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<document ID-DOI="http://dx.doi.org/10.3897/jhr.71.34871" ID-GBIF-Dataset="f3eb6a33-72b8-4101-93f7-1e55da67a1d6" ID-Pensoft-Pub="1314-2607-71-209" ID-Pensoft-UUID="A7DBA71EFF7A590CBDC2993C9CBDA3A0" ID-Zenodo-Dep="3401812" ID-ZooBank="C8F1DA4ED17D4BE4850E692815E17D76" ModsDocID="1314-2607-71-209" checkinTime="1567189230149" checkinUser="pensoft" docAuthor="Wang, Hua-Chuan, Chen, Bin &amp; Li, Ting-Jing" docDate="2019" docId="703D1714A8B55F2C88C1D22EB7470B32" docLanguage="en" docName="JourHymenoptRes 71: 209-224" docOrigin="Journal of Hymenoptera Research 71" docPubDate="2019-08-30" docSource="http://dx.doi.org/10.3897/jhr.71.34871" docTitle="Zethus asperipunctatus Wang &amp; Li 2019, sp. nov." docType="treatment" docUuid="5CB359C4-C337-461C-8224-A42D274B3BD5" docUuidSource="ZooBank" docVersion="4" id="A7DBA71EFF7A590CBDC2993C9CBDA3A0" lastPageNumber="215" masterDocId="A7DBA71EFF7A590CBDC2993C9CBDA3A0" masterDocTitle="Three new species of the genus Zethus Fabricius, 1804 (Hymenoptera, Vespidae, Eumeninae) from China, with an updated key to the Oriental species" masterLastPageNumber="224" masterPageNumber="209" pageNumber="212" updateTime="1678753136558" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new species of the genus Zethus Fabricius, 1804 (Hymenoptera, Vespidae, Eumeninae) from China, with an updated key to the Oriental species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Hua-Chuan</mods:namePart>
<mods:affiliation>Chongqing Key Laboratory of Vector Insects, Chongqing Key Laboratory of Animal Biology, Institute of Entomology &amp; Molecular Biology, College of Life Sciences, Chongqing Normal University, Chongqing, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Bin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5227-7736</mods:nameIdentifier>
<mods:affiliation>Chongqing Key Laboratory of Vector Insects, Chongqing Key Laboratory of Animal Biology, Institute of Entomology &amp; Molecular Biology, College of Life Sciences, Chongqing Normal University, Chongqing, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Ting-Jing</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7175-2697</mods:nameIdentifier>
<mods:affiliation>Chongqing Key Laboratory of Vector Insects, Chongqing Key Laboratory of Animal Biology, Institute of Entomology &amp; Molecular Biology, College of Life Sciences, Chongqing Normal University, Chongqing, China</mods:affiliation>
<mods:nameIdentifier type="email">ltjing1979@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-08-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>71</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>209</mods:start>
<mods:end>224</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.71.34871</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.71.34871</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-71-209</mods:identifier>
<mods:identifier type="ZooBank">C8F1DA4ED17D4BE4850E692815E17D76</mods:identifier>
<mods:identifier type="Pensoft-UUID">A7DBA71EFF7A590CBDC2993C9CBDA3A0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3401812</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159326729" LSID="urn:lsid:zoobank.org:act:5CB359C4-C337-461C-8224-A42D274B3BD5" httpUri="http://treatment.plazi.org/id/703D1714A8B55F2C88C1D22EB7470B32" lastPageId="6" lastPageNumber="215" pageId="3" pageNumber="212">
<subSubSection pageId="3" pageNumber="212" type="nomenclature">
<paragraph pageId="3" pageNumber="212">
<taxonomicName ID-CoL="8GB9Y" LSID="http://zoobank.org/5CB359C4-C337-461C-8224-A42D274B3BD5" authority="Wang &amp; Li" authorityName="Wang &amp; Li" authorityYear="2019" family="Zethus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Zethus asperipunctatus" order="Hymenoptera" pageId="3" pageNumber="212" rank="species" species="asperipunctatus" status="sp. nov.">Zethus asperipunctatus Wang &amp; Li</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="212">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="213" type="description">
<paragraph pageId="4" pageNumber="213">
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="4" pageNumber="213">
<pageBreakToken pageId="4" pageNumber="213" start="start">Figs</pageBreakToken>
13-25
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="213" type="materials_examined">
<paragraph pageId="4" pageNumber="213">
<pageBreakToken pageId="4" pageNumber="213" start="start">Material</pageBreakToken>
examined.
