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data/F7/42/45/F74245F538F41979D6EE628FE0D5B443.xml Normal file
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.245.3416" ID-PMC="PMC3577090" ID-Pensoft-Pub="1313-2970-245-1" ID-Pensoft-UUID="FFFF52503A0AFF882450FFB66D45FF8E" ID-PubMed="23431087" ID-Zenodo-Dep="578462" ModsDocID="1313-2970-245-1" checkinTime="1425642919252" checkinUser="pensoft" docAuthor="Bousquet, Yves" docDate="2012" docId="F74245F538F41979D6EE628FE0D5B443" docLanguage="en" docName="ZooKeys 245: 1-1722" docOrigin="ZooKeys 245" docPubDate="2012-11-28" docSource="http://dx.doi.org/10.3897/zookeys.245.3416" docTitle="Loxandrus nitidulus" docType="treatment" docVersion="10" id="FFFF52503A0AFF882450FFB66D45FF8E" lastPageNumber="492" masterDocId="FFFF52503A0AFF882450FFB66D45FF8E" masterDocTitle="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico" masterLastPageNumber="1722" masterPageNumber="1" pageNumber="492" updateTime="1668154792637" updateUser="ExternalLinkService">
<mods:mods id="AE556E38BAF46F264F3A46079D83F9B9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EA1255381426DE25BBFC03809E0B37BB">
<mods:title id="EE3715A2E20E77AA7342776C81C3FBB9">Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico</mods:title>
</mods:titleInfo>
<mods:name id="73E8854D6C5E1836B65E481699D7B143" type="personal">
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</mods:role>
<mods:namePart id="85E5A882265656A767EF87E865300418">Bousquet, Yves</mods:namePart>
<mods:affiliation id="EAAA3B883F870B92A42DCF4971E36D89">Agriculture and Agri-Food Canada, Central Experimental Farm, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier id="DD68BFA4396234136736A51F33326F07" type="email">bousquety1@yahoo.com</mods:nameIdentifier>
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<mods:typeOfResource id="5FB39A8720F6BC3E688316C66378947B">text</mods:typeOfResource>
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<mods:title id="E04C1CF3618CF0D63D80A0093235D096">ZooKeys</mods:title>
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<mods:part id="C298FD33FD645F74EB516D122860417F">
<mods:date id="83F02F2DAE65FC8C2F80648A79DCB157">2012</mods:date>
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<mods:number id="9340E54B76529B2CACDE662C5B7D8BF4">2012-11-28</mods:number>
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<mods:detail id="29A37B83D552F486DACC09BAA80A8F5F" type="volume">
<mods:number id="0CFF2D9D26EB974ED6837AD7C23B8D7A">245</mods:number>
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<mods:identifier id="609F7162F3D122F1A555EB272D377EF2" type="DOI">http://dx.doi.org/10.3897/zookeys.245.3416</mods:identifier>
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<mods:identifier id="F3528825740078DB3AAA785401BECE07" type="Zenodo-Dep">578462</mods:identifier>
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<treatment id="F74245F538F41979D6EE628FE0D5B443" ID-GBIF-Taxon="127865838" LSID="urn:lsid:plazi:treatment:F74245F538F41979D6EE628FE0D5B443" httpUri="http://treatment.plazi.org/id/F74245F538F41979D6EE628FE0D5B443" lastPageNumber="492" pageId="491" pageNumber="492">
<subSubSection id="584806057904CC6B36D3100BEA67A212" pageId="491" pageNumber="492" type="nomenclature">
<paragraph id="DD69C89821BFF7D243AEDEC10E9371AF" pageId="491" pageNumber="492">
<taxonomicName id="5414AF2DF75E3A5DFA25B5F944806AB6" ID-CoL="8LDHV" LSID="F74245F5-38F4-1979-D6EE-628FE0D5B443" authority="(LeConte, 1846)" baseAuthorityName="LeConte" baseAuthorityYear="1846" class="Insecta" family="Carabidae" genus="Loxandrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Loxandrus nitidulus" order="Coleoptera" pageId="491" pageNumber="492" phylum="Arthropoda" rank="species" species="nitidulus">Loxandrus nitidulus (LeConte, 1846)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="74FAD18F3AE12DE3573F38D5886074C4" pageId="491" pageNumber="492" type="reference_group">
<paragraph id="ABFD78D0CFBAD2588C8D035B4208814F" pageId="491" pageNumber="492">
<taxonomicName id="55A57B5E0E23117881CD8AF27C1D8A6B" class="Hexapoda" family="Carabidae" genus="Argutor" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="491" pageNumber="492" phylum="Arthropoda" rank="species" species="nitidulus">Argutor nitidulus</taxonomicName>
LeConte, 1846b: 339. Type locality: &quot;provinciis australibus&quot; (original citation), herein restricted to Lucedale, George County, Mississippi (see Allen 1972: 153). Four syntypes in MCZ [# 5696].
