907B82FFFA9FFE6FF0DBC455E32A0FE" box="[151,372,375,398]" italics="true" pageId="12" pageNumber="343">
<taxonomicName id="4C731FBEFFA9FFE6FF0DBC455E36A0FE" authorityName="Fanti &amp; Ellenberger" authorityYear="2018" box="[151,368,375,398]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="343" phylum="Arthropoda" rank="genus">Hukawngichthyurus</taxonomicName>
.
</emphasis>
The new species is distinguishable from other
<taxonomicName id="4C731FBEFFA9FFE6FEE9BCAA5D01A0DF" authorityName="Fanti &amp; Ellenberger" authorityYear="2018" box="[371,583,408,431]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFA9FFE6FEE9BCAA5D01A0DF" box="[371,583,408,431]" italics="true" pageId="12" pageNumber="343">Hukawngichthyurus</emphasis>
</taxonomicName>
by its longer pronotum and elytra.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA9FFE6FF0DBCDD5C2CA3CB" pageId="12" pageNumber="343" type="description">
<paragraph id="8BCC643DFFA9FFE6FF0DBCDD5D0FA338" blockId="12.[151,643,495,584]" pageId="12" pageNumber="343">
<emphasis id="B907B82FFFA9FFE6FF0DBCDD5E6EA377" bold="true" box="[151,296,495,519]" pageId="12" pageNumber="343">Description.</emphasis>
Adult, alate, robust. Probable female, for the wide abdomen. Entirely brown with darker elytra. Body length:
<quantity id="4C8BC9D8FFA9FFE6FE6ABF035D03A338" box="[496,581,561,584]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="12" pageNumber="343" unit="mm" value="4.9">4.9 mm</quantity>
.
</paragraph>
<paragraph id="8BCC643DFFA9FFE6FF0DBF5B5C2CA3CB" blockId="12.[151,644,617,1690]" lastBlockId="12.[680,1172,155,699]" pageId="12" pageNumber="343">Head elongated, convex in the anterior part and almost flat posteriorly, slightly restricted behind the eyes, surface finely punctated. Eyes large and prominent, subelliptical, inserted laterally to the head. Rostrum elongated and robust. Mandibles falciform. Maxillary palps 4­ segmented, last palpomere securiform with rounded apex. Labial palps 3­ segmented, last palpomere stout and slightly securiform.Antennae 11­ segmented, short, filiform with the central antennomeres enlarged, reaching almost the half of elytra, insertion of the antennae close to the eyes and in their lower part; scape robust, elongated, club­shaped; antennomere II very short, moniliform­triangular; antennomere III moniliform­globular, slightly irregular; antennomeres IV Velongated, robust; antennomeresVIX filiform, less sturdier than previous ones; antennomere XI filiform, rounded at apex; all antennomeres with not erected and very short setae. Pronotum elongated, slightly narrower than head, anterior margin slightly undulated, posterior margin strongly undulated and slightly bordered, sides straight, surface irregular with a central convex draft. Scutellum triangular. Elytra wider than pronotum, parallel­sided, rounded at apex, rather elongated and revealing the last two abdominal segments, surface without punctation, equipped with few little and thin striae. Metathoracic wings transparent, folded, very long, surpassing the elytra and the last tergites. Metasternum large, stout, sub­quadrate; sternites wide, transverse. Last urite with two asymmetrical apical lobes in a kind of fish tail, one of which is curved and very long with the base slightly enlarged and the center thinner and newly enlarged at apex, while the other one is very short and sturdier. Legs robust, long, with pubescence; coxae elongated and robust; trochanters slightly elongated, globular­shaped with apex rounded; femora robust and slightly curved, sturdier and longer than tibiae; tibiae cylindrical, thin, without spurs at apex. Tarsal formula 5­ 5­ 5; first tarsomere elongated, about 1.2 times longer than second; second tarsomere thin at base and enlarged apically; third tarsomere triangular and shorter than second; fourth tarsomere bilobed with the lobes very long and wide; fifth tarsomere elongated with wider sides; claws simple without denticles or lobes.</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA9FFE6FD32BFE95C32A2FB" pageId="12" pageNumber="343" type="etymology">
<paragraph id="8BCC643DFFA9FFE6FD32BFE95DB4A243" blockId="12.[680,1172,731,819]" pageId="12" pageNumber="343">
<emphasis id="B907B82FFFA9FFE6FD32BFE95C6FA383" bold="true" box="[680,809,731,755]" pageId="12" pageNumber="343">Etymology.</emphasis>
Named in honor of Vincenzo Di Taddeo (Avigliana,
<collectingCountry id="F36424ADFFA9FFE6FCE8BFCE5CE3A263" box="[882,933,764,787]" name="Italy" pageId="12" pageNumber="343">Italy</collectingCountry>
), dear friend of the first author.
</paragraph>
<paragraph id="8BCC643DFFA9FFE6FD32BE615C32A2FB" blockId="12.[680,1172,851,907]" pageId="12" pageNumber="343">
<emphasis id="B907B82FFFA9FFE6FD32BE615C05A21B" bold="true" box="[680,835,851,875]" pageId="12" pageNumber="343">Syninclusions.</emphasis>
Air bubbles (few), detritus (many botanical remains).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFA9FFE6FD32BE995CC7A534" pageId="12" pageNumber="343" type="discussion">
<paragraph id="8BCC643DFFA9FFE6FD32BE995CC7A534" blockId="12.[680,1172,939,1092]" pageId="12" pageNumber="343">
<emphasis id="B907B82FFFA9FFE6FD32BE995C51A2B3" bold="true" box="[680,791,939,963]" pageId="12" pageNumber="343">Remarks.</emphasis>
Piece of clear amber that measures 13x12x
<quantity id="4C8BC9D8FFA9FFE6FD63BEF95C71A292" box="[761,823,971,994]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="343" unit="mm" value="3.0">3 mm</quantity>
. The inclusion is complete except for the missing right metathoracic leg. Furthermore, the right antenna is preserved until the eighth antennomere.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,168 @@
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<mods:title id="D014F062D599885A8A4CBA9E2E40C313">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<subSubSection id="C36937B6FFB0FFFFFD19BDA85CBFA1C2" box="[643,1017,154,178]" pageId="21" pageNumber="352" type="nomenclature">
<paragraph id="8BCC643DFFB0FFFFFD19BDA85CBFA1C2" blockId="21.[643,1017,154,210]" box="[643,1017,154,178]" pageId="21" pageNumber="352">
<heading id="D084D351FFB0FFFFFD19BDA85CBFA1C2" bold="true" box="[643,1017,154,178]" fontSize="10" level="2" pageId="21" pageNumber="352" reason="4">
<emphasis id="B907B82FFFB0FFFFFD19BDA85CBFA1C2" bold="true" box="[643,1017,154,178]" pageId="21" pageNumber="352">
<taxonomicName id="4C731FBEFFB0FFFFFD19BDA85CD8A1C2" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[643,926,154,178]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="352" phylum="Arthropoda" rank="species" species="kachinensis" status="sp. nov.">
<emphasis id="B907B82FFFB0FFFFFD19BDA85CD8A1C2" bold="true" box="[643,926,154,178]" italics="true" pageId="21" pageNumber="352">Sanaungulus kachinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB0FFFFFC3EBDA85CBFA1C2" box="[932,1017,154,178]" pageId="21" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB0FFFFFD19BD895DA6A1A2" box="[643,736,187,210]" pageId="21" pageNumber="352" type="description">
<paragraph id="8BCC643DFFB0FFFFFD19BD895DA6A1A2" blockId="21.[643,1017,154,210]" box="[643,736,187,210]" pageId="21" pageNumber="352">
(
<figureCitation id="134878B8FFB0FFFFFD11BD895D91A1A2" box="[651,727,187,210]" captionStart="Fig" captionStartId="22.[151,190,1665,1688]" captionTargetBox="[151,1171,412,1637]" captionTargetId="figure-115@22.[151,1171,412,1637]" captionTargetPageId="22" captionText="Fig. 12. Sanaungulus kachinensis sp. nov. in Burmese amber. AB: Holotype, lateral views." pageId="21" pageNumber="352">Fig. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB0FFFFFD19BDC35CE4A346" pageId="21" pageNumber="352" type="materials_examined">
<paragraph id="8BCC643DFFB0FFFFFD19BDC35B69A0F8" blockId="21.[643,1134,241,392]" pageId="21" pageNumber="352">
<materialsCitation id="3B1B6E60FFB0FFFFFD19BDC35B69A0F8" collectingDate="2021-12-14" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="21" pageNumber="352" specimenCode="BUB4312" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFB0FFFFFD19BDC35DB7A079" bold="true" box="[643,753,241,265]" pageId="21" pageNumber="352">
<typeStatus id="54C8DA9FFFB0FFFFFD19BDC35DABA079" box="[643,749,241,265]" pageId="21" pageNumber="352" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFB0FFFFFD6DBDC35C76A078" box="[759,816,241,264]" pageId="21" pageNumber="352">Male</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFB0FFFFFB95BDC35B28A078" box="[1039,1134,241,264]" pageId="21" pageNumber="352">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFB0FFFFFD10BC235D9CA058" box="[650,730,273,296]" country="Myanmar" name="Kachin" pageId="21" pageNumber="352">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFB0FFFFFCE9BC235C80A058" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFCE9BC235C80A058" box="[883,966,273,296]" county="Male" municipality="Burmese" name="Catalog" pageId="21" pageNumber="352" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFB0FFFFFD54BC035C10A038" box="[718,854,305,328]" pageId="21" pageNumber="352" value="2021-12-14">
<collectingDate id="EF89BB15FFB0FFFFFD54BC035C10A038" box="[718,854,305,328]" pageId="21" pageNumber="352" value="2021-12-14">
2021 XII
<quantity id="4C8BC9D8FFB0FFFFFCA1BC035C10A038" box="[827,854,305,328]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="21" pageNumber="352" unit="in" value="14.0">14</quantity>
</collectingDate>
</date>
in the
<location id="8EAC32E6FFB0FFFFFC3CBC035DB4A018" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFC3CBC035DB4A018" county="Male" municipality="Burmese" name="Bayerische Staatssammlung" pageId="21" pageNumber="352" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFB0FFFFFC87BC635CF6A018" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFC87BC635CF6A018" box="[797,944,337,360]" county="Male" municipality="Burmese" name="Palaontologie" pageId="21" pageNumber="352" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFB0FFFFFC7EBC635B00A018" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFC7EBC635B00A018" box="[996,1094,337,360]" county="Male" municipality="Burmese" name="Geologie" pageId="21" pageNumber="352" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFB0FFFFFD19BC435D99A0F8" LSID="urn:lsid:plazi:treatment:03DAD52BFFB0FFFDFD19BDA85D18A2C0:8EAC32E6FFB0FFFFFD19BC435D99A0F8" box="[643,735,369,392]" county="Male" municipality="Burmese" name="Muller's" pageId="21" pageNumber="352" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFB0FFFFFC29BC435B62A0F8" box="[947,1060,369,392]" collectionCode="BUB" pageId="21" pageNumber="352">BUB4312</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFB0FFFFFD19BC9A5C93A0AF" blockId="21.[643,1134,424,479]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFD19BC9A5C5AA0B0" bold="true" box="[643,796,424,448]" pageId="21" pageNumber="352">Type locality.</emphasis>
<collectingCountry id="F36424ADFFB0FFFFFCBEBC9A5CD6A0CF" box="[804,912,424,447]" name="Myanmar" pageId="21" pageNumber="352">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFB0FFFFFC01BC9A5B63A0CF" box="[923,1061,424,447]" country="Myanmar" name="Kachin" pageId="21" pageNumber="352">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFB0FFFFFD19BCCC5CE4A346" blockId="21.[643,1134,510,566]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFD19BCCC5C50A366" bold="true" box="[643,790,510,534]" pageId="21" pageNumber="352">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB0FFFFFD19BF675CC8A2BC" pageId="21" pageNumber="352" type="diagnosis">
<paragraph id="8BCC643DFFB0FFFFFD19BF675CC8A2BC" blockId="21.[643,1134,597,972]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFD19BF675C31A31D" bold="true" box="[643,887,597,621]" pageId="21" pageNumber="352">Differential diagnosis.</emphasis>
The new species is characterized by antennae equipped with an antennal process near the base of antennomeres IV X. It differs from
<taxonomicName id="4C731FBEFFB0FFFFFCDCBF845B54A3BD" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[838,1042,694,717]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="352" phylum="Arthropoda" rank="species" species="havai" status="sp. nov.">
<emphasis id="B907B82FFFB0FFFFFCDCBF845B54A3BD" box="[838,1042,694,717]" italics="true" pageId="21" pageNumber="352">Sanaungulus havai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB0FFFFFB81BF875B28A3BC" box="[1051,1134,693,716]" pageId="21" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
based on the head that is less triangular behind the eyes, the pronotum with a median incision, and by antennae with longer rami. It differs from
<taxonomicName id="4C731FBEFFB0FFFFFD19BE045B12A23C" authority="Y. Yang, Liu &amp; Zhao" authorityName="Y. Yang, Liu &amp; Zhao" box="[643,1108,821,845]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="352" phylum="Arthropoda" rank="species" species="ruficollis">
<emphasis id="B907B82FFFB0FFFFFD19BE045C34A23D" box="[643,882,822,845]" italics="true" pageId="21" pageNumber="352">Sanaungulus ruficollis</emphasis>
Y. Yang, Liu &amp; Zhao
</taxonomicName>
in Y. Yang, Zhao, Geiser, W. Zhang, Ren, Bai &amp; Liu, 2022 based on the antennal processes, the pronotum incised, and by longer metathoracic wings (
<bibRefCitation id="EFE219CCFFB0FFFFFD4ABE875CC5A2BC" author="Yang Y. &amp; Zhao W. &amp; Geiser M. F. &amp; Zhang W. &amp; Ren D. &amp; Bai M. &amp; Liu H." box="[720,899,949,972]" pageId="21" pageNumber="352" refId="ref21309" refString="Yang Y., Zhao W., Geiser M. F., Zhang W., Ren D., Bai M., Liu H. 2022. Taxonomic revision of the mid-Cretaceous genus Sanaungulus Fanti, Damgaard et Ellenberger, 2018 (Coleoptera, Cantharidae) from northern Myanmar based on a morphology-based phylogeny. Cretaceous Research 135: 105217 [14 pp. + supplementary data]." type="book" year="2022">Yang et al. 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB0FFFDFD19BEDE5D57A31E" lastPageId="23" lastPageNumber="354" pageId="21" pageNumber="352" type="description">
<paragraph id="8BCC643DFFB0FFFFFD19BEDE5B14A533" blockId="21.[643,1135,1004,1091]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFD19BEDE5C4EA574" bold="true" box="[643,776,1004,1028]" pageId="21" pageNumber="352">Description.</emphasis>
Adult, winged, slender. Male, defined on the shape of the last abdominal segments. Body length about
<quantity id="4C8BC9D8FFB0FFFFFCD6B91E5CE7A533" box="[844,929,1068,1091]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="21" pageNumber="352" unit="mm" value="2.8">2.8 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFB0FFFDFD19B9515D57A31E" blockId="21.[643,1135,1123,1690]" lastBlockId="23.[113,605,155,622]" lastPageId="23" lastPageNumber="354" pageId="21" pageNumber="352">Head elongated, slightly constricted behind the eyes, fitted with small rugosity. Eyes very prominent, roundish, inserted in the upper and dorsolateral part of the head. Mandibles elongated, very robust, falciform.Maxillary palps 4­ segmented, with the last palpomere strongly securiform. Labial palps 3­ segmented, with the last palpomere securiform.Antennae 11­ segmented, pectinate, slender, long, surpassing the apex of the elytra; scape elongated, robust; antennomere II short; antennomere III filiform, about 1.9 times longer than second; antennomeres IVX subequal in length, each equipped near their base with a very long, robust, almost straight antennal process (longer than antennomere), almost rounded apically; antennomere XI filiform, extremely long, rounded apically; all the antennomeres with short pubescence. Pronotum longer than wide, as wide as head, anterior margin rounded and bordered, posterior margin rather straight, corners obtuse, sides almost straight, pronotum disc with a transversal median and deep incision, equipped with short setae. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, as wide as pronotum, sub­parallel, rounded apexes, surface with shallow punctuation and several short setae. Metathoracic wings transparent, long, surpassing the elytra and reaching the last abdominal segments. Sternum elongated, almost straight posteriorly, sternites transverse equipped with short setae; last tergite elongated, wide and strongly globular (spatuliform); last sternite elongated, very thin. Legs thin, long, pubescent; coxae massive, elongated, with rounded apex; trochanters elongated, robust, with rounded apex; femora slightly curved, more robust than tibiae; tibiae cylindrical, thin, without spurs near the apex, pro­ and mesotibiae slightly shorter than pro­ and mesofemora, metatibiae slightly longer than metafemora. Tarsal formula 5­5­5; first tarsomere long; second tarsomere about 1.2 times shorter than first; third tarsomere triangular­shaped, short, straight apically; fourth tarsomere short, strongly bilobed; fifth tarsomere elongated, thin, flat and strongly curved; claws simple, apparently without denticle at the base.</paragraph>
</subSubSection>
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<paragraph id="8BCC643DFFB3FFFCFF0DBBB35B2EA7E8" blockId="22.[151,1128,1665,1688]" box="[151,1128,1665,1688]" pageId="22" pageNumber="353">
Fig. 12.
<taxonomicName id="4C731FBEFFB3FFFCFF6ABBB35EBBA7E8" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[240,509,1665,1688]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="353" phylum="Arthropoda" rank="species" species="kachinensis" status="sp. nov.">
<emphasis id="B907B82FFFB3FFFCFF6ABBB35EBBA7E8" box="[240,509,1665,1688]" italics="true" pageId="22" pageNumber="353">Sanaungulus kachinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB3FFFCFD99BBB35D13A7E8" box="[515,597,1665,1688]" pageId="22" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. AB: Holotype, lateral views.
</paragraph>
</caption>
<subSubSection id="C36937B6FFB2FFFDFFEBBFBF5D18A26A" pageId="23" pageNumber="354" type="etymology">
<paragraph id="8BCC643DFFB2FFFDFFEBBFBF5E19A394" blockId="23.[113,605,653,741]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFFEBBFBF5FB7A3D5" bold="true" box="[113,241,653,677]" pageId="23" pageNumber="354">Etymology.</emphasis>
Derived from the toponym
<collectingRegion id="49B7AADFFFB2FFFDFD8ABFBF5D1AA3D4" box="[528,604,653,676]" country="Myanmar" name="Kachin" pageId="23" pageNumber="354">Kachin</collectingRegion>
(the locality where the amber mines are located) + the Latin suffix ­Á
<emphasis id="B907B82FFFB2FFFDFEB4BFFF5E1EA394" box="[302,344,717,740]" italics="true" pageId="23" pageNumber="354">nsis</emphasis>
.
</paragraph>
<paragraph id="8BCC643DFFB2FFFDFFEBBE315D18A26A" blockId="23.[113,606,771,795]" box="[113,606,771,795]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFFEBBE315E4CA26B" bold="true" box="[113,266,771,795]" pageId="23" pageNumber="354">Syninclusions.</emphasis>
Air bubbles, detritus, one
<taxonomicName id="4C731FBEFFB2FFFDFD91BE315D1CA26A" box="[523,602,771,794]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="354" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB2FFFDFFEBBE0B5D18A2C0" pageId="23" pageNumber="354" type="discussion">
<paragraph id="8BCC643DFFB2FFFDFFEBBE0B5D18A2C0" blockId="23.[113,606,825,944]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFFEBBE0B5F9BA221" bold="true" box="[113,221,825,849]" pageId="23" pageNumber="354">Remarks.</emphasis>
Piece of clear (surface is slightly blurred), golden amber that measures 11x9x
<quantity id="4C8BC9D8FFB2FFFDFD9DBE6B5D1FA200" box="[519,601,857,880]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="354" unit="mm" value="2.5">2.5 mm</quantity>
. The inclusion is complete except for the right antenna preserved until the tenth antennomere.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,172 @@
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<mods:title id="8A372D19211881A3353A1727E0B220DF">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:date id="A40EFD4BD7B9455779313288DE0A1C91">2022</mods:date>
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<treatment id="03DAD52BFFB2FFF3FFEBBED45F9FA2F1" ID-DOI="http://doi.org/10.5281/zenodo.13199819" ID-Zenodo-Dep="13199819" LSID="urn:lsid:plazi:treatment:03DAD52BFFB2FFF3FFEBBED45F9FA2F1" httpUri="http://treatment.plazi.org/id/03DAD52BFFB2FFF3FFEBBED45F9FA2F1" lastPageId="25" lastPageNumber="356" pageId="23" pageNumber="354">
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<paragraph id="8BCC643DFFB2FFFDFFEBBED45D15A28D" blockId="23.[113,595,997,1052]" box="[113,595,997,1022]" pageId="23" pageNumber="354">
<heading id="D084D351FFB2FFFDFFEBBED45D15A28D" bold="true" box="[113,595,997,1022]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="354" reason="4">
<emphasis id="B907B82FFFB2FFFDFFEBBED45D15A28D" bold="true" box="[113,595,997,1022]" pageId="23" pageNumber="354">
<taxonomicName id="4C731FBEFFB2FFFDFFEBBED45EB1A28E" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[113,503,998,1022]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="354" phylum="Arthropoda" rank="species" species="kirstenaeweissbachae" status="sp. nov.">
<emphasis id="B907B82FFFB2FFFDFFEBBED45EB1A28E" bold="true" box="[113,503,998,1022]" italics="true" pageId="23" pageNumber="354">Sanaungulus kirstenaeweissbachae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB2FFFDFE64BED75D15A28D" box="[510,595,997,1021]" pageId="23" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB2FFFDFFEBB9375F88A56C" box="[113,206,1029,1052]" pageId="23" pageNumber="354" type="description">
<paragraph id="8BCC643DFFB2FFFDFFEBB9375F88A56C" blockId="23.[113,595,997,1052]" box="[113,206,1029,1052]" pageId="23" pageNumber="354">
(
<figureCitation id="134878B8FFB2FFFDFFE3B9375F80A56C" box="[121,198,1029,1052]" captionStart="Fig" captionStartId="24.[151,190,1636,1659]" captionTargetBox="[261,1066,151,1606]" captionTargetId="figure-44@24.[261,1066,639,1122]" captionTargetPageId="24" captionText="Fig. 13. Sanaungulus kirstenaeweissbachae sp. nov. in Burmese amber.A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of head." pageId="23" pageNumber="354">Fig. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB2FFFDFFEBB90E5ED6A40E" pageId="23" pageNumber="354" type="materials_examined">
<paragraph id="8BCC643DFFB2FFFDFFEBB90E5FA9A5A2" blockId="23.[113,606,1084,1234]" pageId="23" pageNumber="354">
<materialsCitation id="3B1B6E60FFB2FFFDFFEBB90E5FA9A5A2" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="23" pageNumber="354" specimenCode="BUB4483" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFB2FFFDFFEBB90E5F9BA524" bold="true" box="[113,221,1084,1108]" pageId="23" pageNumber="354">
<typeStatus id="54C8DA9FFFB2FFFDFFEBB90E5F9EA524" box="[113,216,1084,1108]" pageId="23" pageNumber="354" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFB2FFFDFF7BB90E5E76A523" box="[225,304,1084,1107]" pageId="23" pageNumber="354">Female</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFB2FFFDFD9AB90E5D1BA523" box="[512,605,1084,1107]" pageId="23" pageNumber="354">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFB2FFFDFFE3B96E5F8EA503" box="[121,200,1116,1139]" country="Myanmar" name="Kachin" pageId="23" pageNumber="354">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFB2FFFDFEC7B96E5EE9A503" LSID="urn:lsid:plazi:treatment:03DAD52BFFB2FFF3FFEBBED45F9FA2F1:8EAC32E6FFB2FFFDFEC7B96E5EE9A503" box="[349,431,1116,1139]" county="Female" municipality="Burmese" name="Catalog" pageId="23" pageNumber="354" stateProvince="Kachin">Catalog</location>
number
<quantity id="4C8BC9D8FFB2FFFDFD92B96E5FFAA5E2" metricMagnitude="2" metricUnit="m" metricValue="1.525778" pageId="23" pageNumber="354" unit="in" value="6007.0">SNHM­6007 in</quantity>
the
<location id="8EAC32E6FFB2FFFDFF78B9495E4FA5C2" LSID="urn:lsid:plazi:treatment:03DAD52BFFB2FFF3FFEBBED45F9FA2F1:8EAC32E6FFB2FFFDFF78B9495E4FA5C2" county="Female" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="23" pageNumber="354" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FFB2FFFDFEACB9A95ED4A5C2" LSID="urn:lsid:plazi:treatment:03DAD52BFFB2FFF3FFEBBED45F9FA2F1:8EAC32E6FFB2FFFDFEACB9A95ED4A5C2" box="[310,402,1179,1202]" county="Female" municipality="Burmese" name="Muller's" pageId="23" pageNumber="354" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFB2FFFDFFEBB9895FA5A5A2" box="[113,227,1211,1234]" collectionCode="BUB" pageId="23" pageNumber="354">BUB4483</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFB2FFFDFFEBB9C35E82A458" blockId="23.[113,605,1265,1320]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFFEBB9C35E4CA479" bold="true" box="[113,266,1265,1289]" pageId="23" pageNumber="354">Type locality.</emphasis>
<collectingCountry id="F36424ADFFB2FFFDFE89B9C35E39A478" box="[275,383,1265,1288]" name="Myanmar" pageId="23" pageNumber="354">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFB2FFFDFE10B9C35D52A478" box="[394,532,1265,1288]" country="Myanmar" name="Kachin" pageId="23" pageNumber="354">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFB2FFFDFFEBB8755ED6A40E" blockId="23.[113,605,1351,1406]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFFEBB8755E43A42F" bold="true" box="[113,261,1351,1375]" pageId="23" pageNumber="354">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB2FFFDFFEBB8AF5C0FA000" pageId="23" pageNumber="354" type="diagnosis">
<paragraph id="8BCC643DFFB2FFFDFFEBB8AF5C0FA000" blockId="23.[113,605,1437,1683]" lastBlockId="23.[643,1134,155,368]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFFEBB8AF5E30A4C5" bold="true" box="[113,374,1437,1461]" pageId="23" pageNumber="354">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFFB2FFFDFEE5B8AC5E58A4A4" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="354" phylum="Arthropoda" rank="species" species="kirstenaeweissbachae" status="sp. nov.">
<emphasis id="B907B82FFFB2FFFDFEE5B8AC5E58A4A4" italics="true" pageId="23" pageNumber="354">Sanaungulus kirstenaeweissbachae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB2FFFDFEBFB88F5E3EA4A4" box="[293,376,1469,1492]" pageId="23" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C731FBEFFB2FFFDFD9EB88F5FE0A764" authority="Fanti &amp; Damgaard, 2019" authorityName="Fanti &amp; Damgaard" authorityYear="2019" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="354" phylum="Arthropoda" rank="species" species="troelsikloevedali">
<emphasis id="B907B82FFFB2FFFDFD9EB88F5E35A484" italics="true" pageId="23" pageNumber="354">Sanaungulus troelsikloevedali</emphasis>
<bibRefCitation id="EFE219CCFFB2FFFDFEE4B8EF5FE0A764" author="Fanti F. &amp; Damgaard A. L." pageId="23" pageNumber="354" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti &amp; Damgaard, 2019</bibRefCitation>
</taxonomicName>
based on its head that is slightly more transverse and a wider pronotum (
<bibRefCitation id="EFE219CCFFB2FFFDFE56BB2F5FA1A723" author="Fanti F. &amp; Damgaard A. L." pageId="23" pageNumber="354" pagination="101 - 125" refId="ref20361" refString="Fanti F., Damgaard A. L. 2019. New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology 19 (2): 101 - 125." type="journal article" year="2019">Fanti &amp; Damgaard 2019</bibRefCitation>
).
<taxonomicName id="4C731FBEFFB2FFFDFF6FBB0E5E94A703" authority="Fanti, Damgaard &amp; Ellenberger, 2018" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="354" phylum="Arthropoda" rank="species" species="ghitaenoerbyae">
<emphasis id="B907B82FFFB2FFFDFF6FBB0E5D58A723" box="[245,542,1596,1619]" italics="true" pageId="23" pageNumber="354">Sanaungulus ghitaenoerbyae</emphasis>
<bibRefCitation id="EFE219CCFFB2FFFDFDB8BB0E5E94A703" author="Fanti F. &amp; Damgaard A. L. &amp; Ellenberger S." pageId="23" pageNumber="354" pagination="170 - 177" refId="ref20442" refString="Fanti F., Damgaard A. L., Ellenberger S. 2018. Two new genera of Cantharidae from Burmese amber of the Hukawng Valley (Insecta, Coleoptera). Cretaceous Research 86: 170 - 177." type="journal article" year="2018">Fanti, Damgaard &amp; Ellenberger, 2018</bibRefCitation>
</taxonomicName>
has the last antennomeres more robust and shorter than
<taxonomicName id="4C731FBEFFB2FFFDFDB7BB4E5C95A1C2" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[557,979,155,1683]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="354" phylum="Arthropoda" rank="species" species="kirstenaeweissbachae" status="sp. nov.">
<emphasis id="B907B82FFFB2FFFDFDB7BB4E5C95A1C2" box="[557,979,155,1683]" italics="true" pageId="23" pageNumber="354">Sanaungulus kirstenaeweissbachae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB2FFFDFC42BDA95B6FA1C2" box="[984,1065,155,178]" pageId="23" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
(Fanti et al. 2018).
<taxonomicName id="4C731FBEFFB2FFFDFC9ABD895B28A1A2" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[768,1134,187,210]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="354" phylum="Arthropoda" rank="species" species="kirstenaeweissbachae" status="sp. nov.">
<emphasis id="B907B82FFFB2FFFDFC9ABD895B28A1A2" box="[768,1134,187,210]" italics="true" pageId="23" pageNumber="354">Sanaungulus kirstenaeweissbachae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB2FFFDFD19BDE85D93A181" box="[643,725,218,241]" pageId="23" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C731FBEFFB2FFFDFCFABDE85C31A061" authority="Y. Yang, Liu &amp; Bai" authorityName="Y. Yang, Liu &amp; Bai" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="354" phylum="Arthropoda" rank="species" species="emarginaticollis">
<emphasis id="B907B82FFFB2FFFDFCFABDE85DFFA061" italics="true" pageId="23" pageNumber="354">Sanaungulus emarginaticollis</emphasis>
Y.Yang, Liu &amp; Bai
</taxonomicName>
in Y.Yang, Zhao, Geiser, W. Zhang, Ren, Bai &amp; Liu, 2022 by its more transverse and different shape of the pronotum (
<bibRefCitation id="EFE219CCFFB2FFFDFD10BC6B5C78A000" author="Yang Y. &amp; Zhao W. &amp; Geiser M. F. &amp; Zhang W. &amp; Ren D. &amp; Bai M. &amp; Liu H." box="[650,830,345,368]" pageId="23" pageNumber="354" refId="ref21309" refString="Yang Y., Zhao W., Geiser M. F., Zhang W., Ren D., Bai M., Liu H. 2022. Taxonomic revision of the mid-Cretaceous genus Sanaungulus Fanti, Damgaard et Ellenberger, 2018 (Coleoptera, Cantharidae) from northern Myanmar based on a morphology-based phylogeny. Cretaceous Research 135: 105217 [14 pp. + supplementary data]." type="book" year="2022">Yang et al. 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB2FFF3FD19BCBD5D0AA329" lastPageId="25" lastPageNumber="356" pageId="23" pageNumber="354" type="description">
<paragraph id="8BCC643DFFB2FFFDFD19BCBD5B56A096" blockId="23.[643,1134,399,486]" pageId="23" pageNumber="354">
<emphasis id="B907B82FFFB2FFFDFD19BCBD5C48A0D7" bold="true" box="[643,782,399,423]" pageId="23" pageNumber="354">Description.</emphasis>
Adult, winged, rather robust. Female, defined on the basis of the large last sternite. Body length 3.0 mm. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFB2FFF3FD19BF375D0AA329" blockId="23.[643,1135,517,1683]" lastBlockId="25.[113,605,155,601]" lastPageId="25" lastPageNumber="356" pageId="23" pageNumber="354">Head very transverse, constricted (triangular­shaped) behind the eyes, surface irregular with shallow punctation. Eyes prominent, very large, sub­elliptical, inserted in the lateral part of the head. Mandibles elongated, falciform. Maxillary palps 4­ segmented with the last palpomere elongated and securiform. Labial palps 3­ segmented, with the last palpomere securiform. Antennae 11­ segmented, pectinate, long, slightly surpassing the apex of elytra, antennal insertion far from the eyes; scape club­shaped, elongated, very robust; antennomeres IIIII short, subequal in length, swollen apically; antennomeres IV VII extremely long, each equipped at apex with an antennal process which is very long and thin with an emarginate apex; antennomeres VIII X shorter than previous ones, not enlarged and robust, without the antennal process at apex; antennomere XI filiform, not enlarged and robust, long, with rounded apex; all the antennomeres slightly pubescent. Pronotum very transverse, approximately as wide as head, anterior margin undulate and slightly bordered, posterior margin almost straight and feebly bordered, sides rounded, corners obtuse, pronotum disc irregular with very shallow punctation. Scutellum narrow, triangular­shaped, pointed at apex. Elytra short which reveals some abdominal segments, at humeri wider than pronotum, posteriorly parallel­sided and very slightly dehiscent apically, rounded apexes, surface presenting a very coarse and wide punctation gathered in evident striae visible for the entire elytral length. Metathoracic wings transparent, longer than elytra, not reaching the last abdominal segment. Sternum very robust with margin almost straight; sternites transverse, wide and pubescent; last sternite very large and robust, rounded. Legs rather robust, relatively long, slightly pubescent; coxae massive; trochanters very elongated, robust, with rounded apex; femora almost straight, sub­cylindrical, slightly sturdier than tibiae; tibiae, thin, sub­cylindrical, with a small spur near the apex, protibiae approximately as long as profemora, mesotibiae longer than mesofemora, metatibiae shorter than metafemora. Tarsal formula 5­ 5­ 5; first metatarsomere extremely long, about 2.3 times longer than second; third tarsomere short; fourth strongly bilobed; fifth tarsomere elongated, thin, cylindrical and curved; claws simple, pointed, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFBDFFF2FF0DBB565DA7A7EB" pageId="24" pageNumber="355" startId="24.[151,190,1636,1659]" targetBox="[261,1066,151,1606]" targetPageId="24" targetType="figure">
<paragraph id="8BCC643DFFBDFFF2FF0DBB565DA7A7EB" blockId="24.[151,1172,1636,1691]" pageId="24" pageNumber="355">
Fig. 13.
<taxonomicName id="4C731FBEFFBDFFF2FF77BB565D25A70B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[237,611,1636,1659]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="355" phylum="Arthropoda" rank="species" species="kirstenaeweissbachae" status="sp. nov.">
<emphasis id="B907B82FFFBDFFF2FF77BB565D25A70B" box="[237,611,1636,1659]" italics="true" pageId="24" pageNumber="355">Sanaungulus kirstenaeweissbachae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBDFFF2FDF2BB565DFEA70B" box="[616,696,1636,1659]" pageId="24" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of head.
</paragraph>
</caption>
<subSubSection id="C36937B6FFBCFFF3FFEBBF4B5D18A398" pageId="25" pageNumber="356" type="etymology">
<paragraph id="8BCC643DFFBCFFF3FFEBBF4B5D1BA3C1" blockId="25.[113,605,633,689]" pageId="25" pageNumber="356">
<emphasis id="B907B82FFFBCFFF3FFEBBF4B5FB4A3E1" bold="true" box="[113,242,633,657]" pageId="25" pageNumber="356">Etymology.</emphasis>
Named in honor of Kirsten Weissbach, dear friend of the second author (Patrick).
</paragraph>
<paragraph id="8BCC643DFFBCFFF3FFEBBFE35D18A398" blockId="25.[113,606,721,745]" box="[113,606,721,745]" pageId="25" pageNumber="356">
<emphasis id="B907B82FFFBCFFF3FFEBBFE35E4CA399" bold="true" box="[113,266,721,745]" pageId="25" pageNumber="356">Syninclusions.</emphasis>
Air bubbles, detritus, anAlydidae.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBCFFF3FFEBBE3A5F9FA2F1" pageId="25" pageNumber="356" type="discussion">
<paragraph id="8BCC643DFFBCFFF3FFEBBE3A5F9FA2F1" blockId="25.[113,605,776,897]" pageId="25" pageNumber="356">
<emphasis id="B907B82FFFBCFFF3FFEBBE3A5F98A250" bold="true" box="[113,222,776,800]" pageId="25" pageNumber="356">Remarks.</emphasis>
Piece of clear, golden amber in Cabochon that measures 14x10x
<quantity id="4C8BC9D8FFBCFFF3FE13BE1B5E81A230" box="[393,455,809,832]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="356" unit="mm" value="4.0">4 mm</quantity>
. The inclusion is complete, with the head partially covered by emulsion.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,147 @@
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<mods:title id="FD400DD29133C9B555DC47F09A2B1438">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="CE9D84E68BD0BCFBEADBE04791DBCA85">Fanti, Fabrizio</mods:namePart>
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<mods:date id="01237B47B16EC2F6AF6EFD5D2B69779E">2022</mods:date>
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<subSubSection id="C36937B6FFB5FFFAFF0DB8615EAAA41B" box="[151,492,1363,1387]" pageId="16" pageNumber="347" type="nomenclature">
<paragraph id="8BCC643DFFB5FFFAFF0DB8615EAAA41B" blockId="16.[151,492,1363,1418]" box="[151,492,1363,1387]" pageId="16" pageNumber="347">
<heading id="D084D351FFB5FFFAFF0DB8615EAAA41B" bold="true" box="[151,492,1363,1387]" fontSize="10" level="2" pageId="16" pageNumber="347" reason="4">
<emphasis id="B907B82FFFB5FFFAFF0DB8615EAAA41B" bold="true" box="[151,492,1363,1387]" pageId="16" pageNumber="347">
<taxonomicName id="4C731FBEFFB5FFFAFF0DB8615ED7A41B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[151,401,1363,1387]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="347" phylum="Arthropoda" rank="species" species="electrum" status="sp. nov.">
<emphasis id="B907B82FFFB5FFFAFF0DB8615ED7A41B" bold="true" box="[151,401,1363,1387]" italics="true" pageId="16" pageNumber="347">Sanaungulus electrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB5FFFAFE0DB8615EAAA41B" box="[407,492,1363,1387]" pageId="16" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB5FFFAFF0DB8415FA1A4FA" box="[151,231,1395,1418]" pageId="16" pageNumber="347" type="description">
<paragraph id="8BCC643DFFB5FFFAFF0DB8415FA1A4FA" blockId="16.[151,492,1363,1418]" box="[151,231,1395,1418]" pageId="16" pageNumber="347">
(
<figureCitation id="134878B8FFB5FFFAFF05B8415F99A4FA" box="[159,223,1395,1418]" captionStart="Fig" captionStartId="17.[113,152,1638,1661]" captionTargetBox="[113,1134,385,1610]" captionTargetId="figure-30@17.[113,1134,385,1610]" captionTargetPageId="17" captionText="Fig. 9. Sanaungulus electrum sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view." pageId="16" pageNumber="347">Fig. 9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB5FFFAFF0DB8985C81A1A2" pageId="16" pageNumber="347" type="materials_examined">
<paragraph id="8BCC643DFFB5FFFAFF0DB8985E53A731" blockId="16.[151,644,1450,1601]" pageId="16" pageNumber="347">
<materialsCitation id="3B1B6E60FFB5FFFAFF0DB8985E53A731" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="16" pageNumber="347" specimenCode="BUB4325" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFB5FFFAFF0DB8985E40A4B2" bold="true" box="[151,262,1450,1474]" pageId="16" pageNumber="347">
<typeStatus id="54C8DA9FFFB5FFFAFF0DB8985E47A4B2" box="[151,257,1450,1474]" pageId="16" pageNumber="347" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFB5FFFAFE96B8985E03A4B1" box="[268,325,1450,1473]" pageId="16" pageNumber="347">Male</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFB5FFFAFDBEB8985DC5A4B1" box="[548,643,1450,1473]" pageId="16" pageNumber="347">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFB5FFFAFF05B8F85FA8A491" box="[159,238,1482,1505]" country="Myanmar" name="Kachin" pageId="16" pageNumber="347">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFB5FFFAFE19B8F85E93A491" LSID="urn:lsid:plazi:treatment:03DAD52BFFB5FFF8FF0DB8615E9FA3F0:8EAC32E6FFB5FFFAFE19B8F85E93A491" box="[387,469,1482,1505]" county="Male" municipality="Burmese" name="Catalog" pageId="16" pageNumber="347" stateProvince="Kachin">Catalog</location>
number
<quantity id="4C8BC9D8FFB5FFFAFDB4B8F85FA4A771" metricMagnitude="2" metricUnit="m" metricValue="1.52527" pageId="16" pageNumber="347" unit="in" value="6005.0">SNHM­6005 in</quantity>
the
<location id="8EAC32E6FFB5FFFAFE92B8D85E69A751" LSID="urn:lsid:plazi:treatment:03DAD52BFFB5FFF8FF0DB8615E9FA3F0:8EAC32E6FFB5FFFAFE92B8D85E69A751" county="Male" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="16" pageNumber="347" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FFB5FFFAFEC6BB385EF1A751" LSID="urn:lsid:plazi:treatment:03DAD52BFFB5FFF8FF0DB8615E9FA3F0:8EAC32E6FFB5FFFAFEC6BB385EF1A751" box="[348,439,1546,1569]" county="Male" municipality="Burmese" name="Muller's" pageId="16" pageNumber="347" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFB5FFFAFF0DBB185E4FA731" box="[151,265,1578,1601]" collectionCode="BUB" pageId="16" pageNumber="347">BUB4325</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFB5FFFAFF0DBB525EACA7E8" blockId="16.[151,643,1632,1688]" pageId="16" pageNumber="347">
<emphasis id="B907B82FFFB5FFFAFF0DBB525E76A708" bold="true" box="[151,304,1632,1656]" pageId="16" pageNumber="347">Type locality.</emphasis>
<collectingCountry id="F36424ADFFB5FFFAFEA2BB535EE2A708" box="[312,420,1633,1656]" name="Myanmar" pageId="16" pageNumber="347">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFB5FFFAFE2ABB535D7CA708" box="[432,570,1633,1656]" country="Myanmar" name="Kachin" pageId="16" pageNumber="347">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFB5FFFAFD32BDA85C81A1A2" blockId="16.[680,1172,154,210]" pageId="16" pageNumber="347">
<emphasis id="B907B82FFFB5FFFAFD32BDA85C7AA1C2" bold="true" box="[680,828,154,178]" pageId="16" pageNumber="347">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB5FFFAFD32BDC35C7DA0D9" pageId="16" pageNumber="347" type="diagnosis">
<paragraph id="8BCC643DFFB5FFFAFD32BDC35C7DA0D9" blockId="16.[680,1172,241,425]" pageId="16" pageNumber="347">
<emphasis id="B907B82FFFB5FFFAFD32BDC35CD3A079" bold="true" box="[680,917,241,265]" pageId="16" pageNumber="347">Differential diagnosis.</emphasis>
The new species is easily distinguishable by the pronotal shape, with the sides concave at the center and with an expansion/tooth near the first apical third. These features are not present in other species of the genus
<emphasis id="B907B82FFFB5FFFAFD32BCA05C7DA0D9" box="[680,827,402,425]" italics="true" pageId="16" pageNumber="347">
<taxonomicName id="4C731FBEFFB5FFFAFD32BCA05C70A0D9" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[680,822,402,425]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="347" phylum="Arthropoda" rank="genus">Sanaungulus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB5FFF8FD32BCFA5EBAA184" lastPageId="18" lastPageNumber="349" pageId="16" pageNumber="347" type="description">
<paragraph id="8BCC643DFFB5FFFAFD32BCFA5C84A32F" blockId="16.[680,1172,456,607]" pageId="16" pageNumber="347">
<emphasis id="B907B82FFFB5FFFAFD32BCFA5C75A090" bold="true" box="[680,819,456,480]" pageId="16" pageNumber="347">Description.</emphasis>
Adult, winged, slender. Male, defined on the shape of the last tergite that is very elongated, and on the basis of the rounded last sternite. Body length about
<quantity id="4C8BC9D8FFB5FFFAFC4BBF1A5B17A34F" box="[977,1105,552,575]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="16" pageNumber="347" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.93.1 mm</quantity>
(head is folded). Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFB5FFF8FD32BF4D5EBAA184" blockId="16.[680,1173,639,1686]" lastBlockId="18.[151,643,155,244]" lastPageId="18" lastPageNumber="349" pageId="16" pageNumber="347">Head elongated, strongly constricted (triangular­shaped) behind the eyes, fitted with rugosity and punctuation.Eyes prominent, large, roundish, inserted in the upper and dorsolateral part of the head. Mandibles elongated, robust, falciform. Antennae 11­ segmented, filiform, slender, very short and slightly surpassing the humeral zone of the elytra; scape club­shaped; antennomere II very short, globular; antennomeres IIIV, thin, very long, over 2 times longer than second; antennomeres VIX subequal in length, about 2.0 times shorter than previous ones; antennomere XI filiform, slightly longer than previous ones, rounded at apex; all the antennomeres with short pubescence. Pronotum subquadrate, slightly transverse, anterior margin straight and not bordered, posterior margin rather straight, corners obtuse, sides concave at the center and slightly expanded posteriorly and with an expansion/ tooth near the first apical third, pronotum disc smooth. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, wider than pronotum, at humery enlarged and slightly dehiscent posteriorly, rounded apexes, surface smooth without rugosity and only sligthly pubescent. Metathoracic wings transparent, long, surpassing the elytra and reaching the last abdominal segments. Sternum elongated with posterior margin irregular with an expansion at the center that is very small and forked apically, sternites wide and transverse equipped with short setae; penultimate tergite with a long and thin lobe at sides that is curved; last tergite very elongated, initially wide and flat, deeply folded apically, with the apical margin forked and globular at sides; last sternite smaller than last tergite and rounded. Legs thin, relatively short, pubescent; coxae massive, large; trochanters short, globular with rounded apex; femora nearly straight, more robust than tibiae, cylindrical; tibiae cylindrical, thin, without spurs near the apex, metatibiae very longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, second tarsomere similar in size to previous one; third tarsomere short and triangular­shaped; fourth strongly bilobed and very long; fifth tarsomere elongated, thin, slightly flat and curved; claws simple, apparently without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFB4FFFBFFEBBB545FBCA7ED" pageId="17" pageNumber="348" startId="17.[113,152,1638,1661]" targetBox="[113,1134,385,1610]" targetPageId="17" targetType="figure">
<paragraph id="8BCC643DFFB4FFFBFFEBBB545FBCA7ED" blockId="17.[113,1134,1638,1693]" pageId="17" pageNumber="348">
Fig. 9.
<taxonomicName id="4C731FBEFFB4FFFBFF21BB555EE1A70E" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[187,423,1639,1662]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="348" phylum="Arthropoda" rank="species" species="electrum" status="sp. nov.">
<emphasis id="B907B82FFFB4FFFBFF21BB555EE1A70E" box="[187,423,1639,1662]" italics="true" pageId="17" pageNumber="348">Sanaungulus electrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB4FFFBFE37BB545EB8A70D" box="[429,510,1638,1661]" pageId="17" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFB7FFF8FF0DBC265D59A0D4" pageId="18" pageNumber="349" type="etymology">
<paragraph id="8BCC643DFFB7FFF8FF0DBC265D62A01D" blockId="18.[151,643,276,365]" pageId="18" pageNumber="349">
<emphasis id="B907B82FFFB7FFF8FF0DBC265E50A05C" bold="true" box="[151,278,276,300]" pageId="18" pageNumber="349">Etymology.</emphasis>
Derived from the Latin noun “Á
<emphasis id="B907B82FFFB7FFF8FDC1BC265F81A03C" italics="true" pageId="18" pageNumber="349">lectrum</emphasis>
” = yellow amber, succinite. Specific epithet is to be treated as noun in apposition.
</paragraph>
<paragraph id="8BCC643DFFB7FFF8FF0DBCBF5D59A0D4" blockId="18.[151,543,397,421]" box="[151,543,397,421]" pageId="18" pageNumber="349">
<emphasis id="B907B82FFFB7FFF8FF0DBCBF5E7CA0D5" bold="true" box="[151,314,397,421]" pageId="18" pageNumber="349">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB7FFF8FF0DBCF65E9FA3F0" pageId="18" pageNumber="349" type="discussion">
<paragraph id="8BCC643DFFB7FFF8FF0DBCF65E9FA3F0" blockId="18.[151,643,452,640]" pageId="18" pageNumber="349">
<emphasis id="B907B82FFFB7FFF8FF0DBCF65E45A0AC" bold="true" box="[151,259,452,476]" pageId="18" pageNumber="349">Remarks.</emphasis>
Piece of clear, golden amber that measures 12x6x
<quantity id="4C8BC9D8FFB7FFF8FE8EBCD45E15A08D" box="[276,339,486,509]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="349" unit="mm" value="2.0">2 mm</quantity>
. The inclusion lacks four legs and there is only one antenna that is completely detached from the body. Furthermore, the head (and its frons) in the ventral part is covered by an emulsion like white­cotton.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,149 @@
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<mods:title id="FE3D626FFD835817132BFE2F1D123078">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<taxonomicName id="4C731FBEFFB6FFF9FD19BC875C1FA0BD" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[643,857,437,461]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="350" phylum="Arthropoda" rank="species" species="havai" status="sp. nov.">
<emphasis id="B907B82FFFB6FFF9FD19BC875C1FA0BD" bold="true" box="[643,857,437,461]" italics="true" pageId="19" pageNumber="350">Sanaungulus havai</emphasis>
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</emphasis>
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(
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)
</paragraph>
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<materialsCitation id="3B1B6E60FFB6FFF9FD19BF3E5C46A3D3" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="19" pageNumber="350" specimenCode="BUB4453" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFB6FFF9FD19BF3E5DB7A354" bold="true" box="[643,753,524,548]" pageId="19" pageNumber="350">
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.
</emphasis>
<collectingCounty id="62AD1CB1FFB6FFF9FD6DBF3E5C76A353" box="[759,816,524,547]" pageId="19" pageNumber="350">Male</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFB6FFF9FB95BF3E5B28A353" box="[1039,1134,524,547]" pageId="19" pageNumber="350">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFB6FFF9FD10BF1E5D9FA333" box="[650,729,556,579]" country="Myanmar" name="Kachin" pageId="19" pageNumber="350">Kachin</collectingRegion>
) amber piece:
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number
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the
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(ex
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collection number:
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).
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<emphasis id="B907B82FFFB6FFF9FD19BFF05C5AA3AA" bold="true" box="[643,796,706,730]" pageId="19" pageNumber="350">Type locality.</emphasis>
<collectingCountry id="F36424ADFFB6FFF9FCBEBFF15CD6A3AA" box="[804,912,707,730]" name="Myanmar" pageId="19" pageNumber="350">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFB6FFF9FC01BFF15B63A3AA" box="[923,1061,707,730]" country="Myanmar" name="Kachin" pageId="19" pageNumber="350">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
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<emphasis id="B907B82FFFB6FFF9FD19BE2B5C50A241" bold="true" box="[643,790,793,817]" pageId="19" pageNumber="350">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
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<emphasis id="B907B82FFFB6FFF9FD19BE425C31A2F8" bold="true" box="[643,887,880,904]" pageId="19" pageNumber="350">Differential diagnosis.</emphasis>
The new species is characterized by antennae equipped with an antennal process near the base of antennomeres IV X, a feature not present in other
<taxonomicName id="4C731FBEFFB6FFF9FC7EBEE25B28A297" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[996,1134,976,999]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFB6FFF9FC7EBEE25B28A297" box="[996,1134,976,999]" italics="true" pageId="19" pageNumber="350">Sanaungulus</emphasis>
</taxonomicName>
except for the species described below (see differential diagnosis below).
</paragraph>
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<paragraph id="8BCC643DFFB6FFF9FD19B9745B61A5CE" blockId="19.[643,1134,1094,1214]" pageId="19" pageNumber="350">
<emphasis id="B907B82FFFB6FFF9FD19B9745C4BA52E" bold="true" box="[643,781,1094,1118]" pageId="19" pageNumber="350">Description.</emphasis>
Adult, winged, slender. Male, defined on the shape of the last abdominal segments. Body length about
<quantity id="4C8BC9D8FFB6FFF9FC0FB9B55CAEA5EE" box="[917,1000,1159,1182]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="19" pageNumber="350" unit="mm" value="3.2">3.2 mm</quantity>
. Testaceous­brown with black head behind the eyes.
</paragraph>
<paragraph id="8BCC643DFFB6FFFFFD19B9EF5EA8A422" blockId="19.[643,1134,1245,1684]" lastBlockId="21.[113,605,155,1362]" lastPageId="21" lastPageNumber="352" pageId="19" pageNumber="350">Head elongated, strongly constricted (triangular­shaped) behind the eyes, fitted with small rugosity and shallow punctuation. Eyes very prominent, roundish, inserted in the upper and dorsolateral part of the head, inter­ocular dorsal distance about 4.1 times greater than eye diameter. Mandibles elongated, falciform. Maxillary palps 4­ segmented, with the last palpomere securiform. Antennae 11­ segmented, pectinate, slen­ der, long, almost reaching the apex of the elytra; scape very elongated and enlarged apically; antennomere II short, globular; antennomere III thin, filiform, about 2.1 times longer than second; antennomeres IVVII subequal in length, each equipped near their base with a long, thin and curved antennal process, squared apically; antennomeres VIIIX, slightly shorter and stouter than previous ones, each equipped near their base with a long antennal process (stouter than previous ones), almost rounded apically; antennomere XI filiform and robust; antennomeres pubescent. Pronotum longer than wide, slightly wider than head, anterior margin rounded and bordered, posterior margin rather straight, corners obtuse, sides narrower anteriorly, pronotum disc irregular and slightly wrinkled with impressed punctuation. Scutellum triangular­shaped with rounded apex. Elytra short which reveals two abdominal segments, wider than pronotum, at humery enlarged and very slightly dehiscent posteriorly, rounded apexes, surface with strongly impressed punctuation gathered in striae. Metathoracic wings transparent, long, surpassing the elytra and the last abdominal segments. Sternum elongated with posterior margin irregular, sternites transverse equipped with short setae; last tergite elongated, wide and curved and very slightly folded at the apical sides; penultimate sternite with an urophysis at sides wich is rather robust and evident and rounded at apex; last sternite very elongated, curved and thin. Legs thin, very long, pubescent; coxae massive; trochanters elongated; femora nearly straight, more robust than tibiae, cylindrical; tibiae cylindrical, thin, without spurs near the apex, very longer than femora. Tarsal formula 5­ 5­ 5; first tarsomere extremely long and thin; second tarsomere about 2.0 times shorter than first; third tarsomere short and triangular­shaped; fourth strongly bilobed with the lobes very long; fifth tarsomere elongated, thin, slightly flat and curved; claws simple, without denticle at the base.</paragraph>
</subSubSection>
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<paragraph id="8BCC643DFFB1FFFEFF0DBB535D04A7E8" blockId="20.[151,1172,1633,1688]" pageId="20" pageNumber="351">
Fig. 11.
<taxonomicName id="4C731FBEFFB1FFFEFF6ABB535EFAA708" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[240,444,1633,1656]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="351" phylum="Arthropoda" rank="species" species="havai" status="sp. nov.">
<emphasis id="B907B82FFFB1FFFEFF6ABB535EFAA708" box="[240,444,1633,1656]" italics="true" pageId="20" pageNumber="351">Sanaungulus havai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB1FFFEFE59BB535D50A708" box="[451,534,1633,1656]" pageId="20" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, lateral view; C: Holotype, detail of legs.
</paragraph>
</caption>
<subSubSection id="C36937B6FFB0FFFFFFEBB8435D18A48F" pageId="21" pageNumber="352" type="etymology">
<paragraph id="8BCC643DFFB0FFFFFFEBB8435FB9A4B8" blockId="21.[113,605,1393,1480]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFFEBB8435FA8A4F9" bold="true" box="[113,238,1393,1417]" pageId="21" pageNumber="352">Etymology.</emphasis>
Named in honor of Jir í Háva (
<collectingCountry id="F36424ADFFB0FFFFFD81B8435F95A4D8" name="Czech Republic" pageId="21" pageNumber="352">Czech Republic</collectingCountry>
), a well­known specialist of the family
<taxonomicName id="4C731FBEFFB0FFFFFFEBB8835FBDA4B8" box="[113,251,1457,1480]" class="Insecta" family="Dermestidae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="352" phylum="Arthropoda" rank="family">Dermestidae</taxonomicName>
.
</paragraph>
<paragraph id="8BCC643DFFB0FFFFFFEBB8DA5D18A48F" blockId="21.[113,606,1512,1536]" box="[113,606,1512,1536]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFFEBB8DA5E4FA770" bold="true" box="[113,265,1512,1536]" pageId="21" pageNumber="352">Syninclusions.</emphasis>
Air bubbles, detritus, stellate hairs.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB0FFFFFFEBBB2C5E7CA7E6" pageId="21" pageNumber="352" type="discussion">
<paragraph id="8BCC643DFFB0FFFFFFEBBB2C5E7CA7E6" blockId="21.[113,605,1566,1686]" pageId="21" pageNumber="352">
<emphasis id="B907B82FFFB0FFFFFFEBBB2C5FA7A746" bold="true" box="[113,225,1566,1590]" pageId="21" pageNumber="352">Remarks.</emphasis>
Piece of clear, golden amber that measures 8x7x
<quantity id="4C8BC9D8FFB0FFFFFE94BB0D5E0DA726" box="[270,331,1599,1622]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="352" unit="mm" value="3.0">3 mm</quantity>
. The inclusion is complete except for the left posterior leg preserved until the first tarsomere.