</paragraph>
<paragraph pageId="4" pageNumber="213">
<materialsCitation collectingDate="2009-06-07" collectorName="Meicai Wei" country="CHINA" county="Yulong County" latitude="27.38255" location="Hutiaoxia" longLatPrecision="1" longitude="99.88595" municipality="Lijiang City" specimenCount="♀" stateProvince="Yunnan" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
,
<specimenCount>♀</specimenCount>
:
<collectingCountry name="China">CHINA</collectingCountry>
,
<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingMunicipality>Lijiang City</collectingMunicipality>
,
<collectingCounty>Yulong County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:703D1714A8B55F2C88C1D22EB7470B32:EA2199C0020E9A794A257B01C4CA0254" country="CHINA" county="Yulong County" latitude="27.38255" longLatPrecision="1" longitude="99.88595" municipality="Lijiang City" name="Hutiaoxia" stateProvince="Yunnan">Hutiaoxia</location>
,
<geoCoordinate degrees="27" direction="north" minutes="22.953" orientation="latitude" precision="1" value="27.38255">27°22.953'N</geoCoordinate>
,
<geoCoordinate degrees="99" direction="east" minutes="53.157" orientation="longitude" precision="1" value="99.88595">99°53.157'E</geoCoordinate>
,
<collectingDate value="2009-06-07">7.VI.2009</collectingDate>
,
<collectorName>Meicai Wei</collectorName>
leg. (CSUFT).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="215" pageId="4" pageNumber="213" type="description">
<paragraph pageId="4" pageNumber="213">Description.</paragraph>
<paragraph pageId="4" pageNumber="213">
<emphasis bold="true" pageId="4" pageNumber="213">Female</emphasis>
: body length 16.2 mm (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="4" pageNumber="213">Fig. 13</figureCitation>
), fore wing length 13.7 mm. Black, the following parts ferruginous: marking on dorso-inner margin of antennal socket, antennal scape lower area ventrally, pronotum in dorsal view, tegula, dot on lateral of metanotum, apical bands on T1, T2, T3 and middle of S2 (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="4" pageNumber="213">Fig. 13</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="3488104" doi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="4" pageNumber="213" start="Figures 13–25" startId="F2">
<paragraph pageId="4" pageNumber="213">
<emphasis bold="true" pageId="4" pageNumber="213">Figures 13-25.</emphasis>
<taxonomicName authorityName="Wang &amp; Li" authorityYear="2019" family="Zethus" kingdom="Animalia" lsidName="Zethus asperipunctatus" order="Hymenoptera" pageId="4" pageNumber="213" rank="species" species="asperipunctatus">
<emphasis italics="true" pageId="4" pageNumber="213">Zethus asperipunctatus</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="4" pageNumber="213">13</emphasis>
habitus in dorsal view,♀
<emphasis bold="true" pageId="4" pageNumber="213">14</emphasis>
head in frontal view, ♀
<emphasis bold="true" pageId="4" pageNumber="213">15</emphasis>
vertex, ♀
<emphasis bold="true" pageId="4" pageNumber="213">16</emphasis>
clypeus, ♀
<emphasis bold="true" pageId="4" pageNumber="213">17</emphasis>
gena, ♀
<emphasis bold="true" pageId="4" pageNumber="213">18</emphasis>
antenna, ♀
<emphasis bold="true" pageId="4" pageNumber="213">19</emphasis>
mesosoma, ♀
<emphasis bold="true" pageId="4" pageNumber="213">20</emphasis>
propodeum in dorsal view, ♀
<emphasis bold="true" pageId="4" pageNumber="213">21</emphasis>
propodeum in lateral view, ♀
<emphasis bold="true" pageId="4" pageNumber="213">22</emphasis>
metasoma in lateral view, ♀
<emphasis bold="true" pageId="4" pageNumber="213">23</emphasis>
T1, ♀
<emphasis bold="true" pageId="4" pageNumber="213">24</emphasis>
S1, ♀
<emphasis bold="true" pageId="4" pageNumber="213">25</emphasis>
T2, ♀.
