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<mods:title id="9BF1B890AE3DE7B0168AAA91D3E29D5D">Contribution to the Mexican Hyperaspidini: Hyperaspis picta (Gorham, 1899) comb. nov. (Coleoptera: Coccinellidae)</mods:title>
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<taxonomicName id="5AD8EF830E6FFF9502A8F8B6FEFCFF71" authorityName="Szawaryn &amp; Nestor-Arriola" authorityYear="2025" baseAuthorityName="Gorham" baseAuthorityYear="1899" box="[151,353,1811,1837]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="species" species="picta" status="comb. nov.">
<emphasis id="AFAC48120E6FFF9502A8F8B6FEFCFF71" bold="true" box="[151,353,1811,1837]" italics="true" pageId="1" pageNumber="422">Hyperaspis picta</emphasis>
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<figureCitation id="05E388850E6FFF9502A8F892FF65FF0D" box="[151,248,1847,1873]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1982,2006]" captionStartId-1="4.[151,250,1903,1927]" captionStartId-2="5.[151,250,1437,1461]" captionTargetBox-0="[159,1436,591,1947]" captionTargetBox-1="[165,1423,181,1879]" captionTargetBox-2="[151,1435,181,1412]" captionTargetId-0="figure-211@2.[151,1436,591,1959]" captionTargetId-1="figure-39@4.[165,1423,181,1879]" captionTargetId-2="figure-29@5.[151,1435,181,1412]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." captionText-1="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." captionText-2="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi-0="http://doi.org/10.5281/zenodo.14853475" figureDoi-1="http://doi.org/10.5281/zenodo.14853479" figureDoi-2="http://doi.org/10.5281/zenodo.14853481" httpUri-0="https://zenodo.org/record/14853475/files/figure.png" httpUri-1="https://zenodo.org/record/14853479/files/figure.png" httpUri-2="https://zenodo.org/record/14853481/files/figure.png" pageId="1" pageNumber="422">Figs 13</figureCitation>
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<emphasis id="AFAC48120E6FFF9502A8F8DEFEA6FFCF" box="[151,315,1915,1939]" italics="true" pageId="1" pageNumber="422">Cinachyra picta</emphasis>
<bibRefCitation id="F949E9F10E6FFF95037EF8DEFD94FFCF" author="Gorham, H. S." box="[321,521,1915,1939]" pageId="1" pageNumber="422" pagination="257 - 265" refId="ref4171" refString="Gorham, H. S. (1899) Insecta. Coleoptera Vol. VII: Erotylidae, Endomychidae, and Coccinellidae. In: Du Cane Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali - Americana. Published for the editors by R. H. Porter, London, pp. 257-265." type="book chapter" year="1899">Gorham, 1899: 263</bibRefCitation>
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<bibRefCitation id="F949E9F10E6FFF95001EF8DEFC8DFFCF" author="Korschefsky, R." box="[545,784,1915,1939]" pageId="1" pageNumber="422" refId="ref4287" refString="Korschefsky, R. (1931) Pars 118, Coccinellidae. I. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. W. Junk, Berlin, 224 pp." type="book" year="1931">Korschefsky 1931: 109</bibRefCitation>
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;
<treatmentCitation id="1C79B2110E6FFF950125F8DEFB97FFCF" author="Blackwelder, R. E." box="[794,1034,1915,1939]" page="443" pageId="1" pageNumber="422" year="1945">
<bibRefCitation id="F949E9F10E6FFF950125F8DEFB97FFCF" author="Blackwelder, R. E." box="[794,1034,1915,1939]" pageId="1" pageNumber="422" pagination="343 - 550" refId="ref3578" refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 3. Lycidae, Cleridae, Coccinellidae, Tenebrionidae. Bulletin of the United States National Museum, 185 (3), 343-550. https://doi.org/10.5479/si.03629236.185.3" type="journal article" year="1945">Blackwelder 1945: 443</bibRefCitation>
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;
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.
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<emphasis id="AFAC48120E6FFF9502A8F83FFE8CFFED" box="[151,273,1946,1969]" italics="true" pageId="1" pageNumber="422">Semra picta</emphasis>
(
<bibRefCitation id="F949E9F10E6FFF950321F83FFE2BFFEE" author="Gorham, H. S." box="[286,438,1946,1970]" pageId="1" pageNumber="422" pagination="257 - 265" refId="ref4171" refString="Gorham, H. S. (1899) Insecta. Coleoptera Vol. VII: Erotylidae, Endomychidae, and Coccinellidae. In: Du Cane Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali - Americana. Published for the editors by R. H. Porter, London, pp. 257-265." type="book chapter" year="1899">Gorham, 1899</bibRefCitation>
)
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:
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<bibRefCitation id="F949E9F10E6FFF9503F7F83EFD09FFEE" author="Ozdikmen, H." box="[456,660,1946,1972]" pageId="1" pageNumber="422" pagination="25 - 28" refId="ref4769" refString="Ozdikmen, H. (2007) New replacement names for three preoccupied ladybird genera (Coleoptera: Coccinellidae). Munis Entomology &amp; Zoology, 2 (1), 25-28." type="journal article" year="2007">Özdikmen 2007: 25</bibRefCitation>
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;
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<bibRefCitation id="F949E9F10E6FFF9500A1F83FFC21FFEE" author="Nedved, O. &amp; Kovar, I." box="[670,956,1946,1970]" pageId="1" pageNumber="422" pagination="526 - 531" refId="ref4687" refString="Nedved, O. &amp; Kovar, I. (2012) Appendix: List of Genera in Tribes and Subfamilies. In: Hodek, I., van Emden, H. F. &amp; Honek, A. (Eds.), Ecology and Behaviour of the Ladybird Beetles (Coccinellidae). 1 st Edition. Wiley-Blackwell, pp. 526-531. https://doi.org/10.1002/9781118223208.app1" type="book chapter" year="2012">Nedvěd &amp; Kovář 2012: 530</bibRefCitation>
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.
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<emphasis id="AFAC48120E6CFF9602A8FF33FE8FF8EC" bold="true" box="[151,274,150,176]" pageId="2" pageNumber="423">Diagnosis.</emphasis>
<taxonomicName id="5AD8EF830E6CFF960327FF32FE4BF8EC" authorityName="Szawaryn &amp; Nestor-Arriola" authorityYear="2025" baseAuthorityName="Gorham" baseAuthorityYear="1899" box="[280,470,151,176]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="picta" status="comb. nov.">
<emphasis id="AFAC48120E6CFF960327FF32FE4BF8EC" box="[280,470,151,176]" italics="true" pageId="2" pageNumber="423">Hyperaspis picta</emphasis>
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is a very distinctive and unique among all other
<taxonomicName id="5AD8EF830E6CFF960647FF32FACAF8ED" box="[1144,1367,151,177]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="AFAC48120E6CFF960647FF32FB65F8EC" box="[1144,1272,151,176]" italics="true" pageId="2" pageNumber="423">Hyperaspis</emphasis>
species.
