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<mods:title id="1D55F6923E1C3E22AFAE0F822E6C583D">New species of little-known crickets from the subfamily Pteroplistinae (Orthoptera, Grylloidea) from Sabah and contribution to their bioacoustics</mods:title>
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<mods:namePart id="4357517D598ABC3FBA73272D76B7A8EE">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="6B36C07E59A918A4D76C59EBA30621EB">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France</mods:affiliation>
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<mods:namePart id="1951BA03828210D1DB8CCF046979B06F">Gorochov, Andrei</mods:namePart>
<mods:affiliation id="1A502CDEE30A84C64343AB1ABB362D9B">Zoological Institute, Russian Academy of Sciences, 1 Universitetskaya Emb., St Petersburg 199034, Russia</mods:affiliation>
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<mods:namePart id="D70208B04F5E2308B03FBC5A065EC93C">Japir, Razy</mods:namePart>
<mods:affiliation id="00FA12BDF75ECA505C9D6A8593E8F5C0">Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia</mods:affiliation>
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<mods:namePart id="FA2768F387DAC9E9F2881E6D01DFB081">Chung, Arthur Y. C.</mods:namePart>
<mods:affiliation id="9305ABC3AB995009DBCD78AC24E9AF91">Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia</mods:affiliation>
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<mods:namePart id="ECEBD8767BACC7C3E0950091520BB5DB">Robillard, Tony</mods:namePart>
<mods:affiliation id="C6D2ED33C72C49D23320338FB4B7C99F">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France &amp; Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<taxonomicName id="9EF3EED9FFCD3E110C95F964FDA7009E" ID-CoL="4Q2NZ" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[151,599,1690,1717]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="223" phylum="Arthropoda" rank="species" species="lagrecai">
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<emphasis id="6B874948FFCD3E110C95F964FE7D009F" bold="true" box="[151,397,1690,1717]" italics="true" pageId="18" pageNumber="223">Pteroplistes lagrecai</emphasis>
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(
<figureCitation id="C1C889DFFFCD3E110C9DF940FF0B00F2" box="[159,251,1726,1752]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="18" pageNumber="223">Figs 2B</figureCitation>
,
<figureCitation id="C1C889DFFFCD3E110D04F940FED900F2" box="[262,297,1726,1752]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="18" pageNumber="223">3B</figureCitation>
,
<figureCitation id="C1C889DFFFCD3E110D37F940FEA800F2" box="[309,344,1726,1752]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="18" pageNumber="223">4B</figureCitation>
,
<figureCitation id="C1C889DFFFCD3E110D66F940FE6300F3" box="[356,403,1726,1753]" captionStart="FIGURE 15" captionStartId="19.[151,250,1771,1795]" captionTargetBox="[151,1436,181,1747]" captionTargetId="figure-17@19.[151,1436,181,1747]" captionTargetPageId="19" captionText="FIGURE 15. Pteroplistes silam sp. nov. calling on the tree trunk in Mount Silam (A), and Pteroplistes lagrecai Gorochov, 2004 calling on the tree in Sepilok (B)." pageId="18" pageNumber="223">15B</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="79E3B274FFCD3E110C95F8FDFCC30131" box="[151,819,1795,1819]" pageId="18" pageNumber="223">
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<taxonomicName id="9EF3EED9FFCD3E110C95F8FDFDB60131" ID-CoL="4Q2NZ" authority="Gorochov, 2004: 382" authorityName="Gorochov" authorityPageNumber="382" authorityYear="2004" box="[151,582,1795,1819]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="223" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFCD3E110C95F8FDFE990131" box="[151,361,1795,1819]" italics="true" pageId="18" pageNumber="223">Pteroplistes lagrecai</emphasis>
<bibRefCitation id="3D62E8ABFFCD3E110D6DF8FDFDB60131" author="Gorochov, A. V." box="[367,582,1795,1819]" pageId="18" pageNumber="223" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004: 382</bibRefCitation>
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<bibRefCitation id="3D62E8ABFFCD3E110E5EF8FDFCC30131" author="Gorochov, A. V." box="[604,819,1795,1819]" pageId="18" pageNumber="223" pagination="445 - 456" refId="ref7996" refString="Gorochov, A. V. (2022) New taxa of the cricket subfamily Pteroplistinae (Orthoptera: Gryllidae) from Indo Malayan Region. Amurian Zoological Journal, 14 (3), 445 - 456. https: // doi. org / 10.33910 / 2686 - 9519 - 2022 - 14 - 3 - 445 - 456" type="journal article" year="2022">Gorochov, 2022: 447</bibRefCitation>
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<emphasis id="6B874948FFCD3E110C95F8B7FE750149" bold="true" box="[151,389,1865,1891]" pageId="18" pageNumber="223">Material examined.</emphasis>
<materialsCitation id="E99B9F07FFCD3E110D93F8B7FC510181" collectingDate="2022-05-18" collectorName="T. Robillard &amp; M. K. Tan" country="Malaysia" county="Sandakan District" elevation="54" latitude="5.87395" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" pageId="18" pageNumber="223" specimenCount="1" specimenCount-male="1" stateProvince="Sabah">
EAST
<collectingCountry id="21E4D5CAFFCD3E110DE0F8B7FD830149" box="[482,627,1865,1891]" name="Malaysia" pageId="18" pageNumber="223">MALAYSIA</collectingCountry>
: •
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;
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,
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,
<collectingMunicipality id="B9280F20FFCD3E110842F8B7FB680149" box="[1088,1176,1865,1891]" pageId="18" pageNumber="223">Sepilok</collectingMunicipality>
,
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, dipterocarp forest;
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,
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,
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;
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<collectingDate id="3D094A72FFCD3E110F3BF893FC5901AD" box="[825,937,1901,1927]" pageId="18" pageNumber="223" value="2022-05-18">18.v.2022</collectingDate>
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, night (19h); on tree trunk;
<collectorName id="F406F08CFFCD3E1108F0F890FA8C01AD" box="[1266,1404,1901,1928]" pageId="18" pageNumber="223">T. Robillard</collectorName>
&amp;
<collectorName id="F406F08CFFCD3E110C95F86CFEF30186" box="[151,259,1938,1964]" pageId="18" pageNumber="223">M.K. Tan</collectorName>
leg.; TR22-27, call recording video 0062; MNHN-EO1288
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.
</paragraph>
</subSubSection>
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<emphasis id="6B874948FFCD3E110CC5F84BFEDC01E5" bold="true" box="[199,300,1973,1999]" pageId="18" pageNumber="223">Ecology.</emphasis>
This species was found calling on a tree trunk at night, where
<specimenCount id="4FF55ED3FFCD3E110FEBF848FB8101E5" box="[1001,1137,1973,2000]" count="2" pageId="18" pageNumber="223" type="female">two females</specimenCount>
were located near it.
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<emphasis id="6B874948FFCD3E110CC5F827FE9101D9" bold="true" box="[199,353,2009,2035]" pageId="18" pageNumber="223">Distribution.</emphasis>
EAST
<collectingCountry id="21E4D5CAFFCD3E110DB6F827FDBA01D9" box="[436,586,2009,2035]" name="Malaysia" pageId="18" pageNumber="223">MALAYSIA</collectingCountry>
:
<collectingRegion id="9B375BB8FFCD3E110E56F827FD6D01D9" box="[596,669,2009,2035]" country="Malaysia" name="Sabah" pageId="18" pageNumber="223">Sabah</collectingRegion>
: Sepilok.
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<paragraph id="594C955AFFCC3E100C95F915FE1A010D" blockId="19.[151,1437,1771,1831]" pageId="19" pageNumber="224">
<emphasis id="6B874948FFCC3E100C95F915FEEF0129" bold="true" box="[151,287,1771,1795]" pageId="19" pageNumber="224">FIGURE 15.</emphasis>
<taxonomicName id="9EF3EED9FFCC3E100D26F915FE290129" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[292,473,1771,1795]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="224" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFCC3E100D26F915FE290129" box="[292,473,1771,1795]" italics="true" pageId="19" pageNumber="224">Pteroplistes silam</emphasis>
</taxonomicName>
<emphasis id="6B874948FFCC3E100DDCF915FDDE0129" bold="true" box="[478,558,1771,1795]" pageId="19" pageNumber="224">
<taxonomicNameLabel id="70B4F433FFCC3E100DDCF915FDDE0129" box="[478,558,1771,1795]" pageId="19" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
calling on the tree trunk in Mount Silam (A), and
<taxonomicName id="9EF3EED9FFCC3E100826F915FA6D0129" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[1060,1437,1771,1795]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="224" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFCC3E100826F915FB040129" box="[1060,1268,1771,1795]" italics="true" pageId="19" pageNumber="224">Pteroplistes lagrecai</emphasis>
<bibRefCitation id="3D62E8ABFFCC3E1008F8F915FA6D0129" author="Gorochov, A. V." box="[1274,1437,1771,1795]" pageId="19" pageNumber="224" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
calling on the tree in Sepilok (B).
</paragraph>
</caption>
<paragraph id="594C955AFFCC3E170CC5F887FB31074F" blockId="19.[151,1437,1913,2012]" lastBlockId="20.[151,1437,151,357]" lastPageId="20" lastPageNumber="225" pageId="19" pageNumber="224">
<emphasis id="6B874948FFCC3E100CC5F887FDB101B9" bold="true" box="[199,577,1913,1939]" pageId="19" pageNumber="224">
Calling song (
<figureCitation id="C1C889DFFFCC3E100D6CF887FE3E01B9" box="[366,462,1913,1939]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="19" pageNumber="224">Figs 2D</figureCitation>
,
<figureCitation id="C1C889DFFFCC3E100DDFF887FDF001B9" box="[477,512,1913,1939]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="19" pageNumber="224">3D</figureCitation>
,
<figureCitation id="C1C889DFFFCC3E100E0DF887FDC401B9" box="[527,564,1913,1939]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="19" pageNumber="224">4D</figureCitation>
).
</emphasis>
Consists of an echeme-sequence. At 26°C, each echeme-sequence consists two parts, the first being 6±2 (510) isolated syllables, followed by the second being 11±4 (516) echemes made up of two syllables (doublet) each. For the first part, the average syllable duration is 22.1±1.5 ms (19.625.2 ms) and the average interval between consecutive syllables is 0.14±
<geoCoordinate id="3CC7F39DFFCB3E170F38FF69FC8F069B" box="[826,895,151,177]" degrees="0.03" direction="south" orientation="latitude" pageId="20" pageNumber="225" precision="555" value="-0.03">0.03 s</geoCoordinate>
(0.12
<geoCoordinate id="3CC7F39DFFCB3E170FCDFF69FBE6069B" box="[975,1046,151,177]" degrees="0.20" direction="south" orientation="latitude" pageId="20" pageNumber="225" precision="555" value="-0.2">0.20 s</geoCoordinate>
). For the second part, the average echeme duration is 58.8±1.8 ms (57.564.1 ms) and the average silent interval between consecutive echemes is the same as the average silent interval between consecutive syllables in the first part of the song. The average syllable duration of the doublet is 22.4±0.5 ms (21.523.2 ms), barely longer than that in the first part; and the average silent interval between the two syllables is 13.1±0.7 ms (11.814.1 ms). The frequency spectrum is pure-tonal and forms an harmonic series, with the energy peaking at a dominant fundamental frequency of 7.39 kHz.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,308 @@
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<mods:title id="051B08BDE815DFFEF6D9D7E4D30AB0A5">New species of little-known crickets from the subfamily Pteroplistinae (Orthoptera, Grylloidea) from Sabah and contribution to their bioacoustics</mods:title>
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<mods:affiliation id="95CBE5EA41EF5B27AF2817D87BC15264">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France &amp; Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<taxonomicName id="9EF3EED9FFD13E0D0C95FB06FD3E0339" authority="Tan, Gorochov &amp; Robillard" authorityName="Tan, Gorochov &amp; Robillard" authorityYear="2024" box="[151,718,1272,1299]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFD13E0D0C95FB06FE9C0339" bold="true" box="[151,364,1272,1299]" italics="true" pageId="14" pageNumber="219">Pteroplistes silam</emphasis>
Tan, Gorochov &amp; Robillard
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,
<taxonomicNameLabel id="70B4F433FFD13E0D0ED9FB07FCC80339" box="[731,824,1273,1299]" pageId="14" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="11E9C6D1FFD13E0D0C95FAE3FE010312" box="[151,497,1309,1336]" pageId="14" pageNumber="219" type="description">
<paragraph id="594C955AFFD13E0D0C95FAE3FE010312" blockId="14.[151,824,1272,1336]" box="[151,497,1309,1336]" pageId="14" pageNumber="219">
(
<figureCitation id="C1C889DFFFD13E0D0C9DFAE3FF0B031D" box="[159,251,1309,1335]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="14" pageNumber="219">Figs 2C</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0D04FAE3FED9031D" box="[262,297,1309,1335]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="14" pageNumber="219">3C</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0D37FAE3FEA8031D" box="[309,344,1309,1335]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="14" pageNumber="219">4C</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0D66FAE3FE72031D" box="[356,386,1309,1335]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="14" pageNumber="219">13</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0D8CFAE3FE5C031D" box="[398,428,1309,1335]" captionStart="FIGURE 14" captionStartId="18.[151,250,1548,1572]" captionTargetBox="[166,1421,194,1500]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 14. Male genitalia of Pteroplistes silam sp. nov. in posterodorsal (A), dorsal (B), ventral (C) and lateral (D) views, before (A) and after (BD) cleaning with KOH. Scale bar: 1 mm." pageId="14" pageNumber="219">14</figureCitation>
,
<figureCitation id="C1C889DFFFD13E0D0DBAFAE3FE190312" box="[440,489,1309,1336]" captionStart="FIGURE 15" captionStartId="19.[151,250,1771,1795]" captionTargetBox="[151,1436,181,1747]" captionTargetId="figure-17@19.[151,1436,181,1747]" captionTargetPageId="19" captionText="FIGURE 15. Pteroplistes silam sp. nov. calling on the tree trunk in Mount Silam (A), and Pteroplistes lagrecai Gorochov, 2004 calling on the tree in Sepilok (B)." pageId="14" pageNumber="219">15A</figureCitation>
)
</paragraph>
</subSubSection>
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<emphasis id="6B874948FFD13E0D0C95FA9BFE710355" bold="true" box="[151,385,1381,1407]" pageId="14" pageNumber="219">Material examined.</emphasis>
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<typeStatus id="86482BF8FFD13E0D0D8AFA9BFE090355" box="[392,505,1381,1407]" pageId="14" pageNumber="219" type="holotype">Holotype</typeStatus>
:
</emphasis>
EAST
<collectingCountry id="21E4D5CAFFD13E0D0E50FA9BFD180355" box="[594,744,1381,1407]" name="Malaysia" pageId="14" pageNumber="219">MALAYSIA</collectingCountry>
: •
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;
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<collectingMunicipality id="B9280F20FFD13E0D0FB8FA9BFBA00355" box="[954,1104,1381,1407]" pageId="14" pageNumber="219">Mount Silam</collectingMunicipality>
, near
<location id="5C2CC381FFD13E0D0891FA9BFAEA0355" LSID="urn:lsid:plazi:treatment:D15A244CFFD13E130C95FB06FC590193:5C2CC381FFD13E0D0891FA9BFAEA0355" box="[1171,1306,1381,1407]" country="Malaysia" latitude="4.96878" longLatPrecision="1" longitude="118.17189" municipality="Mount Silam" name="Lahad Datu" pageId="14" pageNumber="219" stateProvince="Sabah">Lahad Datu</location>
