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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.129.34988" ID-GBIF-Dataset="7a3be85e-9a8f-4749-9984-1e60c69e67d0" ID-PMC="PMC6717120" ID-Pensoft-Pub="1314-2003-129-1" ID-Pensoft-UUID="BF57C6B3C3065AEE9B4B3D47189C908F" ID-PubMed="31523157" ID-Zenodo-Dep="3382366" ModsDocID="1314-2003-129-1" checkinTime="1566623483646" checkinUser="pensoft" docAuthor="Paton, Alan J., Mwanyambo, Montfort, Govaerts, Rafael H. A., Smitha, Kokkaraniyil, Suddee, Somran, Phillipson, Peter B., Wilson, Trevor C., Forster, Paul I. &amp; Culham, Alastair" docDate="2019" docId="F75E54B95476502EBD582F0E162ADA91" docLanguage="en" docName="PhytoKeys 129: 1-158" docOrigin="PhytoKeys 129" docSource="http://dx.doi.org/10.3897/phytokeys.129.34988" docTitle="Equilabium goetzei Paton &amp; Mwanyambo &amp; Govaerts &amp; Smitha &amp; Suddee &amp; Phillipson &amp; Wilson &amp; Forster &amp; Culham 2019, comb. nov." docType="treatment" docVersion="6" id="BF57C6B3C3065AEE9B4B3D47189C908F" lastPageNumber="99" masterDocId="BF57C6B3C3065AEE9B4B3D47189C908F" masterDocTitle="Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera" masterLastPageNumber="158" masterPageNumber="1" pageNumber="99" updateTime="1668139430914" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paton, Alan J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mwanyambo, Montfort</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Govaerts, Rafael H. A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smitha, Kokkaraniyil</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Suddee, Somran</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Phillipson, Peter B.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wilson, Trevor C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Forster, Paul I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Culham, Alastair</mods:namePart>
</mods:name>
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<mods:date>2019</mods:date>
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<mods:number>129</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.129.34988</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-129-1</mods:identifier>
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<treatment ID-GBIF-Taxon="159244290" LSID="urn:lsid:plazi:treatment:F75E54B95476502EBD582F0E162ADA91" httpUri="http://treatment.plazi.org/id/F75E54B95476502EBD582F0E162ADA91" lastPageNumber="99" pageId="98" pageNumber="99">
<subSubSection pageId="98" pageNumber="99" type="nomenclature">
<paragraph pageId="98" pageNumber="99">
<taxonomicName LSID="F75E54B9-5476-502E-BD58-2F0E162ADA91" authority="(Guerke) Mwany. &amp; A. J. Paton" authorityName="Paton &amp; Mwanyambo &amp; Govaerts &amp; Smitha &amp; Suddee &amp; Phillipson &amp; Wilson &amp; Forster &amp; Culham" authorityYear="2019" baseAuthorityName="Guerke" class="Magnoliopsida" family="Lamiaceae" genus="Equilabium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Equilabium goetzei" order="Lamiales" pageId="98" pageNumber="99" phylum="Tracheophyta" rank="species" species="goetzei" status="comb. nov.">
Equilabium goetzei (
<normalizedToken originalValue="Gürke">Guerke</normalizedToken>
) Mwany. &amp; A.J.Paton
</taxonomicName>
<taxonomicNameLabel pageId="98" pageNumber="99">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="98" pageNumber="99" type="reference_group">
<paragraph pageId="98" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Plectranthus goetzei" order="Lamiales" pageId="98" pageNumber="99" phylum="Tracheophyta" rank="species" species="goetzei">Plectranthus goetzei</taxonomicName>
<normalizedToken originalValue="Gürke">Guerke</normalizedToken>
, Bot. Jahrb. Syst. 28: 469. 1900. Type: Tanzania, Iringa Dist., Rungemba, Mar. 1899, Goetze 724 (holotype: B, destroyed; isotype: BM).
</paragraph>
<paragraph pageId="98" pageNumber="99">
<taxonomicName authorityName="Brummitt &amp; Seyani" authorityYear="1987" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Plectranthus salubenii" order="Lamiales" pageId="98" pageNumber="99" phylum="Tracheophyta" rank="species" species="salubenii">Plectranthus salubenii</taxonomicName>
Brummitt &amp; Seyani, Kew Bull. 42: 690. 1987. Type: Malawi, Katumbi Escarpment, 41.5 km N of Rumphi, 26 Dec. 1970, Pawek 4155 (holotype: K).
</paragraph>
</subSubSection>
<subSubSection pageId="98" pageNumber="99" type="distribution">
<paragraph pageId="98" pageNumber="99">Distribution.</paragraph>
<paragraph pageId="98" pageNumber="99">SW. Tanzania to Zambia.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="9117D8D4F38AF77DD786394BE7BD096E">A new monotypic genus of ant-mimicking Coraebini (Coleoptera: Buprestidae: Agrilinae) from Madagascar</mods:title>
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<heading id="240081D6FF8FFFCAFF61FB2FFDF4FB56" bold="true" box="[151,586,1212,1238]" fontSize="11" level="1" pageId="1" pageNumber="66" reason="1">
<emphasis id="4D83EAA8FF8FFFCAFF61FB2FFDF4FB56" bold="true" box="[151,586,1212,1238]" pageId="1" pageNumber="66">
<taxonomicName id="B8F74D39FF8FFFCAFF61FB2FFE60FB56" ID-CoL="6QRLD" authority="Bellamy, 2008" authorityName="Bellamy" authorityYear="2008" box="[151,478,1212,1238]" class="Insecta" family="Buprestidae" genus="Madecorformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="66" phylum="Arthropoda" rank="species" species="silhouetta" status="sp. nov.">
<emphasis id="4D83EAA8FF8FFFCAFF61FB2FFE60FB56" bold="true" box="[151,478,1212,1238]" italics="true" pageId="1" pageNumber="66">Madecorformica silhouetta</emphasis>
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,
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</emphasis>
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</paragraph>
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<paragraph id="7F4836BAFF8FFFCAFF61FB4CFE99FB76" blockId="1.[151,586,1212,1270]" box="[151,295,1247,1270]" pageId="1" pageNumber="66">
(
<figureCitation id="E7CC2A3FFF8FFFCAFF68FB4CFEA0FB76" box="[158,286,1247,1270]" captionStart="FIGURES 1–6" captionStartId="2.[151,257,1611,1632]" captionTargetBox="[168,1421,263,1581]" captionTargetId="figure-266@2.[151,1436,263,1619]" captionTargetPageId="2" captionText="FIGURES 1–6. Madecorformica silhouetta, gen. &amp; sp. nov. Fig. 1, holotype, dorsal habitus; Fig. 2, holotype, lateral habitus; Fig. 3, closeup of humerus; Fig. 4, holotype, ventral habitus; Figs. 5a, b, elytral silhouette, inverted dorsal aspect; Fig. 6, aedeagus. Scale bars = 1.0 mm; scale equal for Figs. 1, 2, 4." figureDoi="http://doi.org/10.5281/zenodo.5125203" httpUri="https://zenodo.org/record/5125203/files/figure.png" pageId="1" pageNumber="66">Figures 1–6</figureCitation>
)
</paragraph>
<paragraph id="7F4836BAFF8FFFC9FF61FAB7FD43FF51" blockId="1.[151,1436,1316,2033]" lastBlockId="2.[151,1436,152,209]" lastPageId="2" lastPageNumber="67" pageId="1" pageNumber="66">
<emphasis id="4D83EAA8FF8FFFCAFF61FAB7FE5BFABB" box="[151,485,1316,1340]" inLineHeading="true" pageId="1" pageNumber="66" reason="1">
<emphasis id="4D83EAA8FF8FFFCAFF61FAB7FE61FABC" bold="true" box="[151,479,1316,1340]" pageId="1" pageNumber="66">
Description of male
<typeStatus id="A04C8818FF8FFFCAFE8BFAB7FE61FABC" box="[381,479,1316,1340]" pageId="1" pageNumber="66" type="holotype">holotype</typeStatus>
</emphasis>
.