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It can be easily distinguished from congeners by its color pattern (black body surface with large, orange, central irregular macula on elytra) and a very distinctive wrinkled surface of elytra.
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<emphasis id="AFAC48120E6CFF9602F8FEA7FE29F940" bold="true" box="[199,436,258,284]" pageId="2" pageNumber="423">Material examined.</emphasis>
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, Gorh.
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[hand writing]/ B.C.A., Col., VII./Type [red circle]/ HOLOTYPE
<taxonomicName id="5AD8EF830E6CFF960634FE82FA01F91D" authority="Gorham 1899: 263" authorityName="Gorham" authorityPageNumber="263" authorityYear="1899" box="[1035,1436,295,321]" class="Demospongiae" family="Tetillidae" genus="Cinachyra" kingdom="Animalia" order="Tetractinellida" pageId="2" pageNumber="423" phylum="Porifera" rank="species" species="picta">
<emphasis id="AFAC48120E6CFF960634FE82FB5CF91C" box="[1035,1217,295,321]" italics="true" pageId="2" pageNumber="423">Cinachyra picta</emphasis>
<bibRefCitation id="F949E9F10E6CFF9606F7FE82FA01F91D" author="Gorham, H. S." box="[1224,1436,295,321]" pageId="2" pageNumber="423" pagination="257 - 265" refId="ref4171" refString="Gorham, H. S. (1899) Insecta. Coleoptera Vol. VII: Erotylidae, Endomychidae, and Coccinellidae. In: Du Cane Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali - Americana. Published for the editors by R. H. Porter, London, pp. 257-265." type="book chapter" year="1899">Gorham 1899: 263</bibRefCitation>
</taxonomicName>
det.
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2005/
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/
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[NHM] (
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)
</materialsCitation>
.
<emphasis id="AFAC48120E6CFF9601FCFEEFFBE2F938" bold="true" box="[963,1151,330,356]" pageId="2" pageNumber="423">Other material.</emphasis>
<materialsCitation id="2DB09E5D0E6CFF9606B8FEEFFBFFF9F1" collectingDate="2021-11-03" collectionCode="CIUM" collectorName="Col. K. Y. Acosta &amp; J. G. Martinez &amp; V. H. Toledo" country="Mexico" elevation="2302" latitude="19.067093" location="Huitzilac" longLatPrecision="1" longitude="-99.26124" municipality="Tres Marias" pageId="2" pageNumber="423" specimenCode="CIUM-cocc-2662" specimenCount="1" specimenCount-female="1" stateProvince="Morelos">
<emphasis id="AFAC48120E6CFF9606B8FEEFFA9BF938" bold="true" box="[1159,1286,330,356]" pageId="2" pageNumber="423">
<collectingCountry id="E5CFD4900E6CFF9606B8FEEFFA9FF938" box="[1159,1282,330,356]" name="Mexico" pageId="2" pageNumber="423">MEXICO</collectingCountry>
:
</emphasis>
<collectingMunicipality id="7D030E7A0E6CFF960730FEEEFA05F939" box="[1295,1432,331,357]" pageId="2" pageNumber="423">Tres Marias</collectingMunicipality>
,
<location id="9807C2DB0E6CFF9602A8FECAFE9CF9D5" LSID="urn:lsid:plazi:treatment:157125160E6FFF9102A8F8B6FB8EFF8D:9807C2DB0E6CFF9602A8FECAFE9CF9D5" box="[151,257,367,393]" country="Mexico" latitude="19.067093" longLatPrecision="1" longitude="-99.26124" municipality="Tres Marias" name="Huitzilac" pageId="2" pageNumber="423" stateProvince="Morelos">Huitzilac</location>
,
<collectingRegion id="5F1C5AE20E6CFF960332FECAFEF0F9D5" box="[269,365,367,393]" country="Mexico" name="Morelos" pageId="2" pageNumber="423">Morelos</collectingRegion>
,
<geoCoordinate id="F8ECF2C70E6CFF960345FECAFD8EF9D5" box="[378,531,367,393]" degrees="19.067093" direction="north" orientation="latitude" pageId="2" pageNumber="423" precision="1" value="19.067093">19.067093°N</geoCoordinate>
,
<geoCoordinate id="F8ECF2C70E6CFF960020FECAFD27F9D5" box="[543,698,367,393]" degrees="99.261239" direction="west" orientation="longitude" pageId="2" pageNumber="423" precision="1" value="-99.26124">99.261239°W</geoCoordinate>
, alt.
<quantity id="5A2039E50E6CFF9600CBFECAFCD4F9D5" box="[756,841,367,393]" metricMagnitude="3" metricUnit="m" metricValue="2.302" pageId="2" pageNumber="423" unit="m" value="2302.0">
<elevation id="16F573330E6CFF9600CBFECAFCD4F9D5" box="[756,841,367,393]" metricMagnitude="3" metricUnit="m" metricValue="2.302" pageId="2" pageNumber="423" unit="m" value="2302.0">2302 m</elevation>
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/ pine forest, on compound flowers,
<date id="E966B2C00E6CFF9606D8FECAFAFDF9D5" box="[1255,1376,367,393]" pageId="2" pageNumber="423" value="2021-11-03">
<collectingDate id="F9224B280E6CFF9606D8FECAFAFDF9D5" box="[1255,1376,367,393]" pageId="2" pageNumber="423" value="2021-11-03">XI-3-2021</collectingDate>
</date>
,
<collectorName id="302DF1D60E6CFF960752FECAFEBBF9F1" pageId="2" pageNumber="423">Col. K. Y. Acosta</collectorName>
,
<collectorName id="302DF1D60E6CFF96030EFE36FE4CF9F1" box="[305,465,403,429]" pageId="2" pageNumber="423">J. G. Martinez</collectorName>
&amp;
<collectorName id="302DF1D60E6CFF9603CAFE36FD18F9F1" box="[501,645,403,429]" pageId="2" pageNumber="423">V. H. Toledo</collectorName>
/
<specimenCode id="CD7E3C7B0E6CFF9600B1FE36FCC4F9F1" box="[654,857,403,429]" pageId="2" pageNumber="423">CIUM-cocc-2662</specimenCode>
