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<document id="F0D51887CFDF55468DB5CAF71B45CA43" ID-DOI="10.1093/zoolinnean/zlad123" ID-ISSN="0024-4082" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1722843859434" checkinUser="plazi" docAuthor="Lobón-Rovira, Javier, Bauer, Aaron M., Pinto, Pedro Vaz, Trape, Jean-Francois, Conradie, Werner, Kusamba, Chifundera, Júlio, Timóteo, Cael, Garin, Stanley, Edward L., Hughes, Daniel F., Behangana, Mathias, Masudi, Franck M., Pauwels, Olivier S. G. &amp; Greenbaum, Eli" docDate="2024" docId="03B30955FFC0FFE4FF7147FBFDD3F87F" docLanguage="en" docName="zlad123.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 201 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad123" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.6:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="6" docTitle="Lygodactylus depressus , Schmidt 1919" docType="treatment" docVersion="1" lastPageNumber="474" masterDocId="FF8A712DFFD7FFFFFFC2407AFF8CFFB4" masterDocTitle="Integrative revision of the Lygodactylus gutturalis (Bocage, 1873) complex unveils extensive cryptic diversity and traces its evolutionary history" masterLastPageNumber="492" masterPageNumber="447" pageNumber="470" updateTime="1723452314630" updateUser="tatiana">
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<treatment id="03B30955FFC0FFE4FF7147FBFDD3F87F" LSID="urn:lsid:plazi:treatment:03B30955FFC0FFE4FF7147FBFDD3F87F" httpUri="http://treatment.plazi.org/id/03B30955FFC0FFE4FF7147FBFDD3F87F" lastPageId="27" lastPageNumber="474" pageId="23" pageNumber="470">
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<paragraph id="8BA5B843FFC0FFE8FF7147FBFD35F828" blockId="23.[113,697,1921,1984]" box="[179,697,1921,1948]" pageId="23" pageNumber="470">
<taxonomicName id="4C1AC3C0FFC0FFE8FF7147FBFD35F828" ID-CoL="3WQ3G" authority="Schmidt, 1919" authorityName=", Schmidt" authorityYear="1919" box="[179,697,1921,1948]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFC0FFE8FF7147FBFD35F828" box="[179,697,1921,1948]" italics="true" pageId="23" pageNumber="470">
<emphasis id="B96E6451FFC0FFE8FF7147FBFDA4F82F" bold="true" box="[179,552,1921,1947]" italics="true" pageId="23" pageNumber="470">LYGODACTYLUS DEPRESSUS</emphasis>
<bibRefCitation id="EF8BC5B2FFC0FFE8FDEF47FBFD35F828" author="Schmidt KP" box="[557,697,1921,1948]" pageId="23" pageNumber="470" pagination="385 - 624" refId="ref37222" refString="Schmidt KP. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909 - 1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bulletin of the American Museum of Natural History 1919; 39: 385 - 624." type="journal article" year="1919">Schmidt, 1919</bibRefCitation>
</emphasis>
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<paragraph id="8BA5B843FFC0FFE8FFB347D2FD3EF874" blockId="23.[113,697,1921,1984]" box="[113,690,1960,1984]" pageId="23" pageNumber="470">
(
<figureCitation id="1321A4C6FFC0FFE8FFBE47D2FF61F874" box="[124,237,1960,1984]" captionStart-0="Figure 13" captionStart-1="Figure 14" captionStart-2="Figure 15" captionStart-3="Figure 16" captionStartId-0="24.[129,194,1924,1948]" captionStartId-1="25.[113,178,1107,1131]" captionStartId-2="26.[129,194,1836,1860]" captionStartId-3="27.[112,177,961,985]" captionTargetBox-0="[132,1470,1394,1896]" captionTargetBox-1="[115,1458,147,1078]" captionTargetBox-2="[198,1405,144,1808]" captionTargetBox-3="[124,1455,157,932]" captionTargetId-0="figure-744@24.[129,1473,1388,1897]" captionTargetId-1="figure-474@25.[113,1460,144,1079]" captionTargetId-2="figure-10@26.[198,1405,144,1808]" captionTargetId-3="figure-520@27.[114,1458,144,933]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="27" captionText-0="Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC." captionText-1="Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH)." captionText-2="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." captionText-3="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="23" pageNumber="470">Figs 13–16</figureCitation>
,
<tableCitation id="C6988DF8FFC0FFE8FF3547D2FECEF874" box="[247,322,1960,1984]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="23" pageNumber="470">Table 3</tableCitation>
; Supporting Information, Table S6)
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFC0FFE8FCA14498FAA7FA3B" pageId="23" pageNumber="470" type="reference_group">
<paragraph id="8BA5B843FFC0FFE8FCA14498FB48FAAF" blockId="23.[867,1402,1250,1307]" pageId="23" pageNumber="470">
<emphasis id="B96E6451FFC0FFE8FCA14498FB48FAAF" italics="true" pageId="23" pageNumber="470">
<taxonomicName id="4C1AC3C0FFC0FFE8FCA14498FB4EFB48" ID-CoL="3WQ3G" authority="(Schmidt, 1919)" box="[867,1218,1250,1276]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="subSpecies" species="picturatus" subSpecies="depressus">Lygodactylus picturatus depressus</taxonomicName>
:
<bibRefCitation id="EF8BC5B2FFC0FFE8FB0F4498FAFAFB48" author="Loveridge A" box="[1229,1398,1250,1276]" pageId="23" pageNumber="470" pagination="1 - 469" refId="ref35578" refString="Loveridge A. Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology 1947; 98: 1 - 469." type="journal article" year="1947">Loveridge (1947)</bibRefCitation>
,
<bibRefCitation id="EF8BC5B2FFC0FFE8FBDB457BFB4CFAAF" author="Wermuth H" box="[1049,1216,1281,1307]" pageId="23" pageNumber="470" pagination="1 - 246" refId="ref37733" refString="Wermuth H. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Das Tierreich 1965; 80: I - XXIII, 1 - 246." type="journal article" year="1965">Wermuth (1965)</bibRefCitation>
.
</emphasis>
</paragraph>
<paragraph id="8BA5B843FFC0FFE8FC85454CFAA7FA3B" blockId="23.[810,1460,1334,1991]" pageId="23" pageNumber="470">
<emphasis id="B96E6451FFC0FFE8FC85454CFAA7FA3B" italics="true" pageId="23" pageNumber="470">
<taxonomicName id="4C1AC3C0FFC0FFE8FC85454CFBBAFAE4" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[839,1078,1334,1360]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="depressus">Lygodactylus depressus</taxonomicName>
: Pasteur, 1965(1964);
<bibRefCitation id="EF8BC5B2FFC0FFE8FAD5454CFA1EFAE4" author="Kluge AG" box="[1303,1426,1334,1360]" pageId="23" pageNumber="470" pagination="1 - 35" refId="ref35045" refString="Kluge AG. Checklist of gekkonid lizards. Smithsonian Herpetological Information Service 1991; 85: 1 - 35. https: // doi. org / 10.5479 / si. 23317515.85.1" type="journal article" year="1991">Kluge (1991</bibRefCitation>
,
<bibRefCitation id="EF8BC5B2FFC0FFE8FCF7452FFCE6FADB" author="Kluge AG" box="[821,874,1365,1391]" pageId="23" pageNumber="470" refId="ref35077" refString="Kluge AG. Gekkonoid Lizard Taxonomy. San Diego: International Gecko Society, 1993." type="book" year="1993">1993</bibRefCitation>
,
<bibRefCitation id="EF8BC5B2FFC0FFE8FCB0452FFC2BFADB" author="Kluge AG" box="[882,935,1365,1391]" pageId="23" pageNumber="470" pagination="1 - 209" refId="ref35093" refString="Kluge AG. Gekkotan lizard taxonomy. Hamadryad 2001; 26: 1 - 209." type="journal article" year="2001">2001</bibRefCitation>
);
<bibRefCitation id="EF8BC5B2FFC0FFE8FC7E452FFBC9FADB" author="Rosler H" box="[956,1093,1365,1391]" pageId="23" pageNumber="470" pagination="28 - 153" refId="ref36972" refString="Rosler H. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2000; 2: 28 - 153." type="journal article" year="2000">Rösler (2000)</bibRefCitation>
; Chirio and LeBreton (2007);
<bibRefCitation id="EF8BC5B2FFC0FFE8FAA9452FFB95FA3B" author="Roll B &amp; Prohl H &amp; Hoffmann KP" pageId="23" pageNumber="470" pagination="327 - 35" refId="ref36822" refString="Roll B, Prohl H, Hoffmann KP. Multigene phylogenetic analysis of Lygodactylus dwarf geckos (Squamata: Gekkonidae). Molecular Phylogenetics and Evolution 2010; 56: 327 - 35. https: // doi. org / 10.1016 / j. ympev. 2010.02.002" type="journal article" year="2010">Röll et al. (2010)</bibRefCitation>
; de Lisle et al. (2013, 2016)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFC0FFE7FCE845E1FCF5FD6F" lastPageId="24" lastPageNumber="471" pageId="23" pageNumber="470" type="discussion">
<paragraph id="8BA5B843FFC0FFE8FCE845E1FB90F873" blockId="23.[810,1460,1334,1991]" pageId="23" pageNumber="470">
<taxonomicName id="4C1AC3C0FFC0FFE8FCE845E1FB82FA07" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[810,1038,1435,1459]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFC0FFE8FCE845E1FB82FA07" box="[810,1038,1435,1459]" italics="true" pageId="23" pageNumber="470">Lygodactylus depressus</emphasis>
</taxonomicName>
was described from Medje, Haut-Uele Province,
<collectingCountry id="F30DF8D3FFC0FFE8FC5145C0FC47FA66" box="[915,971,1466,1490]" name="Democratic Republic of the Congo" pageId="23" pageNumber="470">DRC</collectingCountry>
, by Schmidt, who described the species as an intermediate form between
<taxonomicName id="4C1AC3C0FFC0FFE8FB9345A0FB43FA45" ID-CoL="3WQ4R" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[1105,1231,1498,1521]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFC0FFE8FB9345A0FB43FA45" box="[1105,1231,1498,1521]" italics="true" pageId="23" pageNumber="470">L. picturatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1AC3C0FFC0FFE8FACA45A0FA0DFA46" ID-CoL="3WQ3T" box="[1288,1409,1498,1522]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC0FFE8FACA45A0FA0DFA46" box="[1288,1409,1498,1522]" italics="true" pageId="23" pageNumber="470">L. gutturalis</emphasis>
</taxonomicName>
: ‘
<emphasis id="B96E6451FFC0FFE8FA5A45A0FA3FFA45" box="[1432,1459,1498,1521]" italics="true" pageId="23" pageNumber="470">…</emphasis>
The relation of this species with
<taxonomicName id="4C1AC3C0FFC0FFE8FBB44580FA38F9A5" ID-CoL="7HDHF" authority="(Peters)" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[1142,1460,1529,1553]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="subSpecies" species="picturatus" subSpecies="picturatus">
<emphasis id="B96E6451FFC0FFE8FBB44580FAD5F9A5" box="[1142,1369,1530,1553]" italics="true" pageId="23" pageNumber="470">L. picturatus picturatus</emphasis>
(Peters)
</taxonomicName>
appears to be close; its coloration is in some respects similar but does not seem to fall within the wide variation … chevrons of the throat equally distinct in the female (only two V’s in three of the
<typeStatus id="54A106E1FFC0FFE8FC06460CFBA6F93A" box="[964,1066,1654,1678]" pageId="23" pageNumber="470" type="paratype">paratypes</typeStatus>
) narrower than in
<taxonomicName id="4C1AC3C0FFC0FFE8FB29460CFACAF93A" ID-CoL="3WQ3T" authority="Venter" authorityName="Venter" box="[1259,1350,1654,1678]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC0FFE8FB29460CFACAF93A" box="[1259,1350,1654,1678]" italics="true" pageId="23" pageNumber="470">gutturalis</emphasis>
</taxonomicName>
… Venter and enlarged subcaudals immaculate yellow in life—an apparently constant distinction from
<taxonomicName id="4C1AC3C0FFC0FFE8FBAA46CFFB81F958" ID-CoL="3WQ3T" authority="' (Schmidt 1919: 466)" authorityName="' (Schmidt" authorityYear="1919" baseAuthorityName="Schmidt" baseAuthorityPageNumber="466" baseAuthorityYear="1919" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="subSpecies" species="picturatus" subSpecies="gutturalis">
<emphasis id="B96E6451FFC0FFE8FBAA46CFFA21F979" box="[1128,1453,1717,1741]" italics="true" pageId="23" pageNumber="470">Lygodactylus picturatus gutturalis</emphasis>
’ (
<bibRefCitation id="EF8BC5B2FFC0FFE8FCF746AEFB89F958" author="Schmidt KP" box="[821,1029,1748,1772]" pageId="23" pageNumber="470" pagination="385 - 624" refId="ref37222" refString="Schmidt KP. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909 - 1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bulletin of the American Museum of Natural History 1919; 39: 385 - 624." type="journal article" year="1919">Schmidt 1919: 466</bibRefCitation>
)
</taxonomicName>
. However,
<bibRefCitation id="EF8BC5B2FFC0FFE8FB4646AEFAF4F958" author="Loveridge A" box="[1156,1400,1748,1772]" pageId="23" pageNumber="470" pagination="1 - 469" refId="ref35578" refString="Loveridge A. Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology 1947; 98: 1 - 469." type="journal article" year="1947">Loveridge (1947: 227)</bibRefCitation>
, relegated this taxon to a subspecies of
<taxonomicName id="4C1AC3C0FFC0FFE8FB5F468EFA97F8BF" ID-CoL="3WQ4R" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[1181,1307,1780,1803]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFC0FFE8FB5F468EFA97F8BF" box="[1181,1307,1780,1803]" italics="true" pageId="23" pageNumber="470">L. picturatus</emphasis>
</taxonomicName>
, stating: ‘
<emphasis id="B96E6451FFC0FFE8FA46468EFA13F8BF" box="[1412,1439,1780,1803]" italics="true" pageId="23" pageNumber="470">…</emphasis>
a somewhat doubtful race, differing from
<taxonomicName id="4C1AC3C0FFC0FFE8FB094769FAAAF89F" ID-CoL="3WQ3T" box="[1227,1318,1811,1835]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC0FFE8FB094769FAAAF89F" box="[1227,1318,1811,1835]" italics="true" pageId="23" pageNumber="470">gutturalis</emphasis>
</taxonomicName>
only in gular and subcaudal markings
<emphasis id="B96E6451FFC0FFE8FBE44749FBCDF8FE" box="[1062,1089,1843,1866]" italics="true" pageId="23" pageNumber="470">…</emphasis>
’. This taxonomic action was ignored by Pasteur, 1965(1964), but followed by
<bibRefCitation id="EF8BC5B2FFC0FFE8FB0A472BFAF4F8DD" author="Wermuth H" box="[1224,1400,1873,1897]" pageId="23" pageNumber="470" pagination="1 - 246" refId="ref37733" refString="Wermuth H. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Das Tierreich 1965; 80: I - XXIII, 1 - 246." type="journal article" year="1965">Wermuth (1965)</bibRefCitation>
, both authors without justification.
<bibRefCitation id="EF8BC5B2FFC0FFE8FB96470BFB52F83D" author="Kluge AG" box="[1108,1246,1905,1929]" pageId="23" pageNumber="470" refId="ref35077" refString="Kluge AG. Gekkonoid Lizard Taxonomy. San Diego: International Gecko Society, 1993." type="book" year="1993">Kluge (1993)</bibRefCitation>
included
<taxonomicName id="4C1AC3C0FFC0FFE8FA824708FA3FF83D" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[1344,1459,1905,1929]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFC0FFE8FA824708FA3FF83D" box="[1344,1459,1905,1929]" italics="true" pageId="23" pageNumber="470">L. depressus</emphasis>
</taxonomicName>
as a full species, in his checklist again without any justification of his taxonomic decision.
</paragraph>
<paragraph id="8BA5B843FFCFFFE7FF5E40EAFA12FE55" blockId="24.[128,778,144,1327]" lastBlockId="24.[825,1474,144,731]" pageId="24" pageNumber="471">
Since then this species has been collected in different regions of the
<collectingRegion id="49DE76A1FFCFFFE7FF0440D5FE80FF73" box="[198,268,175,199]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="24" pageNumber="471">Congo</collectingRegion>
Basin. However, this material was overlooked in many publications that revisited the
<taxonomicName id="4C1AC3C0FFCFFFE7FDC740B5FD08FF52" ID-CoL="3WQ4R" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[517,644,207,230]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFCFFFE7FDC740B5FD08FF52" box="[517,644,207,230]" italics="true" pageId="24" pageNumber="471">L. picturatus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EF8BC5B2FFCFFFE7FD1C40B5FF7CFEB2" author="Roll B &amp; Prohl H &amp; Hoffmann KP" pageId="24" pageNumber="471" pagination="327 - 35" refId="ref36822" refString="Roll B, Prohl H, Hoffmann KP. Multigene phylogenetic analysis of Lygodactylus dwarf geckos (Squamata: Gekkonidae). Molecular Phylogenetics and Evolution 2010; 56: 327 - 35. https: // doi. org / 10.1016 / j. ympev. 2010.02.002" type="journal article" year="2010">
Röll
<emphasis id="B96E6451FFCFFFE7FF434095FF38FEB2" box="[129,180,238,262]" italics="true" pageId="24" pageNumber="471">et al.</emphasis>
2010
</bibRefCitation>
,
<bibRefCitation id="EF8BC5B2FFCFFFE7FF3D4094FE5EFEB2" author="Malonza PK &amp; Granthon C &amp; Williams DA" box="[255,466,238,262]" pageId="24" pageNumber="471" pagination="418 - 28" refId="ref35732" refString="Malonza PK, Granthon C, Williams DA. A new species of dwarf gecko in the genus Lygodactylus (Squamata: Gekkonidae) from central Kenya. Zootaxa 2016; 4061: 418 - 28. https: // doi. org / 10.11646 / zootaxa. 4061.4.6" type="journal article" year="2016">
Malonza
<emphasis id="B96E6451FFCFFFE7FEA04095FE19FEB2" box="[354,405,238,262]" italics="true" pageId="24" pageNumber="471">et al.</emphasis>
2016
</bibRefCitation>
,
<bibRefCitation id="EF8BC5B2FFCFFFE7FE234094FD99FEB2" author="Malonza PK &amp; Bauer AM &amp; Granthon C" box="[481,533,238,262]" pageId="24" pageNumber="471" pagination="308 - 20" refId="ref35682" refString="Malonza PK, Bauer AM, Granthon C et al. A new species of gecko of the genus Lygodactylus (Sauria: Gekkonidae) from southeastern Kenya. Zootaxa 2019; 4609: 308 - 20. https: // doi. org / 10.11646 / zootaxa. 4609.2.6" type="journal article" year="2019">2019</bibRefCitation>
). We recovered a genetic clade that is sister to
<taxonomicName id="4C1AC3C0FFCFFFE7FE464174FE75FE91" ID-CoL="3WQ3T" box="[388,505,269,293]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCFFFE7FE464174FE75FE91" box="[388,505,269,293]" italics="true" pageId="24" pageNumber="471">L. gutturalis</emphasis>
</taxonomicName>
, from the rainforest of the
<collectingRegion id="49DE76A1FFCFFFE7FF424157FF4AFEF1" box="[128,198,301,325]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="24" pageNumber="471">Congo</collectingRegion>
Basin (
<figureCitation id="1321A4C6FFCFFFE7FED04157FEC0FEF1" box="[274,332,301,325]" captionStart="Figure 1" captionStartId="9.[114,179,1431,1455]" captionTargetBox="[115,1460,146,1402]" captionTargetId="figure-247@9.[113,1460,144,1403]" captionTargetPageId="9" captionText="Figure 1. Maximum likelihood analysis based on 532 bp of the mitochondrial 16S rRNA gene, for all available samples of the Afro-American Lygodactylus Clade (Clade C of Gippner et al. 2021). Support values (ML BS = Maximum Likelihood bootstrap values; BI PP = Bayesian Inference posterior probabilities) are shown graphically at the nodes according to the colours shown in the inset key." pageId="24" pageNumber="471">Figs 1</figureCitation>
,
<figureCitation id="1321A4C6FFCFFFE7FE944157FEEFFEF1" box="[342,355,301,325]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="24" pageNumber="471">6</figureCitation>
) that differs by
<emphasis id="B96E6451FFCFFFE7FDC24157FD85FEF0" box="[512,521,301,324]" italics="true" pageId="24" pageNumber="471">c</emphasis>
. 9.04% (
<emphasis id="B96E6451FFCFFFE7FDA04156FD05FEF0" box="[610,649,300,324]" italics="true" pageId="24" pageNumber="471">16S</emphasis>
uncorrected p-distance; Table 2) from its sister taxon. Among this material, we included specimens from Lake Tumba, where historical material of
<taxonomicName id="4C1AC3C0FFCFFFE7FF1741F1FEC5FE17" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[213,329,395,419]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FF1741F1FEC5FE17" box="[213,329,395,419]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
was previously collected. Consequently, we revised historical material collected from the
<collectingRegion id="49DE76A1FFCFFFE7FD9341D0FD1BFE76" box="[593,663,426,450]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="24" pageNumber="471">Congo</collectingRegion>
Basin, tentatively attributed to
<taxonomicName id="4C1AC3C0FFCFFFE7FE9F41B0FE59FE55" ID-CoL="3WQ3T" box="[349,469,457,481]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCFFFE7FE9F41B0FE59FE55" box="[349,469,457,481]" italics="true" pageId="24" pageNumber="471">L. gutturalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1AC3C0FFCFFFE7FDCE41B0FD0EFE55" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[524,642,457,481]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FDCE41B0FD0EFE55" box="[524,642,457,481]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
(Supporting Information, Table S2). After morphological examination and comparison of preserved specimens (faded), we observed a complete overlap between specimens ascribed to these two taxa, failing to identify any morphological differences based on pholidosis or gular ornamentation. In his description of
<taxonomicName id="4C1AC3C0FFCFFFE7FD37421DFE45FD29" ID-CoL="3WQ3G" authority=", Schmidt (1919: 466)" authorityName=", Schmidt" authorityPageNumber="466" authorityYear="1919" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FD37421DFF56FD29" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
,
<bibRefCitation id="EF8BC5B2FFCFFFE7FF2B42FFFE45FD29" author="Schmidt KP" box="[233,457,645,669]" pageId="24" pageNumber="471" pagination="385 - 624" refId="ref37222" refString="Schmidt KP. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909 - 1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bulletin of the American Museum of Natural History 1919; 39: 385 - 624." type="journal article" year="1919">Schmidt (1919: 466)</bibRefCitation>
</taxonomicName>
remarked: ‘… coloration dark blueish grey, irregularly mottled with black, more heavily anteriorly …’. However, he also made reference to colour variation recorded by Herbert O. Lang (collector): ‘…
<specimenCount id="9D1C73CAFFCFFFE7FD89429EFD57FD4F" box="[587,731,739,763]" count="1" pageId="24" pageNumber="471" type="generic">one specimen</specimenCount>
was entirely black when caught, turning blueish grey when injected …’. We observed that
<specimenCount id="9D1C73CAFFCFFFE7FE9A4358FE63FC8E" box="[344,495,802,826]" count="2" pageId="24" pageNumber="471" type="generic">two specimens</specimenCount>
allocated to
<taxonomicName id="4C1AC3C0FFCFFFE7FDAE4359FD53FC8E" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[620,735,802,826]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FDAE4359FD53FC8E" box="[620,735,802,826]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
and collected at Lotende (a river at Mabali, the site of a colonial-era research station at Lake Tumba;
<bibRefCitation id="EF8BC5B2FFCFFFE7FE22431AFDE6FCCC" author="Marlier G" box="[480,618,864,888]" pageId="24" pageNumber="471" pagination="352 - 85" refId="ref35781" refString="Marlier G. Recherches hydrobiologiques au lac Tumba (Congo Belge, Province de l'Equateur). Hydrobiologia 1958; 10: 352 - 85. https: // doi. org / 10.1007 / BF 00142195" type="journal article" year="1958">Marlier 1958</bibRefCitation>
), Lake Tumba (RMCA 1981.065.R.005) by Laurent, exhibit both colour morphs described for
<taxonomicName id="4C1AC3C0FFCFFFE7FE9F43DAFE43FC03" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[349,463,927,951]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FE9F43DAFE43FC03" box="[349,463,927,951]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
(mottled and uniform). In contrast,
<specimenCount id="9D1C73CAFFCFFFE7FF7A43C5FEC3FC62" box="[184,335,958,982]" count="2" pageId="24" pageNumber="471" type="generic">two specimens</specimenCount>
(RMCA R.8575/A and RMCA R.8575/B) collected from Flandria,
<collectingRegion id="49DE76A1FFCFFFE7FE4843A5FDC2FC42" box="[394,590,990,1018]" country="Democratic Republic of the Congo" name="Equateur" pageId="24" pageNumber="471">Équateur Province</collectingRegion>
,
<collectingCountry id="F30DF8D3FFCFFFE7FD9E43A4FD1EFC42" box="[604,658,990,1014]" name="Democratic Republic of the Congo" pageId="24" pageNumber="471">DRC</collectingCountry>
(~
<quantity id="4CE215A6FFCFFFE7FD7143A4FC85FC42" box="[691,777,990,1014]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="471" unit="km" value="110.0">110 km</quantity>
north-east from the previous locality) and identified by de Witte in 1923 (RMCA unpublished data) as
<taxonomicName id="4C1AC3C0FFCFFFE7FDE34467FD16FB80" ID-CoL="3WQ3T" box="[545,666,1052,1076]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCFFFE7FDE34467FD16FB80" box="[545,666,1052,1076]" italics="true" pageId="24" pageNumber="471">L. gutturalis</emphasis>
</taxonomicName>
, show the same two dorsal patterns as the previous specimens mentioned above (
<figureCitation id="1321A4C6FFCFFFE7FF0E4421FE9CFBC7" box="[204,272,1115,1139]" captionStart="Figure 13" captionStartId="24.[129,194,1924,1948]" captionTargetBox="[132,1470,1394,1896]" captionTargetId="figure-744@24.[129,1473,1388,1897]" captionTargetPageId="24" captionText="Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC." pageId="24" pageNumber="471">Fig. 13</figureCitation>
). Two additional specimens (RMCA 1038/B and 1038/C) were collected from the Ituri Forest (without a precise locality within this region, but in the same region as the
<typeStatus id="54A106E1FFCFFFE7FD1944E0FC85FB06" box="[731,777,1178,1202]" pageId="24" pageNumber="471">type</typeStatus>
locality of
<taxonomicName id="4C1AC3C0FFCFFFE7FF2D44C0FEE9FB65" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[239,357,1209,1233]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FF2D44C0FEE9FB65" box="[239,357,1209,1233]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
) and tentatively assigned to
<taxonomicName id="4C1AC3C0FFCFFFE7FD5044C0FC85FB65" ID-CoL="3WQ3T" box="[658,777,1209,1233]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCFFFE7FD5044C0FC85FB65" box="[658,777,1209,1233]" italics="true" pageId="24" pageNumber="471">L. gutturalis</emphasis>
</taxonomicName>
by de Witte in 1923 (RMCA unpublished data). However, these specimens have morphological characters that are intermediate between these two taxa. Thus, material from
<collectingCountry id="F30DF8D3FFCFFFE7FD64456DFD50FA9B" box="[678,732,1303,1327]" name="Democratic Republic of the Congo" pageId="24" pageNumber="471">DRC</collectingCountry>
and
<collectingCountry id="F30DF8D3FFCFFFE7FCFB40EAFC0EFF1C" box="[825,898,144,168]" name="Angola" pageId="24" pageNumber="471">Angola</collectingCountry>
(previously attributed to
<taxonomicName id="4C1AC3C0FFCFFFE7FB4440EAFB76FF1C" ID-CoL="3WQ3T" box="[1158,1274,144,168]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCFFFE7FB4440EAFB76FF1C" box="[1158,1274,144,168]" italics="true" pageId="24" pageNumber="471">L. gutturalis</emphasis>
</taxonomicName>
), shows substantial morphological overlap with topotypic material of
<taxonomicName id="4C1AC3C0FFCFFFE7FA8440CAFA30FF73" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[1350,1468,175,199]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FA8440CAFA30FF73" box="[1350,1468,175,199]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
, making it impossible to differentiate them from the
<typeStatus id="54A106E1FFCFFFE7FA9640B5FA0EFF53" box="[1364,1410,207,231]" pageId="24" pageNumber="471">type</typeStatus>
series (
<figureCitation id="1321A4C6FFCFFFE7FC864094FC1DFEB1" box="[836,913,237,262]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." pageId="24" pageNumber="471">Figs 15</figureCitation>
,
<figureCitation id="1321A4C6FFCFFFE7FC634094FC37FEB2" box="[929,955,238,262]" captionStart="Figure 16" captionStartId="27.[112,177,961,985]" captionTargetBox="[124,1455,157,932]" captionTargetId="figure-520@27.[114,1458,144,933]" captionTargetPageId="27" captionText="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="24" pageNumber="471">16</figureCitation>
). The specimen collected from
<collectingCountry id="F30DF8D3FFCFFFE7FAD14094FAD0FEB2" box="[1299,1372,238,262]" name="Angola" pageId="24" pageNumber="471">Angola</collectingCountry>
(MNCN 50772) has a dark blueish coloration
<emphasis id="B96E6451FFCFFFE7FB074174FB71FE91" box="[1221,1277,269,293]" italics="true" pageId="24" pageNumber="471">in life</emphasis>
(
<figureCitation id="1321A4C6FFCFFFE7FAD24177FAE7FE91" box="[1296,1387,269,293]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." pageId="24" pageNumber="471">Fig. 15G</figureCitation>
), as described in
<taxonomicName id="4C1AC3C0FFCFFFE7FC644157FB97FEF0" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[934,1051,300,324]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FC644157FB97FEF0" box="[934,1051,300,324]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
. Consequently, due to the high morphological and geographic overlap, we consider that this clade, sister to
<emphasis id="B96E6451FFCFFFE7FC964116FC67FE37" box="[852,1003,363,387]" italics="true" pageId="24" pageNumber="471">
<taxonomicName id="4C1AC3C0FFCFFFE7FC964116FC45FE37" ID-CoL="3WQ3T" box="[852,969,363,387]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="gutturalis">L. gutturalis</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFCFFFE7FC124116FC67FE37" box="[976,1003,364,387]" pageId="24" pageNumber="471" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1321A4C6FFCFFFE7FC394111FBBFFE37" box="[1019,1075,363,387]" captionStart="Figure 1" captionStartId="9.[114,179,1431,1455]" captionTargetBox="[115,1460,146,1402]" captionTargetId="figure-247@9.[113,1460,144,1403]" captionTargetPageId="9" captionText="Figure 1. Maximum likelihood analysis based on 532 bp of the mitochondrial 16S rRNA gene, for all available samples of the Afro-American Lygodactylus Clade (Clade C of Gippner et al. 2021). Support values (ML BS = Maximum Likelihood bootstrap values; BI PP = Bayesian Inference posterior probabilities) are shown graphically at the nodes according to the colours shown in the inset key." pageId="24" pageNumber="471">Fig. 1</figureCitation>
), represents
<taxonomicName id="4C1AC3C0FFCFFFE7FB7B4116FAA1FE37" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[1209,1325,363,387]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FB7B4116FAA1FE37" box="[1209,1325,363,387]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
. Nevertheless, we recommend caution until topotypic material can be included in a phylogenetic context, because the possibility of further cryptic diversification within this group cannot be excluded.
</paragraph>
<paragraph id="8BA5B843FFCFFFE7FC974192FCF5FD6F" blockId="24.[825,1474,144,731]" pageId="24" pageNumber="471">
A photographic record from Molondou,
<collectingRegion id="49DE76A1FFCFFFE7FAF64193FA32FDB4" box="[1332,1470,488,512]" country="Cameroon" name="East" pageId="24" pageNumber="471">East Region</collectingRegion>
,
<collectingCountry id="F30DF8D3FFCFFFE7FCFB4272FC25FD94" box="[825,937,520,544]" name="Cameroon" pageId="24" pageNumber="471">Cameroon</collectingCountry>
, reported by Chirio and LeBreton (2007), and attributed to
<taxonomicName id="4C1AC3C0FFCFFFE7FC6C4252FBAEFD8B" authorityName=", Schmidt" authorityYear="1919" box="[942,1058,551,575]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FC6C4252FBAEFD8B" box="[942,1058,551,575]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
, does not agree with the diagnostic dark blueish coloration mentioned by
<bibRefCitation id="EF8BC5B2FFCFFFE7FB44423CFAA7FDEA" author="Schmidt KP" box="[1158,1323,582,606]" pageId="24" pageNumber="471" pagination="385 - 624" refId="ref37222" refString="Schmidt KP. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909 - 1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bulletin of the American Museum of Natural History 1919; 39: 385 - 624." type="journal article" year="1919">Schmidt (1919)</bibRefCitation>
. Therefore, because of the highly diverse character of the
<taxonomicName id="4C1AC3C0FFCFFFE7FB37421CFADAFDC9" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[1269,1366,614,637]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFCFFFE7FB37421CFADAFDC9" box="[1269,1366,614,637]" italics="true" pageId="24" pageNumber="471">picturatus</emphasis>
</taxonomicName>
subgroup, records from
<collectingCountry id="F30DF8D3FFCFFFE7FC0442FFFBB8FD29" box="[966,1076,645,669]" name="Cameroon" pageId="24" pageNumber="471">Cameroon</collectingCountry>
and other localities west and north of the Ubangi and
<collectingCountry id="F30DF8D3FFCFFFE7FC2642DEFBA6FD08" box="[996,1066,676,700]" name="Democratic Republic of the Congo" pageId="24" pageNumber="471">Congo</collectingCountry>
rivers might belong to an undescribed taxon.
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCFFFE6FCFB437FFD28FA33" lastPageId="25" lastPageNumber="472" pageId="24" pageNumber="471" type="materials_examined">
<paragraph id="8BA5B843FFCFFFE7FCFB437FFBE1FCE9" blockId="24.[825,1474,773,861]" pageId="24" pageNumber="471">
<materialsCitation id="3B72B21EFFCFFFE7FCFB437FFBE1FCE9" collectingDate="1914-07-05" collectionCode="AMNH" collectorName="Herbert O. Lang." country="Democratic Republic of the Congo" county="Haut-Uele Province" location="Medje" municipality="Ituri Forest" pageId="24" pageNumber="471" specimenCode="AMNH 10345" specimenCount="1" typeStatus="holotype">
<emphasis id="B96E6451FFCFFFE7FCFB437FFC1EFCA9" box="[825,914,773,797]" italics="true" pageId="24" pageNumber="471">
<typeStatus id="54A106E1FFCFFFE7FCFB437FFC1EFCA9" box="[825,914,773,797]" pageId="24" pageNumber="471" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="1321A4C6FFCFFFE7FC65437FFC60FCAA" box="[935,1004,773,798]" captionStart="Figure 14" captionStartId="25.[113,178,1107,1131]" captionTargetBox="[115,1458,147,1078]" captionTargetId="figure-474@25.[113,1460,144,1079]" captionTargetPageId="25" captionText="Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH)." pageId="24" pageNumber="471">Fig. 14</figureCitation>
):
<specimenCode id="DBBC1038FFCFFFE7FBC6437CFB11FCA9" box="[1028,1181,773,798]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="471">AMNH 10345</specimenCode>
, adult male with original tail, collected at
<location id="8EC5EE98FFCFFFE7FC71435FFC7AFC89" LSID="urn:lsid:plazi:treatment:03B30955FFC0FFE4FF7147FBFDD3F87F:8EC5EE98FFCFFFE7FC71435FFC7AFC89" box="[947,1014,805,829]" country="Democratic Republic of the Congo" county="Haut-Uele Province" municipality="Ituri Forest" name="Medje" pageId="24" pageNumber="471">Medje</location>
,
<collectingMunicipality id="6BC12239FFCFFFE7FC3D435FFBF8FC88" box="[1023,1140,805,829]" pageId="24" pageNumber="471">Ituri Forest</collectingMunicipality>
,
<collectingCounty id="62C4C0CFFFCFFFE7FBBF435FFAC6FC89" box="[1149,1354,805,829]" pageId="24" pageNumber="471">Haut-Uele Province</collectingCounty>
,
<collectingCountry id="F30DF8D3FFCFFFE7FA91435FFA07FC89" box="[1363,1419,805,829]" name="Democratic Republic of the Congo" pageId="24" pageNumber="471">DRC</collectingCountry>
, on
<date id="FFA49E83FFCFFFE7FA77435FFC16FCE8" pageId="24" pageNumber="471" value="1914-07-05">
<collectingDate id="EFE0676BFFCFFFE7FA77435FFC16FCE8" pageId="24" pageNumber="471" value="1914-07-05">5 July 1914</collectingDate>
</date>
by
<collectorName id="26EFDD95FFCFFFE7FC7F433EFBE1FCE9" box="[957,1133,836,861]" pageId="24" pageNumber="471">Herbert O. Lang.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BA5B843FFCFFFE6FCFB4307FD28FA33" blockId="24.[825,1474,893,1325]" lastBlockId="25.[113,762,1235,1416]" lastPageId="25" lastPageNumber="472" pageId="24" pageNumber="471">
<materialsCitation id="3B72B21EFFCFFFE7FCFB4307FC1CFC01" country="Democratic Republic of the Congo" location="DRC" pageId="24" pageNumber="471" specimenCount="22">
<emphasis id="B96E6451FFCFFFE7FCFB4307FA85FC21" box="[825,1289,893,918]" italics="true" pageId="24" pageNumber="471">
<typeStatus id="54A106E1FFCFFFE7FCFB4307FBEFFC21" box="[825,1123,893,917]" isOtherTypeStatus="true" pageId="24" pageNumber="471">Additional material examined</typeStatus>
(
<specimenCount id="9D1C73CAFFCFFFE7FBB64304FB71FC21" box="[1140,1277,894,918]" count="12" pageId="24" pageNumber="471" type="generic">12 specimens</specimenCount>
):
</emphasis>
<emphasis id="B96E6451FFCFFFE7FAD94304FAAAFC22" bold="true" box="[1307,1318,894,918]" pageId="24" pageNumber="471">•</emphasis>
<emphasis id="B96E6451FFCFFFE7FAED4304FAEEFC21" box="[1327,1378,894,917]" italics="true" pageId="24" pageNumber="471">
<collectingCountry id="F30DF8D3FFCFFFE7FAED4304FAEEFC21" box="[1327,1378,894,917]" name="Democratic Republic of the Congo" pageId="24" pageNumber="471">DRC</collectingCountry>
</emphasis>
(
<specimenCount id="9D1C73CAFFCFFFE7FAB14304FC05FC01" count="10" pageId="24" pageNumber="471" type="generic">10 specimens</specimenCount>
)
</materialsCitation>
:
<materialsCitation id="3B72B21EFFCFFFE7FC5943E7FA0CFC01" box="[923,1408,925,949]" collectionCode="RMCA, R" country="Democratic Republic of the Congo" pageId="24" pageNumber="471" specimenCode="AMNH 10346" specimenCount="1">
<collectionCode id="ED0B2086FFCFFFE7FC5943E7FC6BFC01" box="[923,999,925,949]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="24" pageNumber="471" type="Museum">RMCA</collectionCode>
<collectionCode id="ED0B2086FFCFFFE7FC2D43E7FB8DFC01" box="[1007,1025,925,949]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24" pageNumber="471">R</collectionCode>
.3216 (formerly
<specimenCode id="DBBC1038FFCFFFE7FB6F43E7FAC7FC01" box="[1197,1355,925,949]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="471">AMNH 10346</specimenCode>
) and
</materialsCitation>
<materialsCitation id="3B72B21EFFCFFFE7FA4A43E7FB5DFC47" collectingDate="1914-07-05" collectionCode="MCZ" collectorName="Herbert O. Lang." country="Democratic Republic of the Congo" county="Haut-Uele Province" location="Medje" municipality="Ituri Forest" pageId="24" pageNumber="471" specimenCode="R45987, AMNH 10344" specimenCount="1">
<collectionCode id="ED0B2086FFCFFFE7FA4A43E7FA4EFC01" box="[1416,1474,925,949]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="24" pageNumber="471" type="Museum">MCZ</collectionCode>
<specimenCode id="DBBC1038FFCFFFE7FCFB43C6FC07FC60" box="[825,907,956,980]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24" pageNumber="471">R45987</specimenCode>
(formerly
<specimenCode id="DBBC1038FFCFFFE7FBC543C6FB26FC60" box="[1031,1194,956,980]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="471">AMNH 10344</specimenCode>
), males (
<typeStatus id="54A106E1FFCFFFE7FAD343C6FAF0FC60" box="[1297,1404,956,980]" pageId="24" pageNumber="471" type="paratype">paratypes</typeStatus>
), with same collection data as the holotype
</materialsCitation>
;
<materialsCitation id="3B72B21EFFCFFFE7FB2343A6FAF9FC47" box="[1249,1397,987,1011]" collectionCode="UTEP" country="Democratic Republic of the Congo" pageId="24" pageNumber="471" specimenCode="UTEP 22579" specimenCount="1">
<specimenCode id="DBBC1038FFCFFFE7FB2343A6FAF9FC47" box="[1249,1397,987,1011]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="24" pageNumber="471" type="Museum">UTEP 22579</specimenCode>
</materialsCitation>
/
<materialsCitation id="3B72B21EFFCFFFE7FABF43A1FB7AFBA7" collectionCode="UTEP, ELI" country="Democratic Republic of the Congo" pageId="24" pageNumber="471" specimenCode="UTEP 22582, ELI 2128, ELI 2152" specimenCount="1">
<specimenCode id="DBBC1038FFCFFFE7FABF43A1FCF6FBA7" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="24" pageNumber="471" type="Museum">UTEP 22582</specimenCode>
(
<specimenCode id="DBBC1038FFCFFFE7FC494381FC7BFBA7" box="[907,1015,1019,1043]" collectionCode="ELI" pageId="24" pageNumber="471">ELI 2128</specimenCode>
/
<specimenCode id="DBBC1038FFCFFFE7FBCA4381FBFEFBA7" box="[1032,1138,1019,1043]" collectionCode="ELI" pageId="24" pageNumber="471">ELI 2152</specimenCode>
), males, and
</materialsCitation>
<materialsCitation id="3B72B21EFFCFFFE7FAC24381FA35FBA7" box="[1280,1465,1019,1043]" collectionCode="UTEP" country="Democratic Republic of the Congo" pageId="24" pageNumber="471" specimenCode="UTEP 22580-81" specimenCount="1">
<specimenCode id="DBBC1038FFCFFFE7FAC24381FA35FBA7" box="[1280,1465,1019,1043]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="24" pageNumber="471" type="Museum">UTEP 22580–81</specimenCode>
</materialsCitation>
/
<materialsCitation id="3B72B21EFFCFFFE7FCFB4461FBDEFB24" collectingDate="2013-07-06" collectionCode="UTEP, ELI" collectorName="Eli Greenbaum" country="Democratic Republic of the Congo" elevation="311" latitude="-0.7465" location="Lake Tumba" longLatPrecision="7" longitude="18.2243" municipality="Npenda Village" pageId="24" pageNumber="471" specimenCode="UTEP 22583, ELI 2129-30, ELI 2153" specimenCount="1" stateProvince="Equateur">
<specimenCode id="DBBC1038FFCFFFE7FCFB4461FC49FB86" box="[825,965,1050,1074]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="24" pageNumber="471" type="Museum">UTEP 22583</specimenCode>
(
<specimenCode id="DBBC1038FFCFFFE7FC164460FBEBFB86" box="[980,1127,1050,1074]" collectionCode="ELI" pageId="24" pageNumber="471">ELI 2129–30</specimenCode>
/
<specimenCode id="DBBC1038FFCFFFE7FBB44461FB52FB86" box="[1142,1246,1050,1074]" collectionCode="ELI" pageId="24" pageNumber="471">ELI 2153</specimenCode>
), females, collected at
<collectingMunicipality id="6BC12239FFCFFFE7FCFB4443FC56FBE5" box="[825,986,1081,1105]" pageId="24" pageNumber="471">Npenda Village</collectingMunicipality>
, north-east of
<location id="8EC5EE98FFCFFFE7FBAD4443FB78FBE5" LSID="urn:lsid:plazi:treatment:03B30955FFC0FFE4FF7147FBFDD3F87F:8EC5EE98FFCFFFE7FBAD4443FB78FBE5" box="[1135,1268,1081,1105]" country="Democratic Republic of the Congo" latitude="-0.7465" longLatPrecision="7" longitude="18.2243" municipality="Npenda Village" name="Lake Tumba" pageId="24" pageNumber="471" stateProvince="Equateur">Lake Tumba</location>
,
<collectingRegion id="49DE76A1FFCFFFE7FB3C4441FA32FBE5" box="[1278,1470,1081,1110]" country="Democratic Republic of the Congo" name="Equateur" pageId="24" pageNumber="471">Équateur Province</collectingRegion>
,
<geoCoordinate id="EE2EDE84FFCFFFE7FCFB4423FC17FBC5" box="[825,923,1113,1137]" degrees="00.7465" direction="south" orientation="latitude" pageId="24" pageNumber="471" precision="5" value="-0.7465">S00.7465</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFCFFFE7FC654423FB81FBC5" box="[935,1037,1113,1137]" degrees="18.2243" direction="east" orientation="longitude" pageId="24" pageNumber="471" precision="5" value="18.2243">E18.2243</geoCoordinate>
,
<elevation id="00375F70FFCFFFE7FBDA4423FB03FBC5" box="[1048,1167,1113,1137]" metricMagnitude="2" metricUnit="m" metricValue="3.11" pageId="24" pageNumber="471" unit="m" value="311.0">
<quantity id="4CE215A6FFCFFFE7FBDA4423FBD7FBC4" box="[1048,1115,1113,1137]" metricMagnitude="2" metricUnit="m" metricValue="3.11" pageId="24" pageNumber="471" unit="m" value="311.0">311 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCFFFE7FB7B4423FABDFBC5" box="[1209,1329,1113,1137]" pageId="24" pageNumber="471" value="2013-07-06">
<collectingDate id="EFE0676BFFCFFFE7FB7B4423FABDFBC5" box="[1209,1329,1113,1137]" pageId="24" pageNumber="471" value="2013-07-06">6 July 2013</collectingDate>
</date>
by locals and brought to
<collectorName id="26EFDD95FFCFFFE7FC724402FBDEFB24" box="[944,1106,1144,1168]" pageId="24" pageNumber="471">Eli Greenbaum</collectorName>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFCFFFE7FB9C4403FA32FB7B" collectingDate="2010-02-13" collectionCode="UTEP, ELI" collectorName="Chifundera Kusamba" country="Democratic Republic of the Congo" elevation="300" latitude="-0.8" location="Lake Tumba" longLatPrecision="785" longitude="18.15" pageId="24" pageNumber="471" specimenCode="UTEP 22578, ELI 1547" specimenCount="1" stateProvince="Equateur">
<specimenCode id="DBBC1038FFCFFFE7FB9C4403FB66FB24" box="[1118,1258,1144,1168]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="24" pageNumber="471" type="Museum">UTEP 22578</specimenCode>
(
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), female, collected at
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,
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,
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.