</paragraph>
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</treatment>
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<paragraph id="8BCC643DFFB7FFF8FF0DBF855DC5A3BF" blockId="18.[151,643,695,751]" box="[151,643,695,719]" pageId="18" pageNumber="349">
<heading id="D084D351FFB7FFF8FF0DBF855DC5A3BF" bold="true" box="[151,643,695,719]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="349" reason="4">
<emphasis id="B907B82FFFB7FFF8FF0DBF855DC5A3BF" bold="true" box="[151,643,695,719]" pageId="18" pageNumber="349">
<taxonomicName id="4C731FBEFFB7FFF8FF0DBF855D6FA3BF" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[151,553,695,719]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="franziskaeweissbachae" status="sp. nov.">
<emphasis id="B907B82FFFB7FFF8FF0DBF855D6FA3BF" bold="true" box="[151,553,695,719]" italics="true" pageId="18" pageNumber="349">Sanaungulus franziskaeweissbachae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB7FFF8FDB5BF855DC5A3BF" box="[559,643,695,719]" pageId="18" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB7FFF8FF0DBFEA5FB2A39F" box="[151,244,728,751]" pageId="18" pageNumber="349" type="description">
<paragraph id="8BCC643DFFB7FFF8FF0DBFEA5FB2A39F" blockId="18.[151,643,695,751]" box="[151,244,728,751]" pageId="18" pageNumber="349">
(
<figureCitation id="134878B8FFB7FFF8FF05BFEA5FAAA39F" box="[159,236,728,751]" captionStart="Fig" captionStartId="18.[151,190,1665,1688]" captionTargetBox="[151,1172,1023,1635]" captionTargetId="figure-344@18.[151,1172,1023,1635]" captionTargetPageId="18" captionText="Fig. 10. Sanaungulus franziskaeweissbachae sp. nov. in Burmese amber. Holotype, lateral view." pageId="18" pageNumber="349">Fig. 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB7FFF8FF0DBE3D5C81A058" pageId="18" pageNumber="349" type="materials_examined">
<paragraph id="8BCC643DFFB7FFF8FF0DBE3D5F98A2BA" blockId="18.[151,643,783,970]" pageId="18" pageNumber="349">
<materialsCitation id="3B1B6E60FFB7FFF8FF0DBE3D5F98A2BA" collectingDate="2021-12" collectionCode="BUB" county="Probably" location="Catalog" municipality="Burmese" pageId="18" pageNumber="349" specimenCode="BUB 3899" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFB7FFF8FF0DBE3D5E40A257" bold="true" box="[151,262,783,807]" pageId="18" pageNumber="349">
<typeStatus id="54C8DA9FFFB7FFF8FF0DBE3D5E47A257" box="[151,257,783,807]" pageId="18" pageNumber="349" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFB7FFF8FE8ABE3D5E37A256" box="[272,369,783,806]" pageId="18" pageNumber="349">Probably</collectingCounty>
male, adult specimen in a
<collectingMunicipality id="6BA8FE47FFB7FFF8FF35BE025E54A237" box="[175,274,816,839]" pageId="18" pageNumber="349">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFB7FFF8FEBDBE025E3EA237" box="[295,376,816,839]" country="Myanmar" name="Kachin" pageId="18" pageNumber="349">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFB7FFF8FDB6BE025DC5A237" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FDB6BE025DC5A237" box="[556,643,816,839]" county="Probably" municipality="Burmese" name="Catalog" pageId="18" pageNumber="349" stateProvince="Kachin">Catalog</location>
number SNSB­BSPG
<date id="FFCD42FDFFB7FFF8FE1EBE635EA3A218" box="[388,485,849,872]" pageId="18" pageNumber="349" value="2021-12">
<collectingDate id="EF89BB15FFB7FFF8FE1EBE635EA3A218" box="[388,485,849,872]" pageId="18" pageNumber="349" value="2021-12">2021 XII</collectingDate>
</date>
<quantity id="4C8BC9D8FFB7FFF8FE71BE635D64A218" box="[491,546,849,872]" metricMagnitude="0" metricUnit="m" metricValue="1.8288" pageId="18" pageNumber="349" unit="in" value="72.0">72 in</quantity>
the
<location id="8EAC32E6FFB7FFF8FDD5BE635ED1A2F9" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FDD5BE635ED1A2F9" county="Probably" municipality="Burmese" name="Bayerische Staatssammlung" pageId="18" pageNumber="349" stateProvince="Kachin">Bayerische Staatssammlung</location>
für
<location id="8EAC32E6FFB7FFF8FE58BE405D13A2F9" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FE58BE405D13A2F9" box="[450,597,882,905]" county="Probably" municipality="Burmese" name="Palaontologie" pageId="18" pageNumber="349" stateProvince="Kachin">Paläontologie</location>
und
<location id="8EAC32E6FFB7FFF8FF0DBEA15FBEA2DA" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FF0DBEA15FBEA2DA" box="[151,248,915,938]" county="Probably" municipality="Burmese" name="Geologie" pageId="18" pageNumber="349" stateProvince="Kachin">Geologie</location>
(ex
<location id="8EAC32E6FFB7FFF8FEBEBEA15E38A2DA" LSID="urn:lsid:plazi:treatment:03DAD52BFFB7FFF9FF0DBF855B20A00F:8EAC32E6FFB7FFF8FEBEBEA15E38A2DA" box="[292,382,915,938]" county="Probably" municipality="Burmese" name="Muller's" pageId="18" pageNumber="349" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFB7FFF8FDD6BEA15F97A2BA" collectionCode="BUB" pageId="18" pageNumber="349">BUB 3899</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFB7FFF8FD32BDA85CBDA1A2" blockId="18.[680,1172,154,210]" pageId="18" pageNumber="349">
<emphasis id="B907B82FFFB7FFF8FD32BDA85C07A1C2" bold="true" box="[680,833,154,178]" pageId="18" pageNumber="349">Type locality.</emphasis>
<collectingCountry id="F36424ADFFB7FFF8FCD0BDA95CF0A1C2" box="[842,950,155,178]" name="Myanmar" pageId="18" pageNumber="349">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFB7FFF8FC5BBDA95B0DA1C2" box="[961,1099,155,178]" country="Myanmar" name="Kachin" pageId="18" pageNumber="349">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFB7FFF8FD32BDC35C81A058" blockId="18.[680,1172,241,296]" pageId="18" pageNumber="349">
<emphasis id="B907B82FFFB7FFF8FD32BDC35C7AA079" bold="true" box="[680,828,241,265]" pageId="18" pageNumber="349">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB7FFF8FD32BC7A5C3BA32F" pageId="18" pageNumber="349" type="diagnosis">
<paragraph id="8BCC643DFFB7FFF8FD32BC7A5C3BA32F" blockId="18.[680,1172,328,607]" pageId="18" pageNumber="349">
<emphasis id="B907B82FFFB7FFF8FD32BC7A5CE0A010" bold="true" box="[680,934,328,352]" pageId="18" pageNumber="349">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFFB7FFF8FC35BC7A5C12A00F" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="franziskaeweissbachae" status="sp. nov.">
<emphasis id="B907B82FFFB7FFF8FC35BC7A5C12A00F" italics="true" pageId="18" pageNumber="349">Sanaungulus franziskaeweissbachae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB7FFF8FCC0BC5A5CECA00F" box="[858,938,360,383]" pageId="18" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C731FBEFFB7FFF8FBF9BC5A5C78A0EF" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="nilsi">
<emphasis id="B907B82FFFB7FFF8FBF9BC5A5C78A0EF" italics="true" pageId="18" pageNumber="349">Sanaungulus nilsi</emphasis>
</taxonomicName>
(described in the present work), but the new species differs based on its more robost second antennomere and much longer metathoracic wings. Based on these characters and others of lesser importance, the
<specimenCount id="9D75AFB4FFB7FFF8FBBCBF3A5D99A34F" count="2" pageId="18" pageNumber="349" type="generic">two specimens</specimenCount>
are not considered here as male and female of the same species.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB7FFF9FD32BF4C5EA8A7E2" lastPageId="19" lastPageNumber="350" pageId="18" pageNumber="349" type="description">
<paragraph id="8BCC643DFFB7FFF8FD32BF4C5C09A386" blockId="18.[680,1172,638,758]" pageId="18" pageNumber="349">
<emphasis id="B907B82FFFB7FFF8FD32BF4C5C72A3E6" bold="true" box="[680,820,638,662]" pageId="18" pageNumber="349">Description.</emphasis>
Adult, winged, slender. Probably male, defined on the basis of the last abdominal segment triangular shaped. Body length
<quantity id="4C8BC9D8FFB7FFF8FBDABF8D5BD6A3A6" box="[1088,1168,703,726]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="349" unit="mm" value="2.8">2.8 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFB7FFF9FD32BE275EA8A7E2" blockId="18.[680,1172,789,972]" lastBlockId="19.[113,605,155,1682]" lastPageId="19" lastPageNumber="350" pageId="18" pageNumber="349">Head elongated, slightly constricted (triangular­shaped) behind the eyes, equipped with pubescence and shallow punctation. Eyes prominent, extremely large, elongated, inserted in the lateral part of the head.Mandibles elongated, very robust, strongly curved, falciform, pointed apicallly, without teeth. Maxillary palps 4­ segmented with the last palpomere very elongated and securiform. Labial palps 3­ segmented, with the last palpomere elongated and securiform.Antennae 11­ segmented, filiform, rather robust, short, approximately reaching the half of elytra, antennal insertion far from the eyes; scape robust, very elongated, slightly club­shaped; antennomeres IIIII short, thinner than first, antennomere II suboval and more robust than antennomere III; antennomeres IVIX subequal, filiform, very elongated; antennomere X slightly shorter than previous ones; antennomere XI filiform, rounded at apex; all the antennomeres with short pubescence. Pronotum very elongated, slightly narrower than head, anterior margin slightly roundish, posterior margin slightly undulated and slightly bordered, sides straight and bordered, corners obtuse, pronotum disc pubescent and slighlty convex near the posterior margin. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, at humeri slightly wider than pronotum, posteriorly parallel­sided and rather dehiscent apically, rounded apexes, surface pubescent and with a very small rugosity. Metathoracic wings transparent, very long, longer than elytra and surpassing the last abdominal segments. Sternum elongated with posterior margin rounded, sternites narrow and transverse, last tergite elongated with pointed apex which has the margin rounded, last sternite shorter and narrower than last tergite with pointed apex.Legs thin, extremely long, pubescent; coxae extraordinary massive, elongated, very enlarged; trochanters elongated, with rounded apex; femora slightly curved, more robust than tibiae, sub­cylindrical; tibiae flattened, thin, with a small and obtuse spur near the apex, protibiae very shorter than profemora, meso­ and metatibiae very longer than meso­ and metafemora. Tarsal formula 5­ 5­ 5; first protarsomere long, about 3.0 times longer than second; second protarsomere robust, enlarged apically; third protarsomere short, bilobed with pointed apex; fourth protarsomere strongly bilobed with a pointed and short lobe at each corners; fifth protarsomere elongated, thin, flat and curved; claws simple, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFB7FFF8FF0DBBB35BD2A7E8" box="[151,1172,1665,1688]" pageId="18" pageNumber="349" startId="18.[151,190,1665,1688]" targetBox="[151,1172,1023,1635]" targetPageId="18" targetType="figure">
<paragraph id="8BCC643DFFB7FFF8FF0DBBB35BD2A7E8" blockId="18.[151,1172,1665,1688]" box="[151,1172,1665,1688]" pageId="18" pageNumber="349">
Fig. 10.
<taxonomicName id="4C731FBEFFB7FFF8FF6ABBB35D33A7E8" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[240,629,1665,1688]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="franziskaeweissbachae" status="sp. nov.">
<emphasis id="B907B82FFFB7FFF8FF6ABBB35D33A7E8" box="[240,629,1665,1688]" italics="true" pageId="18" pageNumber="349">Sanaungulus franziskaeweissbachae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB7FFF8FDE1BBB35D8BA7E8" box="[635,717,1665,1688]" pageId="18" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. Holotype, lateral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFB6FFF9FD19BDA85B4DA078" pageId="19" pageNumber="350" type="etymology">
<paragraph id="8BCC643DFFB6FFF9FD19BDA85B29A1A2" blockId="19.[643,1135,154,210]" pageId="19" pageNumber="350">
<emphasis id="B907B82FFFB6FFF9FD19BDA85C46A1C2" bold="true" box="[643,768,154,178]" pageId="19" pageNumber="350">Etymology.</emphasis>
Named in honor of Franziska Weissbach, dear friend of the second author (Patrick).
</paragraph>
<paragraph id="8BCC643DFFB6FFF9FD19BDC35B4DA078" blockId="19.[643,1035,241,265]" box="[643,1035,241,265]" pageId="19" pageNumber="350">
<emphasis id="B907B82FFFB6FFF9FD19BDC35C60A079" bold="true" box="[643,806,241,265]" pageId="19" pageNumber="350">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB6FFF9FD19BC1A5B20A00F" pageId="19" pageNumber="350" type="discussion">
<paragraph id="8BCC643DFFB6FFF9FD19BC1A5B20A00F" blockId="19.[643,1134,296,383]" pageId="19" pageNumber="350">
<emphasis id="B907B82FFFB6FFF9FD19BC1A5DB4A030" bold="true" box="[643,754,296,320]" pageId="19" pageNumber="350">Remarks.</emphasis>
Piece of clear, golden amber that measures 11x10x
<quantity id="4C8BC9D8FFB6FFF9FCA7BC7A5C38A02F" box="[829,894,328,351]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="350" unit="mm" value="2.0">2 mm</quantity>
.The inclusion is complete, but well visible only in the lateral view.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,151 @@
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<mods:title id="B2E9551354F167C494B13777ABF9BDD9">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:date id="F7B8DF499E29014D1693EFEF26FE2D9C">2022</mods:date>
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<paragraph id="8BCC643DFFB9FFF6FD32BC155B06A04F" blockId="28.[680,1088,295,350]" box="[680,1088,295,319]" pageId="28" pageNumber="359">
<heading id="D084D351FFB9FFF6FD32BC155B06A04F" bold="true" box="[680,1088,295,319]" fontSize="10" level="2" pageId="28" pageNumber="359" reason="4">
<emphasis id="B907B82FFFB9FFF6FD32BC155B06A04F" bold="true" box="[680,1088,295,319]" pageId="28" pageNumber="359">
<taxonomicName id="4C731FBEFFB9FFF6FD32BC155CA3A04F" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[680,997,295,319]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="359" phylum="Arthropoda" rank="species" species="myitkyinaensis" status="sp. nov.">
<emphasis id="B907B82FFFB9FFF6FD32BC155CA3A04F" bold="true" box="[680,997,295,319]" italics="true" pageId="28" pageNumber="359">Sanaungulus myitkyinaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB9FFF6FC71BC155B06A04F" box="[1003,1088,295,319]" pageId="28" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB9FFF6FD32BC755C43A02E" box="[680,773,327,350]" pageId="28" pageNumber="359" type="description">
<paragraph id="8BCC643DFFB9FFF6FD32BC755C43A02E" blockId="28.[680,1088,295,350]" box="[680,773,327,350]" pageId="28" pageNumber="359">
(
<figureCitation id="134878B8FFB9FFF6FD2ABC755DBBA02E" box="[688,765,327,350]" captionStart="Fig" captionStartId="29.[113,152,1636,1659]" captionTargetBox="[113,1134,380,1605]" captionTargetId="figure-125@29.[113,1134,380,1605]" captionTargetPageId="29" captionText="Fig. 16. Sanaungulus myitkyinaensis sp. nov. in Burmese amber.A: Holotype, dorso­lateral view; B: Holotype, ventro­lateral view." pageId="28" pageNumber="359">Fig. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB9FFF6FD32BC4F5C81A3CF" pageId="28" pageNumber="359" type="materials_examined">
<paragraph id="8BCC643DFFB9FFF6FD32BC4F5CC2A363" blockId="28.[680,1172,381,531]" pageId="28" pageNumber="359">
<materialsCitation id="3B1B6E60FFB9FFF6FD32BC4F5CC2A363" collectionCode="BUB" county="Probable" location="Catalog" municipality="Burmese" pageId="28" pageNumber="359" specimenCode="BUB4462" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFB9FFF6FD32BC4F5C51A0E5" bold="true" box="[680,791,381,405]" pageId="28" pageNumber="359">
<typeStatus id="54C8DA9FFFB9FFF6FD32BC4F5C54A0E5" box="[680,786,381,405]" pageId="28" pageNumber="359" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFB9FFF6FC84BC4F5C3BA0E4" box="[798,893,381,404]" pageId="28" pageNumber="359">Probable</collectingCounty>
male, adult specimen in a
<collectingMunicipality id="6BA8FE47FFB9FFF6FD32BCAF5C42A0C4" box="[680,772,413,436]" pageId="28" pageNumber="359">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFB9FFF6FC8ABCAF5C18A0C4" box="[784,862,413,436]" country="Myanmar" name="Kachin" pageId="28" pageNumber="359">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFB9FFF6FC6ABCAF5B06A0C4" LSID="urn:lsid:plazi:treatment:03DAD52BFFB9FFF4FD32BC155E5FA546:8EAC32E6FFB9FFF6FC6ABCAF5B06A0C4" box="[1008,1088,413,436]" county="Probable" municipality="Burmese" name="Catalog" pageId="28" pageNumber="359" stateProvince="Kachin">Catalog</location>
number SNHM­
<quantity id="4C8BC9D8FFB9FFF6FD62BC8F5C01A0A4" box="[760,839,445,468]" metricMagnitude="2" metricUnit="m" metricValue="1.52654" pageId="28" pageNumber="359" unit="in" value="6010.0">6010 in</quantity>
the
<location id="8EAC32E6FFB9FFF6FCF5BC8F5CE7A084" LSID="urn:lsid:plazi:treatment:03DAD52BFFB9FFF4FD32BC155E5FA546:8EAC32E6FFB9FFF6FCF5BC8F5CE7A084" county="Probable" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="28" pageNumber="359" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FFB9FFF6FC56BCEF5B60A084" LSID="urn:lsid:plazi:treatment:03DAD52BFFB9FFF4FD32BC155E5FA546:8EAC32E6FFB9FFF6FC56BCEF5B60A084" box="[972,1062,477,500]" county="Probable" municipality="Burmese" name="Muller's" pageId="28" pageNumber="359" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFB9FFF6FC92BCCE5C3FA363" box="[776,889,508,531]" collectionCode="BUB" pageId="28" pageNumber="359">BUB4462</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFB9FFF6FD32BF005CBDA319" blockId="28.[680,1172,562,617]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FD32BF005C07A33A" bold="true" box="[680,833,562,586]" pageId="28" pageNumber="359">Type locality.</emphasis>
<collectingCountry id="F36424ADFFB9FFF6FCD0BF015CF0A33A" box="[842,950,563,586]" name="Myanmar" pageId="28" pageNumber="359">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFB9FFF6FC5BBF015B0DA33A" box="[961,1099,563,586]" country="Myanmar" name="Kachin" pageId="28" pageNumber="359">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFB9FFF6FD32BFBA5C81A3CF" blockId="28.[680,1172,648,703]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FD32BFBA5C7AA3D0" bold="true" box="[680,828,648,672]" pageId="28" pageNumber="359">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB9FFF6FD32BFEC5CE1A2E4" pageId="28" pageNumber="359" type="diagnosis">
<paragraph id="8BCC643DFFB9FFF6FD32BFEC5CE1A2E4" blockId="28.[680,1172,734,916]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FD32BFEC5CE2A386" bold="true" box="[680,932,734,758]" pageId="28" pageNumber="359">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFFB9FFF6FC36BFED5DBEA266" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="359" phylum="Arthropoda" rank="species" species="myitkyinaensis" status="sp. nov.">
<emphasis id="B907B82FFFB9FFF6FC36BFED5DBEA266" italics="true" pageId="28" pageNumber="359">Sanaungulus myitkyinaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB9FFF6FD65BFCD5C17A266" box="[767,849,767,790]" pageId="28" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by pectinated antennae with an antennal process in antennomeres VIX. The new species is easily distinguishable from other
<taxonomicName id="4C731FBEFFB9FFF6FC0DBE6C5B67A205" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[919,1057,862,885]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFB9FFF6FC0DBE6C5B67A205" box="[919,1057,862,885]" italics="true" pageId="28" pageNumber="359">Sanaungulus</emphasis>
</taxonomicName>
by its extremely elongated head.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB9FFF4FD32BE865E05A382" lastPageId="30" lastPageNumber="361" pageId="28" pageNumber="359" type="description">
<paragraph id="8BCC643DFFB9FFF6FD32BE865B2EA57A" blockId="28.[680,1172,948,1034]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FD32BE865C74A2BC" bold="true" box="[680,818,948,972]" pageId="28" pageNumber="359">Description.</emphasis>
Adult, winged, rather robust. Probable male, defined on the basis of long antennae. Body length
<quantity id="4C8BC9D8FFB9FFF6FCF9BEC15CF1A57A" box="[867,951,1011,1034]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="28" pageNumber="359" unit="mm" value="3.3">3.3 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFB9FFF4FD32B9185E05A382" blockId="28.[680,1172,1066,1692]" lastBlockId="30.[151,643,155,754]" lastPageId="30" lastPageNumber="361" pageId="28" pageNumber="359">Head extremely long, curved, surface smooth equipped with only some long hairs. Eyes relatively small, very slightly sub­elliptical, inserted in the upper and lateral part of the head. Mandibles elongated, falciform, slender with a pointed apex. Maxillary palps 4­ segmented with the last palpomere securiform. Labial palps 3­ segmented, with the last palpomere securiform.Antennae 11­ segmented, pectinate, very long, almost reaching the last abdominal segment, antennal insertion in the eyes proximity; scape club­shaped; antennomere II short, slightly swollen apically; antennomere III elongated, filiform, about 1.4 times longer than second; antennomeres IVV extremely long, without antennal process, enlarged (in a shape of a draft) apically, antennomere V the longest; antennomere VI elongated, as long as antennomere IV, equipped near the base with a long antennal process which is very long and rounded apically; antennomere VII shorter than previous one, equipped with a long antennal process near the base; antennomeres VIIIX sub­equal, shorter and sturdier than previous one, each equipped near the base with an antennal process shorter and slightly sturdier than others; antennomere XI rather robust, with slightly squadrate and pointed apex; all the antennomeres pubescent. Pronotum elongated, slightly narrower than head, anterior margin straight and strongly bordered, posterior margin slightly undulate (expecially in the middle) and strongly bordered, sides straight and bordered, corners rounded, pronotum disc almost flat with short pubescence. Elytra short which reveals various abdominal segments, wider than pronotum, gibbous at humeri, parallel­sided, rounded apexes, surface pubescent with confused and impressed punctuation. Metathoracic wings longer than elytra, reaching the last abdominal segments. Sternites transverse, last sternite very wide. Legs thin, long, pubescent; coxae massive; trochanters elongated; femora almost straight, cylindrical, sturdier than tibiae; tibiae, thin, cylindrical, long, without a spur at apex, protibiae shorter than profemora, meso­ and metatibiae longer than meso­ and metafemora. Tarsal formula 5­ 5­ 5; first tarsomere very long, about 2.0 times longer than second; third tarsomere robust; fourth strongly bilobed; fifth tarsomere elongated, thin, curved; claws simple, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFB8FFF7FFEBBB565E89A7EB" pageId="29" pageNumber="360" startId="29.[113,152,1636,1659]" targetBox="[113,1134,380,1605]" targetPageId="29" targetType="figure">
<paragraph id="8BCC643DFFB8FFF7FFEBBB565E89A7EB" blockId="29.[113,1134,1636,1691]" pageId="29" pageNumber="360">
Fig. 16.
<taxonomicName id="4C731FBEFFB8FFF7FF5DBB565EB2A70B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[199,500,1636,1659]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="360" phylum="Arthropoda" rank="species" species="myitkyinaensis" status="sp. nov.">
<emphasis id="B907B82FFFB8FFF7FF5DBB565EB2A70B" box="[199,500,1636,1659]" italics="true" pageId="29" pageNumber="360">Sanaungulus myitkyinaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFB8FFF7FE63BB565D0CA70B" box="[505,586,1636,1659]" pageId="29" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorso­lateral view; B: Holotype, ventro­lateral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFBBFFF4FF0DBE235D59A2EF" pageId="30" pageNumber="361" type="etymology">
<paragraph id="8BCC643DFFBBFFF4FF0DBE235E8DA218" blockId="30.[151,643,785,873]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DBE235E5EA259" bold="true" box="[151,280,785,809]" pageId="30" pageNumber="361">Etymology.</emphasis>
Derived from the toponym Myitkyina (district where the amber mines are located) + the Latin suffix ­Á
<emphasis id="B907B82FFFBBFFF4FE00BE605E82A219" box="[410,452,850,873]" italics="true" pageId="30" pageNumber="361">nsis</emphasis>
.
</paragraph>
<paragraph id="8BCC643DFFBBFFF4FF0DBEBA5D59A2EF" blockId="30.[151,543,904,928]" box="[151,543,904,928]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DBEBA5E7CA2D0" bold="true" box="[151,314,904,928]" pageId="30" pageNumber="361">Syninclusions.</emphasis>
Air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFF4FF0DBE8C5E5FA546" pageId="30" pageNumber="361" type="discussion">
<paragraph id="8BCC643DFFBBFFF4FF0DBE8C5E5FA546" blockId="30.[151,643,958,1078]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DBE8C5E45A2A6" bold="true" box="[151,259,958,982]" pageId="30" pageNumber="361">Remarks.</emphasis>
Piece of clear, golden amber that measures 13x10x
<quantity id="4C8BC9D8FFBBFFF4FEBCBEED5E20A286" box="[294,358,991,1014]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="361" unit="mm" value="3.0">3 mm</quantity>
. The inclusion is complete but the head is bent and the left proleg is not well visible.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,150 @@
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<mods:title id="F48D32108748990F2DE22213B168398A">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="B1601E115C03A017850090D5BE7D21AE">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="F824C4DB51F438599CB905AA27F0E53C">Müller, Patrick</mods:namePart>
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<subSubSection id="C36937B6FFBBFFF4FF0DB95A5E8FA5F0" box="[151,457,1128,1152]" pageId="30" pageNumber="361" type="nomenclature">
<paragraph id="8BCC643DFFBBFFF4FF0DB95A5E8FA5F0" blockId="30.[151,457,1128,1183]" box="[151,457,1128,1152]" pageId="30" pageNumber="361">
<heading id="D084D351FFBBFFF4FF0DB95A5E8FA5F0" bold="true" box="[151,457,1128,1152]" fontSize="10" level="2" pageId="30" pageNumber="361" reason="4">
<emphasis id="B907B82FFFBBFFF4FF0DB95A5E8FA5F0" bold="true" box="[151,457,1128,1152]" pageId="30" pageNumber="361">
<taxonomicName id="4C731FBEFFBBFFF4FF0DB95A5E2BA5F0" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[151,365,1128,1152]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="361" phylum="Arthropoda" rank="species" species="nalae" status="sp. nov.">
<emphasis id="B907B82FFFBBFFF4FF0DB95A5E2BA5F0" bold="true" box="[151,365,1128,1152]" italics="true" pageId="30" pageNumber="361">Sanaungulus nalae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBBFFF4FEEEB95A5E8FA5F0" box="[372,457,1128,1152]" pageId="30" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFF4FF0DB9BA5FB2A5EF" box="[151,244,1160,1183]" pageId="30" pageNumber="361" type="description">
<paragraph id="8BCC643DFFBBFFF4FF0DB9BA5FB2A5EF" blockId="30.[151,457,1128,1183]" box="[151,244,1160,1183]" pageId="30" pageNumber="361">
(
<figureCitation id="134878B8FFBBFFF4FF05B9BA5FAAA5EF" box="[159,236,1160,1183]" captionStart="Fig" captionStartId="31.[113,152,1636,1659]" captionTargetBox="[223,1026,151,1606]" captionTargetId="figure-37@31.[223,1026,637,1120]" captionTargetPageId="31" captionText="Fig. 17. Sanaungulus nalae sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of head and pronotum." pageId="30" pageNumber="361">Fig. 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFF4FF0DB98D5EF0A773" pageId="30" pageNumber="361" type="materials_examined">
<paragraph id="8BCC643DFFBBFFF4FF0DB98D5E53A426" blockId="30.[151,644,1215,1366]" pageId="30" pageNumber="361">
<materialsCitation id="3B1B6E60FFBBFFF4FF0DB98D5E53A426" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="30" pageNumber="361" specimenCode="BUB4033" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFBBFFF4FF0DB98D5E40A5A7" bold="true" box="[151,262,1215,1239]" pageId="30" pageNumber="361">
<typeStatus id="54C8DA9FFFBBFFF4FF0DB98D5E47A5A7" box="[151,257,1215,1239]" pageId="30" pageNumber="361" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFBBFFF4FE96B98D5E03A5A6" box="[268,325,1215,1238]" pageId="30" pageNumber="361">Male</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFBBFFF4FDBEB98D5DC5A5A6" box="[548,643,1215,1238]" pageId="30" pageNumber="361">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFBBFFF4FF05B9ED5FA8A586" box="[159,238,1247,1270]" country="Myanmar" name="Kachin" pageId="30" pageNumber="361">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFBBFFF4FE19B9ED5E93A586" LSID="urn:lsid:plazi:treatment:03DAD52BFFBBFFCAFF0DB95A5D72A26B:8EAC32E6FFBBFFF4FE19B9ED5E93A586" box="[387,469,1247,1270]" county="Male" municipality="Burmese" name="Catalog" pageId="30" pageNumber="361" stateProvince="Kachin">Catalog</location>
number
<quantity id="4C8BC9D8FFBBFFF4FDB4B9ED5FA7A466" metricMagnitude="2" metricUnit="m" metricValue="1.526794" pageId="30" pageNumber="361" unit="in" value="6011.0">SNHM­6011 in</quantity>
the
<location id="8EAC32E6FFBBFFF4FE9DB9CD5E69A446" LSID="urn:lsid:plazi:treatment:03DAD52BFFBBFFCAFF0DB95A5D72A26B:8EAC32E6FFBBFFF4FE9DB9CD5E69A446" county="Male" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="30" pageNumber="361" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FFBBFFF4FEC6B82D5EF1A446" LSID="urn:lsid:plazi:treatment:03DAD52BFFBBFFCAFF0DB95A5D72A26B:8EAC32E6FFBBFFF4FEC6B82D5EF1A446" box="[348,439,1311,1334]" county="Male" municipality="Burmese" name="Muller's" pageId="30" pageNumber="361" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFBBFFF4FF0DB80D5E4FA426" box="[151,265,1343,1366]" collectionCode="BUB" pageId="30" pageNumber="361">BUB4033</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFBBFFF4FF0DB8475EACA4DD" blockId="30.[151,643,1397,1453]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DB8475E76A4FD" bold="true" box="[151,304,1397,1421]" pageId="30" pageNumber="361">Type locality.</emphasis>
<collectingCountry id="F36424ADFFBBFFF4FEA2B8445EE2A4FD" box="[312,420,1398,1421]" name="Myanmar" pageId="30" pageNumber="361">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFBBFFF4FE2AB8445D7CA4FD" box="[432,570,1398,1421]" country="Myanmar" name="Kachin" pageId="30" pageNumber="361">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFBBFFF4FF0DB8FE5EF0A773" blockId="30.[151,643,1484,1539]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DB8FE5E6CA494" bold="true" box="[151,298,1484,1508]" pageId="30" pageNumber="361">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFF4FF0DBB115B4BA182" pageId="30" pageNumber="361" type="diagnosis">
<paragraph id="8BCC643DFFBBFFF4FF0DBB115B4BA182" blockId="30.[151,644,1571,1690]" lastBlockId="30.[680,1172,155,242]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DBB115ED6A74B" bold="true" box="[151,400,1571,1595]" pageId="30" pageNumber="361">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFFBBFFF4FE0FBB115D19A74A" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[405,607,1571,1594]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="361" phylum="Arthropoda" rank="species" species="nalae" status="sp. nov.">
<emphasis id="B907B82FFFBBFFF4FE0FBB115D19A74A" box="[405,607,1571,1594]" italics="true" pageId="30" pageNumber="361">Sanaungulus nalae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBBFFF4FDFEBB115F83A72A" pageId="30" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
differs from other
<taxonomicName id="4C731FBEFFBBFFF4FE08BB715D5AA72A" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[402,540,1603,1626]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="361" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFBBFFF4FE08BB715D5AA72A" box="[402,540,1603,1626]" italics="true" pageId="30" pageNumber="361">Sanaungulus</emphasis>
</taxonomicName>
with filiform antennae for a combination of characters: Head partially covered by pronotum, smooth elytra, and a pronotal shape with the sides wider near the posterior margin and narrower from the anterior margin to the middle.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFCAFD32BC235F88A0A4" lastPageId="32" lastPageNumber="363" pageId="30" pageNumber="361" type="description">
<paragraph id="8BCC643DFFBBFFF4FD32BC235B73A0F8" blockId="30.[680,1172,273,392]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FD32BC235C68A059" bold="true" box="[680,814,273,297]" pageId="30" pageNumber="361">Description.</emphasis>
Adult, winged, slender. Male, defined on the shape of the last sternite which is triangular, and narrower than the last tergite. Body length
<quantity id="4C8BC9D8FFBBFFF4FCABBC435CC3A0F8" box="[817,901,369,392]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="30" pageNumber="361" unit="mm" value="5.2">5.2 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFBBFFCAFD32BC9A5F88A0A4" blockId="30.[680,1173,424,1692]" lastBlockId="32.[151,643,155,468]" lastPageId="32" lastPageNumber="363" pageId="30" pageNumber="361">Head transverse, wide, slightly convex in the middle, constricted behind the eyes, smooth with short pubescence. Eyes prominent, very large, roundish and slightly subelliptic, inserted in the lateral part of the head, inter­ocular dorsal distance about 2.1 times greater than the eye diameter. Mandibles falciform. Maxillary palps 4­ segmented with the last palpomere very elongated, securiform. Labial palps 3­ segmented. Antennae 11­ segmented, filiform, short, slightly surpassing the half of elytra and almost reaching the half of abdomen, antennal insertion in the eyes proximity; scape elongated, robust; antennomere II very short, subovate; antennomere III thinner and about 1.8 times longer than second; antennomeres IVV subequal, slightly longer than antennomere III; antennomere VI as long as the previous ones, slightly thinner; antennomeres VIIX filiform, subequal; antennomere XI filiform, slightly sturdier than previous ones, rounded at apex; all the antennomeres with short pubescence. Pronotum transverse, wider than head, narrow anteriorly, anterior margin rounded and strongly bordered, posterior margin almost straight, sides enlarged near the posterior margin and narrower from the anterior margin to the middle, posterior corners rounded, pronotum disc smooth and pubescent. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, at humeri wider than pronotum, humeral zone slightly gibbous, posteriorly parallel­sided, rounded apexes, surface smooth and sligthly pubescent equipped with short setae. Metathoracic wings infuscate, very longer than elytra, not reaching the apex of abdomen. Sternum elongated with posterior margin rounded, sternites narrow and transverse, last tergite wide with posterior margin rounded, last sternite triangular and narrower than last tergite. Legs thin, relatively short, pubescent; coxae massive, elongated, curved; trochanters elongated, curved, roundish at apex; femora more robust than tibiae, nearly straight; tibiae cylindrical, thin, without spurs near the apex, longer than femora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 2.8 times longer than second; third tarsomere short, triangular; fourth strongly bilobed; fifth tarsomere elongated, thin, flat and curved; claws simple, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFBAFFF5FFEBBB565D80A7EB" pageId="31" pageNumber="362" startId="31.[113,152,1636,1659]" targetBox="[223,1026,151,1606]" targetPageId="31" targetType="figure">
<paragraph id="8BCC643DFFBAFFF5FFEBBB565D80A7EB" blockId="31.[113,1134,1636,1691]" pageId="31" pageNumber="362">
Fig. 17.
<taxonomicName id="4C731FBEFFBAFFF5FF51BB565ED1A70B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[203,407,1636,1659]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="362" phylum="Arthropoda" rank="species" species="nalae" status="sp. nov.">
<emphasis id="B907B82FFFBAFFF5FF51BB565ED1A70B" box="[203,407,1636,1659]" italics="true" pageId="31" pageNumber="362">Sanaungulus nalae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBAFFF5FE04BB565EB7A70B" box="[414,497,1636,1659]" pageId="31" pageNumber="362" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of head and pronotum.
</paragraph>
</caption>
<subSubSection id="C36937B6FF85FFCAFF0DBCC65FA9A3F3" pageId="32" pageNumber="363" type="etymology">
<paragraph id="8BCC643DFF85FFCAFF0DBCC65D78A35C" blockId="32.[151,643,500,556]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DBCC65E5EA37C" bold="true" box="[151,280,500,524]" pageId="32" pageNumber="363">Etymology.</emphasis>
Named in honor of Nala Müller, daughter of the second author (Patrick).
</paragraph>
<paragraph id="8BCC643DFF85FFCAFF0DBF795FA9A3F3" blockId="32.[151,643,587,643]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DBF795E7CA313" bold="true" box="[151,314,587,611]" pageId="32" pageNumber="363">Syninclusions.</emphasis>
<taxonomicName id="4C731FBEFF85FFCAFED9BF7E5ED5A313" box="[323,403,588,611]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="363" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
(partial), air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF85FFCAFF0DBF905D72A26B" pageId="32" pageNumber="363" type="discussion">
<paragraph id="8BCC643DFF85FFCAFF0DBF905D72A26B" blockId="32.[151,643,674,795]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DBF905E40A3CA" bold="true" box="[151,262,674,698]" pageId="32" pageNumber="363">Remarks.</emphasis>
Piece of clear, honey colored amber that measures 10x10x
<quantity id="4C8BC9D8FF85FFCAFEE6BFF15E94A3AA" box="[380,466,707,730]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="32" pageNumber="363" unit="mm" value="4.5">4.5 mm</quantity>
. The inclusion is complete. The specimen is ventrally covered by a white and large cotton­like emulsion.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,147 @@
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<mods:title id="C395146346FE6B37CF6FFD743736CDA2">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="8C4C5BE3D4EF8C8FE04E29184C48BC0C">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="B0CA49ADD5C5258179A4DC1099267BC0">Müller, Patrick</mods:namePart>
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<mods:date id="4A04CA36EEA56856D147AC6972B8CC7A">2022</mods:date>
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<subSubSection id="C36937B6FFBCFFF3FFEBBE8A5EEFA2BF" box="[113,425,951,976]" pageId="25" pageNumber="356" type="nomenclature">
<paragraph id="8BCC643DFFBCFFF3FFEBBE8A5EEFA2BF" blockId="25.[113,425,951,1007]" box="[113,425,951,976]" pageId="25" pageNumber="356">
<heading id="D084D351FFBCFFF3FFEBBE8A5EEFA2BF" bold="true" box="[113,425,951,976]" fontSize="10" level="2" pageId="25" pageNumber="356" reason="4">
<emphasis id="B907B82FFFBCFFF3FFEBBE8A5EEFA2BF" bold="true" box="[113,425,951,976]" pageId="25" pageNumber="356">
<taxonomicName id="4C731FBEFFBCFFF3FFEBBE8A5E08A2A0" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[113,334,952,976]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="356" phylum="Arthropoda" rank="species" species="leniae" status="sp. nov.">
<emphasis id="B907B82FFFBCFFF3FFEBBE8A5E08A2A0" bold="true" box="[113,334,952,976]" italics="true" pageId="25" pageNumber="356">Sanaungulus leniae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBCFFF3FECEBE855EEFA2BF" box="[340,425,951,975]" pageId="25" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBCFFF3FFEBBEEA5F88A29F" box="[113,206,984,1007]" pageId="25" pageNumber="356" type="description">
<paragraph id="8BCC643DFFBCFFF3FFEBBEEA5F88A29F" blockId="25.[113,425,951,1007]" box="[113,206,984,1007]" pageId="25" pageNumber="356">
(
<figureCitation id="134878B8FFBCFFF3FFE3BEEA5F80A29F" box="[121,198,984,1007]" captionStart="Fig" captionStartId="26.[151,190,1638,1661]" captionTargetBox="[262,1066,151,1605]" captionTargetId="figure-49@26.[262,1066,636,1118]" captionTargetPageId="26" captionText="Fig. 14. Sanaungulus leniae sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of right metathoracic leg and right antenna." pageId="25" pageNumber="356">Fig. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBCFFF3FFEBB93D5ED6A428" pageId="25" pageNumber="356" type="materials_examined">
<paragraph id="8BCC643DFFBCFFF3FFEBB93D5EFBA5D9" blockId="25.[113,605,1039,1193]" pageId="25" pageNumber="356">
<materialsCitation id="3B1B6E60FFBCFFF3FFEBB93D5EFBA5D9" collectionCode="BUB" county="Sex" location="Catalog" municipality="Burmese" pageId="25" pageNumber="356" specimenCode="BUB4315" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFBCFFF3FFEBB93D5FA6A557" bold="true" box="[113,224,1039,1063]" pageId="25" pageNumber="356">
<typeStatus id="54C8DA9FFFBCFFF3FFEBB93D5F9DA557" box="[113,219,1039,1063]" pageId="25" pageNumber="356" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFBCFFF3FF7DB93D5E49A556" box="[231,271,1039,1062]" pageId="25" pageNumber="356">Sex</collectingCounty>
undefined, adult specimen in a
<collectingMunicipality id="6BA8FE47FFBCFFF3FFEBB9025F8BA537" box="[113,205,1072,1095]" pageId="25" pageNumber="356">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFBCFFF3FF43B9025E61A537" box="[217,295,1072,1095]" country="Myanmar" name="Kachin" pageId="25" pageNumber="356">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFBCFFF3FE23B9025D4FA537" LSID="urn:lsid:plazi:treatment:03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1:8EAC32E6FFBCFFF3FE23B9025D4FA537" box="[441,521,1072,1095]" county="Sex" municipality="Burmese" name="Catalog" pageId="25" pageNumber="356" stateProvince="Kachin">Catalog</location>
number SNHM­
<quantity id="4C8BC9D8FFBCFFF3FF50B9625E67A517" box="[202,289,1104,1127]" metricMagnitude="2" metricUnit="m" metricValue="1.5260319999999998" pageId="25" pageNumber="356" unit="in" value="6008.0">6008 in</quantity>
the
<location id="8EAC32E6FFBCFFF3FEC0B9625E8FA5F8" LSID="urn:lsid:plazi:treatment:03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1:8EAC32E6FFBCFFF3FEC0B9625E8FA5F8" county="Sex" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="25" pageNumber="356" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FFBCFFF3FD9AB9435D1AA5F8" LSID="urn:lsid:plazi:treatment:03DAD52BFFBCFFF1FFEBBE8A5D1BA3D1:8EAC32E6FFBCFFF3FD9AB9435D1AA5F8" box="[512,604,1137,1160]" county="Sex" municipality="Burmese" name="Muller's" pageId="25" pageNumber="356" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFBCFFF3FEDAB9A05EF7A5D9" box="[320,433,1170,1193]" collectionCode="BUB" pageId="25" pageNumber="356">BUB4315</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFBCFFF3FFEBB9FA5E82A470" blockId="25.[113,605,1224,1280]" pageId="25" pageNumber="356">
<emphasis id="B907B82FFFBCFFF3FFEBB9FA5E4CA590" bold="true" box="[113,266,1224,1248]" pageId="25" pageNumber="356">Type locality.</emphasis>
<collectingCountry id="F36424ADFFBCFFF3FE89B9FB5E39A590" box="[275,383,1225,1248]" name="Myanmar" pageId="25" pageNumber="356">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFBCFFF3FE10B9FB5D52A590" box="[394,532,1225,1248]" country="Myanmar" name="Kachin" pageId="25" pageNumber="356">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFBCFFF3FFEBB8125ED6A428" blockId="25.[113,605,1312,1368]" pageId="25" pageNumber="356">
<emphasis id="B907B82FFFBCFFF3FFEBB8125E43A448" bold="true" box="[113,261,1312,1336]" pageId="25" pageNumber="356">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBCFFF3FFEBB84A5FF2A7E4" pageId="25" pageNumber="356" type="diagnosis">
<paragraph id="8BCC643DFFBCFFF3FFEBB84A5FF2A7E4" blockId="25.[113,605,1400,1684]" pageId="25" pageNumber="356">
<emphasis id="B907B82FFFBCFFF3FFEBB84A5E2CA4E0" bold="true" box="[113,362,1400,1424]" pageId="25" pageNumber="356">Differential diagnosis.</emphasis>
The non­dehiscent and wide elytra make this new species easily distinguishable from all other
<taxonomicName id="4C731FBEFFBCFFF3FE0CB8885D66A4A1" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[406,544,1466,1489]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="356" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFBCFFF3FE0CB8885D66A4A1" box="[406,544,1466,1489]" italics="true" pageId="25" pageNumber="356">Sanaungulus</emphasis>
</taxonomicName>
. This character could easily make one think we this is a new genus; unfortunately, the specimen has a crumpled abdomen and we cannot evaluate the length of the elytra. Only the discovery of one or more similar specimens will clarify the taxon.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBCFFF1FD19BDA95EA8A007" lastPageId="27" lastPageNumber="358" pageId="25" pageNumber="356" type="description">
<paragraph id="8BCC643DFFBCFFF3FD19BDA95C67A183" blockId="25.[643,1134,155,243]" pageId="25" pageNumber="356">
<emphasis id="B907B82FFFBCFFF3FD19BDA95C4BA1C3" bold="true" box="[643,781,155,179]" pageId="25" pageNumber="356">Description.</emphasis>
Adult, winged, rather robust. Sex undefined. Body length
<quantity id="4C8BC9D8FFBCFFF3FC1CBD895C9AA1A2" box="[902,988,187,210]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="356" unit="mm" value="2.1">2.1 mm</quantity>
. Entirely reddish­brownish.
</paragraph>
<paragraph id="8BCC643DFFBCFFF1FD19BC205EA8A007" blockId="25.[643,1137,274,1685]" lastBlockId="27.[113,605,155,375]" lastPageId="27" lastPageNumber="358" pageId="25" pageNumber="356">Head transverse, wide, slightly convex in the apex, constricted behind the eyes, covered by pronotum, smooth with short pubescence. Eyes prominent, large, convex, roundish­subelliptic, inserted in the lateral­upper part of the head, inter­ocular dorsal distance about 3.9 times greater than the eye diameter. Mandibles falciform, slender. Maxillary palps 4­ segmented with the last palpomere securiform. Labial palps 3­ segmented. Antennae 11­ segmented, filiform, short, not reaching the half of elytra, antennal insertion in frons, far from eyes; scape elongated, robust, curved and distorted; antennomere II short, robust, enlarged and globular from the middle to the apex; antennomere III thinner and longer than second; antennomere IV slightly shorter than antennomere III; antennomeres V VIII filiform and sub­equal in length; antennomeres IXXI globular­elongated, sturdier than previous ones, with antennomere IX clavate, antennomere X shorter than previous one and antennomere XI rounded at apex; all the antennomeres with very short pubescence. Pronotum transverse, as long as head (eyes are slightly extruded), anterior margin very slightly curved and not bordered, posterior margin almost straight and not bordered, sides strongly curved, corners rounded, pronotum disc smooth and flat andpubescent.Scutellumsmall, triangular­shaped, with pointed apex. Elytra elongate, wider than pronotum and head, subparallel­sided with apex enlarged, strongly rounded apexes, surface smooth without punctaction, and sligthly pubescent equipped with short and not erected setae. Metathoracic wings transparent, almost completely covered by elytra, as long as elytra or very slightly longer. Sternum very elongated and subquadrate with posterior margin almost straight, sternites transverse, last tergite elongated, narrower than previous ones and rounded at apex. Legs thin, rather long, pubescent; coxae massive, elongated, curved; trochanters elongated, curved, roundish at apex; femora very sturdier than tibiae, nearly straight; tibiae cylindrical, thin, without spurs near the apex, metatibiae longer than metafemora. Tarsal formula 5­ 5­ 5; first tarsomere very elongated and thin; second tarsomere about 1.3 times shorter than first, third tarsomere shorter than second; fourth slightly bilobed; fifth tarsomere elongated, thin, flat and curved; claws simple, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFBFFFF0FF0DBB545C83A7ED" pageId="26" pageNumber="357" startId="26.[151,190,1638,1661]" targetBox="[262,1066,151,1605]" targetPageId="26" targetType="figure">
<paragraph id="8BCC643DFFBFFFF0FF0DBB545C83A7ED" blockId="26.[151,1172,1638,1693]" pageId="26" pageNumber="357">
Fig. 14.
<taxonomicName id="4C731FBEFFBFFFF0FF6ABB545E84A70D" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[240,450,1638,1661]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="357" phylum="Arthropoda" rank="species" species="leniae" status="sp. nov.">
<emphasis id="B907B82FFFBFFFF0FF6ABB545E84A70D" box="[240,450,1638,1661]" italics="true" pageId="26" pageNumber="357">Sanaungulus leniae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBFFFF0FE52BB545D5DA70D" box="[456,539,1638,1661]" pageId="26" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of right metathoracic leg and right antenna.
</paragraph>
</caption>
<subSubSection id="C36937B6FFBEFFF1FFEBBCA55E0AA358" pageId="27" pageNumber="358" type="etymology">
<paragraph id="8BCC643DFFBEFFF1FFEBBCA55D5EA0BF" blockId="27.[113,605,407,463]" pageId="27" pageNumber="358">
<emphasis id="B907B82FFFBEFFF1FFEBBCA55FB4A0DF" bold="true" box="[113,242,407,431]" pageId="27" pageNumber="358">Etymology.</emphasis>
Named in honor of Leni Müller, daughter of the second author (Patrick).
</paragraph>
<paragraph id="8BCC643DFFBEFFF1FFEBBCDD5E0AA358" blockId="27.[113,605,495,552]" pageId="27" pageNumber="358">
<emphasis id="B907B82FFFBEFFF1FFEBBCDD5E52A377" bold="true" box="[113,276,495,519]" pageId="27" pageNumber="358">Syninclusions.</emphasis>
Two Psocoptera, one
<taxonomicName id="4C731FBEFFBEFFF1FD9CBCC25D1FA377" box="[518,601,496,519]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
, air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBEFFF1FFEBBF7A5D1BA3D1" pageId="27" pageNumber="358" type="discussion">
<paragraph id="8BCC643DFFBEFFF1FFEBBF7A5D1BA3D1" blockId="27.[113,605,584,673]" pageId="27" pageNumber="358">
<emphasis id="B907B82FFFBEFFF1FFEBBF7A5F98A310" bold="true" box="[113,222,584,608]" pageId="27" pageNumber="358">Remarks.</emphasis>
Piece of clear, golden amber in Cabochon cut 27x14x
<quantity id="4C8BC9D8FFBEFFF1FEBCBF5B5E21A3F0" box="[294,359,617,640]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="358" unit="mm" value="5.0">5 mm</quantity>
. The inclusion is complete, but its abdomen is folded and crumpled.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,164 @@
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<mods:title id="50680D27F4356D211A4C9FA987BFB0AE">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:date id="1FE4BBA7DC6AF0504E6047C2DCF1E1AC">2022</mods:date>
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<treatment id="03DAD52BFFBEFFF6FFEBBFEA5B10A181" LSID="urn:lsid:plazi:treatment:03DAD52BFFBEFFF6FFEBBFEA5B10A181" httpUri="http://treatment.plazi.org/id/03DAD52BFFBEFFF6FFEBBFEA5B10A181" lastPageId="28" lastPageNumber="359" pageId="27" pageNumber="358">
<subSubSection id="C36937B6FFBEFFF1FFEBBFEA5EBAA39F" box="[113,508,727,752]" pageId="27" pageNumber="358" type="nomenclature">
<paragraph id="8BCC643DFFBEFFF1FFEBBFEA5EBAA39F" blockId="27.[113,508,727,784]" box="[113,508,727,752]" pageId="27" pageNumber="358">
<heading id="D084D351FFBEFFF1FFEBBFEA5EBAA39F" bold="true" box="[113,508,727,752]" fontSize="10" level="2" pageId="27" pageNumber="358" reason="4">
<emphasis id="B907B82FFFBEFFF1FFEBBFEA5EBAA39F" bold="true" box="[113,508,727,752]" pageId="27" pageNumber="358">
<taxonomicName id="4C731FBEFFBEFFF1FFEBBFEA5EE7A380" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[113,417,728,752]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="myanmaricus" status="sp. nov.">
<emphasis id="B907B82FFFBEFFF1FFEBBFEA5EE7A380" bold="true" box="[113,417,728,752]" italics="true" pageId="27" pageNumber="358">Sanaungulus myanmaricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBEFFF1FE3DBFE55EBAA39F" box="[423,508,727,751]" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBEFFF1FFEBBFCB5F88A260" box="[113,206,761,784]" pageId="27" pageNumber="358" type="description">
<paragraph id="8BCC643DFFBEFFF1FFEBBFCB5F88A260" blockId="27.[113,508,727,784]" box="[113,206,761,784]" pageId="27" pageNumber="358">
(
<figureCitation id="134878B8FFBEFFF1FFE3BFCB5F80A260" box="[121,198,761,784]" captionStart="Fig" captionStartId="27.[113,152,1665,1688]" captionTargetBox="[113,1134,1022,1634]" captionTargetId="figure-363@27.[113,1134,1022,1634]" captionTargetPageId="27" captionText="Fig. 15. Sanaungulus myanmaricus sp. nov. in Burmese amber. Holotype, lateral view." pageId="27" pageNumber="358">Fig. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBEFFF1FFEBBE025CE4A058" pageId="27" pageNumber="358" type="materials_examined">
<paragraph id="8BCC643DFFBEFFF1FFEBBE025EFBA2BA" blockId="27.[113,605,816,970]" pageId="27" pageNumber="358">
<materialsCitation id="3B1B6E60FFBEFFF1FFEBBE025EFBA2BA" collectionCode="BUB" county="Probable" location="Catalog" municipality="Burmese" pageId="27" pageNumber="358" specimenCode="BUB4166" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFBEFFF1FFEBBE025FA6A238" bold="true" box="[113,224,816,840]" pageId="27" pageNumber="358">
<typeStatus id="54C8DA9FFFBEFFF1FFEBBE025F9DA238" box="[113,219,816,840]" pageId="27" pageNumber="358" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFBEFFF1FF7DBE025E00A237" box="[231,326,816,839]" pageId="27" pageNumber="358">Probable</collectingCounty>
male, adult specimen in a
<collectingMunicipality id="6BA8FE47FFBEFFF1FFEBBE635F8BA218" box="[113,205,849,872]" pageId="27" pageNumber="358">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFBEFFF1FF43BE635E61A218" box="[217,295,849,872]" country="Myanmar" name="Kachin" pageId="27" pageNumber="358">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFBEFFF1FE23BE635D4FA218" LSID="urn:lsid:plazi:treatment:03DAD52BFFBEFFF6FFEBBFEA5B10A181:8EAC32E6FFBEFFF1FE23BE635D4FA218" box="[441,521,849,872]" county="Probable" municipality="Burmese" name="Catalog" pageId="27" pageNumber="358" stateProvince="Kachin">Catalog</location>
number SNHM­
<quantity id="4C8BC9D8FFBEFFF1FF50BE405E67A2F9" box="[202,289,882,905]" metricMagnitude="2" metricUnit="m" metricValue="1.526286" pageId="27" pageNumber="358" unit="in" value="6009.0">6009 in</quantity>
the
<location id="8EAC32E6FFBEFFF1FEC0BE405E8FA2DA" LSID="urn:lsid:plazi:treatment:03DAD52BFFBEFFF6FFEBBFEA5B10A181:8EAC32E6FFBEFFF1FEC0BE405E8FA2DA" county="Probable" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="27" pageNumber="358" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FFBEFFF1FD9ABEA15D1AA2DA" LSID="urn:lsid:plazi:treatment:03DAD52BFFBEFFF6FFEBBFEA5B10A181:8EAC32E6FFBEFFF1FD9ABEA15D1AA2DA" box="[512,604,915,938]" county="Probable" municipality="Burmese" name="Muller's" pageId="27" pageNumber="358" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFBEFFF1FEDABE815EF7A2BA" box="[320,433,947,970]" collectionCode="BUB" pageId="27" pageNumber="358">BUB4166</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFBEFFF1FD19BDA85C93A1A2" blockId="27.[643,1134,154,210]" pageId="27" pageNumber="358">
<emphasis id="B907B82FFFBEFFF1FD19BDA85C5AA1C2" bold="true" box="[643,796,154,178]" pageId="27" pageNumber="358">Type locality.</emphasis>
<collectingCountry id="F36424ADFFBEFFF1FCBEBDA95CD6A1C2" box="[804,912,155,178]" name="Myanmar" pageId="27" pageNumber="358">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFBEFFF1FC01BDA95B63A1C2" box="[923,1061,155,178]" country="Myanmar" name="Kachin" pageId="27" pageNumber="358">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFBEFFF1FD19BDC35CE4A058" blockId="27.[643,1134,241,296]" pageId="27" pageNumber="358">
<emphasis id="B907B82FFFBEFFF1FD19BDC35C50A079" bold="true" box="[643,790,241,265]" pageId="27" pageNumber="358">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBEFFF1FD19BC7A5C62A30F" pageId="27" pageNumber="358" type="diagnosis">
<paragraph id="8BCC643DFFBEFFF1FD19BC7A5C62A30F" blockId="27.[643,1134,328,639]" pageId="27" pageNumber="358">
<emphasis id="B907B82FFFBEFFF1FD19BC7A5C3CA010" bold="true" box="[643,890,328,352]" pageId="27" pageNumber="358">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFFBEFFF1FCE4BC7A5DFEA00F" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="myanmaricus" status="sp. nov.">
<emphasis id="B907B82FFFBEFFF1FCE4BC7A5DFEA00F" italics="true" pageId="27" pageNumber="358">Sanaungulus myanmaricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBEFFF1FD5ABC5A5C55A00F" box="[704,787,360,383]" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by a very elongated pronotum with the surface slightly undulate, a feature present only in
<taxonomicName id="4C731FBEFFBEFFF1FC11BC9A5B0FA0CF" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[907,1097,424,447]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="nilsi" status="sp. nov.">
<emphasis id="B907B82FFFBEFFF1FC11BC9A5B0FA0CF" box="[907,1097,424,447]" italics="true" pageId="27" pageNumber="358">Sanaungulus nilsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBEFFF1FBD5BC9A5DF6A0AF" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C731FBEFFBEFFF1FD7DBCFA5CB0A0AF" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[743,1014,456,479]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="kachinensis" status="sp. nov.">
<emphasis id="B907B82FFFBEFFF1FD7DBCFA5CB0A0AF" box="[743,1014,456,479]" italics="true" pageId="27" pageNumber="358">Sanaungulus kachinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBEFFF1FC64BCFA5B14A0AF" box="[1022,1106,456,479]" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="4C731FBEFFBEFFF1FBC0BCFA5DF6A08F" authorityName="Fanti &amp; Müller" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="nilsi">
<emphasis id="B907B82FFFBEFFF1FBC0BCFA5DF6A08F" italics="true" pageId="27" pageNumber="358">S. nilsi</emphasis>
</taxonomicName>
has a pronotum more concave in the middle and more bordered at the margins (especially at the posterior margin), while
<taxonomicName id="4C731FBEFFBEFFF1FC4FBF1A5B28A34F" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[981,1134,552,575]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="kachinensis">
<emphasis id="B907B82FFFBEFFF1FC4FBF1A5B28A34F" box="[981,1134,552,575]" italics="true" pageId="27" pageNumber="358">S. kachinensis</emphasis>
</taxonomicName>
has a pronotum that is shorter and with a transversal incision.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBEFFF6FD19BFAC5D1CA482" lastPageId="28" lastPageNumber="359" pageId="27" pageNumber="358" type="description">
<paragraph id="8BCC643DFFBEFFF1FD19BFAC5DBBA266" blockId="27.[643,1134,670,790]" pageId="27" pageNumber="358">
<emphasis id="B907B82FFFBEFFF1FD19BFAC5C48A3C6" bold="true" box="[643,782,670,694]" pageId="27" pageNumber="358">Description.</emphasis>
Adult, winged, slender. Probable male, defined on the basis of slender abdomen and long antennae. Body length
<quantity id="4C8BC9D8FFBEFFF1FC40BFED5B69A386" box="[986,1071,735,758]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="358" unit="mm" value="3.2">3.2 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFBEFFF6FD19BE075D1CA482" blockId="27.[643,1135,821,972]" lastBlockId="28.[151,644,155,1522]" lastPageId="28" lastPageNumber="359" pageId="27" pageNumber="358">Head transverse, constricted (triangular­shaped) behind the eyes, equipped with pubescence and punctation. Eyes prominent, large, roundish, inserted in the upper and lateral part of the head. Mandibles elongated, falciform. Maxillary palps 4­ segmented with the last palpomere securiform. Labial palps 3­ segmented, with the last palpomere elongated and strongly securiform.Antennae 11­ segmented, pectinate, surpassing half of elytra and reaching half of abdomen, antennal insertion in the eyes proximity; scape slender, elongated, slightly enlarged apically; antennomere II short, swollen apically; antennomere III elongated, filiform, about 1.8 times longer than second; antennomere IV longer and stouter than previous one; antennomeres VIX extremely long, each equipped near the base with a long antennal process which is very long and rounded apically; antennomere X shorter than previous ones, equipped with an antennal process shorter than others and slightly squadrate at the base; antennomere XI short and robust, with rounded apex; all the antennomeres pubescent. Pronotum very elongated, narrower than head, anterior margin undulate, posterior margin extend backward in the middle and slightly bordered, sides straight and bordered, corners obtuse, pronotum disc irregular with small concavity and drafts. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, at humeri wider than pronotum, posteriorly parallel­sided and slightly dehiscent apically, rounded apexes, surface pubescent. Metathoracic wings transparent, slightly longer than elytra. Sternum elongated, sternites transverse, last abdominal segment wide and with rounded apex. Legs thin, extremely long, pubescent; coxae massive; trochanters elongated, with rounded apex; femora almost straight, cylindrical, slightly sturdier than tibiae; tibiae, thin, cylindrical, extremely long, with a spur near the apex, pro­ and mesotibiae shorter than pro­ and mesofemora, metatibiae as long as metafemora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 2.0 times longer than second; third tarsomere short, triangular­shaped; fourth strongly bilobed; fifth tarsomere extremely elongated, thin, flat and curved; claws simple, long, pointed, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFBEFFF1FFEBBBB35B4CA7E8" box="[113,1034,1665,1688]" pageId="27" pageNumber="358" startId="27.[113,152,1665,1688]" targetBox="[113,1134,1022,1634]" targetPageId="27" targetType="figure">
<paragraph id="8BCC643DFFBEFFF1FFEBBBB35B4CA7E8" blockId="27.[113,1034,1665,1688]" box="[113,1034,1665,1688]" pageId="27" pageNumber="358">
Fig. 15.