/
<specimenCount id="8BDE5F890E6CFF96015EFE36FCF3F9F1" box="[865,878,403,429]" count="1" pageId="2" pageNumber="423" type="female"></specimenCount>
[
<collectionCode id="FBC90CC50E6CFF960144FE36FC57F9F1" box="[891,970,403,429]" pageId="2" pageNumber="423">CIUM</collectionCode>
] (
<figureCitation id="05E388850E6CFF9601DFFE36FBC7F9F1" box="[992,1114,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2006]" captionTargetBox="[159,1436,591,1947]" captionTargetId="figure-211@2.[151,1436,591,1959]" captionTargetPageId="2" captionText="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." figureDoi="http://doi.org/10.5281/zenodo.14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="2" pageNumber="423">Fig. 1D, E</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D5C2C78B0E6CFF9602F8FE13FD39F98D" box="[199,676,438,465]" pageId="2" pageNumber="423" type="distribution">
<paragraph id="9D6794000E6CFF9602F8FE13FD39F98D" blockId="2.[151,1437,150,537]" box="[199,676,438,465]" pageId="2" pageNumber="423">
<emphasis id="AFAC48120E6CFF9602F8FE13FEFCF98C" bold="true" box="[199,353,438,464]" pageId="2" pageNumber="423">Distribution.</emphasis>
<collectingCountry id="E5CFD4900E6CFF960358FE12FE5CF98D" box="[359,449,439,465]" name="Mexico" pageId="2" pageNumber="423">Mexico</collectingCountry>
:
<collectingRegion id="5F1C5AE20E6CFF9603F3FE12FDA9F98D" box="[460,564,439,465]" country="Mexico" name="Guerrero" pageId="2" pageNumber="423">Guerrero</collectingRegion>
,
<collectingRegion id="5F1C5AE20E6CFF960000FE12FD3DF98D" box="[575,672,439,465]" country="Mexico" name="Morelos" pageId="2" pageNumber="423">Morelos</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="D5C2C78B0E6CFF9102F8FE7FFB8EFF8D" lastPageId="5" lastPageNumber="426" pageId="2" pageNumber="423" type="description">
<paragraph id="9D6794000E6CFF9602F8FE7FFC91FA45" blockId="2.[151,1437,150,537]" pageId="2" pageNumber="423">
<emphasis id="AFAC48120E6CFF9602F8FE7FFEE8F9A8" bold="true" box="[199,373,474,500]" pageId="2" pageNumber="423">Redescription.</emphasis>
Length =
<quantity id="5A2039E50E6CFF9603DAFE7FFDD7F9A9" box="[485,586,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="2" pageNumber="423" unit="mm" value="2.45">2.45 mm</quantity>
, width =
<quantity id="5A2039E50E6CFF96008FFE7EFC8BF9A9" box="[688,790,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="2" pageNumber="423" unit="mm" value="1.93">1.93 mm</quantity>
; TL/EW = 1.27; PL/PW = 0.40; EL/EW = 0.94. Body oval (
<figureCitation id="05E388850E6CFF9602A0FE5AFE82FA45" box="[159,287,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2006]" captionTargetBox="[159,1436,591,1947]" captionTargetId="figure-211@2.[151,1436,591,1959]" captionTargetPageId="2" captionText="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." figureDoi="http://doi.org/10.5281/zenodo.14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="2" pageNumber="423">Figs 1A, D</figureCitation>
,
<figureCitation id="05E388850E6CFF960315FE5AFECDFA45" box="[298,336,511,537]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="2" pageNumber="423">2A</figureCitation>
), strongly convex (
<figureCitation id="05E388850E6CFF960018FE5AFD03FA45" box="[551,670,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2006]" captionTargetBox="[159,1436,591,1947]" captionTargetId="figure-211@2.[151,1436,591,1959]" captionTargetPageId="2" captionText="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." figureDoi="http://doi.org/10.5281/zenodo.14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="2" pageNumber="423">Fig. 1C, E</figureCitation>
), winged.
</paragraph>
<caption id="C9A7C4880E6CFF9602A8F81BFBEBFFA7" ID-DOI="http://doi.org/10.5281/zenodo.14853475" ID-Zenodo-Dep="14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="2" pageNumber="423" startId="2.[151,250,1982,2006]" targetBox="[159,1436,591,1947]" targetPageId="2" targetType="figure">
<paragraph id="9D6794000E6CFF9602A8F81BFBEBFFA7" blockId="2.[151,1436,1982,2043]" pageId="2" pageNumber="423">
<emphasis id="AFAC48120E6CFF9602A8F81BFE8EFF8B" bold="true" box="[151,275,1982,2007]" pageId="2" pageNumber="423">FIGURE 1.</emphasis>
<taxonomicName id="5AD8EF830E6CFF960326F81AFDE9FF8B" authority="(Gorham, 1899)" authorityName="Szawaryn &amp; Nestor-Arriola" authorityYear="2025" baseAuthorityName="Gorham" baseAuthorityYear="1899" box="[281,628,1983,2007]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="AFAC48120E6CFF960326F81AFE5BFF8A" box="[281,454,1983,2006]" italics="true" pageId="2" pageNumber="423">Hyperaspis picta</emphasis>
(
<bibRefCitation id="F949E9F10E6CFF9603EAF81AFDF6FF8B" author="Gorham, H. S." box="[469,619,1983,2007]" pageId="2" pageNumber="423" pagination="257 - 265" refId="ref4171" refString="Gorham, H. S. (1899) Insecta. Coleoptera Vol. VII: Erotylidae, Endomychidae, and Coccinellidae. In: Du Cane Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali - Americana. Published for the editors by R. H. Porter, London, pp. 257-265." type="book chapter" year="1899">Gorham, 1899</bibRefCitation>
)
</taxonomicName>
: (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels.