;
<geoCoordinate id="3CC7F39DFFD13E0D0924FA9BFA680355" box="[1318,1432,1381,1407]" degrees="4.96878" direction="north" orientation="latitude" pageId="14" pageNumber="219" precision="1" value="4.96878">N4.96878</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFD13E0D0C95FA77FED10389" box="[151,289,1417,1443]" degrees="118.17189" direction="east" orientation="longitude" pageId="14" pageNumber="219" precision="1" value="118.17189">E118.17189</geoCoordinate>
, 736.1±
<quantity id="9E0B38BFFFD13E0D0D7FFA77FE360389" box="[381,454,1417,1444]" metricMagnitude="0" metricUnit="m" metricValue="5.4" pageId="14" pageNumber="219" unit="m" value="5.4">
<elevation id="D2DE7269FFD13E0D0D7FFA77FE360389" box="[381,454,1417,1444]" metricMagnitude="0" metricUnit="m" metricValue="5.4" pageId="14" pageNumber="219" unit="m" value="5.4">5.4 m</elevation>
</quantity>
.a.s.l.; calling on tree trunk;
<date id="2D4DB39AFFD13E0D0F07FA77FC860389" box="[773,886,1417,1443]" pageId="14" pageNumber="219" value="2022-05-12">
<collectingDate id="3D094A72FFD13E0D0F07FA77FC860389" box="[773,886,1417,1443]" pageId="14" pageNumber="219" value="2022-05-12">12.v.2022</collectingDate>
</date>
, 21h56;
<collectorName id="F406F08CFFD13E0D0FD8FA77FBBA0389" box="[986,1098,1417,1443]" pageId="14" pageNumber="219">M.K. Tan</collectorName>
,
<collectorName id="F406F08CFFD13E0D0855FA77FB110389" box="[1111,1249,1417,1443]" pageId="14" pageNumber="219">T. Robillard</collectorName>
&amp;
<collectorName id="F406F08CFFD13E0D090BFA77FA920389" box="[1289,1378,1417,1443]" pageId="14" pageNumber="219">R. Japir</collectorName>
leg.; SBH.22.42 (
<collectionCode id="3FE20D9FFFD13E0D0D21FA53FEAE03ED" box="[291,350,1453,1479]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="14" pageNumber="219">FRC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD13E0D0CC5FA2FFD7D0179" pageId="14" pageNumber="219" type="diagnosis">
<paragraph id="594C955AFFD13E0D0CC5FA2FFD7D0179" blockId="14.[151,1437,1381,1911]" pageId="14" pageNumber="219">
<emphasis id="6B874948FFD13E0D0CC5FA2FFEB203C1" bold="true" box="[199,322,1489,1515]" pageId="14" pageNumber="219">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="9EF3EED9FFD13E0D0EB3FA2FFB9403C1" authority="Saussure, 1877" authorityName="Saussure" authorityYear="1877" box="[689,1124,1489,1515]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD13E0D0EB3FA2FFC5F03C1" box="[689,943,1489,1515]" italics="true" pageId="14" pageNumber="219">Pteroplistes acinaceus</emphasis>
Saussure, 1877
</taxonomicName>
from Malay Peninsula and
<taxonomicName id="9EF3EED9FFD13E0D0C95FA0BFCE50025" authority="Tan, Gorochov &amp; Wahab, 2019" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[151,789,1525,1551]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="bruneiensis">
<emphasis id="6B874948FFD13E0D0C95FA0BFE560025" box="[151,422,1525,1551]" italics="true" pageId="14" pageNumber="219">Pteroplistes bruneiensis</emphasis>
<bibRefCitation id="3D62E8ABFFD13E0D0DADFA0BFCE50025" author="Tan, M. K. &amp; Gorochov, A. V. &amp; Wahab, R. A. &amp; Japir, R. &amp; Chung, A. Y. C." box="[431,789,1525,1551]" pageId="14" pageNumber="219" pagination="101 - 117" refId="ref8234" refString="Tan, M. K., Gorochov, A. V., Wahab, R. A., Japir, R. &amp; Chung, A. Y. C. (2019) New taxa of crickets (Orthoptera: Grylloidea: Gryllinae, Phaloriinae and Pteroplistinae) from northern Borneo (Belait and Sandakan). Zootaxa, 4661 (1), 101 - 117. https: // doi. org / 10.11646 / zootaxa. 4661.1.4" type="journal article" year="2019">Tan, Gorochov &amp; Wahab, 2019</bibRefCitation>
</taxonomicName>
from Borneo in the ectophallic fold [rachis] of the male genitalia with a pair of rather large and sclerotized processes on its dorsal surface (dp) (before the apical part) but distinguished from them by these processes thickened, almost straight and with acute apices (these processes in
<taxonomicName id="9EF3EED9FFD13E0D0C95F99CFED50051" authorityName="Saussure" authorityYear="1877" box="[151,293,1634,1659]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD13E0D0C95F99CFED50051" box="[151,293,1634,1659]" italics="true" pageId="14" pageNumber="219">P. acinaceus</emphasis>
</taxonomicName>
with characteristic hooks distally, and in
<taxonomicName id="9EF3EED9FFD13E0D0F0AF99CFC570051" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[776,935,1633,1659]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="bruneiensis">
<emphasis id="6B874948FFD13E0D0F0AF99CFC570051" box="[776,935,1633,1659]" italics="true" pageId="14" pageNumber="219">P. bruneiensis</emphasis>
</taxonomicName>
, they are triangular). It is also similar to
<taxonomicName id="9EF3EED9FFD13E0D098AF99CFEF700B5" authorityName="Saussure" authorityYear="1877" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD13E0D098AF99CFEF700B5" italics="true" pageId="14" pageNumber="219">P. acinaceus</emphasis>
</taxonomicName>
,
<taxonomicName id="9EF3EED9FFD13E0D0D15F978FE4500B5" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[279,437,1669,1695]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="bruneiensis">
<emphasis id="6B874948FFD13E0D0D15F978FE4500B5" box="[279,437,1669,1695]" italics="true" pageId="14" pageNumber="219">P. bruneiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF3EED9FFD13E0D0DEDF97BFD2900B5" authorityName="Gorochov" authorityYear="2004" box="[495,729,1669,1695]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFD13E0D0DEDF97BFD2900B5" box="[495,729,1669,1695]" italics="true" pageId="14" pageNumber="219">Pteroplistes lagrecai</emphasis>
</taxonomicName>
by the shapes of the anal plate having a pair of small lobules (hooks), but differs from them by the lobules spaced widely apart (instead of nearly touching each other in
<taxonomicName id="9EF3EED9FFD13E0D098AF954FEE800CD" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="bruneiensis">
<emphasis id="6B874948FFD13E0D098AF954FEE800CD" italics="true" pageId="14" pageNumber="219">P. bruneiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF3EED9FFD13E0D0D52F933FDC800CD" authorityName="Gorochov" authorityYear="2004" box="[336,568,1741,1767]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFD13E0D0D52F933FDC800CD" box="[336,568,1741,1767]" italics="true" pageId="14" pageNumber="219">Pteroplistes lagrecai</emphasis>
</taxonomicName>
) and with the apices of the lobules slightly enlarged (instead of tapering into acute apices in
<taxonomicName id="9EF3EED9FFD13E0D0D48F90CFE270121" authorityName="Saussure" authorityYear="1877" box="[330,471,1778,1803]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD13E0D0D48F90CFE270121" box="[330,471,1778,1803]" italics="true" pageId="14" pageNumber="219">P. acinaceus</emphasis>
</taxonomicName>
). The new species also differs from congeners from
<collectingCountry id="21E4D5CAFFD13E0D083BF90FFB830121" box="[1081,1139,1777,1803]" name="India" pageId="14" pageNumber="219">India</collectingCountry>
(
<taxonomicName id="9EF3EED9FFD13E0D0887F90FFA660121" authorityName="Jaiswara" authorityYear="2014" box="[1157,1430,1777,1803]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="masinagudi">
<emphasis id="6B874948FFD13E0D0887F90FFA660121" box="[1157,1430,1777,1803]" italics="true" pageId="14" pageNumber="219">Pteroplistes masinagudi</emphasis>
</taxonomicName>
,
<taxonomicName id="9EF3EED9FFD13E0D0C95F8EBFE7B0105" authorityName="Jaiswara" authorityYear="2014" box="[151,395,1813,1839]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="kervasae">
<emphasis id="6B874948FFD13E0D0C95F8EBFE7B0105" box="[151,395,1813,1839]" italics="true" pageId="14" pageNumber="219">Pteroplistes kervasae</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF3EED9FFD13E0D0DCAF8EBFD330105" authorityName="Bolivar" authorityYear="1900" box="[456,707,1813,1839]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="219" phylum="Arthropoda" rank="species" species="platycleis">
<emphasis id="6B874948FFD13E0D0DCAF8EBFD330105" box="[456,707,1813,1839]" italics="true" pageId="14" pageNumber="219">Pteroplistes platycleis</emphasis>
</taxonomicName>
) by the head dorsum without distinct pale bands behind eyes, shapes of male anal plate and male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD13E0D0CC5F8A3FC28015D" box="[199,984,1885,1911]" pageId="14" pageNumber="219" type="etymology">
<paragraph id="594C955AFFD13E0D0CC5F8A3FC28015D" blockId="14.[151,1437,1381,1911]" box="[199,984,1885,1911]" pageId="14" pageNumber="219">
<emphasis id="6B874948FFD13E0D0CC5F8A3FEBE015D" bold="true" box="[199,334,1885,1911]" pageId="14" pageNumber="219">Etymology.</emphasis>
The species is named after Mount Silam, its
<typeStatus id="86482BF8FFD13E0D0F45F8A3FC87015D" box="[839,887,1885,1911]" pageId="14" pageNumber="219">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="0D8CC5D2FFD03E0C0C95F8A6FF3001BF" pageId="15" pageNumber="220" startId="15.[151,250,1880,1904]" targetBox="[151,1436,181,1857]" targetPageId="15" targetType="figure">
<paragraph id="594C955AFFD03E0C0C95F8A6FF3001BF" blockId="15.[151,1436,1880,1941]" pageId="15" pageNumber="220">
<emphasis id="6B874948FFD03E0C0C95F8A6FED1015B" bold="true" box="[151,289,1880,1905]" pageId="15" pageNumber="220">FIGURE 12.</emphasis>
<taxonomicName id="9EF3EED9FFD03E0C0D2BF8A7FE0B015B" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[297,507,1881,1905]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="220" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD03E0C0D2BF8A7FE0B015B" box="[297,507,1881,1905]" italics="true" pageId="15" pageNumber="220">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD03E0C0E01F8A7FDA5015B" bold="true" box="[515,597,1881,1905]" pageId="15" pageNumber="220">
<taxonomicNameLabel id="70B4F433FFD03E0C0E01F8A7FDA5015B" box="[515,597,1881,1905]" pageId="15" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: a male with two females on a tree trunk in Sepilok (A); the male started calling (B).
</paragraph>
</caption>
<subSubSection id="11E9C6D1FFCF3E130CC5FF68FED300CB" pageId="16" pageNumber="221" type="description">
<paragraph id="594C955AFFCF3E130CC5FF68FB3C044B" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5FF68FEA9069A" bold="true" box="[199,345,150,176]" pageId="16" pageNumber="221">Description.</emphasis>
Body distinctly dorsoventrally compressed, with head and pronotum finely pubescent. Head rostrum 1.3 times as wide as scape, with apex truncated (
<figureCitation id="C1C889DFFFCF3E130F34FF45FC5106FF" box="[822,929,187,213]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13A</figureCitation>
). Head wider than high in facial view (
<figureCitation id="C1C889DFFFCF3E13096DFF45FF3906D3" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13C</figureCitation>
). Maxillary palps elongated, with apical segment widened apically and subequal to third segment in length, and with subapical segment longest and slightly broadened apically (
<figureCitation id="C1C889DFFFCF3E130FB7FEFDFBEE0737" box="[949,1054,259,285]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13B</figureCitation>
). Eyes elongated in profile view, slightly protruding anteriorly (
<figureCitation id="C1C889DFFFCF3E130DE5FED9FDBB076B" box="[487,587,295,321]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13B</figureCitation>
); median ocellus small and rounded; lateral ocelli larger and more elongated; fenestrae between scapes rounded, small like median ocellus (
<figureCitation id="C1C889DFFFCF3E130F58FEB5FC32074F" box="[858,962,331,357]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13C</figureCitation>
). Pronotal disc 1.3 times as wide as long, finely pubescent along anterior and posterior margins; anterior margin of disc concave; posterior margin of disc straight; lateral margins sub straight and parallel (
<figureCitation id="C1C889DFFFCF3E130ECDFE6DFCC90787" box="[719,825,403,429]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13A</figureCitation>
). Pronotal lateral lobe 2.1 times as long as high, with ventral margin rising posteriorly (
<figureCitation id="C1C889DFFFCF3E130E19FE49FD7407FB" box="[539,644,439,465]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13B</figureCitation>
). TI with minute oval tympana on both sides; legs I and II generally pubescent (especially along ventral margin) and with a few stout setae usually located along dorsal margin, and their tarsus with a row of stout setae on ventral surface; FIII pubescent and flattened, without ventral spines; TIII also pubescent, with about 18 small and stout spines on each dorsal side, and with 2 long ventral apical spurs on inner margin and 4 shorter other apical spurs; hind basitarsus with about 7 inner and 7 outer denticles.
</paragraph>
<paragraph id="594C955AFFCF3E130CC5FD95FE8A021F" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5FD95FEFB04AF" bold="true" box="[199,267,619,645]" pageId="16" pageNumber="221">Male.</emphasis>
Metanotal gland with two lateral areas cream coloured, broadly triangular and slightly raised (
<figureCitation id="C1C889DFFFCF3E13092AFD95FA6004AF" box="[1320,1424,619,645]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13A</figureCitation>
). FW extending beyond abdominal apex, with dorsal field slightly longer than lateral field (
<figureCitation id="C1C889DFFFCF3E130895FD71FAB20483" box="[1175,1346,655,681]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Figs 13D, 13E</figureCitation>
); dorsal tegminal field (
<figureCitation id="C1C889DFFFCF3E130D44FD4DFE1F04E7" box="[326,495,691,717]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Figs 13D, 13E</figureCitation>
) in harp area with 6 oblique veins, mostly almost straight and parallel (except for posterior one which slightly sinuous); mirror in this field large and rounded, as wide as long, with anterior margin roundly angular, with posterior margin somewhat more widely rounded, and with two parallel dividing veins (these dividing veins arcuate and located close to one another); apical field about as long as length of mirror, separated from mirror by distinctive transverse cell (
<figureCitation id="C1C889DFFFCF3E130E83FCBDFCDD0577" box="[641,813,835,861]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Figs 13D, 13E</figureCitation>
); lateral field wide, with about 20 branches of Sc and about 10 cross-veins (some veins faint) between R and M, as well as with R and M mostly parallel and strongly converging towards the apex (
<figureCitation id="C1C889DFFFCF3E130DF1FC75FD50058F" box="[499,672,907,933]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Figs 13D, 13E</figureCitation>
). Hind wings slightly surpassing FWs. Anal plate broadly tongueshaped and slightly bilobed apically, with a pair of small lobules (hooks) in central part of dorsal surface but not very near each other; each of these lobules pointing inwards and tapering into subacute apex (
<figureCitation id="C1C889DFFFCF3E1308ACFC2DFAE105C7" box="[1198,1297,979,1005]" captionStart="FIGURE 13" captionStartId="17.[151,250,1744,1768]" captionTargetBox="[152,1436,181,1720]" captionTargetId="figure-17@17.[151,1436,181,1720]" captionTargetPageId="17" captionText="FIGURE 13. Pteroplistes silam sp. nov.: head and pronotum in dorsal (A) and lateral (B) views, face in anterior view (C), left (D) and right (E) FWs in dorsal view, supra-anal plate in dorsal view (F). Scale bars: 5 mm (A, D, E); 2 mm (rest)." pageId="16" pageNumber="221">Fig. 13F</figureCitation>
). Subgenital plate trapezoidal, about as long as wide; anterior margin wide, tapers slightly posteriorly, and with posterior margin deeply emarginated.