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,
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,
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<quantity id="4CE215A6FFCFFFE7FCFB44CDFC0CFB7A" box="[825,896,1207,1231]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="471" unit="m" value="300.0">300 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCFFFE7FC2B44CDFB26FB7B" box="[1001,1194,1207,1231]" pageId="24" pageNumber="471" value="2010-02-13">
<collectingDate id="EFE0676BFFCFFFE7FC2B44CDFB26FB7B" box="[1001,1194,1207,1231]" pageId="24" pageNumber="471" value="2010-02-13">13 February 2010</collectingDate>
</date>
by
<collectorName id="26EFDD95FFCFFFE7FB1844CDFA32FB7B" box="[1242,1470,1207,1231]" pageId="24" pageNumber="471">Chifundera Kusamba</collectorName>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFCFFFE7FCFB44ADFB67FA99" collectingDate="2015-07-04" collectionCode="UTEP, ELI" collectorName="Eli Greenbaum" country="Democratic Republic of the Congo" elevation="455" latitude="-2.75128" location="Katopa" longLatPrecision="1" longitude="25.10403" pageId="24" pageNumber="471" specimenCode="UTEP 22595, ELI 3624" specimenCount="1" stateProvince="Maniema">
<specimenCode id="DBBC1038FFCFFFE7FCFB44ADFC4EFB5A" box="[825,962,1238,1262]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="24" pageNumber="471" type="Museum">UTEP 22595</specimenCode>
(
<specimenCode id="DBBC1038FFCFFFE7FC0C44ACFBBAFB5A" box="[974,1078,1238,1262]" collectionCode="ELI" pageId="24" pageNumber="471">ELI 3624</specimenCode>
), female, collected at
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,
<collectingRegion id="49DE76A1FFCFFFE7FAA344ACFC19FAB9" country="Democratic Republic of the Congo" name="Maniema" pageId="24" pageNumber="471">Maniema Province</collectingRegion>
,
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,
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,
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<quantity id="4CE215A6FFCFFFE7FB5C448FFB68FAB9" box="[1182,1252,1269,1293]" metricMagnitude="2" metricUnit="m" metricValue="4.55" pageId="24" pageNumber="471" unit="m" value="455.0">455 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCFFFE7FA85448FFA4EFAB9" box="[1351,1474,1269,1293]" pageId="24" pageNumber="471" value="2015-07-04">
<collectingDate id="EFE0676BFFCFFFE7FA85448FFA4EFAB9" box="[1351,1474,1269,1293]" pageId="24" pageNumber="471" value="2015-07-04">4 July 2015</collectingDate>
</date>
by locals and brought to
<collectorName id="26EFDD95FFCFFFE7FB8B456FFB67FA99" box="[1097,1259,1301,1325]" pageId="24" pageNumber="471">Eli Greenbaum</collectorName>
</materialsCitation>
;
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<specimenCode id="DBBC1038FFCFFFE7FB3B456FFA0BFA99" box="[1273,1415,1301,1325]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="24" pageNumber="471" type="Museum">UTEP 22597</specimenCode>
(
<specimenCode id="DBBC1038FFCFFFE6FA55456FFF26FB5F" collectionCode="ELI" lastPageId="25" lastPageNumber="472" pageId="24" pageNumber="471">ELI 3585</specimenCode>
), male collected from
<collectorName id="26EFDD95FFCEFFE6FE6144A9FE63FB5E" box="[419,495,1235,1258]" pageId="25" pageNumber="472">Katopa</collectorName>
</materialsCitation>
,
<materialsCitation id="3B72B21EFFCEFFE6FE3C44A9FDA2FA9D" collectingDate="2015-07-02" collectionCode="ICCN" collectorName="Eli Greenbaum." country="Angola" elevation="450" latitude="-2.74769" location="Camp" longLatPrecision="1" longitude="25.10323" pageId="25" pageNumber="472" specimenCount="2" stateProvince="Maniema">
<collectionCode id="ED0B2086FFCEFFE6FE3C44A9FDB3FB5F" box="[510,575,1235,1259]" pageId="25" pageNumber="472">ICCN</collectionCode>
<location id="8EC5EE98FFCEFFE6FD8844A9FD06FB5F" LSID="urn:lsid:plazi:treatment:03B30955FFC0FFE4FF7147FBFDD3F87F:8EC5EE98FFCEFFE6FD8844A9FD06FB5F" box="[586,650,1235,1259]" country="Angola" latitude="-2.74769" longLatPrecision="1" longitude="25.10323" name="Camp" pageId="25" pageNumber="472" stateProvince="Maniema">Camp</location>
,
<collectingRegion id="49DE76A1FFCEFFE6FD5B44A9FF41FABE" country="Democratic Republic of the Congo" name="Maniema" pageId="25" pageNumber="472">Maniema Province</collectingRegion>
,
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,
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,
<elevation id="00375F70FFCEFFE6FE0E4488FDCCFABE" box="[460,576,1266,1290]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="25" pageNumber="472" unit="m" value="450.0">
<quantity id="4CE215A6FFCEFFE6FE0E4488FD82FABE" box="[460,526,1266,1290]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="25" pageNumber="472" unit="m" value="450.0">450 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCEFFE6FDA54488FD57FABE" box="[615,731,1266,1290]" pageId="25" pageNumber="472" value="2015-07-02">
<collectingDate id="EFE0676BFFCEFFE6FDA54488FD57FABE" box="[615,731,1266,1290]" pageId="25" pageNumber="472" value="2015-07-02">2 July 2015</collectingDate>
</date>
by
<collectorName id="26EFDD95FFCEFFE6FFB3456BFE99FA9D" box="[113,277,1297,1321]" pageId="25" pageNumber="472">Eli Greenbaum.</collectorName>
<emphasis id="B96E6451FFCEFFE6FEDE4568FEABFA9E" bold="true" box="[284,295,1298,1322]" pageId="25" pageNumber="472">•</emphasis>
<emphasis id="B96E6451FFCEFFE6FEED456BFEF8FA9D" box="[303,372,1297,1321]" italics="true" pageId="25" pageNumber="472">
<collectingCountry id="F30DF8D3FFCEFFE6FEED456BFEF8FA9D" box="[303,372,1297,1321]" name="Angola" pageId="25" pageNumber="472">Angola</collectingCountry>
</emphasis>
(
<specimenCount id="9D1C73CAFFCEFFE6FE414568FDABFA9D" box="[387,551,1297,1322]" count="2" pageId="25" pageNumber="472" type="generic">two specimens</specimenCount>
)
</materialsCitation>
:
<materialsCitation id="3B72B21EFFCEFFE6FDFA4568FD2CFA33" collectingDate="2021-09-10" collectionCode="MNCN" collectorName="Pedro Vaz Pinto &amp; Timoteo Julio" country="Democratic Republic of the Congo" elevation="7" latitude="-9.19518" location="Barra do Cuanza" longLatPrecision="1" longitude="13.20327" pageId="25" pageNumber="472" specimenCode="MNCN 50771-72" specimenCount="1" stateProvince="Luanda">
<specimenCode id="DBBC1038FFCEFFE6FDFA4568FD76FA9D" box="[568,762,1297,1322]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="25" pageNumber="472" type="Museum">MNCN 50771–72</specimenCode>
(P1-307 and P1-306), males, collected at
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,
<collectingRegion id="49DE76A1FFCEFFE6FFB3452AFEA5FADC" box="[113,297,1360,1384]" country="Angola" name="Luanda" pageId="25" pageNumber="472">Luanda Province</collectingRegion>
,
<geoCoordinate id="EE2EDE84FFCEFFE6FEF9452AFE26FADC" box="[315,426,1360,1384]" degrees="09.19518" direction="south" orientation="latitude" pageId="25" pageNumber="472" precision="1" value="-9.19518">S09.19518</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFCEFFE6FE7E452AFDA2FADC" box="[444,558,1360,1384]" degrees="13.20327" direction="east" orientation="longitude" pageId="25" pageNumber="472" precision="1" value="13.20327">E13.20327</geoCoordinate>
,
<elevation id="00375F70FFCEFFE6FD82452BFD25FADC" box="[576,681,1360,1384]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="472" unit="m" value="7.0">
<quantity id="4CE215A6FFCEFFE6FD82452BFDFCFADC" box="[576,624,1361,1384]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="472" unit="m" value="7.0">7 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCEFFE6FD22452AFE95FA3C" pageId="25" pageNumber="472" value="2021-09-10">
<collectingDate id="EFE0676BFFCEFFE6FD22452AFE95FA3C" pageId="25" pageNumber="472" value="2021-09-10">10 September 2021</collectingDate>
</date>
by
<collectorName id="26EFDD95FFCEFFE6FEFF4515FE6CFA33" box="[317,480,1391,1415]" pageId="25" pageNumber="472">Pedro Vaz Pinto</collectorName>
and
<collectorName id="26EFDD95FFCEFFE6FDD24515FD2CFA33" box="[528,672,1391,1415]" pageId="25" pageNumber="472">Timoteo Julio</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF65E8CBFFCFFFE7FF4347FEFD3DF80D" pageId="24" pageNumber="471" startId="24.[129,194,1924,1948]" targetBox="[132,1470,1394,1896]" targetPageId="24" targetType="figure">
<paragraph id="8BA5B843FFCFFFE7FF4347FEFD3DF80D" blockId="24.[129,1446,1924,1980]" pageId="24" pageNumber="471">
<emphasis id="B96E6451FFCFFFE7FF4347FEFF69F828" bold="true" box="[129,229,1924,1948]" pageId="24" pageNumber="471">Figure 13.</emphasis>
Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of
<taxonomicName id="4C1AC3C0FFCFFFE7FAF447FFFA2EF828" authorityName=", Schmidt" authorityYear="1919" box="[1334,1442,1924,1948]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="471" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCFFFE7FAF447FFFA2EF828" box="[1334,1442,1924,1948]" italics="true" pageId="24" pageNumber="471">L. depressus</emphasis>
</taxonomicName>
, collected in sympatry at Flandria, Équateur Province, DRC.
</paragraph>
</caption>
<caption id="DF65E8CBFFCEFFE6FFB34429FB85FB17" pageId="25" pageNumber="472" startId="25.[113,178,1107,1131]" targetBox="[115,1458,147,1078]" targetPageId="25" targetType="figure">
<paragraph id="8BA5B843FFCEFFE6FFB34429FB85FB17" blockId="25.[113,1444,1107,1187]" pageId="25" pageNumber="472">
<emphasis id="B96E6451FFCEFFE6FFB34429FF5AFBDF" bold="true" box="[113,214,1107,1131]" pageId="25" pageNumber="472">Figure 14.</emphasis>
Holotype of
<taxonomicName id="4C1AC3C0FFCEFFE6FE90442EFE32FBDF" authorityName=", Schmidt" authorityYear="1919" box="[338,446,1107,1131]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCEFFE6FE90442EFE32FBDF" box="[338,446,1107,1131]" italics="true" pageId="25" pageNumber="472">L. depressus</emphasis>
</taxonomicName>
(AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). C, Details of cloacal region. D, Details of left manus. Photos provided by Lauren Vonnahme: American Museum of Natural History, New York City (AMNH).
</paragraph>
</caption>
<subSubSection id="C300EBC8FFCEFFE4FFB345D0FDB6F9B5" lastPageId="27" lastPageNumber="474" pageId="25" pageNumber="472" type="diagnosis">
<paragraph id="8BA5B843FFCEFFE6FFB345D0FD87F94E" blockId="25.[113,763,1449,1974]" pageId="25" pageNumber="472">
<emphasis id="B96E6451FFCEFFE6FFB345D0FE41FA75" box="[113,461,1449,1473]" italics="true" pageId="25" pageNumber="472">
Diagnosis:
<taxonomicName id="4C1AC3C0FFCEFFE6FF2945D3FE41FA75" authorityName=", Schmidt" authorityYear="1919" box="[235,461,1449,1473]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="depressus">Lygodactylus depressus</taxonomicName>
</emphasis>
is a large-sized
<taxonomicName id="4C1AC3C0FFCEFFE6FDB945D3FD76FA75" box="[635,762,1449,1473]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="genus">
<emphasis id="B96E6451FFCEFFE6FDB945D3FD76FA75" box="[635,762,1449,1473]" italics="true" pageId="25" pageNumber="472">Lygodactylus</emphasis>
</taxonomicName>
with a maximum SVL
<quantity id="4CE215A6FFCEFFE6FEAB45B2FE45FA54" box="[361,457,1480,1504]" metricMagnitude="-2" metricUnit="m" metricValue="3.78" pageId="25" pageNumber="472" unit="mm" value="37.8">37.8 mm</quantity>
(mean 35.6 ±
<quantity id="4CE215A6FFCEFFE6FDAE45B2FD4CFA54" box="[620,704,1480,1504]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="25" pageNumber="472" unit="mm" value="1.4">1.4 mm</quantity>
) that has the typical gular pattern of the
<taxonomicName id="4C1AC3C0FFCEFFE6FE2D459DFDC6FA4B" box="[495,586,1511,1535]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCEFFE6FE2D459DFDC6FA4B" box="[495,586,1511,1535]" italics="true" pageId="25" pageNumber="472">gutturalis</emphasis>
</taxonomicName>
group. Seven to nine supralabials and 6–7 infralabials. Dorsal pholidosis with granular scales that become flattened, larger, and imbricate in original tails. Large triangular mental followed by 2–3 symmetric postmental scales (
<figureCitation id="1321A4C6FFCEFFE6FE44461FFE42F9C9" box="[390,462,1637,1661]" captionStart="Figure 16" captionStartId="27.[112,177,961,985]" captionTargetBox="[124,1455,157,932]" captionTargetId="figure-520@27.[114,1458,144,933]" captionTargetPageId="27" captionText="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="25" pageNumber="472">Fig. 16</figureCitation>
). Males with 7–8 precloacal pores. Ventral pholidosis with large, flattened, and imbricate scales. Ventral scales usually with small denticulation posteriorly. Digits elongated with five terminal scansors on the fourth toe (Supporting Information, Table S6).
</paragraph>
<paragraph id="8BA5B843FFCEFFE6FF4F477BFDF3F802" blockId="25.[113,763,1449,1974]" pageId="25" pageNumber="472">
Like other members of the
<taxonomicName id="4C1AC3C0FFCEFFE6FE634778FD9AF8AD" box="[417,534,1793,1817]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCEFFE6FE634778FD9AF8AD" box="[417,534,1793,1817]" italics="true" pageId="25" pageNumber="472">L. gutturalis</emphasis>
</taxonomicName>
subgroup, this species can be easily differentiated from
<taxonomicName id="4C1AC3C0FFCEFFE6FE7A475BFDA6F88D" authorityName="Gunther" authorityYear="1893" box="[440,554,1825,1849]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFCEFFE6FE7A475BFDA6F88D" box="[440,554,1825,1849]" italics="true" pageId="25" pageNumber="472">L. angularis</emphasis>
</taxonomicName>
group members and from other members of the
<taxonomicName id="4C1AC3C0FFCEFFE6FE4B473BFE66F8EC" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[393,490,1857,1880]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFCEFFE6FE4B473BFE66F8EC" box="[393,490,1857,1880]" italics="true" pageId="25" pageNumber="472">picturatus</emphasis>
</taxonomicName>
subgroup by the gular pattern and dorsal colour pattern (
<figureCitation id="1321A4C6FFCEFFE6FE6C4725FE6AF8C3" box="[430,486,1887,1911]" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." pageId="25" pageNumber="472">Fig. 5</figureCitation>
). It should be noted that
<taxonomicName id="4C1AC3C0FFCEFFE6FD24471AFF47F823" authorityName=", Schmidt" authorityYear="1919" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCEFFE6FD24471AFF47F823" italics="true" pageId="25" pageNumber="472">L. depressus</emphasis>
</taxonomicName>
has a dark blueish grey dorsum
<emphasis id="B96E6451FFCEFFE6FDD94705FDDDF823" box="[539,593,1919,1943]" italics="true" pageId="25" pageNumber="472">in life</emphasis>
, unlike the light blueish dorsal coloration in the
<taxonomicName id="4C1AC3C0FFCEFFE6FE7147E5FD98F802" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[435,532,1951,1974]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFCEFFE6FE7147E5FD98F802" box="[435,532,1951,1974]" italics="true" pageId="25" pageNumber="472">picturatus</emphasis>
</taxonomicName>
subgroup.
</paragraph>
<paragraph id="8BA5B843FFCEFFE6FC8744A9FBCEF88A" blockId="25.[810,1460,1235,1854]" pageId="25" pageNumber="472">
<taxonomicName id="4C1AC3C0FFCEFFE6FC8744A9FBABFB5F" authorityName=", Schmidt" authorityYear="1919" box="[837,1063,1235,1259]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCEFFE6FC8744A9FBABFB5F" box="[837,1063,1235,1259]" italics="true" pageId="25" pageNumber="472">Lygodactylus depressus</emphasis>
</taxonomicName>
can be differentiated from other species within the
<taxonomicName id="4C1AC3C0FFCEFFE6FC074488FBACFABE" box="[965,1056,1266,1290]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCEFFE6FC074488FBACFABE" box="[965,1056,1266,1290]" italics="true" pageId="25" pageNumber="472">gutturalis</emphasis>
</taxonomicName>
subgroup by having a dark blueish grey coloration of the dorsum, with some specimens having a mottled pattern on the dorsum, vs. brownish or light grey without a mottled pattern in
<taxonomicName id="4C1AC3C0FFCEFFE6FBDA452BFB18FADC" box="[1048,1172,1360,1384]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCEFFE6FBDA452BFB18FADC" box="[1048,1172,1360,1384]" italics="true" pageId="25" pageNumber="472">L. gutturalis</emphasis>
</taxonomicName>
. However, blackish specimens can only be differentiated by subtle morphometric and meristic data, being almost impossible to differentiate in the field.
<taxonomicName id="4C1AC3C0FFCEFFE6FCA145D4FBC8FA72" authorityName=", Schmidt" authorityYear="1919" box="[867,1092,1454,1478]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCEFFE6FCA145D4FBC8FA72" box="[867,1092,1454,1478]" italics="true" pageId="25" pageNumber="472">Lygodactylus depressus</emphasis>
</taxonomicName>
can be partially differentiated from
<taxonomicName id="4C1AC3C0FFCEFFE6FCE845B4FC13FA51" box="[810,927,1485,1509]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCEFFE6FCE845B4FC13FA51" box="[810,927,1485,1509]" italics="true" pageId="25" pageNumber="472">L. gutturalis</emphasis>
</taxonomicName>
as follows: eye proportionally smaller in
<taxonomicName id="4C1AC3C0FFCEFFE6FAFD45B4FA3FFA51" authorityName=", Schmidt" authorityYear="1919" box="[1343,1459,1485,1509]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCEFFE6FAFD45B4FA3FFA51" box="[1343,1459,1485,1509]" italics="true" pageId="25" pageNumber="472">L. depressus</emphasis>
</taxonomicName>
(OD/HL ≤ 0.25 vs.
<quantity id="4CE215A6FFCEFFE6FBC14597FB0BF9B1" box="[1027,1159,1517,1541]" metricMagnitude="-3" metricUnit="m" metricValue="7.239" metricValueMax="7.874" metricValueMin="6.604" pageId="25" pageNumber="472" unit="in" value="0.285" valueMax="0.31" valueMin="0.26">0.26–0.31 in</quantity>
<taxonomicName id="4C1AC3C0FFCEFFE6FB534597FA86F9B1" box="[1169,1290,1517,1541]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCEFFE6FB534597FA86F9B1" box="[1169,1290,1517,1541]" italics="true" pageId="25" pageNumber="472">L. gutturalis</emphasis>
</taxonomicName>
); and narrower snout (IN/HW ≤ 0.29 vs.
<quantity id="4CE215A6FFCEFFE6FBFB4676FB36F990" box="[1081,1210,1548,1572]" metricMagnitude="-3" metricUnit="m" metricValue="8.128" metricValueMax="8.636" metricValueMin="7.62" pageId="25" pageNumber="472" unit="in" value="0.32" valueMax="0.34" valueMin="0.3">0.30–0.34 in</quantity>
<taxonomicName id="4C1AC3C0FFCEFFE6FB024677FAB9F990" box="[1216,1333,1548,1572]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCEFFE6FB024677FAB9F990" box="[1216,1333,1548,1572]" italics="true" pageId="25" pageNumber="472">L. gutturalis</emphasis>
</taxonomicName>
). It also differs by a minimum of
<emphasis id="B96E6451FFCEFFE6FBD64656FB91F9F7" box="[1044,1053,1580,1603]" italics="true" pageId="25" pageNumber="472">c</emphasis>
. 9.04% uncorrected p-distance for
<emphasis id="B96E6451FFCEFFE6FA4E4651FA3FF9F7" box="[1420,1459,1579,1603]" italics="true" pageId="25" pageNumber="472">16S</emphasis>
(Table 2), lacks any nuclear haplotype sharing with
<taxonomicName id="4C1AC3C0FFCEFFE6FAFF4631FA3EF9D7" box="[1341,1458,1611,1635]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCEFFE6FAFF4631FA3EF9D7" box="[1341,1458,1611,1635]" italics="true" pageId="25" pageNumber="472">L. gutturalis</emphasis>
</taxonomicName>
in
<emphasis id="B96E6451FFCEFFE6FC8A4610FC08F936" box="[840,900,1642,1666]" italics="true" pageId="25" pageNumber="472">RAG1</emphasis>
(
<figureCitation id="1321A4C6FFCEFFE6FC5B4610FC59F936" box="[921,981,1642,1666]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability&gt; 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="25" pageNumber="472">Fig. 2</figureCitation>
), and habitat (rainforest for
<taxonomicName id="4C1AC3C0FFCEFFE6FAC94611FA0FF936" authorityName=", Schmidt" authorityYear="1919" box="[1291,1411,1642,1666]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCEFFE6FAC94611FA0FF936" box="[1291,1411,1642,1666]" italics="true" pageId="25" pageNumber="472">L. depressus</emphasis>
</taxonomicName>
and sub-Saharan savannah for
<taxonomicName id="4C1AC3C0FFCEFFE6FBF746F0FB26F915" box="[1077,1194,1673,1697]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCEFFE6FBF746F0FB26F915" box="[1077,1194,1673,1697]" italics="true" pageId="25" pageNumber="472">L. gutturalis</emphasis>
</taxonomicName>
). It can also be differentiated from
<taxonomicName id="4C1AC3C0FFCEFFE6FC5646D3FB89F974" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[916,1029,1704,1728]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFCEFFE6FC5646D3FB89F974" box="[916,1029,1704,1728]" italics="true" pageId="25" pageNumber="472">L. dysmicus</emphasis>
</taxonomicName>
by having fewer precloacal pores (7–8 vs.
<quantity id="4CE215A6FFCEFFE6FCE846B2FCDEF954" box="[810,850,1736,1760]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="25" pageNumber="472" unit="in" value="9.0">9 in</quantity>
<taxonomicName id="4C1AC3C0FFCEFFE6FC9846B3FC40F954" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[858,972,1736,1760]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFCEFFE6FC9846B3FC40F954" box="[858,972,1736,1760]" italics="true" pageId="25" pageNumber="472">L. dysmicus</emphasis>
</taxonomicName>
) and nostril never in contact with rostral scale vs. nostril contacting the rostral in
<taxonomicName id="4C1AC3C0FFCEFFE6FB4F4692FB72F94B" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[1165,1278,1767,1791]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="472" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFCEFFE6FB4F4692FB72F94B" box="[1165,1278,1767,1791]" italics="true" pageId="25" pageNumber="472">L. dysmicus</emphasis>
</taxonomicName>
. For a distinction from other species not included above, described herein, see the respective diagnoses below.
</paragraph>
<paragraph id="8BA5B843FFCEFFE4FCE84725FDDCFABF" blockId="25.[810,1459,1887,1974]" lastBlockId="27.[113,763,1048,1291]" lastPageId="27" lastPageNumber="474" pageId="25" pageNumber="472">
<emphasis id="B96E6451FFCEFFE6FCE84725FC51F8C3" box="[810,989,1887,1911]" italics="true" pageId="25" pageNumber="472">Coloration: In life</emphasis>
(
<figureCitation id="1321A4C6FFCEFFE6FC2C4725FBA2F8C3" box="[1006,1070,1887,1911]" captionStart="Figure 15" captionStartId="26.[129,194,1836,1860]" captionTargetBox="[198,1405,144,1808]" captionTargetId="figure-10@26.[198,1405,144,1808]" captionTargetPageId="26" captionText="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." pageId="25" pageNumber="472">Fig.15</figureCitation>
), dorsal colour is highly variable; predominantly dark grey, with lighter grey blotches surrounded by black dots on the flanks, which can form a diffuse dorsolateral band; dark
<emphasis id="B96E6451FFCCFFE4FF2F4463FF71FB84" box="[237,253,1049,1072]" italics="true" pageId="27" pageNumber="474">V</emphasis>
-shaped line between the eyes (absent in darker specimens), and black line from snout to anterior insertion of the forelimb; tail with diffuse bars of lighter grey; throat usually with white background and black chevrons; venter vivid yellow, orange, or cream from tail tip to posterior part of the gular region.
<emphasis id="B96E6451FFCCFFE4FF6944CFFEB4FB78" box="[171,312,1205,1228]" italics="true" pageId="27" pageNumber="474">In preservative</emphasis>
(
<typeStatus id="54A106E1FFCCFFE4FE8444CFFE26FB79" box="[326,426,1205,1229]" pageId="27" pageNumber="474" type="holotype">holotype</typeStatus>
;
<figureCitation id="1321A4C6FFCCFFE4FE7644CFFE76FB79" box="[436,506,1205,1229]" captionStart="Figure 14" captionStartId="25.[113,178,1107,1131]" captionTargetBox="[115,1458,147,1078]" captionTargetId="figure-474@25.[113,1460,144,1079]" captionTargetPageId="25" captionText="Figure 14. Holotype of L. depressus (AMNH 10345) from Medje, Ituri Forest, Haut-Uele Province, DRC. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom).C, Details of cloacal region.D, Details of left manus.Photos provided by Lauren Vonnahme:American Museum of Natural History, New York City (AMNH)." pageId="27" pageNumber="474">Fig. 14</figureCitation>
), dorsum grey with dark brown dots from head to midbody; venter uniform light cream to yellow with whitish colouration on all digits.
</paragraph>
<caption id="DF65E8CBFFCDFFE5FF434756FC23F82C" pageId="26" pageNumber="473" startId="26.[129,194,1836,1860]" targetBox="[198,1405,144,1808]" targetPageId="26" targetType="figure">
<paragraph id="8BA5B843FFCDFFE5FF434756FC23F82C" blockId="26.[129,1451,1836,1944]" pageId="26" pageNumber="473">
<emphasis id="B96E6451FFCDFFE5FF434756FF69F8F0" bold="true" box="[129,229,1836,1860]" pageId="26" pageNumber="473">Figure 15.</emphasis>
Specimens of
<taxonomicName id="4C1AC3C0FFCDFFE5FEAE4757FE54F8F0" authorityName=", Schmidt" authorityYear="1919" box="[364,472,1836,1860]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="473" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCDFFE5FEAE4757FE54F8F0" box="[364,472,1836,1860]" italics="true" pageId="26" pageNumber="473">L. depressus</emphasis>
</taxonomicName>
in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G. (A–C), C.K. (D), P.V.P. (E–G), and J.L.R. (H).
</paragraph>
</caption>
<caption id="DF65E8CBFFCCFFE4FFB243BBFEF8FC41" pageId="27" pageNumber="474" startId="27.[112,177,961,985]" targetBox="[124,1455,157,932]" targetPageId="27" targetType="figure">
<paragraph id="8BA5B843FFCCFFE4FFB243BBFEF8FC41" blockId="27.[112,1456,961,1013]" pageId="27" pageNumber="474">
<emphasis id="B96E6451FFCCFFE4FFB243BBFF58FC6D" bold="true" box="[112,212,961,985]" pageId="27" pageNumber="474">Figure 16.</emphasis>
Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in
<taxonomicName id="4C1AC3C0FFCCFFE4FBE843B8FB1AFC6D" authorityName=", Schmidt" authorityYear="1919" box="[1066,1174,961,985]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="474" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCCFFE4FBE843B8FB1AFC6D" box="[1066,1174,961,985]" italics="true" pageId="27" pageNumber="474">L. depressus</emphasis>
</taxonomicName>
. Red points denote males and blue points denote females.
</paragraph>
</caption>
<paragraph id="8BA5B843FFCCFFE4FFB34557FDB6F9B5" blockId="27.[113,762,1325,1537]" pageId="27" pageNumber="474">
<emphasis id="B96E6451FFCCFFE4FFB34557FF58FAF0" box="[113,212,1325,1348]" italics="true" pageId="27" pageNumber="474">Variation:</emphasis>
Meristic and morphometric data are summarized and depicted in the Supporting Information (Table S6) and
<figureCitation id="1321A4C6FFCCFFE4FD6D4536FF3FFA37" captionStart-0="Figure 15" captionStart-1="Figure 16" captionStartId-0="26.[129,194,1836,1860]" captionStartId-1="27.[112,177,961,985]" captionTargetBox-0="[198,1405,144,1808]" captionTargetBox-1="[124,1455,157,932]" captionTargetId-0="figure-10@26.[198,1405,144,1808]" captionTargetId-1="figure-520@27.[114,1458,144,933]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionText-0="Figure 15. Specimens of L. depressus in life or freshly euthanized from (A) Npenda, Équateur Province, DRC (UTEP 22580), (B) Katopa, Maniema Province, DRC (UTEP 22597), (C) Lake Tumba, Équateur Province, DRC (UTEP 22578), and (E–G) Barra do Cuanza, Cuanza Sul Province, Angola (MNCN 50772). Photo of habitat at (D) Lake Tumba, Équateur Province, DRC and (H) Barra do Cuanza, Cuanza Sul Province, Angola. Photographs by E.G.(A–C), C.K. (D), P.V.P.(E–G), and J.L.R. (H)." captionText-1="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="27" pageNumber="474">Figures 15–16</figureCitation>
.
<taxonomicName id="4C1AC3C0FFCCFFE4FF024511FE2CFA37" authorityName=", Schmidt" authorityYear="1919" box="[192,416,1387,1411]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="474" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCCFFE4FF024511FE2CFA37" box="[192,416,1387,1411]" italics="true" pageId="27" pageNumber="474">Lygodactylus depressus</emphasis>
</taxonomicName>
is the most variable taxon within the
<taxonomicName id="4C1AC3C0FFCCFFE4FF5545F1FE87FA17" box="[151,267,1419,1443]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="474" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCCFFE4FF5545F1FE87FA17" box="[151,267,1419,1443]" italics="true" pageId="27" pageNumber="474">L. gutturalis</emphasis>
</taxonomicName>
group with respect to coloration, particularly the gular pattern (
<figureCitation id="1321A4C6FFCCFFE4FEC645D0FEC6FA76" box="[260,330,1450,1474]" captionStart="Figure 16" captionStartId="27.[112,177,961,985]" captionTargetBox="[124,1455,157,932]" captionTargetId="figure-520@27.[114,1458,144,933]" captionTargetPageId="27" captionText="Figure 16. Dorsal and ventral views of head showing gular ornamentation and pholidosis variation in L. depressus. Red points denote males and blue points denote females." pageId="27" pageNumber="474">Fig. 16</figureCitation>
). Some specimens lose the dorsal pattern, resulting in a uniform dorsal coloration. However, other specimens have a mottled dorsal pattern (
<figureCitation id="1321A4C6FFCCFFE4FE264592FDA5F9B5" box="[484,553,1512,1537]" captionStart="Figure 13" captionStartId="24.[129,194,1924,1948]" captionTargetBox="[132,1470,1394,1896]" captionTargetId="figure-744@24.[129,1473,1388,1897]" captionTargetPageId="24" captionText="Figure 13. Dorsolateral and ventral variation of two male specimens [RMCA R.8575/A (left) and RMCA R.8575/B (right)] of L. depressus, collected in sympatry at Flandria, Équateur Province, DRC." pageId="27" pageNumber="474">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCCFFE4FFB34658FD2FF942" pageId="27" pageNumber="474" type="distribution">
<paragraph id="8BA5B843FFCCFFE4FFB34658FD2FF942" blockId="27.[113,763,1570,1782]" pageId="27" pageNumber="474">
<emphasis id="B96E6451FFCCFFE4FFB34658FEEDF98E" box="[113,353,1570,1594]" italics="true" pageId="27" pageNumber="474">Habitat and distribution</emphasis>
(
<figureCitation id="1321A4C6FFCCFFE4FEBA4658FE39F98E" box="[376,437,1570,1594]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="27" pageNumber="474">Figs 6</figureCitation>
,
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): This species is widely distributed within the
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Basin, from the Ituri Forest in the north-east, through the heart of the
<collectingCountry id="F30DF8D3FFCCFFE4FE1F461BFDAFF9CD" box="[477,547,1633,1657]" name="Democratic Republic of the Congo" pageId="27" pageNumber="474">Congo</collectingCountry>
River at Npenda and Lake Tumba, south to
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at the southernmost extreme of its range. The material from
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was collected from mangroves growing at the mouth of the Cuanza River. However, all material from
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was collected in dense Congolian rainforest.
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<emphasis id="B96E6451FFCCFFE4FFB3476DFE9BF89B" box="[113,279,1815,1839]" italics="true" pageId="27" pageNumber="474">Natural history:</emphasis>
Individuals in
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were observed actively moving and hunting between the branches and trunk of trees during the day, but due to the difficult access to the mangroves, specimens were collected at night while sleeping on thin branches. Specimens from Lotende,
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, were collected in the canopy, as recorded by Laurent in his field notes.
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<emphasis id="B96E6451FFC2FFEAFF0C470DFD99F825" bold="true" box="[206,533,1911,1937]" italics="true" pageId="21" pageNumber="468">LYGODACTYLUS DYSMICUS</emphasis>
<bibRefCitation id="EF8BC5B2FFC2FFEAFDD8470DFD11F825" author="Perret JL" box="[538,669,1911,1937]" pageId="21" pageNumber="468" pagination="47 - 60" refId="ref36360" refString="Perret JL. Les Gekkonidae du Cameroun, avec la description de deux sous - especes nouvelles. Revue Suisse de Zoologie 1963; 70: 47 - 60. https: // doi. org / 10.5962 / bhl. part. 117919" type="journal article" year="1963">Perret, 1963</bibRefCitation>
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(
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<taxonomicName id="4C1AC3C0FFC2FFEAFCB9462BFB41F9DF" authorityName="Perret" authorityYear="1963" box="[891,1229,1617,1643]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="468" phylum="Chordata" rank="subSpecies" species="angularis" subSpecies="dysmicus">Lygodactylus angularis dysmicus</taxonomicName>
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<bibRefCitation id="EF8BC5B2FFC2FFEAFB1A4628FAD1F9DF" author="Perret JL" box="[1240,1373,1617,1643]" pageId="21" pageNumber="468" pagination="47 - 60" refId="ref36360" refString="Perret JL. Les Gekkonidae du Cameroun, avec la description de deux sous - especes nouvelles. Revue Suisse de Zoologie 1963; 70: 47 - 60. https: // doi. org / 10.5962 / bhl. part. 117919" type="journal article" year="1963">Perret (1963)</bibRefCitation>
.
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<emphasis id="B96E6451FFC2FFEAFCF346EAFAF5F95D" italics="true" pageId="21" pageNumber="468">
<taxonomicName id="4C1AC3C0FFC2FFEAFCF346EAFB08F91E" ID-CoL="7HDH7" authorityName="Perret" authorityYear="1963" box="[817,1156,1680,1706]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="468" phylum="Chordata" rank="subSpecies" species="gutturalis" subSpecies="dysmicus">Lygodactylus gutturalis dysmicus</taxonomicName>
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<bibRefCitation id="EF8BC5B2FFC2FFEAFB4D46EBFAAEF91E" author="Pasteur G" box="[1167,1314,1680,1706]" pageId="21" pageNumber="468" pagination="1 - 132" refId="ref36328" refString="Pasteur G. Recherches sur l'evolution des lygodactyles, lezards Afro- Malgaches actuels. Travaux de l'Institut Scientifique Cherifien, Zoologie 1965 (1964); 29: 1 - 132." type="journal article" year="1964">Pasteur (1964)</bibRefCitation>
;
<bibRefCitation id="EF8BC5B2FFC2FFEAFAEF46EAFA24F91E" author="Kluge AG" box="[1325,1448,1680,1706]" pageId="21" pageNumber="468" pagination="1 - 35" refId="ref35045" refString="Kluge AG. Checklist of gekkonid lizards. Smithsonian Herpetological Information Service 1991; 85: 1 - 35. https: // doi. org / 10.5479 / si. 23317515.85.1" type="journal article" year="1991">Kluge (1991</bibRefCitation>
,
<bibRefCitation id="EF8BC5B2FFC2FFEAFCF146D5FCE4F97D" author="Kluge AG" box="[819,872,1711,1737]" pageId="21" pageNumber="468" refId="ref35077" refString="Kluge AG. Gekkonoid Lizard Taxonomy. San Diego: International Gecko Society, 1993." type="book" year="1993">1993</bibRefCitation>
,
<bibRefCitation id="EF8BC5B2FFC2FFEAFCB246D5FC29F97D" author="Kluge AG" box="[880,933,1711,1737]" pageId="21" pageNumber="468" pagination="1 - 209" refId="ref35093" refString="Kluge AG. Gekkotan lizard taxonomy. Hamadryad 2001; 26: 1 - 209." type="journal article" year="2001">2001</bibRefCitation>
);
<bibRefCitation id="EF8BC5B2FFC2FFEAFC7946D5FB23F97D" author="Schatti B &amp; Perret J" box="[955,1199,1711,1737]" pageId="21" pageNumber="468" pagination="357 - 70" refId="ref37084" refString="Schatti B, Perret J. Catalogue revise des types d'amphibiens et de reptiles du Museum d'histoire naturelle de Geneve. Revue Suisse de Zoologie 1997; 104: 357 - 70." type="journal article" year="1997">Schätti and Perret (1997)</bibRefCitation>
;
<bibRefCitation id="EF8BC5B2FFC2FFEAFB7846CAFAD2F97D" author="LeBreton M" box="[1210,1374,1711,1737]" pageId="21" pageNumber="468" refId="ref35386" refString="LeBreton M. A Working Checklist of the Herpetofauna of Cameroon. Amsterdam: UICN: Netherlands Committee for IUCN, 1999." type="book" year="1999">LeBreton (1999)</bibRefCitation>
;
<bibRefCitation id="EF8BC5B2FFC2FFEAFAA846D5FC6FF95D" author="Schatti B &amp; Perret J &amp; Mariaux J" pageId="21" pageNumber="468" refId="ref37117" refString="Schatti B, Perret J, Mariaux J. 2002. Catalogue commente des Types d'Amphibiens et de Reptiles du Museum d'Histoire Naturelle de Geneve. Geneve: Museum d'histoire naturelle Geneve. Available from: http: // www. ville-ge. ch / mhng / erpi / cat 1. html [accessed 10 August 2022]" type="url" year="2002">Schätti et al. (2002)</bibRefCitation>
;
<bibRefCitation id="EF8BC5B2FFC2FFEAFC2C46B4FBEFF95D" author="Roll B" box="[1006,1123,1742,1769]" pageId="21" pageNumber="468" pagination="645 - 8" refId="ref36800" refString="Roll B. Lygodactylus gutturalis (Bocage). Sauria (Suppl.) 2005; 27: 645 - 8." type="journal article" year="2005">Röll (2005)</bibRefCitation>
; de Lisle et al. (2013, 2016).
</emphasis>
</paragraph>
<paragraph id="8BA5B843FFC2FFEAFCEE477CFA3DF894" blockId="21.[812,1457,1798,1824]" box="[812,1457,1798,1824]" pageId="21" pageNumber="468">
<emphasis id="B96E6451FFC2FFEAFCEE477CFA3DF894" box="[812,1457,1798,1824]" italics="true" pageId="21" pageNumber="468">
<taxonomicName id="4C1AC3C0FFC2FFEAFCEE477CFA90F894" ID-CoL="7HDH7" authorityName="Perret" authorityYear="1963" box="[812,1308,1798,1824]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="468" phylum="Chordata" rank="subSpecies" species="gutturalis" subGenus="Lygodactylus" subSpecies="dysmicus">Lygodactylus (Lygodactylus) gutturalis dysmicus</taxonomicName>
:
<bibRefCitation id="EF8BC5B2FFC2FFEAFAE5477CFA21F894" author="Rosler H" box="[1319,1453,1798,1824]" pageId="21" pageNumber="468" pagination="28 - 153" refId="ref36972" refString="Rosler H. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2000; 2: 28 - 153." type="journal article" year="2000">Rösler (2000)</bibRefCitation>
.
</emphasis>
</paragraph>
<paragraph id="8BA5B843FFC2FFEAFCA8473AFAFFF8EE" blockId="21.[874,1395,1856,1882]" box="[874,1395,1856,1882]" pageId="21" pageNumber="468">
<emphasis id="B96E6451FFC2FFEAFCA8473AFAFFF8EE" box="[874,1395,1856,1882]" italics="true" pageId="21" pageNumber="468">
<taxonomicName id="4C1AC3C0FFC2FFEAFCA8473AFBD9F8EE" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[874,1109,1856,1882]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="468" phylum="Chordata" rank="species" species="dysmicus">Lygodactylus dysmicus</taxonomicName>
: Chirio and LeBreton (2007).
</emphasis>
</paragraph>
<paragraph id="8BA5B843FFC2FFEAFCF0470DFB19F804" blockId="21.[818,1451,1911,1968]" pageId="21" pageNumber="468">
<emphasis id="B96E6451FFC2FFEAFCF0470DFB19F804" italics="true" pageId="21" pageNumber="468">
<taxonomicName id="4C1AC3C0FFC2FFEAFCF0470DFB07F825" authority="(Perret, 1963)" box="[818,1163,1911,1937]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="468" phylum="Chordata" rank="subSpecies" species="picturatus" subSpecies="dysmicus">Lygodactylus picturatus dysmicus</taxonomicName>
: Thys van den
<bibRefCitation id="EF8BC5B2FFC2FFEAFAD7470DFB1DF804" author="Thys van den Audenaerde DFE" pageId="21" pageNumber="468" pagination="163 - 77" refId="ref37501" refString="Thys van den Audenaerde DFE. Les Gekkonidae de l'Afrique centrale. Revue de Zoologie et de Botanique Africaines 1967; 76: 163 - 77." type="journal article" year="1967">den Audenaerde (1967)</bibRefCitation>
.
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF65E8CBFFC1FFE9FF4244C9FB66FAB7" pageId="22" pageNumber="469" startId="22.[128,193,1203,1227]" targetBox="[129,1473,144,1175]" targetPageId="22" targetType="figure">
<paragraph id="8BA5B843FFC1FFE9FF4244C9FB66FAB7" blockId="22.[128,1474,1203,1284]" pageId="22" pageNumber="469">
<emphasis id="B96E6451FFC1FFE9FF4244C9FF68FB7F" bold="true" box="[128,228,1203,1227]" pageId="22" pageNumber="469">Figure 11.</emphasis>
Specimens of
<taxonomicName id="4C1AC3C0FFC1FFE9FEAE44CEFE54FB7F" box="[364,472,1203,1227]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC1FFE9FEAE44CEFE54FB7F" box="[364,472,1203,1227]" italics="true" pageId="22" pageNumber="469">L. gutturalis</emphasis>
</taxonomicName>
in life from (A–C) Guinea Bissau and (D–E) Senegal. Photos of habitat at (F) Sambatchur, Bafata Region, Guinea Bissau, (G) Kalan Kalan, Kankan Region, Guinea, (H) Guiguisso, Donga Department, Benin, (I) Nsonsomea, Sunyani Region, Ghana, and (J) Mlomp, Zinguinchor Region, Senegal. Photographs by J.F.T. (A–C and F–J) and Alberto Sanchez-Vialas (D–E).