</paragraph>
</caption>
<paragraph id="9D6794000E6DFF9702F8FF32FA9AF9A9" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="424">
Head black with two orange maculae located at base of the antennae, close to anterior margin of clypeus (
<figureCitation id="05E388850E6DFF970750FF32FF25F889" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2006]" captionTargetBox="[159,1436,591,1947]" captionTargetId="figure-211@2.[151,1436,591,1959]" captionTargetPageId="2" captionText="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." figureDoi="http://doi.org/10.5281/zenodo.14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="3" pageNumber="424">Fig. 1B</figureCitation>
) (in female specimen the head is entirely black,
<figureCitation id="05E388850E6DFF9700EDFF1EFCBBF889" box="[722,806,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2006]" captionTargetBox="[159,1436,591,1947]" captionTargetId="figure-211@2.[151,1436,591,1959]" captionTargetPageId="2" captionText="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." figureDoi="http://doi.org/10.5281/zenodo.14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="3" pageNumber="424">Fig. 1E</figureCitation>
); pronotum black with orange-yellow margin of anterior corners and two barely visible small orange maculae on either side of pronotum (
<figureCitation id="05E388850E6DFF970610FF7AFB35F8A5" box="[1071,1192,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2006]" captionTargetBox="[159,1436,591,1947]" captionTargetId="figure-211@2.[151,1436,591,1959]" captionTargetPageId="2" captionText="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." figureDoi="http://doi.org/10.5281/zenodo.14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="3" pageNumber="424">Fig. 1B, C</figureCitation>
) (in female specimen anterior maculae are connected and form irregular ring-like macule in anterior pronotal projections,
<figureCitation id="05E388850E6DFF970739FEA6FAC6F941" box="[1286,1371,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2006]" captionTargetBox="[159,1436,591,1947]" captionTargetId="figure-211@2.[151,1436,591,1959]" captionTargetPageId="2" captionText="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." figureDoi="http://doi.org/10.5281/zenodo.14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="3" pageNumber="424">Fig. 1E</figureCitation>
), two elliptical orange spots in pronotal discal area; elytra black with three pairs of small orange maculae (anterior, mediolateral, and posterior) and a large orange irregular maculae in the central part of elytra (
<figureCitation id="05E388850E6DFF9706BDFEEEFA9CF939" box="[1154,1281,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2006]" captionTargetBox="[159,1436,591,1947]" captionTargetId="figure-211@2.[151,1436,591,1959]" captionTargetPageId="2" captionText="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." figureDoi="http://doi.org/10.5281/zenodo.14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="3" pageNumber="424">Fig. 1A, D</figureCitation>
); ventral side black to dark brown with abdomen brown-orange; legs orange. Head and pronotum with surface rugged, appearing dull, with distinct sculpture (
<figureCitation id="05E388850E6DFF9703DFFE36FDC5F9F1" box="[480,600,403,429]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig 2AD</figureCitation>
); elytra distinctly and densely wrinkled, wrinkles more deeply marked in posterior part of elytra (
<figureCitation id="05E388850E6DFF9703A2FE12FD88F98D" box="[413,533,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2A, E</figureCitation>
). Head sparsely covered with small single punctae (
<figureCitation id="05E388850E6DFF97066EFE12FB35F98D" box="[1105,1192,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2B</figureCitation>
), pronotum and elytra covered with small, shallow double punctae (one bearing minute seta, second without seta) (
<figureCitation id="05E388850E6DFF97069EFE7EFB67F9A9" box="[1185,1274,475,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="9D6794000E6DFF9702F8FE5AFE20FC05" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="424">
Head partially concealed by pronotum; ventral antennal grooves distinct, short and straight (
<figureCitation id="05E388850E6DFF970680FE5AFA89FA45" box="[1215,1300,511,537]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2G</figureCitation>
). Eyes (
<figureCitation id="05E388850E6DFF970750FE5AFF47FA61" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2D, G</figureCitation>
) large, finely faceted, moderately convex, dorsally with inner orbits arcuate; ocular canthus absent; interocular distance about 0.50 times as wide as head width across eyes; interfacetal setae minute, sparsely distributed. Antennal insertions invisible from above (
<figureCitation id="05E388850E6DFF97002BFDCEFDECFAD9" box="[532,625,619,645]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2D</figureCitation>
), covered with lateral extensions of clypeus. Anterior clypeal margin slightly produced, with rounded lateral lobes extending anteriorly, emarginate medially. Antenna short, about 0.44 times as long as width of head capsule, composed of 11 antennomeres (
<figureCitation id="05E388850E6DFF97018AFD16FB96FA91" box="[949,1035,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2G</figureCitation>
); scape enlarged, distinctly roundly projected anteriorly; pedicel narrower than scape, subquadrate, barrel-shaped; antennomere 3 shorter than pedicel, antennomeres 35 elongate, antennomeres 68 trapezoidal, broadening anteriorly, antennomeres 911 forming elongated, fusiform antennal club; antennomere 9 distinctly elongate, much longer than two apical antennomeres combined; antennomere 10 obliquely truncate apically, terminal antennomere short, sub-triangular. Labrum truncate at apex. Mandible apically bidentate. Maxillary cardo transverse with outer margin projecting laterally, reaching slightly outside of mouth cavity (
<figureCitation id="05E388850E6DFF970032FC2EFDFEFBF9" box="[525,611,907,933]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2F</figureCitation>
); maxillary palp short with terminal palpomere slightly widened apically, obliquely truncate at apex. Gula with distinct, central large pore (
<figureCitation id="05E388850E6DFF9701B1FC0AFC7BFB95" box="[910,998,943,969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2F</figureCitation>
). Submentum transverse with anterior margin straight; mentum cordate in shape, anterior margin with distinctly projected lobes, ventral surface with lateral longitudinal carinae; prementum transverse; labial palps separated by distance about equal to width of palpiger; penultimate palpomere large, elongate; ultimate palpomere conical, distinctly narrower and shorter than penultimate one (
<figureCitation id="05E388850E6DFF970365FB9AFE2DFC05" box="[346,432,1087,1113]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="9D6794000E6DFF9702F8FBC6FBCFFD6D" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="424">
Pronotum with anterior corners rounded, slightly projected, not swollen with regular border; anterior margin without bordering line in median part; posterior angle with short, oblique line, posterior margin bordered medially (
<figureCitation id="05E388850E6DFF9702A0FB0EFF65FC99" box="[159,248,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2006]" captionTargetBox="[159,1436,591,1947]" captionTargetId="figure-211@2.[151,1436,591,1959]" captionTargetPageId="2" captionText="FIGURE 1. Hyperaspis picta (Gorham, 1899): (A) male holotype, habitus dorsal, (B) male holotype, habitus frontal, (C) male holotype, habitus lateral, (D) female, habitus dorsolateral, (E) female, frontal, (F) holotype labels." figureDoi="http://doi.org/10.5281/zenodo.14853475" httpUri="https://zenodo.org/record/14853475/files/figure.png" pageId="3" pageNumber="424">Figs 1B</figureCitation>
,
<figureCitation id="05E388850E6DFF97033CFB0EFEB7FC99" box="[259,298,1195,1221]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">2D</figureCitation>
). Prothoracic hypomeron smooth; notosternal suture distinct; prosternal process parallel sided, truncate apically; its surface with short, parallel, lateral carinae (
<figureCitation id="05E388850E6DFF970138FB6AFCC0FCB5" box="[775,861,1231,1257]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2F</figureCitation>
). Prosternum in front of coxa shorter than procoxal longitudinal diameter at the same position; anterior pronotal margin with complete bordering line; procoxal cavity transverse without anterior bordering line, with apically rounded lateral slit (
<figureCitation id="05E388850E6DFF9701CFFAB2FBD8FD6D" box="[1008,1093,1303,1329]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="9D6794000E6DFF9702F8FA9EFB88FE0D" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="424">
Mesoventrite short, transverse, anteriorly emargined with complete raised border; mesoventral process at median width of coxa about as wide as corresponding mesocoxal diameter; meso-metaventral articulation with suture visible; junction straight (
<figureCitation id="05E388850E6DFF970036FA26FD1EFDC1" box="[521,643,1411,1437]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2F, H</figureCitation>
). Scutellar shield sub-triangular (
<figureCitation id="05E388850E6DFF9701C1FA26FBC6FDC1" box="[1022,1115,1411,1437]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2A</figureCitation>
). Elytra with sides rounded; lateral margins narrow, not or barely visible from above; humeral calli present (
<figureCitation id="05E388850E6DFF9701C2FA02FBCBFD9D" box="[1021,1110,1447,1473]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2A</figureCitation>
); elytral epipleuron relatively wide, incomplete apically, inner margin with incomplete bordering line; with foveae for reception of mid and hind femora in repose (
<figureCitation id="05E388850E6DFF970352FA4AFE56FE55" box="[365,459,1519,1545]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2H</figureCitation>
). Metaventrite with discrimen long but incomplete anteriorly; metaventral postcoxal lines joined at middle forming straight line, roundly recurved and complete laterally (
<figureCitation id="05E388850E6DFF97066DF9B6FB31FE71" box="[1106,1196,1555,1581]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2H</figureCitation>
); metaventral surface coarsely punctate, with microreticulation on lateral sides; metepimeron distinct.