</paragraph>
<paragraph id="594C955AFFCF3E130CC5FBC1FE6A00B3" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
Male genitalia as shown in
<figureCitation id="C1C889DFFFCF3E130E1FFBC1FD870273" box="[541,631,1087,1113]" captionStart="FIGURE 14" captionStartId="18.[151,250,1548,1572]" captionTargetBox="[166,1421,194,1500]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 14. Male genitalia of Pteroplistes silam sp. nov. in posterodorsal (A), dorsal (B), ventral (C) and lateral (D) views, before (A) and after (BD) cleaning with KOH. Scale bar: 1 mm." pageId="16" pageNumber="221">Fig. 14</figureCitation>
. Pseudepiphallus [epiphallus] almost H-shaped, with transverse but moderately long median bridge between lateral parts, moderately short posterolateral lobes and long anterolateral parts; pseudepiphallic lophi [posterolateral epiphallic lobes] lamellate, with inner margin strongly sclerotized and concave, with lateral margin distinctly convex, and with apical part forming small hook directed dorso-externally; median bridge (mb) in shape of parallelogram, with anterior margin 1.9 times as wide as long; anterolateral parts of pseudepiphallus lamellate, sinuate and divergent, anteriorly fused with rami. Ectophallic fold [rachis (= guiding rod)] very strongly sclerotized and directed almost perpendicularly downwards in relation to the longitudinal axis of genitalia, with distal half having anterior margin slightly convex, with apical third tapering into acute apex, with basal part having upper process (up) almost perpendicular to ectophallic fold (this process distally bifurcated into two short triangular lobes with subacute apices); dorsum of upper process with pair of dark lateral sclerites (dp) projecting laterally and posteriorly, these projections connect with ectophallic apodemes. Ectophallic apodemes [endoparameres] strongly sclerotized, slender, with middle parts almost touching each other, and with anterior parts in shape of long arcuate ribbons curved downwards and aside (
<figureCitation id="C1C889DFFFCF3E130F58FA11FC300023" box="[858,960,1519,1545]" captionStart="FIGURE 14" captionStartId="18.[151,250,1548,1572]" captionTargetBox="[166,1421,194,1500]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 14. Male genitalia of Pteroplistes silam sp. nov. in posterodorsal (A), dorsal (B), ventral (C) and lateral (D) views, before (A) and after (BD) cleaning with KOH. Scale bar: 1 mm." pageId="16" pageNumber="221">Fig. 14B</figureCitation>
). Endophallic sclerite [formula (= mold of spermatophore attachment plate)] consists of three small but elongated plates (a pair of lateral oblique plates and unpaired transverse one) as well as a pair of larger and longer but oblique and isolated plates in anteroventral parts of genitalia. Rami rather short and narrow, strongly (arcuately) curved, lamellate and with anterior end tapering almost into acute apex.
</paragraph>
<paragraph id="594C955AFFCF3E130CC5F95DFED300CB" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5F95DFE8E0097" bold="true" box="[199,382,1699,1725]" pageId="16" pageNumber="221">Measurements.</emphasis>
<specimenCount id="4FF55ED3FFCF3E130D9BF95DFE5E0097" box="[409,430,1699,1725]" count="1" pageId="16" pageNumber="221" type="male"></specimenCount>
<typeStatus id="86482BF8FFCF3E130DB5F95DFDED0097" box="[439,541,1699,1725]" pageId="16" pageNumber="221" type="holotype">holotype</typeStatus>
: BL = 14.4; BWL = 18.8; HL = 1.9; PronL = 3.3; PronW = 4.5; FWL = 15.3; FWW = 6.6.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFCF3E130CC5F915FBE4012F" box="[199,1044,1770,1797]" pageId="16" pageNumber="221" type="biology_ecology">
<paragraph id="594C955AFFCF3E130CC5F915FBE4012F" blockId="16.[151,1437,150,1977]" box="[199,1044,1770,1797]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5F915FEDC012F" bold="true" box="[199,300,1771,1797]" pageId="16" pageNumber="221">Ecology.</emphasis>
This species was found calling on a tree trunk at night (
<figureCitation id="C1C889DFFFCF3E130F9DF915FBF8012F" box="[927,1032,1770,1797]" captionStart="FIGURE 15" captionStartId="19.[151,250,1771,1795]" captionTargetBox="[151,1436,181,1747]" captionTargetId="figure-17@19.[151,1436,181,1747]" captionTargetPageId="19" captionText="FIGURE 15. Pteroplistes silam sp. nov. calling on the tree trunk in Mount Silam (A), and Pteroplistes lagrecai Gorochov, 2004 calling on the tree in Sepilok (B)." pageId="16" pageNumber="221">Fig. 15A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFCF3E130CC5F8F0FC590193" pageId="16" pageNumber="221" type="distribution">
<paragraph id="594C955AFFCF3E130CC5F8F0FCCA0103" blockId="16.[151,1437,150,1977]" box="[199,826,1806,1833]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5F8F0FE910102" bold="true" box="[199,353,1806,1832]" pageId="16" pageNumber="221">Distribution.</emphasis>
EAST
<collectingCountry id="21E4D5CAFFCF3E130DB6F8F1FDBA0103" box="[436,586,1807,1833]" name="Malaysia" pageId="16" pageNumber="221">MALAYSIA</collectingCountry>
:
<collectingRegion id="9B375BB8FFCF3E130E56F8F1FD6D0103" box="[596,669,1807,1833]" country="Malaysia" name="Sabah" pageId="16" pageNumber="221">Sabah</collectingRegion>
: Mount Silam
</paragraph>
<paragraph id="594C955AFFCF3E130CC5F8CCFC590193" blockId="16.[151,1437,150,1977]" pageId="16" pageNumber="221">
<emphasis id="6B874948FFCF3E130CC5F8CCFDC50166" bold="true" box="[199,565,1842,1869]" pageId="16" pageNumber="221">
Calling song (
<figureCitation id="C1C889DFFFCF3E130D6BF8CCFE360166" box="[361,454,1842,1868]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="16" pageNumber="221">Figs 2C</figureCitation>
,
<figureCitation id="C1C889DFFFCF3E130DD1F8CCFE060166" box="[467,502,1842,1868]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="16" pageNumber="221">3C</figureCitation>
,
<figureCitation id="C1C889DFFFCF3E130E00F8CCFDD70166" box="[514,551,1842,1868]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="16" pageNumber="221">4C</figureCitation>
).
</emphasis>
Consists of a trill made up of long syllables irregular spaced-apart. At 23.6°C, average syllable duration is 26.8±1.1 ms (25.629.7 ms) and average silent interval between consecutive syllables is 115.8±40.1 ms (89.8249.2 ms). The frequency spectrum is pure-tonal and forms a clear harmonic series, with the energy peaking at a dominant fundamental frequency of 5.34 kHz.
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,141 @@
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Genus
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<bibRefCitation id="3D62E8ABFFD33E0F0E92FA0BFCC50027" author="Kirby, W. F." box="[656,821,1525,1549]" pageId="12" pageNumber="217" refId="ref8188" refString="Kirby, W. F. (1906) A Synonymic Catalogue of Orthoptera. Vol. 2. Orthoptera Saltatoria. Part I. (Achetidae et Phasgonuridae). Trustees of the British Museum, London, VIII + 562 + 25 pp." type="book" year="1906">Kirby, 1906: 62</bibRefCitation>
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;
<treatmentCitation id="D852B34BFFD33E0F0F43FA0BFBFA0027" author="Chopard, L." box="[833,1034,1525,1549]" page="254" pageId="12" pageNumber="217" year="1968">
<bibRefCitation id="3D62E8ABFFD33E0F0F43FA0BFBFA0027" author="Chopard, L." box="[833,1034,1525,1549]" pageId="12" pageNumber="217" refId="ref7631" refString="Chopard, L. (1968) Fam. Gryllidae: Subfam. Mogoplistinae, Myrmecophilinae, Scleropterinae, Cachoplistinae, Pteroplistinae, Pentacentrinae, Phalangopsinae, Trigonidiinae, Eneopterinae; Fam. Oecanthidae, Gryllotalpidae. In: Beier, M. (Ed.), Orthopterorum Catalogus. Vol. 12. Junk, ' s-Gravenhage, The Hague, pp. 213 - 500." type="book" year="1968">Chopard 1968: 254</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="D852B34BFFD33E0F0814FA0BFB150027" author="Chopard, L." box="[1046,1253,1525,1549]" page="230" pageId="12" pageNumber="217" year="1969">
<bibRefCitation id="3D62E8ABFFD33E0F0814FA0BFB150027" author="Chopard, L." box="[1046,1253,1525,1549]" pageId="12" pageNumber="217" refId="ref7700" refString="Chopard, L. (1969) Grylloidea. In: Sewell, R. B. S. (Ed.), The fauna of India and the adjacent countries. Orthoptera. Vol. 2. Zoological Survey of India and Baptist Mission Press, Calcutta, pp. 1 - 421." type="book" year="1969">Chopard, 1969: 230</bibRefCitation>
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;
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;
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<bibRefCitation id="3D62E8ABFFD33E0F0CF9F9EDFD5C0006" author="Jaiswara, R. &amp; Desutter-Grandcolas, L." box="[251,684,1555,1580]" pageId="12" pageNumber="217" pagination="96 - 108" refId="ref8112" refString="Jaiswara, R., &amp; Desutter-Grandcolas, L. (2014) Revision of the genus Pteroplistes in India, with the description of two new species Pteroplistes kervasae Jaiswara, n. sp. and Pteroplistes masinagudi Jaiswara, n. sp. (Orthoptera, Grylloidea, Pteroplistinae). Zootaxa, 3814 (1), 96 - 108. https: // doi. org / 10.11646 / zootaxa. 3814.1.5" type="journal article" year="2014">Jaiswara &amp; Desutter-Grandcolas, 2014: 98</bibRefCitation>
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;
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Tan
<emphasis id="6B874948FFD33E0F0EE0F9EAFCE50001" box="[738,789,1555,1579]" italics="true" pageId="12" pageNumber="217">et al.</emphasis>
, 2019
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; 10;
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<bibRefCitation id="3D62E8ABFFD33E0F0F8AF9EDFBAF0006" author="Gorochov, A. V." box="[904,1119,1555,1580]" pageId="12" pageNumber="217" pagination="445 - 456" refId="ref7996" refString="Gorochov, A. V. (2022) New taxa of the cricket subfamily Pteroplistinae (Orthoptera: Gryllidae) from Indo Malayan Region. Amurian Zoological Journal, 14 (3), 445 - 456. https: // doi. org / 10.33910 / 2686 - 9519 - 2022 - 14 - 3 - 445 - 456" type="journal article" year="2022">Gorochov, 2022: 447</bibRefCitation>
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<emphasis id="6B874948FFD33E0F0C95F9A4FEC50059" bold="true" box="[151,309,1625,1652]" pageId="12" pageNumber="217">
<typeStatus id="86482BF8FFD33E0F0C95F9A4FF22005E" box="[151,210,1626,1652]" pageId="12" pageNumber="217">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="9EF3EED9FFD33E0F0D3EF9A4FD54005E" ID-CoL="4Q2NV" authority="Saussure" authorityName="Saussure" authorityYear="1877" box="[316,676,1626,1652]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="217" phylum="Arthropoda" rank="species" species="acinaceus">
<emphasis id="6B874948FFD33E0F0D3EF9A4FDC70059" box="[316,567,1626,1652]" italics="true" pageId="12" pageNumber="217">Pteroplistes acinaceus</emphasis>
Saussure
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, by subsequent designation
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@ -0,0 +1,371 @@
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<heading id="02042236FFD73E0B0C95FAF0FCAC0303" bold="true" box="[151,860,1294,1321]" fontSize="11" level="1" pageId="8" pageNumber="213" reason="1">
<emphasis id="6B874948FFD73E0B0C95FAF0FCAC0303" bold="true" box="[151,860,1294,1321]" pageId="8" pageNumber="213">
<taxonomicName id="9EF3EED9FFD73E0B0C95FAF0FD010303" authority="Tan, Gorochov &amp; Robillard" authorityName="Tan, Gorochov &amp; Robillard" authorityYear="2024" box="[151,753,1294,1321]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD73E0B0C95FAF0FE7E0303" bold="true" box="[151,398,1294,1321]" italics="true" pageId="8" pageNumber="213">Tembelingiola kabili</emphasis>
Tan, Gorochov &amp; Robillard
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<paragraph id="594C955AFFD73E0B0C95FACDFE190367" blockId="8.[151,860,1294,1357]" box="[151,489,1331,1357]" pageId="8" pageNumber="213">
(
<figureCitation id="C1C889DFFFD73E0B0C9DFACDFF0B0367" box="[159,251,1331,1357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="8" pageNumber="213">Figs 2B</figureCitation>
,
<figureCitation id="C1C889DFFFD73E0B0D04FACDFED90367" box="[262,297,1331,1357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="8" pageNumber="213">3B</figureCitation>
,
<figureCitation id="C1C889DFFFD73E0B0D37FACDFEA80367" box="[309,344,1331,1357]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="8" pageNumber="213">4B</figureCitation>
,
<figureCitation id="C1C889DFFFD73E0B0D66FACDFE6C0367" box="[356,412,1331,1357]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="8" pageNumber="213">7EI</figureCitation>
,
<figureCitation id="C1C889DFFFD73E0B0DAAFACDFE120367" box="[424,482,1331,1357]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStartId-0="12.[151,250,1350,1374]" captionStartId-1="13.[151,250,1588,1612]" captionStartId-2="14.[151,250,1166,1190]" captionStartId-3="15.[151,250,1880,1904]" captionTargetBox-0="[176,1425,193,1326]" captionTargetBox-1="[151,1436,345,1564]" captionTargetBox-2="[151,1436,181,1143]" captionTargetBox-3="[151,1436,181,1857]" captionTargetId-0="figure-20@12.[151,1436,181,1326]" captionTargetId-1="figure-44@13.[151,1436,345,1564]" captionTargetId-2="figure-20@14.[151,1436,181,1143]" captionTargetId-3="figure-17@15.[151,1436,181,1857]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="FIGURE 9. Habitus of Tembelingiola kabili sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." captionText-1="FIGURE 10. Male genitalia of Tembelingiola kabili sp. nov. in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm." captionText-2="FIGURE 11. Habitus of Tembelingiola kabili sp. nov., female in dorsal (A) and lateral (B) views. Scale bars: 2 mm." captionText-3="FIGURE 12. Tembelingiola kabili sp. nov.: a male with two females on a tree trunk in Sepilok (A); the male started calling (B)." pageId="8" pageNumber="213">912</figureCitation>
)
</paragraph>
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<emphasis id="6B874948FFD73E0B0C95FA85FE7703BF" bold="true" box="[151,391,1403,1429]" pageId="8" pageNumber="213">Material examined.</emphasis>
<materialsCitation id="E99B9F07FFD73E0B0D96FA85FD7A03F7" collectingDate="2022-05-18" collectionCode="FRC" collectorName="T. Robillard &amp; M. K. Tan" country="Malaysia" county="Sandakan District" elevation="54" latitude="5.87395" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" pageId="8" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
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<typeStatus id="86482BF8FFD73E0B0D96FA85FDF503BF" box="[404,517,1403,1429]" pageId="8" pageNumber="213" type="holotype">Holotype</typeStatus>
:
</emphasis>
EAST
<collectingCountry id="21E4D5CAFFD73E0B0E69FA85FCF103BF" box="[619,769,1403,1429]" name="Malaysia" pageId="8" pageNumber="213">MALAYSIA</collectingCountry>
:
<specimenCount id="4FF55ED3FFD73E0B0F10FA85FCD803BF" box="[786,808,1403,1429]" count="1" pageId="8" pageNumber="213" type="female"></specimenCount>
;
<collectingRegion id="9B375BB8FFD73E0B0F3EFA85FC3903BF" box="[828,969,1403,1429]" country="Malaysia" name="Sabah" pageId="8" pageNumber="213">Sabah State</collectingRegion>
,
<collectingCounty id="B02DEDD6FFD73E0B0FD9FA85FB5C03BF" box="[987,1196,1403,1429]" pageId="8" pageNumber="213">Sandakan District</collectingCounty>
,
<collectingMunicipality id="B9280F20FFD73E0B08BCFA85FAE603BF" box="[1214,1302,1403,1429]" pageId="8" pageNumber="213">Sepilok</collectingMunicipality>
,
<location id="5C2CC381FFD73E0B092AFA85FE920393" LSID="urn:lsid:plazi:treatment:D15A244CFFD73E080C95FAF0FBDF01F4:5C2CC381FFD73E0B092AFA85FE920393" country="Malaysia" county="Sandakan District" latitude="5.87395" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" name="Rainforest Discovery Centre" pageId="8" pageNumber="213" stateProvince="Sabah">Rainforest Discovery Centre</location>
, dipterocarp forest;