<taxonomicName id="4C731FBEFFBEFFF1FF50BBB35EACA7E8" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[202,490,1665,1688]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="358" phylum="Arthropoda" rank="species" species="myanmaricus" status="sp. nov.">
<emphasis id="B907B82FFFBEFFF1FF50BBB35EACA7E8" box="[202,490,1665,1688]" italics="true" pageId="27" pageNumber="358">Sanaungulus myanmaricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBEFFF1FE6ABBB35D05A7E8" box="[496,579,1665,1688]" pageId="27" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. Holotype, lateral view.
</paragraph>
</caption>
<subSubSection id="C36937B6FFB9FFF6FF0DBB235E85A7EF" pageId="28" pageNumber="359" type="etymology">
<paragraph id="8BCC643DFFB9FFF6FF0DBB235DC2A738" blockId="28.[151,644,1553,1608]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FF0DBB235E5EA759" bold="true" box="[151,280,1553,1577]" pageId="28" pageNumber="359">Etymology.</emphasis>
Derived from the toponym
<collectingCountry id="F36424ADFFB9FFF6FDA6BB235F86A738" name="Myanmar" pageId="28" pageNumber="359">Myanmar</collectingCountry>
(nation where the amber mines are located).
</paragraph>
<paragraph id="8BCC643DFFB9FFF6FF0DBB5A5E85A7EF" blockId="28.[151,643,1640,1695]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FF0DBB5A5E7DA7F0" bold="true" box="[151,315,1640,1664]" pageId="28" pageNumber="359">Syninclusions.</emphasis>
Air bubbles, detritus, stellate hairs, an unidentified insect.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFB9FFF6FD32BDA85B10A181" pageId="28" pageNumber="359" type="discussion">
<paragraph id="8BCC643DFFB9FFF6FD32BDA85B10A181" blockId="28.[680,1172,154,241]" pageId="28" pageNumber="359">
<emphasis id="B907B82FFFB9FFF6FD32BDA85C52A1C2" bold="true" box="[680,788,154,178]" pageId="28" pageNumber="359">Remarks.</emphasis>
Piece of clear, golden amber that measures 18x11x
<quantity id="4C8BC9D8FFB9FFF6FCAEBD885C30A1A1" box="[820,886,186,209]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="359" unit="mm" value="4.0">4 mm</quantity>
. The inclusion is complete but well visible only laterally (one side).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,203 @@
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<mods:title id="B3E758ABE75D45A887FFDDF459497A17">Four new species of the genus Metapocyrtus Heller, 1912 (Coleoptera: Curculionidae: Pachyrhynchini) from Mindanao Island, Philippines</mods:title>
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<subSubSection id="C3566567B445B03D31870D4FE532FE26" pageId="8" pageNumber="291" type="nomenclature">
<paragraph id="8BF336ECB445B03D31870D4FE532FE26" blockId="8.[189,662,331,416]" pageId="8" pageNumber="291">
<heading id="D0BB8180B445B03D31870D4FE532FE26" bold="true" centered="true" fontSize="10" level="2" pageId="8" pageNumber="291" reason="4">
<emphasis id="B938EAFEB445B03D31870D4FE532FE26" bold="true" pageId="8" pageNumber="291">
<taxonomicName id="4C4C4D6FB445B03D31870D4FE56DFE25" ID-CoL="C3BV8" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="291" phylum="Arthropoda" rank="species" species="pseudolanaoensis" status="sp. nov." subGenus="Trachycyrtus">
<emphasis id="B938EAFEB445B03D31870D4FE56DFE25" bold="true" italics="true" pageId="8" pageNumber="291">Metapocyrtus (Trachycyrtus) pseudolanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B445B03D301E0D6DE532FE26" box="[292,379,361,385]" pageId="8" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3566567B445B03D31870D8CE571FE07" box="[189,312,392,416]" pageId="8" pageNumber="291" type="description">
<paragraph id="8BF336ECB445B03D31870D8CE571FE07" blockId="8.[189,662,331,416]" box="[189,312,392,416]" pageId="8" pageNumber="291">
(
<figureCitation id="13772A69B445B03D31FF0D8CE55FFE07" box="[197,278,392,416]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A M. (M.) tetramaculatus sp. nov.; B M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C M. (M.) fulvus sp. nov.; D M. (T.) unalineatus sp. nov.; E M. (T.) pseudolanaoensis sp. nov.; F M. (M.) lanaoensis Bollino, 2021." pageId="8" pageNumber="291">Fig. 1E</figureCitation>
,
<figureCitation id="13772A69B445B03D30180D8CE579FE07" box="[290,304,392,416]" captionStart="Fig" captionStartId="9.[114,153,1355,1379]" captionTargetBox="[114,1043,171,1349]" captionTargetId="figure-118@9.[114,1043,171,1349]" captionTargetPageId="9" captionText="Fig. 7. Habitus of M. (T.) pseudolanaoensis sp. nov., male, holotype; A dorsal view; D lateral view; B aedegal body, lateral view; sternite IX." pageId="8" pageNumber="291">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3566567B445B03C33810CB5E6F1FFD1" lastPageId="9" lastPageNumber="292" pageId="8" pageNumber="291" type="materials_examined">
<paragraph id="8BF336ECB445B03D33810CB5E0D9FEC4" blockId="8.[699,1178,177,355]" pageId="8" pageNumber="291">
<emphasis id="B938EAFEB445B03D33810CB5E720FF6E" bold="true" box="[699,873,177,201]" pageId="8" pageNumber="291">Type material.</emphasis>
<materialsCitation id="3B243CB1B445B03D324C0CB5E79FFE82" collectingDate="2015-12" country="Philippines" county="Sibagat" location="Mindanao" municipality="Agusan" pageId="8" pageNumber="291" specimenCount="1" typeStatus="holotype">
<emphasis id="B938EAFEB445B03D324C0CB5E078FF6E" bold="true" box="[886,1073,177,201]" pageId="8" pageNumber="291">
<typeStatus id="54F7884EB445B03D324C0CB5E796FF6E" box="[886,991,177,201]" pageId="8" pageNumber="291" type="holotype">Holotype</typeStatus>
, male:
</emphasis>
<collectingCountry id="F35B767CB445B03D357C0CB6E754FF4F" name="Philippines" pageId="8" pageNumber="291">PHILIPPINES</collectingCountry>
/
<location id="8E936037B445B03D32790CD4E7FBFF4F" LSID="urn:lsid:plazi:treatment:03E587FAB445B03E31870D4FE5CDFA9F:8E936037B445B03D32790CD4E7FBFF4F" box="[835,946,208,232]" country="Philippines" county="Sibagat" municipality="Agusan" name="Mindanao" pageId="8" pageNumber="291">Mindanao</location>
,
<collectingMunicipality id="6B97AC96B445B03D32FF0CD4E050FF4F" box="[965,1049,208,232]" pageId="8" pageNumber="291">Agusan</collectingMunicipality>
,
<collectingCounty id="62924E60B445B03D35170CD4E034FF4F" box="[1069,1149,208,232]" pageId="8" pageNumber="291">Sibagat</collectingCounty>
/
<date id="FFF2102CB445B03D33810CEBE76DFEA0" box="[699,804,239,263]" pageId="8" pageNumber="291" value="2015-12">
<collectingDate id="EFB6E9C4B445B03D33810CEBE76DFEA0" box="[699,804,239,263]" pageId="8" pageNumber="291" value="2015-12">XII. 2015</collectingDate>
</date>
/ local collector leg. (white rectangular label, printed) //
</materialsCitation>
<typeStatus id="54F7884EB445B03D32D00D09E0C9FE82" box="[1002,1152,269,293]" pageId="8" pageNumber="291" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C4C4D6FB445B03D33810D29E795FEE2" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[699,988,301,325]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="291" phylum="Arthropoda" rank="species" species="pseudolanaoensis" subGenus="Trachycyrtus">
<emphasis id="B938EAFEB445B03D33810D29E795FEE2" box="[699,988,301,325]" italics="true" pageId="8" pageNumber="291">M. (T.) pseudolanaoensis</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023 (red rectangular label, printed).
</paragraph>
<caption id="DF336664B445B03D31870A1BE6B3F9F1" pageId="8" pageNumber="291" startId="8.[189,228,1567,1591]" targetBox="[194,1144,422,1537]" targetPageId="8" targetType="figure">
<paragraph id="8BF336ECB445B03D31870A1BE07EF990" blockId="8.[189,1079,1567,1622]" box="[189,1079,1567,1592]" pageId="8" pageNumber="291">
Fig. 6. Habitus of
<taxonomicName id="4C4C4D6FB445B03D30BA0A24E604F99F" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[384,589,1568,1592]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="291" phylum="Arthropoda" rank="species" species="unalineatus" status="sp. nov." subGenus="Trachycyrtus">
<emphasis id="B938EAFEB445B03D30BA0A24E604F99F" box="[384,589,1568,1592]" italics="true" pageId="8" pageNumber="291">M. (T.) unalineatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B445B03D336E0A1BE6E1F990" box="[596,680,1567,1591]" pageId="8" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
A dorsal view, male, holotype; B
</paragraph>
<paragraph id="8BF336ECB445B03D31870A3AE6B3F9F1" blockId="8.[189,1079,1567,1622]" box="[189,762,1598,1622]" pageId="8" pageNumber="291">dorsal view, female; C lateral view, male, holotype.</paragraph>
</caption>
<paragraph id="8BF336ECB445B03D35560AA2E0DDF919" blockId="8.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="8" pageNumber="291">291</paragraph>
<paragraph id="8BF336ECB444B03C333E0C67E6F1FFD1" blockId="9.[516,696,99,118]" box="[516,696,99,118]" pageId="9" pageNumber="292">
<emphasis id="B938EAFEB444B03C333E0C67E6F1FFD1" box="[516,696,99,118]" italics="true" pageId="9" pageNumber="292">Rukmane-Bārbale A.</emphasis>
</paragraph>
</subSubSection>
<caption id="DF336664B444B03C3148094FE681FA25" pageId="9" pageNumber="292" startId="9.[114,153,1355,1379]" targetBox="[114,1043,171,1349]" targetPageId="9" targetType="figure">
<paragraph id="8BF336ECB444B03C3148094FE681FA25" blockId="9.[114,1078,1355,1410]" pageId="9" pageNumber="292">
Fig. 7. Habitus of
<taxonomicName id="4C4C4D6FB444B03C300E0948E60BFAC3" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[308,578,1356,1380]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="292" phylum="Arthropoda" rank="species" species="pseudolanaoensis" status="sp. nov." subGenus="Trachycyrtus">
<emphasis id="B938EAFEB444B03C300E0948E60BFAC3" box="[308,578,1356,1380]" italics="true" pageId="9" pageNumber="292">M. (T.) pseudolanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B444B03C3370094FE6E9FAC4" box="[586,672,1355,1379]" pageId="9" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
, male, holotype; A dorsal view; D lateral view; B aedegal body, lateral view; sternite IX.
</paragraph>
</caption>
<subSubSection id="C3566567B444B03E314809A3E5AEFBC6" lastPageId="11" lastPageNumber="294" pageId="9" pageNumber="292" type="description">
<paragraph id="8BF336ECB444B03C314809A3E510F9BC" blockId="9.[114,593,1447,1563]" pageId="9" pageNumber="292">
<emphasis id="B938EAFEB444B03C314809A3E598FA18" bold="true" box="[114,465,1447,1471]" pageId="9" pageNumber="292">
Description.
<typeStatus id="54F7884EB444B03C303709A3E53EFA18" box="[269,375,1447,1471]" pageId="9" pageNumber="292" type="holotype">Holotype</typeStatus>
, male.
</emphasis>
(
<figureCitation id="13772A69B444B03C30DE09A3E601FA18" box="[484,584,1447,1471]" captionStart="Fig" captionStartId="9.[114,153,1355,1379]" captionTargetBox="[114,1043,171,1349]" captionTargetId="figure-118@9.[114,1043,171,1349]" captionTargetPageId="9" captionText="Fig. 7. Habitus of M. (T.) pseudolanaoensis sp. nov., male, holotype; A dorsal view; D lateral view; B aedegal body, lateral view; sternite IX." pageId="9" pageNumber="292">Fig. 7A</figureCitation>
) Dimensions: LB: 6,6, LR: 1,4, WR: 1,1, LP: 2,2, WP: 2,1, LE: 4,4, WE: 2,9. N = 1 for all measurements.
</paragraph>
<paragraph id="8BF336ECB444B03C31480A45E652F9D0" blockId="9.[114,593,1601,1655]" pageId="9" pageNumber="292">
<emphasis id="B938EAFEB444B03C31480A45E4B3F9FE" bold="true" box="[114,250,1601,1625]" pageId="9" pageNumber="292">Coloration.</emphasis>
Integument black, shiny, with markings of round, shiny orange scales.
</paragraph>
<paragraph id="8BF336ECB444B03F31480AA2E6D5FF4F" blockId="9.[114,154,1702,1726]" lastBlockId="10.[189,668,178,232]" lastPageId="10" lastPageNumber="293" pageId="9" pageNumber="292">
292
<emphasis id="B938EAFEB444B03C334A09A3E6F3FA18" bold="true" box="[624,698,1447,1471]" pageId="9" pageNumber="292">Head.</emphasis>
Dorsal surface slightly rugose, with single orange scales; surface between eyes with short white setae at basal part; genae with longer, white to metallic green setae; moderate medial longitudinal groove from base of the eyes to base anterior margin of prothorax; transverse, moderate furrow at base of eyes confluent with medial longitudinal groove, not reaching lateral margins;
</paragraph>
<paragraph id="8BF336ECB447B03F33810CB6E75BFF4F" blockId="10.[699,1179,178,232]" pageId="10" pageNumber="293">eyes moderate, weakly convex from outline of head.</paragraph>
<caption id="DF336664B447B03F31870956E013FACD" box="[189,1114,1362,1386]" pageId="10" pageNumber="293" startId="10.[189,228,1362,1386]" targetBox="[205,964,238,1342]" targetPageId="10" targetType="figure">
<paragraph id="8BF336ECB447B03F31870956E013FACD" blockId="10.[189,1114,1362,1386]" box="[189,1114,1362,1386]" pageId="10" pageNumber="293">
Fig. 8. Habitus of
<taxonomicName id="4C4C4D6FB447B03F30BA0956E6AEFACD" authority="Bollino, 2021" authorityName="Bollino" authorityYear="2021" box="[384,743,1362,1386]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="lanaoensis" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB447B03F30BA0956E603FACD" box="[384,586,1362,1386]" italics="true" pageId="10" pageNumber="293">M. (M.) lanaoensis</emphasis>
Bollino, 2021
</taxonomicName>
; A dorsal view; B lateral view.
</paragraph>
</caption>
<paragraph id="8BF336ECB447B03F3187098BE659F9F8" blockId="10.[189,668,1423,1631]" pageId="10" pageNumber="293">
<emphasis id="B938EAFEB447B03F3187098BE566FA00" bold="true" box="[189,303,1423,1447]" pageId="10" pageNumber="293">Rostrum.</emphasis>
(
<figureCitation id="13772A69B447B03F307D098BE5ECFA00" box="[327,421,1423,1447]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A M. (M.) tetramaculatus sp. nov.; B M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C M. (M.) fulvus sp. nov.; D M. (T.) unalineatus sp. nov.; E M. (T.) pseudolanaoensis sp. nov.; F M. (M.) lanaoensis Bollino, 2021." pageId="10" pageNumber="293">Fig. 1E</figureCitation>
) longer than wide (LR/WR: 1,27), granulate in all length; short white setae at basal part; dorsum with shallow medial longitudinal furrow and corresponding shallow concavity; in dorsal contour straight; lateral parts covered with long white to shiny green setae.
</paragraph>
<paragraph id="8BF336ECB447B03F3381098BE76FF985" blockId="10.[699,1179,1423,1570]" pageId="10" pageNumber="293">
<emphasis id="B938EAFEB447B03F3381098BE766FA00" bold="true" box="[699,815,1423,1447]" pageId="10" pageNumber="293">Antennae</emphasis>
with funicular segment I 1,5 times longer than II, segment II nearly two times longer than III, segments III VI as long as wide, segment VII larger, wider than long.
</paragraph>
<paragraph id="8BF336ECB447B03F33810A43E0D3F9D9" blockId="10.[699,1178,1607,1662]" pageId="10" pageNumber="293">
<emphasis id="B938EAFEB447B03F33810A43E77EF9F8" bold="true" box="[699,823,1607,1631]" pageId="10" pageNumber="293">Prothorax</emphasis>
sub globular, nearly same width and length (LP/WP: 1,04), strongly rugose,
</paragraph>
<paragraph id="8BF336ECB447B03E35560AA2E579FE07" blockId="10.[1132,1172,1702,1726]" lastBlockId="11.[114,593,178,416]" lastPageId="11" lastPageNumber="294" pageId="10" pageNumber="293">293 without setae or pubescence on disc; widest just at the middle; with the following markings of round, shiny orange shales: 1) transverse line along anterior margin in all length; 2) transverse line just after the middle in all length; shallow longitudinal medial groove on disc; anterior margin with short pubescence.</paragraph>
<paragraph id="8BF336ECB446B03E31480DC1E4ADFCEA" blockId="11.[114,593,453,845]" pageId="11" pageNumber="294">
<emphasis id="B938EAFEB446B03E31480DC1E488FE7A" bold="true" box="[114,193,453,477]" pageId="11" pageNumber="294">Elytra</emphasis>
sub-ellipsoidal, (LE/WE: 1,51), wider than prothorax, twice as long as prothorax (WE/WP: 1,38; LE/LP: 2,0); rugose in all length, without expressed puncture rows; with short brown setae along suture from apical ½ to apex; each elytron with the following scally markings of round shiny orange scales: 1) two irregular spots at basal part; 2) 4-5 smaller spots at medial part; 3) three elongated longitudinal spots at apical part; 4) one longitudinal line at apical part near suture. Male genitalia as shown in
<figureCitation id="13772A69B446B03E31480F31E4F9FCEA" box="[114,176,821,845]" captionStart="Fig" captionStartId="9.[114,153,1355,1379]" captionTargetBox="[114,1043,171,1349]" captionTargetId="figure-118@9.[114,1043,171,1349]" captionTargetPageId="9" captionText="Fig. 7. Habitus of M. (T.) pseudolanaoensis sp. nov., male, holotype; A dorsal view; D lateral view; B aedegal body, lateral view; sternite IX." pageId="11" pageNumber="294">Fig. 7</figureCitation>
B-C.
</paragraph>
<paragraph id="8BF336ECB446B03E31480F77E578FC2C" blockId="11.[114,305,883,907]" box="[114,305,883,907]" pageId="11" pageNumber="294">Female unknown.</paragraph>
<paragraph id="8BF336ECB446B03E31480FB4E5AEFBC6" blockId="11.[114,593,944,1122]" pageId="11" pageNumber="294">
<emphasis id="B938EAFEB446B03E31480FB4E522FC6F" bold="true" box="[114,363,944,968]" pageId="11" pageNumber="294">Differential analyses.</emphasis>
The new species by scally markings on prothorax and elytra can be referred to
<taxonomicName id="4C4C4D6FB446B03E302F0FEAE4EEFB83" authority="Bollino, 2021" authorityName="Bollino" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="294" phylum="Arthropoda" rank="species" species="lanaoensis" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB446B03E302F0FEAE5A7FBA1" box="[277,494,1006,1030]" italics="true" pageId="11" pageNumber="294">M. (M.) lanaoensis</emphasis>
Bollino, 2021
</taxonomicName>
(
<figureCitation id="13772A69B446B03E318D0808E4B0FB83" box="[183,249,1036,1060]" captionStart="Fig" captionStartId="10.[189,228,1362,1386]" captionTargetBox="[205,964,238,1342]" captionTargetId="figure-142@10.[189,991,232,1355]" captionTargetPageId="10" captionText="Fig. 8. Habitus of M. (M.) lanaoensis Bollino, 2021; A dorsal view; B lateral view." pageId="11" pageNumber="294">Fig. 8</figureCitation>
,
<figureCitation id="13772A69B446B03E303C0808E51AFB83" box="[262,339,1036,1060]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A M. (M.) tetramaculatus sp. nov.; B M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C M. (M.) fulvus sp. nov.; D M. (T.) unalineatus sp. nov.; E M. (T.) pseudolanaoensis sp. nov.; F M. (M.) lanaoensis Bollino, 2021." pageId="11" pageNumber="294">Fig. 1F</figureCitation>
) but can be easily distinguished by features of rostrum that refers species to subgenera
<emphasis id="B938EAFEB446B03E306E084EE5A9FBC5" box="[340,480,1098,1122]" italics="true" pageId="11" pageNumber="294">Trachycyrtus</emphasis>
.
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<paragraph id="8BF336ECB446B03E31480883E491FB1A" blockId="11.[114,593,1159,1213]" pageId="11" pageNumber="294">
<emphasis id="B938EAFEB446B03E31480883E543FB38" bold="true" box="[114,266,1159,1183]" pageId="11" pageNumber="294">Distribution.</emphasis>
Mindanao Island, Agusan Province.
</paragraph>
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<subSubSection id="C3566567B446B03E314808E7E5CDFA9F" pageId="11" pageNumber="294" type="etymology">
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<emphasis id="B938EAFEB446B03E314808E7E4B3FB5C" bold="true" box="[114,250,1251,1275]" pageId="11" pageNumber="294">Etymology.</emphasis>
The specific epithet is related to similar general appearance to
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<emphasis id="B938EAFEB446B03E30CB0906E4A2FA9F" italics="true" pageId="11" pageNumber="294">M. (M.) lanaoensis</emphasis>
Bollino, 2021
</taxonomicName>
.
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<emphasis id="B938EAFEB44AB032334A082FE006FBE4" bold="true" box="[624,1103,1067,1091]" italics="true" pageId="7" pageNumber="290">Metapocyrtus (Trachycyrtus) unalineatus</emphasis>
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<paragraph id="8BF336ECB44AB032334A084DE68DFBC6" blockId="7.[624,1103,1067,1152]" box="[624,708,1097,1121]" pageId="7" pageNumber="290">sp. nov.</paragraph>
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<paragraph id="8BF336ECB44AB032334A086CE779FB27" blockId="7.[624,1103,1067,1152]" box="[624,816,1128,1152]" pageId="7" pageNumber="290">
(
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,
<figureCitation id="13772A69B44AB03233E2086CE6ACFB27" box="[728,741,1128,1152]" captionStart="Fig" captionStartId="5.[114,153,1533,1557]" captionTargetBox="[120,1022,855,1503]" captionTargetId="figure-308@5.[114,1032,846,1526]" captionTargetPageId="5" captionText="Fig. 4. Male genitalia of selected species, A, C, E aedegal body in lateral view; B, D, F sternite IX; A, B M. (M.) tetramaculatus sp. nov., C, D M. (M.) fulvus sp. nov., E, F M. (T.) unalineatus sp. nov." pageId="7" pageNumber="290">4</figureCitation>
E-F, 6)
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<emphasis id="B938EAFEB44AB032334A08A1E722FB1A" bold="true" box="[624,875,1189,1213]" pageId="7" pageNumber="290">Differential analyses.</emphasis>
The new species is referred to subgenera
<emphasis id="B938EAFEB44AB032325708C1E7B0FB7A" box="[877,1017,1221,1245]" italics="true" pageId="7" pageNumber="290">Trachycyrtus</emphasis>
by following characters of rostrum: transverse groove that is not extending to lateral edges, shallow medial ridge at apical part and rounded lateral edges. Along subgeneric conquerors, the new species is unique in having longitudinal scale line along suture of elytra while rest of the species have transverse line markings, spots or single dispersed scales.
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<paragraph id="8BF336ECB44AB032334A0A11E75FF9EB" blockId="7.[624,1103,1557,1612]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB032334A0A11E741F98A" bold="true" box="[624,776,1557,1581]" pageId="7" pageNumber="290">Distribution.</emphasis>
Mindanao Island, Lanao Province, Wao.
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<emphasis id="B938EAFEB445B03D31870CB5E50FFF6E" bold="true" box="[189,326,177,201]" pageId="8" pageNumber="291">Etymology.</emphasis>
The specific epithet is a latinised designation of one scally longitudinallinealong suture of elytra. One una, line lineata.
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<emphasis id="B938EAFEB44CB034334A0D13E6D9FEE9" bold="true" italics="true" pageId="1" pageNumber="284">Metapocyrtus (Metapocyrtus) tetramaculatus</emphasis>
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(
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,
<figureCitation id="13772A69B44CB03433E20D50E6AEFECB" box="[728,743,340,364]" captionStart="Fig" captionStartId="2.[189,228,1386,1410]" captionTargetBox="[193,1172,171,1380]" captionTargetId="figure-114@2.[193,1172,171,1380]" captionTargetPageId="2" captionText="Fig. 2. Habitus of M. (M.) tetramaculatus sp. nov. A dorsal view, male, holotype; B dorsal view, female; C lateral view, male, holotype." pageId="1" pageNumber="284">2</figureCitation>
,
<figureCitation id="13772A69B44CB03433C80D50E765FECB" box="[754,812,340,364]" captionStart="Fig" captionStartId="5.[114,153,1533,1557]" captionTargetBox="[120,1022,855,1503]" captionTargetId="figure-308@5.[114,1032,846,1526]" captionTargetPageId="5" captionText="Fig. 4. Male genitalia of selected species, A, C, E aedegal body in lateral view; B, D, F sternite IX; A, B M. (M.) tetramaculatus sp. nov., C, D M. (M.) fulvus sp. nov., E, F M. (T.) unalineatus sp. nov." pageId="1" pageNumber="284">4A,B</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF336664B44CB03431480813E545FB2C" pageId="1" pageNumber="284" startId="1.[114,153,1047,1071]" targetBox="[114,1096,447,1040]" targetPageId="1" targetType="figure">
<paragraph id="8BF336ECB44CB03431480813E050FB88" blockId="1.[114,1070,1047,1163]" box="[114,1049,1047,1071]" pageId="1" pageNumber="284">
Fig. 1. Dorsal view of rostrum of selected species: A
<taxonomicName id="4C4C4D6FB44CB03433850813E7FEFB88" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[703,951,1047,1071]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB03433850813E7FEFB88" box="[703,951,1047,1071]" italics="true" pageId="1" pageNumber="284">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44CB03432840813E05CFB88" box="[958,1045,1047,1071]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
;
</paragraph>
<paragraph id="8BF336ECB44CB03431480831E545FB2C" blockId="1.[114,1070,1047,1163]" pageId="1" pageNumber="284">
B
<taxonomicName id="4C4C4D6FB44CB03431A40832E6ECFBEA" authority="Cabras &amp; Medina, 2021" authorityName="Cabras &amp; Medina" authorityYear="2021" box="[158,677,1077,1102]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="flavomaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB03431A40832E5D3FBE9" box="[158,410,1078,1102]" italics="true" pageId="1" pageNumber="284">M. (M.) flavomaculatus</emphasis>
Cabras &amp; Medina, 2021
</taxonomicName>
; C
<taxonomicName id="4C4C4D6FB44CB03433E60832E73DFBE9" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[732,884,1078,1102]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="fulvus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB03433E60832E73DFBE9" box="[732,884,1078,1102]" italics="true" pageId="1" pageNumber="284">M. (M.) fulvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44CB03432410831E79BFBEA" box="[891,978,1077,1101]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
; D
<taxonomicName id="4C4C4D6FB44CB03435300832E555FBCA" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="unalineatus" status="sp. nov." subGenus="Trachycyrtus">
<emphasis id="B938EAFEB44CB03435300832E555FBCA" italics="true" pageId="1" pageNumber="284">M. (T.) unalineatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44CB03430190850E533FBCB" box="[291,378,1108,1132]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
; E
<taxonomicName id="4C4C4D6FB44CB03430950851E6F7FBCA" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[431,702,1109,1133]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="pseudolanaoensis" status="sp. nov." subGenus="Trachycyrtus">
<emphasis id="B938EAFEB44CB03430950851E6F7FBCA" box="[431,702,1109,1133]" italics="true" pageId="1" pageNumber="284">M. (T.) pseudolanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44CB03433FF0850E754FBCB" box="[709,797,1108,1132]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
; F
<taxonomicName id="4C4C4D6FB44CB034326B0851E54EFB2C" authority="Bollino, 2021" authorityName="Bollino" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="lanaoensis" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB034326B0851E055FBCA" box="[849,1052,1109,1133]" italics="true" pageId="1" pageNumber="284">M. (M.) lanaoensis</emphasis>
Bollino, 2021
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3566567B44CB034314808B4E683FAC5" pageId="1" pageNumber="284" type="materials_examined">
<paragraph id="8BF336ECB44CB034314808B4E644FAC5" blockId="1.[114,593,1200,1378]" pageId="1" pageNumber="284">
<emphasis id="B938EAFEB44CB034314808B4E569FB6F" bold="true" box="[114,288,1200,1224]" pageId="1" pageNumber="284">Type material.</emphasis>
<materialsCitation id="3B243CB1B44CB034301608B4E513FA83" collectingDate="2015-09" country="Philippines" county="Sibagat" location="Mindanao" municipality="Agusan" pageId="1" pageNumber="284" specimenCount="1" typeStatus="holotype">
<emphasis id="B938EAFEB44CB034301608B4E5AEFB6F" bold="true" box="[300,487,1200,1224]" pageId="1" pageNumber="284">
<typeStatus id="54F7884EB44CB034301608B4E5DCFB6F" box="[300,405,1200,1224]" pageId="1" pageNumber="284" type="holotype">Holotype</typeStatus>
, male:
</emphasis>
<collectingCountry id="F35B767CB44CB03430C708B4E49DFB40" name="Philippines" pageId="1" pageNumber="284">PHILIPPINES</collectingCountry>
/
<location id="8E936037B44CB03431D308CBE511FB40" LSID="urn:lsid:plazi:treatment:03E587FAB44CB036334A0D13E6E7FA76:8E936037B44CB03431D308CBE511FB40" box="[233,344,1231,1255]" country="Philippines" county="Sibagat" municipality="Agusan" name="Mindanao" pageId="1" pageNumber="284">Mindanao</location>
,
<collectingMunicipality id="6B97AC96B44CB034305908CBE5FEFB40" box="[355,439,1231,1255]" pageId="1" pageNumber="284">Agusan</collectingMunicipality>
,
<collectingCounty id="62924E60B44CB03430F808CBE65BFB40" box="[450,530,1231,1255]" pageId="1" pageNumber="284">Sibagat</collectingCounty>
/
<date id="FFF2102CB44CB034331D08CBE4E1FAA2" pageId="1" pageNumber="284" value="2015-09">
<collectingDate id="EFB6E9C4B44CB034331D08CBE4E1FAA2" pageId="1" pageNumber="284" value="2015-09">IX. 2015</collectingDate>
</date>
/ local collector leg. (white rectangular label, printed) //
</materialsCitation>
<typeStatus id="54F7884EB44CB034304B0908E64EFA83" box="[369,519,1292,1316]" pageId="1" pageNumber="284" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C4C4D6FB44CB03433170909E514FAE4" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="tetramaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB03433170909E514FAE4" italics="true" pageId="1" pageNumber="284">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023 (red rectangular label, printed).
</paragraph>
<paragraph id="8BF336ECB44CB03431480983E603F9BD" blockId="1.[114,593,1415,1562]" pageId="1" pageNumber="284">
<emphasis id="B938EAFEB44CB03431480983E536FA38" bold="true" box="[114,383,1415,1439]" pageId="1" pageNumber="284">
<typeStatus id="54F7884EB44CB03431480983E4A1FA38" box="[114,232,1415,1439]" pageId="1" pageNumber="284" type="paratype">Paratypes</typeStatus>
: (
<specimenCount id="9D4AFD65B44CB03431C50983E53AFA38" box="[255,371,1415,1439]" count="3" pageId="1" pageNumber="284" type="female">3 females</specimenCount>
):
</emphasis>
<materialsCitation id="3B243CB1B44CB03430A80983E53EFA7B" collectingDate="2015-09" country="Philippines" county="Mindanao" location="Sibagat" municipality="Agusan" pageId="1" pageNumber="284" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F35B767CB44CB03430A80983E671FA38" box="[402,568,1415,1439]" name="Philippines" pageId="1" pageNumber="284">PHILIPPINES</collectingCountry>
/
<collectingCounty id="62924E60B44CB034314809A1E4A8FA1A" box="[114,225,1445,1469]" pageId="1" pageNumber="284">Mindanao</collectingCounty>
,
<collectingMunicipality id="6B97AC96B44CB03431C909A1E50EFA1A" box="[243,327,1445,1469]" pageId="1" pageNumber="284">Agusan</collectingMunicipality>
,
<location id="8E936037B44CB034306009A1E5E3FA1A" LSID="urn:lsid:plazi:treatment:03E587FAB44CB036334A0D13E6E7FA76:8E936037B44CB034306009A1E5E3FA1A" box="[346,426,1445,1469]" country="Philippines" county="Mindanao" municipality="Agusan" name="Sibagat" pageId="1" pageNumber="284">Sibagat</location>
/
<date id="FFF2102CB44CB03430F709A1E67CFA1A" box="[461,565,1445,1469]" pageId="1" pageNumber="284" value="2015-09">
<collectingDate id="EFB6E9C4B44CB03430F709A1E67CFA1A" box="[461,565,1445,1469]" pageId="1" pageNumber="284" value="2015-09">IX. 2015</collectingDate>
</date>
/ local collector leg. //
</materialsCitation>
<materialsCitation id="3B243CB1B44CB03430B709C0E603F9BD" collectingDate="2015-10" country="Philippines" location="Cabanglasan" municipality="Mindanao" pageId="1" pageNumber="284" specimenCount="1" stateProvince="Bukidnon" typeStatus="paratype">
<collectingCountry id="F35B767CB44CB03430B709C0E67AFA7B" box="[397,563,1476,1500]" name="Philippines" pageId="1" pageNumber="284">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="6B97AC96B44CB034314809E7E4A8FA5C" box="[114,225,1507,1531]" pageId="1" pageNumber="284">Mindanao</collectingMunicipality>
,
<collectingRegion id="4988F80EB44CB03431C209E7E52DFA5C" box="[248,356,1507,1531]" country="Philippines" name="Bukidnon" pageId="1" pageNumber="284">Bukidnon</collectingRegion>
,
<location id="8E936037B44CB034304109E7E64AFA5C" LSID="urn:lsid:plazi:treatment:03E587FAB44CB036334A0D13E6E7FA76:8E936037B44CB034304109E7E64AFA5C" box="[379,515,1507,1531]" country="Philippines" municipality="Mindanao" name="Cabanglasan" pageId="1" pageNumber="284" stateProvince="Bukidnon">Cabanglasan</location>
/
<date id="FFF2102CB44CB034330B09E7E4E1F9BD" pageId="1" pageNumber="284" value="2015-10">
<collectingDate id="EFB6E9C4B44CB034330B09E7E4E1F9BD" pageId="1" pageNumber="284" value="2015-10">X. 2015</collectingDate>
</date>
/ local collector leg.// PHILIP-
</materialsCitation>
</paragraph>
<paragraph id="8BF336ECB44CB03431480AA2E4D3F919" blockId="1.[114,154,1702,1726]" box="[114,154,1702,1726]" pageId="1" pageNumber="284">284</paragraph>
<paragraph id="8BF336ECB44CB034334A08B4E683FAC5" blockId="1.[624,1103,1200,1378]" pageId="1" pageNumber="284">
<materialsCitation id="3B243CB1B44CB034334A08B4E006FAE4" collectingDate="2022-08" collectionCode="PINES" location="Cabanglasan" municipality="Mindanao" pageId="1" pageNumber="284" specimenCount="1" stateProvince="Bukidnon" typeStatus="paratype">
<collectionCode id="ED5DAE29B44CB034334A08B4E6F3FB6F" box="[624,698,1200,1224]" pageId="1" pageNumber="284">PINES</collectionCode>
/
<collectingMunicipality id="6B97AC96B44CB03433F508B4E777FB6F" box="[719,830,1200,1224]" pageId="1" pageNumber="284">Mindanao</collectingMunicipality>
,
<collectingRegion id="4988F80EB44CB034327308B4E7FCFB6F" box="[841,949,1200,1224]" country="Philippines" name="Bukidnon" pageId="1" pageNumber="284">Bukidnon</collectingRegion>
,
<location id="8E936037B44CB03432FA08B4E006FB6F" LSID="urn:lsid:plazi:treatment:03E587FAB44CB036334A0D13E6E7FA76:8E936037B44CB03432FA08B4E006FB6F" box="[960,1103,1200,1224]" municipality="Mindanao" name="Cabanglasan" pageId="1" pageNumber="284" stateProvince="Bukidnon">Cabanglasan</location>
/
<date id="FFF2102CB44CB03433B908CBE6B1FB40" box="[643,760,1231,1255]" pageId="1" pageNumber="284" value="2022-08">
<collectingDate id="EFB6E9C4B44CB03433B908CBE6B1FB40" box="[643,760,1231,1255]" pageId="1" pageNumber="284" value="2022-08">VIII. 2022</collectingDate>
</date>
/ local collector leg. (white rectangular label, printed), with additional red label:
<typeStatus id="54F7884EB44CB03433CA0908E7CAFA83" box="[752,899,1292,1316]" pageId="1" pageNumber="284" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C4C4D6FB44CB034329C0909E697FAE4" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="tetramaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB034329C0909E697FAE4" italics="true" pageId="1" pageNumber="284">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023 (all in
</materialsCitation>
<collectionCode id="ED5DAE29B44CB034334A094EE6F6FAC5" box="[624,703,1354,1378]" pageId="1" pageNumber="284">DUBC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44CB036334A0983E78DFB7B" lastPageId="3" lastPageNumber="286" pageId="1" pageNumber="284" type="description">
<paragraph id="8BF336ECB44CB034334A0983E71EFA5C" blockId="1.[624,1103,1415,1531]" pageId="1" pageNumber="284">
<emphasis id="B938EAFEB44CB034334A0983E798FA38" bold="true" box="[624,977,1415,1439]" pageId="1" pageNumber="284">
Description.
<typeStatus id="54F7884EB44CB03432310983E73CFA38" box="[779,885,1415,1439]" pageId="1" pageNumber="284" type="holotype">Holotype</typeStatus>
, Male.
</emphasis>
(
<figureCitation id="13772A69B44CB03432D90983E00FFA38" box="[995,1094,1415,1439]" captionStart="Fig" captionStartId="2.[189,228,1386,1410]" captionTargetBox="[193,1172,171,1380]" captionTargetId="figure-114@2.[193,1172,171,1380]" captionTargetPageId="2" captionText="Fig. 2. Habitus of M. (M.) tetramaculatus sp. nov. A dorsal view, male, holotype; B dorsal view, female; C lateral view, male, holotype." pageId="1" pageNumber="284">Fig. 2A</figureCitation>
) Dimensions: LB: 6,1, LR: 1,1, WR: 0,9, LP: 2,1, WP: 1,9, LE: 4,0, WE: 2,8. N = 1 for all measurements.
</paragraph>
<paragraph id="8BF336ECB44FB03731FD0C67E0C0FFD1" blockId="2.[199,1161,99,118]" box="[199,1161,99,118]" pageId="2" pageNumber="285">
<emphasis id="B938EAFEB44FB03731FD0C67E0C0FFD1" box="[199,1161,99,118]" italics="true" pageId="2" pageNumber="285">
Four new species of the genus
<taxonomicName id="4C4C4D6FB44FB03730F50C67E6FCFFD1" authority="Heller, 1912" authorityName="Heller" authorityYear="1912" box="[463,693,99,118]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
Metapocyrtus
<bibRefCitation id="EFDD4B1DB44FB03733730C67E6FCFFD1" author="Heller K. M." box="[585,693,99,118]" pageId="2" pageNumber="285" pagination="295 - 402" refId="ref4196" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science, 7 (5): 295 - 402." type="journal article" year="1912">Heller, 1912</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C4C4D6FB44FB03733FB0C67E76DFFD1" authorityName="Linnaeus" authorityYear="1758" box="[705,804,99,118]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C4C4D6FB44FB03732140C67E7E2FFD1" box="[814,939,99,118]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
:
<taxonomicName id="4C4C4D6FB44FB037328F0C67E074FFD1" authorityName="Schoenherr" authorityYear="1826" box="[949,1085,99,118]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="tribe" tribe="Pachyrhynchini">Pachyrhynchini</taxonomicName>
) from …
</emphasis>
</paragraph>
<caption id="DF336664B44FB0373187096EE6B3FA06" pageId="2" pageNumber="285" startId="2.[189,228,1386,1410]" targetBox="[193,1172,171,1380]" targetPageId="2" targetType="figure">
<paragraph id="8BF336ECB44FB0373187096EE6B3FA06" blockId="2.[189,1122,1386,1441]" pageId="2" pageNumber="285">
Fig. 2. Habitus of
<taxonomicName id="4C4C4D6FB44FB03730BA096FE631FA24" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[384,632,1387,1411]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44FB03730BA096FE631FA24" box="[384,632,1387,1411]" italics="true" pageId="2" pageNumber="285">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44FB0373345096EE69DFA25" box="[639,724,1386,1410]" pageId="2" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
A dorsal view, male, holotype; B dorsal view, female; C lateral view, male, holotype.
</paragraph>
</caption>
<paragraph id="8BF336ECB44FB037318709C2E519F9BC" blockId="2.[189,668,1478,1563]" pageId="2" pageNumber="285">
<emphasis id="B938EAFEB44FB037318709C2E50FFA79" bold="true" box="[189,326,1478,1502]" pageId="2" pageNumber="285">Coloration.</emphasis>
Integument black, shiny, with markings of round, metallic green to turquoise scales.
</paragraph>
<paragraph id="8BF336ECB44FB03731870A45E0DDF9DF" blockId="2.[189,662,1601,1656]" lastBlockId="2.[699,1178,1478,1656]" pageId="2" pageNumber="285">
<emphasis id="B938EAFEB44FB03731870A45E54EF9FE" bold="true" box="[189,263,1601,1625]" pageId="2" pageNumber="285">Head.</emphasis>
Dorsal surface smooth, with longitudinal line of green to turquoise scales; surface between eyes with sparse white to greenish setae; genae with patch of metallic green to turquoise scales and green hair-like scales; shallow medial longitudinal groov from base of the eyes to base anterior margin of prothorax; transverse, moderate fur-
</paragraph>
<paragraph id="8BF336ECB44FB03735560AA2E0DDF919" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="285">285</paragraph>
<paragraph id="8BF336ECB44EB03631480CB6E5EEFEA0" blockId="3.[114,592,178,263]" pageId="3" pageNumber="286">row at base of eyes confluent with medial longitudinal groove; eyes large, strongly convex from outline of head.</paragraph>
<paragraph id="8BF336ECB44EB03631480D28E58DFDFF" blockId="3.[114,593,300,600]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB03631480D28E4ADFEE3" bold="true" box="[114,228,300,324]" pageId="3" pageNumber="286">Rostrum.</emphasis>
(
<figureCitation id="13772A69B44EB03631C10D28E512FEE3" box="[251,347,300,324]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A M. (M.) tetramaculatus sp. nov.; B M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C M. (M.) fulvus sp. nov.; D M. (T.) unalineatus sp. nov.; E M. (T.) pseudolanaoensis sp. nov.; F M. (M.) lanaoensis Bollino, 2021." pageId="3" pageNumber="286">Fig. 1A</figureCitation>
) longer than wide (LR/WR: 1,22), smooth at posterior half, slightly punctured at anterior half; green to white setae from apical ½ to base; dorsum with moderate medial longitudinal furrow and corresponding shallow concavity; in dorsal contour straight; in lateral contourraised dorsally at base, then gradually declined to apex; lateral parts covered with long white to shiny green setae.
</paragraph>
<paragraph id="8BF336ECB44EB03631480E79E514FC2C" blockId="3.[114,592,637,907]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB03631480E79E4AFFD32" bold="true" box="[114,230,637,661]" pageId="3" pageNumber="286">Antennae</emphasis>
with antennal scrobe not reaching margin of eye, curved downwards; antennal scape reaching anterior margin of prothorax; funicular segment I nearly three times as long as wide, slightly longer than II, segment II nearly two times longer than III, segments III VI as long as wide, segment VII larger, as longas wide, club 2,5 times as long as wide.
</paragraph>
<paragraph id="8BF336ECB44EB03631480FB4E4BFFA14" blockId="3.[114,593,944,1459]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB03631480FB4E4A7FC6F" bold="true" box="[114,238,944,968]" pageId="3" pageNumber="286">Prothorax</emphasis>
sub cylindrical, slightly longer than wide (LP/WP: 1,1), strongly punctured, with corresponding, short, white setae; widest just before the middle; in dorsal contour narrowed along anterior margin, widened to just before the middle, gradually narrowed to before posterior margin, straight to posterior margin; with the following markings of round, metallic green to turquoise shales: 1) two spots along anterior margin on disc, each redirected laterally; 2) two spots at basal part, from middle to before posterior margin on disc, each redirected laterally; anterior groove confluent at disc, slightly redirected to middle; posterior groove confluent at disc, curved upwards at lateral parts.
</paragraph>
<paragraph id="8BF336ECB44EB036314809DCE619F9EB" blockId="3.[114,593,1496,1612]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036314809DCE488FA57" bold="true" box="[114,193,1496,1520]" pageId="3" pageNumber="286">Elytra</emphasis>
sub-ellipsoidal, (LE/WE: 1,42), wider than prothorax nearly twice as long as prothorax (WE/WP: 1,47; LE/LP: 1,9); punctured in all length, without expressed
</paragraph>
<paragraph id="8BF336ECB44EB03631480AA2E7D5FE07" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1103,178,416]" pageId="3" pageNumber="286">286 puncture rows; in dorsal contour widest just before the middle, moderately convex; each elytron with the following scally markings of round, metallic green to turquoise scales: 1) two sub basal spots; 2) three spots at the medial part; 3) three spots along apical 1/3; 4) longitudinal apical stripe; goldish, thick setae from apical ½ to apex.</paragraph>
<paragraph id="8BF336ECB44EB036334A0DC1E6C1FDBC" blockId="3.[624,1103,453,539]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0DC1E6A0FE7A" bold="true" box="[624,745,453,477]" pageId="3" pageNumber="286">Fore tibia</emphasis>
without denticles along internal margin. Male genitalia as shown in
<figureCitation id="13772A69B44EB03632CD0DE0E6CDFDBC" captionStart="Fig" captionStartId="5.[114,153,1533,1557]" captionTargetBox="[120,1022,855,1503]" captionTargetId="figure-308@5.[114,1032,846,1526]" captionTargetPageId="5" captionText="Fig. 4. Male genitalia of selected species, A, C, E aedegal body in lateral view; B, D, F sternite IX; A, B M. (M.) tetramaculatus sp. nov., C, D M. (M.) fulvus sp. nov., E, F M. (T.) unalineatus sp. nov." pageId="3" pageNumber="286">Fig.4A, B</figureCitation>
.
</paragraph>
<paragraph id="8BF336ECB44EB036334A0E44E68EFD13" blockId="3.[624,1103,576,692]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0E44E699FDFF" bold="true" box="[624,720,576,600]" pageId="3" pageNumber="286">Female.</emphasis>
Elytra wider with strongly expressed apex, legs not as thick as in males, otherwise as in males; dorsal habitus as in
<figureCitation id="13772A69B44EB036334A0E98E68AFD13" box="[624,707,668,692]" captionStart="Fig" captionStartId="2.[189,228,1386,1410]" captionTargetBox="[193,1172,171,1380]" captionTargetId="figure-114@2.[193,1172,171,1380]" captionTargetPageId="2" captionText="Fig. 2. Habitus of M. (M.) tetramaculatus sp. nov. A dorsal view, male, holotype; B dorsal view, female; C lateral view, male, holotype." pageId="3" pageNumber="286">Fig. 2B</figureCitation>
.
</paragraph>
<paragraph id="8BF336ECB44EB036334A0EDDE78DFB7B" blockId="3.[624,1103,729,1245]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0EDDE723FD56" bold="true" box="[624,874,729,753]" pageId="3" pageNumber="286">Differential analyses.</emphasis>
The new species by black body colour, small size and presence of four scally spots on disc of prothorax, two at basal part and two along apical margin, is similar to
<taxonomicName id="4C4C4D6FB44EB03632050F51E7CDFC2C" authority="Cabras &amp; Medina, 2021" authorityName="Cabras &amp; Medina" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="flavomaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB03632050F51E001FCCA" box="[831,1096,853,877]" italics="true" pageId="3" pageNumber="286">M. (M.) flavomaculatus</emphasis>
Cabras &amp; Medina, 2021
</taxonomicName>
, the new species can be easily distinguished by the following characters: 1) large, strongly prominent eyes in
<taxonomicName id="4C4C4D6FB44EB03633E90FCBE794FC40" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[723,989,975,999]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB03633E90FCBE794FC40" box="[723,989,975,999]" italics="true" pageId="3" pageNumber="286">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44EB03632D60FCBE000FC40" box="[1004,1097,975,999]" pageId="3" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
instead of relatively small, weakly prominent eyes in
<taxonomicName id="4C4C4D6FB44EB03633C30809E696FBE4" authority="Cabras &amp; Medina" authorityName="Cabras &amp; Medina" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="flavomaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB03633C30809E7BEFB82" box="[761,1015,1037,1061]" italics="true" pageId="3" pageNumber="286">M. (M.) flavomaculatus</emphasis>
Cabras &amp; Medina
</taxonomicName>
(
<figureCitation id="13772A69B44EB03633CB082FE70FFBE4" box="[753,838,1067,1091]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A M. (M.) tetramaculatus sp. nov.; B M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C M. (M.) fulvus sp. nov.; D M. (T.) unalineatus sp. nov.; E M. (T.) pseudolanaoensis sp. nov.; F M. (M.) lanaoensis Bollino, 2021." pageId="3" pageNumber="286">Fig. 1B</figureCitation>
); 2) scally markings of green to turquoise colour in
<taxonomicName id="4C4C4D6FB44EB0363297084EE697FB27" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB0363297084EE697FB27" italics="true" pageId="3" pageNumber="286">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44EB03633DD086CE777FB27" box="[743,830,1128,1152]" pageId="3" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
instead of yellow ochre spots in
<taxonomicName id="4C4C4D6FB44EB03633EA0883E741FB1A" authority="Cabras &amp; Medina, 2021" authorityName="Cabras &amp; Medina" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="flavomaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB03633EA0883E79DFB38" box="[720,980,1159,1183]" italics="true" pageId="3" pageNumber="286">M. (M.) flavomaculatus</emphasis>
Cabras &amp; Medina, 2021
</taxonomicName>
(
<figureCitation id="13772A69B44EB036322108A1E728FB1A" box="[795,865,1189,1213]" captionStart="Fig" captionStartId="4.[189,228,1083,1107]" captionTargetBox="[189,1091,171,1077]" captionTargetId="figure-205@4.[189,1091,171,1077]" captionTargetPageId="4" captionText="Fig. 3. Habitus of M. (M.) flavomaculatus Cabras &amp; Medina, 2021, female; A dorsal view; B lateral view." pageId="3" pageNumber="286">Fig. 3</figureCitation>
); 3) narrow body of
<taxonomicName id="4C4C4D6FB44EB036334A08C1E720FB7A" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[624,873,1221,1245]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB036334A08C1E720FB7A" box="[624,873,1221,1245]" italics="true" pageId="3" pageNumber="286">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44EB036325508C0E78DFB7B" box="[879,964,1220,1244]" pageId="3" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44EB036334A0905E058FA9F" pageId="3" pageNumber="286" type="distribution">
<paragraph id="8BF336ECB44EB036334A0905E058FA9F" blockId="3.[624,1096,1281,1336]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0905E741FABE" bold="true" box="[624,776,1281,1305]" pageId="3" pageNumber="286">Distribution.</emphasis>
North Central Mindanao Island,
<collectingRegion id="4988F80EB44EB03633910924E75DFA9F" box="[683,788,1312,1336]" country="Philippines" name="Bukidnon" pageId="3" pageNumber="286">Bukidnon</collectingRegion>
and Agusan Provinces.
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44EB036334A0959E6E7FA76" pageId="3" pageNumber="286" type="etymology">
<paragraph id="8BF336ECB44EB036334A0959E6E7FA76" blockId="3.[624,1103,1373,1489]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0959E6B1FAD2" bold="true" box="[624,760,1373,1397]" pageId="3" pageNumber="286">Etymology.</emphasis>
The specific epithet is a latinised designation of four scally spots on disc of prothorax. Tetra four, maculata spots.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="6FD7FAE6246EF1D39A44780DAACDCDF4">Four new species of the genus Metapocyrtus Heller, 1912 (Coleoptera: Curculionidae: Pachyrhynchini) from Mindanao Island, Philippines</mods:title>
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<heading id="D0BB8180B44EB036334A09F3E006F9A8" bold="true" box="[624,1103,1527,1551]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="286" reason="4">
<emphasis id="B938EAFEB44EB036334A09F3E006F9A8" bold="true" box="[624,1103,1527,1551]" pageId="3" pageNumber="286">
<taxonomicName id="4C4C4D6FB44EB036334A09F3E05FF9A8" ID-CoL="C3BVJ" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[624,1046,1527,1551]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="fulvus" status="sp." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB036334A09F3E05FF9A8" bold="true" box="[624,1046,1527,1551]" italics="true" pageId="3" pageNumber="286">Metapocyrtus (Metapocyrtus) fulvus</emphasis>
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</emphasis>
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<emphasis id="B938EAFEB44EB036334A0A11E6E9F98A" bold="true" box="[624,672,1557,1581]" pageId="3" pageNumber="286">nov.</emphasis>
</paragraph>
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<subSubSection id="C3566567B44EB031334A0A30E0C0FFD1" lastPageId="4" lastPageNumber="287" pageId="3" pageNumber="286" type="description">
<paragraph id="8BF336ECB44EB036334A0A30E77CF9EB" blockId="3.[624,1103,1527,1612]" box="[624,821,1588,1612]" pageId="3" pageNumber="286">
(
<figureCitation id="13772A69B44EB03633420A30E683F9EB" box="[632,714,1588,1612]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A M. (M.) tetramaculatus sp. nov.; B M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C M. (M.) fulvus sp. nov.; D M. (T.) unalineatus sp. nov.; E M. (T.) pseudolanaoensis sp. nov.; F M. (M.) lanaoensis Bollino, 2021." pageId="3" pageNumber="286">Fig. 1C</figureCitation>
,
<figureCitation id="13772A69B44EB03633EC0A30E6AAF9EB" box="[726,739,1588,1612]" captionStart="Fig" captionStartId="5.[114,153,1533,1557]" captionTargetBox="[120,1022,855,1503]" captionTargetId="figure-308@5.[114,1032,846,1526]" captionTargetPageId="5" captionText="Fig. 4. Male genitalia of selected species, A, C, E aedegal body in lateral view; B, D, F sternite IX; A, B M. (M.) tetramaculatus sp. nov., C, D M. (M.) fulvus sp. nov., E, F M. (T.) unalineatus sp. nov." pageId="3" pageNumber="286">4</figureCitation>
C-D, 5)
</paragraph>
<paragraph id="8BF336ECB449B03131FD0C67E0C0FFD1" blockId="4.[199,1161,99,118]" box="[199,1161,99,118]" pageId="4" pageNumber="287">
<emphasis id="B938EAFEB449B03131FD0C67E0C0FFD1" box="[199,1161,99,118]" italics="true" pageId="4" pageNumber="287">
Four new species of the genus
<taxonomicName id="4C4C4D6FB449B03130F50C67E6FCFFD1" authority="Heller, 1912" authorityName="Heller" authorityYear="1912" box="[463,693,99,118]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
Metapocyrtus
<bibRefCitation id="EFDD4B1DB449B03133730C67E6FCFFD1" author="Heller K. M." box="[585,693,99,118]" pageId="4" pageNumber="287" pagination="295 - 402" refId="ref4196" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science, 7 (5): 295 - 402." type="journal article" year="1912">Heller, 1912</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C4C4D6FB449B03133FB0C67E76DFFD1" authorityName="Linnaeus" authorityYear="1758" box="[705,804,99,118]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="287" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C4C4D6FB449B03132140C67E7E2FFD1" box="[814,939,99,118]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
:
<taxonomicName id="4C4C4D6FB449B031328F0C67E074FFD1" authorityName="Schoenherr" authorityYear="1826" box="[949,1085,99,118]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="tribe" tribe="Pachyrhynchini">Pachyrhynchini</taxonomicName>
) from …
</emphasis>
</paragraph>
</subSubSection>
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<paragraph id="8BF336ECB449B0313187083FE5F9FBD5" blockId="4.[189,1109,1083,1138]" pageId="4" pageNumber="287">
Fig. 3. Habitus of
<taxonomicName id="4C4C4D6FB449B03130BA0838E7CDFBF4" authority="Cabras &amp; Medina, 2021" authorityName="Cabras &amp; Medina" authorityYear="2021" box="[384,900,1083,1108]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="flavomaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB449B03130BA0838E632FBF3" box="[384,635,1084,1108]" italics="true" pageId="4" pageNumber="287">M. (M.) flavomaculatus</emphasis>
Cabras &amp; Medina, 2021
</taxonomicName>
, female; A dorsal view; B lateral view.