</paragraph>
</caption>
<paragraph lastPageId="6" lastPageNumber="215" pageId="4" pageNumber="213">
<emphasis bold="true" italics="true" pageId="4" pageNumber="213">Head.</emphasis>
Head with long setae, their length distinctly longer than 2
<normalizedToken originalValue="×">x</normalizedToken>
posterior ocellar diameter; head about 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as wide as high in frontal view (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="4" pageNumber="213">Fig. 14</figureCitation>
), about 1.7
<normalizedToken originalValue="×">x</normalizedToken>
wider than long in dorsal view (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="4" pageNumber="213">Fig. 15</figureCitation>
); mandible with four blunt teeth and dense setae, its outer surface with coarse punctures; clypeus convex in lateral view and about 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as wide as high in frontal view, with basal margin almost straight, minutely bi-dentate apically, depressed space between teeth with a median carina, width of truncation 1/3
<normalizedToken originalValue="×">x</normalizedToken>
width of clypeus between inner eye margins, clypeus with dense and long setae and dense punctures (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="4" pageNumber="213">Fig. 16</figureCitation>
); frons with long setae and dense punctures; vertex and gena with dense punctures; gena slightly wider than eye in lateral view and without longi
<pageBreakToken pageId="5" pageNumber="214" start="start">tudinal</pageBreakToken>
carina (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="5" pageNumber="214">Fig. 17</figureCitation>
); distance from posterior ocelli to apical margin of vertex 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as distance from posterior ocelli to inner eye margin (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="5" pageNumber="214">Fig. 15</figureCitation>
); occipital carina complete (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="5" pageNumber="214">Fig. 15</figureCitation>
); antennal scape 2.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as its apical width, A3 1.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as
<pageBreakToken pageId="6" pageNumber="215" start="start">its</pageBreakToken>
maximum width, A4 slightly longer than its maximum width, A5-11 wider than long, A12 bullet-shaped, as long as its basal width (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 18</figureCitation>
).
</paragraph>
<paragraph pageId="6" pageNumber="215">
<emphasis bold="true" italics="true" pageId="6" pageNumber="215">Mesosoma.</emphasis>
Length of mesosoma about 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide in dorsal view (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 19</figureCitation>
); pronotal carina complete, pronotum with distinct humeral angle and reticulate punctures dorsally; notaulix complete, mesoscutum with dense punctures, gradually shallow from the anterior to posterior portion; tegula smooth, with dense and short setae on its anterior and posterior lobe; epicnemial carina distinct; mesopleura with deep and dense punctures; scutellum flattened, and with shallow punctures and middle longitudinal furrow almost invisible, distance between punctures about as wide as punctures diameter; punctures on metanotum like those on scutellum, but deeper; metapleuron almost smooth; propodeum dull, with lateral carina and without submedian carina, its dorsolateral surface with reticulate striae (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 20</figureCitation>
), submarginal carina produced into lamella above propodeal valvulae, lateral surface of propodeum with sparse and fine punctures (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 21</figureCitation>
); orifice angled dorsally (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 20</figureCitation>
).