<geoCoordinate id="3CC7F39DFFD73E0B0E4BFA61FD390390" box="[585,713,1439,1466]" degrees="05.87395" direction="north" orientation="latitude" pageId="8" pageNumber="213" precision="1" value="5.87395">N05.87395</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFD73E0B0ED4FA61FC900393" box="[726,864,1439,1465]" degrees="117.93871" direction="east" orientation="longitude" pageId="8" pageNumber="213" precision="1" value="117.93871">E117.93871</geoCoordinate>
,
<quantity id="9E0B38BFFFD73E0B0F6CFA61FC5C0393" box="[878,940,1439,1466]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="213" unit="m" value="54.0">
<elevation id="D2DE7269FFD73E0B0F6CFA61FC5C0393" box="[878,940,1439,1466]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="213" unit="m" value="54.0">54 m</elevation>
</quantity>
;
<date id="2D4DB39AFFD73E0B0FB8FA61FBDB0393" box="[954,1067,1439,1465]" pageId="8" pageNumber="213" value="2022-05-18">
<collectingDate id="3D094A72FFD73E0B0FB8FA61FBDB0393" box="[954,1067,1439,1465]" pageId="8" pageNumber="213" value="2022-05-18">18.v.2022</collectingDate>
</date>
, night; on tree trunk (h=
<quantity id="9E0B38BFFFD73E0B094EFA61FA800393" box="[1356,1392,1439,1465]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="213" unit="m" value="1.0">
<elevation id="D2DE7269FFD73E0B094EFA61FA800393" box="[1356,1392,1439,1465]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="213" unit="m" value="1.0">1m</elevation>
</quantity>
);
<collectorName id="F406F08CFFD73E0B0984FA61FEF103F7" pageId="8" pageNumber="213">T. Robillard</collectorName>
&amp;
<collectorName id="F406F08CFFD73E0B0D27FA3DFE6003F7" box="[293,400,1475,1501]" pageId="8" pageNumber="213">M.K. Tan</collectorName>
leg.; TR22-30 (
<collectionCode id="3FE20D9FFFD73E0B0E41FA3DFD8E03F7" box="[579,638,1475,1501]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="8" pageNumber="213">FRC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="594C955AFFD73E0B0CC5FA19FE090047" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="213">
<materialsCitation id="E99B9F07FFD73E0B0CC5FA19FE050047" collectingDate="2022-05-18" collectorName="T. Robillard &amp; M. K. Tan &amp; Pap" country="Malaysia" county="Sandakan District" elevation="54" latitude="5.87395" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" pageId="8" pageNumber="213" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="6B874948FFD73E0B0CC5FA19FECF002B" bold="true" box="[199,319,1511,1537]" pageId="8" pageNumber="213">
<typeStatus id="86482BF8FFD73E0B0CC5FA19FECF002B" box="[199,319,1511,1537]" pageId="8" pageNumber="213" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="4FF55ED3FFD73E0B0D4DFA19FE84002B" box="[335,372,1511,1537]" count="1" pageId="8" pageNumber="213" type="male">1♂</specimenCount>
,
<specimenCount id="4FF55ED3FFD73E0B0D7DFA19FE50002B" box="[383,416,1511,1537]" count="1" pageId="8" pageNumber="213" type="female">1♀</specimenCount>
): EAST
<collectingCountry id="21E4D5CAFFD73E0B0E05FA19FD6D002B" box="[519,669,1511,1537]" name="Malaysia" pageId="8" pageNumber="213">MALAYSIA</collectingCountry>
: •
<specimenCount id="4FF55ED3FFD73E0B0EB4FA19FD2B002B" box="[694,731,1511,1537]" count="1" pageId="8" pageNumber="213" type="male">1♂</specimenCount>
;
<collectingRegion id="9B375BB8FFD73E0B0EE5FA19FC9D002B" box="[743,877,1511,1537]" country="Malaysia" name="Sabah" pageId="8" pageNumber="213">Sabah State</collectingRegion>
,
<collectingCounty id="B02DEDD6FFD73E0B0F75FA19FBB1002B" box="[887,1089,1511,1537]" pageId="8" pageNumber="213">Sandakan District</collectingCounty>
,
<collectingMunicipality id="B9280F20FFD73E0B0849FA19FB53002B" box="[1099,1187,1511,1537]" pageId="8" pageNumber="213">Sepilok</collectingMunicipality>
,
<location id="5C2CC381FFD73E0B08ACFA19FF15000F" LSID="urn:lsid:plazi:treatment:D15A244CFFD73E080C95FAF0FBDF01F4:5C2CC381FFD73E0B08ACFA19FF15000F" country="Malaysia" county="Sandakan District" latitude="5.87395" longLatPrecision="1" longitude="117.93871" municipality="Sepilok" name="Rainforest Discovery Centre" pageId="8" pageNumber="213" stateProvince="Sabah">Rainforest Discovery Centre</location>
, dipterocarp forest;
<geoCoordinate id="3CC7F39DFFD73E0B0DCAF9F5FDB8000C" box="[456,584,1547,1574]" degrees="05.87395" direction="north" orientation="latitude" pageId="8" pageNumber="213" precision="1" value="5.87395">N05.87395</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFD73E0B0E56F9F5FD2D000F" box="[596,733,1547,1573]" degrees="117.93871" direction="east" orientation="longitude" pageId="8" pageNumber="213" precision="1" value="117.93871">E117.93871</geoCoordinate>
,
<quantity id="9E0B38BFFFD73E0B0EE8F9F5FCD7000F" box="[746,807,1547,1574]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="213" unit="m" value="54.0">
<elevation id="D2DE7269FFD73E0B0EE8F9F5FCD7000F" box="[746,807,1547,1574]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="213" unit="m" value="54.0">54 m</elevation>
</quantity>
;
<date id="2D4DB39AFFD73E0B0F36F9F5FC54000F" box="[820,932,1547,1573]" pageId="8" pageNumber="213" value="2022-05-18">
<collectingDate id="3D094A72FFD73E0B0F36F9F5FC54000F" box="[820,932,1547,1573]" pageId="8" pageNumber="213" value="2022-05-18">18.v.2022</collectingDate>
</date>
, night; on tree trunk (h=
<quantity id="9E0B38BFFFD73E0B08BCF9F5FB11000F" box="[1214,1249,1547,1573]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="213" unit="m" value="1.0">
<elevation id="D2DE7269FFD73E0B08BCF9F5FB11000F" box="[1214,1249,1547,1573]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="213" unit="m" value="1.0">1m</elevation>
</quantity>
);
<collectorName id="F406F08CFFD73E0B08F5F9F5FA8F000F" box="[1271,1407,1547,1573]" pageId="8" pageNumber="213">T. Robillard</collectorName>
&amp;
<collectorName id="F406F08CFFD73E0B0C95F9D1FEF30063" box="[151,259,1583,1609]" pageId="8" pageNumber="213">M.K. Tan</collectorName>
leg.; call recording: video files 6566, sound files
<collectorName id="F406F08CFFD73E0B0F40F9D1FC9E0063" box="[834,878,1583,1609]" pageId="8" pageNumber="213">Pap</collectorName>
268271, TR22-30 (MNHN-EO1277) •
<specimenCount id="4FF55ED3FFD73E0B0932F9D1FAA50063" box="[1328,1365,1583,1609]" count="1" pageId="8" pageNumber="213" type="female">1♀</specimenCount>
; same information (MNHN-EO1278)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD73E0B0CC5F989FD2B016F" pageId="8" pageNumber="213" type="diagnosis">
<paragraph id="594C955AFFD73E0B0CC5F989FD2B016F" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="213">
<emphasis id="6B874948FFD73E0B0CC5F989FEB200BB" bold="true" box="[199,322,1655,1681]" pageId="8" pageNumber="213">Diagnosis.</emphasis>
This species differs from all congeners by the male genitalia with its pseudepiphallic lophi [posterolateral epiphallic lobes] straight (bent in
<taxonomicName id="9EF3EED9FFD73E0B0EA5F962FCF0009F" authorityName="Gorochov" authorityYear="2004" box="[679,768,1691,1717]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="6B874948FFD73E0B0EA5F962FCF0009F" box="[679,768,1691,1717]" italics="true" pageId="8" pageNumber="213">T. plana</emphasis>
</taxonomicName>
and
<taxonomicName id="9EF3EED9FFD73E0B0F32F962FC30009F" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[816,960,1691,1717]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="species" species="belaitensis">
<emphasis id="6B874948FFD73E0B0F32F962FC30009F" box="[816,960,1691,1717]" italics="true" pageId="8" pageNumber="213">T.belaitensis</emphasis>
</taxonomicName>
) or narrow (narrower than in
<taxonomicName id="9EF3EED9FFD73E0B08FEF962FA7B009F" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[1276,1419,1691,1717]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="species" species="biaculeata">
<emphasis id="6B874948FFD73E0B08FEF962FA7B009F" box="[1276,1419,1691,1717]" italics="true" pageId="8" pageNumber="213">T.biaculeata</emphasis>
</taxonomicName>
); sclerotized bridge connecting the two lateral parts of pseudepiphallus distinctly longer; pseudepiphallic parameres [ectoparameres] membraneous; ectophallic fold [rachis] with two long and slender anterolateral ribbons; endophallic sclerite [formula] nearly divided into two compact lateral parts (each part somewhat U-shaped); additional plates rather long and narrow as well as obliquely located.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD73E0B0CC5F8B1FD4F01A7" pageId="8" pageNumber="213" type="etymology">
<paragraph id="594C955AFFD73E0B0CC5F8B1FD4F01A7" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="213">
<emphasis id="6B874948FFD73E0B0CC5F8B1FEBE0143" bold="true" box="[199,334,1871,1897]" pageId="8" pageNumber="213">Etymology.</emphasis>
The species name refers to the Kabili Monster tree, an Obah Suluk tree (
<taxonomicName id="9EF3EED9FFD73E0B0875F8B1FAB10143" box="[1143,1345,1871,1897]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Shorea" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="213" phylum="Tracheophyta" rank="species" species="pauciflora">
<emphasis id="6B874948FFD73E0B0875F8B1FAB10143" box="[1143,1345,1871,1897]" italics="true" pageId="8" pageNumber="213">Shorea pauciflora</emphasis>
</taxonomicName>
) near to where the species was found; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD73E080CC5F869FADC008A" lastPageId="11" lastPageNumber="216" pageId="8" pageNumber="213" type="description">
<paragraph id="594C955AFFD73E0B0CC5F869FA8D01D3" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="213">
<emphasis id="6B874948FFD73E0B0CC5F869FEA9019B" bold="true" box="[199,345,1943,1969]" pageId="8" pageNumber="213">Description.</emphasis>
General appearance, including tegminal structure, very similar to
<taxonomicName id="9EF3EED9FFD73E0B0851F869FA81019B" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[1107,1393,1943,1969]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD73E0B0851F869FA81019B" box="[1107,1393,1943,1969]" italics="true" pageId="8" pageNumber="213">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD73E0B0979F869FF3801FF" bold="true" pageId="8" pageNumber="213">
<taxonomicNameLabel id="70B4F433FFD73E0B0979F869FF3801FF" pageId="8" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but with following differences (
<figureCitation id="C1C889DFFFD73E0B0E31F845FD8D01FF" box="[563,637,1979,2005]" captionStart="FIGURE 9" captionStartId="12.[151,250,1350,1374]" captionTargetBox="[176,1425,193,1326]" captionTargetId="figure-20@12.[151,1436,181,1326]" captionTargetPageId="12" captionText="FIGURE 9. Habitus of Tembelingiola kabili sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="8" pageNumber="213">
Figs
<geoCoordinate id="3CC7F39DFFD73E0B0E6FF845FD8D01FF" box="[621,637,1979,2005]" degrees="9, 7" direction="east" orientation="longitude" pageId="8" pageNumber="213" precision="5555" value="9.7">9</geoCoordinate>
</figureCitation>
,
<figureCitation id="C1C889DFFFD73E0B0E8BF845FD2001FF" box="[649,720,1979,2005]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="8" pageNumber="213">7EG</figureCitation>
): rostrum between antennal cavities wider, 1.3 times wider than scapes (
<figureCitation id="C1C889DFFFD73E0B0CF2F821FEB701D3" box="[240,327,2015,2041]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="8" pageNumber="213">Fig. 7E</figureCitation>
); maxillary palp with apical segment slightly more elongate; pronotum less pubescent (
<figureCitation id="C1C889DFFFD73E0B091BF821FA8001D3" box="[1305,1392,2015,2041]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="8" pageNumber="213">Fig. 7E</figureCitation>
).
</paragraph>
<caption id="0D8CC5D2FFD63E0A0C95FB6DFAB50280" box="[151,1349,1171,1195]" pageId="9" pageNumber="214" startId="9.[151,250,1171,1195]" targetBox="[151,1423,195,1147]" targetPageId="9" targetType="figure">
<paragraph id="594C955AFFD63E0A0C95FB6DFAB50280" blockId="9.[151,1349,1171,1195]" box="[151,1349,1171,1195]" pageId="9" pageNumber="214">
<emphasis id="6B874948FFD63E0A0C95FB6DFEE30281" bold="true" box="[151,275,1171,1195]" pageId="9" pageNumber="214">FIGURE 6.</emphasis>
Habitus of
<taxonomicName id="9EF3EED9FFD63E0A0D88FB6DFD7B0281" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[394,651,1171,1195]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="214" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD63E0A0D88FB6DFD7B0281" box="[394,651,1171,1195]" italics="true" pageId="9" pageNumber="214">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD63E0A0E90FB6DFD120281" bold="true" box="[658,738,1171,1195]" pageId="9" pageNumber="214">
<taxonomicNameLabel id="70B4F433FFD63E0A0E90FB6DFD120281" box="[658,738,1171,1195]" pageId="9" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male in dorsal (A) and lateral (B) views. Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="594C955AFFD63E0A0CC5FB11FE4500D0" blockId="9.[151,1437,1263,2035]" pageId="9" pageNumber="214">
<emphasis id="6B874948FFD63E0A0CC5FB11FEFB0323" bold="true" box="[199,267,1263,1289]" pageId="9" pageNumber="214">Male.</emphasis>
FW brown, reaching abdominal apex, 2.2 times as long as wide (
<figureCitation id="C1C889DFFFD63E0A0FE1FB11FBCD0323" box="[995,1085,1263,1289]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="9" pageNumber="214">Fig. 7H</figureCitation>
). Dorsal field more slender, 2.2 times as long as wide, with only 4 harp veins, and with mirror longer than wide; with a faint diagonal vein; harps with 4 oblique veins including 2 long, parallel sinusoidal posterior veins fused anteriorly, and 2 anterior shorter and fainter one; mirror approximately 1.1 times longer than wide, separated by two dividing veins, dividing veins mostly straight but slightly bent at anal end.Apical field moderately short (
<figureCitation id="C1C889DFFFD63E0A0F7AFA83FC2003BD" box="[888,976,1405,1431]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="9" pageNumber="214">Fig. 7H</figureCitation>
). Lateral field with about 7 branches of Sc and 9 cross veins between R and M. Hind wings not exposed (
<figureCitation id="C1C889DFFFD63E0A0F43FA5FFC740391" box="[833,900,1441,1467]" captionStart="FIGURE 9" captionStartId="12.[151,250,1350,1374]" captionTargetBox="[176,1425,193,1326]" captionTargetId="figure-20@12.[151,1436,181,1326]" captionTargetPageId="12" captionText="FIGURE 9. Habitus of Tembelingiola kabili sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="9" pageNumber="214">Fig. 9</figureCitation>
). Genitalia as shown in
<figureCitation id="C1C889DFFFD63E0A088EFA5FFB2D0391" box="[1164,1245,1441,1467]" captionStart="FIGURE 10" captionStartId="13.[151,250,1588,1612]" captionTargetBox="[151,1436,345,1564]" captionTargetId="figure-44@13.[151,1436,345,1564]" captionTargetPageId="13" captionText="FIGURE 10. Male genitalia of Tembelingiola kabili sp. nov. in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm." pageId="9" pageNumber="214">Fig. 10</figureCitation>
. Pseudepiphallus [epiphallus] consisting of two lateral parts connected with each other by broad bridge; pseudepiphallic lophi [posterolateral epiphallic lobes] dorso-ventrally compressed, straight, diverging slightly posteriorly, and with apices roundly truncated; between these lophi, pseudepiphallus posteriorly angularly concave. Pseudepiphallic parameres [ectoparameres] membraneous. Ectophallic fold [rachis] semi-tube-like, narrowing posteriorly into acute apex, slightly surpassing pseudepiphallic bridge, and with a pair of long and slender anterolateral sclerotized ribbons. Endophallic sclerite [formula] lamellate and somewhat transverse but compact, almost divided into two U-shaped lateral parts (each part with medial half more sclerotized and lateral half almost semi-sclerotized. Rami weakly sclerotized, fused with epiphallus. Additional lateral sclerotized plates (p) very long and slender, obliquely located, slightly curved in profile.