</paragraph>
</subSubSection>
<subSubSection id="C5266B585EDBFBB57B0A7F6E78D6D805" pageId="491" pageNumber="492" type="distribution">
<paragraph id="D782B8249B062C8CDC3E270DA429A7E1" pageId="491" pageNumber="492">Distribution.</paragraph>
<paragraph id="06594C7C51F60E52C8B0B066673F1928" pageId="491" pageNumber="492">This species ranges from northern Indiana to eastern Iowa, south to northern Louisiana and southwestern Alabama [see Allen 1972: Fig. 161). One specimen simply labeled from Florida is known (Allen 1972: 153).</paragraph>
</subSubSection>
<subSubSection id="B181FE4EBFB854C91A38B064E6A5B3BB" pageId="491" pageNumber="492" type="records">
<paragraph id="41A7CBB1EEF5A3D303B7F23F0BDF3C4E" pageId="491" pageNumber="492">Records.</paragraph>
<paragraph id="1A653F858ED4E8DF0A58D292ED44EB7C" pageId="491" pageNumber="492">
<emphasis id="EE7F5F7E9BF366EA216599FD6142EB56" bold="true" pageId="491" pageNumber="492">USA</emphasis>
: AL, AR, IA, IL, IN, LA, MO, MS, TN [FL]
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.944.53099" ID-GBIF-Dataset="19d82844-0ce4-4edd-9bd7-e4a8b62fc533" ID-PMC="PMC7340628" ID-Pensoft-Pub="1313-2970-944-129" ID-Pensoft-UUID="D3B78C8AAD25541380CB22698BDA841B" ID-PubMed="32684777" ID-ZooBank="F00B2CAF5D9548F59C580DD95AAC9B8B" ModsDocID="1313-2970-944-129" checkinTime="1593557116768" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Beenen, Ron" docDate="2020" docId="F7427B541E0954FA802D255150E29E03" docLanguage="en" docName="ZooKeys 944: 129-146" docOrigin="ZooKeys 944" docSource="http://dx.doi.org/10.3897/zookeys.944.53099" docTitle="Taiwanoshaira taipingshanensis Lee &amp; Beenen 2020, sp. nov." docType="treatment" docUuid="1AE3EA77-43D4-420C-A3F3-E92F3CA65A9B" docUuidSource="ZooBank" docVersion="4" id="D3B78C8AAD25541380CB22698BDA841B" lastPageNumber="129" masterDocId="D3B78C8AAD25541380CB22698BDA841B" masterDocTitle="Taiwanoshaira Lee &amp; Beenen, a new genus and first record of moss-inhabiting Galerucinae sensu stricto (Coleoptera, Chrysomelidae) from Taiwan" masterLastPageNumber="146" masterPageNumber="129" pageNumber="129" updateTime="1668169024658" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taiwanoshaira Lee &amp; Beenen, a new genus and first record of moss-inhabiting Galerucinae sensu stricto (Coleoptera, Chrysomelidae) from Taiwan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
<mods:affiliation>Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-1996-0557</mods:nameIdentifier>
<mods:nameIdentifier type="email">chifeng@tari.gov.tw</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beenen, Ron</mods:namePart>
<mods:affiliation>Martinus Nijhoffhove 51, NL- 3437 ZP Nieuwegein, The Netherlands</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
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<mods:number>944</mods:number>
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<mods:start>129</mods:start>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.944.53099</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-944-129</mods:identifier>
<mods:identifier type="ZooBank">F00B2CAF5D9548F59C580DD95AAC9B8B</mods:identifier>
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<treatment ID-GBIF-Taxon="164844950" LSID="urn:lsid:zoobank.org:act:1AE3EA77-43D4-420C-A3F3-E92F3CA65A9B" httpUri="http://treatment.plazi.org/id/F7427B541E0954FA802D255150E29E03" lastPageNumber="129" pageId="0" pageNumber="129">
<subSubSection pageId="0" pageNumber="129" type="nomenclature">
<paragraph pageId="0" pageNumber="129">
<taxonomicName LSID="F7427B54-1E09-54FA-802D-255150E29E03" authority="Lee &amp; Beenen, 2020" authorityName="Lee &amp; Beenen" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Taiwanoshaira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Taiwanoshaira taipingshanensis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="taipingshanensis" status="sp. nov.">Taiwanoshaira taipingshanensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="129">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of Taiwanoshaira taipingshanensis sp. nov. and T. tsoui sp. nov. A T. taipingshanensis sp. nov., male, dorsal view B same, ventral view C same, female, dorsal view D T. tsoui sp. nov., male, dorsal view E same, ventral view F same, female, dorsal view." figureDoi="10.3897/zookeys.944.53099.figure6" httpUri="https://binary.pensoft.net/fig/426849" pageId="0" pageNumber="129">Figures 6A-C</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="types">