</emphasis>
Myrmecomorphic; small (7.0 mm maximum length x
<quantity id="B80F9B5FFF8FFFCAFBD9FAB7FB3FFABB" box="[1071,1153,1316,1339]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="66" unit="mm" value="2.3">2.3 mm</quantity>
width across apical 1/3 of elytra), elongate-ovoid, subcylindrical; integument generally black with elytra bearing a mix of short erect setae, both black and off-white, forming a pattern resembling a silhouette of one pair of inverse facing heads (
<figureCitation id="E7CC2A3FFF8FFFCAFB48FAF9FA96FA01" box="[1214,1320,1386,1409]" captionStart="FIGURES 1–6" captionStartId="2.[151,257,1611,1632]" captionTargetBox="[168,1421,263,1581]" captionTargetId="figure-266@2.[151,1436,263,1619]" captionTargetPageId="2" captionText="FIGURES 1–6. Madecorformica silhouetta, gen. &amp; sp. nov. Fig. 1, holotype, dorsal habitus; Fig. 2, holotype, lateral habitus; Fig. 3, closeup of humerus; Fig. 4, holotype, ventral habitus; Figs. 5a, b, elytral silhouette, inverted dorsal aspect; Fig. 6, aedeagus. Scale bars = 1.0 mm; scale equal for Figs. 1, 2, 4." figureDoi="http://doi.org/10.5281/zenodo.5125203" httpUri="https://zenodo.org/record/5125203/files/figure.png" pageId="1" pageNumber="66">Fig. 5a, b</figureCitation>
); posterior portion of metaventrite and metepiventrite a deep iridescent maroon; underside with short, appressed squamose setae most dense on thoracic ventrites; head with frontovertex broad, flat on vertex; frons with short, shallow longitudinal depression between eyes; eyes large, inner margins diverging dorsally; circumocular groove extends from beyond dorsalmost point along inner margin to beyond ventral-most point; bisinuate supra-antennal groove above large antennal cavities; antennal cavities narrowly separated; clypeal margin broadly, deeply, arcuately emarginate; gena with feeble transverse depression for basal antennomeres in repose; distal margin of gena broadly rounded; antennae with scape narrow, elongate, feebly arcuate, nearly as long as 2+3+4 together, triangularly serrate from antennomere 5; right antennae missing antennomeres 10–11; pronotum 1.7 times wider than long, widest near midpoint, narrowing to posterior margin; anterior margin feebly convex; posterior margin bisinuate on either side of prescutellar lobe; disc transversely convex on anterior 1/2, otherwise flattened posteriorly and laterally; lateral margin entire; one elongate, narrow premarginal carina on either side parallel to margin; scutellum elongate cordiform; elytra much wider both across humeri and again at about apical 1/3; humeri strongly, subangularly elevated, with angled peaks slightly diverging from each other; disc evenly, transversely convex; lateral margin narrowing posterior to margin below humeral swelling, before widening at about mid-point then narrowing again from apical 1/3 to narrowly, separately rounded apices; epipleuron extends along entire length, widest below humerus, subparallel to opposite metacoxa, then much narrower and extending to apex; pygidium not visible beyond elytra, with apical margin entire; proventrite with mentonniere very feebly produced, angularly bilobed; disc evenly transversely convex; process broad, subparallel between procoxae, apex feebly trilobed; abdominal ventrites expanded laterally, ventrite 2 wider than 1 or 3–5, widest medially; ventrite 5 entire, without terminal sexual morphology; legs with tibiae flattened, having arcuate outer edge; tarsi with tarsomeres 1–4 with at least partial ventral pulvilli; tarsal claws appendiculate; aedeagus as in
<figureCitation id="E7CC2A3FFF8CFFC9FD4FFF29FD44FF51" box="[697,762,186,209]" captionStart="FIGURES 1–6" captionStartId="2.[151,257,1611,1632]" captionTargetBox="[168,1421,263,1581]" captionTargetId="figure-266@2.[151,1436,263,1619]" captionTargetPageId="2" captionText="FIGURES 1–6. Madecorformica silhouetta, gen. &amp; sp. nov. Fig. 1, holotype, dorsal habitus; Fig. 2, holotype, lateral habitus; Fig. 3, closeup of humerus; Fig. 4, holotype, ventral habitus; Figs. 5a, b, elytral silhouette, inverted dorsal aspect; Fig. 6, aedeagus. Scale bars = 1.0 mm; scale equal for Figs. 1, 2, 4." figureDoi="http://doi.org/10.5281/zenodo.5125203" httpUri="https://zenodo.org/record/5125203/files/figure.png" pageId="2" pageNumber="67">Fig. 6</figureCitation>
.
</paragraph>
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<paragraph id="7F4836BAFF8CFFC9FF61F9D8FDB4F91C" blockId="2.[151,1436,1611,1692]" pageId="2" pageNumber="67">
<emphasis id="4D83EAA8FF8CFFC9FF61F9D8FE8CF9E0" bold="true" box="[151,306,1611,1632]" pageId="2" pageNumber="67">FIGURES 1–6.</emphasis>
<taxonomicName id="B8F74D39FF8CFFC9FEC9F9DFFDFAF9E1" authorityName="Bellamy" authorityYear="2008" box="[319,580,1612,1633]" class="Insecta" family="Buprestidae" genus="Madecorformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="silhouetta">
<emphasis id="4D83EAA8FF8CFFC9FEC9F9DFFDFAF9E1" box="[319,580,1612,1633]" italics="true" pageId="2" pageNumber="67">Madecorformica silhouetta</emphasis>
</taxonomicName>
, gen. &amp; sp. nov. Fig. 1, holotype, dorsal habitus; Fig. 2, holotype, lateral habitus; Fig. 3, closeup of humerus; Fig. 4, holotype, ventral habitus; Figs. 5a, b, elytral silhouette, inverted dorsal aspect; Fig. 6, aedeagus. Scale bars = 1.0 mm; scale equal for Figs. 1, 2, 4.
</paragraph>
</caption>
<paragraph id="7F4836BAFF8CFFC9FF30F955FBD9F8A3" blockId="2.[151,1436,1734,2035]" pageId="2" pageNumber="67">
<emphasis id="4D83EAA8FF8CFFC9FF30F955FE86F95D" box="[198,312,1734,1758]" inLineHeading="true" pageId="2" pageNumber="67" reason="1">
<emphasis id="4D83EAA8FF8CFFC9FF30F955FE8FF95E" bold="true" box="[198,305,1734,1758]" pageId="2" pageNumber="67">Variation</emphasis>
.
</emphasis>
In the short
<typeStatus id="A04C8818FF8CFFC9FE35F955FE4FF95D" box="[451,497,1734,1757]" pageId="2" pageNumber="67">type</typeStatus>
series, the variation is only in size and proportion. Males vary: 6.2–7.3 x
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; females vary: 6.8–8.2 x 2.4–3.0 mm. The pair of silhouettes on the elytra vary slightly, allowing the imagination to consider that these may look like the head of a baboon or a chameleon, etc. (e.g.
<figureCitation id="E7CC2A3FFF8CFFC9FC38F89FFBE4F8A3" box="[974,1114,1804,1827]" captionStart="FIGURES 1–6" captionStartId="2.[151,257,1611,1632]" captionTargetBox="[168,1421,263,1581]" captionTargetId="figure-266@2.[151,1436,263,1619]" captionTargetPageId="2" captionText="FIGURES 1–6. Madecorformica silhouetta, gen. &amp; sp. nov. Fig. 1, holotype, dorsal habitus; Fig. 2, holotype, lateral habitus; Fig. 3, closeup of humerus; Fig. 4, holotype, ventral habitus; Figs. 5a, b, elytral silhouette, inverted dorsal aspect; Fig. 6, aedeagus. Scale bars = 1.0 mm; scale equal for Figs. 1, 2, 4." figureDoi="http://doi.org/10.5281/zenodo.5125203" httpUri="https://zenodo.org/record/5125203/files/figure.png" pageId="2" pageNumber="67">Figs. 1, 5a, b</figureCitation>
).
</paragraph>
<paragraph id="7F4836BAFF8CFFC9FF30F8BDFAC9F80B" blockId="2.[151,1436,1734,2035]" pageId="2" pageNumber="67">
<emphasis id="4D83EAA8FF8CFFC9FF30F8BDFE04F8C5" box="[198,442,1838,1862]" inLineHeading="true" pageId="2" pageNumber="67" reason="1">
<emphasis id="4D83EAA8FF8CFFC9FF30F8BDFE0CF8C6" bold="true" box="[198,434,1838,1862]" pageId="2" pageNumber="67">Specimens examined</emphasis>
.
</emphasis>
<materialsCitation id="CF9F3CE7FF8CFFC9FE36F8BDFCF5F80B" collectionCode="MNHN" collectorName="Distr. Mananara N." country="Madagascar" pageId="2" pageNumber="67" specimenCount="27" specimenCount-female="6" specimenCount-male="8" typeStatus="holotype">
<typeStatus id="A04C8818FF8CFFC9FE36F8BDFD9DF8C5" box="[448,547,1838,1861]" pageId="2" pageNumber="67" type="holotype">Holotype</typeStatus>
<specimenCount id="69F1FD33FF8CFFC9FDDCF8A3FD84F8C0" box="[554,570,1840,1856]" pageId="2" pageNumber="67" type="male">♂</specimenCount>
(
<collectionCode id="19E6AE7FFF8CFFC9FDBEF8BDFD1EF8C5" box="[584,672,1838,1861]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="67">MNHN</collectionCode>
):
<collectingCountry id="07E0762AFF8CFFC9FD45F8BDFC8AF8C5" box="[691,820,1838,1861]" name="Madagascar" pageId="2" pageNumber="67">Madagascar</collectingCountry>
Est.,
<collectorName id="D202536CFF8CFFC9FC86F8BDFB81F8C5" box="[880,1087,1838,1861]" pageId="2" pageNumber="67">Distr. Mananara N.</collectorName>
, Seranambe, II-[19]64, Vadon et Peyrieras;
<specimenCount id="69F1FD33FF8CFFC9FEFEF8C2FE2DF8E8" box="[264,403,1873,1896]" pageId="2" pageNumber="67" type="generic" typeStatus="paratypes">13 paratypes</specimenCount>
: 8, same data as holotype, except
<specimenCount id="69F1FD33FF8CFFC9FD0EF8C2FCA5F8E2" box="[760,795,1873,1896]" pageId="2" pageNumber="67" type="female">1 ♀</specimenCount>
, I-[19]64;
<specimenCount id="69F1FD33FF8CFFC9FC79F8C2FC7CF8E2" box="[911,962,1873,1896]" pageId="2" pageNumber="67" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="69F1FD33FF8CFFC9FC26F8C2FC4DF8E2" box="[976,1011,1873,1896]" pageId="2" pageNumber="67" type="female">1 ♀</specimenCount>
, II-[19]65;
<specimenCount id="69F1FD33FF8CFFC9FB99F8C2FB2CF8E2" box="[1135,1170,1873,1896]" pageId="2" pageNumber="67" type="male">1 ♂</specimenCount>
,
<specimenCount id="69F1FD33FF8CFFC9FB69F8C2FB6DF8E2" box="[1183,1235,1873,1896]" pageId="2" pageNumber="67" type="female">2 ♀♀</specimenCount>
, without date;
<specimenCount id="69F1FD33FF8CFFC9FA84F8C2FA2BF8E2" box="[1394,1429,1873,1896]" pageId="2" pageNumber="67" type="male">1 ♂</specimenCount>
, Antanambe, II-[19]67;
<specimenCount id="69F1FD33FF8CFFC9FE79F8E7FE7DF805" box="[399,451,1908,1931]" pageId="2" pageNumber="67" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="69F1FD33FF8CFFC9FE26F8E7FDBAF805" box="[464,516,1908,1931]" pageId="2" pageNumber="67" type="female">2 ♀♀</specimenCount>
, Mont Antampona, iii-[19]67
</materialsCitation>
.