</paragraph>
<paragraph id="594C955AFFD63E0A0CC5F8FDFDD701D8" blockId="9.[151,1437,1263,2035]" pageId="9" pageNumber="214">
<emphasis id="6B874948FFD63E0A0CC5F8FDFE660137" bold="true" box="[199,406,1795,1821]" pageId="9" pageNumber="214">
Female (
<figureCitation id="C1C889DFFFD63E0A0D33F8FDFE770137" box="[305,391,1795,1821]" captionStart="FIGURE 11" captionStartId="14.[151,250,1166,1190]" captionTargetBox="[151,1436,181,1143]" captionTargetId="figure-20@14.[151,1436,181,1143]" captionTargetPageId="14" captionText="FIGURE 11. Habitus of Tembelingiola kabili sp. nov., female in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="9" pageNumber="214">Fig. 11</figureCitation>
).
</emphasis>
Colouration and structure of body similar to male. Tegminal dorsal field with 4 oblique longitudinal branches and with numerous and slightly irregular cross-veins (some cross-veins rather long and very oblique, i.e., situated almost longitudinally); most-anal branch after middle splits into two sub-branches (
<figureCitation id="C1C889DFFFD63E0A093DF8B4FA7F014E" box="[1343,1423,1866,1892]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="9" pageNumber="214">Fig. 7I</figureCitation>
). Lateral field with 3 branches of Sc and with cross-veins between R and M indistinct. Subgenital plate wide but gradually narrowing backwards, with almost widely truncate apex having shallow and rather wide postero-median notch. Ovipositor dark rufous, weakly arcuate in profile, with middle part rather high (wide) and distal third gradually narrowing to acute apex (
<figureCitation id="C1C889DFFFD63E0A0DB6F826FDEA01D8" box="[436,538,2008,2034]" captionStart="FIGURE 11" captionStartId="14.[151,250,1166,1190]" captionTargetBox="[151,1436,181,1143]" captionTargetId="figure-20@14.[151,1436,181,1143]" captionTargetPageId="14" captionText="FIGURE 11. Habitus of Tembelingiola kabili sp. nov., female in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="9" pageNumber="214">Fig. 11B</figureCitation>
).
</paragraph>
<caption id="0D8CC5D2FFD53E090C95F869FE0101D2" pageId="10" pageNumber="215" startId="10.[151,250,1943,1967]" targetBox="[154,1419,181,1900]" targetPageId="10" targetType="figure">
<paragraph id="594C955AFFD53E090C95F869FE0101D2" blockId="10.[151,1436,1943,2040]" pageId="10" pageNumber="215">
<emphasis id="6B874948FFD53E090C95F869FEE2019A" bold="true" box="[151,274,1943,1968]" pageId="10" pageNumber="215">FIGURE 7.</emphasis>
<taxonomicName id="9EF3EED9FFD53E090D15F869FDE70185" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[279,535,1943,1967]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="215" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD53E090D15F869FDE70185" box="[279,535,1943,1967]" italics="true" pageId="10" pageNumber="215">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD53E090E1FF866FD9C019A" bold="true" box="[541,620,1944,1968]" pageId="10" pageNumber="215">
<taxonomicNameLabel id="70B4F433FFD53E090E1FF866FD9C019A" box="[541,620,1944,1968]" pageId="10" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(AD) and
<taxonomicName id="9EF3EED9FFD53E090EE1F869FC420185" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[739,946,1943,1967]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="215" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD53E090EE1F869FC420185" box="[739,946,1943,1967]" italics="true" pageId="10" pageNumber="215">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD53E090FB5F866FBF7019A" bold="true" box="[951,1031,1944,1968]" pageId="10" pageNumber="215">
<taxonomicNameLabel id="70B4F433FFD53E090FB5F866FBF7019A" box="[951,1031,1944,1968]" pageId="10" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest).
</paragraph>
</caption>
<caption id="0D8CC5D2FFD43E080C95FA72FF3603E2" pageId="11" pageNumber="216" startId="11.[151,250,1420,1444]" targetBox="[174,1404,193,1378]" targetPageId="11" targetType="figure">
<paragraph id="594C955AFFD43E080C95FA72FF3603E2" blockId="11.[151,1437,1420,1480]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080C95FA72FEE3038F" bold="true" box="[151,275,1420,1445]" pageId="11" pageNumber="216">FIGURE 8.</emphasis>
Male genitalia of
<taxonomicName id="9EF3EED9FFD43E080DCFFA73FD3F038F" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[461,719,1421,1445]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="216" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD43E080DCFFA73FD3F038F" box="[461,719,1421,1445]" italics="true" pageId="11" pageNumber="216">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD43E080ED7FA73FCD6038F" bold="true" box="[725,806,1421,1445]" pageId="11" pageNumber="216">
<taxonomicNameLabel id="70B4F433FFD43E080ED7FA73FCD6038F" box="[725,806,1421,1445]" pageId="11" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="594C955AFFD43E080CC5F9E5FADC008A" blockId="11.[151,1437,1562,2015]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080CC5F9E5FE8E001F" bold="true" box="[199,382,1563,1589]" pageId="11" pageNumber="216">Measurements.</emphasis>
<specimenCount id="4FF55ED3FFD43E080D96F9E5FE59001F" box="[404,425,1563,1589]" count="1" pageId="11" pageNumber="216" type="male"></specimenCount>
<typeStatus id="86482BF8FFD43E080DADF9E4FDE5001E" box="[431,533,1562,1588]" pageId="11" pageNumber="216" type="holotype">holotype</typeStatus>
: BL = 8.0; BWL = 9.3; HL = 1.2; PronL = 2.2; PronW = 2.5; FWL = 6.1; FWW = 2.8; FIIIL = 5.8; TIIIL = 3.6; TaIIIL = 2.8 •
<specimenCount id="4FF55ED3FFD43E080E78F9C0FD7F0072" box="[634,655,1598,1624]" count="1" pageId="11" pageNumber="216" type="female"></specimenCount>
<typeStatus id="86482BF8FFD43E080E94F9C0FD0B0072" box="[662,763,1598,1624]" pageId="11" pageNumber="216" type="paratype">paratype</typeStatus>
: BL = 7.9; BWL = 9.5; HL = 1.0; PronL = 2.4; PronW = 2.6; FWL = 6.4; FWW = 1.8; FIIIL = 6.9; TIIIL = 3.9; TaIIIL = 3.0; OL = 5.4 •
<specimenCount id="4FF55ED3FFD43E080FD8F99CFC1F0056" box="[986,1007,1634,1660]" count="1" pageId="11" pageNumber="216" type="female"></specimenCount>
<typeStatus id="86482BF8FFD43E080FF7F99CFBAA0056" box="[1013,1114,1634,1660]" pageId="11" pageNumber="216" type="paratype">paratype</typeStatus>
: BL = 7.3; BWL = 9.5; HL = 1.1; PronL = 2.4; PronW = 2.7; FWL = 6.4; FWW = 1.7; FIIIL = 6.2; TIIIL = 3.5; TaIIIL = 2.9; OL = 5.2
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD43E080CC5F956FA6800E8" box="[199,1432,1704,1731]" pageId="11" pageNumber="216" type="biology_ecology">
<paragraph id="594C955AFFD43E080CC5F956FA6800E8" blockId="11.[151,1437,1562,2015]" box="[199,1432,1704,1731]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080CC5F956FEDC00E8" bold="true" box="[199,300,1704,1730]" pageId="11" pageNumber="216">Ecology.</emphasis>
A male was observed producing short echemes behind
<specimenCount id="4FF55ED3FFD43E080F98F957FBD200E8" box="[922,1058,1704,1731]" count="2" pageId="11" pageNumber="216" type="female">two females</specimenCount>
on a tree trunk at night (
<figureCitation id="C1C889DFFFD43E080935F956FA7A00E8" box="[1335,1418,1704,1730]" captionStart="FIGURE 12" captionStartId="15.[151,250,1880,1904]" captionTargetBox="[151,1436,181,1857]" captionTargetId="figure-17@15.[151,1436,181,1857]" captionTargetPageId="15" captionText="FIGURE 12. Tembelingiola kabili sp. nov.: a male with two females on a tree trunk in Sepilok (A); the male started calling (B)." pageId="11" pageNumber="216">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD43E080CC5F935FBDF01F4" pageId="11" pageNumber="216" type="distribution">
<paragraph id="594C955AFFD43E080CC5F935FD0D00CC" blockId="11.[151,1437,1562,2015]" box="[199,765,1739,1766]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080CC5F935FE9100CF" bold="true" box="[199,353,1739,1765]" pageId="11" pageNumber="216">Distribution.</emphasis>
EAST
<collectingCountry id="21E4D5CAFFD43E080DB6F932FDBA00CC" box="[436,586,1740,1766]" name="Malaysia" pageId="11" pageNumber="216">MALAYSIA</collectingCountry>
:
<collectingRegion id="9B375BB8FFD43E080E56F932FD6D00CC" box="[596,669,1740,1766]" country="Malaysia" name="Sabah" pageId="11" pageNumber="216">Sabah</collectingRegion>
: Sepilok
</paragraph>
<paragraph id="594C955AFFD43E080CC5F911FBDF01F4" blockId="11.[151,1437,1562,2015]" pageId="11" pageNumber="216">
<emphasis id="6B874948FFD43E080CC5F911FDC20123" bold="true" box="[199,562,1775,1801]" pageId="11" pageNumber="216">
Calling song (
<figureCitation id="C1C889DFFFD43E080D68F911FE350123" box="[362,453,1775,1801]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="11" pageNumber="216">Figs 2B</figureCitation>
,
<figureCitation id="C1C889DFFFD43E080DD0F911FE040123" box="[466,500,1775,1801]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="11" pageNumber="216">3B</figureCitation>
,
<figureCitation id="C1C889DFFFD43E080E02F911FDD30123" box="[512,547,1775,1801]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="11" pageNumber="216">4B</figureCitation>
).