</paragraph>
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<subSubSection id="C3566567B449B0313187089CE633F9A6" pageId="4" pageNumber="287" type="materials_examined">
<paragraph id="8BF336ECB449B0313187089CE590FAEE" blockId="4.[189,668,1176,1353]" pageId="4" pageNumber="287">
<emphasis id="B938EAFEB449B0313187089CE522FB17" bold="true" box="[189,363,1176,1200]" pageId="4" pageNumber="287">Type material.</emphasis>
<materialsCitation id="3B243CB1B449B0313042089CE5ECFAAC" collectingDate="2016-01" country="Philippines" location="Intavas" municipality="Mindanao" pageId="4" pageNumber="287" specimenCount="1" stateProvince="Bukidnon" typeStatus="holotype">
<emphasis id="B938EAFEB449B0313042089CE67AFB17" bold="true" box="[376,563,1176,1200]" pageId="4" pageNumber="287">
<typeStatus id="54F7884EB449B0313042089CE5A8FB17" box="[376,481,1176,1200]" pageId="4" pageNumber="287" type="holotype">Holotype</typeStatus>
, male:
</emphasis>
<collectingCountry id="F35B767CB449B0313372089CE556FB69" name="Philippines" pageId="4" pageNumber="287">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="6B97AC96B449B031300E08B2E5EAFB69" box="[308,419,1206,1230]" pageId="4" pageNumber="287">Mindanao</collectingMunicipality>
,
<collectingRegion id="4988F80EB449B031309708B2E650FB69" box="[429,537,1206,1230]" country="Philippines" name="Bukidnon" pageId="4" pageNumber="287">Bukidnon</collectingRegion>
,
<location id="8E936037B449B031331E08B2E638FB69" LSID="urn:lsid:plazi:treatment:03E587FAB44EB032334A09F3E692FBA2:8E936037B449B031331E08B2E638FB69" box="[548,625,1206,1230]" country="Philippines" municipality="Mindanao" name="Intavas" pageId="4" pageNumber="287" stateProvince="Bukidnon">Intavas</location>
/
<date id="FFF2102CB449B03133BC08B2E4BAFB4A" pageId="4" pageNumber="287" value="2016-01">
<collectingDate id="EFB6E9C4B449B03133BC08B2E4BAFB4A" pageId="4" pageNumber="287" value="2016-01">I. 2016</collectingDate>
</date>
/ local collector leg. (white rectangular label, printed) //
</materialsCitation>
<typeStatus id="54F7884EB449B031308608F7E61BFAAC" box="[444,594,1267,1291]" pageId="4" pageNumber="287" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C4C4D6FB449B031334308F0E57FFA8C" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="fulvus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB449B031334308F0E57FFA8C" italics="true" pageId="4" pageNumber="287">M. (M.) fulvus</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023 (red rectangular label, printed).
</paragraph>
<paragraph id="8BF336ECB449B0313187096AE633F9A6" blockId="4.[189,668,1390,1537]" pageId="4" pageNumber="287">
<materialsCitation id="3B243CB1B449B0313187096AE56AFA45" collectingDate="2016-04" country="Philippines" location="Cabanglasan" municipality="Mindanao" pageId="4" pageNumber="287" specimenCount="1" specimenCount-female="1" stateProvince="Bukidnon" typeStatus="paratype">
<emphasis id="B938EAFEB449B0313187096AE588FA21" bold="true" box="[189,449,1390,1414]" pageId="4" pageNumber="287">
<typeStatus id="54F7884EB449B0313187096AE57AFA21" box="[189,307,1390,1414]" pageId="4" pageNumber="287" type="paratype">Paratypes</typeStatus>
: (
<specimenCount id="9D4AFD65B449B031306A096AE5F3FA21" box="[336,442,1390,1414]" count="1" pageId="4" pageNumber="287" type="female">1 female</specimenCount>
)
</emphasis>
<collectingCountry id="F35B767CB449B03130E2096AE636FA21" box="[472,639,1390,1414]" name="Philippines" pageId="4" pageNumber="287">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="6B97AC96B449B03131870989E565FA02" box="[189,300,1421,1445]" pageId="4" pageNumber="287">Mindanao</collectingMunicipality>
,
<collectingRegion id="4988F80EB449B031307B0989E5E4FA02" box="[321,429,1421,1445]" country="Philippines" name="Bukidnon" pageId="4" pageNumber="287">Bukidnon</collectingRegion>
,
<location id="8E936037B449B03130F80989E603FA02" LSID="urn:lsid:plazi:treatment:03E587FAB44EB032334A09F3E692FBA2:8E936037B449B03130F80989E603FA02" box="[450,586,1421,1445]" country="Philippines" municipality="Mindanao" name="Cabanglasan" pageId="4" pageNumber="287" stateProvince="Bukidnon">Cabanglasan</location>
/
<date id="FFF2102CB449B03133490989E4BBFA64" pageId="4" pageNumber="287" value="2016-04">
<collectingDate id="EFB6E9C4B449B03133490989E4BBFA64" pageId="4" pageNumber="287" value="2016-04">IV. 2016</collectingDate>
</date>
/ local collector leg., with additional red label
</materialsCitation>
:
<materialsCitation id="3B243CB1B449B031300009CEE63FF9A6" collectionCode="DUBC" country="Philippines" pageId="4" pageNumber="287" specimenCount="1" stateProvince="Bukidnon" typeStatus="paratype">
<typeStatus id="54F7884EB449B031300009CEE584FA45" box="[314,461,1482,1506]" pageId="4" pageNumber="287" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C4C4D6FB449B03130D609CFE6C7FA44" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[492,654,1483,1507]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="fulvus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB449B03130D609CFE6C7FA44" box="[492,654,1483,1507]" italics="true" pageId="4" pageNumber="287">M. (M.) fulvus</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023 (all in
<collectionCode id="ED5DAE29B449B031331A09EDE626F9A6" box="[544,623,1513,1537]" pageId="4" pageNumber="287">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3566567B449B03331870A22E5D9FA2D" lastPageId="6" lastPageNumber="289" pageId="4" pageNumber="287" type="description">
<paragraph id="8BF336ECB449B03131870A22E6D5F9FA" blockId="4.[189,668,1574,1629]" pageId="4" pageNumber="287">
<emphasis id="B938EAFEB449B03131870A22E657F999" bold="true" box="[189,542,1574,1598]" pageId="4" pageNumber="287">
Description.
<typeStatus id="54F7884EB449B03130620A22E58BF999" box="[344,450,1574,1598]" pageId="4" pageNumber="287" type="holotype">Holotype</typeStatus>
, Male.
</emphasis>
(
<figureCitation id="13772A69B449B031330A0A22E6DAF999" box="[560,659,1574,1598]" captionStart="Fig" captionStartId="6.[189,228,1240,1264]" captionTargetBox="[189,1069,171,1234]" captionTargetId="figure-157@6.[189,1069,171,1234]" captionTargetPageId="6" captionText="Fig. 5. Habitus of M. (M.) fulvu ssp. nov. A dorsal view, male, holotype; B dorsal view, female; C lateral view, male, holotype." pageId="4" pageNumber="287">Fig. 5A</figureCitation>
) Dimensions: LB: 8,6, LR: 1,6, WR: 1,2,
</paragraph>
<paragraph id="8BF336ECB449B0313381089CE7EAFB69" blockId="4.[699,1178,1176,1230]" pageId="4" pageNumber="287">LP: 2,5, WP: 2,8, LE: 5,3, WE: 3,4. N = 1 for all measurements.</paragraph>
<paragraph id="8BF336ECB449B031338108F7E017FAEE" blockId="4.[699,1178,1267,1353]" pageId="4" pageNumber="287">
<emphasis id="B938EAFEB449B031338108F7E70DFAAC" bold="true" box="[699,836,1267,1291]" pageId="4" pageNumber="287">Coloration.</emphasis>
Prothorax, elytra, head, femur, and tarsus reddish, brown to black; body surface shiny, without scally markings.
</paragraph>
<paragraph id="8BF336ECB449B0313381096AE001F9DC" blockId="4.[699,1178,1390,1659]" pageId="4" pageNumber="287">
<emphasis id="B938EAFEB449B0313381096AE74CFA21" bold="true" box="[699,773,1390,1414]" pageId="4" pageNumber="287">Head.</emphasis>
Dorsal surface moderately punctured; surface between eyes and genae with sparse white setae; forehead flattish; shallow medial longitudinal furrow from base of the eyes to before anterior margin of prothorax; transverse, shallow furrow at base of the eyes confluent with medial longitudinal furrow; eyes medium-sized, slightly convex from outline of head.
</paragraph>
<paragraph id="8BF336ECB449B03035560AA2E561FE18" blockId="4.[1132,1172,1702,1726]" lastBlockId="5.[114,593,177,447]" lastPageId="5" lastPageNumber="288" pageId="4" pageNumber="287">
287
<emphasis id="B938EAFEB448B03031480CB5E4ADFF6E" bold="true" box="[114,228,177,201]" pageId="5" pageNumber="288">Rostrum.</emphasis>
(
<figureCitation id="13772A69B448B03031C10CB6E513FF6D" box="[251,346,178,202]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A M. (M.) tetramaculatus sp. nov.; B M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C M. (M.) fulvus sp. nov.; D M. (T.) unalineatus sp. nov.; E M. (T.) pseudolanaoensis sp. nov.; F M. (M.) lanaoensis Bollino, 2021." pageId="5" pageNumber="288">Fig. 1C</figureCitation>
) longer than wide (LR/WR: 1,1,33), punctured in all length; white setae in all length; dorsum with moderate medial longitudinal furrow from middle of apical part to base, with shallow concavity; in dorsal contour nearly straight, slightly widened along basal ½; in lateral contour straight from base, decreased from apical ½ to apex.
</paragraph>
<paragraph id="8BF336ECB448B03031480DE0E537FD13" blockId="5.[114,593,484,692]" pageId="5" pageNumber="288">
<emphasis id="B938EAFEB448B03031480DE0E4AFFE5B" bold="true" box="[114,230,484,508]" pageId="5" pageNumber="288">Antennae</emphasis>
with antennal scape slightly longer than funicle, scape reaching beyond hind margin of eye; rare white setae at anterior half; funicular segment 1,5 times longer than II, segment II 1,5 times longer than III, segments III V as long as wide, segments VI - VII wider than long.
</paragraph>
<paragraph id="8BF336ECB448B03031480EDDE7AAFE25" blockId="5.[114,593,729,845]" lastBlockId="5.[624,1103,178,386]" pageId="5" pageNumber="288">
<emphasis id="B938EAFEB448B03031480EDDE4A7FD56" bold="true" box="[114,238,729,753]" pageId="5" pageNumber="288">Prothorax</emphasis>
sub globular, slightly wider than long (WP/LP: 1,12), strongly rugose, covered with white, short setae in all length and longer setae along anterior margin; widest just at the middle, in dorsal contour narrowed along anterior margin, gradually widened middle, gradually narrowed to before posterior margin, straight to posteri- or margin; with pronounced, continuous anterior and posterior groove; with shallow medial longitudinal groove at disc.
</paragraph>
<paragraph id="8BF336ECB448B030334A0DA3E7F9FD32" blockId="5.[624,1103,423,661]" pageId="5" pageNumber="288">
<emphasis id="B938EAFEB448B030334A0DA3E6F6FE18" bold="true" box="[624,703,423,447]" pageId="5" pageNumber="288">Elytra</emphasis>
sub-ovate, (LE/WE: 1,55), wider than prothorax and more than twice as long as prothorax (WE/WP: 1,21; LE/LP: 2,12); striate punctate in all length; puncture rows expressed near suture and more chaotic to lateral margin; in dorsal contour widest just at the middle, weakly convex; short white setae corresponding puncture.
</paragraph>
<paragraph id="8BF336ECB448B030334A0EBFE706FCEA" blockId="5.[624,1103,699,845]" pageId="5" pageNumber="288">
<emphasis id="B938EAFEB448B030334A0EBFE699FD74" bold="true" box="[624,720,699,723]" pageId="5" pageNumber="288">Femora</emphasis>
and tibia covered with sparse white setae from middle towards anterior margin; fore-tibia with denticles along internal margin, mucronate. Male genitalia as shown in
<figureCitation id="13772A69B448B03033E20F31E702FCEA" box="[728,843,821,845]" captionStart="Fig" captionStartId="5.[114,153,1533,1557]" captionTargetBox="[120,1022,855,1503]" captionTargetId="figure-308@5.[114,1032,846,1526]" captionTargetPageId="5" captionText="Fig. 4. Male genitalia of selected species, A, C, E aedegal body in lateral view; B, D, F sternite IX; A, B M. (M.) tetramaculatus sp. nov., C, D M. (M.) fulvus sp. nov., E, F M. (T.) unalineatus sp. nov." pageId="5" pageNumber="288">Fig. 4C, D</figureCitation>
.
</paragraph>
<caption id="DF336664B448B030314809F9E5D2F9F5" pageId="5" pageNumber="288" startId="5.[114,153,1533,1557]" targetBox="[120,1022,855,1503]" targetPageId="5" targetType="figure">
<paragraph id="8BF336ECB448B030314809F9E5D2F9F5" blockId="5.[114,1081,1533,1618]" pageId="5" pageNumber="288">
Fig. 4. Male genitalia of selected species, A, C, E aedegal body in lateral view; B, D, F sternite IX; A, B
<taxonomicName id="4C4C4D6FB448B03030040A18E67EF993" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[318,567,1564,1588]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB448B03030040A18E67EF993" box="[318,567,1564,1588]" italics="true" pageId="5" pageNumber="288">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B448B03033040A1FE6DDF994" box="[574,660,1563,1587]" pageId="5" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
, C, D
<taxonomicName id="4C4C4D6FB448B03033D60A18E7CCF993" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[748,901,1564,1588]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="fulvus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB448B03033D60A18E7CCF993" box="[748,901,1564,1588]" italics="true" pageId="5" pageNumber="288">M. (M.) fulvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B448B03032B10A1FE7ABF994" box="[907,994,1563,1587]" pageId="5" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
, E, F
<taxonomicName id="4C4C4D6FB448B03031480A3EE576F9F5" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[114,319,1594,1618]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="unalineatus" status="sp. nov." subGenus="Trachycyrtus">
<emphasis id="B938EAFEB448B03031480A3EE576F9F5" box="[114,319,1594,1618]" italics="true" pageId="5" pageNumber="288">M. (T.) unalineatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B448B030307C0A3EE5D2F9F5" box="[326,411,1594,1618]" pageId="5" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BF336ECB448B03031480AA2E4D3F919" blockId="5.[114,154,1702,1726]" box="[114,154,1702,1726]" pageId="5" pageNumber="288">288</paragraph>
<paragraph id="8BF336ECB44BB03331FD0C67E0C0FFD1" blockId="6.[199,1161,99,118]" box="[199,1161,99,118]" pageId="6" pageNumber="289">
<emphasis id="B938EAFEB44BB03331FD0C67E0C0FFD1" box="[199,1161,99,118]" italics="true" pageId="6" pageNumber="289">
Four new species of the genus
<taxonomicName id="4C4C4D6FB44BB03330F50C67E6FCFFD1" authority="Heller, 1912" authorityName="Heller" authorityYear="1912" box="[463,693,99,118]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="genus">
Metapocyrtus
<bibRefCitation id="EFDD4B1DB44BB03333730C67E6FCFFD1" author="Heller K. M." box="[585,693,99,118]" pageId="6" pageNumber="289" pagination="295 - 402" refId="ref4196" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science, 7 (5): 295 - 402." type="journal article" year="1912">Heller, 1912</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C4C4D6FB44BB03333FB0C67E76DFFD1" authorityName="Linnaeus" authorityYear="1758" box="[705,804,99,118]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="6" pageNumber="289" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C4C4D6FB44BB03332140C67E7E2FFD1" box="[814,939,99,118]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
:
<taxonomicName id="4C4C4D6FB44BB033328F0C67E074FFD1" authorityName="Schoenherr" authorityYear="1826" box="[949,1085,99,118]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="tribe" tribe="Pachyrhynchini">Pachyrhynchini</taxonomicName>
) from …
</emphasis>
</paragraph>
<caption id="DF336664B44BB033318708DCE638FAA8" pageId="6" pageNumber="289" startId="6.[189,228,1240,1264]" targetBox="[189,1069,171,1234]" targetPageId="6" targetType="figure">
<paragraph id="8BF336ECB44BB033318708DCE638FAA8" blockId="6.[189,1170,1240,1295]" pageId="6" pageNumber="289">
Fig. 5. Habitus of
<emphasis id="B938EAFEB44BB03330BA08DDE657FB56" box="[384,542,1241,1265]" italics="true" pageId="6" pageNumber="289">
<taxonomicName id="4C4C4D6FB44BB03330BA08DDE644FB56" box="[384,525,1241,1265]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="fulvu" subGenus="Metapocyrtus">M. (M.) fulvu</taxonomicName>
s
</emphasis>
sp. nov. A dorsal view, male, holotype; B dorsal view, female; C lateral view, male, holotype.
</paragraph>
</caption>
<paragraph id="8BF336ECB44BB03331870930E5D9FA2D" blockId="6.[189,668,1332,1418]" pageId="6" pageNumber="289">
<emphasis id="B938EAFEB44BB03331870930E557FAEB" bold="true" box="[189,286,1332,1356]" pageId="6" pageNumber="289">Female.</emphasis>
Dorsal habitus as shown in
<figureCitation id="13772A69B44BB03333560930E497FACC" captionStart="Fig" captionStartId="6.[189,228,1240,1264]" captionTargetBox="[189,1069,171,1234]" captionTargetId="figure-157@6.[189,1069,171,1234]" captionTargetPageId="6" captionText="Fig. 5. Habitus of M. (M.) fulvu ssp. nov. A dorsal view, male, holotype; B dorsal view, female; C lateral view, male, holotype." pageId="6" pageNumber="289">Fig. 5B</figureCitation>
, elytra wider, apex of elytral declivity with tuft of bristles.
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44BB033318709ABE778FA60" pageId="6" pageNumber="289" type="materials_examined">
<paragraph id="8BF336ECB44BB033318709ABE643F9C7" blockId="6.[189,668,1455,1632]" pageId="6" pageNumber="289">
<emphasis id="B938EAFEB44BB033318709ABE522FA60" bold="true" box="[189,363,1455,1479]" pageId="6" pageNumber="289">Type material.</emphasis>
<materialsCitation id="3B243CB1B44BB033304209ABE5C7F984" collectingDate="2016-06" country="Philippines" county="Wao" location="Mindanao" municipality="Lanao" pageId="6" pageNumber="289" specimenCount="1" typeStatus="holotype">
<emphasis id="B938EAFEB44BB033304209ABE67AFA60" bold="true" box="[376,563,1455,1479]" pageId="6" pageNumber="289">
<typeStatus id="54F7884EB44BB033304209ABE5A8FA60" box="[376,481,1455,1479]" pageId="6" pageNumber="289" type="holotype">Holotype</typeStatus>
, male:
</emphasis>
<collectingCountry id="F35B767CB44BB033337209ABE556FA41" name="Philippines" pageId="6" pageNumber="289">PHILIPPINES</collectingCountry>
/
<location id="8E936037B44BB033307909CAE5FBFA41" LSID="urn:lsid:plazi:treatment:03E587FAB44EB032334A09F3E692FBA2:8E936037B44BB033307909CAE5FBFA41" box="[323,434,1486,1510]" country="Philippines" county="Wao" municipality="Lanao" name="Mindanao" pageId="6" pageNumber="289">Mindanao</location>
,
<collectingMunicipality id="6B97AC96B44BB03330FE09CAE640FA41" box="[452,521,1486,1510]" pageId="6" pageNumber="289">Lanao</collectingMunicipality>
,
<collectingCounty id="62924E60B44BB033332609CAE607FA41" box="[540,590,1486,1510]" pageId="6" pageNumber="289">Wao</collectingCounty>
/
<date id="FFF2102CB44BB033334909CAE4BAF9A3" pageId="6" pageNumber="289" value="2016-06">
<collectingDate id="EFB6E9C4B44BB033334909CAE4BAF9A3" pageId="6" pageNumber="289" value="2016-06">VI. 2016</collectingDate>
</date>
/ local collector leg. (white rectangular label, printed) //
</materialsCitation>
<typeStatus id="54F7884EB44BB03330A70A0FE67AF984" box="[413,563,1547,1571]" pageId="6" pageNumber="289" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C4C4D6FB44BB03333700A0FE509F9E5" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="unalineatus" subGenus="Trachycyrtus">
<emphasis id="B938EAFEB44BB03333700A0FE509F9E5" italics="true" pageId="6" pageNumber="289">M. (T.) unalineatus</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023 (red rectangular label, printed).
</paragraph>
<paragraph id="8BF336ECB44BB03333810930E778FA60" blockId="6.[699,1178,1332,1479]" pageId="6" pageNumber="289">
<materialsCitation id="3B243CB1B44BB03333810930E7C7FA2D" pageId="6" pageNumber="289" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B938EAFEB44BB03333810930E056FAEB" bold="true" box="[699,1055,1332,1356]" pageId="6" pageNumber="289">
<typeStatus id="54F7884EB44BB03333810930E778FAEB" box="[699,817,1332,1356]" pageId="6" pageNumber="289" type="paratype">Paratypes</typeStatus>
: (
<specimenCount id="9D4AFD65B44BB03332700930E7D3FAEB" box="[842,922,1332,1356]" count="1" pageId="6" pageNumber="289" type="male">1 male</specimenCount>
,
<specimenCount id="9D4AFD65B44BB03332900930E05AFAEB" box="[938,1043,1332,1356]" count="1" pageId="6" pageNumber="289" type="female">1 female</specimenCount>
):
</emphasis>
both with the same location label as
<typeStatus id="54F7884EB44BB03332CE0957E01DFACC" box="[1012,1108,1363,1387]" pageId="6" pageNumber="289" type="holotype">holotype</typeStatus>
, with additional red label
</materialsCitation>
:
<materialsCitation id="3B243CB1B44BB033329A0976E764FA60" collectionCode="DUBC" pageId="6" pageNumber="289" specimenCount="1" typeStatus="paratype">
<typeStatus id="54F7884EB44BB033329A0976E07AFA2D" box="[928,1075,1394,1418]" pageId="6" pageNumber="289" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C4C4D6FB44BB03335730976E777FA0E" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="species" species="unalineatus" subGenus="Trachycyrtus">
<emphasis id="B938EAFEB44BB03335730976E777FA0E" italics="true" pageId="6" pageNumber="289">M. (T.) unalineatus</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023 (all in
<collectionCode id="ED5DAE29B44BB03333ED09ABE76FFA60" box="[727,806,1455,1479]" pageId="6" pageNumber="289">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44BB032338109E8E731FCB7" lastPageId="7" lastPageNumber="290" pageId="6" pageNumber="289" type="description">
<paragraph id="8BF336ECB44BB033338109E8E7EAF9C7" blockId="6.[699,1178,1516,1632]" pageId="6" pageNumber="289">
<emphasis id="B938EAFEB44BB033338109E8E055F9A3" bold="true" box="[699,1052,1516,1540]" pageId="6" pageNumber="289">
Description.
<typeStatus id="54F7884EB44BB033326C09E8E789F9A3" box="[854,960,1516,1540]" pageId="6" pageNumber="289" type="holotype">Holotype</typeStatus>
, Male.
</emphasis>
(
<figureCitation id="13772A69B44BB033351409E8E0D8F9A3" box="[1070,1169,1516,1540]" captionStart="Fig" captionStartId="8.[189,228,1567,1591]" captionTargetBox="[194,1144,422,1537]" captionTargetId="figure-115@8.[189,1160,416,1561]" captionTargetPageId="8" captionText="Fig. 6. Habitus of M. (T.) unalineatus sp. nov. A dorsal view, male, holotype; B dorsal view, female; C lateral view, male, holotype." pageId="6" pageNumber="289">Fig. 6A</figureCitation>
) Dimensions: LB: 4,8, LR: 0,9, WR: 0,6, LP: 1,9, WP: 1,6, LE: 2,9, WE: 2,0. N = 1 for all measurements.
</paragraph>
<paragraph id="8BF336ECB44BB03335560AA2E0DDF919" blockId="6.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="6" pageNumber="289">289</paragraph>
<paragraph id="8BF336ECB44AB03231480CB5E53EFEA0" blockId="7.[114,592,177,263]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB03231480CB5E4B3FF6E" bold="true" box="[114,250,177,201]" pageId="7" pageNumber="290">Coloration.</emphasis>
Body black, strongly shiny, with markings of round, metallic pink, greenish to silver scales.
</paragraph>
<paragraph id="8BF336ECB44AB03231480D28E5C8FD9D" blockId="7.[114,593,300,570]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB03231480D28E4F5FEE3" bold="true" box="[114,188,300,324]" pageId="7" pageNumber="290">Head.</emphasis>
Dorsal surface smooth, with single colourful scales; surface between eyes and genae with rare silver setae; genae without scally markings; shallow medial longitudinal groove in all length; transverse, moderate furrow at base of eyes, not reaching lateral margins, confluent with medial longitudinal groove; eyes large, strongly convex from outline of head.
</paragraph>
<paragraph id="8BF336ECB44AB03231480E5BE664FCB7" blockId="7.[114,593,607,784]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB03231480E5BE4ADFDD0" bold="true" box="[114,228,607,631]" pageId="7" pageNumber="290">Rostrum.</emphasis>
(
<figureCitation id="13772A69B44AB03231C10E5BE515FDD0" box="[251,348,607,631]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A M. (M.) tetramaculatus sp. nov.; B M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C M. (M.) fulvus sp. nov.; D M. (T.) unalineatus sp. nov.; E M. (T.) pseudolanaoensis sp. nov.; F M. (M.) lanaoensis Bollino, 2021." pageId="7" pageNumber="290">Fig. 1D</figureCitation>
) longer than wide (LR/WR: 1,5), rugose in all length; silver setae in all length; dorsum with moderate medial longitudinal furrow and corresponding shallow concavity; in dorsal contour straight; in lateral contour nearly straight.
</paragraph>
<paragraph id="8BF336ECB44AB03231480F31E514FBA2" blockId="7.[114,593,821,1029]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB03231480F31E4AFFCEA" bold="true" box="[114,230,821,845]" pageId="7" pageNumber="290">Antennae</emphasis>
with antennal scape reaching behind margin of eye; funicular segment I nearly four times as long as wide, nearly two times longer than II, segment II more than two times longer than III, segments III VII as long as wide, club wide, two times as long as wide.
</paragraph>
<paragraph id="8BF336ECB44AB0323148082FE5E6FB5C" blockId="7.[114,592,1067,1275]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB0323148082FE4A7FBE4" bold="true" box="[114,238,1067,1091]" pageId="7" pageNumber="290">Prothorax</emphasis>
sub globular, slightly longer than wide (LP/WP: 1,18), moderately punctured with corresponding, short, silver setae; widest just at the middle; without scally markings; with strongly pronounces, continuous anterior and posterior groove; anterior margin curved anteriorly.
</paragraph>
<paragraph id="8BF336ECB44AB03231480924E4B5F9CC" blockId="7.[114,593,1312,1643]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB03231480924E488FA9F" bold="true" box="[114,193,1312,1336]" pageId="7" pageNumber="290">Elytra</emphasis>
sub-ellipsoidal, (LE/WE: 1,45), wider than prothorax and 1,5 times as long as prothorax (WE/WP: 1,25; LE/LP: 1,52); moderately punctured in all length, with expressed puncture rows and corresponding short setae; in dorsal contour widest just before the middle, slightly convex; with the following scally markings: 1) one longitudinal line of scales along suture; 2) longitudinal line of scales along lateral margin of each elytron.
</paragraph>
<paragraph id="8BF336ECB44AB03231480AA2E04BFEC4" blockId="7.[114,154,1702,1726]" lastBlockId="7.[624,1103,177,355]" pageId="7" pageNumber="290">
290
<emphasis id="B938EAFEB44AB032334A0CB5E6E5FF6E" bold="true" box="[624,684,177,201]" pageId="7" pageNumber="290">Tibia</emphasis>
strongly widened from before middle to anterior 2/3; with long, rare silver setae from basal ½ to anterior edge; reddish in colour at basal part; tibia with shorter setae, without denticles along internal margin. Male genitalia as shown in
<figureCitation id="13772A69B44AB03232AE0D4FE79BFEC4" box="[916,978,331,355]" captionStart="Fig" captionStartId="5.[114,153,1533,1557]" captionTargetBox="[120,1022,855,1503]" captionTargetId="figure-308@5.[114,1032,846,1526]" captionTargetPageId="5" captionText="Fig. 4. Male genitalia of selected species, A, C, E aedegal body in lateral view; B, D, F sternite IX; A, B M. (M.) tetramaculatus sp. nov., C, D M. (M.) fulvus sp. nov., E, F M. (T.) unalineatus sp. nov." pageId="7" pageNumber="290">Fig. 4</figureCitation>
E-F.
</paragraph>
<paragraph id="8BF336ECB44AB032334A0D8CE04BFE5B" blockId="7.[624,1103,392,508]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB032334A0D8CE699FE07" bold="true" box="[624,720,392,416]" pageId="7" pageNumber="290">Female.</emphasis>
Elytra wider, more strongly round- ed, with strongly expressed apex, with brown, thick setae along apex; otherwise as in males; dorsal habitus as in
<figureCitation id="13772A69B44AB03232910DE0E7B7FE5B" box="[939,1022,484,508]" captionStart="Fig" captionStartId="8.[189,228,1567,1591]" captionTargetBox="[194,1144,422,1537]" captionTargetId="figure-115@8.[189,1160,416,1561]" captionTargetPageId="8" captionText="Fig. 6. Habitus of M. (T.) unalineatus sp. nov. A dorsal view, male, holotype; B dorsal view, female; C lateral view, male, holotype." pageId="7" pageNumber="290">Fig. 6B</figureCitation>
.
</paragraph>
<paragraph id="8BF336ECB44AB032334A0E25E731FCB7" blockId="7.[624,1103,545,784]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB032334A0E25E722FD9E" bold="true" box="[624,875,545,569]" pageId="7" pageNumber="290">Differential analyses.</emphasis>
The new species is unique in appearance and different from all known members of the genus, by brown - reddish colour of prothorax and general body shape new species can be referred to
<taxonomicName id="4C4C4D6FB44AB032334A0EBFE05EFD74" authority="Cabras &amp; Medina, 2021" authorityName="Cabras &amp; Medina" authorityYear="2021" box="[624,1047,699,723]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="290" phylum="Arthropoda" rank="species" species="ged" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44AB032334A0EBFE749FD74" box="[624,768,699,723]" italics="true" pageId="7" pageNumber="290">M. (M.) ged</emphasis>
Cabras &amp; Medina, 2021
</taxonomicName>
, but can be easily distinguished by bare body without scally markings.
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44AB032334A0F31E69FFCCB" pageId="7" pageNumber="290" type="distribution">
<paragraph id="8BF336ECB44AB032334A0F31E69FFCCB" blockId="7.[624,1103,821,876]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB032334A0F31E741FCEA" bold="true" box="[624,776,821,845]" pageId="7" pageNumber="290">Distribution.</emphasis>
Mindanao Island,
<collectingRegion id="4988F80EB44AB03232E50F31E69BFCCB" country="Philippines" name="Bukidnon" pageId="7" pageNumber="290">Bukidnon Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44AB032334A0F95E692FBA2" pageId="7" pageNumber="290" type="etymology">
<paragraph id="8BF336ECB44AB032334A0F95E692FBA2" blockId="7.[624,1102,913,1029]" pageId="7" pageNumber="290">
<emphasis id="B938EAFEB44AB032334A0F95E6B1FC0E" bold="true" box="[624,760,913,937]" pageId="7" pageNumber="290">Etymology.</emphasis>
The specific epithet is a latinised name of specific body colouration of
<taxonomicName id="4C4C4D6FB44AB032334A0FCBE754FC40" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[624,797,975,999]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="290" phylum="Arthropoda" rank="species" species="fulvus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44AB032334A0FCBE754FC40" box="[624,797,975,999]" italics="true" pageId="7" pageNumber="290">M. (M.) fulvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44AB032321E0FCBE7C9FC40" box="[804,896,975,999]" pageId="7" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
Fulva reddish, brownish.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,238 @@
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<paragraph id="8BF036C09D5EFF9FFEACFC5AFDC15F52" blockId="46.[302,762,996,1084]" pageId="46" pageNumber="47">
<heading id="D0B881AC9D5EFF9FFEACFC5AFDC15F52" fontSize="10" level="2" pageId="46" pageNumber="47" reason="7">
<taxonomicName id="4C4F4D439D5EFF9FFEACFC5AFD4A58B2" authorityName="Muilwijk &amp; Felix &amp; Anichtchenko" authorityYear="2022" box="[302,695,996,1020]" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="kabaki" status="sp. nov." subGenus="Dichromenta">
<emphasis id="B93BEAD29D5EFF9FFEACFC5AFE4458B2" bold="true" box="[302,441,996,1020]" italics="true" pageId="46" pageNumber="47">Merizomena</emphasis>
(
<emphasis id="B93BEAD29D5EFF9FFE4BFC5AFDA658B2" bold="true" box="[457,603,996,1020]" italics="true" pageId="46" pageNumber="47">Dichromenta</emphasis>
) kabaki
</taxonomicName>
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urn:lsid:zoobank.org:act:
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<subSubSection id="C355654B9D5EFF9FFEACFB9BFE335F72" box="[302,462,1061,1084]" pageId="46" pageNumber="47" type="description">
<paragraph id="8BF036C09D5EFF9FFEACFB9BFE335F72" blockId="46.[302,762,996,1084]" box="[302,462,1061,1084]" pageId="46" pageNumber="47">
(
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)
</paragraph>
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<subSubSection id="C355654B9D5EFF9FFEACFBDAFD8F5E53" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BF036C09D5EFF9FFEACFBDAFD8F5E53" blockId="46.[302,813,1124,1309]" pageId="46" pageNumber="47">
<emphasis id="B93BEAD29D5EFF9FFEACFBDAFE245F32" bold="true" box="[302,473,1124,1148]" pageId="46" pageNumber="47">Type material.</emphasis>
<materialsCitation id="3B273C9D9D5EFF9FFE61FBDBFD8F5E53" collectionCode="R, RISNB" collectorName="I. Sc." county="Chodshent" determinerName="G. Fagel" location="Muilwijk" municipality="Solsky" pageId="46" pageNumber="47" specimenCount="2" stateProvince="Mus" typeStatus="holotype">
<typeStatus id="54F488629D5EFF9FFE61FBDBFDB75F32" box="[483,586,1125,1148]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
, labelled: “
<collectingCounty id="62914E4C9D5EFF9FFD6AFBDBFE9A5FD3" pageId="46" pageNumber="47">Chodshent</collectingCounty>
// Reg.
<collectingRegion id="498BF8229D5EFF9FFE3EFB38FE125FD3" box="[444,495,1158,1181]" country="Turkey" name="Mus" pageId="46" pageNumber="47">Mus</collectingRegion>
. Hist. Nat. / Belg. I. G. 10294 //
<determinerName id="7E2CDA049D5EFF9FFEC0FB18FE645FF3" box="[322,409,1190,1213]" pageId="46" pageNumber="47">G. Fagel</determinerName>
det. /
<taxonomicName id="4C4F4D439D5EFF9FFE5AFB18FDE25FF3" authorityName="Chaudoir" authorityYear="1842" box="[472,543,1190,1213]" class="Insecta" family="Carabidae" genus="Glycia" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Glycia</taxonomicName>
/ bicolor /
<collectingMunicipality id="6B94ACBA9D5EFF9FFD0CFB18FD2A5FF3" box="[654,727,1190,1213]" pageId="46" pageNumber="47">Solsky</collectingMunicipality>
//
<taxonomicName id="4C4F4D439D5EFF9FFD72FB18FD8D5F93" authorityName="Muilwijk &amp; Felix &amp; Anichtchenko" authorityYear="2022" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="kabaki" status="sp. nov." subGenus="Dichromenta">Merizomena (Dichromenta) kabaki</taxonomicName>
<emphasis id="B93BEAD29D5EFF9FFDF7FB7BFD4D5F93" bold="true" box="[629,688,1221,1245]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A20857A99D5EFF9FFDF7FB7BFD4D5F93" box="[629,688,1221,1245]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
<emphasis id="B93BEAD29D5EFF9FFD37FB78FD415F93" box="[693,700,1222,1245]" italics="true" pageId="46" pageNumber="47">/</emphasis>
<location id="8E90601B9D5EFF9FFD40FB78FCD55F93" LSID="urn:lsid:plazi:treatment:03E687D69D5EFF80FEACFC5AFC675BEA:8E90601B9D5EFF9FFD40FB78FCD55F93" box="[706,808,1222,1245]" county="Chodshent" municipality="Solsky" name="Muilwijk" pageId="46" pageNumber="47" stateProvince="Mus">Muilwijk</location>
, Felix &amp; Anichtchenko, 2022 //
<typeStatus id="54F488629D5EFF9FFDFBFB58FD205FB3" box="[633,733,1254,1277]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
(red) // Coll.
<collectionCode id="ED5EAE059D5EFF9FFEE8FAB8FE825E53" box="[362,383,1286,1309]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="46" pageNumber="47">R</collectionCode>
.
<collectorName id="26BA53169D5EFF9FFE08FAB8FE3F5E53" box="[394,450,1286,1309]" pageId="46" pageNumber="47">I. Sc.</collectorName>
N. B.”,
<collectionCode id="ED5EAE059D5EFF9FFD99FAB8FD935E53" box="[539,622,1286,1309]" pageId="46" pageNumber="47">RISNB</collectionCode>
.
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</paragraph>
</subSubSection>
<subSubSection id="C355654B9D5EFF81FEACFAFBFD125A16" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
<paragraph id="8BF036C09D5EFF9FFEACFAFBFCD05D73" blockId="46.[302,813,1349,1597]" pageId="46" pageNumber="47">
<emphasis id="B93BEAD29D5EFF9FFEACFAFBFE445E13" bold="true" box="[302,441,1349,1373]" pageId="46" pageNumber="47">Description.</emphasis>
A small sized species, (BL
<quantity id="4CB79B259D5EFF9FFD54FAF8FCD55E13" box="[726,808,1350,1373]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="46" pageNumber="47" unit="mm" value="6.5">6.5 mm</quantity>
, BW
<quantity id="4CB79B259D5EFF9FFEDCFAD8FE4E5E33" box="[350,435,1382,1405]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="47" unit="mm" value="2.5">2.5 mm</quantity>
). Habitus as in
<figureCitation id="13742A459D5EFF9FFDD0FAD8FD5B5E33" box="[594,678,1382,1405]" captionStart="Fig" captionStartId="47.[227,266,1033,1056]" captionTargetBox="[226,1284,189,998]" captionTargetId="figure-10@47.[226,1284,189,998]" captionTargetPageId="47" captionText="Fig. 9.1 Merizomena kabaki habitus, holotype, dorsal view, scale 5 mm." pageId="46" pageNumber="47">Fig. 9.1</figureCitation>
. Head brownish, pronotum brownish with yellowish lateral gutter, basal half of elytra and lateral elytral gutter yellowish, apex of elytra brown, with an extension anteriorly along the suture; small appendages yellowish. Ventral side of head and thorax yellowish, ventrites slightly darker. Dorsal side shiny.
</paragraph>
<paragraph id="8BF036C09D5EFF9FFEACF9DBFAE95F52" blockId="46.[302,813,1637,1757]" lastBlockId="46.[850,1361,869,1052]" pageId="46" pageNumber="47">
<emphasis id="B93BEAD29D5EFF9FFEACF9DBFE8F5D33" bold="true" box="[302,370,1637,1661]" pageId="46" pageNumber="47">Head.</emphasis>
Narrower than pronotum (HW/PM: 0.90). Eyes convex. Tempora, half the length of an eye (T/E: 0.5). Frontal furrows surpressed. Frons with a small and shallow median impression. Vertex on each side with one seta diagonally behind the eyes. Frons and vertex covered with punctures, coarser on posterior side of vertex and very sparce hairs; base of frons with some punctures. Microsculpture consisting of micropunctures on frons. Antennomere 3 as long as antennomere 1.
</paragraph>
<paragraph id="8BF036C09D5EFF9FFCD0FBFAFB305E92" blockId="46.[850,1361,1092,1500]" pageId="46" pageNumber="47">
<emphasis id="B93BEAD29D5EFF9FFCD0FBFAFC565F12" bold="true" box="[850,939,1092,1116]" pageId="46" pageNumber="47">Thorax.</emphasis>
Pronotum cordate (PM/PL: 1.18), widest at approximately the end of anterior third, lateral sides regularly narrowed posteriorly, in basal sixth straight to the nearly straight hind angles (PM/ PS: 1.26, PM/PB: 1.27). Base lobed, oblique towards the hind angles. Lateral gutter narrow and unclear separated from the flat disc. Basal foveae slightly deepened, unified with lateral gutter. Median line impressed. Surface covered with punctures, denser along the lateral sides. Microsculpture consisting of very superficial irregular meshes. Metepisterna about 3.5 times as long as wide, narrowed posteriorly.
</paragraph>
<paragraph id="8BF036C09D5EFF9FFCD0F9BAFAAD5D92" blockId="46.[850,1361,1540,1756]" pageId="46" pageNumber="47">
<emphasis id="B93BEAD29D5EFF9FFCD0F9BAFC345D52" bold="true" box="[850,969,1540,1564]" pageId="46" pageNumber="47">Abdomen.</emphasis>
Elytra subrectangular, slightly elongated, moderately flat, (EL/EW: 1.63); widest at apical quarter. Regularly rounded to the suture. Sutural angle subrectangular, without dent. Disc moderately flat. Striae embedded, punctured. Interstriae slightly convex; with one row of punctures, only in the outer striae with a few fine
</paragraph>
<paragraph id="8BF036C09D5FFF9EFDCDFF2FFC675BEA" blockId="47.[591,922,145,164]" box="[591,922,145,164]" pageId="47" pageNumber="48">
<emphasis id="B93BEAD29D5FFF9EFDCDFF2FFC675BEA" box="[591,922,145,164]" italics="true" pageId="47" pageNumber="48">Muilwijk J., Felix R., Anichtchenko A.</emphasis>
</paragraph>
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<paragraph id="8BF036C09D5FFF9EFF61FBB7FC155F6E" blockId="47.[227,1000,1033,1056]" box="[227,1000,1033,1056]" pageId="47" pageNumber="48">
Fig. 9.1
<taxonomicName id="4C4F4D439D5FFF9EFEBEFBB7FDEC5F6E" authorityName="Muilwijk &amp; Felix &amp; Anichtchenko" authorityYear="2022" box="[316,529,1033,1056]" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kabaki">
<emphasis id="B93BEAD29D5FFF9EFEBEFBB7FDEC5F6E" box="[316,529,1033,1056]" italics="true" pageId="47" pageNumber="48">Merizomena kabaki</emphasis>
</taxonomicName>
habitus, holotype, dorsal view, scale 5 mm.
</paragraph>
</caption>
<paragraph id="8BF036C09D5FFF9EFC84FBE2FB935F9B" blockId="47.[774,1285,1116,1237]" pageId="47" pageNumber="48">setae. Microsculpture absent.Ventrites with yellow, decumbent hairs, denser on lateral sides and on anal ventrite. Anal ventrite with three pairs of marginal setae, incised at middle.</paragraph>
<paragraph id="8BF036C09D5FFF9EFC84FB41FBCD5E37" blockId="47.[774,1285,1279,1401]" pageId="47" pageNumber="48">
<emphasis id="B93BEAD29D5FFF9EFC84FB41FCBE5E59" bold="true" box="[774,835,1279,1303]" pageId="47" pageNumber="48">Legs.</emphasis>
Moderately slender, with meso­ and metatibiae longer than corresponding tarsi. Ventral side of meso­ and metatarsi 14 with dense short bristles. Claws with 5 teeth.
</paragraph>
<paragraph id="8BF036C09D5FFF9EFC84FA1DFBC85D31" blockId="47.[774,1285,1443,1663]" pageId="47" pageNumber="48">
<emphasis id="B93BEAD29D5FFF9EFC84FA1DFC5F5EF5" bold="true" box="[774,930,1443,1467]" pageId="47" pageNumber="48">Median lobe.</emphasis>
Dorsal side in horizontal view curved with a more or less rectilinear part in the middle, ventral side slightly bisinuate (
<figureCitation id="13742A459D5FFF9EFB21FA5BFB045EB2" box="[1187,1273,1509,1532]" captionStart="Fig" captionStartId="47.[227,266,1033,1056]" captionTargetBox="[226,1284,189,998]" captionTargetId="figure-10@47.[226,1284,189,998]" captionTargetPageId="47" captionText="Fig. 9.1 Merizomena kabaki habitus, holotype, dorsal view, scale 5 mm." pageId="47" pageNumber="48">Fig. 9.2</figureCitation>
). Length
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. Endophallus with two copulatory pieces, one copulatory piece comma shaped and the other thin, short and straight. Right paramere with a wide basal part.
</paragraph>
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<paragraph id="8BF036C09D5FFF9EFF61F99AFD7B5D35" blockId="47.[227,737,1572,1659]" pageId="47" pageNumber="48">
Fig. 9.2
<taxonomicName id="4C4F4D439D5FFF9EFEBAF99AFDF75D75" authorityName="Muilwijk &amp; Felix &amp; Anichtchenko" authorityYear="2022" box="[312,522,1572,1595]" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kabaki">
<emphasis id="B93BEAD29D5FFF9EFEBAF99AFDF75D75" box="[312,522,1572,1595]" italics="true" pageId="47" pageNumber="48">Merizomena kabaki</emphasis>
</taxonomicName>
aedeagus, holotype, rl = right lateral view, rp = right­, lp = left paramere, scale 1 mm.jpg, scale 1 mm.
</paragraph>
</caption>
<paragraph id="8BF036C09D5FFF9EFC84F914FBBC5D8F" blockId="47.[774,1089,1706,1730]" box="[774,1089,1706,1730]" pageId="47" pageNumber="48">
<emphasis id="B93BEAD29D5FFF9EFC84F914FC375D8C" bold="true" box="[774,970,1706,1730]" pageId="47" pageNumber="48">Female genitalia.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BF036C09D40FF81FEACFF7EFD125A16" blockId="48.[302,813,192,344]" pageId="48" pageNumber="49">
<emphasis id="B93BEAD29D40FF81FEACFF7EFDFA5B96" bold="true" box="[302,519,192,216]" pageId="48" pageNumber="49">Comparative notes.</emphasis>
<taxonomicName id="4C4F4D439D40FF81FD89FF7FFD8F5B96" authorityName="Muilwijk &amp; Felix &amp; Anichtchenko" authorityYear="2022" box="[523,626,193,216]" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kabaki" status="sp. nov.">
<emphasis id="B93BEAD29D40FF81FD89FF7FFD8F5B96" box="[523,626,193,216]" italics="true" pageId="48" pageNumber="49">M. kabaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857A99D40FF81FDF5FF7FFD505B96" box="[631,685,193,216]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
differs from all other known species of this subgenus by the combination of small body size, more oblong elytra, inner elytral interstriae without hairs and two copulatory pieces in the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C355654B9D40FF81FEACFE3EFB595BB6" pageId="48" pageNumber="49" type="etymology">
<paragraph id="8BF036C09D40FF81FEACFE3EFCD05AF6" blockId="48.[302,813,384,440]" pageId="48" pageNumber="49">
<emphasis id="B93BEAD29D40FF81FEACFE3EFE4D5AD6" bold="true" box="[302,432,384,408]" pageId="48" pageNumber="49">Etymology.</emphasis>
This new species is named after our friend and colleague, the entomologist Ilya I.
</paragraph>
<paragraph id="8BF036C09D40FF81FCD0FF7FFB595BB6" blockId="48.[850,1360,193,248]" pageId="48" pageNumber="49">
Kabak (
<collectingRegion id="498BF8229D40FF81FC2BFF7FFBBB5B96" box="[937,1094,193,216]" country="Russia" name="Saint Petersburg" pageId="48" pageNumber="49">St. Petersburg</collectingRegion>
), who made an essential contribution to this manuscript.
</paragraph>
</subSubSection>
<subSubSection id="C355654B9D40FF81FCD0FE9EFADF5A36" pageId="48" pageNumber="49" type="discussion">
<paragraph id="8BF036C09D40FF81FCD0FE9EFADF5A36" blockId="48.[850,1360,288,376]" pageId="48" pageNumber="49">
<emphasis id="B93BEAD29D40FF81FCD0FE9EFC285A76" bold="true" box="[850,981,288,312]" pageId="48" pageNumber="49">Comments.</emphasis>
This species was erroneously identified as
<taxonomicName id="4C4F4D439D40FF81FC2AFEFFFB5E5A16" authority="Solsky. It" authorityName="Solsky. It" box="[936,1187,321,344]" class="Insecta" family="Carabidae" genus="Glycia" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B93BEAD29D40FF81FC2AFEFFFBC35A16" box="[936,1086,321,344]" italics="true" pageId="48" pageNumber="49">Glycia bicolor</emphasis>
Solsky. It
</taxonomicName>
is unknown why the label “
<typeStatus id="54F488629D40FF81FC3CFEDFFBDA5A36" box="[958,1063,353,376]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
” was added. (
<figureCitation id="13742A459D40FF81FB3CFEDFFAEB5A36" box="[1214,1302,353,376]" captionStart="Fig" captionStartId="47.[227,266,1033,1056]" captionTargetBox="[226,1284,189,998]" captionTargetId="figure-10@47.[226,1284,189,998]" captionTargetPageId="47" captionText="Fig. 9.1 Merizomena kabaki habitus, holotype, dorsal view, scale 5 mm." pageId="48" pageNumber="49">Fig. 9.3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C355654B9D40FF80FCD0FE1EFC675BEA" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="distribution">
<paragraph id="8BF036C09D40FF81FCD0FE1EFBA65A96" blockId="48.[850,1361,416,472]" pageId="48" pageNumber="49">
<emphasis id="B93BEAD29D40FF81FCD0FE1EFC1F5AF6" bold="true" box="[850,994,416,440]" pageId="48" pageNumber="49">Distribution.</emphasis>
Only known from the
<typeStatus id="54F488629D40FF81FB48FE1FFB0A5AF6" box="[1226,1271,417,440]" pageId="48" pageNumber="49">type</typeStatus>
location in
<collectingCountry id="F35876509D40FF81FCEFFE7FFC0B5A96" box="[877,1014,449,472]" name="Tajikistan" pageId="48" pageNumber="49">Tadzhikistan</collectingCountry>
(
<figureCitation id="13742A459D40FF81FB87FE7FFBB35A96" box="[1029,1102,449,472]" captionStart="MAP 9" captionStartId="48.[302,360,1725,1748]" captionTargetBox="[310,1360,1125,1689]" captionTargetId="figure-18@48.[302,1360,1124,1692]" captionTargetPageId="48" captionText="MAP 9. Merizomena (Dichromenta) kabaki sp.n." pageId="48" pageNumber="49">Map 9</figureCitation>
).
</paragraph>
<caption id="DF3066489D40FF81FEACFBA7FCF85F7E" box="[302,773,1049,1072]" pageId="48" pageNumber="49" startId="48.[302,341,1049,1072]" targetBox="[302,1360,484,1014]" targetPageId="48" targetType="figure">
<paragraph id="8BF036C09D40FF81FEACFBA7FCF85F7E" blockId="48.[302,773,1049,1072]" box="[302,773,1049,1072]" pageId="48" pageNumber="49">
Fig. 9.3
<taxonomicName id="4C4F4D439D40FF81FE0AFBA7FDA05F7E" authorityName="Muilwijk &amp; Felix &amp; Anichtchenko" authorityYear="2022" box="[392,605,1049,1072]" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kabaki">
<emphasis id="B93BEAD29D40FF81FE0AFBA7FDA05F7E" box="[392,605,1049,1072]" italics="true" pageId="48" pageNumber="49">Merizomena kabaki</emphasis>
</taxonomicName>
label holotype.
</paragraph>
</caption>
<caption id="DF3066489D40FF81FEACF903FCBC5D9B" box="[302,833,1725,1749]" pageId="48" pageNumber="49" startId="48.[302,360,1725,1748]" targetBox="[310,1360,1125,1689]" targetPageId="48" targetType="figure">
<paragraph id="8BF036C09D40FF81FEACF903FCBC5D9B" blockId="48.[302,833,1725,1749]" box="[302,833,1725,1749]" pageId="48" pageNumber="49">
MAP 9.
<taxonomicName id="4C4F4D439D40FF81FE0BF900FCF85D9B" authorityName="Muilwijk &amp; Felix &amp; Anichtchenko" authorityYear="2022" box="[393,773,1725,1749]" class="Insecta" family="Carabidae" genus="Merizomena" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kabaki" status="sp. nov." subGenus="Dichromenta">
<emphasis id="B93BEAD29D40FF81FE0BF900FDED5D9B" box="[393,528,1726,1749]" italics="true" pageId="48" pageNumber="49">Merizomena</emphasis>
(
<emphasis id="B93BEAD29D40FF81FDA2F900FD505D9B" box="[544,685,1726,1749]" italics="true" pageId="48" pageNumber="49">Dichromenta</emphasis>
)
<emphasis id="B93BEAD29D40FF81FD3FF900FCF85D9B" box="[701,773,1726,1749]" italics="true" pageId="48" pageNumber="49">kabaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857A99D40FF81FC8EF903FCBC5D9B" box="[780,833,1725,1749]" pageId="48" pageNumber="49" rank="species">sp.n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BF036C09D41FF80FDCDFF2FFC675BEA" blockId="49.[591,922,145,164]" box="[591,922,145,164]" pageId="49" pageNumber="50">
<emphasis id="B93BEAD29D41FF80FDCDFF2FFC675BEA" box="[591,922,145,164]" italics="true" pageId="49" pageNumber="50">Muilwijk J., Felix R., Anichtchenko A.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,69 @@
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<emphasis id="B93BEAD29D79FFB8FF61FCDBFDEE5833" bold="true" box="[227,531,869,893]" italics="true" pageId="9" pageNumber="10">Merizomena (s.str.) tricolor</emphasis>
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Subgenus
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<emphasis id="B93BEAD29D7EFFBFFC45FA74FBA45EAC" bold="true" box="[967,1113,1482,1506]" italics="true" pageId="14" pageNumber="15">Dichromenta</emphasis>
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species
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<emphasis id="B93BEAD29D7EFFBFFC6BF9B2FB8D5D6D" box="[1001,1136,1548,1571]" italics="true" pageId="14" pageNumber="15">Merizomena</emphasis>
(
<emphasis id="B93BEAD29D7EFFBFFB01F9B2FAED5D6D" box="[1155,1296,1548,1571]" italics="true" pageId="14" pageNumber="15">Dichromenta</emphasis>
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<emphasis id="B93BEAD29D7EFFBFFAA6F9B2FC735D0A" italics="true" pageId="14" pageNumber="15">castanea</emphasis>
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<emphasis id="B93BEAD29D7EFFBFFCD0F9D0FC245DC8" bold="true" box="[850,985,1646,1670]" pageId="14" pageNumber="15">Description.</emphasis>
Small to medium sized
<taxonomicName id="4C4F4D439D7EFFBFFB4DF9D0FAE65DCB" authorityName="Bonelli" authorityYear="1810" box="[1231,1307,1646,1669]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="tribe" tribe="Lebiini">Lebiini</taxonomicName>
(BL:
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); body moderately flat. Head and pronotum monochromous yellowish to reddish, elytra
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<emphasis id="B93BEAD29D7FFFBEFDCDFF2FFC675BEA" box="[591,922,145,164]" italics="true" pageId="15" pageNumber="16">Muilwijk J., Felix R., Anichtchenko A.</emphasis>
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<paragraph id="8BF036C09D7FFFBEFF61FF7FFD1F5A57" blockId="15.[227,738,193,281]" pageId="15" pageNumber="16">bicolored: base yellowish­red, distal part dark, without metallic lustre, ventral side yellowish, apex more or less brownish; appendages yellowish.</paragraph>
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<emphasis id="B93BEAD29D7FFFBEFF61FEFDFEDA5A15" bold="true" box="[227,295,323,347]" pageId="15" pageNumber="16">Head.</emphasis>
Elongate, tempora with short setae. Frons and vertex with punctures and pubescence. Supraorbital furrow sharp. Maxillary palpi with basal segment widened apically, not thickened. Ultimate palpomere fusiform, slightly longer than penultimate one, with a few small hairs. Penultimate labial palpomere with two long setae on anterior margin.Apical segment of labial palpi securiform, with stout hairs. Gula convex. Mentum punctate at lateral sides, with two setae at anterior margin. Mentum tooth long, slightly shorter than lateral lobes, broadly rounded at apex. Genae finely punctate, with fine erect hairs. Antennae slender, exceeding elytral base approximately by two antennomeres.
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<emphasis id="B93BEAD29D7FFFBEFF61FCF2FEC2582A" bold="true" box="[227,319,844,868]" pageId="15" pageNumber="16">Thorax.</emphasis>
Pronotum cordate slightly transverse; covered with punctures and pubescence; lateral gutter wide. Anterior margin subrectilinear. Both anterior and posterior margin with a fringe of fine hairs. Anterior angles broadly rounded. Base without margin in middle, lateral margin with hairs. Without basal transverse impression. Lateral sides with one seta in basal angle, and one (two) approximately at widest point. Prosternal process finely marginated, with a few long setae. Metacoxae with two long setae. Prosternum, proepisterna, meso­ and mesasternum and meso­ and metepisterna with scattered punctures and hairs, denser on proepisterna and meso­ and metepisterna. Metepisterna long. Elytral apical border with a few fine punctures.
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<emphasis id="B93BEAD29D7FFFBEFF61FACAFEA75EC2" bold="true" box="[227,346,1396,1420]" pageId="15" pageNumber="16">Abdomen.</emphasis>
Elytra relative long (approximately 3.5 times longer than the pronotum) and wide (two times wider than the pronotum), moderately convex. Basal border complete. Sutural apical angle nearly at right angle, with or without a small denticle. Striae slightly embedded; striae 7 and 8 curve to sutural apex. Umbilicate series continuous, consisting of 1518 setiferous pores; one preapical pore located at apex of stria 3. Ventrites pubescent, denser on anal ventrite, anal ventrite with 5 pairs of setae along apical margin.
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<emphasis id="B93BEAD29D7FFFBEFC84FF7FFCBC5B97" bold="true" box="[774,833,193,217]" pageId="15" pageNumber="16">Legs.</emphasis>
More slender than in the nominate subgenus, especially metatibiae and tarsi. Dorsal surface of tarsi provided with sparse long hairs.
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<emphasis id="B93BEAD29D7FFFBEFC84FEFDFC5C5A15" bold="true" box="[774,929,323,347]" pageId="15" pageNumber="16">Median lobe.</emphasis>
Long, endophallus with 1 or 2 copulatory pieces.