<paragraph pageId="0" pageNumber="129">Types</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">
(
<emphasis italics="true" pageId="0" pageNumber="129">N</emphasis>
= 77).
<emphasis italics="true" pageId="0" pageNumber="129">Holotype</emphasis>
</emphasis>
♂ (TARI): Taiwan. Ilan: Taipingshan (太平山), 5.VIII.2015, leg. Y.-T. Chung.
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Paratypes</emphasis>
.
</emphasis>
5♂♂, 15♀♀ (TARI), same as holotype; 10♂♂, 16♀♀ (TARI), Yuanyanghu (鴛鴦湖), 19.VIII.2010, leg. S.-S. Li; 4♂♂, 5♀♀ (TARI), same locality, 22.VIII.2011, leg. C.-F. Lee; 2♂♂, 8♀♀ (2♀♀: TARI; 2♂♂, 2♀♀: RBCN), same but with &quot;leg. M.-H. Tsou&quot;; 5♂♂, 6♀♀ (TARI), same but with &quot;leg. H. Lee&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="description">
<paragraph pageId="0" pageNumber="129">Description.</paragraph>
<paragraph pageId="0" pageNumber="129">
Length 4.0-5.7 mm, width 2.6-3.4 mm. General color dark brown or blackish-brown (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of Taiwanoshaira taipingshanensis sp. nov. and T. tsoui sp. nov. A T. taipingshanensis sp. nov., male, dorsal view B same, ventral view C same, female, dorsal view D T. tsoui sp. nov., male, dorsal view E same, ventral view F same, female, dorsal view." figureDoi="10.3897/zookeys.944.53099.figure6" httpUri="https://binary.pensoft.net/fig/426849" pageId="0" pageNumber="129">6A-C</figureCitation>
). Antenna (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7A</figureCitation>
) filiform in males, ratios of lengths of antennomeres I to XI 1.0: 0.5: 0.6: 0.7: 0.7: 0.7: 0.7: 0.7: 0.6: 0.7: 0.8; ratios of lengths to widths from antennomeres I to XI 3.0: 2.0: 2.2: 3.0: 3.2: 3.2: 3.3: 3.5: 3.0: 3.1: 3.4; similar in females, ratio of lengths of antennomeres I to XI (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7B</figureCitation>
) 1.0: 0.4: 0.5: 0.8: 0.7: 0.7: 0.7: 0.7: 0.7: 0.7: 0.8; ratios of lengths to widths from antennomeres I to XI 3.4: 2.3: 2.3: 3.9: 3.7: 3.7: 4.0: 3.7: 3.4: 3.5: 4.2. Pronotum 1.61-1.63 times wider than long; lateral margins slightly rounded; disc with fine punctures bearing tiny setae. Procoxal cavities almost closed. Elytra 1.11-1.20 times longer than wide; disc with dense, coarse punctures and longitudinal ridges not covered with punctures; apices tapering in both sexes. Tarsomeres I of front legs slightly swollen in males, not modified in females. Penis (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7C-E</figureCitation>
) wide, about 5.5 times longer than wide; parallel sided and strongly curved in lateral view, apex narrowly rounded, base with shallow median notch; tectum broad from apical 1/6 to middle, apex truncate; ventral surface with large opening. Endophallic spiculae complex (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7F, G</figureCitation>
) with median endophallic spiculae slender, apically bifurcate, and straight in lateral view; with one pair of small sclerites near base. Gonocoxae (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7H, I</figureCitation>
) short; apex of each gonocoxa widely rounded, with eight to ten long setae along apical margin, basal margin irregular. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7J, K</figureCitation>
) short and well sclerotized, with several short setae along apical margin, spiculum long. Spermathecal receptaculum (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7L</figureCitation>
) strongly swollen; pump extremely slender and curved; sclerotized spermathecal duct long.