</emphasis>
Consists of echeme-sequence, each made up of 6 to 8 echemes. At 26°C, each echeme-sequence has an average duration of 0.84±
<geoCoordinate id="3CC7F39DFFD43E080EE3F8EDFCD70107" box="[737,807,1811,1837]" degrees="0.08" direction="south" orientation="latitude" pageId="11" pageNumber="216" precision="555" value="-0.08">0.08 s</geoCoordinate>
(0.67
<geoCoordinate id="3CC7F39DFFD43E080F7AF8EDFC4F0107" box="[888,959,1811,1837]" degrees="0.99" direction="south" orientation="latitude" pageId="11" pageNumber="216" precision="555" value="-0.99">0.99 s</geoCoordinate>
). The silent interval between consecutive echeme-sequences is 11.9±
<geoCoordinate id="3CC7F39DFFD43E080DC8F8C8FDF0017B" box="[458,512,1846,1873]" degrees="5.7" direction="south" orientation="latitude" pageId="11" pageNumber="216" precision="5555" value="-5.7">5.7 s</geoCoordinate>
(7.4
<geoCoordinate id="3CC7F39DFFD43E080E40F8C8FD78017A" box="[578,648,1846,1872]" degrees="29.7" direction="south" orientation="latitude" pageId="11" pageNumber="216" precision="5555" value="-29.7">29.7 s</geoCoordinate>
). Each echeme consists of 3 syllables and has an average duration of 56.2±1.4 ms (53.659 ms). The silent interval between consecutive echemes is 66.4±3.5 ms (60.572.1 ms). Within each echeme, the average syllable duration is 11.5±0.5 ms (10.512.2 ms) and the average silent interval between consecutive syllables is 9.5±0.5 ms (8.610.2 ms). The frequency spectrum is pure-tonal and forms a clear harmonic series, with the energy peaking at a dominant fundamental frequency of 5.91 kHz.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,432 @@
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<mods:title id="A41F73DB94E14341007BF6E7C25234ED">New species of little-known crickets from the subfamily Pteroplistinae (Orthoptera, Grylloidea) from Sabah and contribution to their bioacoustics</mods:title>
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<mods:namePart id="EEDC8745D43B3EEA7DC23D71CA4FEC23">Tan, Ming Kai</mods:namePart>
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<mods:affiliation id="2795E6CA38B67655C595514CE1BBF677">Zoological Institute, Russian Academy of Sciences, 1 Universitetskaya Emb., St Petersburg 199034, Russia</mods:affiliation>
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<mods:namePart id="904F84725281D586E02B3A91B5F1DA0C">Japir, Razy</mods:namePart>
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<mods:affiliation id="A62EA24BAD0382B0085CB8BBF1134118">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore &amp; Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France &amp; Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<mods:number id="EA43D4C0CCA21B15EE28D9F2B305195B">2</mods:number>
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<mods:start id="2C7692E930DD465BB975F83013430AA7">205</mods:start>
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<mods:location id="0E6969D86A8703B91493B4D32611E9B5">
<mods:url id="7EC3BCF9C3EE3A3B65B491BDC0F5AABF">http://dx.doi.org/10.11646/zootaxa.5506.2.3</mods:url>
</mods:location>
<mods:classification id="49464D216AFD9D2E5A3C28DF2E7B29AE">journal article</mods:classification>
<mods:identifier id="6B217FCFC9FA23E71641615649D43187" type="DOI">10.11646/zootaxa.5506.2.3</mods:identifier>
<mods:identifier id="DFE776E81441E21777BF9237B6F57382" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FCA203443FB761BF71249924E1151681" type="ZooBank">8D2FAB83-6C2B-424E-BDA3-9EB430B9EDDC</mods:identifier>
</mods:mods>
<treatment id="D15A244CFFDB3E0B0C95FE17FD6A0737" LSID="urn:lsid:plazi:treatment:D15A244CFFDB3E0B0C95FE17FD6A0737" httpUri="http://treatment.plazi.org/id/D15A244CFFDB3E0B0C95FE17FD6A0737" lastPageId="8" lastPageNumber="213" pageId="4" pageNumber="209">
<subSubSection id="11E9C6D1FFDB3E070C95FE17FC65042E" box="[151,917,489,516]" pageId="4" pageNumber="209" type="nomenclature">
<paragraph id="594C955AFFDB3E070C95FE17FC65042E" blockId="4.[151,917,489,553]" box="[151,917,489,516]" pageId="4" pageNumber="209">
<heading id="02042236FFDB3E070C95FE17FC65042E" bold="true" box="[151,917,489,516]" fontSize="11" level="1" pageId="4" pageNumber="209" reason="1">
<emphasis id="6B874948FFDB3E070C95FE17FC65042E" bold="true" box="[151,917,489,516]" pageId="4" pageNumber="209">
<taxonomicName id="9EF3EED9FFDB3E070C95FE17FCDA0429" authority="Tan, Gorochov &amp; Robillard" authorityName="Tan, Gorochov &amp; Robillard" authorityYear="2024" box="[151,810,489,516]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFDB3E070C95FE17FE37042E" bold="true" box="[151,455,489,516]" italics="true" pageId="4" pageNumber="209">Tembelingiola biaculeata</emphasis>
Tan, Gorochov &amp; Robillard
</taxonomicName>
,
<taxonomicNameLabel id="70B4F433FFDB3E070F35FE14FC65042E" box="[823,917,490,516]" pageId="4" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E070C95FDF0FDFA0402" box="[151,522,526,553]" pageId="4" pageNumber="209" type="description">
<paragraph id="594C955AFFDB3E070C95FDF0FDFA0402" blockId="4.[151,917,489,553]" box="[151,522,526,553]" pageId="4" pageNumber="209">
(
<figureCitation id="C1C889DFFFDB3E070C9DFDF0FF0C0402" box="[159,252,526,552]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="4" pageNumber="209">Figs 2A</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070D0AFDF0FEDC0402" box="[264,300,526,552]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="4" pageNumber="209">3A</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070D3AFDF0FEAD0402" box="[312,349,526,552]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="4" pageNumber="209">4A</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070D6AFDF0FE880403" box="[360,376,526,553]" captionStart="FIGURE 5" captionStartId="8.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-78@8.[151,1435,309,1165]" captionTargetPageId="8" captionText="FIGURE 5. Male Tembelingiola biaculeata sp. nov. in its natural habitat in Sepilok." pageId="4" pageNumber="209">5</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070D86FDF0FE640402" box="[388,404,526,552]" captionStart="FIGURE 6" captionStartId="9.[151,250,1171,1195]" captionTargetBox="[151,1423,195,1147]" captionTargetId="figure-17@9.[151,1436,178,1147]" captionTargetPageId="9" captionText="FIGURE 6. Habitus of Tembelingiola biaculeata sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="4" pageNumber="209">6</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070DA2FDF0FE180402" box="[416,488,526,552]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">7AD</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070DF1FDF0FDF10402" box="[499,513,526,552]" captionStart="FIGURE 8" captionStartId="11.[151,250,1420,1444]" captionTargetBox="[174,1404,193,1378]" captionTargetId="figure-17@11.[151,1436,181,1397]" captionTargetPageId="11" captionText="FIGURE 8. Male genitalia of Tembelingiola biaculeata sp. nov. in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm." pageId="4" pageNumber="209">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E070C95FDABFD7C050E" pageId="4" pageNumber="209" type="materials_examined">
<paragraph id="594C955AFFDB3E070C95FDABFC560492" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070C95FDABFE750445" bold="true" box="[151,389,597,623]" pageId="4" pageNumber="209">Material examined.</emphasis>
<materialsCitation id="E99B9F07FFDB3E070D93FDA8FC560492" collectingDate="2022-05-18" collectionCode="FRC" collectorName="M. K. Tan &amp; T. Robillard" country="Malaysia" county="Sandakan District" latitude="5.87552" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.94131" municipality="Sepilok" pageId="4" pageNumber="209" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="6B874948FFDB3E070D93FDA8FDF7045A" bold="true" box="[401,519,598,624]" pageId="4" pageNumber="209">
<typeStatus id="86482BF8FFDB3E070D93FDA8FDF2045A" box="[401,514,598,624]" pageId="4" pageNumber="209" type="holotype">Holotype</typeStatus>
:
</emphasis>
EAST
<collectingCountry id="21E4D5CAFFDB3E070E66FDA8FD0A045A" box="[612,762,598,624]" name="Malaysia" pageId="4" pageNumber="209">MALAYSIA</collectingCountry>
: •
<specimenCount id="4FF55ED3FFDB3E070F1FFDA8FCC5045A" box="[797,821,598,624]" count="1" pageId="4" pageNumber="209" type="male"></specimenCount>
;
<collectingRegion id="9B375BB8FFDB3E070F44FDA8FC21045A" box="[838,977,598,624]" country="Malaysia" name="Sabah" pageId="4" pageNumber="209">Sabah State</collectingRegion>
,
<collectingCounty id="B02DEDD6FFDB3E070FE2FDA8FB40045A" box="[992,1200,598,624]" pageId="4" pageNumber="209">Sandakan District</collectingCounty>
,
<collectingMunicipality id="B9280F20FFDB3E0708BDFDA8FAE7045A" box="[1215,1303,598,624]" pageId="4" pageNumber="209">Sepilok</collectingMunicipality>
,
<location id="5C2CC381FFDB3E07092AFDA8FE9204BE" LSID="urn:lsid:plazi:treatment:D15A244CFFDB3E0B0C95FE17FD6A0737:5C2CC381FFDB3E07092AFDA8FE9204BE" country="Malaysia" county="Sandakan District" latitude="5.87552" longLatPrecision="1" longitude="117.94131" municipality="Sepilok" name="Rainforest Discovery Centre" pageId="4" pageNumber="209" stateProvince="Sabah">Rainforest Discovery Centre</location>
, dipterocarp forest;
<geoCoordinate id="3CC7F39DFFDB3E070E48FD84FD4C04BF" box="[586,700,634,661]" degrees="5.87552" direction="north" orientation="latitude" pageId="4" pageNumber="209" precision="1" value="5.87552">N5.87552</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFDB3E070EC8FD84FCA304BE" box="[714,851,634,660]" degrees="117.94131" direction="east" orientation="longitude" pageId="4" pageNumber="209" precision="1" value="117.94131">E117.94131</geoCoordinate>
, 62.6±6.0 m.a.s.l.;
<date id="2D4DB39AFFDB3E070833FD84FB5204BE" box="[1073,1186,634,660]" pageId="4" pageNumber="209" value="2022-05-18">
<collectingDate id="3D094A72FFDB3E070833FD84FB5204BE" box="[1073,1186,634,660]" pageId="4" pageNumber="209" value="2022-05-18">18.v.2022</collectingDate>
</date>
, 22h36; on foliage of shrub near ground;
<collectorName id="F406F08CFFDB3E070D70FD60FE2D0492" box="[370,477,670,696]" pageId="4" pageNumber="209">M.K. Tan</collectorName>
&amp;
<collectorName id="F406F08CFFDB3E070E02FD60FD770492" box="[512,647,670,696]" pageId="4" pageNumber="209">T. Robillard</collectorName>
leg.; SBH.22.159 (
<collectionCode id="3FE20D9FFFDB3E070F62FD60FC6B0492" box="[864,923,670,696]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="4" pageNumber="209">FRC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="594C955AFFDB3E070CC5FD3CFD7C050E" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<materialsCitation id="E99B9F07FFDB3E070CC5FD3CFD78050E" collectingDate="2022-05-19" collectionCode="ZRC" collectorName="M. K. Tan &amp; T. Robillard" country="Malaysia" county="Sandakan District" elevation="7" latitude="5.87536" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.93977" municipality="Sepilok" pageId="4" pageNumber="209" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="6B874948FFDB3E070CC5FD3CFECD04F6" bold="true" box="[199,317,706,732]" pageId="4" pageNumber="209">
<typeStatus id="86482BF8FFDB3E070CC5FD3CFEC804F6" box="[199,312,706,732]" pageId="4" pageNumber="209" type="paratype">Paratype</typeStatus>
:
</emphasis>
EAST
<collectingCountry id="21E4D5CAFFDB3E070D95FD3CFDDD04F6" box="[407,557,706,732]" name="Malaysia" pageId="4" pageNumber="209">MALAYSIA</collectingCountry>
: •
<specimenCount id="4FF55ED3FFDB3E070E4CFD3CFD9604F6" box="[590,614,706,732]" count="1" pageId="4" pageNumber="209" type="male"></specimenCount>
;
<collectingRegion id="9B375BB8FFDB3E070E77FD3CFD0E04F6" box="[629,766,706,732]" country="Malaysia" name="Sabah" pageId="4" pageNumber="209">Sabah State</collectingRegion>
,
<collectingCounty id="B02DEDD6FFDB3E070F0EFD3CFC2A04F6" box="[780,986,706,732]" pageId="4" pageNumber="209">Sandakan District</collectingCounty>
,
<collectingMunicipality id="B9280F20FFDB3E070FEAFD3CFBB004F6" box="[1000,1088,706,732]" pageId="4" pageNumber="209">Sepilok</collectingMunicipality>
,
<location id="5C2CC381FFDB3E07084DFD3CFA6904F6" LSID="urn:lsid:plazi:treatment:D15A244CFFDB3E0B0C95FE17FD6A0737:5C2CC381FFDB3E07084DFD3CFA6904F6" box="[1103,1433,706,732]" country="Malaysia" county="Sandakan District" latitude="5.87536" longLatPrecision="1" longitude="117.93977" municipality="Sepilok" name="Rainforest Discovery Centre" pageId="4" pageNumber="209" stateProvince="Sabah">Rainforest Discovery Centre</location>
, dipterocarp forest;
<geoCoordinate id="3CC7F39DFFDB3E070D6DFD18FE11052B" box="[367,481,742,769]" degrees="5.87536" direction="north" orientation="latitude" pageId="4" pageNumber="209" precision="1" value="5.87536">N5.87536</geoCoordinate>
,
<geoCoordinate id="3CC7F39DFFDB3E070DEFFD18FD87052A" box="[493,631,742,768]" degrees="117.93977" direction="east" orientation="longitude" pageId="4" pageNumber="209" precision="1" value="117.93977">E117.93977</geoCoordinate>
, 57.6±
<quantity id="9E0B38BFFFDB3E070EC6FD18FCFC052A" box="[708,780,742,769]" metricMagnitude="0" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="209" unit="m" value="6.8">
<elevation id="D2DE7269FFDB3E070EC6FD18FCFC052A" box="[708,780,742,769]" metricMagnitude="0" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="209" unit="m" value="6.8">6.8 m</elevation>
</quantity>
.a.s.l.;
<date id="2D4DB39AFFDB3E070F52FD18FC31052A" box="[848,961,742,768]" pageId="4" pageNumber="209" value="2022-05-19">
<collectingDate id="3D094A72FFDB3E070F52FD18FC31052A" box="[848,961,742,768]" pageId="4" pageNumber="209" value="2022-05-19">19.v.2022</collectingDate>
</date>
, 21h06; among branches and twigs;
<collectorName id="F406F08CFFDB3E070963FD18FF31050E" pageId="4" pageNumber="209">M.K. Tan</collectorName>
&amp;
<collectorName id="F406F08CFFDB3E070CE6FCF4FE9B050E" box="[228,363,778,804]" pageId="4" pageNumber="209">T. Robillard</collectorName>
leg.; SBH.22.165 (
<collectionCode id="3FE20D9FFFDB3E070E41FCF4FD8F050E" box="[579,639,778,804]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="209">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E070CC5FCD3FBC802A6" pageId="4" pageNumber="209" type="diagnosis">
<paragraph id="594C955AFFDB3E070CC5FCD3FDEA059E" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070CC5FCD3FEB2056D" bold="true" box="[199,322,813,839]" pageId="4" pageNumber="209">Diagnosis.</emphasis>
This species differs from
<taxonomicName id="9EF3EED9FFDB3E070E71FCD0FBEB0562" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[627,1051,814,840]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="6B874948FFDB3E070E71FCD0FCAB0562" box="[627,859,814,840]" italics="true" pageId="4" pageNumber="209">Tembelingiola plana</emphasis>
<bibRefCitation id="3D62E8ABFFDB3E070F67FCD0FBEB0562" author="Gorochov, A. V." box="[869,1051,814,840]" pageId="4" pageNumber="209" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
from Peninsular
<collectingCountry id="21E4D5CAFFDB3E0708E6FCD0FABB0562" box="[1252,1355,814,840]" name="Malaysia" pageId="4" pageNumber="209">Malaysia</collectingCountry>
by the pseudepiphallic paramere [ectoparamere] thicker (less slender), directed downwards (but not backwards), with acute and hooked apical part; and the ectophallic fold [rachis] narrowing posteriorly into acute apex and slightly surpassing pseudepiphallic bridge.
</paragraph>
<paragraph id="594C955AFFDB3E070CC5FC40FBC802A6" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
It also differs from
<taxonomicName id="9EF3EED9FFDB3E070DB3FC40FBBD05F2" authority="Tan, Gorochov &amp; Wahab, 2019" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[433,1101,958,984]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="belaitensis">
<emphasis id="6B874948FFDB3E070DB3FC40FD2405F2" box="[433,724,958,984]" italics="true" pageId="4" pageNumber="209">Tembelingiola belaitensis</emphasis>
<bibRefCitation id="3D62E8ABFFDB3E070EDCFC40FBBD05F2" author="Tan, M. K. &amp; Gorochov, A. V. &amp; Wahab, R. A. &amp; Japir, R. &amp; Chung, A. Y. C." box="[734,1101,958,984]" pageId="4" pageNumber="209" pagination="101 - 117" refId="ref8234" refString="Tan, M. K., Gorochov, A. V., Wahab, R. A., Japir, R. &amp; Chung, A. Y. C. (2019) New taxa of crickets (Orthoptera: Grylloidea: Gryllinae, Phaloriinae and Pteroplistinae) from northern Borneo (Belait and Sandakan). Zootaxa, 4661 (1), 101 - 117. https: // doi. org / 10.11646 / zootaxa. 4661.1.4" type="journal article" year="2019">Tan, Gorochov &amp; Wahab, 2019</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="21E4D5CAFFDB3E07089BFC40FA8505F2" box="[1177,1397,958,984]" name="Brunei" pageId="4" pageNumber="209">Brunei Darussalam</collectingCountry>
by the pseudepiphallic lophi [posterolateral epiphallic lobes] directed partly externally (but not curved internally in distal part); pseudepiphallic parameres [ectoparameres] completely sclerotized and somewhat longer but directed downwards (in
<taxonomicName id="9EF3EED9FFDB3E070D4DFBD5FE18026E" authorityName="Tan, Gorochov &amp; Wahab" authorityYear="2019" box="[335,488,1066,1092]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="209" phylum="Arthropoda" rank="species" species="belaitensis">
<emphasis id="6B874948FFDB3E070D4DFBD5FE18026E" box="[335,488,1066,1092]" italics="true" pageId="4" pageNumber="209">T. belaitensis</emphasis>
</taxonomicName>
, they are in the form of membranous lobules directed backwards and with thin sclerotized ribbon ventrally); ectophallic fold [rachis] much shorter and less strongly sclerotized; and endophallic sclerite [formula] clearly less wide and not divided into a pair of sclerotized plates.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E070CC5FB68FB5102FE" pageId="4" pageNumber="209" type="etymology">
<paragraph id="594C955AFFDB3E070CC5FB68FB5102FE" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070CC5FB68FEBE029A" bold="true" box="[199,334,1174,1200]" pageId="4" pageNumber="209">Etymology.</emphasis>
The species name refers to the acute apices at the posterior end of both the pseudepiphallic parameres [ectoparameres]; bi- referring to two (a pair), and aculeatus = with spike in Latin.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDB3E040CC5FB23FC7D013F" lastPageId="7" lastPageNumber="212" pageId="4" pageNumber="209" type="description">
<paragraph id="594C955AFFDB3E070CC5FB23FA800082" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070CC5FB23FEA902DD" bold="true" box="[199,345,1245,1271]" pageId="4" pageNumber="209">Description.</emphasis>
Small-sized, body well pubescent, brown with head, pronotum and abdomen with rufous tinge (
<figureCitation id="C1C889DFFFDB3E070C9DFAFCFF180337" box="[159,232,1282,1309]" captionStart="FIGURE 5" captionStartId="8.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-78@8.[151,1435,309,1165]" captionTargetPageId="8" captionText="FIGURE 5. Male Tembelingiola biaculeata sp. nov. in its natural habitat in Sepilok." pageId="4" pageNumber="209">Figs 5</figureCitation>
,
<figureCitation id="C1C889DFFFDB3E070CF6FAFCFEF40336" box="[244,260,1282,1308]" captionStart="FIGURE 6" captionStartId="9.[151,250,1171,1195]" captionTargetBox="[151,1423,195,1147]" captionTargetId="figure-17@9.[151,1436,178,1147]" captionTargetPageId="9" captionText="FIGURE 6. Habitus of Tembelingiola biaculeata sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="4" pageNumber="209">6</figureCitation>
). Rostrum between antennal cavities 1.1 times wider than scapes (
<figureCitation id="C1C889DFFFDB3E070FF2FAFCFBBA0336" box="[1008,1098,1282,1308]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7A</figureCitation>
), angularly-rounded in profile (
<figureCitation id="C1C889DFFFDB3E070C9DFAD8FF08036A" box="[159,248,1318,1344]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7B</figureCitation>
). Maxillary palps pale coloured, with apical, subapical and third segments of subequal lengths; with apical segment elongated-triangular, very broadly expanded apically (
<figureCitation id="C1C889DFFFDB3E070F5EFAB4FC45034E" box="[860,949,1354,1380]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7B</figureCitation>
). Head wider than high in facial view. Face dark brown (
<figureCitation id="C1C889DFFFDB3E070D24FA90FE8E03A2" box="[294,382,1390,1416]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7C</figureCitation>
); median ocellus present, round; fenestrae between the scapes, anterior of median ocellus, also round and similar in size as median ocellus. Pronotal dorsal disc densely pubescent, weakly transverse, 1.11.3 times as wide as long, faintly widening posteriorly; anterior and posterior margins fairly straight to faintly concave and convex, respectively and with numerous long and thick setae (
<figureCitation id="C1C889DFFFDB3E070F82FA24FC2A03DE" box="[896,986,1498,1524]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7A</figureCitation>
). Lateral lobe 1.7 times as long as wide; posterior margin oblique, with numerous long and thick setae (more so than other parts of lateral lobe); ventral margin raised posteriorly; vaguely darker than dorsal disc (
<figureCitation id="C1C889DFFFDB3E070F35F9DCFC610016" box="[823,913,1570,1596]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7B</figureCitation>
). Inner tympanum on TI large, elongated oval (opened); outer tympanum undeveloped. TIII with only numerous small dorsal denticles; with 3 pairs of apical spurs, inner ones typically longer than outer ones; hind basitarsus with 2 rows of dorsal spines (each row with 56 spines). Legs generally pale brown. FIII pale brown, but distal part and knee dark rufous, and TIII dark rufous.