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<emphasis id="B93BEAD29D7FFFBEFC84FE1AFC315AF2" bold="true" box="[774,972,420,444]" pageId="15" pageNumber="16">Female genitalia.</emphasis>
Gonocoxite 1 without setae, gonocoxite 2 cone shaped with setae in the apical quarter. Laterotergite IX curved; wider and less strongly chitinized as in species from the nominate subgenus. The spermatheca is divided in a pedicel ending in the bursa copulatrix and beginning at the point of the short branch of the spermatheca in which the duct of the spermathecal gland flows and a thicker distal part. The duct of spermathecal gland is very long and thin. The spermatheca is covered with dense villi in the middle part.
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<emphasis id="B93BEAD29D7FFFBEFC84FCF2FC17582A" bold="true" box="[774,1002,844,868]" pageId="15" pageNumber="16">Comparative notes.</emphasis>
Species of this subgenus differ from species of the nominate subgenus by the elytral base completely marginated, the more elongated and flat body, the slenderer appendages, and yellowish colour of head and ventral side.
</paragraph>
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<emphasis id="B93BEAD29D7FFFBEFC84FBB1FC7B5F69" bold="true" box="[774,902,1039,1063]" pageId="15" pageNumber="16">Etymology.</emphasis>
The name of the subgenus is derived from the bicolour dorsum.
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<subSubSection id="C35965422F14916CEE2A75E4FE11A3C3" box="[114,451,177,202]" pageId="3" pageNumber="252" type="nomenclature">
<paragraph id="8BFC36C92F14916CEE2A75E4FE11A3C3" blockId="3.[114,451,177,232]" box="[114,451,177,202]" pageId="3" pageNumber="252">
<heading id="D0B481A52F14916CEE2A75E4FE11A3C3" bold="true" box="[114,451,177,202]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="252" reason="4">
<emphasis id="B937EADB2F14916CEE2A75E4FE11A3C3" bold="true" box="[114,451,177,202]" pageId="3" pageNumber="252">
<taxonomicName id="4C434D4A2F14916CEE2A75E4FEB7A3C0" authorityName="Barševskis" authorityYear="2023" box="[114,357,178,202]" class="Insecta" family="Cerambycidae" genus="Cleptometopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="pepitoi" status="sp. nov.">
<emphasis id="B937EADB2F14916CEE2A75E4FEB7A3C0" bold="true" box="[114,357,178,202]" italics="true" pageId="3" pageNumber="252">Cleptometopus pepitoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457A02F14916CEF3475E7FE11A3C3" box="[364,451,177,201]" pageId="3" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35965422F14916CEE2A7586FC10A081" pageId="3" pageNumber="252" type="description">
<paragraph id="8BFC36C92F14916CEE2A7586FF10A3E2" blockId="3.[114,451,177,232]" box="[114,194,208,232]" pageId="3" pageNumber="252">
(
<figureCitation id="13782A4C2F14916CEE227586FF6BA3E2" box="[122,185,208,232]" captionStart="Fig" captionStartId="3.[114,153,1009,1033]" captionTargetBox="[122,578,273,962]" captionTargetId="figure-11@3.[114,585,263,971]" captionTargetPageId="3" captionText="Fig. 3. Cleptometopus pepitoi sp. nov. [holotype]" pageId="3" pageNumber="252">Fig. 3</figureCitation>
)
</paragraph>
<paragraph id="8BFC36C92F14916CEC2875E4FC1DA24E" blockId="3.[624,1103,178,324]" pageId="3" pageNumber="252">covered with dense grey pubescence. Surface of head covered with dense grey pubescence and punctation. Head without longitudinal middle line, except basal portion of head behind antennae.</paragraph>
<paragraph id="8BFC36C92F14916CEC28743CFCC3A19F" blockId="3.[624,1103,362,661]" pageId="3" pageNumber="252">Labrum brown, shiny, pubescent. Clypeus brown, narrow, transverse, shiny. Mandibles dark, shiny, with acute apices, with very fine, punctures in basal part, covered with grey pubescence in basal portion. Antennae dark-brown, very long, covered with very fine pubescence; basal antennomere cylindric, black, with sparse fine punctures and pubescence and with row of bristles.</paragraph>
<paragraph id="8BFC36C92F14916CEC2877EDFC10A081" blockId="3.[624,1102,699,907]" pageId="3" pageNumber="252">Pronotum almost sub-cylindrical, glossy, black. Pronotal disc dorsally with coarse dense punctures. Lateral sides of pronotal disc with elongated band of yellow pubescence. Basal portion of pronotum without acute angles. Frontal and basal portions of pronotum slightly neck-shaped.</paragraph>
</subSubSection>
<caption id="DF3C66412F14916CEE2A76A7FF33A72D" pageId="3" pageNumber="252" startId="3.[114,153,1009,1033]" targetBox="[122,578,273,962]" targetPageId="3" targetType="figure">
<paragraph id="8BFC36C92F14916CEE2A76A7FF33A72D" blockId="3.[114,587,1009,1063]" pageId="3" pageNumber="252">
Fig. 3.
<taxonomicName id="4C434D4A2F14916CEE8076A7FE04A702" authorityName="Barševskis" authorityYear="2023" box="[216,470,1009,1033]" class="Insecta" family="Cerambycidae" genus="Cleptometopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="pepitoi" status="sp. nov.">
<emphasis id="B937EADB2F14916CEE8076A7FE04A702" box="[216,470,1009,1033]" italics="true" pageId="3" pageNumber="252">Cleptometopus pepitoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457A02F14916CEFB176A7FD99A703" box="[489,587,1009,1033]" pageId="3" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
[holotype]
</paragraph>
</caption>
<subSubSection id="C35965422F14916CEE2A711AFED2A673" pageId="3" pageNumber="252" type="materials_examined">
<paragraph id="8BFC36C92F14916CEE2A711AFEC0A76E" blockId="3.[114,593,1100,1309]" box="[114,274,1100,1124]" pageId="3" pageNumber="252">
<emphasis id="B937EADB2F14916CEE2A711AFEC0A76E" bold="true" box="[114,274,1100,1124]" pageId="3" pageNumber="252">Type material</emphasis>
</paragraph>
<paragraph id="8BFC36C92F14916CEE2A713DFEDFA617" blockId="3.[114,593,1100,1309]" pageId="3" pageNumber="252">
<emphasis id="B937EADB2F14916CEE2A713DFF31A789" bold="true" box="[114,227,1131,1155]" pageId="3" pageNumber="252">
<typeStatus id="54F8886B2F14916CEE2A713DFF0CA789" box="[114,222,1131,1155]" pageId="3" pageNumber="252" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male.
<collectingCountry id="F35476592F14916CEF1A713DFE6FA789" box="[322,445,1131,1155]" name="Philippines" pageId="3" pageNumber="252">Philippines</collectingCountry>
,
<collectingRegion id="4987F82B2F14916CEF89713DFDFCA789" box="[465,558,1131,1155]" country="Philippines" name="Palawan" pageId="3" pageNumber="252">Palawan</collectingRegion>
, / Roxas, 09.2022, / local collector leg. &lt;handwritten&gt; //
<typeStatus id="54F8886B2F14916CEF6E71FFFE0AA7CB" box="[310,472,1193,1217]" pageId="3" pageNumber="252" type="holotype">HOLOTYPUS</typeStatus>
: /
<emphasis id="B937EADB2F14916CEFA271FFFEE4A7D5" italics="true" pageId="3" pageNumber="252">
<taxonomicName id="4C434D4A2F14916CEFA271FFFF19A7D5" authorityName="Thomson" authorityYear="1864" class="Insecta" family="Cerambycidae" genus="Cleptometopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="genus">Cleptometopus</taxonomicName>
/
<taxonomicName id="4C434D4A2F14916CEEB4719EFEE4A7D5" authorityName="Barševskis" authorityYear="2023" box="[236,310,1224,1247]" class="Insecta" family="Cerambycidae" genus="Cleptometopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="pepitoi" status="sp. nov.">pepitoi</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20457A02F14916CEF1A7191FE4EA7D5" box="[322,412,1223,1247]" pageId="3" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
/ A. Barševskis det. 2023. &lt;red label, handwritten&gt;. Deposited in
<collectionCode id="ED52AE0C2F14916CEEE47053FEDBA617" box="[188,265,1285,1309]" pageId="3" pageNumber="252">DUBC</collectionCode>
.
</paragraph>
<paragraph id="8BFC36C92F14916CEE2A7014FED2A673" blockId="3.[114,585,1346,1401]" pageId="3" pageNumber="252">
<emphasis id="B937EADB2F14916CEE2A7014FEAEA650" bold="true" box="[114,380,1346,1370]" pageId="3" pageNumber="252">General distribution:</emphasis>
<collectingCountry id="F35476592F14916CEFD67014FDD9A650" box="[398,523,1346,1370]" name="Philippines" pageId="3" pageNumber="252">Philippines</collectingCountry>
:
<collectingRegion id="4987F82B2F14916CEC7E7014FF60A673" country="Philippines" name="Palawan" pageId="3" pageNumber="252">Palawan</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C35965422F14916BEE2A70C8FDECA3E2" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252" type="description">
<paragraph id="8BFC36C92F14916CEE2A70C8FDC2A518" blockId="3.[114,592,1438,1554]" pageId="3" pageNumber="252">
<emphasis id="B937EADB2F14916CEE2A70C8FED1A6BC" bold="true" box="[114,259,1438,1462]" pageId="3" pageNumber="252">Description.</emphasis>
Body elongate, very narrow, dark, lustrous, surface with brown, elongated, interrupted, lines. Body length:
<quantity id="4CBB9B2C2F14916CEC71708DFF4CA518" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="252" unit="mm" value="9.5">9.5 mm</quantity>
, maximal width of elytra:
<quantity id="4CBB9B2C2F14916CEFEF70ACFDDFA518" box="[439,525,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="252" unit="mm" value="1.9">1.9 mm</quantity>
.
</paragraph>
<paragraph id="8BFC36C92F14916CEE2A7361FD82A564" blockId="3.[114,593,1591,1646]" pageId="3" pageNumber="252">Head flat, wide, with almost parallel sides, with convex eyes. Cheeks not extended,</paragraph>
<paragraph id="8BFC36C92F14916CEE2A73F0FC8EA72E" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1103,944,1060]" pageId="3" pageNumber="252">
252 Scutellum small, rounded apically, covered with yellow pubescence.
<emphasis id="B937EADB2F14916CEDD27686FBC9A0ED" box="[906,1051,975,999]" italics="true" pageId="3" pageNumber="252">Pars stridens</emphasis>
not visible, because it is under the posterior edge of the pronotum.
</paragraph>
<paragraph id="8BFC36C92F14916CEC28711FFD36A6B9" blockId="3.[624,1103,1097,1459]" pageId="3" pageNumber="252">Elytra dark-brown, glossy, with relatively coarse, slightly impressed punctures in rows. Each elytron with slightly - developed humps between shoulders and suturae. Dorsal part of elytra behind middle with wide impression. Dorsal surface of elytra behind middle with four elongated brown short lines, covered with yellow pubescence and whith brown lateral portion behind shoulders. Apical part of elytra smoth, with large sharp extended projection.</paragraph>
<paragraph id="8BFC36C92F14916BEC28708EFDECA3E2" blockId="3.[624,1103,1496,1612]" lastBlockId="4.[189,661,178,232]" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252">Ventral surface of body black, with pubescence. Legs relatively short, dark-brown, slightly shiny, covered with very fine pubescence. Inner sides of tibia with fine yellow pubescence. Tarsomeres dark-brown, covered by fine pubescence.</paragraph>
</subSubSection>
<subSubSection id="C35965422F13916BEEE5745BFE78A152" pageId="4" pageNumber="253" type="diagnosis">
<paragraph id="8BFC36C92F13916BEEE5745BFE78A152" blockId="4.[189,668,269,600]" pageId="4" pageNumber="253">
<emphasis id="B937EADB2F13916BEEE5745BFE1AA22F" bold="true" box="[189,456,269,293]" pageId="4" pageNumber="253">Differential diagnosis.</emphasis>
The new species differs from
<taxonomicName id="4C434D4A2F13916BEF34747AFEB3A269" authority="Newman, 1842" authorityName="Newman" authorityYear="1842" class="Insecta" family="Cerambycidae" genus="Cleptometopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="camuripes">
<emphasis id="B937EADB2F13916BEF34747AFD47A24E" box="[364,661,300,324]" italics="true" pageId="4" pageNumber="253">Cleptometopus camuripes</emphasis>
Newman, 1842
</taxonomicName>
by a different pattern on the elytra: dorsal surface of elytra behind middle with four elongated brown short lines, covered with yellow pubescence and whith brown lateral portion behind shoulders, but
<taxonomicName id="4C434D4A2F13916BEF3974B2FE21A2F6" authorityName="Newman" authorityYear="1842" box="[353,499,484,508]" class="Insecta" family="Cerambycidae" genus="Cleptometopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="camuripes">
<emphasis id="B937EADB2F13916BEF3974B2FE21A2F6" box="[353,499,484,508]" italics="true" pageId="4" pageNumber="253">C. camuripes</emphasis>
</taxonomicName>
dorsal surface of elytra in the middle part with four short yellov lines and without yellow lateral portion behind shoulders.
</paragraph>
</subSubSection>
<subSubSection id="C35965422F13916BEEE5772BFEEFA047" pageId="4" pageNumber="253" type="etymology">
<paragraph id="8BFC36C92F13916BEEE5772BFEEFA047" blockId="4.[189,669,637,845]" pageId="4" pageNumber="253">
<emphasis id="B937EADB2F13916BEEE5772BFEEDA19F" bold="true" box="[189,319,637,661]" pageId="4" pageNumber="253">Etymology.</emphasis>
Patronym. This species is named after my good colleague Mark John Pepito (University of Mindanao,
<collectingRegion id="4987F82B2F13916BEC1277EDFD4AA1D9" box="[586,664,699,723]" country="Philippines" name="Davao" pageId="4" pageNumber="253">Davao</collectingRegion>
,
<collectingCountry id="F35476592F13916BEEE5778CFEE5A1F8" box="[189,311,730,754]" name="Philippines" pageId="4" pageNumber="253">Philippines</collectingCountry>
) in appreciation of excellent cooperation, and in gratitude for his contribution to
<taxonomicName id="4C434D4A2F13916BEFDC7641FE2BA025" authorityName="Linnaeus" authorityYear="1758" box="[388,505,791,815]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
research in
<collectingCountry id="F35476592F13916BEEE57663FEEBA047" box="[189,313,821,845]" name="Philippines" pageId="4" pageNumber="253">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,153 @@
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<mods:title id="830D2668DD70AA4FEC0FF628A988BE70">Two new species of long-horned beetles (Coleoptera: Cerambycidae) from the Philippines</mods:title>
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<heading id="D0B481A52F16916EEE2A7148FE1FA73C" bold="true" box="[114,461,1054,1078]" fontSize="10" level="2" pageId="1" pageNumber="250" reason="4">
<emphasis id="B937EADB2F16916EEE2A7148FE1FA73C" bold="true" box="[114,461,1054,1078]" pageId="1" pageNumber="250">
<taxonomicName id="4C434D4A2F16916EEE2A7148FEBDA73C" authorityName="Barševskis" authorityYear="2023" box="[114,367,1054,1078]" class="Insecta" family="Cerambycidae" genus="Chydaeopsis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="torrejosae" status="sp. nov.">
<emphasis id="B937EADB2F16916EEE2A7148FEBDA73C" bold="true" box="[114,367,1054,1078]" italics="true" pageId="1" pageNumber="250">Chydaeopsis torrejosae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457A02F16916EEF2E7148FE1FA73C" box="[374,461,1054,1078]" pageId="1" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35965422F16916EEE2A716BFF13A75F" box="[114,193,1085,1109]" pageId="1" pageNumber="250" type="description">
<paragraph id="8BFC36C92F16916EEE2A716BFF13A75F" blockId="1.[114,461,1054,1109]" box="[114,193,1085,1109]" pageId="1" pageNumber="250">
(
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)
</paragraph>
</subSubSection>
<subSubSection id="C35965422F16916EEE2A712CFF68A6AC" pageId="1" pageNumber="250" type="materials_examined">
<paragraph id="8BFC36C92F16916EEE2A712CFEC0A798" blockId="1.[114,593,1146,1355]" box="[114,274,1146,1170]" pageId="1" pageNumber="250">
<emphasis id="B937EADB2F16916EEE2A712CFEC0A798" bold="true" box="[114,274,1146,1170]" pageId="1" pageNumber="250">Type material</emphasis>
</paragraph>
<paragraph id="8BFC36C92F16916EEE2A71CFFE5CA641" blockId="1.[114,593,1146,1355]" pageId="1" pageNumber="250">
<emphasis id="B937EADB2F16916EEE2A71CFFF31A7BB" bold="true" box="[114,227,1177,1201]" pageId="1" pageNumber="250">
<typeStatus id="54F8886B2F16916EEE2A71CFFF0CA7BB" box="[114,222,1177,1201]" pageId="1" pageNumber="250" type="holotype">Holotype</typeStatus>
:
</emphasis>
male.
<collectingCountry id="F35476592F16916EEF6171CFFE61A7BB" box="[313,435,1177,1201]" name="Philippines" pageId="1" pageNumber="250">Philippines</collectingCountry>
, Panay Isl. /
<collectingRegion id="4987F82B2F16916EEE2A71EEFF19A7DA" box="[114,203,1208,1232]" country="Philippines" name="Antique" pageId="1" pageNumber="250">Antique</collectingRegion>
, 09.2018 / local collector leg. &lt;handwritten&gt; //
<typeStatus id="54F8886B2F16916EEF2B7180FDC7A7E4" box="[371,533,1238,1262]" pageId="1" pageNumber="250" type="holotype">HOLOTYPUS</typeStatus>
: /
<emphasis id="B937EADB2F16916EEE2A71A3FE41A607" box="[114,403,1269,1293]" italics="true" pageId="1" pageNumber="250">
<taxonomicName id="4C434D4A2F16916EEE2A71A3FF28A607" authorityName="Pascoe" authorityYear="1864" box="[114,250,1269,1293]" class="Insecta" family="Cerambycidae" genus="Chydaeopsis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="genus">Chydaeopsis</taxonomicName>
/
<taxonomicName id="4C434D4A2F16916EEF7E71A0FE41A607" authorityName="Barševskis" authorityYear="2023" box="[294,403,1270,1293]" class="Insecta" family="Cerambycidae" genus="Chydaeopsis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="torrejosae" status="sp. nov.">torrejosae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20457A02F16916EEFFD71A3FDD7A607" box="[421,517,1269,1293]" pageId="1" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
/ A. Barševskis det. 2023. &lt;red label, handwritten&gt;. Deposited in
<collectionCode id="ED52AE0C2F16916EEF657065FE58A641" box="[317,394,1331,1355]" pageId="1" pageNumber="250">DUBC</collectionCode>
.
</paragraph>
<paragraph id="8BFC36C92F16916EEE2A7039FF68A6AC" blockId="1.[114,586,1391,1446]" pageId="1" pageNumber="250">
<emphasis id="B937EADB2F16916EEE2A7039FEA1A68D" bold="true" box="[114,371,1391,1415]" pageId="1" pageNumber="250">General distribution:</emphasis>
<collectingCountry id="F35476592F16916EEF237026FE2AA682" box="[379,504,1392,1416]" name="Philippines" pageId="1" pageNumber="250">Philippines</collectingCountry>
: Panay Island.
</paragraph>
</subSubSection>
<subSubSection id="C35965422F16916DEE2A709DFC97A22F" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250" type="description">
<paragraph id="8BFC36C92F16916EEE2A709DFEC4A54A" blockId="1.[114,593,1483,1600]" pageId="1" pageNumber="250">
<emphasis id="B937EADB2F16916EEE2A709DFED1A6E9" bold="true" box="[114,259,1483,1507]" pageId="1" pageNumber="250">Description.</emphasis>
Body elongate, black, surface with dark and silver-grey pubescence (
<figureCitation id="13782A4C2F16916EEC7870BDFFADA52B" captionStart="Fig" captionStartId="1.[624,663,1015,1039]" captionTargetBox="[624,1092,175,978]" captionTargetId="figure-192@1.[624,1099,171,978]" captionTargetPageId="1" captionText="Fig. 1. Chydaeopsis torrejosae sp. nov. [holotype]" pageId="1" pageNumber="250">Fig. 1</figureCitation>
). Body length:
<quantity id="4CBB9B2C2F16916EEF72735FFEADA52B" box="[298,383,1545,1569]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="250" unit="mm" value="9.1">9.1 mm</quantity>
, maximal width of elytra:
<quantity id="4CBB9B2C2F16916EEEE4737EFEC0A54A" box="[188,274,1576,1600]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="250" unit="mm" value="2.6">2.6 mm</quantity>
.
</paragraph>
<paragraph id="8BFC36C92F16916DEE2A73F0FEFCA3E2" blockId="1.[114,154,1702,1726]" lastBlockId="2.[189,668,178,232]" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250">250 Head flat, wide, with almost parallel sides, with slightly convex eyes. Cheeks not extended, covered with silver-grey sparse pubescence. Surface of head shiny, with fine punctation under dense pubescence. Middle portion of head with longitudinal thin line, slightly impressed in the middle part between eyes. Labrum pubescent, punctated, covered with pubescence. Clypeus brown, narrow, transverse, shiny. Mandibles shiny, dark brown, massive, with acute apices, with very fine, sparse punctures in basal part, covered with very sparse silver-grey pubescence. Antennae thin, dark-brown, very long, about twice as long as the body, covered with dense dark pubescence; basal antennomere thickened, with sparse fine punctures and pubescence. 1st, 3rd, 4th and 5th antennomeres with row of bristles.</paragraph>
<paragraph id="8BFC36C92F15916DEEE5745BFD84A130" blockId="2.[189,668,269,570]" pageId="2" pageNumber="251">Pronotum dark-brown, almost cylindrical, convex and matt, with very fine microsculpture. Frontal and basal portions of pronotum slightly neck-shaped narrowed, without basal angles. Pronotal disc black, dorsally covered with dense fine dark pubescence, with middle line, covered with white pubescence and form narrow elongated band. Lateral portions of pronotal disc covered with dense white pubescence.</paragraph>
<paragraph id="8BFC36C92F15916DECE375E4FC97A22F" blockId="2.[699,1178,178,293]" pageId="2" pageNumber="251">bands behind the middle and before apex of elytra. Along the suturae before middle elytra of new species without two oval brown spots.</paragraph>
</subSubSection>
<subSubSection id="C35965422F15916DECE3741CFEC3A6DB" pageId="2" pageNumber="251" type="etymology">
<paragraph id="8BFC36C92F15916DECE3741CFCE9A2F6" blockId="2.[699,1178,330,508]" pageId="2" pageNumber="251">
<emphasis id="B937EADB2F15916DECE3741CFCEFA268" bold="true" box="[699,829,330,354]" pageId="2" pageNumber="251">Etymology.</emphasis>
Patronym. This species is named after the my good colleague Chrestine Torrejos (University of Mindanao,
<collectingRegion id="4987F82B2F15916DECE374F1FCD1A2B5" box="[699,771,423,447]" country="Philippines" name="Davao" pageId="2" pageNumber="251">Davao</collectingRegion>
,
<collectingCountry id="F35476592F15916DED4874F1FC58A2B5" box="[784,906,423,447]" name="Philippines" pageId="2" pageNumber="251">Philippines</collectingCountry>
) in gratitude to her contribution to
<taxonomicName id="4C434D4A2F15916DED377493FC36A2D7" authorityName="Linnaeus" authorityYear="1758" box="[879,996,453,477]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
research in
<collectingCountry id="F35476592F15916DECE374B2FCE5A2F6" box="[699,823,484,508]" name="Philippines" pageId="2" pageNumber="251">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8BFC36C92F15916DEEE57709FEFEA1F8" blockId="2.[189,668,607,754]" pageId="2" pageNumber="251">
Scutellum small, rounded apically, covered with white dense pubescence, without middle impression.
<emphasis id="B937EADB2F15916DEFC977CBFDCCA1BE" box="[401,542,668,692]" italics="true" pageId="2" pageNumber="251">Pars stridens</emphasis>
not visible because it is under the posterior edge of the pronotum.
</paragraph>
<paragraph id="8BFC36C92F15916DEEE57641FE6FA632" blockId="2.[189,669,791,1336]" pageId="2" pageNumber="251">Elytra dark brown, behind middle darkened, matt, with relatively coarse punctures in regular rows, slightly impressed in basal part. Each elytron with well-developed flattened hump behind scutellum around suturae. Elytral shoulders not extended, weekly developed. Dorsal part of elytra and suturae in basal portion before middle paler and covered with very fine pubesceence, except brown 2nd - 3rd intervals, covered by dark pubescence. Other portions of elytra dark and covered with dark pubescence. Middle portion of elytra toward apex with dark-brown suturae, darkened dorsally. Apical portions of elytra dorsally with two small pale spots. Apex of elytra triangular, near suture with longer pale, dense pubescence.</paragraph>
<paragraph id="8BFC36C92F15916DEEE5700BFEC3A6DB" blockId="2.[189,669,1373,1489]" pageId="2" pageNumber="251">Ventral surface of body black, with white pubescence in lateral portions. Legs dark-brown, covered with dark pubescence and bristles.</paragraph>
</subSubSection>
<subSubSection id="C35965422F15916DEEE570A0FB46A5B4" pageId="2" pageNumber="251" type="diagnosis">
<paragraph id="8BFC36C92F15916DEEE570A0FD47A561" blockId="2.[189,668,1526,1643]" pageId="2" pageNumber="251">
<emphasis id="B937EADB2F15916DEEE570A0FE1AA504" bold="true" box="[189,456,1526,1550]" pageId="2" pageNumber="251">Differential diagnosis.</emphasis>
The new species differs from
<taxonomicName id="4C434D4A2F15916DEF087340FF21A546" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" class="Insecta" family="Cerambycidae" genus="Chydaeopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="luzonica">
<emphasis id="B937EADB2F15916DEF087340FDEDA524" box="[336,575,1558,1582]" italics="true" pageId="2" pageNumber="251">Chydaeopsis luzonica</emphasis>
Heller, 1923
</taxonomicName>
(
<figureCitation id="13782A4C2F15916DEF5D7362FE9AA546" box="[261,328,1588,1612]" captionStart="Fig" captionStartId="2.[699,738,1573,1597]" captionTargetBox="[699,1168,539,1536]" captionTargetId="figure-316@2.[699,1168,539,1536]" captionTargetPageId="2" captionText="Fig. 2. Chydaeopsis luzonica Heller, 1923 [holotype] (Berzak 2023)" pageId="2" pageNumber="251">Fig. 2</figureCitation>
) by a different pattern on the elytra: there is no black wide transversal
</paragraph>
<caption id="DF3C66412F15916DECE37373FC18A556" pageId="2" pageNumber="251" startId="2.[699,738,1573,1597]" targetBox="[699,1168,539,1536]" targetPageId="2" targetType="figure">
<paragraph id="8BFC36C92F15916DECE37373FC18A556" blockId="2.[699,1172,1573,1628]" pageId="2" pageNumber="251">
Fig. 2.
<taxonomicName id="4C434D4A2F15916DED497370FB46A537" authority="Heller, 1923" authorityName="Heller" authorityYear="1923" box="[785,1172,1573,1598]" class="Insecta" family="Cerambycidae" genus="Chydaeopsis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="luzonica">
<emphasis id="B937EADB2F15916DED497370FC2CA534" box="[785,1022,1574,1598]" italics="true" pageId="2" pageNumber="251">Chydaeopsis luzonica</emphasis>
Heller, 1923
</taxonomicName>
[holotype] (
<bibRefCitation id="EFD24B382F15916DED607312FC11A556" author="Berzak L." box="[824,963,1604,1628]" pageId="2" pageNumber="251" refId="ref1918" refString="Berzak L. 2023. Old World Cerambycidae Catalog. http: // bezbycids. com / byciddb / wdefaul t. asp? w = o [accessed: 01.11. 2023]." type="url" year="2023">Berzak 2023</bibRefCitation>
)
</paragraph>
</caption>
<paragraph id="8BFC36C92F15916DEA3473F0FB46A5B4" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="251">251</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,241 @@
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<mods:title id="E177954E874CC933109D61F0C1D39554">Two new species ofCaciaNewman (Coleoptera, Cerambycidae, Lamiinae) from the Mindoro Biogeographic Region of the Philippines</mods:title>
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<mods:namePart id="7CB4F587EFB865CA740B549E9269DD9C">Medina, Milton Norman</mods:namePart>
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<mods:namePart id="98D4336E089015897770BB025FF973AA">Cabras, Analyn</mods:namePart>
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<mods:date id="7B7D79858853AA84C5E4A13883EDE11A">2023</mods:date>
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<treatment id="03EF2F64FFF1DC62ED77D6F6FC5DFEC8" LSID="urn:lsid:plazi:treatment:03EF2F64FFF1DC62ED77D6F6FC5DFEC8" httpUri="http://treatment.plazi.org/id/03EF2F64FFF1DC62ED77D6F6FC5DFEC8" lastPageId="6" lastPageNumber="247" pageId="3" pageNumber="244">
<subSubSection id="C35CCDF9FFF1DC67ED77D6F6FC59FA19" box="[624,975,1434,1459]" pageId="3" pageNumber="244" type="nomenclature">
<paragraph id="8BF99E72FFF1DC67ED77D6F6FC59FA19" blockId="3.[624,975,1434,1489]" box="[624,975,1434,1459]" pageId="3" pageNumber="244">
<heading id="D0B1291EFFF1DC67ED77D6F6FC59FA19" bold="true" box="[624,975,1434,1459]" fontSize="10" level="2" pageId="3" pageNumber="244" reason="4">
<emphasis id="B9324260FFF1DC67ED77D6F6FC59FA19" bold="true" box="[624,975,1434,1459]" pageId="3" pageNumber="244">
<taxonomicName id="4C46E5F1FFF1DC67ED77D6F6FCE7FA18" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" box="[624,881,1435,1459]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="aeschyae" status="sp. nov." subGenus="Cacia">
<emphasis id="B9324260FFF1DC67ED77D6F6FCE7FA18" bold="true" box="[624,881,1435,1459]" italics="true" pageId="3" pageNumber="244">Cacia (Cacia) aeschyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF1DC67EC7FD6F7FC59FA19" box="[888,975,1434,1458]" pageId="3" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF1DC67ED77D6D4FD43FA7A" box="[624,725,1465,1489]" pageId="3" pageNumber="244" type="description">
<paragraph id="8BF99E72FFF1DC67ED77D6D4FD43FA7A" blockId="3.[624,975,1434,1489]" box="[624,725,1465,1489]" pageId="3" pageNumber="244">
<emphasis id="B9324260FFF1DC67ED77D6D4FD43FA7A" bold="true" box="[624,725,1465,1489]" pageId="3" pageNumber="244">
(
<figureCitation id="137D82F7FFF1DC67ED70D6D4FD5BFA7A" box="[631,717,1465,1489]" captionStart="Fig" captionStartId="4.[189,228,1509,1533]" captionTargetBox="[255,1112,829,1455]" captionTargetId="figure-270@4.[229,1132,815,1472]" captionTargetPageId="4" captionText="Fig. 2. Genitalia ofCacia (Ipocregyes) katrinaesp. nov.: A-C. Whole system of genitalia, A. Lateral aspect, B. Ventral aspect, C. Dorsal aspect. D-F. Aedeagus, D. Dorsal aspect, E. Ventral aspect, F. Lateral aspect. G. Tegmen." pageId="3" pageNumber="244">Fig. 2A</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF1DC60ED77D69BFEE2FF43" lastPageId="4" lastPageNumber="245" pageId="3" pageNumber="244" type="materials_examined">
<paragraph id="8BF99E72FFF1DC60ED77D69BFEE2FF43" blockId="3.[624,1103,1526,1643]" lastBlockId="4.[189,668,178,232]" lastPageId="4" lastPageNumber="245" pageId="3" pageNumber="244">
<materialsCitation id="3B2E942FFFF1DC60ED77D69BFEE2FF43" collectingDate="2017-06" collectionCode="MMCP, PNM" country="Philippines" lastPageId="4" lastPageNumber="245" location="Puerto Gallera" municipality="Southern Luzon" pageId="3" pageNumber="244" specimenCount="1" stateProvince="Mindoro Oriental" typeStatus="holotype">
<emphasis id="B9324260FFF1DC67ED77D69BFC91F9A5" bold="true" box="[624,775,1526,1550]" pageId="3" pageNumber="244">
<typeStatus id="54FD20D0FFF1DC67ED77D69BFC91F9A5" box="[624,775,1526,1550]" pageId="3" pageNumber="244" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
male:
<collectingCountry id="F351DEE2FFF1DC67EC78D69AFB8AF9A4" box="[895,1052,1527,1551]" name="Philippines" pageId="3" pageNumber="244">PHILIPPINES</collectingCountry>
<collectingMunicipality id="6B9D0408FFF1DC67ED77D578FCB6F986" box="[624,800,1557,1581]" pageId="3" pageNumber="244">Southern Luzon</collectingMunicipality>
,
<collectingRegion id="49825090FFF1DC67EC2BD578FC7FF986" box="[812,1001,1557,1581]" country="Philippines" name="Mindoro Oriental" pageId="3" pageNumber="244">Oriental Mindoro</collectingRegion>
/
<location id="8E99C8A9FFF1DC67EB04D57BFD28F9E7" LSID="urn:lsid:plazi:treatment:03EF2F64FFF1DC62ED77D6F6FC5DFEC8:8E99C8A9FFF1DC67EB04D57BFD28F9E7" country="Philippines" municipality="Southern Luzon" name="Puerto Gallera" pageId="3" pageNumber="244" stateProvince="Mindoro Oriental">Puerto Gallera</location>
/
<date id="FFF8B8B2FFF1DC67EDE9D559FCD7F9E7" box="[750,833,1588,1612]" pageId="3" pageNumber="244" value="2017-06">
<collectingDate id="EFBC415AFFF1DC67EDE9D559FCD7F9E7" box="[750,833,1588,1612]" pageId="3" pageNumber="244" value="2017-06">vi.2017</collectingDate>
</date>
, loc. collector leg. /
<collectionCode id="ED5706B7FFF1DC67ED77D53EFD55F9C0" box="[624,707,1619,1643]" pageId="3" pageNumber="244">MMCP</collectionCode>
, printed on red card. Type specimen will be deposited at the Philippine National Museum (
<collectionCode id="ED5706B7FFF6DC60EE2ED3BDFEFFFF43" box="[297,361,208,232]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="4" pageNumber="245">PNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF6DC61EFBAD260FC14FC63" lastPageId="5" lastPageNumber="246" pageId="4" pageNumber="245" type="description">
<paragraph id="8BF99E72FFF6DC60EFBAD260FE79FE0B" blockId="4.[189,668,269,416]" pageId="4" pageNumber="245">
<emphasis id="B9324260FFF6DC60EFBAD260FD83FE8E" bold="true" box="[189,533,269,293]" pageId="4" pageNumber="245">Description. Measurements:</emphasis>
<typeStatus id="54FD20D0FFF6DC60ED35D260FD0AFE8E" box="[562,668,269,293]" pageId="4" pageNumber="245" type="holotype">Holotype</typeStatus>
male: LB (LH+LP+LE): mm. LH:
<quantity id="4CBE3397FFF6DC60ED3BD241FD0EFEEF" box="[572,664,300,324]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="245" unit="mm" value="1.5">1.5 mm</quantity>
. WH:
<quantity id="4CBE3397FFF6DC60EFFBD226FEC1FEC9" box="[252,343,331,355]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="245" unit="mm" value="2.5">2.5 mm</quantity>
. LG: 1.0 mm. LL:
<quantity id="4CBE3397FFF6DC60ED3ED226FD0FFEC9" box="[569,665,331,355]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="245" unit="mm" value="0.5">0.5 mm</quantity>
. WL:
<quantity id="4CBE3397FFF6DC60EFFDD204FEC2FE2A" box="[250,340,361,385]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="245" unit="mm" value="0.5">0.5 mm</quantity>
. LP: 2.0 mm. WP: 3.0 mm. LE: 8.0 mm. WEH: 4.0 mm.
</paragraph>
<paragraph id="8BF99E72FFF6DC60EFBAD2A8FE70FDB0" blockId="4.[189,668,453,539]" pageId="4" pageNumber="245">
<emphasis id="B9324260FFF6DC60EFBAD2A8FEADFE76" bold="true" box="[189,315,453,477]" pageId="4" pageNumber="245">Teguments</emphasis>
head, prothorax, and elytradark brownish; underside of the body, antennae, and legs are light brownish.
</paragraph>
<paragraph id="8BF99E72FFF6DC60EFBAD12DFB15FD78" blockId="4.[189,668,576,785]" lastBlockId="4.[699,1179,178,723]" pageId="4" pageNumber="245">
<emphasis id="B9324260FFF6DC60EFBAD12DFE96FDF3" bold="true" box="[189,256,576,600]" pageId="4" pageNumber="245">Head</emphasis>
broader than long; vertex, frons, and genae densely covered with yellowish recumbent pubescence; frons with deep puncturations arranged in random; longitudinal line visible from base to apex; vertex with thick band of yellowish pubescence from base up to margin between frons; genae wider than long; eyes dark brown as long as wide; apical margin of head slightly concave, lined with long erect yellowish setae; clypeus light brownish, lustrous, glabrous; labrum broader than long, densely covered with semi-erect yellowish setae; mandibles lustrous, dark brownish, basal half with puncturations, with lateral depression, and covered with semi-erect yellowish setae. Scape slightly robust towards apex with cicatrix, covered with fined recumbent yellowish pubescence with moderate erect long yellowish setae. Antennomeres III to XI cylindrical, covered with fine recumbent yellowish pubescence, underside with numerous long erect yellowish setae; Antennomere III longer than IV and V combined; antennomeres VI to XI of the same length.
</paragraph>
<caption id="DF39CEFAFFF6DC60EFBAD688FD34F990" pageId="4" pageNumber="245" startId="4.[189,228,1509,1533]" targetBox="[255,1112,829,1455]" targetPageId="4" targetType="figure">
<paragraph id="8BF99E72FFF6DC60EFBAD688FD34F990" blockId="4.[189,1179,1509,1595]" pageId="4" pageNumber="245">
Fig. 2. Genitalia of
<taxonomicName id="4C46E5F1FFF6DC60EE92D68BFD5BFA55" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" box="[405,717,1510,1534]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="katrinae" status="sp. nov." subGenus="Ipocregyes">
<emphasis id="B9324260FFF6DC60EE92D68BFD5BFA55" box="[405,717,1510,1534]" italics="true" pageId="4" pageNumber="245">Cacia (Ipocregyes) katrinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF6DC60EDC9D68BFCBEFA56" box="[718,808,1510,1533]" pageId="4" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
: A-C. Whole system of genitalia, A. Lateral aspect, B. Ventral aspect, C. Dorsal aspect. D-F. Aedeagus, D. Dorsal aspect, E. Ventral aspect, F. Lateral aspect. G. Tegmen.
</paragraph>
</caption>
<paragraph id="8BF99E72FFF6DC61EB6BD5CBFC9EFE8E" blockId="4.[1132,1172,1702,1726]" lastBlockId="5.[624,1103,178,293]" lastPageId="5" lastPageNumber="246" pageId="4" pageNumber="245">245 Femora robust; femora, tibia, and tarsi covered with fine recumbent yellowish pubescence with few short erect yellowish setae. Claws simple.</paragraph>
<paragraph id="8BF99E72FFF7DC61ED77D227FC19FDF3" blockId="5.[624,1103,330,600]" pageId="5" pageNumber="246">
<emphasis id="B9324260FFF7DC61ED77D227FC84FEC9" bold="true" box="[624,786,330,354]" pageId="5" pageNumber="246">Mesosternum</emphasis>
quadrangular, with fine recumbent setae; mesepimeron, mesepisternum, metepisternum, metasternum, and abdominal ventrites densely covered with recumbent yellowish pubescence; mesepimeron slightly wider than long; metepisternum rectangular; ventrite I as broad as ventrite V; ventrite I broader than ventrites II, III, and IV individually.
</paragraph>
<caption id="DF39CEFAFFF7DC61EF75D050FE1EFC39" pageId="5" pageNumber="246" startId="5.[114,153,829,853]" targetBox="[119,587,190,797]" targetPageId="5" targetType="figure">
<paragraph id="8BF99E72FFF7DC61EF75D050FE1EFC39" blockId="5.[114,587,829,914]" pageId="5" pageNumber="246">
Fig. 3. Habitus of
<taxonomicName id="4C46E5F1FFF7DC61EE44D050FDDDFCFE" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" box="[323,587,829,853]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="aeschyae" status="sp. nov." subGenus="Cacia">
<emphasis id="B9324260FFF7DC61EE44D050FDDDFCFE" box="[323,587,829,853]" italics="true" pageId="5" pageNumber="246">Cacia (Cacia) aeschyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF7DC61EF75D031FF59FCD8" box="[114,207,860,883]" pageId="5" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
: A. Dorsal aspect, B. Ventral aspect, C. Lateral, D. Frons.
</paragraph>
</caption>
<paragraph id="8BF99E72FFF7DC61EF75D0DAFF42FA5C" blockId="5.[114,593,951,1527]" pageId="5" pageNumber="246">
<emphasis id="B9324260FFF7DC61EF75D0DAFE52FC64" bold="true" box="[114,452,951,975]" pageId="5" pageNumber="246">Pronotum and Propleuron</emphasis>
with deep puncturations; pronotal disc slightly raised, broadest at the middle; prothorax densely covered with recumbent yellowish pubescence with few erect yellowish setae arranged in random. Elytra twice longer than wide, with dense puncturations from base towards post-median, finer towards apex. Humeri angled and slightly raised. Elytra broadest at humeri tapering towards apex, with three transverse bands of recumbent yellowish pubescence: a transverse band at the base and humeri, a pre-median band from margin to suture, and thick postmedi- an band towards apex. Elytral margin lined with long erect yellowish setae from the lateral side of humeri to apex. Scutellum triangular covered with recumbent yellowish setae.
</paragraph>
<paragraph id="8BF99E72FFF7DC61EF75D571FDDDF9D9" blockId="5.[114,592,1564,1650]" pageId="5" pageNumber="246">
<emphasis id="B9324260FFF7DC61EF75D571FEEDF99F" bold="true" box="[114,379,1564,1588]" pageId="5" pageNumber="246">Procoxa and metacoxa</emphasis>
are robust, raised, and covered with fine recumbent pubescence with few erect yellowish setae.
</paragraph>
<paragraph id="8BF99E72FFF7DC61EF75D5CBFD3AFC21" blockId="5.[114,154,1702,1726]" lastBlockId="5.[624,1103,637,906]" pageId="5" pageNumber="246">
246
<emphasis id="B9324260FFF7DC61ED77D110FD70FD3E" bold="true" box="[624,742,637,661]" pageId="5" pageNumber="246">Genitalia.</emphasis>
Aedeagus (3.0 mm) recurved when viewed laterally, broadest at pre-median, tapering towards apex, apex slightly blunt; tegmen (
<quantity id="4CBE3397FFF7DC61EC3FD1B4FC06FD59" box="[824,912,729,754]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="246" unit="mm" value="3.5">3.5 mm</quantity>
) longer than aedeagus, parameres four times longer than wide (1.0 mm,
<quantity id="4CBE3397FFF7DC61EC11D07AFCEFFC84" box="[790,889,791,815]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="246" unit="mm" value="0.25">0.25 mm</quantity>
), parallel-sided not meeting from base to apex; apex of parameres lined with long soft erect yellowish setae.
</paragraph>
<paragraph id="8BF99E72FFF7DC61ED77D0DDFC14FC63" blockId="5.[624,898,944,968]" box="[624,898,944,968]" pageId="5" pageNumber="246">
<emphasis id="B9324260FFF7DC61ED77D0DDFC95FC63" bold="true" box="[624,771,944,968]" pageId="5" pageNumber="246">Adult female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF7DC62ED77D080FE87FE8E" lastPageId="6" lastPageNumber="247" pageId="5" pageNumber="246" type="diagnosis">
<paragraph id="8BF99E72FFF7DC62ED77D080FE87FE8E" blockId="5.[624,1103,1005,1643]" lastBlockId="6.[189,662,178,293]" lastPageId="6" lastPageNumber="247" pageId="5" pageNumber="246">
<emphasis id="B9324260FFF7DC61ED77D080FCFAFBAE" bold="true" box="[624,876,1005,1029]" pageId="5" pageNumber="246">Differential diagnosis</emphasis>
. This new species belongs to
<taxonomicName id="4C46E5F1FFF7DC61EDE1D761FCBAFB8F" authorityName="Newman" authorityYear="1842" box="[742,812,1036,1060]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B9324260FFF7DC61EDE1D761FCBAFB8F" box="[742,812,1036,1060]" italics="true" pageId="5" pageNumber="246">Cacia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF7DC61EC2AD760FC27FB8F" box="[813,945,1036,1060]" pageId="5" pageNumber="246" sensu="stricto">sēnsū strictō</taxonomicNameLabel>
for having an apical spine at antennomere III, and no apical spines in IV and VI. This species is close to
<taxonomicName id="4C46E5F1FFF7DC61EDC9D705FB89FB2B" authority="Breuning, 1947" authorityName="Breuning" authorityYear="1947" box="[718,1055,1128,1152]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="B9324260FFF7DC61EDC9D705FCE5FB2B" box="[718,883,1128,1152]" italics="true" pageId="5" pageNumber="246">C. trimaculata</emphasis>
<bibRefCitation id="EFD7E383FFF7DC61EC71D705FB89FB2B" author="Breuning S. von" box="[886,1055,1128,1152]" pageId="5" pageNumber="246" pagination="1 - 68" refId="ref3317" refString="247 Breuning S. von. 1947. Nouvelles formes de Longicornes du Musee de Stockholm. Arkiv for Zoologi, 39 A (6), 1 - 68." type="journal article" year="1947">Breuning, 1947</bibRefCitation>
</taxonomicName>
for having three bands of whitish pubescence at the elytra but can be easily differentiated based on the pronotal bands (
<taxonomicName id="4C46E5F1FFF7DC61ECC4D7A9FBDFFB77" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" box="[963,1097,1220,1244]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="aeschyae" status="sp. nov.">
<emphasis id="B9324260FFF7DC61ECC4D7A9FBDFFB77" box="[963,1097,1220,1244]" italics="true" pageId="5" pageNumber="246">C. aeschyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF7DC61ED77D78EFD5CFB51" box="[624,714,1251,1274]" pageId="5" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
fully covered with yellowish pubescence vs. thick median longitudinal band of whitish pubescence in
<taxonomicName id="4C46E5F1FFF7DC61ECCDD64DFD0FFAFC" authorityName="Breuning" authorityYear="1947" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="B9324260FFF7DC61ECCDD64DFD0FFAFC" italics="true" pageId="5" pageNumber="246">C. trimaculata</emphasis>
</taxonomicName>
). Both species also differ in terms of elytral bands (thinner bands in
<taxonomicName id="4C46E5F1FFF7DC61ECC2D633FBDEFADD" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" box="[965,1096,1374,1398]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="aeschyae" status="sp. nov.">
<emphasis id="B9324260FFF7DC61ECC2D633FBDEFADD" box="[965,1096,1374,1398]" italics="true" pageId="5" pageNumber="246">C. aeschyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF7DC61ED77D610FD5FFA3F" box="[624,713,1405,1428]" pageId="5" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
vs. thick circular bands in
<taxonomicName id="4C46E5F1FFF7DC61EB04D611FD77FA18" authorityName="Breuning" authorityYear="1947" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="triomaculata">
<emphasis id="B9324260FFF7DC61EB04D611FD77FA18" italics="true" pageId="5" pageNumber="246">C. triomaculata</emphasis>
</taxonomicName>
).
<taxonomicName id="4C46E5F1FFF7DC61EDFFD6F6FC34FA18" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" box="[760,930,1435,1459]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="aeschyae" status="sp. nov.">
<emphasis id="B9324260FFF7DC61EDFFD6F6FC34FA18" box="[760,930,1435,1459]" italics="true" pageId="5" pageNumber="246">Cacia aeschyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF7DC61ECADD6F6FC69FA19" box="[938,1023,1435,1458]" pageId="5" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
is distinct for having a body tapering towards apex more robust in
<taxonomicName id="4C46E5F1FFF7DC61EC57D6B5FC6CFA5B" authorityName="Breuning" authorityYear="1947" box="[848,1018,1496,1520]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="triomaculata">
<emphasis id="B9324260FFF7DC61EC57D6B5FC6CFA5B" box="[848,1018,1496,1520]" italics="true" pageId="5" pageNumber="246">C. triomaculata</emphasis>
</taxonomicName>
. Finally, antennomere VI of
<taxonomicName id="4C46E5F1FFF7DC61EC60D69AFB86F9A4" authorityName="Breuning" authorityYear="1947" box="[871,1040,1527,1551]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="triomaculata">
<emphasis id="B9324260FFF7DC61EC60D69AFB86F9A4" box="[871,1040,1527,1551]" italics="true" pageId="5" pageNumber="246">C. triomaculata</emphasis>
</taxonomicName>
covered with whitish pubescence, absent in
<taxonomicName id="4C46E5F1FFF7DC61EB37D57BFD44F9E7" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="aeschyae" status="sp. nov.">
<emphasis id="B9324260FFF7DC61EB37D57BFD44F9E7" italics="true" pageId="5" pageNumber="246">C. aeschyae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF7DC61EDE7D558FCAAF9E7" box="[736,828,1589,1612]" pageId="5" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
This new species also differs from
<taxonomicName id="4C46E5F1FFF7DC62EC14D53EFF65FF61" authority="Hudepohl, 1989" authorityName="Hudepohl" authorityYear="1989" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" lastPageId="6" lastPageNumber="247" order="Coleoptera" pageId="5" pageNumber="246" phylum="Arthropoda" rank="species" species="estrellae">
<emphasis id="B9324260FFF7DC61EC14D53EFC5CF9C0" box="[787,970,1619,1643]" italics="true" pageId="5" pageNumber="246">Cacia estrellae</emphasis>
<bibRefCitation id="EFD7E383FFF7DC62ECDFD53EFF65FF61" author="Hudepohl K. E." lastPageId="6" lastPageNumber="247" pageId="5" pageNumber="246" pagination="473 - 505" refId="ref3674" refString="Hudepohl K. E. 1989. UbersudostasiatischeCerambyciden VI (Coleoptera, Cerambycidae). Entomofauna, 10 (31): 473 - 505." type="journal article" year="1989">Hüdepohl, 1989</bibRefCitation>
</taxonomicName>
which can be found in the same biogeographic region for having a muchtapered body form and no two-toned femora.
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF4DC62EDBCD3DCFCAFFEAC" pageId="6" pageNumber="247" type="etymology">
<paragraph id="8BF99E72FFF4DC62EDBCD3DCFCAFFEAC" blockId="6.[699,1179,177,263]" pageId="6" pageNumber="247">
<emphasis id="B9324260FFF4DC62EDBCD3DCFCA0FF62" bold="true" box="[699,822,177,201]" pageId="6" pageNumber="247">Etymology</emphasis>
. The new species is named after Aeschy Rose Medina, the daughter of the first author.
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF4DC62EDBCD241FC5DFEC8" pageId="6" pageNumber="247" type="distribution">
<paragraph id="8BF99E72FFF4DC62EDBCD241FC5DFEC8" blockId="6.[699,1178,300,355]" pageId="6" pageNumber="247">
<emphasis id="B9324260FFF4DC62EDBCD241FCD0FEEF" bold="true" box="[699,838,300,324]" pageId="6" pageNumber="247">Distribution</emphasis>
.
<collectingCountry id="F351DEE2FFF4DC62EC51D241FC58FEEF" box="[854,974,300,324]" name="Philippines" pageId="6" pageNumber="247">Philippines</collectingCountry>
(Southern Luzon,
<collectingRegion id="49825090FFF4DC62EDBCD226FCEFFEC8" box="[699,889,331,355]" country="Philippines" name="Mindoro Oriental" pageId="6" pageNumber="247">Oriental Mindoro</collectingRegion>
, Baco).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="9D865FA7961F0B74F1F2343183DDCE1C">Two new species ofCaciaNewman (Coleoptera, Cerambycidae, Lamiinae) from the Mindoro Biogeographic Region of the Philippines</mods:title>
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<mods:date id="854A0842904F7E3797B0447E322BDF7A">2023</mods:date>
<mods:detail id="990663230B2B96C5DA634E2CC6871AFC" type="volume">
<mods:number id="E9C7C1358242D94EA482C81271983E45">23</mods:number>
</mods:detail>
<mods:detail id="71DA77D38B0752294F5F47295AD2F031" type="issue">
<mods:number id="AF187B33D0B5F23D33663D7769D6DBAB">2</mods:number>
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<mods:start id="89ED786EE10F0A765B91318581E85071">241</mods:start>
<mods:end id="D7B6704C9259455A1B71947B172F8D29">248</mods:end>
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<mods:location id="5654F5FCC7668AF484D9A990F1903AEF">
<mods:url id="51E1D66D0CFAAB5107F356B8F30647AC">http://dx.doi.org/10.1186/s12879-024-09640-2</mods:url>
</mods:location>
<mods:classification id="9B756CC9148ED5DE974D85B88563B51E">journal article</mods:classification>
<mods:identifier id="CDBF420C804CEB971BC0367005DB2F5A" type="DOI">10.59893/bjc.23(2).010</mods:identifier>
<mods:identifier id="CE2323F08A9208BC91E057F2CB7CB07B" type="ISSN">1407-8619</mods:identifier>
</mods:mods>
<treatment id="03EF2F64FFF3DC67ED77D52DFC16FADD" LSID="urn:lsid:plazi:treatment:03EF2F64FFF3DC67ED77D52DFC16FADD" httpUri="http://treatment.plazi.org/id/03EF2F64FFF3DC67ED77D52DFC16FADD" lastPageId="3" lastPageNumber="244" pageId="1" pageNumber="242">
<subSubSection id="C35CCDF9FFF3DC65ED77D52DFC69F9F3" box="[624,1023,1600,1624]" pageId="1" pageNumber="242" type="nomenclature">
<paragraph id="8BF99E72FFF3DC65ED77D52DFC69F9F3" blockId="1.[624,1023,1600,1655]" box="[624,1023,1600,1624]" pageId="1" pageNumber="242">
<heading id="D0B1291EFFF3DC65ED77D52DFC69F9F3" bold="true" box="[624,1023,1600,1624]" fontSize="10" level="2" pageId="1" pageNumber="242" reason="4">
<emphasis id="B9324260FFF3DC65ED77D52DFC69F9F3" bold="true" box="[624,1023,1600,1624]" pageId="1" pageNumber="242">
<taxonomicName id="4C46E5F1FFF3DC65ED77D52DFC37F9F3" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" box="[624,929,1600,1624]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="242" phylum="Arthropoda" rank="species" species="katrinae" status="sp. nov." subGenus="Ipocregyes">
<emphasis id="B9324260FFF3DC65ED77D52DFC37F9F3" bold="true" box="[624,929,1600,1624]" italics="true" pageId="1" pageNumber="242">Cacia (Ipocregyes) katrinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF3DC65ECAFD52DFC69F9F3" box="[936,1023,1600,1624]" pageId="1" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF3DC65ED77D532FD42F9DC" box="[624,724,1631,1655]" pageId="1" pageNumber="242" type="description">
<paragraph id="8BF99E72FFF3DC65ED77D532FD42F9DC" blockId="1.[624,1023,1600,1655]" box="[624,724,1631,1655]" pageId="1" pageNumber="242">
<emphasis id="B9324260FFF3DC65ED77D532FD42F9DC" bold="true" box="[624,724,1631,1655]" pageId="1" pageNumber="242">
(
<figureCitation id="137D82F7FFF3DC65ED70D532FD5AF9DC" box="[631,716,1631,1655]" captionStart="Fig" captionStartId="2.[189,228,780,804]" captionTargetBox="[201,675,178,729]" captionTargetId="figure-477@2.[189,680,171,774]" captionTargetPageId="2" captionText="Fig. 1. Habitus ofCacia (Ipocregyes) katrinae sp. nov.: A. Dorsal aspect, B. Ventral aspect, C. Frons, D. Lateral aspect." pageId="1" pageNumber="242">Fig. 1A</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF39CEFAFFF0DC66EFBAD061FDA5FCCA" pageId="2" pageNumber="243" startId="2.[189,228,780,804]" targetBox="[201,675,178,729]" targetPageId="2" targetType="figure">
<paragraph id="8BF99E72FFF0DC66EFBAD061FDA5FCCA" blockId="2.[189,668,780,866]" pageId="2" pageNumber="243">
Fig. 1. Habitus of
<taxonomicName id="4C46E5F1FFF0DC66EE83D061FF72FCE8" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="243" phylum="Arthropoda" rank="species" species="katrinae" status="sp. nov." subGenus="Ipocregyes">
<emphasis id="B9324260FFF0DC66EE83D061FF72FCE8" italics="true" pageId="2" pageNumber="243">Cacia (Ipocregyes) katrinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF0DC66EFE8D046FEDCFCE9" box="[239,330,811,834]" pageId="2" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
: A. Dorsal aspect, B. Ventral aspect, C. Frons, D. Lateral aspect.
</paragraph>
</caption>
<subSubSection id="C35CCDF9FFF0DC66EFBAD0EBFDD7FBDE" pageId="2" pageNumber="243" type="materials_examined">
<paragraph id="8BF99E72FFF0DC66EFBAD0EBFEC0FC50" blockId="2.[189,668,902,1019]" pageId="2" pageNumber="243">
<materialsCitation id="3B2E942FFFF0DC66EFBAD0EBFEC0FC50" collectingDate="2017-06" collectionCode="MMCP" collectorName="Collector" country="Philippines" location="Baco" municipality="Southern Luzon" pageId="2" pageNumber="243" specimenCount="1" stateProvince="Mindoro Oriental" typeStatus="holotype">
<emphasis id="B9324260FFF0DC66EFBAD0EBFEC2FC35" bold="true" box="[189,340,902,926]" pageId="2" pageNumber="243">
<typeStatus id="54FD20D0FFF0DC66EFBAD0EBFEC2FC35" box="[189,340,902,926]" pageId="2" pageNumber="243" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
. male:
<collectingCountry id="F351DEE2FFF0DC66EEC8D0EAFDFAFC34" box="[463,620,903,927]" name="Philippines" pageId="2" pageNumber="243">PHILIPPINES</collectingCountry>
<collectingMunicipality id="6B9D0408FFF0DC66EFBAD0C8FEFBFC16" box="[189,365,933,957]" pageId="2" pageNumber="243">Southern Luzon</collectingMunicipality>
,
<collectingRegion id="49825090FFF0DC66EE7DD0C8FDA0FC16" box="[378,566,933,957]" country="Philippines" name="Mindoro Oriental" pageId="2" pageNumber="243">Oriental Mindoro</collectingRegion>
/
<location id="8E99C8A9FFF0DC66ED48D0CBFD10FC16" LSID="urn:lsid:plazi:treatment:03EF2F64FFF3DC67ED77D52DFC16FADD:8E99C8A9FFF0DC66ED48D0CBFD10FC16" box="[591,646,934,957]" country="Philippines" municipality="Southern Luzon" name="Baco" pageId="2" pageNumber="243" stateProvince="Mindoro Oriental">Baco</location>
/
<date id="FFF8B8B2FFF0DC66EFBAD0A9FE86FC77" box="[189,272,964,988]" pageId="2" pageNumber="243" value="2017-06">
<collectingDate id="EFBC415AFFF0DC66EFBAD0A9FE86FC77" box="[189,272,964,988]" pageId="2" pageNumber="243" value="2017-06">vi.2017</collectingDate>
</date>
, loc.