</paragraph>
<paragraph id="9D6794000E6DFF9702F8F9FEFAFEFEE1" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="424">
Legs with trochanters angulate (
<figureCitation id="05E388850E6DFF970076F9FEFD5BFE29" box="[585,710,1627,1653]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2H, I</figureCitation>
); pro- and mid femora broad, hind femora slightly swollen; tibiae slender, with distinct apical carina present, tibial apices without spurs (
<figureCitation id="05E388850E6DFF970636F9DAFBC7FEC5" box="[1033,1114,1663,1689]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2I</figureCitation>
); tarsi with four tarsomeres, pseudotrimerous; tarsal claws in female and male with large quadrate basal tooth (
<figureCitation id="05E388850E6DFF97060FF906FB1CFEE1" box="[1072,1153,1699,1725]" captionStart="FIGURE 2" captionStartId="4.[151,250,1903,1927]" captionTargetBox="[165,1423,181,1879]" captionTargetId="figure-39@4.[165,1423,181,1879]" captionTargetPageId="4" captionText="FIGURE 2. Hyperaspis picta (Gorham, 1899) SEM illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw." figureDoi="http://doi.org/10.5281/zenodo.14853479" httpUri="https://zenodo.org/record/14853479/files/figure.png" pageId="3" pageNumber="424">Fig. 2J</figureCitation>
); empodium absent.
</paragraph>
<paragraph id="9D6794000E6DFF9702F8F962FAA9FFE5" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="424">
Abdomen with six ventrites in both sexes (
<figureCitation id="05E388850E6DFF9700A2F962FD68FEBD" box="[669,757,1735,1761]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="3" pageNumber="424">Fig. 3A</figureCitation>
); ventrite 1 long, about as long as ventrites 2 and 3 combined, ventrite 25 sub-equal in length; abdominal postcoxal lines separate medially, incomplete, posteriorly almost reaching hind margin of ventrite 1 and slightly recurved; hind margin of ventrite
<quantity id="5A2039E50E6DFF970609F8ABFBFEFF75" box="[1078,1123,1806,1833]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="424" unit="in" value="5.0">5 in</quantity>
male truncate (
<figureCitation id="05E388850E6DFF97072AF8AAFAECFF75" box="[1301,1393,1807,1833]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="3" pageNumber="424">Fig. 3A</figureCitation>
), in female straight to slightly emarginate; ventrite
<quantity id="5A2039E50E6DFF970093F896FD44FF11" box="[684,729,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="3" pageNumber="424" unit="in" value="6.0">6 in</quantity>
male apically truncate (
<figureCitation id="05E388850E6DFF9701D4F896FBDAFF11" box="[1003,1095,1843,1869]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="3" pageNumber="424">Fig. 3A</figureCitation>
); ventrite
<quantity id="5A2039E50E6DFF970684F896FB7AFF11" box="[1211,1255,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="3" pageNumber="424" unit="in" value="6.0">6 in</quantity>
female apically rounded, in both sexes tergite VIII apically rounded (
<figureCitation id="05E388850E6DFF97013DF8F2FCC0FF2D" box="[770,861,1879,1905]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="3" pageNumber="424">Fig. 3B</figureCitation>
). In both sexes ventrite 6 with lateral pockets for accommodation of laterally bent tergite VIII, lateral margins of tergite VIII partially covers ventrite 6 and is visible ventrally. Small glandular pores between ventrites 23 and 34 present, almost inconspicuous in females.
</paragraph>
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<paragraph id="9D6794000E6AFF9002A8F8CAFCB0FFAF" blockId="4.[151,1437,1903,2035]" pageId="4" pageNumber="425">
<emphasis id="AFAC48120E6AFF9002A8F8CAFE8BFFDB" bold="true" box="[151,278,1903,1927]" pageId="4" pageNumber="425">FIGURE 2.</emphasis>
<taxonomicName id="5AD8EF830E6AFF90031FF8D5FD20FFDB" authority="(Gorham, 1899) SEM" authorityName="SEM" baseAuthorityName="Gorham" baseAuthorityYear="1899" box="[288,701,1903,1927]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="AFAC48120E6AFF90031FF8D5FE4DFFDB" box="[288,464,1904,1927]" italics="true" pageId="4" pageNumber="425">Hyperaspis picta</emphasis>
(
<bibRefCitation id="F949E9F10E6AFF9003DEF8CAFDE5FFDB" author="Gorham, H. S." box="[481,632,1903,1927]" pageId="4" pageNumber="425" pagination="257 - 265" refId="ref4171" refString="Gorham, H. S. (1899) Insecta. Coleoptera Vol. VII: Erotylidae, Endomychidae, and Coccinellidae. In: Du Cane Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali - Americana. Published for the editors by R. H. Porter, London, pp. 257-265." type="book chapter" year="1899">Gorham, 1899</bibRefCitation>
) SEM
</taxonomicName>
illustrations of the holotype: (A) habitus dorsal, (B) magnification of microsculpture and pores on pronotum, (C) magnification of microsculpture and pores on elytra, (D) pronotum and head, (E) apical half of elytra, (F) head ventral, hypomeron, prosternum, mesoventrite, (G) head ventral and antenna, (H) meso- and metaventrite, epipleura and legs, (I) metaleg, (J) mesotarsal claw.