</paragraph>
<paragraph pageId="6" pageNumber="215">
<emphasis bold="true" italics="true" pageId="6" pageNumber="215">Metasoma.</emphasis>
T1 about 2.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, with medial carina from basal margin to one-third of the tergum, gradually widening from one-fourth of the base, then narrowly toward apex, with maximum width about 3.3
<normalizedToken originalValue="×">x</normalizedToken>
its basal width, T1 with coarse and dense punctures (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 23</figureCitation>
) and lateral carina of T1 present wholly (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 24</figureCitation>
); BS of S1 without longitudinal carina and with sparse punctures, lateral portion of AE with weak striae and sporadic punctures (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 24</figureCitation>
); T2 without distinct petiole, gradually swollen from the base to midpoint and nearly straight to apex in lateral view, with developed apical lamella, about 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide in dorsal view, T2 with dense punctures, which gradually sparse from the base to apex, distance between punctures on the base distinctly less than its diameter and 1-2
<normalizedToken originalValue="×">x</normalizedToken>
larger than puncture diameter on apical margin (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 25</figureCitation>
); S2 swollen from the base to near midpoint, subsequent portion nearly straight in profile (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 22</figureCitation>
), punctures on the base of S2 large, punctures on subsequent portion like those on T2, but punctures on lateral portion dense, apical margin of S2 with lamella, T3 and S3 with dense and small punctures, T3 with thick apical lamella, and subsequent terga and sterna with small punctures, distance between punctures about as wide as its diameter.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="215" type="male">
<paragraph pageId="6" pageNumber="215">Male.</paragraph>
<paragraph pageId="6" pageNumber="215">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="215" type="distribution">
<paragraph pageId="6" pageNumber="215">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="215">China: Yunnan.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="215" type="remarks">
<paragraph pageId="6" pageNumber="215">Remarks.</paragraph>
<paragraph pageId="6" pageNumber="215">
This new species resembles
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="Z. nigerrimus" order="Hymenoptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis italics="true" pageId="6" pageNumber="215">Z. nigerrimus</emphasis>
</taxonomicName>
Gusenleitner, 2001 from China, Malaysia, Laos and Vietnam, with which it has the following common characters: clypeus minutely bi-dentate apically, depressed space between teeth with a median ridge (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 16</figureCitation>
), T1 about 2.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 23</figureCitation>
), T2 without distinct petiole (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 25</figureCitation>
), S2 not tuberculate in profile (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 22</figureCitation>
). It differs from
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="Z. nigerrimus" order="Hymenoptera" pageId="6" pageNumber="215" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis italics="true" pageId="6" pageNumber="215">Z. nigerrimus</emphasis>
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and all other members of the genus by the following character combination: body with ferruginous marking (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 13</figureCitation>
), gena without carina (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 17</figureCitation>
); BS of S1 with sparse punctures (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 24</figureCitation>
), S2 blunt angulate in profile (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Fig. 22</figureCitation>
), and propodeum and metasoma dull and with dense setae and punctures (
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">Figs 13</figureCitation>
,
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">21</figureCitation>
,
<figureCitation captionStart="Figures 13–25" captionStartId="F2" captionText="Figures 13 - 25. Zethus asperipunctatus sp. nov., holotype 13 habitus in dorsal view, ♀ 14 head in frontal view, ♀ 15 vertex, ♀ 16 clypeus, ♀ 17 gena, ♀ 18 antenna, ♀ 19 mesosoma, ♀ 20 propodeum in dorsal view, ♀ 21 propodeum in lateral view, ♀ 22 metasoma in lateral view, ♀ 23 T 1, ♀ 24 S 1, ♀ 25 T 2, ♀." figureDoi="10.3897/jhr.71.34871.figures13-25" httpUri="https://binary.pensoft.net/fig/328712" pageId="6" pageNumber="215">22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="215" type="etymology">
<paragraph pageId="6" pageNumber="215">Etymology.</paragraph>
<paragraph pageId="6" pageNumber="215">
The specific name
<taxonomicName authorityName="Wang &amp; Li" authorityYear="2019" family="Zethus" kingdom="Animalia" lsidName="Zethus asperipunctatus" order="Hymenoptera" pageId="6" pageNumber="215" rank="species" species="asperipunctatus">
<emphasis italics="true" pageId="6" pageNumber="215">Zethus asperipunctatus</emphasis>
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is derived from two Latin words:
<emphasis italics="true" pageId="6" pageNumber="215">asper</emphasis>
and
<emphasis italics="true" pageId="6" pageNumber="215">punctatus</emphasis>
, referring to metasoma dull and with dense punctures.