<materialsCitation id="CF9F3CE7FF8CFFC9FCA3F8E7FACDF80B" box="[853,1395,1908,1931]" collectionCode="MNHN, NMPC, CLBC" country="Madagascar" pageId="2" pageNumber="67" specimenCount="1" typeStatus="paratype">
<typeStatus id="A04C8818FF8CFFC9FCA3F8E7FC03F80B" box="[853,957,1908,1931]" pageId="2" pageNumber="67" type="paratype">Paratypes</typeStatus>
deposited in
<collectionCode id="19E6AE7FFF8CFFC9FBBAF8E7FB1EF80B" box="[1100,1184,1908,1931]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="67">MNHN</collectionCode>
,
<collectionCode id="19E6AE7FFF8CFFC9FB5DF8E7FB49F80B" box="[1195,1271,1908,1931]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="67">NMPC</collectionCode>
and
<collectionCode id="19E6AE7FFF8CFFC9FADCF8E7FACDF80B" box="[1322,1395,1908,1931]" pageId="2" pageNumber="67">CLBC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="37ED6531FF8CFFC9FF30F805FAB5F82D" box="[198,1291,1942,1966]" pageId="2" pageNumber="67" type="etymology">
<paragraph id="7F4836BAFF8CFFC9FF30F805FAB5F82D" blockId="2.[151,1436,1734,2035]" box="[198,1291,1942,1966]" pageId="2" pageNumber="67">
<emphasis id="4D83EAA8FF8CFFC9FF30F805FEF8F82D" box="[198,326,1942,1966]" inLineHeading="true" pageId="2" pageNumber="67" reason="1">
<emphasis id="4D83EAA8FF8CFFC9FF30F805FE81F82E" bold="true" box="[198,319,1942,1966]" pageId="2" pageNumber="67">Etymology</emphasis>
.
</emphasis>
The specific epithet
<taxonomicName id="B8F74D39FF8CFFC9FDD3F804FD30F82E" authorityName="Bellamy" authorityYear="2008" box="[549,654,1943,1966]" class="Insecta" family="Buprestidae" genus="Madecorformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="silhouetta">
<emphasis id="4D83EAA8FF8CFFC9FDD3F804FD30F82E" box="[549,654,1943,1966]" italics="true" pageId="2" pageNumber="67">silhouetta</emphasis>
</taxonomicName>
is chosen for the elytral pattern of facing heads on the disc.
</paragraph>
</subSubSection>
<subSubSection id="37ED6531FF8CFFC9FF30F82AFDBDF873" pageId="2" pageNumber="67" type="discussion">
<paragraph id="7F4836BAFF8CFFC9FF30F82AFDBDF873" blockId="2.[151,1436,1734,2035]" pageId="2" pageNumber="67">
<emphasis id="4D83EAA8FF8CFFC9FF30F82AFE8BF850" box="[198,309,1977,2001]" inLineHeading="true" pageId="2" pageNumber="67" reason="1">
<emphasis id="4D83EAA8FF8CFFC9FF30F82AFE90F851" bold="true" box="[198,302,1977,2001]" pageId="2" pageNumber="67">Remarks</emphasis>
.
</emphasis>
This new species differs, and can be distinguished, from other lookalike, putative ant-mimics of the genus
<emphasis id="4D83EAA8FF8CFFC9FF61F84FFE93F873" box="[151,301,2012,2035]" italics="true" pageId="2" pageNumber="67">Entomogaster</emphasis>
as noted in
<tableCitation id="32750301FF8CFFC9FE5BF84FFE42F873" box="[429,508,2012,2035]" captionStart="TABLE 1" captionStartId="1.[151,241,567,591]" captionText="TABLE 1. Diagnostic character states for Entomogaster and Madecorformica." pageId="2" pageNumber="67">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

253
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<paragraph id="7F4836BBAE4DFFC6FF5CF8F8F346E259" blockId="1.[151,976,1831,1925]" box="[151,512,1831,1858]" pageId="1" pageNumber="550">
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<emphasis id="4D83EAA9AE4DFFC6FF5CF8F8F346E259" bold="true" box="[151,512,1831,1858]" pageId="1" pageNumber="550">
Genus
<taxonomicName id="B8F74D38AE4DFFC6FF25F8F7F0CCE25A" ID-CoL="9J22T" authority="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo, 2022" authorityName="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo" authorityYear="2022" box="[238,394,1831,1857]" class="Insecta" family="Scatopsidae" genus="Afroscatopse" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="550" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="4D83EAA9AE4DFFC6FF25F8F7F0CCE25A" bold="true" box="[238,394,1831,1857]" italics="true" pageId="1" pageNumber="550">Afroscatopse</emphasis>
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<taxonomicNameLabel id="56B057D2AE4DFFC6FE59F8F8F346E259" box="[402,512,1832,1858]" pageId="1" pageNumber="550" rank="genus">gen. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="7F4836BBAE4DFFC6FF5CF8BDF2FDE29E" blockId="1.[151,976,1831,1925]" box="[151,955,1900,1925]" pageId="1" pageNumber="550">
<typeStatus id="A04C8819AE4DFFC6FF5CF8BDF18FE29E" box="[151,201,1901,1925]" pageId="1" pageNumber="550">Type</typeStatus>
species
<taxonomicName id="B8F74D38AE4DFFC6FED4F8BDF0B3E29F" authority="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo, 2022" authorityName="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo" authorityYear="2022" box="[287,501,1900,1925]" class="Insecta" family="Scatopsidae" genus="Afroscatopse" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="550" phylum="Arthropoda" rank="species" species="haennii" status="sp. nov.">
<emphasis id="4D83EAA9AE4DFFC6FED4F8BDF0B3E29F" box="[287,501,1900,1925]" italics="true" pageId="1" pageNumber="550">Afroscatopse haennii</emphasis>
</taxonomicName>
<emphasis id="4D83EAA9AE4DFFC6FE30F8BDF30AE29E" bold="true" box="[507,588,1901,1925]" pageId="1" pageNumber="550">
<taxonomicNameLabel id="56B057D2AE4DFFC6FE30F8BDF30AE29E" box="[507,588,1901,1925]" pageId="1" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, present designation and monotypy.
</paragraph>
</subSubSection>
<subSubSection id="37ED6530AE4DFFC5FF5CF862F281E5CE" lastPageId="2" lastPageNumber="551" pageId="1" pageNumber="550" type="diagnosis">
<paragraph id="7F4836BBAE4DFFC5FF5CF862F281E5CE" blockId="1.[151,1436,1970,2039]" lastBlockId="2.[151,1437,151,1011]" lastPageId="2" lastPageNumber="551" pageId="1" pageNumber="550">
<emphasis id="4D83EAA9AE4DFFC6FF5CF862F04CE2D7" bold="true" box="[151,266,1970,1996]" pageId="1" pageNumber="550">Diagnosis</emphasis>
. The genus is distinct in having very broad wings with thick veins, enlarged thick basal ⅓ of costal vein, base of M
<subScript id="E37334FEAE4DFFC6FEC0F839F055E2EC" attach="left" box="[267,275,2025,2039]" fontSize="6" pageId="1" pageNumber="550">1</subScript>
reduced, eyes dichoptic, long antennae with flagellum composed of 9 flagellomeres, wing membrane without macrotrichia, and tergite
<quantity id="B80F9B5EAE4EFFC5FDC6FF47F37EE5AA" box="[525,568,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="2" pageNumber="551" unit="in" value="7.0">7 in</quantity>
male genitalia divided into two long and broad lobes. The strongly thickened costa is unique apomorhic character within the family. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="37ED6530AE4EFFC5FF0CFF0FF189E406" pageId="2" pageNumber="551" type="etymology">
<paragraph id="7F4836BBAE4EFFC5FF0CFF0FF189E406" blockId="2.[151,1437,151,1011]" pageId="2" pageNumber="551">
<emphasis id="4D83EAA9AE4EFFC5FF0CFF0FF00EE5E2" bold="true" box="[199,328,223,249]" pageId="2" pageNumber="551">Etymology</emphasis>
. The generic name is combination of two words: Africa and generic name
<taxonomicName id="B8F74D38AE4EFFC5FB46FF0FF5B7E5E2" authorityName="Geoffroy" authorityYear="1762" box="[1165,1265,223,249]" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FB46FF0FF5B7E5E2" box="[1165,1265,223,249]" italics="true" pageId="2" pageNumber="551">Scatopse</emphasis>
</taxonomicName>
. Gender: feminine.