</paragraph>
</caption>
<subSubSection id="C300EBC8FFC1FFE9FF434534FC56FA32" pageId="22" pageNumber="469" type="discussion">
<paragraph id="8BA5B843FFC1FFE9FF434534FC56FA32" blockId="22.[129,778,1358,1977]" lastBlockId="22.[825,1475,1358,1414]" pageId="22" pageNumber="469">
<bibRefCitation id="EF8BC5B2FFC1FFE9FF434534FE98FAD2" author="Perret JL" box="[129,276,1358,1382]" pageId="22" pageNumber="469" pagination="47 - 60" refId="ref36360" refString="Perret JL. Les Gekkonidae du Cameroun, avec la description de deux sous - especes nouvelles. Revue Suisse de Zoologie 1963; 70: 47 - 60. https: // doi. org / 10.5962 / bhl. part. 117919" type="journal article" year="1963">Perret (1963)</bibRefCitation>
described
<emphasis id="B96E6451FFC1FFE9FE4F4535FDACFAD2" box="[397,544,1358,1382]" italics="true" pageId="22" pageNumber="469">
L. a.
<taxonomicName id="4C1AC3C0FFC1FFE9FE0B4534FDACFAD2" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[457,544,1358,1382]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="dysmicus">dysmicus</taxonomicName>
</emphasis>
from Foulassi, South Region,
<collectingCountry id="F30DF8D3FFC1FFE9FF154517FECBFA31" box="[215,327,1389,1413]" name="Cameroon" pageId="22" pageNumber="469">Cameroon</collectingCountry>
, as the western race of
<taxonomicName id="4C1AC3C0FFC1FFE9FDF84514FD23FA31" authorityName="Gunther" authorityYear="1893" box="[570,687,1389,1413]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFC1FFE9FDF84514FD23FA31" box="[570,687,1389,1413]" italics="true" pageId="22" pageNumber="469">L. angularis</emphasis>
</taxonomicName>
that differs from the nominotypical form by its gular ornamentation. However, Perret never compared this species to
<taxonomicName id="4C1AC3C0FFC1FFE9FDB445D7FD61FA70" box="[630,749,1452,1476]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC1FFE9FDB445D7FD61FA70" box="[630,749,1452,1476]" italics="true" pageId="22" pageNumber="469">L. gutturalis</emphasis>
</taxonomicName>
in neighbouring areas. In 1991, Kluge considered
<emphasis id="B96E6451FFC1FFE9FDA745B6FD63FA57" box="[613,751,1483,1507]" italics="true" pageId="22" pageNumber="469">
L. a.
<taxonomicName id="4C1AC3C0FFC1FFE9FD5A45B1FD63FA57" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[664,751,1483,1507]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="dysmicus">dysmicus</taxonomicName>
</emphasis>
as a subspecies of
<taxonomicName id="4C1AC3C0FFC1FFE9FEDE4591FE1CF9B6" box="[284,400,1514,1538]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC1FFE9FEDE4591FE1CF9B6" box="[284,400,1514,1538]" italics="true" pageId="22" pageNumber="469">L. gutturalis</emphasis>
</taxonomicName>
, although he did not provide any justification. This taxonomic change was followed by other authors until Chirio and LeBreton (2007) considered
<taxonomicName id="4C1AC3C0FFC1FFE9FDA44650FD56F9F5" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[614,730,1577,1601]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC1FFE9FDA44650FD56F9F5" box="[614,730,1577,1601]" italics="true" pageId="22" pageNumber="469">L. dysmicus</emphasis>
</taxonomicName>
as a full species in their ‘
<emphasis id="B96E6451FFC1FFE9FE8B4632FDFFF9D4" box="[329,627,1608,1632]" italics="true" pageId="22" pageNumber="469">
<taxonomicName id="4C1AC3C0FFC1FFE9FE8B4632FE65F9D4" authority="des reptiles" authorityName="des Reptiles" box="[329,489,1608,1632]" class="Reptilia" family="Cetoniidae" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="490" phylum="Arthropoda" rank="genus">Atlas des reptiles</taxonomicName>
du
<collectingCountry id="F30DF8D3FFC1FFE9FDCE4633FDFFF9D4" box="[524,627,1609,1632]" name="Cameroon" pageId="22" pageNumber="469">Cameroun</collectingCountry>
</emphasis>
’ again without justification. Until then, the only known record of this species was the
<typeStatus id="54A106E1FFC1FFE9FF1146FDFEBFF92B" box="[211,307,1671,1695]" pageId="22" pageNumber="469" type="holotype">holotype</typeStatus>
; however, Chirio and LeBreton (2007) added a new photographic record from Nglochifen, West Region,
<collectingCountry id="F30DF8D3FFC1FFE9FF4346BCFF7DF96A" box="[129,241,1734,1758]" name="Cameroon" pageId="22" pageNumber="469">Cameroon</collectingCountry>
, in their book. Herein, based on the osteological evidence mentioned above, and the presence of terminal scansors on the tail tip, which are absent in the
<taxonomicName id="4C1AC3C0FFC1FFE9FDCB477FFDF1F8A8" authorityName="Gunther" authorityYear="1893" box="[521,637,1796,1820]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFC1FFE9FDCB477FFDF1F8A8" box="[521,637,1796,1820]" italics="true" pageId="22" pageNumber="469">L. angularis</emphasis>
</taxonomicName>
group, we regard
<taxonomicName id="4C1AC3C0FFC1FFE9FF71475FFEA8F888" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[179,292,1828,1852]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC1FFE9FF71475FFEA8F888" box="[179,292,1828,1852]" italics="true" pageId="22" pageNumber="469">L. dysmicus</emphasis>
</taxonomicName>
to be part of the
<taxonomicName id="4C1AC3C0FFC1FFE9FE16475FFDC5F888" box="[468,585,1828,1852]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC1FFE9FE16475FFDC5F888" box="[468,585,1828,1852]" italics="true" pageId="22" pageNumber="469">L. gutturalis</emphasis>
</taxonomicName>
subgroup. Despite the lack of molecular data, distinctive morphology supports the validation of
<taxonomicName id="4C1AC3C0FFC1FFE9FEF44719FE27F8CE" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[310,427,1890,1914]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC1FFE9FEF44719FE27F8CE" box="[310,427,1890,1914]" italics="true" pageId="22" pageNumber="469">L. dysmicus</emphasis>
</taxonomicName>
as a full species, as suggested by Chirio and LeBreton (2007) and
<bibRefCitation id="EF8BC5B2FFC1FFE9FE2647F8FDFEF82E" author="Kluge AG" box="[484,626,1922,1946]" pageId="22" pageNumber="469" pagination="1 - 35" refId="ref35045" refString="Kluge AG. Checklist of gekkonid lizards. Smithsonian Herpetological Information Service 1991; 85: 1 - 35. https: // doi. org / 10.5479 / si. 23317515.85.1" type="journal article" year="1991">Kluge (1991)</bibRefCitation>
. Based on the detailed description provided by
<bibRefCitation id="EF8BC5B2FFC1FFE9FE2E47D8FD0CF80D" author="Perret JL" box="[492,640,1953,1977]" pageId="22" pageNumber="469" pagination="47 - 60" refId="ref36360" refString="Perret JL. Les Gekkonidae du Cameroun, avec la description de deux sous - especes nouvelles. Revue Suisse de Zoologie 1963; 70: 47 - 60. https: // doi. org / 10.5962 / bhl. part. 117919" type="journal article" year="1963">Perret (1963)</bibRefCitation>
, we provide only a diagnosis and comparison with
<emphasis id="B96E6451FFC1FFE9FB024535FADBFAD2" box="[1216,1367,1358,1382]" italics="true" pageId="22" pageNumber="469">
<taxonomicName id="4C1AC3C0FFC1FFE9FB024535FAB9FAD2" box="[1216,1333,1358,1382]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="gutturalis">L. gutturalis</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFC1FFE9FAF94535FADBFAD2" box="[1339,1367,1359,1382]" pageId="22" pageNumber="469" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
and the
<taxonomicName id="4C1AC3C0FFC1FFE9FA6C4535FC1FFA31" authorityName="Gunther" authorityYear="1893" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFC1FFE9FA6C4535FC1FFA31" italics="true" pageId="22" pageNumber="469">L. angularis</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFC1FFE9FCFB45D6FBCDF996" pageId="22" pageNumber="469" type="materials_examined">
<paragraph id="8BA5B843FFC1FFE9FCFB45D6FBCDF996" blockId="22.[825,1474,1452,1570]" pageId="22" pageNumber="469">
<materialsCitation id="3B72B21EFFC1FFE9FCFB45D6FBCDF996" collectionCode="MHNG" country="Cameroon" elevation="665" location="Foulassi" municipality="South Region" pageId="22" pageNumber="469" specimenCode="MHNG 1005.72" specimenCount="1" typeStatus="holotype">
<emphasis id="B96E6451FFC1FFE9FCFB45D6FC15FA70" box="[825,921,1452,1476]" italics="true" pageId="22" pageNumber="469">
<typeStatus id="54A106E1FFC1FFE9FCFB45D6FC18FA70" box="[825,916,1452,1476]" pageId="22" pageNumber="469" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DBBC1038FFC1FFE9FC6E45D7FBD1FA70" box="[940,1117,1452,1477]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="22" pageNumber="469" type="Museum">MHNG 1005.72</specimenCode>
, adult male, collected from forest at
<location id="8EC5EE98FFC1FFE9FC9445B6FC27FA50" LSID="urn:lsid:plazi:treatment:03B30955FFC2FFE8FF0C470DFDDEF8D6:8EC5EE98FFC1FFE9FC9445B6FC27FA50" box="[854,939,1484,1508]" country="Cameroon" municipality="South Region" name="Foulassi" pageId="22" pageNumber="469">Foulassi</location>
,
<collectingMunicipality id="6BC12239FFC1FFE9FC7B45B6FBC1FA50" box="[953,1101,1484,1508]" pageId="22" pageNumber="469">South Region</collectingMunicipality>
,
<collectingCountry id="F30DF8D3FFC1FFE9FB9945B6FB45FA50" box="[1115,1225,1484,1508]" name="Cameroon" pageId="22" pageNumber="469">Cameroon</collectingCountry>
(~
<elevation id="00375F70FFC1FFE9FB2B45B6FAE7FA50" box="[1257,1387,1484,1508]" metricMagnitude="2" metricUnit="m" metricValue="6.65" pageId="22" pageNumber="469" unit="m" value="665.0">
<quantity id="4CE215A6FFC1FFE9FB2B45B6FABFFA57" box="[1257,1331,1484,1508]" metricMagnitude="2" metricUnit="m" metricValue="6.65" pageId="22" pageNumber="469" unit="m" value="665.0">665 m</quantity>
a.s.l.
</elevation>
), at the north-west limit of the Congo Basin. Specimen fixed in formalin, now preserved in ethanol.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFC1FFE8FCFB4630FE23F987" lastPageId="23" lastPageNumber="470" pageId="22" pageNumber="469" type="diagnosis">
<paragraph id="8BA5B843FFC1FFE8FCFB4630FE23F987" blockId="22.[825,1475,1610,1978]" lastBlockId="23.[113,763,1250,1587]" lastPageId="23" lastPageNumber="470" pageId="22" pageNumber="469">
<emphasis id="B96E6451FFC1FFE9FCFB4630FC15F9D5" box="[825,921,1610,1633]" italics="true" pageId="22" pageNumber="469">Diagnosis</emphasis>
(
<figureCitation id="1321A4C6FFC1FFE9FC6D4630FC76F9D6" box="[943,1018,1610,1634]" captionStart="Figure 12" captionStartId="23.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-438@23.[114,1458,144,1094]" captionTargetPageId="23" captionText="Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R." pageId="22" pageNumber="469">Fig. 12</figureCitation>
): As noted by
<bibRefCitation id="EF8BC5B2FFC1FFE9FB6A4630FAB0F9D6" author="Perret JL" box="[1192,1340,1610,1634]" pageId="22" pageNumber="469" pagination="47 - 60" refId="ref36360" refString="Perret JL. Les Gekkonidae du Cameroun, avec la description de deux sous - especes nouvelles. Revue Suisse de Zoologie 1963; 70: 47 - 60. https: // doi. org / 10.5962 / bhl. part. 117919" type="journal article" year="1963">Perret (1963)</bibRefCitation>
, the species differs from the
<taxonomicName id="4C1AC3C0FFC1FFE9FC2B4610FBEDF935" authorityName="Gunther" authorityYear="1893" box="[1001,1121,1641,1665]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFC1FFE9FC2B4610FBEDF935" box="[1001,1121,1641,1665]" italics="true" pageId="22" pageNumber="469">L. angularis</emphasis>
</taxonomicName>
group mainly by the gular ornamentation, which consists of two
<emphasis id="B96E6451FFC1FFE9FB7946F2FB40F914" bold="true" box="[1211,1228,1672,1696]" pageId="22" pageNumber="469">∩</emphasis>
-shaped chevrons, one within the other, and a central mark vs. two
<emphasis id="B96E6451FFC1FFE9FB3D46D2FA83F90B" box="[1279,1295,1704,1727]" italics="true" pageId="22" pageNumber="469">V</emphasis>
-shaped marks or broken pattern (in
<taxonomicName id="4C1AC3C0FFC1FFE9FC3D46B2FBEFF96B" box="[1023,1123,1735,1759]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="heeneni">
<emphasis id="B96E6451FFC1FFE9FC3D46B2FBEFF96B" box="[1023,1123,1735,1759]" italics="true" pageId="22" pageNumber="469">L. heeneni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1AC3C0FFC1FFE9FB5A46B2FAAFF96B" baseAuthorityName="Laurent" baseAuthorityYear="1952" box="[1176,1315,1735,1759]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="paurospilus">
<emphasis id="B96E6451FFC1FFE9FB5A46B2FAAFF96B" box="[1176,1315,1735,1759]" italics="true" pageId="22" pageNumber="469">L. paurospilus</emphasis>
</taxonomicName>
) chevrons that converge posteriorly. It also differs from the
<taxonomicName id="4C1AC3C0FFC1FFE9FADD469CFAF6F94A" authorityName="Gunther" authorityYear="1893" box="[1311,1402,1766,1790]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFC1FFE9FADD469CFAF6F94A" box="[1311,1402,1766,1790]" italics="true" pageId="22" pageNumber="469">angularis</emphasis>
</taxonomicName>
group by having a reduced, almost vestigial postorbitofrontal vs. a well-developed postorbitofrontal bone in the
<taxonomicName id="4C1AC3C0FFC1FFE9FADF475FFAF4F889" authorityName="Gunther" authorityYear="1893" box="[1309,1400,1829,1853]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFC1FFE9FADF475FFAF4F889" box="[1309,1400,1829,1853]" italics="true" pageId="22" pageNumber="469">angularis</emphasis>
</taxonomicName>
group.
<taxonomicName id="4C1AC3C0FFC1FFE9FCFB473EFB90F8E8" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[825,1052,1860,1884]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC1FFE9FCFB473EFB90F8E8" box="[825,1052,1860,1884]" italics="true" pageId="22" pageNumber="469">Lygodactylus dysmicus</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C1AC3C0FFC1FFE9FA85473FFA4EF8E8" box="[1351,1474,1860,1884]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC1FFE9FA85473FFA4EF8E8" box="[1351,1474,1860,1884]" italics="true" pageId="22" pageNumber="469">L. gutturalis</emphasis>
</taxonomicName>
by having more precloacal pores (9 vs. 5–8); nostril in narrow contact with rostral scales vs. never in contact in
<taxonomicName id="4C1AC3C0FFC1FFE9FA8147FEFA31F82F" box="[1347,1469,1923,1947]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="469" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC1FFE9FA8147FEFA31F82F" box="[1347,1469,1923,1947]" italics="true" pageId="22" pageNumber="469">L. gutturalis</emphasis>
</taxonomicName>
. We have also recorded some minor differences as follows: fewer lamellae under fourth toe (4 vs. 5–6); a greater number of ventral scales across the body (21 vs. 15–20); and a greater number of scales between the eyes (25 vs. 17–24). It is noteworthy that
<taxonomicName id="4C1AC3C0FFC0FFE8FFB3453BFF69FAEC" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[113,229,1344,1368]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC0FFE8FFB3453BFF69FAEC" box="[113,229,1344,1368]" italics="true" pageId="23" pageNumber="470">L. dysmicus</emphasis>
</taxonomicName>
is small [SVL
<quantity id="4CE215A6FFC0FFE8FE40453AFE6CFAE3" box="[386,480,1344,1368]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="23" pageNumber="470" unit="mm" value="27.6">27.6 mm</quantity>
vs. 33.3 ±
<quantity id="4CE215A6FFC0FFE8FD97453AFD2AFAE3" box="[597,678,1344,1368]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="470" unit="mm" value="1.8">1.8 mm</quantity>
(mean) in
<taxonomicName id="4C1AC3C0FFC0FFE8FF4E451AFE8FFAC3" box="[140,259,1375,1399]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC0FFE8FF4E451AFE8FFAC3" box="[140,259,1375,1399]" italics="true" pageId="23" pageNumber="470">L. gutturalis</emphasis>
</taxonomicName>
], with a paedomorphic skull that has an unfused frontal bone, braincase, and compound bone + surangular (
<figureCitation id="1321A4C6FFC0FFE8FD164505FFF2FA02" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." pageId="23" pageNumber="470">Fig. 5</figureCitation>
). However, the well-developed state of the precloacal pores suggest sexual maturity of this specimen (
<bibRefCitation id="EF8BC5B2FFC0FFE8FDE245C7FD4AFA61" author="Rhen T &amp; Sakata JT &amp; Crews D" box="[544,710,1469,1493]" pageId="23" pageNumber="470" pagination="289 - 96" refId="ref36740" refString="Rhen T, Sakata JT, Crews D. Effects of gonadal sex and incubation temperature on the ontogeny of gonadal steroid concentrations and secondary sex structures in leopard geckos, Eublepharis macularius. General and Comparative Endocrinology 2005; 142: 289 - 96. https: // doi. org / 10.1016 / j. ygcen. 2005.01.018" type="journal article" year="2005">
Rhen
<emphasis id="B96E6451FFC0FFE8FD9C45C4FD01FA61" box="[606,653,1469,1493]" italics="true" pageId="23" pageNumber="470">et al.</emphasis>
2005
</bibRefCitation>
;
<figureCitation id="1321A4C6FFC0FFE8FD1645C7FF16FA41" captionStart="Figure 12" captionStartId="23.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-438@23.[114,1458,144,1094]" captionTargetPageId="23" captionText="Figure 12. Holotype of L. dysmicus (MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R." pageId="23" pageNumber="470">Fig. 12B</figureCitation>
). Finally,
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<emphasis id="B96E6451FFC0FFE8FEC245A7FEFDFA41" box="[256,369,1501,1525]" italics="true" pageId="23" pageNumber="470">L. dysmicus</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C1AC3C0FFC0FFE8FD4645A7FD77FA41" box="[644,763,1501,1525]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC0FFE8FD4645A7FD77FA41" box="[644,763,1501,1525]" italics="true" pageId="23" pageNumber="470">L. gutturalis</emphasis>
</taxonomicName>
based on its distribution in lowland Congolian rainforest vs. the sub-Saharan savannah (
<figureCitation id="1321A4C6FFC0FFE8FEA44661FE13F987" box="[358,415,1563,1587]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="23" pageNumber="470">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF65E8CBFFC0FFE8FFB34418FF48FB06" pageId="23" pageNumber="470" startId="23.[113,178,1122,1146]" targetBox="[114,1458,144,1094]" targetPageId="23" targetType="figure">
<paragraph id="8BA5B843FFC0FFE8FFB34418FF48FB06" blockId="23.[113,1455,1122,1202]" pageId="23" pageNumber="470">
<emphasis id="B96E6451FFC0FFE8FFB34418FF5AFBCE" bold="true" box="[113,214,1122,1146]" pageId="23" pageNumber="470">Figure 12.</emphasis>
Holotype of
<taxonomicName id="4C1AC3C0FFC0FFE8FE904419FE36FBCE" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[338,442,1122,1146]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC0FFE8FE904419FE36FBCE" box="[338,442,1122,1146]" italics="true" pageId="23" pageNumber="470">L. dysmicus</emphasis>
</taxonomicName>
(MHNG 1005.72) from Foulassi, South Region, Cameroon. A, Ventral and dorsal view of body. B, Details of cloacal region. C, Details of left pes. D, Detailed view of head in both lateral views and ventral and dorsal views (from top to bottom). Photos by J.L.R.
</paragraph>
</caption>
<subSubSection id="C300EBC8FFC0FFE8FFB3462FFDA6F8BD" pageId="23" pageNumber="470" type="distribution">
<paragraph id="8BA5B843FFC0FFE8FFB3462FFDA6F8BD" blockId="23.[113,762,1620,1801]" pageId="23" pageNumber="470">
<emphasis id="B96E6451FFC0FFE8FFB3462FFEF9F9D9" box="[113,373,1621,1645]" italics="true" pageId="23" pageNumber="470">Habitat and distribution</emphasis>
(
<figureCitation id="1321A4C6FFC0FFE8FE4D462FFE5EF9D8" box="[399,466,1620,1645]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="23" pageNumber="470">Fig. 6</figureCitation>
):
<taxonomicName id="4C1AC3C0FFC0FFE8FE36462FFD56F9D9" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[500,730,1621,1645]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC0FFE8FE36462FFD56F9D9" box="[500,730,1621,1645]" italics="true" pageId="23" pageNumber="470">Lygodactylus dysmicus</emphasis>
</taxonomicName>
is only known from the
<typeStatus id="54A106E1FFC0FFE8FE99460EFE05F938" box="[347,393,1652,1676]" pageId="23" pageNumber="470">type</typeStatus>
locality in Foulassi, South Region,
<collectingCountry id="F30DF8D3FFC0FFE8FFB346E9FF6DF91F" box="[113,225,1683,1707]" name="Cameroon" pageId="23" pageNumber="470">Cameroon</collectingCountry>
, and an additional photographic record (unconfirmed) from Nglochifen, West Region,
<collectingCountry id="F30DF8D3FFC0FFE8FDC146C9FDFFF97F" box="[515,627,1715,1739]" name="Cameroon" pageId="23" pageNumber="470">Cameroon</collectingCountry>
.
<taxonomicName id="4C1AC3C0FFC0FFE8FDB946C8FF44F95E" baseAuthorityName="Perret" baseAuthorityYear="1963" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="470" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC0FFE8FDB946C8FF44F95E" italics="true" pageId="23" pageNumber="470">Lygodactylus dysmicus</emphasis>
</taxonomicName>
is considered a rainforest species distributed near the northern limit of the Congolian Rainforest.
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFC0FFE8FFB34750FDDEF8D6" pageId="23" pageNumber="470" type="biology_ecology">
<paragraph id="8BA5B843FFC0FFE8FFB34750FDDEF8D6" blockId="23.[113,762,1834,1890]" pageId="23" pageNumber="470">
<emphasis id="B96E6451FFC0FFE8FFB34750FE82F8F6" box="[113,270,1834,1858]" italics="true" pageId="23" pageNumber="470">Natural history:</emphasis>
Diurnal gecko that lives on trees in dense semideciduous forests (Chirio and LeBreton 2007).
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="A875A5271D406888510B57461949BEBF">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="AEA1CC314442A5A8B4FBBB5DF6EB4F47">
<mods:date id="6A458B7E21C938B1C9119BEF1E8E05BB">2024</mods:date>
<mods:detail id="88D0589CD645C5D37BEA0A6B0E31E2C7" type="series">
<mods:title id="8576CD72C4F01F1022730D6456CB720F">Zool. J. Linn. Soc.</mods:title>
</mods:detail>
<mods:detail id="3C66A07A7CF7177FE24415C278580AC8" type="pubDate">
<mods:number id="9AA61541D7A048B41BAF134241E058F6">2023-09-23</mods:number>
</mods:detail>
<mods:detail id="3BECE47AAD34B6E2206F8ADF0C010412" type="volume">
<mods:number id="D911733715FF72B169352DB6ABCF5E29">201</mods:number>
</mods:detail>
<mods:detail id="62F8D121E7AC91CBC98ED36C2B04A4A3" type="issue">
<mods:number id="87035DB202BF19A0D6DE673A29D7823E">2</mods:number>
</mods:detail>
<mods:extent id="237124D60F0EC1EFD4634B204E043004" unit="page">
<mods:start id="E333FB707961C6FFFB80F0DA49BFC92D">447</mods:start>
<mods:end id="D22D24ABDFA85DA4E4CAECA5D4B42441">492</mods:end>
</mods:extent>
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<mods:location id="3BE0D1169C71BD7B2B8C91F3684EC9F5">
<mods:url id="8FE218EACD7C1FF422C23E80AFCE707E">http://dx.doi.org/10.1093/zoolinnean/zlad123</mods:url>
</mods:location>
<mods:classification id="58CD2B0304B1851997C2D4856406B86B">journal article</mods:classification>
<mods:identifier id="8B614A69CDC289202C0D88F7B9BDC639" type="DOI">10.1093/zoolinnean/zlad123</mods:identifier>
<mods:identifier id="9510DC67563D1847C4FF16FAA102BA2C" type="ISSN">0024-4082</mods:identifier>
</mods:mods>
<treatment id="03B30955FFCAFFDEFF1F46B0FD56FE68" LSID="urn:lsid:plazi:treatment:03B30955FFCAFFDEFF1F46B0FD56FE68" httpUri="http://treatment.plazi.org/id/03B30955FFCAFFDEFF1F46B0FD56FE68" lastPageId="33" lastPageNumber="480" pageId="29" pageNumber="476">
<subSubSection id="C300EBC8FFCAFFE2FF1F46B0FD1EF950" box="[221,658,1738,1764]" pageId="29" pageNumber="476" type="nomenclature">
<paragraph id="8BA5B843FFCAFFE2FF1F46B0FD1EF950" blockId="29.[221,658,1738,1764]" box="[221,658,1738,1764]" pageId="29" pageNumber="476">
<heading id="D0ED0F2FFFCAFFE2FF1F46B0FD1EF950" allCaps="true" bold="true" box="[221,658,1738,1764]" centered="true" fontSize="11" level="1" pageId="29" pageNumber="476" reason="1">
<emphasis id="B96E6451FFCAFFE2FF1F46B0FD1EF950" bold="true" box="[221,658,1738,1764]" pageId="29" pageNumber="476">
<taxonomicName id="4C1AC3C0FFCAFFE2FF1F46B0FDA7F950" ID-CoL="5HP8" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[221,555,1738,1764]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="hramoja" status="sp. nov.">
<emphasis id="B96E6451FFCAFFE2FF1F46B0FDA7F950" bold="true" box="[221,555,1738,1764]" italics="true" pageId="29" pageNumber="476">LYGODACTYLUS HRAMOJA</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFCAFFE2FDED46B0FD1EF950" box="[559,658,1738,1764]" pageId="29" pageNumber="476" rank="species">SP.NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCAFFE2FFB3468BFE16F8D5" pageId="29" pageNumber="476" type="description">
<paragraph id="8BA5B843FFCAFFE2FFB3468BFD76F8BD" blockId="29.[113,762,1777,1801]" box="[113,762,1777,1801]" pageId="29" pageNumber="476">
(
<figureCitation id="1321A4C6FFCAFFE2FFBE468BFF67F8BD" box="[124,235,1777,1801]" captionStart-0="Figure 19" captionStart-1="Figure 20" captionStartId-0="31.[115,180,1898,1922]" captionStartId-1="32.[128,193,1898,1922]" captionTargetBox-0="[114,1458,920,1870]" captionTargetBox-1="[129,1473,449,1870]" captionTargetId-0="figure-423@31.[114,1458,920,1870]" captionTargetId-1="figure-170@32.[129,1473,449,1870]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." captionText-1="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="29" pageNumber="476">Figs 19–20</figureCitation>
,
<tableCitation id="C6988DF8FFCAFFE2FF37468BFEB3F8BD" box="[245,319,1777,1801]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="29" pageNumber="476">Table 3</tableCitation>
; Supporting Information,
<figureCitation id="1321A4C6FFCAFFE2FD8B468BFD01F8BD" box="[585,653,1777,1801]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="29" pageNumber="476">Fig. S7</figureCitation>
; Table S8)
</paragraph>
<paragraph id="8BA5B843FFCAFFE2FFB34750FE16F8D5" blockId="29.[113,762,1834,1889]" pageId="29" pageNumber="476">
<emphasis id="B96E6451FFCAFFE2FFB34750FEC1F8F6" box="[113,333,1834,1858]" italics="true" pageId="29" pageNumber="476">Zoobank registration:</emphasis>
https://zoobank.org/
<uuid id="FFBC8296FFCAFFE2FDF64750FE16F8D5" pageId="29" pageNumber="476">F64BBDF9-A337- 4130-A65D-B3E2481112D4</uuid>
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCAFFE2FF5247FBFA85FAFA" pageId="29" pageNumber="476" type="reference_group">
<paragraph id="8BA5B843FFCAFFE2FF5247FBFD10F80F" blockId="29.[144,732,1921,1979]" pageId="29" pageNumber="476">
<emphasis id="B96E6451FFCAFFE2FF5247FBFD10F80F" italics="true" pageId="29" pageNumber="476">
<taxonomicName id="4C1AC3C0FFCAFFE2FF5247FBFE62F82F" ID-CoL="3WQ3T" authorityName="' (Schmidt" authorityYear="1919" box="[144,494,1921,1947]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="subSpecies" species="picturatus" subSpecies="gutturalis">Lygodactylus picturatus gutturalis</taxonomicName>
:
<bibRefCitation id="EF8BC5B2FFCAFFE2FE3B47FBFD19F82F" author="Schmidt KP" box="[505,661,1921,1947]" pageId="29" pageNumber="476" pagination="385 - 624" refId="ref37222" refString="Schmidt KP. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909 - 1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bulletin of the American Museum of Natural History 1919; 39: 385 - 624." type="journal article" year="1919">Schmidt (1919)</bibRefCitation>
[part];
<bibRefCitation id="EF8BC5B2FFCAFFE2FF1347DBFEE8F80E" author="Pasteur G" box="[209,356,1952,1978]" pageId="29" pageNumber="476" pagination="1 - 132" refId="ref36328" refString="Pasteur G. Recherches sur l'evolution des lygodactyles, lezards Afro- Malgaches actuels. Travaux de l'Institut Scientifique Cherifien, Zoologie 1965 (1964); 29: 1 - 132." type="journal article" year="1964">Pasteur (1964)</bibRefCitation>
[part];
<bibRefCitation id="EF8BC5B2FFCAFFE2FE6D47DAFDDAF80E" author="Wermuth H" box="[431,598,1952,1978]" pageId="29" pageNumber="476" pagination="1 - 246" refId="ref37733" refString="Wermuth H. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Das Tierreich 1965; 80: I - XXIII, 1 - 246." type="journal article" year="1965">Wermuth (1965)</bibRefCitation>
[part].
</emphasis>
</paragraph>
<paragraph id="8BA5B843FFCAFFE2FCEC456EFA85FAFA" blockId="29.[810,1459,1300,1645]" pageId="29" pageNumber="476">
<emphasis id="B96E6451FFCAFFE2FCEC456EFA85FAFA" italics="true" pageId="29" pageNumber="476">
<taxonomicName id="4C1AC3C0FFCAFFE2FCEC456EFB92FA9A" ID-CoL="3WQ3T" authority="(Bocage, 1873)" box="[814,1054,1300,1326]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="gutturalis">Lygodactylus gutturalis</taxonomicName>
:
<bibRefCitation id="EF8BC5B2FFCAFFE2FBEB456EFB12FA9B" author="Roll B" box="[1065,1182,1300,1327]" pageId="29" pageNumber="476" pagination="645 - 8" refId="ref36800" refString="Roll B. Lygodactylus gutturalis (Bocage). Sauria (Suppl.) 2005; 27: 645 - 8." type="journal article" year="2005">Röll (2005)</bibRefCitation>
[part];
<bibRefCitation id="EF8BC5B2FFCAFFE2FB28456EFA23FA9B" author="Spawls S &amp; Howell K &amp; Hinkel H &amp; Menegon M" box="[1258,1455,1300,1327]" pageId="29" pageNumber="476" refId="ref37334" refString="Spawls S, Howell K, Hinkel H, Menegon M. 2018. Field Guide to East African Reptiles. London: Bloomsbury." type="book" year="2018">Spawls et al. (2018)</bibRefCitation>
[part]; Behangana et al. (2020)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCAFFE2FCE84520FAE1F9D9" pageId="29" pageNumber="476" type="discussion">
<paragraph id="8BA5B843FFCAFFE2FCE84520FAE1F9D9" blockId="29.[810,1459,1300,1645]" pageId="29" pageNumber="476">
<emphasis id="B96E6451FFCAFFE2FCE84520FB0DFAC6" bold="true" box="[810,1153,1370,1394]" pageId="29" pageNumber="476">
<taxonomicName id="4C1AC3C0FFCAFFE2FCE84520FBA8FAC6" ID-CoL="C7JCW" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[810,1060,1370,1394]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFCAFFE2FCE84520FBA8FAC6" bold="true" box="[810,1060,1370,1394]" italics="true" pageId="29" pageNumber="476">Lygodactylus karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFCAFFE2FBEE4520FB0DFAC6" box="[1068,1153,1370,1394]" pageId="29" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from the northern Albertine Rift in the Lendu Plateau of
<collectingCountry id="F30DF8D3FFCAFFE2FB014500FB75FA26" box="[1219,1273,1402,1426]" name="Democratic Republic of the Congo" pageId="29" pageNumber="476">DRC</collectingCountry>
and several highlands of
<collectingCountry id="F30DF8D3FFCAFFE2FC4145E3FC59FA05" box="[899,981,1433,1457]" name="Uganda" pageId="29" pageNumber="476">Uganda</collectingCountry>
, and it is the sister taxon to
<emphasis id="B96E6451FFCAFFE2FB3845E3FA3FFA05" bold="true" box="[1274,1459,1432,1457]" pageId="29" pageNumber="476">
<taxonomicName id="4C1AC3C0FFCAFFE2FB3845E3FAD6FA04" ID-CoL="C7JCX" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1274,1370,1432,1456]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFCAFFE2FB3845E3FAD6FA04" bold="true" box="[1274,1370,1432,1456]" italics="true" pageId="29" pageNumber="476">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFCAFFE2FAA245E3FA3FFA05" box="[1376,1459,1433,1457]" pageId="29" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in clade (B1), from which it differs by
<emphasis id="B96E6451FFCAFFE2FB6F45C3FB3AFA64" box="[1197,1206,1465,1488]" italics="true" pageId="29" pageNumber="476">c</emphasis>
. 6.07% for the
<emphasis id="B96E6451FFCAFFE2FA8D45C2FAFAFA64" box="[1359,1398,1464,1488]" italics="true" pageId="29" pageNumber="476">16S</emphasis>
mitochondrial gene (Table 2) and a lack of nuclear haplotype sharing in
<emphasis id="B96E6451FFCAFFE2FC86458DFC0DF9BB" box="[836,897,1527,1551]" italics="true" pageId="29" pageNumber="476">RAG1</emphasis>
(
<figureCitation id="1321A4C6FFCAFFE2FC50458DFC77F9BB" box="[914,1019,1527,1551]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability&gt; 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="29" pageNumber="476">Fig. 2B–C</figureCitation>
). The new species differs from
<taxonomicName id="4C1AC3C0FFCAFFE2FAFF4582FA3EF9BB" ID-CoL="3WQ3T" box="[1341,1458,1527,1551]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCAFFE2FAFF4582FA3EF9BB" box="[1341,1458,1527,1551]" italics="true" pageId="29" pageNumber="476">L. gutturalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1AC3C0FFCAFFE2FC9A466DFC42F99A" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[856,974,1558,1582]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCAFFE2FC9A466DFC42F99A" box="[856,974,1558,1582]" italics="true" pageId="29" pageNumber="476">L. depressus</emphasis>
</taxonomicName>
by
<emphasis id="B96E6451FFCAFFE2FC34466DFC73F99A" box="[1014,1023,1559,1582]" italics="true" pageId="29" pageNumber="476">c</emphasis>
. 12.49% and 10.45%, respectively, for the
<emphasis id="B96E6451FFCAFFE2FCE8464FFCDDF9F9" box="[810,849,1589,1613]" italics="true" pageId="29" pageNumber="476">16S</emphasis>
(uncorrected p-distance) mitochondrial gene (Table 2) and a lack of nuclear haplotype sharing in
<emphasis id="B96E6451FFCAFFE2FB6A462FFB69F9D9" box="[1192,1253,1621,1645]" italics="true" pageId="29" pageNumber="476">RAG1</emphasis>
(
<figureCitation id="1321A4C6FFCAFFE2FB37462FFAD0F9D9" box="[1269,1372,1621,1645]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability&gt; 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="29" pageNumber="476">Fig. 2B–C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCAFFE1FCE846F4FD35FA0C" lastPageId="30" lastPageNumber="477" pageId="29" pageNumber="476" type="materials_examined">
<paragraph id="8BA5B843FFCAFFE2FCE846F4FB5BF897" blockId="29.[810,1459,1678,1827]" pageId="29" pageNumber="476">
<materialsCitation id="3B72B21EFFCAFFE2FCE846F4FB5BF897" collectingDate="2015-07-05" collectionCode="UTEP, DFH" collectorName="Daniel F. Hughes &amp; Wilber Lukwago &amp; Mathias Behangana." country="Uganda" county="Northern Region" elevation="1193" latitude="3.83156" location="Agoro Town" longLatPrecision="1" longitude="33.01974" municipality="Imatong Foothills" pageId="29" pageNumber="476" specimenCode="UTEP 22590, DFH 593" specimenCount="1" typeStatus="holotype">
<emphasis id="B96E6451FFCAFFE2FCE846F4FC06F912" box="[810,906,1678,1702]" italics="true" pageId="29" pageNumber="476">
<typeStatus id="54A106E1FFCAFFE2FCE846F4FC09F912" box="[810,901,1678,1702]" pageId="29" pageNumber="476" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DBBC1038FFCAFFE2FC5C46F5FBA0F912" box="[926,1068,1678,1702]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="476" type="Museum">UTEP 22590</specimenCode>
(
<specimenCode id="DBBC1038FFCAFFE2FBFF46F4FB20F912" box="[1085,1196,1678,1702]" collectionCode="DFH" pageId="29" pageNumber="476">DFH 593</specimenCode>
), male with original tail, collected on a large tree at the edge of a village at
<location id="8EC5EE98FFCAFFE2FAE946D4FA23F971" LSID="urn:lsid:plazi:treatment:03B30955FFCAFFDEFF1F46B0FD56FE68:8EC5EE98FFCAFFE2FAE946D4FA23F971" box="[1323,1455,1710,1734]" country="Uganda" county="Northern Region" latitude="3.83156" longLatPrecision="1" longitude="33.01974" municipality="Imatong Foothills" name="Agoro Town" pageId="29" pageNumber="476">Agoro Town</location>
,
<collectingMunicipality id="6BC12239FFCAFFE2FCE846B7FC61F951" box="[810,1005,1741,1765]" pageId="29" pageNumber="476">Imatong Foothills</collectingMunicipality>
,
<collectingCounty id="62C4C0CFFFCAFFE2FBC346B7FB33F951" box="[1025,1215,1741,1765]" pageId="29" pageNumber="476">Northern Region</collectingCounty>
,
<collectingCountry id="F30DF8D3FFCAFFE2FB1146B7FAA9F951" box="[1235,1317,1741,1765]" name="Uganda" pageId="29" pageNumber="476">Uganda</collectingCountry>
,
<geoCoordinate id="EE2EDE84FFCAFFE2FAFB46B7FA23F951" box="[1337,1455,1741,1765]" degrees="03.83156" direction="north" orientation="latitude" pageId="29" pageNumber="476" precision="1" value="3.83156">N03.83156</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFCAFFE2FCE84696FC10F8B0" box="[810,924,1772,1796]" degrees="33.01974" direction="east" orientation="longitude" pageId="29" pageNumber="476" precision="1" value="33.01974">E33.01974</geoCoordinate>
,
<elevation id="00375F70FFCAFFE2FC6A4696FBA0F8B0" box="[936,1068,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.193" pageId="29" pageNumber="476" unit="m" value="1193.0">
<quantity id="4CE215A6FFCAFFE2FC6A4696FC75F8B0" box="[936,1017,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.193" pageId="29" pageNumber="476" unit="m" value="1193.0">1193 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCAFFE2FB954696FB43F8B0" box="[1111,1231,1772,1796]" pageId="29" pageNumber="476" value="2015-07-05">
<collectingDate id="EFE0676BFFCAFFE2FB954696FB43F8B0" box="[1111,1231,1772,1796]" pageId="29" pageNumber="476" value="2015-07-05">5 July 2015</collectingDate>
</date>
by
<collectorName id="26EFDD95FFCAFFE2FB3A4696FA23F8B0" box="[1272,1455,1772,1796]" pageId="29" pageNumber="476">Daniel F. Hughes</collectorName>
,
<collectorName id="26EFDD95FFCAFFE2FCE84771FC58F897" box="[810,980,1803,1827]" pageId="29" pageNumber="476">Wilber Lukwago</collectorName>
, and
<collectorName id="26EFDD95FFCAFFE2FBCA4771FB5BF897" box="[1032,1239,1803,1827]" pageId="29" pageNumber="476">Mathias Behangana.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BA5B843FFCAFFE1FCE8473CFD35FA0C" blockId="29.[810,1459,1861,1979]" lastBlockId="30.[129,778,1284,1465]" lastPageId="30" lastPageNumber="477" pageId="29" pageNumber="476">
<emphasis id="B96E6451FFCAFFE2FCE8473CFBB8F8E8" box="[810,1076,1861,1885]" italics="true" pageId="29" pageNumber="476">
<typeStatus id="54A106E1FFCAFFE2FCE8473CFC05F8E9" box="[810,905,1862,1885]" pageId="29" pageNumber="476" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D1C73CAFFCAFFE2FC5E473FFBA4F8E8" box="[924,1064,1861,1884]" count="6" pageId="29" pageNumber="476" type="generic">six specimens</specimenCount>
):
</emphasis>
<materialsCitation id="3B72B21EFFCAFFE2FB89473FFCDCF8C8" collectionCode="UTEP, DFH" pageId="29" pageNumber="476" specimenCode="UTEP 22588-89, DFH 591-92" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFCAFFE2FB89473FFA8EF8E9" box="[1099,1282,1861,1885]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="476" type="Museum">UTEP 22588–89</specimenCode>
(
<specimenCode id="DBBC1038FFCAFFE2FAD7473FFA20F8E9" box="[1301,1452,1861,1885]" collectionCode="DFH" pageId="29" pageNumber="476">DFH 591–92</specimenCode>
) and
</materialsCitation>
<materialsCitation id="3B72B21EFFCAFFE2FC94471FFC5FF82F" collectingDate="2015-07-05" collectionCode="UTEP, DFH" collectorName="Daniel F. Hughes &amp; Wilber Lukwago &amp; Mathias Behangana." country="Uganda" county="Northern Region" elevation="1193" latitude="3.83156" location="Agoro Town" longLatPrecision="1" longitude="33.01974" municipality="Imatong Foothills" pageId="29" pageNumber="476" specimenCode="UTEP 22591, DFH 594" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFCAFFE2FC94471FFC6CF8C8" box="[854,992,1892,1916]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="476" type="Museum">UTEP 22591</specimenCode>
(
<specimenCode id="DBBC1038FFCAFFE2FC2F471EFBD4F8C8" box="[1005,1112,1892,1916]" collectionCode="DFH" pageId="29" pageNumber="476">DFH 594</specimenCode>
), males, with same collection data as the holotype
</materialsCitation>
;
<materialsCitation id="3B72B21EFFCAFFE1FC2247FEFD34FA8F" collectingDate="2015-07-05" collectionCode="UTEP, DFH" collectorName="Daniel F. Hughes &amp; Wilber Lukwago &amp; Mathias Behangana" country="Uganda" county="Northern Region" elevation="1153" lastPageId="30" lastPageNumber="477" latitude="3.80548" location="Agoro Town" longLatPrecision="1" longitude="32.98836" municipality="Imatong Foothills" pageId="29" pageNumber="476" specimenCode="UTEP 22592, DFH 641" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFCAFFE2FC2247FEFBE3F82F" box="[992,1135,1923,1947]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="476" type="Museum">UTEP 22592</specimenCode>
(
<specimenCode id="DBBC1038FFCAFFE2FB4247FEFB62F82F" box="[1152,1262,1923,1948]" collectionCode="DFH" pageId="29" pageNumber="476">DFH 641</specimenCode>
), female, collected at
<location id="8EC5EE98FFCAFFE2FC8747D9FC46F80E" LSID="urn:lsid:plazi:treatment:03B30955FFCAFFDEFF1F46B0FD56FE68:8EC5EE98FFCAFFE2FC8747D9FC46F80E" box="[837,970,1955,1979]" country="Uganda" county="Northern Region" latitude="3.80548" longLatPrecision="1" longitude="32.98836" municipality="Imatong Foothills" name="Agoro Town" pageId="29" pageNumber="476">Agoro Town</location>
,
<collectingMunicipality id="6BC12239FFCAFFE2FC1447D9FB1CF80F" box="[982,1168,1955,1979]" pageId="29" pageNumber="476">Imatong Foothills</collectingMunicipality>
,
<collectingCounty id="62C4C0CFFFCAFFE2FB5E47D9FADEF80F" box="[1180,1362,1955,1979]" pageId="29" pageNumber="476">Northern Region</collectingCounty>
,
<collectingCountry id="F30DF8D3FFCAFFE2FA9C47D9FA3CF80F" box="[1374,1456,1955,1979]" name="Uganda" pageId="29" pageNumber="476">Uganda</collectingCountry>
,
<geoCoordinate id="EE2EDE84FFC9FFE1FF43457EFF7BFAA8" box="[129,247,1284,1308]" degrees="03.80548" direction="north" orientation="latitude" pageId="30" pageNumber="477" precision="1" value="3.80548">N03.80548</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFC9FFE1FEC0457EFEFFFAA8" box="[258,371,1284,1308]" degrees="32.98836" direction="east" orientation="longitude" pageId="30" pageNumber="477" precision="1" value="32.98836">E32.98836</geoCoordinate>
,
<elevation id="00375F70FFC9FFE1FEBC457EFD8CFAA8" box="[382,512,1284,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.153" pageId="30" pageNumber="477" unit="m" value="1153.0">
<quantity id="4CE215A6FFC9FFE1FEBC457EFE42FAAF" box="[382,462,1284,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.153" pageId="30" pageNumber="477" unit="m" value="1153.0">1153 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFC9FFE1FDEB457EFD13FAA8" box="[553,671,1284,1308]" pageId="30" pageNumber="477" value="2015-07-05">
<collectingDate id="EFE0676BFFC9FFE1FDEB457EFD13FAA8" box="[553,671,1284,1308]" pageId="30" pageNumber="477" value="2015-07-05">5 July 2015</collectingDate>
</date>
by
<collectorName id="26EFDD95FFC9FFE1FD07457EFF62FA8F" pageId="30" pageNumber="477">Daniel F. Hughes</collectorName>
,
<collectorName id="26EFDD95FFC9FFE1FF394559FE26FA8F" box="[251,426,1315,1339]" pageId="30" pageNumber="477">Wilber Lukwago</collectorName>
, and
<collectorName id="26EFDD95FFC9FFE1FE2A4559FD34FA8F" box="[488,696,1315,1339]" pageId="30" pageNumber="477">Mathias Behangana</collectorName>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFC9FFE1FD04455EFE6DFAEE" collectionCode="UTEP, DFH" country="Uganda" pageId="30" pageNumber="477" specimenCode="UTEP 22593, DFH 130" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFC9FFE1FD04455EFF4EFAEE" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="30" pageNumber="477" type="Museum">UTEP 22593</specimenCode>
(
<specimenCode id="DBBC1038FFC9FFE1FF164538FECFFAEE" box="[212,323,1346,1370]" collectionCode="DFH" pageId="30" pageNumber="477">DFH 130</specimenCode>
), juvenile, and
</materialsCitation>
<materialsCitation id="3B72B21EFFC9FFE1FE2E4539FD39FA0C" collectingDate="2015-05-31" collectionCode="UTEP, DFH" collectorName="Daniel F. Hughes &amp; Mathias Behangana" country="Uganda" elevation="1730" latitude="1.8227" location="Mount Kadam" longLatPrecision="1" longitude="34.74603" municipality="Northern Region" pageId="30" pageNumber="477" specimenCode="UTEP 22594, DFH 131" specimenCount="1" stateProvince="Nakapiripirit" typeStatus="paratype">
<specimenCode id="DBBC1038FFC9FFE1FE2E4539FDF7FAEE" box="[492,635,1346,1370]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="30" pageNumber="477" type="Museum">UTEP 22594</specimenCode>
(
<specimenCode id="DBBC1038FFC9FFE1FD4F4538FD71FAEE" box="[653,765,1346,1370]" collectionCode="DFH" pageId="30" pageNumber="477">DFH 131</specimenCode>
), female, collected at
<collectingRegion id="49DE76A1FFC9FFE1FE9D4518FE6AFACE" box="[351,486,1378,1402]" country="Uganda" name="Nakapiripirit" pageId="30" pageNumber="477">Nakapiripirit</collectingRegion>
,
<location id="8EC5EE98FFC9FFE1FE354518FD1BFACE" LSID="urn:lsid:plazi:treatment:03B30955FFCAFFDEFF1F46B0FD56FE68:8EC5EE98FFC9FFE1FE354518FD1BFACE" box="[503,663,1378,1402]" country="Uganda" latitude="1.8227" longLatPrecision="1" longitude="34.74603" municipality="Northern Region" name="Mount Kadam" pageId="30" pageNumber="477" stateProvince="Nakapiripirit">Mount Kadam</location>
,
<collectingMunicipality id="6BC12239FFC9FFE1FD6A4518FF41FA2D" pageId="30" pageNumber="477">Northern Region</collectingMunicipality>
,
<collectingCountry id="F30DF8D3FFC9FFE1FF1845FBFEA0FA2D" box="[218,300,1409,1433]" name="Uganda" pageId="30" pageNumber="477">Uganda</collectingCountry>
,
<geoCoordinate id="EE2EDE84FFC9FFE1FEFA45FBFE23FA2D" box="[312,431,1409,1433]" degrees="01.82270" direction="north" orientation="latitude" pageId="30" pageNumber="477" precision="1" value="1.8227">N01.82270</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFC9FFE1FE7E45FBFDA1FA2D" box="[444,557,1409,1433]" degrees="34.74603" direction="east" orientation="longitude" pageId="30" pageNumber="477" precision="1" value="34.74603">E34.74603</geoCoordinate>
,
<elevation id="00375F70FFC9FFE1FDF845FBFD4EFA2D" box="[570,706,1409,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="30" pageNumber="477" unit="m" value="1730.0">
<quantity id="4CE215A6FFC9FFE1FDF845FBFD01FA2D" box="[570,653,1409,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="30" pageNumber="477" unit="m" value="1730.0">1730 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFC9FFE1FD2D45FBFF6BFA0C" pageId="30" pageNumber="477" value="2015-05-31">
<collectingDate id="EFE0676BFFC9FFE1FD2D45FBFF6BFA0C" pageId="30" pageNumber="477" value="2015-05-31">31 May 2015</collectingDate>
</date>
by
<collectorName id="26EFDD95FFC9FFE1FEC845DAFE36FA0C" box="[266,442,1440,1464]" pageId="30" pageNumber="477">Daniel F. Hughes</collectorName>
and
<collectorName id="26EFDD95FFC9FFE1FE2945DAFD39FA0C" box="[491,693,1440,1464]" pageId="30" pageNumber="477">Mathias Behangana</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF65E8CBFFC9FFE1FF434412FEC7FB60" pageId="30" pageNumber="477" startId="30.[129,194,1128,1152]" targetBox="[129,1473,144,1100]" targetPageId="30" targetType="figure">
<paragraph id="8BA5B843FFC9FFE1FF434412FEC7FB60" blockId="30.[129,1430,1127,1236]" pageId="30" pageNumber="477">
<emphasis id="B96E6451FFC9FFE1FF434412FF69FB34" bold="true" box="[129,229,1128,1152]" pageId="30" pageNumber="477">Figure 18.</emphasis>
Specimens of
<emphasis id="B96E6451FFC9FFE1FEAF441DFD0CFB34" bold="true" box="[365,640,1127,1152]" pageId="30" pageNumber="477">
<taxonomicName id="4C1AC3C0FFC9FFE1FEAF441DFDBCFBCB" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[365,560,1127,1151]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFC9FFE1FEAF441DFDBCFBCB" bold="true" box="[365,560,1127,1151]" italics="true" pageId="30" pageNumber="477">Lygodactylus kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFC9FFE1FDF74412FD0CFB34" box="[565,640,1128,1152]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi. Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D).