</paragraph>
</caption>
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<paragraph id="9D6794000E6BFF9102A8FA38FB6DFDA0" blockId="5.[151,1436,1437,1533]" pageId="5" pageNumber="426">
<emphasis id="AFAC48120E6BFF9102A8FA38FE8FFDE9" bold="true" box="[151,274,1437,1461]" pageId="5" pageNumber="426">FIGURE 3.</emphasis>
<taxonomicName id="5AD8EF830E6BFF910326FA3BFDF0FDE9" authority="(Gorham, 1899)" authorityName="SEM" baseAuthorityName="Gorham" baseAuthorityYear="1899" box="[281,621,1437,1461]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="AFAC48120E6BFF910326FA3BFE58FDE9" box="[281,453,1438,1461]" italics="true" pageId="5" pageNumber="426">Hyperaspis picta</emphasis>
(
<bibRefCitation id="F949E9F10E6BFF9103ECFA38FDFBFDE9" author="Gorham, H. S." box="[467,614,1437,1461]" pageId="5" pageNumber="426" pagination="257 - 265" refId="ref4171" refString="Gorham, H. S. (1899) Insecta. Coleoptera Vol. VII: Erotylidae, Endomychidae, and Coccinellidae. In: Du Cane Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali - Americana. Published for the editors by R. H. Porter, London, pp. 257-265." type="book chapter" year="1899">Gorham, 1899</bibRefCitation>
)
</taxonomicName>
details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X.
</paragraph>
</caption>
<paragraph id="9D6794000E6BFF9102F8F9EAFBA2FF41" blockId="5.[151,1437,1615,2002]" pageId="5" pageNumber="426">
Male terminalia and genitalia. Sternite IX (
<figureCitation id="05E388850E6BFF910080F9EAFC8DFE35" box="[703,784,1615,1641]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="5" pageNumber="426">Fig. 3J</figureCitation>
) with additional sclerite at base of apodeme in form of inverted V; apodeme narrow, simple, rod-like; tergite X short, transverse. Tegmen in inner view (
<figureCitation id="05E388850E6BFF9106FAF9D6FA87FED1" box="[1221,1306,1651,1677]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="5" pageNumber="426">Fig. 3G</figureCitation>
) with penis guide asymmetrical, petal shaped; parameres (
<figureCitation id="05E388850E6BFF910098F932FCB8FEED" box="[679,805,1687,1714]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="5" pageNumber="426">Fig. 3G, H</figureCitation>
) articulated with phallobase, well developed, distinctly longer than penis guide, asymmetric, with fringe of dense, long setae on apices and outer margins; tegminal strut rod-like. Penis basal capsule with inner arm well developed, outer arm slightly reduced; penis of equal diameter along the entire length; penis tip lightly sclerotized, partially membranous (
<figureCitation id="05E388850E6BFF9101DBF8A6FBAFFF41" box="[996,1074,1795,1821]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="5" pageNumber="426">Fig. 3I</figureCitation>
).
</paragraph>
<paragraph id="9D6794000E6BFF9102F8F882FB8EFF8D" blockId="5.[151,1437,1615,2002]" pageId="5" pageNumber="426">
Female genitalia. Sperm duct longer than spermatheca; infundibulum absent; spermatheca compound, basal unit elongate (
<figureCitation id="05E388850E6BFF910303F8EEFE0FFF39" box="[316,402,1867,1893]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="5" pageNumber="426">Fig. 3F</figureCitation>
) and apical portion spherical (
<figureCitation id="05E388850E6BFF9100CEF8EEFCD5FF39" box="[753,840,1867,1893]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="5" pageNumber="426">Fig. 3E</figureCitation>
) connected by narrow duct, spermathecal accessory gland present (
<figureCitation id="05E388850E6BFF910303F8CAFE0EFFD5" box="[316,403,1903,1929]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="5" pageNumber="426">Fig. 3C</figureCitation>
). Coxites sub-quadrate, deeply excavated at base, truncate on apices; apical margins covered with long setae (
<figureCitation id="05E388850E6BFF91036FF836FE35FFF1" box="[336,424,1939,1965]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="5" pageNumber="426">Fig. 3C</figureCitation>
); styli present. Proctiger large, transverse, with anterior margin emarginated, and posterior margin almost straight, slightly emarginated, covered with long setae (
<figureCitation id="05E388850E6BFF910193F812FB9BFF8D" box="[940,1030,1975,2001]" captionStart="FIGURE 3" captionStartId="5.[151,250,1437,1461]" captionTargetBox="[151,1435,181,1412]" captionTargetId="figure-29@5.[151,1435,181,1412]" captionTargetPageId="5" captionText="FIGURE 3. Hyperaspis picta (Gorham, 1899) details of abdomen and genitalia: (A) abdomen, male, (B) tergite VIII, male, (C) female terminalia and genitalia, ventral, (D) female terminalia, dorsal, (E) spherical part of spermatheca, (F) elongate part of spermatheca, (G) tegmen, inner, (H) tegmen, lateral, (I) penis, lateral, (J) male abdominal segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.14853481" httpUri="https://zenodo.org/record/14853481/files/figure.png" pageId="5" pageNumber="426">Fig. 3D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,171 @@
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<heading id="C62F236C0E6FFF9502A8FB6BFD5EFCB4" bold="true" box="[151,707,1230,1257]" fontSize="11" level="1" pageId="1" pageNumber="422" reason="1">
<emphasis id="AFAC48120E6FFF9502A8FB6BFD5EFCB4" bold="true" box="[151,707,1230,1257]" pageId="1" pageNumber="422">
Genus
<taxonomicName id="5AD8EF830E6FFF9502D1FB6AFE67FCB4" ID-CoL="62V33" authority="Chevrolat" authorityName="Chevrolat" authorityYear="1836" box="[238,506,1230,1257]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="AFAC48120E6FFF9502D1FB6AFEE8FCB5" bold="true" box="[238,373,1231,1257]" italics="true" pageId="1" pageNumber="422">Hyperaspis</emphasis>
Chevrolat
</taxonomicName>
in Dejean, 1836
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<subSubSection id="D5C2C78B0E6FFF9502A8FAB1FDFFFD77" box="[151,610,1299,1323]" pageId="1" pageNumber="422" type="reference_group">