</paragraph>
</subSubSection>
<subSubSection id="37ED6530AE4EFFC5FF0CFEF6F5B2E6F6" pageId="2" pageNumber="551" type="discussion">
<paragraph id="7F4836BBAE4EFFC5FF0CFEF6F0BFE4CA" blockId="2.[151,1437,151,1011]" pageId="2" pageNumber="551">
<emphasis id="4D83EAA9AE4EFFC5FF0CFEF6F003E45B" bold="true" box="[199,325,294,320]" pageId="2" pageNumber="551">Discussion</emphasis>
. According to Dalton de Souza Amorim (personal comm.) the new species described below does not need new genus and should be placed in the worldwide distributed genus
<taxonomicName id="B8F74D38AE4EFFC5FC32FE9BF400E47E" authority="Enderlein, 1926" authorityName="Enderlein" authorityYear="1926" box="[1017,1350,331,357]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FC32FE9BF5CBE47E" box="[1017,1165,331,357]" italics="true" pageId="2" pageNumber="551">Colobostema</emphasis>
<bibRefCitation id="1B664B4AAE4EFFC5FB58FE9BF400E47E" author="Enderlein, G." box="[1171,1350,331,357]" pageId="2" pageNumber="551" pagination="137 - 142" refId="ref6100" refString="Enderlein, G. (1926) Zur Kenntnis der Scatopsiden. Zoologischer Anzeiger, 68, 137 - 142." type="journal article" year="1926">Enderlein, 1926</bibRefCitation>
</taxonomicName>
including 65 extant species (
<bibRefCitation id="1B664B4AAE4EFFC5FE5FFEBFF3E9E492" author="Haenni, J. - P. &amp; Amorim, D. de" box="[404,687,367,393]" pageId="2" pageNumber="551" pagination="641 - 651" refId="ref6936" refString="Haenni, J. - P. &amp; Amorim, D. de S. (2017) Scatopsidae (minute black scavenger flies or dung midges). In: Kirk-Spriggs, A. H. &amp; Sinclair, B. (Eds.), Manual of Afrotropical Diptera, Volume 2. Nematocerous Diptera and lower Brachycera. Suricata. Vol. 5. South African National Biodiversity Institute, Pretoria. pp. 641 - 651." type="book chapter" year="2017">Haenni &amp; Amorim 2017</bibRefCitation>
). In our opinion supported by Jean-Paul Haenni (personal comm.) this new genus with unique costal vein is the most distinct within the tribe
<taxonomicName id="B8F74D38AE4EFFC5FC2AFE43F5DCE4B6" authorityName="Amorim" authorityYear="1994" box="[993,1178,403,429]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="tribe" tribe="Colobostematini">Colobostematini</taxonomicName>
including 6 extant and 1 fossil genera (
<bibRefCitation id="1B664B4AAE4EFFC5FE82FE67F0ABE4CA" author="Amorim, D. de" box="[329,493,439,465]" pageId="2" pageNumber="551" pagination="1 - 17" refId="ref5173" refString="Amorim, D. de S. (1998) Amber fossil Scatopsidae (Diptera: Psychodomorpha). I. Considerations on described taxa, Procolob- ostema roseni, new species, from Dominican amber, and the position of Procolobostema in the family. American Museum Novitates, 3227, 1 - 17. [http: // hdl. handle. net / 2246 / 3499]" type="journal article" year="1998">Amorim 1998</bibRefCitation>
).
</paragraph>
<paragraph id="7F4836BBAE4EFFC5FF0CFE0BF23EE60D" blockId="2.[151,1437,151,1011]" pageId="2" pageNumber="551">
<taxonomicName id="B8F74D38AE4EFFC5FF0CFE0BF01EE4EE" authority="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo, 2022" authorityName="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo" authorityYear="2022" box="[199,344,475,501]" class="Insecta" family="Scatopsidae" genus="Afroscatopse" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="4D83EAA9AE4EFFC5FF0CFE0BF01EE4EE" box="[199,344,475,501]" italics="true" pageId="2" pageNumber="551">Afroscatopse</emphasis>
</taxonomicName>
<emphasis id="4D83EAA9AE4EFFC5FE95FE0BF080E4EE" bold="true" box="[350,454,475,501]" pageId="2" pageNumber="551">
<taxonomicNameLabel id="56B057D2AE4EFFC5FE95FE0BF080E4EE" box="[350,454,475,501]" pageId="2" pageNumber="551" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
has long antennae and broad thorax not compressed laterally, rounded head with dichoptic eyes and wing membrane without macrotrichia; this places the genus within the tribe
<taxonomicName id="B8F74D38AE4EFFC5FBBFFE2FF46BE702" authorityName="Amorim" authorityYear="1994" box="[1140,1325,511,537]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="tribe" tribe="Colobostematini">Colobostematini</taxonomicName>
(
<bibRefCitation id="1B664B4AAE4EFFC5FAF0FE2FF194E726" author="Amorim, D. de" pageId="2" pageNumber="551" pagination="1 - 17" refId="ref5173" refString="Amorim, D. de S. (1998) Amber fossil Scatopsidae (Diptera: Psychodomorpha). I. Considerations on described taxa, Procolob- ostema roseni, new species, from Dominican amber, and the position of Procolobostema in the family. American Museum Novitates, 3227, 1 - 17. [http: // hdl. handle. net / 2246 / 3499]" type="journal article" year="1998">Amorim 1998</bibRefCitation>
,
<bibRefCitation id="1B664B4AAE4EFFC5FF14FDF3F030E726" author="Haenni, J. - P." box="[223,374,547,573]" pageId="2" pageNumber="551" pagination="199 - 242" refId="ref6879" refString="Haenni, J. - P. (2013) A revision of the West Palaearctic species of Colobostema Enderlein, 1926 (Diptera, Scatopsidae). Part I. European subregion. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 86, 199 - 242. https: // doi. org / 10.5169 / seals- 40307" type="journal article" year="2013">Haenni 2013</bibRefCitation>
). The tribe includes the following genera (
<bibRefCitation id="1B664B4AAE4EFFC5FC94FDF3F540E726" author="Amorim, D. de" box="[863,1030,547,573]" pageId="2" pageNumber="551" pagination="1 - 17" refId="ref5173" refString="Amorim, D. de S. (1998) Amber fossil Scatopsidae (Diptera: Psychodomorpha). I. Considerations on described taxa, Procolob- ostema roseni, new species, from Dominican amber, and the position of Procolobostema in the family. American Museum Novitates, 3227, 1 - 17. [http: // hdl. handle. net / 2246 / 3499]" type="journal article" year="1998">Amorim 1998</bibRefCitation>
):
<taxonomicName id="B8F74D38AE4EFFC5FBD7FDF3F428E726" authority="Enderlein, 1926" authorityName="Enderlein" authorityYear="1926" box="[1052,1390,547,573]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FBD7FDF3F5F6E726" box="[1052,1200,547,573]" italics="true" pageId="2" pageNumber="551">Colobostema</emphasis>
<bibRefCitation id="1B664B4AAE4EFFC5FB72FDF3F428E726" author="Enderlein, G." box="[1209,1390,547,573]" pageId="2" pageNumber="551" pagination="137 - 142" refId="ref6100" refString="Enderlein, G. (1926) Zur Kenntnis der Scatopsiden. Zoologischer Anzeiger, 68, 137 - 142." type="journal article" year="1926">Enderlein, 1926</bibRefCitation>
</taxonomicName>
(65 species, worldwide),
<taxonomicName id="B8F74D38AE4EFFC5FE4EFD97F3F8E77A" authority="Enderlein, 1912" authorityName="Enderlein" authorityYear="1912" box="[389,702,583,609]" class="Insecta" family="Scatopsidae" genus="Holoplagia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FE4EFD97F343E77A" box="[389,517,583,609]" italics="true" pageId="2" pageNumber="551">Holoplagia</emphasis>
<bibRefCitation id="1B664B4AAE4EFFC5FDC0FD97F3F8E77A" author="Enderlein, G." box="[523,702,583,609]" pageId="2" pageNumber="551" pagination="261 - 282" refId="ref6071" refString="Enderlein, G. (1912) Zur Kenntnis der Zygophthalmen. Uber die Gruppierung der Sciariden und Scatopsiden. Zoologischer Anzeiger, 40, 261 - 282." type="journal article" year="1912">Enderlein, 1912</bibRefCitation>
</taxonomicName>
(10 species, Afrotropical, Australasian, Neotropical, Oriental and Palaearctic regions),
<taxonomicName id="B8F74D38AE4EFFC5FE4EFDBBF379E79E" authority="Cook" authorityName="Cook" box="[389,575,619,645]" class="Insecta" family="Scatopsidae" genus="Ferneiella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FE4EFDBBF0BCE79E" box="[389,506,619,645]" italics="true" pageId="2" pageNumber="551">Ferneiella</emphasis>
Cook
</taxonomicName>
in Freeman, 1985 (3 extant species, Australasian, Palaearctic regions; 1 extinct from Eocene French amber),