</paragraph>
</caption>
<subSubSection id="C300EBC8FFC9FFE0FF43459AFE1CFF7C" lastPageId="31" lastPageNumber="478" pageId="30" pageNumber="477" type="diagnosis">
<paragraph id="8BA5B843FFC9FFE1FF43459AFDF5F8E4" blockId="30.[129,778,1503,1967]" pageId="30" pageNumber="477">
<emphasis id="B96E6451FFC9FFE1FF43459AFE63FA43" box="[129,495,1503,1527]" italics="true" pageId="30" pageNumber="477">
Diagnosis:
<taxonomicName id="4C1AC3C0FFC9FFE1FF3445A5FE63FA43" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[246,495,1503,1527]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFC9FFE1FF3445A5FE63FA43" bold="true" box="[246,495,1503,1527]" pageId="30" pageNumber="477">Lygodactylus karamoja</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A25DD92AFFC9FFE1FE35459AFDC6FA4C" box="[503,586,1504,1528]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
is the sister taxon of
<emphasis id="B96E6451FFC9FFE1FF594585FEDCF9A3" bold="true" box="[155,336,1535,1559]" pageId="30" pageNumber="477">
<taxonomicName id="4C1AC3C0FFC9FFE1FF594585FF75F9A3" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[155,249,1535,1559]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFC9FFE1FF594585FF75F9A3" bold="true" box="[155,249,1535,1559]" italics="true" pageId="30" pageNumber="477">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFC9FFE1FF3C4585FEDCF9A3" box="[254,336,1535,1559]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
that is also a large-sized
<taxonomicName id="4C1AC3C0FFC9FFE1FD8B4585FD44F9A3" box="[585,712,1535,1559]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="genus">
<emphasis id="B96E6451FFC9FFE1FD8B4585FD44F9A3" box="[585,712,1535,1559]" italics="true" pageId="30" pageNumber="477">Lygodactylus</emphasis>
</taxonomicName>
[maximum
<collectionCode id="ED0B2086FFC9FFE1FF054664FF78F982" box="[199,244,1566,1590]" pageId="30" pageNumber="477">SVL</collectionCode>
<quantity id="4CE215A6FFC9FFE1FF3C4664FED1F982" box="[254,349,1566,1590]" metricMagnitude="-2" metricUnit="m" metricValue="3.77" pageId="30" pageNumber="477" unit="mm" value="37.7">37.7 mm</quantity>
(mean 34.4 ±
<quantity id="4CE215A6FFC9FFE1FE3D4664FDD4F982" box="[511,600,1566,1590]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="30" pageNumber="477" unit="mm" value="2.9">2.9 mm</quantity>
)] and shares the gular patterning of the
<taxonomicName id="4C1AC3C0FFC9FFE1FEBE4644FE79F9E2" box="[380,501,1598,1622]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC9FFE1FEBE4644FE79F9E2" box="[380,501,1598,1622]" italics="true" pageId="30" pageNumber="477">L. gutturalis</emphasis>
</taxonomicName>
subgroup. Seven to eight supralabials and 5–7 infralabials. Dorsal pholidosis with granular scales that become flattened, larger, and imbricate on original tails. Large triangular mental followed by usually two (or occasionally three) symmetric postmental scales (Supporting Information,
<figureCitation id="1321A4C6FFC9FFE1FECE46A0FED9F946" box="[268,341,1754,1778]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="30" pageNumber="477">Fig. S7</figureCitation>
). Ventral pholidosis with large, flattened, and imbricate scales. Ventral scales usually denticulated posteriorly. Six terminal scansors on the tail tip. Digits elongated with five terminal scansors on the fourth toe (
<tableCitation id="C6988DF8FFC9FFE1FDDC4742FDE4F8E4" box="[542,616,1848,1872]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="30" pageNumber="477">Table 3</tableCitation>
).
</paragraph>
<paragraph id="8BA5B843FFC9FFE1FF5E4722FA4FFACE" blockId="30.[129,778,1503,1967]" lastBlockId="30.[825,1475,1283,1965]" pageId="30" pageNumber="477">
Like other
<taxonomicName id="4C1AC3C0FFC9FFE1FEDF4722FE10F8C4" box="[285,412,1880,1904]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="genus">
<emphasis id="B96E6451FFC9FFE1FEDF4722FE10F8C4" box="[285,412,1880,1904]" italics="true" pageId="30" pageNumber="477">Lygodactylus</emphasis>
</taxonomicName>
within the
<taxonomicName id="4C1AC3C0FFC9FFE1FDE84722FD09F8C4" box="[554,645,1880,1904]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC9FFE1FDE84722FD09F8C4" box="[554,645,1880,1904]" italics="true" pageId="30" pageNumber="477">gutturalis</emphasis>
</taxonomicName>
group, this species can be easily differentiated from members of the
<taxonomicName id="4C1AC3C0FFC9FFE1FD374702FF57F81A" authorityName="Gunther" authorityYear="1893" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFC9FFE1FD374702FF57F81A" italics="true" pageId="30" pageNumber="477">L. angularis</emphasis>
</taxonomicName>
group by its characteristic
<emphasis id="B96E6451FFC9FFE1FE2447ECFE7BF81A" bold="true" box="[486,503,1942,1966]" pageId="30" pageNumber="477">∩</emphasis>
<emphasis id="B96E6451FFC9FFE1FE3547EDFE72F81A" box="[503,510,1943,1966]" italics="true" pageId="30" pageNumber="477">-</emphasis>
shaped gular chevrons (vs. V-shaped gular chevrons in
<taxonomicName id="4C1AC3C0FFC9FFE1FB9A457EFB40FAAF" authorityName="Gunther" authorityYear="1893" box="[1112,1228,1283,1307]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFC9FFE1FB9A457EFB40FAAF" box="[1112,1228,1283,1307]" italics="true" pageId="30" pageNumber="477">L. angularis</emphasis>
</taxonomicName>
group), and from other members of the
<taxonomicName id="4C1AC3C0FFC9FFE1FC27455EFBEEFA8F" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[997,1122,1316,1339]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFC9FFE1FC27455EFBEEFA8F" box="[997,1122,1316,1339]" italics="true" pageId="30" pageNumber="477">L. picturatus</emphasis>
</taxonomicName>
group based on dorsal coloration (light brown with five or six laterodorsal cream ocelli vs. usually blueish dorsum with yellow to white head in
<taxonomicName id="4C1AC3C0FFC9FFE1FB3A4518FAFEFACD" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[1272,1394,1378,1401]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFC9FFE1FB3A4518FAFEFACD" box="[1272,1394,1378,1401]" italics="true" pageId="30" pageNumber="477">L. picturatus</emphasis>
</taxonomicName>
group).
</paragraph>
<paragraph id="8BA5B843FFC9FFE0FC9745FAFE1CFF7C" blockId="30.[825,1475,1283,1965]" lastBlockId="31.[113,762,145,200]" lastPageId="31" lastPageNumber="478" pageId="30" pageNumber="477">
<emphasis id="B96E6451FFC9FFE1FC9745FAFB26FA2D" bold="true" box="[853,1194,1408,1433]" pageId="30" pageNumber="477">
<taxonomicName id="4C1AC3C0FFC9FFE1FC9745FAFBC2FA2C" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[853,1102,1408,1432]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFC9FFE1FC9745FAFBC2FA2C" bold="true" box="[853,1102,1408,1432]" italics="true" pageId="30" pageNumber="477">Lygodactylus karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFC9FFE1FB9445FBFB26FA2D" box="[1110,1194,1409,1433]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from other species within the
<taxonomicName id="4C1AC3C0FFC9FFE1FB8D45DAFB26FA0C" box="[1103,1194,1440,1464]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC9FFE1FB8D45DAFB26FA0C" box="[1103,1194,1440,1464]" italics="true" pageId="30" pageNumber="477">gutturalis</emphasis>
</taxonomicName>
subgroup by only minor morphometric and meristic data, reflecting its cryptic nature. However, we provide some characters that are putatively diagnostic.
<emphasis id="B96E6451FFC9FFE1FC404584FB5CF9A2" bold="true" box="[898,1232,1534,1558]" pageId="30" pageNumber="477">
<taxonomicName id="4C1AC3C0FFC9FFE1FC404584FBF5F9A2" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[898,1145,1534,1558]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFC9FFE1FC404584FBF5F9A2" bold="true" box="[898,1145,1534,1558]" italics="true" pageId="30" pageNumber="477">Lygodactylus karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFC9FFE1FBBC4584FB5CF9A2" box="[1150,1232,1534,1558]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C1AC3C0FFC9FFE1FA904585FA4EF9A2" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[1362,1474,1534,1558]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC9FFE1FA904585FA4EF9A2" box="[1362,1474,1534,1558]" italics="true" pageId="30" pageNumber="477">L. dysmicus</emphasis>
</taxonomicName>
by its larger size (maximum
<collectionCode id="ED0B2086FFC9FFE1FB9C4664FB07F982" box="[1118,1163,1566,1590]" pageId="30" pageNumber="477">SVL</collectionCode>
<quantity id="4CE215A6FFC9FFE1FB504664FB62F981" box="[1170,1262,1566,1590]" metricMagnitude="-2" metricUnit="m" metricValue="3.77" pageId="30" pageNumber="477" unit="mm" value="37.7">37.7 mm</quantity>
vs.
<quantity id="4CE215A6FFC9FFE1FAD44667FAFBF982" box="[1302,1399,1565,1590]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="30" pageNumber="477" unit="mm" value="27.6">27.6 mm</quantity>
); fewer precloacal pores (8 vs. 9); nostril not in contact with rostral scale (vs. nostril contacting rostral); and lower number of ventral scales across the body (16–18 vs.
<quantity id="4CE215A6FFC9FFE1FB004606FB7BF927" box="[1218,1271,1659,1684]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="30" pageNumber="477" unit="in" value="21.0">21 in</quantity>
<taxonomicName id="4C1AC3C0FFC9FFE1FB3C4606FAFCF927" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[1278,1392,1659,1683]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFC9FFE1FB3C4606FAFCF927" box="[1278,1392,1659,1683]" italics="true" pageId="30" pageNumber="477">L. dysmicus</emphasis>
</taxonomicName>
). It can be easily distinguished from
<emphasis id="B96E6451FFC9FFE1FB9D46E1FA99F907" bold="true" box="[1119,1301,1690,1715]" pageId="30" pageNumber="477">
<taxonomicName id="4C1AC3C0FFC9FFE1FB9D46E1FB32F906" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1119,1214,1690,1714]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFC9FFE1FB9D46E1FB32F906" bold="true" box="[1119,1214,1690,1714]" italics="true" pageId="30" pageNumber="477">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFC9FFE1FB0146E1FA99F907" box="[1219,1301,1691,1715]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the gular pattern that comprises two or three thinner
<emphasis id="B96E6451FFC9FFE1FB3846C0FA87F966" bold="true" box="[1274,1291,1722,1746]" pageId="30" pageNumber="477">∩</emphasis>
-shaped chevrons that never extend beyond the posterior part of the lower jaw vs. three thick
<emphasis id="B96E6451FFC9FFE1FC6E4683FC31F8A5" bold="true" box="[940,957,1785,1809]" pageId="30" pageNumber="477">∩</emphasis>
-shaped chevrons reaching the chest; there are usually two symmetrical postmental scales vs. usually three smaller postmental scales; and a proportionally smaller orbital diameter (
<collectionCode id="ED0B2086FFC9FFE1FC83472DFCE7F8DB" box="[833,875,1879,1903]" pageId="30" pageNumber="477">OD</collectionCode>
/HL 0.19–0.23 vs.
<quantity id="4CE215A6FFC9FFE1FBF1472DFB39F8DB" box="[1075,1205,1879,1903]" metricMagnitude="-3" metricUnit="m" metricValue="6.096" metricValueMax="6.604" metricValueMin="5.587999999999999" pageId="30" pageNumber="477" unit="in" value="0.24" valueMax="0.26" valueMin="0.22">0.22–0.26 in</quantity>
<emphasis id="B96E6451FFC9FFE1FB7F472DFAF4F8DB" bold="true" box="[1213,1400,1878,1903]" pageId="30" pageNumber="477">
<taxonomicName id="4C1AC3C0FFC9FFE1FB7F472DFA91F8DA" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1213,1309,1878,1902]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFC9FFE1FB7F472DFA91F8DA" bold="true" box="[1213,1309,1878,1902]" italics="true" pageId="30" pageNumber="477">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFC9FFE1FAE6472DFAF4F8DB" box="[1316,1400,1879,1903]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Also,
<emphasis id="B96E6451FFC9FFE1FCFB470CFB95F83A" bold="true" box="[825,1049,1910,1934]" pageId="30" pageNumber="477">
<taxonomicName id="4C1AC3C0FFC9FFE1FCFB470CFC31F83A" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[825,957,1910,1934]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFC9FFE1FCFB470CFC31F83A" bold="true" box="[825,957,1910,1934]" italics="true" pageId="30" pageNumber="477">L. karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFC9FFE1FC07470CFB95F83A" box="[965,1049,1910,1934]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated in the same way as
<taxonomicName id="4C1AC3C0FFC9FFE1FCFB47ECFC22F819" box="[825,942,1941,1965]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFC9FFE1FCFB47ECFC22F819" box="[825,942,1941,1965]" italics="true" pageId="30" pageNumber="477">L. gutturalis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C1AC3C0FFC9FFE1FC2F47ECFBEDF819" authorityName=", Schmidt" authorityYear="1919" box="[1005,1121,1941,1965]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFC9FFE1FC2F47ECFBEDF819" box="[1005,1121,1941,1965]" italics="true" pageId="30" pageNumber="477">L. depressus</emphasis>
</taxonomicName>
, based on gular patterning (always two or three
<emphasis id="B96E6451FFC8FFE0FF3940EBFE80FF1D" bold="true" box="[251,268,145,169]" pageId="31" pageNumber="478">∩</emphasis>
-shaped chevrons vs. one
<emphasis id="B96E6451FFC8FFE0FDDA40EBFDA5FF1D" bold="true" box="[536,553,145,169]" pageId="31" pageNumber="478">∩</emphasis>
-shaped followed by one posterior central mark).
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFC8FFE0FFB34093FEECFE2A" pageId="31" pageNumber="478" type="etymology">
<paragraph id="8BA5B843FFC8FFE0FFB34093FEECFE2A" blockId="31.[113,762,233,414]" pageId="31" pageNumber="478">
<emphasis id="B96E6451FFC8FFE0FFB34093FF6DFEB5" box="[113,225,233,257]" italics="true" pageId="31" pageNumber="478">Etymology:</emphasis>
The name
<taxonomicName id="4C1AC3C0FFC8FFE0FEA64093FE4BFEB5" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[356,455,233,257]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="478" phylum="Chordata" rank="species" species="karamoja">
‘
<emphasis id="B96E6451FFC8FFE0FEAB4093FE4BFEB5" box="[361,455,233,257]" italics="true" pageId="31" pageNumber="478">karamoja</emphasis>
</taxonomicName>
’ is a noun in apposition and refers to the Karamoja region in north-eastern
<collectingCountry id="F30DF8D3FFC8FFE0FDA14173FD3EFE95" box="[611,690,265,289]" name="Uganda" pageId="31" pageNumber="478">Uganda</collectingCountry>
where many individuals of this species have been found. The species is named in honour of this arid region, which is occupied by the Karamojong people who are mostly nomadic pastoralists related to the Maasai in
<collectingCountry id="F30DF8D3FFC8FFE0FEDA41FCFED0FE2A" box="[280,348,390,414]" name="Kenya" pageId="31" pageNumber="478">Kenya</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFC8FFDEFFB341BAFE7BFE91" lastPageId="33" lastPageNumber="480" pageId="31" pageNumber="478" type="description">
<paragraph id="8BA5B843FFC8FFE0FFB341BAFBCCFD08" blockId="31.[113,763,447,847]" lastBlockId="31.[810,1459,144,700]" pageId="31" pageNumber="478">
<emphasis id="B96E6451FFC8FFE0FFB341BAFE1EFE63" box="[113,402,447,471]" italics="true" pageId="31" pageNumber="478">
Description of the
<typeStatus id="54A106E1FFC8FFE0FE8241C5FE1EFE63" box="[320,402,447,471]" pageId="31" pageNumber="478" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="1321A4C6FFC8FFE0FE7E41C5FD80FE63" box="[444,524,447,471]" captionStart="Figure 19" captionStartId="31.[115,180,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-423@31.[114,1458,920,1870]" captionTargetPageId="31" captionText="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="31" pageNumber="478">Fig. 19</figureCitation>
): Measurements and meristic characters of the
<typeStatus id="54A106E1FFC8FFE0FEB841A4FE5AFE42" box="[378,470,478,502]" pageId="31" pageNumber="478" type="holotype">holotype</typeStatus>
are presented in Supporting Information, Table S8. Adult male, with a snout–vent length (
<collectionCode id="ED0B2086FFC8FFE0FFBB4267FF20FD81" box="[121,172,541,565]" pageId="31" pageNumber="478">SVL</collectionCode>
) of
<quantity id="4CE215A6FFC8FFE0FF1F4267FEB2FD81" box="[221,318,541,565]" metricMagnitude="-2" metricUnit="m" metricValue="3.4299999999999997" pageId="31" pageNumber="478" unit="mm" value="34.3">34.3 mm</quantity>
and an original tail length slightly larger (TL =
<quantity id="4CE215A6FFC8FFE0FF7B4246FEAFFDE1" box="[185,291,572,597]" metricMagnitude="-2" metricUnit="m" metricValue="3.61" pageId="31" pageNumber="478" unit="mm" value="36.1">36.1 mm</quantity>
). Body slender, nape moderately distinct. Head as broad as body and moderate head length (
<collectionCode id="ED0B2086FFC8FFE0FD584226FD46FDC0" box="[666,714,604,628]" pageId="31" pageNumber="478">HW</collectionCode>
/HL 0.72). Canthus rostralis not prominent. Eye diameter
<quantity id="4CE215A6FFC8FFE0FD6B4201FD76FD27" box="[681,762,635,659]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="31" pageNumber="478" unit="mm" value="1.9">1.9 mm</quantity>
with circular pupil. Ear to eye distance twice the orbital diameter (
<quantity id="4CE215A6FFC8FFE0FF6B42C0FE8FFD66" box="[169,259,698,722]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="31" pageNumber="478" unit="mm" value="3.2">3.2 mm</quantity>
). Snout rounded and slightly pointed. Granular scales on frontal area larger than occipital scales, with 22 small granular scales between the eyes. Dorsal scales granular from rostral to tail. Rostral undivided, in contact with 1st supralabial, prenasals, and one large internasal scale. Eight supralabials and 5–6 infralabials. Prenasal scale present and in contact with 1st supralabial. Nostril circular, bordered by 1st supralabial, prenasal, and one supranasal. Postnasal not contacting nostril, separated by 1st supralabial, which it contacts posteriorly. Four rows of scales between supralabials and orbit. Mental large, triangular, and rounded, bordered posteriorly by two large rounded symmetric postmental scales. Six asymmetrical post-postmental scales. Gular scales granular, rounded, and slightly smaller than ventral scales. Ventral scales imbricate, rounded, and larger than dorsal scales, with 16 scales rows across the venter. Body relatively robust and slightly elongated (
<collectionCode id="ED0B2086FFC8FFE0FB5C41B3FB5EFE55" box="[1182,1234,457,481]" pageId="31" pageNumber="478">TRL</collectionCode>
/
<collectionCode id="ED0B2086FFC8FFE0FB1941B3FA82FE55" box="[1243,1294,457,481]" pageId="31" pageNumber="478">SVL</collectionCode>
0.42). Tail with 34 enlarged transverse scales and six pairs of terminal scansors at the tip. Eight precloacal pores. Fore- and hind limbs moderately short, stout; forearm medium-sized (
<collectionCode id="ED0B2086FFC8FFE0FB1F425DFB71FD8B" box="[1245,1277,551,575]" pageId="31" pageNumber="478">FL</collectionCode>
/
<collectionCode id="ED0B2086FFC8FFE0FAC4425DFAB4FD8B" box="[1286,1336,551,575]" pageId="31" pageNumber="478">SVL</collectionCode>
0.16); tibia short (CL/
<collectionCode id="ED0B2086FFC8FFE0FC55423DFC46FDEB" box="[919,970,583,607]" pageId="31" pageNumber="478">SVL</collectionCode>
0.17). Digits elongated and unwebbed with 5–6 terminal scansors. Thumb rudimentary with a small terminal claw. Relative length of digits: I &lt;II =
<collectionCode id="ED0B2086FFC8FFE0FB0942FCFB50FD29" box="[1227,1244,646,669]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="478" type="Museum">V</collectionCode>
&lt;III &lt;IV (manus); I &lt;II &lt;
<collectionCode id="ED0B2086FFC8FFE0FCBC42DFFC03FD08" box="[894,911,677,700]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="478" type="Museum">V</collectionCode>
&lt;III &lt;IV (pes).
</paragraph>
<paragraph id="8BA5B843FFC8FFDFFCE842A4FDE8FEF1" blockId="31.[810,1459,734,852]" lastBlockId="32.[129,778,144,325]" lastPageId="32" lastPageNumber="479" pageId="31" pageNumber="478">
<emphasis id="B96E6451FFC8FFE0FCE842A4FC60FD42" box="[810,1004,734,758]" italics="true" pageId="31" pageNumber="478">Coloration: In life</emphasis>
(
<figureCitation id="1321A4C6FFC8FFE0FBC142A4FBC2FD42" box="[1027,1102,734,758]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="31" pageNumber="478">Fig. 20</figureCitation>
), dorsal coloration from light to dark brown to greyish or almost black (
<figureCitation id="1321A4C6FFC8FFE0FB0B4287FAAFFCA1" box="[1225,1315,765,789]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="31" pageNumber="478">Fig. 20D</figureCitation>
), with lighter cream-beige lines with five or six lighter cream dots surrounded by black flanks. Some specimens have lighter cream-beige dots along the vertebral line, giving them a granitic appearance (
<figureCitation id="1321A4C6FFF7FFDFFD2140EAFF27FF73" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="32" pageNumber="479">Fig. 20A</figureCitation>
). Black line from nostril to the anterior insertion of the forelimb. Gular region white and two or three black,
<emphasis id="B96E6451FFF7FFDFFDAE40B5FDF1FF53" bold="true" box="[620,637,207,231]" pageId="32" pageNumber="479">∩</emphasis>
-shaped chevrons. Venter uniformly orange or white. Tip of tail and digits dark brown.
<emphasis id="B96E6451FFF7FFDFFF124174FED2FE91" box="[208,350,270,293]" italics="true" pageId="32" pageNumber="479">In preservative</emphasis>
(
<typeStatus id="54A106E1FFF7FFDFFEAE4177FE5CFE91" box="[364,464,269,293]" pageId="32" pageNumber="479" type="holotype">holotype</typeStatus>
;
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): dorsum dark brown and venter with uniform light cream coloration.
</paragraph>
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<paragraph id="8BA5B843FFC8FFE0FFB14710FDDAF80E" blockId="31.[113,1443,1898,1979]" pageId="31" pageNumber="478">
<emphasis id="B96E6451FFC8FFE0FFB14710FF54F837" bold="true" box="[115,216,1898,1923]" pageId="31" pageNumber="478">Figure 19.</emphasis>
Holotype of
<emphasis id="B96E6451FFC8FFE0FE964711FD96F837" bold="true" box="[340,538,1898,1923]" pageId="31" pageNumber="478">
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<emphasis id="B96E6451FFC8FFE0FE964711FE40F836" bold="true" box="[340,460,1898,1922]" italics="true" pageId="31" pageNumber="478">L. karamoja</emphasis>
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</emphasis>
(UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda. A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.
</paragraph>
</caption>
<paragraph id="8BA5B843FFF7FFDFFF43411DFAEAFF53" blockId="32.[129,778,358,413]" lastBlockId="32.[825,1474,144,231]" pageId="32" pageNumber="479">
<emphasis id="B96E6451FFF7FFDFFF43411DFF68FECA" box="[129,228,359,382]" italics="true" pageId="32" pageNumber="479">Variation:</emphasis>
The meristic and morphometric data are summarized in Supporting Information, Table S8. Coloration of this species seems to be consistent, with gular coloration slightly lighter in females. Number and shape of postmentals and internasals vary between specimens (Supporting Information,
<figureCitation id="1321A4C6FFF7FFDFFAD240B5FADAFF53" box="[1296,1366,207,231]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="32" pageNumber="479">Fig. S7</figureCitation>
).
</paragraph>
<paragraph id="8BA5B843FFF7FFDEFCFB4172FE7BFE91" blockId="32.[825,1474,263,414]" lastBlockId="33.[113,762,144,293]" lastPageId="33" lastPageNumber="480" pageId="32" pageNumber="479">
<emphasis id="B96E6451FFF7FFDFFCFB4172FBA1FE94" box="[825,1069,264,288]" italics="true" pageId="32" pageNumber="479">Habitat and distribution</emphasis>
(
<figureCitation id="1321A4C6FFF7FFDFFB8A4172FB0BFE94" box="[1096,1159,264,288]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="32" pageNumber="479">Figs 6</figureCitation>
,
<figureCitation id="1321A4C6FFF7FFDFFB564172FB22FE94" box="[1172,1198,264,288]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." pageId="32" pageNumber="479">20</figureCitation>
<emphasis id="B96E6451FFF7FFDFFB6C4172FA4EFEAB" box="[1198,1474,263,287]" italics="true" pageId="32" pageNumber="479">
):
<taxonomicName id="4C1AC3C0FFF7FFDFFB0B417DFA4EFEAB" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1225,1474,263,287]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="479" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFF7FFDFFB0B417DFA4EFEAB" bold="true" box="[1225,1474,263,287]" pageId="32" pageNumber="479">Lygodactylus karamoja</emphasis>
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<taxonomicNameLabel id="A25DD92AFFF7FFDFFCFB415DFC1CFE8B" box="[825,912,295,319]" pageId="32" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
has been recorded from mid-elevation savannahs of the northern Albertine Rift to the Karamoja Region, eastern
<collectingCountry id="F30DF8D3FFF7FFDFFCFB411CFC07FECA" box="[825,907,358,382]" name="Uganda" pageId="32" pageNumber="479">Uganda</collectingCountry>
, an arid area characterized by mostly woodland savannah habitat with several isolated mountain ranges. This suggests that specimens from western
<collectingCountry id="F30DF8D3FFF6FFDEFEB340EAFE3EFF1C" box="[369,434,144,168]" name="Kenya" pageId="33" pageNumber="480">Kenya</collectingCountry>
and southern
<collectingCountry id="F30DF8D3FFF6FFDEFD8540EAFD47FF1C" box="[583,715,144,168]" name="South Sudan" pageId="33" pageNumber="480">South Sudan</collectingCountry>
also belong to this species. If confirmed, this species would have a distribution that is similar to the lacertid
<taxonomicName id="4C1AC3C0FFF6FFDEFD9740B5FE0CFEB2" authority="(Greenbaum et al. 2018)" baseAuthorityName="Greenbaum" baseAuthorityYear="2018" class="Reptilia" family="Lacertidae" genus="Adolfus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="jacksoni">
<emphasis id="B96E6451FFF6FFDEFD9740B5FD76FF53" box="[597,762,207,231]" italics="true" pageId="33" pageNumber="480">Adolfus jacksoni</emphasis>
(Greenbaum
<emphasis id="B96E6451FFF6FFDEFEC34095FEBAFEB2" box="[257,310,238,262]" italics="true" pageId="33" pageNumber="480">et al.</emphasis>
2018)
</taxonomicName>
. However, we recommend caution until molecular data become available.
</paragraph>
</subSubSection>
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<emphasis id="B96E6451FFF7FFDFFF424710FF69F837" bold="true" box="[128,229,1898,1923]" pageId="32" pageNumber="479">Figure 20.</emphasis>
Specimens of
<emphasis id="B96E6451FFF7FFDFFEAE4711FDBFF837" bold="true" box="[364,563,1898,1923]" pageId="32" pageNumber="479">
<taxonomicName id="4C1AC3C0FFF7FFDFFEAE4711FE68F836" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[364,484,1898,1922]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="479" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFF7FFDFFEAE4711FE68F836" bold="true" box="[364,484,1898,1922]" italics="true" pageId="32" pageNumber="479">L. karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF7FFDFFE2A4711FDBFF837" box="[488,563,1899,1923]" pageId="32" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda. Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H.
</paragraph>
</caption>
<subSubSection id="C300EBC8FFF6FFDEFFB3413DFD56FE68" pageId="33" pageNumber="480" type="biology_ecology">
<paragraph id="8BA5B843FFF6FFDEFFB3413DFD56FE68" blockId="33.[113,762,327,476]" pageId="33" pageNumber="480">
<emphasis id="B96E6451FFF6FFDEFFB3413DFE9DFEEA" box="[113,273,327,351]" italics="true" pageId="33" pageNumber="480">Natural history:</emphasis>
A diurnal and arboreal species that is always found on trees.
<collectionCode id="ED0B2086FFF6FFDEFED3411CFED8FEC9" box="[273,340,358,381]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="33" pageNumber="480" type="Museum">UTEP</collectionCode>
22592, a gravid female, was found in July on a tree at night.
<collectionCode id="ED0B2086FFF6FFDEFEE341FCFEE8FE29" box="[289,356,390,413]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="33" pageNumber="480" type="Museum">UTEP</collectionCode>
22593–94 were found during the day on trees trunks
<emphasis id="B96E6451FFF6FFDEFF2C41DFFF7BFE08" box="[238,247,421,444]" italics="true" pageId="33" pageNumber="480">c</emphasis>
.
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above ground in an open clearing of grass and rocky outcrops that were surrounded by dry forest (
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).
</paragraph>
</subSubSection>
</treatment>
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<mods:number id="F817D138AFC7E0B3F096AF79ECB1ED79">2</mods:number>
</mods:detail>
<mods:extent id="5ABDEA2C568E1228D63A8EF956AEC960" unit="page">
<mods:start id="16F313BD1FE055C0A2749C934180BDDC">447</mods:start>
<mods:end id="2EBAF5D5F0A545E673C23E16A7448450">492</mods:end>
</mods:extent>
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<mods:location id="147CFFC3FDADADD4C4C8EF69B02AC5F5">
<mods:url id="134B83242B3E355223693D4544F0F543">http://dx.doi.org/10.1093/zoolinnean/zlad123</mods:url>
</mods:location>
<mods:classification id="2797708EB3A14283DB4BDB3B8A483D63">journal article</mods:classification>
<mods:identifier id="7787201C6A5347395A7A8E360B6A7124" type="DOI">10.1093/zoolinnean/zlad123</mods:identifier>
<mods:identifier id="A34E00213CC1395A742ED8E9196ED131" type="ISSN">0024-4082</mods:identifier>
</mods:mods>
<treatment id="03B30955FFCCFFE2FC6E4462FF66F92B" LSID="urn:lsid:plazi:treatment:03B30955FFCCFFE2FC6E4462FF66F92B" httpUri="http://treatment.plazi.org/id/03B30955FFCCFFE2FC6E4462FF66F92B" lastPageId="29" lastPageNumber="476" pageId="27" pageNumber="474">
<subSubSection id="C300EBC8FFCCFFE4FC6E4462FABDFB86" box="[940,1329,1047,1074]" pageId="27" pageNumber="474" type="nomenclature">
<paragraph id="8BA5B843FFCCFFE4FC6E4462FABDFB86" blockId="27.[810,1459,1047,1110]" box="[940,1329,1047,1074]" pageId="27" pageNumber="474">
<emphasis id="B96E6451FFCCFFE4FC6E4462FABDFB86" bold="true" box="[940,1329,1047,1074]" pageId="27" pageNumber="474">
<taxonomicName id="4C1AC3C0FFCCFFE4FC6E4462FB41FB86" ID-CoL="C7JCX" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[940,1229,1048,1074]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="477" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFCCFFE4FC6E4462FB41FB86" bold="true" box="[940,1229,1048,1074]" italics="true" pageId="27" pageNumber="474">LYGODACTYLUS KIBERA</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFCCFFE4FB10446DFABDFB86" box="[1234,1329,1047,1074]" pageId="27" pageNumber="474" rank="species">SP.NOV.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCCFFE4FCE84444FBD7FB1B" pageId="27" pageNumber="474" type="description">
<paragraph id="8BA5B843FFCCFFE4FCE84444FA3FFBE2" blockId="27.[810,1459,1047,1110]" box="[810,1459,1086,1110]" pageId="27" pageNumber="474">
(
<figureCitation id="1321A4C6FFCCFFE4FCF74444FC28FBE2" box="[821,932,1086,1110]" captionStart-0="Figure 17" captionStart-1="Figure 18" captionStartId-0="29.[113,178,1122,1146]" captionStartId-1="30.[129,194,1128,1152]" captionTargetBox-0="[114,1458,144,1094]" captionTargetBox-1="[129,1473,144,1100]" captionTargetId-0="figure-359@29.[114,1458,144,1094]" captionTargetId-1="figure-444@30.[129,1473,144,1100]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus." captionText-1="Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D)." pageId="27" pageNumber="474">Figs 17–18</figureCitation>
,
<tableCitation id="C6988DF8FFCCFFE4FC6C4444FC74FBE2" box="[942,1016,1086,1110]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="27" pageNumber="474">Table 3</tableCitation>
; Supporting Information,
<figureCitation id="1321A4C6FFCCFFE4FAC34444FACAFBE2" box="[1281,1350,1086,1110]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="27" pageNumber="474">Fig. S6</figureCitation>
; Table S7)
</paragraph>
<paragraph id="8BA5B843FFCCFFE4FCE84402FBD7FB1B" blockId="27.[810,1459,1144,1199]" pageId="27" pageNumber="474">
<emphasis id="B96E6451FFCCFFE4FCE84402FB84FB3B" box="[810,1032,1144,1168]" italics="true" pageId="27" pageNumber="474">Zoobank registration:</emphasis>
https://zoobank.org/
<uuid id="FFBC8296FFCCFFE4FB364402FBD7FB1B" pageId="27" pageNumber="474">26F76FD8-043A-4226-BDD5-E971235D474C</uuid>
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCCFFE4FCFE44B4FBD5F9D2" pageId="27" pageNumber="474" type="reference_group">
<paragraph id="8BA5B843FFCCFFE4FCFE44B4FB1CFA9C" blockId="27.[810,1459,1230,1320]" pageId="27" pageNumber="474">
<emphasis id="B96E6451FFCCFFE4FCFE44B4FB1CFA9C" italics="true" pageId="27" pageNumber="474">
<taxonomicName id="4C1AC3C0FFCCFFE4FCFE44B4FB16FB5C" ID-CoL="3WQ3T" authorityName="' (Schmidt" authorityYear="1919" box="[828,1178,1230,1256]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="474" phylum="Chordata" rank="subSpecies" species="picturatus" subSpecies="gutturalis">Lygodactylus picturatus gutturalis</taxonomicName>
:
<bibRefCitation id="EF8BC5B2FFCCFFE4FB6744B4FACDFB5D" author="Schmidt KP" box="[1189,1345,1230,1257]" pageId="27" pageNumber="474" pagination="385 - 624" refId="ref37222" refString="Schmidt KP. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909 - 1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bulletin of the American Museum of Natural History 1919; 39: 385 - 624." type="journal article" year="1919">Schmidt (1919)</bibRefCitation>
[part]; de Witte (1953) [part]; Pasteur, 1965(1964) [part];
<bibRefCitation id="EF8BC5B2FFCCFFE4FACE4494FA3FFABC" author="Wermuth H" box="[1292,1459,1262,1288]" pageId="27" pageNumber="474" pagination="1 - 246" refId="ref37733" refString="Wermuth H. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Das Tierreich 1965; 80: I - XXIII, 1 - 246." type="journal article" year="1965">Wermuth (1965)</bibRefCitation>
[part].
</emphasis>
</paragraph>
<paragraph id="8BA5B843FFCCFFE4FCEC4536FB1CFA32" blockId="27.[810,1459,1356,1638]" pageId="27" pageNumber="474">
<emphasis id="B96E6451FFCCFFE4FCEC4536FB1CFA32" italics="true" pageId="27" pageNumber="474">
<taxonomicName id="4C1AC3C0FFCCFFE4FCEC4536FB92FAD2" ID-CoL="3WQ3T" authority="(Bocage, 1873)" box="[814,1054,1356,1382]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="474" phylum="Chordata" rank="species" species="gutturalis">Lygodactylus gutturalis</taxonomicName>
:
<bibRefCitation id="EF8BC5B2FFCCFFE4FBEB4536FB12FAD2" author="Roll B" box="[1065,1182,1356,1382]" pageId="27" pageNumber="474" pagination="645 - 8" refId="ref36800" refString="Roll B. Lygodactylus gutturalis (Bocage). Sauria (Suppl.) 2005; 27: 645 - 8." type="journal article" year="2005">Röll (2005)</bibRefCitation>
[part];
<bibRefCitation id="EF8BC5B2FFCCFFE4FB284536FA23FAD2" author="Spawls S &amp; Howell K &amp; Hinkel H &amp; Menegon M" box="[1258,1455,1356,1382]" pageId="27" pageNumber="474" refId="ref37334" refString="Spawls S, Howell K, Hinkel H, Menegon M. 2018. Field Guide to East African Reptiles. London: Bloomsbury." type="book" year="2018">Spawls et al. (2018)</bibRefCitation>
[part].
</emphasis>
</paragraph>
<paragraph id="8BA5B843FFCCFFE4FCE845E8FBD5F9D2" blockId="27.[810,1459,1356,1638]" pageId="27" pageNumber="474">
<emphasis id="B96E6451FFCCFFE4FCE845E8FBB2FA1D" box="[810,1086,1426,1450]" italics="true" pageId="27" pageNumber="474">
<taxonomicName id="4C1AC3C0FFCCFFE4FCE845E8FC61FA1E" ID-CoL="C7JCX" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[810,1005,1426,1450]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="474" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">Lygodactylus kibera</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFCCFFE4FC3745E8FBB2FA1D" box="[1013,1086,1426,1449]" pageId="27" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to a distinctive clade (B1), from the Albertine Rift of
<collectingCountry id="F30DF8D3FFCCFFE4FBED45CBFB08FA7D" box="[1071,1156,1457,1481]" name="Burundi" pageId="27" pageNumber="474">Burundi</collectingCountry>
and eastern
<collectingCountry id="F30DF8D3FFCCFFE4FAC245CBFAB4FA7D" box="[1280,1336,1457,1481]" name="Democratic Republic of the Congo" pageId="27" pageNumber="474">DRC</collectingCountry>
, which clusters as sister to another candidate new species from the northern Albertine Rift in the Lendu Plateau of
<collectingCountry id="F30DF8D3FFCCFFE4FB01458AFB75F9BC" box="[1219,1273,1520,1544]" name="Democratic Republic of the Congo" pageId="27" pageNumber="474">DRC</collectingCountry>
and several highlands of
<collectingCountry id="F30DF8D3FFCCFFE4FC474675FC5BF993" box="[901,983,1551,1575]" name="Uganda" pageId="27" pageNumber="474">Uganda</collectingCountry>
, but differs from it by
<emphasis id="B96E6451FFCCFFE4FB04466AFB43F993" box="[1222,1231,1552,1575]" italics="true" pageId="27" pageNumber="474">c</emphasis>
. 6.07% in
<emphasis id="B96E6451FFCCFFE4FA824675FAEBF993" box="[1344,1383,1551,1575]" italics="true" pageId="27" pageNumber="474">16S</emphasis>
uncorrected p-distance (Table 2), and a lack of nuclear haplotype sharing in
<emphasis id="B96E6451FFCCFFE4FC564634FC5CF9D2" box="[916,976,1614,1638]" italics="true" pageId="27" pageNumber="474">RAG1</emphasis>
(
<figureCitation id="1321A4C6FFCCFFE4FC224634FBC4F9D2" box="[992,1096,1614,1638]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability&gt; 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="27" pageNumber="474">Fig. 2B–C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCCFFE3FCE846F8FD9EFDB5" lastPageId="28" lastPageNumber="475" pageId="27" pageNumber="474" type="materials_examined">
<paragraph id="8BA5B843FFCCFFE4FCE846F8FBB1F8A3" blockId="27.[810,1459,1666,1815]" pageId="27" pageNumber="474">
<materialsCitation id="3B72B21EFFCCFFE4FCE846F8FBB1F8A3" collectingDate="2011-12-20" collectionCode="UTEP, ELI" country="Burundi" elevation="1660" latitude="-3.06974" location="Mpishi" longLatPrecision="1" longitude="29.48445" municipality="Kibira National Park" pageId="27" pageNumber="474" specimenCode="UTEP 22566, ELI 1145" specimenCount="1" stateProvince="Bubanza" typeStatus="holotype">
<emphasis id="B96E6451FFCCFFE4FCE846F8FC06F92E" box="[810,906,1666,1690]" italics="true" pageId="27" pageNumber="474">
<typeStatus id="54A106E1FFCCFFE4FCE846F8FC09F92E" box="[810,901,1666,1690]" pageId="27" pageNumber="474" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DBBC1038FFCCFFE4FC5546F9FBAEF92E" box="[919,1058,1666,1690]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="27" pageNumber="474" type="Museum">UTEP 22566</specimenCode>
(
<specimenCode id="DBBC1038FFCCFFE4FBED46F8FB14F92E" box="[1071,1176,1666,1690]" collectionCode="ELI" pageId="27" pageNumber="474">ELI 1145</specimenCode>
), a male with a ventral incision, collected in a village near montane forest at
<location id="8EC5EE98FFCCFFE4FAF346D8FAF7F90E" LSID="urn:lsid:plazi:treatment:03B30955FFCCFFE2FC6E4462FF66F92B:8EC5EE98FFCCFFE4FAF346D8FAF7F90E" box="[1329,1403,1698,1722]" country="Burundi" latitude="-3.06974" longLatPrecision="1" longitude="29.48445" municipality="Kibira National Park" name="Mpishi" pageId="27" pageNumber="474" stateProvince="Bubanza">Mpishi</location>
, near
<collectingMunicipality id="6BC12239FFCCFFE4FCE846BBFB8AF96D" box="[810,1030,1729,1753]" pageId="27" pageNumber="474">Kibira National Park</collectingMunicipality>
,
<collectingRegion id="49DE76A1FFCCFFE4FBD046BBFB43F96D" box="[1042,1231,1729,1753]" country="Burundi" name="Bubanza" pageId="27" pageNumber="474">Bubanza Province</collectingRegion>
,
<collectingCountry id="F30DF8D3FFCCFFE4FB1E46BBFABFF96D" box="[1244,1331,1729,1753]" name="Burundi" pageId="27" pageNumber="474">Burundi</collectingCountry>
,
<geoCoordinate id="EE2EDE84FFCCFFE4FAFD46BBFA3CF96D" box="[1343,1456,1729,1753]" degrees="03.06974" direction="south" orientation="latitude" pageId="27" pageNumber="474" precision="1" value="-3.06974">S03.06974</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFCCFFE4FCE8469AFC10F94C" box="[810,924,1760,1784]" degrees="29.48445" direction="east" orientation="longitude" pageId="27" pageNumber="474" precision="1" value="29.48445">E29.48445</geoCoordinate>
,
<elevation id="00375F70FFCCFFE4FC6A469AFBA1F94C" box="[936,1069,1760,1784]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="27" pageNumber="474" unit="m" value="1660.0">
<quantity id="4CE215A6FFCCFFE4FC6A469AFC75F94C" box="[936,1017,1760,1784]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="27" pageNumber="474" unit="m" value="1660.0">1660 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCCFFE4FB9A469AFAADF94C" box="[1112,1313,1760,1784]" pageId="27" pageNumber="474" value="2011-12-20">
<collectingDate id="EFE0676BFFCCFFE4FB9A469AFAADF94C" box="[1112,1313,1760,1784]" pageId="27" pageNumber="474" value="2011-12-20">20 December 2011</collectingDate>
</date>
by locals and brought to Eli Greenbaum.