</paragraph>
<caption doi="10.3897/zookeys.944.53099.figure6" httpUri="https://binary.pensoft.net/fig/426849" pageId="0" pageNumber="129" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 6.</emphasis>
Habitus of
<taxonomicName authorityName="Lee &amp; Beenen" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Taiwanoshaira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Taiwanoshaira taipingshanensis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="taipingshanensis">
<emphasis italics="true" pageId="0" pageNumber="129">Taiwanoshaira taipingshanensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="T. tsoui" pageId="0" pageNumber="129" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="129">T. tsoui</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
<taxonomicName lsidName="T. taipingshanensis" pageId="0" pageNumber="129" rank="species" species="taipingshanensis">
<emphasis italics="true" pageId="0" pageNumber="129">T. taipingshanensis</emphasis>
</taxonomicName>
sp. nov., male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
same, ventral view
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
same, female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
<taxonomicName lsidName="T. tsoui" pageId="0" pageNumber="129" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="129">T. tsoui</emphasis>
</taxonomicName>
sp. nov., male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="129">E</emphasis>
same, ventral view
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
same, female, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="variation">
<paragraph pageId="0" pageNumber="129">Variation.</paragraph>
<paragraph pageId="0" pageNumber="129">
Female genitalic characters are variable among different localities. The apices of the gonocoxae are widely rounded in specimens from Taipingshan (太平山) (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7I</figureCitation>
) but tapering in those from Yuanyanhu (鴛鴦湖) (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7H</figureCitation>
). The apex of abdominal ventrite VIII is shorter in specimens from Taipingshan (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7K</figureCitation>
) than those from Yuanyanhu (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7J</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 7.</emphasis>
Diagnostic characters of
<taxonomicName authorityName="Lee &amp; Beenen" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Taiwanoshaira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Taiwanoshaira taipingshanensis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="taipingshanensis">
<emphasis italics="true" pageId="0" pageNumber="129">Taiwanoshaira taipingshanensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
antenna, male
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
antenna, female
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
penis, apex, dorsal view
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
penis, base, dorsal view
<emphasis bold="true" pageId="0" pageNumber="129">E</emphasis>
penis, lateral view
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
endophallic sclerites, dorsal view
<emphasis bold="true" pageId="0" pageNumber="129">G</emphasis>
ditto, lateral view
<emphasis bold="true" pageId="0" pageNumber="129">H</emphasis>
gonocoxae, from Yuanyanghu (鴛鴦湖)
<emphasis bold="true" pageId="0" pageNumber="129">I</emphasis>
same, from Taipingshan (太平山)
<emphasis bold="true" pageId="0" pageNumber="129">J</emphasis>
abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖)
<emphasis bold="true" pageId="0" pageNumber="129">K</emphasis>
same, from Taipingshan (太平山)
<emphasis bold="true" pageId="0" pageNumber="129">L</emphasis>
spermatheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Lee &amp; Beenen" authorityYear="2020" class="Insecta" family="Chrysomelidae" genus="Taiwanoshaira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Taiwanoshaira taipingshanensis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="taipingshanensis">
<emphasis italics="true" pageId="0" pageNumber="129">Taiwanoshaira taipingshanensis</emphasis>