<taxonomicName id="B8F74D38AE4EFFC5FDEAFD5FF27CE7B2" authority="Haenni, 1998" authorityName="Haenni" authorityYear="1998" box="[545,826,655,681]" class="Insecta" family="Scatopsidae" genus="Efcookella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FDEAFD5FF3DFE7B2" box="[545,665,655,681]" italics="true" pageId="2" pageNumber="551">Efcookella</emphasis>
<bibRefCitation id="1B664B4AAE4EFFC5FD69FD5FF27CE7B2" author="Haenni, J. - P." box="[674,826,655,681]" pageId="2" pageNumber="551" refId="ref6820" refString="Haenni, J. - P. (1998) Scatopsidae. In: Merz, B., Bachli, G., Haenni, J. - P. &amp; Gonseth, Y. (Eds.), Fauna Helvetica. Vol. 1. Diptera- Checklist, Neuchatel, pp. 141 - 144." type="book" year="1998">Haenni, 1998</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="B8F74D38AE4EFFC5FC97FD5FF535E7B2" authority="Freeman, 1985" authorityName="Freeman" authorityYear="1985" box="[860,1139,654,681]" class="Dothideomycetes" family="Cookellaceae" genus="Cookella" kingdom="Fungi" order="Chordeumatida" pageId="2" pageNumber="551" phylum="Ascomycota" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FC97FD5FF287E7B2" box="[860,961,655,681]" italics="true" pageId="2" pageNumber="551">Cookella</emphasis>
Freeman, 1985
</taxonomicName>
(1 extant, Palaearctic Region, 2 fossil from Eocene of
<collectingCountry id="07E0762BAE4EFFC5FE22FD63F373E7D6" box="[489,565,691,717]" name="France" pageId="2" pageNumber="551">France</collectingCountry>
(Oise amber) and
<collectingCountry id="07E0762BAE4EFFC5FCC2FD63F279E7D6" box="[777,831,691,717]" name="United States of America" pageId="2" pageNumber="551">USA</collectingCountry>
(Florissant)),
<taxonomicName id="B8F74D38AE4EFFC5FC2BFD63F464E7D6" authority="Cook, 1971" authorityName="Cook" authorityYear="1971" box="[992,1314,691,717]" class="Insecta" family="Scatopsidae" genus="Procolobostema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FC2BFD63F5D1E7D6" box="[992,1175,691,717]" italics="true" pageId="2" pageNumber="551">Procolobostema</emphasis>
<bibRefCitation id="1B664B4AAE4EFFC5FB54FD63F464E7D6" author="Cook, E. F." box="[1183,1314,691,717]" pageId="2" pageNumber="551" pagination="57 - 61" refId="ref5752" refString="Cook, E. F. (1971) Fossil Scatopsidae in Mexican amber (Diptera). University of California Publication in Entomology, 63, 57 - 61." type="journal article" year="1971">Cook, 1971</bibRefCitation>
</taxonomicName>
(extinct, 4 species, Miocene, Mexican and Dominican ambers),
<taxonomicName id="B8F74D38AE4EFFC5FD25FD07F517E7EA" authority="Freeman, 1990" authorityName="Freeman" authorityYear="1990" box="[750,1105,727,753]" class="Insecta" family="Scatopsidae" genus="Borneoscatopse" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FD25FD07F2E7E7EB" box="[750,929,727,752]" italics="true" pageId="2" pageNumber="551">Borneoscatopse</emphasis>
<bibRefCitation id="1B664B4AAE4EFFC5FC62FD07F517E7EA" author="Freeman, P." box="[937,1105,727,753]" pageId="2" pageNumber="551" pagination="9 - 19" refId="ref6326" refString="Freeman, P. (1990) Redescription of seven Oriental species of Scatopsidae described by F. W. Edwards in the genus Scatopse. The Entomologist's Monthly Magazine, 126, 9 - 19." type="journal article" year="1990">Freeman, 1990</bibRefCitation>
</taxonomicName>
(1 species, Oriental Region), and
<taxonomicName id="B8F74D38AE4EFFC5FF0CFD2BF0A2E60E" authority="Edwards, 1928" authorityName="Edwards" authorityYear="1928" box="[199,484,763,789]" class="Insecta" family="Scatopsidae" genus="Lumpuria" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FF0CFD2BF073E60F" box="[199,309,763,788]" italics="true" pageId="2" pageNumber="551">Lumpuria</emphasis>
<bibRefCitation id="1B664B4AAE4EFFC5FEF7FD2BF0A2E60E" author="Edwards, F. W." box="[316,484,763,789]" pageId="2" pageNumber="551" pagination="1 - 139" refId="ref5997" refString="Edwards, F. W. (1928) Diptera Nematocera from the Federated Malay States Museum. Journal of the Federated Malay States Museums, 14, 1 - 139." type="journal article" year="1928">Edwards, 1928</bibRefCitation>
</taxonomicName>
(5 extant species, Oriental Region).
</paragraph>
<paragraph id="7F4836BBAE4EFFC5FF0CFCCFF5B2E6F6" blockId="2.[151,1437,151,1011]" pageId="2" pageNumber="551">
<taxonomicName id="B8F74D38AE4EFFC5FF0CFCCFF001E622" authorityName="Enderlein" authorityYear="1912" box="[199,327,799,825]" class="Insecta" family="Scatopsidae" genus="Holoplagia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FF0CFCCFF001E622" box="[199,327,799,825]" italics="true" pageId="2" pageNumber="551">Holoplagia</emphasis>
</taxonomicName>
and
<taxonomicName id="B8F74D38AE4EFFC5FEB6FCCFF0B3E622" authorityName="Haenni" authorityYear="1998" box="[381,501,799,825]" class="Insecta" family="Scatopsidae" genus="Efcookella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FEB6FCCFF0B3E622" box="[381,501,799,825]" italics="true" pageId="2" pageNumber="551">Efcookella</emphasis>
</taxonomicName>
differ in having crossvein connecting M
<subScript id="E37334FEAE4EFFC5FC71FCE1F284E624" attach="left" box="[954,962,817,831]" fontSize="6" pageId="2" pageNumber="551">1</subScript>
and R
<subScript id="E37334FEAE4EFFC5FBC1FCE1F562E624" attach="left" box="[1034,1060,817,831]" fontSize="6" pageId="2" pageNumber="551">4+5</subScript>
. The latter genus has slender thorax.
<taxonomicName id="B8F74D38AE4EFFC5FF02FC93F071E647" authorityName="Edwards" authorityYear="1928" box="[201,311,835,860]" class="Insecta" family="Scatopsidae" genus="Lumpuria" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FF02FC93F071E647" box="[201,311,835,860]" italics="true" pageId="2" pageNumber="551">Lumpuria</emphasis>
</taxonomicName>
has hollowed occiput.
<taxonomicName id="B8F74D38AE4EFFC5FDF1FC93F388E646" authorityName="Enderlein" authorityYear="1926" box="[570,718,835,861]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FDF1FC93F388E646" box="[570,718,835,861]" italics="true" pageId="2" pageNumber="551">Colobostema</emphasis>
</taxonomicName>
has CuA with double smooth curve, veins M
<subScript id="E37334FEAE4EFFC5FB02FC85F597E678" attach="left" box="[1225,1233,853,867]" fontSize="6" pageId="2" pageNumber="551">1</subScript>
and M
<subScript id="E37334FEAE4EFFC5FAEBFC85F46EE678" attach="left" box="[1312,1320,853,867]" fontSize="6" pageId="2" pageNumber="551">2</subScript>
vanishing before wing margin, flagellum with 8 flagellomeres, wide thorax, medium sized with body length
<quantity id="B80F9B5EAE4EFFC5FB17FCB6F426E69A" box="[1244,1376,870,897]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="2" pageNumber="551" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5-2.5 mm</quantity>
. Fossil
<taxonomicName id="B8F74D38AE4EFFC5FF73FC5BF029E6BE" authorityName="Cook" authorityYear="1971" box="[184,367,907,933]" class="Insecta" family="Scatopsidae" genus="Procolobostema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FF73FC5BF029E6BE" box="[184,367,907,933]" italics="true" pageId="2" pageNumber="551">Procolobostema</emphasis>
</taxonomicName>
has CuA and M
<subScript id="E37334FEAE4EFFC5FDEFFC4DF36AE6B0" attach="left" box="[548,556,925,939]" fontSize="6" pageId="2" pageNumber="551">4</subScript>
gradually C-curved, M
<subScript id="E37334FEAE4EFFC5FCF8FC4DF27DE6B0" attach="left" box="[819,827,925,939]" fontSize="6" pageId="2" pageNumber="551">1</subScript>
and M
<subScript id="E37334FEAE4EFFC5FC41FC4DF2D4E6B0" attach="left" box="[906,914,925,939]" fontSize="6" pageId="2" pageNumber="551">2</subScript>
reaching wing margin, base of M
<subScript id="E37334FEAE4EFFC5FAC0FC4DF455E6B0" attach="left" box="[1291,1299,925,939]" fontSize="6" pageId="2" pageNumber="551">1</subScript>
complete, 8 flagellomeres, wide thorax.
<taxonomicName id="B8F74D38AE4EFFC5FE04FC7FF302E6D2" box="[463,580,943,969]" class="Insecta" family="Scatopsidae" genus="Ferneiella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FE04FC7FF302E6D2" box="[463,580,943,969]" italics="true" pageId="2" pageNumber="551">Ferneiella</emphasis>
</taxonomicName>
has elongated scutum and 8 flagellomeres.