</materialsCitation>
</paragraph>
<paragraph id="8BA5B843FFCCFFE3FCE84740FD9EFDB5" blockId="27.[810,1459,1849,1998]" lastBlockId="28.[129,778,144,513]" lastPageId="28" lastPageNumber="475" pageId="27" pageNumber="474">
<emphasis id="B96E6451FFCCFFE4FCE84740FBBFF8E4" box="[810,1075,1849,1873]" italics="true" pageId="27" pageNumber="474">
<typeStatus id="54A106E1FFCCFFE4FCE84740FC05F8E5" box="[810,905,1850,1873]" pageId="27" pageNumber="474" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D1C73CAFFCCFFE4FC5F4743FBABF8E4" box="[925,1063,1849,1873]" count="11" pageId="27" pageNumber="474" type="generic">11 specimens</specimenCount>
):
</emphasis>
<emphasis id="B96E6451FFCCFFE4FB8A4743FBDFF8E5" bold="true" box="[1096,1107,1849,1873]" pageId="27" pageNumber="474">•</emphasis>
<materialsCitation id="3B72B21EFFCCFFE4FB9C4743FAEEF8E5" box="[1118,1378,1849,1873]" country="Burundi" location="Burundi" pageId="27" pageNumber="474" specimenCount="10" typeStatus="paratype">
<emphasis id="B96E6451FFCCFFE4FB9C4743FB23F8E5" box="[1118,1199,1849,1873]" italics="true" pageId="27" pageNumber="474">
<collectingCountry id="F30DF8D3FFCCFFE4FB9C4743FB23F8E5" box="[1118,1199,1849,1873]" name="Burundi" pageId="27" pageNumber="474">Burundi</collectingCountry>
</emphasis>
(
<specimenCount id="9D1C73CAFFCCFFE4FB004743FAD7F8E5" box="[1218,1371,1849,1873]" count="10" pageId="27" pageNumber="474" type="generic">10 specimens</specimenCount>
)
</materialsCitation>
:
<materialsCitation id="3B72B21EFFCCFFE4FAAD4743FB55F8C4" collectionCode="UTEP, ELI" country="Burundi" pageId="27" pageNumber="474" specimenCode="UTEP 22567-69, ELI 1146-48" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFCCFFE4FAAD4743FC1FF8C4" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="27" pageNumber="474" type="Museum">UTEP 22567–69</specimenCode>
(
<specimenCode id="DBBC1038FFCCFFE4FC644722FBB1F8C4" box="[934,1085,1880,1904]" collectionCode="ELI" pageId="27" pageNumber="474">ELI 1146–48</specimenCode>
), females, and
</materialsCitation>
<materialsCitation id="3B72B21EFFCCFFE4FB274723FAE4F83B" collectingDate="2011-12-20" collectionCode="UTEP, ELI" country="Burundi" elevation="1660" latitude="-3.06974" location="Mpishi" longLatPrecision="1" longitude="29.48445" municipality="Kibira National Park" pageId="27" pageNumber="474" specimenCode="UTEP 22570, ELI 1149" specimenCount="1" stateProvince="Bubanza" typeStatus="paratype">
<specimenCode id="DBBC1038FFCCFFE4FB274723FAF9F8C4" box="[1253,1397,1880,1904]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="27" pageNumber="474" type="Museum">UTEP 22570</specimenCode>
(
<specimenCode id="DBBC1038FFCCFFE4FA4A4722FCEFF824" collectionCode="ELI" pageId="27" pageNumber="474">ELI 1149</specimenCode>
), male with the same collection data as the holotype
</materialsCitation>
;
<materialsCitation id="3B72B21EFFCCFFE4FAAD4702FB36F81B" collectionCode="UTEP, ELI" country="Burundi" pageId="27" pageNumber="474" specimenCode="UTEP 22571-73, ELI 1195-97" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFCCFFE4FAAD4702FC1FF81B" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="27" pageNumber="474" type="Museum">UTEP 22571–73</specimenCode>
(
<specimenCode id="DBBC1038FFCCFFE4FC6147EDFBBAF81B" box="[931,1078,1943,1967]" collectionCode="ELI" pageId="27" pageNumber="474">ELI 1195–97</specimenCode>
), males, and
</materialsCitation>
<materialsCitation id="3B72B21EFFCCFFE3FB0147EDFDB6FF53" collectingDate="2011-12-21" collectionCode="UTEP, ELI" collectorName="Wandege M. Muninga &amp; Eli Greenbaum" country="Burundi" elevation="1705" lastPageId="28" lastPageNumber="475" latitude="-3.06749" location="Kibira National Park" longLatPrecision="1" longitude="29.4856" municipality="Mpishi" pageId="27" pageNumber="474" specimenCode="UTEP 22574-75, ELI 1199-98" specimenCount="1" stateProvince="Bubanza" typeStatus="paratype">
<specimenCode id="DBBC1038FFCCFFE4FB0147EDFAF4F81B" box="[1219,1400,1943,1967]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="27" pageNumber="474" type="Museum">UTEP 22574–75</specimenCode>
(
<specimenCode id="DBBC1038FFCCFFE4FA4A47EDFC07F87A" collectionCode="ELI" pageId="27" pageNumber="474">ELI 1199–98</specimenCode>
), a male and a female, respectively, collected in a banana field at
<collectingMunicipality id="6BC12239FFCBFFE3FF1040EAFE90FF1C" box="[210,284,144,168]" pageId="28" pageNumber="475">Mpishi</collectingMunicipality>
, near
<location id="8EC5EE98FFCBFFE3FE9D40EAFDB7FF1C" LSID="urn:lsid:plazi:treatment:03B30955FFCCFFE2FC6E4462FF66F92B:8EC5EE98FFCBFFE3FE9D40EAFDB7FF1C" box="[351,571,144,168]" country="Burundi" latitude="-3.06749" longLatPrecision="1" longitude="29.4856" municipality="Mpishi" name="Kibira National Park" pageId="28" pageNumber="475" stateProvince="Bubanza">Kibira National Park</location>
,
<collectingRegion id="49DE76A1FFCBFFE3FD8A40EAFC89FF1C" box="[584,773,144,168]" country="Burundi" name="Bubanza" pageId="28" pageNumber="475">Bubanza Province</collectingRegion>
,
<geoCoordinate id="EE2EDE84FFCBFFE3FF4340D5FF7DFF73" box="[129,241,175,199]" degrees="03.06749" direction="south" orientation="latitude" pageId="28" pageNumber="475" precision="1" value="-3.06749">S03.06749</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFCBFFE3FF3940D5FEE0FF73" box="[251,364,175,199]" degrees="29.48560" direction="east" orientation="longitude" pageId="28" pageNumber="475" precision="1" value="29.4856">E29.48560</geoCoordinate>
,
<elevation id="00375F70FFCBFFE3FEB540D5FE76FF73" box="[375,506,175,199]" metricMagnitude="3" metricUnit="m" metricValue="1.705" pageId="28" pageNumber="475" unit="m" value="1705.0">
<quantity id="4CE215A6FFCBFFE3FEB540D5FE4BFF73" box="[375,455,175,199]" metricMagnitude="3" metricUnit="m" metricValue="1.705" pageId="28" pageNumber="475" unit="m" value="1705.0">1705 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCBFFE3FDE040D5FD65FF73" box="[546,745,175,199]" pageId="28" pageNumber="475" value="2011-12-21">
<collectingDate id="EFE0676BFFCBFFE3FDE040D5FD65FF73" box="[546,745,175,199]" pageId="28" pageNumber="475" value="2011-12-21">21 December 2011</collectingDate>
</date>
by
<collectorName id="26EFDD95FFCBFFE3FF4340B5FEEAFF53" box="[129,358,207,231]" pageId="28" pageNumber="475">Wandege M. Muninga</collectorName>
and
<collectorName id="26EFDD95FFCBFFE3FE5840B5FDB6FF53" box="[410,570,207,231]" pageId="28" pageNumber="475">Eli Greenbaum</collectorName>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFCBFFE3FD8740B5FD3EFEF1" collectingDate="2011-12-16" collectionCode="UTEP, ELI" collectorName="Wandege M. Muninga &amp; Eli Greenbaum" country="Burundi" elevation="811" latitude="-3.38236" location="City" longLatPrecision="1" longitude="29.36419" pageId="28" pageNumber="475" specimenCode="UTEP 22576, ELI 1071" specimenCount="1" stateProvince="Bujumbura Mairie" typeStatus="paratype">
<specimenCode id="DBBC1038FFCBFFE3FD8740B5FD5DFF52" box="[581,721,206,230]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="28" pageNumber="475" type="Museum">UTEP 22576</specimenCode>
(
<specimenCode id="DBBC1038FFCBFFE3FD1D40B5FF35FEB2" collectionCode="ELI" pageId="28" pageNumber="475">ELI 1071</specimenCode>
), female, collected at
<collectingRegion id="49DE76A1FFCBFFE3FE5B4094FD81FEB2" box="[409,525,238,262]" country="Burundi" name="Bujumbura Mairie" pageId="28" pageNumber="475">Bujumbura</collectingRegion>
<location id="8EC5EE98FFCBFFE3FDD14094FDCCFEB2" LSID="urn:lsid:plazi:treatment:03B30955FFCCFFE2FC6E4462FF66F92B:8EC5EE98FFCBFFE3FDD14094FDCCFEB2" box="[531,576,238,262]" country="Burundi" latitude="-3.38236" longLatPrecision="1" longitude="29.36419" name="City" pageId="28" pageNumber="475" stateProvince="Bujumbura Mairie">City</location>
, Bujumbura Mairie Province,
<geoCoordinate id="EE2EDE84FFCBFFE3FF2A4177FED4FE91" box="[232,344,269,293]" degrees="03.38236" direction="south" orientation="latitude" pageId="28" pageNumber="475" precision="1" value="-3.38236">S03.38236</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFCBFFE3FEA04177FE58FE91" box="[354,468,269,293]" degrees="29.36419" direction="east" orientation="longitude" pageId="28" pageNumber="475" precision="1" value="29.36419">E29.36419</geoCoordinate>
,
<elevation id="00375F70FFCBFFE3FE1D4177FDD8FE91" box="[479,596,269,293]" metricMagnitude="2" metricUnit="m" metricValue="8.11" pageId="28" pageNumber="475" unit="m" value="811.0">
<quantity id="4CE215A6FFCBFFE3FE1D4177FDAEFE91" box="[479,546,269,293]" metricMagnitude="2" metricUnit="m" metricValue="8.11" pageId="28" pageNumber="475" unit="m" value="811.0">811 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCBFFE3FDBF4177FF39FEF1" pageId="28" pageNumber="475" value="2011-12-16">
<collectingDate id="EFE0676BFFCBFFE3FDBF4177FF39FEF1" pageId="28" pageNumber="475" value="2011-12-16">16 December 2011</collectingDate>
</date>
by
<collectorName id="26EFDD95FFCBFFE3FF264157FE5EFEF1" box="[228,466,301,325]" pageId="28" pageNumber="475">Wandege M. Muninga</collectorName>
and
<collectorName id="26EFDD95FFCBFFE3FDCC4157FD3EFEF1" box="[526,690,301,325]" pageId="28" pageNumber="475">Eli Greenbaum</collectorName>
</materialsCitation>
.
<emphasis id="B96E6451FFCBFFE3FD024157FD47FEF1" bold="true" box="[704,715,301,325]" pageId="28" pageNumber="475">•</emphasis>
<materialsCitation id="3B72B21EFFCBFFE3FD144157FEABFED0" country="Democratic Republic of the Congo" location="DRC" pageId="28" pageNumber="475" specimenCount="1" typeStatus="paratype">
<emphasis id="B96E6451FFCBFFE3FD144157FC85FEF0" box="[726,777,301,324]" italics="true" pageId="28" pageNumber="475">
<collectingCountry id="F30DF8D3FFCBFFE3FD144157FC85FEF0" box="[726,777,301,324]" name="Democratic Republic of the Congo" pageId="28" pageNumber="475">DRC</collectingCountry>
</emphasis>
(
<specimenCount id="9D1C73CAFFCBFFE3FF4A4136FEACFED0" box="[136,288,332,356]" count="1" pageId="28" pageNumber="475" type="generic">one specimen</specimenCount>
)
</materialsCitation>
:
<materialsCitation id="3B72B21EFFCBFFE3FEF34136FF58FE76" collectingDate="2008-06-15" collectionCode="UTEP, EBG" country="DRC" elevation="1260" latitude="-2.71471" location="Bukavu-Uvira" longLatPrecision="1" longitude="28.94641" municipality="N'Komo River" pageId="28" pageNumber="475" specimenCode="UTEP 22586, EBG 1556" specimenCount="1" stateProvince="Sud-Kivu" typeStatus="paratype">
<specimenCode id="DBBC1038FFCBFFE3FEF34136FE31FED0" box="[305,445,332,356]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="28" pageNumber="475" type="Museum">UTEP 22586</specimenCode>
(
<specimenCode id="DBBC1038FFCBFFE3FE094136FDCDFED0" box="[459,577,332,356]" collectionCode="EBG" pageId="28" pageNumber="475">EBG 1556</specimenCode>
), male, collected in a gallery forest at
<collectingMunicipality id="6BC12239FFCBFFE3FE874111FE6FFE37" box="[325,483,363,387]" pageId="28" pageNumber="475">N’Komo River</collectingMunicipality>
, road.
<location id="8EC5EE98FFCBFFE3FDEC4111FD32FE37" LSID="urn:lsid:plazi:treatment:03B30955FFCCFFE2FC6E4462FF66F92B:8EC5EE98FFCBFFE3FDEC4111FD32FE37" box="[558,702,363,387]" country="DRC" latitude="-2.71471" longLatPrecision="1" longitude="28.94641" municipality="N'Komo River" name="Bukavu-Uvira" pageId="28" pageNumber="475" stateProvince="Sud-Kivu">Bukavu-Uvira</location>
,
<collectingRegion id="49DE76A1FFCBFFE3FD0E4111FE98FE17" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="28" pageNumber="475">South Kivu Province</collectingRegion>
,
<geoCoordinate id="EE2EDE84FFCBFFE3FEDC41F1FE01FE17" box="[286,397,395,419]" degrees="02.71471" direction="south" orientation="latitude" pageId="28" pageNumber="475" precision="1" value="-2.71471">S02.71471</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFCBFFE3FE5A41F0FD85FE16" box="[408,521,394,418]" degrees="28.94641" direction="east" orientation="longitude" pageId="28" pageNumber="475" precision="1" value="28.94641">E28.94641</geoCoordinate>
,
<elevation id="00375F70FFCBFFE3FDD141F0FD18FE17" box="[531,660,394,419]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="28" pageNumber="475" unit="m" value="1260.0">
<quantity id="4CE215A6FFCBFFE3FDD141F0FDEEFE16" box="[531,610,394,418]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="28" pageNumber="475" unit="m" value="1260.0">1260 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFCBFFE3FD7941F0FF39FE76" pageId="28" pageNumber="475" value="2008-06-15">
<collectingDate id="EFE0676BFFCBFFE3FD7941F0FF39FE76" pageId="28" pageNumber="475" value="2008-06-15">15 June 2008</collectingDate>
</date>
by
</materialsCitation>
<materialsCitation id="3B72B21EFFCBFFE3FF1941D0FD81FDB5" collectorName="Luhumyo &amp; Chifundera Kusamba &amp; Mwenebatu M. Aristote &amp; Wandege M. Muninga" country="Mauritius" location="Maurice" pageId="28" pageNumber="475" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F30DF8D3FFCBFFE3FF1941D0FEBCFE76" box="[219,304,426,450]" name="Mauritius" pageId="28" pageNumber="475">Maurice</collectingCountry>
<collectorName id="26EFDD95FFCBFFE3FEF541D0FE10FE76" box="[311,412,426,450]" pageId="28" pageNumber="475">Luhumyo</collectorName>
,
<collectorName id="26EFDD95FFCBFFE3FE6541D0FD09FE76" box="[423,645,426,450]" pageId="28" pageNumber="475">Chifundera Kusamba</collectorName>
,
<collectorName id="26EFDD95FFCBFFE3FD5241D0FF76FE55" pageId="28" pageNumber="475">Mwenebatu M. Aristote</collectorName>
,
<collectorName id="26EFDD95FFCBFFE3FEC741B3FE7CFE55" box="[261,496,457,481]" pageId="28" pageNumber="475">Wandege M. Muninga</collectorName>
, John Akuku, Felix Akuku, Asukulu M’Mema, and Eli Greenbaum
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCBFFE3FF434259FA28FED7" pageId="28" pageNumber="475" type="diagnosis">
<paragraph id="8BA5B843FFCBFFE3FF434259FF65FC27" blockId="28.[129,778,546,1228]" pageId="28" pageNumber="475">
<emphasis id="B96E6451FFCBFFE3FF434259FF6BFD8E" box="[129,231,547,570]" italics="true" pageId="28" pageNumber="475">Diagnosis:</emphasis>
A large
<taxonomicName id="4C1AC3C0FFCBFFE3FE9D4258FE52FD8E" box="[351,478,546,570]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="genus">
<emphasis id="B96E6451FFCBFFE3FE9D4258FE52FD8E" box="[351,478,546,570]" italics="true" pageId="28" pageNumber="475">Lygodactylus</emphasis>
</taxonomicName>
[maximum
<collectionCode id="ED0B2086FFCBFFE3FDA94258FD14FD8E" box="[619,664,546,570]" pageId="28" pageNumber="475">SVL</collectionCode>
<quantity id="4CE215A6FFCBFFE3FD644258FC85FD8D" box="[678,777,546,570]" metricMagnitude="-2" metricUnit="m" metricValue="3.77" pageId="28" pageNumber="475" unit="mm" value="37.7">37.7 mm</quantity>
(mean 34.7 ±
<quantity id="4CE215A6FFCBFFE3FED4423BFEE2FDED" box="[278,366,577,601]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="475" unit="mm" value="2.5">2.5 mm</quantity>
)], that shares a similar distinctive gular chevron ornamentation with the
<taxonomicName id="4C1AC3C0FFCBFFE3FE1D421BFDDBFDCD" box="[479,599,609,633]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCBFFE3FE1D421BFDDBFDCD" box="[479,599,609,633]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
</taxonomicName>
subgroup. It has 7–9 supralabials and 5–7 infralabials. Dorsal pholidosis with granular scales that become flattened, larger, and imbricate on original tails. Large triangular mental followed by usually three (occasionally two) symmetrical postmental scales (Supporting Information,
<figureCitation id="1321A4C6FFCBFFE3FECD4287FED7FCA1" box="[271,347,765,789]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="28" pageNumber="475">Fig. S6</figureCitation>
). Nostril never in contact with rostral. Ventral pholidosis with large, flattened, imbricate scales. Five to six terminal scansors on the tail tip. Digits elongated with five terminal scansors on the fourth toe (Supporting Information, Table S7).
</paragraph>
<paragraph id="8BA5B843FFCBFFE3FF5E43E0FD21FB78" blockId="28.[129,778,546,1228]" pageId="28" pageNumber="475">
This species may be easily differentiated from the
<taxonomicName id="4C1AC3C0FFCBFFE3FD5443E1FC85FC06" authorityName="Gunther" authorityYear="1893" box="[662,777,922,946]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFCBFFE3FD5443E1FC85FC06" box="[662,777,922,946]" italics="true" pageId="28" pageNumber="475">L. angularis</emphasis>
</taxonomicName>
group by the characteristic
<emphasis id="B96E6451FFCBFFE3FE5343C3FE2EFC65" bold="true" box="[401,418,953,977]" pageId="28" pageNumber="475">∩</emphasis>
<emphasis id="B96E6451FFCBFFE3FE6043C0FE25FC65" box="[418,425,954,977]" italics="true" pageId="28" pageNumber="475">-</emphasis>
shaped gular pattern as
<taxonomicName id="4C1AC3C0FFCBFFE3FD5743C0FC85FC65" box="[661,777,953,977]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCBFFE3FD5743C0FC85FC65" box="[661,777,953,977]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
</taxonomicName>
(see
<taxonomicName id="4C1AC3C0FFCBFFE3FF7443A3FEBCFC44" box="[182,304,984,1008]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCBFFE3FF7443A3FEBCFC44" box="[182,304,984,1008]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
</taxonomicName>
diagnosis). It can be differentiated from
<taxonomicName id="4C1AC3C0FFCBFFE3FD3743A3FF7CFBA4" baseAuthorityName="Laurent" baseAuthorityYear="1952" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="paurospilus">
<emphasis id="B96E6451FFCBFFE3FD3743A3FF7CFBA4" italics="true" pageId="28" pageNumber="475">L. paurospilus</emphasis>
</taxonomicName>
, found in the same region, on the basis of three
<emphasis id="B96E6451FFCBFFE3FF43446DFF1EFB9B" bold="true" box="[129,146,1047,1071]" pageId="28" pageNumber="475">∩</emphasis>
-shaped thick chevrons reaching the chest vs. two
<emphasis id="B96E6451FFCBFFE3FD6B4462FD35FB9B" box="[681,697,1048,1071]" italics="true" pageId="28" pageNumber="475">
<collectionCode id="ED0B2086FFCBFFE3FD6B4462FD35FB9B" box="[681,697,1048,1071]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="28" pageNumber="475" type="Museum">V</collectionCode>
</emphasis>
-shaped broken gular chevrons in
<taxonomicName id="4C1AC3C0FFCBFFE3FE48444CFE75FBFA" baseAuthorityName="Laurent" baseAuthorityYear="1952" box="[394,505,1078,1102]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="paurospilus">
<emphasis id="B96E6451FFCBFFE3FE48444CFE75FBFA" box="[394,505,1078,1102]" italics="true" pageId="28" pageNumber="475">paurospilus</emphasis>
</taxonomicName>
; and by having a reduced, almost vestigial postorbitofrontal bone vs. well-developed postorbitofrontal in
<taxonomicName id="4C1AC3C0FFCBFFE3FE96440CFE53FB39" baseAuthorityName="Laurent" baseAuthorityYear="1952" box="[340,479,1141,1165]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="paurospilus">
<emphasis id="B96E6451FFCBFFE3FE96440CFE53FB39" box="[340,479,1141,1165]" italics="true" pageId="28" pageNumber="475">L. paurospilus</emphasis>
</taxonomicName>
. It can also be differentiated from other members of the
<taxonomicName id="4C1AC3C0FFCBFFE3FE6344EFFD91FB18" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[417,541,1173,1196]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFCBFFE3FE6344EFFD91FB18" box="[417,541,1173,1196]" italics="true" pageId="28" pageNumber="475">L. picturatus</emphasis>
</taxonomicName>
group based on dorsal coloration and gular pattern (see
<taxonomicName id="4C1AC3C0FFCBFFE3FE1344CEFDC9FB78" box="[465,581,1204,1228]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCBFFE3FE1344CEFDC9FB78" box="[465,581,1204,1228]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
</taxonomicName>
account).
</paragraph>
<paragraph id="8BA5B843FFCBFFE3FF434497FA28FED7" blockId="28.[129,778,1261,1974]" lastBlockId="28.[825,1474,143,355]" pageId="28" pageNumber="475">
<taxonomicName id="4C1AC3C0FFCBFFE3FF434497FEDEFAB1" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[129,338,1261,1285]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">Lygodactylus kibera</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFCBFFE3FE9E4494FE20FAB1" box="[348,428,1262,1285]" pageId="28" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
can be differentiated from other species within the
<taxonomicName id="4C1AC3C0FFCBFFE3FE8C4577FE45FA90" box="[334,457,1292,1316]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCBFFE3FE8C4577FE45FA90" box="[334,457,1292,1316]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
</taxonomicName>
subgroup by subtle morphometric and meristic features. This species is best regarded as cryptic, but we provide some characters that are diagnostically useful.
<emphasis id="B96E6451FFCBFFE3FF0C4510FD89FA36" bold="true" box="[206,517,1386,1410]" pageId="28" pageNumber="475">
<taxonomicName id="4C1AC3C0FFCBFFE3FF0C4510FE2AFA36" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[206,422,1386,1410]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFCBFFE3FF0C4510FE2AFA36" bold="true" box="[206,422,1386,1410]" italics="true" pageId="28" pageNumber="475">Lygodactylus kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFCBFFE3FE6D4510FD89FA36" box="[431,517,1386,1410]" pageId="28" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C1AC3C0FFCBFFE3FD564511FC86FA36" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[660,778,1386,1410]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFCBFFE3FD564511FC86FA36" box="[660,778,1386,1410]" italics="true" pageId="28" pageNumber="475">L. dysmicus</emphasis>
</taxonomicName>
by its larger size (maximum
<collectionCode id="ED0B2086FFCBFFE3FE6545F0FE58FA16" box="[423,468,1418,1442]" pageId="28" pageNumber="475">SVL</collectionCode>
<quantity id="4CE215A6FFCBFFE3FE1945F0FDB4FA15" box="[475,568,1418,1442]" metricMagnitude="-2" metricUnit="m" metricValue="3.77" pageId="28" pageNumber="475" unit="mm" value="37.7">37.7 mm</quantity>
vs.
<quantity id="4CE215A6FFCBFFE3FDA345F0FD4FFA16" box="[609,707,1418,1442]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="28" pageNumber="475" unit="mm" value="27.6">27.6 mm</quantity>
); gular patterning always with three
<emphasis id="B96E6451FFCBFFE3FE6E45D3FE31FA75" bold="true" box="[428,445,1449,1473]" pageId="28" pageNumber="475">∩</emphasis>
<emphasis id="B96E6451FFCBFFE3FE7F45D0FE48FA75" box="[445,452,1450,1473]" italics="true" pageId="28" pageNumber="475">-</emphasis>
shaped thick chevrons reaching the chest (vs. two thinner
<emphasis id="B96E6451FFCBFFE3FE4845B2FE17FA54" bold="true" box="[394,411,1480,1504]" pageId="28" pageNumber="475">∩</emphasis>
-shaped chevrons, that never extend beyond the posterior part of the lower jaw); fewer precloacal pores (7–8 vs. 9); usually three symmetrical postmental scales vs. two; nostril never in contact with rostral scale (vs. nostril contacting rostral); and lower number of ventral scales across the body (16–18 vs.
<quantity id="4CE215A6FFCBFFE3FE9B461FFE02F9C9" box="[345,398,1637,1661]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="28" pageNumber="475" unit="in" value="21.0">21 in</quantity>
<taxonomicName id="4C1AC3C0FFCBFFE3FE57461CFD8BF9C9" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[405,519,1637,1661]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFCBFFE3FE57461CFD8BF9C9" box="[405,519,1637,1661]" italics="true" pageId="28" pageNumber="475">L. dysmicus</emphasis>
</taxonomicName>
). It can be distinguished from
<emphasis id="B96E6451FFCBFFE3FF7946FFFEDAF928" box="[187,342,1668,1692]" italics="true" pageId="28" pageNumber="475">
<taxonomicName id="4C1AC3C0FFCBFFE3FF7946FFFEBEF928" box="[187,306,1668,1692]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">L. gutturalis</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFCBFFE3FEF946FFFEDAF928" box="[315,342,1669,1692]" pageId="28" pageNumber="475" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
by its slightly larger size [maximum
<collectionCode id="ED0B2086FFCBFFE3FD1E46FEFC85F928" box="[732,777,1668,1692]" pageId="28" pageNumber="475">SVL</collectionCode>
<quantity id="4CE215A6FFCBFFE3FF4346DEFF51F90F" box="[129,221,1700,1724]" metricMagnitude="-2" metricUnit="m" metricValue="3.77" pageId="28" pageNumber="475" unit="mm" value="37.7">37.7 mm</quantity>
(mean 34.7 ±
<quantity id="4CE215A6FFCBFFE3FEB746D9FE4AF908" box="[373,454,1699,1724]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="475" unit="mm" value="2.5">2.5 mm</quantity>
) vs.
<quantity id="4CE215A6FFCBFFE3FE3746D9FDDDF90F" box="[501,593,1699,1723]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="28" pageNumber="475" unit="mm" value="36.2">36.2 mm</quantity>
(mean 33.3 ±
<quantity id="4CE215A6FFCBFFE3FD2846DEFF3EF96F" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="28" pageNumber="475" unit="mm" value="1.8">1.8 mm</quantity>
)]; snout proportionally narrower (
<collectionCode id="ED0B2086FFCBFFE3FDDF46B9FDB0F96F" box="[541,572,1731,1755]" pageId="28" pageNumber="475">IN</collectionCode>
/
<collectionCode id="ED0B2086FFCBFFE3FD8646B9FDF9F96F" box="[580,629,1731,1755]" pageId="28" pageNumber="475">HW</collectionCode>
0.20–0.27 vs. 0.29–0.34) with usually one large internasal scale vs. two smaller internasal scales (this character has shown to be variable in both species). It also differs from
<taxonomicName id="4C1AC3C0FFCBFFE3FE654758FD91F88D" box="[423,541,1825,1849]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCBFFE3FE654758FD91F88D" box="[423,541,1825,1849]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1AC3C0FFCBFFE3FD904758FD44F88D" authorityName=", Schmidt" authorityYear="1919" box="[594,712,1825,1849]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCBFFE3FD904758FD44F88D" box="[594,712,1825,1849]" italics="true" pageId="28" pageNumber="475">L. depressus</emphasis>
</taxonomicName>
based on gular pattern always having three thick
<emphasis id="B96E6451FFCBFFE3FD81473AFDD8F8EC" bold="true" box="[579,596,1856,1880]" pageId="28" pageNumber="475">∩</emphasis>
-shaped chevrons reaching the chest [vs. two or three (in
<taxonomicName id="4C1AC3C0FFCBFFE3FDD8471AFD1CF8C3" box="[538,656,1887,1911]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCBFFE3FDD8471AFD1CF8C3" box="[538,656,1887,1911]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
</taxonomicName>
), or one or two (in
<taxonomicName id="4C1AC3C0FFCBFFE3FF1747FAFEC6F823" authorityName=", Schmidt" authorityYear="1919" box="[213,330,1919,1943]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCBFFE3FF1747FAFEC6F823" box="[213,330,1919,1943]" italics="true" pageId="28" pageNumber="475">L. depressus</emphasis>
</taxonomicName>
) thinner
<emphasis id="B96E6451FFCBFFE3FE724705FE4DF823" bold="true" box="[432,449,1919,1943]" pageId="28" pageNumber="475">∩</emphasis>
-shaped chevrons that never extend beyond the posterior part of the lower jaw]; proportionally more elongated head (HL/
<collectionCode id="ED0B2086FFCBFFE3FB8D40EAFB0EFF1C" box="[1103,1154,144,168]" pageId="28" pageNumber="475">SVL</collectionCode>
0.27–0.30 vs. 0.24–0.26) with larger eyes (
<collectionCode id="ED0B2086FFCBFFE3FC7140D5FC51FF73" box="[947,989,175,199]" pageId="28" pageNumber="475">OD</collectionCode>
/HL 0.24–0.27 vs. 0.20–0.23) than
<taxonomicName id="4C1AC3C0FFCBFFE3FA8C40CAFA4EFF73" authorityName=", Schmidt" authorityYear="1919" box="[1358,1474,175,199]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFCBFFE3FA8C40CAFA4EFF73" box="[1358,1474,175,199]" italics="true" pageId="28" pageNumber="475">L. depressus</emphasis>
</taxonomicName>
(
<figureCitation id="1321A4C6FFCBFFE3FC8640B4FCF0FF52" box="[836,892,206,230]" captionStart="Figure 4" captionStartId="15.[113,178,747,771]" captionTargetBox="[117,1455,147,717]" captionTargetId="figure-570@15.[114,1458,144,720]" captionTargetPageId="15" captionText="Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section." pageId="28" pageNumber="475">Fig. 4</figureCitation>
;
<tableCitation id="C6988DF8FFCBFFE3FC4B40B4FC5FFF52" box="[905,979,206,230]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="28" pageNumber="475">Table 3</tableCitation>
). The new species occurs in mid-elevation moist forest, agricultural fields, and human habitations, vs. lowland dry sub-Saharan savannah (
<taxonomicName id="4C1AC3C0FFCBFFE3FBF34174FB24FE91" box="[1073,1192,269,293]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFCBFFE3FBF34174FB24FE91" box="[1073,1192,269,293]" italics="true" pageId="28" pageNumber="475">L. gutturalis</emphasis>
</taxonomicName>
) and lowland rainforest of the
<collectingCountry id="F30DF8D3FFCBFFE3FCA24156FC2AFEF0" box="[864,934,300,324]" name="Democratic Republic of the Congo" pageId="28" pageNumber="475">Congo</collectingCountry>
Basin (
<taxonomicName id="4C1AC3C0FFCBFFE3FC364157FBE9FEF0" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[1012,1125,300,324]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFCBFFE3FC364157FBE9FEF0" box="[1012,1125,300,324]" italics="true" pageId="28" pageNumber="475">L. dysmicus</emphasis>
</taxonomicName>
; see
<figureCitation id="1321A4C6FFCBFFE3FB554156FB43FEF0" box="[1175,1231,300,324]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="28" pageNumber="475">Fig. 6</figureCitation>
). For a distinction with other species described below, see their respective diagnoses.
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCBFFE3FCFB41FFFA4DFE4F" pageId="28" pageNumber="475" type="etymology">
<paragraph id="8BA5B843FFCBFFE3FCFB41FFFA4DFE4F" blockId="28.[825,1474,389,507]" pageId="28" pageNumber="475">
<emphasis id="B96E6451FFCBFFE3FCFB41FFFC24FE29" box="[825,936,389,413]" italics="true" pageId="28" pageNumber="475">Etymology:</emphasis>
The name
<taxonomicName id="4C1AC3C0FFCBFFE3FBE641FFFBEAFE29" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1060,1126,389,413]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="475" phylum="Chordata" rank="species" species="kibera">
‘
<emphasis id="B96E6451FFCBFFE3FBE841FFFBEAFE29" box="[1066,1126,389,413]" italics="true" pageId="28" pageNumber="475">kibera</emphasis>
</taxonomicName>
’ derives from the word ‘kibira’ or ‘kibera’ in Kinubi—a Sudanese Arabic-based creole language spoken in some regions of
<collectingCountry id="F30DF8D3FFCBFFE3FB9341B9FB24FE6F" box="[1105,1192,451,475]" name="Burundi" pageId="28" pageNumber="475">Burundi</collectingCountry>
,
<collectingCountry id="F30DF8D3FFCBFFE3FB7641BEFB7BFE68" box="[1204,1271,452,476]" name="Kenya" pageId="28" pageNumber="475">Kenya</collectingCountry>
, and
<collectingCountry id="F30DF8D3FFCBFFE3FAED41B9FA0CFE6F" box="[1327,1408,451,475]" name="Uganda" pageId="28" pageNumber="475">Uganda</collectingCountry>
—that means ‘forest’, the main habitat
<typeStatus id="54A106E1FFCBFFE3FBBA4199FB2AFE4F" box="[1144,1190,483,507]" pageId="28" pageNumber="475">type</typeStatus>
associated with the species.
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFCBFFE2FCFB4267FE56F9BC" lastPageId="29" lastPageNumber="476" pageId="28" pageNumber="475" type="description">
<paragraph id="8BA5B843FFCBFFE3FCFB4267FBDAFA15" blockId="28.[825,1475,540,1441]" pageId="28" pageNumber="475">
<emphasis id="B96E6451FFCBFFE3FCFB4267FBD7FD80" box="[825,1115,540,564]" italics="true" pageId="28" pageNumber="475">
Description of the
<typeStatus id="54A106E1FFCBFFE3FBC54266FBD7FD80" box="[1031,1115,540,564]" pageId="28" pageNumber="475" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="1321A4C6FFCBFFE3FB414266FB5EFD80" box="[1155,1234,540,564]" captionStart="Figure 17" captionStartId="29.[113,178,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-359@29.[114,1458,144,1094]" captionTargetPageId="29" captionText="Figure 17. Holotype of Lygodactylus kibera sp. nov. (UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus." pageId="28" pageNumber="475">Fig. 17</figureCitation>
): Measurements and meristic characters of the
<typeStatus id="54A106E1FFCBFFE3FB834241FB13FDE7" box="[1089,1183,571,595]" pageId="28" pageNumber="475" type="holotype">holotype</typeStatus>
are presented in Supporting Information, Table S7. Adult male, with a snout–vent length (
<collectionCode id="ED0B2086FFCBFFE3FC834200FCF9FD26" box="[833,885,634,658]" pageId="28" pageNumber="475">SVL</collectionCode>
) of
<quantity id="4CE215A6FFCBFFE3FC654200FB85FD26" box="[935,1033,634,658]" metricMagnitude="-2" metricUnit="m" metricValue="3.69" pageId="28" pageNumber="475" unit="mm" value="36.9">36.9 mm</quantity>
and a slightly larger original tail length (TL =
<quantity id="4CE215A6FFCBFFE3FC4142E3FC60FD06" box="[899,1004,665,690]" metricMagnitude="-2" metricUnit="m" metricValue="4.06" pageId="28" pageNumber="475" unit="mm" value="40.6">40.6 mm</quantity>
). Body slender, nape moderately distinct. Head as broad as body, and moderate head length (
<collectionCode id="ED0B2086FFCBFFE3FAA342C3FA1EFD65" box="[1377,1426,697,721]" pageId="28" pageNumber="475">HW</collectionCode>
/HL 0.70). Canthus rostralis not prominent. Eye diameter
<quantity id="4CE215A6FFCBFFE3FAAE42A2FA32FD44" box="[1388,1470,728,752]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="475" unit="mm" value="2.5">2.5 mm</quantity>
, with circular pupil. Ear to eye distance slightly larger than orbit diameter (
<quantity id="4CE215A6FFCBFFE3FC1C436DFBBAFC9B" box="[990,1078,791,815]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="28" pageNumber="475" unit="mm" value="3.7">3.7 mm</quantity>
). Snout rounded and slightly pointed. Frontal granular scales larger than occipital scales. Dorsal scales granular from rostral to tail. Rostral undivided, in contact with 1st supralabial, prenasals, and one large internasal scale. Eight to nine supralabials and six infralabials. Prenasal scale in contact with 1st supralabial. Nostril circular, bordered by 1st supralabial, prenasal, one supranasal, and one postnasal. Four rows of scales between supralabials and the orbit. Mental large, triangular, and rounded posteriorly, with two large rounded postmental scales separated by one small rounded postmental scale. Five post-postmental scales. Gular scales granular, rounded, and slightly smaller than ventral scales. Ventral scales large, imbricate, with 17 scales rows across the venter. Body relatively robust and slightly elongated (
<collectionCode id="ED0B2086FFCBFFE3FB6F44D4FB6EFB72" box="[1197,1250,1198,1222]" pageId="28" pageNumber="475">TRL</collectionCode>
/
<collectionCode id="ED0B2086FFCBFFE3FB2944D4FA92FB72" box="[1259,1310,1198,1222]" pageId="28" pageNumber="475">SVL</collectionCode>
0.43). Tail with 51 enlarged transverse scales and six pairs of terminal scansors on the tip. Seven precloacal pores. Fore- and hind limbs moderately short, stout; forearm medium-sized (
<collectionCode id="ED0B2086FFCBFFE3FAD94576FAB0FA90" box="[1307,1340,1292,1316]" pageId="28" pageNumber="475">FL</collectionCode>
/
<collectionCode id="ED0B2086FFCBFFE3FA864576FAFBFA90" box="[1348,1399,1292,1316]" pageId="28" pageNumber="475">SVL</collectionCode>
0.14); tibia short (CL/
<collectionCode id="ED0B2086FFCBFFE3FC244551FB96FAF7" box="[998,1050,1323,1347]" pageId="28" pageNumber="475">SVL</collectionCode>
0.16). Digits elongated and unwebbed with 5–6 terminal scansors. Thumb rudimentary with a small claw. Relative length of digits: I &lt;II =
<collectionCode id="ED0B2086FFCBFFE3FB154510FB64FA35" box="[1239,1256,1386,1409]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="28" pageNumber="475" type="Museum">V</collectionCode>
&lt;III &lt;IV (manus); I &lt;II &lt;
<collectionCode id="ED0B2086FFCBFFE3FC4D45F0FC2CFA15" box="[911,928,1418,1441]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="28" pageNumber="475" type="Museum">V</collectionCode>
&lt;III &lt;IV (pes).
</paragraph>
<paragraph id="8BA5B843FFCBFFE3FCFB45B2FA1BF94E" blockId="28.[825,1474,1480,1786]" pageId="28" pageNumber="475">
<emphasis id="B96E6451FFCBFFE3FCFB45B2FC74FA54" box="[825,1016,1480,1504]" italics="true" pageId="28" pageNumber="475">Coloration: In life</emphasis>
(
<figureCitation id="1321A4C6FFCBFFE3FBCE45B2FBDAFA55" box="[1036,1110,1480,1505]" captionStart="Figure 18" captionStartId="30.[129,194,1128,1152]" captionTargetBox="[129,1473,144,1100]" captionTargetId="figure-444@30.[129,1473,144,1100]" captionTargetPageId="30" captionText="Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D)." pageId="28" pageNumber="475">Fig. 18</figureCitation>
), dorsal coloration brownish grey with light cream-beige lines from nape to tail on each side of the dorsum interspersed by five or six lighter cream dots surrounded by black flanks. Black line from nostril to the anterior insertion of the forelimb. Gular region with white coloration and three black
<emphasis id="B96E6451FFCBFFE3FCFB461FFCC6F9C9" bold="true" box="[825,842,1637,1661]" pageId="28" pageNumber="475">∩</emphasis>
-shaped chevrons. First and second chevrons in contact. Venter uniform orange from second chevrons to anterior portion of the tail and extending onto the hind limbs. Ventral surface of tail tip and digits whitish.
<emphasis id="B96E6451FFCBFFE3FBC646BEFB18F96F" box="[1028,1172,1732,1755]" italics="true" pageId="28" pageNumber="475">In preservative</emphasis>
(
<typeStatus id="54A106E1FFCBFFE3FB6746B9FA85F96F" box="[1189,1289,1731,1755]" pageId="28" pageNumber="475" type="holotype">holotype</typeStatus>
;
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) dorsum dark brown and venter with uniform light cream coloration.
</paragraph>
<paragraph id="8BA5B843FFCBFFE2FCFB475AFE56F9BC" blockId="28.[825,1474,1823,1973]" lastBlockId="29.[113,763,1300,1544]" lastPageId="29" lastPageNumber="476" pageId="28" pageNumber="475">
<emphasis id="B96E6451FFCBFFE3FCFB475AFC10F883" box="[825,924,1824,1847]" italics="true" pageId="28" pageNumber="475">Variation:</emphasis>
The meristic characters of the head and body of this species are variable (see Supporting Information, Table S7). Coloration of this species seems to be consistent, with the gular coloration slightly lighter in females. First and second gular chevrons may be in contact or not (Supporting Information,
<figureCitation id="1321A4C6FFCBFFE3FAAF47E6FA3DF800" box="[1389,1457,1948,1972]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="28" pageNumber="475">Fig. S6</figureCitation>
).
<emphasis id="B96E6451FFCAFFE2FFB3456FFEE2FA99" box="[113,366,1301,1325]" italics="true" pageId="29" pageNumber="476">Habitat and distribution</emphasis>
(
<figureCitation id="1321A4C6FFCAFFE2FE50456FFE59FA99" box="[402,469,1301,1325]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="29" pageNumber="476">Figs 6</figureCitation>
,
<figureCitation id="1321A4C6FFCAFFE2FE25456FFD8DFA99" box="[487,513,1301,1325]" captionStart="Figure 18" captionStartId="30.[129,194,1128,1152]" captionTargetBox="[129,1473,144,1100]" captionTargetId="figure-444@30.[129,1473,144,1100]" captionTargetPageId="30" captionText="Figure 18. Specimens of Lygodactylus kibera sp. nov. in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi.Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D)." pageId="29" pageNumber="476">18</figureCitation>
):
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<emphasis id="B96E6451FFCAFFE2FDE2456EFD76FA98" bold="true" box="[544,762,1300,1324]" italics="true" pageId="29" pageNumber="476">Lygodactylus kibera</emphasis>
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is known from the Albertine Rift adjacent to Lake Tanganyika in
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and
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. Specimens from Mpishi,
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, were collected near montane forest in banana fields. The specimen from
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,
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, was collected inside a hotel in the middle of the city. The specimen from N’Komo River,
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, was found on a tree near a small river between secondary gallery forest and grassland.
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<paragraph id="8BA5B843FFCAFFE2FFB34418FE4AFB06" blockId="29.[113,1442,1122,1202]" pageId="29" pageNumber="476">
<emphasis id="B96E6451FFCAFFE2FFB34418FF5AFBCE" bold="true" box="[113,214,1122,1146]" pageId="29" pageNumber="476">Figure 17.</emphasis>
Holotype of
<emphasis id="B96E6451FFCAFFE2FE904418FDE9FBCE" bold="true" box="[338,613,1122,1146]" pageId="29" pageNumber="476">
<taxonomicName id="4C1AC3C0FFCAFFE2FE904418FD9AFBCE" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[338,534,1122,1146]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFCAFFE2FE904418FD9AFBCE" bold="true" box="[338,534,1122,1146]" italics="true" pageId="29" pageNumber="476">Lygodactylus kibera</emphasis>
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</emphasis>
(UTEP 22566) from Mpishi, near Kibira National Park, Bubanza Province, Burundi. A, Dorsal and ventral view of body. B, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Details of cloacal region (C), (D) tail tip, and (E) left manus.
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<emphasis id="B96E6451FFCAFFE2FFB34653FE9FF9F5" box="[113,275,1577,1601]" italics="true" pageId="29" pageNumber="476">Natural history:</emphasis>
An arboreal species with diurnal habits frequently found on the trunk or branches of trees, and also around anthropogenically altered areas such as buildings and plantations.
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<subSubSection id="C300EBC8FFF3FFDBFF5C4499FD66FA39" pageId="36" pageNumber="483" type="description">
<paragraph id="8BA5B843FFF3FFDBFF5C4499FD60FB4F" blockId="36.[158,748,1251,1275]" box="[158,748,1251,1275]" pageId="36" pageNumber="483">
(
<figureCitation id="1321A4C6FFF3FFDBFF6B4499FE95FB4F" box="[169,281,1251,1275]" captionStart-0="Figure 23" captionStart-1="Figure 24" captionStartId-0="37.[113,178,1204,1228]" captionStartId-1="38.[129,194,1232,1256]" captionTargetBox-0="[114,1458,144,1176]" captionTargetBox-1="[129,1473,144,1205]" captionTargetId-0="figure-366@37.[114,1458,144,1176]" captionTargetId-1="figure-368@38.[129,1473,144,1205]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionText-0="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." captionText-1="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="36" pageNumber="483">Figs 23–24</figureCitation>
,
<tableCitation id="C6988DF8FFF3FFDBFEE64499FEE3FB4F" box="[292,367,1251,1275]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="36" pageNumber="483">Table 3</tableCitation>
; Supporting Information, Table S10)
</paragraph>
<paragraph id="8BA5B843FFF3FFDBFF434566FE38FAE7" blockId="36.[129,777,1308,1363]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFF434566FED2FA80" box="[129,350,1308,1332]" italics="true" pageId="36" pageNumber="483">Zoobank registration:</emphasis>
https://zoobank.org/
<uuid id="FFBC8296FFF3FFDBFD8E4566FE38FAE7" pageId="36" pageNumber="483">411D833D-2017- 4930-BD11-B783AAD6A9B7</uuid>
</paragraph>
<paragraph id="8BA5B843FFF3FFDBFF634509FD66FA39" blockId="36.[161,746,1395,1421]" box="[161,746,1395,1421]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFF634509FD66FA39" box="[161,746,1395,1421]" italics="true" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFF634509FE73FA39" authorityName="' (Schmidt" authorityYear="1919" box="[161,511,1395,1421]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="subSpecies" species="picturatus" subSpecies="gutturalis">Lygodactylus picturatus gutturalis</taxonomicName>
: de Witte (1953) [part].
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFF3FFDBFF5B45CBFD90F939" pageId="36" pageNumber="483" type="reference_group">
<paragraph id="8BA5B843FFF3FFDBFF5B45CBFD1AFA5F" blockId="36.[129,778,1457,1677]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFF5B45CBFD1AFA5F" italics="true" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFF5B45CBFE05FA7F" ID-CoL="3WQ3T" authority="(Bocage, 1873)" box="[153,393,1457,1483]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">Lygodactylus gutturalis</taxonomicName>
:
<bibRefCitation id="EF8BC5B2FFF3FFDBFE5745C8FDABFA7F" author="Pasteur G" box="[405,551,1457,1483]" pageId="36" pageNumber="483" pagination="1 - 132" refId="ref36328" refString="Pasteur G. Recherches sur l'evolution des lygodactyles, lezards Afro- Malgaches actuels. Travaux de l'Institut Scientifique Cherifien, Zoologie 1965 (1964); 29: 1 - 132." type="journal article" year="1964">Pasteur (1964)</bibRefCitation>
[part]; Broadley and Cotterill (2004) [part];
<bibRefCitation id="EF8BC5B2FFF3FFDBFE1E45ABFDDDFA5F" author="Roll B" box="[476,593,1489,1515]" pageId="36" pageNumber="483" pagination="645 - 8" refId="ref36800" refString="Roll B. Lygodactylus gutturalis (Bocage). Sauria (Suppl.) 2005; 27: 645 - 8." type="journal article" year="2005">Röll (2005)</bibRefCitation>
[part].
</emphasis>
</paragraph>
<paragraph id="8BA5B843FFF3FFDBFF43458DFD90F939" blockId="36.[129,778,1457,1677]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFF43458DFE41F9BB" bold="true" box="[129,461,1527,1551]" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFF43458DFEE6F9BB" ID-CoL="C7JCT" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[129,362,1527,1551]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFF43458DFEE6F9BB" bold="true" box="[129,362,1527,1551]" italics="true" pageId="36" pageNumber="483">Lygodactylus gamblei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFEB7458DFE41F9BB" box="[373,461,1527,1551]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a miombo-savannah form within subgroup C, described above.It also differs by a minimum of
<emphasis id="B96E6451FFF3FFDBFF5F464DFF2AF9FA" box="[157,166,1591,1614]" italics="true" pageId="36" pageNumber="483">c</emphasis>
. 9.96% for the
<emphasis id="B96E6451FFF3FFDBFE88464CFEFDF9FA" box="[330,369,1590,1614]" italics="true" pageId="36" pageNumber="483">16S</emphasis>
mitochondrial gene (Table 2) from its sister taxon
<emphasis id="B96E6451FFF3FFDBFF3E462CFE61F9D9" bold="true" box="[252,493,1621,1645]" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFF3E462CFE19F9D9" ID-CoL="C7JCY" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[252,405,1621,1645]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFF3E462CFE19F9D9" bold="true" box="[252,405,1621,1645]" italics="true" pageId="36" pageNumber="483">L. leopardinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFE58462FFE61F9D9" box="[410,493,1621,1645]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1321A4C6FFF3FFDBFE3F462FFDC7F9D9" box="[509,587,1621,1645]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability&gt; 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="36" pageNumber="483">Fig. 2A</figureCitation>
) and lacks nuclear haplotype sharing in
<emphasis id="B96E6451FFF3FFDBFE94460FFE1FF939" box="[342,403,1653,1677]" italics="true" pageId="36" pageNumber="483">RAG1</emphasis>
(
<figureCitation id="1321A4C6FFF3FFDBFE61460FFD87F939" box="[419,523,1653,1677]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability&gt; 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="36" pageNumber="483">Fig. 2B–C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFF3FFD9FF4346D4FC56F8A3" lastPageId="38" lastPageNumber="485" pageId="36" pageNumber="483" type="materials_examined">
<paragraph id="8BA5B843FFF3FFDBFF4346D4FE72F890" blockId="36.[129,778,1710,1828]" pageId="36" pageNumber="483">
<materialsCitation id="3B72B21EFFF3FFDBFF4346D4FE72F890" collectingDate="2010-01-19" collectionCode="UTEP, ELI" country="Democratic Republic of the Congo" elevation="634" latitude="-7.29363" location="Manono" longLatPrecision="1" longitude="27.39472" municipality="Tanganyika Province" pageId="36" pageNumber="483" specimenCode="UTEP 22587, ELI 340" specimenCount="1" typeStatus="holotype">
<emphasis id="B96E6451FFF3FFDBFF4346D4FF6DF972" box="[129,225,1710,1734]" italics="true" pageId="36" pageNumber="483">
<typeStatus id="54A106E1FFF3FFDBFF4346D4FF50F972" box="[129,220,1710,1734]" pageId="36" pageNumber="483" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DBBC1038FFF3FFDBFF3246D4FEF0F972" box="[240,380,1710,1734]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="36" pageNumber="483" type="Museum">UTEP 22587</specimenCode>
(
<specimenCode id="DBBC1038FFF3FFDBFE4846D4FE64F972" box="[394,488,1710,1734]" collectionCode="ELI" pageId="36" pageNumber="483">ELI 340</specimenCode>
), male with regenerated tail and ventral incision, collected at
<location id="8EC5EE98FFF3FFDBFE0F46B4FDAAF951" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFE0F46B4FDAAF951" box="[461,550,1742,1765]" country="Democratic Republic of the Congo" latitude="-7.29363" longLatPrecision="1" longitude="27.39472" municipality="Tanganyika Province" name="Manono" pageId="36" pageNumber="483">Manono</location>
,
<collectingMunicipality id="6BC12239FFF3FFDBFDEC46B7FC89F951" box="[558,773,1741,1765]" pageId="36" pageNumber="483">Tanganyika Province</collectingMunicipality>
,
<collectingCountry id="F30DF8D3FFF3FFDBFF434697FF35F8B1" box="[129,185,1773,1797]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
,
<geoCoordinate id="EE2EDE84FFF3FFDBFF024697FEBCF8B1" box="[192,304,1773,1797]" degrees="07.29363" direction="south" orientation="latitude" pageId="36" pageNumber="483" precision="1" value="-7.29363">S07.29363</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFF3FFDBFEFA4697FE26F8B1" box="[312,426,1773,1797]" degrees="27.39472" direction="east" orientation="longitude" pageId="36" pageNumber="483" precision="1" value="27.39472">E27.39472</geoCoordinate>
,
<elevation id="00375F70FFF3FFDBFE704697FDAFF8B1" box="[434,547,1773,1797]" metricMagnitude="2" metricUnit="m" metricValue="6.34" pageId="36" pageNumber="483" unit="m" value="634.0">
<quantity id="4CE215A6FFF3FFDBFE704697FE7FF8B0" box="[434,499,1773,1797]" metricMagnitude="2" metricUnit="m" metricValue="6.34" pageId="36" pageNumber="483" unit="m" value="634.0">634 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFF3FFDBFD854697FD60F8B1" box="[583,748,1773,1797]" pageId="36" pageNumber="483" value="2010-01-19">
<collectingDate id="EFE0676BFFF3FFDBFD854697FD60F8B1" box="[583,748,1773,1797]" pageId="36" pageNumber="483" value="2010-01-19">19 January 2010</collectingDate>
</date>
by locals and brought to Eli Greenbaum.