<collectorName id="26B3FBA4FFF0DC66EE49D0A9FE27FC77" box="[334,433,964,988]" pageId="2" pageNumber="243">Collector</collectorName>
leg. /
<collectionCode id="ED5706B7FFF0DC66EEFCD0A9FDD8FC77" box="[507,590,964,988]" pageId="2" pageNumber="243">MMCP</collectionCode>
, print- ed on redcard.
</materialsCitation>
</paragraph>
<paragraph id="8BF99E72FFF0DC66EFBAD74DFDD7FBDE" blockId="2.[189,668,1056,1141]" pageId="2" pageNumber="243">
<materialsCitation id="3B2E942FFFF0DC66EFBAD74DFEEAFBFC" collectionCode="DUBC" pageId="2" pageNumber="243" specimenCount="2" typeStatus="paratype">
<emphasis id="B9324260FFF0DC66EFBAD74DFEC3FB93" bold="true" box="[189,341,1056,1080]" pageId="2" pageNumber="243">
<typeStatus id="54FD20D0FFF0DC66EFBAD74DFEC3FB93" box="[189,341,1056,1080]" pageId="2" pageNumber="243" type="paratype">PARATYPE</typeStatus>
</emphasis>
female: same label as the
<typeStatus id="54FD20D0FFF0DC66EFBAD752FE8DFBFC" box="[189,283,1087,1111]" pageId="2" pageNumber="243" type="holotype">holotype</typeStatus>
/
<collectionCode id="ED5706B7FFF0DC66EE37D752FEEAFBFC" box="[304,380,1087,1111]" pageId="2" pageNumber="243">DUBC</collectionCode>
</materialsCitation>
, printed on red card. Type specimen will be deposited at
<collectionCode id="ED5706B7FFF0DC66EEF8D733FDAAFBDE" box="[511,572,1118,1141]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="2" pageNumber="243">PNM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF0DC67EFBAD7F7FD23FE57" lastPageId="3" lastPageNumber="244" pageId="2" pageNumber="243" type="description">
<paragraph id="8BF99E72FFF0DC66EFBAD7F7FDDBFA03" blockId="2.[189,668,1178,1448]" pageId="2" pageNumber="243">
<emphasis id="B9324260FFF0DC66EFBAD7F7FD83FB19" bold="true" box="[189,533,1178,1202]" pageId="2" pageNumber="243">Description. Measurements:</emphasis>
<typeStatus id="54FD20D0FFF0DC66ED35D7F6FD0AFB18" box="[562,668,1179,1203]" pageId="2" pageNumber="243" type="holotype">Holotype</typeStatus>
male: LB (LH+LP+LE):
<quantity id="4CBE3397FFF0DC66EED7D7D4FDA2FB7A" box="[464,564,1209,1233]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="243" unit="mm" value="11.5">11.5 mm</quantity>
. LH:
<quantity id="4CBE3397FFF0DC66ED73D7D4FF7CFB5B" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="243" unit="mm" value="1.5">1.5 mm</quantity>
. WH: 3.0 mm. LG: 1.0 mm. LL: 1.0 mm. WL: 1.0 mm. LP: 2.0 mm. WP: 3.0 mm. LE: 8.0 mm. WEH:
<quantity id="4CBE3397FFF0DC66EED1D678FDBBFA86" box="[470,557,1301,1325]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="243" unit="mm" value="4.5">4.5 mm</quantity>
.
<typeStatus id="54FD20D0FFF0DC66ED3ED67BFD0AFA85" box="[569,668,1302,1326]" pageId="2" pageNumber="243" type="paratype">Paratype</typeStatus>
female: LB (LH+LP+LE): 12.0 mm. LH:
<quantity id="4CBE3397FFF0DC66EFBAD63EFE81FAC1" box="[189,279,1363,1387]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="243" unit="mm" value="1.5">1.5 mm</quantity>
. WH: 3.0 mm. LG:
<quantity id="4CBE3397FFF0DC66ED02D63EFDC9FAC1" box="[517,607,1363,1387]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="243" unit="mm" value="1.5">1.5 mm</quantity>
. LL: 1.0 mm. WL: 1.0 mm. LP: 2.0 mm. WP:
<quantity id="4CBE3397FFF0DC66EFBAD6FDFE84FA03" box="[189,274,1424,1448]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="243" unit="mm" value="3.5">3.5 mm</quantity>
. LE:
<quantity id="4CBE3397FFF0DC66EE4CD6FDFE36FA03" box="[331,416,1424,1448]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="243" unit="mm" value="8.5">8.5 mm</quantity>
. WEH: 5.0 mm.
</paragraph>
<paragraph id="8BF99E72FFF0DC66EFBAD6A0FF69F9CB" blockId="2.[189,668,1485,1632]" pageId="2" pageNumber="243">
<emphasis id="B9324260FFF0DC66EFBAD6A0FE4AFA4E" bold="true" box="[189,476,1485,1509]" pageId="2" pageNumber="243">Adult male. Teguments</emphasis>
head including frons, genae, and antennae matte black; underside of the body, femora, clypeus, legs, pronotum, and elytra dark brown to black.
</paragraph>
<paragraph id="8BF99E72FFF0DC66EDBCD3DCFBAEFCC7" blockId="2.[699,1179,177,876]" pageId="2" pageNumber="243">
<emphasis id="B9324260FFF0DC66EDBCD3DCFD6EFF62" bold="true" box="[699,760,177,201]" pageId="2" pageNumber="243">Head</emphasis>
twice wider than long. The junction between the frons and vertex is slightly raised with a very shallow longitudinal slit along the midline. Vertex with two parallel white and yellowish bands of pubescence from the base up to the apex of the frons. Genae longer than wide, densely covered with whitish and yellowish recumbent pubescence, with few erect black and brownish setae. Posterior end of genae with a shallow dent covered with whitish recumbent setae and few black erect setae. Eyes matte black, lower eye lobesas long as wide, back of the eyes densely covered with semi-erect brownish setae. The apical margin of the head is lined with long whitish setae. Clypeus light brown, glabrous, and lustrous. Labrum matte black densely covered with whitish and yellowish recumbent setae with few erect black setae. Mandible matte black, base with dense black recumbent setae, lateral side with deep depression covered with white and black setae.
</paragraph>
<paragraph id="8BF99E72FFF0DC66EDBCD0FCFB02FADE" blockId="2.[699,1179,913,1397]" pageId="2" pageNumber="243">
<emphasis id="B9324260FFF0DC66EDBCD0FCFCBEFC02" bold="true" box="[699,808,913,937]" pageId="2" pageNumber="243">Antennae</emphasis>
slightly longer than the body; scape slightly robust, rugose, with apical cicatrix, densely covered with whitish recumbent pubescence with black erect setae; with dense micropunctures. Scape, pedicle, antennomeres III to V are densely covered with black or brownish erect setae, the rest of antennomeres are covered with few erect setae. Antennomeres III and IV with whitish pubescence near the base; antennomere III longer than IV; antennomere V entirely covered with whitish pubescence; tuft of black setae at antennomere IV; antennomeres III to XI cylindrical; antennomeres VI to XI densely covered with fine recumbent pubescence, with few black erect setae.
</paragraph>
<paragraph id="8BF99E72FFF0DC66EDBCD6F7FB0CF9E7" blockId="2.[699,1178,1434,1612]" pageId="2" pageNumber="243">
<emphasis id="B9324260FFF0DC66EDBCD6F7FCAEFA19" bold="true" box="[699,824,1434,1458]" pageId="2" pageNumber="243">Prothorax.</emphasis>
Prothorax slightly rugose, covered with fine black recumbent pubescence, lined with 3 longitudinal bands of whitish and yellowish pubescence as follows: along the midline of pronotum, and a thick band at each lower end of propleura. Prosternum
</paragraph>
<paragraph id="8BF99E72FFF0DC66EB6BD5CBFB02F915" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="243">243</paragraph>
<paragraph id="8BF99E72FFF1DC67EF75D3DFFF7FFF43" blockId="3.[114,593,178,232]" pageId="3" pageNumber="244">densely covered with black and whitish erect setae.</paragraph>
<paragraph id="8BF99E72FFF1DC67EF75D260FE57FC85" blockId="3.[114,593,269,815]" pageId="3" pageNumber="244">
<emphasis id="B9324260FFF1DC67EF75D260FF2CFE8E" bold="true" box="[114,186,269,293]" pageId="3" pageNumber="244">Elytra</emphasis>
. Elyra is longer than wide, parallel-sided, and smoothly rounded at the apex. The humeral angle is slightly raised; the basal third of elytra rugose with fine punctures, finer towards the apex. Lateral side from humerus up to basal third with apical declivity, smoother towards the apex. With three bands of whitish to yellowish recumbent pubescence as follows: a short longitudinal band near the humeral angle, a thick pre-median band from near the lateral margin to the elytral suture, and a thick post-median band lined from the elytral margin to suture interrupted at the middle in elytral margin forming a broken circle (
<figureCitation id="137D82F7FFF1DC67EEE3D1D6FDD4FD78" box="[484,578,699,723]" captionStart="Fig" captionStartId="2.[189,228,780,804]" captionTargetBox="[201,675,178,729]" captionTargetId="figure-477@2.[189,680,171,774]" captionTargetPageId="2" captionText="Fig. 1. Habitus ofCacia (Ipocregyes) katrinae sp. nov.: A. Dorsal aspect, B. Ventral aspect, C. Frons, D. Lateral aspect." pageId="3" pageNumber="244">Fig. 1A</figureCitation>
). Elytral margin lined with erect black setae, apex rounded covered with thick whitish and yellowish recumbent setae.
</paragraph>
<paragraph id="8BF99E72FFF1DC67EF75D039FF37FAB2" blockId="3.[114,593,852,1305]" pageId="3" pageNumber="244">
<emphasis id="B9324260FFF1DC67EF75D039FF31FCC7" bold="true" box="[114,167,852,876]" pageId="3" pageNumber="244">Legs</emphasis>
. Procoxa and metacoxa raised, covered with semi-erect whitish setae. Femora and protibiarobust, covered with fine whitish pubescence with few semi-erect whitish or yellowish setae. Basal half of mesotibia, metatibia, and tarsi are covered with fine whitish semi-erect setae, apical half with black semi-erect setae. Claws dark brown, simple. Mesosternum, mesepisternum, metepisternum, metasternum, mesepimeron and abdominal ventrites are densely covered with whitish and yellowish recumbent pubescence and semi-erect setae; ventrite V is broader than ventrites I to IV individually.
</paragraph>
<paragraph id="8BF99E72FFF1DC67EF75D653FDC6F9C0" blockId="3.[114,593,1342,1643]" pageId="3" pageNumber="244">
<emphasis id="B9324260FFF1DC67EF75D653FF4DFAFD" bold="true" box="[114,219,1342,1366]" pageId="3" pageNumber="244">Genitalia</emphasis>
. Whole system of genitalia recurved when viewed laterally. Aedeagus and tegmen are the same length (4.0 mm); endophallus (8.0 mm) is twice longer than the aedeagus and tegmen individually. Parameres are bottle-shaped, separated at the apex, apex lined with soft long yellowish setae. Aedeagus parallel-sided, broader at midline, lanceolate towards the slightly blunt apex. Basal third of endophallus
</paragraph>
<paragraph id="8BF99E72FFF1DC67EF75D5CBFB86FF43" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1103,178,232]" pageId="3" pageNumber="244">244 broader with thick bands of sclerites and forming two lines towards the midline.</paragraph>
<paragraph id="8BF99E72FFF1DC67ED77D260FD23FE57" blockId="3.[624,1103,269,508]" pageId="3" pageNumber="244">
<emphasis id="B9324260FFF1DC67ED77D260FC80FE8E" bold="true" box="[624,790,269,293]" pageId="3" pageNumber="244">Adult female.</emphasis>
There is no morphological difference between the male and female except for the slight difference in body size and anadditional small band of whitish pubescence at the lateral side of the elytra near the humeral angle. Tegument of the underside of the body is darker in females than in males.
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF1DC67ED77D14CFC02FB34" pageId="3" pageNumber="244" type="diagnosis">
<paragraph id="8BF99E72FFF1DC67ED77D14CFC02FB34" blockId="3.[624,1103,545,1183]" pageId="3" pageNumber="244">
<emphasis id="B9324260FFF1DC67ED77D14CFD72FD92" bold="true" box="[624,740,545,569]" pageId="3" pageNumber="244">Diagnosis.</emphasis>
This new species belongs to sub-species
<emphasis id="B9324260FFF1DC67EDF5D12CFCF1FDF3" box="[754,871,577,600]" italics="true" pageId="3" pageNumber="244">Ipocregyes</emphasis>
for having no apical spines in antennomeres III or IV, antennomere III with a tuft of hairs, and the inner side of antennomere III is not inflated. The closest congener of the new species is
<taxonomicName id="4C46E5F1FFF1DC67EB27D1D6FC5AFD59" authority="Medina et al. 2023" authorityName="Medina" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="manobo">
<emphasis id="B9324260FFF1DC67EB27D1D6FD67FD59" italics="true" pageId="3" pageNumber="244">Cacia manobo</emphasis>
<bibRefCitation id="EFD7E383FFF1DC67EDFCD1B7FC5AFD59" author="Medina M. N. D. &amp; Vitali F. &amp; Barsevskis A." box="[763,972,730,754]" pageId="3" pageNumber="244" refId="ref3700" refString="Medina M. N. D., Vitali F., Barsevskis A. 2023. Catalog of the genus Cacia Newman (Coleoptera, Cerambycidae, Lamiinae) in the Philippines with description of two new species. Zootaxa 5231 (5): 537 - 551. DOI: https: // doi. org / 10.11646 / zootaxa. 5231.5.3" type="book" year="2023">Medina et al. 2023</bibRefCitation>
</taxonomicName>
for having similar body form and elytral bands but can be easily differentiated in terms of the pronotal bands (
<taxonomicName id="4C46E5F1FFF1DC67EC0BD05BFC1CFCE5" box="[780,906,822,846]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="manobo">
<emphasis id="B9324260FFF1DC67EC0BD05BFC1CFCE5" box="[780,906,822,846]" italics="true" pageId="3" pageNumber="244">C. manobo</emphasis>
</taxonomicName>
thickly covered with whitish and yellowish pubescence with V-like black pubescence along the apical median area vs. a thinner band of whitish pubescence along the midline from base to apex in
<taxonomicName id="4C46E5F1FFF1DC67EDC2D0A2FCABFC4C" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" box="[709,829,975,999]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="katrinae" status="sp. nov.">
<emphasis id="B9324260FFF1DC67EDC2D0A2FCABFC4C" box="[709,829,975,999]" italics="true" pageId="3" pageNumber="244">C. katrinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF1DC67EC43D0A2FC0BFC4C" box="[836,925,975,999]" pageId="3" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
). A thick tuft of hairs at antennomere IV is absent in
<taxonomicName id="4C46E5F1FFF1DC67EB37D083FD50FB8F" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="manobo">
<emphasis id="B9324260FFF1DC67EB37D083FD50FB8F" italics="true" pageId="3" pageNumber="244">C. manobo</emphasis>
</taxonomicName>
, present in
<taxonomicName id="4C46E5F1FFF1DC67EC43D761FC2BFB8F" authorityName="Medina &amp; Cabras &amp; Barševskis" authorityYear="2023" box="[836,957,1036,1060]" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="katrinae" status="sp. nov.">
<emphasis id="B9324260FFF1DC67EC43D761FC2BFB8F" box="[836,957,1036,1060]" italics="true" pageId="3" pageNumber="244">C. katrinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201FF1BFFF1DC67ECC3D760FB8EFB8F" box="[964,1048,1037,1060]" pageId="3" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
The new species is also close to
<taxonomicName id="4C46E5F1FFF1DC67ECBAD746FCB4FBCA" authority="Hudepohl, 1989" authorityName="Hudepohl" authorityYear="1989" class="Insecta" family="Cerambycidae" genus="Cacia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="marionae">
<emphasis id="B9324260FFF1DC67ECBAD746FBDEFBE8" box="[957,1096,1067,1091]" italics="true" pageId="3" pageNumber="244">C. marionae</emphasis>
<bibRefCitation id="EFD7E383FFF1DC67ED77D724FCB4FBCA" author="Hudepohl K. E." box="[624,802,1097,1121]" pageId="3" pageNumber="244" pagination="473 - 505" refId="ref3674" refString="Hudepohl K. E. 1989. UbersudostasiatischeCerambyciden VI (Coleoptera, Cerambycidae). Entomofauna, 10 (31): 473 - 505." type="journal article" year="1989">Hüdepohl, 1989</bibRefCitation>
</taxonomicName>
but can be easily distinguished by their differences in the head, pronotal, and elytral bands.
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF1DC67ED77D7A9FD63FAB2" pageId="3" pageNumber="244" type="etymology">
<paragraph id="8BF99E72FFF1DC67ED77D7A9FD63FAB2" blockId="3.[624,1103,1220,1305]" pageId="3" pageNumber="244">
<emphasis id="B9324260FFF1DC67ED77D7A9FD6EFB77" bold="true" box="[624,760,1220,1244]" pageId="3" pageNumber="244">Etymology.</emphasis>
This new species is named after Katrina Barševska, the daughter of the third author.
</paragraph>
</subSubSection>
<subSubSection id="C35CCDF9FFF1DC67ED77D653FC16FADD" pageId="3" pageNumber="244" type="distribution">
<paragraph id="8BF99E72FFF1DC67ED77D653FC16FADD" blockId="3.[624,1102,1342,1398]" pageId="3" pageNumber="244">
<emphasis id="B9324260FFF1DC67ED77D653FC94FAFD" bold="true" box="[624,770,1342,1366]" pageId="3" pageNumber="244">Distribution.</emphasis>
<collectingCountry id="F351DEE2FFF1DC67EC0DD652FC14FAFC" box="[778,898,1343,1367]" name="Philippines" pageId="3" pageNumber="244">Philippines</collectingCountry>
(Southern Luzon,
<collectingRegion id="49825090FFF1DC67ED77D630FCB8FADE" box="[624,814,1373,1397]" country="Philippines" name="Mindoro Oriental" pageId="3" pageNumber="244">Oriental Mindoro</collectingRegion>
, Baco).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,235 @@
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<mods:title id="E63D8AE2CD653901E737EF40D7169AFB">A new species of the genus Apocyrtus Erichson, 1834 (Coleoptera: Curculionidae: Pachyrhynchini) from the Marinduque Island, Philippines</mods:title>
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<mods:date id="507B5330069D5A2689FD38CB87F3555A">2022</mods:date>
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<treatment id="03F587E5B230FFA0FFB9FEB8FF1493F3" LSID="urn:lsid:plazi:treatment:03F587E5B230FFA0FFB9FEB8FF1493F3" httpUri="http://treatment.plazi.org/id/03F587E5B230FFA0FFB9FEB8FF1493F3" lastPageId="3" lastPageNumber="432" pageId="1" pageNumber="430">
<subSubSection id="C3466578B230FFA2FFB9FEB8FE339027" box="[113,501,324,348]" pageId="1" pageNumber="430" type="nomenclature">
<paragraph id="8BE336F3B230FFA2FFB9FEB8FE339027" blockId="1.[113,501,324,380]" box="[113,501,324,348]" pageId="1" pageNumber="430">
<heading id="D0AB819FB230FFA2FFB9FEB8FE339027" bold="true" box="[113,501,324,348]" fontSize="10" level="2" pageId="1" pageNumber="430" reason="4">
<emphasis id="B928EAE1B230FFA2FFB9FEB8FE339027" bold="true" box="[113,501,324,348]" pageId="1" pageNumber="430">
<taxonomicName id="4C5C4D70B230FFA2FFB9FEB8FE5F9027" ID-CoL="B7QPF" authorityName="Rukmane-Brbale" authorityYear="2022" box="[113,409,324,348]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="marinduquensis" status="sp. nov.">
<emphasis id="B928EAE1B230FFA2FFB9FEB8FE5F9027" bold="true" box="[113,409,324,348]" italics="true" pageId="1" pageNumber="430">Apocyrtus marinduquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B579AB230FFA2FE57FEB8FE339027" box="[415,501,324,348]" pageId="1" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3466578B230FFA2FFB9FE99FE9A9007" box="[113,348,357,380]" pageId="1" pageNumber="430" type="description">
<paragraph id="8BE336F3B230FFA2FFB9FE99FE9A9007" blockId="1.[113,501,324,380]" box="[113,348,357,380]" pageId="1" pageNumber="430">
(
<figureCitation id="13672A76B230FFA2FFB1FE99FF2A9007" box="[121,236,357,380]" captionStart="Figure 1" captionStartId="1.[113,183,1633,1656]" captionTargetBox="[116,1133,810,1601]" captionTargetId="figure-229@1.[113,1134,808,1602]" captionTargetPageId="1" captionText="Figure 1. Dorsal view of Apocyrtus species. A Apocyrtus auroraensis Rukmane­Brbale, 2021. B Apocyrtus marinduquensis sp. nov., male. C Apocyrtus marinduquensis sp. nov., female." pageId="1" pageNumber="430">Fig. 1BC</figureCitation>
,
<figureCitation id="13672A76B230FFA2FF3EFE99FE939007" box="[246,341,357,380]" captionStart="Figure 2" captionStartId="2.[151,221,1601,1624]" captionTargetBox="[223,1104,785,1571]" captionTargetId="figure-314@2.[219,1107,783,1572]" captionTargetPageId="2" captionText="Figure 2. A, CE. Genitalia of Apocyrtus marinduquensis sp. nov. A aedeagus in lateral view, C sternite IX in dorsal view, D ovipositor in dorsal view, E spermatheca; B aedeagus of Apocyrtus auroraensis in dorsal view." pageId="1" pageNumber="430">2A, CE</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3466578B230FFA2FFB9FE60FE27938A" pageId="1" pageNumber="430" type="materials_examined">
<paragraph id="8BE336F3B230FFA2FFB9FE60FEA9932C" blockId="1.[113,605,412,599]" pageId="1" pageNumber="430">
<emphasis id="B928EAE1B230FFA2FFB9FE60FEDC90CF" bold="true" box="[113,282,412,436]" pageId="1" pageNumber="430">Type material.</emphasis>
<materialsCitation id="3B343CAEB230FFA2FEEBFE60FEA9932C" collectingDate="2021-06" country="Philippines" location="Buenavista" pageId="1" pageNumber="430" specimenCount="2" stateProvince="Marinduque" typeStatus="holotype">
<typeStatus id="54E78851B230FFA2FEEBFE60FE4C90C8" box="[291,394,412,435]" pageId="1" pageNumber="430" type="holotype">Holotype</typeStatus>
. Male. “
<collectingCountry id="F34B7663B230FFA2FE2FFE60FF6590AF" name="Philippines" pageId="1" pageNumber="430">PHILIPPINES</collectingCountry>
/
<collectingRegion id="4998F811B230FFA2FF7BFE41FEF390AF" box="[179,309,445,468]" country="Philippines" name="Marinduque" pageId="1" pageNumber="430">Marinduque</collectingRegion>
,
<location id="8E836028B230FFA2FEF5FE41FE7690AF" LSID="urn:lsid:plazi:treatment:03F587E5B230FFA0FFB9FEB8FF1493F3:8E836028B230FFA2FEF5FE41FE7690AF" box="[317,432,445,468]" country="Philippines" name="Buenavista" pageId="1" pageNumber="430" stateProvince="Marinduque">Buenavista</location>
/
<date id="FFE21033B230FFA2FE77FE41FDDC90AF" box="[447,538,445,468]" pageId="1" pageNumber="430" value="2021-06">
<collectingDate id="EFA6E9DBB230FFA2FE77FE41FDDC90AF" box="[447,538,445,468]" pageId="1" pageNumber="430" value="2021-06">VI. 2021</collectingDate>
</date>
/ local collector leg.” (typed on a white rectangular label); “
<typeStatus id="54E78851B230FFA2FF0AFE03FE96936D" box="[194,336,511,534]" pageId="1" pageNumber="430" type="holotype">HOLOTYPE</typeStatus>
/ Male /
<taxonomicName id="4C5C4D70B230FFA2FE6FFE03FF1A934C" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="marinduquensis">
<emphasis id="B928EAE1B230FFA2FE6FFE03FF1A934C" italics="true" pageId="1" pageNumber="430">Apocyrtus marinduquensis</emphasis>
</taxonomicName>
Rukmane­Brbale, 2022” (typed on a red rectangular label).
</materialsCitation>
</paragraph>
<paragraph id="8BE336F3B230FFA2FFB9FD8BFE27938A" blockId="1.[113,605,631,754]" pageId="1" pageNumber="430">
<emphasis id="B928EAE1B230FFA2FFB9FD8BFF3393F4" bold="true" box="[113,245,631,655]" pageId="1" pageNumber="430">
<typeStatus id="54E78851B230FFA2FFB9FD8BFF2593F4" box="[113,227,631,655]" pageId="1" pageNumber="430" type="paratype">Paratypes</typeStatus>
(
</emphasis>
3, 1): All with the same white label as
<typeStatus id="54E78851B230FFA2FF06FD65FEE893CB" box="[206,302,665,688]" pageId="1" pageNumber="430" type="holotype">holotype</typeStatus>
, with additional red rectangular label: “
<typeStatus id="54E78851B230FFA2FF3CFD45FEBC93AB" box="[244,378,697,720]" pageId="1" pageNumber="430" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C5C4D70B230FFA2FE43FD46FF079389" authorityName="Rukmane-Brbale" authorityYear="2022" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="marinduquensis">
<emphasis id="B928EAE1B230FFA2FE43FD46FF079389" italics="true" pageId="1" pageNumber="430">Apocyrtus marinduquensis</emphasis>
</taxonomicName>
Rukmane­Brbale, 2022”.
</paragraph>
</subSubSection>
<subSubSection id="C3466578B230FFA2FD4BFF66FC2A91C9" box="[643,1004,154,178]" pageId="1" pageNumber="430" type="distribution">
<paragraph id="8BE336F3B230FFA2FD4BFF66FC2A91C9" blockId="1.[643,1004,154,178]" box="[643,1004,154,178]" pageId="1" pageNumber="430">
<emphasis id="B928EAE1B230FFA2FD4BFF66FCD391C9" bold="true" box="[643,789,154,178]" pageId="1" pageNumber="430">Distribution.</emphasis>
<collectingRegion id="4998F811B230FFA2FCD3FF67FC5891C9" box="[795,926,155,178]" country="Philippines" name="Marinduque" pageId="1" pageNumber="430">Marinduque</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C3466578B230FFA0FD4BFF2DFE089369" lastPageId="3" lastPageNumber="432" pageId="1" pageNumber="430" type="description">
<paragraph id="8BE336F3B230FFA2FD4BFF2DFB9B90B3" blockId="1.[643,1137,209,456]" pageId="1" pageNumber="430">
<emphasis id="B928EAE1B230FFA2FD4BFF2DFCCD9192" bold="true" box="[643,779,209,233]" pageId="1" pageNumber="430">Description.</emphasis>
Male. Measurements: LB: 8.0­ 8.9 (
<typeStatus id="54E78851B230FFA2FD42FF0DFD2F9073" box="[650,745,241,264]" pageId="1" pageNumber="430" type="holotype">holotype</typeStatus>
8.4, mean 8.48); LP: 2.2­ 2.4 (
<typeStatus id="54E78851B230FFA2FBFDFF0DFD779053" pageId="1" pageNumber="430" type="holotype">holotype</typeStatus>
2.2, mean 2.3); LE: 5.3­ 5.8 (
<typeStatus id="54E78851B230FFA2FC29FEEDFB869053" box="[993,1088,273,296]" pageId="1" pageNumber="430" type="holotype">holotype</typeStatus>
5.6, mean 5.58); LR: 1.7­ 1.8 (
<typeStatus id="54E78851B230FFA2FC53FECDFC3C9033" box="[923,1018,305,328]" pageId="1" pageNumber="430" type="holotype">holotype</typeStatus>
1.8, mean 1.78); WP: 2.6­ 2.8 (
<typeStatus id="54E78851B230FFA2FC91FEADFC7E9013" box="[857,952,337,360]" pageId="1" pageNumber="430" type="holotype">holotype</typeStatus>
2.6, mean 2.75); WE: 4.7­ 4.9 (
<typeStatus id="54E78851B230FFA2FCC5FE8DFCAC90F3" box="[781,874,369,392]" pageId="1" pageNumber="430" type="holotype">holotype</typeStatus>
4.7, mean 4.8); WR: 1.2­ 1.4 (
<typeStatus id="54E78851B230FFA2FD67FE6DFCCA90D3" box="[687,780,401,424]" pageId="1" pageNumber="430" type="holotype">holotype</typeStatus>
1.3, mean 1.3). N=4 for all measurements. Dorsal habitus as shown in
<figureCitation id="13672A76B230FFA2FBCFFE4DFB9F90B3" box="[1031,1113,433,456]" captionStart="Figure 1" captionStartId="1.[113,183,1633,1656]" captionTargetBox="[116,1133,810,1601]" captionTargetId="figure-229@1.[113,1134,808,1602]" captionTargetPageId="1" captionText="Figure 1. Dorsal view of Apocyrtus species. A Apocyrtus auroraensis Rukmane­Brbale, 2021. B Apocyrtus marinduquensis sp. nov., male. C Apocyrtus marinduquensis sp. nov., female." pageId="1" pageNumber="430">Fig. 1B</figureCitation>
.
</paragraph>
<paragraph id="8BE336F3B230FFA2FD4BFE14FBA99324" blockId="1.[643,1135,488,607]" pageId="1" pageNumber="430">Body except abdomen black, strongly shiny. Elytra and underside reddish. Markings of shiny, metallic green, yellow or gold round to recumbent scales on elytra, underside and prothorax.</paragraph>
<paragraph id="8BE336F3B230FFA1FD4BFD83FDE29347" blockId="1.[643,1134,639,758]" lastBlockId="2.[151,643,155,572]" lastPageId="2" lastPageNumber="431" pageId="1" pageNumber="430">Head nearly smooth, slightly pubescent, without scally margings laterally; forehead wrinkled between eyes, bulging at basal part, impressed at apical part, 1.5 times as wide as eye width; eyes small, prominent from the outline of the head; peak just in the middle; rostrum with moderate pubescence, in dorsal contour straigth, longer than wide, LR/WR: 1.38; dorsally with transverse groove just before the base of each eye, and longitudinal medial groove from middle of rostrum to basal 1/3 of forehead; antennae with basal antennomere three times as long as wide, 1.5 times longer than antennomere II; antennomere II 1.5 times as long as wide, 1.5 times longer than antennomere III; antennomeres III VII subequal in length and width, as long as wide; club two times as long as wide.</paragraph>
<caption id="DF23667BB230FFA2FFB9F99DFBA497E3" pageId="1" pageNumber="430" startId="1.[113,183,1633,1656]" targetBox="[116,1133,810,1601]" targetPageId="1" targetType="figure">
<paragraph id="8BE336F3B230FFA2FFB9F99DFBA497E3" blockId="1.[113,1135,1632,1688]" pageId="1" pageNumber="430">
Figure 1. Dorsal view of
<taxonomicName id="4C5C4D70B230FFA2FEBFF99DFD939703" box="[375,597,1632,1656]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B928EAE1B230FFA2FEBFF99DFE259703" box="[375,483,1633,1656]" italics="true" pageId="1" pageNumber="430">Apocyrtus</emphasis>
species.
<emphasis id="B928EAE1B230FFA2FD8AF99CFD939703" bold="true" box="[578,597,1632,1656]" pageId="1" pageNumber="430">A</emphasis>
</taxonomicName>
<taxonomicName id="4C5C4D70B230FFA2FDA5F99DFC989703" authorityName="Rukmane-Bārbale" authorityYear="2021" box="[621,862,1633,1656]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="auroraensis">
<emphasis id="B928EAE1B230FFA2FDA5F99DFC989703" box="[621,862,1633,1656]" italics="true" pageId="1" pageNumber="430">Apocyrtus auroraensis</emphasis>
</taxonomicName>
Rukmane­Brbale, 2021.
<emphasis id="B928EAE1B230FFA2FFB9F97CFF4597E3" bold="true" box="[113,131,1664,1688]" pageId="1" pageNumber="430">B</emphasis>
<taxonomicName id="4C5C4D70B230FFA2FF56F97DFE7C97E3" authorityName="Rukmane-Brbale" authorityYear="2022" box="[158,442,1665,1688]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="marinduquensis" status="sp. nov.">
<emphasis id="B928EAE1B230FFA2FF56F97DFE7C97E3" box="[158,442,1665,1688]" italics="true" pageId="1" pageNumber="430">Apocyrtus marinduquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B579AB230FFA2FE09F97DFDD597E3" box="[449,531,1665,1688]" pageId="1" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
, male.
<emphasis id="B928EAE1B230FFA2FDA9F97CFDB297E3" bold="true" box="[609,628,1664,1688]" pageId="1" pageNumber="430">C</emphasis>
<taxonomicName id="4C5C4D70B230FFA2FD47F97DFC6A97E3" authorityName="Rukmane-Brbale" authorityYear="2022" box="[655,940,1665,1688]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="marinduquensis" status="sp. nov.">
<emphasis id="B928EAE1B230FFA2FD47F97DFC6A97E3" box="[655,940,1665,1688]" italics="true" pageId="1" pageNumber="430">Apocyrtus marinduquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B579AB230FFA2FC7BF97DFBC297E3" box="[947,1028,1665,1688]" pageId="1" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
, female.
</paragraph>
</caption>
<paragraph id="8BE336F3B233FFA1FF5FFDA0FC979069" blockId="2.[151,643,604,726]" lastBlockId="2.[680,1172,155,274]" pageId="2" pageNumber="431">Prothorax strongly rugose, wider than long, WP/ LP: 1.18, without grooves, mingled with single metallic scales chaotically dispersed; in dorsal contour slightly impressed at subapical part, widest just after the middle, decreased to subbasal part and straight to basal margin; anterior edge dorsally straight, posterior edge strongly curved apically.</paragraph>
<paragraph id="8BE336F3B233FFA1FD60FECDFCA09093" blockId="2.[680,1173,305,488]" pageId="2" pageNumber="431">
Elytra subspherical, longer than wide, LE/WE: 1.19, LE/LP: 2.55, WE/WP: 1.81, with moderate rugosity, without clearly expressed intervals, with markings of chaotically dispersed single scales; widest just at the middle. Genitalia as shown in
<figureCitation id="13672A76B233FFA1FCC7FE2DFCA59093" box="[783,867,465,488]" captionStart="Figure 2" captionStartId="2.[151,221,1601,1624]" captionTargetBox="[223,1104,785,1571]" captionTargetId="figure-314@2.[219,1107,783,1572]" captionTargetPageId="2" captionText="Figure 2. A, CE. Genitalia of Apocyrtus marinduquensis sp. nov. A aedeagus in lateral view, C sternite IX in dorsal view, D ovipositor in dorsal view, E spermatheca; B aedeagus of Apocyrtus auroraensis in dorsal view." pageId="2" pageNumber="431">Fig. 2A</figureCitation>
.
</paragraph>
<paragraph id="8BE336F3B233FFA1FD60FDF4FCFC93A4" blockId="2.[680,1172,520,735]" pageId="2" pageNumber="431">
Female. Measurements: LB: 10.1; LP: 2.2; LE: 7.1; LR: 1.7; WP: 2.7; WE: 5.3; WR: 1.2. N = 1 for all measurements. Elytra with strongly extended apex, in dorsal contour with impression along basal 1/2, otherwise as in males. Dorsal habitus as shown in
<figureCitation id="13672A76B233FFA1FCBFFD54FC0193C4" box="[887,967,680,703]" captionStart="Figure 1" captionStartId="1.[113,183,1633,1656]" captionTargetBox="[116,1133,810,1601]" captionTargetId="figure-229@1.[113,1134,808,1602]" captionTargetPageId="1" captionText="Figure 1. Dorsal view of Apocyrtus species. A Apocyrtus auroraensis Rukmane­Brbale, 2021. B Apocyrtus marinduquensis sp. nov., male. C Apocyrtus marinduquensis sp. nov., female." pageId="2" pageNumber="431">Fig. 1C</figureCitation>
. Genitalia as shown in
<figureCitation id="13672A76B233FFA1FD0CFD34FCF393A4" box="[708,821,712,735]" captionStart="Figure 2" captionStartId="2.[151,221,1601,1624]" captionTargetBox="[223,1104,785,1571]" captionTargetId="figure-314@2.[219,1107,783,1572]" captionTargetPageId="2" captionText="Figure 2. A, CE. Genitalia of Apocyrtus marinduquensis sp. nov. A aedeagus in lateral view, C sternite IX in dorsal view, D ovipositor in dorsal view, E spermatheca; B aedeagus of Apocyrtus auroraensis in dorsal view." pageId="2" pageNumber="431">Fig. 2DE</figureCitation>
.
</paragraph>
<caption id="DF23667BB233FFA1FF5FF9BDFDED97E3" pageId="2" pageNumber="431" startId="2.[151,221,1601,1624]" targetBox="[223,1104,785,1571]" targetPageId="2" targetType="figure">
<paragraph id="8BE336F3B233FFA1FF5FF9BDFDED97E3" blockId="2.[151,1172,1600,1688]" pageId="2" pageNumber="431">
Figure 2.
<emphasis id="B928EAE1B233FFA1FF36F9BCFE979723" bold="true" box="[254,337,1600,1624]" pageId="2" pageNumber="431">A, CE</emphasis>
. Genitalia of
<taxonomicName id="4C5C4D70B233FFA1FE2CF9BDFCC49723" authorityName="Rukmane-Brbale" authorityYear="2022" box="[484,770,1601,1624]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="marinduquensis" status="sp. nov.">
<emphasis id="B928EAE1B233FFA1FE2CF9BDFCC49723" box="[484,770,1601,1624]" italics="true" pageId="2" pageNumber="431">Apocyrtus marinduquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B579AB233FFA1FCCFF9BDFC9C9723" box="[775,858,1601,1624]" pageId="2" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B928EAE1B233FFA1FCA8F9BCFCB59723" bold="true" box="[864,883,1600,1624]" pageId="2" pageNumber="431">A</emphasis>
aedeagus in lateral view,
<emphasis id="B928EAE1B233FFA1FF5FF99CFF6C9703" bold="true" box="[151,170,1632,1656]" pageId="2" pageNumber="431">C</emphasis>
sternite IX in dorsal view,
<emphasis id="B928EAE1B233FFA1FE2AF99CFE339703" bold="true" box="[482,501,1632,1656]" pageId="2" pageNumber="431">D</emphasis>
ovipositor in dorsal view,
<emphasis id="B928EAE1B233FFA1FCEFF99CFCFF9703" bold="true" box="[807,825,1632,1656]" pageId="2" pageNumber="431">E</emphasis>
spermatheca;
<emphasis id="B928EAE1B233FFA1FC20F99CFC3C9703" bold="true" box="[1000,1018,1632,1656]" pageId="2" pageNumber="431">B</emphasis>
aedeagus of
<taxonomicName id="4C5C4D70B233FFA1FF5FF97DFE4F97E3" authorityName="Rukmane-Bārbale" authorityYear="2021" box="[151,393,1665,1688]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="auroraensis">
<emphasis id="B928EAE1B233FFA1FF5FF97DFE4F97E3" box="[151,393,1665,1688]" italics="true" pageId="2" pageNumber="431">Apocyrtus auroraensis</emphasis>
</taxonomicName>
in dorsal view.
</paragraph>
</caption>
<paragraph id="8BE336F3B232FFA0FFB9FF66FD9A91A9" blockId="3.[113,605,154,530]" pageId="3" pageNumber="432">
<emphasis id="B928EAE1B232FFA0FFB9FF66FE9291C9" bold="true" box="[113,340,154,178]" pageId="3" pageNumber="432">Differential analyses.</emphasis>
The new species is similar to
<taxonomicName id="4C5C4D70B232FFA0FF5AFF47FEFC91A9" authorityName="Rukmane-Bārbale" authorityYear="2021" box="[146,314,187,210]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="auroraensis">
<emphasis id="B928EAE1B232FFA0FF5AFF47FEFC91A9" box="[146,314,187,210]" italics="true" pageId="3" pageNumber="432">A. auroraensis</emphasis>
</taxonomicName>
Rukmane­Brbale, 2021
</paragraph>
<paragraph id="8BE336F3B232FFA0FFB9FF27FE089369" blockId="3.[113,605,154,530]" pageId="3" pageNumber="432">
(
<figureCitation id="13672A76B232FFA0FFB1FF27FF0A9189" box="[121,204,219,242]" captionStart="Figure 1" captionStartId="1.[113,183,1633,1656]" captionTargetBox="[116,1133,810,1601]" captionTargetId="figure-229@1.[113,1134,808,1602]" captionTargetPageId="1" captionText="Figure 1. Dorsal view of Apocyrtus species. A Apocyrtus auroraensis Rukmane­Brbale, 2021. B Apocyrtus marinduquensis sp. nov., male. C Apocyrtus marinduquensis sp. nov., female." pageId="3" pageNumber="432">Fig. 1A</figureCitation>
) by body colour and green scales on body, but is easily distinguishable by the following characters: 1) elytra of
<taxonomicName id="4C5C4D70B232FFA0FE5CFEE7FDED9049" authorityName="Rukmane-Bārbale" authorityYear="2021" box="[404,555,283,306]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="auroraensis">
<emphasis id="B928EAE1B232FFA0FE5CFEE7FDED9049" box="[404,555,283,306]" italics="true" pageId="3" pageNumber="432">A. auroraensis</emphasis>
</taxonomicName>
with longitudinal hollows in all lenght that lack rugose elytra of
<taxonomicName id="4C5C4D70B232FFA0FF1FFEA7FE649009" authorityName="Rukmane-Brbale" authorityYear="2022" box="[215,418,347,370]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="marinduquensis" status="sp. nov.">
<emphasis id="B928EAE1B232FFA0FF1FFEA7FE649009" box="[215,418,347,370]" italics="true" pageId="3" pageNumber="432">A. marinduquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B579AB232FFA0FE64FEA7FDC69009" box="[428,512,347,370]" pageId="3" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
; 2) posterior edge of prothorax more strongly curved apically in
<taxonomicName id="4C5C4D70B232FFA0FF29FE67FE6590C9" authorityName="Rukmane-Brbale" authorityYear="2022" box="[225,419,411,434]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="marinduquensis" status="sp. nov.">
<emphasis id="B928EAE1B232FFA0FF29FE67FE6590C9" box="[225,419,411,434]" italics="true" pageId="3" pageNumber="432">A. marinduquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B579AB232FFA0FE6FFE67FE3390C9" box="[423,501,411,434]" pageId="3" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
; 3) antennae with longer basal antennomere and wider club in
<taxonomicName id="4C5C4D70B232FFA0FF77FE27FE429089" authorityName="Rukmane-Brbale" authorityYear="2022" box="[191,388,475,498]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="marinduquensis" status="sp. nov.">
<emphasis id="B928EAE1B232FFA0FF77FE27FE429089" box="[191,388,475,498]" italics="true" pageId="3" pageNumber="432">A. marinduquensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B579AB232FFA0FE42FE27FE1F9089" box="[394,473,475,498]" pageId="3" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
; 4) different shape of aedegal body (
<figureCitation id="13672A76B232FFA0FEA5FE07FE079369" box="[365,449,507,530]" captionStart="Figure 2" captionStartId="2.[151,221,1601,1624]" captionTargetBox="[223,1104,785,1571]" captionTargetId="figure-314@2.[219,1107,783,1572]" captionTargetPageId="2" captionText="Figure 2. A, CE. Genitalia of Apocyrtus marinduquensis sp. nov. A aedeagus in lateral view, C sternite IX in dorsal view, D ovipositor in dorsal view, E spermatheca; B aedeagus of Apocyrtus auroraensis in dorsal view." pageId="3" pageNumber="432">Fig. 2B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3466578B232FFA0FFB9FDCDFF1493F3" pageId="3" pageNumber="432" type="etymology">
<paragraph id="8BE336F3B232FFA0FFB9FDCDFF1493F3" blockId="3.[113,605,561,648]" pageId="3" pageNumber="432">
<emphasis id="B928EAE1B232FFA0FFB9FDCDFF349332" bold="true" box="[113,242,561,585]" pageId="3" pageNumber="432">Etymology.</emphasis>
The species name is dedicated to geographic locality where species is originally reported.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="7FD41F6622F67AE9A842711F46BD0552">New synonymy, combinations and country records in Carabidae Latreille, 1802 (Coleoptera)</mods:title>
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<paragraph id="8BE036D9200B6757B0D5FA8AFDF6104D" blockId="2.[151,575,1308,1332]" box="[151,575,1308,1332]" pageId="2" pageNumber="481">
<heading id="D0A881B5200B6757B0D5FA8AFDF6104D" bold="true" box="[151,575,1308,1332]" fontSize="10" level="2" pageId="2" pageNumber="481" reason="4">
<taxonomicName id="4C5F4D5A200B6757B0D5FA8AFDF6104D" ID-CoL="76GGM" authority="(Reitter, 1897)" baseAuthorityName="Reitter" baseAuthorityYear="1897" box="[151,575,1308,1332]" class="Insecta" family="Carabidae" genus="Penetretus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="481" phylum="Arthropoda" rank="species" species="andalusicus">
<emphasis id="B92BEACB200B6757B0D5FA8AFDF6104D" bold="true" box="[151,575,1308,1332]" pageId="2" pageNumber="481">
<emphasis id="B92BEACB200B6757B0D5FA8AFE5E104D" bold="true" box="[151,407,1308,1332]" italics="true" pageId="2" pageNumber="481">Penetretus andalusicus</emphasis>
(Reitter, 1897)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3456552200B6757B0D5FAC6FB5C102A" pageId="2" pageNumber="481" type="reference_group">
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<emphasis id="B92BEACB200B6757B0D5FAC6FE9E101E" box="[151,343,1360,1383]" italics="true" pageId="2" pageNumber="481">Penetretus imitator</emphasis>
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<taxonomicNameLabel id="A21857B0200B6757B27FFAC6FD4A1011" box="[573,643,1360,1384]" pageId="2" pageNumber="481">syn. n.</taxonomicNameLabel>
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</paragraph>
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<materialsCitation id="3B373C84200B6757B0D5FA12FB58102A" country="Spain" county="Study" location="Species" municipality="Sierra Nevada mountains" pageId="2" pageNumber="481" specimenCount="1">
<location id="8E806002200B6757B0D5FA12FF2C10E2" LSID="urn:lsid:plazi:treatment:03F687CF200B6757B0D5FA8AFB5C102A:8E806002200B6757B0D5FA12FF2C10E2" box="[151,229,1412,1435]" country="Spain" county="Study" municipality="Sierra Nevada mountains" name="Species" pageId="2" pageNumber="481">Species</location>
was described by single male from
<location id="8E806002200B6757B20AFA12FF2910C2" LSID="urn:lsid:plazi:treatment:03F687CF200B6757B0D5FA8AFB5C102A:8E806002200B6757B20AFA12FF2910C2" country="Spain" county="Study" municipality="Sierra Nevada mountains" name="Cerro Pelado" pageId="2" pageNumber="481">Cerro Pelado</location>
(
<collectingMunicipality id="6B84ACA3200B6757B0B0FA32FDC710C2" box="[242,526,1444,1467]" pageId="2" pageNumber="481">Sierra Nevada mountains</collectingMunicipality>
,
<collectingCountry id="F3487649200B6757B25EFA32FD9210C2" box="[540,603,1444,1467]" name="Spain" pageId="2" pageNumber="481">Spain</collectingCountry>
) as species closely related to
<taxonomicName id="4C5F4D5A200B6757B1D5FA52FF101082" ID-CoL="76GGM" authority="(Reitter, 1897)" baseAuthorityName="Reitter" baseAuthorityYear="1897" class="Insecta" family="Carabidae" genus="Penetretus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="481" phylum="Arthropoda" rank="species" species="andalusicus">
<emphasis id="B92BEACB200B6757B1D5FA52FDE310A2" box="[407,554,1476,1499]" italics="true" pageId="2" pageNumber="481">P. andalusicus</emphasis>
(Reitter, 1897)
</taxonomicName>
.
<collectingCounty id="62814E55200B6757B0A4FA72FEED1082" box="[230,292,1508,1531]" pageId="2" pageNumber="481">Study</collectingCounty>
of extensive material, collected by author during
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locality and other regions of Sierra Navada, revealed that both species dont show differences in external features, aedeagus and inflated endophallus structure.All the distinguishing features indicated in the description are within the limits of individual variability. New synonymy is proposed here
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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<document id="45E76A475220F3DA68CFFF44B628B220" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722656488740" checkinUser="felipe" docAuthor="Barševskis, Arvīds" docDate="2009" docId="0D4E87E58D74FFC91B69A6CCFB26FB04" docLanguage="en" docName="BalticJColeopterol.9.1.1-4.pdf" docOrigin="Baltic Journal of Coleopterology 9 (1)" docStyle="DocumentStyle:DECCBB8D96F758554CCC8A51AE82E69F.1:BalticJColeop.2007-2011.journal_article" docStyleId="DECCBB8D96F758554CCC8A51AE82E69F" docStyleName="BalticJColeop.2007-2011.journal_article" docStyleVersion="1" docTitle="Notiophilus anichtchenkoi Barševskis, 2009, sp. n." docType="treatment" docVersion="2" lastPageNumber="3" masterDocId="F177FF9D8D74FFCB181FA30BFFE2FFF3" masterDocTitle="Notiophilus anichtchenkoi sp. n. (Coleoptera: Carabidae) - a new ground beetle species from China" masterLastPageNumber="4" masterPageNumber="1" pageNumber="1" updateTime="1722690798507" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="91980E7110B98336EC40B886B2DB8629">Notiophilus anichtchenkoi sp. n. (Coleoptera: Carabidae) - a new ground beetle species from China</mods:title>
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<mods:date id="D80CEEAE7A7D7527A05BF51962BD01B3">2009</mods:date>
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<treatment id="0D4E87E58D74FFC91B69A6CCFB26FB04" ID-DOI="http://doi.org/10.5281/zenodo.13199461" ID-Zenodo-Dep="13199461" LSID="urn:lsid:plazi:treatment:0D4E87E58D74FFC91B69A6CCFB26FB04" httpUri="http://treatment.plazi.org/id/0D4E87E58D74FFC91B69A6CCFB26FB04" lastPageId="2" lastPageNumber="3" pageId="0" pageNumber="1">
<subSubSection id="CDFD65788D74FFCB1B69A6CCFB3FFA2C" box="[886,1245,1479,1503]" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="855836F38D74FFCB1B69A6CCFB3FFA2C" blockId="0.[886,1245,1479,1503]" box="[886,1245,1479,1503]" pageId="0" pageNumber="1">
<heading id="DE10819F8D74FFCB1B69A6CCFB3FFA2C" bold="true" box="[886,1245,1479,1503]" fontSize="10" level="2" pageId="0" pageNumber="1" reason="4">
<emphasis id="B793EAE18D74FFCB1B69A6CCFB3FFA2C" bold="true" box="[886,1245,1479,1503]" pageId="0" pageNumber="1">
<taxonomicName id="42E74D708D74FFCB1B69A6CCFB7CFA2C" ID-CoL="6SBXB" authorityName="Barševskis" authorityYear="2009" box="[886,1182,1479,1503]" class="Insecta" family="Carabidae" genus="Notiophilus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anichtchenkoi" status="sp. nov.">
<emphasis id="B793EAE18D74FFCB1B69A6CCFB7CFA2C" bold="true" box="[886,1182,1479,1503]" italics="true" pageId="0" pageNumber="1">Notiophilus anichtchenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACA0579A8D74FFCB1CBCA6CCFB3FFA2C" box="[1187,1245,1479,1503]" pageId="0" pageNumber="1" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CDFD65788D74FFC91B69A50CFE79FC64" lastPageId="2" lastPageNumber="3" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="855836F38D74FFCB1B69A50CFC16F9AC" blockId="0.[886,1397,1543,1631]" pageId="0" pageNumber="1">
<materialsCitation id="358F3CAE8D74FFCB1B69A50CFC16F9AC" collectingDate="2007-07-22" collectionCode="V" collectorName="Patrikeev" country="China" county="Sinin" elevation="3000" location="Quinhai prov." municipality="Lajishan Mts." pageId="0" pageNumber="1" specimenCount="1" typeStatus="holotype">
<emphasis id="B793EAE18D74FFCB1B69A50CFC06F9EC" bold="true" box="[886,996,1543,1567]" pageId="0" pageNumber="1">
<typeStatus id="5A5C88518D74FFCB1B69A50CFC3DF9EC" box="[886,991,1543,1567]" pageId="0" pageNumber="1" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male.
<collectingCountry id="FDF076638D74FFCB1C33A50CFB92F9EC" box="[1068,1136,1543,1567]" name="China" pageId="0" pageNumber="1">China</collectingCountry>
,
<location id="803860288D74FFCB1C65A50CFAECF9EC" LSID="urn:lsid:plazi:treatment:0D4E87E58D74FFC91B69A6CCFB26FB04:803860288D74FFCB1C65A50CFAECF9EC" box="[1146,1294,1543,1567]" country="China" county="Sinin" municipality="Lajishan Mts." name="Quinhai prov." pageId="0" pageNumber="1">Quinhai prov.</location>
,
<collectingMunicipality id="653CAC898D74FFCB1D08A50CFC4AF9CC" pageId="0" pageNumber="1">Lajishan Mts.</collectingMunicipality>
,
<collectingCounty id="6C394E7F8D74FFCB1BAEA52CFC08F9CC" box="[945,1002,1575,1599]" pageId="0" pageNumber="1">Sinin</collectingCounty>
v.,
<quantity id="421F9B168D74FFCB1C12A52CFBBCF9CC" box="[1037,1118,1575,1599]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="1" unit="m" value="3000.0">
<elevation id="0ECAD1C08D74FFCB1C12A52CFBBCF9CC" box="[1037,1118,1575,1599]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="1" unit="m" value="3000.0">3000 m</elevation>
</quantity>
,
<date id="F15910338D74FFCB1C77A52CFB09F9CC" box="[1128,1259,1575,1599]" pageId="0" pageNumber="1" value="2007-07-22">
<collectingDate id="E11DE9DB8D74FFCB1C77A52CFB09F9CC" box="[1128,1259,1575,1599]" pageId="0" pageNumber="1" value="2007-07-22">22.VII.2007</collectingDate>
</date>
,
<collectionCode id="E3F6AE368D74FFCB1CEAA523FAEAF9CC" box="[1269,1288,1576,1599]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="0" pageNumber="1" type="Museum">V</collectionCode>
.
<collectorName id="281253258D74FFCB1D0EA52CFA97F9CC" box="[1297,1397,1575,1599]" pageId="0" pageNumber="1">Patrikeev</collectorName>
leg. (
<figureCitation id="1DDC2A768D74FFCB1BB7A54CFC01F9AC" box="[936,995,1607,1631]" captionStart="Fig" captionStartId="1.[262,302,988,1012]" captionText="Fig. 1. Holotype of N. anichtchenkoi sp. n." pageId="0" pageNumber="1">Fig. 1</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="D198667B8D75FFCA1919A0D7FD32FC07" box="[262,720,988,1012]" pageId="1" pageNumber="2" startId="1.[262,302,988,1012]" targetType="figure">
<paragraph id="855836F38D75FFCA1919A0D7FD32FC07" blockId="1.[262,720,988,1012]" box="[262,720,988,1012]" pageId="1" pageNumber="2">
Fig. 1. Holotype of
<taxonomicName id="42E74D708D75FFCA19C9A0D6FD72FC07" authorityName="Barševskis" authorityYear="2009" box="[470,656,988,1012]" class="Insecta" family="Carabidae" genus="Notiophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="anichtchenkoi" status="sp. nov.">N. anichtchenkoi</taxonomicName>
<taxonomicNameLabel id="ACA0579A8D75FFCA1A89A0D6FD32FC07" box="[662,720,989,1012]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="D198667B8D75FFCA1B2EA0D7FB17FC07" box="[817,1269,988,1013]" pageId="1" pageNumber="2" startId="1.[817,857,988,1012]" targetBox="[931,1180,225,906]" targetPageId="1" targetType="figure">
<paragraph id="855836F38D75FFCA1B2EA0D7FB17FC07" blockId="1.[817,1269,988,1013]" box="[817,1269,988,1013]" pageId="1" pageNumber="2">
Fig. 2. Paratype of
<taxonomicName id="42E74D708D75FFCA1BE2A0D6FB54FC07" authorityName="Barševskis" authorityYear="2009" box="[1021,1206,988,1012]" class="Insecta" family="Carabidae" genus="Notiophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="anichtchenkoi" status="sp. nov.">N. anichtchenkoi</taxonomicName>
<taxonomicNameLabel id="ACA0579A8D75FFCA1CA4A0D6FB17FC07" box="[1211,1269,989,1012]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="D198667B8D75FFCA1919A572FD1AF962" box="[262,760,1657,1681]" pageId="1" pageNumber="2" startId="1.[262,302,1657,1681]" targetType="figure">
<paragraph id="855836F38D75FFCA1919A572FD1AF962" blockId="1.[262,760,1657,1681]" box="[262,760,1657,1681]" pageId="1" pageNumber="2">Fig. 3. Measurements of pronotum (Holotype)</paragraph>
</caption>
<caption id="D198667B8D75FFCA1B35A574FB12F964" box="[810,1264,1663,1687]" pageId="1" pageNumber="2" startId="1.[810,850,1663,1687]" targetType="figure">
<paragraph id="855836F38D75FFCA1B35A574FB12F964" blockId="1.[810,1264,1663,1687]" box="[810,1264,1663,1687]" pageId="1" pageNumber="2">Fig. 4. Measurements of elytra (Holotype)</paragraph>
</caption>
<caption id="D198667B8D76FFC9194DA197FC7CFD47" box="[338,926,668,692]" pageId="2" pageNumber="3" startId="2.[338,378,668,692]" targetType="figure">
<paragraph id="855836F38D76FFC9194DA197FC7CFD47" blockId="2.[338,926,668,692]" box="[338,926,668,692]" pageId="2" pageNumber="3">
Fig. 5.
<emphasis id="B793EAE18D76FFC919BFA197FDE8FD47" box="[416,522,668,692]" italics="true" pageId="2" pageNumber="3">Aedeagus</emphasis>
of
<taxonomicName id="42E74D708D76FFC91A32A196FD04FD47" authorityName="Barševskis" authorityYear="2009" box="[557,742,668,692]" class="Insecta" family="Carabidae" genus="Notiophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="anichtchenkoi" status="sp. nov.">N. anichtchenkoi</taxonomicName>
<taxonomicNameLabel id="ACA0579A8D76FFC91AF1A196FCC4FD47" box="[750,806,668,692]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
(Holotype)
</paragraph>
</caption>
<paragraph id="855836F38D76FFC9194DA1F4FCB2FCE4" blockId="2.[338,848,767,791]" box="[338,848,767,791]" pageId="2" pageNumber="3">
<emphasis id="B793EAE18D76FFC9194DA1F4FE5DFCE4" bold="true" box="[338,447,767,791]" pageId="2" pageNumber="3">
<typeStatus id="5A5C88518D76FFC9194DA1F4FE59FCE4" box="[338,443,767,791]" pageId="2" pageNumber="3" type="paratype">Paratype</typeStatus>
.