<taxonomicName id="B8F74D38AE4EFFC5FBE0FC7FF598E6D3" authorityName="Freeman" authorityYear="1990" box="[1067,1246,943,968]" class="Insecta" family="Scatopsidae" genus="Borneoscatopse" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="4D83EAA9AE4EFFC5FBE0FC7FF598E6D3" box="[1067,1246,943,968]" italics="true" pageId="2" pageNumber="551">Borneoscatopse</emphasis>
</taxonomicName>
has wide thorax, 8 flagellomeres, stem of haltere lacking setae, radial cell broad and long, M
<subScript id="E37334FEAE4EFFC5FC2FFC35F2AAE6E8" attach="left" box="[996,1004,997,1011]" fontSize="6" pageId="2" pageNumber="551">1</subScript>
complete without spur.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="2EB1D7303A2209BD945561922316B180" ID-CLB-Dataset="20626" ID-DOI="10.11646/zootaxa.5099.5.3" ID-GBIF-Dataset="39cc7488-ad5a-4372-9920-216ffac600db" ID-ISSN="1175-5326" ID-Zenodo-Dep="6110208" ID-ZooBank="4E2A384E-B908-4D59-AC1D-CF5F6474967D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644997773826" checkinUser="plazi" docAuthor="Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta &amp; Szwedo, Jacek" docDate="2022" docId="F75E87ADAE4FFFC2FF5CF906F5CAE79E" docLanguage="en" docName="zootaxa.5099.5.3.pdf" docOrigin="Zootaxa 5099 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Afroscatopse haennii Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo 2022, sp. nov." docType="treatment" docUuid="EF05224A-51D4-408B-B4B7-FE389A786D12" docUuidSource="ZooBank" docVersion="9" lastPageNumber="554" masterDocId="0B67FFD5AE4CFFC7FFCBFFD0F146E51B" masterDocTitle="A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae)" masterLastPageNumber="562" masterPageNumber="549" pageNumber="552" updateTime="1699312098752" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A3BD887C7D57D36D19A64F7777BB0018">A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae)</mods:title>
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<mods:namePart id="FE523DA872F4744A3BAE62337EE99429">Szadziewski, Ryszard</mods:namePart>
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<mods:namePart id="5FB42887966E3373712A998285FF7693">Bojarski, Błażej</mods:namePart>
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<mods:namePart id="50A838F926F2B5C94305084E5E66F785">Sontag, Elżbieta</mods:namePart>
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<mods:affiliation id="09BA5C920114E60A060D1623FC6367FE">elzbieta. sontag @ ug. edu. pl; https: // orcid. org / 0000 - 0002 - 5966 - 8334</mods:affiliation>
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<subSubSection id="37ED6530AE4FFFC4FF5CF906F0BBE3EA" box="[151,509,1750,1777]" pageId="3" pageNumber="552" type="nomenclature">
<paragraph id="7F4836BBAE4FFFC4FF5CF906F0BBE3EA" blockId="3.[151,965,1750,1850]" box="[151,509,1750,1777]" pageId="3" pageNumber="552">
<heading id="240081D7AE4FFFC4FF5CF906F0BBE3EA" bold="true" box="[151,509,1750,1777]" fontSize="11" level="1" pageId="3" pageNumber="552" reason="1">
<emphasis id="4D83EAA9AE4FFFC4FF5CF906F0BBE3EA" bold="true" box="[151,509,1750,1777]" pageId="3" pageNumber="552">
<taxonomicName id="B8F74D38AE4FFFC4FF5CF906F0DEE3EA" authority="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo, 2022" authorityName="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo" authorityYear="2022" box="[151,408,1750,1777]" class="Insecta" family="Scatopsidae" genus="Afroscatopse" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="552" phylum="Arthropoda" rank="species" species="haennii" status="sp. nov.">
<emphasis id="4D83EAA9AE4FFFC4FF5CF906F0DEE3EA" bold="true" box="[151,408,1750,1777]" italics="true" pageId="3" pageNumber="552">Afroscatopse haennii</emphasis>
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<taxonomicNameLabel id="56B057D2AE4FFFC4FE54F907F0BBE3EA" box="[415,509,1751,1777]" pageId="3" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="37ED6530AE4FFFC4FF5CF92BF283E221" pageId="3" pageNumber="552" type="description">
<paragraph id="7F4836BBAE4FFFC4FF5CF92BF04DE20E" blockId="3.[151,965,1750,1850]" box="[151,267,1787,1813]" pageId="3" pageNumber="552">
(
<figureCitation id="E7CC2A3EAE4FFFC4FF54F92BF044E20E" box="[159,258,1787,1813]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,2008,2032]" captionStartId-1="3.[151,250,1608,1632]" captionStartId-2="4.[151,250,1896,1920]" captionTargetBox-0="[172,1415,1026,1982]" captionTargetBox-1="[151,1436,181,1585]" captionTargetBox-2="[151,1436,849,1873]" captionTargetId-0="figure-403@2.[172,1415,1026,1982]" captionTargetId-1="figure-19@3.[151,1436,181,1585]" captionTargetId-2="figure-359@4.[151,1436,849,1873]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Afroscatopse haennii gen. et sp. nov., holotype male, dorsal view." captionText-1="FIGURE 2. Afroscatopse haennii gen. et sp. nov., holotype male, head and wing.A—head with antennae, B—vertex and frons, C—antenna, D—wing." captionText-2="FIGURE 3. Afroscatopse haennii gen. et sp. nov., male genitalia. A—ventral view, phot., B—ventral aspect, drawing, C—lat- erodorsal view, phot., D—laterodorsal aspect of tergite 7, and gonocoxite. Abbreviations: st7—abdominal sternite 7, tg7—ab- dominal tergite 7, gx—gonocoxite." figureDoi-0="http://doi.org/10.5281/zenodo.6110210" figureDoi-1="http://doi.org/10.5281/zenodo.6110212" figureDoi-2="http://doi.org/10.5281/zenodo.6110214" httpUri-0="https://zenodo.org/record/6110210/files/figure.png" httpUri-1="https://zenodo.org/record/6110212/files/figure.png" httpUri-2="https://zenodo.org/record/6110214/files/figure.png" pageId="3" pageNumber="552">Figs 1–3</figureCitation>
)
</paragraph>
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urn:lsid:zoobank.org:act:
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<subSubSection id="37ED6530AE4FFFC4FF5CF8B6F26BE29A" box="[151,813,1894,1921]" pageId="3" pageNumber="552" type="diagnosis">
<paragraph id="7F4836BBAE4FFFC4FF5CF8B6F26BE29A" blockId="3.[151,1436,1894,2030]" box="[151,813,1894,1921]" pageId="3" pageNumber="552">
<emphasis id="4D83EAA9AE4FFFC4FF5CF8B6F054E29B" bold="true" box="[151,274,1894,1920]" pageId="3" pageNumber="552">Diagnosis.</emphasis>
As for genus, as it is only known species so far.
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<subSubSection id="37ED6530AE4FFFC2FF0CF85AF0D3E492" lastPageId="5" lastPageNumber="554" pageId="3" pageNumber="552" type="description">
<paragraph id="7F4836BBAE4FFFC3FF0CF85AF206E4CA" blockId="3.[151,1436,1894,2030]" lastBlockId="4.[151,1437,151,825]" lastPageId="4" lastPageNumber="553" pageId="3" pageNumber="552">
<emphasis id="4D83EAA9AE4FFFC4FF0CF85AF01FE2BF" bold="true" box="[199,345,1930,1956]" pageId="3" pageNumber="552">Description.</emphasis>
<emphasis id="4D83EAA9AE4FFFC4FEAAF85BF0DDE2BE" box="[353,411,1931,1957]" italics="true" pageId="3" pageNumber="552">Male</emphasis>
(
<figureCitation id="E7CC2A3EAE4FFFC4FE60F85BF0AAE2BE" box="[427,492,1931,1957]" captionStart="FIGURE 1" captionStartId="2.[151,250,2008,2032]" captionTargetBox="[172,1415,1026,1982]" captionTargetId="figure-403@2.[172,1415,1026,1982]" captionTargetPageId="2" captionText="FIGURE 1. Afroscatopse haennii gen. et sp. nov., holotype male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6110210" httpUri="https://zenodo.org/record/6110210/files/figure.png" pageId="3" pageNumber="552">Fig. 1</figureCitation>
) large, with total length about 7.0 mm, whole body including wings dull, not shining, mostly black; pleurae more pale, scutum at scutellum with 3 yellowish elliptic spots covering also scutellum (2 lateral and 1 median), anterior shoulders also yellowish. Head rounded, as wide as high. Eyes emarginated at antennal bases, separated above by distance of an ocellus; hairs between ommatidia not visible (
<figureCitation id="E7CC2A3EAE48FFC3FB02FF47F401E5AA" box="[1225,1351,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-19@3.[151,1436,181,1585]" captionTargetPageId="3" captionText="FIGURE 2. Afroscatopse haennii gen. et sp. nov., holotype male, head and wing.A—head with antennae, B—vertex and frons, C—antenna, D—wing." figureDoi="http://doi.org/10.5281/zenodo.6110212" httpUri="https://zenodo.org/record/6110212/files/figure.png" pageId="4" pageNumber="553">Figs 2A, B</figureCitation>
). Three well developed ocelli present (