</materialsCitation>
</paragraph>
<paragraph id="8BA5B843FFF3FFDBFF43473CFB58FCED" blockId="36.[129,778,1861,1980]" lastBlockId="36.[825,1475,144,858]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFF43473CFE1BF8E9" box="[129,407,1862,1885]" italics="true" pageId="36" pageNumber="483">
<typeStatus id="54A106E1FFF3FFDBFF43473CFF6CF8E9" box="[129,224,1862,1885]" pageId="36" pageNumber="483" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D1C73CAFFF3FFDBFF30473CFE07F8E9" box="[242,395,1862,1885]" count="9" pageId="36" pageNumber="483" type="generic">nine specimens</specimenCount>
):
</emphasis>
<materialsCitation id="3B72B21EFFF3FFDBFE6F473CFAE1FF1C" collectingDate="2010-01-18" collectionCode="UTEP, ELI" collectorName="Mulongo" country="Democratic Republic of the Congo" elevation="875" latitude="-7.65509" location="Haut-Lomami Province" longLatPrecision="1" longitude="27.34027" municipality="Aristote" pageId="36" pageNumber="483" specimenCode="UTEP 22584, ELI 293" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFF3FFDBFE6F473CFDB0F8E9" box="[429,572,1861,1885]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="36" pageNumber="483" type="Museum">UTEP 22584</specimenCode>
(
<specimenCode id="DBBC1038FFF3FFDBFD8C473FFD22F8E9" box="[590,686,1861,1885]" collectionCode="ELI" pageId="36" pageNumber="483">ELI 293</specimenCode>
), female with original tail and ventral incision, collected at
<collectorName id="26EFDD95FFF3FFDBFD65471FFC8AF8C9" box="[679,774,1893,1917]" pageId="36" pageNumber="483">Mulongo</collectorName>
,
<location id="8EC5EE98FFF3FFDBFF4347FEFEF4F828" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFF4347FEFEF4F828" box="[129,376,1924,1948]" country="Democratic Republic of the Congo" latitude="-7.65509" longLatPrecision="1" longitude="27.34027" name="Haut-Lomami Province" pageId="36" pageNumber="483">Haut-Lomami Province</location>
,
<collectingCountry id="F30DF8D3FFF3FFDBFE4747FEFE31F828" box="[389,445,1924,1948]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
,
<geoCoordinate id="EE2EDE84FFF3FFDBFE0847FEFDB6F828" box="[458,570,1924,1948]" degrees="07.65509" direction="south" orientation="latitude" pageId="36" pageNumber="483" precision="1" value="-7.65509">S07.65509</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFF3FFDBFD8447FEFD34F828" box="[582,696,1924,1948]" degrees="27.34027" direction="east" orientation="longitude" pageId="36" pageNumber="483" precision="1" value="27.34027">E27.34027</geoCoordinate>
,
<elevation id="00375F70FFF3FFDBFD0747FEFF21F80F" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="36" pageNumber="483" unit="m" value="875.0">
<quantity id="4CE215A6FFF3FFDBFD0747FEFC86F82F" box="[709,778,1924,1948]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="36" pageNumber="483" unit="m" value="875.0">875 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFF3FFDBFF1147D9FEF7F80F" box="[211,379,1955,1980]" pageId="36" pageNumber="483" value="2010-01-18">
<collectingDate id="EFE0676BFFF3FFDBFF1147D9FEF7F80F" box="[211,379,1955,1980]" pageId="36" pageNumber="483" value="2010-01-18">18 January 2010</collectingDate>
</date>
by
<collectorName id="26EFDD95FFF3FFDBFE6247D9FDF2F80F" box="[416,638,1955,1979]" pageId="36" pageNumber="483">Chifundera Kusamba</collectorName>
,
<collectorName id="26EFDD95FFF3FFDBFD4A47D9FD6BF80F" box="[648,743,1955,1979]" pageId="36" pageNumber="483">Wandege</collectorName>
M.
<collectorName id="26EFDD95FFF3FFDBFCFB40EAFC1BFF1C" box="[825,919,144,168]" pageId="36" pageNumber="483">Muninga</collectorName>
,
<collectorName id="26EFDD95FFF3FFDBFC6040EAFB97FF1C" box="[930,1051,144,168]" pageId="36" pageNumber="483">Mwenebatu</collectorName>
M.
<collectingMunicipality id="6BC12239FFF3FFDBFB8640EAFB14FF1C" box="[1092,1176,144,168]" pageId="36" pageNumber="483">Aristote</collectingMunicipality>
, and
<location id="8EC5EE98FFF3FFDBFB0C40EAFAE1FF1C" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFB0C40EAFAE1FF1C" box="[1230,1389,144,168]" country="Democratic Republic of the Congo" municipality="Aristote" name="Eli Greenbaum" pageId="36" pageNumber="483">Eli Greenbaum</location>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFF3FFDBFABA40EAFCFCFEF1" collectingDate="1947-10-04" collectingDateMax="1947-10-08" collectingDateMin="1947-10-04" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="715" location="Haut-Katanga Province" municipality="Upemba National Park" pageId="36" pageNumber="483" specimenCode="RBINS 2721, RBINS 6157" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFF3FFDBFABA40EAFCE1FF73" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2721</specimenCode>
(formerly under
<specimenCode id="DBBC1038FFF3FFDBFBEB40D5FB3BFF73" box="[1065,1207,175,199]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6157</specimenCode>
), adult male, collected at
<collectorName id="26EFDD95FFF3FFDBFCFB40B5FC43FF53" box="[825,975,207,231]" pageId="36" pageNumber="483">Rivière Kande</collectorName>
, affluent de la rive gauche
<collectorName id="26EFDD95FFF3FFDBFB2740B5FAE7FF53" box="[1253,1387,207,231]" pageId="36" pageNumber="483">de la Lupiala</collectorName>
et sousaffluent de la rive droite de la
<collectorName id="26EFDD95FFF3FFDBFBBD4094FB33FEB2" box="[1151,1215,238,262]" pageId="36" pageNumber="483">Lufira</collectorName>
,
<collectingMunicipality id="6BC12239FFF3FFDBFB094094FA33FEB2" box="[1227,1471,238,262]" pageId="36" pageNumber="483">Upemba National Park</collectingMunicipality>
,
<location id="8EC5EE98FFF3FFDBFCFB4174FBA5FE91" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFCFB4174FBA5FE91" box="[825,1065,269,294]" country="Democratic Republic of the Congo" municipality="Upemba National Park" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
,
<collectingCountry id="F30DF8D3FFF3FFDBFBF34177FBE5FE91" box="[1073,1129,269,293]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
,
<elevation id="00375F70FFF3FFDBFBB34177FA9BFE91" box="[1137,1303,269,293]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="7.3" metricValueMin="7.0" pageId="36" pageNumber="483" unit="m" value="715.0" valueMax="730.0" valueMin="700.0">
<quantity id="4CE215A6FFF3FFDBFBB34177FB6BFE91" box="[1137,1255,269,293]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="7.3" metricValueMin="7.0" pageId="36" pageNumber="483" unit="m" value="715.0" valueMax="730.0" valueMin="700.0">700–730 m</quantity>
a.s.l.
</elevation>
, on
<date id="FFA49E83FFF3FFDBFA824177FCFCFEF1" pageId="36" pageNumber="483" value="1947-10-04" valueMax="1947-10-08" valueMin="1947-10-04">
<collectingDate id="EFE0676BFFF3FFDBFA824177FCFCFEF1" pageId="36" pageNumber="483" value="1947-10-04" valueMax="1947-10-08" valueMin="1947-10-04">4–8 October 1947</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFF3FFDBFCBE4157FC41FE17" collectingDate="1947-10-28" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="700" location="Haut-Katanga Province" pageId="36" pageNumber="483" specimenCode="RBINS 2722, RBINS 6160" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFF3FFDBFCBE4157FB8EFEF1" box="[892,1026,301,325]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2722</specimenCode>
(formerly under
<specimenCode id="DBBC1038FFF3FFDBFB754157FACEFEF0" box="[1207,1346,300,325]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6160</specimenCode>
), adult male collected at
<collectorName id="26EFDD95FFF3FFDBFC734136FBDCFED0" box="[945,1104,332,356]" pageId="36" pageNumber="483">Rivière Lukawe</collectorName>
, affluent de la rive droite
<collectorName id="26EFDD95FFF3FFDBFA894136FA33FED0" box="[1355,1471,332,356]" pageId="36" pageNumber="483">de la Lufira</collectorName>
,
<collectorName id="26EFDD95FFF3FFDBFCFB4111FC37FE37" box="[825,955,363,387]" pageId="36" pageNumber="483">Upemba NP</collectorName>
,
<location id="8EC5EE98FFF3FFDBFC044116FB35FE37" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFC044116FB35FE37" box="[966,1209,363,388]" country="Democratic Republic of the Congo" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
,
<collectingCountry id="F30DF8D3FFF3FFDBFB064111FB70FE37" box="[1220,1276,363,387]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
,
<elevation id="00375F70FFF3FFDBFAC54111FAF2FE37" box="[1287,1406,363,387]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="483" unit="m" value="700.0">
<quantity id="4CE215A6FFF3FFDBFAC54111FAC7FE37" box="[1287,1355,363,387]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="483" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFF3FFDBFA6A4111FC41FE17" pageId="36" pageNumber="483" value="1947-10-28">
<collectingDate id="EFE0676BFFF3FFDBFA6A4111FC41FE17" pageId="36" pageNumber="483" value="1947-10-28">28 October 1947</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFF3FFDBFC1A41F1FB06FE16" box="[984,1162,394,419]" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 2723" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFF3FFDBFC1A41F1FBD1FE17" box="[984,1117,395,419]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2723</specimenCode>
and
</materialsCitation>
<materialsCitation id="3B72B21EFFF3FFDBFB5241F1FC52FD94" collectingDate="1948-06-03" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="890" location="Haut-Katanga Province" pageId="36" pageNumber="483" specimenCode="RBINS 6124" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFF3FFDBFB5241F1FA9AFE16" box="[1168,1302,394,419]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6124</specimenCode>
(formerly under
<specimenCode id="DBBC1038FFF3FFDBFCFB41D0FC4DFE76" box="[825,961,426,450]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6124</specimenCode>
), male and female, respectively, collected at
<collectorName id="26EFDD95FFF3FFDBFAB541D0FA33FE76" box="[1399,1471,426,450]" pageId="36" pageNumber="483">Munoi</collectorName>
, bifurcation de la rivière
<collectorName id="26EFDD95FFF3FFDBFBF741B3FB08FE55" box="[1077,1156,457,481]" pageId="36" pageNumber="483">Lupiala</collectorName>
, affluent de la rive droite
<collectorName id="26EFDD95FFF3FFDBFA5241B3FCF5FDB4" pageId="36" pageNumber="483">de la Lufira</collectorName>
,
<collectorName id="26EFDD95FFF3FFDBFC414192FB88FDB4" box="[899,1028,488,512]" pageId="36" pageNumber="483">Upemba NP</collectorName>
,
<location id="8EC5EE98FFF3FFDBFBCD4193FA8CFDB4" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFBCD4193FA8CFDB4" box="[1039,1280,488,513]" country="Democratic Republic of the Congo" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
,
<collectingCountry id="F30DF8D3FFF3FFDBFAC94193FACFFDB5" box="[1291,1347,489,513]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
,
<elevation id="00375F70FFF3FFDBFA8F4193FA4EFDB4" box="[1357,1474,488,513]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="483" unit="m" value="890.0">
<quantity id="4CE215A6FFF3FFDBFA8F4193FA1CFDB4" box="[1357,1424,489,513]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="483" unit="m" value="890.0">890 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFF3FFDBFC9E4272FC52FD94" box="[860,990,520,544]" pageId="36" pageNumber="483" value="1948-06-03">
<collectingDate id="EFE0676BFFF3FFDBFC9E4272FC52FD94" box="[860,990,520,544]" pageId="36" pageNumber="483" value="1948-06-03">3 June 1948</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFF3FFDBFC284272FABEFD29" collectingDate="1947-05-28" collectingDateMax="1947-05-31" collectingDateMin="1947-05-28" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="890" location="Haut-Katanga Province" pageId="36" pageNumber="483" specimenCode="RBINS 2725-27, RBINS 6122" specimenCount="12" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCode id="DBBC1038FFF3FFDBFC284272FB1BFD94" box="[1002,1175,520,544]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2725–27</specimenCode>
(formerly in a series of
<specimenCount id="9D1C73CAFFF3FFDBFA574272FC28FD8B" count="9" pageId="36" pageNumber="483" type="generic">nine specimens</specimenCount>
under
<specimenCode id="DBBC1038FFF3FFDBFC2D425DFBF5FD8B" box="[1007,1145,551,575]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6122</specimenCode>
),
<specimenCount id="9D1C73CAFFF3FFDBFB4E4252FB79FD8B" box="[1164,1269,551,575]" count="2" pageId="36" pageNumber="483" type="male">two males</specimenCount>
and
<specimenCount id="9D1C73CAFFF3FFDBFAEB4252FA17FD8B" box="[1321,1435,551,575]" count="1" pageId="36" pageNumber="483" type="female">one female</specimenCount>
, respectively, collected at
<collectorName id="26EFDD95FFF3FFDBFBE7423CFBE1FDEA" box="[1061,1133,582,606]" pageId="36" pageNumber="483">Munoi</collectorName>
, bifurcation de la rivière
<collectorName id="26EFDD95FFF3FFDBFAB2423CFA33FDEA" box="[1392,1471,582,606]" pageId="36" pageNumber="483">Lupiala</collectorName>
, affluent de la rive droite
<collectorName id="26EFDD95FFF3FFDBFBEC421CFB2FFDCA" box="[1070,1187,614,638]" pageId="36" pageNumber="483">de la Lufira</collectorName>
,
<collectorName id="26EFDD95FFF3FFDBFB69421CFAA6FDC9" box="[1195,1322,614,638]" pageId="36" pageNumber="483">Upemba NP</collectorName>
,
<location id="8EC5EE98FFF3FFDBFAF1421CFC19FD29" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFAF1421CFC19FD29" country="Democratic Republic of the Congo" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
,
<collectingCountry id="F30DF8D3FFF3FFDBFC6242FFFC54FD29" box="[928,984,645,669]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
,
<elevation id="00375F70FFF3FFDBFC2042FFFBDBFD29" box="[994,1111,645,669]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="483" unit="m" value="890.0">
<quantity id="4CE215A6FFF3FFDBFC2042FFFBA9FD29" box="[994,1061,645,669]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="36" pageNumber="483" unit="m" value="890.0">890 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFF3FFDBFBBD42FFFABEFD29" box="[1151,1330,645,670]" pageId="36" pageNumber="483" value="1947-05-28" valueMax="1947-05-31" valueMin="1947-05-28">
<collectingDate id="EFE0676BFFF3FFDBFBBD42FFFABEFD29" box="[1151,1330,645,670]" pageId="36" pageNumber="483" value="1947-05-28" valueMax="1947-05-31" valueMin="1947-05-28">28–31 May 1947</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFF3FFDBFAFF42FFFBBFFCAF" collectingDate="1949-01-31" collectionCode="RBINS" country="Democratic Republic of the Congo" elevation="700" location="Haut-Katanga Province" municipality="Upemba" pageId="36" pageNumber="483" specimenCode="RBINS 2728, RBINS 6156" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBC1038FFF3FFDBFAFF42FFFA4EFD29" box="[1341,1474,645,669]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 2728</specimenCode>
(formerly under
<specimenCode id="DBBC1038FFF3FFDBFC3442DFFB04FD08" box="[1014,1160,676,701]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6156</specimenCode>
), collected at
<collectorName id="26EFDD95FFF3FFDBFAED42DEFA32FD08" box="[1327,1470,676,700]" pageId="36" pageNumber="483">Kaswabilenga</collectorName>
, région du cours inférieur
<collectorName id="26EFDD95FFF3FFDBFBFC42BEFB48FD68" box="[1086,1220,708,732]" pageId="36" pageNumber="483">de la Lupiala</collectorName>
, affluent de la rive droite
<collectorName id="26EFDD95FFF3FFDBFCFB4299FC3CFD4F" box="[825,944,739,763]" pageId="36" pageNumber="483">de la Lufira</collectorName>
,
<collectingMunicipality id="6BC12239FFF3FFDBFC7B4299FB9DFD4F" box="[953,1041,739,763]" pageId="36" pageNumber="483">Upemba</collectingMunicipality>
NP,
<location id="8EC5EE98FFF3FFDBFB81429EFAB9FD4F" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFB81429EFAB9FD4F" box="[1091,1333,739,764]" country="Democratic Republic of the Congo" municipality="Upemba" name="Haut-Katanga Province" pageId="36" pageNumber="483">Haut-Katanga Province</location>
,
<collectingCountry id="F30DF8D3FFF3FFDBFAFD4299FAFBFD4F" box="[1343,1399,739,763]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
,
<elevation id="00375F70FFF3FFDBFA424299FCE9FCAE" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="483" unit="m" value="700.0">
<quantity id="4CE215A6FFF3FFDBFA424299FA4FFD4F" box="[1408,1475,739,763]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="483" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFF3FFDBFC484379FBBFFCAF" box="[906,1075,771,795]" pageId="36" pageNumber="483" value="1949-01-31">
<collectingDate id="EFE0676BFFF3FFDBFC484379FBBFFCAF" box="[906,1075,771,795]" pageId="36" pageNumber="483" value="1949-01-31">31 January 1949</collectingDate>
</date>
</materialsCitation>
. All
<typeStatus id="54A106E1FFF3FFDBFB9D4379FB34FCAF" box="[1119,1208,771,795]" pageId="36" pageNumber="483" type="paratype">paratype</typeStatus>
material from
<materialsCitation id="3B72B21EFFF3FFDBFA8E4379FB5CFCED" collectionCode="RBINS" collectorName="G. - F. de Witte" country="Democratic Republic of the Congo" location="Katanga region" pageId="36" pageNumber="483" specimenCount="1" stateProvince="Katanga" typeStatus="paratype">
<collectionCode id="ED0B2086FFF3FFDBFA8E4379FA1BFCAF" box="[1356,1431,771,795]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS</collectionCode>
was collected by
<collectorName id="26EFDD95FFF3FFDBFC784358FBC6FC8E" box="[954,1098,802,826]" pageId="36" pageNumber="483">G.-F. de Witte</collectorName>
during his expeditions in the former
<collectingRegion id="49DE76A1FFF3FFDBFCFB4338FC58FCED" box="[825,980,833,858]" country="Democratic Republic of the Congo" name="Katanga" pageId="36" pageNumber="483">Katanga region</collectingRegion>
between 1946 and 1949
</materialsCitation>
.
</paragraph>
<paragraph id="8BA5B843FFF3FFDBFCFB4300FA03FB39" blockId="36.[825,1475,890,1165]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFCFB4300FC73FC26" box="[825,1023,890,914]" italics="true" pageId="36" pageNumber="483">Additional material:</emphasis>
<materialsCitation id="3B72B21EFFF3FFDBFBCB4301FC1AFC65" collectingDate="1932-09" collectionCode="RMCA, R" collectorName="G. F. Overlaet" country="Democratic Republic of the Congo" location="Kapanga" municipality="Haut-Katanga Province" pageId="36" pageNumber="483" specimenCount="1">
<collectionCode id="ED0B2086FFF3FFDBFBCB4301FBD9FC27" box="[1033,1109,891,915]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="36" pageNumber="483" type="Museum">RMCA</collectionCode>
<collectionCode id="ED0B2086FFF3FFDBFB9B4301FBE0FC27" box="[1113,1132,891,915]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="36" pageNumber="483">R</collectionCode>
.8933, male, collected at
<location id="8EC5EE98FFF3FFDBFAA14301FA33FC27" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFAA14301FA33FC27" box="[1379,1471,891,915]" country="Democratic Republic of the Congo" municipality="Haut-Katanga Province" name="Kapanga" pageId="36" pageNumber="483">Kapanga</location>
,
<collectingMunicipality id="6BC12239FFF3FFDBFCFB43E0FBBFFC06" box="[825,1075,922,946]" pageId="36" pageNumber="483">Haut-Katanga Province</collectingMunicipality>
,
<collectingCountry id="F30DF8D3FFF3FFDBFB8743E0FBF1FC06" box="[1093,1149,922,946]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
, in
<date id="FFA49E83FFF3FFDBFB7343E0FAEEFC06" box="[1201,1378,922,946]" pageId="36" pageNumber="483" value="1932-09">
<collectingDate id="EFE0676BFFF3FFDBFB7343E0FAEEFC06" box="[1201,1378,922,946]" pageId="36" pageNumber="483" value="1932-09">September 1932</collectingDate>
</date>
by
<collectorName id="26EFDD95FFF3FFDBFA5543E0FC1AFC65" pageId="36" pageNumber="483">G.F. Overlaet</collectorName>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFF3FFDBFC6343C3FBDFFC65" box="[929,1107,953,977]" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 6121" specimenCount="1">
<specimenCode id="DBBC1038FFF3FFDBFC6343C3FBAAFC65" box="[929,1062,953,977]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6121</specimenCode>
and
</materialsCitation>
<materialsCitation id="3B72B21EFFF3FFDBFB9843C3FC35FC44" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 20749, RBINS 6121" specimenCount="1">
<specimenCode id="DBBC1038FFF3FFDBFB9843C3FB60FC65" box="[1114,1260,953,977]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 20749</specimenCode>
(formerly in
<specimenCode id="DBBC1038FFF3FFDBFAB543C3FCE1FC44" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6121</specimenCode>
series)
</materialsCitation>
,
<materialsCitation id="3B72B21EFFF3FFDBFC0143A2FA32FB9B" collectingDate="1945-07-25" collectingDateMax="1945-07-27" collectingDateMin="1945-07-25" collectorName="de la Grande Kafwe &amp; G. - F. de Witte" country="Democratic Republic of the Congo" county="Haut-Katanga Province" location="Kambi" municipality="Upemba" pageId="36" pageNumber="483" specimenCount="1">
male and female, respectively, collected at
<location id="8EC5EE98FFF3FFDBFAB543A2FA33FC44" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFAB543A2FA33FC44" box="[1399,1471,984,1008]" country="Democratic Republic of the Congo" county="Haut-Katanga Province" municipality="Upemba" name="Kambi" pageId="36" pageNumber="483">Kambi</location>
, affluent
<collectorName id="26EFDD95FFF3FFDBFC4C4382FBD0FBA4" box="[910,1116,1016,1040]" pageId="36" pageNumber="483">de la Grande Kafwe</collectorName>
(
<location id="8EC5EE98FFF3FFDBFBA84382FB66FBA4" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFBA84382FB66FBA4" box="[1130,1258,1016,1040]" country="Democratic Republic of the Congo" county="Haut-Katanga Province" municipality="Upemba" name="Masombwe" pageId="36" pageNumber="483">Masombwe</location>
),
<collectingMunicipality id="6BC12239FFF3FFDBFB394382FADFFBA4" box="[1275,1363,1016,1040]" pageId="36" pageNumber="483">Upemba</collectingMunicipality>
NP,
<collectingCounty id="62C4C0CFFFF3FFDBFA444382FC63FB9B" pageId="36" pageNumber="483">Haut-Katanga Province</collectingCounty>
,
<collectingCountry id="F30DF8D3FFF3FFDBFC3B446DFBBDFB9B" box="[1017,1073,1047,1071]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
, on
<date id="FFA49E83FFF3FFDBFB9E446DFA8AFB9B" box="[1116,1286,1047,1071]" pageId="36" pageNumber="483" value="1945-07-25" valueMax="1945-07-27" valueMin="1945-07-25">
<collectingDate id="EFE0676BFFF3FFDBFB9E446DFA8AFB9B" box="[1116,1286,1047,1071]" pageId="36" pageNumber="483" value="1945-07-25" valueMax="1945-07-27" valueMin="1945-07-25">25–27 July 1945</collectingDate>
</date>
by
<collectorName id="26EFDD95FFF3FFDBFAE9446DFA32FB9B" box="[1323,1470,1047,1071]" pageId="36" pageNumber="483">G.-F. de Witte</collectorName>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFF3FFDBFCFB444CFB82FBFA" box="[825,1038,1078,1102]" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 6139" specimenCount="1">
<specimenCode id="DBBC1038FFF3FFDBFCFB444CFC4CFBFA" box="[825,960,1078,1102]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6139</specimenCode>
(male)
</materialsCitation>
and
<materialsCitation id="3B72B21EFFF3FFDBFB84444CFB82FBDA" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="36" pageNumber="483" specimenCode="RBINS 20748, RBINS 6139" specimenCount="1">
<specimenCode id="DBBC1038FFF3FFDBFB84444CFB56FBFA" box="[1094,1242,1078,1102]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 20748</specimenCode>
(female) (formerly in
<specimenCode id="DBBC1038FFF3FFDBFCFB442CFC4DFBDA" box="[825,961,1110,1134]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="36" pageNumber="483" type="Museum">RBINS 6139</specimenCode>
series)
</materialsCitation>
,
<materialsCitation id="3B72B21EFFF3FFDBFBDE442CFA07FB39" collectingDate="1945-08" collectorName="Upemba NP &amp; G. - F. de Witte" country="Democratic Republic of the Congo" location="Mabwe" municipality="Haut-Lomami Province" pageId="36" pageNumber="483" specimenCount="1">
collected at
<location id="8EC5EE98FFF3FFDBFB5F442CFB66FBDA" LSID="urn:lsid:plazi:treatment:03B30955FFF3FFD9FF3444C6FC56F8A3:8EC5EE98FFF3FFDBFB5F442CFB66FBDA" box="[1181,1258,1110,1134]" country="Democratic Republic of the Congo" municipality="Haut-Lomami Province" name="Mabwe" pageId="36" pageNumber="483">Mabwe</location>
,
<collectorName id="26EFDD95FFF3FFDBFB34442CFAF6FBD9" box="[1270,1402,1110,1134]" pageId="36" pageNumber="483">Upemba NP</collectorName>
,
<collectingMunicipality id="6BC12239FFF3FFDBFA44442CFC7EFB39" pageId="36" pageNumber="483">Haut-Lomami Province</collectingMunicipality>
,
<collectingCountry id="F30DF8D3FFF3FFDBFC39440FFBBFFB39" box="[1019,1075,1141,1165]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
, in
<date id="FFA49E83FFF3FFDBFB97440FFB5BFB39" box="[1109,1239,1141,1165]" pageId="36" pageNumber="483" value="1945-08">
<collectingDate id="EFE0676BFFF3FFDBFB97440FFB5BFB39" box="[1109,1239,1141,1165]" pageId="36" pageNumber="483" value="1945-08">August 1945</collectingDate>
</date>
by
<collectorName id="26EFDD95FFF3FFDBFB38440FFA07FB39" box="[1274,1419,1141,1165]" pageId="36" pageNumber="483">G.-F. de Witte</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA5B843FFF3FFDBFCFB44D5FBC9FA16" blockId="36.[825,1475,1198,1974]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFCFB44D5FC13FB72" box="[825,927,1199,1222]" italics="true" pageId="36" pageNumber="483">Diagnosis:</emphasis>
A large
<taxonomicName id="4C1AC3C0FFF3FFDBFBC044D4FB0DFB72" box="[1026,1153,1198,1222]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="genus">
<emphasis id="B96E6451FFF3FFDBFBC044D4FB0DFB72" box="[1026,1153,1198,1222]" italics="true" pageId="36" pageNumber="483">Lygodactylus</emphasis>
</taxonomicName>
[
<collectionCode id="ED0B2086FFF3FFDBFB4D44D4FB4DFB72" box="[1167,1217,1198,1222]" pageId="36" pageNumber="483">SVL</collectionCode>
to
<quantity id="4CE215A6FFF3FFDBFB2744D4FACFFB72" box="[1253,1347,1198,1222]" metricMagnitude="-2" metricUnit="m" metricValue="3.9799999999999995" pageId="36" pageNumber="483" unit="mm" value="39.8">39.8 mm</quantity>
(mean 36.1 ±
<quantity id="4CE215A6FFF3FFDBFC8D44B4FC28FB52" box="[847,932,1230,1254]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="36" pageNumber="483" unit="mm" value="2.1">2.1 mm</quantity>
)] with 6–9 supralabials and 6–7 infralabials. As with other
<taxonomicName id="4C1AC3C0FFF3FFDBFCB84497FC75FAB1" box="[890,1017,1261,1285]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="genus">
<emphasis id="B96E6451FFF3FFDBFCB84497FC75FAB1" box="[890,1017,1261,1285]" italics="true" pageId="36" pageNumber="483">Lygodactylus</emphasis>
</taxonomicName>
within the
<taxonomicName id="4C1AC3C0FFF3FFDBFBB94494FB78FAB1" box="[1147,1268,1261,1285]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF3FFDBFBB94494FB78FAB1" box="[1147,1268,1261,1285]" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
</taxonomicName>
group, this species can be easily differentiated from the
<taxonomicName id="4C1AC3C0FFF3FFDBFB774577FAA0FA90" authorityName="Gunther" authorityYear="1893" box="[1205,1324,1292,1316]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFF3FFDBFB774577FAA0FA90" box="[1205,1324,1292,1316]" italics="true" pageId="36" pageNumber="483">L. angularis</emphasis>
</taxonomicName>
group species by the gular ornamentation, dorsal pattern, and lack of terminal scansors on the tail tip, and from other members of the
<taxonomicName id="4C1AC3C0FFF3FFDBFCFB4511FC39FA36" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[825,949,1387,1410]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFF3FFDBFCFB4511FC39FA36" box="[825,949,1387,1410]" italics="true" pageId="36" pageNumber="483">L. picturatus</emphasis>
</taxonomicName>
group based on dorsal coloration and gular pattern (see
<taxonomicName id="4C1AC3C0FFF3FFDBFCAB45F0FC51FA16" box="[873,989,1418,1442]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF3FFDBFCAB45F0FC51FA16" box="[873,989,1418,1442]" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
</taxonomicName>
account).
</paragraph>
<paragraph id="8BA5B843FFF3FFDAFC9745D2FF54FA5C" blockId="36.[825,1475,1198,1974]" lastBlockId="37.[113,763,1331,1512]" lastPageId="37" lastPageNumber="484" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFC9745D2FB13FA75" bold="true" box="[853,1183,1448,1473]" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFC9745D2FBB1FA74" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[853,1085,1448,1472]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFC9745D2FBB1FA74" bold="true" box="[853,1085,1448,1472]" italics="true" pageId="36" pageNumber="483">Lygodactylus gamblei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFB8A45D3FB13FA75" box="[1096,1183,1449,1473]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from other members within the
<taxonomicName id="4C1AC3C0FFF3FFDBFB8145B3FB3AFA54" box="[1091,1206,1480,1504]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF3FFDBFB8145B3FB3AFA54" box="[1091,1206,1480,1504]" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
</taxonomicName>
subgroup by the following combination of characters: dorsal coloration lacking dorsolateral series of cream ocelli on flanks (present in
<taxonomicName id="4C1AC3C0FFF3FFDBFAEA4672FA12F9AB" box="[1320,1438,1543,1567]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF3FFDBFAEA4672FA12F9AB" box="[1320,1438,1543,1567]" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
</taxonomicName>
,
<emphasis id="B96E6451FFF3FFDBFA6E467DFC71F98A" bold="true" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFA6E467DFC13F98A" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFA6E467DFC13F98A" bold="true" italics="true" pageId="36" pageNumber="483">L. karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFC6A465CFC71F98A" box="[936,1021,1574,1598]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B96E6451FFF3FFDBFBC9465DFB46F98A" bold="true" box="[1035,1226,1574,1598]" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFBC9465DFBE1F98A" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1035,1133,1574,1598]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFBC9465DFBE1F98A" bold="true" box="[1035,1133,1574,1598]" italics="true" pageId="36" pageNumber="483">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFBB7465CFB46F98A" box="[1141,1226,1574,1598]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C1AC3C0FFF3FFDBFAC5465DFAF7F98A" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[1287,1403,1574,1598]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFF3FFDBFAC5465DFAF7F98A" box="[1287,1403,1574,1598]" italics="true" pageId="36" pageNumber="483">L. dysmicus</emphasis>
</taxonomicName>
), with two parallel dorsolateral lines from head to forelimbs, the lower line extending on to the anterior side of the forelimb, reaching the cubital fossa (
<figureCitation id="1321A4C6FFF3FFDBFC3046FEFBC0F928" box="[1010,1100,1668,1692]" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." pageId="36" pageNumber="483">Fig. 25B</figureCitation>
); males usually with three
<emphasis id="B96E6451FFF3FFDBFAA046FEFAFFF928" bold="true" box="[1378,1395,1668,1692]" pageId="36" pageNumber="483">∩</emphasis>
<emphasis id="B96E6451FFF3FFDBFAB146FFFAF6F928" box="[1395,1402,1669,1692]" italics="true" pageId="36" pageNumber="483">-</emphasis>
shaped chevrons (vs. one or two in
<taxonomicName id="4C1AC3C0FFF3FFDBFB9546DEFB47F90F" authorityName=", Schmidt" authorityYear="1919" box="[1111,1227,1699,1723]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFF3FFDBFB9546DEFB47F90F" box="[1111,1227,1699,1723]" italics="true" pageId="36" pageNumber="483">L. depressus</emphasis>
</taxonomicName>
; a broken chevron in
<emphasis id="B96E6451FFF3FFDBFA6E46DEFB86F96F" bold="true" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFA6E46DEFC39F96E" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFA6E46DEFC39F96E" bold="true" italics="true" pageId="36" pageNumber="483">L. leopardinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFC7B46B9FB86F96F" box="[953,1034,1731,1755]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; and the exceptional inverted
<emphasis id="B96E6451FFF3FFDBFAFA46BEFACBF96F" box="[1336,1351,1732,1755]" italics="true" pageId="36" pageNumber="483">Y</emphasis>
-shaped pattern in
<emphasis id="B96E6451FFF3FFDBFC474699FBF7F94E" bold="true" box="[901,1147,1762,1786]" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFC474699FBADF94E" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[901,1057,1762,1786]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFC474699FBADF94E" bold="true" box="[901,1057,1762,1786]" italics="true" pageId="36" pageNumber="483">L. mirabundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFBEA4698FBF7F94E" box="[1064,1147,1762,1786]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and thicker gular lines than
<taxonomicName id="4C1AC3C0FFF3FFDBFA6C4699FC18F8AD" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF3FFDBFA6C4699FC18F8AD" italics="true" pageId="36" pageNumber="483">L. gutturalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1AC3C0FFF3FFDBFC634778FB98F8AD" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[929,1044,1793,1817]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFF3FFDBFC634778FB98F8AD" box="[929,1044,1793,1817]" italics="true" pageId="36" pageNumber="483">L. dysmicus</emphasis>
</taxonomicName>
, and
<emphasis id="B96E6451FFF3FFDBFB8F4778FC65F88D" bold="true" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFB8F4778FB5CF8AD" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1101,1232,1793,1817]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="karamoja" status="sp. nov">
<emphasis id="B96E6451FFF3FFDBFB8F4778FB5CF8AD" bold="true" box="[1101,1232,1793,1817]" italics="true" pageId="36" pageNumber="483">L. karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFB154778FAA9F8AE" box="[1239,1317,1794,1818]" pageId="36" pageNumber="483" rank="species">sp. nov</taxonomicNameLabel>
..
<taxonomicName id="4C1AC3C0FFF3FFDBFAF5477BFC00F88C" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFAF5477BFC00F88C" bold="true" italics="true" pageId="36" pageNumber="483">Lygodactylus gamblei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFC56475BFC65F88D" box="[916,1001,1825,1849]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
always has three postmental scales, with the lateral pair separated by a posterior extension of the mental (
<figureCitation id="1321A4C6FFF3FFDBFC864725FC2DF8CC" box="[836,929,1887,1912]" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." pageId="36" pageNumber="483">Fig. 25A</figureCitation>
). It can be differentiated from
<emphasis id="B96E6451FFF3FFDBFB1D471AFA31F8CC" bold="true" box="[1247,1469,1887,1912]" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFB1D471AFAEEF8C3" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1247,1378,1887,1911]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFB1D471AFAEEF8C3" bold="true" box="[1247,1378,1887,1911]" italics="true" pageId="36" pageNumber="483">L. karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFAAB471AFA31F8CC" box="[1385,1469,1888,1912]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B96E6451FFF3FFDBFCFB4705FC7CF823" bold="true" box="[825,1008,1918,1943]" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFCFB4705FC14F822" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[825,920,1918,1942]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFCFB4705FC14F822" bold="true" box="[825,920,1918,1942]" italics="true" pageId="36" pageNumber="483">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFC5C4705FC7CF823" box="[926,1008,1919,1943]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<emphasis id="B96E6451FFF3FFDBFBE54705FA9BF823" bold="true" box="[1063,1303,1918,1943]" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFBE54705FB33F822" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1063,1215,1918,1942]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFBE54705FB33F822" bold="true" box="[1063,1215,1918,1942]" italics="true" pageId="36" pageNumber="483">L. leopardinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFB074705FA9BF823" box="[1221,1303,1919,1943]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a proportionally larger tibia (CL/
<collectionCode id="ED0B2086FFF3FFDBFBA147E4FB1AF802" box="[1123,1174,1950,1974]" pageId="36" pageNumber="483">SVL</collectionCode>
≥ 0.19 vs. CL/
<collectionCode id="ED0B2086FFF3FFDBFAF547E4FAE4F802" box="[1335,1384,1950,1974]" pageId="36" pageNumber="483">SVL</collectionCode>
≤ 0.17). As in its sister taxon
<emphasis id="B96E6451FFF2FFDAFE8A454EFDB4FAF8" bold="true" box="[328,568,1331,1356]" pageId="37" pageNumber="484">
<taxonomicName id="4C1AC3C0FFF2FFDAFE8A454EFE6CFAFF" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[328,480,1331,1355]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
<emphasis id="B96E6451FFF2FFDAFE8A454EFE6CFAFF" bold="true" box="[328,480,1331,1355]" italics="true" pageId="37" pageNumber="484">L. leopardinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF2FFDAFE24454EFDB4FAF8" box="[486,568,1332,1356]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, females lack gular patterning (present in all the other members of the group,
<figureCitation id="1321A4C6FFF2FFDAFD164529FF2DFA3E" captionStart="Figure 25" captionStartId="40.[129,194,1561,1585]" captionTargetBox="[137,1469,148,1531]" captionTargetId="figure-207@40.[128,1475,144,1533]" captionTargetPageId="40" captionText="Figure 25. A, Dorsal and ventral views of head, showing gular ornamentation and pholidosis variation in L. gamblei sp. nov. B, Specimens of L. gamblei sp. nov. in lateral view, showing its characteristic scapular pattern. Red points denote males and blue points denote females. Photos by J.L.R." pageId="37" pageNumber="484">Fig. 25A</figureCitation>
).
<emphasis id="B96E6451FFF2FFDAFF704508FE61FA3E" bold="true" box="[178,493,1394,1418]" pageId="37" pageNumber="484">
<taxonomicName id="4C1AC3C0FFF2FFDAFF704508FE1AFA3E" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[178,406,1394,1418]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
<emphasis id="B96E6451FFF2FFDAFF704508FE1AFA3E" bold="true" box="[178,406,1394,1418]" italics="true" pageId="37" pageNumber="484">Lygodactylus gamblei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF2FFDAFE594508FE61FA3E" box="[411,493,1394,1418]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from
<emphasis id="B96E6451FFF2FFDAFFB345E8FEE8FA1E" bold="true" box="[113,356,1425,1450]" pageId="37" pageNumber="484">
<taxonomicName id="4C1AC3C0FFF2FFDAFFB345E8FE80FA1D" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[113,268,1425,1449]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF2FFDAFFB345E8FE80FA1D" bold="true" box="[113,268,1425,1449]" italics="true" pageId="37" pageNumber="484">L. mirabundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF2FFDAFED045E8FEE8FA1E" box="[274,356,1426,1450]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a lower number of precloacal pores (7–9 vs. 10) and by subtle morphometric differences (
<tableCitation id="C6988DF8FFF2FFDAFFBE45AAFF4BFA5C" box="[124,199,1488,1512]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="37" pageNumber="484">Table 3</tableCitation>
).
</paragraph>
<caption id="DF65E8CBFFF2FFDAFFB344CEFADFFAB0" pageId="37" pageNumber="484" startId="37.[113,178,1204,1228]" targetBox="[114,1458,144,1176]" targetPageId="37" targetType="figure">
<paragraph id="8BA5B843FFF2FFDAFFB344CEFADFFAB0" blockId="37.[113,1458,1203,1284]" pageId="37" pageNumber="484">
<emphasis id="B96E6451FFF2FFDAFFB344CEFF5AFB78" bold="true" box="[113,214,1204,1228]" pageId="37" pageNumber="484">Figure 23.</emphasis>
A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of
<emphasis id="B96E6451FFF2FFDAFCBD44CEFBD6FB78" bold="true" box="[895,1114,1203,1228]" pageId="37" pageNumber="484">
<taxonomicName id="4C1AC3C0FFF2FFDAFCBD44CEFB87FB7F" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[895,1035,1203,1227]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
<emphasis id="B96E6451FFF2FFDAFCBD44CEFB87FB7F" bold="true" box="[895,1035,1203,1227]" italics="true" pageId="37" pageNumber="484">L. leopardinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF2FFDAFBCD44CEFBD6FB78" box="[1039,1114,1204,1228]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of
<emphasis id="B96E6451FFF2FFDAFF4B4496FEB3FAB0" bold="true" box="[137,319,1259,1284]" pageId="37" pageNumber="484">
<taxonomicName id="4C1AC3C0FFF2FFDAFF4B4496FF7CFAB7" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[137,240,1259,1283]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="484" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
<emphasis id="B96E6451FFF2FFDAFF4B4496FF7CFAB7" bold="true" box="[137,240,1259,1283]" italics="true" pageId="37" pageNumber="484">L. gamblei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF2FFDAFF364496FEB3FAB0" box="[244,319,1260,1284]" pageId="37" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G.
</paragraph>
</caption>
<paragraph id="8BA5B843FFF2FFDAFFB34672FEB4F928" blockId="37.[113,762,1544,1693]" pageId="37" pageNumber="484">
<emphasis id="B96E6451FFF2FFDAFFB34672FF6DF994" box="[113,225,1544,1568]" italics="true" pageId="37" pageNumber="484">Etymology:</emphasis>
We name this new species after the American evolutionary biologist and herpetologist Tony Gamble of Marquette University, in recognition of his substantial contributions to the evolutionary biology of geckos. The name is a patronym formed in the genitive case.
</paragraph>
<paragraph id="8BA5B843FFF2FFD9FFB346D7FC86FAD8" blockId="37.[113,762,1708,1952]" lastBlockId="38.[129,778,1333,1388]" lastPageId="38" lastPageNumber="485" pageId="37" pageNumber="484">
<emphasis id="B96E6451FFF2FFDAFFB346D7FE1EF970" box="[113,402,1708,1732]" italics="true" pageId="37" pageNumber="484">
Description of the
<typeStatus id="54A106E1FFF2FFDAFE8246D6FE1EF970" box="[320,402,1708,1732]" pageId="37" pageNumber="484" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="1321A4C6FFF2FFDAFE7F46D6FD81F970" box="[445,525,1708,1732]" captionStart="Figure 24" captionStartId="38.[129,194,1232,1256]" captionTargetBox="[129,1473,144,1205]" captionTargetId="figure-368@38.[129,1473,144,1205]" captionTargetPageId="38" captionText="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="37" pageNumber="484">Fig. 24</figureCitation>
): Measurements and meristic characters of the
<typeStatus id="54A106E1FFF2FFDAFEB846B1FE59F957" box="[378,469,1739,1763]" pageId="37" pageNumber="484" type="holotype">holotype</typeStatus>
are presented in Supporting Information, Table S10. Adult male, with a snout–vent length (
<collectionCode id="ED0B2086FFF2FFDAFFBB4770FF20F896" box="[121,172,1802,1826]" pageId="37" pageNumber="484">SVL</collectionCode>
) of 35.0 mm and a regenerated tail of 31.0 mm.Body slender and nape slightly distinct. Head slightly broader than body, and moderate head length (
<collectionCode id="ED0B2086FFF2FFDAFEA94733FE17F8D5" box="[363,411,1865,1889]" pageId="37" pageNumber="484">HW</collectionCode>
/HL 0.66). Canthus rostralis not prominent. Eye diameter
<quantity id="4CE215A6FFF2FFDAFEB04712FE40F834" box="[370,460,1896,1920]" metricMagnitude="-3" metricUnit="m" metricValue="2.51" pageId="37" pageNumber="484" unit="mm" value="2.51">2.51 mm</quantity>
with circular pupil. Ear to eye distance slightly larger than the orbit diameter (
<quantity id="4CE215A6FFF2FFDAFD8E47FDFD22F814" box="[588,686,1927,1952]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" pageId="37" pageNumber="484" unit="mm" value="3.26">3.26 mm</quantity>
). Snout rounded and moderately pointed. Granular scale of frontal larger than occipital scales, with 25 small interorbital granular scales. Dorsal scales granular from rostral to tail. Rostral undivided, in narrow contact with nostril. Two small symmetrical internasals. Seven supralabials and six infralabials. Prenasal contacts the 1st supralabial. Nostril circular and surrounded by 1st supralabial, prenasal, one supranasal, and one postnasal. Four or five rows of scales between supralabials and the orbit. Mental large, triangular, and posteriorly in contact with three rounded symmetrical postmental scales. Mental scales extending between the lateral postmentals reaching ~50% of the medial surface suture of the postmentals. Five smaller post-postmental scales. Gular scales granular, rounded, and smaller than ventral scales. Ventrals imbricate, denticulate, and larger than dorsal scales, with 19 scale rows across the venter. Regenerated tail with 30 enlarged subcaudal scales, some fragmented. Body relatively robust and slightly elongated (
<collectionCode id="ED0B2086FFF2FFDAFC274753FB95F8F5" box="[997,1049,1833,1857]" pageId="37" pageNumber="484">TRL</collectionCode>
/
<collectionCode id="ED0B2086FFF2FFDAFBE34753FBDFF8F5" box="[1057,1107,1833,1857]" pageId="37" pageNumber="484">SVL</collectionCode>
0.42). Eight precloacal pores. Fore- and hind limbs moderately short and stout; forearm of medium length (
<collectionCode id="ED0B2086FFF2FFDAFCB8471DFC16F8CB" box="[890,922,1895,1919]" pageId="37" pageNumber="484">FL</collectionCode>
/
<collectionCode id="ED0B2086FFF2FFDAFC60471DFC58F8CB" box="[930,980,1895,1919]" pageId="37" pageNumber="484">SVL</collectionCode>
0.16); tibia moderately long (CL/
<collectionCode id="ED0B2086FFF2FFDAFAF9471DFAE1F8CB" box="[1339,1389,1895,1919]" pageId="37" pageNumber="484">SVL</collectionCode>
0.19). Digits elongated and unwebbed with 5–6 terminal scansors. Thumb rudimentary with a small terminal claw.Relative length of digits: I &lt;II =
<collectionCode id="ED0B2086FFF1FFD9FED4452EFEABFADF" box="[278,295,1364,1387]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="38" pageNumber="485" type="Museum">V</collectionCode>
&lt;III &lt;IV (manus); I &lt;II &lt;
<collectionCode id="ED0B2086FFF1FFD9FD89452EFDD0FADF" box="[587,604,1364,1387]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="38" pageNumber="485" type="Museum">V</collectionCode>
&lt;III &lt;IV (pes).