</paragraph>
<paragraph id="594C955AFFDB3E070CC5F94CFBBD01AA" blockId="4.[151,1437,597,1920]" pageId="4" pageNumber="209">
<emphasis id="6B874948FFDB3E070CC5F94CFEFB00E6" bold="true" box="[199,267,1714,1740]" pageId="4" pageNumber="209">Male.</emphasis>
FW light brown; barely reaching abdominal apex, 1.81.9 times as long as wide (
<figureCitation id="C1C889DFFFDB3E0708AAF94CFAF300E6" box="[1192,1283,1714,1740]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7D</figureCitation>
). Dorsal field with diagonal vein long and slightly sinuous; 5 oblique veins (3 long, parallel sinusoidal posterior ones and 2 anterior shorter and fainter ones); mirror approximately 1.1 times as wide as long, separated by two dividing veins, dividing veins mostly straight but at anal end slightly bent; with moderately short apical area (
<figureCitation id="C1C889DFFFDB3E0708A5F8E0FAF10112" box="[1191,1281,1822,1848]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[154,1419,181,1900]" captionTargetId="figure-20@10.[153,1434,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Tembelingiola biaculeata sp. nov. (AD) and Tembelingiola kabili sp. nov. (E I): head and pronotum in dorsal (A, E) and lateral (B, F) views, face in anterior view (C, G), male FW in dorsal view (D, H), female FWs in dorsal view (I). Scale bars: 2 mm (D, H, I), 1 mm (rest)." pageId="4" pageNumber="209">Fig. 7D</figureCitation>
). Lateral field of FW with about 9 branches of Sc and 12 cross veins between R and M. Hind wings not exposed (
<figureCitation id="C1C889DFFFDB3E0708E3F8BCFAD40176" box="[1249,1316,1858,1884]" captionStart="FIGURE 6" captionStartId="9.[151,250,1171,1195]" captionTargetBox="[151,1423,195,1147]" captionTargetId="figure-17@9.[151,1436,178,1147]" captionTargetPageId="9" captionText="FIGURE 6. Habitus of Tembelingiola biaculeata sp. nov., male in dorsal (A) and lateral (B) views. Scale bars: 2 mm." pageId="4" pageNumber="209">Fig. 6</figureCitation>
). Thoracic sternites brown. Subgenital plate with apex having an angular notched in the middle.
</paragraph>
<caption id="0D8CC5D2FFDA3E060C95F9BEFD0300EF" pageId="5" pageNumber="210" startId="5.[151,250,1600,1624]" targetBox="[161,1426,191,1567]" targetPageId="5" targetType="figure">
<paragraph id="594C955AFFDA3E060C95F9BEFD0300EF" blockId="5.[151,1436,1600,1733]" pageId="5" pageNumber="210">
<emphasis id="6B874948FFDA3E060C95F9BEFEE60072" bold="true" box="[151,278,1600,1624]" pageId="5" pageNumber="210">FIGURE 2.</emphasis>
Power spectra of the calling songs of the pteroplistine cricket species:
<taxonomicName id="9EF3EED9FFDA3E06080AF9BFFAFC0073" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[1032,1292,1601,1625]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFDA3E06080AF9BFFAFC0073" box="[1032,1292,1601,1625]" italics="true" pageId="5" pageNumber="210">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFDA3E060914F9BFFA990073" bold="true" box="[1302,1385,1601,1625]" pageId="5" pageNumber="210">
<taxonomicNameLabel id="70B4F433FFDA3E060914F9BFFA990073" box="[1302,1385,1601,1625]" pageId="5" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A),
<taxonomicName id="9EF3EED9FFDA3E060C95F99BFE980057" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[151,360,1637,1661]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFDA3E060C95F99BFE980057" box="[151,360,1637,1661]" italics="true" pageId="5" pageNumber="210">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFDA3E060D6DF99BFE310057" bold="true" box="[367,449,1637,1661]" pageId="5" pageNumber="210">
<taxonomicNameLabel id="70B4F433FFDA3E060D6DF99BFE310057" box="[367,449,1637,1661]" pageId="5" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B),
<taxonomicName id="9EF3EED9FFDA3E060DF5F99BFD5E0057" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[503,686,1637,1661]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFDA3E060DF5F99BFD5E0057" box="[503,686,1637,1661]" italics="true" pageId="5" pageNumber="210">Pteroplistes silam</emphasis>
</taxonomicName>
<emphasis id="6B874948FFDA3E060EB7F99BFCF60057" bold="true" box="[693,774,1637,1661]" pageId="5" pageNumber="210">
<taxonomicNameLabel id="70B4F433FFDA3E060EB7F99BFCF60057" box="[693,774,1637,1661]" pageId="5" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(C),
<taxonomicName id="9EF3EED9FFDA3E060F3FF99BFB4B0057" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[829,1211,1637,1661]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFDA3E060F3FF99BFBFF0057" box="[829,1039,1637,1661]" italics="true" pageId="5" pageNumber="210">Pteroplistes lagrecai</emphasis>
<bibRefCitation id="3D62E8ABFFDA3E060815F99BFB4B0057" author="Gorochov, A. V." box="[1047,1211,1637,1661]" pageId="5" pageNumber="210" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
(D) and
<taxonomicName id="9EF3EED9FFDA3E06091AF99BFE11008B" authority="Gorochov &amp; Tan, 2012" authorityName="Gorochov &amp; Tan" authorityYear="2012" class="Insecta" family="Pteroplistidae" genus="Singapuriola" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="separata">
<emphasis id="6B874948FFDA3E06091AF99BFF00008A" italics="true" pageId="5" pageNumber="210">Singapuriola separata</emphasis>
<bibRefCitation id="3D62E8ABFFDA3E060CF4F977FE11008B" author="Gorochov, A. V. &amp; Tan, M. K." box="[246,481,1673,1697]" pageId="5" pageNumber="210" pagination="18 - 34" refId="ref8057" refString="Gorochov, A. V. &amp; Tan, M. K. (2012) New crickets of the subfamilies Phaloriinae and Pteroplistinae (Orthoptera: Gryllidae) from Singapore. Zootaxa, 3525 (1), 18 - 34. https: // doi. org / 10.11646 / zootaxa. 3525.1.2" type="journal article" year="2012">Gorochov &amp; Tan, 2012</bibRefCitation>
</taxonomicName>
(E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black).
</paragraph>
</caption>
<paragraph id="594C955AFFDA3E050CC5F8E9FC9E0737" blockId="5.[151,1437,1815,2021]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="211" pageId="5" pageNumber="210">
Genitalia as shown in
<figureCitation id="C1C889DFFFDA3E060DC2F8E9FDF4011B" box="[448,516,1815,1841]" captionStart="FIGURE 8" captionStartId="11.[151,250,1420,1444]" captionTargetBox="[174,1404,193,1378]" captionTargetId="figure-17@11.[151,1436,181,1397]" captionTargetPageId="11" captionText="FIGURE 8. Male genitalia of Tembelingiola biaculeata sp. nov. in dorsal (A), ventral (B) and lateral (C) views. Scale bar: 0.5 mm." pageId="5" pageNumber="210">Fig. 8</figureCitation>
. Pseudepiphallus [epiphallus] divided into two lateral parts. Pseudepiphallic lophi [posterolateral epiphallic lobes] dorso-ventrally compressed; basal third broad, after basal third taper and bent externally, with external margins concave; apex roundly truncated. Pseudepiphallic lophi [epiphallic lateral lobes] connected by narrow bridge; pseudepiphallus with anterior part transversely straight. Pseudepiphallic parameres [ectoparameres] strongly sclerotized, elongated, not directed downwards and with hooks apically; hook with apex acute, not reaching apex of pseudepiphallic lophi. Ectophallic fold [rachis] semi-tube-like, approximately triangular with basal part having two lateral plates; ectophallic fold narrowing posteriorly into acute apex and slightly surpassing pseudepiphallic bridge. Endophallic sclerite [formula] lamellate, transverse and strongly sclerotized with lateral tongues at anterior and posterior parts. Rami fused with pseudepiphallus. Additional lateral sclerotized plate (p) elongated-pyriform with posterior apex narrow and subacute.
</paragraph>
<caption id="0D8CC5D2FFD93E050C95F918FDB20140" pageId="6" pageNumber="211" startId="6.[151,250,1766,1790]" targetBox="[162,1425,320,1732]" targetPageId="6" targetType="figure">
<paragraph id="594C955AFFD93E050C95F918FDB20140" blockId="6.[151,1437,1766,1898]" pageId="6" pageNumber="211">
<emphasis id="6B874948FFD93E050C95F918FEE600D4" bold="true" box="[151,278,1766,1790]" pageId="6" pageNumber="211">FIGURE 3.</emphasis>
Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of
<taxonomicName id="9EF3EED9FFD93E050C95F8F4FE660108" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[151,406,1802,1826]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD93E050C95F8F4FE660108" box="[151,406,1802,1826]" italics="true" pageId="6" pageNumber="211">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD93E050D98F8F4FE180108" bold="true" box="[410,488,1802,1826]" pageId="6" pageNumber="211">
<taxonomicNameLabel id="70B4F433FFD93E050D98F8F4FE180108" box="[410,488,1802,1826]" pageId="6" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A), two echeme-sequences of
<taxonomicName id="9EF3EED9FFD93E050F1CF8F4FC1C0108" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[798,1004,1802,1826]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD93E050F1CF8F4FC1C0108" box="[798,1004,1802,1826]" italics="true" pageId="6" pageNumber="211">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD93E050FF2F8F4FBCE0108" bold="true" box="[1008,1086,1802,1826]" pageId="6" pageNumber="211">
<taxonomicNameLabel id="70B4F433FFD93E050FF2F8F4FBCE0108" box="[1008,1086,1802,1826]" pageId="6" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B), continuous trill of
<taxonomicName id="9EF3EED9FFD93E050926F8F4FF3E016C" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFD93E050926F8F4FF3E016C" italics="true" pageId="6" pageNumber="211">Pteroplistes silam</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD93E050CD1F8D0FED2016C" bold="true" box="[211,290,1838,1862]" pageId="6" pageNumber="211">
<taxonomicNameLabel id="70B4F433FFD93E050CD1F8D0FED2016C" box="[211,290,1838,1862]" pageId="6" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(C), an echeme-sequence of
<taxonomicName id="9EF3EED9FFD93E050E43F8D0FC48016C" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[577,952,1838,1862]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFD93E050E43F8D0FCE1016C" box="[577,785,1838,1862]" italics="true" pageId="6" pageNumber="211">Pteroplistes lagrecai</emphasis>
<bibRefCitation id="3D62E8ABFFD93E050F14F8D0FC48016C" author="Gorochov, A. V." box="[790,952,1838,1862]" pageId="6" pageNumber="211" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
(D) and three echemes of
<taxonomicName id="9EF3EED9FFD93E0508B9F8D0FE720140" authority="Gorochov &amp; Tan, 2012" authorityName="Gorochov &amp; Tan" authorityYear="2012" class="Insecta" family="Pteroplistidae" genus="Singapuriola" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="separata">
<emphasis id="6B874948FFD93E0508B9F8D0FA6D016C" box="[1211,1437,1838,1862]" italics="true" pageId="6" pageNumber="211">Singapuriola separata</emphasis>
<bibRefCitation id="3D62E8ABFFD93E050C95F8ACFE720140" author="Gorochov, A. V. &amp; Tan, M. K." box="[151,386,1874,1898]" pageId="6" pageNumber="211" pagination="18 - 34" refId="ref8057" refString="Gorochov, A. V. &amp; Tan, M. K. (2012) New crickets of the subfamilies Phaloriinae and Pteroplistinae (Orthoptera: Gryllidae) from Singapore. Zootaxa, 3525 (1), 18 - 34. https: // doi. org / 10.11646 / zootaxa. 3525.1.2" type="journal article" year="2012">Gorochov &amp; Tan, 2012</bibRefCitation>
</taxonomicName>
(E). Scale bar: 1 s.