<figureCitation id="E7CC2A3EAE48FFC3FE24FF6BF301E5CE" box="[495,583,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-19@3.[151,1436,181,1585]" captionTargetPageId="3" captionText="FIGURE 2. Afroscatopse haennii gen. et sp. nov., holotype male, head and wing.A—head with antennae, B—vertex and frons, C—antenna, D—wing." figureDoi="http://doi.org/10.5281/zenodo.6110212" httpUri="https://zenodo.org/record/6110212/files/figure.png" pageId="4" pageNumber="553">Fig. 2B</figureCitation>
). Antennae much longer than head, a little bit longer then scutum (
<figureCitation id="E7CC2A3EAE48FFC3FAFEFF6BF4D6E5CE" box="[1333,1424,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-19@3.[151,1436,181,1585]" captionTargetPageId="3" captionText="FIGURE 2. Afroscatopse haennii gen. et sp. nov., holotype male, head and wing.A—head with antennae, B—vertex and frons, C—antenna, D—wing." figureDoi="http://doi.org/10.5281/zenodo.6110212" httpUri="https://zenodo.org/record/6110212/files/figure.png" pageId="4" pageNumber="553">Fig. 2A</figureCitation>
). Scape and pedicel of antenna short, each with irregularly distributed setulae (
<figureCitation id="E7CC2A3EAE48FFC3FBC9FF0FF51AE5E2" box="[1026,1116,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-19@3.[151,1436,181,1585]" captionTargetPageId="3" captionText="FIGURE 2. Afroscatopse haennii gen. et sp. nov., holotype male, head and wing.A—head with antennae, B—vertex and frons, C—antenna, D—wing." figureDoi="http://doi.org/10.5281/zenodo.6110212" httpUri="https://zenodo.org/record/6110212/files/figure.png" pageId="4" pageNumber="553">Fig. 2C</figureCitation>
). Flagellum
<quantity id="B80F9B5EAE48FFC3FB21FF0EF417E5E2" box="[1258,1361,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="553" unit="mm" value="1.5">1.50 mm</quantity>
, much longer than head and a little bit longer than scutum, composed of 9 uncompressed flagellomeres; flagellomere 1 longest, cylindrical, almost 2 times longer than next flagellomere; flagellomeres 2-5 cylindrical as long as broad, flagellomeres 6-8 more or less spherical, flagellomere 9 cylindrical 1.5 times longer than broad; each flagellomere with more or less distinct rings of short spine-like setae, flagellomere 1 with 4-5 irregular rings, flagellomeres 2-7 and 9 with 2 distinct rings, and flagellomere 8 with 1 ring (
<figureCitation id="E7CC2A3EAE48FFC3FC8BFE43F2DDE4B6" box="[832,923,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-19@3.[151,1436,181,1585]" captionTargetPageId="3" captionText="FIGURE 2. Afroscatopse haennii gen. et sp. nov., holotype male, head and wing.A—head with antennae, B—vertex and frons, C—antenna, D—wing." figureDoi="http://doi.org/10.5281/zenodo.6110212" httpUri="https://zenodo.org/record/6110212/files/figure.png" pageId="4" pageNumber="553">Fig. 2C</figureCitation>
). Frons bare, separated from clypeus bearing setae (
<figureCitation id="E7CC2A3EAE48FFC3FF15FE67F071E4CA" box="[222,311,439,465]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-19@3.[151,1436,181,1585]" captionTargetPageId="3" captionText="FIGURE 2. Afroscatopse haennii gen. et sp. nov., holotype male, head and wing.A—head with antennae, B—vertex and frons, C—antenna, D—wing." figureDoi="http://doi.org/10.5281/zenodo.6110212" httpUri="https://zenodo.org/record/6110212/files/figure.png" pageId="4" pageNumber="553">Fig. 2B</figureCitation>
). Maxillary palpus one-segmented, very short.
</paragraph>
<paragraph id="7F4836BBAE48FFC3FF0CFE0BF2F5E77A" blockId="4.[151,1437,151,825]" pageId="4" pageNumber="553">
Thorax of
<typeStatus id="A04C8819AE48FFC3FEF0FE0BF0DBE4EE" box="[315,413,475,501]" pageId="4" pageNumber="553" type="holotype">holotype</typeStatus>
with proportions of length/width/height: 26/22/27. Scutum almost as long (26) as wide (22); widest at wing bases, narrower at head (
<figureCitation id="E7CC2A3EAE48FFC3FDA9FE2FF3E3E702" box="[610,677,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,2008,2032]" captionTargetBox="[172,1415,1026,1982]" captionTargetId="figure-403@2.[172,1415,1026,1982]" captionTargetPageId="2" captionText="FIGURE 1. Afroscatopse haennii gen. et sp. nov., holotype male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6110210" httpUri="https://zenodo.org/record/6110210/files/figure.png" pageId="4" pageNumber="553">Fig. 1</figureCitation>
). Scutum bare except for lateral margins bearing irregular rows of short setulae; scutellum bare, without marginal setae. Spiracular sclerite distinct and setose, almost rectangular, with central opening; postpronotum bearing short setae, katepisternum bare.
</paragraph>
<paragraph id="7F4836BBAE48FFC3FF0CFDBBF5C4E622" blockId="4.[151,1437,151,825]" pageId="4" pageNumber="553">
Wing very broad, almost 2 times as long as broad, length
<quantity id="B80F9B5EAE48FFC3FC8AFDBAF2A7E79E" box="[833,993,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="4.91" metricValueMax="5.14" metricValueMin="4.68" pageId="4" pageNumber="553" unit="mm" value="4.91" valueMax="5.14" valueMin="4.68">4.68-5.14 mm</quantity>
. Subcosta very short, not reaching costa at wing margin (
<figureCitation id="E7CC2A3EAE48FFC3FE86FD5FF0E1E7B2" box="[333,423,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-19@3.[151,1436,181,1585]" captionTargetPageId="3" captionText="FIGURE 2. Afroscatopse haennii gen. et sp. nov., holotype male, head and wing.A—head with antennae, B—vertex and frons, C—antenna, D—wing." figureDoi="http://doi.org/10.5281/zenodo.6110212" httpUri="https://zenodo.org/record/6110212/files/figure.png" pageId="4" pageNumber="553">Fig. 2D</figureCitation>
). Costal vein swollen on proximal 1/3, R
<subScript id="E37334FEAE48FFC3FCA7FD71F2C3E7B4" attach="left" box="[876,901,673,687]" fontSize="6" pageId="4" pageNumber="553">2+3</subScript>
lost. R
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long, reaching margin near wing apex. Medial vein M
<subScript id="E37334FEAE48FFC3FEF6FD15F003E7C8" attach="left" box="[317,325,709,723]" fontSize="6" pageId="4" pageNumber="553">1</subScript>
angled, reduced on proximal part; a short spur at angle present; M
<subScript id="E37334FEAE48FFC3FBE6FD15F573E7C8" attach="left" box="[1069,1077,709,723]" fontSize="6" pageId="4" pageNumber="553">2</subScript>
well developed from transverse vein r-m to wing apex; M
<subScript id="E37334FEAE48FFC3FE73FD39F097E7EC" attach="left" box="[440,465,745,759]" fontSize="6" pageId="4" pageNumber="553">3+4</subScript>
or M
<subScript id="E37334FEAE48FFC3FDC6FD39F353E7EC" attach="left" box="[525,533,745,759]" fontSize="6" pageId="4" pageNumber="553">4</subScript>
and CuA well visible, both gradually C-curved; all veins reaching wing margin. Wing membrane with distinct microtrichia; macrotrichia present only on costal vein and on dorsal surface of R
<subScript id="E37334FEAE48FFC3FA46FCDDF4D3E600" attach="left" box="[1421,1429,781,795]" fontSize="6" pageId="4" pageNumber="553">1</subScript>
, absent on wing membrane and other veins. Stem of haltere with a row of spine-like setae.
</paragraph>
<caption id="2B886633AE48FFC3FF5CF8B8F0BDE2D3" ID-DOI="http://doi.org/10.5281/zenodo.6110214" ID-Zenodo-Dep="6110214" httpUri="https://zenodo.org/record/6110214/files/figure.png" pageId="4" pageNumber="553" startId="4.[151,250,1896,1920]" targetBox="[151,1436,849,1873]" targetPageId="4">
<paragraph id="7F4836BBAE48FFC3FF5CF8B8F0BDE2D3" blockId="4.[151,1436,1896,1992]" pageId="4" pageNumber="553">
<emphasis id="4D83EAA9AE48FFC3FF5CF8B8F055E29B" bold="true" box="[151,275,1896,1920]" pageId="4" pageNumber="553">FIGURE 3.</emphasis>
<taxonomicName id="B8F74D38AE48FFC3FED2F8B9F0A9E29B" authority="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo, 2022" authorityName="Szadziewski &amp; Bojarski &amp; Sontag &amp; Szwedo" authorityYear="2022" box="[281,495,1896,1921]" class="Insecta" family="Scatopsidae" genus="Afroscatopse" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="553" phylum="Arthropoda" rank="species" species="haennii" status="gen. et sp. nov.">
<emphasis id="4D83EAA9AE48FFC3FED2F8B9F0A9E29B" box="[281,495,1896,1921]" italics="true" pageId="4" pageNumber="553">Afroscatopse haennii</emphasis>
</taxonomicName>
<emphasis id="4D83EAA9AE48FFC3FE3EF8B9F3D4E29A" bold="true" box="[501,658,1897,1921]" pageId="4" pageNumber="553">
<taxonomicNameLabel id="56B057D2AE48FFC3FE3EF8B9F3D4E29A" box="[501,658,1897,1921]" pageId="4" pageNumber="553" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, male genitalia. A—ventral view, phot., B—ventral aspect, drawing, C—laterodorsal view, phot., D—laterodorsal aspect of tergite 7, and gonocoxite. Abbreviations: st7—abdominal sternite 7, tg7—abdominal tergite 7, gx—gonocoxite.