</paragraph>
<caption id="DF65E8CBFFF1FFD9FF4344AAFABCFAB0" pageId="38" pageNumber="485" startId="38.[129,194,1232,1256]" targetBox="[129,1473,144,1205]" targetPageId="38" targetType="figure">
<paragraph id="8BA5B843FFF1FFD9FF4344AAFABCFAB0" blockId="38.[129,1456,1232,1285]" pageId="38" pageNumber="485">
<emphasis id="B96E6451FFF1FFD9FF4344AAFF69FB5D" bold="true" box="[129,229,1232,1257]" pageId="38" pageNumber="485">Figure 24.</emphasis>
Holotype of
<emphasis id="B96E6451FFF1FFD9FEA344ABFD9BFB5D" bold="true" box="[353,535,1232,1257]" pageId="38" pageNumber="485">
<taxonomicName id="4C1AC3C0FFF1FFD9FEA344ABFE44FB5C" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[353,456,1232,1256]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="485" phylum="Chordata" rank="species" species="gamblei" status="sp. nov.">
<emphasis id="B96E6451FFF1FFD9FEA344ABFE44FB5C" bold="true" box="[353,456,1232,1256]" italics="true" pageId="38" pageNumber="485">L. gamblei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF1FFD9FE0E44ABFD9BFB5D" box="[460,535,1233,1257]" pageId="38" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UTEP 22587) from Manono, Tanganyika Province, DRC. A, Dorsal and ventral view of body. B, Details of the cloacal region. C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R.
</paragraph>
</caption>
<paragraph id="8BA5B843FFF1FFD9FF4345F7FD3FF96A" blockId="38.[129,778,1421,1758]" pageId="38" pageNumber="485">
<emphasis id="B96E6451FFF1FFD9FF4345F7FEA2FA11" box="[129,302,1421,1445]" italics="true" pageId="38" pageNumber="485">Coloration: In life</emphasis>
(
<figureCitation id="1321A4C6FFF1FFD9FEFF45F7FE52FA11" box="[317,478,1421,1445]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="38" pageNumber="485">Fig. 23A–D, F–I</figureCitation>
), dorsal colour light brown to grey, with dark brown mottled pattern in the female; male with three rows of interrupted, but conspicuous black stripes on lateral surface from head to neck, converging at the anterior insertion of the forearm, giving the impression of a single large blotch; male and female with a black-brown line from snout to ear aperture; venter uniform orange-yellowish from posterior part of gular to cloaca, and cream to white elsewhere; gular immaculate white in female, and white in male with three thick and black
<emphasis id="B96E6451FFF1FFD9FD6846F2FD37F914" bold="true" box="[682,699,1672,1696]" pageId="38" pageNumber="485">∩</emphasis>
<emphasis id="B96E6451FFF1FFD9FD7946F3FD4EF914" box="[699,706,1673,1696]" italics="true" pageId="38" pageNumber="485">-</emphasis>
shaped chevrons.
<emphasis id="B96E6451FFF1FFD9FF2446D2FEFCF90B" box="[230,368,1704,1727]" italics="true" pageId="38" pageNumber="485">In preservative</emphasis>
(
<typeStatus id="54A106E1FFF1FFD9FEB946DDFE52F90B" box="[379,478,1703,1727]" pageId="38" pageNumber="485" type="holotype">holotype</typeStatus>
;
<figureCitation id="1321A4C6FFF1FFD9FE2446DDFDA6F90B" box="[486,554,1703,1727]" captionStart="Figure 24" captionStartId="38.[129,194,1232,1256]" captionTargetBox="[129,1473,144,1205]" captionTargetId="figure-368@38.[129,1473,144,1205]" captionTargetPageId="38" captionText="Figure 24. Holotype of L. gamblei sp. nov. (UTEP 22587) from Manono, Tanganyika Province, DRC.A, Dorsal and ventral view of body.B, Details of the cloacal region.C, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="38" pageNumber="485">Fig. 24</figureCitation>
): similar coloration as in life, slightly darker, with gular pattern more brownish.
</paragraph>
<paragraph id="8BA5B843FFF1FFD9FF43477BFC3CFA3F" blockId="38.[129,778,1792,1972]" lastBlockId="38.[825,1474,1332,1419]" pageId="38" pageNumber="485">
<emphasis id="B96E6451FFF1FFD9FF43477BFF68F8AC" box="[129,228,1793,1816]" italics="true" pageId="38" pageNumber="485">Variation:</emphasis>
All
<typeStatus id="54A106E1FFF1FFD9FEDC477AFE0DF8AC" box="[286,385,1792,1816]" pageId="38" pageNumber="485" type="paratype">paratypes</typeStatus>
largely agree with the
<typeStatus id="54A106E1FFF1FFD9FDBA477AFD5AF8AC" box="[632,726,1792,1816]" pageId="38" pageNumber="485" type="holotype">holotype</typeStatus>
, and their measurements are summarized in
<collectorName id="26EFDD95FFF1FFD9FDCC4765FC89F883" box="[526,773,1823,1847]" pageId="38" pageNumber="485">Supporting Information</collectorName>
,
<collectorName id="26EFDD95FFF1FFD9FF434744FF43F8E3" box="[129,207,1854,1879]" pageId="38" pageNumber="485">Table S</collectorName>
10.
<collectorName id="26EFDD95FFF1FFD9FF3A4744FE93F8E2" box="[248,287,1854,1878]" pageId="38" pageNumber="485">The</collectorName>
specimen
<specimenCode id="DBBC1038FFF1FFD9FE514745FD96F8E2" box="[403,538,1854,1879]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="38" pageNumber="485" type="Museum">RBINS 6138</specimenCode>
has a bold dorsal pattern from head to midbody.
<collectorName id="26EFDD95FFF1FFD9FE5C4724FE22F8C1" box="[414,430,1886,1909]" pageId="38" pageNumber="485">It</collectorName>
is worth noting that all the female specimens examined from
<collectorName id="26EFDD95FFF1FFD9FE5A4707FE72F821" box="[408,510,1917,1941]" pageId="38" pageNumber="485">de Witte’s</collectorName>
(1953) expedition lack a gular pattern, in contradiction to
<collectorName id="26EFDD95FFF1FFD9FE1D47E6FDC9F800" box="[479,581,1948,1972]" pageId="38" pageNumber="485">de Witte’s</collectorName>
(1953) statement.
<collectorName id="26EFDD95FFF1FFD9FCFB454FFC1BFAF8" box="[825,919,1333,1356]" pageId="38" pageNumber="485">However</collectorName>
, we noted that some young male specimens were catalogued as females, which could have misled
<collectorName id="26EFDD95FFF1FFD9FAD5452EFAFFFAD8" box="[1303,1395,1364,1388]" pageId="38" pageNumber="485">de Witte</collectorName>
to this conclusion.
</paragraph>
<paragraph id="8BA5B843FFF1FFD9FCFB45D6FC19F9D5" blockId="38.[825,1474,1452,1633]" pageId="38" pageNumber="485">
<emphasis id="B96E6451FFF1FFD9FCFB45D6FBA5FA70" box="[825,1065,1452,1476]" italics="true" pageId="38" pageNumber="485">Habitat and distribution</emphasis>
(
<figureCitation id="1321A4C6FFF1FFD9FBFD45D6FBF0FA70" box="[1087,1148,1452,1476]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="38" pageNumber="485">Figs 6</figureCitation>
,
<figureCitation id="1321A4C6FFF1FFD9FB4A45D7FB2EFA71" box="[1160,1186,1453,1477]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="38" pageNumber="485">23</figureCitation>
):
<emphasis id="B96E6451FFF1FFD9FB7845D6FCEBFA50" bold="true" pageId="38" pageNumber="485">
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<emphasis id="B96E6451FFF1FFD9FB7845D6FA12FA70" bold="true" box="[1210,1438,1452,1476]" italics="true" pageId="38" pageNumber="485">Lygodactylus gamblei</emphasis>
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is known from Manono, Upemba National Park, and other nearby areas in south-eastern
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. The species is associated with Miombo Woodland forest and gallery forest between 500 and ~
<quantity id="4CE215A6FFF1FFD9FCB74650FC4BF9F5" box="[885,967,1578,1602]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="485" unit="m" value="1500.0">1500 m</quantity>
a.s.l. This species is sympatric with
<taxonomicName id="4C1AC3C0FFF1FFD9FA834651FA29F9F6" box="[1345,1445,1578,1602]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="485" phylum="Chordata" rank="species" species="heeneni">
<emphasis id="B96E6451FFF1FFD9FA834651FA29F9F6" box="[1345,1445,1578,1602]" italics="true" pageId="38" pageNumber="485">L. heeneni</emphasis>
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in this area.
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<emphasis id="B96E6451FFF1FFD9FCFB46F8FC55F92E" box="[825,985,1666,1690]" italics="true" pageId="38" pageNumber="485">Natural history:</emphasis>
A diurnal and arboreal species. The
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(UTEP 22587) was collected active during the daytime with an air temperature of 45 °C and one of the
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(UTEP 22584) was found scampering around a tree close to the ground in an open area.
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<emphasis id="B96E6451FFF5FFDDFC534104FA8AFE2C" bold="true" box="[913,1286,382,408]" italics="true" pageId="34" pageNumber="481">LYGODACTYLUS LEOPARDINUS</emphasis>
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(
<figureCitation id="1321A4C6FFF5FFDDFCAA41DFFC54FE09" box="[872,984,421,445]" captionStart-0="Figure 22" captionStart-1="Figure 23" captionStartId-0="35.[113,178,1896,1920]" captionStartId-1="37.[113,178,1204,1228]" captionTargetBox-0="[114,1458,918,1868]" captionTargetBox-1="[114,1458,144,1176]" captionTargetId-0="figure-448@35.[114,1458,918,1868]" captionTargetId-1="figure-366@37.[114,1458,144,1176]" captionTargetPageId-0="35" captionTargetPageId-1="37" captionText-0="Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." captionText-1="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="34" pageNumber="481">Figs 22–23</figureCitation>
,
<tableCitation id="C6988DF8FFF5FFDDFC2141DFFBA2FE09" box="[995,1070,421,445]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="34" pageNumber="481">Table 3</tableCitation>
; Supporting Information, Table S9)
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<emphasis id="B96E6451FFF5FFDDFCFB41A4FB9AFE42" box="[825,1046,478,502]" italics="true" pageId="34" pageNumber="481">Zoobank registration:</emphasis>
https://zoobank.org/
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<emphasis id="B96E6451FFF5FFDDFC974266FBEFFD80" bold="true" box="[853,1123,540,564]" italics="true" pageId="34" pageNumber="481">Lygodactylus leopardinus</emphasis>
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is the rainforest member within subgroup C, and the most morphologically distinguishable species within the
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.
<emphasis id="B96E6451FFF5FFDDFB834221FB10FDC7" box="[1089,1180,603,627]" italics="true" pageId="34" pageNumber="481">gutturalis</emphasis>
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subgroup. It also differs by a minimum of
<emphasis id="B96E6451FFF5FFDDFC7C4201FC4BFD26" box="[958,967,635,658]" italics="true" pageId="34" pageNumber="481">c</emphasis>
. 10.10% for
<emphasis id="B96E6451FFF5FFDDFB844200FBE0FD26" box="[1094,1132,634,658]" italics="true" pageId="34" pageNumber="481">16S</emphasis>
uncorrected p-distance (Table 2) from related taxa, and it lacks nuclear haplotype sharing in
<emphasis id="B96E6451FFF5FFDDFA4742E0FA4EFD06" box="[1413,1474,666,690]" italics="true" pageId="34" pageNumber="481">RAG1</emphasis>
(
<figureCitation id="1321A4C6FFF5FFDDFC8642C3FC20FD65" box="[836,940,697,721]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability&gt; 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="34" pageNumber="481">Fig. 2B–C</figureCitation>
).
</paragraph>
<paragraph id="8BA5B843FFF5FFDDFCFB4289FA88FC3C" blockId="34.[825,1474,754,904]" pageId="34" pageNumber="481">
<materialsCitation id="3B72B21EFFF5FFDDFCFB4289FA88FC3C" collectingDate="2013-07-18" collectionCode="UTEP, ELI" collectorName="Chifundera Kusamba &amp; Wandege M. Muninga &amp; Mwenebatu M. Aristote &amp; Eli Greenbaum." country="Democratic Republic of the Congo" elevation="305" latitude="-0.25939" location="Balolombo Village" longLatPrecision="1" longitude="19.63575" municipality="Busira River" pageId="34" pageNumber="481" specimenCode="UTEP 22577, ELI 2330" specimenCount="1" stateProvince="Equateur" typeStatus="holotype">
<emphasis id="B96E6451FFF5FFDDFCFB4289FC15FCBF" box="[825,921,755,779]" italics="true" pageId="34" pageNumber="481">
<typeStatus id="54A106E1FFF5FFDDFCFB4289FC18FCBF" box="[825,916,755,779]" pageId="34" pageNumber="481" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DBBC1038FFF5FFDDFC644289FBBDFCBE" box="[934,1073,754,778]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="34" pageNumber="481" type="Museum">UTEP 22577</specimenCode>
(
<specimenCode id="DBBC1038FFF5FFDDFBFD4289FB25FCBF" box="[1087,1193,755,779]" collectionCode="ELI" pageId="34" pageNumber="481">ELI 2330</specimenCode>
), male with broken tail and ventral incision, collected at
<location id="8EC5EE98FFF5FFDDFBAE4368FAA3FC9E" LSID="urn:lsid:plazi:treatment:03B30955FFF5FFDBFC534104FEB5FB28:8EC5EE98FFF5FFDDFBAE4368FAA3FC9E" box="[1132,1327,786,810]" country="Democratic Republic of the Congo" latitude="-0.25939" longLatPrecision="1" longitude="19.63575" municipality="Busira River" name="Balolombo Village" pageId="34" pageNumber="481" stateProvince="Equateur">Balolombo Village</location>
,
<collectingMunicipality id="6BC12239FFF5FFDDFAFE4368FA32FC9E" box="[1340,1470,786,810]" pageId="34" pageNumber="481">Busira River</collectingMunicipality>
,
<collectingRegion id="49DE76A1FFF5FFDDFCFB4349FC7BFCFD" box="[825,1015,817,846]" country="Democratic Republic of the Congo" name="Equateur" pageId="34" pageNumber="481">Équateur Province</collectingRegion>
,
<collectingCountry id="F30DF8D3FFF5FFDDFBC2434BFBB4FCFD" box="[1024,1080,817,841]" name="Democratic Republic of the Congo" pageId="34" pageNumber="481">DRC</collectingCountry>
,
<geoCoordinate id="EE2EDE84FFF5FFDDFB82434BFB3CFCFD" box="[1088,1200,817,841]" degrees="00.25939" direction="south" orientation="latitude" pageId="34" pageNumber="481" precision="1" value="-0.25939">S00.25939</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFF5FFDDFB7A434BFAA6FCFD" box="[1208,1322,817,841]" degrees="19.63575" direction="east" orientation="longitude" pageId="34" pageNumber="481" precision="1" value="19.63575">E19.63575</geoCoordinate>
,
<elevation id="00375F70FFF5FFDDFAF0434BFA2FFCFD" box="[1330,1443,817,841]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="34" pageNumber="481" unit="m" value="305.0">
<quantity id="4CE215A6FFF5FFDDFAF0434BFAFEFCFD" box="[1330,1394,817,841]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="34" pageNumber="481" unit="m" value="305.0">305 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFF5FFDDFCFB432BFC33FCDD" box="[825,959,849,873]" pageId="34" pageNumber="481" value="2013-07-18">
<collectingDate id="EFE0676BFFF5FFDDFCFB432BFC33FCDD" box="[825,959,849,873]" pageId="34" pageNumber="481" value="2013-07-18">18 July 2013</collectingDate>
</date>
by
<collectorName id="26EFDD95FFF5FFDDFC2A432BFB44FCDD" box="[1000,1224,849,873]" pageId="34" pageNumber="481">Chifundera Kusamba</collectorName>
,
<collectorName id="26EFDD95FFF5FFDDFB16432BFA32FCDD" box="[1236,1470,849,873]" pageId="34" pageNumber="481">Wandege M. Muninga</collectorName>
,
<collectorName id="26EFDD95FFF5FFDDFCFB430AFBA2FC3C" box="[825,1070,880,904]" pageId="34" pageNumber="481">Mwenebatu M. Aristote</collectorName>
, and
<collectorName id="26EFDD95FFF5FFDDFBA0430AFA88FC3C" box="[1122,1284,880,904]" pageId="34" pageNumber="481">Eli Greenbaum.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BA5B843FFF5FFDDFCFB43D0FA0FFBB4" blockId="34.[825,1474,937,1024]" pageId="34" pageNumber="481">
<materialsCitation id="3B72B21EFFF5FFDDFCFB43D0FB4EFC75" box="[825,1218,937,961]" collectionCode="UTEP" pageId="34" pageNumber="481" specimenCode="UTEP 22596, ELI 3584" specimenCount="1" typeStatus="paratype">
<emphasis id="B96E6451FFF5FFDDFCFB43D0FC14FC75" box="[825,920,938,961]" italics="true" pageId="34" pageNumber="481">
<typeStatus id="54A106E1FFF5FFDDFCFB43D0FC1EFC75" box="[825,914,938,961]" pageId="34" pageNumber="481" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCode id="DBBC1038FFF5FFDDFC6643D0FBBAFC75" box="[932,1078,937,961]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="34" pageNumber="481" type="Museum">UTEP 22596</specimenCode>
(
<specimenCode id="DBBC1038FFF5FFDDFB8B43D3FB35FC75" box="[1097,1209,937,961]" collectionCode="ELI" pageId="34" pageNumber="481">ELI 3584</specimenCode>
)
</materialsCitation>
, female, collected from Katopa,
<materialsCitation id="3B72B21EFFF5FFDDFC5743B3FAF3FBB4" collectingDate="2015-07-02" collectionCode="ICCN" country="Democratic Republic of the Congo" elevation="450" latitude="-2.74769" location="Camp" longLatPrecision="1" longitude="25.10323" pageId="34" pageNumber="481" specimenCount="1" stateProvince="Maniema" typeStatus="paratype">
<collectionCode id="ED0B2086FFF5FFDDFC5743B3FC5AFC55" box="[917,982,969,993]" pageId="34" pageNumber="481">ICCN</collectionCode>
<location id="8EC5EE98FFF5FFDDFC2343B3FBADFC55" LSID="urn:lsid:plazi:treatment:03B30955FFF5FFDBFC534104FEB5FB28:8EC5EE98FFF5FFDDFC2343B3FBADFC55" box="[993,1057,969,993]" country="Democratic Republic of the Congo" latitude="-2.74769" longLatPrecision="1" longitude="25.10323" name="Camp" pageId="34" pageNumber="481" stateProvince="Maniema">Camp</location>
,
<collectingRegion id="49DE76A1FFF5FFDDFBF243B3FB74FC55" box="[1072,1272,969,993]" country="Democratic Republic of the Congo" name="Maniema" pageId="34" pageNumber="481">Maniema Province</collectingRegion>
,
<collectingCountry id="F30DF8D3FFF5FFDDFACA43B3FACCFC55" box="[1288,1344,969,993]" name="Democratic Republic of the Congo" pageId="34" pageNumber="481">DRC</collectingCountry>
,
<geoCoordinate id="EE2EDE84FFF5FFDDFA8D43B2FA33FC54" box="[1359,1471,968,992]" degrees="02.74769" direction="south" orientation="latitude" pageId="34" pageNumber="481" precision="1" value="-2.74769">S02.74769</geoCoordinate>
,
<geoCoordinate id="EE2EDE84FFF5FFDDFCFB4392FC27FBB4" box="[825,939,1000,1024]" degrees="25.10323" direction="east" orientation="longitude" pageId="34" pageNumber="481" precision="1" value="25.10323">E25.10323</geoCoordinate>
,
<elevation id="00375F70FFF5FFDDFC764392FBAAFBB4" box="[948,1062,1000,1024]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="34" pageNumber="481" unit="m" value="450.0">
<quantity id="4CE215A6FFF5FFDDFC764392FC79FC4B" box="[948,1013,1000,1024]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="34" pageNumber="481" unit="m" value="450.0">450 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFF5FFDDFB894392FB32FBB4" box="[1099,1214,1000,1024]" pageId="34" pageNumber="481" value="2015-07-02">
<collectingDate id="EFE0676BFFF5FFDDFB894392FB32FBB4" box="[1099,1214,1000,1024]" pageId="34" pageNumber="481" value="2015-07-02">2 July 2015</collectingDate>
</date>
by Eli Greenbaum
</materialsCitation>
.
</paragraph>
<paragraph id="8BA5B843FFF5FFDDFCFB445BFB46FB41" blockId="34.[825,1474,1057,1269]" pageId="34" pageNumber="481">
<emphasis id="B96E6451FFF5FFDDFCFB445BFB89FB8D" box="[825,1029,1057,1081]" italics="true" pageId="34" pageNumber="481">Additional material:</emphasis>
<materialsCitation id="3B72B21EFFF5FFDDFBD2445BFB47FB8D" box="[1040,1227,1057,1081]" collectionCode="RMCA" pageId="34" pageNumber="481" specimenCode="RMCA 1981.065" specimenCount="1">
<specimenCode id="DBBC1038FFF5FFDDFBD2445BFB47FB8D" box="[1040,1227,1057,1081]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="34" pageNumber="481" type="Museum">RMCA 1981.065</specimenCode>
</materialsCitation>
.
<materialsCitation id="3B72B21EFFF5FFDDFB0C445BFC02FB23" collectingDate="1955-08-17" collectingDateMax="1955-08-18" collectingDateMin="1955-08-17" collectionCode="R" collectorName="Raymond Laurent" country="Democratic Republic of the Congo" location="River Lotende" municipality="Mabali" pageId="34" pageNumber="481" specimenCount="1" stateProvince="Equateur">
<collectionCode id="ED0B2086FFF5FFDDFB0C445BFB6CFB8D" box="[1230,1248,1057,1081]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="34" pageNumber="481">R</collectionCode>
.0004, male with original tail, collected at
<location id="8EC5EE98FFF5FFDDFBD1443BFB29FBED" LSID="urn:lsid:plazi:treatment:03B30955FFF5FFDBFC534104FEB5FB28:8EC5EE98FFF5FFDDFBD1443BFB29FBED" box="[1043,1189,1089,1113]" country="Democratic Republic of the Congo" municipality="Mabali" name="River Lotende" pageId="34" pageNumber="481" stateProvince="Equateur">River Lotende</location>
(canopy),
<collectingMunicipality id="6BC12239FFF5FFDDFAD1443BFAD0FBED" box="[1299,1372,1089,1113]" pageId="34" pageNumber="481">Mabali</collectingMunicipality>
,
<collectingRegion id="49DE76A1FFF5FFDDFAA64438FC19FBCC" country="Democratic Republic of the Congo" name="Equateur" pageId="34" pageNumber="481">Équateur Province</collectingRegion>
,
<collectingCountry id="F30DF8D3FFF5FFDDFC63441AFC55FBCC" box="[929,985,1120,1144]" name="Democratic Republic of the Congo" pageId="34" pageNumber="481">DRC</collectingCountry>
, between
<date id="FFA49E83FFF5FFDDFB83441AFAB9FBCC" box="[1089,1333,1120,1144]" pageId="34" pageNumber="481" value="1955-08-17" valueMax="1955-08-18" valueMin="1955-08-17">
<collectingDate id="EFE0676BFFF5FFDDFB83441AFAB9FBCC" box="[1089,1333,1120,1144]" pageId="34" pageNumber="481" value="1955-08-17" valueMax="1955-08-18" valueMin="1955-08-17">17 and 18 August 1955</collectingDate>
</date>
by
<collectorName id="26EFDD95FFF5FFDDFA9E441AFC02FB23" pageId="34" pageNumber="481">Raymond Laurent</collectorName>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFF5FFDDFC5E4405FAE0FB03" collectingDate="1921-07-27" collectionCode="RMCA, R" collectorName="Henri Schouteden" country="Democratic Republic of the Congo" location="Basongo" pageId="34" pageNumber="481" specimenCount="1" stateProvince="Kasai-Oriental">
<collectionCode id="ED0B2086FFF5FFDDFC5E4405FC64FB23" box="[924,1000,1151,1175]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="34" pageNumber="481" type="Museum">RMCA</collectionCode>
<collectionCode id="ED0B2086FFF5FFDDFC304405FB89FB23" box="[1010,1029,1151,1175]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="34" pageNumber="481">R</collectionCode>
.2947, female, collected at
<location id="8EC5EE98FFF5FFDDFAE144FAFAF0FB2C" LSID="urn:lsid:plazi:treatment:03B30955FFF5FFDBFC534104FEB5FB28:8EC5EE98FFF5FFDDFAE144FAFAF0FB2C" box="[1315,1404,1152,1176]" country="Democratic Republic of the Congo" name="Basongo" pageId="34" pageNumber="481" stateProvince="Kasai-Oriental">Basongo</location>
,
<collectingRegion id="49DE76A1FFF5FFDDFA484405FC19FB03" country="Democratic Republic of the Congo" name="Kasai-Oriental" pageId="34" pageNumber="481">Kasai Province</collectingRegion>
,
<collectingCountry id="F30DF8D3FFF5FFDDFC6244E5FC54FB03" box="[928,984,1183,1207]" name="Democratic Republic of the Congo" pageId="34" pageNumber="481">DRC</collectingCountry>
, on
<date id="FFA49E83FFF5FFDDFBC144E5FB0AFB03" box="[1027,1158,1183,1207]" pageId="34" pageNumber="481" value="1921-07-27">
<collectingDate id="EFE0676BFFF5FFDDFBC144E5FB0AFB03" box="[1027,1158,1183,1207]" pageId="34" pageNumber="481" value="1921-07-27">27 July 1921</collectingDate>
</date>
by
<collectorName id="26EFDD95FFF5FFDDFB6E44E5FAE0FB03" box="[1196,1388,1183,1207]" pageId="34" pageNumber="481">Henri Schouteden</collectorName>
</materialsCitation>
;
<materialsCitation id="3B72B21EFFF5FFDDFAB444E5FB49FB41" collectionCode="RMCA, R" collectorName="J. Stevenart" country="Democratic Republic of the Congo" location="Yokamba" municipality="Boende" pageId="34" pageNumber="481" specimenCount="1" stateProvince="Tshuapa Province">
<collectionCode id="ED0B2086FFF5FFDDFAB444E5FA4EFB03" box="[1398,1474,1183,1207]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="34" pageNumber="481" type="Museum">RMCA</collectionCode>
<collectionCode id="ED0B2086FFF5FFDDFCFB44C4FCC0FB62" box="[825,844,1214,1238]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="34" pageNumber="481">R</collectionCode>
.16752, male, collected at
<location id="8EC5EE98FFF5FFDDFB4044C4FB6CFB62" LSID="urn:lsid:plazi:treatment:03B30955FFF5FFDBFC534104FEB5FB28:8EC5EE98FFF5FFDDFB4044C4FB6CFB62" box="[1154,1248,1214,1238]" country="Democratic Republic of the Congo" municipality="Boende" name="Yokamba" pageId="34" pageNumber="481" stateProvince="Tshuapa Province">Yokamba</location>
(
<collectingMunicipality id="6BC12239FFF5FFDDFB3B44C4FADCFB62" box="[1273,1360,1214,1238]" pageId="34" pageNumber="481">Boende</collectingMunicipality>
),
<collectingRegion id="49DE76A1FFF5FFDDFAAE44C4FC19FB41" inferredBySuffix="true" pageId="34" pageNumber="481">Tshuapa Province</collectingRegion>
,
<collectingCountry id="F30DF8D3FFF5FFDDFC5D44A7FC5BFB41" box="[927,983,1245,1269]" name="Democratic Republic of the Congo" pageId="34" pageNumber="481">DRC</collectingCountry>
, in 1953 by
<collectorName id="26EFDD95FFF5FFDDFB9244A4FB49FB41" box="[1104,1221,1245,1269]" pageId="34" pageNumber="481">J. Stevenart</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA5B843FFF5FFDDFCFB455AFC68FA67" blockId="34.[825,1475,1311,1994]" pageId="34" pageNumber="481">
<emphasis id="B96E6451FFF5FFDDFCFB455AFC13FA83" box="[825,927,1312,1335]" italics="true" pageId="34" pageNumber="481">Diagnosis:</emphasis>
This species represents a moderately sized
<taxonomicName id="4C1AC3C0FFF5FFDDFCFB4544FC34FAE2" box="[825,952,1342,1366]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="genus">
<emphasis id="B96E6451FFF5FFDDFCFB4544FC34FAE2" box="[825,952,1342,1366]" italics="true" pageId="34" pageNumber="481">Lygodactylus</emphasis>
</taxonomicName>
[maximum SVL
<quantity id="4CE215A6FFF5FFDDFBAB4545FB48FAE2" box="[1129,1220,1343,1367]" metricMagnitude="-2" metricUnit="m" metricValue="3.47" pageId="34" pageNumber="481" unit="mm" value="34.7">34.7 mm</quantity>
(mean 33.6 ±
<quantity id="4CE215A6FFF5FFDDFA984545FA23FAE3" box="[1370,1455,1343,1367]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="34" pageNumber="481" unit="mm" value="1.3">1.3 mm</quantity>
)], with males having the most distinctive gular patterning of the
<taxonomicName id="4C1AC3C0FFF5FFDDFA6C4525FC18FA21" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF5FFDDFA6C4525FC18FA21" italics="true" pageId="34" pageNumber="481">L. gutturalis</emphasis>
</taxonomicName>
group (described below,
<figureCitation id="1321A4C6FFF5FFDDFB514507FB47FA21" box="[1171,1227,1405,1429]" captionStart="Figure 3" captionStartId="14.[129,194,1167,1191]" captionTargetBox="[131,1462,158,1136]" captionTargetId="figure-369@14.[126,1476,144,1139]" captionTargetPageId="14" captionText="Figure 3. Comparative figure including dorsal (or dorsolateral) and ventral views of preserved holotypes (with exception of Lygodactylus gutturalis that is represented by a paratype) of all species within the L.gutturalis subgroup used in the current study." pageId="34" pageNumber="481">Fig. 3</figureCitation>
). Females without gular pattern. Eight supralabials and 6–7 infralabials. Males with seven precloacal pores.
</paragraph>
<paragraph id="8BA5B843FFF5FFDCFC9745A1FDE4FF73" blockId="34.[825,1475,1311,1994]" lastBlockId="35.[113,763,144,669]" lastPageId="35" lastPageNumber="482" pageId="34" pageNumber="481">
This species can be differentiated from all taxa of the
<taxonomicName id="4C1AC3C0FFF5FFDDFAAA45A1FA4EFA47" authorityName="Gunther" authorityYear="1893" box="[1384,1474,1499,1523]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFF5FFDDFAAA45A1FA4EFA47" box="[1384,1474,1499,1523]" italics="true" pageId="34" pageNumber="481">angularis</emphasis>
</taxonomicName>
group taxa by the same osteological differences described above (e.g. postorbitofrontal reduced almost vestigial vs. well developed in the
<taxonomicName id="4C1AC3C0FFF5FFDDFC6D4643FB85F9E5" authorityName="Gunther" authorityYear="1893" box="[943,1033,1593,1617]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFF5FFDDFC6D4643FB85F9E5" box="[943,1033,1593,1617]" italics="true" pageId="34" pageNumber="481">angularis</emphasis>
</taxonomicName>
group) and a unique leopard-like pattern on the dorsum. Also, it can be easily differentiated from
<taxonomicName id="4C1AC3C0FFF5FFDDFA8D4623FA4EF9C4" authorityName="Gunther" authorityYear="1893" box="[1359,1474,1624,1648]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFF5FFDDFA8D4623FA4EF9C4" box="[1359,1474,1624,1648]" italics="true" pageId="34" pageNumber="481">L. angularis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1AC3C0FFF5FFDDFCAA4603FC54F924" authorityName="Marques, Ceriaco, Buehler, Bandeira, Janota &amp; Bauer" authorityYear="2020" box="[872,984,1656,1680]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="baptistai">
<emphasis id="B96E6451FFF5FFDDFCAA4603FC54F924" box="[872,984,1656,1680]" italics="true" pageId="34" pageNumber="481">L. baptistai</emphasis>
</taxonomicName>
based on the gular patterning (broken pattern vs. V-shaped pattern in
<taxonomicName id="4C1AC3C0FFF5FFDDFBEB46E2FB11F91B" authorityName="Gunther" authorityYear="1893" box="[1065,1181,1687,1711]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFF5FFDDFBEB46E2FB11F91B" box="[1065,1181,1687,1711]" italics="true" pageId="34" pageNumber="481">L. angularis</emphasis>
</taxonomicName>
and the unique pattern of
<taxonomicName id="4C1AC3C0FFF5FFDDFA6C46E2FC01F97A" authorityName="Marques, Ceriaco, Buehler, Bandeira, Janota &amp; Bauer" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="baptistai">
<emphasis id="B96E6451FFF5FFDDFA6C46E2FC01F97A" italics="true" pageId="34" pageNumber="481">L. baptistai</emphasis>
</taxonomicName>
described above; see
<figureCitation id="1321A4C6FFF5FFDDFBB746CCFB23F97A" box="[1141,1199,1718,1742]" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." pageId="34" pageNumber="481">Fig. 5</figureCitation>
). The species can be confused with
<taxonomicName id="4C1AC3C0FFF5FFDDFC6446ACFB8BF95A" box="[934,1031,1750,1774]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="heeneni">
<emphasis id="B96E6451FFF5FFDDFC6446ACFB8BF95A" box="[934,1031,1750,1774]" italics="true" pageId="34" pageNumber="481">L. heeneni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1AC3C0FFF5FFDDFBF746ACFB30F95A" baseAuthorityName="Laurent" baseAuthorityYear="1952" box="[1077,1212,1750,1774]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="paurospilus">
<emphasis id="B96E6451FFF5FFDDFBF746ACFB30F95A" box="[1077,1212,1750,1774]" italics="true" pageId="34" pageNumber="481">L. paurospilus</emphasis>
</taxonomicName>
based on the broken gular ornamentation; however, it can be differentiated from them as follows: unique leopard-like dorsal pattern in
<emphasis id="B96E6451FFF5FFDDFAC5476FFCEBF8F8" bold="true" pageId="34" pageNumber="481">
<taxonomicName id="4C1AC3C0FFF5FFDDFAC5476FFA13F898" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[1287,1439,1812,1836]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
<emphasis id="B96E6451FFF5FFDDFAC5476FFA13F898" bold="true" box="[1287,1439,1812,1836]" italics="true" pageId="34" pageNumber="481">L. leopardinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF5FFDDFA61476FFCEBF8F8" pageId="34" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(vs. light brown dorsal background with a light cream vertebral line of blotches in
<taxonomicName id="4C1AC3C0FFF5FFDDFBD9472EFBF0F8DF" box="[1051,1148,1875,1899]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="heeneni">
<emphasis id="B96E6451FFF5FFDDFBD9472EFBF0F8DF" box="[1051,1148,1875,1899]" italics="true" pageId="34" pageNumber="481">L. heeneni</emphasis>
</taxonomicName>
(
<figureCitation id="1321A4C6FFF5FFDDFB494729FB4FF8DF" box="[1163,1219,1875,1899]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="34" pageNumber="481">Fig. 7</figureCitation>
), similar to
<taxonomicName id="4C1AC3C0FFF5FFDDFAF9472EFA4EF8DF" baseAuthorityName="Laurent" baseAuthorityYear="1952" box="[1339,1474,1875,1899]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="paurospilus">
<emphasis id="B96E6451FFF5FFDDFAF9472EFA4EF8DF" box="[1339,1474,1875,1899]" italics="true" pageId="34" pageNumber="481">L. paurospilus</emphasis>
</taxonomicName>
(based on preserved specimens;
<figureCitation id="1321A4C6FFF5FFDDFB464708FB37F83E" box="[1156,1211,1906,1930]" captionStart="Figure 8" captionStartId="19.[115,180,1122,1146]" captionTargetBox="[114,1458,144,1094]" captionTargetId="figure-377@19.[114,1458,144,1094]" captionTargetPageId="19" captionText="Figure 8. Paratype of L. paurospilus [RMCA (=MRAC) R.27409] from Haute Lubitshako, South Kivu Province, DRC. A, Ventral and dorsal views of body. B, Details of head in ventral, dorsal, and lateral views (from top to bottom). C, Details of cloacal region. D, Details of right manus. Photos by J.L.R." pageId="34" pageNumber="481">Fig. 8</figureCitation>
; Supporting Information,
<figureCitation id="1321A4C6FFF5FFDDFCFB47E8FC0CF81E" box="[825,896,1938,1962]" captionStart="Figure 4" captionStartId="15.[113,178,747,771]" captionTargetBox="[117,1455,147,717]" captionTargetId="figure-570@15.[114,1458,144,720]" captionTargetPageId="15" captionText="Figure 4. A, PCA plots of the first principal component (PC1) vs. the second principal component (PC2). Species are represented by different colours included in the legend. B, Boxplots (top whisker–maximum value; lower whisker–minimum value; dark horizontal line–median) of the body size (SVL) and linear measurements analysed in this study.Values in all the variables with the exception of SVL, are represented using the Least-squares means interaction plot (function Lsmip, package lsmeans). For abbreviations and further details see Material and methods section." pageId="34" pageNumber="481">Fig. S4</figureCitation>
); broken pattern that converges anteriorly (vs. a broken pattern that converges posteriorly (
<emphasis id="B96E6451FFF5FFDDFB6747C8FB39F87D" box="[1189,1205,1970,1993]" italics="true" pageId="34" pageNumber="481">V</emphasis>
-shaped) in
<taxonomicName id="4C1AC3C0FFF5FFDDFAF047C8FA19F87D" box="[1330,1429,1969,1993]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="heeneni">
<emphasis id="B96E6451FFF5FFDDFAF047C8FA19F87D" box="[1330,1429,1969,1993]" italics="true" pageId="34" pageNumber="481">L. heeneni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1AC3C0FFF4FFDCFFB340EBFF73FF1C" baseAuthorityName="Laurent" baseAuthorityYear="1952" box="[113,255,144,168]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="paurospilus">
<emphasis id="B96E6451FFF4FFDCFFB340EBFF73FF1C" box="[113,255,144,168]" italics="true" pageId="35" pageNumber="482">L. paurospilus</emphasis>
</taxonomicName>
); fewer precloacal pores than
<taxonomicName id="4C1AC3C0FFF4FFDCFD8940EBFD3EFF1C" box="[587,690,144,168]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="heeneni">
<emphasis id="B96E6451FFF4FFDCFD8940EBFD3EFF1C" box="[587,690,144,168]" italics="true" pageId="35" pageNumber="482">L. heeneni</emphasis>
</taxonomicName>
(7 vs. 9–10); and osteological features described above.
</paragraph>
<paragraph id="8BA5B843FFF4FFDCFF4F40B4FE27FD29" blockId="35.[113,763,144,669]" pageId="35" pageNumber="482">
<emphasis id="B96E6451FFF4FFDCFF4F40B4FE60FF53" bold="true" box="[141,492,206,231]" pageId="35" pageNumber="482">
<taxonomicName id="4C1AC3C0FFF4FFDCFF4F40B4FE14FF52" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[141,408,206,230]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
<emphasis id="B96E6451FFF4FFDCFF4F40B4FE14FF52" bold="true" box="[141,408,206,230]" italics="true" pageId="35" pageNumber="482">Lygodactylus leopardinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF4FFDCFE5940B5FE60FF53" box="[411,492,207,231]" pageId="35" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the most distinctive species within the
<taxonomicName id="4C1AC3C0FFF4FFDCFED04094FEE1FEB2" box="[274,365,238,262]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF4FFDCFED04094FEE1FEB2" box="[274,365,238,262]" italics="true" pageId="35" pageNumber="482">gutturalis</emphasis>
</taxonomicName>
subgroup and the only species within the
<taxonomicName id="4C1AC3C0FFF4FFDCFF5A4174FF75FE91" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[152,249,270,293]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFF4FFDCFF5A4174FF75FE91" box="[152,249,270,293]" italics="true" pageId="35" pageNumber="482">picturatus</emphasis>
</taxonomicName>
group with a broken gular pattern. It can be easily distinguished from other closely related species based on its unique gular pattern and the leopard-like dorsal pattern, which is not present in related taxa. Females lack gular patterning (vs. weakly present in other members of the
<taxonomicName id="4C1AC3C0FFF4FFDCFDD041F1FDE1FE17" box="[530,621,395,419]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF4FFDCFDD041F1FDE1FE17" box="[530,621,395,419]" italics="true" pageId="35" pageNumber="482">gutturalis</emphasis>
</taxonomicName>
subgroup). It can also be differentiated based on minor meristic differences as follows: greater number of scales between the eyes (30–36 vs.
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<taxonomicName id="4C1AC3C0FFF4FFDCFF174193FEC7FDB5" box="[213,331,489,513]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF4FFDCFF174193FEC7FDB5" box="[213,331,489,513]" italics="true" pageId="35" pageNumber="482">L. gutturalis</emphasis>
</taxonomicName>
,
<quantity id="4CE215A6FFF4FFDCFE9A4192FE01FDB5" box="[344,397,488,513]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="35" pageNumber="482" unit="in" value="25.0">25 in</quantity>
<taxonomicName id="4C1AC3C0FFF4FFDCFE564193FD8AFDB5" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[404,518,489,513]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFF4FFDCFE564193FD8AFDB5" box="[404,518,489,513]" italics="true" pageId="35" pageNumber="482">L. dysmicus</emphasis>
</taxonomicName>
,
<quantity id="4CE215A6FFF4FFDCFDD04193FDE3FDB5" box="[530,623,489,513]" metricMagnitude="-1" metricUnit="m" metricValue="5.968999999999999" metricValueMax="7.112" metricValueMin="4.826" pageId="35" pageNumber="482" unit="in" value="23.5" valueMax="28.0" valueMin="19.0">19–28 in</quantity>
<emphasis id="B96E6451FFF4FFDCFDB44193FF2CFD94" bold="true" pageId="35" pageNumber="482">
<taxonomicName id="4C1AC3C0FFF4FFDCFDB44193FD5AFDB4" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[630,726,488,512]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFF4FFDCFDB44193FD5AFDB4" bold="true" box="[630,726,488,512]" italics="true" pageId="35" pageNumber="482">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF4FFDCFD1E4193FF2CFD94" pageId="35" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<quantity id="4CE215A6FFF4FFDCFF694272FE8BFD94" box="[171,263,520,544]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.588" metricValueMin="4.826" pageId="35" pageNumber="482" unit="in" value="20.5" valueMax="22.0" valueMin="19.0">19–22 in</quantity>
<emphasis id="B96E6451FFF4FFDCFECC4272FE7CFD94" bold="true" box="[270,496,519,544]" pageId="35" pageNumber="482">
<taxonomicName id="4C1AC3C0FFF4FFDCFECC4272FE1DFDAB" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[270,401,519,543]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFF4FFDCFECC4272FE1DFDAB" bold="true" box="[270,401,519,543]" italics="true" pageId="35" pageNumber="482">L. karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF4FFDCFE554272FE67FD94" box="[407,491,520,544]" pageId="35" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
and
<quantity id="4CE215A6FFF4FFDCFDE14272FDD4FD94" box="[547,600,520,544]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="35" pageNumber="482" unit="in" value="19.0">19 in</quantity>
<emphasis id="B96E6451FFF4FFDCFD9D4272FF4EFD8B" bold="true" pageId="35" pageNumber="482">
<taxonomicName id="4C1AC3C0FFF4FFDCFD9D4272FD76FDAB" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[607,762,519,543]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF4FFDCFD9D4272FD76FDAB" bold="true" box="[607,762,519,543]" italics="true" pageId="35" pageNumber="482">L. mirabundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF4FFDCFFB3425DFF4EFD8B" box="[113,194,551,575]" pageId="35" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); and lower number of ventral scales at midbody (14–15 vs.
<quantity id="4CE215A6FFF4FFDCFF53423CFF67FDEA" box="[145,235,582,606]" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="5.08" metricValueMin="3.81" pageId="35" pageNumber="482" unit="in" value="17.5" valueMax="20.0" valueMin="15.0">15–20 in</quantity>
<taxonomicName id="4C1AC3C0FFF4FFDCFF33423DFEE9FDEA" box="[241,357,582,606]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF4FFDCFF33423DFEE9FDEA" box="[241,357,582,606]" italics="true" pageId="35" pageNumber="482">L. gutturalis</emphasis>
</taxonomicName>
,
<quantity id="4CE215A6FFF4FFDCFEB2423DFE2EFDEA" box="[368,418,582,607]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="35" pageNumber="482" unit="in" value="21.0">21 in</quantity>
<taxonomicName id="4C1AC3C0FFF4FFDCFE6A423DFD94FDEA" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[424,536,582,606]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFF4FFDCFE6A423DFD94FDEA" box="[424,536,582,606]" italics="true" pageId="35" pageNumber="482">L. dysmicus</emphasis>
</taxonomicName>
,
<quantity id="4CE215A6FFF4FFDCFDE0423CFDF0FDEA" box="[546,636,582,606]" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="4.826" metricValueMin="4.064" pageId="35" pageNumber="482" unit="in" value="17.5" valueMax="19.0" valueMin="16.0">16–19 in</quantity>
<taxonomicName id="4C1AC3C0FFF4FFDCFD43423DFD78FDEA" authorityName=", Schmidt" authorityYear="1919" box="[641,756,582,606]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFF4FFDCFD43423DFD78FDEA" box="[641,756,582,606]" italics="true" pageId="35" pageNumber="482">L. depressus</emphasis>
</taxonomicName>
,
<quantity id="4CE215A6FFF4FFDCFFB3421CFF40FDCA" box="[113,204,614,638]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="35" pageNumber="482" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
<emphasis id="B96E6451FFF4FFDCFF10421CFE0AFDCA" bold="true" box="[210,390,613,638]" pageId="35" pageNumber="482">
<taxonomicName id="4C1AC3C0FFF4FFDCFF10421CFEBCFDC9" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[210,304,613,637]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFF4FFDCFF10421CFEBCFDC9" bold="true" box="[210,304,613,637]" italics="true" pageId="35" pageNumber="482">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF4FFDCFEF7421CFE0AFDCA" box="[309,390,614,638]" pageId="35" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<quantity id="4CE215A6FFF4FFDCFE53421CFE60FDCA" box="[401,492,614,638]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="35" pageNumber="482" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
<emphasis id="B96E6451FFF4FFDCFE33421CFD45FDCA" bold="true" box="[497,713,613,638]" pageId="35" pageNumber="482">
<taxonomicName id="4C1AC3C0FFF4FFDCFE33421CFDFFFDC9" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[497,627,613,637]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFF4FFDCFE33421CFDFFFDC9" bold="true" box="[497,627,613,637]" italics="true" pageId="35" pageNumber="482">L. karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF4FFDCFDBA421CFD45FDCA" box="[632,713,614,638]" pageId="35" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<quantity id="4CE215A6FFF4FFDCFFB342FFFF29FD29" box="[113,165,645,669]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="35" pageNumber="482" unit="in" value="21.0">21 in</quantity>
<emphasis id="B96E6451FFF4FFDCFF6842FCFE17FD29" bold="true" box="[170,411,645,669]" pageId="35" pageNumber="482">
<taxonomicName id="4C1AC3C0FFF4FFDCFF6842FCFEC8FD29" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[170,324,645,669]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF4FFDCFF6842FCFEC8FD29" bold="true" box="[170,324,645,669]" italics="true" pageId="35" pageNumber="482">L. mirabundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF4FFDCFE8B42FFFE17FD29" box="[329,411,645,669]" pageId="35" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="8BA5B843FFF4FFDCFFB342B6FD4CFCB0" blockId="35.[113,762,716,772]" pageId="35" pageNumber="482">
<emphasis id="B96E6451FFF4FFDCFFB342B6FF6DFD50" box="[113,225,716,740]" italics="true" pageId="35" pageNumber="482">Etymology:</emphasis>
The name
<taxonomicName id="4C1AC3C0FFF4FFDCFE9042B7FE44FD50" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[338,456,716,741]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="482" phylum="Chordata" rank="species" species="leopardinus">
‘
<emphasis id="B96E6451FFF4FFDCFE9A42B6FE44FD50" box="[344,456,716,740]" italics="true" pageId="35" pageNumber="482">leopardinus</emphasis>
</taxonomicName>
’ is an adjective referring to the leopard-like dorsal pattern present in males of this species.