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</treatment>
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Genus
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<paragraph id="F82B36B6FFC2FA5DBFF2FCD5F440FCA4" blockId="5.[143,280,800,823]" box="[143,280,800,823]" pageId="5" pageNumber="358">
<emphasis id="CAE0EAA4FFC2FA5DBFF2FCD5F440FCA4" box="[143,280,800,823]" italics="true" pageId="5" pageNumber="358">
<typeStatus id="272F8814FFC2FA5DBFF2FCD5F59DFCA4" box="[143,197,800,823]" pageId="5" pageNumber="358">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="F82B36B6FFC2FA5DBFF2FCB2F6CFFCCD" blockId="5.[143,919,838,863]" box="[143,919,838,863]" pageId="5" pageNumber="358">
<taxonomicName id="3F944D35FFC2FA5DBFF2FCB2F60DFCCD" authority="Rapp, 1835 in de Ferussac and d'Orbigny, 1834" authorityName="Rapp, 1835 in de Ferussac and d'Orbigny" authorityYear="1834" box="[143,853,838,863]" class="Cephalopoda" family="Octopodidae" genus="Octopus" kingdom="Animalia" order="Octopoda" pageId="5" pageNumber="358" phylum="Mollusca" rank="species" species="indicus">
<emphasis id="CAE0EAA4FFC2FA5DBFF2FCB2F465FCCC" box="[143,317,839,863]" italics="true" pageId="5" pageNumber="358">Octopus indicus</emphasis>
Rapp,
<quantity id="3F6C9B53FFC2FA5DBEF3FCB3F4B8FCCD" box="[398,480,838,862]" metricMagnitude="1" metricUnit="m" metricValue="4.6609" pageId="5" pageNumber="358" unit="in" value="1835.0">1835 in</quantity>
de Ferussac and d’Orbigny, 1834
</taxonomicName>
–1848
</paragraph>
<paragraph id="F82B36B6FFC2FA5DBFF2FC51F5A3FC28" blockId="5.[143,251,932,955]" box="[143,251,932,955]" pageId="5" pageNumber="358">
<emphasis id="CAE0EAA4FFC2FA5DBFF2FC51F5A3FC28" box="[143,251,932,955]" italics="true" pageId="5" pageNumber="358">Diagnosis</emphasis>
</paragraph>
<paragraph id="F82B36B6FFC2FA5DBFF2FC3EF431FBD1" blockId="5.[143,1103,971,1090]" pageId="5" pageNumber="358">
Octopods with biserial arm suckers; eight small mucous pouches, each with a small pore, in the oral surface of the webs with one located between each of the arm bases; hectocotylus of mature males with tiny ligula, calamus absent or barely visible (after
<bibRefCitation id="9C054B47FFC2FA5DBFF2FBDFF405FBD1" author="Liao JX &amp; Lu, CC" box="[143,349,1066,1090]" pageId="5" pageNumber="358" pagination="1 - 10" refId="ref6109" refString="Liao JX, Lu, CC. 2009. A new species of Cistopus (Cephalopoda: Octopodidae) from Taiwan and morphology of mucous pouches. J Mollusc Stud. 75 (3): 1 - 10." type="journal article" year="2009">Liao and Lu 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>