<paragraph id="9D6794000E6FFF9502A8FAB1FDFFFD77" blockId="1.[151,1437,1299,1479]" box="[151,610,1299,1323]" pageId="1" pageNumber="422">
<treatmentCitationGroup id="BDC8B32E0E6FFF9502A8FAB1FDFFFD77" box="[151,610,1299,1323]" pageId="1" pageNumber="422">
<taxonomicName id="5AD8EF830E6FFF9502A8FAB1FEEAFD77" ID-CoL="62V33" authority="Chevrolat" authorityName="Chevrolat" authorityYear="1836" box="[151,375,1299,1323]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="AFAC48120E6FFF9502A8FAB1FE97FD77" box="[151,266,1300,1323]" italics="true" pageId="1" pageNumber="422">Hyperaspis</emphasis>
Chevrolat
</taxonomicName>
in Dejean, 1836: 435.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D5C2C78B0E6FFF950056FAB1FE07FD16" pageId="1" pageNumber="422" type="type_taxon">
<paragraph id="9D6794000E6FFF950056FAB1FE07FD16" blockId="1.[151,1437,1299,1479]" pageId="1" pageNumber="422">
<typeStatus id="42632AA20E6FFF950056FAB1FD06FD70" box="[617,667,1300,1324]" pageId="1" pageNumber="422">Type</typeStatus>
species:
<taxonomicName id="5AD8EF830E6FFF9500C3FAB6FBFFFD77" authority="Herbst, 1783" authorityName="Herbst" authorityYear="1783" box="[764,1122,1299,1323]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="species" species="reppensis">
<emphasis id="AFAC48120E6FFF9500C3FAB6FC4FFD77" box="[764,978,1299,1323]" italics="true" pageId="1" pageNumber="422">Coccinella reppensis</emphasis>
<bibRefCitation id="F949E9F10E6FFF9501E4FAB6FBFFFD77" author="Herbst, J. F. W." box="[987,1122,1299,1323]" pageId="1" pageNumber="422" pagination="1 - 68" refId="ref4236" refString="Herbst, J. F. W. (1783) Kritishes Verzeichniss meiner Insecten-Sammlung. Vol. 3. In: Fuessly, J. C. (Ed.), Archiv der Insectengeschichte. H. Steiner, Zurich und Winterthur, pp. 1-68." type="book chapter" year="1783">Herbst, 1783</bibRefCitation>
</taxonomicName>
, by subsequent designation of
<treatmentCitation id="1C79B2110E6FFF9502F8FA97FE0BFD16" author="Thomson, C. G." box="[199,406,1330,1354]" page="161" pageId="1" pageNumber="422" year="1859">
<bibRefCitation id="F949E9F10E6FFF9502F8FA97FE0BFD16" author="Thomson, C. G." box="[199,406,1330,1354]" pageId="1" pageNumber="422" refId="ref5259" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom. I. Berlingska Boktryckeriet, Lund, 290 pp. https://doi.org/10.5962/bhl.title.138677" type="book" year="1859">Thomson 1859: 161</bibRefCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D5C2C78B0E6FFF9502A8FAF5FB2EFD98" pageId="1" pageNumber="422" type="reference_group">
<paragraph id="9D6794000E6FFF9502A8FAF5FEBAFDDB" blockId="1.[151,1437,1299,1479]" pageId="1" pageNumber="422">
<treatmentCitationGroup id="BDC8B32E0E6FFF9502A8FAF5FE46FD35" box="[151,475,1360,1385]" pageId="1" pageNumber="422">
<treatmentCitation id="1C79B2110E6FFF9502A8FAF5FE4AFD35" author="LeConte, J. L." box="[151,471,1360,1385]" page="238" pageId="1" pageNumber="422" year="1850">
<taxonomicName id="5AD8EF830E6FFF9502A8FAF5FE4AFD35" authority="LeConte, 1850: 238" authorityName="LeConte" authorityPageNumber="238" authorityYear="1850" box="[151,471,1360,1385]" class="Insecta" family="Coccinellidae" genus="Oxynychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="AFAC48120E6FFF9502A8FAF5FE9BFD34" box="[151,262,1360,1384]" italics="true" pageId="1" pageNumber="422">Oxynychus</emphasis>
<bibRefCitation id="F949E9F10E6FFF950333FAF4FE4AFD35" author="LeConte, J. L." box="[268,471,1360,1385]" pageId="1" pageNumber="422" pagination="201 - 242" refId="ref4405" refString="LeConte, J. L. (1850) General remarks upon the Coleoptera of Lake Superior. In: Agassiz, L. (Ed.), Lake Superior: its character, vegetation, and animals, compared with those of other similar regions. Gould, Kendall &amp; Lincoln, Boston, pp. 201-242." type="book chapter" year="1850">LeConte, 1850: 238</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
<treatmentCitationGroup id="BDC8B32E0E6FFF9503DFFAF4FEBAFDDB" pageId="1" pageNumber="422">
<typeStatus id="42632AA20E6FFF9503DFFAF4FD8FFD34" box="[480,530,1361,1384]" pageId="1" pageNumber="422">Type</typeStatus>
species:
<taxonomicName id="5AD8EF830E6FFF950052FAF5FC49FD34" authority="LeConte, 1850" authorityName="LeConte" authorityYear="1850" box="[621,980,1360,1385]" class="Insecta" family="Coccinellidae" genus="Oxynychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="species" species="moerens">
<emphasis id="AFAC48120E6FFF950052FAF5FCAAFD34" box="[621,823,1360,1384]" italics="true" pageId="1" pageNumber="422">Oxynychus moerens</emphasis>
<bibRefCitation id="F949E9F10E6FFF950103FAF4FC49FD34" author="LeConte, J. L." box="[828,980,1360,1385]" pageId="1" pageNumber="422" pagination="201 - 242" refId="ref4405" refString="LeConte, J. L. (1850) General remarks upon the Coleoptera of Lake Superior. In: Agassiz, L. (Ed.), Lake Superior: its character, vegetation, and animals, compared with those of other similar regions. Gould, Kendall &amp; Lincoln, Boston, pp. 201-242." type="book chapter" year="1850">LeConte, 1850</bibRefCitation>
</taxonomicName>
, by monotypy. Synonymized by
<treatmentCitation id="1C79B2110E6FFF950721FAF5FEBEFDDB" author="Dobzhansky, T. G." page="78" pageId="1" pageNumber="422" year="1941">
<bibRefCitation id="F949E9F10E6FFF950721FAF5FEBEFDDB" author="Dobzhansky, T. G." pageId="1" pageNumber="422" pagination="1 - 94" refId="ref3940" refString="Dobzhansky, T. G. (1941) Beetles of the genus Hyperaspis inhabiting the United States. Smithsonian Miscellaneous Collections, 101 (6), 1-94." type="journal article" year="1941">Dobzhansky 1941: 78</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9D6794000E6FFF9502A8FA2BFB7FFDF9" blockId="1.[151,1437,1299,1479]" box="[151,1250,1422,1446]" pageId="1" pageNumber="422">
<treatmentCitationGroup id="BDC8B32E0E6FFF9502A8FA2BFB7FFDF9" box="[151,1250,1422,1446]" pageId="1" pageNumber="422">