</paragraph>
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<emphasis id="B96E6451FFF4FFDCFFB3435CFE1EFC89" box="[113,402,805,829]" italics="true" pageId="35" pageNumber="482">
Description of the
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</emphasis>
(
<figureCitation id="1321A4C6FFF4FFDCFE7E435FFD80FC89" box="[444,524,805,829]" captionStart="Figure 22" captionStartId="35.[113,178,1896,1920]" captionTargetBox="[114,1458,918,1868]" captionTargetId="figure-448@35.[114,1458,918,1868]" captionTargetPageId="35" captionText="Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="35" pageNumber="482">Fig. 22</figureCitation>
): Measurements and meristic characters of the
<typeStatus id="54A106E1FFF4FFDCFEB8433EFE5AFCE8" box="[378,470,836,860]" pageId="35" pageNumber="482" type="holotype">holotype</typeStatus>
are presented in Supporting Information, Table S9.
<tableCitation id="C6988DF8FFF4FFDCFBDE40EAFBE5FF1C" box="[1052,1129,144,168]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="35" pageNumber="482">Table 3</tableCitation>
. Adult male, with a snout–vent length (SVL) of
<quantity id="4CE215A6FFF4FFDCFC1B40D5FBBAFF73" box="[985,1078,175,199]" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" pageId="35" pageNumber="482" unit="mm" value="32.4">32.4 mm</quantity>
and a broken tail. Body slender and nape slightly distinct. Head as broad as body, and moderate head length (HW/HL 0.63). Canthus rostralis not prominent. Eye diameter 2.0 mm with circular pupil. Ear to eye distance almost twice the orbit diameter (
<quantity id="4CE215A6FFF4FFDCFB424157FB5BFEF1" box="[1152,1239,301,325]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="35" pageNumber="482" unit="mm" value="3.4">3.4 mm</quantity>
). Snout rounded and moderately pointed. Frontal granular scales larger than occipital scales, with 36 small granular scales between the eyes. Dorsal scales granular from rostral to tail. Rostral undivided, in slight contact with nostril. Three small symmetrical internasals. Eight supralabials and seven infralabials. Prenasal not in contact with the 1st supralabial. Nostril circular, in narrow contact with the rostral and bordered by 1st supralabial, prenasal, one supranasal, and one postnasal. Four rows of scales between supralabials and the orbit. Mental large, triangular, and in contact posteriorly with two rounded and symmetrical postmental scales. Five smaller post-postmental scales. Gular scales granular and smaller than ventral scales. Ventrals imbricate, rounded, and larger than dorsal scales, in 14 scale rows across the venter. Body relatively robust and slightly elongated (TRL/SVL 0.41). Seven precloacal pores. Fore- and hind limbs moderately short and stout; forearm intermediate (FL/SVL 0.15); tibia short (CL/SVL 0.17). Digits elongated, unwebbed, with 5–6 terminal scansors. Thumb rudimentary with a small terminal claw. Relative length of digits: I &lt;II = V &lt;III &lt;IV (manus); I &lt;II &lt;V &lt;III &lt;IV (pes).
</paragraph>
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<paragraph id="8BA5B843FFF4FFDCFFB34712FE80F80C" blockId="35.[113,1451,1896,1977]" pageId="35" pageNumber="482">
<emphasis id="B96E6451FFF4FFDCFFB34712FF5AF834" bold="true" box="[113,214,1896,1920]" pageId="35" pageNumber="482">Figure 22.</emphasis>
Holotype of
<emphasis id="B96E6451FFF4FFDCFE904713FDA2F835" bold="true" box="[338,558,1896,1921]" pageId="35" pageNumber="482">
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<emphasis id="B96E6451FFF4FFDCFE904713FE53F834" bold="true" box="[338,479,1896,1920]" italics="true" pageId="35" pageNumber="482">L. leopardinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF4FFDCFE214713FDA2F835" box="[483,558,1897,1921]" pageId="35" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC. A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R.
</paragraph>
</caption>
<paragraph id="8BA5B843FFF3FFDBFF434093FD96FDED" blockId="36.[129,778,233,601]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFF434093FEB6FEB5" box="[129,314,233,257]" italics="true" pageId="36" pageNumber="483">Coloration: In life</emphasis>
(
<typeStatus id="54A106E1FFF3FFDBFE894093FE23FEB5" box="[331,431,233,257]" pageId="36" pageNumber="483" type="holotype">holotype</typeStatus>
;
<figureCitation id="1321A4C6FFF3FFDBFE794093FDA3FEB5" box="[443,559,233,257]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="36" pageNumber="483">Fig. 23A–B</figureCitation>
), dorsal background colour light grey to brown, with a black reticulate dorsal pattern from snout to anterior insertion of the hind limbs; light cream blotches surrounded by black dots from eye to anterior insertion of the hind limb on each side of dorsum; tail brownish dorsally with light cream ocelli; throat white with two
<emphasis id="B96E6451FFF3FFDBFD9841FFFDE7FE29" bold="true" box="[602,619,389,413]" pageId="36" pageNumber="483">∩</emphasis>
<emphasis id="B96E6451FFF3FFDBFDA941FCFDFEFE29" box="[619,626,390,413]" italics="true" pageId="36" pageNumber="483">-</emphasis>
shaped broken dark chevrons; venter uniformly white.
<emphasis id="B96E6451FFF3FFDBFDE941DFFD31FE08" box="[555,701,421,444]" italics="true" pageId="36" pageNumber="483">In preservative</emphasis>
(
<typeStatus id="54A106E1FFF3FFDBFD0D41DFFF3EFE68" pageId="36" pageNumber="483" type="holotype">holotype</typeStatus>
;
<figureCitation id="1321A4C6FFF3FFDBFF7F41BEFE89FE68" box="[189,261,452,476]" captionStart="Figure 22" captionStartId="35.[113,178,1896,1920]" captionTargetBox="[114,1458,918,1868]" captionTargetId="figure-448@35.[114,1458,918,1868]" captionTargetPageId="35" captionText="Figure 22. Holotype of L. leopardinus sp. nov. (UTEP 22577) from Balolombo Village, Busira River, Équateur Province, DRC.A, Dorsal and ventral view of body. B, Details of right manus and (C) pes. D, Detailed view of head in ventral, dorsal, and lateral views (from top to bottom). Photos by J.L.R." pageId="36" pageNumber="483">Fig. 22</figureCitation>
), dorsum uniformly dark grey from snout to tail. Venter with silvery appearance, becoming brownish on tail and fore- and hind limbs; black reticulated dorsal pattern visible, with two white paravertebral lines (one on each side) from eyes to midbody; digits dark brown ventrally.
</paragraph>
<paragraph id="8BA5B843FFF3FFDBFF434201FD75FC9B" blockId="36.[129,778,635,816]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFF434201FF68FD26" box="[129,228,635,658]" italics="true" pageId="36" pageNumber="483">Variation:</emphasis>
In contrast to the
<typeStatus id="54A106E1FFF3FFDBFE6E4201FD86FD27" box="[428,522,635,659]" pageId="36" pageNumber="483" type="holotype">holotype</typeStatus>
, the female
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has a dorsal coloration in life that is entirely black with an orangish tail and dorsolateral dots in paravertebral lines that dissipate posteriorly, without a gular pattern (
<figureCitation id="1321A4C6FFF3FFDBFE1242A2FDC5FD45" box="[464,585,728,753]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="36" pageNumber="483">Fig. 23C–D</figureCitation>
). The lack of gular pattern is corroborated
<emphasis id="B96E6451FFF3FFDBFEAC4283FE74FCA4" box="[366,504,761,784]" italics="true" pageId="36" pageNumber="483">in preservative</emphasis>
, in the
<typeStatus id="54A106E1FFF3FFDBFD834282FD10FCA4" box="[577,668,760,784]" pageId="36" pageNumber="483" type="paratype">paratype</typeStatus>
, and in another female specimen (
<collectionCode id="ED0B2086FFF3FFDBFEB6436DFE4FFC9B" box="[372,451,791,815]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="36" pageNumber="483" type="Museum">RMCA</collectionCode>
<collectionCode id="ED0B2086FFF3FFDBFE0A436DFE57FC9B" box="[456,475,791,815]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="36" pageNumber="483">R</collectionCode>
.2947) from
<location id="8EC5EE98FFF3FFDBFD994362FD38FC84" LSID="urn:lsid:plazi:treatment:03B30955FFF5FFDBFC534104FEB5FB28:8EC5EE98FFF3FFDBFD994362FD38FC84" box="[603,692,792,816]" country="Democratic Republic of the Congo" name="Basongo" pageId="36" pageNumber="483">Basongo</location>
,
<collectingCountry id="F30DF8D3FFF3FFDBFD7F436DFD79FC9B" box="[701,757,791,815]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">DRC</collectingCountry>
.
</paragraph>
<paragraph id="8BA5B843FFF3FFDBFF43432AFF64FB90" blockId="36.[128,778,848,1060]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFF43432AFEE3FCDC" box="[129,367,848,872]" italics="true" pageId="36" pageNumber="483">Habitat and distribution</emphasis>
(
<figureCitation id="1321A4C6FFF3FFDBFE46432AFE4CFCDC" box="[388,448,848,872]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." pageId="36" pageNumber="483">Figs 6</figureCitation>
,
<figureCitation id="1321A4C6FFF3FFDBFE09432BFE69FCDD" box="[459,485,849,873]" captionStart="Figure 23" captionStartId="37.[113,178,1204,1228]" captionTargetBox="[114,1458,144,1176]" captionTargetId="figure-366@37.[114,1458,144,1176]" captionTargetPageId="37" captionText="Figure 23. A–B, Holotype (UTEP 22577) and (C–D) paratype (UTEP 22596) of L. leopardinus sp. nov. in life and photo of habitat from (E) type locality at Balolombo Village, Busira River, Équateur Province, DRC. F–G, Holotype (UTEP 22587) and (H–I) paratype (UTEP 22584) of L. gamblei sp. nov. in life and photo of habitat from (J) type locality at Manono, Tanganyika Province, DRC. Photographs by E.G." pageId="36" pageNumber="483">23</figureCitation>
):
<emphasis id="B96E6451FFF3FFDBFE3F432AFF55FC3C" bold="true" pageId="36" pageNumber="483">
<taxonomicName id="4C1AC3C0FFF3FFDBFE3F432AFC86FCDC" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[509,778,848,872]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="leopardinus" status="sp. nov.">
<emphasis id="B96E6451FFF3FFDBFE3F432AFC86FCDC" bold="true" box="[509,778,848,872]" italics="true" pageId="36" pageNumber="483">Lygodactylus leopardinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF3FFDBFF42430AFF55FC3C" box="[128,217,880,904]" pageId="36" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from
<collectingCountry id="F30DF8D3FFF3FFDBFE51430AFE55FC3C" box="[403,473,880,904]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">Congo</collectingCountry>
Basin primary rainforest at Balolombo, Katopa (Lomani River,
<collectingCountry id="F30DF8D3FFF3FFDBFE3143F5FDB5FC13" box="[499,569,911,935]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">Congo</collectingCountry>
River affluent), and Basongo. This pattern suggests a large distribution in the
<collectingCountry id="F30DF8D3FFF3FFDBFD0143D5FC85FC73" box="[707,777,943,967]" name="Democratic Republic of the Congo" pageId="36" pageNumber="483">Congo</collectingCountry>
Basin like its sister taxon,
<taxonomicName id="4C1AC3C0FFF3FFDBFE5043B5FD85FC52" authorityName=", Schmidt" authorityYear="1919" box="[402,521,974,998]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="483" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFF3FFDBFE5043B5FD85FC52" box="[402,521,974,998]" italics="true" pageId="36" pageNumber="483">L. depressus</emphasis>
</taxonomicName>
, which is found in sympatry at Lake Tumba and Katopa. However, it seems to be less abundant.
</paragraph>
<paragraph id="8BA5B843FFF3FFDBFF42443CFEB5FB28" blockId="36.[128,778,1094,1180]" pageId="36" pageNumber="483">
<emphasis id="B96E6451FFF3FFDBFF42443CFE93FBEA" box="[128,287,1094,1118]" italics="true" pageId="36" pageNumber="483">Natural history:</emphasis>
A diurnal and arboreal rainforest species. The
<typeStatus id="54A106E1FFF3FFDBFF42441FFF50FBC9" box="[128,220,1125,1149]" pageId="36" pageNumber="483" type="holotype">holotype</typeStatus>
was collected early in the morning on a tree about
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above the ground.
</paragraph>
</subSubSection>
</treatment>
</document>

398
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<mods:namePart id="C02616BF2BFB2847B1D198B4F1792A97">Cael, Garin</mods:namePart>
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<mods:namePart id="62826282959F13BD13E998CA22F7CCA4">Stanley, Edward L.</mods:namePart>
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<mods:namePart id="73DF76E00BC3849687DCB2473E870025">Hughes, Daniel F.</mods:namePart>
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<mods:namePart id="0F71BEC7D7597C51EAD1AF2DF9153EA1">Behangana, Mathias</mods:namePart>
</mods:name>
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<mods:namePart id="FF66439A4AAFF28CF5F30E170FD5771D">Masudi, Franck M.</mods:namePart>
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<mods:namePart id="47810CC09E97BBE798B288E78E0F4120">Pauwels, Olivier S. G.</mods:namePart>
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<mods:namePart id="04B2B035AC9189E5F7F7DC7DBFE8B8B2">Greenbaum, Eli</mods:namePart>
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<mods:typeOfResource id="BE98303876F14103EAE447F75FE4F15C">text</mods:typeOfResource>
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<mods:titleInfo id="B131695B291028B6DA78E5FC2146F1D0">
<mods:title id="21B1941E05F583D292615162522E835F">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="50CBF9D02F4F70F69E1CD6024E47F640">
<mods:date id="0F0B10B6505DA77187B0EB6C5BB13202">2024</mods:date>
<mods:detail id="41361F49930C602A939D6650DDED8B2C" type="series">
<mods:title id="6099615CE81DF8A0097F857F1F1F7E3C">Zool. J. Linn. Soc.</mods:title>
</mods:detail>
<mods:detail id="A2F807740A5C968E4297BA67F0A2A61D" type="pubDate">
<mods:number id="85EBE810CD31855ADDA970100FF4D743">2023-09-23</mods:number>
</mods:detail>
<mods:detail id="5ACA23002A5A48AA2CECBFC314466F05" type="volume">
<mods:number id="3981271C8D91A9CC1008F6CF283494E3">201</mods:number>
</mods:detail>
<mods:detail id="71E482E3106E57BCF1560D1CFA7C61C6" type="issue">
<mods:number id="2C63A8E2B3D00C0F8AE9B0881FAD9F76">2</mods:number>
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<mods:extent id="6C2958E8278788FA6D05552B962718D2" unit="page">
<mods:start id="0E15AF685C2E2F3EFD18CF0E3CEE417B">447</mods:start>
<mods:end id="B118F483EE465CBEC95555F3D51C5202">492</mods:end>
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<mods:location id="ADD424AC75203EF49A81782594F8D3D7">
<mods:url id="649ED4DEE31F4699A29C6D8E6B09FE7C">http://dx.doi.org/10.1093/zoolinnean/zlad123</mods:url>
</mods:location>
<mods:classification id="323284BB704FF42D8466706FCE9B35B6">journal article</mods:classification>
<mods:identifier id="4CB8A938FB74473FB5023D8262916ECE" type="DOI">10.1093/zoolinnean/zlad123</mods:identifier>
<mods:identifier id="A3FDBA8A6900B48CE620A165BE894110" type="ISSN">0024-4082</mods:identifier>
</mods:mods>
<treatment id="03B30955FFF6FFDDFF0E427BFC3FFEEA" LSID="urn:lsid:plazi:treatment:03B30955FFF6FFDDFF0E427BFC3FFEEA" httpUri="http://treatment.plazi.org/id/03B30955FFF6FFDDFF0E427BFC3FFEEA" lastPageId="34" lastPageNumber="481" pageId="33" pageNumber="480">
<subSubSection id="C300EBC8FFF6FFDEFF0E427BFD13FDAF" box="[204,671,513,539]" pageId="33" pageNumber="480" type="nomenclature">
<paragraph id="8BA5B843FFF6FFDEFF0E427BFD13FDAF" blockId="33.[204,671,513,539]" box="[204,671,513,539]" pageId="33" pageNumber="480">
<heading id="D0ED0F2FFFF6FFDEFF0E427BFD13FDAF" allCaps="true" bold="true" box="[204,671,513,539]" centered="true" fontSize="11" level="1" pageId="33" pageNumber="480" reason="1">
<emphasis id="B96E6451FFF6FFDEFF0E427BFD13FDAF" bold="true" box="[204,671,513,539]" pageId="33" pageNumber="480">
<taxonomicName id="4C1AC3C0FFF6FFDEFF0E427BFDB7FDAF" ID-CoL="C7JCZ" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[204,571,513,539]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="481" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF6FFDEFF0E427BFDB7FDAF" bold="true" box="[204,571,513,539]" italics="true" pageId="33" pageNumber="480">LYGODACTYLUS MIRABUNDUS</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF6FFDEFD82427BFD13FDAF" box="[576,671,513,539]" pageId="33" pageNumber="480" rank="species">SP.NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C300EBC8FFF6FFDDFE9F4252FC3FFEEA" lastPageId="34" lastPageNumber="481" pageId="33" pageNumber="480" type="description">
<paragraph id="8BA5B843FFF6FFDEFE9F4252FD83FDF4" blockId="33.[349,527,552,576]" box="[349,527,552,576]" pageId="33" pageNumber="480">
(
<figureCitation id="1321A4C6FFF6FFDEFEA54252FE20FDF4" box="[359,428,552,576]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="33" pageNumber="480">Fig. 21</figureCitation>
;
<tableCitation id="C6988DF8FFF6FFDEFE7B4252FD88FDF4" box="[441,516,552,576]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="33" pageNumber="480">Table 3</tableCitation>
)
</paragraph>
<paragraph id="8BA5B843FFF6FFDEFFB3421BFE20FD2C" blockId="33.[113,762,609,853]" pageId="33" pageNumber="480">
<emphasis id="B96E6451FFF6FFDEFFB3421BFEDFFDCD" box="[113,339,609,633]" italics="true" pageId="33" pageNumber="480">Zoobank registration:</emphasis>
https://zoobank.org/
<uuid id="FFBC8296FFF6FFDEFD83421BFE20FD2C" pageId="33" pageNumber="480">F4934B38-B746- 470D-B05B-BE56C1DEC7FE</uuid>
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<paragraph id="8BA5B843FFF6FFDEFF4F42DAFED5FCE1" blockId="33.[113,762,609,853]" pageId="33" pageNumber="480">
<emphasis id="B96E6451FFF6FFDEFF4F42DAFE71FD0C" bold="true" box="[141,509,672,696]" pageId="33" pageNumber="480">
<taxonomicName id="4C1AC3C0FFF6FFDEFF4F42DAFE2CFD0C" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[141,416,672,696]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF6FFDEFF4F42DAFE2CFD0C" bold="true" box="[141,416,672,696]" italics="true" pageId="33" pageNumber="480">Lygodactylus mirabundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF6FFDEFE6A42DAFE71FD0C" box="[424,509,672,696]" pageId="33" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the last taxon within subgroup B, described above as B2. It is sister to
<emphasis id="B96E6451FFF6FFDEFD5542BAFF4AFD43" bold="true" pageId="33" pageNumber="480">
<taxonomicName id="4C1AC3C0FFF6FFDEFD5542BAFD76FD63" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[663,762,703,727]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFF6FFDEFD5542BAFD76FD63" bold="true" box="[663,762,703,727]" italics="true" pageId="33" pageNumber="480">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF6FFDEFFB342A5FF4AFD43" box="[113,198,735,759]" pageId="33" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="B96E6451FFF6FFDEFF3F42A5FE53FD43" bold="true" box="[253,479,734,759]" pageId="33" pageNumber="480">
<taxonomicName id="4C1AC3C0FFF6FFDEFF3F42A5FE0EFD42" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[253,386,734,758]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFF6FFDEFF3F42A5FE0EFD42" bold="true" box="[253,386,734,758]" italics="true" pageId="33" pageNumber="480">L. karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF6FFDEFE4842A5FE53FD43" box="[394,479,735,759]" pageId="33" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, from which it differs by a minimum of 8.93% for the
<emphasis id="B96E6451FFF6FFDEFE4E4284FE3FFCA2" box="[396,435,766,790]" italics="true" pageId="33" pageNumber="480">16S</emphasis>
(uncorrected p-distance) mitochondrial gene (Table 2), and it lacks nuclear haplotype sharing for
<emphasis id="B96E6451FFF6FFDEFF564347FF5DFCE1" box="[148,209,829,853]" italics="true" pageId="33" pageNumber="480">RAG1</emphasis>
(
<figureCitation id="1321A4C6FFF6FFDEFF234347FEC4FCE1" box="[225,328,829,853]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability&gt; 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." pageId="33" pageNumber="480">Fig. 2B–C</figureCitation>
).
</paragraph>
<paragraph id="8BA5B843FFF6FFDEFCE840EAFBAFFE91" blockId="33.[810,1459,144,293]" pageId="33" pageNumber="480">
<materialsCitation id="3B72B21EFFF6FFDEFCE840EAFBAFFE91" collectingDate="2015-05-08" collectionCode="UTEP, CFS" collectorName="Wandege M. Muninga &amp; Chifundera Kusamba &amp; Mwenebatu M. Aristote." country="Democratic Republic of the Congo" elevation="551" latitude="-3.23949" location="Katako Kombe" longLatPrecision="1" longitude="24.25117" municipality="Sankuru Province" pageId="33" pageNumber="480" specimenCode="UTEP 22585, CFS 1142" specimenCount="1" typeStatus="holotype">
<emphasis id="B96E6451FFF6FFDEFCE840EAFC06FF1C" box="[810,906,144,168]" italics="true" pageId="33" pageNumber="480">
<typeStatus id="54A106E1FFF6FFDEFCE840EAFC09FF1C" box="[810,901,144,168]" pageId="33" pageNumber="480" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DBBC1038FFF6FFDEFC5E40EAFBA5FF1C" box="[924,1065,144,168]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="33" pageNumber="480" type="Museum">UTEP 22585</specimenCode>
(
<specimenCode id="DBBC1038FFF6FFDEFBFB40EAFB24FF1C" box="[1081,1192,144,168]" collectionCode="CFS" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="33" pageNumber="480">CFS 1142</specimenCode>
w), male without tail and with a ventral incision, collected at
<location id="8EC5EE98FFF6FFDEFB6C40D5FAC3FF73" LSID="urn:lsid:plazi:treatment:03B30955FFF6FFDDFF0E427BFC3FFEEA:8EC5EE98FFF6FFDEFB6C40D5FAC3FF73" box="[1198,1359,175,199]" country="Democratic Republic of the Congo" latitude="-3.23949" longLatPrecision="1" longitude="24.25117" municipality="Sankuru Province" name="Katako Kombe" pageId="33" pageNumber="480">Katako Kombe</location>
,
<collectingMunicipality id="6BC12239FFF6FFDEFA9F40D5FC0AFF53" pageId="33" pageNumber="480">Sankuru Province</collectingMunicipality>
,
<collectingCountry id="F30DF8D3FFF6FFDEFC5040B5FC46FF53" box="[914,970,207,231]" name="Democratic Republic of the Congo" pageId="33" pageNumber="480">DRC</collectingCountry>
,
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,
<geoCoordinate id="EE2EDE84FFF6FFDEFB9340B4FB4FFF52" box="[1105,1219,206,230]" degrees="24.25117" direction="east" orientation="longitude" pageId="33" pageNumber="480" precision="1" value="24.25117">E24.25117</geoCoordinate>
,
<elevation id="00375F70FFF6FFDEFB0C40B4FACAFF53" box="[1230,1350,206,231]" metricMagnitude="2" metricUnit="m" metricValue="5.51" pageId="33" pageNumber="480" unit="m" value="551.0">
<quantity id="4CE215A6FFF6FFDEFB0C40B4FA9EFF52" box="[1230,1298,206,230]" metricMagnitude="2" metricUnit="m" metricValue="5.51" pageId="33" pageNumber="480" unit="m" value="551.0">551 m</quantity>
a.s.l.
</elevation>
on
<date id="FFA49E83FFF6FFDEFAB340B5FCD2FEB2" pageId="33" pageNumber="480" value="2015-05-08">
<collectingDate id="EFE0676BFFF6FFDEFAB340B5FCD2FEB2" pageId="33" pageNumber="480" value="2015-05-08">8 May 2015</collectingDate>
</date>
by
<collectorName id="26EFDD95FFF6FFDEFC504094FB0AFEB2" box="[914,1158,238,262]" pageId="33" pageNumber="480">Wandege M. Muninga</collectorName>
,
<collectorName id="26EFDD95FFF6FFDEFB554094FAF0FEB2" box="[1175,1404,238,262]" pageId="33" pageNumber="480">Chifundera Kusamba</collectorName>
, and
<collectorName id="26EFDD95FFF6FFDEFCE84177FBAFFE91" box="[810,1059,269,293]" pageId="33" pageNumber="480">Mwenebatu M. Aristote.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BA5B843FFF6FFDEFCE8413DFA97FD8E" blockId="33.[810,1459,326,852]" pageId="33" pageNumber="480">
<emphasis id="B96E6451FFF6FFDEFCE8413DFB4DFEEA" box="[810,1217,326,350]" italics="true" pageId="33" pageNumber="480">
Diagnosis:
<taxonomicName id="4C1AC3C0FFF6FFDEFC68413CFB4DFEEA" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[938,1217,326,350]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF6FFDEFC68413CFB4DFEEA" bold="true" box="[938,1217,326,350]" pageId="33" pageNumber="480">Lygodactylus mirabundus</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A25DD92AFFF6FFDEFB0F413DFAAAFEEB" box="[1229,1318,327,351]" pageId="33" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
represents a moderately sized
<taxonomicName id="4C1AC3C0FFF6FFDEFC25411CFBEAFECA" box="[999,1126,358,382]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="B96E6451FFF6FFDEFC25411CFBEAFECA" box="[999,1126,358,382]" italics="true" pageId="33" pageNumber="480">Lygodactylus</emphasis>
</taxonomicName>
(SVL
<quantity id="4CE215A6FFF6FFDEFB77411CFA95FECA" box="[1205,1305,358,382]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" pageId="33" pageNumber="480" unit="mm" value="34.8">34.8 mm</quantity>
), with a gular pattern that is similar to the
<taxonomicName id="4C1AC3C0FFF6FFDEFB9441FCFB41FE29" box="[1110,1229,389,413]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF6FFDEFB9441FCFB41FE29" box="[1110,1229,389,413]" italics="true" pageId="33" pageNumber="480">L. gutturalis</emphasis>
</taxonomicName>
group. Seven to eight supralabials and six infralabials. Large triangular mental followed by three symmetrical postmental scales (
<figureCitation id="1321A4C6FFF6FFDEFACC41BEFADFFE68" box="[1294,1363,452,476]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="33" pageNumber="480">Fig. 21</figureCitation>
). Ventral pholidosis with large, flattened, and imbricate scales. Ventral scales usually denticulated posteriorly. Digits elongated with five terminal scansors under the fourth toe (
<tableCitation id="C6988DF8FFF6FFDEFB024258FA86FD8E" box="[1216,1290,546,570]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." pageId="33" pageNumber="480">Table 3</tableCitation>
).
</paragraph>
<paragraph id="8BA5B843FFF6FFDEFC87423BFACCFCA1" blockId="33.[810,1459,326,852]" pageId="33" pageNumber="480">
Like other
<taxonomicName id="4C1AC3C0FFF6FFDEFC71423BFBBEFDED" box="[947,1074,577,601]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="B96E6451FFF6FFDEFC71423BFBBEFDED" box="[947,1074,577,601]" italics="true" pageId="33" pageNumber="480">Lygodactylus</emphasis>
</taxonomicName>
within the
<taxonomicName id="4C1AC3C0FFF6FFDEFB634238FA99FDED" box="[1185,1301,577,601]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF6FFDEFB634238FA99FDED" box="[1185,1301,577,601]" italics="true" pageId="33" pageNumber="480">L. gutturalis</emphasis>
</taxonomicName>
group, this species can be easily differentiated from members of the
<taxonomicName id="4C1AC3C0FFF6FFDEFA82421BFA3FFDCC" authorityName="Gunther" authorityYear="1893" box="[1344,1459,608,632]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFF6FFDEFA82421BFA3FFDCC" box="[1344,1459,608,632]" italics="true" pageId="33" pageNumber="480">L. angularis</emphasis>
</taxonomicName>
group by its characteristic
<emphasis id="B96E6451FFF6FFDEFBED42FAFBCCFD2C" bold="true" box="[1071,1088,640,664]" pageId="33" pageNumber="480">∩</emphasis>
<emphasis id="B96E6451FFF6FFDEFB8242FBFBCBFD2C" box="[1088,1095,641,664]" italics="true" pageId="33" pageNumber="480">-</emphasis>
shaped gular chevrons (vs. V-shaped gular in the
<taxonomicName id="4C1AC3C0FFF6FFDEFC6742DAFB95FD03" authorityName="Gunther" authorityYear="1893" box="[933,1049,671,695]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B96E6451FFF6FFDEFC6742DAFB95FD03" box="[933,1049,671,695]" italics="true" pageId="33" pageNumber="480">L. angularis</emphasis>
</taxonomicName>
group), and from other members of the
<taxonomicName id="4C1AC3C0FFF6FFDEFCE842C5FC28FD62" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[810,932,703,726]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFF6FFDEFCE842C5FC28FD62" box="[810,932,703,726]" italics="true" pageId="33" pageNumber="480">L. picturatus</emphasis>
</taxonomicName>
group based on dorsal coloration (light brown with five or six laterodorsal cream ocelli vs. usually blueish dorsum with yellow to white head in the
<taxonomicName id="4C1AC3C0FFF6FFDEFBB64284FB62FCA1" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[1140,1262,766,789]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="picturatus">
<emphasis id="B96E6451FFF6FFDEFBB64284FB62FCA1" box="[1140,1262,766,789]" italics="true" pageId="33" pageNumber="480">L. picturatus</emphasis>
</taxonomicName>
group).
</paragraph>
<paragraph id="8BA5B843FFF6FFDDFC874366FE5FFDB5" blockId="33.[810,1459,326,852]" lastBlockId="34.[128,778,144,513]" lastPageId="34" lastPageNumber="481" pageId="33" pageNumber="480">
<emphasis id="B96E6451FFF6FFDEFC874366FB39FC81" bold="true" box="[837,1205,796,821]" pageId="33" pageNumber="480">
<taxonomicName id="4C1AC3C0FFF6FFDEFC874366FBD4FC80" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[837,1112,796,820]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF6FFDEFC874366FBD4FC80" bold="true" box="[837,1112,796,820]" italics="true" pageId="33" pageNumber="480">Lygodactylus mirabundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF6FFDEFBA24367FB39FC81" box="[1120,1205,797,821]" pageId="33" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a cryptic species and can be differentiated from other species within the
<taxonomicName id="4C1AC3C0FFF6FFDEFAFE4347FA3EFCE0" box="[1340,1458,828,852]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF6FFDEFAFE4347FA3EFCE0" box="[1340,1458,828,852]" italics="true" pageId="33" pageNumber="480">L. gutturalis</emphasis>
</taxonomicName>
subgroup by only a few morphometric and meristic characters. It can be differentiated from other
<taxonomicName id="4C1AC3C0FFF5FFDDFE2240CAFDD9FF73" box="[480,597,175,199]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF5FFDDFE2240CAFDD9FF73" box="[480,597,175,199]" italics="true" pageId="34" pageNumber="481">L. gutturalis</emphasis>
</taxonomicName>
subgroup species described above as follows: more precloacal pores [ten vs. 5–8 (in
<taxonomicName id="4C1AC3C0FFF5FFDDFF654095FE91FEB2" box="[167,285,238,262]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="gutturalis">
<emphasis id="B96E6451FFF5FFDDFF654095FE91FEB2" box="[167,285,238,262]" italics="true" pageId="34" pageNumber="481">L. gutturalis</emphasis>
</taxonomicName>
), nine (in
<taxonomicName id="4C1AC3C0FFF5FFDDFE534095FD88FEB2" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[401,516,238,262]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="dysmicus">
<emphasis id="B96E6451FFF5FFDDFE534095FD88FEB2" box="[401,516,238,262]" italics="true" pageId="34" pageNumber="481">L. dysmicus</emphasis>
</taxonomicName>
), seven (in
<taxonomicName id="4C1AC3C0FFF5FFDDFD414095FD74FEB2" authorityName=", Schmidt" authorityYear="1919" box="[643,760,238,262]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="depressus">
<emphasis id="B96E6451FFF5FFDDFD414095FD74FEB2" box="[643,760,238,262]" italics="true" pageId="34" pageNumber="481">L. depressus</emphasis>
</taxonomicName>
), 7–8 (in
<emphasis id="B96E6451FFF5FFDDFF134174FE09FE91" bold="true" box="[209,389,269,293]" pageId="34" pageNumber="481">
<taxonomicName id="4C1AC3C0FFF5FFDDFF134174FEA3FE91" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[209,303,269,293]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
<emphasis id="B96E6451FFF5FFDDFF134174FEA3FE91" bold="true" box="[209,303,269,293]" italics="true" pageId="34" pageNumber="481">L. kibera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF5FFDDFEF14177FE09FE91" box="[307,389,269,293]" pageId="34" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), and eight (in
<emphasis id="B96E6451FFF5FFDDFDE34174FD75FE91" bold="true" box="[545,761,269,293]" pageId="34" pageNumber="481">
<taxonomicName id="4C1AC3C0FFF5FFDDFDE34174FD2EFE91" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[545,674,269,293]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
<emphasis id="B96E6451FFF5FFDDFDE34174FD2EFE91" bold="true" box="[545,674,269,293]" italics="true" pageId="34" pageNumber="481">L. karamoja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF5FFDDFD6A4177FD75FE91" box="[680,761,269,293]" pageId="34" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), and a unique gular pattern comprising black markings on the mandibular margin that extend to the posterior region of the jaw, an inverted
<emphasis id="B96E6451FFF5FFDDFF3C4116FE81FE37" box="[254,269,364,387]" italics="true" pageId="34" pageNumber="481">Y</emphasis>
-shaped chevron that converges anteriorly, and a posterior middle line between the two chevron branches (vs. a
<emphasis id="B96E6451FFF5FFDDFF4241D0FF1DFE76" bold="true" box="[128,145,426,450]" pageId="34" pageNumber="481">∩</emphasis>
-shaped pattern in all the described species above). For comparison with other species not included above, see the diagnoses in the species descriptions below.
</paragraph>
<caption id="DF65E8CBFFF6FFDEFFB34710FF48F80E" pageId="33" pageNumber="480" startId="33.[113,178,1898,1922]" targetBox="[114,1458,920,1870]" targetPageId="33" targetType="figure">
<paragraph id="8BA5B843FFF6FFDEFFB34710FF48F80E" blockId="33.[113,1439,1898,1979]" pageId="33" pageNumber="480">
<emphasis id="B96E6451FFF6FFDEFFB34710FF5AF837" bold="true" box="[113,214,1898,1923]" pageId="33" pageNumber="480">Figure 21.</emphasis>
Holotype of
<emphasis id="B96E6451FFF6FFDEFE904711FDBDF837" bold="true" box="[338,561,1898,1923]" pageId="33" pageNumber="480">
<taxonomicName id="4C1AC3C0FFF6FFDEFE904711FE6EF836" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[338,482,1898,1922]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF6FFDEFE904711FE6EF836" bold="true" box="[338,482,1898,1922]" italics="true" pageId="33" pageNumber="480">L. mirabundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF6FFDEFE244711FDBDF837" box="[486,561,1899,1923]" pageId="33" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.
</paragraph>
</caption>
<paragraph id="8BA5B843FFF5FFDDFF424258FE05FD2C" blockId="34.[128,777,546,664]" pageId="34" pageNumber="481">
<emphasis id="B96E6451FFF5FFDDFF424258FF7CFD8E" box="[128,240,546,570]" italics="true" pageId="34" pageNumber="481">Etymology:</emphasis>
The name
<taxonomicName id="4C1AC3C0FFF5FFDDFEBF4258FE7AFD8E" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[381,502,546,570]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="mirabundus">
‘
<emphasis id="B96E6451FFF5FFDDFE404258FE7AFD8E" box="[386,502,546,570]" italics="true" pageId="34" pageNumber="481">mirabundus</emphasis>
</taxonomicName>
’ is a Latin adjective that means ‘astonishing or surprising’. The species is only known from a unique location in a transition zone between dry savannah and the Congolian Rainforest.
</paragraph>
<paragraph id="8BA5B843FFF5FFDDFF4242C0FF32F98A" blockId="34.[128,778,697,1598]" pageId="34" pageNumber="481">
<emphasis id="B96E6451FFF5FFDDFF4242C0FE0CFD65" box="[128,384,697,721]" italics="true" pageId="34" pageNumber="481">
Description of the
<typeStatus id="54A106E1FFF5FFDDFEEC42C3FE0CFD65" box="[302,384,697,721]" pageId="34" pageNumber="481" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="1321A4C6FFF5FFDDFE5742C3FE55FD65" box="[405,473,697,721]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="34" pageNumber="481">Fig. 21</figureCitation>
): Measurements and meristic characters of the
<typeStatus id="54A106E1FFF5FFDDFEF142A2FE03FD44" box="[307,399,728,752]" pageId="34" pageNumber="481" type="holotype">holotype</typeStatus>
are presented in. Adult male, with a snout–vent length (SVL) of
<quantity id="4CE215A6FFF5FFDDFE6E4282FD86FCBB" box="[428,522,760,784]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" pageId="34" pageNumber="481" unit="mm" value="34.8">34.8 mm</quantity>
and a broken tail. Body slender, nape almost indistinct. Head as broad as body, and moderate head length (HW/HL 0.79). Canthus rostralis not prominent. Eye diameter
<quantity id="4CE215A6FFF5FFDDFE51432CFE6AFCD9" box="[403,486,854,878]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="481" unit="mm" value="1.8">1.8 mm</quantity>
with circular pupil. Ear to eye distance almost double the orbit diameter (
<quantity id="4CE215A6FFF5FFDDFDA6430FFD37FC39" box="[612,699,885,909]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="34" pageNumber="481" unit="mm" value="3.6">3.6 mm</quantity>
). Snout rounded and slightly pointed. Frontal granular scales larger than occipital scales, with 19 small granular scales between the eyes. Dorsal scales granular from rostral to tail. Rostral undivided, in contact with 1st supralabial, prenasals, and one large internasal scale. Seven to eight supralabials and six infralabials. Prenasal scale present and in contact with 1st supralabial. Nostril circular and not in contact with the rostral and surrounded by 1st supralabial, prenasal, and one supranasal. Postnasal not contacting the nostril, separated by 1st supralabial, which it contacts posteriorly. Four rows of scales between supralabials and orbit. Mental large, triangular, and posteriorly in contact with three rounded, symmetric postmental scales. Five barely symmetric post-postmental scales. Gular scales granular, rounded, and slightly smaller than ventral scales. Ventral scales large and imbricate with 21 scales rows across the venter. Body relatively robust and slightly elongated (TRL/SVL 0.44). Ten precloacal pores. Fore- and hind limbs moderately short, stout; forearm medium-sized (FL/SVL 0.14); tibia short (CL/SVL 0.17). Digits elongated and unwebbed with 5–6 terminal scansors. Thumb rudimentary with a small terminal claw. Relative length of digits: I &lt;II = V &lt;III &lt;IV (manus); I &lt;II &lt;V &lt;III &lt;IV (pes).
</paragraph>
<paragraph id="8BA5B843FFF5FFDDFF42461AFD82F8C6" blockId="34.[128,778,1632,1906]" pageId="34" pageNumber="481">
<emphasis id="B96E6451FFF5FFDDFF42461AFEBEF9CC" box="[128,306,1632,1656]" italics="true" pageId="34" pageNumber="481">Coloration: In life</emphasis>
, unknown.
<emphasis id="B96E6451FFF5FFDDFE68461AFDBBF9C3" box="[426,567,1632,1655]" italics="true" pageId="34" pageNumber="481">In preservative</emphasis>
(
<typeStatus id="54A106E1FFF5FFDDFD86461AFD24F9CC" box="[580,680,1632,1656]" pageId="34" pageNumber="481" type="holotype">holotype</typeStatus>
;
<figureCitation id="1321A4C6FFF5FFDDFD73461AFD7BF9CC" box="[689,759,1632,1656]" captionStart="Figure 21" captionStartId="33.[113,178,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-400@33.[114,1458,920,1870]" captionTargetPageId="33" captionText="Figure 21. Holotype of L. mirabundus sp. nov. (UTEP 22585) from Katako Kombe, Sankuru Province, DRC. A, Dorsal and ventral view of body. B, Details of cloacal region and (C) right pes. D, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." pageId="34" pageNumber="481">Fig. 21</figureCitation>
): dorsum uniform dark brown-grey with lateral black spots barely visible; black spotted marks on upper and lower lips; fore- and hind limbs slightly lighter dorsally; black line from nostril to anterior insertion of the forelimb; venter uniform light cream all along the body with gular pattern of two lines (one on each side) that do not connect anteriorly, followed by an inverted
<emphasis id="B96E6451FFF5FFDDFD694766FD36F887" box="[683,698,1820,1843]" italics="true" pageId="34" pageNumber="481">Y</emphasis>
-shaped pattern with middle line between the branches of the
<emphasis id="B96E6451FFF5FFDDFD694746FD36F8E7" box="[683,698,1852,1875]" italics="true" pageId="34" pageNumber="481">Y</emphasis>
-shaped chevron. Lamellae whitish on all digits.
</paragraph>
<paragraph id="8BA5B843FFF5FFDDFF4247EEFAD5FEB1" blockId="34.[128,777,1939,1995]" lastBlockId="34.[825,1474,144,261]" pageId="34" pageNumber="481">
<emphasis id="B96E6451FFF5FFDDFF4247EEFEE3F818" box="[128,367,1940,1964]" italics="true" pageId="34" pageNumber="481">Habitat and distribution</emphasis>
(
<figureCitation id="1321A4C6FFF5FFDDFE4747EEFE31F81F" box="[389,445,1939,1964]" captionStart="Figure 5" captionStartId="16.[129,194,1857,1881]" captionTargetBox="[196,1409,145,1827]" captionTargetId="figure-10@16.[193,1409,143,1829]" captionTargetPageId="16" captionText="Figure 5. Gular and skull osteology comparisons between holotype material or representative material of all members of the L. gutturalis subgroup and the L. angularis group mentioned in this work.From top to bottom, ventral view gular section; dorsal, lateral, and ventral view of the skull; lateral, dorsal, medial, and ventral view of jaw; and number of tooth loci. PM = premaxilla, MX(R) = right maxilla, MX(L) = left maxilla, DN(R) = right dentary of the jaw." pageId="34" pageNumber="481">Fig. 5</figureCitation>
):
<materialsCitation id="3B72B21EFFF5FFDDFE1447E9FAD9FEB1" country="Democratic Republic of the Congo" elevation="2" location="The" pageId="34" pageNumber="481" specimenCount="1" typeStatus="holotype">
<emphasis id="B96E6451FFF5FFDDFE1447E9FF23F87F" bold="true" pageId="34" pageNumber="481">
<taxonomicName id="4C1AC3C0FFF5FFDDFE1447E9FD69F81F" authorityName="Lobón-Rovira &amp; Bauer &amp; Pinto &amp; Trape &amp; Conradie &amp; Kusamba &amp; Júlio &amp; Cael &amp; Stanley &amp; Hughes &amp; Behangana &amp; Masudi &amp; Pauwels &amp; Greenbaum" authorityYear="2024" box="[470,741,1939,1963]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="481" phylum="Chordata" rank="species" species="mirabundus" status="sp. nov.">
<emphasis id="B96E6451FFF5FFDDFE1447E9FD69F81F" bold="true" box="[470,741,1939,1963]" italics="true" pageId="34" pageNumber="481">Lygodactylus mirabundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25DD92AFFF5FFDDFD2947EEFF23F87F" pageId="34" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the type locality in
<collectingCountry id="F30DF8D3FFF5FFDDFD8247C9FDF4F87F" box="[576,632,1971,1995]" name="Democratic Republic of the Congo" pageId="34" pageNumber="481">DRC</collectingCountry>
.
<location id="8EC5EE98FFF5FFDDFD4347C9FD24F87F" LSID="urn:lsid:plazi:treatment:03B30955FFF6FFDDFF0E427BFC3FFEEA:8EC5EE98FFF5FFDDFD4347C9FD24F87F" box="[641,680,1971,1995]" country="Democratic Republic of the Congo" name="The" pageId="34" pageNumber="481">The</location>
<typeStatus id="54A106E1FFF5FFDDFD6F47C9FC85F87F" box="[685,777,1971,1995]" pageId="34" pageNumber="481" type="holotype">holotype</typeStatus>
was found on a large tree ~
<quantity id="4CE215A6FFF5FFDDFBFC40EAFBE9FF13" box="[1086,1125,144,168]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="481" unit="m" value="2.0">
<elevation id="00375F70FFF5FFDDFBFC40EAFBE9FF13" box="[1086,1125,144,168]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="481" unit="m" value="2.0">2 m</elevation>
</quantity>
above the ground at the edge of the transition zone between Miombo Woodland and the Congolian Rainforest. The association of this taxon with a particular biome will remain uncertain until more material is collected
</materialsCitation>
.
</paragraph>
<paragraph id="8BA5B843FFF5FFDDFCFB415DFC3FFEEA" blockId="34.[825,1474,295,350]" pageId="34" pageNumber="481">
<emphasis id="B96E6451FFF5FFDDFCFB415DFC5BFE8B" box="[825,983,295,319]" italics="true" pageId="34" pageNumber="481">Natural history:</emphasis>
Nothing is known about the natural history of this species.
</paragraph>
</subSubSection>
</treatment>
</document>