</taxonomicName>
sp. nov. is easily separated from other congeners by the presence of longitudinal ridges on the elytra (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of Taiwanoshaira taipingshanensis sp. nov. and T. tsoui sp. nov. A T. taipingshanensis sp. nov., male, dorsal view B same, ventral view C same, female, dorsal view D T. tsoui sp. nov., male, dorsal view E same, ventral view F same, female, dorsal view." figureDoi="10.3897/zookeys.944.53099.figure6" httpUri="https://binary.pensoft.net/fig/426849" pageId="0" pageNumber="129">6A, C</figureCitation>
) (lacking longitudinal ridges in others (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Taiwanoshaira chujoi comb. nov. A male, paratype, from Alishan (阿里山), dorsal view B same, ventral view C female, paratype, from Tapan (達邦), dorsal view D male, from Bilu Divine Tree (碧綠神木), dorsal view E same, ventral view F female, from Bilu Divine Tree (碧綠神木), dorsal view." figureDoi="10.3897/zookeys.944.53099.figure3" httpUri="https://binary.pensoft.net/fig/426846" pageId="0" pageNumber="129">3A, C-D, F</figureCitation>
;
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of Taiwanoshaira taipingshanensis sp. nov. and T. tsoui sp. nov. A T. taipingshanensis sp. nov., male, dorsal view B same, ventral view C same, female, dorsal view D T. tsoui sp. nov., male, dorsal view E same, ventral view F same, female, dorsal view." figureDoi="10.3897/zookeys.944.53099.figure6" httpUri="https://binary.pensoft.net/fig/426849" pageId="0" pageNumber="129">6D, F</figureCitation>
)), almost closed procoxal cavities (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Generic characters for Taiwanoshaira. A prothorax, ventral view, T. taipingshanensis sp. nov. B prothorax, ventral view, T. tsoui sp. nov. C last two abdominal ventrites, dorsal view, male, T. tsoui sp. nov. D elytron, ventral view, T. tsoui sp. nov." figureDoi="10.3897/zookeys.944.53099.figure2" httpUri="https://binary.pensoft.net/fig/426845" pageId="0" pageNumber="129">2A</figureCitation>
) (widely open procoxal cavities (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Generic characters for Taiwanoshaira. A prothorax, ventral view, T. taipingshanensis sp. nov. B prothorax, ventral view, T. tsoui sp. nov. C last two abdominal ventrites, dorsal view, male, T. tsoui sp. nov. D elytron, ventral view, T. tsoui sp. nov." figureDoi="10.3897/zookeys.944.53099.figure2" httpUri="https://binary.pensoft.net/fig/426845" pageId="0" pageNumber="129">2B</figureCitation>
) in others), entirely black legs (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of Taiwanoshaira taipingshanensis sp. nov. and T. tsoui sp. nov. A T. taipingshanensis sp. nov., male, dorsal view B same, ventral view C same, female, dorsal view D T. tsoui sp. nov., male, dorsal view E same, ventral view F same, female, dorsal view." figureDoi="10.3897/zookeys.944.53099.figure6" httpUri="https://binary.pensoft.net/fig/426849" pageId="0" pageNumber="129">6A-C</figureCitation>
) (yellowish-brown legs with dark apices of femora and bases of tibiae in others (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Taiwanoshaira chujoi comb. nov. A male, paratype, from Alishan (阿里山), dorsal view B same, ventral view C female, paratype, from Tapan (達邦), dorsal view D male, from Bilu Divine Tree (碧綠神木), dorsal view E same, ventral view F female, from Bilu Divine Tree (碧綠神木), dorsal view." figureDoi="10.3897/zookeys.944.53099.figure3" httpUri="https://binary.pensoft.net/fig/426846" pageId="0" pageNumber="129">3</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of Taiwanoshaira taipingshanensis sp. nov. and T. tsoui sp. nov. A T. taipingshanensis sp. nov., male, dorsal view B same, ventral view C same, female, dorsal view D T. tsoui sp. nov., male, dorsal view E same, ventral view F same, female, dorsal view." figureDoi="10.3897/zookeys.944.53099.figure6" httpUri="https://binary.pensoft.net/fig/426849" pageId="0" pageNumber="129">6D-F</figureCitation>