</paragraph>
</caption>
<paragraph id="7F4836BBAE49FFC2FF0CFF47F382E5E2" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="554">
Legs relatively stout, without modifications. Tibial spurs absent. Tarsi short, 5-segmented, tarsomere 1 as long as 2-4 together, claws equal, simple, empodium well developed; tarsus of hind leg about
<quantity id="B80F9B5EAE49FFC2FBB5FF6BF592E5CE" box="[1150,1236,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="554" unit="mm" value="0.9">0.9 mm</quantity>
long, proportions of tarsomeres as follows: 1—37, 2-4—35, 5—27.
</paragraph>
<paragraph id="7F4836BBAE49FFC2FF0CFED3F3CBE47E" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="554">
Genitalia barely visible, not rotated, symmetrical. Sternite 7 with 4 caudal projections (
<figureCitation id="E7CC2A3EAE49FFC2FB46FED3F5A2E406" box="[1165,1252,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1896,1920]" captionTargetBox="[151,1436,849,1873]" captionTargetId="figure-359@4.[151,1436,849,1873]" captionTargetPageId="4" captionText="FIGURE 3. Afroscatopse haennii gen. et sp. nov., male genitalia. A—ventral view, phot., B—ventral aspect, drawing, C—lat- erodorsal view, phot., D—laterodorsal aspect of tergite 7, and gonocoxite. Abbreviations: st7—abdominal sternite 7, tg7—ab- dominal tergite 7, gx—gonocoxite." figureDoi="http://doi.org/10.5281/zenodo.6110214" httpUri="https://zenodo.org/record/6110214/files/figure.png" pageId="5" pageNumber="554">Fig. 3B</figureCitation>
), tergite 7 divided into 2 long and blunt, bare, divergent lobes (
<figureCitation id="E7CC2A3EAE49FFC2FD6EFEF7F260E45A" box="[677,806,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1896,1920]" captionTargetBox="[151,1436,849,1873]" captionTargetId="figure-359@4.[151,1436,849,1873]" captionTargetPageId="4" captionText="FIGURE 3. Afroscatopse haennii gen. et sp. nov., male genitalia. A—ventral view, phot., B—ventral aspect, drawing, C—lat- erodorsal view, phot., D—laterodorsal aspect of tergite 7, and gonocoxite. Abbreviations: st7—abdominal sternite 7, tg7—ab- dominal tergite 7, gx—gonocoxite." figureDoi="http://doi.org/10.5281/zenodo.6110214" httpUri="https://zenodo.org/record/6110214/files/figure.png" pageId="5" pageNumber="554">Figs 3A–D</figureCitation>
); gonoxite broad with pointed apical projection, covered with long setae, gonostyli reduced (
<figureCitation id="E7CC2A3EAE49FFC2FDEDFE9BF3C6E47E" box="[550,640,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1896,1920]" captionTargetBox="[151,1436,849,1873]" captionTargetId="figure-359@4.[151,1436,849,1873]" captionTargetPageId="4" captionText="FIGURE 3. Afroscatopse haennii gen. et sp. nov., male genitalia. A—ventral view, phot., B—ventral aspect, drawing, C—lat- erodorsal view, phot., D—laterodorsal aspect of tergite 7, and gonocoxite. Abbreviations: st7—abdominal sternite 7, tg7—ab- dominal tergite 7, gx—gonocoxite." figureDoi="http://doi.org/10.5281/zenodo.6110214" httpUri="https://zenodo.org/record/6110214/files/figure.png" pageId="5" pageNumber="554">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph id="7F4836BBAE49FFC2FF0CFEBFF0D3E492" blockId="5.[151,1437,150,645]" box="[199,405,367,393]" pageId="5" pageNumber="554">
<emphasis id="4D83EAA9AE49FFC2FF0CFEBFF058E492" bold="true" box="[199,286,367,393]" pageId="5" pageNumber="554">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="37ED6530AE49FFC2FF0CFE42F03AE725" pageId="5" pageNumber="554" type="materials_examined">
<paragraph id="7F4836BBAE49FFC2FF0CFE42F03AE725" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="554">
<emphasis id="4D83EAA9AE49FFC2FF0CFE42F0EDE4B7" bold="true" box="[199,427,402,428]" pageId="5" pageNumber="554">Material examined</emphasis>
.
<materialsCitation id="CF9F3CE6AE49FFC2FE70FE43F3ADE4B5" ID-GBIF-Occurrence="3467094302" box="[443,747,403,430]" pageId="5" pageNumber="554" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="A04C8819AE49FFC2FE70FE43F365E4B6" box="[443,547,403,429]" pageId="5" pageNumber="554" type="holotype">Holotype</typeStatus>
<specimenCount id="69F1FD32AE49FFC2FDE7FE43F324E4B6" box="[556,610,403,429]" pageId="5" pageNumber="554" type="male">male</specimenCount>
(complete),
</materialsCitation>
<materialsCitation id="CF9F3CE6AE49FFC2FD38FE43F568E4B5" ID-GBIF-Occurrence="3467094301" box="[755,1070,403,430]" pageId="5" pageNumber="554" typeStatus="paratype">
<typeStatus id="A04C8819AE49FFC2FD38FE43F215E4B6" box="[755,851,403,429]" pageId="5" pageNumber="554" type="paratype">paratype</typeStatus>
male (incomplete)
</materialsCitation>
, and
<materialsCitation id="CF9F3CE6AE49FFC2FBA0FE43F080E4CA" ID-GBIF-Occurrence="3467094303" pageId="5" pageNumber="554">head of third specimen not included to the type series</materialsCitation>
,
<collectionCode id="19E6AE7EAE49FFC2FE1FFE67F359E4CA" box="[468,543,439,465]" pageId="5" pageNumber="554">MAIG</collectionCode>
6013. Syninclusions:
<taxonomicName id="B8F74D38AE49FFC2FCD6FE67F2DAE4CA" authorityName="Linnaeus" authorityYear="1758" box="[797,924,439,465]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="B8F74D38AE49FFC2FC62FE67F505E4CA" box="[937,1091,439,465]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
1,
<taxonomicName id="B8F74D38AE49FFC2FBA1FE67F5A3E4CA" box="[1130,1253,439,465]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
:
<taxonomicName id="B8F74D38AE49FFC2FB3AFE67F40CE4CA" box="[1265,1354,439,465]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="family">Miridae</taxonomicName>
1, Hymenoptera:
<taxonomicName id="B8F74D38AE49FFC2FED5FE0BF085E4EE" authorityName="Haliday" authorityYear="1833" box="[286,451,475,501]" class="Insecta" family="Ceraphronidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="family">Ceraphronidae</taxonomicName>
1, Isoptera 1 winged, Acari at least 2;
<taxonomicName id="B8F74D38AE49FFC2FC4CFE0BF54BE4EE" authorityName="Newman" authorityYear="1834" box="[903,1037,475,501]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="family">Scatopsidae</taxonomicName>
1;
<taxonomicName id="B8F74D38AE49FFC2FBFEFE0BF5FAE4EE" authorityName="Linnaeus" authorityYear="1758" box="[1077,1212,475,501]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
1. Abdomen of the
<typeStatus id="A04C8819AE49FFC2FF5CFE2FF1BFE702" box="[151,249,511,537]" pageId="5" pageNumber="554" type="holotype">holotype</typeStatus>
is covered by milky veil. Till now such a veil was observed only in not heated Baltic amber inclusions (
<bibRefCitation id="1B664B4AAE49FFC2FF54FDF3F036E725" author="Szadziewski, R." box="[159,368,547,574]" pageId="5" pageNumber="554" pagination="92 - 97" refId="ref8070" refString="Szadziewski, R. (2006) Forgeries of Baltic amber inclusions [2001]. In: Kosmowska-Ceranowicz, B. &amp; Gierlowski, W. (Eds.), Amber - views, opinions. Scientific seminars AMBERIF - International Fair of Amber, Jewellery and Gemstones 1994 - 2005. The International Amber Association, The Museum of the Earth of the Polish Academy of Sciences. Gdansk Inter- national Fair Co., Gdansk, Warsaw, pp. 92 - 97." type="book chapter" year="2006">Szadziewski 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="37ED6530AE49FFC2FF0CFD97F5CAE79E" pageId="5" pageNumber="554" type="etymology">
<paragraph id="7F4836BBAE49FFC2FF0CFD97F5CAE79E" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="554">
<emphasis id="4D83EAA9AE49FFC2FF0CFD97F008E77A" bold="true" box="[199,334,583,609]" pageId="5" pageNumber="554">Etymology.</emphasis>
The new species is dedicated to Jean-Paul Haenni of the Muséum d’histoire naturelle,
<collectingRegion id="BD33F859AE49FFC2FAEFFD97F4DEE77A" box="[1316,1432,583,609]" country="Switzerland" name="Neuchatel" pageId="5" pageNumber="554">Neuchâtel</collectingRegion>
,
<collectingCountry id="07E0762BAE49FFC2FF5CFDBBF066E79E" box="[151,288,619,645]" name="Switzerland" pageId="5" pageNumber="554">Switzerland</collectingCountry>
, in recognition of his important contributions to the taxonomy of
<taxonomicName id="B8F74D38AE49FFC2FC35FDBBF5C1E79E" authorityName="Newman" authorityYear="1834" box="[1022,1159,619,645]" class="Insecta" family="Scatopsidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="family">Scatopsidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>