</emphasis>
Male, labelled as the
<typeStatus id="5A5C88518D76FFC91A82A1F4FD15FCE4" box="[669,759,767,791]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
(
<figureCitation id="1DDC2A768D76FFC91B1BA1F4FCA1FCE4" box="[772,835,767,791]" captionStart="Fig" captionStartId="1.[817,857,988,1012]" captionTargetBox="[931,1180,225,906]" captionTargetId="figure-358@1.[851,1263,554,576]" captionTargetPageId="1" captionText="Fig. 2. Paratype of N. anichtchenkoi sp. n." pageId="2" pageNumber="3">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="855836F38D76FFC9194DA034FE79FC64" blockId="2.[338,848,831,919]" pageId="2" pageNumber="3">
<materialsCitation id="358F3CAE8D76FFC9194DA034FE75FC64" country="Latvia" location="Daugavpils" pageId="2" pageNumber="3" specimenCount="2" stateProvince="Daugavpils" typeStatus="holotype">
<typeStatus id="5A5C88518D76FFC9194DA034FE5EFCA4" box="[338,444,831,855]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
&amp;
<typeStatus id="5A5C88518D76FFC919F6A04BFDADFCAB" box="[489,591,832,856]" pageId="2" pageNumber="3" type="paratype">paratype</typeStatus>
are deposited in the authors collection (abbrev. ABC) in
<collectingRegion id="4723F8118D76FFC91ACEA054FCAEFC84" box="[721,844,863,887]" country="Latvia" name="Daugavpils" pageId="2" pageNumber="3">Daugavpils</collectingRegion>
,
<collectingCountry id="FDF076638D76FFC9194DA074FE75FC64" box="[338,407,895,919]" name="Latvia" pageId="2" pageNumber="3">Latvia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CDFD65788D76FFC9194DA0B4FB2AFBA4" pageId="2" pageNumber="3" type="description">
<paragraph id="855836F38D76FFC9194DA0B4FB3AFCC4" blockId="2.[338,849,959,1687]" lastBlockId="2.[886,1397,767,823]" pageId="2" pageNumber="3">
<emphasis id="B793EAE18D76FFC9194DA0B4FE2DFC24" bold="true" box="[338,463,959,983]" pageId="2" pageNumber="3">Description</emphasis>
. Body length
<quantity id="421F9B168D76FFC91A45A0B4FD19FC24" box="[602,763,959,983]" metricMagnitude="-3" metricUnit="m" metricValue="6.045" metricValueMax="6.16" metricValueMin="5.93" pageId="2" pageNumber="3" unit="mm" value="6.045" valueMax="6.16" valueMin="5.93">5.93 6.16 mm</quantity>
. Surface flat, shiny, dark bronze-coloured, monochromatic, without yellow spots. Ventral side black, shiny, with bronze-coloured lustre. Head short, width over eyes usually as wide as pronotum (
<figureCitation id="1DDC2A768D76FFC91AE0A734FCA3FBA4" box="[767,833,1087,1111]" captionStart="Fig" captionStartId="1.[262,302,1657,1681]" captionText="Fig. 3. Measurements of pronotum (Holotype)" pageId="2" pageNumber="3">Fig. 3</figureCitation>
). Head width
<quantity id="421F9B168D76FFC919D5A754FD8EFB84" box="[458,620,1119,1143]" metricMagnitude="-3" metricUnit="m" metricValue="1.725" metricValueMax="1.74" metricValueMin="1.71" pageId="2" pageNumber="3" unit="mm" value="1.725" valueMax="1.74" valueMin="1.71">1.71 1.74 mm</quantity>
. Labrum with rounded front side and 6th setiferous punctures, not excised. 1st palpomere, apex of apical palpomere, basal part of 2nd 4th antennomeres yellow. Frons with 9 10 a little irregular keels. Pronotum length
<quantity id="421F9B168D76FFC91982A7F4FDAFFAE4" box="[413,589,1279,1303]" metricMagnitude="-3" metricUnit="m" metricValue="1.105" metricValueMax="1.11" metricValueMin="1.1" pageId="2" pageNumber="3" unit="mm" value="1.105" valueMax="1.11" valueMin="1.1">1.10 1.11 mm</quantity>
, width
<quantity id="421F9B168D76FFC91A82A7F4FCAEFAE4" box="[669,844,1279,1303]" metricMagnitude="-3" metricUnit="m" metricValue="1.73" metricValueMax="1.74" metricValueMin="1.72" pageId="2" pageNumber="3" unit="mm" value="1.73" valueMax="1.74" valueMin="1.72">1.72 1.74 mm</quantity>
. Pronotal sides distinctly sinuated before acute hind angles. Surface of pronotum deeply punctated and wrinkled. Bottom side of thorax deeply punctated. Elytra (
<figureCitation id="1DDC2A768D76FFC91A74A674FD4CFA64" box="[619,686,1407,1431]" captionStart="Fig" captionStartId="1.[810,850,1663,1687]" captionText="Fig. 4. Measurements of elytra (Holotype)" pageId="2" pageNumber="3">Fig. 4</figureCitation>
) length
<quantity id="421F9B168D76FFC91B14A674FE4EFA44" metricMagnitude="-3" metricUnit="m" metricValue="3.425" metricValueMax="3.49" metricValueMin="3.36" pageId="2" pageNumber="3" unit="mm" value="3.425" valueMax="3.49" valueMin="3.36">3.36 3.49 mm</quantity>
, width
<quantity id="421F9B168D76FFC919EDA694FDAFFA44" box="[498,589,1439,1463]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="3" unit="mm" value="2.1">2.10 mm</quantity>
. 3rd 7th interval of elytra with very strong isodiametrical microsculpture. 1st 2nd and marginal intervals smooth, shiny, only in basal and apical parts with microsculpture. 2nd interval has equal width as 3rd 5th together, 5th interval is narrower than 4th or 6th. Elytra with single dorsal (
<typeStatus id="5A5C88518D76FFC91A00A554FD9CF984" box="[543,638,1631,1655]" pageId="2" pageNumber="3" type="paratype">paratype</typeStatus>
right elytron in 4th interval with two dorsal pores) and two apical pores. Legs bichromatic, with reddish brown tibia.
<emphasis id="B793EAE18D76FFC91B69A014FC3FFCC4" box="[886,989,799,823]" italics="true" pageId="2" pageNumber="3">Aedeagus</emphasis>
(
<figureCitation id="1DDC2A768D76FFC91BF6A014FBC3FCC4" box="[1001,1057,799,823]" captionStart="Fig" captionStartId="2.[338,378,668,692]" captionText="Fig. 5. Aedeagus of N. anichtchenkoi sp. n.(Holotype)" pageId="2" pageNumber="3">Fig.5</figureCitation>
) length
<quantity id="421F9B168D76FFC91C6AA014FB36FCC4" box="[1141,1236,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="2" pageNumber="3" unit="mm" value="1.72">1.72 mm</quantity>
.
</paragraph>
<paragraph id="855836F38D76FFC91B69A054FB2AFBA4" blockId="2.[886,1396,863,1111]" pageId="2" pageNumber="3">
<emphasis id="B793EAE18D76FFC91B69A054FC1EFC84" bold="true" box="[886,1020,863,887]" pageId="2" pageNumber="3">Comparison</emphasis>
. Similar to
<taxonomicName id="42E74D708D76FFC91C6DA06BFAC2FC84" authority="Bars." authorityName="Bars." box="[1138,1312,863,887]" class="Insecta" family="Carabidae" genus="Notiophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="katrinae">
<emphasis id="B793EAE18D76FFC91C6DA06BFB07FC84" box="[1138,1253,863,887]" italics="true" pageId="2" pageNumber="3">N. katrinae</emphasis>
Barš.
</taxonomicName>
(
<collectingCountry id="FDF076638D76FFC91D33A054FA92FC84" box="[1324,1392,863,887]" name="China" pageId="2" pageNumber="3">China</collectingCountry>
: N
<collectingRegion id="4723F8118D76FFC91B92A074FC04FC64" box="[909,998,895,919]" country="China" name="Sichuan" pageId="2" pageNumber="3">Sichuan</collectingRegion>
), but differs by large body length, other coloration of legs, number of frontal keels and other characters. Very similar to
<taxonomicName id="42E74D708D76FFC91CFDA0CBFC50FC04" authority="Bates" authorityName="Bates" authorityYear="1881" class="Insecta" family="Carabidae" genus="Notiophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="specularis">
<emphasis id="B793EAE18D76FFC91CFDA0CBFA96FC24" box="[1250,1396,959,983]" italics="true" pageId="2" pageNumber="3">N. specularis</emphasis>
Bates
</taxonomicName>
in body shape (Central America:
<collectingCountry id="FDF076638D76FFC91D04A0D4FA8DFC04" box="[1307,1391,991,1015]" name="Mexico" pageId="2" pageNumber="3">Mexico</collectingCountry>
,
<collectingCountry id="FDF076638D76FFC91B69A0F4FC0AFBE4" box="[886,1000,1023,1047]" name="Guatemala" pageId="2" pageNumber="3">Guatemala</collectingCountry>
etc.), but differs with rounded labrum, number of frontal keels, different form of pronotum and other characters.
</paragraph>
</subSubSection>
<subSubSection id="CDFD65788D76FFC91B69A774FB26FB04" pageId="2" pageNumber="3" type="etymology">
<paragraph id="855836F38D76FFC91B69A774FB26FB04" blockId="2.[886,1396,1151,1271]" pageId="2" pageNumber="3">
<emphasis id="B793EAE18D76FFC91B69A774FC0CFB64" bold="true" box="[886,1006,1151,1175]" pageId="2" pageNumber="3">Etymology</emphasis>
. This new species is named in honour of my colleague, carabidologist Aleksander Anichtchenko (Institute of Systematic Biology,
<collectingRegion id="4723F8118D76FFC91B69A7D4FC12FB04" box="[886,1008,1247,1271]" country="Latvia" name="Daugavpils" pageId="2" pageNumber="3">Daugavpils</collectingRegion>
University,
<collectingCountry id="FDF076638D76FFC91C70A7D4FB5AFB04" box="[1135,1208,1247,1271]" name="Latvia" pageId="2" pageNumber="3">Latvia</collectingCountry>
).
</paragraph>
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View file

@ -0,0 +1,146 @@
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<taxonomicName id="65FF6DB71A038739FDD408CDD8FFFF26" ID-CoL="4KHC8" authority="Newman, 1842" authorityName="Newman" authorityYear="1842" box="[624,944,177,202]" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="notata">
<emphasis id="908BCA261A038739FDD408CDD8FFFF26" bold="true" box="[624,944,177,202]" pageId="3" pageNumber="78">
<emphasis id="908BCA261A038739FDD408CDD84DFF25" bold="true" box="[624,770,178,202]" italics="true" pageId="3" pageNumber="78">Plocia notata</emphasis>
<bibRefCitation id="C66E6BC51A038739FCAD08CED8FFFF26" author="Newman M." box="[777,944,177,201]" pageId="3" pageNumber="78" pagination="288 - 293" refId="ref4137" refString="Newman M. 1842. Cerambycitum insularum manillarum a dom. Cuming captorum enumeratio digesta. The Entomologist, London 18: 288 - 293." type="journal article" year="1842">Newman, 1842</bibRefCitation>
</emphasis>
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<paragraph id="A24016341A038739FDD40890D9ADFE32" blockId="3.[624,1103,239,723]" pageId="3" pageNumber="78">
<bibRefCitation id="C66E6BC51A038739FDD40890D81CFEE8" author="Newman M." box="[624,851,239,263]" pageId="3" pageNumber="78" pagination="288 - 293" refId="ref4137" refString="Newman M. 1842. Cerambycitum insularum manillarum a dom. Cuming captorum enumeratio digesta. The Entomologist, London 18: 288 - 293." type="journal article" year="1842">Newman, 1842: 292</bibRefCitation>
; Thomsom, 1864: 45;
<bibRefCitation id="C66E6BC51A038739FDD40972D834FECA" author="Lacordaire J. T." box="[624,891,269,294]" pageId="3" pageNumber="78" pagination="411 - 930" refId="ref3660" refString="Lacordaire J. T. 1872. Histoire Naturelle des Insectes. Genera of the Beetles or methodical and critical exposition of all the genera proposed so far in this or- der of insects. Family LXVIII. Longhorn beetles. (continued). Subfamily III. LAMIIDES. Paris. Encyclopedic Library of Roret. 9 (2): 411 - 930. http: // www. biodiversitylibrary. org / ite m / 103435" type="journal article" year="1872">Lacordaire, 1872: 610</bibRefCitation>
; Gemmainger &amp; Harold, 1873: 3103;
<bibRefCitation id="C66E6BC51A038739FCFE0953DF0CFEAB" author="Heyrovsky L." box="[858,1091,300,324]" pageId="3" pageNumber="78" pagination="19 - 20" refId="ref3529" refString="Heyrovsky L. 1935. Epaphra albicornis sp. n. (Col., Ceramb.). Casopis ceske spolecnosti entomologicke, Praha 32 (1): 19 - 20." type="journal article" year="1935">Heyrovsky, 1935: 19</bibRefCitation>
, 20;
<bibRefCitation id="C66E6BC51A038739FD3B0934D81BFE8C" author="Kano T." box="[671,852,331,355]" pageId="3" pageNumber="78" refId="ref3625" refString="Kano T. 1939. Second contribution of the knowledge of the coleopterous fauna" type="book" year="1939">Kano, 1939: 32</bibRefCitation>
; Breuning, 1949: 49;
<bibRefCitation id="C66E6BC51A038739FDD40915D81FFE6D" author="Breuning S." box="[624,848,362,386]" pageId="3" pageNumber="78" pagination="109 - 182" refId="ref3076" refString="Breuning S. 1960. Catalogue des Lamiaires du Monde (Col. Ceramb.). Verlag des Museums G. Frey, Tutzingbei Munchen, (3): 109 - 182." type="journal article" year="1960">Breuning, 1960:139</bibRefCitation>
;
<bibRefCitation id="C66E6BC51A038739FCC70915DF06FE6D" author="Breuning S." box="[867,1097,362,386]" pageId="3" pageNumber="78" pagination="1 - 80" refId="ref3111" refString="Breuning S. 1964. Tribus Apomecynini Lac. Die Apomecynini der asiatischaustralischen Region. Abhandlungen und Berichteaus dem staatlichen Museum fur Tierkunde in Dresden, 30 (1): 1 - 80." type="journal article" year="1964">Breuning, 1964: 79</bibRefCitation>
;
<bibRefCitation id="C66E6BC51A038739FDD409F6D856FE4F" author="Hua L." box="[624,793,392,416]" pageId="3" pageNumber="78" pagination="1 - 612" refId="ref3563" refString="Hua L. 2002. List of Chinese Insects. Zhongshan (Sun Yat-sen) University Press, Guangzhou. List of Chinese Insects 2: 1 - 612." type="journal article" year="2002">Hua, 2002: 216</bibRefCitation>
;
<bibRefCitation id="C66E6BC51A038739FC8109F7D9C1FE50" author="Tavakilian G. L. &amp; Jiroux E" pageId="3" pageNumber="78" pagination="76 - 81" refId="ref4295" refString="Tavakilian G. L., Jiroux E 2015. New nomenclatural changes for 2015 (Coleoptera, Cerambycidae). Les Cahiers Magellanes (NS) 20: 76 - 81." type="journal article" year="2015">Tavakilian &amp; Jiroux, 2015: 78</bibRefCitation>
; Barsevskis, 2017: 185;
<bibRefCitation id="C66E6BC51A038739FC3209D8D991FE32" author="Weigel A. &amp; Skale A." pageId="3" pageNumber="78" pagination="345 - 362" refId="ref4483" refString="Weigel A., Skale A. 2021. On the taxonomy, synonymy and faunistics of the Apomecynini of the Asian-Australian region (Coleoptera: Cerambycidae: Lamiinae). Part 8. In Dmitry Telnov, Biodiversity, Biogeography and Nature conservation in Wallacea and New Guinea. The Entomological Society of Latvia, Riga. 4: 345 - 362." type="journal article" year="2021">Weigel &amp; Skale, 2021: 351</bibRefCitation>
.
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<emphasis id="908BCA261A038739FDD4099BD8B6FE13" bold="true" box="[624,1017,484,508]" pageId="3" pageNumber="78">Type information and deposition:</emphasis>
<taxonomicName id="65FF6DB71A038739FBA6099BD824FDF4" authority="Newman, 1842" authorityName="Newman" authorityYear="1842" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="notata">
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<bibRefCitation id="C66E6BC51A038739FD640A7CD824FDF4" author="Newman M." box="[704,875,515,539]" pageId="3" pageNumber="78" pagination="288 - 293" refId="ref4137" refString="Newman M. 1842. Cerambycitum insularum manillarum a dom. Cuming captorum enumeratio digesta. The Entomologist, London 18: 288 - 293." type="journal article" year="1842">Newman, 1842</bibRefCitation>
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,
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,
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;
<taxonomicName id="65FF6DB71A038739FDD40A5DDF48FDD6" authority="Heyrovsky 1935" authorityName="Heyrovsky" authorityYear="1935" box="[624,1031,545,570]" class="Insecta" family="Cerambycidae" genus="Epaphra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="albicornis">
<emphasis id="908BCA261A038739FDD40A5DD80BFDD5" box="[624,836,546,570]" italics="true" pageId="3" pageNumber="78">Epaphra albicornis</emphasis>
<bibRefCitation id="C66E6BC51A038739FCEB0A5DDF48FDD6" author="Heyrovsky L." box="[847,1031,545,570]" pageId="3" pageNumber="78" pagination="19 - 20" refId="ref3529" refString="Heyrovsky L. 1935. Epaphra albicornis sp. n. (Col., Ceramb.). Casopis ceske spolecnosti entomologicke, Praha 32 (1): 19 - 20." type="journal article" year="1935">Heyrovsky 1935</bibRefCitation>
</taxonomicName>
,
<typeStatus id="7D44A8961A038739FBB20A5DD9E7FDB6" pageId="3" pageNumber="78" type="syntype">Syntypes</typeStatus>
(4), Prague National Museum.
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<emphasis id="908BCA261A038739FDD40A20D9ABFD98" bold="true" box="[624,740,607,631]" pageId="3" pageNumber="78">Synonym:</emphasis>
<taxonomicName id="65FF6DB71A038739FD550A20DF05FD98" authority="Newman, 1842" authorityName="Newman" authorityYear="1842" box="[753,1098,607,631]" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="notata">
<emphasis id="908BCA261A038739FD550A20D8C6FD98" box="[753,905,607,631]" italics="true" pageId="3" pageNumber="78">Plocia notata</emphasis>
<bibRefCitation id="C66E6BC51A038739FC330A20DF05FD98" author="Newman M." box="[919,1098,607,631]" pageId="3" pageNumber="78" pagination="288 - 293" refId="ref4137" refString="Newman M. 1842. Cerambycitum insularum manillarum a dom. Cuming captorum enumeratio digesta. The Entomologist, London 18: 288 - 293." type="journal article" year="1842">Newman, 1842</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="65FF6DB71A038739FDD40A01D8B5FD7A" authority="Heyrovsky 1935" authorityName="Heyrovsky" authorityYear="1935" box="[624,1018,637,662]" class="Insecta" family="Cerambycidae" genus="Epaphra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="albicornis">
<emphasis id="908BCA261A038739FDD40A01D80FFD79" box="[624,832,638,662]" italics="true" pageId="3" pageNumber="78">Epaphra albicornis</emphasis>
<bibRefCitation id="C66E6BC51A038739FCE30A02D8B5FD7A" author="Heyrovsky L." box="[839,1018,637,661]" pageId="3" pageNumber="78" pagination="19 - 20" refId="ref3529" refString="Heyrovsky L. 1935. Epaphra albicornis sp. n. (Col., Ceramb.). Casopis ceske spolecnosti entomologicke, Praha 32 (1): 19 - 20." type="journal article" year="1935">Heyrovsky 1935</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<emphasis id="908BCA261A038739FDD40AE3D9B4FD5B" bold="true" box="[624,763,668,692]" pageId="3" pageNumber="78">Distribution</emphasis>
:
<collectingCountry id="DAE856A41A038739FCAB0AE3D8C8FD5B" box="[783,903,668,692]" name="Philippines" pageId="3" pageNumber="78">Philippines</collectingCountry>
(Luzon:
<collectingRegion id="603BD8D61A038739FC510AE3DF05FD5B" box="[1013,1098,668,692]" country="Philippines" name="Manila" pageId="3" pageNumber="78">Manila</collectingRegion>
; Mt. Makiling,
<collectingRegion id="603BD8D61A038739FCA80AC4D82EFD3C" box="[780,865,699,723]" country="Philippines" name="Laguna" pageId="3" pageNumber="78">Laguna</collectingRegion>
).
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<heading id="F908A1581A038739FDD40B68DF4DFCC0" bold="true" box="[624,1026,791,815]" fontSize="10" level="2" pageId="3" pageNumber="78" reason="4">
<taxonomicName id="65FF6DB71A038739FDD40B68DF4DFCC0" ID-CoL="BS8WW" authority="(Breuning, 1965)" baseAuthorityName="Breuning" baseAuthorityYear="1965" box="[624,1026,791,815]" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="908BCA261A038739FDD40B68DF4DFCC0" bold="true" box="[624,1026,791,815]" pageId="3" pageNumber="78">
<emphasis id="908BCA261A038739FDD40B68D874FCC0" bold="true" box="[624,827,791,815]" italics="true" pageId="3" pageNumber="78">Plocia puncticollis</emphasis>
(
<bibRefCitation id="C66E6BC51A038739FCED0B68D8B5FCC0" author="Breuning S." box="[841,1018,791,815]" pageId="3" pageNumber="78" pagination="177 - 182" refId="ref3146" refString="Breuning S. 1965. Neue Cerambycidenaus den Sammlungen des Zoologischen Museums der Humboldt-Universitat zu Berlin (Coleoptera, Cerambycidae) Funfter Teil. Mitteilungenaus dem Zoologischen Museum in Berlin, 41 (2): 177 - 182." type="journal article" year="1965">Breuning, 1965</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A24016341A038739FDD40B2BD9ADFC64" blockId="3.[624,1103,852,1091]" pageId="3" pageNumber="78">
<bibRefCitation id="C66E6BC51A038739FDD40B2BD828FC83" author="Breuning S." box="[624,871,852,876]" pageId="3" pageNumber="78" pagination="177 - 182" refId="ref3146" refString="Breuning S. 1965. Neue Cerambycidenaus den Sammlungen des Zoologischen Museums der Humboldt-Universitat zu Berlin (Coleoptera, Cerambycidae) Funfter Teil. Mitteilungenaus dem Zoologischen Museum in Berlin, 41 (2): 177 - 182." type="journal article" year="1965">Breuning, 1965: 182</bibRefCitation>
;
<bibRefCitation id="C66E6BC51A038739FCDB0B2BD991FC64" author="Weigel A. &amp; Skale A." pageId="3" pageNumber="78" pagination="345 - 362" refId="ref4483" refString="Weigel A., Skale A. 2021. On the taxonomy, synonymy and faunistics of the Apomecynini of the Asian-Australian region (Coleoptera: Cerambycidae: Lamiinae). Part 8. In Dmitry Telnov, Biodiversity, Biogeography and Nature conservation in Wallacea and New Guinea. The Entomological Society of Latvia, Riga. 4: 345 - 362." type="journal article" year="2021">Weigel &amp; Skale, 2021: 353</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EAE545BF1A038739FDD40BEED852FBCB" pageId="3" pageNumber="78" type="materials_examined">
<paragraph id="A24016341A038739FDD40BEED86EFC09" blockId="3.[624,1103,852,1091]" pageId="3" pageNumber="78">
<emphasis id="908BCA261A038739FDD40BEED8BFFC46" bold="true" box="[624,1008,913,937]" pageId="3" pageNumber="78">Type information and deposition:</emphasis>
<taxonomicName id="65FF6DB71A038739FC530BEDD9EDFC27" authorityName="Breuning" authorityYear="1939" class="Insecta" family="Cerambycidae" genus="Parepilysta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="908BCA261A038739FC530BEDD9EDFC27" italics="true" pageId="3" pageNumber="78">Parepilysta</emphasis>
</taxonomicName>
(
<materialsCitation id="12971C691A038739FD1A0BCFD86EFC09" collectionCode="ZMB" pageId="3" pageNumber="78" specimenCount="1" typeStatus="holotype">
s. s.)
<taxonomicName id="65FF6DB71A038739FCA50BCFDF06FC27" authority="Breuning, 1965" authorityName="Breuning" authorityYear="1965" box="[769,1097,944,968]" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="908BCA261A038739FCA50BCFD831FC27" box="[769,894,944,968]" italics="true" pageId="3" pageNumber="78">puncticollis</emphasis>
<bibRefCitation id="C66E6BC51A038739FC340BCFDF06FC27" author="Breuning S." box="[912,1097,944,968]" pageId="3" pageNumber="78" pagination="177 - 182" refId="ref3146" refString="Breuning S. 1965. Neue Cerambycidenaus den Sammlungen des Zoologischen Museums der Humboldt-Universitat zu Berlin (Coleoptera, Cerambycidae) Funfter Teil. Mitteilungenaus dem Zoologischen Museum in Berlin, 41 (2): 177 - 182." type="journal article" year="1965">Breuning, 1965</bibRefCitation>
</taxonomicName>
,
<typeStatus id="7D44A8961A038739FDD40BB0D999FC08" box="[624,726,975,999]" pageId="3" pageNumber="78" type="holotype">Holotype</typeStatus>
,
<collectionCode id="C4EE8EF11A038739FD450BB0D851FC09" box="[737,798,975,998]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="78" type="Museum">ZMB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="A24016341A038739FDD40B92D852FBCB" blockId="3.[624,1103,852,1091]" pageId="3" pageNumber="78">
<emphasis id="908BCA261A038739FDD40B92D9ABFBEA" bold="true" box="[624,740,1005,1029]" pageId="3" pageNumber="78">Synonym:</emphasis>
<taxonomicName id="65FF6DB71A038739FD500B91D820FBE9" authorityName="Breuning" authorityYear="1939" box="[756,879,1006,1030]" class="Insecta" family="Cerambycidae" genus="Parepilysta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="908BCA261A038739FD500B91D820FBE9" box="[756,879,1006,1030]" italics="true" pageId="3" pageNumber="78">Parepilysta</emphasis>
</taxonomicName>
(s. s.)
<taxonomicName id="65FF6DB71A038739FC680B91D856FBCB" authority="Breuning, 1965" authorityName="Breuning" authorityYear="1965" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="908BCA261A038739FC680B91DF06FBE9" box="[972,1097,1006,1030]" italics="true" pageId="3" pageNumber="78">puncticollis</emphasis>
<bibRefCitation id="C66E6BC51A038739FDD40C73D856FBCB" author="Breuning S." box="[624,793,1036,1060]" pageId="3" pageNumber="78" pagination="177 - 182" refId="ref3146" refString="Breuning S. 1965. Neue Cerambycidenaus den Sammlungen des Zoologischen Museums der Humboldt-Universitat zu Berlin (Coleoptera, Cerambycidae) Funfter Teil. Mitteilungenaus dem Zoologischen Museum in Berlin, 41 (2): 177 - 182." type="journal article" year="1965">Breuning, 1965</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
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<emphasis id="908BCA261A038739FDD40C54D84BFBAC" bold="true" box="[624,772,1067,1091]" pageId="3" pageNumber="78">Distribution:</emphasis>
<collectingCountry id="DAE856A41A038739FCAE0C54D8CDFBAC" box="[778,898,1067,1091]" name="Philippines" pageId="3" pageNumber="78">Philippines</collectingCountry>
(Luzon).
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View file

@ -0,0 +1,119 @@
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<mods:title id="8366ADCD7A60A8549C53D36F45903366">Catalog of the genus Plocia Newman, 1842 (Cerambycidae: Lamiinae: Apomecynini) of the Philippines with the description of a new species from Davao De Oro Mindanao Island</mods:title>
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<heading id="F908A1581A038739FDD40C17D8BCFB6F" bold="true" box="[624,1011,1128,1152]" fontSize="10" level="2" pageId="3" pageNumber="78" reason="4">
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<emphasis id="908BCA261A038739FDD40C17D8BCFB6F" bold="true" box="[624,1011,1128,1152]" pageId="3" pageNumber="78">
<emphasis id="908BCA261A038739FDD40C17D867FB6F" bold="true" box="[624,808,1128,1152]" italics="true" pageId="3" pageNumber="78">Plocia splendens</emphasis>
(Hüdepohl, 1995)
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<bibRefCitation id="C66E6BC51A038739FDD40CDAD813FB52" author="Hudephol K. E." box="[624,860,1189,1213]" pageId="3" pageNumber="78" pagination="281 - 316" refId="ref3593" refString="Hudephol K. E. 1995 Uber sudostasiatische Cerambyciden XIII (Coleoptera, Cerambycidae). Entomofauna Zeitschrift fur Entomologie, Ansfelden, 16 (14): 281 - 316." type="journal article" year="1995">Hudephol, 1995: 285</bibRefCitation>
; Tavakilan &amp; Jiroux, 2015: 78;
<bibRefCitation id="C66E6BC51A038739FD780CBBDF4DFB33" author="Weigel A. &amp; Skale A." box="[732,1026,1220,1244]" pageId="3" pageNumber="78" pagination="345 - 362" refId="ref4483" refString="Weigel A., Skale A. 2021. On the taxonomy, synonymy and faunistics of the Apomecynini of the Asian-Australian region (Coleoptera: Cerambycidae: Lamiinae). Part 8. In Dmitry Telnov, Biodiversity, Biogeography and Nature conservation in Wallacea and New Guinea. The Entomological Society of Latvia, Riga. 4: 345 - 362." type="journal article" year="2021">Weigel &amp; Skale, 2021: 353</bibRefCitation>
.
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<paragraph id="A24016341A038739FDD40C9CD850FAD7" blockId="3.[624,1103,1189,1459]" pageId="3" pageNumber="78">
<emphasis id="908BCA261A038739FDD40C9CDF07FB14" bold="true" box="[624,1096,1251,1275]" pageId="3" pageNumber="78">Type Information and deposition:</emphasis>
<taxonomicName id="65FF6DB71A038739FDD40D7EDF06FAF6" authority="Hudephol 1995" authorityName="Hudephol" authorityYear="1995" box="[624,1097,1281,1305]" class="Insecta" family="Cerambycidae" genus="Mimoplocia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="splendens">
<emphasis id="908BCA261A038739FDD40D7ED836FAF6" box="[624,889,1281,1305]" italics="true" pageId="3" pageNumber="78">Mimoplocia splendens</emphasis>
<materialsCitation id="12971C691A038739FC2F0D7EDF06FAF6" box="[907,1097,1281,1305]" collectionCode="ZSM" pageId="3" pageNumber="78" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="C66E6BC51A038739FC2F0D7EDF06FAF6" author="Hudephol K. E." box="[907,1097,1281,1305]" pageId="3" pageNumber="78" pagination="281 - 316" refId="ref3593" refString="Hudephol K. E. 1995 Uber sudostasiatische Cerambyciden XIII (Coleoptera, Cerambycidae). Entomofauna Zeitschrift fur Entomologie, Ansfelden, 16 (14): 281 - 316." type="journal article" year="1995">Hudephol 1995</bibRefCitation>
</materialsCitation>
</taxonomicName>
,
<typeStatus id="7D44A8961A038739FDD40D5FD999FAD7" box="[624,726,1312,1336]" pageId="3" pageNumber="78" type="holotype">Holotype</typeStatus>
,
<collectionCode id="C4EE8EF11A038739FD450D5FD854FAD7" box="[737,795,1312,1336]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="78" type="Museum">ZSM</collectionCode>
.
</paragraph>
<paragraph id="A24016341A038739FDD40D40D9A1FA9A" blockId="3.[624,1103,1189,1459]" pageId="3" pageNumber="78">
<emphasis id="908BCA261A038739FDD40D40D9ABFAB8" bold="true" box="[624,740,1343,1367]" pageId="3" pageNumber="78">Synonym:</emphasis>
<taxonomicName id="65FF6DB71A038739FD5E0D40D9A5FA9A" authority="Hudephol 1965" authorityName="Hudephol" authorityYear="1965" class="Insecta" family="Cerambycidae" genus="Mimoplocia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="splendens">
<emphasis id="908BCA261A038739FD5E0D40DF4DFAB8" box="[762,1026,1343,1367]" italics="true" pageId="3" pageNumber="78">Mimoplocia splendens</emphasis>
Hudephol 1965
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EAE545BF1A038739FDD40D03D80DFA5C" pageId="3" pageNumber="78" type="distribution">
<paragraph id="A24016341A038739FDD40D03D80DFA5C" blockId="3.[624,1103,1189,1459]" pageId="3" pageNumber="78">
<emphasis id="908BCA261A038739FDD40D03D84BFA7B" bold="true" box="[624,772,1404,1428]" pageId="3" pageNumber="78">Distribution:</emphasis>
<collectingCountry id="DAE856A41A038739FCA80D03D8CBFA7B" box="[780,900,1404,1428]" name="Philippines" pageId="3" pageNumber="78">Philippines</collectingCountry>
(Luzon: Sibuyan;
<collectingRegion id="603BD8D61A038739FDD40DE4D874FA5C" box="[624,827,1435,1459]" country="Philippines" name="Romblon" pageId="3" pageNumber="78">Romblon Province</collectingRegion>
)
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,272 @@
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<mods:title id="C556AEDC74E2F09E1BFDB580982E8EFD">Catalog of the genus Plocia Newman, 1842 (Cerambycidae: Lamiinae: Apomecynini) of the Philippines with the description of a new species from Davao De Oro Mindanao Island</mods:title>
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<mods:namePart id="92485EEC2ECCE2A5AC70E38459DC8686">Medina, Milton Norman D.</mods:namePart>
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</mods:name>
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<mods:role id="A2C8FCBBFF7DE8246BFB17BEEB441291">
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<mods:namePart id="46EFB879D08103DCC696E74A71791B37">Loguinsa, Jules T.</mods:namePart>
</mods:name>
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<mods:namePart id="468F8A92C8BD4C2A7DBFC4B57084BC61">Macosang, Monsur M.</mods:namePart>
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<mods:namePart id="D019DAB546CC4570FF04D352E1C27DAA">Panangcad, Jenie Ann M.</mods:namePart>
</mods:name>
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<mods:date id="13DE7D5BF63ADCCAFAA61679144D4156">2023</mods:date>
<mods:detail id="E47037F83FA248639BAFEAAC4CDBD5D2" type="series">
<mods:title id="8E3C408621B87D41E4DC66F0938D54AD">ISSI</mods:title>
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<treatment id="2A56A7221A03873CFDD40E3ED9C0FB52" LSID="urn:lsid:plazi:treatment:2A56A7221A03873CFDD40E3ED9C0FB52" httpUri="http://treatment.plazi.org/id/2A56A7221A03873CFDD40E3ED9C0FB52" lastPageId="6" lastPageNumber="81" pageId="3" pageNumber="78">
<subSubSection id="EAE545BF1A038739FDD40E3ED8AFF9B7" box="[624,992,1600,1625]" pageId="3" pageNumber="78" type="nomenclature">
<paragraph id="A24016341A038739FDD40E3ED8AFF9B7" blockId="3.[624,992,1600,1655]" box="[624,992,1600,1625]" pageId="3" pageNumber="78">
<heading id="F908A1581A038739FDD40E3ED8AFF9B7" bold="true" box="[624,992,1600,1625]" fontSize="10" level="2" pageId="3" pageNumber="78" reason="4">
<emphasis id="908BCA261A038739FDD40E3ED8AFF9B7" bold="true" box="[624,992,1600,1625]" pageId="3" pageNumber="78">
<taxonomicName id="65FF6DB71A038739FDD40E3ED8CDF9B7" authority="Medina" authorityName="Medina &amp; Avergonzado &amp; Loguinsa &amp; Macosang &amp; Panangcad" authorityYear="2023" box="[624,898,1600,1625]" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="maglanai" status="sp. nov.">
<emphasis id="908BCA261A038739FDD40E3ED86BF9B6" bold="true" box="[624,804,1601,1625]" italics="true" pageId="3" pageNumber="78">Plocia maglanai</emphasis>
Medina
</taxonomicName>
<taxonomicNameLabel id="8BB8775D1A038739FC2C0E3FD8AFF9B7" box="[904,992,1600,1624]" pageId="3" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="EAE545BF1A03873EFDD40E20DFDBF951" box="[624,1172,1631,1726]" lastPageId="4" lastPageNumber="79" pageId="3" pageNumber="78" type="description">
<paragraph id="A24016341A038739FDD40E20D9F9F998" blockId="3.[624,992,1600,1655]" box="[624,694,1631,1655]" pageId="3" pageNumber="78">
<heading id="F908A1581A038739FDD40E20D9F9F998" bold="true" box="[624,694,1631,1655]" fontSize="9" level="2" pageId="3" pageNumber="78" reason="6">
<emphasis id="908BCA261A038739FDD40E20D9F9F998" bold="true" box="[624,694,1631,1655]" pageId="3" pageNumber="78">
<figureCitation id="3AC40AB11A038739FDD40E20D9FFF998" box="[624,688,1631,1655]" captionStart="Fig" captionStartId="4.[189,228,1540,1564]" captionTargetBox="[199,1154,203,1503]" captionTargetId="figure-34@4.[188,1171,170,1533]" captionTargetPageId="4" captionText="Fig. 1. Habitus ofPlocia maglanai sp. nov.: A. Dorsal, B. Ventral, C. Lateral, D. Frons." pageId="3" pageNumber="78">Fig. 1</figureCitation>
.
</emphasis>
</heading>
</paragraph>
<caption id="F68046BC1A04873EFF190E7BDF28F9F4" box="[189,1127,1540,1564]" pageId="4" pageNumber="79" startId="4.[189,228,1540,1564]" targetBox="[199,1154,203,1503]" targetPageId="4" targetType="figure">
<paragraph id="A24016341A04873EFF190E7BDF28F9F4" blockId="4.[189,1127,1540,1564]" box="[189,1127,1540,1564]" pageId="4" pageNumber="79">
Fig. 1. Habitus of
<taxonomicName id="65FF6DB71A04873EFEDB0E7BD97EF9F3" authorityName="Medina &amp; Avergonzado &amp; Loguinsa &amp; Macosang &amp; Panangcad" authorityYear="2023" box="[383,561,1540,1564]" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="maglanai" status="sp. nov.">
<emphasis id="908BCA261A04873EFEDB0E7BD97EF9F3" box="[383,561,1540,1564]" italics="true" pageId="4" pageNumber="79">Plocia maglanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8BB8775D1A04873EFD930E7BD9C0F9F4" box="[567,655,1540,1563]" pageId="4" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
: A. Dorsal, B. Ventral, C. Lateral, D. Frons.
</paragraph>
</caption>
<paragraph id="A24016341A04873EFBDD0ED9DFDBF951" blockId="4.[1145,1172,1702,1726]" box="[1145,1172,1702,1726]" pageId="4" pageNumber="79">79</paragraph>
</subSubSection>
<subSubSection id="EAE545BF1A05873CFFD608CEDAF5FE13" lastPageId="6" lastPageNumber="81" pageId="5" pageNumber="80" type="materials_examined">
<paragraph id="A24016341A05873FFFD608CED90EFE8C" blockId="5.[114,593,177,539]" pageId="5" pageNumber="80">
<materialsCitation id="12971C691A05873FFFD608CED90EFE8C" collectionCode="MMCP, PNM" collectorName="de Oro &amp; E. Avergonzado &amp; Philippine National Museum" country="Philippines" elevation="800" location="Pantukan" municipality="Mindanao" pageId="5" pageNumber="80" specimenCount="1" stateProvince="Davao" typeStatus="holotype">
<emphasis id="908BCA261A05873FFFD608CEDA46FF26" bold="true" box="[114,265,177,201]" pageId="5" pageNumber="80">
<typeStatus id="7D44A8961A05873FFFD608CEDA46FF26" box="[114,265,177,201]" pageId="5" pageNumber="80" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
female:
<collectingCountry id="DAE856A41A05873FFE2C08CDD96AFF25" box="[392,549,178,202]" name="Philippines" pageId="5" pageNumber="80">PHILIPPINES</collectingCountry>
<collectingMunicipality id="42248C4E1A05873FFFD608AFDBAEFF07" box="[114,225,208,232]" pageId="5" pageNumber="80">Mindanao</collectingMunicipality>
,
<collectingRegion id="603BD8D61A05873FFF4808AEDA7EFF07" box="[236,305,209,232]" country="Philippines" name="Davao" pageId="5" pageNumber="80">Davao</collectingRegion>
<collectorName id="0F0A73E21A05873FFE9C08AFDACDFF07" box="[312,386,208,232]" pageId="5" pageNumber="80">de Oro</collectorName>
/
<location id="A72040EF1A05873FFE3208AFDAB5FF07" LSID="urn:lsid:plazi:treatment:2A56A7221A03873CFDD40E3ED9C0FB52:A72040EF1A05873FFE3208AFDAB5FF07" box="[406,506,208,232]" country="Philippines" municipality="Mindanao" name="Pantukan" pageId="5" pageNumber="80" stateProvince="Davao">Pantukan</location>
/ Brgy. Tagugpo / VIII.28.2023 /
<quantity id="6507BBD11A05873FFE740890D902FEE8" box="[464,589,239,263]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="80" unit="m" value="800.0">
<elevation id="29D2F1071A05873FFE740890D902FEE8" box="[464,589,239,263]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="80" unit="m" value="800.0">800 masl</elevation>
</quantity>
.
<collectorName id="0F0A73E21A05873FFFD60972DA57FECA" box="[114,280,269,293]" pageId="5" pageNumber="80">E.Avergonzado</collectorName>
leg. /
<collectionCode id="C4EE8EF11A05873FFED50971DA8BFEC9" box="[369,452,270,294]" pageId="5" pageNumber="80">MMCP</collectionCode>
, printed on red card. Type specimen will be deposited at the
<collectorName id="0F0A73E21A05873FFF170934DAA8FE8D" box="[179,487,331,355]" pageId="5" pageNumber="80">Philippine National Museum</collectorName>
(
<collectionCode id="C4EE8EF11A05873FFE510934D97AFE8C" box="[501,565,331,355]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="5" pageNumber="80">PNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="A24016341A05873FFFD60916D958FDF4" blockId="5.[114,593,177,539]" pageId="5" pageNumber="80">
<materialsCitation id="12971C691A05873FFFD60916DA7FFE4F" collectionCode="MMCP" pageId="5" pageNumber="80" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="908BCA261A05873FFFD60916DA55FE6E" bold="true" box="[114,282,361,385]" pageId="5" pageNumber="80">
<typeStatus id="7D44A8961A05873FFFD60916DA59FE6E" box="[114,278,361,385]" pageId="5" pageNumber="80" type="paratype">PARATYPES</typeStatus>
:
</emphasis>
<specimenCount id="B4F9DDBD1A05873FFE8D0915DAD5FE6D" box="[297,410,362,386]" count="2" pageId="5" pageNumber="80" type="female">2 females</specimenCount>
, same label as
<typeStatus id="7D44A8961A05873FFFD609F7DB9DFE4F" box="[114,210,392,416]" pageId="5" pageNumber="80" type="holotype">holotype</typeStatus>
,
<collectionCode id="C4EE8EF11A05873FFF7909F7DA7FFE4F" box="[221,304,392,416]" pageId="5" pageNumber="80">MMCP</collectionCode>
</materialsCitation>
.
<materialsCitation id="12971C691A05873FFE9F09F7D95CFDF4" collectionCode="DUBC" country="Philippines" county="Female" location="Maragusan" municipality="Mindanao" pageId="5" pageNumber="80" specimenCount="1" stateProvince="Compostela" typeStatus="paratype">
<collectingCounty id="4B216EB81A05873FFE9F09F7DAC1FE4F" box="[315,398,392,416]" pageId="5" pageNumber="80">Female</collectingCounty>
:
<collectingCountry id="DAE856A41A05873FFE3C09F7D97AFE4F" box="[408,565,392,416]" name="Philippines" pageId="5" pageNumber="80">PHILIPPINES</collectingCountry>
<collectingMunicipality id="42248C4E1A05873FFFD609D8DBAEFE50" box="[114,225,423,447]" pageId="5" pageNumber="80">Mindanao</collectingMunicipality>
, /
<collectingRegion id="603BD8D61A05873FFF5A09D8DA9FFE50" box="[254,464,423,447]" country="Philippines" name="Compostela" pageId="5" pageNumber="80">Compostela Valley</collectingRegion>
, /
<location id="A72040EF1A05873FFE4A09D8DBD7FE32" LSID="urn:lsid:plazi:treatment:2A56A7221A03873CFDD40E3ED9C0FB52:A72040EF1A05873FFE4A09D8DBD7FE32" country="Philippines" county="Female" municipality="Mindanao" name="Maragusan" pageId="5" pageNumber="80" stateProvince="Compostela">Maragusan</location>
, /
<location id="A72040EF1A05873FFF1709B9DBAAFE32" LSID="urn:lsid:plazi:treatment:2A56A7221A03873CFDD40E3ED9C0FB52:A72040EF1A05873FFF1709B9DBAAFE32" box="[179,229,454,477]" country="Philippines" county="Female" municipality="Mindanao" name="New" pageId="5" pageNumber="80" stateProvince="Compostela">New</location>
<collectingRegion id="603BD8D61A05873FFF4909BADA7EFE32" box="[237,305,453,477]" country="Philippines" name="Albay" pageId="5" pageNumber="80">Albay</collectingRegion>
, 03.2017. / local collector leg./
<location id="A72040EF1A05873FFF10099BDA52FE13" LSID="urn:lsid:plazi:treatment:2A56A7221A03873CFDD40E3ED9C0FB52:A72040EF1A05873FFF10099BDA52FE13" box="[180,285,484,508]" country="Philippines" county="Female" municipality="Mindanao" name="Specimen" pageId="5" pageNumber="80" stateProvince="Compostela">Specimen</location>
deposited at
<location id="A72040EF1A05873FFE75099BDAE2FDF4" LSID="urn:lsid:plazi:treatment:2A56A7221A03873CFDD40E3ED9C0FB52:A72040EF1A05873FFE75099BDAE2FDF4" country="Philippines" county="Female" municipality="Mindanao" name="Daugavpils University Beetles Collection" pageId="5" pageNumber="80" stateProvince="Compostela">Daugavpils University Beetles Collection</location>
(
<collectionCode id="C4EE8EF11A05873FFE1F0A7CD944FDF4" box="[443,523,515,539]" pageId="5" pageNumber="80">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A24016341A05873FFFD60A3FDA3BFD5B" blockId="5.[114,593,576,692]" pageId="5" pageNumber="80">
Description. Dimensions of
<typeStatus id="7D44A8961A05873FFE0B0A3FD95EFDB7" box="[431,529,576,600]" pageId="5" pageNumber="80" type="holotype">holotype</typeStatus>
: LB:
<quantity id="6507BBD11A05873FFFD60A20DB98FD99" box="[114,215,607,631]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="5" pageNumber="80" unit="mm" value="11.5">11.5 mm</quantity>
. WH:
<quantity id="6507BBD11A05873FFE840A20DA37FD99" box="[288,376,607,631]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="80" unit="mm" value="1.5">1.5 mm</quantity>
. LL:
<quantity id="6507BBD11A05873FFE110A20D942FD99" box="[437,525,607,631]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="80" unit="mm" value="0.8">0.8 mm</quantity>
. WL: 1.0 mm.
<collectionCode id="C4EE8EF11A05873FFF7F0A01DBB1FD7A" box="[219,254,638,661]" country="Russia" name="Laboratory of Palaeontology" pageId="5" pageNumber="80">LP</collectionCode>
: 2-.0 mm. WP:
<quantity id="6507BBD11A05873FFE1E0A02D95BFD7A" box="[442,532,637,661]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="80" unit="mm" value="2.5">2.5 mm</quantity>
.
<collectionCode id="C4EE8EF11A05873FFD870A01D902FD7A" box="[547,589,638,661]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="5" pageNumber="80">LE</collectionCode>
:
<quantity id="6507BBD11A05873FFFD60AE3DB88FD5B" box="[114,199,668,692]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="80" unit="mm" value="8.5">8.5 mm</quantity>
. WEH: 4.0 mm.
</paragraph>
<paragraph id="A24016341A05873FFFD60AA6D90FFA3E" blockId="5.[114,593,729,1489]" pageId="5" pageNumber="80">
<emphasis id="908BCA261A05873FFFD60AA6DAEBFD1E" bold="true" box="[114,420,729,753]" pageId="5" pageNumber="80">Adult female. Teguments</emphasis>
in the Head, pronotum, elytra, and legs are matte black; antennae and underside of head are dark brown.
<emphasis id="908BCA261A05873FFF6B0B4ADA43FCA2" bold="true" box="[207,268,821,845]" pageId="5" pageNumber="80">Head</emphasis>
dorsally wider than long; frons, vertex, genae with puncturations and lateral sides covered with recumbent yellowish pubescence; deeper puncturations at frons arranged in random; dorsal median side of vertex free of yellowish recumbent pubescence; one black erect orbital setae; base of frons lined with 3-4 semi-erect black setae; genae lustrous, wider than long; clypleus lustrous, glabrous, rectangular; mandible short robust at base, covered with few long yellowish erect setae.
<emphasis id="908BCA261A05873FFDBB0CF8DB8BFB52" bold="true" pageId="5" pageNumber="80">Antennae</emphasis>
short, reaching until apical third of elytra, densely covered with semi-erect short yellowish setae from scape to antennomere XI; scape robust, normal without apical cicatrix, apex rounded, a little longer than head; pedicle slightly elongated; antennomere III as long as antennomere IV; basal half of antennomere III and IV covered with whitish pubescence; antennomere VII fully covered with whitish pubescence.
</paragraph>
<paragraph id="A24016341A05873FFFD60D88D91EF984" blockId="5.[114,593,1527,1643]" pageId="5" pageNumber="80">
<emphasis id="908BCA261A05873FFFD60D88DBA8F9E0" bold="true" box="[114,231,1527,1551]" pageId="5" pageNumber="80">Prothorax</emphasis>
. Pronotum and Propleuron covered with deep puncturations; pronotum wider than long, median line of pronotum free of yellowish pubescence, lateral sides
</paragraph>
<paragraph id="A24016341A05873FFFD60ED9D8ABFE8C" blockId="5.[114,141,1702,1726]" lastBlockId="5.[624,1102,178,355]" pageId="5" pageNumber="80">80 of pronotum to pleuron covered with yellowish pubescence in line with the yellowish pubescence of the head, with few erect black setae at propleuron. Prosternum covered with fine yellowish recumbent pubescence, bottle-shaped in the middle.</paragraph>
<paragraph id="A24016341A05873FFDD409F7D80FFC45" blockId="5.[624,1103,392,938]" pageId="5" pageNumber="80">
<emphasis id="908BCA261A05873FFDD409F7D9F0FE4F" bold="true" box="[624,703,392,416]" pageId="5" pageNumber="80">Elytra</emphasis>
twice wider than long, covered with fine semi-erect and recumbent yellowish and whitish pubescence with pronounced yellowish recumbent pubescence at basal third. Elytra densely covered with puncturations at the basal half, apical half is devoid of puncturations. At least 4-5 visible striae from elytral suture, gradually diminishing from the apical half towards apex. With a distinct lateral band of whitish pubescence at the middle in line with antennomere VII (fully covered with whitish pubescence). At least five longitudinal bands of whitish pubescence scattered at the apical third and elytral apex. Elytral apex caudate with a pointed sutural spine covered with semi-erect whitish and yellowish pubescence.
</paragraph>
<paragraph id="A24016341A05873FFDD40BB0DF0BFB52" blockId="5.[624,1103,975,1213]" pageId="5" pageNumber="80">
<emphasis id="908BCA261A05873FFDD40BB0D9EAFC08" bold="true" box="[624,677,975,999]" pageId="5" pageNumber="80">Legs</emphasis>
. Coxa, underside of tibia, profemora, and mesofemora are covered with fine yellowish recumbent pubescence; dorsal side of tibia and tarsi with semi-erect whitish and yellowish pubescence; basal half of metafemora with whitish recumbent pubescence, apical half with yellowish pubescence. Claws simple, light brown, glabrous.
</paragraph>
<paragraph id="A24016341A05873CFDD40C9CDBB8FE51" blockId="5.[624,1103,1251,1643]" lastBlockId="6.[189,668,178,446]" lastPageId="6" lastPageNumber="81" pageId="5" pageNumber="80">
<emphasis id="908BCA261A05873FFDD40C9CD882FB14" bold="true" box="[624,973,1251,1275]" pageId="5" pageNumber="80">Mesepimeron, metepisternum,</emphasis>
and lateral sides of
<emphasis id="908BCA261A05873FFD7A0D7ED83AFAF6" bold="true" box="[734,885,1281,1305]" pageId="5" pageNumber="80">metasternum</emphasis>
with deep puncturations arranged in random, covered with yellowish recumbent pubescence, more pronounced at the lateral sides of metasternum. Mesepisternum devoid of puncturations, densely covered with fine black and yellowish pubescence. Mesosternum quadrangular, almost cube-liked, with fine yellowish pubescence. Metasternum wider than long, median line devoid of puncturations and yellowish pubescence with prominent medial concavity. Abdominal ventrite I as long as wide as metasternum; lateral sides of ventrites I to
<collectionCode id="C4EE8EF11A06873CFE5608AED94AFF07" box="[498,517,209,232]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="81" type="Museum">V</collectionCode>
densely covered with yellowish and bluish recumbent pubescence; median plane of ventrites I and II with very fine yellowish pubescence; ventrites III and IV devoid of pubescence; ventrite
<collectionCode id="C4EE8EF11A06873CFEBE0915DA62FE6E" box="[282,301,362,385]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="81" type="Museum">V</collectionCode>
with brownish setae with semi-erect whitish setae at lateral side near the apex.
</paragraph>
<paragraph id="A24016341A06873CFF19099BDAF5FE13" blockId="6.[189,442,484,508]" box="[189,442,484,508]" pageId="6" pageNumber="81">
<emphasis id="908BCA261A06873CFF19099BDA06FE13" bold="true" box="[189,329,484,508]" pageId="6" pageNumber="81">Adult male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="EAE545BF1A06873CFF190A5EDB90FC65" pageId="6" pageNumber="81" type="diagnosis">
<paragraph id="A24016341A06873CFF190A5EDB90FC65" blockId="6.[189,669,545,906]" pageId="6" pageNumber="81">
<emphasis id="908BCA261A06873CFF190A5EDA8EFDD6" bold="true" box="[189,449,545,569]" pageId="6" pageNumber="81">Differential diagnosis.</emphasis>
The new species is unique amongst the congeners of
<taxonomicName id="65FF6DB71A06873CFDF40A3FDBA5FD99" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="81" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="908BCA261A06873CFDF40A3FD9DAFDB7" box="[592,661,576,600]" italics="true" pageId="6" pageNumber="81">Plocia</emphasis>
spp.
</taxonomicName>
in the
<collectingCountry id="DAE856A41A06873CFE900A20DAE0FD98" box="[308,431,607,631]" name="Philippines" pageId="6" pageNumber="81">Philippines</collectingCountry>
. It is distinct for having no whitish or yellowish macules in the elytra but having a transverse band of whitish pubescence in the middle, and a head including frons highly punctate. It is somewhat close to
<taxonomicName id="65FF6DB71A06873CFEF60A86D93FFCFF" authority="(Newman, 1842)" baseAuthorityName="Newman" baseAuthorityYear="1842" box="[338,624,760,784]" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="81" phylum="Arthropoda" rank="species" species="notata">
<emphasis id="908BCA261A06873CFEF60A86DAFBFCFF" box="[338,436,761,784]" italics="true" pageId="6" pageNumber="81">P. notata</emphasis>
(
<bibRefCitation id="C66E6BC51A06873CFE660A87D926FCFF" author="Newman M." box="[450,617,760,784]" pageId="6" pageNumber="81" pagination="288 - 293" refId="ref4137" refString="Newman M. 1842. Cerambycitum insularum manillarum a dom. Cuming captorum enumeratio digesta. The Entomologist, London 18: 288 - 293." type="journal article" year="1842">Newman, 1842</bibRefCitation>
)
</taxonomicName>
for having a whitish pubescence at antennomeres IV, but distinct from
<taxonomicName id="65FF6DB71A06873CFDA70B49D925FCA2" baseAuthorityName="Newman" baseAuthorityYear="1842" box="[515,618,822,845]" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="81" phylum="Arthropoda" rank="species" species="notata">
<emphasis id="908BCA261A06873CFDA70B49D925FCA2" box="[515,618,822,845]" italics="true" pageId="6" pageNumber="81">P. notata</emphasis>
</taxonomicName>
for being two-toned tomentose at antennomere III.