)), sexually dimorphic protarsi I (uniform protarsi I in both sexes of others), tapering elytral apices of both sexes (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of Taiwanoshaira taipingshanensis sp. nov. and T. tsoui sp. nov. A T. taipingshanensis sp. nov., male, dorsal view B same, ventral view C same, female, dorsal view D T. tsoui sp. nov., male, dorsal view E same, ventral view F same, female, dorsal view." figureDoi="10.3897/zookeys.944.53099.figure6" httpUri="https://binary.pensoft.net/fig/426849" pageId="0" pageNumber="129">6A-C</figureCitation>
) (tapering elytra apices in only males of others (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Taiwanoshaira chujoi comb. nov. A male, paratype, from Alishan (阿里山), dorsal view B same, ventral view C female, paratype, from Tapan (達邦), dorsal view D male, from Bilu Divine Tree (碧綠神木), dorsal view E same, ventral view F female, from Bilu Divine Tree (碧綠神木), dorsal view." figureDoi="10.3897/zookeys.944.53099.figure3" httpUri="https://binary.pensoft.net/fig/426846" pageId="0" pageNumber="129">3</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of Taiwanoshaira taipingshanensis sp. nov. and T. tsoui sp. nov. A T. taipingshanensis sp. nov., male, dorsal view B same, ventral view C same, female, dorsal view D T. tsoui sp. nov., male, dorsal view E same, ventral view F same, female, dorsal view." figureDoi="10.3897/zookeys.944.53099.figure6" httpUri="https://binary.pensoft.net/fig/426849" pageId="0" pageNumber="129">6D-F</figureCitation>
)). In addition, most genitalic characters of this new species are diagnostic, including the extremely strongly curved penis (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7C-E</figureCitation>
) (moderately or slightly curved in others (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Taiwanoshaira chujoi comb. nov. A antenna, male B antenna, female C penis, dorsal view D penis, lateral view E gonocoxae, from Tapan (達邦) F gonocoxae, from Pilu (碧綠) G abdominal ventrite VIII H spermatheca." figureDoi="10.3897/zookeys.944.53099.figure4" httpUri="https://binary.pensoft.net/fig/426847" pageId="0" pageNumber="129">4C, D</figureCitation>
;
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Taiwanoshaira tsoui sp. nov. A antenna, male B antenna, female C penis, dorsal view D penis, lateral view E endophallic sclerites, from Meifeng (梅峰), dorsal view F ditto, lateral view G same, from Peitungyanshan (北東眼山), dorsal H ditto, lateral view I samel, from Tahsuehshan (大雪山), J ditto, laeral view K same, form Taipingshan (太平山), dorsal L ditto, lateral view M gonocoxae, from Hsiaofengkou (小風口) N same, from Taipingshan (太平山) O abdominal ventrite VIII, from Peitungyanshan (北東眼山) P same, from Taipingshan (太平山) Q spermatheca." figureDoi="10.3897/zookeys.944.53099.figure8" httpUri="https://binary.pensoft.net/fig/426851" pageId="0" pageNumber="129">8C, D</figureCitation>
), irregular base of gonocoxae (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7H, I</figureCitation>
) (narrowed base of gonocoxae in others (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Taiwanoshaira chujoi comb. nov. A antenna, male B antenna, female C penis, dorsal view D penis, lateral view E gonocoxae, from Tapan (達邦) F gonocoxae, from Pilu (碧綠) G abdominal ventrite VIII H spermatheca." figureDoi="10.3897/zookeys.944.53099.figure4" httpUri="https://binary.pensoft.net/fig/426847" pageId="0" pageNumber="129">4E, F</figureCitation>
;
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Taiwanoshaira tsoui sp. nov. A antenna, male B antenna, female C penis, dorsal view D penis, lateral view E endophallic sclerites, from Meifeng (梅峰), dorsal view F ditto, lateral view G same, from Peitungyanshan (北東眼山), dorsal H ditto, lateral view I samel, from Tahsuehshan (大雪山), J ditto, laeral view K same, form Taipingshan (太平山), dorsal L ditto, lateral view M gonocoxae, from Hsiaofengkou (小風口) N same, from Taipingshan (太平山) O abdominal ventrite VIII, from Peitungyanshan (北東眼山) P same, from Taipingshan (太平山) Q spermatheca." figureDoi="10.3897/zookeys.944.53099.figure8" httpUri="https://binary.pensoft.net/fig/426851" pageId="0" pageNumber="129">8M, N</figureCitation>