<treatmentCitation id="1C79B2110E6FFF9502A8FA2BFE4CFDFA" author="Gorham, H. S." box="[151,465,1422,1446]" page="263" pageId="1" pageNumber="422" year="1899">
<taxonomicName id="5AD8EF830E6FFF9502A8FA2BFE4CFDFA" ID-CoL="84MD7" authority="Gorham, 1899: 263" authorityName="Gorham" authorityPageNumber="263" authorityYear="1899" box="[151,465,1422,1446]" class="Demospongiae" family="Tetillidae" genus="Cinachyra" kingdom="Animalia" order="Tetractinellida" pageId="1" pageNumber="422" phylum="Porifera" rank="genus">
<emphasis id="AFAC48120E6FFF9502A8FA2BFE9FFDFA" box="[151,258,1422,1446]" italics="true" pageId="1" pageNumber="422">Cinachyra</emphasis>
<bibRefCitation id="F949E9F10E6FFF950337FA2BFE4CFDFA" author="Gorham, H. S." box="[264,465,1422,1446]" pageId="1" pageNumber="422" pagination="257 - 265" refId="ref4171" refString="Gorham, H. S. (1899) Insecta. Coleoptera Vol. VII: Erotylidae, Endomychidae, and Coccinellidae. In: Du Cane Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali - Americana. Published for the editors by R. H. Porter, London, pp. 257-265." type="book chapter" year="1899">Gorham, 1899: 263</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="42632AA20E6FFF9503E5FA2BFD91FDFA" box="[474,524,1422,1446]" pageId="1" pageNumber="422">Type</typeStatus>
species:
<taxonomicName id="5AD8EF830E6FFF950055FA2BFC35FDFA" authority="Gorham, 1899" authorityName="Gorham" authorityYear="1899" box="[618,936,1422,1446]" class="Demospongiae" family="Tetillidae" genus="Cinachyra" kingdom="Animalia" order="Tetractinellida" pageId="1" pageNumber="422" phylum="Porifera" rank="species" species="picta">
<emphasis id="AFAC48120E6FFF950055FA2BFC93FDF9" box="[618,782,1422,1446]" italics="true" pageId="1" pageNumber="422">Cinachyra picta</emphasis>
<bibRefCitation id="F949E9F10E6FFF95012BFA2BFC35FDFA" author="Gorham, H. S." box="[788,936,1422,1446]" pageId="1" pageNumber="422" pagination="257 - 265" refId="ref4171" refString="Gorham, H. S. (1899) Insecta. Coleoptera Vol. VII: Erotylidae, Endomychidae, and Coccinellidae. In: Du Cane Godman, F. &amp; Salvin, O. (Eds.), Biologia Centrali - Americana. Published for the editors by R. H. Porter, London, pp. 257-265." type="book chapter" year="1899">Gorham, 1899</bibRefCitation>
</taxonomicName>
, by monotypy.
<emphasis id="AFAC48120E6FFF95067BFA2BFB46FDFA" bold="true" box="[1092,1243,1422,1446]" pageId="1" pageNumber="422">New synonym</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9D6794000E6FFF9502A8FA08FB2EFD98" blockId="1.[151,1437,1299,1479]" box="[151,1203,1452,1479]" pageId="1" pageNumber="422">
<treatmentCitationGroup id="BDC8B32E0E6FFF9502A8FA08FB2EFD98" box="[151,1203,1452,1479]" pageId="1" pageNumber="422">
<treatmentCitation id="1C79B2110E6FFF9502A8FA08FE2DFD98" author="Ozdikmen, H." box="[151,432,1452,1479]" page="26" pageId="1" pageNumber="422" year="2007">
<taxonomicName id="5AD8EF830E6FFF9502A8FA08FE2DFD98" ID-CoL="CGDF7" authority="Ozdikmen, 2007: 26" authorityName="Ozdikmen" authorityPageNumber="26" authorityYear="2007" box="[151,432,1452,1479]" class="Insecta" family="Coccinellidae" genus="Semra" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="AFAC48120E6FFF9502A8FA08FF45FD98" box="[151,216,1453,1476]" italics="true" pageId="1" pageNumber="422">Semra</emphasis>
<bibRefCitation id="F949E9F10E6FFF9502E1FA0BFE2DFD98" author="Ozdikmen, H." box="[222,432,1452,1479]" pageId="1" pageNumber="422" pagination="25 - 28" refId="ref4769" refString="Ozdikmen, H. (2007) New replacement names for three preoccupied ladybird genera (Coleoptera: Coccinellidae). Munis Entomology &amp; Zoology, 2 (1), 25-28." type="journal article" year="2007">Özdikmen, 2007: 26</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Replacement name nec.
<taxonomicName id="5AD8EF830E6FFF95008DFA09FC3CFD98" ID-CoL="84MD7" authority="Sollas, 1866" authorityName="Sollas" authorityYear="1866" box="[690,929,1452,1476]" class="Demospongiae" family="Tetillidae" genus="Cinachyra" kingdom="Animalia" order="Tetractinellida" pageId="1" pageNumber="422" phylum="Porifera" rank="genus">
<emphasis id="AFAC48120E6FFF95008DFA09FC80FD98" box="[690,797,1452,1476]" italics="true" pageId="1" pageNumber="422">Cinachyra</emphasis>
Sollas, 1866
</taxonomicName>
(Porifera).
<emphasis id="AFAC48120E6FFF950629FA08FB30FD99" bold="true" box="[1046,1197,1453,1477]" pageId="1" pageNumber="422">New synonym</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D5C2C78B0E6FFF9502A8FA57FE9BFE9D" pageId="1" pageNumber="422" type="diagnosis">
<paragraph id="9D6794000E6FFF9502A8FA57FE9BFE9D" blockId="1.[151,1436,1522,1729]" pageId="1" pageNumber="422">
<emphasis id="AFAC48120E6FFF9502A8FA57FE8FFE50" bold="true" box="[151,274,1522,1548]" pageId="1" pageNumber="422">Diagnosis.</emphasis>
Body surface of
<taxonomicName id="5AD8EF830E6FFF9503E1FA56FDC3FE50" authorityName="Chevrolat" authorityYear="1836" box="[478,606,1523,1548]" class="Insecta" family="Coccinellidae" genus="Hyperaspis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="AFAC48120E6FFF9503E1FA56FDC3FE50" box="[478,606,1523,1548]" italics="true" pageId="1" pageNumber="422">Hyperaspis</emphasis>
</taxonomicName>
appearing glabrous; antennae composed of 10 or 11 antennomeres; eye canthus absent; prosternal process usually with a pair of parallel carinae; protibiae usually slender, rarely flanged; elytral epipleuron excavated for reception of mid and hind femoral apices; postcoxal lines on the ventrite 1 incomplete; posterior margin of apical ventrite in males usually modified; penis guide asymmetric; parameres of different length; coxites in females sub-rectangular, basally excavated; spermatheca complex, usually composed of two parts.
</paragraph>
</subSubSection>
</treatment>
</document>