</paragraph>
</subSubSection>
<subSubSection id="EAE545BF1A06873CFF190BCFDA5DFB8E" pageId="6" pageNumber="81" type="etymology">
<paragraph id="A24016341A06873CFF190BCFDA5DFB8E" blockId="6.[189,668,944,1121]" pageId="6" pageNumber="81">
<emphasis id="908BCA261A06873CFF190BCFDA09FC27" bold="true" box="[189,326,944,968]" pageId="6" pageNumber="81">Etymology.</emphasis>
<taxonomicName id="65FF6DB71A06873CFEEB0BCFD947FC27" authorityName="Medina &amp; Avergonzado &amp; Loguinsa &amp; Macosang &amp; Panangcad" authorityYear="2023" box="[335,520,944,968]" class="Insecta" family="Cerambycidae" genus="Plocia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="81" phylum="Arthropoda" rank="species" species="maglanai" status="sp. nov.">
<emphasis id="908BCA261A06873CFEEB0BCFD947FC27" box="[335,520,944,968]" italics="true" pageId="6" pageNumber="81">Plocia maglanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8BB8775D1A06873CFDBC0BCED93AFC27" box="[536,629,945,968]" pageId="6" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
is named after Dr. Wenifredo Maglana, a medical doctor and a conservationist who dedicated his life to protecting watersheds and natural reserves in
<collectingRegion id="603BD8D61A06873CFE1F0C54D94FFBAD" box="[443,512,1067,1090]" country="Philippines" name="Davao" pageId="6" pageNumber="81">Davao</collectingRegion>
de Oro,
<collectingCountry id="DAE856A41A06873CFDC60C54DA41FB8E" name="Philippines" pageId="6" pageNumber="81">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="EAE545BF1A06873CFF190CF8D9C0FB52" pageId="6" pageNumber="81" type="distribution">
<paragraph id="A24016341A06873CFF190CF8D9C0FB52" blockId="6.[189,661,1159,1214]" pageId="6" pageNumber="81">
<emphasis id="908BCA261A06873CFF190CF8DA00FB70" bold="true" box="[189,335,1159,1183]" pageId="6" pageNumber="81">Distribution.</emphasis>
<collectingCountry id="DAE856A41A06873CFEF90CF8DA96FB70" box="[349,473,1159,1183]" name="Philippines" pageId="6" pageNumber="81">Philippines</collectingCountry>
: Mindanao (
<collectingRegion id="603BD8D61A06873CFDC80CF8DBABFB52" country="Philippines" name="Davao" pageId="6" pageNumber="81">Davao</collectingRegion>
de Oro, Pantukan,
<collectingRegion id="603BD8D61A06873CFE160CDAD9CCFB52" box="[434,643,1189,1213]" country="Philippines" name="Compostela" pageId="6" pageNumber="81">Compostela Valley</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="4F99845C45A1ED5AE7B1D913C72CD29D" ID-DOI="10.59893/bjc.23(2).012" ID-ISSN="1407-8619" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657859528" checkinUser="felipe" docAuthor="Barševskis, Arvīds &amp; Medina, Milton Norman" docDate="2023" docId="3D56784D2419211136646780177A4A7F" docLanguage="en" docName="BalticJColeopterol.23.2.255-258.pdf" docOrigin="Baltic Journal of Coleopterology (Basel, Switzerland) 23 (2)" docSource="http://dx.doi.org/10.1007/s00345-024-05124-9" docStyle="DocumentStyle:514B3A4252435B0AFD496C900C6D7572.1:BalticJColeop.2023-.journal_article" docStyleId="514B3A4252435B0AFD496C900C6D7572" docStyleName="BalticJColeop.2023-.journal_article" docStyleVersion="1" docTitle="Cristaphanes palawanus Barševskis &amp; Medina, 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="256" masterDocId="C16F0035241821103414677515584941" masterDocTitle="New species of the genus Cristaphanes Aurivillius, 1916 (Coleoptera: Cerambycidae) from Palawan" masterLastPageNumber="258" masterPageNumber="255" pageNumber="256" updateTime="1722691045174" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-NC-4.0">
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<paragraph id="B540C95B24192111366467801683484C" blockId="1.[624,987,245,299]" box="[624,987,245,269]" pageId="1" pageNumber="256">
<heading id="EE087E3724192111366467801683484C" bold="true" box="[624,987,245,269]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="256" reason="4">
<emphasis id="878B154924192111366467801683484C" bold="true" box="[624,987,245,269]" pageId="1" pageNumber="256">
<taxonomicName id="72FFB2D824192111366467801625484C" authorityName="Barševskis &amp; Medina" authorityYear="2023" box="[624,893,245,269]" class="Insecta" family="Cerambycidae" genus="Cristaphanes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="palawanus" status="sp. nov.">
<emphasis id="878B154924192111366467801625484C" bold="true" box="[624,893,245,269]" italics="true" pageId="1" pageNumber="256">Cristaphanes palawanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CB8A83224192111379167801683484C" box="[901,987,245,269]" pageId="1" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FDE59AD02419211136646666177A4A7F" pageId="1" pageNumber="256" type="description">
<paragraph id="B540C95B241921113664666617E7486A" blockId="1.[624,987,245,299]" box="[624,703,275,299]" pageId="1" pageNumber="256">
(
<figureCitation id="2DC4D5DE24192111366C666617EE486A" box="[632,694,275,299]" captionStart="Fig" captionStartId="1.[624,663,1131,1155]" captionTargetBox="[635,1074,314,1116]" captionTargetId="figure-316@1.[624,1085,300,1124]" captionTargetPageId="1" captionText="Fig. 1. Cristaphanes palawanus sp. nov. (holotype)" pageId="1" pageNumber="256">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="B540C95B241921113466654A177A4A7F" blockId="1.[114,593,575,830]" pageId="1" pageNumber="256">
In last months, many scientists have published new taxonomic and distribution data about the longhorn beetle fauna of the Philippine archipelago, including
<bibRefCitation id="D16EB4AA2419211135A065D717144BFB" author="Vives E." box="[436,588,674,698]" pageId="1" pageNumber="256" pagination="1 - 5" refId="ref1456" refString="Vives E. 2023 a. Especies nuevas o interesantes de longicornios de Filipinas (Pars 24). (Coleoptera, Cerambycidae). Faunitaxys, 11 (74), 2023: 1 - 5." type="journal article" year="2023">Vives (2023a</bibRefCitation>
,
<bibRefCitation id="D16EB4AA24192111346665B615EF4B9A" author="Vives E." box="[114,183,707,731]" pageId="1" pageNumber="256" pagination="18 - 32" refId="ref1494" refString="Vives E. 2023 b. Especies nuevas o interesantes de longicornios de Filipinas (Pars 28) (Coleoptera, Cerambycidae). Les Cahiers Magellanes, NS, 46: 18 - 32." type="journal article" year="2023">2023b</bibRefCitation>
, 2023c), Yokoi (2023),
<bibRefCitation id="D16EB4AA2419211135D165B614214BBD" author="Medina M. N. &amp; Vitali F. &amp; Barsevskis A." pageId="1" pageNumber="256" pagination="537 - 551" refId="ref1371" refString="Medina M. N., Vitali F., Barsevskis A., 2023. Catalog of the genus Cacia Newman (Coleoptera, Cerambycidae, Lamiinae) in the Philippines with description of two new species. Zootaxa, 5231 (5): 537 - 551." type="journal article" year="2023">Medina, Vitali, Barševskis (2023)</bibRefCitation>
and others. This shows that the longhorn beetle fauna of the Philippine archipelago is poorly studied.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="36CF52793C2A82DD339C79CB67B50EC7">Revisional notes on the genus Microhoria Chevrolat, 1877 (Insecta: Coleoptera: Anthicidae) from the Eastern Mediterranean and Turkey, with new descriptions, an annotated catalogue, and a key</mods:title>
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<taxonomicName id="1F9C4D45ED03FFB4FD1BFA2D67B3FA18" authorityName="Telnov" authorityYear="2022" box="[643,919,1463,1487]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="270" phylum="Arthropoda" rank="species" species="paxvobiscum" status="sp. nov.">
<emphasis id="EAE8EAD4ED03FFB4FD1BFA2D67B3FA18" bold="true" box="[643,919,1463,1487]" italics="true" pageId="75" pageNumber="270">Microhoria paxvobiscum</emphasis>
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</emphasis>
(
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)
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<emphasis id="EAE8EAD4ED03FFB4FD1BF9856798F9E0" bold="true" box="[643,956,1567,1591]" pageId="75" pageNumber="270">Type material designated.</emphasis>
<materialsCitation id="68F43C9BED03FFB4FC5FF985604AF941" collectionCode="FMNH" collectorName="R. Linnavuori" country="Iraq" location="Najaf" pageId="75" pageNumber="270" specimenCount="1" stateProvince="An Najaf" typeStatus="holotype">
<typeStatus id="07278864ED03FFB4FC5FF985600AF9E1" box="[967,1070,1567,1590]" pageId="75" pageNumber="270" type="holotype">Holotype</typeStatus>
male
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:
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an
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Shabakah [stroke through by hand] Waqisah [printed] 1. 5. [handwritten] 198 [printed] 0 [handwritten]
<collectorName id="75695310ED03FFB4FC4BF9E5604AF941" box="[979,1134,1663,1686]" pageId="75" pageNumber="270">R. Linnavuori</collectorName>
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</paragraph>
<paragraph id="D82336C6ED04FFB3FF0FFF0165ECFF05" blockId="76.[151,643,155,609]" pageId="76" pageNumber="271">
[
<materialsCitation id="68F43C9BED04FFB3FF06FF0165E7FF05" collectorName="Zool. Helsinki Loan No." location="Mus" pageId="76" pageNumber="271" specimenCount="1" stateProvince="Mus">
printed] //
<collectingRegion id="1A58F824ED04FFB3FE95FF016564FF65" box="[269,320,155,178]" country="Turkey" name="Mus" pageId="76" pageNumber="271">Mus</collectingRegion>
.
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C­99 1583 [printed, label yellow]
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.
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<subSubSection id="9086654DED04FFB3FF0FFF7865ADFE6E" pageId="76" pageNumber="271" type="etymology">
<paragraph id="D82336C6ED04FFB3FF0FFF7865ADFE6E" blockId="76.[151,643,155,609]" pageId="76" pageNumber="271">
<emphasis id="EAE8EAD4ED04FFB3FF0FFF78653CFF2D" bold="true" box="[151,280,226,250]" pageId="76" pageNumber="271">Etymology.</emphasis>
The name of this new taxon from war­ and unrests­hit
<collectingCountry id="A08B7656ED04FFB3FEE3FE986582FECE" box="[379,422,258,281]" name="Iraq" pageId="76" pageNumber="271">Iraq</collectingCountry>
is derived from the combination of two words of the common Christian salutation “
<emphasis id="EAE8EAD4ED04FFB3FED1FED965C4FE8D" box="[329,480,323,346]" italics="true" pageId="76" pageNumber="271">pax vobiscum</emphasis>
” (“peace with you” in Latin) to highlight the desire of Iraqi people for peace. Noun in the nominative case, standing in apposition.
</paragraph>
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<subSubSection id="9086654DED04FFB2FF0FFE506604FA79" lastPageId="77" lastPageNumber="272" pageId="76" pageNumber="271" type="description">
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<emphasis id="EAE8EAD4ED04FFB3FF0FFE506506FE35" bold="true" box="[151,290,458,482]" pageId="76" pageNumber="271">Description.</emphasis>
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male, total body length
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. Head
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long, across compound eyes
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wide, pronotum
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long, maximum width
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, elytra
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long, maximum combined width
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. Dorsum uniformly dark brown with indistinct ore shine. Tibiae and tarsi, antennomeres 15 yellowish. Venter uniformly brown. Head ovoid, about 1.21.3× as long as wide, tempora subparallel, head base broadly rounded, posterior temporal angles rounded. Compound eye moderate, longer than tempus, moderately protruding from lateral outline of head. Head dorsum moderately glossy, punctures small and sparse. Intervening spaces generally smooth and glossy, in part microreticulate, about twice as wide as punctures. Dorsal cranial setae greyish, moderately long and dense, appressed to subdecumbent. Antennae slightly enlarged in apical third. Penultimate antennomere cylindrical, longer than wide. Terminal antennomere elongate, bluntly pointed, about 2.52.6× as long as penultimate antennomere, as long combined length of antennomeres 9 10. Pronotum about 1.2× as long as wide, distinctly narrower than head across eyes, anterior margin broadly rounded, medially truncate. Pronotal disc moderately glossy, flattened in dorsal aspect. Lateral margins slightly converging in posterior half. Latero­basal pronotal fovea narrow, not supplemented with dense setae. Pronotal punctures oval, shallow, somewhat denser than those on head. Intervening spaces as wide as to twice as wide as punctures. Pronotal setae similar to as those on head. Elytra elongate, about 1.71.8× as long as wide, laterally subparallel, subtruncate at apex, dorsally flattened. Humerus broadly rounded. Apex of elytron modified, elytral surface somewhat bulged in dorsal at posterior median apical angle, slightly produced posteriad on suture. Elytral surface slightly glossy and smooth, punctures larger than those on forebody, but not much deeper. Intervening spaces generally as wide as punctures, in part distinctly microscopically wrinkled. Elytral setae yellowish, moderately long, rather dense, generally appressed, slightly curved, directed posteriad. Metathoracic wings fully developed. Legs without modifications, tibial terminal spurs paired. Tergite VII subtruncate at posterior margin. Morphological sternite VII broadly rounded at posterior margin, medially subtruncate. Tergite VIII with a row of long setae at posterior margin. Aedeagus (
<figureCitation id="40A72A43ED05FFB2FDAAFB28649AFB3E" captionStart="Fig" captionStartId="76.[151,190,1606,1629]" captionTargetBox="[153,1169,653,1575]" captionTargetId="figure-219@76.[151,1172,645,1576]" captionTargetPageId="76" captionText="Fig. 40. Microhoria paxvobiscum sp. nov. male holotype.A Habitus, dorsal view; B Forebody, dorsal view; C Spiculum gastrale; D Aedeagus, ventral view; E Apical portion of aedeagus [not to scale]." pageId="77" pageNumber="272">Fig. 40CD</figureCitation>
) cylindrical. Tegmen apex bilobate; fused baculi well­developed, extending apicad to near two thirds of tegmen; endophallic armature of four longer, strong, slightly curved spines and a group of 56 shorter, somewhat thicker spines each lobate preapically.
</paragraph>
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<paragraph id="D82336C6ED04FFB3FF0FF9DC650EF94A" blockId="76.[151,1172,1606,1693]" pageId="76" pageNumber="271">
Fig. 40.
<taxonomicName id="1F9C4D45ED04FFB3FF75F9DC65D2F98A" authorityName="Telnov" authorityYear="2022" box="[237,502,1606,1629]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="271" phylum="Arthropoda" rank="species" species="paxvobiscum" status="sp. nov.">
<emphasis id="EAE8EAD4ED04FFB3FF75F9DC65D2F98A" box="[237,502,1606,1629]" italics="true" pageId="76" pageNumber="271">Microhoria paxvobiscum</emphasis>
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<emphasis id="EAE8EAD4ED04FFB3FE63F9DC666BF989" bold="true" box="[507,591,1606,1630]" pageId="76" pageNumber="271">
<taxonomicNameLabel id="F1DB57AFED04FFB3FE63F9DC666BF989" box="[507,591,1606,1630]" pageId="76" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype. A Habitus, dorsal view; B Forebody, dorsal view; C Spiculum gastrale; D Aedeagus, ventral view; E Apical portion of aedeagus [not to scale].
</paragraph>
</caption>
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<emphasis id="EAE8EAD4ED05FFB2FFE9FA0D6571FA78" bold="true" box="[113,341,1431,1455]" pageId="77" pageNumber="272">Sexual dimorphism.</emphasis>
Female is unknow.
</paragraph>
</subSubSection>
<subSubSection id="9086654DED05FFB2FFE9FA2464F3F943" pageId="77" pageNumber="272" type="diagnosis">
<paragraph id="D82336C6ED05FFB2FFE9FA2464F3F943" blockId="77.[113,605,155,1684]" pageId="77" pageNumber="272">
<emphasis id="EAE8EAD4ED05FFB2FFE9FA246549FA01" bold="true" box="[113,365,1470,1494]" pageId="77" pageNumber="272">Differential diagnosis.</emphasis>
<taxonomicName id="1F9C4D45ED05FFB2FEECFA2564B9FA21" authorityName="Telnov" authorityYear="2022" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="272" phylum="Arthropoda" rank="species" species="paxvobiscum" status="sp. nov.">
<emphasis id="EAE8EAD4ED05FFB2FEECFA2564B9FA21" italics="true" pageId="77" pageNumber="272">Microhoria paxvobiscum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1DB57AFED05FFB2FF3BFA4464D7FA22" box="[163,243,1502,1525]" pageId="77" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="1F9C4D45ED05FFB2FE17FA45662FFA21" authorityName="Warren" authorityYear="1895" box="[399,523,1503,1526]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="272" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="EAE8EAD4ED05FFB2FE17FA45662FFA21" box="[399,523,1503,1526]" italics="true" pageId="77" pageNumber="272">M. olivacea</emphasis>
</taxonomicName>
species group and is externally very similar to a number of brown, ore shiny congeners and can only be separated from them due to the specific shape of aedeagus and the structure of the endophallic armature.
</paragraph>
</subSubSection>
<subSubSection id="9086654DED05FFB2FD1BFF01677CFF65" box="[643,856,155,179]" pageId="77" pageNumber="272" type="biology_ecology">
<paragraph id="D82336C6ED05FFB2FD1BFF01677CFF65" blockId="77.[643,1134,155,251]" box="[643,856,155,179]" pageId="77" pageNumber="272">
<emphasis id="EAE8EAD4ED05FFB2FD1BFF0166C7FF64" bold="true" box="[643,739,155,179]" pageId="77" pageNumber="272">Ecology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="9086654DED05FFB2FD1BFF59669FFF2C" pageId="77" pageNumber="272" type="distribution">
<paragraph id="D82336C6ED05FFB2FD1BFF59669FFF2C" blockId="77.[643,1134,155,251]" pageId="77" pageNumber="272">
<emphasis id="EAE8EAD4ED05FFB2FD1BFF596731FF0C" bold="true" box="[643,789,195,219]" pageId="77" pageNumber="272">Distribution.</emphasis>
Southern
<collectingCountry id="A08B7656ED05FFB2FC1FFF596796FF0D" box="[903,946,195,218]" name="Iraq" pageId="77" pageNumber="272">Iraq</collectingCountry>
(
<collectingRegion id="1A58F824ED05FFB2FC5AFF596694FF2C" country="Iraq" name="An Najaf" pageId="77" pageNumber="272">Najaf Governorate</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
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@ -0,0 +1,395 @@
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<mods:title id="0E33B8D0598A8A6BFCC279D42C33B6F9">Revisional notes on the genus Microhoria Chevrolat, 1877 (Insecta: Coleoptera: Anthicidae) from the Eastern Mediterranean and Turkey, with new descriptions, an annotated catalogue, and a key</mods:title>
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<mods:title id="301C8089F2355190B4B86E993039504A">Baltic Journal of Coleopterology</mods:title>
</mods:titleInfo>
<mods:part id="5C9D247B6B76CD97FCBF76A5DDD6E004">
<mods:date id="CB55AE1137BDA7C26F79A0D878C31792">2022</mods:date>
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<mods:classification id="75FD35CB2DDA2BE66060049405F3B91C">journal article</mods:classification>
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<subSubSection id="9086654DED0AFFBDFF0FFC2C65FCFBFA" pageId="66" pageNumber="261" type="nomenclature">
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<emphasis id="EAE8EAD4ED0AFFBDFF0FFC2C6517FC3A" bold="true" pageId="66" pageNumber="261">
<heading id="836B81AAED0AFFBDFF0FFC2C64F3FC3A" bold="true" centered="true" fontSize="10" level="2" pageId="66" pageNumber="261" reason="4">
<taxonomicName id="1F9C4D45ED0AFFBDFF0FFC2C64F3FC3A" authority="Telnov et Degiovanni" authorityName="Telnov et Degiovanni" authorityYear="2022" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="261" phylum="Arthropoda" rank="species" species="machairodus" status="sp. nov.">
<emphasis id="EAE8EAD4ED0AFFBDFF0FFC2C6592FC19" bold="true" box="[151,438,950,974]" italics="true" pageId="66" pageNumber="261">Microhoria machairodus</emphasis>
Telnov et Degiovanni
</taxonomicName>
</heading>
<taxonomicNameLabel id="F1DB57AFED0AFFBDFF46FC4F6517FC3A" box="[222,307,981,1005]" pageId="66" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="40A72A43ED0AFFBDFEDAFC4C65B4FC3A" box="[322,400,982,1005]" captionStart="Fig" captionStartId="67.[113,152,1537,1560]" captionTargetBox="[117,1131,509,1506]" captionTargetId="figure-6@67.[113,1134,503,1506]" captionTargetPageId="67" captionText="Fig. 35. Microhoria machairodus Telnov et Degiovanni sp. nov. male paratypes. A Habitus, dorsal view; B Spiculum gastrale; C Aedeagus, lateral view; D ditto, lateral view with endophallic armature inverted; E Apex of endophallic armature, inverted; FG Apical portion of aedeagus; H Apical portion of endophallic armature, not inverted [not to scale; image C courtesy Augusto Degiovanni, Bubano, Italy]." pageId="66" pageNumber="261">Fig. 35</figureCitation>
,
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)
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urn:lsid:zoobank.org:act:
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</subSubSection>
<subSubSection id="9086654DED0AFFBDFF0FFBA76766FE87" pageId="66" pageNumber="261" type="materials_examined">
<paragraph id="D82336C6ED0AFFBDFF0FFBA7655DFB03" blockId="66.[151,644,949,1692]" pageId="66" pageNumber="261">
<emphasis id="EAE8EAD4ED0AFFBDFF0FFBA765F4FB82" bold="true" box="[151,464,1085,1109]" pageId="66" pageNumber="261">Type material designated.</emphasis>
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<typeStatus id="07278864ED0AFFBDFE43FBA76666FB83" box="[475,578,1085,1108]" pageId="66" pageNumber="261" type="holotype">Holotype</typeStatus>
male
<collectionCode id="BE8DAE03ED0AFFBDFF0FFBC764F2FBA3" box="[151,214,1117,1140]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="66" pageNumber="261">NME</collectionCode>
:
<collectingCountry id="A08B7656ED0AFFBDFF46FBC7656DFBA3" box="[222,329,1117,1140]" name="Turkey" pageId="66" pageNumber="261">TURKEY</collectingCountry>
S,
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,
<location id="DD43601DED0AFFBDFE62FBC76658FBA3" LSID="urn:lsid:plazi:treatment:503587D0ED0AFFBBFF0FFC2C64C5FB75:DD43601DED0AFFBDFE62FBC76658FBA3" box="[506,636,1117,1140]" country="Turkey" latitude="36.450832" longLatPrecision="19" longitude="33.161114" name="Gulnar distr." pageId="66" pageNumber="261" stateProvince="Mersin">Gülnar distr.</location>
,
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,
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<collectingDate id="BC66E9EEED0AFFBDFDF8FBE764DEFB63" pageId="66" pageNumber="261" value="2017-06" valueMax="2017-08-31" valueMin="2017-06-00">vi viii.2017</collectingDate>
</date>
, hollow
<taxonomicName id="1F9C4D45ED0AFFBDFECEFB0465F3FB63" box="[342,471,1181,1205]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="66" pageNumber="261" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="EAE8EAD4ED0AFFBDFECEFB04658BFB62" box="[342,431,1182,1205]" italics="true" pageId="66" pageNumber="261">Quercus</emphasis>
sp.
</taxonomicName>
, leg.
<collectorName id="75695310ED0AFFBDFD89FB076643FB63" box="[529,615,1181,1204]" pageId="66" pageNumber="261">M. Avci</collectorName>
&amp;
<collectorName id="75695310ED0AFFBDFF0FFB27652AFB03" box="[151,270,1213,1236]" pageId="66" pageNumber="261">S. Göktepe</collectorName>
[printed].
</materialsCitation>
</paragraph>
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<typeStatus id="07278864ED0AFFBDFF0FFB7F64DCFB2B" box="[151,248,1253,1276]" pageId="66" pageNumber="261" type="paratype">Paratypes</typeStatus>
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.
<materialsCitation id="68F43C9BED0AFFBDFE12FB7F663BFB2B" box="[394,543,1253,1276]" collectionCode="BMNH" pageId="66" pageNumber="261" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFE12FB7F65E8FB2B" box="[394,460,1253,1276]" count="1" pageId="66" pageNumber="261" type="male">1 male</specimenCount>
<collectionCode id="BE8DAE03ED0AFFBDFE48FB7F663BFB2B" box="[464,543,1253,1276]" pageId="66" pageNumber="261">BMNH</collectionCode>
</materialsCitation>
,
<materialsCitation id="68F43C9BED0AFFBDFDBEFB7F650CFACB" collectionCode="IUIT" pageId="66" pageNumber="261" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFDBEFB7F664CFB2B" box="[550,616,1253,1276]" count="1" pageId="66" pageNumber="261" type="male">1 male</specimenCount>
&amp;
<specimenCount id="CE9AFD4FED0AFFBDFF0FFA9F64D5FACB" box="[151,241,1285,1308]" count="1" pageId="66" pageNumber="261" type="female">1 female</specimenCount>
<collectionCode id="BE8DAE03ED0AFFBDFF6EFA9F650CFACB" box="[246,296,1285,1308]" pageId="66" pageNumber="261">IUIT</collectionCode>
</materialsCitation>
,
<materialsCitation id="68F43C9BED0AFFBDFEABFA9F6592FACB" box="[307,438,1285,1308]" collectionCode="DTC" pageId="66" pageNumber="261" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFEABFA9F655CFACB" box="[307,376,1285,1308]" count="1" pageId="66" pageNumber="261" type="male">1 male</specimenCount>
<collectionCode id="BE8DAE03ED0AFFBDFEE5FA9F6592FACB" box="[381,438,1285,1308]" pageId="66" pageNumber="261">DTC</collectionCode>
</materialsCitation>
: same label as
<typeStatus id="07278864ED0AFFBDFDD3FA9F64E3FAEB" pageId="66" pageNumber="261" type="holotype">holotype</typeStatus>
;
<materialsCitation id="68F43C9BED0AFFBDFF48FABF664DFA4B" collectingDate="1993-07-05" collectorName="J. Jelinek &amp; G. Sama" country="Turkey" location="Iel Aydinlar" municipality="Uzuncaburc" pageId="66" pageNumber="261" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFF48FABF6531FAEB" box="[208,277,1317,1340]" count="1" pageId="66" pageNumber="261" type="male">1 male</specimenCount>
&amp; 1 femaleADC:
<collectingCountry id="A08B7656ED0AFFBDFE48FABF663DFAEB" box="[464,537,1317,1340]" name="Turkey" pageId="66" pageNumber="261">Turkey</collectingCountry>
, 1.VII.83,
<collectingMunicipality id="3847ACBCED0AFFBDFF0FFADF653CFA8B" box="[151,280,1349,1372]" pageId="66" pageNumber="261">Uzuncaburc</collectingMunicipality>
<locationDeviation id="C9912FE4ED0AFFBDFEBAFADF65C2FA8B" box="[290,486,1349,1372]" location="Silifke" pageId="66" pageNumber="261">
<quantity id="1F649B23ED0AFFBDFEBAFADF654CFA8B" box="[290,360,1349,1372]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="66" pageNumber="261" unit="km" value="30.0">30 km</quantity>
NE Silifke
</locationDeviation>
<collectorName id="75695310ED0AFFBDFE68FADF6673FA8B" box="[496,599,1349,1372]" pageId="66" pageNumber="261">J. Jelinek</collectorName>
lgt. [printed]; 1 maleADC:TR [
<collectingCountry id="A08B7656ED0AFFBDFE3FFAFF65D1FAAB" box="[423,501,1381,1404]" name="Turkey" pageId="66" pageNumber="261">Turkey</collectingCountry>
]
<location id="DD43601DED0AFFBDFD8FFAFF6497FA4B" LSID="urn:lsid:plazi:treatment:503587D0ED0AFFBBFF0FFC2C64C5FB75:DD43601DED0AFFBDFD8FFAFF6497FA4B" country="Turkey" municipality="Uzuncaburc" name="Iel Aydinlar" pageId="66" pageNumber="261">Iel Aydinlar</location>
m 1000
<collectorName id="75695310ED0AFFBDFE8DFA1F654BFA4B" box="[277,367,1413,1436]" pageId="66" pageNumber="261">G. Sama</collectorName>
4/
<date id="AC221006ED0AFFBDFE16FA1F6624FA4B" box="[398,512,1413,1436]" pageId="66" pageNumber="261" value="1993-07-05">
<collectingDate id="BC66E9EEED0AFFBDFE16FA1F6624FA4B" box="[398,512,1413,1436]" pageId="66" pageNumber="261" value="1993-07-05">5.VII.1993</collectingDate>
</date>
[printed]
</materialsCitation>
;
<materialsCitation id="68F43C9BED0AFFBDFDEDFA1F653DFA6B" collectionCode="ADC" country="Turkey" pageId="66" pageNumber="261" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFDEDFA1F64F1FA6B" count="2" pageId="66" pageNumber="261" type="male">2 males</specimenCount>
<collectionCode id="BE8DAE03ED0AFFBDFF45FA3F653DFA6B" box="[221,281,1445,1468]" pageId="66" pageNumber="261">ADC</collectionCode>
</materialsCitation>
:
<materialsCitation id="68F43C9BED0AFFBDFEBEFA3F6539F9CB" collectingDate="2010-05-24" collectorName="M. Uliana" country="Turkey" elevation="1080" latitude="36.744724" location="Aktepe" longLatPrecision="19" longitude="36.429447" pageId="66" pageNumber="261" specimenCount="1" typeStatus="paratype">
<collectingCountry id="A08B7656ED0AFFBDFEBEFA3F6555FA6B" box="[294,369,1445,1468]" name="Turkey" pageId="66" pageNumber="261">Turkey</collectingCountry>
, prov., Hatay,
<quantity id="1F649B23ED0AFFBDFD8CFA3F6669FA6B" box="[532,589,1445,1468]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="66" pageNumber="261" unit="km" value="7.0">7 km</quantity>
NW
<location id="DD43601DED0AFFBDFF0FFA5F64C3FA0B" LSID="urn:lsid:plazi:treatment:503587D0ED0AFFBBFF0FFC2C64C5FB75:DD43601DED0AFFBDFF0FFA5F64C3FA0B" box="[151,231,1477,1500]" country="Turkey" latitude="36.744724" longLatPrecision="19" longitude="36.429447" name="Aktepe" pageId="66" pageNumber="261">Aktepe</location>
,
<quantity id="1F649B23ED0AFFBDFF6EFA5F656FFA0B" box="[246,331,1477,1500]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="66" pageNumber="261" unit="m" value="1080.0">
<elevation id="53B1D1F5ED0AFFBDFF6EFA5F656FFA0B" box="[246,331,1477,1500]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="66" pageNumber="261" unit="m" value="1080.0">1080 m</elevation>
</quantity>
<geoCoordinate id="BDA85001ED0AFFBDFECDFA5F65CCFA0B" box="[341,488,1477,1500]" degrees="36" direction="north" minutes="44" orientation="latitude" pageId="66" pageNumber="261" precision="15" seconds="41" value="36.744724">36°4441 N</geoCoordinate>
,
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<date id="AC221006ED0AFFBDFF0FFA7F6525FA2B" box="[151,257,1509,1532]" pageId="66" pageNumber="261" value="2010-05-24">
<collectingDate id="BC66E9EEED0AFFBDFF0FFA7F6525FA2B" box="[151,257,1509,1532]" pageId="66" pageNumber="261" value="2010-05-24">24.V.2010</collectingDate>
</date>
leg.
<collectorName id="75695310ED0AFFBDFEB7FA7F65B2FA2B" box="[303,406,1509,1532]" pageId="66" pageNumber="261">M. Uliana</collectorName>
[printed]; 1 maleADC:
<collectingCountry id="A08B7656ED0AFFBDFF0FF99F64C5F9CB" box="[151,225,1541,1564]" name="Turkey" pageId="66" pageNumber="261">Turkey</collectingCountry>
, prov
</materialsCitation>
.,
<materialsCitation id="68F43C9BED0AFFBDFEA9F99F64DCF98B" collectingDate="2010-05-25" collectorName="M. Uliana" country="Turkey" elevation="1625" latitude="37.016666" location="Arslankoy" longLatPrecision="1184" longitude="34.25" pageId="66" pageNumber="261" specimenCount="1" stateProvince="Mersin" typeStatus="paratype">
<collectingRegion id="1A58F824ED0AFFBDFEA9F99F655BF9CB" box="[305,383,1541,1564]" country="Turkey" name="Mersin" pageId="66" pageNumber="261">Mersin</collectingRegion>
, env.,
<location id="DD43601DED0AFFBDFE58F99F6609F9CB" LSID="urn:lsid:plazi:treatment:503587D0ED0AFFBBFF0FFC2C64C5FB75:DD43601DED0AFFBDFE58F99F6609F9CB" box="[448,557,1541,1564]" country="Turkey" latitude="37.016666" longLatPrecision="1184" longitude="34.25" name="Arslankoy" pageId="66" pageNumber="261" stateProvince="Mersin">Arslankoy</location>
,
<geoCoordinate id="BDA85001ED0AFFBDFDA2F99F6489F9EB" degrees="37" direction="north" minutes="01" orientation="latitude" pageId="66" pageNumber="261" precision="925" value="37.016666">37°01 N</geoCoordinate>
,
<geoCoordinate id="BDA85001ED0AFFBDFF2EF9BF6531F9EB" box="[182,277,1573,1596]" degrees="34" direction="east" minutes="15" orientation="longitude" pageId="66" pageNumber="261" precision="925" value="34.25">34°15E</geoCoordinate>
,
<quantity id="1F649B23ED0AFFBDFE86F9BF654BF9EB" box="[286,367,1573,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="66" pageNumber="261" unit="m" value="1625.0">
<elevation id="53B1D1F5ED0AFFBDFE86F9BF654BF9EB" box="[286,367,1573,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="66" pageNumber="261" unit="m" value="1625.0">1625 m</elevation>
</quantity>
,
<date id="AC221006ED0AFFBDFEE0F9BF65C2F9EB" box="[376,486,1573,1596]" pageId="66" pageNumber="261" value="2010-05-25">
<collectingDate id="BC66E9EEED0AFFBDFEE0F9BF65C2F9EB" box="[376,486,1573,1596]" pageId="66" pageNumber="261" value="2010-05-25">25.V.2010</collectingDate>
</date>
leg.
<collectorName id="75695310ED0AFFBDFD8FF9BF66A7F9EB" box="[535,643,1573,1596]" pageId="66" pageNumber="261">M. Uliana</collectorName>
[printed]
</materialsCitation>
;
<materialsCitation id="68F43C9BED0AFFBDFE9BF9DF65AAF98B" box="[259,398,1605,1628]" collectionCode="ADC" country="Turkey" pageId="66" pageNumber="261" specimenCount="1" specimenCount-male="1" stateProvince="Mersin" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFE9BF9DF6568F98B" box="[259,332,1605,1628]" count="1" pageId="66" pageNumber="261" type="male">1 male</specimenCount>
<collectionCode id="BE8DAE03ED0AFFBDFECAF9DF65AAF98B" box="[338,398,1605,1628]" pageId="66" pageNumber="261">ADC</collectionCode>
</materialsCitation>
:
<materialsCitation id="68F43C9BED0AFFBDFE00F9DF64DCF94B" collectingDate="2011-06-02" collectorName="F. Angelini" country="Turkey" location="Km" municipality="Mut" pageId="66" pageNumber="261" specimenCount="1" stateProvince="Mersin" typeStatus="paratype">
<collectingCountry id="A08B7656ED0AFFBDFE00F9DF65CCF98B" box="[408,488,1605,1628]" name="Turkey" pageId="66" pageNumber="261">Turkey</collectingCountry>
:
<collectingRegion id="1A58F824ED0AFFBDFE6BF9DF665FF98B" box="[499,635,1605,1628]" country="Turkey" name="Mersin" pageId="66" pageNumber="261">Mersin prov.</collectingRegion>
,
<collectingMunicipality id="3847ACBCED0AFFBDFF0FF9FF64E2F9AB" box="[151,198,1637,1660]" pageId="66" pageNumber="261">Mut</collectingMunicipality>
,
<location id="DD43601DED0AFFBDFF48F9FF64DCF9AB" LSID="urn:lsid:plazi:treatment:503587D0ED0AFFBBFF0FFC2C64C5FB75:DD43601DED0AFFBDFF48F9FF64DCF9AB" box="[208,248,1637,1660]" country="Turkey" municipality="Mut" name="Km" pageId="66" pageNumber="261" stateProvince="Mersin">Km</location>
10­ 15 N,
<date id="AC221006ED0AFFBDFEFAF9FF65FFF9AB" box="[354,475,1637,1660]" pageId="66" pageNumber="261" value="2011-06-02">
<collectingDate id="BC66E9EEED0AFFBDFEFAF9FF65FFF9AB" box="[354,475,1637,1660]" pageId="66" pageNumber="261" value="2011-06-02">02.VI.2011</collectingDate>
</date>
<collectorName id="75695310ED0AFFBDFE78F9FF6671F9AB" box="[480,597,1637,1660]" pageId="66" pageNumber="261">F. Angelini</collectorName>
leg. [printed]
</materialsCitation>
;
<materialsCitation id="68F43C9BED0AFFBDFE9BF91F6598F94B" box="[259,444,1669,1692]" collectionCode="BMNH" country="Turkey" pageId="66" pageNumber="261" specimenCount="10" specimenCount-male="10" stateProvince="Mersin" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFE9BF91F6546F94B" box="[259,354,1669,1692]" count="10" pageId="66" pageNumber="261" type="male">10 males</specimenCount>
<collectionCode id="BE8DAE03ED0AFFBDFEF1F91F6598F94B" box="[361,444,1669,1692]" pageId="66" pageNumber="261">BMNH</collectionCode>
</materialsCitation>
,
<materialsCitation id="68F43C9BED0AFFBDFE5EF91F6643F94B" box="[454,615,1669,1692]" collectionCode="NME" country="Turkey" pageId="66" pageNumber="261" specimenCount="10" specimenCount-male="10" stateProvince="Mersin" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFE5EF91F6601F94B" box="[454,549,1669,1692]" count="10" pageId="66" pageNumber="261" type="male">10 males</specimenCount>
<collectionCode id="BE8DAE03ED0AFFBDFDB4F91F6643F94B" box="[556,615,1669,1692]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="66" pageNumber="261">NME</collectionCode>
</materialsCitation>
&amp;
<materialsCitation id="68F43C9BED0AFFBDFD30FF016766FF65" box="[680,834,155,178]" collectionCode="DTC" country="Turkey" pageId="66" pageNumber="261" specimenCount="8" specimenCount-male="8" stateProvince="Mersin" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFD30FF0166DAFF65" box="[680,766,155,178]" count="8" pageId="66" pageNumber="261" type="male">8 males</specimenCount>
<collectionCode id="BE8DAE03ED0AFFBDFC90FF016766FF65" box="[776,834,155,178]" pageId="66" pageNumber="261">DTC</collectionCode>
</materialsCitation>
:
<materialsCitation id="68F43C9BED0AFFBDFCD7FF016019FF26" collectingDate="2011-06-07" collectorName="A. Napolov &amp; I. Roma" country="Turkey" elevation="490" latitude="36.704445" location="Erdemli" longLatPrecision="19" longitude="34.29111" pageId="66" pageNumber="261" specimenCount="1" typeStatus="paratype">
<collectingCountry id="A08B7656ED0AFFBDFCD7FF01679CFF65" box="[847,952,155,178]" name="Turkey" pageId="66" pageNumber="261">TURKEY</collectingCountry>
,
<quantity id="1F649B23ED0AFFBDFC51FF01602BFF65" box="[969,1039,155,178]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="66" pageNumber="261" unit="km" value="10.0">10 km</quantity>
N
<location id="DD43601DED0AFFBDFBAFFF0160B4FF65" LSID="urn:lsid:plazi:treatment:503587D0ED0AFFBBFF0FFC2C64C5FB75:DD43601DED0AFFBDFBAFFF0160B4FF65" box="[1079,1168,155,178]" country="Turkey" latitude="36.704445" longLatPrecision="19" longitude="34.29111" name="Erdemli" pageId="66" pageNumber="261">Erdemli</location>
,
<geoCoordinate id="BDA85001ED0AFFBDFD30FF206716FF06" box="[680,818,186,209]" degrees="36" direction="north" minutes="42" orientation="latitude" pageId="66" pageNumber="261" precision="15" seconds="16" value="36.704445">36°4216 N</geoCoordinate>
,
<geoCoordinate id="BDA85001ED0AFFBDFCA3FF2067E5FF06" box="[827,961,186,209]" degrees="34" direction="east" minutes="17" orientation="longitude" pageId="66" pageNumber="261" precision="15" seconds="28" value="34.29111">34°1728E</geoCoordinate>
,
<quantity id="1F649B23ED0AFFBDFC53FF20602AFF06" box="[971,1038,186,209]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="66" pageNumber="261" unit="m" value="490.0">
<elevation id="53B1D1F5ED0AFFBDFC53FF20602AFF06" box="[971,1038,186,209]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="66" pageNumber="261" unit="m" value="490.0">490 m</elevation>
</quantity>
,
<date id="AC221006ED0AFFBDFB8FFF2060B4FF06" box="[1047,1168,186,209]" pageId="66" pageNumber="261" value="2011-06-07">
<collectingDate id="BC66E9EEED0AFFBDFB8FFF2060B4FF06" box="[1047,1168,186,209]" pageId="66" pageNumber="261" value="2011-06-07">07.VI.2011</collectingDate>
</date>
, leg.
<collectorName id="75695310ED0AFFBDFD4FFF406772FF26" box="[727,854,218,241]" pageId="66" pageNumber="261">A. Napolov</collectorName>
&amp;
<collectorName id="75695310ED0AFFBDFCE4FF4067F0FF26" box="[892,980,218,241]" pageId="66" pageNumber="261">I. Roma</collectorName>
[printed]
</materialsCitation>
;
<materialsCitation id="68F43C9BED0AFFBDFBD2FF4066C5FEC6" collectionCode="DTC" country="Turkey" pageId="66" pageNumber="261" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="CE9AFD4FED0AFFBDFBD2FF4060B0FF26" box="[1098,1172,218,241]" count="1" pageId="66" pageNumber="261" type="male">1 male</specimenCount>
<collectionCode id="BE8DAE03ED0AFFBDFD30FF6066C5FEC6" box="[680,737,250,273]" pageId="66" pageNumber="261">DTC</collectionCode>
</materialsCitation>
:
<materialsCitation id="68F43C9BED0AFFBDFD74FF60671AFE87" collectingDate="2011-06-03" collectionCode="IEL" collectorName="Lg. &amp; Napolov &amp; Roma" country="Turkey" elevation="425" location="6 - 7 km NW Erdemli" pageId="66" pageNumber="261" specimenCount="1" typeStatus="paratype">
<collectingCountry id="A08B7656ED0AFFBDFD74FF60677CFEC6" box="[748,856,250,273]" name="Turkey" pageId="66" pageNumber="261">TURKEY</collectingCountry>
S, (
<collectionCode id="BE8DAE03ED0AFFBDFC05FF6067ECFEC6" box="[925,968,250,273]" pageId="66" pageNumber="261">IEL</collectionCode>
)
<location id="DD43601DED0AFFBDFC42FF6066C3FEE6" LSID="urn:lsid:plazi:treatment:503587D0ED0AFFBBFF0FFC2C64C5FB75:DD43601DED0AFFBDFC42FF6066C3FEE6" country="Turkey" name="6 - 7 km NW Erdemli" pageId="66" pageNumber="261">
<quantity id="1F649B23ED0AFFBDFC42FF60600BFEC6" box="[986,1071,250,273]" metricMagnitude="3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="66" pageNumber="261" unit="km" value="6.5" valueMax="7.0" valueMin="6.0">67 km</quantity>
NW Erdemli
</location>
, h=
<quantity id="1F649B23ED0AFFBDFC97FE8067A0FEE6" box="[783,900,282,305]" metricMagnitude="2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="66" pageNumber="261" unit="m" value="425.0" valueMax="450.0" valueMin="400.0">
<elevation id="53B1D1F5ED0AFFBDFC97FE8067A0FEE6" box="[783,900,282,305]" metricMagnitude="2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="66" pageNumber="261" unit="m" value="425.0" valueMax="450.0" valueMin="400.0">400­ 450 m</elevation>
</quantity>
,
<date id="AC221006ED0AFFBDFC17FE8067D9FEE6" box="[911,1021,282,305]" pageId="66" pageNumber="261" value="2011-06-03">
<collectingDate id="BC66E9EEED0AFFBDFC17FE8067D9FEE6" box="[911,1021,282,305]" pageId="66" pageNumber="261" value="2011-06-03">3.VI.2011</collectingDate>
</date>
,
<collectorName id="75695310ED0AFFBDFB9FFE806008FEE6" box="[1031,1068,282,305]" pageId="66" pageNumber="261">Lg.</collectorName>
A.
<collectorName id="75695310ED0AFFBDFBC9FE8066EEFE87" pageId="66" pageNumber="261">Napolov</collectorName>
&amp; I.
<collectorName id="75695310ED0AFFBDFD60FEA3671AFE87" box="[760,830,313,336]" pageId="66" pageNumber="261">Roma</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9086654DED0AFFBDFD30FEFA605EFDC1" pageId="66" pageNumber="261" type="etymology">
<paragraph id="D82336C6ED0AFFBDFD30FEFA605EFDC1" blockId="66.[680,1173,155,1684]" pageId="66" pageNumber="261">
<emphasis id="EAE8EAD4ED0AFFBDFD30FEFA670DFEAF" bold="true" box="[680,809,352,376]" pageId="66" pageNumber="261">Etymology.</emphasis>
Named after
<emphasis id="EAE8EAD4ED0AFFBDFC26FEFB606FFEAF" box="[958,1099,353,376]" italics="true" pageId="66" pageNumber="261">Machairodus</emphasis>
Kaup, 1833, the extinct genus of enigmatic sabretoothed felids of the late Miocene. The genus name derived from the combination of Greek “
<emphasis id="EAE8EAD4ED0AFFBDFD2CFE456737FE20" bold="true" box="[692,787,479,503]" pageId="66" pageNumber="261">macai´ra</emphasis>
” (knife) + “
<emphasis id="EAE8EAD4ED0AFFBDFC0EFE4567F1FE20" bold="true" box="[918,981,479,503]" pageId="66" pageNumber="261">o´dou´ j</emphasis>
” (tooth). Noun in the nominative case, standing in apposition.
</paragraph>
</subSubSection>
<subSubSection id="9086654DED0AFFBBFD30FDBC64CEFD72" lastPageId="68" lastPageNumber="263" pageId="66" pageNumber="261" type="description">
<paragraph id="D82336C6ED0AFFBCFD30FDBC604AFEA5" blockId="66.[680,1173,155,1684]" lastBlockId="67.[643,1134,155,466]" lastPageId="67" lastPageNumber="262" pageId="66" pageNumber="261">
<emphasis id="EAE8EAD4ED0AFFBDFD30FDBC6717FDE9" bold="true" box="[680,819,550,574]" pageId="66" pageNumber="261">Description.</emphasis>
<typeStatus id="07278864ED0AFFBDFCA1FDBD67B8FDE9" box="[825,924,551,574]" pageId="66" pageNumber="261" type="holotype">Holotype</typeStatus>
male, total body length
<quantity id="1F649B23ED0AFFBDFD30FDDC672DFD8A" box="[680,777,582,605]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="66" pageNumber="261" unit="mm" value="2.35">2.35 mm</quantity>
. Head
<quantity id="1F649B23ED0AFFBDFCD5FDDC67B9FD8A" box="[845,925,582,605]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="66" pageNumber="261" unit="mm" value="0.5">0.5 mm</quantity>
long, across compound eyes
<quantity id="1F649B23ED0AFFBDFD42FDFC6703FDAA" box="[730,807,614,637]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="66" pageNumber="261" unit="mm" value="0.5">0.5 mm</quantity>
wide, pronotum
<quantity id="1F649B23ED0AFFBDFC57FDFC6038FDAA" box="[975,1052,614,637]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="66" pageNumber="261" unit="mm" value="0.4">0.4 mm</quantity>
long, maximum width
<quantity id="1F649B23ED0AFFBDFCBFFD1C675DFD4A" box="[807,889,646,669]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="66" pageNumber="261" unit="mm" value="0.4">0.4 mm</quantity>
, elytra
<quantity id="1F649B23ED0AFFBDFC5DFD1C6030FD4A" box="[965,1044,646,669]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="66" pageNumber="261" unit="mm" value="1.3">1.3 mm</quantity>
long, maximum combined width
<quantity id="1F649B23ED0AFFBDFC05FD3F67D7FD6B" box="[925,1011,677,700]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="66" pageNumber="261" unit="mm" value="0.8">0.8 mm</quantity>
. Selected male
<typeStatus id="07278864ED0AFFBDFD30FD5F6729FD0B" box="[680,781,709,732]" pageId="66" pageNumber="261" type="paratype">paratypes</typeStatus>
<quantity id="1F649B23ED0AFFBDFC8AFD5F67B5FD0B" box="[786,913,709,732]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="66" pageNumber="261" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.22.4 mm</quantity>
, female
<typeStatus id="07278864ED0AFFBDFC7EFD5F6064FD0B" box="[998,1088,709,732]" pageId="66" pageNumber="261" type="paratype">paratype</typeStatus>
<quantity id="1F649B23ED0AFFBDFBDDFD5F60B7FD0B" box="[1093,1171,709,732]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="66" pageNumber="261" unit="mm" value="2.3">2.3 mm</quantity>
long. Dorsum and venter uniformly black­brown. Antennae (except darkened antennomeres 8 11), palps and legs yellowish brown (femora only slightly darker than tibiae). Head slightly ovoid, about 1.11.2× as long as wide, head base rounded in one broad arc with tempora, posterior temporal angles rounded. Compound eye moderate, about as long as tempus, moderately protruding from lateral outline of head. Head dorsum moderately glossy, punctures distinct, moderately deep. Intervening spaces generally smooth and glossy, as wide as to twice as wide as punctures. Dorsal cranial setae greyish to yellowish, sparse, inconspicuous.Antennae slightly enlarged in apical third. Penultimate antennomere widened distally, nearly as long as wide. Terminal antennomere elongate, bluntly pointed, about 2.5× as long as penultimate antennomere, about 1.11.2× as long combined length of antennomeres 910. Pronotum about as long as wide, distinctly narrower than head across eyes, subtruncate at anterior margin. Pronotal disc moderately glossy, flattened in dorsal aspect. Lateral margins moderately converging in posterior half. Latero­basal pronotal fovea broad, laterally supplemented with moderately long, curved, whitish setae. Punctures and dorsal setae similar as those on head, punctures somewhat denser on median part of pronotal disc. Intervening spaces about as wide as punctures. Elytra elongate, about 1.71.8× as long as wide, laterally parallel­sided or broadly rounded, broadly rounded at apex, dorsally flattened, with a feeble postbasal transverse impression. Humerus broadly rounded.Apex of elytron modified, with elongate, acutely pointed, narrow denticle at opening of gland channel. Surface glossy and smooth, punctures somewhat larger than those on forebody, shallow. Intervening spaces generally as wide as to 3× as wide as punctures. Elytral setae greyish to yellowish, moderately long and dense, generally slightly subdecumbent to appressed, directed posteriad.Metathoracic wings fully developed. Legs without modifications, tibial terminal spurs paired. Tergite VII truncate at posterior margin. Morphological sternite VII broadly rounded at posterior margin.Aedeagus
</paragraph>
<paragraph id="D82336C6ED0BFFBBFD1BFEE165F6FEE4" blockId="67.[643,1134,155,466]" lastBlockId="68.[151,643,155,1186]" lastPageId="68" lastPageNumber="263" pageId="67" pageNumber="262">
(
<figureCitation id="40A72A43ED0BFFBCFD12FEE16727FE45" box="[650,771,379,402]" captionStart="Fig" captionStartId="67.[113,152,1537,1560]" captionTargetBox="[117,1131,509,1506]" captionTargetId="figure-6@67.[113,1134,503,1506]" captionTargetPageId="67" captionText="Fig. 35. Microhoria machairodus Telnov et Degiovanni sp. nov. male paratypes. A Habitus, dorsal view; B Spiculum gastrale; C Aedeagus, lateral view; D ditto, lateral view with endophallic armature inverted; E Apex of endophallic armature, inverted; FG Apical portion of aedeagus; H Apical portion of endophallic armature, not inverted [not to scale; image C courtesy Augusto Degiovanni, Bubano, Italy]." pageId="67" pageNumber="262">Fig. 35CH</figureCitation>
) short and broad, tegmen apex short, hooked; fused baculi inconspicuous, extending apicad to third length of tegmen; endophallic armature of two strong, sabre­like curved spines and a double row of acute angulate, small denticles. Two additional, long, weakly chitinous spines are comparatively short in some specimens, including the
<typeStatus id="07278864ED0CFFBBFEF5FE8665EAFEE4" box="[365,462,284,307]" pageId="68" pageNumber="263" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="8CE3664EED0BFFBCFFE9F99B667DF94F" pageId="67" pageNumber="262" startId="67.[113,152,1537,1560]" targetBox="[117,1131,509,1506]" targetPageId="67" targetType="figure">
<paragraph id="D82336C6ED0BFFBCFFE9F99B667DF94F" blockId="67.[113,1134,1536,1688]" pageId="67" pageNumber="262">
Fig. 35.
<taxonomicName id="1F9C4D45ED0BFFBCFF56F99B66EAF9CF" authority="Telnov et Degiovanni" authorityName="Telnov et Degiovanni" authorityYear="2022" box="[206,718,1537,1560]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="262" phylum="Arthropoda" rank="species" species="machairodus" status="sp. nov.">
<emphasis id="EAE8EAD4ED0BFFBCFF56F99B65FEF9CF" box="[206,474,1537,1560]" italics="true" pageId="67" pageNumber="262">Microhoria machairodus</emphasis>
Telnov et Degiovanni
</taxonomicName>
<emphasis id="EAE8EAD4ED0BFFBCFD4EF99A6709F9CF" bold="true" box="[726,813,1536,1560]" pageId="67" pageNumber="262">
<taxonomicNameLabel id="F1DB57AFED0BFFBCFD4EF99A6709F9CF" box="[726,813,1536,1560]" pageId="67" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male paratypes. A Habitus, dorsal view; B Spiculum gastrale; C Aedeagus, lateral view; D ditto, lateral view with endophallic armature inverted; E Apex of endophallic armature, inverted; FG Apical portion of aedeagus; H Apical portion of endophallic armature, not inverted [not to scale; image C courtesy Augusto Degiovanni, Bubano, Italy].
</paragraph>
</caption>
<paragraph id="D82336C6ED0CFFBBFF0FFEDE654CFE6B" blockId="68.[151,643,155,1186]" pageId="68" pageNumber="263">
<emphasis id="EAE8EAD4ED0CFFBBFF0FFEDE6550FE8B" bold="true" box="[151,372,324,348]" pageId="68" pageNumber="263">Sexual dimorphism.</emphasis>
Female is externally similar to male, elytron apex without modifications, rounded. Female tergite VII broadly rounded at posterior margin.
</paragraph>
<paragraph id="D82336C6ED0CFFBBFF0FFE5664CEFD72" blockId="68.[151,643,155,1186]" pageId="68" pageNumber="263">
<emphasis id="EAE8EAD4ED0CFFBBFF0FFE566588FE33" bold="true" box="[151,428,460,484]" pageId="68" pageNumber="263">Interspecific variability.</emphasis>
The number of the pale antennomeres varies, in some specimens antennomeres one or one and two are dark, antennomeres 35 or 37 darkened. Intensity of dorsal punctures on forebody varies, punctures stronger and denser in some paratypic specimens.
</paragraph>
</subSubSection>
<subSubSection id="9086654DED0CFFBBFF0FFD2F64EBFC58" pageId="68" pageNumber="263" type="diagnosis">
<paragraph id="D82336C6ED0CFFBBFF0FFD2F64EBFC58" blockId="68.[151,643,155,1186]" pageId="68" pageNumber="263">
<emphasis id="EAE8EAD4ED0CFFBBFF0FFD2F65BDFD1A" bold="true" box="[151,409,693,717]" pageId="68" pageNumber="263">Differential diagnosis.</emphasis>
<taxonomicName id="1F9C4D45ED0CFFBBFE3CFD2C6598FD3A" authority="Telnov &amp; Degiovanni" authorityName="Telnov &amp; Degiovanni" authorityYear="2022" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="263" phylum="Arthropoda" rank="species" species="machairodus" status="sp. nov.">
<emphasis id="EAE8EAD4ED0CFFBBFE3CFD2C64F7FD3A" italics="true" pageId="68" pageNumber="263">Microhoria machairodus</emphasis>
Telnov &amp; Degiovanni
</taxonomicName>
<taxonomicNameLabel id="F1DB57AFED0CFFBBFE59FD4C6636FD3A" box="[449,530,726,749]" pageId="68" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="1F9C4D45ED0CFFBBFF24FD6D6560FCD9" authorityName="P. Bonadona" baseAuthorityName="W. L. E. Schmidt" baseAuthorityYear="1842" box="[188,324,759,782]" class="Insecta" family="Anthicidae" genus="Microhoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="263" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="EAE8EAD4ED0CFFBBFF24FD6D6560FCD9" box="[188,324,759,782]" italics="true" pageId="68" pageNumber="263">M. terminata</emphasis>
</taxonomicName>
species group and is externally very similar to a number of uniformly dark congeners and can only be differentiated due to the peculiar endophallic armature of the male aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="9086654DED0CFFBBFF0FFC3A6516FB8D" pageId="68" pageNumber="263" type="biology_ecology">
<paragraph id="D82336C6ED0CFFBBFF0FFC3A6516FB8D" blockId="68.[151,643,155,1186]" pageId="68" pageNumber="263">
<emphasis id="EAE8EAD4ED0CFFBBFF0FFC3A64D7FC6F" bold="true" box="[151,243,928,952]" pageId="68" pageNumber="263">Ecology.</emphasis>
Collected using flight interception traps installed on trunks of hollow veteran
<taxonomicName id="1F9C4D45ED0CFFBBFDB1FC5B6492FC2F" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="68" pageNumber="263" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="EAE8EAD4ED0CFFBBFDB1FC5B66A7FC0F" box="[553,643,961,984]" italics="true" pageId="68" pageNumber="263">Quercus</emphasis>
sp.
</taxonomicName>
trees at elevations of
<quantity id="1F649B23ED0CFFBBFE0FFC7B660DFC2F" box="[407,553,993,1016]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="68" pageNumber="263" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">14001500 m</quantity>
(
<figureCitation id="40A72A43ED0CFFBBFDAEFC7B64EFFBCE" captionStart="Fig" captionStartId="109.[113,152,1601,1624]" captionTargetBox="[287,959,151,1571]" captionTargetId="figure-44@109.[287,959,151,1571]" captionTargetPageId="109" captionText="Fig. 55. Veteran tree­connected habitats of Microhoria in Turkey.A Sampling plot near Antakya, Turkey; BC Sampling plots near Erdemli, Turkey [images courtesy Nicklas Jansson, Linköping University, Linköping, Sweden]." pageId="68" pageNumber="263">Fig. 55 BC</figureCitation>
). Additionally, numerous specimens collected from lower vegetation and tree foliage at
<quantity id="1F649B23ED0CFFBBFF2AFBD9650AFB8D" box="[178,302,1091,1114]" metricMagnitude="2" metricUnit="m" metricValue="4.45" metricValueMax="4.9" metricValueMin="4.0" pageId="68" pageNumber="263" unit="m" value="445.0" valueMax="490.0" valueMin="400.0">
400
<specimenCount id="CE9AFD4FED0CFFBBFF7FFBD9650AFB8D" box="[231,302,1091,1114]" count="490" pageId="68" pageNumber="263" type="generic">490 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="9086654DED0CFFBBFF0FFBF064C5FB75" pageId="68" pageNumber="263" type="distribution">
<paragraph id="D82336C6ED0CFFBBFF0FFBF064C5FB75" blockId="68.[151,643,155,1186]" pageId="68" pageNumber="263">
<emphasis id="EAE8EAD4ED0CFFBBFF0FFBF0650FFB55" bold="true" box="[151,299,1130,1154]" pageId="68" pageNumber="263">Distribution.</emphasis>
Southern
<collectingCountry id="A08B7656ED0CFFBBFE07FBF165CFFB55" box="[415,491,1131,1154]" name="Turkey" pageId="68" pageNumber="263">Turkey</collectingCountry>
(
<collectingRegion id="1A58F824ED0CFFBBFE65FBF164F1FB75" country="Turkey" name="Mersin" pageId="68" pageNumber="263">Mersin Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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