)), and long sclerotized spermathecal duct (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Taiwanoshaira taipingshanensis sp. nov. A antenna, male B antenna, female C penis, apex, dorsal view D penis, base, dorsal view E penis, lateral view F endophallic sclerites, dorsal view G ditto, lateral view H gonocoxae, from Yuanyanghu (鴛鴦湖) I same, from Taipingshan (太平山) J abdominal ventrite VIII, from Yuanyanghu (鴛鴦湖) K same, from Taipingshan (太平山) L spermatheca." figureDoi="10.3897/zookeys.944.53099.figure7" httpUri="https://binary.pensoft.net/fig/426850" pageId="0" pageNumber="129">7L</figureCitation>
) (short sclerotized spermathecal duct in others (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Taiwanoshaira chujoi comb. nov. A antenna, male B antenna, female C penis, dorsal view D penis, lateral view E gonocoxae, from Tapan (達邦) F gonocoxae, from Pilu (碧綠) G abdominal ventrite VIII H spermatheca." figureDoi="10.3897/zookeys.944.53099.figure4" httpUri="https://binary.pensoft.net/fig/426847" pageId="0" pageNumber="129">4H</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Taiwanoshaira tsoui sp. nov. A antenna, male B antenna, female C penis, dorsal view D penis, lateral view E endophallic sclerites, from Meifeng (梅峰), dorsal view F ditto, lateral view G same, from Peitungyanshan (北東眼山), dorsal H ditto, lateral view I samel, from Tahsuehshan (大雪山), J ditto, laeral view K same, form Taipingshan (太平山), dorsal L ditto, lateral view M gonocoxae, from Hsiaofengkou (小風口) N same, from Taipingshan (太平山) O abdominal ventrite VIII, from Peitungyanshan (北東眼山) P same, from Taipingshan (太平山) Q spermatheca." figureDoi="10.3897/zookeys.944.53099.figure8" httpUri="https://binary.pensoft.net/fig/426851" pageId="0" pageNumber="129">8Q</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="host">
<paragraph pageId="0" pageNumber="129">Host plants.</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Lee &amp; Beenen" authorityYear="2020" family="Mniaceae" lsidName="" pageId="0" pageNumber="129" rank="family">Mniaceae</taxonomicName>
:
<taxonomicName class="Bryopsida" family="Mniaceae" genus="Plagiomnium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Plagiomnium vesicatum" order="Bryales" pageId="0" pageNumber="129" phylum="Bryophyta" rank="species" species="vesicatum">
<emphasis italics="true" pageId="0" pageNumber="129">Plagiomnium vesicatum</emphasis>
</taxonomicName>
(Besch.) T.J. Kop. We observed that adults fed on leaves of host plants (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitat photographs. A microhabitat for Taiwanoshaira taipingshanensis sp. nov. and T. tsoui sp. nov. at Yuanyang Lake (鴛鴦湖) B active adults of T. chujoi comb. nov. inside moss cushions at Pilu (畢祿) C adult of T. taipingshanensis sp. nov. feeding on leaves of Plagiomnium vesicatum at Yuanyang Lake (鴛鴦湖) D adults of T. taipingshanensis sp. nov. mating at Yuanyang Lake (鴛鴦湖) E adult of T. tsoui sp. nov. at Hsiaofengkou (小風口) F adult of T. tsoui sp. nov. at Meifeng (梅峰)." figureDoi="10.3897/zookeys.944.53099.figure1" httpUri="https://binary.pensoft.net/fig/426844" pageId="0" pageNumber="129">1C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="etymology">
<paragraph pageId="0" pageNumber="129">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="129">This new species is named for its type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="distribution">
<paragraph pageId="0" pageNumber="129">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="129">
Known from two localities in northern Taiwan (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of Taiwanoshaira species, solid line: 1000 m, broken line: 2000 m. Blue Dots T. chujoi comb. nov. Red Dots T. tsoui sp. nov. Green Dots T. taipingshanensis sp. nov." figureDoi="10.3897/zookeys.944.53099.figure5" httpUri="https://binary.pensoft.net/fig/426848" pageId="0" pageNumber="129">5</figureCitation>
). This new species is sympatric with
<taxonomicName lsidName="T. tsoui" pageId="0" pageNumber="129" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="129">T. tsoui</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
</treatment>
</document>