</paragraph>
</caption>
<caption id="0D8CC5D2FFD83E040C95FA22FD310016" pageId="7" pageNumber="212" startId="7.[151,250,1500,1524]" targetBox="[162,1425,192,1466]" targetPageId="7" targetType="figure">
<paragraph id="594C955AFFD83E040C95FA22FD310016" blockId="7.[151,1436,1500,1597]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040C95FA22FEEA03DE" bold="true" box="[151,282,1500,1524]" pageId="7" pageNumber="212">FIGURE 4.</emphasis>
Oscillograms of the syllables of the pteroplistine cricket species:
<taxonomicName id="9EF3EED9FFD83E040FFBFA23FAF103DF" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[1017,1281,1501,1525]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="biaculeata" status="sp. nov.">
<emphasis id="6B874948FFD83E040FFBFA23FAF103DF" box="[1017,1281,1501,1525]" italics="true" pageId="7" pageNumber="212">Tembelingiola biaculeata</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD83E04090CFA23FA9503DF" bold="true" box="[1294,1381,1501,1525]" pageId="7" pageNumber="212">
<taxonomicNameLabel id="70B4F433FFD83E04090CFA23FA9503DF" box="[1294,1381,1501,1525]" pageId="7" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A),
<taxonomicName id="9EF3EED9FFD83E040C95F9FFFE980033" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[151,360,1537,1561]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD83E040C95F9FFFE980033" box="[151,360,1537,1561]" italics="true" pageId="7" pageNumber="212">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD83E040D6DF9FFFE310033" bold="true" box="[367,449,1537,1561]" pageId="7" pageNumber="212">
<taxonomicNameLabel id="70B4F433FFD83E040D6DF9FFFE310033" box="[367,449,1537,1561]" pageId="7" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B),
<taxonomicName id="9EF3EED9FFD83E040DF5F9FFFD5E0033" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[503,686,1537,1561]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="silam" status="sp. nov.">
<emphasis id="6B874948FFD83E040DF5F9FFFD5E0033" box="[503,686,1537,1561]" italics="true" pageId="7" pageNumber="212">Pteroplistes silam</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD83E040EB7F9FFFCF60033" bold="true" box="[693,774,1537,1561]" pageId="7" pageNumber="212">
<taxonomicNameLabel id="70B4F433FFD83E040EB7F9FFFCF60033" box="[693,774,1537,1561]" pageId="7" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(C),
<taxonomicName id="9EF3EED9FFD83E040F3FF9FFFB4B0033" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[829,1211,1537,1561]" class="Insecta" family="Pteroplistidae" genus="Pteroplistes" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="lagrecai">
<emphasis id="6B874948FFD83E040F3FF9FFFBFF0033" box="[829,1039,1537,1561]" italics="true" pageId="7" pageNumber="212">Pteroplistes lagrecai</emphasis>
<bibRefCitation id="3D62E8ABFFD83E040815F9FFFB4B0033" author="Gorochov, A. V." box="[1047,1211,1537,1561]" pageId="7" pageNumber="212" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
(D) and
<taxonomicName id="9EF3EED9FFD83E04091AF9FFFE110017" authority="Gorochov &amp; Tan, 2012" authorityName="Gorochov &amp; Tan" authorityYear="2012" class="Insecta" family="Pteroplistidae" genus="Singapuriola" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="separata">
<emphasis id="6B874948FFD83E04091AF9FFFF000016" italics="true" pageId="7" pageNumber="212">Singapuriola separata</emphasis>
<bibRefCitation id="3D62E8ABFFD83E040CF4F9DBFE110017" author="Gorochov, A. V. &amp; Tan, M. K." box="[246,481,1573,1597]" pageId="7" pageNumber="212" pagination="18 - 34" refId="ref8057" refString="Gorochov, A. V. &amp; Tan, M. K. (2012) New crickets of the subfamilies Phaloriinae and Pteroplistinae (Orthoptera: Gryllidae) from Singapore. Zootaxa, 3525 (1), 18 - 34. https: // doi. org / 10.11646 / zootaxa. 3525.1.2" type="journal article" year="2012">Gorochov &amp; Tan, 2012</bibRefCitation>
</taxonomicName>
(E). Scale bar: 0.05 s.
</paragraph>
</caption>
<paragraph id="594C955AFFD83E040CC5F971FE6A0083" blockId="7.[151,1437,1679,2030]" box="[199,410,1679,1705]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F971FED50083" bold="true" box="[199,293,1679,1705]" pageId="7" pageNumber="212">Female.</emphasis>
Unknown
</paragraph>
<paragraph id="594C955AFFD83E040CC5F94DFC7D013F" blockId="7.[151,1437,1679,2030]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F94DFE8E00E7" bold="true" box="[199,382,1715,1741]" pageId="7" pageNumber="212">Measurements.</emphasis>
<specimenCount id="4FF55ED3FFD83E040D94F94DFE5B00E7" box="[406,427,1715,1741]" count="1" pageId="7" pageNumber="212" type="male"></specimenCount>
<typeStatus id="86482BF8FFD83E040DB0F94DFDE800E7" box="[434,536,1715,1741]" pageId="7" pageNumber="212" type="holotype">holotype</typeStatus>
: BL = 8.9; BWL = 9.6; HL = 1.9; PronL = 2.2; PronW = 2.8; FWL = 5.9; FWW = 3.3; FIIIL = 6.3; TIIIL = 4.0; TaIIIL = 3.1 •
<specimenCount id="4FF55ED3FFD83E040E9FF929FD4200DB" box="[669,690,1751,1777]" count="1" pageId="7" pageNumber="212" type="male"></specimenCount>
<typeStatus id="86482BF8FFD83E040EBBF929FCEE00DB" box="[697,798,1751,1777]" pageId="7" pageNumber="212" type="paratype">paratype</typeStatus>
: BL = 8.3; BWL = 9.4; HL = 1.3; PronL = 2.3; PronW = 2.6; FWL = 5.8; FWW = 3.1; FIIIL = 5.9; TIIIL = 3.6; TaIIIL = 3.0.
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD83E040CC5F8E1FBB70174" pageId="7" pageNumber="212" type="biology_ecology">
<paragraph id="594C955AFFD83E040CC5F8E1FBB70174" blockId="7.[151,1437,1679,2030]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F8E1FEDC0113" bold="true" box="[199,300,1823,1849]" pageId="7" pageNumber="212">Ecology.</emphasis>
This species occurs in sympatry with
<taxonomicName id="9EF3EED9FFD83E040ED1F8E1FC480113" authorityName="Tan &amp; Gorochov &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2024" box="[723,952,1823,1849]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="212" phylum="Arthropoda" rank="species" species="kabili" status="sp. nov.">
<emphasis id="6B874948FFD83E040ED1F8E1FC480113" box="[723,952,1823,1849]" italics="true" pageId="7" pageNumber="212">Tembelingiola kabili</emphasis>
</taxonomicName>
<emphasis id="6B874948FFD83E040FBCF8E1FBE70113" bold="true" box="[958,1047,1823,1849]" pageId="7" pageNumber="212">
<taxonomicNameLabel id="70B4F433FFD83E040FBCF8E1FBE70113" box="[958,1047,1823,1849]" pageId="7" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<specimenCount id="4FF55ED3FFD83E04081FF8E1FB780113" box="[1053,1160,1823,1849]" count="1" pageId="7" pageNumber="212" type="male">One male</specimenCount>
individual was found on a foliage of a low-shrub, whereas another male was found on a dead branch (
<figureCitation id="C1C889DFFFD83E040FF6F8BDFBC90174" box="[1012,1081,1859,1886]" captionStart="FIGURE 5" captionStartId="8.[151,250,1188,1212]" captionTargetBox="[151,1435,309,1165]" captionTargetId="figure-78@8.[151,1435,309,1165]" captionTargetPageId="8" captionText="FIGURE 5. Male Tembelingiola biaculeata sp. nov. in its natural habitat in Sepilok." pageId="7" pageNumber="212">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFD83E0B0CC5F899FD6A0737" lastPageId="8" lastPageNumber="213" pageId="7" pageNumber="212" type="distribution">
<paragraph id="594C955AFFD83E040CC5F899FD0D01AB" blockId="7.[151,1437,1679,2030]" box="[199,765,1895,1921]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F899FE9101AB" bold="true" box="[199,353,1895,1921]" pageId="7" pageNumber="212">Distribution.</emphasis>
EAST
<collectingCountry id="21E4D5CAFFD83E040DB6F899FDBA01AB" box="[436,586,1895,1921]" name="Malaysia" pageId="7" pageNumber="212">MALAYSIA</collectingCountry>
:
<collectingRegion id="9B375BB8FFD83E040E56F899FD6D01AB" box="[596,669,1895,1921]" country="Malaysia" name="Sabah" pageId="7" pageNumber="212">Sabah</collectingRegion>
: Sepilok
</paragraph>
<paragraph id="594C955AFFD83E040CC5F875FA6D01C7" blockId="7.[151,1437,1679,2030]" pageId="7" pageNumber="212">
<emphasis id="6B874948FFD83E040CC5F875FDCB018E" bold="true" box="[199,571,1930,1957]" pageId="7" pageNumber="212">
Calling song (
<figureCitation id="C1C889DFFFD83E040D6EF874FE3A018F" box="[364,458,1930,1957]" captionStart="FIGURE 2" captionStartId="5.[151,250,1600,1624]" captionTargetBox="[161,1426,191,1567]" captionTargetId="figure-17@5.[151,1436,181,1577]" captionTargetPageId="5" captionText="FIGURE 2. Power spectra of the calling songs of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). The grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." pageId="7" pageNumber="212">Figs 2A</figureCitation>
,
<figureCitation id="C1C889DFFFD83E040DDAF875FE0B018F" box="[472,507,1931,1957]" captionStart="FIGURE 3" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[162,1425,320,1732]" captionTargetId="figure-71@6.[151,1436,309,1743]" captionTargetPageId="6" captionText="FIGURE 3. Oscillograms of the calling songs of the pteroplistine cricket species: three echemes with different lengths of Tembelingiola biaculeata sp. nov. (A), two echeme-sequencesof Tembelingiola kabili sp. nov. (B), continuous trill of Pteroplistes silam sp. nov. (C), an echeme-sequence of Pteroplistes lagrecai Gorochov, 2004 (D) and three echemes of Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 1 s." pageId="7" pageNumber="212">3A</figureCitation>
,
<figureCitation id="C1C889DFFFD83E040E0BF874FDDE018E" box="[521,558,1930,1956]" captionStart="FIGURE 4" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[162,1425,192,1466]" captionTargetId="figure-17@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 4. Oscillograms of the syllables of the pteroplistine cricket species: Tembelingiola biaculeata sp. nov. (A), Tembelingiola kabili sp. nov. (B), Pteroplistes silam sp. nov. (C), Pteroplistes lagrecai Gorochov, 2004 (D) and Singapuriola separata Gorochov &amp; Tan, 2012 (E). Scale bar: 0.05 s." pageId="7" pageNumber="212">4A</figureCitation>
).
</emphasis>
Consists of either short or long echemes. At 31.5031.75°C, a short echeme has a duration of 0.26±
<geoCoordinate id="3CC7F39DFFD83E040DA3F851FE1701E3" box="[417,487,1967,1993]" degrees="0.02" direction="south" orientation="latitude" pageId="7" pageNumber="212" precision="555" value="-0.02">0.02 s</geoCoordinate>
(0.23
<geoCoordinate id="3CC7F39DFFD83E040E35F851FD8D01E3" box="[567,637,1967,1993]" degrees="0.29" direction="south" orientation="latitude" pageId="7" pageNumber="212" precision="555" value="-0.29">0.29 s</geoCoordinate>
) and consists of an average of nine syllables (810 syllables). A long echeme has a duration of 3.02±
<geoCoordinate id="3CC7F39DFFD83E040DFAF82DFDCB01C7" box="[504,571,2003,2029]" degrees="0.48" direction="south" orientation="latitude" pageId="7" pageNumber="212" precision="555" value="-0.48">0.48 s</geoCoordinate>
(2.50
<geoCoordinate id="3CC7F39DFFD83E040E88F82DFD3E01C7" box="[650,718,2003,2030]" degrees="3.77" direction="south" orientation="latitude" pageId="7" pageNumber="212" precision="555" value="-3.77">3.77 s</geoCoordinate>
) and consists of an average of 101 syllables (82127 syllables).
</paragraph>
<paragraph id="594C955AFFD73E0B0C95FF69FD6A0737" blockId="8.[151,1437,151,285]" pageId="8" pageNumber="213">
The silent interval between consecutive echemes is 1.7±
<geoCoordinate id="3CC7F39DFFD73E0B0F09FF69FCB1069B" box="[779,833,151,177]" degrees="0.8" direction="south" orientation="latitude" pageId="8" pageNumber="213" precision="5555" value="-0.8">0.8 s</geoCoordinate>
(0.9
<geoCoordinate id="3CC7F39DFFD73E0B0F83FF69FC48069B" box="[897,952,151,177]" degrees="3.4" direction="south" orientation="latitude" pageId="8" pageNumber="213" precision="5555" value="-3.4">3.4 s</geoCoordinate>
). Within each echeme, the average syllable duration is 18.8±2.1 ms (14.021.9 ms) and the average silent interval between consecutive syllables is 11.1±2.9 ms (7.317.3 ms). The frequency spectrum is pure-tonal and forms a harmonics, with the energy peaking at a dominant frequency of 5.66±0.09 kHz (5.535.72 kHz).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,111 @@
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Genus
<taxonomicName id="9EF3EED9FFDD3E010CECF8CCFD950166" ID-CoL="8NZX6" authority="Gorochov, 2004" authorityName="Gorochov" authorityYear="2004" box="[238,613,1842,1869]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis id="6B874948FFDD3E010CECF8CCFE6A0167" bold="true" box="[238,410,1842,1869]" italics="true" pageId="2" pageNumber="207">Tembelingiola</emphasis>
<bibRefCitation id="3D62E8ABFFDD3E010DA3F8CCFD950166" author="Gorochov, A. V." box="[417,613,1842,1868]" pageId="2" pageNumber="207" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDD3E010C95F889FDF401A5" box="[151,516,1911,1935]" pageId="2" pageNumber="207" type="reference_group">
<paragraph id="594C955AFFDD3E010C95F889FDF401A5" blockId="2.[151,516,1911,1935]" box="[151,516,1911,1935]" pageId="2" pageNumber="207">
<treatmentCitationGroup id="79E3B274FFDD3E010C95F889FDF401A5" box="[151,516,1911,1935]" pageId="2" pageNumber="207">
<treatmentCitation id="D852B34BFFDD3E010C95F889FDF401A5" author="Gorochov, A. V." box="[151,516,1911,1935]" page="384" pageId="2" pageNumber="207" year="2004">
<taxonomicName id="9EF3EED9FFDD3E010C95F889FDF401A5" ID-CoL="8NZX6" authority="Gorochov, 2004: 384" authorityName="Gorochov" authorityPageNumber="384" authorityYear="2004" box="[151,516,1911,1935]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis id="6B874948FFDD3E010C95F889FED701A5" box="[151,295,1911,1935]" italics="true" pageId="2" pageNumber="207">Tembelingiola</emphasis>
<bibRefCitation id="3D62E8ABFFDD3E010D2CF889FDF401A5" author="Gorochov, A. V." box="[302,516,1911,1935]" pageId="2" pageNumber="207" pagination="379 - 396" refId="ref7847" refString="Gorochov, A. V. (2004) Review of the subfamily Pteroplistinae (Orthoptera Gryllidae). Memorie della Societa Entomologica Italiana, 82 (2), 379 - 396." type="journal article" year="2004">Gorochov, 2004: 384</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="11E9C6D1FFDD3E010C95F843FC5F01FD" box="[151,943,1981,2007]" pageId="2" pageNumber="207" type="type_taxon">
<paragraph id="594C955AFFDD3E010C95F843FC5F01FD" blockId="2.[151,943,1981,2007]" box="[151,943,1981,2007]" pageId="2" pageNumber="207">
<emphasis id="6B874948FFDD3E010C95F843FEC501FD" bold="true" box="[151,309,1981,2007]" pageId="2" pageNumber="207">
<typeStatus id="86482BF8FFDD3E010C95F843FF2201FD" box="[151,210,1981,2007]" pageId="2" pageNumber="207">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="9EF3EED9FFDD3E010D3EF843FD6A01FD" ID-CoL="5598Q" authority="Gorochov" authorityName="Gorochov" authorityYear="2004" box="[316,666,1981,2007]" class="Insecta" family="Pteroplistidae" genus="Tembelingiola" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="6B874948FFDD3E010D3EF843FDD201FD" box="[316,546,1981,2007]" italics="true" pageId="2" pageNumber="207">Tembelingiola plana</emphasis>
Gorochov
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
</treatment>
</document>